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<treatment LSID="urn:lsid:plazi:treatment:01B57E23612355FA935C5619AF1F8B6E" httpUri="http://treatment.plazi.org/id/01B57E23612355FA935C5619AF1F8B6E" lastPageNumber="71" pageId="0" pageNumber="71">
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<treatment id="01B57E23612355FA935C5619AF1F8B6E" ID-GBIF-Taxon="195868763" LSID="urn:lsid:plazi:treatment:01B57E23612355FA935C5619AF1F8B6E" httpUri="http://treatment.plazi.org/id/01B57E23612355FA935C5619AF1F8B6E" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
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8.
<taxonomicName LSID="01B57E23-6123-55FA-935C-5619AF1F8B6E" authority="Morvan, 1970" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii" subGenus="Eremoderus">Platyderus (Eremoderus) davatchii Morvan, 1970</taxonomicName>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">Fig. 17</figureCitation>
<subSubSection id="2CE182247DB1B51F121437CD5DE31F9D" pageId="0" pageNumber="71" type="description">
<paragraph id="6654F85C677B24956271EE5FA72F6FA8" pageId="0" pageNumber="71">
<figureCitation id="FE92039AD7FC73E10EB62B95F30E1BB4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">Fig. 17</figureCitation>
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<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">Platyderus davatchii</taxonomicName>
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<paragraph id="21936B6849A2364EA64651A58DEB1F8B" pageId="0" pageNumber="71">
<taxonomicName id="82F68172A7EBE919DDE87AFA00F7A84C" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">Platyderus davatchii</taxonomicName>
Morvan, 1970: 194 (type locality:
<normalizedToken originalValue="“l’Elburz">&quot;l'Elburz</normalizedToken>
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, environs
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; alt. 2 700 m...
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torrent, versan nord&quot;).
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<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="794246512D8256BA1B2F4102EEB30685" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="108D74F84A7F95A887A1643104FC685B" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="7107595AE5045320737F90FE06C65F1E" pageId="0" pageNumber="71">
The type locality in the Iranian province of Qazvin, in Qazvin County, in the vicinities of Alamut Castle, 2700 m, may conditionally confine to the high area of the Central Alborz Range situated northward of the upper current of
<normalizedToken originalValue="Shāh">Shah</normalizedToken>
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River, eastward of the Ovan Lake and westward of the upper currents of Seh Hazar River. It may concern the area situated north-northeast of ruins of the Alamut Castle.
</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="71">Figure 9.</emphasis>
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<emphasis id="226A9C62C714C558B528DF65590C79B3" bold="true" pageId="0" pageNumber="71">Figure 9.</emphasis>
Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe).
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authority="(Platyderus)" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">Platyderus (Platyderus)</taxonomicName>
<emphasis id="58137B07B463A4C4B0086854E5D26B37" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="5DB26B8C61801AB8F4C661B8625928D6" authority="(Platyderus)" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">Platyderus (Platyderus)</taxonomicName>
sp., male specimen, Boz
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Mtn., Turkey;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">P. (Eremoderus) afghanistanicus</taxonomicName>
<emphasis id="0F1F8EF127A27968D274C1EC14D3133E" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="72FF28987E3B5E34911995D329C72135" class="Insecta" kingdom="Animalia" lsidName="P. (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">P. (Eremoderus) afghanistanicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
<emphasis id="4758753657C832EB815B40197289404E" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="4552734E56453699FEA7A35A2F12F616" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="546CD75C3289EAC21A9D8B7AA8B9E06F" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="B62DCDDC6C80FE79D5870F33AFA49AEB" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName authority="insignitus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="E." subSpecies="insignitus">P. (E.) insignitus insignitus</taxonomicName>
<emphasis id="3B18AFEC9BF2517B2280CEF06331BABC" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="2F174CC3F4CD3BC85A327275B5647108" authority="insignitus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="E." subSpecies="insignitus">P. (E.) insignitus insignitus</taxonomicName>
Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
<emphasis id="02285E524AF611BBE56688795CE0478C" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="8E309FF6332C3540189A0364DD90BC40" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="A51963E2A8086B053DC8FF641B3E967D" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="BFE5E2CDC72911CCB3E9E1B1490832E6" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="0D16DA2D7E1AD84F9D707EF103BB2577" bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName id="849348536E5175920B582C95DB325E72" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), male specimen, Naẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<subSubSection id="78EECD68FEB5553B584217F6F3796B2A" pageId="0" pageNumber="71" type="references">
<paragraph id="5A8C781D325E59ECFB6DB5AC0DCED29A" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="987BEC82438727D9943E72E6AB5ED848" pageId="0" pageNumber="71">
<taxonomicName id="F75D14C2F192821423642215D40C88FE" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="9E3C68F3AC424D4C778A8DED4F41237B" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1969.21081" author="Morvan, P" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="192 - 198" refId="B40" refString="Morvan, P, 1970. Contribution a la connaissance des coleopteres carabiques de l'Iran. Bulletin de la Societe Entomologique de France 74 (1969): 192 - 198, DOI: https://doi.org/10.3406/bsef.1969.21081" title="Contribution a la connaissance des coleopteres carabiques de l'Iran." url="https://doi.org/10.3406/bsef.1969.21081" volume="74" year="1970">Morvan 1970</bibRefCitation>
<bibRefCitation id="BF571460D27BE15577171C4833EEF4F3" DOI="https://doi.org/10.3406/bsef.1969.21081" author="Morvan, P" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="192 - 198" refId="B40" refString="Morvan, P, 1970. Contribution a la connaissance des coleopteres carabiques de l'Iran. Bulletin de la Societe Entomologique de France 74 (1969): 192 - 198, DOI: https://doi.org/10.3406/bsef.1969.21081" title="Contribution a la connaissance des coleopteres carabiques de l'Iran." url="https://doi.org/10.3406/bsef.1969.21081" volume="74" year="1970">Morvan 1970</bibRefCitation>
:
<bibRefCitation author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan 1974</bibRefCitation>
<bibRefCitation id="6CB9BE7C40840D8CBAE832159FC8E843" author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan 1974</bibRefCitation>
: 149;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="BCBF26E8A58BE17D8845DC41E42144B3" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="A578AB143B8148BF20A5EA06AFAC2832" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 522;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="C1A9001EB05BC3D09651658AC19BC1DB" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 395;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4024.1.1" author="Azadbakhsh, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="1 - 108" refId="B9" refString="Azadbakhsh, S, Nozari, J, 2015. Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae). Zootaxa 4024 (1): 1 - 108, DOI: https://doi.org/10.11646/zootaxa.4024.1.1" title="Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae)." url="https://doi.org/10.11646/zootaxa.4024.1.1" volume="4024" year="2015">Azadbakhsh and Nozari 2015</bibRefCitation>
<bibRefCitation id="A13E74AAC7645CFCFA4908E3F0F06C1C" DOI="https://doi.org/10.11646/zootaxa.4024.1.1" author="Azadbakhsh, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="1 - 108" refId="B9" refString="Azadbakhsh, S, Nozari, J, 2015. Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae). Zootaxa 4024 (1): 1 - 108, DOI: https://doi.org/10.11646/zootaxa.4024.1.1" title="Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae)." url="https://doi.org/10.11646/zootaxa.4024.1.1" volume="4024" year="2015">Azadbakhsh and Nozari 2015</bibRefCitation>
: 84;
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="EE0080C6329D57403125564AEA210EE3" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 757.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="6061EE122773F9C52C5ECEBAD0EB926B" pageId="0" pageNumber="71" type="type material">
<paragraph id="7D4436A49166E89FD6871A9B078FDD94" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="7B04554CCC4F11DF7A67184597D694DE" pageId="0" pageNumber="71">
<emphasis id="793B26367F6C5432CDF2AC6E1D8C916D" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♀ preserved in private collection Pierre Morvan (Carentoir, France). Not examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">None.</paragraph>
<subSubSection id="B838BA74ECF3D6DFF54A55B6662A77E1" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="857E3D3D54AD3B1CD6A44AF1F686436F" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="33EA63911476D0055B5F8DCFE285D783" pageId="0" pageNumber="71">None.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">The holotype was collected under a large stone near a stream, on a north slope at ca. 2700 m altitude.</paragraph>
<subSubSection id="3DA0B3EC8564ED15CA5A68D7E24F076F" pageId="0" pageNumber="71" type="habitat">
<paragraph id="05D3ADC838F8E4F4C5464B37844C570B" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="BDEF742AC7414F4386FC3D9B796ED5AD" pageId="0" pageNumber="71">The holotype was collected under a large stone near a stream, on a north slope at ca. 2700 m altitude.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="963ED921EB1A370D7B89627A15447D4B" pageId="0" pageNumber="71" type="distribution">
<paragraph id="E4DA975B36AA4B528E5DB02D66BD494C" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="CAD83B48E9D19D83FC21CB7041BE4AFF" pageId="0" pageNumber="71">
Only known from type location in North Iran: Central Alborz Range (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="24E439805B306FD2CF21690AAD091BCB" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="41D8F459F3E413C26E52E5B96B002F85" pageId="0" pageNumber="71" type="notes">
<paragraph id="EEF2FEE4F541DF4AD05FD3BDBF53B389" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="E6B9BFBC1F0CCD78F54E157B26A1D976" pageId="0" pageNumber="71">
Based on the habitus drawing and features noted in the original description (
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1969.21081" author="Morvan, P" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="192 - 198" refId="B40" refString="Morvan, P, 1970. Contribution a la connaissance des coleopteres carabiques de l'Iran. Bulletin de la Societe Entomologique de France 74 (1969): 192 - 198, DOI: https://doi.org/10.3406/bsef.1969.21081" title="Contribution a la connaissance des coleopteres carabiques de l'Iran." url="https://doi.org/10.3406/bsef.1969.21081" volume="74" year="1970">Morvan 1970</bibRefCitation>
<bibRefCitation id="94B4B25FD40875764294BD26708AA6B6" DOI="https://doi.org/10.3406/bsef.1969.21081" author="Morvan, P" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="192 - 198" refId="B40" refString="Morvan, P, 1970. Contribution a la connaissance des coleopteres carabiques de l'Iran. Bulletin de la Societe Entomologique de France 74 (1969): 192 - 198, DOI: https://doi.org/10.3406/bsef.1969.21081" title="Contribution a la connaissance des coleopteres carabiques de l'Iran." url="https://doi.org/10.3406/bsef.1969.21081" volume="74" year="1970">Morvan 1970</bibRefCitation>
: 194, fig. 6),
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
<taxonomicName id="D01D250B2883F3A9F2E447797FB113A3" class="Insecta" kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="6D89EA0089D7A1453EB851698FDAA429" italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
</taxonomicName>
is without doubt an
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="516D3CD326A52D8E5DC48AF0488BAC44" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B606B16A52B5C6E1A5329A4681C88C0B" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
-species. For the time being, it is the westernmost and the least known representative of the &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="D3A480A50C4CFC8B1F083D41A87BFD7C" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="22EF0A65C40CEF26B40EAC2ABC9AD790" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; species group. Study of the holotype or topotypical male specimens of
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<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
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<emphasis id="872A5BD052CE14C7330C1C39FF1C0268" italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
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is a key issue that can help to decide whether its closest neighbor,
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<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
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, is a separate species or its synonym. The distance between the population from Alamut Region (Qazvin County), referred to the former species, and the population south of
<normalizedToken originalValue="Kelārdascht">Kelardascht</normalizedToken>
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[= Hasankif; = Rudbarak] (Chalus County:
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District), referred to the latter species is about 40-50 km in straight line.
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The record about
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<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
<taxonomicName id="9A6F36AC9A25E6C6E221D80AB6178BE0" class="Insecta" kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="F14F32EA731962272424D31D3A805809" italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
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from &quot;10 km S Hasan Keif&quot; (
<bibRefCitation author="Lohaj, R" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" pagination="7 - 14" refId="B32" refString="Lohaj, R, Mlejnek, R, 2007. Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria. Acta Societatis Zoologicae Bohemicae 71: 7 - 14" title="Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria." volume="71" year="2007">Lohaj and Mlejnek 2007</bibRefCitation>
<bibRefCitation id="AFC6577C23870AC8F9E32E452114C8E2" author="Lohaj, R" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" pagination="7 - 14" refId="B32" refString="Lohaj, R, Mlejnek, R, 2007. Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria. Acta Societatis Zoologicae Bohemicae 71: 7 - 14" title="Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria." volume="71" year="2007">Lohaj and Mlejnek 2007</bibRefCitation>
) actually concerns
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<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="11BFCB3C180E0B7AFF63951AB6050519" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="2717B436C965BFB4B94FB83D837209F8" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
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.
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204
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@ -1,170 +1,170 @@
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16.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9E</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11D</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15C</figureCitation>
<subSubSection id="029EA41C6A9A346743D7D33B1C7F3552" pageId="0" pageNumber="71" type="description">
<paragraph id="B97C0624EA5786CF9655F1C1465B3621" pageId="0" pageNumber="71">
<figureCitation id="0BECF6297ABD438A0DBEB9185F5C3FA3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4A</figureCitation>
<figureCitation id="0ECA1C17DA80CDA05FD5088CD975074E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9E</figureCitation>
<figureCitation id="5054B787B5F39590B4E9D89B02A04D3C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11D</figureCitation>
<figureCitation id="F4EA4BA4FB35CC1191FA407D6FE369D2" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="E4CC0A989BA876F21401CBE50F7AAFC8" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="A3B0EE932819DF3985DDE08084946849" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="440791AAC02C398EB8D248E495B011FF" pageId="0" pageNumber="71">
This species differs from
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
<taxonomicName id="E701DB55A2E78051218E24F17BA0472C" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis id="3BEDABE74C95EABD0B1E919653C75EEB" italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
in pronotum less constricted to base (PW/PB: 1.12-1.17, vs. PW/PB: 1.21-1.28) and with apex appreciably narrower than base (PA/PB: 0.80-0.85, vs. PA/PB: 0.88-0.92). Median lobe of aedeagus somewhat longer than that in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="5C08BA12A9EF918418C90196646ED088" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="45072C1E06EB741BEA0D078D38ACD82A" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
(s.l.), with ventral sclerite longer and not forming a distal kink (Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
<figureCitation id="E7B656442016EC02C0EC3DEB961AD6C1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11D</figureCitation>
<figureCitation id="8CD7A82F124CB2C1E19B66017BD03F25" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="redescription">
<paragraph pageId="0" pageNumber="71">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="B5B421415E5BA4E3D89D65B45EDBAD9A" pageId="0" pageNumber="71" type="redescription">
<paragraph id="B93102F948F2592A8A538FD6AC7590AB" pageId="0" pageNumber="71">Redescription.</paragraph>
<paragraph id="0CBD5E99ECC57B3565755C3158AD8EFE" pageId="0" pageNumber="71">
<emphasis id="803B44674F8F3DB32DAC754743CB101F" bold="true" pageId="0" pageNumber="71">
<emphasis id="EB3AE6D528FC1107984852A05AFD9455" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Moderately large species (BL: 6.60-8.70 mm; BW: 2.20-3.00 mm), with elongate, narrow and moderately convex body (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4A</figureCitation>
<figureCitation id="25821D0586FF2426FAC9D0538E6ABB62" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4A</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="AA18EC65A25D6BA6E3EFF48C12183455" bold="true" pageId="0" pageNumber="71">
<emphasis id="31554544057CA94AF9929C25E25532CD" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
<tableCitation id="E3A7EAA29EBB3485DE7A17A32F807053" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="841091A276DC12BD34ECDA2A286832A1" bold="true" pageId="0" pageNumber="71">
<emphasis id="1690A1588B5144490BB23C3F95AD4A09" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Chestnut-colored as palpi, with antennae and legs usually lighter than rest of body, head somewhat darker than pronotum and elytra. Integument shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="13FA9FDABB74097324C5FC5EDBB77D52" bold="true" pageId="0" pageNumber="71">
<emphasis id="31E345D5CEE848FA8DEE97C3445062B1" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum and elytra with isodiametric sculpticells. Ventral surface with impressed slight stretched isodiametric sculpticells or with microsculpture less apparent or scarcely-visible. Head impunctate, usually smooth, but lateral areas adjacent to frontal furrows sometimes with fine wrinkles. Surface of pronotum impunctate, largely smooth, basal area medially behind posterior transverse impression longitudinally wrinkled, rarely apical part medially in front of anterior transverse impression also wrinkled; basal foveae and adjacent lateral areas smooth. Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled in whole, 2-5 finely wrinkled at sides.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="22BC660D5AC12FD9F4F6CDEDE3E637BE" bold="true" pageId="0" pageNumber="71">
<emphasis id="7730527BB2309856FA42A1B509FEE48D" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About two thirds as wide as pronotum (PW/HW= 1.35-1.45). Eye long, convex. Labrum slightly shorter than clypeus, subrectangular, with anterior margin concave medially. Frontoclypeal suture more or less distinct in middle, indistinct or disappeared at sides. Frontal furrows very small, shallow, punctiform. Paraorbital sulci fine, backward not reaching level of posterior supraorbital pore.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="F7662CFD1EB9C49985F7F309EF6FF841" bold="true" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
. Pronotum about a tenth to a fifth wider than long (PW/PL= 1.06-1.17), widest point at second quarter. Anterior transverse impression absent, posterior transverse impression very shallow, barely distinct. Sides convex medially and anteriorly, slightly concave to base (somewhat more concave than in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="71C1A221D9AA9F8E3B6BD398179043E2" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="6A0DBE3B6CD1C3054468590F32E409B1" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
); anterior bead present laterally, reduced or lack in medial eight to tenth; basal bead present laterally, reduced to absent in medial fifth to half. Metepisternum slightly longer than wide or nearly as long as wide, MA/MI= 0.79-0.86.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="076FF38974E1936BCDE745B1785C1634" bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
. Oblong, about one-and-a-half times, or a little more, as long as elytra (EL/EW= 1.57-1.65), two-and-a-half times as long as pronotum (EL/PL= 2.41-2.61), and a half as wide as pronotum (EW/PW= 1.40-1.51), with widest point at medial third. Parascutellar striole and striae 1-7 rather shallow, impunctate or slightly punctate, stria 8 deeper than other striae; parascutellar striole not joining stria 1; bases of striae 1-7 mostly reduced, not reaching basal bead, rarely reaching it. Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 or 17 punctures on each side.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="D9FA9B14E86EA2048666DDB0646C32A3" bold="true" pageId="0" pageNumber="71">
<emphasis id="F6A89643D8C77EF859C8EA10B0751647" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur mostly with 5-6 setiferous punctures on anterior side ventrally, rarely with 4 punctures. Anterior side of metafemur ventrally mostly with one seta in basal third and one in medial third or apical half (single specimen with two setae in basal half and two in apical half on one femur).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="BC66EC6F0B44D3D726155B796DA763E7" bold="true" pageId="0" pageNumber="71">
<emphasis id="616CEB6DB92AEDABDFBDD9D90D2D3833" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view arcuate, with narrow basal bulb, long shaft and short, straight apex (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
<figureCitation id="BF3A7D25B1774D23445D5B6B86FE461C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
); same in ventral view straight, about 3.6-3.7 times longer than wide; apical lamella (dorsal view) short, almost symmetrical, rounded at tip and slightly concave at each side. Internal sac in lateral view (Fig.
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<figureCitation id="4B81317805C63E4D9EA4E7E014BB1B94" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
) with ventral sclerite long, slightly broadened distally; same in ventral view (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11D</figureCitation>
<figureCitation id="52CB3532089337BB8969116384E9B56D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11D</figureCitation>
), with dorsal sclerite large, forming two small left-sided protuberances and a broad right-sided protuberance, and with ventral sclerite narrow, straight and slightly broadened distally.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="3218E3B57DA13304C8CBBA5309E5727B" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15C</figureCitation>
<figureCitation id="FA7258D43CFA11F1BF7A68E7EBE0C641" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15C</figureCitation>
). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>

540
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2.
<taxonomicName LSID="194F053F-FAD4-5F5B-9889-0DF47AA646B8" authority="Maran, 1940" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">
<taxonomicName id="F012AEE70C4D3EC7B4E16AEA69B9835C" ID-CoL="4JQLP" LSID="194F053F-FAD4-5F5B-9889-0DF47AA646B8" authority="Maran, 1940" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">
Platyderus (Eremoderus) weiratheri
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<normalizedToken id="1EDCEDD3326AFAC7255CA3405AD0A631" originalValue="Mařan">Maran</normalizedToken>
, 1940
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1B</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7B</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8B</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10B</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12B</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13A</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
<subSubSection id="C1F896FAE46A784845F874F6FEA8344A" pageId="0" pageNumber="71" type="description">
<paragraph id="68A8ADB177171054AED35CF48E7D7569" pageId="0" pageNumber="71">
<figureCitation id="CB5D975D28FE1B26ED110B0F7C072B3F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1B</figureCitation>
<figureCitation id="266F8A446C0423663A630E827D208814" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5A</figureCitation>
<figureCitation id="32F5E7EF6AD1C7B8C9EDD5EE1AFDE5A1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7B</figureCitation>
<figureCitation id="02E75477AA4282CC40DB9519094C213C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8B</figureCitation>
<figureCitation id="B169A6662AA14FC3208EE0E5FFE96399" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10B</figureCitation>
<figureCitation id="154C519F4CE140EAFBED109B27FCD5E5" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12B</figureCitation>
<figureCitation id="72E208C954729B46F453FFF2228713C5" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13A</figureCitation>
<figureCitation id="CE10BBB25038AFA2F4B90D58519D1D24" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="reference_group">
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">Platyderus weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<subSubSection id="E2E1B5553C7740369AF45059489C44D9" pageId="0" pageNumber="71" type="reference_group">
<paragraph id="98168085DEB160162017EC2FAB285BA5" pageId="0" pageNumber="71">
<taxonomicName id="F45B665733CA5FFDDA6D12CB4DBC2114" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">Platyderus weiratheri</taxonomicName>
<normalizedToken id="D230F2DCF2D333A1FA896D5648DFB506" originalValue="Mařan">Maran</normalizedToken>
, 1940: 25 (type locality: &quot;Lydien in montibus Tmolos&quot; [=
<normalizedToken originalValue="Bozdağlar">Bozdaglar</normalizedToken>
<normalizedToken id="05A848734CC8347E19B6BDD0F4C7FBB4" originalValue="Bozdağlar">Bozdaglar</normalizedToken>
], Turkey).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="note on type locality">
<paragraph pageId="0" pageNumber="71">Note on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="6D62DC3C64EC74583DDDF43073FC8AF7" pageId="0" pageNumber="71" type="note on type locality">
<paragraph id="20EB4521EDEE320050D296F2467D0805" pageId="0" pageNumber="71">Note on type locality.</paragraph>
<paragraph id="CA34C24EFD224DE7DC10ACEE5BD27312" pageId="0" pageNumber="71">
The mount
<normalizedToken originalValue="Bozdağlar">Bozdaglar</normalizedToken>
<normalizedToken id="02E2A2AB3E2A10AFB2628ACC3C8BE721" originalValue="Bozdağlar">Bozdaglar</normalizedToken>
(Ancient Greek name:
<normalizedToken originalValue="Tmōlos">Tmōlos</normalizedToken>
<normalizedToken id="16B6E9E5EA25C76FD61BF282EBF6B5E9" originalValue="Tmōlos">Tmōlos</normalizedToken>
), with maximal elevation 2159 m a.s.l., is situated in the southwestern part of the Anatolian Peninsula.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<subSubSection id="4B2587E46AB391D6E6B30B39C6E0F758" pageId="0" pageNumber="71" type="references">
<paragraph id="E14D589103422ACF5972280F7C39EC7F" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="70DD136A6883FCB06771EB57FE96EFA8" pageId="0" pageNumber="71">
<taxonomicName id="FB83236573C27FB46251192AA5136B2E" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="05197794BD0213F8660849B8F96F4E4E" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
:
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" refId="B26" refString="Jedlicka, A, 1963. Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40-41(1962-1963): 16-22." title="Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40 - 41 (1962 - 1963): 16 - 22." year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<bibRefCitation id="DB7D24C24B6DA8A436AC8E28BE00EA2D" author="Jedlicka, A" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" refId="B26" refString="Jedlicka, A, 1963. Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40-41(1962-1963): 16-22." title="Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40 - 41 (1962 - 1963): 16 - 22." year="1963">
<normalizedToken id="E4DF086DE4813FE6566CD4A0F2570E4F" originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 22;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="A268F972616A224D3C921AECB1ACA0DA" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="967DF2FFFAA56571B415CE7B626102B8" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 523;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="72A6E9C8D6877D511C6A8169A184F07B" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 396;
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="13B1CC5AA4848A954BB9873A1DB39DF9" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 760.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="FB54559C655B04BAEFB4D20A43E409CF" pageId="0" pageNumber="71" type="type material">
<paragraph id="D130F105FDA4EEF1C9D6A83533021D82" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="639724FE72D820FE4AC4557445F2DCAA" pageId="0" pageNumber="71">
Should consist of two male and three female specimens (
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<normalizedToken id="63275BBFBB24ADC6C4CE9E28982141F5" originalValue="Mařan">Maran</normalizedToken>
ibid.) all of them syntypes according to Art. 73.2.1. (ICZN 1999). From these specimens, we found only one male syntype with extracted genitalia, here designated as lectotype. This specimen and its genitalia (with missing urite) are glued on two separate white cards on the same pin. The specimen designated herewith as lectotype is labelled as follows: '♂ [w, p] // Tmolos-Gbg., Lydien, / West-Klainasien. / Weirather, Innsbruck [w, p] // TYPUS [r, p] // Mus. Nat. Pragae / Inv. 927 [o, h/p] //
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">Platyderus weiratheri</taxonomicName>
<taxonomicName id="66F9EFED6B7992CE814458E68B2D7C17" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">Platyderus weiratheri</taxonomicName>
m. Dr.
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<normalizedToken id="9E028EC778AAB80A70608B73E5D6E6BD" originalValue="Mařan">Maran</normalizedToken>
det. [w, h&amp;p] // lectotype
<normalizedToken originalValue="label’">label'</normalizedToken>
<normalizedToken id="9C561AABA7C1DBA82D881DCACAD4F08E" originalValue="label’">label'</normalizedToken>
[NMPC]. The depositories and present state of remaining four type specimens remain unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Turkey</emphasis>
<subSubSection id="A70E45D63D708558858B489F6641204C" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="C1C304D38661C5B25B8B0BF0C5F7F12D" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="EF5A394525EF19751CC295D8367EEB87" pageId="0" pageNumber="71">
<emphasis id="AF12A6E9BF4E00DD83A94BE1CCAAB56A" bold="true" pageId="0" pageNumber="71">Turkey</emphasis>
:
<normalizedToken originalValue="Bozdağlar">Bozdaglar</normalizedToken>
<normalizedToken id="D47AF5D58A7A5197750490D3D71D307C" originalValue="Bozdağlar">Bozdaglar</normalizedToken>
(= Mount Tmolus) situated on territories of districts of
<normalizedToken originalValue="İzmir">
<collectingRegion country="Turkey" name="Izmir">Izmir</collectingRegion>
<normalizedToken id="2729DD7F1F5B377EFBB140E3B913F10D" originalValue="İzmir">
<collectingRegion id="E3EA0AABDCEF96EA74FA5AACF76B8135" country="Turkey" name="Izmir">Izmir</collectingRegion>
</normalizedToken>
,
<collectingRegion country="Turkey" name="Manisa">Manisa</collectingRegion>
<collectingRegion id="C167969C875BA270C1E7F7FDDB2FA9CE" country="Turkey" name="Manisa">Manisa</collectingRegion>
and
<collectingRegion country="Turkey" name="Aydin">Aydin</collectingRegion>
<collectingRegion id="176105B7EB8ECDF5CD0216F284A44E6A" country="Turkey" name="Aydin">Aydin</collectingRegion>
:
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Tmolos-Gbg., Lydien, West-Kleinasien., Weirather, Innsbruck" determinerName="J. Mueller" location="Coll." specimenCount="1" specimenCount-male="1" stateProvince="Mus">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="EB6ACBF7F6E16D9807B8650764830693" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Tmolos-Gbg., Lydien, West-Kleinasien., Weirather, Innsbruck" determinerName="J. Mueller" location="Coll." specimenCount="1" specimenCount-male="1" stateProvince="Mus">
<specimenCount id="937D50EE607B51CAF80AE75C92D22C28" type="male">1♂</specimenCount>
, '
<collectorName>Tmolos-Gbg.</collectorName>
<collectorName id="7AC25BE5CC24A372FEB0C7472CF2940F">Tmolos-Gbg.</collectorName>
,
<collectorName>Lydien</collectorName>
<collectorName id="B2E1A44E804C35D251FCF7571AFC36F5">Lydien</collectorName>
, /
<collectorName>West-Kleinasien.</collectorName>
<collectorName id="481B8B7AF18DEF61A5216882ED261518">West-Kleinasien.</collectorName>
/
<collectorName>Weirather</collectorName>
<collectorName id="F9B798DF8F2DC550014789ED81ED3BA0">Weirather</collectorName>
,
<collectorName>Innsbruck</collectorName>
<collectorName id="AAC191C233B7C58E0ECA30137E7984B8">Innsbruck</collectorName>
//
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="16B8C0DB4DC97248316B0CC4A639799E" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
spec.? det.
<determinerName>
<determinerName id="A8E8187E0F3320E96EE7238C9E15463F">
J.
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
<normalizedToken id="6F80B7249D6DDDDE4F0878791DE8C628" originalValue="Müller">Mueller</normalizedToken>
</determinerName>
(vergl. mit cyprius!) //
<location LSID="urn:lsid:plazi:treatment:194F053FFAD45F5B98890DF47AA646B8:690707EF46B9B3360CB9EE1C3D7E4EDD" name="Coll." stateProvince="Mus">Coll.</location>
<collectingRegion country="Turkey" name="Mus">Mus</collectingRegion>
<location id="690707EF46B9B3360CB9EE1C3D7E4EDD" LSID="urn:lsid:plazi:treatment:194F053FFAD45F5B98890DF47AA646B8:690707EF46B9B3360CB9EE1C3D7E4EDD" name="Coll." stateProvince="Mus">Coll.</location>
<collectingRegion id="39C4DAB79D04409DA2C0DC12172B415B" country="Turkey" name="Mus">Mus</collectingRegion>
</materialsCitation>
. / Vindob.' (NMW);
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Tmolos-Gbg., Lydien, West-Kleinasien., Weirather, Innsbruck" location="Coll." specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mus">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="75CBA545FD01211F1E3E84CACE4E4E3E" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Tmolos-Gbg., Lydien, West-Kleinasien., Weirather, Innsbruck" location="Coll." specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mus">
<specimenCount id="91395700504120932E93AC29C78D90B6" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="C0C4A7479D7EAAD8716222DA0EB120F7" type="female">1♀</specimenCount>
, '
<collectorName>Tmolos-Gbg.</collectorName>
<collectorName id="1A452D9B3B60AC2B4F51253BB7F394DC">Tmolos-Gbg.</collectorName>
,
<collectorName>Lydien</collectorName>
<collectorName id="5CC1425F5B677BEEAA3DBD557EA4F348">Lydien</collectorName>
, /
<collectorName>West-Kleinasien.</collectorName>
<collectorName id="9D9A1C6DC85F20EC0AEBED3A9EC5B01F">West-Kleinasien.</collectorName>
/
<collectorName>Weirather</collectorName>
<collectorName id="9DF82ABDCC878AF66B74B7C8A9CB422B">Weirather</collectorName>
,
<collectorName>Innsbruck</collectorName>
<collectorName id="34DA3DBFED8AD2C688E4F9B43660A545">Innsbruck</collectorName>
//
<location LSID="urn:lsid:plazi:treatment:194F053FFAD45F5B98890DF47AA646B8:E16118E79267371557474A703F63075B" name="Coll." stateProvince="Mus">Coll.</location>
<collectingRegion country="Turkey" name="Mus">Mus</collectingRegion>
<location id="E16118E79267371557474A703F63075B" LSID="urn:lsid:plazi:treatment:194F053FFAD45F5B98890DF47AA646B8:E16118E79267371557474A703F63075B" name="Coll." stateProvince="Mus">Coll.</location>
<collectingRegion id="91B93878FD5996FB221D7F077A8E2908" country="Turkey" name="Mus">Mus</collectingRegion>
</materialsCitation>
. / Vindob.' (NMW);
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Tmolos-Gbg., Lydien, West-Kleinasien., Weirather" location="Innsbruck'" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Mus">
<specimenCount type="male">2♂♂</specimenCount>
<materialsCitation id="868CACB486CED105F9D86758C82DD637" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Tmolos-Gbg., Lydien, West-Kleinasien., Weirather" location="Innsbruck'" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Mus">
<specimenCount id="7143A3BFA9EDFA571C92C9A6C723A6AE" type="male">2♂♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="7A4D4E2E1A49EC254839E45564E70865" type="female">1♀</specimenCount>
, '
<collectorName>Tmolos-Gbg.</collectorName>
<collectorName id="B36ACFE8E3419986DF68E0434EAE047C">Tmolos-Gbg.</collectorName>
,
<collectorName>Lydien</collectorName>
<collectorName id="8CBD5ACBD22085031B97AFE7FE4B7123">Lydien</collectorName>
, /
<collectorName>West-Kleinasien.</collectorName>
<collectorName id="18F2A3D4BE31461A6FF6445753C7C1F9">West-Kleinasien.</collectorName>
/
<collectorName>Weirather</collectorName>
<collectorName id="E84194FEF9D8F7E7C3F4F03E86FB821C">Weirather</collectorName>
,
<normalizedToken originalValue="Innsbruck’">
<location LSID="urn:lsid:plazi:treatment:194F053FFAD45F5B98890DF47AA646B8:8EA83D9B52D3CBCAF4E5273E1C95CAC3" name="Innsbruck'" stateProvince="Mus">Innsbruck'</location>
<normalizedToken id="E0D5C3F3C579AF7690C71D2624C847BD" originalValue="Innsbruck’">
<location id="8EA83D9B52D3CBCAF4E5273E1C95CAC3" LSID="urn:lsid:plazi:treatment:194F053FFAD45F5B98890DF47AA646B8:8EA83D9B52D3CBCAF4E5273E1C95CAC3" name="Innsbruck'" stateProvince="Mus">Innsbruck'</location>
</normalizedToken>
(NMW)
</materialsCitation>
;
<materialsCitation collectingDate="1995-05-05" collectorName="Boz" country="TURQUIE" elevation="1175" location="Izmir" specimenCount="1" specimenCount-male="1" stateProvince="Izmir">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="601442AC519D6B514C04FCE90DB518E3" collectingDate="1995-05-05" collectorName="Boz" country="TURQUIE" elevation="1175" location="Izmir" specimenCount="1" specimenCount-male="1" stateProvince="Izmir">
<specimenCount id="E929C4F2ACDC0E64F3BBB6EE0DC2F9C7" type="male">1♂</specimenCount>
, '
<collectorName>Boz</collectorName>
<normalizedToken originalValue="dağ">dag</normalizedToken>
<collectorName id="B770F08896CC2251000BFB440C51F142">Boz</collectorName>
<normalizedToken id="FA6A8816BAF305454AEDEFFDC0996884" originalValue="dağ">dag</normalizedToken>
, vt N /
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" unit="m" value="1175.0" valueMax="1200.0" valueMin="1150.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" unit="m" value="1175.0" valueMax="1200.0" valueMin="1150.0">1150-1200 m</elevation>
<quantity id="12811CFF5FF4E300A0D365886C8C0778" metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" unit="m" value="1175.0" valueMax="1200.0" valueMin="1150.0">
<elevation id="7FAB4016E634C4AC1617582E7A4C60DC" metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" unit="m" value="1175.0" valueMax="1200.0" valueMin="1150.0">1150-1200 m</elevation>
</quantity>
/
<collectingDate value="1995-05-05">5-5-1995</collectingDate>
<collectingDate id="D7064A4B7C15CC0DACEB5D7767930873" value="1995-05-05">5-5-1995</collectingDate>
//
<collectingCountry name="Turkey">TURQUIE</collectingCountry>
<collectingCountry id="669DE74570209B0CD2D3B2A5EB2E1A3E" name="Turkey">TURQUIE</collectingCountry>
/
<collectingRegion country="Turkey" name="Izmir">Izmir</collectingRegion>
<collectingRegion id="871C9F9C3EA4954C891E2059A9E8E284" country="Turkey" name="Izmir">Izmir</collectingRegion>
</materialsCitation>
/ C. JEANNE // Collection /
<normalizedToken originalValue="Machard’">Machard'</normalizedToken>
<normalizedToken id="DB372862EBA6340D6814B3672B8E3470" originalValue="Machard’">Machard'</normalizedToken>
(cMAC);
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Turkey" location="Izmir" specimenCount="3" specimenCount-female="3" stateProvince="Izmir">
<specimenCount type="female">3♀♀</specimenCount>
<materialsCitation id="A743ED2D3AA4B4A9AE2EC59BCDA5480A" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Turkey" location="Izmir" specimenCount="3" specimenCount-female="3" stateProvince="Izmir">
<specimenCount id="6803601C33994A9BEC961D8D846F145B" type="female">3♀♀</specimenCount>
,
<normalizedToken originalValue="ʻTURKEY">ʻTURKEY</normalizedToken>
<normalizedToken id="FAFE37EDB26B09BC9BA777B3AD85D4AB" originalValue="ʻTURKEY">ʻTURKEY</normalizedToken>
, vil.
<collectingRegion country="Turkey" name="Izmir">Izmir</collectingRegion>
<collectingRegion id="CCD873D68D508CC213A8399FB30238E3" country="Turkey" name="Izmir">Izmir</collectingRegion>
</materialsCitation>
, Boz /
<normalizedToken originalValue="dağlari">daglari</normalizedToken>
<normalizedToken id="737225FC244C2F9FA722B00EFCF28BEC" originalValue="dağlari">daglari</normalizedToken>
, Boz
<normalizedToken originalValue="dağ">dag</normalizedToken>
<normalizedToken originalValue="köy">koey</normalizedToken>
<normalizedToken id="A667BDB809F8756D6DF8C94C96115508" originalValue="dağ">dag</normalizedToken>
<normalizedToken id="3CBE6780CAB5C5B6A07FF1C764EE487A" originalValue="köy">koey</normalizedToken>
env. /
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500-1700 m</quantity>
<quantity id="815F931BF21F3BB285205781FF95D4AA" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500-1700 m</quantity>
a.s.l. / 30.5.-3.6.̍03, R.Lohaj lgt.' (NMNHS, cWR).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="6D09EE74E3291CC8B4B0A82117C13479" pageId="0" pageNumber="71">
TME:
<specimenCount type="generic">11 specimens</specimenCount>
<specimenCount id="E461448A9AABF6BACF71C022CDDD50C0" type="generic">11 specimens</specimenCount>
. TGE:
<specimenCount type="male">2♂♂</specimenCount>
<specimenCount id="A73A86F2736F6503C39869323F1DFAC0" type="male">2♂♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="26868519B328115EBA3416F70DE164F7" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="C47F2810D4E5E388D144788F6D7652DA" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="ADDAE3FA10264480D307A8A2CD8AE5BC" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="C3E39DF780749AD7465C3B19E389DFD3" pageId="0" pageNumber="71">
See
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="BC623EF3CAA07DF1C7342D5B111940BA" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="134A6D37581E6097DFEF4B9E6FCF72D3" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="4BD85A0334EC3B7976B23A1A3AB31ED9" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="redescription">
<paragraph pageId="0" pageNumber="71">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="70BADCBE615D6801462ACD3BF7FB72AF" pageId="0" pageNumber="71" type="redescription">
<paragraph id="09FB5D4FB4A25D0F0E548FE675881CF0" pageId="0" pageNumber="71">Redescription.</paragraph>
<paragraph id="687C1EE276207F01DC53DF617E635D8F" pageId="0" pageNumber="71">
<emphasis id="20BB1F4C2FCDAD8E133337085B200201" bold="true" pageId="0" pageNumber="71">
<emphasis id="8C8EFB4E08A098E684CB8D2B4BE7357B" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="BD570D7DB30BF965A274AA57D73609DB" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="7D2B3C906AE73937BF7ED59B51167886" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 7.00-8.10 mm; BW: 2.35-2.60 mm), with rather elongate, moderately convex body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1B</figureCitation>
<figureCitation id="C296DC3430E7E4FC66BB7AD659210897" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="E34136A9D376732358372DA47378D540" bold="true" pageId="0" pageNumber="71">
<emphasis id="89546DBE8CA7D5CF636423545B4EE6EA" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
<tableCitation id="684C2124D9F27FA98A800B5E8221C435" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="13F7A6B21F78E8AB5BA5887E88526096" bold="true" pageId="0" pageNumber="71">
<emphasis id="F52AE9F34F3EB22BF161F5E79720ACC7" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body dorsally and ventrally dark reddish, head and pronotum mostly lighter than elytra, appendages (antennae, palpi and legs) rufous, lighter colored than dorsal surface. Integument moderately shiny, head and pronotum slightly shinier than elytra.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="C9641C34583BD333D191D61902452E81" bold="true" pageId="0" pageNumber="71">
<emphasis id="146FA96C5DB403C5472EF4F370694079" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum without regular microreticulation. Elytral intervals and scutellum with distinct isodiametric sculpticells; basal margin and lateral gutter with such sculpticells less evident or without microsculpture. Ventral surface with hardly visible isodiametric (proepisternum) or slightly transverse sculpticells (abdominal ventrites at sides) or mostly sculpticells not apparent. Head with some punctures and wrinkles only clypeus and frons, labrum, disc and vertex without punctures and wrinkles. Pronotum in basolateral part coarsely and sensely punctate, basal punctures reaching or not reaching the anterior half laterally, disc and anteroapical part mostly smooth, anterior third medially finely punctate and wrinkled. Elytral intervals mostly with a row of punctures, rarely without punctation. Abdominal ventrites 1-5 wrinkled at sides.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="102F42C37EBAF42A0CDB26CCC4CD15AB" bold="true" pageId="0" pageNumber="71">
<emphasis id="341473A464D4BB763F4F36A127DD1D60" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
One-third or narrower than pronotum wide (PW/HW= 1.32-1.44). Eyes slightly convex. Labrum subrectangular, slightly shorter than clypeus, with anterior margin concave. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows barely distinct, small and shallow. Paraorbital sulci straight, moderately deep, ending backwards slightly before level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="62EE2283E8104EDE97F405BBD795EF95" bold="true" pageId="0" pageNumber="71">
<emphasis id="2B2A92DCD624C182E15E174CDBDA7B96" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about one eighth to one quarter wider than long (PW/PL= 1.13-1.27), with widest point at second quarter. Anterior and posterior transverse impressions slightly distinct medially. Sides sinuate, convex medially and anteriorly, slightly to hardly concave posteriorly; anterior border lack in medial 1/10. Metepisternum about as long as wide, MA/MI= 0.95-1.02.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="4A90FA0D514082C6706AC2E1563DCD04" bold="true" pageId="0" pageNumber="71">
<emphasis id="D4257CFC4AC3CF772E1F10B2E7C85D92" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about two thirds longer than wide (EL/EW= 1.63-1.71), two and a half times or longer than pronotum (EL/PL= 2.55-2.83), and a third wider than pronotum (EW/PW= 1.29-1.40), with widest point at medial third or third quarter. Parascutellar striole and striae well-impressed, more coarsely punctate than in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="08B64D0166B3EDD92B47E94B9195C460" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="EBCE78CC5502103167B3D32E92C1942B" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
sp. nov. and less coarsely than in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="B1BF411F3697E236A4D0B7D516C5BA3A" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="F746682955021D62DAB914DF8C82003B" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov.; striole short, not joining stria 1; striae 1-6 reaching or almost reaching basal bead, 7 reduced before. Interval 3 with three discal setiferous punctures on each elytron. Umbilicate setiferous series consisting of 15-16 (rarely 17) punctures on each elytron.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="67FF0E189EA61946AB4C5F3C90EEC98E" bold="true" pageId="0" pageNumber="71">
<emphasis id="AA9D91E7FB90B0764C706EA1D605DF3E" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one seta in medial third. Anterior side of mesofemur ventrally mostly with 4 equidistant setiferous punctures, rarely with 3 or 5 punctures (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5A</figureCitation>
<figureCitation id="983C2C6C1C4E5CFAC1E74BBF44DAD896" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5A</figureCitation>
). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="CEC0997A937B316BDD62DA3818E740F7" bold="true" pageId="0" pageNumber="71">
<emphasis id="470180EFD9B1AA0BC0ADBE500EA408A8" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX with proximal margin almost asymmetrical (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7B</figureCitation>
<figureCitation id="F38C9C23B1B9DFEAE503320B49C9E35C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7B</figureCitation>
). Median lobe of aedeagus in lateral view as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="27F934F75B755793DD251CA6CC0A0CE2" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="00A60B52CB2C6CBDBEB1D03E47A3F140" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
, with shaft little longer (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8B</figureCitation>
<figureCitation id="AEE8B5A66CF4326115BA61C51EEE077C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8B</figureCitation>
); median lobe in ventral view straight, about 3.5 times longer than wide (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10B</figureCitation>
<figureCitation id="1938E70D09F082FE3D9FC9C3C10109E2" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10B</figureCitation>
); apical lamella (dorsal view) as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="CE65BDEBCB70580AA08BDD507175FFA5" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="E0B54496F720DB47564D7128030DEC0B" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
. Internal sac as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="B04CC5E072FFDDB0F9E4FF7FBD481F13" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="421AB44E09C6F3B99D4D671B422004D8" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
, only ventral sclerite narrowed toward both proximal and distal ends, with widest point in middle (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10B</figureCitation>
<figureCitation id="B024DEA42A659D84A2D3004874914D35" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10B</figureCitation>
). Right paramere on Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12B</figureCitation>
<figureCitation id="7DB4B0853E56D7866FA98D982D46D165" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12B</figureCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="CDEC1231FFB90553BFC76C0A3B712470" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13A</figureCitation>
<figureCitation id="BB59DBD415B8CFFF0AFDB140B01E9370" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13A</figureCitation>
). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 2.</emphasis>
<caption id="9E777D603170B8B220F0B94C8FEF4DB4" doi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71" start="Figure 2" startId="F2">
<paragraph id="78B8EAA27518C641B17D5D7D9E625DBD" pageId="0" pageNumber="71">
<emphasis id="8C5AB990BA2AA5C3AD211354032DDABB" bold="true" pageId="0" pageNumber="71">Figure 2.</emphasis>
Dorsal habitus.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authority="" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus (Eremoderus)</taxonomicName>
<emphasis id="0D7CCCA2ADE1AB6C6B7E4499690A9AD9" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="51B8F44CA4FF6B166F843DA4BCE328DB" authority="" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus (Eremoderus)</taxonomicName>
sp., female specimen, Karabet Pass, Turkey;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
<emphasis id="E38565C6C5F51EB22B366B812C431BA4" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="268C2890949EA3BFD34245E419D00F45" class="Insecta" kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi" subGenus="E.">P. (E.) klapperichi</taxonomicName>
<emphasis id="A6C5462963346F796D6EF9493D5D786A" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="014FC7025E2D1BF69CE4B28F1E33B3FA" class="Insecta" kingdom="Animalia" lsidName="P. (E.) klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi" subGenus="E.">P. (E.) klapperichi</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
<emphasis id="9A8985E5BE421EFF52976CED05661B4D" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="78165708CF5B2F197881BCEF858B2B15" class="Insecta" kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
<emphasis id="E15C4AD41FE68E1D8653FBCCF9E6EBBB" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="98E20BB25E84FC4052BDDCBB3FF3E961" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">Nothing is known about bionomics of this species.</paragraph>
<subSubSection id="857D1F28536268C18915738F860E5526" pageId="0" pageNumber="71" type="habitat">
<paragraph id="0D3A9BFC9B056F4ACD02557C07CF2DC6" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="01917F6263C3A8F8C0576D9E55A84067" pageId="0" pageNumber="71">Nothing is known about bionomics of this species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="FF24048FFF0ED79D5E23E6E8AAADD24C" pageId="0" pageNumber="71" type="distribution">
<paragraph id="9BD0F96A6AC7AAF6705E6D48A992E9AD" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="397074758F1F308AAE3AD40161487AC9" pageId="0" pageNumber="71">
Apparently endemic to the mountains of the
<normalizedToken originalValue="Bozdağlar">Bozdaglar</normalizedToken>
<normalizedToken id="9E680CBF9B59DBAA2C119A796E603759" originalValue="Bozdağlar">Bozdaglar</normalizedToken>
Mountain, in Southwest Turkey (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
<figureCitation id="EC1313626A63E114C583ED177DA1B2DB" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="E735893D4BCD58BB403695E28BB1C059" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">Platyderus Eremoderus iranicus</taxonomicName>
<emphasis id="1A9C6F6495876DA58093AD29C4CA4221" bold="true" pageId="0" pageNumber="71">
<emphasis id="44F4C0BE7D8986933F5E1F4086425972" italics="true" pageId="0" pageNumber="71">
<taxonomicName id="C73F22E1EC06C99D110C7514776D34D2" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">Platyderus Eremoderus iranicus</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">Platyderus Eremoderus vanensis</taxonomicName>
<taxonomicName id="99AC1CB161C4556A125453191B37BB2B" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">Platyderus Eremoderus vanensis</taxonomicName>
</emphasis>
&quot; group
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
<paragraph id="354AF632C3AAD58F3D825C8A68C34567" pageId="0" pageNumber="71">
<emphasis id="049935EAB1DCC2AAA631719C182E7EC3" bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
Includes species with basal foveae of pronotum and adjacent lateral areas coarsely and densely punctate with punctures at sides usually reaching anterior half. The prosternum laterally and proepisternum also coarsely and densely punctate. The ventral sclerite of internal sac of median lobe in ventral view is elongate, relatively wide, with straight distal end; the same in lateral view is significantly widened anteriorly having maximum width at the distal third about three or more times larger than width at the proximal third (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C-F</figureCitation>
<figureCitation id="C589C968754080CA35058003A57C583C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C-F</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10C-F</figureCitation>
<figureCitation id="D4C8AF9495666412FDDE51ABFDA635C3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10C-F</figureCitation>
). Most specimens lack also the first elytral discal puncture and have apical gonocoxite with proximal dorsolateral ensiform seta finer than the distal seta.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="55E3C149CCC0ED9D579BB1C0C21F9413" pageId="0" pageNumber="71">
In addition, the Iranian congeners of the &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<taxonomicName id="0CCDDEA904ED2B0DAB965DAD1FC8F24D" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="44D295B7FC41C7CB364D617DD6E59D93" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
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<emphasis id="A62165061CC2C4B937E43376D96E680F" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
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&quot; group differ from those of the &quot;
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="EAE413958F0503D338DB7C2AA7ADA1C9" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="42D51FB3FF7FF698E93F65E22445F624" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
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&quot; group in: (1) sides of pronotum nearly straight to the posterior angles (vs. sides of pronotum concave to the posterior angles) and (2) pronotum appreciably wider than long (PW/PL&gt;1.20, vs. PW/PL &lt;1.20). For further differences between taxa of the group dealt with here and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="FC974A50E436AE2F11D9825F0F9503D0" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="C09E087288FC2D48407C47C97A807839" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
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see
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="D335CE95C5606B200075554DE9C1EA67" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
under the &quot;
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
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<emphasis id="81DE7BF6E953CC081682FFA3CE87E3FB" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
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&quot; species group.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Habitat.</emphasis>
<paragraph id="B310F01C8BC7858B4434D2F7B97ED105" pageId="0" pageNumber="71">
<emphasis id="DFD17DCADFD317B900AC7D8CFD1C3CDE" bold="true" pageId="0" pageNumber="71">Habitat.</emphasis>
The habitats of the
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="3D92BF436ACC5718382DB7E0AB4F2911" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="30075FE75D920A4427C86D4E3A95D0E4" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
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-species in the Zagros (Iran) consist mostly of subalpine slopes, partly near snowfields, and seldom of cultivated grasslands and small orchards. Outside the cultivated grasslands along the rivers and streams the vegetation is xerophilic. The altitude of the area where
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="107E9D1CB88B980297C906DE3947DE16" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="B8458DDEDA37BCC433694863CD6A4D69" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. was collected is appreciably lower than that of the localities of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
<taxonomicName id="5042A6617C472FEC46EE378D9E961103" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="C99F6E76106B53ACBDB55204A61CA052" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="7D358A436809C919F50361B6D2BBACD5" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<emphasis id="4F7F2731FA0A83563407900E8F830F14" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="FCB5A67AE29F5C9F15AF2EAB47ACC793" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="F475DCCF90FB18057F5AE9A78F951A5A" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
&quot; subgroup
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
<paragraph id="0C8574FA72663A4201522479C78CFBB3" pageId="0" pageNumber="71">
<emphasis id="08B0B4FE56CDB5D52E4635A913DF0F94" bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
This complex includes two allopatric, high altitudinal species,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="394470D20E25E34D5082E4079587FD00" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="6B9F9758299CAE9E3BF3A1BA4DFFAF5C" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
<taxonomicName id="541EC7B488BA190F6058D2069C93CAB3" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="64557CFC0874420EAADBA76610D8D417" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
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with parascutellar striole, striae 1-8 scarcely punctate and moderately impressed and quite short and crooked right parameres (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12C, D</figureCitation>
<figureCitation id="B31CECA8F5797AA75A138E0A51355FC4" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12C, D</figureCitation>
).
</paragraph>
</subSubSection>

