|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; fFeuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+subalpin (
+ |
+||
|
+ |
+
+sehr
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+452501 | +
| = | +
+ |
+
+ |
+733 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+nicht
+ |
+|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+|
|---|---|
|
+ |
+Senegal; Yemen | +
|
+ |
+Uganda; Yemen | +
|
+ |
+Ghana; Yemen | +
|
+ |
+Ghana; Oman, Yemen | +
|
+ |
+|
|
+ |
+Lebanon | +
|
+ |
+Oman | +
|
+ |
+Yemen | +
|
+ |
+United Arab Emirates, Yemen | +
|
+ |
+Saudi Arabia | +
|
+ |
+Oman, United Arab Emirates | +
|
+ |
+Yemen | +
|
+ |
+Oman | +
|
+ |
+Israel | +
|
+ |
+Egypt | +
|
+ |
+Egypt | +
|
+ |
+Egypt | +
|
+ |
+|
|
+ |
+Algeria, Afghanistan, Europe, South Africa; Egypt, Israel, Lebanon, United Arab Emirates, Yemen | +
|
+ |
+Afghanistan, Azerbaijan, Europe, Georgia, Kazakhstan, Turkmenistan, Tajikistan, Uzbekistan; Israel, Lebanon | +
|
+ |
+Azerbaijan, Central Russia, Europe, Kyrgyzstan; Israel | +
|
+ |
+Azerbaijan, Ukraine; Turkey | +
|
+ |
+Tajikistan, Turkmenistan; Oman, United Arab Emirates | +
|
+ |
+Azerbaijan, Europe, Kazakhstan, Mongolia, S Russia, Tajikistan; Iran, Iraq, Turkey | +
|
+ |
+Armenia, Georgia, Kazakhstan, Europe; Turkey, Israel | +
|
+ |
+Armenia, Georgia, Europe, Tunisia; Turkey | +
|
+ |
+China, Europe, Japan, Kazakhstan, Kyrgyzstan, Russia; Israel | +
|
+ |
+|
|
+ |
+North America, Azerbaijan, N China, Europe, Georgia, Japan, N and NW Russia, Tajikistan; Turkey | +
| - | +P1 | +P2 | +P3 | +P4 | +P5 | +
| Exopod | +0,1,022 | +0,1,122 | +0,1,122 | +0,1,022 | +3 | +
| Endopod | +0,210 | +0,111 | +0,111 | +020 | +5 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|
| Habit | +Small trees, branches drooping, young parts densely hairy | +Small trees, branches spreading, young parts pubescent | +Shrubs, branches spreading, young parts glabrous | +Shrubs, branches spreading, young parts strigose | +
| Leaf shape and size | +Oblong or obovate, caudate acuminate at apex, 6.0–12.0 × 2.5– 4.0 cm | +Oblong-elliptic or oblong- lanceolate, obtuse or acute at apex, 4.0–9.0 ×1.5–3.0 cm | +Ovate or ovate- lanceolate, acute to shortly acuminate at apex, 5.0–7.0 × 2.0–2.7 cm | +Elliptic-oblong to oblong-lanceolate, acute or slightly acuminate at apex, 4.5–9.0 × 2.5–2.8 cm | +
| Position of intra- marginal looping nerves | +Sub-marginal | +Marginal | +Marginal | +Marginal | +
| Petiole | +Glabrous, 2–3 mm long | +Pubescent, 1–2 mm long | +Pubescent, 1–2 mm long | +Tomentose, 1-2 mm long | +
| Flowers | +Pseudo-terminal, greenish yellow | +Axillary, green | +Leaf-opposed, white | +Axillary, greenish pink | +
| Pedicel | +2–4 mm long, glabrous, greenish | +3–6 mm long, glabrous, greenish | +0.7–1.5 cm long, glabrous, greenish | +2–3 mm long, pubescent, reddish | +
| Sepals | +Glabrous, sub-equal | +Pubescent outside, equal | +Pubescent outside, equal | +Glabrous, sub-equal | +
| Outer petals | +Ovate, hairy on margins | +Ovate, glabrous | +Linear-lanceolate, ciliate | +Lanceolate, glabrous | +
| Inner petals | +2.0–2.4 × 0.8–1.3 cm | +1.2–1.6 × 0.6–0.7 cm | +1.5–2.0 × 0.6–1.0 cm | +1.0–1.7 × 0.3–0.5 cm | +
| Stamens | +With included connectives | +With connectives rounded apiculate at top | +With connectives slightly apiculate at top | +With connectives rounded apiculate at top | +
| Carpels | +Linear-oblong, glabrous | +Linear-oblong, densely pilose | +Ovate, glabrous | +Oblong, grey tomentose | +
| Monocarps | +Obovoid, apex projecting, more or less obturbinate, glabrous, deep pink | +Ovoid or obovoid, apex acute or obtuse, silky pubescent, red | +Sub-globose, apex obtuse, glabrous, red | +Obliquely ellipsoid or oblong, apex apiculate, pubescent, red | +
| Leg measurements: | +||||
|---|---|---|---|---|
| Femur Patella | +Tibia | +Metatarsus | +Tarsus | +Total | +
| I 3.9 1.55 | +4.3 | +4.2 | +2.2 | +16.15 | +
| II 3.9 1.5 | +3.8 | +3.85 | +2.2 | +15.25 | +
| III 3.65 1.45 | +3.35 | +4.05 | +2.05 | +14,55 | +
| IV 4.9 1.45 | +4.55 | +5.55 | +2.45 | +18.90 | +
| Leg measurement: | +||||
|---|---|---|---|---|
| Femur Patella | +Tibia | +Metatarsus | +Tarsus | +Total length | +
| I 2.55 1.2 | +2.3 | +2.1 | +1.35 | +9.50 | +
| II 2.25 1.1 | +1.95 | +1.9 | +1.22 | +8.42 | +
| III 2.1 1.1 | +1.62 | +2 | +1.1 | +7.92 | +
| IV 2.85 1.2 | +2.67 | +3.05 | +1.4 | +11.17 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+sehr warm-kollin (nur an
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
|
+ |
+1.7 | +1.8 | +|||
|
+ |
+14.7 | +1.1 | +|||
|
+ |
+17.3 | +7.9 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+unter-alpin, supra-subalpin und ober-subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
|
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|
| Leaves | +elliptic | +obovate or elliptic-ovate | +elliptic-obovate or oblanceolate | +
| Lateral vein nerves | +6–8 pairs | +10–15 pairs | +5–6 pairs | +
| Petioles | +2–5 mm long | +6–10 mm long | +3–8 mm long | +
| Flower at anthesis (diameter) | +0.7–1 cm across | +ca.2 cm across | +ca.1 cm across | +
| Bracteoles | +oblong | +lanceolate | +linear | +
| Bracteoles length | +1.6–1.9 mm long | +ca. 2 mm long | +ca.3 mm long | +
| Calyx tube length | +0.9–1.5 mm long | +2–3 mm long | +4–6 mm long | +
| Calyx lobe width | +1.6–2.5 mm across | +4–5 mm across | +ca. 5 mm across | +
| Calyx lobes indumentum | +Rusty brown long hairs on the abaxial side and floccose white pubescent on adaxial side. | +tomentose without, glabrous within | +Fulvous tomentose, glabrous within | +
| Petals | +broadly elliptic | +orbicular | +orbicular | +
| Seeds | +1-2 | +1-2 | +many | +
| Species | +CR distal armature | +Atl segs (setae) | +Ant terminal armature | +Labrum proc. | +Setae on Mxl lobes Inner Outer | +|
|---|---|---|---|---|---|---|
|
+ |
+CR absent | +1 (5) | +Claw | +2 | +2 | +2 | +
|
+ |
+Process | +1 (ca. 7) | +2 setae | +6 | +2 | +3 | +
|
+ |
+0 | +2 (6) | +2 proc. | +5 | +2 | +3 | +
|
+ |
+Spine | +2 (4) | +2 proc. | +0 | +2 | +2 | +
|
+ |
+CR absent | +1 (5) | +Claw | +6 | +1 | +2 | +
|
+ |
+Process | +2 (7) | +2 proc. | +6 | +2 | +3 | +
|
+ |
+- | +1 (5) | +Claw | +2 | +2 | +0 | +
|
+ |
+Spine | +1 (6) | +Claw | +0 | +2 | +2 | +
|
+ |
+Seta | +1 (?) | +Claw | +? | +2 | +2 | +
|
+ |
+Seta | +1 (7) | +Claw | +6 | +2 | +3 | +
|
+ |
+Spine | +2 (ca. 7) | +2 spines | +8 | +2 | +3 | +
|
+ |
+Spine | +2 (10) | +2 spines | +0 | +2 | +3 | +
|
+ |
+Seta | +1 (8) | +Spine | +1 | +2 | +2 | +
|
+ |
+0 | +1 (7) | +0 | +4 | +2 | +1 | +
|
+ |
+0 | +2 (8) | +Claw | +3 | +2 | +2 | +
|
+ |
+Spine | +2 (7) | +Claw | +6 | +5 | +3 | +
|
+ |
+Spine | +1 (10) | +2 spines | +6 | +2 | +3 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1: | +0-0 | +1-0 | +I-0; I, I, 3 | +0-0; 0-1; 1, 1, 3 | +
| Leg 2: | +0-0 | +1-0 | +I-1; I-1; II, I, 4 | +0-1; 0-1; 1, 2, 3 | +
| Leg 3: | +0-0 | +1-0 | +I-1; I-1; II, I, 5 | +0-1; 0-1; 1, 2, 3 | +
| Leg 4: | +0-0 | +1-0 | +I-1; I-1; II, I, 4 | +0-1; 0-2; 1, 2, 2 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1: | +0-0 | +1-I | +I-0; I, I, 3 | +0-0; 0-0; 1, 2, 2 | +
| Leg 2: | +0-0 | +1-0 | +I-1; I-1; II, I, 4 | +0-1; 0-2; 1, 2, 3 | +
| Leg 3: | +0-0 | +1-0 | +I-1; I-1; II, I, 5 | +0-1; 0-2; 1, 2, 3 | +
| Leg 4: | +0-0 | +1-0 | +I-1; I-1; II, I, 4 | +0-1; 0-2; 1, 2, 2 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1: | +0-0 | +1-I | +I-I (1-0); I, I, 3 | +0-0; 0-0 (0-1); 1, 2, 2 | +
| Leg 2: | +0-0 | +1-0 | +I-1; I-1; II, I, 4 | +0-1; 0-2 (0-1); 1, 2, 3 | +
| Leg 3: | +0-0 | +1-0 | +I-1; I-1; II, I, 5 | +0-1; 0-2 (0-1); 1, 2, 3 | +
| Leg 4: | +0-0 | +1-0 | +I-1; I-1; II, I, 4 | +0-1; 0-2; 1, 2, 2 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-0 | +VI, 1 | +8 | +
| Leg 2 | +0-0 | +1-0 | +VI, 1 | +9 | +
| Legs 3 & 4 | +0-0 | +0-0 | +VI, 1 | +7 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +V | +1; 6 | +5 | +1; 6 | +
| Leg 2 | +IV | +1; III+3 | +4 | +1; 6 | +
| Leg 3 | +III | +1; I+3 | +3 | +1; 4 | +
| Leg 4 | +IV | +1; III+1 | +3 | +1; 4 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-0 | +VI | +II | +
| Leg 2 | +0-0 | +1-0 | +V | +II | +
| Legs 3 & 4 | +0-0 | +1-0 | +IV | +(endopod lacking) | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-0 | +V, 1 | +6 | +
| Leg 2 | +0-0 | +1-0 | +IV, 1 | +II+ 3 | +
| Leg 3 | +0-0 | +1-0 | +IV, 1 | +II+ 1 | +
| Leg 4 | +0-0 | +1-0 | +IV, 1 | +I+ 1 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +0-0 | +VI, 1 | +1, 2, 5 | +
| Leg 2 | +0-0 | +1-0 | +VI, 1 | +1, 2, 5 | +
| Legs 3 & 4 | +0-0 | +1-0 | +VI, 1 | +1, 2, 4 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Legs 1 & 2 | +0-0 | +1-0 | +VI, 1 | +7 | +
| Legs 3 & 4 | +0-0 | +1-0 | +VI, 1 | +6 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-0 | +VI | +7 | +
| Leg 2 | +0-0 | +1-0 | +VI | +6 | +
| Leg 3 | +0-0 | +1-0 | +VI | +4 | +
| Leg 4 | +0-0 | +0-0 | +VI | +3 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Legs 1 & 2 | +0-0 | +1-0 | +VII | +8 | +
| Legs 3 & 4 | +0-0 | +1-0 | +VII | +7 | +
| Protopod | +Exopod | +Endopod | +|
|---|---|---|---|
| Leg 1 | +1-0 | +V | +8 | +
| Leg 2 | +1-0 | +V | +3 | +
| Legs 3 & 4 | +1-0 | +V | +2 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-0 | +I-0; I-1; III, I, 4 | +0-1; 2 | +
| Leg 2 | +0-0 | +1-0 | +I-0; I-1; III, I, 5 | +0-1; 0-2; I, 3 | +
| Leg 3 | +0-1 | +1-0 | +I-0; I-1; III, I, 5 | +0-1; 0-2; I, 2 | +
| Leg 4 | +0-1 | +1-0 | +I-0; I-1; III, I, 5 | +0-1; I, 4 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-0 | +I-0; I-0; III, I, 4 | +II, 6 | +
| Leg 2 | +0-0 | +1-0 | +I-0; I-1; II, I, 5 | +0-1; 0-2; 0, II, 3 | +
| Leg 3 | +0-1 | +1-0 | +I-0; I-1; II, I, 5 | +0-1; 0-2; 0, II, 2 | +
| Leg 4 | +0-1 | +1-0 | +I-0; I-1; II, I, 5 | +0-1; 0, II, 3 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-0 | +VI | +8 | +
| Leg 2 | +0-0 | +1-0 | +VI | +3 | +
| Legs 3 & 4 | +0-0 | +1-0 | +VI | +2 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +8 | +6 | +8 | +
| Leg 2 | +VI | +1; 7 | +5 | +1; 7 | +
| Leg 3 | +VI | +1; II+3 | +5 | +1; I+4 | +
| Leg 4 | +V | +1; 5 | +4 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VI+1 | +1; 7 | +7 | +1; 7 | +
| Leg 2 | +VI | +1; 7 | +7 | +1; 7 | +
| Leg 3 | +VI | +1; 5 | +6 | +1; 5 | +
| Leg 4 | +VII | +1; 5 | +6 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +1; 7 | +6 | +1; 7 | +
| Leg 2 | +VI | +1; III+4 | +5 | +1; 7 | +
| Leg 3 | +VI | +(missing) | +5 | +1; 5 | +
| Leg 4 | +V | +1; 5 | +5 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +1; 7 | +6 | +1; 7 | +
| Leg 2 | +VI | +1; III+4 | +5 | +1; 6 | +
| Leg 3 Leg 4 | +VI V | +1; II+3 1; II+3 | +5 4 | +1; 5 1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +1; 7 | +6 | +1; 7 | +
| Leg 2 | +VI | +1; III+4 | +5 | +1; 7 | +
| Leg 3 | +VI | +1; II+3 | +5 | +1; 5 | +
| Leg 4 | +VI | +1; II+3 | +4 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +1; 7 | +7 | +1; 7 | +
| Leg 2 | +VI | +1; I+5 | +6 | +1; 6 | +
| Leg 3 | +VI | +1; I+4 | +6 | +1; 5 | +
| Leg 4 | +V | +1; I+4 | +5 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +V+1 | +1; 7 | +5 | +1; 7 | +
| Leg 2 | +V+1 | +1; 7 | +5 | +1; 7 | +
| Leg 3 | +VI | +1; 5 | +5 | +1; 5 | +
| Leg 4 | +IV | +1; 5 | +4 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +1; 6 | +7 | +1; 7 | +
| Leg 2 | +VI | +1; I+5 | +5 | +1; 6 | +
| Leg 3 | +VI | +1; I+4 | +5 | +1; 5 | +
| Leg 4 | +IV | +1; I+4 | +4 | +1; 5 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; III, I, III | +0-0; I, II, I | +
| Leg 2 | +0-0 | +1-0 | +I-0; III, I, III+1 | +0-0; I, II, II | +
| Legs 3 & 4 | +0-1 | +1-0 | +I-0; III, I, IV | +I, II, II | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; III, I, III | +0-0; I, II, I | +
| Leg 2 | +0-0 | +1-0 | +I-0; III, I, III | +0-0; I, II, II | +
| Leg 3 | +0-1 | +1-0 | +1-0; III, I, III | +0-0; I, II, II | +
| Leg 4 | +0-1 | +1-0 | +1-0; III, I, II | +0-0; I, II, II | +
| Armature of legs 1-4 | +|||||
