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<document ID-CLB-Dataset="297102" ID-DOI="10.5281/zenodo.7316519" ID-GBIF-Dataset="38aee501-3c26-4686-9c32-a0991b651db7" ID-ISBN="0-8018-8221-4" checkinTime="1670905099958" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="C7BA8239239D4F982C1C248018296588" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Artiodactyla.637-722.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press" docTitle="Addax nasomaculatus" docType="treatment" docVersion="4" id="AB6A4FBA464988A6B3A53F4B8CD52B4B" lastPageNumber="717" masterDocId="AB6A4FBA464988A6B3A53F4B8CD52B4B" masterDocTitle="Order Artiodactyla" masterLastPageNumber="722" masterPageNumber="637" pageNumber="717" updateTime="1716783024384" updateUser="ExternalLinkService">
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<taxonomicName id="71AF77EF59A8404225A952AC1814BCBC" ID-CoL="652YZ" authority="(de Blainville 1816)" baseAuthorityName="de Blainville" baseAuthorityYear="1816" class="Mammalia" family="Bovidae" genus="Addax" kingdom="Animalia" order="Artiodactyla" pageNumber="717" phylum="Chordata" rank="species" species="nasomaculatus">
<emphasis id="7B406F43CEC1734BDAF6CBBB1CDDFCFF" italics="true" pageNumber="717">Addax nasomaculatus</emphasis>
(de Blainville 1816)
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<paragraph id="EDD7AF68E8E5814E4D40813D65F619A9" pageNumber="717">
<treatmentCitation id="C447AB354CC8512BE95D80A0197B5A83" author="de Blainville" authority="de Blainville 1816" authorityName="de Blainville" authorityYear="1816" class="Mammalia" family="Bovidae" genus="Cerophorus" journal="Bull. Sci. Soc. Philom. Paris" kingdom="Animalia" order="Artiodactyla" page="75" pageNumber="717" phylum="Chordata" rank="species" species="nasomaculata" title="Cerophorus nasomaculata" volume="1816" year="1816">
<taxonomicName id="A1788C52B6E4C5E98B63996DC677E2E3" authority="de Blainville 1816" authorityName="de Blainville" authorityYear="1816" class="Mammalia" family="Bovidae" genus="Cerophorus" kingdom="Animalia" order="Artiodactyla" pageNumber="717" phylum="Chordata" rank="species" species="nasomaculata">
<emphasis id="4FACF0127990AF5B27AE9F06F36F6E68" italics="true" pageNumber="717">[Cerophorus] nasomaculata</emphasis>
de Blainville 1816
</taxonomicName>
,
<bibCitation id="EF4B8CCD8BE63EA72F284F9AD684442D" author="de Blainville" journal="Bull. Sci. Soc. Philom. Paris" pageNumber="717" pagination="75" title="Cerophorus nasomaculata" volume="1816" year="1816">Bull. Sci. Soc. Philom. Paris, 1816: 75</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7ECE56EA5AA1777DE9CDAE0ED6AF9338" pageNumber="717" type="materials_examined">
<paragraph id="7FCC5E1D965FB17BAAC843826C5AEB37" pageNumber="717">
<emphasis id="06CE6BB0A98FE8779F4E2385758E2F72" inLineHeading="true" pageNumber="717">Type Locality:</emphasis>
<materialsCitation id="F00B096157A3374E5EADEE819586FB50" ID-GBIF-Occurrence="4867329444" pageNumber="717">No locality given. Here selected as the Tunisian Sahara.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="26C684EE749581ABB841A185F4468ECD" pageNumber="717" type="vernacular_names">
<paragraph id="28E6CF6975AC78FEB0009B290B29B8AC" pageNumber="717">
<emphasis id="D09912CF168DB6153E96ACA009B85589" inLineHeading="true" pageNumber="717">Vernacular Names:</emphasis>
<vernacularName id="1609FB84D00C961C56332532C43A0428" language="eng" pageNumber="717">Addax</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="EB8AFE986856E6FCE19085C20BD0A08B" pageNumber="717" type="distribution">
<paragraph id="8EC15914765C3447EED2F10FC615F0BC" pageNumber="717">
<emphasis id="A9991F984C96B33FFC397CC71886AF56" inLineHeading="true" pageNumber="717">Distribution:</emphasis>
Extinct in
<collectingCountry id="59F00FA10977FBF4ACE956C1CFAB1AAD" name="Algeria">Algeria</collectingCountry>
,
<collectingCountry id="CFC0EAF11AF2D202ECEB91A936CF1C62" name="Egypt">Egypt</collectingCountry>
,
<collectingCountry id="DAB786E5FC7B5D8EF0488EFB3830FA32" name="Libya">Libya</collectingCountry>
,
<collectingCountry id="CBA19F292DBE600CC99676A431D65D89" name="Morocco">Morocco</collectingCountry>
,
<collectingCountry id="4A52D80CA6495CAA40E4E03CDD9E1C1A" name="Tunisia">Tunisia</collectingCountry>
, and probably
<collectingCountry id="DAD136C3F91767A5E9CBF50A99203841" name="Sudan">Sudan</collectingCountry>
. Vagrants still enter
<collectingCountry id="CDF4469EB59E80A2D68506889035C71E" name="Algeria">Algeria</collectingCountry>
and
<collectingCountry id="D9B44620F80FE5E0578478F7F3C6D01D" name="Sudan">Sudan</collectingCountry>
. Survives in
<collectingCountry id="0DD66A1EECA1D3A3F34188CB1CF652B5" name="Chad">Chad</collectingCountry>
, N
<collectingCountry id="19B02058E4099030C61EC0EDBAECA079" name="Serbia">Mali</collectingCountry>
,
<collectingCountry id="E0243E26798B7A2CC6A9EF3AEFA722E2" name="Mauritania">Mauritania</collectingCountry>
, and
<collectingCountry id="271F333535350C50FEA003A68EC17554" name="Niger">Niger</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6620791A6195619F68DC3EBE8236C934" pageNumber="717" type="conservation">
<paragraph id="289A0A8CF6E5E82D5FD650BFCC8D34D2" pageNumber="717">
<emphasis id="FB58ACB0714C3749309B0CCE52164347" inLineHeading="true" pageNumber="717">Conservation:</emphasis>
Nearly extinct in wild (
<bibRefCitation id="685FF5BDDB5FDE3AEB453886C46F1BBD" author="East, R." refId="ref91247" refString="East, R. (ed.). 1990. Antelopes. Global survey and regional action plans. Part 3. West and central Africa. I. U. C. N., Gland, Switzerland, 171 pp." year="1990">East, 1990</bibRefCitation>
).
<collectionCode id="142D3C8E7ABB1F7D05AEFBC06AB55C16">CITES</collectionCode>
Appendix I;
<collectingCountry id="C82DAB7207366EA5A56B8DE687027557" name="United States of America">U.S.</collectingCountry>
<collectionCode id="0B01FF66973737B4257EE002B5D85088" country="Brazil" lsid="urn:lsid:biocol.org:col:14319" name="Universidade de São Paulo" type="Herbarium">ESA</collectionCode>
Proposed Endangered;
<collectionCode id="F9EEDEC9B1D599F9D39AAD09A07D1E0C">IUCN</collectionCode>
Critically Endangered.
</paragraph>
</subSubSection>
<subSubSection id="D823C0738F59A0F4EB6972EC389EABE9" pageNumber="717" type="discussion">
<paragraph id="E362D97C6DD0690C4502008FE227DCF9" pageNumber="717">
<emphasis id="5D2901DF137661AB39294174066C4CF6" inLineHeading="true" pageNumber="717">Discussion:</emphasis>
The
<typeStatus id="5DC42AFB330F6851C2DC824BE98F81C2">syntypes</typeStatus>
were observed by de Blainville in Bullock's Pantherion or Museum and the Museum of the Royal College of Surgeons, both in London,
<collectingCountry id="27FE8693F36C3ACC2D077CA3DE1B8BFD" name="United Kingdom">UK</collectingCountry>
. C. H.
<bibRefCitation id="03CCF9C1882E28416A88657BAD5EB2E6" author="Smith, C. H." refId="ref330696" refString="Smith, C. H. 1827. The seventh order of the Mammalia. The Ruminantia. Pp. 1 - 428, in The animal kingdom arranged in conformity with its organization, by the Baron Cuvier, member of the Institute of France, &amp; c. &amp; c. &amp; c. with additional descriptions of all the species hitherto named, and of many not before noticed (E. Griffith, C. H. [amilton] Smith, and E. Pidgeon, eds.). G. B. Whitaker, London, 4: 1 - 498." year="1827">Smith (1827)</bibRefCitation>
suggested the specimens came from
<collectingCountry id="E2D8AEE246898E08BE84587D8C02FCEE" name="Guinea-Bissau">Guinea</collectingCountry>
or Western Africa; Lydekker (1914
<emphasis id="84E78D15102E2EFC9F8B08809CFEA15C" italics="true" pageNumber="717">b</emphasis>
:148) stated that the type locality was probably Senegambia. These authors provided no evidence to support their conclusions and from the discussion in Sclater and Thomas (1898), it seems more probable that British hunters or collectors obtained
<taxonomicName id="A9F4A496107BA2920D3A4CE76AC07BAD" authorityName="Laurillard" authorityYear="1841" class="Mammalia" family="Bovidae" genus="Addax" kingdom="Animalia" order="Artiodactyla" phylum="Chordata" rank="genus">Addax</taxonomicName>
from the Tunisian Sahara, to which the type locality is here restricted.
</paragraph>
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Genus
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<taxonomicName id="CC0B0B82A80F6200C7E871935DA3BBA0" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">Tychepsephus</taxonomicName>
Waterhouse, 1876: 15.
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<paragraph id="2EE0840F66B2AAF288489EBDA864414D" pageId="0" pageNumber="23">
<taxonomicName id="180ECAC5AB91281C0F5A50FEFF52B4A8" authority="" class="Insecta" family="Psephenidae" genus="Ectopria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectopria (Chilectopria)" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="subGenus" subGenus="Chilectopria">Ectopria (Chilectopria)</taxonomicName>
Pic, 1947: 3-4, syn. nov.
</paragraph>
<paragraph id="C72250B067F39C5F9FEF73CB4A9BA685" pageId="0" pageNumber="23">
<taxonomicName id="8D2DB2E3CDBAE609A0517F832D606B9C" family="Psephenidae" genus="Tychepselaphus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tychepselaphus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">Tychepselaphus</taxonomicName>
: Philippi, 1887: 665, lapsus calami.
