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<document ID-DOI="http://dx.doi.org/10.3897/vz.73.e101372" ID-Pensoft-Pub="2625-8498-73-717" ID-Pensoft-UUID="318B657A15AB5708B3C35FC1A82B4945" ID-ZooBank="8F3D5EDA2F184E5CA53E2F7741FF1339" ModsDocID="2625-8498-73-717" checkinTime="1695893367890" checkinUser="pensoft" docAuthor="Szyndlar, Zbigniew &amp; Georgalis, Georgios L." docDate="2023" docId="ECD7035EB4A8B24DE84A465999861AC6" docLanguage="en" docName="VertZool 73: 717-886" docOrigin="Vertebrate Zoology 73" docPubDate="2023-09-27" docSource="http://dx.doi.org/10.3897/vz.73.e101372" docTitle="Brachyophidium Wall 1921" docType="treatment" docVersion="1" id="318B657A15AB5708B3C35FC1A82B4945" lastPageNumber="717" masterDocId="318B657A15AB5708B3C35FC1A82B4945" masterDocTitle="An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column" masterLastPageNumber="886" masterPageNumber="717" pageNumber="717" updateTime="1695893367890" updateUser="pensoft">
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<mods:title id="127735C5C520C7FA29AEFB7D7C7A2A0C">An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column</mods:title>
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<mods:namePart id="9ACA1B47B751BD33BB1A8BA6DFAE63D6">Szyndlar, Zbigniew</mods:namePart>
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<mods:namePart id="031A61E5CFE934D2ED4FFB922910D69C">Georgalis, Georgios L.</mods:namePart>
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<mods:title id="FDC79F334B05BBBB9C005F443EE41793">Vertebrate Zoology</mods:title>
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<mods:date id="230004374616EAD2DCE413919C5387CF">2023</mods:date>
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<mods:number id="91F0937CAEC9F6CB80966FD4E337BC8C">73</mods:number>
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<taxonomicName id="254B0CE53D58D29622B0E1B4DD297BC7" authority="Wall, 1921" authorityName="Wall" authorityYear="1921" class="Squamata" family="Uropeltidae" genus="Brachyophidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyophidium" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="genus">Brachyophidium Wall, 1921</taxonomicName>
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<subSubSection id="19D2280929FB90325F092659FC976E02" pageId="0" pageNumber="717" type="materials_examined">
<paragraph id="5176CE29D6DCC3A89C115105C777E77F" pageId="0" pageNumber="717">Material examined.</paragraph>
<paragraph id="1FBCBE2FDF16DE0DCAA546CCBD1ED8C0" pageId="0" pageNumber="717">
<taxonomicName id="2A420344DDC1DEC580318ECE83D2815F" authorityName="Wall" authorityYear="1921" class="Squamata" family="Uropeltidae" genus="Brachyophidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyophidium rhodogaster" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="DE39F634F6B8151BB4A53A31D353513D" italics="true" pageId="0" pageNumber="717">Brachyophidium rhodogaster</emphasis>
</taxonomicName>
Wall, 1921 (MCZ Herp R-18073).
</paragraph>
</subSubSection>
<subSubSection id="238E91C2F8A0D98AB31786B677425F13" pageId="0" pageNumber="717" type="description">
<paragraph id="8D22711A821B5CB1303981B37CEAA1FD" pageId="0" pageNumber="717">Description (Figs 43-44).</paragraph>
<paragraph id="291B1FECAB2F03F709444584D860EAB0" pageId="0" pageNumber="717">
<emphasis id="3468DB385B3966BA563A8DE7FFAD62C0" bold="true" pageId="0" pageNumber="717">
<emphasis id="A8A257AF10568488C8D45D8859FFCE94" italics="true" pageId="0" pageNumber="717">Trunk vertebrae</emphasis>
.
</emphasis>
Centrum elongate; neural arch depressed; posterior median notch of the neural arch absent or very shallow; neural spine vestigial and restricted to the posteriormost part of the neural arch, disappearing in mid- and posterior trunk vertebrae; high-angled prezygapophyses; prezygapophyseal accessory processes moderate in length; zygosphene narrow with strongly concave margins in dorsal view; hypapophyses spine-like, disappearing at the level of V 40 to V 50; moderately developed flattened haemal keel appears on posterior trunk vertebrae; paracotylar foramina absent; an asymmetrical subcentral foramen can be occasionally present.
</paragraph>
<caption id="C3EB8E51DAB98238AFD6110C5A7E1E69" doi="10.3897/vz.73.e101372.figure43" httpUri="https://binary.pensoft.net/fig/912933" pageId="0" pageNumber="717" start="Figure 43" startId="F43">
<paragraph id="EE11367E0B13CA18DB4747B002E1EB6C" pageId="0" pageNumber="717">
<emphasis id="40CEC2C864472BDE59F003DAC4932CCE" bold="true" pageId="0" pageNumber="717">Figure 43.</emphasis>
<taxonomicName id="44CC2AF4E7897E93F4A4BE35B520C9E0" class="Reptilia" family="Uropeltidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="family">Uropeltidae</taxonomicName>
:
<taxonomicName id="6C2E321C1FEABBD7F86D4B65AAA7B516" authorityName="Wall" authorityYear="1921" class="Squamata" family="Uropeltidae" genus="Brachyophidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyophidium rhodogaster" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="A41C0FA58F0AE45067A912C265EF37D9" italics="true" pageId="0" pageNumber="717">Brachyophidium rhodogaster</emphasis>
</taxonomicName>
(MCZ Herp R-18073), trunk vertebrae.
