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<document id="5B0235D4A0125C2AB467B87F499D7BB4" ID-DOI="10.11646/phytotaxa.253.2.6" ID-ISSN="1179-3163" ID-Zenodo-Dep="13676033" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1725407256268" checkinUser="felipe" docAuthor="Crespo, Manuel B. & Alonso, Ángeles" docDate="2016" docId="03A64112C9064F01729EF8C4FAF5B928" docLanguage="en" docName="phytotaxa.253.2.6.pdf" docOrigin="Phytotaxa 253 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.253.2.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Ximenesia encelioides , Cavanilles 1793" docType="treatment" docVersion="1" lastPageNumber="164" masterDocId="FF9F396AC9064F027216FFF6FFF5BC57" masterDocTitle="Remarks and clarifications on the type material of Ximenesia encelioides (Verbesina encelioides; Asteraceae, Heliantheae), with a new lectotype designation" masterLastPageNumber="165" masterPageNumber="161" pageNumber="161" updateTime="1733828708999" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="7BF507691AF63C6AD482C9D087343ECE">
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<mods:title id="23B4435B63C89F426F9AEA013631CB9B">Remarks and clarifications on the type material of Ximenesia encelioides (Verbesina encelioides; Asteraceae, Heliantheae), with a new lectotype designation</mods:title>
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<mods:namePart id="C2F03E6F45B9D9B786C13E8F88B4F4A3">Crespo, Manuel B.</mods:namePart>
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<mods:namePart id="298B4192AA2EE4857EB731E84C7F7BD4">Alonso, Ángeles</mods:namePart>
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<mods:title id="7191EC3F5B4BCBBC49C85E234DDD9CD8">Phytotaxa</mods:title>
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<mods:date id="E20C1395F614A53F89E1B05543D51236">2016</mods:date>
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<mods:number id="A4EAB87E6FB35CBBC35D5EBFAC4EA10B">2016-03-23</mods:number>
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<mods:number id="F263A1E348821665CCB9FF9FAB57F336">253</mods:number>
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<mods:identifier id="0EB0B9B93A2CA6599E7E3DAAE34179CD" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03A64112C9064F01729EF8C4FAF5B928" LSID="urn:lsid:plazi:treatment:03A64112C9064F01729EF8C4FAF5B928" httpUri="http://treatment.plazi.org/id/03A64112C9064F01729EF8C4FAF5B928" lastPageId="3" lastPageNumber="164" pageId="0" pageNumber="161">
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<subSubSection id="C315A38FC9064F02729EF8C4FC7DBB1C" box="[136,904,1842,1867]" pageId="0" pageNumber="161" type="nomenclature">
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<paragraph id="8BB0F004C9064F02729EF8C4FC7DBB1C" blockId="0.[136,1452,1805,1903]" box="[136,904,1842,1867]" pageId="0" pageNumber="161">
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<taxonomicName id="4C0F8B87C9064F02729EF8C4FD8ABB1C" ID-CoL="7G9NK" authority="Cavanilles (1793: 60)" authorityName=", Cavanilles" authorityPageNumber="60" authorityYear="1793" box="[136,639,1842,1867]" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="0" pageNumber="161" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9064F02729EF8C4FE76BB1C" box="[136,387,1842,1867]" italics="true" pageId="0" pageNumber="161">Ximenesia encelioides</emphasis>
|
||||
<bibRefCitation id="EF9E8DF5C9064F02739DF8C4FD8ABB1C" author="Cavanilles, A. J." box="[395,639,1842,1867]" pageId="0" pageNumber="161" refId="ref2168" refString="Cavanilles, A. J. (1793) Icones et descriptiones plantarum, quae aut sponte in Hispania crescunt, aut in hortis hospitantur, vol. 2 (3). Regia Typographia, Matriti [Madrid], 83 pp., 100 pl." type="book" year="1793">Cavanilles (1793: 60)</bibRefCitation>
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||||
</taxonomicName>
|
||||
[sphalm. “
|
||||
<emphasis id="B97B2C16C9064F0270EBF8C4FC86BB1C" box="[765,883,1842,1867]" italics="true" pageId="0" pageNumber="161">enceloides</emphasis>
|
||||
”]
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C315A38FC9064F027187F8C2FE7FBB38" pageId="0" pageNumber="161" type="reference_group">
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<paragraph id="8BB0F004C9064F027187F8C2FE7FBB38" blockId="0.[136,1452,1805,1903]" pageId="0" pageNumber="161">
|
||||
<emphasis id="B97B2C16C9064F027187F8C2FC54BB1D" box="[913,929,1844,1866]" italics="true" pageId="0" pageNumber="161">≡</emphasis>
|
||||