548
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@ -1,513 +1,513 @@
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<treatment LSID="urn:lsid:plazi:treatment:1AEE6C1E66755EF2B86CF9E2E9048806" httpUri="http://treatment.plazi.org/id/1AEE6C1E66755EF2B86CF9E2E9048806" lastPageNumber="71" pageId="0" pageNumber="71">
<subSubSection pageId="0" pageNumber="71" type="nomenclature">
<paragraph pageId="0" pageNumber="71">
<treatment id="1AEE6C1E66755EF2B86CF9E2E9048806" ID-GBIF-Taxon="195868760" LSID="urn:lsid:plazi:treatment:1AEE6C1E66755EF2B86CF9E2E9048806" httpUri="http://treatment.plazi.org/id/1AEE6C1E66755EF2B86CF9E2E9048806" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Sphodrini">
<subSubSection id="E3ACFC517A820C33F078E85618BE12CA" pageId="0" pageNumber="71" type="nomenclature">
<paragraph id="1BE5BC296543C143472E6EB304B3446A" pageId="0" pageNumber="71">
Subgenus
<taxonomicName LSID="1AEE6C1E-6675-5EF2-B86C-F9E2E9048806" authority="Jeanne, 1996" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus Jeanne, 1996</taxonomicName>
<taxonomicName id="D077E72BA18481A321B10748F5692FC2" LSID="1AEE6C1E-6675-5EF2-B86C-F9E2E9048806" authority="Jeanne, 1996" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus Jeanne, 1996</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type species">
<paragraph pageId="0" pageNumber="71">Type species.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Reiche &amp; Saulcy" authorityYear="1855" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">
<emphasis italics="true" pageId="0" pageNumber="71">Feronia languida</emphasis>
<subSubSection id="881CD18168B2548EFB6D1035860F2E08" pageId="0" pageNumber="71" type="type species">
<paragraph id="8008AE9CA0428A670DBEF1A49B976CF4" pageId="0" pageNumber="71">Type species.</paragraph>
<paragraph id="FEA685399A849C7C3F87AEA206DBB34B" pageId="0" pageNumber="71">
<taxonomicName id="D622B4F62EBD17EDF43C8257A77E306A" authorityName="Reiche &amp; Saulcy" authorityYear="1855" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">
<emphasis id="0E9CF9ED2C4BC822D73D1E37229F43E7" italics="true" pageId="0" pageNumber="71">Feronia languida</emphasis>
</taxonomicName>
Reiche &amp; Saulcy, 1855
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="4D7DEA6E10E2505B4210FD454F2517D1" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="BDB98600FA8C875C48B9A1757F6DE7A0" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="8FA3E91B96A60C59116C9B2D5283E1F9" pageId="0" pageNumber="71">
Within
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="11DD5D067959E3C275F72BFFCA2247F5" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="63D5A1E403688D912DD8B375B1E60828" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
, the members of
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="E18D5BA12FE83FF5B8AB70611BF71E00" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="EC3DD9CB20B6523668DBB528D7EE8614" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
are separated by the following set of distinctive characters: ventral sclerite of median lobe of aedeagus narrow and long (best seen at ventral view, Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A-I</figureCitation>
<figureCitation id="8EED55A5B9C7D9D9A67AD67AE392C071" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A-I</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11A-G</figureCitation>
<figureCitation id="C6AE0AE96152AA24CA1AED22B1F4222A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11A-G</figureCitation>
); seminal canal and receptaculum of comparable lengths (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13A-F</figureCitation>
<figureCitation id="C7A8D8BA4BABE2015662471B08744E88" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13A-F</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14A-C, E, F</figureCitation>
<figureCitation id="43A98F06E56D31A8DDDF962959111CCE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14A-C, E, F</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15A-F</figureCitation>
<figureCitation id="E11130CE9F8C811F0C1A41F29CDFBA80" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15A-F</figureCitation>
); ventral margin of anterior side of mesofemur with four or more, rarely three (specimens of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="27FF998A14681298BD17924E83EC7608" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="A7842230F98757BEA64FFE2E3AE8C743" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<taxonomicName id="F54D6635C1EB152D116D54F9CBEC5302" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="77D8B14AFFFE4C073DE957143C7BF774" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
) setiferous punctures ventrally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5A-I</figureCitation>
<figureCitation id="4486718857C5956F1D7F1F2CA927D3FE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5A-I</figureCitation>
); proximal margin, i.e. the
<normalizedToken originalValue="“apex”">&quot;apex&quot;</normalizedToken>
<normalizedToken id="836C765FB8CD3F01EAFE752FD187D27A" originalValue="“apex”">&quot;apex&quot;</normalizedToken>
of urite IX symmetrical or nearly symmetrical (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7A-M</figureCitation>
<figureCitation id="33A58E92AC22800864AB47820BB19695" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7A-M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and micropunctation</emphasis>
<subSubSection id="985F6FBD0C709A5BE59751EF94F7ED95" pageId="0" pageNumber="71" type="description">
<paragraph id="1087C9A2ED4FD87417B1283993C4C8B5" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="2667408B3418F8CD0FA30C2652980471" pageId="0" pageNumber="71">
<emphasis id="EDED0B25BE23338247D4D5CAB5C9CDE1" bold="true" pageId="0" pageNumber="71">
<emphasis id="0FE5DA13AFCD585685A677C15BB08F64" italics="true" pageId="0" pageNumber="71">Microsculpture and micropunctation</emphasis>
.
</emphasis>
Dorsal surface of head with regular isodiametric to slightly transverse sculpticells that are more or less reduced on posterior half of clypeus and disc (most species) or with complete microreticulation of isodiametric sculpticells (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="CF8C5DFF4931336C03C288E31D18D6CC" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="B6FD667746A395FB90DFDC9324CE86A0" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group), labrum with coarser such sculpticells than remainder of head. Elytral microreticulation distinct, usually more engraved and larger than on head and pronotum, represented by regular isodiametric sculpticells (some specimens of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="47F11AF032F032BC864081E3DCD9FF62" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="3D756A462A07660369FBAD482FF618A3" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. without micropunctation on elytral intervals 1-5). Dorsal surface of head and pronotum with scattered micropunctation hardly visible below magnification 90
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="01B37375E848D802170910C8C7AC666F" originalValue="×">x</normalizedToken>
, elytra with coarser, more distinct micropunctation, visible at magnification below 90
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="55B85E760C9A21CABDE2123095F6F713" originalValue="×">x</normalizedToken>
(micropunctation absent on dorsal surface of head and elytra in most taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="86E860FC8D1560EDD426157760947243" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="1051A62E532528BF83F98708A2F3293C" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group, except for head of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="9CC7EF5BF97D23A569AA22006F464F7F" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="CE3CC9CD7ECC8A25506EC3715A7D8BF5" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Punctation</emphasis>
<emphasis id="9C1F4A5F9793790788554F2159C47D50" bold="true" pageId="0" pageNumber="71">
<emphasis id="7D47B8B19FE8D1341273DF797DD7E02C" italics="true" pageId="0" pageNumber="71">Punctation</emphasis>
.
</emphasis>
Basal foveae of pronotum and adjacent lateral areas moderately punctate, punctures at sides usually not reaching anterior half (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<taxonomicName id="E421BB8FDB50E35A4E611A1926732EC7" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="C8633D8DC3498676296932EE368EF0BD" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
, taxa of &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
<taxonomicName id="46D9B3B19382E18ECCEE338EDDBC87FA" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis id="5E661FD08252A8BDE495EAA2CD89711D" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
</taxonomicName>
&quot; and &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="0D2FCF44D877F7F31DD896596B682684" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="C25E010F798A5AB79AA3C8FD65FF2692" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; groups), coarsely and densely punctate as punctures at sides usually reaching anterior half (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="FE1500AB0901003F64FB9DE51A513C97" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="06A018933DD24FDB1C5A5C460B52C8DB" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
n.sp., taxa of &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<taxonomicName id="7B5232B599D85EF4692179858D91B09B" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="D4D6E96CB1D9417E8A6D3EDC742479C5" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
<taxonomicName id="C2DAAD2DB566B106167E0346E3077282" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis id="BA5BF98EF773F0571B2289F84C349CEB" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
</taxonomicName>
&quot; group), or scarcely punctate to impunctate (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
<taxonomicName id="39758E09C0A0FE376D2F8C5A4EF31BA9" class="Insecta" kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis id="2E004A97155C64F7B8A986858F0721C1" italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
</taxonomicName>
sp. nov., taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="D842BB6379FFB3D5B74BF1DA54620BE6" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="AE663AA0B496481A311F0FF5A907B48B" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group). Prosternum laterally and proepisternum moderately (most species) to coarsely, densely and deeply punctate (taxa of &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<taxonomicName id="D5B79AEE3520F132F3BA90F7F7784DD2" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="125476AC1522102C7E95883F347C11F6" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
<taxonomicName id="108D8B4827E50DAC6313489B6344E7D2" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis id="53C36A59D8A3B3B0E1540AD36F5C6A83" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
</taxonomicName>
&quot; group) or finely, sparsely and shallowly punctate to impunctate (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="FA688EA3F2F25497838F950F652A9E94" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="E8FB6F123623C4B2C692A63DB5A8C710" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group). Prosternum medially and metasternum medially smooth. Mesepisternum, metasternum laterally and metepisternum indistinctly (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
<taxonomicName id="EEC4E7D73318DF8B073615A1DC77382E" class="Insecta" kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis id="D288536FFDAF0FD25A49289AB947434C" italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
</taxonomicName>
, taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="331FD6CC5AE356608A215DF3D4A29F55" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="2F65BD56150D88081DA0810EB75E2817" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; groups) or clearly punctate (taxa of resting species groups, as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="E133DC502DBB3CFD4B0F356E1265EB52" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="25A8D88FB49BDB2976A96D3209EB2DC6" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. more coarsely than in other species). Abdominal ventrite 1 punctate and/or wrinkled, sometimes impunctate, ventrites 2-3 wrinkled, punctate or impunctate at sides, smooth medially, ventrites 4-5 mostly impunctate and smooth, but sometimes finely wrinkled at sides, 6 usually impunctate and smooth in whole (ventrites 1-4 coarsely punctate at sides in eastern populations of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="74ECE0B528D580FD905F5D68CAF35276" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="E72684311A5F17AFFFD5CDE5B4F7C235" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov.).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Chaetotaxy</emphasis>
<emphasis id="2C4CA30FD2BBE67D7E101D00E2139BD2" bold="true" pageId="0" pageNumber="71">
<emphasis id="6D903E29AF043F607290CA49E7C633A9" italics="true" pageId="0" pageNumber="71">Chaetotaxy</emphasis>
.
</emphasis>
Labrum with six equidistant setae, lateral two longer than medial four. Clypeus with two long anterolateral setae. Two supraorbital setae each side. Stipes with anterior and posterior setae, former about half the length of latter. Pair of long setae on apical margin of ligula. Penultimate labial palpomere with two long setae on internal margin and 2-4 rather short apical setae; terminal labial palpomere with some scattered, very short and fine hairs. Mentum with two short and fine paramedial setae (sometimes broken off). Submentum with two long medial setae and two very short and fine lateral setae (occurring in most species but often broken off) or without lateral setae (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
<taxonomicName id="01ADB6D7723FF21B0D3BD1A2E0049E2A" class="Insecta" kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="9823C5699C8C7E359875A1F1E43D82B3" italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
</taxonomicName>
sp. nov.). Pronotum with one lateral seta at second quarter and one basal seta near posterior angles. Elytron with parascutellar seta at junction of angular base of stria 1, puncture small; interval 3 with three (rarely two) discal setiferous punctures (in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="3D3FD4D1FBD8684F3BF3310C8AA4EF8A" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="296ACD23CA1726C9CA441D1B6C296EAA" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov., punctures hardly discernible because of coarse and deep macropunctation of interval), first one adjoining stria 3 (rarely in midst of interval 3 -
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="4460A5F1F6F0B2CB945F8F0E886C0319" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="5C1D14BCD1303D8DB76E54DB07FC38F7" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
sp. nov., adjoining stria 2 -
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="6C01530BE19F17EBFA658A5017397A9F" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="FAECADB425E5D87BD6CED615F5053000" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. or absent - in most specimens of &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<taxonomicName id="82A916498CC4BF0D5C38ADC90CDD5B39" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="9D3A05BCA3803595A4BF870FFED28581" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
<taxonomicName id="15E54163181A45CCDEF37BF81FED0256" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis id="4735E45612480A159AAC3B0DA8083CFE" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
</taxonomicName>
&quot; group), second and third punctures adjoining stria 2, first puncture situated in anterior third, second puncture in medial third or third quarter, and third puncture (lacking in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="1E405E9745432E02DE315BA614BF66CB" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="8F0DD854C20FCA6F8C23E4C8AAD46F93" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
sp. nov.) in posterior third to fifth; apical portion of stria 7 with two setiferous punctures, as posterior one situated closer to lateral gutter than to suture; umbilicate series consists of 16-17 (rarely 15 or 18 on one elytron) uniform punctures, not forming compact groups, anterior three punctures on lateral gutter, remaining ones on stria 8. Posterior side of profemur with three (most species) or more (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="001520890AD52D59B36062A56B4F51E3" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="BDF6C7DF7CAA7DE0B64849A4233E78E4" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="C07EDD429844783A3A61F965E28687C8" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="C09A022DF27C5D946975480F5CB1958E" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="6016B46441965DDAFF55A1CB9CE1CAAF" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="77A1AD2710E1B857AE29B4AC56FFB329" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="C2290C8BC2D4811EDA4D94F8DD29109E" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="F36024ED7E92A30B8BE412D5536BF47F" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
) long setae, one to two in basal third, one to two in medial third (near ventral edge), and one in proximal fifth (near knee). Anterior side of mesofemur ventrally usually with four or more setae, rarely with three setae; mesocoxa surface vaguely (most species) or densely pubescent (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="A3EFF2FDEAD40C9DE0262C2253D3B592" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="E5D11F421F29F66E60C93D28C6D9896E" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group), with one posteromedial seta and one (most species) or one to four lateral setae (taxa of &quot;
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<taxonomicName id="BE48BEB997D44783AF35A3700B30C51D" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="B8C43C5D73050FA5C8651A646972669E" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
&quot; and &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="755FB5D88ACADED2EC2820274F372F6B" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="6B66E65D84368437480EF790061C460C" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; groups); mesotrochanter with seta. Anterior side of metafemur ventrally mostly with one seta in basal third and one in medial third (additional setae existing in specimens of taxa from &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="A5DB32CC963B9109186873036FE820D1" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="5458682A6B52DC007F3F7754EF748DAC" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group). Abdominal ventrites 3-5 with ambulatory setae; last ventrite with two setae in male and female.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="97CC3D9C6E5B35179A51FE4CED7C9911" bold="true" pageId="0" pageNumber="71">
<emphasis id="51A28AD96996307B6A10F23A940C3A0F" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
Eyes subconvex, long, each with length little exceeding length of scapus; tempora oblique, straight or convexly converging to neck. Labrum as long as or slightly shorter than clypeus. Clypeus slightly concave in front, rarely straight (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
<taxonomicName id="D36E3F6BD9A0F2E61FCB2846CC803BAC" class="Insecta" kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis id="05C59882DD250A228029BCB497B662B3" italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
</taxonomicName>
sp. nov.). Mentum and submentum separate by distinct labial suture; mentum wide, short, deeply emarginate, without paramedial pits, median tooth, prominent, rounded anteriorly and bordered at base, epilobes well-exceeding median tooth in front, with posterior border medially concave.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="3152766E5DA8297E4189D4BCF349E3B0" bold="true" pageId="0" pageNumber="71">
<emphasis id="1979F5001B840B5438D53471BEAE42E4" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Disc of pronotum barely to moderately convex; median line well-impressed, long but not reaching anterior and posterior margin; base with one sublinear fovea each side, impressed on basal third, foveae well-impressed (most species) or shallowly impressed (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="3E85820D0426E8130FA921BF591005B8" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="BDE96BF15579DB4C84821605B00508BB" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group); anterior margin narrower than posterior margin, slightly concave each side, median projection covering
<emphasis italics="true" pageId="0" pageNumber="71">pars stridens</emphasis>
<emphasis id="171AEF748FD9BDB558FA391408C0B55A" italics="true" pageId="0" pageNumber="71">pars stridens</emphasis>
(stridulatory organs); anterior bead narrow, present laterally, impressed, present, lacking or indistinctly impressed in medial eighth to tenth; anterior angles well-projecting, rounded at tip; sides convex, straight or concave to base, lateral beads fine, complete to posterior angles; posterior margin concave in middle, slightly convex near angles; basal bead present laterally, present or reduced medially; posterior angles obtuse, projecting or not projecting laterally, rounded at tip. Prosternal process bordered, subovate, rounded at apex. Metepisternum narrowed behind, its internal margin as long as, or longer than, anterior margin.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="221DFCCED478AE52A4C45C783092F28A" bold="true" pageId="0" pageNumber="71">
<emphasis id="D200CD820C0B1458B440E5C180F3AE73" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elytra coalesced along suture, widest at middle third or third quarter. Disc convex; humeri widely rounded. Basal bead arcuate medially, reaching or almost reaching scutellum, laterally forming obtuse angle with lateral margin and no denticle at humerus. Sides from basal margin towards middle third smoothly widened, from there to apex roundly narrowed. Parascutellar striole, if present, short, joining (rarely) or not joining (mostly) with stria 1; striae 1-6 usually reaching or almost reaching basal bead, 7 reaching or not reaching to it; base of stria 8 ending at third umbilicate puncture; striae 1, 7 and 8 apically separately joining lateral gutter; striae 1 and 2 mostly fused shortly before apex and reaching lateral gutter (if stria 2 not joining stria 1, it ends before lateral gutter); pairs of striae 3-4 and 5-6 fused before apex but neither joined stria reach lateral gutter (joined stria of pair 3-4 ending more apically than that of pair 5-6; in two specimens of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="B3402D0A6DB4FD0173A108AD464EAF92" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="AB3210FE708410778F8ECF707C896F04" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov., following aberration exists: striae 3 and 4 posteriorly, each of them fusing separately with stria 2 and joined stria reaching lateral gutter); parascutellar striola and striae 1-8 impunctate or finely punctate, shallow (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="5E16EA90D0D3E5E67E02A748FDCF0498" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="20B05BE8803D687C3535B92E6E48E504" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group), finely to moderately punctate and impressed (most species), or coarsely punctate and impressed (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanesis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanesis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanesis</emphasis>
<taxonomicName id="3E31E45C516AB41E6F986610C1FCD53B" class="Insecta" kingdom="Animalia" lsidName="P. vanesis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanesis">
<emphasis id="341E39552A26B5819ABC6332B0C78A68" italics="true" pageId="0" pageNumber="71">P. vanesis</emphasis>
</taxonomicName>
sp. nov.). Intervals subconvex (most species) to nearly flat (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="2627CC9D830DECFCD0DE6E4016FFCC53" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="D1393BA447F60FE34444B6D3C450E0B0" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group). Metathoracic wings reduced, scales-like, or absent.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Abdomen</emphasis>
<emphasis id="145F5FCA0465DFC4E8783229D9A7629F" bold="true" pageId="0" pageNumber="71">
<emphasis id="399AB0FDDE86D161C9E0F947A1CB36D3" italics="true" pageId="0" pageNumber="71">Abdomen</emphasis>
.
</emphasis>
Ventrites 2 and 3 coalesced, suture between them mostly present, rarely lacking.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="DF630D74CCA8394FAB2B48FA23DF3FA6" bold="true" pageId="0" pageNumber="71">
<emphasis id="83CF3CB5F535317C275CFDE5F36433FB" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Moderately to considerably long and slender. Tarsomeres 1-5 convex and smooth (most species; Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6D, E</figureCitation>
<figureCitation id="07BA3883285D279FD7DEFA2CEA8DC76F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6D, E</figureCitation>
) or partly flattened and slightly longitudinally grooved (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="2EE45A5CA49703C86E381129DB7888C7" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="4C0989D16E2B73497D6F99D099B46078" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="95002C8B1D59D764256A02533CF8D67A" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="81AB17BFE251546613C979053E212EAC" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
; Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6A-C</figureCitation>
<figureCitation id="4984547FEF366119AB49A735E0278DFD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6A-C</figureCitation>
) on dorsal side, with two rows of setae on ventral side; male protarsomeres 1-3 distinctly dilated, with biseriate adhesive vestiture ventrally; mesotarsomeres 1-3 and metatarsomeres 1-3 laterally grooved or ungrooved.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="D480A09AB4BF342A2B50A2A83B726BBC" bold="true" pageId="0" pageNumber="71">
<emphasis id="6AC58E524B2DF92A76579B559EA92BCA" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX suboval, with proximal margin not or slightly asymmetrical (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7A-M</figureCitation>
<figureCitation id="FC7EB4C94C97BF9CE53A6E48C5C8BD11" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7A-M</figureCitation>
). Median lobe of aedeagus in lateral view curved ventrally, with basal bulb protruding dorsally and having a well-developed, thick aileron and apex straight or turned up (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8A-I</figureCitation>
<figureCitation id="603AB599B44AFB3FCD4EC5D568B9C0E3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8A-I</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B-H</figureCitation>
<figureCitation id="1DEF965B049544D8B18C12A244C761B3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B-H</figureCitation>
); same in ventral view straight, subcylindrical, with long shaft (part restricted between basal bulb and apical lamella) and apical lamella more or less symmetrical (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A-I</figureCitation>
<figureCitation id="E4F823D6ADC477E080F0CA730E3020CE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A-I</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11A-G</figureCitation>
<figureCitation id="21E7C94474EC49DE6CB84C6D26F481F9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11A-G</figureCitation>
). Internal sac well-differentiated, containing two sclerites, a less sclerotized (reticulate), globular dorsal sclerite and a more sclerotized, elongate ventral sclerite (respectively, light grey colored and dark grey colored on Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8</figureCitation>
<figureCitation id="0E836258203C8E76DA576A8CB350B61B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8</figureCitation>
-
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11</figureCitation>
<figureCitation id="E353F304DDEEF963D630005E8794A84C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11</figureCitation>
); in lateral view dorsal sclerite rounded (often a denser, saddle-shaped structure at proximal position is distinguished), and ventral sclerite is drop-shaped, both sclerites hardly distinctive to each other in lateral position than in ventral position; in ventral view dorsal sclerite is laterally rounded (forming one or two short curves from left side and one long curve from right side), and ventral sclerite is long and more or less narrow, with distal end straight or nearly straight (taxa of most species groups) or curved to left (&quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
<taxonomicName id="391F18C1B6DCD96A46F9D1AFB832D4CF" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis id="8787DC30B21555E6357E2F69762F1AF3" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
</taxonomicName>
&quot; and &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="6AFA9F6F26CD77CD06A4CB43A4147538" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="E755B9E1A700006C2FAA9903B0BA8C90" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; groups) and a position within internal sac parallel to longitudinal axis of median lobe (most species) or oblique to it (&quot;
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<taxonomicName id="5F512AB1654D0A340D6B213583D0617B" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="56646A8A8A4F1BEC74268DAC5A2AF1F8" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
&quot; group). Right paramere short, thin, more or less concave ventrally (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12A-N</figureCitation>
<figureCitation id="DA10059B5EE88DD07253CA0E9BE957A2" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12A-N</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="D7800A4E7579B5BBB96E0E4319087091" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13A-F</figureCitation>
<figureCitation id="4516B81B56ACA937B4B465DA67C2DC08" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13A-F</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14A-C, E, F</figureCitation>
<figureCitation id="BA3BB99711E01FDF8C77477983976B2D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14A-C, E, F</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15A-F</figureCitation>
<figureCitation id="F0D14A91B5C24197534D2E2342CE2963" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15A-F</figureCitation>
). Basal gonocoxite large to very large (five to ten times larger than apical gonocoxite), unsetose. Apical gonocoxite small, short, with narrow, falcate apex (mostly pointed, rarely blunt or rounded), one (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="C3114C713CE25904401FFC18FF4EA2E4" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="902701A8B69F039993E63F6EB8D0DD4F" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group, except for
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
<taxonomicName id="34A657E5EFB844079DCB1B5F57C80F3C" class="Insecta" kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis id="FBCBCC2D0AEC742DAC1B1BBF18BEA8C9" italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov.) or two large dorsolateral ensiform setae (taxa of &quot;
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<taxonomicName id="455FBBCFF99A2A2268205B96C055BA3C" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="CDBF6D4509A7508E7AFD28C80A9DDC7F" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
&quot;, &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<taxonomicName id="36FEE03D4676BD6E0F6AC6F6C2B9F95A" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="206E365BD05F155444F3FFD53B81E8B6" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
<taxonomicName id="C49CEF04D8E4A5160F3F4F4B46BAF5D9" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis id="FE732D6DBBF4373454A591D6C0DA4D41" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
</taxonomicName>
&quot;, &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
<taxonomicName id="D5A47663234CAE2AAD2AF8D3273CD625" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis id="E8365FB9E7DA16BDF36C09DB219D13C3" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
</taxonomicName>
&quot;, &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="DE7F73C618E588253231587A6FDFAAF3" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="842F31380C547F8ABC48AC0EEFD6558F" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; group plus
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
<taxonomicName id="0B0D8109B652036E1F9B86AFA7D48AE7" class="Insecta" kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis id="1BD1A5912F66D7F3BA8B5F1BC0B90A54" italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov.), one large dorsomedial ensiform seta, and two long and fine apical nematiform setae in sensory pit (nematiform setae not found only in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="CF40E3280B0396216BFAF6D3DE84DD2B" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="6AA75AD304CCFFB365EA7B786FA495DD" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="31017DF2DE7669D2C55B0A388F75148E" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="403236BCED69C9EE8112321343C331D8" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
). Bursa copulatrix small to moderately large, subconical or subquadrate, one-chambered in most species, two-chambered in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="BB119D742EE9437458781B6F6384BADB" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="910ECF901E063F1F38BC7E980B9E40DF" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. Spermatheca relatively long, moderately differentiated, formed by seminal canal and receptaculum of similar length, seminal canal narrower, somewhat curved, receptaculum wide and straight; spermathecal gland very large, subelongate to round, with well-differentiated atrium; spermathecal canal short, connected in apical second fourth of spermatheca, either in basal third (taxa of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="A2FABD6888E2C51C1C4E0AD4371ACCB9" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="FFADBF4DF79FB09A3E0A3BC56FFFD815" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group, except
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
<taxonomicName id="8754E579B799723AC73317C9F6723BDF" class="Insecta" kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis id="740EF1A8EBC1B8C3F2BE7C267D4CAE8D" italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus</emphasis>
<taxonomicName id="D7C449F7389927ED61D13A2ACA10776F" class="Insecta" kingdom="Animalia" lsidName="P. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis id="BF606FA1BB41E4081E214517800A3A5C" italics="true" pageId="0" pageNumber="71">P. insignitus</emphasis>
</taxonomicName>
) or medial third of receptaculum (two foregoing species plus taxa of remaining species groups).
</paragraph>
<caption doi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 1.</emphasis>
<caption id="818B9CC7226A23CD4C177432D0803493" doi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71" start="Figure 1" startId="F1">
<paragraph id="50BDB8C53E08E4F28BE48CDBFD4B66AD" pageId="0" pageNumber="71">
<emphasis id="584A8A98670729FE191C615B18E9B841" bold="true" pageId="0" pageNumber="71">Figure 1.</emphasis>
Dorsal habitus.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
<emphasis id="65BBF48AAA405A30741233584D0A662B" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="68A8E5FB292D97B074221CC0A130066B" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<emphasis id="7B539413B8C68561CE3F61B8DD3521C7" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="6369FCF90E8AE2947CC7D94840EBB273" class="Insecta" kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken id="EC9CFED77866814756C077D9F69CA8EE" originalValue="Mařan">Maran</normalizedToken>
, 1940, lectotype and its labels;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
<emphasis id="9DE2C5CE7C0A704AE669A4F2B78754C1" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="F84AC5E86C024CD16DCC880DE57EC0DF" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
<emphasis id="F220E1ECE742A0647F4ED5BD6836A366" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="E383893CBCBCECE5D2ECE51E7828AAC4" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
<emphasis id="99EA643E4092944F739313E233FE464F" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="20E62B2C2C1D127E89B7D7762C6BF54E" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
<emphasis id="A7737BC407888ADF7AF9F41D9DD4DA0B" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="7AE4346D8A69864005173B88B1903DB2" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., holotype. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="71">
<paragraph id="D51CF41F08152B5CA5E7F5B2548C1808" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<emphasis id="1E417512160999CD3DC6B7EB9BD0BC85" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="A3A9E9CE4A1FAC9C0918C4A83D726E42" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="C0E2271810FCE650D4756E64515973B2" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
&quot; species group
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
<paragraph id="CF6780A2ECB4472B89C481DC8A8A1733" pageId="0" pageNumber="71">
<emphasis id="C37363FB90C76684AC7659D5C54971BA" bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
Representatives of this group share one trait, the ventral sclerite of internal sac obliquely situated with respect to the main axis of the median lobe in the ventral view and distinctly bent in middle in lateral view (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A, B</figureCitation>
<figureCitation id="EDB5F7540E57A7CCDC5EF36CBF3415D7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A, B</figureCitation>
). Species from the other groups of
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="3DBB6D8B5CDED079EBE3FA2D9CFCA489" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="F7647214373EC95A9CE72D3A86166183" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
have a ventral sclerite parallel to the main axis of the median lobe in ventral view and straight in the lateral view (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10C-I</figureCitation>
<figureCitation id="3FEE3CB43FB666D4C0C6B08ACB4C77EB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10C-I</figureCitation>
). This trait divides the subgenus into two groups that may separate, one including the taxa of the &quot;
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<taxonomicName id="3D0F672543FB253B90337292E12292F7" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="9B84F40319303B01CB780A26966ED4B3" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
&quot; group, and another - taxa of the remaining species groups.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
<paragraph id="BCEE884C31456CA1922E44BA77050D9E" pageId="0" pageNumber="71">
<emphasis id="E00BA59A1D5D316A5E1EB27C2052EE6B" bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
The group includes
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<taxonomicName id="E9110F16800BF3FA08BF0DC1C101B9B4" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="7A3D25BFBFC108390D63E24C38F24F2B" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
from Southwest Turkey and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<taxonomicName id="9259B9D1986DA065B8FF3F555C1FE0CA" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="CC7AFFCCEAB97FC121DFF9CBD661F1C9" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
sp. nov. from Kalymnos Island (Greece, Dodecanese; see Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
<figureCitation id="59720F76CC687755C09CA22D8D322024" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
). The first species inhabits high altitudes in the western Toros Range, whereas the second one is a dweller of Mediterranean scrub habitat at lower altitudes.
</paragraph>
</subSubSection>

392
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@ -1,339 +1,339 @@
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12.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3A</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5G</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7H</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9B</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11A</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12I</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
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<figureCitation id="E316441730D03DAF916EA177554EE703" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3A</figureCitation>
<figureCitation id="B5CDA6024E00C5AAA5CB91587114BBD5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5G</figureCitation>
<figureCitation id="CD1F9DFD06EE3090BDB8D9EE1D905018" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7H</figureCitation>
<figureCitation id="21D2CE41A0B790DABA110DA460F3B2A8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9B</figureCitation>
<figureCitation id="33A47440BC29C4557CCEA52E9F0862BB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11A</figureCitation>
<figureCitation id="C67C8EB52FB8425020F7670B6A353380" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12I</figureCitation>
<figureCitation id="755F5B8C8D70A074F6D5CCA9605D131E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">&quot;Afghanistan, Habatah, 1.300 m&quot;.</paragraph>
<subSubSection id="4A776560F78F9A5FED402612D922FF4E" pageId="0" pageNumber="71" type="type locality">
<paragraph id="7893DC76B5EA26309222C525AE18D3E0" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="A7E0C36F7557668F3D8DC9BD10EC09D8" pageId="0" pageNumber="71">&quot;Afghanistan, Habatah, 1.300 m&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="04DC8C2339B118F54BF040463694EFFA" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="65B51D47D230752E856124A399FD5FF6" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="678CDDF40F3C515BF0EE8368BBE17589" pageId="0" pageNumber="71">
The precise location remains unestablished as far as several possibilities for the name
<normalizedToken originalValue="“Habatah”">&quot;Habatah&quot;</normalizedToken>
<normalizedToken id="F166CA41B381E04D09120F760BE08B9A" originalValue="“Habatah”">&quot;Habatah&quot;</normalizedToken>
exist: Haybatay Ghar;
<normalizedToken originalValue="Haība">Haība</normalizedToken>
<normalizedToken id="310719F417BAA1174D9BC319A7F2FA20" originalValue="Haība">Haība</normalizedToken>
ṯaī Ghun
<normalizedToken originalValue="ḏêy">ḏey</normalizedToken>
<normalizedToken id="7F6E2DFCAACB9C8777C74A07FE0EBD5C" originalValue="ḏêy">ḏey</normalizedToken>
[= Haibatai Ghundey]; and Haybati. Given the altitude of 1300 m a.s. l. pointed on the label, a quite possible location may be the upper course of Gomal River, Afghan Province of Paktika (see Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="5A63500E0FDFC3FD5C7A63C590AB5737" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="2F9FF735BC836BBFD29E5CB34096B287" pageId="0" pageNumber="71" type="type material">
<paragraph id="6465C5BFEBACFB07117AB827796E27DF" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="0E440DD3BCCDA392D68DCF79549F845D" pageId="0" pageNumber="71">
<emphasis id="C5F000C06C6F84977CE8DC2FBF7E5258" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂,
<normalizedToken originalValue="‘Afghanistan’">'Afghanistan'</normalizedToken>
<normalizedToken id="C75972E11D4735EE2C171AFD71C95594" originalValue="‘Afghanistan’">'Afghanistan'</normalizedToken>
[w, p] // Habatah / 1.300m 17.6.1964 / leg. Kullman [w, h] // genus /? / det. Ing.
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<normalizedToken id="21575EC5DD1D251078B77992D2472727" originalValue="Jedlička">Jedlicka</normalizedToken>
[w, h&amp;p] // Collectio /
<normalizedToken originalValue="Moravské">Moravske</normalizedToken>
<normalizedToken id="BF3388A54DD10BC7F2098254C39071DB" originalValue="Moravské">Moravske</normalizedToken>
museum, / Brno [w, p]' (MMBC).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 1 specimen. TGE: 1♂.</paragraph>
<paragraph id="10C077F950D4D353D17DEDC962EE70F7" pageId="0" pageNumber="71">TME: 1 specimen. TGE: 1♂.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="6A34EB3D361998AEB795ADA3414F1335" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="CEEBEC77D2D0A78066D356499B355737" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="9DB6D3836299A295BA68F27A12D18108" pageId="0" pageNumber="71">
This species is distinguished from all other members of the subgenus by: (1) yellow-brown color of body; (2) ventral surface largely impunctate; (3) pronotum with posterior angles not projecting laterally and basal bead complete; (4) metepisternum as long as wide; and (5) ventral sclerite of median lobe in lateral view not appreciably broadened distally (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B</figureCitation>
<figureCitation id="55D6A185754A6462BB59B91F732F9F42" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="6553969C61B2E22C0E27EF4AAE9AD8BE" pageId="0" pageNumber="71" type="description">
<paragraph id="702F205CD0FEC6C72AA72239A38E06F5" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="5367419F9F5887B996A22EF2E35689FC" pageId="0" pageNumber="71">
<emphasis id="CC0A704295878DB1151BED4130C63D36" bold="true" pageId="0" pageNumber="71">
<emphasis id="B7D209CE6E9067AA5D517C3D10BF7619" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of small size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="EC8C746038316A58415A4D1760131FEA" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="5A04EC4033B0E62775D61E8D916100B9" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
(BL: 6.80 mm; BW: 2.25 mm), with oblong, moderately convex body (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3A</figureCitation>
<figureCitation id="2ED151581A56B06CE962AD0B447DB681" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3A</figureCitation>
). Measurements and ratios. See Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
<tableCitation id="961C50100BE8CAC63C754ACCEAC6B392" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="51D6525D8D89DCEB1436000428707BC2" bold="true" pageId="0" pageNumber="71">
<emphasis id="3DC7A5010EAD615F70A02733FEB94B8F" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body dorsally and ventrally, including appendages yellow-brown. Head, pronotum and ventral surface moderately shiny, elytra less shiny than dorsal surface of head and pronotum.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="D81F4696E49D35DFA54836526CFB47E8" bold="true" pageId="0" pageNumber="71">
<emphasis id="B08C0A4241755BFD37D60B0DF443C95A" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum with isodiametric sculpticells on posterolateral parts and slightly transverse ones on anterior third, with disc having scarcely visible microsculpture. Ventral surface with scarcely-visible regular isodiametric (proepisternum, ventrites laterally) or slightly transverse sculpticells (metacoxae, ventrites medially), or microsculpture not apparent (remaining part). Pronotum almost impunctate, with a few large punctures near posterior angles and several longitudinal wrinkles on anterior transverse impression. Ventral surface, including mesepisternum, metepisternum and abdominal ventrites impunctate; prosternum medially, proepisternum and metasternum laterally with shallow punctures; abdominal ventrites 1-2 wrinkled submedially.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="405F4484BC418CB00A024AF0200F2AE3" bold="true" pageId="0" pageNumber="71">
<emphasis id="82F841CF7519B2DD1FEC2052C1A92E9F" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
More than one-third narrower than pronotum wide (PW/HW= 1.40). Antennae long, with last three antennomeres exceeding base of pronotum. Eyes slightly convex. Labrum subrectangular, with anterior margin straight. Frontal furrows short, shallow. Paraorbital sulci shallow, narrow, ending before level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="92F14AEDAF3F1F16D4E925130EB66C9F" bold="true" pageId="0" pageNumber="71">
<emphasis id="E5C4C74C32838D15BA209C837EF2AA95" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about a quarter wider than long (PW/PL= 1.24), widest at second quarter. Anterior transverse impressions distinct, posterior one well-impressed laterally reaching basal foveae. Sides not sinuate, convex anteriorly, nearly straight posteriorly; anterior bead present laterally, absent in medial 1/10; lateral and basal beads present throughout. Metepisternum nearly as long as wide, MA/MI= 0.95-0.97.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="7CE521BFE47DE1B4AB9335F918406D35" bold="true" pageId="0" pageNumber="71">
<emphasis id="888A704AD167DE60E16371FDD36E8E73" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and two thirds as long as elytra width (EL/EW= 1.63), two and three quarters as long as pronotum (EL/PL= 2.78), and one third as wide as pronotum (EW/PW= 1.36), with widest point at medial third. Striae moderately punctate and impressed; parascutellar striole nearly connected with stria 1; base of stria 1 very short, joining stria 2 and reaching parascutellar pore; striole and basal portion of stria 1 slightly impressed, remaining part of stria 1 and other striae well-impressed. Interval 3 with three discal setiferous punctures each side. Umbilicate setiferous punctures 16 each side.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="859C83FE4BA1E553E47A6C01E76725F2" bold="true" pageId="0" pageNumber="71">
<emphasis id="D58DF215E023500D56EA3B1113B09B57" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Anterior side of right mesofemur ventrally with five (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5G</figureCitation>
<figureCitation id="2C02DBB29D74C7CC719953FE76624C51" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5G</figureCitation>
), left one with four setiferous punctures. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="F39CB60A91DE8616F21D57D91502CA89" bold="true" pageId="0" pageNumber="71">
<emphasis id="F84F18DB74FA6184F4DF81D702EF57CD" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX small, oval, with proximal margin nearly symmetrical (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7H</figureCitation>
<figureCitation id="3B00A56627D41156E0D577DEFB43F123" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7H</figureCitation>
). Median lobe of aedeagus in lateral view slender, with narrow basal bulb, broad shaft with proximal part slightly constricted, and short, straight apex (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B</figureCitation>
<figureCitation id="CF696C661634209D1B7E99695752EBA0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B</figureCitation>
); median lobe in ventral view straight, about 4 times as long as wide; apical lamella (dorsal view) short, symmetrical, rounded at tip, with sides straight to very slightly convex. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B</figureCitation>
<figureCitation id="0A80D5A58AD89FE7B4EC08758640E022" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9B</figureCitation>
) with ventral sclerite broad and nearly equally wide in its length; same in ventral view (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11A</figureCitation>
<figureCitation id="14CEF030B7A2C2C49D0090A78D42655E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11A</figureCitation>
), with dorsal sclerite seemingly separated from ventral sclerite, and ventral sclerite narrow, widened distally. Right paramere fine, moderately concave ventrally (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12I</figureCitation>
<figureCitation id="29803E5B8A0E3C9E3C2740E6A383E1D0" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12I</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="681D84FC63705E9F7B3E0408AFD3917D" bold="true" pageId="0" pageNumber="71">
<emphasis id="242BE19B73CEAEB35F70C55C498CB5A2" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
.
</emphasis>
Unknown.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 12.</emphasis>
<caption id="4EB60FD7FDCC4A898663198BE1896B68" doi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71" start="Figure 12" startId="F12">
<paragraph id="0B2B6F7FC55AEDE46023B62EE14DC6B7" pageId="0" pageNumber="71">
<emphasis id="3DAD7D617A46C918364493DBE9FAE959" bold="true" pageId="0" pageNumber="71">Figure 12.</emphasis>
Right paramere, external face.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
<emphasis id="5CB6B04EB161C4963C355C29C47CBEF1" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="1B7FBB4DB692C5E5594BD52354E61770" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<emphasis id="04A8F853B8BADD7F9B8C778D41B11E46" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="A61B907C9690434ABC7B9C9A0F7D48CD" class="Insecta" kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken id="05842BB23913B03CFF6F46F0246F9B2D" originalValue="Mařan">Maran</normalizedToken>
, 1940, lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
<emphasis id="CEBDCF4D1FF45A1F0BFEE3EB174B06A6" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="7D07A01FB0A4649E024B88C37C8A6448" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
<emphasis id="E96D8C282B4E2E7C1AC8961869E09EC4" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="B7E43E81F4FCD494AEA2C09CA5AC7B1A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
<emphasis id="7340A44490393E932A27B4150F2A6C19" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="BC15D57FBA0C7C6A5F84E1E22E302394" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
<emphasis id="61BF4893E71DA81686C24C0AB4F1E8E9" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="F40CBBE50376011E97A958B2A627340A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
<emphasis id="0F2F976C33F9C4965954B432F112581D" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="5324347B43A458DADDC8E1384E000675" class="Insecta" kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
<emphasis id="9481C092559FA5AA3F34FFD00486D80C" bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName id="47B95990DE161379FB563AC4AEE8734A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, male specimen, Tochal, Tehran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="E.">P. (E.) afghanistanicus</taxonomicName>
<emphasis id="52280D39CA891AF525728DA06E8AA7D3" bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName id="627695FBF7ADC72FDEE07406E745AC45" class="Insecta" kingdom="Animalia" lsidName="P. (E.) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="E.">P. (E.) afghanistanicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
<emphasis id="4FCEBA1D2194481C5BBF346F884BACD6" bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName id="A70890F8B16890C3E73B804ACDE94213" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia;
<emphasis bold="true" pageId="0" pageNumber="71">K.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="2FA51E4FC15FBCE12632F5E219EEEA25" bold="true" pageId="0" pageNumber="71">K.</emphasis>
<taxonomicName id="AAF2979F24A411DB2698CAA61AF118AD" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel;
<emphasis bold="true" pageId="0" pageNumber="71">L.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
<emphasis id="E2DBFEE2E4F17359DD2C8B82B40A22EF" bold="true" pageId="0" pageNumber="71">L.</emphasis>
<taxonomicName id="382FEC92654D7988CC165485E84AB7EF" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">M.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="82E131210DAF833F905D3C10E1BAD565" bold="true" pageId="0" pageNumber="71">M.</emphasis>
<taxonomicName id="991B053751B5F863A4BC474B42B13BD3" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">N.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="D8BD64EC240FA0138FF1CE1CA5956948" bold="true" pageId="0" pageNumber="71">N.</emphasis>
<taxonomicName id="9B736A82C8848F9EE63E574032A81FA0" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), male paralectotype,
<normalizedToken originalValue="“Syria”">&quot;Syria&quot;</normalizedToken>
<normalizedToken id="82F2F6DB0AFF16490A9DA2D0E061202A" originalValue="“Syria”">&quot;Syria&quot;</normalizedToken>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="4D28A626000FA65056055024FF7D33FB" pageId="0" pageNumber="71" type="distribution">
<paragraph id="ABFB6803631A7C6500A1B801CAF875A4" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="E0BBCF980D0357A6B5D78755B9ACBD81" pageId="0" pageNumber="71">
Afghanistan (?
<normalizedToken originalValue="“Habatah”">&quot;Habatah&quot;</normalizedToken>
<normalizedToken id="2E382714B39313A687C1E5AA07A0E29B" originalValue="“Habatah”">&quot;Habatah&quot;</normalizedToken>
); Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="B8B36E03A858ABBEBA8703B37025BEA5" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="43847FB1E6BD0FB181044B85C860BF60" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<emphasis id="8B47A2DF1935FF39EAC22F59E36E190E" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="C0006C41CED3F3777CE3E65FDEE4FEB2" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="0CF58AD4E7774A8DB18A650A0DF98978" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; species group
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
<paragraph id="92D7380F2AB4963F813B610F3513A9C8" pageId="0" pageNumber="71">
<emphasis id="DF956B8E6C487B8B65AF71987EA50EA4" bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
Species are distinct from those of the other
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="F1798B0077ED3D80433E715D9BD9F9FB" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="62C313B1482AB7130FAF21F5DFDC0DAA" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
species groups by: (1) dorsal surface of body with complete or nearly complete microreticulation of isodiametric sculpticells, with transverse sculpticells lacking or highly restricted; (2) basal foveae of pronotum shallowly impressed, foveae and adjacent lateral areas impunctate or scarcely punctate; (3) basal bead of pronotum reduced to absent in basal 1/3 to 1/2; (4) elytral striae scarcely impressed, impunctate or slightly punctate, and elytral intervals rather flat; (5) mesocoxa with two to four long lateral setae, rarely one seta (polysetose state of character shared only by taxa of &quot;
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<taxonomicName id="AF3759DBF8CA8586A1D1BB05E17BF2C6" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="EFD339AA5C04A0E50018E7262936A3F9" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
&quot; group).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="23BB922639B538374C6754082489C109" pageId="0" pageNumber="71">
In addition, the species have anterior side of the mesofemur ventrally with five to seven (rarely four) setiferous punctures. In addition, all taxa save
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
<taxonomicName id="0853FB3E945AB31539A3A343E6506DD4" class="Insecta" kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis id="C211CBB47A68BB6905BD6E7F90B9BB66" italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov. share apical gonocoxite with one dorsomedial ensiform seta.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
<paragraph id="8A2DD8B310EB70E5A5528AD4472D6581" pageId="0" pageNumber="71">
<emphasis id="65DD984A66AFA623940049843A47A82F" bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
This group includes seven species,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="ED88895B187665512965903AD8B3B75F" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="BA5BBC1E6DCC5ECA196CD05404B51A9D" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
(Reiche &amp; Saulcy, 1855),
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="F0839E5809245F5233AADE6175D38DC5" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="62305CCEFA72CD15E9747991D7A713CB" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
Karsch, 1881,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus</emphasis>
<taxonomicName id="234AA721CC18B541C03308ADEE9C1BE9" class="Insecta" kingdom="Animalia" lsidName="P. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis id="83887E4D74E2F4A54DC5A9C3A5896A95" italics="true" pageId="0" pageNumber="71">P. insignitus</emphasis>
</taxonomicName>
Bedel, 1902,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="3724A44FCE2EF457C4817CF83BDAFFAF" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="5F2C37024692FAF0F164578A3F5D3698" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
<taxonomicName id="D34DCFC4638A858176ED573613D1BE9B" class="Insecta" kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis id="ED005E479945397AFA76CE85A77BCD61" italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="DF6FEE9C17A6BDA186CCA086F0E05832" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="79429A1D17D7CFD40115A61FD4F6328E" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="931B8421CA5936D469807F45B533B15C" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="66BE2DE0DC0A5EA27B338966740940EF" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. (Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
<figureCitation id="961009F3094C6B961D694FDA237D0CA8" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
<figureCitation id="CA4465BD19E2F2B23C99D81E6B6558C7" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
). Most probably, the Macaronesian species
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. alticola" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="71">P. alticola</emphasis>
<taxonomicName id="908253D3A6C2EBC2EB558605CFD8E0CE" class="Insecta" kingdom="Animalia" lsidName="P. alticola" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="C6EEDE2204B92517264CB876D7AF8691" italics="true" pageId="0" pageNumber="71">P. alticola</emphasis>
</taxonomicName>
Wollaston, 1864 and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lancerottensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lancerottensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. lancerottensis</emphasis>
<taxonomicName id="7C73DA662C0CA5C5BBF37CBEE7C34B13" class="Insecta" kingdom="Animalia" lsidName="P. lancerottensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lancerottensis">
<emphasis id="9ADC05E8154A7F0BB7D769B63873BFA9" italics="true" pageId="0" pageNumber="71">P. lancerottensis</emphasis>
</taxonomicName>
Israelson, 1990 also belong to this group (see
<bibRefCitation author="Israelson, G" journalOrPublisher="Vieraea" pageId="0" pageNumber="71" pagination="165 - 167" refId="B24" refString="Israelson, G, 1990. A Platyderus Stephens (Col., Carabidae) from Lanzarote, Canary Islands. Vieraea 19: 165 - 167" title="A Platyderus Stephens (Col., Carabidae) from Lanzarote, Canary Islands." volume="19" year="1990">Israelson 1990</bibRefCitation>
<bibRefCitation id="E843B910262F0C5D350866A35643D51E" author="Israelson, G" journalOrPublisher="Vieraea" pageId="0" pageNumber="71" pagination="165 - 167" refId="B24" refString="Israelson, G, 1990. A Platyderus Stephens (Col., Carabidae) from Lanzarote, Canary Islands. Vieraea 19: 165 - 167" title="A Platyderus Stephens (Col., Carabidae) from Lanzarote, Canary Islands." volume="19" year="1990">Israelson 1990</bibRefCitation>
: 166, fig. 4).
</paragraph>
</subSubSection>

102
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<paragraph id="576C0FE8E61C8A4D3B3DFFB0AFC042BA" pageId="13" pageNumber="14">Taxon classification Animalia Pulmonata Plectopylidae</paragraph>
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@ -103,7 +103,7 @@ Godwin-Austen new synonym: Proceedings of the Zoological Society of London: 610,
1878
<taxonomicName id="CF293AEF39A28F16F8BC8B75EB9403EA" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Plectopylis) serica" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="serica" subGenus="Plectopylis">Helix (Plectopylis) serica</taxonomicName>
, - Nevill: Hand list of
<taxonomicName id="2ADB0ABCE0C2CB15D07B29AEF0ABBE49" lsidName="Helicoidea" pageId="13" pageNumber="14" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
<taxonomicName id="2ADB0ABCE0C2CB15D07B29AEF0ABBE49" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helicoidea" pageId="13" pageNumber="14" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
in the Indian Museum, Calcutta...: 71. [&quot;Hengdan Peak and Burrail&quot;].
</paragraph>
<paragraph id="7D6B32C8EBEF8913D1BAE6C21E1F0920" pageId="13" pageNumber="14">
@ -199,13 +199,13 @@ in the Indian Museum, Calcutta...: 71. [&quot;Hengdan Peak and Burrail&quot;].
<paragraph id="D18A8F43C3D75407A3B388C133BB98C1" pageId="13" pageNumber="14">Types.</paragraph>
<paragraph id="6B7C9AF318E7F130762F68342B5D99E5" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Khunho, H.S. Naga Hills, leg. Godwin-Austen, NHMUK 1903.7.1.741 (8 syntypes of
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<taxonomicName id="EDE0A57005237547D0F59BBAA07378E6" genus="Helicoidea" lsidName="Helicoidea serica" pageId="13" pageNumber="14" rank="species" species="serica">serica</taxonomicName>
, Figure 5B); Hengdan P., Naga Hills, leg. Godwin-Austen, NHMUK
<pageBreakToken id="C2D898B49850B7DC5F853321322FB811" pageId="14" pageNumber="15" start="start">1903.7.1.744</pageBreakToken>
(6 syntypes of
<taxonomicName id="598339ED5C1E8FB6A815F3BD99AD81F0" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea serica" order="Cycloneritida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="serica">serica</taxonomicName>
<taxonomicName id="598339ED5C1E8FB6A815F3BD99AD81F0" genus="Helicoidea" lsidName="Helicoidea serica" pageId="14" pageNumber="15" rank="species" species="serica">serica</taxonomicName>
, Figure 5C); Munipur Hills, head of the Ihang valley, Munipur, leg. Godwin-Austen, NHMUK 1903.7.1.742. (6 syntypes of
<taxonomicName id="C5A9289769E2AF99AD684EC57FE41EAE" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea munipurensis" order="Cycloneritida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="munipurensis">munipurensis</taxonomicName>
<taxonomicName id="C5A9289769E2AF99AD684EC57FE41EAE" genus="Helicoidea" lsidName="Helicoidea munipurensis" pageId="14" pageNumber="15" rank="species" species="munipurensis">munipurensis</taxonomicName>
, Figure 5D).
</paragraph>
</subSubSection>
@ -213,7 +213,7 @@ Khunho, H.S. Naga Hills, leg. Godwin-Austen, NHMUK 1903.7.1.741 (8 syntypes of
<paragraph id="0F3166C295A0B7710AEC94F5E22FED23" pageId="14" pageNumber="15">Additional material examined.</paragraph>
<paragraph id="33D5DCEA6FB87FB8E3E3B099802578F6" pageId="14" pageNumber="15">
Naga Hills, coll. Godwin-Austen, NHMUK 1903.7.1.743/4 (under the name
<taxonomicName id="F4977AB4EB49EA1631FDA0B1EC0BC2B8" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea munipurensis" order="Cycloneritida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="munipurensis">munipurensis</taxonomicName>
<taxonomicName id="F4977AB4EB49EA1631FDA0B1EC0BC2B8" genus="Helicoidea" lsidName="Helicoidea munipurensis" pageId="14" pageNumber="15" rank="species" species="munipurensis">munipurensis</taxonomicName>
); Japvo Peak, Nr. Kohima, Naga Hills, NHMUK 20150128/8; Lhota Naga, coll. Godwin-Austen, NHMUK 1903.7.1.745/6; no locality, leg. Maxwell, coll. Godwin-Austen, NHMUK 20150129/5; India, Hengdan Peak, NHMUK 1891.3.17.356-357/2; India, NHMUK 1874.4.26.2/2; Khasi Hills, coll. W. Blanford, NHMUK 1906.2.2.360/2.
</paragraph>
</subSubSection>
@ -300,7 +300,7 @@ described
and
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in the same publication. He did not mention the differences between the two species. According to the illustrations and the identification key in the original description, the upper end of the lamella in
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<taxonomicName id="6B43EF291F2331ECFA248BC7E6016B17" genus="Helicoidea" lsidName="Helicoidea munipurensis" pageId="15" pageNumber="16" rank="species" species="munipurensis">munipurensis</taxonomicName>
is more elongated anteriorly than that of
<taxonomicName id="9C36D0414A773AB16E4145C5BAC57E47" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix serica" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="serica">Helix serica</taxonomicName>
. Two shells of
@ -310,11 +310,11 @@ were opened from the Hengdan sample, and both had an anteriorly elongated plica.
specimens. In the Khunho sample four shells were opened, three having no or very slight upper elongation, but one
<pageBreakToken id="932CDA5A65EA9A0C1EC2359B125E1B86" pageId="16" pageNumber="17" start="start">had</pageBreakToken>
an as long plica as in typical
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<taxonomicName id="E82D01E6806E8C0C323A20168CB4E1C0" genus="Helicoidea" lsidName="Helicoidea munipurensis" pageId="16" pageNumber="17" rank="species" species="munipurensis">munipurensis</taxonomicName>
shells. Examining the type specimens of the two species we have not found significant differences. The width of the umbilicus and the height of the spire showed some variability. Therefore we synonymize
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<taxonomicName id="3B58ACFC65FB3F8E91F70F3251A1D4E4" genus="Helicoidea" lsidName="Helicoidea munipurensis" pageId="16" pageNumber="17" rank="species" species="munipurensis">munipurensis</taxonomicName>
with
<taxonomicName id="F90AA3C0CD4FF9E09A1B2FEE870A05B2" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea serica" order="Cycloneritida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="serica">serica</taxonomicName>
<taxonomicName id="F90AA3C0CD4FF9E09A1B2FEE870A05B2" genus="Helicoidea" lsidName="Helicoidea serica" pageId="16" pageNumber="17" rank="species" species="serica">serica</taxonomicName>
. We choose
<taxonomicName id="1057E0693441CC185FD70832D4BF510F" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Plectopylis) serica" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="serica" subGenus="Plectopylis">Helix (Plectopylis) serica</taxonomicName>
to be the valid specific name.

98
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<paragraph id="4574E06676C79894230E90599D9CFAE6" pageId="28" pageNumber="29">Taxon classification Animalia Pulmonata Plectopylidae</paragraph>
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@ -101,7 +101,7 @@ W. Blanford: Journal of the Asiatic Society of Bengal, 39 (2): 17-18, Plate 3, f
1878
<taxonomicName id="79E2ECABB16A8CDF8DECC2074AFE80A4" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Plectopylis) macromphalus" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="macromphalus" subGenus="Plectopylis">Helix (Plectopylis) macromphalus</taxonomicName>
, - Nevill: Hand list of
<taxonomicName id="971876894EFAF9A7BE6332AFEAE59398" lsidName="Helicoidea" pageId="29" pageNumber="30" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
<taxonomicName id="971876894EFAF9A7BE6332AFEAE59398" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helicoidea" pageId="29" pageNumber="30" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
in the Indian Museum, Calcutta...: 71.
</paragraph>
<paragraph id="0E568F16891307569472EDD2DF6B9A55" pageId="29" pageNumber="30">
@ -185,9 +185,9 @@ Gude new synonym: Records of the Indian Museum, 8: 506-507, Plate 41, figs 2
<paragraph id="96CE14E2CA64EE9BB3F258E17CB70012" pageId="29" pageNumber="30">Types.</paragraph>
<paragraph id="72ECF7C1D99444BBE7676950A32DB0E6" pageId="29" pageNumber="30">
Darjiling, coll. W. Blanford, NHMUK 1906.1.1.754. (holotype of
<taxonomicName id="763AC473948BAB9255074F600E76B9F5" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea macromphalus" order="Cycloneritida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="macromphalus">macromphalus</taxonomicName>
<taxonomicName id="763AC473948BAB9255074F600E76B9F5" genus="Helicoidea" lsidName="Helicoidea macromphalus" pageId="29" pageNumber="30" rank="species" species="macromphalus">macromphalus</taxonomicName>
, Figure 23A); Yamne Valley, Abor Hills, leg. C.F.G. Oakes, R.E., NHMUK 1903.7.1.3124. (holotype of
<taxonomicName id="9F58A3FF8612098D8150299210612DD9" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea gregorsoni" order="Cycloneritida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="gregorsoni">gregorsoni</taxonomicName>
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, Figure 23B).
</paragraph>
<caption id="B82433C4E01351344EA7EC6A1E2787CD" pageId="29" pageNumber="30">
@ -197,11 +197,11 @@ Figure 23. Shells of
species. A
<taxonomicName id="E031C335A2AD644F21B185355AD1B268" class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella macromphalus" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="macromphalus">Endothyrella macromphalus</taxonomicName>
(W. Blanford, 1870), NHMUK 1906.1.1.754. (holotype of
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) B
<taxonomicName id="BD3C99F666A58F88D17733F0D12D7957" class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella macromphalus" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="macromphalus">Endothyrella macromphalus</taxonomicName>
, NHMUK 1903.7.1.3124. (holotype of
<taxonomicName id="2E6F6ECFF6EF9DA57BB1A65F069CB5E7" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea gregorsoni" order="Cycloneritida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="gregorsoni">gregorsoni</taxonomicName>
<taxonomicName id="2E6F6ECFF6EF9DA57BB1A65F069CB5E7" genus="Helicoidea" lsidName="Helicoidea gregorsoni" pageId="29" pageNumber="30" rank="species" species="gregorsoni">gregorsoni</taxonomicName>
) C
<taxonomicName id="92CF59CE6B970969AEF77FAD6469AC91" class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella robustistriata" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="robustistriata">Endothyrella robustistriata</taxonomicName>
sp. n., NHMUK 1903.7.1.767. D
@ -255,7 +255,7 @@ has more depressed shells than
. Moreover,
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shells are smooth on the ventral side, whereas most
<taxonomicName id="B44469721B237B2E0076847D0F0925FB" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea blanda" order="Cycloneritida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="blanda">blanda</taxonomicName>
<taxonomicName id="B44469721B237B2E0076847D0F0925FB" genus="Helicoidea" lsidName="Helicoidea blanda" pageId="29" pageNumber="30" rank="species" species="blanda">blanda</taxonomicName>
shells have hairs, or in case of corroded
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specimens, holes which indicate the
@ -295,7 +295,7 @@ is very similar to typical
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specimens. The main difference is that the palatal plicae are not divided
<pageBreakToken id="6020597BFB7F76219171A7DA253B7783" pageId="30" pageNumber="31" start="start">in</pageBreakToken>
<taxonomicName id="78AEB78DCA72B9DFEAEA70AE62C1D2D0" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea gregorsoni" order="Cycloneritida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="gregorsoni">gregorsoni</taxonomicName>
<taxonomicName id="78AEB78DCA72B9DFEAEA70AE62C1D2D0" genus="Helicoidea" lsidName="Helicoidea gregorsoni" pageId="30" pageNumber="31" rank="species" species="gregorsoni">gregorsoni</taxonomicName>
, and the base is less glossy (rather weakly ribbed). In our view these minor difference are not sufficient for species level distinction, especially because
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is a relatively variable species inhabiting wide geographical range. Very little is known about the distribution of specimens having divided or undivided palatal plicae. Therefore, until more information becomes available,

186
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<treatment LSID="urn:lsid:plazi:treatment:477FF92FD6B85C81B29C93316C73A878" httpUri="http://treatment.plazi.org/id/477FF92FD6B85C81B29C93316C73A878" lastPageNumber="71" pageId="0" pageNumber="71">
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<paragraph id="5EABAEEDEF76282593BC19A5C1F38916" pageId="0" pageNumber="71">
16.2.
<taxonomicName LSID="477FF92F-D6B8-5C81-B29C-93316C73A878" authority="Lagar Mascaro, 1978" authorityName="Lagar Mascaro" authorityYear="1978" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) insignitus subsp. presaharensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="Eremoderus" subSpecies="presaharensis">
<taxonomicName id="1F7E16D5B48E87010C88EDC34DF45396" LSID="477FF92F-D6B8-5C81-B29C-93316C73A878" authority="Lagar Mascaro, 1978" authorityName="Lagar Mascaro" authorityYear="1978" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) insignitus subsp. presaharensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="Eremoderus" subSpecies="presaharensis">
Platyderus (Eremoderus) insignitus presaharensis Lagar
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, 1978
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<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">Fig. 18</figureCitation>
<subSubSection id="95B0AED6CE95E3AC0B7FE8539417328C" pageId="0" pageNumber="71" type="description">
<paragraph id="549A6F277B6B4BDBCE67BE5BEEE50D71" pageId="0" pageNumber="71">
<figureCitation id="FC78DFBC476A36E02D25EDE45F691044" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">Fig. 18</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Lagar Mascaro" authorityYear="1978" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus subsp. presaharensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presaharensis">Platyderus insignitus presaharensis</taxonomicName>
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<paragraph id="0F49CB8594D705401FD90847F7B6F624" pageId="0" pageNumber="71">
<taxonomicName id="6C79BD08AD622264DCC0E0BE65EDA3DE" authorityName="Lagar Mascaro" authorityYear="1978" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus subsp. presaharensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presaharensis">Platyderus insignitus presaharensis</taxonomicName>
Lagar
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, 1978: 31 (type locality: &quot;cueva [cave] Kef Aziza, Tazzouguert&quot;, Morocco).
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<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Lagar Mascaro" authorityYear="1978" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus subsp. presahariensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presahariensis">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus insignitus presahariensis</emphasis>
<subSubSection id="EB7BCB3595F3CE79D8A54359C01E372E" pageId="0" pageNumber="71" type="references">
<paragraph id="09192512D617A48AE94A4C42F8D7488E" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="35218CE203607986CA0C621C39062A99" pageId="0" pageNumber="71">
<taxonomicName id="94C6CC2432F565614DA7296C9647DD24" authorityName="Lagar Mascaro" authorityYear="1978" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus subsp. presahariensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presahariensis">
<emphasis id="5406C869EB32A212808B870AF39965B7" italics="true" pageId="0" pageNumber="71">Platyderus insignitus presahariensis</emphasis>
</taxonomicName>
[sic]:
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="759AAB2D84C06593DAE2A8ECECEC7B24" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 756.
</paragraph>
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<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="217162C723741FD1223C2B19DA580ACD" pageId="0" pageNumber="71" type="type material">
<paragraph id="45615E648A5497AAEB4415EFBBF344A4" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="A024B8EA224CFC02E6E7C1FF3F57C50C" pageId="0" pageNumber="71">
Holotype probably in Museu de
<normalizedToken originalValue="Ciències">Ciencies</normalizedToken>
<normalizedToken id="EFCDCF93916548EDA501EC449A205169" originalValue="Ciències">Ciencies</normalizedToken>
Naturals de Barcelona. Not examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">None.</paragraph>
<subSubSection id="815AA4A5F5E4094A40B6020F6C9E5033" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="B640464F774FDC0E423A4AA0931C414D" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="84C5F6B286FA1BCEB1C0D011462AC5F8" pageId="0" pageNumber="71">None.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="AB6278E896B62A5D909DD660BCD3D415" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="F18CBF557CAD8A468659FDDFD22D72D1" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="D19D2DDEE8CFBD05D754B071F793A1A8" pageId="0" pageNumber="71">
Differs from nominotypical subspecies by pronotum sides more parallel to pronotum base and not sinuate before posterior angles, which are right and rounded, somewhat longer antennae, with antennomeres 9-11 exceeding pronotum base, and elytral striae well-impressed (
<bibRefCitation author="Lagar Mascaro, A" journalOrPublisher="Bulleti de la Unio Excursionista de Catalunya" pageId="0" pageNumber="71" pagination="19 - 20" refId="B30" refString="Lagar Mascaro, A, 1978. Un nuevo Platyderus del Marroc. Coleoptera Pterostichidae. Excursionisme. Bulleti de la Unio Excursionista de Catalunya 41: 19 - 20" title="Un nuevo Platyderus del Marroc. Coleoptera Pterostichidae. Excursionisme." volume="41" year="1978">
<bibRefCitation id="2A85E401765D700D3EB7F327A534E29A" author="Lagar Mascaro, A" journalOrPublisher="Bulleti de la Unio Excursionista de Catalunya" pageId="0" pageNumber="71" pagination="19 - 20" refId="B30" refString="Lagar Mascaro, A, 1978. Un nuevo Platyderus del Marroc. Coleoptera Pterostichidae. Excursionisme. Bulleti de la Unio Excursionista de Catalunya 41: 19 - 20" title="Un nuevo Platyderus del Marroc. Coleoptera Pterostichidae. Excursionisme." volume="41" year="1978">
Lagar
<normalizedToken originalValue="Mascaró">Mascaro</normalizedToken>
<normalizedToken id="5C2B310F876F1365AE7CC1D9B4303ABC" originalValue="Mascaró">Mascaro</normalizedToken>
1978
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: 31).
</paragraph>
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<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">The holotype was collected in the cave Kef Aziza, a 4.5 km long cave near river Oued Guir and village of Ksar Tazougart, one of the longest caves in Morocco. It seems to be a trogloxene or subtroglophile that tends to temporarily inhabit subterranean habitats, but also strongly associated with aboveground habitats.</paragraph>
<subSubSection id="B4AE7DCF02BAEDD764DDC838388C2EE4" pageId="0" pageNumber="71" type="habitat">
<paragraph id="2AD0E6EA49B16E8FF478FDC7BBC9E7E4" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="B4BB4F0BC6D8120E8221BDF1C42E280C" pageId="0" pageNumber="71">The holotype was collected in the cave Kef Aziza, a 4.5 km long cave near river Oued Guir and village of Ksar Tazougart, one of the longest caves in Morocco. It seems to be a trogloxene or subtroglophile that tends to temporarily inhabit subterranean habitats, but also strongly associated with aboveground habitats.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="273F0BAB82BB032B931B74F670F5B205" pageId="0" pageNumber="71" type="distribution">
<paragraph id="19B236DBC1F182A81174AAA197449E65" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="A6DC276883595E3964CE3C2D5332D295" pageId="0" pageNumber="71">
Errachidia Province,
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<normalizedToken id="802B2D049491D6F922C26BB6491ABB53" originalValue="Drâa-Tafilalet">Draa-Tafilalet</normalizedToken>
Region, Eastern-Central Morocco (Fig.
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<figureCitation id="EC2B72C65A0CA6613874FC4488FC3DCC" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
).
</paragraph>
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94
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@ -134,7 +134,7 @@ Martens: Malakozoologische
1878
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, - Nevill: Hand list of
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in the Indian Museum
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: 71.
@ -206,7 +206,7 @@ in the Indian Museum
1899c
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(under
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), - Gude: Science Gossip, 6: 148.
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@ -218,7 +218,7 @@ in the Indian Museum
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1899d
<taxonomicName id="9D17E2A72442167284DD3A19BC613EBC" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea pettos" order="Cycloneritida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="pettos">pettos</taxonomicName>
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, - Gude: Science Gossip, 6: 177.
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@ -226,7 +226,7 @@ in the Indian Museum
1907
<taxonomicName id="464104528308BDC803E26F58021FAEC0" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis pinacis" order="Stylommatophora" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="pinacis">Plectopylis pinacis</taxonomicName>
, - Godwin-Austen: Land and freshwater
<taxonomicName id="38B5BE6099B6FA61724EDF1CBCFD2AD4" lsidName="Helicoidea" pageId="35" pageNumber="36" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
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of India
<normalizedToken id="CF83F52DAC9703D8649E93F0DAAF8BAA" originalValue="…">...</normalizedToken>
: 203-204.
@ -343,7 +343,7 @@ sp. n.
mentioned that the central tooth is larger than that of
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(=
<taxonomicName id="0D4710E43ED65D644958EF293BFADEFA" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea bensoni" order="Cycloneritida" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="bensoni">bensoni</taxonomicName>
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), and that its shape is similar to that of the laterals.
<bibRefCitation id="469C8AF8D648AED2A15E8F17231C00CB" author="Godwin-Austen, HH" journalOrPublisher="Taylor and Francis, London" pageId="47" pageNumber="48" title="Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean; Supplementary to Masers Theobald and Hanley's Conchologia Indica" url="http://www.biodiversitylibrary.org/item/46344page/7/mode/1up" year="1889 - 1914">Godwin-Austen (1889-1914)</bibRefCitation>
gave an accurate description and drawings of the teeth. According to his drawings the morphology of the teeth of