</paragraph>
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<taxonomicName id="AFF45D58ECB93A9182F0E2E31DA930F1" class="Insecta" family="Psephenidae" genus="Tychepsephenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tychepsephenus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">Tychepsephenus</taxonomicName>
: Zaitzev, 1910: 4, lapsus calami.
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<taxonomicName id="A6FF4611675464038255AD63FA40E7F0" class="Insecta" family="Psephenidae" genus="Tychepsephenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tychepsephenus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">Tychepsephenus</taxonomicName>
: Blackwelder, 1944: 274, lapsus calami.
</paragraph>
<paragraph id="1E25634AF6490820451DC33D557D87B9" pageId="0" pageNumber="23">
<taxonomicName id="AB1AB7A981AD02F8F0B89FAE0CAB1EC8" class="Insecta" family="Psephenidae" genus="Tychepsephenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tychepsephenus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">Tychepsephenus</taxonomicName>
: Artigas, 1963: 6, lapsus calami.
</paragraph>
<paragraph id="00DA36CE2D8AA9AD4B05EDDD8DE963C4" pageId="0" pageNumber="23">
<taxonomicName id="831DDEA9B2CAFADB6EDD659A38997BD3" class="Insecta" family="Psephenidae" genus="Tychepsephenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tychepsephenus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">Tychepsephenus</taxonomicName>
: Moroni, 1985: 173, lapsus calami.
</paragraph>
<paragraph id="F3B66C85BE9A72FC2CE2B5FBA8A71387" pageId="0" pageNumber="23">
<taxonomicName id="79A0BC7267001877779FAF03EB206816" class="Insecta" family="Psephenidae" genus="Tychepsephenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tychepsephenus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">Tychepsephenus</taxonomicName>
: Ashworth &amp; Hoganson, 1987: 879, lapsus calami.
</paragraph>
<paragraph id="C69C462C8FDD221AA13F19A730F5D952" pageId="0" pageNumber="23">
<taxonomicName id="5684C205EB0D2CFF6789EFFA1581872E" class="Insecta" family="Psephenidae" genus="Tychepsephenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tychepsephenus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">Tychepsephenus</taxonomicName>
:
<bibRefCitation id="78AB6C783DF154DC9E4469AF9381F362" DOI="https://dx.doi.org/10.1016/B978-0-12-801223-6.00027.5" author="Passos, MI" editor="Hamada, N" journalOrPublisher="4 th edn. Academic Press, Salt Lake City" pageId="0" pageNumber="23" pagination="583 - 598" refId="B16" refString="Passos, MI, Manzo, V, Maier, CA, 2018. Chapter 15.6. Families Dryopidae, Elmidae, and Psephenidae. In: Hamada, N, Thorp, J, Rogers, DC, Eds., Thorp and Covich's Freshwater Invertebrates - Volume III - Keys to Neotropical Hexapoda. 4th edn. Academic Press, Salt Lake City: 583 - 598, DOI: https://dx.doi.org/10.1016/B978-0-12-801223-6.00027.5" title="Chapter 15.6. Families Dryopidae, Elmidae, and Psephenidae." url="https://dx.doi.org/10.1016/B978-0-12-801223-6.00027.5" volumeTitle="Thorp and Covich's Freshwater Invertebrates - Volume III - Keys to Neotropical Hexapoda." year="2018">Passos et al., 2018</bibRefCitation>
: 598, lapsus calami.
</paragraph>
</subSubSection>
<subSubSection id="30C2261899BBE1B3F531E49354A2A4C6" pageId="0" pageNumber="23" type="type species">
<paragraph id="1FE40F743E136EBF50672ED24E699D51" pageId="0" pageNumber="23">Type species.</paragraph>
<paragraph id="1874142D1C3D22F98664E35B2025B362" pageId="0" pageNumber="23">
<taxonomicName id="F9BC9A54830AFBC9B7785495E648043F" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="E926F11D5C14E17C8F6B0FE280AC6CB1" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
Waterhouse, 1876, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="6ACB4012A62B51CBBAB27FE0DAF36803" pageId="0" pageNumber="23" type="etymology">
<paragraph id="31AD478528D1A8F0938043A2F7DE2A97" pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph id="BA95DF32E3B507E5565933BE1768D8A0" pageId="0" pageNumber="23">
<bibRefCitation id="5FFE86F18C088C776645ECD69555F2FD" DOI="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" author="Waterhouse, CO" journalOrPublisher="Transactions of the Royal Entomological Society" pageId="0" pageNumber="23" pagination="11 - 25" refId="B25" refString="Waterhouse, CO, 1876. III. On various new genera and species of Coleoptera. Transactions of the Royal Entomological Society 1876 (1): 11 - 25, DOI: https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" title="III. On various new genera and species of Coleoptera." url="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" volume="1876" year="1876">Waterhouse (1876)</bibRefCitation>
did not explain the etymology of the genus. However,
<emphasis id="6CBD17A70356B3C928CBF393E64EA01E" italics="true" pageId="0" pageNumber="23">Tyche</emphasis>
(Gr.) refers to the goddess of fortune; and the suffix, -
<emphasis id="83076D274B3D93D98E33E796CBCF5AE3" italics="true" pageId="0" pageNumber="23">psephus</emphasis>
, from
<emphasis id="82572415ABD3A4C3A34A10423A372DB8" italics="true" pageId="0" pageNumber="23">psephenos</emphasis>
(Gr.), meaning &quot;dark, obscure,&quot; has been used as the stem for multiple psephenid genera.
</paragraph>
</subSubSection>
<subSubSection id="79747620EA1D36191AE96827E48BC1F0" pageId="0" pageNumber="23" type="adult diagnosis">
<paragraph id="E60B7E14DA7A1DED75DF3F5173D9D094" pageId="0" pageNumber="23">Adult diagnosis.</paragraph>
<paragraph id="7F7F149990ED8867F00FA7D68A5E2F29" pageId="0" pageNumber="23">
The following characters used to distinguish
<taxonomicName id="62AC95C459A66FBD54D6733F96354BB1" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="0EF105A0EF8D1001A1CE2352D4C5C814" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
from the neotropical eubriine genera
<taxonomicName id="68D606EF3F83D5B192D8C7DC6B79BB2D" authorityName="Guerin-Meneville" authorityYear="1861" class="Insecta" family="Psephenidae" genus="Dicranopselaphus" kingdom="Animalia" lsidName="Dicranopselaphus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="4DFC8D27532CB7D0A03510005007C41A" italics="true" pageId="0" pageNumber="23">Dicranopselaphus</emphasis>
</taxonomicName>
,
<taxonomicName id="398BB511B33611AF707A5F9314E149CF" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="FDC231950E2814F240D2CDD55D8958AF" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
, and
<taxonomicName id="AE2B26B642635BBB96AED91507DBCEBE" authorityName="Shepard &amp; Barr" authorityYear="2014" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neoeubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="DC8E8E1C273919B7C74FFA80D4C98D40" italics="true" pageId="0" pageNumber="23">Neoeubria</emphasis>
</taxonomicName>
are for the most part taken from
<bibRefCitation id="0C9E47DEA175F6DD3981E9982161A842" DOI="https://doi.org/10.1111/j.1365-3113.2006.00374.x" author="Lee, C-F" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="23" pagination="502 - 538" refId="B13" refString="Lee, C-F, Sato, M, Shepard, WD, Jaech, MA, 2007. Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters. Systematic Entomology 32 (3): 502 - 538, DOI: https://doi.org/10.1111/j.1365-3113.2006.00374.x" title="Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters." url="https://doi.org/10.1111/j.1365-3113.2006.00374.x" volume="32" year="2007">Lee et al. (2007</bibRefCitation>
,
<bibRefCitation id="8DBD9C13BBF28914EC9AFF290E33942D" DOI="https://doi.org/10.1515/9783110373929" author="Lee, C-F" editor="Beutel, RG" journalOrPublisher="2 nd edn. Walter de Gruyter, Berlin" pageId="0" pageNumber="23" pagination="615 - 628" refId="B14" refString="Lee, C-F, Jaech, MA, Beutel, RG, 2016. Psephenidae Lacordaire, 1854. In: Beutel, RG, Leschen, RAB, Eds., Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics. 2nd edn. Walter de Gruyter, Berlin: 615 - 628, DOI: https://doi.org/10.1515/9783110373929" title="Psephenidae Lacordaire, 1854." url="https://doi.org/10.1515/9783110373929" volumeTitle="Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics." year="2016">2016</bibRefCitation>
): 1) antenna of the male weakly serrate (pectinate in
<taxonomicName id="C975A80637DF64EFADB1DF5221D77045" authorityName="Shepard &amp; Barr" authorityYear="2014" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neoeubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9253B56109016D31B6239D2846007FEE" italics="true" pageId="0" pageNumber="23">Neoeubria</emphasis>
</taxonomicName>
); 2) pronotum with serrate lateral margins (not serrate in
<taxonomicName id="8E58AF260DDBD8E18AF2FDB0F4D5E0F2" authorityName="Guerin-Meneville" authorityYear="1861" class="Insecta" family="Psephenidae" genus="Dicranopselaphus" kingdom="Animalia" lsidName="Dicranopselaphus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="1199AB1C3E032CB4ECD3D7EB55D7499E" italics="true" pageId="0" pageNumber="23">Dicranopselaphus</emphasis>
</taxonomicName>
and
<taxonomicName id="AAD5F3B20ACA8A81E0C7A6E4BA7F467A" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="1C767F0A2003FB82F54328D32CB5B88E" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
); 3) mesoventrite with a median longitudinal sulcus (absent in
<taxonomicName id="EE8674D8BD70FC9D2BB6230099E8CF46" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="F1795531E6694420264126DF59A4FDEB" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
and some
<taxonomicName id="99615A0F61F337FD646BDA36BCCDF8C1" authorityName="Guerin-Meneville" authorityYear="1861" class="Insecta" family="Psephenidae" genus="Dicranopselaphus" kingdom="Animalia" lsidName="Dicranopselaphus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="E3987B476BBC84DE4D51718BFAE09703" italics="true" pageId="0" pageNumber="23">Dicranopselaphus</emphasis>
</taxonomicName>
); and 4) metaventrite with a transverse suture (vestigial in other eubriines except
<taxonomicName id="1A515F22FB90238C1790704A6E15C9A6" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="0B7F5E95E3A0D117BDB28D5631E99108" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
). For the original characterization of adults of
<taxonomicName id="59F56B90615555517C9080EA19CC9129" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="16196AFE35255D762350755A313B1149" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
, see
<bibRefCitation id="33CDE9985A4C96632D693149BBDCB4F1" DOI="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" author="Waterhouse, CO" journalOrPublisher="Transactions of the Royal Entomological Society" pageId="0" pageNumber="23" pagination="11 - 25" refId="B25" refString="Waterhouse, CO, 1876. III. On various new genera and species of Coleoptera. Transactions of the Royal Entomological Society 1876 (1): 11 - 25, DOI: https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" title="III. On various new genera and species of Coleoptera." url="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" volume="1876" year="1876">Waterhouse (1876</bibRefCitation>
: 15-16).
