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<document id="978B9214E80DF1982D98EE8C52106AF0" ID-DOI="10.5281/zenodo.14681715" ID-ISSN="0253-116X" ID-Zenodo-Dep="14681715" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737165364000" checkinUser="felipe" docAuthor="Schmid, Christian &amp; Wolf-H" docDate="2024" docId="03A387D5FFD0FFAFD9CE08B50B80BB52" docLanguage="en" docName="LinzerbiolBeitr.56.1.301-311.pdf" docOrigin="Linzer biologische Beiträge 56 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.12:LinzerbiolBeitr.2006-.journal_article.type1" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article.type1" docStyleVersion="12" docTitle="Carinostigmus marocensis" docType="treatment" docVersion="2" lastPageNumber="303" masterDocId="FF9AFFADFFD1FFADD96D091E0A78BA53" masterDocTitle="New and rarely collected digger wasps from Morocco with description of two new species (Hymenoptera, Spheciformes)" masterLastPageNumber="311" masterPageNumber="301" pageNumber="302" updateTime="1737165766856" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="82548EC3D339252D7B5B798E58201685">New and rarely collected digger wasps from Morocco with description of two new species (Hymenoptera, Spheciformes)</mods:title>
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<mods:namePart id="D175D233FE026C4D9F9E5F8335C0D83D">Schmid, Christian</mods:namePart>
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<mods:date id="730E0F6EE7692BEB0990393625DEED71">2024</mods:date>
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<treatment id="03A387D5FFD0FFAFD9CE08B50B80BB52" ID-DOI="http://doi.org/10.5281/zenodo.14681717" ID-Zenodo-Dep="14681717" LSID="urn:lsid:plazi:treatment:03A387D5FFD0FFAFD9CE08B50B80BB52" httpUri="http://treatment.plazi.org/id/03A387D5FFD0FFAFD9CE08B50B80BB52" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302">
<subSubSection id="C3106548FFD0FFACD9CE08B50998BB93" box="[163,992,426,449]" pageId="1" pageNumber="302" type="nomenclature">
<paragraph id="8BB536C3FFD0FFACD9CE08B50998BB93" blockId="1.[163,992,426,449]" box="[163,992,426,449]" pageId="1" pageNumber="302">
<heading id="D0FD81AFFFD0FFACD9CE08B50998BB93" box="[163,992,426,449]" fontSize="8" level="2" pageId="1" pageNumber="302" reason="2">
<taxonomicName id="4C0A4D40FFD0FFACD9CE08B50815BB93" ID-CoL="RDQV" authority="(TSUNEKI, 1956)" baseAuthorityName="TSUNEKI" baseAuthorityYear="1956" box="[163,621,426,448]" class="Insecta" family="Crabronidae" genus="Carinostigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="marocensis">
<emphasis id="B97EEAD1FFD0FFACD9CE08B5085DBBEC" bold="true" box="[163,549,426,448]" pageId="1" pageNumber="302">
<emphasis id="B97EEAD1FFD0FFACD9CE08B50BC1BB93" bold="true" box="[163,441,427,448]" italics="true" pageId="1" pageNumber="302">Carinostigmus marocensis</emphasis>
(
<bibRefCitation id="EF9B4B32FFD0FFACD8AA08B4085DBBEC" author="TSUNEKI K." box="[455,549,426,448]" pageId="1" pageNumber="302" pagination="263 - 264" refId="ref3979" refString="TSUNEKI K. (1956): A new species of Stigmus from Morocco (Hymen., Sphecidae, Pemphredoninae). - Entomologische Berichten 16: 263 - 264." type="journal article" year="1956">TSUNEKI</bibRefCitation>
</emphasis>
,
<emphasis id="B97EEAD1FFD0FFACDB5F08B40815BB93" bold="true" box="[562,621,426,448]" pageId="1" pageNumber="302">1956)</emphasis>
</taxonomicName>
<emphasis id="B97EEAD1FFD0FFACDB1E08B50998BB93" bold="true" box="[627,992,426,449]" pageId="1" pageNumber="302">
– description of male (
<figureCitation id="13312A46FFD0FFACDA0F08B409CABB93" box="[866,946,426,448]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="1" pageNumber="302">figs 1-3</figureCitation>
,
<figureCitation id="13312A46FFD0FFACDAD208B409A1BB93" box="[959,985,426,448]" captionStart="Fig" captionStartId="10.[163,196,843,865]" captionTargetBox="[163,997,207,833]" captionTargetId="figure-33@10.[163,997,207,833]" captionTargetPageId="10" captionText="Fig. 21: Distribution of Carinostigmus marocensis (blue), Spilomena canariensis (red) and the African distribution of Entomognathus fortuitus (green)." figureDoi="http://doi.org/10.5281/zenodo.14681727" httpUri="https://zenodo.org/record/14681727/files/figure.png" pageId="1" pageNumber="302">21</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3106548FFD0FFACD9CE08CE0BCDB84F" pageId="1" pageNumber="302" type="reference_group">
<paragraph id="8BB536C3FFD0FFACD9CE08CE09CFBBB6" blockId="1.[163,1070,463,541]" box="[163,951,463,486]" pageId="1" pageNumber="302">
<treatmentCitationGroup id="AB1A11EDFFD0FFACD9CE08CE09CFBBB6" box="[163,951,463,486]" pageId="1" pageNumber="302">
<treatmentCitation id="0AAB10D2FFD0FFACD9CE08CE0860BBB6" author="TSUNEKI K." box="[163,536,463,485]" page="263" pageId="1" pageNumber="302" year="1956">
<taxonomicName id="4C0A4D40FFD0FFACD9CE08CE0860BBB6" ID-CoL="52NMS" authority="TSUNEKI, 1956: 263" authorityName="TSUNEKI" authorityPageNumber="263" authorityYear="1956" box="[163,536,463,485]" class="Insecta" family="Crabronidae" genus="Stigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="marocensis">
<emphasis id="B97EEAD1FFD0FFACD9CE08CE0B2FBBB6" box="[163,343,464,485]" italics="true" pageId="1" pageNumber="302">Stigmus marocensis</emphasis>
<materialsCitation id="3B623C9EFFD0FFACD83008CE0860BBB6" box="[349,536,463,485]" collectionCode="RMNH" country="Morocco" location="Morocco" pageId="1" pageNumber="302" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<bibRefCitation id="EF9B4B32FFD0FFACD83008CE0860BBB6" author="TSUNEKI K." box="[349,536,463,485]" pageId="1" pageNumber="302" pagination="263 - 264" refId="ref3979" refString="TSUNEKI K. (1956): A new species of Stigmus from Morocco (Hymen., Sphecidae, Pemphredoninae). - Entomologische Berichten 16: 263 - 264." type="journal article" year="1956">
<smallCapsWord id="8D53A01FFFD0FFACD83008CE0BD7BBB7" baselines="480,480" box="[349,431,464,485]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Tsuneki" pageId="1" pageNumber="302">TSUNEKI</smallCapsWord>
, 1956: 263
</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="9D0CFD4AFFD0FFACDB4C08CE084BBBB6" box="[545,563,464,485]" count="1" pageId="1" pageNumber="302" type="female">♀</specimenCount>
.
<typeStatus id="54B18861FFD0FFACDB5308CE08EEBBB5" box="[574,662,464,486]" pageId="1" pageNumber="302" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D0CFD4AFFD0FFACDBF308CE08D7BBB6" box="[670,687,464,485]" count="1" pageId="1" pageNumber="302" type="female">♀</specimenCount>
,
<collectingCountry id="F31D7653FFD0FFACDBD608CE0968BBB6" box="[699,784,464,485]" name="Morocco" pageId="1" pageNumber="302">Morocco</collectingCountry>
: Agadir (RMNH).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB536C3FFD0FFACD9CE08EE0BCDB84F" blockId="1.[163,1070,463,541]" pageId="1" pageNumber="302">
M a t e r i a l e x a m i n e d: ♁
<date id="FFB41003FFD0FFACDB6A08F1081DB856" box="[519,613,495,517]" pageId="1" pageNumber="302" value="2024-03-23">23.iii.2024</date>
<collectingCountry id="F31D7653FFD0FFACDB0208EE08B8B856" box="[623,704,496,517]" name="Morocco" pageId="1" pageNumber="302">Morocco</collectingCountry>
, Dakhla-Oued Ed-Dahab, Dakhla town
<geoCoordinate id="EE3E5004FFD0FFACD9BD0B190B64B84E" box="[208,284,519,541]" degrees="27.168" direction="north" orientation="latitude" pageId="1" pageNumber="302" precision="55" value="27.168">27.168N</geoCoordinate>
<geoCoordinate id="EE3E5004FFD0FFACD84C0B190B0AB84E" box="[289,370,519,541]" degrees="15.920" direction="west" orientation="longitude" pageId="1" pageNumber="302" precision="55" value="-15.92">15.920W</geoCoordinate>
(CSE).
</paragraph>
</subSubSection>
<subSubSection id="C3106548FFD0FFAFD9CF0B330B80BB52" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302" type="description">
<paragraph id="8BB536C3FFD0FFACD9CF0B3309B0B8A6" blockId="1.[162,1070,556,757]" pageId="1" pageNumber="302">
R e m a r k
<taxonomicName id="4C0A4D40FFD0FFACD8550B320BB5B812" authorityName="Tsuneki" authorityYear="1954" box="[312,461,556,577]" class="Insecta" family="Crabronidae" genus="Carinostigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAD1FFD0FFACD8550B320BB5B812" box="[312,461,556,577]" italics="true" pageId="1" pageNumber="302">Carinostigmus</emphasis>
</taxonomicName>
is a small genus, and most species are distributed in southern and eastern Africa and in southeastern Asia, a few also in the Palaearctic region. Species from the Ethiopian region can be identified with
<bibRefCitation id="EF9B4B32FFD0FFACDA140B760E52B82D" author="LECLERQ J." box="[889,1066,616,638]" pageId="1" pageNumber="302" pagination="17 - 62" refId="ref3644" refString="LECLERQ J. (1959): Pemphredoninae (Hymenoptera Sphecoidea) in Exploration du Parc National de l'Upemba. I. Mission G. F. de Witte en collaboration avec ADAM W., JANSSENS A., VAN MEEL L. et R. VERHEYEN (1946 - 1949), Fasc. 53 (2): 17 - 62." type="journal article" year="1959">LECLERQ (1959)</bibRefCitation>
, southeastern Asian species with
<bibRefCitation id="EF9B4B32FFD0FFACD89B0B9808ECB8C8" author="TSUNEKI K." box="[502,660,645,667]" pageId="1" pageNumber="302" pagination="1 - 38" refId="ref3927" refString="TSUNEKI K. (1954): The genus Stigmus PANZER of Europe and Asia, with description of eight new species (Hymenoptera, Sphecidae). - Memoirs of the Faculty of Liberal Arts, Fukui University (Series II, Natural Science) 3: 1 - 38." type="journal article" year="1954">TSUNEKI (1954</bibRefCitation>
a) and others. Three species are known from the western Palaearctic region, and only one,
<emphasis id="B97EEAD1FFD0FFACDBC10BBA093CB8EA" box="[684,836,676,697]" italics="true" pageId="1" pageNumber="302">
<taxonomicName id="4C0A4D40FFD0FFACDBC10BBA0938B8EA" authorityName="TSUNEKI" authorityYear="1956" box="[684,832,676,697]" class="Insecta" family="Crabronidae" genus="Stigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="marocensis">C. marocensis</taxonomicName>
,
</emphasis>
in northwest Africa. It was therefore only known by the female
<typeStatus id="54B18861FFD0FFACDB3F0BDC08D4B884" box="[594,684,706,727]" pageId="1" pageNumber="302" type="paratype">paratype</typeStatus>
, described from
<collectingCountry id="F31D7653FFD0FFACDA300BDF09DFB884" box="[861,935,705,727]" name="Morocco" pageId="1" pageNumber="302">Agadir</collectingCountry>
. The male is here described for the first time. It agree in general with the female description.
</paragraph>
<paragraph id="8BB536C3FFD0FFACD9CE0A1B0BA3B9BF" blockId="1.[163,1070,773,1004]" pageId="1" pageNumber="302">
With the identification key of
<bibRefCitation id="EF9B4B32FFD0FFACD8B60A1B08FEB948" author="LECLERQ J." box="[475,646,773,795]" pageId="1" pageNumber="302" pagination="17 - 62" refId="ref3644" refString="LECLERQ J. (1959): Pemphredoninae (Hymenoptera Sphecoidea) in Exploration du Parc National de l'Upemba. I. Mission G. F. de Witte en collaboration avec ADAM W., JANSSENS A., VAN MEEL L. et R. VERHEYEN (1946 - 1949), Fasc. 53 (2): 17 - 62." type="journal article" year="1959">LECLERQ (1959)</bibRefCitation>
, the male keyes out with
<taxonomicName id="4C0A4D40FFD0FFACDAE20A1B0B3BB96B" authority="(R. TURNER, 1912)" baseAuthorityName="R. TURNER" baseAuthorityYear="1912" class="Insecta" family="Crabronidae" genus="Carinostigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="gueinzius">
<emphasis id="B97EEAD1FFD0FFACDAE20A1B0E71B948" box="[911,1033,773,795]" italics="true" pageId="1" pageNumber="302">C. gueinzius</emphasis>
(R. TURNER, 1912)
</taxonomicName>
. This species could not be examined. However, according to the description, the only detectable difference in the male are the setae on the underside of the mandible, which are said to be arranged in clusters in
<taxonomicName id="4C0A4D40FFD0FFACDB9E0A410914B927" baseAuthorityName="R. TURNER" baseAuthorityYear="1912" box="[755,876,863,884]" class="Insecta" family="Crabronidae" genus="Carinostigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="gueinzius">
<emphasis id="B97EEAD1FFD0FFACDB9E0A410914B927" box="[755,876,863,884]" italics="true" pageId="1" pageNumber="302">C. gueinzius</emphasis>
</taxonomicName>
, but are solitary in
<taxonomicName id="4C0A4D40FFD0FFACD9CE0A630B4EB9C1" authorityName="TSUNEKI" authorityYear="1956" box="[163,310,893,914]" class="Insecta" family="Crabronidae" genus="Stigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="marocensis">
<emphasis id="B97EEAD1FFD0FFACD9CE0A630B4EB9C1" box="[163,310,893,914]" italics="true" pageId="1" pageNumber="302">C. marocensis</emphasis>
</taxonomicName>
. Otherwise both agree. As
<taxonomicName id="4C0A4D40FFD0FFACDB3A0A6308AAB9C1" baseAuthorityName="R. TURNER" baseAuthorityYear="1912" box="[599,722,893,914]" class="Insecta" family="Crabronidae" genus="Carinostigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="gueinzius">
<emphasis id="B97EEAD1FFD0FFACDB3A0A6308AAB9C1" box="[599,722,893,914]" italics="true" pageId="1" pageNumber="302">C. gueinzius</emphasis>
</taxonomicName>
is widespread in southern Africa and known from West Africa and also reaches the
<collectingCountry id="F31D7653FFD0FFACDBC80A840966B9E3" box="[677,798,922,944]" name="Cape Verde" pageId="1" pageNumber="302">Cape Verde</collectingCountry>
Islands (
<bibRefCitation id="EF9B4B32FFD0FFACDA170A840E5AB9FC" author="PULAWSKI W. J." box="[890,1058,922,943]" pageId="1" pageNumber="302" refId="ref3748" refString="PULAWSKI W. J. (2024): https: // researcharchive. calacademy. org / research / entomology / entomology _ resources / hymenoptera / sphecidae / genera /" type="url" year="2024">PULAWSKI 2024</bibRefCitation>
), both species should be compared and it is not entirely impossible that
<taxonomicName id="4C0A4D40FFD0FFACDAEF0AA70E6CB99D" authorityName="TSUNEKI" authorityYear="1956" box="[898,1044,953,974]" class="Insecta" family="Crabronidae" genus="Stigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="marocensis">
<emphasis id="B97EEAD1FFD0FFACDAEF0AA70E6CB99D" box="[898,1044,953,974]" italics="true" pageId="1" pageNumber="302">C. marocensis</emphasis>
</taxonomicName>
is synonymous with
<taxonomicName id="4C0A4D40FFD0FFACD8310AC80BADB9BF" baseAuthorityName="R. TURNER" baseAuthorityYear="1912" box="[348,469,982,1004]" class="Insecta" family="Crabronidae" genus="Carinostigmus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="gueinzius">
<emphasis id="B97EEAD1FFD0FFACD8310AC80BADB9BF" box="[348,469,982,1004]" italics="true" pageId="1" pageNumber="302">C. gueinzius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB536C3FFD0FFACD9CE0AE20BB9BF81" blockId="1.[163,1070,1020,1490]" pageId="1" pageNumber="302">
D e s c r i p t i o n o f m a l e: 4.0 mm. Color: Black. Mandible except brown apex, labrum and pronotal lobe yellow.
<collectingCountry id="F31D7653FFD0FFACDB520D040827BE7C" box="[575,607,1050,1071]" name="American Samoa" pageId="1" pageNumber="302">AS</collectingCountry>
1-2,
<collectingCountry id="F31D7653FFD0FFACDBFA0D0408CFBE7C" box="[663,695,1050,1071]" name="American Samoa" pageId="1" pageNumber="302">AS</collectingCountry>
3-8 below, fore- and midtarsi and – tibiae brownish-yellow (
<figureCitation id="13312A46FFD0FFACD8F60D260BA9BE1E" box="[411,465,1080,1102]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="1" pageNumber="302">fig. 1</figureCitation>
). Morphology: Apex of mandible with two teeth, mandible below with 6 pale setae, as long as mandibular diameter. Labrum medially deeply emarginate, ACM slightly rounded, clypeus finely crosswise striae. Frons below scape deeply impressed, with small spine, crosswise striate. Inner eye margin with strong furrow, ending at level of foreocellus, clearly cross-ribbed (
<figureCitation id="13312A46FFD0FFACDA590DB1090ABE97" box="[820,882,1199,1221]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="1" pageNumber="302">fig. 3</figureCitation>
). Head divergent behind eye. Occipital carina strong. Gena striate.
<collectingCountry id="F31D7653FFD0FFACDBBF0DD3088ABEB1" box="[722,754,1229,1250]" name="American Samoa" pageId="1" pageNumber="302">AS</collectingCountry>
3-9 widened downwards to roundish. Pronotum 0,3x as long as apical width. Anteriorly with strong lamella, mediolaterally each with 5 coarse oblique rugae. Large (cross-ribbed) part of notauli short, ending at the top at the end of the arch of the mesonotum, pronlonged by a thin line until mid-mesonotum. Mesonotum apically with triangular impression, including 3 rugae. Base of scutellum with deep furrow, also distinctly fielded. Propodeal dorsum anteriorly and medially fielded and shiny, laterally with large polished areas. Petiole somewhat longer than hindtibia (7:6), dorsally somewhat impressed with lateral edge. Tergites polished and shiny.
</paragraph>
<paragraph id="8BB536C3FFD0FFACD9CE0CFC0930BC60" blockId="1.[163,1070,1506,1587]" pageId="1" pageNumber="302">
D i s t r i b u t i o n: Only known from the
<typeStatus id="54B18861FFD0FFACDB100CFC08D1BFA4" box="[637,681,1506,1527]" pageId="1" pageNumber="302">type</typeStatus>
location
<collectingCountry id="F31D7653FFD0FFACDA600CFC092BBFAB" box="[781,851,1506,1528]" name="Morocco" pageId="1" pageNumber="302">Agadir</collectingCountry>
and from Dakhla in
<collectingCountry id="F31D7653FFD0FFACD9CE0F1E0A86BC46" box="[163,254,1536,1557]" name="Morocco" pageId="1" pageNumber="302">Morocco</collectingCountry>
(
<figureCitation id="13312A46FFD0FFACD8630F1E0B2EBC46" box="[270,342,1536,1558]" captionStart="Fig" captionStartId="10.[163,196,843,865]" captionTargetBox="[163,997,207,833]" captionTargetId="figure-33@10.[163,997,207,833]" captionTargetPageId="10" captionText="Fig. 21: Distribution of Carinostigmus marocensis (blue), Spilomena canariensis (red) and the African distribution of Entomognathus fortuitus (green)." figureDoi="http://doi.org/10.5281/zenodo.14681727" httpUri="https://zenodo.org/record/14681727/files/figure.png" pageId="1" pageNumber="302">fig. 21</figureCitation>
). It can be expected that the species occurs also in other places along the coast of the atlantic Ocean and can also be found further south.
