Add updates up until 2025-01-20 15:59:22

This commit is contained in:
ggserver 2025-01-20 16:01:33 +00:00
parent 50112d6d96
commit 379825a3b3
73 changed files with 4348 additions and 4276 deletions

View file

@ -1,65 +1,66 @@
<document id="9B3343138F2B0CD23616218C23138741" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFA0FFEC62EBFD6BFBC8F905" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa berlandi Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov" docType="treatment" docVersion="1" lastPageNumber="239" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="231" updateTime="1737369942339" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
<mods:mods id="129FA99327C4E4834C652FF3C7793525" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3D236C77E84616DEA9E62207A1E9420D">
<mods:title id="3FD21F425D09F690E96FCCB7B8C5DD08">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
<document id="022074F8C156DB47028050A8EFB15F91" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14702734" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFA0FFEC62EBFD6BFBC8F905" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa berlandi Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov" docType="treatment" docVersion="2" lastPageNumber="239" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="231" updateTime="1737387624970" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="900C9344FA37040B57AE6B50DFACBD28" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B68CC011BED3EC3956F970E9DC50B780">
<mods:title id="D86F0DD5EC9122A977EF88C85E7C40A4">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
</mods:titleInfo>
<mods:name id="7C4BE13D04808963E2D816A98614F48C" type="personal">
<mods:role id="480343A311BE7A74BD7DF2C537AC4077">
<mods:roleTerm id="3301FDDAAD6E43DE7893661AB59FAA19">Author</mods:roleTerm>
<mods:name id="F890C103E3C651DBCAA3FE0390B9688E" type="personal">
<mods:role id="8489CC7E3F8020D15B34195ABB0590BE">
<mods:roleTerm id="80D8AFC704ECF55683272F92279402AD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0FF0522958649143264DF830E2287159">Martínez, Leonel</mods:namePart>
<mods:affiliation id="978028989523D20102F5040DEC9A774F">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
<mods:namePart id="D0202333EA9AAF43146680D24C51F46F">Martínez, Leonel</mods:namePart>
<mods:affiliation id="D98BB9D8570B45769107A13D2CC775DA">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
</mods:name>
<mods:name id="F17830D988064EFDC57D1F6F7F1307F1" type="personal">
<mods:role id="A608CF70CFDEDD4A0325351A04F555AE">
<mods:roleTerm id="7A2D357C7339741A443DFE2C466A9B0B">Author</mods:roleTerm>
<mods:name id="82B42C19802FCA711757225B1608F1D0" type="personal">
<mods:role id="E998B0FFE7AA63E565C7DC198CBE0888">
<mods:roleTerm id="8B6AA7C04A142D985ECE8367838788E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8236C5D060A381BC74273DCF1B717F9F">Kochalka, John A.</mods:namePart>
<mods:affiliation id="45BCD542AA8CD8A3AB0A81D40738EF0C">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
<mods:namePart id="8D5ED62CF97BD569C0CF33C6B6333834">Kochalka, John A.</mods:namePart>
<mods:affiliation id="ACB4ABD985E6FB695A8E3044549CE91F">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
</mods:name>
<mods:name id="28A6E75A59B872A0EBF731F222C209C6" type="personal">
<mods:role id="6D06C3D56B4F15E2566366CDDC2338CC">
<mods:roleTerm id="09D9249451FF1C5F762958F3B5CBB6A7">Author</mods:roleTerm>
<mods:name id="F39F4D4788A6063DE0D767C0FE1A1B43" type="personal">
<mods:role id="B0CEA55D61A0294EAE5A376C6CE0FEF2">
<mods:roleTerm id="57B55EEDB7F5CF7922D37DE916B1413E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="431046ED1BC53BA9D192376B5FA21146">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="675B5E8E304030739105DC9B3A0541A0">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
<mods:namePart id="DFE87327999F7A3A0B0F508076BE75F0">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="F208ED2827ED6A9177668B3EAAB14E56">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
</mods:name>
<mods:name id="58A5C34775BD9E272065279090CEF71D" type="personal">
<mods:role id="8C31B67C8D4A4866F631B6EBA3049E3C">
<mods:roleTerm id="FFCCC81BC087095240FD6209D9DF7AF7">Author</mods:roleTerm>
<mods:name id="B12D9E08D9061612AAAF04028B150CED" type="personal">
<mods:role id="37CDC01978C07FE557D55FC96DAECF4C">
<mods:roleTerm id="EF79F3423D890AA8387F68B94E8258EA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="398936432E018CEA17D814286828E8D2">Ramírez, Martin</mods:namePart>
<mods:affiliation id="C7F41E06BB52E6ECEE9A9C95E94873A6">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
<mods:namePart id="C3048F9278293D6A23854AC2328A0299">Ramírez, Martin</mods:namePart>
<mods:affiliation id="CADE06879F97434B2E1C4BD0355353CB">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E0B0F6F8564D8E007B599E297D66057F">text</mods:typeOfResource>
<mods:relatedItem id="ABF09E6218EF2413EE6207A94E7CD72E" type="host">
<mods:titleInfo id="83482A5AF5EBB9385A7965CB60432A6C">
<mods:title id="EAAED817ADF4E3696F41EEE60B531595">Zootaxa</mods:title>
<mods:typeOfResource id="7BBFEEA9BC2EA25F0EF36BE7D91B93FF">text</mods:typeOfResource>
<mods:relatedItem id="8FEC60627FF0C3ECEEA3D8A6DF1453EF" type="host">
<mods:titleInfo id="DBAA7E40ABBD6D200D3B470423EC875C">
<mods:title id="FDBA5E13BB9234D628CA154FBAC5498D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="52C6AC37F0281B0234175FA489E7D271">
<mods:date id="C3B8C6DDB24448D245B59F3A8B12128C">2025</mods:date>
<mods:detail id="683500CBFF7758153567A8E4D75CD507" type="pubDate">
<mods:number id="907321F4B700EBC10CB267C5B01A1AF2">2025-01-08</mods:number>
<mods:part id="64D80B7A363A58055D1A8EB2C728CFDD">
<mods:date id="73B507E6671A0BF64912C8D264B95EF3">2025</mods:date>
<mods:detail id="A4D17F29BAE79257A2785B4B41A35080" type="pubDate">
<mods:number id="703362FC727EE0C104EDA0116F0E5893">2025-01-08</mods:number>
</mods:detail>
<mods:detail id="2D35E512F5F50BE6A4AE0967B8EDA08F" type="volume">
<mods:number id="F32B075031D5BD1D747BCA64E93D731C">5566</mods:number>
<mods:detail id="5CEAC333E130A25DB576430C510C42EC" type="volume">
<mods:number id="18C9037FF62DB7A247EBA66A85D451DD">5566</mods:number>
</mods:detail>
<mods:detail id="4F49D60B5BC8F939B685680739AAD3EB" type="issue">
<mods:number id="14D80837BEB06DBB488C30F57FB05B00">2</mods:number>
<mods:detail id="A8DA02E562E51D7590020877123F4198" type="issue">
<mods:number id="C566E45B4A9552060BF6670D1FCFB3FC">2</mods:number>
</mods:detail>
<mods:extent id="565F33F5AC69B9C1769E26153010B46F" unit="page">
<mods:start id="2AAFDC5778DDA267F57CBC558EA8A52C">201</mods:start>
<mods:end id="37A5E678B3CED9D7AD0E970B28D63D20">242</mods:end>
<mods:extent id="3C9BEB38094AE34B02E8D3D71EC0724D" unit="page">
<mods:start id="C730C9574B0DCFF4E5804C022CE58359">201</mods:start>
<mods:end id="1286D0185E16C91D1B6E2D0D38568DF9">242</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="BC7740490C7C828397EB020355953D92">
<mods:url id="7D9BA6CACC798D40E9180977DCDCC452">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
<mods:location id="E429B0B5652130404CC70FE7732A60FB">
<mods:url id="22BADB8BD153E21262722A7E53C9C2D4">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
</mods:location>
<mods:classification id="649BC48F61676E3EDD73CD8BD6073D2B">journal article</mods:classification>
<mods:identifier id="DE6A54C2FF696A131A84160F4340748B" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="030792040077E95B409A3A90EE346768" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5985FBE246F6D6A1626F2A05909415C7" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
<mods:classification id="4095193B55437C4FED0E57618B8FDCB6">journal article</mods:classification>
<mods:identifier id="FCC6ABDE551CD1AC5E0A8A4013C8ADDD" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="AF2221ADB1E0AC6B82E4C73FAF40F532" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5C9005BBB04AD547FEADAA7D04F2E8A5" type="Zenodo-Dep">14702734</mods:identifier>
<mods:identifier id="A0DAC6C7A5DA91B4126B2C50C695E2C2" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
</mods:mods>
<treatment id="4D76240CFFA0FFEC62EBFD6BFBC8F905" LSID="urn:lsid:plazi:treatment:4D76240CFFA0FFEC62EBFD6BFBC8F905" httpUri="http://treatment.plazi.org/id/4D76240CFFA0FFEC62EBFD6BFBC8F905" lastPageId="38" lastPageNumber="239" pageId="30" pageNumber="231">
<subSubSection id="8DC5C691FFA0FFD462EBFD6BFE3FFD71" box="[151,411,654,681]" pageId="30" pageNumber="231" type="nomenclature">
@ -76,7 +77,7 @@
</subSubSection>
<subSubSection id="8DC5C691FFA0FFD462EBFD57FE9CFD15" box="[151,312,691,717]" pageId="30" pageNumber="231" type="description">
<paragraph id="C560951AFFA0FFD462EBFD57FE9CFD15" blockId="30.[151,411,654,717]" box="[151,312,691,717]" pageId="30" pageNumber="231">
<figureCitation id="5DE4899FFFA0FFD462EBFD57FE9CFD15" box="[151,312,691,717]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStart-2="FIGURE 25" captionStart-3="FIGURE 26" captionStart-4="FIGURE 27" captionStart-5="FIGURE 28" captionStart-6="FIGURE 29" captionStartId-0="31.[151,250,1883,1907]" captionStartId-1="32.[151,250,1879,1903]" captionStartId-2="33.[151,250,1836,1860]" captionStartId-3="34.[151,250,1866,1890]" captionStartId-4="35.[151,250,1779,1803]" captionStartId-5="36.[151,250,1731,1755]" captionStartId-6="37.[151,250,1517,1541]" captionTargetBox-0="[159,1426,193,1860]" captionTargetBox-1="[159,1425,181,1853]" captionTargetBox-2="[151,1436,212,1812]" captionTargetBox-3="[158,1427,185,1840]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetBox-6="[163,1429,193,1485]" captionTargetId-0="figure-17@31.[151,1436,181,1860]" captionTargetId-1="figure-22@32.[151,1436,181,1855]" captionTargetId-2="figure-17@33.[151,1436,181,1812]" captionTargetId-3="figure-22@34.[151,1435,181,1842]" captionTargetId-4="figure-17@35.[151,1435,181,1756]" captionTargetId-5="figure-22@36.[151,1435,181,1708]" captionTargetId-6="figure-17@37.[151,1436,181,1493]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionTargetPageId-5="36" captionTargetPageId-6="37" captionText-0="FIGURE 23. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 24. Josa berlandi sp. nov., female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-2="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." captionText-3="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." captionText-5="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." captionText-6="FIGURE 29. Geographical records of spiders in South America above 3000 masl, classified by altitude intervals." pageId="30" pageNumber="231">Figures 2329</figureCitation>
<figureCitation id="5DE4899FFFA0FFD462EBFD57FE9CFD15" box="[151,312,691,717]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStart-2="FIGURE 25" captionStart-3="FIGURE 26" captionStart-4="FIGURE 27" captionStart-5="FIGURE 28" captionStart-6="FIGURE 29" captionStartId-0="31.[151,250,1883,1907]" captionStartId-1="32.[151,250,1879,1903]" captionStartId-2="33.[151,250,1836,1860]" captionStartId-3="34.[151,250,1866,1890]" captionStartId-4="35.[151,250,1779,1803]" captionStartId-5="36.[151,250,1731,1755]" captionStartId-6="37.[151,250,1517,1541]" captionTargetBox-0="[159,1426,193,1860]" captionTargetBox-1="[159,1425,181,1853]" captionTargetBox-2="[151,1436,212,1812]" captionTargetBox-3="[158,1427,185,1840]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetBox-6="[163,1429,193,1485]" captionTargetId-0="figure-17@31.[151,1436,181,1860]" captionTargetId-1="figure-22@32.[151,1436,181,1855]" captionTargetId-2="figure-17@33.[151,1436,181,1812]" captionTargetId-3="figure-22@34.[151,1435,181,1842]" captionTargetId-4="figure-17@35.[151,1435,181,1756]" captionTargetId-5="figure-22@36.[151,1435,181,1708]" captionTargetId-6="figure-17@37.[151,1436,181,1493]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionTargetPageId-5="36" captionTargetPageId-6="37" captionText-0="FIGURE 23. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 24. Josa berlandi sp. nov., female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-2="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." captionText-3="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." captionText-5="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." captionText-6="FIGURE 29. Geographical records of spiders in South America above 3000 masl, classified by altitude intervals." figureDoi-0="http://doi.org/10.5281/zenodo.14702815" figureDoi-1="http://doi.org/10.5281/zenodo.14702821" figureDoi-2="http://doi.org/10.5281/zenodo.14702825" figureDoi-3="http://doi.org/10.5281/zenodo.14702829" figureDoi-4="http://doi.org/10.5281/zenodo.14702831" figureDoi-5="http://doi.org/10.5281/zenodo.14702833" figureDoi-6="http://doi.org/10.5281/zenodo.14702837" httpUri-0="https://zenodo.org/record/14702815/files/figure.png" httpUri-1="https://zenodo.org/record/14702821/files/figure.png" httpUri-2="https://zenodo.org/record/14702825/files/figure.png" httpUri-3="https://zenodo.org/record/14702829/files/figure.png" httpUri-4="https://zenodo.org/record/14702831/files/figure.png" httpUri-5="https://zenodo.org/record/14702833/files/figure.png" httpUri-6="https://zenodo.org/record/14702837/files/figure.png" pageId="30" pageNumber="231">Figures 2329</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA0FFD462EBFD10FB39FC1B" pageId="30" pageNumber="231" type="materials_examined">
@ -171,21 +172,21 @@ The specific epithet is a patronymic in honor of the late French arachnologist L
<paragraph id="C560951AFFA0FFD462BBFBF0FC36FB3B" blockId="30.[151,1437,756,2043]" pageId="30" pageNumber="231">
<emphasis id="F7AB4908FFA0FFD462BBFBF0FEE6FBF6" bold="true" box="[199,322,1044,1070]" pageId="30" pageNumber="231">Diagnosis.</emphasis>
Males are easily distinguished from other species by the bifid external section of the conductor, with beak-shaped anterior tip, longer than the posterior one, which is sharp and thin, elongated subapical protrusion (
<figureCitation id="5DE4899FFFA0FFD46716FBDCFF6EFBAF" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46716FBDCFF6EFBAF" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="30" pageNumber="231">Figs 25A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD462ABFBB8FEAEFBAE" box="[215,266,1116,1142]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD462ABFBB8FEAEFBAE" box="[215,266,1116,1142]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="30" pageNumber="231">26A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD4636BFBB8FE2BFBAE" box="[279,399,1116,1142]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27AB, D</figureCitation>
<figureCitation id="5DE4899FFFA0FFD4636BFBB8FE2BFBAE" box="[279,399,1116,1142]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">27AB, D</figureCitation>
), median apophysis bifid, laminar, projected, with the anterior tip longer than the posterior one (
<figureCitation id="5DE4899FFFA0FFD462AEFB64FED9FB42" box="[210,381,1152,1179]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">Fig. 27B, F, H</figureCitation>
<figureCitation id="5DE4899FFFA0FFD462AEFB64FED9FB42" box="[210,381,1152,1179]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">Fig. 27B, F, H</figureCitation>
), tegulum prolaterobasally projected (
<figureCitation id="5DE4899FFFA0FFD4614DFB64FC3BFB43" box="[817,927,1152,1179]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD4614DFB64FC3BFB43" box="[817,927,1152,1179]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="30" pageNumber="231">Figs 25A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD461D1FB64FC45FB42" box="[941,993,1152,1178]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD461D1FB64FC45FB42" box="[941,993,1152,1178]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="30" pageNumber="231">26A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46192FB64FB87FB42" box="[1006,1059,1152,1178]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46192FB64FB87FB42" box="[1006,1059,1152,1178]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">27A</figureCitation>
). Females are distinguished by a bilobed median field folded toward the median notch, wide lateral lobes with basally rounded posterior sclerotized projections, and an arch-shaped narrow median notch (
<figureCitation id="5DE4899FFFA0FFD46082FB2CFC22FB3B" box="[766,902,1224,1251]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25EF</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46082FB2CFC22FB3B" box="[766,902,1224,1251]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="30" pageNumber="231">Figs 25EF</figureCitation>
).
