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<document id="EE22DB353148CD852BF136BE89504FC0" ID-DOI="10.3897/phytokeys.251.136715" ID-publisher-id="136715" URI-arpha="E68FB9D2-0824-5B4F-B123-3F1CB7D0FD81" article-type="research-article" checkinTime="1737991346399" checkinUser="pensoft" docAuthor="Villanueva-Tamayo, Boris, Paz-López, Carlos &amp; Ariza-Cortés, William" docDate="2025" docId="7C7C234D44D75FFC8D69DB94BDCDD3DF" docLanguage="en" docName="PhytoKeys 251: 189-201" docOrigin="PhytoKeys 251" docSource="https://phytokeys.pensoft.net/article/136715/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Compsoneura crassitepala Villanueva, Paz-Lopez &amp; W. Ariza 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="201" masterDocId="E68FB9D208245B4FB1233F1CB7D0FD81" masterDocTitle="A new tree species of Compsoneura (Myristicaceae) from the Andean forests on the Eastern Cordillera Range, Colombia" masterLastPageNumber="201" masterPageNumber="189" pageNumber="189" updateTime="1737991708000" updateUser="ExternalLinkService">
<mods:mods id="5CEB89EA2197C81712F4096800444E44" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE2CB2A16C40A1EC384F04341D6CBE03">
<mods:title id="871149AB70D941EED54AB10ED54B2086">A new tree species of Compsoneura (Myristicaceae) from the Andean forests on the Eastern Cordillera Range, Colombia</mods:title>
</mods:titleInfo>
<mods:name id="11EEAFF900D13A75D834B946021F82AA" type="personal">
<mods:role id="F730BF8359C42D7CFBD542754ED86F3D">
<mods:roleTerm id="F7F6AA4C2235023FE6FEE8D9794B3042">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE44E38DD813651ED76B1F76AF4E693B">Villanueva-Tamayo, Boris</mods:namePart>
<mods:nameIdentifier id="2744D608EAA6A79DCF54BD83F85A461B" type="ORCID">0000-0002-6929-3572</mods:nameIdentifier>
<mods:affiliation id="C407DDA48BCE1BD2F98027EB53C393AB">Jardín Botánico de Bogotá “ José Celestino Mutis ”, Av. José Celestino Mutis # 68 - 95, Bogotá, Colombia</mods:affiliation>
</mods:name>
<mods:name id="4DB4D5BA1EB073B0FBAAD4F371D5178B" type="personal">
<mods:role id="522090C10C3381F13B85DF49B4406721">
<mods:roleTerm id="2E30B942AFF0F790AEC4FD0770A24E0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0391EF41027D5E0FAAD5767646358E01">Paz-López, Carlos</mods:namePart>
<mods:nameIdentifier id="B79C7079A0299C0D4D4BA798B5DA6322" type="ORCID">https://orcid.org/0009-0000-0071-2380</mods:nameIdentifier>
<mods:affiliation id="F16F3D2BCD97C6DB82222ECB1205C12D">Herbario Forestal (UDBC), Universidad Distrital Francisco José de Caldas, Carrera 5 Este No. 15 - 82, Bogotá, Colombia</mods:affiliation>
</mods:name>
<mods:name id="FD95764E25CEE106615329FC4A51AF1F" type="personal">
<mods:role id="35D6A566AFAF6B249965CB520CB9801A">
<mods:roleTerm id="0BE5B95432E05949E0F31D3934BBE9BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E29C6DEC6A814E8F555053486DE2125">Ariza-Cortés, William</mods:namePart>
<mods:nameIdentifier id="CDC306C6BF3AB01880DD73EE45753974" type="ORCID">0000-0002-8423-8256</mods:nameIdentifier>
<mods:affiliation id="A5921EA6507B54795F613859417AF74F">Herbario Forestal (UDBC), Universidad Distrital Francisco José de Caldas, Carrera 5 Este No. 15 - 82, Bogotá, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="81D97CFD773E45311BF91AEEA8F32ED7">text</mods:typeOfResource>
<mods:relatedItem id="D0C57406FC7480B92F2DBD41FA318A3B" type="host">
<mods:titleInfo id="7405CA5EAFE52C777903399231D44887">
<mods:title id="C40DAB834A547965AE93A50B7F841D92">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="F5E73DB7C1369E29F72F66BE8B96DD24">
<mods:date id="AC90A5D43858D879C129CCB79E3F04A8">2025</mods:date>
<mods:detail id="CF32ED12281D24AAEAD46796C6F18808" type="pubDate">
<mods:number id="D5E80FD6C1287FA8B84B7C9579037F89">2025-01-27</mods:number>
</mods:detail>
<mods:detail id="9EB8780010A914E5B3124CEDF0D575CD" type="volume">
<mods:number id="84F54866EA9EAEDE7CA708F01AF41E45">251</mods:number>
</mods:detail>
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<mods:start id="70F3C13C668410E7765128E2568DE7A6">189</mods:start>
<mods:end id="B33C843EE2C8FF248CFEB242CD7AB4DA">201</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F3DA7FB1061F6B1ADC57F867419BC025">journal article</mods:classification>
<mods:identifier id="31ED6D1B75C6AB15E82968F3FF192837" type="DOI">10.3897/phytokeys.251.136715</mods:identifier>
</mods:mods>
<treatment id="7C7C234D44D75FFC8D69DB94BDCDD3DF" ID-DOI="http://doi.org/10.5281/zenodo.14749574" ID-Zenodo-Dep="14749574" ID-arpha="7C7C234D-44D7-5FFC-8D69-DB94BDCDD3DF" ID-ipni="urn:lsid:ipni.org:names:77355777-1" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF" LSID-IPNI="urn:lsid:ipni.org:names:77355777-1" httpUri="http://treatment.plazi.org/id/7C7C234D44D75FFC8D69DB94BDCDD3DF">
<subSubSection id="87363F686F14043297E13F73267324A7" type="nomenclature">
<paragraph id="25E6453BFAB85E7F08C5584989A4049E">
<taxonomicName id="AA6812296E391C7B63E206E99A8E3B32" authority="Villanueva, Paz-Lopez &amp; W. Ariza" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala" status="sp. nov.">
<emphasis id="2DF575E6249198FE01F79D2656E5B07C" italics="true">Compsoneura crassitepala</emphasis>
Villanueva, Paz-López &amp; W. Ariza
</taxonomicName>
<taxonomicNameLabel id="E5A700439386136163BFC27C7C105120" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="11021B8A430E6FAD29B7D89749282B7B" type="description">
<paragraph id="91E47917894618ADBDC64C0DA9B7D2F3">
<figureCitation id="E8880EEF99800D2B1E93515D62E4E03C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Compsoneura crassitepala A staminate flower B pistillate flower C fruit without pericarp showing the seed with one valve removed (left), fruit entire (right) D terminal branch with a pistillate inflorescence showing two leaves on the upper surface and one leaf on the lower surface (B. Villanueva-T et al. 6734) E trichomes on abaxial leaf blade F trichome (B. Villanueva-T et al. 6734). Illustration by Manuela Sánchez (JBB)." figureDoi="10.3897/phytokeys.251.136715.figure1" httpUri="https://binary.pensoft.net/fig/1234654">Figs 1</figureCitation>
,
<figureCitation id="F795B1ED9899DB377B6A4575AF945FB2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Compsoneura crassitepala A branches with staminate inflorescences and leaves showing both faces (C. Paz 2158 UDBC) B terminal branch bearing pistillate inflorescence (B Villanueva-T. et al. 6734) C underside of ferruginous leaf (B Villanueva-T. et al. 6734) D pistillate inflorescence (B Villanueva-T. et al. 6734) E staminate flower (C. Paz 2158 UDBC) F androecium visible by side cut (C. Paz 2158 UDBC) G pistil, showing the thick perianth (B Villanueva-T. et al. 6734) H fruit sulcate, densely ferruginous pubescent with stellate trichomes and vestigial tepals, showing the sulcate surface (B Villanueva-T et al. 6734) I external view of open pericarp, showing the sulcate surface (B Villanueva-T et al. 6734) J internal view of open pericarp (B Villanueva-T et al. 6734) K open fruit showing seed and pericarp thickness (B Villanueva-T et al. 6734) L seed side view without aril (B Villanueva-T et al. 6734) M seed without aril from above (B Villanueva-T et al. 6734) N detail of the thin and lignified aril (B Villanueva-T et al. 6734). Photos by Boris Villanueva-T., Lankester plate by Daniel Amaya-Jiménez." figureDoi="10.3897/phytokeys.251.136715.figure2" httpUri="https://binary.pensoft.net/fig/1234655">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAEAE" type="material">
<paragraph id="31C2DA77F2C60C30B6E14D91349431FA">
<heading id="44421A781BAF93CF58ED3EF56F0F9F21" reason="title">
<typeStatus id="E793C920062A8064AE82289955690025">Type</typeStatus>
.