</paragraph>
<paragraph id="5BA08B81A37B611411CBA811978BDFD1" pageId="0" pageNumber="23">
<taxonomicName id="C804FC85BBC1B8E644EDFA6AA9432883" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="57C0EF8C559642CFC11B783E95B68508" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
and the Australian genus
<taxonomicName id="ECE4D4F8AE3817B5FAD62D4DD24C5C8B" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="3F897BD6DB618D79A6FBC3F5F5B08FD3" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
are closely related sister genera which, in the adult stage, do not have many good characters to separate them. One obvious difference is that in
<taxonomicName id="2B6E37A0FCEE1F32AB0E715AB620FA5A" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="5E710D1488E6177075D70528A49FB087" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
the apical margins of abdominal ventrites 2-4 are entire, whereas in
<taxonomicName id="EC5152CECB3BCC95F8952276285D30A1" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B39BEB8F3497347733A90C7EC2DB9DC7" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
they are serrate. Of course, there is also the geographical difference with each occurring on different continents:
<taxonomicName id="2234D1200D31A5784800246073DF9D5A" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="B6BBF172D584A31E18C49BF950FA11A9" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
in South America and
<taxonomicName id="8240F28E65B34E933588DA818FF7FF56" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="54C2FA27DC9171A8BEE58531F0CB1DBF" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
in Australia.
</paragraph>
</subSubSection>
<subSubSection id="E954FE446F3CB55B9E934176530CDB68" pageId="0" pageNumber="23" type="description">
<paragraph id="4C254D095E90A955343F0A484FE8773F" pageId="0" pageNumber="23">Adult description.</paragraph>
<paragraph id="68A4534DFFD2158308C557ECD0292A41" pageId="0" pageNumber="23">
<bibRefCitation id="CBC524C8CA11AF9775E76976235AABA5" DOI="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" author="Waterhouse, CO" journalOrPublisher="Transactions of the Royal Entomological Society" pageId="0" pageNumber="23" pagination="11 - 25" refId="B25" refString="Waterhouse, CO, 1876. III. On various new genera and species of Coleoptera. Transactions of the Royal Entomological Society 1876 (1): 11 - 25, DOI: https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" title="III. On various new genera and species of Coleoptera." url="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" volume="1876" year="1876">Waterhouse (1876)</bibRefCitation>
described the genus
<taxonomicName id="CED169418A0FA5ED054F08FD0F58257B" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="26F6A520AD78E1A40C1BF15FD704750F" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
in English, unlike the type species description which is in Latin. His description is adequate for identifying the genus. However, since Waterhouse had only a female specimen, he did not know that his species,
<taxonomicName id="ED0367A488FD90F24B4AA8E1907128AB" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="F48816538EE80DFFCCF4D44115BAFC2F" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
, is sexually dimorphic, with males having a distinctive patch of setae on the abdominal venter. This character is also present in
<taxonomicName id="4E27492BB913673ABF08915E33706884" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="EFE9165488E5C07EA38786C21A0F7B61" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
sp. nov. The dorsal patterning also differs between the sexes of both species.
</paragraph>
</subSubSection>
<subSubSection id="E6843936E712BA7E1F47819D69E4239F" pageId="0" pageNumber="23" type="remarks">
<paragraph id="F85254A5D91680797E15EAB70DDC6DE9" pageId="0" pageNumber="23">Remark, adult males.</paragraph>
<paragraph id="B0BCA596051F0DACE23D28F5B3AF0F1D" pageId="0" pageNumber="23">A new morphological note is that males have a sperm pump composed of a heavily muscularized ejaculatory duct which is situated medially, anterior to the aedeagus, bent around itself in an S-shape and coming from the juncture of the paired testes. The sperm pump is as short as or shorter than the aedeagus.</paragraph>
</subSubSection>
<subSubSection id="DA8ADB92A32C16B28FF6AD5BF2DC08AA" pageId="0" pageNumber="23" type="larval diagnosis">
<paragraph id="4C177D4A85645511F9E963BDE5713229" pageId="0" pageNumber="23">Larval diagnosis.</paragraph>
<paragraph id="34AD02A1D6B3AED2201587FBD7AAFEE6" pageId="0" pageNumber="23">
The following characters used to distinguish
<taxonomicName id="E5A838816FEE4E554F59884FA7431181" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="7E5F77E7AAA0380047D87C1E92489BC4" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
from the neotropical eubriine genera
<taxonomicName id="F8818389EFD6839D24D96F6FA721F646" authorityName="Guerin-Meneville" authorityYear="1861" class="Insecta" family="Psephenidae" genus="Dicranopselaphus" kingdom="Animalia" lsidName="Dicranopselaphus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C8886C59A92CCDA37BA9E90C5C519906" italics="true" pageId="0" pageNumber="23">Dicranopselaphus</emphasis>
</taxonomicName>
,
<taxonomicName id="3751CBBCD19CA9A0D57A714B7D268104" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="57F3378211E9EC806AC7B46C45C04B37" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
, and
<taxonomicName id="C4A87EDE06AC215DB20592F9A98158C3" authorityName="Shepard &amp; Barr" authorityYear="2014" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neoeubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="2889C40A17D1EE114BAA68EEE5020B81" italics="true" pageId="0" pageNumber="23">Neoeubria</emphasis>
</taxonomicName>
are for the most part taken from
<bibRefCitation id="4F760F15D294DB891DB2F2ACE3C869D3" DOI="https://doi.org/10.1111/j.1365-3113.2006.00374.x" author="Lee, C-F" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="23" pagination="502 - 538" refId="B13" refString="Lee, C-F, Sato, M, Shepard, WD, Jaech, MA, 2007. Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters. Systematic Entomology 32 (3): 502 - 538, DOI: https://doi.org/10.1111/j.1365-3113.2006.00374.x" title="Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters." url="https://doi.org/10.1111/j.1365-3113.2006.00374.x" volume="32" year="2007">Lee et al. (2007</bibRefCitation>
,
<bibRefCitation id="783CBE7EDDE7F6FE206D55D81EA7BA6E" DOI="https://doi.org/10.1515/9783110373929" author="Lee, C-F" editor="Beutel, RG" journalOrPublisher="2 nd edn. Walter de Gruyter, Berlin" pageId="0" pageNumber="23" pagination="615 - 628" refId="B14" refString="Lee, C-F, Jaech, MA, Beutel, RG, 2016. Psephenidae Lacordaire, 1854. In: Beutel, RG, Leschen, RAB, Eds., Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics. 2nd edn. Walter de Gruyter, Berlin: 615 - 628, DOI: https://doi.org/10.1515/9783110373929" title="Psephenidae Lacordaire, 1854." url="https://doi.org/10.1515/9783110373929" volumeTitle="Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics." year="2016">2016</bibRefCitation>
): 1) antennomere 1 subequal to antennomere 2 (much shorter in other eubriines except
<taxonomicName id="9C9311F85A820647EE2809540368D2F1" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="EB8E726F47B693827ACC969FA242969F" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
); 2) maxillary and labial palpi with 4 and 3 palpomeres, respectively (
<taxonomicName id="5A7F9ED7FCAEA7FCFE56A9EA10951672" authorityName="Guerin-Meneville" authorityYear="1861" class="Insecta" family="Psephenidae" genus="Dicranopselaphus" kingdom="Animalia" lsidName="Dicranopselaphus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="1EAE1912E0A6ADE36E98459CC3C481AA" italics="true" pageId="0" pageNumber="23">Dicranopselaphus</emphasis>
</taxonomicName>
and
<taxonomicName id="9B6C9F28968F5622711B5002B4FAF431" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="199895EF59EF003222E23B8604FD8982" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
with 3 and 2, respectively); 3) longitudinal medial suture from thorax to abdominal segment VII (not in other eubriines except
<taxonomicName id="8CB3DA3DAECCBD9BB66B897FBFD6D5DB" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="02E52750BFC6241CF40771536C4B5B47" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="AA24D8A781D6D6C0443A24EFD7C8BF99" pageId="0" pageNumber="23">
Larval characters separating
<taxonomicName id="183D079317D8FF8FF96081D3E2970A9D" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="2F96194EFEEF7C9FD390E31740B10EB2" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
from its Australian sister genus
<taxonomicName id="0EEDF37CE5B746643E39B4F518372A64" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="91DC25A3D82B0A4EE7230744A9B568AD" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
include the following: 1) setae on the posterior margin of the thoracic and abdominal tergites hair-like in
<taxonomicName id="980D5531C7BD516F708EAE4C0894492E" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="3530BCDE1B806C91E0173C50214F6B7F" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
and absent in
<taxonomicName id="ED0EF5019EFB8991E8ADD0D0B8DE4C72" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C05A2E8463D306164F4639A43522A73E" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
; 2) paired longitudinal rows of setae or sensillae near the dorsal midline in
<taxonomicName id="E14844EE949762478EAC444D8BBD4EE8" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="59C6E5377ED3079783578F9B426961AE" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
, but not in
<taxonomicName id="C68458BC3481707267988F6521B9AC12" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="26FFA55267C208B9BEFB9870225D5265" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
; and 3) gin traps on the abdominal tergites of
<taxonomicName id="F57AD9C3744E996CA483FF8F445A6DCC" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="60F4C9D5A574FE3851C41C04FB349F50" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
, but not on
<taxonomicName id="55304E80EA9CB70896C207E24FB5E465" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="394A3FCF331B2C1DAC2C7B9F80CA0847" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="70FA6FA7F9BBD4402F795DED721925CC" pageId="0" pageNumber="23" type="description">
<paragraph id="28592504E180164B5CCE0AB918698EF3" pageId="0" pageNumber="23">Larval descriptions.</paragraph>
<paragraph id="3007382C8E1CDD062E0AC5718096EC43" pageId="0" pageNumber="23">
<taxonomicName id="2C20FEB5ECEAF810E398C57DEE310EEE" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="FD0DAB0F56543A587A996EC8BB00CC08" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
larvae were previously well-described by
<bibRefCitation id="6A30156B371E28494D36D9A1540908B3" author="Lataste, F" journalOrPublisher="Actes de la Societe Scientifique du Chili" pageId="0" pageNumber="23" pagination="103 - 106" refId="B11" refString="Lataste, F, 1897a. Presentation d'une larve de Pseudo-Nevroptere d'aspect crustaceen. Actes de la Societe Scientifique du Chili 7: 103 - 106" title="Presentation d'une larve de Pseudo-Nevroptere d'aspect crustaceen." volume="7" year="1897 a">Lataste (1897a)</bibRefCitation>
and
<bibRefCitation id="B002C3144406B3F0F860D123158BAF41" author="Artigas, J" journalOrPublisher="Gayana (Concepcion)" pageId="0" pageNumber="23" pagination="3 - 11" refId="B1" refString="Artigas, J, 1963. Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae). Gayana (Concepcion) 1963 (8): 3 - 11" title="Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae)." volume="1963" year="1963">Artigas (1963)</bibRefCitation>
. Our collections revealed the presence of two distinct larval morphotypes (Figs
<figureCitation id="D3871240B509DA6E35B89291371392B5" captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Tychepsephus larvae 1 morph 1 a dorsal view b ventral view 2 morph 2 a dorsal view b ventral view." figureDoi="10.3897/zookeys.1164.103184.figures1-2" httpUri="https://binary.pensoft.net/fig/855185" pageId="0" pageNumber="23">1</figureCitation>
,
<figureCitation id="DC6F53FD006B667DCA82B088C426CF7B" captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Tychepsephus larvae 1 morph 1 a dorsal view b ventral view 2 morph 2 a dorsal view b ventral view." figureDoi="10.3897/zookeys.1164.103184.figures1-2" httpUri="https://binary.pensoft.net/fig/855185" pageId="0" pageNumber="23">2</figureCitation>
) which have not been associated with adults and are therefore unnamed.