<taxonomicName id="4C0F8B87C9064F0271BFF8C6FE7FBB38" authority="(Cav.) Benth. & Hook. f. ex Gray (1876: 350)" authorityName="Benth. & Hook. f. ex A. Gray" authorityPageNumber="350" authorityYear="1876" baseAuthorityName="Cav." class="Magnoliopsida" family="Asteraceae" genus="Verbesina" kingdom="Plantae" order="Asterales" pageId="0" pageNumber="161" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9064F0271BFF8C6FB57BB1C" bold="true" box="[937,1186,1840,1867]" italics="true" pageId="0" pageNumber="161">Verbesina encelioides</emphasis>
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||||
(Cav.) Benth. & Hook. f. ex
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||||
<bibRefCitation id="EF9E8DF5C9064F0272DEF8A0FE7FBB38" author="Gray, A." box="[200,394,1878,1903]" pageId="0" pageNumber="161" pagination="288 - 443" refId="ref2310" refString="Gray, A. (1876) Compositae. In: Whitney, J. D. (Ed.) Geological survey of California. Botany, vol. 1. Welch, Bigelow & Co., University Press, Cambridge, Massachusetts, pp. 288 - 443." type="book chapter" year="1876">Gray (1876: 350)</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C315A38FC9064F02729EF856FC68BBE2" box="[136,925,1952,1973]" pageId="0" pageNumber="161" type="distribution">
|
||||
<paragraph id="8BB0F004C9064F02729EF856FC68BBE2" blockId="0.[136,1095,1952,2009]" box="[136,925,1952,1973]" pageId="0" pageNumber="161">
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||||
<emphasis id="B97B2C16C9064F02729EF856FF2EBBE2" bold="true" box="[136,219,1952,1973]" pageId="0" pageNumber="161">Ind. loc.</emphasis>
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||||
:—“Habitat in
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||||
<collectingRegion id="49CB3EE6C9064F02737CF856FE43BBE2" box="[362,438,1952,1973]" country="Mexico" name="Mexico" pageId="0" pageNumber="161">Mexico</collectingRegion>
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, unde introducta in Regium hortum Matritensem”.
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C315A38FC9064F02729EF832FBB2BB8E" box="[136,1095,1988,2009]" pageId="0" pageNumber="161" type="materials_examined">
|
||||
<paragraph id="8BB0F004C9064F02729EF832FBB2BB8E" blockId="0.[136,1095,1952,2009]" box="[136,1095,1988,2009]" pageId="0" pageNumber="161">
|
||||
<emphasis id="B97B2C16C9064F02729EF832FF18BB8E" bold="true" box="[136,237,1988,2009]" pageId="0" pageNumber="161">
|
||||
<typeStatus id="54B44EA6C9064F02729EF832FF18BB8E" box="[136,237,1988,2009]" pageId="0" pageNumber="161" type="lectotype">Lectotype</typeStatus>
|
||||
</emphasis>
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||||
(designated here):—[icon]
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<bibRefCitation id="EF9E8DF5C9064F0273E1F832FD6BBB8E" author="Cavanilles, A. J." box="[503,670,1988,2009]" pageId="0" pageNumber="161" refId="ref2168" refString="Cavanilles, A. J. (1793) Icones et descriptiones plantarum, quae aut sponte in Hispania crescunt, aut in hortis hospitantur, vol. 2 (3). Regia Typographia, Matriti [Madrid], 83 pp., 100 pl." type="book" year="1793">Cavanilles (1793</bibRefCitation>
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||||
: tab. 178), “
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<taxonomicName id="4C0F8B87C9064F027105F832FC1CBB8E" authorityName=", Cavanilles" authorityYear="1793" box="[787,1001,1988,2009]" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="0" pageNumber="161" phylum="Tracheophyta" rank="species" species="encelioides">
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||||
<emphasis id="B97B2C16C9064F027105F832FC1CBB8E" box="[787,1001,1988,2009]" italics="true" pageId="0" pageNumber="161">Ximenesia encelioides</emphasis>
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</taxonomicName>
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” (
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<figureCitation id="1334EC81C9064F027617F832FBCCBB8E" box="[1025,1081,1988,2009]" captionStart="FIGURE 1" captionStartId="1.[136,229,1925,1946]" captionTargetBox="[151,1436,207,1881]" captionTargetId="figure-14@1.[151,1436,190,1900]" captionTargetPageId="1" captionText="FIGURE 1. Ximenesia encelioides Cav. [Verbesina encelioides (Cav.) Benth.& Hook.f. exA.Gray]. Illustration in the protologue (as “Tab. 178”), designated here as lectotype of the name. Reproduced with permission (Biblioteca del Real Jardín Botánico, CSIC, Madrid)." figureDoi="http://doi.org/10.5281/zenodo.13676035" httpUri="https://zenodo.org/record/13676035/files/figure.png" pageId="0" pageNumber="161">Fig. 1</figureCitation>
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).