|---|---|---|---|---|---|
| Species | +Right Exp | +Right Enp | +Left Exp | +Left Enp | +Source | +
|
+ |
+7-8-9-9 | +8-8-7-6 | +7-8-9-9 | +8-8-7-6 | +
+ |
+
|
+ |
+7-6-6-6 | +8-8-6-6 | +7-6-6-6 | +8-9-7-6 | +
+ |
+
|
+ |
+7-6-6-6 | +8-8-7-6 | +6-5-5-6 | +7-8-7-6 | +
+ |
+
|
+ |
+6-6-?-5 | +8-8-6-6 | +6-6-?-5 | +8-8-6-7 | +
+ |
+
|
+ |
+7-6-6-6 | +8-7-8-6 | +6-5-5-5 | +7-9-7-6 | +
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+Present study | +
|
+ |
+7-6-6-6 | +8-9-6-6 | +7-6-6-6 | +8-9-7-6 | +Present study | +
|
+ |
+7-6-6-6 | +8-9-7-6 | +7-5-5-5 | +8-9-7-6 | +Present study | +
|
+ |
+6-5-6-6 | +8-9-6-5 | +6-5-5-6 | +7-8-7-6 | +Present study | +
|
+ |
+6-6-6-6 | +8-9-7-6 | +5-5-5-5 | +8-9-7-6 | +Present study | +
|
+ |
+7-6-6-6* | +8-8-6-6 | +7-6-6-6 | +8-8-6-6 | +
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|
+ |
+6-6-6-5 | +8-7-6-6 | +5-5-5-4 | +8-8-6-6 | +
+ |
+
|
+ |
+6-5-5-5 | +7-7-6-6 | +6-4-4-5 | +7-7-6-6 | +
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
| Armature of legs 1-4 | +|||||
|---|---|---|---|---|---|
| Species | +Right Exp | +Right Enp | +Left Exp | +Left Enp | +Source | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+Present study | +
|
+ |
+7-6-6-5 | +8-8-6-6 | +6-5-5-4 | +8-7-6-6 | +Present study | +
|
+ |
+7-6-6-5 | +8-8-6-6 | +6-5-5-5 | +8-8-6-6 | +
+ |
+
|
+ |
+7-6-6-5 | +8-8-?-6 | +6-5-5-5 | +8-8-6-6 | +Present study | +
|
+ |
+7-6-6-5 | +8-7-6-6 | +7-6-6-5 | +8-7-6-6 | +Present study | +
|
+ |
+7-6-6-6 | +8-8-6-6 | +6-5-5-4 | +8-8-6-6 | +Present study | +
|
+ |
+7-6-6-4 | +7-7-6-6 | +7-5-5-4 | +8-7-6-6 | +Present study | +
|
+ |
+6-5-5-4 | +8-7-5-5 | +5-4-4-3 | +8-7-5-6 | +Ooishi & Illg (1989) | +
|
+ |
+6-5-5-4 | +8-7-5-5 | +5-4-4-3 | +9-7-5-5 | +
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|
+ |
+6-6-6-4 | +8-8-6-6 | +5-5-5-4 | +8-8-6-6 | +Present study | +
|
+ |
+5-4-3-4 | +7-7-5-5 | +5-4-3-3 | +7-7-5-5 | +Present study | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +I; VI | +1; 7 | +1; 5 | +1; 7 | +
| Leg 2 | +I; V | +1; 8 | +1; 5 | +1; 8 | +
| Leg 3 | +I; V | +1; 6 | +1; 5 | +1; 6 | +
| Leg 4 | +I; V | +1; 5 | +1; 5 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +1; 7 | +7 | +1; 7 | +
| Leg 2 | +VI | +1; 8 | +5 | +1; 8 | +
| Leg 3 | +VI | +I; 6 | +5 | +1; 6 | +
| Leg 4 | +VI | +I; 5 | +5 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VII | +1; 7 | +7 | +1; 7 | +
| Leg 2 | +I; V | +1; 8 | +1; 5 | +1; 8 | +
| Leg 3 | +I; V | +1; 5 | +1; 5 | +1; 6 | +
| Leg 4 | +I; V | +1; 5 | +1; 5 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VI | +1; 7 | +5 | +1; 7 | +
| Leg 2 | +VI | +1; 8 | +5 | +1; 8 | +
| Leg 3 | +I; V | +1; 6 | +1; 4 | +1; 6 | +
| Leg 4 | +I; V | +1; 5 | +1; 4 | +1; 5 | +
| Right exopod | +Right endopod | +Left exopod | +Left endopod | +|
|---|---|---|---|---|
| Leg 1 | +VI | +1; 7 | +IV+2 | +1; 6 | +
| Leg 2 | +V | +1; I+7 | +III+2 | +1; 7 | +
| Leg 3 | +I; V | +1; 5 | +1; 4 | +1; I+5 | +
| Leg 4 | +I; V | +1; 4 | +1; 5 | +1; 5 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0; | +1-I | +
+I-0;
+ |
+
+0-0; 0,
+ |
+
| Leg 2 | +0-0 | +1-0 | +
+I-0;
+ |
+
+0-0; I,
+ |
+
| Leg 3 | +0-0 | +1-0 | +
+I-0;
+ |
+
+0-1; I,
+ |
+
| Leg 4 | +0-0 | +1-0 | +
+I-0;
+ |
+
+0-1; 0,
+ |
+
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; III, I, III | +0-0; I II, 1 | +
| Leg 2 | +0-0 | +1-0 | +I-0; III, I, III | +0-0; I, III, 3 | +
| Leg 3 | +0-0 | +1-0 | +I-0; III, I, III | +0-1; I, III, 3 | +
| Leg 4 | +0-0 | +1-0 | +I-0; III, I, II | +0-1; 0, II, 2 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +
+I-0;
+ |
+
+0-0; I,
+ |
+
| Leg 2 | +0-0 | +1-0 | +
+I-0;
+ |
+
+0-0; 0,
+ |
+
| Leg 3 | +0-0 | +1-0 | +
+I-0;
+ |
+
+0-1; 0,
+ |
+
| Leg 4 | +0-0 | +1-0 | +
+I-0;
+ |
+
+0-1; 0,
+ |
+
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; III, II, 0 | +0-0; 0, II, 1 | +
| Leg 2 | +0-0 | +1-0 | +I-0; II, II, 0 | +0-0; 0, II, 3 | +
| Leg 3 | +0-0 | +1-0 | +I-0; II, II, 0 | +0-1; 0, III, 3 | +
| Leg 4 | +0-0 | +1-0 | +I-0; I, II, 0 | +0-1; 0, II, 2 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
| Legs 1-4 | +0-0 | +1-0 | +I-0; III, 0 | +0-0; 0 | +
|---|
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; III, 0 | +0-0; 0, II, 0 | +
| Legs 2-4 | +0-0 | +1-0 | +I-0; III, 0 | +0-0; III, II, 0 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; III, I, III | +0-0; I, II, I | +
| Legs 2 & 3 | +0-0 | +1-0 | +I-0; III, I, III+1 | +0-0; I, II, I+1 | +
| Leg 4 | +0-1 | +1-0 | +I-0; III, I, III+1 | +0-0; I, II, I+1 | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; II, I, II | +0-0; IV | +
| Legs 2 & 3 | +0-0 | +1-0 | +I-0; hook | +0-0; II, II, II | +
| Leg 4 | +0-0 | +1-0 | +I-0; hook | +0-1; II, II, I | +
| Coxa | +Basis | +Exopod | +Endopod | +|
|---|---|---|---|---|
| Leg 1 | +0-0 | +1-I | +I-0; III, I, III | +0-0; I, II, I | +
| Legs 2-4 | +0-0 | +1-0 | +I-0; III, I, III | +0-0; I, II, I | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+133750 | +
| Species | +Colony growth | +Branching type | +Terminal branches length (cm) | +Polyp distribution rows | +Pseudoanastomosis | +Calyx height elevation (mm) | +Calyx form | +Colony Color | +Outer coenenchymal and calyx dominant spindles | +Coenenchymal and calycular spindles maximum size (mm) | +Inner coenenchymal spindles | +Anthocodial sclerites | +Sclerites color | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+bu | +irr, lb | +2.8 | +c, im | +no | +1.9 | +el | +ro | +ls | +0.5 | +slr, ws | +lb, wr | +am, lo, ro, py | +
|
+ |
+bu | +irr, lb | +6-3 | +c | +no | +2.8-3 | +sl | +rb | +uss | +2.4 | +ws | +r, bs | +o, lb | +
|
+ |
+bu | +irr | +1.5-4 | +c | +no | +1-1.2 | +sl | +rb, w, bi | +uss | +2 | +ws | +ws, wr, bs | +w, rb, py | +
|
+ |
+fa | +ob | +8 | +s | +no | +0.6-1 | +sl | +lo | +uss | +4.1 | +ws | +r, ps | +am, lo | +
|
+ |
+fl | +irr, lb | +1-5 | +c, im | +no | +0.7-1.2 | +sl | +db/w | +ls | +1 | +ws | +lb, wr | +rb, am | +
|
+ |
+fl | +di | +4 | +c | +no | +2.6 | +t | +lb | +cl, cs | +1.3 | +tbs | +cl | +br, cl, py, y | +
|
+ |
+fl | +lb | +9.5 | +im | +no | +1 | +sl | +o | +ls, tbs | +1.2 | +ws | +lb | +cl, o, y, py | +
|
+ |
+fl | +lb | +9 | +im | +yes | +1 | +sl | +br-r | +ls | +1.7 | +ae/de-ws | +tbr-ae | +db, rb | +
|
+ |
+fl | +lb | +8.7 | +im | +no | +3 | +t | +g | +uss, tbs | +2.5 | +str, cs | +tbr, lr | +g, am, cl | +
|
+ |
+|||||||||
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+|||||||||
|
+ |
+|||||||||
|
+ |
+Body | +dorsally | +smooth | +(Fig. | +3A). | +Head | +rounded. | +Pereonite | +1 | +
|
+ |
+name | +refers | +to the | +Indian | +Ocean, | +the | +ocean | +where | +the | +
|---|---|---|---|---|---|---|---|---|---|
| + | specimens | +where collected. + +||||||||
|
+ |
+|||||||||
| Holotype male | +|||||||||
|
+ |
+
| Outer plate | +6 setae | +6 setae | +45 setae | +56 setae | +
|---|---|---|---|---|
| Distal projection on penultimate article of palp | +Present | +Present | +Absent | +Present | +
| Gnathopod 1 grasping spines | +1 | +1 | +2 | +1 | +
| Proximal projection of the propodus of gnathopod 2 male | +Triangular | +Triangular | +Rectangular | +Rectangular | +
| Pereopods 3 and 4 | +2articulate | +1articulate | +2articulate | +2articulate | +
| Pereopod 5 | + | +Normal | +Reduced | +Normal | +
| Male abdominal appendages | +1articulate | +2articulate | +1articulate | +1articulate | +
| Discussion | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
| 9 | +7,8 | +4 | +6,1 | +4,5 | +13,5 | +5,7 | +
| 8 | +6,2 | +3 | +4,6 | +3,5 | +12 | +4,8 | +
| Present | +Absent | +Absent | +Absent | +Absent | +Present | +Present | +
| 1 pair | +1 pair | +1 pair | +1 pair | +1 pair | +Absent | +1 pair | +
| 1011 | +14 | +67 | +67 | +68 | +11 | +11 | +
| 10 | +10 | +67 | +67 | +6 | +9 | +7 | +
| 2 | +2 | +2 | +2 | +2 | +2 | +2 | +
| Present | +Present | +Absent | +Present | +Absent | +Absent | +Absent | +
| 1x1, x=310 | +1x1, x=9 | +1 or 2 setae | +1x1, x=1 | +2 setae | +1x1, x=11 | +1x1, x=45 | +
| 5toothed | +5toothed | +5toothed | +5toothed | +5toothed | +5toothed | +5toothed | +
| 5toothed | +5toothed | +5toothed | +5toothed | +5toothed | +5toothed | +5toothed | +
| 5toothed | +4toothed | +5toothed | +5toothed | +5toothed | +5toothed | +5toothed | +
| Serrate | +Serrate | +Serrate | +Serrate4toothed | +Serrate | +Serrate | +Serrate | +
| Absent | +Present | +Absent | +Present | +Absent | +Absent | +Present | +
| 56 | +5 | +45 | +7 | +5 | +6 | +6 | +
| 4 | +4 | +34 | +4 | +4 | +4 | +5 | +
| 57 | +3 | +56 | +8 | +4 | +5 | +4 | +
| 45 | +4 | +45 | +6 | +4 | +5 | +5 | +
| no “tooth” | +1 “tooth” | +no “tooth” | +1 “tooth” | +1 “tooth” | +no “tooth” | +1 “tooth” | +
| 4 setae | +4 setae | +34 setae | +4 setae | +34 setae | +3 setae | +3 setae | +
| 57 setae | +45 setae | +12 setae | +6 setae | +7 setae | +45 setae | +5 setae | +
| Absent | +Absent | +Absent | +Absent | +Present | +Absent | +Present | +
| 2 | +2 | +1 | +1 | +1 | +2 | +1 | +
| Rectangular | +Rectangular | +Triangular | +Triangular | +Rectangular | +Rectangular | +Rectangular | +
| 2articulate | +2articulate | +2articulate | +1articulate | +2articulate | +2articulate | +1 or 2articulate | +
| Normal | +Normal | +Normal | + | +Normal | +Normal | +Normal | +
| 1articulate | +2articulate | +1articulate | +2articulate | +1articulate | +1articulate | +1articulate | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Characters | +
+ |
+
+*
+ |
+
+**
+ |
+
+***
+ |
+
|---|---|---|---|---|
| Petiole length; indumentum | +2.5-7 cm long; puberulous | +3-9 cm long; pubescent | +1-2 cm long; sparsely pilose | +1-2 cm long; adpressed hairy | +
| Lamina; length by width (cm) | +
+lanceolate, oblanceolate or broadly oblanceolate; 15-24
+ |
+
+ovate oblong to lanceolate, rarely ovate; 7.5-33
+ |
+
+narrowly elliptic to lanceolate-elliptic; 14-25
+ |
+
+oblanceolate, oblong-oblanceolate, or elliptic oblong; 11.2-17.5
+ |
+
| Lamina base; sinus depth (mm) | +cordate; 2-3 | +cordate; 5-10 | +narrowly auriculate; 5-7 | +narrowly, slightly cordate | +
| Pinnate lateral vein pairs | +6-7 | +4-6 | +9-10 | +7-8 | +
| Inflorescence | +cauline; peduncle 15.5-17 cm long, divided into 4-5 internodes of different lengths | +in axils of foliage leaves, rarely cauline; peduncle up to 2 cm long | +in axils of foliage leaves, peduncle 2-3 cm long | +in axils of foliage leaves, peduncle ca 10 cm long | +
| Bract indumentum | +pilose | +puberulous | +- | +pilose | +
| Perianth | +
+tube geniculately curved, utricle cylindric, ca 30
+ |
+
+tube geniculately curved, utricle ovoid tubular, 35-30
+ |
+
+tube geniculately curved, basal portion of tube 30-40
+ |
+
+tube geniculately curved, utricle ovoid, 8-10
+ |
+
|
+ |
+Peninsular Malaysia | +
+Sumatra, Java, †Peninsular Malaysia (
+ |
+
+China: Guangdong, Guangxi, Yunnan (
+ |
+
+North Eastern Thailand (
+ |
+
|
+ |
+(PCA) | +of shell | +ratios | +of | +
+ |
+
+and
+ |
+
|---|---|---|---|---|---|---|
|
+ |
+= angle | +of the last | +whorl, | +LW | += length width ratio, RD | += relative diameter, | +
| RSH = relative height of spire. | +
| 25 | +
+ |
+
+ |
+0 | +1 | +0 | +1 | +0 | +0 | +0 | +2.36 | +5.01 | +0.50 | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 26 | +
+ |
+
+ |
+0 | +1 | +0 | +1 | +0 | +0 | +0 | +2.18 | +5.80 | +0.64 | +
| 27 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +0 | +0 | +0 | +2.63 | +14.57 | +0.48 | +
| 28 | +
+ |
+
+ |
+1 | +1 | +0 | +1 | +0 | +0 | +0 | +1.59 | +5.81 | +0.34 | +
| 29 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +0 | +0 | +0 | +1.37 | +4.15 | +0.35 | +
| 30 | +
+ |
+
+ |
+0 | +0 | +0 | +1 | +0 | +0 | +0 | +1.72 | +6.73 | +0.55 | +
| 31 | +
+ |
+
+ |
+0 | +1 | +0 | +0 | +0 | +0 | +0 | +2.03 | +5.57 | +0.50 | +
| 32 | +
+ |
+
+ |
+0 | +? | +? | +? | +? | +? | +? | +? | +? | +? | +
| 33 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +0 | +0 | +0 | +6.67 | +14.84 | +0.41 | +
| 34 | +
+ |
+
+ex. gr.
