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<mods:title id="8FC483B05CEE0560FEC8CB148C794787">A new species of Guadua (Poaceae: Bambusoideae: Bambuseae: Guaduinae) from the Colombian Amazon</mods:title>
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<mods:namePart id="0425E3266C5EEF219FC8AA7B7E3D835C">Londoño, Ximena</mods:namePart>
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<mods:affiliation id="B3BA8B005D250007F912B4D2FA7C30CC">Sociedad Colombiana del Bambú, Quindió, Colombia.</mods:affiliation>
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<mods:namePart id="AE27489D4255F07B7C45D91B3961E288">Ruiz-Sanchez, Eduardo</mods:namePart>
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<mods:affiliation id="4C905C7668E7DCD51821D6BC1DBBAFEA">Departamento de Botánica y Zoología, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara, Camino Ing. Ramón Padilla Sánchez 2100, Nextipac, Zapopan, Jalisco, 45200, Mexico. &amp; Laboratorio Nacional de Identificación y Caracterización Vegetal (LaniVeg), Instituto de Botánica, Universidad de Guadalajara, Camino Ing. Ramón Padilla Sánchez 2100, Nextipac, Zapopan, Jalisco, 45200, Mexico.</mods:affiliation>
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<emphasis id="B8F6A36A9458FFA656BF7436FD56F981" bold="true" box="[136,727,1620,1647]" pageId="1" pageNumber="228">
<taxonomicName id="4D8204FB9458FFA656BF7436FE7BF981" authority="Londono" authorityName="Londono" box="[136,506,1620,1647]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="1" pageNumber="228" phylum="Tracheophyta" rank="species" species="bambuzoniae" status="sp. nov.">
<emphasis id="B8F6A36A9458FFA656BF7436FE07F980" bold="true" box="[136,390,1620,1647]" italics="true" pageId="1" pageNumber="228">Guadua bambuzoniae</emphasis>
Londoño
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,
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<emphasis id="B8F6A36A9458FFA654307436FDA1F981" bold="true" box="[519,544,1620,1646]" italics="true" pageId="1" pageNumber="228">sp</emphasis>
.
<emphasis id="B8F6A36A9458FFA654197436FDD9F981" bold="true" box="[558,600,1620,1646]" italics="true" pageId="1" pageNumber="228">nov</emphasis>
.
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<figureCitation id="12B963FD9458FFA654517436FD50F981" box="[614,721,1620,1646]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[136,229,1990,2012]" captionStartId-1="4.[136,229,1798,1820]" captionStartId-2="5.[136,229,1916,1938]" captionTargetBox-0="[177,1412,356,1957]" captionTargetBox-1="[184,1412,190,1774]" captionTargetBox-2="[164,1423,281,1892]" captionTargetId-0="figure-110@3.[171,1417,350,1963]" captionTargetId-1="figure-20@4.[164,1423,187,1776]" captionTargetId-2="figure-67@5.[164,1423,281,1892]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Guadua bambuzoniae. A. Foliage leaf complement. B. Detail of the apical region of the leaf sheath, adaxial view. C. Nodal region of the culm with solitary bud. D. Thorny branch segment. E. Culm leaves. F. Section of culm leaves upper portion showing blade and inner ligule. Based on: AB: Londoño &amp; S. Morales 2094; CF: Londoño 2032." captionText-1="FIGURE 2. Guadua bambuzoniae. A. Synflorescence terminating a leafy branch. B. Pseudospikelet. C. Basal gemmiparous bract. DE. Lemma, adaxial and abaxial view. F. Palea, close ventral view, G. Palea, abaxial view with winged keels. H. Androecium of 6 stamens surrounding the gynoecium. I. Stamen. J. Lodicules, K. Inmature gynoecium. Based on: Londoño &amp; S. Morales 2094." captionText-2="FIGURE 3. Guadua bambuzoniae. A. Habitat and habit. B. Pachymorph rhizome with the lowest culm internode. C. Culm leaf adaxial surfaces from two different portions of the culm. D. Node showing a single bud and supra- and infranodal white bands. E. Synflorescence showing 3 pseudospikelets. Pictures by Ximena Londoño." figureDoi-0="http://doi.org/10.5281/zenodo.13213004" figureDoi-1="http://doi.org/10.5281/zenodo.13213006" figureDoi-2="http://doi.org/10.5281/zenodo.13213008" httpUri-0="https://zenodo.org/record/13213004/files/figure.png" httpUri-1="https://zenodo.org/record/13213006/files/figure.png" httpUri-2="https://zenodo.org/record/13213008/files/figure.png" pageId="1" pageNumber="228">Figs. 13</figureCitation>
.
</emphasis>
</heading>
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:—
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.
<collectingRegion id="4846B19A9458FFA6575974C2FE3EF95A" box="[366,447,1696,1717]" country="Colombia" name="Caqueta" pageId="1" pageNumber="228">Caquetá</collectingRegion>
, Cartagena del Chaira, vereda Los Andes, Finca La Diosa del Bambú,
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,
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,
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,
<date id="FE3C59B89458FFA656EB74A6FE16F936" box="[220,407,1732,1754]" pageId="1" pageNumber="228" value="2022-09-29">29 September 2022</date>
(fl),
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X. Londoño &amp;
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. Morales 2094
</emphasis>
(
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!,
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<collectionCode id="EC93E7BD9458FFA655C674A6FBBCF936" box="[1009,1085,1732,1753]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="1" pageNumber="228" type="Herbarium">HUAZ</collectionCode>
!).
</paragraph>
<paragraph id="8A3D7F789458FFA656BF748AFD73F8A9" blockId="1.[136,1452,1696,1862]" pageId="1" pageNumber="228">
<taxonomicName id="4D8204FB9458FFA656BF748AFEDDF912" box="[136,348,1768,1789]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="1" pageNumber="228" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A9458FFA656BF748AFEDDF912" box="[136,348,1768,1789]" italics="true" pageId="1" pageNumber="228">Guadua bambuzoniae</emphasis>
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differs from
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<emphasis id="B8F6A36A9458FFA657E8748AFD10F912" box="[479,657,1768,1789]" italics="true" pageId="1" pageNumber="228">G. macrospiculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4D8204FB9458FFA654F5748AFCB4F912" authorityName="Londono &amp; Clark" authorityYear="2002" box="[706,821,1768,1789]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="1" pageNumber="228" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B8F6A36A9458FFA654F5748AFCB4F912" box="[706,821,1768,1789]" italics="true" pageId="1" pageNumber="228">G. uncinata</emphasis>
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in having culms erect at the base and arched from the mid-culm (
<emphasis id="B8F6A36A9458FFA656F2756EFE83F8CE" box="[197,258,1804,1825]" italics="true" pageId="1" pageNumber="228">versus</emphasis>
climbing or scandent) culm sheath with usually single falcate auricle (
<emphasis id="B8F6A36A9458FFA65595756EFC37F8CE" box="[930,950,1804,1825]" italics="true" pageId="1" pageNumber="228">vs</emphasis>
. two falcate auricles or auricles absent), lemma with densely ciliate margins (
<emphasis id="B8F6A36A9458FFA6579F7552FE3DF8AA" box="[424,444,1840,1861]" italics="true" pageId="1" pageNumber="228">vs</emphasis>
. lemmas with smooth margins).
</paragraph>
<paragraph id="8A3D7F789458FFA456BF7516FA14FEA6" blockId="1.[136,1451,1908,2079]" lastBlockId="3.[136,1452,158,329]" lastPageId="3" lastPageNumber="230" pageId="1" pageNumber="228">
Woody and thorny bamboo.
<emphasis id="B8F6A36A9458FFA657E07516FDCDF861" bold="true" box="[471,588,1908,1934]" pageId="1" pageNumber="228">Rhizomes</emphasis>
sympodial, pachymorph; the rhizome proper 912(18) ×
<quantity id="4D7AD29D9458FFA652C87516FADBF860" box="[1279,1370,1908,1935]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="1" pageNumber="228" unit="cm" value="8.0" valueMax="9.0" valueMin="7.0">79 cm</quantity>
; apical rhizome segments
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wide, basal rhizome segments
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wide; the rhizome neck relatively short,
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long; support rhizome necks that push down (or support rhizome necks like “feet”) 2 to 3 on each side of the rhizome,
<quantity id="4D7AD29D9458FFA656BF7582FF5AF814" box="[136,219,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="228" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">24 cm</quantity>
in diam.; adventitious roots emerging from rhizome segments
<quantity id="4D7AD29D9458FFA655AB7582FB8AF814" box="[924,1035,2016,2043]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="228" unit="cm" value="22.5" valueMax="25.0" valueMin="20.0">2025 cm</quantity>
long.