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</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF70A08CC9074F03729EF873FA80BBE9" ID-DOI="http://doi.org/10.5281/zenodo.13676035" ID-Zenodo-Dep="13676035" httpUri="https://zenodo.org/record/13676035/files/figure.png" pageId="1" pageNumber="162" startId="1.[136,229,1925,1946]" targetBox="[151,1436,207,1881]" targetPageId="1" targetType="figure">
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||||
<paragraph id="8BB0F004C9074F03729EF873FA80BBE9" blockId="1.[136,1452,1925,1982]" pageId="1" pageNumber="162">
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<emphasis id="B97B2C16C9074F03729EF873FF08BBCD" bold="true" box="[136,253,1925,1946]" pageId="1" pageNumber="162">FIGURE 1.</emphasis>
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<taxonomicName id="4C0F8B87C9074F037317F873FDF3BBCD" authority="Cav." authorityName=", Cavanilles" authorityYear="1793" box="[257,518,1925,1946]" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="encelioides">
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||||
<emphasis id="B97B2C16C9074F037317F873FE20BBCD" box="[257,469,1925,1946]" italics="true" pageId="1" pageNumber="162">Ximenesia encelioides</emphasis>
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Cav.
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</taxonomicName>
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[
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<taxonomicName id="4C0F8B87C9074F037004F873FBC0BBCD" authority="(Cav.) Benth. & Hook. f. ex A. Gray" authorityName="Benth. & Hook. f. ex A. Gray" authorityYear="1876" baseAuthorityName="Cav." box="[530,1077,1925,1946]" class="Magnoliopsida" family="Asteraceae" genus="Verbesina" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9074F037004F873FD14BBCD" box="[530,737,1925,1946]" italics="true" pageId="1" pageNumber="162">Verbesina encelioides</emphasis>
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(Cav.) Benth. & Hook.f. ex A.Gray
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</taxonomicName>
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]. Illustration in the protologue (as “Tab. 178”), designated here as lectotype of the name. Reproduced with permission (Biblioteca del Real Jardín Botánico, CSIC, Madrid).
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</paragraph>
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</caption>
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<subSubSection id="C315A38FC9044F01729EFF56FAF5B928" lastPageId="3" lastPageNumber="164" pageId="2" pageNumber="163" type="discussion">
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<paragraph id="8BB0F004C9044F00729EFF56FE8CBCA8" blockId="2.[136,1453,160,759]" pageId="2" pageNumber="163">
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||||
<emphasis id="B97B2C16C9044F00729EFF56FF3DBCE0" bold="true" box="[136,200,160,183]" pageId="2" pageNumber="163">Notes</emphasis>
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:—It is worth mentioning that contrarily to argumentation by
|
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<bibRefCitation id="EF9E8DF5C9044F00717FFF56FC1BBCE0" author="Hind, D. J. N." box="[873,1006,160,183]" pageId="2" pageNumber="163" refId="ref2362" refString="Hind, D. J. N. (2011) An annotated preliminary checklist of the Compositae of Bolivia. Vers. 2. Available from: www. kew. org / science / tropamerica / boliviacompositae / checklist. pdf (accessed January 2016)" type="journal volume" year="2011">Hind (2011)</bibRefCitation>
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and according to Art. 46.5 Ex. 29 of the ICN, the full authority of the combination
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<taxonomicName id="4C0F8B87C9044F00705CFF32FCC4BC8C" authorityName="Benth. & Hook. f. ex A. Gray" authorityYear="1876" baseAuthorityName="Cav." box="[586,817,196,219]" class="Magnoliopsida" family="Asteraceae" genus="Verbesina" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="163" phylum="Tracheophyta" rank="species" species="encelioides">
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<emphasis id="B97B2C16C9044F00705CFF32FCC4BC8C" box="[586,817,196,219]" italics="true" pageId="2" pageNumber="163">Verbesina encelioides</emphasis>
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</taxonomicName>
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should be cited “Benth. & Hook.f. ex A.Gray”, as currently shown in
|
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<bibRefCitation id="EF9E8DF5C9044F0072E6FF1EFE80BCA8" author="IPNI" box="[240,373,232,255]" pageId="2" pageNumber="163" refId="ref2410" refString="IPNI (2016) International Plant Names Index. Available from: http: // www. ipni. org (accessed January 2016)" type="url" year="2016">IPNI (2016)</bibRefCitation>
|
||||
.
|
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</paragraph>
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<paragraph id="8BB0F004C9044F0072ABFEFAFC2EBD3C" blockId="2.[136,1453,160,759]" pageId="2" pageNumber="163">
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Cavanilles’s material at MA was first housed among the historical collections (“
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<emphasis id="B97B2C16C9044F00760EFEFAFA96BD74" box="[1048,1379,268,291]" italics="true" pageId="2" pageNumber="163">Antiquum generale Herbarium</emphasis>
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”), and later placed in the collection “
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<emphasis id="B97B2C16C9044F0073D3FEC6FD5BBD10" box="[453,686,304,327]" italics="true" pageId="2" pageNumber="163">Typi Cavanillesianum</emphasis>
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” (http://herbario.rjb.csic.es/). Those materials were manually numbered in 1990, and later digitalised and barcoded in
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<date id="FFB1D6C4C9044F007079FEA2FD02BD3C" box="[623,759,340,363]" pageId="2" pageNumber="163" value="2009-08">August 2009</date>
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(Noya pers. comm.).