</paragraph>
</caption>
<caption id="6F68C29E356854523B787109A2769400" doi="10.3897/vz.73.e101372.figure44" httpUri="https://binary.pensoft.net/fig/912934" pageId="0" pageNumber="717" start="Figure 44" startId="F44">
<paragraph id="5378654262DCB964AF3CF028075424FB" pageId="0" pageNumber="717">
<emphasis id="0D8CD43FC2392DD13B577D976EC80CA0" bold="true" pageId="0" pageNumber="717">Figure 44.</emphasis>
<taxonomicName id="3DA82920B161F862BECDAAA50C33FB69" class="Reptilia" family="Uropeltidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="family">Uropeltidae</taxonomicName>
:
<taxonomicName id="283D735405FB8CC8130130486907AAB5" authorityName="Wall" authorityYear="1921" class="Squamata" family="Uropeltidae" genus="Brachyophidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyophidium rhodogaster" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="42076A37521B9D27037E1BDA75ECF817" italics="true" pageId="0" pageNumber="717">Brachyophidium rhodogaster</emphasis>
</taxonomicName>
(MCZ Herp R-18073), posterior trunk, cloacal, and caudal vertebrae.
</paragraph>
</caption>
<paragraph id="769342D0E7CF90A69E760B20D243817C" pageId="0" pageNumber="717">
<emphasis id="E050146113F71E7531A39B928C8E7038" bold="true" pageId="0" pageNumber="717">
<emphasis id="8ED809C76FCB98FF7393313959B60989" italics="true" pageId="0" pageNumber="717">Trunk</emphasis>
/
<emphasis id="323D5E2732CED74D9BEFC4C9D87A0801" italics="true" pageId="0" pageNumber="717">caudal transition</emphasis>
</emphasis>
. The haemal keel becomes progressively larger in the last trunk to anterior cloacal portion of the column, eventually developing into a prominent hypapophysis in posterior cloacal and all following caudal vertebrae.
</paragraph>
<paragraph id="83D33AD72017F4ED1BCB0CF7D70ABE5A" pageId="0" pageNumber="717">
<emphasis id="4AC8C5062162D48A7B38A82EA7DC54D4" bold="true" pageId="0" pageNumber="717">
<emphasis id="F810C0D2975AD65F752A435E8B75FF5B" italics="true" pageId="0" pageNumber="717">Number of vertebrae</emphasis>
.
</emphasis>
<taxonomicName id="7B1D2EC084ABE637DD4AEDF4AA16C854" authorityName="Wall" authorityYear="1921" class="Squamata" family="Uropeltidae" genus="Brachyophidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyophidium rhodogaster" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="7A82BBBA35E81965F5228E0959AFDE53" italics="true" pageId="0" pageNumber="717">Brachyophidium rhodogaster</emphasis>
</taxonomicName>
(MCZ Herp R-18073):?145 (?133+3+9+; a few anterior trunk vertebrae lacking).
</paragraph>
<paragraph id="5341F4DA77CD2E8F565FBB251322BAAC" pageId="0" pageNumber="717">
Data from literature:
<taxonomicName id="2A3B0767E56E840FE0037490E944EFB7" authorityName="Wall" authorityYear="1921" class="Squamata" family="Uropeltidae" genus="Brachyophidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachyophidium rhodogaster" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="AB9BEB84DCC1F22DE508E1C6907F82FC" italics="true" pageId="0" pageNumber="717">Brachyophidium rhodogaster</emphasis>
</taxonomicName>
: 138-143 trunk and cloacal vertebrae plus 9-12 caudal vertebrae (
<bibRefCitation id="B2989A28BF67443F6051E4D50C99FB3F" author="Alexander, AA" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="717" pagination="171 - 190" refId="B10" refString="Alexander, AA, Gans, C, 1966. The pattern of dermal-vertebral correlation in snakes and amphisbaenians. Zoologische Mededelingen 31: 171 - 190" title="The pattern of dermal-vertebral correlation in snakes and amphisbaenians." volume="31" year="1966">Alexander and Gans 1966</bibRefCitation>
).
