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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.63.8181" ID-PMC="PMC4956930" ID-Pensoft-Pub="1314-2003-63-77" ID-Pensoft-UUID="5844FFA9FFD4FF818A03FB22FFAE354C" ID-PubMed="27489480" ID-Zenodo-Dep="899025" ModsDocID="1314-2003-63-77" checkinTime="1464921839276" checkinUser="pensoft" docAuthor="Rabeler, Richard K. &amp; Wagner, Warren L." docDate="2016" docId="BA12743166BB5B418B562BF3F0394BFE" docLanguage="en" docName="PhytoKeys 63: 77-97" docOrigin="PhytoKeys 63" docPubDate="2016-06-02" docSource="http://dx.doi.org/10.3897/phytokeys.63.8181" docTitle="Odontostemma fimbriatum (Mattf.) Rabeler &amp; W. L. Wagner 2016, comb. nov." docType="treatment" docVersion="3" id="5844FFA9FFD4FF818A03FB22FFAE354C" lastPageNumber="83" masterDocId="5844FFA9FFD4FF818A03FB22FFAE354C" masterDocTitle="New combinations in Odontostemma (Caryophyllaceae)" masterLastPageNumber="97" masterPageNumber="77" pageNumber="83" updateTime="1668141699817" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New combinations in Odontostemma (Caryophyllaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rabeler, Richard K.</mods:namePart>
<mods:affiliation>University of Michigan Herbarium - EEB, 3600 Varsity Drive, Ann Arbor, MI 48108 - 2228, USA</mods:affiliation>
<mods:nameIdentifier type="email">rabeler@umich.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wagner, Warren L.</mods:namePart>
</mods:name>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-06-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>63</mods:number>
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<mods:start>77</mods:start>
<mods:end>97</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.63.8181</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.63.8181</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-63-77</mods:identifier>
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<treatment ID-GBIF-Taxon="182227072" LSID="urn:lsid:plazi:treatment:BA12743166BB5B418B562BF3F0394BFE" httpUri="http://treatment.plazi.org/id/BA12743166BB5B418B562BF3F0394BFE" lastPageNumber="83" pageId="6" pageNumber="83">
<subSubSection pageId="6" pageNumber="83" type="nomenclature">
<paragraph pageId="6" pageNumber="83">
<taxonomicName LSID="BA127431-66BB-5B41-8B56-2BF3F0394BFE" authority="(Mattf.) Rabeler &amp; W. L. Wagner" authorityName="(Mattf.) Rabeler &amp; W. L. Wagner" authorityYear="2016" baseAuthorityName="Mattf." class="Magnoliopsida" family="Caryophyllaceae" genus="Odontostemma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontostemma fimbriatum" order="Caryophyllales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="fimbriatum" status="comb. nov.">Odontostemma fimbriatum (Mattf.) Rabeler &amp; W.L. Wagner</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="83">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="83" type="reference_group">
<paragraph pageId="6" pageNumber="83">
<taxonomicName authorityName="Rabeler &amp; Wagner" authorityYear="2016" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arenaria fimbriata" order="Caryophyllales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="fimbriata" status="comb. nov.">Arenaria fimbriata</taxonomicName>
Mattf., Notizbl. Bot. Gart. Berlin-Dahlem 11: 335. 1932;
<taxonomicName authorityName="Rabeler &amp; Wagner" authorityYear="2016" class="Magnoliopsida" family="Caryophyllaceae" genus="Cerastium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cerastium fimbriatum" order="Caryophyllales" pageId="6" pageNumber="83" phylum="Tracheophyta" rank="species" species="fimbriatum" status="comb. nov.">Cerastium fimbriatum</taxonomicName>
E. Pritz., Bot. Jahrb. Syst. 29: 320. 1900, not Ledeb. (1815), nom. illeg.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="83" type="materials_examined">
<paragraph pageId="6" pageNumber="83">
<typeStatus>Syntypes</typeStatus>
.
</paragraph>
<paragraph pageId="6" pageNumber="83">
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="China" location="Sichuan" specimenCount="1" stateProvince="Sichuan" typeStatus="syntype">
<collectingCountry name="China">China</collectingCountry>
:
<collectingRegion country="China" name="Sichuan">Sichuan</collectingRegion>
?: highest regions, Tai pa Shan, August, Giraldi 1187 (B, presumably destroyed), Giraldi 1188 (B, presumably destroyed)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="762E342B104125D7935F3DFF5947F810" ENCODING="UTF-8" ID-DOI="https://doi.org/10.5281/zenodo.291971" ID-GBIF-Dataset="b91775a1-1123-41b5-9430-8a87164f966f" ID-ISBN="978-0-9506207-7-0" ID-Zenodo-Dep="291971" checkinTime="1588973768064" checkinUser="admin" docAuthor="Jarvis, Charlie" docDate="2007" docId="BA12B33E0E0DB9E85BE12D4C2395C6A5" docLanguage="en" docName="ch7.gg.S.xml" docOrigin="Order out of Chaos. Linnaean Plant Types and their Types, London: Linnaean Society of London in association with the Natural History Museum" docTitle="Solidago canadensis var. scabra Torr. &amp; A. Gray" docType="treatment" docVersion="11" lastPageNumber="863" masterDocId="D1B3EBBB44279C4C23B70DF5C0E42704" masterDocTitle="Chapter 7: Linnaean Plant Names and their Types (part S)" masterLastPageNumber="877" masterPageNumber="806" pageNumber="863" updateTime="1710905156370" updateUser="ExternalLinkService">
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<mods:title id="2987C4BA7BF5D6D5E1712444EBAD4C2D">Chapter 7: Linnaean Plant Names and their Types (part S)</mods:title>
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<mods:affiliation id="EB8B45DEAB228C203E80EA5CFD8C6B9A">Department of Botany, Natural History Museum, Cromwell Road, London, UK</mods:affiliation>
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<mods:title id="0654819C417AAC7028D4E67997D55383">Order out of Chaos. Linnaean Plant Types and their Types</mods:title>
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<treatment id="BA12B33E0E0DB9E85BE12D4C2395C6A5" ID-DOI="http://doi.org/10.5281/zenodo.4374183" ID-GBIF-Taxon="164016665" ID-Zenodo-Dep="4374183" LSID="urn:lsid:plazi:treatment:BA12B33E0E0DB9E85BE12D4C2395C6A5" httpUri="http://treatment.plazi.org/id/BA12B33E0E0DB9E85BE12D4C2395C6A5" lastPageNumber="863" pageNumber="863">
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<paragraph id="E8F5630D4B295CCBCBD1A8B328301079" pageNumber="863">
<treatmentCitation id="405A6F0DFD3DAE4AAEE10D187652FD35" author="Linnaeus" pageNumber="878" year="1753">
<bibCitation id="2F41B3D60C181C03B8C4FAB4F6E5E7C1" author="Linnaeus" pagination="878" title="Solidago altissima" volumeTitle="Species Plantarum 2" year="1753">
<taxonomicName id="D3D308FAF6950466EFB875DF0341C1F4" ID-CoL="4Y752" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Solidago" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="altissima">
<emphasis id="1E088A2BEF788E5390EF9E760157E967" bold="true" pageNumber="863">Solidago altissima</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="C436A08BF4FBC97516A984C22BAB225B">
<emphasis id="C120431E456645B2554B6863B03364ED" italics="true" pageNumber="863">Species Plantarum</emphasis>
2
</volumeTitle>
: 878. 1753
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6D1585519F66450FEBAA8DCE4A656D6E" pageNumber="863" type="distribution">
<paragraph id="FFF4B809B896125BCBD460E35BFF510E" pageNumber="863">&quot;Habitat in America septentrionali.&quot; RCN: 6348.</paragraph>
</subSubSection>
<subSubSection id="403A26BD1DB3199CBE474B8E031DE623" pageNumber="863" type="materials_examined">
<paragraph id="2146F8A6A668AFFBDB802C003B0A11BF" pageNumber="863">
<typeStatus id="CC05CB5426B4C72C6279F5506E698D01">Lectotype</typeStatus>
(Reveal &amp; al. in
<emphasis id="63CBEF4957FAA97B74AE3763FE2465B5" italics="true" pageNumber="863">Huntia</emphasis>
7: 238. 1987): [icon]
<emphasis id="732B1F126892D468A3E47C4AE7D300EB" italics="true" pageNumber="863">&quot;Virga aurea Marilandica, spicis florum racemosis, foliis integris, scabris&quot;</emphasis>
in Martyn, Hist. Pl. Rar.: 13, t. 13. 1728.
</paragraph>
</subSubSection>
<subSubSection id="CCEF6C3500B236E4320DE9CF0A44B9E3" pageNumber="863" type="nomenclature">
<paragraph id="F3C9D577C6875C1D394FB391BDB5E0BE" pageNumber="863">
Current name:
<taxonomicName id="3AC0FF80D512C7D6C5A18C611B8E73A8" ID-CoL="8R3MX" authority="Torr. &amp; A. Gray" authorityName="Torr. &amp; A. Gray" class="Magnoliopsida" family="Asteraceae" genus="Solidago" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="variety" species="canadensis" variety="scabra">
<emphasis id="EBDA7D9A352458156A9105F349909328" italics="true" pageNumber="863">
<emphasis id="8229F37A3FE56A0B254A4146699AB1F9" bold="true" italics="true" pageNumber="863">Solidago canadensis</emphasis>
</emphasis>
L. var
<emphasis id="2D7D7020C8814EE9A0E65DD3DBEF0570" italics="true" pageNumber="863">
<emphasis id="2B0FA166FC7D66443F51ADDD569B0255" bold="true" italics="true" pageNumber="863">scabra</emphasis>
</emphasis>
Torr. &amp; A. Gray
</taxonomicName>
(
<taxonomicName id="200BDEC5A59579AB8C61CD77EAA53100" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="95C11EDAC51EC5D4C07AEE147E05EF6D" pageNumber="863" type="discussion">
<paragraph id="9AD5D60912FE387876E0781EB77AE003" pageNumber="863">
<emphasis id="9B47925FF4D9EF22CD82761EFD89157A" italics="true" pageNumber="863">Note:</emphasis>
Gray (in
<emphasis id="FD4C4442174D5B4467B529672E0BDF30" italics="true" pageNumber="863">Proc. Amer. Acad. Arts</emphasis>
17: 177. 1882), in a study of the
<taxonomicName id="21CED18563431952C37FD790EF2E9174" class="Magnoliopsida" family="Asteraceae" genus="Solidago" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="genus">
<emphasis id="ADA4D21581954398E3C5B1C1A3F8FD11" italics="true" pageNumber="863">Solidago</emphasis>
</taxonomicName>
material in the Linnaean herbarium, concluded that the specimens associated with this name &quot;are confounded&quot; and that the cited Martyn plate is &quot;the true original of the Linnaean species&quot;.
