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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.392.6328" ID-PMC="PMC3974435" ID-Pensoft-Pub="1313-2970-392-1" ID-Pensoft-UUID="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" ID-PubMed="24715782" ID-Zenodo-Dep="578044" ModsDocID="1313-2970-392-1" checkinTime="1451246138192" checkinUser="pensoft" docAuthor="Breure, Abraham S. H. &amp; Ablett, Jonathan D." docDate="2014" docId="6DBB3192730707FE636B5D2EA75163A6" docLanguage="en" docName="ZooKeys 392: 1-367" docOrigin="ZooKeys 392" docPubDate="2014-03-21" docSource="http://dx.doi.org/10.3897/zookeys.392.6328" docTitle="Bulimus clathratus Pfeiffer 1858" docType="treatment" docVersion="5" id="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" lastPageNumber="37" masterDocId="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" masterDocTitle="Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London" masterLastPageNumber="367" masterPageNumber="1" pageNumber="37" updateTime="1668158078298" updateUser="ExternalLinkService">
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<mods:title>Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Breure, Abraham S. H.</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">bbreure@xs4all.nl</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Natural History Museum, Division of Higher Invertebrates, London, SW 7 5 BD, UK</mods:affiliation>
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<taxonomicName LSID="6DBB3192-7307-07FE-636B-5D2EA75163A6" authority="Pfeiffer, 1858" authorityName="Pfeiffer" authorityYear="1858" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Bulimus clathratus" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="clathratus">Bulimus clathratus Pfeiffer, 1858</taxonomicName>
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Drymaeus species. A-C Drymaeus (Drymaeus) cuzcoensis (Reeve, 1849), lectotype NHMUK 1975453 (H = 36.6) D-F Drymaeus (Drymaeus) clathratus (Pfeiffer, 1858), lectotype NHMUK 1975449 (H = 30.0) G-I Drymaeus (Drymaeus) praetextus (Reeve, 1849), lectotype NHMUK 198340 (H = 39.0) J-L Drymaeus (Drymaeus) alabastrinus da Costa, 1906, holotype NHMUK 1907.11.21.16 (H = 33.0). All enlarged." httpUri="https://binary.pensoft.net/fig/26711" pageId="36" pageNumber="37">Figs 43D-F</figureCitation>
<figureCitation captionStart="Figure L13" captionStartId="F13L" captionText="Figure L 13. Labels. i Bulimus citronellus Angas, 1879 ii Bulimus clarus Pfeiffer, 1857 iii Bulimus clathratus Pfeiffer, 1858 iv Bulimus coagulatus Reeve, 1849 v Bulimus coarctatus Pfeiffer, 1845 vi Bulimulus compactus Fulton, 1902." httpUri="https://binary.pensoft.net/fig/26764" pageId="36" pageNumber="37">, L13iii</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="reference_group">
<paragraph pageId="36" pageNumber="37">
<taxonomicName authorityName="Pfeiffer" authorityYear="1858" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="clathratus">Bulimus clathratus</taxonomicName>
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Malakozoologische Blaetter" pageId="187" pageNumber="188" pagination="256 - 258" refId="B130" refString="Pfeiffer, L, 1858a. Descriptions of ten new species of Bulimus, from the collection of H. Cuming, Esq. In: Proceedings of the Zoological Society of London.: 256 - 258" title="Descriptions of ten new species of Bulimus, from the collection of H. Cuming, Esq." volumeTitle="Proceedings of the Zoological Society of London." year="1858 a">Pfeiffer 1858a</bibRefCitation>
: 258;
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Malakozoologische Blaetter" pageId="187" pageNumber="188" pagination="19 - 53" refId="B132" refString="Pfeiffer, L, 1859a. Nachtraege zum zweiten Supplemente meiner Monographia Heliceorum (Fortsetzung). Malakozoologische Blaetter 6: 19 - 53" title="Nachtraege zum zweiten Supplemente meiner Monographia Heliceorum (Fortsetzung)." volume="6" year="1859 a">Pfeiffer 1859a</bibRefCitation>
: 47.
