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<document id="B030113C57C3CDF095450D4F160C77AF" ID-CLB-Dataset="46019" ID-DOI="10.11646/zootaxa.3786.2.3" ID-GBIF-Dataset="172de54d-d273-4a3f-a866-5fe601fbef4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="224463" ID-ZooBank="975E11EF-B1CE-4FF2-9159-91F1AB474D60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460408216353" checkinUser="plazi" docAuthor="Promdam, Rueangrit, Nabhitabhata, Jaruwat &amp; Galil, Bella S." docDate="2014" docId="EC1AD51D8A3D2F32FF16FF37FE23FD3E" docLanguage="en" docName="zt03786p140.pdf" docOrigin="Zootaxa 3786 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleusia huilianae Promdam, Nabhitabhata &amp; Galil, 2014, n. sp." docType="treatment" docVersion="9" lastPageNumber="140" masterDocId="1023AD658A3C2F37FF81FFA0FF9EFFBB" masterDocTitle="A new species of Coleusia Galil, 2006 (Decapoda: Brachyura: Leucosiidae) from southern Asia" masterLastPageNumber="140" masterPageNumber="135" pageNumber="136" updateTime="1698300095810" updateUser="ExternalLinkService">
<mods:mods id="700E30E250FDF92ECC9D2B68AF55471E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="693F369FD687FBA4522AA261C27BFC8A">
<mods:title id="F56B05A624C037C419131847D0EE9ACC">A new species of Coleusia Galil, 2006 (Decapoda: Brachyura: Leucosiidae) from southern Asia</mods:title>
</mods:titleInfo>
<mods:name id="878AC808EF982B5A4E4B0B6F1D78ABD2" type="personal">
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<mods:namePart id="C9501AB27E04F05BD6BBE1DDA6FB63EA">Promdam, Rueangrit</mods:namePart>
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<mods:namePart id="F8741C0CABEB5DEA0728A64A3C927349">Nabhitabhata, Jaruwat</mods:namePart>
</mods:name>
<mods:name id="68FC22503B48240067BDB0D79E925E45" type="personal">
<mods:role id="903B785E1263A356B2FE66519AF81F8E">
<mods:roleTerm id="894C1D61EA216CCC629DFB68B8550ADE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="64A4086B6F83A220F3D1DAC3FDF7FC05">Galil, Bella S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D5007A7D0737CDD2E04643B79ED7E9CC">text</mods:typeOfResource>
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<mods:title id="098831B362541F0C05F2C1C28F70A1B2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="355714EEE712AB49FAAEB6772F558AE1">
<mods:date id="F4FC383DC19EDE969C4FC6C64DA9685D">2014</mods:date>
<mods:detail id="519A5E121D24FDF3CC1900F8D6DE7F8A" type="volume">
<mods:number id="6806CE43825045C64D857F79EDD24F20">3786</mods:number>
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<mods:start id="A6969B49FAB45E0A0D3E4F449C04961D">135</mods:start>
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<mods:classification id="F18B514B2439563AE8421AFD85145733">journal article</mods:classification>
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<mods:identifier id="E69C1012A9F9C6E1CB825A1D5D941836" type="DOI">10.11646/zootaxa.3786.2.3</mods:identifier>
<mods:identifier id="69C8B16E5B75ABBC969AEBCD158DA752" type="GBIF-Dataset">172de54d-d273-4a3f-a866-5fe601fbef4c</mods:identifier>
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</mods:mods>
<treatment id="EC1AD51D8A3D2F32FF16FF37FE23FD3E" ID-DOI="http://doi.org/10.5281/zenodo.6130028" ID-GBIF-Taxon="119422491" ID-Zenodo-Dep="6130028" LSID="urn:lsid:plazi:treatment:EC1AD51D8A3D2F32FF16FF37FE23FD3E" httpUri="http://treatment.plazi.org/id/EC1AD51D8A3D2F32FF16FF37FE23FD3E" lastPageId="5" lastPageNumber="140" pageId="1" pageNumber="136">
<subSubSection id="2CA937808A3D2F36FF16FF37FE8CFF68" pageId="1" pageNumber="136" type="nomenclature">
<paragraph id="640C640B8A3D2F36FF16FF37FE5BFF09" blockId="1.[151,453,151,211]" box="[151,453,151,178]" pageId="1" pageNumber="136">
<heading id="3F44D3678A3D2F36FF16FF37FE5BFF09" bold="true" box="[151,453,151,178]" fontSize="11" level="1" pageId="1" pageNumber="136" reason="1">
<emphasis id="56C7B8198A3D2F36FF16FF37FE5BFF09" bold="true" box="[151,453,151,178]" pageId="1" pageNumber="136">
<taxonomicName id="A3B31F888A3D2F36FF16FF37FEE5FF0A" ID-CoL="X2C6" box="[151,379,151,177]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A3D2F36FF16FF37FEE5FF0A" bold="true" box="[151,379,151,177]" italics="true" pageId="1" pageNumber="136">Coleusia huilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DF405628A3D2F36FE02FF38FE5BFF09" box="[387,453,152,178]" pageId="1" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="640C640B8A3D2F36FF16FF1AFE8CFF68" blockId="1.[151,453,151,211]" box="[151,274,186,211]" pageId="1" pageNumber="136">
(
<figureCitation id="FC88788E8A3D2F36FF1EFF1AFE97FF68" box="[159,265,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1885,1907]" captionStartId-1="2.[151,250,1601,1623]" captionStartId-2="3.[151,250,1969,1991]" captionStartId-3="4.[151,250,1228,1250]" captionTargetBox-0="[322,1264,1070,1852]" captionTargetBox-1="[151,1436,193,1579]" captionTargetBox-2="[189,1393,429,1942]" captionTargetBox-3="[259,1317,196,1197]" captionTargetId-0="figure@1.[322,1264,1070,1853]" captionTargetId-1="figure@2.[151,1436,193,1579]" captionTargetId-2="figure@3.[189,1394,417,1942]" captionTargetId-3="figure@4.[191,1403,157,1217]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Coleusia huilianae n. sp., paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094). Dorsal view of carapace, colours in life." captionText-1="FIGURE 2. Coleusia huilianae n. sp., Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514 - 333 - 0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax." captionText-2="FIGURE 3. Left male first gonopod of Coleusia huilianae n. sp., paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094). A, dorsal view; B, ventral view." captionText-3="FIGURE 4. Left male first gonopod of Coleusia huilianae n. sp., paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094), setae denuded. Scales: A = 3.0 mm; B, C = 1.0 mm." httpUri-0="https://zenodo.org/record/224464/files/figure.png" httpUri-1="https://zenodo.org/record/224465/files/figure.png" httpUri-2="https://zenodo.org/record/224466/files/figure.png" httpUri-3="https://zenodo.org/record/224467/files/figure.png" pageId="1" pageNumber="136">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2CA937808A3D2F36FF16FEA3FD84FECF" pageId="1" pageNumber="136" type="reference_group">