</heading>
</paragraph>
<paragraph id="B47BA3609CDC4F83E42A5B06C437CDBD">
<collectingCountry id="C77E69496EFB9F3A8D0ECE2085799494" name="Colombia">Colombia</collectingCountry>
•
<materialsCitation id="9C580DC733BDBC0E6514C875A5019331" collectingDate="2023-07-29" collectorName="Carlos A. Paz" country="Brazil" county="El" elevation="1720" latitude="5.4130554" location="Municipio de Coper" longLatPrecision="1" longitude="-74.04417" municipality="Vereda Cucunuba" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca" typeStatus="holotype">
<collectingRegion id="5494585C038BCCC8A61E4CA7D1BBFBCF" country="Colombia" name="Boyaca">Boyacá</collectingRegion>
:
<location id="AD5464488B64F69E67192F9ED6A1AFD6" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:AD5464488B64F69E67192F9ED6A1AFD6" country="Brazil" county="El" latitude="5.4130554" longLatPrecision="1" longitude="-74.04417" municipality="Vereda Cucunuba" name="Municipio de Coper" stateProvince="Boyaca">Municipio de Coper</location>
,
<collectingMunicipality id="BA1599DB4983D9A5579F150ADF53CF58">Vereda Cucunubá</collectingMunicipality>
, finca
<collectingCounty id="C8E4577D137CD754B96D7D4E33006D04">El</collectingCounty>
<collectingCountry id="4868FB6104907B53EBC33D05E24C81B3" name="Brazil">Brasil</collectingCountry>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJEAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.044167,5.413056]}">
<geoCoordinate id="58BAAD6C51B610BB04D47FA98D58B860" degrees="5" direction="north" minutes="24" orientation="latitude" precision="1" seconds="47.0" value="5.4130554">5 ° 24 ' 47.0 &quot; N</geoCoordinate>
,
<geoCoordinate id="7548D908223CBABEBE1163CA322FBCAE" degrees="74" direction="west" minutes="02" orientation="longitude" precision="1" seconds="39.00" value="-74.04417">74 ° 02 ' 39.00 W</geoCoordinate>
</named-content>
,
<elevation id="FD1F8BB8D833F1943396298553D8D822" metricMagnitude="3" metricUnit="m" metricValue="1.72" unit="m" value="1720.0">
<quantity id="01A66F53E3706C5A093F9332C01E9542" metricMagnitude="3" metricUnit="m" metricValue="1.72" unit="m" value="1720.0">1720 m</quantity>
a. s. l.
</elevation>
,
<date id="A6BA8423DF94641CA6146B6246DEB32C" value="2023-07-29">
<collectingDate id="1EA5D895466BDEF730AF5355CE634810" value="2023-07-29">29 July 2023</collectingDate>
</date>
(fl
<specimenCount id="7AD9549724A1362CA837837F34F29E52" count="1" type="male">♂</specimenCount>
),
<emphasis id="E7F1142E0D9D6120D122A0F90222E9A3" italics="true">
<collectorName id="D8F7AFAF78C58BAA58917F8BDEDE7B0E">Carlos A. Paz</collectorName>
et al.
</emphasis>
2158 (
<typeStatus id="81C370518020494E245DA06FA6993D23" type="holotype">holotype</typeStatus>
: UDBC!, barcode no. 49999;
<typeStatus id="97494D1621B542264B183D882A2F1645" type="isotype">isotypes</typeStatus>
: COL!, FMB!, JAUM!, JBB!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQEAE" type="diagnosis">
<paragraph id="C93511270A7D30FEFD5C2626054A5D84">
<heading id="56E91D525035491460A5CD0BA9609A9D" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="2968E4E739A060B6CABB1632B1C2D362">
This new species can be distinguished from other species by its staminate and pistillate flowers, which have a spherical and fleshy perianth. It is similar to
<taxonomicName id="CD87C14841FE4602B262B33D73923D63" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="nallarettiana">
<emphasis id="F9FFA8C4344A2C39C98667400E2F29C0" italics="true">C. nallarettiana</emphasis>
</taxonomicName>
in its weakly brochidodromous secondary venation (only the first two veins are non-anastomosed) and differs in pedunculate fruits (vs. sessile fruits) and leaf blades oblong or elliptic-oblong (vs. leaf blades elliptic) and is similar to
<taxonomicName id="77D47EB4029CBA9D3544161F6ECB3A4B" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="cuatrecasasii">
<emphasis id="ACA1BC18378BA2992B3DFA99AF4D11FB" italics="true">C. cuatrecasasii</emphasis>
</taxonomicName>
in slightly brochidodromous secondary venation and trichomes minute 2 - branched. However, it differs in leaf oblong or elliptic-oblong (vs. leaf blades obovate to orbiculate), abaxial lamina densely ferruginous tomentose (vs. abaxial lamina glabrescent to glabrous when mature) leaf blade coppery when dry (vs. leaf blades that dry red-brown).
</paragraph>
<caption id="3B313F3C3D2B57D8BAFDA7478A49B230" ID-DOI="10.3897/phytokeys.251.136715.figure1" ID-arpha="5197E671-43FB-5DDE-B1C4-FC3458BEA4C4" httpUri="https://binary.pensoft.net/fig/1234654" startId="F1">
<paragraph id="DFCE2D69B4A2D6E890C7CFFFEDAFFD12">
<label id="B9328349B339F3B5DA7104986CF60ACA">Figure 1.</label>
</paragraph>
<paragraph id="53A688EC665BAA4248D69815C2DC3050">
<taxonomicName id="D04DE400F5576A7C1182DAF8681BFD93" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="C9514F5734E9474ACA134C3C7382ECFC" italics="true">Compsoneura crassitepala</emphasis>
</taxonomicName>
<emphasis id="E648CB04DC63F0C57ADD41AA7C60468C" bold="true">A</emphasis>
staminate flower
<emphasis id="FE1440C38C767DA7F750D7FEA2248770" bold="true">B</emphasis>
pistillate flower
<emphasis id="F65AB06D189CBBAC5B7880BEF318F4C9" bold="true">C</emphasis>
fruit without pericarp showing the seed with one valve removed (left), fruit entire (right)
<emphasis id="89F40B2D023922AC1DC92DB6C79C1EA2" bold="true">D</emphasis>
terminal branch with a pistillate inflorescence showing two leaves on the upper surface and one leaf on the lower surface (
<emphasis id="7B79049821D73A23FC44398EAEFF3CAC" italics="true">B. Villanueva-T et al. 6734</emphasis>
)
<emphasis id="17B1578899C859BB31F3A1499F379209" bold="true">E</emphasis>
trichomes on abaxial leaf blade
<emphasis id="36C9F96B512A50B9745024864539E590" bold="true">F</emphasis>
trichome (
<emphasis id="6BBDA98DFD72274DDE8227EEAAF3098E" italics="true">B. Villanueva-T et al. 6734</emphasis>
). Illustration by Manuela Sánchez (JBB).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EUGAE" type="description">
<paragraph id="929608331255ADEF8B0A1304A661FA7A">
<heading id="9199CFBFCF9A1B98CFB73CAB88EDCE0E" reason="title">Description.</heading>
</paragraph>
<paragraph id="9C81780471F1D84AD7E8D469086A5E68">
Dioecious tree,
<quantity id="0430F3BA76356E44B668367F9082C808" metricMagnitude="1" metricUnit="m" metricValue="1.8" unit="m" value="18.0">18 m</quantity>
tall, pyramidal crown, rhytidome slightly fissured and detachable in plates, the inner bark exuding sap red-hyaline; wood red-cream colour; branchlets ferruginous when dry,
<quantity id="811EBC2ED7581DC4E746F8B16E4376A4" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="11.3" metricValueMin="5.3" unit="mm" value="8.3" valueMax="11.3" valueMin="5.3">5.3–11.3 mm</quantity>
wide, tomentose with dendritic trichomes, when young densely ferruginous tomentellous, hairs dendritic trichomes with only one axis, densely brown ferruginous when old, densely covered with lenticels, 0.7–1.6 ×
<quantity id="C17CA82BBC78981C689D8C36B33074B1" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
.
