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<mods:title id="30156D45ECCDC640DD58DD3473617AEA">A review of Crassignatha (Araneae, Symphytognathidae)</mods:title>
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<mods:namePart>Lin, Yucheng</mods:namePart>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<treatment ID-GBIF-Taxon="168821878" LSID="urn:lsid:zoobank.org:act:0FF6F414-70C7-4404-9352-7AE564CA4971" httpUri="http://treatment.plazi.org/id/C75BACDD6F6458F79DFDC80D68C860DA" lastPageNumber="63" pageId="0" pageNumber="63">
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<taxonomicName LSID="C75BACDD-6F64-58F7-9DFD-C80D68C860DA" authority="Y. Lin &amp; S. Li" authorityName="Y. Lin &amp; S. Li" authorityYear="2020" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha gucheng" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="gucheng" status="sp. nov.">Crassignatha gucheng Y. Lin &amp; S. Li</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Crassignatha gucheng sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure9" httpUri="https://binary.pensoft.net/fig/472594" pageId="0" pageNumber="63">Figs 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">, 10</figureCitation>
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">, 38</figureCitation>
<treatment id="C75BACDD6F6458F79DFDC80D68C860DA" ID-GBIF-Taxon="168821878" LSID="urn:lsid:zoobank.org:act:0FF6F414-70C7-4404-9352-7AE564CA4971" httpUri="http://treatment.plazi.org/id/C75BACDD6F6458F79DFDC80D68C860DA" lastPageNumber="63" pageId="0" pageNumber="63" scope_family="Symphytognathidae" scope_order="Araneae">
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<taxonomicName id="561417F8DBF7F255DF9DF830F3C776B2" ID-CoL="98ZDX" LSID="C75BACDD-6F64-58F7-9DFD-C80D68C860DA" authority="Y. Lin &amp; S. Li" authorityName="Y. Lin &amp; S. Li" authorityYear="2020" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha gucheng" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="gucheng" status="sp. nov.">Crassignatha gucheng Y. Lin &amp; S. Li</taxonomicName>
<taxonomicNameLabel id="A6F5A5A3AF9F88E08228304386882C1D" pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
<figureCitation id="A3D3B5FB4DCF9DFD44461FFC90C38310" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Crassignatha gucheng sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure9" httpUri="https://binary.pensoft.net/fig/472594" pageId="0" pageNumber="63">Figs 9</figureCitation>
<figureCitation id="E4C5DE28292043E1FA918381A565D536" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">, 10</figureCitation>
<figureCitation id="BA35E6BCE45CA25B394B685CA2408014" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">, 38</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="63">Type material.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
<subSubSection id="FC3F7B871F1F7385F1EDF8FEFC9B5F28" pageId="0" pageNumber="63" type="type material">
<paragraph id="511F3C522DA00A8E91A576E104A196E6" pageId="0" pageNumber="63">Type material.</paragraph>
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<emphasis id="8EA4929AD4373C1FAD3FA7390DC541F3" bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
♂ (NHMSU Ar 018) and
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<emphasis id="FCEB8067AF1E01CA11AD894A00AA3E15" bold="true" italics="true" pageId="0" pageNumber="63">paratypes</emphasis>
2♂ 5♀ (NHMSU Ar 019-025),
<emphasis bold="true" pageId="0" pageNumber="63">China</emphasis>
<emphasis id="014A7E7FD1747B808263AF8E540E981E" bold="true" pageId="0" pageNumber="63">China</emphasis>
: Yunnan Province, Longling County, Longjiang Township, Xiaoheishan Nature Reserve, Gucheng Mountain, in good forest (
<geoCoordinate degrees="24.82886" direction="north" orientation="latitude" precision="1" value="24.82886">24.82886°N</geoCoordinate>
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,
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<geoCoordinate id="7D355291819DA4CB4939E82819E5DDFB" degrees="98.75917" direction="east" orientation="longitude" precision="1" value="98.75917">98.75917°E</geoCoordinate>
; 2010 m), 22.VIII.2018, Y. Lin et al. leg.; 1♀ (NHMSU-HA132) used for sequencing, GenBank: MT992014, same data as for preceding.