<emphasis id="B8F6A36A9458FFA652657582FB14F815" bold="true" box="[1106,1173,2016,2042]" pageId="1" pageNumber="228">Culm</emphasis>
(4)
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tall,
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diam., erect at the base, arched in the middle 1/3 and very arched in the apical 1/3, dull green in color, covered by whitish or straw-brown pubescence when young, cylindrical, tapered at apex; internodes (9)1524 (34) cm long, terete, the internodes of the lower 1/3 and the middle 1/3 hollow, those of the upper 1/3 solid, wall thickness in the 1/3 lower portion
<quantity id="4D7AD29D945BFFA557647284FE3FFEEE" box="[339,446,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="2" pageNumber="229" unit="cm" value="1.3" valueMax="1.6" valueMin="1.0">11.6 cm</quantity>
, wall thickness in the 1/3 middle portion
<quantity id="4D7AD29D945BFFA555A67284FB92FEEE" box="[913,1043,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="2" pageNumber="229" unit="cm" value="0.6" valueMax="0.8" valueMin="0.4">0.40.8 cm</quantity>
; nodes solitary, the single nodal line horizontal, with a band of white, appressed hairs above and below the nodal line, upper band
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wide, lower band
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wide, the supranodal ridge manifest when adult;
<emphasis id="B8F6A36A945BFFA5555A734CFC1DFEA7" bold="true" box="[877,924,302,328]" pageId="2" pageNumber="229">bud</emphasis>
single, triangular, ca. 1.8 ×
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, covered by a prophyllum; prophyll pubescent, ca.
<quantity id="4D7AD29D945BFFA5540A7330FD0AFE82" box="[573,651,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="229" unit="cm" value="1.6">1.6 cm</quantity>
long, covered by appressed brown hairs, the shoulders of the prophyll ciliate, cilia purple when young then brown.
<emphasis id="B8F6A36A945BFFA554417314FC87FE7F" bold="true" box="[630,774,374,400]" pageId="2" pageNumber="229">Culm leaves</emphasis>
<quantity id="4D7AD29D945BFFA5553B7314FCFBFE7E" box="[780,890,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" metricValueMax="4.6" metricValueMin="2.8" pageId="2" pageNumber="229" unit="cm" value="37.0" valueMax="46.0" valueMin="28.0">2846 cm</quantity>
long,
<quantity id="4D7AD29D945BFFA555897314FBADFE7E" box="[958,1068,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="2" pageNumber="229" unit="cm" value="14.0" valueMax="16.0" valueMin="12.0">1216 cm</quantity>
wide at base, generally surpassing the internode, coriaceous, deciduous, triangular-elongated;
<emphasis id="B8F6A36A945BFFA5551573F8FCF1FE5B" bold="true" box="[802,880,410,436]" pageId="2" pageNumber="229">sheath</emphasis>
2033 ×
<quantity id="4D7AD29D945BFFA555EA73F8FBCEFE5A" box="[989,1103,410,437]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="2" pageNumber="229" unit="cm" value="14.0" valueMax="16.0" valueMin="12.0">1216 cm</quantity>
, 2.5 to 2.8 times as long as the blade, abaxially pubescent, covered by
<specimenCount id="9C84B4F1945BFFA5547C73DDFD38FE36" box="[587,697,447,473]" count="2" pageId="2" pageNumber="229" type="generic" typeStatus="types">two types</specimenCount>
of hairs: a) hispid, rigid, sparse, antrorse, hyaline or brown hairs,
<quantity id="4D7AD29D945BFFA556BF7380FE92FE12" box="[136,275,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="2" pageNumber="229" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 mm</quantity>
long, removable, and b) short, appressed, fine, hyaline hairs, rough to the touch, adaxially glabrous, shiny, marginally ciliate, cilia hyaline or brown,
<quantity id="4D7AD29D945BFFA554807064FCADFDCE" box="[695,812,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="229" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">11.2 mm</quantity>
long, bearing a fimbriate auricle or not; auricle falcate, (0.4)0.71 ×
<quantity id="4D7AD29D945BFFA557107048FE27FDAA" box="[295,422,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="229" unit="cm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 cm</quantity>
, marginally fimbriate, the fimbriae ivory, shiny, wavy,
<quantity id="4D7AD29D945BFFA552307048FB05FDAA" box="[1031,1156,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="2" pageNumber="229" unit="cm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 cm</quantity>
long; inner ligule
<quantity id="4D7AD29D945BFFA553677048FA2DFDAA" box="[1360,1452,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="229" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, truncate, straight or slightly incurved at the middle or sigmoidal, usually ending before the margin, adaxially glabrous, shiny, abaxially puberulous, the margin finely ciliolate, cilia very short, hyaline, ca.
<quantity id="4D7AD29D945BFFA552A47010FB69FD62" box="[1171,1256,626,653]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="229" unit="mm" value="0.1">0.1 mm</quantity>
long;
<emphasis id="B8F6A36A945BFFA5531B7010FAECFD63" bold="true" box="[1324,1389,626,652]" pageId="2" pageNumber="229">blade</emphasis>
613 ×
<quantity id="4D7AD29D945BFFA5569670F4FF78FD5E" box="[161,249,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="229" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">57 cm</quantity>
, triangular, erect, persistent, abaxially glabrescent, covered by short, hyaline, sparse trichomes, adaxially conspicuously nerved and pubescent between nerves, covered by short brown hairs between nerves, densely pubescent at the lower portion, marginally ciliate, cilia
<quantity id="4D7AD29D945BFFA5544470BCFD62FD16" box="[627,739,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="2" pageNumber="229" unit="mm" value="1.75" valueMax="3.0" valueMin="0.5">0.53 mm</quantity>
long, brown, conspicuous towards the lower portion, mucronate at the apex, the mucro ca.
<quantity id="4D7AD29D945BFFA5579A7160FD9EFCF2" box="[429,543,770,797]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="229" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long, strong.
<emphasis id="B8F6A36A945BFFA5548E7160FCB6FCF3" bold="true" box="[697,823,770,796]" pageId="2" pageNumber="229">Branching</emphasis>
intravaginal, typically with one strong dominant branch that comes out ca. 45° angle, occasionally in the upper part developing 2 branches per node, the primary branch
<quantity id="4D7AD29D945BFFA5535B7144FF1FFC8A" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="229" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">11.5 m</quantity>
long, branching orders 2 to 3, developing thorny branches at almost all culm nodes, thorns straight to slightly curved, with (1)36 thorns/node.
<emphasis id="B8F6A36A945BFFA5543F710CFD2CFC67" bold="true" box="[520,685,878,904]" pageId="2" pageNumber="229">Foliage leaves</emphasis>
borne in lax complements of 718 leaves;
<emphasis id="B8F6A36A945BFFA552B3710CFB53FC67" bold="true" box="[1156,1234,878,904]" pageId="2" pageNumber="229">sheath</emphasis>
green when young, later stramineous, abaxially glabrous, ciliate on the margins, bearing fimbriae at the summit, cilia dark, purple to brown, fimbriae
<quantity id="4D7AD29D945BFFA5577171D4FE0EFC3F" box="[326,399,950,976]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="2.032" metricValueMin="1.016" pageId="2" pageNumber="229" unit="in" value="6.0" valueMax="8.0" valueMin="4.0">48 in</quantity>
number, deciduous, whitish,
<quantity id="4D7AD29D945BFFA554E971D4FCCAFC3E" box="[734,843,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="2" pageNumber="229" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">812 mm</quantity>
long, margins smooth;
<emphasis id="B8F6A36A945BFFA5526E71D4FB63FC3F" bold="true" box="[1113,1250,950,976]" pageId="2" pageNumber="229">inner ligule</emphasis>
ca.
<quantity id="4D7AD29D945BFFA5532471D4FAEBFC3E" box="[1299,1386,950,977]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="229" unit="mm" value="0.5">0.5 mm</quantity>
long, membranaceous, truncate, puberulous, stramineous to dark, margin glabrous;
<emphasis id="B8F6A36A945BFFA555DD71B8FBF0FC1B" bold="true" box="[1002,1137,986,1012]" pageId="2" pageNumber="229">outer ligule</emphasis>
<quantity id="4D7AD29D945BFFA5524F71B8FB7EFC1A" box="[1144,1279,986,1013]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="229" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.10.2 mm</quantity>
long, glabrous, shiny, marginally smooth;
<emphasis id="B8F6A36A945BFFA55798719CFDD3FBF7" bold="true" box="[431,594,1022,1048]" pageId="2" pageNumber="229">pseudopetiole</emphasis>
<quantity id="4D7AD29D945BFFA5546E719CFD34FBF6" box="[601,693,1022,1049]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="2" pageNumber="229" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">25 mm</quantity>
long, glabrous on both surfaces, usually pulvinate;
<emphasis id="B8F6A36A945BFFA552D8719CFAB1FBF7" bold="true" box="[1263,1328,1022,1048]" pageId="2" pageNumber="229">blade</emphasis>
(8)1318 (20) ×
<quantity id="4D7AD29D945BFFA556D47640FECEFBD2" box="[227,335,1058,1085]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="229" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 cm</quantity>
,
<collectionCode id="EC93E7BD945BFFA5576D7640FEEFFBD3" box="[346,366,1058,1084]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="2" pageNumber="229" type="Herbarium">L</collectionCode>
:
<collectionCode id="EC93E7BD945BFFA557457640FE13FBD3" box="[370,402,1058,1084]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="2" pageNumber="229" type="Herbarium">W</collectionCode>
=89:1, linear to linear-lanceolate, 1119 nerved, marginally scabrous on one side, scabrid on the other side, apically acuminate, adaxially glabrescent, with scattered, sparsely strigose, hyaline hairs ca.
<quantity id="4D7AD29D945BFFA553637624FA2DFB8E" box="[1364,1452,1094,1121]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="229" unit="mm" value="0.5">0.5 mm</quantity>
long, with 23 rows of prickle-hairs along one edge, abaxially pubescent, covered with scattered, sparse, hyaline or stramineous hairs ca.