<bibRefCitation id="C29CFE6D4AFA4EFE9F3D5D37E52C0C60" author="Artigas, J" journalOrPublisher="Gayana (Concepcion)" pageId="0" pageNumber="23" pagination="3 - 11" refId="B1" refString="Artigas, J, 1963. Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae). Gayana (Concepcion) 1963 (8): 3 - 11" title="Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae)." volume="1963" year="1963">Artigas (1963)</bibRefCitation>
illustrated a larva that corresponds to our larval morph 2, below.
</paragraph>
<caption id="8C5BA227A8079D7CED86753A0A9A3E06" doi="10.3897/zookeys.1164.103184.figures1-2" httpUri="https://binary.pensoft.net/fig/855185" pageId="0" pageNumber="23" start="Figures 1, 2" startId="F1">
<paragraph id="63A11B22D80EC969D787B6103B63894A" pageId="0" pageNumber="23">
<emphasis id="DB9708A58089414AB2F276679E30A80D" bold="true" pageId="0" pageNumber="23">Figures 1, 2.</emphasis>
<taxonomicName id="7F582D814C23CF0B610074A9B18A6DC1" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="2EEF618300004E048A09C7FAE0EB4315" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
larvae
<emphasis id="8EE8299C085B0162D844BB6E2354B4EE" bold="true" pageId="0" pageNumber="23">1</emphasis>
morph 1
<emphasis id="ECECDCB1F9330681A50398DADBE20EF1" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="D33D8791CA0B6D17638A0AC57512B2B4" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view
<emphasis id="795F0502D315749CB479AF839373D60F" bold="true" pageId="0" pageNumber="23">2</emphasis>
morph 2
<emphasis id="4F3DDB2C583E09237CFCC0294B8885DF" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="34EEA430EB0FB083E0AD5B1C413FAAA8" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="66BDB798269C0B9FFB88DD63243E0019" pageId="0" pageNumber="23">
<emphasis id="811D734F9D8DB0AC08C18C359F1BF7F9" bold="true" pageId="0" pageNumber="23">
<taxonomicName id="0D54F5FFC94AA293DF2B88815AA490B5" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="0484D2471082248BAB8DD363B1E65628" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
larval morph 1 (Fig.
<figureCitation id="084B1BBD33384C9D1E23ECF811D51A46" captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Tychepsephus larvae 1 morph 1 a dorsal view b ventral view 2 morph 2 a dorsal view b ventral view." figureDoi="10.3897/zookeys.1164.103184.figures1-2" httpUri="https://binary.pensoft.net/fig/855185" pageId="0" pageNumber="23">1</figureCitation>
).
</emphasis>
Body shape elongate-oval; color brown and red-brown with variable yellow patterning; tergites darker than paratergites. Body margined with a long, dense fringe of golden yellow, white-tipped setae. Dorsal surface with very small, scattered cuticular beads. Longitudinal medial suture from middle of pronotum to AB VIII tergite. Mesothoracic tergite to AB VII tergite each with a pair of large, prominent, dark tubercles straddling the medial suture. Abdominal tergite VIII clasping AB IX laterally; AB VIII with a pair of large, spiracular tubercles on posterior margin at bases of paratergites. Abdominal tergite IX subquadrate, flattened, apex rounded. Paratergites generally rectangular, more than twice as wide as long, anterolateral margin curved. Ventral surfaces lacking tubercles and cuticular beads. Abdominal ventrites I-VIII with sternopleural sutures. Operculum as long as wide, widest just anterior to apex; apex broadly rounded.
</paragraph>
<paragraph id="85ED35D219E9D49227A2BE0AC34D738E" pageId="0" pageNumber="23">
<emphasis id="BEF61A69FD95EE53FDDDC876F10B079D" bold="true" pageId="0" pageNumber="23">
<taxonomicName id="11F83700431DDBE7E581D3C91369309F" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="9E31A4C9A76E63AB47743D58EA5DAD71" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
larval morph 2 (Fig.
<figureCitation id="6A96718E276052178FEB0C3606A35320" captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Tychepsephus larvae 1 morph 1 a dorsal view b ventral view 2 morph 2 a dorsal view b ventral view." figureDoi="10.3897/zookeys.1164.103184.figures1-2" httpUri="https://binary.pensoft.net/fig/855185" pageId="0" pageNumber="23">2</figureCitation>
).
</emphasis>
Body shape elongate-oval; color brown to red-brown with yellow areas of variable size, shape, and position, often along midline of tergites and base of paratergites. Body margined with a long, dense fringe of yellow-brown, white-tipped setae. Dorsal surface sculptured with shallow, irregularly shaped depressions of varying sizes; cuticle with numerous, round, dark, flat-topped tubercles arranged in curvilinear shapes often encircling the depressions. Longitudinal medial suture from middle of pronotum to AB VII tergite. Tergites without pairs of prominent tubercles at the midline. Abdominal tergites each with an irregular, transverse line of tubercles; paratergites often with a longitudinal line of tubercles near the midline. Abdominal tergite VIII with a pair of large spiracular tubercles on posterior margin at base of paratergites. Abdominal tergite IX subelliptical, convex, sometimes sculptured and laterally angulate. Paratergites I-VIII paddle-shaped, twice as wide as long, narrower at base than apex, anterolateral margin curved. Abdominal ventrites with anterior and posterior transverse lines of small, faint, brown tubercles. Abdominal ventrites I-VIII with sternopleural sutures. Operculum nearly circular from midline to apex.
</paragraph>
</subSubSection>
<subSubSection id="37FF55089DC0DD1B2CAE3228F4074820" pageId="0" pageNumber="23" type="remarks">
<paragraph id="1834494BED433F601D2F356F13280A6E" pageId="0" pageNumber="23">Remarks, larvae.</paragraph>
<paragraph id="BE6CDA42FF78B657542852921A623620" pageId="0" pageNumber="23">The most obvious differences between the larval morphs are: 1) the presence on the dorsum of numerous dark tubercles, some in curvilinear patterns (morph 2); 2) the presence of pairs of large, prominent tubercles at the midline (morph 1); 3) the shape of AB IX tergite; and 4) the shape of the operculum. The dorsal morphology of larval morph 2 is quite variable in regard to the number and arrangement of tubercles and the amount of sculpturing. Because of this, it would not be surprising if another undescribed species is discovered with further sampling of the adult habitat.</paragraph>
<paragraph id="3F520DF3E49BFF80C9DC07FC29B80A7A" pageId="0" pageNumber="23">Some larvae have an abundance of peritrich protozoans attached to the venter in a scattered fashion on both sclerites and membranes.</paragraph>
</subSubSection>
<subSubSection id="E181D7FBABDFE19650929E3C3008E2AC" pageId="0" pageNumber="23" type="description">
<paragraph id="C30973533FF34658450911209751AB8F" pageId="0" pageNumber="23">Pupal description</paragraph>
<paragraph id="3FF8844781F8FF0F93FF1249945DB81C" pageId="0" pageNumber="23">
<emphasis id="D2175C042376B714DE238927E090139B" bold="true" pageId="0" pageNumber="23">
(pupa of larval morph 2) (Figs
<figureCitation id="364FD62D8E12E9119BF8C78B16A095B9" captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Tychepsephus pupa and larval exuvium 3 pupa a dorsal view b ventral view 4 exuvium of last larval instar, ventral view." figureDoi="10.3897/zookeys.1164.103184.figures3-4" httpUri="https://binary.pensoft.net/fig/855186" pageId="0" pageNumber="23">3</figureCitation>
,
<figureCitation id="0063105F7DF8E4B30381A52F656B8A11" captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Tychepsephus pupa and larval exuvium 3 pupa a dorsal view b ventral view 4 exuvium of last larval instar, ventral view." figureDoi="10.3897/zookeys.1164.103184.figures3-4" httpUri="https://binary.pensoft.net/fig/855186" pageId="0" pageNumber="23">4</figureCitation>
).
</emphasis>
Pupa (Fig.
<figureCitation id="69F8AA8FD24F140B19CFD7E8CE41B1C9" captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Tychepsephus pupa and larval exuvium 3 pupa a dorsal view b ventral view 4 exuvium of last larval instar, ventral view." figureDoi="10.3897/zookeys.1164.103184.figures3-4" httpUri="https://binary.pensoft.net/fig/855186" pageId="0" pageNumber="23">3</figureCitation>
) under exuvium of last larval instar (Fig.
<figureCitation id="161156A47D28B8D9CAB62966AE51C35D" captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Tychepsephus pupa and larval exuvium 3 pupa a dorsal view b ventral view 4 exuvium of last larval instar, ventral view." figureDoi="10.3897/zookeys.1164.103184.figures3-4" httpUri="https://binary.pensoft.net/fig/855186" pageId="0" pageNumber="23">4</figureCitation>
). Exuvium 8.3 mm long; entire dorsum intact; venter with abdominal ventrites anterior to AB II separated from tergum and reflexed, remainder of exuvium intact. Pupa 7.1 mm long, exarate, unsclerotized, color golden yellow. Pronotum projecting anteriorly, covering head; entire margin with very long setae. Elytra and abdominal segments I-IX with long setae on lateral margins. Abdominal tergites I-VIII each with two faint, transverse rows of round tubercles, at anterior 1/3 and near posterior margin, lateral to midline on each side. Abdominal tergite IX with apical margin nearly truncate, each posterolateral angle with a short spine. Paratergites separate; I reduced; II larger, projecting weakly anteriorly; III-VI each longer than II, projecting posteriorly, each with an anterobasal spiracular tubercle; VII similar but with a lateral spiracular tubercle. Ventrally, AB II-VI with sternopleural sutures; II-VIII each with a faint, transverse row of round tubercles near posterior margin.
