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<figureCitation id="131E2A55FFE0FFBAFD53F90021D3F932" box="[682,750,1705,1731]" captionStart="Fig" captionStartId="21.[189,232,1788,1814]" captionTargetBox="[228,1364,265,1702]" captionTargetId="figure@21.[228,1364,265,1743]" captionTargetPageId="21" captionText="Fig. 6. AD. Aphyosemion aurantiacum Chirio, Legros &amp; Agnèse sp. nov. A. Adult, ♁, from locality 12, not preserved. Photo O. Buisson. B. Adult, ♀, from locality 12, not preserved. Photo O. Buisson. C. Holotype, adult, ♁, from Wézé Spring (MRAC 2016-019-P-64). D. Paratype, adult, ♀, from Wézé Spring (MRAC 2016-019-P-65-73). EH. A. rubrogaster Chirio, Legros &amp; Agnèse sp. nov. E. Adult, ♁, from locality 16, not preserved. Photo O. Buisson. F. Adult, ♀, from locality 16, not preserved. Photo O. Buisson. G. Holotype, adult, ♁, from the Niengé River (MRAC 2016-019-P-93). H. Paratype, adult, ♀, from the Niengé River (MRAC 2016-019-P-94-108)." figureDoi="http://doi.org/10.5281/zenodo.3825065" httpUri="https://zenodo.org/record/3825065/files/figure.png" pageId="20" pageNumber="21">Fig. 6</figureCitation>
EH,
<tableCitation id="C6A7036BFFE0FFBAFCCAF90020B7F932" box="[819,906,1705,1731]" captionStart="Table 8" captionStartId="24.[189,258,268,294]" captionText="Table 8. Meristic and morphometric data of Aphyosemion rubrogaster Chirio, Legros &amp; Agnèse sp. nov. See Table 3 for the abbreviations used." pageId="20" pageNumber="21">Table 8</tableCitation>
<tableCitation id="C6A7036BFFE0FFBAFCCAF90020B7F932" box="[819,906,1705,1731]" captionStart="Table 8" captionStartId="24.[189,258,268,294]" captionTargetPageId="24" captionText="Table 8. Meristic and morphometric data of Aphyosemion rubrogaster Chirio, Legros &amp; Agnèse sp. nov. See Table 3 for the abbreviations used." pageId="20" pageNumber="21">Table 8</tableCitation>
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@ -143,7 +143,7 @@ a.s.l.
leg. (
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@ -181,7 +181,7 @@ Chirio, Legros &amp; Agnèse
. Paratype, adult, ♀, from the Niengé River (MRAC 2016-019-P-94-108).
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<paragraph id="8B9A36D0FFE2FFB8FF44FEA521D8FEB9" blockId="22.[189,1398,267,329]" pageId="22" pageNumber="23">
<emphasis id="B951EAC2FFE2FFB8FF44FEA52218FED7" bold="true" box="[189,293,268,294]" pageId="22" pageNumber="23">Table 7.</emphasis>
Meristic and morphometric data of

View file

@ -0,0 +1,219 @@
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<heading id="D0A5819F8E45D52ED097FBD7FD33FB8D" box="[231,676,1108,1135]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="53" reason="2">
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<emphasis id="B926EAE18E45D52ED097FBD7FE43FB8D" bold="true" box="[231,468,1108,1135]" italics="true" pageId="4" pageNumber="53">Plesiotortrix edwardsi</emphasis>
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(
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)
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<emphasis id="B926EAE18E45D52ED0F4FB2EFEDAFB27" box="[132,333,1197,1221]" italics="true" pageId="4" pageNumber="53">Plesiotortrix edwardsi</emphasis>
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. —
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<emphasis id="B926EAE18E45D52ED1F2FB5EFDDDFB17" box="[386,586,1245,1269]" italics="true" pageId="4" pageNumber="53">Plesiotortrix edwardsi</emphasis>
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(new genus and species); [
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<emphasis id="B926EAE18E45D52ED10CFB7BFDD4FAF2" box="[380,579,1272,1296]" italics="true" pageId="4" pageNumber="53">Plesiotortrix edwardsi</emphasis>
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<bibRefCitation id="EFC34B028E45D52ED220FB7BFC90FAF2" author="RAGE J. - C. &amp; AUGE M." box="[592,775,1272,1296]" pageId="4" pageNumber="53" pagination="199 - 216" refId="ref7953" refString="RAGE J. - C. &amp; AUGE M. 1993. - Squamates from the Cenozoic of the western part of Europe. A review. Revue de Paleobiologie, volume special 7: 199 - 216." type="journal article" year="1993">Rage &amp; Augé 1993</bibRefCitation>
(nomen dubium). Note that the “Taxonomic history” section here deals only with novel nomenclatural acts, new taxonomic combinations, or novel taxonomic renderings of the taxon, similarly to the style proposed in other recent reptilian papers (e.g.