<quantity id="4D7AD29D945BFFA5579376ECFE7CFB46" box="[420,509,1166,1193]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="229" unit="mm" value="0.5">0.5 mm</quantity>
long, tessellate, the midnerve prominent in the middle lower portion and slightly asymmetric.
<emphasis id="B8F6A36A945BFFA5572F76D0FE55FB23" bold="true" box="[280,468,1202,1228]" pageId="2" pageNumber="229">Pseudospikelets</emphasis>
(3)
<quantity id="4D7AD29D945BFFA5543D76D0FDDCFB22" box="[522,605,1202,1229]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="2" pageNumber="229" unit="cm" value="6.5" valueMax="8.0" valueMin="5.0">58 cm</quantity>
× (3)
<quantity id="4D7AD29D945BFFA5549076D0FC89FB22" box="[679,776,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="229" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
, linear-lanceolate, usually straight sometimes slighly curved dorsally, green with purple when young, later stramineous, consisting of a subtending bract, a prophyll, 24(6) basal gemmiparous bracts, 02 sterile lemmas and 610(12) fertile florets, and ending in a rudimentary floret; the terminal pseudospikelet lacking an immediately subtending prophyll;
<emphasis id="B8F6A36A945BFFA5551A777CFC08FAD7" bold="true" box="[813,905,1310,1336]" pageId="2" pageNumber="229">rachilla</emphasis>
segments 2.55 ×
<quantity id="4D7AD29D945BFFA55260777CFB18FAD6" box="[1111,1177,1310,1337]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="229" unit="mm" value="2.0">2 mm</quantity>
, pubescent in the middle upper portion, covered by short hyaline hairs, basally glabrous, usually longer between fertile florets, not disarticulating easily at the junction with the floret;
<emphasis id="B8F6A36A945BFFA5540F7704FC90FA6F" bold="true" box="[568,785,1382,1408]" pageId="2" pageNumber="229">subtending bracts</emphasis>
varying in size and shape throughout the main axis, with blades varying from fully developed and similar to small foliage leaf blade to reduced apicules, gemmiparous or not, the blades deciduous, the sheath abaxially pubescent towards the margins and glabrous on the dorsal part, ciliolate on the margins and bearing fimbriae or not at the summit;
<emphasis id="B8F6A36A945BFFA554C777B0FCD7FA03" bold="true" box="[752,854,1490,1516]" pageId="2" pageNumber="229">prophyll</emphasis>
13 ×
<quantity id="4D7AD29D945BFFA555AA77B0FBA7FA02" box="[925,1062,1490,1517]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.5" metricValueMin="1.1" pageId="2" pageNumber="229" unit="mm" value="1.8" valueMax="2.5" valueMin="1.1">1.12.5 mm</quantity>
, abaxially and adaxially pubescent in the middle upper portion, basally glabrous, with winged ciliolate keels;
<emphasis id="B8F6A36A945BFFA555E27794FA93F9FF" bold="true" box="[981,1298,1526,1552]" pageId="2" pageNumber="229">basal gemmiparous bracts</emphasis>
26, 45(8) × 34(5) mm wide, ovate to ovate-lanceolate, sometimes empty, the apex apiculate with a mucro
<quantity id="4D7AD29D945BFFA552C17478FAEBF9DA" box="[1270,1386,1562,1589]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="229" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long, 1217-nerved, abaxially pubescent towards the margins and otherwise glabrous, adaxially pubescent toward the apex, the rest glabrous, shiny and tessellate, the margins ciliolate;
<emphasis id="B8F6A36A945BFFA5550B7400FC6BF993" bold="true" box="[828,1002,1634,1660]" pageId="2" pageNumber="229">sterile lemmas</emphasis>
810 ×
<quantity id="4D7AD29D945BFFA5527B7400FB2CF992" box="[1100,1197,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="229" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
, rounded on the back, acuminate at the apex with a strong mucro
<quantity id="4D7AD29D945BFFA5544474E4FD67F94E" box="[627,742,1670,1697]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="229" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long, 1517-nerved, abaxially pubescent towards the margins, dorsally glabrous, adaxially pubescent towards the apex, otherwise glabrous and shiny, marginally densely ciliate, cilia brown in color, always enclosing a rudimentary palea.
<emphasis id="B8F6A36A945BFFA554CB74ACFCCEF907" bold="true" box="[764,847,1742,1768]" pageId="2" pageNumber="229">Florets</emphasis>
green and purplish in the middle basal portion, later stramineous, strongly attached at the base with a glabrous green band, ca.
<quantity id="4D7AD29D945BFFA5558E7490FC72F8E3" box="[953,1011,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="229" unit="mm" value="1.0">1mm</quantity>
long;
<emphasis id="B8F6A36A945BFFA5520F7490FB08F8E3" bold="true" box="[1080,1161,1778,1804]" pageId="2" pageNumber="229">lemma</emphasis>
1113 ×
<quantity id="4D7AD29D945BFFA552C77490FADAF8E2" box="[1264,1371,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="2" pageNumber="229" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
, ovate-lanceolate, rounded on the back, the apex acuminate with a strong mucro ca.
<quantity id="4D7AD29D945BFFA5523E7574FBCCF8DE" box="[1033,1101,1814,1841]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="229" unit="mm" value="1.0">1 mm</quantity>
long, 1821-nerved, abaxially pubescent towards the margins, otherwise glabrous, green with purple, later stramineous, adaxially pubescent on the apex, otherwise glabrous and shiny, the margins densely ciliate, cilia hyaline or brown, completely embracing the palea;
<emphasis id="B8F6A36A945BFFA556E675E0FE90F873" bold="true" box="[209,273,1922,1948]" pageId="2" pageNumber="229">palea</emphasis>
1012 ×
<quantity id="4D7AD29D945BFFA5574075E0FE54F872" box="[375,469,1922,1949]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="229" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
, stramineous, slightly shorter than the lemma, the keel wings laterally exceed the lemma, the sulcus between the keels
<quantity id="4D7AD29D945BFFA557EE75C4FDB8F82E" box="[473,569,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="229" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
wide, 58-nerved, with transverse veins, densely pubescent in the 3/4 upper portion, the ¼ lower portion glabrous, enfolding margins puberulous toward the 1/3 upper portion, otherwise glabrous, 57-nerved, the keels winged, ending at the apex, the wings
<quantity id="4D7AD29D945BFFA5551E758CFC1AF7E6" box="[809,923,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="229" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
wide, 13-nerved, adaxially puberulent toward the apex, abaxially glabrous and shiny, finely ciliolate on the margins towards the upper middle part, the cilia hyaline.
<emphasis id="B8F6A36A945AFFA456BF72FCFF7AFF57" bold="true" box="[136,251,158,184]" pageId="3" pageNumber="230">Lodicules</emphasis>
3, membranous, hyaline, many-nerved,
<quantity id="4D7AD29D945AFFA454F172FCFCBBFF56" box="[710,826,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="3" pageNumber="230" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.53 mm</quantity>
long,
<quantity id="4D7AD29D945AFFA455B472FCFC77FF56" box="[899,1014,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="230" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
wide, short-ciliolate on the upper part of the margin, the apex acute, the anterior pair a little longer and wider than the posterior one.
<emphasis id="B8F6A36A945AFFA4528E72A0FA9FFF33" bold="true" box="[1209,1310,194,220]" pageId="3" pageNumber="230">Stamens</emphasis>
6, filaments filiform, free;
<emphasis id="B8F6A36A945AFFA457187284FE08FEEF" bold="true" box="[303,393,230,256]" pageId="3" pageNumber="230">anthers</emphasis>
68 ×
<quantity id="4D7AD29D945AFFA457D27284FDDCFEEE" box="[485,605,230,257]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="230" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 mm</quantity>
, sagittate at the base, apiculate at the apex of each theca, ochre.
<emphasis id="B8F6A36A945AFFA453687284FA2AFEEF" bold="true" box="[1375,1451,230,256]" pageId="3" pageNumber="230">Ovary</emphasis>
(immature) 12 ×
<quantity id="4D7AD29D945AFFA4576F7368FE31FECA" box="[344,432,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="230" unit="mm" value="0.5">0.5 mm</quantity>
, fusiform, densely hispidulous, with a thickened, glabrous, basal support,
<quantity id="4D7AD29D945AFFA452C07368FAEBFECA" box="[1271,1386,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="3" pageNumber="230" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 mm</quantity>
long;
<emphasis id="B8F6A36A945AFFA456BF734CFF3FFEA7" bold="true" box="[136,190,302,328]" pageId="3" pageNumber="230">style</emphasis>
1, ca.
<quantity id="4D7AD29D945AFFA4573F734CFEC8FEA6" box="[264,329,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="230" unit="mm" value="1.0">1 mm</quantity>
long, densely antrorse-hispidulous with 3 plumose stigmas
<quantity id="4D7AD29D945AFFA455DF734CFBC4FEA6" box="[1000,1093,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="230" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long, hyaline.
<emphasis id="B8F6A36A945AFFA452DB734CFAABFEA7" bold="true" box="[1260,1322,302,328]" pageId="3" pageNumber="230">Fruit</emphasis>
not seen.