</paragraph>
<paragraph id="8BB536C3FFD3FFAFD9CE09D10B80BB52" blockId="2.[163,1070,207,257]" pageId="2" pageNumber="303">H a b i t a t: The specimen was catched in a small garden area at the western border of Dakhla town on flowering plants.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="61D5000E6F0EFDDAB69E96CB48C42058">New and rarely collected digger wasps from Morocco with description of two new species (Hymenoptera, Spheciformes)</mods:title>
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<taxonomicName id="4C0A4D40FFD3FFAFD9CE083508EDBB12" ID-CoL="6FGDN" authority="(VANDER LINDEN, 1829)" baseAuthorityName="VANDER LINDEN" baseAuthorityYear="1829" box="[163,661,299,321]" class="Insecta" family="Crabronidae" genus="Entomognathus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="B97EEAD1FFD3FFAFD9CE08350BF7BB12" bold="true" box="[163,399,299,321]" italics="true" pageId="2" pageNumber="303">Entomognathus brevis</emphasis>
(VANDER LINDEN, 1829)
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(
<figureCitation id="13312A46FFD3FFAFDBCF083508A2BB12" box="[674,730,299,321]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="2" pageNumber="303">fig. 4</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BB536C3FFD3FFAFD9CE084E0E56BB35" blockId="2.[163,1070,335,437]" box="[163,1070,335,358]" pageId="2" pageNumber="303">
<taxonomicName id="4C0A4D40FFD3FFAFD9CE084E085FBB35" ID-CoL="Z3CZ" authority="VANDER LINDEN, 1829: 72" authorityName="VANDER LINDEN" authorityPageNumber="72" authorityYear="1829" box="[163,551,336,358]" class="Insecta" family="Crabronidae" genus="Crabro" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="B97EEAD1FFD3FFAFD9CE084E0B5BBB36" box="[163,291,336,357]" italics="true" pageId="2" pageNumber="303">Crabro brevis</emphasis>
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, 1829: 72
</taxonomicName>
,
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, ♁.
<typeStatus id="54B18861FFD3FFAFDB06084F08BABB35" box="[619,706,337,358]" pageId="2" pageNumber="303" type="syntype">Syntypes</typeStatus>
:
<collectingCountry id="F31D7653FFD3FFAFDBA6084E0964BB35" box="[715,796,336,358]" name="Belgium" pageId="2" pageNumber="303">Belgium</collectingCountry>
:
<collectingRegion id="49CEF821FFD3FFAFDA4B084E0905BB35" box="[806,893,336,358]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="2" pageNumber="303">Bruxelles</collectingRegion>
and
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: Bologna
</paragraph>
<paragraph id="8BB536C3FFD3FFAFD9A408790E56BBCE" blockId="2.[163,1070,335,437]" pageId="2" pageNumber="303">
(Inst. Roy. Sci. Nat.
<collectingCountry id="F31D7653FFD3FFAFD8EE08790BAFBB2E" box="[387,471,359,381]" name="Belgium" pageId="2" pageNumber="303">Belgique</collectingCountry>
,
<collectingRegion id="49CEF821FFD3FFAFD88D08790843BB2E" box="[480,571,359,381]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="2" pageNumber="303">Bruxelles</collectingRegion>
). M a t e r i a l e x a m i n e d: ♁
<date id="FFB41003FFD3FFAFDB6708990819BBCE" box="[522,609,391,413]" pageId="2" pageNumber="303" value="2015-04-16">16.4.2015</date>
<quantity id="4CF29B26FFD3FFAFDB00089908D3BBCE" box="[621,683,391,413]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="303" unit="km" value="80.0">80 km</quantity>
NE
<collectingCountry id="F31D7653FFD3FFAFDB8D08990967BBCE" box="[736,799,391,413]" name="Morocco" pageId="2" pageNumber="303">Agadir</collectingCountry>
, Talmakant,
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<geoCoordinate id="EE3E5004FFD3FFAFDA8508990E56BBCE" box="[1000,1070,391,413]" degrees="8.956" direction="west" orientation="longitude" pageId="2" pageNumber="303" precision="55" value="-8.956">8.956W</geoCoordinate>
</paragraph>
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(CSE);
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<date id="FFB41003FFD3FFAFD84508810B07BBE7" box="[296,383,415,436]" pageId="2" pageNumber="303" value="2020-04-24">
<collectingDate id="EFF0E9EBFFD3FFAFD84508810B07BBE7" box="[296,383,415,436]" pageId="2" pageNumber="303" value="2020-04-24">24.4.2020</collectingDate>
</date>
Sidi Slimane,
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(
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)
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.
</paragraph>
<paragraph id="8BB536C3FFD3FFAFD9CF08DA0B97B94F" blockId="2.[162,1070,451,796]" pageId="2" pageNumber="303">
R e m a r k
<taxonomicName id="4C0A4D40FFD3FFAFD85908DD0877BB8B" baseAuthorityName="VANDER LINDEN" baseAuthorityYear="1829" box="[308,527,451,472]" class="Insecta" family="Crabronidae" genus="Entomognathus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="B97EEAD1FFD3FFAFD85908DD0877BB8B" box="[308,527,451,472]" italics="true" pageId="2" pageNumber="303">Entomognathus brevis</emphasis>
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was already mentioned for the fauna of
<collectingCountry id="F31D7653FFD3FFAFDAD808DA0E74BB8A" box="[949,1036,452,473]" name="Morocco" pageId="2" pageNumber="303">Morocco</collectingCountry>
by some authors (see
<bibRefCitation id="EF9B4B32FFD3FFAFD83808FF0B87BBA5" author="PULAWSKI W. J." box="[341,511,481,502]" pageId="2" pageNumber="303" refId="ref3748" refString="PULAWSKI W. J. (2024): https: // researcharchive. calacademy. org / research / entomology / entomology _ resources / hymenoptera / sphecidae / genera /" type="url" year="2024">PULAWSKI, 2024</bibRefCitation>
), but it seems to be very rare. We could only examine
<specimenCount id="9D0CFD4AFFD3FFAFDD6708FF0B72B847" count="2" pageId="2" pageNumber="303" type="generic">two specimens</specimenCount>
. The specimens from
<collectingCountry id="F31D7653FFD3FFAFD89208E0082EB840" box="[511,598,510,531]" name="Morocco" pageId="2" pageNumber="303">Morocco</collectingCountry>
differ distinctly by colour pattern from the European population (
<figureCitation id="13312A46FFD3FFAFD8EE0B050BC5B863" box="[387,445,539,561]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="2" pageNumber="303">fig. 4</figureCitation>
). Morphology is similar in all details. Therefore, there is no reason to see two species here. The first genetic barcoding (with the CO1 gene) of the Moroccan specimens results in a separate clade compared to examined specimens from
<collectingCountry id="F31D7653FFD3FFAFD9CE0B6D0A87B8DB" box="[163,255,627,648]" name="Germany" pageId="2" pageNumber="303">Germany</collectingCountry>
, which however is closely related with European species (SCHMID- EGGER et al. 2018). These results therefore do not justify a separate species status. SCHMID- EGGER et al. (2018) found 4.1% intraspecific distance in the species with a separate BIN of single moroccan specimens, but a second barcode of another specimen from
<collectingCountry id="F31D7653FFD3FFAFDA0B0BD209C4B8B2" box="[870,956,716,737]" name="Morocco" pageId="2" pageNumber="303">Morocco</collectingCountry>
even shifts the result and only gives a common clade with the European specimens with a minimal gap in between (Schmid-Egger, unpubl.).
</paragraph>
<paragraph id="8BB536C3FFD3FFAFD9CE0A320B55B980" blockId="2.[163,1070,811,980]" pageId="2" pageNumber="303">
D e s c r i p t i o n: Color pattern of male and female from
<collectingCountry id="F31D7653FFD3FFAFDA4F0A3209F9B912" box="[802,897,812,833]" name="Morocco" pageId="2" pageNumber="303">Morocco</collectingCountry>
: Black, with the following parts yellow:
<collectingCountry id="F31D7653FFD3FFAFD8F10A570BB8B90D" box="[412,448,841,862]" name="American Samoa" pageId="2" pageNumber="303">AS</collectingCountry>
1 (scape), tibiae, tarsi (the latter dark yellow), apical half of fore- and midfemora, apex of hindfemur, pronotum and pronotal lobe, scutellum, metanotum. European and Asian specimens have usually mesosoma all black except pronotal lobe and tegula,
<collectingCountry id="F31D7653FFD3FFAFD8C30ABF0BAAB9E5" box="[430,466,929,950]" name="American Samoa" pageId="2" pageNumber="303">AS</collectingCountry>
1 is only yellow below, femora are black and tibiae partly black (
<figureCitation id="13312A46FFD3FFAFD98A0AA00B58B980" box="[231,288,958,980]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="2" pageNumber="303">fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B97EEAD1FFD3FFAFD9CE0AE00BD3BE47" bold="true" box="[163,427,1022,1044]" italics="true" pageId="2" pageNumber="303">Entomognathus fortuitus</emphasis>
(KOHL
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– new record for
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(
<figureCitation id="13312A46FFD3FFAFDA030AE009C7BE40" box="[878,959,1022,1044]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="2" pageNumber="303">figs 5-6</figureCitation>
,
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)
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<emphasis id="B97EEAD1FFD3FFAFD9CE0D3D0B40BE6B" box="[163,312,1059,1080]" italics="true" pageId="2" pageNumber="303">Crabro fortuitus</emphasis>
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, 1915: 314
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.
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:
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,
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: no specific locality (NHMW)
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M a t e r i a l e x a m i n e d:
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<date id="FFB41003FFD3FFAFD8940D5D0829BE0B" box="[505,593,1091,1112]" pageId="2" pageNumber="303" value="1997-03-19">19.3.1997</date>
Ht. Atlas,
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SE Ait Ourir
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(leg. S. Blank, coll CSE).
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R e m a r k
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<emphasis id="B97EEAD1FFD3FFAFD8560D61084BBEC7" box="[315,563,1151,1172]" italics="true" pageId="2" pageNumber="303">Entomognathus fortuitus</emphasis>
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was previously only know from
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and
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. The present record is the first in
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and Africa (
<figureCitation id="13312A46FFD3FFAFDA5F0D830901BEE1" box="[818,889,1181,1203]" captionStart="Fig" captionStartId="10.[163,196,843,865]" captionTargetBox="[163,997,207,833]" captionTargetId="figure-33@10.[163,997,207,833]" captionTargetPageId="10" captionText="Fig. 21: Distribution of Carinostigmus marocensis (blue), Spilomena canariensis (red) and the African distribution of Entomognathus fortuitus (green)." figureDoi="http://doi.org/10.5281/zenodo.14681727" httpUri="https://zenodo.org/record/14681727/files/figure.png" pageId="2" pageNumber="303">fig. 21</figureCitation>
). The species can be identified with
<bibRefCitation id="EF9B4B32FFD3FFAFD80C0DA4084CBE9C" author="BITSCH J. &amp; BARBIER Y. &amp; GAYUBO S. F. &amp; JACOBS H. - J. &amp; LECLERCQ J. &amp; K. SCHMIDT" box="[353,564,1210,1232]" pageId="2" pageNumber="303" refId="ref3417" refString="BITSCH J., BARBIER Y., GAYUBO S. F., JACOBS H. - J., LECLERCQ J. &amp; K. SCHMIDT (2020): Faune de France. Volume 1. Generalites - Heterogynaidae, Ampulicidae, Sphecidae, Crabronidae (1 re partie). - Federation Francaise des Societes de Sciences Naturelles, Paris. 370 pp." type="book" year="2020">BITSCH et al. (2020)</bibRefCitation>
. It is unique amoung western Palaeartcic species by the lack of the epicnemial carina (
<figureCitation id="13312A46FFD3FFAFDB440DC9081CBEBE" box="[553,612,1239,1261]" captionStart="Figs 1-6" captionStartId="7.[163,203,1258,1280]" captionTargetBox="[163,1069,208,1243]" captionTargetId="figure-51@7.[163,1069,208,1245]" captionTargetPageId="7" captionText="Figs 1-6: (1) Carinostigmus marocensis, ♁ lateral; (2) Carinostigmus marocensis, ♁ dorsal; (3) Carinostigmus marocensis, ♁ face; (4) Entomognathus brevis, ♁ dorsal; (5) Entomognathus fortuitus, ♀ dorsal; (6) Entomognathus fortuitus, ♀ mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.14681719" httpUri="https://zenodo.org/record/14681719/files/figure.png" pageId="2" pageNumber="303">fig. 6</figureCitation>
). Mesopleuron is basally therefore rounded, and also mate, compared to the shiny mesopleuron in remaining species.
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(
<figureCitation id="13312A46FFD5FFA9DA320DB009B2BE90" box="[863,970,1198,1220]" captionStart="Figs 14-20" captionStartId="9.[644,684,1262,1284]" captionTargetBox="[166,1072,202,1583]" captionTargetId="figure-1@9.[166,1072,202,1583]" captionTargetPageId="9" captionText="Figs 14-20: (14) Miscophus smaraensis, ♁ lateral; (15) Miscophus smaraensis, ♁ dorsal; (16) Miscophus smaraensis, ♁ face; (17) Miscophus smaraensis, ♁ first AS; (18) Miscophus smaraensis, ♁ S8, in dorsal view; (19) Spilomena canariensis, ♀ lateral; (20) Spilomena canariensis, ♀ face." figureDoi="http://doi.org/10.5281/zenodo.14681723" httpUri="https://zenodo.org/record/14681723/files/figure.png" pageId="4" pageNumber="305">figs 19-20</figureCitation>
,
<figureCitation id="13312A46FFD5FFA9DABB0DB00988BE97" box="[982,1008,1198,1220]" captionStart="Fig" captionStartId="10.[163,196,843,865]" captionTargetBox="[163,997,207,833]" captionTargetId="figure-33@10.[163,997,207,833]" captionTargetPageId="10" captionText="Fig. 21: Distribution of Carinostigmus marocensis (blue), Spilomena canariensis (red) and the African distribution of Entomognathus fortuitus (green)." figureDoi="http://doi.org/10.5281/zenodo.14681727" httpUri="https://zenodo.org/record/14681727/files/figure.png" pageId="4" pageNumber="305">21</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3106548FFD5FFA9D9CE0DCA0B70BF64" pageId="4" pageNumber="305" type="reference_group">
<paragraph id="8BB536C3FFD5FFA9D9CE0DCA0863BF51" blockId="4.[163,1072,1234,1335]" pageId="4" pageNumber="305">
<taxonomicName id="4C0A4D40FFD5FFA9D9CE0DCA084CBEBA" ID-CoL="4Z44B" authority="BISCHOFF, 1937 b: 2" authorityName="BISCHOFF" authorityPageNumber="2" authorityYear="1937" box="[163,564,1236,1258]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="canariensis">
<emphasis id="B97EEAD1FFD5FFA9D9CE0DCA0B16BEB9" box="[163,366,1236,1258]" italics="true" pageId="4" pageNumber="305">Spilomena canariensis</emphasis>
<materialsCitation id="3B623C9EFFD5FFA9D8150DCB084CBEBA" box="[376,564,1236,1258]" country="Spain" location="Orotava" municipality="Tenerife" pageId="4" pageNumber="305" specimenCount="2" specimenCount-female="2" stateProvince="Canarias" typeStatus="holotype">
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, 1937b: 2
</materialsCitation>
</taxonomicName>
,
<specimenCount id="9D0CFD4AFFD5FFA9DB2F0DCB082BBEB9" box="[578,595,1237,1258]" count="1" pageId="4" pageNumber="305" type="female">♀</specimenCount>
, ♁.
<typeStatus id="54B18861FFD5FFA9DBEE0DCA08A0BEB9" box="[643,728,1236,1258]" pageId="4" pageNumber="305" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D0CFD4AFFD5FFA9DB880DCB088EBEB9" box="[741,758,1237,1258]" count="1" pageId="4" pageNumber="305" type="female">♀</specimenCount>
,
<collectingCountry id="F31D7653FFD5FFA9DA680DCA0942BEB9" box="[773,826,1236,1258]" name="Spain" pageId="4" pageNumber="305">Spain</collectingCountry>
:
<collectingRegion id="49CEF821FFD5FFA9DA2A0DCA09AABEB9" box="[839,978,1236,1258]" country="Spain" name="Canarias" pageId="4" pageNumber="305">Canary Islands</collectingRegion>
:
<collectingMunicipality id="6BD1ACB9FFD5FFA9DAB20DCA0E54BEB9" box="[991,1068,1236,1258]" pageId="4" pageNumber="305">Tenerife</collectingMunicipality>
:
<location id="8ED56018FFD5FFA9D9BD0DF20B6FBF52" LSID="urn:lsid:plazi:treatment:03A387D5FFD5FFA8D9CE0DB008D4BB51:8ED56018FFD5FFA9D9BD0DF20B6FBF52" box="[208,279,1260,1281]" country="Spain" municipality="Tenerife" name="Orotava" pageId="4" pageNumber="305" stateProvince="Canarias">Orotava</location>
(Zool. Mus. Univ. Helsinki.).
</paragraph>
<paragraph id="8BB536C3FFD5FFA9D9CE0C150B70BF64" blockId="4.[163,1072,1234,1335]" pageId="4" pageNumber="305">
M a t e r i a l e x a m i n e d:
<specimenCount id="9D0CFD4AFFD5FFA9D8990C15087DBF73" box="[500,517,1291,1312]" count="1" pageId="4" pageNumber="305" type="female">♀</specimenCount>
<date id="FFB41003FFD5FFA9DB790C150801BF72" box="[532,633,1291,1313]" pageId="4" pageNumber="305" value="2024-03-21">21.iii.2024</date>
, Laâyoune-Sakia, Smara
<geoCoordinate id="EE3E5004FFD5FFA9DA110C1509B3BF72" box="[892,971,1291,1313]" degrees="26.746" direction="north" orientation="latitude" pageId="4" pageNumber="305" precision="55" value="26.746">26.746N</geoCoordinate>
<geoCoordinate id="EE3E5004FFD5FFA9DAB70C150E55BF72" box="[986,1069,1291,1313]" degrees="11.681" direction="west" orientation="longitude" pageId="4" pageNumber="305" precision="55" value="-11.681">11.681W</geoCoordinate>
(CSE).
</paragraph>
</subSubSection>
<subSubSection id="C3106548FFD5FFA8D9CE0C5808D4BB51" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305" type="description">
<paragraph id="8BB536C3FFD5FFA9D9CE0C580B5BBFE0" blockId="4.[163,1070,1349,1460]" pageId="4" pageNumber="305">
R e c o g n i t i o n: The species can be identified with
<bibRefCitation id="EF9B4B32FFD5FFA9DB890C5809E3BF08" author="DOLLFUSS H." box="[740,923,1349,1371]" pageId="4" pageNumber="305" pagination="481 - 510" refId="ref3517" refString="DOLLFUSS H. (1986): Eine Revision der Gattung Spilomena Shuckard der westlichen und zentralen palaarktischen Region (Hymenoptera, Sphecidae). - Annalen des Naturhistorischen Museum in Wien 88 - 89 B: 481 - 510." type="book chapter" year="1986">DOLLFUSS (1986)</bibRefCitation>
. The female is unique by having a distinct emargination on ACM, mandible is brown with a reddish apex, propodeal enclosure is not surrounded by a keel, and T6 has two longitudinal keels (
<figureCitation id="13312A46FFD5FFA9D9C70C800B6FBFE0" box="[170,279,1438,1460]" captionStart="Figs 14-20" captionStartId="9.[644,684,1262,1284]" captionTargetBox="[166,1072,202,1583]" captionTargetId="figure-1@9.[166,1072,202,1583]" captionTargetPageId="9" captionText="Figs 14-20: (14) Miscophus smaraensis, ♁ lateral; (15) Miscophus smaraensis, ♁ dorsal; (16) Miscophus smaraensis, ♁ face; (17) Miscophus smaraensis, ♁ first AS; (18) Miscophus smaraensis, ♁ S8, in dorsal view; (19) Spilomena canariensis, ♀ lateral; (20) Spilomena canariensis, ♀ face." figureDoi="http://doi.org/10.5281/zenodo.14681723" httpUri="https://zenodo.org/record/14681723/files/figure.png" pageId="4" pageNumber="305">fig. 19, 20</figureCitation>
).