<normalizedToken id="4492477F8EC084F3C7F52FAE722C60C7" originalValue="Gray’s">Gray's</normalizedToken>
conclusion was accepted and followed by Fernald (in
<emphasis id="D49F0D482175A55BD16145EC5175DEB1" italics="true" pageNumber="863">Rhodora</emphasis>
11: 91. 1908), and formalised by Reveal &amp; al. (in
<emphasis id="0F2CDCACFEE85D5EBD2E7949F8ADD5D7" italics="true" pageNumber="863">Huntia</emphasis>
7: 238. 1987), and accepted here. Mackenzie (in
<emphasis id="903ACC9FBE45E5784B16FDDC946278B2" italics="true" pageNumber="863">Rhodora</emphasis>
29: 73. 1927) claimed that
<normalizedToken id="0EE1D4A5BCD0742A2F8F237C13865DB9" originalValue="Linnaeus’">Linnaeus'</normalizedToken>
original concept (based on what was in cultivation in Uppsala) was of
<taxonomicName id="FEAA06BF69AA223B2DB4E1593E568F25" authority="Mill." authorityName="Mill. Gray" authorityYear="1768" class="Magnoliopsida" family="Asteraceae" genus="Solidago" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="rugosa">
<emphasis id="EB3CFCD66DE6AC4439184D007B5DFD3E" italics="true" pageNumber="863">S. rugosa</emphasis>
Mill.
</taxonomicName>
and proposed that usage of
<taxonomicName id="81ECA27387F3BA5D49ACB4BE453C0221" authorityName="Linnaeus" authorityYear="1753" baseAuthorityName="Muhl. ex Willd." class="Magnoliopsida" family="Asteraceae" genus="Solidago" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="canadensis">
<emphasis id="ECB5D2CA7A8DAE32CCBCC4894C0B4C83" italics="true" pageNumber="863">S. canadensis</emphasis>
</taxonomicName>
should revert to that. However, Mackenzie did not typify the name to effect this change. Taylor &amp; Taylor (in
<taxonomicName id="F40196CC45B2B2021CB01D3B25BAA4B1" class="Magnoliopsida" family="Malvaceae" genus="Sida" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="D200EF260C0F005A9D6A112F37FEE9B7" italics="true" pageNumber="863">Sida</emphasis>
</taxonomicName>
10: 231. 1984) indicated unspecified material in LINN as
<normalizedToken id="815344FC57883B4403F0E91DA23400D6" originalValue="“type”">&quot;type&quot;</normalizedToken>
but did not distinguish between the several sheets noted by Gray. These collections are not part of a single gathering so Art. 9.15 does not apply.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="58E1CFDC2188DE5161A5D1298E014FD9" type="Zenodo-Dep">233137</mods:identifier>
<mods:identifier id="8A796C3C2D2AC1AD985D7C973414B565" type="ZooBank">CD453319-1BE9-4A9A-B25E-6A70D003512E</mods:identifier>
</mods:mods>
<treatment id="BA12BA2CFFE5FFA851C3235857E3FBFB" ID-DOI="http://doi.org/10.5281/zenodo.6103016" ID-GBIF-Taxon="119609745" ID-Zenodo-Dep="6103016" LSID="urn:lsid:plazi:treatment:BA12BA2CFFE5FFA851C3235857E3FBFB" httpUri="http://treatment.plazi.org/id/BA12BA2CFFE5FFA851C3235857E3FBFB" lastPageId="13" lastPageNumber="249" pageId="9" pageNumber="245">
<subSubSection id="7AA158B1FFE5FFAC51C32358544DFA83" pageId="9" pageNumber="245" type="nomenclature">
<paragraph id="32040B3AFFE5FFAC51C32358571AFAA2" blockId="9.[151,638,1340,1399]" box="[151,638,1340,1366]" pageId="9" pageNumber="245">
<heading id="694CBC56FFE5FFAC51C32358571AFAA2" bold="true" box="[151,638,1340,1366]" fontSize="11" level="1" pageId="9" pageNumber="245" reason="1">
<emphasis id="00CFD728FFE5FFAC51C32358571AFAA2" bold="true" box="[151,638,1340,1366]" pageId="9" pageNumber="245">
<taxonomicName id="F5BB70B9FFE5FFAC51C32358577DFAA2" ID-CoL="4R2GL" box="[151,537,1340,1366]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="245" phylum="Arthropoda" rank="species" species="paululus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFE5FFAC51C323585598FAA2" bold="true" box="[151,252,1340,1366]" italics="true" pageId="9" pageNumber="245">Quedius</emphasis>
(
<emphasis id="00CFD728FFE5FFAC5059235854FAFAA2" bold="true" box="[269,414,1340,1366]" italics="true" pageId="9" pageNumber="245">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE5FFAC50FB2358577DFAA2" bold="true" box="[431,537,1340,1366]" italics="true" pageId="9" pageNumber="245">paululus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE5FFAC53742358571AFAA2" box="[544,638,1340,1366]" pageId="9" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="32040B3AFFE5FFAC51C3233A544DFA83" blockId="9.[151,638,1340,1399]" box="[151,297,1374,1399]" pageId="9" pageNumber="245">
(
<figureCitation id="AA8017BFFFE5FFAC51CB233A5444FA83" box="[159,288,1374,1399]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="9" pageNumber="245">Figs. 4, 4-1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE5FFAC51C323C25758FA17" pageId="9" pageNumber="245" type="materials_examined">
<paragraph id="32040B3AFFE5FFAC51C323C25758FA17" blockId="9.[151,1437,1446,1975]" pageId="9" pageNumber="245">
<emphasis id="00CFD728FFE5FFAC51C323C25423FA4B" bold="true" box="[151,327,1446,1471]" pageId="9" pageNumber="245">
<typeStatus id="ED00B598FFE5FFAC51C323C255B6FA4B" box="[151,210,1446,1471]" pageId="9" pageNumber="245">Type</typeStatus>
material.
</emphasis>
<typeStatus id="ED00B598FFE5FFAC500423C254DCFA4B" box="[336,440,1446,1471]" pageId="9" pageNumber="245" type="holotype">Holotype</typeStatus>
♂,
<emphasis id="00CFD728FFE5FFAC50BC23C257DCFA4B" bold="true" box="[488,696,1446,1471]" pageId="9" pageNumber="245">
<collectingCountry id="4AAC4BAAFFE5FFAC50BC23C2572DFA4B" box="[488,585,1446,1471]" name="China" pageId="9" pageNumber="245">CHINA</collectingCountry>
: Sichuan
</emphasis>
Province: Baoxing County, Guobayan,
<quantity id="F543A6DFFFE5FFAC55D023C351B9FA4A" box="[1156,1245,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="9" pageNumber="245" unit="m" value="3270.0">3270 m</quantity>
,
<date id="46052DFAFFE5FFAC55BF23C350FCFA4B" box="[1259,1432,1447,1471]" pageId="9" pageNumber="245" value="2001-07-01" valueMax="2001-07-04" valueMin="2001-07-01">1–4. VII. 2001</date>
, coll. Xiaodong Yu, Hongzhang Zhou.
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE5FFA85193238A54F4FD5C" lastPageId="13" lastPageNumber="249" pageId="9" pageNumber="245" type="description">
<paragraph id="32040B3AFFE5FFAC5193238A5063F9DF" blockId="9.[151,1437,1446,1975]" pageId="9" pageNumber="245">
<emphasis id="00CFD728FFE5FFAC5193238A543DF9F3" bold="true" box="[199,345,1518,1543]" pageId="9" pageNumber="245">Description.</emphasis>
Head black, pronotum dark brown, both strongly iridescent; scutellum and elytra dark brown; abdomen entirely dark brown; antennae and mouth part dark brown; legs dark brown, tibiae blackish.
</paragraph>
<paragraph id="32040B3AFFE5FFAC519320535021F9BB" blockId="9.[151,1437,1446,1975]" box="[199,1349,1590,1615]" pageId="9" pageNumber="245">
BL =
<quantity id="F543A6DFFFE5FFAC505C20535404F9BA" box="[264,352,1591,1615]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="245" unit="mm" value="7.8">7.8 mm</quantity>
, BW =
<quantity id="F543A6DFFFE5FFAC50E32053576BF9BA" box="[439,527,1591,1615]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="245" unit="mm" value="1.8">1.8 mm</quantity>
, HL/PL/EL = 1.00: 1.38: 1.80, HW/PW/EW/AW = 1.00: 1.34: 1.57: 1.39
</paragraph>
<paragraph id="32040B3AFFE5FFAC5193203E56D6F843" blockId="9.[151,1437,1446,1975]" pageId="9" pageNumber="245">
Head (
<figureCitation id="AA8017BFFFE5FFAC5046203E5431F987" box="[274,341,1626,1651]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="9" pageNumber="245">Fig. 4</figureCitation>
A) round, wider than long, HW/HL = 1.15; eye large and convex, in dorsal view tempora shorter than length of eye, sharply narrowed posteriad, HEL/HTL = 2.24; two additional setiferous punctures between anterior frontal setiferous punctures; posterior frontal setiferous puncture situated distinctly behind the level of posteriomedial margin of eye, separated by distance longer than diameter of puncture; one smaller setiferous puncture between it and temporal setiferous puncture at posterior margin of eye; temporal setiferous puncture situated slightly closer to posterior margin of eye than to nuchal constriction, with several small setiferous punctures arranged in an oblique curve behind it; two basal setiferous punctures situated very close to nuchal constriction; head with very fine and dense microsculpture of transverse waves and irregular meshes. Antenna robust, with segment I longer than segment II or III, segment III distinctly longer than segment II, segments IV–VI, XI slightly longer than wide, segments VII–X slightly wider than long.