</paragraph>
</subSubSection>
@ -196,36 +197,36 @@ Males are easily distinguished from other species by the bifid external section
(
<typeStatus id="1A642BB8FFA0FFD463CFFB08FDBDFADE" box="[435,537,1260,1286]" pageId="30" pageNumber="231" type="holotype">holotype</typeStatus>
, MUSENUV-Ar-2675; vchLAM-253) (
<figureCitation id="5DE4899FFFA0FFD46199FB08FB9BFADE" box="[997,1087,1260,1286]" captionStart="FIGURE 23" captionStartId="31.[151,250,1883,1907]" captionTargetBox="[159,1426,193,1860]" captionTargetId="figure-17@31.[151,1436,181,1860]" captionTargetPageId="31" captionText="FIGURE 23. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="30" pageNumber="231">Fig. 23</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46199FB08FB9BFADE" box="[997,1087,1260,1286]" captionStart="FIGURE 23" captionStartId="31.[151,250,1883,1907]" captionTargetBox="[159,1426,193,1860]" captionTargetId="figure-17@31.[151,1436,181,1860]" captionTargetPageId="31" captionText="FIGURE 23. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." figureDoi="http://doi.org/10.5281/zenodo.14702815" httpUri="https://zenodo.org/record/14702815/files/figure.png" pageId="30" pageNumber="231">Fig. 23</figureCitation>
): Total length 4.18. Carapace length 2.35, width 1.73, height 0.89. Clypeus length 0.15. Sternum length 1.20, width 0.83. Legs. Leg I: ti length 1.79/width 0.31, mt 1.45. II: ti 1.72/0.28, mt 1.43. III: ti 1.35/0.24, mt 1.35. IV: ti 1.71/0.30, mt 1.73. Spines: legs III, femur d 1-1-1, p 2ap, r 0-d1-d1; tibia v p1-2-2, p 1-d1-1-0, r 1-d1-1-0; metatarsus v 2-2-0, p d1-0. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 0-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1- 2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. IV=III but tibia v 2-2-2, metatarsus v 2-2-2. Abdomen: length 1.92, width 1.01, spiracleepigastrium 0.67. Color: carapace dark brown, with cervical groves spotted in black and converging in the large black patch on fovea. Lateral region with two poorly defined dark bands on sides, covered by abundant white setae. Ocular area on a black patch, eyes bordered in black rings. Clypeus brown, low with abundant white setae. Chelicerae brown with black stripes and fine setae on frontal edge. Endites, labium, and sternum brown with some dark spots; labium more pigmented. Legs coxae-tarsi brown, femora with dark longitudinal bands, tibia-metatarsi with rings of white setae; tarsi uniformly brown. Articles with abundant white setae. Abdomen dehydrated, with a pattern of faint light spots on a dark gray background, covered by abundant white setae. Venter uniformly dark gray. Palp: tegulum with very developed median hematodochae (
<figureCitation id="5DE4899FFFA0FFD4615EF924FC28F903" box="[802,908,1728,1755]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD4615EF924FC28F903" box="[802,908,1728,1755]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="30" pageNumber="231">Figs 25A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD461E9F924FC6CF902" box="[917,968,1728,1754]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD461E9F924FC6CF902" box="[917,968,1728,1754]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="30" pageNumber="231">26A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD461ADF924FBA2F902" box="[977,1030,1728,1754]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A</figureCitation>
<figureCitation id="5DE4899FFFA0FFD461ADF924FBA2F902" box="[977,1030,1728,1754]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">27A</figureCitation>
). Median apophysis very large, bifid, with anterior branch long, laminar, with projected prolateral tip, posterior branch membranous at base and with the external border sub-rounded (
<figureCitation id="5DE4899FFFA0FFD46399F8ECFDD6F8FB" box="[485,626,1800,1827]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25AB</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46399F8ECFDD6F8FB" box="[485,626,1800,1827]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="30" pageNumber="231">Figs 25AB</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46002F8ECFD76F8FA" box="[638,722,1800,1826]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26AC</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46002F8ECFD76F8FA" box="[638,722,1800,1826]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="30" pageNumber="231">26AC</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD460A1F8ECFCD7F8FA" box="[733,883,1800,1827]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A, B, D, F</figureCitation>
<figureCitation id="5DE4899FFFA0FFD460A1F8ECFCD7F8FA" box="[733,883,1800,1827]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">27A, B, D, F</figureCitation>
). Conductor small, rounded, with projected base, external section apically wide and bifid, with a swollen, elongated projection; anterior tip of external section large, curved, and longer than posterior; posterior tip short and sharp; internal section ending as a sharp tip, converging with the posterior tip of external section (
<figureCitation id="5DE4899FFFA0FFD4600EF890FCA5F857" box="[626,769,1908,1935]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25AB</figureCitation>
<figureCitation id="5DE4899FFFA0FFD4600EF890FCA5F857" box="[626,769,1908,1935]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="30" pageNumber="231">Figs 25AB</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46172F890FCC6F856" box="[782,866,1908,1934]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26AC</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46172F890FCC6F856" box="[782,866,1908,1934]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="30" pageNumber="231">26AC</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46112F890FBACF856" box="[878,1032,1908,1935]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A, B, D, F</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46112F890FBACF856" box="[878,1032,1908,1935]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">27A, B, D, F</figureCitation>
). Paramedian apophysis large, cup-shaped, heavily sclerotized, concave, apically acuminated (
<figureCitation id="5DE4899FFFA0FFD46156F87CFC08F86A" box="[810,940,1944,1970]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">Fig. 27C, E</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46156F87CFC08F86A" box="[810,940,1944,1970]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">Fig. 27C, E</figureCitation>
) Embolus short, tape-shaped at base, coiled, with poorly developed laminar projection, apically filiform; basal process small, flat, retrolateral region pointed in posterior region, prolateral region oval, poorly pronounced external keel (
<figureCitation id="5DE4899FFFA0FFD461ACF804FBFBF823" box="[976,1119,2016,2043]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25AD</figureCitation>
<figureCitation id="5DE4899FFFA0FFD461ACF804FBFBF823" box="[976,1119,2016,2043]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="30" pageNumber="231">Figs 25AD</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46615F804FB1AF822" box="[1129,1214,2016,2042]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26AC</figureCitation>
<figureCitation id="5DE4899FFFA0FFD46615F804FB1AF822" box="[1129,1214,2016,2042]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="30" pageNumber="231">26AC</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD466B4F804FB4CF822" box="[1224,1256,2016,2042]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27</figureCitation>
<figureCitation id="5DE4899FFFA0FFD466B4F804FB4CF822" box="[1224,1256,2016,2042]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="30" pageNumber="231">27</figureCitation>
).
</paragraph>
<caption id="91A0C592FFA1FFD562EBF8BFFDECF840" pageId="31" pageNumber="232" startId="31.[151,250,1883,1907]" targetBox="[159,1426,193,1860]" targetPageId="31" targetType="figure">
<caption id="91A0C592FFA1FFD562EBF8BFFDECF840" ID-DOI="http://doi.org/10.5281/zenodo.14702815" ID-Zenodo-Dep="14702815" httpUri="https://zenodo.org/record/14702815/files/figure.png" pageId="31" pageNumber="232" startId="31.[151,250,1883,1907]" targetBox="[159,1426,193,1860]" targetPageId="31" targetType="figure">
<paragraph id="C560951AFFA1FFD562EBF8BFFDECF840" blockId="31.[151,1436,1883,1944]" pageId="31" pageNumber="232">
<emphasis id="F7AB4908FFA1FFD562EBF8BFFEBAF8AB" bold="true" box="[151,286,1883,1907]" pageId="31" pageNumber="232">FIGURE 23.</emphasis>
<taxonomicName id="02DFEE99FFA1FFD5635FF8B8FE08F8AC" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[291,428,1884,1908]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
@ -237,7 +238,7 @@ Males are easily distinguished from other species by the bifid external section
, male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FF9EFFEA62EBF8B3FD3CF84C" pageId="32" pageNumber="233" startId="32.[151,250,1879,1903]" targetBox="[159,1425,181,1853]" targetPageId="32" targetType="figure">
<caption id="91A0C592FF9EFFEA62EBF8B3FD3CF84C" ID-DOI="http://doi.org/10.5281/zenodo.14702821" ID-Zenodo-Dep="14702821" httpUri="https://zenodo.org/record/14702821/files/figure.png" pageId="32" pageNumber="233" startId="32.[151,250,1879,1903]" targetBox="[159,1425,181,1853]" targetPageId="32" targetType="figure">
<paragraph id="C560951AFF9EFFEA62EBF8B3FD3CF84C" blockId="32.[151,1436,1879,1940]" pageId="32" pageNumber="233">
<emphasis id="F7AB4908FF9EFFEA62EBF8B3FE86F8B7" bold="true" box="[151,290,1879,1903]" pageId="32" pageNumber="233">FIGURE 24.</emphasis>
<taxonomicName id="02DFEE99FF9EFFEA6357F8BCFE1CF8B7" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,440,1879,1903]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
@ -249,7 +250,7 @@ Males are easily distinguished from other species by the bifid external section
, female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FF9FFFEB62EBF8C8FBB1F80C" pageId="33" pageNumber="234" startId="33.[151,250,1836,1860]" targetBox="[151,1436,212,1812]" targetPageId="33" targetType="figure">
<caption id="91A0C592FF9FFFEB62EBF8C8FBB1F80C" ID-DOI="http://doi.org/10.5281/zenodo.14702825" ID-Zenodo-Dep="14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="33" pageNumber="234" startId="33.[151,250,1836,1860]" targetBox="[151,1436,212,1812]" targetPageId="33" targetType="figure">
<paragraph id="C560951AFF9FFFEB62EBF8C8FBB1F80C" blockId="33.[151,1436,1836,2005]" pageId="33" pageNumber="234">
<emphasis id="F7AB4908FF9FFFEB62EBF8C8FE80F89C" bold="true" box="[151,292,1836,1860]" pageId="33" pageNumber="234">FIGURE 25.</emphasis>
<taxonomicName id="02DFEE99FF9FFFEB6352F8C9FE19F89C" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[302,445,1836,1860]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
@ -263,7 +264,7 @@ Males are easily distinguished from other species by the bifid external section
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FF9CFFE862EBF8AEFC9DF82A" pageId="34" pageNumber="235" startId="34.[151,250,1866,1890]" targetBox="[158,1427,185,1840]" targetPageId="34" targetType="figure">
<caption id="91A0C592FF9CFFE862EBF8AEFC9DF82A" ID-DOI="http://doi.org/10.5281/zenodo.14702829" ID-Zenodo-Dep="14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="34" pageNumber="235" startId="34.[151,250,1866,1890]" targetBox="[158,1427,185,1840]" targetPageId="34" targetType="figure">
<paragraph id="C560951AFF9CFFE862EBF8AEFC9DF82A" blockId="34.[151,1437,1866,2034]" pageId="34" pageNumber="235">
<emphasis id="F7AB4908FF9CFFE862EBF8AEFE80F8BA" bold="true" box="[151,292,1866,1890]" pageId="34" pageNumber="235">FIGURE 26.</emphasis>
<taxonomicName id="02DFEE99FF9CFFE86352F8AFFE19F8BA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[302,445,1866,1890]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
@ -277,7 +278,7 @@ AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct.
</paragraph>
</caption>
<caption id="91A0C592FF9DFFE962EBF917FC32F844" pageId="35" pageNumber="236" startId="35.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="35" targetType="figure">
<caption id="91A0C592FF9DFFE962EBF917FC32F844" ID-DOI="http://doi.org/10.5281/zenodo.14702831" ID-Zenodo-Dep="14702831" httpUri="https://zenodo.org/record/14702831/files/figure.png" pageId="35" pageNumber="236" startId="35.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="35" targetType="figure">
<paragraph id="C560951AFF9DFFE962EBF917FC32F844" blockId="35.[151,1436,1779,1948]" pageId="35" pageNumber="236">
<emphasis id="F7AB4908FF9DFFE962EBF917FE87F8D4" bold="true" box="[151,291,1779,1804]" pageId="35" pageNumber="236">FIGURE 27.</emphasis>
<taxonomicName id="02DFEE99FF9DFFE96357F910FE1CF8D4" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,440,1780,1804]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
@ -291,7 +292,7 @@ AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; d
C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FF9AFFEE62EBF927FAFDF8FC" pageId="36" pageNumber="237" startId="36.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="36" targetType="figure">
<caption id="91A0C592FF9AFFEE62EBF927FAFDF8FC" ID-DOI="http://doi.org/10.5281/zenodo.14702833" ID-Zenodo-Dep="14702833" httpUri="https://zenodo.org/record/14702833/files/figure.png" pageId="36" pageNumber="237" startId="36.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="36" targetType="figure">
<paragraph id="C560951AFF9AFFEE62EBF927FAFDF8FC" blockId="36.[151,1436,1731,1828]" pageId="36" pageNumber="237">
<emphasis id="F7AB4908FF9AFFEE62EBF927FE85F904" bold="true" box="[151,289,1731,1756]" pageId="36" pageNumber="237">FIGURE 28.</emphasis>
<taxonomicName id="02DFEE99FF9AFFEE635BF920FE17F904" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[295,435,1732,1756]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
@ -305,7 +306,7 @@ C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus;
AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca.
</paragraph>
</caption>
<caption id="91A0C592FF9BFFEF62EBFA09FAB2F9DD" box="[151,1302,1517,1541]" pageId="37" pageNumber="238" startId="37.[151,250,1517,1541]" targetBox="[163,1429,193,1485]" targetPageId="37" targetType="figure">
<caption id="91A0C592FF9BFFEF62EBFA09FAB2F9DD" ID-DOI="http://doi.org/10.5281/zenodo.14702837" ID-Zenodo-Dep="14702837" box="[151,1302,1517,1541]" httpUri="https://zenodo.org/record/14702837/files/figure.png" pageId="37" pageNumber="238" startId="37.[151,250,1517,1541]" targetBox="[163,1429,193,1485]" targetPageId="37" targetType="figure">
<paragraph id="C560951AFF9BFFEF62EBFA09FAB2F9DD" blockId="37.[151,1302,1517,1541]" box="[151,1302,1517,1541]" pageId="37" pageNumber="238">
<emphasis id="F7AB4908FF9BFFEF62EBFA09FE84F9DD" bold="true" box="[151,288,1517,1541]" pageId="37" pageNumber="238">FIGURE 29.</emphasis>
Geographical records of spiders in South America above 3000 masl, classified by altitude intervals.
