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<document ID-DOI="http://dx.doi.org/10.3897/jhr.81.57792" ID-GBIF-Dataset="7c1bea40-8c93-4ba1-9faa-97fcf81ffdad" ID-Pensoft-Pub="1314-2607-81-43" ID-Pensoft-UUID="D48C8138C7E35800953C86D433828718" ID-Zenodo-Dep="4584245" ID-ZooBank="6E3EC3E6F3EF4AA59B2A10C4D719CE19" ModsDocID="1314-2607-81-43" checkinTime="1614373120647" checkinUser="pensoft" docAuthor="Olmi, Massimo, Chen, Hua-Yan, Shih, Chungkun, Mueller, Patrick, Capradossi, Leonardo, Ren, Dong, Perkovsky, Evgeny E. &amp; Guglielmino, Adalgisa" docDate="2021" docId="EC11012F24245F6398D34D94E886C4C0" docLanguage="en" docName="JourHymenoptRes 81: 43-55" docOrigin="Journal of Hymenoptera Research 81" docPubDate="2021-02-25" docSource="http://dx.doi.org/10.3897/jhr.81.57792" docTitle="Hybristodryinus zaifui Chen, Olmi &amp; Perkovsky 2021, sp. nov." docType="treatment" docUuid="6455A84B-B0A6-42E6-818E-F7B4F6586BE8" docUuidSource="ZooBank" docVersion="4" id="D48C8138C7E35800953C86D433828718" lastPageNumber="43" masterDocId="D48C8138C7E35800953C86D433828718" masterDocTitle="New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts" masterLastPageNumber="55" masterPageNumber="43" pageNumber="43" updateTime="1678755185687" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Olmi, Massimo</mods:namePart>
<mods:affiliation>Tropical Entomology Research Center, Viterbo 01100, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Hua-Yan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0382-1635</mods:nameIdentifier>
<mods:affiliation>State Key Laboratory of Biocontrol, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
<mods:nameIdentifier type="email">huayanc@scbg.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shih, Chungkun</mods:namePart>
<mods:affiliation>College of Life Sciences, Capital Normal University, Haidian District, Beijing 100048, China &amp; Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mueller, Patrick</mods:namePart>
<mods:affiliation>Friedhofstrasse 9, Kaeshofen 66894, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Capradossi, Leonardo</mods:namePart>
<mods:affiliation>Via Pericle Scriboni 28, Tuscania 01017, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ren, Dong</mods:namePart>
<mods:affiliation>College of Life Sciences, Capital Normal University, Haidian District, Beijing 100048, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perkovsky, Evgeny E.</mods:namePart>
<mods:affiliation>Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev 01601, Ukraine &amp; Borissiak Paleontological Institute of the Russian Academy of Sciences, Moscow 117997, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guglielmino, Adalgisa</mods:namePart>
<mods:affiliation>Department of Agriculture and Forest Sciences (DAFNE), University of Tuscia, Viterbo 01100, Italy</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>81</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>43</mods:start>
<mods:end>55</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.81.57792</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.81.57792</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-81-43</mods:identifier>
<mods:identifier type="ZooBank">6E3EC3E6F3EF4AA59B2A10C4D719CE19</mods:identifier>
<mods:identifier type="Pensoft-UUID">D48C8138C7E35800953C86D433828718</mods:identifier>
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<treatment ID-GBIF-Taxon="179918098" LSID="urn:lsid:zoobank.org:act:6455A84B-B0A6-42E6-818E-F7B4F6586BE8" httpUri="http://treatment.plazi.org/id/EC11012F24245F6398D34D94E886C4C0" lastPageNumber="43" pageId="0" pageNumber="43">
<subSubSection pageId="0" pageNumber="43" type="nomenclature">
<paragraph pageId="0" pageNumber="43">
<taxonomicName LSID="http://zoobank.org/6455A84B-B0A6-42E6-818E-F7B4F6586BE8" authority="Chen, Olmi &amp; Perkovsky" authorityName="Chen, Olmi &amp; Perkovsky" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus zaifui" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="zaifui" status="sp. nov.">
Hybristodryinus zaifui Chen
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hybristodryinus zaifui sp. nov., female, holotype A fore leg, 5 t = protarsomere 5, Ec = enlarged claw B claw C sting." figureDoi="10.3897/jhr.81.57792.figure3" httpUri="https://binary.pensoft.net/fig/512988" pageId="0" pageNumber="43">, Olmi</figureCitation>
&amp; Perkovsky
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="description">