<bibRefCitation id="EFC34B028E45D52ED2E5FAE1FC97FA9B" author="JOYCE W. G." box="[661,768,1377,1402]" pageId="4" pageNumber="53" pagination="21 - 56" refId="ref6925" refString="JOYCE W. G. 2016. - A review of the fossil record of turtles of the clade Pan-Chelydridae. Bulletin of the Peabody Museum of Natural History 57 (1): 21 - 56. https: // doi. org / 10.3374 / 014.057.0103" type="journal article" year="2016">Joyce 2016</bibRefCitation>
, 2017;
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;
<bibRefCitation id="EFC34B028E45D52ED1CAFAFFFD18FA76" author="GEORGALIS G. L. &amp; RABI M. &amp; SMITH K. T." box="[442,655,1404,1428]" pageId="4" pageNumber="53" refId="ref6328" refString="GEORGALIS G. L., RABI M. &amp; SMITH K. T. 2021 c. - Taxonomic revision of the snakes of the genera Palaeopython and Paleryx (Serpentes, Constrictores) from the Paleogene of Europe. Swiss Journal of Palaeontology 140 (18), 140 p. https: // doi. org / 10.1186 / s 13358 - 021 - 00224 - 0" type="book" year="2021">
Georgalis
<emphasis id="B926EAE18E45D52ED26CFAFEFDDCFA76" box="[540,587,1404,1428]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
2021c
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).
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. —
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), three articulated trunk vertebrae (
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1884
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; pl. II.6, II.6a; this paper:
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;
<figureCitation id="13692A768E45D52ED2F8FA62FD02FA1B" box="[648,661,1505,1529]" captionStart="FIG" captionStartId="6.[811,822,549,566]" captionTargetBox="[853,1337,230,473]" captionTargetId="figure-1044@6.[812,1180,213,436]" captionTargetPageId="6" captionText="FIG. 2. — A, B, Original lithograph of the holotype articulated trunk vertebrae (MNHN.F.QU16332) of Plesiotortrix edwardsi Rochebrune, 1884 in anterior (A) and dorsal (B) views. Modified from Rochebrune (1884: pl. II). Note the inaccuracies of both images, compared with the actual specimen, as shown in the photographs in Figure 1 above." pageId="4" pageNumber="53">2</figureCitation>
)
</materialsCitation>
.