<emphasis id="A62C61C62A84D417698E450B865B682B" bold="true" italics="true">Leaves</emphasis>
simple, alternate;
<emphasis id="D02F98C33A4E3DB03919F9734032D2A8" bold="true" italics="true">petioles</emphasis>
cylindrical stout, light ferruginous when fresh or coppery when dry, 12.0–13.4 ×
<quantity id="3CB7F6320D40AB5E4C24FB07EF0A5C51" metricMagnitude="-3" metricUnit="m" metricValue="4.449999999999999" metricValueMax="4.8" metricValueMin="4.1" unit="mm" value="4.449999999999999" valueMax="4.8" valueMin="4.1">4.1–4.8 mm</quantity>
;
<emphasis id="ED88B725FED3CA4A193CE2BEFBB27A21" bold="true" italics="true">leaf blades</emphasis>
coriaceous or strongly coriaceous when dried, oblong or elliptic-oblong, 21.0–25.5 ×
<quantity id="B78E356074206B7B01A0FD7CCB38978B" metricMagnitude="-2" metricUnit="m" metricValue="7.35" metricValueMax="8.2" metricValueMin="6.5" unit="cm" value="7.35" valueMax="8.2" valueMin="6.5">6.5–8.2 cm</quantity>
, width at ¼ length 5.5–7.0 cm, width at ½ length
<quantity id="42F199ACECEA01B128D8136A7015949D" metricMagnitude="-2" metricUnit="m" metricValue="7.3" metricValueMax="8.1" metricValueMin="6.5" unit="cm" value="7.3" valueMax="8.1" valueMin="6.5">6.5–8.1 cm</quantity>
, width at ¾ length
<quantity id="E22F2D18525DCC7275014C8DB798149F" metricMagnitude="-2" metricUnit="m" metricValue="5.6" metricValueMax="6.2" metricValueMin="5.0" unit="cm" value="5.6" valueMax="6.2" valueMin="5.0">5–6.2 cm</quantity>
; base rounded or obtuse or slightly attenuate, not revolute; apex acute, acuminate or apiculate, acumen
<quantity id="1F16D112091A5B0B7F598EAD3127F988" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="8.1" metricValueMin="4.5" unit="mm" value="6.3" valueMax="8.1" valueMin="4.5">4.5–8.1 mm</quantity>
; margin entire;
<emphasis id="05F542A5F603B609AA0B79CC3CACF682" bold="true" italics="true">adaxial</emphasis>
lamina surface shiny, light green or yellow-green when fresh, coppery when dry, with dispersed dendritic trichomes in young leaves;
<emphasis id="C24C32E5ED9090CFCA1BBCBF509B6505" bold="true" italics="true">abaxial</emphasis>
lamina surface dull, densely ferruginous tomentose, dendritic trichomes, light orange when fresh, yellowish-brown when dry; costa raised above near the base, scattered dendritic trichomes on young leaves and at the base of older leaves, raised; secondary veins 11–16 per side, spaced
<quantity id="4D9A47B42227414A43B1F8438584943B" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.6" metricValueMin="0.6" unit="mm" value="11.0" valueMax="16.0" valueMin="6.0">6–16 mm</quantity>
, slight arcuate ascending, the first two veins not anastomosing completely or only between, the next veins strongly brochidodromus.
<emphasis id="D5790B2FA0320CE7E4352D35ACD8BAD4" bold="true" italics="true">Staminate inflorescences</emphasis>
: 3.1–6.1 ×
<quantity id="DFEE2B5B4C888A978E4BBCFA3FD81A94" metricMagnitude="-2" metricUnit="m" metricValue="1.955" metricValueMax="2.6" metricValueMin="1.31" unit="cm" value="1.955" valueMax="2.6" valueMin="1.31">1.31–2.6 cm</quantity>
long, rachis direction in a zig-zag pattern, axillary, 1–2 - paniculate, densely ferruginous tomentellous with dendritic trichomes, peduncle 5.2–13.3 ×
<quantity id="8562E04DF4CE07E72B2C45BBF44F3E37" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.7" metricValueMin="1.7" unit="mm" value="2.2" valueMax="2.7" valueMin="1.7">1.7–2.7 mm</quantity>
, main axes with 3–8 ramifications, alternate.
<emphasis id="199B9E6AB76FBCB11A78FECB7187A27F" bold="true" italics="true">Staminate flower buds</emphasis>
: subglobose densely ferruginous tomentose.
<emphasis id="BEAD948BD9605849C8844E45B3D1C791" bold="true" italics="true">Staminate flowers</emphasis>
: subglobose or obovoid-globose, in fascicles of 4–8 flowers, fleshy when fresh and dry, densely ferruginous tomentellous, with dendritic trichomes on the outer side, thinly ferruginous, covered by dendritic trichomes inside, borne on a receptacle
<quantity id="B0F01C5419F0A6160519724B292E9662" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="3.4" metricValueMin="1.5" unit="mm" value="2.45" valueMax="3.4" valueMin="1.5">1.5–3.4 mm</quantity>
wide; pedicels 0.5–0.6 ×
<quantity id="5DB002CF40C34D4E6EBFB62D8D3EAE1E" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="cm" value="0.11" valueMax="0.12" valueMin="0.1">0.1–0.12 cm</quantity>
; perianth 3.9–4.6 ×
<quantity id="C58A0C011B67CD2A1CDB6235DAA93048" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.5" metricValueMin="3.6" unit="mm" value="4.05" valueMax="4.5" valueMin="3.6">3.6–4.5 mm</quantity>
, globose; lobes 3–4, fleshy, erect, triangular or broadly triangular, apex reflexed when mature, 1.5–1.8 ×
<quantity id="8C90A43B038435B5EF98729A21CC67F3" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" unit="mm" value="2.15" valueMax="2.4" valueMin="1.9">1.9–2.4 mm</quantity>
, connate by ⅔;
<emphasis id="AC5BF181A74454C1723BCC7589D4276A" bold="true" italics="true">androecium</emphasis>
:
<quantity id="E816BD562F580FF3C37627434B6AB27E" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.8–2.3 mm</quantity>
long, filament column 0.9–2.1 ×
<quantity id="28645BF48B2545BF1AA81174230C68A1" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.8" metricValueMin="0.6" unit="mm" value="1.2" valueMax="1.8" valueMin="0.6">0.6–1.8 mm</quantity>
, with a ferruginous, tomentose ring of dendritic trichomes at the base, anthers 6, dorsally adnate by ⅓, strongly erect ascending from base to apex 0.7–1.1 ×
<quantity id="188F45E02BBAED9F35C58BF79B7BF045" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
obtuse to apex.
<emphasis id="9058748C099925B81B6E562987791DDD" bold="true" italics="true">Pistillate inflorescences</emphasis>
: 1.9–2.1 ×
<quantity id="018CAD7349572165077732E8F6C9B849" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" unit="cm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 cm</quantity>
, rachis direction in a zig-zag pattern, axillary, 1–2 paniculate, densely ferruginous tomentellous with dendritic trichomes, peduncle 5.2–7.4 ×
<quantity id="91C58D338F25DA7972FEACCB98442B73" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.2" metricValueMin="2.3" unit="mm" value="2.75" valueMax="3.2" valueMin="2.3">2.3–3.2 mm</quantity>
, main axes with 1–2 ramifications, alternate.
<emphasis id="FC4EFD455CE72BD191038C294C81E534" bold="true" italics="true">Pistillate flower buds</emphasis>
: subglobose densely ferruginous tomentose.
<emphasis id="B1FECD0EC3266D414A4C3EE13FED75E0" bold="true" italics="true">Pistillate flowers</emphasis>
: subglobose or obovoid-globose, in fascicles of 4–5 flowers, fleshy when fresh and dry, densely ferruginous tomentellous, with dendritic trichomes on the outer side, thinly ferruginous, covered by dendritic trichomes inside; borne on a receptacle
<quantity id="DD8700911A2CA3B68B6D231E643E2596" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" unit="mm" value="2.8" valueMax="3.1" valueMin="2.5">2.5–3.1 mm</quantity>
wide, pedicels 0.1–0.5 ×
<quantity id="DFA009A68DD32E13F8C15EF8D32CA765" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" unit="cm" value="0.135" valueMax="0.2" valueMin="0.07">0.07–0.2 cm</quantity>
; perianth fleshy when fresh and dry, 5.3–5.7 ×
<quantity id="0F907C6B7667F7FE69689A3C022228E6" metricMagnitude="-3" metricUnit="m" metricValue="5.7" metricValueMax="5.9" metricValueMin="5.5" unit="mm" value="5.7" valueMax="5.9" valueMin="5.5">5.5–5.9 mm</quantity>
, lobes 3–4, fleshy, erect, triangular or broadly triangular, with lobes slightly curved at apex at maturity, 0.8–1.3 ×
<quantity id="A5ED069F1B9CF3933E00926D86E67B81" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" unit="mm" value="0.95" valueMax="1.3" valueMin="0.6">0.6–1.3 mm</quantity>
, connate by ⅓, densely ferruginous tomentellous with dendritic trichomes in both sides; ovary, ovate, tannish-brown, cover by densely ferruginous tomentellous, fresh or dry, 2.8–3.7 ×
<quantity id="399668D5350E5FBE985F7F931900099F" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2–2.8 mm</quantity>
; stigma bilobed, dark brown, 0.4–0.9 ×
<quantity id="0C5C05C376DAC5BE1B50AD0BB963F4A2" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" unit="mm" value="0.55" valueMax="0.8" valueMin="0.3">0.3–0.8 mm</quantity>
. Infructescences axillary, solitary or paired,
<quantity id="B43943FD25C48B70A6BF89C6BCD30369" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="8.5" metricValueMin="5.0" unit="cm" value="6.75" valueMax="8.5" valueMin="5.0">5–8.5 cm</quantity>
long; peduncle 11.2–11.9 ×
<quantity id="900CE01177F239DD3A8839364070114C" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="5.3" metricValueMin="3.3" unit="mm" value="4.3" valueMax="5.3" valueMin="3.3">3.3–5.3 mm</quantity>
, terete or subterete, with short tomentellous dendritic trichomes; fruits 1–2 per infructescence.