+ |
+0 | +0 | +0 | +0 | +1 | +0 | +0 | +2.06 | +5.15 | +0.46 | +
| 35 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +0 | +0 | +2.37 | +6.21 | +0.62 | +
| 36 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +1 | +0 | +0 | +1.36 | +4.41 | +0.50 | +
| 37 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +1 | +0 | +0 | +1.93 | +5.93 | +0.35 | +
| 38 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +0 | +0 | +0 | +2.64 | +5.25 | +0.48 | +
| 39 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +0 | +0 | +0 | +2.37 | +8.52 | +0.37 | +
| 40 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +0 | +0 | +2.23 | +4.93 | +0.57 | +
| 41 | +
+ |
+
+ |
+? | +? | +? | +? | +? | +? | +? | +? | +? | +? | +
| 42 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +0 | +0 | +1.79 | +5.04 | +0.45 | +
| 43 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +0 | +0 | +1.98 | +4.27 | +0.51 | +
| 44 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +0 | +0 | +2.45 | +6.36 | +0.48 | +
| 45 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +0 | +0 | +2.97 | +12.88 | +0.38 | +
| 46 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +0 | +0 | +0 | +2.80 | +7.37 | +0.54 | +
| 47 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +1 | +1 | +2.71 | +7.94 | +0.67 | +
| 48 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +1 | +1 | +2.60 | +6.08 | +0.48 | +
| 49 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +1 | +1 | +2.48 | +7.23 | +0.57 | +
| 50 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +1 | +0 | +0 | +1.59 | +5.26 | +0.35 | +
| 51 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +1 | +0 | +0 | +1.22 | +3.63 | +0.29 | +
| 52 | +
+ |
+
+ |
+1 | +0 | +1 | +0 | +1 | +0 | +0 | +2.10 | +5.36 | +0.41 | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 53 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +1 | +0 | +0 | +1.92 | +6.08 | +0.29 | +
| 54 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +1 | +0 | +0 | +1.41 | +4.02 | +0.36 | +
| 55 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +1 | +0 | +0 | +1.78 | +5.80 | +0.53 | +
| 56 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +1 | +0 | +0 | +1.72 | +3.63 | +0.45 | +
| 57 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +1 | +0 | +0 | +2.91 | +4.86 | +0.57 | +
| 58 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +1 | +0 | +0 | +? | +? | +? | +
| 59 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +1 | +0 | +0 | +1.27 | +5.04 | +0.27 | +
| 60 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +1 | +1 | +2.92 | +7.87 | +0.52 | +
| 61 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +1 | +0 | +3.82 | +8.89 | +0.54 | +
| 62 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +1 | +0 | +3.65 | +9.50 | +0.51 | +
| 63 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +1 | +0 | +1.70 | +4.42 | +0.29 | +
| 64 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +1 | +0 | +1.23 | +6.37 | +0.22 | +
| 65 | +
+ |
+
+ |
+1 | +1 | +1 | +0 | +1 | +1 | +0 | +1.13 | +5.16 | +0.20 | +
| 66 | +
+ |
+
+ |
+0 | +0 | +0 | +0 | +0 | +1 | +0 | +0.93 | +4.92 | +0.21 | +
| 67 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +1 | +0 | +0.88 | +5.05 | +0.20 | +
| 68 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +1 | +0 | +1.81 | +3.53 | +0.37 | +
| 69 | +
+ |
+
+ |
+0 | +1 | +1 | +0 | +0 | +1 | +0 | +0.87 | +5.02 | +0.11 | +
| 70 | +
+ |
+
+ |
+0 | +0 | +1 | +0 | +0 | +1 | +0 | +1.54 | +6.63 | +0.20 | +
| 71 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +0 | +1 | +0 | +1.80 | +5.55 | +0.27 | +
| 72 | +
+ |
+
+ |
+1 | +1 | +0 | +1 | +1 | +1 | +0 | +2.42 | +4.51 | +0.25 | +
| 73 | +
+ |
+
+ |
+1 | +1 | +0 | +0 | +1 | +1 | +0 | +? | +? | +? | +
| 74 | +
+ |
+
+ |
+1 | +1 | +0 | +1 | +1 | +0 | +0 | +? | +? | +? | +
| SK | +
| Saskatchewan: 49.9037°, -109.5909° | +
|
+ |
+
| Collector | +Locality | +Number | +
|---|---|---|
| Arnoux | +Wallis (Oua-Horn) | +1 | +
| Le Guillou | +Oceania | +1 | +
| Quoy & Gaimard | +Tongatabou | +2 (ad. and subad.) | +
| Quoy & Gaimard | +Tongatabou | +1 | +
| Arnoux | +Tongatabou | +1 | +
| Unknown | +Oceania | +1 | +
| MNHN-RA present tag | +Locality | +Sex | +Species | +Collector / date | +Old catalogue number | +Dum. Bibr. ca. 1864 catal. | +
|---|---|---|---|---|---|---|
| 2371 | +Not given | +F | +Bb | +
+Second exped. Dumont d’Urville (1837– 1840) on
+ |
+1736 | +Yes | +
| 2372 | +Tongatapu | +F? | +Bf | +
+First expedition Dumont d’Urville (1826– 1829) on
+ |
+1739 | +Yes Specimen measured in Duméril & Bibron, 1837 page 229 | +
| 5283 | +Fiji | +M | +Bb | +Filhol (around 1875) | +1736a | +No | +
| 6809 | +Tongatapu | +M | +Bf | +Arnoux (1845) | +1737 | +Yes | +
| 6811 | +No | +F | +Bb | +
+Second exped. Dumont d’Urville on
+ |
+1735 | +Yes | +
| 6812 | +No | +M | +Bf | +No | +1738 | +Yes “Origin?” | +
| 6813 | +Tongatapu | +M | +Bf | +
+First exp. Dumont d’Urville on
+ |
+1740 | +Yes | +
| MNHN-RA Species | +SVL | +TL | +Total length | +Pattern fitting illustration | +
|---|---|---|---|---|
| 2371 Bb | +17.3 | +6.5+(reg.) | +/ | +No | +
| 2372 Bf | +19.2 | +53.5 | +72.7 | +No | +
| 5283 Bb | +14.5 | +42.5 | +67 | +No | +
| 6809 Bf | +16 | +51.9 | +67.9 | +No | +
| 6811 Bb | +15.7 | +33.5 | +49.2 | +No | +
| 6812 Bf | +17 | +53.8 | +70.8 | +More-or-less | +
| 6813 Bf | +16.2 | +48.4 | +64.6 | +No | +
| Character | +
+ |
+Mean, Sd ♀ | +Range ♀ | +Mean, Sd 6 | +Range 6 | +Mean, Sd ♀+6 | +Range ♀+6 | +
|---|---|---|---|---|---|---|---|
| MSW5 | +19♀/156 | +152, 16.7 | +130–200 | +132, 13.2 | +99–145 | +143, 18.3 | +99–200 | +
| MSW5/TL | +19♀/156 | +30, 0.03 | +30–31 | +30, 0.04 | +20–36 | +30, 0.03 | +20–36 | +
| SW10 | +19♀/156 | +131, 15.7 | +103–166 | +110, 12.8 | +84–127 | +122, 17.7 | +84–166 | +
| SW10/TL | +19♀/156 | +26, 0.02 | +26–27 | +25, 0.03 | +19–28 | +26, 0.03 | +19–31 | +
| S1 | +19♀/156 | +78, 8.5 | +64–96 | +64, 6.5 | +54–79 | +72, 10.3 | +54–96 | +
| S2 | +19♀/156 | +49, 5.9 | +41–63 | +43, 4.8 | +36–51 | +47, 6.1 | +36–63 | +
| S3 | +19♀/156 | +53, 7.1 | +42–72 | +46, 3.8 | +40–54 | +50, 6.7 | +40–72 | +
| S4 | +19♀/156 | +59, 8.9 | +47–84 | +51, 3.5 | +43–58 | +55, 8.2 | +43–84 | +
| S5 | +19♀/156 | +62, 6.6 | +55–78 | +51, 2.8 | +46–57 | +57, 7.7 | +46–78 | +
| S6 | +19♀/156 | +64, 7.8 | +53–85 | +52, 4.5 | +46–61 | +58, 8.8 | +46–85 | +
| S7 | +19♀/156 | +67, 7 | +55–82 | +56, 3.5 | +50–63 | +62, 7.8 | +50–82 | +
| S8 | +19♀/156 | +69, 7.5 | +58–88 | +58, 3.1 | +50–62 | +64, 8.3 | +50–88 | +
| S9 | +19♀/156 | +73, 9.2 | +62–99 | +61, 3.8 | +55–68 | +68, 9.3 | +55–99 | +
| S10 | +19♀/156 | +77, 9.3 | +52–92 | +60, 5.6 | +54–74 | +70, 11.5 | +52–92 | +
| S11 | +19♀/156 | +32, 4.9 | +25–41 | +35, 5.8 | +22–43 | +33, 5.5 | +22–43 | +
| LTS | +17♀/146 | +46, 10.3 | +31–67 | +137, 14.2 | +104–162 | +87, 47.8 | +31–162 | +
| LTS/TL | +17♀/146 | +9, 0.02 | +7–11 | +31, 0.04 | +24–37 | +19, 0.1 | +7–37 | +
| Segment | +MD | +PD | +SD | +LD | +LV | +VL | +VM | +
|---|---|---|---|---|---|---|---|
| 1 | +ce* | +ss | +ss x2 | +ss | +|||
| 2 | +ce* | +se, ss | +ss | +se, ss | +se | +se (♀) | +tu (6), ss | +
| 3 | +ce* | +se, ss | +ss | +se, ss | +se | +se, ss | +|
| 4 | +ce* | +se, ss | +ss | +se, ss | +se | +se, ss | +|
| 5 | +ce* | +se, ss | +ss | +se, ss | +se | +se, ss | +|
| 6 | +ce* | +se, ss | +ss | +se | +se | +se, ss | +|
| 7 | +ce* | +se, ss | +ss | +se, ss | +se | +se, ss | +|
| 8 | +ce* | +se*, ss | +ss x2 | +se, ss | +se | +se, ss | +|
| 9 | +ce* | +se, ss | +ss x2 | +se, ss | +se, ne | +se, ss | +|
| 10 | +ss x2 | +se | +|||||
| 11 | +lts | +ps x2 (6) | +
| Measurement | +BM 9.11.19.1 (female), holotype | +MN 91624 (male) | +
|---|---|---|
| FL | +44.5 | +43.4 | +
| EL | +27.5 | +27.2 | +
| WMLE | +4.6 | +- | +
| MAL | +12.1 | +11.9 | +
| MAN | +6.9 | +7.1 | +
| COH | +4.6 | +- | +
| GLS | +18.3 | +16.9 | +
| CCL | +16.0 | +14.9 | +
| CIL | +17.0 | +- | +
| BAL | +15.2 | +- | +
| ZYG | +11.2 | +10.3 | +
| MAB | +9.1 | +8.9 | +
| BCB | +8.3 | +8.9 | +
| POB | +4.5 | +4.3 | +
| BAC | +5.0 | +4.5 | +
| BAM | +7.1 | +6.1 | +
| MTL | +6.4 | +5.8 | +
| M1M3 | +4.1 | +3.8 | +
| WFH | +4.1 | +- | +
| Diagnostic characters | +
+ |
+
+ |
+
|---|---|---|
| Prebranchial chaetiger | +3 | +|
| Branchial pairs | +10 to 13 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Measurements | +Holotype | +Paratypes | +Mean | +SD | +N | +
|---|---|---|---|---|---|
| Min.—Max. | +|||||
|
+ |
+|||||
| Total length (TL) | +160.0 | +104.0–145.0 | +- | +- | +10 | +
| Length to end of anal fin (LEA) | +113.0 | +76.0–115.0 | +- | +- | +10 | +
| Head length (HL) | +16.7 | +11.4–15.5 | +- | +- | +10 | +
|
+ |
+|||||
| Head length | +13.0 | +13.1–14.9 | +13.9 | +1.0 | +10 | +
| Snout-to-opercle length | +12.8 | +11.2–14.3 | +13.0 | +0.9 | +10 | +
| Preanal distance | +17.5 | +14.7–19.2 | +17.2 | +1.5 | +10 | +
| Prepectoral distance | +15.1 | +12.9–15.8 | +14.5 | +0.8 | +10 | +
| Snout-to-anus length | +7.7 | +7.0–8.2 | +7.5 | +0.4 | +10 | +
| Body depth at pectoral-fin tip | +15.4 | +12.1–16.2 | +14.7 | +1.3 | +10 | +
| Body depth at anal-fin origin | +14.0 | +11.0–15.1 | +13.2 | +1.1 | +10 | +
| Body width | +7.51 | +6.4–8.1 | +7.1 | +0.7 | +10 | +
| Anus to anal-fin origin distance | +10.7 | +7.0–11.6 | +9.7 | +1.4 | +10 | +
| Anal-fin length | +82.3 | +75.4–84.3 | +80.9 | +2.8 | +10 | +
| Pectoral-fin length | +8.8 | +7.5–8.8 | +8.4 | +0.4 | +10 | +
| Snout-to-occiput length | +11.7 | +9.5–12.6 | +11.5 | +0.9 | +10 | +
| Caudal-filament length | +41.5 | +22.8–38.9 | +32.3 | +5.6 | +10 | +
| Caudal filament width | +0.7 | +0.6–0.7 | +0.7 | +0.1 | +10 | +
| Caudal filament depth | +1.5 | +1.1–1.4 | +1.3 | +0.2 | +10 | +
|
+ |
+|||||
| Snout length | +33.6 | +32.2–35.2 | +33.8 | +0.8 | +10 | +
| Internasal distance | +10.8 | +9.3–11.9 | +10.3 | +0.7 | +10 | +
| Snout-to-posterior naris distance | +23.6 | +21.5–25.9 | +23.2 | +1.3 | +10 | +
| Posterior naris-to-orbit distance | +9.6 | +8.5–10.5 | +9.5 | +0.7 | +10 | +
| Internarial width | +15.8 | +13.6–18.8 | +15.6 | +1.5 | +10 | +
| Head width at opercle | +51.8 | +47.4–53.6 | +50.4 | +2.2 | +10 | +
| Head width at orbits | +41.6 | +36.1–42.3 | +40.2 | +2.1 | +10 | +
| Orbital diameter | +12.4 | +10.6–13.3 | +12.0 | +0.8 | +10 | +
| Postorbital distance | +58.2 | +56.8–62.1 | +58.9 | +1.3 | +10 | +
| Eye-to-opercle length | +47.6 | +44.5–48.4 | +46.4 | +1.2 | +10 | +
| Opercular opening | +25.8 | +21.0–25.0 | +23.0 | +1.5 | +10 | +
| Suborbital depth | +31.8 | +25.8–32.3 | +29.0 | +2.0 | +10 | +
| Interorbital distance | +25.9 | +25.2–30.1 | +28.0 | +1.7 | +10 | +
| Head depth at supraoccipital | +65.9 | +63.7–70.0 | +66.6 | +2.1 | +10 | +