</paragraph>
</subSubSection>
<caption id="DEFD2FF0945AFFA456BF75A4FC2EF7CB" ID-DOI="http://doi.org/10.5281/zenodo.13213004" ID-Zenodo-Dep="13213004" httpUri="https://zenodo.org/record/13213004/files/figure.png" pageId="3" pageNumber="230" startId="3.[136,229,1990,2012]" targetBox="[177,1412,356,1957]" targetPageId="3" targetType="figure">
<paragraph id="8A3D7F78945AFFA456BF75A4FC2EF7CB" blockId="3.[136,1452,1990,2084]" pageId="3" pageNumber="230">
<emphasis id="B8F6A36A945AFFA456BF75A4FF7FF834" bold="true" box="[136,254,1990,2012]" pageId="3" pageNumber="230">FIGURE 1.</emphasis>
<taxonomicName id="4D8204FB945AFFA4573375A4FE59F834" box="[260,472,1990,2011]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="3" pageNumber="230" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945AFFA4573375A4FE59F834" box="[260,472,1990,2011]" italics="true" pageId="3" pageNumber="230">Guadua bambuzoniae</emphasis>
</taxonomicName>
. A. Foliage leaf complement. B. Detail of the apical region of the leaf sheath, adaxial view. C. Nodal region of the culm with solitary bud. D. Thorny branch segment. E. Culm leaves. F. Section of culm leaves upper portion showing blade and inner ligule. Based on: AB:
<emphasis id="B8F6A36A945AFFA457FE7A6CFD5AF7CC" box="[457,731,2062,2084]" italics="true" pageId="3" pageNumber="230">Londoño &amp; S. Morales 2094</emphasis>
; CF:
<emphasis id="B8F6A36A945AFFA4552A7A6CFC28F7CC" box="[797,937,2062,2083]" italics="true" pageId="3" pageNumber="230">Londoño 2032</emphasis>
.
</paragraph>
</caption>
<caption id="DEFD2FF0945DFFA356BF7564FB73F88C" ID-DOI="http://doi.org/10.5281/zenodo.13213006" ID-Zenodo-Dep="13213006" httpUri="https://zenodo.org/record/13213006/files/figure.png" pageId="4" pageNumber="231" startId="4.[136,229,1798,1820]" targetBox="[184,1412,190,1774]" targetPageId="4" targetType="figure">
<paragraph id="8A3D7F78945DFFA356BF7564FB73F88C" blockId="4.[136,1452,1798,1892]" pageId="4" pageNumber="231">
<emphasis id="B8F6A36A945DFFA356BF7564FF7FF8F4" bold="true" box="[136,254,1798,1820]" pageId="4" pageNumber="231">FIGURE 2.</emphasis>
<taxonomicName id="4D8204FB945DFFA357337564FE59F8F4" box="[260,472,1798,1819]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="4" pageNumber="231" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945DFFA357337564FE59F8F4" box="[260,472,1798,1819]" italics="true" pageId="4" pageNumber="231">Guadua bambuzoniae</emphasis>
</taxonomicName>
. A. Synflorescence terminating a leafy branch. B. Pseudospikelet. C. Basal gemmiparous bract. DE. Lemma, adaxial and abaxial view. F. Palea, close ventral view, G. Palea, abaxial view with winged keels. H. Androecium of 6 stamens surrounding the gynoecium. I. Stamen. J. Lodicules, K. Inmature gynoecium. Based on:
<emphasis id="B8F6A36A945DFFA355ED752CFB6DF88C" box="[986,1260,1870,1891]" italics="true" pageId="4" pageNumber="231">Londoño &amp; S. Morales 2094</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C2982CF3945DFFAF568A75F1FCD9FE13" lastPageId="8" lastPageNumber="235" pageId="4" pageNumber="231" type="etymology">
<paragraph id="8A3D7F78945DFFA3568A75F1FD6DF7F7" blockId="4.[136,1451,1938,2072]" pageId="4" pageNumber="231">
<emphasis id="B8F6A36A945DFFA3568A75F1FEE5F842" bold="true" box="[189,356,1939,1965]" pageId="4" pageNumber="231">Etymology:—</emphasis>
The specific epithet honors the Bambuzonía Project, funded by the International Bamboo and Rattan Organization—INBAR, and the International Fund for Agricultural Development—IFAD for the Amazon region of
<collectingCountry id="F2953FE8945DFFA356CC75B8FEEDF81B" box="[251,364,2010,2036]" name="Colombia" pageId="4" pageNumber="231">Colombia</collectingCountry>
,
<collectingCountry id="F2953FE8945DFFA3574C75B8FE5BF81B" box="[379,474,2010,2036]" name="Ecuador" pageId="4" pageNumber="231">Ecuador</collectingCountry>
, and
<collectingCountry id="F2953FE8945DFFA3542D75B9FDD0F81A" box="[538,593,2011,2037]" name="Peru" pageId="4" pageNumber="231">Peru</collectingCountry>
. The project was executed between 2020 and 2023 and contributed to the knowledge and development of bamboo in this region.
</paragraph>
<paragraph id="8A3D7F78945CFFA2568A72FFFB43FEEF" blockId="5.[136,1451,157,257]" pageId="5" pageNumber="232">
<emphasis id="B8F6A36A945CFFA2568A72FFFD87FF58" bold="true" box="[189,518,157,184]" pageId="5" pageNumber="232">Habitat and distribution:—</emphasis>
This species is distributed in the southeastern part of
<collectingCountry id="F2953FE8945CFFA2526772FCFB40FF57" box="[1104,1217,158,184]" name="Colombia" pageId="5" pageNumber="232">Colombia</collectingCountry>
, in the department of
<collectingRegion id="4846B19A945CFFA256BF72A0FF66FF33" box="[136,231,194,220]" country="Colombia" name="Caqueta" pageId="5" pageNumber="232">Caquetá</collectingRegion>
, between
<quantity id="4D7AD29D945CFFA2576572A0FE4EFF33" box="[338,463,194,221]" metricMagnitude="2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="5" pageNumber="232" unit="m" value="240.0" valueMax="250.0" valueMin="230.0">230250 m</quantity>
asl (
<figureCitation id="12B963FD945CFFA2543772A0FDC5FF33" box="[512,580,194,220]" captionStart="FIGURE 4" captionStartId="6.[136,229,1871,1893]" captionTargetBox="[151,1436,190,1842]" captionTargetId="figure-20@6.[151,1436,190,1847]" captionTargetPageId="6" captionText="FIGURE 4. Geographical distribution map of Guadua bambuzoniae. Country code: CO = Colombia, BR = Brazil, EC = Ecuador, PA = Panama, PE = Peru and VE = Venezuela." figureDoi="http://doi.org/10.5281/zenodo.13213010" httpUri="https://zenodo.org/record/13213010/files/figure.png" pageId="5" pageNumber="232">Fig. 4</figureCitation>
). It grows near streams and “cananguchales” (vegetation formations dominated by
<taxonomicName id="4D8204FB945CFFA2569C7285FDF6FEEE" authority="Linnaeus (1782: 454))" authorityName="Linnaeus" authorityPageNumber="454" authorityYear="1782" box="[171,631,230,257]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="232" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="B8F6A36A945CFFA2569C7285FEF2FEEF" box="[171,371,230,256]" italics="true" pageId="5" pageNumber="232">Mauritia flexuosa</emphasis>
Linnaeus (1782: 454))
</taxonomicName>
, in flooded areas, under shade or in full sun (
<figureCitation id="12B963FD945CFFA252477284FB35FEEF" box="[1136,1204,230,256]" captionStart="FIGURE 3" captionStartId="5.[136,229,1916,1938]" captionTargetBox="[164,1423,281,1892]" captionTargetId="figure-67@5.[164,1423,281,1892]" captionTargetPageId="5" captionText="FIGURE 3. Guadua bambuzoniae. A. Habitat and habit. B. Pachymorph rhizome with the lowest culm internode. C. Culm leaf adaxial surfaces from two different portions of the culm. D. Node showing a single bud and supra- and infranodal white bands. E. Synflorescence showing 3 pseudospikelets. Pictures by Ximena Londoño." figureDoi="http://doi.org/10.5281/zenodo.13213008" httpUri="https://zenodo.org/record/13213008/files/figure.png" pageId="5" pageNumber="232">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DEFD2FF0945CFFA256BF751EFD36F834" ID-DOI="http://doi.org/10.5281/zenodo.13213008" ID-Zenodo-Dep="13213008" httpUri="https://zenodo.org/record/13213008/files/figure.png" pageId="5" pageNumber="232" startId="5.[136,229,1916,1938]" targetBox="[164,1423,281,1892]" targetPageId="5" targetType="figure">
<paragraph id="8A3D7F78945CFFA256BF751EFD36F834" blockId="5.[136,1452,1916,2011]" pageId="5" pageNumber="232">
<emphasis id="B8F6A36A945CFFA256BF751EFF7FF87E" bold="true" box="[136,254,1916,1938]" pageId="5" pageNumber="232">FIGURE 3.</emphasis>
<taxonomicName id="4D8204FB945CFFA25732751FFE58F87D" box="[261,473,1917,1938]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="5" pageNumber="232" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945CFFA25732751FFE58F87D" box="[261,473,1917,1938]" italics="true" pageId="5" pageNumber="232">Guadua bambuzoniae</emphasis>
</taxonomicName>
. A. Habitat and habit. B. Pachymorph rhizome with the lowest culm internode. C. Culm leaf adaxial surfaces from two different portions of the culm. D. Node showing a single bud and supra- and infranodal white bands. E. Synflorescence showing 3 pseudospikelets. Pictures by Ximena Londoño.