</paragraph>
<caption id="66C45BB2FFE6FFAF51C32131501DF87E" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="10" pageNumber="246" targetBox="[179,1418,203,1855]" targetPageId="10">
<paragraph id="32040B3AFFE6FFAF51C32131501DF87E" blockId="10.[151,1436,1877,1930]" pageId="10" pageNumber="246">
<emphasis id="00CFD728FFE6FFAF51C321315472F89E" bold="true" box="[151,278,1877,1899]" pageId="10" pageNumber="246">FIGURE 4.</emphasis>
<taxonomicName id="F5BB70B9FFE6FFAF50742131570FF89F" box="[288,619,1877,1899]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="246" phylum="Arthropoda" rank="species" species="paululus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFE6FFAF507421315410F89F" box="[288,372,1877,1899]" italics="true" pageId="10" pageNumber="246">Quedius</emphasis>
(
<emphasis id="00CFD728FFE6FFAF50D321315766F89F" box="[391,514,1877,1899]" italics="true" pageId="10" pageNumber="246">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE6FFAF53402131570FF89F" box="[532,619,1877,1899]" italics="true" pageId="10" pageNumber="246">paululus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE6FFAF5321213257ADF89F" box="[629,713,1878,1899]" pageId="10" pageNumber="246" rank="species">
<emphasis id="00CFD728FFE6FFAF5321213257ADF89F" bold="true" box="[629,713,1878,1899]" pageId="10" pageNumber="246">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)
</paragraph>
</caption>
<caption id="66C45BB2FFE7FFAE51C321C554A3F800" httpUri="https://zenodo.org/record/233144/files/figure.png" pageId="11" pageNumber="247" targetBox="[164,1422,193,1931]" targetPageId="11">
<paragraph id="32040B3AFFE7FFAE51C321C554A3F800" blockId="11.[151,1436,1953,2036]" pageId="11" pageNumber="247">
<emphasis id="00CFD728FFE7FFAE51C321C5544FF842" bold="true" box="[151,299,1953,1975]" pageId="11" pageNumber="247">FIGURE 4-1.</emphasis>
<taxonomicName id="F5BB70B9FFE7FFAE506121C5571BF843" box="[309,639,1953,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="247" phylum="Arthropoda" rank="species" species="paululus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFE7FFAE506121C554EDF843" box="[309,393,1953,1975]" italics="true" pageId="11" pageNumber="247">Quedius</emphasis>
(
<emphasis id="00CFD728FFE7FFAE50C821C55772F843" box="[412,534,1953,1975]" italics="true" pageId="11" pageNumber="247">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE7FFAE537C21C5571BF843" box="[552,639,1953,1975]" italics="true" pageId="11" pageNumber="247">paululus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE7FFAE53DE21C557BAF842" box="[650,734,1953,1974]" pageId="11" pageNumber="247" rank="species">
<emphasis id="00CFD728FFE7FFAE53DE21C557BAF842" bold="true" box="[650,734,1953,1974]" pageId="11" pageNumber="247">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. male sternite VIII; B. male sternite IX; C. male tergite X; D. aedeagus, lateral view; E. aedeagus, parameral view; F. underside of paramere; G. apical portion of median lobe, parameral view. (Scale bars = 0.20 mm).
</paragraph>
</caption>
<caption id="66C45BB2FFE0FFA951C323A651CFFA03" httpUri="https://zenodo.org/record/233145/files/figure.png" pageId="12" pageNumber="248" targetBox="[151,1436,193,1453]" targetPageId="12">
<paragraph id="32040B3AFFE0FFA951C323A651CFFA03" blockId="12.[151,1436,1474,1527]" pageId="12" pageNumber="248">
<emphasis id="00CFD728FFE0FFA951C323A65471FA2C" bold="true" box="[151,277,1474,1496]" pageId="12" pageNumber="248">FIGURE 5.</emphasis>
Distribution of
<taxonomicName id="F5BB70B9FFE0FFA950EA23A6570CFA2C" box="[446,616,1474,1496]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="elpenor">
<emphasis id="00CFD728FFE0FFA950EA23A6570CFA2C" box="[446,616,1474,1496]" italics="true" pageId="12" pageNumber="248">Quedius elpenor</emphasis>
</taxonomicName>
species. ●
<taxonomicName id="F5BB70B9FFE0FFA953B523A756ECFA2C" box="[737,904,1474,1496]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="biprominulus" status="sp. nov.">
<emphasis id="00CFD728FFE0FFA953B523A756ECFA2C" box="[737,904,1474,1496]" italics="true" pageId="12" pageNumber="248">Q. biprominulus</emphasis>
</taxonomicName>
<emphasis id="00CFD728FFE0FFA952C523A75167FA2C" bold="true" box="[913,1027,1475,1496]" pageId="12" pageNumber="248">
<taxonomicNameLabel id="1BFC6A53FFE0FFA952C523A75680FA2C" box="[913,996,1475,1496]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
; ■
</emphasis>
<taxonomicName id="F5BB70B9FFE0FFA9555823A7511DFA2C" box="[1036,1145,1474,1496]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="elpenor">
<emphasis id="00CFD728FFE0FFA9555823A7511DFA2C" box="[1036,1145,1474,1496]" italics="true" pageId="12" pageNumber="248">Q. elpenor</emphasis>
</taxonomicName>
;
<emphasis id="00CFD728FFE0FFA955DE23A751C7FA2C" bold="true" box="[1162,1187,1475,1496]" pageId="12" pageNumber="248">▲</emphasis>
<taxonomicName id="F5BB70B9FFE0FFA955F823A75065FA2C" box="[1196,1281,1474,1496]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="fusus" status="sp. nov.">
<emphasis id="00CFD728FFE0FFA955F823A75065FA2C" box="[1196,1281,1474,1496]" italics="true" pageId="12" pageNumber="248">Q. fusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE0FFA9545E23A75038FA2C" box="[1290,1372,1475,1496]" pageId="12" pageNumber="248" rank="species">
<emphasis id="00CFD728FFE0FFA9545E23A75038FA2C" bold="true" box="[1290,1372,1475,1496]" pageId="12" pageNumber="248">sp. nov.</emphasis>
</taxonomicNameLabel>
; +
<taxonomicName id="F5BB70B9FFE0FFA954D723A755BBFA03" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="00CFD728FFE0FFA954D723A755BBFA03" italics="true" pageId="12" pageNumber="248">Q. kozlovi</emphasis>
</taxonomicName>
; #
<taxonomicName id="F5BB70B9FFE0FFA951AB23865431FA03" box="[255,341,1505,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="ladas">
<emphasis id="00CFD728FFE0FFA951AB23865431FA03" box="[255,341,1505,1527]" italics="true" pageId="12" pageNumber="248">Q. ladas</emphasis>
</taxonomicName>
; %
<taxonomicName id="F5BB70B9FFE0FFA9502923865499FA03" box="[381,509,1506,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="menippus">
<emphasis id="00CFD728FFE0FFA9502923865499FA03" box="[381,509,1506,1527]" italics="true" pageId="12" pageNumber="248">Q. menippus</emphasis>
</taxonomicName>
; &amp;
<taxonomicName id="F5BB70B9FFE0FFA953712386571FFA03" box="[549,635,1506,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="numa">
<emphasis id="00CFD728FFE0FFA953712386571FFA03" box="[549,635,1506,1527]" italics="true" pageId="12" pageNumber="248">Q. numa</emphasis>
</taxonomicName>
; @
<taxonomicName id="F5BB70B9FFE0FFA953F323865678FA03" box="[679,796,1505,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="paululus" status="sp. nov.">
<emphasis id="00CFD728FFE0FFA953F323865678FA03" box="[679,796,1505,1527]" italics="true" pageId="12" pageNumber="248">Q. paululus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE0FFA9527723855610FA02" box="[803,884,1505,1526]" pageId="12" pageNumber="248" rank="species">
<emphasis id="00CFD728FFE0FFA9527723855610FA02" bold="true" box="[803,884,1505,1526]" pageId="12" pageNumber="248">sp. nov.</emphasis>
</taxonomicNameLabel>
; ¥
<taxonomicName id="F5BB70B9FFE0FFA952C02386516DFA03" box="[916,1033,1505,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="phormio">
<emphasis id="00CFD728FFE0FFA952C02386516DFA03" box="[916,1033,1505,1527]" italics="true" pageId="12" pageNumber="248">Q. phormio</emphasis>
</taxonomicName>
; §
<taxonomicName id="F5BB70B9FFE0FFA9557E238651C2FA03" box="[1066,1190,1505,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="248" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="00CFD728FFE0FFA9557E238651C2FA03" box="[1066,1190,1505,1527]" italics="true" pageId="12" pageNumber="248">Q. tibetanus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="32040B3AFFE0FFA95193207B548CF933" blockId="12.[151,1437,1566,2023]" pageId="12" pageNumber="248">
Pronotum (
<figureCitation id="AA8017BFFFE0FFA95013207A54E8F9C3" box="[327,396,1566,1591]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="12" pageNumber="248">Fig. 4</figureCitation>
B) wider than long, PW/PL = 1.11, slightly narrowed anteriad, posterolateral and posterior margins continuously and broadly rounded, lateral margins not explanate; three setiferous punctures in each dorsal and sublateral row, one extra setiferous puncture between each dorsal and sublateral row, last sublateral row puncture situated behind level of large lateral setiferous puncture; surface of pronotum with microsculpture similar to that of head, but even finer.
</paragraph>
<paragraph id="32040B3AFFE0FFA9519320B65181F91F" blockId="12.[151,1437,1566,2023]" box="[199,1253,1746,1771]" pageId="12" pageNumber="248">
Scutellum (
<figureCitation id="AA8017BFFFE0FFA9501C20B654EFF91F" box="[328,395,1746,1771]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="12" pageNumber="248">Fig. 4</figureCitation>
C) with very fine and dense microsculpture of transverse waves, impunctate.