<paragraph id="640C640B8A3D2F36FF16FEA3FC86FE8C" blockId="1.[151,1435,258,372]" pageId="1" pageNumber="136">
<treatmentCitationGroup id="44A343258A3D2F36FF16FEA3FC86FE8C" pageId="1" pageNumber="136">
<taxonomicName id="A3B31F888A3D2F36FF16FEA3FE61FEA3" ID-CoL="84FXW" authority="Alcock 1896: 220" authorityName="Alcock" authorityPageNumber="220" authorityYear="1896" box="[151,511,258,280]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A3D2F36FF16FEA3FEA2FEA3" box="[151,316,259,280]" italics="true" pageId="1" pageNumber="136">Leucosia urania</emphasis>
,
<treatmentCitation id="E512421A8A3D2F36FEC9FEA2FE61FEA3" author="Alcock" box="[328,511,258,280]" page="220" pageId="1" pageNumber="136" year="1896">
<bibRefCitation id="002219FA8A3D2F36FEC9FEA2FE50FEA3" author="Alcock" box="[328,462,258,280]" pageId="1" pageNumber="136" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2. The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, Calcutta, 65 (2), 134 - 296." type="journal article" year="1896">Alcock 1896</bibRefCitation>
: 220
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="E512421A8A3D2F36FD9CFEA2FD2FFEA3" author="Ihle" box="[541,689,258,280]" page="316" pageId="1" pageNumber="136" year="1918">
<bibRefCitation id="002219FA8A3D2F36FD9CFEA2FD1FFEA3" author="Ihle" box="[541,641,258,280]" pageId="1" pageNumber="136" refString="Ihle, J. E. W. (1918) Die Decapoda Brachyura der Siboga-Expedition. III. Oxystomata: Calappidae, Leucosiidae, Raninidae. Siboga Expeditie, 39 (B 2), 159 - 322." type="journal article" year="1918">Ihle 1918</bibRefCitation>
: 316
</treatmentCitation>
(part) (not
<taxonomicName id="A3B31F888A3D2F36FCA4FEA3FBD8FEA3" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[805,1094,258,280]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A3D2F36FCA4FEA3FC24FEA3" box="[805,954,259,280]" italics="true" pageId="1" pageNumber="136">Cancer urania</emphasis>
Herbst, 1801
</taxonomicName>
).
<taxonomicName id="A3B31F888A3D2F36FF16FE82FE6CFE8C" authority="Chen &amp; Turkay" authorityName="Chen &amp; Turkay" box="[151,498,289,311]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="56C7B8198A3D2F36FF16FE82FEDBFE8C" box="[151,325,289,311]" italics="true" pageId="1" pageNumber="136">Leucosia grandis</emphasis>
, Chen &amp; Türkay
</taxonomicName>
, in
<bibRefCitation id="002219FA8A3D2F36FD97FE81FD4DFE8C" author="Chen" box="[534,723,289,311]" pageId="1" pageNumber="136" refString="Chen, H. &amp; Sun, H. (2002) Arthropoda Crustacea. Brachyura. Marine primitive crabs. Fauna Sinica. Invertebrata, 30, Science Press, Beijing, 597 pp." type="book" year="2002">Chen &amp; Sun, 2002</bibRefCitation>
(part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="640C640B8A3D2F36FF16FEE1FD84FECF" blockId="1.[151,1435,258,372]" pageId="1" pageNumber="136">
<treatmentCitationGroup id="44A343258A3D2F36FF16FEE1FD84FECF" pageId="1" pageNumber="136">
Not
<taxonomicName id="A3B31F888A3D2F36FF42FEE0FEE3FEED" box="[195,381,320,342]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A3D2F36FF42FEE0FEE3FEED" box="[195,381,320,342]" italics="true" pageId="1" pageNumber="136">Leucosides urania</emphasis>
</taxonomicName>
or
<taxonomicName id="A3B31F888A3D2F36FE21FEE1FC89FEED" ID-CoL="84FXW" authority="Rathbun 1910: 309" authorityName="Rathbun" authorityPageNumber="309" authorityYear="1910" box="[416,791,320,342]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A3D2F36FE21FEE1FDD8FEED" box="[416,582,321,342]" italics="true" pageId="1" pageNumber="136">Leucosia urania</emphasis>
,
<treatmentCitation id="E512421A8A3D2F36FDD2FEE0FC89FEED" author="Rathbun" box="[595,791,320,342]" page="309" pageId="1" pageNumber="136" year="1910">
<bibRefCitation id="002219FA8A3D2F36FDD2FEE0FD7BFEED" author="Rathbun" box="[595,741,320,342]" pageId="1" pageNumber="136" refString="Rathbun, M. J. (1910) Brachyura. The Danish Expedition to Siam 1899 - 1900, V. Det Kongelige Danske Videnskabernes Selskabs Skrifter, 7. Raekke, Naturvidenskabelig og Mathematisk Afdeling, 4, 301 - 367." type="journal article" year="1910">Rathbun 1910</bibRefCitation>
: 309
</treatmentCitation>
</taxonomicName>
, pl. 1, fig. 1.—
<treatmentCitation id="E512421A8A3D2F36FC2AFEE0FBEBFEED" author="Naiyanetr" box="[939,1141,320,342]" page="58" pageId="1" pageNumber="136" year="1998">
<bibRefCitation id="002219FA8A3D2F36FC2AFEE0FBCDFEED" author="Naiyanetr" box="[939,1107,320,342]" pageId="1" pageNumber="136" refString="Naiyanetr, P. (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, 161 pp." type="book" year="1998">Naiyanetr 1998</bibRefCitation>
: 58
</treatmentCitation>
(in list); 2007: 69 (in list) (=
<taxonomicName id="A3B31F888A3D2F36FF46FEFEFD88FECF" ID-CoL="4XYP4" authority="Miers, 1877" authorityName="Miers" authorityYear="1877" box="[199,534,350,372]" class="Malacostraca" family="Leucosiidae" genus="Soceulia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="56C7B8198A3D2F36FF46FEFEFEE6FECF" box="[199,376,350,372]" italics="true" pageId="1" pageNumber="136">Soceulia brunnea</emphasis>
(Miers, 1877))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2CA937808A3D2F36FF16FE01FD40FD61" pageId="1" pageNumber="136" type="materials_examined">
<paragraph id="640C640B8A3D2F36FF16FE01FAA4FDD5" blockId="1.[151,1436,417,1018]" pageId="1" pageNumber="136">
<emphasis id="56C7B8198A3D2F36FF16FE01FEE7FE01" bold="true" box="[151,377,417,442]" pageId="1" pageNumber="136">Material examined</emphasis>
.