</paragraph>
</caption>
<caption id="DEFD2FF0945FFFA156BF752DFD92F866" ID-DOI="http://doi.org/10.5281/zenodo.13213010" ID-Zenodo-Dep="13213010" httpUri="https://zenodo.org/record/13213010/files/figure.png" pageId="6" pageNumber="233" startId="6.[136,229,1871,1893]" targetBox="[151,1436,190,1842]" targetPageId="6" targetType="figure">
<paragraph id="8A3D7F78945FFFA156BF752DFD92F866" blockId="6.[136,1452,1871,1929]" pageId="6" pageNumber="233">
<emphasis id="B8F6A36A945FFFA156BF752DFF7FF88B" bold="true" box="[136,254,1871,1893]" pageId="6" pageNumber="233">FIGURE 4.</emphasis>
Geographical distribution map of
<taxonomicName id="4D8204FB945FFFA1547A752DFCA0F88B" box="[589,801,1871,1892]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="6" pageNumber="233" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945FFFA1547A752DFCA0F88B" box="[589,801,1871,1892]" italics="true" pageId="6" pageNumber="233">Guadua bambuzoniae</emphasis>
</taxonomicName>
. Country code: CO = Colombia, BR = Brazil, EC = Ecuador, PA = Panama, PE = Peru and VE = Venezuela.
</paragraph>
</caption>
<paragraph id="8A3D7F78945FFFA1568A75D5FE79F83D" blockId="6.[136,1452,1975,2074]" box="[189,504,1975,2002]" pageId="6" pageNumber="233">
<emphasis id="B8F6A36A945FFFA1568A75D5FE1BF83D" bold="true" box="[189,410,1975,2002]" pageId="6" pageNumber="233">Common name:—</emphasis>
<taxonomicName id="4D8204FB945FFFA157AE75DAFE75F83D" authorityName="Kunth" authorityYear="1822" box="[409,500,1976,2002]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="6" pageNumber="233" phylum="Tracheophyta" rank="genus">Guadua</taxonomicName>
.
</paragraph>
<paragraph id="8A3D7F78945FFFA0568A75BEFF5DFF33" blockId="6.[136,1452,1975,2074]" lastBlockId="7.[136,1452,158,653]" lastPageId="7" lastPageNumber="234" pageId="6" pageNumber="233">
<emphasis id="B8F6A36A945FFFA1568A75BEFE9BF819" bold="true" box="[189,282,2012,2038]" pageId="6" pageNumber="233">Uses:—</emphasis>
The small straight and tubular portion of the culm and the excess of branches with spines at almost all nodes make
<taxonomicName id="4D8204FB945FFFA157237A62FD8BF7F5" box="[276,522,2048,2074]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="6" pageNumber="233" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945FFFA157237A62FD8BF7F5" box="[276,522,2048,2074]" italics="true" pageId="6" pageNumber="233">Guadua bambuzoniae</emphasis>
</taxonomicName>
have little applicability. However, poles can be extracted from the first 2 or 3 basal meters to make fences, to poke or harass cattle, to make furniture, or to replace the decorative poles in the porch of the houses.
</paragraph>
<paragraph id="8A3D7F78945EFFA0568A7284FA06FECB" blockId="7.[136,1452,158,653]" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA0568A7284FEDEFEEF" bold="true" box="[189,351,230,256]" pageId="7" pageNumber="234">Phenology:—</emphasis>
<taxonomicName id="4D8204FB945EFFA057687284FDD2FEEF" box="[351,595,230,256]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945EFFA057687284FDD2FEEF" box="[351,595,230,256]" italics="true" pageId="7" pageNumber="234">Guadua bambuzoniae</emphasis>
</taxonomicName>
is able to flowering during a 2 or 3 year period without a dying, according with local people. Flowering culms retain foliage leaves and the fruit production is scarce (Londoño pers. observation).
</paragraph>
<paragraph id="8A3D7F78945EFFA0568A734CFBEAFE7F" blockId="7.[136,1452,158,653]" pageId="7" pageNumber="234">
<materialsCitation id="3AEA7525945EFFA0568A734CFB66FE83" collectingDate="2020-10-11" collectionCode="HUAZ" collectorName="X. Londono" country="Colombia" elevation="230" latitude="1.3125" location="Finca Camberos" longLatPrecision="21" longitude="-74.849724" municipality="Cartagena del Chaira" pageId="7" pageNumber="234" specimenCount="2" stateProvince="Caqueta">
<emphasis id="B8F6A36A945EFFA0568A734CFD66FEA7" bold="true" box="[189,743,302,328]" pageId="7" pageNumber="234">
<typeStatus id="5539C1DA945EFFA0568A734CFDB5FEA7" box="[189,564,302,328]" isOtherTypeStatus="true" pageId="7" pageNumber="234">Additional specimens examined</typeStatus>
(
<typeStatus id="5539C1DA945EFFA05476734CFD3CFEA7" box="[577,701,302,328]" pageId="7" pageNumber="234" type="paratype">paratypes</typeStatus>
):—
</emphasis>
<collectingCountry id="F2953FE8945EFFA054D0734CFC01FEA7" box="[743,896,302,328]" name="Colombia" pageId="7" pageNumber="234">COLOMBIA</collectingCountry>
.
<collectingRegion id="4846B19A945EFFA055BE734CFC69FEA7" box="[905,1000,302,328]" country="Colombia" name="Caqueta" pageId="7" pageNumber="234">Caquetá</collectingRegion>
,
<collectingMunicipality id="6A59E502945EFFA055C6734CFB5CFEA7" box="[1009,1245,302,328]" pageId="7" pageNumber="234">Cartagena del Chaira</collectingMunicipality>
, Vda. Guadualosa,
<location id="8F5D29A3945EFFA056BF7330FEC1FE83" LSID="urn:lsid:plazi:treatment:022BCE6E9458FFAF56BF7436FCD9FE13:8F5D29A3945EFFA056BF7330FEC1FE83" box="[136,320,338,364]" country="Colombia" latitude="1.3125" longLatPrecision="21" longitude="-74.849724" municipality="Cartagena del Chaira" name="Finca Camberos" pageId="7" pageNumber="234" stateProvince="Caqueta">Finca Camberos</location>
,
<geoCoordinate id="EFB619BF945EFFA0577D7330FE46FE82" box="[330,455,338,365]" degrees="1" direction="north" minutes="18" orientation="latitude" pageId="7" pageNumber="234" precision="15" seconds="45" value="1.3125">1°1845”N</geoCoordinate>
,
<geoCoordinate id="EFB619BF945EFFA057E57330FDE6FE82" box="[466,615,338,365]" degrees="74" direction="west" minutes="50" orientation="longitude" pageId="7" pageNumber="234" precision="15" seconds="59" value="-74.849724">74°5059” W</geoCoordinate>
,
<elevation id="01AF984B945EFFA054467330FD72FE83" box="[625,755,338,365]" metricMagnitude="2" metricUnit="m" metricValue="2.303" pageId="7" pageNumber="234" unit="m" value="230.3">
<quantity id="4D7AD29D945EFFA054467330FD4AFE82" box="[625,715,338,365]" metricMagnitude="2" metricUnit="m" metricValue="2.303" pageId="7" pageNumber="234" unit="m" value="230.3">230.3 m</quantity>
asl
</elevation>
,
<date id="FE3C59B8945EFFA054CA7330FC39FE83" box="[765,952,338,364]" pageId="7" pageNumber="234" value="2020-10-11">
<collectingDate id="EE78A050945EFFA054CA7330FC39FE83" box="[765,952,338,364]" pageId="7" pageNumber="234" value="2020-10-11">11 October 2020</collectingDate>
</date>
,
<emphasis id="B8F6A36A945EFFA055F57331FB00FE83" box="[962,1153,338,364]" italics="true" pageId="7" pageNumber="234">
<collectorName id="27771AAE945EFFA055F57331FBC5FE83" box="[962,1092,338,364]" pageId="7" pageNumber="234">X. Londoño</collectorName>
2032
</emphasis>
(
<collectionCode id="EC93E7BD945EFFA052B97330FB5EFE83" box="[1166,1247,338,364]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="7" pageNumber="234" type="Herbarium">HUAZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3AEA7525945EFFA052C17330FBE6FE7F" collectingDate="2022-02-28" collectionCode="S, HUAZ" collectorName="X. Londono &amp; Morales" country="Colombia" latitude="1.3125" location="Finca Camberos" longLatPrecision="15" longitude="-74.84978" pageId="7" pageNumber="234" specimenCount="1">
<location id="8F5D29A3945EFFA052C17330FA2DFE83" LSID="urn:lsid:plazi:treatment:022BCE6E9458FFAF56BF7436FCD9FE13:8F5D29A3945EFFA052C17330FA2DFE83" box="[1270,1452,338,364]" country="Colombia" latitude="1.3125" longLatPrecision="15" longitude="-74.84978" name="Finca Camberos" pageId="7" pageNumber="234">Finca Camberos</location>
<geoCoordinate id="EFB619BF945EFFA056BF7314FE8CFE7E" box="[136,269,374,401]" degrees="1" direction="north" minutes="18" orientation="latitude" pageId="7" pageNumber="234" precision="15" seconds="45" value="1.3125">1°1845” N</geoCoordinate>
,
<geoCoordinate id="EFB619BF945EFFA0572F7314FE3CFE7E" box="[280,445,374,401]" degrees="74" direction="west" minutes="50" orientation="longitude" pageId="7" pageNumber="234" precision="1" seconds="59.2" value="-74.84978">74°5059.2”W</geoCoordinate>
,
<date id="FE3C59B8945EFFA057FF7314FD13FE7F" box="[456,658,374,400]" pageId="7" pageNumber="234" value="2022-02-28">
<collectingDate id="EE78A050945EFFA057FF7314FD13FE7F" box="[456,658,374,400]" pageId="7" pageNumber="234" value="2022-02-28">28 February 2022</collectingDate>
</date>
,
<emphasis id="B8F6A36A945EFFA054AA7315FC7DFE7F" box="[669,1020,374,401]" italics="true" pageId="7" pageNumber="234">
<collectorName id="27771AAE945EFFA054AA7315FCA1FE7F" box="[669,800,374,400]" pageId="7" pageNumber="234">X. Londoño</collectorName>
&amp;
<collectionCode id="EC93E7BD945EFFA055737315FCD4FE7E" box="[836,853,375,401]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="7" pageNumber="234" type="Herbarium">S</collectionCode>
.