|
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</paragraph>
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<paragraph id="8BB0F004C9044F0072ABFE8EFBD4BEDC" blockId="2.[136,1453,160,759]" pageId="2" pageNumber="163">
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In a study of Cavanilles’s historical collections at MA,
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<bibRefCitation id="EF9E8DF5C9044F007126FE8EFC2EBDD8" author="Garilleti, R." box="[816,987,376,399]" pageId="2" pageNumber="163" pagination="6 - 248" refId="ref2271" refString="Garilleti, R. (1993) Herbarium cavanillesianum seu enumeratio plantarum exsiccatarum aliquo modo ad novitatis cavanillesianas pertinentium, quae in Horti Regii Matritensis atque Londinensis Societatis Linnaeanae herbariis asservantur. Fontqueria 38: 6 - 248." type="journal article" year="1993">Garilleti (1993)</bibRefCitation>
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remarked that part of the material in the voucher MA-00476508!, which then included two unmarked sheets, should correspond to the gathering indicated in the protologue, and therefore should be considered original material. However, as a result of recent digitalisation, both sheets are currently barcoded MA-00476508 and MA-00476508–2 (
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<figureCitation id="1334EC81C9044F0070E4FE12FCC6BDAC" box="[754,819,484,507]" captionStart="FIGURE 2" captionStartId="2.[136,229,1794,1815]" captionTargetBox="[163,1426,794,1761]" captionTargetId="figure-294@2.[151,1436,784,1768]" captionTargetPageId="2" captionText="FIGURE 2.Part of the probable original material of Ximenesia encelioides Cav.[Verbesinaencelioides (Cav.) Benth.&Hook.f.exA.Gray], including three different gatherings. A. Voucher MA-00476508! with material lacking original labels; B. Voucher MA-00476508−2! which bears the historical label handwritten by Cavanilles, among others. Reproduced with permission (Herbario del Real Jardín Botánico, CSIC, Madrid)." figureDoi="http://doi.org/10.5281/zenodo.13676039" httpUri="https://zenodo.org/record/13676039/files/figure.png" pageId="2" pageNumber="163">Fig. 2</figureCitation>
|
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), a sequence that is opposite to that of
|
||||
<bibRefCitation id="EF9E8DF5C9044F0076C5FE12FA8FBDAC" author="Garilleti, R." box="[1235,1402,484,507]" pageId="2" pageNumber="163" pagination="6 - 248" refId="ref2271" refString="Garilleti, R. (1993) Herbarium cavanillesianum seu enumeratio plantarum exsiccatarum aliquo modo ad novitatis cavanillesianas pertinentium, quae in Horti Regii Matritensis atque Londinensis Societatis Linnaeanae herbariis asservantur. Fontqueria 38: 6 - 248." type="journal article" year="1993">Garilleti (1993)</bibRefCitation>
|
||||
(see additional details in
|
||||
<bibRefCitation id="EF9E8DF5C9044F007375FDFEFDE6BE48" author="Juan, A. & Hind, D. J. N. & Alonso, M. A." box="[355,531,520,543]" pageId="2" pageNumber="163" pagination="298 - 300" refId="ref2435" refString="Juan, A., Hind, D. J. N. & Alonso, M. A. (2015) Lectotypification of the name Ximenesia encelioides, basionym of Verbesina encelioides (Asteraceae: Heliantheae: Verbesininae). Phytotaxa 238 (3): 298 - 300. http: // dx. doi. org / 10.11646 / phytotaxa. 238.3.11" type="journal article" year="2015">
|
||||
Juan
|
||||
<emphasis id="B97B2C16C9044F00738DFDFEFE27BE48" box="[411,466,520,543]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
|
||||
2015
|
||||
</bibRefCitation>
|
||||
). Other two additional vouchers of
|
||||
<taxonomicName id="4C0F8B87C9044F007186FDFEFB43BE48" authority="Cav." authorityName=", Cavanilles" authorityYear="1793" box="[912,1206,520,543]" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="163" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9044F007186FDFEFB8BBE48" box="[912,1150,520,543]" italics="true" pageId="2" pageNumber="163">Ximenesia encelioides</emphasis>
|
||||
Cav.
|
||||
</taxonomicName>
|
||||
also conserved among the Cavanillesian types at
|
||||
<collectingRegion id="49CB3EE6C9044F0073B0FDDAFE00BE14" box="[422,501,556,579]" country="Spain" name="Madrid" pageId="2" pageNumber="163">Madrid</collectingRegion>
|
||||
(MA barcodes MA-00476509! and MA-00476510!; images available at http://herbario. rjb.csic.es/imagenes2/basesdedatos.html) are not relevant for typification since they were collected after publication of that name in 1793, though bring interesting data for
|
||||
<typeStatus id="54B44EA6C9044F007090FD82FD1DBEDC" box="[646,744,628,651]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
|
||||
designation as shown below.