</paragraph>
<caption id="FD0C2151ADE27E007BB140B483B030EB" doi="10.3897/zookeys.1164.103184.figures3-4" httpUri="https://binary.pensoft.net/fig/855186" pageId="0" pageNumber="23" start="Figures 3, 4" startId="F2">
<paragraph id="B6853CCB6ABB4B70193F9EACE50D51AD" pageId="0" pageNumber="23">
<emphasis id="05DF48113E1EA86A3E3EF8D39448050E" bold="true" pageId="0" pageNumber="23">Figures 3, 4.</emphasis>
<taxonomicName id="DC786669AC5BDF6273286A46D5AC512B" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="7621C8B519AA09F1579BDEE330DFA2C4" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
pupa and larval exuvium
<emphasis id="A28E8271EB86415FA01B4D0C893FF92B" bold="true" pageId="0" pageNumber="23">3</emphasis>
pupa
<emphasis id="8F481BA19C818F2EC67DE96DD7C35935" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="F36470CF04930E3544A66F72111F5090" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view
<emphasis id="1E50FBA72B08B3B1306953520457FF7D" bold="true" pageId="0" pageNumber="23">4</emphasis>
exuvium of last larval instar, ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="113D7535FCA166896936AB11AF5BF168" pageId="0" pageNumber="23" type="remarks">
<paragraph id="0C2232E69A797CB4B3409CA0B4083BB0" pageId="0" pageNumber="23">Remarks, pupa.</paragraph>
<paragraph id="2F1F3FC31827DCB7B9AC172FD86D0A6A" pageId="0" pageNumber="23">
This rare specimen was provided to WDS by
<normalizedToken id="D18A886C6A34BDA07064B5CCDB389F50" originalValue="Tomás">Tomas</normalizedToken>
<normalizedToken id="0FAA9BC6124154E4AF8A59FCAF537AA1" originalValue="Čekalović">Cekalovic</normalizedToken>
who collected it as a larva at Estero
<normalizedToken id="2F540446B6117D74E63B0D28D8A5083C" originalValue="Nonguén">Nonguen</normalizedToken>
in
<normalizedToken id="1765055ECCE9EF4CFE54D13422A3AD4C" originalValue="Concepción">Concepcion</normalizedToken>
Province (
<normalizedToken id="E10C6C2261619AA5AC8BBF97ED4B0450" originalValue="Región">Region</normalizedToken>
VIII,
<normalizedToken id="0A940A44B98431C10DDC24CEAF6CA0E9" originalValue="Bío">Bio</normalizedToken>
<normalizedToken id="B77656EADA1133DFEE02DF9C66B1838A" originalValue="Bío">Bio</normalizedToken>
) on 27 January 1996. He kept it alive for more than nine months until it pupated on 10 November 1996.
</paragraph>
<paragraph id="E53CFC285F8E0F815719B16EA109E442" pageId="0" pageNumber="23">
In comparison with sister genus
<taxonomicName id="2150D49E186A9A8FA58E870FE11F010A" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="E126508A74305B5FF90FC31926CD9C93" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
, pupal characters separating the two genera include the following: 1) gin traps in
<taxonomicName id="A1C4648EE6C23B650C6EA1568D8647F1" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C8F94FDD72110E4F4EF8D35BF44813F1" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
but absent in
<taxonomicName id="89BC80409BD88792A8801BFE0778D9FC" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="C492641B6B295D7E0F21F12814715C89" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
; 2) spiracle on abdominal paratergite II reduced in
<taxonomicName id="7440EE19FDDCD87631EACF7B1E5D872C" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="EB048B1E0F64D98D94CDD8CE73D53538" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
but not in
<taxonomicName id="E8FAFC06B52D7DBA0820FA98524956EB" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="61BAD92235DD3F1EF4767F8B897B2C28" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
; 3) spiracles on all paratergites located at the anterior base in
<taxonomicName id="34AD3FE2C9699BCBDCFEA065465A4894" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="72AF7F253620BDE00DEB1478F17FA408" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
but mid-dorsally in
<taxonomicName id="2BAB21EF28CB4FB03E75E48DE38A2235" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="2E455F9EC190C1B4DA730269FCC77994" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
; 4) paratergites in
<taxonomicName id="0EF973FDF1D93D955BC6658C8F6CAAC0" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="BAA2446920224D72914286AC3899F4EB" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
much longer than in
<taxonomicName id="A0FB3740C5CCC0B43B8D27B10C849235" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="FEB80028BA27204D4CC0DBA41AFB2F6B" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
; and 5) apex of abdominal segment IX with a median projection in
<taxonomicName id="22D389F89033BC70C832AED6E443677A" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="BB60F47A76DCBFD0436CD997431BCC1A" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
but lacking in
<taxonomicName id="77DE8558EEBC8FBFA07A68439CC350E6" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="61299DBAADED19F267614B331730CDD0" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
. The latter two characters probably aid the pupa of
<taxonomicName id="08ED9B62BA29E5477A39812676CCC9B9" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sclerocyphon" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="05CCE4343834C3C757BF06A7FB8C7BA7" italics="true" pageId="0" pageNumber="23">Sclerocyphon</emphasis>
</taxonomicName>
in crawling out from under the larval exuvium (
<bibRefCitation id="C8570A069252E2BAFF7E274DE8D7A16B" DOI="https://doi.org/10.1111/j.1440-6055.1981.tb01048.x" author="Smith, JA" journalOrPublisher="Journal of the Australian Entomological Society" pageId="0" pageNumber="23" pagination="277 - 288" refId="B21" refString="Smith, JA, 1981. Two Tasmanian species of Sclerocyphon Blackburn (Coleoptera: Psephenidae) with notes on their life history and distribution. Journal of the Australian Entomological Society 20 (4): 277 - 288, DOI: https://doi.org/10.1111/j.1440-6055.1981.tb01048.x" title="Two Tasmanian species of Sclerocyphon Blackburn (Coleoptera: Psephenidae) with notes on their life history and distribution." url="https://doi.org/10.1111/j.1440-6055.1981.tb01048.x" volume="20" year="1981">Smith 1981</bibRefCitation>
;
<bibRefCitation id="3526C39D1FD00BD01D7EB98802D66BFD" DOI="https://doi.org/10.1071/AJZS119" author="Davis, JA" journalOrPublisher="United States Printing Office, Washington, D. C." pageId="0" pageNumber="23" refId="B6" refString="Davis, JA, 1986. Revision of the Australian Psephenidae (Coleoptera): Systematics, phylogeny and historical biogeography. Australian Journal of Zoology. Supplementary Series (119): 1-97. https://doi.org/10.1071/AJZS119" title="Revision of the Australian Psephenidae (Coleoptera): Systematics, phylogeny and historical biogeography. Australian Journal of Zoology. Supplementary Series (119): 1 - 97." url="https://doi.org/10.1071/AJZS119" year="1986">Davis 1986</bibRefCitation>
). The pupa of
<taxonomicName id="B7911C0301EC8F6F057E873EC9A60F0E" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="BD1F615E0281CA5F7F466EE39254162D" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
remains under the larval exuvium until the adult emerges.
