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<mods:title id="3392993AC32A9F9C0A441B89D2928F15">Three new species in Calamus sect. Podocephalus (Arecaceae: Calamoideae) from the Philippines, Indonesia, and Papua New Guinea</mods:title>
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<mods:title id="F3B9D257A4C7725FDD6980FC9DB39D14">Three new species in Calamus sect. Podocephalus (Arecaceae: Calamoideae) from the Philippines, Indonesia, and Papua New Guinea</mods:title>
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<treatment id="038387EF23351C64ACD023B7FB24FCA3" ID-DOI="http://doi.org/10.5281/zenodo.15212843" ID-Zenodo-Dep="15212843" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3" httpUri="http://treatment.plazi.org/id/038387EF23351C64ACD023B7FB24FCA3" lastPageId="3" lastPageNumber="72" pageId="1" pageNumber="70">
<subSubSection id="C330657223351C66ACD023B7FD00FF6D" box="[151,675,222,247]" pageId="1" pageNumber="70" type="nomenclature">
<treatment id="038387EF23351C64ACD023B7FB24FCA3" ID-DOI="http://doi.org/10.5281/zenodo.15212843" ID-Zenodo-Dep="15212843" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3" httpUri="http://treatment.plazi.org/id/038387EF23351C64ACD023B7FB24FCA3" lastPageId="3" lastPageNumber="72" pageId="1" pageNumber="70" scope_family="Arecaceae">
<paragraph id="8B9536F923351C66ACD023B7FD00FF6D" blockId="1.[151,1436,222,319]" box="[151,675,222,247]" pageId="1" pageNumber="70">
<taxonomicName id="4C2A4D7A23351C66ACD023B7FDE2FF6C" authority="Fernando" authorityName="Fernando. A, Leaf" authorityYear="2014" box="[151,577,222,247]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="70" phylum="Chordata" rank="species" species="daemonoropoides" status="sp. nov.">
<emphasis id="B95EEAEB23351C66ACD023B7FE6DFF6C" bold="true" box="[151,462,222,247]" italics="true" pageId="1" pageNumber="70">Calamus daemonoropoides</emphasis>
@ -51,19 +50,20 @@ Fernando
<taxonomicNameLabel id="A26D579023351C66AE0B23B6FD00FF6D" box="[588,675,223,246]" pageId="1" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C330657223351C66AEEE23B7FE75FEA4" pageId="1" pageNumber="70" type="materials_examined">
<paragraph id="8B9536F923351C66AEEE23B7FE75FEA4" blockId="1.[151,1436,222,319]" pageId="1" pageNumber="70">
<collectingCounty id="62F44E7523351C66AEEE23B7FD40FF6C" box="[681,739,222,247]" pageId="1" pageNumber="70">Type</collectingCounty>
:—
<materialsCitation id="3B423CA423351C66AF4423B7FEEAFEA4" collectingDate="1984-03-02" collectionCode="LBC" collectorName="Fernando" country="Philippines" elevation="230" location="Quezon National Park" municipality="Atimonan" pageId="1" pageNumber="70" specimenCount="1" stateProvince="Quezon Province" typeStatus="holotype">
<materialsCitation id="3B423CA423351C66AF4423B7FEEAFEA4" pageId="1" pageNumber="70">
<collectingCountry id="F33D766923351C66AF4423B7FC0EFF6C" box="[771,941,222,247]" name="Philippines" pageId="1" pageNumber="70">PHILIPPINES</collectingCountry>
. Luzon Island,
.
<collectingMunicipality id="6BF1AC8323351C66AFFF23B6FBEEFF6C" box="[952,1101,222,247]" pageId="1" pageNumber="70">Luzon Island</collectingMunicipality>
,
<collectingRegion id="49EEF81B23351C66A81F23B7FABFFF6C" box="[1112,1308,222,247]" country="Philippines" name="Quezon" pageId="1" pageNumber="70">Quezon Province</collectingRegion>
,
<collectingMunicipality id="6BF1AC8323351C66A96123B7FA3BFF6C" box="[1318,1432,222,247]" pageId="1" pageNumber="70">Atimonan</collectingMunicipality>
<location id="8EF5602223351C66A96123B7FA3BFF6C" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223351C66A96123B7FA3BFF6C" box="[1318,1432,222,247]" country="Philippines" county="Type" municipality="Luzon Island" name="Atimonan" pageId="1" pageNumber="70" stateProvince="Quezon">Atimonan</location>
,
<location id="8EF5602223351C66ACD0226BFE39FE80" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223351C66ACD0226BFE39FE80" box="[151,410,258,283]" country="Philippines" municipality="Atimonan" name="Quezon National Park" pageId="1" pageNumber="70" stateProvince="Quezon Province">Quezon National Park</location>
<collectingRegion id="49EEF81B23351C66ACD0226BFF53FE80" box="[151,240,258,283]" country="Philippines" name="Quezon" pageId="1" pageNumber="70">Quezon</collectingRegion>
<location id="8EF5602223351C66ACBF226BFE39FE80" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223351C66ACBF226BFE39FE80" box="[248,410,258,283]" country="Philippines" county="Type" municipality="Luzon Island" name="National Park" pageId="1" pageNumber="70" stateProvince="Quezon">National Park</location>
, in lowland evergreen rain forest on gentle slope,
<quantity id="4CD29B1C23351C66AFA7226BFB8FFE81" box="[992,1068,258,283]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="70" unit="m" value="230.0">
<elevation id="0007D1CA23351C66AFA7226BFB8FFE81" box="[992,1068,258,283]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="70" unit="m" value="230.0">230 m</elevation>
@ -73,25 +73,15 @@ Fernando
<collectingDate id="EFD0E9D123351C66A87E226BFB4FFE80" box="[1081,1260,258,283]" pageId="1" pageNumber="70" value="1984-03-02">02 March 1984</collectingDate>
</date>
,
<emphasis id="B95EEAEB23351C66A8BF226BFA3FFE80" box="[1272,1436,258,283]" italics="true" pageId="1" pageNumber="70">
<collectorName id="26DF532F23351C66A8BF226BFACAFE80" box="[1272,1385,258,283]" pageId="1" pageNumber="70">Fernando</collectorName>
386
</emphasis>
<emphasis id="B95EEAEB23351C66A8BF226BFA3FFE80" box="[1272,1436,258,283]" italics="true" pageId="1" pageNumber="70">Fernando 386</emphasis>
(
<typeStatus id="5491885B23351C66ACD9224FFEA3FEA4" box="[158,256,294,319]" pageId="1" pageNumber="70" type="holotype">holotype</typeStatus>
<collectionCode id="ED3BAE3C23351C66AD40224FFEE6FEA4" box="[263,325,294,319]" pageId="1" pageNumber="70">LBC</collectionCode>
!
LBC!
</materialsCitation>
,
<materialsCitation id="3B423CA423351C66AD14224FFE6BFEA4" box="[339,456,294,319]" collectingDate="1984-03-02" collectionCode="K" collectorName="Fernando" country="Philippines" elevation="230" location="Quezon National Park" municipality="Atimonan" pageId="1" pageNumber="70" specimenCount="1" stateProvince="Quezon Province" typeStatus="isotype">
<typeStatus id="5491885B23351C66AD14224FFE01FEA4" box="[339,418,294,319]" pageId="1" pageNumber="70">isotype</typeStatus>
<collectionCode id="ED3BAE3C23351C66ADEE224FFE67FEA4" box="[425,452,294,319]" pageId="1" pageNumber="70">K</collectionCode>
!
</materialsCitation>
<materialsCitation id="3B423CA423351C66AD14224FFE6BFEA4" box="[339,456,294,319]" pageId="1" pageNumber="70">isotype K!</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330657223351C66ACD02206FBDCFE7A" pageId="1" pageNumber="70" type="diagnosis">
<paragraph id="8B9536F923351C66ACD02206FBDCFE7A" blockId="1.[151,1436,367,481]" pageId="1" pageNumber="70">
Similar to
<taxonomicName id="4C2A4D7A23351C66AD4F2206FC20FE1E" authority="(Beccari 1902: 225) Beccari (1911: 232)" authorityName="Beccari" authorityPageNumber="232" authorityYear="1911" baseAuthorityName="Beccari" baseAuthorityPageNumber="225" baseAuthorityYear="1902" box="[264,899,367,389]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="70" phylum="Chordata" rank="species" species="erinaceus">
@ -109,8 +99,6 @@ long arranged along the margins and mid-costa on adaxial surface; the very promi
<quantity id="4CD29B1C23351C66ACAE22A2FEBFFE7B" box="[233,284,459,480]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="70" unit="cm" value="4.0">4 cm</quantity>
) and reflexed female rachilla, shorter bracts, and more dense fruits along the rachilla.
</paragraph>
</subSubSection>
<subSubSection id="C330657223351C66ACD02164FC93F975" pageId="1" pageNumber="70" type="description">
<paragraph id="8B9536F923351C66ACD02164FC93F975" blockId="1.[151,1437,525,1954]" pageId="1" pageNumber="70">
Clustering, moderately robust rattan, climbing to
<quantity id="4CD29B1C23351C66AEA62164FCBFFDBE" box="[737,796,525,550]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="70" unit="m" value="15.0">15 m</quantity>
@ -204,8 +192,6 @@ long, recurved, the 3 stigmatic lobes persistent in young fruits. Mature fruit y
<figureCitation id="13112A7C23351C66AE8725BCFCBDF975" box="[704,798,1749,1774]" captionStart="FIGURE 1" captionStartId="2.[151,245,1912,1934]" captionTargetBox="[211,1381,212,1878]" captionTargetId="figure-114@2.[189,1397,191,1891]" captionTargetPageId="2" captionText="FIGURE 1. Calamus daemonoropoides Fernando. A, Leaf sheath showing dense partial whorls of acicular spines and prominent knee with sharp bulge. B, Young, newly expanding leaf. C, Partial male inflorescence. D, Distal portion of young infructescence. E, Fruiting rachillae. Scale bar = 1 cm. A, Dransfield 5472 (LBC); B, Fernando 1732 (LBC); C, Fernando 386 (LBC); D, Yap 001 (PUH); E, Fernando 1824 (LBC). AC and E Photos by E.S. Fernando; D Photo by S.L. Yap." figureDoi="http://doi.org/10.5281/zenodo.5138218" httpUri="https://zenodo.org/record/5138218/files/figure.png" pageId="1" pageNumber="70">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330657223351C66AC812590FD30F8AD" pageId="1" pageNumber="70" type="distribution">
<paragraph id="8B9536F923351C66AC812590FD30F8AD" blockId="1.[151,1437,525,1954]" pageId="1" pageNumber="70">
<emphasis id="B95EEAEB23351C66AC812590FE54F889" bold="true" box="[198,503,1785,1810]" pageId="1" pageNumber="70">Distribution and habitat:</emphasis>
—Endemic to the
@ -216,8 +202,6 @@ long, recurved, the 3 stigmatic lobes persistent in young fruits. Mature fruit y
<quantity id="4CD29B1C23351C66AD902474FDBDF8AE" box="[471,542,1821,1846]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="70" unit="m" value="450.0">450 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="C330657223351C66AC812428FC81F8C1" box="[198,802,1857,1882]" pageId="1" pageNumber="70" type="vernacular_names">
<paragraph id="8B9536F923351C66AC812428FC81F8C1" blockId="1.[151,1437,525,1954]" box="[198,802,1857,1882]" pageId="1" pageNumber="70">
<emphasis id="B95EEAEB23351C66AC812428FE71F8C1" bold="true" box="[198,466,1857,1882]" pageId="1" pageNumber="70">Local names and uses:</emphasis>
@ -226,8 +210,6 @@ elevation.
<collectingRegion id="49EEF81B23351C66AE112428FCB4F8C1" box="[598,791,1857,1882]" country="Philippines" name="Isabela" pageId="1" pageNumber="70">Isabela Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C330657223351C66AC81240CFACCF839" pageId="1" pageNumber="70" type="etymology">
<paragraph id="8B9536F923351C66AC81240CFACCF839" blockId="1.[151,1437,525,1954]" pageId="1" pageNumber="70">
<emphasis id="B95EEAEB23351C66AC81240CFEF2F8E5" bold="true" box="[198,337,1893,1918]" pageId="1" pageNumber="70">Etymology:</emphasis>
—The epithet
@ -243,7 +225,6 @@ Blume
<bibRefCitation id="EFBB4B0823351C66AFBC24E0FAC1F839" author="Roemer, J. J. &amp; Schultes, J. A." box="[1019,1378,1929,1954]" pageId="1" pageNumber="70" refId="ref5455" refString="Roemer, J. J. &amp; Schultes, J. A. (1830). Systema Vegetabilium: Secundum Classes, Ordines, Genera, Species, ed. 15, Volume 7, Part 2, pp. 1725 - 1815. Sumtibus J. G. Cottae, Stuttgardtiae. http: // dx. doi. org / 10.5962 / bhl. title. 825" type="book" year="1830">Roemer &amp; Schultes 1830: 1333</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF55667123361C65ACD02411FB83F87E" ID-DOI="http://doi.org/10.5281/zenodo.5138218" ID-Zenodo-Dep="5138218" httpUri="https://zenodo.org/record/5138218/files/figure.png" pageId="2" pageNumber="71" startId="2.[151,245,1912,1934]" targetBox="[211,1381,212,1878]" targetPageId="2">
<paragraph id="8B9536F923361C65ACD02411FB83F87E" blockId="2.[151,1436,1912,2022]" pageId="2" pageNumber="71">
<emphasis id="B95EEAEB23361C65ACD02411FEADF815" bold="true" box="[151,270,1912,1934]" pageId="2" pageNumber="71">FIGURE 1.</emphasis>
@ -290,18 +271,19 @@ Photos by E.S. Fernando;
Photo by S.L. Yap.
</paragraph>
</caption>
<subSubSection id="C330657223371C64AC8123FEFB35FDA6" pageId="3" pageNumber="72" type="materials_examined">
<paragraph id="8B9536F923371C64AC8123FEFB35FDA6" blockId="3.[151,1436,151,824]" pageId="3" pageNumber="72">
<emphasis id="B95EEAEB23371C64AC8123FEFDF2FF2B" bold="true" box="[198,593,151,176]" pageId="3" pageNumber="72">Additional specimens examined:</emphasis>
<materialsCitation id="3B423CA423371C64AE2A23FEFC96FF4E" collectingDate="2005-05-17" collectionCode="PUH" collectorName="S. Yap" country="Philippines" elevation="120" location="Sierra Madre Mountains" municipality="Palanan" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Isabela Province">
<materialsCitation id="3B423CA423371C64AE2A23FEFC96FF4E" pageId="3" pageNumber="72">
<collectingCountry id="F33D766923371C64AE2A23FEFCB6FF2B" box="[621,789,151,176]" name="Philippines" pageId="3" pageNumber="72">PHILIPPINES</collectingCountry>
. Luzon Island,
.
