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<document id="96FAAC35FA70B63697303A8DAA023AFE" ID-DOI="10.11646/zootaxa.5642.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="15818151" ID-ZooBank="6CAAAAD8-0347-43C8-B3E9-D7011B436C19" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1751790876585" checkinUser="plazi" docAuthor="Hopkins, Paul, Kosterin, Oleg E., Phan, Quoc Toan & Keetapithchayakul, Tosaphol Saetung" docDate="2025" docId="03F7FE24FFC3FFFBFF771BCEE4D7FDAC" docLanguage="en" docName="zootaxa.5642.5.3.pdf" docOrigin="Zootaxa 5642 (5)" docSource="https://doi.org/10.11646/zootaxa.5642.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lestes praemorsus" docType="treatment" docVersion="1" lastPageNumber="472" masterDocId="FFCE865CFFD1FFEEFFC21A21E640FF91" masterDocTitle="Taxonomic reconsideration of Lestes dorothea Fraser, 1924, L. decipiens Kirby, 1893, bona species, L. praecellens Lieftinck, 1937 and L. praemorsus Hagen in Selys, 1862 (Odonata, Lestidae)" masterLastPageNumber="475" masterPageNumber="451" pageNumber="469" updateTime="1751999258314" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:titleInfo id="53F325CEEA8EE40717EAAB35469C7697">
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<mods:title id="6CF5AD0C071C452006CC11E5C271BC67">Taxonomic reconsideration of Lestes dorothea Fraser, 1924, L. decipiens Kirby, 1893, bona species, L. praecellens Lieftinck, 1937 and L. praemorsus Hagen in Selys, 1862 (Odonata, Lestidae)</mods:title>
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<mods:roleTerm id="D3FA7C12A89104FFC2D2DFD7008F293F">Author</mods:roleTerm>
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<mods:namePart id="0350873C55745EE02DC63FF05B85784A">Hopkins, Paul</mods:namePart>
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<mods:affiliation id="493812E414E30B29C479375DCD0149E4">9 New street, Penryn, Cornwall, TR 10 8 EB, United Kingdom</mods:affiliation>
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<mods:namePart id="50803C087063356449A1AF29B491C813">Kosterin, Oleg E.</mods:namePart>
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<mods:affiliation id="7871A227D706F4093CAB23EB5F097A07">Institute of Cytology and Genetics SB RAS, Academician Lavrentyev ave. 10, Novosibirsk, 630090, Russia</mods:affiliation>
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<mods:namePart id="E60F1A7777B2BD0D41C902478632E282">Phan, Quoc Toan</mods:namePart>
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<mods:affiliation id="278958851BBE81C306907C70277B01BD">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
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<mods:namePart id="234FB3721E2614DAAE90E57647A8C5FC">Keetapithchayakul, Tosaphol Saetung</mods:namePart>
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<mods:affiliation id="81F87D53C1D78C848CE98CF9B0FB46AE">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
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<mods:title id="18150F5DD29233C538FBEC954A1ED57F">Zootaxa</mods:title>
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<mods:date id="068D56EE417BBED6A4D5F8BF52CC81D7">2025</mods:date>
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<mods:number id="298C29674D24F307568B2109122819EE">2025-06-03</mods:number>
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<mods:classification id="91A2597321E14FFD2D228BC286FADAA6">journal article</mods:classification>
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<mods:identifier id="E8639E373634BA904344A93F48DCDE83" type="DOI">10.11646/zootaxa.5642.5.3</mods:identifier>
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<mods:identifier id="339D4C033DBDF2E3603D38D67EC4C66D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F7FE24FFC3FFFBFF771BCEE4D7FDAC" LSID="urn:lsid:plazi:treatment:03F7FE24FFC3FFFBFF771BCEE4D7FDAC" httpUri="http://treatment.plazi.org/id/03F7FE24FFC3FFFBFF771BCEE4D7FDAC" lastPageId="21" lastPageNumber="472" pageId="18" pageNumber="469">
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<subSubSection id="C3441CB9FFC3FFFCFF771BCEE4F1FD98" box="[181,689,494,521]" pageId="18" pageNumber="469" type="nomenclature">
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<emphasis id="B92A9320FFC3FFFCFF771BCEE733FD98" bold="true" box="[181,371,494,521]" italics="true" pageId="18" pageNumber="469">Lestes dorothea</emphasis>
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||||
versus
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<taxonomicName id="4C5E34B1FFC3FFFCFE131BCEE4F1FD98" ID-CoL="6PQRQ" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[465,689,495,521]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFE131BCEE4F1FD98" bold="true" box="[465,689,495,521]" italics="true" pageId="18" pageNumber="469">Lestes praemorsus</emphasis>
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</taxonomicName>
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<subSubSection id="C3441CB9FFC3FFFBFF551817E4D7FDAC" lastPageId="21" lastPageNumber="472" pageId="18" pageNumber="469" type="multiple">
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<paragraph id="8BE14F32FFC3FFFCFF551817E708FC04" blockId="18.[151,1437,566,1313]" pageId="18" pageNumber="469">
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According to
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<bibRefCitation id="EFCF32C3FFC3FFFCFEF71816E7A1FDC0" author="van Tol, J." box="[309,481,566,593]" pageId="18" pageNumber="469" pagination="3 - 263" refId="ref15497" refString="van Tol, J. (1992) An annotated index to names of Odonata used in publications by M. A. Lieftinck. Zoologische Verhandelingen, 279, 3-263." type="journal article" year="1992">van Tol (1992)</bibRefCitation>
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||||
, M.A. Lieftinck never mentioned
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<taxonomicName id="4C5E34B1FFC3FFFCFCA81816E5AEFDC1" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[874,1006,566,592]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
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||||
<emphasis id="B92A9320FFC3FFFCFCA81816E5AEFDC1" box="[874,1006,566,592]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
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||||
</taxonomicName>
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in his publications. At the same time from the textual annotations and very detailed drawings in
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<bibRefCitation id="EFCF32C3FFC3FFFCFCFA187BE5B1FDE4" author="Lieftinck, M. A." box="[824,1009,602,629]" pageId="18" pageNumber="469" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
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one can see that the anal appendages of
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<taxonomicName id="4C5E34B1FFC3FFFCFF74185EE719FD09" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[182,345,639,664]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
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||||
<emphasis id="B92A9320FFC3FFFCFF74185EE719FD09" box="[182,345,639,664]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
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</taxonomicName>
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(his figs 19–20, reproduced here as
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<figureCitation id="136553B7FFC3FFFCFD34185FE522FD09" box="[758,866,638,664]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1a–b</figureCitation>
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) are very similar to
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<taxonomicName id="4C5E34B1FFC3FFFCFB8A185EE28BFD09" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[1096,1227,638,664]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
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<emphasis id="B92A9320FFC3FFFCFB8A185EE28BFD09" box="[1096,1227,638,664]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
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</taxonomicName>
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(
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<figureCitation id="136553B7FFC3FFFCFB19185FE307FD08" box="[1243,1351,638,665]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="18" pageNumber="469">Fig. 5a, c</figureCitation>
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), while substantially differ from
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<taxonomicName id="4C5E34B1FFC3FFFCFE6F1882E46CFD2D" authorityName="Kirby" authorityYear="1890" box="[429,556,674,700]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="decipens">
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<emphasis id="B92A9320FFC3FFFCFE6F1882E46CFD2D" box="[429,556,674,700]" italics="true" pageId="18" pageNumber="469">L. decipens</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFCF32C3FFC3FFFCFDF81883E4A5FD2C" author="Lieftinck, M. A." box="[570,741,674,701]" pageId="18" pageNumber="469" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949</bibRefCitation>
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: figs 18–19, reproduced here as
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<figureCitation id="136553B7FFC3FFFCFB8F1883E2F9FD2D" box="[1101,1209,674,700]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1c–d</figureCitation>
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; see also
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<figureCitation id="136553B7FFC3FFFCFAE11883E3D0FD2C" box="[1315,1424,674,701]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="18" pageNumber="469">Fig. 5e–g</figureCitation>
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). Both
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<taxonomicName id="4C5E34B1FFC3FFFCFF1618E6E735FD71" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[212,373,711,736]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
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<emphasis id="B92A9320FFC3FFFCFF1618E6E735FD71" box="[212,373,711,736]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5E34B1FFC3FFFCFE6918E6E46CFD71" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[427,556,710,736]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
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<emphasis id="B92A9320FFC3FFFCFE6918E6E46CFD71" box="[427,556,710,736]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
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</taxonomicName>
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have the paraprocts the same diverging and concealed under the cerci in dorsal view. In lateral view, the paraproct of
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<taxonomicName id="4C5E34B1FFC3FFFCFD8018CAE4A3FC95" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[578,739,747,772]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
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<emphasis id="B92A9320FFC3FFFCFD8018CAE4A3FC95" box="[578,739,747,772]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFD3118CBE520FC95" box="[755,864,746,772]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1a–b</figureCitation>
|
||||
) also has the distal part as in
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFB6918CAE36DFC95" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[1195,1325,746,772]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFB6918CAE36DFC95" box="[1195,1325,746,772]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFAFF18CBE3CEFC94" box="[1341,1422,746,773]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="18" pageNumber="469">Fig. 5c</figureCitation>
|
||||
), that is narrower than in
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFE68192EE473FCB9" authorityName="Kirby" authorityYear="1893" box="[426,563,782,808]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC3FFFCFE68192EE473FCB9" box="[426,563,782,808]" italics="true" pageId="18" pageNumber="469">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFD86192FE4F2FCB9" box="[580,690,782,808]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1c–d</figureCitation>
|
||||
), although in Lieftinck’s figures of the latter (
|
||||
<figureCitation id="136553B7FFC3FFFCFB7D192FE36CFCB9" box="[1215,1324,782,808]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1c–d</figureCitation>
|
||||
) is not as broad as in the Cambodian specimen (
|
||||
<figureCitation id="136553B7FFC3FFFCFD8B1913E4DDFCDC" box="[585,669,818,845]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="18" pageNumber="469">
|
||||
Fig.
|
||||
<quantity id="4CA6E2D7FFC3FFFCFDBC1913E4DDFCDC" box="[638,669,818,845]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="18" pageNumber="469" unit="g" value="5.0">5g</quantity>
|
||||
</figureCitation>
|
||||
). Even the lateral view of the cerci is very similar in
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFB3B1912E3DCFCDD" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1273,1436,819,844]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFB3B1912E3DCFCDD" box="[1273,1436,819,844]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFF5D1977E74DFCE1" box="[159,269,854,880]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1a–b</figureCitation>
|
||||
) and
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFE8F1976E790FCE1" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[333,464,854,880]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFE8F1976E790FCE1" box="[333,464,854,880]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFE221977E472FCE0" box="[480,562,854,881]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="18" pageNumber="469">Fig. 5c</figureCitation>
|
||||
), with the inner expansion well seen, while it is scarcely seen in
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFAD61976E3DCFCE1" authorityName="Kirby" authorityYear="1893" box="[1300,1436,854,880]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC3FFFCFAD61976E3DCFCE1" box="[1300,1436,854,880]" italics="true" pageId="18" pageNumber="469">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFF5D195BE750FC05" box="[159,272,890,916]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Figs 1c–d</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFC3FFFCFEDE195BE77BFC04" box="[284,315,890,917]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="18" pageNumber="469">
|
||||
<quantity id="4CA6E2D7FFC3FFFCFEDE195BE77BFC04" box="[284,315,890,917]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="18" pageNumber="469" unit="g" value="5.0">5g</quantity>
|
||||
</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC3FFFCFF0519BFE29FFBFD" blockId="18.[151,1437,566,1313]" pageId="18" pageNumber="469">
|
||||
These facts suggest that it is
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFDC119BEE4C4FC29" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[515,644,926,952]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFDC119BEE4C4FC29" box="[515,644,926,952]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
rather than
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFCC519BEE5CEFC29" authorityName="Kirby" authorityYear="1893" box="[775,910,926,952]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC3FFFCFCC519BEE5CEFC29" box="[775,910,926,952]" italics="true" pageId="18" pageNumber="469">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
which would more naturally be considered as a subspecies or synonym of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFE7E19E2E41DFC4D" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[444,605,963,988]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFE7E19E2E41DFC4D" box="[444,605,963,988]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
. It is difficult to evaluate the difference between
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFBB819E2E35BFC4D" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1146,1307,963,988]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFBB819E2E35BFC4D" box="[1146,1307,963,988]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFAEB19E3E3D4FC4D" box="[1321,1428,962,988]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1a–b</figureCitation>
|
||||
) and
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFF0719C6E706FB91" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[197,326,998,1024]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFF0719C6E706FB91" box="[197,326,998,1024]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC3FFFCFE9619C7E7FCFB90" box="[340,444,998,1025]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="18" pageNumber="469">Fig. 5a, c</figureCitation>
|
||||
), if any, without specimens of the former in our disposal.
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFBF519C6E298FB91" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1079,1240,999,1024]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFBF519C6E298FB91" box="[1079,1240,999,1024]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
was described by a female specimen (
|
||||
<bibRefCitation id="EFCF32C3FFC3FFFCFEB71E2BE7B5FBB5" author="Selys Longchamps, M. E. de" box="[373,501,1034,1060]" pageId="18" pageNumber="469" pagination="288 - 338" refId="ref15384" refString="Selys Longchamps, M. E. de (1862) Synopsis des Agrionines. Seconde legion: Lestes. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 13, 288-338." type="journal article" year="1862">Selys 1862</bibRefCitation>
|
||||
), but it would be more informative to compare males. So, topotypic male specimens of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFF771E0EE716FBD9" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[181,342,1071,1096]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFF771E0EE716FBD9" box="[181,342,1071,1096]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<collectingRegion id="499A81D0FFC3FFFCFE581E0FE7A9FBD9" box="[410,489,1070,1096]" country="Philippines" name="Manila" pageId="18" pageNumber="469">Manila</collectingRegion>
|
||||
should be compared with the
|
||||
<typeStatus id="54E5F190FFC3FFFCFCF51E0FE5D9FBD9" box="[823,921,1070,1096]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
|
||||
and other specimens of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFB651E0EE368FBD9" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[1191,1320,1070,1096]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFB651E0EE368FBD9" box="[1191,1320,1070,1096]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
. This is to be a matter of further study. For the time being we prefer to retain
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFCBD1E72E241FBFD" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[895,1025,1106,1132]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFCBD1E72E241FBFD" box="[895,1025,1106,1132]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
in the species rank.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC3FFFCFF051E56E4AFFAB1" blockId="18.[151,1437,566,1313]" pageId="18" pageNumber="469">
|
||||
In the current sense,
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFE711E56E476FB01" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[435,566,1142,1168]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFE711E56E476FB01" box="[435,566,1142,1168]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
co-occurs with
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFD2E1E56E534FB01" authorityName="Kirby" authorityYear="1893" box="[748,884,1142,1168]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC3FFFCFD2E1E56E534FB01" box="[748,884,1142,1168]" italics="true" pageId="18" pageNumber="469">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
in the vast area from the Western Ghats of
|
||||
<collectingCountry id="F3490FA2FFC3FFFCFAA11E57E3DDFB01" box="[1379,1437,1142,1168]" name="India" pageId="18" pageNumber="469">India</collectingCountry>
|
||||
to Indochina, without ‘transitory’ specimens known. As being nearly identical in the general appearance, except for the end of abdomen in both sexes, they obviously are closely related to each other. Nevertheless, their broad cooccurrence suggests that they are full species reproductively isolated from each other, and the strong difference in their male paraprocts may contribute to this isolation.
|
||||
</paragraph>
|
||||
<subSection id="E2D156D9FFC3FFFBFF551F52E4D7FDAC" lastPageId="21" lastPageNumber="472" pageId="18" pageNumber="469" type="multiple">
|
||||
<paragraph id="8BE14F32FFC3FFFCFF551F52E418FA1C" blockId="18.[151,600,1394,1421]" box="[151,600,1394,1421]" pageId="18" pageNumber="469">
|
||||
<heading id="D0A9F85EFFC3FFFCFF551F52E418FA1C" bold="true" box="[151,600,1394,1421]" fontSize="11" level="1" pageId="18" pageNumber="469" reason="1">
|
||||
<emphasis id="B92A9320FFC3FFFCFF551F52E418FA1C" bold="true" box="[151,600,1394,1421]" pageId="18" pageNumber="469">
|
||||
7. The problem of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFEBF1F52E418FA1C" authorityName="Lieftinck" authorityYear="1937" box="[381,600,1394,1421]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC3FFFCFEBF1F52E418FA1C" bold="true" box="[381,600,1394,1421]" italics="true" pageId="18" pageNumber="469">Lestes praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC3FFFDFF551F9BE47FFF44" blockId="18.[151,1437,1466,2033]" lastBlockId="19.[151,1436,151,213]" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
|
||||
Another potential junior synonym of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFDF91F9AE49DFA45" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[571,733,1467,1492]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFDF91F9AE49DFA45" box="[571,733,1467,1492]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2192E5BFFC3FFFCFD271F9AE560FA44" box="[741,800,1467,1493]" pageId="18" pageNumber="469" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
and/or
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFCB41F9AE5B8FA45" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[886,1016,1466,1492]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFCB41F9AE5B8FA45" box="[886,1016,1466,1492]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(depending on the resolution of their taxonomic relationships, see above) is
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFD981FFEE56BFA69" authorityName="Lieftinck" authorityYear="1937" box="[602,811,1502,1528]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC3FFFCFD981FFEE56BFA69" box="[602,811,1502,1528]" italics="true" pageId="18" pageNumber="469">Lestes praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
, described from south and south-west
|
||||
<collectingRegion id="499A81D0FFC3FFFCFB321FFEE362FA68" box="[1264,1314,1503,1529]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="469">Java</collectingRegion>
|
||||
(
|
||||
<bibRefCitation id="EFCF32C3FFC3FFFCFAF61FFFE690F98C" author="Lieftinck, M. A." pageId="18" pageNumber="469" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck 1937</bibRefCitation>
|
||||
) and also known from
|
||||
<collectingRegion id="499A81D0FFC3FFFCFE0E1C22E469F98C" box="[460,553,1539,1565]" country="Indonesia" name="Sumatera Utara" pageId="18" pageNumber="469">Sumatra</collectingRegion>
|
||||
(
|
||||
<bibRefCitation id="EFCF32C3FFC3FFFCFDF51C22E4ADF98C" author="Dow, R. A. & Choong, C. Y. & Grinang, J. & Lupiyaningdyah, P. & Ngiam, R. W. J. & Kalkman, V. J." box="[567,749,1538,1565]" pageId="18" pageNumber="469" pagination="1 - 122" refId="ref13545" refString="Dow, R. A., Choong, C. Y., Grinang, J., Lupiyaningdyah, P., Ngiam, R. W. J. & Kalkman, V. J. (2024) Checklist of the Odonata (Insecta) of Sundaland and Wallacea (Malaysia, Singapore, Brunei, Indonesia and Timor Leste). Zootaxa, 5460 (1), 1-122. https://doi.org/10.11646/zootaxa.5460.1.1" type="journal article" year="2024">
|
||||
Dow
|
||||
<emphasis id="B92A9320FFC3FFFCFDB61C22E4E4F98D" box="[628,676,1538,1564]" italics="true" pageId="18" pageNumber="469">et al</emphasis>
|
||||
. 2024
|
||||
</bibRefCitation>
|
||||
). The photographs of the
|
||||
<typeStatus id="54E5F190FFC3FFFCFBC11C23E228F98D" box="[1027,1128,1538,1564]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
|
||||
, male, and
|
||||
<typeStatus id="54E5F190FFC3FFFCFB201C23E37FF98D" box="[1250,1343,1538,1564]" pageId="18" pageNumber="469" type="allotype">allotype</typeStatus>
|
||||
, female, of this species are shown in
|
||||
<figureCitation id="136553B7FFC3FFFCFE181C07E418F9D0" box="[474,600,1574,1601]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStartId-0="21.[151,250,1403,1427]" captionStartId-1="22.[151,250,1320,1344]" captionTargetBox-0="[189,1398,597,1380]" captionTargetBox-1="[189,1398,345,1297]" captionTargetId-0="figure-204@21.[189,1398,597,1380]" captionTargetId-1="figure-105@22.[189,1398,345,1297]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 14. The allotype, female (a), and holotype, male (b), of Lestes praecellens, from S. Java, Tijamar, near Tijdaoen, 5 xi 1935. M. Bartels, preserved in the RMNH collection, and their labels (c) (photos by Charlotte Hartong and Anna Ruijbroek)." captionText-1="FIGURE 15. The end of abdomen of the holotype, male (a–c), and allotype, female (d–e) of Lestes praecellens, in dorsal (a, d), lateral (b, e) and ventral (c) views (photos by Charlotte Hartong and Anna Ruijbroek)." figureDoi-0="http://doi.org/10.5281/zenodo.15818182" figureDoi-1="http://doi.org/10.5281/zenodo.15818184" httpUri-0="https://zenodo.org/record/15818182/files/figure.png" httpUri-1="https://zenodo.org/record/15818184/files/figure.png" pageId="18" pageNumber="469">Figs 14–15</figureCitation>
|
||||
and very well correspond to the precise drawings by
|
||||
<bibRefCitation id="EFCF32C3FFC3FFFCFB7E1C07E338F9D0" author="Lieftinck, M. A." box="[1212,1400,1574,1601]" pageId="18" pageNumber="469" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck (1937)</bibRefCitation>
|
||||
. In the original description this species was compared only to specimens of “
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFC161C6AE22BF9F5" authorityName="" (Fraser" authorityYear="1933" box="[980,1131,1611,1636]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsa">
|
||||
<emphasis id="B92A9320FFC3FFFCFC161C6AE22BF9F5" box="[980,1131,1611,1636]" italics="true" pageId="18" pageNumber="469">L. praemorsa</emphasis>
|
||||
</taxonomicName>
|
||||
” from Malay Archipelago, which actually were
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFE431C4EE449F919" authorityName="Kirby" authorityYear="1893" box="[385,521,1646,1672]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC3FFFCFE431C4EE449F919" box="[385,521,1646,1672]" italics="true" pageId="18" pageNumber="469">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<bibRefCitation id="EFCF32C3FFC3FFFCFDD51C4FE490F918" author="Lieftinck, M. A." box="[535,720,1646,1673]" pageId="18" pageNumber="469" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck (1937)</bibRefCitation>
|
||||
indicated three diagnostic differences of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFB631C4EE300F919" authorityName="Lieftinck" authorityYear="1937" box="[1185,1344,1646,1672]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC3FFFCFB631C4EE300F919" box="[1185,1344,1646,1672]" italics="true" pageId="18" pageNumber="469">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFA441C4EE741F93D" authorityName="Kirby" authorityYear="1893" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC3FFFCFA441C4EE741F93D" italics="true" pageId="18" pageNumber="469">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
in the sense of this paper: (i) absence of black spots on the metopleurae (metepimeron and metepisternum) and lower surface of synthorax (
|
||||
<bibRefCitation id="EFCF32C3FFC3FFFCFDC21C97E4ECF940" author="Lieftinck, M. A." box="[512,684,1718,1745]" pageId="18" pageNumber="469" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck 1937</bibRefCitation>
|
||||
: fig. 2, reproduced here as
|
||||
<figureCitation id="136553B7FFC3FFFCFC181C97E265F941" box="[986,1061,1718,1744]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1i</figureCitation>
|
||||
;
|
||||
<figureCitation id="136553B7FFC3FFFCFBF01C97E2EEF941" box="[1074,1198,1718,1744]" captionStart="FIGURE 14" captionStartId="21.[151,250,1403,1427]" captionTargetBox="[189,1398,597,1380]" captionTargetId="figure-204@21.[189,1398,597,1380]" captionTargetPageId="21" captionText="FIGURE 14. The allotype, female (a), and holotype, male (b), of Lestes praecellens, from S. Java, Tijamar, near Tijdaoen, 5 xi 1935. M. Bartels, preserved in the RMNH collection, and their labels (c) (photos by Charlotte Hartong and Anna Ruijbroek)." figureDoi="http://doi.org/10.5281/zenodo.15818182" httpUri="https://zenodo.org/record/15818182/files/figure.png" pageId="18" pageNumber="469">Fig. 14a–b</figureCitation>
|
||||
); (ii) the black dorsal stripe on S2–S7 not constricted before the posterior widening (
|
||||
<figureCitation id="136553B7FFC3FFFCFC8D1CFBE58AF965" box="[847,970,1754,1780]" captionStart="FIGURE 14" captionStartId="21.[151,250,1403,1427]" captionTargetBox="[189,1398,597,1380]" captionTargetId="figure-204@21.[189,1398,597,1380]" captionTargetPageId="21" captionText="FIGURE 14. The allotype, female (a), and holotype, male (b), of Lestes praecellens, from S. Java, Tijamar, near Tijdaoen, 5 xi 1935. M. Bartels, preserved in the RMNH collection, and their labels (c) (photos by Charlotte Hartong and Anna Ruijbroek)." figureDoi="http://doi.org/10.5281/zenodo.15818182" httpUri="https://zenodo.org/record/15818182/files/figure.png" pageId="18" pageNumber="469">Fig. 14a–b</figureCitation>
|
||||
), and (iii) paraprocts “divaricate, abruptly turning into narrow, finger-shaped processes not visible in dorsal view” (
|
||||
<bibRefCitation id="EFCF32C3FFC3FFFCFC031CDFE2D4F889" author="Lieftinck, M. A." box="[961,1172,1790,1817]" pageId="18" pageNumber="469" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck 1937: 63</bibRefCitation>
|
||||
, fig. 3, reproduced here as
|
||||
<figureCitation id="136553B7FFC3FFFCFF771D03E6BEF8AD" box="[181,254,1826,1852]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="18" pageNumber="469">Fig. 1j</figureCitation>
|
||||
;
|
||||
<figureCitation id="136553B7FFC3FFFCFEC91D03E73FF8AC" box="[267,383,1826,1853]" captionStart="FIGURE 15" captionStartId="22.[151,250,1320,1344]" captionTargetBox="[189,1398,345,1297]" captionTargetId="figure-105@22.[189,1398,345,1297]" captionTargetPageId="22" captionText="FIGURE 15. The end of abdomen of the holotype, male (a–c), and allotype, female (d–e) of Lestes praecellens, in dorsal (a, d), lateral (b, e) and ventral (c) views (photos by Charlotte Hartong and Anna Ruijbroek)." figureDoi="http://doi.org/10.5281/zenodo.15818184" httpUri="https://zenodo.org/record/15818184/files/figure.png" pageId="18" pageNumber="469">Fig 15b–c</figureCitation>
|
||||
). It is clear that the character (iii) is the same as in
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFC6F1D02E26EF8AD" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[941,1070,1826,1852]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFC6F1D02E26EF8AD" box="[941,1070,1826,1852]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFBA11D02E343F8AD" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1123,1283,1827,1852]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC3FFFCFBA11D02E343F8AD" box="[1123,1283,1827,1852]" italics="true" pageId="18" pageNumber="469">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
. The
|
||||
<typeStatus id="54E5F190FFC3FFFCFA801D03E3DCF8AD" box="[1346,1436,1826,1852]" pageId="18" pageNumber="469" type="allotype">allotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFF7A1D66E77BF8F1" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[184,315,1862,1888]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFF7A1D66E77BF8F1" box="[184,315,1862,1888]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
exhibits the character (i) of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFD461D66E565F8F1" authorityName="Lieftinck" authorityYear="1937" box="[644,805,1862,1888]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC3FFFCFD461D66E565F8F1" box="[644,805,1862,1888]" italics="true" pageId="18" pageNumber="469">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
as having no black spots on the metapleurae (
|
||||
<figureCitation id="136553B7FFC3FFFCFAF91D67E3CFF8F0" box="[1339,1423,1862,1889]" captionStart="FIGURE 7" captionStartId="12.[151,250,1055,1079]" captionTargetBox="[151,1435,181,1031]" captionTargetId="figure-20@12.[151,1435,181,1031]" captionTargetPageId="12" captionText="FIGURE 7. Thorax of the holotype, male (a,b), and allotype, female (c,d) of Lestes dorothea, in lateral (a, c) and dorsal (b, d) views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818165" httpUri="https://zenodo.org/record/15818165/files/figure.png" pageId="18" pageNumber="469">Fig. 7c</figureCitation>
|
||||
). Perhaps the same is the
|
||||
<typeStatus id="54E5F190FFC3FFFCFE631D4BE446F815" box="[417,518,1898,1924]" pageId="18" pageNumber="469" type="holotype">holotype</typeStatus>
|
||||
, although the metepisternum is somewhat obscured by pruinescence (
|
||||
<figureCitation id="136553B7FFC3FFFCFACD1D4BE320F814" box="[1295,1376,1898,1925]" captionStart="FIGURE 7" captionStartId="12.[151,250,1055,1079]" captionTargetBox="[151,1435,181,1031]" captionTargetId="figure-20@12.[151,1435,181,1031]" captionTargetPageId="12" captionText="FIGURE 7. Thorax of the holotype, male (a,b), and allotype, female (c,d) of Lestes dorothea, in lateral (a, c) and dorsal (b, d) views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818165" httpUri="https://zenodo.org/record/15818165/files/figure.png" pageId="18" pageNumber="469">Fig. 7a</figureCitation>
|
||||
). But all our specimens of
|
||||
<emphasis id="B92A9320FFC3FFFCFEBC1DAEE446F839" box="[382,518,1934,1960]" italics="true" pageId="18" pageNumber="469">
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFEBC1DAEE442F839" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[382,514,1934,1960]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">L. dorothea</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
including the
|
||||
<specimenCount id="9D5884BBFFC3FFFCFD641DAEE557F839" box="[678,791,1934,1961]" count="2" pageId="18" pageNumber="469" type="male">two males</specimenCount>
|
||||
from
|
||||
<collectingRegion id="499A81D0FFC3FFFCFC991DAFE5E8F839" box="[859,936,1934,1960]" country="India" name="Kerala" pageId="18" pageNumber="469">Kerala</collectingRegion>
|
||||
, the Western Ghats, have two roundish black spots at the lower margin of the metepimeron (e.g.
