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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="88D168D0245B5B6D98C171AA188EEE32" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole vadum Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="F85769B8-4A08-4652-8686-CFD9B7F3A620" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salata, Sebastian</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>949</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>185</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="165090172" LSID="urn:lsid:zoobank.org:act:F85769B8-4A08-4652-8686-CFD9B7F3A620" httpUri="http://treatment.plazi.org/id/88D168D0245B5B6D98C171AA188EEE32" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="88D168D0-245B-5B6D-98C1-71AA188EEE32" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vadum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vadum" status="sp. nov.">Pheidole vadum</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">Figs 59A-F</figureCitation>
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64Q</figureCitation>
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Distribution. Pheidole renirano sp. nov. (A). Pheidole sava sp. nov. (B). Pheidole sikorae Forel (C). Pheidole sofia sp. nov. (D). Pheidole sparsa sp. nov. (E). Pheidole tampony sp. nov. (F). Pheidole trichotos sp. nov. (G). Pheidole tsaravoniana sp. nov. (H). Pheidole vadum sp. nov. (I). Pheidole veteratrix Forel (J). Pheidole volontany sp. nov. (K). Pheidole vony sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure67" httpUri="https://binary.pensoft.net/fig/432236" pageId="0" pageNumber="1">, 67I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
Madagascar. • 1 major worker; Antananarivo; 3 km 41°NE Andranomay, 11.5 km 147°SSE Anjozorobe; -18.47333, 47.96; alt. 1300 m; 5 Dec 2000; Fisher et al. leg.; montane rainforest, ex dead twig above ground; BLF02436; CASENT0427789 (CASC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
• 4w., 1s.; same data as for holotype; CASENT0427788, CASENT0427791, CASENT0427785 (CASC, MHNG).
</paragraph>
<caption doi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1" start="Figure 59" startId="F59">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 59.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vadum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vadum</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0427785) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0427789).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Moderately large species.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Head in full-face view oval, not widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval, ventral and dorsal faces convex, occipital cleft very shallow; sides of the head with dense, moderately long, suberect pilosity; medial frons with moderately dense, thin, longitudinal to irregular and interrupted rugae, rugae in posteromedial part slightly directed outward, interspaces shiny with sparse and distinct rugofoveolae; occipital lobes and area posterolateral from eyes never smooth; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and not connected by concavity; mesosoma with thick and sparse foveolae; lateral sides of pronotum and propodeum with smooth notches; body yellowish brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Head foveolate; anteromedial part of frons with smooth notch; area posterolateral from eyes predominantly smooth; scape, when laid back, surpassing the posterior head margin by one-fifth of its length; promesonotum moderately high and short; promesonotal groove absent; propodeal spines minute and triangular; mesosoma with thick foveolae; body yellowish brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 2): HL: 0.96, 1.04; HW: 0.92, 1.01; SL: 0.64, 0.63; EL: 0.12, 0.13; WL: 0.94, 1.0; PSL: 0.17, 0.18; MTL: 0.61, 0.62; PNW: 0.38, 0.44; PTW: 0.13, 0.14; PPW: 0.23, 0.25; CI: 103.5, 103.0; SI: 69.7, 62.4; PSLI: 17.7, 17.4; PPI: 57.8, 53.6; PNI: 41.0, 43.4; MTI: 66.0, 61.2.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
In full-face view oval, not widening posteriorly, with anterior and posterior sides convex (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59B</figureCitation>
); occipital cleft very shallow. In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with dense, moderately long, suberect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with moderately dense, thin, longitudinal to irregular and interrupted rugae, rugae in posteromedial part slightly directed outward, interspaces shiny with sparse and distinct rugofoveolae; anterolateral sides with longitudinal, thin, and relatively dense rugae; posterolateral sides with irregular, thin, and relatively dense rugae; interspaces shiny with relatively dense and distinct rugofoveolae. Occipital lobes with sparse, thick, and irregular rugae; interspaces rugofoveolate. Gena with relatively sparse, thick, and longitudinal rugae and rugofoveolate interspaces. Area posterolateral from eyes shiny, with dense rugofoveolae. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59B, D</figureCitation>
). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64Q</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, moderately narrow, with acute apex; humeral area laterally weakly produced (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59D</figureCitation>
). Surface shiny with thick and sparse foveolae; lateral sides of pronotum and propodeum with smooth notches. Pilosity sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
Shiny with fine and sparse foveolae; node smooth, low, triangular, with rounded and thin apex, in rear view node dorsoventrally slightly concave; pilosity moderately sparse and erect (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
Shiny and smooth; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
Shiny and smooth; pilosity moderately dense, long, and erect (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Yellowish brown; mandibles and gaster slightly darker; legs yellowish (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59D, F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 4): HL: 0.52-0.59 (0.56); HW: 0.46-0.51 (0.49); SL: 0.55-0.59 (0.56); EL: 0.1-0.11 (0.11); WL: 0.72-0.73 (0.73); PSL: 0.07-0.08 (0.08); MTL: 0.47-0.51 (0.49); PNW: 0.3-0.33 (0.31); PTW: 0.07-0.09 (0.08); PPW: 0.13-0.14 (0.13); CI: 112.7-116.7 (114.1); SI: 111.3-117.9 (114.9); PSLI: 11.3-15.9 (13.6); PPI: 53.0-62.8 (59.3); PNI: 57.6-65.0 (62.2); MTI: 98.8-101.2 (100.5).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
Cephalic margin indistinctly convex or straight (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59A</figureCitation>
). Pilosity relatively sparse, long, decumbent to suberect. Sculpture foveolate; anteromedial part of frons with smooth notch; area posterolateral from eyes predominantly smooth. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59A, C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines minute, triangular (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59C</figureCitation>
). Sculpture shiny and with thick foveolae. Pilosity very sparse, moderately long, and erect (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
Short, low, and relatively flat; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
With sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Yellowish brown, vertex slightly darker (Fig.