</paragraph>
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<smallCapsWord id="8D0BA02F8E45D52ED169F995FEA9F9C5" baselines="1571" box="[281,318,1558,1575]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="age" pageId="4" pageNumber="53">AGE</smallCapsWord>
. — Similar to most other fossil specimens collected during the 19th century, there are no precise locality data for the
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of
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<emphasis id="B926EAE18E45D52ED164F9C6FE42F9BF" box="[276,469,1605,1629]" italics="true" pageId="4" pageNumber="53">Plesiotortrix edwardsi</emphasis>
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, apart from the general information that it originates from the Phosphorites du Quercy. However, this is still vague, taking into consideration that the Phosphorites du Quercy include at least 170 fissure filling localities, distributed over a broad geographic area, encompassing large parts of the current departments of Lot, Tarn-et-Garonne, Tarn, and Aveyron, all in the administrative region of Occitanie (
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; (
<bibRefCitation id="EFC34B028E45D52ED22AF967FF21F8F4" author="SIGE B. &amp; HUGUENEY M." pageId="4" pageNumber="53" pagination="207 - 227" refId="ref8256" refString="SIGE B. &amp; HUGUENEY M. 2006. - Les micromammiferes des gisements a Phosphate du Quercy (SW France). Strata 13: 207 - 227." type="journal article" year="2006">Sigé &amp; Hugueney 2006</bibRefCitation>
;
<bibRefCitation id="EFC34B028E45D52ED0B6F97DFE37F8F4" author="GEORGALIS G. L. &amp; CERNANSKY A. &amp; KLEMBARA J." box="[198,416,1790,1814]" pageId="4" pageNumber="53" pagination="219 - 293" refId="ref6176" refString="GEORGALIS G. L., CERNANSKY A. &amp; KLEMBARA J. 2021 a. - Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material. Geodiversitas 43 (9): 219 - 293. https: // doi. org / 10.5252 / geodiversitas 2021 v 43 a 9" type="journal article" year="2021">
Georgalis
<emphasis id="B926EAE18E45D52ED15AF97CFECCF8F4" box="[298,347,1790,1814]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
2021a
</bibRefCitation>
, c, 2023;
<bibRefCitation id="EFC34B028E45D52ED278F97DFD56F8F4" author="PELISSIE T. &amp; ORLIAC M. J. &amp; ANTOINE P. - O. &amp; BLOT V. &amp; ESCARGUEL G." box="[520,705,1790,1814]" pageId="4" pageNumber="53" pagination="573 - 585" refId="ref7543" refString="PELISSIE T., ORLIAC M. J., ANTOINE P. - O., BLOT V. &amp; ESCARGUEL G. 2021. - Beyond Eocene and Oligocene epochs: The Causses Du Quercy Geopark and the Grande Coupure. Geoconservation Research 4: 573 - 585." type="journal article" year="2021">
Pelissié
<emphasis id="B926EAE18E45D52ED225F97CFD11F8F4" box="[597,646,1790,1814]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
2021
</bibRefCitation>
). They also stratigraphically span over a considerable time period, from the early Eocene (MP 8+9) until the Early Miocene (MN 3); however, most of the respective fossiliferous localities range between the late middle Eocene (MP 16) and the late Oligocene (MP 28) (
<bibRefCitation id="EFC34B028E45D52ED2A6F8EBFF20F878" author="RAGE J. - C." pageId="4" pageNumber="53" pagination="161 - 173" refId="ref7847" refString="RAGE J. - C. 2006. - The lower vertebrates from the Eocene and Oligocene of the Phosphorites du Quercy (France): an overview, in PELISSIE T. &amp; SIGE B. (eds), 30 millions d'annees de biodiversite dynamique dans le paleokarst du Quercy, Journees Bernard Geze. Strata 13: 161 - 173." type="journal article" year="2006">Rage 2006</bibRefCitation>
;