</paragraph>
<paragraph id="32040B3AFFE0FFA95193209256BEF88F" blockId="12.[151,1437,1566,2023]" pageId="12" pageNumber="248">
Elytra (
<figureCitation id="AA8017BFFFE0FFA9504F2092543AF8FB" box="[283,350,1782,1807]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="12" pageNumber="248">Fig. 4</figureCitation>
C) relatively long, as wide as long, EW/EL = 1.00, ESL/EL = 0.57, slightly broadened posteriad, each elytron with surface evenly covered with moderately dense and coarse setiferous punctures, transverse interspaces between punctures about twice as wide as diameter of punctures, surface between punctures with microsculpture of dense microscopic punctulation. Wings fully developed.
</paragraph>
<paragraph id="32040B3AFFE0FFA9519321E25077F836" blockId="12.[151,1437,1566,2023]" pageId="12" pageNumber="248">Abdominal tergite II impunctate; setiferous punctures of other tergites much finer and denser than those of elytra, almost evenly distributed on each tergite; tergite VII with whitish apical seam of palisade setae.</paragraph>
<paragraph id="32040B3AFFE0FFA8519321AA5663FD70" blockId="12.[151,1437,1566,2023]" lastBlockId="13.[151,1436,151,1040]" lastPageId="13" lastPageNumber="249" pageId="12" pageNumber="248">
Male first four segments of fore tarsus strongly dilated, sub-bilobed, each heavily covered with modified pale setae ventrally, segment II slightly wider than apex of tibia; sternite VII with a very inconspicuous medioapical emargination at apical margin, a very small triangular area around the emargination impunctate; tergite VIII with basal ridge complete, nearly straight, without any long seta; sternite VIII (
<figureCitation id="AA8017BFFFE1FFA8528926BB5150FF0C" box="[989,1076,223,248]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="13" pageNumber="249">Fig. 4-1</figureCitation>
A) with basal ridge very vague, with two long setae on each side, apical margin with a deep and wide medioapical emargination, a relatively large acute triangular area in front of the emargination impunctate; sternite IX (
<figureCitation id="AA8017BFFFE1FFA852B42743515DFEB4" box="[992,1081,295,320]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="13" pageNumber="249">Fig. 4-1</figureCitation>
B) with basal portion wide and short, with two slightly differentiated setae on each side of apical margin; tergite X (
<figureCitation id="AA8017BFFFE1FFA8553A272851ACFE90" box="[1134,1224,332,357]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="13" pageNumber="249">Fig. 4-1</figureCitation>
C) with basal side broadly and shallowly concave, apical margin widely rounded; aedeagus in lateral view (
<figureCitation id="AA8017BFFFE1FFA855D8270B5181FE7C" box="[1164,1253,367,392]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="13" pageNumber="249">Fig. 4-1</figureCitation>
D) with apex of paramere not quite reaching that of median lobe, median lobe with apical 1/4 distinctly attenuate and curved toward paramere, with a short median carina at apex pointing toward paramere side; aedeagus in parameral view (
<figureCitation id="AA8017BFFFE1FFA8540427D355C5FE00" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="13" pageNumber="249">Fig. 4- 1</figureCitation>
E) with paramere moderately narrowed laterally at basal 1/4, and then gradually widened to apical 1/4, apical 1/4 gradually narrowed forming subtriangular apex, lateral margins thinned and widely expanded; median lobe wide at base, then evenly narrowed to acute apex (
<figureCitation id="AA8017BFFFE1FFA853C024405783FDC8" box="[660,743,548,573]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="13" pageNumber="249">Figs. 4</figureCitation>
E, 4-1G); apical portion of paramere with two apical setae clustered at middle and three subapical setae on each side, underside with 19 very small sensory peg setae arranged in two longitudinal groups along sides (
<figureCitation id="AA8017BFFFE1FFA85307240857C5FD70" box="[595,673,620,645]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[179,1418,203,1855]" captionTargetId="figure@10.[164,1422,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Distichalius) paululus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233143/files/figure.png" pageId="13" pageNumber="249">Figs. 4</figureCitation>
D, 4-1F).
</paragraph>
<paragraph id="32040B3AFFE1FFA8519324EB54F4FD5C" blockId="13.[151,1436,151,1040]" box="[199,400,655,680]" pageId="13" pageNumber="249">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE1FFA8519324D05741FD39" box="[199,549,692,717]" pageId="13" pageNumber="249" type="distribution">
<paragraph id="32040B3AFFE1FFA8519324D05741FD39" blockId="13.[151,1436,151,1040]" box="[199,549,692,717]" pageId="13" pageNumber="249">
<emphasis id="00CFD728FFE1FFA8519324D05404FD39" bold="true" box="[199,352,692,717]" pageId="13" pageNumber="249">Distribution.</emphasis>
<collectingCountry id="4AAC4BAAFFE1FFA8503324D054CEFD39" box="[359,426,692,717]" name="China" pageId="13" pageNumber="249">China</collectingCountry>
(Sichuan).
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE1FFA8519324B3577BFC33" pageId="13" pageNumber="249" type="diagnosis">
<paragraph id="32040B3AFFE1FFA8519324B3577BFC33" blockId="13.[151,1436,151,1040]" pageId="13" pageNumber="249">
<emphasis id="00CFD728FFE1FFA8519324B35425FD04" bold="true" box="[199,321,727,752]" pageId="13" pageNumber="249">Diagnosis.</emphasis>
This new species and
<taxonomicName id="F5BB70B9FFE1FFA8530424BC56A0FD04" authority="Smetana" authorityName="Smetana" box="[592,964,728,752]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="249" phylum="Arthropoda" rank="species" species="numa" subGenus="Distichalius">
<emphasis id="00CFD728FFE1FFA8530424BC5708FD1B" box="[592,620,728,751]" italics="true" pageId="13" pageNumber="249">Q.</emphasis>
(
<emphasis id="00CFD728FFE1FFA8532B24BC5662FD04" box="[639,774,728,752]" italics="true" pageId="13" pageNumber="249">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE1FFA8524F24BD563DFD04" box="[795,857,729,752]" italics="true" pageId="13" pageNumber="249">numa</emphasis>
Smetana
</taxonomicName>
are slightly similar in male genitalia by having aedeagus in relatively similar shape and sensory peg setae on underside of paramere in small size, but they can be distinguished by the new species having paramere relatively wider in apical half, distinctly covering apical portion of median lobe except for apex, lateral margins thinned and expanded, and median lobe evenly narrowed to acute apex; whereas
<taxonomicName id="F5BB70B9FFE1FFA850C4250C566DFC74" authority="Smetana" authorityName="Smetana" box="[400,777,872,896]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="249" phylum="Arthropoda" rank="species" species="numa" subGenus="Distichalius">
<emphasis id="00CFD728FFE1FFA850C4250C54CFFC8B" box="[400,427,872,895]" italics="true" pageId="13" pageNumber="249">Q.</emphasis>
(
<emphasis id="00CFD728FFE1FFA85095250C572CFC74" box="[449,584,872,896]" italics="true" pageId="13" pageNumber="249">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE1FFA8530A250D57F8FC74" box="[606,668,873,896]" italics="true" pageId="13" pageNumber="249">numa</emphasis>
Smetana
</taxonomicName>
has paramere relatively narrower in apical half, barely covering apical portion of median lobe, lateral margins not thinned or expanded, and median lobe suddenly narrowed into rod-like, acute apex.
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE1FFA8519325B057E3FBFB" pageId="13" pageNumber="249" type="etymology">
<paragraph id="32040B3AFFE1FFA8519325B057E3FBFB" blockId="13.[151,1436,151,1040]" pageId="13" pageNumber="249">
<emphasis id="00CFD728FFE1FFA8519325B0542AFC19" bold="true" box="[199,334,980,1005]" pageId="13" pageNumber="249">Etymology.</emphasis>
The specific name is from Latin adjective
<taxonomicName id="F5BB70B9FFE1FFA8526725B056F0FC18" box="[819,916,980,1004]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="249" phylum="Arthropoda" rank="species" species="paululus">
<emphasis id="00CFD728FFE1FFA8526725B056F0FC18" box="[819,916,980,1004]" italics="true" pageId="13" pageNumber="249">paululus</emphasis>
</taxonomicName>
(very small), referring to the small size of the sensory peg setae on underside of paramere.
</paragraph>
</subSubSection>
</treatment>
</document>

230
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<heading id="694CBC56FFEAFFA351C324985730FCE2" bold="true" box="[151,596,764,790]" fontSize="11" level="1" pageId="6" pageNumber="242" reason="1">
<emphasis id="00CFD728FFEAFFA351C324985730FCE2" bold="true" box="[151,596,764,790]" pageId="6" pageNumber="242">
<taxonomicName id="F5BB70B9FFEAFFA351C324985494FCE2" ID-CoL="4R2FS" box="[151,496,764,790]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="242" phylum="Arthropoda" rank="species" species="fusus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFEAFFA351C324985598FCE2" bold="true" box="[151,252,764,790]" italics="true" pageId="6" pageNumber="242">Quedius</emphasis>
(
<emphasis id="00CFD728FFEAFFA35059249854FAFCE2" bold="true" box="[269,414,764,790]" italics="true" pageId="6" pageNumber="242">Distichalius</emphasis>
)
<emphasis id="00CFD728FFEAFFA350FB24985494FCE2" bold="true" box="[431,496,764,790]" italics="true" pageId="6" pageNumber="242">fusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFEAFFA350A324985730FCE2" box="[503,596,764,790]" pageId="6" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="32040B3AFFEAFFA351C3257A544DFCC3" blockId="6.[151,596,764,823]" box="[151,297,798,823]" pageId="6" pageNumber="242">
(
<figureCitation id="AA8017BFFFEAFFA351CB257A547AFCC3" box="[159,286,798,823]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="6" pageNumber="242">Figs. 3, 3-1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFEAFFA351C325025019FBFA" pageId="6" pageNumber="242" type="materials_examined">
<paragraph id="32040B3AFFEAFFA351C325025019FBFA" blockId="6.[151,1437,870,1975]" pageId="6" pageNumber="242">
<emphasis id="00CFD728FFEAFFA351C325025422FC8B" bold="true" box="[151,326,870,895]" pageId="6" pageNumber="242">
<typeStatus id="ED00B598FFEAFFA351C3250255B6FC8B" box="[151,210,870,895]" pageId="6" pageNumber="242">Type</typeStatus>
material.