</paragraph>
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1066.69697" ID-GBIF-Dataset="89be094e-6732-48a6-b5aa-74053d16f748" ID-GBIF-Taxon="189599982" ID-Pensoft-Pub="1313-2970-1066-1" ID-Pensoft-UUID="BD84F4C3157550C9B64120B2BE53F01A" ID-ZooBank="133CE040A5AF44F1BC9A558C2F06A8AA" ModsDocID="1313-2970-1066-1" checkinTime="1635480455043" checkinUser="pensoft" docAuthor="Filander, Zoleka N., Kitahara, Marcelo V., Cairns, Stephen D., Sink, Kerry J. &amp; Lombard, Amanda T." docDate="2021" docId="ECD71DB1B92AFF9AB42EDEFF6E392A53" docLanguage="en" docName="ZooKeys 1066: 1-198" docOrigin="ZooKeys 1066" docPubDate="2021-10-28" docSource="http://dx.doi.org/10.3897/zookeys.1066.69697" docTitle="Trochocyathus (Trochocyathus) Milne-Edwards &amp; Haime 1848" docType="treatment" docVersion="3" id="BD84F4C3157550C9B64120B2BE53F01A" lastPageNumber="1" masterDocId="BD84F4C3157550C9B64120B2BE53F01A" masterDocTitle="Azooxanthellate Scleractinia (Cnidaria, Anthozoa) from South Africa" masterLastPageNumber="198" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1635513010010" updateUser="ExternalLinkService">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Filander, Zoleka N.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6905-4440</mods:nameIdentifier>
<mods:affiliation>Biodiversity and Coastal Research, Oceans and Coasts, Department of Environment, Forestry, and Fisheries, Cape Town, South Africa &amp; Zoology Department, Nelson Mandela University, Port Elizabeth, South Africa</mods:affiliation>
<mods:nameIdentifier type="email">zfilander@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kitahara, Marcelo V.</mods:namePart>
<mods:affiliation>Universidade Federal de Sao Paulo, Departamento de Ciencias do Mar, Santos, Brazil &amp; Centro de Biologia Marinha, Universidade de Sao Paulo, Sao Sebastiao, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cairns, Stephen D.</mods:namePart>
<mods:affiliation>Department of Invertebrate Zoology, Smithsonian Institution, Washington DC, USA</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sink, Kerry J.</mods:namePart>
<mods:affiliation>South African National Biodiversity Institute, Cape Town, South Africa &amp; Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lombard, Amanda T.</mods:namePart>
<mods:affiliation>Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth, South Africa</mods:affiliation>
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<mods:date>2021</mods:date>
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<mods:number>2021-10-28</mods:number>
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<mods:number>1066</mods:number>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ZooBank">133CE040A5AF44F1BC9A558C2F06A8AA</mods:identifier>
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<treatment ID-GBIF-Taxon="189599982" LSID="urn:lsid:plazi:treatment:ECD71DB1B92AFF9AB42EDEFF6E392A53" httpUri="http://treatment.plazi.org/id/ECD71DB1B92AFF9AB42EDEFF6E392A53" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName authority="(Trochocyathus) Milne-Edwards &amp; Haime, 1848" authorityName="Milne-Edwards &amp; Haime" authorityYear="1848" baseAuthorityName="Trochocyathus" class="Anthozoa" family="Caryophylliidae" genus="Trochocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trochocyathus (Trochocyathus)" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="subGenus" subGenus="Trochocyathus">Trochocyathus (Trochocyathus) Milne-Edwards &amp; Haime, 1848</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Milne-Edwards &amp; Haime" authorityYear="1848" class="Anthozoa" family="Turbinoliidae" genus="Trochocyathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trochocyathus" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Trochocyathus</emphasis>
</taxonomicName>
lacking basal costal spines and with other than discoidal coralla.
</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Anthozoa" family="Deltocyathidae" genus="Turbinolia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turbinolia mitrata" order="Scleractinia" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="mitrata">
<emphasis italics="true" pageId="0" pageNumber="1">Turbinolia mitrata</emphasis>
</taxonomicName>
Goldfuss, 1827, by subsequent designation (
<bibRefCitation author="Milne-Edwards, H" journalOrPublisher="Palaeontological Society, London" pageId="0" pageNumber="1" refId="B189" refString="Milne-Edwards, H, Haime, J, 1850b. A monograph on the British fossil Corals. Palaeontological Society, London" title="A monograph on the British fossil Corals." year="1850 b">Milne-Edwards and Haime 1850b</bibRefCitation>
).
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</treatment>
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</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>88</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>972</mods:end>
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<treatment ID-GBIF-Taxon="159364198" LSID="urn:lsid:plazi:treatment:ECD79690304B395D60BC158486E1B4EE" httpUri="http://treatment.plazi.org/id/ECD79690304B395D60BC158486E1B4EE" lastPageNumber="217" pageId="216" pageNumber="217">
<subSubSection pageId="216" pageNumber="217" type="nomenclature">
<paragraph pageId="216" pageNumber="217">
Tribe
<taxonomicName authority="Thomson, 1860" authorityName="Thomson" authorityYear="1860" lsidName="" pageId="216" pageNumber="217" rank="tribe" tribe="Tetraopini">Tetraopini Thomson, 1860</taxonomicName>
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</subSubSection>
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<paragraph pageId="216" pageNumber="217">
<taxonomicName genus="Tetraopesitae" lsidName="Tetraopesitae" pageId="216" pageNumber="217" rank="genus">Tetraopesitae</taxonomicName>
J. Thomson, 1860a: 66 [stem: Tetraop-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Tetraopes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetraopes" order="Coleoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="genus">Tetraopes</taxonomicName>
<normalizedToken originalValue="Schönherr">Schoenherr</normalizedToken>
, 1817. Comment: incorrect original stem formation, not in prevailing usage.
