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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.519.8784" ID-GBIF-Dataset="0e6858a4-6dae-4dfb-a6fb-787124f42c32" ID-PMC="PMC4591602" ID-Pensoft-Pub="1313-2970-519-1" ID-PubMed="26448705" ID-ZBK="3004DC4729C54C6CA061234F9410D7D3" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-519-1" ModsDocOrigin="ZooKeys 519" ModsDocTitle="Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae)" checkinTime="1451244019430" checkinUser="pensoft" docAuthor="Li, Shuqiang & Zonstein, Sergei" docDate="2015" docId="E59FF3101B42DE2D4E5185DF0CA370CC" docLanguage="en" docName="ZooKeys 519: 1-32" docOrigin="ZooKeys 519" docSource="http://dx.doi.org/10.3897/zookeys.519.8784" docTitle="Raveniola bellula Li & Zonstein, 2015, sp. n." docType="treatment" docUuid="CAB030BD-41D7-43E9-978F-0C0FE340CE19" docUuidSource="ZooBank" docVersion="4" lastPageNumber="6" masterDocId="D67CFFD4FFDFD43BFFBEFF94FFA89A2A" masterDocTitle="Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="5" updateTime="1668160637374" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zonstein, Sergei</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>519</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>32</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.519.8784</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.519.8784</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-519-1</mods:identifier>
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||||
<mods:identifier type="ZBK">3004DC4729C54C6CA061234F9410D7D3</mods:identifier>
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||||
<mods:identifier type="ZooBank">3004DC4729C54C6CA061234F9410D7D3</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152062163" LSID="urn:lsid:zoobank.org:act:CAB030BD-41D7-43E9-978F-0C0FE340CE19" httpUri="http://treatment.plazi.org/id/E59FF3101B42DE2D4E5185DF0CA370CC" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<subSubSection pageId="4" pageNumber="5" type="multiple">
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<paragraph pageId="4" pageNumber="5">Taxon classification Animalia Araneae Nemesiidae</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="nomenclature">
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName LSID="http://zoobank.org/CAB030BD-41D7-43E9-978F-0C0FE340CE19" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola bellula" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bellula">Raveniola bellula</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="5">sp. n.</taxonomicNameLabel>
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Figs 3, 4, 5
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="type material">
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Holotype ♂ - CHINA, Yunnan Province, Mengla County, Xishuangbanna, Menglun Town, primary tropical seasonal rainforest in Menglun Nature Reserve [
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="21.957417">21°57.445'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="101.21661">101°12.997'E</geoCoordinate>
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, 744 m], January 16-31, 2007, G. Zheng (IZCAS). Paratypes: 26♂, 2♀ (IZCAS), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="etymology">
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<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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The specific name is taken from the Latin adjective
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<normalizedToken originalValue="“bellulus”">"bellulus"</normalizedToken>
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(the diminutive form of
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<normalizedToken originalValue="“bellus”">"bellus"</normalizedToken>
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), which means
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<normalizedToken originalValue="“beautiful”">"beautiful"</normalizedToken>
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and refers to the perfect shape of the palpal bulb.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="diagnosis">
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<paragraph pageId="5" pageNumber="6">Diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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This new species is similar to
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola chayi" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chayi">Raveniola chayi</taxonomicName>
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sp. n. and
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola yunnanensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yunnanensis">Raveniola yunnanensis</taxonomicName>
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but can be distinguished by having a considerably shorter embolus than that in
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola chayi" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chayi">Raveniola chayi</taxonomicName>
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sp. n. (Figs 3
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 5
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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; cf. Figs 6
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 7
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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), by possessing a longer cymbium and a less twisted embolus than
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola yunnanensis" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yunnanensis">Raveniola yunnanensis</taxonomicName>
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, as well as by having a ventral abdominal pattern and completely reduced PLS (Fig. 3
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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; cf.
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<bibRefCitation author="Zonstein, S" journalOrPublisher="ZooKeys" pageId="17" pageNumber="18" pagination="71 - 99" title="A review of the genus Raveniola (Araneae, Nemesiidae) in China, with notes on allied genera and description of four new species from Yunnan." url="10.3897/zookeys.211.3060" volume="211" year="2012">Zonstein and Marusik 2012</bibRefCitation>
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: figs 35, 43). Females are characterised by the unique shape of the receptacles, divided into a long, digitiform inner branch and a short, knob-shaped outer branch (Figs 4A, 5D). The habitus and the abdominal pattern of
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola bellula" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bellula">Raveniola bellula</taxonomicName>
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sp. n. resemble that of
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Sinopesa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinopesa maculata" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculata">Sinopesa maculata</taxonomicName>
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, but it is distinguished by generic characters, such as well-developed body and leg setation and by much longer and denser tarsal scopula.