<bibRefCitation id="EFC34B028E45D52ED0B6F801FE24F878" author="SIGE B. &amp; HUGUENEY M." box="[198,435,1922,1947]" pageId="4" pageNumber="53" pagination="207 - 227" refId="ref8256" refString="SIGE B. &amp; HUGUENEY M. 2006. - Les micromammiferes des gisements a Phosphate du Quercy (SW France). Strata 13: 207 - 227." type="journal article" year="2006">Sigé &amp; Hugueney 2006</bibRefCitation>
;
<bibRefCitation id="EFC34B028E45D52ED1B2F801FD0CF879" author="GEORGALIS G. L. &amp; CERNANSKY A. &amp; KLEMBARA J." box="[450,667,1922,1947]" pageId="4" pageNumber="53" pagination="219 - 293" refId="ref6176" refString="GEORGALIS G. L., CERNANSKY A. &amp; KLEMBARA J. 2021 a. - Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material. Geodiversitas 43 (9): 219 - 293. https: // doi. org / 10.5252 / geodiversitas 2021 v 43 a 9" type="journal article" year="2021">
Georgalis
<emphasis id="B926EAE18E45D52ED256F800FDC0F878" box="[550,599,1922,1946]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
2021a
</bibRefCitation>
, c;
<bibRefCitation id="EFC34B028E45D52ED2B2F801FF7AF857" author="PELISSIE T. &amp; ORLIAC M. J. &amp; ANTOINE P. - O. &amp; BLOT V. &amp; ESCARGUEL G." pageId="4" pageNumber="53" pagination="573 - 585" refId="ref7543" refString="PELISSIE T., ORLIAC M. J., ANTOINE P. - O., BLOT V. &amp; ESCARGUEL G. 2021. - Beyond Eocene and Oligocene epochs: The Causses Du Quercy Geopark and the Grande Coupure. Geoconservation Research 4: 573 - 585." type="journal article" year="2021">
Pelissié
<emphasis id="B926EAE18E45D52ED0F4F81DFF23F857" box="[132,180,1949,1973]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
2021
</bibRefCitation>
), though still important finds have also been recovered from the older (early Eocene) and younger (Early Miocene) sites (e.g. Čerňanský 2023;
<bibRefCitation id="EFC34B028E45D52ED113F850FDD4F808" author="CERNANSKY A. &amp; DAZA J. D. &amp; TABUCE R. &amp; SAXTON E. &amp; VIDALENC D." box="[355,579,2002,2029]" pageId="4" pageNumber="53" pagination="695 - 708" refId="ref5597" refString="CERNANSKY A., DAZA J. D., TABUCE R., SAXTON E. &amp; VIDALENC D. 2023 a. - An early Eocene pan-gekkotan from France could represent an extra squamate group that survived the K / Pg extinction. Acta Palaeontologica Polonica 68 (4): 695 - 708. https: // doi. org / 10.4202 / app. 01083.2023" type="journal article" year="2023">
Čerňanský
<emphasis id="B926EAE18E45D52ED1A2F850FD96F808" box="[466,513,2002,2026]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
2023a
</bibRefCitation>
, b).
</paragraph>
</subSubSection>
<subSubSection id="C34865788E45D52ED35DFF5BFA27F809" pageId="4" pageNumber="53" type="description">
<paragraph id="8BED36F38E45D52ED35DFF5BFC2AFF12" blockId="4.[811,1458,216,2028]" box="[813,957,216,242]" pageId="4" pageNumber="53">
<smallCapsWord id="8D0BA02F8E45D52ED35DFF5BFC2AFF12" baselines="235,235" box="[813,957,216,242]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="4" pageNumber="53">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BED36F38E45D52ED35DFF7BFBD6F8CE" blockId="4.[811,1458,216,2028]" pageId="4" pageNumber="53">
The
<typeStatus id="54E988518E45D52ED311FF7BFC52FEF0" box="[865,965,248,274]" pageId="4" pageNumber="53" type="holotype">holotype</typeStatus>
and only known specimen of
<taxonomicName id="4C524D708E45D52ED55DFF7BFC1DFED3" authorityName="Rochebrune" authorityYear="1884" class="Squamata" family="Boidae" genus="Plesiotortrix" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="edwardsi">
<emphasis id="B926EAE18E45D52ED55DFF7BFC1DFED3" italics="true" pageId="4" pageNumber="53">Plesiotortrix edwardsi</emphasis>
</taxonomicName>
consists of three articulated trunk vertebrae. The three vertebrae are relatively well preserved, but still have parts of their prezygapophyses and neural spines damaged and their paradiapophyses are eroded (