<collectorName id="27771AAE945EFFA055577314FC3CFE7F" box="[864,957,374,400]" pageId="7" pageNumber="234">Morales</collectorName>
2093
</emphasis>
(
<collectionCode id="EC93E7BD945EFFA0523C7314FBDEFE7F" box="[1035,1119,374,400]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="7" pageNumber="234" type="Herbarium">HUAZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8A3D7F78945EFFA0568A73F8FB06FD63" blockId="7.[136,1452,158,653]" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA0568A73F8FEA7FE5B" bold="true" box="[189,294,410,436]" pageId="7" pageNumber="234">Notes:—</emphasis>
<taxonomicName id="4D8204FB945EFFA0571173F8FD9DFE5B" box="[294,540,410,436]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945EFFA0571173F8FD9DFE5B" box="[294,540,410,436]" italics="true" pageId="7" pageNumber="234">Guadua bambuzoniae</emphasis>
</taxonomicName>
inhabits swampy habitats, as do
<taxonomicName id="4D8204FB945EFFA055B973F9FBDCFE5B" authorityName="Londono &amp; Clark" authorityYear="2002" box="[910,1117,410,437]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="macrospiculata">
<emphasis id="B8F6A36A945EFFA055B973F9FBDCFE5B" box="[910,1117,410,437]" italics="true" pageId="7" pageNumber="234">G. macrospiculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4D8204FB945EFFA052A373F8FA99FE5B" authorityName="Londono &amp; Clark" authorityYear="2002" box="[1172,1304,410,436]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="uncinata">
G.
<emphasis id="B8F6A36A945EFFA0528173F9FA99FE5B" box="[1206,1304,411,436]" italics="true" pageId="7" pageNumber="234">uncinata</emphasis>
</taxonomicName>
, species also found in this region of
<collectingCountry id="F2953FE8945EFFA057BA73DCFE7EFE37" box="[397,511,446,472]" name="Colombia" pageId="7" pageNumber="234">Colombia</collectingCountry>
. With these two species,
<taxonomicName id="4D8204FB945EFFA0552D73DDFC53FE37" box="[794,978,446,473]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945EFFA0552D73DDFC53FE37" box="[794,978,446,473]" italics="true" pageId="7" pageNumber="234">G. bambuzoniae</emphasis>
</taxonomicName>
shares green spikelets with purple lemma when young, a pubescent ovary and style densely covered with hispid and antrorse hairs, and the presence of 3 hyaline plumose stigmas. However,
<taxonomicName id="4D8204FB945EFFA057FD7065FD02FDCF" box="[458,643,518,545]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945EFFA057FD7065FD02FDCF" box="[458,643,518,545]" italics="true" pageId="7" pageNumber="234">G. bambuzoniae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4D8204FB945EFFA0552B7065FC6CFDCF" authorityName="Londono &amp; Clark" authorityYear="2002" box="[796,1005,518,545]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="macrospiculata">
<emphasis id="B8F6A36A945EFFA0552B7065FC6CFDCF" box="[796,1005,518,545]" italics="true" pageId="7" pageNumber="234">G. macrospiculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4D8204FB945EFFA052107065FB2CFDCF" authorityName="Londono &amp; Clark" authorityYear="2002" box="[1063,1197,519,545]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B8F6A36A945EFFA052107065FB2CFDCF" box="[1063,1197,519,545]" italics="true" pageId="7" pageNumber="234">G. uncinata</emphasis>
</taxonomicName>
because: a) the culms are erect at the base and arched from the mid-culm (
<emphasis id="B8F6A36A945EFFA054E27049FC9CFDAB" box="[725,797,555,580]" italics="true" pageId="7" pageNumber="234">versus</emphasis>
climbing or scandent); b) the blade of the culm leaf is not swollen (
<emphasis id="B8F6A36A945EFFA056C6702DFE89FD87" box="[241,264,591,616]" italics="true" pageId="7" pageNumber="234">vs</emphasis>
. swollen); c) the culm leaf sheath usually has a falcate and fimbriate auricle (
<emphasis id="B8F6A36A945EFFA052B0702DFB24FD87" box="[1159,1189,591,616]" italics="true" pageId="7" pageNumber="234">vs.</emphasis>
two falcate auricles or auricles absent); and d) the margins of the lemma are densely ciliate (
<emphasis id="B8F6A36A945EFFA055A47011FC2BFD63" box="[915,938,627,652]" italics="true" pageId="7" pageNumber="234">vs</emphasis>
. smooth) (
<tableCitation id="C7004AC3945EFFA052157010FBF7FD63" box="[1058,1142,626,652]" captionStart="TABLE 1" captionStartId="7.[136,229,698,724]" captionText="TABLE 1. Morphological comparison of Guadua bambuzoniae with morphologically related species. Characters of G. macrospiculata and G. uncinata were based on Londoño &amp; L.G. Clark (2002)." pageId="7" pageNumber="234">Table 1</tableCitation>
).
</paragraph>
<caption id="DEFD2FF0945EFFA056BF70D8FB93FD17" pageId="7" pageNumber="234" startId="7.[136,229,698,724]" targetType="table">
<paragraph id="8A3D7F78945EFFA056BF70D8FB93FD17" blockId="7.[136,1451,698,761]" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056BF70D8FF7AFD3B" bold="true" box="[136,251,698,724]" pageId="7" pageNumber="234">TABLE 1</emphasis>
. Morphological comparison of
<taxonomicName id="4D8204FB945EFFA0545370D8FCDAFD3B" box="[612,859,698,724]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945EFFA0545370D8FCDAFD3B" box="[612,859,698,724]" italics="true" pageId="7" pageNumber="234">Guadua bambuzoniae</emphasis>
</taxonomicName>
with morphologically related species. Characters of
<taxonomicName id="4D8204FB945EFFA056BF70BDFED6FD17" authorityName="Londono &amp; Clark" authorityYear="2002" box="[136,343,734,761]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="macrospiculata">
<emphasis id="B8F6A36A945EFFA056BF70BDFED6FD17" box="[136,343,734,761]" italics="true" pageId="7" pageNumber="234">G. macrospiculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4D8204FB945EFFA057B970BDFD93FD17" authorityName="Londono &amp; Clark" authorityYear="2002" box="[398,530,735,761]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B8F6A36A945EFFA057B970BDFD93FD17" box="[398,530,735,761]" italics="true" pageId="7" pageNumber="234">G. uncinata</emphasis>
</taxonomicName>
were based on Londoño &amp; L.G. Clark (2002).