</paragraph>
</subSubSection>
<subSubSection id="4FE63FD1DC24F6CC520147DD1E8CA134" pageId="0" pageNumber="23" type="tychepsephus geographic and seasonal distribution">
<paragraph id="EA20D36B44B7FA9651C2D363C1E50754" pageId="0" pageNumber="23">
<taxonomicName id="3CDDDA7E8BB955C5EAC54C07D402A086" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="E58859776DA9107AE6557BB1E9D59C79" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
geographic and seasonal distribution
</paragraph>
<paragraph id="CF4672BDBFC852FB95C26877CD98CB3C" pageId="0" pageNumber="23">
In Chile, published localities describe
<taxonomicName id="F4613378E23F28CD09634CFC4843ECFB" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="9F2A6EB70D533C1C52CA85FA89ED66B2" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
occurring from
<normalizedToken id="81583413DCC4DE2583D7E57E2BA6D586" originalValue="Peñaflor">Penaflor</normalizedToken>
(
<normalizedToken id="61EA26A0500768A678A2D0F702095B3D" originalValue="Región">Region</normalizedToken>
Metropolitana) in central Chile (
<bibRefCitation id="6388A9C6F4EB562BF608E1317CA5E75A" author="Lataste, F" journalOrPublisher="Actes de la Societe Scientifique du Chili" pageId="0" pageNumber="23" pagination="103 - 106" refId="B11" refString="Lataste, F, 1897a. Presentation d'une larve de Pseudo-Nevroptere d'aspect crustaceen. Actes de la Societe Scientifique du Chili 7: 103 - 106" title="Presentation d'une larve de Pseudo-Nevroptere d'aspect crustaceen." volume="7" year="1897 a">Lataste 1897a</bibRefCitation>
), south to
<normalizedToken id="89DE46479C6EDED2DFAE89ADE80F475A" originalValue="Chillán">Chillan</normalizedToken>
,
<normalizedToken id="8D5FC9F8BADDB3F15BB469701980BF61" originalValue="Tomé">Tome</normalizedToken>
and Arauco (
<normalizedToken id="2AC75C8A5C43ED820FF3BCA075CA6F06" originalValue="Región">Region</normalizedToken>
VIII,
<normalizedToken id="147654A3023D5E9622FD1F9AFA6CFD01" originalValue="Bío">Bio</normalizedToken>
<normalizedToken id="931182406E14F883899606124D601169" originalValue="Bío">Bio</normalizedToken>
;
<normalizedToken id="8986835039E6A68A8833AC89655EE2B7" originalValue="Región">Region</normalizedToken>
XVI,
<normalizedToken id="6091783E65F38344D9E2CF477824EC08" originalValue="Ñuble">Nuble</normalizedToken>
) (
<bibRefCitation id="6F620394EEDA46872222D3D5272AAA28" author="Artigas, J" journalOrPublisher="Gayana (Concepcion)" pageId="0" pageNumber="23" pagination="3 - 11" refId="B1" refString="Artigas, J, 1963. Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae). Gayana (Concepcion) 1963 (8): 3 - 11" title="Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae)." volume="1963" year="1963">Artigas 1963</bibRefCitation>
), and west of Puerto Varas (
<normalizedToken id="A3B68062225077F9B3BFC57C7919843D" originalValue="Región">Region</normalizedToken>
X, Los Lagos) (
<bibRefCitation id="011BE0584F51A397BC225ECAABF785B9" DOI="https://doi.org/10.1093/aesa/80.6.865" author="Ashworth, AC" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="23" pagination="865 - 895" refId="B2" refString="Ashworth, AC, Hoganson, JW, 1987. Coleoptera bioassociations along an elevational gradient in the Lake Region of southern Chile, and comments on the postglacial development of the fauna. Annals of the Entomological Society of America 80 (6): 865 - 895, DOI: https://doi.org/10.1093/aesa/80.6.865" title="Coleoptera bioassociations along an elevational gradient in the Lake Region of southern Chile, and comments on the postglacial development of the fauna." url="https://doi.org/10.1093/aesa/80.6.865" volume="80" year="1987">Ashworth and Hoganson 1987</bibRefCitation>
). Data records from the MNNC include adult specimens from the Andes and foothills in Reserva Nacional Altos de Lircay (
<normalizedToken id="A5E09DB8238B7F33EC380A899CFC8B0A" originalValue="Región">Region</normalizedToken>
VII, del Maule), Cordillera
<normalizedToken id="580F7E545EF5D52B65B9D277896F4617" originalValue="Chillán">Chillan</normalizedToken>
(
<normalizedToken id="98DA3166219F59EE7C9302271191D5C6" originalValue="Región">Region</normalizedToken>
XVI,
<normalizedToken id="A79BFCDA13434A44A02DC6D6A2FE5E45" originalValue="Ñuble">Nuble</normalizedToken>
), Parque Nacional
<normalizedToken id="12995FD8DCA73DCC6FD013DAD0DB1EEB" originalValue="Conguillío">Conguillio</normalizedToken>
(
<normalizedToken id="6B81CDB0D72A09D1ABA680D3BCD3371B" originalValue="Región">Region</normalizedToken>
IX,
<normalizedToken id="49F8D4006194C1B90AB85DA7781BC531" originalValue="Araucanía">Araucania</normalizedToken>
) and Parque Nacional Vicente
<normalizedToken id="A82C2E1B34548F318EFB5500D8F3611A" originalValue="Pérez">Perez</normalizedToken>
<taxonomicName id="50A70A68CA9C6BDA4B0A5734B610A7A6" authorityName="Waterhouse" authorityYear="1876" kingdom="Animalia" lsidName="" order="Rosales" pageId="0" pageNumber="23" rank="order">Rosales</taxonomicName>
(
<normalizedToken id="7199BEF455BD856040CCA132AB31B1AD" originalValue="Región">Region</normalizedToken>
X, Los Lagos), as well as specimens from near the Pacific Coast in Quirihue (
<normalizedToken id="30738A261CFFE3CB5359AAB294C500EA" originalValue="Región">Region</normalizedToken>
XVI,
<normalizedToken id="226C8BDA08E276B393DBE287815940A9" originalValue="Ñuble">Nuble</normalizedToken>
) and Valdivia (
<normalizedToken id="A1BFBE8D0D7BE41CADA2CE64EA60E664" originalValue="Región">Region</normalizedToken>
XIV, Los
<normalizedToken id="BBBEB5156267BDBF3D588750DB4E621F" originalValue="Ríos">Rios</normalizedToken>
) (M. Elgueta, in litt.). See Appendix 2 for detailed locality information regarding the above records. Collections by the authors of adult and larval
<taxonomicName id="C37740FA6618547F97DFB7E031139C7D" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="A10786218852DD53E87CE50BC7F7638C" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
, from regions VIII (
<normalizedToken id="0242EE5AC94B2198E5FC5F0B23C0B9AC" originalValue="Bío">Bio</normalizedToken>
<normalizedToken id="21CC6F9ABA9D8BD89A72FB8DC3CFFEBA" originalValue="Bío">Bio</normalizedToken>
) through XI (
<normalizedToken id="8A26413EE4EE3E5E16B3FB6040DBEB6A" originalValue="Aysén">Aysen</normalizedToken>
), plus regions XIV (Los
<normalizedToken id="83D72183F1444B1513F520AD3B85EE76" originalValue="Ríos">Rios</normalizedToken>
) and XVI (
<normalizedToken id="60C41F968D44DFC11B581420519F5508" originalValue="Ñuble">Nuble</normalizedToken>
), are listed in Appendix 1.
</paragraph>
<paragraph id="AEAC18C28288B14801BB09B3471646D4" pageId="0" pageNumber="23">
Larval records from Provincia del
<normalizedToken id="12BDD8CD5220EB28FAEC0C2E11540DD1" originalValue="Neuquén">Neuquen</normalizedToken>
, Argentina, indicate that
<taxonomicName id="43A6E8D399DB67F1ED137ED77FE219FB" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="89CE7FC0E9396C99BEE4E720BF568C8E" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
also occurs on the east front of the Andes.
<bibRefCitation id="81FAD3F22FB378E4BE6EC5A912B57955" author="Wais, IR" journalOrPublisher="Acta Limnologica Brasiliensia" pageId="0" pageNumber="23" pagination="829 - 845" refId="B23" refString="Wais, IR, 1990. A checklist of the benthic macroinvertebrates of the Negro River Basin, Patagonia, Argentina, including an approach to their functional feeding groups. Acta Limnologica Brasiliensia 3: 829 - 845" title="A checklist of the benthic macroinvertebrates of the Negro River Basin, Patagonia, Argentina, including an approach to their functional feeding groups." volume="3" year="1990">Wais (1990</bibRefCitation>
,
<bibRefCitation id="363B372CB4B9F90F7C682DFBC9393D97" author="Wais, IR" journalOrPublisher="Boletin Mosquero" pageId="0" pageNumber="23" pagination="27 - 33" refId="B24" refString="Wais, IR, 1995. El alimento - presa que encontraron las truchas en la Argentina. Boletin Mosquero 19: 27 - 33" title="El alimento - presa que encontraron las truchas en la Argentina." volume="19" year="1995">1995</bibRefCitation>
) reported T
<emphasis id="A9CBB83B7E86EBC25124EE5E3DA27CD6" italics="true" pageId="0" pageNumber="23">ychepsephus</emphasis>
(listed as
<taxonomicName id="4DA0FFE6E159F15CEC9965770F694047" class="Insecta" family="Psephenidae" genus="Chilectopria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chilectopria grandis" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="BA38BF7E584D3347A695CD76BE2CB7AB" italics="true" pageId="0" pageNumber="23">Chilectopria grandis</emphasis>
</taxonomicName>
) from the Rio Meliquina in the Rio Negro Basin north of San Carlos de Bariloche.
<normalizedToken id="56E3C1293DA294A5C924083884CDB611" originalValue="Nicolás">Nicolas</normalizedToken>
<normalizedToken id="D119375018BD48801306DAA4A8093E8B" originalValue="Román">Roman</normalizedToken>
(N.
<normalizedToken id="E6F42398557F4CAF49A6EFEEECBF7B9D" originalValue="Román">Roman</normalizedToken>
, in litt.) has more recently reported finding a larva (morph 2) (Fig.
<figureCitation id="319591803AFFC61F7B50EE143F01A27F" captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Non-Chilean larvae 5 Tychepsephus larva from Argentina a dorsal view b ventral view. Images provided by Nicolas Roman (CONICET) 6 larva from French Guiana, possibly Tychepsephus a dorsal view b ventral view. Images provided by Robert Sites (UMC)." figureDoi="10.3897/zookeys.1164.103184.figures5-6" httpUri="https://binary.pensoft.net/fig/855187" pageId="0" pageNumber="23">5</figureCitation>
) near San Martin de los Andes. More surprisingly, in the EMEC there is a larval eubriine from French Guiana (Fig.
<figureCitation id="683D2D254DFC7BDD8AACF9B4AE031F39" captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Non-Chilean larvae 5 Tychepsephus larva from Argentina a dorsal view b ventral view. Images provided by Nicolas Roman (CONICET) 6 larva from French Guiana, possibly Tychepsephus a dorsal view b ventral view. Images provided by Robert Sites (UMC)." figureDoi="10.3897/zookeys.1164.103184.figures5-6" httpUri="https://binary.pensoft.net/fig/855187" pageId="0" pageNumber="23">6</figureCitation>
) that greatly resembles the larva of
<taxonomicName id="C074E23E2F81EA0622FA18C9028BB7AA" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="4CCF9445570646FE2C5D38852A784057" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
. If this is actually a
<taxonomicName id="BB8B627720CEDDD2B28737EBE3F71203" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="11424D2F516C933E35B243B7E63B0AF6" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
larva, the geographic distribution of the genus would be significantly expanded (see Discussion).
</paragraph>
<caption id="9CA0394CCCBFF8DA1E1F9EFDD4DA4A06" doi="10.3897/zookeys.1164.103184.figures5-6" httpUri="https://binary.pensoft.net/fig/855187" pageId="0" pageNumber="23" start="Figures 5, 6" startId="F3">
<paragraph id="E5D7CDB75C4C7204619BAA0004C2FACF" pageId="0" pageNumber="23">
<emphasis id="F5FD9E95AEA6E3BED061AFD81C0AF213" bold="true" pageId="0" pageNumber="23">Figures 5, 6.</emphasis>
Non-Chilean larvae
<emphasis id="5E03D76A8F7864922C26760B3005CCCE" bold="true" pageId="0" pageNumber="23">5</emphasis>
<taxonomicName id="5F7F6651AF59566CCDA6E79712D41A22" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="FB635FACAB88C222382D2DE63D412615" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
larva from Argentina
<emphasis id="5609A4ED5B603BCFF22637A9BB1103F1" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="32A6048D7557E13DE640FABDAE871277" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view. Images provided by
<normalizedToken id="E5E088EAA450EAC7C3C59C39CA7160FE" originalValue="Nicolás">Nicolas</normalizedToken>
<normalizedToken id="045ABD83BECE3D4CB0A7BBF83B107ED1" originalValue="Román">Roman</normalizedToken>
(CONICET)
<emphasis id="DDED4570A8EDD4E73E07DD94456E9AB2" bold="true" pageId="0" pageNumber="23">6</emphasis>
larva from French Guiana, possibly
<taxonomicName id="8CC958D0C78F25C656AE88ED5620A181" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="54773747AE34C4228CF3D7D8DE1A7B18" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
<emphasis id="B98FD1FA047C2851C0E5856967637C47" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="CF5EDEB1B88BA2DC382E174A4F3594D5" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view. Images provided by Robert Sites (UMC).