</paragraph>
<paragraph id="8BB536C3FFD5FFA9D9CE0CDD097ABC63" blockId="4.[163,1070,1475,1585]" pageId="4" pageNumber="305">
D i s t r i b u t i o n
<taxonomicName id="4C0A4D40FFD5FFA9D8E10CDD080CBF8B" authorityName="BISCHOFF" authorityYear="1937" box="[396,628,1475,1496]" class="Insecta" family="Crabronidae" genus="Spilomena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="canariensis">
<emphasis id="B97EEAD1FFD5FFA9D8E10CDD080CBF8B" box="[396,628,1475,1496]" italics="true" pageId="4" pageNumber="305">Spilomena canariensis</emphasis>
</taxonomicName>
was previously considered endemic to the Canary Islands (
<bibRefCitation id="EF9B4B32FFD5FFA9D8220CFF0B87BFA5" author="DOLLFUSS H." box="[335,511,1504,1526]" pageId="4" pageNumber="305" pagination="481 - 510" refId="ref3517" refString="DOLLFUSS H. (1986): Eine Revision der Gattung Spilomena Shuckard der westlichen und zentralen palaarktischen Region (Hymenoptera, Sphecidae). - Annalen des Naturhistorischen Museum in Wien 88 - 89 B: 481 - 510." type="book chapter" year="1986">DOLLFUSS, 1986</bibRefCitation>
). It has now also been discovered on Madeira (
<bibRefCitation id="EF9B4B32FFD5FFA9DA960CFF0AA2BC40" author="SMIT J." pageId="4" pageNumber="305" pagination="165 - 180" refId="ref3896" refString="SMIT J. (2000): The wasps of Madeira (Hymenoptera: Chrysididae, Pompilidae, Vespidae, Sphecidae). - Entomofauna 21: 165 - 180." type="journal article" year="2000">SMIT 2000</bibRefCitation>
). This is the first record for
<collectingCountry id="F31D7653FFD5FFA9D8940CE0082BBC40" box="[505,595,1534,1555]" name="Morocco" pageId="4" pageNumber="305">Marocco</collectingCountry>
and thus for the African mainland (
<figureCitation id="13312A46FFD5FFA9DAD50CE00983BC40" box="[952,1019,1534,1556]" captionStart="Fig" captionStartId="10.[163,196,843,865]" captionTargetBox="[163,997,207,833]" captionTargetId="figure-33@10.[163,997,207,833]" captionTargetPageId="10" captionText="Fig. 21: Distribution of Carinostigmus marocensis (blue), Spilomena canariensis (red) and the African distribution of Entomognathus fortuitus (green)." figureDoi="http://doi.org/10.5281/zenodo.14681727" httpUri="https://zenodo.org/record/14681727/files/figure.png" pageId="4" pageNumber="305">fig. 21</figureCitation>
). For detailed records of Canary Islands see
<bibRefCitation id="EF9B4B32FFD5FFA9DB780F050887BC63" author="HOHMANN H. &amp; LA ROCHE F. &amp; ORTEGA G. &amp; J. BARQUIN" box="[533,767,1563,1585]" pageId="4" pageNumber="305" pagination="493 - 894" refId="ref3558" refString="HOHMANN H., LA ROCHE F., ORTEGA G. &amp; J. BARQUIN (1993): Bienen, Wespen und Ameisen der Kanarischen Inseln - Abejas, avispas y hormigas de las Islas Canarias - Bees, wasps and ants of the Canary Islands (Insecta: Hymenoptera: Aculeata). - Veroffentlichungen aus dem Ubersee-Museum Bremen. Naturwissenschaften 12, vol. 1: 1 - 465, XII pls., vol. 2: 493 - 894." type="journal article" year="1993">HOHMANN et al. (1993)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB536C3FFD4FFA8D9CE09CE08D4BB51" blockId="5.[163,1070,207,259]" pageId="5" pageNumber="306">
H a b i t a t: The specimen was caught on a flowering
<emphasis id="B97EEAD1FFD4FFA8DB9E09CE0931BAB6" box="[755,841,208,229]" italics="true" pageId="5" pageNumber="306">Tamarix</emphasis>
shrub, at the western border of the town, in a semiarid and very dry area.
</paragraph>
</subSubSection>
</treatment>
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500
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<emphasis id="1BD53B019348FFA4FF24A5F5FEFCF0ED" bold="true" box="[163,260,385,406]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FE94A5F4FE62F0ED" bold="true" box="[275,410,384,406]" italics="true" pageId="2" pageNumber="367">Leptolambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE2FA5F5FE08F0ED" bold="true" box="[424,496,385,406]" italics="true" pageId="2" pageNumber="367">natrun</emphasis>
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?
<taxonomicName id="EEA19C909348FFA4FF34A5D2FDFFF0C7" authority="SCHAUM, 1857" authorityName="SCHAUM" authorityYear="1857" box="[179,519,422,444]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
<emphasis id="1BD53B019348FFA4FF34A5D2FE8CF0C0" box="[179,372,422,443]" italics="true" pageId="2" pageNumber="367">Hydroporus lernaeus</emphasis>
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, 1857
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;
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<bibRefCitation id="4D309AE29348FFA4FD97A5D2FD57F0C7" author="SHARP D." box="[528,687,422,444]" pageId="2" pageNumber="367" pagination="179 - 1003" refId="ref6342" refString="SHARP D. (1882): On aquatic carnivorous Coleoptera or Dytiscidae. - Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">
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1882: 405
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(partim).
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?
<emphasis id="1BD53B019348FFA4FF3EA5B4FE20F0AE" box="[185,472,448,469]" italics="true" pageId="2" pageNumber="367">Coelambus parallellogrammus</emphasis>
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<emphasis id="1BD53B019348FFA4FE63A5B4FDCAF0AE" box="[484,562,448,469]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FDC2A5B4FD22F0AD" author="SCHAUM H." box="[581,730,448,470]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FDC2A5B4FD6FF0AE" baselines="464,465" box="[581,663,448,469]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
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)
</taxonomicName>
;
<smallCapsWord id="2FF871CF9348FFA4FD77A5B5FCD6F0AF" baselines="465,464" box="[752,814,449,470]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Bedel" pageId="2" pageNumber="367">BEDEL</smallCapsWord>
1895-1925: 339, footnote (partim).
</paragraph>
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<treatmentCitationGroup id="09B1C03D9348FFA4FF24A586FF0DF34C" pageId="2" pageNumber="367">
<emphasis id="1BD53B019348FFA4FF24A586FEF1F37C" box="[163,265,498,519]" italics="true" pageId="2" pageNumber="367">Coelambus</emphasis>
<taxonomicName id="EEA19C909348FFA4FE96A586FDFFF373" authority="(SCHAUM, 1857)" baseAuthorityName="SCHAUM" baseAuthorityYear="1857" box="[273,519,498,520]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
<emphasis id="1BD53B019348FFA4FE96A586FEA7F37C" box="[273,351,498,519]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FEE8A586FDF9F373" author="SCHAUM H." box="[367,513,498,520]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FEE8A586FE39F37C" baselines="514,515" box="[367,449,498,519]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="A800C1029348FFA4FD94A587FD18F373" author="REGIMBART M." box="[531,736,498,520]" page="35" pageId="2" pageNumber="367" year="1895">
<bibRefCitation id="4D309AE29348FFA4FD94A587FD18F373" author="REGIMBART M." box="[531,736,498,520]" pageId="2" pageNumber="367" pagination="1 - 244" refId="ref6276" refString="REGIMBART M. (1895): Revision des Dytiscidae et Gyrinidae d'Afrique et Madagascar et iles voisines. - Memoires de la Societe Entomologiques de Belgique 4: 1 - 244." type="journal article" year="1895">
<smallCapsWord id="2FF871CF9348FFA4FD94A587FD7CF37C" baselines="515,514" box="[531,644,498,520]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Regimbart" pageId="2" pageNumber="367">RÉGIMBART</smallCapsWord>
1895: 35
</bibRefCitation>
</treatmentCitation>
(partim);
<treatmentCitation id="A800C1029348FFA4FCC6A587FBD6F373" author="ZIMMERMANN A." box="[833,1070,498,520]" page="151" pageId="2" pageNumber="367" year="1919">
<bibRefCitation id="4D309AE29348FFA4FCC6A587FBD6F373" author="ZIMMERMANN A." box="[833,1070,498,520]" pageId="2" pageNumber="367" pagination="68 - 249" refId="ref6528" refString="ZIMMERMANN A. (1919): Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem. - Archiv fur Naturgeschichte 83 (1917) (A 12): 68 - 249." type="journal article" year="1919">
<smallCapsWord id="2FF871CF9348FFA4FCC6A587FC3FF37C" baselines="515,515" box="[833,967,499,520]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zimmermann" pageId="2" pageNumber="367">ZIMMERMANN</smallCapsWord>
1919: 151
</bibRefCitation>
</treatmentCitation>
(partim); 1920: 68 (partim); 1930: 101 (partim);
<treatmentCitation id="A800C1029348FFA4FD14A67EFCB5F364" author="GUIGNOT F." box="[659,845,522,544]" page="332" pageId="2" pageNumber="367" year="1959">
<bibRefCitation id="4D309AE29348FFA4FD14A67EFCB5F364" author="GUIGNOT F." box="[659,845,522,544]" pageId="2" pageNumber="367" pagination="323 - 648" refId="ref5985" refString="GUIGNOT F. (1959): Revision des hydrocanthares d'Afrique (Coleoptera Dytiscoidea). 2. - Annales du Musee Royal du Congo Belge Serie 8 vo (Sciences Zoologiques) 78: 323 - 648." type="journal article" year="1959">
<smallCapsWord id="2FF871CF9348FFA4FD14A67EFD13F364" baselines="538,539" box="[659,747,522,543]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Guignot" pageId="2" pageNumber="367">GUIGNOT</smallCapsWord>
1959: 332
</bibRefCitation>
</treatmentCitation>
(partim);
<treatmentCitation id="A800C1029348FFA4FC2DA67EFF09F34C" author="ALFIERI A." page="33" pageId="2" pageNumber="367" year="1976">
<bibRefCitation id="4D309AE29348FFA4FC2DA67EFF09F34C" author="ALFIERI A." pageId="2" pageNumber="367" pagination="1 - 361" refId="ref5595" refString="ALFIERI A. (1976): The Coleoptera of Egypt. - Memoires de la Societe Entomologique d'Egypte 5: i-xvi, 1 - 361." type="journal article" year="1976">
<smallCapsWord id="2FF871CF9348FFA4FC2DA67EFC0CF365" baselines="538,538" box="[938,1012,522,543]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Alfieri" pageId="2" pageNumber="367">ALFIERI</smallCapsWord>
1976: 33
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="291EE7139348FFA4FF24A649FE2DF311" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
<treatmentCitationGroup id="09B1C03D9348FFA4FF24A649FE2DF311" pageId="2" pageNumber="367">
<taxonomicName id="EEA19C909348FFA4FF24A649FD82F329" ID-CoL="8LZGZ" authority="(SCHAUM, 1857)" baseAuthorityName="SCHAUM" baseAuthorityYear="1857" box="[163,634,572,594]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Coelambus">
<emphasis id="1BD53B019348FFA4FF24A649FF00F329" box="[163,248,573,594]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FE8EA648FE97F32A" box="[265,367,572,593]" italics="true" pageId="2" pageNumber="367">Coelambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE06A648FE37F32A" box="[385,463,572,593]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FE67A648FD8BF329" author="SCHAUM H." box="[480,627,572,594]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FE67A648FDCAF32A" baselines="588,589" box="[480,562,572,593]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="A800C1029348FFA4FD0FA649FC9CF329" author="ZALAT S. &amp; SALEH AHMED R. &amp; ANGUS R. B." box="[648,868,572,594]" page="26" pageId="2" pageNumber="367" year="2000">
<bibRefCitation id="4D309AE29348FFA4FD0FA649FC9CF329" author="ZALAT S. &amp; SALEH AHMED R. &amp; ANGUS R. B." box="[648,868,572,594]" pageId="2" pageNumber="367" pagination="1 - 107" refId="ref6482" refString="ZALAT S., SALEH AHMED R., ANGUS R. B. &amp; A. KASCHEF (2000): Diving beetles (Coleoptera: Dytiscidae and Noteridae) of Egypt. - Egyptian Journal of Natural History 2: 1 - 107." type="journal article" year="2000">
<smallCapsWord id="2FF871CF9348FFA4FD0FA649FD30F32A" baselines="589,589" box="[648,712,573,594]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zalat" pageId="2" pageNumber="367">ZALAT</smallCapsWord>
et al. 2000: 26
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="A800C1029348FFA4FCF5A649FBD5F32A" author="NILSSON A. N." box="[882,1069,572,594]" page="207" pageId="2" pageNumber="367" year="2001">
<bibRefCitation id="4D309AE29348FFA4FCF5A649FBD5F32A" author="NILSSON A. N." box="[882,1069,572,594]" pageId="2" pageNumber="367" pagination="1 - 395" refId="ref6087" refString="NILSSON A. N. (2001): Dytiscidae (Coleoptera). - In: World Catalogue of insects 3: 1 - 395." type="journal article" year="2001">
<smallCapsWord id="2FF871CF9348FFA4FCF5A649FC3BF32A" baselines="589,589" box="[882,963,573,594]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Nilsson" pageId="2" pageNumber="367">NILSSON</smallCapsWord>
2001: 207
</bibRefCitation>
</treatmentCitation>
(partim); 2003: 72 (partim).
</treatmentCitationGroup>
</paragraph>
<paragraph id="291EE7139348FFA4FF24A61BFEA8F3E7" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
<taxonomicName id="EEA19C909348FFA4FF24A61BFD81F3FF" ID-CoL="8LZGZ" authority="(SCHAUM, 1857)" baseAuthorityName="SCHAUM" baseAuthorityYear="1857" box="[163,633,622,644]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
<emphasis id="1BD53B019348FFA4FF24A61BFF00F3FF" box="[163,248,623,644]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FE82A61AFE82F3F8" box="[261,378,622,643]" italics="true" pageId="2" pageNumber="367">Leptolambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE0FA61AFE2EF3F8" box="[392,470,622,643]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FE64A61BFD8BF3FF" author="SCHAUM H." box="[483,627,622,644]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FE64A61BFDCDF3F8" baselines="639,639" box="[483,565,623,644]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
)
</taxonomicName>
;
<smallCapsWord id="2FF871CF9348FFA4FD04A61BFD2CF3F8" baselines="639,639" box="[643,724,623,644]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Nilsson" pageId="2" pageNumber="367">NILSSON</smallCapsWord>
&amp;
<smallCapsWord id="2FF871CF9348FFA4FD74A61BFCC9F3F8" baselines="639,638" box="[755,817,622,644]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hajek" pageId="2" pageNumber="367">HÁJEK</smallCapsWord>
2024a: 46 (partim); 2024b: 197 (partim).
</paragraph>
<paragraph id="291EE7139348FFA4FF24A6D5FD9AF39D" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
T y p e l o c a l i t y: Wadi El Natrun (also Wadi Natrun, Wadi an-Natrun, Wadi Natroun), ca.
<quantity id="EE594AF69348FFA4FF5FA6CDFEE7F3B5" box="[216,287,696,718]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="367" unit="km" value="100.0">100 km</quantity>
NW
<collectingRegion id="EB6529F19348FFA4FED3A6CCFE71F3B5" box="[340,393,696,718]" country="Egypt" name="Al Qahirah" pageId="2" pageNumber="367">Cairo</collectingRegion>
, ca.
<geoCoordinate id="4C9581D49348FFA4FE32A6CDFE01F3B5" box="[437,505,697,718]" degrees="30.41" direction="north" orientation="latitude" pageId="2" pageNumber="367" precision="555" value="30.41">30.41N</geoCoordinate>
<geoCoordinate id="4C9581D49348FFA4FD87A6CDFDBCF3B5" box="[512,580,697,718]" degrees="30.30" direction="east" orientation="longitude" pageId="2" pageNumber="367" precision="555" value="30.3">30.30E</geoCoordinate>
. N o t e s: This wadi is about
<quantity id="EE594AF69348FFA4FCE0A6CCFC6FF3B5" box="[871,919,696,718]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="367" unit="m" value="25.0">25 m</quantity>
below sea level and about
<quantity id="EE594AF69348FFA4FEB1A6A5FE9DF39D" box="[310,357,721,742]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="367" unit="m" value="38.0">38 m</quantity>
below the level of the Nile.
</paragraph>
</subSubSection>
<subSubSection id="61BBB4989348FFA2FF24A685FC2EF778" lastPageId="4" lastPageNumber="369" pageId="2" pageNumber="367" type="description">
<paragraph id="291EE7139348FFA4FF24A685FDE5F575" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
<materialsCitation id="99C9ED4E9348FFA4FF24A685FC39F265" collectorName="Coll. Alfieri &amp; Coll. F. Guignot" country="Egypt" location="Wadi Natroun" pageId="2" pageNumber="367" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
M a t e r i a l e x a m i n e d:
<typeStatus id="F61A59B19348FFA4FE6FA684FDB8F27D" box="[488,576,752,774]" pageId="2" pageNumber="367" type="holotype">Holotype</typeStatus>
:
<specimenCount id="3FA72C9A9348FFA4FDCBA685FDA5F27D" box="[588,605,753,774]" count="1" pageId="2" pageNumber="367" type="female">♀</specimenCount>
, &quot;
<location id="2C7EB1C89348FFA4FDF6A684FD0FF27D" LSID="urn:lsid:plazi:treatment:A10856059348FFA2FF24A5F5FC2EF778:2C7EB1C89348FFA4FDF6A684FD0FF27D" box="[625,759,752,774]" country="Egypt" name="Wadi Natroun" pageId="2" pageNumber="367">Wadi Natroun</location>
, 2.5.[19]27&quot; [hw Guignot], &quot;
<collectorName id="845482C59348FFA4FB87A684FEF4F265" pageId="2" pageNumber="367">Coll. Alfieri</collectorName>
,
<collectingCountry id="51B6A7839348FFA4FE9FA77DFEA1F265" box="[280,345,777,798]" name="Egypt" pageId="2" pageNumber="367">Egypte</collectingCountry>
&quot; [printed], &quot;
<specimenCount id="3FA72C9A9348FFA4FE4AA77DFE25F265" box="[461,477,777,798]" count="1" pageId="2" pageNumber="367" type="female">♀</specimenCount>
&quot;, &quot;Muséum Paris, 1960,
<collectorName id="845482C59348FFA4FD4DA77CFC9CF265" box="[714,868,776,798]" pageId="2" pageNumber="367">Coll. F. Guignot</collectorName>
&quot; [printed]
</materialsCitation>
;
<materialsCitation id="99C9ED4E9348FFA4FC49A77CFBD2F235" collectionCode="MNHN" collectorName="Guignot &amp; Coelambus" country="Egypt" determinerName="H. Fery" pageId="2" pageNumber="367" specimenCount="1" typeStatus="holotype">
&quot;
<typeStatus id="F61A59B19348FFA4FC53A77CFBD2F265" box="[980,1066,776,798]" pageId="2" pageNumber="367" type="holotype">Holotype</typeStatus>
,
<taxonomicName id="EEA19C909348FFA4FF57A755FE1EF24D" authorityName="Fery" authorityYear="2024" box="[208,486,800,822]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="natrun" status="sp. nov." subGenus="Leptolambus">
<emphasis id="1BD53B019348FFA4FF57A755FED9F24D" box="[208,289,801,822]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FEA9A754FE67F24E" box="[302,415,800,821]" italics="true" pageId="2" pageNumber="367">Leptolambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE2BA755FE1EF24D" box="[428,486,801,822]" italics="true" pageId="2" pageNumber="367">natrun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E6867A9348FFA4FE6AA755FDD6F24D" box="[493,558,801,822]" pageId="2" pageNumber="367" rank="species">nov.sp.</taxonomicNameLabel>
,
<determinerName id="DCC20BD79348FFA4FDB0A755FD82F24D" box="[567,634,801,822]" pageId="2" pageNumber="367">H. Fery</determinerName>
det. 2024&quot; [red, printed] (
<collectionCode id="4FB07FD69348FFA4FCE7A755FC5EF24D" box="[864,934,801,822]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="367">MNHN</collectionCode>
) [for labels see
<figureCitation id="B19AFB969348FFA4FF57A74CFEFDF235" box="[208,261,824,846]" captionStart="Fig" captionStartId="9.[163,195,1508,1530]" captionTargetBox="[164,1067,201,1493]" captionTargetId="figure-23@9.[164,1067,201,1493]" captionTargetPageId="9" captionText="Fig. 1: Habitus of Hygrotus (Leptolambus) natrun nov.sp. and labels (holotype from Wadi El Natrun, Egypt)." figureDoi="http://doi.org/10.5281/zenodo.14681677" httpUri="https://zenodo.org/record/14681677/files/figure.png" pageId="2" pageNumber="367">Fig. 1</figureCitation>
; specimen until now kept in coll.
<collectorName id="845482C59348FFA4FDB1A74CFD86F235" box="[566,638,824,846]" pageId="2" pageNumber="367">Guignot</collectorName>
under
<emphasis id="1BD53B019348FFA4FD3AA74CFCD8F236" box="[701,800,824,845]" italics="true" pageId="2" pageNumber="367">
<collectorName id="845482C59348FFA4FD3AA74CFCD8F236" box="[701,800,824,845]" pageId="2" pageNumber="367">Coelambus</collectorName>
</emphasis>
<taxonomicName id="EEA19C909348FFA4FCA0A74CFBD9F236" box="[807,1057,824,846]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="variety" species="lernaeus" variety="orthogrammus">
<emphasis id="1BD53B019348FFA4FCA0A74CFC8AF236" box="[807,882,824,845]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
var.
