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<mods:title id="4870023F0C83EDA74E15C423F55F6CB2">New records on the rich loriciferan fauna of Trezen ar Skoden (Roscoff, France): Description of two new species of Nanaloricus and the new genus Scutiloricus</mods:title>
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<mods:namePart id="DD4143A574C905466BEA5AFCA1DC625C">Neves, Ricardo Cardoso</mods:namePart>
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<emphasis id="B94BDF0769005162FD9FF883C52CF8A8" box="[565,631,1804,1827]" italics="true" pageId="23" pageNumber="24">Genus</emphasis>
.
<taxonomicName id="4C3F789669005162FD29F884C5A7F8A8" ID-CoL="9X5PD" authorityName="Neves &amp; Kristensen &amp; Møbjerg" authorityYear="2021" box="[643,764,1803,1827]" family="Nanaloricidae" genus="Scutiloricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Nanaloricida" pageId="23" pageNumber="24" phylum="Loricifera" rank="genus" status="gen. nov.">
<emphasis id="B94BDF0769005162FD29F884C5A7F8A8" box="[643,764,1803,1827]" italics="true" pageId="23" pageNumber="24">Scutiloricus</emphasis>
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(Type species:
<taxonomicName id="4C3F789669005162FD7AF8DFC4D3F8E3" authorityName="Neves &amp; Kristensen &amp; Møbjerg" authorityYear="2021" box="[720,904,1872,1896]" family="Nanaloricidae" genus="Scutiloricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Nanaloricida" pageId="23" pageNumber="24" phylum="Loricifera" rank="species" species="hugoi" status="gen. et sp. nov.">
<emphasis id="B94BDF0769005162FD7AF8DFC4D3F8E3" box="[720,904,1872,1896]" italics="true" pageId="23" pageNumber="24">Scutiloricus hugoi</emphasis>
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<taxonomicNameLabel id="A278627C69005162FC24F8DEC376F8E2" box="[910,1069,1873,1897]" pageId="23" pageNumber="24" rank="species">gen. et sp. nov.</taxonomicNameLabel>
)
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<emphasis id="B94BDF07690F516DFD9FFF5FC5AAFF63" bold="true" box="[565,753,208,232]" pageId="24" pageNumber="25">Genus diagnosis.</emphasis>
<emphasis id="B94BDF07690F516DFCACFF40C410FF6C" box="[774,843,207,231]" italics="true" pageId="24" pageNumber="25">Adults</emphasis>
. (1) mouth cone with 8 oral ridges of different length and characterized by posterior sclerotized oral furca and absence of oral stylets; (2) introvert with 9 rows of scalids; (3) first row with eight clavoscalids that differ between males (multiform, broad and flat, all branched except for the midventral pair) and females (four-segmented, slender, unbranched); (4) second row with 9 four-segmented, leg-like spinoscalids; (5) third row with 7 two-segmented, feather-like scalids; (6) fourth row with 16 spinoscalids of
<specimenCount id="9D39C89C690F516DFA83FEF2C2CAFE1E" box="[1321,1425,381,405]" count="2" pageId="24" pageNumber="25" type="generic" typeStatus="types">two types</specimenCount>
: 8 three-segmented, leg-like scalids (
<typeStatus id="5484BDB7690F516DFC90FE2FC432FE33" box="[826,873,416,440]" pageId="24" pageNumber="25">type</typeStatus>
A) alternate with 8 three-segmented spinoscalids with a feather-like distal segment (
<typeStatus id="5484BDB7690F516DFC93FE4DC433FE51" box="[825,872,450,474]" pageId="24" pageNumber="25">type</typeStatus>
B); (7) fifth to seventh rows all similar, each row with 30 leg-like, three-segmented scalids; (8) eighth row with 30 long, unsegmented spinoscalids whith a small conical base; (9) ninth row with 30 oval, plate-like scalids with several minute teeth; (10) neck with 8 single trichoscalids alternating with 7 double trichoscalids, with both the single and each of the double trichoscalids protruding from a single trichoscalid plate; (11) square lorica composed of six cuticular plates with reinforced anterior margins, honeycomb sculpturing and bearing a total of 14 large anterior spikes; (12) anterior spikes with 34 transverse cuticular ridges, except for the lateral spikes, which are reinforced internally by thick cuticle with a triangle wave-shape; (13) 10 flosculi (4 dorsolateral pairs and 1 dorsal pair) present posteriorly on the lorica; (14) posterior region of the lorica characterized by a slightly invaginated anal field from which a small anal cone protrudes posteroventrally, flanked by a pair of small spurs; (15) posterior region of the lorica also characterized by two pairs of pores: a dorsolateral (gonopores?) pair and a ventral (gland outlets?) pair; (16) internally, females are characterized by a pair of seminal receptacles (filled with mature spermatozoa) located in the abdomen and postero-dorsally to the ovaries. Larval stages were not found.
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<emphasis id="B94BDF07690F516DFD9FFC45C5F8FC69" box="[565,675,970,994]" italics="true" pageId="24" pageNumber="25">Etymology</emphasis>
. The generic name is composed by the Latin words scuti (= shield) and lorica (corset), masculine gender.
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Genus:
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Kristensen, 1983
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(Type species:
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<emphasis id="B94BDF0769145176FD7AFC44C4F4FC68" box="[720,943,971,995]" italics="true" pageId="3" pageNumber="4">Nanaloricus mysticus</emphasis>
Kristensen, 1983
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<mods:title id="0E228DD24185901DC1F6638D4ACE9B83">New Late Cretaceous mammals from the Intertrappean beds of Rangapur, India and paleobiogeographic framework</mods:title>
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<paragraph id="8BBB36A8A163FFADA25B0CB0FE71A758" blockId="6.[102,733,1260,1358]" pageId="6" pageNumber="337">
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<taxonomicName id="4C044D2BA163FFADA25B0CB0FE71A758" authority="Prasad, Jaeger, Sahni, Gheerbrant, and Khajuria, 1994" authorityName="Prasad, Jaeger, Sahni, Gheerbrant, and Khajuria" authorityYear="1994" class="Mammalia" family="Adapisoriculidae" genus="Deccanolestes" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="337" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B970EABAA163FFADA25B0CB0FE95A77C" box="[102,287,1260,1289]" italics="true" pageId="6" pageNumber="337">Deccanolestes</emphasis>
cf.
<emphasis id="B970EABAA163FFADA3720CB0FE37A77C" box="[335,445,1260,1289]" italics="true" pageId="6" pageNumber="337">robustus</emphasis>
<bibRefCitation id="EF954B59A163FFADA3FB0CB0FE71A758" author="Prasad, G. V. R. &amp; Jaeger, J. J. &amp; Sahni, A. &amp; Gheerbrant, E. &amp; Khajuria, C. K." pageId="6" pageNumber="337" pagination="260 - 277" refId="ref16988" refString="Prasad, G. V. R., Jaeger, J. J., Sahni, A., Gheerbrant, E., and Khajuria, C. K. 1994. Eutherian mammals from the Upper Cretaceous (Maastrichtian) Intertrappean Beds of Naskal, Andhra Pradesh, India. Journal of Vertebrate Paleontology 14: 260 - 277." type="journal article" year="1994">Prasad, Jaeger, Sahni, Gheerbrant, and Khajuria, 1994</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBB36A8A163FFADA25B0D64FF73A73B" blockId="6.[102,733,1260,1358]" box="[102,249,1336,1358]" pageId="6" pageNumber="337">
<tableCitation id="C6860313A163FFADA25B0D64FF3AA73B" box="[102,176,1336,1358]" captionStart="Table 2" captionStartId="5.[102,157,1298,1320]" captionText="Table 2. Measurements (in mm) and ratios of Rangapur and Naskal lower molars. Abbreviations as follows:L, length; TRW, trigonid width; TAW, talonid width; PHL, posthypocristid length; PEL, postentocristid length; PDH, protoconid height; MDH, metaconid height; AML, premetaconid length; TAH, talonid height; TAL, talonid length. Naskal data from Prasad et al. (1994)." pageId="6" pageNumber="337">Table 2</tableCitation>
,
<figureCitation id="133F2A2DA163FFADA2870D64FF7EA73B" box="[186,244,1336,1358]" captionStart="Fig" captionStartId="6.[102,135,655,674]" captionTargetBox="[102,1485,227,627]" captionTargetId="figure-717@6.[102,1485,227,627]" captionTargetPageId="6" captionText="Fig. 4. Deccanolestes cf. robustus. Right m1 or m2 (ITV/R/Mm6) in occlusal (A, B), lingual (C), and labial (D) views. Scale bar 0.5 mm." pageId="6" pageNumber="337">Fig. 4</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E6523A163FFADA25B0D3DFC52A487" pageId="6" pageNumber="337" type="materials_examined">
<paragraph id="8BBB36A8A163FFADA25B0D3DFDB5A70C" blockId="6.[102,575,1377,1401]" box="[102,575,1377,1401]" pageId="6" pageNumber="337">
<emphasis id="B970EABAA163FFADA25B0D3DFF42A70C" box="[102,200,1377,1401]" italics="true" pageId="6" pageNumber="337">Material</emphasis>
.—ITV/
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/Mm6, right m1 or m2.
</paragraph>
<paragraph id="8BBB36A8A163FFADA25B0DD5FD3CA7B4" blockId="6.[102,774,1417,1474]" pageId="6" pageNumber="337">
<emphasis id="B970EABAA163FFADA25B0DD5FF61A7D4" box="[102,235,1417,1441]" italics="true" pageId="6" pageNumber="337">Distribution</emphasis>
.—Late Cretaceous (Maastrichtian) Intertrappean beds of Naskal and Rangapur,
<collectingRegion id="49C0F84AA163FFADA3850DF5FDE6A7B4" box="[440,620,1449,1473]" country="India" name="Andhra Pradesh" pageId="6" pageNumber="337">Andhra Pradesh</collectingRegion>
,
<collectingCountry id="F3137638A163FFADA04B0DF5FD38A7B4" box="[630,690,1449,1473]" name="India" pageId="6" pageNumber="337">India</collectingCountry>
.
</paragraph>
<paragraph id="8BBB36A8A163FFADA25B0D8EFB3BA13C" blockId="6.[102,775,1490,2026]" lastBlockId="6.[812,1485,721,1417]" pageId="6" pageNumber="337">
<emphasis id="B970EABAA163FFADA25B0D8EFF6DA79C" box="[102,231,1490,1513]" italics="true" pageId="6" pageNumber="337">Description</emphasis>
.—ITV/
<collectionCode id="ED15AE6DA163FFADA3060D8EFEC7A79F" box="[315,333,1490,1514]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="337">R</collectionCode>
/Mm6 is identified as a right lower molar, either m1 or m2. The roots are missing and a small portion of the lingual base of the crown is eroded. The crown is much larger than the other Rangapur specimens (
<tableCitation id="C6860313A163FFADA05E0E6EFD4BA43F" box="[611,705,1586,1610]" captionStart="Table 2" captionStartId="5.[102,157,1298,1320]" captionText="Table 2. Measurements (in mm) and ratios of Rangapur and Naskal lower molars. Abbreviations as follows:L, length; TRW, trigonid width; TAW, talonid width; PHL, posthypocristid length; PEL, postentocristid length; PDH, protoconid height; MDH, metaconid height; AML, premetaconid length; TAH, talonid height; TAL, talonid length. Naskal data from Prasad et al. (1994)." pageId="6" pageNumber="337">Table 2</tableCitation>
). The trigonid is just wider than and over oneandahalf times taller than the talonid. Cusps on the trigonid are well spaced and somewhat inflated. The protoconid is the tallest cusp, but only slightly taller than the metaconid. The paraconid is small, labial, and slightly anteriorly projecting but not crestlike. The metaconid is more distally positioned than the protoconid, and its apex is somewhat lingually deflected. The protoconid is connected to the metaconid by a high protocristid, and to the paraconid by a very low paracristid. A broad notch separates the metaconid and paraconid. The anterolabial face of trigonid is slightly worn but preserves a faint anterior labial cingulid. It runs subvertically from the base of the crown below the protoconid, anterolingually toward the paraconid. It terminates below the notch formed by the paracristid. An anterior lingual cuspule is absent but the inflated base of the paraconid may have served the same function. In labial view, the trigonid appears tipped anteriorly (
<figureCitation id="133F2A2DA163FFADA6710B6DFB2EA13C" box="[1100,1188,817,841]" captionStart="Fig" captionStartId="6.[102,135,655,674]" captionTargetBox="[102,1485,227,627]" captionTargetId="figure-717@6.[102,1485,227,627]" captionTargetPageId="6" captionText="Fig. 4. Deccanolestes cf. robustus. Right m1 or m2 (ITV/R/Mm6) in occlusal (A, B), lingual (C), and labial (D) views. Scale bar 0.5 mm." pageId="6" pageNumber="337">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8BBB36A8A163FFADA16F0B0EFBF2A7FC" blockId="6.[812,1485,721,1417]" pageId="6" pageNumber="337">
In occlusal view, the posterior face of the trigonid contacts the talonid at an oblique angle (
<figureCitation id="133F2A2DA163FFADA6FF0B2DFAB3A1FC" box="[1218,1337,881,905]" captionStart="Fig" captionStartId="6.[102,135,655,674]" captionTargetBox="[102,1485,227,627]" captionTargetId="figure-717@6.[102,1485,227,627]" captionTargetPageId="6" captionText="Fig. 4. Deccanolestes cf. robustus. Right m1 or m2 (ITV/R/Mm6) in occlusal (A, B), lingual (C), and labial (D) views. Scale bar 0.5 mm." pageId="6" pageNumber="337">Fig. 4A, B</figureCitation>
). The talonid basin is wide but not very deep. It slopes downward in an anterolingual direction and is only weakly closed by a low entocristid on the lingual side. The talonid notch is deep. On the labial side, the cristid obliqua is higher than the entocristid and contacts the base of the trigonid just lingual to the protocristid notch below the base of the metaconid. A faint trace of a distal metacristid is present. The acute angle between the cristid obliqua and the posterior face of the trigonid forms a wellexcavated hypoflexid. The hypoconid is more anterior than the entoconid and is labially shifted. The hypoconulid is closer to the entoconid than it is to the hypoconid. The hypoconid is the tallest of the talonid cusps, followed by the hypoconulid, and then the entoconid. In posterior view, the talonid is canted lingually with respect to the trigonid, and the labial margins of the talonid and protoconid are convex. A postcingulid is absent.
</paragraph>
<paragraph id="8BBB36A8A163FFADA1110DC7FC52A487" blockId="6.[812,1486,1434,1778]" pageId="6" pageNumber="337">
<emphasis id="B970EABAA163FFADA1110DC7FC2DA7C7" box="[812,935,1435,1458]" italics="true" pageId="6" pageNumber="337">Discussion</emphasis>
.—In most respects, ITV/
<collectionCode id="ED15AE6DA163FFADA6FE0DC6FB5FA7C7" box="[1219,1237,1434,1458]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="337">R</collectionCode>
/Mm6 is very similar to the only known lower molar of
<taxonomicName id="4C044D2BA163FFADA6F00DE7FADEA7A7" authorityName="Prasad, Jaeger, Sahni, Gheerbrant, and Khajuria" authorityYear="1994" box="[1229,1364,1466,1490]" class="Mammalia" family="Adapisoriculidae" genus="Deccanolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="337" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B970EABAA163FFADA6F00DE7FADEA7A7" box="[1229,1364,1466,1490]" italics="true" pageId="6" pageNumber="337">D. robustus</emphasis>
</taxonomicName>
(
<collectionCode id="ED15AE6DA163FFADA75A0DE6FA17A7A7" box="[1383,1437,1466,1490]" country="USA" httpUri="http://grbio.org/cool/xxmh-b3dj" name="Vertebrate Paleontology Laboratory" pageId="6" pageNumber="337" type="Museum">VPL</collectionCode>
/JU/
<collectionCode id="ED15AE6DA163FFADA1110D86FCF0A787" box="[812,890,1498,1522]" pageId="6" pageNumber="337">NKIM</collectionCode>
/14, m1). It differs from the latter in both size and in the arrangement of the talonid cusps. ITV/
<collectionCode id="ED15AE6DA163FFADA7370DA6FA96A467" box="[1290,1308,1530,1554]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="337">R</collectionCode>
/Mm6 is about 24% larger than
<collectionCode id="ED15AE6DA163FFADA1D30E46FBAEA447" box="[1006,1060,1562,1586]" country="USA" httpUri="http://grbio.org/cool/xxmh-b3dj" name="Vertebrate Paleontology Laboratory" pageId="6" pageNumber="337" type="Museum">VPL</collectionCode>
/JU/
<collectionCode id="ED15AE6DA163FFADA6690E46FB15A447" box="[1108,1183,1562,1586]" pageId="6" pageNumber="337">NKIM</collectionCode>
/14 and has a hypoconulid that is closer to the entoconid than it is to the hypoconid (
<collectionCode id="ED15AE6DA163FFADA1090E06FCE0A407" box="[820,874,1626,1650]" country="USA" httpUri="http://grbio.org/cool/xxmh-b3dj" name="Vertebrate Paleontology Laboratory" pageId="6" pageNumber="337" type="Museum">VPL</collectionCode>
/JU/
<collectionCode id="ED15AE6DA163FFADA1A70E06FC6CA407" box="[922,998,1626,1650]" pageId="6" pageNumber="337">NKIM</collectionCode>
/14 has the reverse arrangement). These differences may be due to preservational, intraspecific, or interspecific variation. Because sample sizes are still very small, we tentatively refer this specimen to
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<emphasis id="B970EABAA163FFADA6F70EE7FAC7A4A7" box="[1226,1357,1722,1746]" italics="true" pageId="6" pageNumber="337">D. robustus</emphasis>
</taxonomicName>
rather than to a new taxon.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="4C044D2BA16DFFA3A25B0B97FE59A19D" authorityName="Rana &amp; Wilson" authorityYear="2003" box="[102,467,971,1000]" class="Mammalia" family="Adapisoriculidae" genus="Sahnitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="8" pageNumber="339" phylum="Chordata" rank="species" species="rangapurensis" status="sp. nov.">
<emphasis id="B970EABAA16DFFA3A25B0B97FE59A19D" box="[102,467,971,1000]" italics="true" pageId="8" pageNumber="339">Sahnitherium rangapurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357C1A16DFFA3A3E60B97FDB5A192" box="[475,575,971,999]" pageId="8" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BBB36A8A16DFFA3A25B0BAFFF73A67C" blockId="8.[102,575,971,1033]" box="[102,249,1011,1033]" pageId="8" pageNumber="339">
<tableCitation id="C6860313A16DFFA3A25B0BAFFF24A67C" box="[102,174,1011,1033]" captionStart="Table 1" captionStartId="4.[102,157,231,253]" captionText="Table 1. Measurements (in mm) and ratios of Rangapur and Naskal upper molars. Abbreviations as follows: AW, anterior width; PW, posterior width; PW1, posterior width sensu Butler (1990); PRW, protocone shelf width; MCL, postmetacrista length; BL, buccal length; PRL, protocone shelf length; PRH, protocone height; PAH, paracone height; MEH, metacone height. Naskal data from Prasad et al. (1994)." pageId="8" pageNumber="339">Table 1</tableCitation>
,
<figureCitation id="133F2A2DA16DFFA3A2870BAFFF7EA67C" box="[186,244,1011,1033]" captionStart="Fig" captionStartId="8.[812,845,1273,1292]" captionTargetBox="[812,1484,227,1235]" captionTargetId="figure-683@8.[812,1485,227,1235]" captionTargetPageId="8" captionText="Fig. 6. Sahnitherium rangapurensis. Right M1 or M2 (ITV/R/Mm1), holotype in occlusal (A, B), labial (C), anterior (D), and posterior (E) views. Scale bar 0.5 mm." pageId="8" pageNumber="339">Fig. 6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E6523A16DFFA2A25B0C47FD77A7C5" lastPageId="9" lastPageNumber="340" pageId="8" pageNumber="339" type="description">
<paragraph id="8BBB36A8A16DFFA3A25B0C47FDE6A638" blockId="8.[102,775,1051,1197]" pageId="8" pageNumber="339">
<emphasis id="B970EABAA16DFFA3A25B0C47FEABA645" box="[102,289,1051,1073]" italics="true" pageId="8" pageNumber="339">Derivation of name</emphasis>
:
<taxonomicName id="4C044D2BA16DFFA3A3130C47FE33A645" authorityName="Rana &amp; Wilson" authorityYear="2003" box="[302,441,1051,1072]" class="Mammalia" family="Adapisoriculidae" genus="Sahnitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="8" pageNumber="339" phylum="Chordata" rank="species" species="rangapurensis">
<emphasis id="B970EABAA16DFFA3A3130C47FE33A645" box="[302,441,1051,1072]" italics="true" pageId="8" pageNumber="339">rangapurensis</emphasis>
</taxonomicName>
<materialsCitation id="3B6C3CF5A16DFFA3A3820C47FDE2A638" country="India" location="Andhra Pradesh" pageId="8" pageNumber="339" specimenCount="1" stateProvince="Andhra Pradesh">
from the village of Rangapur near the
<typeStatus id="54BF880AA16DFFA3A2B40C6BFF38A638" box="[137,178,1079,1101]" pageId="8" pageNumber="339">type</typeStatus>
locality in the state of
<collectingRegion id="49C0F84AA16DFFA3A3B30C6BFDA0A638" box="[398,554,1079,1101]" country="India" name="Andhra Pradesh" pageId="8" pageNumber="339">Andhra Pradesh</collectingRegion>
,
<collectingCountry id="F3137638A16DFFA3A0090C6BFDE2A638" box="[564,616,1079,1101]" name="India" pageId="8" pageNumber="339">India</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBB36A8A16DFFA3A25B0C05FD4FA61A" blockId="8.[102,775,1051,1197]" box="[102,709,1113,1135]" pageId="8" pageNumber="339">
<materialsCitation id="3B6C3CF5A16DFFA3A25B0C05FD4FA61A" box="[102,709,1113,1135]" collectionCode="R" pageId="8" pageNumber="339" specimenCount="1" typeStatus="holotype">
<emphasis id="B970EABAA16DFFA3A25B0C05FF34A61A" box="[102,190,1113,1135]" italics="true" pageId="8" pageNumber="339">
<typeStatus id="54BF880AA16DFFA3A25B0C05FF34A61A" box="[102,190,1113,1135]" pageId="8" pageNumber="339" type="holotype">Holotype</typeStatus>
</emphasis>
: ITV/
<collectionCode id="ED15AE6DA16DFFA3A2C40C05FE83A61A" box="[249,265,1113,1135]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="339">R</collectionCode>
/Mm1, isolated right upper molar, M1 or M2.
</materialsCitation>
</paragraph>
<paragraph id="8BBB36A8A16DFFA3A25B0C20FE56A6D8" blockId="8.[102,775,1051,1197]" pageId="8" pageNumber="339">
<emphasis id="B970EABAA16DFFA3A25B0C20FF6BA6E4" box="[102,225,1147,1169]" italics="true" pageId="8" pageNumber="339">Type locality</emphasis>
: Terminal Cretaceous (Maastrichtian) Intertrappean horizon, Rangapur,
<collectingRegion id="49C0F84AA16DFFA3A2C30CCBFE13A6D8" box="[254,409,1175,1197]" country="India" name="Andhra Pradesh" pageId="8" pageNumber="339">Andhra Pradesh</collectingRegion>
,
<collectingCountry id="F3137638A16DFFA3A39E0CCBFE5DA6D8" box="[419,471,1175,1197]" name="India" pageId="8" pageNumber="339">India</collectingCountry>
.
</paragraph>
<paragraph id="8BBB36A8A16DFFA3A25B0CE3FDBEA682" blockId="8.[102,774,1215,1271]" pageId="8" pageNumber="339">
<emphasis id="B970EABAA16DFFA3A25B0CE3FF61A6A2" box="[102,235,1215,1239]" italics="true" pageId="8" pageNumber="339">Distribution</emphasis>
.—Late Cretaceous (Maastrichtian) Intertrappean beds of Rangapur,
<collectingRegion id="49C0F84AA16DFFA3A30B0C83FE63A682" box="[310,489,1247,1271]" country="India" name="Andhra Pradesh" pageId="8" pageNumber="339">Andhra Pradesh</collectingRegion>
,
<collectingCountry id="F3137638A16DFFA3A3C90C83FDBAA682" box="[500,560,1247,1271]" name="India" pageId="8" pageNumber="339">India</collectingCountry>
.
</paragraph>
<caption id="DF7B6620A16DFFA3A1110CA5FB98A730" pageId="8" pageNumber="339" startId="8.[812,845,1273,1292]" targetBox="[812,1484,227,1235]" targetPageId="8" targetType="figure">
<paragraph id="8BBB36A8A16DFFA3A1110CA5FB98A730" blockId="8.[812,1486,1273,1349]" pageId="8" pageNumber="339">
Fig. 6.
