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<document id="332967CB3181FE570ADA0BCAF6D8E26F" ID-DOI="10.37828/em.2024.71.15" ID-ISSN="2336-9744" ID-Zenodo-Dep="13249314" ID-ZooBank="urn:lsid:zoobank.org:pub:5216E2D4-C04C-44DD-92E5-422F228C959B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1722974861239" checkinUser="felipe" docAuthor="Legalov, Andrei A., Vasilenko, Dmitry V. &amp; Perkovsky, Evgeny E." docDate="2024" docId="03FC9C22FFD7FF8427C3A7F7FBF4F99F" docLanguage="en" docName="EcolMontenegrina.71.165-171.pdf" docOrigin="Ecologica Montenegrina 71" docSource="http://dx.doi.org/10.37828/em.2024.71.15" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="3" docTitle="Trematosphindus kirejtshuki Legalov, Vasilenko &amp; Perkovsky, 2024, sp. n." docType="treatment" docUuid="AC889191-A6B0-4EDF-A99F-4ED65F1F4F67" docUuidSource="ZooBank" docVersion="1" lastPageNumber="168" masterDocId="FFC5E45AFFD6FF87277EA429FFA4FFA3" masterDocTitle="A new species of the genus Trematosphindus Li et Cai, 2021 (Coleoptera: Sphindidae) from Cenomanian Taimyr amber" masterLastPageNumber="171" masterPageNumber="165" pageNumber="166" updateTime="1727787188064" updateUser="guilherme" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4E4BDFA9798CA5F14AF2EE5F1021302A">A new species of the genus Trematosphindus Li et Cai, 2021 (Coleoptera: Sphindidae) from Cenomanian Taimyr amber</mods:title>
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<mods:namePart id="02CECE120254387B5C7D68383426B77A">Legalov, Andrei A.</mods:namePart>
<mods:affiliation id="329A23CF069396C9D87A004377CFDA40">Institute of Systematics and Ecology of Animals, SB RAS, Frunze Street 11, Novosibirsk, 630091, Russia;. &amp; Altai State University, Lenina Street 61, Barnaul, 656049, Russia; &amp; Tomsk State University, Lenina Prospekt 36, 634050 Tomsk, Russia;</mods:affiliation>
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<mods:namePart id="3F0FA0AF0973602B5833483B65EA7351">Vasilenko, Dmitry V.</mods:namePart>
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<mods:affiliation id="95D57E6B9F5307639CC00B31D1384B2B">Laboratory of Arthropods, A. A. Borissiak Paleontological Institute of the Russian Academy of Sciences, 117647 Moscow, Russia; https: // orcid. org / 0000 - 0002 - 4827 - 7290 &amp; Paleontological Laboratory, Cherepovets State University, 162600 Cherepovets, Russia;</mods:affiliation>
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<mods:namePart id="F94CEB544D1ACD3FE1FE318E98724C62">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="DFA1F409F7FB997FBA4AC7984C7C9113">Natural History Museum of Denmark, 2100 Copenhagen, Denmark; https: // orcid. org / 0000 - 0002 - 7959 - 4379 * Corresponding author. E-mail: fossilweevils @ gmail. com; https: // orcid. org / 0000 - 0001 - 7347 - 8169</mods:affiliation>
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<taxonomicName id="4C5556B7FFD7FF8627C3A7F7FDA8FC5B" authorityName="Legalov &amp; Vasilenko &amp; Perkovsky" authorityYear="2024" box="[189,524,990,1016]" class="Insecta" family="Sphindidae" genus="Trematosphindus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="kirejtshuki" status="sp. nov.">