<figureCitation captionStart="Figure 59" captionStartId="F59" captionText="Figure 59. Pheidole vadum sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0427785) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0427789)." figureDoi="10.3897/zookeys.949.51269.figure59" httpUri="https://binary.pensoft.net/fig/432228" pageId="0" pageNumber="1">59C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Latin for shallow in reference to very shallow occipital cleft.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at 1300 m in elevation, in montane rainforest. Nest was located in a dead twig above ground.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vadum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vadum</emphasis>
</taxonomicName>
sp. nov. is a member of the group of species characterised by major workers with head in full-face view oval and not widening posteriorly and medial part of frons with thick, moderately sparse, irregular rugae or medial frons with moderately dense, thin, longitudinal anteriorly to irregular posteriorly, interrupted rugae and very shallow occipital cleft. Minor workers of this group have a short and moderately high promesonotum and dark body colouration ranging from orange to brown. The group consists of three species:
<taxonomicName lsidName="P. vadum" pageId="0" pageNumber="1" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">P. vadum</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. analavelona" pageId="0" pageNumber="1" rank="species" species="analavelona">
<emphasis italics="true" pageId="0" pageNumber="1">P. analavelona</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName lsidName="P. ambohimanga" pageId="0" pageNumber="1" rank="species" species="ambohimanga">
<emphasis italics="true" pageId="0" pageNumber="1">P. ambohimanga</emphasis>
</taxonomicName>
sp. nov.
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vadum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vadum</emphasis>
</taxonomicName>
sp. nov. is known from the vicinity of Antananarivo and is sympatric with
<taxonomicName lsidName="P. ambohimanga" pageId="0" pageNumber="1" rank="species" species="ambohimanga">
<emphasis italics="true" pageId="0" pageNumber="1">P. ambohimanga</emphasis>
</taxonomicName>
sp. nov. Its majors can be easily separated from
<taxonomicName lsidName="P. ambohimanga" pageId="0" pageNumber="1" rank="species" species="ambohimanga">
<emphasis italics="true" pageId="0" pageNumber="1">P. ambohimanga</emphasis>
</taxonomicName>
sp. nov. based on presence of moderately dense, thin, longitudinal anteriorly to irregular posteriorly, interrupted rugae on medial frons and very shallow occipital cleft; minor workers can be separated based on predominantly foveolate head sculpture lacking additional rugae. Additionally, minors of
<taxonomicName lsidName="P. vadum" pageId="0" pageNumber="1" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">P. vadum</emphasis>
</taxonomicName>
sp. nov. can be confused with workers of other members of the
<taxonomicName lsidName="sikorae" pageId="0" pageNumber="1" rank="species" species="sikorae">
<emphasis italics="true" pageId="0" pageNumber="1">sikorae</emphasis>
</taxonomicName>
group, especially with workers of
<taxonomicName lsidName="P. sava" pageId="0" pageNumber="1" rank="species" species="sava">
<emphasis italics="true" pageId="0" pageNumber="1">P. sava</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. sparsa" pageId="0" pageNumber="1" rank="species" species="sparsa">
<emphasis italics="true" pageId="0" pageNumber="1">P. sparsa</emphasis>
</taxonomicName>
sp. nov., both known from the northern part of the island. However, minor workers of
<taxonomicName lsidName="P. vadum" pageId="0" pageNumber="1" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">P. vadum</emphasis>
</taxonomicName>
sp. nov. can be separated based on the combination of the following characters: body yellowish brown, head and mesosoma predominantly foveolate with no additional sculpture, scape, when laid back, surpassing the posterior head margin by one-fifth of its length, promesonotum short and moderately high, and promesonotal groove absent.