@ -316,28 +317,28 @@ Geographical records of spiders in South America above 3000 masl, classified by
(
<typeStatus id="1A642BB8FF9BFFEF6351F9B5FE36F9B3" box="[301,402,1617,1643]" pageId="37" pageNumber="238" type="paratype">paratype</typeStatus>
, MUSENUV-Ar-2676; vchLAM-255). (
<figureCitation id="5DE4899FFF9BFFEF6122F9B4FC12F9B3" box="[862,950,1616,1643]" captionStart="FIGURE 24" captionStartId="32.[151,250,1879,1903]" captionTargetBox="[159,1425,181,1853]" captionTargetId="figure-22@32.[151,1436,181,1855]" captionTargetPageId="32" captionText="FIGURE 24. Josa berlandi sp. nov., female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="37" pageNumber="238">Fig. 24</figureCitation>
<figureCitation id="5DE4899FFF9BFFEF6122F9B4FC12F9B3" box="[862,950,1616,1643]" captionStart="FIGURE 24" captionStartId="32.[151,250,1879,1903]" captionTargetBox="[159,1425,181,1853]" captionTargetId="figure-22@32.[151,1436,181,1855]" captionTargetPageId="32" captionText="FIGURE 24. Josa berlandi sp. nov., female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." figureDoi="http://doi.org/10.5281/zenodo.14702821" httpUri="https://zenodo.org/record/14702821/files/figure.png" pageId="37" pageNumber="238">Fig. 24</figureCitation>
): Total length 4.09. Carapace length 1.73, width 1.31, height 0.67. Clypeus length 0.08. Sternum length 1.06, width 0.71. Legs. Leg I: ti length 0.97/width 0.23, mt 0.70. II: ti 0.90/0.21, mt 0.69. III: ti 0.71/0.21, mt 0.66. IV: ti 1.12/0.22, mt 1,01. Spines: legs III, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v p1- 2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 2.53, width 1.54, spiracleepigastrium 0.80. Color pattern as in male, but dark brown, abdominal and legs spots more defined. Sternum and labium darker. Epigyne: anterior plate sclerotized with accentuated anterior excavations, median excavation long and narrow (
<figureCitation id="5DE4899FFF9BFFEF6036F8A8FD13F8BF" box="[586,695,1868,1895]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="37" pageNumber="238">Figs 25E</figureCitation>
<figureCitation id="5DE4899FFF9BFFEF6036F8A8FD13F8BF" box="[586,695,1868,1895]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="37" pageNumber="238">Figs 25E</figureCitation>
,
<figureCitation id="5DE4899FFF9BFFEF60BAF8A8FD5EF8BE" box="[710,762,1868,1894]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="37" pageNumber="238">26D</figureCitation>
<figureCitation id="5DE4899FFF9BFFEF60BAF8A8FD5EF8BE" box="[710,762,1868,1894]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="37" pageNumber="238">26D</figureCitation>
). Median field totally divided in two sub-rounded section interrupted by a large medial depression (
<figureCitation id="5DE4899FFF9BFFEF6002F894FCA0F853" box="[638,772,1904,1931]" captionStart="FIGURE 28" captionStartId="36.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-22@36.[151,1435,181,1708]" captionTargetPageId="36" captionText="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="37" pageNumber="238">
<figureCitation id="5DE4899FFF9BFFEF6002F894FCA0F853" box="[638,772,1904,1931]" captionStart="FIGURE 28" captionStartId="36.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-22@36.[151,1435,181,1708]" captionTargetPageId="36" captionText="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702833" httpUri="https://zenodo.org/record/14702833/files/figure.png" pageId="37" pageNumber="238">
Fig. 28E
<emphasis id="F7AB4908FF9BFFEF609FF895FD55F853" bold="true" box="[739,753,1905,1931]" pageId="37" pageNumber="238"></emphasis>
F
</figureCitation>
). Lateral lobes with sclerotized extensions wide, swollen and apically rounded (
<figureCitation id="5DE4899FFF9BFFEF63E0F870FDA1F877" box="[412,517,1940,1967]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="37" pageNumber="238">Fig. 25E</figureCitation>
<figureCitation id="5DE4899FFF9BFFEF63E0F870FDA1F877" box="[412,517,1940,1967]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="37" pageNumber="238">Fig. 25E</figureCitation>
). Copulatory ducts short joining with the accessory bulb ducts at medial level. Spermathecae rounded in median position oriented towards lateral region. Ducts of accessory bulbs long, thin, convergent (
<figureCitation id="5DE4899FFF9BFFEF635EF838FE02F82F" box="[290,422,2012,2039]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="37" pageNumber="238">Figs 25EF</figureCitation>
<figureCitation id="5DE4899FFF9BFFEF635EF838FE02F82F" box="[290,422,2012,2039]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702825" httpUri="https://zenodo.org/record/14702825/files/figure.png" pageId="37" pageNumber="238">Figs 25EF</figureCitation>
,
<figureCitation id="5DE4899FFF9BFFEF63CDF838FE59F82E" box="[433,509,2012,2038]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="37" pageNumber="238">26EF</figureCitation>
<figureCitation id="5DE4899FFF9BFFEF63CDF838FE59F82E" box="[433,509,2012,2038]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702829" httpUri="https://zenodo.org/record/14702829/files/figure.png" pageId="37" pageNumber="238">26EF</figureCitation>
,
<figureCitation id="5DE4899FFF9BFFEF6074F839FD8CF82F" box="[520,552,2013,2039]" captionStart="FIGURE 28" captionStartId="36.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-22@36.[151,1435,181,1708]" captionTargetPageId="36" captionText="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="37" pageNumber="238">28</figureCitation>
<figureCitation id="5DE4899FFF9BFFEF6074F839FD8CF82F" box="[520,552,2013,2039]" captionStart="FIGURE 28" captionStartId="36.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-22@36.[151,1435,181,1708]" captionTargetPageId="36" captionText="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702833" httpUri="https://zenodo.org/record/14702833/files/figure.png" pageId="37" pageNumber="238">28</figureCitation>
).
</paragraph>
<caption id="91A0C592FF98FFEC62EBFA09FAFBF9DD" box="[151,1375,1517,1541]" pageId="38" pageNumber="239" startId="38.[151,250,1517,1541]" targetBox="[163,1429,193,1485]" targetPageId="38" targetType="figure">
<caption id="91A0C592FF98FFEC62EBFA09FAFBF9DD" ID-DOI="http://doi.org/10.5281/zenodo.14702841" ID-Zenodo-Dep="14702841" box="[151,1375,1517,1541]" httpUri="https://zenodo.org/record/14702841/files/figure.png" pageId="38" pageNumber="239" startId="38.[151,250,1517,1541]" targetBox="[163,1429,193,1485]" targetPageId="38" targetType="figure">
<paragraph id="C560951AFF98FFEC62EBFA09FAFBF9DD" blockId="38.[151,1375,1517,1541]" box="[151,1375,1517,1541]" pageId="38" pageNumber="239">
<emphasis id="F7AB4908FF98FFEC62EBFA09FE84F9DD" bold="true" box="[151,288,1517,1541]" pageId="38" pageNumber="239">FIGURE 30.</emphasis>
Geographical records of
@ -352,7 +353,7 @@ Males (n=3): total length: 4.084.18; carapace length: 2.102.35, width: 1.5
Only known from Sierra Nevada de Santa Marta
<collectingCountry id="BDC8D58AFF98FFEC61F2F927FC58F905" box="[910,1020,1731,1757]" name="Colombia" pageId="38" pageNumber="239">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFF98FFEC6670F927FBFBF905" box="[1036,1119,1731,1757]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="38" pageNumber="239">Fig. 31</figureCitation>
<figureCitation id="5DE4899FFF98FFEC6670F927FBFBF905" box="[1036,1119,1731,1757]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." figureDoi="http://doi.org/10.5281/zenodo.14702843" httpUri="https://zenodo.org/record/14702843/files/figure.png" pageId="38" pageNumber="239">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,65 +1,66 @@
<document id="2453A579842476465CA5D825C28C4076" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFA8FFD462EBFADFFB5CFDE5" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa olimpica Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="231" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="223" updateTime="1737369942339" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
<mods:mods id="574C0B685CE71130DD9CDB57B7B62463" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="00CEDA1EEAA51B516C25DFB87B8A7770">
<mods:title id="F304DC65159AA2D3779B8B62F147F71A">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
<document id="8FE6B92F57FD316C1A878198C6F5DDA0" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14702734" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFA8FFD462EBFADFFB5CFDE5" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa olimpica Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="231" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="223" updateTime="1737387624970" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6A15AD3A910EAFDD038B36FC755CA45E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4EE780452E3B2EC916324D42AC78FECC">
<mods:title id="F407D10BA16BA6D78B1BC5A415812B0F">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
</mods:titleInfo>
<mods:name id="1556F039FCD530C8B9A7316C19E78470" type="personal">
<mods:role id="CDFEEA5892C7B29D53762B1AC9E7A7CA">
<mods:roleTerm id="63D9FB9CC0CC6C90C86FB586779FF882">Author</mods:roleTerm>
<mods:name id="71139A857D4B2D3723EF3401F3B764D1" type="personal">
<mods:role id="2B0421F38EC086EC6EABAB512AE36F62">
<mods:roleTerm id="A54352722B75DBF87CB741C9106F0107">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9617FB43068345851B2A7F8B434A4293">Martínez, Leonel</mods:namePart>
<mods:affiliation id="078C74968424C998758902EBB7DDE533">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
<mods:namePart id="75C44E281012AD578E4B493029DD2814">Martínez, Leonel</mods:namePart>
<mods:affiliation id="31B650181B0FD60C0EC29F30D19C7BA9">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
</mods:name>
<mods:name id="43C7F995612BB63531345B1CAA4F6EC9" type="personal">
<mods:role id="1426078F8B00BFAEC9DB8B1D0AFEEBCF">
<mods:roleTerm id="939BBDDCD97A85843F030B43733AB782">Author</mods:roleTerm>
<mods:name id="88A1751A2B3D6CB562E168BB197D8B2B" type="personal">
<mods:role id="EBFCA58F6B69D05E763222B3284C88D2">
<mods:roleTerm id="118EA4D05EB70896A8F5317BBE4A9CF1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFD4C212C16A57E5AF3706EFB73CDE78">Kochalka, John A.</mods:namePart>
<mods:affiliation id="0EEFF070DF7B860D82EDE979CB074C72">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
<mods:namePart id="57363E6D13A1E5FBBC49002C8291A8C0">Kochalka, John A.</mods:namePart>
<mods:affiliation id="8ED5A4B6FDB2FDAA9E8A4F7939A480CF">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
</mods:name>
<mods:name id="60FCE257D0226A935172B8F0D6F2100D" type="personal">
<mods:role id="FCEFE810FB694FD0C6AB44576EE8F81E">
<mods:roleTerm id="C8BFCE3535C993DF34835A887D8EAFE6">Author</mods:roleTerm>
<mods:name id="018F5730185DAF6B86198B8A0B9CD079" type="personal">
<mods:role id="899052E11FA6E3759CEFDF678ED4E8F1">
<mods:roleTerm id="7A00E19DDD6212159BC143A761F5B7BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4DDAA74596A28EE275415F09AE64864">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="433DF690456FABE3489EC8C23420CE6A">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
<mods:namePart id="ACF018C5C44274A8179DF57FBEB801C4">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="774FFB105413F4B5F12C9A6A94DDC946">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
</mods:name>
<mods:name id="0A9873D00333D07817DE3C5820A32655" type="personal">
<mods:role id="A6A4AC392B8ECF68855BC5D9FE93E76D">
<mods:roleTerm id="4E6A6A4F0893D84D47292967F7465A63">Author</mods:roleTerm>
<mods:name id="73403C620CB78AFE491AD40F6B12764A" type="personal">
<mods:role id="095BD9C107DA45EE994D2F0A79D4F26F">
<mods:roleTerm id="61B312E4AB701BB516D826C332846A37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99FA2B3D5A5D4631D4DB64695A850F6D">Ramírez, Martin</mods:namePart>
<mods:affiliation id="9918628317D77B1452813EA7D580B9E8">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
<mods:namePart id="EDABD5FE496801EB4CEA6F8CAB3DCF17">Ramírez, Martin</mods:namePart>
<mods:affiliation id="0BF259B3EB2C819DF51FD795362386F5">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3CB261B2FE93D3D757F6F15FE8C7209D">text</mods:typeOfResource>
<mods:relatedItem id="9D34D9BF47211B456DAF4A7DA6C5FAF5" type="host">
<mods:titleInfo id="180336EE5D8340414C6A719AB27E84C7">
<mods:title id="76BC8B740F95D843698DDB80EC976B86">Zootaxa</mods:title>
<mods:typeOfResource id="60387459F9E224136DE5D67BC12FEEEC">text</mods:typeOfResource>
<mods:relatedItem id="586B3AFDEE79A1B4D3F6528479BD7867" type="host">
<mods:titleInfo id="018A95C8B6F4B1D5B00539CA956030FB">
<mods:title id="BE6CC2B20896B75C5E184CEA0FB5768D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5301583A3DA224A91E0C083C3AE67901">
<mods:date id="B1EF4676BB71E692C2195F0C5D0608F9">2025</mods:date>
<mods:detail id="627DCFF1BA754A3C6C04DAB5CEC3E834" type="pubDate">
<mods:number id="0272E73AC80A981E9D0C40024F47B4E5">2025-01-08</mods:number>
<mods:part id="AB0732B968C04B84D04CDFEFA33D72A3">
<mods:date id="76FCC8F050B9D52412E165974189B8FC">2025</mods:date>
<mods:detail id="4A97462172B3F04B03CD5031D51F0ECB" type="pubDate">
<mods:number id="D97796E9D0B6671F044932982600F122">2025-01-08</mods:number>
</mods:detail>
<mods:detail id="ADA5EC1B24D627430B03477FC268397C" type="volume">
<mods:number id="43B49C0AFCAD418DE1FE7ABBC7768441">5566</mods:number>
<mods:detail id="1E83CD720849E2310701132A4FAAEA7C" type="volume">
<mods:number id="5CF6CB3AF00E43D841E091AF55F82867">5566</mods:number>
</mods:detail>
<mods:detail id="E06F3977D0AC47ED4D88848D8F6EAF53" type="issue">
<mods:number id="9B98D825375A69B7D5025B3A327965DC">2</mods:number>
<mods:detail id="EA97029266A0D732F6A9D8820C4B0BC3" type="issue">
<mods:number id="F76F42B6FC5F18D639CD508CB0D29F19">2</mods:number>
</mods:detail>
<mods:extent id="E4BF97F8F0FBAFA41B22FA710307349E" unit="page">
<mods:start id="D85B9ADD3075F021DCCE5A8123099D2F">201</mods:start>
<mods:end id="BB59ADBDE3637FFB9FB4F133EDFBB47E">242</mods:end>
<mods:extent id="FF8C34BBF407E46F5B2EB6D253DCF7EF" unit="page">
<mods:start id="D3A60AA4F03A3FB298CF81A7618F8936">201</mods:start>
<mods:end id="2DB7E55B865BD0700F867AC66A6E561F">242</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="62CEEFA2B64917955D63502DA0DECE7D">
<mods:url id="AD903D758BF672299BC563DD07435A03">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
<mods:location id="96D7A173D95948C2724812E19587C602">
<mods:url id="FBF40DDB5158B978EC9E3A71FD339322">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
</mods:location>
<mods:classification id="5DC4CC05217EA1CAD03E6D33DB00994C">journal article</mods:classification>
<mods:identifier id="90C1253BA27DF0229ECF0FBDF24C54A7" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="CBCCB0AB036669258304B4E34E38FE2E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="148A0AE6121FDA3CB48E145999DEFEA8" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
<mods:classification id="AF2A0AE67F42C85BB94A7ACC95C6EF73">journal article</mods:classification>
<mods:identifier id="BBE0B3643DC63CEE6248A1D0238F83B2" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="63EB7CA62B4233A5539F91AB34105E74" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="54502D95AA8F8CDE66ACAD87044D41FC" type="Zenodo-Dep">14702734</mods:identifier>
<mods:identifier id="A081A859B2D5DA004FE6EB29BC8900F8" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
</mods:mods>
<treatment id="4D76240CFFA8FFD462EBFADFFB5CFDE5" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5" httpUri="http://treatment.plazi.org/id/4D76240CFFA8FFD462EBFADFFB5CFDE5" lastPageId="30" lastPageNumber="231" pageId="22" pageNumber="223">
<subSubSection id="8DC5C691FFA8FFDC62EBFADFFE07FA8D" box="[151,419,1338,1365]" pageId="22" pageNumber="223" type="nomenclature">
@ -76,7 +77,7 @@
</subSubSection>
<subSubSection id="8DC5C691FFA8FFDC62EBFABBFEB0FAA1" box="[151,276,1375,1401]" pageId="22" pageNumber="223" type="description">
<paragraph id="C560951AFFA8FFDC62EBFABBFEB0FAA1" blockId="22.[151,419,1338,1401]" box="[151,276,1375,1401]" pageId="22" pageNumber="223">
<figureCitation id="5DE4899FFFA8FFDC62EBFABBFEB0FAA1" box="[151,276,1375,1401]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStart-4="FIGURE 21" captionStart-5="FIGURE 22" captionStartId-0="24.[151,250,1880,1904]" captionStartId-1="25.[151,250,1879,1903]" captionStartId-2="26.[151,250,1851,1875]" captionStartId-3="27.[151,250,1858,1882]" captionStartId-4="28.[151,250,1779,1803]" captionStartId-5="29.[151,250,1731,1755]" captionTargetBox-0="[155,1428,182,1847]" captionTargetBox-1="[158,1429,192,1849]" captionTargetBox-2="[151,1428,212,1819]" captionTargetBox-3="[162,1427,189,1832]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetId-0="figure-22@24.[151,1436,181,1856]" captionTargetId-1="figure-17@25.[151,1436,181,1855]" captionTargetId-2="figure-22@26.[151,1436,181,1828]" captionTargetId-3="figure-17@27.[158,1430,181,1834]" captionTargetId-4="figure-22@28.[151,1435,181,1756]" captionTargetId-5="figure-17@29.[151,1435,181,1708]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionTargetPageId-4="28" captionTargetPageId-5="29" captionText-0="FIGURE 17. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 18. Josa olimpica sp. nov., female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum." captionText-2="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." captionText-3="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." captionText-5="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">Figs 1722</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC62EBFABBFEB0FAA1" box="[151,276,1375,1401]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStart-4="FIGURE 21" captionStart-5="FIGURE 22" captionStartId-0="24.[151,250,1880,1904]" captionStartId-1="25.[151,250,1879,1903]" captionStartId-2="26.[151,250,1851,1875]" captionStartId-3="27.[151,250,1858,1882]" captionStartId-4="28.[151,250,1779,1803]" captionStartId-5="29.[151,250,1731,1755]" captionTargetBox-0="[155,1428,182,1847]" captionTargetBox-1="[158,1429,192,1849]" captionTargetBox-2="[151,1428,212,1819]" captionTargetBox-3="[162,1427,189,1832]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetId-0="figure-22@24.[151,1436,181,1856]" captionTargetId-1="figure-17@25.[151,1436,181,1855]" captionTargetId-2="figure-22@26.[151,1436,181,1828]" captionTargetId-3="figure-17@27.[158,1430,181,1834]" captionTargetId-4="figure-22@28.[151,1435,181,1756]" captionTargetId-5="figure-17@29.[151,1435,181,1708]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionTargetPageId-4="28" captionTargetPageId-5="29" captionText-0="FIGURE 17. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 18. Josa olimpica sp. nov., female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum." captionText-2="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." captionText-3="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." captionText-5="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.14702789" figureDoi-1="http://doi.org/10.5281/zenodo.14702793" figureDoi-2="http://doi.org/10.5281/zenodo.14702797" figureDoi-3="http://doi.org/10.5281/zenodo.14702803" figureDoi-4="http://doi.org/10.5281/zenodo.14702807" figureDoi-5="http://doi.org/10.5281/zenodo.14702809" httpUri-0="https://zenodo.org/record/14702789/files/figure.png" httpUri-1="https://zenodo.org/record/14702793/files/figure.png" httpUri-2="https://zenodo.org/record/14702797/files/figure.png" httpUri-3="https://zenodo.org/record/14702803/files/figure.png" httpUri-4="https://zenodo.org/record/14702807/files/figure.png" httpUri-5="https://zenodo.org/record/14702809/files/figure.png" pageId="22" pageNumber="223">Figs 1722</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA8FFDD62EBFA43FA3DFEBD" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223" type="materials_examined">
@ -463,35 +464,35 @@ Males can be distinguished from those of other species of
<emphasis id="F7AB4908FFA9FFDD6190FE53FBBBFE08" box="[1004,1055,439,464]" italics="true" pageId="23" pageNumber="224">Josa</emphasis>
</taxonomicName>
by the following combination of characters: bifid external section of the conductor with anterior branch shorter than posterior, which is long and sharp (
<figureCitation id="5DE4899FFFA9FFDD6298FE1BFED4FDC1" box="[228,368,511,537]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19AB</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6298FE1BFED4FDC1" box="[228,368,511,537]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="23" pageNumber="224">Figs 19AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6301FE1BFE75FDC1" box="[381,465,511,537]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20AB</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6301FE1BFE75FDC1" box="[381,465,511,537]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">20AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD63A1FE1BFDDCFDC1" box="[477,632,511,537]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21AB, D, H</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD63A1FE1BFDDCFDC1" box="[477,632,511,537]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">21AB, D, H</figureCitation>
); embolus describing a loop around basal process, with a thin and long laminar projection (
<figureCitation id="5DE4899FFFA9FFDD6307FDC7FE4CFDE5" box="[379,488,547,573]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">Figs 20C</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6307FDC7FE4CFDE5" box="[379,488,547,573]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">Figs 20C</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD638BFDC7FDE7FDE5" box="[503,579,547,573]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21HI</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD638BFDC7FDE7FDE5" box="[503,579,547,573]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">21HI</figureCitation>
), and bifid median apophysis with many small denticles on external edge (
<figureCitation id="5DE4899FFFA9FFDD62E3FDA3FEA5FDB9" box="[159,257,583,609]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">Fig. 21F</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD62E3FDA3FEA5FDB9" box="[159,257,583,609]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">Fig. 21F</figureCitation>
, see inset). Females of this species are distinguished from those of other species of
<taxonomicName id="02DFEE99FFA9FFDD66C8FDA3FB43FDB8" authorityName="Keyserling" authorityYear="1891" box="[1204,1255,583,608]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="F7AB4908FFA9FFDD66C8FDA3FB43FDB8" box="[1204,1255,583,608]" italics="true" pageId="23" pageNumber="224">Josa</emphasis>
</taxonomicName>
by the shape of the posterior sclerotized projections of lateral lobes, which are narrow, handle-like, arched, and with deep rounded anterior depressions (
<figureCitation id="5DE4899FFFA9FFDD63F5FD6BFE56FD71" box="[393,498,655,681]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19E</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD63F5FD6BFE56FD71" box="[393,498,655,681]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="23" pageNumber="224">Figs 19E</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6383FD6BFD96FD71" box="[511,562,655,681]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20D</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6383FD6BFD96FD71" box="[511,562,655,681]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">20D</figureCitation>
), the median field visible in dorsal view (
<figureCitation id="5DE4899FFFA9FFDD6677FD6BFBD0FD71" box="[1035,1140,655,681]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19E</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6677FD6BFBD0FD71" box="[1035,1140,655,681]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="23" pageNumber="224">Figs 19E</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD66FDFD6BFB6BFD71" box="[1153,1231,655,681]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">22EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD66FDFD6BFB6BFD71" box="[1153,1231,655,681]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702809" httpUri="https://zenodo.org/record/14702809/files/figure.png" pageId="23" pageNumber="224">22EF</figureCitation>
), small, elongated spermathecae, laterally positioned at the median region of the epigyne (
<figureCitation id="5DE4899FFFA9FFDD61CBFD57FB9FFD15" box="[951,1083,691,717]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD61CBFD57FB9FFD15" box="[951,1083,691,717]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="23" pageNumber="224">Figs 19EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD663AFD57FB36FD15" box="[1094,1170,691,717]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD663AFD57FB36FD15" box="[1094,1170,691,717]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">20EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD66E1FD57FB19FD15" box="[1181,1213,691,717]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">22</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD66E1FD57FB19FD15" box="[1181,1213,691,717]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702809" httpUri="https://zenodo.org/record/14702809/files/figure.png" pageId="23" pageNumber="224">22</figureCitation>
).
</paragraph>
</subSubSection>
@ -502,23 +503,23 @@ by the shape of the posterior sclerotized projections of lateral lobes, which ar
(
<typeStatus id="1A642BB8FFA9FFDD63CFFD33FDBDFD29" box="[435,537,727,753]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
, MUSENUV-Ar-2682; vchLAM-260) (
<figureCitation id="5DE4899FFFA9FFDD6199FD33FB9AFD29" box="[997,1086,727,753]" captionStart="FIGURE 17" captionStartId="24.[151,250,1880,1904]" captionTargetBox="[155,1428,182,1847]" captionTargetId="figure-22@24.[151,1436,181,1856]" captionTargetPageId="24" captionText="FIGURE 17. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="23" pageNumber="224">Fig. 17</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6199FD33FB9AFD29" box="[997,1086,727,753]" captionStart="FIGURE 17" captionStartId="24.[151,250,1880,1904]" captionTargetBox="[155,1428,182,1847]" captionTargetId="figure-22@24.[151,1436,181,1856]" captionTargetPageId="24" captionText="FIGURE 17. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." figureDoi="http://doi.org/10.5281/zenodo.14702789" httpUri="https://zenodo.org/record/14702789/files/figure.png" pageId="23" pageNumber="224">Fig. 17</figureCitation>
): Total length 8.89. Carapace length 4.35, width 3.29, height 1.07. Clypeus length 0.12. Sternum length 2.31, width 1.58. Legs. Leg I: ti length 3.84/width 0.49, mt 3.20. II: ti 3.82/0.52, mt 3.13. III: ti 3.12/0.45, mt 3.29. IV: ti 3.95/0.54, mt 4.11. Spines: legs III, femur d 1-1-1, p 0-1d2ap, r 0-d1-d1; tibia v 2-2-2; metatarsus v 2-0-0. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1- d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 4.31, width 2.52, spiracleepigastrium 1.72. Color: carapace brown, with two wide longitudinal discontinuous dark brown stripes, extending to fovea and marginally covered by abundant white setae; eyes bordered by black rings, triangular band emerging from posterior side of PLE and PME. Clypeus brown, low with abundant white setae. Chelicerae brown with black spots and fine setae on frontal edge. Endites light brown, light at anterior side. Labium and sternum dark brown. Legs coxae light brown, femora-tarsi brown with some dark patches, patellae less pigmented. All articles with black dots, stripes, and covered by few white and black setae, abundant white setae between coxae. Abdomen with pattern of black patches on lighter brown background, sub-rounded by thin band; anterior region of dorsum medially with a wide longitudinal dark band extended to submedial region, followed by five wide and very short chevrons, lateral regions with many dots. Abdomen covered with few fine white setae, anterior region with a dense bunch of setae. Venter dark gray with many light patches. Palp: tegulum without tegular lobe. Median apophysis very wide, bifid, with anterior hook-shaped tip, posterior region with a denticulated edge (
<figureCitation id="5DE4899FFFA9FFDD607AFAF3FDCEFAE9" box="[518,618,1303,1329]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">Fig. 21F</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD607AFAF3FDCEFAE9" box="[518,618,1303,1329]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">Fig. 21F</figureCitation>
). Conductor large, rounded, external section apically bifid (
<figureCitation id="5DE4899FFFA9FFDD6779FAF3FA33FAE9" box="[1285,1431,1303,1329]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs. 19AB</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6779FAF3FA33FAE9" box="[1285,1431,1303,1329]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="23" pageNumber="224">Figs. 19AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD62EBFADFFF4FFA8D" box="[151,235,1339,1365]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20AB</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD62EBFADFFF4FFA8D" box="[151,235,1339,1365]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">20AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6284FADFFED4FA8D" box="[248,368,1339,1365]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21AB, D</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6284FADFFED4FA8D" box="[248,368,1339,1365]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">21AB, D</figureCitation>
), with posterior tip long, sharp, anterior tip shorter with small apical denticles, dorsal section sharp (
<figureCitation id="5DE4899FFFA9FFDD6298FABBFEE8FAA1" box="[228,332,1375,1401]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">Fig. 21B</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6298FABBFEE8FAA1" box="[228,332,1375,1401]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">Fig. 21B</figureCitation>
). Paramedian apophysis short, heavily sclerotized, sharp, narrow. Embolus apically filiform, with a narrow laminar projection extending to subapical side; basal process small, reticulate, posterior region rounded, with pronounced external keel (
<figureCitation id="5DE4899FFFA9FFDD6386FA43FDD9FA19" box="[506,637,1447,1473]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6386FA43FDD9FA19" box="[506,637,1447,1473]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="23" pageNumber="224">Figs 19EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD60F5FA43FD71FA19" box="[649,725,1447,1473]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD60F5FA43FD71FA19" box="[649,725,1447,1473]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">20EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD60A3FA43FD5AFA19" box="[735,766,1447,1473]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD60A3FA43FD5AFA19" box="[735,766,1447,1473]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">21</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFA9FFDD62BBFA2FFDD0F895" blockId="23.[151,1437,150,1869]" pageId="23" pageNumber="224">
@ -526,22 +527,22 @@ by the shape of the posterior sclerotized projections of lateral lobes, which ar
(
<typeStatus id="1A642BB8FFA9FFDD6350FA2FFE35FA3D" box="[300,401,1483,1509]" pageId="23" pageNumber="224" type="paratype">paratype</typeStatus>
, MUSENUV-Ar-2683; vchLAM-261). (
<figureCitation id="5DE4899FFFA9FFDD6126FA2FFC0BFA3D" box="[858,943,1483,1509]" captionStart="FIGURE 18" captionStartId="25.[151,250,1879,1903]" captionTargetBox="[158,1429,192,1849]" captionTargetId="figure-17@25.[151,1436,181,1855]" captionTargetPageId="25" captionText="FIGURE 18. Josa olimpica sp. nov., female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum." pageId="23" pageNumber="224">Fig. 18</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6126FA2FFC0BFA3D" box="[858,943,1483,1509]" captionStart="FIGURE 18" captionStartId="25.[151,250,1879,1903]" captionTargetBox="[158,1429,192,1849]" captionTargetId="figure-17@25.[151,1436,181,1855]" captionTargetPageId="25" captionText="FIGURE 18. Josa olimpica sp. nov., female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum." figureDoi="http://doi.org/10.5281/zenodo.14702793" httpUri="https://zenodo.org/record/14702793/files/figure.png" pageId="23" pageNumber="224">Fig. 18</figureCitation>
): Total length 11.18. Carapace length 5.98, width 4.16, height 1.72. Clypeus length 0.28. Sternum length 2.90, width 2.20. Legs. Leg I: ti length 3.43/width 0.70, mt 2.86. II: ti 4.12/0.65, mt 3.26. III: ti 3.24/0.63, mt 3.49. IV: ti 4.39/0.60, mt 3.97. Spines: legs III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; tibia v 2-2-2; metatarsus v 2bas. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v p1-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1- 1-0-1, IV=III but femur r 1d. Abdomen: length 6.08, width 3.96, spiracleepigastrium 2.15. Color pattern as in male, but more pigmented. Lateral regions of abdomen with many dots. Epigyne: median field transverse, wide, oval in posterior view (
<figureCitation id="5DE4899FFFA9FFDD63E3F923FD81F939" box="[415,549,1735,1761]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">Fig. 22EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD63E3F923FD81F939" box="[415,549,1735,1761]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702809" httpUri="https://zenodo.org/record/14702809/files/figure.png" pageId="23" pageNumber="224">Fig. 22EF</figureCitation>
). Lateral lobes with sclerotized extensions wide, rounded, with small, rounded concavities at anterior side. Spermathecae small, oval, lateromedially placed. Copulatory ducts long, thin, coiled along longitudinal axis, oriented in parallel, anterior region rounded (
<figureCitation id="5DE4899FFFA9FFDD61D9F8EBFBAAF8F1" box="[933,1038,1807,1833]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">Figs 20E</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD61D9F8EBFBAAF8F1" box="[933,1038,1807,1833]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">Figs 20E</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6666F8EBFBD4F8F1" box="[1050,1136,1807,1833]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21AB</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD6666F8EBFBD4F8F1" box="[1050,1136,1807,1833]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="23" pageNumber="224">21AB</figureCitation>
). Ducts of accessory bulbs long, convergent (
<figureCitation id="5DE4899FFFA9FFDD631EF8D7FE41F895" box="[354,485,1843,1869]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD631EF8D7FE41F895" box="[354,485,1843,1869]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." figureDoi="http://doi.org/10.5281/zenodo.14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="23" pageNumber="224">Figs 19EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD638DF8D7FD99F895" box="[497,573,1843,1869]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20EF</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD638DF8D7FD99F895" box="[497,573,1843,1869]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="23" pageNumber="224">20EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD603BF8D7FDC3F895" box="[583,615,1843,1869]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">22</figureCitation>
<figureCitation id="5DE4899FFFA9FFDD603BF8D7FDC3F895" box="[583,615,1843,1869]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702809" httpUri="https://zenodo.org/record/14702809/files/figure.png" pageId="23" pageNumber="224">22</figureCitation>
).
</paragraph>
<caption id="91A0C592FFA6FFD262EBF8BCFD3CF84C" pageId="24" pageNumber="225" startId="24.[151,250,1880,1904]" targetBox="[155,1428,182,1847]" targetPageId="24" targetType="figure">
<caption id="91A0C592FFA6FFD262EBF8BCFD3CF84C" ID-DOI="http://doi.org/10.5281/zenodo.14702789" ID-Zenodo-Dep="14702789" httpUri="https://zenodo.org/record/14702789/files/figure.png" pageId="24" pageNumber="225" startId="24.[151,250,1880,1904]" targetBox="[155,1428,182,1847]" targetPageId="24" targetType="figure">
<paragraph id="C560951AFFA6FFD262EBF8BCFD3CF84C" blockId="24.[151,1436,1880,1940]" pageId="24" pageNumber="225">
<emphasis id="F7AB4908FFA6FFD262EBF8BCFE87F8A8" bold="true" box="[151,291,1880,1904]" pageId="24" pageNumber="225">FIGURE 17.</emphasis>
<taxonomicName id="02DFEE99FFA6FFD26351F8BDFE1BF8A8" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[301,447,1880,1904]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
@ -553,7 +554,7 @@ by the shape of the posterior sclerotized projections of lateral lobes, which ar
, male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFA7FFD362EBF8B3FE50F84B" pageId="25" pageNumber="226" startId="25.[151,250,1879,1903]" targetBox="[158,1429,192,1849]" targetPageId="25" targetType="figure">
<caption id="91A0C592FFA7FFD362EBF8B3FE50F84B" ID-DOI="http://doi.org/10.5281/zenodo.14702793" ID-Zenodo-Dep="14702793" httpUri="https://zenodo.org/record/14702793/files/figure.png" pageId="25" pageNumber="226" startId="25.[151,250,1879,1903]" targetBox="[158,1429,192,1849]" targetPageId="25" targetType="figure">
<paragraph id="C560951AFFA7FFD362EBF8B3FE50F84B" blockId="25.[151,1436,1879,1939]" pageId="25" pageNumber="226">
<emphasis id="F7AB4908FFA7FFD362EBF8B3FE86F8B7" bold="true" box="[151,290,1879,1903]" pageId="25" pageNumber="226">FIGURE 18.</emphasis>
<taxonomicName id="02DFEE99FFA7FFD36357F8BCFE1FF8B7" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,443,1879,1903]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
@ -565,7 +566,7 @@ by the shape of the posterior sclerotized projections of lateral lobes, which ar
, female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFA4FFD062EBF8DFFBB2F83C" pageId="26" pageNumber="227" startId="26.[151,250,1851,1875]" targetBox="[151,1428,212,1819]" targetPageId="26" targetType="figure">
<caption id="91A0C592FFA4FFD062EBF8DFFBB2F83C" ID-DOI="http://doi.org/10.5281/zenodo.14702797" ID-Zenodo-Dep="14702797" httpUri="https://zenodo.org/record/14702797/files/figure.png" pageId="26" pageNumber="227" startId="26.[151,250,1851,1875]" targetBox="[151,1428,212,1819]" targetPageId="26" targetType="figure">
<paragraph id="C560951AFFA4FFD062EBF8DFFBB2F83C" blockId="26.[151,1436,1851,2020]" pageId="26" pageNumber="227">
<emphasis id="F7AB4908FFA4FFD062EBF8DFFEBBF88C" bold="true" box="[151,287,1851,1876]" pageId="26" pageNumber="227">FIGURE 19.</emphasis>
<taxonomicName id="02DFEE99FFA4FFD06358F8D8FE15F88C" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[292,433,1852,1876]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
@ -579,7 +580,7 @@ by the shape of the posterior sclerotized projections of lateral lobes, which ar
AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FFA5FFD162EBF8A6FBAFF832" pageId="27" pageNumber="228" startId="27.[151,250,1858,1882]" targetBox="[162,1427,189,1832]" targetPageId="27" targetType="figure">
<caption id="91A0C592FFA5FFD162EBF8A6FBAFF832" ID-DOI="http://doi.org/10.5281/zenodo.14702803" ID-Zenodo-Dep="14702803" httpUri="https://zenodo.org/record/14702803/files/figure.png" pageId="27" pageNumber="228" startId="27.[151,250,1858,1882]" targetBox="[162,1427,189,1832]" targetPageId="27" targetType="figure">
<paragraph id="C560951AFFA5FFD162EBF8A6FBAFF832" blockId="27.[151,1436,1858,2027]" pageId="27" pageNumber="228">
<emphasis id="F7AB4908FFA5FFD162EBF8A6FEBBF882" bold="true" box="[151,287,1858,1882]" pageId="27" pageNumber="228">FIGURE 20.</emphasis>
<taxonomicName id="02DFEE99FFA5FFD16358F8A7FE15F882" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[292,433,1858,1882]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
@ -593,7 +594,7 @@ AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertil
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct.
</paragraph>
</caption>
<caption id="91A0C592FFA2FFD662EBF917FEA8F818" pageId="28" pageNumber="229" startId="28.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="28" targetType="figure">
<caption id="91A0C592FFA2FFD662EBF917FEA8F818" ID-DOI="http://doi.org/10.5281/zenodo.14702807" ID-Zenodo-Dep="14702807" httpUri="https://zenodo.org/record/14702807/files/figure.png" pageId="28" pageNumber="229" startId="28.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="28" targetType="figure">
<paragraph id="C560951AFFA2FFD662EBF917FEA8F818" blockId="28.[151,1437,1779,1984]" pageId="28" pageNumber="229">
<emphasis id="F7AB4908FFA2FFD662EBF917FE85F8D4" bold="true" box="[151,289,1779,1804]" pageId="28" pageNumber="229">FIGURE 21.</emphasis>
<taxonomicName id="02DFEE99FFA2FFD66354F910FE13F8D4" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[296,439,1780,1804]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
@ -607,7 +608,7 @@ AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; d
C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FFA3FFD762EBF927FED0F890" pageId="29" pageNumber="230" startId="29.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="29" targetType="figure">
<caption id="91A0C592FFA3FFD762EBF927FED0F890" ID-DOI="http://doi.org/10.5281/zenodo.14702809" ID-Zenodo-Dep="14702809" httpUri="https://zenodo.org/record/14702809/files/figure.png" pageId="29" pageNumber="230" startId="29.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="29" targetType="figure">
<paragraph id="C560951AFFA3FFD762EBF927FED0F890" blockId="29.[151,1437,1731,1864]" pageId="29" pageNumber="230">
<emphasis id="F7AB4908FFA3FFD762EBF927FE87F904" bold="true" box="[151,291,1731,1756]" pageId="29" pageNumber="230">FIGURE 22.</emphasis>
<taxonomicName id="02DFEE99FFA3FFD76350F920FE19F904" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[300,445,1732,1756]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
@ -632,7 +633,7 @@ Males (n=5): total length: 7.5111.35; carapace length: 3.754.35, width: 2.
Only known from Sierra Nevada de Santa Marta,
<collectingCountry id="BDC8D58AFFA0FFD461E9FEE7FBA7FEC5" box="[917,1027,259,285]" name="Colombia" pageId="30" pageNumber="231">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFFA0FFD4666FFEE7FBC2FEC5" box="[1043,1126,259,285]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="30" pageNumber="231">Fig. 31</figureCitation>
<figureCitation id="5DE4899FFFA0FFD4666FFEE7FBC2FEC5" box="[1043,1126,259,285]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." figureDoi="http://doi.org/10.5281/zenodo.14702843" httpUri="https://zenodo.org/record/14702843/files/figure.png" pageId="30" pageNumber="231">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFA0FFD462BBFEC2FB5CFDE5" blockId="30.[151,1437,150,573]" pageId="30" pageNumber="231">

View file

@ -1,65 +1,66 @@
<document id="D5D99B526D86442527107E67F4E7D2E9" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFB1FFDC62EBFC13FBD7FB31" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa nadineae Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="223" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="216" updateTime="1737369942339" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
<mods:mods id="5F46D0027A2A6732F41BDD12D2E77F38" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="37D75FB2CC2054B9CD6B21EDDB6E811C">
<mods:title id="9C1AE8869BD78E917E6ACFF1AB5BE500">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
<document id="D8214CAC55253145BA69AF86A631A106" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14702734" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFB1FFDC62EBFC13FBD7FB31" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa nadineae Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="223" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="216" updateTime="1737387624970" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="715D93FA0FC31DAD4B30745F065542FF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="05E10EC958BB5C09944F676FD095EEE1">
<mods:title id="B38E621D7661DB885F080D2226AF37E7">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
</mods:titleInfo>
<mods:name id="22D769C9C03707ABC4655CC31B2DBD60" type="personal">
<mods:role id="8D9E0F63B91055A9EE91F5F697A8B312">
<mods:roleTerm id="BDB3A1F1F30452E0389C4A97DEA305E6">Author</mods:roleTerm>
<mods:name id="8734F08952CA73A90D886CE7DD4AE715" type="personal">
<mods:role id="49218192B4A72543D6F6396FB1B7B54F">
<mods:roleTerm id="55D8BAE20F9E04CBCF3A3AB0AD12D8E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7CE08896FBE11AF0B691AEC5230C117B">Martínez, Leonel</mods:namePart>
<mods:affiliation id="69FE04FE9875997C0CEE1E6439534B62">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
<mods:namePart id="D9073D0CED7B70A21F94FBF92A4E3AF5">Martínez, Leonel</mods:namePart>
<mods:affiliation id="B10800BAFBD9135D8988FFA762D79A56">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
</mods:name>
<mods:name id="C1483530EF5897ABDC297B3293CF78E7" type="personal">
<mods:role id="E8CF7229463A5282BB18E1C884D58C1D">
<mods:roleTerm id="A19BD55921A32935D7DD185272853102">Author</mods:roleTerm>
<mods:name id="F9838476DC30774349CDA13F5BEA3AA4" type="personal">
<mods:role id="BBFD7678EE599D7DB5225DF68CCCFFD8">
<mods:roleTerm id="8D6DC9206FFE20AF755772FA9C6CFF8B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E4B0A74F13DE009DB60E907668E94DC">Kochalka, John A.</mods:namePart>
<mods:affiliation id="C84B1649D51849637777B1B2AA745B22">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
<mods:namePart id="F0288C077584E7F520DE9FFA2339A29B">Kochalka, John A.</mods:namePart>
<mods:affiliation id="29AE76D9735E547F03AC63C0741C6AFA">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
</mods:name>
<mods:name id="4CF166767279E40F1C3590C3EF080494" type="personal">
<mods:role id="A4095137412322FBAD135FCFD425464E">
<mods:roleTerm id="61B227A934399FD2B2DF1A6883F0EFFE">Author</mods:roleTerm>
<mods:name id="DC37CE4D48EF2149545437E20DEC52D6" type="personal">
<mods:role id="1C062F030046E199BC8727D97CCD835F">
<mods:roleTerm id="E8DBD515C075285D107D9EFE2148D2D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D94203A24136F3E73AD67B87B100990">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="29C1AA22E8B72AC78E2EBC4C85B9C1EF">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
<mods:namePart id="EC162A99C24DE77A4CE37049891A12A7">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="1C54C27371AB83AD7756E9C4C3FAB7DB">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
</mods:name>
<mods:name id="CBD82E1DA6CA09843231F0ED6F72A1A5" type="personal">
<mods:role id="4B33DBC16B238E21D42C3A85E1E53593">
<mods:roleTerm id="CF44E629D35BE5ED4611266775430EF7">Author</mods:roleTerm>
<mods:name id="6FA9682647B76E99C9863D1164D6D0EC" type="personal">
<mods:role id="15BA989054B6D616A38BA586AE568DE4">
<mods:roleTerm id="CB4A8AB2E2561CE01E077BA0BF93D875">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19597DD42C0A11ED73D69D6F02E88776">Ramírez, Martin</mods:namePart>
<mods:affiliation id="3D72E8E41F7F582C9E61F7E0E15630F4">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
<mods:namePart id="B918EAF8118A5E39B91CD4E574559B28">Ramírez, Martin</mods:namePart>
<mods:affiliation id="F8128EA6D964332E915966771252600F">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1DE7562ECA2648C4D3D7DAEBCEBD3A5D">text</mods:typeOfResource>
<mods:relatedItem id="26046ABDCBF6D75A906FF047F5955BB1" type="host">
<mods:titleInfo id="6FD714B4B88AA709E748874DA5120CF9">
<mods:title id="F9D4E1BA19CBABEA4EEAD66B0D67E4E9">Zootaxa</mods:title>
<mods:typeOfResource id="453BB052568B4D6EB8141FE7CB33E07F">text</mods:typeOfResource>
<mods:relatedItem id="A7C7B99EED2E4100C8E55B06915BF8B8" type="host">
<mods:titleInfo id="CC523990599403499A368CD4BBBC3397">
<mods:title id="A0336A386DB03B18F3161ABB7B31CB9E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D116F25C133FB2C73172E3008E25FD6B">
<mods:date id="1D59A709967A34A5AC860CD7EB372547">2025</mods:date>
<mods:detail id="CBF14E9C0334191B8199D9E20B3777A0" type="pubDate">
<mods:number id="2F5D7191FFFDAFC51323E5E8B823AB0D">2025-01-08</mods:number>
<mods:part id="48489392E45B7ABA84BB56BB322D8FFE">
<mods:date id="E6C8ACC8650DE24A0278AC4055AF956F">2025</mods:date>
<mods:detail id="C3ECD83D8D4475BA1705B6AE9CCCFBFA" type="pubDate">
<mods:number id="5585BA1296F0A751225651F6EA4A5C03">2025-01-08</mods:number>
</mods:detail>
<mods:detail id="4947D05952A236316FC3C94B39B00B78" type="volume">
<mods:number id="9CF1CE7481B6B51BD87F611186C79B09">5566</mods:number>
<mods:detail id="51D13409FD8E34901F6AB4B38A129E3F" type="volume">
<mods:number id="721AC4ED55CE78C2CDD74A9F12731B57">5566</mods:number>
</mods:detail>
<mods:detail id="5883CF3B12CADCDCE38D21D339DD47CD" type="issue">
<mods:number id="5BA3FC834B4ACE51596306546A377CDF">2</mods:number>
<mods:detail id="C9FDA4FB65E9C5A93EEB55697785E170" type="issue">
<mods:number id="A05BBBD96F0C280932A95138A520E6D7">2</mods:number>
</mods:detail>
<mods:extent id="22520A8274196AB92DF739C20A569742" unit="page">
<mods:start id="E2FBCFA7C24FDA35845DD4F57210EFAA">201</mods:start>
<mods:end id="3F25F24431B675F07F44E538490E3BF0">242</mods:end>
<mods:extent id="8759B81F82703B024507B95E4232C27C" unit="page">
<mods:start id="08ECE38FAF0E5281C828110F16763C09">201</mods:start>
<mods:end id="9DDDCA69CF05522B4C7E031DE9C9A6C4">242</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="BE40813FCFF02826BCF57A571CBD03EC">
<mods:url id="EA73082A3168A182C1CFA90E8AF7C6E2">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
<mods:location id="CD376F83BE21FB9E440A03099A9E837B">
<mods:url id="428E9214CD145BD8E22518EF97615725">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
</mods:location>
<mods:classification id="4F7592730B65788B406F461610CE37D8">journal article</mods:classification>
<mods:identifier id="CBBD1C9638ED652922F09D623B5EA865" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="9CA82574C2584F5F750990502E8352EA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4E5C6D45D9ADD5D6D2DB54DC33DAB62D" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
<mods:classification id="61A37D57575345295593691CDCB5FD14">journal article</mods:classification>
<mods:identifier id="5E34F5727BB37008BEB9974E18DFFCC4" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="271418481A25F296D04A5CFB9AE294B0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="83D9742C63333BC7DCB6890E41FC3CBC" type="Zenodo-Dep">14702734</mods:identifier>
<mods:identifier id="9CA92987782BA4652F3226FDB89A5C15" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
</mods:mods>
<treatment id="4D76240CFFB1FFDC62EBFC13FBD7FB31" LSID="urn:lsid:plazi:treatment:4D76240CFFB1FFDC62EBFC13FBD7FB31" httpUri="http://treatment.plazi.org/id/4D76240CFFB1FFDC62EBFC13FBD7FB31" lastPageId="22" lastPageNumber="223" pageId="15" pageNumber="216">
<subSubSection id="8DC5C691FFB1FFC562EBFC13FE0EFBC9" box="[151,426,1014,1041]" pageId="15" pageNumber="216" type="nomenclature">
@ -76,7 +77,7 @@
</subSubSection>
<subSubSection id="8DC5C691FFB1FFC562EBFBFFFE9CFBED" box="[151,312,1051,1077]" pageId="15" pageNumber="216" type="description">
<paragraph id="C560951AFFB1FFC562EBFBFFFE9CFBED" blockId="15.[151,426,1014,1077]" box="[151,312,1051,1077]" pageId="15" pageNumber="216">
<figureCitation id="5DE4899FFFB1FFC562EBFBFFFE9CFBED" box="[151,312,1051,1077]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStart-5="FIGURE 16" captionStartId-0="16.[151,250,1882,1906]" captionStartId-1="17.[151,250,1882,1906]" captionStartId-2="18.[151,250,1836,1860]" captionStartId-3="19.[151,250,1866,1890]" captionStartId-4="20.[151,250,1779,1803]" captionStartId-5="21.[151,250,1731,1755]" captionTargetBox-0="[159,1427,189,1858]" captionTargetBox-1="[159,1423,186,1853]" captionTargetBox-2="[151,1412,182,1812]" captionTargetBox-3="[155,1432,185,1840]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetId-0="figure-22@16.[151,1436,181,1858]" captionTargetId-1="figure-17@17.[151,1436,181,1858]" captionTargetId-2="figure-22@18.[151,1436,181,1812]" captionTargetId-3="figure-17@19.[151,1435,181,1842]" captionTargetId-4="figure-22@20.[151,1435,181,1756]" captionTargetId-5="figure-17@21.[151,1435,181,1708]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionText-0="FIGURE 11. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 12. Josa nadineae sp. nov., female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-2="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." captionText-3="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." captionText-5="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Figures 1116</figureCitation>
<figureCitation id="5DE4899FFFB1FFC562EBFBFFFE9CFBED" box="[151,312,1051,1077]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStart-5="FIGURE 16" captionStartId-0="16.[151,250,1882,1906]" captionStartId-1="17.[151,250,1882,1906]" captionStartId-2="18.[151,250,1836,1860]" captionStartId-3="19.[151,250,1866,1890]" captionStartId-4="20.[151,250,1779,1803]" captionStartId-5="21.[151,250,1731,1755]" captionTargetBox-0="[159,1427,189,1858]" captionTargetBox-1="[159,1423,186,1853]" captionTargetBox-2="[151,1412,182,1812]" captionTargetBox-3="[155,1432,185,1840]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetId-0="figure-22@16.[151,1436,181,1858]" captionTargetId-1="figure-17@17.[151,1436,181,1858]" captionTargetId-2="figure-22@18.[151,1436,181,1812]" captionTargetId-3="figure-17@19.[151,1435,181,1842]" captionTargetId-4="figure-22@20.[151,1435,181,1756]" captionTargetId-5="figure-17@21.[151,1435,181,1708]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionText-0="FIGURE 11. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 12. Josa nadineae sp. nov., female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-2="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." captionText-3="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." captionText-5="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.14702773" figureDoi-1="http://doi.org/10.5281/zenodo.14702777" figureDoi-2="http://doi.org/10.5281/zenodo.14702781" figureDoi-3="http://doi.org/10.5281/zenodo.14702783" figureDoi-4="http://doi.org/10.5281/zenodo.14702785" figureDoi-5="http://doi.org/10.5281/zenodo.14702787" httpUri-0="https://zenodo.org/record/14702773/files/figure.png" httpUri-1="https://zenodo.org/record/14702777/files/figure.png" httpUri-2="https://zenodo.org/record/14702781/files/figure.png" httpUri-3="https://zenodo.org/record/14702783/files/figure.png" httpUri-4="https://zenodo.org/record/14702785/files/figure.png" httpUri-5="https://zenodo.org/record/14702787/files/figure.png" pageId="15" pageNumber="216">Figures 1116</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB1FFC562EBFBB8FCE6FAAB" pageId="15" pageNumber="216" type="materials_examined">
@ -204,29 +205,29 @@ Males resemble those of
)
</taxonomicName>
by the short and tape-shaped embolus (
<figureCitation id="5DE4899FFFB1FFC562E3FA0CFF4AF9DA" box="[159,238,1512,1538]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Fig. 13</figureCitation>
<figureCitation id="5DE4899FFFB1FFC562E3FA0CFF4AF9DA" box="[159,238,1512,1538]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="15" pageNumber="216">Fig. 13</figureCitation>
EF, compare with
<bibRefCitation id="A14EE8EBFFB1FFC563AEFA0CFD15F9DA" author="Duperre, N." box="[466,689,1512,1538]" pageId="15" pageNumber="216" pagination="118 - 277" refId="ref14337" refString="Duperre, N. (2023) New light on some historical type specimens - in relation to the South American spider (Araneae) fauna-. New Zealand Journal of Zoology, 50 (1), 118 - 277. https: // doi. org / 10.1080 / 03014223.2022.2123835" type="journal article" year="2023">Dupérré, 2023: 139</bibRefCitation>
, fig. 16AC; Poy. et. al. 2023: 6, fig. 3E). It can be distinguished from the latter species by the following combination of characters: laminar projection of the embolus larger (
<figureCitation id="5DE4899FFFB1FFC56716F9E8FF48F992" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Figs 13AD</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56716F9E8FF48F992" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="15" pageNumber="216">Figs 13AD</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56285F9D4FEE8F992" box="[249,332,1584,1610]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="15" pageNumber="216">14AB</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56285F9D4FEE8F992" box="[249,332,1584,1610]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="15" pageNumber="216">14AB</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56325F9D4FE07F992" box="[345,419,1584,1611]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="15" pageNumber="216">15B, I</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56325F9D4FE07F992" box="[345,419,1584,1611]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="15" pageNumber="216">15B, I</figureCitation>
), bifid external section of the conductor with both branches interrupted by a deep groove (
<figureCitation id="5DE4899FFFB1FFC562E3F9B0FE8EF9B7" box="[159,298,1620,1647]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="15" pageNumber="216">Fig. 15AB</figureCitation>
<figureCitation id="5DE4899FFFB1FFC562E3F9B0FE8EF9B7" box="[159,298,1620,1647]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="15" pageNumber="216">Fig. 15AB</figureCitation>
), posterior branch of the conductor protruded, with many small spicules and longer than the anterior branch, which is laminar and apically irregular with small reticulations, bifid median apophysis with small scales on the external edge (
<figureCitation id="5DE4899FFFB1FFC56318F978FE62F96F" box="[356,454,1692,1719]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="15" pageNumber="216">Fig. 15F</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56318F978FE62F96F" box="[356,454,1692,1719]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="15" pageNumber="216">Fig. 15F</figureCitation>
), and pronounced retrolateral basal lobe (
<figureCitation id="5DE4899FFFB1FFC561EFF978FC59F96E" box="[915,1021,1692,1718]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Figs 13A</figureCitation>
<figureCitation id="5DE4899FFFB1FFC561EFF978FC59F96E" box="[915,1021,1692,1718]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="15" pageNumber="216">Figs 13A</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56675F978FB99F96E" box="[1033,1085,1692,1718]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="15" pageNumber="216">14A</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56675F978FB99F96E" box="[1033,1085,1692,1718]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="15" pageNumber="216">14A</figureCitation>
). The females are distinguished by the spermathecae converging on the median line, touching each other, the obliquely oriented posterior borders of the epigyne and the copulatory ducts anteriorly arched, widening towards the anterior part of vulva (
<figureCitation id="5DE4899FFFB1FFC56768F900FA3CF926" box="[1300,1432,1764,1790]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Figs 13EF</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56768F900FA3CF926" box="[1300,1432,1764,1790]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="15" pageNumber="216">Figs 13EF</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC562EBF8ECFF46F8FA" box="[151,226,1800,1826]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="15" pageNumber="216">14EF</figureCitation>
<figureCitation id="5DE4899FFFB1FFC562EBF8ECFF46F8FA" box="[151,226,1800,1826]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="15" pageNumber="216">14EF</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56292F8ECFEA9F8FA" box="[238,269,1800,1826]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="15" pageNumber="216">16</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56292F8ECFEA9F8FA" box="[238,269,1800,1826]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702787" httpUri="https://zenodo.org/record/14702787/files/figure.png" pageId="15" pageNumber="216">16</figureCitation>
).
</paragraph>
</subSubSection>
@ -237,34 +238,34 @@ EF, compare with
(
<typeStatus id="1A642BB8FFB1FFC563CFF8C8FDBDF89E" box="[435,537,1836,1862]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
, MUSENUV-Ar-2678; vchLAM-249) (
<figureCitation id="5DE4899FFFB1FFC5619AF8C8FB9AF89E" box="[998,1086,1836,1862]" captionStart="FIGURE 11" captionStartId="16.[151,250,1882,1906]" captionTargetBox="[159,1427,189,1858]" captionTargetId="figure-22@16.[151,1436,181,1858]" captionTargetPageId="16" captionText="FIGURE 11. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="15" pageNumber="216">Fig. 11</figureCitation>
<figureCitation id="5DE4899FFFB1FFC5619AF8C8FB9AF89E" box="[998,1086,1836,1862]" captionStart="FIGURE 11" captionStartId="16.[151,250,1882,1906]" captionTargetBox="[159,1427,189,1858]" captionTargetId="figure-22@16.[151,1436,181,1858]" captionTargetPageId="16" captionText="FIGURE 11. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." figureDoi="http://doi.org/10.5281/zenodo.14702773" httpUri="https://zenodo.org/record/14702773/files/figure.png" pageId="15" pageNumber="216">Fig. 11</figureCitation>
): Total length 5.75. Carapace length 2.95, width 2.13, height 1.10. Clypeus length 0.13. Sternum length 1.45, width 0.98. Legs. Leg I: ti length 1.79/width 0.36, mt 1.39. II: ti 1.70/0.35, mt 1.46. III: ti 1.72/0.38, mt 1.75. IV: ti 2.74/0.44, mt 2.59. Spines: legs III, femur d 1-1-1, p 2ap, r 0-d1-d1; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0; metatarsus v 2-0-0, p d1-0. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 2.81, width 1.56, spiracleepigastrium 1.01. Color: carapace brown, marginally darker, with two wide black degraded bands, converging and narrow anteriorly. Ocular area dark brown with eyes bordered on a black patch. Lateral regions of carapace upholstered with small fine white setae. Clypeus brown, low with some white setae. Chelicerae brown with fine setae and dark patches on frontal edge. Endites light brown, with faint medial dark patch. Labium brown reddish, anteriorly lighter. Sternum brown with faint black patches. Legs uniformly brown, covered by white setae, mainly between coxae. Abdomen dorsum with a pattern of black spots on a brown background, surrounded by a thin band, extended to medial region; dorsum with an anterior medial dark patch with posterior extensions, followed by irregular spots. Abdomen covered with fine white setae; anterior region with a dense bunch of setae, marginal ring covered by black setae. Venter uniformly brown with light and dark dots, dark patches surrounding the spinnerets base. Palp: tegulum with a basal small lobe, with granulose texture (
<figureCitation id="5DE4899FFFA8FFDC6316FE8BFE73FE51" box="[362,471,367,393]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13A</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6316FE8BFE73FE51" box="[362,471,367,393]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="22" pageNumber="223">Figs 13A</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6398FE8BFDB3FE51" box="[484,535,367,393]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14A</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6398FE8BFDB3FE51" box="[484,535,367,393]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="22" pageNumber="223">14A</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6058FE8AFDDFFE51" box="[548,635,366,393]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">15A, G</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6058FE8AFDDFFE51" box="[548,635,366,393]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="22" pageNumber="223">15A, G</figureCitation>
). Median apophysis long, laminar, ending in a very short hook-shaped tip, with many denticles (
<figureCitation id="5DE4899FFFA8FFDC63CEFE77FDB6FE75" box="[434,530,402,429]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">Fig. 15F</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC63CEFE77FDB6FE75" box="[434,530,402,429]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="22" pageNumber="223">Fig. 15F</figureCitation>
, see square). Conductor small, rounded, ventral section apically bifid, with many narrow furrows, anterior tip apically laminar with several very small denticles, posterior tip with small granules, posterior tip slightly longer than anterior one; base of the conductor with a sharp, spine-shaped projection (
<figureCitation id="5DE4899FFFA8FFDC674EFE3FFF08FDC1" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">Fig. 15A, B</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC674EFE3FFF08FDC1" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="22" pageNumber="223">Fig. 15A, B</figureCitation>
). Paramedian apophysis cup-shaped, concave (
<figureCitation id="5DE4899FFFA8FFDC60B4FE1BFCF3FDC1" box="[712,855,511,537]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13AB</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC60B4FE1BFCF3FDC1" box="[712,855,511,537]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="22" pageNumber="223">Figs 13AB</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6118FE1BFC1CFDC1" box="[868,952,511,537]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14AB</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6118FE1BFC1CFDC1" box="[868,952,511,537]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="22" pageNumber="223">14AB</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC61BAFE1AFBFBFDC1" box="[966,1119,510,537]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">15AB, D, F</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC61BAFE1AFBFBFDC1" box="[966,1119,510,537]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="22" pageNumber="223">15AB, D, F</figureCitation>
). Embolus tape-shaped, not describing a loop, very short, with large laminar projection (
<figureCitation id="5DE4899FFFA8FFDC613EFDC7FC47FDE5" box="[834,995,546,573]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">Fig. 15B, G, I</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC613EFDC7FC47FDE5" box="[834,995,546,573]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="22" pageNumber="223">Fig. 15B, G, I</figureCitation>
), apically twisted; basal process small, flat, sclerotized, with poorly pronounced external keel and medial depression (
<figureCitation id="5DE4899FFFA8FFDC667AFDA3FB37FDB9" box="[1030,1171,583,609]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13AD</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC667AFDA3FB37FDB9" box="[1030,1171,583,609]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="22" pageNumber="223">Figs 13AD</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC66E3FDA3FB57FDB9" box="[1183,1267,583,609]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14AC</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC66E3FDA3FB57FDB9" box="[1183,1267,583,609]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="22" pageNumber="223">14AC</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6682FDA2FAB9FDB9" box="[1278,1309,582,609]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">15</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6682FDA2FAB9FDB9" box="[1278,1309,582,609]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="22" pageNumber="223">15</figureCitation>
).
</paragraph>
<caption id="91A0C592FFAEFFDA62EBF8BEFD3CF84E" pageId="16" pageNumber="217" startId="16.[151,250,1882,1906]" targetBox="[159,1427,189,1858]" targetPageId="16" targetType="figure">
<caption id="91A0C592FFAEFFDA62EBF8BEFD3CF84E" ID-DOI="http://doi.org/10.5281/zenodo.14702773" ID-Zenodo-Dep="14702773" httpUri="https://zenodo.org/record/14702773/files/figure.png" pageId="16" pageNumber="217" startId="16.[151,250,1882,1906]" targetBox="[159,1427,189,1858]" targetPageId="16" targetType="figure">
<paragraph id="C560951AFFAEFFDA62EBF8BEFD3CF84E" blockId="16.[151,1436,1882,1942]" pageId="16" pageNumber="217">
<emphasis id="F7AB4908FFAEFFDA62EBF8BEFE86F8AA" bold="true" box="[151,290,1882,1906]" pageId="16" pageNumber="217">FIGURE 11.</emphasis>
<taxonomicName id="02DFEE99FFAEFFDA6357F8BFFE65F8AA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,449,1882,1906]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
@ -276,7 +277,7 @@ EF, compare with
, male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFAFFFDB62EBF8BEFD3CF84E" pageId="17" pageNumber="218" startId="17.[151,250,1882,1906]" targetBox="[159,1423,186,1853]" targetPageId="17" targetType="figure">
<caption id="91A0C592FFAFFFDB62EBF8BEFD3CF84E" ID-DOI="http://doi.org/10.5281/zenodo.14702777" ID-Zenodo-Dep="14702777" httpUri="https://zenodo.org/record/14702777/files/figure.png" pageId="17" pageNumber="218" startId="17.[151,250,1882,1906]" targetBox="[159,1423,186,1853]" targetPageId="17" targetType="figure">
<paragraph id="C560951AFFAFFFDB62EBF8BEFD3CF84E" blockId="17.[151,1436,1882,1942]" pageId="17" pageNumber="218">
<emphasis id="F7AB4908FFAFFFDB62EBF8BEFE86F8AA" bold="true" box="[151,290,1882,1906]" pageId="17" pageNumber="218">FIGURE 12.</emphasis>
<taxonomicName id="02DFEE99FFAFFFDB6356F8BFFE1AF8AA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[298,446,1882,1906]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
@ -288,7 +289,7 @@ EF, compare with
, female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFACFFD862EBF8C8FBEEF80C" pageId="18" pageNumber="219" startId="18.[151,250,1836,1860]" targetBox="[151,1412,182,1812]" targetPageId="18" targetType="figure">
<caption id="91A0C592FFACFFD862EBF8C8FBEEF80C" ID-DOI="http://doi.org/10.5281/zenodo.14702781" ID-Zenodo-Dep="14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="18" pageNumber="219" startId="18.[151,250,1836,1860]" targetBox="[151,1412,182,1812]" targetPageId="18" targetType="figure">
<paragraph id="C560951AFFACFFD862EBF8C8FBEEF80C" blockId="18.[151,1436,1836,2005]" pageId="18" pageNumber="219">
<emphasis id="F7AB4908FFACFFD862EBF8C8FE87F89C" bold="true" box="[151,291,1836,1860]" pageId="18" pageNumber="219">FIGURE 13.</emphasis>
<taxonomicName id="02DFEE99FFACFFD86351F8C9FE67F89C" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[301,451,1836,1860]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
@ -302,7 +303,7 @@ EF, compare with
C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe.
</paragraph>
</caption>
<caption id="91A0C592FFADFFD962EBF8AEFBF9F82A" pageId="19" pageNumber="220" startId="19.[151,250,1866,1890]" targetBox="[155,1432,185,1840]" targetPageId="19" targetType="figure">
<caption id="91A0C592FFADFFD962EBF8AEFBF9F82A" ID-DOI="http://doi.org/10.5281/zenodo.14702783" ID-Zenodo-Dep="14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="19" pageNumber="220" startId="19.[151,250,1866,1890]" targetBox="[155,1432,185,1840]" targetPageId="19" targetType="figure">
<paragraph id="C560951AFFADFFD962EBF8AEFBF9F82A" blockId="19.[151,1436,1866,2035]" pageId="19" pageNumber="220">
<emphasis id="F7AB4908FFADFFD962EBF8AEFE83F8BA" bold="true" box="[151,295,1866,1890]" pageId="19" pageNumber="220">FIGURE 14.</emphasis>
<taxonomicName id="02DFEE99FFADFFD96348F8AFFE68F8BA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[308,460,1866,1890]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
@ -316,7 +317,7 @@ C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct.
</paragraph>
</caption>
<caption id="91A0C592FFAAFFDE62EBF917FD70F818" pageId="20" pageNumber="221" startId="20.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="20" targetType="figure">
<caption id="91A0C592FFAAFFDE62EBF917FD70F818" ID-DOI="http://doi.org/10.5281/zenodo.14702785" ID-Zenodo-Dep="14702785" httpUri="https://zenodo.org/record/14702785/files/figure.png" pageId="20" pageNumber="221" startId="20.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="20" targetType="figure">
<paragraph id="C560951AFFAAFFDE62EBF917FD70F818" blockId="20.[151,1437,1779,1984]" pageId="20" pageNumber="221">
<emphasis id="F7AB4908FFAAFFDE62EBF917FE84F8D4" bold="true" box="[151,288,1779,1804]" pageId="20" pageNumber="221">FIGURE 15.</emphasis>
<taxonomicName id="02DFEE99FFAAFFDE635BF910FE1EF8D4" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[295,442,1780,1804]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
@ -330,7 +331,7 @@ AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; d
C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum.
</paragraph>
</caption>
<caption id="91A0C592FFABFFDF62EBF927FB0AF8FC" pageId="21" pageNumber="222" startId="21.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="21" targetType="figure">
<caption id="91A0C592FFABFFDF62EBF927FB0AF8FC" ID-DOI="http://doi.org/10.5281/zenodo.14702787" ID-Zenodo-Dep="14702787" httpUri="https://zenodo.org/record/14702787/files/figure.png" pageId="21" pageNumber="222" startId="21.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="21" targetType="figure">
<paragraph id="C560951AFFABFFDF62EBF927FB0AF8FC" blockId="21.[151,1436,1731,1828]" pageId="21" pageNumber="222">
<emphasis id="F7AB4908FFABFFDF62EBF927FE87F904" bold="true" box="[151,291,1731,1756]" pageId="21" pageNumber="222">FIGURE 16.</emphasis>
<taxonomicName id="02DFEE99FFABFFDF6351F920FE66F904" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[301,450,1732,1756]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
@ -349,23 +350,23 @@ dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, media
(
<typeStatus id="1A642BB8FFA8FFDC6351FD8FFE36FD5D" box="[301,402,619,645]" pageId="22" pageNumber="223" type="paratype">paratype</typeStatus>
, MUSENUV-Ar-2679; vchLAM-251). (
<figureCitation id="5DE4899FFFA8FFDC6122FD8FFC12FD5D" box="[862,950,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[159,1423,186,1853]" captionTargetId="figure-17@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 12. Josa nadineae sp. nov., female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="22" pageNumber="223">Fig. 12</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6122FD8FFC12FD5D" box="[862,950,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[159,1423,186,1853]" captionTargetId="figure-17@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 12. Josa nadineae sp. nov., female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." figureDoi="http://doi.org/10.5281/zenodo.14702777" httpUri="https://zenodo.org/record/14702777/files/figure.png" pageId="22" pageNumber="223">Fig. 12</figureCitation>
): Total length 7.02. Carapace length 3.19, width 2.23, height 1.17. Clypeus length 0.13. Sternum length 1.59, width 1.06. Legs. Leg I: ti length 1.61/width 0.39, mt 1.21. II: ti 1.63/0.34, mt 1.26. III: ti 1.36/0.37, mt 1.56. IV: ti 1.95/0.38, mt 1.50. Spines: legs III, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 3.63, width 2.69, spiracleepigastrium 1.53. Color pattern as in male, but more pigmented, abdominal spots more defined. Sternum and labium more pigmented. Legs with defined longitudinal dark bands on dorsal edge, and many dots on ventral one. Epigyne: anterior plate sclerotized with deep and rounded anterior excavations, laterally elongated, median depression small, quadrangular, posteriorly positioned (
<figureCitation id="5DE4899FFFA8FFDC6126FC6FFC66FC7D" box="[858,962,907,933]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13E</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6126FC6FFC66FC7D" box="[858,962,907,933]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="22" pageNumber="223">Figs 13E</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC61B0FC6FFBA5FC7D" box="[972,1025,907,933]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14D</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC61B0FC6FFBA5FC7D" box="[972,1025,907,933]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="22" pageNumber="223">14D</figureCitation>
). Median field transverse, large, oval in posterior view with medial depression (
<figureCitation id="5DE4899FFFA8FFDC60FDFC4BFCACFC11" box="[641,776,943,969]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">Fig. 16EF</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC60FDFC4BFCACFC11" box="[641,776,943,969]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702787" httpUri="https://zenodo.org/record/14702787/files/figure.png" pageId="22" pageNumber="223">Fig. 16EF</figureCitation>
). Lateral lobes with sclerotized extensions wide, apically rounded (
<figureCitation id="5DE4899FFFA8FFDC637FFC37FE2CFC35" box="[259,392,979,1005]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">Fig. 16EF</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC637FFC37FE2CFC35" box="[259,392,979,1005]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702787" httpUri="https://zenodo.org/record/14702787/files/figure.png" pageId="22" pageNumber="223">Fig. 16EF</figureCitation>
). Copulatory ducts long, with a very wide posterior coil, laterally expanded in anterior side, joining with the accessory bulb ducts at medial level. Spermathecae small, rounded, in median position, oriented towards median septum (
<figureCitation id="5DE4899FFFA8FFDC63CCFBFFFDB2FBED" box="[432,534,1051,1077]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">Fig. 14E</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC63CCFBFFFDB2FBED" box="[432,534,1051,1077]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="22" pageNumber="223">Fig. 14E</figureCitation>
). Ducts of accessory bulbs long, thin, convergent (
<figureCitation id="5DE4899FFFA8FFDC6637FBFFFB6BFBED" box="[1099,1231,1051,1077]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13EF</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC6637FBFFFB6BFBED" box="[1099,1231,1051,1077]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="22" pageNumber="223">Figs 13EF</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC66A7FBFFFA82FBED" box="[1243,1318,1051,1077]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14EF</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC66A7FBFFFA82FBED" box="[1243,1318,1051,1077]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="22" pageNumber="223">14EF</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC674DFBFFFAF4FBED" box="[1329,1360,1051,1077]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">16</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC674DFBFFFAF4FBED" box="[1329,1360,1051,1077]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702787" httpUri="https://zenodo.org/record/14702787/files/figure.png" pageId="22" pageNumber="223">16</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFA8FFDC62BBFBDAFEB1FB1D" blockId="22.[151,1437,151,1257]" pageId="22" pageNumber="223">
@ -379,7 +380,7 @@ Males (n=3): total length: 5.405.75; carapace length: 2.472.97, width: 1.8
Only known from Sierra Nevada de Santa Marta,
<collectingCountry id="BDC8D58AFFA8FFDC61E9FB2BFBA7FB31" box="[917,1027,1231,1257]" name="Colombia" pageId="22" pageNumber="223">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFFA8FFDC666FFB2BFBC2FB31" box="[1043,1126,1231,1257]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="22" pageNumber="223">Fig. 31</figureCitation>
<figureCitation id="5DE4899FFFA8FFDC666FFB2BFBC2FB31" box="[1043,1126,1231,1257]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." figureDoi="http://doi.org/10.5281/zenodo.14702843" httpUri="https://zenodo.org/record/14702843/files/figure.png" pageId="22" pageNumber="223">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,65 +1,66 @@
<document id="03F5D866D325DFAB3904B0DDAC7FBC57" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFB2FFC562EBFA2FFBFAFC7D" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa samaria Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="216" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="213" updateTime="1737369942339" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
<mods:mods id="B533B0B059F21981DFB632831A22D905" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B5ACEE9A795404A9A7BF164996244042">
<mods:title id="072FF30F099D9ABC7E97D073BF3FC37F">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
<document id="4AB08DD1B006268CD4B94294FAE5B8C0" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14702734" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFB2FFC562EBFA2FFBFAFC7D" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa samaria Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="216" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="213" updateTime="1737387624970" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A2B6EBB2B3DEFA5FC69287008092103A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E3882DEB49073374B5A07262C125751D">
<mods:title id="CAD4C1F72324EBABB39C1586759C78B9">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
</mods:titleInfo>
<mods:name id="76EA0AA6F039C35E0C804951281F1F0B" type="personal">
<mods:role id="9EC84028AC9F0E42BC48BBF634F77B60">
<mods:roleTerm id="CC4049DE7A3BF564D3FF86635E2E3A37">Author</mods:roleTerm>
<mods:name id="EBF7818041E31BEA2E938F4BE9C6871F" type="personal">
<mods:role id="59EE8D1881865E24DF1D97894209C948">
<mods:roleTerm id="33D419A002052C79917FE3F9418464EA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC6B14BBA522B2314DA2AC72AF623591">Martínez, Leonel</mods:namePart>
<mods:affiliation id="B4918B6C5EB7D22AE18B30F8B741FEFA">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
<mods:namePart id="89D006CCEBE9B2D48AC4FA1E27DACCF9">Martínez, Leonel</mods:namePart>
<mods:affiliation id="3876B86C1C3ABDD8B6E5988724135F4C">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
</mods:name>
<mods:name id="1C053B8607CEEFA3FA8DBF85111BCE3A" type="personal">
<mods:role id="4C0B9F0483EFA1EAC572786527C23DEC">
<mods:roleTerm id="CDDEC1F078F28E0938A72FB86666271F">Author</mods:roleTerm>
<mods:name id="490477684B322EFF818B20ED486FFCEE" type="personal">
<mods:role id="1CE9928E5384A330370A53F6E114A3FD">
<mods:roleTerm id="2B9292B05A6CA64F0F8E3EA98DED532C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02C677A31006ECEE4AF02C3953B5A037">Kochalka, John A.</mods:namePart>
<mods:affiliation id="626AA6B3A06863DDB8A9004BA3AD9FD0">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
<mods:namePart id="616B18A0BAD6E36892B070DF862DA846">Kochalka, John A.</mods:namePart>
<mods:affiliation id="3F7E6717C0523E8FD5DE7335FA46BC47">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
</mods:name>
<mods:name id="BFAF5B6815CF706C0DB9A207A6EB6F81" type="personal">
<mods:role id="61D6CB33E145794FD00840A2D88AFB27">
<mods:roleTerm id="577A5B296DBB27E4657DCFAFE98081B9">Author</mods:roleTerm>
<mods:name id="B3302D8970A9AA8E344CFA539F46B8EE" type="personal">
<mods:role id="3E06E479DD39D9FB835F84840CEB84D9">
<mods:roleTerm id="6D63EED1B77EA05072C4E5F5BDE8AC85">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7C2A72774407AE338CEA1CDE8EFC3441">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="C54F2E77027EE90D11A756E01E7C0D85">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
<mods:namePart id="D2AFBF462E63F521BBA6C725301A5FD9">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="A46E73E26C421088FEB34041D8BFF4FE">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
</mods:name>
<mods:name id="FDDA3E0446493106435D3E32DF54657B" type="personal">
<mods:role id="0500BDA6F8BE2554A278121A7DB2232F">
<mods:roleTerm id="5E9B80419760D28A6AE1B5F50D6FFDEA">Author</mods:roleTerm>
<mods:name id="F10CB52067B4055CB4E18B8C5ED6E2BC" type="personal">
<mods:role id="D48B29461B9392E0632C400BD4478D9B">
<mods:roleTerm id="724B77EEA4CCA4E155B7A508747F9470">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9970DDCC2FF65271C8C0C87775E204C9">Ramírez, Martin</mods:namePart>