|
||||
<figureCitation id="136553B7FFC3FFFCFD111D93E555F85C" box="[723,789,1970,1997]" captionStart="FIGURE 2" captionStartId="5.[151,250,1122,1146]" captionTargetBox="[151,1436,181,1098]" captionTargetId="figure-18@5.[151,1436,181,1098]" captionTargetPageId="5" captionText="FIGURE 2. Male of Lestes dorothea. 8 vii 2024. Seima Forest, Cambodia (PH)." figureDoi="http://doi.org/10.5281/zenodo.15818155" httpUri="https://zenodo.org/record/15818155/files/figure.png" pageId="18" pageNumber="469">Fig. 2</figureCitation>
|
||||
) and some black underside the thorax. As to the character (ii), the dorsal black stripe on S2–7 is somewhat constricted in the Cambodian and Thai males of
|
||||
<taxonomicName id="4C5E34B1FFC3FFFCFB2C1DF6E330F861" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[1262,1392,2006,2032]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC3FFFCFB2C1DF6E330F861" box="[1262,1392,2006,2032]" italics="true" pageId="18" pageNumber="469">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
but almost not constricted, as in
|
||||
<taxonomicName id="4C5E34B1FFC2FFFDFE161AB6E432FF20" authorityName="Lieftinck" authorityYear="1937" box="[468,626,151,177]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC2FFFDFE161AB6E432FF20" box="[468,626,151,177]" italics="true" pageId="19" pageNumber="470">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
, in the
|
||||
<specimenCount id="9D5884BBFFC2FFFDFD011AB6E529FF20" box="[707,873,151,177]" count="2" pageId="19" pageNumber="470" type="generic">two specimens</specimenCount>
|
||||
from
|
||||
<collectingRegion id="499A81D0FFC2FFFDFC691AB6E5B9FF20" box="[939,1017,151,177]" country="India" name="Kerala" pageId="19" pageNumber="470">Kerala</collectingRegion>
|
||||
. Perhaps the characters (i) and (ii) are not so reliable, at least in
|
||||
<taxonomicName id="4C5E34B1FFC2FFFDFE741A9AE478FF44" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[438,568,187,213]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC2FFFDFE741A9AE478FF44" box="[438,568,187,213]" italics="true" pageId="19" pageNumber="470">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF211FBAFFC2FFFDFF551D48E787F87C" ID-DOI="http://doi.org/10.5281/zenodo.15818178" ID-Zenodo-Dep="15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="19" pageNumber="470" startId="19.[151,250,1897,1921]" targetBox="[263,1324,237,1860]" targetPageId="19" targetType="figure">
|
||||
<paragraph id="8BE14F32FFC2FFFDFF551D48E787F87C" blockId="19.[151,1437,1897,2029]" pageId="19" pageNumber="470">
|
||||
<emphasis id="B92A9320FFC2FFFDFF551D48E75BF810" bold="true" box="[151,283,1897,1921]" pageId="19" pageNumber="470">FIGURE 12</emphasis>
|
||||
. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of
|
||||
<taxonomicName id="4C5E34B1FFC2FFFDFB8D1D4BE2B2F810" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[1103,1266,1897,1921]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC2FFFDFB8D1D4BE2B2F810" box="[1103,1266,1897,1921]" italics="true" pageId="19" pageNumber="470">Lestes dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and
|
||||
<taxonomicName id="4C5E34B1FFC2FFFDFD7A1DAFE574F834" authorityName="Kirby" authorityYear="1893" box="[696,820,1933,1957]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC2FFFDFD7A1DAFE574F834" box="[696,820,1933,1957]" italics="true" pageId="19" pageNumber="470">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF211FBAFFC5FFFAFF551F99E30AFA41" ID-DOI="http://doi.org/10.5281/zenodo.15818180" ID-Zenodo-Dep="15818180" box="[151,1354,1464,1489]" httpUri="https://zenodo.org/record/15818180/files/figure.png" pageId="20" pageNumber="471" startId="20.[151,250,1464,1488]" targetBox="[164,1423,181,1440]" targetPageId="20" targetType="figure">
|
||||
<paragraph id="8BE14F32FFC5FFFAFF551F99E30AFA41" blockId="20.[151,1354,1464,1489]" box="[151,1354,1464,1489]" pageId="20" pageNumber="471">
|
||||
<emphasis id="B92A9320FFC5FFFAFF551F99E759FA41" bold="true" box="[151,281,1464,1488]" pageId="20" pageNumber="471">FIGURE 13</emphasis>
|
||||
. A male of
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFE4D1F98E475FA41" authorityName="Kirby" authorityYear="1893" box="[399,565,1464,1488]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC5FFFAFE4D1F98E475FA41" box="[399,565,1464,1488]" italics="true" pageId="20" pageNumber="471">Lestes decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(species rank restored in this paper). 23 vi 2024. Siem Pang, Cambodia (PH).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BE14F32FFC5FFFAFF051C03E57CF915" blockId="20.[151,1437,1570,2029]" pageId="20" pageNumber="471">
|
||||
The
|
||||
<typeStatus id="54E5F190FFC5FFFAFF3C1C03E718F9AD" box="[254,344,1570,1596]" pageId="20" pageNumber="471" type="allotype">allotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFE471C02E467F9AD" authorityName="Lieftinck" authorityYear="1937" box="[389,551,1570,1596]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC5FFFAFE471C02E467F9AD" box="[389,551,1570,1596]" italics="true" pageId="20" pageNumber="471">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC5FFFAFDF81C03E4E1F9AD" box="[570,673,1570,1596]" captionStart="FIGURE 14" captionStartId="21.[151,250,1403,1427]" captionTargetBox="[189,1398,597,1380]" captionTargetId="figure-204@21.[189,1398,597,1380]" captionTargetPageId="21" captionText="FIGURE 14. The allotype, female (a), and holotype, male (b), of Lestes praecellens, from S. Java, Tijamar, near Tijdaoen, 5 xi 1935. M. Bartels, preserved in the RMNH collection, and their labels (c) (photos by Charlotte Hartong and Anna Ruijbroek)." figureDoi="http://doi.org/10.5281/zenodo.15818182" httpUri="https://zenodo.org/record/15818182/files/figure.png" pageId="20" pageNumber="471">Figs 14a</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFC5FFFAFD731C03E4B7F9AC" box="[689,759,1570,1597]" captionStart="FIGURE 15" captionStartId="22.[151,250,1320,1344]" captionTargetBox="[189,1398,345,1297]" captionTargetId="figure-105@22.[189,1398,345,1297]" captionTargetPageId="22" captionText="FIGURE 15. The end of abdomen of the holotype, male (a–c), and allotype, female (d–e) of Lestes praecellens, in dorsal (a, d), lateral (b, e) and ventral (c) views (photos by Charlotte Hartong and Anna Ruijbroek)." figureDoi="http://doi.org/10.5281/zenodo.15818184" httpUri="https://zenodo.org/record/15818184/files/figure.png" pageId="20" pageNumber="471">15d–e</figureCitation>
|
||||
) does not show noticeable differences from females of
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFA441C02E6BBF9F1" baseAuthorityName="Fraser" baseAuthorityYear="1933" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC5FFFAFA441C02E6BBF9F1" italics="true" pageId="20" pageNumber="471">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC5FFFAFECE1C67E716F9F1" box="[268,342,1606,1632]" captionStart="FIGURE 8" captionStartId="13.[151,250,1302,1326]" captionTargetBox="[177,1411,181,1278]" captionTargetId="figure-18@13.[177,1411,181,1278]" captionTargetPageId="13" captionText="FIGURE 8. The allotype of Lestes dorothea, from S. India, Greenfields, Sidapur. 26 iv1923, F.C. Fraser, preserved in the BMNH collection: a–dorsal general view; b–labels, c,d–end of abdomen in lateral and dorsal views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818167" httpUri="https://zenodo.org/record/15818167/files/figure.png" pageId="20" pageNumber="471">Figs 8</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFC5FFFAFEA71C67E7EBF9F1" box="[357,427,1606,1632]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="20" pageNumber="471">12a–e</figureCitation>
|
||||
), both sharing the pruinosity of the end of abdomen confined to S10, gradually tapering cerci with black tips and a concave ovipositor lower margin.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC5FFFAFF051CAEE244F8CC" blockId="20.[151,1437,1570,2029]" pageId="20" pageNumber="471">
|
||||
A sound difference between
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFDC71CAEE4E3F939" authorityName="Lieftinck" authorityYear="1937" box="[517,675,1678,1704]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC5FFFAFDC71CAEE4E3F939" box="[517,675,1678,1704]" italics="true" pageId="20" pageNumber="471">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFD1B1CAEE51AF939" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[729,858,1678,1704]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC5FFFAFD1B1CAEE51AF939" box="[729,858,1678,1704]" italics="true" pageId="20" pageNumber="471">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
is seen in the paraproct shape in lateral view, which in
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFF741C92E717F95D" authorityName="Lieftinck" authorityYear="1937" box="[182,343,1714,1740]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC5FFFAFF741C92E717F95D" box="[182,343,1714,1740]" italics="true" pageId="20" pageNumber="471">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
has a narrow base gradually tapering to the distal part (
|
||||
<bibRefCitation id="EFCF32C3FFC5FFFAFC1E1C93E2CAF95D" author="Lieftinck, M. A." box="[988,1162,1714,1740]" pageId="20" pageNumber="471" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck 1937</bibRefCitation>
|
||||
: fig. 3; reproduced here in
|
||||
<figureCitation id="136553B7FFC5FFFAFF741CF7E742F961" box="[182,258,1750,1776]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="20" pageNumber="471">Fig. 1j</figureCitation>
|
||||
;
|
||||
<figureCitation id="136553B7FFC5FFFAFED31CF7E732F960" box="[273,370,1750,1777]" captionStart="FIGURE 15" captionStartId="22.[151,250,1320,1344]" captionTargetBox="[189,1398,345,1297]" captionTargetId="figure-105@22.[189,1398,345,1297]" captionTargetPageId="22" captionText="FIGURE 15. The end of abdomen of the holotype, male (a–c), and allotype, female (d–e) of Lestes praecellens, in dorsal (a, d), lateral (b, e) and ventral (c) views (photos by Charlotte Hartong and Anna Ruijbroek)." figureDoi="http://doi.org/10.5281/zenodo.15818184" httpUri="https://zenodo.org/record/15818184/files/figure.png" pageId="20" pageNumber="471">Fig. 15b</figureCitation>
|
||||
), but a thick base abruptly narrowing to the distal part in
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFBCA1CF6E2CBF961" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[1032,1163,1750,1776]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC5FFFAFBCA1CF6E2CBF961" box="[1032,1163,1750,1776]" italics="true" pageId="20" pageNumber="471">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC5FFFAFB5E1CF7E2B3F960" box="[1180,1267,1750,1777]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="20" pageNumber="471">Figs 5c</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFC5FFFAFAC31CF7E35DF961" box="[1281,1309,1750,1776]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[295,1293,181,1908]" captionTargetId="figure-18@11.[295,1293,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. The holotype of Lestes dorothea, from S. India, Coorg, Mercara, 17 iv1923, F.C. Fraser, male, preserved in the BMNH collection: a–dorsal general view; b–labels, c-e–the end of abdomen in dorsal, ventral and lateral views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818163" httpUri="https://zenodo.org/record/15818163/files/figure.png" pageId="20" pageNumber="471">6e</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFC5FFFAFAE91CF7E315F961" box="[1323,1365,1750,1776]" captionStart="FIGURE 11" captionStartId="16.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1373]" captionTargetId="figure-20@16.[151,1436,181,1376]" captionTargetPageId="16" captionText="FIGURE 11. The anal appendages of two male specimens of Lestes dorothea from India, Western Ghats, Kerala, Iduikki District: a–c–a pond at 10.0951° N, 76.9319° E, 4 vi 2024; d–f–a pool under a rocky slope by Mankulam Power House, 10.0965° N, 76.9345° E, 19 vi 2024; a–d–dorsal view; b–e–ventral view; c–f–lateral view." figureDoi="http://doi.org/10.5281/zenodo.15818176" httpUri="https://zenodo.org/record/15818176/files/figure.png" pageId="20" pageNumber="471">11c</figureCitation>
|
||||
). This character was not discussed by
|
||||
<bibRefCitation id="EFCF32C3FFC5FFFAFE3E1CDBE4F8F885" author="Lieftinck, M. A." box="[508,696,1786,1812]" pageId="20" pageNumber="471" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck (1937)</bibRefCitation>
|
||||
. Strikingly, one of two our male specimens from
|
||||
<collectingRegion id="499A81D0FFC5FFFAFB2A1CDBE375F885" box="[1256,1333,1786,1812]" country="India" name="Kerala" pageId="20" pageNumber="471">Kerala</collectingRegion>
|
||||
, Western Ghats–that from Mankulam Power Station, of
|
||||
<date id="FFE069F2FFC5FFFAFD631D3FE55DF8A9" box="[673,797,1822,1848]" pageId="20" pageNumber="471" value="2024-06-19">19 vi 2024</date>
|
||||
, has the paraprocts (
|
||||
<figureCitation id="136553B7FFC5FFFAFBC01D3FE21CF8A9" box="[1026,1116,1822,1848]" captionStart="FIGURE 11" captionStartId="16.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1373]" captionTargetId="figure-20@16.[151,1436,181,1376]" captionTargetPageId="16" captionText="FIGURE 11. The anal appendages of two male specimens of Lestes dorothea from India, Western Ghats, Kerala, Iduikki District: a–c–a pond at 10.0951° N, 76.9319° E, 4 vi 2024; d–f–a pool under a rocky slope by Mankulam Power House, 10.0965° N, 76.9345° E, 19 vi 2024; a–d–dorsal view; b–e–ventral view; c–f–lateral view." figureDoi="http://doi.org/10.5281/zenodo.15818176" httpUri="https://zenodo.org/record/15818176/files/figure.png" pageId="20" pageNumber="471">Fig. 11f</figureCitation>
|
||||
) with a much narrower base exactly as in
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFEE81D62E789F8CD" authorityName="Lieftinck" authorityYear="1937" box="[298,457,1858,1884]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC5FFFAFEE81D62E789F8CD" box="[298,457,1858,1884]" italics="true" pageId="20" pageNumber="471">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFC5FFFAFE1A1D63E464F8CD" box="[472,548,1858,1884]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="20" pageNumber="471">Fig. 1i</figureCitation>
|
||||
)! This fact can be interpreted in two ways:
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC5FFFBFF051D47E342FF44" blockId="20.[151,1437,1570,2029]" lastBlockId="21.[151,1437,151,573]" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471">
|
||||
1. A more likely option is that the thickness of the paraproct shape base has a great intra-species variability. This matter is to be further investigated. Since the only currently recognisable difference between
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFB6D1DAAE30EF835" authorityName="Lieftinck" authorityYear="1937" box="[1199,1358,1930,1956]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC5FFFAFB6D1DAAE30EF835" box="[1199,1358,1930,1956]" italics="true" pageId="20" pageNumber="471">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFA441DAAE6BBF859" baseAuthorityName="Fraser" baseAuthorityYear="1933" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC5FFFAFA441DAAE6BBF859" italics="true" pageId="20" pageNumber="471">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
is confined to this character only, its intra-species variability would mean that
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFB431D8EE361F859" authorityName="Lieftinck" authorityYear="1937" box="[1153,1313,1966,1992]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC5FFFAFB431D8EE361F859" box="[1153,1313,1966,1992]" italics="true" pageId="20" pageNumber="471">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
is a junior subjective synonym of
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFE631DF2E465F87D" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[417,549,2002,2028]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC5FFFAFE631DF2E465F87D" box="[417,549,2002,2028]" italics="true" pageId="20" pageNumber="471">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
, and maybe also to
|
||||
<taxonomicName id="4C5E34B1FFC5FFFAFCCD1DF2E5F2F87D" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[783,946,2003,2028]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC5FFFAFCCD1DF2E5F2F87D" box="[783,946,2003,2028]" italics="true" pageId="20" pageNumber="471">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
(see above). It should be added that in his later work,
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFEDB1AB6E792FF20" author="Lieftinck, M. A." box="[281,466,151,177]" pageId="21" pageNumber="472" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
|
||||
depicted the male anal appendages of
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFC451AB6E26AFF21" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[903,1066,151,176]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFC4FFFBFC451AB6E26AFF21" box="[903,1066,151,176]" italics="true" pageId="21" pageNumber="472">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
and mentioned
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFB1D1AB6E33FFF20" authorityName="Lieftinck" authorityYear="1937" box="[1247,1407,151,177]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC4FFFBFB1D1AB6E33FFF20" box="[1247,1407,151,177]" italics="true" pageId="21" pageNumber="472">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
in another context but, unfortunately, did not compare them, in spite of the striking structural similarity.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC4FFFBFF051AFEE4F4FED0" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="472">
|
||||
2. A less likely option is that
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFDC51AFEE4E5FF68" authorityName="Lieftinck" authorityYear="1937" box="[519,677,223,249]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFC4FFFBFDC51AFEE4E5FF68" box="[519,677,223,249]" italics="true" pageId="21" pageNumber="472">L. praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
is a species different from
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFC101AFEE212FF68" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[978,1106,223,249]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC4FFFBFC101AFEE212FF68" box="[978,1106,223,249]" italics="true" pageId="21" pageNumber="472">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
and differing from it in much narrower male paraprocts. It this case its range extends from
|
||||
<collectingRegion id="499A81D0FFC4FFFBFC831B22E533FE8C" box="[833,883,259,285]" country="Indonesia" name="Jawa Timur" pageId="21" pageNumber="472">Java</collectingRegion>
|
||||
to the Indian Peninsula, where it does not exhibit the character (i) and co-occurs with
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFDE91B06E4EDFED0" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[555,685,295,321]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC4FFFBFDE91B06E4EDFED0" box="[555,685,295,321]" italics="true" pageId="21" pageNumber="472">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC4FFFBFF051B6AE468FE18" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="472">
|
||||
Hence, the actual status of
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFE331B6AE432FEF4" baseAuthorityName="Fraser" baseAuthorityYear="1933" box="[497,626,331,357]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC4FFFBFE331B6AE432FEF4" box="[497,626,331,357]" italics="true" pageId="21" pageNumber="472">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
is a matter of future research which should involve numerous specimens originating from
|
||||
<collectingCountry id="F3490FA2FFC4FFFBFE951B4EE7D1FE18" box="[343,401,367,393]" name="India" pageId="21" pageNumber="472">India</collectingCountry>
|
||||
to
|
||||
<collectingCountry id="F3490FA2FFC4FFFBFE761B4EE465FE18" box="[436,549,367,393]" name="Indonesia" pageId="21" pageNumber="472">Indonesia</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFC4FFFBFF051BB2E4D7FDAC" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="472">
|
||||
Two more species have been described by M.A. Lieftinck as related to
|
||||
<emphasis id="B92A9320FFC4FFFBFC131BB2E23BFE3D" box="[977,1147,403,428]" italics="true" pageId="21" pageNumber="472">
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFC131BB2E234FE3D" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[977,1140,403,428]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="praemorsus">L. praemorsus</taxonomicName>
|
||||
:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFB421BB2E68FFE40" authority="Lieftinck, 1932" authorityName="Lieftinck" authorityYear="1932" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="pertinax">
|
||||
<emphasis id="B92A9320FFC4FFFBFB421BB2E369FE3D" box="[1152,1321,403,428]" italics="true" pageId="21" pageNumber="472">Lestes pertinax</emphasis>
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFAED1BB2E68FFE40" author="Lieftinck, M. A." pageId="21" pageNumber="472" pagination="485 - 602" refId="ref14986" refString="Lieftinck, M. A. (1932) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part I. Descriptions of new genera and species of the families Lestidae and Agrionidae. Nova Guinea, 15 (3), 485-602." type="journal article" year="1932">Lieftinck, 1932</bibRefCitation>
|
||||
</taxonomicName>
|
||||
from New
|
||||
<collectingCountry id="F3490FA2FFC4FFFBFE971B96E7E6FE40" box="[341,422,439,465]" name="Guinea-Bissau" pageId="21" pageNumber="472">Guinea</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFE771B96E423FE40" author="Lieftinck, M. A." box="[437,611,439,465]" pageId="21" pageNumber="472" pagination="485 - 602" refId="ref14986" refString="Lieftinck, M. A. (1932) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part I. Descriptions of new genera and species of the families Lestidae and Agrionidae. Nova Guinea, 15 (3), 485-602." type="journal article" year="1932">Lieftinck 1932</bibRefCitation>
|
||||
), also known from Malay Peninsula (
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFBCD1B96E23BFE40" author="Orr, A. G." box="[1039,1147,438,465]" pageId="21" pageNumber="472" refId="ref15216" refString="Orr, A. G. (2005) Dragonflies of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 127 pp." type="book" year="2005">Orr 2005</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFB4B1B96E304FE40" author="Dow, R. A. & Choong, C. Y. & Grinang, J. & Lupiyaningdyah, P. & Ngiam, R. W. J. & Kalkman, V. J." box="[1161,1348,439,465]" pageId="21" pageNumber="472" pagination="1 - 122" refId="ref13545" refString="Dow, R. A., Choong, C. Y., Grinang, J., Lupiyaningdyah, P., Ngiam, R. W. J. & Kalkman, V. J. (2024) Checklist of the Odonata (Insecta) of Sundaland and Wallacea (Malaysia, Singapore, Brunei, Indonesia and Timor Leste). Zootaxa, 5460 (1), 1-122. https://doi.org/10.11646/zootaxa.5460.1.1" type="journal article" year="2024">
|
||||
Dow
|
||||
<emphasis id="B92A9320FFC4FFFBFB0A1B96E2BBFE40" box="[1224,1275,439,465]" italics="true" pageId="21" pageNumber="472">et al</emphasis>
|
||||
. 2024
|
||||
</bibRefCitation>
|
||||
) and
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFA441B96E7F0FE64" authority="Lieftinck, 1940" authorityName="Lieftinck" authorityYear="1940" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="praevius">
|
||||
<emphasis id="B92A9320FFC4FFFBFA441B96E6B7FE65" italics="true" pageId="21" pageNumber="472">L. praevius</emphasis>
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFEC31BFAE7F0FE64" author="Lieftinck, M. A." box="[257,432,475,501]" pageId="21" pageNumber="472" pagination="337 - 390" refId="ref15063" refString="Lieftinck, M. A. (1940) Descriptions and records of South-East Asiatic Odonata (II). Treubia, 17 (4), 337-390. https://doi.org/10.14203/treubia.v17i4.2581" type="journal article" year="1940">Lieftinck, 1940</bibRefCitation>
|
||||
</taxonomicName>
|
||||
from Borneo and Enggano (
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFD3B1BFAE5E7FE64" author="Lieftinck, M. A." box="[761,935,475,501]" pageId="21" pageNumber="472" pagination="337 - 390" refId="ref15063" refString="Lieftinck, M. A. (1940) Descriptions and records of South-East Asiatic Odonata (II). Treubia, 17 (4), 337-390. https://doi.org/10.14203/treubia.v17i4.2581" type="journal article" year="1940">Lieftinck 1940</bibRefCitation>
|
||||
). Fortunately, both they well differ from all above discussed taxa in having the cerci evenly curved and the paraprocts simply conical, as in
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFB221BDEE329FD88" authorityName="Kirby" authorityYear="1893" box="[1248,1385,511,537]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFC4FFFBFB221BDEE329FD88" box="[1248,1385,511,537]" italics="true" pageId="21" pageNumber="472">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
, but divaricate, as in
|
||||
<taxonomicName id="4C5E34B1FFC4FFFBFE8D1802E4D3FDAC" authority="(Lieftinck 1940)" baseAuthorityName="Lieftinck" baseAuthorityYear="1940" box="[335,659,547,573]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFC4FFFBFE8D1802E790FDAC" box="[335,464,547,573]" italics="true" pageId="21" pageNumber="472">L. dorothea</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFCF32C3FFC4FFFBFE1D1802E4CAFDAC" author="Lieftinck, M. A." box="[479,650,547,573]" pageId="21" pageNumber="472" pagination="337 - 390" refId="ref15063" refString="Lieftinck, M. A. (1940) Descriptions and records of South-East Asiatic Odonata (II). Treubia, 17 (4), 337-390. https://doi.org/10.14203/treubia.v17i4.2581" type="journal article" year="1940">Lieftinck 1940</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSection>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
526
data/03/F7/FE/03F7FE24FFD2FFE8FF051FBEE45AFB30.xml
Normal file
526
data/03/F7/FE/03F7FE24FFD2FFE8FF051FBEE45AFB30.xml
Normal file
|
|
@ -0,0 +1,526 @@
|
|||
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<mods:title id="AEB38156302DDD8790F9763D5D3663A3">Taxonomic reconsideration of Lestes dorothea Fraser, 1924, L. decipiens Kirby, 1893, bona species, L. praecellens Lieftinck, 1937 and L. praemorsus Hagen in Selys, 1862 (Odonata, Lestidae)</mods:title>
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<mods:name id="629ADE5C2C88C066DD4B75E23CF5BEF2" type="personal">
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<mods:namePart id="04553FE0CF1D5D13B74B1F355BB628F8">Hopkins, Paul</mods:namePart>
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<mods:namePart id="427422B5D1679DD7A297633B95FECC2E">Kosterin, Oleg E.</mods:namePart>
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<mods:affiliation id="133F673FF821E1A3F947A6BAF70F97F1">Institute of Cytology and Genetics SB RAS, Academician Lavrentyev ave. 10, Novosibirsk, 630090, Russia</mods:affiliation>
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</mods:role>
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<mods:namePart id="B2942E6258E5185109BEAC1EA7BF1B76">Phan, Quoc Toan</mods:namePart>
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<mods:affiliation id="9DB43CC303EE409AD1FEA005A91BCBE3">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
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<mods:namePart id="BDCD013384746DDC01CDB599936D4D24">Keetapithchayakul, Tosaphol Saetung</mods:namePart>
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<mods:affiliation id="BE11627746514F23C3ACB7F0004BC303">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
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<mods:title id="BFA6AEF5A2CF8FE8A53AB0F63E776A3F">Zootaxa</mods:title>
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<mods:part id="C8DF6FCCFD824BBA86E018088037F736">
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<mods:date id="C18948FB0A166D8014417D4DF747E884">2025</mods:date>
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<mods:number id="371D7AE1C2D3D4BA097CA3B8310F7F05">2025-06-03</mods:number>
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</mods:mods>
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<treatment id="03F7FE24FFD2FFE8FF051FBEE45AFB30" LSID="urn:lsid:plazi:treatment:03F7FE24FFD2FFE8FF051FBEE45AFB30" httpUri="http://treatment.plazi.org/id/03F7FE24FFD2FFE8FF051FBEE45AFB30" lastPageId="6" lastPageNumber="457" pageId="3" pageNumber="454">
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<paragraph id="8BE14F32FFD2FFEDFF051FBEE4F6FA29" blockId="3.[151,1437,151,1465]" box="[199,694,1438,1465]" pageId="3" pageNumber="454">
|
||||
<emphasis id="B92A9320FFD2FFEDFF051FBEE4F6FA29" bold="true" box="[199,694,1438,1465]" pageId="3" pageNumber="454">
|
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<taxonomicName id="4C5E34B1FFD2FFEDFF051FBEE455FA29" authority="Kirby, 1893" authorityName="Kirby" authorityYear="1893" box="[199,533,1438,1465]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFF051FBEE7C1FA28" bold="true" box="[199,385,1438,1465]" italics="true" pageId="3" pageNumber="454">Lestes decipiens</emphasis>
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFE4A1FBFE455FA29" author="Kirby, J." box="[392,533,1438,1464]" pageId="3" pageNumber="454" pagination="545 - 566" refId="ref14162" refString="Kirby, J. (1893) Catalogue of the described Neuroptera Odonata (Dragonflies) of Ceylon, with descritions of new species. The Journal of the Linnaean Society of London, Zoology, 24, 545-566. https://doi.org/10.1111/j.1096-3642.1894.tb02495.x" type="journal article" year="1893">Kirby, 1893</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, bona species
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSection id="E2D156D9FFD2FFE8FF551FC6E204FF68" lastPageId="6" lastPageNumber="457" pageId="3" pageNumber="454" type="multiple">
|
||||
<subSubSection id="C3441CB9FFD2FFEDFF551FC6E7F4F990" box="[151,436,1511,1537]" pageId="3" pageNumber="454" type="discussion">
|
||||
<paragraph id="8BE14F32FFD2FFEDFF551FC6E7F4F990" blockId="3.[151,436,1511,1537]" box="[151,436,1511,1537]" pageId="3" pageNumber="454">
|
||||
<heading id="D0A9F85EFFD2FFEDFF551FC6E7F4F990" bold="true" box="[151,436,1511,1537]" fontSize="11" level="1" pageId="3" pageNumber="454" reason="1">
|
||||
<emphasis id="B92A9320FFD2FFEDFF551FC6E7F4F990" bold="true" box="[151,436,1511,1537]" pageId="3" pageNumber="454">4. Cambodian findings</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3441CB9FFD2FFE8FF551C11E204FF68" lastPageId="6" lastPageNumber="457" pageId="3" pageNumber="454" type="multiple">
|
||||
<paragraph id="8BE14F32FFD2FFEDFF551C11E48EF802" blockId="3.[151,1436,1584,1940]" pageId="3" pageNumber="454">
|
||||
Of the above mentioned taxa of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFDCA1C10E4C6F9DA" box="[520,646,1585,1611]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="undetermined">
|
||||
<emphasis id="B92A9320FFD2FFEDFDCA1C10E40EF9DB" box="[520,590,1585,1610]" italics="true" pageId="3" pageNumber="454">Lestes</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
, only
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFD0F1C10E5A3F9DB" authorityName="Kirby" authorityYear="1894" box="[717,995,1584,1610]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="subSpecies" species="praemorsus" subSpecies="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFD0F1C10E5A3F9DB" box="[717,995,1584,1610]" italics="true" pageId="3" pageNumber="454">L. praemorsus decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
was hitherto reported from
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFAE01C11E3D8F9DB" box="[1314,1432,1584,1610]" name="Cambodia" pageId="3" pageNumber="454">Cambodia</collectingCountry>
|
||||
, for the first time by Roland
|
||||
<emphasis id="B92A9320FFD2FFEDFE141C77E448F9FE" box="[470,520,1621,1647]" italics="true" pageId="3" pageNumber="454">et al</emphasis>
|
||||
. (2011), not identified to subspecies, later in many particular publications by the second author (further in the text OEK).