<emphasis id="56C7B8198A3D2F36FE06FE01FD51FE01" bold="true" box="[391,719,417,442]" pageId="1" pageNumber="136">
<collectingCountry id="1CA4249B8A3D2F36FE06FE01FE6FFE01" box="[391,497,417,442]" name="Thailand" pageId="1" pageNumber="136">
<emphasis id="56C7B8198A3D2F36FE06FE01FE6FFE01" bold="true" box="[391,497,417,442]" italics="true" pageId="1" pageNumber="136">Thailand</emphasis>
</collectingCountry>
(
<emphasis id="56C7B8198A3D2F36FD83FE01FD51FE01" bold="true" box="[514,719,417,442]" italics="true" pageId="1" pageNumber="136">
Gulf of
<collectingCountry id="1CA4249B8A3D2F36FDDDFE01FD58FE01" box="[604,710,417,442]" name="Thailand" pageId="1" pageNumber="136">Thailand</collectingCountry>
)
</emphasis>
</emphasis>
.
<typeStatus id="BB08DAA98A3D2F36FD5CFE01FCD7FE01" box="[733,841,417,442]" pageId="1" pageNumber="136" type="holotype">Holotype</typeStatus>
. Ko Raet, Sattahip, Chon Buri Province,
<date id="100D42CB8A3D2F36FA96FE01FA09FE01" box="[1303,1431,417,442]" pageId="1" pageNumber="136" value="2012-05-18">18.05.2012</date>
, leg. P. Wisespongpand,
<specimenCount id="72B5AF828A3D2F36FE20FE66FE55FE66" box="[417,459,454,478]" pageId="1" pageNumber="136" type="generic">1 ♂</specimenCount>
(
<quantity id="A34BC9EE8A3D2F36FE5BFE65FDDFFE65" box="[474,577,453,478]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="1" pageNumber="136" unit="mm" value="35.3">35.3 mm</quantity>
) (PSUNHM 211514-333-0041).
<typeStatus id="BB08DAA98A3D2F36FC32FE66FBBAFE65" box="[947,1060,454,478]" pageId="1" pageNumber="136" type="paratype">Paratypes</typeStatus>
. Ko Phai, Ko Rin and Ko Khram, Artisanal fishery of Ko Lan, Chon Buri Province, bottom gill net,
<date id="100D42CB8A3D2F36FC0EFE49FB8EFDBA" box="[911,1040,489,513]" pageId="1" pageNumber="136" value="2012-10-19">19.10.2012</date>
, leg. P. Wisespongpand,
<specimenCount id="72B5AF828A3D2F36FAACFE49FAC7FDBA" box="[1325,1369,489,513]" pageId="1" pageNumber="136" type="generic">1 ♂</specimenCount>
(
<quantity id="A34BC9EE8A3D2F36FAEBFE49FF5AFD9D" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="1" pageNumber="136" unit="mm" value="38.5">38.5 mm</quantity>
) (KUMF-ART0012); Pattani Province,
<date id="100D42CB8A3D2F36FD02FDADFC9AFD9D" box="[643,772,525,550]" pageId="1" pageNumber="136" value="1984-07-25">25.07.1984</date>
, leg. Bencha Mapha,
<typeStatus id="BB08DAA98A3D2F36FC77FDAEFBC8FD9D" box="[1014,1110,526,550]" pageId="1" pageNumber="136" type="paratype">paratype</typeStatus>
of
<taxonomicName id="A3B31F888A3D2F36FBFAFDAEFF29FDF1" authority="Chen &amp; Tu" authorityName="Chen &amp; Tu" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="56C7B8198A3D2F36FBFAFDAEFAA3FD9E" box="[1147,1341,525,549]" italics="true" pageId="1" pageNumber="136">Leucosia grandis</emphasis>
Chen &amp; Tu
</taxonomicName>
?rkay,
<specimenCount id="72B5AF828A3D2F36FE81FD91FEB2FDF2" box="[256,300,561,585]" pageId="1" pageNumber="136" type="generic">1 ♂</specimenCount>
(
<quantity id="A34BC9EE8A3D2F36FEBDFD91FE3AFDF2" box="[316,420,561,585]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="1" pageNumber="136" unit="mm" value="37.2">37.2 mm</quantity>
) (
<collectionCode id="02A2FCCE8A3D2F36FE3AFD91FE6DFDF2" box="[443,499,561,585]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="136">ZRC</collectionCode>
2000.0094); Pattani fishing port,
<date id="100D42CB8A3D2F36FCF5FD91FC6AFDF1" box="[884,1012,561,586]" pageId="1" pageNumber="136" value="1998-10-25">25.10.1998</date>
, leg. H.H. Tan,
<specimenCount id="72B5AF828A3D2F36FB28FD91FB4AFDF2" box="[1193,1236,561,585]" pageId="1" pageNumber="136" type="generic">1 ♂</specimenCount>
(
<quantity id="A34BC9EE8A3D2F36FB65FD91FAD2FDF2" box="[1252,1356,561,585]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="1" pageNumber="136" unit="mm" value="37.2">37.2 mm</quantity>
) (
<collectionCode id="02A2FCCE8A3D2F36FAE5FD91FA02FDF2" box="[1380,1436,561,585]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="136">ZRC</collectionCode>
1999.0304); 0 6.2011, leg. P.K.L from aquarium dealer in Bangkok. Ng,
<specimenCount id="72B5AF828A3D2F36FC3FFDF6FC77FDD6" box="[958,1001,598,622]" pageId="1" pageNumber="136" type="generic">1 ♂</specimenCount>
(
<quantity id="A34BC9EE8A3D2F36FC76FDF6FBC1FDD5" box="[1015,1119,598,622]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="1" pageNumber="136" unit="mm" value="37.2">37.2 mm</quantity>
) (
<collectionCode id="02A2FCCE8A3D2F36FBF5FDF6FB32FDD5" box="[1140,1196,598,622]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="136">ZRC</collectionCode>
2011.0662).