</emphasis>
<typeStatus id="ED00B598FFEAFFA35019250254D1FC8B" box="[333,437,870,895]" pageId="6" pageNumber="242" type="holotype">Holotype</typeStatus>
♂,
<emphasis id="00CFD728FFEAFFA350B6250257C2FC8B" bold="true" box="[482,678,870,895]" pageId="6" pageNumber="242">
<collectingCountry id="4AAC4BAAFFEAFFA350B625025727FC8B" box="[482,579,870,895]" name="China" pageId="6" pageNumber="242">CHINA</collectingCountry>
: Beijing
</emphasis>
City: Mentougou District, Dongling Mountain,
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,
<date id="46052DFAFFEAFFA354262502544CFC57" pageId="6" pageNumber="242" value="2012-09-15" valueMax="2012-09-18" valueMin="2012-09-15">15– 18. IX. 2012</date>
, coll. Xiaodong Yu.
<typeStatus id="ED00B598FFEAFFA3535A25EF57E4FC57" box="[526,640,907,931]" pageId="6" pageNumber="242" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="4AAC4BAAFFEAFFA353DE25EE5788FC57" box="[650,748,906,931]" name="China" pageId="6" pageNumber="242">CHINA</collectingCountry>
: Beijing
</emphasis>
City:
<specimenCount id="24BDC0B3FFEAFFA352C025EF56A4FC56" box="[916,960,907,931]" pageId="6" pageNumber="242" type="generic">1 ♂</specimenCount>
, Mentougou District, Xiaolongmen Forest Park,
<quantity id="F543A6DFFFEAFFA3518D25CB5455FC32" box="[217,305,943,967]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="242" unit="m" value="1800.0">1800 m</quantity>
,
<date id="46052DFAFFEAFFA3506925CB54B1FC33" box="[317,469,943,967]" pageId="6" pageNumber="242" value="1993-08-07">7. VIII. 1993</date>
, coll. Hongzhang Zhou; 1 ♀, Mentougou District, Xiaolongmen Forest Park,
<date id="46052DFAFFEAFFA3541725CB55B6FC1F" pageId="6" pageNumber="242" value="1997-07-22">22. VII. 1997</date>
, coll. Haisheng Zhou;
<specimenCount id="24BDC0B3FFEAFFA3508125B75766FC1E" box="[469,514,979,1003]" pageId="6" pageNumber="242" type="generic">1 ♂</specimenCount>
, 1 ♀, Mentougou District, Dongling Mountain,
<quantity id="F543A6DFFFEAFFA3557225B651A0FC1E" box="[1062,1220,978,1003]" metricMagnitude="3" metricUnit="m" metricValue="1.3995" metricValueMax="1.509" metricValueMin="1.29" pageId="6" pageNumber="242" unit="m" value="1399.5" valueMax="1509.0" valueMin="1290.0">1290–1509 m</quantity>
,
<date id="46052DFAFFEAFFA3558625B650FCFC1F" box="[1234,1432,978,1003]" pageId="6" pageNumber="242" value="2012-07-15" valueMax="2012-07-18" valueMin="2012-07-15">15–18. VII. 2012</date>
, coll. Xiaodong Yu; 1 ♀, Mentougou District, Dongling Mountain,
<quantity id="F543A6DFFFEAFFA352D4259256B3FBFA" box="[896,983,1014,1038]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="6" pageNumber="242" unit="m" value="1225.0">1225 m</quantity>
,
<date id="46052DFAFFEAFFA352B7259351F9FBFB" box="[995,1181,1015,1039]" pageId="6" pageNumber="242" value="2012-09-14" valueMax="2012-09-17" valueMin="2012-09-14">14–17. IX. 2012</date>
, coll. Xiaodong Yu.
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFEAFFAC5193227E559EFC3C" lastPageId="9" lastPageNumber="245" pageId="6" pageNumber="242" type="description">
<paragraph id="32040B3AFFEAFFA35193227E5139FB6B" blockId="6.[151,1437,870,1975]" pageId="6" pageNumber="242">
<emphasis id="00CFD728FFEAFFA35193227E543DFBC7" bold="true" box="[199,345,1050,1075]" pageId="6" pageNumber="242">Description.</emphasis>
Head black; pronotum and scutellum dark brown; elytra dark brown, some with humeral angle, suture and posterior margin slightly paler; abdomen dark brown, each tergite with posterior margin distinctly paler; antennae dark brown, with basal half of antennomere II yellowish brown, labrum and mandibles dark reddish brown, maxillary and labial palpi dark brown; legs dark brown, tibiae blackish brown.
</paragraph>
<paragraph id="32040B3AFFEAFFA3519322CF5046FB37" blockId="6.[151,1437,870,1975]" box="[199,1314,1194,1219]" pageId="6" pageNumber="242">
BL =
<quantity id="F543A6DFFFEAFFA3505C22CE5404FB36" box="[264,352,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="242" unit="mm" value="7.5">7.5 mm</quantity>
, BW =
<quantity id="F543A6DFFFEAFFA350E322CE576BFB36" box="[439,527,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="242" unit="mm" value="1.6">1.6 mm</quantity>
, HL/PL/EL = 1.00:1.26:1.50, HW/PW/EW/AW = 1.00:1.25:1.47:1.42
</paragraph>
<paragraph id="32040B3AFFEAFFA3519322AA542AF9BB" blockId="6.[151,1437,870,1975]" pageId="6" pageNumber="242">
Head (
<figureCitation id="AA8017BFFFEAFFA3504522AA5436FB13" box="[273,338,1230,1255]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="6" pageNumber="242">Fig. 3</figureCitation>
A) round, about as wide as long, HW/HL = 1.05; eye moderately large and convex, in dorsal view tempora shorter than length of eye, gradually narrowed posteriad, HEL/HTL = 2.02; two additional setiferous punctures between anterior frontal setiferous punctures; posterior frontal setiferous puncture situated slightly behind the level of posteriomedial margin of eye, separated by distance about twice as long as diameter of puncture; one smaller setiferous puncture between it and temporal setiferous puncture at posterior margin of eye; temporal setiferous puncture situated in midway between posterior margin of eye and nuchal constriction of head, with several small setiferous punctures arranged in an oblique curve below it; two basal setiferous punctures situated much closer to nuchal constriction than to posterior frontal setiferous puncture; head with fine and dense microsculpture of transverse waves and irregular meshes. Antenna robust, with segment I longer than segment II or III, segment III slightly longer than segment II, segments IV, XI slightly longer than wide, segments V–X slightly wider than long.
</paragraph>
<paragraph id="32040B3AFFEAFFA35193203F57D0F8F6" blockId="6.[151,1437,870,1975]" pageId="6" pageNumber="242">
Pronotum (
<figureCitation id="AA8017BFFFEAFFA3501C203E54E9F987" box="[328,397,1626,1651]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="6" pageNumber="242">Fig. 3</figureCitation>
B) about as wide as long, PW/PL = 1.05, narrowed anteriad, posterolateral and posterior margins continuously and broadly rounded, lateral margins not explanate; three setiferous punctures in each dorsal and sublateral row, one extra setiferous puncture between each dorsal and sublateral row, last sublateral row puncture situated behind level of large lateral setiferous puncture; surface of pronotum with very fine and dense microsculpture of transverse and oblique waves.
</paragraph>
<paragraph id="32040B3AFFEAFFA35193216A5181F8D3" blockId="6.[151,1437,870,1975]" box="[199,1253,1806,1831]" pageId="6" pageNumber="242">
Scutellum (
<figureCitation id="AA8017BFFFEAFFA3501C216A54EDF8D3" box="[328,393,1806,1831]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="6" pageNumber="242">Fig. 3</figureCitation>
C) with very fine and dense microsculpture of transverse waves, impunctate.
</paragraph>
<paragraph id="32040B3AFFEAFFA3519321565736F843" blockId="6.[151,1437,870,1975]" pageId="6" pageNumber="242">
Elytra (
<figureCitation id="AA8017BFFFEAFFA350482156543BF8BF" box="[284,351,1842,1867]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="6" pageNumber="242">Fig. 3</figureCitation>
C) moderately long, about as wide as long, EW/EL = 1.03, ESL/EL = 0.52, slightly broadened posteriad, each elytron with surface covered with moderately dense and coarse setiferous punctures, transverse interspaces between punctures about twice as wide as diameter of punctures, surface between punctures without microsculpture. Wings fully developed.