<paragraph pageId="0" pageNumber="43">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">Figures 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hybristodryinus zaifui sp. nov., female, holotype A fore leg, 5 t = protarsomere 5, Ec = enlarged claw B claw C sting." figureDoi="10.3897/jhr.81.57792.figure3" httpUri="https://binary.pensoft.net/fig/512988" pageId="0" pageNumber="43">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="diagnosis">
<paragraph pageId="0" pageNumber="43">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="43">
Female: macropterous, with disc of pronotum shaped as an isosceles triangle; notauli incomplete, reaching about 0.7
<normalizedToken originalValue="×">x</normalizedToken>
length of mesoscutum (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2C</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="4584249" doi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 2.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
right,
<taxonomicName authorityName="Chen, Olmi &amp; Perkovsky" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus zaifui" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="zaifui">
<emphasis italics="true" pageId="0" pageNumber="43">Hybristodryinus zaifui</emphasis>
</taxonomicName>
sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of
<taxonomicName authorityName="Shcherbakov" authorityYear="2007" class="Insecta" family="Perforissidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="family">Perforissidae</taxonomicName>
(
<taxonomicName authorityName="Shcherbakov" authorityYear="2007" class="Insecta" family="Perforissidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="family">Perforissidae</taxonomicName>
), ventral view
<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
left,
<taxonomicName authorityName="Chen, Olmi &amp; Perkovsky" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus zaifui" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="zaifui">
<emphasis italics="true" pageId="0" pageNumber="43">Hybristodryinus zaifui</emphasis>
</taxonomicName>
sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of
<taxonomicName authorityName="Shcherbakov" authorityYear="2007" lsidName="" pageId="0" pageNumber="43" rank="subFamily" subFamily="Cixitettiginae">Cixitettiginae</taxonomicName>
, habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="43">C, D</emphasis>
<taxonomicName authorityName="Chen, Olmi &amp; Perkovsky" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus zaifui" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="zaifui">
<emphasis italics="true" pageId="0" pageNumber="43">Hybristodryinus zaifui</emphasis>
</taxonomicName>
sp. nov., female, holotype:
<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
head and mesosoma, dorsal view
<emphasis bold="true" pageId="0" pageNumber="43">D</emphasis>
head and mesosoma, latero-ventral view
<emphasis bold="true" pageId="0" pageNumber="43">E</emphasis>
antenna, arrows indicate the ADOs in the antennomeres 6-10 (one ADO per antennomere, except two ADOs in antennomere 10)
<emphasis bold="true" pageId="0" pageNumber="43">F</emphasis>
wings, 1Cu = cubital 1 cell, 2r-rs&amp;Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="description">
<paragraph pageId="0" pageNumber="43">Description.</paragraph>
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Female.</emphasis>
Macropterous (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2A, B</figureCitation>
); length 1.8 mm. Apparently completely brown-testaceous. Antenna clavate, apparently almost hairless; antennomeres in following proportions: 6:4:5:4:4:3.5:3.5:3:3:4; antenna about three times as long as head (37:12), reaching metapectal-propodeal disc. ADOs apparently present in antennomeres 6-10 (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2E</figureCitation>
). Flagellomeres longer than wide; antennomere 9 about three times as long as broad. Head excavated, apparently unsculptured; clypeus not visible; mandible not visible; face slightly concave; occipital carina complete (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2C</figureCitation>
); vertex without two longitudinal keels connecting lateral ocelli to occipital carina; occiput deeply excavated; eye normally bulging; ocellar ratio: POL:OL:OOL:OPL = 1:2:2:1; temple about twice as long as OOL (4:2); greatest breadth of lateral ocellus longer than POL (2:1); frontal line apparently complete; palpal formula probably 6/3 (labial palpus not distinct). Pronotum crossed by anterior deep transverse impression between anterior collar and disc; disc humped, apparently isosceles triangle shaped, not deeply medially longitudinally excavated; posterior collar absent; anterior surface of pronotum not sculptured by longitudinal striae; sculpture of disc not distinct; pronotum shorter than head (6:12); anterior collar of pronotum much shorter than disc (2:4); pronotal tubercle reaching tegula. Epicnemium not visible. Mesoscutum apparently unsculptured, longer than pronotum (10:6). Notauli incomplete, reaching about 0.7