<location id="8EF5602223371C64AF6323F1FC19FF2B" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AF6323F1FC19FF2B" box="[804,954,151,176]" country="Philippines" name="Luzon Island" pageId="3" pageNumber="72">Luzon Island</location>
,
<collectingRegion id="49EEF81B23371C64AF8C23FEFB2EFF2B" box="[971,1165,151,176]" country="Philippines" name="Isabela" pageId="3" pageNumber="72">Isabela Province</collectingRegion>
:
<collectingMunicipality id="6BF1AC8323371C64A8DC23FEFB54FF2B" box="[1179,1271,151,176]" pageId="3" pageNumber="72">Palanan</collectingMunicipality>
,
<location id="8EF5602223371C64A94223FEFEB4FF4E" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64A94223FEFEB4FF4E" country="Philippines" municipality="Palanan" name="Sierra Madre Mountains" pageId="3" pageNumber="72" stateProvince="Isabela Province">Sierra Madre Mountains</location>
<location id="8EF5602223371C64A94223FEFEB4FF4E" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64A94223FEFEB4FF4E" country="Philippines" municipality="Palanan" name="Sierra Madre Mountains" pageId="3" pageNumber="72" stateProvince="Isabela">Sierra Madre Mountains</location>
,
<date id="FF94103923371C64AD6423D5FE60FF4E" box="[291,451,188,213]" pageId="3" pageNumber="72" value="2005-05-17">
<collectingDate id="EFD0E9D123371C64AD6423D5FE60FF4E" box="[291,451,188,213]" pageId="3" pageNumber="72" value="2005-05-17">17 May 2005</collectingDate>
@ -315,17 +297,15 @@ Photo by S.L. Yap.
<collectorName id="26DF532F23371C64AE1223D5FD3DFF4F" box="[597,670,188,212]" pageId="3" pageNumber="72">S. Yap</collectorName>
001
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AEA423D5FC8BFF4E" box="[739,808,188,213]" pageId="3" pageNumber="72">PUH</collectionCode>
!)
(PUH!)
</materialsCitation>
.
<materialsCitation id="3B423CA423371C64AF0623D5FD81FF63" collectingDate="1983-08-17" collectionCode="K" collectorName="Fernando" elevation="300" location="Lucapon" municipality="Sta. Cruz" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Zambales Province">
<materialsCitation id="3B423CA423371C64AF0623D5FD81FF63" pageId="3" pageNumber="72">
<collectingRegion id="49EEF81B23371C64AF0623D5FB84FF4E" box="[833,1063,188,213]" country="Philippines" name="Zambales" pageId="3" pageNumber="72">Zambales Province</collectingRegion>
:
<collectingMunicipality id="6BF1AC8323371C64A87423D5FB00FF4E" box="[1075,1187,188,213]" pageId="3" pageNumber="72">Sta. Cruz</collectingMunicipality>
,
<location id="8EF5602223371C64A8F723D5FABBFF48" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64A8F723D5FABBFF48" box="[1200,1304,188,211]" country="Philippines" municipality="Sta. Cruz" name="Lucapon" pageId="3" pageNumber="72" stateProvince="Zambales Province">Lucapon</location>
<location id="8EF5602223371C64A8F723D5FABBFF48" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64A8F723D5FABBFF48" box="[1200,1304,188,211]" country="Philippines" municipality="Sta. Cruz" name="Lucapon" pageId="3" pageNumber="72" stateProvince="Zambales">Lucapon</location>
,
<quantity id="4CD29B1C23371C64A96223D5FAD1FF48" box="[1317,1394,188,213]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="72" unit="m" value="300.0">
<elevation id="0007D1CA23371C64A96223D5FAD1FF48" box="[1317,1394,188,213]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="72" unit="m" value="300.0">300 m</elevation>
@ -339,15 +319,13 @@ Photo by S.L. Yap.
<collectorName id="26DF532F23371C64AD7923B6FE12FF63" box="[318,433,223,248]" pageId="3" pageNumber="72">Fernando</collectorName>
305
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64ADBF23B6FDB7FF63" box="[504,532,223,248]" pageId="3" pageNumber="72">K</collectionCode>
!)
(K!)
</materialsCitation>
.
<materialsCitation id="3B423CA423371C64AE7723B6FA94FF63" box="[560,1335,223,248]" collectingDate="1915-04" collectionCode="BM" collectorName="A. Loher" location="Batingtingan" municipality="Batingtingan" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Rizal Province">
<materialsCitation id="3B423CA423371C64AE7723B6FA94FF63" box="[560,1335,223,248]" pageId="3" pageNumber="72">
<collectingRegion id="49EEF81B23371C64AE7723B6FD47FF63" box="[560,740,223,248]" country="Philippines" name="Rizal" pageId="3" pageNumber="72">Rizal Province</collectingRegion>
:
<collectingMunicipality id="6BF1AC8323371C64AEB723B6FC28FF63" box="[752,907,223,248]" country="Philippines" name="Batingtingan" pageId="3" pageNumber="72" stateProvince="Rizal">Batingtingan</collectingMunicipality>
<location id="8EF5602223371C64AEB723B6FC28FF63" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AEB723B6FC28FF63" box="[752,907,223,248]" country="Philippines" name="Batingtingan" pageId="3" pageNumber="72" stateProvince="Rizal">Batingtingan</location>
,
<date id="FF94103923371C64AFDF23B6FBBFFF63" box="[920,1052,223,248]" pageId="3" pageNumber="72" value="1915-04">
<collectingDate id="EFD0E9D123371C64AFDF23B6FBBFFF63" box="[920,1052,223,248]" pageId="3" pageNumber="72" value="1915-04">April 1915</collectingDate>
@ -357,17 +335,15 @@ Photo by S.L. Yap.
<collectorName id="26DF532F23371C64A86E2389FB33FF63" box="[1065,1168,223,248]" pageId="3" pageNumber="72">A. Loher</collectorName>
12973
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64A8B323B6FA88FF63" box="[1268,1323,223,248]" pageId="3" pageNumber="72">BM</collectionCode>
!)
(BM!)
</materialsCitation>
.
<materialsCitation id="3B423CA423371C64A9022389FB20FE86" collectionCode="BH, CAHUP" collectorName="Moore &amp; Pancho" elevation="200" location="Mt Makiling" municipality="Los Banos" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Laguna Province">
<materialsCitation id="3B423CA423371C64A9022389FB20FE86" pageId="3" pageNumber="72">
<collectingRegion id="49EEF81B23371C64A9022389FEA6FE86" country="Philippines" name="Laguna" pageId="3" pageNumber="72">Laguna Province</collectingRegion>
:
<collectingMunicipality id="6BF1AC8323371C64AD56226DFE31FE80" box="[273,402,260,283]" pageId="3" pageNumber="72">Los Baños</collectingMunicipality>
,
<location id="8EF5602223371C64ADE7226DFD98FE86" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64ADE7226DFD98FE86" box="[416,571,260,285]" country="Philippines" municipality="Los Banos" name="Mt Makiling" pageId="3" pageNumber="72" stateProvince="Laguna Province">Mt Makiling</location>
<location id="8EF5602223371C64ADE7226DFD98FE86" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64ADE7226DFD98FE86" box="[416,571,260,285]" country="Philippines" municipality="Los Banos" name="Mt Makiling" pageId="3" pageNumber="72" stateProvince="Laguna">Mt Makiling</location>
,
<quantity id="4CD29B1C23371C64AE00226DFD36FE80" box="[583,661,260,285]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="72" unit="m" value="200.0">
<elevation id="0007D1CA23371C64AE00226DFD36FE80" box="[583,661,260,285]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="72" unit="m" value="200.0">200 m</elevation>
@ -379,14 +355,10 @@ Photo by S.L. Yap.
<collectorName id="26DF532F23371C64AF5F226DFCD2FE86" box="[792,881,260,285]" pageId="3" pageNumber="72">Pancho</collectorName>
9393
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AF80226DFC5BFE86" box="[967,1016,260,285]" pageId="3" pageNumber="72">BH</collectionCode>
!,
<collectionCode id="ED3BAE3C23371C64A84D226DFBD4FE86" box="[1034,1143,260,285]" pageId="3" pageNumber="72">CAHUP</collectionCode>
!)
(BH!, CAHUP!)
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64A8D6226DFD20FEDB" collectingDate="1979-04-01" collectionCode="K, LBC, PNH" collectorName="Dransfield" elevation="400" pageId="3" pageNumber="72" specimenCount="1">
<materialsCitation id="3B423CA423371C64A8D6226DFD20FEDB" pageId="3" pageNumber="72">
<quantity id="4CD29B1C23371C64A8D6226DFB7CFE80" box="[1169,1247,260,285]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="72" unit="m" value="400.0">
<elevation id="0007D1CA23371C64A8D6226DFB7CFE80" box="[1169,1247,260,285]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="72" unit="m" value="400.0">400 m</elevation>
</quantity>
@ -399,16 +371,10 @@ Photo by S.L. Yap.
<collectorName id="26DF532F23371C64ACD0224EFEB6FEDB" box="[151,277,295,320]" pageId="3" pageNumber="72">Dransfield</collectorName>
et al. 5467
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64ADF6224EFE6EFEDB" box="[433,461,295,320]" pageId="3" pageNumber="72">K</collectionCode>
!,
<collectionCode id="ED3BAE3C23371C64AD98224EFD83FEDB" box="[479,544,295,320]" pageId="3" pageNumber="72">LBC</collectionCode>
!,
<collectionCode id="ED3BAE3C23371C64AE75224EFDD5FEDB" box="[562,630,295,320]" pageId="3" pageNumber="72">PNH</collectionCode>
!)
(K!, LBC!, PNH!)
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64AED7224EFB6BFEDB" box="[656,1224,295,320]" collectingDate="2003-03-13" collectionCode="LBC" collectorName="Fernando" elevation="450" pageId="3" pageNumber="72" specimenCount="1">
<materialsCitation id="3B423CA423371C64AED7224EFB6BFEDB" box="[656,1224,295,320]" pageId="3" pageNumber="72">
<quantity id="4CD29B1C23371C64AED7224EFD7DFEA4" box="[656,734,295,320]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="72" unit="m" value="450.0">
<elevation id="0007D1CA23371C64AED7224EFD7DFEA4" box="[656,734,295,320]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="72" unit="m" value="450.0">450 m</elevation>
</quantity>
@ -421,12 +387,10 @@ et al. 5467
<collectorName id="26DF532F23371C64AFF6224EFB86FEDB" box="[945,1061,295,320]" pageId="3" pageNumber="72">Fernando</collectorName>
1626
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64A83E224EFB18FEDB" box="[1145,1211,295,320]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
(LBC!)
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64A890224EFE16FEFE" collectingDate="2004-08-18" collectionCode="LBC" collectorName="Fernando" pageId="3" pageNumber="72" specimenCount="1">
<materialsCitation id="3B423CA423371C64A890224EFE16FEFE" pageId="3" pageNumber="72">
<date id="FF94103923371C64A890224EFA3BFEDB" box="[1239,1432,295,320]" pageId="3" pageNumber="72" value="2004-08-18">
<collectingDate id="EFD0E9D123371C64A890224EFA3BFEDB" box="[1239,1432,295,320]" pageId="3" pageNumber="72" value="2004-08-18">18 August 2004</collectingDate>
</date>
@ -435,17 +399,16 @@ et al. 5467
<collectorName id="26DF532F23371C64ACD02225FEAFFEFE" box="[151,268,332,357]" pageId="3" pageNumber="72">Fernando</collectorName>
1732
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AD232225FE0AFEFE" box="[356,425,332,357]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
(LBC!)
</materialsCitation>
.
<materialsCitation id="3B423CA423371C64AD842225FE71FE13" collectingDate="1979-09-01" collectionCode="K, LBC" collectorName="Fernando" elevation="300" location="Quezon National Park" municipality="Atimonan" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Quezon Province">
<materialsCitation id="3B423CA423371C64AD842225FE71FE13" pageId="3" pageNumber="72">
<collectingRegion id="49EEF81B23371C64AD842225FD3AFEFE" box="[451,665,332,357]" country="Philippines" name="Quezon" pageId="3" pageNumber="72">Quezon Province</collectingRegion>
:
<collectingMunicipality id="6BF1AC8323371C64AEE22225FCBCFEFE" box="[677,799,332,357]" pageId="3" pageNumber="72">Atimonan</collectingMunicipality>
<collectorName id="26DF532F23371C64AEE22225FCBCFEFE" box="[677,799,332,357]" pageId="3" pageNumber="72">Atimonan</collectorName>
,
<location id="8EF5602223371C64AF692225FB9CFEFE" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AF692225FB9CFEFE" box="[814,1087,332,357]" municipality="Atimonan" name="Quezon National Park" pageId="3" pageNumber="72" stateProvince="Quezon Province">Quezon National Park</location>
<collectingRegion id="49EEF81B23371C64AF692225FC29FEFE" box="[814,906,332,357]" country="Philippines" name="Quezon" pageId="3" pageNumber="72">Quezon</collectingRegion>
<collectingMunicipality id="6BF1AC8323371C64AFD32225FB9CFEFE" box="[916,1087,332,357]" pageId="3" pageNumber="72">National Park</collectingMunicipality>
,
<quantity id="4CD29B1C23371C64A80B2225FB38FEF8" box="[1100,1179,332,357]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="72" unit="m" value="300.0">
<elevation id="0007D1CA23371C64A80B2225FB38FEF8" box="[1100,1179,332,357]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="72" unit="m" value="300.0">300 m</elevation>
@ -455,18 +418,11 @@ et al. 5467
<collectingDate id="EFD0E9D123371C64A8EE2225FA34FEFE" box="[1193,1431,332,357]" pageId="3" pageNumber="72" value="1979-09-01">01 September 1979</collectingDate>
</date>
,
<emphasis id="B95EEAEB23371C64ACD02206FEE3FE13" box="[151,320,367,392]" italics="true" pageId="3" pageNumber="72">
<collectorName id="26DF532F23371C64ACD02206FEA8FE13" box="[151,267,367,392]" pageId="3" pageNumber="72">Fernando</collectorName>
166
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AD132206FED3FE13" box="[340,368,367,392]" pageId="3" pageNumber="72">K</collectionCode>
!,
<collectionCode id="ED3BAE3C23371C64ADC42206FE66FE13" box="[387,453,367,392]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
<emphasis id="B95EEAEB23371C64ACD02206FEE3FE13" box="[151,320,367,392]" italics="true" pageId="3" pageNumber="72">Fernando 166</emphasis>
(K!, LBC!)