</paragraph>
</subSubSection>
</treatment>
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@ -0,0 +1,381 @@
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<taxonomicName LSID="urn:lsid:ipni.org:names:177582-1" authority="Bieb. 1808" authorityYear="1808" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anthemis ruthenica" order="Asterales" pageId="0" pageNumber="1018" phylum="Tracheophyta" rank="species" species="ruthenica">Anthemis ruthenica Bieb. 1808</taxonomicName>
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<subSubSection pageId="0" pageNumber="1018" type="materials_examined">
<paragraph pageId="0" pageNumber="1018">Materials</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="1996-6-7" collectionCode="FRU" collectorName="M. Pimenov, E. Kluykov, G. Lazkov" country="Kyrgyzstan" latitude="42.6733" location="Northern side of Kyrgyz Ala-Too, near Belogorka [Tosh-Bulak] Village" longitude="74.2415" pageId="0" pageNumber="1018" specimenCode="1" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">1</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">M.Pimenov, E.Kluykov, G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:9C3982A35841C4BC3631CDC6A7D589B8" country="Kyrgyzstan" latitude="42.6733" longitude="74.2415" name="Northern side of Kyrgyz Ala-Too, near Belogorka [Tosh-Bulak] Village" pageId="0" pageNumber="1018">Northern side of Kyrgyz Ala-Too, near Belogorka [Tosh-Bulak] Village</location>
; decimalLatitude:
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; decimalLongitude:
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; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="1996-6-7">06/07/1996</collectingDate>
; year: 1996; month: 6; day: 7; fieldNumber: 29; eventRemarks: alien plant; Record Level: collectionID: 88420; institutionCode:
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</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="1996-6-7" collectionCode="LE" collectorName="M. Pimenov, E. Kluykov, G. Lazkov" country="Kyrgyzstan" latitude="42.6733" location="Northern side of Kyrgyz Ala-Too, near Belogorka [Tosh-Bulak] Village" longitude="74.2415" pageId="0" pageNumber="1018" specimenCode="2" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">2</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">M.Pimenov, E.Kluykov, G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:94BC95E74701FE194544E787191C9C7D" country="Kyrgyzstan" latitude="42.6733" longitude="74.2415" name="Northern side of Kyrgyz Ala-Too, near Belogorka [Tosh-Bulak] Village" pageId="0" pageNumber="1018">Northern side of Kyrgyz Ala-Too, near Belogorka [Tosh-Bulak] Village</location>
; decimalLatitude:
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; decimalLongitude:
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; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="1996-6-7">06/07/1996</collectingDate>
; year: 1996; month: 6; day: 7; fieldNumber: 29; eventRemarks: alien plant; Record Level: collectionID: 90565; institutionCode:
<collectionCode pageId="0" pageNumber="1018">LE</collectionCode>
; basisOfRecord: specimen
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="1965-7-21" collectionCode="FRU" collectorName="anonym" country="Kyrgyzstan" latitude="42.7493" location="Northern side of Kyrgyz Ala-Too, near Cholok Village" longitude="73.5036" pageId="0" pageNumber="1018" specimenCode="3" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">3</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">anonym</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:B995B6388D48FA7D0091250D813A1EFA" country="Kyrgyzstan" latitude="42.7493" longitude="73.5036" name="Northern side of Kyrgyz Ala-Too, near Cholok Village" pageId="0" pageNumber="1018">Northern side of Kyrgyz Ala-Too, near Cholok Village</location>
; decimalLatitude:
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; decimalLongitude:
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; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="1965-7-21">07/21/1965</collectingDate>
; year: 1965; month: 7; day: 21; fieldNumber: 1281; eventRemarks: alien plant; Record Level: collectionID: 88420; institutionCode:
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; basisOfRecord: specimen
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="1970-5-14" collectionCode="FRU" collectorName="Sudnitsyna" country="Kazakhstan" latitude="42.8744" location="Vicinity of Jambyl [Taraz] Town, foothills" longitude="71.4376" pageId="0" pageNumber="1018" specimenCode="4" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
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; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kazakhstan</collectingCountry>
; stateProvince: Taraz Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:5FF2383B472A27D53525C0587FA316C2" country="Kazakhstan" latitude="42.8744" longitude="71.4376" name="Vicinity of Jambyl [Taraz] Town, foothills" pageId="0" pageNumber="1018">Vicinity of Jambyl [Taraz] Town, foothills</location>
; decimalLatitude:
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; decimalLongitude:
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; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="1970-5-14">05/14/1970</collectingDate>
; year: 1970; month: 5; day: 14; habitat: clayey slopes, semidesert; eventRemarks: alien plant; Record Level: collectionID: 88420; institutionCode:
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; basisOfRecord: specimen
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="1968-6-2" collectionCode="FRU" collectorName="R. Sultanova" country="Kyrgyzstan" latitude="42.7297" location="Pokrovka Village" longitude="71.607" pageId="0" pageNumber="1018" specimenCode="5" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1018">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">5</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">R.Sultanova</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince: Talas Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:013FC70871ACB855E4F4AEBD07E51914" country="Kyrgyzstan" latitude="42.7297" longitude="71.607" name="Pokrovka Village" pageId="0" pageNumber="1018">Pokrovka Village</location>
; decimalLatitude:
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; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="1018" value="71.607">71.607</geoCoordinate>
; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="1968-6-2">06/02/1968</collectingDate>
; year: 1968; month: 6; day: 2; habitat: roadside; eventRemarks: alien plant; Record Level: collectionID: 88420; institutionCode:
<collectionCode pageId="0" pageNumber="1018">FRU</collectionCode>
; basisOfRecord: specimen
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="05/01/1998" collectionCode="FRU" collectorName="G. Lazkov" country="Kyrgyzstan" latitude="42.6739" location="Ala-Archa River, Kashka-Suu Village" longitude="74.5275" pageId="0" pageNumber="1018" specimenCode="6" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1018">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">6</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
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; decimalLatitude:
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; decimalLongitude:
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; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
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; basisOfRecord: specimen
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="1990-5-18" collectionCode="FRU" collectorName="S. Sheremetova &amp; G. Lazkov" country="Kyrgyzstan" latitude="42.7055" location="right side of Aspara River, 4 km upstream Granitogorsk Town" longitude="73.458" pageId="0" pageNumber="1018" specimenCode="7" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
<collectorName pageId="0" pageNumber="1018">S.Sheremetova &amp; G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:861042110FEDA377CD13AD77172E3334" country="Kyrgyzstan" latitude="42.7055" longitude="73.458" name="right side of Aspara River, 4 km upstream Granitogorsk Town" pageId="0" pageNumber="1018">right side of Aspara River, 4 km upstream Granitogorsk Town</location>
; decimalLatitude:
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; decimalLongitude:
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; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="1990-5-18">05/18/1990</collectingDate>
; year: 1990; month: 5; day: 18; habitat: gravelly slope along the road; eventRemarks: alien plant; Record Level: collectionID: 88420; institutionCode:
<collectionCode pageId="0" pageNumber="1018">FRU</collectionCode>
; basisOfRecord: specimen
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="1978-6-17" collectionCode="FRU" collectorName="E. S. Poliakova" country="Kyrgyzstan" latitude="42.6529" location="Kara-Balta River, Taldy-Bulak" longitude="73.9064" pageId="0" pageNumber="1018" specimenCode="8" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
<collectorName pageId="0" pageNumber="1018">E.S.Poliakova</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
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; decimalLatitude:
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; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="1018" value="73.9064">73.9064</geoCoordinate>
; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="1978-6-17">06/17/1978</collectingDate>
; year: 1978; month: 6; day: 17; habitat: by the road; eventRemarks: alien plant; Record Level: collectionID: 88420; institutionCode:
<collectionCode pageId="0" pageNumber="1018">FRU</collectionCode>
; basisOfRecord: specimen
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="01/06/2010-01/09/2010" collectorName="G. Lazkov" country="Kyrgyzstan" latitude="42.8694" location="Kant Town" longitude="74.8419" pageId="0" pageNumber="1018" specimenCode="9" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1018">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">9</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:16D8536679C25D6769175170C424DB5C" country="Kyrgyzstan" latitude="42.8694" longitude="74.8419" name="Kant Town" pageId="0" pageNumber="1018">Kant Town</location>
; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="1018" value="42.8694">42.8694</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="1018" value="74.8419">74.8419</geoCoordinate>
; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="01/06/2010-01/09/2010">01/06/2010-01/09/2010</collectingDate>
; year: 2010; eventRemarks: alien plant; Record Level: basisOfRecord: observation
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="01/06/2010-01/09/2010" collectorName="G. Lazkov" country="Kyrgyzstan" latitude="42.8105" location="Tokmok Town" longitude="75.2758" pageId="0" pageNumber="1018" specimenCode="10" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1018">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">10</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:9BFDCF7FC9377324DAF7A10E3E4507E0" country="Kyrgyzstan" latitude="42.8105" longitude="75.2758" name="Tokmok Town" pageId="0" pageNumber="1018">Tokmok Town</location>
; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="1018" value="42.8105">42.8105</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="1018" value="75.2758">75.2758</geoCoordinate>
; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="01/06/2010-01/09/2010">01/06/2010-01/09/2010</collectingDate>
; year: 2010; eventRemarks: alien plant; Record Level: basisOfRecord: observation
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="01/06/2010-01/09/2010" collectorName="G. Lazkov" country="Kyrgyzstan" latitude="42.7595" location="Kemin Town" longitude="75.6875" pageId="0" pageNumber="1018" specimenCode="11" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1018">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">11</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:B659488F34494B2D7F8399F828A490C9" country="Kyrgyzstan" latitude="42.7595" longitude="75.6875" name="Kemin Town" pageId="0" pageNumber="1018">Kemin Town</location>
; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="1018" value="42.7595">42.7595</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="1018" value="75.6875">75.6875</geoCoordinate>
; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 01/01/1998; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="01/06/2010-01/09/2010">01/06/2010-01/09/2010</collectingDate>
; year: 2010; eventRemarks: alien plant; Record Level: basisOfRecord: observation
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1018">