</paragraph>
<paragraph id="640C640B8A3D2F36FF46FDD9FCDFFD0D" blockId="1.[151,1436,417,1018]" pageId="1" pageNumber="136">
<emphasis id="56C7B8198A3D2F36FF46FDD9FE2AFD29" bold="true" box="[199,436,633,658]" pageId="1" pageNumber="136">Additional material</emphasis>
:
<collectingCountry id="1CA4249B8A3D2F36FE45FDD9FD9CFD29" box="[452,514,633,658]" name="India" pageId="1" pageNumber="136">
<emphasis id="56C7B8198A3D2F36FE45FDD9FD9CFD29" bold="true" box="[452,514,633,658]" italics="true" pageId="1" pageNumber="136">India</emphasis>
</collectingCountry>
. Dhanushkodi, Rameshwaram, discards on shore,
<date id="100D42CB8A3D2F36FBBEFDDAFB5EFD29" box="[1087,1216,634,658]" pageId="1" pageNumber="136" value="2009-07-31">31.07.2009</date>
, leg. V.S. Kakati,
<specimenCount id="72B5AF828A3D2F36FA0FFDDAFF32FD0E" pageId="1" pageNumber="136" type="generic">1 ♂</specimenCount>
(
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) (
<collectionCode id="02A2FCCE8A3D2F36FEB6FD3EFE0EFD0D" box="[311,400,670,694]" httpUri="http://grbio.org/cool/mve2-984h" name="See FMRI" pageId="1" pageNumber="136">CMFRI</collectionCode>
<collectionCode id="02A2FCCE8A3D2F36FE16FD3DFE21FD0D" box="[407,447,669,694]" pageId="1" pageNumber="136">ED</collectionCode>
.5.6.4.1), 1♀ (
<quantity id="A34BC9EE8A3D2F36FDDBFD3EFD5EFD0D" box="[602,704,670,694]" metricMagnitude="-2" metricUnit="m" metricValue="3.42" pageId="1" pageNumber="136" unit="mm" value="34.2">34.2 mm</quantity>
) (
<collectionCode id="02A2FCCE8A3D2F36FD56FD3EFCABFD0D" box="[727,821,670,694]" httpUri="http://grbio.org/cool/mve2-984h" name="See FMRI" pageId="1" pageNumber="136">CMFRI</collectionCode>
).
</paragraph>
<paragraph id="640C640B8A3D2F36FF16FD61FD40FD61" blockId="1.[151,1436,417,1018]" box="[151,734,705,730]" pageId="1" pageNumber="136">
<bibRef id="BD13F0FD8A3D2F36FF16FD61FD40FD61" box="[151,734,705,730]" pageId="1" pageNumber="136" type="book chapter">
<title id="16B78F3D8A3D2F36FF16FD61FE93FD61" box="[151,269,705,730]" pageId="1" pageNumber="136">
<collectingCountry id="1CA4249B8A3D2F36FF16FD61FE93FD61" box="[151,269,705,730]" name="Singapore" pageId="1" pageNumber="136">
<emphasis id="56C7B8198A3D2F36FF16FD61FE93FD61" bold="true" box="[151,269,705,730]" italics="true" pageId="1" pageNumber="136">Singapore</emphasis>
</collectingCountry>
</title>
.
<year id="101B25588A3D2F36FE9AFD62FEC8FD61" box="[283,342,706,730]" pageId="1" pageNumber="136">1983</year>
,
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<pagination id="5D7A9AFF8A3D2F36FEE0FD62FEF1FD61" box="[353,367,706,730]" pageId="1" pageNumber="136" type="pagination">1</pagination>
♂
</specimenCount>
<journalOrPublisher id="BE726EC78A3D2F36FE13FD61FD44FD61" box="[402,730,705,730]" pageId="1" pageNumber="136">
(
<quantity id="A34BC9EE8A3D2F36FE1BFD61FD9FFD61" box="[410,513,705,730]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="1" pageNumber="136" unit="mm" value="35.5">35.5 mm</quantity>
) (
<collectionCode id="02A2FCCE8A3D2F36FD96FD62FDD1FD61" box="[535,591,706,730]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="136">ZRC</collectionCode>
1988.2237)
</journalOrPublisher>
.
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="2CA937808A3D2F33FF46FD45FBBFF915" lastPageId="4" lastPageNumber="139" pageId="1" pageNumber="136" type="description">
<paragraph id="640C640B8A3D2F36FF46FD45FD6FFC41" blockId="1.[151,1436,417,1018]" pageId="1" pageNumber="136">
<emphasis id="56C7B8198A3D2F36FF46FD45FECFFD45" bold="true" box="[199,337,741,766]" pageId="1" pageNumber="136">Description</emphasis>
. Carapace (
<figureCitation id="FC88788E8A3D2F36FE56FD45FDBBFD45" box="[471,549,741,766]" captionStart="FIGURE 1" captionStartId="1.[151,250,1885,1907]" captionTargetBox="[322,1264,1070,1852]" captionTargetId="figure@1.[322,1264,1070,1853]" captionTargetPageId="1" captionText="FIGURE 1. Coleusia huilianae n. sp., paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/224464/files/figure.png" pageId="1" pageNumber="136">Figs. 1</figureCitation>
,
<figureCitation id="FC88788E8A3D2F36FDB2FD46FDDCFD45" box="[563,578,742,766]" captionStart="FIGURE 2" captionStartId="2.[151,250,1601,1623]" captionTargetBox="[151,1436,193,1579]" captionTargetId="figure@2.[151,1436,193,1579]" captionTargetPageId="2" captionText="FIGURE 2. Coleusia huilianae n. sp., Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514 - 333 - 0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/224465/files/figure.png" pageId="1" pageNumber="136">2</figureCitation>
) subpentagonal, globose, 1.1 times as long as wide; dorsal surface glabrous, smooth, sparsely punctate; regions indistinct. Frontal region produced, upcurved, deeply concave medially, laterally constricted; frontal margin rounded. Anterolateral surface slightly inflated anteriorly, margin sinuous, with obsolescent granules distally. Lateral angle of carapace rounded, overhanging thoracic sinus, margin prominently beaded, granules lozenge-shaped. Posterolateral margins of carapace rounded, smooth. Epimeral margin invisible in dorsal view, gutter-like anteriorly, narrowing posteriorly, closely beaded marginal granules decreasing in size posteriorly, meeting posterior margin at an angle. Posterior margin sinuous in male, slightly rounded in female, minutely granular; deflexed posterior surface smooth.