|
||||
</paragraph>
|
||||
<paragraph id="8BB0F004C9044F0072ABFD6EFA7CBEA0" blockId="2.[136,1453,160,759]" pageId="2" pageNumber="163">
|
||||
In the protologue of
|
||||
<taxonomicName id="4C0F8B87C9044F00738BFD6EFCF7BEF8" authority=", Cavanilles (1793)" authorityName=", Cavanilles" authorityYear="1793" box="[413,770,664,687]" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="163" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9044F00738BFD6EFDC1BEF8" box="[413,564,664,687]" italics="true" pageId="2" pageNumber="163">X. encelioides</emphasis>
|
||||
,
|
||||
<bibRefCitation id="EF9E8DF5C9044F007054FD6EFCF7BEF8" author="Cavanilles, A. J." box="[578,770,664,687]" pageId="2" pageNumber="163" refId="ref2168" refString="Cavanilles, A. J. (1793) Icones et descriptiones plantarum, quae aut sponte in Hispania crescunt, aut in hortis hospitantur, vol. 2 (3). Regia Typographia, Matriti [Madrid], 83 pp., 100 pl." type="book" year="1793">Cavanilles (1793)</bibRefCitation>
|
||||
</taxonomicName>
|
||||
did not cite either
|
||||
<typeStatus id="54B44EA6C9044F0071C7FD6EFBDBBEF8" box="[977,1070,664,687]" pageId="2" pageNumber="163" type="holotype">holotype</typeStatus>
|
||||
or
|
||||
<typeStatus id="54B44EA6C9044F007642FD6EFB40BEF8" box="[1108,1205,664,687]" pageId="2" pageNumber="163" type="syntype">syntypes</typeStatus>
|
||||
. Therefore the original material of that name, from which a new
|
||||
<typeStatus id="54B44EA6C9044F00702AFD4AFD6BBE84" box="[572,670,700,723]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
|
||||
should be designated, is constituted by the illustration in “Tab. 178” (
|
||||
<figureCitation id="1334EC81C9044F007796FD4AFF61BEA0" captionStart="FIGURE 1" captionStartId="1.[136,229,1925,1946]" captionTargetBox="[151,1436,207,1881]" captionTargetId="figure-14@1.[151,1436,190,1900]" captionTargetPageId="1" captionText="FIGURE 1. Ximenesia encelioides Cav. [Verbesina encelioides (Cav.) Benth.& Hook.f. exA.Gray]. Illustration in the protologue (as “Tab. 178”), designated here as lectotype of the name. Reproduced with permission (Biblioteca del Real Jardín Botánico, CSIC, Madrid)." figureDoi="http://doi.org/10.5281/zenodo.13676035" httpUri="https://zenodo.org/record/13676035/files/figure.png" pageId="2" pageNumber="163">Fig. 1</figureCitation>
|
||||
) and some of the fragments in the sheets kept in MA currently barcoded MA-00476508 and MA-00476508–2 (
|
||||
<figureCitation id="1334EC81C9044F00772DFD16FA89BEA0" box="[1339,1404,736,759]" captionStart="FIGURE 2" captionStartId="2.[136,229,1794,1815]" captionTargetBox="[163,1426,794,1761]" captionTargetId="figure-294@2.[151,1436,784,1768]" captionTargetPageId="2" captionText="FIGURE 2.Part of the probable original material of Ximenesia encelioides Cav.[Verbesinaencelioides (Cav.) Benth.&Hook.f.exA.Gray], including three different gatherings. A. Voucher MA-00476508! with material lacking original labels; B. Voucher MA-00476508−2! which bears the historical label handwritten by Cavanilles, among others. Reproduced with permission (Herbario del Real Jardín Botánico, CSIC, Madrid)." figureDoi="http://doi.org/10.5281/zenodo.13676039" httpUri="https://zenodo.org/record/13676039/files/figure.png" pageId="2" pageNumber="163">Fig. 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF70A08CC9044F00729EF8F4FEEABBD4" ID-DOI="http://doi.org/10.5281/zenodo.13676039" ID-Zenodo-Dep="13676039" httpUri="https://zenodo.org/record/13676039/files/figure.png" pageId="2" pageNumber="163" startId="2.[136,229,1794,1815]" targetBox="[163,1426,794,1761]" targetPageId="2" targetType="figure">
|
||||
<paragraph id="8BB0F004C9044F00729EF8F4FEEABBD4" blockId="2.[136,1452,1794,1923]" pageId="2" pageNumber="163">
|
||||
<emphasis id="B97B2C16C9044F00729EF8F4FF09BB40" bold="true" box="[136,252,1794,1815]" pageId="2" pageNumber="163">FIGURE 2.</emphasis>
|
||||
Part of the probable original material of
|
||||
<taxonomicName id="4C0F8B87C9044F007063F8F4FC8CBB40" authority="Cav." authorityName=", Cavanilles" authorityYear="1793" box="[629,889,1794,1815]" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="163" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9044F007063F8F4FCBCBB40" box="[629,841,1794,1815]" italics="true" pageId="2" pageNumber="163">Ximenesia encelioides</emphasis>
|
||||
Cav.