<materialsCitation collectingDate="2013-6-22" collectionCode="FRU" collectorName="G. Lazkov" country="Kyrgyzstan" latitude="42.798332" location="E of Kara-Balta Town, factory area (under construction)" longitude="73.88861" pageId="0" pageNumber="1018" specimenCode="12" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1018">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1018">12</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1018">G.Lazkov</collectorName>
; Taxon: family: Asteraceae; genus: Anthemis; specificEpithet: ruthenica; taxonRank: species; scientificNameAuthorship: Bieb.; Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="1018">Kyrgyzstan</collectingCountry>
; stateProvince:
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region; locality:
<location LSID="urn:lsid:plazi:treatment:ECD7ABE43CD3FC2A16D9529A98B818AD:AD2CCA9F40685EF3843C1066FD6AA338" country="Kyrgyzstan" latitude="42.798332" longitude="73.88861" name="E of Kara-Balta Town, factory area (under construction)" pageId="0" pageNumber="1018">E of Kara-Balta Town, factory area (under construction)</location>
; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="1018" value="42.798333">42.798333</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="1018" value="73.888611">73.888611</geoCoordinate>
; Identification: identifiedBy: G.A.Lazkov; dateIdentified: 22/06/2013; Event: eventDate:
<collectingDate pageId="0" pageNumber="1018" value="2013-6-22">22/06/2013</collectingDate>
; year: 2013; month: 6; day: 22; habitat: ruderal places; eventRemarks: alien plant; Record Level: collectionID: 88420; institutionCode:
<collectionCode pageId="0" pageNumber="1018">FRU</collectionCode>
; basisOfRecord: specimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1018" type="native distribution and occurrence in central asia">
<paragraph pageId="0" pageNumber="1018">Native distribution and occurrence in Central Asia</paragraph>
<paragraph pageId="0" pageNumber="1018">
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Anthemis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anthemis ruthenica" order="Asterales" pageId="0" pageNumber="1018" phylum="Tracheophyta" rank="species" species="ruthenica">Anthemis ruthenica</taxonomicName>
is an annual or biennial species native to southeastern Europe and the Caucasus (
<bibRefCitation author="Tzvelev, N. N." editor="Tzvelev, N. N." journalOrPublisher="Science Publishers, St. Petersburg" pageId="0" pageNumber="1018" title="Flora of the European part of the USSR" year="1994">Tzvelev 1994</bibRefCitation>
). There are very few published records of this alien species from Central Asia. Earlier it was known from Magtymguly Village (Kopetdagh Range, Turkmenistan), western parts of Talas and Kyrgyz Ranges in Kazakhstan, and along Shakhimardan River in Uzbekistan (
<bibRefCitation author="Kamelin, R. V." editor="Adylov, T. A." journalOrPublisher="Academy of Sciences of the Uzbek SSR, Tashkent" pageId="0" pageNumber="1018" title="Conspectus florae Asiae Mediae" year="1993">Kamelin and Kovalevskaya 1993</bibRefCitation>
). The localities in Kazakhstan are very close to the border of Kyrgyzstan, and this proximity is confirmed by a specimen at FRU that was collected at the distance of ca 10 km (vicinity of Jambyl [Taraz] Town, foothills, clayey slopes, semidesert, 14.05.1970, I.Sudnitsyna). The locality in Uzbekistan is situated in the enclave surrounded by the territory of Kyrgyzstan. The presence of this species in Kyrgyzstan was therefore expected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1018" type="occurrence in kyrgyzstan">
<paragraph pageId="0" pageNumber="1018">Occurrence in Kyrgyzstan</paragraph>
<paragraph pageId="0" pageNumber="1018">
The species was first reported from Kyrgyzstan on the basis of a single recent collection from Tosh-Bulak [formerly Belogorka] village on the northern side of Kyrgyz Range (
<bibRefCitation author="Lazkov, G. A." journalOrPublisher="Botanical Journal (St. Petersburg)" pageId="0" pageNumber="1018" pagination="125 - 126" title="A new species and new records in Asteraceae from Kyrgyzstan" volume="84" year="1999">Lazkov 1999</bibRefCitation>
). A further revision of collections at FRU revealed a number of earlier specimens of the species collected along the foothills of the northern side of Kyrgyz Range between the border with Kazakhstan and Kara-Balta Town. The species has been most frequently collected at the country border. Besides, a single herbarium specimen was collected in the Talas Depression, also very close to the border with Kazakhstan. These records make an extension of the previously known distribution area.
</paragraph>
<paragraph pageId="0" pageNumber="1018">
The species is also known from the eastern part of the
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Depression. It was observed by G.Lazkov (without voucher specimens) near Kant Town, Tokmok Town, and Kemin Town of
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Region.
</paragraph>
<paragraph pageId="0" pageNumber="1018">
At present, in Kyrgyzstan
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Anthemis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anthemis ruthenica" order="Asterales" pageId="0" pageNumber="1018" phylum="Tracheophyta" rank="species" species="ruthenica">Anthemis ruthenica</taxonomicName>
is widespread and locally abundant in the
<normalizedToken originalValue="Chüy">Chuey</normalizedToken>
Depression and the western part of the Talas Depression (including foothills), completely covering the lowlands of northern Kyrgyzstan (Fig. 10). It typically occurs in wastelands, on roadsides and dry riversides. The occurrence of this species in Kyrgyzstan is not new but has been very poorly known; previously it was misidentified in collections of FRU as
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Pyrethrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pyrethrum transiliense" order="Asterales" pageId="0" pageNumber="1018" phylum="Tracheophyta" rank="species" species="transiliense">Pyrethrum transiliense</taxonomicName>
(Herd.) Regel &amp; Schmalh. The species has a clear tendency to get established in new territories and may potentially spread much further. Our latest observation comes from the vicinity of Kara-Balta Town from which the species was first recorded as late as 1978; vigorous stands were observed in the new factory area (Fig. 1). At present the species is truly abundant in disturbed lands of various kind around the town (Fig. 2).