</paragraph>
<caption id="30CC34838A3D2F36FF16F8FDFDB2F829" httpUri="https://zenodo.org/record/224464/files/figure.png" pageId="1" pageNumber="136" targetBox="[322,1264,1070,1852]" targetPageId="1">
<paragraph id="640C640B8A3D2F36FF16F8FDFDB2F829" blockId="1.[151,1436,1885,1938]" pageId="1" pageNumber="136">
<emphasis id="56C7B8198A3D2F36FF16F8FDFE91F8C8" bold="true" box="[151,271,1885,1907]" pageId="1" pageNumber="136">FIGURE 1</emphasis>
.
<taxonomicName id="A3B31F888A3D2F36FEA1F8FDFE7AF8C8" box="[288,484,1885,1907]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A3D2F36FEA1F8FDFE7AF8C8" box="[288,484,1885,1907]" italics="true" pageId="1" pageNumber="136">Coleusia huilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DF405628A3D2F36FE6FF8FEFDB4F8C8" box="[494,554,1886,1907]" pageId="1" pageNumber="136" rank="species">
<emphasis id="56C7B8198A3D2F36FE6FF8FEFDB4F8C8" bold="true" box="[494,554,1886,1907]" pageId="1" pageNumber="136">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094). Dorsal view of carapace, colours in life.
</paragraph>
</caption>
<caption id="30CC34838A3E2F35FF16F9E1FC6EF92F" httpUri="https://zenodo.org/record/224465/files/figure.png" pageId="2" pageNumber="137" targetBox="[151,1436,193,1579]" targetPageId="2">
<paragraph id="640C640B8A3E2F35FF16F9E1FC6EF92F" blockId="2.[151,1435,1601,1684]" pageId="2" pageNumber="137">
<emphasis id="56C7B8198A3E2F35FF16F9E1FF64F9EC" bold="true" box="[151,250,1601,1623]" pageId="2" pageNumber="137">FIGURE</emphasis>
2.
<taxonomicName id="A3B31F888A3E2F35FE9CF9E1FE40F9EC" box="[285,478,1601,1623]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A3E2F35FE9CF9E1FE40F9EC" box="[285,478,1601,1623]" italics="true" pageId="2" pageNumber="137">Coleusia huilianae</emphasis>
</taxonomicName>
<emphasis id="56C7B8198A3E2F35FE69F9E1FDBDF9ED" bold="true" box="[488,547,1601,1622]" pageId="2" pageNumber="137">
<taxonomicNameLabel id="4DF405628A3E2F35FE69F9E1FDBDF9ED" box="[488,547,1601,1622]" pageId="2" pageNumber="137" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514- 333-0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax.
</paragraph>
</caption>
<paragraph id="640C640B8A3E2F34FF46F91BFE33FE33" blockId="2.[151,1436,1723,2036]" lastBlockId="3.[151,1436,151,392]" lastPageId="3" lastPageNumber="138" pageId="2" pageNumber="137">
Antennule obliquely folded into fossa, fossa sealed by basal antennular segment. Antennae short, inserted between antennular fossa, orbit. Orbits small, rounded, outer orbital margin unisutured, anterior margin of efferent branchial channel forms part of lower orbital margin. Eyes retractable. External maxillipeds concealing trapezoid buccal opening, smooth; endopod merus triangulate, as long as ischium; in female, endopod with setose fringe lengthwise, ischium medially elevated, keel-like. Thoracic sinus deep, tomentose, anterior margin level with base of coxopodite of third maxillipeds, anteriorly defined by overhanging beaded margin of pterygostomian region; ventral margin with row of oval granules medially, decreasing in size posteriorly (
<figureCitation id="FC88788E8A3E2F35FBDDF833FB3AF817" box="[1116,1188,1939,1964]" captionStart="FIGURE 2" captionStartId="2.[151,250,1601,1623]" captionTargetBox="[151,1436,193,1579]" captionTargetId="figure@2.[151,1436,193,1579]" captionTargetPageId="2" captionText="FIGURE 2. Coleusia huilianae n. sp., Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514 - 333 - 0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/224465/files/figure.png" pageId="2" pageNumber="137">Fig. 2</figureCitation>
B). Thoracic sternites smooth, shiny; sternites 1–4 fused; sternites 4–8 with medially interrupted sutures between them. Abdominal cavity nearly reaching buccal cavity, lateral margins prominently ridged medially, ridge fitting into notch between somites 5, 6. Male abdomen smooth; somite 1 transversely narrow; somite 2 narrow, spindle-shaped; somites 3–6 fused, proximally inflated, furrowed lengthwise; somite 6 separated from proximal segments by suture, with lateral margins evenly curved, narrowing distally, with prominent triangular denticle on proximal quarter; telson triangular, as wide as long (
<figureCitation id="FC88788E8A3F2F34FE4CFEA4FD8FFEA7" box="[461,529,260,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1601,1623]" captionTargetBox="[151,1436,193,1579]" captionTargetId="figure@2.[151,1436,193,1579]" captionTargetPageId="2" captionText="FIGURE 2. Coleusia huilianae n. sp., Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514 - 333 - 0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/224465/files/figure.png" pageId="3" pageNumber="138">Fig. 2</figureCitation>
F). Female abdomen with somites 3–6 fused, greatly enlarged, shield-like, telson triangular. First male gonopod elongated, shaft stout, sinuous, tightly coiled 3 times on its axis, ultimate coil swollen, setose; apical process cornute, digitate, curved externally (
<figureCitation id="FC88788E8A3F2F34FC38FEECFB90FEDF" box="[953,1038,332,357]" captionStart="FIGURE 3" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[189,1393,429,1942]" captionTargetId="figure@3.[189,1394,417,1942]" captionTargetPageId="3" captionText="FIGURE 3. Left male first gonopod of Coleusia huilianae n. sp., paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094). A, dorsal view; B, ventral view." httpUri="https://zenodo.org/record/224466/files/figure.png" pageId="3" pageNumber="138">Figs. 3</figureCitation>
,
<figureCitation id="FC88788E8A3F2F34FBA1FEECFBAFFEDF" box="[1056,1073,332,356]" captionStart="FIGURE 4" captionStartId="4.[151,250,1228,1250]" captionTargetBox="[259,1317,196,1197]" captionTargetId="figure@4.[191,1403,157,1217]" captionTargetPageId="4" captionText="FIGURE 4. Left male first gonopod of Coleusia huilianae n. sp., paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094), setae denuded. Scales: A = 3.0 mm; B, C = 1.0 mm." httpUri="https://zenodo.org/record/224467/files/figure.png" pageId="3" pageNumber="138">4</figureCitation>
). Second male gonopod short, slender, apex scoop-like.