</paragraph>
</caption>
<paragraph id="8A3D7F78945EFFA056A77168FA25F803" pageId="7" pageNumber="234">
<table id="F8828DD8945E005856A77168FA25F811" box="[144,1444,778,2046]" gridcols="4" gridrows="16" pageId="7" pageNumber="234">
<tr id="34B27D3A945E005856A77168FA25FCF2" box="[144,1444,778,797]" gridrow="0" pageId="7" pageNumber="234">
<th id="77631446945E005856A77168FE54FCF2" box="[144,469,778,797]" gridcol="0" gridrow="0" pageId="7" pageNumber="234">Character</th>
<th id="77631446945E005857C57168FC90FCF2" box="[498,785,778,797]" gridcol="1" gridrow="0" pageId="7" pageNumber="234">
<taxonomicName id="4D8204FB945EFFA057C57168FD00FCF2" box="[498,641,778,797]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A945EFFA057C57168FD00FCF2" bold="true" box="[498,641,778,797]" italics="true" pageId="7" pageNumber="234">G. bambuzoniae</emphasis>
</taxonomicName>
</th>
<th id="77631446945E0058551C7168FBFEFCF2" box="[811,1151,778,797]" gridcol="2" gridrow="0" pageId="7" pageNumber="234">
<taxonomicName id="4D8204FB945EFFA0551C7168FC4DFCF2" authorityName="Londono &amp; Clark" authorityYear="2002" box="[811,972,778,797]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="macrospiculata">
<emphasis id="B8F6A36A945EFFA0551C7168FC4DFCF2" bold="true" box="[811,972,778,797]" italics="true" pageId="7" pageNumber="234">G. macrospiculata</emphasis>
</taxonomicName>
</th>
<th id="77631446945E005852B97168FA25FCF2" box="[1166,1444,778,797]" gridcol="3" gridrow="0" pageId="7" pageNumber="234">
<taxonomicName id="4D8204FB945EFFA052B97168FB76FCF2" authority="Habit" authorityName="Habit" box="[1166,1271,778,797]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="7" pageNumber="234" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B8F6A36A945EFFA052B97168FB76FCF2" bold="true" box="[1166,1271,778,797]" italics="true" pageId="7" pageNumber="234">G. uncinata</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="34B27D3A945E005856A77151FA25FC85" box="[144,1444,819,874]" gridrow="1" pageId="7" pageNumber="234">
<th id="77631446945E005856A77151FE54FC85" box="[144,469,819,874]" gridcol="0" gridrow="1" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A77127FF45FCB7" bold="true" box="[144,196,837,856]" pageId="7" pageNumber="234">Habit</emphasis>
</th>
<td id="77631446945E005857C57151FC90FC85" box="[498,785,819,874]" gridcol="1" gridrow="1" pageId="7" pageNumber="234">Erect at the base, arched at mid- culm</td>
<td id="77631446945E0058551C7151FBFEFC85" box="[811,1151,819,874]" gridcol="2" gridrow="1" pageId="7" pageNumber="234">Climing or scandent</td>
<td id="77631446945E005852B97151FA25FC85" box="[1166,1444,819,874]" gridcol="3" gridrow="1" pageId="7" pageNumber="234">Erect at the base, scandent at mid-culm</td>
</tr>
<tr id="34B27D3A945E005856A771F3FA25FC4A" box="[144,1444,913,933]" gridrow="2" pageId="7" pageNumber="234">
<th id="77631446945E005856A771F3FE54FC4A" box="[144,469,913,933]" gridcol="0" gridrow="2" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A771F3FE4DFC4B" bold="true" box="[144,460,913,933]" pageId="7" pageNumber="234">Culms: lenght (m) × diameter (cm)</emphasis>
</th>
<td id="77631446945E005857C571F3FC90FC4A" box="[498,785,913,933]" gridcol="1" gridrow="2" pageId="7" pageNumber="234">(4)810 × 14</td>
<td id="77631446945E0058551C71F3FBFEFC4A" box="[811,1151,913,933]" gridcol="2" gridrow="2" pageId="7" pageNumber="234">815 × 1.84</td>
<td id="77631446945E005852B971F3FA25FC4A" box="[1166,1444,913,933]" gridcol="3" gridrow="2" pageId="7" pageNumber="234">38 × 24(6)</td>
</tr>
<tr id="34B27D3A945E005856A771AEFA25FBEC" box="[144,1444,972,1027]" gridrow="3" pageId="7" pageNumber="234">
<th id="77631446945E005856A771AEFE54FBEC" box="[144,469,972,1027]" gridcol="0" gridrow="3" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A771BCFEFBFC1E" bold="true" box="[144,378,990,1009]" pageId="7" pageNumber="234">Culm internode condition</emphasis>
</th>
<td id="77631446945E005857C571AEFC90FBEC" box="[498,785,972,1027]" gridcol="1" gridrow="3" pageId="7" pageNumber="234">Basal internodes hollow, mid- culm internodes solid</td>
<td id="77631446945E0058551C71AEFBFEFBEC" box="[811,1151,972,1027]" gridcol="2" gridrow="3" pageId="7" pageNumber="234">Solid</td>
<td id="77631446945E005852B971AEFA25FBEC" box="[1166,1444,972,1027]" gridcol="3" gridrow="3" pageId="7" pageNumber="234">Basal internodes solid, mid- culm internodes hollow</td>
</tr>
<tr id="34B27D3A945E005856A77678FA25FBB8" box="[144,1444,1050,1111]" gridrow="4" pageId="7" pageNumber="234">
<th id="77631446945E005856A77678FE54FBB8" box="[144,469,1050,1111]" gridcol="0" gridrow="4" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A77678FED0FBB8" bold="true" pageId="7" pageNumber="234">Culm internode indument Thorns number/node</emphasis>
</th>
<td id="77631446945E005857C57678FC90FBB8" box="[498,785,1050,1111]" gridcol="1" gridrow="4" pageId="7" pageNumber="234">Pubescent 16</td>
<td id="77631446945E0058551C7678FBFEFBB8" box="[811,1151,1050,1111]" gridcol="2" gridrow="4" pageId="7" pageNumber="234">Slightly estrigose 13</td>
<td id="77631446945E005852B97678FA25FBB8" box="[1166,1444,1050,1111]" gridcol="3" gridrow="4" pageId="7" pageNumber="234">Densely pubescent 15</td>
</tr>
<tr id="34B27D3A945E005856A7761CFA25FB7D" box="[144,1444,1150,1170]" gridrow="5" pageId="7" pageNumber="234">
<th id="77631446945E005856A7761CFE54FB7D" box="[144,469,1150,1170]" gridcol="0" gridrow="5" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A7761CFE4EFB7E" bold="true" box="[144,463,1150,1170]" pageId="7" pageNumber="234">Culm leaf sheath abaxial indument</emphasis>
</th>
<td id="77631446945E005857C5761CFC90FB7D" box="[498,785,1150,1170]" gridcol="1" gridrow="5" pageId="7" pageNumber="234">Estrigose</td>
<td id="77631446945E0058551C761CFBFEFB7D" box="[811,1151,1150,1170]" gridcol="2" gridrow="5" pageId="7" pageNumber="234">Scabrid</td>
<td id="77631446945E005852B9761CFA25FB7D" box="[1166,1444,1150,1170]" gridcol="3" gridrow="5" pageId="7" pageNumber="234">Densely pubescent</td>
</tr>
<tr id="34B27D3A945E005856A776DBFA25FAF5" box="[144,1444,1209,1306]" gridrow="6" pageId="7" pageNumber="234">
<th id="77631446945E005856A776DBFE54FAF5" box="[144,469,1209,1306]" gridcol="0" gridrow="6" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A776DBFE39FAF5" bold="true" pageId="7" pageNumber="234">
Culm leaf auricles,
<taxonomicName id="4D8204FB945EFFA0577776DBFE14FB23" box="[320,405,1209,1228]" form="and" pageId="7" pageNumber="234" rank="form">form and</taxonomicName>
size (mm length × mm wide) Foliage sheath abaxila indument
</emphasis>
</th>
<td id="77631446945E005857C576DBFC90FAF5" box="[498,785,1209,1306]" gridcol="1" gridrow="6" pageId="7" pageNumber="234">Usually 1 auricle present, falcate, 410 × 23 Glabrous</td>
<td id="77631446945E0058551C76DBFBFEFAF5" box="[811,1151,1209,1306]" gridcol="2" gridrow="6" pageId="7" pageNumber="234">Auricles absent or 2 present, falcate, 13 × 11.2 Densely pubescent</td>
<td id="77631446945E005852B976DBFA25FAF5" box="[1166,1444,1209,1306]" gridcol="3" gridrow="6" pageId="7" pageNumber="234">2 auricles present, falcate, 46 × 11.2 Usually glabrous</td>
</tr>
<tr id="34B27D3A945E005856A77753FA25FA63" box="[144,1444,1329,1420]" gridrow="7" pageId="7" pageNumber="234">
<th id="77631446945E005856A77753FE54FA63" box="[144,469,1329,1420]" gridcol="0" gridrow="7" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A77721FF69FA95" bold="true" pageId="7" pageNumber="234">Pseudopetiole length (mm) and indument</emphasis>
</th>
<td id="77631446945E005857C57753FC90FA63" box="[498,785,1329,1420]" gridcol="1" gridrow="7" pageId="7" pageNumber="234">25; glabrous on both surfaces</td>
<td id="77631446945E0058551C7753FBFEFA63" box="[811,1151,1329,1420]" gridcol="2" gridrow="7" pageId="7" pageNumber="234">11.5; adaxially hispidulous, abaxially hirsute to glabrescent</td>
<td id="77631446945E005852B97753FA25FA63" box="[1166,1444,1329,1420]" gridcol="3" gridrow="7" pageId="7" pageNumber="234">36; adaxially glabrous and puberulous at the base, abaxially glabrous</td>
</tr>
<tr id="34B27D3A945E005856A777C1FA25FA35" box="[144,1444,1443,1498]" gridrow="8" pageId="7" pageNumber="234">
<th id="77631446945E005856A777C1FE54FA35" box="[144,469,1443,1498]" gridcol="0" gridrow="8" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A777C1FF73FA35" bold="true" pageId="7" pageNumber="234">Foliage leave blade: length (cm) × width (cm)</emphasis>
</th>
<td id="77631446945E005857C577C1FC90FA35" box="[498,785,1443,1498]" gridcol="1" gridrow="8" pageId="7" pageNumber="234">820 × 11.5</td>