</paragraph>
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Soula, 2008: 10.
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<bibRefCitation author="Ohaus, F" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="1" pagination="283 - 329" refId="B266" refString="Ohaus, F, 1905. Beitraege zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329" title="Beitraege zur Kenntniss der amerikanischen Ruteliden." volume="66" year="1905">Ohaus 1905</bibRefCitation>
: 324, by monotypy.
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113
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<emphasis id="0B11FF1282ED94DDEA952F51428EFF17" italics="true" pageNumber="457">Hesperoptenus (Hesperoptenus) doriae</emphasis>
Peters 1868
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<emphasis id="4F78F1064D0DDB03FE6E8F9DA8C2A08F" italics="true" pageNumber="457">Hesperoptenus (Hesperoptenus) doriae</emphasis>
Peters 1868
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,
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Monatsb. K. Preuss. Akad. Wiss.
<collectingRegion id="25B5AD4CD0A4D3AB7D29EE4D2392AA6C" country="Germany" name="Berlin">Berlin</collectingRegion>
, 1868: 626
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.
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<paragraph id="4D61919F2E55EEECAAEB05AE96244E84" pageNumber="457">
<emphasis id="E812880535AA9F6203D48AD06E742246" inLineHeading="true" pageNumber="457">Type Locality:</emphasis>
<materialsCitation id="17494B1AC64950665D12A283BB73CC6B" ID-GBIF-Occurrence="4867287623" country="Malaysia" pageNumber="457">
<collectingCountry id="45CFCEB5414242AA34AAFD6CA58765C0" name="Malaysia">Malaysia</collectingCountry>
, Borneo,
<collectingCountry id="B3DB4A1778927D94BABCE7263F89BD0B" name="Malaysia">Sarawak</collectingCountry>
.
</materialsCitation>
</paragraph>
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<subSubSection id="03C2657FC8C85B86DB4258078CFDC4CA" pageNumber="457" type="vernacular_names">
<paragraph id="993A9F103DEC0D64E566050AC61A2365" pageNumber="457">
<emphasis id="E486AE3542EFBBB41BD02E0B1B231BAB" inLineHeading="true" pageNumber="457">Vernacular Names:</emphasis>
<vernacularName id="5748BCE8B92939458FEA731AAAF91E9F" language="eng" pageNumber="457">False Serotine Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="7ADD8082B1E0B4278DA1DCB94023F9CD" pageNumber="457" type="distribution">
<paragraph id="7C5C9EC7397A9780EB513BDE25426A0A" pageNumber="457">
<emphasis id="A39E3DDC9F61FFC7C1706B6EDE39A27F" inLineHeading="true" pageNumber="457">Distribution:</emphasis>
Borneo, Malay Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="37CCE806E248AD549F6C00B405DE3978" pageNumber="457" type="conservation">
<paragraph id="8BD15F687AAB2689CF5E5F9E0635C5E3" pageNumber="457">
<emphasis id="EDB5992457AFE80FFE9D36C85F6F3996" inLineHeading="true" pageNumber="457">Conservation:</emphasis>
<collectionCode id="2A4B5E9570901F35DA3C5329CBEC5DF6">IUCN</collectionCode>
2003 and
<collectionCode id="553CCCCC6E3BFEB4B3986BBC03EA3EB0">IUCN</collectionCode>
/
<collectionCode id="5999A696A94A80D19C06EDE81B8B5466" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) – Endangered.
</paragraph>
</subSubSection>
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<paragraph id="7319DE2C98A924F11B517657FFFCE7B1" pageNumber="457">
<emphasis id="E28EFA4CC0B83F3105A49FD182775739" inLineHeading="true" pageNumber="457">Discussion:</emphasis>
Subgenus
<taxonomicName id="07473674C581FDBF48660A96C9158AC9" authorityName="Peters" authorityYear="1868" class="Mammalia" family="Vespertilionidae" genus="Hesperoptenus" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="genus">
<emphasis id="CEC27C87F59A27926A796CAAE940AD54" italics="true" pageNumber="457">Hesperoptenus</emphasis>
</taxonomicName>
.