</paragraph>
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<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 9.</emphasis>
<taxonomicName authorityName="Y. Lin &amp; S. Li" authorityYear="2020" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha gucheng" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="gucheng">
<emphasis italics="true" pageId="0" pageNumber="63">Crassignatha gucheng</emphasis>
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<emphasis id="569D84C98D5A84E8985FE18E520969C0" bold="true" pageId="0" pageNumber="63">Figure 9.</emphasis>
<taxonomicName id="E2D8AFFEEE0C9264BE2F9C5D5CB5E7AC" authorityName="Y. Lin &amp; S. Li" authorityYear="2020" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha gucheng" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="gucheng">
<emphasis id="86E9AFB117D9C01EF921CDD6DC746181" italics="true" pageId="0" pageNumber="63">Crassignatha gucheng</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
<emphasis id="A95D7ECE66B742AF7826962E28C12262" bold="true" pageId="0" pageNumber="63">A</emphasis>
male habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
<emphasis id="047437A492C02AE93E124C5D06CE2E06" bold="true" pageId="0" pageNumber="63">B</emphasis>
male habitus, ventral
<emphasis bold="true" pageId="0" pageNumber="63">C</emphasis>
<emphasis id="CFE8C8E72F3E8038330AC1EF04F83FDA" bold="true" pageId="0" pageNumber="63">C</emphasis>
male habitus, lateral
<emphasis bold="true" pageId="0" pageNumber="63">D</emphasis>
<emphasis id="39BEB1A10D32E9191BD1007877CD6C7C" bold="true" pageId="0" pageNumber="63">D</emphasis>
female habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="63">E</emphasis>
<emphasis id="BCFFBDC8D01F88F584B77633376CE1CE" bold="true" pageId="0" pageNumber="63">E</emphasis>
female habitus, ventral
<emphasis bold="true" pageId="0" pageNumber="63">F</emphasis>
<emphasis id="44B80072FDFA862B47DF6FF2B4D6219F" bold="true" pageId="0" pageNumber="63">F</emphasis>
female habitus, lateral. Scale bars: 0.50 mm (
<emphasis bold="true" pageId="0" pageNumber="63">A-F</emphasis>
<emphasis id="34D0A8F4021A35D64E441F161364D29A" bold="true" pageId="0" pageNumber="63">A-F</emphasis>
).
</paragraph>
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<paragraph pageId="0" pageNumber="63">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="63">
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<paragraph id="F31263B0AF29310521EA219A0DD5B933" pageId="0" pageNumber="63">Diagnosis.</paragraph>
<paragraph id="50E90C5134269F8B8C397B22F8005811" pageId="0" pageNumber="63">
The male of
<taxonomicName lsidName="C. gucheng" pageId="0" pageNumber="63" rank="species" species="gucheng">
<emphasis italics="true" pageId="0" pageNumber="63">C. gucheng</emphasis>
<taxonomicName id="01B48F4E811DF5E10432D2C5D29B020A" lsidName="C. gucheng" pageId="0" pageNumber="63" rank="species" species="gucheng">
<emphasis id="4798A6A74EF2E0089AA3190CAD9AEC2B" italics="true" pageId="0" pageNumber="63">C. gucheng</emphasis>
</taxonomicName>
sp. nov. is similar to that of
<taxonomicName lsidName="C. pianma" pageId="0" pageNumber="63" rank="species" species="pianma">
<emphasis italics="true" pageId="0" pageNumber="63">C. pianma</emphasis>
<taxonomicName id="67A09B0823CFB4C104B39C66DD2F9448" lsidName="C. pianma" pageId="0" pageNumber="63" rank="species" species="pianma">
<emphasis id="C5F2A7B292007F5B4428BE23FFAA4389" italics="true" pageId="0" pageNumber="63">C. pianma</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. quanqu" pageId="0" pageNumber="63" rank="species" species="quanqu">
<emphasis italics="true" pageId="0" pageNumber="63">C. quanqu</emphasis>
<taxonomicName id="C06DF74545BEC38F21368ED0CE35D552" lsidName="C. quanqu" pageId="0" pageNumber="63" rank="species" species="quanqu">
<emphasis id="51A91B9F55C0B7BF8A6F99C8F612514B" italics="true" pageId="0" pageNumber="63">C. quanqu</emphasis>
</taxonomicName>
but can be distinguished by the narrower embolic base and spine-like cymbial tooth vs. a wider embolic base and spur-like cymbial tooth in the latter two (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10A</figureCitation>
<figureCitation id="80FECE296BDF482D3CB019EFDB465F65" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10A</figureCitation>
). The female of
<taxonomicName lsidName="C. gucheng" pageId="0" pageNumber="63" rank="species" species="gucheng">
<emphasis italics="true" pageId="0" pageNumber="63">C. gucheng</emphasis>
<taxonomicName id="553918EFF8A3F3272DC74EFA5B65C507" lsidName="C. gucheng" pageId="0" pageNumber="63" rank="species" species="gucheng">
<emphasis id="1CAB1C54ED5CEA166F3E420395B01528" italics="true" pageId="0" pageNumber="63">C. gucheng</emphasis>
</taxonomicName>
sp. nov. differs from that of
<taxonomicName lsidName="C. dongnai" pageId="0" pageNumber="63" rank="species" species="dongnai">
<emphasis italics="true" pageId="0" pageNumber="63">C. dongnai</emphasis>
<taxonomicName id="04833001A33D99ADC426C3FA06B39152" lsidName="C. dongnai" pageId="0" pageNumber="63" rank="species" species="dongnai">
<emphasis id="13574882FC3B81D87CE6D8BF17D29484" italics="true" pageId="0" pageNumber="63">C. dongnai</emphasis>
</taxonomicName>
sp. nov. by the copulatory ducts twisted twice at the center of vulva, only once in the latter (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10F, G</figureCitation>
<figureCitation id="ED736CAB152E31F65C1EE4370BC2DCD8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10F, G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="description">
<paragraph pageId="0" pageNumber="63">Description.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Male</emphasis>
<subSubSection id="8C8F436AF7458F5A80FC5A802EADE3F3" pageId="0" pageNumber="63" type="description">
<paragraph id="5A02A12B4EFAD18B42C279904F3CD20E" pageId="0" pageNumber="63">Description.</paragraph>
<paragraph id="394BD6255032212914EBF579E5F8FABE" pageId="0" pageNumber="63">
<emphasis id="A6A3C9CBDF7C440FD2E108C507644154" bold="true" pageId="0" pageNumber="63">Male</emphasis>
(holotype). Total length 0.72. Carapace 0.32 long, 0.36 wide, 0.40 high. Clypeus 0.12 high. Sternum 0.24 long, 0.24 wide. Abdomen 0.44 long, 0.44 wide, 0.44 high. Length of legs: I 1.28 (0.38, 0.14, 0.32, 0.20, 0.24); II 0.98 (0.26, 0.14, 0.26, 0.14, 0.18); III 0.78 (0.20, 0.10, 0.16, 0.12, 0.20); IV 0.94 (0.30, 0.12, 0.18, 0.14, 0.20).
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Somatic characters</emphasis>
<paragraph id="C026C23C377035CC3198F7B981F944B0" pageId="0" pageNumber="63">
<emphasis id="77DFC1B487B4F7C1FB789FAC1A353319" bold="true" pageId="0" pageNumber="63">Somatic characters</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Crassignatha gucheng sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure9" httpUri="https://binary.pensoft.net/fig/472594" pageId="0" pageNumber="63">9A-C</figureCitation>
<figureCitation id="C2E3253590457736D2581CA1280A68A5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Crassignatha gucheng sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure9" httpUri="https://binary.pensoft.net/fig/472594" pageId="0" pageNumber="63">9A-C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Coloration</emphasis>
<emphasis id="42C2BF91270512213073E395382808E7" bold="true" italics="true" pageId="0" pageNumber="63">Coloration</emphasis>
: carapace pale brown, marginally darker. Mouthparts and sternum pale brown. Legs pale yellow, with faint pigmentation on each segment distally. Abdomen pale dorsally, brown-yellow laterally and ventrally, dark posteriorly.
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Prosoma</emphasis>
<emphasis id="052FF7B1914D49BF235A69017A69CCEF" bold="true" italics="true" pageId="0" pageNumber="63">Prosoma</emphasis>
: carapace nearly rounded, cephalic area elevated, cervical groove distinct, thoracic area and clypeus sculptured, clypeus concave. ALE protruded, PER recurved. Chelicerae bears setae anteriorly. Labium semicircular. Sternum heart shaped, slightly swollen, surface rough, truncated posteriorly.
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Legs</emphasis>
<emphasis id="F8824BEE596E0D6C4799B16148D0C13F" bold="true" italics="true" pageId="0" pageNumber="63">Legs</emphasis>
: with dorsal seta on each patella distally, with two setae on each tibia subproximally. Tibia II with two clasping spurs.
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Abdomen</emphasis>
<emphasis id="7B95025C954E19CE2F5357DDDC7F5F91" bold="true" italics="true" pageId="0" pageNumber="63">Abdomen</emphasis>
: anteriorly rounded, posteriorly subquadrate, lateral scutum inconspicuous. Spinnerets slightly sclerotized, lack circular plate.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Palp</emphasis>
<paragraph id="B102BBAEE049DE560DD23619F43D3ACC" pageId="0" pageNumber="63">
<emphasis id="C0BED0A80C0643140377792D2DD55490" bold="true" italics="true" pageId="0" pageNumber="63">Palp</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10A-C</figureCitation>
<figureCitation id="5386EBE01D3B548B7E61B2E2FBA1BB35" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10A-C</figureCitation>
): relatively large, no less than 1/3 carapace size. A few setae on cymbium distally. Apical cymbial tooth caniniform. Tegulum smooth, broad, globular. Plate-like median apophysis with a distal sclerotized process. Embolic membrane slender, laminar, translucent, arises from base of embolus. Embolus short and rigid, terminus blunt.
</paragraph>
<caption doi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 10.</emphasis>
<taxonomicName authorityName="Y. Lin &amp; S. Li" authorityYear="2020" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha gucheng" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="gucheng">
<emphasis italics="true" pageId="0" pageNumber="63">Crassignatha gucheng</emphasis>
<caption id="AC49CAFE7F97AAE099B47369F5A60D55" doi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63" start="Figure 10" startId="F10">
<paragraph id="5C680C01D8411D4A5F96659EC536ADC5" pageId="0" pageNumber="63">
<emphasis id="DBD9043EFBB42CDDC70AF1DB0B28C166" bold="true" pageId="0" pageNumber="63">Figure 10.</emphasis>
<taxonomicName id="9B7E7D14A543249969884D6D83348825" authorityName="Y. Lin &amp; S. Li" authorityYear="2020" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha gucheng" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="gucheng">
<emphasis id="62E124C2B69FBAAC4C14DB5E27277E94" italics="true" pageId="0" pageNumber="63">Crassignatha gucheng</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
<emphasis id="5B66E7BBA6AA02591FAF09C490E83872" bold="true" pageId="0" pageNumber="63">A</emphasis>
male palp, prolateral
<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
<emphasis id="3E48D5A0E3A77B6236B7BF55CC2B64B0" bold="true" pageId="0" pageNumber="63">B</emphasis>
male palp, ventral
<emphasis bold="true" pageId="0" pageNumber="63">C</emphasis>
<emphasis id="15BDC14589820D1277D8AAEA6065AB1A" bold="true" pageId="0" pageNumber="63">C</emphasis>
male palp, retrolateral
<emphasis bold="true" pageId="0" pageNumber="63">D</emphasis>
<emphasis id="D2C1EB7725CC5C2E0A73B1ACA755457B" bold="true" pageId="0" pageNumber="63">D</emphasis>
epigyne, ventral
<emphasis bold="true" pageId="0" pageNumber="63">E</emphasis>
<emphasis id="EF31E5F1AD2D28A2D3172CEEB3A2FF1C" bold="true" pageId="0" pageNumber="63">E</emphasis>
epigyne, lateral
<emphasis bold="true" pageId="0" pageNumber="63">F</emphasis>
<emphasis id="17A2D77D365E596EE3D43D265B55AAF5" bold="true" pageId="0" pageNumber="63">F</emphasis>
vulva, ventral
<emphasis bold="true" pageId="0" pageNumber="63">G</emphasis>
<emphasis id="33B050D28B778A48064767FF99F35765" bold="true" pageId="0" pageNumber="63">G</emphasis>
vulva, dorsal. Scale bars: 0.10 mm (
<emphasis bold="true" pageId="0" pageNumber="63">A-G</emphasis>
<emphasis id="9280CBF63BF7E61468CA2674F1425AA9" bold="true" pageId="0" pageNumber="63">A-G</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Female</emphasis>
<paragraph id="22AA983BFD349B9A6D89F42786FBB994" pageId="0" pageNumber="63">
<emphasis id="A5D765337216C414F76FA5687CFB584E" bold="true" pageId="0" pageNumber="63">Female</emphasis>
(one of the paratypes). Total length 1.08. Carapace 0.44 long, 0.40 wide, 0.40 high. Clypeus 0.12 high. Sternum 0.28 long, 0.24 wide. Abdomen 0.72 long, 0.72 wide, 0.84 high. Length of legs: I 1.48 (0.56, 0.16, 0.40, 0.14, 0.22); II 1.20 (0.38, 0.16, 0.34, 0.12, 0.20); III 0.84 (0.28, 0.14, 0.18, 0.10, 0.14); IV 1.1 (0.38, 0.14, 0.28, 0.18, 0.12).