<taxonomicName id="4C044D2BA16DFFA3A14A0CA5FBF5A779" authorityName="Rana &amp; Wilson" authorityYear="2003" box="[887,1151,1273,1292]" class="Mammalia" family="Adapisoriculidae" genus="Sahnitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="8" pageNumber="339" phylum="Chordata" rank="species" species="rangapurensis">
<emphasis id="B970EABAA16DFFA3A14A0CA5FBF5A779" box="[887,1151,1273,1292]" italics="true" pageId="8" pageNumber="339">Sahnitherium rangapurensis</emphasis>
</taxonomicName>
. Right M1 or M2 (ITV/R/Mm1), holotype in occlusal (
<emphasis id="B970EABAA16DFFA3A1C60D49FB81A75D" bold="true" box="[1019,1035,1301,1320]" pageId="8" pageNumber="339">A</emphasis>
,
<emphasis id="B970EABAA16DFFA3A6260D49FBA0A75D" bold="true" box="[1051,1066,1301,1320]" pageId="8" pageNumber="339">B</emphasis>
), labial (
<emphasis id="B970EABAA16DFFA3A6B90D49FB1EA75D" bold="true" box="[1156,1172,1301,1320]" pageId="8" pageNumber="339">C</emphasis>
), anterior (
<emphasis id="B970EABAA16DFFA3A73E0D49FA99A75D" bold="true" box="[1283,1299,1301,1320]" pageId="8" pageNumber="339">D</emphasis>
), and posterior (
<emphasis id="B970EABAA16DFFA3A78B0D49FA4FA75D" bold="true" box="[1462,1477,1301,1320]" pageId="8" pageNumber="339">E</emphasis>
) views. Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BBB36A8A16DFFA3A25B0D54FD17A535" blockId="8.[102,775,1288,1856]" pageId="8" pageNumber="339">
<emphasis id="B970EABAA16DFFA3A25B0D54FEE8A755" box="[102,354,1288,1312]" italics="true" pageId="8" pageNumber="339">Differential diagnosis</emphasis>
.—Similar in size and in some morphological aspects to
<taxonomicName id="4C044D2BA16DFFA3A3580D74FDE9A735" box="[357,611,1320,1344]" class="Mammalia" family="Adapisoriculidae" genus="Deccanolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="8" pageNumber="339" phylum="Chordata" rank="species" species="hislopi">
<emphasis id="B970EABAA16DFFA3A3580D74FDE9A735" box="[357,611,1320,1344]" italics="true" pageId="8" pageNumber="339">Deccanolestes hislopi</emphasis>
</taxonomicName>
but differs in that it is not as transversely wide; the postmetacrista is stronger, longer, and carries the socalled cusp “C”; a stylocone is not present; the metastylar area is more posterolabially expanded and larger than the parastylar region; the ectoflexus is shallower; the labial cusps are labiolingually compressed, convex in labial view, and with steeper lingual slopes; paraconule larger and more anterolingually placed; and the protocone height is greater and its labial aspect meets the crown at a steeper angle. The taxon is also diagnosed by a paracone slightly higher and more lingually expanded than the metacone; bases of the labial cusps slightly appressed (moderate zalambdodonty); conular region not anteroposteriorly constricted; conules winglike with weak internal crests; protocone welldeveloped and somewhat anteroposteriorly expanded; the apex of the protocone anterolabially recumbent; and no cingula present.
</paragraph>
<paragraph id="8BBB36A8A16DFFA3A25B0F0EFBB4A79F" blockId="8.[102,775,1874,2026]" lastBlockId="8.[812,1485,1394,2026]" pageId="8" pageNumber="339">
<emphasis id="B970EABAA16DFFA3A25B0F0EFF60A51C" box="[102,234,1874,1897]" italics="true" pageId="8" pageNumber="339">Description</emphasis>
.—ITV/R/Mm1 is identified as a right upper molar, either M1 or M2 (
<figureCitation id="133F2A2DA16DFFA3A3B80F2EFE41A5FF" box="[389,459,1906,1930]" captionStart="Fig" captionStartId="8.[812,845,1273,1292]" captionTargetBox="[812,1484,227,1235]" captionTargetId="figure-683@8.[812,1485,227,1235]" captionTargetPageId="8" captionText="Fig. 6. Sahnitherium rangapurensis. Right M1 or M2 (ITV/R/Mm1), holotype in occlusal (A, B), labial (C), anterior (D), and posterior (E) views. Scale bar 0.5 mm." pageId="8" pageNumber="339">Fig. 6</figureCitation>
). The posterolabial and lingual roots are preserved. The enamel surface of the crown is pitted in some regions, and a small break can be seen in labial view at the anterolabial end of the crown (
<figureCitation id="133F2A2DA16DFFA3A0120F8EFD0FA59F" box="[559,645,2002,2026]" captionStart="Fig" captionStartId="8.[812,845,1273,1292]" captionTargetBox="[812,1484,227,1235]" captionTargetId="figure-683@8.[812,1485,227,1235]" captionTargetPageId="8" captionText="Fig. 6. Sahnitherium rangapurensis. Right M1 or M2 (ITV/R/Mm1), holotype in occlusal (A, B), labial (C), anterior (D), and posterior (E) views. Scale bar 0.5 mm." pageId="8" pageNumber="339">Fig. 6C</figureCitation>
). The break removed an anterolabial root and perhaps a parastylar hook and parastyle, if they existed. Otherwise, the labial margin of the crown, including the region where a stylocone would be preserved, is unaffected.
</paragraph>
<paragraph id="8BBB36A8A16DFFA2A16F0DAEFD80A075" blockId="8.[812,1485,1394,2026]" lastBlockId="9.[102,776,232,1123]" lastPageId="9" lastPageNumber="340" pageId="8" pageNumber="339">
Although the break slightly altered the buccal length and perhaps the anterior width, the crown was clearly not transversely wide (
<tableCitation id="C6860313A16DFFA3A1F90E6EFB9DA43F" box="[964,1047,1586,1610]" captionStart="Table 1" captionStartId="4.[102,157,231,253]" captionText="Table 1. Measurements (in mm) and ratios of Rangapur and Naskal upper molars. Abbreviations as follows: AW, anterior width; PW, posterior width; PW1, posterior width sensu Butler (1990); PRW, protocone shelf width; MCL, postmetacrista length; BL, buccal length; PRL, protocone shelf length; PRH, protocone height; PAH, paracone height; MEH, metacone height. Naskal data from Prasad et al. (1994)." pageId="8" pageNumber="339">Table 1</tableCitation>
). The main cusps of the crown are moderately tall and conical. The paracone is the tallest cusp, but it is only slightly taller than the metacone and protocone. The apices of the labial cusps are somewhat labiolingually compressed. The bases of these cusps are fused just above the level of the stylar shelf (moderate zalambdodonty). Centrocristae are weak. The base of the paracone is somewhat lingually expanded. The labial aspect of the paracone is flat to convex and gently slopes toward the labial margin of the tooth encroaching on the parastylar region. The labial aspect of the metacone is less convex and slopes more steeply toward the labial margin of the crown. The metastylar region slopes and then flattens but is not gutterlike. Also, this region is slightly posterolabially expanded. The postmetacrista is a distinct ridge that runs along a shallow gradient posterolabially toward the metastylar region. A metacrista cusp (cusp “C”) is present. The preparacrista is a less distinct ridge that runs along a steep gradient anterolabially toward the parastylar region. A faint trace of a raised ectocingulum is preserved on the labial margin of the stylar shelf, but a stylocone or accessory cusps are not present. The parastylar region (except a parastylar hook if one existed) is significantly smaller than the metastylar region. The ectoflexus forms a shallow curve.
</paragraph>
<paragraph id="8BBB36A8A16CFFA2A2B60A57FD4EA617" blockId="9.[102,776,232,1123]" pageId="9" pageNumber="340">
The paraconule is larger than the metaconule. It is shifted anteriorly and positioned midway between the paracone and protocone. The metaconule is positioned closer to the metacone than to the protocone. The postmetaconule wing terminates at or before the lingual base of the metacone. The preparaconule wing extends labially to the parastylar region. The preparaconule crista runs labially and dorsally along the anterior margin of the crown and becomes a faint trace near the parastylar region. Faint traces of internal wings are present. The trigon basin is deep, especially so toward the labial cusps. The step down to the trigon basin from the metaconule is small and from the paraconule it is negligible. The base of the protocone is somewhat lingually expanded and anteroposteriorly widened. The apex of the protocone is tall and anterolabially deflected. The labial face of the protocone is nearly perpendicular to the rest of the crown, as are the lingual faces of the paracone and metacone (
<figureCitation id="133F2A2DA16CFFA2A3C80C57FDC7A657" box="[501,589,1034,1059]" captionStart="Fig" captionStartId="8.[812,845,1273,1292]" captionTargetBox="[812,1484,227,1235]" captionTargetId="figure-683@8.[812,1485,227,1235]" captionTargetPageId="8" captionText="Fig. 6. Sahnitherium rangapurensis. Right M1 or M2 (ITV/R/Mm1), holotype in occlusal (A, B), labial (C), anterior (D), and posterior (E) views. Scale bar 0.5 mm." pageId="9" pageNumber="340">Fig. 6D</figureCitation>
). In occlusal outline, there is little to no anteroposterior waisting of the crown in the conular region (
<figureCitation id="133F2A2DA16CFFA2A3640C17FE3BA617" box="[345,433,1098,1123]" captionStart="Fig" captionStartId="8.[812,845,1273,1292]" captionTargetBox="[812,1484,227,1235]" captionTargetId="figure-683@8.[812,1485,227,1235]" captionTargetPageId="8" captionText="Fig. 6. Sahnitherium rangapurensis. Right M1 or M2 (ITV/R/Mm1), holotype in occlusal (A, B), labial (C), anterior (D), and posterior (E) views. Scale bar 0.5 mm." pageId="9" pageNumber="340">Fig. 6A</figureCitation>
). No cingula are present.
</paragraph>
<paragraph id="8BBB36A8A16CFFA2A25B0C24FD77A7C5" blockId="9.[102,775,1144,1456]" pageId="9" pageNumber="340">
<emphasis id="B970EABAA16CFFA2A25B0C24FF6BA6FA" box="[102,225,1144,1167]" italics="true" pageId="9" pageNumber="340">Discussion</emphasis>
.—The specimen (ITV/R/Mm1) is similar in size and in some morphological aspects to the M1 of
<taxonomicName id="4C044D2BA16CFFA2A0A40CC5FF7DA6A5" class="Mammalia" family="Adapisoriculidae" genus="Deccanolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="9" pageNumber="340" phylum="Chordata" rank="species" species="hislopi">
<emphasis id="B970EABAA16CFFA2A0A40CC5FF7DA6A5" italics="true" pageId="9" pageNumber="340">Deccanolestes hislopi</emphasis>
</taxonomicName>
(VPL/JU/NKIM/10), but it differs in a number of important ways outlined in the diagnosis. Based on evaluation of upper molars from other Cretaceous eutherians, we judge these differences to be greater than preservational, interspecific, or even intrageneric variation. Thus, we erect a new genus and species for the single isolated upper molar because we are unaware of any other Cretaceous or Paleocene boreosphenidan or australosphenidan with this morphology.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B965EA9AAB2E1E3C26D8E5D3FC1DFDE1" bold="true" box="[689,976,488,512]" pageId="34">Rhamphichthys heleios</emphasis>
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<emphasis id="B965EA9AAB2E1E3C26D8E5D3FC1DFDE1" bold="true" box="[689,976,488,512]" pageId="34" pageNumber="35">Rhamphichthys heleios</emphasis>
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(
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<figureCitation id="132A2A0DAB2E1E3C270CE608FC2CFDAA" box="[869,993,563,587]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStart-3="FIGURE 29" captionStartId-0="34.[488,561,1836,1856]" captionStartId-1="35.[488,561,1276,1296]" captionStartId-2="35.[488,561,1798,1818]" captionStartId-3="36.[488,561,716,736]" captionTargetBox-0="[488,1459,1446,1813]" captionTargetBox-1="[488,1459,585,1254]" captionTargetBox-2="[488,1459,1403,1784]" captionTargetBox-3="[488,1459,192,694]" captionTargetId-0="figure-336@34.[488,1459,1446,1813]" captionTargetId-1="figure-76@35.[488,1459,585,1254]" captionTargetId-2="figure-77@35.[488,1459,1403,1784]" captionTargetId-3="figure-355@36.[488,1459,192,694]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionTargetPageId-2="35" captionTargetPageId-3="36" captionText-0="FIGURE 26 | Holotype of Rhamphichthys heleios, INPA 42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil." captionText-1="FIGURE 27 | Detail of the head of holotype of Rhamphichthys heleios, INPA 42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil." captionText-2="FIGURE 28 | Paratype of Rhamphichthys heleios, juvenile, MCP 45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted)." captionText-3="FIGURE 29 | Detail of the head of paratype of Rhamphichthys heleios, MCP 45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted)." figureDoi-0="http://doi.org/10.5281/zenodo.13326949" figureDoi-1="http://doi.org/10.5281/zenodo.13326951" figureDoi-2="http://doi.org/10.5281/zenodo.13326953" figureDoi-3="http://doi.org/10.5281/zenodo.13326955" httpUri-0="https://zenodo.org/record/13326949/files/figure.png" httpUri-1="https://zenodo.org/record/13326951/files/figure.png" httpUri-2="https://zenodo.org/record/13326953/files/figure.png" httpUri-3="https://zenodo.org/record/13326955/files/figure.png" pageId="34" pageNumber="35">Fig. 2629</figureCitation>
;
<tableCitation id="C6930333AB2E1E3C2785E608FBF8FDAA" box="[1004,1077,563,587]" captionStart="TABLE 7" captionStartId="37.[488,550,193,213]" captionText="TABLE 7 | Morphometric and meristic data for Rhamphichthys heleios. H = holotype, SD = standard deviation, n = number of specimens." pageId="34">Tab. 7</tableCitation>
<tableCitation id="C6930333AB2E1E3C2785E608FBF8FDAA" box="[1004,1077,563,587]" captionStart="TABLE 7" captionStartId="37.[488,550,193,213]" captionText="TABLE 7 | Morphometric and meristic data for Rhamphichthys heleios. H = holotype, SD = standard deviation, n = number of specimens." pageId="34" pageNumber="35">Tab. 7</tableCitation>
)
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(
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description, type-locality:
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,
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,
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, confluence of
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and
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<collectingCounty id="62CF4E04AB2E1E3C27BAE69CFBA2FD5B" box="[979,1135,679,698]" pageId="34" pageNumber="35">Costa do Catalão</collectingCounty>
,
<geoCoordinate id="EE25504FAB2E1E3C2011E69CFB73FD5B" box="[1144,1214,679,698]" degrees="03" direction="south" minutes="09" orientation="latitude" pageId="34" precision="925" value="-3.15">03°09S</geoCoordinate>
<geoCoordinate id="EE25504FAB2E1E3C20ADE69CFAD7FD5B" box="[1220,1306,679,698]" degrees="59" direction="west" minutes="54" orientation="longitude" pageId="34" precision="925" value="-59.9">59°54W</geoCoordinate>
<geoCoordinate id="EE25504FAB2E1E3C2011E69CFB73FD5B" box="[1144,1214,679,698]" degrees="03" direction="south" minutes="09" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="-3.15">03°09S</geoCoordinate>
<geoCoordinate id="EE25504FAB2E1E3C20ADE69CFAD7FD5B" box="[1220,1306,679,698]" degrees="59" direction="west" minutes="54" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="-59.9">59°54W</geoCoordinate>
). —Ferraris
<emphasis id="B965EA9AAB2E1E3C21EDE69CFA65FD5B" box="[1412,1448,679,698]" italics="true" pageId="34">et al</emphasis>
<emphasis id="B965EA9AAB2E1E3C21EDE69CFA65FD5B" box="[1412,1448,679,698]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 2017:28 (listed). —Giora, Carvalho, 2018:1060 (accessory electric organ anatomy). —Janzen
<emphasis id="B965EA9AAB2E1E3C2122E6F6FABDFD3E" box="[1355,1392,716,735]" italics="true" pageId="34">et al</emphasis>
<emphasis id="B965EA9AAB2E1E3C2122E6F6FABDFD3E" box="[1355,1392,716,735]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 2022: tab. 1 (barcode library).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30B6503AB2E1E3C266FE703FD6CFB41" pageId="34" type="diagnosis">
<paragraph id="8BAE3688AB2E1E3C266FE703FD6CFB41" blockId="34.[488,1461,824,1184]" pageId="34">
<emphasis id="B965EA9AAB2E1E3C266FE703FD4BFCB1" bold="true" box="[518,646,824,848]" pageId="34">Diagnosis.</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C26FDE703FC45FCB1" authorityName="Carvalho &amp; Albert" authorityYear="2015" box="[660,904,824,848]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2E1E3C26FDE703FC45FCB1" box="[660,904,824,848]" italics="true" pageId="34">Rhamphichthys heleios</emphasis>
<subSubSection id="C30B6503AB2E1E3C266FE703FD6CFB41" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BAE3688AB2E1E3C266FE703FD6CFB41" blockId="34.[488,1461,824,1184]" pageId="34" pageNumber="35">
<emphasis id="B965EA9AAB2E1E3C266FE703FD4BFCB1" bold="true" box="[518,646,824,848]" pageId="34" pageNumber="35">Diagnosis.</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C26FDE703FC45FCB1" authorityName="Carvalho &amp; Albert" authorityYear="2015" box="[660,904,824,848]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2E1E3C26FDE703FC45FCB1" box="[660,904,824,848]" italics="true" pageId="34" pageNumber="35">Rhamphichthys heleios</emphasis>
</taxonomicName>
differs from all congeners by the presence of irregular dark rounded blotches along the lateral body surface over the lateral line, sometimes connected forming an irregular longitudinal dark stripe,
<emphasis id="B965EA9AAB2E1E3C217EE7BFFAFCFC7A" box="[1303,1329,900,923]" italics="true" pageId="34">vs.</emphasis>
<emphasis id="B965EA9AAB2E1E3C217EE7BFFAFCFC7A" box="[1303,1329,900,923]" italics="true" pageId="34" pageNumber="35">vs.</emphasis>
diagonally displaced dark bands on the ventrolateral surface of the body over this region sometimes faded in
<emphasis id="B965EA9AAB2E1E3C2625E7F4FD0FFC07" box="[588,706,974,998]" italics="true" pageId="34">
<taxonomicName id="4C114D0BAB2E1E3C2625E7F4FD72FC07" box="[588,703,974,998]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="lineatus">R. lineatus</taxonomicName>
<emphasis id="B965EA9AAB2E1E3C2625E7F4FD0FFC07" box="[588,706,974,998]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="4C114D0BAB2E1E3C2625E7F4FD72FC07" box="[588,703,974,998]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="lineatus">R. lineatus</taxonomicName>
.
</emphasis>
It differs from
<taxonomicName id="4C114D0BAB2E1E3C2706E7F4FC12FC07" box="[879,991,974,998]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B965EA9AAB2E1E3C2706E7F4FC12FC07" box="[879,991,974,998]" italics="true" pageId="34">R. lineatus</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C2706E7F4FC12FC07" box="[879,991,974,998]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B965EA9AAB2E1E3C2706E7F4FC12FC07" box="[879,991,974,998]" italics="true" pageId="34" pageNumber="35">R. lineatus</emphasis>
</taxonomicName>
by an overall dark,
<emphasis id="B965EA9AAB2E1E3C20AAE7F4FB13FC07" box="[1219,1246,975,998]" italics="true" pageId="34">vs.</emphasis>
<emphasis id="B965EA9AAB2E1E3C20AAE7F4FB13FC07" box="[1219,1246,975,998]" italics="true" pageId="34" pageNumber="35">vs.</emphasis>
light color pattern of the body. It differs from
<taxonomicName id="4C114D0BAB2E1E3C274CE7CFFC53FBEA" box="[805,926,1012,1035]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B965EA9AAB2E1E3C274CE7CFFC53FBEA" box="[805,926,1012,1035]" italics="true" pageId="34">R. rostratus</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C274CE7CFFC53FBEA" box="[805,926,1012,1035]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B965EA9AAB2E1E3C274CE7CFFC53FBEA" box="[805,926,1012,1035]" italics="true" pageId="34" pageNumber="35">R. rostratus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D0BAB2E1E3C27B0E7CFFB97FBEA" box="[985,1114,1012,1035]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="apurensis">
<emphasis id="B965EA9AAB2E1E3C27B0E7CFFB97FBEA" box="[985,1114,1012,1035]" italics="true" pageId="34">R. apurensis</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C27B0E7CFFB97FBEA" box="[985,1114,1012,1035]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="apurensis">
<emphasis id="B965EA9AAB2E1E3C27B0E7CFFB97FBEA" box="[985,1114,1012,1035]" italics="true" pageId="34" pageNumber="35">R. apurensis</emphasis>
</taxonomicName>
by a short snout (50.555.1% of HL),
<emphasis id="B965EA9AAB2E1E3C262AE022FD90FBD1" box="[579,605,1049,1072]" italics="true" pageId="34">vs.</emphasis>
<emphasis id="B965EA9AAB2E1E3C262AE022FD90FBD1" box="[579,605,1049,1072]" italics="true" pageId="34" pageNumber="35">vs.</emphasis>
long snout (58.465.1% of HL); by low number of anal-fin rays (320345),
<emphasis id="B965EA9AAB2E1E3C2581E004FDCFFBB7" box="[488,514,1087,1110]" italics="true" pageId="34">vs.</emphasis>
<emphasis id="B965EA9AAB2E1E3C2581E004FDCFFBB7" box="[488,514,1087,1110]" italics="true" pageId="34" pageNumber="35">vs.</emphasis>
high number of anal-fin rays (347455). It differs from
<taxonomicName id="4C114D0BAB2E1E3C20E4E004FAD0FBB7" box="[1165,1309,1086,1110]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="drepanium">
<emphasis id="B965EA9AAB2E1E3C20E4E004FAD0FBB7" box="[1165,1309,1086,1110]" italics="true" pageId="34">R. drepanium</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C20E4E004FAD0FBB7" box="[1165,1309,1086,1110]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="drepanium">
<emphasis id="B965EA9AAB2E1E3C20E4E004FAD0FBB7" box="[1165,1309,1086,1110]" italics="true" pageId="34" pageNumber="35">R. drepanium</emphasis>
</taxonomicName>
,
<taxonomicName id="4C114D0BAB2E1E3C2147E004FA47FBB7" box="[1326,1418,1086,1110]" family="Apteronotidae" genus="Sternarchorhamphus" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="hahni">
<emphasis id="B965EA9AAB2E1E3C2147E004FA47FBB7" box="[1326,1418,1086,1110]" italics="true" pageId="34">R. hahni</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C2147E004FA47FBB7" box="[1326,1418,1086,1110]" family="Apteronotidae" genus="Sternarchorhamphus" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="hahni">
<emphasis id="B965EA9AAB2E1E3C2147E004FA47FBB7" box="[1326,1418,1086,1110]" italics="true" pageId="34" pageNumber="35">R. hahni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C114D0BAB2E1E3C21F2E004FDF2FB9A" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B965EA9AAB2E1E3C21F2E004FDF2FB9A" italics="true" pageId="34">R. rostratus</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C21F2E004FDF2FB9A" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B965EA9AAB2E1E3C21F2E004FDF2FB9A" italics="true" pageId="34" pageNumber="35">R. rostratus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C114D0BAB2E1E3C2626E05FFD1DFB9A" box="[591,720,1124,1147]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="apurensis">
<emphasis id="B965EA9AAB2E1E3C2626E05FFD1DFB9A" box="[591,720,1124,1147]" italics="true" pageId="34">R. apurensis</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C2626E05FFD1DFB9A" box="[591,720,1124,1147]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="apurensis">
<emphasis id="B965EA9AAB2E1E3C2626E05FFD1DFB9A" box="[591,720,1124,1147]" italics="true" pageId="34" pageNumber="35">R. apurensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C114D0BAB2E1E3C277DE05FFC63FB9A" authorityName="Castelnau" authorityYear="1855" box="[788,942,1123,1147]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="pantherinus">
<emphasis id="B965EA9AAB2E1E3C277DE05FFC63FB9A" box="[788,942,1123,1147]" italics="true" pageId="34">R. pantherinus</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C277DE05FFC63FB9A" authorityName="Castelnau" authorityYear="1855" box="[788,942,1123,1147]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="pantherinus">
<emphasis id="B965EA9AAB2E1E3C277DE05FFC63FB9A" box="[788,942,1123,1147]" italics="true" pageId="34" pageNumber="35">R. pantherinus</emphasis>
</taxonomicName>
by the absence of dark transversal saddles on the mid dorsum.