</paragraph>
</caption>
<paragraph id="838875B3CEA01C076A8F22B9FEE14501" pageId="0" pageNumber="23">
A summation of the locality data for adults and larvae reveals a geographic distribution of
<taxonomicName id="E159CC312C5D909B389405594AFEF512" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="2AA231FC4E8A36E1DD94E4A6E461B789" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
in the middle third of Chile, including both the Andes and the Coast Range, and on the eastern front of the central Andes in
<normalizedToken id="1A8DF8DFACC397AE29679FA0EED2930E" originalValue="Neuquén">Neuquen</normalizedToken>
Province, Argentina (Appendices 1, 2). The distribution map (Fig.
<figureCitation id="C110E96853E3885D32178551E907CFB0" captionStart="Figure 7" captionStartId="F4" captionText="Figure 7. Map of Chile with enlarged section showing the geographic distribution of species-verified adult Tychepsephus specimens: T. cekalovici, yellow triangles; T. felix, violet squares; both species, red circles." figureDoi="10.3897/zookeys.1164.103184.figure7" httpUri="https://binary.pensoft.net/fig/855188" pageId="0" pageNumber="23">7</figureCitation>
) represents species-verified adult records only.
</paragraph>
<caption id="8237210C57384F7F21EF7FE7D5447A60" doi="10.3897/zookeys.1164.103184.figure7" httpUri="https://binary.pensoft.net/fig/855188" pageId="0" pageNumber="23" start="Figure 7" startId="F4">
<paragraph id="50A4EC0F724D739CEC7A24F3491A8A8F" pageId="0" pageNumber="23">
<emphasis id="FE921915BE815684B6715D48ADF0D71B" bold="true" pageId="0" pageNumber="23">Figure 7.</emphasis>
Map of Chile with enlarged section showing the geographic distribution of species-verified adult
<taxonomicName id="ACD26A2E6FE859EAE2508FF9C1A450CF" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="52E1FB4014DBE525C762414BC5F07089" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
specimens:
<taxonomicName id="CF61275183C492AB0DAAFD75853F3537" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="68D05FF400EACF868D90A465CCF409CD" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
, yellow triangles;
<taxonomicName id="591166C10B146181759CB9C10E88A012" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="2329DB75C9779783A05063AC57AE5756" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
, violet squares; both species, red circles.
</paragraph>
</caption>
<paragraph id="9B940D7F336FAA5F3D333B77A76DAB70" pageId="0" pageNumber="23">
The adults examined for this study were collected from November through January, during the austral summer. Larvae are present year-round.
<normalizedToken id="82AB835E15F5518926648D8DF423A77A" originalValue="Tomás">Tomas</normalizedToken>
<normalizedToken id="6DC41B6509DC77127E9BA07F8AC741A5" originalValue="Čekalović">Cekalovic</normalizedToken>
collected larvae in January and August, and
<bibRefCitation id="22BE651039FA51DB52ABCF3173E8C31C" DOI="https://doi.org/10.4067/S0718-19572012000100003" author="Fierro, P" journalOrPublisher="Revista de Biologia Marina y Oceanografia" pageId="0" pageNumber="23" pagination="23 - 33" refId="B9" refString="Fierro, P, Bertran, C, Mercado, M, Pena-Cortes, F, Tapia, J, Hauenstein, E, Vargas-Chacoff, L, 2012. Benthic macroinvertebrate assemblages as indicators of water quality applying a modified biotic index in a spatio-seasonal context in a coastal basin of southern Chile. Revista de Biologia Marina y Oceanografia 47 (1): 23 - 33, DOI: https://doi.org/10.4067/S0718-19572012000100003" title="Benthic macroinvertebrate assemblages as indicators of water quality applying a modified biotic index in a spatio-seasonal context in a coastal basin of southern Chile." url="https://doi.org/10.4067/S0718-19572012000100003" volume="47" year="2012">Fierro et al. (2012)</bibRefCitation>
reported collecting larval
<taxonomicName id="B79A178DA1A4394DD3372D6D97BE0E4D" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="8CE7A9BC258929396013144CC113A11B" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
in all seasons except winter.
</paragraph>
</subSubSection>
<subSubSection id="BF564BF41AD70AED2C76FEC6697F421E" pageId="0" pageNumber="23" type="biology_ecology">
<paragraph id="A011B8797B6890C3F34E4D8B96D9FFE4" pageId="0" pageNumber="23">
<taxonomicName id="32E982D3ED0475351EA3513B6EFA8CD5" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="19EF270BFD1BCF07C10FB2C06641EAC4" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
habitat
</paragraph>
<paragraph id="C9DDC717697A4C2023073545212173C9" pageId="0" pageNumber="23">
We have collected larvae and/or adults of
<taxonomicName id="E43C4AA795C336EE1024825463605E90" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="E61271C2E97E9CA75CE5D8B23B265E2C" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
across an elevational range of 15-1685 m at streams and rivers in Chile. In general, these watercourses were small to medium-sized and rather shallow, with moderate current, and with clear or often brown, tannin-stained water. This is in contrast to
<bibRefCitation id="BB4F877B990C0164BFD6D450CA518663" DOI="https://doi.org/10.5772/intechopen.89077" author="Zarges, CV" editor="Devlin, A" journalOrPublisher="IntechOpen, London" pageId="0" pageNumber="23" pagination="1 - 30" refId="B27" refString="Zarges, CV, Fierro, P, Olmos, V, 2019. Freshwater invertebrates of southwestern South America: Diversity, biogeography, and threats. In: Devlin, A, Pan, J, Shah, MM, Eds., Inland Waters - Dynamics and Ecology. IntechOpen, London: 1 - 30, DOI: https://doi.org/10.5772/intechopen.89077" title="Freshwater invertebrates of southwestern South America: Diversity, biogeography, and threats." url="https://doi.org/10.5772/intechopen.89077" volumeTitle="Inland Waters - Dynamics and Ecology." year="2019">Zarges et al. (2019)</bibRefCitation>
who reported them from areas with &quot;high slope and high water currents.&quot; Label data from specimens collected by
<normalizedToken id="6D4C1AEBBF02B00C5824BFA49A94059B" originalValue="Tomás">Tomas</normalizedToken>
<normalizedToken id="FBDB44AE327F51BD6A3D6080AE69263F" originalValue="Čekalović">Cekalovic</normalizedToken>
show larvae living in temperatures of 10-14 °C. Surprisingly, some them were found in humus under
<taxonomicName id="3A9D07AFE75529561D98C061A5891785" class="Liliopsida" family="Poaceae" genus="Chusquea" kingdom="Plantae" lsidName="Chusquea quila" order="Poales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="quila">
<emphasis id="1394D07FDC30EC050DBD9862A075E6F2" italics="true" pageId="0" pageNumber="23">Chusquea quila</emphasis>
</taxonomicName>
[Kunth (
<taxonomicName id="3F5EE9CC2924BAAA9D5C34E9A1FB1CE5" authorityName="Kunth" authorityYear="1822" class="Liliopsida" family="Poaceae" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) bamboo], 2-3 m from a river.
</paragraph>
<paragraph id="17B2DE64682E7525302F7E5653633E4D" pageId="0" pageNumber="23">
Examples of the small to medium-sized streams and small, shallow rivers in which adult
<taxonomicName id="9F2EF540939D7B9111CCF2B326600CA4" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="CDD365D5E9677B641AC6283FB355DA70" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
have been collected by the authors, described below, include
<normalizedToken id="479A522BD8BDE6BF16372EDBE10AC83F" originalValue="Río">Rio</normalizedToken>
Colegual,
<normalizedToken id="7910D5AE5F774368D43EED984A36A6D1" originalValue="Río">Rio</normalizedToken>
Contaco,
<normalizedToken id="657B9614DB6915665960CFCE82C49C64" originalValue="Río">Rio</normalizedToken>
Oroco, and an unnamed tributary of the
<normalizedToken id="5E6A5E47C144E5EA25BB791030496E11" originalValue="Río">Rio</normalizedToken>
Blanco (Figs
<figureCitation id="EF9C4CB6227D78071A975115DDAFDE3C" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">8</figureCitation>
,
<figureCitation id="290DE741DF8FE3767184A8E873076E36" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">9</figureCitation>
,
<figureCitation id="E08414883370003D496CB731A6855850" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">11</figureCitation>
).
</paragraph>
<paragraph id="DDAC65CA43105309E9216AAD42C7574E" pageId="0" pageNumber="23">
<emphasis id="956960DD9EDF0EFAC6AAC3F1EB285E93" bold="true" pageId="0" pageNumber="23">
<normalizedToken id="4756029556C6A9DC442BC64E516DB691" originalValue="Río">Rio</normalizedToken>
Colegual
</emphasis>
(Fig.
<figureCitation id="F69EE1F2F8F790CF1F326E3B74A0614F" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">8</figureCitation>
), located west of Puerto Varas at an elevation of ~ 200 m, is a medium-sized stream with moderate to slow flow over cobble and gravel coated with brown algae. Pools with laminar water flow are interrupted by areas of shallow riffles. Riparian vegetation overhangs some of the banks. At the time of sampling, the water was cool, clear, and brown-stained (Fig.
<figureCitation id="6899F9722342F6D50804C57C3DEB3D4D" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">8b</figureCitation>
). The watercourse is situated in mostly flat terrain bordered by cleared fields (Fig.
<figureCitation id="EFFFFD0540EE4CAB0A98274CA4735833" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">8a</figureCitation>
) and is partly shaded at the collection site near the bridge. Adults of two species were readily collected by sweeping and beating the riparian vegetation, which consisted of willows, streamside grasses and forbs. Blacklight sampling yielded no specimens. Many larvae were present in the substrate of the riffles, including both morphs.
</paragraph>
<paragraph id="5B799A087C8FCB4ECE1AB7AD3D91DB0E" pageId="0" pageNumber="23">
<emphasis id="8D8B770682D89EE388E435C60E113D80" bold="true" pageId="0" pageNumber="23">
<normalizedToken id="5065102F9B8B767F1BDC8E2A808752C8" originalValue="Río">Rio</normalizedToken>
Contaco (=
<normalizedToken id="A9CB1A4A5756A2102C49BB02B46E3716" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="BF79BFC331C831434C7D7C09618F81E6" originalValue="Tranallaquín">Tranallaquin</normalizedToken>
)
</emphasis>
(Fig.