</paragraph>
<paragraph pageId="36" pageNumber="37">
<taxonomicName baseAuthorityName="L.Pfeiffer" baseAuthorityYear="1858" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="clathratus" subGenus="Drymaeus">Drymaeus (Drymaeus) clathratus</taxonomicName>
;
<bibRefCitation author="Breure, ASH" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="181" pageNumber="182" pagination="1 - 215" refId="B15" refString="Breure, ASH, 1979. Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen Leiden 168: 1 - 215" title="Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca)." volume="168" year="1979">Breure 1979</bibRefCitation>
: 107 (lectotype designation);
<bibRefCitation author="Breure, ASH" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="181" pageNumber="182" pagination="1 - 111" refId="B25" refString="Breure, ASH, Eskens, AAC, 1981. Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda), II. Zoologische Verhandelingen Leiden 186: 1 - 111" title="Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda), II." volume="186" year="1981">Breure and Eskens 1981</bibRefCitation>
: 14.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="type locality">
<paragraph pageId="36" pageNumber="37">Type locality.</paragraph>
<paragraph pageId="36" pageNumber="37">
&quot;Province of Patas, Andes of Peru (
<emphasis italics="true" pageId="36" pageNumber="37">Dr. Farris</emphasis>
)&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="label">
<paragraph pageId="36" pageNumber="37">Label.</paragraph>
<paragraph pageId="36" pageNumber="37">
&quot;Province of Patas, / Andes of Peru / Dr Farris&quot;; taxon label in
<normalizedToken originalValue="Pfeiffers">Pfeiffer's</normalizedToken>
handwriting.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="dimensions">
<paragraph pageId="36" pageNumber="37">Dimensions.</paragraph>
<paragraph pageId="36" pageNumber="37">&quot;Long. 30, diam. 11 mill.&quot;; figured specimen herein H 30, D 12.2, W 7.8.</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="type material">
<paragraph pageId="36" pageNumber="37">Type material.</paragraph>
<paragraph pageId="36" pageNumber="37">NHMUK 1975449, lectotype (Cuming coll.).</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="remarks">
<paragraph pageId="36" pageNumber="37">Remarks.</paragraph>
<paragraph pageId="36" pageNumber="37">
This hitherto unfigured taxon is known from a single specimen. The current systematic position follows
<bibRefCitation author="Richardson, CL" journalOrPublisher="Tryonia" pageId="189" pageNumber="190" pagination="1 - 458" refId="B164" refString="Richardson, CL, 1995. Bulimulidae: catalog of species. Tryonia 28: 1 - 458" title="Bulimulidae: catalog of species." volume="28" year="1995">Richardson (1995</bibRefCitation>
: 110).
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="current systematic position">
<paragraph pageId="36" pageNumber="37">Current systematic position.</paragraph>
<paragraph pageId="36" pageNumber="37">
<taxonomicName authorityName="Tryon" authorityYear="1867" class="Gastropoda" family="Bulimulidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="36" pageNumber="37" phylum="Mollusca" rank="family">Bulimulidae</taxonomicName>
,
<taxonomicName baseAuthorityName="L.Pfeiffer" baseAuthorityYear="1858" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="clathratus" subGenus="Drymaeus">
<emphasis italics="true" pageId="36" pageNumber="37">Drymaeus (Drymaeus) clathratus</emphasis>
</taxonomicName>
(Pfeiffer, 1858).