</paragraph>
<caption id="66C45BB2FFEBFFA251C321315074F87E" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="7" pageNumber="243" targetBox="[177,1417,210,1855]" targetPageId="7">
<paragraph id="32040B3AFFEBFFA251C321315074F87E" blockId="7.[151,1435,1877,1930]" pageId="7" pageNumber="243">
<emphasis id="00CFD728FFEBFFA251C321315476F89E" bold="true" box="[151,274,1877,1899]" pageId="7" pageNumber="243">FIGURE 3.</emphasis>
<taxonomicName id="F5BB70B9FFEBFFA2504D2131575DF89F" box="[281,569,1877,1899]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="243" phylum="Arthropoda" rank="species" species="fusus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFEBFFA2504D21315409F89F" box="[281,365,1877,1899]" italics="true" pageId="7" pageNumber="243">Quedius</emphasis>
(
<emphasis id="00CFD728FFEBFFA2502821315492F89F" box="[380,502,1877,1899]" italics="true" pageId="7" pageNumber="243">Distichalius</emphasis>
)
<emphasis id="00CFD728FFEBFFA253512131575DF89F" box="[517,569,1877,1899]" italics="true" pageId="7" pageNumber="243">fusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFEBFFA25314213257F4F89F" box="[576,656,1878,1899]" pageId="7" pageNumber="243" rank="species">
<emphasis id="00CFD728FFEBFFA25314213257F4F89F" bold="true" box="[576,656,1878,1899]" pageId="7" pageNumber="243">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)
</paragraph>
</caption>
<caption id="66C45BB2FFE4FFAD51C321C55758F800" httpUri="https://zenodo.org/record/233142/files/figure.png" pageId="8" pageNumber="244" targetBox="[196,1391,193,1931]" targetPageId="8">
<paragraph id="32040B3AFFE4FFAD51C321C55758F800" blockId="8.[151,1436,1953,2036]" pageId="8" pageNumber="244">
<emphasis id="00CFD728FFE4FFAD51C321C55443F842" bold="true" box="[151,295,1953,1975]" pageId="8" pageNumber="244">FIGURE 3-1.</emphasis>
<taxonomicName id="F5BB70B9FFE4FFAD507A21C5572AF843" box="[302,590,1953,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="244" phylum="Arthropoda" rank="species" species="fusus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFE4FFAD507A21C554E6F843" box="[302,386,1953,1975]" italics="true" pageId="8" pageNumber="244">Quedius</emphasis>
(
<emphasis id="00CFD728FFE4FFAD50C521C5576FF843" box="[401,523,1953,1975]" italics="true" pageId="8" pageNumber="244">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE4FFAD534E21C5572AF843" box="[538,590,1953,1975]" italics="true" pageId="8" pageNumber="244">fusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE4FFAD530021C557C0F842" box="[596,676,1953,1974]" pageId="8" pageNumber="244" rank="species">
<emphasis id="00CFD728FFE4FFAD530021C557C0F842" bold="true" box="[596,676,1953,1974]" pageId="8" pageNumber="244">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. male sternite VIII; B. male sternite IX; C. male tergite X; D. underside of paramere; E. apical portion of median lobe, parameral view; F. aedeagus, lateral view; G. aedeagus, parameral view; H. female tergite X. (Scale bars = 0.20 mm).
</paragraph>
</caption>
<paragraph id="32040B3AFFE5FFAC519326F35602FF03" blockId="9.[151,1436,151,1256]" pageId="9" pageNumber="245">Abdominal tergite II impunctate; setiferous punctures of other tergites finer than those of elytra, becoming slightly sparser toward posterior margin of each tergite, and generally becoming slightly so toward apex of abdomen; tergite VII with whitish apical seam of palisade setae.</paragraph>
<paragraph id="32040B3AFFE5FFAC5193276050E3FCE0" blockId="9.[151,1436,151,1256]" pageId="9" pageNumber="245">
Male first four segments of fore tarsus strongly dilated, sub-bilobed, each heavily covered with modified pale setae ventrally, segment II slightly wider than apex of tibia; sternite VII with very inconspicuous medioapical emargination at apical margin, a small acute triangular area in front of the emargination impunctate; tergite VIII with basal ridge complete, nearly straight, with one long seta on each side; sternite VIII (
<figureCitation id="AA8017BFFFE5FFAC55F7270B519FFE7C" box="[1187,1275,367,392]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Fig. 3-1</figureCitation>
A) with basal ridge complete, nearly straight, with two long setae on each side, apical margin with a deep and wide medioapical emargination, a relatively large acute triangular area in front of the emargination impunctate; sternite IX (
<figureCitation id="AA8017BFFFE5FFAC541B27D355C6FE00" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Fig. 3- 1</figureCitation>
B) with basal portion long and narrow, strongly curved, with a very shallow middle emargination apically; tergite X (
<figureCitation id="AA8017BFFFE5FFAC51E8279B5470FDEC" box="[188,276,511,536]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Fig. 3-1</figureCitation>
C) with basal side broadly and shallowly concave, apical margin widely rounded; aedeagus in lateral view (
<figureCitation id="AA8017BFFFE5FFAC51B42440545EFDC8" box="[224,314,548,573]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Fig. 3-1</figureCitation>
F) with apex of paramere distinctly protruding beyond that of median lobe, median lobe with a minute hook-shaped process at apex and an inconspicuous narrow median carina along midline from subapex to apical 1/4, both pointing toward paramere side; aedeagus in parameral view (
<figureCitation id="AA8017BFFFE5FFAC55452408510CFD70" box="[1041,1128,620,645]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Fig. 3-1</figureCitation>
G) with paramere strongly narrowed laterally in middle, with apical half fusiform, rounded at apex, covering apical 1/3 of median lobe; median lobe wide at base, then gradually narrowed to basal 2/3, apical 1/3 nearly parallel-sided, with tiny subacute apex (
<figureCitation id="AA8017BFFFE5FFAC518924B3544EFD04" box="[221,298,727,752]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Figs. 3</figureCitation>
E, 3-1E); apical portion of paramere with four apical setae and two to three subapical setae on each side, underside with numerous sensory peg setae arranged in a longitudinal band along midline (
<figureCitation id="AA8017BFFFE5FFAC559A2498507EFCE0" box="[1230,1306,764,789]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Figs. 3</figureCitation>
D, 3-1D).
</paragraph>
<paragraph id="32040B3AFFE5FFAC5193257B559EFC3C" blockId="9.[151,1436,151,1256]" pageId="9" pageNumber="245">
Female first four segments of fore tarsus similar to those of male, but less dilated, segment II about as wide as apex of tibia; tergite VIII with basal ridge complete, nearly straight, with one long seta on each side; sternite VIII with basal ridge complete, with two long setae on each side; tergite X (
<figureCitation id="AA8017BFFFE5FFAC528D25035155FC74" box="[985,1073,871,896]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[177,1417,210,1855]" captionTargetId="figure@7.[166,1420,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Quedius (Distichalius) fusus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233141/files/figure.png" pageId="9" pageNumber="245">Fig. 3-1</figureCitation>
H) with basal side broadly and shallowly concave, with subtriangular area in middle distinctly raised and strongly pigmented, apical margin rounded.
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE5FFAC519325B0577AFC19" box="[199,542,980,1005]" pageId="9" pageNumber="245" type="distribution">
<paragraph id="32040B3AFFE5FFAC519325B0577AFC19" blockId="9.[151,1436,151,1256]" box="[199,542,980,1005]" pageId="9" pageNumber="245">
<emphasis id="00CFD728FFE5FFAC519325B05404FC19" bold="true" box="[199,352,980,1005]" pageId="9" pageNumber="245">Distribution.</emphasis>
<collectingCountry id="4AAC4BAAFFE5FFAC503325B054CEFC19" box="[359,426,980,1005]" name="China" pageId="9" pageNumber="245">China</collectingCountry>
(Beijing).
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE5FFAC5193259351BFFB54" pageId="9" pageNumber="245" type="diagnosis">
<paragraph id="32040B3AFFE5FFAC5193259351BFFB54" blockId="9.[151,1436,151,1256]" pageId="9" pageNumber="245">
<emphasis id="00CFD728FFE5FFAC519325935425FBE4" bold="true" box="[199,321,1015,1040]" pageId="9" pageNumber="245">Diagnosis.</emphasis>
This new species is the only one without microscopic punctulation on elytral surface in
<taxonomicName id="F5BB70B9FFE5FFAC5412259C50FFFBE4" box="[1350,1435,1016,1040]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="245" phylum="Arthropoda" rank="species" species="elpenor">
<emphasis id="00CFD728FFE5FFAC5412259C50FFFBE4" box="[1350,1435,1016,1040]" italics="true" pageId="9" pageNumber="245">elpenor</emphasis>
</taxonomicName>
group, but the presence of the small setiferous puncture at posterior margin of eye on head and the extra setiferous puncture between dorsal and sublateral row on pronotum, and the shape of the aedeagus show that it is doubtlessly a member of
<taxonomicName id="F5BB70B9FFE5FFAC5066220054E3FB88" box="[306,391,1124,1148]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="245" phylum="Arthropoda" rank="species" species="elpenor">
<emphasis id="00CFD728FFE5FFAC5066220054E3FB88" box="[306,391,1124,1148]" italics="true" pageId="9" pageNumber="245">elpenor</emphasis>
</taxonomicName>
group. It can be distinguished from other similar species easily by the unique shape of the fusiform paramere and the distinct banded pattern of sensory peg setae on underside of paramere.