| Head depth at orbits | +46.1 | +46.6–51.4 | +47.8 | +1.5 | +10 | +
| Mouth length | +24.6 | +19.9–23.8 | +21.9 | +1.5 | +10 | +
| Oral width | +23.7 | +23.5–26.0 | +24.7 | +1.0 | +10 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|---|---|---|---|---|---|
|
+ |
+A | +A* | +A* | +A | +A | +
|
+ |
+A | +A | +A | +A* | +A | +
|
+ |
+A | +A | +A | +A | +A | +
|
+ |
+A | +A | +A | +A | +A* | +
|
+ |
+A | +P | +P | +A* | +P | +
|
+ |
+P | +P | +P | +P | +P | +
|
+ |
+P | +P | +P | +P | +P | +
| Character | +IEBR 4741 (TQ.2018.140) | +IEBR 4742 (TQ.2018.141) | +IEBR 4743 (TQ.2018.96) | +Min–Max | +
|---|---|---|---|---|
| Holotype | +Paratype | +Paratype | +||
| Sex | +male | +male | +female | +|
| SVL | +43.58 | +43.45 | +41.44 | +41.44–43.58 | +
| TaL | +37.35 | +36.36 | +32.13 | +32.13–37.35 | +
| TrunkL | +21.29 | +21.27 | +20.89 | +20.89–21.29 | +
| EyeD | +2.53 | +2.61 | +2.68 | +2.53–2.68 | +
| HeadL | +10.65 | +10.22 | +9.96 | +9.96–10.65 | +
| HeadW | +7.36 | +7.09 | +6.87 | +6.87–7.36 | +
| NarEye | +3.07 | +3.15 | +2.81 | +2.81–3.15 | +
| SnEye | +3.81 | +4.02 | +3.54 | +3.54–4.02 | +
| SnW | +1.33 | +1.12 | +0.96 | +0.96–1.33 | +
| EarD | +0.69 | +0.55 | +0.79 | +0.55–0.79 | +
| Trunk/SVL | +0.49 | +0.49 | +0.50 | +0.49–0.50 | +
| HeadL/SVL | +0.24 | +0.24 | +0.24 | +0.24 | +
| HeadW/SVL | +0.17 | +0.16 | +0.17 | +0.16–0.17 | +
| HeadW/HeadL | +0.69 | +0.69 | +0.69 | +0.69 | +
| SnEye/HeadL | +0.36 | +0.39 | +0.36 | +0.36–0.39 | +
| NarEye/HeadL | +0.29 | +0.31 | +0.28 | +0.28–0.31 | +
| EyeD/HeadL | +0.24 | +0.26 | +0.27 | +0.24–0.27 | +
| SnW/HeadL | +0.12 | +0.11 | +0.10 | +0.10–0.12 | +
| EyeD/NarEye | +0.82 | +0.83 | +0.95 | +0.82–0.95 | +
| SnW/HeadW | +0.18 | +0.16 | +0.14 | +0.14–0.18 | +
| Dorsal scale rows | +22 | +18 | +23 | +18–23 | +
| Ventral scale rows | +9 | +9 | +13 | +9–13 | +
| Cloacal spurs (CloacS) | +1 | +1 | +1 | +1 | +
| Circumnasal (CircN) | +3/3 | +2/3 | +3/3 | +2–3 | +
| Scales between supranasals (SnS) | +4 | +6 | +4 | +4–6 | +
| Supralabials (Sublab) | +11/11 | +11/12 | +12/10 | +10–12 | +
| Infralabials (Inflab) | +10/9 | +11/10 | +11/10 | +9–11 | +
| Chin scale (Chin) | +8 | +8 | +9 | +8–9 | +
| Lamella formula of | +||||
| forelimbs (pairs only) | +3-4-5-4 | +3-4-5-4 | +3-4-5-4 | +3-4-5-4 | +
| hindlimbs (pairs only) | +4-5-5-5 | +4-5-5-5 | +4-5-5-5 | +4-5-5-5 | +
| First digit of | +||||
| forelimbs | +3 | +3 | +3 | +3 | +
| hindlimbs | +3 | +3 | +3 | +3 | +
| Pores | +24 | +22 | +0 | +22–24 (in males) | +
| Subcaudals enlarged, plate-like (1) or not (0) | +0 | +0 | +0 | +0 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+179650 | +
| KEINE ANGABE | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
| Kein internationaler, nationaler oder kantonaler Schutz | +
| Ch. / Stage | +L. | +P. | +D. (♀) | +D. (♂) | +A. (♀) | +A. (♂) | +
|---|---|---|---|---|---|---|
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | +- | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | +- | ++ | ++ | ++ | ++ | +
|
+ |
+- | +- | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | +- | ++ | ++ | ++ | ++ | +
|
+ |
+- | +- | ++ | +- | ++ | +- | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | +- | ++ | +- | ++ | +- | +
|
+ |
++ | ++ | ++ | ++ | ++ | ++ | +
|
+ |
+- | +- | ++ | +- | ++ | +- | +
| Spermadactyl | +- | +- | +- | +- | +- | ++ | +
| Macrosetae on basitarsus IV | +- | ++ | ++ | ++ | ++ | ++ | +
| Femora I-IV* | +10,7,5,- | +10,7,5,4 | +12,10,6,6 | +12,10,6,6 | +12,10,6,6 | +12,10,6,6 | +
| Genua I-IV* | +8,6,6,- | +8,6,6,5 | +10,7,7,7 | +10,7,7,7 | +10,7,7,7 | +10,7,7,7 | +
| Tibiae I-IV* | +8,7,7,- | +8,7,7,6 | +10,7,7,6 | +10,7,7,6 | +10,7,7,6 | +10,7,7,6 | +
|
+4.2 -
+ |
+
|
+4.2.4 -
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+337700 | +
| = | +
+ |
+
+ |
+198 | +
| = | +
+ |
+
+ |
+173 | +
| = | +
+ |
+
+ |
+173 | +
| = | +
+ |
+
+ |
+337700 | +
| = | +
+ |
+
+ |
+1192 | +
| = | +
+ |
+
+ |
+1027 | +
| = | +
+ |
+
+ |
+337700 | +
| = | +
+ |
+
+ |
+398 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+nicht
+ |
+|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+L - Leitart | +weitere Informationen | +
| This study | +
+ |
+||
|---|---|---|---|
|
+Non-types Yonaguni-jima island
+ |
+Holotype Philippines | +
+Paratypes Western Pacific
+ |
+|
| Standard length (SL; mm) | +31.6–38.2 | +35.4 | +30.4–51.0 | +
| Dorsal-fin rays | +III, 23 | +III, 23 | +III, 22–23 | +
| Anal-fin rays | +III, 14 | +III, 14 | +III, 14 | +
| Pectoral-fin rays | +16–17 | +17 | +16–19 | +
| Upper procurrent caudal-fin rays | +6 | +— | +6–7 | +
| Lower procurrent caudal-fin rays | +5 | +5 | +5–6 | +
| Total caudal-fin rays | +29 | +28 | +28–30 | +
| Scales in lateral series | +35 | +35 | +29–37 | +
| Anterior lateral-line scales | +24–28 | +28 | +24–31 | +
| Anterior lateral line terminating beneath segmented dorsal-fin ray | +15–18th | +17th | +15–21st | +
| Posterior lateral-line scales | +5–9+1 | +7+1 | +3–13+0–2 | +
| Scale rows between lateral lines | +3 | +3 | +2–4 | +
| Horizontal scale rows above anal-fin origin | +14 | +15 | +14–18 | +
| Circumpeduncular scales | +14–16 | +14 | +14–16 | +
| Pre-dorsal-fin scale rows | +14–15 | +15 | +13–18 | +
| Gill rakers | +4–5+11=15–16 | +4+10=14 | +3–5+9–11=13–16 | +
| Pseudobranch filaments | +10 | +10 | +7–14 | +
| Circumorbital pores | +17–18 | +20 | +16–26 | +
| Preopercular pores | +8 | +10 | +8–16 | +
| Dentary pores | +4 | +4 | +4–5 | +
| Posterior interorbital pores | +0 | +0 | +0 | +
| This study | +
+ |
+||
|---|---|---|---|
|
+Non-types Yonaguni-jima island
+ |
+Holotype Philippines | +
+Paratypes Western Pacific
+ |
+|
| Standard length (SL; mm) | +31.6–38.2 | +35.4 | +30.4–51.0 | +
| Head length | +26.4–27.3 | +25.1 | +23.1–27.0 | +
| Orbit diameter | +8.5–9.0 | +9.9 | +8.0–10.9 | +
| Snout length | +6.0–6.5 | +5.1 | +4.8–6.3 | +
| Fleshy interorbital width | +4.1–5.2 | +4.8 | +4.5–5.6 | +
| Bony interorbital width | +3.0–3.9 | +3.1 | +2.8–3.7 | +
| Body width | +14.2–14.6 | +12.4 | +11.4–14.8 | +
| Distance from snout tip to posterior tip of retroarticular bone | +13.9–14.9 | +14.7 | +13.2–15.1 | +
| Pre-dorsal-fin length | +33.9–35.1 | +32.8 | +30.5–35.2 | +
| Pre-pelvic-fin length | +32.0–34.3 | +32.2 | +29.6–33.4 | +
| Distance from posterior tip of retroarticular bone to pelvic-fin origin | +17.7–21.5 | +18.6 | +17.3–20.5 | +
| Distance from dorsal-fin origin to pelvic-fin origin | +27.6–28.2 | +28.0 | +26.9–30.6 | +
| Distance from dorsal-fin origin to base of middle part of dorsal-fin ray | +35.8–36.4 | +37.3 | +35.0–37.8 | +
| Distance from dorsal-fin origin to anal-fin origin | +39.9–40.7 | +41.5 | +39.4–42.5 | +
| Distance from pelvic-fin origin to anal-fin origin | +21.5–25.9 | +26.3 | +25.3–30.6 | +
| Distance from middle of dorsal-fin ray base to end of dorsal-fin base | +24.9–26.3 | +26.3 | +23.3–27.3 | +
| Distance from middle of dorsal-fin ray base to anal-fin origin | +27.5–29.8 | +28.8 | +26.0–27.9 | +
| Distance from anal-fin origin to end of dorsal-fin base | +37.4–38.3 | +37.3 | +35.0–39.0 | +
| Anal-fin base length | +31.0–31.9 | +30.8 | +28.1–32.0 | +
| Distance from end of dorsal-fin base to end of anal-fin base | +16.1–16.5 | +15.8 | +13.3–16.8 | +
| Distance from end of dorsal-fin base to end of dorsal edge of caudal peduncle | +10.9–11.1 | +10.2 | +8.9–11.4 | +
| Distance from end of dorsal-fin base to end of ventral edge of caudal peduncle | +16.9–18.7 | +18.1 | +15.9–18.7 | +
| Distance from end of anal-fin base to end of dorsal edge of caudal peduncle | +18.3–19.6 | +18.9 | +17.1–20.1 | +
| Distance from end of anal-fin base to end of ventral edge of caudal peduncle | +10.7–11.5 | +11.9 | +10.7–12.9 | +
| 1st dorsal-fin spine length | +1.9–2.4 | +2.3 | +1.7–3.2 | +
| 2nd dorsal-fin spine length | +3.4–4.4 | +4.5 | +4.1–5.7 | +
| 3rd dorsal-fin spine length | +5.8–7.0 | +7.1 | +5.8–7.7 | +
| 1st segmented dorsal-fin ray length | +10.8–11.5 | +13.0 | +9.9–13.8 | +
| Length of 4th-to-last segmented dorsal-fin ray | +16.4 | +18.9 | +17.7–21.8 | +
| 1st anal-fin spine length | +1.9–2.2 | +2.3 | +1.6–3.9 | +
| 2nd anal-fin spine length | +4.1–4.7 | +4.8 | +3.9–6.7 | +
| 3rd anal-fin spine length | +6.3–7.1 | +7.6 | +5.8–9.0 | +
| First segmented anal-fin ray length | +10.8–11.7 | +12.1 | +10.6–13.6 | +
| Length of 4th-to-last segmented anal-fin ray | +16.2–18.7 | +18.1 | +17.1–20.8 | +
| Third pectoral-fin ray length | +16.1–18.6 | +18.4 | +14.5–19.3 | +
| Pelvic-fin spine length | +9.8–9.9 | +9.9 | +8.8–11.0 | +
| Second segmented pelvic-fin ray length | +21.5–23.4 | +23.2 | +21.2–24.4 | +
| Caudal-fin length | +25.7–27.5 | +28.2 | +25.2–32.3 | +
| n= | +b.l. | +w.l. | +A.R. | +sR | +sR1 | +sR4+5 | +sq | +ac | +|
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+1 | +6.5 | +2.8 | +2.6 | +n/a | +n/a | +n/a | +8 | +0 | +
|
+ |
+1–2 | +6.5–6.9 | +2.5–2.7 | +2.4 | +23–26 | +22–23 | +0–1 | +11 | +0 | +
|
+ |
+? | +3.2 | +2.8 | +2.5 | +n/a | +n/a | +n/a | +n/a | +n/a | +
|
+ |
+9 | +3.5–4.0 | +1.8–2.0 | +2.6 | +32 | +28 | +22–24 | +10–12 | +2 | +
|
+ |
+1–2 | +6.0–6.2 | +2.8–2.9 | +n/a | +27–33 | +23–27 | +17 | +6–9 | +0 | +
|
+ |
+1 | +6 | +2.4 | +2.0 | +30 | +16 | +14 | +4 | +0 | +
| continued. | +
| dc | +pa | +l.apn | +sct | +LR1 | +TIX setae gs.l. gs.R. | +
|---|---|---|---|---|---|
|
+ |
+3 | +n/a | +6 | +n/a | +?0 220 3.1 | +
|
+ |
+3 | +4–6 | +5–7 | +n/a | +3–6 205–210 3.3 | +
|
+ |
+n/a | +n/a | +n/a | +1.4 | +n/a n/a 3.0 | +
|
+ |
+9–10 | +3–4 n/a | +8 | +1.4 | +n/a | +n/a 3.2 | +
|---|---|---|---|---|---|---|
|
+ |
+9–10 | +3 3–5 | +9–12 | +n/a | +3–5 | +200–215 3.6–3.8 | +
|
+ |
+9–10 | +3 6 | +9 | +n/a | +6 | +170–175 3.0–3.2 | +
| 1 | +Antero-inferior corner of pronotum with distinct tooth-like process (Fig. 1D) | +2 | +
| - | +Antero-inferior corner of pronotum without tooth-like process (Fig. 1E) | +4 | +
| 2 | +Head (Fig. 12B), sides of petiole and terga 1 and 2 of gaster smooth and polished, integument with bluish luster (Fig. 12A); southeastern coast of Brazil | +
+ |
+
| - | +Head, lateral sides of the petiole and terga 1 and 2 of gaster finely micro-punctate/scaled (Fig. 12B) | +3 | +
| 3 | +Total body length under 3 cm; Bolivia, Paraguay, northern Argentina and southern Brazil | +
+ |
+
| - | +Total body length over 3 cm; Brazil, Peru, and Guyana | +
+ |
+
| 4 | +
+Body covered in bristle-like setae which are not flagellate (Fig. 1C);
+ |
+
+ |
+
| - | +Body covered in fine and flagellate setae (Fig. 1B) | +5 | +
| 5 | +
+Appressed golden yellow pubescence present on frons (Fig. 1A); Colombia south to
+ |
+
+ |
+
| - | +Pubescence absent from frons or not golden yellow | +6 | +