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Somatic characters</emphasis>
<paragraph id="C1B0D714E655257B562A459C68C1DB14" pageId="0" pageNumber="63">
<emphasis id="A8F78F77248FD4B11FC983F0DE331B4B" bold="true" pageId="0" pageNumber="63">Somatic characters</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Crassignatha gucheng sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure9" httpUri="https://binary.pensoft.net/fig/472594" pageId="0" pageNumber="63">9D-F</figureCitation>
<figureCitation id="D8E9DD7951F45DF0DB91C09AB9059B51" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Crassignatha gucheng sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure9" httpUri="https://binary.pensoft.net/fig/472594" pageId="0" pageNumber="63">9D-F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Coloration</emphasis>
<emphasis id="103DD88DA56AC41D5F2C21072F4071E4" bold="true" italics="true" pageId="0" pageNumber="63">Coloration</emphasis>
: carapace roughly the same as in male. Legs pale brown. Abdomen with black pigmentation laterally, posteriorly, and ventrally.
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Prosoma</emphasis>
<emphasis id="62ABA62EC242318BAA2760B5B694C304" bold="true" italics="true" pageId="0" pageNumber="63">Prosoma</emphasis>
: carapace pear shaped, sculptured, cephalic area slightly lower than in male. PER recurved. Mouthparts and sternum as in male.
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Abdomen</emphasis>
<emphasis id="F9CD0A287E0834A09205D2096FEB9FB2" bold="true" italics="true" pageId="0" pageNumber="63">Abdomen</emphasis>
: anteriorly rounded and posteriorly subquadrate, surface with weakly sclerotized hairy patches. Spinnerets as in male.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Epigyne</emphasis>
<paragraph id="C8F22D4A1448614812EA94258237D526" pageId="0" pageNumber="63">
<emphasis id="F088C74FC7BA6764D209F6FB35E4C653" bold="true" italics="true" pageId="0" pageNumber="63">Epigyne</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10D-G</figureCitation>
<figureCitation id="1DE40B675F3E65ACA7EE52E6C39B0ACD" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha gucheng sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure10" httpUri="https://binary.pensoft.net/fig/472595" pageId="0" pageNumber="63">10D-G</figureCitation>
): epigynal area distinctly sclerotized, bears a few setae. Scape large, finger-like, protruded, copulatory opening located at its terminus. Internal structures faintly visible via translucent tegument. Spermathecae globose, strongly sclerotized, widely separated by 1.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="F655EFE33CE79E499AB61AAD15B1821C" originalValue="×">x</normalizedToken>
their diameter. Fertilization duct originates at medial posterior margin of spermathecae, extends to posterior margin of epigyne. Copulatory ducts long, connected to dorsal surface of spermathecae, extends below spermathecae toward the vulval center, and forms two return paths, then turns downward to copulatory opening.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="etymology">
<paragraph pageId="0" pageNumber="63">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="63">The specific name is derived from the type locality; noun in apposition.</paragraph>
<subSubSection id="E40C700B570DC817FF2176AB49A36536" pageId="0" pageNumber="63" type="etymology">
<paragraph id="4699A6A51A2A79B26BF0E7AF28E5CCA5" pageId="0" pageNumber="63">Etymology.</paragraph>
<paragraph id="03218936DB38F769B3D898F49F76C6F8" pageId="0" pageNumber="63">The specific name is derived from the type locality; noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="distribution">
<paragraph pageId="0" pageNumber="63">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="63">
<subSubSection id="A54112FDBCC09DBF33312E537D183142" pageId="0" pageNumber="63" type="distribution">
<paragraph id="123D59D2C1D8470A1C9C22E4C45AC871" pageId="0" pageNumber="63">Distribution.</paragraph>
<paragraph id="1484420BA8FF88058F51B50E99022ADE" pageId="0" pageNumber="63">
China (Yunnan), Vietnam (Fig.
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">38</figureCitation>
<figureCitation id="E076F42F7B06BF6D796182A5BA2FEF88" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">38</figureCitation>
).
</paragraph>
</subSubSection>