</paragraph>
<paragraph pageId="0" pageNumber="1018">Since most of the older records originated from the territories neighbouring with Kazakhstan, we conclude that the species was imported from that country in mid-Soviet times. The oldest record documented by a specimen collected in 1965 came indeed from Cholok-Aryk Village at the very border with Kazakhstan.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1018" type="invasion status in kyrgyzstan">
<paragraph pageId="0" pageNumber="1018">Invasion status in Kyrgyzstan</paragraph>
<paragraph pageId="0" pageNumber="1018">Established alien, naturalised (self-sustaining with established populations of a high number of individuals) in human-made and disturbed habitats at the regional scale. Spreading and invasive, locally replacing other species.</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.85584" ID-Pensoft-Pub="1860-1324-2-139" ID-Pensoft-UUID="03EFF5CBC2CF53CFA27D350B268657C3" ID-ZooBank="73B92B049B454237B81497A21DB59468" ModsDocID="1860-1324-69-2-139" checkinTime="1658439162963" checkinUser="pensoft" docAuthor="Kobor, Peter" docDate="2022" docId="ECD7F4512F9C5F27ABFD5798BAEAB4C4" docLanguage="en" docName="DeutEntomolZeit 69(2): 139-150" docOrigin="Deutsche Entomologische Zeitschrift 69 (2)" docPubDate="2022-07-21" docSource="http://dx.doi.org/10.3897/dez.69.85584" docTitle="Umbrageocoris malipatili Kóbor 2022, sp. nov." docType="treatment" docUuid="A5568832-84E7-4502-8741-97398F57F841" docUuidSource="ZooBank" docVersion="1" id="03EFF5CBC2CF53CFA27D350B268657C3" lastPageNumber="139" masterDocId="03EFF5CBC2CF53CFA27D350B268657C3" masterDocTitle="A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae)" masterLastPageNumber="150" masterPageNumber="139" pageNumber="139" updateTime="1658439162963" updateUser="pensoft">
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<mods:title>A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kobor, Peter</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0266-7303</mods:nameIdentifier>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Eoetvoes Lorand Research Network, 15 Herman Otto street, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">kobor.peter@atk.hu</mods:nameIdentifier>
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<mods:number>2022-07-21</mods:number>
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<mods:number>69</mods:number>
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<mods:number>2</mods:number>
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<mods:start>139</mods:start>
<mods:end>150</mods:end>
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<treatment LSID="urn:lsid:zoobank.org:act:A5568832-84E7-4502-8741-97398F57F841" httpUri="http://treatment.plazi.org/id/ECD7F4512F9C5F27ABFD5798BAEAB4C4" lastPageNumber="139" pageId="0" pageNumber="139">
<subSubSection pageId="0" pageNumber="139" type="nomenclature">
<paragraph pageId="0" pageNumber="139">
<taxonomicName LSID="https://zoobank.org/A5568832-84E7-4502-8741-97398F57F841" authority="Kóbor, 2022" authorityName="Kóbor" authorityYear="2022" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris malipatili" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="malipatili" status="sp. nov.">Umbrageocoris malipatili</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="139">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="description">
<paragraph pageId="0" pageNumber="139">
<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Exoskeletal characters of Umbrageocoris: 1 - 2. Labial trough (indicated by arrow) of U. kondorosyi (1) and U. malipatili sp. nov. (2), 3 - 4. Metathoracic wing (arrow indicates hamus) of U. kondorosyi (3) and U. elegantulus (4) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures1-4" httpUri="https://binary.pensoft.net/fig/718124" pageId="0" pageNumber="139">Figs 2</figureCitation>
<figureCitation captionStart="Figures 510" captionStartId="F2" captionText="Figures 5 - 10. Exoskeletal characters of Umbrageocoris: peritreme of metathoracic scent efferent apparatus of U. kondorosyi (5), U. maai maai (6), U. boonei sp. nov. (7), U. malipatili sp. nov. (8), U. elegantulus (9), and U. woodwardi (10) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures5-10" httpUri="https://binary.pensoft.net/fig/718125" pageId="0" pageNumber="139">, 7</figureCitation>
<figureCitation captionStart="Figures 1924" captionStartId="F5" captionText="Figures 19 - 24. Dorsal habitus of Umbrageocoris species: 19. U. kondorosyi (holotype, HNHM), 20. U. maai maai (holotype, BPBM), 21. U. maai timorensis (holotype, BPBM), 22. U. boonei sp. nov. (holotype, BPBM), 23. U. malipatili sp. nov. (holotype, MNHN), 24. U. woodwardi (SEMC) (scale bar = 1 mm for all images)." figureDoi="10.3897/dez.69.85584.figures19-24" httpUri="https://binary.pensoft.net/fig/718128" pageId="0" pageNumber="139">, 23</figureCitation>
<figureCitation captionStart="Figures 2728" captionStartId="F7" captionText="Figures 27 - 28. Distribution map of Umbrageocoris species: 27. Indomalayan species (U. maai ssp. and U. boonei sp. nov.), 28. Australasian species (U. elegantulus, U. kondorosyi, U. malipatili sp. nov. and U. woodwardi)." figureDoi="10.3897/dez.69.85584.figures27-28" httpUri="https://binary.pensoft.net/fig/718130" pageId="0" pageNumber="139">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="materials_examined">
<paragraph pageId="0" pageNumber="139">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="139">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="GALLES" location="GALLES" specimenCount="1" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">
<typeStatus>Holotype</typeStatus>
</emphasis>
, f (MNHN): &quot;MUSEUM PARIS / NOUV.