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</paragraph>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figure 3.
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola bellula" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bellula">Raveniola bellula</taxonomicName>
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sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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); 2.0 mm (D, G); 1.0 mm (E, F).
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figure 4.
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola bellula" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bellula">Raveniola bellula</taxonomicName>
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sp. n., female paratype. A vulva, dorsal view B genital area, ventral view. Scale bars: 0.25 mm.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figure 5.
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola bellula" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bellula">Raveniola bellula</taxonomicName>
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sp. n., male holotype (
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) and female paratype (D). A palp, prolateral view B palp, ventral view C palp, retrolateral view D vulva, dorsal view. Scale bars: 0.25 mm.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figure 6.
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola chayi" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chayi">Raveniola chayi</taxonomicName>
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sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view D habitus, dorsal view E leg I, ventral view F leg II, ventral view G habitus, ventral view. Scale bars: 0.5 mm (
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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); 2.0 mm (
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<normalizedToken originalValue="D–G">D-G</normalizedToken>
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).
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figure 7.
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<taxonomicName class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raveniola chayi" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chayi">Raveniola chayi</taxonomicName>
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sp. n., male holotype. A palp, prolateral view B palp, ventral view C palp, retrolateral view. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="description">
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<paragraph pageId="5" pageNumber="6">Description.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Male (holotype): TL 7.90, CL 3.50, CW 2.55, AL 3.60, AW 2.25. Eye diameters and interdistances: AME 0.20, ALE 0.24, PLE 0.16, PME 0.12,
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<normalizedToken originalValue="AME–AME">AME-AME</normalizedToken>
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0.09,
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<normalizedToken originalValue="AME–ALE">AME-ALE</normalizedToken>
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0.04,
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<normalizedToken originalValue="PME–PME">PME-PME</normalizedToken>
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0.35,
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<normalizedToken originalValue="PME–PLE">PME-PLE</normalizedToken>
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0.03. Leg lengths: I: 10.04 (2.75+1.65+2.65+1.80+1.55), II: 9.70 (2.75+1.30+2.30+1.85+1.50), III: 9.05 (2.50+1.15+1.65+2.25+1.50), IV: 12.40 (3.25+1.25+3.10+3.15+1.65). Labium, sternum and maxillae as shown in Fig. 3G. Maxillae with ca. 20 cuspules. Prosoma, palps and legs light brown. Spinnerets light grey. Light yellowish grey abdomen with darker (brown) dorsal and ventral pattern (Fig. 3D, G). Metatarsus I very slightly curved (Fig. 3E). PMS absent; apical segment of PLS digitiform (Fig. 3D, G). Palpal tibia moderately long, subcylindrical; bulb long, oval; embolus gradually tapering to a slender point; cymbium with four short, stout spines (Figs 3
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 5
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Female. TL 6.75, CL 3.25, CW 2.50, AL 2.75, AW 2.40; body and legs colouration as in male. Eye diameters and interdistances: AME 0.22, ALE 0.24, PME 0.10, PLE 0.15,
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<normalizedToken originalValue="AME–AME">AME-AME</normalizedToken>
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0.08,
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<normalizedToken originalValue="AME–ALE">AME-ALE</normalizedToken>
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0.05,
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<normalizedToken originalValue="PME–PME">PME-PME</normalizedToken>
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0.36,
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<normalizedToken originalValue="PME–PLE">PME-PLE</normalizedToken>
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0.04, Leg lengths: I: 10.15 (2.65+1.60+2.55+1.90+1.45), II: 9.55 (2.65+1.45+2.25+1.75+1.45), III: 8.75 (2.45+1.10+1.50+2.25+1.45), IV: 12.00 (3.20+1.15+3.00+3.10+1.55). Genital area as in Fig. 4B. Receptacles divided into a long, digitiform inner branch and a short, knob-shaped outer branch that is bent forward (Figs 4A, 5D).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="variation">
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<paragraph pageId="5" pageNumber="6">Variation.</paragraph>
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<paragraph pageId="5" pageNumber="6">Total length of males and females: 6.75-7.93 (n=8).</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="distribution">
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<paragraph pageId="5" pageNumber="6">Distribution.</paragraph>
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<paragraph pageId="5" pageNumber="6">The species is known only from the type locality.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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