<figureCitation id="13692A768E45D52ED4BAFEF4FA90FE73" box="[1226,1287,375,401]" captionStart="FIG" captionStartId="5.[132,143,1985,2002]" captionTargetBox="[141,1446,215,1852]" captionTargetId="figure-81@5.[703,1262,759,1307]" captionTargetPageId="5" captionText="FIG. 1. — A-F, Holotype articulated trunk vertebrae (MNHN.F.QU16332) of Plesiotortrix edwardsi Rochebrune,1884 in anterior (A), posterior (B), ventral (C), dorsal (D), right lateral (E), and left lateral (F) views. Scale bar: 5 mm." pageId="4" pageNumber="53">Fig. 1</figureCitation>
). The vertebrae are moderately small, with an average centrum length (CL) for each vertebra of approximately
<quantity id="4CAA9B168E45D52ED4EEFE34FB65FE33" box="[1182,1266,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="53" unit="mm" value="3.3">3.3 mm</quantity>
and a neural arch width (NAW) of 3.9 (ratio CL / NAW approximately 0.8). In anterior view (
<figureCitation id="13692A768E45D52ED382FE74FBDEFDF3" box="[1010,1097,503,529]" captionStart="FIG" captionStartId="5.[132,143,1985,2002]" captionTargetBox="[141,1446,215,1852]" captionTargetId="figure-81@5.[703,1262,759,1307]" captionTargetPageId="5" captionText="FIG. 1. — A-F, Holotype articulated trunk vertebrae (MNHN.F.QU16332) of Plesiotortrix edwardsi Rochebrune,1884 in anterior (A), posterior (B), ventral (C), dorsal (D), right lateral (E), and left lateral (F) views. Scale bar: 5 mm." pageId="4" pageNumber="53">Fig. 1A</figureCitation>
), the zygosphene is straight and relatively thin. The neural canal is trapezoidal in shape. The prezygapophyses are only slightly inclined, but they clearly reach above the level of the floor of the neural canal. The cotyle is large and circular. Deep fossae are present on each lateral side of the cotyle; no paracotylar foramina are present. In posterior view (
<figureCitation id="13692A768E45D52ED38DFD35FBC5FD33" box="[1021,1106,694,721]" captionStart="FIG" captionStartId="5.[132,143,1985,2002]" captionTargetBox="[141,1446,215,1852]" captionTargetId="figure-81@5.[703,1262,759,1307]" captionTargetPageId="5" captionText="FIG. 1. — A-F, Holotype articulated trunk vertebrae (MNHN.F.QU16332) of Plesiotortrix edwardsi Rochebrune,1884 in anterior (A), posterior (B), ventral (C), dorsal (D), right lateral (E), and left lateral (F) views. Scale bar: 5 mm." pageId="4" pageNumber="53">Fig. 1B</figureCitation>
), the neural arch is moderately depressed, with a vaulting ratio (
<emphasis id="B926EAE18E45D52ED4F6FD54FB2CFD12" box="[1158,1211,727,752]" italics="true" pageId="4" pageNumber="53">sensu</emphasis>
<bibRefCitation id="EFC34B028E45D52ED4B0FD55FA30FD12" author="GEORGALIS G. L. &amp; RABI M. &amp; SMITH K. T." box="[1216,1447,726,752]" pageId="4" pageNumber="53" refId="ref6328" refString="GEORGALIS G. L., RABI M. &amp; SMITH K. T. 2021 c. - Taxonomic revision of the snakes of the genera Palaeopython and Paleryx (Serpentes, Constrictores) from the Paleogene of Europe. Swiss Journal of Palaeontology 140 (18), 140 p. https: // doi. org / 10.1186 / s 13358 - 021 - 00224 - 0" type="book" year="2021">
Georgalis
<emphasis id="B926EAE18E45D52ED55CFD54FAC9FD12" box="[1324,1374,726,752]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
2021c
</bibRefCitation>
) equal to 0.38. The parapophyseal portion of the paradiapophysis (fully preserved only in the right paradiapophysis of the third vertebra) extends ventrally below the level of the cotyle and condyle. The condyle is large and slightly elliptical. In dorsal view (