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64ADA72206FBC2FE13" box="[480,1121,367,392]" collectingDate="1979-04-04" collectionCode="LBC" collectorName="Dransfield" elevation="200" pageId="3" pageNumber="72" specimenCount="1">
<materialsCitation id="3B423CA423371C64ADA72206FBC2FE13" box="[480,1121,367,392]" pageId="3" pageNumber="72">
<quantity id="4CD29B1C23371C64ADA72206FD8DFE1C" box="[480,558,367,392]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="72" unit="m" value="200.0">
<elevation id="0007D1CA23371C64ADA72206FD8DFE1C" box="[480,558,367,392]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="72" unit="m" value="200.0">200 m</elevation>
</quantity>
@ -475,16 +431,11 @@ et al. 5467
<collectingDate id="EFD0E9D123371C64AE7B2206FD4BFE13" box="[572,744,367,392]" pageId="3" pageNumber="72" value="1979-04-04">04 April 1979</collectingDate>
</date>
,
<emphasis id="B95EEAEB23371C64AEB22206FC5CFE13" box="[757,1023,367,392]" italics="true" pageId="3" pageNumber="72">
<collectorName id="26DF532F23371C64AEB22206FCD7FE13" box="[757,884,367,392]" pageId="3" pageNumber="72">Dransfield</collectorName>
et al. 5472
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64A8562206FBF7FE13" box="[1041,1108,367,392]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
<emphasis id="B95EEAEB23371C64AEB22206FC5CFE13" box="[757,1023,367,392]" italics="true" pageId="3" pageNumber="72">Dransfield et al. 5472</emphasis>
(LBC!)
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64A8282206FE01FE36" collectingDate="2006-10-13" collectionCode="LBC" collectorName="Fernando" elevation="220" pageId="3" pageNumber="72" specimenCount="1">
<materialsCitation id="3B423CA423371C64A8282206FE01FE36" pageId="3" pageNumber="72">
<quantity id="4CD29B1C23371C64A8282206FB1EFE1C" box="[1135,1213,367,392]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="72" unit="m" value="220.0">
<elevation id="0007D1CA23371C64A8282206FB1EFE1C" box="[1135,1213,367,392]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="72" unit="m" value="220.0">220 m</elevation>
</quantity>
@ -493,16 +444,11 @@ et al. 5472
<collectingDate id="EFD0E9D123371C64A88C2206FA3BFE13" box="[1227,1432,367,392]" pageId="3" pageNumber="72" value="2006-10-13">13 October 2006</collectingDate>
</date>
,
<emphasis id="B95EEAEB23371C64ACD022FDFEE6FE36" box="[151,325,404,429]" italics="true" pageId="3" pageNumber="72">
<collectorName id="26DF532F23371C64ACD022FDFEA6FE36" box="[151,261,404,429]" pageId="3" pageNumber="72">Fernando</collectorName>
1824
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AD1122FDFE36FE36" box="[342,405,404,429]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
<emphasis id="B95EEAEB23371C64ACD022FDFEE6FE36" box="[151,325,404,429]" italics="true" pageId="3" pageNumber="72">Fernando 1824</emphasis>
(LBC!)
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64ADE922FDFC9FFE36" box="[430,828,404,429]" collectionCode="K, LBC" collectorName="Baja-Lapis" elevation="300" pageId="3" pageNumber="72" specimenCount="1">
<materialsCitation id="3B423CA423371C64ADE922FDFC9FFE36" box="[430,828,404,429]" pageId="3" pageNumber="72">
<quantity id="4CD29B1C23371C64ADE922FDFE5BFE30" box="[430,504,404,429]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="72" unit="m" value="300.0">
<elevation id="0007D1CA23371C64ADE922FDFE5BFE30" box="[430,504,404,429]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="72" unit="m" value="300.0">300 m</elevation>
</quantity>
@ -511,15 +457,13 @@ et al. 5472
<collectorName id="26DF532F23371C64AE4322FCFD23FE37" box="[516,640,405,428]" pageId="3" pageNumber="72">Baja-Lapis</collectorName>
010
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AE8322FDFD7CFE36" box="[708,735,404,429]" pageId="3" pageNumber="72">K</collectionCode>
!,
<collectionCode id="ED3BAE3C23371C64AEB722FDFC93FE36" box="[752,816,404,429]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
(K!, LBC!)
</materialsCitation>
;
<materialsCitation id="3B423CA423371C64AF0F22FDFEB9FE4B" collectingDate="1987-04-15" collectionCode="LBC" collectorName="Natividad" location="Real, Kawayan Watershed" pageId="3" pageNumber="72" specimenCount="1">
<location id="8EF5602223371C64AF0F22FDFBD6FE36" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AF0F22FDFBD6FE36" box="[840,1141,404,429]" name="Real, Kawayan Watershed" pageId="3" pageNumber="72">Real, Kawayan Watershed</location>
<materialsCitation id="3B423CA423371C64AF0F22FDFEB9FE4B" pageId="3" pageNumber="72">
<collectorName id="26DF532F23371C64AF0F22FDFCDDFE36" box="[840,894,404,429]" pageId="3" pageNumber="72">Real</collectorName>
,
<location id="8EF5602223371C64AFCD22FDFBD6FE36" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AFCD22FDFBD6FE36" box="[906,1141,404,429]" country="Philippines" municipality="National Park" name="Kawayan Watershed" pageId="3" pageNumber="72" stateProvince="Quezon">Kawayan Watershed</location>
,
<date id="FF94103923371C64A8C722FDFA81FE36" box="[1152,1314,404,429]" pageId="3" pageNumber="72" value="1987-04-15">
<collectingDate id="EFD0E9D123371C64A8C722FDFA81FE36" box="[1152,1314,404,429]" pageId="3" pageNumber="72" value="1987-04-15">15 April 1987</collectingDate>
@ -529,17 +473,15 @@ et al. 5472
<collectorName id="26DF532F23371C64A96A22FDFA38FE36" box="[1325,1435,404,429]" pageId="3" pageNumber="72">Natividad</collectorName>
s.n
</emphasis>
. (
<collectionCode id="ED3BAE3C23371C64AC8922DEFEADFE4B" box="[206,270,439,464]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
. (LBC!)
</materialsCitation>
.
<materialsCitation id="3B423CA423371C64AD6222DEFF47FE6E" collectingDate="1985-07-14" collectionCode="K, LBC" collectorName="Fernando" elevation="150" location="Bicol National Park" municipality="Basud" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Camarines Norte Province">
<materialsCitation id="3B423CA423371C64AD6222DEFF47FE6E" pageId="3" pageNumber="72">
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,
<collectingMunicipality id="6BF1AC8323371C64AE1B22DEFD06FE4B" box="[604,677,439,464]" pageId="3" pageNumber="72">Basud</collectingMunicipality>
,
<location id="8EF5602223371C64AEF722DEFC32FE4B" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AEF722DEFC32FE4B" box="[688,913,439,464]" country="Philippines" municipality="Basud" name="Bicol National Park" pageId="3" pageNumber="72" stateProvince="Camarines Norte Province">Bicol National Park</location>
<location id="8EF5602223371C64AEF722DEFC32FE4B" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AEF722DEFC32FE4B" box="[688,913,439,464]" country="Philippines" municipality="Basud" name="Bicol National Park" pageId="3" pageNumber="72" stateProvince="Camarines Norte">Bicol National Park</location>
,
<quantity id="4CD29B1C23371C64AFDA22DEFBBDFE54" box="[925,1054,439,464]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="72" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<elevation id="0007D1CA23371C64AFDA22DEFBBDFE54" box="[925,1054,439,464]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="72" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100200 m</elevation>
@ -553,27 +495,21 @@ s.n
<collectorName id="26DF532F23371C64A88F22DEFA95FE4B" box="[1224,1334,439,464]" pageId="3" pageNumber="72">Fernando</collectorName>
545
</emphasis>
(
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!,
<collectionCode id="ED3BAE3C23371C64ACD022B5FF74FE6E" box="[151,215,476,501]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
(K!, LBC!)
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64ACB522B4FD95FE6E" box="[242,566,476,501]" collectionCode="K, PNH" collectorName="Baja-Lapis" pageId="3" pageNumber="72" specimenCount="1">
<materialsCitation id="3B423CA423371C64ACB522B4FD95FE6E" box="[242,566,476,501]" pageId="3" pageNumber="72">
<emphasis id="B95EEAEB23371C64ACB522B4FE06FE6E" box="[242,421,476,501]" italics="true" pageId="3" pageNumber="72">
<collectorName id="26DF532F23371C64ACB522B4FED1FE6F" box="[242,370,477,500]" pageId="3" pageNumber="72">Baja-Lapis</collectorName>
036
</emphasis>
(
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!,
<collectionCode id="ED3BAE3C23371C64ADA222B5FD8AFE6E" box="[485,553,476,501]" pageId="3" pageNumber="72">PNH</collectionCode>
!)
(K!, PNH!)
</materialsCitation>
.
<materialsCitation id="3B423CA423371C64AE0422B5FE00FD83" collectingDate="2011-02-03" collectionCode="LBC" collectorName="Fernando" elevation="300" location="Silago" municipality="Silago" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Southern Leyte Province">
Leyte Island,
<materialsCitation id="3B423CA423371C64AE0422B5FE00FD83" pageId="3" pageNumber="72">
<collectingRegion id="49EEF81B23371C64AE0422B5FD26FE6F" box="[579,645,476,500]" country="Philippines" name="Southern Leyte" pageId="3" pageNumber="72">Leyte</collectingRegion>
<collectorName id="26DF532F23371C64AECA22B5FD74FE6E" box="[653,727,476,501]" pageId="3" pageNumber="72">Island</collectorName>
,
<collectingRegion id="49EEF81B23371C64AEA322B5FBAAFE6E" box="[740,1033,476,501]" country="Philippines" name="Southern Leyte" pageId="3" pageNumber="72">Southern Leyte Province</collectingRegion>
:
<collectingMunicipality id="6BF1AC8323371C64A85322B5FBC2FE6E" box="[1044,1121,476,501]" pageId="3" pageNumber="72">Silago</collectingMunicipality>
@ -590,16 +526,15 @@ Leyte Island,
<collectorName id="26DF532F23371C64ACD02296FEA5FD83" box="[151,262,511,536]" pageId="3" pageNumber="72">Fernando</collectorName>
2351
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AD112296FE35FD83" box="[342,406,511,536]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
(LBC!)
</materialsCitation>
.
<materialsCitation id="3B423CA423371C64ADE92296FEEAFDA6" collectingDate="1984-07-18" collectionCode="K, LBC" collectorName="Fernando" elevation="150" location="Trento" municipality="Trento" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Agusan del Sur Province">
Mindanao Island,
<materialsCitation id="3B423CA423371C64ADE92296FEEAFDA6" pageId="3" pageNumber="72">
<location id="8EF5602223371C64ADE92296FDCCFD83" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64ADE92296FDCCFD83" box="[430,623,511,536]" country="Philippines" municipality="Silago" name="Mindanao Island" pageId="3" pageNumber="72" stateProvince="Southern Leyte">Mindanao Island</location>
,
<collectingRegion id="49EEF81B23371C64AE3C2296FC39FD83" box="[635,922,511,536]" country="Philippines" name="Agusan del Sur" pageId="3" pageNumber="72">Agusan del Sur Province</collectingRegion>
:
<collectingMunicipality id="6BF1AC8323371C64AFE22169FC51FD8C" box="[933,1010,512,535]" pageId="3" pageNumber="72">Trento</collectingMunicipality>
<location id="8EF5602223371C64AFE22169FC51FD8C" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AFE22169FC51FD8C" box="[933,1010,512,535]" country="Philippines" name="Trento" pageId="3" pageNumber="72" stateProvince="Agusan del Sur">Trento</location>
,
<quantity id="4CD29B1C23371C64AFBA2296FBDCFD8C" box="[1021,1151,511,536]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="72" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<elevation id="0007D1CA23371C64AFBA2296FBDCFD8C" box="[1021,1151,511,536]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="72" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100200 m</elevation>
@ -613,17 +548,13 @@ Mindanao Island,
<collectorName id="26DF532F23371C64A96A2296FA3FFD83" box="[1325,1436,511,536]" pageId="3" pageNumber="72">Fernando</collectorName>
418
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64AC95214DFF4EFDA6" box="[210,237,548,573]" pageId="3" pageNumber="72">K</collectionCode>
!,
<collectionCode id="ED3BAE3C23371C64ACBB214DFE9FFDA6" box="[252,316,548,573]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
(K!, LBC!)
</materialsCitation>
;
<materialsCitation id="3B423CA423371C64AD13214DFB31FDA6" box="[340,1170,548,573]" collectingDate="1987-10-03" collectionCode="LBC" collectorName="Fernando" location="Bislig" municipality="Bislig" pageId="3" pageNumber="72" specimenCount="1" stateProvince="Surigao del Sur Province">
<materialsCitation id="3B423CA423371C64AD13214DFB31FDA6" box="[340,1170,548,573]" pageId="3" pageNumber="72">
<collectingRegion id="49EEF81B23371C64AD13214DFDCDFDA6" box="[340,622,548,573]" country="Philippines" name="Surigao del Sur" pageId="3" pageNumber="72">Surigao del Sur Province</collectingRegion>
,
<collectingMunicipality id="6BF1AC8323371C64AE3C214DFD62FDA6" box="[635,705,548,573]" country="Philippines" name="Bislig" pageId="3" pageNumber="72" stateProvince="Surigao del Sur">Bislig</collectingMunicipality>
<location id="8EF5602223371C64AE3C214DFD62FDA6" LSID="urn:lsid:plazi:treatment:038387EF23351C64ACD023B7FB24FCA3:8EF5602223371C64AE3C214DFD62FDA6" box="[635,705,548,573]" country="Philippines" name="Bislig" pageId="3" pageNumber="72" stateProvince="Surigao del Sur">Bislig</location>
,
<date id="FF94103923371C64AE8B214DFC28FDA6" box="[716,907,548,573]" pageId="3" pageNumber="72" value="1987-10-03">
<collectingDate id="EFD0E9D123371C64AE8B214DFC28FDA6" box="[716,907,548,573]" pageId="3" pageNumber="72" value="1987-10-03">03 October 1987</collectingDate>
@ -633,14 +564,10 @@ Mindanao Island,
<collectorName id="26DF532F23371C64AFD1214DFBA7FDA6" box="[918,1028,548,573]" pageId="3" pageNumber="72">Fernando</collectorName>
728
</emphasis>
(
<collectionCode id="ED3BAE3C23371C64A801214DFB25FDA6" box="[1094,1158,548,573]" pageId="3" pageNumber="72">LBC</collectionCode>
!)