|
||||
</taxonomicName>
|
||||
[
|
||||
<taxonomicName id="4C0F8B87C9044F007192F8F4FA54BB40" authority="(Cav.) Benth. & Hook. f. ex A. Gray" authorityName="Benth. & Hook. f. ex A. Gray" authorityYear="1876" baseAuthorityName="Cav." box="[900,1441,1794,1815]" class="Magnoliopsida" family="Asteraceae" genus="Verbesina" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="163" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9044F007192F8F4FBA7BB40" box="[900,1106,1794,1815]" italics="true" pageId="2" pageNumber="163">Verbesina encelioides</emphasis>
|
||||
(Cav.) Benth. & Hook.f. ex A.Gray
|
||||
</taxonomicName>
|
||||
], including three different gatherings. A. Voucher MA-00476508! with material lacking original labels; B. Voucher MA-00476508
|
||||
<emphasis id="B97B2C16C9044F00779CF8DEFA6DBB6D" box="[1418,1432,1832,1850]" italics="true" pageId="2" pageNumber="163">−</emphasis>
|
||||
2! which bears the historical label handwritten by Cavanilles, among others. Reproduced with permission (Herbario del Real Jardín Botánico, CSIC, Madrid).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BB0F004C9044F0172ABF842FC6EBD10" blockId="2.[136,1453,1972,2067]" lastBlockId="3.[136,1453,160,1407]" lastPageId="3" lastPageNumber="164" pageId="2" pageNumber="163">
|
||||
As correctly described in
|
||||
<bibRefCitation id="EF9E8DF5C9044F0073DAF842FD73BB9C" author="Juan, A. & Hind, D. J. N. & Alonso, M. A." box="[460,646,1972,1995]" pageId="2" pageNumber="163" pagination="298 - 300" refId="ref2435" refString="Juan, A., Hind, D. J. N. & Alonso, M. A. (2015) Lectotypification of the name Ximenesia encelioides, basionym of Verbesina encelioides (Asteraceae: Heliantheae: Verbesininae). Phytotaxa 238 (3): 298 - 300. http: // dx. doi. org / 10.11646 / phytotaxa. 238.3.11" type="journal article" year="2015">
|
||||
Juan
|
||||
<emphasis id="B97B2C16C9044F007014F842FDC2BB9C" box="[514,567,1972,1995]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
|
||||
(2015)
|
||||
</bibRefCitation>
|
||||
, those two sheets at MA include material collected in the flowerbeds of Real Jardín Botánico de
|
||||
<collectingRegion id="49CB3EE6C9044F00734DF82EFE5FBBB8" box="[347,426,2008,2031]" country="Spain" name="Madrid" pageId="2" pageNumber="163">Madrid</collectingRegion>
|
||||
at three different times. On the one hand, MA-00476508 (
|
||||
<figureCitation id="1334EC81C9044F00763FF82EFB8ABBB8" box="[1065,1151,2008,2031]" captionStart="FIGURE 2" captionStartId="2.[136,229,1794,1815]" captionTargetBox="[163,1426,794,1761]" captionTargetId="figure-294@2.[151,1436,784,1768]" captionTargetPageId="2" captionText="FIGURE 2.Part of the probable original material of Ximenesia encelioides Cav.[Verbesinaencelioides (Cav.) Benth.&Hook.f.exA.Gray], including three different gatherings. A. Voucher MA-00476508! with material lacking original labels; B. Voucher MA-00476508−2! which bears the historical label handwritten by Cavanilles, among others. Reproduced with permission (Herbario del Real Jardín Botánico, CSIC, Madrid)." figureDoi="http://doi.org/10.5281/zenodo.13676039" httpUri="https://zenodo.org/record/13676039/files/figure.png" pageId="2" pageNumber="163">Fig. 2A</figureCitation>
|
||||
) includes six fragments but does not bear any original label, it showing only two modern typewritten labels referring to the Cavanillesian type locality. On the other hand, MA-00476508–2 (
|
||||
<figureCitation id="1334EC81C9054F017027FF56FD72BCE0" box="[561,647,160,183]" captionStart="FIGURE 2" captionStartId="2.[136,229,1794,1815]" captionTargetBox="[163,1426,794,1761]" captionTargetId="figure-294@2.[151,1436,784,1768]" captionTargetPageId="2" captionText="FIGURE 2.Part of the probable original material of Ximenesia encelioides Cav.[Verbesinaencelioides (Cav.) Benth.&Hook.f.exA.Gray], including three different gatherings. A. Voucher MA-00476508! with material lacking original labels; B. Voucher MA-00476508−2! which bears the historical label handwritten by Cavanilles, among others. Reproduced with permission (Herbario del Real Jardín Botánico, CSIC, Madrid)." figureDoi="http://doi.org/10.5281/zenodo.13676039" httpUri="https://zenodo.org/record/13676039/files/figure.png" pageId="3" pageNumber="164">Fig. 2B</figureCitation>
|
||||
) shows four fragments and several labels with data useful for
|
||||
<typeStatus id="54B44EA6C9054F01775CFF56FA59BCE0" box="[1354,1452,160,183]" pageId="3" pageNumber="164" type="lectotype">lectotype</typeStatus>
|
||||
selection. Two of them are identical to the labels on the voucher MA-00476509 and were handwritten in 1801 and 1841 by José Demetrio Rodríguez, a disciple of Cavanilles. Another label is in Cavanilles’s hand and matches the information in the protologue. Unfortunately, none of those three labels is clearly connected to the corresponding plant fragments, a fact that hinders the unequivocal identification of the true Cavanillesian gathering.