</paragraph>
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</treatment>
</document>

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<document ID-CLB-Dataset="297104" ID-DOI="10.5281/zenodo.7316519" ID-GBIF-Dataset="908f4357-4e0b-40f8-8fb2-9e04c0b37137" ID-ISBN="0-8018-8221-4" checkinTime="1670905233444" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="88D1D3A2995E14E03FF18379AB1DBCE2" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Primates.111-184.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press" docTitle="Cebus albifrons subsp. versicolor Pucheran 1845" docType="treatment" docVersion="4" id="B0294F6FEAA127D846D57464E1129452" lastPageNumber="136" masterDocId="B0294F6FEAA127D846D57464E1129452" masterDocTitle="Order Primates" masterLastPageNumber="184" masterPageNumber="111" pageNumber="136" updateTime="1716785509035" updateUser="ExternalLinkService">
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<mods:title id="DAD661798344003B124758443237FEC2">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1</mods:title>
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<taxonomicName id="B4C4B6AFAEB28DF251FE6DA95B40D042" ID-CoL="RZ24" authority="Pucheran 1845" authorityName="Pucheran" authorityYear="1845" class="Mammalia" family="Cebidae" genus="Cebus" kingdom="Animalia" order="Primates" pageNumber="136" phylum="Chordata" rank="subSpecies" species="albifrons" subSpecies="versicolor">
<emphasis id="A2CA92B93CF45510EB3EFFB986F9FAA9" italics="true" pageNumber="136">Cebus albifrons</emphasis>
subsp.
<emphasis id="6D512AD270A8A10D1C311D62C7271283" italics="true" pageNumber="136">versicolor</emphasis>
Pucheran 1845
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<emphasis id="EC0D29AB78A65B11D76A27060C4C0A80" inLineHeading="true" pageNumber="136">Synonyms:</emphasis>
<taxonomicName id="8590E7C934FFE47FDAF34FCCBC1DC85E" authority="Hershkovitz 1849" authorityName="Hershkovitz" authorityYear="1849" class="Mammalia" family="Cebidae" genus="Cebus" kingdom="Animalia" order="Primates" pageNumber="136" phylum="Chordata" rank="subSpecies" species="albifrons" subSpecies="adustus">
<emphasis id="A661E37987F117251F81D6FE6293F5F8" italics="true" pageNumber="136">Cebus albifrons</emphasis>
subsp.
<emphasis id="07A5711642D3EF1B0D542FE3EEF67C49" italics="true" pageNumber="136">adustus</emphasis>
Hershkovitz 1849
</taxonomicName>
;
<taxonomicName id="2D396B6B169134935B3425FEC297E751" authority="Hershkovitz 1949" authorityName="Hershkovitz" authorityYear="1949" class="Mammalia" family="Cebidae" genus="Cebus" kingdom="Animalia" order="Primates" pageNumber="136" phylum="Chordata" rank="subSpecies" species="albifrons" subSpecies="cesarae">
<emphasis id="C6CFCE0CD4DD83756C2E16E256CC5C9B" italics="true" pageNumber="136">Cebus albifrons</emphasis>
subsp.
<emphasis id="EC50CB8D828EA12ADC39C9C2B8E4BB04" italics="true" pageNumber="136">cesarae</emphasis>
Hershkovitz 1949
</taxonomicName>
;
<taxonomicName id="A14DC260506902B4FCCB26C60A7B7FB2" authority="Gray 1865" authorityName="Gray" authorityYear="1865" class="Mammalia" family="Cebidae" genus="Cebus" kingdom="Animalia" order="Primates" pageNumber="136" phylum="Chordata" rank="subSpecies" species="albifrons" subSpecies="leucocephalus">
<emphasis id="CD0BB0A0F335814906BA79385BA86BD2" italics="true" pageNumber="136">Cebus albifrons</emphasis>
subsp.
<emphasis id="6226683B258FB3B889FE962CA77705DC" italics="true" pageNumber="136">leucocephalus</emphasis>
Gray 1865
</taxonomicName>
;
<taxonomicName id="12077E418EA5A281DFFD088AEC8BC0BC" authority="Elliot 1909" authorityName="Elliot" authorityYear="1909" class="Mammalia" family="Cebidae" genus="Cebus" kingdom="Animalia" order="Primates" pageNumber="136" phylum="Chordata" rank="subSpecies" species="albifrons" subSpecies="malitiosus">
<emphasis id="FD2A3338AB556F85F281C1DB2C687A3A" italics="true" pageNumber="136">Cebus albifrons</emphasis>
subsp.
<emphasis id="16A53E58B392836F38ECF682EB6E79AD" italics="true" pageNumber="136">malitiosus</emphasis>
Elliot 1909
</taxonomicName>
;
<taxonomicName id="082F2CBBF0BE612E4E18FE4A46803A82" authority="Hershkovitz 1949" authorityName="Hershkovitz" authorityYear="1949" class="Mammalia" family="Cebidae" genus="Cebus" kingdom="Animalia" order="Primates" pageNumber="136" phylum="Chordata" rank="subSpecies" species="albifrons" subSpecies="pleei">
<emphasis id="70771D923F9253AF991D6CE3046171B9" italics="true" pageNumber="136">Cebus albifrons</emphasis>
subsp.
<emphasis id="B7237B77B189A1C5C3C180ED90A4328B" italics="true" pageNumber="136">pleei</emphasis>
Hershkovitz 1949
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.
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