| 6 | +Sides of head, lateral sides of the petiole and terga 1 and 2 of gaster finely micro-punctate/scaled (Fig. 12B); in profile antero-dorsal edge of petiole bulging (Fig. 1G); northeast Brazil | +
+ |
+
| - | +Sides of head, sides of the petiole and terga 1 and 2 of gaster smooth, shiny and polished with no microsculpturing, integument with bluish luster (Fig. 12A); in profile petiole with even dorsal corners (Fig. 1F); eastern Bolivia, northern Paraguay and southwestern Brazil | +
+ |
+
| 1 | +Total body length less than 15 mm | +
+ |
+
| - | +Total body length greater than 15 mm | +2 | +
| 2 | +Subpetiolar process in form of spine; pronotum heavily sculptured; palp formula 5:3 | +
+ |
+
| - | +Subpetiolar process without spine; pronotum shiny, microsculptured; palp formula 4:3 | +3 | +
| 3 | +Ocelli protruding on occipital margin of head (Fig. 4A-D) | +4 | +
| - | +Ocelli not protruding on occipital margin of head (Fig. 4E); Bolivia, Paraguay, northern Argentina and southern Brazil | +
+ |
+
| 4 | +Setae on funiculus long and erect (Fig. 4F, G) | +5 | +
| - | +Setae on funiculus short, stiff and subdecumbent, or minute pubescence present (Fig. 4H, I, J) | +6 | +
| 5 | +Digitus volsellaris with toothless lobe at distal end (Fig. 10C); Brazil, Peru, and Guyana | +
+ |
+
| - | +Digitus volsellaris without lobe at distal end (Fig. 10A); northeast Brazil | +
+ |
+
| 6 | +
+Pygidium terminating in narrow, elongate spine (Fig. 4M); penis valve of aedeagus wedge-shaped in lateral view (Fig. 11E); Colombia south to
+ |
+
+ |
+
| - | +Pygidium terminating in short, broad, triangular angle (Fig. 4N); penis valve of aedeagus with distal flange and triangular ventral lobe (Fig. 11B); Mato Grosso do Sul, Brazil | +
+ |
+
| 1 | +Esquina antero-inferior del pronoto con proceso en forma de diente distinto (Fig. 1D) | +2 | +
| - | +Esquina antero-inferior del pronoto sin proceso en forma de diente (Fig. 1E) | +4 | +
| 2 | +
+Cabeza, lados laterales del
+ |
+
+ |
+
| - | +
+Cabeza, lados laterales del
+ |
+3 | +
| 3 | +Largo total menos de 3 cms | +
+ |
+
| - | +
+Largo total
+ |
+
+ |
+
| 4 | +Cuerpo cubierto con pelos gruesos, no flagelados (Fig. 1C) | +
+ |
+
|
+ |
+Cuerpo cubierto con pelos finos y flagelados (Fig. 1B) | +5 | +
| 5 | +Frente con pubescencia recostada y amarillo-dorada (Fig. 1A) | +
+ |
+
| - | +Frente sin pubescencia recostada y amarilla | +6 | +
| 6 | +
+Lados de la cabeza, lados del
+ |
+
+ |
+
| - | +
+Lados de la cabeza, lados del
+ |
+
+ |
+
| 1 | +Largo total menos de 15 mm | +
+ |
+
| - | +
+Largo total
+ |
+2 | +
| 2 | +Proceso subpetiolar en forma de espina; pronoto fuertemente esculturado; formula palpal 5:3 | +
+ |
+
| - | +Proceso subpetiolar no en forma de espina; pronoto liso, microesculturado; formula palpal 4:3 | +3 | +
| 3 | +Ocelos muy hinchados en el margen occipital de la cabeza (Fig. 4A-D) | +4 | +
| - | +Ocelos no muy hinchados en el margen occipital de la cabeza (Fig. 4E) | +
+ |
+
| 4 | +
+ |
+5 | +
| - | +
+ |
+6 | +
| 5 | +
+ |
+
+ |
+
| - | +
+Digito del volsela sin
+ |
+
+ |
+
| 6 | +
+Pigidio terminando en un espina, alargada y delgada (Fig. 4M);
+ |
+
+ |
+
| - | +
+Pigidio terminando en una espina corta y ancha, en forma de
+ |
+
+ |
+
| 1 | +Esquina antero-inferior do pronoto com processo em forma de dente distinto (Fig. 1D) | +2 | +
| - | +Esquina antero-inferior do pronoto sem processo em forma de dente (Fig. 1E) | +4 | +
|
+ |
+
+ |
+
+ |
+
| - | +
+Cabeza, lados laterais do
+ |
+3 | +
| 3 | +Comprido total menos 3 cms | +
+ |
+
| - | +Comprido total mais de 3 cms | +
+ |
+
| 4 | +
+Corpo cobrido com cogumelos em forma de cerdas, que
+ |
+
+ |
+
| - | +Corpo cobrido com cogumelos finos e flagelados (Fig. 1B) | +5 | +
| 5 | +Frons com pubescencia prendida e amarela-dourada (Fig. 1A) | +
+ |
+
| - | +Frons sem pubescencia amarelo prendido | +6 | +
| 6 | +
+Lados da
+ |
+
+ |
+
| - | +
+Lados da
+ |
+
+ |
+
| 1 | +Comprido total menos de 15mm | +
+ |
+
| - | +Comprido total mais de 15 mm | +2 | +
| 2 | +Proceso subpetiolar em forma de espinha; pronoto fortemente esculturado; formula palpular 5:3 | +
+ |
+
| - | +
+Processo subpetiolar
+ |
+3 | +
| 3 | +
+Ocelos na margem occipital da
+ |
+4 | +
| - | +
+Ocelos na margem occipital da
+ |
+
+ |
+
| 4 | +
+ |
+5 | +
| - | +
+ |
+6 | +
| 5 | +
+ |
+
+ |
+
| - | +
+ |
+
+ |
+
| 6 | +
+Pigidio acabado numa espinha, alongada e delgada (Fig. 4M);
+ |
+
+ |
+
| - | +
+Pigidio acabado numa espinha apara e alarga, em forma de
+ |
+
+ |
+
| Specimens No. | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+summarizing of measurement (minimum-maximum, mean
+ |
+
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Sex | +||||||||||||||
| SVL | +||||||||||||||
| HDL | +||||||||||||||
| HDW | +||||||||||||||
| SNT | +||||||||||||||
| IND | +||||||||||||||
| IOD | +||||||||||||||
| ED | +||||||||||||||
| TD | +||||||||||||||
| TED | +||||||||||||||
| HND | +||||||||||||||
| RAD | +||||||||||||||
| FTL | +||||||||||||||
| TIB | +||||||||||||||
| HDL/SVL | +||||||||||||||
| HDW/SVL | +||||||||||||||
| HDW/HDL | +||||||||||||||
| SNT/HDL | +||||||||||||||
| SNT/SVL | +||||||||||||||
| IND/HDW | +||||||||||||||
| IOD/HDW | +||||||||||||||
| ED/HDL | +||||||||||||||
| ED/SVL | +||||||||||||||
| TD/ED | +||||||||||||||
| TED/TD | +||||||||||||||
| HND/SVL | +||||||||||||||
| RAD/SVL | +||||||||||||||
| FTL/SVL | +||||||||||||||
| TIB/SVL | +
| one-note call (n = 25) | +two-note call (n = 59) | +three-note call (n = 3) | +
|---|---|---|
| Call duration (ms) | +||
| Note duration (ms) | +st | +st | +
| nd | +nd | +|
| rd | +||
| Note rise time (ms) | +st | +st | +
| nd | +nd | +|
| rd | +||
| Note interval (ms) | +st | +|
| nd | +||
| Call PF (Hz) | +||
| Note PF (Hz) | +st | +st | +
| nd | +nd | +|
| rd | +||
| Note IQR-BW (Hz) | +st | +st | +
| nd | +nd | +|
| rd | +
| Group/species | +Body colour | +Antennalsegments | +Propodeum | +Crenulae | +
|---|---|---|---|---|
|
+ |
+Rather strongly | +4 6, ♁ 4 | +Relatively long; | +Large, overlapping; or small, | +
| pigmented | +or short | +nonoverlapping | +||
|
+ |
+Greyish or dusky; black | +4 6, ♁ 6 | +Relatively long; | +Elongate, relatively narrow, | +
| cephalic | +or short | +nonoverlapping; or | +||
| pigmentation | +rounded, slightly | +|||
| overlapping | +||||
|
+ |
+Generally yellow with | +4 6, ♁ 6 | +Relatively short | +Small, nonoverlapping | +
| coarse, dark thoracic | +||||
| setae | +||||
|
+ |
+Yellow | +4 6, ♁ 6 | +Relatively long | +Large, overlapping | +
|
+ |
+Yellow | +4 6, ♁ 6 | +Very long | +Small, nonoverlapping | +
|
+ |
+Yellow | +4 5, ♁ 5 | +Short | +Small, nonoverlapping, | +
| generally forming one | +||||
| continuous row | +||||
|
+ |
+Yellow | +4 6, ♁ 5 | +Short | +Large,distinctlyelongate, | +
| sp.n. | +sharply tooth-shaped, | +|||
| nonoverlapping | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+177900 | +
| = | +
+ |
+
+ |
+177900 | +
| = | +
+ |
+
+ |
+2801 | +
| = | +
+ |
+
+ |
+177900 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
| Carapace general shape | +Trapezoidal | +Squarish | +Squarish | +Squarish | +Squarish | +Squarish | +Squarish but | +Squarish | +Squarish but | +
| (cw./cl. ratio) | +(mean 1.06, | +(mean 1.09, | +(mean 1.13, | +(mean 1.12, | +(mean 1.11, | +(mean 1.13, | +relatively | +(mean 1.14, | +relatively | +
| SD ± 0.12, n | +SD ± 0.06, n | +SD ± 0.03, n | +SD ± 0.02, n | +SD ± 0.02, | +SD ± 0.03, n | +wide | +SD ± 0.05, n | +wide | +|
| = 4) | += 3) | += 17) | += 8) | +n = 10) | += 5) | +(mean 1.20, | += 10) | +(mean 1.17, | +|
| (e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. 7A) | +(e.g. Fig. 9A) | +(e.g. Fig. | +SD ± 0.00, n | +(e.g. Fig. | +SD ± 0.03, n | +|
| 1A) | +3A) | +4A) | +11A) | += 23) | +15A) | += 6) | +|||
| (e.g. Fig. | +(e.g. Fig. | +||||||||
| 13A) | +17A) | +||||||||
| Carapace granulation on frontal | +Finely | +Finely | +Finely | +Coarsely | +Finely | +Finely | +Finely | +Finely | +Finely | +
| region | +granular | +granular | +granular | +granular | +granular | +granular | +granular | +granular | +granular | +
| (e.g. Fig.1C) | +(e.g. Fig. 3F) | +(e.g. Fig. | +(e.g. Fig. 7C) | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +|
| 4C) | +9A) | +11C) | +13C) | +15C) | +17C) | +||||
| Shape of frontal border in dorsal | +Nearly | +Concave in | +Concave in | +Strongly | +Concave in | +Strongly | +Strongly | +Concave in | +Concave in | +
| view | +straight | +medial part | +medial part | +concave in | +medial part | +concave in | +concave in | +medial part | +medial part | +
| (e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +medial part | +(e.g. Fig. 9C) | +medial part | +medial part | +(e.g. Fig. | +(e.g. Fig. | +|
| 1A) | +3A) | +4A) | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +15A) | +17A) | +||
| 7A) | +11A) | +13A) | +|||||||
| Granulation on outer surface | +Relatively | +Relatively | +Relatively | +Relatively | +Relatively | +Relatively | +Relatively | +Relatively | +Relatively | +
| of palm | +robust | +minute | +robust (Fig. | +robust (Fig. | +minute (Fig. | +minute (Fig. | +robust (Fig. | +robust (Fig. | +robust | +
| (Fig. 19A) | +(Fig. 3D) | +19B) | +19C) | +19D) | +19E) | +19F) | +19G) | +(Fig. 19H) | +|
| Male pectinated crests on palm | +2 | +2 | +2 or 3 | +2 | +2 or 3 | +2 | +2 | +2 | +2 | +
| (Fig. 2C) | +(Fig. 3E) | +(most 2) | +(Fig. 8C) | +(most 2) | +(Fig. 12C) | +(Fig. 14C) | +(Fig. 16C) | +(Fig. 18C) | +|
| (Fig. 5C) | +(Fig. 10C) | +||||||||
| Male dactylar tubercles | +7 | +9 | +10–15 | +6–8 | +9–12 | +7 or 8 | +9 | +10 | +11 | +
| (e.g. Fig. | +(e.g. Fig. | +(average 11) | +(average 7) | +(average 10) | +(average 8) | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +|
| 19A) | +3D) | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +19F) | +19G) | +19H) | +|
| 19B) | +19C) | +19D) | +19E) | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
| Dorsal margin of ambulatory | +Straight | +Straight | +Straight | +Upturned | +Straight | +Straight to | +Straight to | +Straight to | +Straight | +
|---|---|---|---|---|---|---|---|---|---|
| merus | +(e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +distally | +(e.g. Fig. | +slightly | +slightly | +slightly | +(e.g. Fig. | +
| 1E) | +3G) | +2 0A) | +(e.g. Fig. | +2 0C) | +upturned | +upturned | +upturned | +2 0G) | +|
| 2 0B) | +distally | +distally | +distally | +||||||
| (e.g. Fig. | +(e.g. Fig. | +(e.g. Fig. | +|||||||
| 20D) | +20E) | +20F) | +
| Merus of P3 | +Very broad | +Broad | +Broad | +Broad | +Broad | +Slender | +Broad | +Broad | +Very broad | +
|---|---|---|---|---|---|---|---|---|---|