|
||||
</paragraph>
|
||||
<paragraph id="8BB0F004C9054F0172ABFEA2FE32BD80" blockId="3.[136,1453,160,1407]" pageId="3" pageNumber="164">
|
||||
Joining several successive gatherings of the same
|
||||
<typeStatus id="54B44EA6C9054F0170C6FEA2FD0BBD3C" box="[720,766,340,363]" pageId="3" pageNumber="164">type</typeStatus>
|
||||
plant in a single voucher appears to be a common practice in the 19
|
||||
<superScript id="7C7A5D4CC9054F0172B5FE80FF5ABDD3" attach="left" box="[163,175,374,388]" fontSize="6" pageId="3" pageNumber="164">th</superScript>
|
||||
century in MA, perhaps due to the scarcity of paper and/or space in the herbarium. This fact sometimes makes difficult identification of original material of plants collected in the Spanish expeditions around the New World (see, for instance,
|
||||
<bibRefCitation id="EF9E8DF5C9054F01729EFE36FE49BD80" author="Crespo, M. B. & Pena-Martin, C." box="[136,444,448,471]" pageId="3" pageNumber="164" pagination="215 - 221" refId="ref2216" refString="Crespo, M. B. & Pena-Martin, C. (2014) Typification of Pascalia glauca (Heliantheae, Asteraceae), with historical notes on that name. Phytotaxa 186 (4): 215 - 221. http: // dx. doi. org / 10.11646 / phytotaxa. 186.4.5" type="journal article" year="2014">Crespo & Pena-Martín 2014</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BB0F004C9054F0172ABFE12FBD7BEDC" blockId="3.[136,1453,160,1407]" pageId="3" pageNumber="164">
|
||||
Although today it is not possible to trace the real source and collection date of the fragments in that “
|
||||
<typeStatus id="54B44EA6C9054F0176E1FE12FAA8BDAC" box="[1271,1373,484,507]" pageId="3" pageNumber="164" type="lectotype">lectotype</typeStatus>
|
||||
” sheet, most of them (as well as those in MA-00476509) appear to belong to descendants of the individuals grown in
|
||||
<collectingRegion id="49CB3EE6C9054F01774BFDFEFA59BE48" box="[1373,1452,520,543]" country="Spain" name="Madrid" pageId="3" pageNumber="164">Madrid</collectingRegion>
|
||||
and described by Cavanilles (or even some of these were “clonotypes”), and they are relevant to the authentic material. Furthermore, all specimens in those vouchers are in good state or conservation and are undoubtedly identifiable with the current
|
||||
<taxonomicName id="4C0F8B87C9054F0172CCFD82FE37BEDC" authorityName="Benth. & Hook. f. ex A. Gray" authorityYear="1876" baseAuthorityName="Cav." box="[218,450,628,651]" class="Magnoliopsida" family="Asteraceae" genus="Verbesina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="164" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9054F0172CCFD82FE37BEDC" box="[218,450,628,651]" italics="true" pageId="3" pageNumber="164">Verbesina encelioides</emphasis>
|
||||
</taxonomicName>
|
||||
, and hence the application of the name is not threatened.
|
||||
</paragraph>
|
||||
<paragraph id="8BB0F004C9054F0172ABFD6EFDE9BF34" blockId="3.[136,1453,160,1407]" pageId="3" pageNumber="164">
|
||||
A detailed comparison of the illustration in the protologue (
|
||||
<figureCitation id="1334EC81C9054F017153FD6EFC71BEF8" box="[837,900,664,687]" captionStart="FIGURE 1" captionStartId="1.[136,229,1925,1946]" captionTargetBox="[151,1436,207,1881]" captionTargetId="figure-14@1.[151,1436,190,1900]" captionTargetPageId="1" captionText="FIGURE 1. Ximenesia encelioides Cav. [Verbesina encelioides (Cav.) Benth.& Hook.f. exA.Gray]. Illustration in the protologue (as “Tab. 178”), designated here as lectotype of the name. Reproduced with permission (Biblioteca del Real Jardín Botánico, CSIC, Madrid)." figureDoi="http://doi.org/10.5281/zenodo.13676035" httpUri="https://zenodo.org/record/13676035/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
|
||||
) with every single fragment in MA-00476508 and MA-00476508–2 does not bring a perfect match, which will facilitate lectotypification. However, part of the fragment in the lower central side of MA-00476508 or the one on the left side of that sheet are partially very similar to Cavanilles’s drawing, the former also bearing fruiting heads that could have served for preparing “Tab. 178”. These facts would probably suggest that Cavanilles’s artwork either was drawn from fragments that were discarded later or would represent only part or parts of the fragments in the original material.