</paragraph>
<caption id="30CC34838A3F2F34FF16F811FD4BF85D" httpUri="https://zenodo.org/record/224466/files/figure.png" pageId="3" pageNumber="138" targetBox="[189,1393,429,1942]" targetPageId="3">
<paragraph id="640C640B8A3F2F34FF16F811FD4BF85D" blockId="3.[151,1436,1969,2022]" pageId="3" pageNumber="138">
<emphasis id="56C7B8198A3F2F34FF16F811FE90F87D" bold="true" box="[151,270,1969,1991]" pageId="3" pageNumber="138">FIGURE 3</emphasis>
. Left male first gonopod of
<taxonomicName id="A3B31F888A3F2F34FDB8F811FD65F87C" box="[569,763,1969,1991]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A3F2F34FDB8F811FD65F87C" box="[569,763,1969,1991]" italics="true" pageId="3" pageNumber="138">Coleusia huilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DF405628A3F2F34FC82F812FCA0F87C" box="[771,830,1970,1991]" pageId="3" pageNumber="138" rank="species">
<emphasis id="56C7B8198A3F2F34FC82F812FCA0F87C" bold="true" box="[771,830,1970,1991]" pageId="3" pageNumber="138">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094). A, dorsal view; B, ventral view.
</paragraph>
</caption>
<caption id="30CC34838A382F33FF16FB6CFC3FFABA" httpUri="https://zenodo.org/record/224467/files/figure.png" pageId="4" pageNumber="139" targetBox="[259,1317,196,1197]" targetPageId="4">
<paragraph id="640C640B8A382F33FF16FB6CFC3FFABA" blockId="4.[151,1436,1228,1281]" pageId="4" pageNumber="139">
<emphasis id="56C7B8198A382F33FF16FB6CFE90FB5A" bold="true" box="[151,270,1228,1250]" pageId="4" pageNumber="139">FIGURE 4</emphasis>
. Left male first gonopod of
<taxonomicName id="A3B31F888A382F33FDB8FB6CFD65FB59" box="[569,763,1228,1250]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A382F33FDB8FB6CFD65FB59" box="[569,763,1228,1250]" italics="true" pageId="4" pageNumber="139">Coleusia huilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DF405628A382F33FC82FB6DFCA0FB59" box="[771,830,1229,1250]" pageId="4" pageNumber="139" rank="species">
<emphasis id="56C7B8198A382F33FC82FB6DFCA0FB59" bold="true" box="[771,830,1229,1250]" pageId="4" pageNumber="139">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094), setae denuded. Scales: A = 3.0 mm; B, C = 1.0 mm.
</paragraph>
</caption>
<paragraph id="640C640B8A382F33FF46FA8DFBBFF915" blockId="4.[151,1436,1325,2034]" pageId="4" pageNumber="139">
Chelipeds subequal, robust, longer in adult males than in females. Cheliped merus trigonal; anterior, posterior margins lined with perliform granules, diminishing in size distally on posterior margin, reaching two-thirds posterior margin (
<figureCitation id="FC88788E8A382F33FEEBFAD5FE2CFA35" box="[362,434,1397,1422]" captionStart="FIGURE 2" captionStartId="2.[151,250,1601,1623]" captionTargetBox="[151,1436,193,1579]" captionTargetId="figure@2.[151,1436,193,1579]" captionTargetPageId="2" captionText="FIGURE 2. Coleusia huilianae n. sp., Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514 - 333 - 0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/224465/files/figure.png" pageId="4" pageNumber="139">Fig. 2</figureCitation>
C); upper surface with coalesced cluster of granules proximally, followed by 4 large perliform granules, distally smooth; lower surface pitted, granular proximally, few small granules proximally on posterior margin. Carpus inflated, smooth. Propodus lenticular, slightly longer as wide, upper margin rounded, smooth; lower margin with row of minute granules, lower external surface minutely punctuate (
<figureCitation id="FC88788E8A382F33FB3FFA41FA9EFA41" box="[1214,1280,1505,1530]" captionStart="FIGURE 2" captionStartId="2.[151,250,1601,1623]" captionTargetBox="[151,1436,193,1579]" captionTargetId="figure@2.[151,1436,193,1579]" captionTargetPageId="2" captionText="FIGURE 2. Coleusia huilianae n. sp., Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514 - 333 - 0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/224465/files/figure.png" pageId="4" pageNumber="139">Fig. 2</figureCitation>
D). Fingers as long as propodus, gaping, cutting edges, finely teethed. Ambulatory legs similar in shape, gradually decreasing in length from first to fourth. Coxal condyles rounded. Meri subcylindrical; meri 1-4 with row of minute granules on upper margin, obsolescent on anterior merus; curved row of granules on lower margin, last merus with prominently granular crest proximally. Carpi rounded, obsolescent denticle on upper surface. Propodi dorsally, ventrally carinate, expanded, foliaceous (
<figureCitation id="FC88788E8A382F33FE78F935FDA2F915" box="[505,572,1685,1710]" captionStart="FIGURE 2" captionStartId="2.[151,250,1601,1623]" captionTargetBox="[151,1436,193,1579]" captionTargetId="figure@2.[151,1436,193,1579]" captionTargetPageId="2" captionText="FIGURE 2. Coleusia huilianae n. sp., Holotype male (carapace length 35.3 mm, carapace width 32.6) (PSUNHM 211514 - 333 - 0041). A, dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of left merus of cheliped; D, view of lower margin of palm; E, fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/224465/files/figure.png" pageId="4" pageNumber="139">Fig. 2</figureCitation>
E). Dactyli longer than propodi, lanceolate.
</paragraph>
</subSubSection>
<subSubSection id="2CA937808A382F33FF46F919FB6BF94D" pageId="4" pageNumber="139" type="etymology">
<paragraph id="640C640B8A382F33FF46F919FB6BF94D" blockId="4.[151,1436,1325,2034]" pageId="4" pageNumber="139">
<emphasis id="56C7B8198A382F33FF46F919FED8F969" bold="true" box="[199,326,1721,1746]" pageId="4" pageNumber="139">Etymology</emphasis>
. The name honours a fellow carcinologist, the late Chen Hui Lian of the Institute of Oceanology, Chinese Academy of Science in Qingdao,
<collectingCountry id="1CA4249B8A382F33FDF3F97DFD26F94D" box="[626,696,1757,1782]" name="China" pageId="4" pageNumber="139">China</collectingCountry>
, who had published extensively on leucosiid crabs.
</paragraph>
</subSubSection>
<subSubSection id="2CA937808A382F32FF46F8A1FE9EFE16" lastPageId="5" lastPageNumber="140" pageId="4" pageNumber="139" type="discussion">
<paragraph id="640C640B8A382F33FF46F8A1FCF6F849" blockId="4.[151,1436,1325,2034]" pageId="4" pageNumber="139">
<emphasis id="56C7B8198A382F33FF46F8A1FEAAF8A1" bold="true" box="[199,308,1793,1818]" pageId="4" pageNumber="139">Remarks</emphasis>
.
<bibRefCitation id="002219FA8A382F33FEC0F8A1FD85F8A1" author="Alcock" box="[321,539,1793,1818]" pageId="4" pageNumber="139" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2. The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, Calcutta, 65 (2), 134 - 296." type="journal article" year="1896">Alcock (1896: 220)</bibRefCitation>
listed a single female specimen of
<taxonomicName id="A3B31F888A382F33FC20F8A2FBC9F8A2" box="[929,1111,1794,1817]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A382F33FC20F8A2FBC9F8A2" box="[929,1111,1794,1817]" italics="true" pageId="4" pageNumber="139">Leucosia urania</emphasis>
</taxonomicName>
collected in the Andaman Is. Its size (
<quantity id="A34BC9EE8A382F33FF7CF886FEC8F885" box="[253,342,1830,1854]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="139" unit="mm" value="38.0">38 mm</quantity>
), foliaceous meropodites and color pattern fit
<taxonomicName id="A3B31F888A382F33FCF6F886FBCEF885" box="[887,1104,1830,1854]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A382F33FCF6F886FBCEF885" box="[887,1104,1830,1854]" italics="true" pageId="4" pageNumber="139">Coleusia huilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DF405628A382F33FBDAF885FB00F885" box="[1115,1182,1829,1854]" pageId="4" pageNumber="139" rank="species">
<emphasis id="56C7B8198A382F33FBDAF885FB00F885" bold="true" box="[1115,1182,1829,1854]" pageId="4" pageNumber="139">n. sp.</emphasis>
</taxonomicNameLabel>
<bibRefCitation id="002219FA8A382F33FB2BF885FA02F885" author="Rathbun" box="[1194,1436,1829,1854]" pageId="4" pageNumber="139" refString="Rathbun, M. J. (1910) Brachyura. The Danish Expedition to Siam 1899 - 1900, V. Det Kongelige Danske Videnskabernes Selskabs Skrifter, 7. Raekke, Naturvidenskabelig og Mathematisk Afdeling, 4, 301 - 367." type="journal article" year="1910">Rathbun (1910: 309)</bibRefCitation>
identified material collected by the Danish Expedition to
<collectingCountry id="1CA4249B8A382F33FCA0F8E9FCC5F8D9" box="[801,859,1865,1890]" name="Thailand" pageId="4" pageNumber="139">Siam</collectingCountry>
from the islands of Koh Chang, Koh Lan and Koh Kahdat as
<taxonomicName id="A3B31F888A382F33FE95F8CEFE78F83D" box="[276,486,1902,1926]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A382F33FE95F8CEFE78F83D" box="[276,486,1902,1926]" italics="true" pageId="4" pageNumber="139">Leucosides urania</emphasis>
</taxonomicName>
. Upon examination, however, her specimens were identified as
<taxonomicName id="A3B31F888A382F33FB55F8CEFE46F811" authority="(Miers, 1877) (Galil, 2006b)" authorityName="Miers" authorityYear="1877" baseAuthorityName="Miers" baseAuthorityYear="1877" class="Malacostraca" family="Leucosiidae" genus="Soceulia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="56C7B8198A382F33FB55F8CEFA02F83D" box="[1236,1436,1902,1926]" italics="true" pageId="4" pageNumber="139">Soceulia brunnea</emphasis>
(Miers, 1877) (
<bibRefCitation id="002219FA8A382F33FEBFF831FE4FF811" author="Galil" box="[318,465,1937,1962]" pageId="4" pageNumber="139" refString="Galil, B. S. (2006 b) Contributions to the knowledge of Leucosiidae VI. Soceulia gen. nov. (Crustacea: Brachyura). Zoologische Mededelingen, 80, 71 - 79." type="journal article" year="2006" yearSuffix="b">Galil, 2006b</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="002219FA8A382F33FE63F831FD09F811" author="Ihle" box="[482,663,1937,1962]" pageId="4" pageNumber="139" refString="Ihle, J. E. W. (1918) Die Decapoda Brachyura der Siboga-Expedition. III. Oxystomata: Calappidae, Leucosiidae, Raninidae. Siboga Expeditie, 39 (B 2), 159 - 322." type="journal article" year="1918">Ihle (1918: 316)</bibRefCitation>
listed
<taxonomicName id="A3B31F888A382F33FD61F832FCD6F812" box="[736,840,1938,1961]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A382F33FD61F832FD71F812" box="[736,751,1938,1961]" italics="true" pageId="4" pageNumber="139">L</emphasis>
.
<emphasis id="56C7B8198A382F33FD7CF832FCD6F812" box="[765,840,1938,1961]" italics="true" pageId="4" pageNumber="139">urania</emphasis>
</taxonomicName>
among the Indo-West Pacific leucosiids and cited the records by Alcock and Rathbun.
<bibRefCitation id="002219FA8A382F33FD8DF815FD58F875" author="Naiyanetr" box="[524,710,1973,1998]" pageId="4" pageNumber="139" refString="Naiyanetr, P. (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, 161 pp." type="book" year="1998">Naiyanetr (1998</bibRefCitation>
,
<bibRefCitation id="002219FA8A382F33FD52F816FC93F875" author="Naiyanetr" box="[723,781,1974,1998]" pageId="4" pageNumber="139" refString="Naiyanetr, P. (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">2007</bibRefCitation>
) listed
<taxonomicName id="A3B31F888A382F33FCE0F817FC54F875" box="[865,970,1975,1998]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A382F33FCE0F817FCEEF875" box="[865,880,1975,1998]" italics="true" pageId="4" pageNumber="139">L</emphasis>
.
<emphasis id="56C7B8198A382F33FCFEF817FC54F875" box="[895,970,1975,1998]" italics="true" pageId="4" pageNumber="139">urania</emphasis>
</taxonomicName>
in
<collectingCountry id="1CA4249B8A382F33FC71F815FBC8F875" box="[1008,1110,1973,1998]" name="Thailand" pageId="4" pageNumber="139">Thailand</collectingCountry>
, citing the localities cited by Rathbun, and doubtlessly referring to her misidentified material.
</paragraph>
<paragraph id="640C640B8A392F32FF46FF38FE9EFE16" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="140">
<taxonomicName id="A3B31F888A392F32FF46FF38FE05FF0B" box="[199,411,152,176]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A392F32FF46FF38FE05FF0B" box="[199,411,152,176]" italics="true" pageId="5" pageNumber="140">Coleusia huilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DF405628A392F32FE20FF37FE41FF0B" box="[417,479,151,176]" pageId="5" pageNumber="140" rank="species">
<emphasis id="56C7B8198A392F32FE20FF37FE41FF0B" bold="true" box="[417,479,151,176]" pageId="5" pageNumber="140">n. sp.</emphasis>
</taxonomicNameLabel>
, like the closely allied
<taxonomicName id="A3B31F888A392F32FD60FF38FC67FF0B" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[737,1017,151,176]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A392F32FD60FF38FCD3FF14" box="[737,845,152,175]" italics="true" pageId="5" pageNumber="140">C. urania</emphasis>
(Herbst, 1801)
</taxonomicName>
, differs from their cogeners in having prominently granular pereiopodal meri and expanded, leaf-like pereiopodal propodi.
<taxonomicName id="A3B31F888A392F32FBD0FF1CFAB8FF6F" box="[1105,1318,188,212]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="huilianae" status="sp. nov.">
<emphasis id="56C7B8198A392F32FBD0FF1CFAB8FF6F" box="[1105,1318,188,212]" italics="true" pageId="5" pageNumber="140">Coleusia huilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DF405628A392F32FAACFF1CFAF2FF6E" box="[1325,1388,188,213]" pageId="5" pageNumber="140" rank="species">
<emphasis id="56C7B8198A392F32FAACFF1CFAF2FF6E" bold="true" box="[1325,1388,188,213]" pageId="5" pageNumber="140">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="A3B31F888A392F32FF16FF40FE9CFF4C" box="[151,258,224,247]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A392F32FF16FF40FE9CFF4C" box="[151,258,224,247]" italics="true" pageId="5" pageNumber="140">C. urania</emphasis>
</taxonomicName>
have a superficially similar colour pattern consisting of a broad pale stripe medially on the carapace, two large rust-colored blotches posteriorly, and two smaller blotches on the posterolateral margins. The new species differs from
<taxonomicName id="A3B31F888A392F32FEA7FE88FE0DFE84" box="[294,403,296,319]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="56C7B8198A392F32FEA7FE88FE0DFE84" box="[294,403,296,319]" italics="true" pageId="5" pageNumber="140">C. urania</emphasis>
</taxonomicName>
, however, in the shape of the apical process of the first male pleopod which is digitate and curved laterally in the former, beak-like and curved interiorly in the latter; granulation on the posterior and lower margins of cheliped and ambulatory meri smaller and sparser, and entirely missing from lower external surface of the palm.
</paragraph>
</subSubSection>
<subSubSection id="2CA937808A392F32FF46FE17FE23FD3E" pageId="5" pageNumber="140" type="distribution">
<paragraph id="640C640B8A392F32FF46FE17FBCBFE6B" blockId="5.[151,1437,151,645]" box="[199,1109,439,464]" pageId="5" pageNumber="140">
<emphasis id="56C7B8198A392F32FF46FE17FEC7FE6B" bold="true" box="[199,345,439,464]" pageId="5" pageNumber="140">Distribution</emphasis>
.
<collectingCountry id="1CA4249B8A392F32FEE6FE17FE3FFE6B" box="[359,417,439,464]" name="India" pageId="5" pageNumber="140">India</collectingCountry>
(Tamil Nadu and Andaman Is.), Gulf of
<collectingCountry id="1CA4249B8A392F32FCEAFE17FC4FFE6B" box="[875,977,439,464]" name="Thailand" pageId="5" pageNumber="140">Thailand</collectingCountry>
,
<collectingCountry id="1CA4249B8A392F32FC5DFE17FBCFFE6B" box="[988,1105,439,464]" name="Singapore" pageId="5" pageNumber="140">Singapore</collectingCountry>
.
</paragraph>
<paragraph id="640C640B8A392F32FF46FE7CFE23FD3E" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="140">
<emphasis id="56C7B8198A392F32FF46FE7CFE85FE4E" bold="true" box="[199,283,476,501]" pageId="5" pageNumber="140">Colour</emphasis>
(
<figureCitation id="FC88788E8A392F32FEB1FE7CFEE9FE4F" box="[304,375,476,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1885,1907]" captionTargetBox="[322,1264,1070,1852]" captionTargetId="figure@1.[322,1264,1070,1853]" captionTargetPageId="1" captionText="FIGURE 1. Coleusia huilianae n. sp., paratype male (carapace length 37.2 mm, carapace width 33.3) (ZRC 2000.0094). Dorsal view of carapace, colours in life." httpUri="https://zenodo.org/record/224464/files/figure.png" pageId="5" pageNumber="140">Fig. 1</figureCitation>
). Dorsal surface of carapace olive green, paler laterally and posteriorly; a whitish stripe proceeding from the front to the gastric region. Two large rust coloured spots on the posterior part of the carapace, two smaller spots near lateral margin above first ambulatory leg. The anterior margins of the thoracic sinus, the basal portion of the fingers, and a ring on each segment of the ambulatory legs, are all bright orange colored. Ventral surface paler grey.
</paragraph>
</subSubSection>
</treatment>
</document>