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFDA71C58E498F902" box="[613,728,1657,1683]" name="Cambodia" pageId="3" pageNumber="454">Cambodia</collectingCountry>
|
||||
is an Indochinese country, the flora and fauna of which is still rather poorly studied as compared to the adjacent
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFD0C1CBFE572F929" box="[718,818,1694,1720]" name="Thailand" pageId="3" pageNumber="454">Thailand</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFCA81CBFE58DF929" box="[874,973,1694,1720]" name="Vietnam" pageId="3" pageNumber="454">Vietnam</collectingCountry>
|
||||
. However, for the second decade of XXI century, focused studies of the fauna of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFD941CE2E4F5F94C" box="[598,693,1731,1757]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||||
of
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFD181CE2E510F94C" box="[730,848,1731,1757]" name="Cambodia" pageId="3" pageNumber="454">Cambodia</collectingCountry>
|
||||
, mostly by OEK, have resulted in 204 species (three of which are still unpublished and seven not identified or described) registered in that country with the number of species expected to approach 300 (
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFDDD1D2DE485F8B7" author="Kosterin, O. E." box="[543,709,1804,1830]" pageId="3" pageNumber="454" pagination="1 - 19" refId="ref14740" refString="Kosterin, O. E. (2024) Photographic recording dragonflies and damselflies (Odonata) of Cambodia by public over a decade. International Dragonfly Fund Report, 189, 1-19." type="journal article" year="2024">Kosterin 2024</bibRefCitation>
|
||||
). Since about one third of Cambodian species of dragonflies and damselflies had still to be revealed, the first author (further in the text
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFC691D11E590F8DB" box="[939,976,1840,1866]" name="Philippines" pageId="3" pageNumber="454">PH</collectingCountry>
|
||||
) undertook two expeditions to the northeastern and eastern
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFEB91D74E7AEF8FE" box="[379,494,1877,1903]" name="Cambodia" pageId="3" pageNumber="454">Cambodia</collectingCountry>
|
||||
in 2022 (
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFDA21D74E544F8FE" author="Hopkins, P." box="[608,772,1877,1903]" pageId="3" pageNumber="454" pagination="36 - 56" refId="ref13819" refString="Hopkins, P. (2024) A first survey of the dragonflies (Odonata) of Siem Pang Wildlife Sanctuary, northeast Cambodia. Cambodian Journal of Natural History, 2024 (1), 36-56." type="journal article" year="2024">Hopkins 2024</bibRefCitation>
|
||||
) and 2024. During the latter one,
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFB531D77E355F8FE" authorityName="Fraser" authorityYear="1924" box="[1169,1301,1877,1903]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD2FFEDFB531D77E355F8FE" box="[1169,1301,1877,1903]" italics="true" pageId="3" pageNumber="454">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
was found, which had not been reported for
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFDC71D58E438F802" box="[517,632,1913,1939]" name="Cambodia" pageId="3" pageNumber="454">Cambodia</collectingCountry>
|
||||
before.
|
||||
</paragraph>
|
||||
<caption id="DF211FBAFFD5FFEAFF551C45E5B3F875" ID-DOI="http://doi.org/10.5281/zenodo.15818153" ID-Zenodo-Dep="15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="4" pageNumber="455" startId="4.[151,250,1636,1660]" targetBox="[400,1192,184,1607]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8BE14F32FFD5FFEAFF551C45E5B3F875" blockId="4.[151,1437,1636,2020]" pageId="4" pageNumber="455">
|
||||
<emphasis id="B92A9320FFD5FFEAFF551C45E750F9ED" bold="true" box="[151,272,1636,1660]" pageId="4" pageNumber="455">FIGURE 1</emphasis>
|
||||
. Drawings of
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFE681C44E45EF9ED" box="[426,542,1637,1660]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="undetermined">
|
||||
<emphasis id="B92A9320FFD5FFEAFE681C44E7A9F9ED" box="[426,489,1637,1660]" italics="true" pageId="4" pageNumber="455">Lestes</emphasis>
|
||||
spp.
|
||||
</taxonomicName>
|
||||
(all males but i-right female) from literature, with original identifications: a, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFA9F1C44E7CEF931" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="praemorsus" subSpecies="praemorsus">
|
||||
<emphasis id="B92A9320FFD5FFEAFA9F1C44E7CEF931" italics="true" pageId="4" pageNumber="455">Lestes praemorsus praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
”, Philippines, Luzon, Bilucao (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFD131CA9E52DF930" author="Lieftinck, M. A." box="[721,877,1672,1697]" pageId="4" pageNumber="455" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949</bibRefCitation>
|
||||
: fig. 19); b, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFC351CA8E374F931" box="[1015,1332,1673,1696]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="praemorsus" subSpecies="praemorsus">
|
||||
<emphasis id="B92A9320FFD5FFEAFC351CA8E374F931" box="[1015,1332,1673,1696]" italics="true" pageId="4" pageNumber="455">Lestes praemorsus praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
”, N. New Guinea, Hollandia (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFEA21C8DE7BDF954" author="Lieftinck, M. A." box="[352,509,1708,1733]" pageId="4" pageNumber="455" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949</bibRefCitation>
|
||||
: fig. 20); c, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFD451C8CE5EDF955" authorityName="Kirby" authorityYear="1894" box="[647,941,1708,1732]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="praemorsus" subSpecies="decipiens">
|
||||
<emphasis id="B92A9320FFD5FFEAFD451C8CE5EDF955" box="[647,941,1708,1732]" italics="true" pageId="4" pageNumber="455">Lestes praemorsus decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
”, India, Coorg, Gonocopal (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFB101C8DE32EF954" author="Lieftinck, M. A." box="[1234,1390,1708,1733]" pageId="4" pageNumber="455" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949</bibRefCitation>
|
||||
: fig. 17); d, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFF321CF0E458F979" authorityName="Kirby" authorityYear="1894" box="[240,536,1744,1768]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="praemorsus" subSpecies="decipiens">
|
||||
<emphasis id="B92A9320FFD5FFEAFF321CF0E458F979" box="[240,536,1744,1768]" italics="true" pageId="4" pageNumber="455">Lestes praemorsus decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
”, Indonesia, Java, Buitenzorg (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFCA21CF1E5BDF978" author="Lieftinck, M. A." box="[864,1021,1744,1769]" pageId="4" pageNumber="455" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949</bibRefCitation>
|
||||
: fig. 18); e, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFB4E1CF0E370F979" authorityName="Fraser" authorityYear="1924" box="[1164,1328,1744,1768]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD5FFEAFB4E1CF0E370F979" box="[1164,1328,1744,1768]" italics="true" pageId="4" pageNumber="455">Lestes dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
” from the original description (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFEAF1CD4E7ABF89C" author="Fraser, F. C." box="[365,491,1781,1805]" pageId="4" pageNumber="455" pagination="423 - 522" refId="ref13634" refString="Fraser, F. C. (1924) A survey of the odonate (dragonfly) fauna of western India with special remarks on the genera of Macromia and Idionyx and descriptions of thirty new species. Records of the Indian Museum, 26 (5), 423-522, pls. XXV - XXVII. https://doi.org/10.26515/rzsi/v26/i5/1924/162660" type="journal article" year="1924">Fraser 1924</bibRefCitation>
|
||||
: plate XXVI, fig. 8); f, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFD2F1CD4E5E2F89D" authorityName="Hagen" authorityYear="1862" box="[749,930,1781,1804]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="praemorsa">
|
||||
<emphasis id="B92A9320FFD5FFEAFD2F1CD4E5E2F89D" box="[749,930,1781,1804]" italics="true" pageId="4" pageNumber="455">Lestes praemorsa</emphasis>
|
||||
</taxonomicName>
|
||||
” (in fact
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFBC51CD4E2E9F89D" authorityName="Fraser" authorityYear="1924" box="[1031,1193,1780,1804]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD5FFEAFBC51CD4E2E9F89D" box="[1031,1193,1780,1804]" italics="true" pageId="4" pageNumber="455">Lestes dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
) (Fraser 1934: fig. 12); g, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFF7F1D38E720F8A1" authorityName="Fraser" authorityYear="1924" box="[189,352,1816,1840]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD5FFEAFF7F1D38E720F8A1" box="[189,352,1816,1840]" italics="true" pageId="4" pageNumber="455">Lestes dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
” (in fact
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFE071D38E42CF8A1" authorityName="Kirby" authorityYear="1893" box="[453,620,1816,1840]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD5FFEAFE071D38E42CF8A1" box="[453,620,1816,1840]" italics="true" pageId="4" pageNumber="455">Lestes decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFAD81D38E3CCF8A0" box="[1306,1420,1817,1841]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="undefined-1">
|
||||
<emphasis id="B92A9320FFD5FFEAFAD81D38E319F8A1" box="[1306,1369,1817,1840]" italics="true" pageId="4" pageNumber="455">Lestes</emphasis>
|
||||
sp.1
|
||||
</taxonomicName>
|
||||
”, Laos, Champasak Province, Bolaven Plateau, Paksong (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFD0A1D1DE5B4F8C4" author="Yokoi, N. & Souphanthong, V." box="[712,1012,1852,1877]" pageId="4" pageNumber="455" refId="ref15573" refString="Yokoi, N. & Souphanthong, V. (2014) A List of Lao Dragonflies. Kyoei Printing Co Ltd., Koriyama, 110 pp." type="book" year="2014">Yokoi & Souphanthong 2014</bibRefCitation>
|
||||
: fig. 5); i-j, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFBB11D1CE36DF8C5" authorityName="Lieftinck" authorityYear="1937" box="[1139,1325,1852,1876]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFD5FFEAFBB11D1CE36DF8C5" box="[1139,1325,1852,1876]" italics="true" pageId="4" pageNumber="455">Lestes praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFC4E1D41E269F8E8" author="Lieftinck, M. A." box="[908,1065,1888,1913]" pageId="4" pageNumber="455" pagination="55 - 119" refId="ref15035" refString="Lieftinck, M. A. (1937) Descriptions and records of South-East Asiatic Odonata. Treubia, 16 (1), 55-119." type="journal article" year="1937">Lieftinck 1937</bibRefCitation>
|
||||
: figs 2–3); k–l, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFB1F1D40E6B7F80D" authorityName="Kirby" authorityYear="1894" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="subSpecies" species="praemorsus" subSpecies="decipiens">
|
||||
<emphasis id="B92A9320FFD5FFEAFB1F1D40E6B7F80D" italics="true" pageId="4" pageNumber="455">Lestes praemorsus decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
”, Taiwan (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFEA51DA5E7B9F80D" author="Asahina, S." box="[359,505,1924,1948]" pageId="4" pageNumber="455" pagination="2 - 13" refId="ref13478" refString="Asahina, S. (1985) A list of the Odonata recorded from Thailand, Part VIII. Lestidae. Cho Cho, 8 (8), 2-13." type="journal article" year="1985">Asahina 1985</bibRefCitation>
|
||||
: figs 25–26); m, “
|
||||
<taxonomicName id="4C5E34B1FFD5FFEAFD761DA4E240F80C" authority="" (Orr 2005: 28)" authorityName="" (Orr" authorityYear="2005" baseAuthorityName="Orr" baseAuthorityPageNumber="28" baseAuthorityYear="2005" box="[692,1024,1924,1949]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD5FFEAFD761DA4E516F80D" box="[692,854,1924,1948]" italics="true" pageId="4" pageNumber="455">Lestes dorothea</emphasis>
|
||||
” (
|
||||
<bibRefCitation id="EFCF32C3FFD5FFEAFCAD1DA4E5B9F80C" author="Orr, A. G." box="[879,1017,1924,1949]" pageId="4" pageNumber="455" refId="ref15216" refString="Orr, A. G. (2005) Dragonflies of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 127 pp." type="book" year="2005">Orr 2005: 28</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
; a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF211FBAFFD4FFEBFF551E43E588FBEB" ID-DOI="http://doi.org/10.5281/zenodo.15818155" ID-Zenodo-Dep="15818155" box="[151,968,1122,1146]" httpUri="https://zenodo.org/record/15818155/files/figure.png" pageId="5" pageNumber="456" startId="5.[151,250,1122,1146]" targetBox="[151,1436,181,1098]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8BE14F32FFD4FFEBFF551E43E588FBEB" blockId="5.[151,968,1122,1146]" box="[151,968,1122,1146]" pageId="5" pageNumber="456">
|
||||
<emphasis id="B92A9320FFD4FFEBFF551E43E74DFBEB" bold="true" box="[151,269,1122,1146]" pageId="5" pageNumber="456">FIGURE 2</emphasis>
|
||||
. Male of
|
||||
<emphasis id="B92A9320FFD4FFEBFEAD1E42E457FBEB" box="[367,535,1122,1146]" italics="true" pageId="5" pageNumber="456">
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFEAD1E42E453FBEB" authorityName="" (Orr" authorityYear="2005" box="[367,531,1122,1146]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="dorothea">Lestes dorothea</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
8 vii 2024. Seima Forest, Cambodia (PH).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BE14F32FFD4FFEBFF051EEFE596FA75" blockId="5.[151,1437,1230,2012]" pageId="5" pageNumber="456">
|
||||
On
|
||||
<date id="FFE069F2FFD4FFEBFF311EEFE724FB79" box="[243,356,1230,1256]" pageId="5" pageNumber="456" value="2024-06-08">8 vi 2024</date>
|
||||
, whilst exploring an area of the Seima Forest in Modulkiri Province,
|
||||
<collectingCountry id="F3490FA2FFD4FFEBFB451EEFE2BDFB79" box="[1159,1277,1230,1256]" name="Cambodia" pageId="5" pageNumber="456">Cambodia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F3490FA2FFD4FFEBFAC81EEFE36EFB79" box="[1290,1326,1230,1256]" name="Philippines" pageId="5" pageNumber="456">PH</collectingCountry>
|
||||
observed up to
|
||||
<specimenCount id="9D5884BBFFD4FFEBFF1F1ED3E775FA9D" box="[221,309,1266,1292]" count="4" pageId="5" pageNumber="456" type="male">4 males</specimenCount>
|
||||
of a
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFEB51ED2E457FA9D" box="[375,535,1266,1292]" pageId="5" pageNumber="456">
|
||||
<emphasis id="B92A9320FFD4FFEBFEB51ED2E7FDFA9D" box="[375,445,1267,1292]" italics="true" pageId="5" pageNumber="456">Lestes</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
(241078616 and 243313496). No collecting was made. However, preliminary identification of the photographs and video taken based on the pruinescence of the abdominal tip according to Kompier (2025) resulted in
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFE121F1AE4C1FAC5" authorityName="" (Orr" authorityYear="2005" box="[464,641,1338,1364]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD4FFEBFE121F1AE4C1FAC5" box="[464,641,1338,1364]" italics="true" pageId="5" pageNumber="456">Lestes dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD4FFEBFD521F1BE4B6FAC5" box="[656,758,1338,1364]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="5.[151,250,1122,1146]" captionStartId-1="7.[151,250,1545,1569]" captionTargetBox-0="[151,1436,181,1098]" captionTargetBox-1="[151,1435,237,1521]" captionTargetId-0="figure-18@5.[151,1436,181,1098]" captionTargetId-1="figure-51@7.[151,1435,237,1521]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="FIGURE 2. Male of Lestes dorothea. 8 vii 2024. Seima Forest, Cambodia (PH)." captionText-1="FIGURE 3. Another male of Lestes dorothea. 8 vii June 2024. Seima Forest, Cambodia (PH)." figureDoi-0="http://doi.org/10.5281/zenodo.15818155" figureDoi-1="http://doi.org/10.5281/zenodo.15818157" httpUri-0="https://zenodo.org/record/15818155/files/figure.png" httpUri-1="https://zenodo.org/record/15818157/files/figure.png" pageId="5" pageNumber="456">Figs 2–3</figureCitation>
|
||||
), a species not previously recorded in
|
||||
<collectingCountry id="F3490FA2FFD4FFEBFB661F1BE35BFAC5" box="[1188,1307,1338,1364]" name="Cambodia" pageId="5" pageNumber="456">Cambodia</collectingCountry>
|
||||
. They were found in a flooded area adjacent to the N76 road, less than
|
||||
<quantity id="4CA6E2D7FFD4FFEBFCF11F7FE53BFAE9" box="[819,891,1374,1400]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="456" unit="km" value="10.0">10 km</quantity>
|
||||
apart from the border with
|
||||
<collectingCountry id="F3490FA2FFD4FFEBFB761F7FE353FAE9" box="[1204,1299,1374,1400]" name="Vietnam" pageId="5" pageNumber="456">Vietnam</collectingCountry>
|
||||
(
|
||||
<geoCoordinate id="EE6A29F5FFD4FFEBFAE01F7FE3D8FAE9" box="[1314,1432,1374,1400]" degrees="12.190" direction="north" orientation="latitude" pageId="5" pageNumber="456" precision="55" value="12.19">12.190º N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD4FFEBFF551FA3E75EFA0D" box="[151,286,1410,1436]" degrees="106.996" direction="east" orientation="longitude" pageId="5" pageNumber="456" precision="55" value="106.996">106.996 º E</geoCoordinate>
|
||||
). The habitat was within evergreen dipterocarp forest and shaded, although some sunshine reached the forest floor in places. There were a deep area of stagnant water with both muddy margins and rich vegetation and an area of slow flowing flood water emanating from a small stream (
|
||||
<figureCitation id="136553B7FFD4FFEBFCB51FEBE589FA75" box="[887,969,1482,1508]" captionStart="FIGURE 4" captionStartId="8.[151,250,1552,1576]" captionTargetBox="[151,1436,237,1528]" captionTargetId="figure-47@8.[151,1436,237,1528]" captionTargetPageId="8" captionText="FIGURE 4. Habitats of (a) Lestes dorothea. Seima Forest, Cambodia, and (b) of L. dorothea, L. decipiens (species rank restored in this paper) and L. nigriceps; 15 km north of Sen Momorom, Cambodia (b) (PH)." figureDoi="http://doi.org/10.5281/zenodo.15818159" httpUri="https://zenodo.org/record/15818159/files/figure.png" pageId="5" pageNumber="456">Fig. 4a</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD4FFE8FF051FCFE204FF68" blockId="5.[151,1437,1230,2012]" lastBlockId="6.[151,1436,151,249]" lastPageId="6" lastPageNumber="457" pageId="5" pageNumber="456">
|
||||
This finding inspired OEK to reconsider his Cambodian specimens (
|
||||
<specimenCount id="9D5884BBFFD4FFEBFC011FCFE26AF999" box="[963,1066,1518,1544]" count="12" pageId="5" pageNumber="456" type="male">12 males</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D5884BBFFD4FFEBFBF71FCFE2D8F999" box="[1077,1176,1518,1544]" count="1" pageId="5" pageNumber="456" type="female">1 female</specimenCount>
|
||||
) collected in preceding years and previously identified as
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFDDA1C32E48FF9BD" authorityName="Kirby" authorityYear="1893" box="[536,719,1554,1580]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD4FFEBFDDA1C32E48FF9BD" box="[536,719,1554,1580]" italics="true" pageId="5" pageNumber="456">Lestes decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
and available for examination (seven more such specimens had been earlier donated to Naturalis Biodiversity Center, Leiden and so could not be checked). They originated from eight localities in
|
||||
<collectingRegion id="499A81D0FFD4FFEBFEA01C7BE79CF9E5" box="[354,476,1626,1652]" country="Cambodia" name="Koh Kong" pageId="5" pageNumber="456">Koh Kong</collectingRegion>
|
||||
, Preah Sihanouk,
|
||||
<collectingRegion id="499A81D0FFD4FFEBFD6B1C7BE546F9E5" box="[681,774,1626,1652]" country="Cambodia" name="Kampot" pageId="5" pageNumber="456">Kampot</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD4FFEBFCD01C7BE5DDF9E5" box="[786,925,1626,1652]" country="Cambodia" name="Stung Treng" pageId="5" pageNumber="456">Stung Treng</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="499A81D0FFD4FFEBFC171C7BE215F9E5" box="[981,1109,1626,1652]" country="Cambodia" name="Mondolkiri" pageId="5" pageNumber="456">Mondulkiri</collectingRegion>
|
||||
Provinces of
|
||||
<collectingCountry id="F3490FA2FFD4FFEBFB311C7BE32AF9E5" box="[1267,1386,1626,1652]" name="Cambodia" pageId="5" pageNumber="456">Cambodia</collectingCountry>
|
||||
. For 10 of them that identification was confirmed but
|
||||
<specimenCount id="9D5884BBFFD4FFEBFD771C5FE559F909" box="[693,793,1662,1688]" count="1" pageId="5" pageNumber="456" type="male">one male</specimenCount>
|
||||
turned out to be in fact
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFBDD1C5EE2E0F909" authorityName="" (Orr" authorityYear="2005" box="[1055,1184,1662,1688]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD4FFEBFBDD1C5EE2E0F909" box="[1055,1184,1662,1688]" italics="true" pageId="5" pageNumber="456">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
, which is superficially similar to
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFEC11C82E7C8F92D" authorityName="Kirby" authorityYear="1893" box="[259,392,1698,1724]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD4FFEBFEC11C82E7C8F92D" box="[259,392,1698,1724]" italics="true" pageId="5" pageNumber="456">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
but differs strongly from it by the paraprocts(
|
||||
<figureCitation id="136553B7FFD4FFEBFCBA1C83E5A3F92C" box="[888,995,1698,1725]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="5" pageNumber="456">Fig.5a–d</figureCitation>
|
||||
; note that in this figure, the pruinescence is absent as washed off by acetone). It was collected on
|
||||
<date id="FFE069F2FFD4FFEBFCCE1CE7E5C4F971" box="[780,900,1734,1760]" pageId="5" pageNumber="456" value="2018-06-27">
|
||||
27 vi
|
||||
<quantity id="4CA6E2D7FFD4FFEBFC8E1CE7E5C4F971" box="[844,900,1734,1760]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="5" pageNumber="456" unit="in" value="2018.0">2018</quantity>
|
||||
</date>
|
||||
in a large open swamp with thick sedge,
|
||||
<quantity id="4CA6E2D7FFD4FFEBFA971CE7E3DCF971" box="[1365,1436,1734,1761]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="456" unit="km" value="15.0">15 km</quantity>
|
||||
NNE of Sen Momorom (
|
||||
<geoCoordinate id="EE6A29F5FFD4FFEBFE6C1CCBE463F895" box="[430,547,1770,1797]" degrees="12.550" direction="north" orientation="latitude" pageId="5" pageNumber="456" precision="55" value="12.55">12.550º N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD4FFEBFDED1CCBE4E9F895" box="[559,681,1770,1797]" degrees="107.258" direction="east" orientation="longitude" pageId="5" pageNumber="456" precision="55" value="107.258">107.258º E</geoCoordinate>
|
||||
) (
|
||||
<figureCitation id="136553B7FFD4FFEBFD031CCBE554F895" box="[705,788,1770,1796]" captionStart="FIGURE 4" captionStartId="8.[151,250,1552,1576]" captionTargetBox="[151,1436,237,1528]" captionTargetId="figure-47@8.[151,1436,237,1528]" captionTargetPageId="8" captionText="FIGURE 4. Habitats of (a) Lestes dorothea. Seima Forest, Cambodia, and (b) of L. dorothea, L. decipiens (species rank restored in this paper) and L. nigriceps; 15 km north of Sen Momorom, Cambodia (b) (PH)." figureDoi="http://doi.org/10.5281/zenodo.15818159" httpUri="https://zenodo.org/record/15818159/files/figure.png" pageId="5" pageNumber="456">Fig. 4b</figureCitation>
|
||||
), designated as locality M
|
||||
<quantity id="4CA6E2D7FFD4FFEBFBF91CCBE233F895" box="[1083,1139,1770,1796]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="5" pageNumber="456" unit="in" value="12.0">12 in</quantity>
|
||||
<bibRefCitation id="EFCF32C3FFD4FFEBFBB81CCBE379F895" author="Kosterin, O. E." box="[1146,1337,1770,1796]" pageId="5" pageNumber="456" pagination="185 - 224" refId="ref14616" refString="Kosterin, O. E. (2020 a) Miscellaneous faunal data on Odonata of Cambodia. International Dragonfly Fund Report, 154, 185-224." type="journal article" year="2020">Kosterin (2020a)</bibRefCitation>