</paragraph>
</subSubSection>
<subSubSection id="C30B6503AB2E1E3A266FE0E8FC4BF8FA" lastPageId="36" pageId="34" type="description">
<paragraph id="8BAE3688AB2E1E3D266FE0E8FBA2FDE1" blockId="34.[488,1459,1235,1371]" lastBlockId="35.[488,1459,190,512]" lastPageId="35" pageId="34">
<emphasis id="B965EA9AAB2E1E3C266FE0E8FD51FB0A" bold="true" box="[518,668,1235,1259]" pageId="34">Description.</emphasis>
<subSubSection id="C30B6503AB2E1E3A266FE0E8FC4BF8FA" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
<paragraph id="8BAE3688AB2E1E3D266FE0E8FBA2FDE1" blockId="34.[488,1459,1235,1371]" lastBlockId="35.[488,1459,190,512]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="B965EA9AAB2E1E3C266FE0E8FD51FB0A" bold="true" box="[518,668,1235,1259]" pageId="34" pageNumber="35">Description.</emphasis>
Morphometrics and meristics given in
<tableCitation id="C6930333AB2E1E3C200AE0E8FB7DFB0A" box="[1123,1200,1235,1259]" captionStart="TABLE 7" captionStartId="37.[488,550,193,213]" captionText="TABLE 7 | Morphometric and meristic data for Rhamphichthys heleios. H = holotype, SD = standard deviation, n = number of specimens." pageId="34">Tab. 7</tableCitation>
<tableCitation id="C6930333AB2E1E3C200AE0E8FB7DFB0A" box="[1123,1200,1235,1259]" captionStart="TABLE 7" captionStartId="37.[488,550,193,213]" captionText="TABLE 7 | Morphometric and meristic data for Rhamphichthys heleios. H = holotype, SD = standard deviation, n = number of specimens." pageId="34" pageNumber="35">Tab. 7</tableCitation>
. Adult body size small to moderate compared with other congeners, maximum size
<quantity id="4CE99B6DAB2E1E3C20DDE0C2FAD8FAF1" box="[1204,1301,1273,1297]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="34" unit="mm" value="400.0">400 mm</quantity>
<quantity id="4CE99B6DAB2E1E3C20DDE0C2FAD8FAF1" box="[1204,1301,1273,1297]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="35" unit="mm" value="400.0">400 mm</quantity>
LEA. Mouth subterminal. Snout relatively short and robust, size about half of head length. Dorsal profile of snout strongly convex from snout tip to nares; concave from this point to eye and convex from this point to supraoccipital. Anterior nares positioned terminally; posterior nares located closer to snout than eyes at about one third of length. Body profile almost straight to slightly concave dorsally. Ventral body profile slightly concave from analfin origin to end of anal fin. Greatest body depth slightly at vertical of posterior end of anal fin or posterior to end of body cavity. Body tapering posterior to mid body. Eyes relatively large and positioned laterally. Urogenital papilla developed and anteriorly positioned below eyes in specimens larger than
<quantity id="4CE99B6DAB2F1E3D20FEE5A5FB3FFE54" box="[1175,1266,414,438]" metricMagnitude="-1" metricUnit="m" metricValue="3.3" pageId="35" unit="mm" value="330.0">330 mm</quantity>
<quantity id="4CE99B6DAB2F1E3D20FEE5A5FB3FFE54" box="[1175,1266,414,438]" metricMagnitude="-1" metricUnit="m" metricValue="3.3" pageId="35" pageNumber="36" unit="mm" value="330.0">330 mm</quantity>
LEA. Urogenital papilla large. Posterior gas bladder reduced with thickened walls. Caudal appendage laterally compressed, its depth about three times its width.
</paragraph>
<caption id="DF6E6600AB2E1E3C2581E317FC19F884" pageId="34" startId="34.[488,561,1836,1856]" targetBox="[488,1459,1446,1813]" targetPageId="34" targetType="figure">
<paragraph id="8BAE3688AB2E1E3C2581E317FC19F884" blockId="34.[488,1459,1836,1893]" pageId="34">
<emphasis id="B965EA9AAB2E1E3C2581E317FDA9F8A1" bold="true" box="[488,612,1836,1856]" pageId="34">FIGURE 26 |</emphasis>
<caption id="DF6E6600AB2E1E3C2581E317FC19F884" ID-DOI="http://doi.org/10.5281/zenodo.13326949" ID-Zenodo-Dep="13326949" httpUri="https://zenodo.org/record/13326949/files/figure.png" pageId="34" pageNumber="35" startId="34.[488,561,1836,1856]" targetBox="[488,1459,1446,1813]" targetPageId="34" targetType="figure">
<paragraph id="8BAE3688AB2E1E3C2581E317FC19F884" blockId="34.[488,1459,1836,1893]" pageId="34" pageNumber="35">
<emphasis id="B965EA9AAB2E1E3C2581E317FDA9F8A1" bold="true" box="[488,612,1836,1856]" pageId="34" pageNumber="35">FIGURE 26 |</emphasis>
Holotype of
<taxonomicName id="4C114D0BAB2E1E3C26B6E317FC2FF8A1" authority=", INPA" authorityName="INPA" box="[735,994,1836,1856]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2E1E3C26B6E317FC69F8A1" bold="true" box="[735,932,1836,1856]" italics="true" pageId="34">Rhamphichthys heleios</emphasis>
<taxonomicName id="4C114D0BAB2E1E3C26B6E317FC2FF8A1" authority=", INPA" authorityName="INPA" box="[735,994,1836,1856]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2E1E3C26B6E317FC69F8A1" bold="true" box="[735,932,1836,1856]" italics="true" pageId="34" pageNumber="35">Rhamphichthys heleios</emphasis>
, INPA
</taxonomicName>
42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil.
</paragraph>
</caption>
<caption id="DF6E6600AB2F1E3D2581E0C7FBBDFAD4" pageId="35" startId="35.[488,561,1276,1296]" targetBox="[488,1459,585,1254]" targetPageId="35" targetType="figure">
<paragraph id="8BAE3688AB2F1E3D2581E0C7FBBDFAD4" blockId="35.[487,1459,1276,1333]" pageId="35">
<emphasis id="B965EA9AAB2F1E3D2581E0C7FD90FAF1" bold="true" box="[488,605,1276,1296]" pageId="35">FIGURE 27 |</emphasis>
<caption id="DF6E6600AB2F1E3D2581E0C7FBBDFAD4" ID-DOI="http://doi.org/10.5281/zenodo.13326951" ID-Zenodo-Dep="13326951" httpUri="https://zenodo.org/record/13326951/files/figure.png" pageId="35" pageNumber="36" startId="35.[488,561,1276,1296]" targetBox="[488,1459,585,1254]" targetPageId="35" targetType="figure">
<paragraph id="8BAE3688AB2F1E3D2581E0C7FBBDFAD4" blockId="35.[487,1459,1276,1333]" pageId="35" pageNumber="36">
<emphasis id="B965EA9AAB2F1E3D2581E0C7FD90FAF1" bold="true" box="[488,605,1276,1296]" pageId="35" pageNumber="36">FIGURE 27 |</emphasis>
Detail of the head of holotype of
<taxonomicName id="4C114D0BAB2F1E3D27E5E0C7FB40FAF1" authority=", INPA" authorityName="INPA" box="[908,1165,1276,1296]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="35" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2F1E3D27E5E0C7FB9CFAF1" bold="true" box="[908,1105,1276,1296]" italics="true" pageId="35">Rhamphichthys heleios</emphasis>
<taxonomicName id="4C114D0BAB2F1E3D27E5E0C7FB40FAF1" authority=", INPA" authorityName="INPA" box="[908,1165,1276,1296]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2F1E3D27E5E0C7FB9CFAF1" bold="true" box="[908,1105,1276,1296]" italics="true" pageId="35" pageNumber="36">Rhamphichthys heleios</emphasis>
, INPA
</taxonomicName>
42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil.
</paragraph>
</caption>
<caption id="DF6E6600AB2F1E3D2581E33DFB59F8A1" pageId="35" startId="35.[488,561,1798,1818]" targetBox="[488,1459,1403,1784]" targetPageId="35" targetType="figure">
<paragraph id="8BAE3688AB2F1E3D2581E33DFB59F8A1" blockId="35.[488,1460,1798,1856]" pageId="35">
<emphasis id="B965EA9AAB2F1E3D2581E33DFDACF8FB" bold="true" box="[488,609,1798,1818]" pageId="35">FIGURE 28 |</emphasis>
<caption id="DF6E6600AB2F1E3D2581E33DFB59F8A1" ID-DOI="http://doi.org/10.5281/zenodo.13326953" ID-Zenodo-Dep="13326953" httpUri="https://zenodo.org/record/13326953/files/figure.png" pageId="35" pageNumber="36" startId="35.[488,561,1798,1818]" targetBox="[488,1459,1403,1784]" targetPageId="35" targetType="figure">
<paragraph id="8BAE3688AB2F1E3D2581E33DFB59F8A1" blockId="35.[488,1460,1798,1856]" pageId="35" pageNumber="36">
<emphasis id="B965EA9AAB2F1E3D2581E33DFDACF8FB" bold="true" box="[488,609,1798,1818]" pageId="35" pageNumber="36">FIGURE 28 |</emphasis>
Paratype of
<taxonomicName id="4C114D0BAB2F1E3D26BFE33CFC51F8FA" box="[726,924,1799,1819]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="35" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2F1E3D26BFE33CFC51F8FA" bold="true" box="[726,924,1799,1819]" italics="true" pageId="35">Rhamphichthys heleios</emphasis>
<taxonomicName id="4C114D0BAB2F1E3D26BFE33CFC51F8FA" box="[726,924,1799,1819]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB2F1E3D26BFE33CFC51F8FA" bold="true" box="[726,924,1799,1819]" italics="true" pageId="35" pageNumber="36">Rhamphichthys heleios</emphasis>
</taxonomicName>
, juvenile, MCP 45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted).
</paragraph>
</caption>
<caption id="DF6E6600AB281E3A2581E6F7FAD0FCE4" pageId="36" startId="36.[488,561,716,736]" targetBox="[488,1459,192,694]" targetPageId="36" targetType="figure">
<paragraph id="8BAE3688AB281E3A2581E6F7FAD0FCE4" blockId="36.[488,1459,716,773]" pageId="36">
<emphasis id="B965EA9AAB281E3A2581E6F7FDA8FD01" bold="true" box="[488,613,716,736]" pageId="36">FIGURE 29 |</emphasis>
<caption id="DF6E6600AB281E3A2581E6F7FAD0FCE4" ID-DOI="http://doi.org/10.5281/zenodo.13326955" ID-Zenodo-Dep="13326955" httpUri="https://zenodo.org/record/13326955/files/figure.png" pageId="36" pageNumber="37" startId="36.[488,561,716,736]" targetBox="[488,1459,192,694]" targetPageId="36" targetType="figure">
<paragraph id="8BAE3688AB281E3A2581E6F7FAD0FCE4" blockId="36.[488,1459,716,773]" pageId="36" pageNumber="37">
<emphasis id="B965EA9AAB281E3A2581E6F7FDA8FD01" bold="true" box="[488,613,716,736]" pageId="36" pageNumber="37">FIGURE 29 |</emphasis>
Detail of the head of paratype of
<taxonomicName id="4C114D0BAB281E3A27C3E6F7FB65FD01" authority=", MCP" authorityName="MCP" box="[938,1192,716,736]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="36" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB281E3A27C3E6F7FBA2FD01" bold="true" box="[938,1135,716,736]" italics="true" pageId="36">Rhamphichthys heleios</emphasis>
<taxonomicName id="4C114D0BAB281E3A27C3E6F7FB65FD01" authority=", MCP" authorityName="MCP" box="[938,1192,716,736]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB281E3A27C3E6F7FBA2FD01" bold="true" box="[938,1135,716,736]" italics="true" pageId="36" pageNumber="37">Rhamphichthys heleios</emphasis>
, MCP
</taxonomicName>
45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted).
</paragraph>
</caption>
<paragraph id="8BAE3688AB281E3A266FE765FA84FBB7" blockId="36.[487,1460,862,1110]" pageId="36">
<paragraph id="8BAE3688AB281E3A266FE765FA84FBB7" blockId="36.[487,1460,862,1110]" pageId="36" pageNumber="37">
Body entirely covered by scales in adults, anterior and dorsal areas naked in juveniles. Scales cycloid elongate in shape posteriorly, length twice as depth. Lateral line complete with ossified tubules, anterior portion of presenting ventral rami. Pectoral fin with 1619 (mode 19) rays, dorsal rays longer than ventral rays, distal fin margin slightly rounded. Anal fin with 320345 (median 338) rays. Unbranched 2233 analfin rays anteriorly. Analfin origin at vertical of anterior portion of opercle. Precaudal vertebrae 19 (
<tableCitation id="C6930333AB281E3A267BE005FD90FBB7" box="[530,605,1086,1110]" captionStart="TABLE 5" captionStartId="20.[488,550,193,213]" captionText="TABLE 5 | Number of specimens observed for precaudal vertebrae (PCV) and caudal vertebrae (CV) counts in Rhamphichthys." pageId="36">Tab. 5</tableCitation>
<tableCitation id="C6930333AB281E3A267BE005FD90FBB7" box="[530,605,1086,1110]" captionStart="TABLE 5" captionStartId="20.[488,550,193,213]" captionText="TABLE 5 | Number of specimens observed for precaudal vertebrae (PCV) and caudal vertebrae (CV) counts in Rhamphichthys." pageId="36" pageNumber="37">Tab. 5</tableCitation>
). Displaced hemal spines 12 (n = 1). Sexual dimorphism unknown.
</paragraph>
<paragraph id="8BAE3688AB281E3A266FE0B3FB16F981" blockId="36.[487,1459,1160,1632]" pageId="36">
<emphasis id="B965EA9AAB281E3A266FE0B3FCDCFB40" bold="true" box="[518,785,1160,1185]" pageId="36">Coloration in alcohol.</emphasis>
<paragraph id="8BAE3688AB281E3A266FE0B3FB16F981" blockId="36.[487,1459,1160,1632]" pageId="36" pageNumber="37">
<emphasis id="B965EA9AAB281E3A266FE0B3FCDCFB40" bold="true" box="[518,785,1160,1185]" pageId="36" pageNumber="37">Coloration in alcohol.</emphasis>
Ground color of dorsal and lateral surfaces of head and body light brown to pale yellow (
<figureCitation id="132A2A0DAB281E3A2770E095FC53FB27" box="[793,926,1198,1222]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStart-3="FIGURE 29" captionStartId-0="34.[488,561,1836,1856]" captionStartId-1="35.[488,561,1276,1296]" captionStartId-2="35.[488,561,1798,1818]" captionStartId-3="36.[488,561,716,736]" captionTargetBox-0="[488,1459,1446,1813]" captionTargetBox-1="[488,1459,585,1254]" captionTargetBox-2="[488,1459,1403,1784]" captionTargetBox-3="[488,1459,192,694]" captionTargetId-0="figure-336@34.[488,1459,1446,1813]" captionTargetId-1="figure-76@35.[488,1459,585,1254]" captionTargetId-2="figure-77@35.[488,1459,1403,1784]" captionTargetId-3="figure-355@36.[488,1459,192,694]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionTargetPageId-2="35" captionTargetPageId-3="36" captionText-0="FIGURE 26 | Holotype of Rhamphichthys heleios, INPA 42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil." captionText-1="FIGURE 27 | Detail of the head of holotype of Rhamphichthys heleios, INPA 42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil." captionText-2="FIGURE 28 | Paratype of Rhamphichthys heleios, juvenile, MCP 45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted)." captionText-3="FIGURE 29 | Detail of the head of paratype of Rhamphichthys heleios, MCP 45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted)." pageId="36">Figs. 2629</figureCitation>
<figureCitation id="132A2A0DAB281E3A2770E095FC53FB27" box="[793,926,1198,1222]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStart-3="FIGURE 29" captionStartId-0="34.[488,561,1836,1856]" captionStartId-1="35.[488,561,1276,1296]" captionStartId-2="35.[488,561,1798,1818]" captionStartId-3="36.[488,561,716,736]" captionTargetBox-0="[488,1459,1446,1813]" captionTargetBox-1="[488,1459,585,1254]" captionTargetBox-2="[488,1459,1403,1784]" captionTargetBox-3="[488,1459,192,694]" captionTargetId-0="figure-336@34.[488,1459,1446,1813]" captionTargetId-1="figure-76@35.[488,1459,585,1254]" captionTargetId-2="figure-77@35.[488,1459,1403,1784]" captionTargetId-3="figure-355@36.[488,1459,192,694]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionTargetPageId-2="35" captionTargetPageId-3="36" captionText-0="FIGURE 26 | Holotype of Rhamphichthys heleios, INPA 42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil." captionText-1="FIGURE 27 | Detail of the head of holotype of Rhamphichthys heleios, INPA 42309, 375 mm LEA, confluence of rio Amazonas and Solimões at Costa do Catalão, Amazonas, Brazil." captionText-2="FIGURE 28 | Paratype of Rhamphichthys heleios, juvenile, MCP 45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted)." captionText-3="FIGURE 29 | Detail of the head of paratype of Rhamphichthys heleios, MCP 45545, 133 mm LEA, ilha do Martelo, Tefé, Amazonas, Brazil. Picture of the right side (figure horizontally inverted)." figureDoi-0="http://doi.org/10.5281/zenodo.13326949" figureDoi-1="http://doi.org/10.5281/zenodo.13326951" figureDoi-2="http://doi.org/10.5281/zenodo.13326953" figureDoi-3="http://doi.org/10.5281/zenodo.13326955" httpUri-0="https://zenodo.org/record/13326949/files/figure.png" httpUri-1="https://zenodo.org/record/13326951/files/figure.png" httpUri-2="https://zenodo.org/record/13326953/files/figure.png" httpUri-3="https://zenodo.org/record/13326955/files/figure.png" pageId="36" pageNumber="37">Figs. 2629</figureCitation>
). Head with scattered large dark brown blotches of about eye; snout mostly dark, ventral margin less pigmented. Dorsum of body dark, presenting darker irregular marks laterally displaced not forming saddles. Presence of large dark blotches over the midlateral surface of body running from humeral region to a vertical of the end of anal fin; blotches sometimes coalescent forming an irregular longitudinal stripe. Lateral line clearer forming an inconspicuous light longitudinal stripe. Two pairs of whitish stripes running on the lateral surface of body, one over and another below the midlateral region presenting the dark blotches. Lateroventral region brownish with darker rounded or vermiculous blotches over pterygiophores region. Anal fin mostly clear or hyaline, with scattered chromatophores, these forming round blotches on the proximal and posterior regions. Pectoral fin clear with darkish irregular bars. Caudal appendage light brown with scattered darker blotches.
</paragraph>
<paragraph id="8BAE3688AB281E3A266FE2A8FC4BF8FA" blockId="36.[488,1459,1683,1819]" pageId="36">
<emphasis id="B965EA9AAB281E3A266FE2A8FC8AF94A" bold="true" box="[518,839,1683,1707]" pageId="36">Geographical distribution.</emphasis>
<taxonomicName id="4C114D0BAB281E3A2726E2A8FBF0F94A" box="[847,1085,1683,1707]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="36" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB281E3A2726E2A8FBF0F94A" box="[847,1085,1683,1707]" italics="true" pageId="36">Rhamphichthys heleios</emphasis>
<paragraph id="8BAE3688AB281E3A266FE2A8FC4BF8FA" blockId="36.[488,1459,1683,1819]" pageId="36" pageNumber="37">
<emphasis id="B965EA9AAB281E3A266FE2A8FC8AF94A" bold="true" box="[518,839,1683,1707]" pageId="36" pageNumber="37">Geographical distribution.</emphasis>
<taxonomicName id="4C114D0BAB281E3A2726E2A8FBF0F94A" box="[847,1085,1683,1707]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB281E3A2726E2A8FBF0F94A" box="[847,1085,1683,1707]" italics="true" pageId="36" pageNumber="37">Rhamphichthys heleios</emphasis>
</taxonomicName>
is known from the Solimões and Amazon Rivers in
<collectingCountry id="F3067618AB281E3A26D1E283FD37F931" box="[696,762,1720,1744]" name="Brazil" pageId="36">Brazil</collectingCountry>
<collectingCountry id="F3067618AB281E3A26D1E283FD37F931" box="[696,762,1720,1744]" name="Brazil" pageId="36" pageNumber="37">Brazil</collectingCountry>
upstream of the mouth of the Madeira River (
<figureCitation id="132A2A0DAB281E3A2097E283FA9DF930" box="[1278,1360,1720,1745]" captionStart="FIGURE 30" captionStartId="37.[136,209,1873,1893]" captionTargetBox="[136,1459,1122,1851]" captionTargetId="figure-365@37.[136,1459,1122,1851]" captionTargetPageId="37" captionText="FIGURE 30 | Distribution of Rhamphichthys heleios based on examined museum specimens. Red dot represents the type-locality." pageId="36">Fig. 30</figureCitation>
<figureCitation id="132A2A0DAB281E3A2097E283FA9DF930" box="[1278,1360,1720,1745]" captionStart="FIGURE 30" captionStartId="37.[136,209,1873,1893]" captionTargetBox="[136,1459,1122,1851]" captionTargetId="figure-365@37.[136,1459,1122,1851]" captionTargetPageId="37" captionText="FIGURE 30 | Distribution of Rhamphichthys heleios based on examined museum specimens. Red dot represents the type-locality." figureDoi="http://doi.org/10.5281/zenodo.13326957" httpUri="https://zenodo.org/record/13326957/files/figure.png" pageId="36" pageNumber="37">Fig. 30</figureCitation>
;
<tableCitation id="C6930333AB281E3A2132E283FA6EF930" box="[1371,1443,1720,1745]" captionStart="TABLE 1" captionStartId="9.[136,198,193,213]" captionText="TABLE 1 | Distributional data of species of Rhamphichthys in cis-Andean drainages of South America. Numbers corresponds to FEOW of Abell et al. (2008): Orinoco Llanos-307; Orinoco Guiana Shield-308; Orinoco Delta-309; Essequibo-310; Guianas-311; Negro-314; Amazonas Guiana Shield-315; Amazonas Lowlands-316; Mamoré-318; Guaporé-Itenez-319; Xingu-322; Amazonas estuary and coastal drainages-323; Tocantins- Araguaia-324; Parnaíba-325; Lower Uruguay-332; Chaco-342; Paraguay-343; Upper Paraná-344; and Lower Paraná-345. Asterisk (*) indicate the type-locality. Asterisks (**) indicates that Rhamphichthys hahni is allochthonous in the Upper Paraná (344) ecoregion." pageId="36">Tab. 1</tableCitation>
<tableCitation id="C6930333AB281E3A2132E283FA6EF930" box="[1371,1443,1720,1745]" captionStart="TABLE 1" captionStartId="9.[136,198,193,213]" captionText="TABLE 1 | Distributional data of species of Rhamphichthys in cis-Andean drainages of South America. Numbers corresponds to FEOW of Abell et al. (2008): Orinoco Llanos-307; Orinoco Guiana Shield-308; Orinoco Delta-309; Essequibo-310; Guianas-311; Negro-314; Amazonas Guiana Shield-315; Amazonas Lowlands-316; Mamoré-318; Guaporé-Itenez-319; Xingu-322; Amazonas estuary and coastal drainages-323; Tocantins- Araguaia-324; Parnaíba-325; Lower Uruguay-332; Chaco-342; Paraguay-343; Upper Paraná-344; and Lower Paraná-345. Asterisk (*) indicate the type-locality. Asterisks (**) indicates that Rhamphichthys hahni is allochthonous in the Upper Paraná (344) ecoregion." pageId="36" pageNumber="37">Tab. 1</tableCitation>
). It is also known from a single locality in the Guaporé River in
<collectingCountry id="F3067618AB281E3A20C8E2E5FB2BF917" box="[1185,1254,1758,1782]" name="Brazil" pageId="36">Brazil</collectingCountry>
<collectingCountry id="F3067618AB281E3A20C8E2E5FB2BF917" box="[1185,1254,1758,1782]" name="Brazil" pageId="36" pageNumber="37">Brazil</collectingCountry>
. It inhabits mostly lakes in the floodplain of large rivers.
</paragraph>
</subSubSection>
<caption id="DF6E6600AB291E3B2581E4FAFCF1FF1A" pageId="37" startId="37.[488,550,193,213]" targetType="table">
<paragraph id="8BAE3688AB291E3B2581E4FAFCF1FF1A" blockId="37.[488,1459,193,251]" pageId="37">
<emphasis id="B965EA9AAB291E3B2581E4FAFD9CFF34" bold="true" box="[488,593,193,213]" pageId="37">TABLE 7 |</emphasis>
<caption id="DF6E6600AB291E3B2581E4FAFCF1FF1A" pageId="37" pageNumber="38" startId="37.[488,550,193,213]" targetType="table">
<paragraph id="8BAE3688AB291E3B2581E4FAFCF1FF1A" blockId="37.[488,1459,193,251]" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B2581E4FAFD9CFF34" bold="true" box="[488,593,193,213]" pageId="37" pageNumber="38">TABLE 7 |</emphasis>
Morphometric and meristic data for
<taxonomicName id="4C114D0BAB291E3B27ACE4FAFB42FF34" box="[965,1167,193,213]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="37" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB291E3B27ACE4FAFB42FF34" bold="true" box="[965,1167,193,213]" italics="true" pageId="37">Rhamphichthys heleios</emphasis>
<taxonomicName id="4C114D0BAB291E3B27ACE4FAFB42FF34" box="[965,1167,193,213]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB291E3B27ACE4FAFB42FF34" bold="true" box="[965,1167,193,213]" italics="true" pageId="37" pageNumber="38">Rhamphichthys heleios</emphasis>
</taxonomicName>
. H = holotype, SD = standard deviation, n = number of specimens.
</paragraph>
</caption>
<paragraph id="8BAE3688AB291E3B27FCE520FA49FBF3" pageId="37">
<table id="F911C428AB29E1E12586E521FA47FBF3" box="[495,1418,282,1042]" colsContinueFrom="28.[495,1422,691,1394]" colsContinueIn="41.[495,1418,857,1653]" gridcols="6" gridrows="8" pageId="37">
<tr id="352134CAAB29E1E12586E521FA47FECD" box="[495,1418,282,300]" gridrow="0" pageId="37" rowspan-0="1">
<th id="76F05DB6AB29E1E127E2E521FC7DFECD" box="[907,944,282,300]" gridcol="1" gridrow="0" pageId="37">
<emphasis id="B965EA9AAB291E3B27FCE520FC69FECA" bold="true" box="[917,932,283,299]" pageId="37">H</emphasis>
<paragraph id="8BAE3688AB291E3B27FCE520FA49FBF3" pageId="37" pageNumber="38">
<table id="F911C428AB29E1E12586E521FA47FBF3" box="[495,1418,282,1042]" colsContinueFrom="28.[495,1422,691,1394]" colsContinueIn="41.[495,1418,857,1653]" gridcols="6" gridrows="8" pageId="37" pageNumber="38">
<tr id="352134CAAB29E1E12586E521FA47FECD" box="[495,1418,282,300]" gridrow="0" pageId="37" pageNumber="38" rowspan-0="1">
<th id="76F05DB6AB29E1E127E2E521FC7DFECD" box="[907,944,282,300]" gridcol="1" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B27FCE520FC69FECA" bold="true" box="[917,932,283,299]" pageId="37" pageNumber="38">H</emphasis>
</th>
<th id="76F05DB6AB29E1E12064E521FBD4FECD" box="[1037,1049,282,300]" gridcol="2" gridrow="0" pageId="37">
<emphasis id="B965EA9AAB291E3B2064E520FBD4FECA" bold="true" box="[1037,1049,283,299]" pageId="37">n</emphasis>
<th id="76F05DB6AB29E1E12064E521FBD4FECD" box="[1037,1049,282,300]" gridcol="2" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B2064E520FBD4FECA" bold="true" box="[1037,1049,283,299]" pageId="37" pageNumber="38">n</emphasis>
</th>
<th id="76F05DB6AB29E1E12008E521FB79FECD" box="[1121,1204,282,300]" gridcol="3" gridrow="0" pageId="37">
<emphasis id="B965EA9AAB291E3B2004E520FB6AFECA" bold="true" box="[1133,1191,283,299]" pageId="37">Range</emphasis>
<th id="76F05DB6AB29E1E12008E521FB79FECD" box="[1121,1204,282,300]" gridcol="3" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B2004E520FB6AFECA" bold="true" box="[1133,1191,283,299]" pageId="37" pageNumber="38">Range</emphasis>
</th>
<th id="76F05DB6AB29E1E1208EE521FAD6FECD" box="[1255,1307,282,300]" gridcol="4" gridrow="0" pageId="37">
<emphasis id="B965EA9AAB291E3B208EE520FAD6FECA" bold="true" box="[1255,1307,283,299]" pageId="37">Mean</emphasis>
<th id="76F05DB6AB29E1E1208EE521FAD6FECD" box="[1255,1307,282,300]" gridcol="4" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B208EE520FAD6FECA" bold="true" box="[1255,1307,283,299]" pageId="37" pageNumber="38">Mean</emphasis>
</th>
<th id="76F05DB6AB29E1E1210CE521FA47FECD" box="[1381,1418,282,300]" gridcol="5" gridrow="0" pageId="37">
<emphasis id="B965EA9AAB291E3B2102E521FA49FECD" bold="true" box="[1387,1412,282,300]" pageId="37">SD</emphasis>
<th id="76F05DB6AB29E1E1210CE521FA47FECD" box="[1381,1418,282,300]" gridcol="5" gridrow="0" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B2102E521FA49FECD" bold="true" box="[1387,1412,282,300]" pageId="37" pageNumber="38">SD</emphasis>
</th>
</tr>
<tr id="352134CAAB29E1E12586E578FA47FEB4" box="[495,1418,323,341]" gridrow="1" pageId="37">
<th id="76F05DB6AB29E1E12586E578FCCDFEB4" box="[495,768,323,341]" gridcol="0" gridrow="1" pageId="37">Length to end of anal fin (LEA)</th>
<td id="76F05DB6AB29E1E127E2E578FC7DFEB4" box="[907,944,323,341]" gridcol="1" gridrow="1" pageId="37">333</td>
<td id="76F05DB6AB29E1E12064E578FBD4FEB4" box="[1037,1049,323,341]" gridcol="2" gridrow="1" pageId="37">5</td>
<td id="76F05DB6AB29E1E12008E578FB79FEB4" box="[1121,1204,323,341]" gridcol="3" gridrow="1" pageId="37">320400</td>
<td id="76F05DB6AB29E1E1208EE578FAD6FEB4" box="[1255,1307,323,341]" gridcol="4" gridrow="1" pageId="37">355.3</td>
<td id="76F05DB6AB29E1E1210CE578FA47FEB4" box="[1381,1418,323,341]" gridcol="5" gridrow="1" pageId="37"></td>
<tr id="352134CAAB29E1E12586E578FA47FEB4" box="[495,1418,323,341]" gridrow="1" pageId="37" pageNumber="38">
<th id="76F05DB6AB29E1E12586E578FCCDFEB4" box="[495,768,323,341]" gridcol="0" gridrow="1" pageId="37" pageNumber="38">Length to end of anal fin (LEA)</th>
<td id="76F05DB6AB29E1E127E2E578FC7DFEB4" box="[907,944,323,341]" gridcol="1" gridrow="1" pageId="37" pageNumber="38">333</td>
<td id="76F05DB6AB29E1E12064E578FBD4FEB4" box="[1037,1049,323,341]" gridcol="2" gridrow="1" pageId="37" pageNumber="38">5</td>
<td id="76F05DB6AB29E1E12008E578FB79FEB4" box="[1121,1204,323,341]" gridcol="3" gridrow="1" pageId="37" pageNumber="38">320400</td>
<td id="76F05DB6AB29E1E1208EE578FAD6FEB4" box="[1255,1307,323,341]" gridcol="4" gridrow="1" pageId="37" pageNumber="38">355.3</td>
<td id="76F05DB6AB29E1E1210CE578FA47FEB4" box="[1381,1418,323,341]" gridcol="5" gridrow="1" pageId="37" pageNumber="38"></td>
</tr>
<tr id="352134CAAB29E1E12586E557FA47FE9F" box="[495,1418,364,382]" gridrow="2" pageId="37">
<th id="76F05DB6AB29E1E12586E557FA47FE9F" box="[495,1418,364,382]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="37">
<emphasis id="B965EA9AAB291E3B2586E556FD49FE9F" bold="true" box="[495,644,364,382]" pageId="37">Percents of LEA</emphasis>
<tr id="352134CAAB29E1E12586E557FA47FE9F" box="[495,1418,364,382]" gridrow="2" pageId="37" pageNumber="38">
<th id="76F05DB6AB29E1E12586E557FA47FE9F" box="[495,1418,364,382]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B2586E556FD49FE9F" bold="true" box="[495,644,364,382]" pageId="37" pageNumber="38">Percents of LEA</emphasis>
</th>
</tr>
<tr id="352134CAAB29E1E12586E5ADFA47FD97" box="[495,1418,406,630]" gridrow="3" pageId="37">
<th id="76F05DB6AB29E1E12586E5ADFCCDFD97" box="[495,768,406,630]" gridcol="0" gridrow="3" pageId="37">Anal-fin length Body depth Pectoral-fin length Head length Caudal filament length Caudal filament depth</th>
<td id="76F05DB6AB29E1E127E2E5ADFC7DFD97" box="[907,944,406,630]" gridcol="1" gridrow="3" pageId="37">90.1 7.8 4.8 12.0 16.9 1.7</td>
<td id="76F05DB6AB29E1E12064E5ADFBD4FD97" box="[1037,1049,406,630]" gridcol="2" gridrow="3" pageId="37">6 6 6 6 3 5</td>
<td id="76F05DB6AB29E1E12008E5ADFB79FD97" box="[1121,1204,406,630]" gridcol="3" gridrow="3" pageId="37">88.090.1 7.58.8 4.35.1 11.413.0 12.716.9 1.52.0</td>
<td id="76F05DB6AB29E1E1208EE5ADFAD6FD97" box="[1255,1307,406,630]" gridcol="4" gridrow="3" pageId="37">89.1 8.2 4.8 12.4 14.5 1.8</td>
<td id="76F05DB6AB29E1E1210CE5ADFA47FD97" box="[1381,1418,406,630]" gridcol="5" gridrow="3" pageId="37">0.8 0.5 0.2 0.6 2.2 0.1</td>
<tr id="352134CAAB29E1E12586E5ADFA47FD97" box="[495,1418,406,630]" gridrow="3" pageId="37" pageNumber="38">
<th id="76F05DB6AB29E1E12586E5ADFCCDFD97" box="[495,768,406,630]" gridcol="0" gridrow="3" pageId="37" pageNumber="38">Anal-fin length Body depth Pectoral-fin length Head length Caudal filament length Caudal filament depth</th>
<td id="76F05DB6AB29E1E127E2E5ADFC7DFD97" box="[907,944,406,630]" gridcol="1" gridrow="3" pageId="37" pageNumber="38">90.1 7.8 4.8 12.0 16.9 1.7</td>
<td id="76F05DB6AB29E1E12064E5ADFBD4FD97" box="[1037,1049,406,630]" gridcol="2" gridrow="3" pageId="37" pageNumber="38">6 6 6 6 3 5</td>
<td id="76F05DB6AB29E1E12008E5ADFB79FD97" box="[1121,1204,406,630]" gridcol="3" gridrow="3" pageId="37" pageNumber="38">88.090.1 7.58.8 4.35.1 11.413.0 12.716.9 1.52.0</td>
<td id="76F05DB6AB29E1E1208EE5ADFAD6FD97" box="[1255,1307,406,630]" gridcol="4" gridrow="3" pageId="37" pageNumber="38">89.1 8.2 4.8 12.4 14.5 1.8</td>
<td id="76F05DB6AB29E1E1210CE5ADFA47FD97" box="[1381,1418,406,630]" gridcol="5" gridrow="3" pageId="37" pageNumber="38">0.8 0.5 0.2 0.6 2.2 0.1</td>
</tr>
<tr id="352134CAAB29E1E12586E6B6FA47FD7E" box="[495,1418,653,671]" gridrow="4" pageId="37">
<th id="76F05DB6AB29E1E12586E6B6FA47FD7E" box="[495,1418,653,671]" colspan="6" colspanRight="5" gridcol="0" gridrow="4" pageId="37">
<emphasis id="B965EA9AAB291E3B2586E6B5FD09FD7F" bold="true" box="[495,708,653,671]" pageId="37">Percent of head length</emphasis>
<tr id="352134CAAB29E1E12586E6B6FA47FD7E" box="[495,1418,653,671]" gridrow="4" pageId="37" pageNumber="38">
<th id="76F05DB6AB29E1E12586E6B6FA47FD7E" box="[495,1418,653,671]" colspan="6" colspanRight="5" gridcol="0" gridrow="4" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B2586E6B5FD09FD7F" bold="true" box="[495,708,653,671]" pageId="37" pageNumber="38">Percent of head length</emphasis>
</th>
</tr>
<tr id="352134CAAB29E1E12586E68DFA47FC77" box="[495,1418,694,918]" gridrow="5" pageId="37">
<th id="76F05DB6AB29E1E12586E68DFCCDFC77" box="[495,768,694,918]" gridcol="0" gridrow="5" pageId="37">Interorbital distance Snout length Postorbital length Eye diameter Post. nares length Branchial opening</th>
<td id="76F05DB6AB29E1E127E2E68DFC7DFC77" box="[907,944,694,918]" gridcol="1" gridrow="5" pageId="37">12.3 52.8 45.9 5.4 19.0 22.4</td>
<td id="76F05DB6AB29E1E12064E68DFBD4FC77" box="[1037,1049,694,918]" gridcol="2" gridrow="5" pageId="37">6 6 6 6 6 6</td>
<td id="76F05DB6AB29E1E12008E68DFB79FC77" box="[1121,1204,694,918]" gridcol="3" gridrow="5" pageId="37">10.314.9 50.555.1 41.145.9 5.16.5 16.219.0 16.524.4</td>
<td id="76F05DB6AB29E1E1208EE68DFAD6FC77" box="[1255,1307,694,918]" gridcol="4" gridrow="5" pageId="37">12.2 52.5 44.1 5.8 17.9 21.8</td>
<td id="76F05DB6AB29E1E1210CE68DFA47FC77" box="[1381,1418,694,918]" gridcol="5" gridrow="5" pageId="37">1.7 1.5 1.7 0.5 0.9 2.7</td>
<tr id="352134CAAB29E1E12586E68DFA47FC77" box="[495,1418,694,918]" gridrow="5" pageId="37" pageNumber="38">
<th id="76F05DB6AB29E1E12586E68DFCCDFC77" box="[495,768,694,918]" gridcol="0" gridrow="5" pageId="37" pageNumber="38">Interorbital distance Snout length Postorbital length Eye diameter Post. nares length Branchial opening</th>
<td id="76F05DB6AB29E1E127E2E68DFC7DFC77" box="[907,944,694,918]" gridcol="1" gridrow="5" pageId="37" pageNumber="38">12.3 52.8 45.9 5.4 19.0 22.4</td>
<td id="76F05DB6AB29E1E12064E68DFBD4FC77" box="[1037,1049,694,918]" gridcol="2" gridrow="5" pageId="37" pageNumber="38">6 6 6 6 6 6</td>
<td id="76F05DB6AB29E1E12008E68DFB79FC77" box="[1121,1204,694,918]" gridcol="3" gridrow="5" pageId="37" pageNumber="38">10.314.9 50.555.1 41.145.9 5.16.5 16.219.0 16.524.4</td>
<td id="76F05DB6AB29E1E1208EE68DFAD6FC77" box="[1255,1307,694,918]" gridcol="4" gridrow="5" pageId="37" pageNumber="38">12.2 52.5 44.1 5.8 17.9 21.8</td>
<td id="76F05DB6AB29E1E1210CE68DFA47FC77" box="[1381,1418,694,918]" gridcol="5" gridrow="5" pageId="37" pageNumber="38">1.7 1.5 1.7 0.5 0.9 2.7</td>
</tr>
<tr id="352134CAAB29E1E12586E796FA47FC5E" box="[495,1418,941,959]" gridrow="6" pageId="37">
<th id="76F05DB6AB29E1E12586E796FA47FC5E" box="[495,1418,941,959]" colspan="6" colspanRight="5" gridcol="0" gridrow="6" pageId="37">
<emphasis id="B965EA9AAB291E3B2586E796FD8BFC5E" bold="true" box="[495,582,941,959]" pageId="37">Meristics</emphasis>
<tr id="352134CAAB29E1E12586E796FA47FC5E" box="[495,1418,941,959]" gridrow="6" pageId="37" pageNumber="38">
<th id="76F05DB6AB29E1E12586E796FA47FC5E" box="[495,1418,941,959]" colspan="6" colspanRight="5" gridcol="0" gridrow="6" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B2586E796FD8BFC5E" bold="true" box="[495,582,941,959]" pageId="37" pageNumber="38">Meristics</emphasis>
</th>
</tr>
<tr id="352134CAAB29E1E12586E7ECFA47FBF3" box="[495,1418,983,1042]" gridrow="7" pageId="37">
<th id="76F05DB6AB29E1E12586E7ECFCCDFBF3" box="[495,768,983,1042]" gridcol="0" gridrow="7" pageId="37">Anal-fin rays Pectoral-fin rays</th>
<td id="76F05DB6AB29E1E127E2E7ECFC7DFBF3" box="[907,944,983,1042]" gridcol="1" gridrow="7" pageId="37">342 17</td>
<td id="76F05DB6AB29E1E12064E7ECFBD4FBF3" box="[1037,1049,983,1042]" gridcol="2" gridrow="7" pageId="37">5 6</td>
<td id="76F05DB6AB29E1E12008E7ECFB79FBF3" box="[1121,1204,983,1042]" gridcol="3" gridrow="7" pageId="37">320345 1619</td>
<td id="76F05DB6AB29E1E1208EE7ECFAD6FBF3" box="[1255,1307,983,1042]" gridcol="4" gridrow="7" pageId="37">357.8 17.1</td>
<td id="76F05DB6AB29E1E1210CE7ECFA47FBF3" box="[1381,1418,983,1042]" gridcol="5" gridrow="7" pageId="37">15.0 0.7</td>
<tr id="352134CAAB29E1E12586E7ECFA47FBF3" box="[495,1418,983,1042]" gridrow="7" pageId="37" pageNumber="38">
<th id="76F05DB6AB29E1E12586E7ECFCCDFBF3" box="[495,768,983,1042]" gridcol="0" gridrow="7" pageId="37" pageNumber="38">Anal-fin rays Pectoral-fin rays</th>
<td id="76F05DB6AB29E1E127E2E7ECFC7DFBF3" box="[907,944,983,1042]" gridcol="1" gridrow="7" pageId="37" pageNumber="38">342 17</td>
<td id="76F05DB6AB29E1E12064E7ECFBD4FBF3" box="[1037,1049,983,1042]" gridcol="2" gridrow="7" pageId="37" pageNumber="38">5 6</td>
<td id="76F05DB6AB29E1E12008E7ECFB79FBF3" box="[1121,1204,983,1042]" gridcol="3" gridrow="7" pageId="37" pageNumber="38">320345 1619</td>
<td id="76F05DB6AB29E1E1208EE7ECFAD6FBF3" box="[1255,1307,983,1042]" gridcol="4" gridrow="7" pageId="37" pageNumber="38">357.8 17.1</td>
<td id="76F05DB6AB29E1E1210CE7ECFA47FBF3" box="[1381,1418,983,1042]" gridcol="5" gridrow="7" pageId="37" pageNumber="38">15.0 0.7</td>
</tr>
</table>
</paragraph>
<caption id="DF6E6600AB291E3B24E1E36AFAF9F884" box="[136,1332,1873,1893]" pageId="37" startId="37.[136,209,1873,1893]" targetBox="[136,1459,1122,1851]" targetPageId="37" targetType="figure">
<paragraph id="8BAE3688AB291E3B24E1E36AFAF9F884" blockId="37.[136,1332,1873,1893]" box="[136,1332,1873,1893]" pageId="37">
<emphasis id="B965EA9AAB291E3B24E1E36AFECCF884" bold="true" box="[136,257,1873,1893]" pageId="37">FIGURE 30 |</emphasis>
<caption id="DF6E6600AB291E3B24E1E36AFAF9F884" ID-DOI="http://doi.org/10.5281/zenodo.13326957" ID-Zenodo-Dep="13326957" box="[136,1332,1873,1893]" httpUri="https://zenodo.org/record/13326957/files/figure.png" pageId="37" pageNumber="38" startId="37.[136,209,1873,1893]" targetBox="[136,1459,1122,1851]" targetPageId="37" targetType="figure">
<paragraph id="8BAE3688AB291E3B24E1E36AFAF9F884" blockId="37.[136,1332,1873,1893]" box="[136,1332,1873,1893]" pageId="37" pageNumber="38">
<emphasis id="B965EA9AAB291E3B24E1E36AFECCF884" bold="true" box="[136,257,1873,1893]" pageId="37" pageNumber="38">FIGURE 30 |</emphasis>
Distribution of
<taxonomicName id="4C114D0BAB291E3B25F1E36AFD90F884" box="[408,605,1873,1893]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="37" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB291E3B25F1E36AFD90F884" bold="true" box="[408,605,1873,1893]" italics="true" pageId="37">Rhamphichthys heleios</emphasis>
<taxonomicName id="4C114D0BAB291E3B25F1E36AFD90F884" box="[408,605,1873,1893]" family="Rhamphichthyidae" genus="Rhamphichthys" kingdom="Animalia" order="Gymnotiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="heleios">
<emphasis id="B965EA9AAB291E3B25F1E36AFD90F884" bold="true" box="[408,605,1873,1893]" italics="true" pageId="37" pageNumber="38">Rhamphichthys heleios</emphasis>
</taxonomicName>
based on examined museum specimens. Red dot represents the type-locality.
</paragraph>
</caption>
<subSubSection id="C30B6503AB2A1E38266FE485FA79FCB0" pageId="38" type="materials_examined">
<paragraph id="8BAE3688AB2A1E38266FE485FA79FCB0" blockId="38.[488,1460,190,849]" pageId="38">
<emphasis id="B965EA9AAB2A1E38266FE485FC39FF37" bold="true" box="[518,1012,190,214]" pageId="38">Material examined. Amazonas lowlands:</emphasis>
<materialsCitation id="3B793CD5AB2A1E382794E485FD0BFEC0" collectingDate="1998-02-05" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Amazonas" municipality="Iranduba" pageId="38" specimenCode="INPA 42309" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB79EF3AB2A1E382794E485FB5DFF37" box="[1021,1168,190,214]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">INPA 42309</specimenCode>
<subSubSection id="C30B6503AB2A1E38266FE485FA79FCB0" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BAE3688AB2A1E38266FE485FA79FCB0" blockId="38.[488,1460,190,849]" pageId="38" pageNumber="39">
<emphasis id="B965EA9AAB2A1E38266FE485FC39FF37" bold="true" box="[518,1012,190,214]" pageId="38" pageNumber="39">Material examined. Amazonas lowlands:</emphasis>
<materialsCitation id="3B793CD5AB2A1E382794E485FD0BFEC0" collectingDate="1998-02-05" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Amazonas" municipality="Iranduba" pageId="38" pageNumber="39" specimenCode="INPA 42309" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB79EF3AB2A1E382794E485FB5DFF37" box="[1021,1168,190,214]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 42309</specimenCode>
,
<typeStatus id="54AA882AAB2A1E3820F4E485FACEFF37" box="[1181,1283,190,214]" pageId="38" type="holotype">holotype</typeStatus>
<typeStatus id="54AA882AAB2A1E3820F4E485FACEFF37" box="[1181,1283,190,214]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
,
<quantity id="4CE99B6DAB2A1E382179E485FAA2FF34" box="[1296,1391,190,214]" metricMagnitude="-1" metricUnit="m" metricValue="3.75" pageId="38" unit="mm" value="375.0">375 mm</quantity>
<quantity id="4CE99B6DAB2A1E382179E485FAA2FF34" box="[1296,1391,190,214]" metricMagnitude="-1" metricUnit="m" metricValue="3.75" pageId="38" pageNumber="39" unit="mm" value="375.0">375 mm</quantity>
LEA,
<collectingCountry id="F3067618AB2A1E382581E4D8FDE1FF1A" box="[488,556,227,251]" name="Brazil" pageId="38">Brazil</collectingCountry>
<collectingCountry id="F3067618AB2A1E382581E4D8FDE1FF1A" box="[488,556,227,251]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingCounty id="62CF4E04AB2A1E382652E4D8FD7CFF1A" box="[571,689,227,251]" pageId="38">Amazonas</collectingCounty>
<collectingCounty id="62CF4E04AB2A1E382652E4D8FD7CFF1A" box="[571,689,227,251]" pageId="38" pageNumber="39">Amazonas</collectingCounty>
,
<collectingMunicipality id="6BCAACF2AB2A1E3826A9E4D8FCEAFF1A" box="[704,807,227,251]" pageId="38">Iranduba</collectingMunicipality>
<collectingMunicipality id="6BCAACF2AB2A1E3826A9E4D8FCEAFF1A" box="[704,807,227,251]" pageId="38" pageNumber="39">Iranduba</collectingMunicipality>
, confluence of
<location id="8ECE6053AB2A1E3827B6E4D8FBB0FF1A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3827B6E4D8FBB0FF1A" box="[991,1149,227,251]" country="Brazil" county="Amazonas" municipality="Iranduba" name="rio Amazonas" pageId="38">rio Amazonas</location>
<location id="8ECE6053AB2A1E3827B6E4D8FBB0FF1A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3827B6E4D8FBB0FF1A" box="[991,1149,227,251]" country="Brazil" county="Amazonas" municipality="Iranduba" name="rio Amazonas" pageId="38" pageNumber="39">rio Amazonas</location>
and
<location id="8ECE6053AB2A1E3820D4E4D8FAECFF1A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820D4E4D8FAECFF1A" box="[1213,1313,227,251]" country="Brazil" county="Amazonas" municipality="Iranduba" name="Solimoes" pageId="38">Solimões</location>
<location id="8ECE6053AB2A1E3820D4E4D8FAECFF1A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820D4E4D8FAECFF1A" box="[1213,1313,227,251]" country="Brazil" county="Amazonas" municipality="Iranduba" name="Solimoes" pageId="38" pageNumber="39">Solimões</location>
at
<location id="8ECE6053AB2A1E382122E4D8FDF2FEC1" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382122E4D8FDF2FEC1" country="Brazil" county="Amazonas" municipality="Iranduba" name="Costa do Catalao" pageId="38">Costa do Catalão</location>
<location id="8ECE6053AB2A1E382122E4D8FDF2FEC1" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382122E4D8FDF2FEC1" country="Brazil" county="Amazonas" municipality="Iranduba" name="Costa do Catalao" pageId="38" pageNumber="39">Costa do Catalão</location>
,
<date id="FFAF1048AB2A1E382620E533FD0BFEC0" box="[585,710,264,289]" pageId="38" value="1998-02-05">
<collectingDate id="EFEBE9A0AB2A1E382620E533FD0BFEC0" box="[585,710,264,289]" pageId="38" value="1998-02-05">5 Feb 1998</collectingDate>
<date id="FFAF1048AB2A1E382620E533FD0BFEC0" box="[585,710,264,289]" pageId="38" pageNumber="39" value="1998-02-05">
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</date>
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, C. Cox Fernandes.
<materialsCitation id="3B793CD5AB2A1E3827CEE532FBB0FEA7" collectingDate="1998-11-19" collectionCode="INPA" country="Brazil" county="Amazonas" location="ilha da Marchantaria" municipality="Iranduba" pageId="38" specimenCode="INPA 17683, 1" specimenCount="1" typeStatus="paratype">
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<materialsCitation id="3B793CD5AB2A1E3827CEE532FBB0FEA7" collectingDate="1998-11-19" collectionCode="INPA" country="Brazil" county="Amazonas" location="ilha da Marchantaria" municipality="Iranduba" pageId="38" pageNumber="39" specimenCode="INPA 17683, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E3827CEE532FBFBFEC1" box="[935,1078,264,289]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 17683</specimenCode>
,
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<specimenCode id="DBB79EF3AB2A1E382028E532FB82FEC0" box="[1089,1103,265,289]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">1</specimenCode>
,
<typeStatus id="54AA882AAB2A1E382032E532FB70FEC0" box="[1115,1213,265,289]" pageId="38" type="paratype">paratype</typeStatus>
<typeStatus id="54AA882AAB2A1E382032E532FB70FEC0" box="[1115,1213,265,289]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
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<quantity id="4CE99B6DAB2A1E3820A1E532FAE9FEC1" box="[1224,1316,265,289]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="38" unit="mm" value="270.0">270 mm</quantity>
<quantity id="4CE99B6DAB2A1E3820A1E532FAE9FEC1" box="[1224,1316,265,289]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="38" pageNumber="39" unit="mm" value="270.0">270 mm</quantity>
LEA,
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<collectingCountry id="F3067618AB2A1E382102E533FA62FEC1" box="[1387,1455,264,288]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
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<collectingCounty id="62CF4E04AB2A1E382581E515FD93FEA7" box="[488,606,302,326]" pageId="38">Amazonas</collectingCounty>
<collectingCounty id="62CF4E04AB2A1E382581E515FD93FEA7" box="[488,606,302,326]" pageId="38" pageNumber="39">Amazonas</collectingCounty>
,
<collectingMunicipality id="6BCAACF2AB2A1E382605E515FD1EFEA7" box="[620,723,302,326]" pageId="38">Iranduba</collectingMunicipality>
<collectingMunicipality id="6BCAACF2AB2A1E382605E515FD1EFEA7" box="[620,723,302,326]" pageId="38" pageNumber="39">Iranduba</collectingMunicipality>
,
<location id="8ECE6053AB2A1E38268BE515FC1CFEA7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38268BE515FC1CFEA7" box="[738,977,302,326]" country="Brazil" county="Amazonas" municipality="Iranduba" name="ilha da Marchantaria" pageId="38">ilha da Marchantaria</location>
<location id="8ECE6053AB2A1E38268BE515FC1CFEA7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38268BE515FC1CFEA7" box="[738,977,302,326]" country="Brazil" county="Amazonas" municipality="Iranduba" name="ilha da Marchantaria" pageId="38" pageNumber="39">ilha da Marchantaria</location>
,
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<collectingDate id="EFEBE9A0AB2A1E382789E515FBB0FEA7" box="[992,1149,302,326]" pageId="38" value="1998-11-19">19 Nov 1998</collectingDate>
<date id="FFAF1048AB2A1E382789E515FBB0FEA7" box="[992,1149,302,326]" pageId="38" pageNumber="39" value="1998-11-19">
<collectingDate id="EFEBE9A0AB2A1E382789E515FBB0FEA7" box="[992,1149,302,326]" pageId="38" pageNumber="39" value="1998-11-19">19 Nov 1998</collectingDate>
</date>
</materialsCitation>
, C. Cox Fernandes.
<materialsCitation id="3B793CD5AB2A1E382119E515FD24FE70" collectingDate="2000-02-24" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Japura" municipality="Alvaraes" pageId="38" specimenCode="INPA 18321, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382119E515FDE2FE8A" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">INPA 18321</specimenCode>
<materialsCitation id="3B793CD5AB2A1E382119E515FD24FE70" collectingDate="2000-02-24" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Japura" municipality="Alvaraes" pageId="38" pageNumber="39" specimenCode="INPA 18321, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382119E515FDE2FE8A" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 18321</specimenCode>
,
<specimenCode id="DBB79EF3AB2A1E382650E568FD8AFE8A" box="[569,583,339,363]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">1</specimenCode>
<specimenCode id="DBB79EF3AB2A1E382650E568FD8AFE8A" box="[569,583,339,363]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">1</specimenCode>
,
<typeStatus id="54AA882AAB2A1E38263AE56FFD78FE8D" box="[595,693,340,364]" pageId="38" type="paratype">paratype</typeStatus>
<typeStatus id="54AA882AAB2A1E38263AE56FFD78FE8D" box="[595,693,340,364]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
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<quantity id="4CE99B6DAB2A1E3826A9E568FCD1FE8A" box="[704,796,339,363]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="38" unit="mm" value="370.0">370 mm</quantity>
<quantity id="4CE99B6DAB2A1E3826A9E568FCD1FE8A" box="[704,796,339,363]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="38" pageNumber="39" unit="mm" value="370.0">370 mm</quantity>
LEA,
<collectingCountry id="F3067618AB2A1E38270AE568FC6AFE8A" box="[867,935,339,363]" name="Brazil" pageId="38">Brazil</collectingCountry>
<collectingCountry id="F3067618AB2A1E38270AE568FC6AFE8A" box="[867,935,339,363]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingCounty id="62CF4E04AB2A1E3827DBE568FBE5FE8A" box="[946,1064,339,363]" pageId="38">Amazonas</collectingCounty>
<collectingCounty id="62CF4E04AB2A1E3827DBE568FBE5FE8A" box="[946,1064,339,363]" pageId="38" pageNumber="39">Amazonas</collectingCounty>
,
<collectingMunicipality id="6BCAACF2AB2A1E38205BE568FB5FFE8A" box="[1074,1170,339,363]" pageId="38">Alvarães</collectingMunicipality>
<collectingMunicipality id="6BCAACF2AB2A1E38205BE568FB5FFE8A" box="[1074,1170,339,363]" pageId="38" pageNumber="39">Alvarães</collectingMunicipality>
,
<location id="8ECE6053AB2A1E3820F5E568FAC5FE8A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820F5E568FAC5FE8A" box="[1180,1288,339,363]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="rio Japura" pageId="38">rio Japurá</location>
<location id="8ECE6053AB2A1E3820F5E568FAC5FE8A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820F5E568FAC5FE8A" box="[1180,1288,339,363]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="rio Japura" pageId="38" pageNumber="39">rio Japurá</location>
at comunidade
<location id="8ECE6053AB2A1E382581E543FD82FE71" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382581E543FD82FE71" box="[488,591,376,400]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Caborini" pageId="38">Caborini</location>
<location id="8ECE6053AB2A1E382581E543FD82FE71" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382581E543FD82FE71" box="[488,591,376,400]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Caborini" pageId="38" pageNumber="39">Caborini</location>
,
<date id="FFAF1048AB2A1E382632E542FD24FE70" box="[603,745,376,401]" pageId="38" value="2000-02-24">
<collectingDate id="EFEBE9A0AB2A1E382632E542FD24FE70" box="[603,745,376,401]" pageId="38" value="2000-02-24">24 Feb 2000</collectingDate>
<date id="FFAF1048AB2A1E382632E542FD24FE70" box="[603,745,376,401]" pageId="38" pageNumber="39" value="2000-02-24">
<collectingDate id="EFEBE9A0AB2A1E382632E542FD24FE70" box="[603,745,376,401]" pageId="38" pageNumber="39" value="2000-02-24">24 Feb 2000</collectingDate>
</date>
</materialsCitation>
, W. G.
<materialsCitation id="3B793CD5AB2A1E38272CE542FC12FE70" box="[837,991,377,401]" collectionCode="R" country="Brazil" location="Crampton" pageId="38" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED00AE4DAB2A1E38272CE542FC96FE71" box="[837,859,377,400]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38">R</collectionCode>
<materialsCitation id="3B793CD5AB2A1E38272CE542FC12FE70" box="[837,991,377,401]" collectionCode="R" country="Brazil" location="Crampton" pageId="38" pageNumber="39" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED00AE4DAB2A1E38272CE542FC96FE71" box="[837,859,377,400]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="39">R</collectionCode>
.
<location id="8ECE6053AB2A1E38270EE542FC12FE70" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38270EE542FC12FE70" box="[871,991,377,401]" country="Brazil" name="Crampton" pageId="38">Crampton</location>
<location id="8ECE6053AB2A1E38270EE542FC12FE70" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38270EE542FC12FE70" box="[871,991,377,401]" country="Brazil" name="Crampton" pageId="38" pageNumber="39">Crampton</location>
</materialsCitation>
.
<materialsCitation id="3B793CD5AB2A1E382783E542FDA6FE3A" collectingDate="1999-02-03" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Japura" municipality="Alvaraes" pageId="38" specimenCode="INPA 18323, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382783E542FBB6FE70" box="[1002,1147,377,401]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">INPA 18323</specimenCode>
<materialsCitation id="3B793CD5AB2A1E382783E542FDA6FE3A" collectingDate="1999-02-03" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Japura" municipality="Alvaraes" pageId="38" pageNumber="39" specimenCode="INPA 18323, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382783E542FBB6FE70" box="[1002,1147,377,401]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 18323</specimenCode>
,
<specimenCode id="DBB79EF3AB2A1E3820EEE542FB58FE70" box="[1159,1173,377,401]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">1</specimenCode>
<specimenCode id="DBB79EF3AB2A1E3820EEE542FB58FE70" box="[1159,1173,377,401]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">1</specimenCode>
,
<typeStatus id="54AA882AAB2A1E3820CDE542FACBFE70" box="[1188,1286,377,401]" pageId="38" type="paratype">paratype</typeStatus>
<typeStatus id="54AA882AAB2A1E3820CDE542FACBFE70" box="[1188,1286,377,401]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
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<quantity id="4CE99B6DAB2A1E38217BE543FABDFE71" box="[1298,1392,376,400]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" pageId="38" unit="mm" value="530.0">530 mm</quantity>
<quantity id="4CE99B6DAB2A1E38217BE543FABDFE71" box="[1298,1392,376,400]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" pageId="38" pageNumber="39" unit="mm" value="530.0">530 mm</quantity>
LEA,
<collectingCountry id="F3067618AB2A1E382581E5A5FDE1FE57" box="[488,556,414,438]" name="Brazil" pageId="38">Brazil</collectingCountry>
<collectingCountry id="F3067618AB2A1E382581E5A5FDE1FE57" box="[488,556,414,438]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingCounty id="62CF4E04AB2A1E38265FE5A5FD61FE57" box="[566,684,414,438]" pageId="38">Amazonas</collectingCounty>
<collectingCounty id="62CF4E04AB2A1E38265FE5A5FD61FE57" box="[566,684,414,438]" pageId="38" pageNumber="39">Amazonas</collectingCounty>
,
<collectingMunicipality id="6BCAACF2AB2A1E3826DFE5A5FCDBFE57" box="[694,790,414,438]" pageId="38">Alvarães</collectingMunicipality>
<collectingMunicipality id="6BCAACF2AB2A1E3826DFE5A5FCDBFE57" box="[694,790,414,438]" pageId="38" pageNumber="39">Alvarães</collectingMunicipality>
,
<location id="8ECE6053AB2A1E382748E5A5FC43FE57" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382748E5A5FC43FE57" box="[801,910,414,438]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="rio Japura" pageId="38">rio Japurá</location>
<location id="8ECE6053AB2A1E382748E5A5FC43FE57" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382748E5A5FC43FE57" box="[801,910,414,438]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="rio Japura" pageId="38" pageNumber="39">rio Japurá</location>
mouth of
<location id="8ECE6053AB2A1E38206BE5A5FB71FE57" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38206BE5A5FB71FE57" box="[1026,1212,414,438]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Lago Caxinguba" pageId="38">Lago Caxinguba</location>
<location id="8ECE6053AB2A1E38206BE5A5FB71FE57" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38206BE5A5FB71FE57" box="[1026,1212,414,438]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Lago Caxinguba" pageId="38" pageNumber="39">Lago Caxinguba</location>
at
<location id="8ECE6053AB2A1E3820B4E5A5FA62FE57" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820B4E5A5FA62FE57" box="[1245,1455,414,438]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Reserva Mamiraua" pageId="38">Reserva Mamirauá</location>
<location id="8ECE6053AB2A1E3820B4E5A5FA62FE57" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820B4E5A5FA62FE57" box="[1245,1455,414,438]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Reserva Mamiraua" pageId="38" pageNumber="39">Reserva Mamirauá</location>
,
<date id="FFAF1048AB2A1E382581E5F8FDA6FE3A" box="[488,619,451,475]" pageId="38" value="1999-02-03">
<collectingDate id="EFEBE9A0AB2A1E382581E5F8FDA6FE3A" box="[488,619,451,475]" pageId="38" value="1999-02-03">3 Feb 1999</collectingDate>
<date id="FFAF1048AB2A1E382581E5F8FDA6FE3A" box="[488,619,451,475]" pageId="38" pageNumber="39" value="1999-02-03">
<collectingDate id="EFEBE9A0AB2A1E382581E5F8FDA6FE3A" box="[488,619,451,475]" pageId="38" pageNumber="39" value="1999-02-03">3 Feb 1999</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B793CD5AB2A1E382611E5F8FD71FDE1" collectionCode="INPA" collectorName="Maiana, W" country="Brazil" location="rio Japura" municipality="Amazonas" pageId="38" specimenCode="INPA 18331, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382611E5F8FCC6FE3A" box="[632,779,451,475]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">INPA 18331</specimenCode>
<materialsCitation id="3B793CD5AB2A1E382611E5F8FD71FDE1" collectionCode="INPA" collectorName="Maiana, W" country="Brazil" location="rio Japura" municipality="Amazonas" pageId="38" pageNumber="39" specimenCode="INPA 18331, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382611E5F8FCC6FE3A" box="[632,779,451,475]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 18331</specimenCode>
,
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<specimenCode id="DBB79EF3AB2A1E382771E5F8FCEBFE3A" box="[792,806,451,475]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">1</specimenCode>
,
<typeStatus id="54AA882AAB2A1E38275FE5FFFC55FE3D" box="[822,920,452,476]" pageId="38" type="paratype">paratype</typeStatus>
<typeStatus id="54AA882AAB2A1E38275FE5FFFC55FE3D" box="[822,920,452,476]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
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<quantity id="4CE99B6DAB2A1E3827CFE5F8FBC8FE3A" box="[934,1029,451,475]" metricMagnitude="-1" metricUnit="m" metricValue="3.3" pageId="38" unit="mm" value="330.0">330 mm</quantity>
<quantity id="4CE99B6DAB2A1E3827CFE5F8FBC8FE3A" box="[934,1029,451,475]" metricMagnitude="-1" metricUnit="m" metricValue="3.3" pageId="38" pageNumber="39" unit="mm" value="330.0">330 mm</quantity>
LEA,
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<collectingCountry id="F3067618AB2A1E38203BE5F8FB5BFE3A" box="[1106,1174,451,475]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingMunicipality id="6BCAACF2AB2A1E3820CAE5F8FAD4FE3A" box="[1187,1305,451,475]" pageId="38">Amazonas</collectingMunicipality>
<collectingMunicipality id="6BCAACF2AB2A1E3820CAE5F8FAD4FE3A" box="[1187,1305,451,475]" pageId="38" pageNumber="39">Amazonas</collectingMunicipality>
,
<location id="8ECE6053AB2A1E38214FE5F8FA5BFE3A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38214FE5F8FA5BFE3A" box="[1318,1430,451,475]" country="Brazil" municipality="Amazonas" name="rio Japura" pageId="38">rio Japurá</location>
<location id="8ECE6053AB2A1E38214FE5F8FA5BFE3A" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E38214FE5F8FA5BFE3A" box="[1318,1430,451,475]" country="Brazil" municipality="Amazonas" name="rio Japura" pageId="38" pageNumber="39">rio Japurá</location>
at paraná
<collectorName id="26E4535EAB2A1E382653E5D3FD71FDE1" box="[570,700,488,512]" pageId="38">Maiana, W</collectorName>
<collectorName id="26E4535EAB2A1E382653E5D3FD71FDE1" box="[570,700,488,512]" pageId="38" pageNumber="39">Maiana, W</collectorName>
</materialsCitation>
.
<materialsCitation id="3B793CD5AB2A1E3826A0E5D2FC45FDE0" box="[713,904,489,513]" collectionCode="R" country="Brazil" location="Crampton" pageId="38" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED00AE4DAB2A1E3826A0E5D2FD12FDE1" box="[713,735,489,512]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38">R</collectionCode>
<materialsCitation id="3B793CD5AB2A1E3826A0E5D2FC45FDE0" box="[713,904,489,513]" collectionCode="R" country="Brazil" location="Crampton" pageId="38" pageNumber="39" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED00AE4DAB2A1E3826A0E5D2FD12FDE1" box="[713,735,489,512]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="39">R</collectionCode>
. G.
<location id="8ECE6053AB2A1E382779E5D2FC45FDE0" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382779E5D2FC45FDE0" box="[784,904,489,513]" country="Brazil" name="Crampton" pageId="38">Crampton</location>
<location id="8ECE6053AB2A1E382779E5D2FC45FDE0" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382779E5D2FC45FDE0" box="[784,904,489,513]" country="Brazil" name="Crampton" pageId="38" pageNumber="39">Crampton</location>
</materialsCitation>
.
<materialsCitation id="3B793CD5AB2A1E3827FDE5D2FA4FFDC7" collectingDate="2000-07-09" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Solimoes" municipality="Alvaraes" pageId="38" specimenCode="INPA 18332, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E3827FDE5D2FBEBFDE0" box="[916,1062,489,513]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">INPA 18332</specimenCode>
<materialsCitation id="3B793CD5AB2A1E3827FDE5D2FA4FFDC7" collectingDate="2000-07-09" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Solimoes" municipality="Alvaraes" pageId="38" pageNumber="39" specimenCode="INPA 18332, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E3827FDE5D2FBEBFDE0" box="[916,1062,489,513]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 18332</specimenCode>
,
<specimenCode id="DBB79EF3AB2A1E38205AE5D2FB8CFDE0" box="[1075,1089,489,513]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">1</specimenCode>
<specimenCode id="DBB79EF3AB2A1E38205AE5D2FB8CFDE0" box="[1075,1089,489,513]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">1</specimenCode>
,
<typeStatus id="54AA882AAB2A1E382039E5D2FB7FFDE0" box="[1104,1202,489,513]" pageId="38" type="paratype">paratype</typeStatus>
<typeStatus id="54AA882AAB2A1E382039E5D2FB7FFDE0" box="[1104,1202,489,513]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
,
<quantity id="4CE99B6DAB2A1E3820A9E5D3FAD3FDE1" box="[1216,1310,488,512]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="38" unit="mm" value="250.0">250 mm</quantity>
<quantity id="4CE99B6DAB2A1E3820A9E5D3FAD3FDE1" box="[1216,1310,488,512]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="mm" value="250.0">250 mm</quantity>
LEA,
<collectingCountry id="F3067618AB2A1E382102E5D3FA62FDE1" box="[1387,1455,488,512]" name="Brazil" pageId="38">Brazil</collectingCountry>
<collectingCountry id="F3067618AB2A1E382102E5D3FA62FDE1" box="[1387,1455,488,512]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingCounty id="62CF4E04AB2A1E382581E635FD93FDC7" box="[488,606,526,550]" pageId="38">Amazonas</collectingCounty>
<collectingCounty id="62CF4E04AB2A1E382581E635FD93FDC7" box="[488,606,526,550]" pageId="38" pageNumber="39">Amazonas</collectingCounty>
,
<collectingMunicipality id="6BCAACF2AB2A1E382600E635FD04FDC7" box="[617,713,526,550]" pageId="38">Alvarães</collectingMunicipality>
<collectingMunicipality id="6BCAACF2AB2A1E382600E635FD04FDC7" box="[617,713,526,550]" pageId="38" pageNumber="39">Alvarães</collectingMunicipality>
,
<location id="8ECE6053AB2A1E3826BFE635FCACFDC7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3826BFE635FCACFDC7" box="[726,865,526,550]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="rio Solimoes" pageId="38">rio Solimões</location>
<location id="8ECE6053AB2A1E3826BFE635FCACFDC7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3826BFE635FCACFDC7" box="[726,865,526,550]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="rio Solimoes" pageId="38" pageNumber="39">rio Solimões</location>
at
<location id="8ECE6053AB2A1E3827EFE635FBDCFDC7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3827EFE635FBDCFDC7" box="[902,1041,526,550]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Vila Alencar" pageId="38">Vila Alencar</location>
<location id="8ECE6053AB2A1E3827EFE635FBDCFDC7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3827EFE635FBDCFDC7" box="[902,1041,526,550]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Vila Alencar" pageId="38" pageNumber="39">Vila Alencar</location>
at reserva
<location id="8ECE6053AB2A1E3820E5E635FACCFDC7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820E5E635FACCFDC7" box="[1164,1281,526,550]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Mamiraua" pageId="38">Mamirauá</location>
<location id="8ECE6053AB2A1E3820E5E635FACCFDC7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3820E5E635FACCFDC7" box="[1164,1281,526,550]" country="Brazil" county="Amazonas" municipality="Alvaraes" name="Mamiraua" pageId="38" pageNumber="39">Mamirauá</location>
,
<date id="FFAF1048AB2A1E382164E635FA4FFDC7" box="[1293,1410,526,550]" pageId="38" value="2000-07-09">
<collectingDate id="EFEBE9A0AB2A1E382164E635FA4FFDC7" box="[1293,1410,526,550]" pageId="38" value="2000-07-09">9 Jul 2000</collectingDate>
<date id="FFAF1048AB2A1E382164E635FA4FFDC7" box="[1293,1410,526,550]" pageId="38" pageNumber="39" value="2000-07-09">
<collectingDate id="EFEBE9A0AB2A1E382164E635FA4FFDC7" box="[1293,1410,526,550]" pageId="38" pageNumber="39" value="2000-07-09">9 Jul 2000</collectingDate>
</date>
</materialsCitation>
, W. G.
<materialsCitation id="3B793CD5AB2A1E382664E60FFD64FDAA" box="[525,681,563,587]" collectionCode="R" country="Brazil" location="Crampton" pageId="38" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED00AE4DAB2A1E382664E60FFDEEFDAA" box="[525,547,564,587]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38">R</collectionCode>
<materialsCitation id="3B793CD5AB2A1E382664E60FFD64FDAA" box="[525,681,563,587]" collectionCode="R" country="Brazil" location="Crampton" pageId="38" pageNumber="39" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED00AE4DAB2A1E382664E60FFDEEFDAA" box="[525,547,564,587]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="39">R</collectionCode>
.
<location id="8ECE6053AB2A1E382658E608FD64FDAA" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382658E608FD64FDAA" box="[561,681,563,587]" country="Brazil" name="Crampton" pageId="38">Crampton</location>
<location id="8ECE6053AB2A1E382658E608FD64FDAA" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382658E608FD64FDAA" box="[561,681,563,587]" country="Brazil" name="Crampton" pageId="38" pageNumber="39">Crampton</location>
</materialsCitation>
.
<materialsCitation id="3B793CD5AB2A1E3826DFE608FBD2FD90" collectingDate="1998-03-12" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Solimoes" municipality="Manaus" pageId="38" specimenCode="INPA 27611, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E3826DFE608FC87FDAA" box="[694,842,563,587]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">INPA 27611</specimenCode>
<materialsCitation id="3B793CD5AB2A1E3826DFE608FBD2FD90" collectingDate="1998-03-12" collectionCode="INPA" country="Brazil" county="Amazonas" location="rio Solimoes" municipality="Manaus" pageId="38" pageNumber="39" specimenCode="INPA 27611, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E3826DFE608FC87FDAA" box="[694,842,563,587]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 27611</specimenCode>
,
<specimenCode id="DBB79EF3AB2A1E382731E608FCABFDAA" box="[856,870,563,587]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">1</specimenCode>
<specimenCode id="DBB79EF3AB2A1E382731E608FCABFDAA" box="[856,870,563,587]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">1</specimenCode>
,
<typeStatus id="54AA882AAB2A1E38271FE60FFC15FDAD" box="[886,984,564,588]" pageId="38" type="paratype">paratype</typeStatus>
<typeStatus id="54AA882AAB2A1E38271FE60FFC15FDAD" box="[886,984,564,588]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
,
<quantity id="4CE99B6DAB2A1E38278EE608FB8AFDAA" box="[999,1095,563,587]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="38" unit="mm" value="235.0">235 mm</quantity>
<quantity id="4CE99B6DAB2A1E38278EE608FB8AFDAA" box="[999,1095,563,587]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="38" pageNumber="39" unit="mm" value="235.0">235 mm</quantity>
SL,
<collectingCountry id="F3067618AB2A1E382016E608FB0EFDAA" box="[1151,1219,563,587]" name="Brazil" pageId="38">Brazil</collectingCountry>
<collectingCountry id="F3067618AB2A1E382016E608FB0EFDAA" box="[1151,1219,563,587]" name="Brazil" pageId="38" pageNumber="39">Brazil</collectingCountry>
,
<collectingCounty id="62CF4E04AB2A1E3820B8E608FA8AFDAA" box="[1233,1351,563,587]" pageId="38">Amazonas</collectingCounty>
<collectingCounty id="62CF4E04AB2A1E3820B8E608FA8AFDAA" box="[1233,1351,563,587]" pageId="38" pageNumber="39">Amazonas</collectingCounty>
,
<collectingMunicipality id="6BCAACF2AB2A1E38213CE60FFA62FDAA" box="[1365,1455,564,587]" pageId="38">Manaus</collectingMunicipality>
<collectingMunicipality id="6BCAACF2AB2A1E38213CE60FFA62FDAA" box="[1365,1455,564,587]" pageId="38" pageNumber="39">Manaus</collectingMunicipality>
,
<location id="8ECE6053AB2A1E382581E663FDB9FD91" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382581E663FDB9FD91" box="[488,628,600,624]" country="Brazil" county="Amazonas" municipality="Manaus" name="rio Solimoes" pageId="38">rio Solimões</location>
<location id="8ECE6053AB2A1E382581E663FDB9FD91" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382581E663FDB9FD91" box="[488,628,600,624]" country="Brazil" county="Amazonas" municipality="Manaus" name="rio Solimoes" pageId="38" pageNumber="39">rio Solimões</location>
at paraná do
<location id="8ECE6053AB2A1E382778E663FCB0FD91" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382778E663FCB0FD91" box="[785,893,600,624]" country="Brazil" county="Amazonas" municipality="Manaus" name="Xiborena" pageId="38">Xiborena</location>
<location id="8ECE6053AB2A1E382778E663FCB0FD91" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E382778E663FCB0FD91" box="[785,893,600,624]" country="Brazil" county="Amazonas" municipality="Manaus" name="Xiborena" pageId="38" pageNumber="39">Xiborena</location>
,
<date id="FFAF1048AB2A1E3827E2E662FBD2FD90" box="[907,1055,601,625]" pageId="38" value="1998-03-12">
<collectingDate id="EFEBE9A0AB2A1E3827E2E662FBD2FD90" box="[907,1055,601,625]" pageId="38" value="1998-03-12">12 Mar 1998</collectingDate>
<date id="FFAF1048AB2A1E3827E2E662FBD2FD90" box="[907,1055,601,625]" pageId="38" pageNumber="39" value="1998-03-12">
<collectingDate id="EFEBE9A0AB2A1E3827E2E662FBD2FD90" box="[907,1055,601,625]" pageId="38" pageNumber="39" value="1998-03-12">12 Mar 1998</collectingDate>
</date>
</materialsCitation>
, C. Cox Fernandes.
<materialsCitation id="3B793CD5AB2A1E382162E662FCC2FD77" collectingDate="1998-02-05" collectionCode="ANSP" country="Brazil" county="Amazonas" location="rio Amazonas" municipality="Iranduba" pageId="38" specimenCode="ANSP 195254, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382162E662FA62FD91" box="[1291,1455,600,625]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="38">ANSP 195254</specimenCode>
<materialsCitation id="3B793CD5AB2A1E382162E662FCC2FD77" collectingDate="1998-02-05" collectionCode="ANSP" country="Brazil" county="Amazonas" location="rio Amazonas" municipality="Iranduba" pageId="38" pageNumber="39" specimenCode="ANSP 195254, 1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382162E662FA62FD91" box="[1291,1455,600,625]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="38" pageNumber="39">ANSP 195254</specimenCode>
,
<specimenCode id="DBB79EF3AB2A1E382581E645FE3BFD77" box="[488,502,638,662]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="38">1</specimenCode>
<specimenCode id="DBB79EF3AB2A1E382581E645FE3BFD77" box="[488,502,638,662]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="38" pageNumber="39">1</specimenCode>
, collected with holotype
</materialsCitation>
;
<materialsCitation id="3B793CD5AB2A1E382773E645FA7DFD77" box="[794,1456,638,662]" collectingDate="1998-02-05" collectionCode="MZUSP" country="Brazil" county="Amazonas" location="rio Amazonas" municipality="Iranduba" pageId="38" specimenCode="MZUSP 115347" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382773E645FC1BFD77" box="[794,982,638,662]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="38">MZUSP 115347</specimenCode>
<materialsCitation id="3B793CD5AB2A1E382773E645FA7DFD77" box="[794,1456,638,662]" collectingDate="1998-02-05" collectionCode="MZUSP" country="Brazil" county="Amazonas" location="rio Amazonas" municipality="Iranduba" pageId="38" pageNumber="39" specimenCode="MZUSP 115347" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382773E645FC1BFD77" box="[794,982,638,662]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="38" pageNumber="39">MZUSP 115347</specimenCode>
,
<quantity id="4CE99B6DAB2A1E382788E645FB97FD74" box="[993,1114,638,662]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="38" unit="mm" value="1.45">1, 450 mm</quantity>
<quantity id="4CE99B6DAB2A1E382788E645FB97FD74" box="[993,1114,638,662]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="38" pageNumber="39" unit="mm" value="1.45">1, 450 mm</quantity>
LEA, collected with holotype
</materialsCitation>
.
<materialsCitation id="3B793CD5AB2A1E382581E698FC38FD01" collectionCode="INPA" country="Brazil" elevation="352" pageId="38" specimenCode="INPA 42308, 4" specimenCount="4" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382581E698FDB7FD5A" box="[488,634,675,699]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">INPA 42308</specimenCode>
<materialsCitation id="3B793CD5AB2A1E382581E698FC38FD01" collectionCode="INPA" country="Brazil" elevation="352" pageId="38" pageNumber="39" specimenCode="INPA 42308, 4" specimenCount="4" typeStatus="paratype">
<specimenCode id="DBB79EF3AB2A1E382581E698FDB7FD5A" box="[488,634,675,699]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">INPA 42308</specimenCode>
,
<specimenCode id="DBB79EF3AB2A1E3826EEE698FD58FD5A" box="[647,661,675,699]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38">4</specimenCode>
<specimenCode id="DBB79EF3AB2A1E3826EEE698FD58FD5A" box="[647,661,675,699]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="38" pageNumber="39">4</specimenCode>
(
<specimenCount id="9D17FD01AB2A1E3826CFE698FCE2FD5A" box="[678,815,675,699]" count="4" pageId="38" type="generic">4 specimens</specimenCount>
<specimenCount id="9D17FD01AB2A1E3826CFE698FCE2FD5A" box="[678,815,675,699]" count="4" pageId="38" pageNumber="39" type="generic">4 specimens</specimenCount>
measured in
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<tableCitation id="C6930333AB2A1E3827A3E698FBDAFD5A" box="[970,1047,675,699]" captionStart="TABLE 7" captionStartId="37.[488,550,193,213]" captionText="TABLE 7 | Morphometric and meristic data for Rhamphichthys heleios. H = holotype, SD = standard deviation, n = number of specimens." pageId="38" pageNumber="39">
Tab.
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, 333400 mm LEA, 1 cs),
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LEA, collected with the
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LEA,
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,
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<collectingMunicipality id="6BCAACF2AB2A1E382604E6D5FD69FCE7" box="[621,676,750,774]" pageId="38" pageNumber="39">Tefé</collectingMunicipality>
,
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<location id="8ECE6053AB2A1E3826DDE6D5FCA6FCE7" LSID="urn:lsid:plazi:treatment:03B8879EAB2E1E3826D8E5D3FA79FCB0:8ECE6053AB2A1E3826DDE6D5FCA6FCE7" box="[692,875,750,774]" country="Brazil" county="Amazonas" latitude="-3.7802777" longLatPrecision="21" longitude="-64.99139" municipality="Tefe" name="ilha do Martelo" pageId="38" pageNumber="39">ilha do Martelo</location>
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,
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(
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measure in
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Tab.
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, 367 mm
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),
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, Guarajá-
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,
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downstream
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,
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,
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, G. M. Santos.

View file

@ -0,0 +1,68 @@
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Subfamily
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,
</heading>
</paragraph>
<paragraph id="8BAE5107FFB2EE6102652112FC89AE98" blockId="3.[812,1486,1176,1480]" box="[812,876,1395,1424]" pageId="3" pageNumber="164">1922</paragraph>
</subSubSection>
<subSubSection id="C30B028CFFB2EE61026521F7FC7DAEC0" pageId="3" pageNumber="164" type="description">
<paragraph id="8BAE5107FFB2EE61026521F7FC7DAEC0" blockId="3.[812,1486,1176,1480]" pageId="3" pageNumber="164">
Genus
<taxonomicName id="4C112A84FFB2EE61023821F7FC09AEA4" box="[881,1004,1430,1452]" class="Reptilia" family="Tyrannosauridae" genus="Tarbosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="164" phylum="Chordata" rank="genus">
<emphasis id="B9658D15FFB2EE61023821F7FC09AEA4" box="[881,1004,1430,1452]" italics="true" pageId="3" pageNumber="164">Tarbosaurus</emphasis>
</taxonomicName>
monospecific, see diagnosis of the species
<taxonomicName id="4C112A84FFB2EE6104E721F6FC77AEC0" authorityName="&quot; Maleev" authorityYear="1955" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="bataar">
<emphasis id="B9658D15FFB2EE6104E721F6FC77AEC0" italics="true" pageId="3" pageNumber="164">Ta. bataar</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="EB40EA5A274ADA9E277E33DE9284C9ED">New albanerpetontid amphibians from the Early Cretaceous of Morocco and Middle Jurassic of England</mods:title>
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<subSubSection id="E2A86548B31E1B0E9D62FB27BE85FB56" box="[812,1202,1176,1205]" pageId="2" pageNumber="303" type="nomenclature">
<paragraph id="AA0D36C3B31E1B0E9D62FB27BE85FB56" blockId="2.[812,1202,1176,1240]" box="[812,1202,1176,1205]" pageId="2" pageNumber="303">
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<taxonomicName id="6DB24D40B31E1B0E9D62FB27BE72FB56" authorityName="Gardner &amp; Evans &amp; Sigogneau-Russell" authorityYear="2003" box="[812,1093,1176,1205]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="2" pageNumber="303" phylum="Chordata" rank="species" species="unicus" status="sp. nov.">
<emphasis id="98C6EAD1B31E1B0E9D62FB27BE72FB56" box="[812,1093,1176,1205]" italics="true" pageId="2" pageNumber="303">Anoualerpeton unicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83F557AAB31E1B0E9A00FB26BE85FB56" box="[1102,1202,1177,1205]" pageId="2" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E2A86548B31E1B049D62FB7DBB11FDC3" lastPageId="8" lastPageNumber="309" pageId="2" pageNumber="303" type="description">
<paragraph id="AA0D36C3B31E1B0E9D62FB7DB9BBFB3B" blockId="2.[812,1202,1176,1240]" box="[812,908,1218,1240]" pageId="2" pageNumber="303">
<figureCitation id="32892A46B31E1B0E9D62FB7DB959FB3B" box="[812,878,1218,1240]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="2" pageNumber="303">Figs. 1</figureCitation>
,
<figureCitation id="32892A46B31E1B0E9D34FB7DB9BFFB3B" box="[890,904,1218,1240]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="2" pageNumber="303">2</figureCitation>
.
</paragraph>
<paragraph id="AA0D36C3B31E1B0E9D62FB53BE4DFAE1" blockId="2.[812,1226,1260,1312]" box="[812,1146,1260,1282]" pageId="2" pageNumber="303">
Anoual species;
<bibRefCitation id="CE234B32B31E1B0E9D85FB53BE40FAE1" author="Gardner, J. D." box="[971,1143,1260,1282]" pageId="2" pageNumber="303" pagination="12 - 22" refId="ref15363" refString="Gardner, J. D. 2002. Monophyly and intra-generic relationships of the amphibian Albanerpeton. Journal of Vertebrate Paleontology 22: 12 - 22." type="journal article" year="2002">Gardner 2002: 12</bibRefCitation>
.
</paragraph>
<paragraph id="AA0D36C3B31E1B0E9D62FAB5BEFDFAC3" blockId="2.[812,1226,1260,1312]" box="[812,1226,1290,1312]" pageId="2" pageNumber="303">
Anoual albanerpetontid;
<bibRefCitation id="CE234B32B31E1B0E9A55FAB5BEF0FAC3" author="Gardner, J. D." box="[1051,1223,1290,1312]" pageId="2" pageNumber="303" pagination="12 - 22" refId="ref15363" refString="Gardner, J. D. 2002. Monophyly and intra-generic relationships of the amphibian Albanerpeton. Journal of Vertebrate Paleontology 22: 12 - 22." type="journal article" year="2002">Gardner 2002: 14</bibRefCitation>
.
</paragraph>
<paragraph id="AA0D36C3B31E1B0E9D62FA8DBF66FA87" blockId="2.[812,1485,1330,1380]" pageId="2" pageNumber="303">
<emphasis id="98C6EAD1B31E1B0E9D62FA8DB9A4FAAB" box="[812,915,1330,1352]" italics="true" pageId="2" pageNumber="303">Etymology</emphasis>
:
<taxonomicName id="6DB24D40B31E1B0E9DEEFA8CBE14FAAB" authority=", Latin" authorityName="Latin" box="[928,1059,1330,1352]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="2" pageNumber="303" phylum="Chordata" rank="species" species="unicus">
<emphasis id="98C6EAD1B31E1B0E9DEEFA8CB9D3FAAB" box="[928,996,1331,1352]" italics="true" pageId="2" pageNumber="303">Unicus</emphasis>
, Latin
</taxonomicName>
“alone” or “solitary”, referring to this being the only known albanerpetontid species from Gondwana.
</paragraph>
<paragraph id="AA0D36C3B31E1B0E9D62FACBBFADFA21" blockId="2.[812,1486,1396,1819]" pageId="2" pageNumber="303">
<materialsCitation id="1ADA3C9EB31E1B0E9D62FACBBFADFA21" collectionCode="MCM" pageId="2" pageNumber="303" specimenCode="MCM 187" specimenCount="1" typeStatus="holotype">
<emphasis id="98C6EAD1B31E1B0E9D62FACBB9B3FA69" box="[812,900,1396,1418]" italics="true" pageId="2" pageNumber="303">
<typeStatus id="75098861B31E1B0E9D62FACBB9B3FA69" box="[812,900,1396,1418]" pageId="2" pageNumber="303" type="holotype">Holotype</typeStatus>
</emphasis>
: MNHN.
<specimenCode id="FA149EB8B31E1B0E9DACFACBBE7BFA69" box="[994,1100,1396,1418]" collectionCode="MCM" country="France" httpUri="http://grbio.org/cool/i15t-gzyx" name="Institut de Paleontologie, Museum d'Histoire naturelle" pageId="2" pageNumber="303">MCM 187</specimenCode>
, right premaxilla missing most of pars palatinumandlateralmostpartofparsdentalis,withpreservedtoothrow containing three broken teeth and five tooth slots (
<figureCitation id="32892A46B31E1B0E9B41FA13BF53FA21" box="[1295,1380,1452,1474]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="2" pageNumber="303">Figs. 1A</figureCitation>
,
<figureCitation id="32892A46B31E1B0E9B20FA13BFB8FA21" box="[1390,1423,1452,1474]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="2" pageNumber="303">2A</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="AA0D36C3B31E1B0E9D62FA6EBF62F8F8" blockId="2.[812,1486,1396,1819]" pageId="2" pageNumber="303">
<emphasis id="98C6EAD1B31E1B0E9D62FA6EBEB2FA04" box="[812,1157,1489,1511]" italics="true" pageId="2" pageNumber="303">
<typeStatus id="75098861B31E1B0E9D62FA6EB9B3FA04" box="[812,900,1489,1511]" pageId="2" pageNumber="303" type="holotype">Holotype</typeStatus>
locality, horizon, and age
</emphasis>
: Anoual microvertebrate locality, about
<quantity id="6D4A9B26B31E1B0E9D29FA52B998F9E0" box="[871,943,1517,1539]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="303" unit="km" value="100.0">100 km</quantity>
east of the city of Anoual and near a fort called Ksar Met Lili, Talsinnt Province, eastern High Atlas Mountains, eastcentral
<collectingCountry id="D2A57653B31E1B0E9BEAF9B6B956F9D8" name="Morocco" pageId="2" pageNumber="303">Morocco</collectingCountry>
(
<bibRefCitation id="CE234B32B31E1B0E9D3FF99ABE96F9D8" author="Sigogneau-Russell, D. &amp; Evans, S. E. &amp; Levine, J. F. &amp; Russell, D. A." box="[881,1185,1573,1595]" pageId="2" pageNumber="303" refId="ref16596" refString="Sigogneau-Russell, D., Evans, S. E., Levine, J. F., and Russell, D. A. 1998." type="book" year="1998">SigogneauRussell et al. 1998</bibRefCitation>
: fig. 1); unnamed nonmarine limestone lens, Couches Rouges (“red beds”) sandstone. The fossiliferous lens occurs within marine beds in the upper part of the Couches Rouges sandstone (
<bibRefCitation id="CE234B32B31E1B0E9DADF9C6BF35F96C" author="Sigogneau-Russell, D. &amp; Monbaron, M. &amp; de Kaenel, E." box="[995,1282,1657,1679]" pageId="2" pageNumber="303" pagination="461 - 483" refId="ref16555" refString="Sigogneau-Russell, D., Monbaron, M., and de Kaenel, E. 1990. Nouvelles donnees sur le gisement a Mammif e res Mesozo i ques du Haut-Atlas Marocain. Geobios 23: 461 - 483." type="journal article" year="1990">SigogneauRussell et al. 1990</bibRefCitation>
) and is interpreted as having been deposited in a deltaic setting (
<bibRefCitation id="CE234B32B31E1B0E9A92F92AB957F924" author="Sigogneau-Russell, D. &amp; Evans, S. E. &amp; Levine, J. F. &amp; Russell, D. A." pageId="2" pageNumber="303" refId="ref16596" refString="Sigogneau-Russell, D., Evans, S. E., Levine, J. F., and Russell, D. A. 1998." type="book" year="1998">SigogneauRussell et al. 1998</bibRefCitation>
). Nanofossils support an Early Cretaceous age, probably Berriasian, for the lens (
<bibRefCitation id="CE234B32B31E1B0E9DA3F972BF2FF900" author="Sigogneau-Russell, D. &amp; Monbaron, M. &amp; de Kaenel, E." box="[1005,1304,1741,1763]" pageId="2" pageNumber="303" pagination="461 - 483" refId="ref16555" refString="Sigogneau-Russell, D., Monbaron, M., and de Kaenel, E. 1990. Nouvelles donnees sur le gisement a Mammif e res Mesozo i ques du Haut-Atlas Marocain. Geobios 23: 461 - 483." type="journal article" year="1990">SigogneauRussell et al. 1990</bibRefCitation>
;
<bibRefCitation id="CE234B32B31E1B0E9B6BF972B9C7F91C" author="Duffin, C. J. &amp; Sigogneau-Russell, D." pageId="2" pageNumber="303" pagination="175 - 190" refId="ref14586" refString="Duffin, C. J. and Sigogneau-Russell, D. 1993. Fossil shark teeth from the Early Cretaceous of Anoual, Morocco. Belgian Geological Survey, Professional Papers 264: 175 - 190." type="journal article" year="1993">Duffin and SigogneauRussell 1993</bibRefCitation>
). See
<bibRefCitation id="CE234B32B31E1B0E9A62F956BF57F91C" author="Sigogneau-Russell, D. &amp; Evans, S. E. &amp; Levine, J. F. &amp; Russell, D. A." box="[1068,1376,1769,1791]" pageId="2" pageNumber="303" refId="ref16596" refString="Sigogneau-Russell, D., Evans, S. E., Levine, J. F., and Russell, D. A. 1998." type="book" year="1998">SigogneauRussell et al. (1998)</bibRefCitation>
and Evans and SigogneauRussell (2001) for additional information.
</paragraph>
<paragraph id="AA0D36C3B31E1B0F9D62F88DB8F2FEC2" blockId="2.[812,1486,1842,2026]" lastBlockId="3.[102,774,232,289]" lastPageId="3" lastPageNumber="304" pageId="2" pageNumber="303">
<emphasis id="98C6EAD1B31E1B0E9D62F88DBE3CF8AA" box="[812,1035,1842,1866]" italics="true" pageId="2" pageNumber="303">Referred specimens</emphasis>
.—Premaxillae (n = 4): MNHN.MCM 188, 198200; maxillae (n = 5): MNHN.MCM 13, 189, 201203; dentaries (n = 34): MNHN.MCM 4, 5, 1934, 4148, 192197, 204, 205; frontals (n = 3): MNHN.MCM 11, 190, 191; parietal (n = 1): MNHN.MCM 12; articular (n = 1): MNHN.MCM 6; humerus (n = 4): MNHN.MCM 7, 1416; trunk vertebrae (n = 15): MNHN.MCM 8, 9, 3540, 4955; and caudal vertebrae (n = 3): MNHN.MCM 10, 17, 18.
</paragraph>
<paragraph id="AA0D36C3B31F1B0F9E28FE8DB8F8FEA9" blockId="3.[102,719,306,330]" box="[102,719,306,330]" pageId="3" pageNumber="304">
<emphasis id="98C6EAD1B31F1B0F9E28FE8DBAD8FEA9" box="[102,239,306,330]" italics="true" pageId="3" pageNumber="304">Distribution</emphasis>
.—Known only from the
<typeStatus id="75098861B31F1B0F9C44FE8DB85BFEA9" box="[522,620,306,330]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
locality.
</paragraph>
<paragraph id="AA0D36C3B31F1B0F9E28FEE2BBA9FD37" blockId="3.[102,775,348,724]" pageId="3" pageNumber="304">
<emphasis id="98C6EAD1B31F1B0F9E28FEE2BAEFFE97" box="[102,216,349,372]" italics="true" pageId="3" pageNumber="304">Diagnosis</emphasis>
.—Species of
<taxonomicName id="6DB24D40B31F1B0F9F39FEE3B82AFE97" authorityName="Gardner &amp; Evans &amp; Sigogneau-Russell" authorityYear="2003" box="[375,541,348,372]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="3" pageNumber="304" phylum="Chordata" rank="genus">
<emphasis id="98C6EAD1B31F1B0F9F39FEE3B82AFE97" box="[375,541,348,372]" italics="true" pageId="3" pageNumber="304">Anoualerpeton</emphasis>
</taxonomicName>
differing from Middle Jurassic congener described below as follows: suprapalatal pit located more medially in pars dorsalis of premaxilla, just medial to margin for external narial opening and in line with fourth or fifth locus from medial end of tooth row; dorsal opening of palatal foramen in premaxilla below or within base of suprapalatal pit; internasal process on frontals relatively shorter (midline length subequal to width across base) and pointed distally; anterolateral processes on frontals pointed distally and distinct from main body of bone; and ventrolateral crests on frontals convex ventrally to bevelled in transverse view.
</paragraph>
<paragraph id="AA0D36C3B31F1B0F9E28FD58B82AFC7D" blockId="3.[102,775,742,926]" pageId="3" pageNumber="304">
<emphasis id="98C6EAD1B31F1B0F9E28FD58BADDFD1D" box="[102,234,743,766]" italics="true" pageId="3" pageNumber="304">Description</emphasis>
.—None of the 71 catalogued specimens at hand is complete, but the available jaws and frontals document most of the structure of these elements. Because the specimens are small and fragile, in some cases we have left cemented matrix attached to the bone rather than risk irreparable damage by attempting to remove it.
</paragraph>
<paragraph id="AA0D36C3B31F1B0F9E28FC0FB8E3FAAB" blockId="3.[102,775,944,2026]" pageId="3" pageNumber="304">
<emphasis id="98C6EAD1B31F1B0F9E28FC0FBAD7FC2B" box="[102,224,944,968]" italics="true" pageId="3" pageNumber="304">Premaxilla</emphasis>
(
<figureCitation id="32892A46B31F1B0F9EA0FC0FBB59FC2B" box="[238,366,944,968]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Figs. 1A, B</figureCitation>
,
<figureCitation id="32892A46B31F1B0F9F37FC0FBB8EFC2B" box="[377,441,944,968]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">2A, B</figureCitation>
).—
<materialsCitation id="1ADA3C9EB31F1B0F9FADFC0FB8E7FAAB" collectionCode="MCM" pageId="3" pageNumber="304" specimenCode="MCM 187, MCM 188" specimenCount="2" typeStatus="holotype">
The two most nearly complete premaxillae are MNHN.
<specimenCode id="FA149EB8B31F1B0F9FE1FC6FB811FC0B" box="[431,550,976,1000]" collectionCode="MCM" country="France" httpUri="http://grbio.org/cool/i15t-gzyx" name="Institut de Paleontologie, Museum d'Histoire naturelle" pageId="3" pageNumber="304">MCM 187</specimenCode>
(
<typeStatus id="75098861B31F1B0F9C7AFC6FB8A0FC0B" box="[564,663,976,1000]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
) and 188. The
<typeStatus id="75098861B31F1B0F9ED8FC4FBAC1FBEB" box="[150,246,1008,1032]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
(
<figureCitation id="32892A46B31F1B0F9F4AFC4FBB53FBEB" box="[260,356,1008,1032]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Figs. 1A</figureCitation>
,
<figureCitation id="32892A46B31F1B0F9F20FC4FBBA6FBEB" box="[366,401,1008,1032]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">2A</figureCitation>
) is from the right side, is about
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high, and retains an intact pars dorsalis, but lacks the more lingual part of the pars palatinum and the lateral end of the pars dentalis. MNHN.
<specimenCode id="FA149EB8B31F1B0F9F68FBEFBB97FB8B" box="[294,416,1104,1128]" collectionCode="MCM" country="France" httpUri="http://grbio.org/cool/i15t-gzyx" name="Institut de Paleontologie, Museum d'Histoire naturelle" pageId="3" pageNumber="304">MCM 188</specimenCode>
(
<figureCitation id="32892A46B31F1B0F9FFCFBEFB821FB8B" box="[434,534,1104,1128]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Figs. 1B</figureCitation>
,
<figureCitation id="32892A46B31F1B0F9C6BFBEFB870FB8B" box="[549,583,1104,1128]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">2B</figureCitation>
) is from the left side, is about
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high, and lacks the lateral end of the pars dentalis, dorsolateral part of the pars dorsalis, and the lateral and medial parts of the pars palatinum. The former specimen is more robust than the latter and is from a slightly larger individual. The remaining premaxillae are from comparablesized individuals, but preserve only the base of the pars dorsalis and varying amounts of the pars palatinum and pars dentalis
</materialsCitation>
.
</paragraph>
<paragraph id="AA0D36C3B31F1B0F9EC5FAEEB930F96B" blockId="3.[102,775,944,2026]" pageId="3" pageNumber="304">The medial edge of each specimen bears elongate grooves and a flange, indicating that the premaxillae were sutured (i.e., paired) in life with their opposite. MNHN.MCM 187 and 188 show that the pars dorsalis is relatively short and broad, with the ratio of height:width across the suprapalatal pit about 1.4. On MNHN.MCM 187 the dorsal edge of the pars dorsalis bears weak suture marks forcontactwith the nasaland the process is indented laterally above the external narial margin by a narrow, shallow notch for receipt of the lacrimal. In labial aspect about the dorsal onethird of the pars dorsalis on</paragraph>
<paragraph id="AA0D36C3B31F1B0F9E28F92EBB4AF84A" blockId="3.[102,775,944,2026]" pageId="3" pageNumber="304">MNHN.MCM 187 bears a low, indistinct boss that is ornamented with low, relatively broad ridges enclosing shallow, irregular pits. MNHN.MCM 188 preserves the medial edge of an evidently less prominent boss. All specimens show that the remainder of the labial face of the bone is relatively smooth and perforated by small, scattered external nutritive foramina. The lateral corner of the pars dentalis is indented by a smooth facet that, in life, was overlapped labially by a complementary process from the maxilla.</paragraph>
<paragraph id="AA0D36C3B31F1B0F9EC5F80DBF89FDA1" blockId="3.[102,775,944,2026]" lastBlockId="3.[812,1486,232,2026]" pageId="3" pageNumber="304">
MNHN.MCM 187 and 188 are the most informative specimens for documenting the lingual structure of the pars dorsalis. The suprapalatal pit (
<figureCitation id="32892A46B31F1B0F9A33FF57BEE5FEE2" box="[1149,1234,232,257]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Fig. 1A</figureCitation>
<figureCitation id="32892A46B31F1B0F9A9DFF4CBF32FEE7" box="[1235,1285,233,260]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">
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, B
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) lies in the lateral half of the process, medial to the external narial margin and in line with the fourth or fifth tooth position from the medial end of the tooth row, and well dorsal to the pars palatinum. The pit is oval in outline, relatively small (i.e., occupies about two percent of lingual surface area of pars dorsalis), and opens laterolingually. To either side, the suprapalatal pit is bracketed by an indistinct internal strut. A few tiny, sedimentinfilled lateral foramina perforate the pars dorsalis laterally, in the wall for the external narial opening. The remainder of the lingual surface of the pars dorsalis is smooth.
</paragraph>
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None of the
<specimenCount id="BCB4FD4AB31F1B0F9DA9FDF3BEA6FD87" box="[999,1169,588,612]" count="5" pageId="3" pageNumber="304" type="generic">five specimens</specimenCount>
retains an intact pars palatinum, but judging by preserved sections and broken surfaces the process was a lingually broad, horizontal shelf as in other albanerpetontids. MNHN.MCM 188 medially preserves the base of the lingually projecting vomerine process. MNHN.MCM 187, 188, and 198 are useful for documenting and interpreting the openings associated with the pars palatinum. The first
<specimenCount id="BCB4FD4AB31F1B0F9A5BFC91BE8BFCA6" box="[1045,1212,813,837]" count="2" pageId="3" pageNumber="304" type="generic">two specimens</specimenCount>
have two tiny foramina that open dorsally in the junction between the pars palatinum and pars dorsalis, in the vicinity of the suprapalatal pit. The larger and more dorsolabial of these two foramina opens just below the suprapalatal pit in MNHN.MCM 187 (
<figureCitation id="32892A46B31F1B0F9B15FC11BF84FC25" box="[1371,1459,942,966]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Fig. 1A</figureCitation>
<figureCitation id="32892A46B31F1B0F9BFDFC07BF8BFC2A" box="[1459,1468,952,969]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">
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), but more dorsally within the ventral margin of the pit in MNHN.MCM 188 (
<figureCitation id="32892A46B31F1B0F9A5FFC51BE50FBE5" box="[1041,1127,1006,1030]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Fig. 1B</figureCitation>
<figureCitation id="32892A46B31F1B0F9A29FC46BE47FBE9" box="[1127,1136,1017,1034]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">
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). We interpret this foramen as the dorsal opening of the palatal foramen, because in other albanerpetontid taxa, including the Jurassic species described below, the dorsal opening of the palatal foramen is consistently present and closely associated with the ventral margin of the suprapalatal pit (
<bibRefCitation id="CE234B32B31F1B0F9ADDFB31BF72FB44" author="Gardner, J. D." box="[1171,1349,1166,1191]" pageId="3" pageNumber="304" pagination="55 - 70" refId="ref15275" refString="Gardner, J. D. 2000 a. Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45: 55 - 70." type="journal article" year="2000">Gardner 2000a</bibRefCitation>
). In life the suprapalatal pit probably housed a gland (
<bibRefCitation id="CE234B32B31F1B0F9B5BFB10B952FB04" author="Fox, R. C. &amp; Naylor, B. G." pageId="3" pageNumber="304" pagination="118 - 128" refId="ref15075" refString="Fox, R. C. and Naylor, B. G. 1982. A reconsideration of the relationships of the fossil amphibian Albanerpeton. Canadian Journal of Earth Sciences 19: 118 - 128." type="journal article" year="1982">Fox and Naylor 1982</bibRefCitation>
) and the palatal foramen may have carried a duct from the gland into the roof of the mouth (
<bibRefCitation id="CE234B32B31F1B0F9AF1FB50BF5BFAE4" author="Gardner, J. D." box="[1215,1388,1263,1287]" pageId="3" pageNumber="304" pagination="55 - 70" refId="ref15275" refString="Gardner, J. D. 2000 a. Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45: 55 - 70." type="journal article" year="2000">Gardner 2000a</bibRefCitation>
). The inferred ventral opening of the palatal foramen lies in the junction between the pars palatinum and pars dentalis in MNHN.MCM 187 (
<figureCitation id="32892A46B31F1B0F9A5BFAF0BE58FA84" box="[1045,1135,1359,1383]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Fig. 1A</figureCitation>
<figureCitation id="32892A46B31F1B0F9A3EFAE5BE4EFA88" box="[1136,1145,1370,1387]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">
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) and, although not visible in figures published here, slightly more lingually in the ventral face of the pars palatinum in MNHN.MCM 188; these positions are consistent with the pattern in other albanerpetontids. In both specimens the ventral opening of the palatal foramen is displaced medially relative to the dorsal opening of the foramen. Although the palatal foramen is plugged with matrix in both specimens, the relative positions of the dorsal and ventral openings imply that the canal connecting the two openings extends dorsolaterallyventromedially through the pars palatinum. This canal is exposed in the broken surface of the pars palatinum on MNHN.MCM 198 (unfigured) and confirms that the canal extended obliquely through the shelf. As for the smaller and more lingual foramen (= “unknown foramen” in
<figureCitation id="32892A46B31F1B0F9DF4F8AEBE38F8CA" box="[954,1039,1809,1833]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="3" pageNumber="304">Fig. 1A</figureCitation>
<figureCitation id="32892A46B31F1B0F9A41F8A3BE75F8CE" box="[1039,1090,1809,1837]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="3" pageNumber="304">
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, B
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) that opens in the dorsal surface of the pars palatinum, judging by the condition in other albanerpetontids this foramen likely communicates, via a canal extending obliquely through the pars palatinum, with a similarly small foramen that opens in the lingual face of the pars dentalis, medial to and slightly below the ventral opening of the palatal foramen. The function of this smaller foramen is unknown. The pars dentalis is deep. Tiny foramina pierce the lingual face of the pars dentalis above some tooth positions.
<emphasis id="98C6EAD1B3181B089D62F80FB9B4F82B" box="[812,899,1968,1992]" italics="true" pageId="4" pageNumber="305">Maxilla</emphasis>
(
<figureCitation id="32892A46B3181B089DDBF80FB9CEF82A" box="[917,1017,1968,1993]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="4" pageNumber="305">Figs. 1C</figureCitation>
,
<figureCitation id="32892A46B3181B089A47F80FBE1EF82B" box="[1033,1065,1968,1992]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="4" pageNumber="305">2C</figureCitation>
).—The two best preserved maxillae collectively document all but the posteriormost end of the bone. MNHN.MCM 189 (
<figureCitation id="32892A46B3191B099FC4FF57BBDCFEE2" box="[394,491,232,257]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="5" pageNumber="306">Figs. 1C</figureCitation>
,
<figureCitation id="32892A46B3191B099FB9FF56B82FFEE2" box="[503,536,233,257]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="5" pageNumber="306">2C</figureCitation>
) and 201 (unfigured) are broken well behind the internal narial margin and preserve about the anterior twothirds to fourfifths of the bone. The former specimen is from the left side and has the first 17 loci, whereas the latter is from the right side and retains the first 14 loci. MNHN.MCM 189 is about
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long and was probably about
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long when the bone was intact; MNHN.MCM 201 is from a slightly smaller individual. Available specimens resemble other albanerpetontid maxillae as follows: pars facialis elongate and low, becoming shallower posteriorly; nasal process projects dorsally, triangular in outline, and bevelled posteriorly for articulation with lacrimal; premaxillary dorsal and lateral processes prominent, the former a lingually expanded shelf and the latter an anteriorly projecting flange; pars palatinum a lingually broad shelf, narrowing posteriorly, with lingual edge indented anteriorly by concave internal narial margin, and, more posteriorly, bearing shallow trough dorsolingually for articulation with unknown palatal bone(s); and pars dentalis deep, becoming shallower posteriorly. The labial face of the bone is smooth, except for tiny external nutritive foramina scattered across the more anterior part of the pars facialis. The premaxillary lateral process is relatively long (i.e., length greater than height at base) and blunt distally. Unlike many other albanerpetontid maxillae, including the Jurassic material described below, the dorsal surface of the pars palatinum adjacent to the nasal process lacks a saddleshaped bony patch for contact with the base of the lacrimal. Also, the ventral surface of the premaxillary dorsal process lacks the transverse ridge that, in life, abutted against the lateral edge of the maxillary process on the premaxilla when the two bones were articulated. The occlusal edge of the pars dentalis is convex ventrally in labial or lingual outline. In MNHN.MCM 189 the pars dentalis is deepest adjacent to about the sixth locus. Tiny foramina are variably present in the lingual face of the pars dentalis above the teeth. The anterior end of the tooth row lies several loci anterior to the level of the leading edge of the nasal process.
</paragraph>
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<paragraph id="AA0D36C3B3181B089E28F910BF30F865" blockId="4.[102,1486,1711,1927]" pageId="4" pageNumber="305">
Fig. 1. Upper jaws and frontals of
<taxonomicName id="6DB24D40B3181B089FE6F910B846F921" authorityName="Gardner &amp; Evans &amp; Sigogneau-Russell" authorityYear="2003" box="[424,625,1711,1730]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="4" pageNumber="305" phylum="Chordata" rank="species" species="unicus" status="sp. nov.">
<emphasis id="98C6EAD1B3181B089FE6F910B846F921" box="[424,625,1711,1730]" italics="true" pageId="4" pageNumber="305">Anoualerpeton unicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83F557AAB3181B089C37F90FB8F3F920" box="[633,708,1712,1731]" pageId="4" pageNumber="305" rank="species">sp. nov.</taxonomicNameLabel>
, type species; Lower Cretaceous (Berriasian), Anoual, Morocco.
<emphasis id="98C6EAD1B3181B089B6CF910BF05F921" bold="true" box="[1314,1330,1711,1730]" pageId="4" pageNumber="305">A</emphasis>
,
<emphasis id="98C6EAD1B3181B089B71F910BF79F921" bold="true" box="[1343,1358,1711,1730]" pageId="4" pageNumber="305">B</emphasis>
. Premaxillae.
<emphasis id="98C6EAD1B3181B089E28F974BA41F93D" bold="true" box="[102,118,1739,1758]" pageId="4" pageNumber="305">A</emphasis>
. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A
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) and lingual (A
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) views.
<emphasis id="98C6EAD1B3181B089A08F974BE62F93D" bold="true" box="[1094,1109,1739,1758]" pageId="4" pageNumber="305">B</emphasis>
. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B
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) and lingual (B
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) views.
<emphasis id="98C6EAD1B3181B089B4CF958BF25F919" bold="true" box="[1282,1298,1767,1786]" pageId="4" pageNumber="305">C</emphasis>
. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C
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), lingual (C
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), and dorsal (C
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) views.
<emphasis id="98C6EAD1B3181B089A46F8BCBE2FF8F5" bold="true" box="[1032,1048,1795,1814]" pageId="4" pageNumber="305">D</emphasis>
,
<emphasis id="98C6EAD1B3181B089A6CF8BCBE06F8F5" bold="true" box="[1058,1073,1795,1814]" pageId="4" pageNumber="305">E</emphasis>
. Fused frontals.
<emphasis id="98C6EAD1B3181B089A88F8BCBEE1F8F5" bold="true" box="[1222,1238,1795,1814]" pageId="4" pageNumber="305">D</emphasis>
. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D
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) and ventral (D
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) views and closeup of anterior part in right lateral view (D
<subScript id="36363486B3181B089C9FF8FBB8EEF8B1" attach="left" box="[721,729,1860,1874]" fontSize="6" pageId="4" pageNumber="305">3</subScript>
).
<emphasis id="98C6EAD1B3181B089CA5F884B8CDF8AD" bold="true" box="[747,762,1851,1870]" pageId="4" pageNumber="305">E</emphasis>
. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales.
</paragraph>
</caption>
<paragraph id="AA0D36C3B3191B099E28FA00B9D1FB66" blockId="5.[102,775,1470,1752]" lastBlockId="5.[812,1485,232,1158]" pageId="5" pageNumber="306">
<emphasis id="98C6EAD1B3191B099E28FA00BAF5FA35" box="[102,194,1471,1494]" italics="true" pageId="5" pageNumber="306">Dentary</emphasis>
(
<figureCitation id="32892A46B3191B099E9CFA01BB79FA34" box="[210,334,1470,1495]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="5" pageNumber="306">Fig. 2DG</figureCitation>
).—A catalogued size series of 34 broken dentaries is available. The most nearly complete specimen, MNHN.MCM 204 (
<figureCitation id="32892A46B3191B099FCDFA40BBECF9F4" box="[387,475,1535,1559]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="5" pageNumber="306">Fig. 2D</figureCitation>
), is a left dentary that is intact from the symphysis back to the level of the anterior margin of the opening for the Meckelian canal and bears 30 tooth positions with 25 teeth and five tooth slots. The specimen is about
<quantity id="6D4A9B26B3191B099EE1F93FBB3EF974" box="[175,265,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="306" unit="mm" value="3.6">3.6 mm</quantity>
long and would have been about onethird again as long when the bone was complete. In dentaries from Anoual the occlusal edge of the dental parapet is strongly convex dorsally in labial or lingual outline, with the apex labial to the eighthtenth loci. This condition is most pronounced in larger specimens, such as MNHN.MCM 204 and MNHN.MCM 195 (
<figureCitation id="32892A46B3191B099A43FEF6BE54FE82" box="[1037,1123,329,353]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="5" pageNumber="306">Fig. 2D</figureCitation>
<figureCitation id="32892A46B3191B099A2DFEEBBE5BFE86" box="[1123,1132,340,357]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="5" pageNumber="306">
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, E, respectively), but it also occurs in smaller specimens (e.g., MNHN.MCM 4 and 5; unfigured). In other respects, the structure of the dentary is typical for albanerpetontids. In dorsal aspect the bone is broadly curved. The labial face is smooth and a row of up to five or six external nutritive foramina extends along the anterior half of the bone. Below these foramina a low ridge is variably developed; this ridge marks the upper boundary of a shallow scar that extends anteroposteriorly along the lateroventral and ventral surfaces of the bone, for attachment of the intermandibularis musculature. A foramen opens in the underside of the symphysis. The symphyseal face is vertical anteriorly and posteriorly bears one or two symphyseal prongs that, in life, interlocked in a mortiseandtenon fashion with one or two complementary prongs on the opposite dentary. In lingual aspect the dental parapet is moderately deep. The subdental shelf is moderately broad lingually, shallow, and gutter shaped anteriorly, and becomes deeper and narrower posteriorly. No dentary has the area for attachment of the postdentary bones intact, but several specimens, including MNHN.MCM 192 and 193 (
<figureCitation id="32892A46B3191B099A39FC72BEC5FC06" box="[1143,1266,973,997]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="5" pageNumber="306">Fig. 2F, G</figureCitation>
, respectively), preserve enough of this region to show that the opening for the Meckelian canal extends forward below about the posterior onequarter to onethird of the tooth row and that the dorsal edge of the bone behind the tooth row was smooth and lacked a dorsal process.
</paragraph>
<paragraph id="AA0D36C3B3191B0A9D62FB22B824F806" blockId="5.[812,1485,1181,1752]" lastBlockId="6.[102,1485,1862,2021]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
<emphasis id="98C6EAD1B3191B099D62FB22B95DFB56" box="[812,874,1181,1205]" italics="true" pageId="5" pageNumber="306">Teeth</emphasis>
(
<figureCitation id="32892A46B3191B099D37FB22B9CCFB56" box="[889,1019,1181,1205]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="5" pageNumber="306">Figs. 1AC</figureCitation>
<figureCitation id="32892A46B3191B099DB5FB18BE7BFB56" box="[1019,1100,1181,1208]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="5" pageNumber="306">
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; 2AI
</figureCitation>
).—Marginal teeth are characteristic for albanerpetontids in being highly pleurodont, nonpedicellate, straight, closely packed, and in having crowns that are labiolingually compressed and bear three mesiodistallyaligned cuspules. The structure of the tooth crowns is more variable than in species of
<taxonomicName id="6DB24D40B3191B099AE3FA81BF72FAB5" baseAuthorityName="Gardner" baseAuthorityYear="2002" box="[1197,1349,1342,1366]" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="98C6EAD1B3191B099AE3FA81BF72FAB5" box="[1197,1349,1342,1366]" italics="true" pageId="5" pageNumber="306">Albanerpeton</emphasis>
</taxonomicName>
. Crowns on some teeth in
<taxonomicName id="6DB24D40B3191B099D98FAE1BEE6FA95" authorityName="Gardner &amp; Evans &amp; Sigogneau-Russell" authorityYear="2003" box="[982,1233,1374,1398]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="unicus">
<emphasis id="98C6EAD1B3191B099D98FAE1BEE6FA95" box="[982,1233,1374,1398]" italics="true" pageId="5" pageNumber="306">Anoualerpeton unicus</emphasis>
</taxonomicName>
are decidedly chisellike, with the central cuspule relatively low and the occlusal surface of the crown essentially convex in lingual outline; this pattern is typical for
<taxonomicName id="6DB24D40B3191B099A04FA01BED5FA35" baseAuthorityName="Gardner" baseAuthorityYear="2002" box="[1098,1250,1470,1494]" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="98C6EAD1B3191B099A04FA01BED5FA35" box="[1098,1250,1470,1494]" italics="true" pageId="5" pageNumber="306">Albanerpeton</emphasis>
</taxonomicName>
. Some tooth crowns in
<taxonomicName id="6DB24D40B3191B099D06FA60B98AFA15" authorityName="Gardner &amp; Evans &amp; Sigogneau-Russell" authorityYear="2003" box="[840,957,1503,1526]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="unicus">
<emphasis id="98C6EAD1B3191B099D06FA60B98AFA15" box="[840,957,1503,1526]" italics="true" pageId="5" pageNumber="306">An. unicus</emphasis>
</taxonomicName>
are more wedgeshaped in lingual or labial outline, with the central cuspule considerably longer than the mesial and distal cuspules (
<figureCitation id="32892A46B3191B099A28F9A0BEEEF9D4" box="[1126,1241,1567,1591]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="5" pageNumber="306">Fig. 2H, I</figureCitation>
). Similar variation in tooth crown structure also occurs in the Jurassic congener described below and in an indeterminate Early Cretaceous (Berriasian)
<taxonomicName id="6DB24D40B3191B099DF0F93FBE6FF974" box="[958,1112,1664,1688]" class="Amphibia" family="Albanerpetontidae" genus="Celtedens" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="98C6EAD1B3191B099DF0F93FBE1BF97B" box="[958,1068,1664,1688]" italics="true" pageId="5" pageNumber="306">Celtedens</emphasis>
sp.
</taxonomicName>
from Purbeck,
<collectingCountry id="D2A57653B3191B099B5BF93FBF45F97B" box="[1301,1394,1664,1688]" name="United Kingdom" pageId="5" pageNumber="306">England</collectingCountry>
(
<bibRefCitation id="CE234B32B3191B099BC9F93FBE2CF95B" author="Evans, S. E. &amp; McGowan, G. J." pageId="5" pageNumber="306" pagination="103 - 119" refId="ref14838" refString="Evans, S. E. and McGowan, G. J. 2002. An amphibian assemblage from the PurbeckLimestoneGroup. SpecialPapersinPalaeontology 68: 103 - 119." type="journal article" year="2002">Evans and McGowan 2002</bibRefCitation>
: pl. 1: 9); in these European taxa the central cuspule on some teeth is even more elongate, resultincomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view.
<emphasis id="98C6EAD1B31A1B0A9B64F8F9BF0BF8BA" bold="true" box="[1322,1340,1862,1881]" pageId="6" pageNumber="307">H</emphasis>
,
<emphasis id="98C6EAD1B31A1B0A9B09F8F9BF67F8BA" bold="true" box="[1351,1360,1862,1881]" pageId="6" pageNumber="307">I</emphasis>
. Close ups of tooth crowns, both in lingual view.
<emphasis id="98C6EAD1B31A1B0A9FECF8DDBB83F896" bold="true" box="[418,436,1890,1909]" pageId="6" pageNumber="307">H</emphasis>
. MNHN.MCM 199, right premaxilla, crown of tooth at sixth locus from medial end of tooth row.
<emphasis id="98C6EAD1B31A1B0A9B6FF8DDBF1DF896" bold="true" box="[1313,1322,1890,1909]" pageId="6" pageNumber="307">I</emphasis>
. MNHN.MCM 5, right dentary, crown of tooth at eighth locus from anterior end of tooth row.
<emphasis id="98C6EAD1B31A1B0A9D4CF8C1B93AF872" bold="true" box="[770,781,1918,1937]" pageId="6" pageNumber="307">J</emphasis>
,
<emphasis id="98C6EAD1B31A1B0A9D59F8C1B91EF872" bold="true" box="[791,809,1918,1937]" pageId="6" pageNumber="307">K</emphasis>
. Fused frontals, both in ventral view.
<emphasis id="98C6EAD1B31A1B0A9A38F8C1BEB6F872" bold="true" box="[1142,1153,1918,1937]" pageId="6" pageNumber="307">J</emphasis>
. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides.
<emphasis id="98C6EAD1B31A1B0A9A0DF825BE62F84E" bold="true" box="[1091,1109,1946,1965]" pageId="6" pageNumber="307">K</emphasis>
. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone, photographed before damage (cf.,
<figureCitation id="32892A46B31A1B0A9EFBF86DBAC3F806" box="[181,244,2002,2021]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="6" pageNumber="307">Fig 1E</figureCitation>
). Specimens at different scales.
</paragraph>
<caption id="FECD664BB3191B099E28F8A8BFFAF80B" pageId="5" pageNumber="306" startId="5.[102,135,1815,1834]" targetType="figure">
<paragraph id="AA0D36C3B3191B099E28F8A8BFFAF80B" blockId="5.[102,1486,1815,2025]" pageId="5" pageNumber="306">
Fig. 2. Jaws and frontals of
<taxonomicName id="6DB24D40B3191B099F30F8A8B87CF8CA" authorityName="Gardner &amp; Evans &amp; Sigogneau-Russell" authorityYear="2003" box="[382,587,1815,1834]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="unicus" status="sp. nov.">
<emphasis id="98C6EAD1B3191B099F30F8A8B87CF8CA" box="[382,587,1815,1834]" italics="true" pageId="5" pageNumber="306">Anoualerpeton unicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83F557AAB3191B099C1BF8A8B893F8C9" box="[597,676,1815,1834]" pageId="5" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
, type species; Lower Cretaceous (Berriasian), Anoual, Morocco.
<emphasis id="98C6EAD1B3191B099B55F8A8BF1CF8C9" bold="true" box="[1307,1323,1815,1834]" pageId="5" pageNumber="306">A</emphasis>
,
<emphasis id="98C6EAD1B3191B099B72F8A8BF7CF8C9" bold="true" box="[1340,1355,1815,1834]" pageId="5" pageNumber="306">B</emphasis>
. Premaxillae.
<emphasis id="98C6EAD1B3191B099E28F88DBA41F8A6" bold="true" box="[102,118,1842,1861]" pageId="5" pageNumber="306">A</emphasis>
. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A
<subScript id="36363486B3191B099DECF884B99DF8AA" attach="right" box="[930,938,1851,1865]" fontSize="6" pageId="5" pageNumber="306">1</subScript>
) and laterolingual (A
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) views.
<emphasis id="98C6EAD1B3191B099A9AF88DBED4F8A6" bold="true" box="[1236,1251,1842,1861]" pageId="5" pageNumber="306">B</emphasis>
. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B
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) and laterolingual (B
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) and slightly dorsal views.
<emphasis id="98C6EAD1B3191B099F60F8D7BB09F898" bold="true" box="[302,318,1896,1915]" pageId="5" pageNumber="306">C</emphasis>
. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view.
<emphasis id="98C6EAD1B3191B099B72F8D6BF5DF898" bold="true" box="[1340,1386,1896,1916]" pageId="5" pageNumber="306">DG</emphasis>
, dentaries.
<emphasis id="98C6EAD1B3191B099E28F83BBA41F874" bold="true" box="[102,118,1924,1943]" pageId="5" pageNumber="306">D</emphasis>
. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D
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), lingual (D
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), and occlusal (D
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) views.
<emphasis id="98C6EAD1B3191B099D5CF820B916F851" bold="true" box="[786,801,1951,1970]" pageId="5" pageNumber="306">E</emphasis>
. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view.
<emphasis id="98C6EAD1B3191B099C51F805B81AF82E" bold="true" box="[543,557,1978,1997]" pageId="5" pageNumber="306">F</emphasis>
. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view.
<emphasis id="98C6EAD1B3191B099DEFF86AB984F80B" bold="true" box="[929,947,2005,2024]" pageId="5" pageNumber="306">G</emphasis>
. MNHN.MCM 193, left dentary, anteriorly and posteriorly
</paragraph>
</caption>
<paragraph id="AA0D36C3B31B1B0B9E28FF57BA95FC82" blockId="7.[102,775,232,2026]" pageId="7" pageNumber="308">
ing in a more pointed crown. In jaws from Anoual the premaxillary teeth and the more anterior teeth on the maxilla and dentary tend to have wedgeshaped crowns, whereas teeth farther back tend to have more chiselshaped crowns. Tooth counts are typical for albanerpetontids. MNHN.MCM 198 is the only premaxilla with an intact tooth row and preserves ten tooth positions. No maxilla or dentary preserves a complete tooth row, but specimens with anatomically overlapping sections of the row yield estimated, maximum counts of, respectively, 25 and 35 tooth positions. Most specimens preserve evidence of tooth replacement in the form of empty replacement slots and several, such as the maxilla MNHN. MCM 189 (
<figureCitation id="32892A46B31B1B0B9EA1FDD7BB66FD62" box="[239,337,616,641]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Figs. 1C</figureCitation>
<figureCitation id="32892A46B31B1B0B9F1FFDCCBBB8FD62" box="[337,399,617,644]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="7" pageNumber="308">
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, 2C
</figureCitation>
: anteriormost tooth), also have a lingual resorption pit in the base of one or more teeth. The dentary MNHN.MCM 34 (unfigured) preserves a nonfunctional, replacement tooth at the second locus. Teeth on the maxilla and dentary are strongly heterodont in size, with teeth about onethird of the distance along the tooth row from the anterior end being longer than nearby teeth (
<figureCitation id="32892A46B31B1B0B9CDAFC97B8C1FCA2" box="[660,758,808,833]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Figs. 1C</figureCitation>
<figureCitation id="32892A46B31B1B0B9CB8FC8CBAA6FC87" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="7" pageNumber="308">
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, 2D
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).
</paragraph>
<paragraph id="AA0D36C3B31B1B0B9E28FCCDBFF4FDC6" blockId="7.[102,775,232,2026]" lastBlockId="7.[812,1485,232,1351]" pageId="7" pageNumber="308">
<emphasis id="98C6EAD1B31B1B0B9E28FCCDBAF1FC69" box="[102,198,882,906]" italics="true" pageId="7" pageNumber="308">Frontals</emphasis>
(
<figureCitation id="32892A46B31B1B0B9E99FCCDBB69FC6A" box="[215,350,882,906]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Figs. 1D, E</figureCitation>
,
<figureCitation id="32892A46B31B1B0B9F23FCCDBB9CFC69" box="[365,427,882,906]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="7" pageNumber="308">2J, K</figureCitation>
).—The two most nearly complete specimens are figured here: MNHN.MCM 190 (
<figureCitation id="32892A46B31B1B0B9C80FC2DBABEFC29" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Figs. 1D</figureCitation>
,
<figureCitation id="32892A46B31B1B0B9ED4FC0DBA83FC29" box="[154,180,946,970]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="7" pageNumber="308">2J</figureCitation>
) and MNHN.MCM 191 (
<figureCitation id="32892A46B31B1B0B9FAAFC0DB87FFC29" box="[484,584,946,970]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Figs. 1E</figureCitation>
,
<figureCitation id="32892A46B31B1B0B9C17FC0DB848FC29" box="[601,639,946,970]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="7" pageNumber="308">2K</figureCitation>
). The more nearly complete specimen, MNHN.MCM 190, lacks only the distal end of the left anterolateral process and the posterior end of the ventrolateral crests on both sides. The two halves are solidly fused medially. The specimen is
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in midline length and
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wide across the posterior edge, which yields a relative length (ratio of midline length:posterior width) of about 1.5. The internasal process is acuminate in dorsal or ventral outline and relatively broad, with the midline length subequal to the width across the base. The lateral face of the internasal process bears an elongate groove (
<figureCitation id="32892A46B31B1B0B9E20FAADBAF3FAC9" box="[110,196,1298,1322]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Fig. 1D</figureCitation>
<figureCitation id="32892A46B31B1B0B9E8BFAA3BAF9FACE" box="[197,206,1308,1325]" captionStart="Fig" captionStartId="9.[102,135,1780,1799]" captionTargetBox="[106,1481,225,1752]" captionTargetId="figure-8@9.[106,1481,225,1752]" captionTargetPageId="9" captionText="Fig. 3. Jaws of Anoualerpeton priscus sp. nov.; Middle Jurassic (late Bathonian), Kirtlington, England. AC. Premaxillae. A. BMNH R.16336, holotype, left premaxilla, lacking dorsal part of pars dorsalis, lateral part of pars dentalis, and most of pars palatinum and preserving no intact teeth, in lingual (A1) and laterolingual (A2) views. B. BMNH R.16337, right premaxilla, lacking dorsal part of pars dorsalis and medial pars of pars palatinum and pars dentalis, in labial (B1) and lingual (B2) views. C. BMNH R.14157, right premaxilla, lacking dorsal part of pars dorsalis, ventral part of pars dentalis, and lateral end of maxillary process and preserving no intact teeth, in lingual (C1) and dorsal (C2) views, both with hair extending obliquely through palatal foramen, and in occlusal (C3) view. D. BMNH R.16338, left maxilla, missing part of pars dentalis below nasal process and about posterior onefifth of bone, in labial (D1), lingual (D2), and dorsal (D3) views. EH. Dentaries, all in lingual view. E. BMNH R.16344, left dentary, posteriorly incomplete ramus preserving about anterior onequarter of bone. F. BMNH R.16354, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones; G, BMNH R.16356, left dentary, anteriorly and posteriorly incomplete ramus preserving about posterior" pageId="7" pageNumber="308">
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</figureCitation>
) for articulation with the medial edge of the nasal. The anterolateral process is distinct from the main body of the bone and is pointed distally. The more anterior slot between the internasal and anterolateral processes for receipt of the posterior end of the nasal and the more posterior slot between the anterolateral process and the orbital margin for receipt of the posterior end of the prefrontal are both deep. The dorsal margin of the latter slot is shallowly excavated medially. The anterior end of the orbital margin, as demarcated by the posterior end of the slot for receipt of the prefrontal, is approximately in line with the anteroposterior midpoint of the bone. MNHN.MCM 190 is vaguely bellshaped in outline, with the posterior edge nearly two times wider than the distance between the slots for receipt of the prefrontals. Behind the base of the anterolateral process, the lateral edge of the bone first extends posteriorly in a straight line, then curves outwards in a shallow arc at about 25° from the midline. The posterior edge of the bone is transverse and shallowly concave to either side of the midline. Dorsally, the bone is ornamented with shallow, broad polygonal pits enclosed by a network of narrower, low ridges. In ventral view, the ventrolateral crest is relatively narrow—i.e., crest at the anterior limit of the orbital margin is about 0.4 times as wide as the width across the posterior edge of the frontals, between the medial edges of the crest. The ventrolateral crest is somewhat convex ventrally in transverse profile, with the more lateral part along the orbital margin shallowly bevelled and facing ventrolaterally. Posteriorly the medial face of the ventrolateral crest is indented by a facet for receipt of a complementary process from the parietal. Although the ventrolateral crest is broken more posteriorly, this structure probably extended farther back to underlap the parietal, as in other albanerpetontids (e.g.,
<bibRefCitation id="CE234B32B31B1B0B9A62FDB2BF6EFDC6" author="Estes, R. &amp; Hoffstetter, R." box="[1068,1369,525,549]" pageId="7" pageNumber="308" pagination="297 - 343" refId="ref14711" refString="Estes, R. and Hoffstetter, R. 1976. Les Urod e les du Mioc e ne de La Grive- Saint-Alban (Is e re, France). Bulletin du Museum national d'Histoire naturelle, 3 e Serie, no. 398, Sciences de la Terre 57: 297 - 343." type="journal article" year="1976">Estes and Hoffstetter 1976</bibRefCitation>
: pl. 8: 2).
</paragraph>
<paragraph id="AA0D36C3B31B1B0B9D1CFD8FBFFAFC64" blockId="7.[812,1485,232,1351]" pageId="7" pageNumber="308">
MNHN.MCM 191 (
<figureCitation id="32892A46B31B1B0B9A73FD90BE95FDAB" box="[1085,1186,559,584]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Figs. 1E</figureCitation>
,
<figureCitation id="32892A46B31B1B0B9AFDFD8FBEE0FDAB" box="[1203,1239,560,584]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="7" pageNumber="308">2K</figureCitation>
) is a larger, but less nearly complete fused pair of frontals. Originally the specimen lacked only the distal ends of the internasal and left anterolateral processes, the posterior end of the left ventrolateral crest, and the right posterolateral corner (
<figureCitation id="32892A46B31B1B0B9B2CFD10BFF7FD2B" box="[1378,1472,687,712]" captionStart="Fig" captionStartId="5.[102,135,1815,1834]" captionText="Fig. 2. Jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in lingual (A1) and laterolingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in lingual (B1) and laterolingual (B2) and slightly dorsal views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in lingual and slightly ventral view. DG, dentaries. D. MNHN.MCM 204, left dentary, posteriorly incomplete ramus, broken at level of opening for Meckelian canal and preserving about anterior threequarters of bone, in labial (D1), lingual (D2), and occlusal (D3) views. E. MNHN.MCM 195, right dentary, posteriorly incomplete ramus preserving about anterior twofifths of bone, in lingual view. F. MNHN.MCM 192, right dentary, anteriorly and posteriorly incomplete ramus preserving posterior part of tooth row and anterior part of area for attachment of postdentary bones, in lingual view. G. MNHN.MCM 193, left dentary, anteriorly and posteriorly" pageId="7" pageNumber="308">Fig. 2K</figureCitation>
). The specimen was broken during photography and some bone was lost from the median area (
<figureCitation id="32892A46B31B1B0B9A89FD50BF2BFCEB" box="[1223,1308,751,776]" captionStart="Fig" captionStartId="4.[102,135,1711,1730]" captionTargetBox="[108,1478,241,1674]" captionTargetId="figure-49@4.[102,1484,225,1684]" captionTargetPageId="4" captionText="Fig. 1. Upper jaws and frontals of Anoualerpeton unicus sp. nov., type species; Lower Cretaceous (Berriasian), Anoual, Morocco. A, B. Premaxillae. A. MNHN.MCM 187, holotype, right premaxilla, lacking pars palatinum, in labial (A1) and lingual (A2) views. B. MNHN.MCM 188, left premaxilla, lacking dorsolateral part of pars dorsalis and vomerine and maxillary processes on pars palatinum, in labial (B1) and lingual (B2) views. C. MNHN.MCM 189, left maxilla, lacking about posterior onefifth of bone, in labial (C1), lingual (C2), and dorsal (C3) views. D, E. Fused frontals. D. MNHN.MCM 190, nearly complete frontals, lacking distal end of left anterolateral process and posterior end of ventrolateral crests on both sides, entire specimen in dorsal (D1) and ventral (D2) views and closeup of anterior part in right lateral view (D3). E. MNHN.MCM 191, less nearly complete frontals, missing distal tip of internasal process, posterior end of left ventrolateral crest, and right posterolateral corner of bone and showing damage to median portion sustained during photography (cf., Fig. 2K), in dorsal view. White areas are broken surfaces and cross hatches are sand grains. Specimens at different scales." pageId="7" pageNumber="308">Fig. 1E</figureCitation>
). The specimen is
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long in midline length and was probably
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long or slightly longer when complete; the posterior width is estimated at about 3.0 mm. The estimated ratio of midline length:posterior width is 1.6, which compares favorably with
</paragraph>
<paragraph id="AA0D36C3B31B1B0B9D62FC2FBE3AFAA4" blockId="7.[812,1485,232,1351]" pageId="7" pageNumber="308">
MNHN.MCM 190. Based on the assumption that snoutpelvic length in albanerpetontids is about ten times the midline length of the frontals (
<bibRefCitation id="CE234B32B31B1B0B9A61FC70BED6FC04" author="Gardner, J. D." box="[1071,1249,975,999]" pageId="7" pageNumber="308" pagination="529 - 544" refId="ref15179" refString="Gardner, J. D. 1999 b. The amphibian Albanerpeton arthridion and the Aptian - Albian biogeography of albanerpetontids. Palaeontology 42: 529 - 544." type="journal article" year="1999">Gardner 1999b</bibRefCitation>
), MNHN.MCM 191 suggests a snoutpelvic length of about
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. In addition to being larger, MNHN.MCM 191 differs from MNHN.MCM 190 as follows: bone somewhat more triangular in dorsal or ventral outline; laterodorsal margins of internasal process straighter in dorsal or ventral outline (but with midline length of process remaining subequal to width across the base); lateral edge of bone more nearly concave along its entire length and extending laterally at a lesser angle of about 20° in dorsal or ventral outline; and ventral face of ventrolateral crest more bevelled. In other respects, MNHN.MCM 190 and 191 resemble one another.
</paragraph>
<paragraph id="AA0D36C3B31B1B0B9D62FAE1BF0BF995" blockId="7.[812,1486,1374,1654]" pageId="7" pageNumber="308">
<emphasis id="98C6EAD1B31B1B0B9D62FAE1B9E3FA95" box="[812,980,1374,1398]" italics="true" pageId="7" pageNumber="308">Other elements</emphasis>
.—An articular, 15 trunk vertebrae, three caudal vertebrae, and four humeri complete the inventory of albanerpetontid material from Anoual. These are typical for albanerpetontids and do not differ appreciably from homologous specimens reported from the Middle Jurassic (
<bibRefCitation id="CE234B32B31B1B0B9B18FA61B950F9F5" author="McGowan, G. J." pageId="7" pageNumber="308" pagination="227 - 234" refId="ref15663" refString="McGowan, G. J. 1996. Albanerpetontid amphibians from the Jurassic (Bathonian) of southern England. Bulletin of the Museum of Northern Arizona 60: 227 - 234." type="journal article" year="1996">McGowan 1996</bibRefCitation>
), Lower Cretaceous (
<bibRefCitation id="CE234B32B31B1B0B9A02FA41BF67F9F5" author="Estes, R. &amp; Sanchiz, B." box="[1100,1360,1534,1558]" pageId="7" pageNumber="308" pagination="21 - 39" refId="ref14767" refString="Estes, R. and Sanchiz, B. 1982. Early Cretaceous lower vertebrates from Galve (Teruel), Spain. Journal of Vertebrate Paleontology 2: 21 - 39." type="journal article" year="1982">Estes and Sanchíz 1982</bibRefCitation>
;
<bibRefCitation id="CE234B32B31B1B0B9B16FA41B9DCF9D5" author="McGowan, G. J. &amp; Ensom, P. C." pageId="7" pageNumber="308" pagination="113 - 117" refId="ref15799" refString="McGowan, G. J. and Ensom, P. C. 1997. Albanerpetontid amphibians from the Lower Cretaceous of the Isle of Purbeck, Dorset. Proceedings of the Dorset Natural History and Archaeological Society 118: 113 - 117." type="journal article" year="1997">McGowan and Ensom 1997</bibRefCitation>
), and Miocene (
<bibRefCitation id="CE234B32B31B1B0B9AD3F9A1BFF1F9D5" author="Estes, R. &amp; Hoffstetter, R." box="[1181,1478,1566,1590]" pageId="7" pageNumber="308" pagination="297 - 343" refId="ref14711" refString="Estes, R. and Hoffstetter, R. 1976. Les Urod e les du Mioc e ne de La Grive- Saint-Alban (Is e re, France). Bulletin du Museum national d'Histoire naturelle, 3 e Serie, no. 398, Sciences de la Terre 57: 297 - 343." type="journal article" year="1976">Estes and Hoffstetter 1976</bibRefCitation>
) of Europe and from the Lower Cretaceous of North America (
<bibRefCitation id="CE234B32B31B1B0B9D7DF9E1BE4DF995" author="Gardner, J. D. &amp; Averianov, A. O." box="[819,1146,1630,1654]" pageId="7" pageNumber="308" pagination="453 - 467" refId="ref15390" refString="Gardner, J. D. and Averianov, A. O. 1998. Albanerpetontid amphibiansfrom Middle Asia. Acta Palaeontologica Polonica 43: 453 - 467." type="journal article" year="1998">Gardner and Averianov 1998</bibRefCitation>
;
<bibRefCitation id="CE234B32B31B1B0B9AC8F9E1BF06F995" author="Gardner, J. D." box="[1158,1329,1630,1654]" pageId="7" pageNumber="308" pagination="529 - 544" refId="ref15179" refString="Gardner, J. D. 1999 b. The amphibian Albanerpeton arthridion and the Aptian - Albian biogeography of albanerpetontids. Palaeontology 42: 529 - 544." type="journal article" year="1999">Gardner 1999b</bibRefCitation>
).
</paragraph>
<paragraph id="AA0D36C3B31B1B049D62F932BB11FDC3" blockId="7.[812,1485,1677,2026]" lastBlockId="8.[102,775,232,544]" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
<emphasis id="98C6EAD1B31B1B0B9D62F932B9B9F946" box="[812,910,1677,1701]" italics="true" pageId="7" pageNumber="308">Remarks</emphasis>
.—Association of albanerpetontid specimens from Anoual within one species is supported by three lines of evidence. First, the provenance of the specimens is unique—no other albanerpetontids are known from Gondwana. Second, the specimens were collected from a single horizon at one locality. The fossiliferous lens at Anoual is restricted, with a depth of only about
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and an areal extent of some
<quantity id="6D4A9B26B31B1B0B9B31F8EFBFF3F884" box="[1407,1476,1872,1896]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="308" unit="m" value="200.0">200 m</quantity>
<superScript id="5DC79B8BB31B1B0B9B8AF8F2BFFAF8BD" attach="left" box="[1476,1485,1869,1886]" fontSize="7" pageId="7" pageNumber="308">2</superScript>
(
<bibRefCitation id="CE234B32B31B1B0B9D7DF8CFBEBAF86B" author="Sigogneau-Russell, D. &amp; Evans, S. E. &amp; Levine, J. F. &amp; Russell, D. A." box="[819,1165,1904,1928]" pageId="7" pageNumber="308" refId="ref16596" refString="Sigogneau-Russell, D., Evans, S. E., Levine, J. F., and Russell, D. A. 1998." type="book" year="1998">SigogneauRussell et al. 1998</bibRefCitation>
). Third, there is no compelling morphological evidence that more than one taxon of albanerpetontid is represented. The maxillae and dentaries have two complementary attributes that justify their association: occlusal margin of jaw convex and teeth strongly heterodont in size anteriorly. Comparisons with other albanerpetontids known by more extensive series of frontals and premaxillae, particularly species of
<taxonomicName id="6DB24D40B3141B049C78FEF7B8F9FE83" baseAuthorityName="Gardner" baseAuthorityYear="2002" box="[566,718,328,352]" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="8" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="98C6EAD1B3141B049C78FEF7B8F9FE83" box="[566,718,328,352]" italics="true" pageId="8" pageNumber="309">Albanerpeton</emphasis>
</taxonomicName>
(see
<bibRefCitation id="CE234B32B3141B049E28FED7BB2EFE62" author="Gardner, J. D." box="[102,281,360,385]" pageId="8" pageNumber="309" pagination="55 - 70" refId="ref15275" refString="Gardner, J. D. 2000 a. Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45: 55 - 70." type="journal article" year="2000">Gardner 2000a</bibRefCitation>
, b), suggest that differences among the Anoual premaxillae in relative build and proportions and among frontals in overall shape and in details of the internasal process, lateral edge of the bone, and transverse crest are within the expected limits of intraspecific variation for albanerpetontids.
</paragraph>
</subSubSection>
</treatment>
</document>