614
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<mods:namePart>Wrase, David W.</mods:namePart>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:affiliation id="DB53F765562CFDDDE4AB757634D727A9">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
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<mods:namePart>Machard, Patrice</mods:namePart>
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<mods:date>2022</mods:date>
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<paragraph pageId="0" pageNumber="71">
<treatment id="5594C38F7CA55DD48FA088B495F9EE00" ID-GBIF-Taxon="195868757" LSID="urn:lsid:zoobank.org:act:8CA1EB54-9BEA-4E30-83C6-BF3B49C83A13" httpUri="http://treatment.plazi.org/id/5594C38F7CA55DD48FA088B495F9EE00" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
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<paragraph id="7010918C5D2BCD106EE3535E2AD91BC0" pageId="0" pageNumber="71">
9.
<taxonomicName LSID="http://zoobank.org/8CA1EB54-9BEA-4E30-83C6-BF3B49C83A13" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) klapperichi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<taxonomicName id="15F6C30049E46FAC79FE9CBD3FAC1A24" LSID="http://zoobank.org/8CA1EB54-9BEA-4E30-83C6-BF3B49C83A13" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) klapperichi</taxonomicName>
<taxonomicNameLabel id="9A19F72A5B1F9D7091C72D469582E393" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2C</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5E</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
<subSubSection id="47CF2E94A4FBC4F2D82FBADC29D84657" pageId="0" pageNumber="71" type="description">
<paragraph id="720ED6DC0992EC3FB4B5232B161D7A1C" pageId="0" pageNumber="71">
<figureCitation id="9CCF905C722F2ADBE477406ED3B7C400" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2C</figureCitation>
<figureCitation id="FD26AC86EB034AC9E136A02E01D1ABF6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5E</figureCitation>
<figureCitation id="F9C9377069A569F798D34F38141024BB" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Iran, Alborz Mountains, Damavand, 2000 m.</paragraph>
<subSubSection id="BE5DCA5CF62C9F78397D32CFDC440A20" pageId="0" pageNumber="71" type="type locality">
<paragraph id="5A251EE9BC5094BAE7B3BF5F85853972" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="6D3E4F2807B06C47C23B8C0331B457C8" pageId="0" pageNumber="71">Iran, Alborz Mountains, Damavand, 2000 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">The locality is situated in the Province of Tehran, in the Damavand County (Central Alborz Range). Data on the label are insufficient to locate whether the holotype has been collected in the surrounding area of the Damavand City or in foothills of the Mount Damavand.</paragraph>
<caption doi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 10.</emphasis>
<subSubSection id="F0B2684662A43B17BB9A55C25FD5C0D2" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="6512A9FC2AED06E8566DFDAA1915F6CC" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="B09121334EC44A63E33F6587F5BE9E7A" pageId="0" pageNumber="71">The locality is situated in the Province of Tehran, in the Damavand County (Central Alborz Range). Data on the label are insufficient to locate whether the holotype has been collected in the surrounding area of the Damavand City or in foothills of the Mount Damavand.</paragraph>
<caption id="A2623DCD3B39036637E9487C113329E1" doi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71" start="Figure 10" startId="F10">
<paragraph id="5AE8E88DEC22961357A82EE34669CC43" pageId="0" pageNumber="71">
<emphasis id="1AEEC748846ED548553FE6711DC7462F" bold="true" pageId="0" pageNumber="71">Figure 10.</emphasis>
Median lobe of aedeagus, ventral view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
<emphasis id="250AD8D7E168D00C54A4F9C2FE7D3314" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="8FE243F72FF188B00AC07BEADFDE6F91" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<emphasis id="B75DDE96D4AFD86EEB01B1549B0532EF" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="FD2962125DF359AEF62059D1D0251FEA" class="Insecta" kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken id="C9C36965182E61F2082442C8291BF2F1" originalValue="Mařan">Maran</normalizedToken>
, 1940, lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
<emphasis id="E05C6D45B993BD20E478AD85F11FC2BE" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="9330DCB0CF763AF55E6633D58DD2EFB1" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
<emphasis id="CE18205C08753B860955594577784AC5" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="C87E5EAD7348857CE12C43CFB1E64859" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
<emphasis id="809D128187F6A9036080B00C85C1BBD7" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="DF5BCD890368780B3937CCE51C11474B" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
<emphasis id="5D1D6FE335F4FAD976D5100E6DC5C25C" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="0E025B818DD4DF43D519EDC746FADF5B" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
<emphasis id="F6DF1A388022635134B84BBFC44B3C64" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="962BA707CA7D2976842F680E497DAAF8" class="Insecta" kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
<emphasis id="463F37A8CCBC878F8B1DDE74204A91C4" bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName id="1866E0290D393B029B8A13DE3DBD3361" class="Insecta" kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
<emphasis id="BB1CB22EAB0EF7A18AB1D85122655774" bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName id="0CFB60B0A19E85FB25912F3C7A31433F" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, male specimen, Tochal, Tehran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName authority="" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">P. (Platyderus)</taxonomicName>
<emphasis id="D5754136A2E4E0B148DCADF6E9C299A6" bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName id="F1350006A622BE41687F18C18418D57F" authority="" class="Insecta" kingdom="Animalia" lsidName="P. (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">P. (Platyderus)</taxonomicName>
sp., male specimen, vicinity of Izmir City, Turkey;
<emphasis bold="true" pageId="0" pageNumber="71">K.</emphasis>
<taxonomicName authority=")" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (P.)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="P.">P. (P.)</taxonomicName>
<emphasis id="2829630D646F78F944FF1AED87255FC1" bold="true" pageId="0" pageNumber="71">K.</emphasis>
<taxonomicName id="341B93908111E7664FC5988F36AC4257" authority=")" class="Insecta" kingdom="Animalia" lsidName="P. (P.)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="P.">P. (P.)</taxonomicName>
sp., male specimen, Boz
<normalizedToken originalValue="Dağlar">Daglar</normalizedToken>
<normalizedToken id="A75ECC188E3161DE865F108D23F18122" originalValue="Dağlar">Daglar</normalizedToken>
Mtn., Turkey. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="8AC84A599A7C3CC0242AA952AB3F9DA7" pageId="0" pageNumber="71" type="type material">
<paragraph id="A326AE20B1C8EEC43ECB7CDF77A25AF5" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="56B893E92E97B6D6379C9B2D5E9A544F" pageId="0" pageNumber="71">
<emphasis id="2CA79D18300C6FD6173BB2B51F491E3B" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂, 'IRAN, Demavand / 2000 m, Elbursgeb. / 2.VI.1960 / leg. J.
<normalizedToken originalValue="Klapperich’">Klapperich'</normalizedToken>
<normalizedToken id="6A852E687B6AC6E59E4A08F23332B2D8" originalValue="Klapperich’">Klapperich'</normalizedToken>
(HNHM).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 1 specimen. TGE: 0.</paragraph>
<paragraph id="5F73BE96570C3B18A19475CF5746839A" pageId="0" pageNumber="71">TME: 1 specimen. TGE: 0.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">The species is named in honor of the German entomologist and collector of insects Johann Friedrich Klapperich (1913-1987), famous by its very successful expeditions to Southern China, the near and Middle East, and who collected the holotype of this new species.</paragraph>
<caption ID-Table-UUID="25F9C0CC5B2F4E7FD96A07933453D3C4" httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Table 2.</emphasis>
<subSubSection id="ED191A9311AA36AE2D556EE6301319CE" pageId="0" pageNumber="71" type="etymology">
<paragraph id="A58B8E70973816A170C1929CE07A1CC9" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="BE421CEABE77037EC7304A77B90ED6E6" pageId="0" pageNumber="71">The species is named in honor of the German entomologist and collector of insects Johann Friedrich Klapperich (1913-1987), famous by its very successful expeditions to Southern China, the near and Middle East, and who collected the holotype of this new species.</paragraph>
<caption id="25F9C0CC5B2F4E7FD96A07933453D3C4" ID-Table-UUID="25F9C0CC5B2F4E7FD96A07933453D3C4" httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" start="Table 2" startId="T2">
<paragraph id="15A9EEA074306FFDB3156507E0644177" pageId="0" pageNumber="71">
<emphasis id="F684B7F701B9365DD4014BA11AC3E08D" bold="true" pageId="0" pageNumber="71">Table 2.</emphasis>
Morphometric data for species of the '
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
<taxonomicName id="DAA939EDBD5FE5702B9676091F346854" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis id="AE9C325CE36949103792ED15DB6C1044" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
</taxonomicName>
', '
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="673D85CA961AA378FF6ECCC34AF5A587" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="ED50066350E808512405C783E546C12B" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
' and '
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus afghanistanicus</emphasis>
<taxonomicName id="DE92EB774487B825C84C3636C185415C" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">
<emphasis id="DD29BD4BDD8423E6AE58259FCA67A9D0" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus afghanistanicus</emphasis>
</taxonomicName>
' groups of
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="9904B53DC6D447A85B1A6F3B449549BD" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="633D3A652B898C59CD9974A6788ED67F" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="71">
<table pageId="0" pageNumber="71">
<tr pageId="0" pageNumber="71">
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">Species (number of samples)</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BL/mm</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BW/mm</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/HW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PL</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">
<paragraph id="D1853EBF4FBC624C22A270D3E0C816D4" pageId="0" pageNumber="71">
<table id="0D8AF081695393906F914893CA28AF3F" pageId="0" pageNumber="71">
<tr id="9DE8B4207CBCCCFE7B6764B9427FB55E" pageId="0" pageNumber="71">
<th id="6B275D9EB38547344EBCD49A7F1C79C2" colspan="1" pageId="0" pageNumber="71" rowspan="1">Species (number of samples)</th>
<th id="ECDFB2E5D8BDB6B95F9DA5EFBDF01438" colspan="1" pageId="0" pageNumber="71" rowspan="1">BL/mm</th>
<th id="4B70B277555D865BBB6B86215B324C48" colspan="1" pageId="0" pageNumber="71" rowspan="1">BW/mm</th>
<th id="F307E3909A7BE1E5E28A6D4F76C02AD4" colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/HW</th>
<th id="1CB148F754501B06B4B41E07C37D37DF" colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PL</th>
<th id="9FAF9F502ACE2FF87DF36AAEF12AE237" colspan="1" pageId="0" pageNumber="71" rowspan="1">
PW/
<normalizedToken originalValue="PА">PA</normalizedToken>
<normalizedToken id="3DB70AA4720AA460C8B99FED95DBFBB5" originalValue="PА">PA</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PB</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PA/PB</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/EW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EW/PW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/PL</th>
<th id="CC13AD66FA694CC5BAD0BC3979E673E2" colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PB</th>
<th id="74DF091D76C0B60DA39B75F7D7B1B075" colspan="1" pageId="0" pageNumber="71" rowspan="1">PA/PB</th>
<th id="1D1497FFD84BAA9FEFD3BA8989ABFCB4" colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/EW</th>
<th id="70793E948D466F6B8C2BC658DBF0B91C" colspan="1" pageId="0" pageNumber="71" rowspan="1">EW/PW</th>
<th id="CE0DA5F7E565E77F2E988C1718404B41" colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/PL</th>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<tr id="AF60BB5D572AD026CD6E802AA0D62C0F" pageId="0" pageNumber="71">
<td id="254B917C4CCFF750FF8787FC7A27336A" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="33B8B1BD7BE9DDB3882FF83A70C26A4E" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="77C1980A5E2788E834DDD29EF35A43BB" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
sp. nov. (2♂♂, 1♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.05-8.50</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.95-3.05</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37-1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23-1.24</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39-1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.16-1.18</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83-0.84</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.49-1.55</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36-1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.51-2.62</td>
<td id="430BE6026C42AAC2E838DCE528A818BB" colspan="1" pageId="0" pageNumber="71" rowspan="1">8.05-8.50</td>
<td id="DDF57E863D84071D0A085BE33B6C03E9" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.95-3.05</td>
<td id="E40FC462647F4A1BD5214537E4317B1E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37-1.41</td>
<td id="9AD3CEF375512F7E880E9815C022F22A" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23-1.24</td>
<td id="80D16B3844BD1D890D9CE0E1456BB481" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39-1.40</td>
<td id="1D61A41078850212D276E5ECA5217A0B" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.16-1.18</td>
<td id="AD726C3AF85ADB48496A905A55D6EA0E" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83-0.84</td>
<td id="B4F0351414629D63E380596322834F6E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.49-1.55</td>
<td id="3184E9430DA04CCE0A38B420A4B20C06" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36-1.41</td>
<td id="4896B6B92BC8D2805B89E4974413AAE9" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.51-2.62</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<tr id="CFDE4FCFB34F468F007B74BE30E0CBB3" pageId="0" pageNumber="71">
<td id="FCEFCF5EDDB52B377875009451C6834D" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="F75F9B31957E29EDEFE46263892B7712" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="DD7D38086D5E58E49C9E629FC93238C3" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.27</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.00</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.17</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.51</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.58</td>
<td id="50007E2EFF5DB1A00D633C5C506EBD8F" colspan="1" pageId="0" pageNumber="71" rowspan="1">8.27</td>
<td id="62F59B30EEF8D87D949793DA2E790095" colspan="1" pageId="0" pageNumber="71" rowspan="1">3.00</td>
<td id="1E989E27CCCE305A8DE4EF5EEE04D034" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
<td id="62AACE5BA2DF61BB9B62B6DD79A6C0BB" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23</td>
<td id="572E8A8CD5853A1EB4B213E8C2E7E47F" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td id="6B38BB20D56FE6F1E7FC966522952322" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.17</td>
<td id="1E6A533668C75E49F13EB49FE26602F5" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
<td id="B0B52F93949DCAAFC10432BC3F918BF1" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.51</td>
<td id="0907CE9F86C6758354EE2DF6C13EB5CC" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td id="C0EBD0D8DFD0D9AC759100207110A52B" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.58</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<tr id="4843ED8063CD3EA50697704F5F12D5F9" pageId="0" pageNumber="71">
<td id="81B50B6367FD7A1E9733ADB2DDCF3356" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="B4E9DB47E1869CB3FECFC0A7A7DCA206" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="0D5D79F94AA13CBCCA4761F20E6607B2" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
Morvan (2♂♂, 1♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.35-7.90</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45-2.55</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.43-1.51</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.16</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.43-1.49</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23-1.28</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84-0.88</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.69</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.34</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50-2.59</td>
<td id="6CD236B7A1B67D448F55EB8FF6C72754" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.35-7.90</td>
<td id="2C651B2B5A4598AE3905D2EE39B4AD13" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45-2.55</td>
<td id="3062C7C625FF0D2391DD7FA04914A18C" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.43-1.51</td>
<td id="F5A5A00F0FC3AB8C6940CF778D0B373B" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.16</td>
<td id="64CF06EC0B4C80E196B06D5D946F9A9C" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.43-1.49</td>
<td id="3A226CBE92CE70ED8033400E5D8B2DD6" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23-1.28</td>
<td id="65BC9989D25BC104DEA5934288CBFAAA" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84-0.88</td>
<td id="93A8060E9E238BEB2D15CB6D39046611" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.69</td>
<td id="C082C31A47DA3E6CF98FA45670337B21" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.34</td>
<td id="DCA9626EE6C2EA68FEE6F31AEAEE7520" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50-2.59</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<tr id="053790F8CD3574588854A479D7E48AE9" pageId="0" pageNumber="71">
<td id="C736964FB5AC5D872119386F4C1D6002" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="618AA089BB647BC2CB0CDDF0B3760B2F" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="36D5B9A1A891338EFB5E4C73FFE3EA02" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
Morvan (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.53</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.52</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.48</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.46</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.86</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.67</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.33</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.55</td>
<td id="4D69EF31EBEA4784E259B63D816D1CCF" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.53</td>
<td id="ED8FF9F4A36A064877D3EF0AD92FB6CE" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.52</td>
<td id="5F00556C709EA1D5472AD75BC371D9C0" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.48</td>
<td id="33893EB4E73C0826DBCAB0FE23071CFD" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
<td id="408F803D272B36E125278D62B3098367" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.46</td>
<td id="A9FCCDD82FC037F79C825852A200BD7C" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.25</td>
<td id="532EC9F0F75C10565042BC4317A1A5CD" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.86</td>
<td id="F9610AD9B88FBDE495FA7454DCC26747" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.67</td>
<td id="7045A36A3CF208516D705042E86486DC" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.33</td>
<td id="16C5265AED57167ED052A6F34817DC86" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.55</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<tr id="9F37D808160B707B435C345EB4FD61B0" pageId="0" pageNumber="71">
<td id="B362C6A81D3B5458FF1567E92A439D75" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="01CA18470FEE333C9B4A2C2B47537B31" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="D22ACD0660C20A58EF7EF884FCCA6153" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
Morvan (1♂, 1♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.10-7.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31-1.33</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.06-1.12</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34-1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.22</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83-0.91</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.65-1.68</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41-1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.47-2.67</td>
<td id="E8817376278ADA1896D054A9A24C8EE9" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.10-7.20</td>
<td id="A30FE3CB49DB0845DC15371CA7AD95D7" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40</td>
<td id="8FD6E089832A00B85CFCED74EC5F5E8D" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31-1.33</td>
<td id="426D0EB374FB7C21B4B7EC939807F4F2" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.06-1.12</td>
<td id="BE21F4A8A52FF30F87BA929A8DF05791" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34-1.36</td>
<td id="5DF9561E50D4FE6422E19FA1C4A6E20C" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.22</td>
<td id="D05DE66E7A08EBFDE7CDE727C8B83627" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83-0.91</td>
<td id="7DCD2C0FD5F0C8CB9B8F9250166076B0" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.65-1.68</td>
<td id="67761C89171947DC75FA7B3F00D9F1F9" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41-1.42</td>
<td id="A786EC3D8FBE9C1C8135221C11A8ABBE" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.47-2.67</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<tr id="4292C9C04EC890C7726A7CA4253D909C" pageId="0" pageNumber="71">
<td id="E679C0B9A994EADBA5A769410D847F74" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="91853693856F5DA94ECE0C4B5B7BEB57" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="8879CC739BBA64005054DF0C2E6DBF6B" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
Morvan (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.15</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.87</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.67</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.57</td>
<td id="5FA9439F7718969EC12DB399FF8520B1" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.15</td>
<td id="7AD7FDB45080B6066375BB1906A714E1" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40</td>
<td id="30204FD1321494F333991651BC5498D5" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td id="76BCA6394D271D78F5541D58242785B5" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09</td>
<td id="D608889A645AA6420424622DB29263F1" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35</td>
<td id="649FA806E811B8EC45DF6970CAE22024" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td id="C8B73046048E8E9B12103AF7E916C2A3" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.87</td>
<td id="A52DBB8D7ACAA8357745E647972D610E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.67</td>
<td id="140F73B5744E8D9D90ABC755D8CC4B97" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.42</td>
<td id="6C85607E8197EDD1CF82D00B6B7649CF" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.57</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. klapperichii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichii">
<emphasis italics="true" pageId="0" pageNumber="71">P. klapperichii</emphasis>
<tr id="976E9E7799DA0FD6354E5B7414B39521" pageId="0" pageNumber="71">
<td id="A3103E8F4219C2BE7CE5179E3AD92169" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="EBEC7E45A3A5CBF2C56F1647ECB10D91" class="Insecta" kingdom="Animalia" lsidName="P. klapperichii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichii">
<emphasis id="1BAFDA76F8F5C14C19A463753056AE32" italics="true" pageId="0" pageNumber="71">P. klapperichii</emphasis>
</taxonomicName>
sp. nov. (1♂)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.60</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.27</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.70</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60</td>
<td id="E7475C0DFE0F7E9761F73C59DB6DAF56" colspan="1" pageId="0" pageNumber="71" rowspan="1">6.60</td>
<td id="BC2BAD7665B45F4EE62F93F8C64AD942" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.20</td>
<td id="1CEB9F3E77C52A96445C4BDF3CFDFC9E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.27</td>
<td id="3B369535745A1B8B467E3076833D6180" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09</td>
<td id="8D9E4EC4CEA1A3336630EFC774911181" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td id="1FC858C95CEE96AAEC9EE43792CDF904" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
<td id="117FF0047112D5B86EEA963E80F2CF4E" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83</td>
<td id="C41536F4E614F31B6420D5217E2660C5" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.70</td>
<td id="EADE4E6F07828E075B1577127FAC20FA" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td id="D0B4D446B5FABA9507BEC938BC053C60" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
<tr id="A9C387C853A10D28F8EC787CD4A7BBD8" pageId="0" pageNumber="71">
<td id="B61357FED29E9B12487944284EF0AE62" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="B383C8F8FBC48D6747986118A465D470" class="Insecta" kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis id="4E90EBE3B6E388EA01441BACB83FB7B3" italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
</taxonomicName>
sp. nov. (1♂)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.80</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.24</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.49</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.24</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.78</td>
<td id="AE90FD6508DA3B68CDCA1CB7BC8DDA2F" colspan="1" pageId="0" pageNumber="71" rowspan="1">6.80</td>
<td id="AFA561A00B4794447F142BB3CD74EAF8" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25</td>
<td id="8B1CA80A9C97F69189B3758D313A9111" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td id="F8F02D5D87A2C71E60AAAD199CCD6E34" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.24</td>
<td id="DF0F101A00EEA68267BE859895C26D5F" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.49</td>
<td id="2CA2B864B45809211AD261727B347290" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.24</td>
<td id="51CD3D73F1AC49BD394ED70DFBB9D55C" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83</td>
<td id="E76E0FAB3938E3D906B9AFCB0000A5DE" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63</td>
<td id="4F857A7EC09C2971FADDB5879D1C8588" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37</td>
<td id="B4540584876FF5C06D051B215490AFC1" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.78</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<tableNote pageId="0" pageNumber="71">For abbreviations in line 1, see &quot;Abbreviations to measurements and ratios&quot; (section &quot;Material and methods&quot;).</tableNote>
<paragraph id="D15C78903A719522DF3C8CECC391A87B" pageId="0" pageNumber="71">
<tableNote id="5EBD4A9D736A584E8F9FFE0FE628E912" pageId="0" pageNumber="71">For abbreviations in line 1, see &quot;Abbreviations to measurements and ratios&quot; (section &quot;Material and methods&quot;).</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="CCB28FD1DAE60F6ADF1E18243E1ACE1F" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="3A210F0A5071F7B8EE4ACABD80C0FB2A" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="367603A2850E7CEA89FFD884EA3F6347" pageId="0" pageNumber="71">
It differs from other representatives of the &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="C792748D34DA265A15558F0715A38AB5" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="EC4C345462F18D4C23126819F7C12815" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; group by very small size of body (BL &lt;7 mm) and pronotum less wide in relation to head (PW/HW= 1.27). In addition, it differs from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="A15D13D5F31EDEAF18C40C54EE30B0CE" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="FF4F4A28E4829A5973DEFC702C136D54" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
by anterior side of mesofemur ventrally with four setiferous punctures (vs. three setiferous punctures in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="7C7AB5EAF477741685BCDE4818F49ACC" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="BC7472DBA0166031D53BFB360F23DE06" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="36A141C5913A0C4B0528DA824498B3CD" pageId="0" pageNumber="71" type="description">
<paragraph id="1280B79959579F3F15312BD2A40BD3CB" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="68F32FA0BDC9AEFD0B1840387C9D8D9B" pageId="0" pageNumber="71">
<emphasis id="3B49C9D1908BCF973665F61F51445B97" bold="true" pageId="0" pageNumber="71">
<emphasis id="8CAABB0C7C5485582BC7DBB0C7A7AF59" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimen of small size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="46126D29200D53CC1B8584C0AF67337F" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="0F2CE2B19864509A6D5B2220EDA07C84" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species, with elongate and convex body (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2C</figureCitation>
<figureCitation id="6C1118D4D0EF40C88FB6B1AE67FD8409" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="705CE773A002F422474953321418C934" bold="true" pageId="0" pageNumber="71">
<emphasis id="48C619E821CAEE291B5908499C332672" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
<tableCitation id="DE7E9CEF9F46609F79E42F1B27BEAC1E" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="78B81170790101487F66B85603AABC8F" bold="true" pageId="0" pageNumber="71">
<emphasis id="6B06770384E1BE16697FC56FC33E1877" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body uniformly dark brown (castaneous), antennae, palpi and legs reddish-brown. Integument moderately shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="359B67ED60FA0AA50C01782F28073B59" bold="true" pageId="0" pageNumber="71">
<emphasis id="AAB3B2566249470C95358862DA32462D" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Microsculpture of head faint to absent on clypeus and frons, present on vertex. Pronotum with isodiametric sculpticells only posterolaterally, remaining surface without distinct microsculpture. Lateral parts of pro- and metasternum, pro-, mes-, and metepisternum, abdomen and legs with slightly stretched isodiametric sculpticells; ventral part of head, prosternum and metasternum medially without or with reduced sculpticells. Pronotum basal half with several transverse wrinkles.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="19A41E95153FFE48EF7A3CB9FE06988F" bold="true" pageId="0" pageNumber="71">
<emphasis id="553A7EDC3D8080C1F7903430DE68232E" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
Narrower than pronotum (PW/HW= 1.27). Frontal furrows slightly impressed, shapeless, wrinkled. Paraorbital sulci moderately deep, ending at level close to posterior supraorbital pore. Frons behind frontal furrows wrinkled.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="A878B89FE158ADB0902B058F378C7042" bold="true" pageId="0" pageNumber="71">
<emphasis id="71DC547D34A6945C54AD1ACD28475946" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum barely wider than long (PW/PL= 1.09), with widest point at second quarter. Anterior transverse impression distinct, posterior transverse one indistinct between adjacent wrinkles. Sides moderately curved apicad and basad, convexly anteriorly, concavely posteriorly; beads of anterior margin and posterior margin present laterally, very faint to reduced in medial 1/8. Metepisternum somewhat longer than wide, MA/MI about 0.8.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="2AAE458C265AF47B4E1E567773A36344" bold="true" pageId="0" pageNumber="71">
<emphasis id="15120542EABE8B3A96B5015C87A7C55F" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Cylindrical, about one and two thirds as long as wide (EL/EW= 1.70), wider and considerably longer than pronotum (EW/PW= 1.40; EL/PL= 2.60). Stria 7 ending at first umbilicate puncture on left elytron or after reaching first forming a kink connecting with the basal rim on the right elytron. Two or three discal setiferous punctures, anterior puncture near stria 3 on left elytron, and lacking on right elytron, remaining two punctures adjoining stria 2. Umbilicate setiferous series of 15 setiferous punctures on left elytron, with 16 punctures on right elytron.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="A8DBD5741C2388A04997371CB1742F7D" bold="true" pageId="0" pageNumber="71">
<emphasis id="1C49F608F2AD7F6375A4886B02288128" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Anterior side of mesofemur ventrally with four setiferous punctures (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5E</figureCitation>
<figureCitation id="1344E062A3539F5A0B61D20BAB30C31F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5E</figureCitation>
). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="94B526B32D601124E76B57F003EEE01B" bold="true" pageId="0" pageNumber="71">
<emphasis id="E31069A1BCEFD7B12BFA0BF2FDE8CBC5" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Unknown (see
<normalizedToken originalValue="“Notes”">&quot;Notes&quot;</normalizedToken>
<normalizedToken id="241858891A9E331612608B0CD18A89CA" originalValue="“Notes”">&quot;Notes&quot;</normalizedToken>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="BA32F2EC5039B2C61560880E87DA00ED" bold="true" pageId="0" pageNumber="71">
<emphasis id="C21FE81E7CC11BA55690AC234365826D" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="BB48B459089C9B73D40AFF98FC39D45B" pageId="0" pageNumber="71" type="comparisons">
<paragraph id="066B3CC88B68690644D0EF6282ECB560" pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph id="AF2EAF6471BE7238DA33C15EE8E0BA66" pageId="0" pageNumber="71">
The species can be easily distinguished from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="6AF2F37CD1A2022E3460613D9BBF16E4" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="0DFF3FF18D1BB2E5908F9DED9DF150F5" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
by characters noted in section
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="B539280C8B6D6BBE281B6349D6914D8A" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
(under
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="03CCFDF4AC276F2E464D3C9A0B145D2A" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="FB66816576038FEFF3343A6A075FFE9C" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="674CC995EA8DDC64EDA21CC41C00AF06" pageId="0" pageNumber="71" type="habitat">
<paragraph id="EBE4439C13F2E9E37B598ECCC856E332" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="B50B93CD94DC53D910B056272256F7BE" pageId="0" pageNumber="71">
Like other representatives of the &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="96E10759E690C737B59FA79CE9D98639" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="7C12EE786D2AA06838FBD9F2789B6750" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; group,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
<taxonomicName id="2C424A7E52E8F42A8DF3D55B4F8A4E41" class="Insecta" kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="3E67AC0B9B183A82FC8B25B144738AE3" italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
</taxonomicName>
inhabits high-mountain meadows around and above 2000 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="FC0B5B7DD65FB93788D80A604405C6A7" pageId="0" pageNumber="71" type="distribution">
<paragraph id="003593F096C4D45341BD9988183895CC" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="88B61ABD4DCF654F8D23103E2B71A5EF" pageId="0" pageNumber="71">
North Iran, Central Alborz Range, most likely in vicinities of Damavand City (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="5B221B28FDD50CFFD71065B89828CBA4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">The aedeagus with attached parameres and urite IX were lost after the extraction.</paragraph>
<subSubSection id="B5D754834CC7FD0CB18A9A21B4CD77DE" pageId="0" pageNumber="71" type="notes">
<paragraph id="60E1A6F695F0679E54D153185479C7D1" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="D03C4E744764C4717B582A8B222B0C49" pageId="0" pageNumber="71">The aedeagus with attached parameres and urite IX were lost after the extraction.</paragraph>
</subSubSection>
</treatment>
</document>

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15.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3D, F</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5H</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6A, B</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7I, J</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9C, D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11B, C</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12J, K</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15A, B</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
<subSubSection id="D6469DC7516A83171CE53D495AF0ACFA" pageId="0" pageNumber="71" type="description">
<paragraph id="87197DC0089602A5086464917D294666" pageId="0" pageNumber="71">
<figureCitation id="22ED3D4ACD4025612E209F4DDC60B0BB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3D, F</figureCitation>
<figureCitation id="1392E83F4C7610D41D06B31BE58384E9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5H</figureCitation>
<figureCitation id="6B8D9602E175523E7241198679C543FE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6A, B</figureCitation>
<figureCitation id="A7CD08F76E23A2A153A603B2F3DFBC85" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7I, J</figureCitation>
<figureCitation id="F11D97EB658399622857F668668D5ADA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9C, D</figureCitation>
<figureCitation id="C9296947179883CF8E7A95DFB6DF70CA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11B, C</figureCitation>
<figureCitation id="1825677A5F16A3BA6E05869BF65AB701" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12J, K</figureCitation>
<figureCitation id="465BF25C2FAA4D280D90B02A5E4BBE71" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15A, B</figureCitation>
<figureCitation id="7A1E5569D2E1995B0F3536D150F8E9FF" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="E7D6571A98FC2BB624E35183F094732C" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="5F1CC9BE6239D33C1CB7B9AE2F2EEE8D" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="E528E28AAC55A9B19F3CD2B0B5F3BF41" pageId="0" pageNumber="71">
A species of
<taxonomicName authority="" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus (Eremoderus)</taxonomicName>
<taxonomicName id="3A0B3FDBBF62AF989AA8CF04A57DEABB" authority="" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus (Eremoderus)</taxonomicName>
, with mesotarsomeres and metatarsomeres dorsally more or less flattened and grooved (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6A, B</figureCitation>
<figureCitation id="F189906866308B41FCCDE6E236F13FE1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6A, B</figureCitation>
), cordate pronotum, with sides from widest point to base markedly (nominotypical subspecies) or barely concave (ssp.
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="1844B684789F1770582FC86DC4BB59DB" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="173D670E818C979C9E35F24A01462E48" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
) and posterior angles obtuse, with tips laterally more (nominotypical subspecies) or less protruding (ssp.
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="1EB42EC04FCCAF21C617A9AD709F109A" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="ACB522282CAA2718AAD80B5E515BAF68" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
), and median lobe of aedeagus with basal bulb more slanting to shaft (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
<figureCitation id="670362D686C533AE36BF9D8787CB6D3E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="redescription">
<paragraph pageId="0" pageNumber="71">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="98033BABF677886AF007E7BC6B6C60DF" pageId="0" pageNumber="71" type="redescription">
<paragraph id="793431F73E953B37CDA254382CCC5837" pageId="0" pageNumber="71">Redescription.</paragraph>
<paragraph id="DE63A72FDFAB6A3B0ED46B6A7B6DC57F" pageId="0" pageNumber="71">
<emphasis id="C050192CB099947A88FA16A0AEB67020" bold="true" pageId="0" pageNumber="71">
<emphasis id="1DCDB8A7E1B2ADF53D5E8FFAF85BDE5E" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Moderately to large-sized species (BL: 7.60-9.30 mm; BW: 2.70-3.40 mm), with elongate and moderately convex body (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3D, F</figureCitation>
<figureCitation id="93B0BE401E7212622090EDF61B90F38B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="774B584CA10DC8DCA09F55FD26F09B58" bold="true" pageId="0" pageNumber="71">
<emphasis id="3C6C5FDC6A61997008CD3969C2520D2F" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
<tableCitation id="551B9FA5D8160B6B5A9520F2DCDDECB4" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="868D5BC981FD81C6951E727EEB194ECA" bold="true" pageId="0" pageNumber="71">
<emphasis id="E40DF5A69EED5116AD724DAF6FDD74D1" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body, palpi, antennae and most of legs yellow-brown to reddish- brown, with head usually darker than pronotum and elytra and femora lighter than tibiae and tarsomeres. Integument slightly shiny (less shiny than in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="D6EBAD4CDDAB6027760D6F88BE2F4840" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="3DCFFB2830AAD8DCAB1F13C13E48E1E1" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="26C71D2E823F036C3696DDCB8260FF4D" bold="true" pageId="0" pageNumber="71">
<emphasis id="94A9FB322F70624E4B61F6BFFD8F8B78" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum with evident microreticulation, sculpticells regularly isodiametric. Elytral intervals, scutellum and lateral gutterwith distinct isodiametric sculpticells; basal margin with reduced microreticulation. Ventral surface with impressed slight transverse sculpticells or with sculpticells scarcely-visible. Head impunctate, usually smooth, clypeus and areas laterad of frontal furrows sometimes with fine longitudinal wrinkles. Surface of pronotum impunctate, largely smooth, only basal area medially behind posterior transverse impression longitudinally wrinkled, rarely apical part medially in front of anterior transverse impression also wrinkled; basal foveae and adjacent lateral areas smooth, smoother than those in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="93067E9758C78D3FB0060C7E2A8E3D82" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="F772415986A89AB38E8FB0567B2050E9" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
. Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled in whole, 2-5 finely wrinkled at sides.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="A694D7205E9DA9E1AAA078CAEE20F48F" bold="true" pageId="0" pageNumber="71">
<emphasis id="BB901E9997C726A210B022C738D67E2B" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About one-third as wide as pronotum (PW/HW= 1.28-1.41). Eye long, moderately convex. Labrum subrectangular, as long as clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, reduced to disappeared at sides. Frontal furrows small, rather shallow, punctiform. Paraorbital sulci straight, fine, backward surpassing posterior margin of eye, nearly reaching level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="D660F8505963C7A9E450B0BE623AE6B8" bold="true" pageId="0" pageNumber="71">
<emphasis id="D1C19D698F9BD5F2E3D3A72446860273" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum one-tenth to one-fifth wider than long (PW/PL= 1.10-1.21), with widest point at second quarter. Anterior transverse impression absent, posterior transverse impression short, barely distinct. Sides convex medially and anteriorly, slightly concave to straight in base; anterior bead present laterally, lacking or present in medial eight to tenth; basal bead present laterally, reduced to absent in medial fifth to half. Metepisternum slightly longer than wide or nearly as long as wide, MA/MI= 0.87-1.00, with greater values occurring in specimens of ssp.
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="718E2F93D1620D0B0A685AAEC2E34218" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="22BFAD0FC1817899DC5F2FF6C5BDE9F1" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="1F6D93E01AB9581E98D4568121EC7A7D" bold="true" pageId="0" pageNumber="71">
<emphasis id="F87A5AB1CFE85A9505374EC543715D1B" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Oblong, about one-and-a-half times to one and two thirds as long as wide (EL/EW= 1.52-1.72), two-and-a-half times or little more as long as pronotum (EL/PL= 2.46-2.72), and one and a third to a one-and-a-half times as wide as pronotum (EW/PW= 1.35-1.48), with widest point at medial third. Parascutellar striole and striae 1-7 very shallow to indistinct, stria 8 evident, clearly deeper than other striae (nominotypical form) or parascutellar striole and striae 1-7 distinct, slightly impressed, stria 8 somewhat deeper than other striae (ssp.
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="841648D170DB928744CE0696646452DA" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="B5D85A9A346D5C26F444AC575E3DFC7E" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
); parascutellar striole, if present, not joining stria 1; striae 1-8 superficially to moderately punctate; bases of striae 1-7 more or less reduced, not reaching basal bead (nominotypical form) or distinct, reaching basal bead (ssp.
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="84D1B0B001C53C30D0D5EC4CB5517B42" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="7C8EA7B4B52B33AF6168B340878D1904" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
). Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 or 17 punctures on each side.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="666AE40683FF3220DDF0B8BBB322BE75" bold="true" pageId="0" pageNumber="71">
<emphasis id="C4F976109EE4AACAC702091644D2C006" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one or two setae in basal third and one or two in medial third. Mesofemur with 4-7 setiferous punctures on anterior side ventrally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5H</figureCitation>
<figureCitation id="88C70F923A479EBDA55F380EF51EE947" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5H</figureCitation>
). Anterior side of metafemur ventrally with a few long setae, one in basal third and one to three in apical half.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="48081BB624194F0B12BD2B14BC71CF07" bold="true" pageId="0" pageNumber="71">
<emphasis id="3304463B9913792161028E2CE029C28B" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX suboval, with proximal margin slightly asymmetrical, insignificantly turned to left (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7I, J</figureCitation>
<figureCitation id="05863C0D3C7837AC7C312BC36435A7B1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7I, J</figureCitation>
). Median lobe of aedeagus in lateral view slender, with narrow basal bulb, long shaft and short, straight apex (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
<figureCitation id="BC23E6A36E8F699F1CBB78A7568DCA85" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
); median lobe in ventral view straight, 3.4-3.5 times longer than wide (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11B, C</figureCitation>
<figureCitation id="3AC01AFA6CE42E929551AA39C633B5B3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11B, C</figureCitation>
); apical lamella (dorsal view) short, scarcely asymmetrical, rounded at tip, with right side straight and left side slightly concave. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
<figureCitation id="872021723E9DEE0B10889D3202A41572" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
) with ventral sclerite long, broadened and rounded distally; same in ventral view (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11B, C</figureCitation>
<figureCitation id="176D2FFDAE74CD6883F78B81B3DCC4CC" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11B, C</figureCitation>
), with dorsal sclerite large, reticulate, with left-sided and right-sided protuberances distinct, and ventral sclerite narrow and distally very slightly curved to left. Right paramere as on Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12J, K</figureCitation>
<figureCitation id="F9F3EC427B086E6DF794101D76693D6B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12J, K</figureCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="48944FC475AF4EC301803FD9EA23C43F" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15A, B</figureCitation>
<figureCitation id="4D06A1C52F6E34F1A92D102F529D962D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15A, B</figureCitation>
). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="124F985D828777688D6AC58C96990E4F" pageId="0" pageNumber="71" type="notes">
<paragraph id="20726AD7087D775FF2660CEC0690ABD7" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="CAA8B757FBB16B605453BEBD097E3F41" pageId="0" pageNumber="71">
Due to minor morphological differences, the populations from the lower course of Nile River and Southwest Israel are treated as a distinct subspecies of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="B0FD5534214588707C17CBF6CE53C1D3" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="0A02F5D014E827B373C676F678FC6FC2" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
(see Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
<figureCitation id="2EA668B60E1A8797CDA95F189A97F0B7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
). Moreover, no verifiable records of the species are so far known from the vast area between Misrata District (Libya) and Cairo Governorate (Egypt).
</paragraph>
</subSubSection>

258
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1.
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<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<taxonomicName id="AD71530083530A8C1A18108049EEF0F7" LSID="http://zoobank.org/65B653A0-F02D-4F08-B697-41D5DE8E7B79" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
<taxonomicNameLabel id="11169F483D13B743CB212D08F1EB9D7E" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7A</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8A</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10A</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12A</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
<subSubSection id="0A7DDCFD089C8D4E6159949B3AA2A2CB" pageId="0" pageNumber="71" type="description">
<paragraph id="7FF46C9F750833D37C6F8DBF44E31DE8" pageId="0" pageNumber="71">
<figureCitation id="961D8BA76C66D457E58BF498F3C02A54" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1A</figureCitation>
<figureCitation id="5EA568A871F2013BB7C6CEFBB003435E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7A</figureCitation>
<figureCitation id="984F9C301EDA5C65A16AA4D20A776A5D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8A</figureCitation>
<figureCitation id="1FD3C63937E20D74718ED6543B790266" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10A</figureCitation>
<figureCitation id="D255584D002F225A896EA1A493884FE1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12A</figureCitation>
<figureCitation id="C8C26DF7A4B18219F1075F62427766D6" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Greece, Kalymnos Island, Stimenia Cave.</paragraph>
<subSubSection id="3654E2173B06C1ABB52C2C463D04A97F" pageId="0" pageNumber="71" type="type locality">
<paragraph id="274333931A605FDC61B00769A5E7A60A" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="41CDE80ADC41030E63930D03DAD05C90" pageId="0" pageNumber="71">Greece, Kalymnos Island, Stimenia Cave.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="note on type locality">
<paragraph pageId="0" pageNumber="71">Note on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">The Stimenia Cave is situated in the northeastern part of the Vathi Valley, in an area called St. Nikolas, on the Kalymnos Island, belonging to the Dodecanese.</paragraph>
<subSubSection id="F9167F838208F22E002ADDB628FD38C9" pageId="0" pageNumber="71" type="note on type locality">
<paragraph id="569FB84813A1285D57B772B8384E54FC" pageId="0" pageNumber="71">Note on type locality.</paragraph>
<paragraph id="9C45D6608AE9110F9B0997676EFEEE27" pageId="0" pageNumber="71">The Stimenia Cave is situated in the northeastern part of the Vathi Valley, in an area called St. Nikolas, on the Kalymnos Island, belonging to the Dodecanese.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="860DDE570685309B34E60125EB7B4B70" pageId="0" pageNumber="71" type="type material">
<paragraph id="E024374C0E33BB88297C867D34F3BD4B" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="7AE7F46B29F36517571746F86E88C2F5" pageId="0" pageNumber="71">
<emphasis id="43D51A572C787BC194C3AD24AC5CD71C" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂,
<normalizedToken originalValue="ʻ">ʻ</normalizedToken>
<normalizedToken id="A48B32B2457C5947DD3F8240FF8A885B" originalValue="ʻ">ʻ</normalizedToken>
8422, KALYMNOS Stimenia, end / road st18 9/6/2005, Lg
<normalizedToken originalValue="Chatzaki’">Chatzaki'</normalizedToken>
<normalizedToken id="469496F39B43D08AEB41BC2CB5810D07" originalValue="Chatzaki’">Chatzaki'</normalizedToken>
(NHMC).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 1 specimen. TGE: 1♂.</paragraph>
<paragraph id="2F6E17EEF9DC45C1117E86521BD1B8CE" pageId="0" pageNumber="71">TME: 1 specimen. TGE: 1♂.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">Latinized eponym based on the surname of Maria Chatzaki (Democritus University of Thrace, Komotini, Greece), a Greek arachnologist, who collected the holotype.</paragraph>
<subSubSection id="5E197AF91B08028516A36CEB3085F279" pageId="0" pageNumber="71" type="etymology">
<paragraph id="F4AB8247CDF5C1D1A5CB5C952A924057" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="D40689D9D3AD7C06F3C4084DBD49EA36" pageId="0" pageNumber="71">Latinized eponym based on the surname of Maria Chatzaki (Democritus University of Thrace, Komotini, Greece), a Greek arachnologist, who collected the holotype.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
<subSubSection id="A43379D58C5F337FB93D48377C1803C4" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="1C56E0AEC93DED2EFC9FA416A5E4F16B" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="047745C27B036E4CCBA42D9B808C6C75" pageId="0" pageNumber="71">
<taxonomicName id="3721F9687B1688F8AFAA54E49F3B573D" class="Insecta" kingdom="Animalia" lsidName="P. chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="A73C67203EEAFE1D847D964358F1F395" italics="true" pageId="0" pageNumber="71">P. chatzakiae</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<taxonomicName id="B11BE6452DA53B74F7F6431D78891292" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="DC25A002961F1F0477F031FFBD5CAFDC" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
in wider body (BW: 2.85 mm, vs. 2.35-2.60 mm), more coarsely punctate disc of head (vs. finely punctate or nearly smooth disc of head), anterior elytral discal puncture situated in midst of interval 3 (vs. anterior discal puncture adjoining stria 3), and elytra both in relation to elytra width narrower (EL/EW= 1.60, vs. 1.63-1.71) and in relation to pronotum length shorter (EL/PL= 2.49, vs. 2.55-2.83).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="8891A7C3C2F725561FFC396B8227D6C1" pageId="0" pageNumber="71" type="description">
<paragraph id="24E7336AEF43A9CD24ED3900611579FE" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="17D54A40FE13881C0928D098E3B84B49" pageId="0" pageNumber="71">
<emphasis id="38BE847B175A38B6800B6F47956EAAEC" bold="true" pageId="0" pageNumber="71">
<emphasis id="CEC9F2D72431418B066216F29EAFD03F" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Large size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="82690BA2685BFD1055DA0E7EC04CA0D9" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="CDF0686C8D017DF3294C7E5A10A4EE45" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 8.05 mm; BW: 2.85 mm), with elongate, moderately convex body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1A</figureCitation>
<figureCitation id="0C0F5CD861A7D23B48E7D49534B8BAFC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1A</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="D3FD13A9E117776E5D219FFA53929D54" bold="true" pageId="0" pageNumber="71">
<emphasis id="DBAB004C47070AE12E0F25B207ECAD4F" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
<tableCitation id="6F176B78E91CF1A3D1244F38E2C4C1EE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="CC116F720D59F15F0A649FF8F8154BAA" bold="true" pageId="0" pageNumber="71">
<emphasis id="73EA3D76CB309EDF381EA3FDFDC22BE5" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body dorsally and ventrally red-brown, head, pronotum and ventral surface slightly darker than elytra, appendages (antennae, palpi and legs) and elytral epipleura slightly lighter than elytra. Head, pronotum, and ventral surface rather shiny, elytra much less shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="E65F54DF5CA0F8019F2BFD839E64AAF6" bold="true" pageId="0" pageNumber="71">
<emphasis id="B4BC086BA7E087EFBC5C83008731D5AD" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum without regular microreticulation even on anterolateral and posterolateral parts. Elytral intervals, scutellum and basal margin with distinct isodiametric sculpticells; lateral gutter without microsculpture. Ventral surface with scarcely-visible isodiametric sculpticells (proepisternum) or with sculpticells slightly transverse (abdominal ventrites at sides) to not apparent. Head punctate and wrinkled on disc and posterior part of clypeus, labrum impunctate, without wrinkles, anterior part of clypeus and vertex sparsely punctate, without wrinkles. Pronotum coarsely and densely punctate in basal part and laterally, finely and sparsely punctate on disc and anterior part medially. Elytra with scanty micropunctation. Abdominal ventrite 1 moderately wrinkled, ventrites 2-6 smooth.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="016B0A599178CA9178E8BF78B3570566" bold="true" pageId="0" pageNumber="71">
<emphasis id="0D655C5615B0628140F8CF279A82D81D" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
One-third narrower than pronotum wide (PW/HW= 1.34). Eyes almost flat. Labrum subrectangular, as long as clypeus, with anterior margin rather concave. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows subfoveolate, shallow. Paraorbital sulci straight, moderately deep, ended posteriorly after level of posterior margin of eye and slightly before level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="6A53EC603FBF1B0504AE02DF33716819" bold="true" pageId="0" pageNumber="71">
<emphasis id="C53C5C18D4379DC51D38EC9E2EC62519" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum slightly wider than long (PW/PL= 1.19), with widest point at second quarter. Anterior and posterior transverse impressions slightly distinct medially, lacking laterally. Sides sinuate, convex medially and anteriorly, slightly concave posteriorly; anterior border distinctly beaded laterally, indistinctly in medial 1/15; posterior margin finely beaded. Metepisternum as long as wide, MA/MI= 0.97-1.03.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="0318EC3EEB6D3A52E8CF783A69714B1E" bold="true" pageId="0" pageNumber="71">
<emphasis id="95A64A2AC7FD1BB0B1C17B3159789533" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, less than one and two thirds as long as wide (EL/EW= 1.60), two and a half times as long as pronotum (EL/PL= 2.49), and about a third as wide as pronotum (EW/PW= 1.31), with widest point at medial third. Parascutellar striole and striae well-impressed, slightly punctate; parascutellar striole short, not joining stria 1; striae 1-3 (and 4 on right elytron) reaching basal bead, 5-7 not reaching it. Interval 3 with three discal setiferous punctures on left elytron and two on right elytron, first pair of large, foveolate punctures in midst of interval 3, second pair of smaller, punctiform punctures adjoining stria 2, third pair of punctures lack, instead an additional large, foveolate puncture present in midst of interval 3 of left elytron between first and second punctures. Umbilicate setiferous series with 17 punctures on left elytron and 16 punctures on right elytron.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="845309A2ECA2CE681DD5B6E822EB74E0" bold="true" pageId="0" pageNumber="71">
<emphasis id="A22AE952D3F707EDF4A6A0F3163DC91D" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one seta in medial third. Anterior side of left mesofemur ventrally with five, right one with four equidistant setiferous punctures. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="CEF69CB7B3E7364984DB65419468C9EF" bold="true" pageId="0" pageNumber="71">
<emphasis id="EF28BA6165CF7D64513F7E1DB88EA00F" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX suboval, with proximal margin slightly asymmetrical and pointed (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7A</figureCitation>
<figureCitation id="7B2999C0E3AA6D4EF82FEE83B1EDEA36" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7A</figureCitation>
). Median lobe of aedeagus in lateral view, with short basal bulb, long and broad shaft moderately constricted proximally, and a short, straight apex; median lobe in ventral view straight, about 3.5 times longer than wide; apical lamella (dorsal view) short, symmetrical, subacuminate at tip, with sides straight. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8A</figureCitation>
<figureCitation id="24195FF2C9CAEB7078DBA3366C664405" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8A</figureCitation>
) with ventral sclerite partly visible (as a short plate); same in ventral view (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A</figureCitation>
<figureCitation id="1137373CF3E27E861350BF7A00FABC8C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10A</figureCitation>
), with dorsal sclerite situated proximally and surrounding ventral sclerite from behind, ventral sclerite obliquely situated, broadened distally. Right paramere less concave ventrally (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12A</figureCitation>
<figureCitation id="A666B12D9B756BB60148A2846AD70CEC" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12A</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="99909F8B54EFD9432F3A42ED28FF49AD" bold="true" pageId="0" pageNumber="71">
<emphasis id="59D8448EE7F6799CEE2615BBC795BBA5" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">The holotype was collected in a pitfall trap that was set (3 May-9 June 2005) at the end of a street leading to Stimenia Cave, west of Vathys Village (Maria Chatzaki pers. comm.). The habitat where the specimen was found consists of a homogenous degraded phrygana (Mediterranean open scrubland and grass community at low altitude). Approximate GPS coordinates of the location are: 36.9975 26.9607.</paragraph>
<subSubSection id="70E0971446B117C4500C16D4C746879B" pageId="0" pageNumber="71" type="habitat">
<paragraph id="93937BB3D71FBC66821BB0EFFC7DCDFC" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="8D353FF9D8C5090200FC334C4B45F65A" pageId="0" pageNumber="71">The holotype was collected in a pitfall trap that was set (3 May-9 June 2005) at the end of a street leading to Stimenia Cave, west of Vathys Village (Maria Chatzaki pers. comm.). The habitat where the specimen was found consists of a homogenous degraded phrygana (Mediterranean open scrubland and grass community at low altitude). Approximate GPS coordinates of the location are: 36.9975 26.9607.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="0CAC5929F02AD4FFF4CD3BD09805EC75" pageId="0" pageNumber="71" type="distribution">
<paragraph id="A1426444C90A0EAA5BA3FC51E07A2F19" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="E153A93F4F2E6057EABB8A23B3A9B308" pageId="0" pageNumber="71">
Kalymnos Island (Greece, Dodecanese; Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
<figureCitation id="CAACEF9D35462030B79D773EAAA2C6BA" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
).
</paragraph>
</subSubSection>

372
data/79/90/FB/7990FB77E5105D75BB4FA97D31F18594.xml
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<mods:affiliation id="08DC14E542F774D5017338FD93F94200">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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5.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1E</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8E</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10E</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13D, E</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
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<figureCitation id="127286FE560F73DACCC617DA00B73245" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1E</figureCitation>
<figureCitation id="2696A128378861D55EACBF3D0302354C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8E</figureCitation>
<figureCitation id="1FD0D4AA7A5A0FE059E9210E83EBF619" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10E</figureCitation>
<figureCitation id="D96C5ADAACC7C896E97B52FBC58C8BBC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13D, E</figureCitation>
<figureCitation id="182AB8E312FC5A4E9394A9838522FE28" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="36D590911FC607E2811B515CCD78BE47" pageId="0" pageNumber="71" type="type locality">
<paragraph id="6BE33782730967AE9179E876AEF56B74" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="CF0049431E8F2789975AE03CDC7A471B" pageId="0" pageNumber="71">
Turkey, Van Province,
<normalizedToken originalValue="Gevaş">Gevas</normalizedToken>
<normalizedToken id="C7A173F288F701054F144B716CEB757A" originalValue="Gevaş">Gevas</normalizedToken>
Town environs.
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<subSubSection pageId="0" pageNumber="71" type="note on type locality">
<paragraph pageId="0" pageNumber="71">Note on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<normalizedToken originalValue="Gevaş">Gevas</normalizedToken>
<subSubSection id="6ACBC50DB8F702C3542352DD278231B9" pageId="0" pageNumber="71" type="note on type locality">
<paragraph id="343BEBFB35C91559E9529DB1F9248E91" pageId="0" pageNumber="71">Note on type locality.</paragraph>
<paragraph id="B665010B91021DF8800E830630E5F5C4" pageId="0" pageNumber="71">
<normalizedToken id="84D4E459C4BAD6C26BB6E2E5F672138E" originalValue="Gevaş">Gevas</normalizedToken>
Town is situated on the southern shore of Van Lake. The holotype was perhaps collected in vicinities of the town.
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<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="9EA240491B982E7AD889CDF045842E9E" pageId="0" pageNumber="71" type="type material">
<paragraph id="1A47DED39DC8C0F24C1072EC1F298644" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="246EA9B9933225798DE2F060E5A5392A" pageId="0" pageNumber="71">
<emphasis id="778CE5DE7C16C34B201C4257E8A74FE8" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
: 1♂, 'TURCIA or. Van / Gevas 29.6.1993 / lgt. J.
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<normalizedToken id="B7DEE05417173E282B4B277CE9070337" originalValue="Růžička">Ruzicka</normalizedToken>
//
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="B05B72A796EBE56AA14E01C0D5E56DF5" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ cf. / punctiger / (REICHE &amp; SAULCY) / WRASE det. 2008' (cWR). Paratypes: 1♀, 'TURKEY OR. / GEVAS env., 2100-2600m / (CADIR DAGI) / 1993-06-29,
<normalizedToken originalValue="Klíma">Klima</normalizedToken>
<normalizedToken id="7D7966B3320891D7DBFFE15F978D9F9D" originalValue="Klíma">Klima</normalizedToken>
lgt. //
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="C07D210440AF1889285B3AF615DA3832" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ cf. / punctiger / (REICHE &amp; SAULCY) / WRASE det. 2008' (NMNHS); 1♂, 1♀, 'Resadiye 1900m / TR: Bitlis 6.91 / coll. B.
<normalizedToken originalValue="Lassalle’">Lassalle'</normalizedToken>
<normalizedToken id="9E5820034AB3ED0F3E27A76C0F06B752" originalValue="Lassalle’">Lassalle'</normalizedToken>
(cLAS); 1♀, 'CE TURKEY, prov. Bitlis /
<geoCoordinate degrees="38" direction="north" minutes="37" orientation="latitude" precision="925" value="38.616665">38°37'N</geoCoordinate>
<geoCoordinate id="C18DD8C8F30EAB7C1AFE2F1BCA3E8A60" degrees="38" direction="north" minutes="37" orientation="latitude" precision="925" value="38.616665">38°37'N</geoCoordinate>
,
<geoCoordinate degrees="42" direction="east" minutes="16" orientation="longitude" precision="925" value="42.266666">42°16'E</geoCoordinate>
<geoCoordinate id="141D7D4BE8F51ECD36A6092E474FD494" degrees="42" direction="east" minutes="16" orientation="longitude" precision="925" value="42.266666">42°16'E</geoCoordinate>
2290m /
<normalizedToken originalValue="Şentepe">Sentepe</normalizedToken>
<normalizedToken id="AB841C27BF7FBD232324345243699CCA" originalValue="Şentepe">Sentepe</normalizedToken>
env. / Nemrut Mt. - crater / 24.VI.08 lgt. E.Hajdaj // COLL. / E. &amp;
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<taxonomicName id="CF97137CA8C53AB84BD27507050166D8" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
HAJDAJ /
<normalizedToken originalValue="JEŽOV">JEZOV</normalizedToken>
<normalizedToken id="8A96541BF28EB614356EB046AAB71535" originalValue="JEŽOV">JEZOV</normalizedToken>
/ Czech
<normalizedToken originalValue="Republic’">Republic'</normalizedToken>
<normalizedToken id="B7AC6866719447D88639EADF115C8094" originalValue="Republic’">Republic'</normalizedToken>
(cHAJ); 5♂♂, 2♀♀, 'TR, Bitlis, Yelkenli / Van lake, ca. 1800 m / ~
<geoCoordinate degrees="38" direction="north" minutes="28" orientation="latitude" precision="925" value="38.466667">38°28'N</geoCoordinate>
<geoCoordinate id="0F437F0ED2D63C5A55DF4A758B8D78ED" degrees="38" direction="north" minutes="28" orientation="latitude" precision="925" value="38.466667">38°28'N</geoCoordinate>
,
<geoCoordinate degrees="42" direction="east" minutes="32" orientation="longitude" precision="925" value="42.533333">42°32'E</geoCoordinate>
<geoCoordinate id="C9234CE53B81BEE090F177D6537C57F5" degrees="42" direction="east" minutes="32" orientation="longitude" precision="925" value="42.533333">42°32'E</geoCoordinate>
/ 21.IV.-20.V.2014 / pitfall, leg. C.
<normalizedToken originalValue="Reuter’">Reuter'</normalizedToken>
<normalizedToken id="3C9901E4C2094B75106AF0A61A002663" originalValue="Reuter’">Reuter'</normalizedToken>
(NMNHS, cREU, cWR); 1♀, 'col de Buglan 1500m / TR Mus 6.88 / col. B.
<normalizedToken originalValue="Lassalle’">Lassalle'</normalizedToken>
<normalizedToken id="B48EB3CDEEF8E741E45181184F9C1041" originalValue="Lassalle’">Lassalle'</normalizedToken>
(cLAS); 1♂, 'E Turkey, Mus / 8km SE Solhan, Buglan / Gec., 1700-1800m /
<geoCoordinate degrees="38" direction="north" minutes="56" orientation="latitude" precision="925" value="38.933334">38°56'N</geoCoordinate>
<geoCoordinate id="5859EBB69E968AEEE70B794AA8BDCCD9" degrees="38" direction="north" minutes="56" orientation="latitude" precision="925" value="38.933334">38°56'N</geoCoordinate>
,
<geoCoordinate degrees="41" direction="east" minutes="08" orientation="longitude" precision="925" value="41.133335">41°08'E</geoCoordinate>
<geoCoordinate id="F240B25A6115688CD68B1228E3A8A691" degrees="41" direction="east" minutes="08" orientation="longitude" precision="925" value="41.133335">41°08'E</geoCoordinate>
18. - / 20.VI.05 lgt. E.&amp;
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<normalizedToken originalValue="Hajdaj’">Hajdaj'</normalizedToken>
<taxonomicName id="34C808B38C122DE93FF3DCE95639775E" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<normalizedToken id="EA19548702AFCE1C87429E83C7600898" originalValue="Hajdaj’">Hajdaj'</normalizedToken>
(cHAJ); 3♂♂, 5♀♀, 'TR,
<normalizedToken originalValue="Bingöl">Bingoel</normalizedToken>
<normalizedToken id="1FF4FB702B0D8F368FBB3FD5C2A080A0" originalValue="Bingöl">Bingoel</normalizedToken>
/
<normalizedToken originalValue="Muş">Mus</normalizedToken>
<normalizedToken id="B60D34C811B30903066373C4EFB12661" originalValue="Muş">Mus</normalizedToken>
/
<normalizedToken originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken originalValue="Geçidi">Gecidi</normalizedToken>
<normalizedToken id="3E07581BEE1AC5F4162AB420B47DA42F" originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken id="A56B0D9CC34605102B7659B503E07B0F" originalValue="Geçidi">Gecidi</normalizedToken>
, 1640 m / 21.IV. - 11.V.2014 / pitfall, leg. C.
<normalizedToken originalValue="Reuter’">Reuter'</normalizedToken>
<normalizedToken id="7070940EF0D441F2250A34E8490643F1" originalValue="Reuter’">Reuter'</normalizedToken>
(NMNHS, cREU, cWR).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 22 specimens. TGE: 4♂♂, 2♀♀.</paragraph>
<paragraph id="02ACD96074917EB4DA57E5C04D6C43F0" pageId="0" pageNumber="71">TME: 22 specimens. TGE: 4♂♂, 2♀♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">An adjective derived from the geographical name Van.</paragraph>
<subSubSection id="F6AC246A3B76B025BFADA7B0B42550EA" pageId="0" pageNumber="71" type="etymology">
<paragraph id="F9AF94069CA219E8D19D8A9A19D5B35A" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="07A877074DFEC5C38018817D8D52ED60" pageId="0" pageNumber="71">An adjective derived from the geographical name Van.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="89EF3BDF653056509EBA3B988AF4B85A" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="E3550F2BB026552C000D3EA1CC87B7F1" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="0C3EEC7F9AB49F94E81EE2CC6D10B809" pageId="0" pageNumber="71">
The new species is most closely related to
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
<taxonomicName id="36C57071B27C7949CAC9DB40294131F7" class="Insecta" kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis id="EBA1C897BD9B78FAD4B4D8EBB753B346" italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
</taxonomicName>
sp. nov., but easily differs from it by the apical lamella of the median lobe (dorsal and ventral view) that is less symmetrical and shorter (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10E</figureCitation>
<figureCitation id="C7033418CCA1423BA4285BB4722E4182" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10E</figureCitation>
), conical shape of bursa copulatrix (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13D, E</figureCitation>
<figureCitation id="0CB3E936202603E47CE10AF39D1C7140" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13D, E</figureCitation>
), darker coloration of the integument, elytral striae and striole more coarsely punctate and smaller value PW/
<normalizedToken originalValue="PА">PA</normalizedToken>
<normalizedToken id="74AABAECBCF5313982097F0717A345F4" originalValue="PА">PA</normalizedToken>
(1.32-1.41, vs. 1.41-1.42).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="4A62B98FA6013458501C4C09F08FDD54" pageId="0" pageNumber="71" type="description">
<paragraph id="C6FC41D474926E33683C40209C8BB9D1" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="68F326A994623064FD67D719762D2B9C" pageId="0" pageNumber="71">
<emphasis id="F990C4B510E1376384AA9858A64F848F" bold="true" pageId="0" pageNumber="71">
<emphasis id="351163CB6C30DF27E117C6A932D48439" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of middle size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="205920FA9C8D8FF869DF5C51FC75DCBF" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="EE238B2A21F5CA2A6AE22A46853B1BA6" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 6.85-8.40 mm; BW: 2.25-2.70 mm), with subelongate, slender body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1E</figureCitation>
<figureCitation id="D06EE6F12E903BE2896DB0A49E412744" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="EB5465FCB19B99DC622586976D3F7FE9" bold="true" pageId="0" pageNumber="71">
<emphasis id="F0D5E1C159BDC7E0381160145C2F3E94" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
<tableCitation id="47465F20630E3C12989811E410748FCE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="6EE4CDF262B197426FB47E9DB00180C5" bold="true" pageId="0" pageNumber="71">
<emphasis id="EEF569E729996B7CB8DDCE514848FEE6" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body dorsally and ventrally mostly reddish-brown, appendages (antennae, palpi and legs) lighter colored than body, some specimens from
<normalizedToken originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken originalValue="Geçidi">Gecidi</normalizedToken>
<normalizedToken id="E5A6680B452A782AA553E031B2C7F6FD" originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken id="FA817D20B664B659AAC2F3848983678B" originalValue="Geçidi">Gecidi</normalizedToken>
with elytra almost black, and head, pronotum and ventral surface reddish-brown. Integument rather shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="CCBFB129B8FDBF331B2C0B3384B2792C" bold="true" pageId="0" pageNumber="71">
<emphasis id="EF55B152A4FBAFC6C94338DC0FC3AFFC" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum and elytral intervals 1-5 (rarely also 6-7) without distinct sculpticells or with very faint ones, intervals 6-9 mostly (all specimens from Yelkenli) with distinct isodiametric sculpticells. Ventral surface mostly without visible sculpticells, sometimes with scarcely-visible isodiametric meshes. Pronotum coarsely and densely punctate in basal third and adjacent lateral areas, punctures at sides often reaching medial third. Abdominal ventrites 1-4 punctate at sides.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="78734751893C6C68B6DB5BF575623AC1" bold="true" pageId="0" pageNumber="71">
<emphasis id="27E2133D22A6209ED70A1060FFDF4C9E" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About two thirds as wide as pronotum (PW/HW= 1.30-1.43). Antennae long, with last three antennomeres exceeding base of pronotum. Eyes long, subconvex. Labrum subrectangular, with anterior margin slightly concave. Frontal furrows shallow, subfoveolate. Paraorbital sulci moderately deep, ending slightly before level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="541C4D4638574BEF9FA83C26D4AE4BDB" bold="true" pageId="0" pageNumber="71">
<emphasis id="E0335D30BD84726C7F5E25CE88FAE1F2" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum wider than long (PW/PL= 1.13-1.24), widest point at anterior third. Anterior transverse impression barely distinct, posterior one distinct medially. Sides sinuate, convex anteriorly, concave posteriorly; anterior bead reduced in medial 1/10. Metepisternum slightly longer than wide, MA/MI= 0.87-0.93.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="DC25120E7CFE08234D48B4E1817F4457" bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
. Elongate, about one and two thirds as long as wide (EL/EW= 1.61-1.73), two and a half times as long as pronotum (EL/PL= 2.52-2.74), and one third wider than pronotum (EW/PW= 1.22-1.36), with widest point at medial fifth. Parascutellar striole and striae deeply impressed (specimens from
<normalizedToken originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken originalValue="Geçidi">Gecidi</normalizedToken>
<normalizedToken id="98AF4DD97431CE438437934DC3D514D5" originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken id="FD667B8C461D38393575DF4D1D6D1EAF" originalValue="Geçidi">Gecidi</normalizedToken>
with striae 7-8 less impressed than others); parascutellar striole short, not connected with stria 1. Interval 3 with three discal setiferous punctures. Umbilicate setiferous punctures consisting of 16-17 (rarely 18) each side.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="9899F7860C5A2AC4F4DEB99FCE527F20" bold="true" pageId="0" pageNumber="71">
<emphasis id="5D9C48A7E22F0851C3E11DD152280760" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur with three or four setiferous punctures (three specimens with two on one side) on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="C68ACBA10EEDF53350EE3176FFEB801D" bold="true" pageId="0" pageNumber="71">
<emphasis id="3AC1571A2AB17EBE42B51A6DBA77E599" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view, with elongate basal bulb, long and broad shaft moderately constricted proximally, and short, straight apex; median lobe in ventral view straight, about 3.4 times longer than wide; apical lamella (dorsal view) short, symmetrical, rounded at tip, with sides straight or slightly convex. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8E</figureCitation>
<figureCitation id="66882323D2A8FFD08BA4161F287FEF81" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8E</figureCitation>
) with ventral sclerite distinctly broadened and rounded distally; same in ventral view (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10E</figureCitation>
<figureCitation id="7FF977329F3C3DCC052EC55383D817A3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10E</figureCitation>
), with dorsal sclerite forming a slightly sclerotized and short left-sided protuberance and a convex and long right-sided curve, and ventral sclerite narrow and nearly straight (only just scarcely bent to left).
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="1D331EE56F37B185886CA1C264584B83" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13D, E</figureCitation>
<figureCitation id="92D74496517DE06409E5C484BCE6B173" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13D, E</figureCitation>
). Apical gonocoxite with semi-pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="879195933E73673E23EE74F042821DE2" pageId="0" pageNumber="71" type="habitat">
<paragraph id="3F28540BE3482C7B71225AD937465F39" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="18D8C89258206EF6F6C91EB135AF181C" pageId="0" pageNumber="71">
Christoph Reuter (CR) and Bernard Lassalle (BL) kindly provided us with information on habitat preferences. The specimens found by CR near Yelkenli were collected in a stunted oak forest. Underground is karst rock, in the middle of the tiny valley flows a small stream that still carries water, at least in spring. The series found by CR at
<normalizedToken originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken originalValue="Geçidi">Gecidi</normalizedToken>
<normalizedToken id="546CF792B98690FE0F405F79974968C4" originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken id="884521DFA56783F4BB46DDADFC4AC10C" originalValue="Geçidi">Gecidi</normalizedToken>
, 1640 m, was collected in a semi-open landscape with a low, sparse oak forest of rather small, thin trees, a lot of scrubland, in between meadows, at least in some places, again karst rock, weathered rocks, and a lot of foliage. BL declare that he found a female specimen at
<normalizedToken originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken originalValue="Geçidi">Gecidi</normalizedToken>
<normalizedToken id="5CF3757224EFA6CCC3B79136D02BBFA8" originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken id="13C8F8966CE3DAFC029EFF3B0389B677" originalValue="Geçidi">Gecidi</normalizedToken>
, 1500 m, in &quot;oak bushes&quot;; same stated that, in Resadiye one male and one female of the new species was collected in small oak grove in a little valley.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 6.</emphasis>
<caption id="54CE467E5AE8CA15F4E55F3E84EEF034" doi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71" start="Figure 6" startId="F6">
<paragraph id="CE094207C439E5C2133EDD9BD33CCE37" pageId="0" pageNumber="71">
<emphasis id="8A69C19048D98E2D624D84E77FA8888D" bold="true" pageId="0" pageNumber="71">Figure 6.</emphasis>
Metatarsus, dorsal view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authority="brunneus" authorityName="brunneus Karsch" authorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="Eremoderus" subSpecies="brunneus">Platyderus (Eremoderus) brunneus brunneus</taxonomicName>
<emphasis id="1471D5FAA5A2C5896F9DC5179F25E3A5" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="7C3504C134ED48E6669416B7496019C1" authority="brunneus" authorityName="brunneus Karsch" authorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="Eremoderus" subSpecies="brunneus">Platyderus (Eremoderus) brunneus brunneus</taxonomicName>
Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="B6BFCBB7C02F9BF003468A2B1D48A1E2" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="9362B3BD06036C3361E7578FC9DFFFAC" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
<emphasis id="C790453B8D456A698F7D0C231D297E46" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="EA67C2F2C009FD47E47A58EED937844A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
<emphasis id="F3CE1103C8C44F009A98CF6D0A495E03" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="E84BE2615700CD7E35B2F872FBA1A809" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., right metatarsus, holotype;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="5ED1E47FEE85005B242F720D4952759D" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="A6B321BDE20056B57ED23DDAF0E4703F" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Naẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="06D8F3EB8E8D378363CE20D1B40B19FF" pageId="0" pageNumber="71" type="distribution">
<paragraph id="B613CC6318CD13892EBB91C9672FE2BB" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="C7B6A2546CBB89CB03BFE014222D51CF" pageId="0" pageNumber="71">
Turkey (provinces of
<normalizedToken originalValue="Bingöl">Bingoel</normalizedToken>
<normalizedToken id="ED8A35CFCECF8247B894B7229DE544EA" originalValue="Bingöl">Bingoel</normalizedToken>
and
<normalizedToken originalValue="Muş">Mus</normalizedToken>
<normalizedToken id="A3CCE0F493FBC90B165EE02AFC6394CF" originalValue="Muş">Mus</normalizedToken>
:
<normalizedToken originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken originalValue="Geçidi">Gecidi</normalizedToken>
<normalizedToken id="A93E430FFACEBA6FFFA3E986A74A5638" originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken id="95F05DE3A091F0D52320F7E1FEA2FAB5" originalValue="Geçidi">Gecidi</normalizedToken>
; Bitlis Province; Van Province; Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
<figureCitation id="1774B6143AF8EA5CF70F9F8BA833CCF7" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
). In Bitlis Province, the species inhabits the Nemrut volcanic massif west of Bitlis Town and the mountain region south of the Yelkenli (=
<normalizedToken originalValue="Reşadiye">Resadiye</normalizedToken>
<normalizedToken id="464BDB39868F44C7938E1FF06347D49E" originalValue="Reşadiye">Resadiye</normalizedToken>
)
<normalizedToken originalValue="Bucaği">Bucagi</normalizedToken>
<normalizedToken id="D69AFFC58562B3FD4ED18A81E9450881" originalValue="Bucaği">Bucagi</normalizedToken>
. In Van Province,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="3646B7DE8D7CE447CAFA8B40936ADFBA" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="DDB1B0123369BA233472A833D672CA1C" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. lives around
<normalizedToken originalValue="Gevaş">Gevas</normalizedToken>
<normalizedToken id="56162E8F15B95DC3FA18E448197F4502" originalValue="Gevaş">Gevas</normalizedToken>
Town and in the
<normalizedToken originalValue="Çadir">Cadir</normalizedToken>
<normalizedToken originalValue="Daği">Dagi</normalizedToken>
<normalizedToken id="8C97556E83064F6EA383470955BEA615" originalValue="Çadir">Cadir</normalizedToken>
<normalizedToken id="D30F7734E3F6B1F9132A29EBEDAA22AB" originalValue="Daği">Dagi</normalizedToken>
, a massif situated south of the town aforementioned. In the border area of provinces of
<normalizedToken originalValue="Bingöl">Bingoel</normalizedToken>
<normalizedToken id="0C45AC8923A94802D6AC0F00963DE63A" originalValue="Bingöl">Bingoel</normalizedToken>
and
<normalizedToken originalValue="Muş">Mus</normalizedToken>
<normalizedToken id="40A1609CFE0211A5BB879463C0AC01C4" originalValue="Muş">Mus</normalizedToken>
, it lives along the pass
<normalizedToken originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken originalValue="Geçidi">Gecidi</normalizedToken>
<normalizedToken id="3CE54328108E5A04DDEB476D40E8F233" originalValue="Buğlan">Buglan</normalizedToken>
<normalizedToken id="D929657808DC6868CAD718E1CF24943F" originalValue="Geçidi">Gecidi</normalizedToken>
.
</paragraph>
</subSubSection>