<collectingCountry name="United Kingdom">GALLES</collectingCountry>
DU SUD / COLL. NOUALHIER 1898&quot;.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="description">
<paragraph pageId="0" pageNumber="139">Description.</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Colouration</emphasis>
: Head ochraceous, posterior part of eye stalks with irregular brownish marking. Eyes reddish, ocelli hyaline. Antennae missing in the single examined specimen. Labiomeres dark ochraceous. Thorax. Pronotum ochraceous with brownish punctation and an infuscate transversal band across pronotal callosities. Scutellum entirely dark brownish. Hemelytra semi-hyaline, ochraceous with brownish punctation and an irregular infuscate spot on the corium. Thoracic pleurites and sternites dark brownish except prosternal collar, supracoxal lobes and peritreme; propleurite with an ochraceous spot dorsally. Abdomen dark brownish both dorsally and ventrally with a whitish longitudinal band at dorsal part of sternites.
</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Structure</emphasis>
: Head. Median furrow of vertex short, hardly surpassing base of clypeus. Antennomeres missing. Labial trough Y-shaped with suture reduced, reaching approximately midline of head. Relative lengths of labiomeres: 1.00: 0.85: 1.32: 1:38. Thorax. Pronotum with dense and deep punctation except anterior margin, pronotal callosities and posterior margin with humeral angles. Pronotal callosities confluent. Pronotum length to basal width: 1.00: 1.76. Scutellum with trifurcate carina apically reduced. Hemelytra submacropterous. Scutellum length to basal width: 1.00: 1.06. Clavus with an incomplete row of punctures at corial margin. Corium punctate along margin of clavus, Cu, between M-R and costa in apical half. Exocorium slightly widened subapically. Membrane short, not surpassing apex of abdomen.
</paragraph>
<caption doi="10.3897/dez.69.85584.figures11-14" httpUri="https://binary.pensoft.net/fig/718126" pageId="0" pageNumber="139" start="Figures 1114" startId="F3">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figures 11-14.</emphasis>
Exoskeletal characters of
<taxonomicName authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="139">Umbrageocoris</emphasis>
</taxonomicName>
: male pygophore with parameres
<emphasis italics="true" pageId="0" pageNumber="139">in situ</emphasis>
(
<emphasis bold="true" pageId="0" pageNumber="139">11</emphasis>
), female ovipositor
<emphasis italics="true" pageId="0" pageNumber="139">in situ</emphasis>
(
<emphasis bold="true" pageId="0" pageNumber="139">12</emphasis>
), arrangement of abdominal trichobothria on sternite IV (
<emphasis bold="true" pageId="0" pageNumber="139">13</emphasis>
) and sternites V-VII (
<emphasis bold="true" pageId="0" pageNumber="139">14</emphasis>
)
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">
<emphasis italics="true" pageId="0" pageNumber="139">Measurements</emphasis>
.
</emphasis>
Body length: 4.20; head length: 0.53; head width: 1.66; lengths of labiomeres I-IV: 0.36-0.30-0.47-0.49; pronotum length: 0.88; pronotum width: 1.54; scutellum length: 0.88; scutellum width: 0.83.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="diagnosis">
<paragraph pageId="0" pageNumber="139">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName class="Insecta" genus="U." kingdom="Animalia" lsidName="U. malipatili" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="malipatili">
<emphasis italics="true" pageId="0" pageNumber="139">U. malipatili</emphasis>
</taxonomicName>
sp. nov. resembles
<taxonomicName class="Insecta" genus="U." kingdom="Animalia" lsidName="U. boonei" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="boonei">
<emphasis italics="true" pageId="0" pageNumber="139">U. boonei</emphasis>
</taxonomicName>
sp. nov. and both subspecies of
<taxonomicName class="Insecta" genus="U." kingdom="Animalia" lsidName="U. maai" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="maai">
<emphasis italics="true" pageId="0" pageNumber="139">U. maai</emphasis>
</taxonomicName>
in colouration and markings, but its cup-shaped labial trough and the presence of a minute furrow on its clypeus readily distinguishes it from the Indomalayan representatives of the genus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="distribution">
<paragraph pageId="0" pageNumber="139">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="139">The single known specimen is from an unknown locality in New South Wales, Australia.</paragraph>
</subSubSection>
</treatment>
</document>