<figureCitation id="755258F577D07D3AF4D318059410FEF8" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">9</figureCitation>
), located west of Osorno at an elevation of ~ 160 m, is a medium-sized stream with clear water, moderate flow, and a substrate of sand, gravel and rubble with submerged mosses. Larvae were very abundant in the substrate of the riffles, as they also were at a small tributary of the
<normalizedToken id="1115533C75D8F3B38EA2CE1C3FBDF534" originalValue="Río">Rio</normalizedToken>
Contaco at Puente El Avion (Fig.
<figureCitation id="0C0DD0411AD219144976EA4A199C01B8" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">10</figureCitation>
).
</paragraph>
<caption id="D6DCB89572DD3DFDF8DB3FB0EBBABDE0" doi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23" start="Figures 811" startId="F5">
<paragraph id="B9AD28F853DB27804F7BD41070BCF89C" pageId="0" pageNumber="23">
<emphasis id="9F2AAB3B002C492909A6D73A1ED33664" bold="true" pageId="0" pageNumber="23">Figures 8-11.</emphasis>
<taxonomicName id="2FDCC6860AE8E9CE45D5FD1A996E979E" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="CA54216D1B2EBD1C45736E77BAB89A18" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
habitat
<emphasis id="3C2CB8AD4F00E54DF2E56D333DD88710" bold="true" pageId="0" pageNumber="23">8</emphasis>
<normalizedToken id="865C8707697D1AA2393D4843634FBE11" originalValue="Río">Rio</normalizedToken>
Colegual,
<taxonomicName id="4941972CB378DBD71138A526FD412B23" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="C3C84902ACC2501F8D9080BD29BF69F2" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
type locality
<emphasis id="21FE07487324374EC16BDC0E70CF1342" bold="true" pageId="0" pageNumber="23">a</emphasis>
stream with riffle-pool morphology flowing through a mixture of forest and pasture land
<emphasis id="40F29B9471A85731B04A7C1A6384351C" bold="true" pageId="0" pageNumber="23">b</emphasis>
streamside shrubby vegetation, adult habitat
<emphasis id="B593483204F35900CDE4F82CC925A27E" bold="true" pageId="0" pageNumber="23">9</emphasis>
<normalizedToken id="34739D62F05E66A75F9DEB04CA7D964C" originalValue="Río">Rio</normalizedToken>
Contaco
<emphasis id="544CA12A934AE33048CFCF1CDB4B35AE" bold="true" pageId="0" pageNumber="23">10</emphasis>
stream at Puente El Avion
<emphasis id="306142A9CC3F1612C38F4249D9D0ED60" bold="true" pageId="0" pageNumber="23">11</emphasis>
tributary
<normalizedToken id="E4CB64D8867942188352ACEBFA04D9C7" originalValue="Río">Rio</normalizedToken>
Blanco.
</paragraph>
</caption>
<paragraph id="C74E2E1DEBB027032ADDFB8D7F1E679B" pageId="0" pageNumber="23">
<emphasis id="F2BC47A94CFDE7E92BDD20EE10E0F331" bold="true" pageId="0" pageNumber="23">
<normalizedToken id="5292B7C57E0E3F5550BEA201C6961E2F" originalValue="Río">Rio</normalizedToken>
Oroco
</emphasis>
at Puente Hondo, located east of Puerto Montt at an elevation of ~ 30 m, is small and shallow, with cool, brown-stained water and a sand and cobble substrate with aquatic moss.
</paragraph>
<paragraph id="F7C4C998D165CEE61A02FE847E42EF51" pageId="0" pageNumber="23">
<emphasis id="6E66B2EF3AD5F35F7CC852283F11E5F5" bold="true" pageId="0" pageNumber="23">
An unnamed tributary of
<normalizedToken id="FFDC4D50D24C7B4AF91BE70AF299390E" originalValue="Río">Rio</normalizedToken>
Blanco
</emphasis>
(Fig.
<figureCitation id="686E0D67C2AC3EA2AD804FF0F171EB0C" captionStart="Figures 811" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">11</figureCitation>
), located between Caleta Gonzalo and Caleta Santa
<normalizedToken id="B7FF6963CE8355C6E477ECC7DA266577" originalValue="Bárbara">Barbara</normalizedToken>
(north of
<normalizedToken id="50CA60653FDACA7376825A8F016E67BD" originalValue="Chaitén">Chaiten</normalizedToken>
) at an elevation of ~ 160 m, yielded one adult and a small number of larvae. The stream is small, clear, and very cold with moderate current, shallow riffles, and knee-deep pools. It has a substrate of cobbles and sandy gravel with some aquatic moss present.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="8244BC13B6B90321D90F9C9F1F03DCA8">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
</mods:titleInfo>
<mods:part id="5FDFD2C3F4405FCECB65F1CC00501055">
<mods:extent id="CDC8F4EC797574E90E57CA045FC3E4FA" unit="page">
<mods:start id="52872032CF02F7EECF8CA6891A31C97E">1189</mods:start>
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<mods:identifier id="A671747AF340CDA166BF074BD5709DDB" type="ISBN">0-8018-8221-4</mods:identifier>
<mods:identifier id="9AD8C57DDA8C1710D232D06D0279619B" type="DOI">10.5281/zenodo.7316535</mods:identifier>
</mods:mods>
<treatment id="C7BAD2001B010DF0568EE12DE067EAA8" ID-DOI="http://doi.org/10.5281/zenodo.11335596" ID-GBIF-Taxon="231539569" ID-Zenodo-Dep="11335596" LSID="urn:lsid:plazi:treatment:C7BAD2001B010DF0568EE12DE067EAA8" httpUri="http://treatment.plazi.org/id/C7BAD2001B010DF0568EE12DE067EAA8" lastPageNumber="1499" pageNumber="1499" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13001807">
<subSubSection id="B46F7A44D58355B3C2C08ADB4A30E656" pageNumber="1499" type="nomenclature">
<paragraph id="ED83C12E9FE4EC1E9848F974265F3F36" pageNumber="1499">
<taxonomicName id="3EF7F55EA9D1B5E0498D08DF338DF88A" ID-CoL="4Y823" authority="Flannery and Wickler 1990" authorityName="Flannery and Wickler" authorityYear="1990" class="Mammalia" family="Muridae" genus="Solomys" kingdom="Animalia" order="Rodentia" pageNumber="1499" phylum="Chordata" rank="species" species="spriggsarum">
<emphasis id="FDC03990D667858D26D93D4C1C5FE9AF" italics="true" pageNumber="1499">Solomys spriggsarum</emphasis>
Flannery and Wickler 1990
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="72D2C44340D73C7A0ACE39E37030D214" pageNumber="1499" type="reference_group">
<paragraph id="BFCA8BC8CA707A1C98D6F7922A694EF8" pageNumber="1499">
<treatmentCitation id="FD110C6C512C86B9FE863B2C325504FE" author="Flannery and Wickler" authority="Flannery and Wickler 1990" authorityName="Flannery and Wickler" authorityYear="1990" class="Mammalia" family="Muridae" genus="Solomys" journal="Aust. Mammal." kingdom="Animalia" order="Rodentia" page="133" pageNumber="1499" phylum="Chordata" rank="species" species="spriggsarum" title="Solomys spriggsarum" volume="13" year="1990">
<taxonomicName id="911E22F2E39D57A324EE70BE189052CC" ID-CoL="4Y823" authority="Flannery and Wickler 1990" authorityName="Flannery and Wickler" authorityYear="1990" class="Mammalia" family="Muridae" genus="Solomys" kingdom="Animalia" order="Rodentia" pageNumber="1499" phylum="Chordata" rank="species" species="spriggsarum">
<emphasis id="76FF3E0C8860E613DFD692756191F791" italics="true" pageNumber="1499">Solomys spriggsarum</emphasis>
<bibRefCitation id="3273789228BC0A5B3DE463448BE0CDFB" author="Flannery, T. F. &amp; S. Wickler" refId="ref103915" refString="Flannery, T. F., and S. Wickler. 1990. Quaternary murids (Rodentia: Muridae) from Buka Island, Papua New Guinea, with descriptions of two new species. Australian Mammalogy, 13: 127 - 139." year="1990">Flannery and Wickler 1990</bibRefCitation>
</taxonomicName>
,
<bibCitation id="166FCF7887405E77D7379D6CEE419156" author="Flannery and Wickler" journal="Aust. Mammal." pageNumber="1499" pagination="133" title="Solomys spriggsarum" volume="13" year="1990">
<collectingRegion id="F0E0DC9342083B20561656A06BF317EB" country="Cameroon" name="East">Aust</collectingRegion>
. Mammal., 13: 133
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A48E3F86EDBEBD2122E66E2C1C9E8ED0" pageNumber="1499" type="materials_examined">
<paragraph id="DC8F70EDDC551815E92A70F639A8E4F0" pageNumber="1499">
<emphasis id="FE5742D154887CF7831DA376E6820248" inLineHeading="true" pageNumber="1499">Type Locality:</emphasis>
<materialsCitation id="3E0913EF3DC5F642C31D8FA3F7C28348" ID-GBIF-Occurrence="4867324553" pageNumber="1499">Solomon Isls, Buka Isl, Kilu rockshelter.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8B026EAB73DDF77C8445F728C34C683B" pageNumber="1499" type="vernacular_names">
<paragraph id="6A8DB90C509C1E5272638ED8868D60B0" pageNumber="1499">
<emphasis id="1AEC76BCF4773BD8B624D0F344FCD917" inLineHeading="true" pageNumber="1499">Vernacular Names:</emphasis>
<vernacularName id="737E867B6C67066C355F3C52BD4678B4" language="eng" pageNumber="1499">Buka Island Solomys</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="FF17C7B4B554A9F563AC4DBCD986A747" pageNumber="1499" type="distribution">
<paragraph id="9DE2C01387B2A97233D21FF34D6282F3" pageNumber="1499">
<emphasis id="94552C54C83E2A18969CE635EB036855" inLineHeading="true" pageNumber="1499">Distribution:</emphasis>
Endemic to Buka Isl.
</paragraph>
</subSubSection>
<subSubSection id="4D13E2AC74E1CD7F18236457962DCFCD" pageNumber="1499" type="discussion">
<paragraph id="3BC31073E73FB58698B01E9FA2FDCC63" pageNumber="1499">
<emphasis id="3F39A803507229FFD2AD3B0FDC09E9E1" inLineHeading="true" pageNumber="1499">Discussion:</emphasis>
Known only by subfossil archaeological material (
<bibRefCitation id="CFD8885DA42F3F17016402D1D59024CF" author="Flannery, T. F. &amp; S. Wickler" refId="ref103915" refString="Flannery, T. F., and S. Wickler. 1990. Quaternary murids (Rodentia: Muridae) from Buka Island, Papua New Guinea, with descriptions of two new species. Australian Mammalogy, 13: 127 - 139." year="1990">Flannery and Wickler, 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>