482
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@ -1,424 +1,424 @@
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7.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2B</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5D</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7E</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8G</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10G</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12F</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14A</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
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<figureCitation id="11B9C683B5136CED8A2BDD46C7FD9588" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2B</figureCitation>
<figureCitation id="CD26F40E78DBD9F43E569DE4ED86B574" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5D</figureCitation>
<figureCitation id="1155C97CDA711E3A58264A5C33D3AC92" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7E</figureCitation>
<figureCitation id="3E1F60DCB8EBC44EDFC8CFA4BAE523AF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8G</figureCitation>
<figureCitation id="C5AF6EB20951774F68B5FC87892EB3D4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10G</figureCitation>
<figureCitation id="6F7822DD29A2AB1536C43485DCC0EC95" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12F</figureCitation>
<figureCitation id="387B53161B60B79C9DAB2C4CA39AF18E" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14A</figureCitation>
<figureCitation id="A75FDCFE61AA73FD1EFB4AD04DC2FD56" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Iran, Mazandaran Province, Nur County, between Nur City and Lavij Village, 500-1300 m.</paragraph>
<subSubSection id="001439F6F632E36E7406DA5D171CC29E" pageId="0" pageNumber="71" type="type locality">
<paragraph id="BFF8932B279B951042388D28FA1D564E" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="30BE408D3B02ED8DB1C1F941B7CA07CF" pageId="0" pageNumber="71">Iran, Mazandaran Province, Nur County, between Nur City and Lavij Village, 500-1300 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Bernard Lassalle (pers. comm.) stated that the holotype was caught with soil traps in the period 8-25 June 2000. The place of the exposition of traps was: &quot;south of Nur, on the road between Nur and Lavij, in a mixed forest, between 500-1300 m&quot;. However, having in mind that the northern outskirts of Lavij Village are situated at about 600 m altitude, we consider that the real altitude at which the specimen was caught is between 500 and 600 meters.</paragraph>
<subSubSection id="AD89819659BFAFC07FB897EDC4B64EE0" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="6AFE6B29ABACBFFB6DF884B975D2C08C" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="304079EB7D7A4B962463B0E57B52CF3D" pageId="0" pageNumber="71">Bernard Lassalle (pers. comm.) stated that the holotype was caught with soil traps in the period 8-25 June 2000. The place of the exposition of traps was: &quot;south of Nur, on the road between Nur and Lavij, in a mixed forest, between 500-1300 m&quot;. However, having in mind that the northern outskirts of Lavij Village are situated at about 600 m altitude, we consider that the real altitude at which the specimen was caught is between 500 and 600 meters.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="740DF4C7F9D534DAB0657FE892FEF3F1" pageId="0" pageNumber="71" type="type material">
<paragraph id="BE8A4F9D4B74E188D0C6CA2B5EA9C10A" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="05560A3CF5495072252F134676A64949" pageId="0" pageNumber="71">
<emphasis id="F5011C29CD6FA8F829F55959237737E8" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂, 'sud Nur 500-1300m / IR:Mazanderan 6.00 / coll. B.
<normalizedToken originalValue="Lassalle’">Lassalle'</normalizedToken>
<normalizedToken id="DD0A9DF800498F8A03E8B4B6DA605898" originalValue="Lassalle’">Lassalle'</normalizedToken>
(cLAS). Paratypes: 1♂, 'IRAN, Prov. Mazandaran / [IR08-01] Sari County, / Mohammadabad, Elburz Mts., / N-Slope, NE Sangdeh, 1533m, /
<geoCoordinate degrees="36" direction="north" minutes="04" orientation="latitude" precision="1" seconds="06.6" value="36.0685">36°04'06.6&quot;N</geoCoordinate>
<geoCoordinate id="1F2CC72E7B0C41979E79B4FD29A1C9EB" degrees="36" direction="north" minutes="04" orientation="latitude" precision="1" seconds="06.6" value="36.0685">36°04'06.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="53" direction="east" minutes="09" orientation="longitude" precision="1" seconds="57.8" value="53.166058">53°09'57.8&quot;E</geoCoordinate>
<geoCoordinate id="99C4EE946F59ADC45472543D7B480F75" degrees="53" direction="east" minutes="09" orientation="longitude" precision="1" seconds="57.8" value="53.166058">53°09'57.8&quot;E</geoCoordinate>
, /
<taxonomicName class="Magnoliopsida" family="Carabidae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fagus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
<taxonomicName id="71D220A862AD4BDC79BAF8F49F67BE65" class="Insecta" family="Carabidae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fagus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Fagus</taxonomicName>
forest, leaves debris, / sifted, 29.V.2008, leg. A.
<normalizedToken originalValue="Pütz’">Puetz'</normalizedToken>
<normalizedToken id="8C14D4D8667D0BB6E55C8B698A63E1DC" originalValue="Pütz’">Puetz'</normalizedToken>
(cWR); 1♀, 'IRAN, Prov. Mazandaran / [IR08-03A] Sari County, / Mohammadabad Elburz Mts., / N-Slope, E Qolqol, /
<geoCoordinate degrees="36" direction="north" minutes="10" orientation="latitude" precision="1" seconds="26.7" value="36.174084">36°10'26.7&quot;N</geoCoordinate>
<geoCoordinate id="21ECC3F8A7482F132F09518173630263" degrees="36" direction="north" minutes="10" orientation="latitude" precision="1" seconds="26.7" value="36.174084">36°10'26.7&quot;N</geoCoordinate>
,
<geoCoordinate degrees="53" direction="east" minutes="16" orientation="longitude" precision="1" seconds="29.2" value="53.274776">53°16'29.2&quot;E</geoCoordinate>
<geoCoordinate id="0A5216F40C5265A164E56929E6664ED7" degrees="53" direction="east" minutes="16" orientation="longitude" precision="1" seconds="29.2" value="53.274776">53°16'29.2&quot;E</geoCoordinate>
, / 916m, sifted, 30.V.2008, / leg. A.
<normalizedToken originalValue="Pütz’">Puetz'</normalizedToken>
<normalizedToken id="11325133B9FEB7F1D95745AFA21AF07A" originalValue="Pütz’">Puetz'</normalizedToken>
(cPTZ); 1♂, 2♀♀, 'N.IRAN-Mazandaran prov. / Maji to Vemzela rd. 1360 m /
<geoCoordinate degrees="36" direction="north" minutes="07" orientation="latitude" precision="1" seconds="10.8" value="36.119663">36°07'10.8&quot;N</geoCoordinate>
<geoCoordinate id="09348D4FB6C11CF2F7F9A3512C40A45E" degrees="36" direction="north" minutes="07" orientation="latitude" precision="1" seconds="10.8" value="36.119663">36°07'10.8&quot;N</geoCoordinate>
,
<geoCoordinate degrees="53" direction="east" minutes="11" orientation="longitude" precision="1" seconds="50.9" value="53.19747">53°11'50.9&quot;E</geoCoordinate>
<geoCoordinate id="5AB8865DF78EEAFD7801229D69AC2ECD" degrees="53" direction="east" minutes="11" orientation="longitude" precision="1" seconds="50.9" value="53.19747">53°11'50.9&quot;E</geoCoordinate>
/ 1-5.VI.2018,
<normalizedToken originalValue="Václav">Vaclav</normalizedToken>
<normalizedToken originalValue="Čutka">Cutka</normalizedToken>
<normalizedToken id="C41454E38B66B88F6EAE151DAC7A93B1" originalValue="Václav">Vaclav</normalizedToken>
<normalizedToken id="2E707E440E6612EEC2F2138608EC0067" originalValue="Čutka">Cutka</normalizedToken>
leg.' (NMNHS); 1♂, 1♀, 'N.IRAN-Mazandaran prov. /
<normalizedToken originalValue="Sárí-Talarem">Sari-Talarem</normalizedToken>
<normalizedToken id="BE66AD57C8DBF09CEFD009C153C588E8" originalValue="Sárí-Talarem">Sari-Talarem</normalizedToken>
env,
<geoCoordinate degrees="36" direction="north" minutes="13" orientation="latitude" precision="15" seconds="21" value="36.2225">36°13'21&quot;N</geoCoordinate>
<geoCoordinate id="D52FA94D98D74C4C0D0722E430B17D15" degrees="36" direction="north" minutes="13" orientation="latitude" precision="15" seconds="21" value="36.2225">36°13'21&quot;N</geoCoordinate>
,
<geoCoordinate degrees="53" direction="east" minutes="15" orientation="longitude" precision="1" seconds="49.6" value="53.26378">53°15'49.6&quot;E</geoCoordinate>
<geoCoordinate id="C7B3E655E2F7B3786FEE3CD19D032E61" degrees="53" direction="east" minutes="15" orientation="longitude" precision="1" seconds="49.6" value="53.26378">53°15'49.6&quot;E</geoCoordinate>
, 965m, forest / 1-5.VI.2018,
<normalizedToken originalValue="Václav">Vaclav</normalizedToken>
<normalizedToken originalValue="Čutka">Cutka</normalizedToken>
<normalizedToken id="F9B4FEAF13DF45E162455216F7E9E87A" originalValue="Václav">Vaclav</normalizedToken>
<normalizedToken id="CF41311541F088BA7C9E7E78C849FBD3" originalValue="Čutka">Cutka</normalizedToken>
leg.' (cKME); 1♂, 'N.IRAN-Mazandaran prov. / Galugah-Niala env. 1390 m /
<geoCoordinate degrees="36" direction="north" minutes="37" orientation="latitude" precision="1" seconds="38.8" value="36.62744">36°37'38.8&quot;N</geoCoordinate>
<geoCoordinate id="50E3FEFDEB8CC8776077D3CA3C2BA82F" degrees="36" direction="north" minutes="37" orientation="latitude" precision="1" seconds="38.8" value="36.62744">36°37'38.8&quot;N</geoCoordinate>
,
<geoCoordinate degrees="53" direction="east" minutes="50" orientation="longitude" precision="1" seconds="15.6" value="53.837666">53°50'15.6&quot;E</geoCoordinate>
<geoCoordinate id="18B43DE452ED6291F94D2BAA64A20A3C" degrees="53" direction="east" minutes="50" orientation="longitude" precision="1" seconds="15.6" value="53.837666">53°50'15.6&quot;E</geoCoordinate>
/ 4.VI.2018,
<normalizedToken originalValue="Václav">Vaclav</normalizedToken>
<normalizedToken originalValue="Čutka">Cutka</normalizedToken>
<normalizedToken id="E5E98B97AE442FDA9CA30AF4C78FB7D7" originalValue="Václav">Vaclav</normalizedToken>
<normalizedToken id="D49A4026B15CA84987512B4E187A9A92" originalValue="Čutka">Cutka</normalizedToken>
leg.' (cKME).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 9 specimens. TGE: 5♂♂, 1 ♀.</paragraph>
<paragraph id="08E6D1862ADEE532EE449FC1EEAD5200" pageId="0" pageNumber="71">TME: 9 specimens. TGE: 5♂♂, 1 ♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">Latinized patronym based on the surname of Bernard Lassalle (Boissy-les-Perche, France), whose assiduous efforts in the field contributed substantial numbers of Carabid beetles that are very interesting or new to science.</paragraph>
<subSubSection id="642EDD306E473CAA63F69BEA4E83588C" pageId="0" pageNumber="71" type="etymology">
<paragraph id="FBFCB2C48A2C58EAC0A41E54FC46F5C9" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="4779D999B6A4CD345A9FD3F86ACD80B2" pageId="0" pageNumber="71">Latinized patronym based on the surname of Bernard Lassalle (Boissy-les-Perche, France), whose assiduous efforts in the field contributed substantial numbers of Carabid beetles that are very interesting or new to science.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="B9C965B80752D5BEB9132468CC35C924" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="8AA581380CD7EED5BF03754AE07F43D7" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="9EEE37125AFECD2D99929876F8D526A3" pageId="0" pageNumber="71">
This species is distinct from the other species of the subgenus in the following set of characters: (1) large size of body (8.00-8.50 mm, Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
<tableCitation id="982F13CF94430E39AE308E4D0EA83600" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
); (2) deep black color of body and reddish-brown appendages; (3) dorsal surface of head and pronotum basal foveae with more extensive punctation (incl. micropunctation); (4) median lobe of aedeagus (lateral view) less noticeably curved basally, with apex not turned up.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="37578CDF627B4D7BFE96A61CC9007EDB" pageId="0" pageNumber="71">
In addition to the aforementioned characters, the new species differs from species of the &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="5F7F23D6A9162E1FB92258BD078BA1C4" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="CC0FC36533A02A159995AE976DB9BA27" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; group (which together with
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="E31617E27FC15CE3BA7295157704F67B" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="52D14C9689CF230F6B717596809328BA" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
are the only
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="35C7533B32A4E3B1DEA14D1C5655FC45" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="4D466829610F32750AC1DC47D2E147EF" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
-representatives that inhabit Alborz Range) in the wider pronotum (PW/PL&gt;1.20, vs. PW/PL &lt;1.19) with sides to base more convex, the less long elytra (EL/EW &lt;1.58-1.60, vs. EL/EW&gt; 1.60), and the larger and longer median lobe (1.4 mm long and 4.3-4.4 times longer than wide, measurements taken ventrally). The median lobes of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="C276B32E39D5A48DD26F09308CD220EA" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="B7C43435EC601C5E1CCCB035C943B075" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="4BD44C9B34A871ACFD80F8B740DC8C35" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="1C6B876484F8375D2E6F5D0E284EC427" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
are proportionally smaller and less long (1.1-1.3 mm long and 3.7-3.9 times longer than wide) more clearly curved basally, with apex somewhat turned up (lateral view).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="004D5B816B2076AD6EA4CE6D4A6C5A49" pageId="0" pageNumber="71" type="description">
<paragraph id="33F10679F06B5FC2B76B4F6C24A4A960" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="81AFF2A49B96E6E4648AD40FD94CBE93" pageId="0" pageNumber="71">
<emphasis id="766DA8D2BDB56B7926B2B80B87A5049F" bold="true" pageId="0" pageNumber="71">
<emphasis id="9030FF7ABFE6D8C29A4741F68CA2322E" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="0D057106379CD87A0AD8CA17D485A87A" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="7A3D181A55A70CD26174BD6306E7CFA4" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 8.05-8.50 mm; BW: 2.95-3.05 mm), with suboval and subconvex body (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2B</figureCitation>
<figureCitation id="9FD0BED1FAF949F1F3CBCF7492867BB2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="3BC0D149CED8627A8D0A727F82687789" bold="true" pageId="0" pageNumber="71">
<emphasis id="6C01C470D228AA24ECF14A72A48022C9" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
<tableCitation id="AFFD8BBB2B7B4861E9C7996B74E0A657" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="224EC98F18BB3AB489599DE80AE74FA2" bold="true" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
. Head, pronotum and elytra black, ventral part of body dark brown, appendages distinctly lighter than body, reddish-brown. Integument moderately shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="9B39DF7C8247E8957EB015F56F7B56B6" bold="true" pageId="0" pageNumber="71">
<emphasis id="DC76807AA5F54F3480A2F34D31004BDF" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Disc of head with reduced sculpticells. Pronotum with regular isodiametric sculpticells posterolaterally and stretched isodiametric sculpticells anterolaterally, microsculpture very faint to absent on disc. Ventral surface largely with obsolete microsculpture, slightly stretched isodiametric sculpticells present on mentum, submentum and sternal parts laterally, slightly transverse sculpticells present on sternal part medially; gula, abdominal ventrites and legs without microreticulation. Prosternum laterally punctate.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="67B1FD3895477EE68FE4099D117C0117" bold="true" pageId="0" pageNumber="71">
<emphasis id="6FC92E107D6EBC15706C21BD88BD1D92" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About two thirds as wide as pronotum (mean PW/HW= 1.39). Antennae long, with last three antennomeres exceeding base of pronotum. Eyes large and convex. Labrum subrectangular, with anterior margin slightly concave. Frontal furrows slightly impressed, subfoveolate. Paraorbital sulci moderately deep, ending slightly before level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="04C38312D12576E5B829D8255C4D29D8" bold="true" pageId="0" pageNumber="71">
<emphasis id="9049B3D991833664B4F33585AFE1D78E" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum distinctly wider than long (mean PW/PL= 1.23), widest at second quarter. Anterior transverse impressions barely distinct to indistinct, posterior one well-distinct medially. Sides convex anteriorly, slightly to barely concave posteriorly; bead of anterior margin present laterally, reduced in medial 1/10; bead of posterior margin present throughout (in one paratype reduced to absent in medial 1/8). Metepisternum as long as wide, MA/MI about 1.0.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="9F5E0EBAB47DF60D53483B38AD8AC7DD" bold="true" pageId="0" pageNumber="71">
<emphasis id="E4D546610213DAD5EF02FFA3656187EE" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Suboval, one-and-a-half times as long as wide (mean EL/EW= 1.51), wider and much longer in relation to pronotum (mean EW/PW= 1.38; mean EL/PL= 2.58). Stria 7 reaching or ending before. Elytral interval 3 with three discal setiferous punctures, anterior puncture adjoining stria 3, remaining two punctures adjoining stria 2. Umbilicate setiferous series of 16 punctures on each elytron.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="9530C8E31895CB544047A6CD0F934508" bold="true" pageId="0" pageNumber="71">
<emphasis id="D9F11FE54762A4B8C864FBCC8EF68C3E" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Anterior side of mesofemur ventrally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5D</figureCitation>
<figureCitation id="1F91956197087C943B62AFCFED8E43D1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5D</figureCitation>
) mostly with three-four setiferous punctures (one specimen with five punctures on one mesofemur, whereas another with two such also on one mesofemur). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="6E975D18A4C29DCB646A4BDB8FB9EE9E" bold="true" pageId="0" pageNumber="71">
<emphasis id="D586CED3EEA1B50DEC5EB28F05D83441" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX large, with right shoulder straight to apex and proximal margin widely rounded (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7E</figureCitation>
<figureCitation id="26A7D1EACCE9EFB83B3FA77FB7B19ACF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7E</figureCitation>
). Median lobe of aedeagus in lateral view arcuate ventrally, with apex almost straight; same in ventral view straight, ca. 4.3-4.4 times longer than wide, with apical lamella symmetrical. Internal sac in lateral view well-differentiated, with ventral sclerite long and narrow (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8G</figureCitation>
<figureCitation id="F1D876C79C6FA614662931F51FEB5949" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8G</figureCitation>
), in ventral view with dorsal sclerite forming a short left-sided curve and a long right-sided curve, and ventral sclerite straight, narrow, with distal end curved to left (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10G</figureCitation>
<figureCitation id="AFE9DF518CFD8831BD1CD931F1502C21" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10G</figureCitation>
). Right paramere relatively large, rather concave ventrally (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12F</figureCitation>
<figureCitation id="8A3A036E1E0C2AE514D0A8754ABE1F87" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="0699931E935740B25A42A16E11EF5547" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14A</figureCitation>
<figureCitation id="26999ACBA6F3BAE8CCD419307231230D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14A</figureCitation>
). Apical gonocoxite with semi-pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 8.</emphasis>
<caption id="9624D257E49D62BAC17DECB1B3639AF4" doi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71" start="Figure 8" startId="F8">
<paragraph id="572A935EABE2A3213E9B44840D2A9CC6" pageId="0" pageNumber="71">
<emphasis id="615096752F54A0C139C1FA24F39FD1A3" bold="true" pageId="0" pageNumber="71">Figure 8.</emphasis>
Median lobe of aedeagus, left lateral view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
<emphasis id="5722781FD6C946F83BED0770D6CF76B1" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="E89677D56A00743399FF954C906D1AB1" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<emphasis id="211BA605969EBE83D60D707DFA6F64AC" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="ECBF0377D2A9E22E29892EE28F571F5D" class="Insecta" kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken id="3C3FA7CACFF3713958ACCAA4153B546F" originalValue="Mařan">Maran</normalizedToken>
, 1940, lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
<emphasis id="627FD46587E27AB3EEB5C6C55B1DCA53" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="F141AFEF6B6BD4231B45B3F22C032CF6" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
<emphasis id="C29C45F8E40584CBA625EC49E3CEA434" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="538D263A26B59F173A4E253759649775" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
<emphasis id="7CE860E0B50CFC4371B0CF6989F4B006" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="5FD5109D7104DE96BF66C42C33B621B9" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
<emphasis id="9A1DB848418B622AAA4B0FCA78055C04" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="7794EB21125B494030E1C5D25B9FC922" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
<emphasis id="53E6AA7727D7BE0D9ED837129622C55D" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="69AE63D3061A00F7F3FBC6BD0E371166" class="Insecta" kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
<emphasis id="3EEACA2CF88DF94A8EFA1EE2FF08D617" bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName id="9C31D0CA5771BCBAFCA59CFA6A53B660" class="Insecta" kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
<emphasis id="BC0F6654A78FE3D3EC8CBDBDAAB1CD45" bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName id="F4E9A6433FFAC894EA1A8CC936B8EBC7" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, male specimen, Tochal, Tehran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName authority="" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">P. (Platyderus)</taxonomicName>
<emphasis id="1F380755FA1713D4C30B1CC91B44DE60" bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName id="C325A35D3D2E1E3747D0354C2816BD14" authority="" class="Insecta" kingdom="Animalia" lsidName="P. (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">P. (Platyderus)</taxonomicName>
sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="1B9EBD314FB5A4C8F8B80FB7EA25D613" pageId="0" pageNumber="71" type="comparisons">
<paragraph id="EEE9A5852F9137C42D81A88824321F22" pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph id="4A7BA03F90C9C70F9B9BC29A18E8B211" pageId="0" pageNumber="71">
By dorsal coloration, form of pronotum and elytra, the new species is similar to
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. reticulatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis italics="true" pageId="0" pageNumber="71">P. reticulatus</emphasis>
<taxonomicName id="F73C6DAE2B5B98480128412DF5193130" class="Insecta" kingdom="Animalia" lsidName="P. reticulatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="CCA72E4A5AC545331E7CB15899586541" italics="true" pageId="0" pageNumber="71">P. reticulatus</emphasis>
</taxonomicName>
(Chaudoir, 1842), the only member of the nominotypical subgenus that lives in sympatry with
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="92465E6F1CEDE6FA470C2A4CF796C02C" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="F36484AAA74E4F00630A12DAE0AA96BA" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
on north slopes of the Alborz Range. However, the latter has the anterior side of mesofemur ventrally mostly with three-four setae (vs. two such setae in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. reticulatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis italics="true" pageId="0" pageNumber="71">P. reticulatus</emphasis>
<taxonomicName id="93F6F24F2F364D0F235684527AF32783" class="Insecta" kingdom="Animalia" lsidName="P. reticulatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="21FDBBBFC3FF815E8926DDF979A3CFE8" italics="true" pageId="0" pageNumber="71">P. reticulatus</emphasis>
</taxonomicName>
) as well as completely different shapes of the urite and median lobe of aedeagus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="727F842DC3B574DC8AD4E2E043A585C5" pageId="0" pageNumber="71" type="habitat">
<paragraph id="E389C7D64A723B09ABD49C289796DA9E" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="B8199A67428DEDA5387D12A2FA6A6D26" pageId="0" pageNumber="71">
According to the label data, the species occurs on northern slopes in
<taxonomicName authorityName="Lipsky" authorityYear="1898" class="Magnoliopsida" family="Carabidae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fagus orientalis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="71">Fagus orientalis</emphasis>
<taxonomicName id="4A029FC23E2CEDD226AE453C2C0101C5" authorityName="Lipsky" authorityYear="1898" class="Insecta" family="Carabidae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fagus orientalis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="4B62CC732F6939CAAF907A7AAE72CACA" italics="true" pageId="0" pageNumber="71">Fagus orientalis</emphasis>
</taxonomicName>
or mixed forests in altitudes of about 500-1600 m a.s.l. Specimens were collected by sifting leaves debris or with pitfall traps.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="4E050A5EE3F9F8382148417556A2C581" pageId="0" pageNumber="71" type="distribution">
<paragraph id="1E5B2DA8AD4C10E2073F198CD5E81C7D" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="C23CFA01BA22D9D0738EE7B4DB760C99" pageId="0" pageNumber="71">
North Iran: Central Alborz Range (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="0C2057A8CCC70E860D51956FB47014F6" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
). The three records known until today come from three counties of the Province of Mazandaran, Nur County, Sary County and Galugah County. The areas between these counties are probably also populated by
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="A0DAF7963771007B9F6113777F54F13D" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="143927679B57555E594C13AEBF3AA63F" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="6D275A24C778DDBD6F9AF378FC22A843" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<emphasis id="CE06400466B6635E3B8FC51CA2EE0ABC" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="D76DC01CECD50FCE501FC3AEE40611A7" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="18A0B6944E2A516A23BCF45F42890031" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; species group
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
<paragraph id="C502D80ECFD2687576896B72F19022FA" pageId="0" pageNumber="71">
<emphasis id="CC129610146DA6D5A8F30D427EA1A0BD" bold="true" pageId="0" pageNumber="71">Diagnosis.</emphasis>
This group includes
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
<taxonomicName id="33D50E26967D3997CD5F6352694BBE77" class="Insecta" kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="A7AF483E389FF7E6255400AE3B074BCA" italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="F111C2490586088B1E62F45A3815CAB7" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="8DEA4AD0EF6DF8394E5AF29CCC4C28A2" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="C4A1752FE0516E09BFE6FBAE47EFA7FD" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="E8131C6019A780574C0DF04288CA112E" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
<taxonomicName id="5DF37CFCC80762F981EAEC9296B23666" class="Insecta" kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="737B4E3A085BF0D239A41B3994DD60D0" italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
</taxonomicName>
sp. nov., which are adapted to inhabit higher altitudes and open habitats in the Alborz Range (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="6C22646F228A869F11495693855CF818" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
). Compared with
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<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
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<emphasis id="DF0912BF7E4BA8A4CF3079B0AB64BE38" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
sp. nov., the species in question have a more elongate and parallel body of smaller size and lighter color of the integument (see also
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<normalizedToken id="25CB167919D3520CC95F7ED224C23D36" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
under &quot;
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
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<emphasis id="2F2EA7A51784AE2B3194CB6563246B8C" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
</taxonomicName>
&quot; species group).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="563E33070626379EE82C1533E1CBFB00" pageId="0" pageNumber="71">
The species from the &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="3761A67517D462B41690F62686EE3B20" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="5F6DAFCF83F2E09471E757E66D1778C8" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; group differs from those of the &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
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<emphasis id="9B7A965747C74480FF3828A2CFCFD626" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
&quot; subgroup, in: (1) the sides of pronotum are clearly concave toward the posterior angles (vs. sides of pronotum hardly concave or straight toward the posterior angles), (2) PW/PL &lt;1.20 (vs. PW/PL&gt; 1.22), and (3) EL/EW ≥ 1.63 (vs. EL/EW ≤ 1.63).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Habitat.</emphasis>
<paragraph id="2A1D20F5D91ED264A353F82B0AE63F70" pageId="0" pageNumber="71">
<emphasis id="CE0F6D3A07255F59EBF775608375EE9A" bold="true" pageId="0" pageNumber="71">Habitat.</emphasis>
The habitat of the species of the &quot;
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="1179AAA74010B6AD1AB0628A4D2EEC96" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="710CBB68F9FB585F897644786DE66783" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
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&quot; group is similar to that of the two Iranian species from the Zagros Mountains (see
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<normalizedToken id="30BE8583BC1D4D4A95ACFB0E9F6774C2" originalValue="“Habitat”">&quot;Habitat&quot;</normalizedToken>
under &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<taxonomicName id="90E4929C31BF5DF8CAC54E3F15256958" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="F9CC9EBC1D9C91B34C02A9A7FB0245DF" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
<taxonomicName id="311AD924334F4EE505D5F78FBE05B85B" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis id="3CE7C6874F9F83D07E947D8E7CBEB2D7" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
</taxonomicName>
&quot; species group). In the high mountain zone of Elburz the
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="7EE8DA5D4697EB7AB01EA91787AE0261" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="E6CE2C2713B95FF2434777664F39B84A" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
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-species were collected in natural mountain pastures, covered with snow in winter.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
<paragraph id="AB4DD53E1C48BFB7739E3729201C0FB9" pageId="0" pageNumber="71">
<emphasis id="57FA7403FBC054D8FBFA2D7940D0E965" bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
Having not been able to examine the type material of Pierre Morvan, we treated
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
<taxonomicName id="10963A8A546E906D7343318BAB91AD0C" class="Insecta" kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="2057BA6B815E6E027612DAA3B9F2F383" italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
</taxonomicName>
Morvan, 1970,
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<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="6B07D214CC0ABCC2B5F51886EA3C3A0B" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="2B9515C2D9F4E10FA703D80D29A066E8" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
Morvan, 1974 and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="C004A3848F6E86EDCDA74F2BEFED5FE7" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="AC125BBB304625CBF83257A9E131E059" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
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Morvan, 1974 as valid species. Based on original descriptions and new material at hand, we were able to ascertain the identity of the last two species. Without available material for study, the identity of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
<taxonomicName id="1094F8C474C4FFC731F5BC4B7C6840B8" class="Insecta" kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="19418026487C8B5698303F5972297104" italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
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remains unknown for the time being.
</paragraph>

94
data/99/91/BE/9991BEC4035105F87458E370E3B3727C.xml
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@ -1,63 +1,63 @@
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@ -132,7 +132,7 @@ Benson: Journal of the Asiatic Society of Bengal, 5: 351. [not specified. &quot;
1878
<taxonomicName id="3F5FE5C507AE81E02147E1A85FF7ECDA" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Plectopylis) plectostoma" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="plectostoma" subGenus="Plectopylis">Helix (Plectopylis) plectostoma</taxonomicName>
, - Nevill: Hand list of
<taxonomicName id="E1DC0CDBA8D44E5D7563509F5CF8B373" lsidName="Helicoidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
<taxonomicName id="E1DC0CDBA8D44E5D7563509F5CF8B373" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helicoidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
in the Indian Museum
<normalizedToken id="ECEE27FF989BF8B41B5362E610124157" originalValue="…">...</normalizedToken>
: 1: 71. [
@ -212,11 +212,11 @@ Hills&quot;, &quot;Bassein, &amp;c., Pegu&quot;,
<paragraph id="01D107447E11BF08785BEC91B7D72DC6" pageId="37" pageNumber="38">Types:.</paragraph>
<paragraph id="91327B808EFB86DDF07F289EED36B7FB" pageId="37" pageNumber="38">
Darjeeling, coll. MacAndrew ex coll. Benson, UMZC 102160 (7 syntypes of
<taxonomicName id="26EC4BDAB854C7D62065DF1E6799F6E2" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea plectostoma" order="Cycloneritida" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="plectostoma">plectostoma</taxonomicName>
<taxonomicName id="26EC4BDAB854C7D62065DF1E6799F6E2" genus="Helicoidea" lsidName="Helicoidea plectostoma" pageId="37" pageNumber="38" rank="species" species="plectostoma">plectostoma</taxonomicName>
); Darjeeling, coll. MacAndrew ex coll. Benson, UMZC 102155 (1 syntype of
<taxonomicName id="9AE742FD3A26A6EE803A18C113183697" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea plectostoma" order="Cycloneritida" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="plectostoma">plectostoma</taxonomicName>
<taxonomicName id="9AE742FD3A26A6EE803A18C113183697" genus="Helicoidea" lsidName="Helicoidea plectostoma" pageId="37" pageNumber="38" rank="species" species="plectostoma">plectostoma</taxonomicName>
, Figure 13A); Bengal, coll. MacAndrew ex coll. Benson, UMZC 102156 (3 syntypes of
<taxonomicName id="E330D155D27EBAC3CA421E50DFEBD107" class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea plectostoma" order="Cycloneritida" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="plectostoma">plectostoma</taxonomicName>
<taxonomicName id="E330D155D27EBAC3CA421E50DFEBD107" genus="Helicoidea" lsidName="Helicoidea plectostoma" pageId="37" pageNumber="38" rank="species" species="plectostoma">plectostoma</taxonomicName>
).
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@ -351,7 +351,7 @@ Museum samples are labelled from several locations. This species is probably wid
<paragraph id="F8177D920E9ED7B2C0EFD5ED5C941DC7" pageId="40" pageNumber="41">Remarks.</paragraph>
<paragraph id="FFFA64F7ED6731AB43D9A7A1A13B480D" pageId="40" pageNumber="41">
The name &quot;
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<taxonomicName id="D02D130FFA5562BD2DADDB78D337F80A" genus="Helicoidea" lsidName="Helicoidea prodigium" pageId="40" pageNumber="41" rank="species" species="prodigium">prodigium</taxonomicName>
Benson&quot; probably refers to
<taxonomicName id="BD372258525673E85C16C9336E29CBDD" class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella plectostoma" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="plectostoma">Endothyrella plectostoma</taxonomicName>
. It is a manuscript name, which was mentioned several times in the literature (

422
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<paragraph id="15B66FF0EE29CFCE45BDA51453ACF6F4" pageId="0" pageNumber="71">
18.
<taxonomicName LSID="http://zoobank.org/A95FB1A6-43F7-4FFC-A202-05073CC72F90" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) jordanensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<taxonomicName id="5536E0A4C18A6E8D040A0A8CE85B8B92" LSID="http://zoobank.org/A95FB1A6-43F7-4FFC-A202-05073CC72F90" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) jordanensis</taxonomicName>
<taxonomicNameLabel id="CD4A51EADB9B77E79A082E8D2619D7B6" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4C</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6D</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7K</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9F</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11E</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12L</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15E</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
<subSubSection id="75AC2E195E2750BC6C9B5FD4E737EE82" pageId="0" pageNumber="71" type="description">
<paragraph id="1BDD4E6DE059C0A79A029613B6C3B4C0" pageId="0" pageNumber="71">
<figureCitation id="D740A7218F3203A9EADDEF68DDECFDD8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4C</figureCitation>
<figureCitation id="554FF20CDEC9A0F1D1C400B57D4937FF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6D</figureCitation>
<figureCitation id="E58512A772DE56D34E3EFC62F02039E8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7K</figureCitation>
<figureCitation id="A4B079B1C3BCE029E81B4B712244FEDA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9F</figureCitation>
<figureCitation id="FD41A09C2A1BA88F2982DD1151924F94" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11E</figureCitation>
<figureCitation id="8B605D179201F893F040625504BA659B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12L</figureCitation>
<figureCitation id="E786FAC1E5108AB5E793837E58DE2D8A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15E</figureCitation>
<figureCitation id="E5CFC5D6C5FE803022487299D2874D8D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="A58595AAE648CADC2E462787418BD75C" pageId="0" pageNumber="71" type="type locality">
<paragraph id="01A6122F966CC79843E241C5D2D75772" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="904BE4063025F30E649104861E38C53F" pageId="0" pageNumber="71">
Jordan,
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
<normalizedToken id="AC1A5F3369F9C60C75D6555E815E5F6E" originalValue="Ma’an">Ma'an</normalizedToken>
Governorate,
<normalizedToken originalValue="Al-Betrā’">Al-Betra'</normalizedToken>
<normalizedToken id="3D37314762633E74978CED9AA0DA27B8" originalValue="Al-Betrā’">Al-Betra'</normalizedToken>
District, Little Petra, archaeological area, 30.3667, 35.4333.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="643CBAACA18958D455D521E1D9AAB299" pageId="0" pageNumber="71" type="type material">
<paragraph id="312118DEAF029B1766881243DEB0FAB3" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="480F23427972376FD5C9855A59C51B52" pageId="0" pageNumber="71">
<emphasis id="1F6E41DF750DFB85843E121F4EF6550E" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂, 'JORDAN (
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
<normalizedToken id="A475E49587029C00D853398880F2F0B4" originalValue="Ma’an">Ma'an</normalizedToken>
) / Little Petra, ca. 1000 m / archaeological area /
<geoCoordinate degrees="30" direction="north" minutes="22" orientation="latitude" precision="925" value="30.366667">30°22'N</geoCoordinate>
<geoCoordinate id="5ED697446003176516DB25DA26EEF967" degrees="30" direction="north" minutes="22" orientation="latitude" precision="925" value="30.366667">30°22'N</geoCoordinate>
/
<geoCoordinate degrees="35" direction="east" minutes="26" orientation="longitude" precision="925" value="35.433334">35°26'E</geoCoordinate>
<geoCoordinate id="2FC3BD2D078CE0DE65E06EA16FFF1F8F" degrees="35" direction="east" minutes="26" orientation="longitude" precision="925" value="35.433334">35°26'E</geoCoordinate>
/ (narrow shaft beside / stony staircase) / 25.III.2016 Wrase &amp; Laser [12B] [w, p]' (cWR). Paratypes: 1♀, 'Jordanien 1024 m / Prov
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
<normalizedToken id="69ACD8AE49BD0E95A1D8F3201212B2F8" originalValue="Ma’an">Ma'an</normalizedToken>
/ W. Ziegler 9.3.2015 [w, p] // N Wadi Musa / 30°24'.15&quot;N, 35°26'.53&quot;E [w, p]' (cZIEG); 1♂, 'Jordania / Prov.
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
<normalizedToken id="518264D0A9252BE46B879EB4F4D5C234" originalValue="Ma’an">Ma'an</normalizedToken>
/ n Wadi Musa 1023m / W.Ziegler 25.3.2016 [w, p] // Jabal al Bayda (Project / 30°24'291&quot;N, 35°26'743&quot;E [w, p]' (NMNHS); 1♂ 'S-Jordan: Wadi Rum / Qatar Spring /
<geoCoordinate degrees="29.51" direction="north" orientation="latitude" precision="555" value="29.51">29.51°N</geoCoordinate>
<geoCoordinate id="82C74598DA026D17F6767C98F8D96AFE" degrees="29.51" direction="north" orientation="latitude" precision="555" value="29.51">29.51°N</geoCoordinate>
/
<geoCoordinate degrees="35.41" direction="east" orientation="longitude" precision="555" value="35.41">35.41°E</geoCoordinate>
<geoCoordinate id="995E56C72EEDCAB947B7577311940DCA" degrees="35.41" direction="east" orientation="longitude" precision="555" value="35.41">35.41°E</geoCoordinate>
/ 1000-1100 m asl [w, p] // dry wadi with
<taxonomicName class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
<taxonomicName id="1990C5A0A484344FEEF754BF46B33342" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="5DE71A666BCF7076DB3E2CC939149CDC" italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
</taxonomicName>
trees, 21.III.2017 / leg. Th. Assmann [w, p]&quot; (cAL); 1♂,
<normalizedToken originalValue="ʻS-Jordan">ʻS-Jordan</normalizedToken>
<normalizedToken id="C0127CEDE6215AC464F5471AF068A6A2" originalValue="ʻS-Jordan">ʻS-Jordan</normalizedToken>
: Little Petra, ca. 1000m / wadis with Mediterranean trees in vicinity to the archaeological site /
<geoCoordinate degrees="30.34" direction="north" orientation="latitude" precision="555" value="30.34">30.34°N</geoCoordinate>
<geoCoordinate id="C2D45A52336D17D01BE46FFD6BED44CF" degrees="30.34" direction="north" orientation="latitude" precision="555" value="30.34">30.34°N</geoCoordinate>
,
<geoCoordinate degrees="035.46" direction="east" orientation="longitude" precision="555" value="35.46">035.46°E</geoCoordinate>
<geoCoordinate id="996F10970C7B88F96FD82E240FD5CCAB" degrees="035.46" direction="east" orientation="longitude" precision="555" value="35.46">035.46°E</geoCoordinate>
/ 25.III.2016 / leg. Th. Assmann [w, p]' (cAL).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 5 specimens. TGE: 4♂♂, 1♀.</paragraph>
<paragraph id="B19E00D01EE6F53C54C3E9CC9CFAA167" pageId="0" pageNumber="71">TME: 5 specimens. TGE: 4♂♂, 1♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, based on the name of the country in which this species was found.</paragraph>
<subSubSection id="19F898B408994978C067A86A66802474" pageId="0" pageNumber="71" type="etymology">
<paragraph id="A1EA26A91B7A584676CC9C3DF4202319" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="25170663BBD0F7FFD330341498E12C61" pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, based on the name of the country in which this species was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="EA391AF010259535EB9CA92FD42C1730" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="0C1BE937956BBA3B96A30813520E70B6" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="A798CE5A067304A4EEFE3392AD38B224" pageId="0" pageNumber="71">
Differs from all other species of the &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="2E89BE732E04E98CBE998B9BDEF37D3A" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="BABB02B9BE7FF5CA8AF3ABD88D4838A9" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group by the following set of characters: (1) meso- and metatarsomeres dorsally convex and smooth (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6D</figureCitation>
<figureCitation id="991DDED325AFCE75B11638214EA0EE87" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6D</figureCitation>
); (2) posterior angles of pronotum incompletely rounded, with tips protruding laterally (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
<figureCitation id="37B9A0F1AA0A915B1B0BD6CFA1C49FBC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
); and (3) ventral margin of apex of median lobe (lateral view) evidently convex thus apex appears bent up (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
<figureCitation id="552D7F7BF2C325EBFDEFDA50B6E6F6CF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus jordanensis</emphasis>
<paragraph id="489FAB09CA961FC983491EEC955FFAB8" pageId="0" pageNumber="71">
<taxonomicName id="0E3DC674BA05A678FBE11C4222BB3D12" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="67F57B8A3A5189B89C55BB8C0775EEB8" italics="true" pageId="0" pageNumber="71">Platyderus jordanensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="6F7F8BDF7D316733210BBA953065C162" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="91410F42895E5A2B35D778AC3F912FB4" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
share a trait that can be a mark for their close relationships, the apical gonocoxite without nematiform setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="3D19800C3BDCF03A2AB7A9B33AD9D048" pageId="0" pageNumber="71" type="description">
<paragraph id="AC5E6E7168E1B9AE71767BB1DE8BE0D8" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="C2550C5F1249F65300F7061735590E97" pageId="0" pageNumber="71">
<emphasis id="8F88887C869182D9CC3D76F3F8136980" bold="true" pageId="0" pageNumber="71">
<emphasis id="A438108C24AD4F64F004F27C43DF93C7" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="99B144BAF2E8D9B2A3BF2A4C4EC1FE92" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="30CF337387E1F6E84761716B426777FA" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 8.70-9.90 mm; BW: 3.00-3.45 mm), with rather elongate and moderately convex body (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
<figureCitation id="D3B5D2304BD998C1C81983E561D292AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="44DBAEA864FDC38106F971123BCFEE87" bold="true" pageId="0" pageNumber="71">
<emphasis id="6EE1A2EF4FAB99671F24309F616CD1EC" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
<tableCitation id="BC39A972EC8B6321A2161863C03D363C" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="D5BB185DE6DB3E827CFA501C4B63E446" bold="true" pageId="0" pageNumber="71">
<emphasis id="D297A9C5028A717B09EEC0051B60B359" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly light reddish-brown, palpomeres and legs somewhat lighter than rest of body. Integument slightly to moderately shiny, head and pronotum shinier than elytra.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="6073F1E542EF6751C6702FD7E8F74FC2" bold="true" pageId="0" pageNumber="71">
<emphasis id="F7F7FC4779D10D0C8CCDE822658E4BE3" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Surface of pronotum with evident microreticulation, sculpticells regular isodiametric. Elytra (intervals, scutellum, basal margin and lateral gutter) with distinct isodiametric sculpticells. Ventral surface largely with more or less impressed isodiametric or slight transverse sculpticells. Head impunctate, nearly smooth, with a few, fine and shallow wrinkles in and laterally of frontal furrows. Pronotum surface mostly smooth, only basal area between foveae longitudinally wrinkled and adjacent lateral areas with rare and superficial punctures that do not reach anterior half. Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled medially, smooth laterally, other ventrites neither wrinkled nor punctate.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="BF496CA73DE520734FACCFF616B1C146" bold="true" pageId="0" pageNumber="71">
<emphasis id="B1216CDB5EF66AFB60127142A5A7514A" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
One-third or more as wide as pronotum wide (PW/HW= 1.35-1.42). Eyes long, moderately convex. Labrum subrectangular, shorter than clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows small, punctiform. Paraorbital sulci straight, backward surpassing posterior margin of eye, hardly reaching level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="FC80E526AD50D089377B8668073EE74C" bold="true" pageId="0" pageNumber="71">
<emphasis id="563FF8AE512B8FF198D7B3EA7B403871" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about a tenth wider than long (PW/PL= 1.08-1.13), with widest point at medial third. Anterior transverse impression absent; posterior transverse impression barely distinct. Sides sinuate, convex medially and anteriorly, slightly concave posteriorly; anterior bead present laterally, lacking or present in medial 1/10; basal bead present laterally, reduced to absent in medial third or medial half. Metepisternum somewhat longer than wide, MA/MI= 0.88-0.91.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="22E683FDFE2F117E9C90D93B26329F74" bold="true" pageId="0" pageNumber="71">
<emphasis id="F49567D1064B260DE4F2F3FBF34A9D14" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and a half times or little more as long as elytra width (EL/EW= 1.56-1.60), two-and-a-half-times as long as pronotum (EL/PL= 2.41-2.60), and a bit less than a half as wide as pronotum (EW/PW= 1.39-1.46), with widest point at medial third. Parascutellar striole linear, as impressed as other striae, not joining stria 1; striae 1-8 and striole slightly to indistinctly punctate; base of stria 1 ending in parascutellar pore, striae 2-5 reaching or barely not reaching basal bead, 6 and 7 ending little before basal bead. Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 punctures on each elytron in three specimens examined, including holotype.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="D5EDBE95AE75A76BC985E9447FCBF806" bold="true" pageId="0" pageNumber="71">
<emphasis id="403DB74F2F190105C146A8CE0C283195" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one or two setae in basal third and one in medial third. Mesofemora with 6 (holotype) or 5 (both paratypes) setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="685FB4E07941462CB4BD05FCD29FC649" bold="true" pageId="0" pageNumber="71">
<emphasis id="C30B7CD030BD21632A6C5325AE377C32" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX subtriangular, with proximal margin symmetrical and pointed at apex (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7K</figureCitation>
<figureCitation id="3E9DB51B4AD7168E711143AFB041D2CD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7K</figureCitation>
). Median lobe of aedeagus in lateral view slender, with narrow basal bulb, shaft broad with proximal part slightly constricted, and short, straight apex (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
<figureCitation id="B574ACB2686647EA04ADD9CC1DD28291" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
); median lobe in ventral view straight, 3.7 times longer than wide; apical lamella (dorsal view) rather short, nearly symmetrical, rounded at tip, with sides straight to slightly concave. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
<figureCitation id="897D6E91513A2C276C582CE551DBDC8F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9F</figureCitation>
) with ventral sclerite slightly broadened and rounded distally; same in ventral view (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11E</figureCitation>
<figureCitation id="2E916BB7EE326A1CB84FA60E2BC5C4C3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11E</figureCitation>
), with dorsal sclerite large, having small left-sided and large right-sided protuberance, and ventral sclerite straight and rather narrow. Right paramere as on Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12L</figureCitation>
<figureCitation id="A6AF9C5FBE8D5CCF98BEE81105BFF403" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12L</figureCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="9838D52B0C3698E9BF3BF8E3A930289B" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15E</figureCitation>
<figureCitation id="9423DE90DECE2A7F20F131EAE3E7B66F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15E</figureCitation>
). Apical gonocoxite with widely rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="20B84D72FD16B24E6F2391C66F54F6B2" pageId="0" pageNumber="71" type="comparisons">
<paragraph id="BF2CC1D2D310383B19D40122303E4840" pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph id="4F2687583811CC63E2DC3BC706AB700C" pageId="0" pageNumber="71">
In additions to characters mentioned in
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="E0B13E9DDDE8372BF096C0C0354E44F7" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="8AFB5084DE21D67229A5B7F04D5FC0AA" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="FFA496FDF87CC1C46AC9BC69C5508C42" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. differs from geographically close
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferranei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferranei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferranei</emphasis>
<taxonomicName id="DDA64799775BC6EABC57B14B5E0067A8" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. ferranei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferranei">
<emphasis id="14DC3CACD978E4D14F0C8F46E6A1A796" italics="true" pageId="0" pageNumber="71">P. brunneus ferranei</emphasis>
</taxonomicName>
by pronotum less wide (PW/PL: 1.08-1.13, vs. PW/PL: 1.14-1.21), with anterior margin narrower than posterior margin (PA/PB: 0.78-0.87, vs. PA/PB: 0.88-0.92).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="1E099A3323331FAF86CE28FE26814F74" pageId="0" pageNumber="71">
For differences between
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="DD858AA964EEDF06BB96AD9FB1842A03" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="7ADC2EF0239B8598EA83F6F43A2E9985" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
, from one side, and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="E14BAA8F709ED9187E99838CA21037AD" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="A66F73656DD8F0E924E02049472DA457" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="541801AA0B039C6F334F400EDE38ECA7" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="D97553E51ED7968AB8E6889545D5900B" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="A1D1004FAD66E60AA305822A96258D22" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="2B5825E75B3C16AB50D78CC88D658642" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
, from other side, see sections
<normalizedToken originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
<normalizedToken id="B25D1371AEF60DDFC70743C0E71274A2" originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
(under the first two species) and
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="DE1C19CBE62FD77774CBD23BEF8DB2AD" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
(under
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="ABE20D5AD94634996C3DDD5E2E67FC55" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="AE24CA45E6517435B963D56741CEC9A6" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="A800C22679152010F8BF8894D160F344" pageId="0" pageNumber="71" type="habitat">
<paragraph id="F7FD875D869B3A03230F7BF808DD841C" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="0E2520D5ABD0644499E20F265F4FFA7C" pageId="0" pageNumber="71">
In Little Petra
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanansis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanansis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanansis</emphasis>
<taxonomicName id="11A9A4F96641CD61FABFED298F7288F4" class="Insecta" kingdom="Animalia" lsidName="P. jordanansis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanansis">
<emphasis id="F497385A380149D1F95B7E587709D4BC" italics="true" pageId="0" pageNumber="71">P. jordanansis</emphasis>
</taxonomicName>
sp. nov. lives in wadis that have a canyon-like profile and some Mediterranean tree species. There it lives together with
<taxonomicName class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus quadricollisquadricollis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="quadricollisquadricollis">
<emphasis italics="true" pageId="0" pageNumber="71">Laemostenus quadricollisquadricollis</emphasis>
<taxonomicName id="D9EE775D0B548ECE51EEFEC1EC0A6CBA" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus quadricollisquadricollis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="quadricollisquadricollis">
<emphasis id="E5005306A3063C0BF6EBD81A42DF9E28" italics="true" pageId="0" pageNumber="71">Laemostenus quadricollisquadricollis</emphasis>
</taxonomicName>
L. Redtenbacher, 1843,
<taxonomicName authorityName="Piochard de la Brulerie" authorityYear="1876" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus crucifer" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="crucifer">
<emphasis italics="true" pageId="0" pageNumber="71">Trechus crucifer</emphasis>
<taxonomicName id="6E24631D8E9EBCC28DAC72D2C88CA526" authorityName="Piochard de la Brulerie" authorityYear="1876" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus crucifer" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="crucifer">
<emphasis id="7BCD112ED0A593C41FC314E87E405A5E" italics="true" pageId="0" pageNumber="71">Trechus crucifer</emphasis>
</taxonomicName>
Piochard de la
<normalizedToken originalValue="Brûlerie">Brulerie</normalizedToken>
<normalizedToken id="81517CA01572C7D9284FAA998F969C76" originalValue="Brûlerie">Brulerie</normalizedToken>
, 1876,
<taxonomicName authorityName="Motschulsky" authorityYear="1850" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" lsidName="Calathus cinctus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis italics="true" pageId="0" pageNumber="71">Calathus cinctus</emphasis>
<taxonomicName id="07A8501B6A4824AAA546AEBA695FFEB7" authorityName="Motschulsky" authorityYear="1850" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" lsidName="Calathus cinctus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis id="CEBF5F39B8C2CAED667515964787B494" italics="true" pageId="0" pageNumber="71">Calathus cinctus</emphasis>
</taxonomicName>
Motschulsky, 1850,
<taxonomicName authorityName="Menetries" authorityYear="1832" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis andreae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="andreae">
<emphasis italics="true" pageId="0" pageNumber="71">Cymindis andreae</emphasis>
<taxonomicName id="3E51DE434460013CF8102ED2849C063E" authorityName="Menetries" authorityYear="1832" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis andreae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="F3FE9B0706EA08D68C9806997D57125E" italics="true" pageId="0" pageNumber="71">Cymindis andreae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
<normalizedToken id="09328264157CA6C882140C2C6375BB45" originalValue="Ménétriés">Menetries</normalizedToken>
, 1832, and
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus impressus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="impressus">
<emphasis italics="true" pageId="0" pageNumber="71">Carabus impressus</emphasis>
<taxonomicName id="5FF0408293E03BB7627FDE82EDB2DC3F" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus impressus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="0C530A9D901DC52F56AB4FD4870E24D3" italics="true" pageId="0" pageNumber="71">Carabus impressus</emphasis>
</taxonomicName>
Klug, 1832. In Wadi Rum the single specimen was found in a dry wadi with
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Retama" kingdom="Plantae" lsidName="Retama" order="Fabales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Retama</emphasis>
<taxonomicName id="0CC878B092BC7622E842E1505E156152" class="Magnoliopsida" family="Fabaceae" genus="Retama" kingdom="Plantae" lsidName="Retama" order="Fabales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="F26293DD759B1409B4E5D56E9A29B2B1" italics="true" pageId="0" pageNumber="71">Retama</emphasis>
</taxonomicName>
shrubs and single
<taxonomicName class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
<taxonomicName id="0EE6353674A91ADB611609A3FCAE1B7A" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" lsidName="Ficus" order="Rosales" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="8A6B181D1F97B5EBCD4E705F971969C8" italics="true" pageId="0" pageNumber="71">Ficus</emphasis>
</taxonomicName>
trees in the shade of the walls of the head of the canyon-like valley. It occurs together with ground beetles that are mainly typical for desert habitats (
<taxonomicName authorityName="Schatzmayr" authorityYear="1936" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus aegyptiacus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis italics="true" pageId="0" pageNumber="71">Laemostenus aegyptiacus</emphasis>
<taxonomicName id="B1AAE5D30699CC0976A02B5403830F4A" authorityName="Schatzmayr" authorityYear="1936" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus aegyptiacus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis id="9D4AB572F82F81139928469D0D1CDCA7" italics="true" pageId="0" pageNumber="71">Laemostenus aegyptiacus</emphasis>
</taxonomicName>
Schatzmayr, 1936,
<taxonomicName authorityName="Peyerimhoff" authorityYear="1907" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" lsidName="Singilis filicornis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis italics="true" pageId="0" pageNumber="71">Singilis filicornis</emphasis>
<taxonomicName id="C079019AEF83F02DC06A366E19329B92" authorityName="Peyerimhoff" authorityYear="1907" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" lsidName="Singilis filicornis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="36F7228DE3D31474FBB9B9BA5B72B8FC" italics="true" pageId="0" pageNumber="71">Singilis filicornis</emphasis>
</taxonomicName>
Peyerimhoff, 1907,
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis hierichontica" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="hierichontica">
<emphasis italics="true" pageId="0" pageNumber="71">Cymindis hierichontica</emphasis>
<taxonomicName id="9B17A5152F6BACDF69673B80FC913CF6" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis hierichontica" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="hierichontica">
<emphasis id="D4A1B853AC0220DF2BDE76FD32EAA0CE" italics="true" pageId="0" pageNumber="71">Cymindis hierichontica</emphasis>
</taxonomicName>
Reiche &amp; Saulcy, 1855,
<taxonomicName baseAuthorityName="Klug" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" lsidName="Merizomena castanea" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="71">Merizomena castanea</emphasis>
<taxonomicName id="F63FFEF4C06C32F990E2E3AE8DE97EFC" baseAuthorityName="Klug" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" lsidName="Merizomena castanea" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="6CC4670815229D0B583B7AB14A246CF3" italics="true" pageId="0" pageNumber="71">Merizomena castanea</emphasis>
</taxonomicName>
Klug, 1832, and
<taxonomicName authorityName="Bedel" authorityYear="1907" class="Insecta" family="Carabidae" genus="Amara" kingdom="Animalia" lsidName="Amara maindroni" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis italics="true" pageId="0" pageNumber="71">Amara maindroni</emphasis>
<taxonomicName id="1F19F46C0B2F256DC451FDFD018ED038" authorityName="Bedel" authorityYear="1907" class="Insecta" family="Carabidae" genus="Amara" kingdom="Animalia" lsidName="Amara maindroni" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="90D972FEBB8D02FB4F3E21C856089680" italics="true" pageId="0" pageNumber="71">Amara maindroni</emphasis>
</taxonomicName>
Bedel, 1907). A photograph of the habitat of the latter is given by
<bibRefCitation DOI="https://doi.org/10.4081/fe.2017.226" author="Casale, A" journalOrPublisher="Fragmenta Entomologica" pageId="0" pageNumber="71" pagination="13 - 24" refId="B14" refString="Casale, A, Assmann, T, 2017. The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini). Fragmenta Entomologica 49 (1): 13 - 24, DOI: https://doi.org/10.4081/fe.2017.226" title="The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini)." url="https://doi.org/10.4081/fe.2017.226" volume="49" year="2017">Casale and Assmann (2017</bibRefCitation>
<bibRefCitation id="3D0B02C88ADE9681507B292C6BAF236D" DOI="https://doi.org/10.4081/fe.2017.226" author="Casale, A" journalOrPublisher="Fragmenta Entomologica" pageId="0" pageNumber="71" pagination="13 - 24" refId="B14" refString="Casale, A, Assmann, T, 2017. The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini). Fragmenta Entomologica 49 (1): 13 - 24, DOI: https://doi.org/10.4081/fe.2017.226" title="The Sphodrina of the southern Levant (Coleoptera: Carabidae, Sphodrini)." url="https://doi.org/10.4081/fe.2017.226" volume="49" year="2017">Casale and Assmann (2017</bibRefCitation>
: 22, fig. 35).
</paragraph>
<caption doi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 16.</emphasis>
<caption id="B7C2D1EF7B8A4877358839DC3BDD8F62" doi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71" start="Figure 16" startId="F16">
<paragraph id="483983A43D776E02E1EE56E7D509A132" pageId="0" pageNumber="71">
<emphasis id="A82F0EC46609CDFBB8F678D69495C335" bold="true" pageId="0" pageNumber="71">Figure 16.</emphasis>
Distribution of species from subgenus
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus</taxonomicName>
<taxonomicName id="0CF5AE6AA23714C19306BE0F9F4E8C35" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus</taxonomicName>
in the western Aegean Sea Region and Turkey:
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus chatzakiae</emphasis>
<taxonomicName id="55598F8B258B4CB300CD1F4260201A78" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae">
<emphasis id="0ABF9738A31581663F502B848848EE96" italics="true" pageId="0" pageNumber="71">Platyderus chatzakiae</emphasis>
</taxonomicName>
sp. nov. (red circle),
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<taxonomicName id="A2DB52AA7BEC20A2A6F14F82114F409E" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="DE821850421226482554E6315F7BA44B" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
(blue circles),
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="1DBEC94B7AD070B35B2058CD5FDE6A7A" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="6661663CC6C806C50E94CC7E5847CACD" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. (brown circles), and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
<taxonomicName id="52987D2EFE606E6315F0A0B2D597C6E8" class="Insecta" kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis id="BAF8B982B86568A17EFFFFEFB018E2E2" italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
</taxonomicName>
sp. nov. (yellow circle).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="A30A76F02F89038680F6B502C8535FFC" pageId="0" pageNumber="71" type="distribution">
<paragraph id="E011C47E6EBED90C8239CDA528ED8A75" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="0BC5D2F713C7E1659C15AAFB4F4018BE" pageId="0" pageNumber="71">
According to the data available, the species is the representative of the group &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="3100D88B314CC03E6067393CE804D99D" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="52087D64249F0557D6D89A3BBC8D2E21" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; in the southern part of Jordan, stretching from the vicinity of Little Petra to areas close to the Saudi Arabian border (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
<figureCitation id="A560F786574B40FFFAE66A411A8695B8" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
). As Wadi Rum is in the most southern part of Jordan, the species may also occur in Northwest Saudi Arabia.
</paragraph>
</subSubSection>

476
data/A4/50/6F/A4506FC87AE95F68823AFCC37C9ADD51.xml
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@ -1,422 +1,422 @@
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<mods:namePart id="F2FE8BCA09DEA9257779A06F822257F6">Assmann, Thorsten</mods:namePart>
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15.2.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3F</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5H</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6B</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7J</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11C</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12K</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15B</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
<subSubSection id="E7390FE7DE2DA01ADBD609DBC8464C68" pageId="0" pageNumber="71" type="description">
<paragraph id="8A270EC866AE153BC6A415435C672489" pageId="0" pageNumber="71">
<figureCitation id="F4F8AC46FBD8CE235755334919E83767" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3F</figureCitation>
<figureCitation id="B95547CEEA5E4D0EBAA89C4EC70B86C7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5H</figureCitation>
<figureCitation id="E33D41D016D6451BDC8B0E29BEFAEF93" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6B</figureCitation>
<figureCitation id="0D56FE246A50DA1C891F3437A5291FFF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7J</figureCitation>
<figureCitation id="EF10534D9E159249A120C4FA246F89CA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9D</figureCitation>
<figureCitation id="C1358796743AE7B7C7E87B4AEE69FE73" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11C</figureCitation>
<figureCitation id="CCEAA197E4FE0B60CCE13932B6328E23" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12K</figureCitation>
<figureCitation id="415F2CC718CF369C36348E5F70D0946F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15B</figureCitation>
<figureCitation id="F9AB372475F3C66073D48428490D6052" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="reference_group">
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" baseAuthorityName="Reitter" baseAuthorityYear="1909" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei" status="stat. nov.">Platyderus ferrantei</taxonomicName>
<subSubSection id="D18239152023C1C14B4C0181FF9B6F07" pageId="0" pageNumber="71" type="reference_group">
<paragraph id="84CFCCF207AB5B76FDD1516FE3B9C39B" pageId="0" pageNumber="71">
<taxonomicName id="DB6597F50CC57469838356CC2BB17F60" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" baseAuthorityName="Reitter" baseAuthorityYear="1909" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei" status="stat. nov.">Platyderus ferrantei</taxonomicName>
Reitter, 1909: 29 (type locality: Egypt,
<normalizedToken originalValue="“Cairo”">&quot;Cairo&quot;</normalizedToken>
<normalizedToken id="7BEA5F150C0CE4D66C7A151CCF65CDB4" originalValue="“Cairo”">&quot;Cairo&quot;</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="4B7FE9A7A82F64526D0A264FB056B53D" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="6DE639102A9E004C24754B21C460127B" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="8CF5C7D73A5D1B4A2163490A388D79EA" pageId="0" pageNumber="71">
Nothing is known about the exact locality and circumstances of collecting. At time of collecting, the city of Cairo was entirely situated on the east bank of the Nile River. Nevertheless, it is possible for the subspecies
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="321AECD4A8BF57FB500FECE623F8017C" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="E78AB99FDC21663D65C53D1E135084AF" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
to occur on both sides of the Nile River, as moisture should favor the
<normalizedToken originalValue="species’">species'</normalizedToken>
<normalizedToken id="FD2B768DCC80556CD75C22E06452787D" originalValue="species’">species'</normalizedToken>
occurrence and distribution in certain less human-influenced habitats.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" baseAuthorityName="Reitter" baseAuthorityYear="1909" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus ferrantei</emphasis>
<subSubSection id="818EAAA33F3CCF88751058B7231D2222" pageId="0" pageNumber="71" type="references">
<paragraph id="8C4C87643790FBCEF147BB53E319A124" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="A5960E66231A2117E6A8CCF35817B636" pageId="0" pageNumber="71">
<taxonomicName id="46CEA6F7BCFF582718C4858BA93E876E" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" baseAuthorityName="Reitter" baseAuthorityYear="1909" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei">
<emphasis id="1406A9B36ACDD94E041DF161BF18F545" italics="true" pageId="0" pageNumber="71">Platyderus ferrantei</emphasis>
</taxonomicName>
:
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
<bibRefCitation id="ACBF3AE1C3B2C30200F4977A289A3806" author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
: 768;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="840C272703A5A91BB1413244803AEC38" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="CA30B45652190777B20AE721FC4E9F56" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 522;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="76CF371511547B36C1E2F6A521B5AB96" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 395;
<bibRefCitation DOI="https://doi.org/10.15560/8.2.197" author="Abdel-Dayem, MS" journalOrPublisher="Check List" pageId="0" pageNumber="71" pagination="197 - 203" refId="B1" refString="Abdel-Dayem, MS, 2012. An annotated checklist of the endemic Carabidae (Coleoptera) of Egypt. Check List 8 (2): 197 - 203, DOI: https://doi.org/10.15560/8.2.197" title="An annotated checklist of the endemic Carabidae (Coleoptera) of Egypt." url="https://doi.org/10.15560/8.2.197" volume="8" year="2012">Abdel-Dayem 2012</bibRefCitation>
<bibRefCitation id="3F7D991CA56FE17DF9E37C0C02682BB1" DOI="https://doi.org/10.15560/8.2.197" author="Abdel-Dayem, MS" journalOrPublisher="Check List" pageId="0" pageNumber="71" pagination="197 - 203" refId="B1" refString="Abdel-Dayem, MS, 2012. An annotated checklist of the endemic Carabidae (Coleoptera) of Egypt. Check List 8 (2): 197 - 203, DOI: https://doi.org/10.15560/8.2.197" title="An annotated checklist of the endemic Carabidae (Coleoptera) of Egypt." url="https://doi.org/10.15560/8.2.197" volume="8" year="2012">Abdel-Dayem 2012</bibRefCitation>
: 200-201;
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="86F304A826D402BF2DD8CCACF4BE6313" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 758.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" baseAuthorityName="Reitter" baseAuthorityYear="1909" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus ferrantei</emphasis>
<subSubSection id="1CCFC503DAFFA245BD0C13698B85CD1D" pageId="0" pageNumber="71" type="type material">
<paragraph id="DA4F480F55792C1FEDC8A14287962FD8" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="7DCCF9009A61821F56E1BC3B26D178BD" pageId="0" pageNumber="71">
<taxonomicName id="E28C54AF2C6CD9877A0DB5EDD3C92C8B" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" baseAuthorityName="Reitter" baseAuthorityYear="1909" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei">
<emphasis id="E3E1C0C0BB08368B2BEFCE4507AF9C6D" italics="true" pageId="0" pageNumber="71">Platyderus ferrantei</emphasis>
</taxonomicName>
Reitter, 1909. Holotype, female in HNHM, with the following labels: 'Aegiptus / Cairo
<normalizedToken originalValue="Ferrante’">Ferrante'</normalizedToken>
<normalizedToken id="4A5C2AF9D46DB4BC07E5D5DEDF85A45B" originalValue="Ferrante’">Ferrante'</normalizedToken>
[h, w] //
<taxonomicName genus="Ferrantei" lsidName="Ferrantei" pageId="0" pageNumber="71" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Ferrantei</emphasis>
<taxonomicName id="103D13D2792E96FD280C00798521AA0A" genus="Ferrantei" lsidName="Ferrantei" pageId="0" pageNumber="71" rank="genus">
<emphasis id="9624593E515BD810A0DBEA249C96E5B5" italics="true" pageId="0" pageNumber="71">Ferrantei</emphasis>
</taxonomicName>
/ m. / Cairo. [h, w, written by Reitter himself] // coll. Reitter [p, w] // Holotypus 1909 /
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="C52D5E25A05316FF78B98BB62507617A" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="7D5678E9892F7238F7FC34145E71070B" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
/
<taxonomicName genus="Ferrantei" lsidName="Ferrantei" pageId="0" pageNumber="71" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Ferrantei</emphasis>
<taxonomicName id="6098D8318B37038CCCEFC2D2D97ED21B" genus="Ferrantei" lsidName="Ferrantei" pageId="0" pageNumber="71" rank="genus">
<emphasis id="6A35D28C7E3207AD6CB33EAC9E9B6A5A" italics="true" pageId="0" pageNumber="71">Ferrantei</emphasis>
</taxonomicName>
/ Reitter [p, h, w/r, subsequently added]'.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="F9871143C13730E92F04D79384032395" pageId="0" pageNumber="71">
The holotype designations in many type series, stored in HNHM, made by the late
<normalizedToken originalValue="Zoltán">Zoltan</normalizedToken>
<normalizedToken id="47AD694D6259AA74992AEE3779634BE1" originalValue="Zoltán">Zoltan</normalizedToken>
Kaszab (or one of his co-workers) are invalid because in most cases these are syntypes. In this case, the labelling as holotype was correct, as it is only one specimen on which the description was based.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
<subSubSection id="84DDD1A73EC3A907748EF64F46286CAD" pageId="0" pageNumber="71" type="type material">
<paragraph id="426E4D8C5AE1FD49B1CFEB76262B90E9" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="36831178AD11F3843FC5D7850C998422" pageId="0" pageNumber="71">
<taxonomicName id="C2C33349862F0F2F0406A18911003C84" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="0CE8F141EF7B730A5649E3A9926F5E50" italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
</taxonomicName>
Karsch, 1881. One female specimen and one male specimen from type series of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="C43D6F7FB7D133FD9A1B7C3CCB79325D" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="F5C65CF9F02424373458D9FD7A0A263F" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
are conspecific with the holotype of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. ferrantei</emphasis>
<taxonomicName id="CF2CFE8E18DDD5D6374927BAC63021CD" class="Insecta" kingdom="Animalia" lsidName="P. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ferrantei">
<emphasis id="50F09050169B18E7D418D1B716ABA2F3" italics="true" pageId="0" pageNumber="71">P. ferrantei</emphasis>
</taxonomicName>
. They are labelled as follows: 1♀, '2469 [p, w] // Hist.-Coll. (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="C4D5E8AE671F99E1A3BB401857EBE9FC" class="Insecta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 2469 /
<taxonomicName class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Sphodrus</taxonomicName>
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="BB46C5EA612B439A196165DAAE73DCB4" class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Sphodrus</taxonomicName>
<taxonomicName id="91FE04F49CCDF28052D4E5845DCEEF43" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
/ Aegypt., Ehrenberg / Zool. Mus. Berlin [p, w] // SYNTYPE /
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="E889C66CFCAF47F2C574AB9C326C4E74" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="ACA82F789E3902EABF97A3C469D7CCD9" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r] //
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="E612C634B0B3EA8DEFED595D0D57D51C" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
/ m. Karsch* / Egypt Ehrbg. [h, tl]' (MFNB); 1♂, '2470 [p, w] // Syria. Ehr: / lxxiii. 196. [h, y] // Hist.-Coll. (
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="FD5C75E25195F5D2F5F15B88D5882D7D" authorityName="Stephens" authorityYear="1827" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 2470 /
<taxonomicName class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Sphodrus</taxonomicName>
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="9E404AE79C6310F6EB9CE0010BDCB15A" class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Sphodrus</taxonomicName>
<taxonomicName id="B577131D9D388C6C7604236D2798D699" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
/ Syria, Ehrenberg / Zool. Mus. Berlin [p, w] // SYNTYPE /
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="AF72C18B406E67E67ADDE91E1527C862" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="68643AB084253DAFE0C9A9940571B5C7" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]' (MFNB). The male specimen has non-chitinised genitalia and a paler coloration (see
<bibRefCitation DOI="https://doi.org/10.1002/mmnd.18810250108" author="Karsch, F" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="0" pageNumber="71" pagination="41 - 50" refId="B28" refString="Karsch, F, 1881. Die Kaefer der Rohlfs'schen Afrikanischen Expedition 1878-79. Berliner Entomologische Zeitschrift 25: 41 - 50, DOI: https://doi.org/10.1002/mmnd.18810250108" title="Die Kaefer der Rohlfs'schen Afrikanischen Expedition 1878 - 79." url="https://doi.org/10.1002/mmnd.18810250108" volume="25" year="1881">Karsch 1881</bibRefCitation>
<bibRefCitation id="4FFBD086610501FF6C1CEB3FA8D7616B" DOI="https://doi.org/10.1002/mmnd.18810250108" author="Karsch, F" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="0" pageNumber="71" pagination="41 - 50" refId="B28" refString="Karsch, F, 1881. Die Kaefer der Rohlfs'schen Afrikanischen Expedition 1878-79. Berliner Entomologische Zeitschrift 25: 41 - 50, DOI: https://doi.org/10.1002/mmnd.18810250108" title="Die Kaefer der Rohlfs'schen Afrikanischen Expedition 1878 - 79." url="https://doi.org/10.1002/mmnd.18810250108" volume="25" year="1881">Karsch 1881</bibRefCitation>
: 44), indicating the immaturity of the specimen.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 15.</emphasis>
<caption id="CE8A337D9F640A9719ED1F27FB7EA4CA" doi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71" start="Figure 15" startId="F15">
<paragraph id="2B6DA2B1A78D86FAB694C562E44ABC35" pageId="0" pageNumber="71">
<emphasis id="A86B8FF7353BA516688E931EA2C7AC87" bold="true" pageId="0" pageNumber="71">Figure 15.</emphasis>
Spermathecal complex and gonocoxites, ventral view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authority="brunneus" authorityName="brunneus Karsch" authorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="Eremoderus" subSpecies="brunneus">Platyderus (Eremoderus) brunneus brunneus</taxonomicName>
<emphasis id="EBF2597FCA331E5CB3A2C327B1C317AD" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="A79EE08FCEB379CB32002DF78943A955" authority="brunneus" authorityName="brunneus Karsch" authorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="Eremoderus" subSpecies="brunneus">Platyderus (Eremoderus) brunneus brunneus</taxonomicName>
Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="DF10C3A233C79D76CF0FC92CFFB6F3BD" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="A32EF8B0C12D377E6334A8BE2C9CBBA2" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName authority="insignitus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="E." subSpecies="insignitus">P. (E.) insignitus insignitus</taxonomicName>
<emphasis id="3DC73BDC9D6AE0F88BD2BC30CE4A05B5" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="289A7899A45DD62A00001EA7E489CD7A" authority="insignitus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="E." subSpecies="insignitus">P. (E.) insignitus insignitus</taxonomicName>
Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
<emphasis id="0B17A0F73C256787C53284BDB4212BA7" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="8C1EC3AF9CC8F9E38241ACBEAF2456CF" class="Insecta" kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
<emphasis id="A5196D176B22EE39A20C384B06939DA9" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="9AB7BE90F2650D55A824C6E869F50897" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., female paratype, N Wadi Musa,
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
<normalizedToken id="FC145B4B3A7DF490C2E422124ADEF32C" originalValue="Ma’an">Ma'an</normalizedToken>
Governorate, Jordan;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="4614782EBA4B69EB165FDE1A70061076" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="EEC59B19595FBD7E0A2B2B37C083BDC3" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22.xii.1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13</figureCitation>
<figureCitation id="D45CE496830EAFF08943F45E22F75EC2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13</figureCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="2008-03-01" collectorName="Israel, Holot Haluza, Ittai Renan'" country="Israel" location="Southern District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Southern District">
<emphasis bold="true" pageId="0" pageNumber="71">
<collectingCountry name="Israel">Israel</collectingCountry>
<subSubSection id="38A59AE233FA21F56A41832AD594C7B3" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="DED18425216163C3AF975B7EAE5BCD55" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="884775E430D4D5B5A925DAB00906743A" pageId="0" pageNumber="71">
<materialsCitation id="3A01C8A0816334BF89BE3EABF1C2BF3A" collectingDate="2008-03-01" collectorName="Israel, Holot Haluza, Ittai Renan'" country="Israel" location="Southern District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Southern District">
<emphasis id="DCA9E22C89916F99208B83A1C0CA9F29" bold="true" pageId="0" pageNumber="71">
<collectingCountry id="787C64ABA701E620025AB26387C7C835" name="Israel">Israel</collectingCountry>
</emphasis>
:
<collectingRegion country="Israel" name="Southern">Southern District</collectingRegion>
<collectingRegion id="30895EE5CAA775AA40A9B1CF31025550" country="Israel" name="Southern">Southern District</collectingRegion>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="AAC00FC9B558A30D374B80A6A47A16B1" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="D346072ADD84B818F7CCA2852B0C704C" type="female">1♀</specimenCount>
, '
<collectorName>Israel</collectorName>
<collectorName id="45AF961A72DF3475DA655FDCC1DACC9F">Israel</collectorName>
/
<collectorName>Holot Haluza</collectorName>
<collectorName id="B8AE3E027E6A258D48F172EB6CE545CF">Holot Haluza</collectorName>
/
<collectingDate value="2008-03-01">1.iii.2008</collectingDate>
<collectingDate id="2AAF9D7732F70823A02ACA733AC5A8A3" value="2008-03-01">1.iii.2008</collectingDate>
/
<collectorName>
<collectorName id="817E263A49DC46036E925707F862C564">
Ittai
<normalizedToken originalValue="Renan’">Renan'</normalizedToken>
<normalizedToken id="DA4A64BD2F5A84D8BB719E546416E04A" originalValue="Renan’">Renan'</normalizedToken>
</collectorName>
(SMNH-TAU)
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="A18D3DF05602CA813C757327144AF748" type="female">1♀</specimenCount>
, 'Israel / Holot Haluza / 25.IV.
<date bridgedPair="/" value="2009-01">2009 / I.</date>
<normalizedToken originalValue="Renan’">Renan'</normalizedToken>
<date id="E6B9B8A5FDEAF540BB093F1190801086" bridgedPair="/" value="2009-01">2009 / I.</date>
<normalizedToken id="EDF335EFFDE76DC7A4177C2D67FABAB5" originalValue="Renan’">Renan'</normalizedToken>
(SMNH-TAU);
<specimenCount type="female">2♀♀</specimenCount>
<specimenCount id="5D7542F640AE297CFD6995A90552E166" type="female">2♀♀</specimenCount>
, 'Israel / Holot Haluza /
<date value="2010-03-02">2.iii.2010</date>
<date id="CA8336E7AA70A11FE6B12B8DD178E500" value="2010-03-02">2.iii.2010</date>
/ I.
<normalizedToken originalValue="Renan’">Renan'</normalizedToken>
<normalizedToken id="63B05BB66855E745DB6ABF534931B8A5" originalValue="Renan’">Renan'</normalizedToken>
(SMNH-TAU).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="C2048FDE13B2FE1B827A659CAA4DE0F8" pageId="0" pageNumber="71">
TME:
<specimenCount type="generic">8 specimens</specimenCount>
<specimenCount id="0C5D65FE7D8CEC201BB55B1CC6C0F156" type="generic">8 specimens</specimenCount>
. TGE:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="9C84D513B84966CAEF641C36F43FD10A" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="C16151CC9515EA81FF86C936F9F6CF4B" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="C142ABEFAA8F79B7F23B7B825C36EA36" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="FD21765253FBC639B74B7F12AACC8FB9" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="0F1BB1817D8F4814D62594579E4F2975" pageId="0" pageNumber="71">
See
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="605F1863F2C604B32CD8275CC6B20A8E" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
under the nominotypical subspecies.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="530627C5982A6304EE93E19536E8D342" pageId="0" pageNumber="71" type="comparisons">
<paragraph id="C75C0DCA693D23CAAB1EA044046AEB6A" pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph id="AD6CC61D10BB0CFC267B73EF73450529" pageId="0" pageNumber="71">
The subspecies considered here differs from easterly occurring representatives of the &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="BB9DE48DA24CEAF229E865DD487E1B09" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="81EB8D89119B3D2DAD12DC00B3566784" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group, i.e.
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="01B328389C97D6B80EEDF69F7CF02F76" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="057BC0E9B6916662C62C76B3C7113859" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="ECE97DB66C93E19B9CCCC0A243C7FB8F" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="8007E5B1AA8CBBD2FD08086071B588B9" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="31306B807F4759B5B18046DF9DD55FC0" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="515E005EDF6BD07C16AE8ACCF7B9BA06" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov., by the meso- and metatarsomeres dorsally somewhat flattened and grooved. Its median lobe of aedeagus is more arcuate ventrally and with ventral margin of apex straight (this of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="40F16C57E47E17E78EA44B5ADDB7CE2D" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="0C49BA74898DFA2025762609EED3565D" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
is appreciably less arcuate ventrally, whereas that of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="4CC31DBFF1E1677058FD93E5D5EBCA38" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="CB38D1F8850179BD9679A7EE2649E8F6" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
has ventral margin of apex convex). Other differences between
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
<taxonomicName id="A8A475B40BC4FFEF575B88F2243C40A3" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="503A52E337187F5D172BA2A2D71C3C0D" italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="16988F9B2866B27ABBA60EB3E5351E07" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="5633B95C9AEE2860A671A5627A2D98B6" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. are listed in section
<normalizedToken originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
<normalizedToken id="1A06669E79A50D2746FFB3B8919EE30A" originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
under the latter taxon.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="33EAB4B6AF628324E1AC375B6262FE5A" pageId="0" pageNumber="71">
Supplementary distinctions between
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
<taxonomicName id="7AE666542EFA96056FF5C4A480DFE1EB" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="B576583CDC0181B2F8F5BBBC5212730C" italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="D80770F369DC06C52DC04BDC4DB5343F" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="9EE1B11C990BE68BD0B703BD2E74CB84" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov., are: a/ head compared to pronotum wider, PW/HW: 1.29-1.39 (vs. head narrower, PW/HW: 1.41-1.43); b/ pronotum appreciably wider than long, PW/PL: 1.14-1.21, less constricted to apex, PW/PA: 1.34-1.42, with anterior margins compared to posterior margin long, PA/PB: 0.82-0.89 (vs. pronotum less wider, PW/PL: 1.07-1.11, more constricted to apex, PW/PA: 1.44-1.48, with anterior margin compared to posterior margin much narrower, PA/PB: 0.76-0.79).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<paragraph id="BDC0A3196CFA3E669B0219C8ADC8191F" pageId="0" pageNumber="71">
<taxonomicName id="7B648158BFD5AC124185CA1689890E44" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="B9C04FAA72299DBB910384241A4EF9F9" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
<taxonomicName id="9895B1E3B40B27D9D435D598713158D7" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="22D213668F54B6CD6BBB8ADDB1BD3FFD" italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
share meso- and metatarsomeres dorsally flattened and slightly grooved. The two taxa can be reliably diagnosed by a combination of three ratios - PW/PB, EL/EW and EL/PL (Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
<tableCitation id="C21BE046D9D876DEDD10C10084B4CADB" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="A8B63CF13170AD9A5793FF6670E0C6A5" pageId="0" pageNumber="71" type="habitat">
<paragraph id="1471670801AEFCA2D8B28F298A0D92A7" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="D1AB2FA7C65219FBABB6120FD6E4EEBE" pageId="0" pageNumber="71">
The Israeli site is located in the sand dunes of Western Negev. The species lives on the shifting dunes, together with
<taxonomicName authorityName="Assmann &amp; Renan&amp;Wrase" authorityYear="2015" class="Insecta" family="Carabidae" genus="Atlantomasoreus" kingdom="Animalia" lsidName="Atlantomasoreus groneri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="groneri">
<emphasis italics="true" pageId="0" pageNumber="71">Atlantomasoreus groneri</emphasis>
<taxonomicName id="18BD7EAA7CCAD4A87FCB6F11878A91CE" authorityName="Assmann &amp; Renan&amp;Wrase" authorityYear="2015" class="Insecta" family="Carabidae" genus="Atlantomasoreus" kingdom="Animalia" lsidName="Atlantomasoreus groneri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="groneri">
<emphasis id="FB56CB395F4413F961D25740EF29B8DF" italics="true" pageId="0" pageNumber="71">Atlantomasoreus groneri</emphasis>
</taxonomicName>
Assmann, Renan &amp; Wrase, 2015,
<taxonomicName authorityName="Fairmaire" authorityYear="1896" class="Insecta" family="Carabidae" genus="Discoptera" kingdom="Animalia" lsidName="Discoptera arabica" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabica">
<emphasis italics="true" pageId="0" pageNumber="71">Discoptera arabica</emphasis>
<taxonomicName id="FA3D66974421045FB6A2B25745F76AFE" authorityName="Fairmaire" authorityYear="1896" class="Insecta" family="Carabidae" genus="Discoptera" kingdom="Animalia" lsidName="Discoptera arabica" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="7B175A327919E57350E85848C6B27F07" italics="true" pageId="0" pageNumber="71">Discoptera arabica</emphasis>
</taxonomicName>
Fairmaire, 1896,
<taxonomicName class="Insecta" family="Carabidae" genus="Scarites" kingdom="Animalia" lsidName="Scarites striatulus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis italics="true" pageId="0" pageNumber="71">Scarites striatulus</emphasis>
<taxonomicName id="5AC7D7F0941610743141100251833E2C" class="Insecta" family="Carabidae" genus="Scarites" kingdom="Animalia" lsidName="Scarites striatulus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis id="02DDDD0C6DAF5108D250C1E0B9DE932B" italics="true" pageId="0" pageNumber="71">Scarites striatulus</emphasis>
</taxonomicName>
Dejean, 1825,
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Carabidae" genus="Thermophilum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thermophilum sexmaculatum" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="sexmaculatum">
<emphasis italics="true" pageId="0" pageNumber="71">Thermophilum sexmaculatum</emphasis>
<taxonomicName id="BEA37463029CD5BB2AF22C9AC7857AD8" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Carabidae" genus="Thermophilum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thermophilum sexmaculatum" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="sexmaculatum">
<emphasis id="5B080CC3CCB778E60D69E970BA184FF8" italics="true" pageId="0" pageNumber="71">Thermophilum sexmaculatum</emphasis>
</taxonomicName>
(Fabricius, 1787),
<taxonomicName baseAuthorityName="Forskal" baseAuthorityYear="1775" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus serrator" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="serrator">
<emphasis italics="true" pageId="0" pageNumber="71">Graphipterus serrator</emphasis>
<taxonomicName id="6F54A1FDA3011E6661403D7F27FEBA6B" baseAuthorityName="Forskal" baseAuthorityYear="1775" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus serrator" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="serrator">
<emphasis id="741C4D1F53FF21B5E3D8E6DB2311F914" italics="true" pageId="0" pageNumber="71">Graphipterus serrator</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Forsskål">Forsskal</normalizedToken>
<normalizedToken id="B16EDA9972BC676F8E2FE7CDEC6BA9D7" originalValue="Forsskål">Forsskal</normalizedToken>
, 1775) (or
<taxonomicName genus="G." lsidName="G. multiguttatus" pageId="0" pageNumber="71" rank="species" species="multiguttatus">
<emphasis italics="true" pageId="0" pageNumber="71">G. multiguttatus</emphasis>
<taxonomicName id="8F097222C6FACF47A50F2CF4DD3F3F96" lsidName="G. multiguttatus" pageId="0" pageNumber="71" rank="species" species="multiguttatus">
<emphasis id="ACF3AF6B0145C1922159D04BDF277843" italics="true" pageId="0" pageNumber="71">G. multiguttatus</emphasis>
</taxonomicName>
(Olivier, 1790)), and a species of the
<emphasis italics="true" pageId="0" pageNumber="71">
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Cymindis</taxonomicName>
<emphasis id="BDFB3624D0FB3816A09E470408D15380" italics="true" pageId="0" pageNumber="71">
<taxonomicName id="711DE97F0CABDE1500B62F33E093CEBB" class="Insecta" family="Carabidae" genus="Cymindis" kingdom="Animalia" lsidName="Cymindis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Cymindis</taxonomicName>
setifeensis/suturalis
</emphasis>
group(s). For a detailed description of the habitat see those of
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Carabidae" genus="Anthia" kingdom="Animalia" lsidName="Anthia sexmaculata" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis italics="true" pageId="0" pageNumber="71">Anthia sexmaculata</emphasis>
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and
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in
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,
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) and
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. Photographs of the habitat are given by
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, fig. 7b, p. 62) and
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, fig. 14, p. 59).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="93FBE8A4DEA2B8D511E6B7340617F1AA" pageId="0" pageNumber="71" type="distribution">
<paragraph id="3BE62E84FB81F5534AC3C2E6881E00E0" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="48242797DB0062C8C3ACB596750E96B5" pageId="0" pageNumber="71">
Northeast part of Egypt (not yet found in Sinai Peninsula, but possibly occurring there), southwestern part of Israel (Southern District), see Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
<figureCitation id="F06703449CE8A60D1A76F8370D07B86E" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
. The record from Holot Haluza is the first record for
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="BBCE0FE871F221B20C9247BBB1C461E7" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="A2028ED91D658E92A121319FF09A611B" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
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(s.l.) for Israel and for the Asian continent.
</paragraph>

734
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<treatment LSID="urn:lsid:zoobank.org:act:17C88C87-FE7B-46B8-8602-F57CC6318FD0" httpUri="http://treatment.plazi.org/id/A7EB05D68F17564D90265034DD4E3C08" lastPageNumber="71" pageId="0" pageNumber="71">
<subSubSection pageId="0" pageNumber="71" type="nomenclature">
<paragraph pageId="0" pageNumber="71">
<treatment id="A7EB05D68F17564D90265034DD4E3C08" ID-GBIF-Taxon="195868749" LSID="urn:lsid:zoobank.org:act:17C88C87-FE7B-46B8-8602-F57CC6318FD0" httpUri="http://treatment.plazi.org/id/A7EB05D68F17564D90265034DD4E3C08" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
<subSubSection id="4D4C2406B90E2DCF4CBBFD659D1A03C7" pageId="0" pageNumber="71" type="nomenclature">
<paragraph id="0966E82817C960538FBFC8854EB00CCC" pageId="0" pageNumber="71">
3.
<taxonomicName LSID="http://zoobank.org/17C88C87-FE7B-46B8-8602-F57CC6318FD0" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) felixi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<taxonomicName id="7077A2DBBAAFBA33AD7C71E1F55AF3CB" LSID="http://zoobank.org/17C88C87-FE7B-46B8-8602-F57CC6318FD0" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) felixi</taxonomicName>
<taxonomicNameLabel id="8680589E920BDDDC1ACD338FE7806A68" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1C</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5B</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7C</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8C</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10C</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12C</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13B</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
<subSubSection id="0A24F9F2AC665F6296487D02CD61F79F" pageId="0" pageNumber="71" type="description">
<paragraph id="18EB85A3F2B3FCCC66FFC90497A315C2" pageId="0" pageNumber="71">
<figureCitation id="8FB1902FBD47232C9975977614E20FCF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1C</figureCitation>
<figureCitation id="55F368E132E38F936EAC6957BF7F4C8B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5B</figureCitation>
<figureCitation id="B1E5C9F7500856EA002433B41EF97993" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7C</figureCitation>
<figureCitation id="5732B72B23814FDF619D36EABBBE6761" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8C</figureCitation>
<figureCitation id="D5B3B7067E3EC08EF73685CCBCA1DB3A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10C</figureCitation>
<figureCitation id="687153535A2CA691636A5060189A4938" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12C</figureCitation>
<figureCitation id="87E1DD599B28CC88CFA83F8186606EE2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13B</figureCitation>
<figureCitation id="6E55122D327F4BDD740A92FBF122573E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Iran, Chahar Mahal va Bakhtiari Province, 10 km W Naghan Town, 31.9410, 50.6014, 1492-1505 m.</paragraph>
<subSubSection id="E2DA97E374C86FCC7C1993EA34B1CB79" pageId="0" pageNumber="71" type="type locality">
<paragraph id="2BFCF088E135CCE187A6957706D8B589" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="01366C19B086C68410212E463EBE8465" pageId="0" pageNumber="71">Iran, Chahar Mahal va Bakhtiari Province, 10 km W Naghan Town, 31.9410, 50.6014, 1492-1505 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">The distance from Naghan Town to the type locality is about 10 km in straight line and the direction is to the west of the town. The GPS coordinates and height indicated are more or less correct as they well correspond to the river that flows towards the village of Do Polan. In fact, the type locality is closer to the last village than to Naghan Town.</paragraph>
<subSubSection id="AEEED6E14148AA42EBFC8D1A1E8733C8" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="66581E05DE0DFEE002FDCB97DE923383" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="DFA2E81269C3A3760794EA177D0D0CDD" pageId="0" pageNumber="71">The distance from Naghan Town to the type locality is about 10 km in straight line and the direction is to the west of the town. The GPS coordinates and height indicated are more or less correct as they well correspond to the river that flows towards the village of Do Polan. In fact, the type locality is closer to the last village than to Naghan Town.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="0A252BDD3559E112B28EB003BFB84E91" pageId="0" pageNumber="71" type="type material">
<paragraph id="83A2240CA14F9FAD8FC8042B6915FA53" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="F4BF9A1734F88569E3AB70F5DD5668FE" pageId="0" pageNumber="71">
<emphasis id="3E336792846AE56192A07082D6CD5CC4" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂, 'IRAN, /
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="D66FE294DDBA86B39980712279A3703A" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="25DD2A6E9E9CB0698B0919CAA37420AD" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="4395A5258AA35C59ED8AC71CBDC39D02" originalValue="vā">va</normalizedToken>
<normalizedToken id="EB20F769BB13CFD3F4AC24FD45972F34" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
/ 10km W Naghan Town /
<geoCoordinate degrees="31" direction="north" minutes="56" orientation="latitude" precision="15" seconds="27" value="31.940832">31°56'27&quot;N</geoCoordinate>
<geoCoordinate id="4B640405B55B12EE1E8D3621E1BA951A" degrees="31" direction="north" minutes="56" orientation="latitude" precision="15" seconds="27" value="31.940832">31°56'27&quot;N</geoCoordinate>
,
<geoCoordinate degrees="050" direction="east" minutes="36" orientation="longitude" precision="1" seconds="05.5" value="50.601524">050°36'05.5&quot;E</geoCoordinate>
<geoCoordinate id="4C2AD361A6B667C40DB308EFA1BA9FF2" degrees="050" direction="east" minutes="36" orientation="longitude" precision="1" seconds="05.5" value="50.601524">050°36'05.5&quot;E</geoCoordinate>
/ 1492-1505m, 02.04.2007 / leg. Jan
<normalizedToken originalValue="Muilwijk’">Muilwijk'</normalizedToken>
<normalizedToken id="EAA24628EE3F6798DEACDE95150CB1D7" originalValue="Muilwijk’">Muilwijk'</normalizedToken>
(HMIM). Paratypes: 4♀♀, labeled as holotype (cMUI); 1♂, 'IRAN, /
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="F57FA50C791240683EE30EAE59A08359" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="A8D36B5C4C5AD39D51DFF2156B610803" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="1F96F2EE572162A299AB63BDE58138F0" originalValue="vā">va</normalizedToken>
<normalizedToken id="69D582017C0A4ABDA8F7718A27085087" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
/ 10km W Naghan Town /
<geoCoordinate degrees="31" direction="north" minutes="56" orientation="latitude" precision="15" seconds="27" value="31.940832">31°56'27&quot;N</geoCoordinate>
<geoCoordinate id="64C0CF0C83040EE54C69E62689DC0AB7" degrees="31" direction="north" minutes="56" orientation="latitude" precision="15" seconds="27" value="31.940832">31°56'27&quot;N</geoCoordinate>
,
<geoCoordinate degrees="050" direction="east" minutes="36" orientation="longitude" precision="1" seconds="05.5" value="50.601524">050°36'05.5&quot;E</geoCoordinate>
<geoCoordinate id="E41D8B8CB55597CD3DA5BB5744D86CBC" degrees="050" direction="east" minutes="36" orientation="longitude" precision="1" seconds="05.5" value="50.601524">050°36'05.5&quot;E</geoCoordinate>
/ 1492-1505m, 02.04.2007 / leg. R.F.F.L.
<normalizedToken originalValue="Felix’">Felix'</normalizedToken>
<normalizedToken id="5B9526FCFE96E34BE20CCA738ACBD489" originalValue="Felix’">Felix'</normalizedToken>
(cFEL).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 6 specimens. TGE: 2♂♂, 1♀.</paragraph>
<paragraph id="7AFFD9EB9CD77F5629C57414FF5A12A5" pageId="0" pageNumber="71">TME: 6 specimens. TGE: 2♂♂, 1♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">Latinized patronym name after Ron Felix (Berkel-Enschot, Netherlands), an enthusiastic coleopterologist working on the taxonomy of carabid beetles, with great collecting skills, who collected part of the type series of the new species.</paragraph>
<subSubSection id="C03CC2526F1344414909965F770D58D5" pageId="0" pageNumber="71" type="etymology">
<paragraph id="4EC09A2E3A5211E887E96397FEA1CE4C" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="F13954789779B82A359EDA86F0900DA7" pageId="0" pageNumber="71">Latinized patronym name after Ron Felix (Berkel-Enschot, Netherlands), an enthusiastic coleopterologist working on the taxonomy of carabid beetles, with great collecting skills, who collected part of the type series of the new species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="238C1F72D316902D22E8B295558B4257" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="5D3754ED8ADE7B1ADAA4879F852EC0A6" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="D7E7401E9AFD04C53C1FAC18C5899156" pageId="0" pageNumber="71">
It is distinct from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
<taxonomicName id="6656C72541E4D261E130B68BCCB7E5CF" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="5F8BF6B68C13EC375DE1C8681CEF110C" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
sp. nov. in the pronotum significantly wider than head (PW/HW ≥1.45, vs. PW/HW ≤1.45) and elytra noticeably less wide in relation to length (EL/EW ≤1.55, vs. EL/EW ≥1.55). In addition, median lobe of aedeagus (lateral view) of the new species is larger, with shaft broader and apex barely bent up (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
<taxonomicName id="3E82C8D631F2EE96C263C28DCA9C95B6" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="C13A51A6EB6FF6E7411ABC819655CE13" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
has significantly narrower shaft and apex more distinctly bent up; Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C, D</figureCitation>
<figureCitation id="39D1693724E0B6FA181A6BCBAA94CE97" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="049D4065BA313719C13FCD9342503B15" pageId="0" pageNumber="71" type="description">
<paragraph id="4F7036E5942792FF348CB71085781A2A" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="33110E4A6B4EC2F94B2C4E76EDA55C4E" pageId="0" pageNumber="71">
<emphasis id="8A169772E6BDC36372349506C20AEA71" bold="true" pageId="0" pageNumber="71">
<emphasis id="E454A9E94D1E4C624CD210498C2339EF" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of moderate size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="21DC5C7A5971D1738067E30086374727" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="2433847BBAA30BDC5A237A3D3BB2FA60" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 6.00-7.00 mm; BW: 2.20-2.50 mm), with relatively short, moderately convex body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1C</figureCitation>
<figureCitation id="B72E1EECB30A2A98D4D7A3617A9094E9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="17F2A349195B42320C0FC8029B09A447" bold="true" pageId="0" pageNumber="71">
<emphasis id="353944DB9C70D5BB790C33E8B725ABA6" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
<tableCitation id="6CBA952797446A91453606ABF7CB4AC2" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="C4481492F164200F8B0EAEE62C2189EA" bold="true" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
. Integument uniformly reddish-brown, appendages somewhat lighter than body. Surface moderately shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="9182950638159F8007EFAAF13DCAC8E0" bold="true" pageId="0" pageNumber="71">
<emphasis id="FD0F84A699F39A8D123ED11F1384E64E" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Disc of head with less distinct microsculpture. Pronotum with regular isodiametric sculpticells posterolaterally and slightly stretched isodiametric sculpticells anterolaterally, microsculpture faint to absent in middle. Ventral surface (excl. gula) microsculptured, sculpticells isodiametric (mentum, submentum, pro-, mes-, and metepisternum, prosternum and abdominal ventrites laterally) or slightly stretched isodiametric (genae, medial surface of prosternum, metasternum, abdominal ventrites medially and legs).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="E50A1AD79F042CCED996AA8C1A5C27BA" bold="true" pageId="0" pageNumber="71">
<emphasis id="119370AAF44A57D2229C4086D60F1791" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
As wide as a half of pronotum (mean PW/HW= 1.49). Eyes moderately convex. Labrum subrectangular, as long as clypeus, with anterior margin straight. Frontoclypeal suture distinct in middle but very fine, lacking at sides. Frontal furrows shallow, punctiform. Paraorbital sulci moderately deep, ending slightly before or at level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="2D9F5D5E7C98CF421F51101578D3FEFC" bold="true" pageId="0" pageNumber="71">
<emphasis id="84F251BF94267CD2F9D8AABE51A53B1B" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum distinctly wider than long (mean PW/PL= 1.31), with widest point at second quarter. Anterior and posterior transverse impressions indistinct. Anterior bead present medially, very fine. Sides convex anteriorly, nearly straight posteriorly. Metepisternum slightly longer than wide, MA/MI= 0.90-0.95.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="8EE71CB525734B0F0E482F02E5AE5CA9" bold="true" pageId="0" pageNumber="71">
<emphasis id="D8B62E2B01B45C9297CC0EF1C188CD8F" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Cylindrical, one and a half time as long as wide (mean EL/EW= 1.52), wider and quite longer than pronotum (mean EW/PW= 1.31; mean EL/PL= 2.62). Parascutellar striole and striae moderately impressed, wide; striole short, not joining stria 1; striae 1-6 reaching basal bead, 7 reduced before. Following aberration exists in two specimens, striae 3 and 4 posteriorly, each of them fusing separately with stria 2 and joined stria reaches lateral gutter, otherwise striae posteriorly as in subgeneric description. Interval 3 with three or two minute discal setiferous punctures situated as follows: anterior puncture (in most specimens missing on one or both elytron) adjoining stria 2, in middle of interval 3 or adjoining stria 3, remaining two punctures adjoining stria 2. Umbilicate setiferous series mostly consists of 16 punctures on elytron, rarely with 17 punctures.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="BF3BA1C62E68435406B9535102A6CA5A" bold="true" pageId="0" pageNumber="71">
<emphasis id="AD6F571DC0CAC2308E6D0A3EB43B97B2" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur with four setiferous punctures on anterior side ventrally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5B</figureCitation>
<figureCitation id="4214A1606F3820C2BB11D8485E53FE2F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5B</figureCitation>
). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="D448A4A537D7203C08A4B36F91024135" bold="true" pageId="0" pageNumber="71">
<emphasis id="4C024B4D5AF52DE385D1A778C8CFF7B1" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX small, oval, with proximal margin slightly asymmetrical and rounded (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7C</figureCitation>
<figureCitation id="28A8DA2C6F04863586852386C4D967E2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7C</figureCitation>
). Median lobe of aedeagus curved in lateral view, with short basal bulb, long and broad shaft moderately constricted proximally, and apex hardly reversely turned up; median lobe in ventral view straight, ca. 3.7 times longer than wide; apical lamella more or less asymmetrical. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C</figureCitation>
<figureCitation id="DD418E7183219A0D4B4CB8FC4A42498B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8C</figureCitation>
) with ventral sclerite distinctly broadened and rounded distally; same in ventral view (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10C</figureCitation>
<figureCitation id="5EF35C731C8B2D284BD563FA5BE31C2E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10C</figureCitation>
), with dorsal sclerite forming a concave left protuberance and a convex right curve, and ventral sclerite narrow and completely straight. Right paramere rather short and crooked (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12C</figureCitation>
<figureCitation id="DFD9DAE45A7C53F0843A6E0820699133" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="3550A210B340AC72BDE3BBC60C44E429" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13B</figureCitation>
<figureCitation id="B391077C685327AD2DB79439F779A5DF" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13B</figureCitation>
). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 3.</emphasis>
<caption id="6723EF93653C06B8CB9A6E83A71C1A0F" doi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71" start="Figure 3" startId="F3">
<paragraph id="E8175D0F7ECDF55A9E6C73F8578704AF" pageId="0" pageNumber="71">
<emphasis id="582B2461D0AD8527F5BF67332EED2C87" bold="true" pageId="0" pageNumber="71">Figure 3.</emphasis>
Dorsal habitus.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">Platyderus (Eremoderus) afghanistanicus</taxonomicName>
<emphasis id="C81F59A99AE708E9C0E3A3130A21A84F" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="B3C41E95EF1ADDDA2F8EFF8E439E02B3" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">Platyderus (Eremoderus) afghanistanicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus" subGenus="E.">P. (E.) arabicus</taxonomicName>
<emphasis id="98CAA23630BD1F0A504D2F2E4CF196AC" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="C846F58C512DAA8706A376AC73D4B36E" class="Insecta" kingdom="Animalia" lsidName="P. (E.) arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus" subGenus="E.">P. (E.) arabicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" subGenus="E.">P. (E.) brunki</taxonomicName>
<emphasis id="9A8A955C2F86955577A7CE7B923684AC" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="D4E8E8D3407538DC8DB20BB25BBD08A1" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" subGenus="E.">P. (E.) brunki</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
<emphasis id="715B553D89D9AB007F4231526E56752A" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="F2A2AAE462046C748EC0307E452152E1" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
Karsch, 1881, lectotype and its labels;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
<emphasis id="1B3CF0605F90AC5D6A73941350F40EE9" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="A8A6E35EEC35FB1D9A3E3B39D6434883" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="0779C95E1CE9101502D6E94F1D58857A" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="CEEC47048A3639C3276D9CBCEA54A933" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, lectotype and its labels. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="37FCA4E229D05C84B7862AEE27CF9F19" pageId="0" pageNumber="71" type="comparisons">
<paragraph id="1A9BFF4388F4F1BC7E7057FB2C4AF5C5" pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph id="250796FDE8D15A5E1C25DA7EE9F285CD" pageId="0" pageNumber="71">
The new species lives in sympatry with
<taxonomicName baseAuthorityName="Menetries" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">Platyderus (s. str.) umbratus</taxonomicName>
<taxonomicName id="317EB1F4B5DF70BA3652BB9CDFB3AB80" baseAuthorityName="Menetries" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">Platyderus (s. str.) umbratus</taxonomicName>
(
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
<normalizedToken id="A9CF655EF8F36B75AD4977A9281C7600" originalValue="Ménétriés">Menetries</normalizedToken>
, 1832). However,
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="03692BB7DF8583C1CA5088D001F9B2D7" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="A6C7031595EDB0726C2D84AA514C5172" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
has pronotum more coarsely punctate in basolateral parts with punctures almost reaching the lateral margin, mesotibia with four setiferous punctures on its anterior side ventrally, and short urite with apex more symmetrical (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7C</figureCitation>
<figureCitation id="DE4212C3FC83BB5AA568690CD4124096" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7C</figureCitation>
), whereas
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis italics="true" pageId="0" pageNumber="71">P. umbratus</emphasis>
<taxonomicName id="E675F9325909BD1BB66B3F41F3FB0DD8" class="Insecta" kingdom="Animalia" lsidName="P. umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="1DA2C5A456BDB437FAF72985A8E65910" italics="true" pageId="0" pageNumber="71">P. umbratus</emphasis>
</taxonomicName>
possesses pronotum shallowly punctate in base with punctures mostly limited to the basal impressions, mesotibia with two setiferous punctures on its anterior side ventrally, and long urite with apex curved to left (as in Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7O</figureCitation>
<figureCitation id="0DD2CF5D6788217BDC7E9C2CC61DD7E1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7O</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="ADBAD32C4643AAB3D1823CD894BA80BA" httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Table 1.</emphasis>
<caption id="ADBAD32C4643AAB3D1823CD894BA80BA" ID-Table-UUID="ADBAD32C4643AAB3D1823CD894BA80BA" httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" start="Table 1" startId="T1">
<paragraph id="C31CB851BBE153D00884005EA7580FB8" pageId="0" pageNumber="71">
<emphasis id="7497D3329F7ECBF3D9297C227E04C8B4" bold="true" pageId="0" pageNumber="71">Table 1.</emphasis>
Morphometric data for species of the '
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
<taxonomicName id="C11F9B160672ADE2DE0BEA8CB3A8AD7C" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="F0691B6C2ABF9C2DEC67B09959C4639A" italics="true" pageId="0" pageNumber="71">Platyderus weiratheri</emphasis>
</taxonomicName>
' and '
<emphasis italics="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">Platyderus Eremoderus iranicus</taxonomicName>
<emphasis id="F3B2C75E08A53FE6EDCD2A6B7A328828" italics="true" pageId="0" pageNumber="71">
<taxonomicName id="B0FDE4861ED569FEEC23B0E29712C663" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">Platyderus Eremoderus iranicus</taxonomicName>
-
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">Platyderus Eremoderus vanensis</taxonomicName>
<taxonomicName id="E7DF5872771741352CE035ED163ADCDA" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">Platyderus Eremoderus vanensis</taxonomicName>
</emphasis>
' groups of
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="D6EC1E5F2793651FEDC3C3ED0B9E2084" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="1D13245FA83E063DDA66D1C63DE787DB" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="71">
<table pageId="0" pageNumber="71">
<tr pageId="0" pageNumber="71">
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">Species (number of samples)</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BL/mm</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BW/mm</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/HW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PL</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">
<paragraph id="2B1DABB41DF010F6C6FC6960BCE8A3E4" pageId="0" pageNumber="71">
<table id="8C8329815A536DB2FF00091F81B23D50" pageId="0" pageNumber="71">
<tr id="E8AB06ABD35D31FC02F905A2E705ED79" pageId="0" pageNumber="71">
<th id="D6FBB11D34368D7A00534E533DEC0EBC" colspan="1" pageId="0" pageNumber="71" rowspan="1">Species (number of samples)</th>
<th id="DE3227A8F96D1F5B1A38355977AAF57B" colspan="1" pageId="0" pageNumber="71" rowspan="1">BL/mm</th>
<th id="DD54984E65AC10F811F40540113971A4" colspan="1" pageId="0" pageNumber="71" rowspan="1">BW/mm</th>
<th id="E9687ED1831B6634A9A9036A15741DC3" colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/HW</th>
<th id="8EA7738C1265A4D9D7914739709DA1BD" colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PL</th>
<th id="52B54934B7BB90665AF49D94B105274D" colspan="1" pageId="0" pageNumber="71" rowspan="1">
PW/
<normalizedToken originalValue="PА">PA</normalizedToken>
<normalizedToken id="25ED7DF73370CA84F0DCD55FB654E12D" originalValue="PА">PA</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PB</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PA/PB</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/EW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EW/PW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/PL</th>
<th id="0D64BA8F78E6116BE5C896F02E6F0994" colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PB</th>
<th id="22BF5543369F46199778CE275D2FD367" colspan="1" pageId="0" pageNumber="71" rowspan="1">PA/PB</th>
<th id="8646032D0AD727A9F6542EA0051BB23C" colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/EW</th>
<th id="B4AC90045C9126B1EE9080C02FC29DD4" colspan="1" pageId="0" pageNumber="71" rowspan="1">EW/PW</th>
<th id="7E7DCD27097B9EA29EE6CB85E4909706" colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/PL</th>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<tr id="937281CB3C8006E9E4372AC834CE8A53" pageId="0" pageNumber="71">
<td id="3B6A2A0967D75685A6C9919EAC9B9519" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="D0EF4A01C78DB24D5865A0B47935B7F8" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="8F3EA36D6069250AE4F690B4F95BA692" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<normalizedToken id="382DDA3F4AA68A143302E7F09F841377" originalValue="Mařan">Maran</normalizedToken>
(6♂♂, 5♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.00-8.10</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.35-2.60</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.44</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.27</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09-1.19</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.81-0.89</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.71</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.55-2.83</td>
<td id="29907F28D5F9E06B29B95E39466818E8" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.00-8.10</td>
<td id="69B145530ABBF005659662D6EC861F07" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.35-2.60</td>
<td id="D829A7C12D906A2BF3C6100FBA9B7EC1" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.44</td>
<td id="FE1A082658241E018C72E33C4654FADF" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.27</td>
<td id="C7438345A033BE4612313622CF4FB672" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.41</td>
<td id="881BC9D0FF7069D0C95A5A38712DDE0C" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09-1.19</td>
<td id="5ACB0B8BB568074D100BF3DD49FDCBD5" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.81-0.89</td>
<td id="CF03DA1640D9040B187F200AC1193063" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.71</td>
<td id="ADEA651D75547BBF008FEF5AB5E9646D" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.40</td>
<td id="C09F35D8412BCD9F627E896C2139A9BF" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.55-2.83</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<tr id="1CFA9C0C6F03293F245ADC059EFFCC17" pageId="0" pageNumber="71">
<td id="79A053080D81ABFADCA5116F69C844BE" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="5C8219A961CA7BD54D14FC9E700B965C" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="6F8881472EED3DF29C0029F98C2FBE14" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<normalizedToken id="6F92CEDC9F94AB474777576914697A37" originalValue="Mařan">Maran</normalizedToken>
(mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.52</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.67</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.64</td>
<td id="FA040E83E6EFEB331DBF7F53F7E48008" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.52</td>
<td id="1E9A1696DDD306C4F8CE29E3D940CD40" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50</td>
<td id="5861B4291C7F10C84E957799D4C0A663" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td id="7E5B0242EBB81A047303F68236C04D70" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.20</td>
<td id="EBACB1B361AF4D523332230B733E4CAE" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37</td>
<td id="ED023F33FDBF9B337458E28A4C9CBB8F" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
<td id="3A8C50C57CD122CFC31584D1F7556A90" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
<td id="8E61A40037CC0C0CA8A99B67DAC9E3F8" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.67</td>
<td id="ACA5F2CB44F67CDBF5067528AA8F4993" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td id="DFDB788A2443D1DE6DB25573D6F139D5" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.64</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. chatzakii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakii">
<emphasis italics="true" pageId="0" pageNumber="71">P. chatzakii</emphasis>
<tr id="01244DF9F5DB49087AFB8E22BE8910F5" pageId="0" pageNumber="71">
<td id="A160A611196EDAD97DF87BF29E120CFB" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="34FE5C50099A69F96788A25C080C2CA6" class="Insecta" kingdom="Animalia" lsidName="P. chatzakii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakii">
<emphasis id="CD6C12BF9C0B155973450524F6F015CC" italics="true" pageId="0" pageNumber="71">P. chatzakii</emphasis>
</taxonomicName>
(1♂)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.05</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.85</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.49</td>
<td id="2A4FC27C28B7B299567485FB75EE995A" colspan="1" pageId="0" pageNumber="71" rowspan="1">8.05</td>
<td id="A8BF1C1063D16CAFA38E7E38959B1E44" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.85</td>
<td id="3D996E03405EBE1FC532CF9EE4278A9E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
<td id="13B78DA2495CEA9989F8E4AD7EDD8CEB" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td id="53E43AD3015636EA15AA33463E2D2D8A" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td id="B39FE5627C0D39A941DA6879BB1E513D" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td id="0603EEBA2F4F136B9E78A27B9A48CED6" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
<td id="F2C3559E8087663D3BECCB5A309569DE" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.60</td>
<td id="BC853571FBDC5A5F41AA2742311B9E59" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
<td id="2D0B471C30DAFC8C153FF16CD6C9CC0E" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.49</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<tr id="52E49DB1E7FCC84A7F4172F6E7CAF9F3" pageId="0" pageNumber="71">
<td id="B9A02D4AC3933F0BCCC1574D03CD1C9C" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="D3131783E5710B7DACA29183C374DDC8" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="2ACB46CC65E20313650121ECE99284D6" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. (2♂♂, 4♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.00-7.00</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.20-2.50</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.45-1.52</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.25-1.34</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.33-1.43</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.13</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.79-0.83</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.48-1.55</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.35</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60-2.66</td>
<td id="F105191989AFE66BF91A410D47857A82" colspan="1" pageId="0" pageNumber="71" rowspan="1">6.00-7.00</td>
<td id="37275B80DE04AEA3115B91AA6F809BAE" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.20-2.50</td>
<td id="54320990E09A142794D1F838BB48D98A" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.45-1.52</td>
<td id="81CEC4CEAEBC77DBB09AA8624059C56F" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.25-1.34</td>
<td id="69B5083C8041F353F05D294C18ABA5B2" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.33-1.43</td>
<td id="8FCEAECD8907D0A7BA89849C30C8D7D8" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.13</td>
<td id="F4038B7C504A53D35C2C4AD5E41BB209" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.79-0.83</td>
<td id="23DC84683755D19021F44389EF99999F" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.48-1.55</td>
<td id="390A9EB8D05A0589554620435E2A1EF5" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.35</td>
<td id="EF61AD2C357B18373572D12F558310D8" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60-2.66</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<tr id="1203DF93C74ACAF12E6A2ADE1555A43B" pageId="0" pageNumber="71">
<td id="FCAB459C9958CEDC9F9264A80E03FF12" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="5FC6DC94B636EA8B02D6CDA97232DA57" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="D35673A75A8661D67E81978FAF3CAF70" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.58</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.49</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.11</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.80</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.52</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.62</td>
<td id="C88DE9A512277B565803A8E438DF6847" colspan="1" pageId="0" pageNumber="71" rowspan="1">6.58</td>
<td id="632B98697231D6E3294EEC15EC8D3EBB" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40</td>
<td id="495FED51C391FABE94B8640626CE8162" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.49</td>
<td id="3394791142FE48470EC31B4C2999A3BD" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
<td id="54BD4EC8038081B64124031CDA7A92F7" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
<td id="59BB7DA961CEEB08FCA4778E436599C8" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.11</td>
<td id="1633715E6C1E669CEA3DA05FAA47EFAE" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.80</td>
<td id="2AB72E9B2BF6AD5ECD0248A13BA50D64" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.52</td>
<td id="B585DD8C26BD387BFFB7C849DF8B7ABF" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
<td id="4C5ED81F792270D59A30629893956C41" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.62</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
<tr id="131E7E45A89F3EC853C50CB98FF0311C" pageId="0" pageNumber="71">
<td id="C01E66B4A59B22FCC98FE82E90722472" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="9C6F9098FF4494B09A2CA66ED9ECA88B" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="C4961ECC7FFF973D482ADCCDD64D8EFE" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
sp. nov. (19♂♂, 16♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">4.30-6.65</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.50-2.30</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.21-1.31</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30-1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.24</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82-0.93</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.55-1.69</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.24-1.34</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50-2.77</td>
<td id="57252CAE0C64B2EF5A99276303675E95" colspan="1" pageId="0" pageNumber="71" rowspan="1">4.30-6.65</td>
<td id="18781C11529075147614B3547210F84E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.50-2.30</td>
<td id="93257A0E5697A7A49E416028959B3888" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.45</td>
<td id="2351FA7A3EB688C07CE33E1A1DB3FD3C" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.21-1.31</td>
<td id="E26C5697FC13702034EC4238F291EC4E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30-1.42</td>
<td id="F5EBEBEC99F71B0C23FD71ED52717B18" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.24</td>
<td id="411156F8401DDE3ED2CA321A8E790D6D" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82-0.93</td>
<td id="D9B08335099F016039EA1850D73EA9FE" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.55-1.69</td>
<td id="E51EA61EB9437E39D9F01CFAA441DC36" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.24-1.34</td>
<td id="2FDBDF1FE344DBBC66EF5586852B7DFF" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50-2.77</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
<tr id="E7A5C2AF5BC9E384F0B86DEC96934E08" pageId="0" pageNumber="71">
<td id="5CECEB0398AC7E808274F110678A6973" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="CABB34692E6AA2DC5441FF705DF8AC42" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="95E6C4FE3C4A1BC52D720FEAD854FA04" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">5.67</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.95</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.26</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.87</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.59</td>
<td id="56DDC4062CA023AEE556DFEFF5B9987D" colspan="1" pageId="0" pageNumber="71" rowspan="1">5.67</td>
<td id="7D107D83E5CEA6408DEFF34BED1038EC" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.95</td>
<td id="585FD2D1D5F4A67BF4DA5C45C0C10580" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td id="C2A2DFDC13F1148DEBFF807DDC7357F7" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.26</td>
<td id="6E08430D4EF1E3800FB633A80F1A49D0" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td id="3D331352438B0256B1FADC5C2F1F8176" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td id="F3EB2725E28A6518A407438EA693C315" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.87</td>
<td id="B479A9FC7EF685B85815314ACB9B26E0" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.60</td>
<td id="82CE79E2C0203AAFADF6F97C848470FB" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29</td>
<td id="34048B1723C5BFA97EBBE3DE7A24A968" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.59</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<tr id="C5DF9BFF938852BAA4BAEA5E066FBC95" pageId="0" pageNumber="71">
<td id="175BD7BDF04FD6E0CE8DA18220719F73" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="94AAACA5C3ECC1652C449855A762AEFC" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="6075A2427D5582D3E4E43EAFC2162C30" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. (11♂♂, 11♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.85-8.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25-2.70</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30-1.43</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.24</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.14-1.29</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82-0.95</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.61-1.73</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.22-1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.52-2.74</td>
<td id="7D9942C4B907C0ED41CCF7729BA81832" colspan="1" pageId="0" pageNumber="71" rowspan="1">6.85-8.40</td>
<td id="D949698EEF091CD57AB4E5DE7A4678C0" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25-2.70</td>
<td id="58BEA4E974E3E0CEAE083287538AC6F1" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30-1.43</td>
<td id="E276CECE962B7D57CDAE092CAA57729D" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13-1.24</td>
<td id="C8ABDAD07D039FAC949737F5C5736CB8" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32-1.41</td>
<td id="E4BE555B170859EBF5949F82C4555C36" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.14-1.29</td>
<td id="40E5C906EA9EC56A583B14C724A88297" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82-0.95</td>
<td id="D8168B786617AAE771819B4610A99622" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.61-1.73</td>
<td id="39646F901979B2A9DEE45E7AE9BFF1D5" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.22-1.36</td>
<td id="89581EB9DB001CBBEC331987C6129D79" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.52-2.74</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<tr id="70E367A40D9207E41558AC7C02E4DA7E" pageId="0" pageNumber="71">
<td id="0705A4DD1FBEDBF9E6BC76BCA1001473" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="F49189AC7A80B8029EFA54C64CE967DF" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="A74679E0E10582ABD81FAE68DB8C4E4F" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.53</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.68</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60</td>
<td id="6512B541B1BAE2EE869DDC5866382F0B" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.53</td>
<td id="7FCD6AC4B5D728A621CC3AF80E1DBF97" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45</td>
<td id="D4C3EFE98593A607195F6F9D9370BBA9" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td id="9B1143AB2259132E7C1BFDE1F894CC7F" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td id="21414FCDCB270F5C6DB90B2E3A30C76A" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td id="660B234A9B5EFA38735C6C09BEACEC4E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.20</td>
<td id="B4F8E7FE9886F159BDC12A52FF030007" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
<td id="E2E098F4EC92A1D692BD2EFF3508EC7E" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.68</td>
<td id="8B82FB25716B13D8D0E7BA2576932AAC" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30</td>
<td id="DEAA1E78C1C39477498A1C6D683BDCD3" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.60</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">P.</emphasis>
<tr id="209EB955BA9168605632B5359D8829A0" pageId="0" pageNumber="71">
<td id="FED7051FB7A5000AA07790FBCFC51B16" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="1D19CFA56F5271D212BB846F7B0303B4" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="3F325E66DD33FC7A8BD623E54FE1FE53" italics="true" pageId="0" pageNumber="71">P.</emphasis>
</taxonomicName>
sp. (1♀, Karabet Pass)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.05</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.12</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.76</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.67</td>
<td id="BEE74237B97861D12803301B9D154842" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.05</td>
<td id="72D3691DF8CD4ED2FF828E9453A40584" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.25</td>
<td id="B598B17B2E3FBEE7AC3460851CB82CE3" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
<td id="75C5CDA8A127B376288A9AB608250A97" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.12</td>
<td id="340B5992C6E1EC84A206CB5A0D112D95" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34</td>
<td id="EC8D07316A304C2D94570CE5C9160568" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td id="E6EDC6274DF3E30A4D7202A03472CC5A" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88</td>
<td id="3FEFBBB5FD610AEFDF0995439820B99F" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.76</td>
<td id="DC859481BB6B63D0F2EEDE34060C8955" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td id="BB9FB5E06A037CD45141302FAED2828F" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.67</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
<tr id="4923188ED15071C42888747FB39B45B8" pageId="0" pageNumber="71">
<td id="69D22309E621CB7050FE23FD7E26D7FF" colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName id="36E379A6E7A9044E07CF2B7CD577D755" class="Insecta" kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis id="7515AE058E247FB9D5C8FB7E3C0092A4" italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
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sp. nov. (1♂, 1♀)
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<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.30-7.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40-2.50</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38-1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.21-1.25</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41-1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.17-1.21</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83-0.85</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.66</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.63-2.71</td>
<td id="EF35C58093656B9CBC1BF93784F4269D" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.30-7.45</td>
<td id="C03EB5397A4B3EA39548E79A9758233D" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40-2.50</td>
<td id="99FDDBCE20BF8233EE7ECA7941E97FBC" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38-1.42</td>
<td id="35EF3A40CFB83C51D7487D8B238C4ACF" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.21-1.25</td>
<td id="0C5292666B9DEE2FA2C46D59AA682E78" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41-1.42</td>
<td id="CD2261729EE326A0B8C38831AD21935B" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.17-1.21</td>
<td id="218A26BECAA04396AB8D7BFA4DA9D899" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83-0.85</td>
<td id="1C6AD01723446C186A973B3266B07FBC" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63-1.66</td>
<td id="1E0658F21F19BA845BCEFBA837BFBEAA" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td id="B4052646C3450515FE49E1D09F8483B0" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.63-2.71</td>
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<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
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sp. nov. (mean)
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<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.65</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.67</td>
<td id="BADEA6E0222F9F9EA535C4D642896BF6" colspan="1" pageId="0" pageNumber="71" rowspan="1">7.38</td>
<td id="0286C1BC542F694B35CCE26541F6F6FB" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.45</td>
<td id="0EA0BD9D5245CA88D0CE3AC8E93C9ED9" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td id="4840DC11B050F1AF936B6C54E9BD0B8A" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.23</td>
<td id="82249ECFF9DCD369023CB6A4B7EAC067" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.42</td>
<td id="7B6C8246E9FA2E19AA62FBACA87E2711" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.19</td>
<td id="4C5C959EF3B2827122F790CEEC07B6ED" colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
<td id="B7DDCD533537AEA61B021E389D98FCE0" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.65</td>
<td id="39B8A1E02F88480691A883230CDC5925" colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td id="EED288208328C4F8AF6835C486E46364" colspan="1" pageId="0" pageNumber="71" rowspan="1">2.67</td>
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</paragraph>
<paragraph pageId="0" pageNumber="71">
<tableNote pageId="0" pageNumber="71">For abbreviations in line 1, see &quot;Abbreviations to measurements and ratios&quot; (section &quot;Material and methods&quot;).</tableNote>
<paragraph id="21C91573821507A6F877DDFB246F076B" pageId="0" pageNumber="71">
<tableNote id="5BC92DB39E32DFCBB99EFA1F8F7B972C" pageId="0" pageNumber="71">For abbreviations in line 1, see &quot;Abbreviations to measurements and ratios&quot; (section &quot;Material and methods&quot;).</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">The specimens were hand collected by R. Felix and J. Muilwijk close to the village Do Polan along the river course. The vegetation there consisted of (over)-grazed grassland, fields and small orchards; characteristic of a cultivated landscape.</paragraph>
<subSubSection id="567E6D919AB3A875A2363567456540DC" pageId="0" pageNumber="71" type="habitat">
<paragraph id="11651FECEE623CB1770CB7DA95EB925D" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="99F7712DF3B51C69AB601FB782CCF552" pageId="0" pageNumber="71">The specimens were hand collected by R. Felix and J. Muilwijk close to the village Do Polan along the river course. The vegetation there consisted of (over)-grazed grassland, fields and small orchards; characteristic of a cultivated landscape.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="CB653FC55B05C0F5763F04D0AEE3597B" pageId="0" pageNumber="71" type="distribution">
<paragraph id="67E3DD53D27BCF1DABE8DE40BEEA9546" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="72B45010FE6B0A9B3C05E371E9555EA4" pageId="0" pageNumber="71">
Southwest Iran: Central Zagros Mountain Chains (Chahar Mahal va Bakhtiari Province, Kiar County; Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="9F7D2888AC36AD47B6DE096BF5EB209B" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
).
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466
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<treatment id="B13F5DA3FB16552EB4E773FA121E6038" ID-GBIF-Taxon="195868748" LSID="urn:lsid:plazi:treatment:B13F5DA3FB16552EB4E773FA121E6038" httpUri="http://treatment.plazi.org/id/B13F5DA3FB16552EB4E773FA121E6038" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
<subSubSection id="3D548F91DFF313D2E1DEA18B508B0CA9" pageId="0" pageNumber="71" type="nomenclature">
<paragraph id="BA931482D790AAD49758CBD0C57F0681" pageId="0" pageNumber="71">
11.
<taxonomicName LSID="B13F5DA3-FB16-552E-B4E7-73FA121E6038" authority="Morvan, 1974" authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="Eremoderus">Platyderus (Eremoderus) taghizadehi Morvan, 1974</taxonomicName>
<taxonomicName id="DB1790DF98B23F7331953BA55DE50D68" ID-CoL="4JQL9" LSID="B13F5DA3-FB16-552E-B4E7-73FA121E6038" authority="Morvan, 1974" authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="Eremoderus">Platyderus (Eremoderus) taghizadehi Morvan, 1974</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2E</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5F</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7G</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8I</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10I</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12H</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14C</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
<subSubSection id="EC54E67FAAF5306428F9D487C786D91F" pageId="0" pageNumber="71" type="description">
<paragraph id="BCA4E3E0AB2DF3217A20AB839F824449" pageId="0" pageNumber="71">
<figureCitation id="A0CCF01D6EA4F0FDA7ADB7C12D82BACF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2E</figureCitation>
<figureCitation id="C2F28E9F08E2E470A30B3C056C8761BD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5F</figureCitation>
<figureCitation id="ED50448425148BA58E63AC8D8D77978C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7G</figureCitation>
<figureCitation id="8467D81C3E212A99CFC67DBA7E515CA5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8I</figureCitation>
<figureCitation id="F67871210D9DD56ECA30533960FCD6D7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10I</figureCitation>
<figureCitation id="CA59AE7E621F6E6CF93A6825153AC19C" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12H</figureCitation>
<figureCitation id="948FB3D3278468F92F1B61DD6B283230" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14C</figureCitation>
<figureCitation id="D72CB2CA3B41E3C4872B1EB7EF0AE3DF" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="reference_group">
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">Platyderus taghizadehi</taxonomicName>
<subSubSection id="262240561DF8F2B7082D61D3EA2DAB19" pageId="0" pageNumber="71" type="reference_group">
<paragraph id="A710DA63B2DDBF5B430855A40829204C" pageId="0" pageNumber="71">
<taxonomicName id="28B85E8E04C77CBA73BBBF070ADB0D8B" authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">Platyderus taghizadehi</taxonomicName>
Morvan, 1974: 147 (type locality:
<normalizedToken originalValue="“l’Elburz">&quot;l'Elburz</normalizedToken>
<normalizedToken id="A1D1A29F1DF8E5062A09C694B6892727" originalValue="“l’Elburz">&quot;l'Elburz</normalizedToken>
, massif du Sutak-kuh, Dizine 3500 m&quot;).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="8F1C907EE534FFB9D642FDDD68C8877E" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="02B7B9BED767F3A1DED9170D9AFCD511" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="5C233A0B80D62AC81830C5657622CE1E" pageId="0" pageNumber="71">
The Mount Sutak Kuh is situated in the province of Mazandaran (southwestern part of Nur County), and lies in the Central Alborz Range, at ca. 3500 m altitude. Its approximate GPS coordinates are 36.153, 51.456, referring to a point situated ca. 300 m higher than the height Morvan mentioned (ibid.). The type locality locates somewhere between the Sutak Kuh Peak, in the north, and the
<normalizedToken originalValue="Dīzīn">Dīzīn</normalizedToken>
<normalizedToken id="72CC0E059597033F02C846A3E20C87C4" originalValue="Dīzīn">Dīzīn</normalizedToken>
Ski Resort, in the south. Roughly, it may be defined as a site in the plot
<normalizedToken originalValue="Kandovān">Kandovan</normalizedToken>
<normalizedToken id="1D0ADEBB9A86E25C7302BB24CF480E13" originalValue="Kandovān">Kandovan</normalizedToken>
Tunnel -
<normalizedToken originalValue="Āzād">Azad</normalizedToken>
<normalizedToken id="4B2E2783DFB636479474327094A72736" originalValue="Āzād">Azad</normalizedToken>
Kuh Peak - Kholeno Lake -
<normalizedToken originalValue="Dīzīn">Dīzīn</normalizedToken>
<normalizedToken id="1BB3404DFAA9033C303829EFC7D5748A" originalValue="Dīzīn">Dīzīn</normalizedToken>
Ski Resort. Spatially, it is a point geographically situated midway between the type localities of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="271BCAE8A898AD54F8ECE73C9787F231" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="7B893F0A870B34DBE0671F3A21534666" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
(vicinities of Rudbarak) and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
<taxonomicName id="647B6E507F9CF6486D1E8D0C123C15D8" class="Insecta" kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="0A597C4C814255E21F7FA2E7BBCF2FB8" italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
</taxonomicName>
sp. nov. (vicinities of Damavand City).
</paragraph>
<caption doi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 11.</emphasis>
<caption id="14B6F9B47AC3BD04F1DD184ED18F647A" doi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71" start="Figure 11" startId="F11">
<paragraph id="443FE92A49E9BAA03DFF0399AC604C97" pageId="0" pageNumber="71">
<emphasis id="D7003A3C49BC1B20B8BDE015E2D87368" bold="true" pageId="0" pageNumber="71">Figure 11.</emphasis>
Median lobe of aedeagus, ventral view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">Platyderus (Eremoderus) afghanistanicus</taxonomicName>
<emphasis id="0CFF0D39D86A5C4BEC5531591F7CE78F" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="192ED66D6A8B883B95FC234ADF3AD25B" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">Platyderus (Eremoderus) afghanistanicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
<emphasis id="07D0D80AAECC96DB027FB8FC18A240D1" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="EA3AACFA00E68A3AD1C9738A02399581" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="E08C93F547A29EAF4E12697B358707FD" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="6B69B9BB9C64076B4A691FCFA433E730" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName authority="insignitus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="E." subSpecies="insignitus">P. (E.) insignitus insignitus</taxonomicName>
<emphasis id="C4F60C614D410E199D5629DB5B6A7714" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="5400A2C26AD0DC1F2D55C55DAB8368BD" authority="insignitus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="E." subSpecies="insignitus">P. (E.) insignitus insignitus</taxonomicName>
Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
<emphasis id="FF286824691F18BDE031B5DF82CF9BE5" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="6AA201FDAC1853784DCE2B255B5F8187" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="DE190A0FB70000FCEED05F7D3E0633E6" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="7FE623A890E226305736630152422337" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="AAE751E094EE26896E1AD85E4796320C" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="805018BDA8398BC3EF3A6EAFC76BB45B" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), male specimen, Naẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus taghizadehi</emphasis>
<subSubSection id="05011C56AD112C91D97F29811D3F770E" pageId="0" pageNumber="71" type="references">
<paragraph id="F9268B717EC28DB3F214A3C86BBFC4F9" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="5363393F78AB59C2D1551AB1B281D9A3" pageId="0" pageNumber="71">
<taxonomicName id="438135D9C58D4E66AF09087CB21AAC5B" authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="68BA00EC5187ECF10391BE3FF6790B7F" italics="true" pageId="0" pageNumber="71">Platyderus taghizadehi</emphasis>
</taxonomicName>
:
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="C3412754DFF51CEC7F557E9B1F5311EC" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="797E94000EDE4C2A285C54FFF2B9CF9C" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 523;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="CD98FE860F44147FF9EF6FAC2DCDD74C" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 396;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4024.1.1" author="Azadbakhsh, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="1 - 108" refId="B9" refString="Azadbakhsh, S, Nozari, J, 2015. Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae). Zootaxa 4024 (1): 1 - 108, DOI: https://doi.org/10.11646/zootaxa.4024.1.1" title="Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae)." url="https://doi.org/10.11646/zootaxa.4024.1.1" volume="4024" year="2015">Azadbakhsh and Nozari 2015</bibRefCitation>
<bibRefCitation id="9D1370D93761C418FF82048B576D978E" DOI="https://doi.org/10.11646/zootaxa.4024.1.1" author="Azadbakhsh, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="1 - 108" refId="B9" refString="Azadbakhsh, S, Nozari, J, 2015. Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae). Zootaxa 4024 (1): 1 - 108, DOI: https://doi.org/10.11646/zootaxa.4024.1.1" title="Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae)." url="https://doi.org/10.11646/zootaxa.4024.1.1" volume="4024" year="2015">Azadbakhsh and Nozari 2015</bibRefCitation>
: 84;
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="E2CA4CDB911028BF3E74C6F24E575D69" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 759.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="C1A386D8AF62FC321CFFB3C7740AF77A" pageId="0" pageNumber="71" type="type material">
<paragraph id="A2D902DF6BFB5EF79DDB146AC300FF73" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="C656EAF1103E9600FC9CBB8F4B04C05E" pageId="0" pageNumber="71">
<emphasis id="38D7E02C259BC592E03B0F4479705A43" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂ in private collection Pierre Morvan (Carentoir, France). Not examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Muilwijk J, Morvan, Muilwijk" country="Iran" elevation="3450" latitude="36.0335" location="Alburz Dizin" longLatPrecision="1" longitude="51.43347" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tehran Province">
<emphasis bold="true" pageId="0" pageNumber="71">
<collectingCountry name="Iran">Iran</collectingCountry>
<subSubSection id="B8653E42588126EEE2E48AC99CBE96EF" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="17AAC0872CA0BD2F21E603D23038D85D" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="285384E70F6F6B6520BDA9018BBEB2B7" pageId="0" pageNumber="71">
<materialsCitation id="470FC8E7EF3A38B1BBCB46DBDAA2F864" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Muilwijk J, Morvan, Muilwijk" country="Iran" elevation="3450" latitude="36.0335" location="Alburz Dizin" longLatPrecision="1" longitude="51.43347" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tehran Province">
<emphasis id="B86044A30F2FEA854F61BDD286F6320B" bold="true" pageId="0" pageNumber="71">
<collectingCountry id="CA96389EB9C9D19629DE502BAC7DABBA" name="Iran">Iran</collectingCountry>
</emphasis>
:
<collectingRegion country="Iran" name="Tehran">Tehran Province</collectingRegion>
<collectingRegion id="45839DEC82DE5B522394F66E93FFBCE6" country="Iran" name="Tehran">Tehran Province</collectingRegion>
:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="821CE293ADC456680DC2AFC217607CBD" type="female">1♀</specimenCount>
,
<normalizedToken originalValue="ʻIR">ʻIR</normalizedToken>
<location LSID="urn:lsid:plazi:treatment:B13F5DA3FB16552EB4E773FA121E6038:E7A467FE33A9AB606162C00D79F3B19F" country="Iran" latitude="36.0335" longLatPrecision="1" longitude="51.43347" name="Alburz Dizin" stateProvince="Tehran Province">Alburz Dizin</location>
<normalizedToken id="1E0009F33A33F66C470B51FAD54B7A81" originalValue="ʻIR">ʻIR</normalizedToken>
<location id="E7A467FE33A9AB606162C00D79F3B19F" LSID="urn:lsid:plazi:treatment:B13F5DA3FB16552EB4E773FA121E6038:E7A467FE33A9AB606162C00D79F3B19F" country="Iran" latitude="36.0335" longLatPrecision="1" longitude="51.43347" name="Alburz Dizin" stateProvince="Tehran Province">Alburz Dizin</location>
/
<geoCoordinate degrees="36" direction="north" minutes="2" orientation="latitude" precision="1" seconds="00.6" value="36.0335">36°2'00.6&quot;N</geoCoordinate>
<geoCoordinate id="B5F6BB17DB8FACDBED4212ABE492878D" degrees="36" direction="north" minutes="2" orientation="latitude" precision="1" seconds="00.6" value="36.0335">36°2'00.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="26" orientation="longitude" precision="1" seconds="00.5" value="51.43347">51°26'00.5&quot;E</geoCoordinate>
<geoCoordinate id="27F5D5602E4056788550A07481BFBCB5" degrees="51" direction="east" minutes="26" orientation="longitude" precision="1" seconds="00.5" value="51.43347">51°26'00.5&quot;E</geoCoordinate>
/
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" unit="m" value="3450.0" valueMax="3600.0" valueMin="3300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" unit="m" value="3450.0" valueMax="3600.0" valueMin="3300.0">3300-3600m</elevation>
<quantity id="74B058954C40F27CA9EC4C58AD247867" metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" unit="m" value="3450.0" valueMax="3600.0" valueMin="3300.0">
<elevation id="D623ABDDEBAADE4DBE6104DAC5AD8D5F" metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" unit="m" value="3450.0" valueMax="3600.0" valueMin="3300.0">3300-3600m</elevation>
</quantity>
10.06.17 /
<collectorName>Muilwijk J</collectorName>
<collectorName id="1398A842FB9121DE1A7F3758B5E37B10">Muilwijk J</collectorName>
//
<taxonomicName authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">Platyderus taghizadehi</taxonomicName>
<taxonomicName id="9600741950C0D2014D4D2480F9D6AEBF" authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">Platyderus taghizadehi</taxonomicName>
/
<collectorName>Morvan</collectorName>
<collectorName id="E5F9D282D0CF289D9B704185D409AC22">Morvan</collectorName>
/
<collectorName>Muilwijk</collectorName>
<collectorName id="A8EDC5C5E2BD9206427A5843C099A36A">Muilwijk</collectorName>
2017' (cMUI);
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="E80E7172E752480674184C5BF038A625" type="male">1♂</specimenCount>
, 'IR
</materialsCitation>
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Seiedy, Muilwijk, Morvan" country="Iran" elevation="3550" latitude="35.890614" location="Tochal" longLatPrecision="1" longitude="51.406975" specimenCount="1" stateProvince="Tehran">
<collectingRegion country="Iran" name="Tehran">Tehran</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:B13F5DA3FB16552EB4E773FA121E6038:81D9E8F00E420D36654068535B3B1D58" country="Iran" latitude="35.890614" longLatPrecision="1" longitude="51.406975" name="Tochal" stateProvince="Tehran">Tochal</location>
<materialsCitation id="8F3DEEFAB809983B4826847B14EB5563" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Seiedy, Muilwijk, Morvan" country="Iran" elevation="3550" latitude="35.890614" location="Tochal" longLatPrecision="1" longitude="51.406975" specimenCount="1" stateProvince="Tehran">
<collectingRegion id="1625406D92938526B953F3919170907E" country="Iran" name="Tehran">Tehran</collectingRegion>
<location id="81D9E8F00E420D36654068535B3B1D58" LSID="urn:lsid:plazi:treatment:B13F5DA3FB16552EB4E773FA121E6038:81D9E8F00E420D36654068535B3B1D58" country="Iran" latitude="35.890614" longLatPrecision="1" longitude="51.406975" name="Tochal" stateProvince="Tehran">Tochal</location>
/
<geoCoordinate degrees="35" direction="north" minutes="53" orientation="latitude" precision="1" seconds="26.2" value="35.890614">35°53'26.2&quot;N</geoCoordinate>
<geoCoordinate id="F6371E221181254AE1FBEBA5A760482A" degrees="35" direction="north" minutes="53" orientation="latitude" precision="1" seconds="26.2" value="35.890614">35°53'26.2&quot;N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="24" orientation="longitude" precision="1" seconds="25.1" value="51.406975">51°24'25.1&quot;E</geoCoordinate>
<geoCoordinate id="2A3BB106CEEF9E5ADFA07B3F368DA619" degrees="51" direction="east" minutes="24" orientation="longitude" precision="1" seconds="25.1" value="51.406975">51°24'25.1&quot;E</geoCoordinate>
/
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.55" unit="m" value="3550.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.55" unit="m" value="3550.0">3550m</elevation>
<quantity id="7D45F587DDE79FB78CD826F72DDDF605" metricMagnitude="3" metricUnit="m" metricValue="3.55" unit="m" value="3550.0">
<elevation id="F778505AB723C4ABE20336F83BC660B7" metricMagnitude="3" metricUnit="m" metricValue="3.55" unit="m" value="3550.0">3550m</elevation>
</quantity>
27.06.17 /
<collectorName>Seiedy</collectorName>
<collectorName id="04B3BBA6D5A6D672243A07A9E124CD12">Seiedy</collectorName>
/
<collectorName>Muilwijk</collectorName>
<collectorName id="088A8992DBDB299EA4BA42E5954CD855">Muilwijk</collectorName>
//
<taxonomicName authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">Platyderus taghizadehi</taxonomicName>
<taxonomicName id="8B74A5C7B1A97AA65B95D25F21B09FDC" authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">Platyderus taghizadehi</taxonomicName>
/
<collectorName>Morvan</collectorName>
<collectorName id="80A61A8E1C1E835D0520CBD92EA2F4CD">Morvan</collectorName>
/
<collectorName>Muilwijk</collectorName>
<collectorName id="D569083F584B92C81E33572F1F457D18">Muilwijk</collectorName>
2017' (cMUI)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="CE1F8B69BF89178DF1E4CA6D94BF029E" pageId="0" pageNumber="71">
TME:
<specimenCount type="generic">2 specimens</specimenCount>
<specimenCount id="461796F93D25294B9B323A4A5951FDEA" type="generic">2 specimens</specimenCount>
. TGE:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="B617FC26270CBFC68520681B168179D1" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="6F0FDE6A9BBD396BE90123956C58E435" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="0601248433876CA198B8E2264189ECD5" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="6E5E097E24F81D7B856C560603A96DE8" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="F3E9BC0A777ADC69120DD03C71B11979" pageId="0" pageNumber="71">
A species of medium size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="3B42805094E6574F653D8BCA68FC8C33" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="726010F97778A98D7E3836C83C7B5EED" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
, with anterior side of mesofemur ventrally with three setiferous punctures. It is distinct from the closely related
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="2A8DF6A9E500AE49490F7BF3E846B2E9" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="E8FE624774DAF99D958215C64E0110E1" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
by pronotum about a third wider than head (PW/HW= 1.31-1.33), with apex compared with widest point less constricted (PW/PA &lt;1.40), and elytra in relation to pronotum significantly wider (EW/PW&gt;1.38). See also
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="07A89AC8C8E501BF81829DC05D95D6AE" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
under
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="EFC320BE36D7D59E5CA3CD635920B5BB" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="63A5AF40FF83FBE4324F87CF47DC414C" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="redescription">
<paragraph pageId="0" pageNumber="71">Redescription</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="669DFEB027B9513D9296AF8374AED4A0" pageId="0" pageNumber="71" type="redescription">
<paragraph id="12C164B6742D7C93D0B85399E1AB7E9F" pageId="0" pageNumber="71">Redescription</paragraph>
<paragraph id="DF9ECB7CACBF216C7D0185F74902F489" pageId="0" pageNumber="71">
(based on non-type material).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<emphasis id="F320D43A838F97AC4D2EC591BEDDC809" bold="true" pageId="0" pageNumber="71">
<emphasis id="7CAB9CEA920779279121B2CEF4CF3545" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Moderately large-sized specimens for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="EA4C84D7D0E155181AAA73DD76F8A945" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="1E67933B24B835931ABCA32C44DB0F3F" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 7.10-7.20 mm; BW: 2.40 mm), with elongate and slender body (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2E</figureCitation>
<figureCitation id="12D54D89BD10379EBD3E542C8BF02F44" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="7F296314C5B999B272DE24E6319AB718" bold="true" pageId="0" pageNumber="71">
<emphasis id="4E427D7C127057A777EEBE634F55BE6E" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
<tableCitation id="BFC13FAEA5CD9B1D6DB6E19D75A9F012" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="779D1323F44E7124D397437CB67F4A6C" bold="true" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
. Integument of body orange-brown, appendages slightly lighter than body. Surface moderately shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="750501B4B374E82D99082EB709000051" bold="true" pageId="0" pageNumber="71">
<emphasis id="FE56C240695BF0781FE7D00A5320EF6C" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum largely with reduced sculpticells or without microsculpture, traces of isodiametric sculpticells present only laterally. Ventral surface (excluding gula and prosternal process) microsculptured, genae, mentum, submentum, proepisternum and ventrites 2-3 medially with regular isodiametric meshes, abdominal ventrites laterally and legs with slightly stretched isodiametric sculpticells. Prosternum scarcely punctate.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="FF3AB7FA6FE92FBDAC0C3FD3BC18DD72" bold="true" pageId="0" pageNumber="71">
<emphasis id="A9AEE91EF32DCF78ED918E37305052DB" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
Two thirds as wide as pronotum (mean PW/HW= 1.32).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="B5CC3A28EB63A520E836A4895622E115" bold="true" pageId="0" pageNumber="71">
<emphasis id="E05609719234143F752D0DF3738FD500" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
Significantly narrower than pronotum (mean PW/HW= 1.48). Antennae long, with last three antennomeres exceeding base of pronotum. Eyes long, little convex. Labrum subrectangular, with anterior margin concave. Frontal furrows small, subfoveolate. Paraorbital sulci moderately deep, ending slightly before level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="CCD397AF217550F51C8F8A4C3EEC5A14" bold="true" pageId="0" pageNumber="71">
<emphasis id="5D9D1BF199434DE6022F42E42AF6C753" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum sub-quadrate, barely wider than long (mean PW/PL= 1.09), widest at second quarter. Anterior and posterior transverse impressions indistinct. Sides rather sinuate, convex anteriorly, concave posteriorly; mean PW/PA= 1.35, mean PW/PB= 1.18; bead of anterior margin present laterally, reduced in medial 1/6; bead of posterior margin reduced to absent in medial 1/5 to 1/3. Metepisternum longer than wide, MA/MI about 0.9.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="E13CC61C7A673EE18311E154A20C1158" bold="true" pageId="0" pageNumber="71">
<emphasis id="29D99F123968BFAEE9B8F8BBAF57A741" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Long, cylindrical, one and two thirds as long as wide (mean EL/EW= 1.67), wider and much longer than pronotum (mean EW/PW= 1.42; mean EL/PL= 2.57). Stria 7 not reach basal margin. Elytral interval 3 with three discal setiferous punctures (specimen from Dizin with four punctures on left elytron, as an additional one between second and third normal puncture) located as follows: anterior puncture adjoining stria 2, stria 3 or in middle of interval 3, medial puncture adjoining stria 2 or in midst of interval 3, posterior puncture adjoining stria 2. Umbilicate setiferous series consisting of 16 punctures on each elytron in specimen from Tochal, and with 15 punctures on each elytron in specimen from Dizin.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="C0B629F5ADC65CDF50FB29DDC826523C" bold="true" pageId="0" pageNumber="71">
<emphasis id="ECF6E641E82E728666352B03C2DF09E3" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur with three setiferous punctures on anterior side ventrally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5F</figureCitation>
<figureCitation id="321F569EA9AB13373CEE64B98C2411D4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5F</figureCitation>
), but one specimen has two punctures on one mesofemur. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="4017FCAF2D25EFB2CF4C24A7D62A1F12" bold="true" pageId="0" pageNumber="71">
<emphasis id="0B97CE4113B2061817F08023D160B61D" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX small, suboval, with proximal margin slightly asymmetrical (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7G</figureCitation>
<figureCitation id="EB4C04950B8C3597EB1DE04EF8CA6CE8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7G</figureCitation>
). Median lobe of aedeagus in lateral view with shaft shorter than that in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="3C5E3C077D3CB7AC55DD626A3A3FA4A0" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="231FCAB738A5B1D7D631D9CD1FCC3E03" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
and more curved than that in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="040BE87914B09D33E92BF5BA1875654C" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="A9804C9DB760C9E1E8A995674950CFBB" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
sp. nov., and with apex turned up; median lobe in ventral view straight, ca. 3.7 times longer than wide, with apical lamella slightly asymmetrical. Internal sac of same structure as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="E588717AE039FCD563597CE15D7F0285" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="54DCEFE7CDC4006808EBA57B07CC5BD6" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
, but dorsal sclerite in ventral view with left-sided protuberances somewhat larger. Right paramere shorter and more crooked than that of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="3A43BE0401D4F7DC94870951480E59B2" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="D3DB50BDEFE22FEC9CAEF165DC1847D2" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12H</figureCitation>
<figureCitation id="5DEBB40BFF3E90E3E61FEB3BF4B2B870" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12H</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="8DAA610A914C6B0C9F6FE93267FEE50F" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14C</figureCitation>
<figureCitation id="114336F7632019065B9E3C8EF2F4A94F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14C</figureCitation>
). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Bursa copulatrix smaller than that of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="C921EEE969C16E68AAFBA1FA7A79358E" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="820028B459184B38C442522396A7795D" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">All specimens hitherto known were found in open places at high altitudes between 3300 m and 3600 m. The two specimens collected by JM in June 2017 were caught from under stones in rather steep green mountain meadows (without bushes or trees). At the time of its collection the place was not dry, probably due to the thaw just a few weeks before and to some water streamed down.</paragraph>
<subSubSection id="DA4829C0D1B7F0CEA84485A4437F5DCB" pageId="0" pageNumber="71" type="habitat">
<paragraph id="E501F3DB8434DDF268E49BFAA32705BE" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="C673EC458DD05C66DEF72F99568B4347" pageId="0" pageNumber="71">All specimens hitherto known were found in open places at high altitudes between 3300 m and 3600 m. The two specimens collected by JM in June 2017 were caught from under stones in rather steep green mountain meadows (without bushes or trees). At the time of its collection the place was not dry, probably due to the thaw just a few weeks before and to some water streamed down.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="56A93F1B7110286175F55E861FD9F6DE" pageId="0" pageNumber="71" type="distribution">
<paragraph id="24E741931CB160FF3FDFE9F61C259A1B" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="9E900B675F136B6FF146FEEF825965A8" pageId="0" pageNumber="71">
North Iran, Central Alborz Range (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="91C6DBEC01D0D9CB9DEDE4DD04AF7032" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
). The species occurs in the area where three Iranian provinces, Mazandaran, Alborz and Tehran, are touching each other.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="0FF133788D22BABF03F513EFC72F3967" pageId="0" pageNumber="71" type="notes">
<paragraph id="41B22E1C2238952454ABDD5300D2967A" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="B2FA98D6F2BFB926733F3F9AA82B7FB6" pageId="0" pageNumber="71">
Three metric measurements made by
<bibRefCitation author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan (1974</bibRefCitation>
<bibRefCitation id="FB3B6EAA9B267EAB931EB9DCFFD9D119" author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan (1974</bibRefCitation>
: 147), i.e. width (1.15 mm), length (1.13 mm) of pronotum and length of elytra (2.1 mm), are certainly wrong. If these are correct, then we should have a total length of the pronotum and elytra 3.23 mm, but then the length of head should be at least 4 mm as far as the holotype of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="94E90609C162D610607CBE4FEDF5BA84" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="6AECE76C0C667EFFF8AAF072720FAA07" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
has length of body 7.5 mm (ibid.: 147).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="D52756E63421710D21C1BBF1A2AF258A" pageId="0" pageNumber="71">
Being a male specimen, the holotype of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="E56E42E51E09B0848A08D67EF53BDBFB" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="4EF0DEF57F73DA9756BD83231622B1E9" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
(or topotypic males) must be studied for two reasons. Firstly, this investigation can support the current taxonomic affinity with
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="D61D56D9570DC022CB7C3F5C0E4140CC" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="1908749FA4284F0F22C6620338F5AA0B" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
because the distance in a straight line between the population of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="DFE3A7394FBED06A94069D5BC2CD5DB6" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="A70D511A430045D05B3D0A6EB70308A1" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
southwest of Rudarbak and the type locality of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="63164FBDFD5C0296FDC964CCC8845535" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="AE2AF5FDB18710B8AA67EF83E2BE1B5C" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
is only about 50 kilometers and it is unclear whether the Chalus River acts as a biogeographical barrier. Furthermore, it is necessary to confirm the conspecificity of the populations from Sutak Kuh Peak and Tochal Peak (here last referred to as
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="0ACDBB871B59C5F0D6852D6C18D98E70" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="C0998698EA9F03AD6B8823500E644084" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
), so that the two local forms of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="AADBC49BFAE13192233016F6EF34A91A" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="8E5973245ECBA01DFD3D4325D6E5089F" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
<taxonomicName id="A2CC7449DF64672E788FDEB6241C0A54" class="Insecta" kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="0AEAA4F7B177A98248D4F4E52BCFD10E" italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
</taxonomicName>
sp. nov. (from Damavand County).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="F806ABB4E346A2E5FEA5755B943857B5" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus afghanistanicus</emphasis>
<emphasis id="45DCE4A568257EC7A8D4261965658291" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="E13BF6CEA5B2DFECA7FC37B80A0F6812" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="Eremoderus">
<emphasis id="F1F36D6F3F988E4B5EAB0C48D7ABE427" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus afghanistanicus</emphasis>
</taxonomicName>
&quot; species group
</emphasis>

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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1F</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7D</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8F</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10F</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12E</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13F</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
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<figureCitation id="52A82E2C92DDF3C8218C2E7CB373C604" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1F</figureCitation>
<figureCitation id="545C332663684AC7F6266AED6F8BA05E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7D</figureCitation>
<figureCitation id="0D81450575C6D599D73D3EE68EE72AAC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8F</figureCitation>
<figureCitation id="C0AE46343FF5C4AEFED34BDC19AF6496" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10F</figureCitation>
<figureCitation id="9C2347713A3450E924B63FDEEFAC3D2D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12E</figureCitation>
<figureCitation id="BD61B4F2AAF949E687423BDB216D15F2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13F</figureCitation>
<figureCitation id="23744C7B3C4C8434FB7C25F0471831B5" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">, 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="779DA292A9EE65E86DF79F0C8ED973FE" pageId="0" pageNumber="71" type="type locality">
<paragraph id="1171D153B6B45E6B4F082B74D00C07D3" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="808507841EF92C705E3BBE0AAB95BF14" pageId="0" pageNumber="71">
Turkey,
<normalizedToken originalValue="Adıyaman">Adiyaman</normalizedToken>
<normalizedToken id="3061F4110EAE32DD158A123388FC96C4" originalValue="Adıyaman">Adiyaman</normalizedToken>
Province, Nemrut
<normalizedToken originalValue="Dağı">Dagi</normalizedToken>
<normalizedToken id="88A4B7C5C6026FBB2AA40FB5CC1B4002" originalValue="Dağı">Dagi</normalizedToken>
, NE of
<normalizedToken originalValue="Adıyaman">Adiyaman</normalizedToken>
<normalizedToken id="2DF528FFDA9D791770FA69C15ACAE318" originalValue="Adıyaman">Adiyaman</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="334012E56D2D735C2A053E56A50B3DAE" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="8775ACF36CBB479F6229D1B12BAF4E6D" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="AED1457B76E707C7CC5CA81DBD39112B" pageId="0" pageNumber="71">
The Nemrut
<normalizedToken originalValue="Dağı">Dagi</normalizedToken>
<normalizedToken id="81FB5F57A81C5405962D93E91466EC68" originalValue="Dağı">Dagi</normalizedToken>
, a mount in Southeastern Anatolia, elevated over 2150 m a.s.l., not far from the upper reaches of the Euphrates. It belongs to the Taurus Mountains and lies 86 kilometers northeast of
<normalizedToken originalValue="Adıyaman">Adiyaman</normalizedToken>
<normalizedToken id="3F09FB8CCD024181FD21D6176DF8021E" originalValue="Adıyaman">Adiyaman</normalizedToken>
in the province of the same name.
</paragraph>
<paragraph pageId="0" pageNumber="71">The GPS coordinates indicated on original labels are imprecise. They were taken from a map after the time of collecting (V. Vrabec, pers. comm.).</paragraph>
<paragraph id="564C42CBFC5D0E3F4401CDA3A80B0C99" pageId="0" pageNumber="71">The GPS coordinates indicated on original labels are imprecise. They were taken from a map after the time of collecting (V. Vrabec, pers. comm.).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="ED18335F7D848615AF1F093A75578FF5" pageId="0" pageNumber="71" type="type material">
<paragraph id="BCCF393D9AD4F32A3BDF229EAF313B53" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="C1D50B8305CFCCAF8B3D009589B2F764" pageId="0" pageNumber="71">
<emphasis id="582246C3E24ADED4A6B6F187F1951FC4" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂, 'S Turkey: NEMRUT DAGI / (NE from Adiyaman), UTM: DC60 / 38.00N/38.35E, 1700-1900 m / mount. pastures, stone fields / 27.-28.IV.1997, V. Vrabec lgt.' (cWR). Paratype: 1♀, labelled as holotype (NMNHS).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 2 specimens. TGE: 1♂, 1♀.</paragraph>
<paragraph id="634DC3B624CB4401C3AE68464E51781E" pageId="0" pageNumber="71">TME: 2 specimens. TGE: 1♂, 1♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="D99DDDA9D6AC0211B6C9D144F0371ACB" pageId="0" pageNumber="71" type="etymology">
<paragraph id="FDDB445C8D1D0092B4CA415FC559DBA5" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="627784F6EAFA2DD91F87E491335CD1A3" pageId="0" pageNumber="71">
Patronymic, named after
<normalizedToken originalValue="Vladimír">Vladimir</normalizedToken>
<normalizedToken id="6A0F8FD98DDADFE99F1303C0A5058A30" originalValue="Vladimír">Vladimir</normalizedToken>
Vrabec, an entomologist interested in beetles and butterflies, who collected the type series of the new species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="958E1A7A6ACE5B833FE79F4F94196619" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="5C0E6E84BDCABC1B2582170F8F37E5F1" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="6024A773DE78B1040D13BCB001904F7F" pageId="0" pageNumber="71">
It differs from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="1230A7E45B349DB95F2D2A229ED95156" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="FE5BE2ECF0208DFFF6D534C6D20BD471" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. by the uniformly lighter coloration of the body, by the elytra and strioles much less coarsely and deeply punctate, and by slightly higher value of ratio PW/
<normalizedToken originalValue="PА">PA</normalizedToken>
<normalizedToken id="C771D1CDF8F5B8D33EFBF827F442D4D8" originalValue="PА">PA</normalizedToken>
(1.41-1.42, vs. 1.32-1.41). The apical lamella in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
<taxonomicName id="15C3DE0A9E0A3714781241803A4F5087" class="Insecta" kingdom="Animalia" lsidName="P. vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci">
<emphasis id="1975E205105FEDF0646696A2D5D5669C" italics="true" pageId="0" pageNumber="71">P. vrabeci</emphasis>
</taxonomicName>
sp. nov. is also more symmetrical and elongate (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10F</figureCitation>
<figureCitation id="214CDC2A415B07CD3F0E67B0D597C0FB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10F</figureCitation>
) and the bursa copulatrix is rounded (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13F</figureCitation>
<figureCitation id="B9C743274D95AB2F9DB0161256D0BDE8" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="3DCFD3D8C7A4B2BE54F242D1923C1FFA" pageId="0" pageNumber="71" type="description">
<paragraph id="5645A8F9CDDBE2B65BB9450D96EFA065" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="0FD91F6C0AE7B2DF8E65EFB98C2F1BF8" pageId="0" pageNumber="71">
<emphasis id="58A59F11BBE6EC97094322FB5D74C918" bold="true" pageId="0" pageNumber="71">
<emphasis id="AC6A9F7B43003050558A86B8C474D55D" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of relatively large size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="574268307A4CF613381F0C187663283C" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="61B1A72864588D7984D22619A19103F7" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 7.30-7.45 mm; BW: 2.40-2.50 mm), with elongate, moderately convex body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1F</figureCitation>
<figureCitation id="9DCBA12C7B3A005AFAD304672BEB53B8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="06C91C065FE108F479EA938C5AE0E414" bold="true" pageId="0" pageNumber="71">
<emphasis id="32915C68829183826268EC06ADCE8048" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
<tableCitation id="AFF89A59302E19A9227F8D332D4CBCCA" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="D6482D214B67DCA9CFA9EAF7B4CEA5E6" bold="true" pageId="0" pageNumber="71">
<emphasis id="AF723DCF02813B078662D88CE3618B2F" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages rusty red, head slightly darker than pronotum and elytra. Integument moderately shiny, head and pronotum shinier than elytra.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="DB7287C049CB1CEF0B3E5DF380CA931C" bold="true" pageId="0" pageNumber="71">
<emphasis id="2B957868990068E05DB4EA4F2FF5AE88" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum without microreticulation. Elytral intervals and scutellum with distinct isodiametric sculpticells, basal margin and lateral gutterof elytra without evident microsculpture. Ventral surface largely with sculpticells not apparent or with scarcely-visible isodiametric (proepisternum, abdominal ventrites 1-5 laterally) or slightly transverse sculpticells (abdominal ventrite 6). Head with punctures and wrinkles only on posterior part of clypeus and areas in and around frontal furrows, rest of dorsal surface smooth. Pronotum coarsely punctate in basal third, with some punctures at sides reaching medial third; disc and apical third smooth. Elytral intervals with a row of irregular punctures. Abdominal ventrites smooth to very finely wrinkled at sides.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="119133C9292B2056DE440DC1BADB15F4" bold="true" pageId="0" pageNumber="71">
<emphasis id="C3C37F4857CFBA8582BCE596DF4E149C" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
One-third or less as wide as pronotum (PW/HW= 1.38-1.42). Eyes moderately convex. Labrum subrectangular, as long as clypeus, with anterior margin slightly concave. Frontoclypeal suture distinct, impressed in middle, less distinct to obliterate at sides. Frontal furrows moderately distinct and impressed, subfoveolate. Paraorbital sulci straight, moderately deep, backward barely reaching posterior margin of eye, not reaching level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="24ED58D2E361614B54CA2DD5475687D6" bold="true" pageId="0" pageNumber="71">
<emphasis id="134E0C8E6C5F3B01202B7B8142FF235A" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about a quarter wider than long (PW/PL= 1.21-1.25), with widest point at second quarter. Anterior and posterior transverse impressions barely distinct. Sides sinuate, convex medially and anteriorly, concave posteriorly; bead of anterior border present throughout, rather fine; bead of posterior border present laterally and submedially, reduced to lacking at medial 1/10. Metepisternum as long as wide, MA/MI= 0.95-1.01.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="BFF1EF780584F9CBC7147B2CECCF3D3E" bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
. Elongate, about two thirds as long as elytra wide (EL/EW= 1.63-1.66), two times and two thirds as long as pronotum (EL/PL= 2.63-2.71), and one and a third as wide as pronotum (EW/PW= 1.32), with widest point at beginning of third quarter. Parascutellar striole and striae well-impressed, moderately punctate (less coarsely than
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
<taxonomicName id="90BF93516D52CC9C30D0B9E150C36D26" class="Insecta" kingdom="Animalia" lsidName="P. weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri">
<emphasis id="FB1C9F59F75A833606BB1478F5059A59" italics="true" pageId="0" pageNumber="71">P. weiratheri</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="CF46508148E494B9EF4F5D69D6592B8A" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="47E70B60D6A46881C4A1B7D4E7E470E9" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov.); parascutellar striole short, not joining stria 1; striae 1-6 reaching basal bead, 7 shortened a little before. Interval 3 with three discal setiferous punctures, as first lack on left eytron of paratype. Umbilicate setiferous series with 16 punctures on each side in holotype, with 17 punctures on left elytron and 15 ones on right elytron in paratype.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="BCC02D767C5FB2B379433C5D7466EAD0" bold="true" pageId="0" pageNumber="71">
<emphasis id="8302A3A3B6AC1B9B10A9069042BE8CBA" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur with 3-4 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="0712580E92C4D0A65C62C4548DE93099" bold="true" pageId="0" pageNumber="71">
<emphasis id="034A89070B8643139996772DC8CB83D6" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX suboval, with proximal margin slightly asymmetrical and pointed (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7D</figureCitation>
<figureCitation id="015B77E1FC2B308476FCE753AE4EE801" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7D</figureCitation>
). Median lobe of aedeagus in lateral view similar to that of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="F767EC7909F521AF66ACE02B2E0A115A" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="74CB3D68A6585A84B0D318D53306A75F" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
, but slightly narrower and with apex a little longer (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8F</figureCitation>
<figureCitation id="932AF61DFA372477394E46540B7D6D80" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8F</figureCitation>
); same in ventral view straight, about 3.2 - 3.3 times longer than wide (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10F</figureCitation>
<figureCitation id="1E0D336A788B5E65587EC93D86091882" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10F</figureCitation>
); apical lamella (dorsal view) less symmetrical and more elongate than in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="434AC585E3926C085223AE7E966A6C16" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="5F6B44262E518AEFDE33E208D81A4014" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8F</figureCitation>
<figureCitation id="7516C26B803329F256942F486D4D7053" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8F</figureCitation>
) with ventral sclerite distinctly broadened and rounded distally; same in ventral view (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10F</figureCitation>
<figureCitation id="89DC2E97C37164CCE3549C1F8EEB58D8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10F</figureCitation>
), with ventral sclerite somewhat more broadened distally than that of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
<taxonomicName id="21D2B3F93F6819DC6B8659CD15EE966C" class="Insecta" kingdom="Animalia" lsidName="P. vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis">
<emphasis id="D52F59444F2D99CD679360EC0C5CED58" italics="true" pageId="0" pageNumber="71">P. vanensis</emphasis>
</taxonomicName>
sp. nov. Right paramere on Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12E</figureCitation>
<figureCitation id="7130D25E16E158C62584D16E5A05BBCF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12E</figureCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="3EE2EF45C26983858D6ED479FE9A5A20" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13F</figureCitation>
<figureCitation id="A1F208C055104C0EAF30C52BE97EDAE2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13F</figureCitation>
). Apical gonocoxite with semi-pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 7.</emphasis>
<caption id="01960CD550D5CBB08AF8D16394854761" doi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71" start="Figure 7" startId="F7">
<paragraph id="EF751B9E2017DA769C4A522097B77404" pageId="0" pageNumber="71">
<emphasis id="B4DF020DD631E808682665558CAF718D" bold="true" pageId="0" pageNumber="71">Figure 7.</emphasis>
Urite, ventral view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
<emphasis id="D0E097387CCD1293E593D27A9B69CD09" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="28AA670C3F5890FE5B287EBABE6476FD" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) chatzakiae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chatzakiae" subGenus="Eremoderus">Platyderus (Eremoderus) chatzakiae</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<emphasis id="6A0E49F92C0CF9C91F880BFFC6333F66" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="9EA59BA435832A8CA27C6A72EC27E895" class="Insecta" kingdom="Animalia" lsidName="P. (E.) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="E.">P. (E.) weiratheri</taxonomicName>
<normalizedToken id="2885AFDD8DC03FC8CB71DAE91409F12F" originalValue="Mařan">Maran</normalizedToken>
, 1940, E: topotype male specimen;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
<emphasis id="F5CAE3653668D539690CF1DFCDA9BC9F" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="C16D0DB441DB6AB66B0B82587F48745A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
<emphasis id="3E848B4FD3C64BFDAF8D1ED1B3F62E2A" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="47218A9A9A389E05D52ABC42509F9EA0" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
<emphasis id="4FBD971FC0B4A9BDA058AC8EFB403310" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="A4690AB75FC599C369B67EE60DDC5407" class="Insecta" kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
<emphasis id="D32B3AE0CEAF39920A869EE425EA653B" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="D6E00F262AEFAF1C63CD9527AB14028B" class="Insecta" kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
<emphasis id="29F8F5A8246A7FA9E6A69012DA47DD71" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="9BB6AB1BDF749C0106EED4FF3778AF76" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, male specimen, Tochal, Tehran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="E.">P. (E.) afghanistanicus</taxonomicName>
<emphasis id="6AE1F93DD32C5F69C606D0C45CD6C785" bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName id="DF74E7F14BDCA8EE6B9BCFB44A97D0F3" class="Insecta" kingdom="Animalia" lsidName="P. (E.) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="E.">P. (E.) afghanistanicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
<emphasis id="732388EB0D73F23C849A6E6AEF81B261" bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName id="A10E373A66DA60DC2DCBE94642D19438" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="brunneus">P. (E.) brunneus brunneus</taxonomicName>
Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia;
<emphasis bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="F7C91250945C370FF1C4253A58D44382" bold="true" pageId="0" pageNumber="71">J.</emphasis>
<taxonomicName id="BFA755E35B95522CFDE747E4D5F1AE21" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel;
<emphasis bold="true" pageId="0" pageNumber="71">K.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
<emphasis id="58DBF705E56EAD29F38BE9260BE7F88C" bold="true" pageId="0" pageNumber="71">K.</emphasis>
<taxonomicName id="8AD37571727BA9FA2DCB3F43A20F2450" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">L.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="AA75794B276400666B0449F4D6B097A0" bold="true" pageId="0" pageNumber="71">L.</emphasis>
<taxonomicName id="3B2D9C914CCBF8A3FEB05FA8E3F646B4" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">M.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="558ED292F7EFB486B74E2B066B329B3E" bold="true" pageId="0" pageNumber="71">M.</emphasis>
<taxonomicName id="D959715D7C768579C9267909CB55EA00" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), male specimen, Naẖal Prat, Judea and Samaria Area, Israel;
<emphasis bold="true" pageId="0" pageNumber="71">N.</emphasis>
<taxonomicName authority="" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">P. (Platyderus)</taxonomicName>
<emphasis id="0C40E873ECA82DF9B5B038EB24FD196A" bold="true" pageId="0" pageNumber="71">N.</emphasis>
<taxonomicName id="FE3DFC7B6807B712C643822A4E367AD0" authority="" class="Insecta" kingdom="Animalia" lsidName="P. (Platyderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Platyderus">P. (Platyderus)</taxonomicName>
sp., male specimen, Boz
<normalizedToken originalValue="Dağlar">Daglar</normalizedToken>
<normalizedToken id="D38782C2D6B7032C174764DB02DF8EA5" originalValue="Dağlar">Daglar</normalizedToken>
Mtn., Turkey;
<emphasis bold="true" pageId="0" pageNumber="71">O.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (P.) reticulatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="reticulatus" subGenus="P.">P. (P.) reticulatus</taxonomicName>
<emphasis id="DB8195563C47531A59934911F2EE7941" bold="true" pageId="0" pageNumber="71">O.</emphasis>
<taxonomicName id="30432FE8A2FA1AACF67555C4D5A048D8" class="Insecta" kingdom="Animalia" lsidName="P. (P.) reticulatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="reticulatus" subGenus="P.">P. (P.) reticulatus</taxonomicName>
(Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="9EFB637B3AE2A43B946086A2C14FE698" pageId="0" pageNumber="71" type="habitat">
<paragraph id="E4C6C0B1ACDCE8E6DDE6448425DD7F95" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="791628F9381797641CAB1152EBC600B0" pageId="0" pageNumber="71">
Slopes with pastures and stone fields covered with snow patches at the time of collecting at an altitude of 1700-1900 m. Several trees in poor condition and a few flowering wild representatives of
<taxonomicName genus="Hyacintus" lsidName="Hyacintus" pageId="0" pageNumber="71" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Hyacintus</emphasis>
<taxonomicName id="01DBE57E537687248EC4495F27F4E825" genus="Hyacintus" lsidName="Hyacintus" pageId="0" pageNumber="71" rank="genus">
<emphasis id="3BA79C837058F1C9AA70F37C78C7B20A" italics="true" pageId="0" pageNumber="71">Hyacintus</emphasis>
</taxonomicName>
spp. were observed around the place of collecting (V. Vrabec, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="EBF0F10CDC5D380DEB0C48D16D8E160E" pageId="0" pageNumber="71" type="distribution">
<paragraph id="4E10157DEA5D121A7CF3B6A4127CBFAF" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="163A8035BCB826B7E7702C0B223F9A54" pageId="0" pageNumber="71">
Nemrut
<normalizedToken originalValue="Daği">Dagi</normalizedToken>
<normalizedToken id="8C2B8D54BAF457FFEBB9FB418CBE03A9" originalValue="Daği">Dagi</normalizedToken>
(Turkey, Southeastern Anatolia, Province of
<normalizedToken originalValue="Adıyaman">Adiyaman</normalizedToken>
<normalizedToken id="2D1BC237BA4A947FDE1399BFD77C379B" originalValue="Adıyaman">Adiyaman</normalizedToken>
; Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
<figureCitation id="273AC65FBE4489EBBEBBE54C79CDC22B" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of species from subgenus Platyderus Eremoderus in the western Aegean Sea Region and Turkey: Platyderus chatzakiae sp. nov. (red circle), P. weiratheri (blue circles), P. vanensis sp. nov. (brown circles), and P. vrabeci sp. nov. (yellow circle)." figureDoi="10.3897/dez.69.83840.figure16" httpUri="https://binary.pensoft.net/fig/695653" pageId="0" pageNumber="71">16</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="FC10B70B8462A79F5D0DFA2258BADDA8" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
<emphasis id="C5CF8286280EC215031246F89A6680E8" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="B5862515E3DC6E85C413A0EC7F960D41" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis id="DA0897134D3D03B19FA1947C3FC60659" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
</taxonomicName>
&quot; species group
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Notes</emphasis>
<paragraph id="9AF79DEFB32DDE6A4750378C1AA39693" pageId="0" pageNumber="71">
<emphasis id="CD8334D2653F8361BDD6FCFA5FAAF683" bold="true" pageId="0" pageNumber="71">Notes</emphasis>
. The species of the &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
<taxonomicName id="CE157B21780D7C00ADB631CDDDE6C35D" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">
<emphasis id="453BF795876FFFDD589E13CD0CA36C5A" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus lassallei</emphasis>
</taxonomicName>
&quot; and the &quot;
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<taxonomicName id="81AD36DFA2E6F3251448A83865A77FCC" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="D3C321DC0306C3843855C818426E9354" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
&quot; group share a synapomorphy, the distal end of the ventral sclerite of the median lobe is curved to the left (ventral view).
</paragraph>

184
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@ -1,146 +1,146 @@
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<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Fig. 2A</figureCitation>
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<figureCitation id="7A92A7FCE305AD1DEFA95A56C5D78982" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Fig. 2A</figureCitation>
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TME:
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. TGE: 0.
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<paragraph pageId="0" pageNumber="71">Diagnostic features.</paragraph>
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It is distinct from individuals of
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sp. nov. by flatter elytral intervals and less coarsely punctate elytral striae. Measurements and ratios shown on Table
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<paragraph pageId="0" pageNumber="71">Habitat and bionomic notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
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<paragraph id="E37668229D6B7D8AFFDDA9193AD8690A" pageId="0" pageNumber="71">Habitat and bionomic notes.</paragraph>
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The specimen was collected by PM just before the pass Karabet Pass (eastern slope), on the road coming from Yukari-Narlica
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village towards the pass. There is a large cirque on the left with many snowfields and streams descending from them; it was found in gravel between the rivulets, at an altitude of about 2800 m.
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<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
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<paragraph id="703395B3D6334E150313720EEEEB6D35" pageId="0" pageNumber="71">Distribution.</paragraph>
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Karabet Pass [= Karabet
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<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">The single female remains unidentified due to the lack of a male specimen. In addition, the characters of the elytral intervals and striae are not sufficiently diagnostic to propose a new name for this form.</paragraph>
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<paragraph id="6F8BBB850AC8DD66CD8703579B3EB205" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="D7CF7ACBD0030952A525E76F7D4F98B4" pageId="0" pageNumber="71">The single female remains unidentified due to the lack of a male specimen. In addition, the characters of the elytral intervals and striae are not sufficiently diagnostic to propose a new name for this form.</paragraph>
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16.1.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4A</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9E</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11D</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15C</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
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<figureCitation id="FA374366C759A8AD5E1022979E4A0A8B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4A</figureCitation>
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<figureCitation id="F422D22EF113D1B71D99E87AEF506F1B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11D</figureCitation>
<figureCitation id="47FFC01F7CF5EF6D1CA42B24FDDEFD26" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15C</figureCitation>
<figureCitation id="9E1CADFC61131F0CB8457236B236B89E" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">Platyderus insignitus</taxonomicName>
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<taxonomicName id="44A39CEF500EA340A62B212B4F4049D8" authorityName="Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">Platyderus insignitus</taxonomicName>
Bedel, 1902: 211 (type locality:
<normalizedToken originalValue="“Mogador”">&quot;Mogador&quot;</normalizedToken>
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[= Essaouira], Morocco).
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<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus insignitus</emphasis>
<subSubSection id="C8235DF0AF9DB582A5ED02FA3BBB81F4" pageId="0" pageNumber="71" type="references">
<paragraph id="444C2B6C9BC8353C1ABCDF98A4225C26" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="22CB62C8663E4FB7C94B6C4B892A065F" pageId="0" pageNumber="71">
<taxonomicName id="760624792F6B219EE9A3B160B0299A76" authorityName="Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis id="36FDB5EAC896A0FD7D0098C5B9690557" italics="true" pageId="0" pageNumber="71">Platyderus insignitus</emphasis>
</taxonomicName>
:
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
<bibRefCitation id="A58F072650430B33CDE056376AF37648" author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
: 214;
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
<bibRefCitation id="AD5B4B404DB416C8869B242117E02ED7" author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
: 769;
<bibRefCitation author="Antoine, M" journalOrPublisher=", Zoologie)" pageId="0" pageNumber="71" pagination="179 - 314" refId="B5" refString="Antoine, M, 1957. Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N.S., Zoologie) 3: 179 - 314" title="Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N. S." volume="3" year="1957">Antoine 1957</bibRefCitation>
<bibRefCitation id="884C232980AA51D62F00AD0C0F2CFC4B" author="Antoine, M" journalOrPublisher=", Zoologie)" pageId="0" pageNumber="71" pagination="179 - 314" refId="B5" refString="Antoine, M, 1957. Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N.S., Zoologie) 3: 179 - 314" title="Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N. S." volume="3" year="1957">Antoine 1957</bibRefCitation>
: 235-236;
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1996.17271" author="Jeanne, C" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="397 - 412" refId="B25" refString="Jeanne, C, 1996. Le genre Platyderus Stephens: I. - Especes nouvelles de la peninsule Iberique (Coleoptera, Pterostichidae). Bulletin de la Societe Entomologique de France 101 (4): 397 - 412, DOI: https://doi.org/10.3406/bsef.1996.17271" title="Le genre Platyderus Stephens: I. - Especes nouvelles de la peninsule Iberique (Coleoptera, Pterostichidae)." url="https://doi.org/10.3406/bsef.1996.17271" volume="101" year="1996">Jeanne 1996</bibRefCitation>
<bibRefCitation id="341FCD3C81F0CB748611C8963A7FAE23" DOI="https://doi.org/10.3406/bsef.1996.17271" author="Jeanne, C" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="397 - 412" refId="B25" refString="Jeanne, C, 1996. Le genre Platyderus Stephens: I. - Especes nouvelles de la peninsule Iberique (Coleoptera, Pterostichidae). Bulletin de la Societe Entomologique de France 101 (4): 397 - 412, DOI: https://doi.org/10.3406/bsef.1996.17271" title="Le genre Platyderus Stephens: I. - Especes nouvelles de la peninsule Iberique (Coleoptera, Pterostichidae)." url="https://doi.org/10.3406/bsef.1996.17271" volume="101" year="1996">Jeanne 1996</bibRefCitation>
: 398;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="4239F01415C08E194343C8BE6377F645" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="CA2D6F084D166C94E829731F41C687F3" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 521;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="3FE424F4321113322F4FC2DC0479C553" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 395;
<bibRefCitation author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="101 - 102" refId="B36" refString="Machard, P, 2017. Un Platyderus nouveau du Maroc (Coleoptera, Carabidae). Coleopterist 20 (2): 101 - 102" title="Un Platyderus nouveau du Maroc (Coleoptera, Carabidae)." volume="20" year="2017">Machard 2017</bibRefCitation>
<bibRefCitation id="84D46038A46D9E5B2D65CB640217CED9" author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="101 - 102" refId="B36" refString="Machard, P, 2017. Un Platyderus nouveau du Maroc (Coleoptera, Carabidae). Coleopterist 20 (2): 101 - 102" title="Un Platyderus nouveau du Maroc (Coleoptera, Carabidae)." volume="20" year="2017">Machard 2017</bibRefCitation>
: 102;
<bibRefCitation author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="183 - 203" refId="B37" refString="Machard, P, 2019. Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae). Coleopterist 21 (3): 183 - 203" title="Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae)." volume="21" year="2019">Machard 2019</bibRefCitation>
<bibRefCitation id="410FA727E25E1E160310444846801244" author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="183 - 203" refId="B37" refString="Machard, P, 2019. Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae). Coleopterist 21 (3): 183 - 203" title="Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae)." volume="21" year="2019">Machard 2019</bibRefCitation>
: 199.
<taxonomicName authority="insignitus" authorityName="insignitus Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus insignitus insignitus</emphasis>
<taxonomicName id="82F6366681A971B7291245C96DE5301A" authority="insignitus" authorityName="insignitus Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="insignitus">
<emphasis id="AE234054B95E1C4BAEFE3EC4E8B1C900" italics="true" pageId="0" pageNumber="71">Platyderus insignitus insignitus</emphasis>
</taxonomicName>
:
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="25B588337036C798CAA27E1EA2C1E0C6" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 756.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">Five syntypes should be present in MNHN, according to original description. Not examined.</paragraph>
<subSubSection id="36C42F6F988384B4F5A370CC09A10D2D" pageId="0" pageNumber="71" type="type material">
<paragraph id="D24A8125C3F6F21BA5389D7C7CCDD011" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="5669276EDB5D504E0C17564BBA101154" pageId="0" pageNumber="71">Five syntypes should be present in MNHN, according to original description. Not examined.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Morocco</emphasis>
<subSubSection id="4A3B95885733376AEA245671D0476ACC" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="3A5BBB4B9D4F7C29C629A700564F1176" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="8F7E32E653EDE36255E7A414602BA15E" pageId="0" pageNumber="71">
<emphasis id="980DC747CC9D54DCA32B3611B39286FF" bold="true" pageId="0" pageNumber="71">Morocco</emphasis>
:
<materialsCitation collectingDate="2000-03-02" collectingDateMax="2011-05-06" collectingDateMin="2000-03-02" collectorName="El Jadida, M. Egger', S Essaouira, J. Kalab, I. Puchner, Plage Aglou, NW Tiznit" country="MOROCCO" county="Sous-Massa Region" elevation="60" latitude="29.75" location="Casablanca-Settat Region" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="region Doukkala-Abda">
<location LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:A0439F3F0927D48D4F99B67FDD863A2C" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Casablanca-Settat Region" stateProvince="region Doukkala-Abda">Casablanca-Settat Region</location>
<materialsCitation id="DACF73AEB66CD3B1FE7F49D8C8E136B9" collectingDate="2000-03-02" collectingDateMax="2011-05-06" collectingDateMin="2000-03-02" collectorName="El Jadida, M. Egger', S Essaouira, J. Kalab, I. Puchner, Plage Aglou, NW Tiznit" country="MOROCCO" county="Sous-Massa Region" elevation="60" latitude="29.75" location="Casablanca-Settat Region" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="region Doukkala-Abda">
<location id="A0439F3F0927D48D4F99B67FDD863A2C" LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:A0439F3F0927D48D4F99B67FDD863A2C" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Casablanca-Settat Region" stateProvince="region Doukkala-Abda">Casablanca-Settat Region</location>
:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="77202C49B79AB6A290C9A4ADC8123539" type="female">1♀</specimenCount>
, '
<location LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:6C817577EF72619BF69399A9DCED9836" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Marokko" stateProvince="region Doukkala-Abda">Marokko</location>
<location id="6C817577EF72619BF69399A9DCED9836" LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:6C817577EF72619BF69399A9DCED9836" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Marokko" stateProvince="region Doukkala-Abda">Marokko</location>
(
<collectingRegion country="Morocco" name="Doukkala-Abda">region Doukkala-Abda</collectingRegion>
<collectingRegion id="30D989C20DA62476D534A9334CA39A28" country="Morocco" name="Doukkala-Abda">region Doukkala-Abda</collectingRegion>
) /
<location LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:D9055B9A0C197097591B329D2D039324" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Bir Jdid" stateProvince="region Doukkala-Abda">Bir Jdid</location>
<location id="D9055B9A0C197097591B329D2D039324" LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:D9055B9A0C197097591B329D2D039324" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Bir Jdid" stateProvince="region Doukkala-Abda">Bir Jdid</location>
between
<collectorName>
<location LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:F3F4190814B67A0710D47B207022EE13" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="El Jadida" stateProvince="region Doukkala-Abda">El Jadida</location>
<collectorName id="FD42105084A332B5F515F6DABB6106FD">
<location id="F3F4190814B67A0710D47B207022EE13" LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:F3F4190814B67A0710D47B207022EE13" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="El Jadida" stateProvince="region Doukkala-Abda">El Jadida</location>
</collectorName>
/ and
<location LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:007DCE000FF78A8CAD6796F5392413F7" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Casablanca" stateProvince="region Doukkala-Abda">Casablanca</location>
<location id="007DCE000FF78A8CAD6796F5392413F7" LSID="urn:lsid:plazi:treatment:C3A0284CB7B0560D9422C68138948997:007DCE000FF78A8CAD6796F5392413F7" country="MOROCCO" county="Sous-Massa Region" latitude="29.75" longLatPrecision="1225" longitude="-9.9" municipality="Marrakesh-Safi Region" name="Casablanca" stateProvince="region Doukkala-Abda">Casablanca</location>
/
<collectingDate value="2003-02-04">4.2.2003</collectingDate>
<collectingDate id="F8C66E0676247FA7C9D4C69C50CE4667" value="2003-02-04">4.2.2003</collectingDate>
, leg.
<collectorName>
<collectorName id="29B8FA9F2B370D088C5918B370C3F0A0">
M.
<normalizedToken originalValue="Egger’">Egger'</normalizedToken>
<normalizedToken id="88D9B50D0E4FB92602BD27F2F4C468EA" originalValue="Egger’">Egger'</normalizedToken>
</collectorName>
(cWR).
<collectingMunicipality>Marrakesh-Safi Region</collectingMunicipality>
<collectingMunicipality id="117BF3E447DEC9307D53026B8FE26726">Marrakesh-Safi Region</collectingMunicipality>
:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="F53A55182914601BC10640F9276EEBF1" type="female">1♀</specimenCount>
, 'SW-Morocco,
<geoCoordinate degrees="31.28" direction="north" orientation="latitude" precision="555" value="31.28">31.28 N</geoCoordinate>
<geoCoordinate id="44E30444F5C07D3AEE7A253160814F43" degrees="31.28" direction="north" orientation="latitude" precision="555" value="31.28">31.28 N</geoCoordinate>
/
<geoCoordinate degrees="09.46" direction="west" orientation="longitude" precision="555" value="-9.46">09.46W</geoCoordinate>
<geoCoordinate id="ADBB9F36B4426671149793EC1B6DE165" degrees="09.46" direction="west" orientation="longitude" precision="555" value="-9.46">09.46W</geoCoordinate>
, DIABAT (
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="km" value="4.0">4 km</quantity>
<quantity id="57BFD0C04107FEEB653ED65BAB3D13D7" metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="km" value="4.0">4 km</quantity>
/
<collectorName>S Essaouira</collectorName>
<collectorName id="A5F55F92BCC992BBCC3C7A4414612777">S Essaouira</collectorName>
),
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</elevation>
<quantity id="E142CCA9BC4527B2A942491E6563E235" metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">
<elevation id="1A869F6A64F7B9099589112444AA254D" metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</elevation>
</quantity>
//
<collectorName>
<collectorName id="E2C60BC236EFE17F5AE2932E166FFBC7">
J.
<normalizedToken originalValue="Kaláb">Kalab</normalizedToken>
<normalizedToken id="F3E0737ACB51FE3106A5F6AEDF90AAC6" originalValue="Kaláb">Kalab</normalizedToken>
</collectorName>
leg.,
<collectingDate value="2003-11-17">17.XI.2003</collectingDate>
<collectingDate id="50F219275A8F5F74B90B2CB90A8202E9" value="2003-11-17">17.XI.2003</collectingDate>
/ bushes, sandy dunes, /
<normalizedToken originalValue="steppe’">steppe'</normalizedToken>
<normalizedToken id="271E3A0A7040FECDB8CEA7F9DEF7C019" originalValue="steppe’">steppe'</normalizedToken>
(cASL).
<collectingCounty>Sous-Massa Region</collectingCounty>
<collectingCounty id="5010A1C5335AA63B570B4B5F55EEED89">Sous-Massa Region</collectingCounty>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="D3C7107F62A1BF1C9D678094AF2479B0" type="male">1♂</specimenCount>
, 'MAROKKO, s. w. TIZNIT / umg. TIZNIT,
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</elevation>
<quantity id="6046AA55AE8905A2C3BCE53C41834CB3" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<elevation id="2E7860E668583D95707793903E5117F0" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</elevation>
</quantity>
/ Strand-AGLOU /
<collectorName>I. Puchner</collectorName>
<collectorName id="1ECC106FC62EF282272749CCD37D5CC2">I. Puchner</collectorName>
,
<collectingDate value="2000-03-02">2.III.2000</collectingDate>
<collectingDate id="CAF0912E07B22A9F5A8C9C5C3F99EC72" value="2000-03-02">2.III.2000</collectingDate>
' (cDOS);
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="CA5E223929B8DADA9080A8A5D1D480B4" type="female">1♀</specimenCount>
, 'SW-Morocco,
<geoCoordinate degrees="29.48" direction="north" orientation="latitude" precision="555" value="29.48">29.48N</geoCoordinate>
<geoCoordinate id="D8F420D6C0CB824C87B94400C7511D53" degrees="29.48" direction="north" orientation="latitude" precision="555" value="29.48">29.48N</geoCoordinate>
/
<geoCoordinate degrees="09.50" direction="west" orientation="longitude" precision="555" value="-9.5">09.50W</geoCoordinate>
<geoCoordinate id="C5B74C40A06DCA1155E8B7AFB2D1F3E2" degrees="09.50" direction="west" orientation="longitude" precision="555" value="-9.5">09.50W</geoCoordinate>
/ AGLOU PLAGE,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</elevation>
<quantity id="1BCEF39D6F72C5053EF9C8AD09498EC1" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">
<elevation id="46178105B605F24F9D456D036C21A03B" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</elevation>
</quantity>
, /
<collectorName>
<collectorName id="B83B3C91FE53A4506424E6E007375056">
J.
<normalizedToken originalValue="Kaláb">Kalab</normalizedToken>
<normalizedToken id="C0D2B561C8266F94CB1EA5D2D06DD603" originalValue="Kaláb">Kalab</normalizedToken>
</collectorName>
leg.,
<collectingDate value="2004-11-26" valueMax="2004-11-28" valueMin="2004-11-26">26-28.xi.2004</collectingDate>
<collectingDate id="86F7400F499A97B9331ED9C69451C740" value="2004-11-26" valueMax="2004-11-28" valueMin="2004-11-26">26-28.xi.2004</collectingDate>
/ sandy/grassy places, /
<normalizedToken originalValue="half-desert’">half-desert'</normalizedToken>
<normalizedToken id="E490423EA0A1AB24C1FD46B47FAF4421" originalValue="half-desert’">half-desert'</normalizedToken>
(cWR);
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="B279876F2D7E5D6541EDFD5EDA617950" type="female">1♀</specimenCount>
, '
<collectingCountry name="Morocco">MOROCCO</collectingCountry>
<collectingCountry id="07F9F94A6D68132BA5E22AF4C40DDC4B" name="Morocco">MOROCCO</collectingCountry>
,
<collectorName>Plage Aglou</collectorName>
<collectorName id="854C585F51932AF6A86C63F77C629FE5">Plage Aglou</collectorName>
/
<collectorName>NW Tiznit</collectorName>
<collectorName id="30E5E441F68524C4BAD719EB2AC62236">NW Tiznit</collectorName>
/
<geoCoordinate degrees="29" direction="north" minutes="45" orientation="latitude" precision="925" value="29.75">29°45'N</geoCoordinate>
<geoCoordinate id="7A39552EBB4ECD1C93240D5971EBD45A" degrees="29" direction="north" minutes="45" orientation="latitude" precision="925" value="29.75">29°45'N</geoCoordinate>
,
<geoCoordinate degrees="9" direction="west" minutes="54" orientation="longitude" precision="925" value="-9.9">9°54'W</geoCoordinate>
<geoCoordinate id="9FA0B8B1A6B597AC7BDBB435CE86E1C7" degrees="9" direction="west" minutes="54" orientation="longitude" precision="925" value="-9.9">9°54'W</geoCoordinate>
/
<quantity metricMagnitude="0" metricUnit="m" metricValue="6.0" unit="m" value="6.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="6.0" unit="m" value="6.0">6 m</elevation>
<quantity id="C5B092E50EBFA3643EE521F8A4619DB5" metricMagnitude="0" metricUnit="m" metricValue="6.0" unit="m" value="6.0">
<elevation id="482AD284C081E5840D27FCFC6ACA38E1" metricMagnitude="0" metricUnit="m" metricValue="6.0" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<collectingDate value="2011-05-06">6.V.2011</collectingDate>
<collectingDate id="AAAA7AA578C9A766FB4ABE71D92073EA" value="2011-05-06">6.V.2011</collectingDate>
/ lgt.
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<normalizedToken originalValue="Kabátek’">Kabatek'</normalizedToken>
<taxonomicName id="6AF0B51D406730DD73D428EC3C078AA5" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<normalizedToken id="75F617A4F1C120796D9C47CBD914D929" originalValue="Kabátek’">Kabatek'</normalizedToken>
(cWR)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="00D24042A0038959B89337C9AF9E23F3" pageId="0" pageNumber="71">
TME:
<specimenCount type="generic">5 specimens</specimenCount>
<specimenCount id="F42F71AA25C18490552CD8A122C743DD" type="generic">5 specimens</specimenCount>
. TGE:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="EC8A8603830AE50B3BAAC7642C0EB729" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="64ECFB9914CB00E7EF20C5DD0319BED4" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="05BD69993D4D589E161C6BBBE39A9428" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="D91C6FA5C236E255564F6E43C326C33E" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="BEC6E76D146E06CB340746F2285DD227" pageId="0" pageNumber="71">
See
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="ECEB1C887F56CE16A30FDD0D4D5993FF" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus subsp. presaharensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presaharensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus presaharensis</emphasis>
<taxonomicName id="76670451E8463E07BBBB550AC4DBC858" class="Insecta" kingdom="Animalia" lsidName="P. insignitus subsp. presaharensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presaharensis">
<emphasis id="CED66EF29AD2625A54634F9EDE44AAE1" italics="true" pageId="0" pageNumber="71">P. insignitus presaharensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">Nothing is known about the bionomics of this species.</paragraph>
<subSubSection id="DDDF115774EAD87B396B346AD841F3C9" pageId="0" pageNumber="71" type="habitat">
<paragraph id="5C7D4BEAC318078C17CA2714A0B16A14" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="6D338293A0A836772315D137CCC581DD" pageId="0" pageNumber="71">Nothing is known about the bionomics of this species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="00C7343E4DB6EFF1207948E3D872F5B0" pageId="0" pageNumber="71" type="distribution">
<paragraph id="D7F7B709ABF9A8A8D61BAE37A41FD909" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="13796F78F1CED42E8AF85BECF6449E8C" pageId="0" pageNumber="71">
According to the available data,
<taxonomicName authority="insignitus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus insignitus</emphasis>
<taxonomicName id="44B4E2E8E89B01352E4ED1FD06BB7CBE" authority="insignitus" class="Insecta" kingdom="Animalia" lsidName="P. insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="insignitus">
<emphasis id="A048BD6C752AACECFF1C20A2EBA4400A" italics="true" pageId="0" pageNumber="71">P. insignitus insignitus</emphasis>
</taxonomicName>
is endemic to Morocco (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
<figureCitation id="F3A75AE19B0C222E3EFFD622701B1340" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
).
<bibRefCitation author="Antoine, M" journalOrPublisher=", Zoologie)" pageId="0" pageNumber="71" pagination="179 - 314" refId="B5" refString="Antoine, M, 1957. Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N.S., Zoologie) 3: 179 - 314" title="Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N. S." volume="3" year="1957">Antoine (1957</bibRefCitation>
<bibRefCitation id="2B1DF350839F8A5C7003ECAFE3A71074" author="Antoine, M" journalOrPublisher=", Zoologie)" pageId="0" pageNumber="71" pagination="179 - 314" refId="B5" refString="Antoine, M, 1957. Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N.S., Zoologie) 3: 179 - 314" title="Coleopteres carabiques du Maroc (deuxieme partie). Memoires de la Societe des Sciences Naturelles et Physiques du Maroc (N. S." volume="3" year="1957">Antoine (1957</bibRefCitation>
: 236) stated that it is a sublittoral species living in the area between Oum er Rbia River to the north and the Anti-Atlas Mountain Range to the south.
<bibRefCitation author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="101 - 102" refId="B36" refString="Machard, P, 2017. Un Platyderus nouveau du Maroc (Coleoptera, Carabidae). Coleopterist 20 (2): 101 - 102" title="Un Platyderus nouveau du Maroc (Coleoptera, Carabidae)." volume="20" year="2017">Machard (2017</bibRefCitation>
<bibRefCitation id="3E6B30FEFEACE82738065B0A2AB97E96" author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="101 - 102" refId="B36" refString="Machard, P, 2017. Un Platyderus nouveau du Maroc (Coleoptera, Carabidae). Coleopterist 20 (2): 101 - 102" title="Un Platyderus nouveau du Maroc (Coleoptera, Carabidae)." volume="20" year="2017">Machard (2017</bibRefCitation>
,
<bibRefCitation author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="183 - 203" refId="B37" refString="Machard, P, 2019. Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae). Coleopterist 21 (3): 183 - 203" title="Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae)." volume="21" year="2019">2019</bibRefCitation>
<bibRefCitation id="C706FD689C53ED2CE943FCE7A9C7411D" author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="183 - 203" refId="B37" refString="Machard, P, 2019. Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae). Coleopterist 21 (3): 183 - 203" title="Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae)." volume="21" year="2019">2019</bibRefCitation>
) noted that it inhabits the &quot;Littoral atlantique de Casablanca
<normalizedToken originalValue="à">a</normalizedToken>
<normalizedToken id="7E69C39C4AB41CE6EF7011B8FA70165D" originalValue="à">a</normalizedToken>
Agadir&quot;. The new data from the environs of Tiznit show that the species occurs south of the city of Agadir as well.
</paragraph>
</subSubSection>