| (length/width ratio) | +(mean 2.40, | +(mean 2.60, | +(mean 2.48, | +(mean 2.52, | +(mean 2.58, | +(mean 3.05, | +(mean 2.56, | +(mean 2.50, | +(mean 2.34, | +
| SD ± 0.26, n | +SD ± 0.04, n | +SD ± 0.11, n | +SD ± 0.20, n | +SD ± 0.14, n | +SD ± 0.61, n | +SD ± 0.16, n | +SD ± 0.20, n | +SD ± 0.19, n | +|
| = 4) | += 2) | += 12) | += 8) | += 10) | += 5) | += 9) | += 9) | += 6) | +|
| Proodi of P3 | +Broad | +Broad | +Broad | +Broad | +Broad | +Slender | +Slender | +Broad | +Very broad | +
| (length/width ratio) | +(mean 4.13, | +(mean 4.38, | +(mean 4.83, | +(mean 4.99, | +(mean 4.35, | +(mean 5.87, | +(mean 5.65, | +(mean 4.84, | +(mean 3.87, | +
| SD ±0.48, n | +SD ± 0.49, n | +SD ± 0.40, n | +SD ± 0.26, n | +SD ± 0.26, n | +SD ± 0.75, n | +SD ± 0.72, n | +SD ± 0.30, n | +SD ± 0.37, n | +|
| = 4) | += 2) | += 12) | += 8) | += 10) | += 5) | += 9) | += 9) | += 6) | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +sauer (pH 3.5-6.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +2 - Schwerpunktlebensraum | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+219900 | +
| = | +
+ |
+
+ |
+2443 | +
| = | +
+ |
+
+ |
+2615 | +
| = | +
+ |
+
+ |
+2615 | +
| = | +
+ |
+
+ |
+219900 | +
| = | +
+ |
+
+ |
+580 | +
| = | +
+ |
+
+ |
+509 | +
| = | +
+ |
+
+ |
+219900 | +
| = | +
+ |
+
+ |
+2146 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +regional beziehungsweise in der Schweiz ausgestorben (Regionally Extinct) | +|
| Alpennordflanke (NA) | +-- | +|
|
+ |
+vom Aussterben bedroht (Critically Endangered) | +B1ab(iii); B2ab(iii) | +
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +-- | +
|
+ |
+
+2 - Hohe nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+1 - Gering | +
|
+ |
+
+2 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(23.01.2013) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(25.07.2007) | +
|
+ |
+Z - Zielart | +weitere Informationen | +
| Morphometric characters | +Holotype | +
+ |
+
+ |
+
|---|---|---|---|
| Standard length (mm) | +30.6 | +28.6–31.5 | +|
| In percent of standard length | +|||
| Predorsal length | +41.3 | +39.9–44.5 | +27.5±1.0 | +
| Preanal length | +72.2 | +71.3–74.2 | +72.8±1.1 | +
| Prepelvic length | +56.0 | +47.9–56 | +53.0±2.6 | +
| Prepectoral length | +25.8 | +24.3–28.5 | +26.6±1.3 | +
| Length of dorsal fin base | +13.9 | +11.6–15.5 | +13.9±1.2 | +
| Dorsal-fin spine length | +15.3 | +12.3–16.8 | +14.8±1.2 | +
| Length of anal-fin base | +12.1 | +12.1–14.6 | +13.0±0.9 | +
| Pelvic-fin length | +16.6 | +13.2–16.6 | +15.1±1.1 | +
| Pectoral-fin length | +23.2 | +20.1–24.1 | +22.0±1.2 | +
| Pectoral fin spine length | +19.2 | +18.2–21.0 | +52.5±2.4 | +
| Caudal fin length | +22.4 | +20.7–24.5 | +19.7±1.0 | +
| Length of adipose-fin base | +15.6 | +12.1–17.8 | +15.0±1.8 | +
| Dorsal to adipose distance | +14.9 | +13.0–16.4 | +14.6±1.2 | +
| Post-adipose distance | +12.9 | +11.9–15.5 | +13.9±1.1 | +
| Caudal peduncle length | +15.2 | +14.6–18.2 | +16.3±1.2 | +
| Caudal peduncle depth | +9.0 | +9.0–10.5 | +9.7±0.5 | +
| Body depth at anus | +16.5 | +15.6–17.7 | +16.6±0.5 | +
| Head length | +28.2 | +25.9–29.2 | +27.5±1.0 | +
| Head width at pectoral fin origin | +25.2 | +19.7–25.3 | +23.1±2.0 | +
| Head depth just posterior to eye | +15.9 | +13.7–18.9 | +16.6±1.5 | +
| % HL | +|||
| Snout length | +53.8 | +45.3–53.8 | +50.2±2.7 | +
| Interorbital width | +26.8 | +22.7–28.1 | +25.5±1.7 | +
| Eye diameter | +11.6 | +11.5–21.3 | +15.0±3.2 | +
| Nasal barbel length | +19.5 | +8.6–19.5 | +12.5±3.1 | +
| Maxillary barbel length | +47.7 | +47.7–62.9 | +57.9±4.9 | +
| Outer mandibular barbel length | +40.7 | +27.4–54.5 | +37.3±7.6 | +
| Inner mandibular barbel length | +31.1 | +23.6–34.3 | +28.3±3.7 | +
| Mean | +SD | +Holotype | +Paratypes (N=2) | +|
|---|---|---|---|---|
| Total length | +18.67 | +0.15 | +18.5 | +18.7–18.8 | +
| Head length (excluding neck) | +1.67 | +0.05 | +1.6 | +1.7 | +
| Anteocular portion length | +0.5 | +0.0 | +0.5 | +0.5 | +
| Postocular portion length | +0.6 | +0.0 | +0.6 | +0.6 | +
| Head width across eyes | +1.1 | +0.0 | +1.1 | +1.1 | +
| Interocular distance | +0.53 | +0.05 | +0.6 | +0.5 | +
| Transverse width of eye | +0.3 | +0.0 | +0.3 | +0.3 | +
| Antennal segment I length | +10.5 | +0.36 | +10.9 | +10.2–10.4 | +
| Antennal segment II length | +10.97 | +0.23 | +11.1 | +10.7–11.1 | +
| Ant. segment III length | +0.87 | +0.05 | +0.8 | +0.9 | +
| Ant. segment IV length | +2.37 | +0.05 | +2.4 | +2.3–2.4 | +
| Labial segment II length | +0.43 | +0.05 | +0.4 | +0.4–0.5 | +
| Labial segment III length | +0.6 | +0.0 | +0.6 | +0.6 | +
| Labial segment IV length | +1.0 | +0.0 | +1.0 | +1.0 | +
| Fore lobe of pronotum length | +1.53 | +0.05 | +1.5 | +1.5–1.6 | +
| Petiole length | +2.33 | +0.11 | +2.4 | +2.2–2.4 | +
| Hind lobe of pronotum length | +1.93 | +0.05 | +1.9 | +1.9–2.0 | +
| Hind lobe of pronotum max. width | +1.6 | +0.0 | +1.6 | +1.6 | +
| Forewing length | +11.6 | +0.1 | +11.7 | +11.5–11.6 | +
| Hindwing length | +9.6 | +0.1 | +9.7 | +9.5–9.6 | +
| Fore coxa length | +3.23 | +0.15 | +3.1 | +3.2–3.4 | +
| Fore femur length | +7.03 | +0.11 | +7.1 | +6.9–7.1 | +
| Fore tibia length | +6.03 | +0.05 | +6.0 | +6.0–6.1 | +
| Fore tarsus length | +0.5 | +0.0 | +0.5 | +0.5 | +
| Mid femur length | +13.6 | +0.2 | +13.8 | +13.4–13.6 | +
| Mid tibia length | +20.76 | +0.89 | +21.8 | +20.2–20.3 | +
| Mid tarsus length | +0.5 | +0.0 | +0.5 | +0.5 | +
| Hind femur length | +18.23 | +0.25 | +18.2 | +18.0–18.5 | +
| Hind tibia length | +27.0 | +1.32 | +27.5 | +25.5–28.0 | +
| Hind tarsus length | +0.6 | +0.0 | +0.6 | +0.6 | +
| Abdomen length | +9.56 | +0.11 | +9.5 | +9.5–9.7 | +
| Abdomen maximum width | +1.3 | +0.1 | +1.4 | +1.2–1.3 | +
|
+ |
+
+ |
+view, scale bar 2.0 mm, 64, forewings, 65, venation of | +
|---|---|---|
| right forewing, 66, petiole, hind | +lobe of pronotum, left forewing, except | +apex and right hind wing (pointed by the arrow, right | +
| forewing moved laterally). | +
| Mean | +SD | +Maximum | +Minimum | +|
|---|---|---|---|---|
| Total length | +22.14 | +0.72 | +23.0 | +21.1 | +
| Head length (excluding neck) | +1.96 | +0.08 | +2.0 | +1.8 | +
| Anteocular portion length | +0.74 | +0.05 | +0.8 | +0.7 | +
| Postocular portion length | +0.84 | +0.05 | +0.9 | +0.8 | +
| Head width across eyes | +1.0 | +0.0 | +1.0 | +1.0 | +
| Interocular distance | +0.6 | +0.0 | +0.6 | +0.6 | +
| Transverse width of eye | +0.2 | +0.0 | +0.2 | +0.2 | +
| Antennal segment I length | +11.66 | +0.59 | +12.4 | +10.8 | +
| Antennal segment II length | +11.56 | +0.67 | +12.5 | +10.7 | +
| Ant. segment III length | +0.9 | +0.0 | +0.9 | +0.9 | +
|
+Ant. segment IV length
+ |
+2.3 | +0.25 | +2.6 | +2.0 | +
| Labial segment II length | +0.56 | +0.05 | +0.6 | +0.5 | +
| Labial segment III length | +0.7 | +0.0 | +0.7 | +0.7 | +
| Labial segment IV length | +1.16 | +0.05 | +1.2 | +1.1 | +
| Fore lobe of pronotum length | +1.56 | +0.11 | +1.7 | +1.4 | +
| Petiole length | +3.88 | +0.25 | +4.1 | +3.6 | +
| Hind lobe of pronotum length | +1.78 | +0.17 | +2.0 | +1.5 | +
| Hind lobe of pronotum max. width | +1.2 | +0.0 | +1.2 | +1.2 | +
| Forewing length | +7.06 | +0.18 | +7.3 | +6.9 | +
| Hindwing length | +1.4 | +0.12 | +1.6 | +1.3 | +
| Fore coxa length | +3.9 | +0.1 | +4.0 | +3.8 | +
| Fore femur length | +8.58 | +0.4 | +9.1 | +8.0 | +
| Fore tibia length | +7.28 | +0.32 | +7.8 | +6.9 | +
| Fore tarsus length | +0.62 | +0.02 | +0.65 | +0.6 | +
| Mid femur length | +15.78 | +0.54 | +16.5 | +15.2 | +
| Mid tibia length | +23.38 | +1.14 | +24.9 | +22.2 | +
| Mid tarsus length | +0.52 | +0.02 | +0.55 | +0.5 | +
| Hind femur length | +20.34 | +1.08 | +21.6 | +19.0 | +
| Hind tibia length | +31.12 | +1.67 | +33.5 | +29.5 | +
|
+Hind tarsus length
+ |
+0.6 | +0.0 | +0.6 | +0.6 | +
| Abdomen length | +10.5 | +0.38 | +10.9 | +10.0 | +
| Abdomen maximum width | +2.5 | +0.07 | +2.6 | +2.4 | +
|
+ |
+1945, male | +holotype deposited in MNRJ (courtesy of the team of the | +
|---|---|---|
| digitization project of the Entomological collection of | +MNRJ and | +Luiz A. A. Costa), 84, dorsal view, 85, ventral view, 86, | +
| lateral view, 87, labels. | +
| Characters | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|---|---|---|---|---|---|---|
| Type 1 | +Type 2 | +|||||
|
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|
| Mandibular apex | +simple (as in Fig. 1f) | +simple (Fig. 1f) | +bifid (Fig. 1e; rare specimens with simple apex exist) | +
| Lacinia | +simple, with trichoid setae (as in Fig. 1d) | +simple, with trichoid setae (Fig. 1d) | +bilobate, with stout setae distally (Fig. 1c) | +
| Pronotum shape | +flat | +weakly arched | +highly and evenly arched | +
| Pronotal grooves | +largely reduced (Figs 2e, 4a–c) | +well developed (Figs 2c, 4d–h) | +well developed (Figs 2a, 5) | +
| Lateral margins of pronotum | +very weakly arcuate (Figs 4a–c) | +strongly arcuate, strongly narrowed anteriorly (Figs 4d–h) | +evenly rounded (Fig. 5) | +
| Basal pronotal margin | +narrower than elytral bases combined (Figs 3a–c) | +narrower than elytral bases combined (Figs 3e–g) | +ca. as wide as elytral bases combined (Figs 3h–k) | +
| Strial punctures of elytra | +minute, scarcely impressed (Fig. 2b) | +coarse, well impressed (Fig. 2d) | +minute, scarcely impressed (Fig. 2f) | +
| Interval punctures | +ca. as large as strial punctures | +much smaller than strial punctres | +ca. large as strial punctures | +
| Punctures of scutellary stria | +7–9, often become indis- tinct | +4–7, coarse and always distinct | +6–8, always distinct | +
| Legs including claws | +conspicuously long | +conspicuously long | +less obviously long | +
| Metarsomere 5 | +ca. as long as metatar- someres 3–4 combined (sometimes longer) | +much longer than meta- tarsomeres 3–4 combined (sometimes as long as) | +slightly longer or as long as metatarsomeres 3–4 combined | +
| Metatarsomere / tarsal claw | +1.5 | +1.7 | +1.9 | +
| Apex of parameres | +pointed | +pointed | +blunt | +
| Apical portion of median lobe | +equilateral triangle | +isosceles triangle | +isosceles to almost equilat- eral triangle | +
| Phallobase | +short | +short to very long | +short | +
| Species | +
+Male
+ |
+① | +② | +③ | +④ | +⑤ | +⑥ | +⑦ | +⑧ | +
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+35.1–38.2 (37.1 ± 1.3, n = 8) | +0 | +0 | +0 | +0 | +0 | +0 | +0 | +0 | +
|
+ |
+33.1–38.7 (35.3 ± 2.3, n = 6) | +0 | +1 | +0 | +0 | +0 | +1 | +1 | +1 | +
|
+ |
+37.8–50.4 (n = 28) | +1 | +1 | +1 | +1 | +0 | +? | +0 | +0 | +
|
+ |
+39.7–44.2 (41.3 ± 1.8, n = 5) | +0 | +1 | +0 | +1 | +1 | +1 | +1 | +0 | +
| Sl. No | +Parameter | +Best Cut- | +At Best Cut-off | +RMS (FP, | +|