(LBC!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330657223371C64AC81212EFB24FCA3" pageId="3" pageNumber="72" type="discussion">
<paragraph id="8B9536F923371C64AC81212EFB24FCA3" blockId="3.[151,1436,151,824]" pageId="3" pageNumber="72">
<emphasis id="B95EEAEB23371C64AC81212EFEB1FDFB" bold="true" box="[198,274,583,608]" pageId="3" pageNumber="72">Notes:</emphasis>
—In its vegetative characters, especially in the leaf sheath and leaf armature, this rattan is quite unlike any other known species of
@ -674,6 +601,5 @@ in its generally larger, to
<quantity id="4CD29B1C23371C64AFA52195FB95FC88" box="[994,1078,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="72" unit="cm" value="6.5" valueMax="7.0" valueMin="6.0">67 cm</quantity>
diameter sheathed stems, fruits with up to 2122 vertical series of scales, and seeds which are smooth on the convex side.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,46 +1,45 @@
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<treatment id="038387EF23371C62ACD0200EFE02FDFB" ID-DOI="http://doi.org/10.5281/zenodo.15212848" ID-Zenodo-Dep="15212848" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB" httpUri="http://treatment.plazi.org/id/038387EF23371C62ACD0200EFE02FDFB" lastPageId="5" lastPageNumber="74" pageId="3" pageNumber="72" scope_family="Arecaceae">
<paragraph id="8B9536F923371C64ACD0200EFD9BFC1B" blockId="3.[151,1436,871,933]" box="[151,568,871,896]" pageId="3" pageNumber="72">
<taxonomicName id="4C2A4D7A23371C64ACD0200EFE76FC1B" authority="Fernando" authorityName="Fernando. A, Leaf" authorityYear="2014" box="[151,469,871,896]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="72" phylum="Chordata" rank="species" species="parutan" status="sp. nov.">
<emphasis id="B95EEAEB23371C64ACD0200EFEC3FCE4" bold="true" box="[151,352,871,896]" italics="true" pageId="3" pageNumber="72">Calamus parutan</emphasis>
@ -51,19 +50,17 @@ Fernando
<taxonomicNameLabel id="A26D579023371C64ADA62000FD9BFC1B" box="[481,568,873,896]" pageId="3" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C330657223371C64AE78200EFBBCFC3E" pageId="3" pageNumber="72" type="materials_examined">
<paragraph id="8B9536F923371C64AE78200EFBBCFC3E" blockId="3.[151,1436,871,933]" pageId="3" pageNumber="72">
<collectingCounty id="62F44E7523371C64AE78200EFDDAFC1B" box="[575,633,871,896]" pageId="3" pageNumber="72">Type</collectingCounty>
:—
<materialsCitation id="3B423CA423371C64AEDD200EFCDEFC3E" collectingDate="1973-05-21" collectionCode="K" collectorName="Dransfield" country="Indonesia" elevation="50" location="Betiri Nature Reserve" municipality="Sukamade" pageId="3" pageNumber="72" specimenCount="1" stateProvince="East Java" typeStatus="holotype">
<materialsCitation id="3B423CA423371C64AEDD200EFCDEFC3E" pageId="3" pageNumber="72">
<collectingCountry id="F33D766923371C64AEDD200EFC9BFC1B" box="[666,824,871,896]" name="Indonesia" pageId="3" pageNumber="72">INDONESIA</collectingCountry>
.
<collectingRegion id="49EEF81B23371C64AF042001FC13FCE4" box="[835,944,872,895]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="72">East Java</collectingRegion>
,
<collectingMunicipality id="6BF1AC8323371C64AFFC200EFB91FC1B" box="[955,1074,871,896]" pageId="3" pageNumber="72">Sukamade</collectingMunicipality>
,
<location id="8EF5602223371C64A879200EFA95FCE4" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223371C64A879200EFA95FCE4" box="[1086,1334,871,896]" country="Indonesia" county="Type" municipality="Sukamade" name="Betiri Nature Reserve" pageId="3" pageNumber="72" stateProvince="East Java">Betiri Nature Reserve</location>
<location id="8EF5602223371C64A879200EFA95FCE4" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223371C64A879200EFA95FCE4" box="[1086,1334,871,896]" country="Indonesia" county="Type" municipality="Sukamade" name="Betiri Nature Reserve" pageId="3" pageNumber="72" stateProvince="Jawa Timur">Betiri Nature Reserve</location>
, lowland forest, hillslope,
<quantity id="4CD29B1C23371C64AD0920E5FE2AFC38" box="[334,393,908,933]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="72" unit="m" value="50.0">
<elevation id="0007D1CA23371C64AD0920E5FE2AFC38" box="[334,393,908,933]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="72" unit="m" value="50.0">50 m</elevation>
@ -73,25 +70,15 @@ Fernando
<collectingDate id="EFD0E9D123371C64ADD320E5FD89FC3E" box="[404,554,908,933]" pageId="3" pageNumber="72" value="1973-05-21">21 May 1973</collectingDate>
</date>
,
<emphasis id="B95EEAEB23371C64AE7320E5FD44FC3E" box="[564,743,908,933]" italics="true" pageId="3" pageNumber="72">
<collectorName id="26DF532F23371C64AE7320E5FD09FC3E" box="[564,682,908,933]" pageId="3" pageNumber="72">Dransfield</collectorName>
3534
</emphasis>
<emphasis id="B95EEAEB23371C64AE7320E5FD44FC3E" box="[564,743,908,933]" italics="true" pageId="3" pageNumber="72">Dransfield 3534</emphasis>
(
<typeStatus id="5491885B23371C64AEB220E5FCF4FC3E" box="[757,855,908,933]" pageId="3" pageNumber="72" type="holotype">holotype</typeStatus>
<collectionCode id="ED3BAE3C23371C64AF1920E5FCDAFC3E" box="[862,889,908,933]" pageId="3" pageNumber="72">K</collectionCode>
!
K!
</materialsCitation>
,
<materialsCitation id="3B423CA423371C64AFCE20E5FBB2FC3E" box="[905,1041,908,933]" collectingDate="1973-05-21" collectionCode="BO" collectorName="Dransfield" country="Indonesia" elevation="50" location="Betiri Nature Reserve" municipality="Sukamade" pageId="3" pageNumber="72" specimenCount="1" stateProvince="East Java" typeStatus="isotype">
<typeStatus id="5491885B23371C64AFCE20E5FC7AFC3E" box="[905,985,908,933]" pageId="3" pageNumber="72">isotype</typeStatus>
<collectionCode id="ED3BAE3C23371C64AF9820E5FBADFC3E" box="[991,1038,908,933]" pageId="3" pageNumber="72">BO</collectionCode>
!
</materialsCitation>
<materialsCitation id="3B423CA423371C64AFCE20E5FBB2FC3E" box="[905,1041,908,933]" pageId="3" pageNumber="72">isotype BO!</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330657223371C64ACD020BDFC62FB92" pageId="3" pageNumber="72" type="diagnosis">
<paragraph id="8B9536F923371C64ACD020BDFC62FB92" blockId="3.[151,1436,980,1033]" pageId="3" pageNumber="72">
Related to
<taxonomicName id="4C2A4D7A23371C64AD4220BDFD05FC71" authority="Beccari (1902: 198)" authorityName="Beccari" authorityPageNumber="198" authorityYear="1902" box="[261,678,980,1002]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="72" phylum="Chordata" rank="species" species="burckianus">
@ -106,8 +93,6 @@ group of sect.
<emphasis id="B95EEAEB23371C64A80F20BDFB7DFC72" box="[1096,1246,980,1001]" italics="true" pageId="3" pageNumber="72">Podocephalus,</emphasis>
but differing in its fruit with only one seed, and in the shorter (up to ½ times) fruiting rachilla.
</paragraph>
</subSubSection>
<subSubSection id="C330657223371C64ACD0275CFD8EF8FD" pageId="3" pageNumber="72" type="description">
<paragraph id="8B9536F923371C64ACD0275CFD8EF8FD" blockId="3.[151,1436,1077,2038]" pageId="3" pageNumber="72">
Clustering, robust rattan climbing to
<quantity id="4CD29B1C23371C64AE71275CFDCCFBD6" box="[566,623,1077,1102]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="72" unit="m" value="30.0">30 m</quantity>
@ -191,8 +176,6 @@ apart along the axis. Pistillate flower not known. Fruit globose, 0.9 ×
<figureCitation id="13112A7C23371C64ADFA2424FD83F8FD" box="[445,544,1869,1894]" captionStart="FIGURE 2" captionStartId="4.[151,244,1909,1931]" captionTargetBox="[201,1391,218,1877]" captionTargetId="figure-107@4.[185,1409,195,1892]" captionTargetPageId="4" captionText="FIGURE 2. Calamus parutan Fernando. A, Leaf sheath showing dense laminar spines and distinct ocrea. B, Adaxial side of petiole. C, Adaxial side of rachis with proximal leaflets. D, Abaxial side of mid-portion of rachis. E, Portion of male inflorescence showing unarmed tubular bracts. F, Partial infructescence with fruiting rachillae. Scale bar = 1 cm. AD and F, Dransfield 3534 (BO); E, Maier Sarip 120 (BO). All photos by E.S. Fernando." figureDoi="http://doi.org/10.5281/zenodo.5138220" httpUri="https://zenodo.org/record/5138220/files/figure.png" pageId="3" pageNumber="72">Figure 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330657223371C64AC812418FADDF811" box="[198,1406,1905,1930]" pageId="3" pageNumber="72" type="distribution">
<paragraph id="8B9536F923371C64AC812418FADDF811" blockId="3.[151,1436,1077,2038]" box="[198,1406,1905,1930]" pageId="3" pageNumber="72">
<emphasis id="B95EEAEB23371C64AC812418FE51F811" bold="true" box="[198,498,1905,1930]" pageId="3" pageNumber="72">Distribution and habitat:</emphasis>
—Endemic to
@ -203,8 +186,6 @@ and
<collectingRegion id="49EEF81B23371C64AFF52418FC46F811" box="[946,997,1905,1930]" country="Indonesia" name="Bali" pageId="3" pageNumber="72">Bali</collectingRegion>
). In low elevation dry scrub forests.
</paragraph>
</subSubSection>
<subSubSection id="C330657223371C64AC8124FCFB03F835" box="[198,1184,1941,1966]" pageId="3" pageNumber="72" type="vernacular_names">
<paragraph id="8B9536F923371C64AC8124FCFB03F835" blockId="3.[151,1436,1077,2038]" box="[198,1184,1941,1966]" pageId="3" pageNumber="72">
<emphasis id="B95EEAEB23371C64AC8124FCFE71F835" bold="true" box="[198,466,1941,1966]" pageId="3" pageNumber="72">Local names and uses:</emphasis>
@ -217,8 +198,6 @@ and
<collectingRegion id="49EEF81B23371C64A82724FCFB37F835" box="[1120,1172,1941,1966]" country="Indonesia" name="Bali" pageId="3" pageNumber="72">Bali</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C330657223371C64AC8124D0FC47F86D" pageId="3" pageNumber="72" type="etymology">
<paragraph id="8B9536F923371C64AC8124D0FC47F86D" blockId="3.[151,1436,1077,2038]" pageId="3" pageNumber="72">
<emphasis id="B95EEAEB23371C64AC8124D0FEF2F849" bold="true" box="[198,337,1977,2002]" pageId="3" pageNumber="72">Etymology:</emphasis>
—The specific epithet
@ -231,7 +210,6 @@ and
<collectingRegion id="49EEF81B23371C64AFEA24B4FC7CF86D" box="[941,991,2013,2038]" country="Indonesia" name="Bali" pageId="3" pageNumber="72">Bali</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="DF55667123301C63ACD0241CFDF3F879" ID-DOI="http://doi.org/10.5281/zenodo.5138220" ID-Zenodo-Dep="5138220" httpUri="https://zenodo.org/record/5138220/files/figure.png" pageId="4" pageNumber="73" startId="4.[151,244,1909,1931]" targetBox="[201,1391,218,1877]" targetPageId="4">
<paragraph id="8B9536F923301C63ACD0241CFDF3F879" blockId="4.[151,1437,1909,2018]" pageId="4" pageNumber="73">
<emphasis id="B95EEAEB23301C63ACD0241CFEAEF810" bold="true" box="[151,269,1909,1931]" pageId="4" pageNumber="73">FIGURE 2.</emphasis>
@ -264,99 +242,83 @@ and
(BO). All photos by E.S. Fernando.
</paragraph>
</caption>
<subSubSection id="C330657223311C62AC8123FEFB78FEDB" pageId="5" pageNumber="74" type="materials_examined">
<paragraph id="8B9536F923311C62AC8123FEFB78FEDB" blockId="5.[151,1436,151,608]" pageId="5" pageNumber="74">
<emphasis id="B95EEAEB23311C62AC8123FEFDF4FF2B" bold="true" box="[198,599,151,176]" pageId="5" pageNumber="74">Additional specimens examined:</emphasis>
<materialsCitation id="3B423CA423311C62AE3323FEFE4EFF4E" collectionCode="BO, K" collectorName="Warsita Mahyar" country="Indonesia" elevation="100" location="Sukamade, Betiri Nature Reserve" pageId="5" pageNumber="74" specimenCount="1" stateProvince="East Java">
<materialsCitation id="3B423CA423311C62AE3323FEFE4EFF4E" pageId="5" pageNumber="74">
<collectingCountry id="F33D766923311C62AE3323FEFCACFF2B" box="[628,783,151,176]" name="Indonesia" pageId="5" pageNumber="74">INDONESIA</collectingCountry>
.
<collectingRegion id="49EEF81B23311C62AF6623F1FC36FF34" box="[801,917,152,175]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="74">East Java</collectingRegion>
:
<location id="8EF5602223311C62AFE123FEFA96FF34" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AFE123FEFA96FF34" box="[934,1333,151,176]" country="Indonesia" name="Sukamade, Betiri Nature Reserve" pageId="5" pageNumber="74" stateProvince="East Java">Sukamade, Betiri Nature Reserve</location>
<collectingCounty id="62F44E7523311C62AFE123FEFBBDFF2B" box="[934,1054,151,176]" pageId="5" pageNumber="74">Sukamade</collectingCounty>
,
<collectingMunicipality id="6BF1AC8323311C62A87723FEFA96FF34" box="[1072,1333,151,176]" pageId="5" pageNumber="74">Betiri Nature Reserve</collectingMunicipality>
,
<quantity id="4CD29B1C23311C62A90023FEFA34FF34" box="[1351,1431,151,176]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="74" unit="m" value="100.0">
<elevation id="0007D1CA23311C62A90023FEFA34FF34" box="[1351,1431,151,176]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="74" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<emphasis id="B95EEAEB23311C62ACD023D4FED0FF4E" box="[151,371,188,213]" italics="true" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62ACD023D4FEECFF4E" box="[151,335,188,213]" pageId="5" pageNumber="74">Warsita Mahyar</collectorName>
<location id="8EF5602223311C62ACD023D4FEECFF4E" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62ACD023D4FEECFF4E" box="[151,335,188,213]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Warsita Mahyar" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Warsita Mahyar</location>
14
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62ADC323D5FE10FF4E" box="[388,435,188,213]" pageId="5" pageNumber="74">BO</collectionCode>
!,
<collectionCode id="ED3BAE3C23311C62AD8323D5FE7CFF4E" box="[452,479,188,213]" pageId="5" pageNumber="74">K</collectionCode>
!)