</paragraph>
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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8151" ID-Pensoft-Pub="1314-2828-4-8151" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e8151" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" checkinTime="1461288934892" checkinUser="pensoft" docAuthor="Broad, Gavin R., Shaw, Mark R. &amp; Godfray, H. Charles J." docDate="2016" docId="6DBB38B8F2E419787757349A82B8BE15" docLanguage="en" docName="BiodivDatJour 4: e8151" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8151" docTitle="Napamus Papp 1993" docType="treatment" docVersion="3" lastPageNumber="8151" masterDocId="EC41FFB1FFD9782CFFAEFFF87E2EFF92" masterDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" masterLastPageNumber="8151" masterPageNumber="8151" pageNumber="8151" updateTime="1618528331117" updateUser="pensoft">
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<taxonomicName authority="Papp, 1993" authorityName="Papp" authorityYear="1993" class="Insecta" family="Braconidae" genus="Napamus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Napamus" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Napamus Papp, 1993</taxonomicName>
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<subSubSection pageId="0" pageNumber="8151" type="notes">
<paragraph pageId="0" pageNumber="8151">Notes</paragraph>
<paragraph pageId="0" pageNumber="8151">
species of
<taxonomicName class="Insecta" family="Braconidae" genus="Napamus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Napamus" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Napamus</taxonomicName>
excluded from the British and Irish list:
</paragraph>
<paragraph pageId="0" pageNumber="8151">
[vipio (Reinhard, 1880,
<taxonomicName class="Insecta" family="Braconidae" genus="Apanteles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apanteles" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Apanteles</taxonomicName>
)] Mistakenly listed as a British species by
<bibRefCitation author="Kloet, G. S." journalOrPublisher="Stockport by the Authors, Stockport" pageId="0" pageNumber="8151" title="A check list of British insects" year="1945">Kloet and Hincks (1945)</bibRefCitation>
and
<bibRefCitation author="Huddleston, T." editor="Fitton, M. G." journalOrPublisher="Handbooks for the Identification of British Insects 11: ix + 159 pp." pageId="0" pageNumber="8151" title="Kloet and Hincks. A check list of British insects. Part 4: Hymenoptera" year="1978">Huddleston (1978)</bibRefCitation>
.
<bibRefCitation author="Shenefelt, R. D." journalOrPublisher="Hymenopterorum Catalogus (nova editio)" pageId="0" pageNumber="8151" pagination="669 - 812" title="Braconidae 5. Microgasterinae &amp; Ichneutinae" volume="11" year="1973">Shenefelt (1973)</bibRefCitation>
listed it as British on the misunderstanding that
<bibRefCitation author="Morley, C." journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="8151" pagination="133 - 183" title="The Hymenopterous parasites of the British Lepidoptera" volume="81" year="1933">Morley and Rait-Smith (1933)</bibRefCitation>
had produced a catalogue of British
<taxonomicName lsidName="Ichneumonoidea" order="Lepidoptera" pageId="0" pageNumber="8151" rank="order">Lepidoptera</taxonomicName>
-parasitoid asscociations, whereas their listings also included non-British rearings from hosts that occur in Britain.
<bibRefCitation author="Morley, C." journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="8151" pagination="133 - 183" title="The Hymenopterous parasites of the British Lepidoptera" volume="81" year="1933">Morley and Rait-Smith (1933)</bibRefCitation>
cite
<bibRefCitation author="Marshall, T. A." editor="Andre, E." journalOrPublisher="Gray 1891" pageId="0" pageNumber="8151" title="Species des Hymenopteres d'Europe et d'Algerie" year="1896">Marshall (1896)</bibRefCitation>
as the source of host records for vipio, who does not mention Britain.
<taxonomicName class="Insecta" family="Braconidae" genus="Apanteles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apanteles vipio" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species">Apanteles vipio</taxonomicName>
was transferred from
<taxonomicName class="Insecta" family="Braconidae" genus="Illidops" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Illidops" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Illidops</taxonomicName>
by
<bibRefCitation author="Papp, J." journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="8151" pagination="155 - 180" title="New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 4" volume="84" year="1993">Papp (1993)</bibRefCitation>
.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:CC84B06D-AD54-41A7-8237-F2031D57F0A7" authority="Rix &amp; Harvey" class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea thompsoni" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="thompsoni">Austrarchaea thompsoni Rix &amp; Harvey</taxonomicName>
<taxonomicNameLabel pageId="9" pageNumber="10">sp. n.</taxonomicNameLabel>
Figs 101925
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="vernacular name">
<paragraph pageId="9" pageNumber="10">Vernacular name.</paragraph>
<paragraph pageId="9" pageNumber="10">Carbine Tableland Assassin Spider</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="type material">
<paragraph pageId="9" pageNumber="10">Type material.</paragraph>
<paragraph pageId="9" pageNumber="10">
Holotype male: Devils Thumb area, [Daintree National Park (Mossman Gorge Section)], 10 km NW. of Mossman, Queensland, Australia, [
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-16.45">16°27'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="145.28334">145°17'E</geoCoordinate>
], pyrethrum knockdown, tropical rainforest, 1000-1180 m, 10.X.1982, G. Monteith, D. Yeates, G. Thompson (QMB S30840).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="etymology">
<paragraph pageId="9" pageNumber="10">Etymology</paragraph>
<paragraph pageId="9" pageNumber="10">. The specific epithet is a patronym in honour of Geoff Thompson, for his ongoing efforts in collecting and documenting the invertebrate rainforest fauna of the Wet Tropics, and for collecting the only known specimen of this species.</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="diagnosis">
<paragraph pageId="9" pageNumber="10">Diagnosis.</paragraph>
<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea thompsoni" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="thompsoni">Austrarchaea thompsoni</taxonomicName>
can be distinguished from all other
<taxonomicName family="Archaeidae" lsidName="" pageId="9" pageNumber="10" rank="family">Archaeidae</taxonomicName>
from north-eastern Queensland except
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea karenae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="karenae">Austrarchaea karenae</taxonomicName>
,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea tealei" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="tealei">Austrarchaea tealei</taxonomicName>
sp. n. and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea wallacei" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="wallacei">Austrarchaea wallacei</taxonomicName>
by the
<pageBreakToken pageId="10" pageNumber="11" start="start">presence</pageBreakToken>
of a triangular spur on the embolus (Fig. 10D); and from
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea karenae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="karenae">Austrarchaea karenae</taxonomicName>
,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea tealei" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tealei">Austrarchaea tealei</taxonomicName>
sp. n. and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea wallacei" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="wallacei">Austrarchaea wallacei</taxonomicName>
by the very small tegular sclerite 1 (TS 1), which is not visible in ventral view (Fig. 10D), and by the more proximally positioned embolic spur, which is situated near the base of the exposed embolic portion (Fig. 10D).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="description">
<paragraph pageId="10" pageNumber="11">Description.</paragraph>
<paragraph pageId="10" pageNumber="11">
Holotype male: Total length 2.97; leg I femur 3.23; F1/CL ratio 2.74. Cephalothorax dark reddish-brown; legs tan-brown with darker annulations; abdomen mottled grey-brown and beige, with darker brown dorsal scute and sclerites (Fig. 10A). Carapace tall (CH/CL ratio 2.13); 1.18 long, 2.51 high, 1.13 wide,
<normalizedToken originalValue="neck">'neck'</normalizedToken>
0.63 wide; bearing two pairs of rudimentary horns; highest point of pars cephalica (HPC) near posterior third of
<normalizedToken originalValue="head">'head'</normalizedToken>
(ratio of HPC to post-ocular length 0.69), carapace gently sloping posterior to HPC;
<normalizedToken originalValue="head">'head'</normalizedToken>
not strongly elevated dorsally (post-ocular ratio 0.25). Chelicerae with short brush of accessory setae on anterior face of paturon (Fig. 10B). Abdomen 1.64 long, 1.23 wide; with two pairs of dorsal hump-like tubercles (HT 1-4); dorsal scute fused anteriorly to epigastric sclerites, extending posteriorly to first pair of hump-like tubercles; HT 3-4 each covered by separate dorsal sclerites. Unexpanded pedipalp (Figs 10
<normalizedToken originalValue="CE">C-E</normalizedToken>
) of Type A morphology (Fig. 6), with large, retrolaterally directed, arched conductor; embolus distally directed, slightly sinuous, with short triangular spur situated near base of exposed embolic portion, embolus projecting beyond distal rim of conductor by slightly less than 1/2 length of exposed embolic portion; tegular sclerite 3 (TS 3) short, spur-like, with constricted tegular base and sharply pointed, claw-like apex; TS 2-2a looped beneath overhanging retrolateral edge of conductor, TS 2 with rounded, subtriangular apex, TS 2a projecting beyond distal rim of conductor to near tip of embolus; TS 1 very small, obscured by TS 2-3, not visible in ventral view.