<emphasis id="1BD53B019348FFA4FC27A74CFBD9F236" box="[928,1057,824,845]" italics="true" pageId="2" pageNumber="367">orthogrammus</emphasis>
</taxonomicName>
]
</materialsCitation>
.
<materialsCitation id="99C9ED4E9348FFA4FF57A725FDE1F575" collectingDate="1982-09-27" collectingDateMax="1996-04-30" collectingDateMin="1982-09-27" collectionCode="NHMW, CHF, OLML, CLH, ZSM" collectorName="Coll. A. Alfieri &amp; Coll. Gschwendtner &amp; W Desert Oasis Bahariva &amp; W. G. Ullrich &amp; Zimmermann &amp; Samml. A. Zimmermann &amp; Coelambus &amp; Fery &amp; All" country="Egypt" determinerName="Gschwendtner &amp; Fery" latitude="28.33" location="Desert Oasis" longLatPrecision="740" longitude="29.0" municipality="Bahariva Oases" pageId="2" pageNumber="367" specimenCount="2" specimenCount-female="2" stateProvince="Al Qahirah" typeStatus="paratype">
<typeStatus id="F61A59B19348FFA4FF57A725FED4F21D" box="[208,300,849,870]" pageId="2" pageNumber="367" type="paratype">Paratypes</typeStatus>
: 2♁♁, &quot;Ägypten
<date id="5D1FC1D39348FFA4FE42A725FDE6F21D" box="[453,542,849,870]" pageId="2" pageNumber="367" value="1982-09-27">
<collectingDate id="4D5B383B9348FFA4FE42A725FDE6F21D" box="[453,542,849,870]" pageId="2" pageNumber="367" value="1982-09-27">27.9.1982</collectingDate>
</date>
, Wadi Natrun, Kafr Daoud, Sattm.&quot; [hw?] (
<collectionCode id="4FB07FD69348FFA4FC26A725FC15F21D" box="[929,1005,849,870]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="367" type="Museum">NHMW</collectionCode>
,
<collectionCode id="4FB07FD69348FFA4FC71A725FBDDF21D" box="[1014,1061,849,870]" pageId="2" pageNumber="367">CHF</collectionCode>
). 2♁♁, &quot;
<collectingMunicipality id="C97A7D699348FFA4FE89A71CFE64F205" box="[270,412,872,894]" pageId="2" pageNumber="367">Bahariva Oases</collectingMunicipality>
, 20.–24.3 [19]25&quot; [printed], &quot;
<collectingCountry id="51B6A7839348FFA4FD27A71DFD21F205" box="[672,729,873,894]" name="Egypt" pageId="2" pageNumber="367">Egypt</collectingCountry>
, Min. Agric. (
<collectingCountry id="51B6A7839348FFA4FCD1A71DFC76F205" box="[854,910,873,894]" name="Egypt" pageId="2" pageNumber="367">Egypt</collectingCountry>
),
<collectorName id="845482C59348FFA4FC19A71CFBD0F205" box="[926,1064,872,894]" pageId="2" pageNumber="367">Coll. A. Alfieri</collectorName>
&quot; [printed], &quot;
<collectorName id="845482C59348FFA4FEA9A7F4FE1BF2ED" box="[302,483,896,918]" pageId="2" pageNumber="367">Coll. Gschwendtner</collectorName>
&quot; [printed],
<taxonomicName id="EEA19C909348FFA4FDC0A7F5FD0EF2ED" authority="SCHAUM." authorityName="SCHAUM." box="[583,758,896,918]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
&quot;
<emphasis id="1BD53B019348FFA4FDD7A7F4FD63F2EE" box="[592,667,896,917]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
<smallCapsWord id="2FF871CF9348FFA4FD26A7F5FD09F2EE" baselines="913,913" box="[673,753,897,918]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
.
</taxonomicName>
, det.
<determinerName id="DCC20BD79348FFA4FCAFA7F4FC51F2ED" box="[808,937,896,918]" pageId="2" pageNumber="367">Gschwendtner</determinerName>
&quot; [first line hw, second line printed] (
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).
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, &quot;
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W
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Bahariva
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[= Bahariya, ca.
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, ca.
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SW
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],
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<collectingDate id="4D5B383B9348FFA4FD87A7C4FD9CF2BD" box="[512,612,944,966]" pageId="2" pageNumber="367" value="1996-04-30">30.IV.1996</collectingDate>
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<collectorName id="845482C59348FFA4FDEAA7C5FD1FF2BD" box="[621,743,944,966]" pageId="2" pageNumber="367">W.G. Ullrich</collectorName>
&quot; [printed] (
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).
<specimenCount id="3FA72C9A9348FFA4FC13A7C5FC48F2BD" box="[916,944,945,966]" count="1" pageId="2" pageNumber="367" type="female">1♀</specimenCount>
, &quot;
<collectingRegion id="EB6529F19348FFA4FC42A7C4FC04F2BD" box="[965,1020,944,966]" country="Egypt" name="Al Qahirah" pageId="2" pageNumber="367">Kairo</collectingRegion>
&quot; [hw
<collectorName id="845482C59348FFA4FF57A7BCFEB4F2A5" box="[208,332,968,990]" pageId="2" pageNumber="367">Zimmermann</collectorName>
], &quot;
<collectorName id="845482C59348FFA4FEE7A7BCFDBCF2A5" box="[352,580,968,990]" pageId="2" pageNumber="367">Samml. A. Zimmermann</collectorName>
&quot; [printed], &quot;
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<collectorName id="845482C59348FFA4FD37A7BCFCEAF2A6" box="[688,786,968,989]" pageId="2" pageNumber="367">Coelambus</collectorName>
</emphasis>
,
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<emphasis id="1BD53B019348FFA4FC98A7BCFC92F2A6" box="[799,874,968,989]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
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)
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,
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det. 19[91] [hw
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], &quot;SNSB-ZSM
<taxonomicName id="EEA19C909348FFA4FD87A794FD4BF28D" authority=", ID No." authorityName="ID No." box="[512,691,992,1014]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="order">Coleoptera, ID No.</taxonomicName>
000124&quot; [printed] (
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).
<collectorName id="845482C59348FFA4FC33A794FC28F28D" box="[948,976,992,1014]" pageId="2" pageNumber="367">All</collectorName>
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provided with the respective red label
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.
</paragraph>
<paragraph id="291EE7139348FFA4FF25A069FD71F549" blockId="2.[162,649,1053,1075]" box="[162,649,1053,1075]" pageId="2" pageNumber="367">D e s c r i p t i o n o f t h e h o l o t y p e:</paragraph>
<paragraph id="291EE7139348FFA4FF24A037FEA5F5B6" blockId="2.[163,1070,1090,1230]" pageId="2" pageNumber="367">
H a b i t u s elongate oval, sides evenly rounded (
<figureCitation id="B19AFB969348FFA4FD3FA036FD0DF52C" box="[696,757,1090,1112]" captionStart="Fig" captionStartId="9.[163,195,1508,1530]" captionTargetBox="[164,1067,201,1493]" captionTargetId="figure-23@9.[164,1067,201,1493]" captionTargetPageId="9" captionText="Fig. 1: Habitus of Hygrotus (Leptolambus) natrun nov.sp. and labels (holotype from Wadi El Natrun, Egypt)." figureDoi="http://doi.org/10.5281/zenodo.14681677" httpUri="https://zenodo.org/record/14681677/files/figure.png" pageId="2" pageNumber="367">Fig. 1</figureCitation>
), body outline in dorsal view without distinct discontinuity at shoulders. Maximum width near midlength of body. Elytra brownish yellow, with blackish longitudinal vittae. Head and pronotum more reddish yellow, with brownish spots. Entire dorsal surface shiny although weakly reticulated in part.
</paragraph>
<paragraph id="291EE7139348FFA5FF24A0A9FE8DF078" blockId="2.[163,1071,1245,1560]" lastBlockId="3.[163,1069,208,259]" lastPageId="3" lastPageNumber="368" pageId="2" pageNumber="367">H e a d: Anterior margin medially almost straight, but appearing semicircular if observed in perpendicular view on entire body of specimens because anterior margin not visible; anteriorly between eyes with two clypeal grooves. Whole surface finely reticulated, but not matt. Third and fourth antennomeres equal in length, thinner and shorter than first and second as well as rest of them. Punctation on clypeus fine and sparse, diameter of punctures that of one mesh, distance between punctures that of about 5 meshes; on frons punctures larger, diameter that of about three meshes, distance that of about three meshes; on vertex punctures again smaller and sparser; in clypeal groves punctures very dense; line of coarse punctures alongside inner margin of eyes. Clypeus with two ill-defined, brownish interocular spots; above insertions of antennae (between clypeal grooves and anterior margin) shining through brownish; vertex dark brownish. Antennae reddish-yellowish with antennomers progressively darkened distally, beginning with fifth.</paragraph>
<paragraph id="291EE7139349FFA5FF24A567FDEDF368" blockId="3.[163,1093,274,531]" pageId="3" pageNumber="368">P r o n o t u m: Sides slightly rounded, with distinct brownish rim; posterior angles more or less right-angled. Punctation on disc rather fine (as on clypeus), becoming coarser laterally; centre of disc with one small longitudinal puncture; directly behind anterior margin punctation very fine and sparse, but then before disc very coarse; before posterior margin again very coarse; near lateral margins punctation much denser. Few coarser punctures near margin with extremely short seta. Surface of pronotum smooth and shiny, but directly behind anterior margin distinctly reticulated and matt. Disc with one large ill-defined, brownish spot; anterior margin appearing dark brownish because dark vertex of head shining through.</paragraph>
<paragraph id="291EE7139349FFA5FF24A657FD66F550" blockId="3.[162,1070,546,1067]" pageId="3" pageNumber="368">E l y t r a: In dorsal view lateral margin of elytron visible only near elytral base and before apex because else sides of elytra turned inwards. Epipleural rim in lateral view weakly ascending towards humeral angle. Surface smooth in anterior two thirds, behind reticulated, but not distinctly matt. Punctation double; coarse punctures interspersed with small punctures; coarse punctures progressively becoming more numerous posteriad, but near apex punctures smaller. Each elytron in anterior half with two indistinct puncture lines, which coincide more or less with second and fourth vittae; in anterior quarter between fifth vitta and elytral margin some larger punctures indicating a third line; no sutural line perceptible. Most coarser punctures with very short seta, almost imperceptible on disc, otherwise slightly longer; in posterior half a very few punctures with a longer seta; however, generally setation only well visible if adequately illuminated. Each elytron with suture, four discal vittae, a lateral and a submarginal vitta blackish; elytral base medially with blackish stripe. Second vitta reaching base, fourth vitta ending just before base, first and third vittae ending far before base; lateral vitta short, only present in anterior half, and ending far before base; submarginal vitta present only in posterior half, more or less parallel to elytral margin, reaching until apex, and here fused with sutural line. First, second and third vittae fused just before apex, third and fourth vittae short behind midlength of elytra.</paragraph>
<paragraph id="291EE7139349FFA2FF24A04FFCE0F370" blockId="3.[163,1071,1082,1574]" lastBlockId="4.[163,1070,208,523]" lastPageId="4" lastPageNumber="369" pageId="3" pageNumber="368">
V e n t r a l s u r f a c e: Head, prosternum, epipleura and legs light brownish, metaventrite and abdomen predominantly black; metacoxal processes apically, neighbouring small parts of metacoxal plates, first abdominal ventrite, hind margins of third to fifth abdominal ventrites and sixth abdominal ventrite apically shining through brownish. Almost entire venter reticulated, only slightly shining, but not distinctly matt; metaventrite medially without reticulation and here more shiny; small brownish parts of metacoxal plates neighbouring apex of metacoxal processes smooth and shiny. Punctation on metacoxal plates and metaventrite distinct; on metaventrite punctures somewhat larger; punctures absent in small posterior part of metacoxal plates and on areas right and left of medial part of metaventrite; medial part of metaventrite with irregular lines of small punctures; metacoxal processes with sparse small punctures. Abdominal ventrites generally punctured; punctures progressively becoming smaller to last ventrite; second and third ventrites medially fused and here with only very few small punctures. Third to fifth ventrites medially with few densely arranged punctures, these with long and backwardly directed setae. Prosternal process (
<figureCitation id="B19AFB969349FFA5FCE6A1A1FC50F491" box="[865,936,1493,1515]" captionStart="Figs 2-3" captionStartId="10.[163,203,1041,1063]" captionTargetBox="[161,1066,197,1027]" captionTargetId="figure-29@10.[161,1066,197,1027]" captionTargetPageId="10" captionText="Figs 2-3: Prosternal process. (2) Hygrotus (Leptolambus) natrun nov.sp.; (3) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681679" httpUri="https://zenodo.org/record/14681679/files/figure.png" pageId="3" pageNumber="368">Fig. 2</figureCitation>
) lanceolate, longitudinally carinate, with tectiform cross section; apex rounded, sides slightly flattened and here with some punctures, lateral margin rimmed (for comparison see flat process of
<emphasis id="1BD53B01934EFFA2FE92A4A4FEDFF19E" box="[277,295,208,229]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FEB1A4A4FE01F19E" baseAuthorityName="SHARP" baseAuthorityYear="1882" box="[310,505,208,229]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FEB8A4A4FEB5F19E" box="[319,333,208,229]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FEE3A4A4FE01F19E" box="[356,505,208,229]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
in
<figureCitation id="B19AFB96934EFFA2FD99A4A4FDA5F19E" box="[542,605,208,230]" captionStart="Figs 2-3" captionStartId="10.[163,203,1041,1063]" captionTargetBox="[161,1066,197,1027]" captionTargetId="figure-29@10.[161,1066,197,1027]" captionTargetPageId="10" captionText="Figs 2-3: Prosternal process. (2) Hygrotus (Leptolambus) natrun nov.sp.; (3) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681679" httpUri="https://zenodo.org/record/14681679/files/figure.png" pageId="4" pageNumber="369">Fig. 3</figureCitation>
); prosternal column coarsely punctured with small transverse elevation between anterior coxae; medial part of prosternum (before column) vaulted and very coarsely sculptured. Epipleura in anterior third more or less as broad as apex of mesotibia, with some punctures and extremely weak reticulation; evenly narrowed behind; humeral part of epipleura with oblique carina delimiting genicular fossa; carina forming with epipleural rim an angle of about 25° (see details in
<bibRefCitation id="4D309AE2934EFFA2FF24A5F4FE36F0EE" author="VILLASTRIGO A. &amp; RIBERA I. &amp; MANUEL M. &amp; MILLAN A." box="[163,462,384,406]" pageId="4" pageNumber="369" pagination="499 - 529" refId="ref6430" refString="VILLASTRIGO A., RIBERA I., MANUEL M., MILLAN A. &amp; H. FERY (2017): A new classification of the tribe Hygrotini PORTEVIN, 1929 (Coleoptera: Dytiscidae: Hydroporinae). - Zootaxa 4317 (3): 499 - 529." type="journal article" year="2017">VILLASTRIGO et al. (2017))</bibRefCitation>
. Hind margins of metacoxal processes slightly slanted, interlaminary bridge well exposed, forming a more or less triangular (or wing-like shaped) elevation on first and second abdominal ventrites (see details in FERY &amp; RIBERA (2018)). Metacoxal lines diverging forwards, ending before hind margin of metaventrite. Protarsal claws thin, evenly curved and more or less identical.
</paragraph>
<paragraph id="291EE713934EFFA2FF24A66EFD82F3B9" blockId="4.[163,1070,538,707]" pageId="4" pageNumber="369">
G e n i t a l i a: Gonocoxae in ventral view as in
<figureCitation id="B19AFB96934EFFA2FD1BA66EFD21F354" box="[668,729,538,560]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 4</figureCitation>
; in lateral view (
<figureCitation id="B19AFB96934EFFA2FC03A66EFC39F34B" box="[900,961,538,560]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 5</figureCitation>
) distinctly flatter than in
<emphasis id="1BD53B01934EFFA2FEB2A64CFEB6F336" box="[309,334,568,589]" italics="true" pageId="4" pageNumber="369">H.</emphasis>
<taxonomicName id="EEA19C90934EFFA2FED2A64CFE21F336" box="[341,473,568,589]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FED9A64CFE94F336" box="[350,364,568,589]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FE05A64CFE21F336" box="[386,473,568,589]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FE6FA64CFDDFF336" box="[488,551,568,590]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 8</figureCitation>
); proximal portion less curved upwards than in
<emphasis id="1BD53B01934EFFA2FB92A64CFBDFF336" box="[1045,1063,568,589]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FF24A621FED1F311" box="[163,297,597,619]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FF2CA622FF41F310" box="[171,185,598,619]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FF55A621FED1F311" box="[210,297,597,618]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FEBDA621FE63F311" box="[314,411,597,619]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Figs 7, 8</figureCitation>
), but more than in
<emphasis id="1BD53B01934EFFA2FDEAA622FD87F310" box="[621,639,598,619]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FD08A621FCABF311" baseAuthorityName="SHARP" baseAuthorityYear="1882" box="[655,851,597,619]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FD1FA622FD5EF310" box="[664,678,598,619]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FD39A621FCABF311" box="[702,851,597,618]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FCE3A621FC27F311" box="[868,991,597,619]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Figs 10, 11</figureCitation>
); distal margins slightly concavely emarginate (see arrow in
<figureCitation id="B19AFB96934EFFA2FD3AA606FD03F3FC" box="[701,763,626,648]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 4</figureCitation>
). Gonocoxosternum as in
<figureCitation id="B19AFB96934EFFA2FB82A606FF4AF3DE" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 6</figureCitation>
; differences to those of
<emphasis id="1BD53B01934EFFA2FE36A6E4FE32F3DE" box="[433,458,656,677]" italics="true" pageId="4" pageNumber="369">H.</emphasis>
<taxonomicName id="EEA19C90934EFFA2FE55A6E4FDA0F3DE" box="[466,600,656,677]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FE5CA6E4FE11F3DE" box="[475,489,656,677]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FD86A6E4FDA0F3DE" box="[513,600,656,677]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FDEFA6E4FD5FF3DE" box="[616,679,656,678]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 9</figureCitation>
) and
<emphasis id="1BD53B01934EFFA2FD62A6E4FD0FF3DE" box="[741,759,656,677]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FC80A6E4FC32F3DE" baseAuthorityName="SHARP" baseAuthorityYear="1882" box="[775,970,656,677]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FC88A6E4FCE5F3DE" box="[783,797,656,677]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FCB2A6E4FC32F3DE" box="[821,970,656,677]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FC5DA6E4FBDEF3DE" box="[986,1062,656,678]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 12</figureCitation>
) present, but irrelevant for diagnostic purposes.
</paragraph>
<paragraph id="291EE713934EFFA2FF24A6A7FE9EF2CE" blockId="4.[163,1070,722,950]" pageId="4" pageNumber="369">
M a l e s: Externally similar to females, but pro- and mesotarsi broader and anterior (inner) claw of protarsi strongly enlarged and curved (
<figureCitation id="B19AFB96934EFFA2FD49A684FCE1F27E" box="[718,793,752,774]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Fig. 22</figureCitation>
). Median lobe of aedeagus in ventral and lateral view as in
<figureCitation id="B19AFB96934EFFA2FE7DA779FD63F259" box="[506,667,781,803]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 13 and 14</figureCitation>
; apex simple, not bulged as in
<emphasis id="1BD53B01934EFFA2FC61A77AFBF8F258" box="[998,1024,782,803]" italics="true" pageId="4" pageNumber="369">H.</emphasis>
<taxonomicName id="EEA19C90934EFFA2FB8FA779FEC0F244" baseAuthorityName="SHARP" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FB96A77AFBE7F258" box="[1041,1055,782,803]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FF24A75EFEC0F244" box="[163,312,810,831]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FEC1A75EFE14F244" box="[326,492,810,832]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 18 and 18a</figureCitation>
); left paramere as in
<figureCitation id="B19AFB96934EFFA2FD44A75EFCF6F244" box="[707,782,810,832]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Fig. 19</figureCitation>
. Medial part of median lobe in lateral view significantly thinner (see arrow in
<figureCitation id="B19AFB96934EFFA2FD39A73CFCF3F226" box="[702,779,840,862]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Fig. 14</figureCitation>
) than in other two species (compare
<figureCitation id="B19AFB96934EFFA2FE89A711FE40F201" box="[270,440,869,891]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 16 and 18</figureCitation>
); corrugated membrane (see FOSTER &amp; ANGUS (1985: 8)) extending less far distally (compare
<figureCitation id="B19AFB96934EFFA2FDA6A7F6FD13F2EC" box="[545,747,898,920]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 13, 15 and 17</figureCitation>
); for comparison of parameres see
<figureCitation id="B19AFB96934EFFA2FF4EA7D4FE9EF2CE" box="[201,358,928,950]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 20 and 21.</figureCitation>
</paragraph>
<paragraph id="291EE713934EFFA2FF24A7B1FD20F28C" blockId="4.[163,1069,965,1016]" pageId="4" pageNumber="369">
M e a s u r e m e n t s: TL:
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, MW:
<quantity id="EE594AF6934EFFA2FD42A7B1FCB2F2A1" box="[709,842,965,986]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="4" pageNumber="369" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
, TL/MW: 1.93–2.01 (
<typeStatus id="F61A59B1934EFFA2FF2DA796FEF0F283" box="[170,264,994,1016]" pageId="4" pageNumber="369" type="holotype">holotype</typeStatus>
: TL:
<quantity id="EE594AF6934EFFA2FEBAA796FE76F283" box="[317,398,994,1016]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="369" unit="mm" value="3.9">3.9 mm</quantity>
, MW:
<quantity id="EE594AF6934EFFA2FE53A796FDD7F283" box="[468,559,994,1016]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="4" pageNumber="369" unit="mm" value="1.95">1.95 mm</quantity>
, TL/MW: 2.00).
</paragraph>
<paragraph id="291EE713934EFFA2FF24A07CFC62F438" blockId="4.[163,1070,1031,1347]" pageId="4" pageNumber="369">V a r i a b i l i t y: The variation of external morphological characters of the specimens studied is scanty. Worth mentioning are only: the spots on head and pronotum and the elytral vittae can be more blackish; the degree of fusion of the elytral vittae can vary; the small brownish parts on metathorax and abdomen can be darker or even black; the areas with coarser punctures on pronotum can be extended more mediad. The variation in the shape of the median lobe is small, but in some specimens the corrugated membrane extends a little more distally (ventral view). The variation in the shape of the parameres is somewhat greater, particularly because the distal part may be slightly curved inward. This also applies to the other two species and therefore the shape of the parameres is more or less unsuitable for diagnostic purposes. N o t e s: The females studied have a shiny upper surface, but the existence of a matt morph cannot be excluded.</paragraph>
<paragraph id="291EE713934EFFA2FF24A127FC2DF4A5" blockId="4.[163,1070,1362,1502]" pageId="4" pageNumber="369">
D i s t r i b u t i o n: Endemic to
<collectingCountry id="51B6A783934EFFA2FD8CA127FDB3F413" box="[523,587,1363,1384]" name="Egypt" pageId="4" pageNumber="369">Egypt</collectingCountry>
. In
<bibRefCitation id="4D309AE2934EFFA2FDFEA126FCBCF41C" author="ALFIERI A." box="[633,836,1362,1384]" pageId="4" pageNumber="369" pagination="1 - 361" refId="ref5595" refString="ALFIERI A. (1976): The Coleoptera of Egypt. - Memoires de la Societe Entomologique d'Egypte 5: i-xvi, 1 - 361." type="journal article" year="1976">ALFIERI (1976: 33)</bibRefCitation>
is given: &quot;
<emphasis id="1BD53B01934EFFA2FC38A127FEC9F4FE" italics="true" pageId="4" pageNumber="369">
in swamps around
<collectingRegion id="EB6529F1934EFFA2FF71A104FEC9F4FE" box="[246,305,1392,1413]" country="Egypt" name="Al Qahirah" pageId="4" pageNumber="369">Cairo</collectingRegion>
</emphasis>
&quot;. In
<bibRefCitation id="4D309AE2934EFFA2FEEBA104FD91F4FE" author="ZALAT S. &amp; SALEH AHMED R. &amp; ANGUS R. B." box="[364,617,1391,1414]" pageId="4" pageNumber="369" pagination="1 - 107" refId="ref6482" refString="ZALAT S., SALEH AHMED R., ANGUS R. B. &amp; A. KASCHEF (2000): Diving beetles (Coleoptera: Dytiscidae and Noteridae) of Egypt. - Egyptian Journal of Natural History 2: 1 - 107." type="journal article" year="2000">ZALAT et al. (2000: 26)</bibRefCitation>
is given: &quot;
<emphasis id="1BD53B01934EFFA2FD65A104FC1CF4BB" italics="true" pageId="4" pageNumber="369">Nile valley &amp; Delta, eastern &amp; western desert. Months of abundance: April, July-August. Habitat: collected from margins of swamps and marshes of rich vegetation; fresh and brackish water.</emphasis>
&quot; (both under
<taxonomicName id="EEA19C90934EFFA2FF64A1BCFEBCF4A6" box="[227,324,1480,1501]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus">
&quot;
<emphasis id="1BD53B01934EFFA2FF6AA1BCFEBCF4A6" box="[237,324,1480,1501]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
&quot;). Unfortunately, I was not enabled to study respective material.
</paragraph>
<paragraph id="291EE713934EFFA2FF24A199FC2EF778" blockId="4.[163,982,1517,1539]" box="[163,982,1517,1539]" pageId="4" pageNumber="369">
D e r i v a t i o n o m i n i s: The new species is named after the
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locality.
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<taxonomicName id="A8A63CBBFFF1FFCDFF34CC50FDABFDAB" ID-CoL="LD3R" authority="(HANDLIRSCH, 1908)" baseAuthorityName="HANDLIRSCH" baseAuthorityYear="1908" box="[163,594,602,624]" class="Insecta" family="Crabronidae" genus="Bembecinus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="fertoni">
<emphasis id="5DD29B2AFFF1FFCDFF34CC50FDF3FDB4" bold="true" box="[163,522,602,624]" pageId="5" pageNumber="318">
<emphasis id="5DD29B2AFFF1FFCDFF34CC50FE96FDAB" bold="true" box="[163,367,602,624]" italics="true" pageId="5" pageNumber="318">Bembecinus fertoni</emphasis>
(HANDLIRSCH
</emphasis>
,
<emphasis id="5DD29B2AFFF1FFCDFD80CC50FDABFDAB" bold="true" box="[535,594,602,624]" pageId="5" pageNumber="318">1908)</emphasis>
</taxonomicName>
<emphasis id="5DD29B2AFFF1FFCDFDCFCC50FD29FDAB" bold="true" box="[600,720,602,624]" pageId="5" pageNumber="318">(figs 24-27)</emphasis>
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</paragraph>
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<taxonomicName id="A8A63CBBFFF1FFCDFF34CC8AFDFAFD4E" ID-CoL="52SKL" authority="HANDLIRSCH, 1908: 242" authorityName="HANDLIRSCH" authorityPageNumber="242" authorityYear="1908" box="[163,515,640,661]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="fertoni">
<emphasis id="5DD29B2AFFF1FFCDFF34CC8AFEE0FD4E" box="[163,281,640,661]" italics="true" pageId="5" pageNumber="318">Stizus fertoni</emphasis>
<smallCapsWord id="69FFD1E4FFF1FFCDFE88CC8AFE62FD4F" baselines="656,656" box="[287,411,640,661]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Handlirsch" pageId="5" pageNumber="318">HANDLIRSCH</smallCapsWord>
, 1908: 242
</taxonomicName>
,
<specimenCount id="79A08CB1FFF1FFCDFD9ACC8AFDE7FD4E" box="[525,542,640,661]" count="1" pageId="5" pageNumber="318" type="female">♀</specimenCount>
♁.
<typeStatus id="B01DF99AFFF1FFCDFDD7CC8AFD6EFD4E" box="[576,663,640,661]" pageId="5" pageNumber="318" type="syntype">Syntypes</typeStatus>
:
<collectingCountry id="17B107A8FFF1FFCDFD08CC8AFD1EFD4D" box="[671,743,640,662]" name="Algeria" pageId="5" pageNumber="318">Algeria</collectingCountry>
:
<collectingRegion id="AD6289DAFFF1FFCDFD78CC8AFCC1FD4D" box="[751,824,640,662]" country="Algeria" name="Tebessa" pageId="5" pageNumber="318">Tébessa</collectingRegion>
(Vienna).
</paragraph>
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<collectionCode id="09B7DFFDFFF1FFCDFF34CCAEFF4DFD62" box="[163,180,676,697]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="318">R</collectionCode>
e m a r k:
<taxonomicName id="A8A63CBBFFF1FFCDFED6CCA9FDFEFD63" baseAuthorityName="HANDLIRSCH" baseAuthorityYear="1908" box="[321,519,675,696]" class="Insecta" family="Crabronidae" genus="Bembecinus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="fertoni">
<emphasis id="5DD29B2AFFF1FFCDFED6CCA9FDFEFD63" box="[321,519,675,696]" italics="true" pageId="5" pageNumber="318">Bembecinus fertoni</emphasis>
</taxonomicName>
was already recognized and keyed in
<collectionCode id="09B7DFFDFFF1FFCDFC19CCAEFBD4FD63" box="[910,1069,676,697]" pageId="5" pageNumber="318">SCHMID- EGGER</collectionCode>
(2004). No new material was examined. The records from
<collectingCountry id="17B107A8FFF1FFCDFC8DCCCBFC9FFD0C" box="[794,870,705,727]" name="Tunisia" pageId="5" pageNumber="318">Tunisia</collectingCountry>
in
<collectionCode id="09B7DFFDFFF1FFCDFC18CCCBFBD4FD0D" box="[911,1069,705,726]" pageId="5" pageNumber="318">SCHMID- EGGER</collectionCode>
(2004) were misidentified and belong to
<taxonomicName id="A8A63CBBFFF1FFCDFDD4CCD5FD09FD28" authorityName="MORICE" authorityYear="1911" box="[579,752,734,756]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="acanthomerus">
<emphasis id="5DD29B2AFFF1FFCDFDD4CCD5FD09FD28" box="[579,752,734,756]" italics="true" pageId="5" pageNumber="318">B. acanthomerus</emphasis>
</taxonomicName>
. The species must therefore be removed from the list of Tunisian species.
</paragraph>
<paragraph id="6F194738FFF1FFCDFF34CD2BFE25FCEC" blockId="5.[163,476,801,823]" box="[163,476,801,823]" pageId="5" pageNumber="318">
D i s t r i b u t i o n:
<collectingCountry id="17B107A8FFF1FFCDFE1ECD2BFE21FCEC" box="[393,472,801,823]" name="Algeria" pageId="5" pageNumber="318">Algeria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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105
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<paragraph id="6F194738FFF1FFCDFF34CEC7FD0FFF38" blockId="5.[163,758,203,227]" box="[163,758,203,227]" pageId="5" pageNumber="318">
<heading id="3451F054FFF1FFCDFF34CEC7FD0FFF38" bold="true" box="[163,758,203,227]" fontSize="9" level="2" pageId="5" pageNumber="318" reason="3">
<taxonomicName id="A8A63CBBFFF1FFCDFF34CEC7FD8EFF38" ID-CoL="52SJG" authority="(GUSSAKOVSKIJ, 1933)" baseAuthorityName="GUSSAKOVSKIJ" baseAuthorityYear="1933" box="[163,631,203,227]" class="Insecta" family="Crabronidae" genus="Bembecinus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="5DD29B2AFFF1FFCDFF34CEC7FE86FF38" bold="true" box="[163,383,205,227]" italics="true" pageId="5" pageNumber="318">Bembecinus dentipes</emphasis>
<emphasis id="5DD29B2AFFF1FFCDFE12CEC1FD8EFF38" bold="true" box="[389,631,203,227]" pageId="5" pageNumber="318">(GUSSAKOVSKIJ, 1933)</emphasis>
</taxonomicName>
<emphasis id="5DD29B2AFFF1FFCDFDEACEC7FD0FFF38" bold="true" box="[637,758,205,227]" pageId="5" pageNumber="318">(figs 16-18)</emphasis>
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<subSubSection id="27BC14B3FFF1FFCDFF34CEF9FCFEFEE4" pageId="5" pageNumber="318" type="reference_group">
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<taxonomicName id="A8A63CBBFFF1FFCDFF34CEF9FDC9FEDC" ID-CoL="52SJG" authority="GUSSAKOVSKIJ, 1933 b: 289" authorityName="GUSSAKOVSKIJ" authorityPageNumber="289" authorityYear="1933" box="[163,560,242,264]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="5DD29B2AFFF1FFCDFF34CEF9FED0FEDC" box="[163,297,242,264]" italics="true" pageId="5" pageNumber="318">Stizus dentipes</emphasis>
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, 1933b: 289
</taxonomicName>
, ♁,
<collectingCountry id="17B107A8FFF1FFCDFDC1CEF9FD84FED3" box="[598,637,243,264]" name="Iran" pageId="5" pageNumber="318">Iran</collectingCountry>
,
<collectingRegion id="AD6289DAFFF1FFCDFD11CEF9FD28FED3" box="[646,721,243,264]" country="Iran" name="Kerman" pageId="5" pageNumber="318">Kerman</collectingRegion>
, Kucha, Kambil (St. Petersburg).
</paragraph>
<paragraph id="6F194738FFF1FFCDFF34CF18FCFEFEE4" blockId="5.[163,1070,241,320]" pageId="5" pageNumber="318">
M a t e r i a l e x a m i n e d: 1♁,
<date id="1B1861F8FFF1FFCDFD9CCF18FD90FEF3" box="[523,617,274,296]" pageId="5" pageNumber="318" value="1956-06-30">30.vi.1956</date>
SO-Iran (Djiroft), Anbar-Abad
<geoCoordinate id="0A9221FFFFF1FFCDFC12CF18FC2EFEFC" box="[901,983,274,295]" degrees="28.474" direction="north" orientation="latitude" pageId="5" pageNumber="318" precision="55" value="28.474">28.474 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF1FFCDFC49CF18FBD7FEFC" box="[990,1070,274,296]" degrees="57.845" direction="east" orientation="longitude" pageId="5" pageNumber="318" precision="55" value="57.845">57.845 E</geoCoordinate>
(leg. Richter, det. de Beaumont, coll. St. Museum Nat. Stuttgart).
</paragraph>
</subSubSection>
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<paragraph id="6F194738FFF1FFCDFF35CF44FE79FDD1" blockId="5.[162,1070,333,523]" pageId="5" pageNumber="318">
R e m a r k: The species was treated as a doubtfull synonym of
<taxonomicName id="A8A63CBBFFF1FFCDFCF1CF44FBF6FEB9" authorityName="MORICE" authorityYear="1911" box="[870,1039,333,355]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="acanthomerus">
<emphasis id="5DD29B2AFFF1FFCDFCF1CF44FBF6FEB9" box="[870,1039,333,355]" italics="true" pageId="5" pageNumber="318">B. acanthomerus</emphasis>
</taxonomicName>
in SCHMID- EGGER (2004), but was not synonymised due to lack of
<typeStatus id="B01DF99AFFF1FFCDFCDCCF60FC8EFEA4" box="[843,887,362,383]" pageId="5" pageNumber="318">type</typeStatus>
comparison. The here examined male however, is clearly a valid species and differs by its color pattern and morphology from
<taxonomicName id="A8A63CBBFFF1FFCDFE1ECFA8FDCDFE6D" authorityName="MORICE" authorityYear="1911" box="[393,564,417,439]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="acanthomerus">
<emphasis id="5DD29B2AFFF1FFCDFE1ECFA8FDCDFE6D" box="[393,564,417,439]" italics="true" pageId="5" pageNumber="318">B. acanthomerus</emphasis>
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. Because it was found close to the
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area, it is assigned here to
<taxonomicName id="A8A63CBBFFF1FFCDFECCCFB4FE29FE09" authorityName="GUSSAKOVSKIJ" authorityYear="1933" box="[347,464,445,467]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="5DD29B2AFFF1FFCDFECCCFB4FE29FE09" box="[347,464,445,467]" italics="true" pageId="5" pageNumber="318">B. dentipes</emphasis>
</taxonomicName>
, even if the
<typeStatus id="B01DF99AFFF1FFCDFDF5CFB4FD77FE08" box="[610,654,446,467]" pageId="5" pageNumber="318">type</typeStatus>
could not be examined. However, the description matches well. It was already identified and published by
<bibRefCitation id="0B373AC9FFF1FFCDFC03CFD1FF1FFDD1" author="DE BEAUMONT J." pageId="5" pageNumber="318" pagination="1 - 18" refId="ref9960" refString="DE BEAUMONT J. (1970): Sphecidae de l'Iran (Hym.) (Resultats de voyages entomologiques de Willi Richter, Stuttgart, en Iran, en 1954 et 1956). - Stuttgarter Beitrage zur Naturkunde aus dem Staatlichen Museum fur Naturkunde in Stuttgart No. 220: 1 - 18." type="book chapter" year="1970">DE BEAUMONT (1970)</bibRefCitation>
as
<emphasis id="5DD29B2AFFF1FFCDFE9FCFFCFE79FDD1" box="[264,384,501,523]" italics="true" pageId="5" pageNumber="318">
<taxonomicName id="A8A63CBBFFF1FFCDFE9FCFFCFE82FDD1" authorityName="GUSSAKOVSKIJ" authorityYear="1933" box="[264,379,501,523]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="dentipes">B. dentipes</taxonomicName>
.
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D i s t r i b u t i o n: Southern
<collectingCountry id="17B107A8FFF1FFCDFE7CCC11FDEEFDEB" box="[491,535,539,560]" name="Iran" pageId="5" pageNumber="318">Iran</collectingCountry>
.
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273
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<mods:classification id="1F4945F53B85A76FF47909E5F926A7B9">journal article</mods:classification>
<mods:identifier id="68CDEE56F4AB0358B06A5389AD906933" type="DOI">10.5281/zenodo.14681733</mods:identifier>
<mods:identifier id="0EC47484FC2ADDEEB22AB61A82182BFD" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="068BA5B5516D11A7D64BC397C44CFF44" type="Zenodo-Dep">14681733</mods:identifier>
</mods:mods>
<treatment id="E70FF62EFFF7FFCBFF34CCFAFC41FB3D" ID-DOI="http://doi.org/10.5281/zenodo.14681735" ID-Zenodo-Dep="14681735" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D" httpUri="http://treatment.plazi.org/id/E70FF62EFFF7FFCBFF34CCFAFC41FB3D" lastPageNumber="316" pageId="3" pageNumber="316">
<subSubSection id="27BC14B3FFF7FFCBFF34CCFAFD2AFCDD" box="[163,723,752,774]" pageId="3" pageNumber="316" type="nomenclature">
<paragraph id="6F194738FFF7FFCBFF34CCFAFD2AFCDD" blockId="3.[163,723,752,774]" box="[163,723,752,774]" pageId="3" pageNumber="316">
<heading id="3451F054FFF7FFCBFF34CCFAFD2AFCDD" box="[163,723,752,774]" fontSize="8" level="2" pageId="3" pageNumber="316" reason="2">
<taxonomicName id="A8A63CBBFFF7FFCBFF34CCFAFD97FCDD" ID-CoL="LCZS" authority="(MORICE, 1911)" baseAuthorityName="MORICE" baseAuthorityYear="1911" box="[163,622,752,774]" class="Insecta" family="Crabronidae" genus="Bembecinus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="acanthomerus">
<emphasis id="5DD29B2AFFF7FFCBFF34CCFAFDDFFCDE" bold="true" box="[163,550,752,774]" pageId="3" pageNumber="316">
<emphasis id="5DD29B2AFFF7FFCBFF34CCFAFE47FCDD" bold="true" box="[163,446,752,774]" italics="true" pageId="3" pageNumber="316">Bembecinus acanthomerus</emphasis>
(MORICE
</emphasis>
,
<emphasis id="5DD29B2AFFF7FFCBFDA4CCFAFD97FCDD" bold="true" box="[563,622,752,774]" pageId="3" pageNumber="316">1911)</emphasis>
</taxonomicName>
<emphasis id="5DD29B2AFFF7FFCBFDE3CCFAFD2AFCDD" bold="true" box="[628,723,752,774]" pageId="3" pageNumber="316">(figs 1-5)</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="27BC14B3FFF7FFCBFF34CD1FFC8BFBD1" pageId="3" pageNumber="316" type="reference_group">
<paragraph id="6F194738FFF7FFCBFF34CD1FFC6FFCF0" blockId="3.[163,1071,787,1034]" box="[163,918,787,811]" pageId="3" pageNumber="316">
<taxonomicName id="A8A63CBBFFF7FFCBFF34CD1FFDE1FCF0" ID-CoL="52SFW" authority="MORICE, 1911: 116" authorityName="MORICE" authorityPageNumber="116" authorityYear="1911" box="[163,536,789,811]" class="Insecta" family="Crabronidae" genus="Stizus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="acanthomerus">
<emphasis id="5DD29B2AFFF7FFCBFF34CD1FFEA4FCF1" box="[163,349,789,810]" italics="true" pageId="3" pageNumber="316">Stizus acanthomerus</emphasis>
<smallCapsWord id="69FFD1E4FFF7FFCBFEF4CD1FFE49FCF1" baselines="805,806" box="[355,432,789,810]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Morice" pageId="3" pageNumber="316">MORICE</smallCapsWord>
, 1911: 116
</taxonomicName>
,
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♁.