<figureCitation id="13692A768E45D52ED473FCF6FBCDFC72" box="[1027,1114,885,912]" captionStart="FIG" captionStartId="5.[132,143,1985,2002]" captionTargetBox="[141,1446,215,1852]" captionTargetId="figure-81@5.[703,1262,759,1307]" captionTargetPageId="5" captionText="FIG. 1. — A-F, Holotype articulated trunk vertebrae (MNHN.F.QU16332) of Plesiotortrix edwardsi Rochebrune,1884 in anterior (A), posterior (B), ventral (C), dorsal (D), right lateral (E), and left lateral (F) views. Scale bar: 5 mm." pageId="4" pageNumber="53">Fig. 1D</figureCitation>
), the zygosphene possesses two incipient lateral lobes and a wide median lobe. The neural spine commences slightly posteriorly from the level of the zygosphene; its base becomes wider and distinctly saddle-shaped towards the posterior half of the neural arch. The prezygapophyseal articular facets are massive and broad. The interzygapophyseal constriction is deep. In ventral view (
<figureCitation id="13692A768E45D52ED346FBD6FC1BFB8D" box="[822,908,1109,1135]" captionStart="FIG" captionStartId="5.[132,143,1985,2002]" captionTargetBox="[141,1446,215,1852]" captionTargetId="figure-81@5.[703,1262,759,1307]" captionTargetPageId="5" captionText="FIG. 1. — A-F, Holotype articulated trunk vertebrae (MNHN.F.QU16332) of Plesiotortrix edwardsi Rochebrune,1884 in anterior (A), posterior (B), ventral (C), dorsal (D), right lateral (E), and left lateral (F) views. Scale bar: 5 mm." pageId="4" pageNumber="53">Fig. 1C</figureCitation>
), the centrum is wider than long. A distinct haemal keel runs almost throughout the midline of the ventral surface of the centrum. It is relatively thin and has the same width across its whole length; it commences anteriorly at around the ventral lip of the cotyle and terminates posteriorly right prior to the condylar lip. Prezygapophyseal accessory processes are present, with their tip only slightly projecting beyond the prezygapophyseal articular facets. The subcentral grooves are deep. Subcentral foramina are visible, situated next to the haemal keel. The paradiapophyses are eroded in the three vertebrae with the exception of the right paradiapophyses of the last vertebra, which is almost complete. There seems to be no clear differentiation into diapophyses and parapophyses; in the almost completely preserved right paradiapophysis of the third vertebra, it seems that the parapophyseal portion is slightly larger than the diapophyseal one. The postzygapophyseal articular facets are large and their posterior edge is somehow acute. In lateral view (
<figureCitation id="13692A768E45D52ED546F9F0FA35F96F" box="[1334,1442,1651,1677]" captionStart="FIG" captionStartId="5.[132,143,1985,2002]" captionTargetBox="[141,1446,215,1852]" captionTargetId="figure-81@5.[703,1262,759,1307]" captionTargetPageId="5" captionText="FIG. 1. — A-F, Holotype articulated trunk vertebrae (MNHN.F.QU16332) of Plesiotortrix edwardsi Rochebrune,1884 in anterior (A), posterior (B), ventral (C), dorsal (D), right lateral (E), and left lateral (F) views. Scale bar: 5 mm." pageId="4" pageNumber="53">Fig. 1E, F</figureCitation>
), the neural spine is damaged and practically only its base is preserved. A small condylar neck is present. The subcentral ridges are slightly convex. The haemal keel slightly projects ventrally, with its dorsoventral height being higher towards its posteriormost section.
</paragraph>
<paragraph id="8BED36F38E45D52ED333F8B1FA27F809" blockId="4.[811,1458,216,2028]" pageId="4" pageNumber="53">Judging from the absolute shape, the vaulting of the neural arch, the depth of the subcentral grooves, and the width of the haemal keel, I assume that these three vertebrae originate from the mid- or anterior posterior trunk region of the column. This slightly contradictsRochebrunes (1884) original description, who treated them as “vertèbres de la région pelvienne”.</paragraph>
</subSubSection>
</treatment>
</document>