|
||||
and also considered in
|
||||
<bibRefCitation id="EFCF32C3FFD4FFEBFEF41D2FE7A9F8B9" author="Kosterin, O. E." box="[310,489,1806,1832]" pageId="5" pageNumber="456" pagination="561 - 575" refId="ref14481" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561-575. https://doi.org/10.11646/zootaxa.4526.4.8" type="journal article" year="2018">Kosterin (2018)</bibRefCitation>
|
||||
(It should be noted that both papers (
|
||||
<bibRefCitation id="EFCF32C3FFD4FFEBFC501D2FE277F8B9" author="Kosterin, O. E." box="[914,1079,1806,1832]" pageId="5" pageNumber="456" pagination="561 - 575" refId="ref14481" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561-575. https://doi.org/10.11646/zootaxa.4526.4.8" type="journal article" year="2018">Kosterin 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD4FFEBFB861D2FE2CAF8B9" author="Kosterin, O. E." box="[1092,1162,1806,1832]" pageId="5" pageNumber="456" pagination="185 - 224" refId="ref14616" refString="Kosterin, O. E. (2020 a) Miscellaneous faunal data on Odonata of Cambodia. International Dragonfly Fund Report, 154, 185-224." type="journal article" year="2020">2020a</bibRefCitation>
|
||||
) unfortunately provided somewhat wrong coordinates (
|
||||
<geoCoordinate id="EE6A29F5FFD4FFEBFE361D13E435F8DD" box="[500,629,1842,1869]" degrees="12.5771" direction="north" orientation="latitude" pageId="5" pageNumber="456" precision="5" value="12.5771">12.5771º N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD4FFEBFD401D13E54BF8DD" box="[642,779,1842,1869]" degrees="107.2587" direction="east" orientation="longitude" pageId="5" pageNumber="456" precision="5" value="107.2587">107.2587º E</geoCoordinate>
|
||||
) for this place, which comprise an error of
|
||||
<quantity id="4CA6E2D7FFD4FFEBFAC11D13E313F8DD" box="[1283,1363,1842,1868]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="456" unit="km" value="2.8">2.8 km</quantity>
|
||||
to the north). Along with the single male specimen of
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFD661D76E564F8E1" authorityName="" (Orr" authorityYear="2005" box="[676,804,1878,1904]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD4FFEBFD661D76E564F8E1" box="[676,804,1878,1904]" italics="true" pageId="5" pageNumber="456">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD4FFEBFCF01D77E5DFF8E0" box="[818,927,1878,1905]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="5" pageNumber="456">Fig. 5a–d</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D5884BBFFD4FFEBFC701D77E258F8E1" box="[946,1048,1878,1904]" count="2" pageId="5" pageNumber="456" type="male">two male</specimenCount>
|
||||
specimens of the true
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFACD1D76E3D5F8E1" authorityName="Kirby" authorityYear="1893" box="[1295,1429,1878,1904]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD4FFEBFACD1D76E3D5F8E1" box="[1295,1429,1878,1904]" italics="true" pageId="5" pageNumber="456">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
, were collected. So, both species co-occurred at that site. In his report of
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFC1A1D5BE277F805" box="[984,1079,1914,1940]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||||
found at that swamp (
|
||||
<bibRefCitation id="EFCF32C3FFD4FFEBFAF81D5BE69FF829" author="Kosterin, O. E." pageId="5" pageNumber="456" pagination="185 - 224" refId="ref14616" refString="Kosterin, O. E. (2020 a) Miscellaneous faunal data on Odonata of Cambodia. International Dragonfly Fund Report, 154, 185-224." type="journal article" year="2020">Kosterin 2020a</bibRefCitation>
|
||||
), OEK listed
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFE431DBEE44CF829" authorityName="Kirby" authorityYear="1893" box="[385,524,1950,1976]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD4FFEBFE431DBEE44CF829" box="[385,524,1950,1976]" italics="true" pageId="5" pageNumber="456">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(37611012) with the abundance score of 4, which means “abundant (~20–100 [individuals])”. This report was correct, but
|
||||
<taxonomicName id="4C5E34B1FFD4FFEBFD461DE2E545F84D" authorityName="" (Orr" authorityYear="2005" box="[644,773,1986,2012]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD4FFEBFD461DE2E545F84D" box="[644,773,1986,2012]" italics="true" pageId="5" pageNumber="456">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
was missing. This location was also visited by
|
||||
<collectingCountry id="F3490FA2FFD4FFEBFAD41DE3E37AF84D" box="[1302,1338,1986,2012]" name="Philippines" pageId="5" pageNumber="456">PH</collectingCountry>
|
||||
on
|
||||
<date id="FFE069F2FFD4FFE8FAA61DE3E692FF20" lastPageId="6" lastPageNumber="457" pageId="5" pageNumber="456" value="2024-06-13">13 vi 2024</date>
|
||||
, who did not record
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FE711AB6E473FF20" authorityName="" (Orr" authorityYear="2005" box="[435,563,151,177]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD7FFE8FE711AB6E473FF20" box="[435,563,151,177]" italics="true" pageId="6" pageNumber="457">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
but did record
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FD1B1AB6E520FF20" authorityName="Kirby" authorityYear="1893" box="[729,864,151,177]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD7FFE8FD1B1AB6E520FF20" box="[729,864,151,177]" italics="true" pageId="6" pageNumber="457">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(241479436). Rather remarkably,
|
||||
<collectingCountry id="F3490FA2FFD7FFE8FB1C1AB6E342FF20" box="[1246,1282,151,177]" name="Philippines" pageId="6" pageNumber="457">PH</collectingCountry>
|
||||
also recorded
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FF551A9AE79BFF44" authority="Fraser, 1924" authorityName="Fraser" authorityYear="1924" box="[151,475,187,213]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="nigriceps">
|
||||
<emphasis id="B92A9320FFD7FFE8FF551A9AE70BFF45" box="[151,331,187,212]" italics="true" pageId="6" pageNumber="457">Lestes nigriceps</emphasis>
|
||||
<bibRefCitation id="EFCF32C3FFD7FFE8FE931A9AE79BFF44" author="Fraser, F. C." box="[337,475,187,213]" pageId="6" pageNumber="457" pagination="423 - 522" refId="ref13634" refString="Fraser, F. C. (1924) A survey of the odonate (dragonfly) fauna of western India with special remarks on the genera of Macromia and Idionyx and descriptions of thirty new species. Records of the Indian Museum, 26 (5), 423-522, pls. XXV - XXVII. https://doi.org/10.26515/rzsi/v26/i5/1924/162660" type="journal article" year="1924">Fraser, 1924</bibRefCitation>
|
||||
</taxonomicName>
|
||||
at this swamp, without any prior knowledge that OEK had previously visited the place and found this species at exactly the same site in
|
||||
<date id="FFE069F2FFD7FFE8FD7C1AFEE570FF68" box="[702,816,223,249]" pageId="6" pageNumber="457" value="2018-06">June 2018</date>
|
||||
(
|
||||
<bibRefCitation id="EFCF32C3FFD7FFE8FCFD1AFEE5A3FF68" author="Kosterin, O. E." box="[831,995,223,249]" pageId="6" pageNumber="457" pagination="561 - 575" refId="ref14481" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561-575. https://doi.org/10.11646/zootaxa.4526.4.8" type="journal article" year="2018">Kosterin 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD7FFE8FC2D1AFEE278FF68" author="Kosterin, O. E." box="[1007,1080,223,249]" pageId="6" pageNumber="457" pagination="185 - 224" refId="ref14616" refString="Kosterin, O. E. (2020 a) Miscellaneous faunal data on Odonata of Cambodia. International Dragonfly Fund Report, 154, 185-224." type="journal article" year="2020">2020a</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</subSection>
|
||||
<subSection id="E2D156D9FFD7FFE8FF551B6AE45AFB30" pageId="6" pageNumber="457" type="multiple">
|
||||
<subSubSection id="C3441CB9FFD7FFE8FF551B6AE45AFB30" pageId="6" pageNumber="457" type="discussion">
|
||||
<paragraph id="8BE14F32FFD7FFE8FF551B6AE5C8FEF4" blockId="6.[151,904,330,357]" box="[151,904,330,357]" pageId="6" pageNumber="457">
|
||||
<heading id="D0A9F85EFFD7FFE8FF551B6AE5C8FEF4" bold="true" box="[151,904,330,357]" fontSize="11" level="1" pageId="6" pageNumber="457" reason="1">
|
||||
<emphasis id="B92A9320FFD7FFE8FF551B6AE5C8FEF4" bold="true" box="[151,904,330,357]" pageId="6" pageNumber="457">
|
||||
5. Morphological comparison of
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FDEC1B6AE4FAFEF4" authorityName="" (Orr" authorityYear="2005" box="[558,698,330,357]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD7FFE8FDEC1B6AE4FAFEF4" bold="true" box="[558,698,330,357]" italics="true" pageId="6" pageNumber="457">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FD3A1B6AE5C8FEF4" authorityName="Kirby" authorityYear="1893" box="[760,904,330,357]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD7FFE8FD3A1B6AE5C8FEF4" bold="true" box="[760,904,330,357]" italics="true" pageId="6" pageNumber="457">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD7FFE8FF551BB2E447FD88" blockId="6.[151,1437,403,1185]" pageId="6" pageNumber="457">
|
||||
The photographs of morphological details of the
|
||||
<typeStatus id="54E5F190FFD7FFE8FD7B1BB2E55BFE3C" box="[697,795,403,429]" pageId="6" pageNumber="457" type="holotype">holotype</typeStatus>
|
||||
(
|
||||
<figureCitation id="136553B7FFD7FFE8FCEB1BB2E532FE3C" box="[809,882,403,429]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[295,1293,181,1908]" captionTargetId="figure-18@11.[295,1293,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. The holotype of Lestes dorothea, from S. India, Coorg, Mercara, 17 iv1923, F.C. Fraser, male, preserved in the BMNH collection: a–dorsal general view; b–labels, c-e–the end of abdomen in dorsal, ventral and lateral views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818163" httpUri="https://zenodo.org/record/15818163/files/figure.png" pageId="6" pageNumber="457">Figs 6</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFD7FFE8FCBF1BB2E5F5FE3C" box="[893,949,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1055,1079]" captionTargetBox="[151,1435,181,1031]" captionTargetId="figure-20@12.[151,1435,181,1031]" captionTargetPageId="12" captionText="FIGURE 7. Thorax of the holotype, male (a,b), and allotype, female (c,d) of Lestes dorothea, in lateral (a, c) and dorsal (b, d) views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818165" httpUri="https://zenodo.org/record/15818165/files/figure.png" pageId="6" pageNumber="457">7a–b</figureCitation>
|
||||
) and
|
||||
<typeStatus id="54E5F190FFD7FFE8FC311BB2E20DFE3C" box="[1011,1101,403,429]" pageId="6" pageNumber="457" type="allotype">allotype</typeStatus>
|
||||
(
|
||||
<figureCitation id="136553B7FFD7FFE8FB991BB2E28DFE3C" box="[1115,1229,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1055,1079]" captionTargetBox="[151,1435,181,1031]" captionTargetId="figure-20@12.[151,1435,181,1031]" captionTargetPageId="12" captionText="FIGURE 7. Thorax of the holotype, male (a,b), and allotype, female (c,d) of Lestes dorothea, in lateral (a, c) and dorsal (b, d) views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818165" httpUri="https://zenodo.org/record/15818165/files/figure.png" pageId="6" pageNumber="457">Figs 7c–d</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFD7FFE8FB1A1BB2E2A6FE3C" box="[1240,1254,403,429]" captionStart="FIGURE 8" captionStartId="13.[151,250,1302,1326]" captionTargetBox="[177,1411,181,1278]" captionTargetId="figure-18@13.[177,1411,181,1278]" captionTargetPageId="13" captionText="FIGURE 8. The allotype of Lestes dorothea, from S. India, Greenfields, Sidapur. 26 iv1923, F.C. Fraser, preserved in the BMNH collection: a–dorsal general view; b–labels, c,d–end of abdomen in lateral and dorsal views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818167" httpUri="https://zenodo.org/record/15818167/files/figure.png" pageId="6" pageNumber="457">8</figureCitation>
|
||||
) of
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FAD61BB2E3D5FE3C" authorityName="" (Orr" authorityYear="2005" box="[1300,1429,403,429]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD7FFE8FAD61BB2E3D5FE3C" box="[1300,1429,403,429]" italics="true" pageId="6" pageNumber="457">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
, and of the
|
||||
<typeStatus id="54E5F190FFD7FFE8FECE1B96E733FE40" box="[268,371,439,465]" pageId="6" pageNumber="457" type="lectotype">lectotype</typeStatus>
|
||||
(
|
||||
<figureCitation id="136553B7FFD7FFE8FE431B96E784FE40" box="[385,452,439,465]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[467,1121,181,1919]" captionTargetId="figure-20@14.[467,1121,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. The lectotype, male, of Lestes decipens, from Ceylon, Mahagany, 20 ix 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818171" httpUri="https://zenodo.org/record/15818171/files/figure.png" pageId="6" pageNumber="457">Fig. 9</figureCitation>
|
||||
) and female
|
||||
<typeStatus id="54E5F190FFD7FFE8FD961B96E4ABFE40" box="[596,747,439,465]" pageId="6" pageNumber="457" type="paralectotype">paralectotype</typeStatus>
|
||||
(incorrectly designated as “the
|
||||
<typeStatus id="54E5F190FFD7FFE8FB881B96E2E7FE40" box="[1098,1191,439,465]" pageId="6" pageNumber="457" type="allotype">allotype</typeStatus>
|
||||
” by
|
||||
<bibRefCitation id="EFCF32C3FFD7FFE8FB181B96E3DCFE40" author="Kimmins, D. E." box="[1242,1436,439,465]" pageId="6" pageNumber="457" pagination="171 - 205" refId="ref14051" refString="Kimmins, D. E. (1970) A list of the type-specimens of Odonata in the British Museum (Natural History). Part 3. Bulletin of the British Museum (Natural History), Entomology, 24 (6), 171-205. https://doi.org/10.5962/bhl.part.1521" type="journal article" year="1970">Kimmins (1970))</bibRefCitation>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FF761BFAE779FE64" authorityName="Kirby" authorityYear="1893" box="[180,313,475,501]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD7FFE8FF761BFAE779FE64" box="[180,313,475,501]" italics="true" pageId="6" pageNumber="457">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD7FFE8FE851BFAE7D7FE64" box="[327,407,475,501]" captionStart="FIGURE 10" captionStartId="15.[151,250,1944,1968]" captionTargetBox="[455,1133,181,1920]" captionTargetId="figure-18@15.[455,1133,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. A paralectotype, female (incorrectly designated as “the allotype” by Kimmins (1970)) of Lestes decipiens, from Ceylon, 6th Milestone, Nilavelli Road, 9 xii 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818173" httpUri="https://zenodo.org/record/15818173/files/figure.png" pageId="6" pageNumber="457">Fig. 10</figureCitation>
|
||||
), kept in the collection of Natural History Museum, London (BMNH) provided us a reference for the characters of this species.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD7FFE8FF051802E558FD60" blockId="6.[151,1437,403,1185]" pageId="6" pageNumber="457">
|
||||
The labels of the type specimens of
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FDA41802E4A9FDAC" authorityName="" (Orr" authorityYear="2005" box="[614,745,547,573]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD7FFE8FDA41802E4A9FDAC" box="[614,745,547,573]" italics="true" pageId="6" pageNumber="457">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD7FFE8FD381802E513FDAC" box="[762,851,547,573]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[295,1293,181,1908]" captionTargetId="figure-18@11.[295,1293,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. The holotype of Lestes dorothea, from S. India, Coorg, Mercara, 17 iv1923, F.C. Fraser, male, preserved in the BMNH collection: a–dorsal general view; b–labels, c-e–the end of abdomen in dorsal, ventral and lateral views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818163" httpUri="https://zenodo.org/record/15818163/files/figure.png" pageId="6" pageNumber="457">Figs 6b</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFD7FFE8FCA21802E53DFDAC" box="[864,893,547,573]" captionStart="FIGURE 8" captionStartId="13.[151,250,1302,1326]" captionTargetBox="[177,1411,181,1278]" captionTargetId="figure-18@13.[177,1411,181,1278]" captionTargetPageId="13" captionText="FIGURE 8. The allotype of Lestes dorothea, from S. India, Greenfields, Sidapur. 26 iv1923, F.C. Fraser, preserved in the BMNH collection: a–dorsal general view; b–labels, c,d–end of abdomen in lateral and dorsal views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818167" httpUri="https://zenodo.org/record/15818167/files/figure.png" pageId="6" pageNumber="457">8b</figureCitation>
|
||||
) provided more precise information of the type locality than that presented in the original description (
|
||||
<bibRefCitation id="EFCF32C3FFD7FFE8FD3E1866E5C7FDF0" author="Fraser, F. C." box="[764,903,583,609]" pageId="6" pageNumber="457" pagination="423 - 522" refId="ref13634" refString="Fraser, F. C. (1924) A survey of the odonate (dragonfly) fauna of western India with special remarks on the genera of Macromia and Idionyx and descriptions of thirty new species. Records of the Indian Museum, 26 (5), 423-522, pls. XXV - XXVII. https://doi.org/10.26515/rzsi/v26/i5/1924/162660" type="journal article" year="1924">Fraser 1924</bibRefCitation>
|
||||
), and also included the date. The
|
||||
<typeStatus id="54E5F190FFD7FFE8FB3F1866E31FFDF0" box="[1277,1375,583,609]" pageId="6" pageNumber="457" type="holotype">holotype</typeStatus>
|
||||
label says: “S.
|
||||
<collectingCountry id="F3490FA2FFD7FFE8FEC1184AE701FD14" box="[259,321,619,645]" name="India" pageId="6" pageNumber="457">India</collectingCountry>
|
||||
: / Coorg, / Mercara. /
|
||||
<date id="FFE069F2FFD7FFE8FDF5184AE4F6FD14" box="[567,694,619,645]" pageId="6" pageNumber="457" value="1923-04-17">17-iv-1923</date>
|
||||
. / Maj. F.C. Fraser.” (
|
||||
<figureCitation id="136553B7FFD7FFE8FC6F184AE241FD14" box="[941,1025,619,645]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[295,1293,181,1908]" captionTargetId="figure-18@11.[295,1293,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. The holotype of Lestes dorothea, from S. India, Coorg, Mercara, 17 iv1923, F.C. Fraser, male, preserved in the BMNH collection: a–dorsal general view; b–labels, c-e–the end of abdomen in dorsal, ventral and lateral views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818163" httpUri="https://zenodo.org/record/15818163/files/figure.png" pageId="6" pageNumber="457">Fig. 6b</figureCitation>
|
||||
), hence Mercara is the species’ type locality, and the
|
||||
<typeStatus id="54E5F190FFD7FFE8FE9A18AEE7F2FD38" box="[344,434,655,681]" pageId="6" pageNumber="457" type="allotype">allotype</typeStatus>
|
||||
label is as follows: “S.
|
||||
<collectingCountry id="F3490FA2FFD7FFE8FD0E18AEE54AFD38" box="[716,778,655,681]" name="India" pageId="6" pageNumber="457">India</collectingCountry>
|
||||
: / Coorg, / Greenfields, Sidapur. /
|
||||
<date id="FFE069F2FFD7FFE8FB5118AEE351FD38" box="[1171,1297,655,681]" pageId="6" pageNumber="457" value="1923-04-26">26-iv-1923</date>
|
||||
. / Maj. F.C. Fraser.” (
|
||||
<figureCitation id="136553B7FFD7FFE8FF3C1892E710FD5C" box="[254,336,691,717]" captionStart="FIGURE 8" captionStartId="13.[151,250,1302,1326]" captionTargetBox="[177,1411,181,1278]" captionTargetId="figure-18@13.[177,1411,181,1278]" captionTargetPageId="13" captionText="FIGURE 8. The allotype of Lestes dorothea, from S. India, Greenfields, Sidapur. 26 iv1923, F.C. Fraser, preserved in the BMNH collection: a–dorsal general view; b–labels, c,d–end of abdomen in lateral and dorsal views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818167" httpUri="https://zenodo.org/record/15818167/files/figure.png" pageId="6" pageNumber="457">Fig. 8b</figureCitation>
|
||||
). The original identification labels “
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FD201892E5A3FD5C" box="[738,995,691,717]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="subSpecies" species="praemorsa" subSpecies="dorothea">
|
||||
<emphasis id="B92A9320FFD7FFE8FD201892E5A3FD5C" box="[738,995,691,717]" italics="true" pageId="6" pageNumber="457">L. praemorsa dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
” treats this taxon in a subspecific level, which seems to be Fraser’s initial, not published opinion.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD7FFE8FF0518DAE2C6FC34" blockId="6.[151,1437,403,1185]" pageId="6" pageNumber="457">
|
||||
<bibRefCitation id="EFCF32C3FFD7FFE8FF0518DAE71DFC84" author="Kirby, J." box="[199,349,763,789]" pageId="6" pageNumber="457" pagination="545 - 566" refId="ref14162" refString="Kirby, J. (1893) Catalogue of the described Neuroptera Odonata (Dragonflies) of Ceylon, with descritions of new species. The Journal of the Linnaean Society of London, Zoology, 24, 545-566. https://doi.org/10.1111/j.1096-3642.1894.tb02495.x" type="journal article" year="1893">Kirby (1893)</bibRefCitation>
|
||||
did not indicate the
|
||||
<typeStatus id="54E5F190FFD7FFE8FD9618DAE4F6FC84" box="[596,694,763,789]" pageId="6" pageNumber="457" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FD2118DAE52EFC84" authorityName="Kirby" authorityYear="1893" box="[739,878,763,789]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD7FFE8FD2118DAE52EFC84" box="[739,878,763,789]" italics="true" pageId="6" pageNumber="457">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
, so
|
||||
<bibRefCitation id="EFCF32C3FFD7FFE8FC6018DAE21FFC84" author="Kimmins, D. E." box="[930,1119,763,789]" pageId="6" pageNumber="457" pagination="171 - 205" refId="ref14051" refString="Kimmins, D. E. (1970) A list of the type-specimens of Odonata in the British Museum (Natural History). Part 3. Bulletin of the British Museum (Natural History), Entomology, 24 (6), 171-205. https://doi.org/10.5962/bhl.part.1521" type="journal article" year="1970">Kimmins (1970)</bibRefCitation>
|
||||
designated one of its male
|
||||
<typeStatus id="54E5F190FFD7FFE8FF55193EE6BCFCA8" box="[151,252,799,825]" pageId="6" pageNumber="457" type="syntype">syntypes</typeStatus>
|
||||
, that from Mahagany,
|
||||
<date id="FFE069F2FFD7FFE8FE38193EE435FCA8" box="[506,629,799,825]" pageId="6" pageNumber="457" value="1891-09-20">20 ix 1891</date>
|
||||
, as its
|
||||
<typeStatus id="54E5F190FFD7FFE8FD01193EE56DFCA8" box="[707,813,799,825]" pageId="6" pageNumber="457" type="lectotype">lectotype</typeStatus>
|
||||
. At the same time he designated a female
|
||||
<typeStatus id="54E5F190FFD7FFE8FAC5193EE31EFCA8" box="[1287,1374,799,825]" pageId="6" pageNumber="457" type="syntype">syntype</typeStatus>
|
||||
from 6
|
||||
<superScript id="7C2BE27AFFD7FFE8FF671963E6F2FCC1" attach="left" box="[165,178,834,848]" fontSize="6" pageId="6" pageNumber="457">th</superScript>
|
||||
Milestone, Nilavelli Road,
|
||||
<date id="FFE069F2FFD7FFE8FE371962E42EFCCC" box="[501,622,835,861]" pageId="6" pageNumber="457" value="1891-12-09">9 xii 1891</date>
|
||||
, as “
|
||||
<typeStatus id="54E5F190FFD7FFE8FD6B1962E54BFCCC" box="[681,779,835,861]" pageId="6" pageNumber="457" type="allotype">allotype</typeStatus>
|
||||
”. However ICZN does not offer an option of subsequent designations of
|
||||
<typeStatus id="54E5F190FFD7FFE8FE811946E7E8FC10" box="[323,424,871,897]" pageId="6" pageNumber="457" type="allotype">allotypes</typeStatus>
|
||||
after original descriptions.Hence the latter female specimen is just one of the
|
||||
<typeStatus id="54E5F190FFD7FFE8FB361946E3D9FC10" box="[1268,1433,871,897]" pageId="6" pageNumber="457" type="paralectotype">paralectotypes</typeStatus>
|
||||
. (Also an
|
||||
<typeStatus id="54E5F190FFD7FFE8FF3C19AAE718FC34" box="[254,344,907,933]" pageId="6" pageNumber="457" type="allotype">allotype</typeStatus>
|
||||
does not have any taxonomical value above being one of the
|
||||
<typeStatus id="54E5F190FFD7FFE8FBC919AAE23CFC34" box="[1035,1148,907,933]" pageId="6" pageNumber="457" type="paratype">paratypes</typeStatus>
|
||||
.)
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD7FFE8FF05198EE707FC7C" blockId="6.[151,1437,403,1185]" pageId="6" pageNumber="457">
|
||||
We also analysed
|
||||
<specimenCount id="9D5884BBFFD7FFE8FE53198EE443FC58" box="[401,515,943,969]" count="2" pageId="6" pageNumber="457" type="male">two males</specimenCount>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FDEB198EE4EBFC58" authorityName="" (Orr" authorityYear="2005" box="[553,683,943,969]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD7FFE8FDEB198EE4EBFC58" box="[553,683,943,969]" italics="true" pageId="6" pageNumber="457">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<collectingRegion id="499A81D0FFD7FFE8FD32198EE57DFC58" box="[752,829,943,969]" country="India" name="Kerala" pageId="6" pageNumber="457">Kerala</collectingRegion>
|
||||
, which is another Indian state in the southern Western Ghats (
|
||||
<figureCitation id="136553B7FFD7FFE8FF2A19F2E779FC7C" box="[232,313,979,1005]" captionStart="FIGURE 11" captionStartId="16.[151,250,1399,1423]" captionTargetBox="[151,1436,181,1373]" captionTargetId="figure-20@16.[151,1436,181,1376]" captionTargetPageId="16" captionText="FIGURE 11. The anal appendages of two male specimens of Lestes dorothea from India, Western Ghats, Kerala, Iduikki District: a–c–a pond at 10.0951° N, 76.9319° E, 4 vi 2024; d–f–a pool under a rocky slope by Mankulam Power House, 10.0965° N, 76.9345° E, 19 vi 2024; a–d–dorsal view; b–e–ventral view; c–f–lateral view." figureDoi="http://doi.org/10.5281/zenodo.15818176" httpUri="https://zenodo.org/record/15818176/files/figure.png" pageId="6" pageNumber="457">Fig. 11</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD7FFE8FF0519D6E45AFB30" blockId="6.[151,1437,403,1185]" pageId="6" pageNumber="457">
|
||||
Since the Cambodian specimens available to us included only
|
||||
<specimenCount id="9D5884BBFFD7FFE8FCB819D6E5B5FB80" box="[890,1013,1015,1041]" count="1" pageId="6" pageNumber="457" type="female">one female</specimenCount>
|
||||
belonging to this group, preliminarily identified as
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FEE81E3AE7F3FBA4" authorityName="Kirby" authorityYear="1893" box="[298,435,1051,1077]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD7FFE8FEE81E3AE7F3FBA4" box="[298,435,1051,1077]" italics="true" pageId="6" pageNumber="457">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
, we also examined the females of
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FCFF1E3AE580FBA4" authorityName="" (Orr" authorityYear="2005" box="[829,960,1051,1077]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD7FFE8FCFF1E3AE580FBA4" box="[829,960,1051,1077]" italics="true" pageId="6" pageNumber="457">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD7FFE8FC121E3AE261FBA4" box="[976,1057,1051,1077]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="6" pageNumber="457">Fig. 12</figureCitation>
|
||||
a-e) and
|
||||
<taxonomicName id="4C5E34B1FFD7FFE8FB441E3AE34FFBA4" authorityName="Kirby" authorityYear="1893" box="[1158,1295,1051,1077]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD7FFE8FB441E3AE34FFBA4" box="[1158,1295,1051,1077]" italics="true" pageId="6" pageNumber="457">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD7FFE8FADD1E3AE330FBA4" box="[1311,1392,1051,1077]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="6" pageNumber="457">Fig. 12</figureCitation>
|
||||
f-j) from the neighbouring
|
||||
<collectingCountry id="F3490FA2FFD7FFE8FE551E1EE7BBFBC8" box="[407,507,1087,1113]" name="Thailand" pageId="6" pageNumber="457">Thailand</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F3490FA2FFD7FFE8FDEC1E1EE4D0FBC8" box="[558,656,1087,1113]" name="Vietnam" pageId="6" pageNumber="457">Vietnam</collectingCountry>
|
||||
, respectively. Their identification to species was proved by their being collected while in tandems with males. In sum, morphological details were examined in the following specimens and can be commented as follows:
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</subSection>
|
||||
</treatment>
|
||||
</document>
|
||||
464
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Normal file
464
data/03/F7/FE/03F7FE24FFD3FFEDFF5518F7E705FA05.xml
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|
|
@ -0,0 +1,464 @@
|
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<mods:title id="BB07C4C3E008933384B58C3C9F231C17">Taxonomic reconsideration of Lestes dorothea Fraser, 1924, L. decipiens Kirby, 1893, bona species, L. praecellens Lieftinck, 1937 and L. praemorsus Hagen in Selys, 1862 (Odonata, Lestidae)</mods:title>
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<mods:namePart id="54349917AA45F357B86C4FA80D37E344">Hopkins, Paul</mods:namePart>
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<mods:affiliation id="B074081246FCF3D075E2EF663AD72A1A">9 New street, Penryn, Cornwall, TR 10 8 EB, United Kingdom</mods:affiliation>
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<mods:namePart id="CDAA06C27B311ACFDCB7797ADFB13504">Kosterin, Oleg E.</mods:namePart>
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<mods:affiliation id="BC628330592D2444DC058D4104DE4357">Institute of Cytology and Genetics SB RAS, Academician Lavrentyev ave. 10, Novosibirsk, 630090, Russia</mods:affiliation>
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<mods:namePart id="E2915312B66FD7014D9925E1B295798A">Phan, Quoc Toan</mods:namePart>
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<mods:affiliation id="8818A63CD504A4D09594C6A3985AAAC5">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
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<subSubSection id="C3441CB9FFD3FFECFF5518F7E48CFD60" box="[151,716,726,753]" pageId="2" pageNumber="453" type="nomenclature">
|
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<paragraph id="8BE14F32FFD3FFECFF5518F7E48CFD60" blockId="2.[151,716,726,753]" box="[151,716,726,753]" pageId="2" pageNumber="453">
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<heading id="D0A9F85EFFD3FFECFF5518F7E48CFD60" bold="true" box="[151,716,726,753]" fontSize="11" level="1" pageId="2" pageNumber="453" reason="1">
|
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<emphasis id="B92A9320FFD3FFECFF5518F7E48CFD60" bold="true" box="[151,716,726,753]" pageId="2" pageNumber="453">
|
||||
3.
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFF7718F6E452FD61" authority="Kirby, 1893" authorityName="Kirby" authorityYear="1893" box="[181,530,726,753]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD3FFECFF7718F6E737FD60" bold="true" box="[181,375,726,753]" italics="true" pageId="2" pageNumber="453">Lestes decipiens</emphasis>
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFEBD18F7E452FD61" author="Kirby, J." box="[383,530,726,752]" pageId="2" pageNumber="453" pagination="545 - 566" refId="ref14162" refString="Kirby, J. (1893) Catalogue of the described Neuroptera Odonata (Dragonflies) of Ceylon, with descritions of new species. The Journal of the Linnaean Society of London, Zoology, 24, 545-566. https://doi.org/10.1111/j.1096-3642.1894.tb02495.x" type="journal article" year="1893">Kirby, 1893</bibRefCitation>
|
||||
</taxonomicName>
|
||||
is
|
||||
<emphasis id="B92A9320FFD3FFECFDF118F7E48CFD60" bold="true" box="[563,716,726,753]" italics="true" pageId="2" pageNumber="453">bona species</emphasis>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3441CB9FFD3FFECFF55193EE4B9F908" pageId="2" pageNumber="453" type="description">
|
||||
<paragraph id="8BE14F32FFD3FFECFF55193EE2C0FC34" blockId="2.[151,1437,799,2015]" pageId="2" pageNumber="453">
|
||||
In order to simplify further reading by making our taxonomic treatments uniform throughout further text, we have to start with a taxonomic revision based on literature only. Both
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFC9F1962E5BEFCCD" box="[861,1022,835,861]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFC9F1962E52DFCCD" box="[861,877,835,860]" italics="true" pageId="2" pageNumber="453">L</emphasis>
|
||||
.
|
||||
<emphasis id="B92A9320FFD3FFECFCB81962E5BEFCCD" box="[890,1022,835,860]" italics="true" pageId="2" pageNumber="453">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
(sub.
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFB801962E297FCCD" authorityName="Hagen" authorityYear="1862" box="[1090,1239,835,860]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsa">
|
||||
<emphasis id="B92A9320FFD3FFECFB801962E297FCCD" box="[1090,1239,835,860]" italics="true" pageId="2" pageNumber="453">L. praemorsa</emphasis>
|
||||
</taxonomicName>
|
||||
) and
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFAD41962E3DCFCCC" authorityName="Kirby" authorityYear="1893" box="[1302,1436,835,861]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD3FFECFAD41962E3DCFCCC" box="[1302,1436,835,861]" italics="true" pageId="2" pageNumber="453">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
were described at the species rank (
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFDE91946E4ECFC10" author="Selys Longchamps, M. E. de" box="[555,684,871,897]" pageId="2" pageNumber="453" pagination="288 - 338" refId="ref15384" refString="Selys Longchamps, M. E. de (1862) Synopsis des Agrionines. Seconde legion: Lestes. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 13, 288-338." type="journal article" year="1862">Selys 1862</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFD7B1946E57FFC10" author="Kirby, J." box="[697,831,871,897]" pageId="2" pageNumber="453" pagination="545 - 566" refId="ref14162" refString="Kirby, J. (1893) Catalogue of the described Neuroptera Odonata (Dragonflies) of Ceylon, with descritions of new species. The Journal of the Linnaean Society of London, Zoology, 24, 545-566. https://doi.org/10.1111/j.1096-3642.1894.tb02495.x" type="journal article" year="1893">Kirby 1893</bibRefCitation>
|
||||
). Their
|
||||
<typeStatus id="54E5F190FFD3FFECFC5A1946E588FC10" box="[920,968,871,897]" pageId="2" pageNumber="453">type</typeStatus>
|
||||
localities were fairly distant:
|
||||
<collectingRegion id="499A81D0FFD3FFECFADE1946E32EFC10" box="[1308,1390,871,897]" country="Philippines" name="Manila" pageId="2" pageNumber="453">Manila</collectingRegion>
|
||||
, the
|
||||
<collectingCountry id="F3490FA2FFD3FFECFF5519AAE755FC34" box="[151,277,907,933]" name="Philippines" pageId="2" pageNumber="453">Philippines</collectingCountry>
|
||||
for
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFE8619AAE7A5FC35" box="[324,485,907,932]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFE8619AAE7A5FC35" box="[324,485,907,932]" italics="true" pageId="2" pageNumber="453">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
and four localities on the island of
|
||||
<collectingCountry id="F3490FA2FFD3FFECFCB019AAE583FC34" box="[882,963,907,933]" name="Sri Lanka" pageId="2" pageNumber="453">Ceylon</collectingCountry>
|
||||
for
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFC3019AAE239FC34" authorityName="Kirby" authorityYear="1893" box="[1010,1145,907,933]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD3FFECFC3019AAE239FC34" box="[1010,1145,907,933]" italics="true" pageId="2" pageNumber="453">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD3FFECFF05198EE4B9F908" blockId="2.[151,1437,799,2015]" pageId="2" pageNumber="453">
|
||||
The description of
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFE55198EE478FC59" box="[407,568,943,968]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFE55198EE478FC59" box="[407,568,943,968]" italics="true" pageId="2" pageNumber="453">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
was based on a single female (
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFC48198EE249FC58" author="Selys Longchamps, M. E. de" box="[906,1033,943,969]" pageId="2" pageNumber="453" pagination="288 - 338" refId="ref15384" refString="Selys Longchamps, M. E. de (1862) Synopsis des Agrionines. Seconde legion: Lestes. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 13, 288-338." type="journal article" year="1862">Selys 1862</bibRefCitation>
|
||||
), which made its further comparison with other taxa difficult.
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFE6D19F2E432FC7C" author="Fraser, F. C." box="[431,626,979,1005]" pageId="2" pageNumber="453" refId="ref13746" refString="Fraser, F. C. (1933) The Fauna of British India including Ceylon and Burma: Odonata. Vol. 1. Taylor and Francis Ltd., London, 423 pp." type="book" year="1933">Fraser (1933: 33)</bibRefCitation>
|
||||
incorrectly claimed the existence of the male
|
||||
<typeStatus id="54E5F190FFD3FFECFBB819F2E294FC7C" box="[1146,1236,979,1005]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFB3819F2E3DCFC7D" box="[1274,1436,979,1004]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFB3819F2E3DCFC7D" box="[1274,1436,979,1004]" italics="true" pageId="2" pageNumber="453">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
from the current
|
||||
<collectingCountry id="F3490FA2FFD3FFECFEA019D6E791FB80" box="[354,465,1015,1041]" name="Myanmar" pageId="2" pageNumber="453">Myanmar</collectingCountry>
|
||||
, as follows: “The
|
||||
<emphasis id="B92A9320FFD3FFECFD6C19D6E49DFB81" box="[686,733,1015,1040]" italics="true" pageId="2" pageNumber="453">type</emphasis>
|
||||
, a female from
|
||||
<collectingRegion id="499A81D0FFD3FFECFC6319D6E5B3FB80" box="[929,1011,1015,1041]" country="Philippines" name="Manila" pageId="2" pageNumber="453">Manila</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F3490FA2FFD3FFECFBC119D6E2C4FB80" box="[1027,1156,1015,1041]" name="Philippines" pageId="2" pageNumber="453">Philippines</collectingCountry>
|
||||
, is now in the Selysian collection. The
|
||||
<typeStatus id="54E5F190FFD3FFECFE851E3AE7E4FBA4" box="[327,420,1051,1077]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
, a male, also in the same collection, was taken at Puepoli,
|
||||
<collectingCountry id="F3490FA2FFD3FFECFBFC1E3AE2CDFBA4" box="[1086,1165,1051,1077]" name="Myanmar" pageId="2" pageNumber="453">Burma</collectingCountry>
|
||||
, in June”. However, the original description of
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFE581E1EE494FBC8" authority="(Selys 1862)" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[410,724,1087,1113]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFE581E1EE47CFBC9" box="[410,572,1087,1112]" italics="true" pageId="2" pageNumber="453">L. praemorsus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFD8E1E1EE48BFBC8" author="Selys Longchamps, M. E. de" box="[588,715,1087,1113]" pageId="2" pageNumber="453" pagination="288 - 338" refId="ref15384" refString="Selys Longchamps, M. E. de (1862) Synopsis des Agrionines. Seconde legion: Lestes. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 13, 288-338." type="journal article" year="1862">Selys 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
claimed the male to be unknown. But the male specimen from Puepoli cannot be the
|
||||
<typeStatus id="54E5F190FFD3FFECFE561E42E7B1FBEC" box="[404,497,1123,1149]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
, since the
|
||||
<typeStatus id="54E5F190FFD3FFECFDA81E42E484FBEC" box="[618,708,1123,1149]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
is a member of the type series by definition, and the type series contains only specimens upon which the original description was based, regardless of further literature. Hence the additional male specimen in Selys’ collection has no taxonomic value. Fraser’s statement concerning the presumed
|
||||
<typeStatus id="54E5F190FFD3FFECFF551EEEE6B1FB78" box="[151,241,1231,1257]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
was also rejected by
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFE241EEEE4B8FB78" author="Lieftinck, M. A." box="[486,760,1231,1257]" pageId="2" pageNumber="453" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949: 33–34)</bibRefCitation>
|
||||
, as follows: “The first reference known to me in which a
|
||||
<specimenCount id="9D5884BBFFD3FFECFA451EEEE3DCFB78" box="[1415,1436,1231,1257]" count="1" pageId="2" pageNumber="453" type="male">♂</specimenCount>
|
||||
of this insect is mentioned, is a note of De Selys on a single specimen from Puepoli, in
|
||||
<collectingCountry id="F3490FA2FFD3FFECFBA01ED2E2F1FA9C" box="[1122,1201,1267,1293]" name="Myanmar" pageId="2" pageNumber="453">Burma</collectingCountry>
|
||||
, of which this author writes: “Le mâle des Monts Carin pris à Puepoli par M. Fea appartient au type de l’espèce par ses appendices anals supérieurs noirs au bout et aussi à l’extrême base” ” [The male from the Carin Mountains taken at Puepoli by Mr. Fea belongs to the type of the species because of its upper anal appendages black at the tip and also at the extreme base]. (Ann. Mus. civ. Genova (2) 10, 1891, p. 495–496). This statement does not necessarily imply that the two sexes, from such remote localities, are really con-subspecific, and since no description of this
|
||||
<specimenCount id="9D5884BBFFD3FFECFB021F86E295FA50" box="[1216,1237,1447,1473]" count="1" pageId="2" pageNumber="453" type="male">♂</specimenCount>
|
||||
was given I have not followed Fraser, who proclaimed that the Burmese
|
||||
<specimenCount id="9D5884BBFFD3FFECFCCC1FEAE563FA74" box="[782,803,1483,1509]" count="1" pageId="2" pageNumber="453" type="male">♂</specimenCount>
|
||||
is the
|
||||
<typeStatus id="54E5F190FFD3FFECFCB11FEAE58DFA74" box="[883,973,1483,1509]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFC341FEAE23AFA75" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1014,1146,1483,1508]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFC341FEAE23AFA75" box="[1014,1146,1483,1508]" italics="true" pageId="2" pageNumber="453">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
”. It should be added that Lieftinck’s reasons make no sense from the formal reasons: the Puepoli specimen would not be an
|
||||
<typeStatus id="54E5F190FFD3FFECFAC61FCEE31EF998" box="[1284,1374,1519,1545]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
even if it were conspecific with the
|
||||
<typeStatus id="54E5F190FFD3FFECFE301C32E414F9BC" box="[498,596,1555,1581]" pageId="2" pageNumber="453" type="holotype">holotype</typeStatus>
|
||||
and its description were published after the original description. Actually,
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFF551C16E710F9C0" author="Lieftinck, M. A." box="[151,336,1591,1617]" pageId="2" pageNumber="453" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
|
||||
just challenged the opinion by the species’ author, de Selys-Longchamps (based on the coloration of the appendages of different sexes!), that the additional
|
||||
<collectingCountry id="F3490FA2FFD3FFECFCE21C7AE5CDF9E4" box="[800,909,1627,1653]" name="Myanmar" pageId="2" pageNumber="453">Myanmar</collectingCountry>
|
||||
male was conspecific to the
|
||||
<typeStatus id="54E5F190FFD3FFECFB111C7AE375F9E4" box="[1235,1333,1627,1653]" pageId="2" pageNumber="453" type="holotype">holotype</typeStatus>
|
||||
from the
|
||||
<collectingCountry id="F3490FA2FFD3FFECFF551C5EE758F908" box="[151,280,1663,1689]" name="Philippines" pageId="2" pageNumber="453">Philippines</collectingCountry>
|
||||
, a doubt which we fully agree (see below).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3441CB9FFD3FFEDFF051C82E705FA05" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453" type="discussion">
|
||||
<paragraph id="8BE14F32FFD3FFECFF051C82E73BF894" blockId="2.[151,1437,799,2015]" pageId="2" pageNumber="453">
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFF051C82E7DAF92C" author="Kirby, J." box="[199,410,1698,1725]" pageId="2" pageNumber="453" pagination="545 - 566" refId="ref14162" refString="Kirby, J. (1893) Catalogue of the described Neuroptera Odonata (Dragonflies) of Ceylon, with descritions of new species. The Journal of the Linnaean Society of London, Zoology, 24, 545-566. https://doi.org/10.1111/j.1096-3642.1894.tb02495.x" type="journal article" year="1893">Kirby (1893: 566)</bibRefCitation>
|
||||
abstained from providing a factual differential diagnosis of
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFB9A1C82E2A2F92C" authorityName="Kirby" authorityYear="1893" box="[1112,1250,1699,1725]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD3FFECFB9A1C82E2A2F92C" box="[1112,1250,1699,1725]" italics="true" pageId="2" pageNumber="453">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
, providing only the following note: “Allied to
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFE3B1CE6E4D3F971" authorityName="Hagen" authorityYear="1862" box="[505,659,1735,1760]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsa">
|
||||
<emphasis id="B92A9320FFD3FFECFE3B1CE6E4D3F971" box="[505,659,1735,1760]" italics="true" pageId="2" pageNumber="453">L. praemorsa</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<collectingCountry id="F3490FA2FFD3FFECFD671CE6E567F970" box="[677,807,1735,1761]" name="Philippines" pageId="2" pageNumber="453">Philippines</collectingCountry>
|
||||
) and
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFCA91CE6E23FF970" authority=", De Selys" authorityName="De Selys" box="[875,1151,1735,1761]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="quercifolia">
|
||||
<emphasis id="B92A9320FFD3FFECFCA91CE6E246F970" box="[875,1030,1735,1761]" italics="true" pageId="2" pageNumber="453">L. quercifolia</emphasis>
|
||||
, De Selys
|
||||
</taxonomicName>
|
||||
(Menado and
|
||||
<collectingRegion id="499A81D0FFD3FFECFAE91CE6E322F970" box="[1323,1378,1735,1761]" country="Philippines" name="Sulu" pageId="2" pageNumber="453">Sulu</collectingRegion>
|
||||
), but apparently distinct”.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD3FFECFF051D2EE28AF84E" blockId="2.[151,1437,799,2015]" pageId="2" pageNumber="453">
|
||||
<bibRefCitation id="EFCF32C3FFD3FFECFF051D2EE7C5F8B8" author="Fraser, F. C." box="[199,389,1807,1833]" pageId="2" pageNumber="453" refId="ref13746" refString="Fraser, F. C. (1933) The Fauna of British India including Ceylon and Burma: Odonata. Vol. 1. Taylor and Francis Ltd., London, 423 pp." type="book" year="1933">Fraser (1933: 33)</bibRefCitation>
|
||||
assumed
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFE2D1D2EE435F8B8" authorityName="Kirby" authorityYear="1893" box="[495,629,1807,1833]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD3FFECFE2D1D2EE435F8B8" box="[495,629,1807,1833]" italics="true" pageId="2" pageNumber="453">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
as the junior synonym of
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFC4E1D2EE26BF8B9" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[908,1067,1807,1832]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFC4E1D2EE26BF8B9" box="[908,1067,1807,1832]" italics="true" pageId="2" pageNumber="453">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
, with the following note: “Owing to the gradual development of melanosis, a large number of varying forms are found ranging from pale brown with distinct markings in the teneral condition to melanotic forms which are quite black and exhibit no markings. Kirby’s form, known as
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFE8C1D5AE795F804" authorityName="Kirby" authorityYear="1893" box="[334,469,1915,1941]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD3FFECFE8C1D5AE795F804" box="[334,469,1915,1941]" italics="true" pageId="2" pageNumber="453">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
, is merely one such as the last, very heavily pruinosed.” This point of view was based on his above discussion and incorrect assumption of the existence of the male
|
||||
<typeStatus id="54E5F190FFD3FFECFBD21DBEE22AF828" box="[1040,1130,1951,1977]" pageId="2" pageNumber="453" type="allotype">allotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD3FFECFB531D81E373F828" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1169,1331,1952,1977]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD3FFECFB531D81E373F828" box="[1169,1331,1952,1977]" italics="true" pageId="2" pageNumber="453">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
from the current
|
||||
<collectingCountry id="F3490FA2FFD3FFECFF2F1DE4E71CF84E" box="[237,348,1989,2015]" name="Myanmar" pageId="2" pageNumber="453">Myanmar</collectingCountry>
|
||||
. Note that Fraser did not consider structural characters in the above statement.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD2FFEDFF051AB6E727FDD7" blockId="3.[151,1437,151,1465]" pageId="3" pageNumber="454">
|
||||
It was
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFED31AB6E788FF20" author="Lieftinck, M. A." box="[273,456,151,177]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
|
||||
who considered and illustrated both taxa in detail, based on specimens of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFAC21AB6E3C4FF21" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1280,1412,151,176]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" sensu="stricto" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFAC21AB6E3C4FF21" box="[1280,1412,151,176]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2192E5BFFD2FFEDFA481AB6E6F9FF47" pageId="3" pageNumber="454" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
from the
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFEEA1A9AE7E6FF44" box="[296,422,187,213]" name="Philippines" pageId="3" pageNumber="454">Philippines</collectingCountry>
|
||||
and New
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFDD81A9AE42BFF44" box="[538,619,187,213]" name="Guinea-Bissau" pageId="3" pageNumber="454">Guinea</collectingCountry>
|
||||
and specimens of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFCFC1A9AE5E8FF44" authorityName="Kirby" authorityYear="1893" box="[830,936,187,213]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFCFC1A9AE5E8FF44" box="[830,936,187,213]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
from the Kodagu District of
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFB301A9AE36CFF44" box="[1266,1324,187,213]" name="India" pageId="3" pageNumber="454">India</collectingCountry>
|
||||
and Java. His detailed key and fine drawings of the appendages (
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFCC71AC1E5F1FF6B" author="Lieftinck, M. A." box="[773,945,224,250]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949</bibRefCitation>
|
||||
: figs 19–20), reproduced here as
|
||||
<figureCitation id="136553B7FFD2FFEDFAE91AC1E3D8FF6B" box="[1323,1432,224,250]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="3" pageNumber="454">Fig. 1a–d</figureCitation>
|
||||
, show a drastic difference in the paraprocts, which are diverging and concealed under the cerci in dorsal view in
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFF551B0AE75BFED5" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[151,283,299,324]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" sensu="stricto" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFF551B0AE75BFED5" box="[151,283,299,324]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2192E5BFFD2FFEDFEE01B0BE71CFED5" box="[290,348,298,324]" pageId="3" pageNumber="454" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="136553B7FFD2FFEDFEA91B0BE796FED5" box="[363,470,298,324]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="3" pageNumber="454">Fig. 1a–b</figureCitation>
|
||||
) while directed caudad and well seen in dorsal view in
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFBFD1B0BE2E9FED5" authorityName="Kirby" authorityYear="1893" box="[1087,1193,298,324]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFBFD1B0BE2E9FED5" box="[1087,1193,298,324]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD2FFEDFB751B0BE365FED5" box="[1207,1317,298,324]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="3" pageNumber="454">Fig. 1c–d</figureCitation>
|
||||
). In lateral view, the paraproct of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFE5B1B6EE45DFEF9" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[409,541,335,360]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" sensu="stricto" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFE5B1B6EE45DFEF9" box="[409,541,335,360]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2192E5BFFD2FFEDFDE41B6EE41DFEF8" box="[550,605,335,361]" pageId="3" pageNumber="454" sensu="stricto">s. str</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="136553B7FFD2FFEDFDAC1B6EE49CFEF8" box="[622,732,335,361]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="3" pageNumber="454">Fig. 1a–b</figureCitation>
|
||||
) has the distal part narrower than in
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFB461B6EE2AEFEF8" authorityName="Kirby" authorityYear="1893" box="[1156,1262,335,361]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFB461B6EE2AEFEF8" box="[1156,1262,335,361]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136553B7FFD2FFEDFB3D1B6EE32FFEF8" box="[1279,1391,335,361]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="3" pageNumber="454">Fig. 1c–d</figureCitation>
|
||||
). In spite of these drastic differences,
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFDD51B52E492FE1F" author="Lieftinck, M. A." box="[535,722,371,398]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
|
||||
considered the taxon
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFC0D1B52E279FE1C" authorityName="Kirby" authorityYear="1893" box="[975,1081,371,397]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFC0D1B52E279FE1C" box="[975,1081,371,397]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
as a subspecies
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFB3A1B55E741FE23" authorityName="Kirby" authorityYear="1894" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="subSpecies" species="praemorsus" subSpecies="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFB3A1B55E741FE23" italics="true" pageId="3" pageNumber="454">L. praemorsus decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
rather than a full species
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFDE31BB8E4E9FE23" authorityName="Kirby" authorityYear="1893" box="[545,681,408,434]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFDE31BB8E4E9FE23" box="[545,681,408,434]" italics="true" pageId="3" pageNumber="454">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
. This point of view was assumed by later authors (e.g.
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFAD71BB9E71FFE46" author="Hamalainen, M. & Pinratana, A." pageId="3" pageNumber="454" refId="ref13782" refString="Hamalainen, M. & Pinratana, A. (1999) Atlas of the Dragonflies of Thailand. Distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp." type="book" year="1999">Hämäläinen & Pinratana 1999</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFEA81B9CE792FE49" author="Orr, A. G." box="[362,466,445,472]" pageId="3" pageNumber="454" refId="ref15216" refString="Orr, A. G. (2005) Dragonflies of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 127 pp." type="book" year="2005">Orr 2005</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFE1F1B9FE4E0FE46" author="Do, M. C. & Dang, T. T. H." box="[477,672,445,472]" pageId="3" pageNumber="454" refId="ref13511" refString="Do, M. C. & Dang, T. T. H. (2007) Checklist of dragonfly from Vietnam. Vietnam National University Publisher, Hanoi, 182 pp." type="book" year="2007">Do & Dang 2007</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFD6E1B9CE53DFE46" author="Roland, H. - H. & Sacher, T. & Roland, N." box="[684,893,445,472]" pageId="3" pageNumber="454" pagination="1 - 22" refId="ref15327" refString="Roland, H. - H., Sacher, T. & Roland, N. (2010) New records of Odonata for Cambodia - results from a trip through various places of the country November 14 th - December 1 st 2010. International Dragonfly Fund Report, 35, 1-22." type="journal article" year="2010">
|
||||
Roland
|
||||
<emphasis id="B92A9320FFD2FFEDFCC11B9FE574FE46" box="[771,820,445,471]" italics="true" pageId="3" pageNumber="454">et al</emphasis>
|
||||
. 2010
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFC451B9FE27EFE46" author="Tang, H. B. & Wang, L. K. & Hamalainen, M." box="[903,1086,445,472]" pageId="3" pageNumber="454" refId="ref15456" refString="Tang, H. B., Wang, L. K. & Hamalainen, M. (2010) A photographic guide to the dragonflies of Singapore. Raffles Museum of Biodiversity Research, Singapore, 222 pp." type="book" year="2010">
|
||||
Tang
|
||||
<emphasis id="B92A9320FFD2FFEDFC071B9FE5B5FE46" box="[965,1013,445,471]" italics="true" pageId="3" pageNumber="454">et al</emphasis>
|
||||
. 2010
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFB8B1B9CE2AAFE46" author="Kosterin, O. E." box="[1097,1258,445,471]" pageId="3" pageNumber="454" pagination="1 - 108" refId="ref14261" refString="Kosterin, O. E. (2011) Odonata of the Cambodian coastal regions revisited: beginning of dry season 2010. International Dragonfly Fund Report, 40, 1-108." type="journal article" year="2011">Kosterin 2011</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFB371B9CE37FFE46" author="Kosterin, O. E." box="[1269,1343,445,471]" pageId="3" pageNumber="454" pagination="1 - 102" refId="ref14295" refString="Kosterin, O. E. (2012 a) Odonata of the Cambodian coastal regions in late rainy season of 2011. International Dragonfly Fund Report, 45, 1-102." type="journal article" year="2012">2012a</bibRefCitation>
|
||||
,b; 2014; 2016; 2017; 2018; 2020a;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFE0A1BC3E4F2FE6D" author="Kosterin, O. E. & Chartier, G. & Holden, J. & Mey, F. S." box="[456,690,482,508]" pageId="3" pageNumber="454" pagination="150 - 163" refId="ref14824" refString="Kosterin, O. E., Chartier, G., Holden, J. & Mey, F. S. (2012) New records of Odonata from Cambodia, based mostly on photographs. Cambodian Journal of Natural History, 2012, 150-163." type="journal article" year="2012">
|
||||
Kosterin
|
||||
<emphasis id="B92A9320FFD2FFEDFDF01BC2E42DFE6D" box="[562,621,482,508]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
|
||||
2012
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFD7C1BC3E250FE6D" author="Yokoi, N. & Souphanthong, V." box="[702,1040,482,508]" pageId="3" pageNumber="454" refId="ref15573" refString="Yokoi, N. & Souphanthong, V. (2014) A List of Lao Dragonflies. Kyoei Printing Co Ltd., Koriyama, 110 pp." type="book" year="2014">Yokoi & Souphanthong 2014</bibRefCitation>
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||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFBDC1BC3E307FE6D" author="Kosterin, O. E. & Chartier, G." box="[1054,1351,482,508]" pageId="3" pageNumber="454" pagination="1 - 26" refId="ref14776" refString="Kosterin, O. E. & Chartier, G. (2017) Update of 2014 and 2016 to Odonata found at the marshy coast of SW Cambodia including three species added for the country. International Dragonfly Fund Report, 101, 1-26." type="journal article" year="2017">Kosterin & Chartier 2017</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFA971BC3E692FDB0" author="Zhang, H. - M." pageId="3" pageNumber="454" refId="ref15602" refString="Zhang, H. - M. (2019) Dragonflies and damselflies of China. Chongqing University Press, Chongquing, 1450 pp. [in Chinese and English]" type="book" year="2019">Zhang 2019</bibRefCitation>
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||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFF221826E78FFDB0" author="Kalkman, V. J. & Babu, R. & Bedjanic, M. & Conniff, K. & Gyeltshen, M. K. & Khan, M. K. & Subramanian, K. A. & Zia, A. & Orr, A. G." box="[224,463,519,545]" pageId="3" pageNumber="454" pagination="1 - 84" refId="ref13886" refString="Kalkman, V. J., Babu, R., Bedjanic, M., Conniff, K., Gyeltshen, M. K., Khan, M. K., Subramanian, K. A., Zia, A. & Orr, A. G. (2020) Checklist of the dragonflies and damselflies (Insecta: Odonata) of Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka. Zootaxa, 4849 (1), 1-84. https://doi.org/10.11646/zootaxa.4849.1.1" type="journal article" year="2020">
|
||||
Kalkman
|
||||
<emphasis id="B92A9320FFD2FFEDFE921826E7C3FDB0" box="[336,387,519,545]" italics="true" pageId="3" pageNumber="454">et al</emphasis>
|
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. 2020
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFE1E1826E4ACFDB0" author="Kosterin, O. E. & Smith, E." box="[476,748,519,545]" pageId="3" pageNumber="454" pagination="99 - 184" refId="ref14874" refString="Kosterin, O. E. & Smith, E. (2020) Odonata of Phnom Kulen Mts, Cambodia: a preliminary checklist. International Dragonfly Fund Report, 154, 99-184." type="journal article" year="2020">Kosterin & Smith 2020</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFD3B1826E212FDB0" author="Keetapithchayakul, T. S. & Makbun, N. & Rattanachan, K. & Tungpairojwong, N." box="[761,1106,519,545]" pageId="3" pageNumber="454" pagination="1 - 46" refId="ref13991" refString="Keetapithchayakul, T. S., Makbun, N., Rattanachan, K. & Tungpairojwong, N. (2023) Contribution to the knowledge of the Odonata fauna of Khao Yai National Park and the adjacent regions in Thailand. Faunistic Studies in South-east Asian and Pacific Island Odonata, 42, 1-46." type="journal article" year="2023">
|
||||
Keetapithchayakul
|
||||
<emphasis id="B92A9320FFD2FFEDFC161826E247FDB0" box="[980,1031,519,545]" italics="true" pageId="3" pageNumber="454">et al</emphasis>
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. 2023
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFBA21826E35CFDB0" author="Dow, R. A. & Choong, C. Y. & Grinang, J. & Lupiyaningdyah, P. & Ngiam, R. W. J. & Kalkman, V. J." box="[1120,1308,519,545]" pageId="3" pageNumber="454" pagination="1 - 122" refId="ref13545" refString="Dow, R. A., Choong, C. Y., Grinang, J., Lupiyaningdyah, P., Ngiam, R. W. J. & Kalkman, V. J. (2024) Checklist of the Odonata (Insecta) of Sundaland and Wallacea (Malaysia, Singapore, Brunei, Indonesia and Timor Leste). Zootaxa, 5460 (1), 1-122. https://doi.org/10.11646/zootaxa.5460.1.1" type="journal article" year="2024">
|
||||
Dow
|
||||
<emphasis id="B92A9320FFD2FFEDFB5D1826E293FDB0" box="[1183,1235,519,545]" italics="true" pageId="3" pageNumber="454">et al</emphasis>
|
||||
. 2024
|
||||
</bibRefCitation>
|
||||
) and is the current consensus.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD2FFEDFF051870E775FC00" blockId="3.[151,1437,151,1465]" pageId="3" pageNumber="454">
|
||||
However, Lieftinck was far from being certain in this solution, which was just preliminary, as follows from the following citations: “This statement [by Selys, that the Puepoli male was
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFC101857E216FD1E" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[978,1110,630,655]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFC101857E216FD1E" box="[978,1110,630,655]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
, see the citation above] does not necessarily imply that the two sexes, from such remote localities [the
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFC3218BBE22EFD25" box="[1008,1134,666,692]" name="Philippines" pageId="3" pageNumber="454">Philippines</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFB6818BBE35DFD25" box="[1194,1309,666,692]" name="Myanmar" pageId="3" pageNumber="454">Myanmar</collectingCountry>
|
||||
], are really con-subspecific” (
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFEA0189EE425FD48" author="Lieftinck, M. A." box="[354,613,703,729]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949: 33–34</bibRefCitation>
|
||||
). “I prefer for the present to take a very broad view and to recognize two races only; and although the present situation is not wholly satisfactory, any attempt to a further splitting up of the western subspecies
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFE631929E44BFCB3" authorityName="Kirby" authorityYear="1893" box="[417,523,776,802]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFE631929E44BFCB3" box="[417,523,776,802]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
has proved to be futile. On the other hand, a small number of undescribed races seem to exist in the eastern part of the Archipelago, but there is no doubt much intergradation; and a long series of
|
||||
<typeStatus id="54E5F190FFD2FFEDFF551973E744FCFD" box="[151,260,850,876]" pageId="3" pageNumber="454" type="topotype">topotypes</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFEEB1972E7EDFCFD" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[297,429,851,876]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFEEB1972E7EDFCFD" box="[297,429,851,876]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
from the
|
||||
<collectingCountry id="F3490FA2FFD2FFEDFDD91973E4D9FCFD" box="[539,665,850,876]" name="Philippines" pageId="3" pageNumber="454">Philippines</collectingCountry>
|
||||
is necessary to justify the separation of these subspecies.” (
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFAF61973E769FC00" author="Lieftinck, M. A." pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949: 35–36</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD2FFEDFF0519BDE230FBB5" blockId="3.[151,1437,151,1465]" pageId="3" pageNumber="454">
|
||||
On the other hand, he wrote: “The two subspecies are easily separated by differences which may be seen from the following comparison and from the drawings” (
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFD2119E1E5A8FC4A" author="Lieftinck, M. A." box="[739,1000,960,987]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck 1949: 35–36</bibRefCitation>
|
||||
). Curiously, some species assumed by him as distinct in fact exhibited much less difference. For instance, the species described by him 12 years before,
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFF551E2AE725FBB5" authorityName="Lieftinck" authorityYear="1937" box="[151,357,1034,1060]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="praecellens">
|
||||
<emphasis id="B92A9320FFD2FFEDFF551E2AE725FBB5" box="[151,357,1034,1060]" italics="true" pageId="3" pageNumber="454">Lestes praecellens</emphasis>
|
||||
</taxonomicName>
|
||||
, is very similar to
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFDF51E2AE4FBFBB5" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[567,699,1035,1060]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" sensu="stricto" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFDF51E2AE4FBFBB5" box="[567,699,1035,1060]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2192E5BFFD2FFEDFD001E2BE4BDFBB5" box="[706,765,1034,1060]" pageId="3" pageNumber="454" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
but not to
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFCB41E2BE5A0FBB5" authorityName="Kirby" authorityYear="1893" box="[886,992,1034,1060]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFCB41E2BE5A0FBB5" box="[886,992,1034,1060]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
(see below).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD2FFEDFF051E0EE5D3FA90" blockId="3.[151,1437,151,1465]" pageId="3" pageNumber="454">
|
||||
Thus, the preliminary solution of
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFD981E0EE558FBD8" author="Lieftinck, M. A." box="[602,792,1071,1097]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
|
||||
to consider the taxon
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFBE91E0EE2D5FBD8" authorityName="Kirby" authorityYear="1893" box="[1067,1173,1071,1097]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFBE91E0EE2D5FBD8" box="[1067,1173,1071,1097]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
as a subspecies of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFA441E0EE75BFBFC" baseAuthorityName="Selys" baseAuthorityYear="1862" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFA441E0EE75BFBFC" italics="true" pageId="3" pageNumber="454">L. praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
looks strange even for his time and in view of his own earlier works. We find it not justified, while the original species status of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFE6F1E58E474FB03" authorityName="Kirby" authorityYear="1893" box="[429,564,1144,1170]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFE6F1E58E474FB03" box="[429,564,1144,1170]" italics="true" pageId="3" pageNumber="454">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
fits much better the evidence presented by
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFBD31E59E288FB03" author="Lieftinck, M. A." box="[1041,1224,1144,1170]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
|
||||
himself. Further in this paper one will see that the anal appendages of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFD041EBFE506FB26" authorityName="Fraser" authorityYear="1924" box="[710,838,1181,1207]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD2FFEDFD041EBFE506FB26" box="[710,838,1181,1207]" italics="true" pageId="3" pageNumber="454">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
are again nearly identical to
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFB471EBFE349FB26" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1157,1289,1182,1207]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" sensu="stricto" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFB471EBFE349FB26" box="[1157,1289,1182,1207]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2192E5BFFD2FFEDFACD1EBFE309FB29" box="[1295,1353,1182,1208]" pageId="3" pageNumber="454" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
but not to
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFF761EE3E75EFB4D" authorityName="Kirby" authorityYear="1893" box="[180,286,1218,1244]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFF761EE3E75EFB4D" box="[180,286,1218,1244]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
. In fact, that Lieftinck’s solution to consider
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFCDA1EE3E5C2FB4D" authorityName="Kirby" authorityYear="1893" box="[792,898,1218,1244]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFCDA1EE3E5C2FB4D" box="[792,898,1218,1244]" italics="true" pageId="3" pageNumber="454">decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
as a subspecies of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFB9A1EE2E29CFB4D" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1112,1244,1219,1244]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFB9A1EE2E29CFB4D" box="[1112,1244,1219,1244]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
obscured further species comparisons and resulted to a nomenclatural mess in
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFC841EC6E5CCFA91" box="[838,908,1255,1280]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B92A9320FFD2FFEDFC841EC6E5CCFA91" box="[838,908,1255,1280]" italics="true" pageId="3" pageNumber="454">Lestes</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD2FFEDFF051F2AE705FA05" blockId="3.[151,1437,151,1465]" pageId="3" pageNumber="454">
|
||||
Unfortunately, we have no topotypic specimens of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFCD11F2DE5D7FAB4" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[787,919,1292,1317]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" sensu="stricto" species="praemorsus">
|
||||
<emphasis id="B92A9320FFD2FFEDFCD11F2DE5D7FAB4" box="[787,919,1292,1317]" italics="true" pageId="3" pageNumber="454">praemorsus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2192E5BFFD2FFEDFC631F2DE59EFAB7" box="[929,990,1292,1318]" pageId="3" pageNumber="454" sensu="stricto">s. str.</taxonomicNameLabel>
|
||||
from Luzon, an utmost need of which was stated by
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFEFB1F11E7B2FADB" author="Lieftinck, M. A." box="[313,498,1328,1354]" pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949)</bibRefCitation>
|
||||
still 76 years ago. However, based on a detailed key and illustrations by
|
||||
<bibRefCitation id="EFCF32C3FFD2FFEDFAF71F11E69DFAFE" author="Lieftinck, M. A." pageId="3" pageNumber="454" pagination="1 - 271" refId="ref15109" refString="Lieftinck, M. A. (1949) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VII. Results of the Third Archbold expedition 1938 - 1939 and of the Le Roux expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1-271." type="journal article" year="1949">Lieftinck (1949</bibRefCitation>
|
||||
: figs 19–20; our
|
||||
<figureCitation id="136553B7FFD2FFEDFE5C1F74E449FAFE" box="[414,521,1365,1391]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="3" pageNumber="454">Fig. 1a–d</figureCitation>
|
||||
) showing the profound differences in the paraprocts, we restore the species rank of
|
||||
<taxonomicName id="4C5E34B1FFD2FFEDFF741F5AE77DFA05" authorityName="Kirby" authorityYear="1893" box="[182,317,1402,1428]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD2FFEDFF741F5AE77DFA05" box="[182,317,1402,1428]" italics="true" pageId="3" pageNumber="454">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,563 +0,0 @@
|
|||
<document id="F2E2CC8BCDFBA12FFFF5C536FF416846" ID-DOI="10.11646/zootaxa.5642.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="15818151" ID-ZooBank="6CAAAAD8-0347-43C8-B3E9-D7011B436C19" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1751790876585" checkinUser="plazi" docAuthor="Hopkins, Paul, Kosterin, Oleg E., Phan, Quoc Toan & Keetapithchayakul, Tosaphol Saetung" docDate="2025" docId="03F7FE24FFD9FFE7FF551CB5E31AFA9C" docLanguage="en" docName="zootaxa.5642.5.3.pdf" docOrigin="Zootaxa 5642 (5)" docSource="https://doi.org/10.11646/zootaxa.5642.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lestes decipiens Kirby 1893" docType="treatment" docVersion="2" lastPageNumber="460" masterDocId="FFCE865CFFD1FFEEFFC21A21E640FF91" masterDocTitle="Taxonomic reconsideration of Lestes dorothea Fraser, 1924, L. decipiens Kirby, 1893, bona species, L. praecellens Lieftinck, 1937 and L. praemorsus Hagen in Selys, 1862 (Odonata, Lestidae)" masterLastPageNumber="475" masterPageNumber="451" pageNumber="459" updateTime="1751791374561" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||||
<mods:mods id="04499F55AB0414F2E07FB9653A4A9137" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="E7763EA175EC373BF383A40EE5478DD1">
|
||||
<mods:title id="9176D510570B6AAC7B3464A88DAD2E0A">Taxonomic reconsideration of Lestes dorothea Fraser, 1924, L. decipiens Kirby, 1893, bona species, L. praecellens Lieftinck, 1937 and L. praemorsus Hagen in Selys, 1862 (Odonata, Lestidae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="150B584A433134470A98C1A6AD3D1284" type="personal">
|
||||
<mods:role id="7F3A891ABD2AE6A14FD92ABE30130401">
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<mods:roleTerm id="6DCF44DB8C1E97172E0F15B348AE3FB0">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="A104DFBF67B3234865B6602E90387C75">Hopkins, Paul</mods:namePart>
|
||||
<mods:affiliation id="3A1471C0BCA6ECEB124993930E4618A4">9 New street, Penryn, Cornwall, TR 10 8 EB, United Kingdom</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="07748A79BCC9E7981890571C844D6558" type="personal">
|
||||
<mods:role id="9C543CF406A04DD19B3FE338B1561C16">
|
||||
<mods:roleTerm id="B39792A84A56C62DE88EEE8BF3AF0F19">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="F7D60DDC765D8995D072A88AE60189EB">Kosterin, Oleg E.</mods:namePart>
|
||||
<mods:affiliation id="3820D52C2E518EB9E54B1AF311903EF4">Institute of Cytology and Genetics SB RAS, Academician Lavrentyev ave. 10, Novosibirsk, 630090, Russia</mods:affiliation>
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||||
</mods:name>
|
||||
<mods:name id="8787FAED4EEB5A2BC339788D9385AE26" type="personal">
|
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<mods:role id="C3AF149C0A16D5C91C7074F89F83A7FF">
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<mods:roleTerm id="84EFFFACB6797D57F32ABE801DF1EF70">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="88B8424BC60A41871C2D182D40A10473">Phan, Quoc Toan</mods:namePart>
|
||||
<mods:affiliation id="34145B28BE7AC06C4A9BFEED061B441C">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="E5E125D58F76B087A247EA08984BF840" type="personal">
|
||||
<mods:role id="6334756F32CB0EE012FC484C9E648506">
|
||||
<mods:roleTerm id="403C262384E32C2FC2FD5D4FE4239901">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="4AF3DC290FE130D729E41515CFD2CB5A">Keetapithchayakul, Tosaphol Saetung</mods:namePart>
|
||||
<mods:nameIdentifier id="5FDBBA2EBC64BE288C80D72FF443C644" type="ORCID">0000-0001-7565-4701</mods:nameIdentifier>
|
||||
<mods:affiliation id="0013558D6ED1C45E6943E8E427CBAD72">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, Vietnam</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="21CA3A304715A3AC9C6FB3E621249391">text</mods:typeOfResource>
|
||||
<mods:relatedItem id="1259F12560D7521C0555BC4A076E691E" type="host">
|
||||
<mods:titleInfo id="27760651E27C0B188053EDB816B60E0D">
|
||||
<mods:title id="65A16AEF2342A2C185C2C915A62F66EB">Zootaxa</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="A1BAEFA408665091DBCE7990678DF1D6">
|
||||
<mods:date id="987A6D3E03B4BA341DE81A56F3E3E39C">2025</mods:date>
|
||||
<mods:detail id="327E917CD273C37BCF0634005046E423" type="pubDate">
|
||||
<mods:number id="2DD83D394F027AEE7575B20C0A2268A5">2025-06-03</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="5072768F1D6697B3F8ABDB0C491BD580" type="volume">
|
||||
<mods:number id="769D49EEA24AFD8721EEBCE51A16B431">5642</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="3BA1B1B3ACD7F8D1D7A0C450130E6847" type="issue">
|
||||
<mods:number id="30383512FC61405205622421F3A79D56">5</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="5144708452D54D1385630601B2C11F35" unit="page">
|
||||
<mods:start id="B74A22F4155F4472C9833BA27D62ED0E">451</mods:start>
|
||||
<mods:end id="4558AC6194B4838B75334CA187C31F16">475</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="36E14B33652288409429C9C9095901B2">
|
||||
<mods:url id="545DB30F8E81C6D6CF12BB6C044BCF47">https://doi.org/10.11646/zootaxa.5642.5.3</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="8FC94D13CC804341CB1FA90691D36EE4">journal article</mods:classification>
|
||||
<mods:identifier id="5B8F8AE131EB9968396B0118EB2A19BB" type="DOI">10.11646/zootaxa.5642.5.3</mods:identifier>
|
||||
<mods:identifier id="E167E3B99881C7B12391B7CDE15F70E6" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="A7E946CB5043ED3E80357CF5557EF086" type="Zenodo-Dep">15818151</mods:identifier>
|
||||
<mods:identifier id="2EA890CB6C0697C410532D804165B8D8" type="ZooBank">6CAAAAD8-0347-43C8-B3E9-D7011B436C19</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03F7FE24FFD9FFE7FF551CB5E31AFA9C" ID-DOI="http://doi.org/10.5281/zenodo.15818148" ID-Zenodo-Dep="15818148" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C" httpUri="http://treatment.plazi.org/id/03F7FE24FFD9FFE7FF551CB5E31AFA9C" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
|
||||
<subSubSection id="C3441CB9FFD9FFE6FF551CB5E7B4F93F" box="[151,500,1683,1710]" pageId="8" pageNumber="459" type="nomenclature">
|
||||
<paragraph id="8BE14F32FFD9FFE6FF551CB5E7B4F93F" blockId="8.[151,500,1683,1747]" box="[151,500,1683,1710]" pageId="8" pageNumber="459">
|
||||
<heading id="D0A9F85EFFD9FFE6FF551CB5E7B4F93F" bold="true" box="[151,500,1683,1710]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
|
||||
<taxonomicName id="4C5E34B1FFD9FFE6FF551CB5E7B4F93F" authority="Kirby, 1893" authorityName="Kirby" authorityYear="1893" box="[151,500,1683,1710]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD9FFE6FF551CB5E7B4F93F" bold="true" box="[151,500,1683,1710]" pageId="8" pageNumber="459">
|
||||
<emphasis id="B92A9320FFD9FFE6FF551CB5E719F93F" bold="true" box="[151,345,1683,1710]" italics="true" pageId="8" pageNumber="459">Lestes decipiens</emphasis>
|
||||
<bibRefCitation id="EFCF32C3FFD9FFE6FEA31CB2E7B4F93F" author="Kirby, J." box="[353,500,1683,1710]" pageId="8" pageNumber="459" pagination="545 - 566" refId="ref14162" refString="Kirby, J. (1893) Catalogue of the described Neuroptera Odonata (Dragonflies) of Ceylon, with descritions of new species. The Journal of the Linnaean Society of London, Zoology, 24, 545-566. https://doi.org/10.1111/j.1096-3642.1894.tb02495.x" type="journal article" year="1893">Kirby, 1893</bibRefCitation>
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3441CB9FFD9FFE7FF551C99E31AFA9C" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="description">
|
||||
<paragraph id="8BE14F32FFD9FFE6FF551C99E7A7F943" blockId="8.[151,500,1683,1747]" box="[151,487,1720,1747]" pageId="8" pageNumber="459">
|
||||
(
|
||||
<figureCitation id="136553B7FFD9FFE6FF5D1C99E751F943" box="[159,273,1720,1746]" captionStart="FIGURE 1" captionStartId="4.[151,250,1636,1660]" captionTargetBox="[400,1192,184,1607]" captionTargetId="figure-20@4.[393,1194,179,1610]" captionTargetPageId="4" captionText="FIGURE 1. Drawings of Lestes spp. (all males but i-right female) from literature, with original identifications: a, “Lestes praemorsus praemorsus”, Philippines, Luzon, Bilucao (Lieftinck 1949: fig. 19); b, “Lestes praemorsus praemorsus”, N. New Guinea, Hollandia (Lieftinck 1949: fig. 20); c, “Lestes praemorsus decipiens”, India, Coorg, Gonocopal (Lieftinck 1949: fig. 17); d, “Lestes praemorsus decipiens”, Indonesia, Java, Buitenzorg (Lieftinck 1949: fig. 18); e, “Lestes dorothea” from the original description (Fraser 1924: plate XXVI, fig. 8); f, “Lestes praemorsa” (in fact Lestes dorothea) (Fraser 1934: fig. 12); g, “Lestes dorothea” (in fact Lestes decipiens, restored in species rank in this paper) (Fraser 1934: fig. 13); h, “Lestes sp.1”, Laos, Champasak Province, Bolaven Plateau, Paksong (Yokoi & Souphanthong 2014: fig. 5); i-j, “Lestes praecellens”, holotype (i-left,-j) and allotype (i-right) from SW Java;, Oedjoeng Benteng Bay (Lieftinck 1937: figs 2–3); k–l, “Lestes praemorsus decipiens”, Taiwan (Asahina 1985: figs 25–26); m, “Lestes dorothea” (Orr 2005: 28); a-left, b-left, c-left, d-left, e, f-left, g-h, j-right, l-right, m-left–anal appendages in dorsal view; a-right, b-right, c-right, d-right, f-right, j-left–the same in lateral view; i-left, k–thorax in dorsal view; i-right, m-right–the same in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.15818153" httpUri="https://zenodo.org/record/15818153/files/figure.png" pageId="8" pageNumber="459">Figs 1c–d</figureCitation>
|
||||
;
|
||||
<figureCitation id="136553B7FFD9FFE6FEDF1C99E717F942" box="[285,343,1720,1747]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="8" pageNumber="459">5e–h</figureCitation>
|
||||
;
|
||||
<figureCitation id="136553B7FFD9FFE6FEA01C99E7D2F943" box="[354,402,1720,1746]" captionStart="FIGURE 8" captionStartId="13.[151,250,1302,1326]" captionTargetBox="[177,1411,181,1278]" captionTargetId="figure-18@13.[177,1411,181,1278]" captionTargetPageId="13" captionText="FIGURE 8. The allotype of Lestes dorothea, from S. India, Greenfields, Sidapur. 26 iv1923, F.C. Fraser, preserved in the BMNH collection: a–dorsal general view; b–labels, c,d–end of abdomen in lateral and dorsal views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818167" httpUri="https://zenodo.org/record/15818167/files/figure.png" pageId="8" pageNumber="459">8f–j</figureCitation>
|
||||
;
|
||||
<figureCitation id="136553B7FFD9FFE6FE5C1C99E79AF943" box="[414,474,1720,1746]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="14.[151,250,1943,1967]" captionStartId-1="15.[151,250,1944,1968]" captionTargetBox-0="[467,1121,181,1919]" captionTargetBox-1="[455,1133,181,1920]" captionTargetId-0="figure-20@14.[467,1121,181,1919]" captionTargetId-1="figure-18@15.[455,1133,181,1920]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURE 9. The lectotype, male, of Lestes decipens, from Ceylon, Mahagany, 20 ix 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." captionText-1="FIGURE 10. A paralectotype, female (incorrectly designated as “the allotype” by Kimmins (1970)) of Lestes decipiens, from Ceylon, 6th Milestone, Nilavelli Road, 9 xii 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.15818171" figureDoi-1="http://doi.org/10.5281/zenodo.15818173" httpUri-0="https://zenodo.org/record/15818171/files/figure.png" httpUri-1="https://zenodo.org/record/15818173/files/figure.png" pageId="8" pageNumber="459">9–10</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD9FFE7FF551D21E526FD88" blockId="8.[151,1437,1791,2035]" lastBlockId="9.[151,1437,150,1293]" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
|
||||
<emphasis id="B92A9320FFD9FFE6FF551D21E7C0F88B" box="[151,384,1792,1818]" italics="true" pageId="8" pageNumber="459">Specimens examined</emphasis>
|
||||
.
|
||||
<materialsCitation id="3B36456FFFD9FFE6FE521CDEE4A5F88A" box="[400,741,1791,1819]" country="Cambodia" location="Cambodia" pageId="8" pageNumber="459" specimenCount="2" specimenCount-male="2">
|
||||
<emphasis id="B92A9320FFD9FFE6FE521CDEE456F888" bold="true" box="[400,534,1791,1817]" pageId="8" pageNumber="459">
|
||||
<collectingCountry id="F3490FA2FFD9FFE6FE521CDEE451F888" box="[400,529,1791,1817]" name="Cambodia" pageId="8" pageNumber="459">Cambodia</collectingCountry>
|
||||
:
|
||||
</emphasis>
|
||||
<specimenCount id="9D5884BBFFD9FFE6FDDC1D21E41EF88B" box="[542,606,1792,1818]" count="2" pageId="8" pageNumber="459" type="male">2 ♂♂</specimenCount>
|
||||
(
|
||||
<figureCitation id="136553B7FFD9FFE6FDAC1D21E49EF88A" box="[622,734,1792,1819]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="8" pageNumber="459">Fig. 5e–h</figureCitation>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B36456FFFD9FFE6FD301D21E221F8AF" collectingDate="2018-06-27" collectorName="O. Kosterin" country="Cambodia" elevation="570" latitude="12.55" location="15 km NNE of Sen Momorom" longLatPrecision="77" longitude="107.258" pageId="8" pageNumber="459" specimenCount="1" specimenCount-female="1" stateProvince="Mondolkiri">
|
||||
<specimenCount id="9D5884BBFFD9FFE6FD301D21E55DF88B" box="[754,797,1792,1818]" count="1" pageId="8" pageNumber="459" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD9FFE6FCEE1D21E25AF88B" box="[812,1050,1792,1818]" country="Cambodia" name="Mondolkiri" pageId="8" pageNumber="459">Mondulkiri Province</collectingRegion>
|
||||
,
|
||||
<location id="8E8119E9FFD9FFE6FBE41D21E3C4F88B" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD9FFE6FBE41D21E3C4F88B" box="[1062,1412,1792,1819]" country="Cambodia" latitude="12.55" longLatPrecision="77" longitude="107.258" name="15 km NNE of Sen Momorom" pageId="8" pageNumber="459" stateProvince="Mondolkiri">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FBE41D21E22EF88B" box="[1062,1134,1792,1819]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="459" unit="km" value="15.0">15 km</quantity>
|
||||
NNE of Sen Momorom
|
||||
</location>
|
||||
, a large sedgy swamp,
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FEBC1D05E7B5F8AF" box="[382,501,1828,1855]" degrees="12.550" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="55" value="12.55">12.550° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FDC31D05E4C2F8AF" box="[513,642,1828,1855]" degrees="107.258" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="55" value="107.258">107.258° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD9FFE6FD4D1D05E554F8AF" box="[655,788,1828,1855]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="459" unit="m" value="570.0">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FD4D1D05E497F8AF" box="[655,727,1828,1855]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="459" unit="m" value="570.0">570 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD9FFE6FCE31D05E5DDF8AF" box="[801,925,1828,1854]" pageId="8" pageNumber="459" value="2018-06-27">
|
||||
<collectingDate id="EFA4901AFFD9FFE6FCE31D05E5DDF8AF" box="[801,925,1828,1854]" pageId="8" pageNumber="459" value="2018-06-27">27 vi 2018</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD9FFE6FC6B1D05E26CF8AF" box="[937,1068,1828,1854]" pageId="8" pageNumber="459">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD9FFE6FBAF1D05E5ABF817" collectingDate="2016-07-27" collectorName="O. Kosterin" country="Cambodia" elevation="83" latitude="13.8" location="Sam'ang village" longLatPrecision="76" longitude="105.866" municipality="Thala Barivat District" pageId="8" pageNumber="459" specimenCount="1" specimenCount-male="1" stateProvince="Stung Treng">
|
||||
<specimenCount id="9D5884BBFFD9FFE6FBAF1D05E2D2F8AF" box="[1133,1170,1828,1854]" count="1" pageId="8" pageNumber="459" type="male">1♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD9FFE6FB5D1D05E3D8F8AF" box="[1183,1432,1828,1854]" country="Cambodia" name="Stung Treng" pageId="8" pageNumber="459">Stung Treng Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6B85D548FFD9FFE6FF551D69E7D2F8F3" box="[151,402,1864,1890]" pageId="8" pageNumber="459">Thala Barivat District</collectingMunicipality>
|
||||
,
|
||||
<locationDeviation id="9A535610FFD9FFE6FE5D1D69E4A0F8F3" box="[415,736,1864,1891]" location="Stung Treng" pageId="8" pageNumber="459">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FE5D1D69E7BDF8F3" box="[415,509,1864,1891]" metricMagnitude="4" metricUnit="m" metricValue="3.25" pageId="8" pageNumber="459" unit="km" value="32.5">32.5 km</quantity>
|
||||
NNW
|
||||
<collectingRegion id="499A81D0FFD9FFE6FD931D69E4A0F8F3" box="[593,736,1864,1890]" country="Cambodia" name="Stung Treng" pageId="8" pageNumber="459">Stung Treng</collectingRegion>
|
||||
</locationDeviation>
|
||||
,
|
||||
<quantity id="4CA6E2D7FFD9FFE6FD2F1D69E57DF8F3" box="[749,829,1864,1891]" metricMagnitude="3" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="459" unit="km" value="9.5">9.5 km</quantity>
|
||||
NE
|
||||
<location id="8E8119E9FFD9FFE6FCB61D69E26FF8F3" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD9FFE6FCB61D69E26FF8F3" box="[884,1071,1864,1890]" country="Cambodia" latitude="13.8" longLatPrecision="76" longitude="105.866" municipality="Thala Barivat District" name="Sam'ang village" pageId="8" pageNumber="459" stateProvince="Stung Treng">Sam’ang village</location>
|
||||
, 13.798°–
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FB611D69E35BF8F3" box="[1187,1307,1864,1890]" degrees="13.800" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="55" value="13.8">13.800° N</geoCoordinate>
|
||||
, 105.860°–
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FF551D4DE755F817" box="[151,277,1900,1927]" degrees="105.866" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="55" value="105.866">105.866° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD9FFE6FEE21D4DE7FBF817" box="[288,443,1900,1927]" metricMagnitude="1" metricUnit="m" metricValue="8.3" metricValueMax="8.5" metricValueMin="8.1" pageId="8" pageNumber="459" unit="m" value="83.0" valueMax="85.0" valueMin="81.0">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FEE21D4DE7C1F817" box="[288,385,1900,1927]" metricMagnitude="1" metricUnit="m" metricValue="8.3" metricValueMax="8.5" metricValueMin="8.1" pageId="8" pageNumber="459" unit="m" value="83.0" valueMax="85.0" valueMin="81.0">81–85 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
, a rivulet with a pond,
|
||||
<date id="FFE069F2FFD9FFE6FD701D4DE570F817" box="[690,816,1900,1926]" pageId="8" pageNumber="459" value="2016-07-27">
|
||||
<collectingDate id="EFA4901AFFD9FFE6FD701D4DE570F817" box="[690,816,1900,1926]" pageId="8" pageNumber="459" value="2016-07-27">27 vii 2016</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD9FFE6FCFB1D4DE5F9F817" box="[825,953,1900,1926]" pageId="8" pageNumber="459">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD9FFE6FC371D4DE3C3F83B" collectingDate="2010-12-09" collectorName="O. Kosterin" country="Cambodia" elevation="926" latitude="10.6456" location="Pond'" longLatPrecision="7" longitude="104.0392" municipality="Bokor" pageId="8" pageNumber="459" specimenCount="1" specimenCount-male="1" stateProvince="Kampot">
|
||||
<specimenCount id="9D5884BBFFD9FFE6FC371D4DE25FF817" box="[1013,1055,1900,1926]" count="1" pageId="8" pageNumber="459" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD9FFE6FBEB1D4DE2AEF817" box="[1065,1262,1900,1926]" country="Cambodia" name="Kampot" pageId="8" pageNumber="459">Kampot Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6B85D548FFD9FFE6FB3A1D4DE37EF817" box="[1272,1342,1900,1926]" pageId="8" pageNumber="459">Bokor</collectingMunicipality>
|
||||
Plateau,
|
||||
<taxonomicName id="4C5E34B1FFD9FFE6FF551DB1E763F83B" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[151,291,1936,1962]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="praemorsus">‘Praemorsus</taxonomicName>
|
||||
<location id="8E8119E9FFD9FFE6FEE81DB1E730F83B" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD9FFE6FEE81DB1E730F83B" box="[298,368,1936,1962]" country="Cambodia" latitude="10.6456" longLatPrecision="7" longitude="104.0392" municipality="Bokor" name="Pond'" pageId="8" pageNumber="459" stateProvince="Kampot">Pond’</location>
|
||||
,
|
||||
<locationDeviation id="9A535610FFD9FFE6FEB81DB1E4D1F83B" box="[378,657,1936,1962]" location="Bokor Palace" pageId="8" pageNumber="459">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FEB81DB1E787F83B" box="[378,455,1936,1962]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="459" unit="km" value="2.9">2.9 km</quantity>
|
||||
NE Bokor Palace
|
||||
</locationDeviation>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FD5E1DB1E55FF83B" box="[668,799,1936,1963]" degrees="10.6456" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="5" value="10.6456">10.6456° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FCE81DB1E5F7F83B" box="[810,951,1936,1962]" degrees="104.0392" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="5" value="104.0392">104.0392° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD9FFE6FC011DB1E205F83B" box="[963,1093,1936,1962]" metricMagnitude="2" metricUnit="m" metricValue="9.26" pageId="8" pageNumber="459" unit="m" value="926.0">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FC011DB1E249F83B" box="[963,1033,1936,1962]" metricMagnitude="2" metricUnit="m" metricValue="9.26" pageId="8" pageNumber="459" unit="m" value="926.0">926 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD9FFE6FB931DB1E284F83B" box="[1105,1220,1936,1962]" pageId="8" pageNumber="459" value="2010-12-09">
|
||||
<collectingDate id="EFA4901AFFD9FFE6FB931DB1E284F83B" box="[1105,1220,1936,1962]" pageId="8" pageNumber="459" value="2010-12-09">9 xii 2010</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD9FFE6FB0C1DB1E310F83B" box="[1230,1360,1936,1962]" pageId="8" pageNumber="459">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD9FFE6FA4C1DB1E550F863" collectingDate="2011-08-18" collectorName="O. Kosterin" country="Cambodia" elevation="926" latitude="10.6583" location="Popokvil River" longLatPrecision="7" longitude="104.0525" municipality="Bokor" pageId="8" pageNumber="459" specimenCount="1" specimenCount-male="1" stateProvince="Kampot">
|
||||
<specimenCount id="9D5884BBFFD9FFE6FA4C1DB1E6EEF85F" count="1" pageId="8" pageNumber="459" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD9FFE6FF781D95E7C1F85F" box="[186,385,1972,1998]" country="Cambodia" name="Kampot" pageId="8" pageNumber="459">Kampot Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6B85D548FFD9FFE6FE4E1D95E792F85F" box="[396,466,1972,1998]" pageId="8" pageNumber="459">Bokor</collectingMunicipality>
|
||||
Plateau, the
|
||||
<location id="8E8119E9FFD9FFE6FDA31D95E54BF85F" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD9FFE6FDA31D95E54BF85F" box="[609,779,1972,1998]" country="Cambodia" latitude="10.6583" longLatPrecision="7" longitude="104.0525" municipality="Bokor" name="Popokvil River" pageId="8" pageNumber="459" stateProvince="Kampot">Popokvil River</location>
|
||||
at the lower bridge,
|
||||
<locationDeviation id="9A535610FFD9FFE6FC331D95E34AF85F" box="[1009,1290,1972,1998]" location="Bokor Palace" pageId="8" pageNumber="459">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FC331D95E27FF85F" box="[1009,1087,1972,1998]" metricMagnitude="3" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="459" unit="km" value="4.9">4.9 km</quantity>
|
||||
NE Bokor Palace
|
||||
</locationDeviation>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FAD61D95E3D8F85F" box="[1300,1432,1972,1999]" degrees="10.6583" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="5" value="10.6583">10.6583° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD9FFE6FF551DF9E766F863" box="[151,294,2008,2035]" degrees="104.0525" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="5" value="104.0525">104.0525° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD9FFE6FEF01DF9E7F7F863" box="[306,439,2008,2034]" metricMagnitude="2" metricUnit="m" metricValue="9.26" pageId="8" pageNumber="459" unit="m" value="926.0">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FEF01DF9E73AF863" box="[306,378,2008,2034]" metricMagnitude="2" metricUnit="m" metricValue="9.26" pageId="8" pageNumber="459" unit="m" value="926.0">926 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD9FFE6FE011DF9E40EF863" box="[451,590,2008,2034]" pageId="8" pageNumber="459" value="2011-08-18">
|
||||
<collectingDate id="EFA4901AFFD9FFE6FE011DF9E40EF863" box="[451,590,2008,2034]" pageId="8" pageNumber="459" value="2011-08-18">18 viii 2011</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD9FFE6FD9B1DF9E49CF863" box="[601,732,2008,2034]" pageId="8" pageNumber="459">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD9FFE7FCDE1DF9E46DFF44" collectingDate="2013-05-26" collectorName="O. Kosterin" country="Cambodia" elevation="53" lastPageId="9" lastPageNumber="460" latitude="10.674" location="Kbal Chhay Waterfall" longLatPrecision="77" longitude="103.607" municipality="Preah Sihanouk Province" pageId="8" pageNumber="459" specimenCount="1" specimenCount-male="1" stateProvince="Sihanoukville">
|
||||
<specimenCount id="9D5884BBFFD9FFE6FCDE1DF9E508F863" box="[796,840,2008,2034]" count="1" pageId="8" pageNumber="459" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingMunicipality id="6B85D548FFD9FFE6FC971DF9E233F863" box="[853,1139,2008,2034]" pageId="8" pageNumber="459">Preah Sihanouk Province</collectingMunicipality>
|
||||
,
|
||||
<locationDeviation id="9A535610FFD9FFE6FBBD1DF9E3D8F863" box="[1151,1432,2008,2034]" location="Sihanoukville" pageId="8" pageNumber="459">
|
||||
<quantity id="4CA6E2D7FFD9FFE6FBBD1DF9E286F863" box="[1151,1222,2008,2034]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="459" unit="km" value="11.0">11 km</quantity>
|
||||
NE
|
||||
<collectingRegion id="499A81D0FFD9FFE6FB381DF9E3D8F863" box="[1274,1432,2008,2034]" country="Cambodia" name="Sihanoukville" pageId="8" pageNumber="459">Sihanoukville</collectingRegion>
|
||||
</locationDeviation>
|
||||
,
|
||||
<location id="8E8119E9FFD8FFE7FF551AB6E7CDFF20" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FF551AB6E7CDFF20" box="[151,397,151,177]" country="Cambodia" latitude="10.674" longLatPrecision="77" longitude="103.607" municipality="Preah Sihanouk Province" name="Kbal Chhay Waterfall" pageId="9" pageNumber="460" stateProvince="Sihanoukville">Kbal Chhay Waterfall</location>
|
||||
env., a pond at the main river right bank,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FCAF1AB6E5A5FF20" box="[877,997,151,177]" degrees="10.674" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="55" value="10.674">10.674° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FC331AB6E232FF20" box="[1009,1138,151,177]" degrees="103.607" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="55" value="103.607">103.607° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD8FFE7FB421AB7E2B7FF20" box="[1152,1271,150,177]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="460" unit="m" value="53.0">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FB421AB7E2FAFF20" box="[1152,1210,150,177]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="460" unit="m" value="53.0">53 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
, a rivulet with a pond,
|
||||
<date id="FFE069F2FFD8FFE7FF311A9AE728FF44" box="[243,360,187,213]" pageId="9" pageNumber="460" value="2013-05-26">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FF311A9AE728FF44" box="[243,360,187,213]" pageId="9" pageNumber="460" value="2013-05-26">26 v 2013</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD8FFE7FEB61A9AE7B8FF44" box="[372,504,187,213]" pageId="9" pageNumber="460">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD8FFE7FDF81A9AE574FF68" collectingDate="2011-08-25" collectorName="O. Kosterin" country="Cambodia" elevation="371" latitude="11.6703" location="Triangularis" longLatPrecision="7" longitude="103.4086" municipality="Thma Bang village" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" stateProvince="Koh Kong">
|
||||
<specimenCount id="9D5884BBFFD8FFE7FDF81A9AE427FF44" box="[570,615,187,213]" count="1" pageId="9" pageNumber="460" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FDB61A9AE519FF44" box="[628,857,187,213]" country="Cambodia" name="Koh Kong" pageId="9" pageNumber="460">Koh Kong Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6B85D548FFD8FFE7FCA71A9AE201FF44" box="[869,1089,187,213]" pageId="9" pageNumber="460">Thma Bang village</collectingMunicipality>
|
||||
, ‘
|
||||
<location id="8E8119E9FFD8FFE7FB931A9AE29EFF44" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FB931A9AE29EFF44" box="[1105,1246,187,213]" country="Cambodia" latitude="11.6703" longLatPrecision="7" longitude="103.4086" municipality="Thma Bang village" name="Triangularis" pageId="9" pageNumber="460" stateProvince="Koh Kong">Triangularis</location>
|
||||
pond’,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FAF51A9AE6EEFF68" degrees="11.6703" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="5" value="11.6703">11.6703° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FF781AFEE709FF68" box="[186,329,223,249]" degrees="103.4086" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="5" value="103.4086">103.4086° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD8FFE7FE971AFEE79AFF68" box="[341,474,223,249]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="9" pageNumber="460" unit="m" value="371.0">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FE971AFEE7DDFF68" box="[341,413,223,249]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="9" pageNumber="460" unit="m" value="371.0">371 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FE251AFFE432FF68" box="[487,626,222,249]" pageId="9" pageNumber="460" value="2011-08-25">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FE251AFFE432FF68" box="[487,626,222,249]" pageId="9" pageNumber="460" value="2011-08-25">25 viii 2011</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD8FFE7FDBF1AFEE540FF68" box="[637,768,223,249]" pageId="9" pageNumber="460">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD8FFE7FC821AFEE3D8FE8C" collectingDate="2011-08-13" collectorName="O. Kosterin" country="Cambodia" elevation="316" latitude="11.66" location="Aciagrion Rivulet'" longLatPrecision="7" longitude="103.0956" pageId="9" pageNumber="460" specimenCount="3" specimenCount-male="3" stateProvince="Koh Kong">
|
||||
<specimenCount id="9D5884BBFFD8FFE7FC821AFEE5C2FF68" box="[832,898,223,249]" count="3" pageId="9" pageNumber="460" type="male">3 ♂♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FC4D1AFEE232FF68" box="[911,1138,223,249]" country="Cambodia" name="Koh Kong" pageId="9" pageNumber="460">Koh Kong Province</collectingRegion>
|
||||
,
|
||||
<locationDeviation id="9A535610FFD8FFE7FBBC1AFEE3C4FF68" box="[1150,1412,223,249]" location="Koh Kong" pageId="9" pageNumber="460">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FBBC1AFEE286FF68" box="[1150,1222,223,249]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="460" unit="km" value="13.0">13 km</quantity>
|
||||
ENE
|
||||
<collectingRegion id="499A81D0FFD8FFE7FAC91AFEE3C4FF68" box="[1291,1412,223,249]" country="Cambodia" name="Koh Kong" pageId="9" pageNumber="460">Koh Kong</collectingRegion>
|
||||
</locationDeviation>
|
||||
, a grassy swamp at ‘
|
||||
<location id="8E8119E9FFD8FFE7FE9E1B22E473FE8C" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FE9E1B22E473FE8C" box="[348,563,259,285]" country="Cambodia" latitude="11.66" longLatPrecision="7" longitude="103.0956" name="Aciagrion Rivulet'" pageId="9" pageNumber="460" stateProvince="Koh Kong">Aciagrion Rivulet’</location>
|
||||
source,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FD531B22E555FE8C" box="[657,789,259,285]" degrees="11.6600" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="5" value="11.66">11.6600° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FCE21B23E5EFFE8C" box="[800,943,258,285]" degrees="103.0956" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="5" value="103.0956">103.0956° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD8FFE7FC791B22E27FFE8C" box="[955,1087,259,285]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="9" pageNumber="460" unit="m" value="316.0">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FC791B22E242FE8C" box="[955,1026,259,285]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="9" pageNumber="460" unit="m" value="316.0">316 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FB8E1B22E296FE8C" box="[1100,1238,259,285]" pageId="9" pageNumber="460" value="2011-08-13">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FB8E1B22E296FE8C" box="[1100,1238,259,285]" pageId="9" pageNumber="460" value="2011-08-13">13 viii 2011</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD8FFE7FB231B22E324FE8C" box="[1249,1380,259,285]" pageId="9" pageNumber="460">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD8FFE7FF551B06E576FEF4" collectingDate="2011-08-26" collectorName="O. Kosterin" country="Cambodia" elevation="316" latitude="11.584" location="Peam Krasaop village" longLatPrecision="77" longitude="102.985" municipality="Calamorum" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" stateProvince="Koh Kong">
|
||||
<specimenCount id="9D5884BBFFD8FFE7FF551B06E682FED0" box="[151,194,295,321]" count="1" pageId="9" pageNumber="460" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FF0C1B06E7EEFED0" box="[206,430,295,321]" country="Cambodia" name="Koh Kong" pageId="9" pageNumber="460">Koh Kong Province</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FE7B1B06E46EFED0" box="[441,558,295,321]" country="Cambodia" name="Koh Kong" pageId="9" pageNumber="460">Koh Kong</collectingRegion>
|
||||
S suburbs, ‘
|
||||
<collectingMunicipality id="6B85D548FFD8FFE7FD701B06E57BFED0" box="[690,827,295,321]" pageId="9" pageNumber="460">Calamorum</collectingMunicipality>
|
||||
ponds’ E of the road to
|
||||
<location id="8E8119E9FFD8FFE7FB861B06E37BFED0" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FB861B06E37BFED0" box="[1092,1339,295,321]" country="Cambodia" latitude="11.584" longLatPrecision="77" longitude="102.985" municipality="Calamorum" name="Peam Krasaop village" pageId="9" pageNumber="460" stateProvince="Koh Kong">Peam Krasaop village</location>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FA871B07E6EEFEF4" degrees="11.584" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="55" value="11.584">11.584° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FF7E1B6BE77EFEF4" box="[188,318,330,357]" degrees="102.985" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="55" value="102.985">102.985° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD8FFE7FE8E1B6AE794FEF4" box="[332,468,331,357]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="9" pageNumber="460" unit="m" value="316.0">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FE8E1B6AE7D6FEF4" box="[332,406,331,357]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="9" pageNumber="460" unit="m" value="316.0">316 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FE211B6AE431FEF4" box="[483,625,331,357]" pageId="9" pageNumber="460" value="2011-08-26">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FE211B6AE431FEF4" box="[483,625,331,357]" pageId="9" pageNumber="460" value="2011-08-26">26 viii 2011</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD8FFE7FDBC1B6AE543FEF4" box="[638,771,331,357]" pageId="9" pageNumber="460">O. Kosterin</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B36456FFFD8FFE7FCFD1B6BE406FE18" collectingDate="2015-03-15" collectorName="Makbun" country="Thailand" location="Wang Nam Khiao" pageId="9" pageNumber="460" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Ratchasima">
|
||||
<emphasis id="B92A9320FFD8FFE7FCFD1B6BE5F5FEF5" bold="true" box="[831,949,330,356]" pageId="9" pageNumber="460">
|
||||
<collectingCountry id="F3490FA2FFD8FFE7FCFD1B6BE5F0FEF5" box="[831,944,330,356]" name="Thailand" pageId="9" pageNumber="460">Thailand</collectingCountry>
|
||||
:
|
||||
</emphasis>
|
||||
<specimenCount id="9D5884BBFFD8FFE7FC7C1B6AE5AAFEF4" box="[958,1002,331,357]" count="1" pageId="9" pageNumber="460" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FC391B6AE30EFEF4" box="[1019,1358,331,357]" country="Thailand" name="Nakhon Ratchasima" pageId="9" pageNumber="460">Nakhon Ratchasima Province</collectingRegion>
|
||||
,
|
||||
<location id="8E8119E9FFD8FFE7FA9E1B6AE75DFE18" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FA9E1B6AE75DFE18" country="Thailand" name="Wang Nam Khiao" pageId="9" pageNumber="460" stateProvince="Nakhon Ratchasima">Wang Nam Khiao</location>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FEEB1B4FE7E8FE18" box="[297,424,366,393]" pageId="9" pageNumber="460" value="2015-03-15">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FEEB1B4FE7E8FE18" box="[297,424,366,393]" pageId="9" pageNumber="460" value="2015-03-15">15 iii 2015</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD8FFE7FE761B4EE451FE18" box="[436,529,367,393]" pageId="9" pageNumber="460">Makbun</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD8FFE7FD911B4EE280FE18" box="[595,1216,367,393]" collectingDate="2018-08-01" collectorName="Makbun" country="Thailand" location="Muang" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" stateProvince="Krabi">
|
||||
<specimenCount id="9D5884BBFFD8FFE7FD911B4EE4C0FE18" box="[595,640,367,393]" count="1" pageId="9" pageNumber="460" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FD4F1B4EE57BFE18" box="[653,827,367,393]" country="Thailand" name="Krabi" pageId="9" pageNumber="460">Krabi Province</collectingRegion>
|
||||
,
|
||||
<location id="8E8119E9FFD8FFE7FC8A1B4EE5DAFE18" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FC8A1B4EE5DAFE18" box="[840,922,367,393]" country="Thailand" name="Muang" pageId="9" pageNumber="460" stateProvince="Krabi">Muang</location>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FC641B4EE265FE18" box="[934,1061,367,393]" pageId="9" pageNumber="460" value="2018-08-01">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FC641B4EE265FE18" box="[934,1061,367,393]" pageId="9" pageNumber="460" value="2018-08-01">1 viii 2018</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26AB2AE4FFD8FFE7FBF31B4EE2CEFE18" box="[1073,1166,367,393]" pageId="9" pageNumber="460">Makbun</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B36456FFFD8FFE7FB0A1B4FE531FE40" collectingDate="2019-10-16" collectionCode="T, ZCDTU" collectorName="Phan" country="Vietnam" county="Qu" elevation="46" latitude="15.3757" location="Tam My Tay Commune" longLatPrecision="7" longitude="108.5761" municipality="Nui Thanh District" pageId="9" pageNumber="460" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Nam Province">
|
||||
<emphasis id="B92A9320FFD8FFE7FB0A1B4FE377FE19" bold="true" box="[1224,1335,366,392]" pageId="9" pageNumber="460">
|
||||
<collectingCountry id="F3490FA2FFD8FFE7FB0A1B4FE372FE19" box="[1224,1330,366,392]" name="Vietnam" pageId="9" pageNumber="460">Vietnam</collectingCountry>
|
||||
:
|
||||
</emphasis>
|
||||
<specimenCount id="9D5884BBFFD8FFE7FAFD1B4EE3C1FE18" box="[1343,1409,367,393]" count="2" pageId="9" pageNumber="460" type="male">2 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D5884BBFFD8FFE7FA4C1B4EE681FE3C" count="2" pageId="9" pageNumber="460" type="female">2 ♀♀</specimenCount>
|
||||
(collected in tandems;
|
||||
<figureCitation id="136553B7FFD8FFE7FE041BB2E469FE3C" box="[454,553,403,429]" captionStart="FIGURE 8" captionStartId="13.[151,250,1302,1326]" captionTargetBox="[177,1411,181,1278]" captionTargetId="figure-18@13.[177,1411,181,1278]" captionTargetPageId="13" captionText="FIGURE 8. The allotype of Lestes dorothea, from S. India, Greenfields, Sidapur. 26 iv1923, F.C. Fraser, preserved in the BMNH collection: a–dorsal general view; b–labels, c,d–end of abdomen in lateral and dorsal views (photos by Dan Hall)." figureDoi="http://doi.org/10.5281/zenodo.15818167" httpUri="https://zenodo.org/record/15818167/files/figure.png" pageId="9" pageNumber="460">Fig. 8f–j</figureCitation>
|
||||
),
|
||||
<collectingCounty id="628037BEFFD8FFE7FDFF1BB3E420FE3C" box="[573,608,402,429]" pageId="9" pageNumber="460">Qu</collectingCounty>
|
||||
ảng
|
||||
<collectingRegion id="499A81D0FFD8FFE7FD4D1BB2E572FE3C" box="[655,818,403,429]" inferredBySuffix="true" pageId="9" pageNumber="460">Nam Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6B85D548FFD8FFE7FCFC1BB2E253FE3C" box="[830,1043,403,429]" pageId="9" pageNumber="460">Nui Thanh District</collectingMunicipality>
|
||||
,
|
||||
<location id="8E8119E9FFD8FFE7FBDC1BB2E36CFE3C" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FBDC1BB2E36CFE3C" box="[1054,1324,403,429]" country="Vietnam" county="Qu" latitude="15.3757" longLatPrecision="7" longitude="108.5761" municipality="Nui Thanh District" name="Tam My Tay Commune" pageId="9" pageNumber="460" stateProvince="Nam Province">Tam My Tay Commune</location>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FAF41BB3E6EEFE40" degrees="15.3757" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="5" value="15.3757">15.3757° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FF781B97E708FE40" box="[186,328,438,465]" degrees="108.5761" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="5" value="108.5761">108.5761° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD8FFE7FE961B96E78AFE40" box="[340,458,439,465]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="9" pageNumber="460" unit="m" value="46.0">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FE961B96E7CEFE40" box="[340,398,439,465]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="9" pageNumber="460" unit="m" value="46.0">46 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FE151B96E40AFE40" box="[471,586,439,465]" pageId="9" pageNumber="460" value="2019-10-16">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FE151B96E40AFE40" box="[471,586,439,465]" pageId="9" pageNumber="460" value="2019-10-16">16 x 2019</collectingDate>
|
||||
</date>
|
||||
, Q.