<emphasis id="B921F126FFD7FF8627C3A7F7FDA8FC5B" box="[189,524,990,1016]" italics="true" pageId="1" pageNumber="166">Trematosphindus kirejtshuki</emphasis>
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(
<figureCitation id="136E31B1FFD7FF86251CA7F7FD6EFC5B" box="[610,714,990,1016]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="2.[189,265,1136,1159]" captionStartId-1="2.[189,265,2024,2047]" captionStartId-2="3.[189,265,1168,1191]" captionTargetBox-0="[208,1378,189,1116]" captionTargetBox-1="[208,1378,1221,2004]" captionTargetBox-2="[408,1178,189,1148]" captionTargetId-0="figure-42@2.[208,1378,189,1116]" captionTargetId-1="figure-44@2.[208,1378,1221,2004]" captionTargetId-2="figure-272@3.[408,1178,189,1148]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figure 1. Trematosphindus kirejtshuki sp. n., PIN 3426/258, head and prosternum, lateral view, Taimyr amber." captionText-1="Figure 2. Trematosphindus kirejtshuki sp. n., PIN 3426/258, head and prosternum, ventral view, Taimyr amber." captionText-2="Figure 3. Trematosphindus kirejtshuki sp. n., PIN 3426/258, head and pronotum, dorsal view, Taimyr amber." figureDoi-0="http://doi.org/10.5281/zenodo.13249316" figureDoi-1="http://doi.org/10.5281/zenodo.13249320" figureDoi-2="http://doi.org/10.5281/zenodo.13249322" httpUri-0="https://zenodo.org/record/13249316/files/figure.png" httpUri-1="https://zenodo.org/record/13249320/files/figure.png" httpUri-2="https://zenodo.org/record/13249322/files/figure.png" pageId="1" pageNumber="166">Figs. 1-3</figureCitation>
)
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<paragraph id="8BEA2D34FFD7FF8627C3A069FD01FBDF" blockId="1.[189,1399,1088,2025]" pageId="1" pageNumber="166">
<materialsCitation id="3B3D2769FFD7FF8627C3A069FD05FBDF" collectionCode="PIN" country="Russia" county="Taymyrsky Dolgano-Nenetsky District" location="Nizhnyaya Agapa" municipality="Krasnoyarskii Krai" pageId="1" pageNumber="166" specimenCount="1" typeStatus="type">
<emphasis id="B921F126FFD7FF8627C3A069FEDCFBF9" bold="true" box="[189,376,1088,1114]" pageId="1" pageNumber="166">
<typeStatus id="54EE9396FFD7FF8627C3A069FF59FBF9" box="[189,253,1088,1114]" pageId="1" pageNumber="166">Type</typeStatus>
material
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:
<collectionCode id="ED44B5F1FFD7FF8626EFA069FE64FBF9" box="[401,448,1088,1114]" pageId="1" pageNumber="166">PIN</collectionCode>
no. 3426/258,
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,
<collectingCounty id="628B55B8FFD7FF8624ECA069FAD6FBF9" box="[914,1394,1088,1114]" pageId="1" pageNumber="166">Taymyrsky Dolgano-Nenetsky District</collectingCounty>
,
<collectingMunicipality id="6B8EB74EFFD7FF8627C3A04BFE04FBDF" box="[189,416,1122,1148]" pageId="1" pageNumber="166">Krasnoyarskii Krai</collectingMunicipality>
,
<collectingCountry id="F3426DA4FFD7FF8626D2A04BFE59FBDF" box="[428,509,1122,1148]" name="Russia" pageId="1" pageNumber="166">Russia</collectingCountry>
, Cenomanian
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.