|---|---|---|---|---|---|
| Off | +FN) | +||||
| Sensitivity | +Specificity | +||||
| 1 | +GA-both leaf and | +0.0500 | +39.15 | +77.80 | +64.80 | +
| bark | +|||||
| 2 | +GA-BARK | +0.0500 | +7.50 | +77.80 | +95.10 | +
| 3 | +GA-LEAF | +0.0500 | +70.80 | +77.80 | +36.70 | +
| 4 | +EPI- both leaf and | +0.0250 | +94.80 | +88.90 | +12.30 | +
| bark | +|||||
| 5 | +EPI-BARK | +0.0250 | +99.10 | +88.90 | +11.20 | +
| 6 | +EPI-LEAF | +0.0250 | +90.60 | +88.90 | +14.60 | +
| 7 | +CAT- both leaf and | +0.0500 | +37.70 | +77.80 | +66.10 | +
| bark | +|||||
| 8 | +CAT-BARK | +0.0500 | +50.90 | +77.80 | +53.90 | +
| 9 | +CAT-LEAF | +0.0500 | +80.20 | +33.30 | +69.50 | +
| Fe | +Pt | +Ti | +Mt | +Ta | +Total | +|
|---|---|---|---|---|---|---|
| I | +3.0/3.3 | +1.25/1.55 | +2.75/3.35 | +3.0/3.0 | +1.6/1.5 | +11.6/12.7 | +
| II | +2.75/3.3 | +1.2/1.4 | +2.35/2.6 | +2.9/2.9 | +1.45/1.5 | +10.65/11.7 | +
| III | +2.6/3.25 | +1.1/1.35 | +2.2/2.5 | +3.15/2.85 | +1.4/1.5 | +10.45/11.45 | +
| IV | +3.35/4.0 | +1.25/1.55 | +3.05/3.5 | +3.25/5.05 | +1.75/2.0 | +12.65/16.1 | +
| Fe | +Pt | +Ti | +Mt | +|
|---|---|---|---|---|
| I | +3d 2(3)p 2r | +2d | +2p 2r 2-2v | +1p 2-2v | +
| II | +3d 2p 2r | +2d | +2p 2r 2-2v | +2p 1r 2-2v | +
| III | +3d 2p 2r | +2d 1p 1r | +2d 2p 2r 2-2v | +2p 2r 2-2v | +
| IV | +3d 2p 1r | +2d 1p 1r | +2d 2p 2r 2-2v | +2p 2r 2-2v | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|
|
+ |
+Transversely bilobed, the lobes cuneate. The apex retuse with a wide sinus. | +Transversely bilobed, the lobes flabellate with thickened external margins. | +Reniform, with the basal angles narrowly rounded, the apex broadly rounded, shallowly incised. | +Subcordate, with the apex broadly rounded, shortly incised. | +
|
+ |
+Absent. | +Oblong, recurved, pubescent. | +Absent. | +A microscopic lobule in the sinus. | +
|
+ |
+Burgundy | +Vermilion | +Orange | +Vermilion | +
|
+ |
+Ovate, oblique, with a second point midway on the inner margin | +Narrowly triangular, rounded. | +Obliquely subquadrate or oblong, obliquely truncate with the inner angles shortly acuminate-obtuse, the outer angles subacute. | +Obliquely triangular, narrowly rounded at the apex. | +
|
+ |
+Triangular, oblique, with the apex attenuate, incurved. Shorter than the upper lobe. | +Triangular, rounded. ½ the length of the upper lobe. | +Obliquely subquadrate or oblong, obliquely truncate with the inner angles shortly acuminate-obtuse, the outer angles subacute. | +Oblong, oblique at the apex. Shorter than the upper lobe. | +
|
+ |
+Orange, strongly suffused with burgundy at the base | +Crimson with saffron margins. The upper lobe apices sometimes crimson. | +Bright yellow with orange at the base. | +Saffron, suffused with crimson near the base. | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +frisch | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+unter-alpin, supra-subalpin und ober-subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+306350 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +sehr hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+sehr warm-kollin (nur an
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| EmTaCe | +Durchschnitt | +Min.-Max. | +Holotypus | +|
|
+ |
+165,2 | +160,0 - 175 | +168,7 | +|
|
+ |
+63,9 | +65,0 - 70,0 | +70,0 | +|
|
+ |
+42,5 | +40,0 - 45.0 | +43.5 | +|
| Breite Prodorsum | +63,3 | +60,0 - 70,0 | +65,0 | +|
| Breite Na | +89,1 | +83,7 - 100,0 | +90,0 | +|
|
+ |
+33,0 | +31,2 - 35.0 | +33.8 | +|
|
+ |
+15,5 | +13,7 - 17,5 | +15.2 | +|
| Keulenbreite | +5,0 | +- | +5,0 | +|
| Abstand ro | +11,1 | +10,0 - 12,0 | +12,0 | +|
| Abstand la | +18,0 | +17,5 - 20,0 | +- | +|
| Abstand ila | +20,3 | +19,2 - 21,2 | +21,2 | +|
| Abstand c1 | +35.6 | +33.2 - 38,2 | +37.5 | +|
| Abstand e1 | +23,0 | +20,7 - 23.7 | +- | +|
|
+ |
+20,4 | +18,7 - 22,5 | +22,5 | +|
|
+ |
+20,1 | +20,0 - 21,2 | +21,2 | +|
|
+ |
+16,7 | +15,0 - 18,0 | +17.5 | +|
|
+ |
+18,4 | +17,0 - 20,0 | +20,0 | +|
|
+ |
+21,3 | +18,7 - 23,7 | +23,7 | +|
| Abstand la: ro | +1,62 | +EmTaCe | +- | +|
| Abstand ila: la | +1,13 | +EmTaCe | +- | +|
|
+ |
+0,54 | +EmTaCe | +0,54 | +|
|
+ |
+2,36 | +EmTaCe | +- | +|
|
+ |
+1,25 | +EmTaCe | +1,16 | +|
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +sauer (pH 3.5-6.5) | +Temperaturzahl T | +alpin und nival (von der Baumgrenze bis zur Schneegrenze) | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| 4.6 - Grasbrachen | +
|
+4.6.1 - Queckenbrache (
+ |
+
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +unter-montan und ober-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+68300 | +
| = | +
+ |
+
+ |
+629 | +
| = | +
+ |
+
+ |
+859 | +
| = | +
+ |
+
+ |
+859 | +
| = | +
+ |
+
+ |
+68300 | +
| = | +
+ |
+
+ |
+1327 | +
| = | +
+ |
+
+ |
+1130 | +
| = | +
+ |
+
+ |
+68300 | +
| = | +
+ |
+
+ |
+473 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+22900 | +
| = | +
+ |
+
+ |
+22900 | +
| = | +
+ |
+
+ |
+670 | +
| = | +
+ |
+
+ |
+22900 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+nass; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +2 - Schwerpunktlebensraum | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+378600 | +
| = | +
+ |
+
+ |
+2490 | +
| = | +
+ |
+
+ |
+2666 | +
| = | +
+ |
+
+ |
+2666 | +
| = | +
+ |
+
+ |
+378600 | +
| = | +
+ |
+
+ |
+444 | +
| = | +
+ |
+
+ |
+388 | +
| = | +
+ |
+
+ |
+378600 | +
| = | +
+ |
+
+ |
+2391 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +verschollen, vermutlich in der Schweiz ausgestorben (Critically Endangered, Probably Extinct) | +|
| Alpennordflanke (NA) | +verschollen, vermutlich in der Schweiz ausgestorben (Critically Endangered, Probably Extinct) | +|
|
+ |
+vom Aussterben bedroht (Critically Endangered) | +C1 | +
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +-- | +
|
+ |
+
+2 - Hohe nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+1 - Gering | +
|
+ |
+
+2 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(23.01.2013) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(25.07.2007) | +
| - | +Holotype | +Paratypes (n = 12) | +
|---|---|---|
|
+ |
+67.6 | +44.3-71.6 | +
|
+ |
+||
| Body depth | +14.0 | +13.9-18.7 | +
| Caudal peduncle depth | +12.0 | +11.3-13.0 | +
| Body width | +10.2 | +10.2-14.1 | +
| Caudal peduncle width | +3.1 | +2.9-4.7 | +
| Pre-dorsal length | +60.2 | +61.5-66.6 | +
| Pre-pelvic length | +54.5 | +54.9-61.6 | +
| Dorsal-fin base length | +11.7 | +10.8-13.8 | +
| Anal-fin base length | +7.7 | +7.7-9.1 | +
| Caudal-fin length | +17.3 | +14.4-18.5 | +
| Pectoral-fin length | +14.9 | +12.1-15.1 | +
| Pelvic-fin length | +10.3 | +8.7-11.2 | +
| Head length | +20.4 | +19.7-22.8 | +
|
+ |
+||
| Head depth | +45.4 | +46.6-54.4 | +
| Head width | +81.3 | +79.0-89.9 | +
| Snout length | +40.1 | +41.2-45.5 | +
| Interorbital length | +20.7 | +20.5-25.9 | +
| Preorbital length | +15.3 | +13.6-16.7 | +
| Eye diameter | +12.8 | +10.8-13.8 | +
| Morphological feature | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|---|---|
| Source of Information | +
+Holotype: SWIMS-ANN- 18-012; Paratypes: SWIMS-ANN- 18-013 - (
+ |
+
+Holotype: NTM W 29624; Paratypes: NTM W 29619 – NTM 29623 (
+ |
+
+Holotype: ASIZW 0000345-1 (
+ |
+
+Type: USNM. No. 20114 (
+ |
+
+Holotype: ASIZW 0000348-2 (
+ |
+
+Holotype:
+ |
+Holotype: UMTAnn 2149 (this study) | +Holotype: UMTAnn 2177 (this study) | +
|
+Size (mm):
+ |
+3.3–7.0, 2.2–5.3 | +
+ |
+13.9, 5.7 | +
+ |
+8.7, 3.9 | +12.7, 5.95 | +5.25 (3.45–5.85), 2.85 (1.95–3.15) | +2.7 (2.85–4.8), 1.8 (1.65–2.55) | +
| Prostomium: shape | +Bilobed | +Bilobed | +Bilobed | +Bilobed | +Bilobed | +Bilobed | +Bilobed | +Bilobed | +
| Palps: reaching | +Chaetiger 1 | +Chaetiger 1 | +
+ |
+
+ |
+Chaetiger 1 | +
+ |
+
+ |
+Chaetiger 3 | +
| Lateral antennae: reaching | +Chaetiger 1 | +Chaetiger 1 or 2 | +
+ |
+
+ |
+Chaetiger 2 | +
+ |
+Chaetiger 2 | +Chaetiger 4 | +
| Median antennae: reaching | +Chaetiger 1 or 2 | +Chaetiger 1 or 2 | +
+ |
+
+ |
+Chaetiger 3 | +Chaetiger 1 | +Chaetiger 3 | +Chaetiger 5 | +
| Peduncle in prostomial appendages | +Absent | +Absent | +Absent | +Absent | +Absent | +Absent | +Absent | +Absent | +
| Eyes * | +Absent | +Present | +Present | +Absent | +Absent | +Absent | +Present | +Present | +
|
+ |
+5-6 + 5 – 6, 7 + 0, 4 + 8 | +4 + 5, 4–5 + 0, 3–4 + 5 – 6 | +3 + 3, 4 + 0, 4 + 5 | +3 + 3, 4 + 0, 4 + 3 | +4 + 4, 5 + 0, 4 + 8 | +5 + 5, 5 + 0, 4 + 8 | +5 (4–5) + 5 (5–6), 7 (6–7) + 0, 4 (4–5) + 8 (5–8) | +5 (4–5) + 5 (4–6), 5 (4–6) + 0, 3 (3–4) + 6 (7–8) | +
| Branchiae: shape | +Pectinate | +Pectinate | +Palmate | +
+ |
+Pectinate | +Pectinate | +Pectinate | +Pectinate | +
| Branchiae: start chaetiger; last chaetiger before pygidium | +15–35; until pygidium | +16–20; until pygidium | +32; end at chaetiger 281 | +45; end ~ 30 last chaetiger | +26; end at chaetiger 181 | +15; end at chaetiger 399 | +24 (16–27); end ~ 10 last chaetiger | +20 (15–25); until pygidium | +
| Branchial filaments: numbers | +5–10 | +6–7 | +5 | +3 | +3 | +8 | +6 | +5 | +
| Dorsal cirri: shaped | +Conical | +Conical | +
+ |
+Conical | +
+ |
+
+ |
+Digitiform | +Thumb-shape, digitiform | +
| Prechaetal lobe: shaped | +Transverse fold | +Transverse fold | +
+ |
+
+ |
+
+ |
+
+ |
+Transverse fold | +Transverse fold | +
| Chaetal lobe: shaped | +Rounded | +Rounded | +
+ |
+Rounded | +
+ |
+
+ |
+Rounded and conical | +Rounded | +
| Aciculae: shape; colour | +
+ |
+Blunt; black with paler tips | +
+ |
+Blunt; black | +
+ |
+
+ |
+Blunt; black and translucent at distal end | +Blunt; black and translucent at distal end | +
|
+Subacicular limbate chaetae: (
+ |
+Absent; all chaetigers | +Absent; all chaetigers | +Absent; all chaetigers | +Absent; all chaetigers | +Absent; all chaetigers | +Absent; all chaetigers | +Absent; all chaetigers | +Absent; all chaetigers | +
| Pectinate chaetae: number of type * | +4 | +3 | +4 | +
+ |
+3 | +3 | +5 | +4 | +
| Subacicular hook: shape; colour * | +Unidentate and bidentate; amber | +Unidentate; amber to black | +Unidentate and bidentate; yellow | +
+Unidentate;
+ |
+Unidentate and bidentate; brown | +Unidentate | +Unidentate; light brown and translucent at distal end | +Unidentate and bidentate; light brown and translucent at distal end | +
| Subacicular hook: start chaetiger * | +26–58 | +30–38 | +20 | +Present in posterior region (No information on start chaetiger) | +20 | +170 | +37 (26–42) | +25 (21–38) | +
| Subacicular hook: distribution | +Continuous | +Continuous | +Continuous | +
+ |
+Continuous | +Continuous | +Continuous | +Continuous | +
| Species | +Jacana Eco Estate | +Queen Elizabeth Park | +
|---|---|---|
|
+ |
+1-15, 42-52 (25 weeks) | +1-18, 41-52 (29 weeks) | +