</paragraph>
<paragraph pageId="10" pageNumber="11">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="distribution">
<paragraph pageId="10" pageNumber="11">Distribution and habitat.</paragraph>
<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea thompsoni" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="thompsoni">Austrarchaea thompsoni</taxonomicName>
is known only from Devils Thumb, on the Carbine Tableland 10 km west-north-west of Mossman (Figs 19, 25). The single known specimen was collected in high altitude tropical rainforest.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="conservation status">
<paragraph pageId="10" pageNumber="11">Conservation status.</paragraph>
<paragraph pageId="10" pageNumber="11">Unknown (data deficient).</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>De Mattia, Willy</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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<mods:namePart>Reier, Susanne</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haring, Elisabeth</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="6DBBAB53-D1A5-52FA-B9D1-E9432D6C9680" authority="(Rossmaessler, 1836) s. l." authorityName="s. l." baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">
Siciliaria grohmanniana (
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
, 1836) s. l.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="s. l." baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana</emphasis>
</taxonomicName>
forms a subclade with its sister group
<taxonomicName baseAuthorityName="Philippi" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="septemplicata">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria septemplicata</emphasis>
</taxonomicName>
(mean p distance 9.0%). The two subspecies of
<taxonomicName lsidName="S. grohmanniana" pageId="0" pageNumber="1" rank="species" species="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">S. grohmanniana</emphasis>
</taxonomicName>
appear monophyletic in the mt tree (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. BI tree of based on the COI data set of Siciliaria, Sicania and related genera. BI posterior probability values (&gt; 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure4" httpUri="https://binary.pensoft.net/fig/622796" pageId="0" pageNumber="1">4</figureCitation>
), albeit each represented by only a small number of samples:
<taxonomicName authority="grohmanniana" authorityName="grohmanniana (Rossmaessler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana grohmanniana</emphasis>
</taxonomicName>
(one population with three samples),
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. addaurae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="addaurae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana addaurae</emphasis>
</taxonomicName>
ssp. nov. (one population with three samples). Mean distance between the two subspecies was found to be 3.3%.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The shell of
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. addaurae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="addaurae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana addaurae</emphasis>
</taxonomicName>
ssp. nov. was at first considered by
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck (2013b</bibRefCitation>
: 6), as an &quot;intermediate geographic form&quot; of
<taxonomicName lsidName="S. grohmanninana" pageId="0" pageNumber="1" rank="species" species="grohmanninana">
<emphasis italics="true" pageId="0" pageNumber="1">S. grohmanninana</emphasis>
</taxonomicName>
s. s. with &quot;shell not decollate and whorls more weakly ribbed&quot; and (2013b: 10) as: &quot;a
<taxonomicName lsidName="S. grohmanniana" pageId="0" pageNumber="1" rank="species" species="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">S. grohmanniana</emphasis>
</taxonomicName>
with characters of
<taxonomicName lsidName="S. septemplicata" pageId="0" pageNumber="1" rank="species" species="septemplicata">
<emphasis italics="true" pageId="0" pageNumber="1">S. septemplicata</emphasis>
</taxonomicName>
&quot; but without stating what these characters are. He introduced it as the: &quot;Priola form from Priola (= Punta Priola) and San Lorenzo near Palermo&quot; and depicted a shell (
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
: 12, fig. 8). The internal penial sculpturing of
<taxonomicName authority="grohmanniana" authorityName="grohmanniana (Rossmaessler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana grohmanniana</emphasis>
</taxonomicName>
is similar to
<taxonomicName baseAuthorityName="Philippi" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="septemplicata">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria septemplicata</emphasis>
</taxonomicName>
(internal penial sculpturing showing a more or less fringed longitudinal pleats) although the sculpturing of epiphallus and vagina shows greater variability.
</paragraph>
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