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<subSubSection id="C34F7EBFFFD7FF862662A0ADFB61F84A" pageId="1" pageNumber="166" type="description">
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<emphasis id="B921F126FFD7FF862662A0ADFE09FB3D" bold="true" box="[284,429,1156,1182]" pageId="1" pageNumber="166">Description</emphasis>
. Body brownish, covered with sparse setae. Head without distinct grooves. Mandibles large. Eyes large, convex, finely faceted. Frons flattened. Antennae 11-segmented. First and second antennomeres suboval. First antennomere 2.0 times as long as wide in middle. Second antennomere 1.6 times as long as wide in middle, 0.6 times as long as and 0.8 times as narrow as first antennomere. Thirdfourth antennomeres long-conical. Third antennomere about 2.9 times as long as wide at apex, subequal in length and about 0.5 as narrow as second antennomere. Fourth antennomere about 1.8 times as long as wide at apex, about 0.7 times as long as and slightly wider than third antennomere. Fifthsixth antennomeres conical. Fifth antennomere 1.5 times as long as wide at apex, subequal in length and about 1.2 times as wide as fourth antennomere. Sixth antennomere about 0.9 times as long as wide at apex, about 0.7 times as long as and about 1.3 times as wide as fifth antennomere. Seventh and eighth antennomeres wide-conical. Seventh antennomere slightly longer than width at apex, about 1.2 times as long as and subequal in width to sixth antennomere. Eighth antennomere about 0.7 times as long as wide at apex, subequal in length and about 1.5 times as wide as seventh antennomere. Antennal club distinct, consists of nintheleventh antennomeres, shorter than antennomeres 2-7 combined. Ninth antennomere about 0.7 times as long as wide at apex, about 1.7 times as long as and about 1.8 times as wide as eighth antennomere. Tenth antennomere about 0.5 times as long as wide at apex, about 0.9 times as long as and about 1.1 times as wide as ninth antennomere. Eleventh antennomere subequal in length and width in middle, about 1.6 times as long as and about 0.9 times as narrow as tenth antennomere. Pronotum campanulate, subequal to wide at apex, about 0.7 times as long as wide in middle and at base, without large cavities at anterior pronotal angles. Maximum width before base. Disk convex, coarsely punctate. Distances between punctures are much smaller than punctures. Sides rounded. Pronotal lateral edges dentate by quite blunt teeth. Precoxal portion of prosternum elongate, about 1.9 times as long as precoxal cavity length. Postcoxal portion of prosternum short, about 0.6 times as long as precoxal cavity length and about 0.3 times as long as precoxal portion. Distance between procoxae about 0.6 times as narrow as procoxal cavity width. Procoxal cavities closed externally. Tarsi five-segmented. Length of head:
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; length of pronotum:
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.
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<paragraph id="8BEA2D34FFD4FF8527C3A059FAC6FB24" blockId="2.[189,1378,1136,1159]" box="[189,1378,1136,1159]" pageId="2" pageNumber="167">
<emphasis id="B921F126FFD4FF8527C3A059FE87FB24" bold="true" box="[189,291,1136,1159]" pageId="2" pageNumber="167">Figure 1.</emphasis>
<taxonomicName id="4C5556B7FFD4FF852654A059FDFEFB24" authorityName="Legalov &amp; Vasilenko &amp; Perkovsky" authorityYear="2024" box="[298,602,1136,1159]" class="Insecta" family="Sphindidae" genus="Trematosphindus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="kirejtshuki" status="sp. nov.">
<emphasis id="B921F126FFD4FF852654A059FDFEFB24" box="[298,602,1136,1159]" italics="true" pageId="2" pageNumber="167">Trematosphindus kirejtshuki</emphasis>
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<taxonomicNameLabel id="A2124C5DFFD4FF85251FA059FD3FFB24" box="[609,667,1136,1159]" pageId="2" pageNumber="167" rank="species">sp. n.</taxonomicNameLabel>
, PIN 3426/258, head and prosternum, lateral view, Taimyr amber.
</paragraph>
</caption>
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<paragraph id="8BEA2D34FFD4FF8527C3A3C1FACDF85C" blockId="2.[189,1385,2024,2047]" box="[189,1385,2024,2047]" pageId="2" pageNumber="167">
<emphasis id="B921F126FFD4FF8527C3A3C1FE87F85C" bold="true" box="[189,291,2024,2047]" pageId="2" pageNumber="167">Figure 2.</emphasis>
<taxonomicName id="4C5556B7FFD4FF852654A3C1FDFEF85C" authorityName="Legalov &amp; Vasilenko &amp; Perkovsky" authorityYear="2024" box="[298,602,2024,2047]" class="Insecta" family="Sphindidae" genus="Trematosphindus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="kirejtshuki" status="sp. nov.">
<emphasis id="B921F126FFD4FF852654A3C1FDFEF85C" box="[298,602,2024,2047]" italics="true" pageId="2" pageNumber="167">Trematosphindus kirejtshuki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2124C5DFFD4FF85251FA3C1FD3FF85C" box="[609,667,2024,2047]" pageId="2" pageNumber="167" rank="species">sp. n.</taxonomicNameLabel>
, PIN 3426/258, head and prosternum, ventral view, Taimyr amber.