|
+ |
+1-6, 45-52 (14 weeks) | +1-16, 52 (17 weeks) | +
|
+ |
+5-20 (16 weeks) | +1-20, 48-52 (25 weeks) | +
|
+ |
+11 (1 week) | +18 (1 week) | +
|
+ |
+21-22 (2 weeks) | +20-36 (17 weeks) | +
|
+ |
+38 (1 week) | +10-16 (7 weeks) | +
|
+ |
+45-52 (8 weeks) | +45-49 (5 weeks) | +
|
+ |
+51 (1 week) | +13 (1 week) | +
|
+ |
+51 (1 week) | +3-5, 48-52 (8 weeks) | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +trocken | +Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +basisch (pH 6.5->8.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +0 - unbedeutend, keine Bindung. | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+178300 | +
| = | +
+ |
+
+ |
+1940 | +
| = | +
+ |
+
+ |
+1444 | +
| = | +
+ |
+
+ |
+1444 | +
| = | +
+ |
+
+ |
+178300 | +
| = | +
+ |
+
+ |
+2813 | +
| = | +
+ |
+
+ |
+2282 | +
| = | +
+ |
+
+ |
+178300 | +
| = | +
+ |
+
+ |
+1616 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+stark
+ |
+C2a(i) | +
| Mittelland (MP) | +
+stark
+ |
+C2a(i) | +
| Alpennordflanke (NA) | +
+stark
+ |
+C2a(i) | +
|
+ |
+verschollen, vermutlich in der Schweiz ausgestorben (Critically Endangered, Probably Extinct) | +|
|
+ |
+
+stark
+ |
+B2ab(iii) | +
| Westliche Zentralalpen (WA) | +verletzlich (Vulnerable) | +A2c; A4c; C1 | +
|
+ |
+
+4 -
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+1 - Gering | +
|
+ |
+
+1 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(25.07.2007) | +
|
+ |
+
+ |
+(29.11.2005) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(02.03.2005) | +
|
+ |
+Z - Zielart | +weitere Informationen | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +unter-montan und ober-kollin | +||
|
+ |
+-- | +
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| 1 | +Forewing: hairs absent in 3rd submarginal cell and 2nd medial cell, usually also in 2nd submarginal cell and 1st medial cell; S4 in males with a rounded, triangular or elongated brush of dense usually forward-directed bristles, S6 of females medially with raised area which varies among species from inconspicuously roughened, broadly parabolic with well-defined rim and slender spine, to almost a pygidium like structure | +
+ |
+
| - | +Forewing: hairs present in in 3rd submarginal cell, 2nd medial cell and most other cells; S4 in males and S6 in females not modified as described above | +
+ |
+
| 2 | +Female: Integument of paraclypeal areas black; fore and mid femora and tibiae with iridescent blue-green hairs, male: S6 gently convex; apex S7 broadly triangular | +
+ |
+
| - | +Female: Integument of paraclypeal areas partly yellow, white or ivory; hair on fore and mid legs never iridescent, male: S6 with broad depressions either side of midline; apex S7 ovate | +
+ |
+
| 1 | +
+Female: Body size large (> 19 mm), pubescence on head and mesosoma white, metasomal integument predominantly dark red (Fig.
+ |
+
+ |
+
| - | +Not with above combination of characters (20 spp.) | +
+ |
+
| 2 | +Metasomal pubescence predominantly black, some species with light coloured posterior bands of white pubescence on terminal terga (5 spp.) | +
+ |
+
| - | +Metasomal pubescence predominantly orange, brown or grey, with or without an anterior band of black pubescence on T2 (15 spp.) | +
+ |
+
| 3 | +
+Light coloured posterior bands present on all terga (Figs
+ |
+
+ |
+
| - | +
+Light coloured posterior bands lacking, or only on last 1-3 terminal terga (Figs
+ |
+
+ |
+
| 4 | +
+Female: T1 with thin, faint white posterior band, T2-4 with well-developed white posterior hair bands (Fig.
+ |
+
+ |
+
| - | +All terga with well-developed posterior hair bands. Hair bands on T1-2 orange (2 spp.) | +
+ |
+
| 5 | +
+Female: clypeus ivory with two large black marks, which are sometimes combined at the top, scape black. Male: S4 with two adjacent small patches of downward-directed bristles on apicomedial margin and two large comma-shaped patches of bristles on S5 (Fig.
+ |
+
+ |
+
| - | +
+Female: clypeus with two small brown marks, scape with pale yellow mark (Fig.
+ |
+
+ |
+
| 6 | +
+Metasomal pubescence entirely black (Fig.
+ |
+
+ |
+
| - | +
+T4-5 (female), T5-6 (male) with predominantly white hairs (Fig.
+ |
+
+ |
+
| 7 | +Male | +
+ |
+
| - | +Female | +
+ |
+
| 8 | +
+Both S3 and S4 medially with round patch of bristles, the patch on S3 smaller than on S4 (Figs
+ |
+
+ |
+
| - | +
+Only S4 with median patch of bristles of varying size and shape (Figs
+ |
+
+ |
+
| 9 | +
+S2-4 without apicomedial emarginations; S5 apicomedial area deeply emarginated, emargination deeper than wide and almost parallel to body axis (Figs
+ |
+
+ |
+
| - | +
+S2-4 with distinct apicomedial emarginations (Fig.
+ |
+
+ |
+
| 10 | +
+Apicomedial emargination on S5 parallel-sided, slightly deeper than wide (Fig.
+ |
+
+ |
+
| - | +
+Apicomedial emargination on S5 more than twice as deep as wide (Fig.
+ |
+
+ |
+
| 11 | +
+S4 shape of patch of bristles round (Figs
+ |
+
+ |
+
| - | +
+S4 shape of patch of bristles triangular or bell-shaped (Figs
+ |
+
+ |
+
| - | +
+S4 shape of patch of bristles elongated (Figs
+ |
+
+ |
+
| 12 | +
+S4 patch of bristles relatively large, circa 1/7 of width of sternum (Figs
+ |
+
+ |
+
| - | +
+S4 patch of bristles small, less than 1/10 of width of sternum (Fig.
+ |
+
+ |
+
| 13 | +
+Face marks pale yellow, scape brown (Fig.
+ |
+
+ |
+
| - | +
+Face marks ivory, underside of scape with ivory mark (Fig.
+ |
+
+ |
+
| 14 | +
+S4 patch of bristles small, circa 1/5 of width of sternum (Fig.
+ |
+
+ |
+
| - | +
+S4 patch of bristles larger, S5 apicomedial emargination shallow (Figs
+ |
+
+ |
+
| 15 | +
+S4 patch of bristles large, circa 1/3 of width of sternum, bristles brown (Fig.
+ |
+
+ |
+
| - | +S4 patch of bristles less than 1/3 of width of sternum, bristles black (2 spp.) | +
+ |
+
| 16 | +
+S4 patch of bristles triangular (Fig.
+ |
+
+ |
+
| - | +
+S4 patch of bristles bell-shaped (Fig.
+ |
+
+ |
+
| 17 | +
+S4 patch of black bristles broad and circa 1/2-2/3 of sternum width (Fig.
+ |
+
+ |
+
| - | +S4 patch of pale bristles narrow, circa 1/3 of width of sternum, S6 apicomedial area without conspicuous fimbria of black and orange-fringed hairs, T2 anteriorly with area of black hairs much narrower than marginal zone, or without black hairs (4 spp.) | +
+ |
+
| 18 | +
+Face marks ivory/white, elongated patch of bristles on S4 black (Fig.
+ |
+
+ |
+
| - | +Face marks pale to bright yellow, patch of bristles on S4 dark or light brown (3 spp.) | +
+ |
+
| 19 | +
+T2 anteriorly without dark hairs, S5 apicomedial emargination circa 6
+ |
+
+ |
+
| - | +
+T2 anteriorly with entire narrow band of dark hairs, S5 apicomedial emargination less than 4
+ |
+
+ |
+
| 20 | +
+S6 marginal zone smooth without punctures or hairs, some short hairs medially (Fig.
+ |
+
+ |
+
| - | +
+S6 marginal area, smooth with open punctures and hairs, medially with patch of stiff dark hairs (Fig.
+ |
+
+ |
+
| 21 | +Hind leg: tibial scopa on outer surface with white or grey-white hairs (5 spp.) | +
+ |
+
| - | +Hind leg: tibial scopa on outer surface with pale yellow, ochre or orange hairs (8 spp.) | +
+ |
+
| 22 | +Paraclypeal area with pale yellow mark (3 spp.) | +
+ |
+
| - | +Paraclypeal area black (3 spp.) | +
+ |
+
| 23 | +
+Paraclypeal marks small (Fig.
+ |
+
+ |
+
| - | +
+Paraclypeal marks large (Figs
+ |
+
+ |
+
| 24 | +Scutum and T1 pubescence light brown, T2 and T3 anteriorly with black hairs | +
+ |
+
| - | +Scutum and T1 pubescence appears grey, hairs light ochre intermixed with longer black hairs, only T2 with black hairs anteriorly | +
+ |
+
| 25 | +
+Supraclypeal area black, clypeus with yellow inverted T shaped mark (Fig.
+ |
+
+ |
+
| - | +
+Supraclypeal area with yellow or pale yellow mark, clypeus with two vertical brown marks (Fig.
+ |
+
+ |
+
| 26 | +Paraclypeal area with light coloured mark (3 spp.) | +
+ |
+
| - | +Paraclypeal area without light coloured mark (6 spp.) | +
+ |
+
| 27 | +T2 anteriorly with narrow band of black hairs across whole width, sterna with dark hairs | +
+ |
+
| - | +Dark hairs on T2 restricted to antero-lateral corners or lacking, sterna on disk with orange hairs (2 spp.) | +
+ |
+
| 28 | +
+Integument of T3 and following segments anteriorly dark brown, clypeus with two well defined vertical dark orange marks (Fig.
+ |
+
+ |
+
| - | +
+Integument of T3 and following segments anteriorly orange, marks on clypeus undefined and small (Fig.
+ |
+
+ |
+
| 29 | +T2 anteriorly with black hairs (2 spp.) | +
+ |
+
| - | +Pubescence on T2 uniformly coloured (4 spp.) | +
+ |
+
| 30 | +
+Scape orange, T2 anterior with wide area of black hairs, clypeus marks undefined (Fig.
+ |
+
+ |
+
| - | +
+Scape black, apically brown, T2 anteriorly with narrow area of erect black hairs, clypeus with brown horseshoe-like mark (Fig.
+ |
+
+ |
+
| 31 | +
+Clypeus colouration yellow, orange pubescence on terga with iridescent shine, S6 projection broadly parabolic, well defined (Figs
+ |
+
+ |
+
| - | +
+Clypeus colouration pale yellow or ivory, pubescence on terga without iridescent shine, S6 projection rounded, not well defined (Figs
+ |
+
+ |
+
| 32 | +
+Clypeus marked with inverted yellow T (Fig.
+ |
+
+ |
+
| - | +
+Clypeus yellow, with two faint brown patches (Fig.
+ |
+
+ |
+
| 33 | +
+Clypeus with pale yellow inverted T-shape, supraclypeal area black (Fig.
+ |
+
+ |
+
| - | +
+Clypeus colouration ivory, supraclypeal area black with narrow ivory line at base (Fig.
+ |
+
+ |
+
| Leg | +Tr | +Fe | +Ge | +Ti | +Ta | +
|---|---|---|---|---|---|
| v | +dlbv | +lv | +lv12 | +fttcitpuaspvvpll12 | +|
| v | +dlbv | +lv | +lv | +fttcitpuaspv12 | +|
| v | +dev | +l | +lv | +fttcitpuaspv | +|
| v | +dev | +dl | +lv | +fttcpuaspv | +
|
+1.
+ |
+
+ |
+
|
+1*.
+ |
+|
|
+2.
+ |
+|
|
+3.
+ |
+
+ |
+
|
+3*.
+ |
+
+ |
+
|
+2*.
+ |
+
+ |
+
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+136760 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+62600 | +
| = | +
+ |
+
+ |
+292 | +
| = | +
+ |
+
+ |
+1122 | +
| = | +
+ |
+
+ |
+1122 | +
| = | +
+ |
+
+ |
+62600 | +
| = | +
+ |
+
+ |
+62600 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+frisch; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +unter-montan und ober-kollin | +||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| 4.0 - Kunstrasen | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +1 | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+427400 | +
| = | +
+ |
+
+ |
+1120 | +
| = | +
+ |
+
+ |
+632 | +
| = | +
+ |
+
+ |
+632 | +
| = | +
+ |
+
+ |
+427400 | +
| = | +
+ |
+
+ |
+1715 | +
| = | +
+ |
+
+ |
+1429 | +
| = | +
+ |
+
+ |
+427400 | +
| = | +
+ |
+
+ |
+884 | +