<typeStatus id="B01DF99AFFF7FFCBFDD8CD1FFD5CFCF1" box="[591,677,789,810]" pageId="3" pageNumber="316" type="syntype">Syntypes</typeStatus>
:
<collectingCountry id="17B107A8FFF7FFCBFD39CD1FFD0FFCF0" box="[686,758,789,811]" name="Algeria" pageId="3" pageNumber="316">Algeria</collectingCountry>
:
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(Oxford).
</paragraph>
<paragraph id="6F194738FFF7FFCBFF34CD3FFC8BFBD1" blockId="3.[163,1071,787,1034]" pageId="3" pageNumber="316">
<materialsCitation id="DFCE4D65FFF7FFCBFF34CD3FFE20FCB9" collectingDate="2014-06-12" collectionCode="CSE" country="Morocco" latitude="30.551" location="Assaki" longLatPrecision="72" longitude="-8.025" pageId="3" pageNumber="316" specimenCount="2" specimenCount-female="2">
M a t e r i a l e x a m i n e d:
<collectingCountry id="17B107A8FFF7FFCBFE70CD3FFDA3FC91" box="[487,602,821,842]" name="Morocco" pageId="3" pageNumber="316">MOROCCO</collectingCountry>
: 2♁♁
<specimenCount id="79A08CB1FFF7FFCBFD0ECD3FFD3EFC91" box="[665,711,821,842]" count="2" pageId="3" pageNumber="316" type="female">2♀♀</specimenCount>
,
<date id="1B1861F8FFF7FFCBFD45CD3EFCC8FC91" box="[722,817,820,842]" pageId="3" pageNumber="316" value="2014-06-12">
<collectingDate id="0B5C9810FFF7FFCBFD45CD3EFCC8FC91" box="[722,817,820,842]" pageId="3" pageNumber="316" value="2014-06-12">12.vi.2014</collectingDate>
</date>
,
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<quantity id="A85EEADDFFF7FFCBFCABCD3FFC8FFC91" box="[828,886,820,842]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="316" unit="km" value="10.0">10 km</quantity>
E Aoulouz
</locationDeviation>
,
<location id="6A7911E3FFF7FFCBFC7BCD3EFBD3FC91" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFC7BCD3EFBD3FC91" box="[1004,1066,820,842]" country="Morocco" latitude="30.551" longLatPrecision="72" longitude="-8.025" name="Assaki" pageId="3" pageNumber="316">Assaki</location>
,
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<geoCoordinate id="0A9221FFFFF7FFCBFEBFCD46FE8DFCB9" box="[296,372,844,866]" degrees="8.025" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-8.025">8.025 W</geoCoordinate>
(leg.
<collectionCode id="09B7DFFDFFF7FFCBFE30CD47FE2AFCB9" box="[423,467,845,866]" pageId="3" pageNumber="316">CSE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFE74CD41FED6FCA1" collectingDate="2018-04-25" collectionCode="CSE" country="Morocco" latitude="30.468" location="Tafdroust" longLatPrecision="72" longitude="-5.918" pageId="3" pageNumber="316" specimenCount="1">
4♁♁,
<date id="1B1861F8FFF7FFCBFD8DCD46FD8EFCB9" box="[538,631,844,866]" pageId="3" pageNumber="316" value="2018-04-25">
<collectingDate id="0B5C9810FFF7FFCBFD8DCD46FD8EFCB9" box="[538,631,844,866]" pageId="3" pageNumber="316" value="2018-04-25">25.iv.2018</collectingDate>
</date>
<locationDeviation id="7EAB5E1AFFF7FFCBFDEACD46FCDFFCB9" box="[637,806,844,866]" location="Zagora" pageId="3" pageNumber="316">
<quantity id="A85EEADDFFF7FFCBFDEACD46FD4FFCB9" box="[637,694,844,866]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="316" unit="km" value="16.0">16 km</quantity>
NW Zagora
</locationDeviation>
,
<location id="6A7911E3FFF7FFCBFCB8CD46FC7DFCB9" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFCB8CD46FC7DFCB9" box="[815,900,844,866]" country="Morocco" latitude="30.468" longLatPrecision="72" longitude="-5.918" name="Tafdroust" pageId="3" pageNumber="316">Tafdroust</location>
<geoCoordinate id="0A9221FFFFF7FFCBFC1DCD46FC25FCB9" box="[906,988,844,866]" degrees="30.468" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="30.468">30.468 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFC75CD46FBD6FCB9" box="[994,1071,844,866]" degrees="5.918" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-5.918">5.918 W</geoCoordinate>
(leg.
<collectionCode id="09B7DFFDFFF7FFCBFF6ACD6FFED0FCA1" box="[253,297,869,890]" pageId="3" pageNumber="316">CSE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFEAFCD69FC80FCA1" box="[312,889,867,890]" collectingDate="2015-04-12" collectionCode="CSE" country="Morocco" latitude="29.922" location="Tissint" longLatPrecision="73" longitude="-7.328" pageId="3" pageNumber="316" specimenCount="1">
2♁♁,
<date id="1B1861F8FFF7FFCBFEF8CD6EFE34FCA1" box="[367,461,868,890]" pageId="3" pageNumber="316" value="2015-04-12">
<collectingDate id="0B5C9810FFF7FFCBFEF8CD6EFE34FCA1" box="[367,461,868,890]" pageId="3" pageNumber="316" value="2015-04-12">12.iv.2015</collectingDate>
</date>
<quantity id="A85EEADDFFF7FFCBFE45CD6FFDF9FCA1" box="[466,512,868,890]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="316" unit="km" value="2.0">2 km</quantity>
NW
<location id="6A7911E3FFF7FFCBFDA7CD6EFD92FCA1" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFDA7CD6EFD92FCA1" box="[560,619,868,890]" country="Morocco" latitude="29.922" longLatPrecision="73" longitude="-7.328" name="Tissint" pageId="3" pageNumber="316">Tissint</location>
<geoCoordinate id="0A9221FFFFF7FFCBFDE6CD6FFD3AFCA1" box="[625,707,869,890]" degrees="29.922" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="29.922">29.922 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFD5ECD6FFCECFCA1" box="[713,789,869,890]" degrees="7.328" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-7.328">7.328 W</geoCoordinate>
(leg.
<collectionCode id="09B7DFFDFFF7FFCBFCD0CD6FFC8AFCA1" box="[839,883,869,890]" pageId="3" pageNumber="316">CSE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFC14CD69FD50FC49" collectingDate="2014-06-16" collectionCode="CSE" country="Morocco" latitude="29.69" location="Assaka" longLatPrecision="73" longitude="-9.53" pageId="3" pageNumber="316" specimenCount="1">
3♁♁
<date id="1B1861F8FFF7FFCBFC22CD6EFBE8FCA1" box="[949,1041,868,890]" pageId="3" pageNumber="316" value="2014-06-16">
<collectingDate id="0B5C9810FFF7FFCBFC22CD6EFBE8FCA1" box="[949,1041,868,890]" pageId="3" pageNumber="316" value="2014-06-16">16.vi.2014</collectingDate>
</date>
<locationDeviation id="7EAB5E1AFFF7FFCBFB8FCD6FFEBCFC49" location="Tizinit" pageId="3" pageNumber="316">
<quantity id="A85EEADDFFF7FFCBFB8FCD6FFF14FC49" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="316" unit="km" value="20.0">20 km</quantity>
E Tizinit
</locationDeviation>
,
<location id="6A7911E3FFF7FFCBFEC7CD76FE76FC49" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFEC7CD76FE76FC49" box="[336,399,892,914]" country="Morocco" latitude="29.69" longLatPrecision="73" longitude="-9.53" name="Assaka" pageId="3" pageNumber="316">Assaka</location>
<geoCoordinate id="0A9221FFFFF7FFCBFE00CD76FE12FC49" box="[407,491,892,914]" degrees="29.690" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="29.69">29.690 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFE64CD76FDB9FC49" box="[499,576,892,914]" degrees="9.530" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-9.53">9.530 W</geoCoordinate>
(leg.
<collectionCode id="09B7DFFDFFF7FFCBFDE0CD77FD5AFC49" box="[631,675,893,914]" pageId="3" pageNumber="316">CSE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFD23CD77FE2FFC71" collectingDate="2014-06-15" collectionCode="CSE" country="Morocco" latitude="10.5101" location="Tazenakht" longLatPrecision="54" longitude="-7.228" pageId="3" pageNumber="316" specimenCount="1" specimenCount-female="1">
<specimenCount id="79A08CB1FFF7FFCBFD23CD77FD29FC49" box="[692,720,893,914]" count="1" pageId="3" pageNumber="316" type="female">1♀</specimenCount>
<date id="1B1861F8FFF7FFCBFD4FCD76FCCCFC49" box="[728,821,892,914]" pageId="3" pageNumber="316" value="2014-06-15">
<collectingDate id="0B5C9810FFF7FFCBFD4FCD76FCCCFC49" box="[728,821,892,914]" pageId="3" pageNumber="316" value="2014-06-15">15.vi.2014</collectingDate>
</date>
<quantity id="A85EEADDFFF7FFCBFCABCD76FC95FC49" box="[828,876,892,914]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="316" unit="km" value="5.0">5 km</quantity>
NW
<location id="6A7911E3FFF7FFCBFC37CD76FC04FC49" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFC37CD76FC04FC49" box="[928,1021,892,914]" country="Morocco" latitude="10.5101" longLatPrecision="54" longitude="-7.228" name="Tazenakht" pageId="3" pageNumber="316">Tazenakht</location>
, N
<geoCoordinate id="0A9221FFFFF7FFCBFB8ECD77FED8FC71" degrees="10" direction="north" minutes="30.606" orientation="latitude" pageId="3" pageNumber="316" precision="1" value="10.5101">10 30.606 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFEB0CD9FFE8BFC71" box="[295,370,917,938]" degrees="7.228" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-7.228">7.228 W</geoCoordinate>
(leg.
<collectionCode id="09B7DFFDFFF7FFCBFE33CD9FFE29FC71" box="[420,464,917,938]" pageId="3" pageNumber="316">CSE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFE48CD99FEC9FC19" collectingDate="2018-04-25" collectionCode="CSE" country="Morocco" latitude="30.392" location="Boikhlal" longLatPrecision="72" longitude="-5.796" pageId="3" pageNumber="316" specimenCount="2" specimenCount-female="2">
1♁,
<specimenCount id="79A08CB1FFF7FFCBFD92CD9FFDCBFC71" box="[517,562,917,938]" count="2" pageId="3" pageNumber="316" type="female">2♀♀</specimenCount>
,
<date id="1B1861F8FFF7FFCBFDACCD9EFD61FC71" box="[571,664,916,938]" pageId="3" pageNumber="316" value="2018-04-25">
<collectingDate id="0B5C9810FFF7FFCBFDACCD9EFD61FC71" box="[571,664,916,938]" pageId="3" pageNumber="316" value="2018-04-25">25.iv.2018</collectingDate>
</date>
<locationDeviation id="7EAB5E1AFFF7FFCBFD09CD9EFCCAFC71" box="[670,819,916,938]" location="Zagora" pageId="3" pageNumber="316">
<quantity id="A85EEADDFFF7FFCBFD09CD9EFD32FC71" box="[670,715,916,938]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="316" unit="km" value="6.0">6 km</quantity>
NE Zagora
</locationDeviation>
,
<location id="6A7911E3FFF7FFCBFCACCD9EFC7FFC71" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFCACCD9EFC7FFC71" box="[827,902,916,938]" country="Morocco" latitude="30.392" longLatPrecision="72" longitude="-5.796" name="Boikhlal" pageId="3" pageNumber="316">Boikhlal</location>
<geoCoordinate id="0A9221FFFFF7FFCBFC1CCD9FFC24FC71" box="[907,989,917,938]" degrees="30.392" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="30.392">30.392 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFC75CD9EFBD6FC71" box="[994,1071,916,938]" degrees="5.796" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-5.796">5.796 W</geoCoordinate>
(leg.
<collectionCode id="09B7DFFDFFF7FFCBFF6ACDA7FED0FC19" box="[253,297,941,962]" pageId="3" pageNumber="316">CSE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFEADCDA1FBFAFC19" box="[314,1027,939,962]" collectingDate="2014-06-12" collectionCode="CSE" country="Morocco" latitude="30.697" location="Aoulouz" longLatPrecision="72" longitude="-8.156" municipality="Souss river" pageId="3" pageNumber="316" specimenCount="1">
12♁♁
<date id="1B1861F8FFF7FFCBFEE0CDA6FE2DFC19" box="[375,468,940,962]" pageId="3" pageNumber="316" value="2014-06-12">
<collectingDate id="0B5C9810FFF7FFCBFEE0CDA6FE2DFC19" box="[375,468,940,962]" pageId="3" pageNumber="316" value="2014-06-12">12.vi.2014</collectingDate>
</date>
<location id="6A7911E3FFF7FFCBFE4CCDA6FDD0FC19" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFE4CCDA6FDD0FC19" box="[475,553,940,962]" country="Morocco" latitude="30.697" longLatPrecision="72" longitude="-8.156" municipality="Souss river" name="Aoulouz" pageId="3" pageNumber="316">Aoulouz</location>
, bridge on
<collectingMunicipality id="8F7DDD42FFF7FFCBFD1ACDA7FD09FC19" box="[653,752,940,962]" pageId="3" pageNumber="316">Souss river</collectingMunicipality>
<geoCoordinate id="0A9221FFFFF7FFCBFD60CDA6FCB0FC19" box="[759,841,940,962]" degrees="30,697" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="30.697">30,697 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFCC7CDA6FC65FC19" box="[848,924,940,962]" degrees="8.156" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-8.156">8.156 W</geoCoordinate>
(leg.
<collectionCode id="09B7DFFDFFF7FFCBFC46CDA7FC04FC19" box="[977,1021,941,962]" pageId="3" pageNumber="316">CSE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFB9ACDA1FC8DFC01" collectingDate="2014-05-28" collectorName="Kroupa" country="Morocco" latitude="30.051" location="Lake" longLatPrecision="73" longitude="-9.108" pageId="3" pageNumber="316" specimenCount="1" specimenCount-female="1">
1♁,
<specimenCount id="79A08CB1FFF7FFCBFF47CDCFFF15FC01" box="[208,236,965,986]" count="1" pageId="3" pageNumber="316" type="female">1♀</specimenCount>
<date id="1B1861F8FFF7FFCBFF62CDCFFEB5FC01" box="[245,332,965,986]" pageId="3" pageNumber="316" value="2014-05-28">
<collectingDate id="0B5C9810FFF7FFCBFF62CDCFFEB5FC01" box="[245,332,965,986]" pageId="3" pageNumber="316" value="2014-05-28">28.v.2014</collectingDate>
</date>
<location id="6A7911E3FFF7FFCBFEC2CDCEFE78FC01" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFEC2CDCEFE78FC01" box="[341,385,964,986]" country="Morocco" latitude="30.051" longLatPrecision="73" longitude="-9.108" name="Lake" pageId="3" pageNumber="316">Lake</location>
<locationDeviation id="7EAB5E1AFFF7FFCBFE1DCDCFFDC0FC01" box="[394,569,964,986]" location="Ait Baha" pageId="3" pageNumber="316">
<quantity id="A85EEADDFFF7FFCBFE1DCDCFFE42FC01" box="[394,443,964,986]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="316" unit="km" value="2.0">2 km</quantity>
SE Ait Baha
</locationDeviation>
<geoCoordinate id="0A9221FFFFF7FFCBFDD5CDCEFD6EFC01" box="[578,663,964,986]" degrees="30.051" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="30.051">30.051 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFD37CDCFFD09FC01" box="[672,752,965,986]" degrees="9.108" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-9.108">9.108 W</geoCoordinate>
(leg.
<collectorName id="C25322EEFFF7FFCBFCBECDCFFC97FC01" box="[809,878,965,986]" pageId="3" pageNumber="316">Kroupa</collectorName>
)
</materialsCitation>
;
<materialsCitation id="DFCE4D65FFF7FFCBFC17CDC9FD64FC29" collectingDate="2018-04-25" collectionCode="CSE" country="Morocco" latitude="30.312" location="Zagora" longLatPrecision="72" longitude="-5.841" municipality="Qued Draa" pageId="3" pageNumber="316" specimenCount="1">
9♁♁,
<date id="1B1861F8FFF7FFCBFC2CCDCEFBE1FC01" box="[955,1048,964,986]" pageId="3" pageNumber="316" value="2018-04-25">
<collectingDate id="0B5C9810FFF7FFCBFC2CCDCEFBE1FC01" box="[955,1048,964,986]" pageId="3" pageNumber="316" value="2018-04-25">25.iv.2018</collectingDate>
</date>
S
<location id="6A7911E3FFF7FFCBFF47CDD7FEE9FC29" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFF47CDD7FEE9FC29" box="[208,272,989,1010]" country="Morocco" latitude="30.312" longLatPrecision="72" longitude="-5.841" municipality="Qued Draa" name="Zagora" pageId="3" pageNumber="316">Zagora</location>
,
<collectingMunicipality id="8F7DDD42FFF7FFCBFE8BCDD6FE86FC29" box="[284,383,988,1010]" pageId="3" pageNumber="316">Qued Draa</collectingMunicipality>
<geoCoordinate id="0A9221FFFFF7FFCBFE10CDD7FE25FC29" box="[391,476,989,1010]" degrees="30.312" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="30.312">30.312 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFE72CDD6FDCAFC29" box="[485,563,988,1010]" degrees="5.841" direction="west" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="-5.841">5.841 W</geoCoordinate>
(leg.
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)
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.
<materialsCitation id="DFCE4D65FFF7FFCBFD3ECDD7FC94FBD1" collectingDate="2000-06-08" collectorName="Hauser" country="Tunisia" county="Matmata" latitude="33.539" location="Sud-Tunesien" longLatPrecision="72" longitude="9.974" municipality="Ksourberge" pageId="3" pageNumber="316" specimenCount="2" specimenCount-female="2">
<collectingCountry id="17B107A8FFF7FFCBFD3ECDD7FCFCFC29" box="[681,773,989,1010]" name="Tunisia" pageId="3" pageNumber="316">TUNISIA</collectingCountry>
:
<specimenCount id="79A08CB1FFF7FFCBFC98CDD7FCC5FC29" box="[783,828,989,1010]" count="2" pageId="3" pageNumber="316" type="female">2♀♀</specimenCount>
,
<date id="1B1861F8FFF7FFCBFCDECDD7FC5FFC29" box="[841,934,988,1010]" pageId="3" pageNumber="316" value="2000-06-08">
<collectingDate id="0B5C9810FFF7FFCBFCDECDD7FC5FFC29" box="[841,934,988,1010]" pageId="3" pageNumber="316" value="2000-06-08">08.vi.2000</collectingDate>
</date>
<location id="6A7911E3FFF7FFCBFC38CDD6FBD2FC29" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFC38CDD6FBD2FC29" box="[943,1067,988,1010]" country="Tunisia" county="Matmata" latitude="33.539" longLatPrecision="72" longitude="9.974" municipality="Ksourberge" name="Sud-Tunesien" pageId="3" pageNumber="316">Süd-Tunesien</location>
,
<location id="6A7911E3FFF7FFCBFF47CDFEFEC6FBD1" LSID="urn:lsid:plazi:treatment:E70FF62EFFF7FFCBFF34CCFAFC41FB3D:6A7911E3FFF7FFCBFF47CDFEFEC6FBD1" box="[208,319,1012,1034]" country="Tunisia" county="Matmata" latitude="33.539" longLatPrecision="72" longitude="9.974" municipality="Ksourberge" name="Dhargebirge" pageId="3" pageNumber="316">Dhargebirge</location>
,
<collectingMunicipality id="8F7DDD42FFF7FFCBFED0CDFEFE56FBD1" box="[327,431,1012,1034]" pageId="3" pageNumber="316">Ksourberge</collectingMunicipality>
,
<quantity id="A85EEADDFFF7FFCBFE2FCDFFFE1CFBD1" box="[440,485,1012,1034]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="316" unit="km" value="3.0">3 km</quantity>
sw
<collectingCounty id="86783FB4FFF7FFCBFD90CDFFFDACFBD1" box="[519,597,1013,1034]" pageId="3" pageNumber="316">Matmata</collectingCounty>
<geoCoordinate id="0A9221FFFFF7FFCBFDCDCDFEFD52FBD1" box="[602,683,1012,1034]" degrees="33.539" direction="north" orientation="latitude" pageId="3" pageNumber="316" precision="55" value="33.539">33.539 N</geoCoordinate>
<geoCoordinate id="0A9221FFFFF7FFCBFD27CDFFFD0CFBD1" box="[688,757,1013,1034]" degrees="9.974" direction="east" orientation="longitude" pageId="3" pageNumber="316" precision="55" value="9.974">9.974 E</geoCoordinate>
(leg.
<collectorName id="C25322EEFFF7FFCBFCB2CDFFFC9EFBD1" box="[805,871,1013,1034]" pageId="3" pageNumber="316">Hauser</collectorName>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="27BC14B3FFF7FFCBFF35CA13FC41FB3D" pageId="3" pageNumber="316" type="description">
<paragraph id="6F194738FFF7FFCBFF35CA13FEE2FB7F" blockId="3.[162,1070,1049,1188]" pageId="3" pageNumber="316">R e m a r k: The species was treated as a species aggregate with a wide distribution in SCHMID- EGGER (2004). Here the species is re-defined, and refers only to specimens from Northwest Africa, and according to the characteristics in the key. Only newly collected material is listed above, see SCHMID- EGGER (2004) for other records and for detailed description.</paragraph>
<paragraph id="6F194738FFF7FFCBFF34CABEFC41FB3D" blockId="3.[163,1070,1203,1255]" pageId="3" pageNumber="316">
D i s t r i b u t i o n:
<collectingCountry id="17B107A8FFF7FFCBFE1BCABEFE13FB12" box="[396,490,1204,1225]" name="Morocco" pageId="3" pageNumber="316">Morocco</collectingCountry>
,
<collectingCountry id="17B107A8FFF7FFCBFE60CAB9FDBCFB12" box="[503,581,1203,1225]" name="Algeria" pageId="3" pageNumber="316">Algeria</collectingCountry>
, new record for
<collectingCountry id="17B107A8FFF7FFCBFD60CAB9FCBAFB12" box="[759,835,1203,1225]" name="Tunisia" pageId="3" pageNumber="316">Tunisia</collectingCountry>
(the above mentioned record from Matmata was misidentfied as
<taxonomicName id="A8A63CBBFFF7FFCBFDC7CAD8FD49FB3D" baseAuthorityName="HANDLIRSCH" baseAuthorityYear="1908" box="[592,688,1233,1255]" class="Insecta" family="Crabronidae" genus="Bembecinus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="fertoni">
<emphasis id="5DD29B2AFFF7FFCBFDC7CAD8FD49FB3D" box="[592,688,1233,1255]" italics="true" pageId="3" pageNumber="316">B. fertoni</emphasis>
</taxonomicName>
in SCHMID- EGGER 2004).
</paragraph>
</subSubSection>
</treatment>
</document>

179
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<heading id="3A5481B90B46FFE81785FF00FA7D9F99" box="[163,860,205,227]" fontSize="8" level="2" pageId="12" pageNumber="359" reason="2">
<taxonomicName id="A6A34D560B46FFE81785FF00FBAE9F98" ID-CoL="DRNS" authority="SMITH, 1878" authorityName="SMITH" authorityYear="1878" box="[163,655,205,227]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="floridula">
<emphasis id="53D7EAC70B46FFE81785FF00FB719F98" bold="true" box="[163,592,205,227]" pageId="12" pageNumber="359">
<emphasis id="53D7EAC70B46FFE81785FF00F9DF9F99" bold="true" box="[163,254,205,227]" italics="true" pageId="12" pageNumber="359">Andrena</emphasis>
(
<emphasis id="53D7EAC70B46FFE8162BFF00F8BA9F99" bold="true" box="[269,411,205,227]" italics="true" pageId="12" pageNumber="359">incertae sedis</emphasis>
)
<emphasis id="53D7EAC70B46FFE8168FFF00FB259F99" bold="true" box="[425,516,205,227]" italics="true" pageId="12" pageNumber="359">floridula</emphasis>
<bibRefCitation id="05324B240B46FFE8152DFF00FB719F98" author="SMITH F." box="[523,592,205,227]" pageId="12" pageNumber="359" pagination="1 - 22" refId="ref7315" refString="SMITH F. (1878): Hymenoptera. - In: Scientific results of the second Yarkand Mission, based upon the collections and notes of the late Ferdinand Stoliczka. By order of the Government of India. 1878 - 91. pp. 1 - 22. Calcutta (government printing)." type="book chapter" year="1878">SMITH</bibRefCitation>
</emphasis>
,
<emphasis id="53D7EAC70B46FFE8157AFF00FBAE9F98" bold="true" box="[604,655,205,226]" pageId="12" pageNumber="359">1878</emphasis>
</taxonomicName>
<emphasis id="53D7EAC70B46FFE815B3FF00FA7D9F99" bold="true" box="[661,860,205,227]" pageId="12" pageNumber="359">
(
<figureCitation id="F9982A500B46FFE815BAFF00FA2C9F98" box="[668,781,205,227]" captionStart="Figs 17-21" captionStartId="8.[163,203,1500,1522]" captionTargetBox="[166,1072,199,1487]" captionTargetId="figure-53@8.[166,1072,199,1487]" captionTargetPageId="8" captionText="Figs 17-21: (17) Collecting site of Rumbak at 3850 m; (18-19) Andrena mordax MORAWITZ, 1876 female at Mentha flowers (Lamiaceae; photographs taken at Rumbak); (20-21) Andrena floridula SMITH, 1878 female at Asteraceae flowers (photographs taken in Leh gardens; photographs: M. Jacobs)." figureDoi="http://doi.org/10.5281/zenodo.14681699" httpUri="https://zenodo.org/record/14681699/files/figure.png" pageId="12" pageNumber="359">Figs 20-21</figureCitation>
,
<figureCitation id="F9982A500B46FFE8143FFF00FA749F99" box="[793,853,205,227]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">36-43</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="611C36D50B46FFE81785FF39FB689E5B" blockId="12.[163,1070,243,289]" pageId="12" pageNumber="359">
M a t e r i a l e x a m i n e d:
<collectingCountry id="19B476450B46FFE816C0FF3EFB379E73" box="[486,534,243,265]" name="India" pageId="12" pageNumber="359">India</collectingCountry>
: Ladakh, Leh Tehsil, city gardens,
<quantity id="A65B9B300B46FFE8146CFF3EFAAE9E73" box="[842,911,243,265]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="359" unit="m" value="3500.0">3500 m</quantity>
,
<date id="151D10150B46FFE814BEFF3EFD229E73" box="[920,1027,243,265]" pageId="12" pageNumber="359" value="2015-08-30">30.viii.2015</date>
,
<specimenCount id="77A5FD5C0B46FFE8132AFF3EFD089E72" box="[1036,1065,243,264]" count="1" pageId="12" pageNumber="359" type="female">1♀</specimenCount>
, leg. M. Jacobs, coll. Biologiezentrum Linz.
</paragraph>
</subSubSection>
<subSubSection id="29B9655E0B46FFE91784FEFDFB859B6F" lastPageId="13" lastPageNumber="360" pageId="12" pageNumber="359" type="description">
<paragraph id="611C36D50B46FFE81784FEFDFA189D4A" blockId="12.[162,1070,303,560]" pageId="12" pageNumber="359">
D i a g n o s i s:
<taxonomicName id="A6A34D560B46FFE81668FEE2FB229E3E" authorityName="SMITH" authorityYear="1878" box="[334,515,303,324]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="floridula">
<emphasis id="53D7EAC70B46FFE81668FEE2FB229E3E" box="[334,515,303,324]" italics="true" pageId="12" pageNumber="359">Andrena floridula</emphasis>
</taxonomicName>
was described from Drás [= Dras], Kárgil [Kargil], and Leh in Ladakh (
<bibRefCitation id="05324B240B46FFE8166BFE80F8C89E18" author="SMITH F." box="[333,489,333,354]" pageId="12" pageNumber="359" pagination="1 - 22" refId="ref7315" refString="SMITH F. (1878): Hymenoptera. - In: Scientific results of the second Yarkand Mission, based upon the collections and notes of the late Ferdinand Stoliczka. By order of the Government of India. 1878 - 91. pp. 1 - 22. Calcutta (government printing)." type="book chapter" year="1878">SMITH 1878: 2</bibRefCitation>
). It was described only in the male sex, with the flight period given as August-September.
<bibRefCitation id="05324B240B46FFE81539FEA7FAB59E05" author="GUSENLEITNER F. &amp; M. SCHWARZ" box="[543,916,362,383]" pageId="12" pageNumber="359" pagination="1 - 1280" refId="ref6875" refString="GUSENLEITNER F. &amp; M. SCHWARZ (2002): Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). - Entomofauna. Supplement 10: 1 - 1280." type="journal article" year="2002">GUSENLEITNER &amp; SCHWARZ (2002)</bibRefCitation>
gave the
<typeStatus id="BE1888770B46FFE81324FEA6FD0C9EFA" box="[1026,1069,363,384]" pageId="12" pageNumber="359">type</typeStatus>
depository as the NHMUK, but no specimens or record of
<taxonomicName id="A6A34D560B46FFE81436FE45FAA99EE7" authorityName="SMITH" authorityYear="1878" box="[784,904,392,413]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="floridula">
<emphasis id="53D7EAC70B46FFE81436FE45FAA99EE7" box="[784,904,392,413]" italics="true" pageId="12" pageNumber="359">A. floridula</emphasis>
</taxonomicName>
could be found there. Donald Baker (unpublished Phd thesis) suggested that
<typeStatus id="BE1888770B46FFE8142BFE68FA189EC0" box="[781,825,421,442]" pageId="12" pageNumber="359">type</typeStatus>
material could be in the Zoological Survey of
<collectingCountry id="19B476450B46FFE816A6FE0FF8959EA2" box="[384,436,450,472]" name="India" pageId="12" pageNumber="359">India</collectingCountry>
collection in Kolkata which is currently unavailable for study. In any case, although the female was not previously known, the contemporary specimen from Leh is unmistakably
<taxonomicName id="A6A34D560B46FFE81696FE30FB049D68" authorityName="SMITH" authorityYear="1878" box="[432,549,509,530]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="floridula">
<emphasis id="53D7EAC70B46FFE81696FE30FB049D68" box="[432,549,509,530]" italics="true" pageId="12" pageNumber="359">A. floridula</emphasis>
</taxonomicName>
based on the description, illustration (
<bibRefCitation id="05324B240B46FFE8148AFE30FD0B9D68" author="SMITH F." box="[940,1066,509,530]" pageId="12" pageNumber="359" pagination="1 - 22" refId="ref7315" refString="SMITH F. (1878): Hymenoptera. - In: Scientific results of the second Yarkand Mission, based upon the collections and notes of the late Ferdinand Stoliczka. By order of the Government of India. 1878 - 91. pp. 1 - 22. Calcutta (government printing)." type="book chapter" year="1878">SMITH 1878</bibRefCitation>
: figure 4, as
<taxonomicName id="A6A34D560B46FFE8163CFDD6F8AF9D55" authorityName="Smith" authorityYear="1878" box="[282,398,538,560]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="floricula">
<emphasis id="53D7EAC70B46FFE8163CFDD6F8AF9D55" box="[282,398,538,560]" italics="true" pageId="12" pageNumber="359">A. floricula</emphasis>
</taxonomicName>
sic), collecting locality, and flight period.
</paragraph>
<paragraph id="611C36D50B46FFE81785FDF2F8E69C59" blockId="12.[163,1070,575,803]" pageId="12" pageNumber="359">
Although Smith writes that the specimens he described were males (&quot;Mas.-&quot;), the specimen pictured in his figure 4 actually appears to be a female, as there appear to be scopal hairs present on the hind tibiae. In any case, the specimen matches the description with regards to the key criteria which are that the hind legs are covered with silvery pubescence (
<figureCitation id="F9982A500B46FFE81602FD78F84F9DB0" box="[292,366,693,715]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 42</figureCitation>
) and the ovate metasoma has the apical margins testaceous with a thin fringe of whitish pubescence (
<figureCitation id="F9982A500B46FFE816C3FD1FFB149D9D" box="[485,565,722,744]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 43</figureCitation>
). Indeed, the terga have not only their margins lightened white-hyaline, but the transitional area between the margin and the tergal discs is lightened dark orange-red.
</paragraph>
<paragraph id="611C36D50B46FFE81785FCFFFAF69BBC" blockId="12.[163,1070,818,1222]" pageId="12" pageNumber="359">
Examination of this contemporary specimen shows that it presents an unusual combination of characters that defies classification within the existing subgeneric system. The facial foveae are relatively narrow and impressed (
<figureCitation id="F9982A500B46FFE81546FCA0FBF59CF8" box="[608,724,877,899]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Figs 37; 39</figureCitation>
), slightly narrowing ventrally (but not as clearly as in members of the subgenus
<taxonomicName id="A6A34D560B46FFE815ACFC47FA879CE5" authority="HEDICKE, 1933" authorityName="HEDICKE" authorityYear="1933" box="[650,934,906,928]" pageId="12" pageNumber="359" rank="subGenus" subGenus="Euandrena">
<emphasis id="53D7EAC70B46FFE815ACFC47FBDA9CE5" box="[650,763,906,927]" italics="true" pageId="12" pageNumber="359">Euandrena</emphasis>
HEDICKE, 1933
</taxonomicName>
), the head is almost round in frontal view (only 1.1 times wider than long,
<figureCitation id="F9982A500B46FFE81406FC65FA4A9CC7" box="[800,875,936,958]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 37</figureCitation>
), the ocelloccipital distance exceeds two times the diameter of a lateral ocellus (
<figureCitation id="F9982A500B46FFE81424FC08FA6C9CA0" box="[770,845,965,987]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 39</figureCitation>
), with the vertex itself being clearly depressed at the meeting point with the two lateral ocelli (similar to the condition in the subgenus
<taxonomicName id="A6A34D560B46FFE816EBFBCDFA959B6F" authority="HIRASHIMA &amp; TADAUCHI, 1975" authorityName="HIRASHIMA &amp; TADAUCHI" authorityYear="1975" box="[461,948,1023,1045]" pageId="12" pageNumber="359" rank="subGenus" subGenus="Oreomelissa">
<emphasis id="53D7EAC70B46FFE816EBFBCDFB6C9B6F" box="[461,589,1024,1045]" italics="true" pageId="12" pageNumber="359">Oreomelissa</emphasis>
HIRASHIMA &amp; TADAUCHI, 1975
</taxonomicName>
and allied subgenera), the head and mesosoma show weak metallic green reflections (
<figureCitation id="F9982A500B46FFE81488FBD0F99E9B35" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Figs 37; 39- 40</figureCitation>
), the propodeal triangle is poorly delineated but weak lateral carinae and some raised rugae are present laterally and basally (
<figureCitation id="F9982A500B46FFE8151EFB95FBA59B17" box="[568,644,1112,1134]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 41</figureCitation>
), the scopal hairs are weakly plumose on the hind femorae and less so on the hind tibiae (
<figureCitation id="F9982A500B46FFE815A3FBB8FBF19BF0" box="[645,720,1141,1163]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 42</figureCitation>
), and there are no other distinctive characters. Without available male specimens (as the genital capsule is unknown) or any genetic data, it is not possible to confidently place the species within a subgenus.
</paragraph>
<paragraph id="611C36D50B46FFE91785FB18F9DE9E5A" blockId="12.[163,1071,1237,1582]" lastBlockId="13.[163,1070,208,288]" lastPageId="13" lastPageNumber="360" pageId="12" pageNumber="359">
D e s c r i p t i o n. Female. Body length:
<quantity id="A65B9B300B46FFE81564FB18FB5A9B90" box="[578,635,1237,1258]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="359" unit="mm" value="9.0">9 mm</quantity>
(
<figureCitation id="F9982A500B46FFE815AEFB18FBF29B91" box="[648,723,1237,1259]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 36</figureCitation>
). Head: Dark, 1.1 times wider than long (
<figureCitation id="F9982A500B46FFE817F8FB3FF80B9A7D" box="[222,298,1266,1288]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 37</figureCitation>
), integument with weak metallic green reflections most visible on gena and vertex. Clypeus weakly domed, irregularly punctate, punctures basally and laterally separated by 0.5 puncture diameters, medially becoming scattered, separated by 1-3 puncture dimeters, underlying surface polished and shining. Process of labrum broadly trapezoidal, 2 times wider than long, apical margin with extremely deep rounded emargination, emargination reaching ½ of visible length (
<figureCitation id="F9982A500B46FFE815C7FA48FA0A9AE0" box="[737,811,1413,1435]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 38</figureCitation>
). Gena slightly exceeding width of compound eye; ocelloccipital distance 2 times diameter of lateral ocellus (
<figureCitation id="F9982A500B46FFE81323FA6FF99E9AAF" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="12" pageNumber="359">Fig. 39</figureCitation>
); vertex slightly but distinctly depressed at level of posterior margin of lateral ocelli. Foveae dorsally occupying slightly less than ½ space between lateral ocellus and compound eye, narrowly slightly ventrally, ventrally extending to slightly below lower margin of antennal insertions; foveae filled with light brown hairs. Face, gena, vertex, and scape covered with pale whitish pubescence. Antennae basally dark, A4-12 ventrally lightened by presence of brownish scales; A3 slightly exceeding A4+5, shorter than A4+5+6.
</paragraph>
<paragraph id="611C36D50B47FFE91785FEFDF89D9D84" blockId="13.[163,1070,303,766]" pageId="13" pageNumber="360">
Mesosoma: Scutum predominantly polished and shining, weakly shagreened anteriorly, surface irregularly punctate, punctures separated by 0.5-2 puncture diameters; surface with subtle metallic reflections laterally; scutellum more strongly polished and shining and more sparsely punctate, punctures separated by 0.5-4 puncture diameters (
<figureCitation id="F9982A500B47FFE914E9FE45FD029EE7" box="[975,1059,392,414]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="13" pageNumber="360">Fig. 40</figureCitation>
). Pronotum with weak but distinct humeral angle, humeral angle with longitudinal carina. Mesepisternum and dorsolateral parts of propodeum microreticulate, dull, microreticulation overlain by raised network of rugosity; propodeal triangle laterally delineated by fine carina, internal surface slightly depressed, covered with fine network of raised carinae over majority of surface (
<figureCitation id="F9982A500B47FFE9151AFDD7FBA89D55" box="[572,649,538,560]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="13" pageNumber="360">Fig. 41</figureCitation>
). Mesosoma covered with pale hairs, not exceeding length of scape. Propodeal corbicula incomplete, dorsal fringe composed of long plumose pale hairs, internal surface with abundant plumose pale hairs. Legs dark, pubescence light brown. Flocculus complete, strongly produced, composed of white plumose hairs; femoral and tibial scopae composed of bright white-silvery hairs, hairs of femoral scopae somewhat plumose, hairs of tibial scopae obscurely plumose (
<figureCitation id="F9982A500B47FFE914F5FD60FD029DB8" box="[979,1059,685,707]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="13" pageNumber="360">Fig. 42</figureCitation>
). Hind tarsal claws with small inner tooth. Wings hyaline, stigma and venation dark brown, nervulus weakly antefurcal.
</paragraph>
<paragraph id="611C36D50B47FFE91785FCC0FAC89C8A" blockId="13.[163,1071,781,1008]" pageId="13" pageNumber="360">
Metasoma: Tergal discs dark, marginal areas broadly lightened hyaline orange-brown, apical rims lightened hyaline whitish (
<figureCitation id="F9982A500B47FFE9156DFCE7FBBE9C45" box="[587,671,810,832]" captionStart="Figs 36-43" captionStartId="11.[163,203,1491,1513]" captionTargetBox="[184,1041,194,1479]" captionTargetId="figure-67@11.[184,1041,194,1479]" captionTargetPageId="11" captionText="Figs 36-43: Andrena (incertae sedis) floridula SMITH, 1878 female. (36) Habitus, profile view; (37) face, frontal view; (38) process of labrum, frontal view; (39) head, dorsal view; (40) scutum, dorsal view; (41) propodeum, dorsal view; (42) hind leg, profile view; (43) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681705" httpUri="https://zenodo.org/record/14681705/files/figure.png" pageId="13" pageNumber="360">Fig. 43</figureCitation>
). Tergal discs polished and shining, irregularly punctate, punctures separated by 0.5-3 puncture diameters, most dense basally, becoming weaker apically onto tergal margins. Tergal discs with scattered pale pubescence, tergal margins with pubescence longer and forming distinct hairbands on T2- 4, weakly interrupted on T2, complete on T3-4, obscuring underlying surface laterally. Apical fringe of T5 and hairs flanking pygidial plate orange-brown. Pygidial plate rounded triangular, lateral margins slightly raised, internal surface flat and dull, featureless.