<normalizedToken originalValue="×">x</normalizedToken>
length of mesoscutum (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2C</figureCitation>
). Mesoscutellum apparently unsculptured, shorter than mesoscutum (4:10). Metanotum shorter than mesoscutellum (2:4), with sculpture indistinct. Metapectal-propodeal complex shorter than mesoscutum (5:10), with disc reticulate rugose, with areolae very broad, with posterior corners lamina shaped, strongly projected posteriorly (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2D</figureCitation>
); propodeal declivity not visible. Fore wing (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2F</figureCitation>
) apparenly hyaline, not darkened, with the usual venation of
<taxonomicName authorityName="Haliday" authorityYear="1833" lsidName="" pageId="0" pageNumber="43" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
; pterostigma much longer than broad (10:2); second radial cell (2R1) open; stigmal vein (2r-rs&amp;Rs) regularly curved; fore wing with usual three basal cells (C, R, 1Cu) clearly enclosed by pigmented veins. Hind wing not visible. Fore leg ratio: 7 (coxa): 5 (trochanter): 15 (femur): 9 (tibia):? (tarsus only partly visible); enlarged claw (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hybristodryinus zaifui sp. nov., female, holotype A fore leg, 5 t = protarsomere 5, Ec = enlarged claw B claw C sting." figureDoi="10.3897/jhr.81.57792.figure3" httpUri="https://binary.pensoft.net/fig/512988" pageId="0" pageNumber="43">3A, B</figureCitation>
) slightly shorter than protarsomere 5; protrochanter very slender and long, with long and slender proximal stalk, broadened after half-way; protrochanter less than seven times as long as broad (5:1); protarsomere 3 produced into hook; rudimentary claw not visible; arolium not visible; enlarged claw apparently with one subapical tooth, with one row of lamellae (number of lamellae uncertain). Protarsomere 5 (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hybristodryinus zaifui sp. nov., female, holotype A fore leg, 5 t = protarsomere 5, Ec = enlarged claw B claw C sting." figureDoi="10.3897/jhr.81.57792.figure3" httpUri="https://binary.pensoft.net/fig/512988" pageId="0" pageNumber="43">3B</figureCitation>
) with inner side not visible; lamellae of protarsomere 5 not visible. Mid leg ratio: 5 (coxa): 3 (trochanter): 13 (femur): 12 (tibia): 16 (tarsus). Mid trochanter short and broad. Hind leg ratio: 6 (coxa): 3 (trochanter): 18 (femur): 17 (tibia): 21 (tarsus). Metasomal petiole very short. Metasoma (excluding sting (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hybristodryinus zaifui sp. nov., female, holotype A fore leg, 5 t = protarsomere 5, Ec = enlarged claw B claw C sting." figureDoi="10.3897/jhr.81.57792.figure3" httpUri="https://binary.pensoft.net/fig/512988" pageId="0" pageNumber="43">3C</figureCitation>
)) longer than mesosoma (39:27). Tibial spurs 1/1/2.
</paragraph>
<caption ID-Zenodo-Dep="4584251" doi="10.3897/jhr.81.57792.figure3" httpUri="https://binary.pensoft.net/fig/512988" pageId="0" pageNumber="43" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 3.</emphasis>
<taxonomicName authorityName="Chen, Olmi &amp; Perkovsky" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus zaifui" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="zaifui">
<emphasis italics="true" pageId="0" pageNumber="43">Hybristodryinus zaifui</emphasis>
</taxonomicName>
sp. nov., female, holotype
<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
fore leg, 5t = protarsomere 5, Ec = enlarged claw
<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
claw
<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
sting.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="materials_examined">
<paragraph pageId="0" pageNumber="43">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="43">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Myanmar" county="Burmese" location="Northern" municipality="Obtained" specimenCount="1" stateProvince="Kachin State" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="43">
<typeStatus>Holotype</typeStatus>
:
</emphasis>
female (CNN-HYM-MA 2017087),
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
: specimen in mid-Cretaceous
<collectingCounty>Burmese</collectingCounty>
amber (about 99 Ma).