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFE5FFAC519322C854C5FB1C" pageId="9" pageNumber="245" type="etymology">
<paragraph id="32040B3AFFE5FFAC519322C854C5FB1C" blockId="9.[151,1436,151,1256]" pageId="9" pageNumber="245">
<emphasis id="00CFD728FFE5FFAC519322C8542AFB31" bold="true" box="[199,334,1196,1221]" pageId="9" pageNumber="245">Etymology.</emphasis>
The specific name is from Latin noun
<taxonomicName id="F5BB70B9FFE5FFAC524C22C85636FB30" box="[792,850,1196,1220]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="245" phylum="Arthropoda" rank="species" species="fusus">
<emphasis id="00CFD728FFE5FFAC524C22C85636FB30" box="[792,850,1196,1220]" italics="true" pageId="9" pageNumber="245">fusus</emphasis>
</taxonomicName>
(a spindle) in apposition, referring to the fusiform apical half of paramere.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,213 @@
<document id="60E9E188DE5B640FAB8E7F58A5580621" ID-DOI="10.11646/zootaxa.3947.2.6" ID-GBIF-Dataset="51c3606a-2061-4a48-b716-7be8fa053ec1" ID-ISSN="1175-5326" ID-Zenodo-Dep="233137" ID-ZooBank="CD453319-1BE9-4A9A-B25E-6A70D003512E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461040712641" checkinUser="plazi" docAuthor="Cai, Yan-Peng &amp; Zhou, Hong-Zhang" docDate="2015" docId="BA12BA2CFFEFFFA351C327D5519CFD5C" docLanguage="en" docName="zt03947p250.pdf" docOrigin="Zootaxa 3947 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius (Distichalius) biprominulus Cai &amp; Zhou, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="242" masterDocId="462BC254FFECFFA5515426645564FFF4" masterDocTitle="Three new species of Quedius elpenor group (Coleoptera: Staphylinidae: Staphylinini: Quediina) from China" masterLastPageNumber="250" masterPageNumber="236" pageNumber="239" updateTime="1698575494253" updateUser="plazi">
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<mods:title id="3C87587153756114EBAB4D007E0D4226">Three new species of Quedius elpenor group (Coleoptera: Staphylinidae: Staphylinini: Quediina) from China</mods:title>
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<mods:namePart id="346CC7376F820ED60F4DCDE55C8EEBAB">Cai, Yan-Peng</mods:namePart>
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<mods:namePart id="1B6B592831068F6DC223475234D30482">Zhou, Hong-Zhang</mods:namePart>
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<treatment id="BA12BA2CFFEFFFA351C327D5519CFD5C" ID-DOI="http://doi.org/10.5281/zenodo.6103012" ID-GBIF-Taxon="119609743" ID-Zenodo-Dep="6103012" LSID="urn:lsid:plazi:treatment:BA12BA2CFFEFFFA351C327D5519CFD5C" httpUri="http://treatment.plazi.org/id/BA12BA2CFFEFFFA351C327D5519CFD5C" lastPageId="6" lastPageNumber="242" pageId="3" pageNumber="239">
<subSubSection id="7AA158B1FFEFFFA651C327D5544DFE18" pageId="3" pageNumber="239" type="nomenclature">
<paragraph id="32040B3AFFEFFFA651C327D557D3FE3F" blockId="3.[151,695,433,493]" box="[151,695,433,459]" pageId="3" pageNumber="239">
<heading id="694CBC56FFEFFFA651C327D557D3FE3F" bold="true" box="[151,695,433,459]" fontSize="11" level="1" pageId="3" pageNumber="239" reason="1">
<emphasis id="00CFD728FFEFFFA651C327D557D3FE3F" bold="true" box="[151,695,433,459]" pageId="3" pageNumber="239">
<taxonomicName id="F5BB70B9FFEFFFA651C327D55736FE3F" ID-CoL="4R2FD" box="[151,594,433,459]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="239" phylum="Arthropoda" rank="species" species="biprominulus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFEFFFA651C327D55598FE3F" bold="true" box="[151,252,433,459]" italics="true" pageId="3" pageNumber="239">Quedius</emphasis>
(
<emphasis id="00CFD728FFEFFFA6505927D554FAFE3F" bold="true" box="[269,414,433,459]" italics="true" pageId="3" pageNumber="239">Distichalius</emphasis>
)
<emphasis id="00CFD728FFEFFFA650FB27D55736FE3F" bold="true" box="[431,594,433,459]" italics="true" pageId="3" pageNumber="239">biprominulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFEFFFA6530D27D557D3FE3F" box="[601,695,433,459]" pageId="3" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="32040B3AFFEFFFA651C327B0544DFE18" blockId="3.[151,695,433,493]" box="[151,297,468,493]" pageId="3" pageNumber="239">
(
<figureCitation id="AA8017BFFFEFFFA651CB27B0547BFE18" box="[159,287,468,493]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Figs. 2, 2-1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFEFFFA651C32478575AFDAC" pageId="3" pageNumber="239" type="materials_examined">
<paragraph id="32040B3AFFEFFFA651C32478575AFDAC" blockId="3.[151,1437,540,2005]" pageId="3" pageNumber="239">
<emphasis id="00CFD728FFEFFFA651C324785420FDC1" bold="true" box="[151,324,540,565]" pageId="3" pageNumber="239">
<typeStatus id="ED00B598FFEFFFA651C3247855B6FDC1" box="[151,210,540,565]" pageId="3" pageNumber="239">Type</typeStatus>
material.
</emphasis>
<typeStatus id="ED00B598FFEFFFA6501E247854D6FDC1" box="[330,434,540,565]" pageId="3" pageNumber="239" type="holotype">Holotype</typeStatus>
♂,
<emphasis id="00CFD728FFEFFFA65088247857C0FDC1" bold="true" box="[476,676,540,565]" pageId="3" pageNumber="239">
<collectingCountry id="4AAC4BAAFFEFFFA6508824785759FDC1" box="[476,573,540,565]" name="China" pageId="3" pageNumber="239">CHINA</collectingCountry>
: Yunnan
</emphasis>
Province: Zhongdian County, Bitahai Nature Reserve,
<quantity id="F543A6DFFFEFFFA6544424785003FDC0" box="[1296,1383,540,564]" metricMagnitude="3" metricUnit="m" metricValue="3.003" pageId="3" pageNumber="239" unit="m" value="3003.0">3003 m</quantity>
,
<date id="46052DFAFFEFFFA6542624785455FDAC" pageId="3" pageNumber="239" value="2000-07-26" valueMax="2000-07-30" valueMin="2000-07-26">26– 30. VII. 2000</date>
, coll. Hongzhang Zhou.
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFEFFFA35193240054F4FEE9" lastPageId="6" lastPageNumber="242" pageId="3" pageNumber="239" type="description">
<paragraph id="32040B3AFFEFFFA651932400541AFD31" blockId="3.[151,1437,540,2005]" pageId="3" pageNumber="239">
<emphasis id="00CFD728FFEFFFA651932400543DFD89" bold="true" box="[199,345,612,637]" pageId="3" pageNumber="239">Description.</emphasis>
Head and pronotum black; scutellum blackish brown, elytra blackish brown, with inconspicuous dark green metallic hue; abdomen dark brown, each tergite with posterior margin paler; antennae, mouth part and legs blackish brown.
</paragraph>
<paragraph id="32040B3AFFEFFFA6519324B45037FD1C" blockId="3.[151,1437,540,2005]" box="[199,1363,719,744]" pageId="3" pageNumber="239">
BL =
<quantity id="F543A6DFFFEFFFA6505C24B4540AFD13" box="[264,366,720,744]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="3" pageNumber="239" unit="mm" value="10.7">10.7 mm</quantity>
, BW =
<quantity id="F543A6DFFFEFFFA6509124B45779FD13" box="[453,541,720,744]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="239" unit="mm" value="2.4">2.4 mm</quantity>
, HL/PL/EL = 1.00: 1.29: 1.64, HW/PW/EW/AW = 1.00: 1.32: 1.57: 1.41
</paragraph>
<paragraph id="32040B3AFFEFFFA65193249056B2FBA4" blockId="3.[151,1437,540,2005]" pageId="3" pageNumber="239">
Head (
<figureCitation id="AA8017BFFFEFFFA650472490543CFCF8" box="[275,344,756,781]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2</figureCitation>
A) round, slightly wider than long, HW/HL = 1.09; eye moderately large and convex, in dorsal view tempora shorter than length of eye, gradually narrowed posteriad, HEL/HTL = 1.66; two additional setiferous punctures between anterior frontal setiferous punctures; posterior frontal setiferous puncture situated distinctly behind level of posteriomedial margin of eye, closer to posterior medial margin of eye than to nuchal constriction of head; one smaller setiferous puncture between it and temporal setiferous puncture at posterior margin of eye; temporal setiferous puncture situated about midway between posterior margin of eye and nuchal constriction, with several small setiferous punctures arranged in an oblique curve behind it; two basal setiferous punctures situated very close to nuchal constriction; head with very fine and dense microsculpture of transverse and oblique waves. Antenna somewhat robust, with segment I longer than segment II or III, segment III longer than segment II, segments IV–VII, XI longer than wide, segments VIII–X wider than long.
</paragraph>
<paragraph id="32040B3AFFEFFFA651932239545BFAF1" blockId="3.[151,1437,540,2005]" pageId="3" pageNumber="239">
Pronotum (
<figureCitation id="AA8017BFFFEFFFA65012223854EFFB80" box="[326,395,1116,1141]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2</figureCitation>
B) wider than long, PW/PL = 1.12, narrowed anteriad, posterolateral and posterior margins continuously and broadly rounded, lateral margins not explanate; three setiferous punctures in each dorsal and sublateral row, one small extra setiferous puncture between each dorsal and sublateral row, last sublateral row puncture situated behind level of large lateral setiferous puncture; surface of pronotum with microsculpture similar to that of head.
</paragraph>
<paragraph id="32040B3AFFEFFFA65193236B5181FADC" blockId="3.[151,1437,540,2005]" box="[199,1253,1295,1321]" pageId="3" pageNumber="239">
Scutellum (
<figureCitation id="AA8017BFFFEFFFA6501C236B54EEFADC" box="[328,394,1295,1320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2</figureCitation>
C) with very fine and dense microsculpture of transverse waves, impunctate.
</paragraph>
<paragraph id="32040B3AFFEFFFA65193235056BEFA4C" blockId="3.[151,1437,540,2005]" pageId="3" pageNumber="239">
Elytra (
<figureCitation id="AA8017BFFFEFFFA6504923505407FAB8" box="[285,355,1332,1357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2</figureCitation>
C) relatively long, about as wide as long, EW/EL = 1.04, ESL/EL = 0.56, slightly broadened posteriad, each elytron with surface unevenly covered with sparse coarse setiferous punctures, somewhat arranged in longitudinal rows and leaving relatively large areas of elytron impunctate, surface between punctures with microsculpture of dense microscopic punctulation. Wings fully developed.