</paragraph>
<paragraph id="611C36D50B47FFE91785FBCDFB859B6F" blockId="13.[163,676,1023,1045]" box="[163,676,1023,1045]" pageId="13" pageNumber="360">
D i s t r i b u t i o n:
<collectingCountry id="19B476450B47FFE91653FC32F8889B6F" box="[373,425,1023,1045]" name="India" pageId="13" pageNumber="360">India</collectingCountry>
(Ladakh) (
<bibRefCitation id="05324B240B47FFE9153DFBCDFBB99B6F" author="SMITH F." box="[539,664,1024,1045]" pageId="13" pageNumber="360" pagination="1 - 22" refId="ref7315" refString="SMITH F. (1878): Hymenoptera. - In: Scientific results of the second Yarkand Mission, based upon the collections and notes of the late Ferdinand Stoliczka. By order of the Government of India. 1878 - 91. pp. 1 - 22. Calcutta (government printing)." type="book chapter" year="1878">SMITH 1878</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="DCD3A4B5B31DF6EEA978E2B4D193EBC4" type="Zenodo-Dep">14681691</mods:identifier>
</mods:mods>
<treatment id="E90A87C30B4DFFE31785FE9FFA439C57" ID-DOI="http://doi.org/10.5281/zenodo.14681695" ID-Zenodo-Dep="14681695" LSID="urn:lsid:plazi:treatment:E90A87C30B4DFFE31785FE9FFA439C57" httpUri="http://treatment.plazi.org/id/E90A87C30B4DFFE31785FE9FFA439C57" lastPageNumber="354" pageId="7" pageNumber="354">
<subSubSection id="29B9655E0B4DFFE31785FE9FFAE59EC4" pageId="7" pageNumber="354" type="nomenclature">
<paragraph id="611C36D50B4DFFE31785FE9FFAA69E12" blockId="7.[163,903,338,360]" box="[163,903,338,360]" pageId="7" pageNumber="354">
<heading id="3A5481B90B4DFFE31785FE9FFAA69E12" box="[163,903,338,360]" fontSize="8" level="2" pageId="7" pageNumber="354" reason="2">
<taxonomicName id="A6A34D560B4DFFE31785FE9FFB999E12" ID-CoL="DSBF" authority="MORAWITZ, 1876" authorityName="MORAWITZ" authorityYear="1876" box="[163,696,338,360]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="mordax" subGenus="Hoplandrena">
<emphasis id="53D7EAC70B4DFFE31785FE9FFB599E1D" bold="true" box="[163,632,338,360]" pageId="7" pageNumber="354">
<emphasis id="53D7EAC70B4DFFE31785FE9FF9DF9E12" bold="true" box="[163,254,338,360]" italics="true" pageId="7" pageNumber="354">Andrena</emphasis>
(
<emphasis id="53D7EAC70B4DFFE3162BFE9FF8B99E12" bold="true" box="[269,408,338,360]" italics="true" pageId="7" pageNumber="354">Hoplandrena</emphasis>
)
<emphasis id="53D7EAC70B4DFFE31681FE9FF8D69E12" bold="true" box="[423,503,338,360]" italics="true" pageId="7" pageNumber="354">mordax</emphasis>
MORAWITZ
</emphasis>
,
<emphasis id="53D7EAC70B4DFFE315A3FE9EFB999E12" bold="true" box="[645,696,339,360]" pageId="7" pageNumber="354">1876</emphasis>
</taxonomicName>
<emphasis id="53D7EAC70B4DFFE31598FE9FFAA69E12" bold="true" box="[702,903,338,360]" pageId="7" pageNumber="354">
(
<figureCitation id="F9982A500B4DFFE315E3FE9FFA199E12" box="[709,824,338,360]" captionStart="Figs 17-21" captionStartId="8.[163,203,1500,1522]" captionTargetBox="[166,1072,199,1487]" captionTargetId="figure-53@8.[166,1072,199,1487]" captionTargetPageId="8" captionText="Figs 17-21: (17) Collecting site of Rumbak at 3850 m; (18-19) Andrena mordax MORAWITZ, 1876 female at Mentha flowers (Lamiaceae; photographs taken at Rumbak); (20-21) Andrena floridula SMITH, 1878 female at Asteraceae flowers (photographs taken in Leh gardens; photographs: M. Jacobs)." figureDoi="http://doi.org/10.5281/zenodo.14681699" httpUri="https://zenodo.org/record/14681699/files/figure.png" pageId="7" pageNumber="354">Figs 18-19</figureCitation>
;
<figureCitation id="F9982A500B4DFFE31462FE9FFAA09E12" box="[836,897,338,360]" captionStart="Figs 22-27" captionStartId="9.[163,203,1228,1250]" captionTargetBox="[166,1073,197,1214]" captionTargetId="figure-134@9.[166,1073,197,1214]" captionTargetPageId="9" captionText="Figs 22-27: Andrena (Hoplandrena) mordax MORAWITZ, 1876. (22) Female habitus, lateral view; (23) male habitus, lateral view; (24) female face, frontal view; (25) male face, frontal view; (26) female terga, dorsal view; (27) male genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681701" httpUri="https://zenodo.org/record/14681701/files/figure.png" pageId="7" pageNumber="354">22-27</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="611C36D50B4DFFE31785FEB4FAE59EC4" blockId="7.[163,1071,376,446]" pageId="7" pageNumber="354">
<materialsCitation id="D1CB3C880B4DFFE31785FEB4FA829EDC" collectingDate="1975-08-18" collectingDateMax="1975-08-25" collectingDateMin="1975-08-18" collectorName="H. Huss &amp; Biologiezentrum Linz" country="Afghanistan" elevation="3500" location="GrosserPamir" municipality="Issiktal" pageId="7" pageNumber="354" specimenCount="9" specimenCount-female="9">
M a t e r i a l e x a m i n e d:
<collectingCountry id="19B476450B4DFFE316C1FEB5FB7D9EF4" box="[487,604,376,398]" name="Afghanistan" pageId="7" pageNumber="354">Afghanistan</collectingCountry>
:
<location id="647C600E0B4DFFE31579FEB5FBC59EF4" LSID="urn:lsid:plazi:treatment:E90A87C30B4DFFE31785FE9FFA439C57:647C600E0B4DFFE31579FEB5FBC59EF4" box="[607,740,376,398]" country="Afghanistan" municipality="Issiktal" name="GrosserPamir" pageId="7" pageNumber="354">GrosserPamir</location>
,
<collectingMunicipality id="8178ACAF0B4DFFE315C1FEB5FA0A9EF4" box="[743,811,376,398]" pageId="7" pageNumber="354">Issiktal</collectingMunicipality>
[Isikmountain, Isik, Wakhan Corridor],
<quantity id="A65B9B300B4DFFE3160BFE5DF8529EDC" box="[301,371,400,422]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="354" unit="m" value="3500.0">
<elevation id="EA8ED1E60B4DFFE3160BFE5DF8529EDC" box="[301,371,400,422]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="354" unit="m" value="3500.0">3500 m</elevation>
</quantity>
,
<date id="151D10150B4DFFE31658FE5DFB279EDC" box="[382,518,400,422]" pageId="7" pageNumber="354" value="1975-08-18" valueMax="1975-08-25" valueMin="1975-08-18">
<collectingDate id="0559E9FD0B4DFFE31658FE5DFB279EDC" box="[382,518,400,422]" pageId="7" pageNumber="354" value="1975-08-18" valueMax="1975-08-25" valueMin="1975-08-18">18-25.viii.1975</collectingDate>
</date>
,
<specimenCount id="77A5FD5C0B4DFFE31529FE5CFB0D9EDC" box="[527,556,401,422]" count="9" pageId="7" pageNumber="354" type="female">9♀</specimenCount>
, leg.
<collectorName id="CC5653030B4DFFE3157BFE5CFB899EDC" box="[605,680,401,422]" pageId="7" pageNumber="354">H. Huss</collectorName>
, coll.
<collectorName id="CC5653030B4DFFE315F9FE5DFA829EDC" box="[735,931,400,422]" pageId="7" pageNumber="354">Biologiezentrum Linz</collectorName>
</materialsCitation>
;
<materialsCitation id="D1CB3C880B4DFFE3148BFE5DFAE59EC4" collectingDate="2015-09-01" collectorName="M. Jacobs &amp; Maarten Jacobs." country="India" county="Rumbak" elevation="3850" location="Ladakh" municipality="Leh Tehsil" pageId="7" pageNumber="354" specimenCount="2" specimenCount-female="2">
<collectingCountry id="19B476450B4DFFE3148BFE5DFAFC9EDC" box="[941,989,400,422]" name="India" pageId="7" pageNumber="354">India</collectingCountry>
:
<location id="647C600E0B4DFFE314C1FE5DFD0A9EDC" LSID="urn:lsid:plazi:treatment:E90A87C30B4DFFE31785FE9FFA439C57:647C600E0B4DFFE314C1FE5DFD0A9EDC" box="[999,1067,400,422]" country="India" county="Rumbak" municipality="Leh Tehsil" name="Ladakh" pageId="7" pageNumber="354">Ladakh</location>
,
<collectingMunicipality id="8178ACAF0B4DFFE317E9FE65F8119EC4" box="[207,304,424,446]" pageId="7" pageNumber="354">Leh Tehsil</collectingMunicipality>
,
<collectingCounty id="887D4E590B4DFFE3161EFE65F8A49EC4" box="[312,389,424,446]" pageId="7" pageNumber="354">Rumbak</collectingCounty>
,
<quantity id="A65B9B300B4DFFE316ABFE65F8F09EC4" box="[397,465,424,446]" metricMagnitude="3" metricUnit="m" metricValue="3.85" pageId="7" pageNumber="354" unit="m" value="3850.0">
<elevation id="EA8ED1E60B4DFFE316ABFE65F8F09EC4" box="[397,465,424,446]" metricMagnitude="3" metricUnit="m" metricValue="3.85" pageId="7" pageNumber="354" unit="m" value="3850.0">3850 m</elevation>
</quantity>
,
<date id="151D10150B4DFFE316FFFE65FB0C9EC4" box="[473,557,424,446]" pageId="7" pageNumber="354" value="2015-09-01">
<collectingDate id="0559E9FD0B4DFFE316FFFE65FB0C9EC4" box="[473,557,424,446]" pageId="7" pageNumber="354" value="2015-09-01">1.ix.2015</collectingDate>
</date>
, 1♁,
<specimenCount id="77A5FD5C0B4DFFE3157AFE64FB599EC4" box="[604,632,425,446]" count="2" pageId="7" pageNumber="354" type="female">2♀</specimenCount>
, leg.
<collectorName id="CC5653030B4DFFE31581FE64FA209EC4" box="[679,769,424,446]" pageId="7" pageNumber="354">M. Jacobs</collectorName>
, coll.
<collectorName id="CC5653030B4DFFE31413FE64FAE59EC4" box="[821,964,424,446]" pageId="7" pageNumber="354">Maarten Jacobs.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="29B9655E0B4DFFE31784FE00FA439C57" pageId="7" pageNumber="354" type="description">
<paragraph id="611C36D50B4DFFE31784FE00FA9D9D91" blockId="7.[162,1070,461,747]" pageId="7" pageNumber="354">
R e m a r k s:
<taxonomicName id="A6A34D560B4DFFE31613FE00F8C09E98" authorityName="MORAWITZ" authorityYear="1876" box="[309,481,461,482]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="mordax">
<emphasis id="53D7EAC70B4DFFE31613FE00F8C09E98" box="[309,481,461,482]" italics="true" pageId="7" pageNumber="354">Andrena mordax</emphasis>
</taxonomicName>
is an uncommon but typical species of Central Asia. The
<typeStatus id="BE1888770B4DFFE31785FE26F9EE9D7A" box="[163,207,491,512]" pageId="7" pageNumber="354">type</typeStatus>
material (a male) was illustrated by
<bibRefCitation id="05324B240B4DFFE3156AFE27FA7E9E85" author="ASTAFUROVA Y. A. &amp; PROSHCHALYKIN M. Y. &amp; D. A. SIDOROV" box="[588,863,490,512]" pageId="7" pageNumber="354" pagination="105 - 176" refId="ref6439" refString="ASTAFUROVA Y. A., PROSHCHALYKIN M. Y. &amp; D. A. SIDOROV (2022): The bees of the genus Andrena FABRICIUS, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. - Zookeys. 1120: 105 - 176." type="journal article" year="2022">ASTAFUROVA et al. (2022)</bibRefCitation>
, and differs slightly from the male pictured here (
<figureCitation id="F9982A500B4DFFE316E3FDC5FB419D67" box="[453,608,519,542]" captionStart="Figs 22-27" captionStartId="9.[163,203,1228,1250]" captionTargetBox="[166,1073,197,1214]" captionTargetId="figure-134@9.[166,1073,197,1214]" captionTargetPageId="9" captionText="Figs 22-27: Andrena (Hoplandrena) mordax MORAWITZ, 1876. (22) Female habitus, lateral view; (23) male habitus, lateral view; (24) female face, frontal view; (25) male face, frontal view; (26) female terga, dorsal view; (27) male genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681701" httpUri="https://zenodo.org/record/14681701/files/figure.png" pageId="7" pageNumber="354">Figs 23; 25; 25</figureCitation>
), specifically due to the genital capsule which has the penis valves comparatively slightly wider basally, whereas they are slightly more narrow in the Indian specimen (
<figureCitation id="F9982A500B4DFFE316C9FD8FFB1C9D2D" box="[495,573,578,600]" captionStart="Figs 22-27" captionStartId="9.[163,203,1228,1250]" captionTargetBox="[166,1073,197,1214]" captionTargetId="figure-134@9.[166,1073,197,1214]" captionTargetPageId="9" captionText="Figs 22-27: Andrena (Hoplandrena) mordax MORAWITZ, 1876. (22) Female habitus, lateral view; (23) male habitus, lateral view; (24) female face, frontal view; (25) male face, frontal view; (26) female terga, dorsal view; (27) male genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681701" httpUri="https://zenodo.org/record/14681701/files/figure.png" pageId="7" pageNumber="354">Fig. 27</figureCitation>
). Otherwise, the specimens conform well to the concept of
<taxonomicName id="A6A34D560B4DFFE31637FDADF8599D0F" authorityName="MORAWITZ" authorityYear="1876" box="[273,376,608,629]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="mordax">
<emphasis id="53D7EAC70B4DFFE31637FDADF8599D0F" box="[273,376,608,629]" italics="true" pageId="7" pageNumber="354">A. mordax</emphasis>
</taxonomicName>
in both sexes (
<figureCitation id="F9982A500B4DFFE3152DFDADFB5A9D0F" box="[523,635,608,630]" captionStart="Figs 22-27" captionStartId="9.[163,203,1228,1250]" captionTargetBox="[166,1073,197,1214]" captionTargetId="figure-134@9.[166,1073,197,1214]" captionTargetPageId="9" captionText="Figs 22-27: Andrena (Hoplandrena) mordax MORAWITZ, 1876. (22) Female habitus, lateral view; (23) male habitus, lateral view; (24) female face, frontal view; (25) male face, frontal view; (26) female terga, dorsal view; (27) male genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14681701" httpUri="https://zenodo.org/record/14681701/files/figure.png" pageId="7" pageNumber="354">Figs 22-27</figureCitation>
;
<bibRefCitation id="05324B240B4DFFE315A2FDADFA5B9D0F" author="OSYTSHNJUK A. Z. &amp; ROMASENKO L. &amp; BANASZAK J. &amp; E. MOTYKA" box="[644,890,608,630]" pageId="7" pageNumber="354" refId="ref7095" refString="OSYTSHNJUK A. Z., ROMASENKO L., BANASZAK J. &amp; E. MOTYKA (2008): Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, Poland, 233 pp." type="book" year="2008">OSYTSHNJUK et al. 2008</bibRefCitation>
). Specimens were observed foraging on
<emphasis id="53D7EAC70B4DFFE316A6FDB0F8EC9DE8" box="[384,461,637,658]" italics="true" pageId="7" pageNumber="354">Mentha</emphasis>
(
<taxonomicName id="A6A34D560B4DFFE316FCFDB0FB6A9DE9" box="[474,587,637,659]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="354" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
) at the Rumbak site (
<figureCitation id="F9982A500B4DFFE31405FDB0FAB59DE8" box="[803,916,637,659]" captionStart="Figs 17-21" captionStartId="8.[163,203,1500,1522]" captionTargetBox="[166,1072,199,1487]" captionTargetId="figure-53@8.[166,1072,199,1487]" captionTargetPageId="8" captionText="Figs 17-21: (17) Collecting site of Rumbak at 3850 m; (18-19) Andrena mordax MORAWITZ, 1876 female at Mentha flowers (Lamiaceae; photographs taken at Rumbak); (20-21) Andrena floridula SMITH, 1878 female at Asteraceae flowers (photographs taken in Leh gardens; photographs: M. Jacobs)." figureDoi="http://doi.org/10.5281/zenodo.14681699" httpUri="https://zenodo.org/record/14681699/files/figure.png" pageId="7" pageNumber="354">Figs 18-19</figureCitation>
). In addition to reporting
<taxonomicName id="A6A34D560B4DFFE31623FD56F84C9DD5" authorityName="MORAWITZ" authorityYear="1876" box="[261,365,666,688]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="mordax">
<emphasis id="53D7EAC70B4DFFE31623FD56F84C9DD5" box="[261,365,666,688]" italics="true" pageId="7" pageNumber="354">A. mordax</emphasis>
</taxonomicName>
as new for
<collectingCountry id="19B476450B4DFFE316C7FD57FB399DCA" box="[481,536,666,688]" name="India" pageId="7" pageNumber="354">India</collectingCountry>
, the species is reported from
<collectingCountry id="19B476450B4DFFE3141DFD57FA969DCA" box="[827,951,666,688]" name="Afghanistan" pageId="7" pageNumber="354">Afghanistan</collectingCountry>
for the first time from the Wakhan Corridor (Isik mountain, not to be confused with lake Issyk Kul in
<collectingCountry id="19B476450B4DFFE31785FD18F83D9D90" box="[163,284,725,746]" name="Kyrgyzstan" pageId="7" pageNumber="354">Kyrgyzstan</collectingCountry>
), linking populations in Central Asia proper with those in Ladakh.
</paragraph>
<paragraph id="611C36D50B4DFFE31785FD36FA439C57" blockId="7.[163,1070,762,814]" pageId="7" pageNumber="354">
D i s t r i b u t i o n:
<collectingCountry id="19B476450B4DFFE31652FD37F8CB9C6A" box="[372,490,762,784]" name="Kazakhstan" pageId="7" pageNumber="354">Kazakhstan</collectingCountry>
,
<collectingCountry id="19B476450B4DFFE316D5FD37FB449C6A" box="[499,613,762,784]" name="Uzbekistan" pageId="7" pageNumber="354">Uzbekistan</collectingCountry>
,
<collectingCountry id="19B476450B4DFFE31548FD37FBF59C6A" box="[622,724,762,784]" name="Tajikistan" pageId="7" pageNumber="354">Tajikistan</collectingCountry>
,
<collectingCountry id="19B476450B4DFFE315FBFD36FA739C6A" box="[733,850,763,784]" name="Kyrgyzstan" pageId="7" pageNumber="354">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="19B476450B4DFFE3147DFD37FAB89C6A" box="[859,921,762,784]" name="China" pageId="7" pageNumber="354">China</collectingCountry>
,
<collectingCountry id="19B476450B4DFFE31484FD37FD009C6A" box="[930,1057,762,784]" name="Afghanistan" pageId="7" pageNumber="354">Afghanistan</collectingCountry>
*,
<collectingCountry id="19B476450B4DFFE31785FCD5F9FB9C54" box="[163,218,792,814]" name="India" pageId="7" pageNumber="354">India</collectingCountry>
* (Ladakh) (
<bibRefCitation id="05324B240B4DFFE31673FCD5FB6F9C57" author="OSYTSHNJUK A. Z. &amp; ROMASENKO L. &amp; BANASZAK J. &amp; E. MOTYKA" box="[341,590,792,814]" pageId="7" pageNumber="354" refId="ref7095" refString="OSYTSHNJUK A. Z., ROMASENKO L., BANASZAK J. &amp; E. MOTYKA (2008): Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, Poland, 233 pp." type="book" year="2008">OSYTSHNJUK et al. 2008</bibRefCitation>
;
<bibRefCitation id="05324B240B4DFFE3157EFCD5FA769C57" author="ASTAFUROVA Y. A. &amp; PROSHCHALYKIN M. Y. &amp; D. A. SIDOROV" box="[600,855,792,814]" pageId="7" pageNumber="354" pagination="105 - 176" refId="ref6439" refString="ASTAFUROVA Y. A., PROSHCHALYKIN M. Y. &amp; D. A. SIDOROV (2022): The bees of the genus Andrena FABRICIUS, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. - Zookeys. 1120: 105 - 176." type="journal article" year="2022">ASTAFUROVA et al. 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>