<td id="77631446945E0058551C77C1FBFEFA35" box="[811,1151,1443,1498]" gridcol="2" gridrow="8" pageId="7" pageNumber="234">212 × 0.82.8</td>
<td id="77631446945E005852B977C1FA25FA35" box="[1166,1444,1443,1498]" gridcol="3" gridrow="8" pageId="7" pageNumber="234">6.518 × 1.44.2</td>
</tr>
<tr id="34B27D3A945E005856A77793FA25F9C7" box="[144,1444,1521,1576]" gridrow="9" pageId="7" pageNumber="234">
<th id="77631446945E005856A77793FE54F9C7" box="[144,469,1521,1576]" gridcol="0" gridrow="9" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A77461FE0CF9F9" bold="true" box="[144,397,1539,1558]" pageId="7" pageNumber="234">Foliage leaf blade indument</emphasis>
</th>
<td id="77631446945E005857C57793FC90F9C7" box="[498,785,1521,1576]" gridcol="1" gridrow="9" pageId="7" pageNumber="234">Adaxially glabrescent; abaxially pubescent</td>
<td id="77631446945E0058551C7793FBFEF9C7" box="[811,1151,1521,1576]" gridcol="2" gridrow="9" pageId="7" pageNumber="234">Adaxially scabrid to glabrous; abaxially pilose to glabrescent</td>
<td id="77631446945E005852B97793FA25F9C7" box="[1166,1444,1521,1576]" gridcol="3" gridrow="9" pageId="7" pageNumber="234">Adaxially glabrous; abaxially pubescent</td>
</tr>
<tr id="34B27D3A945E005856A7745CFA25F999" box="[144,1444,1598,1654]" gridrow="10" pageId="7" pageNumber="234">
<th id="77631446945E005856A7745CFE54F999" box="[144,469,1598,1654]" gridcol="0" gridrow="10" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A7745CFF59F999" bold="true" pageId="7" pageNumber="234">Pseudospikelet/ coflorescence number</emphasis>
</th>
<td id="77631446945E005857C5745CFC90F999" box="[498,785,1598,1654]" gridcol="1" gridrow="10" pageId="7" pageNumber="234">13</td>
<td id="77631446945E0058551C745CFBFEF999" box="[811,1151,1598,1654]" gridcol="2" gridrow="10" pageId="7" pageNumber="234">usually 13</td>
<td id="77631446945E005852B9745CFA25F999" box="[1166,1444,1598,1654]" gridcol="3" gridrow="10" pageId="7" pageNumber="234">12(15)</td>
</tr>
<tr id="34B27D3A945E005856A774EEFA25F92C" box="[144,1444,1676,1731]" gridrow="11" pageId="7" pageNumber="234">
<th id="77631446945E005856A774EEFE54F92C" box="[144,469,1676,1731]" gridcol="0" gridrow="11" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A774EEFF73F92C" bold="true" pageId="7" pageNumber="234">Pseudospikelet size: length (cm) × width (cm)</emphasis>
</th>
<td id="77631446945E005857C574EEFC90F92C" box="[498,785,1676,1731]" gridcol="1" gridrow="11" pageId="7" pageNumber="234">37 × 0.30.5</td>
<td id="77631446945E0058551C74EEFBFEF92C" box="[811,1151,1676,1731]" gridcol="2" gridrow="11" pageId="7" pageNumber="234">317 × 0.30.5</td>
<td id="77631446945E005852B974EEFA25F92C" box="[1166,1444,1676,1731]" gridcol="3" gridrow="11" pageId="7" pageNumber="234">38 × 0.30.6</td>
</tr>
<tr id="34B27D3A945E005856A77488FA25F911" box="[144,1444,1770,1790]" gridrow="12" pageId="7" pageNumber="234">
<th id="77631446945E005856A77488FE54F911" box="[144,469,1770,1790]" gridcol="0" gridrow="12" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A77488FE51F912" bold="true" box="[144,464,1770,1789]" pageId="7" pageNumber="234">Synflorescence main axis indument</emphasis>
</th>
<td id="77631446945E005857C57488FC90F911" box="[498,785,1770,1790]" gridcol="1" gridrow="12" pageId="7" pageNumber="234">Pubescent</td>
<td id="77631446945E0058551C7488FBFEF911" box="[811,1151,1770,1790]" gridcol="2" gridrow="12" pageId="7" pageNumber="234">Pubescent</td>
<td id="77631446945E005852B97488FA25F911" box="[1166,1444,1770,1790]" gridcol="3" gridrow="12" pageId="7" pageNumber="234">Hispidulos or glabrous</td>
</tr>
<tr id="34B27D3A945E005856A77547FA25F8B2" box="[144,1444,1829,1885]" gridrow="13" pageId="7" pageNumber="234">
<th id="77631446945E005856A77547FE54F8B2" box="[144,469,1829,1885]" gridcol="0" gridrow="13" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A77555FEFBF8A5" bold="true" box="[144,378,1847,1866]" pageId="7" pageNumber="234">Lemma color abaxial side</emphasis>
</th>
<td id="77631446945E005857C57547FC90F8B2" box="[498,785,1829,1885]" gridcol="1" gridrow="13" pageId="7" pageNumber="234">Green with purple centrally, otherwise straw-colored</td>
<td id="77631446945E0058551C7547FBFEF8B2" box="[811,1151,1829,1885]" gridcol="2" gridrow="13" pageId="7" pageNumber="234">Straw-colored or green with purple along the margins to the apex</td>
<td id="77631446945E005852B97547FA25F8B2" box="[1166,1444,1829,1885]" gridcol="3" gridrow="13" pageId="7" pageNumber="234">Green with purple when young then straw-colored</td>
</tr>
<tr id="34B27D3A945E005856A77511FA25F85F" box="[144,1444,1907,1968]" gridrow="14" pageId="7" pageNumber="234">
<th id="77631446945E005856A77511FE54F85F" box="[144,469,1907,1968]" gridcol="0" gridrow="14" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A77511FE0AF85F" bold="true" pageId="7" pageNumber="234">Lemma margin indument Lemma mucro length (mm)</emphasis>
</th>
<td id="77631446945E005857C57511FC90F85F" box="[498,785,1907,1968]" gridcol="1" gridrow="14" pageId="7" pageNumber="234">Denselly ciliate 1</td>
<td id="77631446945E0058551C7511FBFEF85F" box="[811,1151,1907,1968]" gridcol="2" gridrow="14" pageId="7" pageNumber="234">Smoth 0.250.5</td>
<td id="77631446945E005852B97511FA25F85F" box="[1166,1444,1907,1968]" gridcol="3" gridrow="14" pageId="7" pageNumber="234">Smooth (0.2)0.50.7</td>
</tr>
<tr id="34B27D3A945E005856A775A5FA25F811" box="[144,1444,1991,2046]" gridrow="15" pageId="7" pageNumber="234">
<th id="77631446945E005856A775A5FE54F811" box="[144,469,1991,2046]" gridcol="0" gridrow="15" pageId="7" pageNumber="234">
<emphasis id="B8F6A36A945EFFA056A775BBFE54F803" bold="true" box="[144,469,2009,2028]" pageId="7" pageNumber="234">Palea wing size (mm) and indument</emphasis>
</th>
<td id="77631446945E005857C575A5FC90F811" box="[498,785,1991,2046]" gridcol="1" gridrow="15" pageId="7" pageNumber="234">11.5; puberulent abaxially; glabrous adaxially</td>
<td id="77631446945E0058551C75A5FBFEF811" box="[811,1151,1991,2046]" gridcol="2" gridrow="15" pageId="7" pageNumber="234">0.250.5; puberulent on both surfaces</td>
<td id="77631446945E005852B975A5FA25F811" box="[1166,1444,1991,2046]" gridcol="3" gridrow="15" pageId="7" pageNumber="234">0.51; glabrous on both surfaces</td>
</tr>
</table>
</paragraph>
<paragraph id="8A3D7F789451FFAF568A72FCFBBAFE7F" blockId="8.[136,1452,158,508]" pageId="8" pageNumber="235">
<taxonomicName id="4D8204FB9451FFAF568A72FCFE34FF57" box="[189,437,158,184]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="8" pageNumber="235" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A9451FFAF568A72FCFE34FF57" box="[189,437,158,184]" italics="true" pageId="8" pageNumber="235">Guadua bambuzoniae</emphasis>
</taxonomicName>
differs from all other congeners by the following combination of characters: hollow and erect culms at the basal part, and solid and arched culms at the middle and upper part; development of primary branches with thorns at each culm node, oriented at 45° to the central axis; usually with a falcate, prominent, and fimbriate auricle at the upper part of the culm leaf sheath; elongated leaf sheaths, giving a lax appearance to the leaf complement; pseudospikelets linear-lanceolate, usually straight, with the lemmas densely ciliate at the margins and conspicuously mucronate; and paleas with conspicuously winged keels. This adaptation mechanism allows them to thrive in swampy environments and face these
<typeStatus id="5539C1DA9451FFAF54A07315FD53FE7E" box="[663,722,375,401]" pageId="8" pageNumber="235">types</typeStatus>
of challenges that nature offers.
</paragraph>
<paragraph id="8A3D7F789451FFAF568A73F8FCD9FE13" blockId="8.[136,1452,158,508]" pageId="8" pageNumber="235">
Like many species in this Amazon region of
<collectingCountry id="F2953FE89451FFAF548573F8FCA2FE5B" box="[690,803,410,436]" name="Colombia" pageId="8" pageNumber="235">Colombia</collectingCountry>
,
<taxonomicName id="4D8204FB9451FFAF551973F8FBA5FE5B" box="[814,1060,410,436]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="8" pageNumber="235" phylum="Tracheophyta" rank="species" species="bambuzoniae">
<emphasis id="B8F6A36A9451FFAF551973F8FBA5FE5B" box="[814,1060,410,436]" italics="true" pageId="8" pageNumber="235">Guadua bambuzoniae</emphasis>
</taxonomicName>
is endangered due to deforestation in its habitat. It is an ecologically important species for the conservation of swampy areas called “canaguchales” or “aguajales”, and for the biological diversity that is housed there.
</paragraph>
</subSubSection>
</treatment>
</document>