<mods:affiliation id="2F1F8D6D8FE400AFA23B69ECC168CE6D">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
<mods:namePart id="8C90ACD236EC1E8A83057632C5938232">Ramírez, Martin</mods:namePart>
<mods:affiliation id="738CC4665281195ADC3CDF6CD716EC26">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:typeOfResource id="615106C6E7CC5EBCB36832CDEC318A60">text</mods:typeOfResource>
<mods:relatedItem id="31B0656631545898581313E2753239D5" type="host">
<mods:titleInfo id="C7C1264CCEC43FB9303F7E3D08F50212">
<mods:title id="DB24DB533EDDC2C4F534C1C2E9783562">Zootaxa</mods:title>
<mods:typeOfResource id="AE064FC72AB4B7033BA58E86C0420CC6">text</mods:typeOfResource>
<mods:relatedItem id="2DF36906B1C3021C56DA78776D17DCF6" type="host">
<mods:titleInfo id="1B82A3E756321EF6965356E1408E866C">
<mods:title id="4187428414E13AF37847F1E11005CB51">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="470191B519A14000E62C9FCF69298F12">
<mods:date id="120263F218E846B3C5728673C4ADDE06">2025</mods:date>
<mods:detail id="91B508B81A0FDD651579D1A9DB1B00FC" type="pubDate">
<mods:number id="605E38C5481538B174BF200D7E70B69D">2025-01-08</mods:number>
<mods:part id="EE3FB88026613E30310E01798287A890">
<mods:date id="ABCD72B1382F40758F7ED126EC92094D">2025</mods:date>
<mods:detail id="8BFFEC69D4AF279DC7E7D75E5771DB83" type="pubDate">
<mods:number id="E32B56D15717A2FBACB72205816B36BE">2025-01-08</mods:number>
</mods:detail>
<mods:detail id="756E3969BDFA13E7755BB0403062E4D6" type="volume">
<mods:number id="316326B3915B1982DE94F3DAC65ABEDE">5566</mods:number>
<mods:detail id="363479F28A9AEC6C07DBD0324A3D8F67" type="volume">
<mods:number id="D0120265DE88551E3C6BD0AC2DAE222B">5566</mods:number>
</mods:detail>
<mods:detail id="38F79E9DC616E9B2F73EEDF4F8DEFC1C" type="issue">
<mods:number id="DD9A8E3FE0A699DF74C9656686657F94">2</mods:number>
<mods:detail id="EB6321C9EB32A9361439C4142436DC63" type="issue">
<mods:number id="9CE2436CDDC9E2D5A153309E8000078D">2</mods:number>
</mods:detail>
<mods:extent id="4D1FF25678611594EB01513CB11E757A" unit="page">
<mods:start id="A6710F5D12C4381671EC611A64729AB1">201</mods:start>
<mods:end id="E024E2F7F06DD8F318C82A7554DF602D">242</mods:end>
<mods:extent id="B3D299D743837A4167852CBF22C3E737" unit="page">
<mods:start id="95A2751796B9DE199674454CB74049F0">201</mods:start>
<mods:end id="B27BA507E2DD43D9F8E2AADA6F54C71F">242</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4AAEF8AF767DBEA0E1863E89DB8B76AF">
<mods:url id="AF3C6375B8839028F1496C4E3C395A1E">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
<mods:location id="9067A5EBCE630E2B6226A16225F004D8">
<mods:url id="933EF9105C067BB6CEFEC47FAA75EA05">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
</mods:location>
<mods:classification id="C0EEF232EA2C1B5AF6C401722CE1CFA7">journal article</mods:classification>
<mods:identifier id="12FAD4CEE0D47FE74B0822F0242DDFCA" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="7B43BF11640B1954F0F9B02F86ADE00C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B880A91DE246FDD4DFBB2BEF1B114436" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
<mods:classification id="9796D663BF062E682329F1C3692F1285">journal article</mods:classification>
<mods:identifier id="2F5B5720B049607C698991F69AFD3411" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="CED554FDCC7649ACC0B28943A45F640B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="812E7C8EA5734C4CAA224DA4526C5786" type="Zenodo-Dep">14702734</mods:identifier>
<mods:identifier id="CBD07CBA5BE9FF514688820D3276C833" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
</mods:mods>
<treatment id="4D76240CFFB2FFC562EBFA2FFBFAFC7D" LSID="urn:lsid:plazi:treatment:4D76240CFFB2FFC562EBFA2FFBFAFC7D" httpUri="http://treatment.plazi.org/id/4D76240CFFB2FFC562EBFA2FFBFAFC7D" lastPageId="15" lastPageNumber="216" pageId="12" pageNumber="213">
<subSubSection id="8DC5C691FFB2FFC662EBFA2FFE39FA3D" box="[151,413,1483,1509]" pageId="12" pageNumber="213" type="nomenclature">
@ -76,7 +77,7 @@
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662EBFA0BFE8EF9D1" box="[151,298,1519,1545]" pageId="12" pageNumber="213" type="description">
<paragraph id="C560951AFFB2FFC662EBFA0BFE8EF9D1" blockId="12.[151,413,1483,1545]" box="[151,298,1519,1545]" pageId="12" pageNumber="213">
<figureCitation id="5DE4899FFFB2FFC662EBFA0BFE8EF9D1" box="[151,298,1519,1545]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="13.[151,250,1867,1891]" captionStartId-1="14.[151,250,1785,1809]" captionTargetBox-0="[167,1426,189,1841]" captionTargetBox-1="[151,1417,191,1757]" captionTargetId-0="figure-17@13.[151,1436,181,1844]" captionTargetId-1="figure-22@14.[151,1436,181,1761]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 9. Josa samaria sp. nov., female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="12" pageNumber="213">Figures 910</figureCitation>
<figureCitation id="5DE4899FFFB2FFC662EBFA0BFE8EF9D1" box="[151,298,1519,1545]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="13.[151,250,1867,1891]" captionStartId-1="14.[151,250,1785,1809]" captionTargetBox-0="[167,1426,189,1841]" captionTargetBox-1="[151,1417,191,1757]" captionTargetId-0="figure-17@13.[151,1436,181,1844]" captionTargetId-1="figure-22@14.[151,1436,181,1761]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 9. Josa samaria sp. nov., female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.14702767" figureDoi-1="http://doi.org/10.5281/zenodo.14702769" httpUri-0="https://zenodo.org/record/14702767/files/figure.png" httpUri-1="https://zenodo.org/record/14702769/files/figure.png" pageId="12" pageNumber="213">Figures 910</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662EBF9D7FABFF993" box="[151,1307,1587,1611]" pageId="12" pageNumber="213" type="reference_group">
@ -142,11 +143,11 @@ The female is similar to that of
<taxonomicNameLabel id="EC98F473FFB2FFC66120F8EDFC12F8FB" box="[860,950,1801,1827]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the posterior quadrangular notch of the epigyne (
<figureCitation id="5DE4899FFFB2FFC6637EF8C9FECDF89F" box="[258,361,1837,1863]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Fig. 13E</figureCitation>
<figureCitation id="5DE4899FFFB2FFC6637EF8C9FECDF89F" box="[258,361,1837,1863]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="12" pageNumber="213">Fig. 13E</figureCitation>
, compare with
<figureCitation id="5DE4899FFFB2FFC66060F8C9FD22F89F" box="[540,646,1837,1863]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Figs 13E</figureCitation>
<figureCitation id="5DE4899FFFB2FFC66060F8C9FD22F89F" box="[540,646,1837,1863]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." figureDoi="http://doi.org/10.5281/zenodo.14702781" httpUri="https://zenodo.org/record/14702781/files/figure.png" pageId="12" pageNumber="213">Figs 13E</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC660E8F8C9FD62F89F" box="[660,710,1837,1863]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">14D</figureCitation>
<figureCitation id="5DE4899FFFB2FFC660E8F8C9FD62F89F" box="[660,710,1837,1863]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." figureDoi="http://doi.org/10.5281/zenodo.14702783" httpUri="https://zenodo.org/record/14702783/files/figure.png" pageId="12" pageNumber="213">14D</figureCitation>
) but can be distinguished by the posterior lateral margins on a straight transversal line (vs. oblique in
<taxonomicName id="02DFEE99FFB2FFC66028F8B6FD53F8B3" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[596,759,1873,1899]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFB2FFC66028F8B6FD53F8B3" box="[596,759,1873,1899]" italics="true" pageId="12" pageNumber="213">Josa nadineae</emphasis>
@ -156,11 +157,11 @@ by the posterior quadrangular notch of the epigyne (
)
</emphasis>
, and by the copulatory ducts describing two wide posterior coils, and the small spermathecae, anteromedially placed with long fertilization ducts (
<figureCitation id="5DE4899FFFB2FFC666B0F891FAF1F857" box="[1228,1365,1909,1935]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="12" pageNumber="213">Fig. 10AE</figureCitation>
<figureCitation id="5DE4899FFFB2FFC666B0F891FAF1F857" box="[1228,1365,1909,1935]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702769" httpUri="https://zenodo.org/record/14702769/files/figure.png" pageId="12" pageNumber="213">Fig. 10AE</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="91A0C592FFB3FFC762EBF8AFFE55F850" pageId="13" pageNumber="214" startId="13.[151,250,1867,1891]" targetBox="[167,1426,189,1841]" targetPageId="13" targetType="figure">
<caption id="91A0C592FFB3FFC762EBF8AFFE55F850" ID-DOI="http://doi.org/10.5281/zenodo.14702767" ID-Zenodo-Dep="14702767" httpUri="https://zenodo.org/record/14702767/files/figure.png" pageId="13" pageNumber="214" startId="13.[151,250,1867,1891]" targetBox="[167,1426,189,1841]" targetPageId="13" targetType="figure">
<paragraph id="C560951AFFB3FFC762EBF8AFFE55F850" blockId="13.[151,1436,1867,1928]" pageId="13" pageNumber="214">
<emphasis id="F7AB4908FFB3FFC762EBF8AFFEB0F8BC" bold="true" box="[151,276,1867,1892]" pageId="13" pageNumber="214">FIGURE 9.</emphasis>
<taxonomicName id="02DFEE99FFB3FFC76361F8A8FE02F8BB" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[285,422,1868,1891]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="samaria" status="sp. nov.">
@ -172,7 +173,7 @@ by the posterior quadrangular notch of the epigyne (
, female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFB0FFC462EBF91DFDE7F8A5" pageId="14" pageNumber="215" startId="14.[151,250,1785,1809]" targetBox="[151,1417,191,1757]" targetPageId="14" targetType="figure">
<caption id="91A0C592FFB0FFC462EBF91DFDE7F8A5" ID-DOI="http://doi.org/10.5281/zenodo.14702769" ID-Zenodo-Dep="14702769" httpUri="https://zenodo.org/record/14702769/files/figure.png" pageId="14" pageNumber="215" startId="14.[151,250,1785,1809]" targetBox="[151,1417,191,1757]" targetPageId="14" targetType="figure">
<paragraph id="C560951AFFB0FFC462EBF91DFDE7F8A5" blockId="14.[151,1436,1785,1918]" pageId="14" pageNumber="215">
<emphasis id="F7AB4908FFB0FFC462EBF91DFE86F8C9" bold="true" box="[151,290,1785,1809]" pageId="14" pageNumber="215">FIGURE 10.</emphasis>
<taxonomicName id="02DFEE99FFB0FFC46357F91EFE11F8C9" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,437,1786,1809]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="samaria" status="sp. nov.">
@ -193,15 +194,15 @@ AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of ac
(
<typeStatus id="1A642BB8FFB1FFC563BDFF73FD83FF69" box="[449,551,151,177]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
, MNHP; vchLAM-244) (
<figureCitation id="5DE4899FFFB1FFC5613EFF73FC22FF69" box="[834,902,151,177]" captionStart="FIGURE 9" captionStartId="13.[151,250,1867,1891]" captionTargetBox="[167,1426,189,1841]" captionTargetId="figure-17@13.[151,1436,181,1844]" captionTargetPageId="13" captionText="FIGURE 9. Josa samaria sp. nov., female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="15" pageNumber="216">Fig. 9</figureCitation>
<figureCitation id="5DE4899FFFB1FFC5613EFF73FC22FF69" box="[834,902,151,177]" captionStart="FIGURE 9" captionStartId="13.[151,250,1867,1891]" captionTargetBox="[167,1426,189,1841]" captionTargetId="figure-17@13.[151,1436,181,1844]" captionTargetPageId="13" captionText="FIGURE 9. Josa samaria sp. nov., female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." figureDoi="http://doi.org/10.5281/zenodo.14702767" httpUri="https://zenodo.org/record/14702767/files/figure.png" pageId="15" pageNumber="216">Fig. 9</figureCitation>
): Total length 7.33. Carapace length 3.41, width 2.36, height 1.35. Clypeus length 0.19. Sternum length 1.79, width 1.31. Legs. Leg I: ti length 1.84/width 0.42, metatarsus (mt) 1.45. II: ti 1.74/0.40, mt 1.39. III: ti 1.59/0.38, mt 1.55. IV: ti 2.17/0.44, mt 2.41. Spines: legs III, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2r-1-2, p 1-d1-1-0 r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. IV=III but metatarsus v 2-2-2. Abdomen: length 4.03, width 2.33, spiracleepigastrium 0.80. Color: carapace brown, with two wide, longitudinal darker band on sides, narrow in cephalic region. Ocular area on black patch with eyes bordered by black rings. Carapace with lateral region covered by fringe of white setae. Clypeus brown, low with abundant white setae. Chelicerae brown with black patches and fine setae on frontal edge. Endites, labium, and sternum dark brown, labium and endites light in apical region. Sternum with a very thin medial pale stripe. Legs coxae-tarsi dark brown, femora and patellae with longitudinal dark bands. All articles covered by white setae, abundant between coxae. Abdomen with dark brown pattern of spots on a gray background, dorsally with white medial band followed by 4-5 faint light chevrons; lateral regions covered by abundant small dots. Abdomen laterally surrounded by abundant white setae, with a thick bunch of setae on anterior region. Venter uniformly covered by dark and light patches. Epigyne: anterior plate sclerotized with deep and rounded anterior excavations, median notch quadrangular, posteriorly positioned (
<figureCitation id="5DE4899FFFB1FFC563E6FD57FD80FD15" box="[410,548,691,717]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10A, D</figureCitation>
<figureCitation id="5DE4899FFFB1FFC563E6FD57FD80FD15" box="[410,548,691,717]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702769" httpUri="https://zenodo.org/record/14702769/files/figure.png" pageId="15" pageNumber="216">Fig. 10A, D</figureCitation>
). Median field transverse, large, oval in posterior view (
<figureCitation id="5DE4899FFFB1FFC566EEFD57FB5DFD15" box="[1170,1273,691,717]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10C</figureCitation>
<figureCitation id="5DE4899FFFB1FFC566EEFD57FB5DFD15" box="[1170,1273,691,717]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702769" httpUri="https://zenodo.org/record/14702769/files/figure.png" pageId="15" pageNumber="216">Fig. 10C</figureCitation>
). Lateral lobes with sclerotized extensions wide, forming, posterior extensions rounded at base (
<figureCitation id="5DE4899FFFB1FFC56662FD33FB6EFD29" box="[1054,1226,727,753]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10A, CD</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56662FD33FB6EFD29" box="[1054,1226,727,753]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702769" httpUri="https://zenodo.org/record/14702769/files/figure.png" pageId="15" pageNumber="216">Fig. 10A, CD</figureCitation>
). Copulatory ducts long, forming two basal coils, anteriorly medially expanded, joining with the accessory bulb ducts at medial level. Spermathecae small, rounded, medially positioned in the epigyne, oriented towards longitudinal plane. Ducts of accessory bulbs short, wide, convergent (
<figureCitation id="5DE4899FFFB1FFC56018FCA7FD49FC85" box="[612,749,835,861]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10AE</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56018FCA7FD49FC85" box="[612,749,835,861]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14702769" httpUri="https://zenodo.org/record/14702769/files/figure.png" pageId="15" pageNumber="216">Fig. 10AE</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFB1FFC562BBFC83FE21FC59" blockId="15.[151,1437,150,933]" box="[199,389,871,897]" pageId="15" pageNumber="216">
@ -215,7 +216,7 @@ Unknown.
Only known for Sierra Nevada de Santa Marta,
<collectingCountry id="BDC8D58AFFB1FFC56103FC6FFC49FC7D" box="[895,1005,907,933]" name="Colombia" pageId="15" pageNumber="216">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFFB1FFC56181FC6FFBF4FC7D" box="[1021,1104,907,933]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="15" pageNumber="216">Fig. 31</figureCitation>
<figureCitation id="5DE4899FFFB1FFC56181FC6FFBF4FC7D" box="[1021,1104,907,933]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." figureDoi="http://doi.org/10.5281/zenodo.14702843" httpUri="https://zenodo.org/record/14702843/files/figure.png" pageId="15" pageNumber="216">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>

File diff suppressed because one or more lines are too long