<emphasis id="6B304B5562C20285A61948C9C35A29E3" bold="true" italics="true">Fruits</emphasis>
rounded, brown when fresh, with the surface strongly and irregularly sulcate, 4.3–6.2 ×
<quantity id="64C992EF77ACFD32882E6E5E73F89556" metricMagnitude="-2" metricUnit="m" metricValue="5.2" metricValueMax="6.2" metricValueMin="4.2" unit="cm" value="5.2" valueMax="6.2" valueMin="4.2">4.2–6.2 cm</quantity>
, grooves 1.0–5.0 mm deep,
<quantity id="78B695C105229C3A01240A3AC38918D8" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
wide, thinly ferruginous, with dendritic trichomes outside, dehiscent; seed (1), 3.3–4.5 ×
<quantity id="27F19FB74B919DEBF3EEDB88ADB2E70F" metricMagnitude="-2" metricUnit="m" metricValue="3.6999999999999997" metricValueMax="4.1" metricValueMin="3.3" unit="cm" value="3.6999999999999997" valueMax="4.1" valueMin="3.3">3.3–4.1 cm</quantity>
, testa minutely reticulate, shiny, brown, aril thin and lignified when mature, pale brown fresh or dry, lacinate, reduced,
<quantity id="25801D976A1EBB179A3ABDF49D291B26" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 cm</quantity>
long.
</paragraph>
<caption id="11BD9856F6AC113A5D6BFD97F03BEAB2" ID-DOI="10.3897/phytokeys.251.136715.figure2" ID-arpha="3209DF8E-9C04-524D-AE3C-CA7A6969B11F" httpUri="https://binary.pensoft.net/fig/1234655" startId="F2">
<paragraph id="00F5E9D5C8B7982CA2CBD41EE5A64D89">
<label id="C6CD28182964D7530AB6F0A875262E72">Figure 2.</label>
</paragraph>
<paragraph id="088C0386AD69DAB6C78D182005AE3FE5">
<taxonomicName id="CCA0EE512DBA37B62076208BD8755DCC" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="D5D61FF31418C92786B4C467A0F93AE3" italics="true">Compsoneura crassitepala</emphasis>
</taxonomicName>
<emphasis id="A68428215B4425027DF3F4F53E95E7D1" bold="true">A</emphasis>
branches with staminate inflorescences and leaves showing both faces
<emphasis id="B36EB2D685BD87A4DBCC0162F9E30AAD" italics="true">(C. Paz 2158</emphasis>
UDBC)
<emphasis id="BC729A3A87F4321E436E7762142A6BF2" bold="true">B</emphasis>
terminal branch bearing pistillate inflorescence (
<emphasis id="6E0AF80E1665C9E8042A9287DE228857" italics="true">B Villanueva-T. et al. 6734</emphasis>
)
<emphasis id="F5068E5F77D78861559C4CD5F533303C" bold="true">C</emphasis>
underside of ferruginous leaf (
<emphasis id="5AF77813094BBE99136D8DD08418BDE8" italics="true">B Villanueva-T. et al. 6734</emphasis>
)
<emphasis id="2F10C6CD70B117EC4CB1BE32E76A5033" bold="true">D</emphasis>
pistillate inflorescence (
<emphasis id="B0AD05F606D7918C24464377FEE34056" italics="true">B Villanueva-T. et al. 6734</emphasis>
)
<emphasis id="B9E44F9AB06581C54779A84FE3BC48C2" bold="true">E</emphasis>
staminate flower
<emphasis id="20F550F76CF96FCE19B807751979C8AB" italics="true">(C. Paz 2158</emphasis>
UDBC)
<emphasis id="0D9C6B0D997E4CAFE7252578CA4AD049" bold="true">F</emphasis>
androecium visible by side cut
<emphasis id="886E83682ECC32689B14DD40B915829E" italics="true">(C. Paz 2158</emphasis>
UDBC)
<emphasis id="53A6A2348B983BF27D4E68174E65F765" bold="true">G</emphasis>
pistil, showing the thick perianth (
<emphasis id="54FB46BE211077312AED9E2CDF108C5A" italics="true">B Villanueva-T. et al. 6734</emphasis>
)
<emphasis id="39DCBA6DEFD2357A16B8215EB3D4C87E" bold="true">H</emphasis>
fruit sulcate, densely ferruginous pubescent with stellate trichomes and vestigial tepals, showing the sulcate surface (
<emphasis id="0B334420176E14E6E0F74C8EF6CB196B" italics="true">B Villanueva-T et al. 6734</emphasis>
)
<emphasis id="0F253889B4FFD08102FC9CC2D614DFD2" bold="true">I</emphasis>
external view of open pericarp, showing the sulcate surface (
<emphasis id="C933FBB87403EACAA3D447092F7679D6" italics="true">B Villanueva-T et al. 6734</emphasis>
)
<emphasis id="01E60A94E94FC955208E3EE2907E3EB9" bold="true">J</emphasis>
internal view of open pericarp (
<emphasis id="AAA6A7E761B16EBE41DCCD9C7126E53A" italics="true">B Villanueva-T et al. 6734</emphasis>
)
<emphasis id="2BAED0AC3ED045B1EB070CFD7D6BA85B" bold="true">K</emphasis>
open fruit showing seed and pericarp thickness (
<emphasis id="1401F1B017B9E5852A4046715E8E8494" italics="true">B Villanueva-T et al. 6734</emphasis>
)
<emphasis id="3CCD0CA189C8F7E2421E6A9B10E33282" bold="true">L</emphasis>
seed side view without aril (
<emphasis id="EFD4DE439347930AA1E52FDDA0C6DD88" italics="true">B Villanueva-T et al. 6734</emphasis>
)
<emphasis id="2BE7E1AFF1ACE1D13D47BA1474F3DBC0" bold="true">M</emphasis>
seed without aril from above (
<emphasis id="594002AAA3F7AC99ED6BDD1B85A26C65" italics="true">B Villanueva-T et al. 6734</emphasis>
)
<emphasis id="04C32345BF732FFEB544C82DDCF6FB14" bold="true">N</emphasis>
detail of the thin and lignified aril (
<emphasis id="B661E8799430C30E5852491327C4F09F" italics="true">B Villanueva-T et al. 6734</emphasis>
). Photos by Boris Villanueva-T., Lankester plate by Daniel Amaya-Jiménez.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ERKAE" type="distribution">
<paragraph id="B07B1646B5DFA6DD8D9A1D5B08DD6090">
<heading id="FA055247E6FF607AD4F5DE184D3AD091" reason="title">Distribution and habitat.</heading>
</paragraph>
<paragraph id="EE0F9B5532B65328900A617C59A2F0AD">
According to
<bibRefCitation id="96523162A63208D9149623F0B0D4391D" DOI="10.18257/raccefyn.570" author="Cuatrecasas J" firstAuthor="Cuatrecasas" issue="1" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales" pagination="221-268" refId="B3" refString="Cuatrecasas J (1958) Aspectos de la vegetación natural de Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 10 (1): 221–268. https://doi.org/10.18257/raccefyn.570" title="Aspectos de la vegetación natural de Colombia." volume="10" year="1958">Cuatrecasas (1958)</bibRefCitation>
, this species inhabits the sub-Andean forests of the Eastern Cordillera in the
<collectingRegion id="F087A6753070191FB994A9296BF7B224" country="Colombia" name="Boyaca">Boyacá Department</collectingRegion>
, at elevations between 1500 and
<quantity id="A47D14F6367A5020E8F69D0A68DE70E2" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">1900 m</quantity>
. It is endemic to these forests, where it is associated with
<taxonomicName id="93EC85C8F1D8D0EC86BD3DFABD549B30" authority="(Mart.) H. Wendl." authorityName="H. Wendl." baseAuthorityName="Mart." class="Liliopsida" family="Arecaceae" genus="Dictyocaryum" kingdom="Plantae" order="Arecales" phylum="Tracheophyta" rank="species" species="lamarckianum">
<emphasis id="9AE940472F3969D8F2330615DC4AD1B9" italics="true">Dictyocaryum lamarckianum</emphasis>
(Mart.) H. Wendl.