410
data/CB/C3/2D/CBC32D678CEC5274987BE44552D6AA13.xml
View file

@ -1,368 +1,368 @@
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10.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2D</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7F</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8H</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10H</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12G</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14B</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
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<figureCitation id="FA1E0213E1B5890BED688E8D19A18E28" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">Figs 2D</figureCitation>
<figureCitation id="EED1E249F2870DAA85FD7DD36D401EE3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7F</figureCitation>
<figureCitation id="BBFF9DDA59D9AE5C404B2D2DD082F3DB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8H</figureCitation>
<figureCitation id="EA6D56139D0B6125E875511BE44AD8D1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10H</figureCitation>
<figureCitation id="E9535E63593548CE29BBDD928E32E69E" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12G</figureCitation>
<figureCitation id="670EBCD31D400223DED8804A0A5A1138" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14B</figureCitation>
<figureCitation id="00E0BDCB5E5880C6BDC06474307B6959" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
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<taxonomicName authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">Platyderus ledouxi</taxonomicName>
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<paragraph id="5516C6063A69847B8CFA66A72D037998" pageId="0" pageNumber="71">
<taxonomicName id="04FC7423558E329F16F6E593557B3E35" authorityName="Morvan" authorityYear="1974" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">Platyderus ledouxi</taxonomicName>
Morvan, 1974: 149 (type locality:
<normalizedToken originalValue="“l’Elburz">&quot;l'Elburz</normalizedToken>
<normalizedToken id="C31F5B304D46C5186007D24CCEE055DE" originalValue="“l’Elburz">&quot;l'Elburz</normalizedToken>
, province du Mazandaran, massif du Soleyman, Roudbarak, alt: 1800 m, en
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de
<taxonomicName class="Magnoliopsida" family="Carabidae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fagus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
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&quot;).
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<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">The Takht-e Soleyman Massif, a subrange of the Central Alborz Range, is located in the Province of Mazandaran (Chalus County, Keraldasht District). For the type locality Morvan indicated as biotope a beech forest in the vicinity of Rudbarak village, situated at ca. 1800 m altitude. This site lies in the upper valley of the Sardabrud River, northeast of the Takht-e Suleyman Massif.</paragraph>
<subSubSection id="6F35865F8C9C54A3A629621F68208035" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="A014AB9A4FC9940556712F8B55C0553A" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="4EDE51ABF118BA7131361C5FD0FD1AA2" pageId="0" pageNumber="71">The Takht-e Soleyman Massif, a subrange of the Central Alborz Range, is located in the Province of Mazandaran (Chalus County, Keraldasht District). For the type locality Morvan indicated as biotope a beech forest in the vicinity of Rudbarak village, situated at ca. 1800 m altitude. This site lies in the upper valley of the Sardabrud River, northeast of the Takht-e Suleyman Massif.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
<subSubSection id="1946D94F59A95F5B99CF3152A73EF995" pageId="0" pageNumber="71" type="references">
<paragraph id="08AB85CA9B5F6AF460EC00D4BE39153F" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="6D3E30A999FEDCFC33EA5DA863C8D522" pageId="0" pageNumber="71">
<taxonomicName id="3733893AA8B9C673113741A2B7169A99" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="CB93E5B9E5F05673EDB2039E90F7701D" italics="true" pageId="0" pageNumber="71">Platyderus davatchii</emphasis>
</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1969.21081" author="Morvan, P" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="192 - 198" refId="B40" refString="Morvan, P, 1970. Contribution a la connaissance des coleopteres carabiques de l'Iran. Bulletin de la Societe Entomologique de France 74 (1969): 192 - 198, DOI: https://doi.org/10.3406/bsef.1969.21081" title="Contribution a la connaissance des coleopteres carabiques de l'Iran." url="https://doi.org/10.3406/bsef.1969.21081" volume="74" year="1970">Morvan 1970</bibRefCitation>
<bibRefCitation id="CA2DA0176E4AC157FB481F4DF92792B6" DOI="https://doi.org/10.3406/bsef.1969.21081" author="Morvan, P" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="192 - 198" refId="B40" refString="Morvan, P, 1970. Contribution a la connaissance des coleopteres carabiques de l'Iran. Bulletin de la Societe Entomologique de France 74 (1969): 192 - 198, DOI: https://doi.org/10.3406/bsef.1969.21081" title="Contribution a la connaissance des coleopteres carabiques de l'Iran." url="https://doi.org/10.3406/bsef.1969.21081" volume="74" year="1970">Morvan 1970</bibRefCitation>
:
<bibRefCitation author="Lohaj, R" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" pagination="7 - 14" refId="B32" refString="Lohaj, R, Mlejnek, R, 2007. Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria. Acta Societatis Zoologicae Bohemicae 71: 7 - 14" title="Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria." volume="71" year="2007">Lohaj and Mlejnek 2007</bibRefCitation>
<bibRefCitation id="7C6B4D70C5763B43A6F18E20C635D3CA" author="Lohaj, R" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" pagination="7 - 14" refId="B32" refString="Lohaj, R, Mlejnek, R, 2007. Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria. Acta Societatis Zoologicae Bohemicae 71: 7 - 14" title="Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabiade) from Turkey and Syria." volume="71" year="2007">Lohaj and Mlejnek 2007</bibRefCitation>
: 12.
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="631491FA31F70A95194C6765BEB599FE" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="DEB610CCDD426AF85A4BD4A26F1FC595" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
:
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="7631809F2A28F1E15BD24C2086E52A25" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="2B1A2401E0A9A4B6796716179A07FABE" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="4A19BB90854676844D4A4544A88C22D7" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 396;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4024.1.1" author="Azadbakhsh, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="1 - 108" refId="B9" refString="Azadbakhsh, S, Nozari, J, 2015. Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae). Zootaxa 4024 (1): 1 - 108, DOI: https://doi.org/10.11646/zootaxa.4024.1.1" title="Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae)." url="https://doi.org/10.11646/zootaxa.4024.1.1" volume="4024" year="2015">Azadbakhsh and Nozari 2015</bibRefCitation>
<bibRefCitation id="55F1B5A52AFBCC314CA1860FE8AA7738" DOI="https://doi.org/10.11646/zootaxa.4024.1.1" author="Azadbakhsh, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" pagination="1 - 108" refId="B9" refString="Azadbakhsh, S, Nozari, J, 2015. Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae). Zootaxa 4024 (1): 1 - 108, DOI: https://doi.org/10.11646/zootaxa.4024.1.1" title="Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae)." url="https://doi.org/10.11646/zootaxa.4024.1.1" volume="4024" year="2015">Azadbakhsh and Nozari 2015</bibRefCitation>
: 84;
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="14935552466CA39E8441FD223CC2DA90" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 758.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="D6CD7F96B21D33CD88E09D7B531FFDB3" pageId="0" pageNumber="71" type="type material">
<paragraph id="F01ACDE34AF4C20EA07C7FD5E99CD4A4" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="CD048C0239083073C1020E6C3FD306F2" pageId="0" pageNumber="71">
<emphasis id="ABC65377B011500CA994687BADC94BEE" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂ in MNHN, not examined; paratype ♀ in private collection Pierre Morvan (Carentoir, France), not examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="N Iran" country="Iran" location="Mazandaran Province" specimenCount="1" specimenCount-male="1" stateProvince="Mazandaran Province">
<emphasis bold="true" pageId="0" pageNumber="71">
<collectingCountry name="Iran">Iran</collectingCountry>
<subSubSection id="B154ABD4655136803D885C8EB6EAA7DC" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="0B6016C8C0E376B6AA3C9A7244BF7A1A" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="B83F672FA4544D83C9A76515BC608EE5" pageId="0" pageNumber="71">
<materialsCitation id="A0185A81A894F5A686834E580D38C4D4" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="N Iran" country="Iran" location="Mazandaran Province" specimenCount="1" specimenCount-male="1" stateProvince="Mazandaran Province">
<emphasis id="DB123040B5BA506CFC0B0EA797B2BB9D" bold="true" pageId="0" pageNumber="71">
<collectingCountry id="81E9DE028438A13A2D8827A4418D3459" name="Iran">Iran</collectingCountry>
</emphasis>
:
<collectingRegion country="Iran" name="Mazandaran">Mazandaran Province</collectingRegion>
<collectingRegion id="81C686E01467A5889BCF8BFFE49D2BC7" country="Iran" name="Mazandaran">Mazandaran Province</collectingRegion>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="DB2799482747B33AAD7C1D139DD88D4B" type="male">1♂</specimenCount>
, '
<collectorName>N Iran</collectorName>
<collectorName id="AD5B5033192051D57251832BF735F6CA">N Iran</collectorName>
p
</materialsCitation>
.
<materialsCitation collectingDate="2000-06-17" collectorName="Hajdaj E., Collectio, Hajdaj, Morv." country="Iran" determinerName="D. W. Wrase" elevation="2300" latitude="36.416668" location="Hasan Keif" longLatPrecision="1187" longitude="51.033333" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mazandaran">
<normalizedToken originalValue="Mazandarán">
<collectingRegion country="Iran" name="Mazandaran">Mazandaran</collectingRegion>
<materialsCitation id="0DFB5EA3DCB72ED5EA1D2125B35B1CD4" collectingDate="2000-06-17" collectorName="Hajdaj E., Collectio, Hajdaj, Morv." country="Iran" determinerName="D. W. Wrase" elevation="2300" latitude="36.416668" location="Hasan Keif" longLatPrecision="1187" longitude="51.033333" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mazandaran">
<normalizedToken id="2A9A528FDA2E4B6690BFB9D1CC454B9A" originalValue="Mazandarán">
<collectingRegion id="73EB5D395B95D5B7ED6FEBED5DFD039B" country="Iran" name="Mazandaran">Mazandaran</collectingRegion>
</normalizedToken>
/
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
<quantity id="2E2094F3E24F2B25A37BBFDB8BE13D40" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
S
<location LSID="urn:lsid:plazi:treatment:CBC32D678CEC5274987BE44552D6AA13:6712582026365C1461E65DD7A4238047" country="Iran" latitude="36.416668" longLatPrecision="1187" longitude="51.033333" name="Hasan Keif" stateProvince="Mazandaran">Hasan Keif</location>
<location id="6712582026365C1461E65DD7A4238047" LSID="urn:lsid:plazi:treatment:CBC32D678CEC5274987BE44552D6AA13:6712582026365C1461E65DD7A4238047" country="Iran" latitude="36.416668" longLatPrecision="1187" longitude="51.033333" name="Hasan Keif" stateProvince="Mazandaran">Hasan Keif</location>
/
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">2300m</elevation>
<quantity id="6C2407E0453E15EF89A88CC9AD955688" metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">
<elevation id="3CA2C5477BC15EC11452B4169E943766" metricMagnitude="3" metricUnit="m" metricValue="2.3" unit="m" value="2300.0">2300m</elevation>
</quantity>
3625N 5102E [
<geoCoordinate degrees="36" direction="north" minutes="25" orientation="latitude" precision="925" value="36.416668">36°25'N</geoCoordinate>
<geoCoordinate id="FD88E2C6DBE09DEBFBF104271A478F62" degrees="36" direction="north" minutes="25" orientation="latitude" precision="925" value="36.416668">36°25'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="02" orientation="longitude" precision="925" value="51.033333">51°02'E</geoCoordinate>
<geoCoordinate id="521CEBA53899CBFDCFA8BFF241D68CDA" degrees="51" direction="east" minutes="02" orientation="longitude" precision="925" value="51.033333">51°02'E</geoCoordinate>
] /
<collectingDate value="2000-06-17">17.VI.2000</collectingDate>
<collectingDate id="75B938DE7EC58EFF39E84C841A8F3C7A" value="2000-06-17">17.VI.2000</collectingDate>
lgt.
<collectorName>Hajdaj E.</collectorName>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<collectorName id="52BD6AD6AE1DE5978798502BDB1F8AFB">Hajdaj E.</collectorName>
<taxonomicName id="74D8B770333E7CE56AE07CB9BC19A903" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
//
<collectorName>Collectio</collectorName>
<collectorName id="7FA2507827F899C8BF41C9D5D62E409F">Collectio</collectorName>
/
<collectorName>Hajdaj</collectorName>
<collectorName id="F196895D419A12BABC09548B9BFF3465">Hajdaj</collectorName>
//
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="6BB96E51379EB1E79DB3C7F89D7ED0E3" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ cf. /
<taxonomicName authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">Platyderus davatchii</taxonomicName>
<collectorName>Morv.</collectorName>
<taxonomicName id="E7864BB37BC23E5ACB771E1A7DB362C7" authorityName="Morvan" authorityYear="1970" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">Platyderus davatchii</taxonomicName>
<collectorName id="1F370558D6D64FE81B4B6C28CD644ECB">Morv.</collectorName>
/
<determinerName>D.W. Wrase</determinerName>
<determinerName id="221793C5E3F504BDA3A63836F4C7FFD7">D.W. Wrase</determinerName>
det. 01' (cHAJ);
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="4FB996C1FF21534A0F2F2BDEFAA9BBA5" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="4A24F27F4D03832BF2B700BD45BAF0EE" type="female">1♀</specimenCount>
, 'IR
</materialsCitation>
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Seiedy, Muilwijk'" country="Iran" elevation="2500" latitude="36.40003" location="Rudbarak" longLatPrecision="1" longitude="51.035416" specimenCount="1" stateProvince="Mazandaran">
<collectingRegion country="Iran" name="Mazandaran">Mazandaran</collectingRegion>
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10km</quantity>
<materialsCitation id="FCA976B3879E81E063DE06D8475CDD01" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Seiedy, Muilwijk'" country="Iran" elevation="2500" latitude="36.40003" location="Rudbarak" longLatPrecision="1" longitude="51.035416" specimenCount="1" stateProvince="Mazandaran">
<collectingRegion id="8986A6CE1D585D00C7F5680BF1D17BBA" country="Iran" name="Mazandaran">Mazandaran</collectingRegion>
<quantity id="3989500D50CBFE1A59D8FF7926AA72FA" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10km</quantity>
SW /
<location LSID="urn:lsid:plazi:treatment:CBC32D678CEC5274987BE44552D6AA13:DBA818F3DD273755CAE250017B62D324" country="Iran" latitude="36.40003" longLatPrecision="1" longitude="51.035416" name="Rudbarak" stateProvince="Mazandaran">Rudbarak</location>
<geoCoordinate degrees="36" direction="north" minutes="24" orientation="latitude" precision="1" seconds="00.1" value="36.40003">36°24'00.1&quot;N</geoCoordinate>
<location id="DBA818F3DD273755CAE250017B62D324" LSID="urn:lsid:plazi:treatment:CBC32D678CEC5274987BE44552D6AA13:DBA818F3DD273755CAE250017B62D324" country="Iran" latitude="36.40003" longLatPrecision="1" longitude="51.035416" name="Rudbarak" stateProvince="Mazandaran">Rudbarak</location>
<geoCoordinate id="BEEC5056FF4EE45977BB0DE4B05C4B70" degrees="36" direction="north" minutes="24" orientation="latitude" precision="1" seconds="00.1" value="36.40003">36°24'00.1&quot;N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="2" orientation="longitude" precision="1" seconds="07.5" value="51.035416">51°2'07.5&quot;E</geoCoordinate>
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">2500m</elevation>
<geoCoordinate id="B71579D4F7E789AE36AFC2CC96B3089A" degrees="51" direction="east" minutes="2" orientation="longitude" precision="1" seconds="07.5" value="51.035416">51°2'07.5&quot;E</geoCoordinate>
<quantity id="94EC822FA3CCAADF5EB4017E71280D5F" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">
<elevation id="FFDA172BBD4DD0F4730F6A0F62DD57F4" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">2500m</elevation>
</quantity>
/ 16.06.17
<collectorName>Seiedy</collectorName>
<collectorName id="E66C51F942AB4E293FAD91B33BB4D18F">Seiedy</collectorName>
/
<normalizedToken originalValue="Muilwijk’">
<collectorName>Muilwijk'</collectorName>
<normalizedToken id="D8614D7C9C340681772B34AA5D732FB1" originalValue="Muilwijk’">
<collectorName id="897881AB2419DE1F18698E3C8C9C3D57">Muilwijk'</collectorName>
</normalizedToken>
(cMUI)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="4A880EC2CF5025697ADF7FF89D6C14E8" pageId="0" pageNumber="71">
TME:
<specimenCount type="generic">3 specimens</specimenCount>
<specimenCount id="9432A889D78B5EB4D4B193A7BBC8C485" type="generic">3 specimens</specimenCount>
. TGE:
<specimenCount type="male">2♂♂</specimenCount>
<specimenCount id="E047F1C8E040B423A2DF8AB4E32686E2" type="male">2♂♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="F0CF5C0DE8D98AF70C068DED6F953009" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="C4766E3AAEDC6C5E7E144DBC19E6DA13" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="5A91F7D951B4F2A908BC68B5EB1BF421" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="DA17726F7129D6654A61A01FD68DB882" pageId="0" pageNumber="71">
Similar to
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="239446CE7F4A8D3254B74F0C6E10D255" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="01B7FB77675345DAC3A24F9A511FEBF1" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
, but differs from it in pronotum much wider than head (PW/HW&gt;1.40), with apex more constricted compared with widest point (PW/PA&gt;1.40), elytra in relation to pronotum narrower (EW/PW &lt;1.38), and anterior side of mesofemur ventrally with four setiferous punctures. Male specimens of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="988D0EEC53BB0C47DADC6E58FE770BF1" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="C6632F83AA040071F6520A75138BED18" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
can be additionally distinguished from males of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="9798FB8CA61362AC7C5BFDFE91EDCD85" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="650BCFA43FB78F6A08E5B384843A88E2" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
by median lobe at lateral view larger (1.2-1.3 mm, vs. 1.1-1.2 mm, with a longer shaft (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8H</figureCitation>
<figureCitation id="E01BECD79527FC02EB099999847026C3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="redescription">
<paragraph pageId="0" pageNumber="71">Redescription</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="364D665669378D37BB27976F84BEBD71" pageId="0" pageNumber="71" type="redescription">
<paragraph id="516D73B09929FCD054C8AF97492BCD6E" pageId="0" pageNumber="71">Redescription</paragraph>
<paragraph id="F5D61C5DCB728E5AC38F3B39BB850942" pageId="0" pageNumber="71">
(based on non-type material).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<emphasis id="77A0EA884AB9FCF40F83AEAD111229C0" bold="true" pageId="0" pageNumber="71">
<emphasis id="B09F081857927945C9E74AC035DB18B5" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of moderate size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="7BE60B1DE139852A9290A308105113AF" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B54FB63B043BB1BBB9685853E3F5ECA4" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 7.35-7.90 mm; BW: 2.45-2.55 mm), with elongate and subconvex body (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2D</figureCitation>
<figureCitation id="F2D654354EA5FA7C7B8EFF3F510002D9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habitus. A. Platyderus (Eremoderus) sp., female specimen, Karabet Pass, Turkey; B. P. (E.) lassallei, sp. nov., female paratype, E Qolqol, Mazandaran Province, Iran; C. P. (E.) klapperichi, sp. nov., holotype; D. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; E. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure2" httpUri="https://binary.pensoft.net/fig/695639" pageId="0" pageNumber="71">2D</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="DC453C7918233509A3B13947C8A0EB99" bold="true" pageId="0" pageNumber="71">
<emphasis id="081AF4FD69473516BFAE9571CCED0E7C" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
<tableCitation id="A1DA9D69710890572516606693D53FCB" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric data for species of the ' Platyderus Eremoderus lassallei ', ' Platyderus davatchii ' and ' Platyderus Eremoderus afghanistanicus ' groups of Platyderus." httpUri="http://table.plazi.org/id/25F9C0CC5B2F4E7FD96A07933453D3C4" pageId="0" pageNumber="71" tableUuid="25F9C0CC5B2F4E7FD96A07933453D3C4">2</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="A8887519378AC98F07FB84FF75E66EC6" bold="true" pageId="0" pageNumber="71">
<emphasis id="D81820996FAE88BB09D8ACE6F9D59393" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Integument uniformly dark brown, appendages barely lighter than body. Surface moderately shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="9BDECBE694CBDEAE88860FD615CD2E6C" bold="true" pageId="0" pageNumber="71">
<emphasis id="B46C5B620496844E455010A70D22CA6E" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum with regular isodiametric sculpticells posterolaterally and slightly transverse ones anterolaterally, microsculpture faint to absent in middle. Ventral surface (excluding gula) microsculptured, mentum, submentum and proepisternum with regular isodiametric microsculpture, abdominal ventrites and femora with slightly stretched isodiametric sculpticells.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="509569ACFC870C00258AD770BC6420B2" bold="true" pageId="0" pageNumber="71">
<emphasis id="D69EA4C7413F89FBDF42F04CB8F2AA81" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
Significantly narrower than pronotum (mean PW/HW= 1.48). Antennae long, with last three antennomeres exceeding base of pronotum. Eyes long, subconvex. Labrum subrectangular, with anterior margin slightly concave. Frontal furrows small, shallow, subfoveoulate. Paraorbital sulci moderately deep, ending at level of posterior supraorbital pore or lightly before.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="7FC43958BDBF4E945DDF9699647AABDB" bold="true" pageId="0" pageNumber="71">
<emphasis id="35A15ECD977DEBAC8BD4BCC3F808019D" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum slightly wider than long (mean PW/PL= 1.15), widest at second quarter. Anterior and posterior transverse impressions indistinct. Sides rather convex anteriorly, fairly concave posteriorly; mean PW/PA= 1.46, mean PW/PB= 1.25; bead of anterior margin present laterally, reduced in medial 1/8 to 1/10; bead of posterior margin reduced to absent in medial 1/8. Metepisternum slightly longer than wide, MA/MI about 0.9.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="D61FC3BE0B8FE4DEF7B910EE959F48F9" bold="true" pageId="0" pageNumber="71">
<emphasis id="1FF78DE0E890181A7C60370CA274481D" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Long, cylindrical, about one and two thirds as long as wide (mean EL/EW= 1.67), one time and a third as wide as pronotum (mean EW/PW= 1.33; mean EL/PL= 2.55). Stria 7 reaching basal margin. Three elytral discal setiferous punctures in interval 3 (one specimen with four punctures on one elytron, as an additional puncture exists at fourth sixth, between second and third normal punctures), anterior puncture usually adjoining stria 3 (in two specimens adjoining stria 2 on one elytron), remaining two punctures adjoining stria 2 (in specimens from Rudbarak env., right elytron with medial pore adjoining stria 3). Umbilicate setiferous series of 16-17 punctures on each elytron (one specimen with 18 punctures on one elytron).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="0EFEDD559FF522B8ACCC3132073EDA21" bold="true" pageId="0" pageNumber="71">
<emphasis id="5CA64732B8E2C54646197748774C02B2" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one seta in medial third. Anterior side of mesofemur ventrally with four setiferous punctures. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="2E848781E5D28E8C52B68F538A333806" bold="true" pageId="0" pageNumber="71">
<emphasis id="B9C6BF251D84E491FB861005475E640D" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX elongate, suboval, with proximal margin rather pointed (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7F</figureCitation>
<figureCitation id="C7D5EE58DF1BD89E3CC4D743BDD097F4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7F</figureCitation>
). Median lobe of aedeagus in lateral view with shaft longer than in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
<taxonomicName id="7C5BABF53C437D80D405C98A555BAB32" class="Insecta" kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis id="95A300B53856117DD5C9E6D865CEEF69" italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
, and significantly more curved than in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
<taxonomicName id="376BBC5F19260C07500E5620DD1C32A4" class="Insecta" kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="CC7FB148101D0FC1AD17947E5B78417A" italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
, and with apex turned up; median lobe in ventral view straight, ca. 3.85 times longer than wide, with apical lamella nearly symmetrical. Internal sac in lateral view (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8H</figureCitation>
<figureCitation id="9C9226DD3C7A650F801DCAFA6F249740" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8H</figureCitation>
) with ventral sclerite elongate, widened medially and distally; same in ventral view (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10H</figureCitation>
<figureCitation id="D9A41AB696972F0414C3A2D787713037" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10H</figureCitation>
) with dorsal sclerite forming two left-sided protuberances and a large right-sided curve, and ventral sclerite narrow, straight, with distal end curved to left. Right paramere thin, concave ventrally (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12G</figureCitation>
<figureCitation id="2CCE8DDC0DC38B8892BB1797DA815DFE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12G</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="CF865C28300E6D108D721B4F80AD0046" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14B</figureCitation>
<figureCitation id="911E6317A215B3B7812FDD386C70EF5A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14B</figureCitation>
). Apical gonocoxite with blunt apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">All specimens above examined were collected in open places at high altitudes between 2300 m and 2700 m a.s.l. The specimens collected by JM in June 2017 have been caught under stones in a high-mountain meadow covered with lots of flowers, without bushes or trees. The place was definitely not dry during the time of visiting and perhaps covered with snow a few weeks before.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="7808A15C309C13CC10F870B9837384E2" pageId="0" pageNumber="71" type="habitat">
<paragraph id="3479197690BCF3AA7178000CA32E6807" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="4468FCFF14B6EA9ACE4FF8FB7D239948" pageId="0" pageNumber="71">All specimens above examined were collected in open places at high altitudes between 2300 m and 2700 m a.s.l. The specimens collected by JM in June 2017 have been caught under stones in a high-mountain meadow covered with lots of flowers, without bushes or trees. The place was definitely not dry during the time of visiting and perhaps covered with snow a few weeks before.</paragraph>
<paragraph id="770D70384D0D867872188C22E57A55F2" pageId="0" pageNumber="71">
The type series of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Eremoderus) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="Eremoderus">P. (Eremoderus) ledouxi</taxonomicName>
<taxonomicName id="B9884469AA50489301166E89386823DE" class="Insecta" kingdom="Animalia" lsidName="P. (Eremoderus) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="Eremoderus">P. (Eremoderus) ledouxi</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Platyderus) chodjaii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chodjaii" subGenus="Platyderus">P. (Platyderus) chodjaii</taxonomicName>
<taxonomicName id="E3C90E2BB232D0823EA8C0E1334D92EF" class="Insecta" kingdom="Animalia" lsidName="P. (Platyderus) chodjaii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chodjaii" subGenus="Platyderus">P. (Platyderus) chodjaii</taxonomicName>
Morvan, 1974 were collected together (cfr.
<bibRefCitation author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan 1974</bibRefCitation>
<bibRefCitation id="295C121047425074BD1910A6807C28EA" author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan 1974</bibRefCitation>
), demonstrating that the two species occur sympatrically.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="671062E9813D502B807E96433EBAEEAC" pageId="0" pageNumber="71" type="distribution">
<paragraph id="35CA247D783C9317C799ECFFE4CF200A" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="10FE6382DB37AB2191E6CA12044ABCD8" pageId="0" pageNumber="71">
North Iran: the Central Alborz Range, in area of the upper valley of Sardabrud River, northeast of the Takht-e Suleyman Subrange (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="C8278374CEF6837F8C8CEC95802891A4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="58CC53385E9FC0A9C94C2E535EB3044D" pageId="0" pageNumber="71" type="notes">
<paragraph id="A67E158303B002D1278F80F1CE866096" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="A76DC63D5BAD045377EA41CE4FA47452" pageId="0" pageNumber="71">
Although not able to study the holotype, based on the original description of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
<taxonomicName id="99F733CEE3F720EC7B740908CC490360" class="Insecta" kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="F99E1A5BE1027BF653AF871C64C6DD0F" italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
and the illustrations of the median lobe of aedeagus and internal sac (
<bibRefCitation author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan 1974</bibRefCitation>
<bibRefCitation id="87FBB91FC0248D6BBE9BC5252E78C75E" author="Morvan, P" journalOrPublisher="Journal of Entomological Society of Iran" pageId="0" pageNumber="71" pagination="143 - 156" refId="B41" refString="Morvan, P, 1974. Nouveaux coleopteres carabiques d'Iran (6eme note). Journal of Entomological Society of Iran 1: 143 - 156" title="Nouveaux coleopteres carabiques d'Iran (6 eme note)." volume="1" year="1974">Morvan 1974</bibRefCitation>
: 148-149, figs 17-24), we have no doubt that the material examined belongs to this species.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="5E1554CB3BDBF662A9E28A6DC9E85D4B" pageId="0" pageNumber="71">
See also
<normalizedToken originalValue="“Notes”">&quot;Notes&quot;</normalizedToken>
<normalizedToken id="5FC1D481509E05AA44534D24FE247019" originalValue="“Notes”">&quot;Notes&quot;</normalizedToken>
under
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
<taxonomicName id="0B2D77C89FE6C68B6F374972528A06B5" class="Insecta" kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis id="524B3F8406CE84FECE9C584E6EE366F1" italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
</taxonomicName>
Morvan.
</paragraph>

528
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<mods:namePart id="401CBA9928FAB2E69A5A366133945086">Assmann, Thorsten</mods:namePart>
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<mods:affiliation id="41F2B7BFEC55B1DE3A5F1721EDC54103">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
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4.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1D</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5C</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8D</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10D</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12D</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13C</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
<subSubSection id="2633F1EFF0B1410DD140C0F4E6CA28B4" pageId="0" pageNumber="71" type="description">
<paragraph id="B55C46A5FD2E99750B3731210FA9E3D5" pageId="0" pageNumber="71">
<figureCitation id="06D2050E404806BEAC4A3A623494BD4A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1D</figureCitation>
<figureCitation id="5AB27AE93E02DBD2B141C2DEDD8E6445" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5C</figureCitation>
<figureCitation id="FE27E7B42E3DEDF29A50815039C3A597" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8D</figureCitation>
<figureCitation id="EAA501A9224713751E2D16FDCEDF72BA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10D</figureCitation>
<figureCitation id="D1956C969863AFC2AF5F25C3EDE06629" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12D</figureCitation>
<figureCitation id="B5A582E372E4EB45D7FA00C1507D2F44" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13C</figureCitation>
<figureCitation id="6AE0B3F7646B0536E7FC6C626AE9D301" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Iran, Chahar Mahal va Bakhtiari Province, 7 km NE Naghan Town, 31.97472, 50.77694, 2400 m.</paragraph>
<subSubSection id="BB775C2BE6894BC89590234051E2541E" pageId="0" pageNumber="71" type="type locality">
<paragraph id="2C108ECE6F629568FFDB6D4D682D479A" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="FFE426D7E201EC233E32772E81F0D4AC" pageId="0" pageNumber="71">Iran, Chahar Mahal va Bakhtiari Province, 7 km NE Naghan Town, 31.97472, 50.77694, 2400 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="2A30974A247E7A6646C63FD0CF92A197" pageId="0" pageNumber="71" type="type material">
<paragraph id="4F9D60705E36AEA98131E28BAD0A38F5" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="D7A7C3669F9BF9B4CA65BC5D65C8855D" pageId="0" pageNumber="71">
<emphasis id="533F49A49C6F819308F86B2418A31B90" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
1♂, 'IRAN, /
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="0010CB94F2F601CA6C9A6A662587CDAB" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="D418D8A1DB8ED17C01C54817A05D0470" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="0977CC27676AE17B806558B9995290CE" originalValue="vā">va</normalizedToken>
<normalizedToken id="BC262F18498745B82B0791F9629EB212" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
/ 7km NE Naghan Town /
<geoCoordinate degrees="31" direction="north" minutes="58" orientation="latitude" precision="15" seconds="29" value="31.974722">31°58'29&quot;N</geoCoordinate>
<geoCoordinate id="9C041529D174B09765C13704FF835CA9" degrees="31" direction="north" minutes="58" orientation="latitude" precision="15" seconds="29" value="31.974722">31°58'29&quot;N</geoCoordinate>
,
<geoCoordinate degrees="050" direction="east" minutes="46" orientation="longitude" precision="15" seconds="37" value="50.776943">050°46'37&quot;E</geoCoordinate>
<geoCoordinate id="509770EB677F655A6F52ED5D6FA06C28" degrees="050" direction="east" minutes="46" orientation="longitude" precision="15" seconds="37" value="50.776943">050°46'37&quot;E</geoCoordinate>
2400m / 03.04.2007, leg. Jan
<normalizedToken originalValue="Muilwijk’">Muilwijk'</normalizedToken>
<normalizedToken id="2474ADB2B8FE3C8CD9F6DBC913C5678F" originalValue="Muilwijk’">Muilwijk'</normalizedToken>
(HMIM). Paratypes: 1♀, labelled as holotype (cMUI); 1♀, 'IRAN (
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="CD30CE6B3ED58C23B43AAA768D90EA9A" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="050CA976D6C145183E01B06879C35EF8" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="AF95B8FD5C68D5F26FFAF587719AD7D5" originalValue="vā">va</normalizedToken>
<normalizedToken id="562FF6397AE36999AC5266369944BF21" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
) / Zagros Mts. / Boldaghi vill. (nr Choghaklor lake) / 3 km S Sibak, 2600-2700 m /
<geoCoordinate degrees="31" direction="north" minutes="51" orientation="latitude" precision="15" seconds="47" value="31.863056">31°51'47&quot;N</geoCoordinate>
<geoCoordinate id="B252EA66EDEDD4D4110D053E462EE42F" degrees="31" direction="north" minutes="51" orientation="latitude" precision="15" seconds="47" value="31.863056">31°51'47&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="55" orientation="longitude" precision="15" seconds="24" value="50.923336">50°55'24&quot; E</geoCoordinate>
<geoCoordinate id="43F34B5180F5AD8A93C4279072E04741" degrees="50" direction="east" minutes="55" orientation="longitude" precision="15" seconds="24" value="50.923336">50°55'24&quot; E</geoCoordinate>
/ (subalpine slopes with snowfields / under stones) / 20.IV.2018 Wrase &amp; Laser [07]' (cWR); 1♂, 'IRAN (
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="F0CE8E171F2DE187295407D9294F4954" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="9038CE8BFDE4DE446DCADD8ADEA17CA7" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="FAD98227876FECB971045C40134704C9" originalValue="vā">va</normalizedToken>
<normalizedToken id="8050C7000E13F6CA06E52A0E4360EA09" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
) / Zagros Mts., Zard Koh Mt. / Cheri pass 20 km W Samsami / 2775 m,
<geoCoordinate degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
<geoCoordinate id="830706224DA72CF0C0D3DB51E99C294C" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
/
<geoCoordinate degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
<geoCoordinate id="0708910A3E494856F2ACB0DD0E407F00" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
/ (subalpine slopes/under stones) / 21.IV.2018 Wrase &amp; Laser [08]' (cWR); 5♂♂, 1♀, 'IRAN, Zagros Mts., P: / Chahar Mahal va / Bachtiari, Zard Koh Mt. / pass, subalpine slope // 2770m, 21.IV.2018 /
<geoCoordinate degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
<geoCoordinate id="105EAB20E4AF1D98DA479F4C85FB20CD" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
/
<geoCoordinate degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
<geoCoordinate id="2DAC0992AEB0C7D84D95BEEBF108DFDC" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
/ leg. M.
<normalizedToken originalValue="Hartmann’">Hartmann'</normalizedToken>
<normalizedToken id="68E3653CC52A1B5B623F31B8BE90EB8F" originalValue="Hartmann’">Hartmann'</normalizedToken>
(c NME); 10♂♂, 5♀♀,
<geoCoordinate degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
<geoCoordinate id="3320F72027F03AC81C561C33FA8B35B0" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
<geoCoordinate id="BCD6C44B6B8BD53479EAB6BE3FE24949" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
Iran / Zagros Mts., p.
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="CBC2A82839DD4971677DDD9811D64D6A" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="8DAEC9EE492F9A41EC0E28E82153558D" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="69D270E3EA03315FA628D1B7236A35FA" originalValue="vā">va</normalizedToken>
<normalizedToken id="7CA2D5267ABDA57AD1B5E537654FCC64" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Zard Koh Mts., pass / 20km W Samsami, subalpine / slopes 21.IV.2018 2.775m NN / leg.: Schnitter IR49' (NME); 12♂♂, 2♀♀, 'IRAN: Zagros Mts., p.
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="61512F531FB60675378DE949941A3564" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="164A5A0EA61406D2F63683F20659C946" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="B58A10B94E1788686CF451A36EB633F4" originalValue="vā">va</normalizedToken>
<normalizedToken id="06E8C8FED56B92E355EA0E2F26968F44" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Zard Koh Mts.,pass 20km / W Samsami,
<geoCoordinate degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
<geoCoordinate id="DA6982F1EF78F1EA03F0E01DCBB6BEF3" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
<geoCoordinate id="4F496C39829FDC25CED6169644878A17" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
/ 2775m, 21.IV.2018, subalp. slopes / leg. A.Weigel #49' (NME); 1♀, 'IR Chaharmahal and / Bakhtiari Cheri pass / 4.vi.2018 3000m /
<geoCoordinate degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="31" value="32.15861">32°09'31&quot;N</geoCoordinate>
<geoCoordinate id="08807C17ADF033152AB2281C799AC2C3" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="31" value="32.15861">32°09'31&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="47" value="50.17972">50°10'47&quot;E</geoCoordinate>
<geoCoordinate id="05DC41C3A80CF63103F4E21EAC6752B4" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="47" value="50.17972">50°10'47&quot;E</geoCoordinate>
/ Muilwijk J.' (cMUI); 1♂, 'IR Chaharmahal and / Bakhtiari Cheri pass / 5.vi.2018 2450m /
<geoCoordinate degrees="32" direction="north" minutes="10" orientation="latitude" precision="15" seconds="36" value="32.176666">32°10'36&quot;N</geoCoordinate>
<geoCoordinate id="F9CBA0E8B2422A6EDCE1D2EAD00A2815" degrees="32" direction="north" minutes="10" orientation="latitude" precision="15" seconds="36" value="32.176666">32°10'36&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="06" value="50.168335">50°10'06&quot;E</geoCoordinate>
<geoCoordinate id="504D437304ED467E5DD4A9D4FB417E52" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="06" value="50.168335">50°10'06&quot;E</geoCoordinate>
/ Muilwijk J.' (cMUI); 9♀♀, 'IR Chahar Mahaal Va / Bakhtiari Chery pass / 31.v.2019 Muilwijk J. / 2950-3260 m /
<geoCoordinate degrees="32" direction="north" minutes="10" orientation="latitude" precision="15" seconds="51" value="32.180836">32°10'51&quot;N</geoCoordinate>
<geoCoordinate id="22CAC833687D5AB663BA93547501B799" degrees="32" direction="north" minutes="10" orientation="latitude" precision="15" seconds="51" value="32.180836">32°10'51&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="09" orientation="longitude" precision="15" seconds="59" value="50.16639">50°09'59&quot;E</geoCoordinate>
<geoCoordinate id="DBB14EC337B2374C455286BF16B033F5" degrees="50" direction="east" minutes="09" orientation="longitude" precision="15" seconds="59" value="50.16639">50°09'59&quot;E</geoCoordinate>
(cMUI, HMIM); 1♂, 2♀♀, 'IRAN (
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="4D106C356D783996BF3A2A906FF13E2C" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="F8B1B86701249C44D3B139279F052913" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="1401F40A059A8E150659D4BD2B6E6002" originalValue="vā">va</normalizedToken>
<normalizedToken id="D9EB7D999030834330778944FC4547FC" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
) / Zagros Mts., 5 km SW Asad Abad /
<geoCoordinate degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
<geoCoordinate id="F231DD68CEE0A9F2C112DB6E3A878D1D" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
/
<geoCoordinate degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
<geoCoordinate id="337141E15B2C4A14A6C6A35A0FAFA060" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
/ ca 2400 m (subalpine slopes/ / under stones) / 19.IV.2018 Wrase &amp; Laser [03]' (cWR); 3♂♂,
<geoCoordinate degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
<geoCoordinate id="84107A4794293139D94D398AF911DA4A" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
<geoCoordinate id="E1E06787294C6B48B6012AD88F0949E9" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
Iran / Zagros Mts., p.
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="24D902F8DA06E54DF5A3BDF9DEDD40B5" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="DEEF6F5029ADA4714A2701B8B30DF864" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="D75BB33CFB93A9EB0B5F3968716F3A23" originalValue="vā">va</normalizedToken>
<normalizedToken id="1EC1043E6F922B3D216D382C441565D1" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Asad Abad 5 km SW / snow field, high pasture / 19.IV.2018 2.500-2.770m NN / leg.: Schnitter IR 44' (NME); 1♂ (genitalia lost after study), 3♀♀, 'IRAN Zagros Mts., P: Chahar / Mahall va Bachtiari, Asad / Abad 5 km SW,
<geoCoordinate degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
<geoCoordinate id="B31387129408E0AF48AC1FF2F74B1BCC" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
, /
<geoCoordinate degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
<geoCoordinate id="1D61B5C88E99FABCF425B79789C83F2E" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
snow field, high / pasture, 19.IV.2018, 2.500- / 2.770m, leg. A. Weigel #18-04' (NME); 3♂♂, 'IRAN: Zagros Mts., p.
<normalizedToken originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="2AE34B65E6791DB093336B3C916E9279" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="17752E78145C52DA83DC5EECBDC0E835" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken originalValue="vā">va</normalizedToken>
<normalizedToken originalValue="Bachtīārī">Bachtīarī</normalizedToken>
<normalizedToken id="D1C63BFB9BC08991987DC20F4DD7E226" originalValue="vā">va</normalizedToken>
<normalizedToken id="AFD707C107DB73AEF24712A566D3C9DB" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Asad Abad 5 km SW, / 19.IV.2018,
<geoCoordinate degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
<geoCoordinate id="34CF8A40A6D0489760E95068245E21D8" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
<geoCoordinate id="4F23A8EB4358F9AE944D99E72290A1B3" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
/ 2500-2770m, snow field, high / pasture leg. A. Weigel #44' (NME); 1♂, 'IRAN, Zagros Mts., P: / Chahar Mahal va / Bachtiari, Asad Abad / 5 km SW, 2500-2770m // 19.IV.2018, snow field / high pasture, 32°20'30&quot;N, 50°32'59&quot;E / leg. M. Hartmann, #3/18' (NME); 4♂♂, 3♀♀, 'IR Esfahan Kamran Pass / 1.vi.2019 Muilwijl J. / 3000-3300m /
<geoCoordinate degrees="32" direction="north" minutes="45" orientation="latitude" precision="15" seconds="49" value="32.76361">32°45'49&quot;N</geoCoordinate>
<geoCoordinate id="ECE5D799621A25F992492B2798E6A10D" degrees="32" direction="north" minutes="45" orientation="latitude" precision="15" seconds="49" value="32.76361">32°45'49&quot;N</geoCoordinate>
,
<geoCoordinate degrees="50" direction="east" minutes="02" orientation="longitude" precision="15" seconds="34" value="50.042778">50°02'34&quot;E</geoCoordinate>
<geoCoordinate id="FBA0FB1D1DD155D15CB0C88387A59DE3" degrees="50" direction="east" minutes="02" orientation="longitude" precision="15" seconds="34" value="50.042778">50°02'34&quot;E</geoCoordinate>
' (cMUI); 1♂, 'IR Esfahan Pashadagan / 2.vi.2019 / Muilwijl J. 3000-3300m /
<geoCoordinate degrees="32" direction="north" minutes="45" orientation="latitude" precision="15" seconds="48" value="32.763332">32°45'48&quot;N</geoCoordinate>
<geoCoordinate id="B301A05E686C0A707D6ED66EBBD38CA0" degrees="32" direction="north" minutes="45" orientation="latitude" precision="15" seconds="48" value="32.763332">32°45'48&quot;N</geoCoordinate>
,
<geoCoordinate degrees="49" direction="east" minutes="59" orientation="longitude" precision="15" seconds="52" value="49.99778">49°59'52&quot;E</geoCoordinate>
<geoCoordinate id="AD6B9F84C50B1DE52283479982DA92F8" degrees="49" direction="east" minutes="59" orientation="longitude" precision="15" seconds="52" value="49.99778">49°59'52&quot;E</geoCoordinate>
' (cMUI).
</paragraph>
<caption doi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 4.</emphasis>
<caption id="BE3587610FBEE3D7C93C5DBDADB42081" doi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71" start="Figure 4" startId="F4">
<paragraph id="AA1D335EE6AE36D6ECF0FD510233238C" pageId="0" pageNumber="71">
<emphasis id="30923E4D5DC1EE2E5DEF00F35A437205" bold="true" pageId="0" pageNumber="71">Figure 4.</emphasis>
Dorsal habitus.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authority="insignitus" authorityName="insignitus Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="Eremoderus" subSpecies="insignitus">Platyderus (Eremoderus) insignitus insignitus</taxonomicName>
<emphasis id="AB4ABAFCB9D3160D474A0E872398D720" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="B3B0D1FC7DD5895399518DC28B7EDB32" authority="insignitus" authorityName="insignitus Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="Eremoderus" subSpecies="insignitus">Platyderus (Eremoderus) insignitus insignitus</taxonomicName>
Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
<emphasis id="47F17E3DF73E5054F71BE39BBD64F0E5" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="865F06177EF9FC26B5B201D6266469EF" class="Insecta" kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
<emphasis id="338A1524A011A145C3194C6D3CC5380A" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="6669B1FE4CA1B761FED1247B1DFBE493" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="EC0913A171172910F4881A8229FCAECC" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="AE9D76FC751F7D793A961C29414EBB6F" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), lectotype and its labels;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="F62301DA9FDA90BFE454155DFCA984C7" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="26F0E1023703AAD76E499D9D040CB567" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), male specimen, &quot;Jerusalem Syria&quot;, Jerusalem District, Israel. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="71">TME: 72 specimens. TGE: 16♂♂, 3♀♀.</paragraph>
<paragraph id="FE6F8D76F8D0A043B639A781E8D70D7B" pageId="0" pageNumber="71">TME: 72 specimens. TGE: 16♂♂, 3♀♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">Adjective, derived from the name of the country where the species was collected.</paragraph>
<subSubSection id="6BA95D3737F085992C9572419E98BE7D" pageId="0" pageNumber="71" type="etymology">
<paragraph id="2EFDC422E77B4A1BA157DA13265ED319" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="1483EDF18D2792E30C711FC7592B3A28" pageId="0" pageNumber="71">Adjective, derived from the name of the country where the species was collected.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="D89110C98ADA47D97E8C9F75BEB9C568" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="FD140145B0305EB0CBF716F07FB043B0" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="666CF3298DE8876E16AA015497F53D05" pageId="0" pageNumber="71">
See
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="201908CCCEAE13A9B8CD40BA41C8301B" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
under
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="7803A6891CEDB8D3D7C8577C9496F9EA" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="B340ADAA2C5047C9A8798BBDB09E2FF5" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="0137C4AB5C9DCCFB0A3B15D8A256F6A8" pageId="0" pageNumber="71" type="description">
<paragraph id="F37316CD05186C22A17079C3CC5370B0" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="DAE4CD739765378647CFF7014659586C" pageId="0" pageNumber="71">
<emphasis id="E6495CDD8D7F3EA9333AF92FDC41BACB" bold="true" pageId="0" pageNumber="71">
<emphasis id="7DDBECB89C63CC58FA09D8A9B4C3910F" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of small size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="5FDBBF6B7C99ED644AFC861AA9044576" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="5C926A9F0037B335669FD22C99FB7940" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 4.30-6.65 mm; BW: 1.50-2.30 mm), with relatively short, moderately convex body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1D</figureCitation>
<figureCitation id="291F1E8EB6B3DC37960600D165E675DE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1D</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratio</emphasis>
<emphasis id="6C06C1787F7DB7BA8FA1C8E55FEDE6D7" bold="true" pageId="0" pageNumber="71">
<emphasis id="A756914BB66C7148BCB4927FA130E5B2" italics="true" pageId="0" pageNumber="71">Measurements and ratio</emphasis>
s.
</emphasis>
See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
<tableCitation id="97BF5D8525B1FC566E17AB731D822600" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="B735B4103EA779E2C784A04FD1367052" bold="true" pageId="0" pageNumber="71">
<emphasis id="B26AC7AA82FC845C68918DFA6803A34E" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Integument uniformly yellow-brown to reddish-brown, appendages somewhat lighter than body. Surface moderately shiny.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="F199DDAED346ADBA6F72186E415C4BAF" bold="true" pageId="0" pageNumber="71">
<emphasis id="93D1EAEBF5D0EBD3AE50378747970D57" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Clypeus, vertex and disc of head with reduced microsculpture. Pronotum with regular isodiametric sculpticells posterolaterally and scarcely-visible to absent microsculpture in middle and anteriorly. Ventral surface with microsculpture and dorsal punctation as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="CE464DD7813596897CED75E379BC5AF7" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="29428B7C77A86D206D5D92D6929179D5" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="4B1EAB9A2AB914B2BB4CF31B66220C9F" bold="true" pageId="0" pageNumber="71">
<emphasis id="27078162DB9A7934C836071ACD6BC185" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About two thirds as wide as pronotum (mean PW/HW= 1.37). Eyes moderately convex. Labrum subrectangular, as long as or slightly shorter than clypeus, with anterior margin straight. Frontoclypeal suture and frontal furrows as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="8B94DD3B831D43C42AFDEE3ACDD22D15" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="93C2FCB4762445741CA702DDE5AFD41A" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. Paraorbital sulci moderately deep, ending before level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="361A2FF17FCABCD8A9627B49F9163300" bold="true" pageId="0" pageNumber="71">
<emphasis id="5F2275EBB5D2CDDAE64A50D6DDD9D318" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum wider than long (mean PW/PL= 1.27), with widest point at second quarter. Anterior and posterior transverse impressions indistinct. Anterior bead lacking in medial 1/10 to 1/8. Sides convex anteriorly, straight posteriorly. Metepisternum somewhat longer than wide, MA/MI= 0.9-1.0.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="F86C0BFB39D818CB75D959B5C09179B6" bold="true" pageId="0" pageNumber="71">
<emphasis id="1E532F8357BDFEED182461C98DF75B3C" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Cylindrical, one and a half times or more as long as wide (mean EL/EW= 1.61), wider than pronotum and relatively long in relation to pronotum length (mean EW/PW= 1.31; mean EL/PL= 2.66). Parascutellar striole and striae as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="13C287E8C4DC813B89B31167C0A0CD9B" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="9F910B4A7250820ACE5EE3BCA6CF79D8" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. Interval 3 with two or three minute discal setiferous punctures, anterior puncture being mostly absent. Umbilicate setiferous series consisting mostly of 15 or 16 punctures.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="68AF99EA3EADDDA9E37F3FCE6717CE55" bold="true" pageId="0" pageNumber="71">
<emphasis id="2FABA926C876C7CF19A4FEB6D9E9D47C" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Anterior side of mesofemur ventrally with four setiferous punctures (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5C</figureCitation>
<figureCitation id="F83B54D2202EFA2368FFAF4B87A6A46D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5C</figureCitation>
). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="01E37C0E5DA1AB3196364C8937AB81EB" bold="true" pageId="0" pageNumber="71">
<emphasis id="0453F6CA381E12FCDA824C70ACF72C30" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="47B334FF391BF292CAC1D4E6E761D142" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="813FFABCA91FE8CA191E2CFDCB670A52" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov., but significantly narrower, with shaft less protruding dorsally and more constricted proximally (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8D</figureCitation>
<figureCitation id="E58846D34B9FC4F1FDD2C0FC1762FACD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8D</figureCitation>
); median lobe in ventral view straight, ca. 3.7 times longer than wide; apical lamella nearly symmetrical (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10D</figureCitation>
<figureCitation id="E52B42FDC609A4EBF32B3BC769E096F7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10D</figureCitation>
). Internal sac in lateral view as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="7EACB159C45A9A11DC69EBAFB51002D4" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="D03CA187AD51846A6EC4226638115D6A" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov., but with ventral sclerite truncate distally; same in ventral view as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="22D3003C4D2052A5DE9F45348F08FB64" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="7626D3EC5D26ED70C0CA01FFD50A3DFF" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. Right paramere as that of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="0FD6782A0986FB95600FF7BECDEB5222" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="EC0E2937E1E88A013FEA4D353175F836" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. but somewhat finer (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12D</figureCitation>
<figureCitation id="C0714296BCE7DFCD2BB64432AC56BACC" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12D</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="5AA55889EC069378726D5D3125EE8AE8" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13C</figureCitation>
<figureCitation id="040ABA6BCDD6221048C496612E07E88E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13C</figureCitation>
). Apical gonocoxite with blunt apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="2B77E96B32CF7C9BDCC4E7D390264BAE" pageId="0" pageNumber="71" type="habitat">
<paragraph id="B4DF59BA991DEE9ACD5DD8B45589FF61" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="770C9ECBEF054B7BB58BC981F38FA8FA" pageId="0" pageNumber="71">
The holotype and one female paratype were collected along an artificial lake situated in a valley northwest of village Aliabad. They were caught at grassland with lots of stones situated along this lake. The vegetation there consisted of grassland, fields and small orchards - as a whole a cultivated landscape. Other ground beetles collected at this place, together with the new species, were
<taxonomicName authorityName="Osculati" authorityYear="1844" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus maurus subsp. osculatii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="maurus" subSpecies="osculatii">
<emphasis italics="true" pageId="0" pageNumber="71">Carabus maurus osculatii</emphasis>
<taxonomicName id="80EF32C0CF2EF40C1C4AA23A04FBDFDD" authorityName="Osculati" authorityYear="1844" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus maurus subsp. osculatii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="maurus" subSpecies="osculatii">
<emphasis id="958448BAF260FF2C87BF9FD2BCB247D9" italics="true" pageId="0" pageNumber="71">Carabus maurus osculatii</emphasis>
</taxonomicName>
Villa, 1844,
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1868" class="Insecta" family="Carabidae" genus="Poecilus" kingdom="Animalia" lsidName="Poecilus festivus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="festivus">
<emphasis italics="true" pageId="0" pageNumber="71">Poecilus festivus</emphasis>
<taxonomicName id="5DEAFC138CD0CD39936212B736511502" baseAuthorityName="Chaudoir" baseAuthorityYear="1868" class="Insecta" family="Carabidae" genus="Poecilus" kingdom="Animalia" lsidName="Poecilus festivus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="274343B574FFE65D50B2A0635FB68C51" italics="true" pageId="0" pageNumber="71">Poecilus festivus</emphasis>
</taxonomicName>
(Chaudoir, 1868) and
<taxonomicName baseAuthorityName="Heer" baseAuthorityYear="1841" class="Insecta" family="Carabidae" genus="Amblystomus" kingdom="Animalia" lsidName="Amblystomus niger" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="niger">
<emphasis italics="true" pageId="0" pageNumber="71">Amblystomus niger</emphasis>
<taxonomicName id="097B0D19F89146CDEC290A2BC18E76D0" baseAuthorityName="Heer" baseAuthorityYear="1841" class="Insecta" family="Carabidae" genus="Amblystomus" kingdom="Animalia" lsidName="Amblystomus niger" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="937A36F67C359028FF3B5CB54E27CFDA" italics="true" pageId="0" pageNumber="71">Amblystomus niger</emphasis>
</taxonomicName>
(Heer, 1841).
</paragraph>
<paragraph pageId="0" pageNumber="71">Cheri Pass environments were represented by overgrazed grasslands with rocks and stones at lower altitudes and stone slopes with sparse vegetation and snow fields at higher altitudes. Similarly, Kamran Pass habitats were consisted of overgrazed grasslands at lower altitude and of stony (sub)alpine slopes with sparse vegetation and snow fields at higher altitudes.</paragraph>
<paragraph pageId="0" pageNumber="71">Habitats around Asad Abad represent grasslands on subalpine slopes; specimens there were collected under stones, sometimes near edges of snowfields.</paragraph>
<paragraph id="CA2AA32A5C4163292290DA94603BB603" pageId="0" pageNumber="71">Cheri Pass environments were represented by overgrazed grasslands with rocks and stones at lower altitudes and stone slopes with sparse vegetation and snow fields at higher altitudes. Similarly, Kamran Pass habitats were consisted of overgrazed grasslands at lower altitude and of stony (sub)alpine slopes with sparse vegetation and snow fields at higher altitudes.</paragraph>
<paragraph id="256FA30DAFC7A4D19399703D645996E0" pageId="0" pageNumber="71">Habitats around Asad Abad represent grasslands on subalpine slopes; specimens there were collected under stones, sometimes near edges of snowfields.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">Southwest Iran: Central Zagros Mountain Chains. So far, four isolated populations are known: (1) Borujen County; (2) Kuhrang County (type locality); (3) Shahr-e Kord County (locality southwest Asad Abad); (4) Fereydunshahr County (Kamran Pass), the first three in Chahar Mahal va Bakhtiari Province, the fourth one in Isfahan Province (near border area with Chahar Mahal va Bakhtiari Province).</paragraph>
<subSubSection id="35301B938720F4DE746565825BFDDF92" pageId="0" pageNumber="71" type="distribution">
<paragraph id="AB68C52710B59C0CAB4B07DEA7C5BB20" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="E579426763A327FCD66212EF6E2712F0" pageId="0" pageNumber="71">Southwest Iran: Central Zagros Mountain Chains. So far, four isolated populations are known: (1) Borujen County; (2) Kuhrang County (type locality); (3) Shahr-e Kord County (locality southwest Asad Abad); (4) Fereydunshahr County (Kamran Pass), the first three in Chahar Mahal va Bakhtiari Province, the fourth one in Isfahan Province (near border area with Chahar Mahal va Bakhtiari Province).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="53420298D6D6A81D3FDBCEA6DDC2DFF9" pageId="0" pageNumber="71" type="notes">
<paragraph id="D896A0F75E918FCE236065D1CB306C88" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="8AA8DDD4AF52910003932EF60FF78C04" pageId="0" pageNumber="71">
The two specimens collected 7 km northeast of Naghan Town represent the nearest population of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
<taxonomicName id="C77E768AE7E28648CD246C4D9AA09681" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="20133AA4FC6C28E2D80111D95B8E5848" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
sp. nov. to the type locality of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
<taxonomicName id="9EF4B585EB9893A56C961C1970F2E09D" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="E5228728FDCFE5805BF74517B7AF830E" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov., ca. 17 km in a straight line (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
<figureCitation id="654F03018AA6CFA5B8A57BB1BA8CA288" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
). The specimen from the environments of the Choghaklor Lake was collected together with three specimens of
<taxonomicName baseAuthorityName="Menetries" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">Platyderus (s. str.) umbratus</taxonomicName>
<taxonomicName id="58F908B9DEF916484810DCBF58518756" baseAuthorityName="Menetries" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">Platyderus (s. str.) umbratus</taxonomicName>
(Men.).
</paragraph>
<paragraph pageId="0" pageNumber="71">The distance between the population from the type locality (near Cheri Pass) and that one southwest of Asad Abad is ca. 45 km in a straight line. On the other hand, the distance between Cheri Pass and locality northeast of Naghan Town is ca. 62 km in a straight line. Respectively, the distance between the locality southwest of Asad Abad and that one northeast of Naghan Town is about 47 km in a straight line. The distance between the type locality (near Cheri Pass) and the locality of Kamran Pass is about 67 km in a straight line (the same distance is between the populations from the Kamran Pass and that of Asad Abad). The distance between the Kamran Pass (Fereydunshahr County) and the locality northeast of Naghan Town is about 112 km in a straight line.</paragraph>
<caption doi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 5.</emphasis>
<paragraph id="F699B38DC622C20A1B13C99A9F56F0B8" pageId="0" pageNumber="71">The distance between the population from the type locality (near Cheri Pass) and that one southwest of Asad Abad is ca. 45 km in a straight line. On the other hand, the distance between Cheri Pass and locality northeast of Naghan Town is ca. 62 km in a straight line. Respectively, the distance between the locality southwest of Asad Abad and that one northeast of Naghan Town is about 47 km in a straight line. The distance between the type locality (near Cheri Pass) and the locality of Kamran Pass is about 67 km in a straight line (the same distance is between the populations from the Kamran Pass and that of Asad Abad). The distance between the Kamran Pass (Fereydunshahr County) and the locality northeast of Naghan Town is about 112 km in a straight line.</paragraph>
<caption id="60C9768E5594E7C17DC142D58579ABB7" doi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71" start="Figure 5" startId="F5">
<paragraph id="6CD6FF3A75EF78699C9736420E8ED0EF" pageId="0" pageNumber="71">
<emphasis id="55E0F9645EA65A11D15FE9E25A53B04D" bold="true" pageId="0" pageNumber="71">Figure 5.</emphasis>
Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures).
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">Platyderus (Eremoderus) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<emphasis id="3ED81C56EC195D854092DC791E04D2F8" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="F46DB3F1BFDC9C1E5E4CD5A7DF8A8D92" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">Platyderus (Eremoderus) weiratheri</taxonomicName>
<normalizedToken id="81C7FE8B61E63C8BCF7B059A93DEEAD3" originalValue="Mařan">Maran</normalizedToken>
, 1940, female specimen, Boz
<normalizedToken originalValue="dağ">dag</normalizedToken>
<normalizedToken originalValue="köy">koey</normalizedToken>
<normalizedToken id="C696279DA373452A9FA3BEE0C1320393" originalValue="dağ">dag</normalizedToken>
<normalizedToken id="89A9621B2F642EEF72308B8BF4843807" originalValue="köy">koey</normalizedToken>
env., Izmir District, Turkey;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
<emphasis id="8039EF565473717C542719E511C0F190" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="4447411521BEC0760FAE743F91F608AC" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
<emphasis id="BBE89BBAED171813DBA4F1694DBD169D" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="F0658CB01CB51CDFFB7AA1F34082907E" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
<emphasis id="42E3AF97AB1A2A508C67354DD58EAAB4" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="B7922B043E45CA18637C8C34FA5E2963" class="Insecta" kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi" subGenus="E.">P. (E.) klapperichi</taxonomicName>
<emphasis id="EC8A2D9E4C7694B0A69A47B2501E1393" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="AFEA46346CE200117E730F28F97EEBAC" class="Insecta" kingdom="Animalia" lsidName="P. (E.) klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi" subGenus="E.">P. (E.) klapperichi</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
<emphasis id="EACE73164284DC0FB5A34BCE6D8D21D2" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="98AC6949753FE8C36958513E67B01A93" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, male specimen, Tochal, Tehran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="E.">P. (E.) afghanistanicus</taxonomicName>
<emphasis id="F3002E08AF6240E11A90EE0925244BB2" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="442899284229F9E714C80D6E32FCA7F8" class="Insecta" kingdom="Animalia" lsidName="P. (E.) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="E.">P. (E.) afghanistanicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
<emphasis id="A57306EA78C25E287BD810BF33F2CE02" bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName id="5857BE09CF994448A231ED5A0C34A328" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, lectotype;
<emphasis bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
<emphasis id="81BC1435291B956D15820CA59DBF232C" bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName id="95C3CC3C5CE1AD3B9EA0E6B3ED264495" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="71">
<paragraph id="1990771B4AAC5EC239895F3EC41E7754" pageId="0" pageNumber="71">
&quot;
<emphasis bold="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
<emphasis id="818FC640FD26E1049A47E01A6C8EDAB5" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="F3E74EF3BD38D5B34FB96765071B8A0D" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis id="F22B550870FEB1CF4BEB935D06FABD33" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
</taxonomicName>
&quot; subgroup
</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
<paragraph id="265E4111A8B6629CED1D4E48E68B12FA" pageId="0" pageNumber="71">
<emphasis id="A498351687713657A6B55B737AFF3E51" bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
By its parascutellar striole and striae 1-8 more coarsely and densely punctate and more deeply impressed, the species from Eastern Turkey differ from those of the &quot;
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
<taxonomicName id="CB734DA94A9515AE6169C1334AD8F3B3" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="18D3A56F2EF324338854BE8C95DECBE0" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
&quot; subgroup.
</paragraph>