(BO!, K!)
</materialsCitation>
;
<materialsCitation id="3B423CA423311C62ADBC23D5FB2DFF4E" box="[507,1166,188,213]" collectionCode="BO" collectorName="Versteegh" elevation="150" location="Gunung Watangan" municipality="Jember" pageId="5" pageNumber="74" specimenCount="1">
<collectingMunicipality id="6BF1AC8323311C62ADBC23D5FDECFF4E" box="[507,591,188,213]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Jember" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Jember</collectingMunicipality>
<materialsCitation id="3B423CA423311C62ADBC23D5FB2DFF4E" box="[507,1166,188,213]" pageId="5" pageNumber="74">
<location id="8EF5602223311C62ADBC23D5FDECFF4E" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62ADBC23D5FDECFF4E" box="[507,591,188,213]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Jember" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Jember</location>
,
<location id="8EF5602223311C62AE1D23D5FC93FF4F" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AE1D23D5FC93FF4F" box="[602,816,188,213]" country="Indonesia" county="Sukamade" municipality="Jember" name="Gunung Watangan" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Gunung Watangan</location>
<location id="8EF5602223311C62AE1D23D5FC93FF4F" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AE1D23D5FC93FF4F" box="[602,816,188,213]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Gunung Watangan" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Gunung Watangan</location>
,
<quantity id="4CD29B1C23311C62AF7B23D5FC24FF48" box="[828,903,188,213]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="74" unit="m" value="150.0">
<elevation id="0007D1CA23311C62AF7B23D5FC24FF48" box="[828,903,188,213]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="74" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<emphasis id="B95EEAEB23311C62AFD423D5FBE1FF4E" box="[915,1090,188,213]" italics="true" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62AFD423D5FBA2FF4E" box="[915,1025,188,213]" pageId="5" pageNumber="74">Versteegh</collectorName>
<location id="8EF5602223311C62AFD423D5FBA2FF4E" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AFD423D5FBA2FF4E" box="[915,1025,188,213]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Versteegh" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Versteegh</location>
4881
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62A81523D5FB22FF4E" box="[1106,1153,188,213]" pageId="5" pageNumber="74">BO</collectionCode>
!)
(BO!)
</materialsCitation>
,
<materialsCitation id="3B423CA423311C62A8DC23D5FA34FF4E" box="[1179,1431,188,213]" collectionCode="BO" collectorName="Versteegh" pageId="5" pageNumber="74" specimenCount="1">
<materialsCitation id="3B423CA423311C62A8DC23D5FA34FF4E" box="[1179,1431,188,213]" pageId="5" pageNumber="74">
<emphasis id="B95EEAEB23311C62A8DC23D5FAEAFF4E" box="[1179,1353,188,213]" italics="true" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62A8DC23D5FAAAFF4E" box="[1179,1289,188,213]" pageId="5" pageNumber="74">Versteegh</collectorName>
<location id="8EF5602223311C62A8DC23D5FAAAFF4E" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62A8DC23D5FAAAFF4E" box="[1179,1289,188,213]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Versteegh" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Versteegh</location>
4882
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62A91C23D5FA29FF4E" box="[1371,1418,188,213]" pageId="5" pageNumber="74">BO</collectionCode>
!)
(BO!)
</materialsCitation>
,
<materialsCitation id="3B423CA423311C62ACD023B6FE2DFF63" box="[151,398,223,248]" collectionCode="BO" collectorName="Versteegh" pageId="5" pageNumber="74" specimenCount="1">
<materialsCitation id="3B423CA423311C62ACD023B6FE2DFF63" box="[151,398,223,248]" pageId="5" pageNumber="74">
<emphasis id="B95EEAEB23311C62ACD023B6FEE1FF63" box="[151,322,223,248]" italics="true" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62ACD023B6FEA7FF63" box="[151,260,223,248]" pageId="5" pageNumber="74">Versteegh</collectorName>
<location id="8EF5602223311C62ACD023B6FEA7FF63" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62ACD023B6FEA7FF63" box="[151,260,223,248]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Versteegh" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Versteegh</location>
4885
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62AD1523B6FE22FF63" box="[338,385,223,248]" pageId="5" pageNumber="74">BO</collectionCode>
!)
(BO!)
</materialsCitation>
;
<materialsCitation id="3B423CA423311C62ADDD23B6FCB6FF63" box="[410,789,223,248]" collectionCode="BO" collectorName="Versteegh" location="Panarukan" municipality="Panarukan" pageId="5" pageNumber="74" specimenCount="1">
<collectingMunicipality id="6BF1AC8323311C62ADDD23B6FDB0FF63" box="[410,531,223,248]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Panarukan" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Panarukan</collectingMunicipality>
<materialsCitation id="3B423CA423311C62ADDD23B6FCB6FF63" box="[410,789,223,248]" pageId="5" pageNumber="74">
<location id="8EF5602223311C62ADDD23B6FDB0FF63" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62ADDD23B6FDB0FF63" box="[410,531,223,248]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Panarukan" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Panarukan</location>
,
<emphasis id="B95EEAEB23311C62AE5923B6FD6AFF63" box="[542,713,223,248]" italics="true" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62AE5923B6FD28FF63" box="[542,651,223,248]" pageId="5" pageNumber="74">Versteegh</collectorName>
<location id="8EF5602223311C62AE5923B6FD28FF63" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AE5923B6FD28FF63" box="[542,651,223,248]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Versteegh" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Versteegh</location>
4892
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62AE9E23B6FCABFF63" box="[729,776,223,248]" pageId="5" pageNumber="74">BO</collectionCode>
!)
(BO!)
</materialsCitation>
;
<materialsCitation id="3B423CA423311C62AF6623B6FA34FF63" box="[801,1431,223,248]" collectingDate="1940-03-07" collectionCode="BO" collectorName="Versteegh" location="Gunung Baluran" municipality="Gunung Baluran" pageId="5" pageNumber="74" specimenCount="1">
<collectingMunicipality id="6BF1AC8323311C62AF6623B6FC7DFF63" box="[801,990,223,248]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Gunung Baluran" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Gunung Baluran</collectingMunicipality>
<materialsCitation id="3B423CA423311C62AF6623B6FA34FF63" box="[801,1431,223,248]" pageId="5" pageNumber="74">
<location id="8EF5602223311C62AF6623B6FC7DFF63" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AF6623B6FC7DFF63" box="[801,990,223,248]" country="Indonesia" county="Sukamade" municipality="Betiri Nature Reserve" name="Gunung Baluran" pageId="5" pageNumber="74" stateProvince="Jawa Timur">Gunung Baluran</location>
,
<date id="FF94103923311C62AFAF23B6FB36FF63" box="[1000,1173,223,248]" pageId="5" pageNumber="74" value="1940-03-07">
<collectingDate id="EFD0E9D123311C62AFAF23B6FB36FF63" box="[1000,1173,223,248]" pageId="5" pageNumber="74" value="1940-03-07">07 March 1940</collectingDate>
</date>
,
<emphasis id="B95EEAEB23311C62A8E723B6FAEFFF63" box="[1184,1356,223,248]" italics="true" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62A8E723B6FAAEFF63" box="[1184,1293,223,248]" pageId="5" pageNumber="74">Versteegh</collectorName>
4893
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62A91C23B6FA29FF63" box="[1371,1418,223,248]" pageId="5" pageNumber="74">BO</collectionCode>
!)
<emphasis id="B95EEAEB23311C62A8E723B6FAEFFF63" box="[1184,1356,223,248]" italics="true" pageId="5" pageNumber="74">Versteegh 4893</emphasis>
(BO!)
</materialsCitation>
,
<materialsCitation id="3B423CA423311C62ACD0226DFD99FE86" box="[151,570,260,285]" collectingDate="1940-04-25" collectionCode="BO" collectorName="Versteegh" pageId="5" pageNumber="74" specimenCount="1">
<materialsCitation id="3B423CA423311C62ACD0226DFD99FE86" box="[151,570,260,285]" pageId="5" pageNumber="74">
<date id="FF94103923311C62ACD0226DFE94FE86" box="[151,311,260,285]" pageId="5" pageNumber="74" value="1940-04-25">
<collectingDate id="EFD0E9D123311C62ACD0226DFE94FE86" box="[151,311,260,285]" pageId="5" pageNumber="74" value="1940-04-25">25 April 1940</collectingDate>
</date>
@ -365,48 +327,37 @@ and
<collectorName id="26DF532F23311C62AD05226DFE0CFE86" box="[322,431,260,285]" pageId="5" pageNumber="74">Versteegh</collectorName>
4915
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62ADB9226DFD8EFE86" box="[510,557,260,285]" pageId="5" pageNumber="74">BO</collectionCode>
!)
(BO!)
</materialsCitation>
.
<materialsCitation id="3B423CA423311C62AE02226DFEA7FEDB" collectingDate="1973-05-19" collectionCode="BO, L" collectorName="Dransfield" county="Cakik" location="Suaka, Margasatwa" municipality="Gilimanuk" pageId="5" pageNumber="74" specimenCount="1" stateProvince="Bali">
<materialsCitation id="3B423CA423311C62AE02226DFEA7FEDB" pageId="5" pageNumber="74">
<collectingRegion id="49EEF81B23311C62AE02226DFDDBFE86" box="[581,632,260,285]" country="Indonesia" name="Bali" pageId="5" pageNumber="74">Bali</collectingRegion>
:
<collectingCounty id="62F44E7523311C62AEC4226DFD6AFE86" box="[643,713,260,285]" pageId="5" pageNumber="74">Cakik</collectingCounty>
,
<collectingMunicipality id="6BF1AC8323311C62AE93226DFCECFE86" box="[724,847,260,285]" pageId="5" pageNumber="74">Gilimanuk</collectingMunicipality>
,
<location id="8EF5602223311C62AF1E226DFB94FE87" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AF1E226DFB94FE87" box="[857,1079,260,285]" county="Cakik" municipality="Gilimanuk" name="Suaka, Margasatwa" pageId="5" pageNumber="74" stateProvince="Bali">Suaka, Margasatwa</location>
<location id="8EF5602223311C62AF1E226DFC03FE86" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AF1E226DFC03FE86" box="[857,928,260,285]" country="Indonesia" county="Cakik" municipality="Gilimanuk" name="Suaka" pageId="5" pageNumber="74" stateProvince="Bali">Suaka</location>
,
<location id="8EF5602223311C62AFEB226DFB94FE87" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AFEB226DFB94FE87" box="[940,1079,260,284]" country="Indonesia" county="Cakik" municipality="Gilimanuk" name="Margasatwa" pageId="5" pageNumber="74" stateProvince="Bali">Margasatwa</location>
,
<date id="FF94103923311C62A805226DFB7AFE86" box="[1090,1241,260,285]" pageId="5" pageNumber="74" value="1973-05-19">
<collectingDate id="EFD0E9D123311C62A805226DFB7AFE86" box="[1090,1241,260,285]" pageId="5" pageNumber="74" value="1973-05-19">19 May 1973</collectingDate>
</date>
,
<emphasis id="B95EEAEB23311C62A8A2226DFA3FFE86" box="[1253,1436,260,285]" italics="true" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62A8A2226DFAFEFE86" box="[1253,1373,260,285]" pageId="5" pageNumber="74">Dransfield</collectorName>
3519
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62ACE7224EFF6CFEDB" box="[160,207,295,320]" pageId="5" pageNumber="74">BO</collectionCode>
!,
<collectionCode id="ED3BAE3C23311C62AC99224EFF56FEDB" box="[222,245,295,320]" pageId="5" pageNumber="74">L</collectionCode>
!)
<emphasis id="B95EEAEB23311C62A8A2226DFA3FFE86" box="[1253,1436,260,285]" italics="true" pageId="5" pageNumber="74">Dransfield 3519</emphasis>
(BO!, L!)
</materialsCitation>
;
<materialsCitation id="3B423CA423311C62AD56224EFCA4FEDB" box="[273,775,295,320]" collectionCode="BO" collectorName="R. Maier Sarip" location="Gunung Kelalakan" pageId="5" pageNumber="74" specimenCount="1">
<location id="8EF5602223311C62AD56224EFE44FEDB" LSID="urn:lsid:plazi:treatment:038387EF23371C62ACD0200EFE02FDFB:8EF5602223311C62AD56224EFE44FEDB" box="[273,487,295,320]" name="Gunung Kelalakan" pageId="5" pageNumber="74">Gunung Kelalakan</location>
,
<collectorName id="26DF532F23311C62ADB52240FD3AFEA4" box="[498,665,296,320]" pageId="5" pageNumber="74">
<materialsCitation id="3B423CA423311C62AD56224EFCA4FEDB" box="[273,775,295,320]" pageId="5" pageNumber="74">
<collectorName id="26DF532F23311C62AD56224EFDA9FEDB" box="[273,522,295,320]" pageId="5" pageNumber="74">
Gunung Kelalakan,
<emphasis id="B95EEAEB23311C62ADB52240FDA9FEDB" box="[498,522,297,320]" italics="true" pageId="5" pageNumber="74">R.</emphasis>
Maier Sarip
</collectorName>
98 (
<collectionCode id="ED3BAE3C23311C62AE8B224EFD59FEDB" box="[716,762,295,320]" pageId="5" pageNumber="74">BO</collectionCode>
!)
Maier Sarip 98 (BO!)
</materialsCitation>
,
<materialsCitation id="3B423CA423311C62AF54224EFB74FEDB" box="[787,1239,295,320]" collectingDate="1918-07-20" collectionCode="BO" collectorName="R. Maier Sarip" pageId="5" pageNumber="74" specimenCount="1">
<materialsCitation id="3B423CA423311C62AF54224EFB74FEDB" box="[787,1239,295,320]" pageId="5" pageNumber="74">
<date id="FF94103923311C62AF54224EFC04FEDB" box="[787,935,295,320]" pageId="5" pageNumber="74" value="1918-07-20">
<collectingDate id="EFD0E9D123311C62AF54224EFC04FEDB" box="[787,935,295,320]" pageId="5" pageNumber="74" value="1918-07-20">20 July 1918</collectingDate>
</date>
@ -415,14 +366,10 @@ Maier Sarip
<collectorName id="26DF532F23311C62AFF52240FBF9FEA4" box="[946,1114,296,320]" pageId="5" pageNumber="74">R. Maier Sarip</collectorName>
120
</emphasis>
(
<collectionCode id="ED3BAE3C23311C62A8DD224EFB6AFEDB" box="[1178,1225,295,320]" pageId="5" pageNumber="74">BO</collectionCode>
!)
(BO!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330657223311C62AC812225FE02FDFB" pageId="5" pageNumber="74" type="discussion">
<paragraph id="8B9536F923311C62AC812225FE02FDFB" blockId="5.[151,1436,151,608]" pageId="5" pageNumber="74">
<emphasis id="B95EEAEB23311C62AC812225FEB3FEFE" bold="true" box="[198,272,332,357]" inLineHeading="true" pageId="5" pageNumber="74" reason="1">Notes.</emphasis>
Specimens belonging to
@ -493,6 +440,5 @@ are referrable to
</taxonomicName>
<emphasis id="B95EEAEB23311C62ADDA212EFE02FDFB" box="[413,417,583,608]" italics="true" pageId="5" pageNumber="74">.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,328 @@
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<mods:title id="F79C5E901DD05A89ED70A912787E6EDB">Description Of The Larva Of Nicagus Bscurus (Leconte) (Coleoptera: Lucanidae: Nicaginae), With Comments On Its Position In Lucanidae And Notes On The Larval And Adult Habitat</mods:title>
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<mods:affiliation id="F4F5BDFD8F36E9F686215981FBAD6C4D">Department of Entomology University of Wisconsin Madison, Wisconsin 53706, U. S. A.</mods:affiliation>
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<mods:date id="E8BF50B39F01E1B5006712E39BDE1E05">2002</mods:date>
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<treatment id="AD3F87A3FFB16A45FDA5FA2AFE77FC2D" LSID="urn:lsid:plazi:treatment:AD3F87A3FFB16A45FDA5FA2AFE77FC2D" httpUri="http://treatment.plazi.org/id/AD3F87A3FFB16A45FDA5FA2AFE77FC2D" lastPageId="4" lastPageNumber="257" pageId="0" pageNumber="253">
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<heading id="7E6181D9FFB16A41FDA5FA2AFCB8FA48" bold="true" box="[568,782,1484,1505]" centered="true" fontSize="9" level="1" pageId="0" pageNumber="253" reason="1">
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Larval
<taxonomicName id="E2964D36FFB16A41FD10FA2AFCB8FA48" ID-CoL="73XS3" authority="(LECONTE)" baseAuthorityName="LeConte" box="[653,782,1484,1505]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="obscurus">Description</taxonomicName>
</emphasis>
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<paragraph id="252936B5FFB16A41FE80FA12FBBBF997" blockId="0.[259,1091,1524,1652]" pageId="0" pageNumber="253">
Last instar larvae were
<quantity id="E26E9B50FFB16A41FD84FA12FDCFF9A0" box="[537,633,1524,1545]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="0" pageNumber="253" unit="mm" value="17.5">17.5 mm</quantity>
long, maximum width of head capsule
<quantity id="E26E9B50FFB16A41FBBFFA12FE98F98D" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="0" pageNumber="253" unit="mm" value="2.4">2.4 mm</quantity>
, width across dorsum of 2nd abdominal segment
<quantity id="E26E9B50FFB16A41FCA6F9E9FC39F98D" box="[827,911,1551,1572]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="0" pageNumber="253" unit="mm" value="2.9">2.9 mm</quantity>
. Body C­shaped; cranium and clypeus pale yellow, thoracic and abdominal tergites white.
</paragraph>
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<emphasis id="17E2EAA7FFB16A41FE80F9A2FEE9F9F0" bold="true" box="[285,351,1604,1625]" pageId="0" pageNumber="253">Head.</emphasis>
Frontal suture distinctly U­shaped, terminating in a broad opening (
<figureCitation id="BDAD2A30FFB16A41FE97F9B9FEFAF9DD" box="[266,332,1631,1652]" captionStart="Figs" captionStartId="1.[279,322,984,1003]" captionTargetBox="[468,874,266,962]" captionTargetId="figure-12@1.[467,875,265,964]" captionTargetPageId="1" captionText="Figs. 13. Nicagus obscurus. 1) Head, anterodorsal view; 2) antenna; 3) epipharynx, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10103143" httpUri="https://zenodo.org/record/10103143/files/figure.png" pageId="0" pageNumber="253">Fig. 1</figureCitation>
). Frons with paired, long primary frontal setae, paired, long exterior
</paragraph>
<caption id="71E9663DFFB06A40FE8AFC3EFECDFBAD" ID-DOI="http://doi.org/10.5281/zenodo.10103143" ID-Zenodo-Dep="10103143" httpUri="https://zenodo.org/record/10103143/files/figure.png" pageId="1" pageNumber="254" startId="1.[279,322,984,1003]" targetBox="[468,874,266,962]" targetPageId="1">
<paragraph id="252936B5FFB06A40FE8AFC3EFECDFBAD" blockId="1.[255,1088,984,1028]" pageId="1" pageNumber="254">
<emphasis id="17E2EAA7FFB06A40FE8AFC3EFEC3FC42" bold="true" box="[279,373,984,1003]" pageId="1" pageNumber="254">Figs. 13.</emphasis>
<taxonomicName id="E2964D36FFB06A40FE11FC3FFD85FC45" box="[396,563,985,1004]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="254" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="17E2EAA7FFB06A40FE11FC3FFD85FC45" box="[396,563,985,1004]" italics="true" pageId="1" pageNumber="254">Nicagus obscurus</emphasis>
</taxonomicName>
.
<emphasis id="17E2EAA7FFB06A40FDA2FC3EFDE4FC42" bold="true" box="[575,594,984,1003]" pageId="1" pageNumber="254">1)</emphasis>
Head, anterodorsal view;
<emphasis id="17E2EAA7FFB06A40FCD4FC3EFCEAFC42" bold="true" box="[841,860,984,1003]" pageId="1" pageNumber="254">2)</emphasis>
antenna;
<emphasis id="17E2EAA7FFB06A40FC2AFC3EFC7CFC42" bold="true" box="[951,970,984,1003]" pageId="1" pageNumber="254">3)</emphasis>
epipharynx, ventral view.
</paragraph>
</caption>
<paragraph id="252936B5FFB06A43FF62FBAFFE3CFB7D" blockId="1.[255,1088,1097,1652]" lastBlockId="2.[259,1092,1001,1502]" lastPageId="2" lastPageNumber="255" pageId="1" pageNumber="254">
setae, and paired, long anterior angle setae. Clypeus trapezoidal; dorsolateral region of clypeus with several small setae and three long setae on each side; preclypeus and postclypeus lacking setae. Labrum with anterior angles rounded; anterior margin crenulate, setose, surface with posterior­medial pair of long setae; posterior margin with scattered, short setae. Antennae (
<figureCitation id="BDAD2A30FFB06A40FC35FB52FC5DFB60" box="[936,1003,1204,1225]" captionStart="Figs" captionStartId="1.[279,322,984,1003]" captionTargetBox="[468,874,266,962]" captionTargetId="figure-12@1.[467,875,265,964]" captionTargetPageId="1" captionText="Figs. 13. Nicagus obscurus. 1) Head, anterodorsal view; 2) antenna; 3) epipharynx, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10103143" httpUri="https://zenodo.org/record/10103143/files/figure.png" pageId="1" pageNumber="254">Fig. 2</figureCitation>
) 4­segmented; 1st segment short, cylindrical, 1/6 length of 2nd segment; 2nd segment long, clavate; 3rd segment clavate, widening distally, apical surface broadly rounded, bearing large sensory spot; 4th segment narrow, apex with two short setae, and a single long seta laterally. Epipharynx (
<figureCitation id="BDAD2A30FFB06A40FCA9FAF9FCC0FA9D" box="[820,886,1311,1332]" captionStart="Figs" captionStartId="1.[279,322,984,1003]" captionTargetBox="[468,874,266,962]" captionTargetId="figure-12@1.[467,875,265,964]" captionTargetPageId="1" captionText="Figs. 13. Nicagus obscurus. 1) Head, anterodorsal view; 2) antenna; 3) epipharynx, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10103143" httpUri="https://zenodo.org/record/10103143/files/figure.png" pageId="1" pageNumber="254">Fig. 3</figureCitation>
) broadly rounded, anterior margin lined with long, broad setae. Chaetoparia with four distinct, large setae. Phoba with oval pattern of small setae. Pedium with small patch of granules on dexiophoba side. Protophoba with three blunt, short setae. Epipharynx with united tormae and three nesia: right nesium ovate, median nesium elongate, wider at posterior end, slightly tapering toward anterior end, left nesium small, tear shaped. Mandibles black along margins, well sclerotized,
<quantity id="E26E9B50FFB06A40FF62FA3FFEE7FA47" box="[255,337,1497,1518]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="254" unit="mm" value="0.9">0.9 mm</quantity>
long. Left mandible (
<figureCitation id="BDAD2A30FFB06A40FDA4FA3FFD3BFA47" box="[569,653,1497,1518]" captionStart="Figs" captionStartId="2.[282,325,899,918]" captionTargetBox="[473,881,269,875]" captionTargetId="figure-12@2.[467,883,265,878]" captionTargetPageId="2" captionText="Figs. 46. Nicagus obscurus. 4A) Left mandible, dorsal view; 4B) right mandible, dorsal view; 5) right maxilla, ventral view; 6) labium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10103145" httpUri="https://zenodo.org/record/10103145/files/figure.png" pageId="1" pageNumber="254">Fig. 4A</figureCitation>
) in dorsal view with blunt scissoral area, slightly bilobed in appearance; ventral process in lateral view prominent under molar area. Right mandible (
<figureCitation id="BDAD2A30FFB06A40FDACF9E9FD35F98D" box="[561,643,1551,1572]" captionStart="Figs" captionStartId="2.[282,325,899,918]" captionTargetBox="[473,881,269,875]" captionTargetId="figure-12@2.[467,883,265,878]" captionTargetPageId="2" captionText="Figs. 46. Nicagus obscurus. 4A) Left mandible, dorsal view; 4B) right mandible, dorsal view; 5) right maxilla, ventral view; 6) labium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10103145" httpUri="https://zenodo.org/record/10103145/files/figure.png" pageId="1" pageNumber="254">Fig. 4B</figureCitation>
) in dorsal view with scissorial area blunt, weakly bilobed, angled aborally; ventral process in lateral view prominent below molar area. Maxilla (
<figureCitation id="BDAD2A30FFB06A40FDB8F9A2FDD2F9F0" box="[549,612,1604,1625]" captionStart="Figs" captionStartId="2.[282,325,899,918]" captionTargetBox="[473,881,269,875]" captionTargetId="figure-12@2.[467,883,265,878]" captionTargetPageId="2" captionText="Figs. 46. Nicagus obscurus. 4A) Left mandible, dorsal view; 4B) right mandible, dorsal view; 5) right maxilla, ventral view; 6) labium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10103145" httpUri="https://zenodo.org/record/10103145/files/figure.png" pageId="1" pageNumber="254">Fig. 5</figureCitation>
) with galea and lacinia separated; galea thin, tapering to a blunt apex, medially with a pair of setae; lacinia with sclerotized apex, adoral lateral margin lined with nine pairs of setae; maxillary palpus 4­ segmented: 1st segment globular, aboral lateral margin with a single seta; 2nd segment ovate, subequal to 3rd; 3rd segment ovate, ringed distally with four setae; 4th segment ovate, subequal to 2nd and 3rd segments combined, distally with a sensory patch. Labium (
<figureCitation id="BDAD2A30FFB36A43FDD0FBB2FD3BFBC0" box="[589,653,1108,1129]" captionStart="Figs" captionStartId="2.[282,325,899,918]" captionTargetBox="[473,881,269,875]" captionTargetId="figure-12@2.[467,883,265,878]" captionTargetPageId="2" captionText="Figs. 46. Nicagus obscurus. 4A) Left mandible, dorsal view; 4B) right mandible, dorsal view; 5) right maxilla, ventral view; 6) labium, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10103145" httpUri="https://zenodo.org/record/10103145/files/figure.png" pageId="2" pageNumber="255">Fig. 6</figureCitation>
) quadrate, edges slightly sinuate, bearing four distinct, small setae (two medially, two posteriomedially); two V­shaped patches of large setae on anterior margin; labial palpus 2­segmented (appearing almost 3­segmented), last oval, 2× length of first, apically with several circular sensory pits.
</paragraph>
<caption id="71E9663DFFB36A43FE87FC65FC3EFC07" ID-DOI="http://doi.org/10.5281/zenodo.10103145" ID-Zenodo-Dep="10103145" httpUri="https://zenodo.org/record/10103145/files/figure.png" pageId="2" pageNumber="255" startId="2.[282,325,899,918]" targetBox="[473,881,269,875]" targetPageId="2">
<paragraph id="252936B5FFB36A43FE87FC65FC3EFC07" blockId="2.[259,1092,899,942]" pageId="2" pageNumber="255">
<emphasis id="17E2EAA7FFB36A43FE87FC65FECCFC3F" bold="true" box="[282,378,899,918]" pageId="2" pageNumber="255">Figs. 46.</emphasis>
<taxonomicName id="E2964D36FFB36A43FE0CFC65FD8CFC3F" box="[401,570,899,918]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="17E2EAA7FFB36A43FE0CFC65FD8CFC3F" box="[401,570,899,918]" italics="true" pageId="2" pageNumber="255">Nicagus obscurus</emphasis>
</taxonomicName>
.
<emphasis id="17E2EAA7FFB36A43FDD4FC65FDDBFC3F" bold="true" box="[585,621,899,918]" pageId="2" pageNumber="255">4A)</emphasis>
Left mandible, dorsal view;
<emphasis id="17E2EAA7FFB36A43FC1AFC65FC1CFC3F" bold="true" box="[903,938,899,918]" pageId="2" pageNumber="255">4B)</emphasis>
right mandible, dorsal view;
<emphasis id="17E2EAA7FFB36A43FEE3FC7DFE24FC07" bold="true" box="[382,402,923,942]" pageId="2" pageNumber="255">5)</emphasis>
right maxilla, ventral view;
<emphasis id="17E2EAA7FFB36A43FD39FC7DFD01FC07" bold="true" box="[676,695,923,942]" pageId="2" pageNumber="255">6)</emphasis>
labium, ventral view.