</paragraph>
</caption>
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<paragraph id="8BEA2D34FFD5FF8427C3A0B9FAEBFB04" blockId="3.[189,1359,1168,1191]" box="[189,1359,1168,1191]" pageId="3" pageNumber="168">
<emphasis id="B921F126FFD5FF8427C3A0B9FE87FB04" bold="true" box="[189,291,1168,1191]" pageId="3" pageNumber="168">Figure 3.</emphasis>
<taxonomicName id="4C5556B7FFD5FF842654A0B9FDFEFB04" authorityName="Legalov &amp; Vasilenko &amp; Perkovsky" authorityYear="2024" box="[298,602,1168,1191]" class="Insecta" family="Sphindidae" genus="Trematosphindus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="kirejtshuki" status="sp. nov.">
<emphasis id="B921F126FFD5FF842654A0B9FDFEFB04" box="[298,602,1168,1191]" italics="true" pageId="3" pageNumber="168">Trematosphindus kirejtshuki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2124C5DFFD5FF84251FA0B9FD3FFB04" box="[609,667,1168,1191]" pageId="3" pageNumber="168" rank="species">sp. n.</taxonomicNameLabel>
, PIN 3426/258, head and pronotum, dorsal view, Taimyr amber.
</paragraph>
</caption>
<subSubSection id="C34F7EBFFFD5FF842662A0DAFD3EFAAE" box="[284,666,1267,1293]" pageId="3" pageNumber="168" type="distribution">
<paragraph id="8BEA2D34FFD5FF842662A0DAFD3EFAAE" blockId="3.[189,1399,1267,1596]" box="[284,666,1267,1293]" pageId="3" pageNumber="168">
<emphasis id="B921F126FFD5FF842662A0DAFE08FAAE" bold="true" box="[284,428,1267,1293]" pageId="3" pageNumber="168">
<typeStatus id="54EE9396FFD5FF842662A0DAFEF8FAAE" box="[284,348,1267,1293]" pageId="3" pageNumber="168">Type</typeStatus>
strata
</emphasis>
. Dolgan Formation.
</paragraph>
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<subSubSection id="C34F7EBFFFD5FF842662A13DFDF7FAD1" pageId="3" pageNumber="168" type="diagnosis">
<paragraph id="8BEA2D34FFD5FF842662A13DFDF7FAD1" blockId="3.[189,1399,1267,1596]" pageId="3" pageNumber="168">
<emphasis id="B921F126FFD5FF842662A13DFD8CFA8D" bold="true" box="[284,552,1300,1326]" pageId="3" pageNumber="168">Differential diagnosis</emphasis>
. The new species differs from
<taxonomicName id="4C5556B7FFD5FF8424EFA13DFAD3FA8D" authority="Li et Cai, 2021" authorityName="Li et Cai" authorityYear="2021" box="[913,1399,1300,1326]" class="Insecta" family="Sphindidae" genus="Trematosphindus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="B921F126FFD5FF8424EFA13DFB65FA8D" box="[913,1217,1300,1326]" italics="true" pageId="3" pageNumber="168">Trematosphindus newtoni</emphasis>
<bibRefCitation id="EFC450C5FFD5FF8423B9A13DFAD3FA8D" author="Li, Y. - D. &amp; Tihelka, E. &amp; Liu, Z. - H. &amp; Huang, D. - Y. &amp; Cai, C. - Y." box="[1223,1399,1300,1326]" pageId="3" pageNumber="168" pagination="587 - 597" refId="ref2924" refString="Li, Y. - D., Tihelka, E., Liu, Z. - H., Huang, D. - Y., Cai, C. - Y. (2021) New mid-Cretaceous cryptic slime mold beetles and the early evolution of Sphindidae (Coleoptera: Cucujoidea). Arthropod Systematics &amp; Phylogeny, 79, 587 - 597. https: // doi. org / 10.3897 / asp. 79. e 72724" type="journal article" year="2021">
Li et Cai,
<quantity id="4CAD80D1FFD5FF842242A13DFAD3FA8D" box="[1340,1399,1300,1326]" metricMagnitude="1" metricUnit="m" metricValue="5.13334" pageId="3" pageNumber="168" unit="in" value="2021.0">2021</quantity>
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in the pronotum without large cavities at anterior pronotal angles, more blunt teeth on the sides of the pronotum, and finely faceted eyes.