</paragraph>
<paragraph id="32040B3AFFEFFFA6519323A05602F9D0" blockId="3.[151,1437,540,2005]" pageId="3" pageNumber="239">Abdominal tergite II impunctate; setiferous punctures of other tergites much finer and denser than those of elytra, becoming sparser toward posterior margin of each tergite, and generally becoming so toward apex of abdomen; tergite VII with whitish apical seam of palisade setae.</paragraph>
<paragraph id="32040B3AFFEFFFA35193204B5151FF0C" blockId="3.[151,1437,540,2005]" lastBlockId="6.[151,1437,151,680]" lastPageId="6" lastPageNumber="242" pageId="3" pageNumber="239">
Male first four segments of fore tarsus strongly dilated, sub-bilobed, each heavily covered with modified pale setae ventrally, segment II slightly wider than apex of tibia; sternite VII with a very inconspicuous medioapical emargination at apical margin, a very small triangular area in front of the emargination impunctate; tergite VIII with basal ridge complete, straight, without any long seta; sternite VIII (
<figureCitation id="AA8017BFFFEFFFA652B120F8515BF940" box="[997,1087,1692,1717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2-1</figureCitation>
A) with basal ridge complete, nearly straight, with two long setae on each side, apical margin with a deep and wide medioapical emargination, a relatively large acute triangular area in front of the emargination impunctate; sternite IX (
<figureCitation id="AA8017BFFFEFFFA655F02080519BF908" box="[1188,1279,1764,1789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2-1</figureCitation>
B) with basal portion rather wide, with one long seta on each side, apical margin rounded; tergite X (
<figureCitation id="AA8017BFFFEFFFA65520216351A8F8D4" box="[1140,1228,1799,1824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2-1</figureCitation>
C) with basal side broadly and shallowly concave, apical margin widely rounded; aedeagus in lateral view (
<figureCitation id="AA8017BFFFEFFFA655D821485180F8B0" box="[1164,1252,1836,1861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2-1</figureCitation>
D) with apex of paramere not quite reaching that of median lobe, median lobe swollen at about apical 1/4 toward paramere side, with two small hook-shaped processes in characteristic shapes at apex pointing toward paramere side (
<figureCitation id="AA8017BFFFEFFFA654612110501FF878" box="[1333,1403,1908,1933]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2</figureCitation>
E); aedeagus in parameral view (
<figureCitation id="AA8017BFFFEFFFA650B621FC575EF844" box="[482,570,1944,1969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="3" pageNumber="239">Fig. 2-1</figureCitation>
E) with paramere gradually narrowed in general from base to apex, but with middle third inconspicuously dilated laterally, being as wide as median lobe at widest point, apex bluntly rounded; median lobe wide at base, then evenly narrowed, and abruptly constricted into small and narrow apex (
<figureCitation id="AA8017BFFFEAFFA3544F26F3500DFF44" box="[1307,1385,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="6" pageNumber="242">Figs. 2</figureCitation>
F, 2- 1G); apical portion of paramere with two long setae at apex and three subapical setae on each side, underside with 28 sensory peg setae arranged in two parallel longitudinal groups (
<figureCitation id="AA8017BFFFEAFFA352D426BB56ABFF0C" box="[896,975,223,248]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1432,195,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Distichalius) biprominulus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="6" pageNumber="242">Figs. 2</figureCitation>
D, 2-1F).
</paragraph>
<caption id="66C45BB2FFE8FFA151C3213151A4F85C" httpUri="https://zenodo.org/record/233139/files/figure.png" pageId="4" pageNumber="240" targetBox="[151,1432,195,1855]" targetPageId="4">
<paragraph id="32040B3AFFE8FFA151C3213151A4F85C" blockId="4.[151,1436,1877,1960]" pageId="4" pageNumber="240">
<emphasis id="00CFD728FFE8FFA151C321315470F89E" bold="true" box="[151,276,1877,1899]" pageId="4" pageNumber="240">FIGURE 2.</emphasis>
<taxonomicName id="F5BB70B9FFE8FFA1504F213157F4F89F" box="[283,656,1877,1899]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="240" phylum="Arthropoda" rank="species" species="biprominulus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFE8FFA1504F2131540BF89F" box="[283,367,1877,1899]" italics="true" pageId="4" pageNumber="240">Quedius</emphasis>
(
<emphasis id="00CFD728FFE8FFA150D42131549EF89F" box="[384,506,1877,1899]" italics="true" pageId="4" pageNumber="240">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE8FFA1535E213157F4F89F" box="[522,656,1877,1899]" italics="true" pageId="4" pageNumber="240">biprominulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE8FFA153CC2132578DF89F" box="[664,745,1878,1899]" pageId="4" pageNumber="240" rank="species">
<emphasis id="00CFD728FFE8FFA153CC2132578DF89F" bold="true" box="[664,745,1878,1899]" pageId="4" pageNumber="240">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, lateral view (0.2 mm); F. apical portion of median lobe, parameral view (0.2 mm); G. microscopic punctulation on elytra (0.2 mm). (Scale lengths in parentheses)
</paragraph>
</caption>
<caption id="66C45BB2FFE9FFA051C321C554A3F800" httpUri="https://zenodo.org/record/233140/files/figure.png" pageId="5" pageNumber="241" targetBox="[167,1419,193,1931]" targetPageId="5">
<paragraph id="32040B3AFFE9FFA051C321C554A3F800" blockId="5.[151,1436,1953,2036]" pageId="5" pageNumber="241">
<emphasis id="00CFD728FFE9FFA051C321C5544CF842" bold="true" box="[151,296,1953,1975]" pageId="5" pageNumber="241">FIGURE 2-1.</emphasis>
<taxonomicName id="F5BB70B9FFE9FFA0506421C557C1F843" box="[304,677,1953,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="241" phylum="Arthropoda" rank="species" species="biprominulus" status="sp. nov." subGenus="Distichalius">
<emphasis id="00CFD728FFE9FFA0506421C554E0F843" box="[304,388,1953,1975]" italics="true" pageId="5" pageNumber="241">Quedius</emphasis>
(
<emphasis id="00CFD728FFE9FFA050C021C5576AF843" box="[404,526,1953,1975]" italics="true" pageId="5" pageNumber="241">Distichalius</emphasis>
)
<emphasis id="00CFD728FFE9FFA0534B21C557C1F843" box="[543,677,1953,1975]" italics="true" pageId="5" pageNumber="241">biprominulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFC6A53FFE9FFA053F821C5579AF842" box="[684,766,1953,1974]" pageId="5" pageNumber="241" rank="species">
<emphasis id="00CFD728FFE9FFA053F821C5579AF842" bold="true" box="[684,766,1953,1974]" pageId="5" pageNumber="241">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. male sternite VIII; B. male sternite IX; C. male tergite X; D. aedeagus, lateral view; E. aedeagus, parameral view; F. underside of paramere; G. apical portion of median lobe, parameral view. (Scale bars = 0.20 mm).
</paragraph>
</caption>
<paragraph id="32040B3AFFEAFFA35193276054F4FEE9" blockId="6.[151,1437,151,680]" box="[199,400,260,285]" pageId="6" pageNumber="242">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFEAFFA3519327435745FEB4" box="[199,545,295,320]" pageId="6" pageNumber="242" type="distribution">
<paragraph id="32040B3AFFEAFFA3519327435745FEB4" blockId="6.[151,1437,151,680]" box="[199,545,295,320]" pageId="6" pageNumber="242">
<emphasis id="00CFD728FFEAFFA3519327435404FEB4" bold="true" box="[199,352,295,320]" pageId="6" pageNumber="242">Distribution.</emphasis>
<collectingCountry id="4AAC4BAAFFEAFFA35033274354CEFEB4" box="[359,426,295,320]" name="China" pageId="6" pageNumber="242">China</collectingCountry>
(Yunnan).
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFEAFFA35193272851E1FDAB" pageId="6" pageNumber="242" type="diagnosis">
<paragraph id="32040B3AFFEAFFA35193272851E1FDAB" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="242">
<emphasis id="00CFD728FFEAFFA3519327285425FE91" bold="true" box="[199,321,332,357]" pageId="6" pageNumber="242">Diagnosis.</emphasis>
This new species can be distinguished from other similar species easily by elytra with dark green metallic hue in color, with sparser coarse setiferous punctures, somewhat arranged in longitudinal rows, and by the two characteristic small hook-shaped processes at apex of median lobe. However, it is also a little similar to
<taxonomicName id="F5BB70B9FFEAFFA354D527F05763FE24" authority="Smetana" authorityName="Smetana" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="242" phylum="Arthropoda" rank="species" species="phormio" subGenus="Distichalius">
<emphasis id="00CFD728FFEAFFA354D527F050F8FE5F" box="[1409,1436,404,427]" italics="true" pageId="6" pageNumber="242">Q.</emphasis>
(
<emphasis id="00CFD728FFEAFFA351F427DC5443FE24" box="[160,295,440,464]" italics="true" pageId="6" pageNumber="242">Distichalius</emphasis>
)
<emphasis id="00CFD728FFEAFFA3506827DC54FFFE24" box="[316,411,440,464]" italics="true" pageId="6" pageNumber="242">phormio</emphasis>
Smetana
</taxonomicName>
in male genitalia, but it can be distinguished from the latter by having more sensory peg setae on underside of paramere, arranged in two parallel longitudinal groups, median lobe abruptly constricted into small and narrow apex, with two apical processes; whereas
<taxonomicName id="F5BB70B9FFEAFFA352B22464500BFDEC" authority="Smetana" authorityName="Smetana" box="[998,1391,512,536]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="242" phylum="Arthropoda" rank="species" species="phormio" subGenus="Distichalius">
<emphasis id="00CFD728FFEAFFA352B224645166FDE3" box="[998,1026,512,535]" italics="true" pageId="6" pageNumber="242">Q.</emphasis>
(
<emphasis id="00CFD728FFEAFFA35546246451FDFDEC" box="[1042,1177,512,536]" italics="true" pageId="6" pageNumber="242">Distichalius</emphasis>
)
<emphasis id="00CFD728FFEAFFA355FE2464506DFDEC" box="[1194,1289,512,536]" italics="true" pageId="6" pageNumber="242">phormio</emphasis>
Smetana
</taxonomicName>
has fewer sensory peg setae on underside of paramere, arranged in two more laterally spread curved groups, median lobe gradually and evenly narrowed into acute apex with one hook-shaped apical process.
</paragraph>
</subSubSection>
<subSubSection id="7AA158B1FFEAFFA351932408519CFD5C" pageId="6" pageNumber="242" type="etymology">
<paragraph id="32040B3AFFEAFFA351932408519CFD5C" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="242">
<emphasis id="00CFD728FFEAFFA351932408542AFD71" bold="true" box="[199,334,620,645]" pageId="6" pageNumber="242">Etymology.</emphasis>
The specific name is derived from Latin prefix
<emphasis id="00CFD728FFEAFFA352CC240856C9FD70" box="[920,941,620,644]" italics="true" pageId="6" pageNumber="242">bi</emphasis>
- (double) and Latin adjective
<emphasis id="00CFD728FFEAFFA35449240850FFFD70" box="[1309,1435,620,644]" italics="true" pageId="6" pageNumber="242">prominulus</emphasis>
(projecting slightly), referring to the two characteristically shaped processes at apex of median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>