</taxonomicName>
(
<taxonomicName id="01C8609544F6618D74F257FE83CE835A" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
),
<taxonomicName id="2918C25722657636028DFAB3DB625511" authority="Donn. Sm." authorityName="Donn. Sm." class="Magnoliopsida" family="Euphorbiaceae" genus="Conceveiba" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="pleiostemona">
<emphasis id="DEF35576D509552F887B9C92317D9B01" italics="true">Conceveiba pleiostemona</emphasis>
Donn. Sm.
</taxonomicName>
(
<taxonomicName id="33784F3C8C60B10F696B75BF5226C85C" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
) and
<taxonomicName id="4939814E970C0E8DF618D3132F43CB52" authority="(Bonpl.) P. J. F. Guim. &amp; Michelang." authorityName="P. J. F. Guim. &amp; Michelang." baseAuthorityName="Bonpl." class="Magnoliopsida" family="Melastomataceae" genus="Andesanthus" kingdom="Plantae" order="Myrtales" phylum="Tracheophyta" rank="species" species="lepidotus">
<emphasis id="9361D9259970C659822EB139103C8246" italics="true">Andesanthus lepidotus</emphasis>
(Bonpl.) P. J. F. Guim. &amp; Michelang.
</taxonomicName>
(
<taxonomicName id="54828A60E4A33CC4DC3832707023684E" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
) (Fig.
<figureCitation id="947AB7C79CB33123EC5E7FE91DA78882" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Compsoneura crassitepala." figureDoi="10.3897/phytokeys.251.136715.figure3" httpUri="https://binary.pensoft.net/fig/1234656">3</figureCitation>
).
</paragraph>
<caption id="453E571314593671450F30D9183429C7" ID-DOI="10.3897/phytokeys.251.136715.figure3" ID-arpha="2F96C51B-092F-57C1-88C5-D49DB52E58B9" httpUri="https://binary.pensoft.net/fig/1234656" startId="F3">
<paragraph id="1D1692A933A40C75F5D84DEE8141B3F9">
<label id="26E25432DA32C5686142AA1D5E7B60F4">Figure 3.</label>
</paragraph>
<paragraph id="66CE2CA8E0D7DF2CFC163174DBDF6474">
Distribution map of
<taxonomicName id="B5B3638602594389202F1E47300DAEDC" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="94C9090927F436BB80A310EC6CE9F0FF" italics="true">Compsoneura crassitepala</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="B6D0E5216A82648657AE0879FC0CE4B4">
<taxonomicName id="21E2E4CCC6F3E7C30064C6A037AB48BD" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="AF819BF1E42D0AFB63912C15A5E70CA2" italics="true">Compsoneura crassitepala</emphasis>
</taxonomicName>
has been recorded in sub-Andean forest fragments within pasturelands, at the boundary of the
<collectingRegion id="4BD096024D883814A58B4773D7B1D534" country="Colombia" name="Boyaca">Boyacá</collectingRegion>
and
<collectingRegion id="1D50C662AF30B62652A86837F36F3776" country="Colombia" name="Cundinamarca">Cundinamarca</collectingRegion>
Departments, on the western slopes of the Eastern Cordillera in
<collectingCountry id="3A499205905ED3353993812BAC9DF39C" name="Colombia">Colombia</collectingCountry>
. This new species is sympatric with
<taxonomicName id="2EEBA942E91AAA372463BD35FC52BFC2" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="rigidifolia">
<emphasis id="7C00DF39523F067F33440E3AA3CBD9C9" italics="true">C. rigidifolia</emphasis>
</taxonomicName>
, the species recorded at the highest altitude in
<collectingCountry id="23498634DE2DA0CEB7EFBE71BC6F9406" name="Colombia">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6NAE" type="material">
<paragraph id="489F0E2D6AB1FBC2C7B75B64C08DD84B">
<heading id="6D38808E733682AB04583701D1B80086" reason="title">Additional specimens examined.</heading>
</paragraph>
<paragraph id="BFD766CF54CA2ABF577D9355A7B79D2A">
<collectingCountry id="3574410D5F3A69637F9565BE901DF3C6" name="Colombia">Colombia</collectingCountry>
•
<materialsCitation id="2FF6DD2341671FB3EB5F2CB6CC725AC8" collectingDate="2022-06-21" collectorName="M. Betancour" country="Brazil" elevation="1700" latitude="5.4068475" location="Coper" longLatPrecision="1" longitude="-74.04822" municipality="Vereda Cucunuba Finca El" specimenCount="1" specimenCount-female="1" stateProvince="Boyaca">
<collectingRegion id="97C23F45E5A3F8F2359EE58BFB83688B" country="Colombia" name="Boyaca">Boyacá</collectingRegion>
:
<location id="05BB1992556AF29BF6FD8428BCD11867" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:05BB1992556AF29BF6FD8428BCD11867" country="Brazil" latitude="5.4068475" longLatPrecision="1" longitude="-74.04822" municipality="Vereda Cucunuba Finca El" name="Coper" stateProvince="Boyaca">Coper</location>
,
<collectingMunicipality id="1E2533895D111F7ED575EF67D6B51DB2">Vereda Cucunubá Finca El</collectingMunicipality>
<collectingCountry id="97322BFC2952A9ED8E512DEF37B385C0" name="Brazil">Brasil</collectingCountry>
,
<elevation id="AA4D646151211B515C65525C6E8C592E" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity id="851966AAB5DD00A00DB5A8DF73EFA63A" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a. s. l.
</elevation>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIOAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.048217,5.406847]}">
<geoCoordinate id="4A0DC8D352CF73CF0C705180FA286161" degrees="5" direction="north" minutes="24" orientation="latitude" precision="1" seconds="24.65" value="5.4068475">5 ° 24 ' 24.65 &quot; N</geoCoordinate>
,
<geoCoordinate id="6F42984F917CE280A2E04C9C47D51BD7" degrees="74" direction="west" minutes="02" orientation="longitude" precision="1" seconds="53.58" value="-74.04822">74 ° 02 ' 53.58 &quot; W</geoCoordinate>
</named-content>
,
<date id="BB829E6D910B8746AFA37A25B7B3BC30" value="2022-06-21">
<collectingDate id="293CAA549E4067A6EED2FF4C54C99D48" value="2022-06-21">21 Jun 2022</collectingDate>
</date>
(
<specimenCount id="CB2326EFC9426370520A1485BBF66FE6" count="1" type="female">♀</specimenCount>
fl, fr),
<emphasis id="47FAB453AC1C0FA9FD3BC9AF29ADC70A" italics="true">
<collectorName id="43C09BE189C4A183BA1E1BB0A000C363">M. Betancour</collectorName>
et al. 51
</emphasis>
(UDBC!)
</materialsCitation>
; •
<materialsCitation id="2B78F82658BA3928FD4AEE1CF296659D" collectingDate="2022-06-22" collectorName="M. Betancour" country="Brazil" elevation="1700" latitude="5.4067416" location="Ibid" longLatPrecision="1" longitude="-74.04823" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<location id="244A331C301D688EB5577F36C5188050" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:244A331C301D688EB5577F36C5188050" country="Brazil" latitude="5.4067416" longLatPrecision="1" longitude="-74.04823" name="Ibid" stateProvince="Boyaca">Ibid</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESOAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.048231,5.406742]}">
<geoCoordinate id="A6CC9935C4CAD88B627014D9EB2BD4E8" degrees="5" direction="north" minutes="24" orientation="latitude" precision="1" seconds="24.27" value="5.4067416">5 ° 24 ' 24.27 &quot; N</geoCoordinate>
,
<geoCoordinate id="D028C3924D20762E3E534B8193CBEB82" degrees="74" direction="west" minutes="02" orientation="longitude" precision="1" seconds="53.63" value="-74.04823">74 ° 02 ' 53.63 &quot; W</geoCoordinate>
</named-content>
,
<elevation id="B8DF73A9A2CB49848D103D623FA027B5" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity id="1EE06FD8CA7CA7841A9F93FD728362D0" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a. s. l.
</elevation>
,
<date id="BA3001AE05B6101B2E10E3AC47C4DCF1" value="2022-06-22">
<collectingDate id="3165222D5A005A7046949697B75B871C" value="2022-06-22">22 Jun 2022</collectingDate>
</date>
(
<specimenCount id="52CFAAF83C7C6898B2A86D69C61F7EA9" count="1" type="male">♂</specimenCount>
fl),
<emphasis id="2E4976EDD16E5C4E53AF9897CCFCDC72" italics="true">
<collectorName id="ADA6857D67ADD30E0BBA3C6C4A8AFE47">M. Betancour</collectorName>
et al. 58
</emphasis>
(JBB!, UDBC!)
</materialsCitation>
; •
<materialsCitation id="4D47CC0DA5FECB971378E0A0A035DDEB" collectingDate="2023-07-29" country="Brazil" elevation="1582" latitude="5.411528" location="Ibid" longLatPrecision="15" longitude="-74.03694" specimenCount="1" specimenCount-female="1" stateProvince="Boyaca">
<location id="479611FB85404D30F15BC2E6627E2069" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:479611FB85404D30F15BC2E6627E2069" country="Brazil" latitude="5.411528" longLatPrecision="15" longitude="-74.03694" name="Ibid" stateProvince="Boyaca">Ibid</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3OAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.036944,5.411528]}">
<geoCoordinate id="42B30BAD4E8B4C99E68AABB93FA3D855" degrees="5" direction="north" minutes="24" orientation="latitude" precision="1" seconds="41.5" value="5.411528">5 ° 24 ' 41.5 &quot; N</geoCoordinate>
,
<geoCoordinate id="2657F6911C775A19AEA25BAAA5ADFB96" degrees="74" direction="west" minutes="02" orientation="longitude" precision="15" seconds="13" value="-74.03694">74 ° 02 ' 13 &quot; W</geoCoordinate>
</named-content>
,
<elevation id="6E2B598C70B39583C03DE071FF49D5DE" metricMagnitude="3" metricUnit="m" metricValue="1.5819999999999999" unit="m" value="1582.0">
<quantity id="88C6189ED834F0D143A198B9C9782A0B" metricMagnitude="3" metricUnit="m" metricValue="1.5819999999999999" unit="m" value="1582.0">1582 m</quantity>
a. s. l.
</elevation>
,
<date id="25675FCE103117109A43C7AB9E16D8D4" value="2023-07-29">
<collectingDate id="AC7BED31672C3F410606BC6EDB73503F" value="2023-07-29">29 Jul 2023</collectingDate>
</date>
(
<specimenCount id="37583E374BE007C20C819C6B3BDBEC79" count="1" type="female">♀</specimenCount>
fl),
<emphasis id="4127FC31AE190B7FBF6A8938C0B1FD46" italics="true">B. Villanueva-T. et al. 6734</emphasis>
(JBB!, barcode no. 40030)
</materialsCitation>
; •
<materialsCitation id="68F2456084D18C8087F61C0D372E6A76" collectingDate="2023-07-29" country="Brazil" elevation="1575" latitude="5.411667" location="Ibid" longLatPrecision="21" longitude="-74.03667" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<location id="506AC4BC6A35B9516959759624D98426" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:506AC4BC6A35B9516959759624D98426" country="Brazil" latitude="5.411667" longLatPrecision="21" longitude="-74.03667" name="Ibid" stateProvince="Boyaca">Ibid</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGPAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.036667,5.411667]}">
<geoCoordinate id="437808C34522E1EB2D19399F3C7C1979" degrees="5" direction="north" minutes="24" orientation="latitude" precision="15" seconds="42" value="5.411667">5 ° 24 ' 42 &quot; N</geoCoordinate>
,
<geoCoordinate id="5FE15EB892A28AD09715E103169190FA" degrees="74" direction="west" minutes="02" orientation="longitude" precision="15" seconds="12" value="-74.03667">74 ° 02 ' 12 &quot; W</geoCoordinate>
</named-content>
,
<elevation id="252E4765F6EAF589ACF91CE17D951F78" metricMagnitude="3" metricUnit="m" metricValue="1.575" unit="m" value="1575.0">
<quantity id="00799F0C2402E013592C26B916176430" metricMagnitude="3" metricUnit="m" metricValue="1.575" unit="m" value="1575.0">1575 m</quantity>
a. s. l.
</elevation>
,
<date id="5C18F51066ABD7B92696F79F6F363B84" value="2023-07-29">
<collectingDate id="B0C4A2D51D7FD07556AF6B0269070DEA" value="2023-07-29">29 Jul 2023</collectingDate>
</date>
(
<specimenCount id="774A97D1511F949C896E9FAC4E98B534" count="1" type="male">♂</specimenCount>
fl),
<emphasis id="0EF51C29AF46FBA4A95C16E5EF6594E4" italics="true">B. Villanueva-T. et al. 6735</emphasis>
(JBB!, barcode nos. 40031 &amp; 40032)
</materialsCitation>
; •
<materialsCitation id="5C75D04CBB7B4FF6E226BEE3245AF20D" collectingDate="2023-07-29" collectorName="W. Ariza" country="Brazil" elevation="1769" latitude="5.4130554" location="Ibid" longLatPrecision="1" longitude="-74.04417" specimenCount="1" specimenCount-female="1" stateProvince="Boyaca">
<location id="89C1F3B2DED46B98C7DDF5573FA54925" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:89C1F3B2DED46B98C7DDF5573FA54925" country="Brazil" latitude="5.4130554" longLatPrecision="1" longitude="-74.04417" name="Ibid" stateProvince="Boyaca">Ibid</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQPAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.044167,5.413056]}">
<geoCoordinate id="6A3D6B5DD25FF4097FCFE5B0E2A1EC67" degrees="5" direction="north" minutes="24" orientation="latitude" precision="1" seconds="47.0" value="5.4130554">5 ° 24 ' 47.0 &quot; N</geoCoordinate>
,
<geoCoordinate id="F68436435537EFDD62C219487A159257" degrees="74" direction="west" minutes="02" orientation="longitude" precision="1" seconds="39.00" value="-74.04417">74 ° 02 ' 39.00 W</geoCoordinate>
</named-content>
,
<elevation id="765DD55F5A7C7797752D31D37098131F" metricMagnitude="3" metricUnit="m" metricValue="1.769" unit="m" value="1769.0">
<quantity id="281E7DD53BE22BBA3DA478105074AA53" metricMagnitude="3" metricUnit="m" metricValue="1.769" unit="m" value="1769.0">1769 m</quantity>
a. s. l.
</elevation>
,
<date id="D7E1F4BC24A16D1E926FD7D1D5B73BEB" value="2023-07-29">
<collectingDate id="14DD69F8D614E08223C4BA2D60FABD0B" value="2023-07-29">29 Jul 2023</collectingDate>
</date>
(
<specimenCount id="A26FCAE70F2523F3D0883E032583FB78" count="1" type="female">♀</specimenCount>
fl, fr),
<emphasis id="A22D3BEB238B54F4CC4574D94AEB81B3" italics="true">
<collectorName id="14591D77E2456C3A96F919871A5FD796">W. Ariza</collectorName>
et al. 10016
</emphasis>
(COL!, FMB!, JAUM!, JBB!, UDBC!)
</materialsCitation>
; •
<materialsCitation id="58008F7237D205499DA34333B414EAA0" collectingDate="2023-07-29" country="Brazil" elevation="1729" latitude="5.4135556" location="Ibid" longLatPrecision="1" longitude="-74.04325" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<location id="C1B3F50845AE70129A43C8CE58A5D022" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:C1B3F50845AE70129A43C8CE58A5D022" country="Brazil" latitude="5.4135556" longLatPrecision="1" longitude="-74.04325" name="Ibid" stateProvince="Boyaca">Ibid</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1PAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.043250,5.413556]}">
<geoCoordinate id="59D8180C84ECAD135FB2214B438477E3" degrees="5" direction="north" minutes="24" orientation="latitude" precision="1" seconds="48.80" value="5.4135556">5 ° 24 ' 48.80 &quot; N</geoCoordinate>
,
<geoCoordinate id="B45DDDF5E7C8210AA61BE6A691C1DBF4" degrees="74" direction="west" minutes="02" orientation="longitude" precision="1" seconds="35.70" value="-74.04325">74 ° 02 ' 35.70 &quot; W</geoCoordinate>
</named-content>
,
<elevation id="1D1FE817F7B7B80EDBA44A4E856B19E7" metricMagnitude="3" metricUnit="m" metricValue="1.7289999999999999" unit="m" value="1729.0">
<quantity id="378EA34C4D8C789C8D85D16660AF601E" metricMagnitude="3" metricUnit="m" metricValue="1.7289999999999999" unit="m" value="1729.0">1729 m</quantity>
a. s. l.
</elevation>
,
<date id="9128121CE0C47524F5C403EF2AB59294" value="2023-07-29">
<collectingDate id="19C2B166357C93A8CBC7C4A20337169D" value="2023-07-29">29 Jul 2023</collectingDate>
</date>
(
<specimenCount id="54EA280AA2810862CF20B32EA23C5A6F" count="1" type="male">♂</specimenCount>
fl)
<emphasis id="3E2FB5B77934D82BC55801B56332BE91" italics="true">A. Torrejano-M et al. 1356</emphasis>
(JBB!, UDBC!)