<collectingMunicipality>Obtained</collectingMunicipality>
from a mine situated in
<location LSID="urn:lsid:plazi:treatment:EC11012F24245F6398D34D94E886C4C0:3728698A49D2B9C470BF26E1EA7C54BA" country="Myanmar" county="Burmese" municipality="Obtained" name="Northern" stateProvince="Kachin State">Northern</location>
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
,
<collectingRegion country="Myanmar" name="Kachin">Kachin State</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:EC11012F24245F6398D34D94E886C4C0:6E9497E53F62E6D09F8F1F0789AD09D6" country="Myanmar" county="Burmese" municipality="Obtained" name="Tanai Township" stateProvince="Kachin State">Tanai Township</location>
,
<location LSID="urn:lsid:plazi:treatment:EC11012F24245F6398D34D94E886C4C0:7A2A770D9CC0C9EBCF74141C68A4FC94" country="Myanmar" county="Burmese" municipality="Obtained" name="Hukawng Valley" stateProvince="Kachin State">Hukawng Valley</location>
,
<locationDeviation location="Tanai City">SW of Tanai City</locationDeviation>
(CNUB).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="etymology">
<paragraph pageId="0" pageNumber="43">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="43">
The new species is named after the late Prof. Zaifu Xu (South China Agricultural University, Guangzhou, Guangdong, China), for his important contribution to the study of Chinese
<taxonomicName class="Insecta" family="Dryinidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="family">Dryinidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="remarks">
<paragraph pageId="0" pageNumber="43">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="43">
For its isosceles triangle shaped pronotal disc,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. zaifui" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="zaifui">
<emphasis italics="true" pageId="0" pageNumber="43">H. zaifui</emphasis>
</taxonomicName>
is similar to
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. nalae" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="nalae">
<emphasis italics="true" pageId="0" pageNumber="43">H. nalae</emphasis>
</taxonomicName>
Perkovsky, Olmi,
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
&amp; Martynova, 2019,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. concavifrons" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="concavifrons">
<emphasis italics="true" pageId="0" pageNumber="43">H. concavifrons</emphasis>
</taxonomicName>
Perkovsky, Olmi,
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
&amp; Martynova, 2019,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. resinicolus" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="resinicolus">
<emphasis italics="true" pageId="0" pageNumber="43">H. resinicolus</emphasis>
</taxonomicName>
Engel, 2005 and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. ligulatus" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis italics="true" pageId="0" pageNumber="43">H. ligulatus</emphasis>
</taxonomicName>
Perkovsky, Olmi,
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
&amp; Martynova, 2019. However,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. zaifui" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="zaifui">
<emphasis italics="true" pageId="0" pageNumber="43">H. zaifui</emphasis>
</taxonomicName>
has incomplete notauli (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A right, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-dorsal view; left, nymph of Perforissidae (Perforissidae), ventral view B left, Hybristodryinus zaifui sp. nov., female, holotype, habitus, latero-ventral view; right, nymph of Cixitettiginae, habitus, dorsal view C, D Hybristodryinus zaifui sp. nov., female, holotype: C head and mesosoma, dorsal view D head and mesosoma, latero-ventral view E antenna, arrows indicate the ADOs in the antennomeres 6 - 10 (one ADO per antennomere, except two ADOs in antennomere 10) F wings, 1 Cu = cubital 1 cell, 2 r-rs &amp; Rs = stigmal vein, C = costal cell, P = pterostigma, R = radial cell." figureDoi="10.3897/jhr.81.57792.figure2" httpUri="https://binary.pensoft.net/fig/512987" pageId="0" pageNumber="43">2C</figureCitation>
), reaching about 0.7
<normalizedToken originalValue="×">x</normalizedToken>
length of mesoscutum, whereas in the other species the notauli are complete.
</paragraph>
</subSubSection>
</treatment>
</document>

76
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<paragraph id="C3A629CCB77050609E0DEC27FEAC3671" pageId="0" pageNumber="24071">Notes</paragraph>
<paragraph id="334A1F2CF2ECC066E9837D2FBD92DD30" pageId="0" pageNumber="24071">
Widespread but previously unrecorded for Arkansas (
<bibRefCitation id="37C1B7434B3D036598D992651EF93F71" author="Mitchell, T. B." journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="24071" pagination="1 - 44" title="A revision of the genus Megachile in the nearctic region. Part II" volume="61" year="1935">Mitchell 1935</bibRefCitation>
,
<bibRefCitation id="1F38F7E1AD5F27FCC2EFE2C65497171A" author="Bzdyk, E L" journalOrPublisher="ZooKeys" pageId="0" pageNumber="24071" pagination="31 - 61" title="A revision of the MegachilesubgenusLitomegachile Mitchell with an illustrated key and description of a new species (Hymenoptera, Megachilidae, Megachilini)" volume="221" year="2012">Bzdyk 2012</bibRefCitation>
). Opportunistic (Table 1: Sites 2, 3, 15).
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<bibRefCitation author="Prout, L. B." journalOrPublisher="Novitates Zoologicae" pageId="0" pageNumber="5447" pagination="1 - 32" title="New Geometridae." volume="33" year="1926">Prout 1926</bibRefCitation>
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Type status:
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. Occurrence: sex:
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<paragraph pageId="0" pageNumber="5447">Type locality: [West Papua], Mt Goliath, 5000-7000 ft.</paragraph>
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