</paragraph>
</subSubSection>
<subSubSection id="C34F7EBFFFD5FF842662A150FB14FA16" pageId="3" pageNumber="168" type="etymology">
<paragraph id="8BEA2D34FFD5FF842662A150FB14FA16" blockId="3.[189,1399,1267,1596]" pageId="3" pageNumber="168">
<emphasis id="B921F126FFD5FF842662A150FE00FA30" bold="true" box="[284,420,1401,1427]" pageId="3" pageNumber="168">Etymology</emphasis>
. Patronymic. In honour of Dr. Alexander G. Kirejtshuk (Zoological Institute of the Russian Academy of Sciences,
<collectingRegion id="4991E3D6FFD5FF842550A1B2FD71FA16" box="[558,725,1435,1461]" country="Russia" name="Saint Petersburg" pageId="3" pageNumber="168">St. Petersburg</collectingRegion>
,
<collectingCountry id="F3426DA4FFD5FF84259FA1B2FC90FA16" box="[737,820,1435,1461]" name="Russia" pageId="3" pageNumber="168">Russia</collectingCountry>
), who studied fossil
<taxonomicName id="4C5556B7FFD5FF84235BA1B2FB08FA16" box="[1061,1196,1435,1461]" class="Insecta" family="Sphindidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="family">Sphindidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F7EBFFFD5FF842662A194FB46FA74" box="[284,1250,1469,1495]" pageId="3" pageNumber="168" type="distribution">
<paragraph id="8BEA2D34FFD5FF842662A194FB46FA74" blockId="3.[189,1399,1267,1596]" box="[284,1250,1469,1495]" pageId="3" pageNumber="168">
<emphasis id="B921F126FFD5FF842662A194FE12FA74" bold="true" box="[284,438,1469,1495]" pageId="3" pageNumber="168">Localisation</emphasis>
.
<collectingCountry id="F3426DA4FFD5FF8426BAA194FDB3FA74" box="[452,535,1469,1495]" name="Russia" pageId="3" pageNumber="168">Russia</collectingCountry>
: Krasnoyarskii Krai: Taymyrsky Dolgano-Nenetsky District.
</paragraph>
</subSubSection>
<subSubSection id="C34F7EBFFFD5FF842662A1F6FBF4F99F" pageId="3" pageNumber="168" type="discussion">
<paragraph id="8BEA2D34FFD5FF842662A1F6FBF4F99F" blockId="3.[189,1399,1267,1596]" pageId="3" pageNumber="168">
<emphasis id="B921F126FFD5FF842662A1F6FE2AFA5A" bold="true" box="[284,398,1503,1529]" pageId="3" pageNumber="168">Remarks</emphasis>
. The species is placed in the genus
<taxonomicName id="4C5556B7FFD5FF84243EA1F6FB6FFA5A" authority="Li et Cai, 2021" authorityName="Li et Cai" authorityYear="2021" box="[832,1227,1503,1529]" class="Insecta" family="Sphindidae" genus="Trematosphindus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B921F126FFD5FF84243EA1F6FBA8FA5A" box="[832,1036,1503,1529]" italics="true" pageId="3" pageNumber="168">Trematosphindus</emphasis>
<bibRefCitation id="EFC450C5FFD5FF84236BA1F6FB6FFA5A" author="Li, Y. - D. &amp; Tihelka, E. &amp; Liu, Z. - H. &amp; Huang, D. - Y. &amp; Cai, C. - Y." box="[1045,1227,1503,1529]" pageId="3" pageNumber="168" pagination="587 - 597" refId="ref2924" refString="Li, Y. - D., Tihelka, E., Liu, Z. - H., Huang, D. - Y., Cai, C. - Y. (2021) New mid-Cretaceous cryptic slime mold beetles and the early evolution of Sphindidae (Coleoptera: Cucujoidea). Arthropod Systematics &amp; Phylogeny, 79, 587 - 597. https: // doi. org / 10.3897 / asp. 79. e 72724" type="journal article" year="2021">Li et Cai, 2021</bibRefCitation>
</taxonomicName>
because of an antennal club consists of three antennomeres, a head without distinct grooves, 11-segmented antennae, dentate lateral edges of the pronotum, and closed externally procoxal cavities.
</paragraph>
</subSubSection>
</treatment>
</document>