</materialsCitation>
; •
<materialsCitation id="B7C7FE3A5C83DDEC2AA8991996DE197A" collectingDate="2024-03-16" country="Brazil" elevation="1735" latitude="5.4130445" location="Ibid" longLatPrecision="1" longitude="-74.04325" specimenCount="1" stateProvince="Boyaca">
<location id="B99620E9FCEB2554FDD003E8FAE9298C" LSID="urn:lsid:plazi:treatment:7C7C234D44D75FFC8D69DB94BDCDD3DF:B99620E9FCEB2554FDD003E8FAE9298C" country="Brazil" latitude="5.4130445" longLatPrecision="1" longitude="-74.04325" name="Ibid" stateProvince="Boyaca">Ibid</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEQAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-74.043250,5.413044]}">
<geoCoordinate id="8701E7AB23F05BBE2CB51AD23FA0EB33" degrees="5" direction="north" minutes="24" orientation="latitude" precision="1" seconds="46.96" value="5.4130445">5 ° 24 ' 46.96 &quot; N</geoCoordinate>
,
<geoCoordinate id="7903DB065DB630C3D9DD9C556A02465D" degrees="74" direction="west" minutes="02" orientation="longitude" precision="1" seconds="35.70" value="-74.04325">74 ° 02 ' 35.70 &quot; W</geoCoordinate>
</named-content>
,
<elevation id="2443BEC977E55142B35C728D625E6452" metricMagnitude="3" metricUnit="m" metricValue="1.735" unit="m" value="1735.0">
<quantity id="4F70A64934DA2D2708B547B15E309874" metricMagnitude="3" metricUnit="m" metricValue="1.735" unit="m" value="1735.0">1735 m</quantity>
a. s. l.
</elevation>
,
<date id="3CB8BA087E0EA8F954806E2AF87CDA4D" value="2024-03-16">
<collectingDate id="3BBC2B5C032493CF1F0BA75416EDB7CA" value="2024-03-16">16 Mar 2024</collectingDate>
</date>
(fr)
<emphasis id="9DB1300D8F9E832D8BC0B0ADC3A3D10D" italics="true">B. Villanueva-T. et al. 6840</emphasis>
(JBB!, UDBC!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELQAE" type="phenology">
<paragraph id="67F28F765A48D1D7F10CB1577E4BA4D6">
<heading id="63853CC2F6017256D814EF4EA79B639E" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="6FD08DF3518BFF501F5014B1D4E15B5D">Collected with staminate and pistillate flowers, and fruit during June, July and fruit in March.</paragraph>
</subSubSection>
<subSubSection id="SECID0EQQAE" type="conservation status">
<paragraph id="583CF6A7721F7C0A2EBF59260CBB49CC">
<heading id="A0086CA9F6F98393E13887EE2E4D90CB" reason="title">Conservation status.</heading>
</paragraph>
<paragraph id="A8EE8B2B24CC1E88D75E6BC4EADF780D">
<taxonomicName id="416A16BB0420A473006B69348780516B" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="83DDD35C75822B9D8B3BEACB370FADBF" italics="true">Compsoneura crassitepala</emphasis>
</taxonomicName>
has an area of occupancy (
<abbrev id="ABBRID0EBRAE" xlink_title="area of occupancy">AOO</abbrev>
) of
<quantity id="B185421B49A36429DD4A755BB069CC05" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
<superScript id="EA5BCE0A6D33FD576A53BD681BBD5604">2</superScript>
and an extent of occurrence (
<abbrev id="ABBRID0EHRAE" xlink_title="extent of occurrence">EOO</abbrev>
) of approximately
<quantity id="85158A555A36A10B7D30BFB05308A19B" metricMagnitude="6" metricUnit="m" metricValue="4.2" unit="km" value="4200.0">4,200 km</quantity>
<superScript id="408E887CD12D4C93338B7C2BA6527D66">2</superScript>
and, according to the IUCN criteria (
<bibRefCitation id="523D5F94E5069C45756D4E919DCDDD86" refId="B7" refString="IUCN (2024) Guidelines for Using the IUCN Red List Categories and Criteria. Version 16. Prepared by the Standards and Petitions Committee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2024">IUCN 2024</bibRefCitation>
), it should be considered “ Endangered ” (
<abbrev id="ABBRID0ERRAE" xlink_title="“Endangered”">EN</abbrev>
) (B 1 ab). This area encompasses a mosaic of pastures with exploited forests and limited cultivation. Currently, there are no protected areas or policies that support the conservation of the species’ habitat.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVRAE" type="etymology">
<paragraph id="C713BDF9639B3ECE1BB40F863FEEF784">
<heading id="2AE2F589EF275A9B27A51B9E1BA92688" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="15E880258AD86264FD411D5C1487AE96">The specific epithet of this species refers to its unusually thick and fleshy tepals.</paragraph>
</subSubSection>
<subSubSection id="SECID0E1RAE" type="Common names">
<paragraph id="81E53D4F7ACFC59C8932FC52063C8A4A">
<heading id="6853EDEBCB6C731381887EA601CB103A" reason="title">Common names.</heading>
</paragraph>
<paragraph id="B85589311666DBEC564EDB1BAFFF5BDD">
Sangrino (
<collectingCountry id="0AB60984E5150F45646310B18FC05ECE" name="Colombia">Colombia</collectingCountry>
,
<collectingRegion id="D8B32650A025CABE792FEE387851103E" country="Colombia" name="Boyaca">Boyacá</collectingRegion>
;
<emphasis id="99957167AE20F8CC2FFFE8D42F7061A2" italics="true">C. Paz et al. 2158</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECSAE" type="discussion">
<paragraph id="748608B2AB3EEE6733A273A543444D55">
<heading id="7C0D7FED469F479CD5DDA5BC5E97E471" reason="title">Discussion.</heading>
</paragraph>
<paragraph id="B9F48627D639DFB6D2382FA1D0DC694D">
Based on different diagnostic characteristics such as secondary vein, drying colour of leaf blades and pericarps,
<bibRefCitation id="56E4D46FE8FC0D47C40F449D568F5655" author="Janovec PJ" firstAuthor="Janovec" journalOrPublisher="thesis, Texas A &amp; M University, College Station, Texas" refId="B8" refString="Janovec PJ (2000) A systematic study of Compsoneura (Candolle) Warburg, a Neotropical member of the nutmeg family (Myristicaceae). PhD. thesis, Texas A &amp; M University, College Station, Texas, 359 pp." title="A systematic study of Compsoneura (Candolle) Warburg, a Neotropical member of the nutmeg family (Myristicaceae). PhD." year="2000">Janovec (2000)</bibRefCitation>
proposed to divide the genus
<taxonomicName id="DAEDB2F76EA8A70BF96A383AC5AF1514" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="genus">
<emphasis id="0F0D3BD3AE046B9425CDC4F2AEBECC6F" italics="true">Compsoneura</emphasis>
</taxonomicName>
into two sections. He redefined the classic study of Warburg in 1897, but these sections are not formally published. Other species published after 2000 (i. e.
<taxonomicName id="87492C70C85CCA279991C171161D2D0C" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="choibo">
<emphasis id="1F26C2193E6E7FA2CB36E06AD41F7E10" italics="true">C. choibo</emphasis>
</taxonomicName>
and
<taxonomicName id="2D4B076D1992D67328DA3CA6F5C2BF67" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="lapidiflora">
<emphasis id="6BC41BE44E34CB53A469E88333CF4159" italics="true">C. lapidiflora</emphasis>
</taxonomicName>
) do not fit within the two informally proposed sections and, for the last species, its fruit is unknown. Therefore, we prefer to classify the new species within Warburg’s classic concept of the
<emphasis id="61495B47561B58F93767148C8262D11A" italics="true">Eucompsoneura</emphasis>
section, due to its partially fixed anthers to the filament (
<bibRefCitation id="D3B5C7BD8F3589CD65DD8BE734DDF64A" author="Warburg" firstAuthor="Warburg" journalOrPublisher="Nova Acta Academiae Caesareae Leopoldino- Carolinae Germanicae Naturae Curiosorum" pagination="1-680" refId="B27" refString="Warburg O (1897) Monographie der Myristicaceen. Nova Acta Academiae Caesareae Leopoldino- Carolinae Germanicae Naturae Curiosorum 68: 1–680." title="Monographie der Myristicaceen." volume="68" year="1897">Warburg 1897</bibRefCitation>
,
<bibRefCitation id="7A00043385DAB970D3A28E36E446C234" DOI="10.1002/fedr.19050010503" author="Warburg" firstAuthor="Warburg" refId="B28" refString="Warburg O (1905) Myristicaceae Costaricenses (Original diagnosen), Repertorium specierum novarum regni vegetabilis. Selbstverlag des Herausgebers, 71 pp. https://doi.org/10.1002/fedr.19050010503" year="1905">1905</bibRefCitation>
).
</paragraph>
<paragraph id="AF049D7725957ADF39D84FDF0E7B537C">
<taxonomicName id="4966AE016A4FD1B82222183659B09341" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="E0F57AF2B9AED8D8DABD14812B7214A5" italics="true">Compsoneura crassitepala</emphasis>
</taxonomicName>
shows a combination of characters that differentiate it from the other species in the genus. The thin and lignified aril not previously recorded in
<taxonomicName id="6A9827FFF48EF302A9EB6E4293D40D5A" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="genus">
<emphasis id="5E4DB1E2E800A3180656B9C2D777C88C" italics="true">Compsoneura</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="92FCCF6EE28536514E4754C5CCFB198C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Compsoneura crassitepala A branches with staminate inflorescences and leaves showing both faces (C. Paz 2158 UDBC) B terminal branch bearing pistillate inflorescence (B Villanueva-T. et al. 6734) C underside of ferruginous leaf (B Villanueva-T. et al. 6734) D pistillate inflorescence (B Villanueva-T. et al. 6734) E staminate flower (C. Paz 2158 UDBC) F androecium visible by side cut (C. Paz 2158 UDBC) G pistil, showing the thick perianth (B Villanueva-T. et al. 6734) H fruit sulcate, densely ferruginous pubescent with stellate trichomes and vestigial tepals, showing the sulcate surface (B Villanueva-T et al. 6734) I external view of open pericarp, showing the sulcate surface (B Villanueva-T et al. 6734) J internal view of open pericarp (B Villanueva-T et al. 6734) K open fruit showing seed and pericarp thickness (B Villanueva-T et al. 6734) L seed side view without aril (B Villanueva-T et al. 6734) M seed without aril from above (B Villanueva-T et al. 6734) N detail of the thin and lignified aril (B Villanueva-T et al. 6734). Photos by Boris Villanueva-T., Lankester plate by Daniel Amaya-Jiménez." figureDoi="10.3897/phytokeys.251.136715.figure2" httpUri="https://binary.pensoft.net/fig/1234655">2 N</figureCitation>
), the rachis in staminate inflorescence in a zig-zag pattern (Fig.
<figureCitation id="8B8C71C8BDE586AD446DF328D6377847" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Compsoneura crassitepala A branches with staminate inflorescences and leaves showing both faces (C. Paz 2158 UDBC) B terminal branch bearing pistillate inflorescence (B Villanueva-T. et al. 6734) C underside of ferruginous leaf (B Villanueva-T. et al. 6734) D pistillate inflorescence (B Villanueva-T. et al. 6734) E staminate flower (C. Paz 2158 UDBC) F androecium visible by side cut (C. Paz 2158 UDBC) G pistil, showing the thick perianth (B Villanueva-T. et al. 6734) H fruit sulcate, densely ferruginous pubescent with stellate trichomes and vestigial tepals, showing the sulcate surface (B Villanueva-T et al. 6734) I external view of open pericarp, showing the sulcate surface (B Villanueva-T et al. 6734) J internal view of open pericarp (B Villanueva-T et al. 6734) K open fruit showing seed and pericarp thickness (B Villanueva-T et al. 6734) L seed side view without aril (B Villanueva-T et al. 6734) M seed without aril from above (B Villanueva-T et al. 6734) N detail of the thin and lignified aril (B Villanueva-T et al. 6734). Photos by Boris Villanueva-T., Lankester plate by Daniel Amaya-Jiménez." figureDoi="10.3897/phytokeys.251.136715.figure2" httpUri="https://binary.pensoft.net/fig/1234655">2 A</figureCitation>
) and the subglobose or obovoid-globose and fleshy perianth in staminate and pistillate flowers (Fig.
<figureCitation id="55A7BF2A204AA0764572C8CE5AF39C83" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Compsoneura crassitepala A branches with staminate inflorescences and leaves showing both faces (C. Paz 2158 UDBC) B terminal branch bearing pistillate inflorescence (B Villanueva-T. et al. 6734) C underside of ferruginous leaf (B Villanueva-T. et al. 6734) D pistillate inflorescence (B Villanueva-T. et al. 6734) E staminate flower (C. Paz 2158 UDBC) F androecium visible by side cut (C. Paz 2158 UDBC) G pistil, showing the thick perianth (B Villanueva-T. et al. 6734) H fruit sulcate, densely ferruginous pubescent with stellate trichomes and vestigial tepals, showing the sulcate surface (B Villanueva-T et al. 6734) I external view of open pericarp, showing the sulcate surface (B Villanueva-T et al. 6734) J internal view of open pericarp (B Villanueva-T et al. 6734) K open fruit showing seed and pericarp thickness (B Villanueva-T et al. 6734) L seed side view without aril (B Villanueva-T et al. 6734) M seed without aril from above (B Villanueva-T et al. 6734) N detail of the thin and lignified aril (B Villanueva-T et al. 6734). Photos by Boris Villanueva-T., Lankester plate by Daniel Amaya-Jiménez." figureDoi="10.3897/phytokeys.251.136715.figure2" httpUri="https://binary.pensoft.net/fig/1234655">2 F, G</figureCitation>
).
</paragraph>
<paragraph id="4E9A24CF55037E96B9D26EE1D804BB54">
This species shares characteristics with
<taxonomicName id="577D15094C0975754EE6F63D4214A8E5" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="nallarettiana">
<emphasis id="5E0E81A144CA0DEE11366F88CFE79D8B" italics="true">Compsoneura nallarettiana</emphasis>
</taxonomicName>
, such as the slightly brochidodromous secondary venation and the anastomosis from the second vein. However,
<taxonomicName id="FB93A0B6CA31C5D5E2EB0FFF972B403B" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="59E53934BE182FA159EA249E0A058EDE" italics="true">C. crassitepala</emphasis>
</taxonomicName>
can be distinguished by its pedunculate fruits (vs. sessile fruits) and elliptic-oblong leaf blades (vs. elliptic or elliptic-ovate leaves). The new species shares the following characters with
<taxonomicName id="54F8817DF3E463F2A751FB85B946DC05" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="cuatrecasasii">
<emphasis id="37AFE5F794C6E028AB28D1CF968F9B0A" italics="true">C. cuatrecasasii</emphasis>
</taxonomicName>
: oblong or elliptic-oblong leaves and slightly brochidodromous secondary venation.
<taxonomicName id="2EA233849725888FBB31640E85E49E4A" authorityName="Villanueva, Paz-Lopez &amp; W. Ariza" authorityYear="2025" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="crassitepala">
<emphasis id="86AFBA44A063337254676A780AB425B3" italics="true">C. crassitepala</emphasis>
</taxonomicName>
differs from
<taxonomicName id="8B6DF50B30792CCC78F362C790FCEE22" class="Magnoliopsida" family="Myristicaceae" genus="Compsoneura" kingdom="Plantae" order="Magnoliales" phylum="Tracheophyta" rank="species" species="cuatrecasasii">
<emphasis id="481CD50D50A91A878AA4E897DD15A2AF" italics="true">C. cuatrecasasii</emphasis>
</taxonomicName>
by the leaf blades that dry yellowish-brown and have an abaxial lamina densely ferruginous tomentose with dendritic trichomes (vs. leaf blades that dry red-brown and are glabrescent to glabrous when mature on the abaxial lamina with minute 2 - branched trichomes). Additionally, after the revision of herbarium specimens, based on morphological traits such as leaf blade veins and size, perianth lobes and pericarp texture and following the process of
<bibRefCitation id="AAD60EAD97A83AC1397C4372E96798B7" DOI="10.11646/phytotaxa.637.3.3" author="Villanueva-Tamayo B &amp; Cogollo-Pacheco A" firstAuthor="Villanueva-Tamayo" issue="3" journalOrPublisher="Phytotaxa" pagination="251-260" refId="B26" refString="Villanueva-Tamayo B, Cogollo-Pacheco A (2024) A new species of Compsoneura (Myristicaceae) from interandean valleys of middle Magdalena and lower Cauca rivers, Colombia. Phytotaxa 637 (3): 251–260. https://doi.org/10.11646/phytotaxa.637.3.3" title="A new species of Compsoneura (Myristicaceae) from interandean valleys of middle Magdalena and lower Cauca rivers, Colombia." volume="637" year="2024">Villanueva-Tamayo and Cogollo-Pacheco (2024)</bibRefCitation>
, an identification key for the species of the genus is provided.
</paragraph>
</subSubSection>
</treatment>
</document>