</paragraph>
</caption>
<paragraph id="252936B5FFB36A43FE80FB3FFB8FFAF0" blockId="2.[259,1092,1001,1502]" pageId="2" pageNumber="255">
<emphasis id="17E2EAA7FFB36A43FE80FB3FFEC0FB47" bold="true" box="[285,374,1241,1262]" pageId="2" pageNumber="255">Thorax.</emphasis>
Tarsungulus of prothoracic leg (
<figureCitation id="BDAD2A30FFB36A43FD50FB3FFCBAFB47" box="[717,780,1241,1262]" captionStart="Figs" captionStartId="3.[279,322,886,905]" captionTargetBox="[465,877,266,863]" captionTargetId="figure-12@3.[462,881,265,865]" captionTargetPageId="3" captionText="Figs. 711. Nicagus obscurus. 7) Tarsungulus, right prothoracic leg; 8) mesothoracic leg, posterior margin; 9) metathoracic leg, posterior margin; 10) abdominal tergite 7, alar area, lateral view; 11) abdominal segment 10, caudal view." figureDoi="http://doi.org/10.5281/zenodo.10103147" httpUri="https://zenodo.org/record/10103147/files/figure.png" pageId="2" pageNumber="255">Fig. 7</figureCitation>
) with single claw, large setae adjacent to apex; short, blunt, broad seta medially on claw. Mesothoracic leg (
<figureCitation id="BDAD2A30FFB36A43FE97FAE9FEFDFA8D" box="[266,331,1295,1316]" captionStart="Figs" captionStartId="3.[279,322,886,905]" captionTargetBox="[465,877,266,863]" captionTargetId="figure-12@3.[462,881,265,865]" captionTargetPageId="3" captionText="Figs. 711. Nicagus obscurus. 7) Tarsungulus, right prothoracic leg; 8) mesothoracic leg, posterior margin; 9) metathoracic leg, posterior margin; 10) abdominal tergite 7, alar area, lateral view; 11) abdominal segment 10, caudal view." figureDoi="http://doi.org/10.5281/zenodo.10103147" httpUri="https://zenodo.org/record/10103147/files/figure.png" pageId="2" pageNumber="255">Fig. 8</figureCitation>
) with stridulatory area of coxa long, ovate; stridulatory area with vertical line of small, short setae, and grainy surface. Inner metathoracic leg (
<figureCitation id="BDAD2A30FFB36A43FB84FACFFEA7FAF0" captionStart="Figs" captionStartId="3.[279,322,886,905]" captionTargetBox="[465,877,266,863]" captionTargetId="figure-12@3.[462,881,265,865]" captionTargetPageId="3" captionText="Figs. 711. Nicagus obscurus. 7) Tarsungulus, right prothoracic leg; 8) mesothoracic leg, posterior margin; 9) metathoracic leg, posterior margin; 10) abdominal tergite 7, alar area, lateral view; 11) abdominal segment 10, caudal view." figureDoi="http://doi.org/10.5281/zenodo.10103147" httpUri="https://zenodo.org/record/10103147/files/figure.png" pageId="2" pageNumber="255">Fig. 9</figureCitation>
) with oval stridulatory area on trochanter, stridulatory area with 14 ridges.
</paragraph>
<paragraph id="252936B5FFB36A43FE80FAB8FC77FA77" blockId="2.[259,1092,1001,1502]" pageId="2" pageNumber="255">
<emphasis id="17E2EAA7FFB36A43FE80FAB8FE27FADA" bold="true" box="[285,401,1374,1395]" pageId="2" pageNumber="255">Abdomen.</emphasis>
Abdomen consisting of 10, non­plicate segments. Spiracles (
<figureCitation id="BDAD2A30FFB36A43FB84FAB9FEA8FA27" captionStart="Figs" captionStartId="3.[279,322,886,905]" captionTargetBox="[465,877,266,863]" captionTargetId="figure-12@3.[462,881,265,865]" captionTargetPageId="3" captionText="Figs. 711. Nicagus obscurus. 7) Tarsungulus, right prothoracic leg; 8) mesothoracic leg, posterior margin; 9) metathoracic leg, posterior margin; 10) abdominal tergite 7, alar area, lateral view; 11) abdominal segment 10, caudal view." figureDoi="http://doi.org/10.5281/zenodo.10103147" httpUri="https://zenodo.org/record/10103147/files/figure.png" pageId="2" pageNumber="255">Fig. 10</figureCitation>
) cribriform, oriented cephalad. Last (=10th) abdominal segment with anal opening Y­ shaped (
<figureCitation id="BDAD2A30FFB36A43FE48FA72FD94FA00" box="[469,546,1428,1449]" captionStart="Figs" captionStartId="3.[279,322,886,905]" captionTargetBox="[465,877,266,863]" captionTargetId="figure-12@3.[462,881,265,865]" captionTargetPageId="3" captionText="Figs. 711. Nicagus obscurus. 7) Tarsungulus, right prothoracic leg; 8) mesothoracic leg, posterior margin; 9) metathoracic leg, posterior margin; 10) abdominal tergite 7, alar area, lateral view; 11) abdominal segment 10, caudal view." figureDoi="http://doi.org/10.5281/zenodo.10103147" httpUri="https://zenodo.org/record/10103147/files/figure.png" pageId="2" pageNumber="255">Fig. 11</figureCitation>
), bearing two large, ovate lower anal pads caudally, slightly grainy in texture; upper anal pad broadly triangular, conforming with Y­shape; entire caudal region surrounded by circle of large setae.
</paragraph>
</subSubSection>
<subSubSection id="6D8C653EFFB36A45FDA5F9E6FD9FFE97" lastPageId="4" lastPageNumber="257" pageId="2" pageNumber="255" type="discussion">
<paragraph id="252936B5FFB36A43FDA5F9E6FCBBF9BC" blockId="2.[568,781,1536,1557]" box="[568,781,1536,1557]" pageId="2" pageNumber="255">
<heading id="7E6181D9FFB36A43FDA5F9E6FCBBF9BC" bold="true" box="[568,781,1536,1557]" centered="true" fontSize="9" level="1" pageId="2" pageNumber="255" reason="1">
<emphasis id="17E2EAA7FFB36A43FDA5F9E6FCBBF9BC" bold="true" box="[568,781,1536,1557]" pageId="2" pageNumber="255">Generic Placement</emphasis>
</heading>
</paragraph>
<paragraph id="252936B5FFB36A42FE80F9CCFD35FB75" blockId="2.[259,1091,1577,1652]" lastBlockId="3.[255,1088,1033,1652]" lastPageId="3" lastPageNumber="256" pageId="2" pageNumber="255">
<taxonomicName id="E2964D36FFB36A43FE80F9CCFEC5F996" authorityName="Leconte" authorityYear="1861" box="[285,371,1578,1599]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB36A43FE80F9CCFEC5F996" box="[285,371,1578,1599]" italics="true" pageId="2" pageNumber="255">Nicagus</emphasis>
</taxonomicName>
larvae possess all of the traditional synapomorphic characters of
<taxonomicName id="E2964D36FFB36A43FE9EF9A2FEC5F9F0" baseAuthorityName="Lawrence and Newton" baseAuthorityYear="1995" box="[259,371,1604,1625]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
. Previous placement in
<taxonomicName id="E2964D36FFB36A43FDEDF9A2FCDEF9F0" authority="(Kikuta 1986)" baseAuthorityName="Kikuta" baseAuthorityYear="1986" box="[624,872,1604,1625]" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="family">
Trogidae (
<bibRefCitation id="41074B44FFB36A43FD46F9A2FCD4F9F0" author="Kikuta" box="[731,866,1604,1625]" firstAuthor="Kikuta" pageId="2" pageNumber="255" pagination="131 - 138" refId="ref2277" refString="Kikuta, T. 1986. On the higher taxa of the stag beetle family Lucanidae [pp. 131 - 138]. In: Papers on Entomology presented to Professor Takeshiko Nakane in commemoration of his retirement. Japanese Society of Coleopterology, Tokyo." type="book chapter" year="1986">Kikuta 1986</bibRefCitation>
)
</taxonomicName>
was based solely on characters of the adults.
<bibRefCitation id="41074B44FFB36A43FD84F9B9FC8CF9DD" author="Tabana, M. &amp; N. Okuda" box="[537,826,1631,1652]" journalOrPublisher="Gekkan-Mushi" pageId="2" pageNumber="255" pagination="4 - 10" part="256" refId="ref2500" refString="Tabana, M., and N. Okuda. 1992. Notes on Nicagus japonicus Nagel. Gekkan-Mushi 256: 4-10." title="Notes on Nicagus japonicus Nagel" type="journal article" year="1992">Tabana and Okuda (1992)</bibRefCitation>
placed
<taxonomicName id="E2964D36FFB36A43FC05F9B9FB94F9DD" authorityName="Nagel" authorityYear="1928" box="[920,1058,1631,1652]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="17E2EAA7FFB36A43FC05F9B9FB94F9DD" box="[920,1058,1631,1652]" italics="true" pageId="2" pageNumber="255">N. japonicus</emphasis>
</taxonomicName>
in
<taxonomicName id="E2964D36FFB26A42FF62FBEFFEDAFBB7" baseAuthorityName="Lawrence and Newton" baseAuthorityYear="1995" box="[255,364,1033,1054]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
based on five characters of the adults and three characters of the larvae (antennae 4­segmented, abdominal segments without folds, and mesothoracic and metathoracic legs each with a stridulatory organ). They suggested that
<taxonomicName id="E2964D36FFB26A42FEAFFBBCFE3EFBC6" authorityName="Leconte" authorityYear="1861" box="[306,392,1114,1135]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB26A42FEAFFBBCFE3EFBC6" box="[306,392,1114,1135]" italics="true" pageId="3" pageNumber="256">Nicagus</emphasis>
</taxonomicName>
belonged to the lucanid subfamily
<taxonomicName id="E2964D36FFB26A42FC85FBBCFCC9FBC6" authorityName="Macleay" authorityYear="1819" box="[792,895,1114,1135]" pageId="3" pageNumber="256" rank="subFamily" subFamily="Aesalinae">Aesalinae</taxonomicName>
(
<emphasis id="17E2EAA7FFB26A42FC0EFBBCFC7BFBC6" box="[915,973,1114,1135]" italics="true" pageId="3" pageNumber="256">sensu</emphasis>
<bibRefCitation id="41074B44FFB26A42FC44FBBCFE83FB23" author="Holloway" firstAuthor="Holloway" pageId="3" pageNumber="256" pagination="267 - 269" refId="ref2198" refString="Holloway, B. A. 1968. The relationships of Syndesus MacLeay and Sinodendron Schnei- der (Coleoptera: Lucanidae). New Zealand Journal of Science 11: 267 - 269." type="journal article" year="1968">Holloway 1968</bibRefCitation>
) based on similarities of larvae to
<taxonomicName id="E2964D36FFB26A42FD45FB93FC2EFB23" authority="Fabricius" authorityName="Fabricius" authorityYear="1801" box="[728,920,1141,1162]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB26A42FD45FB93FC9FFB23" box="[728,809,1141,1162]" italics="true" pageId="3" pageNumber="256">Aesalus</emphasis>
Fabricius
</taxonomicName>
and
<taxonomicName id="E2964D36FFB26A42FC46FB93FED4FB0C" authority="MacLeay" authorityName="MacLeay" class="Insecta" family="Lucanidae" genus="Ceruchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB26A42FC46FB93FB89FB23" box="[987,1087,1141,1162]" italics="true" pageId="3" pageNumber="256">Ceruchus</emphasis>
MacLeay
</taxonomicName>
in three additional characters: apex of abdomen with three anal lobes, 2nd antennal segment without setae, and left mandible lacking teeth between the 23 apical teeth and molar area.
</paragraph>
<caption id="71E9663DFFB26A42FE8AFC90FC85FC10" ID-DOI="http://doi.org/10.5281/zenodo.10103147" ID-Zenodo-Dep="10103147" httpUri="https://zenodo.org/record/10103147/files/figure.png" pageId="3" pageNumber="256" startId="3.[279,322,886,905]" targetBox="[465,877,266,863]" targetPageId="3">
<paragraph id="252936B5FFB26A42FE8AFC90FC85FC10" blockId="3.[255,1088,886,953]" pageId="3" pageNumber="256">
<emphasis id="17E2EAA7FFB26A42FE8AFC90FE36FC20" bold="true" box="[279,384,886,905]" pageId="3" pageNumber="256">Figs. 711.</emphasis>
<taxonomicName id="E2964D36FFB26A42FE05FC90FD88FC20" box="[408,574,886,905]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="17E2EAA7FFB26A42FE05FC90FD88FC20" box="[408,574,886,905]" italics="true" pageId="3" pageNumber="256">Nicagus obscurus</emphasis>
</taxonomicName>
.
<emphasis id="17E2EAA7FFB26A42FDD7FC90FDEBFC20" bold="true" box="[586,605,886,905]" pageId="3" pageNumber="256">7)</emphasis>
Tarsungulus, right prothoracic leg;
<emphasis id="17E2EAA7FFB26A42FC36FC90FC08FC20" bold="true" box="[939,958,886,905]" pageId="3" pageNumber="256">8)</emphasis>
mesothoracic leg, posterior margin;
<emphasis id="17E2EAA7FFB26A42FE52FC68FE55FC08" bold="true" box="[463,483,910,929]" pageId="3" pageNumber="256">9)</emphasis>
metathoracic leg, posterior margin;
<emphasis id="17E2EAA7FFB26A42FCAAFC68FCE0FC08" bold="true" box="[823,854,910,929]" pageId="3" pageNumber="256">10)</emphasis>
abdominal tergite 7, alar area, lateral view;
<emphasis id="17E2EAA7FFB26A42FE2CFC40FE79FC10" bold="true" box="[433,463,934,953]" pageId="3" pageNumber="256">11)</emphasis>
abdominal segment 10, caudal view.
</paragraph>
</caption>
<paragraph id="252936B5FFB26A45FE87FB04FD9FFE97" blockId="3.[255,1088,1033,1652]" lastBlockId="4.[259,1091,271,318]" lastPageId="4" lastPageNumber="257" pageId="3" pageNumber="256">
<taxonomicName id="E2964D36FFB26A42FE87FB04FE37FB5E" authorityName="Macleay" authorityYear="1819" box="[282,385,1250,1271]" pageId="3" pageNumber="256" rank="subFamily" subFamily="Aesalinae">Aesalinae</taxonomicName>
(
<emphasis id="17E2EAA7FFB26A42FE09FB04FE78FB5E" box="[404,462,1250,1271]" italics="true" pageId="3" pageNumber="256">sensu</emphasis>
<bibRefCitation id="41074B44FFB26A42FE44FB04FD53FB5E" author="Tabana, M. &amp; N. Okuda" box="[473,741,1250,1271]" journalOrPublisher="Gekkan-Mushi" pageId="3" pageNumber="256" pagination="4 - 10" part="256" refId="ref2500" refString="Tabana, M., and N. Okuda. 1992. Notes on Nicagus japonicus Nagel. Gekkan-Mushi 256: 4-10." title="Notes on Nicagus japonicus Nagel" type="journal article" year="1992">Tabana and Okuda 1992</bibRefCitation>
) was based on the inclusion of
<emphasis id="17E2EAA7FFB26A42FF62FB1BFEDDFABB" box="[255,363,1277,1298]" italics="true" pageId="3" pageNumber="256">
<taxonomicName id="E2964D36FFB26A42FF62FB1BFED1FABB" box="[255,359,1277,1298]" class="Insecta" family="Lucanidae" genus="Ceruchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="genus">Ceruchus</taxonomicName>
.