382
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<paragraph id="03E037DCCB7F7330AEF0CEE3F3A24E47" pageId="0" pageNumber="71">
14.
<taxonomicName LSID="http://zoobank.org/C5B5E518-02FF-4E24-A1C1-70F560699170" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) brunki</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<taxonomicName id="0ECCB9A3FBECE250EA7C6ACE5492B027" LSID="http://zoobank.org/C5B5E518-02FF-4E24-A1C1-70F560699170" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) brunki</taxonomicName>
<taxonomicNameLabel id="79EE0F4E2E65C38F580F0200F7990736" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3C</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14F</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
<subSubSection id="EF4EF542AD41BDF96117896548AB61E1" pageId="0" pageNumber="71" type="description">
<paragraph id="419D06C5BBECED231E2AE34446C86E20" pageId="0" pageNumber="71">
<figureCitation id="C8F68E13A8A23E7301C52580441EE82B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3C</figureCitation>
<figureCitation id="47B763F39EE6C89F5BF6069F18E874AE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14F</figureCitation>
<figureCitation id="0181E6B62C14CEB7101D21F8F9F5FB28" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Yemen, 'Amran Governorate, Thula District, between Kaukaban and Shibam, approximate GPS coordinates: 15.503, 43.901.</paragraph>
<subSubSection id="7E61355898DDC8C7C8F8206F0411E19F" pageId="0" pageNumber="71" type="type locality">
<paragraph id="F7E905F9019D4BF13C8D0E990743FE22" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="190CCBE8E4E87B97260A329031A62D2A" pageId="0" pageNumber="71">Yemen, 'Amran Governorate, Thula District, between Kaukaban and Shibam, approximate GPS coordinates: 15.503, 43.901.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="A198F7549074E449D563AC6175374E96" pageId="0" pageNumber="71" type="type material">
<paragraph id="3DD82EA3325B0ED9F2BF8E38854A59CE" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="B808AD9E4C0BE003FD1AE985737A2385" pageId="0" pageNumber="71">
<emphasis id="E6AB8ADCE3702774DCACE6C447536221" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♀, 'YEMEN, ~ 40 km NW Sanaa, / Kaukaban/Shibam, small waste- / waterstream, ruralic vegetation, / sand, 2700-2930 m Handaufsam. /
<geoCoordinate degrees="15" direction="north" minutes="30" orientation="latitude" precision="1" seconds="9.98" value="15.502772">15°30'9.98&quot;N</geoCoordinate>
<geoCoordinate id="038E34B8604E81C2E793C600A4893621" degrees="15" direction="north" minutes="30" orientation="latitude" precision="1" seconds="9.98" value="15.502772">15°30'9.98&quot;N</geoCoordinate>
,
<geoCoordinate degrees="43" direction="east" minutes="54" orientation="longitude" precision="1" seconds="2.66" value="43.90074">43°54'2.66&quot;E</geoCoordinate>
<geoCoordinate id="27201D6A6A3DF56C7C953FE0F8547D2B" degrees="43" direction="east" minutes="54" orientation="longitude" precision="1" seconds="2.66" value="43.90074">43°54'2.66&quot;E</geoCoordinate>
/ 02. Dez. 2009, legit. Ingo Brunk [p, w] // Coll. / I. Brunk / Dresden/Germ [h, pn]' (cBRU).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="1938-02-03" collectorName="El Errein, B. M. Exp., S. W. Arabia., H. Scott, E. B. Britton., Reiche" country="YEMEN" county="Remains" determinerName="E. B. Britton" elevation="2835" location="Haz" municipality="El Errein" specimenCount="1" stateProvince="San'a">
<collectingCounty>Remains</collectingCounty>
<subSubSection id="0101859E89DD6A70FA0B77C4C1D985F2" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="7288BF644BF9AA45826C48D4F9529404" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="BCFB34D9E5AECFFD536C5700A718F636" pageId="0" pageNumber="71">
<materialsCitation id="AFF509186082773F1F7D638E8BB34A32" collectingDate="1938-02-03" collectorName="El Errein, B. M. Exp., S. W. Arabia., H. Scott, E. B. Britton., Reiche" country="YEMEN" county="Remains" determinerName="E. B. Britton" elevation="2835" location="Haz" municipality="El Errein" specimenCount="1" stateProvince="San'a">
<collectingCounty id="3778818EE4F536D38FFB2E7BB2930058">Remains</collectingCounty>
of a specimen of uncertain sex in bad condition (only pronotum and elytra preserved), 'UNDER A STONE //
<collectingCountry name="Yemen">YEMEN</collectingCountry>
<collectingCountry id="AD307D33B06856E4B8C121FE889DDAF5" name="Yemen">YEMEN</collectingCountry>
/
<collectorName>
<collectingMunicipality>El Errein</collectingMunicipality>
<collectorName id="F0D8EFC4A129C9E08395B0C9A27C938A">
<collectingMunicipality id="BE6FC64FC49BCC1064D67E13596FADE7">El Errein</collectingMunicipality>
</collectorName>
, nr.
<location LSID="urn:lsid:plazi:treatment:D816CC852981571EA5B1E555A3DEC960:3E358768E5D3A423AA89691B0268ECCE" country="YEMEN" county="Remains" municipality="El Errein" name="Haz" stateProvince="San'a">Haz</location>
<location id="3E358768E5D3A423AA89691B0268ECCE" LSID="urn:lsid:plazi:treatment:D816CC852981571EA5B1E555A3DEC960:3E358768E5D3A423AA89691B0268ECCE" country="YEMEN" county="Remains" municipality="El Errein" name="Haz" stateProvince="San'a">Haz</location>
, / about
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.5749504" unit="mi" value="16.0">16 miles</quantity>
<quantity id="32C5BDACCC15151EDD1DA48E364BC047" metricMagnitude="4" metricUnit="m" metricValue="2.5749504" unit="mi" value="16.0">16 miles</quantity>
N.W. / of
<normalizedToken originalValue="San’a">
<collectingRegion country="Yemen" name="San'a'">San'a</collectingRegion>
<normalizedToken id="9D4B708F120033C11771FF439A88A41B" originalValue="San’a">
<collectingRegion id="B99147F8F35C73EAAFF38E3642E94B3B" country="Yemen" name="San'a'">San'a</collectingRegion>
</normalizedToken>
, ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">9,300ft.</elevation>
<quantity id="F188696F4162FFA38CF5B99D471D1587" metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">
<elevation id="5691CF37ECC0D8E069DD068F58CB79F1" metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">9,300ft.</elevation>
</quantity>
[=
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">2835 m</elevation>
<quantity id="2B6AB52C74B7534C4E8D70CCF5613BC5" metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">
<elevation id="D98AADB3E537F517D5725F9AFABC724F" metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">2835 m</elevation>
</quantity>
] /
<collectingDate value="1938-02-03">3.ii.1938</collectingDate>
<collectingDate id="D3F830DBE79DF770BD681521AB625C3D" value="1938-02-03">3.ii.1938</collectingDate>
.//
<collectorName>B.M. Exp.</collectorName>
<collectorName id="9381707295BEE83F4CAF60A03714F0B7">B.M. Exp.</collectorName>
to /
<collectorName>S.W. Arabia.</collectorName>
<collectorName id="DF510613F65866AF742604936C828014">S.W. Arabia.</collectorName>
/
<collectorName>H. Scott</collectorName>
<collectorName id="99F812D16BF3643195F5386701B60254">H. Scott</collectorName>
&amp; /
<collectorName>E.B. Britton.</collectorName>
<collectorName id="F4FE4B749CBDCC3790F10D9553589A20">E.B. Britton.</collectorName>
/ B.M. 1938-246, //
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="8D34C6279A6257256A2705D0DFB5F7FE" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
<taxonomicName id="190BBEF500F297211B697DDEB11AD7CC" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
/
<collectorName>Reiche</collectorName>
<collectorName id="E7D03581FEB4410F2C629371EABEAEFF">Reiche</collectorName>
/
<determinerName>E.B. Britton</determinerName>
<determinerName id="99E438DE862692A7035A6C1F75F1CF91">E.B. Britton</determinerName>
det. / 1946' (NHMUK)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="7F416094CF82CD27C52FAE5AEEA09C1C" pageId="0" pageNumber="71">
TME:
<specimenCount type="generic">2 specimens</specimenCount>
<specimenCount id="6CF7949C8494182953170400206750B2" type="generic">2 specimens</specimenCount>
. TGE:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="5DAED92502806DA4345E3CECF5A43384" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">Latinized patronym based on the surname of Ingo Brunk (Dresden, Germany), who collected the first well-preserved representative of the new species.</paragraph>
<subSubSection id="C9B6EAE2A170F4A6D205726BBD830CE7" pageId="0" pageNumber="71" type="etymology">
<paragraph id="033B1F841E1F8C96E7E07C9800DE4B9E" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="6D897BCF2543FB6C215123567970433F" pageId="0" pageNumber="71">Latinized patronym based on the surname of Ingo Brunk (Dresden, Germany), who collected the first well-preserved representative of the new species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunki</emphasis>
<subSubSection id="E61C8889D659B77EC38D5AE911C1A764" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="A53B3C620306195FE879D7ACBCF1AFE1" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="B9377375612234005D1B11FC74EF6BA1" pageId="0" pageNumber="71">
<taxonomicName id="A2AC75101974B65D0271822B20AECF04" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis id="5C83AD0E5AB6D40FC4BFFE0990FB2259" italics="true" pageId="0" pageNumber="71">Platyderus brunki</emphasis>
</taxonomicName>
sp. nov. readily differs from all other
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="3B574C0DC2EA31078CC017900B56E76A" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="9E25AFE420F428512BF6B167F30D7892" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
-taxa in its smallest value in ratio PW/PL (Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
<tableCitation id="74AFD47F8862DC5AE2E478A59F0CBC4D" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
). It is most closely related to
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="AA1669BC90B6D0D178344B25EBCB6D39" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="B6B83DBA332E276B85E8F1DF6B3F59B6" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
; both species are the only two within &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="43ACF063EF816521C633226FC3B26768" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="91097CFC095776D08B05B48496A95F06" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group with darkest coloration of integument and sides of pronotum not sinuate from widest point to base (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="83849AC37FB84B6AB9BD6AACD58E0E8F" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="C3E119084269F072B9A9E23436AC0263" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
has also sides of pronotum not sinuate to base, but much lighter coloration of integument and higher value in ratio EL/PL). The new species is distinct from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="B2554F1FFBB1801BCB89CF20E6B553D3" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="FEBEC51BD7D05E5521E2C953C1BC6432" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov., in: (1) pronotum with anterior angles more rounded at tips (vs. pronotum with anterior angles more pointed at tips, Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3B, C</figureCitation>
<figureCitation id="B6C4D1300B2262741A679C94EFB53994" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3B, C</figureCitation>
); (2) pronotum nearly as long as wide, PW/PL: 1.02-1.05 (vs. pronotum appreciably wider than long, PW/PL: 1.07-1.11); (3) pronotum subcordate, less constricted anteriorly, PW/PA: 1.38-1.42, with greater value in ratio PA/PB: 0.80-0.88 (vs. pronotum subquadrate, more constricted anteriorly, PW/PA: 1.44-1.48, with smaller value in ratio PA/PB: 0.76-0.79); (4) elytra compared to pronotum slightly shorter, EL/PL: 2.40-2.44 (vs. elytra compared to pronotum slightly longer, EL/PL: 2.47-2.56). Females are different from all other representatives of the &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="7F15FE88B52EB3E766E3597C9D3AB5B7" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="B25E456D7D85E87CCD37337819494F6B" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group by the presence of two, instead of one, dorsolateral ensiform setae on the apical gonocoxite (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
<figureCitation id="4D6F55E42F2EB21FA4F1900FF3CEE496" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="A42053743EC3926373B21B894DA16CEA" pageId="0" pageNumber="71" type="description">
<paragraph id="590A9A7E00303D9359AEB612CA33F186" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="7F81BBA8564B7F37D8097B02D321E3E7" pageId="0" pageNumber="71">
<emphasis id="FFC4CD173C06101A67E1C78A61284961" bold="true" pageId="0" pageNumber="71">
<emphasis id="DDCF49BA781D94359A62FE2E3FAE8C2A" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="FAB1C20C448EAE341FDF220342AE778A" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="FEB66EC360187972794F7BCD7F22E36F" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL of holotype: 9.50 mm; BW: 3.30 mm), with elongate, moderately convex body (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3C</figureCitation>
<figureCitation id="BB675E84F5ACC07BB99F6AAA9A37B134" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="47D5BD8718D8F2508DA9FB4B3BA090E1" bold="true" pageId="0" pageNumber="71">
<emphasis id="4D3CAC33A4640BAAAA33CFAA1E9144C2" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
<tableCitation id="FA433765B7A6BED3ED3B22A66B20E690" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="7B0454786CE415BEBF72C7D0EC2D74B2" bold="true" pageId="0" pageNumber="71">
<emphasis id="3BAE866FE8EBF06F6C305B2D78C1D83F" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly reddish-brown, with palpi and antennae slightly lighter. Integument slightly to moderately shiny, head and pronotum shinier than elytra.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="A6B4BAAAA72CF4413D9B6DEA3104C141" bold="true" pageId="0" pageNumber="71">
<emphasis id="C4082DF76557135460015E01D87D7286" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Microreticulation of pronotum and elytra as in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="DE5B896C17951B344EACB3ECB8E09A74" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="940E9746A8D5B99618CC7737627BA0B2" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. Ventral surface largely with well-impressed isodiametric or slight transverse sculpticells, epipleura, mesosternum and middle coxa with sculpticells less apparent. Head (holotype) nearly smooth, impunctate, wrinkles present only on clypeus and frons, including frontal foveae and area surrounding them. Pronotum surface largely smooth, only apical part medially in front of anterior transverse impression and basal area medially behind posterior transverse impression longitudinally wrinkled, as well as each side along lateral margin with a few shallow punctures not reaching anterior half. Elytral intervals without apparent punctation. Abdominal ventrites nearly smooth, impunctate, 3-5 with a few fine wrinkles at sides.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="CB3B591A7FB68068CB1F17D34C2D328E" bold="true" italics="true" pageId="0" pageNumber="71">Head</emphasis>
(holotype). About two-thirds as wide as pronotum (PW/HW of holotype: 1.36). Eyes long, slightly convex. Labrum subrectangular, as long as clypeus, with anterior margin concave. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows shallow, subfoveolate. Paraorbital sulci straight, backward barely reaching posterior margin of eye, not reaching level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="58949ED928EB6EBBCB9774C71347214A" bold="true" pageId="0" pageNumber="71">
<emphasis id="77B2937838F3EA7274A34588FBC4B2A3" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum almost as wide as long (PW/PL= 1.02-1.05), with widest point at second quarter (holotype) or medial fifth (specimen from &quot;El Errein, nr. Haz&quot;). Anterior and posterior transverse impressions slightly distinct medially, indistinct laterally. Sides not sinuate, smoothly convex anteriorly and medially, straight (specimen from &quot;El Errein, nr. Haz&quot;) or convex posteriorly (holotype); anterior bead present laterally, lacking in medial 1/8 to 1/10; basal bead present laterally, reduced to absent in medial half; posterior angles rounded (holotype) or obtuse (specimen from &quot;El Errein, nr. Haz&quot;). Metepisternum as long as wide, MA/MI= 1.00.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="CB80510955717E2862FA7983FB9EF5C7" bold="true" pageId="0" pageNumber="71">
<emphasis id="6D285A1BB35D8D0C2274CCF33383817A" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and a half times as long as elytra (EL/EW= 1.58-1.59), two and a half times as long as pronotum (EL/PL= 2.40-2.44), and a one and half as wide as pronotum (EW/PW= 1.48), with widest point at medial third. Parascutellar striole scarcely distinct, very shallow; striae more impressed than striole, barely punctate; parascutellar striole not joining stria 1; striae 1-5 and sometimes 6 reaching basal bead, 7 ending little before. Interval 3 with three discal setiferous punctures, medial one larger than others, in midst of interval 3 (holotype) or near stria 2 (specimen from &quot;El Errein, nr. Haz&quot;), anterior and posterior punctures smaller, adjoining stria 3 (posterior puncture lacking on right elytron of holotype). Umbilicate setiferous series with 16 or 17 punctures of each elytron.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="1465489AE9958CF6B62E1449AD5406D0" bold="true" pageId="0" pageNumber="71">
<emphasis id="8D4D34967AF1D07713BEC418EE6BEF0E" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Left mesofemur with 5, right one with 4 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="C4109A88AB27201C37A3C1A7849B37EB" bold="true" pageId="0" pageNumber="71">
<emphasis id="2DB31572FEFF39B2CFAB65B62CF00EF3" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Unknown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="79EF9DA28FACA28BB1187B4F2EBC2759" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
<figureCitation id="8B5635DC3BC24F9F83AE40ACD7941576" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="B15DDE8E00D8A72C276542EED704BDD8" pageId="0" pageNumber="71" type="habitat">
<paragraph id="50E3CE1A68BED9B65B5A3238EC7FA67D" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="EA44654BF4ABCECDB91217643E930616" pageId="0" pageNumber="71">
The holotype was collected by sifting the waste-water vegetation and soil at a humid place where waste water falls down (Ingo Brunk, pers. comm.). The correct altitude of the type locality is ca. 2700 m a. s. l. The locality is situated approximately half-way on a very steep walking path starting from Kawkaban (town on the top of the Mount Kawkaban) downwards to Schibam Kawkaban (town on the bottom of the mountain). Together with the holotype, numerous specimens of
<taxonomicName authorityName="Belousov" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus (Arabotrechus) rougemontiellus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="rougemontiellus" subGenus="Arabotrechus">Trechus (Arabotrechus) rougemontiellus</taxonomicName>
<taxonomicName id="C6EBA98C25791DE1D29B78A3B9C2DFB4" authorityName="Belousov" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus (Arabotrechus) rougemontiellus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="rougemontiellus" subGenus="Arabotrechus">Trechus (Arabotrechus) rougemontiellus</taxonomicName>
Belousov, 2017 and few
<taxonomicName class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Bembidion</emphasis>
<taxonomicName id="8BF19F0A1BAAE3AFFFCFCC3A0608205D" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="989544D5A05B78AB436CD4D41A541BA5" italics="true" pageId="0" pageNumber="71">Bembidion</emphasis>
</taxonomicName>
spp. were collected.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 13.</emphasis>
<caption id="50408317C1DC681E2043924A449599DA" doi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71" start="Figure 13" startId="F13">
<paragraph id="186159E279C956FA2199A62EDCEC0693" pageId="0" pageNumber="71">
<emphasis id="FD006047F6F587C19F3286417AC8DC2A" bold="true" pageId="0" pageNumber="71">Figure 13.</emphasis>
Spermathecal complex and gonocoxites, ventral view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">Platyderus (Eremoderus) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
<emphasis id="90E9A2B1F2F3A80079F1C0C8451235BC" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="319AA356741370D0F2232F3F53C9CDD1" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">Platyderus (Eremoderus) weiratheri</taxonomicName>
<normalizedToken id="666869EF0EF24B55279A8DD919AA1FBF" originalValue="Mařan">Maran</normalizedToken>
, 1940, topotype female specimen;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
<emphasis id="A698588B6022E4EDF6D8180138B67C76" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="E6D0EE7A91B1DAD4FEBA088227E7578F" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
<emphasis id="D40D3C2153B59BEF669D2F611DA0E709" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="46B61937022E53A98B5C50D1548D112A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
<emphasis id="2FB1B840AB56345FA8470F019295AC6E" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="6B6977BAAE76F811C8A93DDC22B0BB02" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
<emphasis id="3F6E029497FC236124CE9737A641CA4F" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="C11859DFDE13A0BF2CEFCB96F1B94EC4" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., female paratype, col de Buglan 1500 m, border area of
<normalizedToken originalValue="Bingöl">Bingoel</normalizedToken>
<normalizedToken id="8C3297C53F2DA6541D1D45863D4A0D90" originalValue="Bingöl">Bingoel</normalizedToken>
and
<normalizedToken originalValue="Muş">Mus</normalizedToken>
<normalizedToken id="862C4AD0C536DF16918DDF4C5829AD3E" originalValue="Muş">Mus</normalizedToken>
provinces, Turkey;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
<emphasis id="52F7182B51DAEA63C094B0B6B7145F8E" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="CE21516C59BE6FC4104C1473F4FE1253" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="71">ag</emphasis>
<emphasis id="7844FC20C2ABF780549D2B1093B28612" bold="true" pageId="0" pageNumber="71">ag</emphasis>
- apical gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="71">at</emphasis>
<emphasis id="AB357D7458224FB93904092B295424EE" bold="true" pageId="0" pageNumber="71">at</emphasis>
- atrium;
<emphasis bold="true" pageId="0" pageNumber="71">bc</emphasis>
<emphasis id="FF076DE33BA125144CD405945D76AEE9" bold="true" pageId="0" pageNumber="71">bc</emphasis>
- bursa copulatrix;
<emphasis bold="true" pageId="0" pageNumber="71">bg</emphasis>
<emphasis id="49646EF3B6D836F904FAFC7EDCCAC980" bold="true" pageId="0" pageNumber="71">bg</emphasis>
- basal gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="71">co</emphasis>
<emphasis id="D53ECA72289A33EEC5ABCD61FB2115F2" bold="true" pageId="0" pageNumber="71">co</emphasis>
- common oviduct;
<emphasis bold="true" pageId="0" pageNumber="71">rc</emphasis>
<emphasis id="DFD485FC261A99652A1DA5CCD5E31D19" bold="true" pageId="0" pageNumber="71">rc</emphasis>
- receptaculum;
<emphasis bold="true" pageId="0" pageNumber="71">sg</emphasis>
<emphasis id="7D3A8FC1AADE6F7FD2D17F7616D124F9" bold="true" pageId="0" pageNumber="71">sg</emphasis>
- spermathecal gland;
<emphasis bold="true" pageId="0" pageNumber="71">smc</emphasis>
<emphasis id="DEC34678D75960E5A275008C8DB0F27A" bold="true" pageId="0" pageNumber="71">smc</emphasis>
- seminal canal;
<emphasis bold="true" pageId="0" pageNumber="71">spc</emphasis>
<emphasis id="0B212AC1F581B5617AD9FD9142CBA3DD" bold="true" pageId="0" pageNumber="71">spc</emphasis>
- spermathecal canal.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="95225C4D59ED43D19C69B46D378002D7" pageId="0" pageNumber="71" type="distribution">
<paragraph id="C351A19D3F428694C42B786FC52DE77C" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="E7E91D35E8F29BA3053EEF5B6E38874B" pageId="0" pageNumber="71">
Yemen (
<normalizedToken originalValue="Sana’a">Sana'a</normalizedToken>
<normalizedToken id="480DCD26935B7025D09C49BB924E98F5" originalValue="Sana’a">Sana'a</normalizedToken>
Governorate), see Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
<figureCitation id="5DDF7F7631A71FDC24492FD7001DD606" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
.
</paragraph>
</subSubSection>

516
data/DE/34/C9/DE34C90D9A425CC69E158B25667916DA.xml
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@ -1,460 +1,460 @@
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15.1.
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<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3D, E</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7I</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9C</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11B</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12J</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15A</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
<subSubSection id="5BDFAE0F364737D8303894F0F5646736" pageId="0" pageNumber="71" type="description">
<paragraph id="CB287AB4C35B28ED58C7CCCB32758BD4" pageId="0" pageNumber="71">
<figureCitation id="8521A6262A533FB811EFDC1B97154FFE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3D, E</figureCitation>
<figureCitation id="F18DAD339C617C5F69528114471BECEA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6A</figureCitation>
<figureCitation id="C623B7B0D07726164414C3D7C4370A74" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7I</figureCitation>
<figureCitation id="B90ED59B47947AC7B932741311B7FB7E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9C</figureCitation>
<figureCitation id="BA6C3EAE426EB93803E46C61513D23B5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11B</figureCitation>
<figureCitation id="F3EAD6E32C6C2D9FEC55CA186137C27D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12J</figureCitation>
<figureCitation id="677107EAD294C899D00197895672C59E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15A</figureCitation>
<figureCitation id="8258F08B398F17BB3CCFC6B48A736D01" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="reference_group">
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
<subSubSection id="6FDE57287CF85D519B3D4ACAC3849333" pageId="0" pageNumber="71" type="reference_group">
<paragraph id="D2D580C56184F2F4D54BD715EE938080" pageId="0" pageNumber="71">
<taxonomicName id="D3D40A5B2ADD191140DF4A03FE5A48C9" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Karsch, 1881: 43 (type locality: &quot;Bir Milrha&quot;, Jabal
<normalizedToken originalValue="Tarhūnah">Tarhūnah</normalizedToken>
<normalizedToken id="7D77DCDB3552347AC7B4C58815DCB426" originalValue="Tarhūnah">Tarhūnah</normalizedToken>
[Tarhuna plateau], Libya, based on lectotype designation), stat. rev.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
<paragraph id="3DC83E3E03B6314E77B95928DB11B255" pageId="0" pageNumber="71">
<taxonomicName id="7E3AE012287095FEE3CE078302F6455F" authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
=
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
<taxonomicName id="5B66F0C4D8467A22D70F194AE395A589" authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
Bedel, 1900: 170 (type locality: &quot;Sud de la Tunisie&quot;), syn. n.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="44AF2AEA931BCC817DF0C70359188985" pageId="0" pageNumber="71" type="notes on type locality">
<paragraph id="5CB7938F48800B75F3828D209F68A7F5" pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph id="32BC5EFAF27D53E7124E23F60C38A58D" pageId="0" pageNumber="71">
This taxon was described from &quot;Bir Milrha&quot; (Murqub District, Libya; locality cannot be localized) and &quot;Uadi Mbellem&quot; (Misrata District, Libya; locality with approximate GPS coordinates: 31.167, 15.050) both localities in the
<normalizedToken originalValue="Gebèl">Gebel</normalizedToken>
<normalizedToken originalValue="Tarhúna">Tarhuna</normalizedToken>
<normalizedToken id="7A073DFC6E578ACEB7D4106DA05AE559" originalValue="Gebèl">Gebel</normalizedToken>
<normalizedToken id="0248C3020458A03C767A42234CCED436" originalValue="Tarhúna">Tarhuna</normalizedToken>
Plateaux. As a result of current lectotype designation, the former place becomes the type locality of
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
<taxonomicName id="2CE7C81E384C7A03367AB2EAEF660485" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis id="F03077F741E0298CB9925E1DC0A378FF" italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus elegans</emphasis>
<taxonomicName id="F8D9A4D578F6FD5AD48179B675B21725" authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="EF5D8B94A542BEFFC7A633B19C30F9CE" italics="true" pageId="0" pageNumber="71">Platyderus elegans</emphasis>
</taxonomicName>
was first discovered from &quot;Sud de la Tunisie&quot; (Bedel, 1900), and for the second time - from
<normalizedToken originalValue="“Aïn">&quot;Ain</normalizedToken>
<normalizedToken id="0555EFC738CD78A53B319F542A38C1F0" originalValue="“Aïn">&quot;Ain</normalizedToken>
Segoufta
<normalizedToken originalValue="à">a</normalizedToken>
<normalizedToken originalValue="l’O">l'O</normalizedToken>
<normalizedToken id="347525E073EB4D0F0ECDFFF3C5BCE120" originalValue="à">a</normalizedToken>
<normalizedToken id="C675C71D2BE0890BABD7C0C35D6834B6" originalValue="l’O">l'O</normalizedToken>
. du Dj. Bon-Hedma&quot; (Bedel, 1902); the second location refers to present Jebel Bou-Hedma National Park, situated on Gafsa Governorate and Sidi Bouzid Governorate, Tunisia.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 14.</emphasis>
<caption id="5CB341D192FFF6B7A2F6E6752D846E4C" doi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71" start="Figure 14" startId="F14">
<paragraph id="624F3204C4AC50E7BCAC4527B55F8DF7" pageId="0" pageNumber="71">
<emphasis id="B67158694A72E19E4BFEE53FBFD486F4" bold="true" pageId="0" pageNumber="71">Figure 14.</emphasis>
Spermathecal complex and gonocoxites, ventral view.
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">Platyderus (Eremoderus) lassallei</taxonomicName>
<emphasis id="6B05EDEFF4FC9D042D0061E3850B61AC" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="E0FEE1ADC07AEBE8B0C168418CD88388" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">Platyderus (Eremoderus) lassallei</taxonomicName>
, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol;
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
<emphasis id="72AC5EC5C26949AAABB16CAE6FA89BE1" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="166CCD43D8CE6C85889D55436045A8B9" class="Insecta" kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
<emphasis id="9BC68D93D6CFE234625166792E322AC5" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="DDED80E3E147CC7999469A7B7931CFA7" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran;
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Platyderus) umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus" subGenus="Platyderus">P. (Platyderus) umbratus</taxonomicName>
<emphasis id="16C66466F39769ECEAAF656DCB754A2F" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="F21A33861B952F08636E5EBE7F6DF0DB" class="Insecta" kingdom="Animalia" lsidName="P. (Platyderus) umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus" subGenus="Platyderus">P. (Platyderus) umbratus</taxonomicName>
(
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
<normalizedToken id="1F12B6C09289FC3245BA059F29DA7377" originalValue="Ménétriés">Menetries</normalizedToken>
), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht;
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus" subGenus="E.">P. (E.) arabicus</taxonomicName>
<emphasis id="18B1105B7E12EECEC56F57D16348C0D1" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="B6EC93A9A58473D37685744D1D1CB147" class="Insecta" kingdom="Animalia" lsidName="P. (E.) arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus" subGenus="E.">P. (E.) arabicus</taxonomicName>
, sp. nov., holotype;
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" subGenus="E.">P. (E.) brunki</taxonomicName>
<emphasis id="B692C1423CFE0F3C09AA09A69E6F6F35" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="B745295D54F5A084C8E67856FAA60CC4" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" subGenus="E.">P. (E.) brunki</taxonomicName>
, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13</figureCitation>
<figureCitation id="F4D4A16A63DB092CEE14F4338BB0B425" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13</figureCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus elegans</emphasis>
<subSubSection id="3E64B8BF71AE8B04082B86E14F93AC35" pageId="0" pageNumber="71" type="references">
<paragraph id="F82B7377CF607202FD60A1021744A8B1" pageId="0" pageNumber="71">References.</paragraph>
<paragraph id="5DC2FBFE6920D040D63D8A6B5A49A717" pageId="0" pageNumber="71">
<taxonomicName id="4B367D606906589C8573C20EE142F00A" authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="AC25DFD78FD3546C75AEC321572C6D9F" italics="true" pageId="0" pageNumber="71">Platyderus elegans</emphasis>
</taxonomicName>
:
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
<bibRefCitation id="2DEA1E085B3F6303FA64FD178F65F004" author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
: 211, 214-215;
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
<bibRefCitation id="4CBC05FD777C836659E14A78F4829770" author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
: 768;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
<bibRefCitation id="E34A4FA5CAEEA16D2183B2268CE34AE6" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="C0E88FB1E1333A1E552F6991EF3D4A07" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 522;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="8F617BDC00911356C493C33509D8B951" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 395;
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="9713FB6B0C3BE3DF915A24289A1FA770" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 757.
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
<taxonomicName id="E4B408ABD7CCFEFB70922CD51C51E2F5" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="46E0B2AD2D7430B4BC6BB0350DBF603A" italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
</taxonomicName>
(data for
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus brunneus</emphasis>
<taxonomicName id="DB4717CC81B23886B8E1D38F9A9E9E78" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">
<emphasis id="F051A80605E364F297B09347DDEC032C" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus brunneus</emphasis>
</taxonomicName>
Karsch):
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
<bibRefCitation id="38004FABAC49FD53DFF8748FF7E2C0D4" author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
: 211, 214-215;
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
<bibRefCitation id="68990B0AE5EC0D9ABCE0CA436D55A0A9" author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
: 769;
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
<bibRefCitation id="42B52CCAD02714151A3C205874D8DAF1" author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 521;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="60501A580B31BD58CFE2FCF776CFA486" author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 396;
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
<bibRefCitation id="FBEF45A08AC79C49EF1A1CC741225B9F" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 756.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
<subSubSection id="0377237DE43E381B8E3C5E026B469141" pageId="0" pageNumber="71" type="type material">
<paragraph id="ACF5B74DDD0BD86155F0312CE0DED917" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="50ED8AA02ABBAC67110FA36A4A8B4F1B" pageId="0" pageNumber="71">
<taxonomicName id="FD9905C06752E95A2E10B587A08198B4" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="82CE1438ACB8916BC98EC6DD97D98CA0" italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
</taxonomicName>
Karsch, 1881. The original type series of Karsch consisted of six syntypes, 4♂♂, 2♀♀, all stored in MFNB. Their examination revealed the presence of two diagnosable taxa. One male and one female specimen were found to represent the subspecies
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="315E4C47AE93FFCBBAC09C17E17366BD" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="74FDFEA6018C0393043C83BFA421D5B4" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
(see section &quot;Type material&quot; under
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
<taxonomicName id="21099FEDA6167BD109999AA03ED230F4" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="FA4887E2E705859A305655D9EDA4B32C" italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
). The other four specimens belong to
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
<taxonomicName id="EA28C5540E673E1EA8B9E8B8AFE40DD0" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis id="AC96448407BCD374E0D2C27EB9D3993D" italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
, the first of them designated lectotype and other three paralectotypes: ♂, '60906 [p, w] // Bir Milrha / Exp. Rohlfs. [h, tq] // Hist.-Coll. (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="86EABCD8B76079B25AB41052C3AA663D" class="Insecta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 /
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
<taxonomicName id="D4F45B65051F459A82874F8C5E3E3C26" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Bir Milrha / Zool. Mus. Berlin [p, w] // SYNTYPE /
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="A6B305D1014EE0BC917FDFB7CA4F8DE7" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="C472BF9A893CFB436FC09F373E68AFDB" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'; ♂, 'Uadi
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
<normalizedToken id="5AD22EBFC19849F63192A7822FB243AF" originalValue="M’bellem">M'bellem</normalizedToken>
/ Exp. Rohlfs. [h, tq] // Hist.-Coll. (
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="B0E31DD5B1FFA53C464950A625826C4E" authorityName="Stephens" authorityYear="1827" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 /
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
<taxonomicName id="AC9DEBCB6C322612986BA3771F9A01A5" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Uadi
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
<normalizedToken id="64633CE4E7E7F525FF3A7C2C210E592F" originalValue="M’bellem">M'bellem</normalizedToken>
, Exped. / Rohlfs / Zool. Mus. Berlin [p, w] // SYNTYPE /
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="C734CE757BA7D47A0EFE265F83FCD902" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="3CFF314F59A061EA8212C1F0A991A3D3" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'; ♂, 'gn 2469. [h, w] // Hist.-Coll. (
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="F1BF2BE3318D09B6D9687713D7F5F93E" authorityName="Stephens" authorityYear="1827" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 /
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
<taxonomicName id="E6FD32CB440F3AED1ED93A1D6B9BA82A" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Bir Milrha &amp; Uadi
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
<normalizedToken id="F8FB0B028F31C4063619EEC7DC2F1107" originalValue="M’bellem">M'bellem</normalizedToken>
, / Exped. Rohlfs / Zool. Mus. Berlin [p, w] // SYNTYPE /
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="BEA1C5493178C0BECDB60BB0DFA243D7" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="3A18D58F5EB1584D385A2D9F817D8461" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'; ♀, 'Hist.-Coll. (
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="056CE2BC0FD886B6C8C703223B02F3E5" authorityName="Stephens" authorityYear="1827" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 /
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
<taxonomicName id="568F57288AE0199A760578F4F350BF15" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Bir Milrha &amp; Uadi
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
<normalizedToken id="C64097DC4B76ED18452C0B4C178E86CB" originalValue="M’bellem">M'bellem</normalizedToken>
, / Exped. Rohlfs / Zool. Mus. Berlin [p, w] // SYNTYPE /
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="E665BA61422740BDF11152A90004D520" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
<taxonomicName id="A33EFFDB4761D5A463B1FB94354AA2BF" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="700CBA734C2C13169B23B936EFB9FBA5" pageId="0" pageNumber="71">
The synonymy of
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
<taxonomicName id="19F3C0E3BBC6488B287E111841DDBA1E" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="61DE8F96033E88F5B7744D9A68D38BC2" italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
</taxonomicName>
with
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="91D9DCB9A0C237ACF8B18C6EB44FB2E6" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="4635B65942A323AA5F9408A20788FC9E" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
was proposed by
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel (1902</bibRefCitation>
<bibRefCitation id="9EFD7233D85C447DB99BB40033FB409E" author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel (1902</bibRefCitation>
: 211). Revision of the type material and study of the original descriptions of the both aforementioned forms and that of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
<taxonomicName id="E9E1DAE74AC06BD920D448A1A887D8D4" class="Insecta" kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="1FC8375CA783AB024DD0088E968B7386" italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
</taxonomicName>
, as well the study of other
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
<taxonomicName id="0A9F5B14E65FFF82CF4D0E32C6211841" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="9E67D12810D65AE4983E7858633A05A1" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
-specimens from North Africa and the Levant, revealed that
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
<taxonomicName id="1C97D85746511882765BF355BA909655" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="6DE3716A0A33C1C4B2F4107C50F506D2" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
<taxonomicName id="38AD9CBC8924B5E6495AF289F3BD9C5D" class="Insecta" kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="067CC4A16664D84159C70254136762EF" italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
</taxonomicName>
represent a single species in contrast with
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="A0A84468CDDA16F43F1D5E45AB53B733" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="8970DC863523556E8DB91017D0C376C5" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
which is a separate species not occurring in North Africa.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Tunisia</emphasis>
<subSubSection id="E15AEA40EAC37F994278B50CDE3DE045" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="3774C169452AEB72510EECF9CEB1C731" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="0B8B79D21ED87F447F9AD357F9F5E1F5" pageId="0" pageNumber="71">
<emphasis id="B2BA35C296F9AD5E5C7DFAFC6A62A114" bold="true" pageId="0" pageNumber="71">Tunisia</emphasis>
:
<materialsCitation collectingDate="1998-04-22" collectorName="R. Borovec, Bedel" country="TUNISIA" location="Ben Gardane" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Medenine Governorate">
<collectingRegion country="Tunisia" name="Medenine">Medenine Governorate</collectingRegion>
<materialsCitation id="37FB1487F3B056D724728394C51C77CB" collectingDate="1998-04-22" collectorName="R. Borovec, Bedel" country="TUNISIA" location="Ben Gardane" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Medenine Governorate">
<collectingRegion id="3E1F9133C65D26522ADDE3C22C677F36" country="Tunisia" name="Medenine">Medenine Governorate</collectingRegion>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="2F5BB8F28F5BDACC1F65C9DFBA79AD26" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="0F5477462F2AC933E20CC0BCBEB13C26" type="female">1♀</specimenCount>
, '
<collectingCountry name="Tunisia">TUNISIA</collectingCountry>
<collectingCountry id="AA6E92945BF2551FE4EC8F148AEC0545" name="Tunisia">TUNISIA</collectingCountry>
c. or. /
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
<quantity id="0798DCE00F6122D980BB0E8254301E0F" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
W of
<location LSID="urn:lsid:plazi:treatment:DE34C90D9A425CC69E158B25667916DA:E650DDF8528D0FB64660AEC58F557611" country="TUNISIA" name="Ben Gardane" stateProvince="Medenine Governorate">Ben Gardane</location>
<location id="E650DDF8528D0FB64660AEC58F557611" LSID="urn:lsid:plazi:treatment:DE34C90D9A425CC69E158B25667916DA:E650DDF8528D0FB64660AEC58F557611" country="TUNISIA" name="Ben Gardane" stateProvince="Medenine Governorate">Ben Gardane</location>
/
<collectingDate value="1998-04-22">22.4.1998</collectingDate>
<collectingDate id="0EEACEE764E9230B34F39AEF7A8A3123" value="1998-04-22">22.4.1998</collectingDate>
/
<collectorName>R. Borovec</collectorName>
<collectorName id="4663289759315C30E81BB0B41D787D6F">R. Borovec</collectorName>
lgt. //
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="66F18AD8B0DE485B2026FBDF9D6D7AC1" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
<collectorName>Bedel</collectorName>
<taxonomicName id="085C0730D6402F83FC82B49951D38426" authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
<collectorName id="03C542E6C07101D8B3015E5DA00BB0A2">Bedel</collectorName>
/ JEANNE det. 2003 // COLL. WRASE /
<normalizedToken originalValue="BERLIN’">BERLIN'</normalizedToken>
<normalizedToken id="10A39357FE8BF5187075698B3D7D5285" originalValue="BERLIN’">BERLIN'</normalizedToken>
(cWR)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="2006-02-04" collectorName="Weill'" country="Libya" location="Nuqat al Khams District" specimenCount="1" specimenCount-female="1" stateProvince="Nuqat al Khams District">
<emphasis bold="true" pageId="0" pageNumber="71">
<collectingCountry name="Libya">Libya</collectingCountry>
<paragraph id="E7FD888FAAFDA43354EDC39351B8AC3A" pageId="0" pageNumber="71">
<materialsCitation id="CEE0B9951E7BB41F03452729A4E90AF8" collectingDate="2006-02-04" collectorName="Weill'" country="Libya" location="Nuqat al Khams District" specimenCount="1" specimenCount-female="1" stateProvince="Nuqat al Khams District">
<emphasis id="B90F9BD943C46C6E91C4B0BB7530147C" bold="true" pageId="0" pageNumber="71">
<collectingCountry id="DFF0F222293328035E52B9B6871FE1C3" name="Libya">Libya</collectingCountry>
</emphasis>
:
<collectingRegion country="Libya" name="An Nuqat al Khams">Nuqat al Khams District</collectingRegion>
<collectingRegion id="0ECD1F18EAEC6CA986270014BB84AB81" country="Libya" name="An Nuqat al Khams">Nuqat al Khams District</collectingRegion>
:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="8D6F4A187A803C811C239E1CCC316193" type="female">1♀</specimenCount>
, 'BUKAMASH [near Zuwarah City] /
<collectingDate value="2006-02-04">04-II-2006</collectingDate>
<collectingDate id="2FFD670748C6C6B8D12629C30501439D" value="2006-02-04">04-II-2006</collectingDate>
/ LIBYA
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<normalizedToken originalValue="Weill’">
<collectorName>Weill'</collectorName>
<taxonomicName id="459C98DE60C07EDC501E98B54FCEC3E4" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<normalizedToken id="7CE84363E244D8D5EDB2348CA79D58A9" originalValue="Weill’">
<collectorName id="3B61216B077B6D0CAE977BAE63289FE1">Weill'</collectorName>
</normalizedToken>
(cWEI)
</materialsCitation>
.
<materialsCitation collectingDate="2010-01-30" collectorName="Det., Weill" country="Libya" location="Jafara District" specimenCount="1" specimenCount-female="1" stateProvince="Jafara District">
<collectingRegion inferredBySuffix="true">Jafara District</collectingRegion>
<materialsCitation id="188E53C436BA229A6BD8B2D5D9C54319" collectingDate="2010-01-30" collectorName="Det., Weill" country="Libya" location="Jafara District" specimenCount="1" specimenCount-female="1" stateProvince="Jafara District">
<collectingRegion id="1D7C096B57E26580CCB7CDEF462165C8" inferredBySuffix="true">Jafara District</collectingRegion>
:
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="42C6B7592C3328D80C0A0EE1CD82AE64" type="female">1♀</specimenCount>
, 'AZIZIYAH
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<taxonomicName id="251CE3DE6CD6AB688F0442555CD87D8F" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
Weill /
<collectingDate value="2010-01-30">30-I-2010</collectingDate>
<collectingDate id="FCFF71F70762358435EA89C90994FEFC" value="2010-01-30">30-I-2010</collectingDate>
LIBYE //
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
<taxonomicName id="6EC77363B88E2B8AF33094D540CFE8D4" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
<taxonomicName id="A53EE938D4897EF62653466EAC81FBE3" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
. R &amp; S /
<collectorName>Det.</collectorName>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<collectorName>Weill</collectorName>
<collectorName id="D97918B32B70D254E7DA1C414539C886">Det.</collectorName>
<taxonomicName id="1138CEE9EA72CB177BB32E37844680F0" class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<collectorName id="AECB200EE0B0BB0509AEFFC6EBA9C36A">Weill</collectorName>
2010' (cWEI)
</materialsCitation>
.
<materialsCitation collectingDate="2002-03-02" collectorName="JC Ringenbach Leg." country="Libya" location="Tripoli District" specimenCount="1" specimenCount-male="1" stateProvince="Tripoli District">
<collectingRegion country="Libya" name="Tarabulus">Tripoli District</collectingRegion>
<materialsCitation id="E899C7C531A0144549147AEB8D523BB0" collectingDate="2002-03-02" collectorName="JC Ringenbach Leg." country="Libya" location="Tripoli District" specimenCount="1" specimenCount-male="1" stateProvince="Tripoli District">
<collectingRegion id="725B03EFAB317CE02ACD3F28430D0201" country="Libya" name="Tarabulus">Tripoli District</collectingRegion>
:
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="60F8A88B430A82E1C37E70D50918C84B" type="male">1♂</specimenCount>
, '
<collectingDate value="2002-03-02">02-III-2002</collectingDate>
<collectingDate id="38BD4BAE131EE0145CF7CC266B94B94E" value="2002-03-02">02-III-2002</collectingDate>
/ JANZUR / LIBYA - TRIPOLITANIA //
<collectorName>JC Ringenbach Leg.</collectorName>
<collectorName id="A2D707BBA7DA75E542DF14A4C3DA91DC">JC Ringenbach Leg.</collectorName>
/
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
<taxonomicName id="B95457EE5AE7859940BF60B6C2C4E85E" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="80DF606D6F20A6F28A097EEA5A378EAC" italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
</taxonomicName>
' (cWEI)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="4609C2F989598C36AD46F51C89F1A82D" pageId="0" pageNumber="71">
Three additional records from
<collectingCountry name="Libya">Libya</collectingCountry>
<collectingCountry id="073E93076D10CD53E89E8F1DA446AC55" name="Libya">Libya</collectingCountry>
that concern
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
<taxonomicName id="0A8B40D8AF98EF93EA271449804A26A6" authority="brunneus" class="Insecta" kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis id="4CD765B2499BF2695C03969EEA3AF345" italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
are pointed out for
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="579A527DAF1CD07C59A183F6A56D8AF1" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="41C3DD22553045FFD592D3C2B1C649AC" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
in the webpage http://jcringenbach.free.fr/website/beetles/carabidae/Platyderus_languidus.htm, namely: 'East of
<normalizedToken originalValue="Tajura’">Tajura'</normalizedToken>
<normalizedToken id="01C6B4A95B0ED4B88D4F5504DE9AF4F5" originalValue="Tajura’">Tajura'</normalizedToken>
,
<date value="2002-01-04">04/01/2002</date>
<date id="E748D5DD47E424A1F598C97E0C3AC51B" value="2002-01-04">04/01/2002</date>
, JCR Leg.'; 'Road Yafran-Az Zintan,
<date value="2001-12-25">25/12/2001</date>
<date id="788ADA1AAD5AFA1F7A3E9A0AD4ED18D4" value="2001-12-25">25/12/2001</date>
, JCR Leg.'; 'Sidi As
<normalizedToken originalValue="Sa’ih">Sa'ih</normalizedToken>
<date value="2006-02-03">03/02/2006</date>
<normalizedToken id="CFFD10A349A02801F23ADD6E65A390EA" originalValue="Sa’ih">Sa'ih</normalizedToken>
<date id="1A558ED7AB54DB6D1CE245A5023D6C74" value="2006-02-03">03/02/2006</date>
, PW Leg.'. The webpage is part of the database http://jcringenbach.free.fr/website/beetles/carabidae/carablibya.htm created by Jean-Claude Ringenbach (
<normalizedToken originalValue="Pardies-Piétat">Pardies-Pietat</normalizedToken>
<normalizedToken id="1B0F7A5FFA37CAFC8309AE6DF7C963E8" originalValue="Pardies-Piétat">Pardies-Pietat</normalizedToken>
,
<collectingCountry name="France">France</collectingCountry>
<collectingCountry id="0395139A81F4FB13668141B73A9AF06D" name="France">France</collectingCountry>
) and Patrick Weill (Pau,
<collectingCountry name="France">France</collectingCountry>
<collectingCountry id="389B66D978551CF9E0311A3023D42E05" name="France">France</collectingCountry>
) and representing online checklist of Libyan ground-beetles.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="7806363DA4473E23E66E97F369A9F16B" pageId="0" pageNumber="71">
TME:
<specimenCount type="generic">9 specimens</specimenCount>
<specimenCount id="CE93F409B919249AE9EE3A19F3E0391A" type="generic">9 specimens</specimenCount>
. TGE:
<specimenCount type="male">2♂♂</specimenCount>
<specimenCount id="DCFBE2C325359CC6665A7A06C67C71A8" type="male">2♂♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="E0202A5BC48A8F3506C13873E9E1887B" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="CCC3937F7382921FBFAEE73CBA221D62" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="D76EB3897D0E6658B83B86D992927F4D" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="7FDE65FC5E1B93B7DAE5A2E7CC8F5FD8" pageId="0" pageNumber="71">
Nominotypical subspecies differs from ssp.
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
<taxonomicName id="E41A301AE37458D63F9D775F30DE32D0" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis id="ECBC5B17B69730E2D6529F9F1D173253" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
in parascutellar striole and elytral striae 1-7 very shallow, almost indistinct, with bases not reaching basal bead (vs. parascutellar striole and elytral striae 1-7 distinct, slightly impressed, with bases reaching basal bead). In addition, the former has the pronotal anterior margin proportionally broader, compared to the posterior margin (vs. anterior margin of pronotum proportionally narrower, compared to posterior margin, see PA/PB, Table
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).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">Two of the three Libyan specimens were collected under stones or plant debris. Bukamash and Aziziyah are two biotopes of Tripolitania located in a semi-desert zone (Jaffarah Plain), with a few sparse trees and abundant ground vegetation, mainly in early spring (Patrick Weill, pers. comm.).</paragraph>
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<paragraph id="763A9FFD5F7FDD1DB8AE4AF87409BF94" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="2A56D478C2CFDB8340E66F065C518325" pageId="0" pageNumber="71">Two of the three Libyan specimens were collected under stones or plant debris. Bukamash and Aziziyah are two biotopes of Tripolitania located in a semi-desert zone (Jaffarah Plain), with a few sparse trees and abundant ground vegetation, mainly in early spring (Patrick Weill, pers. comm.).</paragraph>
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<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
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<paragraph id="DD7218C666A0BE056EEF2912ABBADC48" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="6EA2F6CB5286915C581FE0BC2354D3BB" pageId="0" pageNumber="71">
Tunisia, Libya (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
<figureCitation id="D5EC9DB1428AD1F61BAB004B96D37326" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
).
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<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
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<paragraph id="571FE6F899C81C44B41343A77F0C32BB" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="3DDA50C8072AF43003342449324AF5A4" pageId="0" pageNumber="71">
<bibRefCitation id="36CFE26EE1B9872E6A58B0C2701DEFBF" author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka (2017</bibRefCitation>
: 757) placed
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<emphasis italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
<taxonomicName id="EE8C22A6A04FA63CAB73C3B2D6E37477" class="Insecta" kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="EF7E258F40F9DB2A05718116853FFE17" italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
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in
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="D9ADAA83618507D770138F9BA9C70EDD" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
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(s. str.). Present study proved however that the species currently validly known as
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus P. brunneus</emphasis>
<taxonomicName id="371F76299F995EF23A07501DE43FE027" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="279C714FA68B4CD9D6FCC54E9AA92F89" italics="true" pageId="0" pageNumber="71">Platyderus P. brunneus</emphasis>
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is indeed a member of subgenus
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="(Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Eremoderus</taxonomicName>
<taxonomicName id="0B4032BB071227E5A3BE16FF787C8F35" class="Insecta" kingdom="Animalia" lsidName="(Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Eremoderus</taxonomicName>
.
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282
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17.
<taxonomicName LSID="http://zoobank.org/95E89306-626A-468D-BA68-F16489ABB9F3" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) irakensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<taxonomicName id="E4112E0CF492F61CA339EE26AC18FC84" LSID="http://zoobank.org/95E89306-626A-468D-BA68-F16489ABB9F3" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) irakensis</taxonomicName>
<taxonomicNameLabel id="21625B9AAEE70C42714932BAE911AA34" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4B</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6C</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15D</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
<subSubSection id="465AD97BAA664265E0BB803CEB1026CF" pageId="0" pageNumber="71" type="description">
<paragraph id="BC6170DB3CA790D38FF71BEE7FD4BFA8" pageId="0" pageNumber="71">
<figureCitation id="29C53515C5AB5054054B3D6D9D9942E8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4B</figureCitation>
<figureCitation id="667C57D771F485AC97CD149E1C81237E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6C</figureCitation>
<figureCitation id="A01A569C519E89C11F4C795BCD55EE6C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15D</figureCitation>
<figureCitation id="A81588F11A12216DDB4FEC33E1C46A8D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Iraq, Al Anbar Governorate, Ar Rutba District, ca. 115 km east of Ar-Rutbah Town.</paragraph>
<subSubSection id="0A211E92523452922724E4E20BC916A1" pageId="0" pageNumber="71" type="type locality">
<paragraph id="698953688EA00DD9616814168F77A57A" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="D67A7663EC33556FB489C66EB19B4064" pageId="0" pageNumber="71">Iraq, Al Anbar Governorate, Ar Rutba District, ca. 115 km east of Ar-Rutbah Town.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
<subSubSection id="678553EF50BF2DB955FC12827D996ADE" pageId="0" pageNumber="71" type="type material">
<paragraph id="F0C038F468DD8358F35D24090031E519" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="A74EAF94713797DD0DBD6665600C74C5" pageId="0" pageNumber="71">
<emphasis id="DD414B1198D1750EF4CA8937B04C0E25" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♀, 'IRAQ, Al-Anbar / Rutba, singled / 115 km. E of town / 8.I.1978 [w, h] // No. 350 /
<normalizedToken originalValue="Topál">Topal</normalizedToken>
<normalizedToken id="8D3289290655D56D6598CE4C7810C5A2" originalValue="Topál">Topal</normalizedToken>
&amp; Zilahy [w, h]' (HNHM); paratype ♀, 'IRAQ, Al-Anbar / Rutba, singled / at 30 km W of / town, 9.I.1978 [w, h] // No. 351 /
<normalizedToken originalValue="Topál">Topal</normalizedToken>
<normalizedToken id="DA3F61A1D021D2A0BE18E82D71F4E1F5" originalValue="Topál">Topal</normalizedToken>
&amp; Zilahy [w, h]' (NMNHS).
</paragraph>
<paragraph pageId="0" pageNumber="71">TME: 2 specimens. TGE: 1♀.</paragraph>
<paragraph id="6944EEDFEFA22911CA3F3412908BF33D" pageId="0" pageNumber="71">TME: 2 specimens. TGE: 1♀.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, referring to the name of the country in which this new species was found.</paragraph>
<subSubSection id="617F52C03D6618EFFC7D5FB14AC7CFAB" pageId="0" pageNumber="71" type="etymology">
<paragraph id="3CE51B53ED1B8F51873102A918EE1CA8" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="7108342DBB61E68AF15A777D2CD526FA" pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, referring to the name of the country in which this new species was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus irakensis</emphasis>
<subSubSection id="DB84E0B3A940937786BCB9A61843CA5A" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="E0BC7FC199737C9E979B4D126AE31DEA" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="DFCB230D97FA96DA92117D36E08EA0FA" pageId="0" pageNumber="71">
<taxonomicName id="804433C3D97D8B14C4C095379761DDC4" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="B6D7CDF8DAC38F9DE9DA1080113DEE39" italics="true" pageId="0" pageNumber="71">Platyderus irakensis</emphasis>
</taxonomicName>
is distinct from all other species of &quot;
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
<taxonomicName id="3AF36286D66FA9C65C944454CE73A970" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="15C833457F17BA99A147C9AE011B98B9" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group by the orange-brown color of integument (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
<figureCitation id="D8600DC906CA1C41933553F99776C99F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
), mesotarsomeres and metatarsomeres dorsally flattened and slightly grooved (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6C</figureCitation>
<figureCitation id="D730B85777CDCA0BD3613201B0D13C64" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6C</figureCitation>
), pronotum with sides to base slightly convex and posterior angles laterally not prominent, and distal enlargement of bursa copulatrix (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
<figureCitation id="2BF97CDBCD9DADA70DA5DACE9B7AE01C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
); the last character may be an autapomorphy. Among Asian representatives of species group, it is the form with the highest mean values for EL/PL (= 2.69) and EL/EW (= 1.63), and, together with
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="32784EE0CD294C907FAC63E3FBC18B79" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="ECC97239E52B255954EAAE8D37F6B003" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov., the one possessing lowest index for PW/PB (= 1.10).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
<subSubSection id="A874EDF4BEE9E4940C18ECA211DE1E7B" pageId="0" pageNumber="71" type="description">
<paragraph id="B8FC0AD9E057F59725693AE186F076B4" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="3793E59906F5E06FC5A8A6693F30A0C8" pageId="0" pageNumber="71">
<emphasis id="65CBF4F76681606F8C1BA4FCF955241A" bold="true" pageId="0" pageNumber="71">
<emphasis id="F06C283F9CA6C8292F58A8CDC9644B52" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of moderate size for
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
<taxonomicName id="1C8972CBDB4CDD4F0EC5EAB9498785E7" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="DF41C6AE90981F97C3A16C2556C9502F" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 8.30-9.00 mm; BW: 2.90-3.20 mm), with elongate and fairly convex body (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
<figureCitation id="4EE4BE8E93BFD76B5412FBF39708EE02" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
<emphasis id="EEE27C7779F685ACD9A00D97682E159F" bold="true" pageId="0" pageNumber="71">
<emphasis id="E09AD9D4236B50FE1DEC67755F5C6135" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
<tableCitation id="A1C3FAC50812411B85D248BCF5BC904B" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
<emphasis id="3DAE9103AB93A296FAD8AA952D16B3F4" bold="true" pageId="0" pageNumber="71">
<emphasis id="BF81F19B2389E453602BFF29A1017226" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages orange-brown, with head somewhat darker than the rest of the body and ventral surface lighter than dorsal surface. Integument slightly to moderately shiny, head and pronotum as shiny as elytra, ventral surface somewhat shinier than dorsal surface.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
<emphasis id="2EE59135DDC7FD1464DB79E5A373F5C8" bold="true" pageId="0" pageNumber="71">
<emphasis id="F5180657761EB2D636FB458A62E37F4A" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum with evident microreticulation, sculpticells regular isodiametric. Elytral intervals, scutellum and lateral gutter with distinct isodiametric sculpticells, sculpticells of basal margin more or less reduced. Ventral surface with isodiametric or slight transverse sculpticells, less apparent on epipleura and middle coxa. Head impunctate and smooth, only frontal furrows and lateral ends of clypeus, with very fine and short wrinkles. Pronotum surface mostly smooth, only apical part medially in front of anterior transverse impression and basal area medially behind posterior transverse impression longitudinally wrinkled; basal foveae and adjacent lateral areas shallowly punctate. Elytral intervals without evident punctures. Abdominal ventrites 1-3 finely wrinkled at sides, smooth medially.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
<emphasis id="8DB26D5B128ED40CB4794CE3F35663E5" bold="true" pageId="0" pageNumber="71">
<emphasis id="D37F0E030BC3B0EAB7D64325170E2BFB" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About one time and a third as wide as pronotum (PW/HW= 1.30-1.32). Eyes long, moderately convex. Labrum subrectangular, barely shorter than clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, reduced at sides. Frontal furrows punctiform, shallow. Paraorbital sulci straight, fine, backward surpassing posterior margin of eye, not reaching level of posterior supraorbital pore.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
<emphasis id="0F3A6EE49F48E95107FD6964CEC06B1D" bold="true" pageId="0" pageNumber="71">
<emphasis id="2B1A971807F0F21443F426890C7CDEB1" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about a tenth wider than long (PW/PL= 1.13), with widest point at medial third. Anterior transverse impression indistinct; posterior transverse impression indistinct to barely distinct. Sides not sinuate, shallowly convex anteriorly, barely convex posteriorly; anterior bead present laterally, lacking or present in medial ninth to tenth; basal bead present laterally, reduced to absent in medial third. Metepisternum slightly longer than wide, MA/MI= 0.91-0.95.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
<emphasis id="136A1AE4A0388A9DDC22E7C95FB25A47" bold="true" pageId="0" pageNumber="71">
<emphasis id="59B3B44CA1B5BA33CC5CE78213F90B0A" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and two thirds as long as wide (EL/EW= 1.62-1.63), two and two thirds as long as pronotum (EL/PL= 2.65-2.72), and almost one and a half times as wide as pronotum (EW/PW= 1.45-1.47), with widest point at first half of third quarter. Parascutellar striole and striae 1-8 shallowly impressed, superficially to indistinctly punctate; striole short, not joining stria 1; bases of striae not reaching basal bead (except for stria 2 in paratype). Interval 3 with three discal setiferous punctures (posterior puncture lacking on left elytron of holotype), all adjoining stria 2; one additional discal puncture adjoining stria 4 at anterior quarter of right elytron of paratype. Umbilicate setiferous series with 16 punctures on left elytron and 17 on right elytron in holotype, respectively, with 16 and 15 punctures in paratype.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
<emphasis id="A51D94C2ABD3D262530668F5078DA5F5" bold="true" pageId="0" pageNumber="71">
<emphasis id="581451617B75624752E3AF54719B2697" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one or two setae in basal third and one in medial third. Mesofemur mostly with 4 or 5 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with a few long setae, one in basal third and one-two in apical half.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
<emphasis id="185E476C2436FCC266757707EBDFDEA3" bold="true" pageId="0" pageNumber="71">
<emphasis id="94289AF9AB5F31D3ED0000355626B899" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Unknown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
<emphasis id="F3B77E091BD521A1828AC9D995A8BA39" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
<figureCitation id="1199D535025DE4E745A1E841280ED1FC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Bursa copulatrix two-chambered, with a shorter, but wider basal part and a longer, but narrower apical enlargement. Spermathecal canal connected in basal third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="23D5A6A598F79EDAD9DF32810EA017E4" pageId="0" pageNumber="71" type="comparisons">
<paragraph id="26D5ADBF8ECBEDD9629DDF1BF38BF5DD" pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph id="3C7D2B011FDF0D81F0D862323B9825FA" pageId="0" pageNumber="71">
In addition to characters mentioned in
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
<normalizedToken id="8ED3017C9B4ADD3A66E84F303AE46475" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
, the new species differs from
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
<taxonomicName id="B95645334534393F864E514F1C0997C7" class="Insecta" kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="1B2809E79BB636A6E281070F6A8D362F" italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov., by: (1) head wider, compared to pronotum (PW/HW: 1.30-1.32, vs. PW/HW: 1.35-1.42); (2) pronotum with sides less constricted anteriorly and posteriorly (PW/PA: 1.34-1.38 and PW/PB: 1.09-1.11, vs. PW/PA: 1.39-1.52 and PW/PB: 1.15-1.22); (3) elytra longer compared to their width and the length of the pronotum (EL/EW: 1.62-1.63 and EL/PL: 2.65-2.72, vs. EL/EW: 1.56-1.60 and EL/PL: 2.41-2.60).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<paragraph id="B8901E7F04899D0D2AB0DA119C770E57" pageId="0" pageNumber="71">
For differences between
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="0DDA9139A917FF18BCB9A851F7E8AD91" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="AAAA4A03A2962E6510368A78027DFD57" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
<taxonomicName id="BB53620A2F9559B4D5BCB97C412DA7AE" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="EDE7CB25D5B34E9B6C68A51D2084278C" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. and such between
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
<taxonomicName id="75687745B76E9CF9ED74D7994D210A08" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="F8FB7F833C7EA3A347FED0F8A0F44BDE" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
<taxonomicName id="A8BBB69C4087893F0766E73F4C13D301" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="065F1E20E294702C9EB086AAA9E12BEA" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
, see
<normalizedToken originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
<normalizedToken id="58DDD8AB9DF29E9D3F68423A828BD207" originalValue="“Comparisons”">&quot;Comparisons&quot;</normalizedToken>
under the latter species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">Nothing is known about the bionomics of this species.</paragraph>
<subSubSection id="3ED63E3CE282185B862FF9134883C411" pageId="0" pageNumber="71" type="habitat">
<paragraph id="896A593812174DD8E72CA93FA8F70956" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="079B4F48D946D69A2B2D6A472BCF4E75" pageId="0" pageNumber="71">Nothing is known about the bionomics of this species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<subSubSection id="120463BBC54C65B991CD14F12530480F" pageId="0" pageNumber="71" type="distribution">
<paragraph id="482638913415734BAE8097A09A23F31D" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="FF0F012F4829C8D882167533A593F5F8" pageId="0" pageNumber="71">
It is currently known from two localities situated in the Ar Rutba District, which is the largest district of Al Anbar Governorate, Western Iraq (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
<figureCitation id="3CD49227915598816927E24DA782003D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
). The holotype was collected about 115 km east of Ar-Rutbah Town whilst the paratype was found about 30 km west of the same town. Most of the area of the Ar Rutba District is a high plateau.
</paragraph>
</subSubSection>

128
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Genus
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<paragraph pageId="0" pageNumber="71">Type species.</paragraph>
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Audinet-Serville, 1821
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