|
||||
<collectionCode id="ED4FD7F7FFD8FFE7FDB21B96E4C3FE40" box="[624,643,439,465]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="460">T</collectionCode>
|
||||
.
|
||||
<collectorName id="26AB2AE4FFD8FFE7FD4C1B96E486FE40" box="[654,710,439,465]" pageId="9" pageNumber="460">Phan</collectorName>
|
||||
leg. (
|
||||
<collectionCode id="ED4FD7F7FFD8FFE7FCC71B96E529FE40" box="[773,873,439,465]" pageId="9" pageNumber="460">ZCDTU</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD8FFE7FCBE1B96E520FE64" collectingDate="2023-05-24" collectionCode="T, ZCDTU" collectorName="Phan" country="Vietnam" elevation="622" latitude="11.5288" location="Bao Loc" longLatPrecision="7" longitude="107.7082" pageId="9" pageNumber="460" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong">
|
||||
<specimenCount id="9D5884BBFFD8FFE7FCBE1B96E5E7FE40" box="[892,935,439,465]" count="1" pageId="9" pageNumber="460" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FC771B96E2DBFE40" box="[949,1179,439,465]" country="Vietnam" name="Lam Dong" pageId="9" pageNumber="460">Lam Dong Province</collectingRegion>
|
||||
,
|
||||
<location id="8E8119E9FFD8FFE7FB641B96E349FE40" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FB641B96E349FE40" box="[1190,1289,439,465]" country="Vietnam" latitude="11.5288" longLatPrecision="7" longitude="107.7082" name="Bao Loc" pageId="9" pageNumber="460" stateProvince="Lam Dong">Bao Loc</location>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FAD71B97E3D8FE40" box="[1301,1432,438,465]" degrees="11.5288" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="5" value="11.5288">11.5288° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FF551BFAE766FE64" box="[151,294,475,501]" degrees="107.7082" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="5" value="107.7082">107.7082° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD8FFE7FEF01BFAE7F7FE64" box="[306,439,475,501]" metricMagnitude="2" metricUnit="m" metricValue="6.22" pageId="9" pageNumber="460" unit="m" value="622.0">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FEF01BFAE73AFE64" box="[306,378,475,501]" metricMagnitude="2" metricUnit="m" metricValue="6.22" pageId="9" pageNumber="460" unit="m" value="622.0">622 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FE061BFAE478FE64" box="[452,568,475,501]" pageId="9" pageNumber="460" value="2023-05-24">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FE061BFAE478FE64" box="[452,568,475,501]" pageId="9" pageNumber="460" value="2023-05-24">24 v 2023</collectingDate>
|
||||
</date>
|
||||
, Q.
|
||||
<collectionCode id="ED4FD7F7FFD8FFE7FD9C1BFAE431FE64" box="[606,625,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="460">T</collectionCode>
|
||||
.
|
||||
<collectorName id="26AB2AE4FFD8FFE7FDBE1BFAE4F4FE64" box="[636,692,475,501]" pageId="9" pageNumber="460">Phan</collectorName>
|
||||
leg. (
|
||||
<collectionCode id="ED4FD7F7FFD8FFE7FD361BFAE518FE64" box="[756,856,475,501]" pageId="9" pageNumber="460">ZCDTU</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B36456FFFD8FFE7FCAE1BFAE4A8FD88" collectingDate="2024-09-02" collectionCode="T" collectorName="Phan" country="Vietnam" elevation="1097" latitude="15.2091" location="Dak Glei" longLatPrecision="7" longitude="107.7358" pageId="9" pageNumber="460" specimenCount="4" specimenCount-male="4" stateProvince="Kon Tum">
|
||||
<specimenCount id="9D5884BBFFD8FFE7FCAE1BFAE5EDFE64" box="[876,941,475,501]" count="4" pageId="9" pageNumber="460" type="male">4 ♂♂</specimenCount>
|
||||
,
|
||||
<collectingRegion id="499A81D0FFD8FFE7FC781BFAE2D2FE64" box="[954,1170,475,501]" country="Vietnam" name="Kon Tum" pageId="9" pageNumber="460">Kon Tum Province</collectingRegion>
|
||||
,
|
||||
<location id="8E8119E9FFD8FFE7FB5C1BFAE347FE64" LSID="urn:lsid:plazi:treatment:03F7FE24FFD9FFE7FF551CB5E31AFA9C:8E8119E9FFD8FFE7FB5C1BFAE347FE64" box="[1182,1287,475,501]" country="Vietnam" latitude="15.2091" longLatPrecision="7" longitude="107.7358" name="Dak Glei" pageId="9" pageNumber="460" stateProvince="Kon Tum">Dak Glei</location>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FAD11BFBE3D8FE64" box="[1299,1432,474,501]" degrees="15.2091" direction="north" orientation="latitude" pageId="9" pageNumber="460" precision="5" value="15.2091">15.2091° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE6A29F5FFD8FFE7FF551BDFE765FD88" box="[151,293,510,537]" degrees="107.7358" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="5" value="107.7358">107.7358° E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0073A801FFD8FFE7FEF31BDEE783FD88" box="[305,451,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="9" pageNumber="460" unit="m" value="1097.0">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FEF31BDEE7C6FD88" box="[305,390,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="9" pageNumber="460" unit="m" value="1097.0">1097 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="FFE069F2FFD8FFE7FE0D1BDEE47CFD88" box="[463,572,511,537]" pageId="9" pageNumber="460" value="2024-09-02">
|
||||
<collectingDate id="EFA4901AFFD8FFE7FE0D1BDEE47CFD88" box="[463,572,511,537]" pageId="9" pageNumber="460" value="2024-09-02">2 ix 2024</collectingDate>
|
||||
</date>
|
||||
, Q.
|
||||
<collectionCode id="ED4FD7F7FFD8FFE7FDA31BDEE434FD88" box="[609,628,511,537]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="460">T</collectionCode>
|
||||
.
|
||||
<collectorName id="26AB2AE4FFD8FFE7FDBD1BDEE4F7FD88" box="[639,695,511,537]" pageId="9" pageNumber="460">Phan</collectorName>
|
||||
leg.
|
||||
</materialsCitation>
|
||||
(ZCDTU).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD8FFE7FF051802E5E7FC34" blockId="9.[151,1437,150,1293]" pageId="9" pageNumber="460">
|
||||
<emphasis id="B92A9320FFD8FFE7FF051802E705FDAC" box="[199,325,547,573]" italics="true" pageId="9" pageNumber="460">Characters</emphasis>
|
||||
. Male (
|
||||
<figureCitation id="136553B7FFD8FFE7FE581802E446FDAC" box="[410,518,546,573]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="9" pageNumber="460">Fig. 5e–h</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFD8FFE7FDD31802E461FDAC" box="[529,545,547,573]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[467,1121,181,1919]" captionTargetId="figure-20@14.[467,1121,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. The lectotype, male, of Lestes decipens, from Ceylon, Mahagany, 20 ix 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818171" httpUri="https://zenodo.org/record/15818171/files/figure.png" pageId="9" pageNumber="460">9</figureCitation>
|
||||
). The paraprocts are set more or less parallel to each other, so well seen in dorsal view (
|
||||
<figureCitation id="136553B7FFD8FFE7FF1C1866E772FDF0" box="[222,306,582,609]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="9" pageNumber="460">Fig. 5e</figureCitation>
|
||||
). Because of this position, the paraprocts extend well behind the level of the cercus tooth (
|
||||
<figureCitation id="136553B7FFD8FFE7FAFD1866E3D0FDF0" box="[1343,1424,582,609]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="9" pageNumber="460">Fig. 5f</figureCitation>
|
||||
). The length of the paraproct inner margin (as measured from the S10 border to the tip) comprises ca 46 % of the cercus length (as measured from the S10 border to the tip along the insect body axis). In ventral view (
|
||||
<figureCitation id="136553B7FFD8FFE7FAC518AEE315FD38" box="[1287,1365,654,681]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="9" pageNumber="460">Fig. 5f</figureCitation>
|
||||
), their outer margins are convex and inner margins slightly concave, so that their bluntly terminated apical parts, somewhat converge to each other, as seen also in dorsal view (
|
||||
<figureCitation id="136553B7FFD8FFE7FD2318F6E574FD60" box="[737,820,726,753]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="9" pageNumber="460">Fig. 5e</figureCitation>
|
||||
). In the oblique lateroventral view, the paraproct distal part is much broader and less S-like curved than in
|
||||
<taxonomicName id="4C5E34B1FFD8FFE7FD1718DAE516FC84" authorityName="Fraser" authorityYear="1924" box="[725,854,763,789]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="dorothea">
|
||||
<emphasis id="B92A9320FFD8FFE7FD1718DAE516FC84" box="[725,854,763,789]" italics="true" pageId="9" pageNumber="460">L. dorothea</emphasis>
|
||||
</taxonomicName>
|
||||
, with broadly blunt apices (
|
||||
<figureCitation id="136553B7FFD8FFE7FB4C18DAE2A2FC84" box="[1166,1250,762,789]" captionStart="FIGURE 5" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[232,1355,181,1884]" captionTargetId="figure-20@10.[232,1355,181,1884]" captionTargetPageId="10" captionText="FIGURE 5. Anal appendages of male specimens of Lestes dorothea (a–d) and L. praemorsus decipiens (e–h) collected at the same sedge swamp 15 km NNE of Sen Monorom, Mondulkiri Province, Cambodia, on 27 vi 2018; a,e–dorsal view; b,f–ventral view, c–g–lateral view; d–h–oblique lateroventral view. Scale bar = 1 mm. The pruinescence was washed off by acetone (OEK). Asterisk indicates the position of the paraproct tip." figureDoi="http://doi.org/10.5281/zenodo.15818161" httpUri="https://zenodo.org/record/15818161/files/figure.png" pageId="9" pageNumber="460">Fig. 5h</figureCitation>
|
||||
). In lateral view, the paraproct looks evenly tapering and slightly curved up, without distinction into the broader basal and narrower distal part. The appendages of the
|
||||
<typeStatus id="54E5F190FFD8FFE7FDD21962E432FCCC" box="[528,626,835,861]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C5E34B1FFD8FFE7FD571962E55BFCCC" authorityName="Kirby" authorityYear="1893" box="[661,795,835,861]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD8FFE7FD571962E55BFCCC" box="[661,795,835,861]" italics="true" pageId="9" pageNumber="460">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
are damaged so that the right cercus and left paraproct are missing (
|
||||
<figureCitation id="136553B7FFD8FFE7FF3F1946E70FFC10" box="[253,335,871,897]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[467,1121,181,1919]" captionTargetId="figure-20@14.[467,1121,181,1919]" captionTargetPageId="14" captionText="FIGURE 9. The lectotype, male, of Lestes decipens, from Ceylon, Mahagany, 20 ix 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818171" httpUri="https://zenodo.org/record/15818171/files/figure.png" pageId="9" pageNumber="460">Fig. 9e</figureCitation>
|
||||
). However, the remaining paraproct corresponds perfectly to our recent specimens of
|
||||
<taxonomicName id="4C5E34B1FFD8FFE7FACF1946E3D5FC10" authorityName="Kirby" authorityYear="1893" box="[1293,1429,871,897]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="decipiens">
|
||||
<emphasis id="B92A9320FFD8FFE7FACF1946E3D5FC10" box="[1293,1429,871,897]" italics="true" pageId="9" pageNumber="460">L. decipiens</emphasis>
|
||||
</taxonomicName>
|
||||
: it is directed behind and has the same shape, slightly curving inwards.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD8FFE7FF05198EE741FC7C" blockId="9.[151,1437,150,1293]" pageId="9" pageNumber="460">
|
||||
Measurements (mm). Hindwing 19.5–20,
|
||||
<quantity id="4CA6E2D7FFD8FFE7FD5C198EE4ACFC58" box="[670,748,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="5.2578" pageId="9" pageNumber="460" unit="in" value="20.7">20.7 in</quantity>
|
||||
the
|
||||
<typeStatus id="54E5F190FFD8FFE7FCD9198EE5C1FC58" box="[795,897,943,969]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
|
||||
; abdomen with appendages 29–29.5,
|
||||
<quantity id="4CA6E2D7FFD8FFE7FAE7198EE333FC58" box="[1317,1395,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="5.3848" pageId="9" pageNumber="460" unit="in" value="21.2">21.2 in</quantity>
|
||||
the
|
||||
<typeStatus id="54E5F190FFD8FFE7FF5519F2E6BDFC7C" box="[151,253,979,1005]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD8FFE7FF0519D6E705FB54" blockId="9.[151,1437,150,1293]" pageId="9" pageNumber="460">
|
||||
Female (
|
||||
<figureCitation id="136553B7FFD8FFE7FEE819D6E7C3FB80" box="[298,387,1015,1041]" captionStart="FIGURE 10" captionStartId="15.[151,250,1944,1968]" captionTargetBox="[455,1133,181,1920]" captionTargetId="figure-18@15.[455,1133,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. A paralectotype, female (incorrectly designated as “the allotype” by Kimmins (1970)) of Lestes decipiens, from Ceylon, 6th Milestone, Nilavelli Road, 9 xii 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818173" httpUri="https://zenodo.org/record/15818173/files/figure.png" pageId="9" pageNumber="460">Figs 10</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFD8FFE7FE5319D6E790FB80" box="[401,464,1015,1041]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">12f–j</figureCitation>
|
||||
). The cerci in dorsal view are finger-like, narrowing only in apical part, pale entirely (
|
||||
<figureCitation id="136553B7FFD8FFE7FF5D1E3AE743FBA4" box="[159,259,1051,1077]" captionStart="FIGURE 10" captionStartId="15.[151,250,1944,1968]" captionTargetBox="[455,1133,181,1920]" captionTargetId="figure-18@15.[455,1133,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. A paralectotype, female (incorrectly designated as “the allotype” by Kimmins (1970)) of Lestes decipiens, from Ceylon, 6th Milestone, Nilavelli Road, 9 xii 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818173" httpUri="https://zenodo.org/record/15818173/files/figure.png" pageId="9" pageNumber="460">Figs 10e</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFD8FFE7FED31E3AE71BFBA4" box="[273,347,1051,1077]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">
|
||||
<quantity id="4CA6E2D7FFD8FFE7FED31E3AE77BFBA4" box="[273,315,1051,1077]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="9" pageNumber="460" unit="g" value="12.0">12g</quantity>
|
||||
–h
|
||||
</figureCitation>
|
||||
). The ovipositor is but very slightly concave in lateral view (
|
||||
<figureCitation id="136553B7FFD8FFE7FBD31E3AE230FBA4" box="[1041,1136,1051,1077]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">Fig. 12f</figureCitation>
|
||||
); blackish, with pale areas at base (
|
||||
<figureCitation id="136553B7FFD8FFE7FF311E1EE714FBC8" box="[243,340,1087,1113]" captionStart="FIGURE 10" captionStartId="15.[151,250,1944,1968]" captionTargetBox="[455,1133,181,1920]" captionTargetId="figure-18@15.[455,1133,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. A paralectotype, female (incorrectly designated as “the allotype” by Kimmins (1970)) of Lestes decipiens, from Ceylon, 6th Milestone, Nilavelli Road, 9 xii 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818173" httpUri="https://zenodo.org/record/15818173/files/figure.png" pageId="9" pageNumber="460">Fig. 10d</figureCitation>
|
||||
,
|
||||
<figureCitation id="136553B7FFD8FFE7FEA21E1EE7E4FBC8" box="[352,420,1087,1113]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">12f, h</figureCitation>
|
||||
). The genital lamina is blackish-brown at base and pale at posterior margin (
|
||||
<figureCitation id="136553B7FFD8FFE7FB3E1E1EE31DFBC8" box="[1276,1373,1087,1113]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">Fig. 12h</figureCitation>
|
||||
). The dorsal side of S10 black in the studied specimens (
|
||||
<figureCitation id="136553B7FFD8FFE7FD041E42E566FBEC" box="[710,806,1123,1149]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">
|
||||
Fig.
|
||||
<quantity id="4CA6E2D7FFD8FFE7FD3B1E42E566FBEC" box="[761,806,1123,1149]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="9" pageNumber="460" unit="g" value="12.0">12g</quantity>
|
||||
</figureCitation>
|
||||
), dark-brown in the female
|
||||
<typeStatus id="54E5F190FFD8FFE7FB9B1E42E2B0FBEC" box="[1113,1264,1123,1149]" pageId="9" pageNumber="460" type="paralectotype">paralectotype</typeStatus>
|
||||
(
|
||||
<figureCitation id="136553B7FFD8FFE7FB3D1E42E31EFBEC" box="[1279,1374,1123,1149]" captionStart="FIGURE 10" captionStartId="15.[151,250,1944,1968]" captionTargetBox="[455,1133,181,1920]" captionTargetId="figure-18@15.[455,1133,181,1920]" captionTargetPageId="15" captionText="FIGURE 10. A paralectotype, female (incorrectly designated as “the allotype” by Kimmins (1970)) of Lestes decipiens, from Ceylon, 6th Milestone, Nilavelli Road, 9 xii 1891, Yerbury: a–general habitus in lateral view; b–ditto in dorsal view; c–labels, d–end of abdomen in lateral view; e–ditto in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15818173" httpUri="https://zenodo.org/record/15818173/files/figure.png" pageId="9" pageNumber="460">Fig. 10e</figureCitation>
|
||||
). The prothorax hind lobe is almost upright, very scarcely slanting in lateral view (
|
||||
<figureCitation id="136553B7FFD8FFE7FC3A1EA6E212FB30" box="[1016,1106,1159,1185]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">Fig. 12i</figureCitation>
|
||||
) and looks rounded in dorsal view (
|
||||
<figureCitation id="136553B7FFD8FFE7FF1F1E8AE778FB54" box="[221,312,1195,1221]" captionStart="FIGURE 12" captionStartId="19.[151,250,1897,1921]" captionTargetBox="[263,1324,237,1860]" captionTargetId="figure-48@19.[263,1324,237,1874]" captionTargetPageId="19" captionText="FIGURE 12. The end of abdomen (a–c, f–h) and thorax (d–e, i–j) of female specimens of Lestes dorothea (from Thailand, Kanchanaburi Province, Kroeng Krawia) (a–e) and L. decipiens (from Vietnam, Quảng Nam Province, Nui Thanh, Tam My Tay; species rank restored in this paper) (f–j) collected in tandem with males: a, d, f, i–lateral view; b, e, g, j–dorsal view, c–h–ventral view; d mirrored." figureDoi="http://doi.org/10.5281/zenodo.15818178" httpUri="https://zenodo.org/record/15818178/files/figure.png" pageId="9" pageNumber="460">Fig. 12i</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE14F32FFD8FFE7FF051EEEE31AFA9C" blockId="9.[151,1437,150,1293]" pageId="9" pageNumber="460">
|
||||
Measurements (mm). Hindwing 20–21 (mostly 20), 20.0 in the female
|
||||
<typeStatus id="54E5F190FFD8FFE7FC111EEEE22DFB78" box="[979,1133,1231,1257]" pageId="9" pageNumber="460" type="paralectotype">paralectotype</typeStatus>
|
||||
, abdomen with appendages
|
||||
<quantity id="4CA6E2D7FFD8FFE7FF551ED2E6A5FA9C" box="[151,229,1267,1293]" metricMagnitude="-1" metricUnit="m" metricValue="7.0612" pageId="9" pageNumber="460" unit="in" value="27.8">27.8 in</quantity>
|
||||
the
|
||||
<typeStatus id="54E5F190FFD8FFE7FED71ED2E732FA9C" box="[277,370,1267,1293]" pageId="9" pageNumber="460" type="allotype">allotype</typeStatus>
|
||||
, 26.5 and
|
||||
<quantity id="4CA6E2D7FFD8FFE7FE271ED2E474FA9C" box="[485,564,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="460" unit="mm" value="30.0">30 mm</quantity>
|
||||
in Vietnamese specimens, 29–30 (mostly 30) in Cambodian specimens.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
1266
data/03/F7/FE/03F7FE24FFD9FFFFFF551CB5E458F9C0.xml
Normal file
1266
data/03/F7/FE/03F7FE24FFD9FFFFFF551CB5E458F9C0.xml
Normal file
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