</emphasis>
<bibRefCitation id="41074B44FFB26A42FEE8FB1BFD76FABB" author="Howden, H. F. &amp; J. F. Lawrence" box="[373,704,1277,1298]" journalOrPublisher="Canadian Journal of Zoology" pageId="3" pageNumber="256" pagination="1505 - 1510" part="52" refId="ref2232" refString="Howden, H. F., and J. F. Lawrence. 1974. The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera: Lucanidae). Canadian Journal of Zoology 52: 1505-1510." title="The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera: Lucanidae)." type="journal article" year="1974">Howden and Lawrence (1974)</bibRefCitation>
defined the New World
<taxonomicName id="E2964D36FFB26A42FC4FFB1BFB8AFABB" authorityName="Macleay" authorityYear="1819" box="[978,1084,1277,1298]" pageId="3" pageNumber="256" rank="subFamily" subFamily="Aesalinae">Aesalinae</taxonomicName>
, placing
<taxonomicName id="E2964D36FFB26A42FEC8FAFFFE0FFA87" box="[341,441,1305,1326]" class="Insecta" family="Lucanidae" genus="Ceruchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB26A42FEC8FAFFFE0FFA87" box="[341,441,1305,1326]" italics="true" pageId="3" pageNumber="256">Ceruchus</emphasis>
</taxonomicName>
in Syndesinae, and
<taxonomicName id="E2964D36FFB26A42FD0CFAFFFD51FA87" authorityName="Leconte" authorityYear="1861" box="[657,743,1305,1326]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB26A42FD0CFAFFFD51FA87" box="[657,743,1305,1326]" italics="true" pageId="3" pageNumber="256">Nicagus</emphasis>
</taxonomicName>
in the newly formed
<taxonomicName id="E2964D36FFB26A42FC52FAFEFB8AFA84" authorityName="LeConte" authorityYear="1861" box="[975,1084,1304,1325]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="subFamily" subFamily="Nicaginae">Nicaginae</taxonomicName>
. Larvae of
<taxonomicName id="E2964D36FFB26A42FEE9FAD2FE42FAE0" box="[372,500,1332,1353]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="17E2EAA7FFB26A42FEE9FAD2FE42FAE0" box="[372,500,1332,1353]" italics="true" pageId="3" pageNumber="256">N. obscurus</emphasis>
</taxonomicName>
demonstrate several character states which contradict placement within Syndesinae (
<emphasis id="17E2EAA7FFB26A42FDD1FAA9FD30FACD" box="[588,646,1359,1380]" italics="true" pageId="3" pageNumber="256">sensu</emphasis>
<bibRefCitation id="41074B44FFB26A42FD0CFAA9FC7BFACD" author="Howden, H. F. &amp; J. F. Lawrence" box="[657,973,1359,1380]" journalOrPublisher="Canadian Journal of Zoology" pageId="3" pageNumber="256" pagination="1505 - 1510" part="52" refId="ref2232" refString="Howden, H. F., and J. F. Lawrence. 1974. The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera: Lucanidae). Canadian Journal of Zoology 52: 1505-1510." title="The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera: Lucanidae)." type="journal article" year="1974">Howden and Lawrence 1974</bibRefCitation>
): cephalad spiracle orientation, trochanter of metathoracic leg with single longitudinal row of very short transverse ridges, and the appearance of the three anal lobes, as seen in Lucaninae (
<emphasis id="17E2EAA7FFB26A42FE4EFA47FDBBFA1F" box="[467,525,1441,1462]" italics="true" pageId="3" pageNumber="256">sensu</emphasis>
<bibRefCitation id="41074B44FFB26A42FD8AFA46FCF9FA1C" author="Howden, H. F. &amp; J. F. Lawrence" box="[535,847,1440,1461]" journalOrPublisher="Canadian Journal of Zoology" pageId="3" pageNumber="256" pagination="1505 - 1510" part="52" refId="ref2232" refString="Howden, H. F., and J. F. Lawrence. 1974. The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera: Lucanidae). Canadian Journal of Zoology 52: 1505-1510." title="The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera: Lucanidae)." type="journal article" year="1974">Howden and Lawrence 1974</bibRefCitation>
). The lack of teeth on the inner margin of the mandible, reduction of the mesothoracic and metathoracic tarsungulus, and the lack of setal patches on the raster are suggestive of Syndesinae. The number of setae present on the claw resembles what is seen in Syndesinae as well as some species of Lucaninae. The three distinct tubercles on the protophoba are also seen in
<taxonomicName id="E2964D36FFB26A42FD35F9CFFCBAF997" box="[680,780,1577,1598]" class="Insecta" family="Lucanidae" genus="Ceruchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB26A42FD35F9CFFCBAF997" box="[680,780,1577,1598]" italics="true" pageId="3" pageNumber="256">Ceruchus</emphasis>
</taxonomicName>
; however, there are apparent differences. In view of the conflicting larval characters, and in lieu of any other synapomorphic characters to suggest other relationships among the Lucanidae, it appears that continued placement of
<taxonomicName id="E2964D36FFB56A45FC8FFEE9FCDEFE8D" authorityName="Leconte" authorityYear="1861" box="[786,872,271,292]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB56A45FC8FFEE9FCDEFE8D" box="[786,872,271,292]" italics="true" pageId="4" pageNumber="257">Nicagus</emphasis>
</taxonomicName>
in
<taxonomicName id="E2964D36FFB56A45FC0FFEE9FC4AFE8D" authorityName="LeConte" authorityYear="1861" box="[914,1020,271,292]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="subFamily" subFamily="Nicaginae">Nicaginae</taxonomicName>
is the solution for the time being.
</paragraph>
</subSubSection>
<subSubSection id="6D8C653EFFB56A45FE7DFE85FE77FC2D" pageId="4" pageNumber="257" type="distribution">
<paragraph id="252936B5FFB56A45FE7DFE85FCD0FED1" blockId="4.[480,870,355,376]" box="[480,870,355,376]" pageId="4" pageNumber="257">
<heading id="7E6181D9FFB56A45FE7DFE85FCD0FED1" bold="true" box="[480,870,355,376]" centered="true" fontSize="9" level="1" pageId="4" pageNumber="257" reason="1">
<emphasis id="17E2EAA7FFB56A45FE7DFE85FCD0FED1" bold="true" box="[480,870,355,376]" pageId="4" pageNumber="257">Distribution and Notes on Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="252936B5FFB56A45FE80FE76FE80FD06" blockId="4.[259,1092,399,900]" pageId="4" pageNumber="257">
<taxonomicName id="E2964D36FFB56A45FE80FE76FEC5FE0C" authorityName="Leconte" authorityYear="1861" box="[285,371,400,421]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="17E2EAA7FFB56A45FE80FE76FEC5FE0C" box="[285,371,400,421]" italics="true" pageId="4" pageNumber="257">Nicagus</emphasis>
</taxonomicName>
is reported from
<collectingCountry id="5D817625FFB56A45FDAAFE69FDC5FE0D" box="[567,627,399,420]" name="Japan" pageId="4" pageNumber="257">Japan</collectingCountry>
and the eastern and central
<collectingCountry id="5D817625FFB56A45FC30FE69FBF6FE0D" box="[941,1088,399,420]" name="United States of America" pageId="4" pageNumber="257">United States</collectingCountry>
. Adults occur under or around debris along shorelines of large, fresh water bodies. Distribution has historically been poorly reported for
<emphasis id="17E2EAA7FFB56A45FC0DFE23FBA0FE73" box="[912,1046,453,474]" italics="true" pageId="4" pageNumber="257">
<taxonomicName id="E2964D36FFB56A45FC0DFE23FBA4FE73" box="[912,1042,453,474]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="obscurus">N. obscurus</taxonomicName>
.
</emphasis>
Examinations of, and inquires to, various collections and existing faunal lists (
<bibRefCitation id="41074B44FFB56A45FE97FE1CFE05FDA6" author="Blatchley" box="[266,435,506,527]" firstAuthor="Blatchley" pageId="4" pageNumber="257" refId="ref2015" refString="Blatchley, W. S. 1910. Coleoptera or beetles of Indiana. The Nature Publishing Company, Indianapolis, IN. 1386 pp." type="book" year="1910">Blatchley 1910</bibRefCitation>
;
<bibRefCitation id="41074B44FFB56A45FE58FE1CFDDDFDA6" author="Hamilton" box="[453,619,506,527]" firstAuthor="Hamilton" pageId="4" pageNumber="257" pagination="317 - 381" refId="ref2167" refString="Hamilton, J. 1895. Catalogue of the Coleoptera of southwest Pennsylvania, with notes and descriptions. Transactions of the American Entomological Society 22: 317 - 381." type="journal article" year="1895">Hamilton 1895</bibRefCitation>
;
<bibRefCitation id="41074B44FFB56A45FDE1FE1CFCA0FDA6" author="Leonard" box="[636,790,506,527]" firstAuthor="Leonard" pageId="4" pageNumber="257" refId="ref2423" refString="Leonard, M. D., ed. 1926. A list of the insects of New York, with a list of the spiders and certain other allied groups. Cornell University Agricultural Experimental Station, Ithaca, NY. 1121 pp." type="book" year="1926">Leonard 1926</bibRefCitation>
) yielded several new state records, represented by an asterisk:
<emphasis id="17E2EAA7FFB56A45FD1AFDF3FD55FD83" box="[647,739,533,554]" italics="true" pageId="4" pageNumber="257">
<collectingCountry id="5D817625FFB56A45FD1AFDF3FD6BFD83" box="[647,733,533,554]" name="Canada" pageId="4" pageNumber="257">Canada</collectingCountry>
:
</emphasis>
<collectingRegion id="E752F857FFB56A45FD71FDF3FC89FD83" box="[748,831,533,554]" country="Canada" name="Ontario" pageId="4" pageNumber="257">Ontario</collectingRegion>
,
<collectingRegion id="E752F857FFB56A45FCD6FDF3FC28FD83" box="[843,926,533,554]" country="Canada" name="Quebec" pageId="4" pageNumber="257">Québec</collectingRegion>
;
<emphasis id="17E2EAA7FFB56A45FC36FDF3FBF5FD83" box="[939,1091,533,554]" italics="true" pageId="4" pageNumber="257">
<collectingCountry id="5D817625FFB56A45FC36FDF3FB8BFD83" box="[939,1085,533,554]" name="United States of America" pageId="4" pageNumber="257">United States</collectingCountry>
:
</emphasis>
*
<collectingRegion id="E752F857FFB56A45FE90FDC9FED0FDED" box="[269,358,559,580]" country="United States of America" name="Georgia" pageId="4" pageNumber="257">Georgia</collectingRegion>
,
<collectingRegion id="E752F857FFB56A45FEE5FDC9FE7FFDED" box="[376,457,559,580]" country="United States of America" name="Indiana" pageId="4" pageNumber="257">Indiana</collectingRegion>
,
<collectingRegion id="E752F857FFB56A45FE46FDC9FDF2FDED" box="[475,580,559,580]" country="United States of America" name="Maryland" pageId="4" pageNumber="257">Maryland</collectingRegion>
, *
<collectingRegion id="E752F857FFB56A45FDFCFDC9FD7DFDED" box="[609,715,559,580]" country="United States of America" name="Michigan" pageId="4" pageNumber="257">Michigan</collectingRegion>
, *
<collectingRegion id="E752F857FFB56A45FD7AFDC9FC28FDED" box="[743,926,559,580]" country="United States of America" name="New Hampshire" pageId="4" pageNumber="257">New Hampshire</collectingRegion>
, *
<collectingRegion id="E752F857FFB56A45FC27FDC9FB89FDED" box="[954,1087,559,580]" country="United States of America" name="New Jersey" pageId="4" pageNumber="257">New Jersey</collectingRegion>
,
<collectingRegion id="E752F857FFB56A45FE9EFDACFEC4FDF6" box="[259,370,586,607]" country="United States of America" name="New York" pageId="4" pageNumber="257">New York</collectingRegion>
, *
<collectingRegion id="E752F857FFB56A45FE14FDACFE74FDF6" box="[393,450,586,607]" country="United States of America" name="Ohio" pageId="4" pageNumber="257">Ohio</collectingRegion>
,
<collectingRegion id="E752F857FFB56A45FE53FDACFDEAFDF6" box="[462,604,586,607]" country="United States of America" name="Pennsylvania" pageId="4" pageNumber="257">Pennsylvania</collectingRegion>
, *
<collectingRegion id="E752F857FFB56A45FDEEFDACFCAEFDF6" box="[627,792,586,607]" country="United States of America" name="South Carolina" pageId="4" pageNumber="257">South Carolina</collectingRegion>
, *
<collectingRegion id="E752F857FFB56A45FCB0FDACFC3FFDF6" box="[813,905,586,607]" country="United States of America" name="Virginia" pageId="4" pageNumber="257">Virginia</collectingRegion>
, and *
<collectingRegion id="E752F857FFB56A45FC51FDACFB89FDF6" box="[972,1087,586,607]" country="United States of America" name="Wisconsin" pageId="4" pageNumber="257">Wisconsin</collectingRegion>
. Most records were provided by museum curators or noted on the faunal lists. The authors did not attempt to validate these records or obtain specific locality data.
</paragraph>
<paragraph id="252936B5FFB56A45FE80FD53FE77FC2D" blockId="4.[259,1092,399,900]" pageId="4" pageNumber="257">
Adults and larvae from
<collectingRegion id="E752F857FFB56A45FDBDFD53FD3BFD63" box="[544,653,693,714]" country="United States of America" name="Wisconsin" pageId="4" pageNumber="257">Wisconsin</collectingRegion>
were collected concurrently in June. All larvae were found feeding on small diameter branches of driftwood aspen. Larvae were not found within the wood, but in grooves open to the sandy substrate. Larvae were apparently mature upon collection and pupated within their feeding grooves. Adult emergence took place in August. Adults remained hidden during the day under debris well back from the water line. When disturbed they remained motionless and, due to their cryptic coloration, were difficult to locate.
</paragraph>
</subSubSection>
</treatment>
</document>