|
||||
</paragraph>
|
||||
<paragraph id="8BB0F004C9054F0172ABFC86FBD7B808" blockId="3.[136,1453,160,1407]" pageId="3" pageNumber="164">
|
||||
As the vouchers MA-00476508 and MA-00476508–2 include fragments from three different gatherings, designation of any of them as
|
||||
<typeStatus id="54B44EA6C9054F017359FC62FE44BFFC" box="[335,433,916,939]" pageId="3" pageNumber="164" type="lectotype">lectotype</typeStatus>
|
||||
would be speculative and even daring, since no connection with the corresponding labels and collection dates can be unmistakably established. It is worth mentioning here that both biggest fragments in the central and left lower sides of MA-00476508–2, which were suggested as good candidates for
|
||||
<typeStatus id="54B44EA6C9054F0171E1FC2AFBACBFA4" box="[1015,1113,988,1011]" pageId="3" pageNumber="164" type="lectotype">lectotype</typeStatus>
|
||||
by Juan
|
||||
<emphasis id="B97B2C16C9054F0176A1FC2AFB18BFA4" box="[1207,1261,988,1011]" italics="true" pageId="3" pageNumber="164">et al.</emphasis>
|
||||
(2005), look very much alike the biggest fragments on MA-00476509. Those latter were collected either in 1801 or 1841, and therefore they most probably belong to gatherings made after publication of the protologue. Any of the remaining two fragments in MA-00476508–2 can neither be undoubtedly identified as belonging to the 1792 gathering.
|
||||
</paragraph>
|
||||
<paragraph id="8BB0F004C9054F0172ABFB9AFD97B89C" blockId="3.[136,1453,160,1407]" pageId="3" pageNumber="164">
|
||||
Accordingly, it is safer in this case to designate the illustration in “Tab. 178” as
|
||||
<typeStatus id="54B44EA6C9054F017654FB9AFB51B8D4" box="[1090,1188,1132,1155]" pageId="3" pageNumber="164" type="lectotype">lectotype</typeStatus>
|
||||
of the name
|
||||
<taxonomicName id="4C0F8B87C9054F01772AFB9AFEF5B8F0" authorityName=", Cavanilles" authorityYear="1793" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="164" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9054F01772AFB9AFEF5B8F0" italics="true" pageId="3" pageNumber="164">Ximenesia encelioides</emphasis>
|
||||
</taxonomicName>
|
||||
. This avoids involuntary selection of a fragment not corresponding with Cavanilles’s original gathering, and hence not belonging to the original material.
|
||||
</paragraph>
|
||||
<paragraph id="8BB0F004C9054F0172ABFB2EFAF5B928" blockId="3.[136,1453,160,1407]" pageId="3" pageNumber="164">
|
||||
It should be noted that the drawing of
|
||||
<taxonomicName id="4C0F8B87C9054F01705BFB2EFD17B8B8" authorityName=", Cavanilles" authorityYear="1793" box="[589,738,1240,1263]" class="Magnoliopsida" family="Asteraceae" genus="Ximenesia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="164" phylum="Tracheophyta" rank="species" species="encelioides">
|
||||
<emphasis id="B97B2C16C9054F01705BFB2EFD17B8B8" box="[589,738,1240,1263]" italics="true" pageId="3" pageNumber="164">X. encelioides</emphasis>
|
||||
</taxonomicName>
|
||||
here designated as
|
||||
<typeStatus id="54B44EA6C9054F0171B9FB2EFBE4B8B8" box="[943,1041,1240,1263]" pageId="3" pageNumber="164" type="lectotype">lectotype</typeStatus>
|
||||
is incomplete (it depicts the upper part of a stem, lacking the typically opposite lower leaves) and some of the diagnostic features it shows are poor (e.g. lack of leaf indumentum details, unclear structure of capitula, or undetailed features of flowers and fruits). However, the general habit and the structures depicted allow an easy and unambiguous identification of the species as well as a precise application of the name, and therefore an eventual
|
||||
<typeStatus id="54B44EA6C9054F01701DFA9EFDAFB928" box="[523,602,1384,1407]" pageId="3" pageNumber="164" type="epitype">epitype</typeStatus>
|
||||
designation is not necessary as required by Art. 9.8 of the ICN.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue