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<document id="95BFA5470F9FB9256F39E77B099B98C3" ID-DOI="10.1186/s13358-022-00261-3" ID-ISSN="1664-2384" ID-Zenodo-Dep="11999827" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1718653084494" checkinUser="felipe" docAuthor="Schwarzhans, Werner, Klots, Oleksandr, Ryabokon, Tamara &amp; Kovalchuk, Oleksandr" docDate="2022" docId="039D7D5BFE40FFA8F872FDD0FE8EFBF2" docLanguage="en" docName="SwissJPalaeontol.141.18.pdf" docOrigin="Swiss Journal of Palaeontology (18) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00261-3" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Gobius ukrainicus Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk 2022, n. sp." docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="FFA40523FE49FFA2FB5FFFDBFFBAFFF1" masterDocTitle="A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths" masterLastPageNumber="35" masterPageNumber="1" pageNumber="10" updateTime="1722278998376" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="47322052AE02A5C9BFA93D7DDCC37806">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:namePart id="DCE5BCB859FE58E8BB607B09623A0AEC">Schwarzhans, Werner</mods:namePart>
<mods:affiliation id="65C3454D3ED545C1D9F450C9AB3F5550">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="3EA09EA85F67230159F44A178E40E395">Klots, Oleksandr</mods:namePart>
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<mods:namePart id="0D610826FB7F9F66F4310F6B142B9E33">Ryabokon, Tamara</mods:namePart>
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<mods:date id="7C2E906D36C2BD84696BC50967560DE6">2022</mods:date>
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<treatment id="039D7D5BFE40FFA8F872FDD0FE8EFBF2" LSID="urn:lsid:plazi:treatment:039D7D5BFE40FFA8F872FDD0FE8EFBF2" httpUri="http://treatment.plazi.org/id/039D7D5BFE40FFA8F872FDD0FE8EFBF2" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<taxonomicName id="4C34B7CEFE40FFABF872FDD0FBBAFDD2" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[813,1024,523,547]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ukrainicus" status="sp. nov.">
<emphasis id="B940105FFE40FFABF872FDD0FBBAFDD2" bold="true" box="[813,1024,523,547]" italics="true" pageId="9" pageNumber="10">Gobius ukrainicus</emphasis>
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<paragraph id="8B8BCC4DFE40FFABF872FD90FC14FD92" blockId="9.[813,942,587,611]" box="[813,942,587,611]" pageId="9" pageNumber="10">
<figureCitation id="130FD0C8FE40FFABF872FD90FC14FD92" box="[813,942,587,611]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: ae Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, be paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,ce (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).gk Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, gi, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="9" pageNumber="10">
Figure
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k
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</paragraph>
<paragraph id="8B8BCC4DFE40FFABF872FD50FBCDFD32" blockId="9.[813,1436,651,707]" pageId="9" pageNumber="10">
<materialsCitation id="3B5CC610FE40FFABF872FD50FBCDFD32" collectionCode="NMNHU-P" country="Ukraine" location="Shydlivshchyna" municipality="Badenian" pageId="9" pageNumber="10" specimenCode="PI 2558" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548F72EFFE40FFABF872FD50FC34FD52" box="[813,910,651,675]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
</emphasis>
<collectionCode id="ED255488FE40FFABF8C6FD57FBA3FD52" box="[921,1049,652,675]" pageId="9" pageNumber="10">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE40FFABFF7BFD57FBC7FD52" box="[1060,1149,651,675]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="9" pageNumber="10">PI 2558</specimenCode>
,
<location id="8EEB9A96FE40FFABFFD3FD50FA8DFD52" LSID="urn:lsid:plazi:treatment:039D7D5BFE40FFA8F872FDD0FE8EFBF2:8EEB9A96FE40FFABFFD3FD50FA8DFD52" box="[1164,1335,651,675]" country="Ukraine" municipality="Badenian" name="Shydlivshchyna" pageId="9" pageNumber="10">Shydlivshchyna</location>
, western
<collectingCountry id="F3238CDDFE40FFABF872FD70FC3DFD32" box="[813,903,683,707]" name="Ukraine" pageId="9" pageNumber="10">Ukraine</collectingCountry>
, late
<collectingMunicipality id="6BEF5637FE40FFABF8E1FD70FB9CFD32" box="[958,1062,683,707]" pageId="9" pageNumber="10">Badenian</collectingMunicipality>
,
<figureCitation id="130FD0C8FE40FFABFF6FFD70FBCEFD32" box="[1072,1140,683,707]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: ae Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, be paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,ce (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).gk Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, gi, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="9" pageNumber="10">Fig. 4j</figureCitation>
.
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</paragraph>
<paragraph id="8B8BCC4DFE40FFABF872FD37FAEFFCB2" blockId="9.[813,1436,747,835]" pageId="9" pageNumber="10">
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<typeStatus id="548F72EFFE40FFABF872FD37FC22FCF2" box="[813,920,748,771]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
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30 otoliths:
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<specimenCount id="9D3207C4FE40FFABFF42FD30FB18FCF2" box="[1053,1186,747,771]" count="8" pageId="9" pageNumber="10" type="generic">8 specimens</specimenCount>
same data as holotype,
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<specimenCode id="DB926436FE40FFABF8EAFCD7FBB6FCD2" box="[949,1036,779,803]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="9" pageNumber="10">PI 2560</specimenCode>
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;
<materialsCitation id="3B5CC610FE40FFABFF48FCD0FC3AFCB2" collectionCode="NMNHU-P" country="Ukraine" pageId="9" pageNumber="10" specimenCode="PI 2559" specimenCount="11" typeStatus="paratype">
<specimenCount id="9D3207C4FE40FFABFF48FCD0FB14FCD2" box="[1047,1198,779,803]" count="11" pageId="9" pageNumber="10" type="generic">11 specimens</specimenCount>
, Mlyntsi,
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<specimenCode id="DB926436FE40FFABF872FCF7FC3AFCB2" box="[813,896,811,835]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="9" pageNumber="10">PI 2559</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE40FFABF8D4FCF0FAE8FCB2" box="[907,1362,811,835]" collectionCode="NMB" country="Ukraine" pageId="9" pageNumber="10" specimenCode="P1212" specimenCount="11" typeStatus="paratype">
<specimenCount id="9D3207C4FE40FFABF8D4FCF0FB9AFCB2" box="[907,1056,811,835]" count="11" pageId="9" pageNumber="10" type="generic">11 specimens</specimenCount>
, Kozatskyi Yar,
<collectionCode id="ED255488FE40FFABFF96FCF7FABFFCB2" box="[1225,1285,812,835]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="9" pageNumber="10">NMB</collectionCode>
<specimenCode id="DB926436FE40FFABFE54FCF0FAE8FCB2" box="[1291,1362,811,835]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="9" pageNumber="10">P1212</specimenCode>
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.
</paragraph>
<paragraph id="8B8BCC4DFE40FFABF872FCB0FB2BFC72" blockId="9.[813,1169,875,899]" box="[813,1169,875,899]" pageId="9" pageNumber="10">
<emphasis id="B940105FFE40FFABF872FCB0FC24FC72" box="[813,926,875,899]" italics="true" pageId="9" pageNumber="10">Etymology</emphasis>
Named after
<collectingCountry id="F3238CDDFE40FFABFF6DFCB0FB37FC72" box="[1074,1165,875,899]" name="Ukraine" pageId="9" pageNumber="10">Ukraine</collectingCountry>
.
</paragraph>
<paragraph id="8B8BCC4DFE40FFABF872FC77FAF4FB72" blockId="9.[813,1437,938,1155]" pageId="9" pageNumber="10">
<emphasis id="B940105FFE40FFABF872FC77FC2DFC32" box="[813,919,940,963]" italics="true" pageId="9" pageNumber="10">Diagnosis</emphasis>
OL:OH = 1.21.3. Anterior rim inclined at 8085° with short preventral projection; posterior rim with moderately developed postdorsal projection. Dorsal rim regularly curved. Sulcus narrow, relatively short (OL:SuL = 1.952.1), inclined at 1520°. Ostial lobe moderate; subcaudal iugum small, below anterior part of cauda. Inner face flat; outer face markedly convex.
</paragraph>
<paragraph id="8B8BCC4DFE40FFABF872FB77FC05F992" blockId="9.[813,1437,1195,1923]" pageId="9" pageNumber="10">
<emphasis id="B940105FFE40FFABF872FB77FC10FB32" box="[813,938,1196,1219]" italics="true" pageId="9" pageNumber="10">Description</emphasis>
Moderately slender, massive otoliths reaching about
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in length (
<typeStatus id="548F72EFFE40FFABFF99FB10FA9DFB12" box="[1222,1319,1227,1251]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
<quantity id="4CCC61A8FE40FFABFE6BFB10FA2BFB12" box="[1332,1425,1227,1251]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="mm" value="1.8">1.8 mm</quantity>
). OH:OT = 2.42.7. Anterior rim slightly inclined, with very feeble concavity at about tip of ostium and distinct but relatively short preventral angle. Predorsal angle distinct, positioned high on anterior rim. Ventral rim nearly straight, smooth. Dorsal rim regularly curved usually without prominent angles or concavities, sometimes slightly undulating or crenulated, slightly ascending toward posterior, with distinct moderately strong postdorsal projection with rounded tip and not bent outward. Posterior rim below postdorsal projection slightly incised, thereafter nearly vertical, with orthogonal postventral angle.
</paragraph>
<paragraph id="8B8BCC4DFE40FFA8F81DF9B7FDB1FE32" blockId="9.[813,1437,1195,1923]" lastBlockId="10.[151,776,235,452]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Inner face slightly convex, nearly flat, with narrow, somewhat deepened, centrally positioned, distinctly inclined sulcus. SuL:SuH = 2.42.8. Sulcus with moderate to low ostial lobe, sole-shaped, and somewhat pointed anterior tip and more rounded posterior tip, inclined at 1520°. Subcaudal iugum small, knob-like, sometimes poorly discernible (
<figureCitation id="130FD0C8FE40FFABF8CCF8F0FC46F8B2" box="[915,1020,1835,1860]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: ae Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, be paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,ce (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).gk Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, gi, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="9" pageNumber="10">
Fig.
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, h
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), positioned below sulcus indentation marking ostialcaudal joint. Ventral furrow broad, deep, distinct, reaching from anterior to posterior tips of sulcus, more regularly curved than ventral rim of otolith. Ventral field between ventral furrow and sulcus slightly bulged. Dorsal field with long, distinct and relatively broad dorsal depression, ventrally marked by distinct crista superior toward sulcus and dorsally closed to dorsal rim of otolith. Outer face convex, distinctly more convex than inner face, and very regularly shaped, smooth.
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE43FFA8FBC8FE37FE8EFBF2" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8B8BCC4DFE43FFA8FBC8FE37FE8EFBF2" blockId="10.[151,776,491,1027]" pageId="10" pageNumber="11">
<emphasis id="B940105FFE43FFA8FBC8FE37FE66FDF2" box="[151,476,491,515]" italics="true" pageId="10" pageNumber="11">
Discussion
<taxonomicName id="4C34B7CEFE43FFA8FA4BFE30FE66FDF2" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[276,476,491,515]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="ukrainicus">Gobius ukrainicus</taxonomicName>
</emphasis>
is the most common species at Horodok in the Medobory back-reef environment. It belongs to the
<taxonomicName id="4C34B7CEFE43FFA8FA11FDF0FE55FDB2" authorityName="Schwarzhans" authorityYear="2014" box="[334,495,555,579]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="mustus">
<emphasis id="B940105FFE43FFA8FA11FDF0FE55FDB2" box="[334,495,555,579]" italics="true" pageId="10" pageNumber="11">Gobius mustus</emphasis>
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complex and differs from the coeval
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<emphasis id="B940105FFE43FFA8FA54FD97FE2FFD92" box="[267,405,587,611]" italics="true" pageId="10" pageNumber="11">G. bratishkoi</emphasis>
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in the regularly curved dorsal rim (vs. anteriorly with broad concavity), the slightly inclined anterior rim (vs. vertical), and the moderately developed ostial lobe (vs. very low ostial lobe), which results in the sole-shaped sulcus. It also resembles
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<emphasis id="B940105FFE43FFA8F91DFD17FD0EFD12" box="[578,692,716,739]" italics="true" pageId="10" pageNumber="11">G. mustus</emphasis>
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, which occurs in non-reef-related environments during the same time, and differs in a number of subtle but nevertheless stable characters.
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<emphasis id="B940105FFE43FFA8FA04FCF0FD9BFCB2" box="[347,545,811,835]" italics="true" pageId="10" pageNumber="11">Gobius ukrainicus</emphasis>
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differs from
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<emphasis id="B940105FFE43FFA8F9EFFCF7FF02FC92" italics="true" pageId="10" pageNumber="11">G. mustus</emphasis>
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in the regularly curved dorsal rim, the not outward bent postdorsal projection (vs. slightly outward bent), the shorter sulcus (OL:SuL = 1.952.1 vs. 1.651.9), and the ventral furrow clipping the pre- and postventral angles (vs. more strictly following the shape of the ventral rim of the otolith).
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Figure 4
<bibRefCitation id="EFA5B1BCFE40FFABFBB0FA30FF4DF9F2" author="Bannikov, A. F." box="[239,247,1515,1539]" pageId="9" pageNumber="10" pagination="23 - 34" refId="ref19263" refString="Bannikov, A. F. (2014). The systematic composition of the Eocene actinopterygian fish fauna from Monte Bolca, northern Italy, as known to date. Studi e ricerche sui giacimenti terziari di Bolca, XV. Miscellanea Paleontologica, 12, 23 - 34." type="journal article" year="2014">f</bibRefCitation>
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2014
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<emphasis id="B940105FFE40FFABFBBAF9F0FE43F9B2" box="[229,505,1579,1603]" italics="true" pageId="9" pageNumber="10">Gobius reichenbacherae</emphasis>
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—Schwarzhans: pl. 9, figs. 29.
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<paragraph id="8B8BCC4DFE40FFABFBC8F950FE1BF912" blockId="9.[151,775,1675,1763]" pageId="9" pageNumber="10">
2020a
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<emphasis id="B940105FFE40FFABFBB3F950FE46F952" box="[236,508,1675,1699]" italics="true" pageId="9" pageNumber="10">Gobius reichenbacherae</emphasis>
<bibRefCitation id="EFA5B1BCFE40FFABF94FF950FD54F952" author="Schwarzhans, W." box="[528,750,1675,1699]" pageId="9" pageNumber="10" pagination="35 - 69" refId="ref23301" refString="Schwarzhans, W. (2014). Otoliths from the middle Miocene (Serravallian) of the Karaman Basin, Turkey. Cainozoic Research, 14 (1), 35 - 69." type="journal article" year="2014">Schwarzhans, 2014</bibRefCitation>
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— Schwarzhans, Brzobohatý &amp; Radwańska: pl. 4, figs. 69 (and references therein).
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<emphasis id="B940105FFE40FFABFBC8F8D0FF42F8D2" box="[151,248,1803,1827]" italics="true" pageId="9" pageNumber="10">Material</emphasis>
1 otolith, Shydlivshchyna,
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<collectionCode id="ED255488FE40FFABF9FDF8D7FD00F8D2" box="[674,698,1804,1827]" country="Russia" name="Paleontological Institute" pageId="9" pageNumber="10">PI</collectionCode>
2557.
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<emphasis id="B940105FFE40FFABFBC8F897FEB0F892" box="[151,266,1868,1891]" italics="true" pageId="9" pageNumber="10">Discussion</emphasis>
Te single, large otolith of
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in length is readily recognized by the slightly convex inner face and the flat outer face, the parallelogram-like shape, and the postdorsal projection being markedly bent outward. Te sulcus shows a shoe sole shape with moderate ostial lobe, and the subcaudal iugum is clearly discernable despite erosion.
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,
<bibRefCitation id="EFA5B1BCFE40FFABFE5FFE10FAF9FE12" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[1280,1347,459,483]" pageId="9" pageNumber="10" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
).
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<paragraph id="8B8BCC4DFE43FFA8FBC8FBB0FCBDFB72" blockId="10.[151,775,1131,1187]" box="[151,775,1131,1155]" pageId="10" pageNumber="11">
<taxonomicName id="4C34B7CEFE43FFA8FBC8FBB0FCBAFB72" authority="Schwarzhans et al., 2020 a" authorityName="Schwarzhans" authorityYear="2020" box="[151,768,1131,1155]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
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<emphasis id="B940105FFE43FFA8FBC8FBB0FE0DFB72" bold="true" box="[151,439,1131,1155]" italics="true" pageId="10" pageNumber="11">Odondebuenia agiadiae</emphasis>
<bibRefCitation id="EFA5B1BCFE43FFA8FA9BFBB0FD0BFB72" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[452,689,1131,1155]" pageId="10" pageNumber="11" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al.,</bibRefCitation>
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2020a
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,
</paragraph>
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<heading id="D0C37B21FE43FFA8FBC8FB50FF60FB52" box="[151,218,1163,1187]" fontSize="10" level="2" pageId="10" pageNumber="11" reason="7">
<bibRefCitation id="EFA5B1BCFE43FFA8FBC8FB50FF60FB52" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[151,218,1163,1187]" pageId="10" pageNumber="11" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
</heading>
</paragraph>
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<subSubSection id="C32E9FC6FE43FFA8FBC8FB10FD30F892" pageId="10" pageNumber="11" type="description">
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<figureCitation id="130FD0C8FE43FFA8FBC8FB10FEA3FB12" box="[151,281,1227,1251]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="10" pageNumber="11">Figure 5ad</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE43FFA8FBC8FAD0FD9EFAB2" blockId="10.[151,775,1291,1347]" pageId="10" pageNumber="11">
2020a
<taxonomicName id="4C34B7CEFE43FFA8FBB9FAD0FE49FAD2" box="[230,499,1291,1315]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE43FFA8FBB9FAD0FE49FAD2" box="[230,499,1291,1315]" italics="true" pageId="10" pageNumber="11">Odondebuenia agiadiae</emphasis>
</taxonomicName>
—Schwarzhans, Brzobohatý &amp; Radwańska: pl. 6, figs. 1222.
</paragraph>
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<emphasis id="B940105FFE43FFA8FBC8FAB0FF42FA72" box="[151,248,1387,1411]" italics="true" pageId="10" pageNumber="11">Material</emphasis>
7 otoliths:
<specimenCount id="9D3207C4FE43FFA8FADDFAB0FDABFA72" box="[386,529,1387,1411]" count="2" pageId="10" pageNumber="11" type="generic">2 specimens</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE43FFA8F9D9FAB7FCBCFA72" box="[646,774,1388,1411]" pageId="10" pageNumber="11">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE43FFA8FBC8FA57FF15FA52" box="[151,175,1420,1443]" country="Russia" name="Paleontological Institute" pageId="10" pageNumber="11">PI</collectionCode>
2570;
<specimenCount id="9D3207C4FE43FFA8FBA8FA50FE3BFA52" box="[247,385,1419,1443]" count="5" pageId="10" pageNumber="11" type="generic">5 specimens</specimenCount>
, Kozatskyi Yar,
<collectionCode id="ED255488FE43FFA8F973FA57FD16FA52" box="[556,684,1420,1443]" pageId="10" pageNumber="11">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE43FFA8F9EBFA57FD76FA52" box="[692,716,1420,1443]" country="Russia" name="Paleontological Institute" pageId="10" pageNumber="11">PI</collectionCode>
2569 and
<collectionCode id="ED255488FE43FFA8FB99FA77FEB8FA32" box="[198,258,1452,1475]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="10" pageNumber="11">NMB</collectionCode>
1219.
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<emphasis id="B940105FFE43FFA8FBC8FA37FEB0F9F2" box="[151,266,1516,1539]" italics="true" pageId="10" pageNumber="11">Discussion</emphasis>
Otoliths of
<taxonomicName id="4C34B7CEFE43FFA8FAC7FA37FDA3F9F2" box="[408,537,1515,1539]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE43FFA8FAC7FA37FDA3F9F2" box="[408,537,1515,1539]" italics="true" pageId="10" pageNumber="11">O. agiadiae</emphasis>
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are characterized by the preventral projection being stronger than the postdorsal one, a depressed predorsal region, and a small, rather narrow sulcus with reduced sole shape and without a subcaudal iugum. Outline and proportions were found to vary considerably in the original description by Schwarzhans et. al. (2020a); this variability is confirmed here.
<taxonomicName id="4C34B7CEFE43FFA8FB8BF910FE60F912" box="[212,474,1739,1763]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE43FFA8FB8BF910FE60F912" box="[212,474,1739,1763]" italics="true" pageId="10" pageNumber="11">Odondebuenia agiadiae</emphasis>
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is apparently a species that preferred near-reef environments, such as Lakšárská Nová Ves (late Badenian of
<collectingCountry id="F3238CDDFE43FFA8FA90F8D0FD91F8D2" box="[463,555,1803,1827]" name="Slovakia" pageId="10" pageNumber="11">Slovakia</collectingCountry>
) or Węglinek (early Badenian of
<collectingCountry id="F3238CDDFE43FFA8FA77F8F0FECDF8B2" box="[296,375,1835,1859]" name="Poland" pageId="10" pageNumber="11">Poland</collectingCountry>
), and is now also confirmed in the back-reef environment of the Medobory Reef.
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<heading id="D0C37B21FE43FFA8F872FEF0FBE5FEB2" bold="true" box="[813,1119,299,323]" fontSize="10" level="1" pageId="10" pageNumber="11" reason="1">
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Genus
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<emphasis id="B940105FFE43FFA8F822FEF0FBB7FEB2" bold="true" box="[893,1037,299,323]" italics="true" pageId="10" pageNumber="11">Parenypnias</emphasis>
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</emphasis>
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<typeStatus id="548F72EFFE43FFA8F872FEB7FCDBFE72" box="[813,865,364,387]" pageId="10" pageNumber="11">Type</typeStatus>
species
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:
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<emphasis id="B940105FFE43FFA8F8E3FEB7FB0BFE72" box="[956,1201,363,387]" italics="true" pageId="10" pageNumber="11">Parenypnias inauditus</emphasis>
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<emphasis id="B940105FFE43FFA8F872FE70FC24FE32" box="[813,926,427,451]" italics="true" pageId="10" pageNumber="11">Etymology</emphasis>
Derived from the gobiosomatine genus
<taxonomicName id="4C34B7CEFE43FFA8FE02FE77FCE1FE12" authorityName="Jordan &amp; Evermann" authorityYear="1898" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE43FFA8FE02FE77FCE1FE12" italics="true" pageId="10" pageNumber="11">Enypnias</emphasis>
</taxonomicName>
<collectingCountry id="F3238CDDFE43FFA8F83BFE10FC16FE12" box="[868,940,459,483]" name="Jordan" pageId="10" pageNumber="11">Jordan</collectingCountry>
&amp; Evermann, 1898 because of some resemblance to otoliths of that genus, which, however, is regarded as a junior synonym of
<taxonomicName id="4C34B7CEFE43FFA8FFCDFDD0FA26FDD2" authority="Girard, 1858" authorityName="Girard" authorityYear="1858" box="[1170,1436,523,547]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE43FFA8FFCDFDD0FAB6FDD2" box="[1170,1292,523,547]" italics="true" pageId="10" pageNumber="11">Gobiosoma</emphasis>
Girard, 1858
</taxonomicName>
according to
<bibRefCitation id="EFA5B1BCFE43FFA8F8E5FDF0FA9DFDB2" author="Tornabene, L. &amp; Van Tassell, J. L." box="[954,1319,555,579]" pageId="10" pageNumber="11" refId="ref23817" refString="Tornabene, L., &amp; Van Tassell, J. L. (2014). Redescription of the goby genus Gobiosoma (Teleostei: Gobiidae: Gobiosomatini), with the synonymy" type="book" year="2014">Tornabene and van Tassell (2014)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8BCC4DFE43FFA8F872FDB7FBDBFBD2" blockId="10.[813,1437,619,1859]" pageId="10" pageNumber="11">
<emphasis id="B940105FFE43FFA8F872FDB7FC2DFD72" box="[813,919,620,643]" italics="true" pageId="10" pageNumber="11">Diagnosis</emphasis>
An otolith of the family
<taxonomicName id="4C34B7CEFE43FFA8FFEFFDB0FAAFFD72" authorityName="Cuvier" authorityYear="1816" box="[1200,1301,619,643]" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
, supposedly belonging to the Gobiosomatini, with the following combination of diagnostic characters. Otolith size small, up to slightly over
<quantity id="4CCC61A8FE43FFA8F884FD10FB8BFD12" box="[987,1073,715,739]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="mm" value="1.5">1.5 mm</quantity>
in length. Ventral rim of otolith straight, horizontal; anterior rim near vertical or with pointed preventral projection; anterior and posterior rims with deep incisions at anterior and posterior terminations of sulcus. Dorsal rim with distinct middorsal angle and relatively short, mildly to distinctly outward bent postdorsal projection. Inner and outer faces about equally convex or inner face more convex. Sulcus narrow, moderately short, oval, without ostial lobe or subcaudal iugum, inclined at 15°18°. Ventral furrow following outline of ventral rim of otolith.
</paragraph>
<paragraph id="8B8BCC4DFE43FFA8F81DFBF7FA27FA32" blockId="10.[813,1437,619,1859]" pageId="10" pageNumber="11">
<emphasis id="B940105FFE43FFA8F81DFBF7FC0FFBB2" box="[834,949,1068,1091]" italics="true" pageId="10" pageNumber="11">Discussion</emphasis>
Te Gobiosomatini are a group of gobiid fishes that are today only known from the seas around the Americas, where they form one of the most diverse and species rich groups of the family (
<bibRefCitation id="EFA5B1BCFE43FFA8FF91FB50FA37FB52" author="RUber, L. &amp; Van Tassell, J. L. &amp; Zardoya, R." box="[1230,1421,1163,1187]" pageId="10" pageNumber="11" pagination="1584 - 1598" refId="ref22835" refString="RUber, L., Van Tassell, J. L., &amp; Zardoya, R. (2003). Rapid speciation and ecological divergence in the American seven-spined gobies (Gobiidae, Gobiosomatini) inferred from a molecular phylogeny. Evolution, 57 (7), 1584 - 1598." type="journal article" year="2003">Rüber et al., 2003</bibRefCitation>
). Teir otoliths appear inconspicuous at first glance but are characterized by certain traits not usually found in combination in Atlanto-Mediterranean goby groups. Tese traits are the relatively narrow sulcus with or without ostial lobe, the lack of a subcaudal iugum, the outward bent postdorsal projection, and the inner face being as strongly convex as the outer face or more so. In that respect,
<taxonomicName id="4C34B7CEFE43FFA8F8CFFA57FBACFA52" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[912,1046,1420,1443]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE43FFA8F8CFFA57FBACFA52" box="[912,1046,1420,1443]" italics="true" pageId="10" pageNumber="11">Parenypnias</emphasis>
</taxonomicName>
closely resembles otoliths of the
<typeStatus id="548F72EFFE43FFA8F872FA77FCE6FA35" box="[813,860,1452,1476]" pageId="10" pageNumber="11">type</typeStatus>
species of
<taxonomicName id="4C34B7CEFE43FFA8F883FA77FBFAFA32" authorityName="Jordan &amp; Evermann" authorityYear="1898" box="[988,1088,1452,1475]" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE43FFA8F883FA77FBFAFA32" box="[988,1088,1452,1475]" italics="true" pageId="10" pageNumber="11">Enypnias</emphasis>
</taxonomicName>
,
<taxonomicName id="4C34B7CEFE43FFA8FF0FFA77FA27FA32" authority="(Gunther, 1861)" baseAuthorityName="Gunther" baseAuthorityYear="1861" box="[1104,1437,1451,1475]" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="seminudus">
<emphasis id="B940105FFE43FFA8FF0FFA77FB59FA32" box="[1104,1251,1451,1475]" italics="true" pageId="10" pageNumber="11">E. seminudus</emphasis>
(Günther, 1861)
</taxonomicName>
</paragraph>
<paragraph id="8B8BCC4DFE43FFA9F872FA10FD03F93B" blockId="10.[813,1437,619,1859]" lastBlockId="11.[151,775,1682,1898]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
(
<figureCitation id="130FD0C8FE43FFA8F86BFA10FCC4FA12" box="[820,894,1483,1507]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="10" pageNumber="11">Fig. 5j</figureCitation>
) in the outline of the otolith, the pronounced postdorsal projection, and the narrow sulcus with a low ostial lobe. It differs in the rather strongly reduced and small, nearly oval sulcus (vs. sole-shaped or somewhat smoothened sole-shaped) and the ventral furrow following the ventral rim of the otolith (vs. being more regularly curved and clipping the postventral angles/projections). Te otoliths of
<taxonomicName id="4C34B7CEFE43FFA8F888F970FBEBF932" authorityName="Girard" authorityYear="1858" box="[983,1105,1707,1731]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE43FFA8F888F970FBEBF932" box="[983,1105,1707,1731]" italics="true" pageId="10" pageNumber="11">Gobiosoma</emphasis>
</taxonomicName>
with the
<typeStatus id="548F72EFFE43FFA8FFE0F977FB54F935" box="[1215,1262,1708,1732]" pageId="10" pageNumber="11">type</typeStatus>
species
<taxonomicName id="4C34B7CEFE43FFA8FE12F977FC5DF912" authority="(Lacepede, 1800)" baseAuthorityName="Lacepede" baseAuthorityYear="1800" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="bosc">
<emphasis id="B940105FFE43FFA8FE12F977FA26F932" box="[1357,1436,1707,1731]" italics="true" pageId="10" pageNumber="11">G. bosc</emphasis>
(Lacépède, 1800)
</taxonomicName>
(
<figureCitation id="130FD0C8FE43FFA8F8A8F910FBF9F912" box="[1015,1091,1739,1763]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="10" pageNumber="11">Fig. 5k</figureCitation>
) are more similar in respect to the shape and size of the sulcus but differ significantly in the reduced postdorsal projection and the postventral projection being significantly more pronounced than the postdorsal one. In this respect,
<taxonomicName id="4C34B7CEFE42FFA9F972F948FD09F95B" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[557,691,1683,1706]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE42FFA9F972F948FD09F95B" box="[557,691,1683,1706]" italics="true" pageId="11" pageNumber="12">Parenypnias</emphasis>
</taxonomicName>
almost resembles a combination of the two morphotypes.
</paragraph>
<caption id="DF4B9CC5FE42FFA9FBF7FA61FF68F9A2" ID-DOI="http://doi.org/10.5281/zenodo.11999891" ID-Zenodo-Dep="11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="11" pageNumber="12" startId="11.[168,199,1466,1486]" targetBox="[152,1438,237,1629]" targetPageId="11" targetType="figure">
<paragraph id="8B8BCC4DFE42FFA9FBF7FA61FF68F9A2" blockId="11.[168,1421,1466,1619]" pageId="11" pageNumber="12">
<emphasis id="B940105FFE42FFA9FBF7FA61FF61FA3F" bold="true" box="[168,219,1466,1486]" pageId="11" pageNumber="12">Fig. 5</emphasis>
Otoliths of
<emphasis id="B940105FFE42FFA9FA1CFA61FD93FA3F" box="[323,553,1466,1486]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C34B7CEFE42FFA9FA1CFA61FE05FA3F" authorityName="de Buen" authorityYear="1930" box="[323,447,1466,1486]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Odondebuenia</taxonomicName>
,
<taxonomicName id="4C34B7CEFE42FFA9FA99FA61FD93FA3F" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[454,553,1466,1486]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Parenypnias</taxonomicName>
</emphasis>
and
<taxonomicName id="4C34B7CEFE42FFA9F90CFA61FD02FA3F" authorityName="Bleeker" authorityYear="1878" box="[595,696,1466,1486]" family="Gobiidae" genus="Bathygobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE42FFA9F90CFA61FD02FA3F" box="[595,696,1466,1486]" italics="true" pageId="11" pageNumber="12">Bathygobius</emphasis>
</taxonomicName>
?:
<emphasis id="B940105FFE42FFA9F996FA61FD50FA3F" bold="true" box="[713,746,1466,1486]" pageId="11" pageNumber="12">ad</emphasis>
<taxonomicName id="4C34B7CEFE42FFA9F9B0FA61FB28FA3F" authority="Schwarzhans et al., 2020 a" authorityName="Schwarzhans" authorityYear="2020" box="[751,1170,1466,1486]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE42FFA9F9B0FA61FC0FFA3F" box="[751,949,1466,1486]" italics="true" pageId="11" pageNumber="12">Odondebuenia agiadiae</emphasis>
<bibRefCitation id="EFA5B1BCFE42FFA9F8E5FA61FB28FA3F" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[954,1170,1466,1486]" pageId="11" pageNumber="12" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFA5B1BCFE42FFA9FFC5FA61FB75FA3F" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[1178,1231,1466,1486]" pageId="11" pageNumber="12" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
,
<emphasis id="B940105FFE42FFA9FF88FA61FB58FA3F" bold="true" box="[1239,1250,1466,1486]" pageId="11" pageNumber="12">a</emphasis>
(reversed) Kozatskyi Yar,NMB P1219,
<emphasis id="B940105FFE42FFA9FA6BFA0FFEFAFA19" bold="true" box="[308,320,1492,1512]" pageId="11" pageNumber="12">b</emphasis>
(reversed),
<emphasis id="B940105FFE42FFA9FAFEFA0FFE17FA19" bold="true" box="[417,429,1492,1512]" pageId="11" pageNumber="12">d</emphasis>
Kozatskyi Yar, NMNHU-P PI 2569,
<emphasis id="B940105FFE42FFA9F998FA0FFD6BFA19" bold="true" box="[711,721,1492,1512]" pageId="11" pageNumber="12">c</emphasis>
(reversed) Mlyntsi, NMNHU-P PI 2570.
<emphasis id="B940105FFE42FFA9FF4AFA0FFB8DFA19" bold="true" box="[1045,1079,1492,1512]" pageId="11" pageNumber="12">eh</emphasis>
<taxonomicName id="4C34B7CEFE42FFA9FF63FA0FFB55FA19" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[1084,1263,1492,1512]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="inauditus" status="n. gen. et n. sp.">
<emphasis id="B940105FFE42FFA9FF63FA0FFB55FA19" box="[1084,1263,1492,1512]" italics="true" pageId="11" pageNumber="12">Parenypnias inauditus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE42FFA9FFABFA0FFACFFA19" box="[1268,1397,1492,1512]" pageId="11" pageNumber="12" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
,
<emphasis id="B940105FFE42FFA9FE24FA0FFA38FA19" bold="true" box="[1403,1410,1492,1512]" pageId="11" pageNumber="12">f</emphasis>
holotype, Kozatskyi Yar,NMNHU-P PI 2550,
<emphasis id="B940105FFE42FFA9F94DFA34FDA7F9F2" bold="true" box="[530,541,1519,1539]" pageId="11" pageNumber="12">e</emphasis>
,
<emphasis id="B940105FFE42FFA9F979FA34FD88F9F2" bold="true" box="[550,562,1519,1539]" pageId="11" pageNumber="12">g</emphasis>
(reversed) Shydlivshchyna, NMB P1207,
<emphasis id="B940105FFE42FFA9F8DAFA34FC2BF9F2" bold="true" box="[901,913,1519,1539]" pageId="11" pageNumber="12">h</emphasis>
Žižkov 1 well (16161622 m), DGS MU-0395.
<emphasis id="B940105FFE42FFA9FE4FFA34FAAFF9F2" bold="true" box="[1296,1301,1519,1539]" pageId="11" pageNumber="12">i</emphasis>
<taxonomicName id="4C34B7CEFE42FFA9FE45FA34FF5BF9EF" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kiselevi" status="n. gen. et n. sp.">
<emphasis id="B940105FFE42FFA9FE45FA34FF5BF9EF" italics="true" pageId="11" pageNumber="12">Parenypnias kiselevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE42FFA9FBB9F9D1FEDDF9EF" box="[230,359,1546,1566]" pageId="11" pageNumber="12" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
, holotype, Mlyntsi,NMNHU-P PI 2574.
<emphasis id="B940105FFE42FFA9F9F6F9D1FD15F9EF" bold="true" box="[681,687,1546,1566]" pageId="11" pageNumber="12">j</emphasis>
<taxonomicName id="4C34B7CEFE42FFA9F9ECF9D1FCE2F9EF" baseAuthorityName="Gunther" baseAuthorityYear="1861" box="[691,856,1546,1566]" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="seminudus">
<emphasis id="B940105FFE42FFA9F9ECF9D1FCE2F9EF" box="[691,856,1546,1566]" italics="true" pageId="11" pageNumber="12">Enypnias seminudus</emphasis>
</taxonomicName>
(GÜnther, 1861), Recent,USNM 407784,13° 22
<emphasis id="B940105FFE42FFA9FFBDF9D2FB5DF9EC" bold="true" box="[1250,1255,1545,1565]" italics="true" pageId="11" pageNumber="12"></emphasis>
N 87° 52
<emphasis id="B940105FFE42FFA9FE6CF9D2FA82F9EC" bold="true" box="[1331,1336,1545,1565]" italics="true" pageId="11" pageNumber="12"></emphasis>
W.
<emphasis id="B940105FFE42FFA9FE0AF9D1FADAF9EF" bold="true" box="[1365,1376,1546,1566]" pageId="11" pageNumber="12">k</emphasis>
<taxonomicName id="4C34B7CEFE42FFA9FBF7F9FFFE7DF9C9" authority="(Lacepede, 1800)" baseAuthorityName="Lacepede" baseAuthorityYear="1800" box="[168,455,1572,1592]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="bosc">
<emphasis id="B940105FFE42FFA9FBF7F9FFFE8BF9C9" box="[168,305,1572,1592]" italics="true" pageId="11" pageNumber="12">Gobiosoma bosc</emphasis>
(Lacépède,1800)
</taxonomicName>
, Recent (reversed), LACM coll. Fitch, off Rappahannock.
<emphasis id="B940105FFE42FFA9F8C2F9FFFC18F9C9" bold="true" box="[925,930,1572,1592]" pageId="11" pageNumber="12">l</emphasis>
<taxonomicName id="4C34B7CEFE42FFA9F8F8F9FFFBB6F9C9" authorityName="Bleeker" authorityYear="1878" box="[935,1036,1572,1592]" family="Gobiidae" genus="Bathygobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE42FFA9F8F8F9FFFBB6F9C9" box="[935,1036,1572,1592]" italics="true" pageId="11" pageNumber="12">Bathygobius</emphasis>
</taxonomicName>
? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548
</paragraph>
</caption>
<paragraph id="8B8BCC4DFE42FFA9FBF2F909FBDEF89B" blockId="11.[151,775,1682,1898]" lastBlockId="11.[813,1437,1682,1898]" pageId="11" pageNumber="12">
Te Gobiosomatini are considered to represent a group endemic to the tropical to subtropical shores of the Americas (
<bibRefCitation id="EFA5B1BCFE42FFA9FA4EF8C9FE61F8DB" author="RUber, L. &amp; Van Tassell, J. L. &amp; Zardoya, R." box="[273,475,1810,1834]" pageId="11" pageNumber="12" pagination="1584 - 1598" refId="ref22835" refString="RUber, L., Van Tassell, J. L., &amp; Zardoya, R. (2003). Rapid speciation and ecological divergence in the American seven-spined gobies (Gobiidae, Gobiosomatini) inferred from a molecular phylogeny. Evolution, 57 (7), 1584 - 1598." type="journal article" year="2003">Rüber et al., 2003</bibRefCitation>
). Teir outgroup relationship is still elusive and has been discussed in Rüber et. al. (2003). Due to the occurrence of
<taxonomicName id="4C34B7CEFE42FFA9F960F888FD7FF89B" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[575,709,1875,1898]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE42FFA9F960F888FD7FF89B" box="[575,709,1875,1898]" italics="true" pageId="11" pageNumber="12">Parenypnias</emphasis>
</taxonomicName>
away from the extant distribution range and some remaining morphological uncertainties, our allocation to the Gobiosomatini should be considered tentative. Interestingly, however, the extant Gobiosomatini of the genus
<taxonomicName id="4C34B7CEFE42FFA9FE02F928FCE1F8DB" authorityName="Jordan &amp; Evermann" authorityYear="1898" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE42FFA9FE02F928FCE1F8DB" italics="true" pageId="11" pageNumber="12">Enypnias</emphasis>
</taxonomicName>
are mostly associated with reef environments, while those of the genus
<taxonomicName id="4C34B7CEFE42FFA9FF57F8E9FB38F8BB" authorityName="Girard" authorityYear="1858" box="[1032,1154,1842,1866]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE42FFA9FF57F8E9FB38F8BB" box="[1032,1154,1842,1866]" italics="true" pageId="11" pageNumber="12">Gobiosoma</emphasis>
</taxonomicName>
occur in shallow coastal and estuarine environments.
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBC8FF37FDC4FEB2" blockId="12.[151,775,235,323]" pageId="12" pageNumber="13">
<emphasis id="B940105FFE45FFAEFBC8FF37FF5EFEF2" box="[151,228,236,259]" italics="true" pageId="12" pageNumber="13">Species</emphasis>
Two species:
<taxonomicName id="4C34B7CEFE45FFAEFADDFF37FDBEFEF2" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[386,516,235,259]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="inauditus" status="n. sp.">
<emphasis id="B940105FFE45FFAEFADDFF37FDBEFEF2" box="[386,516,235,259]" italics="true" pageId="12" pageNumber="13">P. inauditus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE45FFAEF952FF37FDF0FEF2" box="[525,586,236,259]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
, which is known from the late Badenian of the
<collectingCountry id="F3238CDDFE45FFAEFABFFED0FD31FED2" box="[480,651,267,291]" name="Czech Republic" pageId="12" pageNumber="13">Czech Republic</collectingCountry>
(as revised here) and
<collectingCountry id="F3238CDDFE45FFAEFA5BFEF0FEE4FEB2" box="[260,350,299,323]" name="Ukraine" pageId="12" pageNumber="13">Ukraine</collectingCountry>
, and
<taxonomicName id="4C34B7CEFE45FFAEFAC8FEF7FE44FEB2" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[407,510,299,323]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kiselevi">
<emphasis id="B940105FFE45FFAEFAC8FEF7FE44FEB2" box="[407,510,299,323]" italics="true" pageId="12" pageNumber="13">P. kiselevi</emphasis>
</taxonomicName>
in
<collectingCountry id="F3238CDDFE45FFAEF97FFEF0FDC1FEB2" box="[544,635,299,323]" name="Ukraine" pageId="12" pageNumber="13">Ukraine</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,183 @@
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<mods:title id="31B490E1E585D303239883E281D3BB2B">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<paragraph id="8B8BCC4DFE44FFAFF872FE70FB3DFE32" blockId="13.[813,1159,427,451]" box="[813,1159,427,451]" pageId="13" pageNumber="14">
<heading id="D0C37B21FE44FFAFF872FE70FB3DFE32" bold="true" box="[813,1159,427,451]" fontSize="10" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B940105FFE44FFAFF872FE70FB3DFE32" bold="true" box="[813,1159,427,451]" pageId="13" pageNumber="14">
Genus
<taxonomicName id="4C34B7CEFE44FFAFF822FE70FB8FFE32" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[893,1077,427,451]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B940105FFE44FFAFF822FE70FB8FFE32" bold="true" box="[893,1077,427,451]" italics="true" pageId="13" pageNumber="14">Medoborichthys</emphasis>
</taxonomicName>
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</emphasis>
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<emphasis id="B940105FFE44FFAFF872FE37FC0AFDF2" box="[813,944,492,515]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="548F72EFFE44FFAFF872FE37FCDBFDF2" box="[813,865,492,515]" pageId="13" pageNumber="14">Type</typeStatus>
species
</emphasis>
:
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<emphasis id="B940105FFE44FFAFF8E3FE30FB62FDF2" box="[956,1240,491,515]" italics="true" pageId="13" pageNumber="14">Medoborichthys renesulcis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE44FFAFFF81FE37FAA2FDF2" box="[1246,1304,492,515]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B8BCC4DFE44FFAFF872FDF0FC0CFD72" blockId="13.[813,1436,555,643]" pageId="13" pageNumber="14">
<emphasis id="B940105FFE44FFAFF872FDF0FC24FDB2" box="[813,926,555,579]" italics="true" pageId="13" pageNumber="14">Etymology</emphasis>
Referring to the Medobory barrier reef complex of late Badenian to early Sarmatian s.l. age in western
<collectingCountry id="F3238CDDFE44FFAFF808FDB0FC08FD72" box="[855,946,619,643]" name="Ukraine" pageId="13" pageNumber="14">Ukraine</collectingCountry>
.
</paragraph>
<paragraph id="8B8BCC4DFE44FFAFF872FD77FCC7FB92" blockId="13.[813,1437,683,1123]" pageId="13" pageNumber="14">
<emphasis id="B940105FFE44FFAFF872FD77FC2DFD32" box="[813,919,684,707]" italics="true" pageId="13" pageNumber="14">Diagnosis</emphasis>
An otolith of the family
<taxonomicName id="4C34B7CEFE44FFAFFFEFFD70FAAFFD32" authorityName="Cuvier" authorityYear="1816" box="[1200,1301,683,707]" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
, supposedly belonging to the
<taxonomicName id="4C34B7CEFE44FFAFF8A8FD10FB00FD12" authority="Lineage" authorityName="Lineage" box="[1015,1210,715,739]" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE44FFAFF8A8FD10FBECFD12" box="[1015,1110,715,739]" italics="true" pageId="13" pageNumber="14">Priolepis</emphasis>
Lineage
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, with the following combination of diagnostic characters. Otolith size small, up to slightly over
<quantity id="4CCC61A8FE44FFAFF8A3FCD0FBEAFCD2" box="[1020,1104,779,803]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="14" unit="mm" value="1.5">1.5 mm</quantity>
in length. Otolith shape horizontal rectangular to subrectangular. Anterior and posterior rims straight to slightly inclined. Inner and outer faces nearly equally convex or outer face more strongly though. Sulcus relatively short (OL:SuL = 2.03.2), wide, smoothened sole-shape or reniform, with low or without ostial lobe, without subcaudal iugum, inclined at 815°. Ventral furrow regularly curved, distant and diverging from ventral rim of otolith, connected around sulcus to distinct dorsal depression. Area around sulcus distinctly bulged.
</paragraph>
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<emphasis id="B940105FFE44FFAFF872FB57FC1AFB52" box="[813,928,1164,1187]" italics="true" pageId="13" pageNumber="14">Discussion</emphasis>
At first glance, the otoliths of
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<emphasis id="B940105FFE44FFAFFE4DFB50FCE2FB32" italics="true" pageId="13" pageNumber="14">Medoborichthys</emphasis>
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resemble those of several different lineages. Te otolith shape and characteristics resemble otoliths of the
<taxonomicName id="4C34B7CEFE44FFAFF872FB30FC68FAF2" authorityName="Schwarzhans" authorityYear="2014" box="[813,978,1259,1283]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="mustus">
<emphasis id="B940105FFE44FFAFF872FB30FC68FAF2" box="[813,978,1259,1283]" italics="true" pageId="13" pageNumber="14">Gobius mustus</emphasis>
</taxonomicName>
complex described above, but
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<emphasis id="B940105FFE44FFAFFE63FB30FC39FAD2" italics="true" pageId="13" pageNumber="14">Medoborichthys</emphasis>
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otoliths differ in the lack of a subcaudal iugum, the reduced sole-shaped or reniform sulcus, the shape of the ventral furrow, and the distinctly bulged area around the sulcus. Otoliths of the Ponto-Caspian genus
<taxonomicName id="4C34B7CEFE44FFAFFE17FAB7FCDCFA52" genus="Capsiosoma" pageId="13" pageNumber="14" rank="genus">
<emphasis id="B940105FFE44FFAFFE17FAB7FCDCFA52" italics="true" pageId="13" pageNumber="14">Capsiosoma</emphasis>
</taxonomicName>
resemble
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<emphasis id="B940105FFE44FFAFF884FA50FB32FA52" box="[987,1160,1419,1443]" italics="true" pageId="13" pageNumber="14">Medoborichthys</emphasis>
</taxonomicName>
in the shape of the otolith, the reduced sulcus shape, and the lack of a subcaudal iugum but show neither the deep ventral furrow and dorsal depression nor the bulged area around the sulcus.
<taxonomicName id="4C34B7CEFE44FFAFF872F9D0FC60F9D2" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[813,986,1547,1571]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE44FFAFF872F9D0FC60F9D2" box="[813,986,1547,1571]" italics="true" pageId="13" pageNumber="14">Medoborichthys</emphasis>
</taxonomicName>
resembles otoliths of the
<taxonomicName id="4C34B7CEFE44FFAFFFA4F9D0FCE9F9B5" authority="Lineage" authorityName="Lineage" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE44FFAFFFA4F9D0FAE0F9D2" box="[1275,1370,1547,1571]" italics="true" pageId="13" pageNumber="14">Priolepis</emphasis>
Lineage
</taxonomicName>
, particularly of the genus
<taxonomicName id="4C34B7CEFE44FFAFFF2CF9F0FB68F9B2" box="[1139,1234,1579,1603]" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE44FFAFFF2CF9F0FB68F9B2" box="[1139,1234,1579,1603]" italics="true" pageId="13" pageNumber="14">Priolepis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFA5B1BCFE44FFAFFE52F9F0FC15F992" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." pageId="13" pageNumber="14" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
for photos of extant species), in the reduced sulcus shape, the lack of a subcaudal iugum, and the ventral furrow curving at a distance from the ventral rim of the otoliths and being connected to the dorsal depression and the bulged area around the sulcus. It differs in the less reduced sulcus, particularly in
<taxonomicName id="4C34B7CEFE44FFAFFFE0F937FAF4F8F2" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[1215,1358,1771,1795]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="podolicus" status="n. sp.">
<emphasis id="B940105FFE44FFAFFFE0F937FAF4F8F2" box="[1215,1358,1771,1795]" italics="true" pageId="13" pageNumber="14">M. podolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE44FFAFFE06F937FA22F8F2" box="[1369,1432,1772,1795]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
, and the less narrowed dorsal depression, which in extant species almost forms a continuous furrow connected to the ventral furrow. Tis character state could be regarded as plesiomorphic in respect to the modern genera of the lineage. Extant members of the
<taxonomicName id="4C34B7CEFE47FFACFAADFED0FD17FED2" authority="Lineage" authorityName="Lineage" box="[498,685,267,291]" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE47FFACFAADFED0FDEBFED2" box="[498,593,267,291]" italics="true" pageId="14" pageNumber="15">Priolepis</emphasis>
Lineage
</taxonomicName>
are also mostly associated with reefoidal environments. Due to the uncertainties concerning the sulcus shape, we allocate
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only tentatively to the
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Lineage.
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<emphasis id="B940105FFE47FFACF872FF37FCC0FEF2" box="[813,890,236,259]" italics="true" pageId="14" pageNumber="15">Species</emphasis>
Two species, both from the late Badenian of western
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in the back-reef facies of the Medobory barrier reef:
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<emphasis id="B940105FFE47FFACF8E5FEF0FB6DFEB2" box="[954,1239,299,323]" italics="true" pageId="14" pageNumber="15">Medoborichthys podolicus</emphasis>
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and
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<emphasis id="B940105FFE47FFACFE09FEF0FBB1FE92" italics="true" pageId="14" pageNumber="15">Medoborichthys renesulcis</emphasis>
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@ -0,0 +1,111 @@
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<emphasis id="B940105FFE44FFAFFBC8FC50FE9CFC52" bold="true" box="[151,294,907,931]" italics="true" pageId="13" pageNumber="14">Bathygobius</emphasis>
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? sp.
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<figureCitation id="130FD0C8FE44FFAFFBC8FC10FF4CFC12" box="[151,246,971,995]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="13" pageNumber="14">Figure 5j</figureCitation>
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<emphasis id="B940105FFE44FFAFFBC8FBD0FF42FBD2" box="[151,248,1035,1059]" italics="true" pageId="13" pageNumber="14">Material</emphasis>
1 otolith, Kozatskyi Yar,
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<collectionCode id="ED255488FE44FFAFF9D4FBD7FD19FBD2" box="[651,675,1036,1059]" country="Russia" name="Paleontological Institute" pageId="13" pageNumber="14">PI</collectionCode>
2548.
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<emphasis id="B940105FFE44FFAFFBC8FB97FEAEFB92" box="[151,276,1100,1123]" italics="true" pageId="13" pageNumber="14">Description</emphasis>
Te single, rather well-preserved small otolith of
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in length is characterized as follows. OL:OH = 1.05; OH:OT = 2.5. Ventral and posterior rim straight, meeting in orthogonal angle; postdorsal projection not expanded. Anterior rim slightly inclined at about 80°, straight. Dorsal rim with obtuse, depressed predorsal angle, ascending to broad postdorsal angle followed by short, slightly declining postdorsal section.
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<paragraph id="8B8BCC4DFE44FFAFFBF2FA97FD88F992" blockId="13.[151,776,1099,1635]" pageId="13" pageNumber="14">Inner face slightly convex, with large, wide, slightly deepened, distinctly inclined (20°) centrally positioned sulcus with moderate ostial lobe and no subcaudal iugum. OL:SuL = 1.55; SuL:SuH = 1.85. Ventral furrow deep and distinct, curving more regular than ventral rim of otolith and thus clipping pre- and postventral angles. Dorsal depression wide and deep with distinct crista superior toward sulcus and dorsally open. Outer face more convex than inner face, smooth.</paragraph>
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<emphasis id="B940105FFE44FFAFFBC8F957FEB0F952" box="[151,266,1676,1699]" italics="true" pageId="13" pageNumber="14">Discussion</emphasis>
Te large and wide sulcus of this otolith is quite remarkable and resembles the morphotype found in otoliths of extant species of
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(see
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for figures). However, extant
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</mods:mods>
<treatment id="039D7D5BFE45FFAEFBC8FEB0FC7AFCD2" LSID="urn:lsid:plazi:treatment:039D7D5BFE45FFAEFBC8FEB0FC7AFCD2" httpUri="http://treatment.plazi.org/id/039D7D5BFE45FFAEFBC8FEB0FC7AFCD2" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="C32E9FC6FE45FFAEFBC8FEB0FE5BFE72" box="[151,481,363,387]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B8BCC4DFE45FFAEFBC8FEB0FE5BFE72" blockId="12.[151,481,363,387]" box="[151,481,363,387]" pageId="12" pageNumber="13">
<heading id="D0C37B21FE45FFAEFBC8FEB0FE5BFE72" bold="true" box="[151,481,363,387]" fontSize="10" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B940105FFE45FFAEFBC8FEB0FE5BFE72" bold="true" box="[151,481,363,387]" pageId="12" pageNumber="13">
<taxonomicName id="4C34B7CEFE45FFAEFBC8FEB0FE25FE72" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[151,415,363,387]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="inauditus" status="sp. nov.">
<emphasis id="B940105FFE45FFAEFBC8FEB0FE25FE72" bold="true" box="[151,415,363,387]" italics="true" pageId="12" pageNumber="13">Parenypnias inauditus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE45FFAEFAFBFEB0FE5BFE72" box="[420,481,363,387]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE45FFAEFBC8FE70FC3CFE92" pageId="12" pageNumber="13" type="description">
<paragraph id="8B8BCC4DFE45FFAEFBC8FE70FEA0FE32" blockId="12.[151,282,427,451]" box="[151,282,427,451]" pageId="12" pageNumber="13">
<figureCitation id="130FD0C8FE45FFAEFBC8FE70FEA0FE32" box="[151,282,427,451]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="12" pageNumber="13">Figure 5eh</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBC8FE30FE1BFDD2" blockId="12.[151,775,491,547]" pageId="12" pageNumber="13">
2020a
<taxonomicName id="4C34B7CEFE45FFAEFBACFE30FE35FDF2" box="[243,399,491,515]" family="Gobiidae" genus="Ponticola" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE45FFAEFBACFE30FEE3FDF2" box="[243,345,491,515]" italics="true" pageId="12" pageNumber="13">Ponticola</emphasis>
sp.
</taxonomicName>
—Schwarzhans, Brzobohatý &amp; Radwańska: pl. 4, fig. 18.
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBC8FD90FE5EFD72" blockId="12.[151,775,587,643]" pageId="12" pageNumber="13">
<materialsCitation id="3B5CC610FE45FFAEFBC8FD90FE5EFD72" collectionCode="NMNHU-P" country="Ukraine" location="Kozatskyi Yar" municipality="Badenian" pageId="12" pageNumber="13" specimenCode="PI 2550" specimenCount="5" typeStatus="holotype">
<emphasis id="B940105FFE45FFAEFBC8FD90FF42FD92" box="[151,248,587,611]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="548F72EFFE45FFAEFBC8FD90FF42FD92" box="[151,248,587,611]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
<collectionCode id="ED255488FE45FFAEFA59FD97FE3CFD92" box="[262,390,588,611]" pageId="12" pageNumber="13">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE45FFAEFACAFD97FE4BFD92" box="[405,497,587,611]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="12" pageNumber="13">PI 2550</specimenCode>
,
<location id="8EEB9A96FE45FFAEF95CFD90FD1AFD92" LSID="urn:lsid:plazi:treatment:039D7D5BFE45FFAEFBC8FEB0FC7AFCD2:8EEB9A96FE45FFAEF95CFD90FD1AFD92" box="[515,672,587,611]" country="Ukraine" municipality="Badenian" name="Kozatskyi Yar" pageId="12" pageNumber="13">Kozatskyi Yar</location>
, western
<collectingCountry id="F3238CDDFE45FFAEFBC8FDB0FF4BFD72" box="[151,241,619,643]" name="Ukraine" pageId="12" pageNumber="13">Ukraine</collectingCountry>
, late
<collectingMunicipality id="6BEF5637FE45FFAEFA77FDB0FE2AFD72" box="[296,400,619,643]" pageId="12" pageNumber="13">Badenian</collectingMunicipality>
,
<figureCitation id="130FD0C8FE45FFAEFAC5FDB0FE5AFD72" box="[410,480,619,643]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="12" pageNumber="13">
Fig.
<specimenCount id="9D3207C4FE45FFAEFA97FDB0FE5EFD72" box="[456,484,619,643]" count="5" pageId="12" pageNumber="13" type="generic">5f.</specimenCount>
</figureCitation>
</materialsCitation>
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBC8FD77FDE4FCF2" blockId="12.[151,775,683,771]" pageId="12" pageNumber="13">
<materialsCitation id="3B5CC610FE45FFAEFBC8FD77FF67FD12" collectionCode="NMB" location="Shydlivshchyna" pageId="12" pageNumber="13" specimenCode="P1207" specimenCount="3" typeStatus="paratype">
<emphasis id="B940105FFE45FFAEFBC8FD77FEB8FD32" box="[151,258,684,707]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="548F72EFFE45FFAEFBC8FD77FEB8FD32" box="[151,258,684,707]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
</emphasis>
4 otoliths:
<specimenCount id="9D3207C4FE45FFAEFA20FD70FDB3FD32" box="[383,521,683,707]" count="3" pageId="12" pageNumber="13" type="generic">3 specimens</specimenCount>
,
<location id="8EEB9A96FE45FFAEF94BFD70FD05FD32" LSID="urn:lsid:plazi:treatment:039D7D5BFE45FFAEFBC8FEB0FC7AFCD2:8EEB9A96FE45FFAEF94BFD70FD05FD32" box="[532,703,683,707]" name="Shydlivshchyna" pageId="12" pageNumber="13">Shydlivshchyna</location>
,
<collectionCode id="ED255488FE45FFAEF994FD77FCBDFD32" box="[715,775,684,707]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="12" pageNumber="13">NMB</collectionCode>
<specimenCode id="DB926436FE45FFAEFBC8FD10FF67FD12" box="[151,221,715,739]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="12" pageNumber="13">P1207</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE45FFAEFBB6FD10FDE0FCF2" collectionCode="MU" country="Czech Republic" elevation="1619" location="Badenian" municipality="Zizkov" pageId="12" pageNumber="13" specimenCode="MU-0395" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D3207C4FE45FFAEFBB6FD10FED3FD12" box="[233,361,715,739]" count="1" pageId="12" pageNumber="13" type="generic">1 specimen</specimenCount>
, late
<location id="8EEB9A96FE45FFAEFAF9FD10FDB4FD12" LSID="urn:lsid:plazi:treatment:039D7D5BFE45FFAEFBC8FEB0FC7AFCD2:8EEB9A96FE45FFAEFAF9FD10FDB4FD12" box="[422,526,715,739]" country="Czech Republic" municipality="Zizkov" name="Badenian" pageId="12" pageNumber="13">Badenian</location>
,
<collectingMunicipality id="6BEF5637FE45FFAEF944FD1DFDDCFD12" box="[539,614,710,739]" pageId="12" pageNumber="13">Žižkov</collectingMunicipality>
1 well (
<quantity id="4CCC61A8FE45FFAEF99DFD10FF50FCF2" metricMagnitude="3" metricUnit="m" metricValue="1.619" metricValueMax="1.6219999999999999" metricValueMin="1.616" pageId="12" pageNumber="13" unit="m" value="1619.0" valueMax="1622.0" valueMin="1616.0">
<elevation id="00192B7EFE45FFAEF99DFD10FF50FCF2" metricMagnitude="3" metricUnit="m" metricValue="1.619" metricValueMax="1.6219999999999999" metricValueMin="1.616" pageId="12" pageNumber="13" unit="m" value="1619.0" valueMax="1622.0" valueMin="1616.0">1616 1622 m</elevation>
</quantity>
),
<collectingCountry id="F3238CDDFE45FFAEFBA3FD30FE12FCF2" box="[252,424,747,771]" name="Czech Republic" pageId="12" pageNumber="13">Czech Republic</collectingCountry>
, DGS
<specimenCode id="DB926436FE45FFAEFAB1FD30FDE0FCF2" box="[494,602,747,771]" collectionCode="MU-" pageId="12" pageNumber="13">MU-0395</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBC8FCF0FDEAFC72" blockId="12.[151,775,810,900]" pageId="12" pageNumber="13">
<emphasis id="B940105FFE45FFAEFBC8FCF0FEB2FCB2" box="[151,264,811,835]" italics="true" pageId="12" pageNumber="13">Etymology</emphasis>
From
<taxonomicName id="4C34B7CEFE45FFAEFA01FCF0FDA1FCB2" authority="(Latin)" baseAuthorityName="Latin" box="[350,539,811,835]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="inauditus">inauditus (Latin)</taxonomicName>
= unbeknown, referring to the occurrence of the species outside of the extant distribution range of the Gobiosomatini.
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBC8FC77FDBFFB72" blockId="12.[151,775,938,1155]" pageId="12" pageNumber="13">
<emphasis id="B940105FFE45FFAEFBC8FC77FEBBFC32" box="[151,257,940,963]" italics="true" pageId="12" pageNumber="13">Diagnosis</emphasis>
OL:OH = 1.151.22. Preventral projection pointed; postdorsal projection moderately long, pointed, slightly to moderately bent outward. Dorsal rim regularly curved but with pointed middorsal denticle. Sulcus narrow, short (OL:SuL = 2.22.5), inclined at 1215°. Ostial lobe absent or very weak; no subcaudal iugum. Inner face equal or less bent than outer face.
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBC8FB77FD90F972" blockId="12.[151,776,1195,1923]" pageId="12" pageNumber="13">
<emphasis id="B940105FFE45FFAEFBC8FB77FEAEFB32" box="[151,276,1196,1219]" italics="true" pageId="12" pageNumber="13">Description</emphasis>
Moderately slender, moderately thin, small otoliths reaching
<quantity id="4CCC61A8FE45FFAEFAF1FB10FDB3FB12" box="[430,521,1227,1251]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="13" unit="mm" value="1.6">1.6 mm</quantity>
in length (
<typeStatus id="548F72EFFE45FFAEF9C8FB10FD46FB12" box="[663,764,1227,1251]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
). OH:OT = 2.73.0. Anterior rim inclined at 7783°, with distinct concavity at about level of ostial tip and distinct, mostly sharp and moderately long preventral projection. Predorsal angle distinct, angular, somewhat projecting, positioned moderately high on anterior rim. Ventral rim nearly straight, smooth, slightly undulating in smallest specimen (
<figureCitation id="130FD0C8FE45FFAEFA51FA70FEE1FA32" box="[270,347,1451,1475]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="12" pageNumber="13">Fig. 5h</figureCitation>
). Dorsal rim overall relatively regularly curved, sometimes slightly undulating (
<figureCitation id="130FD0C8FE45FFAEF919FA10FD28FA12" box="[582,658,1483,1507]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="12" pageNumber="13">Fig. 5h</figureCitation>
), with distinct angular or projecting denticle in middorsal position, with distinct moderately strong, sharp postdorsal projection, slightly to moderately bent outward. Posterior rim with distinct incision below postdorsal projection and variably developed postventral angle.
</paragraph>
<paragraph id="8B8BCC4DFE45FFAEFBF2F957FC3CFE92" blockId="12.[151,776,1195,1923]" lastBlockId="12.[813,1437,235,355]" pageId="12" pageNumber="13">Inner face slightly convex to nearly flat, with very narrow, slightly deepened, centrally positioned sulcus. SuL:SuH = 2.63.2. Sulcus without discernable ostial lobe, oval-shaped, with rounded tips, inclined at 1215°. No subcaudal iugum. Ventral furrow broad, distinct, reaching from anterior to posterior tips of sulcus, curvature more or less following ventral rim of otolith. Dorsal field with small, indistinct dorsal depression, ventrally marked by feeble crista superior toward sulcus and dorsally closed to dorsal rim of otolith. Outer face convex, equally or more convex than inner face, regularly shaped, smooth.</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE45FFAEF872FE57FC7AFCD2" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8B8BCC4DFE45FFAEF872FE57FC7AFCD2" blockId="12.[813,1437,395,803]" pageId="12" pageNumber="13">
<emphasis id="B940105FFE45FFAEF872FE57FB11FE52" box="[813,1195,395,419]" italics="true" pageId="12" pageNumber="13">
Discussion
<taxonomicName id="4C34B7CEFE45FFAEF8F1FE57FB11FE52" box="[942,1195,395,419]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="inauditus">Parenypnias inauditus</taxonomicName>
</emphasis>
is easily recognized by certain features of the outline of the otolith, such as the sharply pointed preventral projection, the pointed postdorsal projection, and the middorsal denticle, which, however, is easily affected by erosion and also seems to be developed more like a middorsal angle in specimens smaller than
<quantity id="4CCC61A8FE45FFAEF8E5FD90FBA3FD92" box="[954,1049,587,611]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="12" pageNumber="13" unit="mm" value="1.25">1.25 mm</quantity>
in length (
<figureCitation id="130FD0C8FE45FFAEFFD2FD90FB48FD92" box="[1165,1266,587,612]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="12" pageNumber="13">
Fig.
<quantity id="4CCC61A8FE45FFAEFFE3FD90FB62FD95" box="[1212,1240,587,612]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="12" pageNumber="13" unit="g" value="5.0">5g</quantity>
, h
</figureCitation>
). It differs from the parallel occurring
<taxonomicName id="4C34B7CEFE45FFAEFF76FDB7FB2FFD72" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[1065,1173,619,643]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kiselevi" status="n. sp.">
<emphasis id="B940105FFE45FFAEFF76FDB7FB2FFD72" box="[1065,1173,619,643]" italics="true" pageId="12" pageNumber="13">P. kiselevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE45FFAEFFC0FDB7FB67FD72" box="[1183,1245,620,643]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
described below in the longer sulcus (OL:SuL = 2.02.5 vs. 2.0), the more pronounced preventral projection, the less strongly outward bent postdorsal projection, the sharp otolith rims (vs. thick), and the more slender shape (OL:OH = 1.15 1.22 vs. 1.05).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B940105FFE45FFAEF872FC90FBA3FC92" bold="true" box="[813,1049,843,867]" italics="true" pageId="12" pageNumber="13">Parenypnias kiselevi</emphasis>
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<figureCitation id="130FD0C8FE45FFAEF872FC50FC1CFC52" box="[813,934,907,931]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="12" pageNumber="13">Figure 5h, i</figureCitation>
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,
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, western
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,
<figureCitation id="130FD0C8FE45FFAEF893FC30FBA2FBF2" box="[972,1048,1003,1027]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="12" pageNumber="13">Fig. 5h</figureCitation>
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<emphasis id="B940105FFE45FFAEF872FBF0FC24FBB2" box="[813,926,1067,1091]" italics="true" pageId="12" pageNumber="13">Etymology</emphasis>
Named after Iosif Kiselev, enthusiastic paleoichthyologist amateur who collected a number of fish fossils of different ages in
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<emphasis id="B940105FFE45FFAEF872FB77FC2DFB32" box="[813,919,1196,1219]" italics="true" pageId="12" pageNumber="13">Diagnosis</emphasis>
OL:OH = 1.05. Preventral projection blunt, short; postdorsal projection moderately long, blunt, very strongly bent outward. Dorsal rim ascending to sharp postdorsal angle. Sulcus narrow, short (OL:SuL = 2.0), inclined at 15°. Ostial lobe absent; no subcaudal iugum. Inner face more strongly bent than outer face.
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<emphasis id="B940105FFE45FFAEF872FA57FC10FA52" box="[813,938,1420,1443]" italics="true" pageId="12" pageNumber="13">Description</emphasis>
Small, thick otolith of
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in length (
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). OH:OT = 2.4. Anterior rim vertical, with distinct concavity at about level of ostial tip and distinct preventral angle and relatively low positioned predorsal angle of about equal strength. Ventral rim nearly straight. Dorsal rim ascending straight to sharp postdorsal angle positioned above cauda, followed by slightly concave postdorsal rim toward strongly outward bent postdorsal projection. Posterior rim with distinct and deep incision below postdorsal projection and orthogonal postventral angle. Ventral rim smooth, dorsal rim anteriorly slightly crenulated. Rims very thick and massive.
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<paragraph id="8B8BCC4DFE45FFAFF81DF8D7FE8BFE32" blockId="12.[813,1437,1419,1923]" lastBlockId="13.[151,775,235,451]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">Inner face markedly convex, with very narrow, deepened, centrally positioned sulcus at 15°. OL:SuL= 2.0; SuL:SuH = 2.5. Sulcus without discernable ostial lobe, oval-shaped, with rounded tips, inclined at 15°. No subcaudal iugum. Ventral furrow broad, distinct, reaching from anterior to posterior tips of sulcus, curvature more or less following ventral rim of otolith. Dorsal field with small, indistinct dorsal depression ventrally marked by feeble crista superior toward sulcus and dorsally closed to dorsal rim of otolith. Outer face nearly flat, relatively smooth.</paragraph>
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<emphasis id="B940105FFE44FFAFFBC8FE37FEB0FDF2" box="[151,266,492,515]" italics="true" pageId="13" pageNumber="14">Discussion</emphasis>
Tis small otolith is easily recognized by a number of unique characters, not the least of which are the very thick rims, strongly outward bent postdorsal projection, and small, almost oval sulcus. Te morphological expression is that of a mature fish, and we, therefore, assume that the otolith stems from a small species. For distinction from the second
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species
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, see above.
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<heading id="D0C37B21FE46FFADF872FE70FB1AFE32" bold="true" box="[813,1184,427,451]" fontSize="10" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B940105FFE46FFADF872FE70FB1AFE32" bold="true" box="[813,1184,427,451]" pageId="15" pageNumber="16">
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<emphasis id="B940105FFE46FFADF872FE70FBE7FE32" bold="true" box="[813,1117,427,451]" italics="true" pageId="15" pageNumber="16">Medoborichthys renesulcis</emphasis>
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<paragraph id="8B8BCC4DFE46FFADF872FE30FC17FDF5" blockId="15.[813,941,491,516]" box="[813,941,491,516]" pageId="15" pageNumber="16">
<figureCitation id="130FD0C8FE46FFADF872FE30FC17FDF5" box="[813,941,491,516]" captionStart="Fig" captionStartId="14.[169,200,1796,1816]" captionTargetBox="[152,1438,540,1933]" captionTargetId="figure-92@14.[168,1423,556,1783]" captionTargetPageId="14" captionText="Fig. 6 Otoliths of Medoborichthys and Vanderhorstia: ad Medoborichthys podolicus n. gen. et n. sp., b holotype, NMNHU-P PI 2563, Mlyntsi, a, c, d paratypes,a Mlyntsi, NMNHU-P PI 2564,c Shydlivshchyna,NMNHU-P PI 2565, d (reversed) Kozatskyi Yar, NMB P1215. eg Medoborichthys renesulcis n. gen. et n. sp., e holotype, NMNHU-P PI 2566, Mlyntsi,f, g paratypes,f Kozatskyi Yar, NMB P1216, g (reversed) Mlyntsi, NMNHU-P PI 2567. hj Vanderhorstia prochazkai Schwarzhans et al., 2020a, 2020b, h (reversed), j (reversed) Mlyntsi, NMB P1223, h (reversed) Shydlivshchyna, NMNHU-P PI 2582. k Vanderhorstia sp., Shydlivshchyna, NMB P1224" figureDoi="http://doi.org/10.5281/zenodo.11999907" httpUri="https://zenodo.org/record/11999907/files/figure.png" pageId="15" pageNumber="16">Figure 6eg</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE46FFADF872FDF0FBA3FD92" blockId="15.[813,1436,555,611]" pageId="15" pageNumber="16">
<materialsCitation id="3B5CC610FE46FFADF872FDF0FBA3FD92" collectionCode="NMNHU-P" country="Ukraine" location="Mlyntsi" municipality="Badenian" pageId="15" pageNumber="16" specimenCode="PI 2566" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548F72EFFE46FFADF872FDF0FC34FDB2" box="[813,910,555,579]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
<collectionCode id="ED255488FE46FFADF8CAFDF7FBAFFDB2" box="[917,1045,556,579]" pageId="15" pageNumber="16">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE46FFADFF41FDF7FBCEFDB2" box="[1054,1140,555,579]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="15" pageNumber="16">PI 2566</specimenCode>
,
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, western
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, late
<collectingMunicipality id="6BEF5637FE46FFADF805FD90FC78FD92" box="[858,962,587,611]" pageId="15" pageNumber="16">Badenian</collectingMunicipality>
,
<figureCitation id="130FD0C8FE46FFADF893FD90FBAFFD92" box="[972,1045,587,611]" captionStart="Fig" captionStartId="14.[169,200,1796,1816]" captionTargetBox="[152,1438,540,1933]" captionTargetId="figure-92@14.[168,1423,556,1783]" captionTargetPageId="14" captionText="Fig. 6 Otoliths of Medoborichthys and Vanderhorstia: ad Medoborichthys podolicus n. gen. et n. sp., b holotype, NMNHU-P PI 2563, Mlyntsi, a, c, d paratypes,a Mlyntsi, NMNHU-P PI 2564,c Shydlivshchyna,NMNHU-P PI 2565, d (reversed) Kozatskyi Yar, NMB P1215. eg Medoborichthys renesulcis n. gen. et n. sp., e holotype, NMNHU-P PI 2566, Mlyntsi,f, g paratypes,f Kozatskyi Yar, NMB P1216, g (reversed) Mlyntsi, NMNHU-P PI 2567. hj Vanderhorstia prochazkai Schwarzhans et al., 2020a, 2020b, h (reversed), j (reversed) Mlyntsi, NMB P1223, h (reversed) Shydlivshchyna, NMNHU-P PI 2582. k Vanderhorstia sp., Shydlivshchyna, NMB P1224" figureDoi="http://doi.org/10.5281/zenodo.11999907" httpUri="https://zenodo.org/record/11999907/files/figure.png" pageId="15" pageNumber="16">Fig. 6e</figureCitation>
.
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<materialsCitation id="3B5CC610FE46FFADF872FD57FBFEFD32" collectionCode="NMNHU-P" country="Ukraine" location="Mlyntsi" municipality="Badenian" pageId="15" pageNumber="16" specimenCode="PI 2567" specimenCount="4" typeStatus="paratype">
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<typeStatus id="548F72EFFE46FFADF872FD57FC22FD52" box="[813,920,652,675]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
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6 otoliths:
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same data as holotype,
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<specimenCode id="DB926436FE46FFADF8AEFD77FBFEFD32" box="[1009,1092,683,707]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="15" pageNumber="16">PI 2567</specimenCode>
</materialsCitation>
<materialsCitation id="3B5CC610FE46FFADFF14FD70FABEFD32" box="[1099,1284,683,707]" collectionCode="NMB" country="Ukraine" pageId="15" pageNumber="16" specimenCode="P1217" specimenCount="1" typeStatus="paratype">
and
<collectionCode id="ED255488FE46FFADFF24FD77FB0DFD32" box="[1147,1207,684,707]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="15" pageNumber="16">NMB</collectionCode>
<specimenCode id="DB926436FE46FFADFFE0FD70FABEFD32" box="[1215,1284,683,707]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="15" pageNumber="16">P1217</specimenCode>
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;
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<specimenCount id="9D3207C4FE46FFADFE50FD70FA22FD32" box="[1295,1432,683,707]" count="2" pageId="15" pageNumber="16" type="generic">2 specimens</specimenCount>
, Kozatskyi Yar,
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<specimenCode id="DB926436FE46FFADFF51FD10FBEEFD12" box="[1038,1108,715,739]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="15" pageNumber="16">P1216</specimenCode>
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.
</paragraph>
<paragraph id="8B8BCC4DFE46FFADF872FCD0FCCDFC92" blockId="15.[813,1437,778,867]" pageId="15" pageNumber="16">
<emphasis id="B940105FFE46FFADF872FCD0FC24FCD2" box="[813,926,779,803]" italics="true" pageId="15" pageNumber="16">Etymology</emphasis>
From reneformis (Latin) = reniform in combination with sulcus indicating the reniform shape of the sulcus.
</paragraph>
<paragraph id="8B8BCC4DFE46FFADF872FC57FB88FB52" blockId="15.[813,1437,906,1187]" pageId="15" pageNumber="16">
<emphasis id="B940105FFE46FFADF872FC57FC2DFC52" box="[813,919,908,931]" italics="true" pageId="15" pageNumber="16">Diagnosis</emphasis>
OL:OH = 1.15. Anterior rim inclined at about 80° with depressed predorsal angle; posterior rim with weak developed postdorsal projection. Dorsal rim anteriorly depressed, straight ascending to postdorsal angle. Sulcus small, OL:SuL = 2.33.2, inclined at 810°. Sulcus reniform; no subcaudal iugum. Ventral furrow deep, distant and divergent from ventral otolith rim and connected around sulcus to dorsal depression; area around sulcus distinctly bulged.
</paragraph>
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<emphasis id="B940105FFE46FFADF872FB17FC10FB12" box="[813,938,1228,1251]" italics="true" pageId="15" pageNumber="16">Description</emphasis>
Moderately compressed, subrectangular, massive otoliths reaching about
<quantity id="4CCC61A8FE46FFADFFD4FB30FB51FAF2" box="[1163,1259,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="15" pageNumber="16" unit="mm" value="1.45">1.45 mm</quantity>
in length (
<typeStatus id="548F72EFFE46FFADFE3EFB30FCDAFAD2" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
). OH:OT = 2.22.4. Anterior rim slightly inclined, without concavity, and with short preventral angle. Predorsal angle obtuse, depressed. Ventral rim nearly straight, smooth. Dorsal rim relatively straight, anteriorly depressed and ascending to short, rounded postdorsal projection without or with very feeble postdorsal angle. Posterior rim with concavity below blunt postdorsal projection and orthogonal postventral angle.
</paragraph>
<paragraph id="8B8BCC4DFE46FFADF81DF9D7FC0FF892" blockId="15.[813,1437,1227,1891]" pageId="15" pageNumber="16">Inner face moderately convex, with area around sulcus, interior of ventral furrow, and dorsal depression distinctly bulged. Sulcus wide, relatively shallow, centrally positioned, moderately inclined at 810°. OL:SuL = 2.3 3.2; SuL:SuH = 1.82.0. Sulcus reniform without ostial lobe and without subcaudal iugum. Ventral furrow broad, deep, distinct, regularly curved, distant and divergent from ventral rim of otolith, connected around sulcus to narrow and deep dorsal depression. Outer face convex, more convex than inner face and very regularly shaped and smooth.</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE59FFB2FBC8FF37FE37FD32" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B8BCC4DFE59FFB2FBC8FF37FE37FD32" blockId="16.[151,775,235,707]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2FBC8FF37FD81FEF2" box="[151,571,235,259]" italics="true" pageId="16" pageNumber="17">
Discussion
<taxonomicName id="4C34B7CEFE59FFB2FA47FF30FD81FEF2" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[280,571,235,259]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="renesulcis">Medoborichthys renesulcis</taxonomicName>
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is readily distinguishable from
<taxonomicName id="4C34B7CEFE59FFB2FA15FED7FE63FED2" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[330,473,267,291]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="podolicus">
<emphasis id="B940105FFE59FFB2FA15FED7FE63FED2" box="[330,473,267,291]" italics="true" pageId="16" pageNumber="17">M. podolicus</emphasis>
</taxonomicName>
by the shapes of the otolith and the sulcus, with both character states being considered as apomorphic and closer to extant otoliths of
<taxonomicName id="4C34B7CEFE59FFB2FBE7FEB0FEADFE72" box="[184,279,363,387]" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE59FFB2FBE7FEB0FEADFE72" box="[184,279,363,387]" italics="true" pageId="16" pageNumber="17">Priolepis</emphasis>
</taxonomicName>
. However, Schwarzhans et. al. (2020a) also recorded a single, incomplete otolith of
<taxonomicName id="4C34B7CEFE59FFB2F924FE50FCBDFE52" box="[635,775,395,419]" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE59FFB2F924FE50FD60FE52" box="[635,730,395,419]" italics="true" pageId="16" pageNumber="17">Priolepis</emphasis>
sp.
</taxonomicName>
from the early Badenian of Korytnica,
<collectingCountry id="F3238CDDFE59FFB2F900FE70FD16FE32" box="[607,684,427,451]" name="Poland" pageId="16" pageNumber="17">Poland</collectingCountry>
, that is similar to
<taxonomicName id="4C34B7CEFE59FFB2FA56FE17FE22FE12" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[265,408,459,483]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="renesulcis">
<emphasis id="B940105FFE59FFB2FA56FE17FE22FE12" box="[265,408,459,483]" italics="true" pageId="16" pageNumber="17">M. renesulcis</emphasis>
</taxonomicName>
in most aspects but has an even more reduced sulcus morphology, is distinctly thinner (OH:OT = 2.9 vs. 2.22.4), and shows a less strongly bulged area around the sulcus. Te pattern of this otolith is considered more advanced and representative of a true member of the genus
<taxonomicName id="4C34B7CEFE59FFB2FADAFDB0FE5EFD72" box="[389,484,619,643]" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE59FFB2FADAFDB0FE5EFD72" box="[389,484,619,643]" italics="true" pageId="16" pageNumber="17">Priolepis</emphasis>
</taxonomicName>
; thus, the otolith indicates that the lineage was indeed present in the middle Miocene of the Paratethys.
</paragraph>
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</document>

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<mods:number id="89C1C5861CA47552787C1F088C20AA63">2022-11-17</mods:number>
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<mods:number id="B5035DBD2097BF2DAF50081265145FB3">141</mods:number>
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<mods:url id="768211DD751836683BA8CCD6159E854A">http://dx.doi.org/10.1186/s13358-022-00261-3</mods:url>
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<mods:identifier id="3546DAACE6FD1221042F9A5088B11EDD" type="DOI">10.1186/s13358-022-00261-3</mods:identifier>
<mods:identifier id="CE6207B0ECD70B5E9649031CC143ED2E" type="ISSN">1664-2384</mods:identifier>
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<treatment id="039D7D5BFE47FFADF872FE50FB1AFE72" LSID="urn:lsid:plazi:treatment:039D7D5BFE47FFADF872FE50FB1AFE72" httpUri="http://treatment.plazi.org/id/039D7D5BFE47FFADF872FE50FB1AFE72" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="C32E9FC6FE47FFACF872FE50FB26FE52" box="[813,1180,395,419]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8B8BCC4DFE47FFACF872FE50FB26FE52" blockId="14.[813,1180,395,419]" box="[813,1180,395,419]" pageId="14" pageNumber="15">
<heading id="D0C37B21FE47FFACF872FE50FB26FE52" bold="true" box="[813,1180,395,419]" fontSize="10" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B940105FFE47FFACF872FE50FB26FE52" bold="true" box="[813,1180,395,419]" pageId="14" pageNumber="15">
<taxonomicName id="4C34B7CEFE47FFACF872FE50FBE0FE52" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[813,1114,395,419]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="podolicus" status="sp. nov.">
<emphasis id="B940105FFE47FFACF872FE50FBE0FE52" bold="true" box="[813,1114,395,419]" italics="true" pageId="14" pageNumber="15">Medoborichthys podolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE47FFACFF3FFE50FB26FE52" box="[1120,1180,395,419]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<caption id="DF4B9CC5FE47FFACFBF6F8DFFDD4F872" ID-DOI="http://doi.org/10.5281/zenodo.11999907" ID-Zenodo-Dep="11999907" httpUri="https://zenodo.org/record/11999907/files/figure.png" pageId="14" pageNumber="15" startId="14.[169,200,1796,1816]" targetBox="[152,1438,540,1933]" targetPageId="14" targetType="figure">
<paragraph id="8B8BCC4DFE47FFACFBF6F8DFFDD4F872" blockId="14.[169,1402,1796,1923]" pageId="14" pageNumber="15">
<emphasis id="B940105FFE47FFACFBF6F8DFFF61F8E9" bold="true" box="[169,219,1796,1816]" pageId="14" pageNumber="15">Fig. 6</emphasis>
Otoliths of
<taxonomicName id="4C34B7CEFE47FFACFA1CF8DFFE7DF8E9" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[323,455,1796,1816]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE47FFACFA1CF8DFFE7DF8E9" box="[323,455,1796,1816]" italics="true" pageId="14" pageNumber="15">Medoborichthys</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34B7CEFE47FFACFAAEF8DFFDD9F8E9" authorityName="Smith" authorityYear="1949" box="[497,611,1796,1816]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE47FFACFAAEF8DFFDD9F8E9" box="[497,611,1796,1816]" italics="true" pageId="14" pageNumber="15">Vanderhorstia</emphasis>
</taxonomicName>
:
<emphasis id="B940105FFE47FFACF933F8DFFD37F8E9" bold="true" box="[620,653,1796,1816]" pageId="14" pageNumber="15">ad</emphasis>
<taxonomicName id="4C34B7CEFE47FFACF9CDF8DFFCD3F8E9" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[658,873,1796,1816]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="podolicus" status="n. gen. et n. sp.">
<emphasis id="B940105FFE47FFACF9CDF8DFFCD3F8E9" box="[658,873,1796,1816]" italics="true" pageId="14" pageNumber="15">Medoborichthys podolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE47FFACF832F8DFFC54F8E9" box="[877,1006,1796,1816]" pageId="14" pageNumber="15" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
,
<emphasis id="B940105FFE47FFACF8ACF8DFFC45F8E9" bold="true" box="[1011,1023,1796,1816]" pageId="14" pageNumber="15">b</emphasis>
holotype, NMNHU-P PI 2563, Mlyntsi,
<emphasis id="B940105FFE47FFACFE1EF8DFFAF6F8E9" bold="true" box="[1345,1356,1796,1816]" pageId="14" pageNumber="15">a</emphasis>
,
<emphasis id="B940105FFE47FFACFE0BF8DFFAE4F8E9" bold="true" box="[1364,1374,1796,1816]" pageId="14" pageNumber="15">c</emphasis>
,
<emphasis id="B940105FFE47FFACFE39F8DFFAC8F8E9" bold="true" box="[1382,1394,1796,1816]" pageId="14" pageNumber="15">d</emphasis>
paratypes,
<emphasis id="B940105FFE47FFACFA5BF8C4FEB5F8C2" bold="true" box="[260,271,1823,1843]" pageId="14" pageNumber="15">a</emphasis>
Mlyntsi, NMNHU-P PI 2564,
<emphasis id="B940105FFE47FFACFAA4F8C4FDBFF8C2" bold="true" box="[507,517,1823,1843]" pageId="14" pageNumber="15">c</emphasis>
Shydlivshchyna,NMNHU-P PI 2565,
<emphasis id="B940105FFE47FFACF867F8C4FCFEF8C2" bold="true" box="[824,836,1823,1843]" pageId="14" pageNumber="15">d</emphasis>
(reversed) Kozatskyi Yar, NMB P1215.
<emphasis id="B940105FFE47FFACFF21F8C4FB1AF8C2" bold="true" box="[1150,1184,1823,1843]" pageId="14" pageNumber="15">eg</emphasis>
<taxonomicName id="4C34B7CEFE47FFACFFFAF8C4FAC0F8C2" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[1189,1402,1823,1843]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="renesulcis" status="n. gen. et n. sp.">
<emphasis id="B940105FFE47FFACFFFAF8C4FAC0F8C2" box="[1189,1402,1823,1843]" italics="true" pageId="14" pageNumber="15">Medoborichthys renesulcis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE47FFACFBF6F8E2FE90F8BC" box="[169,298,1849,1869]" pageId="14" pageNumber="15" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
,
<emphasis id="B940105FFE47FFACFA70F8E2FE80F8BC" bold="true" box="[303,314,1849,1869]" pageId="14" pageNumber="15">e</emphasis>
holotype, NMNHU-P PI 2566, Mlyntsi,
<emphasis id="B940105FFE47FFACF924F8E2FD38F8BC" bold="true" box="[635,642,1849,1869]" pageId="14" pageNumber="15">f</emphasis>
,
<emphasis id="B940105FFE47FFACF9D4F8E2FD2DF8BC" bold="true" box="[651,663,1849,1869]" pageId="14" pageNumber="15">g</emphasis>
paratypes,
<emphasis id="B940105FFE47FFACF9A8F8E2FD44F8BC" bold="true" box="[759,766,1849,1869]" pageId="14" pageNumber="15">f</emphasis>
Kozatskyi Yar, NMB P1216,
<emphasis id="B940105FFE47FFACF8BFF8E2FC56F8BC" bold="true" box="[992,1004,1849,1869]" pageId="14" pageNumber="15">g</emphasis>
(reversed) Mlyntsi, NMNHU-P PI 2567.
<emphasis id="B940105FFE47FFACFE6EF8E2FAF4F8BC" bold="true" box="[1329,1358,1849,1869]" pageId="14" pageNumber="15">hj</emphasis>
<taxonomicName id="4C34B7CEFE47FFACFBF6F88FFDEFF899" authority="Schwarzhans et al., 2020 a" authorityName="Schwarzhans" authorityYear="2020" box="[169,597,1876,1896]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="prochazkai">
<emphasis id="B940105FFE47FFACFBF6F88FFEC2F899" box="[169,376,1876,1896]" italics="true" pageId="14" pageNumber="15">Vanderhorstia prochazkai</emphasis>
<bibRefCitation id="EFA5B1BCFE47FFACFA22F88FFDEFF899" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[381,597,1876,1896]" pageId="14" pageNumber="15" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFA5B1BCFE47FFACF902F88FFD28F899" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[605,658,1876,1896]" pageId="14" pageNumber="15" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
,
<emphasis id="B940105FFE47FFACF9C5F88FFD1CF899" bold="true" box="[666,678,1876,1896]" pageId="14" pageNumber="15">h</emphasis>
(reversed),
<emphasis id="B940105FFE47FFACF858F88FFCB7F899" bold="true" box="[775,781,1876,1896]" pageId="14" pageNumber="15">j</emphasis>
(reversed) Mlyntsi, NMB P1223,
<emphasis id="B940105FFE47FFACFF45F88FFB9CF899" bold="true" box="[1050,1062,1876,1896]" pageId="14" pageNumber="15">h</emphasis>
(reversed) Shydlivshchyna, NMNHU-P PI 2582.
<emphasis id="B940105FFE47FFACFB84F8B4FF5CF872" bold="true" box="[219,230,1903,1923]" pageId="14" pageNumber="15">k</emphasis>
<taxonomicName id="4C34B7CEFE47FFACFBB5F8B4FEC1F872" box="[234,379,1903,1923]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE47FFACFBB5F8B4FEE6F872" box="[234,348,1903,1923]" italics="true" pageId="14" pageNumber="15">Vanderhorstia</emphasis>
sp.
</taxonomicName>
, Shydlivshchyna, NMB P1224
</paragraph>
</caption>
<subSubSection id="C32E9FC6FE46FFADFBC8FF30FEC8F972" pageId="15" pageNumber="16" type="description">
<paragraph id="8B8BCC4DFE46FFADFBC8FF30FEA3FEF2" blockId="15.[151,281,235,259]" box="[151,281,235,259]" pageId="15" pageNumber="16">
<figureCitation id="130FD0C8FE46FFADFBC8FF30FEA3FEF2" box="[151,281,235,259]" captionStart="Fig" captionStartId="14.[169,200,1796,1816]" captionTargetBox="[152,1438,540,1933]" captionTargetId="figure-92@14.[168,1423,556,1783]" captionTargetPageId="14" captionText="Fig. 6 Otoliths of Medoborichthys and Vanderhorstia: ad Medoborichthys podolicus n. gen. et n. sp., b holotype, NMNHU-P PI 2563, Mlyntsi, a, c, d paratypes,a Mlyntsi, NMNHU-P PI 2564,c Shydlivshchyna,NMNHU-P PI 2565, d (reversed) Kozatskyi Yar, NMB P1215. eg Medoborichthys renesulcis n. gen. et n. sp., e holotype, NMNHU-P PI 2566, Mlyntsi,f, g paratypes,f Kozatskyi Yar, NMB P1216, g (reversed) Mlyntsi, NMNHU-P PI 2567. hj Vanderhorstia prochazkai Schwarzhans et al., 2020a, 2020b, h (reversed), j (reversed) Mlyntsi, NMB P1223, h (reversed) Shydlivshchyna, NMNHU-P PI 2582. k Vanderhorstia sp., Shydlivshchyna, NMB P1224" figureDoi="http://doi.org/10.5281/zenodo.11999907" httpUri="https://zenodo.org/record/11999907/files/figure.png" pageId="15" pageNumber="16">Figure 6ad</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE46FFADFBC8FEF0FE53FE92" blockId="15.[151,775,299,355]" pageId="15" pageNumber="16">
<materialsCitation id="3B5CC610FE46FFADFBC8FEF0FE53FE92" collectionCode="NMNHU-P" country="Ukraine" location="Kozatskyi Yar" municipality="Badenian" pageId="15" pageNumber="16" specimenCode="PI 2563" specimenCount="1" typeStatus="holotype">
<emphasis id="B940105FFE46FFADFBC8FEF0FF42FEB2" box="[151,248,299,323]" italics="true" pageId="15" pageNumber="16">
<typeStatus id="548F72EFFE46FFADFBC8FEF0FF42FEB2" box="[151,248,299,323]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
<collectionCode id="ED255488FE46FFADFA59FEF7FE3CFEB2" box="[262,390,300,323]" pageId="15" pageNumber="16">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE46FFADFACAFEF7FE4AFEB2" box="[405,496,299,323]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="15" pageNumber="16">PI 2563</specimenCode>
,
<location id="8EEB9A96FE46FFADF95CFEF0FD1AFEB2" LSID="urn:lsid:plazi:treatment:039D7D5BFE47FFADF872FE50FB1AFE72:8EEB9A96FE46FFADF95CFEF0FD1AFEB2" box="[515,672,299,323]" country="Ukraine" municipality="Badenian" name="Kozatskyi Yar" pageId="15" pageNumber="16">Kozatskyi Yar</location>
, western
<collectingCountry id="F3238CDDFE46FFADFBC8FE90FF4BFE92" box="[151,241,331,355]" name="Ukraine" pageId="15" pageNumber="16">Ukraine</collectingCountry>
, late
<collectingMunicipality id="6BEF5637FE46FFADFA77FE90FE2AFE92" box="[296,400,331,355]" pageId="15" pageNumber="16">Badenian</collectingMunicipality>
,
<figureCitation id="130FD0C8FE46FFADFAC5FE90FE5FFE92" box="[410,485,331,355]" captionStart="Fig" captionStartId="14.[169,200,1796,1816]" captionTargetBox="[152,1438,540,1933]" captionTargetId="figure-92@14.[168,1423,556,1783]" captionTargetPageId="14" captionText="Fig. 6 Otoliths of Medoborichthys and Vanderhorstia: ad Medoborichthys podolicus n. gen. et n. sp., b holotype, NMNHU-P PI 2563, Mlyntsi, a, c, d paratypes,a Mlyntsi, NMNHU-P PI 2564,c Shydlivshchyna,NMNHU-P PI 2565, d (reversed) Kozatskyi Yar, NMB P1215. eg Medoborichthys renesulcis n. gen. et n. sp., e holotype, NMNHU-P PI 2566, Mlyntsi,f, g paratypes,f Kozatskyi Yar, NMB P1216, g (reversed) Mlyntsi, NMNHU-P PI 2567. hj Vanderhorstia prochazkai Schwarzhans et al., 2020a, 2020b, h (reversed), j (reversed) Mlyntsi, NMB P1223, h (reversed) Shydlivshchyna, NMNHU-P PI 2582. k Vanderhorstia sp., Shydlivshchyna, NMB P1224" figureDoi="http://doi.org/10.5281/zenodo.11999907" httpUri="https://zenodo.org/record/11999907/files/figure.png" pageId="15" pageNumber="16">Fig. 6b</figureCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8BCC4DFE46FFADFBC8FE57FD45FE12" blockId="15.[151,775,395,483]" pageId="15" pageNumber="16">
<emphasis id="B940105FFE46FFADFBC8FE57FEB8FE52" box="[151,258,396,419]" italics="true" pageId="15" pageNumber="16">
<typeStatus id="548F72EFFE46FFADFBC8FE57FEB8FE52" box="[151,258,396,419]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
30 otoliths:
<materialsCitation id="3B5CC610FE46FFADFACCFE50FEDAFE32" collectionCode="NMB" country="Ukraine" location="Kozatskyi Yar" municipality="Badenian" pageId="15" pageNumber="16" specimenCode="P1215" specimenCount="12" typeStatus="paratype">
<specimenCount id="9D3207C4FE46FFADFACCFE50FD93FE52" box="[403,553,395,419]" count="12" pageId="15" pageNumber="16" type="generic">12 specimens</specimenCount>
same data as holotype,
<collectionCode id="ED255488FE46FFADFB8AFE77FEABFE32" box="[213,273,428,451]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="15" pageNumber="16">NMB</collectionCode>
<specimenCode id="DB926436FE46FFADFA45FE70FEDAFE32" box="[282,352,427,451]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="15" pageNumber="16">P1215</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE46FFADFA33FE70FF74FE12" collectionCode="NMNHU-P" country="Ukraine" pageId="15" pageNumber="16" specimenCode="PI 2564" specimenCount="9" typeStatus="paratype">
<specimenCount id="9D3207C4FE46FFADFA33FE70FE4DFE32" box="[364,503,427,451]" count="9" pageId="15" pageNumber="16" type="generic">9 specimens</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE46FFADF939FE77FD5CFE32" box="[614,742,428,451]" pageId="15" pageNumber="16">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE46FFADF9B0FE77FF74FE12" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="15" pageNumber="16">PI 2564</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE46FFADFB87FE10FD40FE12" box="[216,762,459,483]" collectionCode="NMNHU-P" country="Ukraine" pageId="15" pageNumber="16" specimenCode="PI 2565" specimenCount="9" typeStatus="paratype">
<specimenCount id="9D3207C4FE46FFADFB87FE10FEDAFE12" box="[216,352,459,483]" count="9" pageId="15" pageNumber="16" type="generic">9 specimens</specimenCount>
, Shydlivshchyna,
<collectionCode id="ED255488FE46FFADF940FE17FD25FE12" box="[543,671,460,483]" pageId="15" pageNumber="16">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE46FFADF9F9FE17FD40FE12" box="[678,762,459,483]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="15" pageNumber="16">PI 2565</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BCC4DFE46FFADFBC8FDD0FF4CFDB2" blockId="15.[151,775,523,579]" pageId="15" pageNumber="16">
<emphasis id="B940105FFE46FFADFBC8FDD0FEB2FDD2" box="[151,264,523,547]" italics="true" pageId="15" pageNumber="16">Etymology</emphasis>
Named after the region Podolia of western
<collectingCountry id="F3238CDDFE46FFADFBC8FDF0FF48FDB2" box="[151,242,555,579]" name="Ukraine" pageId="15" pageNumber="16">Ukraine</collectingCountry>
.
</paragraph>
<paragraph id="8B8BCC4DFE46FFADFBC8FDB7FE4BFC92" blockId="15.[151,775,618,867]" pageId="15" pageNumber="16">
<emphasis id="B940105FFE46FFADFBC8FDB7FEBBFD72" box="[151,257,620,643]" italics="true" pageId="15" pageNumber="16">Diagnosis</emphasis>
OL:OH = 1.21.25. Anterior rim inclined at 7580° with short preventral projection; posterior rim with moderately developed postdorsal projection. Dorsal rim relatively shallow. Sulcus wide, inclined at 1015°. Ostial lobe moderate; no subcaudal iugum. Ventral furrow deep, distant and divergent from ventral otolith rim and connected around sulcus to dorsal depression; area around sulcus distinctly bulged.
</paragraph>
<paragraph id="8B8BCC4DFE46FFADFBC8FC57FEE3FAF2" blockId="15.[151,776,907,1667]" pageId="15" pageNumber="16">
<emphasis id="B940105FFE46FFADFBC8FC57FEAEFC52" box="[151,276,908,931]" italics="true" pageId="15" pageNumber="16">Description</emphasis>
Moderately slender, rectangular, massive otoliths reaching about
<quantity id="4CCC61A8FE46FFADFAEEFC70FDB1FC32" box="[433,523,939,963]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="16" unit="mm" value="1.6">1.6 mm</quantity>
in length (
<typeStatus id="548F72EFFE46FFADF9C8FC70FD46FC32" box="[663,764,939,963]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
). OH:OT = 2.22.5. Anterior rim slightly inclined, with feeble concavity at about tip of ostium and distinct but relatively short preventral angle. Predorsal angle distinct, positioned high on anterior rim. Ventral rim nearly straight, smooth. Dorsal rim shallow, with variably developed mostly blunt postdorsal angle, slightly ascending toward posterior, with moderately strong postdorsal projection not bent outward. Posterior rim with distinct concavity below postdorsal projection and orthogonal postventral angle.
</paragraph>
<paragraph id="8B8BCC4DFE46FFADFBF2FAD7FEC8F972" blockId="15.[151,776,907,1667]" pageId="15" pageNumber="16">Inner face moderately convex, with area around sulcus and interior of ventral furrow and dorsal depression distinctly bulged. Sulcus wide, relatively deep, centrally positioned, moderately inclined at 1015°. OL:SuL = 2.0 2.5; SuL:SuH = 1.82.1. Sulcus with moderate ostial lobe, smoothened sole-shaped, with rounded anterior and posterior tips. No subcaudal iugum. Ventral furrow broad, deep, distinct, regularly curved, distant and divergent from ventral rim of otolith, connected around sulcus to moderately wide and deep dorsal depression. Outer face convex, more convex than inner face and very regularly shaped and smooth.</paragraph>
</subSubSection>
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Discussion
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differs from
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<emphasis id="B940105FFE46FFADF9B6F977FEBAF912" italics="true" pageId="15" pageNumber="16">M. renesulcis</emphasis>
</taxonomicName>
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described below in the less reduced and larger sulcus and the anteriorly not depressed predorsal angle (vs. depressed). Te dorsal depression is wider in
<taxonomicName id="4C34B7CEFE46FFADFBEAF8F7FEF8F8B2" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[181,322,1835,1859]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="podolicus">
<emphasis id="B940105FFE46FFADFBEAF8F7FEF8F8B2" box="[181,322,1835,1859]" italics="true" pageId="15" pageNumber="16">M. podolicus</emphasis>
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than in
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<emphasis id="B940105FFE46FFADFAFEF8F7FD8BF8B2" box="[417,561,1835,1859]" italics="true" pageId="15" pageNumber="16">M. renesulcis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE46FFADF966F8F7FDCEF8B2" box="[569,628,1836,1859]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C34B7CEFE46FFADF923F8F0FE8DF892" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="podolicus">
<emphasis id="B940105FFE46FFADF923F8F0FE8DF892" italics="true" pageId="15" pageNumber="16">Medoborichthys podolicus</emphasis>
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is as common as
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<emphasis id="B940105FFE46FFADF947F890FD5EF892" box="[536,740,1867,1891]" italics="true" pageId="15" pageNumber="16">Gobius ukrainicus</emphasis>
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in the assemblage, and both can be confused in atypical or eroded specimens. Consistent differences between the two include the wider sulcus (SuL:SuH = 1.82.1 vs. 2.42.8), the missing subcaudal iugum (vs. present), the shape of the ventral furrow, and the strongly bulged area around the sulcus in
<taxonomicName id="4C34B7CEFE46FFADFF50FEB7FB20FE72" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[1039,1178,363,387]" family="Gobiidae" genus="Medoborichthys" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="podolicus">
<emphasis id="B940105FFE46FFADFF50FEB7FB20FE72" box="[1039,1178,363,387]" italics="true" pageId="15" pageNumber="16">M. podolicus</emphasis>
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.
</paragraph>
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</document>

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@ -0,0 +1,88 @@
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Suborder
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<emphasis id="B940105FFE4EFFA5FBC8F976FF4FF934" box="[151,245,1709,1733]" italics="true" pageId="7" pageNumber="8">Remarks</emphasis>
Te familial arrangement of the families of the
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is still very much in flux. Here, we have kept the
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<subSubSection id="C32E9FC6FE4EFFA5FBC8FCB0FDC4FC72" box="[151,638,875,899]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B8BCC4DFE4EFFA5FBC8FCB0FDC4FC72" blockId="7.[151,638,875,899]" box="[151,638,875,899]" pageId="7" pageNumber="8">
<heading id="D0C37B21FE4EFFA5FBC8FCB0FDC4FC72" box="[151,638,875,899]" fontSize="9" level="2" pageId="7" pageNumber="8" reason="8">
<taxonomicName id="4C34B7CEFE4EFFA5FBC8FCB0FDC4FC72" authority="(Schubert, 1906)" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[151,638,875,899]" family="Lotidae" genus="Onogadus" kingdom="Animalia" order="Gadiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="simplicissimus">
<emphasis id="B940105FFE4EFFA5FBC8FCB0FD80FC72" bold="true" box="[151,570,875,899]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5FBC8FCB0FE01FC72" bold="true" box="[151,443,875,899]" italics="true" pageId="7" pageNumber="8">Onogadus simplicissimus</emphasis>
(
<bibRefCitation id="EFA5B1BCFE4EFFA5FA97FCB0FD80FC72" author="Schubert, R. J." box="[456,570,875,899]" pageId="7" pageNumber="8" pagination="623 - 706" refId="ref23059" refString="Schubert, R. J. (1906). Die Fischotolithen des osterr. - ungar. Tertiars. III. Macruriden und Beryciden. Jahrbuch Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, 56, 623 - 706." type="journal article" year="1906">Schubert,</bibRefCitation>
</emphasis>
1906
<emphasis id="B940105FFE4EFFA5F92AFCB0FDC4FC72" bold="true" box="[629,638,875,899]" pageId="7" pageNumber="8">)</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE4EFFA5FBC8FC70FEE2FBD2" pageId="7" pageNumber="8" type="description">
<paragraph id="8B8BCC4DFE4EFFA5FBC8FC70FEA8FC32" blockId="7.[151,274,939,963]" box="[151,274,939,963]" pageId="7" pageNumber="8">
<figureCitation id="130FD0C8FE4EFFA5FBC8FC70FEA8FC32" box="[151,274,939,963]" captionStart="Fig" captionStartId="6.[168,199,1875,1895]" captionTargetBox="[153,1438,1176,1934]" captionTargetId="figure-236@6.[168,1423,1192,1862]" captionTargetPageId="6" captionText="Fig. 3 Otoliths of Gadiformes: a, b Micromesistius planatus (Bassoli &amp; Schubert, 1906), Mlyntsi, a NMNHU-P PI 2568 and b NMB P1218. cf Onogadus simplicissimus (Schubert, 1906), c (reversed), d, f Shydlivshchyna, NMB P1220, e Mlyntsi, NMNHU-P PI 2572" figureDoi="http://doi.org/10.5281/zenodo.11999867" httpUri="https://zenodo.org/record/11999867/files/figure.png" pageId="7" pageNumber="8">Figure 3cf</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE4EFFA5FBC8FC30FEE2FBD2" blockId="7.[151,776,1003,1059]" pageId="7" pageNumber="8">
1906
<taxonomicName id="4C34B7CEFE4EFFA5FB87FC30FD3CFBF2" box="[216,646,1003,1027]" family="Sciaenidae" genus="Otolithus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="simplicissimus" subGenus="Crenilabrus">
<emphasis id="B940105FFE4EFFA5FB87FC30FE84FBF2" box="[216,318,1003,1027]" italics="true" pageId="7" pageNumber="8">Otolithus</emphasis>
(
<emphasis id="B940105FFE4EFFA5FA0CFC30FE6EFBF2" box="[339,468,1003,1027]" italics="true" pageId="7" pageNumber="8">Crenilabrus</emphasis>
)
<emphasis id="B940105FFE4EFFA5FAB5FC30FD3CFBF2" box="[490,646,1003,1027]" italics="true" pageId="7" pageNumber="8">simplicissimus</emphasis>
</taxonomicName>
—Schubert: pl. 18, figs. 43, 44.
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE4EFFA5FBC8FB90FD19FB72" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="8B8BCC4DFE4EFFA5FBC8FB90FD19FB72" blockId="7.[151,774,1099,1155]" pageId="7" pageNumber="8">
2015
<taxonomicName id="4C34B7CEFE4EFFA5FB87FB90FD0CFB92" authority="(Schubert, 1906)" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[216,694,1099,1123]" family="Lotidae" genus="Onogadus" kingdom="Animalia" order="Gadiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="simplicissimus">
<emphasis id="B940105FFE4EFFA5FB87FB90FE55FB92" box="[216,495,1099,1123]" italics="true" pageId="7" pageNumber="8">Onogadus simplicissimus</emphasis>
(
<bibRefCitation id="EFA5B1BCFE4EFFA5F95BFB90FD15FB92" author="Schubert, R. J." box="[516,687,1099,1123]" pageId="7" pageNumber="8" pagination="623 - 706" refId="ref23059" refString="Schubert, R. J. (1906). Die Fischotolithen des osterr. - ungar. Tertiars. III. Macruriden und Beryciden. Jahrbuch Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, 56, 623 - 706." type="journal article" year="1906">Schubert, 1906</bibRefCitation>
)
</taxonomicName>
—Bratishko et al.: figs. 4.612 (and references therein).
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE4EFFA5FBC8FB70FE5BFA12" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8B8BCC4DFE4EFFA5FBC8FB70FCBCFAF2" blockId="7.[151,775,1195,1283]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5FBC8FB70FF48FB32" box="[151,242,1195,1219]" italics="true" pageId="7" pageNumber="8">Material</emphasis>
<materialsCitation id="3B5CC610FE4EFFA5FBA4FB70FF5DFB12" collectionCode="NMNHU-P" pageId="7" pageNumber="8" specimenCode="PI 2571" specimenCount="1">
8 otoliths:
<specimenCount id="9D3207C4FE4EFFA5FA34FB70FE5FFB32" box="[363,485,1195,1219]" count="1" pageId="7" pageNumber="8" type="generic">1 specimen</specimenCount>
, Kozatskyi Yar,
<collectionCode id="ED255488FE4EFFA5F9D4FB77FCBDFB32" box="[651,775,1196,1219]" pageId="7" pageNumber="8">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE4EFFA5FBC8FB17FF5DFB12" box="[151,231,1227,1251]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI 2571</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE4EFFA5FBADFB10FD16FB12" box="[242,684,1227,1251]" collectionCode="NMNHU-P" pageId="7" pageNumber="8" specimenCode="PI 2572" specimenCount="2">
<specimenCount id="9D3207C4FE4EFFA5FBADFB10FECEFB12" box="[242,372,1227,1251]" count="2" pageId="7" pageNumber="8" type="generic">2 specimens</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE4EFFA5FA86FB17FDEFFB12" box="[473,597,1228,1251]" pageId="7" pageNumber="8">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE4EFFA5F904FB17FD16FB12" box="[603,684,1227,1251]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI 2572</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE4EFFA5F9E9FB10FDEAFAF2" collectionCode="NMNHU-P" pageId="7" pageNumber="8" specimenCode="PI 2573" specimenCount="5">
<specimenCount id="9D3207C4FE4EFFA5F9E9FB10FF6BFAF2" count="5" pageId="7" pageNumber="8" type="generic">5 specimens</specimenCount>
, Shydlivshchyna,
<collectionCode id="ED255488FE4EFFA5FADDFB37FE44FAF2" box="[386,510,1260,1283]" pageId="7" pageNumber="8">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE4EFFA5F95CFB37FDEAFAF2" box="[515,592,1259,1283]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI 2573</specimenCode>
</materialsCitation>
<materialsCitation id="3B5CC610FE4EFFA5F90AFB30FCB8FAF2" box="[597,770,1259,1283]" collectionCode="NMB" pageId="7" pageNumber="8" specimenCode="P1220" specimenCount="1">
and
<collectionCode id="ED255488FE4EFFA5F9DFFB37FD01FAF2" box="[640,699,1260,1283]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="7" pageNumber="8">NMB</collectionCode>
<specimenCode id="DB926436FE4EFFA5F9E0FB30FCB8FAF2" box="[703,770,1259,1283]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="7" pageNumber="8">P1220</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BCC4DFE4EFFA5FBC8FAF7FE5BFA12" blockId="7.[151,775,1323,1507]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5FBC8FAF7FEB0FAB2" box="[151,266,1324,1347]" italics="true" pageId="7" pageNumber="8">Discussion</emphasis>
Tis species was redefined, described in detail, and compared to extant species in Bratishko et. al. (2015).
<taxonomicName id="4C34B7CEFE4EFFA5FBB1FAB0FDB9FA72" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[238,515,1387,1411]" family="Lotidae" genus="Onogadus" kingdom="Animalia" order="Gadiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="simplicissimus">
<emphasis id="B940105FFE4EFFA5FBB1FAB0FDB9FA72" box="[238,515,1387,1411]" italics="true" pageId="7" pageNumber="8">Onogadus simplicissimus</emphasis>
</taxonomicName>
is apparently endemic to the Central and Eastern Paratethys, where it occurs from the late Badenian/Konkian to the Bessarabian (Bratishko et al., ongoing research).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="61613FF40BA8157CAC44534049382F80">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<paragraph id="8B8BCC4DFE4EFFA5FBC8FE30FE56FDF2" blockId="7.[151,492,491,515]" box="[151,492,491,515]" pageId="7" pageNumber="8">
<heading id="D0C37B21FE4EFFA5FBC8FE30FE56FDF2" bold="true" box="[151,492,491,515]" fontSize="10" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B940105FFE4EFFA5FBC8FE30FE56FDF2" bold="true" box="[151,492,491,515]" pageId="7" pageNumber="8">
Genus
<taxonomicName id="4C34B7CEFE4EFFA5FBB8FE30FE56FDF2" ID-CoL="6PF9" authority="Walbaum, 1792" authorityName="Walbaum" authorityYear="1792" box="[231,492,491,515]" family="Phycidae" genus="Phycis" kingdom="Animalia" order="Gadiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE4EFFA5FBB8FE30FE8AFDF2" bold="true" box="[231,304,491,515]" italics="true" pageId="7" pageNumber="8">Phycis</emphasis>
Walbaum, 1792
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE4EFFA5FBC8FDF0FEBDFDB2" box="[151,263,555,579]" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8B8BCC4DFE4EFFA5FBC8FDF0FEBDFDB2" blockId="7.[151,263,555,579]" box="[151,263,555,579]" pageId="7" pageNumber="8">
<heading id="D0C37B21FE4EFFA5FBC8FDF0FEBDFDB2" bold="true" box="[151,263,555,579]" fontSize="10" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="4C34B7CEFE4EFFA5FBC8FDF0FEBDFDB2" box="[151,263,555,579]" family="Phycidae" genus="Phycis" kingdom="Animalia" order="Gadiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE4EFFA5FBC8FDF0FEBDFDB2" bold="true" box="[151,263,555,579]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5FBC8FDF0FF5AFDB2" bold="true" box="[151,224,555,579]" italics="true" pageId="7" pageNumber="8">Phycis</emphasis>
sp.
</emphasis>
</taxonomicName>
</heading>
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</subSubSection>
<subSubSection id="C32E9FC6FE4EFFA5FBC8FDB0FE14FD32" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8B8BCC4DFE4EFFA5FBC8FDB0FE14FD32" blockId="7.[151,775,619,707]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5FBC8FDB0FF42FD72" box="[151,248,619,643]" italics="true" pageId="7" pageNumber="8">Material</emphasis>
A single, small, and poorly preserved specimen from Kozatskyi Yar,
<collectionCode id="ED255488FE4EFFA5FA26FD57FE43FD52" box="[377,505,652,675]" pageId="7" pageNumber="8">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE4EFFA5F95DFD57FDA0FD52" box="[514,538,652,675]" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI</collectionCode>
2575, that cannot be identified to species level.
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<mods:title id="8581A27D272248F065C9460F445AB877">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:namePart id="9F7D33C5C814A65FAA1768A5AF294C56">Schwarzhans, Werner</mods:namePart>
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<mods:namePart id="DF78386AFCB7C33D1AF8FA4B31CFA72E">Ryabokon, Tamara</mods:namePart>
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<mods:date id="97ED8AB42E8641BD5713C03EA7A43BF5">2022</mods:date>
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<heading id="D0C37B21FE4EFFA5F872FE30FB86FDF2" bold="true" box="[813,1084,491,515]" fontSize="10" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="B940105FFE4EFFA5F872FE30FC40FDF2" bold="true" box="[813,1018,491,515]" italics="true" pageId="7" pageNumber="8">Gobius bratishkoi</emphasis>
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<paragraph id="8B8BCC4DFE4EFFA5F872FDF0FC16FDB2" blockId="7.[813,940,555,579]" box="[813,940,555,579]" pageId="7" pageNumber="8">
<figureCitation id="130FD0C8FE4EFFA5F872FDF0FC16FDB2" box="[813,940,555,579]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: ae Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, be paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,ce (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).gk Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, gi, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="7" pageNumber="8">Figure 4ae</figureCitation>
</paragraph>
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<emphasis id="B940105FFE4EFFA5F872FDB0FC34FD72" box="[813,910,619,643]" italics="true" pageId="7" pageNumber="8">
<typeStatus id="548F72EFFE4EFFA5F872FDB0FC34FD72" box="[813,910,619,643]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
<collectionCode id="ED255488FE4EFFA5F8C3FDB7FBA6FD72" box="[924,1052,620,643]" pageId="7" pageNumber="8">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE4EFFA5FF75FDB7FB3CFD72" box="[1066,1158,619,643]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI 2555</specimenCode>
,
<location id="8EEB9A96FE4EFFA5FFC7FDB0FA8FFD72" LSID="urn:lsid:plazi:treatment:039D7D5BFE4EFFABF872FE30FE4EFA72:8EEB9A96FE4EFFA5FFC7FDB0FA8FFD72" box="[1176,1333,619,643]" country="Ukraine" municipality="Badenian" name="Kozatskyi Yar" pageId="7" pageNumber="8">Kozatskyi Yar</location>
, western
<collectingCountry id="F3238CDDFE4EFFA5F872FD50FC3DFD52" box="[813,903,651,675]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
, late
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,
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.
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</paragraph>
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<materialsCitation id="3B5CC610FE4EFFA5F872FD17FC0EFCF2" collectionCode="NMB" country="Ukraine" location="Kozatskyi Yar" municipality="Badenian" pageId="7" pageNumber="8" specimenCode="P1211" specimenCount="5" typeStatus="paratype">
<emphasis id="B940105FFE4EFFA5F872FD17FC22FD12" box="[813,920,716,739]" italics="true" pageId="7" pageNumber="8">
<typeStatus id="548F72EFFE4EFFA5F872FD17FC22FD12" box="[813,920,716,739]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
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7 otoliths:
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same data as holotype,
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<specimenCode id="DB926436FE4EFFA5F830FD30FC0EFCF2" box="[879,948,747,771]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="7" pageNumber="8">P1211</specimenCode>
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;
<materialsCitation id="3B5CC610FE4EFFA5F8E1FD30FA36FCF2" box="[958,1420,747,771]" collectionCode="NMNHU-P" country="Ukraine" pageId="7" pageNumber="8" specimenCode="PI 2556" specimenCount="2" typeStatus="paratype">
<specimenCount id="9D3207C4FE4EFFA5F8E1FD30FBFDFCF2" box="[958,1095,747,771]" count="2" pageId="7" pageNumber="8" type="generic">2 specimens</specimenCount>
, Mlyntsi,
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<specimenCode id="DB926436FE4EFFA5FE68FD37FA36FCF2" box="[1335,1420,747,771]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI 2556</specimenCode>
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.
</paragraph>
<paragraph id="8B8BCC4DFE4EFFA5F872FCF0FA27FC72" blockId="7.[813,1437,811,899]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5F872FCF0FC20FCB2" box="[813,922,811,835]" italics="true" pageId="7" pageNumber="8">Etymology</emphasis>
Named after Andriy Bratishko, Tallahassee, Florida (formerly
<collectingRegion id="49F002AFFE4EFFA5F8E1FC90FBA0FC92" box="[958,1050,843,867]" country="Ukraine" name="Luhans'k" pageId="7" pageNumber="8">Luhansk</collectingRegion>
,
<collectingCountry id="F3238CDDFE4EFFA5FF7AFC90FBC6FC92" box="[1061,1148,843,867]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
), in recognition of his contributions to the fossil fish otoliths of the Eastern Paratethys.
</paragraph>
<paragraph id="8B8BCC4DFE4EFFA5F872FC77FBA4FB52" blockId="7.[813,1437,938,1187]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5F872FC77FC2DFC32" box="[813,919,940,963]" italics="true" pageId="7" pageNumber="8">Diagnosis</emphasis>
OL:OH = 1.21.3. Anterior rim nearly vertical; posterior rim with prominent, rounded postdorsal projection. Dorsal rim with distinct, broad predorsal concavity and narrow postdorsal angle positioned above cauda. Sulcus narrow, relatively short (OL:SuL = 2.12.35), inclined at 1018°. Ostial lobe low; subcaudal iugum small, below anterior part of cauda. Inner face flat; outer face markedly convex.
</paragraph>
<paragraph id="8B8BCC4DFE4EFFA5F872FB17FB11F972" blockId="7.[813,1437,1227,1827]" pageId="7" pageNumber="8">
<emphasis id="B940105FFE4EFFA5F872FB17FC10FB12" box="[813,938,1228,1251]" italics="true" pageId="7" pageNumber="8">Description</emphasis>
Moderately slender, massive gobiid otoliths reaching about
<quantity id="4CCC61A8FE4EFFA5F8BFFB30FB82FAF2" box="[992,1080,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="mm" value="2.2">2.2 mm</quantity>
in length (
<typeStatus id="548F72EFFE4EFFA5FFE3FB30FAA7FAF2" box="[1212,1309,1259,1283]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
<quantity id="4CCC61A8FE4EFFA5FE76FB30FA2BFAF2" box="[1321,1425,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="8" unit="mm" value="2.15">2.15 mm</quantity>
). OH:OT = 2.22.6, decreasing with size. Anterior rim nearly vertical with only minor indentation above level of ostial tip. Preventral angle nearly orthogonal, predorsal angle distinct, positioned high on anterior rim. Ventral rim nearly straight to slightly bent. Dorsal rim with broad concavity across entire predorsal section followed by narrow postdorsal angle positioned above cauda and in turn followed by shorter concavity or flat, inclined section toward distinctly expanded postdorsal projection with rounded tip and not bent outward. Posterior rim below postdorsal projection nearly vertical, with orthogonal postventral angle. All rims smooth.
</paragraph>
<paragraph id="8B8BCC4DFE4EFFABF81DF957FCBDFEF2" blockId="7.[813,1437,1227,1827]" lastBlockId="9.[151,775,235,580]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">Inner face slightly convex, nearly flat, with narrow, somewhat deepened, centrally positioned, distinctly inclined sulcus. SuL:SuH = 2.52.9. Sulcus with very low ostial lobe and rounded anterior and posterior tips, inclined at 1018°. Subcaudal iugum small, often indistinct or indiscernible after even slight erosion</paragraph>
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<emphasis id="B940105FFE41FFAAFBF7F935FF61F8F3" bold="true" box="[168,219,1774,1794]" pageId="8" pageNumber="9">Fig. 4</emphasis>
Otoliths of
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<emphasis id="B940105FFE41FFAAFA1CF935FEC6F8F3" box="[323,380,1774,1794]" italics="true" pageId="8" pageNumber="9">Gobius</emphasis>
spp.
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:
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<taxonomicName id="4C34B7CEFE41FFAAFA8EF935FDE7F8F3" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[465,605,1774,1794]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bratishkoi" status="sp. nov.">
<emphasis id="B940105FFE41FFAAFA8EF935FDE7F8F3" box="[465,605,1774,1794]" italics="true" pageId="8" pageNumber="9">Gobius bratishkoi</emphasis>
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<taxonomicNameLabel id="A273AD24FE41FFAAF93DF935FD2AF8F3" box="[610,656,1774,1794]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
,
<emphasis id="B940105FFE41FFAAF9CAF935FD1AF8F3" bold="true" box="[661,672,1774,1794]" pageId="8" pageNumber="9">a</emphasis>
holotype, NMNHU-P PI 2555, Kozatskyi Yar,
<emphasis id="B940105FFE41FFAAFF50F935FB8BF8F3" bold="true" box="[1039,1073,1774,1794]" pageId="8" pageNumber="9">be</emphasis>
paratypes,
<emphasis id="B940105FFE41FFAAFFCDF935FB24F8F3" bold="true" box="[1170,1182,1774,1794]" pageId="8" pageNumber="9">b</emphasis>
(reversed) Mlyntsi, NMNHU-P PI 2556,
<emphasis id="B940105FFE41FFAAFA15F8D2FED0F8EC" bold="true" box="[330,362,1801,1821]" pageId="8" pageNumber="9">ce</emphasis>
(reversed) Kozatskyi Yar, NMB P1211.
<emphasis id="B940105FFE41FFAAF9FBF8D2FD11F8EC" bold="true" box="[676,683,1801,1821]" pageId="8" pageNumber="9">f</emphasis>
<taxonomicName id="4C34B7CEFE41FFAAF9F0F8D2FBA3F8EC" authority="Schwarzhans, 2014" authorityName="Schwarzhans" authorityYear="2014" box="[687,1049,1801,1821]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="reichenbacherae">
<emphasis id="B940105FFE41FFAAF9F0F8D2FCC9F8EC" box="[687,883,1801,1821]" italics="true" pageId="8" pageNumber="9">Gobius reichenbacherae</emphasis>
<bibRefCitation id="EFA5B1BCFE41FFAAF827F8D2FBA3F8EC" author="Schwarzhans, W." box="[888,1049,1801,1821]" pageId="8" pageNumber="9" pagination="35 - 69" refId="ref23301" refString="Schwarzhans, W. (2014). Otoliths from the middle Miocene (Serravallian) of the Karaman Basin, Turkey. Cainozoic Research, 14 (1), 35 - 69." type="journal article" year="2014">Schwarzhans,2014</bibRefCitation>
</taxonomicName>
, Shydlivshchyna, NMNHU-P PI 2557 (reversed).
<emphasis id="B940105FFE41FFAAFA5BF8F8FE9CF8C6" bold="true" box="[260,294,1827,1847]" pageId="8" pageNumber="9">gk</emphasis>
<taxonomicName id="4C34B7CEFE41FFAAFA74F8F8FE00F8C6" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[299,442,1827,1847]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ukrainicus" status="sp. nov.">
<emphasis id="B940105FFE41FFAAFA74F8F8FE00F8C6" box="[299,442,1827,1847]" italics="true" pageId="8" pageNumber="9">Gobius ukrainicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE41FFAAFAE0F8F8FE57F8C6" box="[447,493,1827,1847]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
,
<emphasis id="B940105FFE41FFAAFAADF8F8FE42F8C6" bold="true" box="[498,504,1827,1847]" pageId="8" pageNumber="9">j</emphasis>
holotype, Shydlivshchyna, NMNHU-P PI 2558,
<emphasis id="B940105FFE41FFAAF8DFF8F8FC26F8C6" bold="true" box="[896,924,1827,1847]" pageId="8" pageNumber="9">gi</emphasis>
,
<emphasis id="B940105FFE41FFAAF8FAF8F8FC0AF8C6" bold="true" box="[933,944,1827,1847]" pageId="8" pageNumber="9">k</emphasis>
paratypes,
<emphasis id="B940105FFE41FFAAFF4FF8F8FBA6F8C6" bold="true" box="[1040,1052,1827,1847]" pageId="8" pageNumber="9">g</emphasis>
,
<emphasis id="B940105FFE41FFAAFF7AF8F8FB90F8C6" bold="true" box="[1061,1066,1827,1847]" pageId="8" pageNumber="9">i</emphasis>
,
<emphasis id="B940105FFE41FFAAFF6DF8F8FB87F8C6" bold="true" box="[1074,1085,1827,1847]" pageId="8" pageNumber="9">k</emphasis>
NMNHU-P PI 2560,
<emphasis id="B940105FFE41FFAAFFBBF8F8FB4AF8C6" bold="true" box="[1252,1264,1827,1847]" pageId="8" pageNumber="9">h</emphasis>
(reversed) Mlyntsi, NMNHU-P PI 2559
</paragraph>
</caption>
<paragraph id="8B8BCC4DFE40FFABFBC8FED0FD2FFDB2" blockId="9.[151,775,235,580]" pageId="9" pageNumber="10">
(
<figureCitation id="130FD0C8FE40FFABFBC0FED0FEB0FED2" box="[159,266,267,291]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: ae Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, be paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,ce (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).gk Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, gi, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="9" pageNumber="10">Fig. 4c, d</figureCitation>
), positioned below sulcus indentation marking ostialcaudal joint. Ventral furrow broad, deep, distinct, reaching from anterior to posterior tips of sulcus, more regularly curved than ventral rim of otolith. Ventral field between ventral furrow and sulcus slightly bulged. Dorsal field with long, narrow dorsal depression, ventrally marked by distinct crista superior toward sulcus and dorsally open to dorsal rim of otolith. Outer face convex, distinctly more convex than inner face and very regularly shaped, smooth.
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE40FFABFBC8FDB7FE4EFA72" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8B8BCC4DFE40FFABFBC8FDB7FE4EFA72" blockId="9.[151,776,619,1412]" pageId="9" pageNumber="10">
<emphasis id="B940105FFE40FFABFBC8FDB7FE62FD72" box="[151,472,619,643]" italics="true" pageId="9" pageNumber="10">
Discussion
<taxonomicName id="4C34B7CEFE40FFABFA4AFDB0FE62FD72" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[277,472,619,643]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bratishkoi">Gobius bratishkoi</taxonomicName>
</emphasis>
is easily recognized by its characteristic dorsal rim with the two concavities, the nearly vertical anterior rim, and the distinct postdorsal projection with rounded tip that is not bent outward. Further distinguishing characteristics include the outer face being more convex than the inner face and the very low ostial lobe of the sulcus.
<taxonomicName id="4C34B7CEFE40FFABFA8CFCF0FD29FCB2" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[467,659,811,835]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bratishkoi">
<emphasis id="B940105FFE40FFABFA8CFCF0FD29FCB2" box="[467,659,811,835]" italics="true" pageId="9" pageNumber="10">Gobius bratishkoi</emphasis>
</taxonomicName>
resembles the coeval
<taxonomicName id="4C34B7CEFE40FFABFA4FFC97FDE0FC92" authority="Schwarzhans, 2014" authorityName="Schwarzhans" authorityYear="2014" box="[272,602,843,867]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
<emphasis id="B940105FFE40FFABFA4FFC97FEC5FC92" box="[272,383,844,867]" italics="true" pageId="9" pageNumber="10">G. mustus</emphasis>
<bibRefCitation id="EFA5B1BCFE40FFABFAD7FC90FDE0FC92" author="Schwarzhans, W." box="[392,602,843,867]" pageId="9" pageNumber="10" pagination="35 - 69" refId="ref23301" refString="Schwarzhans, W. (2014). Otoliths from the middle Miocene (Serravallian) of the Karaman Basin, Turkey. Cainozoic Research, 14 (1), 35 - 69." type="journal article" year="2014">Schwarzhans, 2014</bibRefCitation>
</taxonomicName>
but clearly differs in shape of the dorsal rim, the near vertical anterior rim, the postdorsal projection, the strongly reduced ostial lobe, and the very small subcaudal iugum. However,
<taxonomicName id="4C34B7CEFE40FFABF9B1FC77FEB9FC12" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bratishkoi">
<emphasis id="B940105FFE40FFABF9B1FC77FEB9FC12" italics="true" pageId="9" pageNumber="10">G. bratishkoi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C34B7CEFE40FFABFA4DFC17FE3AFC12" authorityName="Schwarzhans" authorityYear="2014" box="[274,384,972,995]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
<emphasis id="B940105FFE40FFABFA4DFC17FE3AFC12" box="[274,384,972,995]" italics="true" pageId="9" pageNumber="10">G. mustus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C34B7CEFE40FFABFAE0FC17FDE8FC12" authority="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk, 2022" authorityName="Schwarzhans &amp; Klots &amp; Ryabokon &amp; Kovalchuk" authorityYear="2022" box="[447,594,971,995]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ukrainicus" status="n. sp.">
<emphasis id="B940105FFE40FFABFAE0FC17FDE8FC12" box="[447,594,971,995]" italics="true" pageId="9" pageNumber="10">G. ukrainicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273AD24FE40FFABF905FC17FD2FFC12" box="[602,661,972,995]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
described below represent a distinct species group, which cannot be linked to any extant
<taxonomicName id="4C34B7CEFE40FFABFA99FBD0FDD2FBD2" box="[454,616,1035,1059]" pageId="9" pageNumber="10">
<emphasis id="B940105FFE40FFABFA99FBD0FDA9FBD2" box="[454,531,1035,1059]" italics="true" pageId="9" pageNumber="10">Gobius</emphasis>
species
</taxonomicName>
and which we term the
<taxonomicName id="4C34B7CEFE40FFABFBA0FBF0FE1BFBB2" authorityName="Schwarzhans" authorityYear="2014" box="[255,417,1067,1091]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
<emphasis id="B940105FFE40FFABFBA0FBF0FE1BFBB2" box="[255,417,1067,1091]" italics="true" pageId="9" pageNumber="10">Gobius mustus</emphasis>
</taxonomicName>
complex. Among extant
<taxonomicName id="4C34B7CEFE40FFABF9E5FBF0FCBDFBB2" authorityName="Linnaeus" authorityYear="1758" box="[698,775,1067,1091]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE40FFABF9E5FBF0FCBDFBB2" box="[698,775,1067,1091]" italics="true" pageId="9" pageNumber="10">Gobius</emphasis>
</taxonomicName>
otoliths,
<taxonomicName id="4C34B7CEFE40FFABFBA4FB97FE46FB92" authority="Risso, 1810" authorityName="Risso" authorityYear="1810" box="[251,508,1099,1123]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="auratus">
<emphasis id="B940105FFE40FFABFBA4FB97FEC9FB92" box="[251,371,1100,1123]" italics="true" pageId="9" pageNumber="10">G. auratus</emphasis>
Risso, 1810
</taxonomicName>
,
<taxonomicName id="4C34B7CEFE40FFABF954FB97FCB8FB92" authority="Sarato, 1889" authorityName="Sarato" authorityYear="1889" box="[523,770,1099,1123]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="fallax">
<emphasis id="B940105FFE40FFABF954FB97FDD7FB92" box="[523,621,1099,1123]" italics="true" pageId="9" pageNumber="10">G. fallax</emphasis>
Sarato, 1889
</taxonomicName>
, and
<taxonomicName id="4C34B7CEFE40FFABFB95FBB7FDAAFB72" authority="Vinciguerra, 1883" authorityName="Vinciguerra" authorityYear="1883" box="[202,528,1131,1155]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="B940105FFE40FFABFB95FBB7FEFBFB72" box="[202,321,1132,1155]" italics="true" pageId="9" pageNumber="10">G. vittatus</emphasis>
Vinciguerra, 1883
</taxonomicName>
(see
<bibRefCitation id="EFA5B1BCFE40FFABF90FFBB0FCBAFB72" author="Gut, C. &amp; Sanda, R. &amp; Moritz, T. &amp; Reichenbacher, B." box="[592,768,1131,1155]" pageId="9" pageNumber="10" pagination="282 - 323" refId="ref20633" refString="Gut, C., VukiC, J., Sanda, R., Moritz, T., &amp; Reichenbacher, B. (2020). Identification of past and present gobies: Distinguishing Gobius and Pomatoschistus (Teleostei: Gobioidei) species using characters of otoliths, meristics and body morphometry. Contributions to Zoology, 89, 282 - 323." type="journal article" year="2020">Gut et al., 2020</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE40FFABFBC8FB50FE00FB52" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[151,442,1163,1187]" pageId="9" pageNumber="10" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
for figures) share the closest resemblance in terms of their compressed shapes, their inner faces being nearly flat, and their outer faces more convex. However, the degree of thickness of the outer face and the small size of the sulcus distinguish the
<taxonomicName id="4C34B7CEFE40FFABF98EFAD0FEB4FAB2" authorityName="Schwarzhans" authorityYear="2014" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
<emphasis id="B940105FFE40FFABF98EFAD0FEB4FAB2" italics="true" pageId="9" pageNumber="10">Gobius mustus</emphasis>
</taxonomicName>
complex from those extant species and indicate that it may represent an extinct clade that is possibly separate from the genus
<taxonomicName id="4C34B7CEFE40FFABFAFDFAB0FE55FA72" authorityName="Linnaeus" authorityYear="1758" box="[418,495,1387,1411]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE40FFABFAFDFAB0FE55FA72" box="[418,495,1387,1411]" italics="true" pageId="9" pageNumber="10">Gobius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B940105FFE4FFFA4F872FEF0FBF3FEB2" bold="true" box="[813,1097,299,323]" italics="true" pageId="6" pageNumber="7">Micromesistius planatus</emphasis>
(
<bibRefCitation id="EFA5B1BCFE4FFFA4FF08FEF0FA87FEB2" author="Bassoli, G." box="[1111,1341,299,323]" pageId="6" pageNumber="7" pagination="36 - 61" refId="ref19471" refString="Bassoli, G. (1906). Otoliti fossili terziari dell' Emilia. Rivista Italiana di Paleontologia, 12, 36 - 61." type="journal article" year="1906">Bassoli &amp; Schubert,</bibRefCitation>
</emphasis>
1906
<emphasis id="B940105FFE4FFFA4FE27FEF0FA3BFEB2" bold="true" box="[1400,1409,299,323]" pageId="6" pageNumber="7">)</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE4FFFA4F872FEB0FC35FDB2" pageId="6" pageNumber="7" type="description">
<paragraph id="8B8BCC4DFE4FFFA4F872FEB0FC11FE72" blockId="6.[813,939,363,387]" box="[813,939,363,387]" pageId="6" pageNumber="7">
<figureCitation id="130FD0C8FE4FFFA4F872FEB0FC11FE72" box="[813,939,363,387]" captionStart="Fig" captionStartId="6.[168,199,1875,1895]" captionTargetBox="[153,1438,1176,1934]" captionTargetId="figure-236@6.[168,1423,1192,1862]" captionTargetPageId="6" captionText="Fig. 3 Otoliths of Gadiformes: a, b Micromesistius planatus (Bassoli &amp; Schubert, 1906), Mlyntsi, a NMNHU-P PI 2568 and b NMB P1218. cf Onogadus simplicissimus (Schubert, 1906), c (reversed), d, f Shydlivshchyna, NMB P1220, e Mlyntsi, NMNHU-P PI 2572" figureDoi="http://doi.org/10.5281/zenodo.11999867" httpUri="https://zenodo.org/record/11999867/files/figure.png" pageId="6" pageNumber="7">Figure 3a, b</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE4FFFA4F872FE70FB07FE12" blockId="6.[813,1436,427,483]" pageId="6" pageNumber="7">
1906
<taxonomicName id="4C34B7CEFE4FFFA4F834FE70FAA0FE32" box="[875,1306,427,451]" family="Sciaenidae" genus="Otolithus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="variety" species="elegans" subGenus="Phycis" variety="planata">
<emphasis id="B940105FFE4FFFA4F834FE70FC6BFE32" box="[875,977,427,451]" italics="true" pageId="6" pageNumber="7">Otolithus</emphasis>
(
<emphasis id="B940105FFE4FFFA4F8BBFE70FB92FE32" box="[996,1064,427,451]" italics="true" pageId="6" pageNumber="7">Phycis</emphasis>
)
<emphasis id="B940105FFE4FFFA4FF64FE70FB33FE32" box="[1083,1161,427,451]" italics="true" pageId="6" pageNumber="7">elegans</emphasis>
var.
<emphasis id="B940105FFE4FFFA4FF9BFE70FAA0FE32" box="[1220,1306,427,451]" italics="true" pageId="6" pageNumber="7">planata</emphasis>
</taxonomicName>
—Bassoli &amp; Schubert in Bassoli: pl. 1, figs. 11, 12.
</paragraph>
<paragraph id="8B8BCC4DFE4FFFA4F872FDD0FC35FDB2" blockId="6.[813,1437,523,579]" pageId="6" pageNumber="7">
1975
<taxonomicName id="4C34B7CEFE4FFFA4F820FDD7FB69FDD2" authorityName="Holec" authorityYear="1975" box="[895,1235,523,547]" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="schwarzhansi">
<emphasis id="B940105FFE4FFFA4F820FDD7FB69FDD2" box="[895,1235,523,547]" italics="true" pageId="6" pageNumber="7">Micromesistius schwarzhansi</emphasis>
</taxonomicName>
—Holec: pl. 2, figs. 24.
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE4FFFA4F872FDB0FBA4FC92" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8B8BCC4DFE4FFFA4F872FDB0FADEFD52" blockId="6.[813,1436,619,675]" pageId="6" pageNumber="7">
1983
<taxonomicName id="4C34B7CEFE4FFFA4F830FDB7FA8BFD72" authority="(Bassoli, 1906)" baseAuthorityName="Bassoli &amp; Schubert" baseAuthorityYear="1906" box="[879,1329,619,643]" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="planatus">
<emphasis id="B940105FFE4FFFA4F830FDB7FB3AFD72" box="[879,1152,619,643]" italics="true" pageId="6" pageNumber="7">Micromesistius planatus</emphasis>
(
<bibRefCitation id="EFA5B1BCFE4FFFA4FFCAFDB0FA90FD72" author="Bassoli, G." box="[1173,1322,619,643]" pageId="6" pageNumber="7" pagination="36 - 61" refId="ref19471" refString="Bassoli, G. (1906). Otoliti fossili terziari dell' Emilia. Rivista Italiana di Paleontologia, 12, 36 - 61." type="journal article" year="1906">Bassoli, 1906</bibRefCitation>
)
</taxonomicName>
—Nolf &amp; Steurbaut: pl. 4, figs. 1013 (and references therein).
</paragraph>
<paragraph id="8B8BCC4DFE4FFFA4F872FD10FB69FCF2" blockId="6.[813,1436,715,771]" pageId="6" pageNumber="7">
1992
<taxonomicName id="4C34B7CEFE4FFFA4F833FD17FBC0FD12" authorityName="Radwanska" authorityYear="1992" box="[876,1146,716,739]" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="arcuatus">
<emphasis id="B940105FFE4FFFA4F833FD17FBC0FD12" box="[876,1146,716,739]" italics="true" pageId="6" pageNumber="7">Micromesistius arcuatus</emphasis>
</taxonomicName>
—Radwańska: text-fig. 49, pl. 9, figs. 59 (and references therein).
</paragraph>
<paragraph id="8B8BCC4DFE4FFFA4F872FCF0FBA4FC92" blockId="6.[813,1436,811,867]" pageId="6" pageNumber="7">
2010
<taxonomicName id="4C34B7CEFE4FFFA4F832FCF7FA90FCB2" authority="(Bassoli, 1906)" baseAuthorityName="Bassoli &amp; Schubert" baseAuthorityYear="1906" box="[877,1322,811,835]" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="planatus">
<emphasis id="B940105FFE4FFFA4F832FCF7FBC6FCB2" box="[877,1148,811,835]" italics="true" pageId="6" pageNumber="7">Micromesistius planatus</emphasis>
(
<bibRefCitation id="EFA5B1BCFE4FFFA4FFCFFCF0FA99FCB2" author="Bassoli, G." box="[1168,1315,811,835]" pageId="6" pageNumber="7" pagination="36 - 61" refId="ref19471" refString="Bassoli, G. (1906). Otoliti fossili terziari dell' Emilia. Rivista Italiana di Paleontologia, 12, 36 - 61." type="journal article" year="1906">Bassoli, 1906</bibRefCitation>
)
</taxonomicName>
—Schwarzhans: pl. 47, figs. 3, 4.
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE4FFFA5F872FC50FCBDFE32" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8B8BCC4DFE4FFFA4F872FC50FB7CFC32" blockId="6.[813,1436,907,963]" pageId="6" pageNumber="7">
<emphasis id="B940105FFE4FFFA4F872FC50FC34FC52" box="[813,910,907,931]" italics="true" pageId="6" pageNumber="7">Material</emphasis>
2 otoliths, Mlyntsi, OL 7.0 and
<quantity id="4CCC61A8FE4FFFA4FE67FC50FA22FC52" box="[1336,1432,907,931]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="7" unit="mm" value="4.2">4.2 mm</quantity>
,
<collectionCode id="ED255488FE4FFFA4F872FC77FC17FC32" box="[813,941,940,963]" pageId="6" pageNumber="7">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE4FFFA4F8ECFC77FC71FC32" box="[947,971,940,963]" country="Russia" name="Paleontological Institute" pageId="6" pageNumber="7">PI</collectionCode>
2568 and
<collectionCode id="ED255488FE4FFFA4FF65FC77FBCCFC32" box="[1082,1142,940,963]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="6" pageNumber="7">NMB</collectionCode>
P1218.
</paragraph>
<paragraph id="8B8BCC4DFE4FFFA5F872FC37FCBDFE32" blockId="6.[813,1437,1003,1091]" lastBlockId="7.[151,775,235,451]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B940105FFE4FFFA4F872FC37FC25FBF2" box="[813,927,1004,1027]" italics="true" pageId="6" pageNumber="7">Discussion</emphasis>
Otoliths of
<taxonomicName id="4C34B7CEFE4FFFA4FF76FC37FA89FBF2" baseAuthorityName="Bassoli &amp; Schubert" baseAuthorityYear="1906" box="[1065,1331,1003,1027]" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="planatus">
<emphasis id="B940105FFE4FFFA4FF76FC37FA89FBF2" box="[1065,1331,1003,1027]" italics="true" pageId="6" pageNumber="7">Micromesistius planatus</emphasis>
</taxonomicName>
exhibit a very pronounced ontogenetic allometry, whereby specimens of up to about
<quantity id="4CCC61A8FE4FFFA4FF55FBF0FBF2FBB2" box="[1034,1096,1067,1091]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="mm" value="8.0">8 mm</quantity>
in length are relatively thin and flat and those of the largest known specimens, which are over
<quantity id="4CCC61A8FE4EFFA5FB93FED0FEA2FED2" box="[204,280,267,291]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="mm" value="15.0">15 mm</quantity>
in length, are considerably thicker and show a more strongly bent inner face. Transitional morphologies between the size classes are relatively rare. Hence, smaller specimens have been described in the past as
<taxonomicName id="4C34B7CEFE4EFFA5F9D1FEB7FF66FE52" authorityName="Holec" authorityYear="1975" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="schwarzhansi">
<emphasis id="B940105FFE4EFFA5F9D1FEB7FF66FE52" italics="true" pageId="7" pageNumber="8">M. schwarzhansi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C34B7CEFE4EFFA5FBB5FE57FE4BFE52" baseAuthorityName="Bassoli &amp; Schubert" baseAuthorityYear="1906" box="[234,497,395,419]" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="planatus">
<emphasis id="B940105FFE4EFFA5FBB5FE57FE4BFE52" box="[234,497,395,419]" italics="true" pageId="7" pageNumber="8">Micromesistius planatus</emphasis>
</taxonomicName>
is known from the Badenian of the Paratethys and the Tortonian of northern
<collectingCountry id="F3238CDDFE4EFFA5F98DFE70FCB9FE32" box="[722,771,427,451]" name="Italy" pageId="7" pageNumber="8">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,119 @@
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<subSubSection id="C32E9FC6FE58FFB3F872FA62FA92FA20" box="[813,1320,1465,1489]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8B8BCC4DFE58FFB3F872FA62FA92FA20" blockId="17.[813,1320,1465,1489]" box="[813,1320,1465,1489]" pageId="17" pageNumber="18">
<heading id="D0C37B21FE58FFB3F872FA62FA92FA20" box="[813,1320,1465,1489]" fontSize="9" level="2" pageId="17" pageNumber="18" reason="8">
<taxonomicName id="4C34B7CEFE58FFB3F872FA62FA92FA20" authority="(Weiler, 1943)" baseAuthorityName="Weiler" baseAuthorityYear="1943" box="[813,1320,1465,1489]" family="Gobiidae" genus="Economidichthys" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="triangularis">
<emphasis id="B940105FFE58FFB3F872FA62FB5EFA20" bold="true" box="[813,1252,1465,1489]" pageId="17" pageNumber="18">
<emphasis id="B940105FFE58FFB3F872FA62FB38FA20" bold="true" box="[813,1154,1465,1489]" italics="true" pageId="17" pageNumber="18">Economidichthys triangularis</emphasis>
(
<bibRefCitation id="EFA5B1BCFE58FFB3FFD0FA62FB5EFA20" author="Weiler, W." box="[1167,1252,1465,1489]" pageId="17" pageNumber="18" pagination="87 - 115" refId="ref24030" refString="Weiler, W. (1943). Die Otolithen aus dem Jungtertiar SUd- Rumaniens, I. Buglow Und Sarmat. Senckenbergiana Lethaea, 26, 87 - 115." type="journal article" year="1943">Weiler,</bibRefCitation>
</emphasis>
1943
<emphasis id="B940105FFE58FFB3FE40FA62FA92FA20" bold="true" box="[1311,1320,1465,1489]" pageId="17" pageNumber="18">)</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE58FFB0F872FA22FEAAFED2" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8B8BCC4DFE58FFB3F872FA22FC13F9E0" blockId="17.[813,937,1529,1553]" box="[813,937,1529,1553]" pageId="17" pageNumber="18">
<figureCitation id="130FD0C8FE58FFB3F872FA22FC13F9E0" box="[813,937,1529,1553]" captionStart="Fig" captionStartId="17.[168,199,1199,1219]" captionTargetBox="[152,1437,237,1310]" captionTargetId="figure-220@17.[167,1422,253,1186]" captionTargetPageId="17" captionText="Fig. 7 Otoliths of Gobionellidae: ae Deltentosteus aff. telleri (Schubert, 1906), 7a Shydlivshchyna, NMNHU-P PI 2553, b (reversed),d Kozatskyi Yar, NMNHU-P PI 2552,c (reversed), e Mlyntsi, NMB P1209.f, g Economidichthys triangularis (Weiler, 1943), f (reversed) Shydlivshchyna, NMB P1210, g (reversed) Mlyntsi, NMNHU-P PI 2554.h, i Knipowitschia polonica Schwarzhans et al., 2020a, 2020b, h Shydlivshchyna, NMNHU-P PI 2562, i Kozatskyi Yar,NMB P1213. j, k Pomatoschistus elegans (Procházka, 1900), j (reversed) Kozatskyi Yar, NMNHU-P PI 2576, k (reversed) Shydlivshchyna, NMB P1222" figureDoi="http://doi.org/10.5281/zenodo.11999917" httpUri="https://zenodo.org/record/11999917/files/figure.png" pageId="17" pageNumber="18">Figure 7fg</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE58FFB3F872F9E2FB8DF960" blockId="17.[813,1436,1593,1681]" pageId="17" pageNumber="18">
2020a
<taxonomicName id="4C34B7CEFE58FFB3F8DFF9E2FA38F9A0" authority="(Weiler, 1943)" baseAuthorityName="Weiler" baseAuthorityYear="1943" box="[896,1410,1593,1617]" family="Gobiidae" genus="Economidichthys" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="triangularis">
<emphasis id="B940105FFE58FFB3F8DFF9E2FB76F9A0" box="[896,1228,1593,1617]" italics="true" pageId="17" pageNumber="18">Economidichthys triangularis</emphasis>
(
<bibRefCitation id="EFA5B1BCFE58FFB3FFBAF9E2FAC1F9A0" author="Weiler, W." box="[1253,1403,1593,1617]" pageId="17" pageNumber="18" pagination="87 - 115" refId="ref24030" refString="Weiler, W. (1943). Die Otolithen aus dem Jungtertiar SUd- Rumaniens, I. Buglow Und Sarmat. Senckenbergiana Lethaea, 26, 87 - 115." type="journal article" year="1943">Weiler, 1943</bibRefCitation>
)
</taxonomicName>
— Schwarzhans, Brzobohatý &amp; Radwańska: pl. 9, figs. 69 (and references therein).
</paragraph>
<paragraph id="8B8BCC4DFE58FFB3F872F962FAFBF900" blockId="17.[813,1437,1721,1777]" pageId="17" pageNumber="18">
<emphasis id="B940105FFE58FFB3F872F962FC34F920" box="[813,910,1721,1745]" italics="true" pageId="17" pageNumber="18">Material</emphasis>
<materialsCitation id="3B5CC610FE58FFB3F8C9F962FCDEF900" collectionCode="NMNHU-P" pageId="17" pageNumber="18" specimenCode="PI 2554" specimenCount="1">
3 otoliths:
<specimenCount id="9D3207C4FE58FFB3FF52F962FB37F920" box="[1037,1165,1721,1745]" count="1" pageId="17" pageNumber="18" type="generic">1 specimen</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE58FFB3FFA3F961FAC6F920" box="[1276,1404,1722,1745]" pageId="17" pageNumber="18">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE58FFB3FEDAF961FCDEF900" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="17" pageNumber="18">PI 2554</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE58FFB3F832F902FA87F900" box="[877,1341,1753,1777]" collectionCode="NMB" pageId="17" pageNumber="18" specimenCode="P1210" specimenCount="2">
<specimenCount id="9D3207C4FE58FFB3F832F902FC4FF900" box="[877,1013,1753,1777]" count="2" pageId="17" pageNumber="18" type="generic">2 specimens</specimenCount>
, Shydlivshchyna,
<collectionCode id="ED255488FE58FFB3FFEAF901FB4BF900" box="[1205,1265,1754,1777]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="17" pageNumber="18">NMB</collectionCode>
<specimenCode id="DB926436FE58FFB3FFA8F902FA87F900" box="[1271,1341,1753,1777]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="17" pageNumber="18">P1210</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BCC4DFE58FFB3F872F8C1FA27F8A0" blockId="17.[813,1437,1817,1873]" pageId="17" pageNumber="18">
<emphasis id="B940105FFE58FFB3F872F8C1FC1AF8C0" box="[813,928,1818,1841]" italics="true" pageId="17" pageNumber="18">Discussion</emphasis>
Tis small species is easily recognized by its triangular shape. It has commonly been recorded from
</paragraph>
<paragraph id="8B8BCC4DFE5BFFB0FBC8FF30FEAAFED2" blockId="18.[151,775,235,291]" pageId="18" pageNumber="19">the late Badenian and early Sarmatian s.s. of the Central Paratethys.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,251 @@
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<treatment id="039D7D5BFE59FFB3F872FDD0FE2BF8C0" LSID="urn:lsid:plazi:treatment:039D7D5BFE59FFB3F872FDD0FE2BF8C0" httpUri="http://treatment.plazi.org/id/039D7D5BFE59FFB3F872FDD0FE2BF8C0" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C32E9FC6FE59FFB2F872FDD0FAB8FDD2" box="[813,1282,523,547]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B8BCC4DFE59FFB2F872FDD0FAB8FDD2" blockId="16.[813,1282,523,547]" box="[813,1282,523,547]" pageId="16" pageNumber="17">
<heading id="D0C37B21FE59FFB2F872FDD0FAB8FDD2" box="[813,1282,523,547]" fontSize="9" level="2" pageId="16" pageNumber="17" reason="8">
<taxonomicName id="4C34B7CEFE59FFB2F872FDD0FAB8FDD2" authority="(Schubert, 1906)" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[813,1282,523,547]" family="Gobiidae" genus="Deltentosteus" isUncertain="true" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="telleri">
<emphasis id="B940105FFE59FFB2F872FDD0FB04FDD2" bold="true" box="[813,1214,523,547]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2F872FDD0FC70FDD2" bold="true" box="[813,970,523,547]" italics="true" pageId="16" pageNumber="17">Deltentosteus</emphasis>
aff.
<emphasis id="B940105FFE59FFB2F8A6FDD0FB84FDD2" bold="true" box="[1017,1086,523,547]" italics="true" pageId="16" pageNumber="17">telleri</emphasis>
(
<bibRefCitation id="EFA5B1BCFE59FFB2FF13FDD0FB04FDD2" author="Schubert, R. J." box="[1100,1214,523,547]" pageId="16" pageNumber="17" pagination="623 - 706" refId="ref23059" refString="Schubert, R. J. (1906). Die Fischotolithen des osterr. - ungar. Tertiars. III. Macruriden und Beryciden. Jahrbuch Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, 56, 623 - 706." type="journal article" year="1906">Schubert,</bibRefCitation>
</emphasis>
1906
<emphasis id="B940105FFE59FFB2FFA6FDD0FAB8FDD2" bold="true" box="[1273,1282,523,547]" pageId="16" pageNumber="17">)</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE59FFB2F872FD90FAEEFC92" pageId="16" pageNumber="17" type="description">
<paragraph id="8B8BCC4DFE59FFB2F872FD90FC16FD92" blockId="16.[813,940,587,611]" box="[813,940,587,611]" pageId="16" pageNumber="17">
<figureCitation id="130FD0C8FE59FFB2F872FD90FC16FD92" box="[813,940,587,611]" captionStart="Fig" captionStartId="17.[168,199,1199,1219]" captionTargetBox="[152,1437,237,1310]" captionTargetId="figure-220@17.[167,1422,253,1186]" captionTargetPageId="17" captionText="Fig. 7 Otoliths of Gobionellidae: ae Deltentosteus aff. telleri (Schubert, 1906), 7a Shydlivshchyna, NMNHU-P PI 2553, b (reversed),d Kozatskyi Yar, NMNHU-P PI 2552,c (reversed), e Mlyntsi, NMB P1209.f, g Economidichthys triangularis (Weiler, 1943), f (reversed) Shydlivshchyna, NMB P1210, g (reversed) Mlyntsi, NMNHU-P PI 2554.h, i Knipowitschia polonica Schwarzhans et al., 2020a, 2020b, h Shydlivshchyna, NMNHU-P PI 2562, i Kozatskyi Yar,NMB P1213. j, k Pomatoschistus elegans (Procházka, 1900), j (reversed) Kozatskyi Yar, NMNHU-P PI 2576, k (reversed) Shydlivshchyna, NMB P1222" figureDoi="http://doi.org/10.5281/zenodo.11999917" httpUri="https://zenodo.org/record/11999917/files/figure.png" pageId="16" pageNumber="17">Figure 7ae</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2F872FD50FC45FD12" blockId="16.[813,1436,651,739]" pageId="16" pageNumber="17">
2020a
<taxonomicName id="4C34B7CEFE59FFB2F82CFD50FA90FD52" authority="(Schubert, 1906)" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[883,1322,651,675]" family="Gobiidae" genus="Deltentosteus" isUncertain="true" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="telleri">
<emphasis id="B940105FFE59FFB2F82CFD50FBBFFD52" box="[883,1029,651,675]" italics="true" pageId="16" pageNumber="17">Deltentosteus</emphasis>
aff.
<emphasis id="B940105FFE59FFB2FF6DFD50FBCBFD52" box="[1074,1137,651,675]" italics="true" pageId="16" pageNumber="17">telleri</emphasis>
(
<bibRefCitation id="EFA5B1BCFE59FFB2FF21FD50FA99FD52" author="Schubert, R. J." box="[1150,1315,651,675]" pageId="16" pageNumber="17" pagination="623 - 706" refId="ref23059" refString="Schubert, R. J. (1906). Die Fischotolithen des osterr. - ungar. Tertiars. III. Macruriden und Beryciden. Jahrbuch Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, 56, 623 - 706." type="journal article" year="1906">Schubert, 1906</bibRefCitation>
)
</taxonomicName>
—Schwarzhans, Brzobohatý &amp; Radwańska: pl. 8, figs. 1721 (and references therein).
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2F872FCD0FAEEFC92" blockId="16.[813,1437,779,867]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2F872FCD0FC34FCD2" box="[813,910,779,803]" italics="true" pageId="16" pageNumber="17">Material</emphasis>
8 otoliths:
<materialsCitation id="3B5CC610FE59FFB2FF63FCD0FBB6FCB2" collectionCode="NMNHU-P" pageId="16" pageNumber="17" specimenCode="PI 2552" specimenCount="3">
<specimenCount id="9D3207C4FE59FFB2FF63FCD0FB6CFCD2" box="[1084,1238,779,803]" count="3" pageId="16" pageNumber="17" type="generic">3 specimens</specimenCount>
, Kozatskyi Yar,
<collectionCode id="ED255488FE59FFB2F872FCF7FC17FCB2" box="[813,941,812,835]" pageId="16" pageNumber="17">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE59FFB2F8EAFCF7FBB6FCB2" box="[949,1036,811,835]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="16" pageNumber="17">PI 2552</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE59FFB2FF48FCF0FA22FCB2" box="[1047,1432,811,835]" collectionCode="NMB" pageId="16" pageNumber="17" specimenCode="P1209" specimenCount="2">
<specimenCount id="9D3207C4FE59FFB2FF48FCF0FB1BFCB2" box="[1047,1185,811,835]" count="2" pageId="16" pageNumber="17" type="generic">2 specimens</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE59FFB2FE50FCF7FAF1FCB2" box="[1295,1355,812,835]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="16" pageNumber="17">NMB</collectionCode>
<specimenCode id="DB926436FE59FFB2FE0CFCF0FA22FCB2" box="[1363,1432,811,835]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="16" pageNumber="17">P1209</specimenCode>
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;
<materialsCitation id="3B5CC610FE59FFB2F872FC90FAF5FC92" box="[813,1359,843,867]" collectionCode="NMNHU-P" pageId="16" pageNumber="17" specimenCode="PI 2553" specimenCount="3">
<specimenCount id="9D3207C4FE59FFB2F872FC90FC0FFC92" box="[813,949,843,867]" count="3" pageId="16" pageNumber="17" type="generic">3 specimens</specimenCount>
, Shydlivshchyna,
<collectionCode id="ED255488FE59FFB2FF2BFC97FB4EFC92" box="[1140,1268,844,867]" pageId="16" pageNumber="17">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE59FFB2FFA4FC97FAF5FC92" box="[1275,1359,843,867]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="16" pageNumber="17">PI 2553</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE59FFB3F872FC57FE2BF8C0" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B8BCC4DFE59FFB2F872FC57FBFEFB72" blockId="16.[813,1437,907,1923]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2F872FC57FB39FC52" box="[813,1155,907,931]" italics="true" pageId="16" pageNumber="17">
Discussion
<taxonomicName id="4C34B7CEFE59FFB2F8F6FC50FB39FC52" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[937,1155,907,931]" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="telleri">Deltentosteus telleri</taxonomicName>
</emphasis>
was a widely distributed and common species in the middle Miocene of the Tethys and Paratethys and the late Miocene of the Tethys. Te otoliths are easily recognizable by their smooth, convex inner face with a shallow sulcus, which has a much widened ostial region and ostial lobe, and narrowed caudal tip. Te otolith outline does not usually show preventral or postdorsal projections.
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2F81DFB50FAABF8B2" blockId="16.[813,1437,907,1923]" pageId="16" pageNumber="17">
Otoliths of
<taxonomicName id="4C34B7CEFE59FFB2F894FB57FB95FB52" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[971,1071,1163,1187]" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="telleri">
<emphasis id="B940105FFE59FFB2F894FB57FB95FB52" box="[971,1071,1163,1187]" italics="true" pageId="16" pageNumber="17">D. telleri</emphasis>
</taxonomicName>
show a certain degree of variability and considerable allometric ontogenetic growth. Morphological variations of large specimens have been observed by
<bibRefCitation id="EFA5B1BCFE59FFB2F8E1FB30FB33FAF2" author="Radwanska, U." box="[958,1161,1259,1283]" pageId="16" pageNumber="17" pagination="141 - 328" refId="ref22671" refString="Radwanska, U. (1992). Fish otoliths in the Middle Miocene (Badenian) deposits of southern Poland. Acta Geologica Polonica, 42 (3 - 4), 141 - 328." type="journal article" year="1992">Radwańska (1992)</bibRefCitation>
and Schwarzhans et. al. (2020a), particularly in the locality Niskowa in
<collectingCountry id="F3238CDDFE59FFB2FE14FAD0FA22FAD2" box="[1355,1432,1291,1315]" name="Poland" pageId="16" pageNumber="17">Poland</collectingCountry>
; these variations have been tentatively attributed to alterations that may have resulted “from temporary fluctuations in salinity of the coastal waters” that may occur in a marginal marine environment with varying freshwater influx (
<bibRefCitation id="EFA5B1BCFE59FFB2F820FA70FBFBFA32" author="Radwanska, U." box="[895,1089,1451,1475]" pageId="16" pageNumber="17" pagination="141 - 328" refId="ref22671" refString="Radwanska, U. (1992). Fish otoliths in the Middle Miocene (Badenian) deposits of southern Poland. Acta Geologica Polonica, 42 (3 - 4), 141 - 328." type="journal article" year="1992">Radwańska, 1992</bibRefCitation>
). Among the observed alterations in the otoliths from Niskowa were the reduction of the colliculum from the tapering cauda, reduction of the ostial lobe, and differences in otolith shape and thickness. Te largest specimen now available from the Medobory back-reef environment (
<figureCitation id="130FD0C8FE59FFB2FF69F990FB3AF992" box="[1078,1152,1611,1635]" captionStart="Fig" captionStartId="17.[168,199,1199,1219]" captionTargetBox="[152,1437,237,1310]" captionTargetId="figure-220@17.[167,1422,253,1186]" captionTargetPageId="17" captionText="Fig. 7 Otoliths of Gobionellidae: ae Deltentosteus aff. telleri (Schubert, 1906), 7a Shydlivshchyna, NMNHU-P PI 2553, b (reversed),d Kozatskyi Yar, NMNHU-P PI 2552,c (reversed), e Mlyntsi, NMB P1209.f, g Economidichthys triangularis (Weiler, 1943), f (reversed) Shydlivshchyna, NMB P1210, g (reversed) Mlyntsi, NMNHU-P PI 2554.h, i Knipowitschia polonica Schwarzhans et al., 2020a, 2020b, h Shydlivshchyna, NMNHU-P PI 2562, i Kozatskyi Yar,NMB P1213. j, k Pomatoschistus elegans (Procházka, 1900), j (reversed) Kozatskyi Yar, NMNHU-P PI 2576, k (reversed) Shydlivshchyna, NMB P1222" figureDoi="http://doi.org/10.5281/zenodo.11999917" httpUri="https://zenodo.org/record/11999917/files/figure.png" pageId="16" pageNumber="17">Fig. 7a</figureCitation>
) shares most of the characteristics observed in the specimens from Niskowa but stems from a completely different environments (reef-associated vs. marginal marine with freshwater influx). Terefore, it now seems possible that these morphotypes actually represent a different species. However, a wider review of otoliths attributed to
<taxonomicName id="4C34B7CEFE59FFB2FFFDF8D7FABDF8D2" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[1186,1287,1803,1827]" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="telleri">
<emphasis id="B940105FFE59FFB2FFFDF8D7FABDF8D2" box="[1186,1287,1803,1827]" italics="true" pageId="16" pageNumber="17">D. telleri</emphasis>
</taxonomicName>
is necessary before a conclusive assessment can be made.
</paragraph>
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Te otoliths of
<taxonomicName id="4C34B7CEFE59FFB2F8B1F890FB4BF892" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[1006,1265,1867,1891]" family="Gobiidae" genus="Deltentosteus" isUncertain="true" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="telleri">
<emphasis id="B940105FFE59FFB2F8B1F890FB3AF892" box="[1006,1152,1867,1891]" italics="true" pageId="16" pageNumber="17">Deltentosteus</emphasis>
aff.
<emphasis id="B940105FFE59FFB2FFEDF890FB4BF892" box="[1202,1265,1867,1891]" italics="true" pageId="16" pageNumber="17">telleri</emphasis>
</taxonomicName>
described here from the Medobory back reef are also remarkable in that they represent a rather complete ontogenetic sequence, which has thus far rarely been observed in the species, where the fossil record is dominated by adult specimens. Te ontogenetic sequence shows a distinct allometry, with small specimens being much more compressed than large ones. Te ratio OL:OH ranges from
<quantity id="4CCC61A8FE58FFB3F937FA22FCBDF9E0" box="[616,775,1529,1553]" metricMagnitude="-2" metricUnit="m" metricValue="2.6542999999999997" metricValueMax="2.7178" metricValueMin="2.5908" pageId="17" pageNumber="18" unit="in" value="1.045" valueMax="1.07" valueMin="1.02">1.02 to 1.07 in</quantity>
specimens of 1.0
<quantity id="4CCC61A8FE58FFB3FA00F9C2FE09F9C0" box="[351,435,1561,1585]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="18" unit="mm" value="1.4">1.4 mm</quantity>
in length (
<figureCitation id="130FD0C8FE58FFB3F971F9C2FD2FF9C0" box="[558,661,1561,1585]" captionStart="Fig" captionStartId="17.[168,199,1199,1219]" captionTargetBox="[152,1437,237,1310]" captionTargetId="figure-220@17.[167,1422,253,1186]" captionTargetPageId="17" captionText="Fig. 7 Otoliths of Gobionellidae: ae Deltentosteus aff. telleri (Schubert, 1906), 7a Shydlivshchyna, NMNHU-P PI 2553, b (reversed),d Kozatskyi Yar, NMNHU-P PI 2552,c (reversed), e Mlyntsi, NMB P1209.f, g Economidichthys triangularis (Weiler, 1943), f (reversed) Shydlivshchyna, NMB P1210, g (reversed) Mlyntsi, NMNHU-P PI 2554.h, i Knipowitschia polonica Schwarzhans et al., 2020a, 2020b, h Shydlivshchyna, NMNHU-P PI 2562, i Kozatskyi Yar,NMB P1213. j, k Pomatoschistus elegans (Procházka, 1900), j (reversed) Kozatskyi Yar, NMNHU-P PI 2576, k (reversed) Shydlivshchyna, NMB P1222" figureDoi="http://doi.org/10.5281/zenodo.11999917" httpUri="https://zenodo.org/record/11999917/files/figure.png" pageId="17" pageNumber="18">Fig. 7ce</figureCitation>
); the ratio is about
<quantity id="4CCC61A8FE58FFB3FBA9F9E2FEFAF9A0" box="[246,320,1593,1617]" metricMagnitude="-2" metricUnit="m" metricValue="2.921" pageId="17" pageNumber="18" unit="in" value="1.15">1.15 in</quantity>
a specimen of
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in length (
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) and about
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a specimen of
<quantity id="4CCC61A8FE58FFB3F96CF982FD34F980" box="[563,654,1625,1649]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="17" pageNumber="18" unit="mm" value="2.1">2.1 mm</quantity>
in length
</paragraph>
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<paragraph id="8B8BCC4DFE58FFB3FBF7FB74FA38FAE2" blockId="17.[168,1410,1199,1299]" pageId="17" pageNumber="18">
<emphasis id="B940105FFE58FFB3FBF7FB74FF61FB32" bold="true" box="[168,219,1199,1219]" pageId="17" pageNumber="18">Fig. 7</emphasis>
Otoliths of Gobionellidae:
<emphasis id="B940105FFE58FFB3FA9CFB74FE59FB32" bold="true" box="[451,483,1199,1219]" pageId="17" pageNumber="18">ae</emphasis>
<taxonomicName id="4C34B7CEFE58FFB3FAB7FB74FC8EFB32" authority="(Schubert, 1906)" baseAuthorityName="Schubert" baseAuthorityYear="1906" box="[488,820,1199,1219]" family="Gobiidae" genus="Deltentosteus" isUncertain="true" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="telleri">
<emphasis id="B940105FFE58FFB3FAB7FB74FDEDFB32" box="[488,599,1199,1219]" italics="true" pageId="17" pageNumber="18">Deltentosteus</emphasis>
aff.
<emphasis id="B940105FFE58FFB3F926FB74FD1FFB32" box="[633,677,1199,1219]" italics="true" pageId="17" pageNumber="18">telleri</emphasis>
(
<bibRefCitation id="EFA5B1BCFE58FFB3F9F2FB74FC95FB32" author="Schubert, R. J." box="[685,815,1199,1219]" pageId="17" pageNumber="18" pagination="623 - 706" refId="ref23059" refString="Schubert, R. J. (1906). Die Fischotolithen des osterr. - ungar. Tertiars. III. Macruriden und Beryciden. Jahrbuch Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, 56, 623 - 706." type="journal article" year="1906">Schubert, 1906</bibRefCitation>
)
</taxonomicName>
, 7a Shydlivshchyna, NMNHU-P PI 2553,
<emphasis id="B940105FFE58FFB3FFDCFB74FB35FB32" bold="true" box="[1155,1167,1199,1219]" pageId="17" pageNumber="18">b</emphasis>
(reversed),
<emphasis id="B940105FFE58FFB3FFAFFB74FB46FB32" bold="true" box="[1264,1276,1199,1219]" pageId="17" pageNumber="18">d</emphasis>
Kozatskyi Yar, NMNHU-P PI 2552,
<emphasis id="B940105FFE58FFB3FA15FB11FEEEFB2F" bold="true" box="[330,340,1226,1246]" pageId="17" pageNumber="18">c</emphasis>
(reversed),
<emphasis id="B940105FFE58FFB3FAEAFB11FE7AFB2F" bold="true" box="[437,448,1226,1246]" pageId="17" pageNumber="18">e</emphasis>
Mlyntsi, NMB P1209.
<emphasis id="B940105FFE58FFB3F92BFB11FDC1FB2F" bold="true" box="[628,635,1226,1246]" pageId="17" pageNumber="18">f</emphasis>
,
<emphasis id="B940105FFE58FFB3F9DBFB11FD2AFB2F" bold="true" box="[644,656,1226,1246]" pageId="17" pageNumber="18">g</emphasis>
<taxonomicName id="4C34B7CEFE58FFB3F9CBFB11FC43FB2F" authority="(Weiler, 1943)" baseAuthorityName="Weiler" baseAuthorityYear="1943" box="[660,1017,1226,1246]" family="Gobiidae" genus="Economidichthys" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="triangularis">
<emphasis id="B940105FFE58FFB3F9CBFB11FC3BFB2F" box="[660,897,1226,1246]" italics="true" pageId="17" pageNumber="18">Economidichthys triangularis</emphasis>
(
<bibRefCitation id="EFA5B1BCFE58FFB3F8D5FB11FC49FB2F" author="Weiler, W." box="[906,1011,1226,1246]" pageId="17" pageNumber="18" pagination="87 - 115" refId="ref24030" refString="Weiler, W. (1943). Die Otolithen aus dem Jungtertiar SUd- Rumaniens, I. Buglow Und Sarmat. Senckenbergiana Lethaea, 26, 87 - 115." type="journal article" year="1943">Weiler, 1943</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B940105FFE58FFB3FF5EFB11FBB2FB2F" bold="true" box="[1025,1032,1226,1246]" pageId="17" pageNumber="18">f</emphasis>
(reversed) Shydlivshchyna, NMB P1210,
<emphasis id="B940105FFE58FFB3FE04FB11FADDFB2F" bold="true" box="[1371,1383,1226,1246]" pageId="17" pageNumber="18">g</emphasis>
(reversed) Mlyntsi, NMNHU-P PI 2554.
<emphasis id="B940105FFE58FFB3FAB7FB3EFE4EFB08" bold="true" box="[488,500,1253,1273]" pageId="17" pageNumber="18">h</emphasis>
,
<emphasis id="B940105FFE58FFB3FAA2FB3EFDB8FB08" bold="true" box="[509,514,1253,1273]" pageId="17" pageNumber="18">i</emphasis>
<taxonomicName id="4C34B7CEFE58FFB3F958FB3EFC1BFB08" authority="Schwarzhans et al., 2020 a" authorityName="Schwarzhans" authorityYear="2020" box="[519,929,1253,1273]" family="Gobiidae" genus="Knipowitschia" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="polonica">
<emphasis id="B940105FFE58FFB3F958FB3EFD7FFB08" box="[519,709,1253,1273]" italics="true" pageId="17" pageNumber="18">Knipowitschia polonica</emphasis>
<bibRefCitation id="EFA5B1BCFE58FFB3F996FB3EFC1BFB08" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[713,929,1253,1273]" pageId="17" pageNumber="18" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFA5B1BCFE58FFB3F8F6FB3EFC64FB08" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[937,990,1253,1273]" pageId="17" pageNumber="18" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
,
<emphasis id="B940105FFE58FFB3F8B9FB3EFC48FB08" bold="true" box="[998,1010,1253,1273]" pageId="17" pageNumber="18">h</emphasis>
Shydlivshchyna, NMNHU-P PI 2562,
<emphasis id="B940105FFE58FFB3FE7AFB3EFA90FB08" bold="true" box="[1317,1322,1253,1273]" pageId="17" pageNumber="18">i</emphasis>
Kozatskyi Yar,NMB P1213.
<emphasis id="B940105FFE58FFB3FA6BFB24FE80FAE2" bold="true" box="[308,314,1279,1299]" pageId="17" pageNumber="18">j</emphasis>
,
<emphasis id="B940105FFE58FFB3FA1DFB24FEF7FAE2" bold="true" box="[322,333,1279,1299]" pageId="17" pageNumber="18">k</emphasis>
<taxonomicName id="4C34B7CEFE58FFB3FA0EFB24FD14FAE2" authority="(Prochazka, 1900)" baseAuthorityName="Prochazka" baseAuthorityYear="1900" box="[337,686,1279,1299]" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B940105FFE58FFB3FA0EFB24FDA9FAE2" box="[337,531,1279,1299]" italics="true" pageId="17" pageNumber="18">Pomatoschistus elegans</emphasis>
(Procházka, 1900)
</taxonomicName>
,
<emphasis id="B940105FFE58FFB3F9EBFB24FD00FAE2" bold="true" box="[692,698,1279,1299]" pageId="17" pageNumber="18">j</emphasis>
(reversed) Kozatskyi Yar, NMNHU-P PI 2576,
<emphasis id="B940105FFE58FFB3FF73FB24FB8DFAE2" bold="true" box="[1068,1079,1279,1299]" pageId="17" pageNumber="18">k</emphasis>
(reversed) Shydlivshchyna, NMB P1222
</paragraph>
</caption>
<paragraph id="8B8BCC4DFE58FFB3FBC8F9A2FE2BF8C0" blockId="17.[151,776,1369,1841]" pageId="17" pageNumber="18">
(
<figureCitation id="130FD0C8FE58FFB3FBC0F9A2FF4AF960" box="[159,240,1657,1681]" captionStart="Fig" captionStartId="17.[168,199,1199,1219]" captionTargetBox="[152,1437,237,1310]" captionTargetId="figure-220@17.[167,1422,253,1186]" captionTargetPageId="17" captionText="Fig. 7 Otoliths of Gobionellidae: ae Deltentosteus aff. telleri (Schubert, 1906), 7a Shydlivshchyna, NMNHU-P PI 2553, b (reversed),d Kozatskyi Yar, NMNHU-P PI 2552,c (reversed), e Mlyntsi, NMB P1209.f, g Economidichthys triangularis (Weiler, 1943), f (reversed) Shydlivshchyna, NMB P1210, g (reversed) Mlyntsi, NMNHU-P PI 2554.h, i Knipowitschia polonica Schwarzhans et al., 2020a, 2020b, h Shydlivshchyna, NMNHU-P PI 2562, i Kozatskyi Yar,NMB P1213. j, k Pomatoschistus elegans (Procházka, 1900), j (reversed) Kozatskyi Yar, NMNHU-P PI 2576, k (reversed) Shydlivshchyna, NMB P1222" figureDoi="http://doi.org/10.5281/zenodo.11999917" httpUri="https://zenodo.org/record/11999917/files/figure.png" pageId="17" pageNumber="18">Fig. 7a</figureCitation>
). Likewise, the crenulation of the dorsal rim, albeit weak, is only evident in specimens of about
<quantity id="4CCC61A8FE58FFB3F997F942FF50F920" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="17" pageNumber="18" unit="mm" value="1.5" valueMax="1.8" valueMin="1.2">1.2 to 1.8 mm</quantity>
in length (
<figureCitation id="130FD0C8FE58FFB3FA01F962FE7BF920" box="[350,449,1721,1745]" captionStart="Fig" captionStartId="17.[168,199,1199,1219]" captionTargetBox="[152,1437,237,1310]" captionTargetId="figure-220@17.[167,1422,253,1186]" captionTargetPageId="17" captionText="Fig. 7 Otoliths of Gobionellidae: ae Deltentosteus aff. telleri (Schubert, 1906), 7a Shydlivshchyna, NMNHU-P PI 2553, b (reversed),d Kozatskyi Yar, NMNHU-P PI 2552,c (reversed), e Mlyntsi, NMB P1209.f, g Economidichthys triangularis (Weiler, 1943), f (reversed) Shydlivshchyna, NMB P1210, g (reversed) Mlyntsi, NMNHU-P PI 2554.h, i Knipowitschia polonica Schwarzhans et al., 2020a, 2020b, h Shydlivshchyna, NMNHU-P PI 2562, i Kozatskyi Yar,NMB P1213. j, k Pomatoschistus elegans (Procházka, 1900), j (reversed) Kozatskyi Yar, NMNHU-P PI 2576, k (reversed) Shydlivshchyna, NMB P1222" figureDoi="http://doi.org/10.5281/zenodo.11999917" httpUri="https://zenodo.org/record/11999917/files/figure.png" pageId="17" pageNumber="18">Fig. 7b, c</figureCitation>
). Te characteristic reduction of the colliculum in the cauda, however, is a stable trait in all sizes observed and could possibly represent a diagnostically valuable feature.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,126 @@
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<mods:title id="0D0978A5471CBFE3134066862EEA8257">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<paragraph id="8B8BCC4DFE5BFFB0FBC8FB50FDC7FB52" blockId="18.[151,637,1162,1187]" box="[151,637,1162,1187]" pageId="18" pageNumber="19">
<heading id="D0C37B21FE5BFFB0FBC8FB50FDC7FB52" bold="true" box="[151,637,1162,1187]" fontSize="10" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C34B7CEFE5BFFB0FBC8FB50FDC7FB52" authority="(Prochazka, 1900)" baseAuthorityName="Prochazka" baseAuthorityYear="1900" box="[151,637,1162,1187]" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B940105FFE5BFFB0FBC8FB50FDC7FB52" bold="true" box="[151,637,1162,1187]" pageId="18" pageNumber="19">
<emphasis id="B940105FFE5BFFB0FBC8FB50FE1FFB52" bold="true" box="[151,421,1163,1187]" italics="true" pageId="18" pageNumber="19">Pomatoschistus elegans</emphasis>
(Procházka, 1900)
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<figureCitation id="130FD0C8FE5BFFB0F872FF30FC12FEF2" box="[813,936,235,259]" captionStart="Fig" captionStartId="17.[168,199,1199,1219]" captionTargetBox="[152,1437,237,1310]" captionTargetId="figure-220@17.[167,1422,253,1186]" captionTargetPageId="17" captionText="Fig. 7 Otoliths of Gobionellidae: ae Deltentosteus aff. telleri (Schubert, 1906), 7a Shydlivshchyna, NMNHU-P PI 2553, b (reversed),d Kozatskyi Yar, NMNHU-P PI 2552,c (reversed), e Mlyntsi, NMB P1209.f, g Economidichthys triangularis (Weiler, 1943), f (reversed) Shydlivshchyna, NMB P1210, g (reversed) Mlyntsi, NMNHU-P PI 2554.h, i Knipowitschia polonica Schwarzhans et al., 2020a, 2020b, h Shydlivshchyna, NMNHU-P PI 2562, i Kozatskyi Yar,NMB P1213. j, k Pomatoschistus elegans (Procházka, 1900), j (reversed) Kozatskyi Yar, NMNHU-P PI 2576, k (reversed) Shydlivshchyna, NMB P1222" figureDoi="http://doi.org/10.5281/zenodo.11999917" httpUri="https://zenodo.org/record/11999917/files/figure.png" pageId="18" pageNumber="19">Figure 7jk</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE5BFFB0F872FEF0FB8DFE72" blockId="18.[813,1436,299,387]" pageId="18" pageNumber="19">
2020a
<taxonomicName id="4C34B7CEFE5BFFB0F8D9FEF0FA3BFEB2" authority="(Prochazka, 1900)" baseAuthorityName="Prochazka" baseAuthorityYear="1900" box="[902,1409,299,323]" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B940105FFE5BFFB0F8D9FEF0FB2EFEB2" box="[902,1172,299,323]" italics="true" pageId="18" pageNumber="19">Pomatoschistus elegans</emphasis>
(Procházka, 1900)
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— Schwarzhans, Brzobohatý &amp; Radwańska: pl. 9, figs. 2022 (and references therein).
</paragraph>
<paragraph id="8B8BCC4DFE5BFFB0F872FE70FAD3FE12" blockId="18.[813,1436,427,483]" pageId="18" pageNumber="19">
<emphasis id="B940105FFE5BFFB0F872FE70FC34FE32" box="[813,910,427,451]" italics="true" pageId="18" pageNumber="19">Material</emphasis>
<materialsCitation id="3B5CC610FE5BFFB0F8CCFE70FC2FFE12" collectionCode="PI" pageId="18" pageNumber="19" specimenCode="PI 2576" specimenCount="1">
2 otoliths:
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, Kozatskyi Yar, NMNHU- P
<specimenCode id="DB926436FE5BFFB0F81DFE17FC2FFE12" box="[834,917,459,483]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="18" pageNumber="19">PI 2576</specimenCode>
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;
<materialsCitation id="3B5CC610FE5BFFB0F8FFFE10FADFFE12" box="[928,1381,459,483]" collectionCode="NMB" pageId="18" pageNumber="19" specimenCode="P1222" specimenCount="1">
<specimenCount id="9D3207C4FE5BFFB0F8FFFE10FBA7FE12" box="[928,1053,459,483]" count="1" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
, Shydlivshchyna,
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<specimenCode id="DB926436FE5BFFB0FE40FE10FADFFE12" box="[1311,1381,459,483]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="18" pageNumber="19">P1222</specimenCode>
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.
</paragraph>
<paragraph id="8B8BCC4DFE5BFFB0F872FDD7FA9DFC32" blockId="18.[813,1437,523,963]" pageId="18" pageNumber="19">
<emphasis id="B940105FFE5BFFB0F872FDD7FC1AFDD2" box="[813,928,524,547]" italics="true" pageId="18" pageNumber="19">Discussion</emphasis>
Te small otoliths of
<taxonomicName id="4C34B7CEFE5BFFB0FFC5FDD7FAB9FDD2" baseAuthorityName="Prochazka" baseAuthorityYear="1900" box="[1178,1283,523,547]" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B940105FFE5BFFB0FFC5FDD7FAB9FDD2" box="[1178,1283,523,547]" italics="true" pageId="18" pageNumber="19">P. elegans</emphasis>
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, which rarely exceed
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in length, are characterized by a nearly quadratic otolith shape, with the exception of the broad, obtuse middorsal angle, and a flat inner face with a slightly eccentrically positioned sulcus that is narrow and steeply inclined at over 30°.
<taxonomicName id="4C34B7CEFE5BFFB0FFC2FD70FA26FD32" baseAuthorityName="Prochazka" baseAuthorityYear="1900" box="[1181,1436,683,707]" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B940105FFE5BFFB0FFC2FD70FA26FD32" box="[1181,1436,683,707]" italics="true" pageId="18" pageNumber="19">Pomatoschistus elegans</emphasis>
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has been observed relatively rarely in the early Badenian of the Central Paratethys, and these are the first records from the late Badenian. Te specimens of
<taxonomicName id="4C34B7CEFE5BFFB0FE71FCD7FA26FCD2" baseAuthorityName="Prochazka" baseAuthorityYear="1900" box="[1326,1436,779,803]" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B940105FFE5BFFB0FE71FCD7FA26FCD2" box="[1326,1436,779,803]" italics="true" pageId="18" pageNumber="19">P. elegans</emphasis>
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described here from the Medobory back-reef environment differ from those of the early Badenian in being more thickset (OH:OT = 2.32.4 vs. 3.43.6), but this difference is not supported by any other features and, therefore, is regarded as an expression of variability.
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indet.
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Figure
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<emphasis id="B940105FFE5FFFB4FBC8FEF0FF42FEB2" box="[151,248,299,323]" italics="true" pageId="22" pageNumber="23">Material</emphasis>
1 otolith, Shydlivshchyna,
<collectionCode id="ED255488FE5FFFB4F944FEF7FDEDFEB2" box="[539,599,300,323]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="22" pageNumber="23">NMB</collectionCode>
P1221.
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<emphasis id="B940105FFE5FFFB4FBC8FEB7FEAEFE72" box="[151,276,364,387]" italics="true" pageId="22" pageNumber="23">Description</emphasis>
A single, somewhat eroded, massive and roundish otolith of
<quantity id="4CCC61A8FE5FFFB4FADEFE50FE66FE52" box="[385,476,395,419]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="23" unit="mm" value="2.5">2.5 mm</quantity>
in length. OL:OH = 1.12; OH:OT = 2.7. Ventral rim gently curving; dorsal rim high, with distinct predorsal angle. Rostrum broad, blunt, about 24% of OL; no excisura or antirostrum. Posterior rim rounded.
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4FBF2FDF0FDE7FCD2" blockId="22.[151,775,363,1027]" pageId="22" pageNumber="23">
Smooth and convex inner face with slightly supramedian, moderately deepened sulcus. Ostium wider and shorter than cauda; cauda very slightly flexed and reaching close to posterior rim of otolith. CaL:OsL = 1.15; OsH:CaH = 1.6; OL:SuL= 1.1. No ventral furrow; small but distinct dorsal depression. Outer face convex with postcentral umbo and very distinct and unusual outward oriented hump at rostral tip (
<figureCitation id="130FD0C8FE5FFFB4FA8CFCD0FDA7FCD2" box="[467,541,779,803]" captionStart="Fig" captionStartId="21.[168,199,1587,1607]" captionTargetBox="[152,1438,237,1698]" captionTargetId="figure-47@21.[167,1422,253,1574]" captionTargetPageId="21" captionText="Fig. 9 Otoliths of Haemulidae, Serranidae, Sparidae, Leiognathidae and Caproidae:a (reversed)—Brachydeuterus speronatus (Bassoli,1906), Kozatskyi Yar, NMNHU-P PI 2549. b, c Serranidae indet., b (broken and repaired), c (reversed), Kozatskyi Yar,NMNHU-P PI 2579. d, e Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, Mlyntsi, NMNHU-P PI 2578. f (reversed)—Leiognathidae indet., Shydlivshchyna, NMB P1214. g Perciformes indet., Shydlivshchyna,NMB P1221. h (reversed) Capros aper (Linnaeus,1758), Mlyntsi, NMB P1206" figureDoi="http://doi.org/10.5281/zenodo.11999944" httpUri="https://zenodo.org/record/11999944/files/figure.png" pageId="22" pageNumber="23">
Fig.
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</figureCitation>
<figureCitation id="130FD0C8FE5FFFB4F943FCD0FD90FCD2" box="[540,554,779,803]" captionStart="Fig" captionStartId="3.[168,199,1780,1800]" captionTargetBox="[152,1438,237,1839]" captionTargetId="figure-11@3.[170,1419,253,1767]" captionTargetPageId="3" captionText="Fig. 2 Photographs of the investigated outcrops:Mlyntsi (a, b), Kozatskyi Yar (c, d), and Shydlivshchyna (e, f) in general view and close-up.The total length of the shovel handle is 72 cm" figureDoi="http://doi.org/10.5281/zenodo.11999851" httpUri="https://zenodo.org/record/11999851/files/figure.png" pageId="22" pageNumber="23">2</figureCitation>
, g
<figureCitation id="130FD0C8FE5FFFB4F91CFCD0FDEAFCD2" box="[579,592,779,803]" captionStart="Fig" captionStartId="6.[168,199,1875,1895]" captionTargetBox="[153,1438,1176,1934]" captionTargetId="figure-236@6.[168,1423,1192,1862]" captionTargetPageId="6" captionText="Fig. 3 Otoliths of Gadiformes: a, b Micromesistius planatus (Bassoli &amp; Schubert, 1906), Mlyntsi, a NMNHU-P PI 2568 and b NMB P1218. cf Onogadus simplicissimus (Schubert, 1906), c (reversed), d, f Shydlivshchyna, NMB P1220, e Mlyntsi, NMNHU-P PI 2572" figureDoi="http://doi.org/10.5281/zenodo.11999867" httpUri="https://zenodo.org/record/11999867/files/figure.png" pageId="22" pageNumber="23">3</figureCitation>
).
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4FBF2FCF7FE09FBF2" blockId="22.[151,775,363,1027]" pageId="22" pageNumber="23">
<emphasis id="B940105FFE5FFFB4FBF2FCF7FE9AFCB2" box="[173,288,812,835]" italics="true" pageId="22" pageNumber="23">Discussion</emphasis>
Tis otolith is characterized by the extremely unusual outward-oriented hump at the tip of the rostrum, which to the best of our knowledge has not been reported from any other extant or fossil perciform otolith. It cannot be determined at present whether this feature has diagnostic value or represents some kind of teratological deformation.
</paragraph>
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@ -0,0 +1,118 @@
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indet.
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<subSubSection id="C32E9FC6FE5DFFB7F872FBF0FDCEF890" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="8B8BCC4DFE5DFFB6F872FBF0FC37FBB2" blockId="20.[813,909,1067,1091]" box="[813,909,1067,1091]" pageId="20" pageNumber="21">
<figureCitation id="130FD0C8FE5DFFB6F872FBF0FC37FBB2" box="[813,909,1067,1091]" captionStart="Fig" captionStartId="21.[168,199,1587,1607]" captionTargetBox="[152,1438,237,1698]" captionTargetId="figure-47@21.[167,1422,253,1574]" captionTargetPageId="21" captionText="Fig. 9 Otoliths of Haemulidae, Serranidae, Sparidae, Leiognathidae and Caproidae:a (reversed)—Brachydeuterus speronatus (Bassoli,1906), Kozatskyi Yar, NMNHU-P PI 2549. b, c Serranidae indet., b (broken and repaired), c (reversed), Kozatskyi Yar,NMNHU-P PI 2579. d, e Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, Mlyntsi, NMNHU-P PI 2578. f (reversed)—Leiognathidae indet., Shydlivshchyna, NMB P1214. g Perciformes indet., Shydlivshchyna,NMB P1221. h (reversed) Capros aper (Linnaeus,1758), Mlyntsi, NMB P1206" figureDoi="http://doi.org/10.5281/zenodo.11999944" httpUri="https://zenodo.org/record/11999944/files/figure.png" pageId="20" pageNumber="21">Figure 9f</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE5DFFB6F872FBB0FA87FB72" blockId="20.[813,1341,1131,1155]" box="[813,1341,1131,1155]" pageId="20" pageNumber="21">
<emphasis id="B940105FFE5DFFB6F872FBB0FC34FB72" box="[813,910,1131,1155]" italics="true" pageId="20" pageNumber="21">Material</emphasis>
1 otolith, Shydlivshchyna,
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P1214.
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<paragraph id="8B8BCC4DFE5DFFB6F872FB77FB7AFA72" blockId="20.[813,1437,1195,1891]" pageId="20" pageNumber="21">
<emphasis id="B940105FFE5DFFB6F872FB77FC10FB32" box="[813,938,1196,1219]" italics="true" pageId="20" pageNumber="21">Description</emphasis>
A small roundish otolith of
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in length. OL:OH = 1.2; OH:OT = 4.0. Ventral and dorsal rim regularly curved, semicircular, but dorsal rim shorter than ventral rim, both without angles but irregularly crenulated. Rostrum broad, blunt, 22% of OL; excisura and antirostrum distinct, antirostrum much shorter than rostrum. Posterior rim regularly curved.
</paragraph>
<paragraph id="8B8BCC4DFE5DFFB6F81DFA57FC77F952" blockId="20.[813,1437,1195,1891]" pageId="20" pageNumber="21">Inner face distinctly convex in horizontal direction, with moderately deep and long supramedian sulcus. Ostium slightly wider and shorter than cauda; CaL:OsL = 1.05; OsH:CaH = 1.2. Cauda very slightly flexed and posterodorsally widened, reaching close to posterior rim of otolith. OL:SuL = 1.12. No discernable ventral furrow; small but distinct dorsal depression. Outer face mildly concave, smooth except for few short radial furrows.</paragraph>
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<emphasis id="B940105FFE5DFFB6F81DF977FC0FF932" box="[834,949,1708,1731]" italics="true" pageId="20" pageNumber="21">Discussion</emphasis>
Tis characteristic otolith may represent an undescribed species of an undefined leiognathid genus (see
<bibRefCitation id="EFA5B1BCFE5DFFB6F800F930FB95F8F2" author="Lin, C. - H. &amp; Chang, C. - W." box="[863,1071,1771,1795]" pageId="20" pageNumber="21" pagination="1 - 415" refId="ref21462" refString="Lin, C. - H., &amp; Chang, C. - W. (2012). Otolith atlas of Taiwan fishes. National Museum of Marine Biology and Aquarium, 12, 1 - 415." type="journal article" year="2012">Lin &amp; Chang, 2012</bibRefCitation>
for figures of extant otoliths). As for this group, a set of skeletal imprints of leiognathids has been described from the early Miocene (Burdigalian) of
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(
<bibRefCitation id="EFA5B1BCFE5DFFB6F8F8F890FB25F892" author="Kovalchuk, O. &amp; Swidnicka, E. &amp; Stefaniak, K." box="[935,1183,1867,1891]" pageId="20" pageNumber="21" pagination="421 - 428" refId="ref21276" refString="Kovalchuk, O., Swidnicka, E., &amp; Stefaniak, K. (2021). Early Miocene ponyfishes (Acanthuriformes, Leiognathidae) of the Carpathian Basin. Palaeontological Journal, 55 (4), 421 - 428. https: // doi. org / 10.1134 / S 003103012 1040092" type="journal article" year="2021">Kovalchuk et al., 2021</bibRefCitation>
). Tese specimens are morphologically identical and similar in meristic values to
<taxonomicName id="4C34B7CEFE5CFFB7FBE2F8D2FE73F8D0" box="[189,457,1801,1825]" genus="Leiognathoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B940105FFE5CFFB7FBE2F8D2FE73F8D0" box="[189,457,1801,1825]" italics="true" pageId="21" pageNumber="22">Leiognathoides minutus</emphasis>
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described from the Great Caucasian Basin (
<bibRefCitation id="EFA5B1BCFE5CFFB7FA3AF8F2FDA9F8B0" author="Bannikov, A. F." box="[357,531,1833,1857]" pageId="21" pageNumber="22" pagination="112 - 114" refId="ref19160" refString="Bannikov, A. F. (2001). A new genus of ponyfish (Perciformes, Leiognathidae). Journal of Ichthyology, 41 (1), 112 - 114." type="journal article" year="2001">Bannikov, 2001</bibRefCitation>
,
<bibRefCitation id="EFA5B1BCFE5CFFB7F978F8F2FDE1F8B0" author="Bannikov, A. F." box="[551,603,1833,1857]" pageId="21" pageNumber="22" pagination="1 - 244" refId="ref19194" refString="Bannikov, A. F. (2010). Fossil acanthopterygian fishes (Teleostei, Acanthopterygii). In L. P. Tatarinov, E. I. Vorobyeva, &amp; E. N. Kurochkin (Eds.), Fossil vertebrates of Russia and adjacent countries (pp. 1 - 244). GEOS. (in Russian, with English summary)." type="book chapter" year="2010">2010</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE5CFFB7F930F8F2FF71F890" author="Daniltshenko, P. G." pageId="21" pageNumber="22" pagination="115 - 121" refId="ref20057" refString="Daniltshenko, P. G. (1980). Order Perciformes (excluding the Percidae), fossil bony fishes of the USSR. Trudy Paleontologicheskogo Instituta AN SSSR, 178, 115 - 121 and 125 - 169. (in Russian)." type="journal article" year="1980">Daniltshenko, 1980</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE5CFFB7FB87F892FE08F890" author="Micklich, N. &amp; Bannikov, A. F. &amp; Yabumoto, Y." box="[216,434,1865,1889]" pageId="21" pageNumber="22" pagination="375 - 398" refId="ref21688" refString="Micklich, N., Bannikov, A. F., &amp; Yabumoto, Y. (2017). First record of ponyfishes (Perciformes: Leiognathidae) from the Oligocene of the Grube Unterfeld (&quot; Frauenweiler' ') clay pit. Palaontologische Zeitschrift, 91, 375 - 398." type="journal article" year="2017">Micklich et al., 2017</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE5CFFB7FAE1F892FDDFF890" author="Prokofiev, A. M." box="[446,613,1865,1889]" pageId="21" pageNumber="22" refId="ref22625" refString="Prokofiev, A. M. (2002). New and little known perciform fishes from the" type="book" year="2002">Prokofiev, 2002</bibRefCitation>
).
</paragraph>
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<mods:title id="7C12583D2D0C568D7F9BE80544A5CAA2">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="1982414E70A393890356246A9B3B597F">2022</mods:date>
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<mods:number id="05E074B7D2F441FBF46A069410CD8519">141</mods:number>
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<mods:url id="411254138A711FE18BFDEC72EAABB1FD">http://dx.doi.org/10.1186/s13358-022-00261-3</mods:url>
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<treatment id="039D7D5BFE5EFFBAF872FAB0FF60FC52" LSID="urn:lsid:plazi:treatment:039D7D5BFE5EFFBAF872FAB0FF60FC52" httpUri="http://treatment.plazi.org/id/039D7D5BFE5EFFBAF872FAB0FF60FC52" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<subSubSection id="C32E9FC6FE5EFFB5F872FAB0FB17FA72" box="[813,1197,1387,1411]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8B8BCC4DFE5EFFB5F872FAB0FB17FA72" blockId="23.[813,1197,1387,1411]" box="[813,1197,1387,1411]" pageId="23" pageNumber="24">
<heading id="D0C37B21FE5EFFB5F872FAB0FB17FA72" bold="true" box="[813,1197,1387,1411]" fontSize="10" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B940105FFE5EFFB5F872FAB0FB17FA72" bold="true" box="[813,1197,1387,1411]" pageId="23" pageNumber="24">
Genus
<taxonomicName id="4C34B7CEFE5EFFB5F822FAB0FB17FA72" ID-CoL="3ZY5" authority="Rafinesque, 1810" authorityName="Rafinesque" authorityYear="1810" box="[893,1197,1387,1411]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE5EFFB5F822FAB0FC58FA72" bold="true" box="[893,994,1387,1411]" italics="true" pageId="23" pageNumber="24">Dasyatis</emphasis>
Rafinesque, 1810
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</emphasis>
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</paragraph>
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<subSubSection id="C32E9FC6FE5EFFB5F872FA70FC02FA32" box="[813,952,1451,1475]" pageId="23" pageNumber="24" type="multiple">
<paragraph id="8B8BCC4DFE5EFFB5F872FA70FC02FA32" blockId="23.[813,952,1451,1475]" box="[813,952,1451,1475]" pageId="23" pageNumber="24">
<heading id="D0C37B21FE5EFFB5F872FA70FC02FA32" bold="true" box="[813,952,1451,1475]" fontSize="10" level="1" pageId="23" pageNumber="24" reason="1">
<taxonomicName id="4C34B7CEFE5EFFB5F872FA70FC02FA32" box="[813,952,1451,1475]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE5EFFB5F872FA70FC02FA32" bold="true" box="[813,952,1451,1475]" pageId="23" pageNumber="24">
<emphasis id="B940105FFE5EFFB5F872FA70FC28FA32" bold="true" box="[813,914,1451,1475]" italics="true" pageId="23" pageNumber="24">Dasyatis</emphasis>
sp.
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B8BCC4DFE5EFFB5F872FA30FC20F9F2" blockId="23.[813,922,1515,1539]" box="[813,922,1515,1539]" pageId="23" pageNumber="24">
<figureCitation id="130FD0C8FE5EFFB5F872FA30FC20F9F2" box="[813,922,1515,1539]" captionStart="Fig" captionStartId="22.[169,200,1823,1843]" captionTargetBox="[153,1438,1393,1909]" captionTargetId="figure-434@22.[168,1423,1409,1811]" captionTargetPageId="22" captionText="Fig. 10 Fish teeth from the studied localities: ae Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions. f Dasyatis sp., anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)" figureDoi="http://doi.org/10.5281/zenodo.11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="23" pageNumber="24">Figure 10f</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE5EFFB5F872F9F0FA31F9B2" blockId="23.[813,1419,1579,1603]" box="[813,1419,1579,1603]" pageId="23" pageNumber="24">
<emphasis id="B940105FFE5EFFB5F872F9F0FC34F9B2" box="[813,910,1579,1603]" italics="true" pageId="23" pageNumber="24">Material</emphasis>
1 anterior tooth, Mlyntsi,
<collectionCode id="ED255488FE5EFFB5FFF3F9F7FA96F9B2" box="[1196,1324,1580,1603]" pageId="23" pageNumber="24">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE5EFFB5FE6DF9F7FAF0F9B2" box="[1330,1354,1580,1603]" country="Russia" name="Paleontological Institute" pageId="23" pageNumber="24">PI</collectionCode>
2587.
</paragraph>
<paragraph id="8B8BCC4DFE5EFFBAF872F9B7FD4DFDF2" blockId="23.[813,1437,1643,1891]" lastBlockId="24.[151,776,235,931]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="B940105FFE5EFFB5F872F9B7FC10F972" box="[813,938,1644,1667]" italics="true" pageId="23" pageNumber="24">Description</emphasis>
Te tooth is small and wider than it is high (1.50 vs.
<quantity id="4CCC61A8FE5EFFB5F8D0F950FC49F952" box="[911,1011,1675,1699]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="24" unit="mm" value="0.9">0.90 mm</quantity>
). A clearly recognizable bulging transverse crest separates the labial and lingual faces in the middle of the crown. Te globular and non-cuspidate crown suggests that the described specimen belongs to a stingray female (
<bibRefCitation id="EFA5B1BCFE5EFFB5FF47F8D0FB7AF8D2" author="Cappetta, H." box="[1048,1216,1803,1827]" pageId="23" pageNumber="24" refId="ref19805" refString="Cappetta, H. (2012). Handbook of paleoichthyology, Vol. 3 E: Chondrichthyes - Mesozoic and Cenozoic Elasmobranchii: Teeth (p. 512). Verlag Dr. Friedrich Pfeil." type="book" year="2012">Cappetta, 2012</bibRefCitation>
). Te labial visor is partly broken, and no ornamentation can be observed on the labial face. Te smooth lingual face is divided into two marginal lingual areas by a median lingual ridge. Tere are distinct marginal angles, one of which is more rounded, while the other is sharper. Te convex lingual visor partially overhangs the labial face of the root. Te latter is bilobed; its lobes slightly differ in width and are displaced lingually. Tere is a deep but quite narrow notch separating the lobes and reaching half of the root height. Weakly separated and wide root lobes also indicate that the tooth belongs to a female (
<bibRefCitation id="EFA5B1BCFE51FFBAF91CFE30FD52FDF2" author="Cappetta, H." box="[579,744,491,515]" pageId="24" pageNumber="25" refId="ref19805" refString="Cappetta, H. (2012). Handbook of paleoichthyology, Vol. 3 E: Chondrichthyes - Mesozoic and Cenozoic Elasmobranchii: Teeth (p. 512). Verlag Dr. Friedrich Pfeil." type="book" year="2012">Cappetta, 2012</bibRefCitation>
).
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<paragraph id="8B8BCC4DFE51FFBAFBF2FDD7FF60FC52" blockId="24.[151,776,235,931]" pageId="24" pageNumber="25">
<emphasis id="B940105FFE51FFBAFBF2FDD7FE9AFDD2" box="[173,288,524,547]" italics="true" pageId="24" pageNumber="25">Discussion</emphasis>
Te specimen described shares diagnostic features of the genus
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<emphasis id="B940105FFE51FFBAFADCFDF7FE5BFDB2" box="[387,481,556,579]" italics="true" pageId="24" pageNumber="25">Dasyatis</emphasis>
Rafinesque, 1810
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, and differs from representatives of the genus
<taxonomicName id="4C34B7CEFE51FFBAF90EFD97FEB1FD72" authority="Garman, 1913" authorityName="Garman" authorityYear="1913" class="Elasmobranchii" family="Dasyatidae" genus="Taeniurops" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE51FFBAF90EFD97FD70FD92" box="[593,714,588,611]" italics="true" pageId="24" pageNumber="25">Taeniurops</emphasis>
Garman, 1913
</taxonomicName>
, in the non-hollowed labial face lacking sharp crests. However, it cannot be attributed to a particular species due to its poor preservation. Te middle Miocene fossil record of stingrays of the genus
<taxonomicName id="4C34B7CEFE51FFBAF969FD17FD2EFD12" authorityName="Rafinesque" authorityYear="1810" box="[566,660,716,739]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE51FFBAF969FD17FD2EFD12" box="[566,660,716,739]" italics="true" pageId="24" pageNumber="25">Dasyatis</emphasis>
</taxonomicName>
in Europe is represented by
<taxonomicName id="4C34B7CEFE51FFBAFA01FD37FD21FCF2" authority="Cappetta, 1970" authorityName="Cappetta" authorityYear="1970" box="[350,667,747,771]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="delfortriei">
<emphasis id="B940105FFE51FFBAFA01FD37FE51FCF2" box="[350,491,747,771]" italics="true" pageId="24" pageNumber="25">D. delfortriei</emphasis>
<bibRefCitation id="EFA5B1BCFE51FFBAFAABFD30FD21FCF2" author="Cappetta, H." box="[500,667,747,771]" pageId="24" pageNumber="25" pagination="1 - 139" refId="ref19778" refString="Cappetta, H. (1970). Les Selachiens du Miocene de la region de Montpellier. Palaeovertebrata, Memoire extraordinaire, 1 - 139." type="book chapter" year="1970">Cappetta, 1970</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C34B7CEFE51FFBAF9F4FD37FED8FCD2" authority="Cappetta, 1970" authorityName="Cappetta" authorityYear="1970" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="probsti">
<emphasis id="B940105FFE51FFBAF9F4FD37FF0AFCD2" italics="true" pageId="24" pageNumber="25">D. probsti</emphasis>
<bibRefCitation id="EFA5B1BCFE51FFBAFBE5FCD0FED8FCD2" author="Cappetta, H." box="[186,354,779,803]" pageId="24" pageNumber="25" pagination="1 - 139" refId="ref19778" refString="Cappetta, H. (1970). Les Selachiens du Miocene de la region de Montpellier. Palaeovertebrata, Memoire extraordinaire, 1 - 139." type="book chapter" year="1970">Cappetta, 1970</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C34B7CEFE51FFBAFAFAFCD7FD0DFCD2" authority="(Probst, 1877)" baseAuthorityName="Probst" baseAuthorityYear="1877" box="[421,695,779,803]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B940105FFE51FFBAFAFAFCD7FDB4FCD2" box="[421,526,780,803]" italics="true" pageId="24" pageNumber="25">D. rugosa</emphasis>
(Probst, 1877)
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, which have been variably reported from the early Badenian of Korytnica,
<collectingCountry id="F3238CDDFE51FFBAFA4EFC90FEE6FC92" box="[273,348,843,867]" name="Poland" pageId="24" pageNumber="25">Poland</collectingCountry>
(
<bibRefCitation id="EFA5B1BCFE51FFBAFA35FC90FE47FC92" author="Schultz, O." box="[362,509,843,867]" pageId="24" pageNumber="25" pagination="201 - 209" refId="ref23099" refString="Schultz, O. (1977). Elasmobranch and teleost fish remains from the Korytnica Clays (Middle Miocene; Holy Cross Mountains, Poland). Acta Geologica Polonica, 27 (2), 201 - 209." type="journal article" year="1977">Schultz, 1977</bibRefCitation>
,
<bibRefCitation id="EFA5B1BCFE51FFBAF956FC90FD87FC92" author="Schultz, O." box="[521,573,843,867]" pageId="24" pageNumber="25" pagination="287 - 293" refId="ref23140" refString="Schultz, O. (1979). Supplementary notes on elasmobranch and teleost fish remains from the Korytnica Clays (Middle Miocene; Holy Cross Mountains, Poland). Acta Geologica Polonica, 29 (3), 287 - 293." type="journal article" year="1979">1979</bibRefCitation>
) and from the late Badenian of Weissenegg and Wurzing in
<collectingCountry id="F3238CDDFE51FFBAF908FCB0FD1DFC72" box="[599,679,875,899]" name="Austria" pageId="24" pageNumber="25">Austria</collectingCountry>
(
<bibRefCitation id="EFA5B1BCFE51FFBAF9EAFCB0FF71FC52" author="Hilden, H. R." pageId="24" pageNumber="25" pagination="41 - 110" refId="ref20763" refString="Hilden, H. R. (1995). Elasmobranchier (Pisces, Chondrichthyes) aus dem Badenium (Mittleres Miozan) des Steirischen Beckens (Osterreich). Mitteilungen der Abteilung fur Geologie und Palaontologie am Landesmuseum Johanneum, 52 (53), 41 - 110." type="journal article" year="1995">Hilden, 1995</bibRefCitation>
).
</paragraph>
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<mods:affiliation id="5338BA4D5817DEC9EA6AAB1E9FD7134F">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
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<subSubSection id="C88DABE113EA3CC2AD41B83EF80D7B31" type="nomenclature">
@ -99,8 +101,13 @@ Boruah, Narayanan, Deepak &amp; Das
</heading>
</paragraph>
<paragraph id="B61575AA4DECB8C95540F059045F4859">
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Adult female (WII-ADR 971), from Athibung (
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<specimenCount id="1110C6980182BA2D049625A9DA3F6E05" count="1" type="female">Adult female</specimenCount>
(
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), from
<location id="556B3B54E6261C08813B1D540A901F40" LSID="urn:lsid:plazi:treatment:0987463F2E2D509F88594C782A30D269:556B3B54E6261C08813B1D540A901F40" county="Peren District" latitude="25.54199" longLatPrecision="5" longitude="93.6307" name="Athibung" stateProvince="Nagaland">Athibung</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECTAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[93.630700,25.541990]}">
<geoCoordinate id="C48E58CB783A9E1E667ECE6E75C9B6AA" degrees="25.54199" direction="north" orientation="latitude" precision="1" value="25.54199">25.54199 ° N</geoCoordinate>
;
@ -115,12 +122,14 @@ a. s. l.
) (Fig.
<figureCitation id="E533ABC182595ED938F3C30C6D134557" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
),
<location id="FF6612D812061757FC57E42A61FB3A60" LSID="urn:lsid:plazi:treatment:0987463F2E2D509F88594C782A30D269:FF6612D812061757FC57E42A61FB3A60" latitude="25.54199" longLatPrecision="5" longitude="93.6307" name="Peren District" stateProvince="Nagaland">Peren District</location>
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,
<collectingRegion id="9E413FE8D03EB5A5C3AAB2BF5D1D4092" country="India" name="Nagaland">Nagaland</collectingRegion>
, collected by
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and Bitupan Boruah on
and
<collectorName id="BB05A905BC2FA54D9484C2BF852F7083">Bitupan Boruah</collectorName>
on
<date id="227153EA81FE12A333F3BD32F70642D7" value="2021-08-14">
<collectingDate id="3EC9E7BCF7086D4E4C945D513629488F" value="2021-08-14">14 August 2021</collectingDate>
</date>
@ -225,7 +234,9 @@ Holotype of
<emphasis id="4F17B258138F82E2088D92DA6A928304" bold="true">
<taxonomicNameLabel id="7261F079EED76924C624281285129106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(WII-ADR 971).
(
<specimenCode id="EC715245A4BD8530056E107194BA1350">WII-ADR 971</specimenCode>
).
<emphasis id="5430F8A719136F5BDFAB24384D985926" bold="true">A</emphasis>
dorsal and
<emphasis id="4B94EFE71FB7DEF1CF0A3C6F47C87106" bold="true">B</emphasis>
@ -550,11 +561,7 @@ has a genetic divergence of 10.7 % and 22.3 % in the ND 2 gene.
<paragraph id="C2A055C2F6F4AEA250E6D8B9A45B8DA5">
<heading id="545AAB5F77882AB9FF882F5C4B59E367" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C1C74D550F578A7714F3003B736CF333">
The specific epithet is a toponym derived from the name of the hill range “ Barail ” where the
<typeStatus id="8403B414FEEA81ACF2858CDD9587830F">type</typeStatus>
locality of the species lies.
</paragraph>
<paragraph id="C1C74D550F578A7714F3003B736CF333">The specific epithet is a toponym derived from the name of the hill range “ Barail ” where the type locality of the species lies.</paragraph>
</subSubSection>
<subSubSection id="SECID0EBIBK" type="Suggested common name">
<paragraph id="B867BA96A8DB7210668A22F50726F75C">
@ -567,9 +574,7 @@ locality of the species lies.
<heading id="21E9BC883F8FA3AA01577E1C27A45E43" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="52480E70FB1130A6E26EB4A6DCE59551">
During our two-day survey we only located a single individual of this species. Thus, the new species is currently known only from the
<typeStatus id="EAD1E9EDF68DE046B4B0BE2BB0CCB8B5">type</typeStatus>
locality, in Peren District,
During our two-day survey we only located a single individual of this species. Thus, the new species is currently known only from the type locality, in Peren District,
<collectingRegion id="7D3CE58919ED460C35F986413B85302C" country="India" name="Nagaland">Nagaland</collectingRegion>
,
<collectingCountry id="3F15ED38E3DFB440AE8EF583A2F29F16" name="India">India</collectingCountry>
@ -577,9 +582,7 @@ locality, in Peren District,
<quantity id="6C06FA77D4CB18C006A6B24CB3944FDD" metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">2 m</quantity>
from the ground in the Athibung Reserve Forest at approximately 20: 00 hrs on
<date id="08E9E3E3025C754C49F50712CD4C17D7" value="2021-08-14">14 August 2021</date>
. The forest
<typeStatus id="797C4907F968C66CD5481DD696FADD08">type</typeStatus>
is semi-evergreen with relatively little anthropogenic pressure.
. The forest type is semi-evergreen with relatively little anthropogenic pressure.
</paragraph>
</subSubSection>
</treatment>

View file

@ -1,78 +1,80 @@
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<mods:title id="32ED38DE159AA5107D6862C18B799C88">Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India</mods:title>
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@ -99,10 +101,15 @@ Boruah, Narayanan, Lalronunga, Deepak &amp; Das
</heading>
</paragraph>
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Adult male (WII-ADR 1057; Fig.
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<specimenCount id="5A96676A852391AC54F311C212F99A0B" count="1" type="male">Adult male</specimenCount>
(
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; Fig.
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), from Ngengpui Wildlife Sanctuary (
), from
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@ -117,14 +124,18 @@ a. s. l.
) (Fig.
<figureCitation id="3880332778AFA36BFF0C2D1599353545" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
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collected by
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, Bitupan Boruah and Samuel Lalronunga on
,
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and
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on
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@ -146,7 +157,9 @@ collected by
<emphasis id="98DAC7B51925693A33E4D59C2CBB7F64" bold="true">A</emphasis>
<emphasis id="F3D90BE3D97EEBA3C8DEFF2E057C0839" bold="true">J</emphasis>
holotype (ADR-WII 1057),
holotype (
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),
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dorsal and
<emphasis id="D7864A54964BCD14E4267B0005ECE3E1" bold="true">B</emphasis>
@ -166,9 +179,13 @@ right pes,
<emphasis id="DA0479C623A637DA3314FA8CE62398BA" bold="true">I</emphasis>
precloacofemoral pores;
<emphasis id="FBD5865EA2AE63B124C970E6D89E4F8F" bold="true">J</emphasis>
holotype (ADR-WII 1057), and
holotype (
<specimenCode id="3FAC1BF21477368AB586F26353DB8F5C">ADR-WII 1057</specimenCode>
), and
<emphasis id="84D05F05D82233A864CA7A4AAD4D2046" bold="true">K</emphasis>
paratype (WII-ADR 991) in life. Scale bar: 10 mm.
paratype (
<specimenCode id="5B96FB8E94CF577AC07F9FE85AE06EF0">WII-ADR 991</specimenCode>
) in life. Scale bar: 10 mm.
<emphasis id="F0CC3B570F73174CE5E012A84384D1BD" bold="true">J, K</emphasis>
not to scale.
</paragraph>
@ -183,9 +200,15 @@ not to scale.
</paragraph>
<paragraph id="D0CFD4E8796C3198E29114BFF63A9EB4">
<specimenCount id="8CAE3C4A05313DFD2FD3F763D9B9B4AB" count="2" type="adult">Two adult</specimenCount>
females (WII-ADR 991 and WII-ADR 1058) and
females (
<specimenCode id="8A478F6B5458AC2E2C23665DA95DFA9D">WII-ADR 991</specimenCode>
and
<specimenCode id="DABE00B9E9DDB8158838081D1C87E703">WII-ADR 1058</specimenCode>
) and
<specimenCount id="DC1BA10AF099CF6AC67745612ABB086D" count="1" type="subadult">one subadult</specimenCount>
female (WII-ADR 1059) collected from the same locality in Ngengpui Wildlife Sanctuary by the same team on
female (
<specimenCode id="A940522E8EF8089995090CB89C1FAF88">WII-ADR 1059</specimenCode>
) collected from the same locality in Ngengpui Wildlife Sanctuary by the same team on
<date id="2F13415877074AE32BB76A4E4CE4F96F" value="2021-09-07" valueMax="2021-09-08" valueMin="2021-09-07">7 and 8 September 2021</date>
.
</paragraph>
@ -295,7 +318,11 @@ Details of the variations in morphometric and meristic characters of the type se
<typeStatus id="7BB7CC918C817442284EAB77A85A7A87" type="paratype">paratypes</typeStatus>
slightly varied. In the
<specimenCount id="3CF2D2A17A0896245BA9A60942906D33" count="2" type="generic" typeStatus="paratype">two paratypes</specimenCount>
(WII-ADR 1058 and WII-ADR 1059), dorsal bands are broken mid dorsally giving the appearance of enlarged spots (Fig. S 2). Precloacal pores in female are smaller than that of male.
(
<specimenCode id="CD8734A42EA7B55F3243066F9D6EF73F">WII-ADR 1058</specimenCode>
and
<specimenCode id="54BC92C000869203C6165E30A4BB86DB">WII-ADR 1059</specimenCode>
), dorsal bands are broken mid dorsally giving the appearance of enlarged spots (Fig. S 2). Precloacal pores in female are smaller than that of male.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHLCK" type="comparisons">
@ -495,9 +522,7 @@ is 0.8 %.
<paragraph id="469226927F875902E1CCFB888B17437C">
The specific epithet is a toponym derived from the name “ Ngengpui Wildlife Sanctuary ” of
<collectingRegion id="BA1398980158B38BB5F47273A9604F98" country="India" name="Mizoram">Mizoram state</collectingRegion>
from where the
<typeStatus id="66C9BF3FF107A35FD1C445E1EFFEB11B">type</typeStatus>
series of the species were collected.
from where the type series of the species were collected.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUXCK" type="Suggested common name">
@ -517,9 +542,7 @@ series of the species were collected.
<emphasis id="CBFFA2613E31F760F8DA5DB26A850958" bold="true">
<taxonomicNameLabel id="A29B6B0A61186B394694229C7218225D" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
new species is currently only known from the
<typeStatus id="27077758B0BE48388362EC39323B1E03">type</typeStatus>
locality, the Ngengpui Wildlife Sanctuary, Lawngtlai District,
new species is currently only known from the type locality, the Ngengpui Wildlife Sanctuary, Lawngtlai District,
<collectingRegion id="5B9C8A266B21E10253524F2EBC70E953" country="India" name="Mizoram">Mizoram</collectingRegion>
,
<collectingCountry id="6E7C31C7FDABCC656C3D104C1AD2DBA6" name="India">India</collectingCountry>

View file

@ -1,78 +1,80 @@
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@ -124,113 +126,249 @@
(Fig.
<figureCitation id="E6C15AE00D7800CAE646D20BFBA5DCE9" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyrtodactylus cayuensis from India showing the variation in dorsal marking pattern. A WII-ADR 1219 (Balek), B WII-ADR 1681 (Ezengo), C WII-ADR 1199 (Jengging), D WII-ADR 1218 (Balek), E WII-ADR 454 (Jengging), F WII-ADR 1213 (Ramsing), G WII-ADR 698 (Potin), H WII-ADR 3017 (Glaw lake), I precloacal pores in WII-ADR 698, and J precloacal pores in WII-ADR 478. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure10" httpUri="https://binary.pensoft.net/fig/1102276">10</figureCitation>
).
<materialsCitation id="27423835FA9120BC03746BE1907164BC" collectingDate="2021-10-29" collectorName="Bitupan Boruah" county="East Siang District" elevation="450" latitude="28.0624" location="Balek village" longLatPrecision="6" longitude="95.2721" specimenCode="WII-ADR 1219, WII-ADR 1218" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<collectingRegion id="99A66E4C22AA6BF8E1F495EBA10EFD16" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
:
<specimenCount id="207B96D2CFF269066E33A683CAAD9077" count="1" type="male">one male</specimenCount>
(WII-ADR 1219) and
(
<specimenCode id="1804A9900E4D2A535280C0AEF2026CAE">WII-ADR 1219</specimenCode>
) and
<specimenCount id="0A778DBEE8C1EFB27BFF071EC46F7DBA" count="1" type="female">one female</specimenCount>
(WII-ADR 1218) collected from Balek village (
(
<specimenCode id="1716D2BD94936D68FB797DA6A72EC3CA">WII-ADR 1218</specimenCode>
) collected from
<location id="911E10464BE356B0697B1AC21806E265" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:911E10464BE356B0697B1AC21806E265" county="East Siang District" latitude="28.0624" longLatPrecision="6" longitude="95.2721" name="Balek village" stateProvince="Arunachal Pradesh">Balek village</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP2DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.272100,28.062400]}">
<geoCoordinate id="75F2584ACA713396B799F29F9F69F5DB" degrees="28.0624" direction="north" orientation="latitude" precision="5" value="28.0624">28.0624 ° N</geoCoordinate>
;
<geoCoordinate id="BCA63C1A6B619CF6C8232BCA11A09999" degrees="95.2721" direction="east" orientation="longitude" precision="5" value="95.2721">95.2721 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="C59C3841943AC9D1DFF398EB93A9851E" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</quantity>
a. s. l.), East Siang District on
<date id="31BBBA95F5569F63A1A04F48C439847A" value="2021-10-29">29 October 2021</date>
by Bitupan Boruah;
<quantity id="C59C3841943AC9D1DFF398EB93A9851E" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">
<elevation id="ADA5222086D2DDFDC292738CCBD9AF7B" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</elevation>
</quantity>
a. s. l.),
<collectingCounty id="E0F4B81FFFA20B8C648EF952CB3FA546">East Siang District</collectingCounty>
on
<date id="31BBBA95F5569F63A1A04F48C439847A" value="2021-10-29">
<collectingDate id="C7E2CD2C68BFEBAE404D05CD3923EA9E" value="2021-10-29">29 October 2021</collectingDate>
</date>
by
<collectorName id="13127AFA01A633DCAC5B9991972D7A9B">Bitupan Boruah</collectorName>
</materialsCitation>
;
<materialsCitation id="E229B9ABC76B19618239BDD35279C19C" collectingDate="2021-10-27" collectorName="Bitupan Boruah" county="Upper Siang District" elevation="600" latitude="28.6563" location="Ramsing" longLatPrecision="6" longitude="94.9795" municipality="Mouling National Park" specimenCode="WII-ADR 1213" specimenCount="1" specimenCount-male="1">
<specimenCount id="29BB93CF6B2F8B184B29200C53119BAC" count="1" type="male">one male</specimenCount>
(WII-ADR 1213) collected from Ramsing (
(
<specimenCode id="8FB8649EBA539B4D94CE1BABAEA27BB9">WII-ADR 1213</specimenCode>
) collected from
<location id="16ECD38C1363D9D7E8D781AFEB7BA951" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:16ECD38C1363D9D7E8D781AFEB7BA951" county="Upper Siang District" latitude="28.6563" longLatPrecision="6" longitude="94.9795" municipality="Mouling National Park" name="Ramsing">Ramsing</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX2DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[94.979500,28.656300]}">
<geoCoordinate id="F8897D5244653885B97E9D1D28ECD03E" degrees="28.6563" direction="north" orientation="latitude" precision="5" value="28.6563">28.6563 ° N</geoCoordinate>
;
<geoCoordinate id="04D1B3D647C89A53608C01DB4B714064" degrees="94.9795" direction="east" orientation="longitude" precision="5" value="94.9795">94.9795 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="E2579E4E2FCD3554D27DA29809C53FF5" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
a. s. l.), Mouling National Park, Upper Siang District on
<date id="E92BB40106586A5FFB897F30283C97E7" value="2021-10-27">27 October 2021</date>
by Bitupan Boruah;
<quantity id="E2579E4E2FCD3554D27DA29809C53FF5" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<elevation id="BA9CFE34AA33CB8C013180C81BE6164E" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
</quantity>
a. s. l.),
<collectingMunicipality id="4D919A7DAC2AC0D9183E78A8709F6551">Mouling National Park</collectingMunicipality>
,
<collectingCounty id="5F5B8F773D20F4948AB81BE0E703DD0C">Upper Siang District</collectingCounty>
on
<date id="E92BB40106586A5FFB897F30283C97E7" value="2021-10-27">
<collectingDate id="9DD8D2DD18D0DBC1FCCC17C8A3512BEB" value="2021-10-27">27 October 2021</collectingDate>
</date>
by
<collectorName id="673FFB03647E18BE1FCF004792C76000">Bitupan Boruah</collectorName>
</materialsCitation>
;
<materialsCitation id="D165CE4A13A46BBB314F5ACA9E95DFA2" collectingDate="2021-10-26" collectorName="Bitupan Boruah" county="Upper Siang District" elevation="890" latitude="28.534" location="Syrnyup stream" longLatPrecision="6" longitude="95.0305" municipality="Mouling National Park" specimenCode="WII-ADR 1199" specimenCount="1" specimenCount-female="1">
<specimenCount id="56B7E3F556DB4553B1339FE72622FB02" count="1" type="female">one female</specimenCount>
(WII-ADR 1199) collected from Syrnyup stream (
(
<specimenCode id="DCE65F3C20C6479C7CB26FDF81D4A476">WII-ADR 1199</specimenCode>
) collected from
<location id="B9DBB4D3953A70CE46310ECB31ED0880" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:B9DBB4D3953A70CE46310ECB31ED0880" county="Upper Siang District" latitude="28.534" longLatPrecision="6" longitude="95.0305" municipality="Mouling National Park" name="Syrnyup stream">Syrnyup stream</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E62DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.030500,28.534000]}">
<geoCoordinate id="BE2A74361CE825ABA05338B581420200" degrees="28.5340" direction="north" orientation="latitude" precision="5" value="28.534">28.5340 ° N</geoCoordinate>
;
<geoCoordinate id="73979654104F37809B98C863027B4DCC" degrees="95.0305" direction="east" orientation="longitude" precision="5" value="95.0305">95.0305 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="329965E1309EFA79A6BE2D2C4F30306D" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</quantity>
a. s. l.), Jengging, Mouling National Park, Upper Siang District on
<date id="74002D5F7A7C46E2989AE8955CB16213" value="2021-10-26">26 October 2021</date>
by Bitupan Boruah;
<quantity id="329965E1309EFA79A6BE2D2C4F30306D" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">
<elevation id="A705DD74FD0AECCA068ECD6E558E3DC7" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</elevation>
</quantity>
a. s. l.), Jengging,
<collectingMunicipality id="B183E89183816B55A69D7BEEDC1BC36C">Mouling National Park</collectingMunicipality>
,
<collectingCounty id="463E28675463337FBD20375D8F581C07">Upper Siang District</collectingCounty>
on
<date id="74002D5F7A7C46E2989AE8955CB16213" value="2021-10-26">
<collectingDate id="B522F5C184C97DF851266B371C9F6658" value="2021-10-26">26 October 2021</collectingDate>
</date>
by
<collectorName id="0DC957D79B9489ADCB61E480E068E1CA">Bitupan Boruah</collectorName>
</materialsCitation>
;
<materialsCitation id="32CACEF5259ED0ADAC1E828E78F40F0E" collectingDate="2019-10-05" collectorName="Bitupan Boruah" county="Lower Subansiri District" elevation="900" latitude="27.3478" location="Potin" longLatPrecision="6" longitude="93.8497" specimenCode="WII-ADR 695, WII-ADR 696, WII-ADR 697, WII-ADR 698" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="EAB5A5201F674947B54FDCDEF0964A98" count="2" type="male">two males</specimenCount>
(WII-ADR 695 and WII-ADR 696) and
(
<specimenCode id="36DFAD4A3EBF153BA6CCA44A58B830B0">WII-ADR 695</specimenCode>
and
<specimenCode id="83055EEAC2F7167563C8D0B33ECAA251">WII-ADR 696</specimenCode>
) and
<specimenCount id="FB615ABC298C53CF136A47668ADFB78A" count="2" type="female">two females</specimenCount>
(WII-ADR 697 and WII-ADR 698) collected from Potin (
(
<specimenCode id="793BB520EB0EAB885BF909519FEF6426">WII-ADR 697</specimenCode>
and
<specimenCode id="6E95969667AD016CCE6FBEE324603613">WII-ADR 698</specimenCode>
) collected from
<location id="873DE383CB0A4C3B1643BDAFD94BCF3C" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:873DE383CB0A4C3B1643BDAFD94BCF3C" county="Lower Subansiri District" latitude="27.3478" longLatPrecision="6" longitude="93.8497" name="Potin">Potin</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH3DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[93.849700,27.347800]}">
<geoCoordinate id="1BCAEB6108EB205809C1C1F14E4A1871" degrees="27.3478" direction="north" orientation="latitude" precision="5" value="27.3478">27.3478 ° N</geoCoordinate>
,
<geoCoordinate id="6104781A052483D83FE2646A0385D20F" degrees="93.8497" direction="east" orientation="longitude" precision="5" value="93.8497">93.8497 ° E</geoCoordinate>
</named-content>
, elevation
<quantity id="EDCC4D11652435D48665772D276076BE" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</quantity>
a. s. l.), Lower Subansiri District on
<date id="3DB9D5066C035EB365FCE3DDDAD0EF60" value="2019-10-05">5 October 2019</date>
by Bitupan Boruah;
<quantity id="EDCC4D11652435D48665772D276076BE" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<elevation id="AA8969AFB05B5F3F4898F5E51336075B" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
</quantity>
a. s. l.),
<collectingCounty id="AE669176F3D9E8F4ACBB1A95B60F6AE0">Lower Subansiri District</collectingCounty>
on
<date id="3DB9D5066C035EB365FCE3DDDAD0EF60" value="2019-10-05">
<collectingDate id="4C09CCF0DE5A3DBB588CC3615B815C32" value="2019-10-05">5 October 2019</collectingDate>
</date>
by
<collectorName id="3D22E29845D4B355533F52842802CA2A">Bitupan Boruah</collectorName>
</materialsCitation>
;
<materialsCitation id="1DCAB4BF28E5CB3AC207CD45479EE6E5" collectingDate="2022-09-02" collectorName="Abhijit Das &amp; Bitupan Boruah &amp; Naitik G. Patel" county="Lohit District" elevation="1200" latitude="27.696" location="Glaw lake" longLatPrecision="6" longitude="96.4456" municipality="Kamlang Tiger Reserve" specimenCode="WII-ADR 3017" specimenCount="1" specimenCount-female="1">
<specimenCount id="CB5AF0605439ECC8A81E9883A036C211" count="1" type="female">one female</specimenCount>
(WII-ADR 3017) collected near Glaw lake (
(
<specimenCode id="DB78616540335B649D09DB0EC648BC7C">WII-ADR 3017</specimenCode>
) collected near
<location id="062DABF1DC33BCC5C75D555D326179CD" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:062DABF1DC33BCC5C75D555D326179CD" county="Lohit District" latitude="27.696" longLatPrecision="6" longitude="96.4456" municipality="Kamlang Tiger Reserve" name="Glaw lake">Glaw lake</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP3DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.445600,27.696000]}">
<geoCoordinate id="F5115D947AFC324DBEF425E7401A6282" degrees="27.6960" direction="north" orientation="latitude" precision="5" value="27.696">27.6960 ° N</geoCoordinate>
;
<geoCoordinate id="9A314D5B0239E88C73437EAE8BC461A0" degrees="96.4456" direction="east" orientation="longitude" precision="5" value="96.4456">96.4456 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="B883FAB4D2C42BEFB836CC65304E01F5" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
a. s. l.), Kamlang Tiger Reserve, Lohit District on
<date id="0DCBC3E5C73A607C47E838073B1A0244" value="2022-09-02">2 September 2022</date>
by Abhijit Das, Bitupan Boruah and Naitik G. Patel;
<quantity id="B883FAB4D2C42BEFB836CC65304E01F5" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="4A994FEBE3711D04009E269F2E45EB9C" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
a. s. l.),
<collectingMunicipality id="4DE101EFE781DDCDD37AC4B0E5F2FE0A">Kamlang Tiger Reserve</collectingMunicipality>
,
<collectingCounty id="AA2CC8CE1FC5AABD34CC4B7DA50B3B72">Lohit District</collectingCounty>
on
<date id="0DCBC3E5C73A607C47E838073B1A0244" value="2022-09-02">
<collectingDate id="435C1D9A68E9C47BC2DAB07B0DEEC964" value="2022-09-02">2 September 2022</collectingDate>
</date>
by
<collectorName id="B202F211D4D63B2EE998FFE850DAB15F">Abhijit Das</collectorName>
,
<collectorName id="B81F0CD3592E389716072B06CFBC46D2">Bitupan Boruah</collectorName>
and
<collectorName id="45769729D5FB971CBE30562772893BAB">Naitik G. Patel</collectorName>
</materialsCitation>
;
<materialsCitation id="1305104981AC4C12B6A7559E98D03942" collectingDate="2022-08-03" collectorName="Bitupan Boruah" county="Lower Dibang Valley District" elevation="560" latitude="28.1565" location="Ezengo" longLatPrecision="6" longitude="95.8638" specimenCode="WII-ADR 1682, WII-ADR 1681" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="5983550108D8B5E41FDE6B9164BB818A" count="1" type="male">one male</specimenCount>
(WII-ADR 1682) and
(
<specimenCode id="858B2B52BBD3663BFACBC669CDCC3AEE">WII-ADR 1682</specimenCode>
) and
<specimenCount id="1AB4A844CCC6CBF52C5565AD354F2AEE" count="1" type="female">one female</specimenCount>
(WII-ADR 1681) collected from Ezengo (
(
<specimenCode id="5D029F531EF7801E2BE083CA389A276B">WII-ADR 1681</specimenCode>
) collected from
<location id="4ABB099D5D23C291D15947BC877CC84B" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:4ABB099D5D23C291D15947BC877CC84B" county="Lower Dibang Valley District" latitude="28.1565" longLatPrecision="6" longitude="95.8638" name="Ezengo">Ezengo</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX3DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.863800,28.156500]}">
<geoCoordinate id="F70328F07D5627C626CF9F702A505183" degrees="28.1565" direction="north" orientation="latitude" precision="5" value="28.1565">28.1565 ° N</geoCoordinate>
;
<geoCoordinate id="D48505D8A2B7956AAB23489B3B2D144F" degrees="95.8638" direction="east" orientation="longitude" precision="5" value="95.8638">95.8638 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="E6094C6516B8B1883D3E9FD0551BCC67" metricMagnitude="2" metricUnit="m" metricValue="5.6" unit="m" value="560.0">560 m</quantity>
a. s. l.), Lower Dibang Valley District on
<date id="7D91172C428B060DC773E804AB60CC11" value="2022-08-03">3 August 2022</date>
By Bitupan Boruah;
<quantity id="E6094C6516B8B1883D3E9FD0551BCC67" metricMagnitude="2" metricUnit="m" metricValue="5.6" unit="m" value="560.0">
<elevation id="FFC07782A348C9FDE8C1C17D49DC827B" metricMagnitude="2" metricUnit="m" metricValue="5.6" unit="m" value="560.0">560 m</elevation>
</quantity>
a. s. l.),
<collectingCounty id="4894E6527A358A900A2E36654462272A">Lower Dibang Valley District</collectingCounty>
on
<date id="7D91172C428B060DC773E804AB60CC11" value="2022-08-03">
<collectingDate id="40022B40847441137358A16FC6CD20F2" value="2022-08-03">3 August 2022</collectingDate>
</date>
By
<collectorName id="7492C581C0F64676440E4FA1A851A591">Bitupan Boruah</collectorName>
</materialsCitation>
;
<materialsCitation id="56C106E407859FB4D44BE23E50A896B9" collectingDate="2018-10-05" collectorName="Abhijit Das" elevation="760" latitude="28.5499" location="Jengging" longLatPrecision="6" longitude="95.0537" specimenCode="WII-ADR 453, WII-ADR 454, WII-ADR 459" specimenCount="3" specimenCount-female="3">
<specimenCount id="36E563588AA5A6D34FEA761A5BE7CC3A" count="3" type="female">three females</specimenCount>
(WII-ADR 453, WII-ADR 454 and WII-ADR 459) collected from Jengging (
(
<specimenCode id="EC26D2BA284480D353A5AD7F11C5DFC5">WII-ADR 453</specimenCode>
,
<specimenCode id="625128E0700D58BA68B383C3535F4BF6">WII-ADR 454</specimenCode>
and
<specimenCode id="06D299170E1D01E638A9AEF1114760F6">WII-ADR 459</specimenCode>
) collected from
<location id="C34AA3E207E05AF705DCA3EC1CA81F5B" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:C34AA3E207E05AF705DCA3EC1CA81F5B" latitude="28.5499" longLatPrecision="6" longitude="95.0537" name="Jengging">Jengging</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E63DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.053700,28.549900]}">
<geoCoordinate id="98E6CAB88A3E1B58DEE3628294964A8B" degrees="28.5499" direction="north" orientation="latitude" precision="5" value="28.5499">28.5499 ° N</geoCoordinate>
;
<geoCoordinate id="7D9F321ECC03AE9DDE319284FD073E46" degrees="95.0537" direction="east" orientation="longitude" precision="5" value="95.0537">95.0537 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="C4F79E803110B8E81A1353CFCA55D0E8" metricMagnitude="2" metricUnit="m" metricValue="7.6" unit="m" value="760.0">760 m</quantity>
a. s. l.) by Abhijit Das on
<date id="3B5BC1D537033D4E28C6066DB9C3DBB1" value="2018-10-05">5 October 2018</date>
<quantity id="C4F79E803110B8E81A1353CFCA55D0E8" metricMagnitude="2" metricUnit="m" metricValue="7.6" unit="m" value="760.0">
<elevation id="00A8B231ECF7E35B9E1613BF82843190" metricMagnitude="2" metricUnit="m" metricValue="7.6" unit="m" value="760.0">760 m</elevation>
</quantity>
a. s. l.) by
<collectorName id="DF35D793663BAEDFA316355B088AABC1">Abhijit Das</collectorName>
on
<date id="3B5BC1D537033D4E28C6066DB9C3DBB1" value="2018-10-05">
<collectingDate id="C6A82A9F5152350DC3D4FD5A4DEE4622" value="2018-10-05">5 October 2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="162777AAA2204626E7C57FDC84F1C954" collectingDate="2018-10-02" collectorName="Abhijit Das" elevation="410" latitude="28.0945" location="Pasighat" longLatPrecision="6" longitude="95.2682" specimenCode="WII-ADR 478, WII-ADR 473" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="A35CB19E2045DBDF80E2832C93610F27" count="1" type="female">one female</specimenCount>
(WII-ADR 478) and
(
<specimenCode id="D3648F7E07DF123D9CD32CB757FD87F7">WII-ADR 478</specimenCode>
) and
<specimenCount id="BDF452773A67C8CCEE1F3E25336696B3" count="1" type="male">one male</specimenCount>
(WII-ADR 473) collected from
<quantity id="515323FACB6D73565D972F6E03CEE122" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
northwest to Pasighat (
(
<specimenCode id="5B17DED857C029BCE118D4273655FB35">WII-ADR 473</specimenCode>
) collected from
<quantity id="515323FACB6D73565D972F6E03CEE122" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">
<locationDeviation id="FEBC7F023FE72A451A149EA0FA2079EF" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</locationDeviation>
</quantity>
northwest to
<location id="1B733FD36BD57F34F56A9F3B236F5FE5" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C:1B733FD36BD57F34F56A9F3B236F5FE5" latitude="28.0945" longLatPrecision="6" longitude="95.2682" name="Pasighat">Pasighat</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH4DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.268200,28.094500]}">
<geoCoordinate id="7E5A365D295C37F2863527C9CA23DA69" degrees="28.0945" direction="north" orientation="latitude" precision="5" value="28.0945">28.0945 ° N</geoCoordinate>
;
<geoCoordinate id="28D82508028FD3D230C39435D8479B15" degrees="95.2682" direction="east" orientation="longitude" precision="5" value="95.2682">95.2682 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="5B9B9B3D86A396CEDD0E49D29616382A" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</quantity>
a. s. l.) by Abhijit Das on
<date id="EE44B8C8746D60186591CDB47D6ED123" value="2018-10-02">2 October 2018</date>
<quantity id="5B9B9B3D86A396CEDD0E49D29616382A" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">
<elevation id="EEE59AF5CD32C63816F4E9DCBAF335BC" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</elevation>
</quantity>
a. s. l.) by
<collectorName id="31D84AB6119838A2CDD0126338F7D059">Abhijit Das</collectorName>
on
<date id="EE44B8C8746D60186591CDB47D6ED123" value="2018-10-02">
<collectingDate id="41F3574B33AFDE200DFE8C8BAB8380B5" value="2018-10-02">2 October 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="9A6CE0EB7C8B2849EA5740F64A78FD75" ID-DOI="10.3897/vz.74.e124752.figure10" ID-arpha="85FD849D-6749-530E-B888-2043F84A1248" httpUri="https://binary.pensoft.net/fig/1102276" startId="F10">
@ -243,25 +381,37 @@ a. s. l.) by Abhijit Das on
</taxonomicName>
from India showing the variation in dorsal marking pattern.
<emphasis id="9DB13E9CF5B9CE758EB75D173582D238" bold="true">A</emphasis>
WII-ADR 1219 (Balek),
<specimenCode id="9282746426AFD6EE8CA17A9A72840D5A">WII-ADR 1219</specimenCode>
(Balek),
<emphasis id="AC25C0262069E488206AEC361CCC7F6A" bold="true">B</emphasis>
WII-ADR 1681 (Ezengo),
<specimenCode id="3300351D7F84B4E160E9C0D7C2F09F29">WII-ADR 1681</specimenCode>
(Ezengo),
<emphasis id="1735B8C9054955D98F61025001FF9814" bold="true">C</emphasis>
WII-ADR 1199 (Jengging),
<specimenCode id="5B8509CDC35089E98100FB4C0569A3F0">WII-ADR 1199</specimenCode>
(Jengging),
<emphasis id="189317A6F9AA7CE8373D7D7E385DCF63" bold="true">D</emphasis>
WII-ADR 1218 (Balek),
<specimenCode id="AFE338AB78CBB1F703418A459FACA562">WII-ADR 1218</specimenCode>
(Balek),
<emphasis id="FA0BC65F5AC74B70D4DFC503D9016530" bold="true">E</emphasis>
WII-ADR 454 (Jengging),
<specimenCode id="BB27AEB3DF51C61EFA0D96C6A2B530D0">WII-ADR 454</specimenCode>
(Jengging),
<emphasis id="27E3EC9C73D1F0D9A00B6F220C179E30" bold="true">F</emphasis>
WII-ADR 1213 (Ramsing),
<specimenCode id="7E591DF33A12C943F3A4FB5602BA92F0">WII-ADR 1213</specimenCode>
(Ramsing),
<emphasis id="12B99593C367D72F5554E0912229323F" bold="true">G</emphasis>
WII-ADR 698 (Potin),
<specimenCode id="1054EBC6048E9270CADB3404C6D7D8AD">WII-ADR 698</specimenCode>
(Potin),
<emphasis id="E253066F32F753AED84417B3A86F6CA7" bold="true">H</emphasis>
WII-ADR 3017 (Glaw lake),
<specimenCode id="3A38369894F4EC49E022614EE194A356">WII-ADR 3017</specimenCode>
(Glaw lake),
<emphasis id="F067B8AE1E672EEFCE3D7F33FFECF438" bold="true">I</emphasis>
precloacal pores in WII-ADR 698, and
precloacal pores in
<specimenCode id="6EA5577839E098C02795356147D5CF5E">WII-ADR 698</specimenCode>
, and
<emphasis id="ED247180329B79ECB8B428552FEDF8C5" bold="true">J</emphasis>
precloacal pores in WII-ADR 478. Scale bar: 10 mm.
precloacal pores in
<specimenCode id="31394E1BC6873B9C26CCE892A7DB3BA0">WII-ADR 478</specimenCode>
. Scale bar: 10 mm.
<emphasis id="810B37CBE3573B27FA9688CB82640A3E" bold="true">I</emphasis>
,
<emphasis id="1BD501DA45BE36B39D9EAC222F25D318" bold="true">J</emphasis>
@ -369,9 +519,7 @@ ranges between 0.16.4 %. Among the two clades identified by
<heading id="99BCA824112410B94FBB1510B8F0CB21" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="DE222B6093D082C936AFA56C8F3EF1C1">
The
<typeStatus id="56046C7452172E48D70B41084D1869C4">type</typeStatus>
locality,
The type locality,
<collectingRegion id="CCF01467387DFDCD2590598B4DA75269" country="China" name="Xizang">Xizang</collectingRegion>
(
<collectingRegion id="1640CA3865AE026CCA9C597449F4E094" country="China" name="Xizang">Tibet</collectingRegion>

View file

@ -1,78 +1,80 @@
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<treatment id="6B05318011F35D7C95E621A314732FC0" ID-DOI="http://doi.org/10.5281/zenodo.13124035" ID-Zenodo-Dep="13124035" ID-arpha="6B053180-11F3-5D7C-95E6-21A314732FC0" ID-zoobank="https://zoobank.org/C45008F3-D9E2-473A-B72A-DE92D00C73E4" LSID="urn:lsid:zoobank.org:act:C45008F3-D9E2-473A-B72A-DE92D00C73E4" LSID-ZBK="urn:lsid:zoobank.org:act:C45008F3-D9E2-473A-B72A-DE92D00C73E4" httpUri="http://treatment.plazi.org/id/6B05318011F35D7C95E621A314732FC0">
<subSubSection id="09ED2136318458953535773DE83E0C40" type="nomenclature">
@ -99,9 +101,12 @@ Boruah, Narayanan, Deepak &amp; Das
</heading>
</paragraph>
<paragraph id="97399F86377217FB2F8E8F0D7AF8CDB3">
<materialsCitation id="7E7FE9731E7B89E3D7AAD6FBB3F8D56A" collectingDate="2022-05-18" country="India" county="Bitupan Boruah" elevation="650" latitude="27.4595" location="Kamala Valley" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
Adult male (WII-ADR 1416), collected from
<location id="016069FAB6E452DB5B75AF67F673C652" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:016069FAB6E452DB5B75AF67F673C652" country="India" county="Bitupan Boruah" latitude="27.4595" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" name="Kamala Valley" stateProvince="Arunachal Pradesh">Kamala Valley</location>
<materialsCitation id="7E7FE9731E7B89E3D7AAD6FBB3F8D56A" collectingDate="2022-05-18" collectorName="Abhijit Das &amp; Bitupan Boruah" country="India" county="Changlang District" elevation="650" latitude="27.4595" location="Kamala Valley" longLatPrecision="6" longitude="96.4279" municipality="Namdapha Tiger Reserve" specimenCode="WII-ADR 1416" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<specimenCount id="044D8C2B34593DD28CE0524733E7206B" count="1" type="male">Adult male</specimenCount>
(
<specimenCode id="F25FD55028BBD79D852BDA9261806E6A">WII-ADR 1416</specimenCode>
), collected from
<location id="016069FAB6E452DB5B75AF67F673C652" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:016069FAB6E452DB5B75AF67F673C652" country="India" county="Changlang District" latitude="27.4595" longLatPrecision="6" longitude="96.4279" municipality="Namdapha Tiger Reserve" name="Kamala Valley" stateProvince="Arunachal Pradesh">Kamala Valley</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EB6CK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.427900,27.459500]}">
<geoCoordinate id="356961E8EF822186FD330ADC4D1E4829" degrees="27.4595" direction="north" orientation="latitude" precision="5" value="27.4595">27.4595 ° N</geoCoordinate>
@ -115,17 +120,17 @@ elevation
a. s. l.
</elevation>
),
<location id="C885B0EF22DD1794A31BF04675E2F406" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:C885B0EF22DD1794A31BF04675E2F406" country="India" county="Bitupan Boruah" latitude="27.4595" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" name="Namdapha Tiger Reserve" stateProvince="Arunachal Pradesh">Namdapha Tiger Reserve</location>
<collectingMunicipality id="0EAAC732F747F56B82DD6A13C67654C6">Namdapha Tiger Reserve</collectingMunicipality>
,
<location id="E4A3028A8163D85E4C350A92639AAFD3" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:E4A3028A8163D85E4C350A92639AAFD3" country="India" county="Bitupan Boruah" latitude="27.4595" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" name="Changlang District" stateProvince="Arunachal Pradesh">Changlang District</location>
<collectingCounty id="8AAD51C9CD2804072D55AB17656B505F">Changlang District</collectingCounty>
,
<collectingRegion id="20E2ADC38446B988C443265855911E38" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="A6584DC4680CF6BDF07EC84EF1884EA4" name="India">India</collectingCountry>
by
<collectingMunicipality id="77A37FE350048406A86502CF3B73DEEA">Abhijit Das</collectingMunicipality>
<collectorName id="F0A3C1F6D52DD844E3B21C45DCB69ADC">Abhijit Das</collectorName>
and
<collectingCounty id="C85A79F6C94EA4028B5331AF27B836D6">Bitupan Boruah</collectingCounty>
<collectorName id="A873E376021DB3B9850E20CF5A162E56">Bitupan Boruah</collectorName>
on
<date id="C348B582366DE5FFD64907F3ECF29BEC" value="2022-05-18">
<collectingDate id="AFB9BB98FA99EF9B7CAA20A1813CF2DB" value="2022-05-18">18 May 2022</collectingDate>
@ -144,46 +149,98 @@ on
</heading>
</paragraph>
<paragraph id="ABDEAD43EDE1F01394612366B3D4CDDC">
<materialsCitation id="D7D0D1BCAB1F72357281779FD65A885F" collectingDate="2022-05-18" collectorName="Abhijit Das &amp; Bitupan Boruah" country="India" county="Changlang District" elevation="650" latitude="27.4595" location="Kamala Valley" longLatPrecision="6" longitude="96.4279" municipality="Namdapha Tiger Reserve" specimenCode="WII-ADR 1415, WII-ADR 1417" specimenCount="3" specimenCount-adult="2" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="08CA1623AEA3FDD42BC03789E6267BC1" count="2" type="adult">Two adult</specimenCount>
males (WII-ADR 1415, WII-ADR 1417) collected from the same locality as the
<typeStatus id="52FCA7858678FB875BEB952F25F8E1A3" type="holotype">holotype</typeStatus>
on the same date;
<specimenCount id="0EBA2573800C31B0BDE6ED522ED1B6FA" count="1" type="male">males</specimenCount>
(
<specimenCode id="D5FDD4CE4F2BFE5F0BFB46D1E2E6B967">WII-ADR 1415</specimenCode>
,
<specimenCode id="032256FD0E9738A0AD896D9C0BBE06C2">WII-ADR 1417</specimenCode>
) collected from the same locality as the holotype on the same date
</materialsCitation>
;
<materialsCitation id="D4436660B324A2890C6073F9F7EB917F" collectingDate="2022-05-11" collectorName="Abhijit Das &amp; Bitupan Boruah" country="India" county="Changlang District" elevation="400" latitude="27.4942" location="Deban" longLatPrecision="6" longitude="96.3701" municipality="Namdapha Tiger Reserve" specimenCode="WII-ADR 1404" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="1D3E531609EE26BDF27A66241AF4724A" count="1" type="adult">one adult</specimenCount>
female (WII-ADR 1404) collected near Deban (
<specimenCount id="28DE51D46807B6F5AE89A9A7E95892BA" count="1" type="female">female</specimenCount>
(
<specimenCode id="501DD8BB47FACCB2BE986D6E02DA22D0">WII-ADR 1404</specimenCode>
) collected near
<location id="C93A6BA7A217C98C59F54373382F06C3" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:C93A6BA7A217C98C59F54373382F06C3" country="India" county="Changlang District" latitude="27.4942" longLatPrecision="6" longitude="96.3701" municipality="Namdapha Tiger Reserve" name="Deban" stateProvince="Arunachal Pradesh">Deban</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET6CK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.370100,27.494200]}">
<geoCoordinate id="2E6BF5C260830EFE0F4A91D09D1ACFF1" degrees="27.4942" direction="north" orientation="latitude" precision="5" value="27.4942">27.4942 ° N</geoCoordinate>
;
<geoCoordinate id="9F1DAECC4EE6FF4053191F9FB46DDAF6" degrees="96.3701" direction="east" orientation="longitude" precision="5" value="96.3701">96.3701 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="DF9E7E6654A2636A40D7779BFBA416CC" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</quantity>
a. s. l.), Namdapha Tiger Reserve, Changlang District,
<quantity id="DF9E7E6654A2636A40D7779BFBA416CC" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation id="5B7BDF7D2E3292BAA6824CEB6A141F21" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
a. s. l.),
<collectingMunicipality id="A953D023BBAA76E9631A35D63938BDE4">Namdapha Tiger Reserve</collectingMunicipality>
,
<collectingCounty id="11E5D7BA794B3FC1777AC14AD21BD997">Changlang District</collectingCounty>
,
<collectingRegion id="9A92A90AA0264A67C2147FEF7BFDA7B3" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="E511377949896FE8D05F38888D81D3C4" name="India">India</collectingCountry>
by Abhijit Das and Bitupan Boruah on
<date id="A89057C8B6144D5D640508DE0356BA53" value="2022-05-11">11 May 2022</date>
by
<collectorName id="4C0C16FDF1101BEC735274C5D581FC21">Abhijit Das</collectorName>
and
<collectorName id="949B3D00AF8F3DCB832901846C686205">Bitupan Boruah</collectorName>
on
<date id="A89057C8B6144D5D640508DE0356BA53" value="2022-05-11">
<collectingDate id="23D6FD9D1725BCC3921F141CE42999DD" value="2022-05-11">11 May 2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="283E860FBC208D0735DC4A9928CD0A9B" collectingDate="2022-09-11" collectorName="Abhijit Das &amp; Bitupan Boruah" country="India" county="Changlang District" elevation="470" latitude="27.4899" location="Motijheel trail" longLatPrecision="6" longitude="96.3348" municipality="Gibbons' Land" specimenCode="WII-ADR 3067, WII-ADR 3068" specimenCount="3" specimenCount-adult="2" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="141D74BF4D005A226356967F18E82507" count="2" type="adult">two adult</specimenCount>
females (WII-ADR 3067, WII-ADR 3068) collected from Motijheel trail (
<specimenCount id="0292117B6AC23897D7E3B4E38F1AD787" count="1" type="female">females</specimenCount>
(
<specimenCode id="5E2627125075ECAA0C8ECF424C0CD9EF">WII-ADR 3067</specimenCode>
,
<specimenCode id="E9859E397421DBC6E629F02468D745C6">WII-ADR 3068</specimenCode>
) collected from
<location id="079C966D0C6C164E06E8EDBDC679FD5E" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:079C966D0C6C164E06E8EDBDC679FD5E" country="India" county="Changlang District" latitude="27.4899" longLatPrecision="6" longitude="96.3348" municipality="Gibbons' Land" name="Motijheel trail" stateProvince="Arunachal Pradesh">Motijheel trail</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E26CK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.334800,27.489900]}">
<geoCoordinate id="981C8340D3E57524E03BAC6093CC3627" degrees="27.4899" direction="north" orientation="latitude" precision="5" value="27.4899">27.4899 ° N</geoCoordinate>
,
<geoCoordinate id="D80A7B12AE4559C5427AB52A772B3C8A" degrees="96.3348" direction="east" orientation="longitude" precision="5" value="96.3348">96.3348 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="EDF933E331F4678E124DE2BDA270F9A5" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</quantity>
a. s. l.), Gibbons Land, Namdapha Tiger Reserve, Changlang District,
<quantity id="EDF933E331F4678E124DE2BDA270F9A5" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">
<elevation id="CC1AB4193290307206719177D068FD05" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</elevation>
</quantity>
a. s. l.),
<collectingMunicipality id="826661803AA421341A14AB2B347155CB">Gibbons Land</collectingMunicipality>
,
<collectingMunicipality id="53DF41972EC6D5C4D1FEA2ACF4565328">Namdapha Tiger Reserve</collectingMunicipality>
,
<collectingCounty id="C5CCC0D974D985FD97087B6E04623EA6">Changlang District</collectingCounty>
,
<collectingRegion id="218B4A141E95789A7C4783F22F7D5267" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="ED607637F6B0E11B13AA78BA5DDC488C" name="India">India</collectingCountry>
by Abhijit Das and Bitupan Boruah on
<date id="9318970E16A177EBCF0961655A62ADB6" value="2022-09-11">11 September 2022</date>
by
<collectorName id="801C4FDF84851BDCBDFFDE2721002A59">Abhijit Das</collectorName>
and
<collectorName id="A4DCFA02164BCD1FA07483730BF6CDAB">Bitupan Boruah</collectorName>
on
<date id="9318970E16A177EBCF0961655A62ADB6" value="2022-09-11">
<collectingDate id="D9F86D7599A38FAC3F8F42D42144368A" value="2022-09-11">11 September 2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="EAA43C07F1C7533CD6DEE11D44733F82" collectingDate="2022-09-18" collectorName="Abhijit Das &amp; Bitupan Boruah" country="India" county="Changlang District" elevation="480" latitude="27.4878" location="Burma Nullah" longLatPrecision="6" longitude="96.5416" municipality="Namdapha Tiger Reserve" specimenCode="WII-ADR 1790" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="D6A07B524509B399A17720C247AE0F4E" count="1" type="adult">one adult</specimenCount>
female (WII-ADR 1790) collected near
<collectingCountry id="4B71C6FED809D0FEFE8763D962639713" name="Myanmar">Burma</collectingCountry>
Nullah (
<specimenCount id="DCFA886689FB859A36023B2DADB1FE9F" count="1" type="female">female</specimenCount>
(
<specimenCode id="84881DD73077F9012375048AEACD237D">WII-ADR 1790</specimenCode>
) collected near
<location id="E2BBDF23726A65251D1D35BE70F0A6F1" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:E2BBDF23726A65251D1D35BE70F0A6F1" country="India" county="Changlang District" latitude="27.4878" longLatPrecision="6" longitude="96.5416" municipality="Namdapha Tiger Reserve" name="Burma Nullah" stateProvince="Arunachal Pradesh">Burma Nullah</location>
(
<quantity id="E32FD9799914E401015B53E5FF09022A" metricMagnitude="4" metricUnit="m" metricValue="6.437376" unit="mi" value="40.0">40 mile</quantity>
point) (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDADK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.541600,27.487800]}">
@ -192,13 +249,26 @@ point) (
<geoCoordinate id="65EFBA1B38799B371864B3C214766B61" degrees="96.5416" direction="east" orientation="longitude" precision="5" value="96.5416">96.5416 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="9D2B9A25B85DA756FB0F933227428D96" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">480 m</quantity>
a. s. l.), Namdapha Tiger Reserve, Changlang District,
<quantity id="9D2B9A25B85DA756FB0F933227428D96" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">
<elevation id="A7E30C003B2619CDE2F3AC0F3A6737FE" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">480 m</elevation>
</quantity>
a. s. l.),
<collectingMunicipality id="8B8086C8C6DD49C451E2792AAABF9B18">Namdapha Tiger Reserve</collectingMunicipality>
,
<collectingCounty id="8DE8B80FE825ED52C1ABDD947AEADDDC">Changlang District</collectingCounty>
,
<collectingRegion id="24F47448037CA7AA5CB483F4A4BB7FCB" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="6DA73970ADDA3FB26FDC461BC844DD19" name="India">India</collectingCountry>
by Abhijit Das and Bitupan Boruah on
<date id="6AD3105EF9C5983158E2FC2F59CAF407" value="2022-09-18">18 September 2022</date>
by
<collectorName id="E908B0B6130E25B366F4998F31F7CA95">Abhijit Das</collectorName>
and
<collectorName id="B3B25E839337987348781E8F42BE67EB">Bitupan Boruah</collectorName>
on
<date id="6AD3105EF9C5983158E2FC2F59CAF407" value="2022-09-18">
<collectingDate id="C544CA2D3CE0B186F10388AA45A681B2" value="2022-09-18">18 September 2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -207,40 +277,80 @@ by Abhijit Das and Bitupan Boruah on
<heading id="D1ED0B7FAA7E3C93B8AA6226622FCADB" reason="title">Referred materials.</heading>
</paragraph>
<paragraph id="5F3DB67A0F51D5FF026FD5826CE54144">
An adult female (WII-ADR 3060) collected from Sinabrai (
<materialsCitation id="4786D1F32DCA9FCFFDD930B4FADC274D" collectingDate="2022-09-05" collectorName="Bitupan Boruah &amp; Abhijit Das" country="India" county="Lohit District" elevation="470" latitude="27.7434" location="Sinabrai" longLatPrecision="6" longitude="96.3872" municipality="Kamlang Tiger Reserve" specimenCode="WII-ADR 3060" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
An
<specimenCount id="ED983900C1996ABA185B3867D19359BB" count="1" type="female">adult female</specimenCount>
(
<specimenCode id="615859B4004CF42773B02F2939BBA648">WII-ADR 3060</specimenCode>
) collected from
<location id="584D40418BDEB422F11D73F403EF26AF" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:584D40418BDEB422F11D73F403EF26AF" country="India" county="Lohit District" latitude="27.7434" longLatPrecision="6" longitude="96.3872" municipality="Kamlang Tiger Reserve" name="Sinabrai" stateProvince="Arunachal Pradesh">Sinabrai</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERADK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.387200,27.743400]}">
<geoCoordinate id="568417FD29E6E62FF1C96DBF361BBF61" degrees="27.7434" direction="north" orientation="latitude" precision="5" value="27.7434">27.7434 ° N</geoCoordinate>
;
<geoCoordinate id="454F33EF3373862354F0583E73FC6D81" degrees="96.3872" direction="east" orientation="longitude" precision="5" value="96.3872">96.3872 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="792D1D7B387058FF20F7BD645771916D" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</quantity>
a. s. l.), Kamlang Tiger Reserve, Lohit District,
<quantity id="792D1D7B387058FF20F7BD645771916D" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">
<elevation id="A900E416F2FD53CDA755769A011EED55" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</elevation>
</quantity>
a. s. l.),
<collectingMunicipality id="741B4D038F90CD19D30F3FA13379380E">Kamlang Tiger Reserve</collectingMunicipality>
,
<collectingCounty id="D3B1D941E66F8C1EDA06443DF63D4CD4">Lohit District</collectingCounty>
,
<collectingRegion id="2F9F785362B33C113032AFBEEA2C2FDA" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="3A4C4FBB6A39EC1615B02F1652B217B8" name="India">India</collectingCountry>
on
<date id="5AF38BDD10C50A76F60DA449B6A79B3A" value="2022-09-05">5 September 2022</date>
by Bitupan Boruah and Abhijit Das; an adult female (WII-ADR 3281) and an adult male (WII-ADR 3282) collected from Hornbill (
<date id="5AF38BDD10C50A76F60DA449B6A79B3A" value="2022-09-05">
<collectingDate id="33C2D6B51D5AF48B662F6FB22B7B9DDC" value="2022-09-05">5 September 2022</collectingDate>
</date>
by
<collectorName id="E9F941B4F9F352DBFA71D9CFB6CC7184">Bitupan Boruah</collectorName>
and
<collectorName id="20B0CC81A3EB6573160FE63934CAC40C">Abhijit Das</collectorName>
</materialsCitation>
;
<materialsCitation id="AAE281F1E91842EFB4C7A7FB8C13F626" collectingDate="2023-05-10" collectorName="Rajiv N. V." country="India" county="Changlang District" elevation="670" latitude="27.5381" location="Hornbill" longLatPrecision="6" longitude="96.4403" municipality="Namdapha Tiger Reserve" specimenCode="WII-ADR 3281, WII-ADR 3282" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
an
<specimenCount id="A1A0C4AAD0C30A0D4EE5B1C76B0DE303" count="1" type="female">adult female</specimenCount>
(
<specimenCode id="0ACA94659909E4DD0A0F7047B3C0E424">WII-ADR 3281</specimenCode>
) and an
<specimenCount id="11890371F7B5BC7EAFEECE688DE23230" count="1" type="male">adult male</specimenCount>
(
<specimenCode id="5EE3607352A8AD603C7613ACCAB38D72">WII-ADR 3282</specimenCode>
) collected from
<location id="FBF9B7DE8F53FD270544B079E6033767" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:FBF9B7DE8F53FD270544B079E6033767" country="India" county="Changlang District" latitude="27.5381" longLatPrecision="6" longitude="96.4403" municipality="Namdapha Tiger Reserve" name="Hornbill" stateProvince="Arunachal Pradesh">Hornbill</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZADK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.440300,27.538100]}">
<geoCoordinate id="FBCD3653BAA983934A5DDC1F71FC6D78" degrees="27.5381" direction="north" orientation="latitude" precision="5" value="27.5381">27.5381 ° N</geoCoordinate>
;
<geoCoordinate id="A7F213FD7EB9092A0BABEED401953DF2" degrees="96.4403" direction="east" orientation="longitude" precision="5" value="96.4403">96.4403 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="E5181AA3D7199B34DEACB46554A16801" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</quantity>
a. s. l.), Namdapha Tiger Reserve, Changlang District,
<quantity id="E5181AA3D7199B34DEACB46554A16801" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">
<elevation id="C0E89DEB9E57BFC07C7E3656EDA9DB84" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</elevation>
</quantity>
a. s. l.),
<collectingMunicipality id="FF6B3D4461C41E29D265F47E48C82268">Namdapha Tiger Reserve</collectingMunicipality>
,
<collectingCounty id="852EE7ED617BAA527653C4F1D31DCCF4">Changlang District</collectingCounty>
,
<collectingRegion id="CB00BD2DD73E6FC9F6AE1CDEB4930F37" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="2E5ADDDF27246FD866EFDE3A3C06F01C" name="India">India</collectingCountry>
on
<date id="BFFBF276C5EBDD13DE60D0D944EB4E18" value="2023-05-10">10 May 2023</date>
by Rajiv
<taxonomicNameLabel id="841FB27F9F7F3678962C7C0DEE96B224" rank="variety">
N.
<collectionCode id="01534B89436CB073006DC30D33A7124C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.
</taxonomicNameLabel>
<date id="BFFBF276C5EBDD13DE60D0D944EB4E18" value="2023-05-10">
<collectingDate id="E24A3B3882271B8C4E57C013C7394FB2" value="2023-05-10">10 May 2023</collectingDate>
</date>
by
<collectorName id="54AB43EC5E870D4B15BF179E31519FA3">
Rajiv
<taxonomicNameLabel id="841FB27F9F7F3678962C7C0DEE96B224" rank="variety">N. V.</taxonomicNameLabel>
</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5ADK" type="diagnosis">
@ -336,7 +446,8 @@ Tail regenerated (
<paragraph id="77EF1AC4FE58B2DA9D3F7C1A0CE47016">
(Based on
<typeStatus id="B206B4E6C1BFF7FEB954A9482F733A9B" type="paratype">paratype</typeStatus>
WII-ADR 1417) (Fig.
<specimenCode id="14D039D7F540D44737D5A95DE5C8AD99">WII-ADR 1417</specimenCode>
) (Fig.
<figureCitation id="C58C103BD7BF9A93CBA6B7463C767D6F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyrtodactylus namdaphaensis sp. nov. A G holotype (WII-ADR 1416), A dorsal, and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F ventral view of left manus and G left pes; H precloacal pores of paratype WII-ADR 1417; I paratype WII-ADR 1417, and J paratype WII-ADR 1415 in life. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure8" httpUri="https://binary.pensoft.net/fig/1102274">8 I</figureCitation>
); head dorsally brown with irregular dark-brown patches, a pair of pale-yellowish spots on loreal in front of anterodorsal corner of eyes, lips slightly paler than dorsal head colour with irregular pale-yellowish spots; a pale-brown postorbital streak; pale-brown irregular spots on dark-brown background on neck; upper eyelids greyish-brown; slightly dark-brown large spots of irregular size and shape on dorsal and lateral side of the trunk, interspaced with pale-brown patches, dark-brown spots on dorsal side of the trunk giving appearance of continuous stripes up to middle of the trunk; anterior two third of dorsal side of the tail with alternative broad dark-brown and narrow pale-brown bands, those dark-brown bands anteriorly diffused and posterior edge zigzagged, posterior one third brown with dark-brown marbling; limbs brown with irregular pale spots and dark-brown reticulation; digits with alternative dark-brown and pale-brown bands.
</paragraph>
@ -354,7 +465,9 @@ A
</emphasis>
<emphasis id="0936969B886F50CE25738E5B2D7CE0FD" bold="true">G</emphasis>
holotype (WII-ADR 1416),
holotype (
<specimenCode id="5E17D4F3E69FCF0FA7D83562A1FF1DF6">WII-ADR 1416</specimenCode>
),
<emphasis id="896CBAEE5C9BC5B40EE16E68E5ACE118" bold="true">A</emphasis>
dorsal, and
<emphasis id="1FACD4F7828FAC1093812041669319F7" bold="true">B</emphasis>
@ -370,11 +483,17 @@ ventral view of left manus and
<emphasis id="1A782C7EEB92A4791E5525853B6C0101" bold="true">G</emphasis>
left pes;
<emphasis id="34AF92A7CF93E051E307F19A5530FE6A" bold="true">H</emphasis>
precloacal pores of paratype WII-ADR 1417;
precloacal pores of paratype
<specimenCode id="6FBB5C6E80E58F58BEED11D8951F07B2">WII-ADR 1417</specimenCode>
;
<emphasis id="3B204674B7882FFD73BD7E1E29D73CFD" bold="true">I</emphasis>
paratype WII-ADR 1417, and
paratype
<specimenCode id="67640943E6CCFAC4B534C801E1D2AD22">WII-ADR 1417</specimenCode>
, and
<emphasis id="7A19A74013128DA7309F9AF54EC69E39" bold="true">J</emphasis>
paratype WII-ADR 1415 in life. Scale bar: 10 mm.
paratype
<specimenCode id="43E1255828891CDC36CE87527A3FF92B">WII-ADR 1415</specimenCode>
in life. Scale bar: 10 mm.
<emphasis id="B22C2436E46EB813A0302A4F51932537" bold="true">I, J</emphasis>
not to scale.
</paragraph>
@ -384,7 +503,23 @@ not to scale.
<paragraph id="2A7912C654749788ECEFA31FCADE3B5B">
<heading id="05F06E40064E4FE6B3F5B03FE24B401E" reason="title">Morphological variation.</heading>
</paragraph>
<paragraph id="1A9B401CE4780394F8A4D4E915D57F5E">Morphological variations are given in Table S 2. Except for those, the dorsal marking pattern varied among the collected specimens. The dorsal spots of WII-ADR 1417 are in the form of two continuous stripes starting from neck, posteriorly broken; a spot at the middle of occipital region followed by a dark cross bar on the nape; WII-ADR 3067 and WII-ADR 3281 has four dorsal stripes starting from neck to the level of hind limb insertion; WII-ADR 3068 has four stripes on neck, followed by five irregular and zigzag cross bars on dorsum; in WII-ADR 1790, dorsal spots are indistinct and irregular; WII-ADR 1415 has three pairs of dark spots posteriorly bordered with white spots on neck, cross bars on dorsum narrower, irregular and posteriorly with white (Fig. S 2); WII-ADR 3060 has six dark brown cross bands on back between the level of axilla and groin, consists of irregular shaped enlarged individual spots, mid dorsally these spots are remarkably disjunct,. Precloacal pores in females are smaller than that of males.</paragraph>
<paragraph id="1A9B401CE4780394F8A4D4E915D57F5E">
Morphological variations are given in Table S 2. Except for those, the dorsal marking pattern varied among the collected specimens. The dorsal spots of
<specimenCode id="380CE1794B8D7A637B8745968326A174">WII-ADR 1417</specimenCode>
are in the form of two continuous stripes starting from neck, posteriorly broken; a spot at the middle of occipital region followed by a dark cross bar on the nape;
<specimenCode id="698B4C3EBA1FF986463CE656EE4D629C">WII-ADR 3067</specimenCode>
and
<specimenCode id="ACDB05166D0C28719D3DAF3B5CAA426D">WII-ADR 3281</specimenCode>
has four dorsal stripes starting from neck to the level of hind limb insertion;
<specimenCode id="8AAC81715546B52FEBB808A3E3ED822C">WII-ADR 3068</specimenCode>
has four stripes on neck, followed by five irregular and zigzag cross bars on dorsum; in
<specimenCode id="E02C42C0B23770569F262D02B1278643">WII-ADR 1790</specimenCode>
, dorsal spots are indistinct and irregular;
<specimenCode id="C302A0587F927AACA1C95F369D737E20">WII-ADR 1415</specimenCode>
has three pairs of dark spots posteriorly bordered with white spots on neck, cross bars on dorsum narrower, irregular and posteriorly with white (Fig. S 2);
<specimenCode id="DA1123315AA023BBA373917D919EC0D7">WII-ADR 3060</specimenCode>
has six dark brown cross bands on back between the level of axilla and groin, consists of irregular shaped enlarged individual spots, mid dorsally these spots are remarkably disjunct,. Precloacal pores in females are smaller than that of males.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKGDK" type="comparisons">
<paragraph id="036CE7FD74697C82EEE7B6018792E23D">
@ -450,9 +585,7 @@ ranges between 0.15 %.
<heading id="17E1DAFFB68F9BBB0F73FE8C719750CF" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="396F0A99E917C96A131E6A8ADD72AB84">
The specific epithet is a toponym named after its
<typeStatus id="812E0499C4A546FA8F1D77B4ABB10F1A">type</typeStatus>
locality Namdapha Tiger Reserve in
The specific epithet is a toponym named after its type locality Namdapha Tiger Reserve in
<collectingRegion id="C44C9587FFDE5F569F840067BC24077D" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="9163649EFEF2A7B9721E9181E4D3180E" name="India">India</collectingCountry>
@ -485,9 +618,7 @@ a. s. l. inside Namdapha Tiger Reserve. All the localities are south of Noa-Dihi
<collectingCountry id="B5BDE214604BB32968EA456924D21C69" name="India">India</collectingCountry>
. We recorded this species between
<date id="4D1605E801EED748375053DB395509AF" value="2022-05" valueMax="2022-09-31" valueMin="2022-05-00">May and September 2022</date>
. Individuals were recorded on tree bark, ferns, and riparian vegetation and along forest trails between 18: 0023: 00 hrs. The forest
<typeStatus id="D522B6BA5D96E5798ED299C2E45338FF">type</typeStatus>
can be classified as
. Individuals were recorded on tree bark, ferns, and riparian vegetation and along forest trails between 18: 0023: 00 hrs. The forest type can be classified as
<collectingRegion id="DE458EA2BB1FAC62E08F0A00AA53AD3E" country="India" name="Assam">Assam</collectingRegion>
Valley Tropical Evergreen Forest. The area had a distinct understory with a thick covering of leaf-litter. Currently the species has been recorded from 25 Mile,
<collectingCountry id="B8BF08A4A8C9F94AE33622E3E940024C" name="Myanmar">Burma</collectingCountry>

View file

@ -1,78 +1,80 @@
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<mods:namePart id="21B3720DD0FC794FE7C85BC397AC4DB9">Boruah, Bitupan</mods:namePart>
<mods:nameIdentifier id="7575E73F8B76B445FB672D0C81A4CF91" type="ORCID">0000-0001-8829-6069</mods:nameIdentifier>
<mods:affiliation id="7DEC2CC8557075C7B6D1CBFB0F36D8B2">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
<mods:namePart id="7BA8A66493041B5D29B67C5FBE795418">Boruah, Bitupan</mods:namePart>
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<mods:affiliation id="68BB575D30479CC9464A2AF1B4E7015F">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
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<mods:namePart id="DA94DF9270CE52989F74CF6F031FEF2D">Narayanan, Surya</mods:namePart>
<mods:nameIdentifier id="53918460EE8F7F9DE1729B1B17D8CEBB" type="ORCID">0000-0001-9359-2815</mods:nameIdentifier>
<mods:affiliation id="4C23DBA0A609E6E7C35264C65BF9F320">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
<mods:namePart id="D191892F9FD72D220B9299E6CD22B108">Narayanan, Surya</mods:namePart>
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<mods:affiliation id="BCF2196F635F458686B2CFE5DAA1C7CD">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
</mods:name>
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<mods:namePart id="C5B8DCDE5AFA922C7A7F125E4FC445DE">Aravind, Neelavar Ananthram</mods:namePart>
<mods:nameIdentifier id="234C55E857655E2DDF674BC4B36322A1" type="ORCID">0000-0002-4515-8421</mods:nameIdentifier>
<mods:affiliation id="723457748101333C6A1E2B296ACA0897">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India &amp; Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
<mods:namePart id="110EFF26EEBEAFA2D672FC91EE530371">Aravind, Neelavar Ananthram</mods:namePart>
<mods:nameIdentifier id="9449D45B03E5185251964637F7F6ABEA" type="ORCID">0000-0002-4515-8421</mods:nameIdentifier>
<mods:affiliation id="538DCA5E4542D35DADDEFAB715A4F90D">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India &amp; Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
</mods:name>
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@ -99,8 +101,11 @@ Boruah, Narayanan, Deepak &amp; Das
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Adult male (WII-ADR 1596), collected near
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@ -136,7 +141,9 @@ on
<label id="7525362A0A7DCABCD6DA88E762F1751B">Figure 6.</label>
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Holotype (WII-ADR 1596) of
Holotype (
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@ -554,9 +561,7 @@ bent-toed gecko.
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is currently only recorded from the
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locality. We collected a single individual at 18: 00 hrs on
is currently only recorded from the type locality. We collected a single individual at 18: 00 hrs on
<date id="2A81CCF026A391872CDC8CC0E4C271F2" value="2022-07-25">25 July 2022</date>
near Lamdan Kabui village. It was perched on a shrub at a height of approximately
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View file

@ -0,0 +1,747 @@
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Boruah, Narayanan, Deepak &amp; Das
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</paragraph>
</subSubSection>
<subSubSection id="6942BE4B998A2636DECE47BA9D72AB2B" type="description">
<paragraph id="F0B732943FC6C40387484B4A87E85516">
<figureCitation id="765185CB1654C391B1EA949B063DDE4A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">Figure 4</figureCitation>
;
<tableCitation id="E8FA5885DBC629B868F0C921CE246C55" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOVDI" type="Holotype">
<paragraph id="5C5E2E60A3D53D8A678234BAAE141C4B">
<heading id="7AC59DBAEAC1F75523A46D56DAE20C4F" reason="title">
<typeStatus id="8C9D6601413CFCED5126D37D30A20D77" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="8532EB0165DCFABDF0E829B71B95A499">
<materialsCitation id="309CC2CE64088676A4C54689D1A2D5B6" collectedFrom="from a trail beside Forest Colony" collectingDate="2021-08-01" collectorName="Abhijit Das &amp; Bitupan Boruah" country="India" elevation="1300" latitude="25.8994" location="Kiphire Forest Division" longLatPrecision="6" longitude="94.7694" municipality="Kiphire District" specimenCode="WII-ADR 964" specimenCount="1" specimenCount-male="1" stateProvince="Nagaland" typeStatus="holotype">
<specimenCount id="7FDCC1ECB27028C7ED207787B20D333C" count="1" type="male">Adult male</specimenCount>
(
<specimenCode id="8AE08E5EB4A82E917B663BDD6B8CC984">WII-ADR 964</specimenCode>
; Fig.
<figureCitation id="D4AD9E555AE2E9F6B9BC46D11E88CACA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 A E</figureCitation>
),
<collectedFrom id="A2915236A16AEB97F87FD75389B88ECA">from a trail beside Forest Colony</collectedFrom>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2VDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[94.769400,25.899400]}">
<geoCoordinate id="D53F826442D4CE265727048678F1249E" degrees="25.8994" direction="north" orientation="latitude" precision="5" value="25.8994">25.8994 ° N</geoCoordinate>
;
<geoCoordinate id="3FA69315812F783952BDE24F08774B36" degrees="94.7694" direction="east" orientation="longitude" precision="5" value="94.7694">94.7694 ° E</geoCoordinate>
</named-content>
;
<elevation id="C30BAA20B829E9565424A94591EA0D05" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
elevation
<quantity id="285C206AEB1F32774894FF6378DE566F" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</quantity>
a. s. l.
</elevation>
; Fig.
<figureCitation id="83E1D4B3224A6E6052FED73706F76B55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
),
<location id="7BE4A1DB51A7B6BEC76D2A5E4816BF3A" LSID="urn:lsid:plazi:treatment:CAD2A307ECEF5751BF30BB86E58CF3CE:7BE4A1DB51A7B6BEC76D2A5E4816BF3A" country="India" latitude="25.8994" longLatPrecision="6" longitude="94.7694" municipality="Kiphire District" name="Kiphire Forest Division" stateProvince="Nagaland">Kiphire Forest Division</location>
,
<collectingMunicipality id="6FF1644153C380C00DD89FB2DE2417B5">Kiphire District</collectingMunicipality>
,
<collectingRegion id="0CF90506FB6E994271D00180E602EC21" country="India" name="Nagaland">Nagaland</collectingRegion>
,
<collectingCountry id="D7BA8ED0DAB73B5D853F6A6DBD7CEA67" name="India">India</collectingCountry>
; collected by
<collectorName id="8D0A51866B9AFE5D570C734228E40C3E">Abhijit Das</collectorName>
and
<collectorName id="4D09BEC4656FAA9780DF114A49AF886C">Bitupan Boruah</collectorName>
on
<date id="7FA9382C7482B5955F21AE5DC92B2296" value="2021-08-01">
<collectingDate id="D69B2CD56C0185A8683EB7398AC2B2ED" value="2021-08-01">1 August 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="6B88B2D1CD1AEF31BACC5C0A360B9A1B" ID-DOI="10.3897/vz.74.e124752.figure4" ID-arpha="DABDFDAF-7299-516F-8A29-87172C0710E2" httpUri="https://binary.pensoft.net/fig/1102270" startId="F4">
<paragraph id="4EDE8C573D85E2BCB7E21132E1F81173">
<label id="415C02CA2C8E0862A800AB0F487483EB">Figure 4.</label>
</paragraph>
<paragraph id="B212032A0A129C90CE0B73CA4883CD75">
<taxonomicName id="946AB9E7A1669C8CE12E05360FCA00B6" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="F6FC7AC1F87EE5287579F45D572D62E0" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="4145365D4FA0F5E65C9EF85D0BF95A39" bold="true">
<taxonomicNameLabel id="B332C00C6F05D543C18A72A83B6E93AE" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
<emphasis id="671A31BD0999EC7F424AE16FE2E91B3C" bold="true">E</emphasis>
holotype (
<specimenCode id="7460332CE31CAB5CDAB4F4CB4F98C5F0">WII-ADR 964</specimenCode>
),
<emphasis id="1CB5CDE07BD878F43321B2CCD09F853B" bold="true">A</emphasis>
dorsal view,
<emphasis id="A3803197A7227AB7247EF372609FB1AF" bold="true">B</emphasis>
ventral view,
<emphasis id="A8EBECF77AF40C420476A6460237DA73" bold="true">C</emphasis>
dorsal view of head,
<emphasis id="AB66D3E331A9D84DD90BC9FC3E68F981" bold="true">D</emphasis>
ventral view of head,
<emphasis id="AE3E2216532EB8F960D138414E1BEC04" bold="true">E</emphasis>
lateral view of head;
<emphasis id="16B6AD7F60FC7765572C415ADD43893C" bold="true">F</emphasis>
,
<emphasis id="C3826622407A71BEDAD4EFE394506CCF" bold="true">G</emphasis>
dorsal and ventral view of paratype (
<specimenCode id="90EA8A02033FFC40A148EDE0914772E1">WII-ADR 963</specimenCode>
);
<emphasis id="32F9310303BC469C3D21569F0EE0F51A" bold="true">I</emphasis>
ventral view of right manus and
<emphasis id="70C5CD71BA68615D01FD2A72C203C099" bold="true">J</emphasis>
right pes (
<specimenCode id="2670623CECCF651E68755E75AADA2ECF">WII-ADR 964</specimenCode>
);
<emphasis id="68B14F9F66EA7573A44695078BBCBCEB" bold="true">K</emphasis>
holotype in life. Scale bar: 10 mm.
<emphasis id="52EFF49FB98D23E8E4770FB97B6B1A1A" bold="true">J</emphasis>
not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EWXDI" type="Paratype">
<paragraph id="7DE0276CFD10B4EB1B1D825594CE4090">
<heading id="04792C8F8E6D98B92B65949F1924013B" reason="title">
<typeStatus id="AEA6440A4C3A6EA73EDD47FCDA895BA8" type="paratype">Paratype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="306FAFF52A4FBC9DC7F7E0C743EF4E91">
An adult male (
<specimenCode id="9F5318D0D3864E19283ABAFBAB32DE36">WII-ADR 963</specimenCode>
; Fig.
<figureCitation id="CD549BE059D39CFFFCE83EAD6C6A8961" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 F, G</figureCitation>
) collection locality details are same as the
<typeStatus id="6DC8E355E7A33B12C3C726A7BEEBCB9B" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAYDI" type="diagnosis">
<paragraph id="B5046B1797471F3C4F979DD9E22984C6">
<heading id="871B49C03CC4D5352A039382828BB9E9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A53C9EEC33C134BEDCBEB5FD282FC60B">
A medium-sized gecko (
<abbrev id="ABBRID0EGYDI" xlink_title="snout to vent length">SVL</abbrev>
up to at least
<quantity id="500D10E7197D9FEF17A1A8CEC75BC096" metricMagnitude="-2" metricUnit="m" metricValue="6.47" unit="mm" value="64.7">64.7 mm</quantity>
); 10 supralabials; 912 infralabials; 16 bluntly conical and feebly keeled dorsal tubercles; 2629 paravertebral tubercles; 35 or 36 midventral scales rows between the weak ventrolateral folds; no precloacal groove; six or seven precloacal pores in continuous series; 1215 total subdigital lamellae beneath digit IV of pes; dark brown irregular cross bars or reticulation on dorsum.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKYDI" type="description">
<paragraph id="47E0AC54D91A76A73931AF49501EAD5D">
<heading id="104516CA9DCB6705CC9281750190CFFA" reason="title">
Description of the
<typeStatus id="174456A277EAC414E5AA1F066905CC26" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="835DCB072BC230DA71F5AD0598E727CA">
<typeStatus id="96F18E6247D0EFCAB306646334D0315B" type="holotype">Holotype</typeStatus>
well preserved except single incision below left axilla ventrolaterally. Snout-vent length
<quantity id="7D77C098FFC4371856D1E2A8B910AFDC" metricMagnitude="-2" metricUnit="m" metricValue="6.39" unit="mm" value="63.9">63.9 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0EQYDI" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0EUYDI" xlink_title="snout to vent length">SVL</abbrev>
= 0.26), dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0EYYDI" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0E3YDI" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.68), distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (
<abbrev id="ABBRID0EAZDI" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0EEZDI" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.4), longer than orbit (
<abbrev id="ABBRID0EIZDI" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0EMZDI" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.7); nostril nearly rounded, opening directed posterolaterally; ear opening oval and oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous, supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at approximately anterior one third of it; rostral wide, a short groove at the middle on top; rostral connected with nasals, supranasals, internasal and first supralabials; two scale between the supranasals, larger than the rest of the granular snout scales; granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles starting from the level of posterior margin of the upper eyelids, size increases towards nape; supralabial 10 on both side, seven supralabials upto midorbit, size decreases towards angle of jaw; a series of narrow, enlarged scales above the supralabials between nostril and anterior orbital border; mental slightly wider than rostral (RW /
<collectionCode id="BF1CFBF3A1A6CD372D6F0E62C544DA45" country="Netherlands" name="Museum Wasmann">MW</collectionCode>
= 0.8), triangular, connected with first infralabials, inner postmentals; nine infralabials on both sides, size decreases towards angle of jaw; first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals; posterior margin of the inner postmentals bordered by eight granular scales of different size; two rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size decreases, elongated and narrow; rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards the throat where they become imbricate.
</paragraph>
<paragraph id="46012B2CE1FD77C5C3AC9A40F9F3FCA1">
Habitus slender (
<abbrev id="ABBRID0ESZDI" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0EWZDI" xlink_title="snout to vent length">SVL</abbrev>
= 0.2,
<abbrev id="ABBRID0E1ZDI" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0E5ZDI" xlink_title="snout to vent length">SVL</abbrev>
= 0.35), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, up to fourth segment of the tail, size increases towards posterior body and pronounced; 16 dorsal tubercles across mid dorsum; 26 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest on belly; 36 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “
<collectionCode id="24B372FDE76E5802BE796622998EC79C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” shaped continuous series, followed by five unpored, large scales below it, largest at the apex.
</paragraph>
<paragraph id="0AA79FABF5A18751D2E20B81342C4F2B">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EE1DI" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0EI1DI" xlink_title="snout to vent length">SVL</abbrev>
= 0.13,
<abbrev id="ABBRID0EM1DI" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0EQ1DI" xlink_title="snout to vent length">SVL</abbrev>
= 0.16); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 5 + 8 and 5 + 9 respectively; lamellae beneath digit IV of right and left pes (given as basal + distal) is 5 + 10 and 5 + 9 respectively; dorsal scales on forelimbs smooth and subimbricate, small and granular at elbow and proximal end of forearm; forearm scales intermixed with rounded large tubercles; dorsal scales of hindlimbs heterogeneous, intermixed with large, rounded and bluntly conical tubercles; horizontally upper half of the thigh scales are smooth, large and subimbricate, those on lower half small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; scales on ventral side of hindlimbs smaller than those of belly, smooth, cycloid and subimbricate, but on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="0184FF468E2BDAD1150D5C34DEBD0929">
Tail regenerated (
<abbrev id="ABBRID0EW1DI" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="9C39D378FB5CE3C4864D5893AB95227D" metricMagnitude="-2" metricUnit="m" metricValue="6.8" unit="mm" value="68.0">68 mm</quantity>
), slender, gradually tapering towards tip, segments indistinct, dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; large feebly keeled scales upto fourth segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal and widest at the middle, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; two and three bluntly conical spurs on right and left side of the tail base respectively.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E11DI" type="Colouration in life">
<paragraph id="033F0219D647095F574FAF3618A441C5">
<heading id="D5F8E201F8827A4CDE8C351CAB18D48D" reason="title">Colouration in life.</heading>
</paragraph>
<paragraph id="0F044384C055053ED34F66A6F3F69E90">
Top of head intermixed with pale-yellow and pale-brown; dark-brown spots of irregular shape and size on posterior part of head; short dark-brown postorbital streak present; another streak along loreal region; supraciliary yellow; four dark-brown stripes on neck interspaced with cream colour; large spot at the middle on the anterior part of the nape; six dark-brown cross bars of irregular size and shape on dorsum between the level of axilla and groin (Fig.
<figureCitation id="5FF7F36F0F7AE4E2708F89FFE2AAF731" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 J</figureCitation>
); these cross bars intersected on mid dorsum and posteriorly bordered with cream coloured patches; limbs with slightly dark-brown indistinct reticulations intermixed with pale-yellow or cream coloured patches; scales on axillary region and base of forelimbs are with pinkish tinge; digits with alternating brown and pale-yellow bands; anterior half of the tail with alternating dark-brown and light-cream coloured cross bands of irregular shape and size; posterior half (regenerated part) of the tail pale-brown; ventrally head, trunk and limbs whitish; tail with brown mottling.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE2DI" type="Colouration in preservative">
<paragraph id="B968412DB2EE1679C20BCDA197A17C2A">
<heading id="AF9864EFCFE56F2EA8140CFEF8CF6CE5" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="DA76161E7E1AF9F9CA1E033A0679C168">Top of head, back, limbs and tail pale-greyish-brown with dark-brown markings; upper eyelids grey; dorsal marking pattern as it was in life; and ventrally whitish.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ2DI" type="Morphological variation">
<paragraph id="4E56385D83D494FD5966B6B8C2A1736D">
<heading id="1F20E129B20877D4A63BB3AB015259C0" reason="title">Morphological variation.</heading>
</paragraph>
<paragraph id="983820DF28DC4CCAC87E389ADA0B9A89">
Detailed morphometric variations are given in Table S 2. Apart from those, the dorsal colour and marking pattern of the
<typeStatus id="21717126980873B2BB15A1E688889F63" type="paratype">paratype</typeStatus>
varied from that of
<typeStatus id="D106DB6D86377832CF441870A8E2FC60" type="holotype">holotype</typeStatus>
.
<typeStatus id="1CD925445846CC5D0DDFC3DCE0EACA18" type="paratype">Paratype</typeStatus>
has two post orbital streaks on each side, upper one shorter than the lower one; no other spots on occipital region as in the
<typeStatus id="1BEF2C625ADD472960E2B21F4048190E" type="holotype">holotype</typeStatus>
; dorsal cross bands somewhat reticulated (Fig.
<figureCitation id="49198000418951E600801ED5E07F5DEC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cyrtodactylus kiphire sp. nov. A E holotype (WII-ADR 964), A dorsal view, B ventral view, C dorsal view of head, D ventral view of head, E lateral view of head; F, G dorsal and ventral view of paratype (WII-ADR 963); I ventral view of right manus and J right pes (WII-ADR 964); K holotype in life. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure4" httpUri="https://binary.pensoft.net/fig/1102270">4 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ET2DI" type="comparisons">
<paragraph id="D9661BFF7CD371A9469A1CD9D3A5C963">
<heading id="F2DC8F5D7E019F164A3A9694B83BB861" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="D43B67C1DB59215E0996E627D818DCA1">
<taxonomicName id="A9ED2B082567CBD60D77A782D87D5AC7" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="73340120157914E56D4AE01EC5F84F13" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="9E897CE7108631970E02A2D72AFD611C" bold="true">
<taxonomicNameLabel id="65F0A5842EE375D39362B20BCFE0319C" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="42D36C484B0CBCAA593CB40F262E40F8" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aaronbaueri">
<emphasis id="830D9F2D057CFD62209ED90F92EFD5A0" italics="true">C. aaronbaueri</emphasis>
</taxonomicName>
by having larger body size in male,
<abbrev id="ABBRID0ER3DI" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="68385ED4FD2BC53CF38270D9CD5ECDC2" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0EV3DI" xlink_title="snout to vent length">
<collectionCode id="4F4368FA770BE34E4508298FEB2C8ED2">SVL</collectionCode>
</abbrev>
<quantity id="35781095592196AD824B57C0EC1875C5" metricMagnitude="-2" metricUnit="m" metricValue="5.81" metricValueMax="6.2" metricValueMin="5.42" unit="mm" value="58.1" valueMax="62.0" valueMin="54.2">54.262 mm</quantity>
in male), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EZ3DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0E43DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="429F5E63A38F288D86E5E154FF3B72C1">DTR</collectionCode>
</abbrev>
2228), fewer paravertebral tubercles, 2629 (vs.
<abbrev id="ABBRID0EB4DI" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="8D7AC1565ED447B0EF663050DD53F875">PVT</collectionCode>
</abbrev>
3639); differs from
<taxonomicName id="B6F5E5DC95AAFBE6586348E96689BD06" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aunglini">
<emphasis id="C21E0954E42CCD636B3579F23D142CF2" italics="true">C. aunglini</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EQ4DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EU4DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="37F6966BD7E55C8BE41E117FBC4C7615">DTR</collectionCode>
</abbrev>
2126), fewer precloacal pores,
<abbrev id="ABBRID0EY4DI" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0E34DI" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
1213), fewer mid-ventral scale rows,
<abbrev id="ABBRID0EA5DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="E6F66EEFAD126F98FB4D4D47B7C162FB">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EE5DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="D04C97438098684BD09BD443DC338797">MVSR</collectionCode>
</abbrev>
4149), fewer paravertebral tubercles,
<abbrev id="ABBRID0EI5DI" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3645); differs from
<taxonomicName id="548E4F02D9EA3BA377B8557BBA84EC70" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="bengkhuaiai">
<emphasis id="B679918FE006553C6AC0D6837513A3F9" italics="true">C. bengkhuaiai</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EX5DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0E25DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="024797FCC8F95CBBEE18FEB71894BB65">DTR</collectionCode>
</abbrev>
2226), fewer paravertebral tubercles, 2629 (vs.
<abbrev id="ABBRID0E65DI" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="7BF268A492949AE148969AB3EF45FE6E">PVT</collectionCode>
</abbrev>
3541), fewer mid-ventral scale rows,
<abbrev id="ABBRID0ED6DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="B6EDCA3DD9116075BCF70684E50F1219">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EH6DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="998672D351E6EDF4CB3201E4CF179F0D">MVSR</collectionCode>
</abbrev>
3742); differs from
<taxonomicName id="68E2350FA4A0FA9722BE55F84412793A" authorityName="Bauer" authorityYear="2002" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="brevidactylus">
<emphasis id="C64723241CF178ACC3950FA813524218" italics="true">C. brevidactylus</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EW6DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0E16DI" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="5AC732E91A16E1FDC1EFF613705CF15B">DTR</collectionCode>
</abbrev>
27), fewer mid-ventral scale rows,
<abbrev id="ABBRID0E56DI" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="75C7B710CC09C77EBB8CFAD473892B4F">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EDAAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="EB5E2154CDDF917C5E0B5587BDF04FE9">MVSR</collectionCode>
</abbrev>
45), fewer precloacal pores,
<abbrev id="ABBRID0EHAAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0ELAAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
8); differs from
<taxonomicName id="9161941C4496C993F85F69533F023CF2" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chrysopylos">
<emphasis id="DA38365F3D44E564CF9E5A49CBE7B5C5" italics="true">C. chrysopylos</emphasis>
</taxonomicName>
by fewer precloacal pores,
<abbrev id="ABBRID0E1AAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0E5AAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
813), mid ventral scales,
<abbrev id="ABBRID0ECBAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="5501FDC09ED8C70A65F67F64F0E6384F">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EGBAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="79DA4C6F441E83D04D8F774749596D4A">MVSR</collectionCode>
</abbrev>
3755), dorsal tubercle rows,
<abbrev id="ABBRID0EKBAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
2629 (vs.
<abbrev id="ABBRID0EOBAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="5000F93A3AECB76A67A961BF16F4C09C">DTR</collectionCode>
</abbrev>
3035); differs from
<taxonomicName id="92404894FB68DC8A56155096D86F49DF" authorityName="Liu &amp; Rao" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="dianxiensis">
<emphasis id="93E6B41D6E5D0E74B08AAEABE13F0CC5" italics="true">C. dianxiensis</emphasis>
</taxonomicName>
by smaller body size,
<abbrev id="ABBRID0E4BAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="B1DE3820F1B9218CED1AB0D0C0BE0553" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0EBCAK" xlink_title="snout to vent length">
<collectionCode id="F7E51A5ABFCF905FF72B2FC0E70E43EB">SVL</collectionCode>
</abbrev>
<quantity id="1038C09EB9A5B2B0DA22DEC6CA3F1327" metricMagnitude="-2" metricUnit="m" metricValue="7.685" metricValueMax="7.99" metricValueMin="7.38" unit="mm" value="76.85" valueMax="79.9" valueMin="73.8">73.879.9 mm</quantity>
), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EFCAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EJCAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="3C08742B58B11269EEAC323C07CAD33A">DTR</collectionCode>
</abbrev>
18 or 19), 35 or 36 mid ventral scales (vs.
<abbrev id="ABBRID0ENCAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="214E497013A09441C2E21114ADD15CDA">MVSR</collectionCode>
</abbrev>
3741), 26 or 29 paravertebral tubercles (vs.
<abbrev id="ABBRID0ERCAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="173C97DAAD89C90C552B40CFCC05693A">PVT</collectionCode>
</abbrev>
31 or 32); differs from
<taxonomicName id="1873EFE2845B0FAAC9CE9DC05DD1F406" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="gansi">
<emphasis id="0844A91513E4A4C4B0E28AD3678654AA" italics="true">C. gansi</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EADAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EEDAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="D7830C7F101D3A262172DC38B303F931">DTR</collectionCode>
</abbrev>
2025), having fewer precloacal pores,
<abbrev id="ABBRID0EIDAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EMDAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
1629); differs from
<taxonomicName id="F4ABB2165D18AA56A1EFA95CB3FF99ED" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="jaintiaensis">
<emphasis id="C8EB78F97D42605832E655F534B4FEF2" italics="true">C. jaintiaensis</emphasis>
</taxonomicName>
by smaller body size in male,
<abbrev id="ABBRID0E2DAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="CA9CD25A3ABB4E0E3B6E1EB85F7BC56E" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0E6DAK" xlink_title="snout to vent length">
<collectionCode id="B64090FBE7C39B7D0F957878EB8CB6DA">SVL</collectionCode>
</abbrev>
<quantity id="F6E830D78CC87640E94D37ABD2BF76FC" metricMagnitude="-2" metricUnit="m" metricValue="8.765" metricValueMax="8.83" metricValueMin="8.7" unit="mm" value="87.65" valueMax="88.3" valueMin="87.0">8788.3 mm</quantity>
), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EDEAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EHEAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="B3C129AB4F1CDAA5A26EF1B7B9A41CE7">DTR</collectionCode>
</abbrev>
1920), fewer mid-ventral scale rows,
<abbrev id="ABBRID0ELEAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="5D85E15E4FA531D3CA87631BC9842E10">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0EPEAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="4A69F92A686DD119E4343E3F366A4048">MVSR</collectionCode>
</abbrev>
4042), fewer precloacal pores,
<abbrev id="ABBRID0ETEAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EXEAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
1112); differs from
<taxonomicName id="94D182FF5864FC62F56BC3C73A72459F" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="lungleiensis">
<emphasis id="6D846123DC1B7EC197E5F1D630E5697F" italics="true">C. lungleiensis</emphasis>
</taxonomicName>
by smaller body size in male,
<abbrev id="ABBRID0EGFAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="02601F2866564E0EA3B313C09BA19325" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0EKFAK" xlink_title="snout to vent length">
<collectionCode id="3D94CD6B8D4B99EF8610ED7EBAE546B4">SVL</collectionCode>
</abbrev>
<quantity id="7421E4649754BF1A7C650DA8C8295817" metricMagnitude="-2" metricUnit="m" metricValue="6.654999999999999" metricValueMax="6.81" metricValueMin="6.5" unit="mm" value="66.55" valueMax="68.1" valueMin="65.0">6568.1 mm</quantity>
), higher precloacal pores,
<abbrev id="ABBRID0EOFAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0ESFAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
35), fewer mid-ventral scale rows,
<abbrev id="ABBRID0EWFAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="CF0E0358701FA3E2CC99CBFEE5E45D10">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0E1FAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="C71E1D5B14DF1187A21FE42FD10D7798">MVSR</collectionCode>
</abbrev>
3743), fewer dorsal tubercle rows,
<abbrev id="ABBRID0E5FAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs. 2428), fewer paravertebral scales,
<abbrev id="ABBRID0ECGAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3240), fewer subdigital lamellae beneath toe IV,
<abbrev id="ABBRID0EGGAK" xlink_title="count for the apical series, comprising lamellae distal to the digital inflection but not including the ventral claw sheath and the small non lamellar scales between the basal and apical lamellae series, abbreviations for lamellae are as follows: on the pes">TIVLam</abbrev>
1214 (vs.
<abbrev id="ABBRID0ELGAK" xlink_title="count for the apical series, comprising lamellae distal to the digital inflection but not including the ventral claw sheath and the small non lamellar scales between the basal and apical lamellae series, abbreviations for lamellae are as follows: on the pes">TIVLam</abbrev>
1618); differs from
<taxonomicName id="13DA08BF8E9DFD4FD126CAAB68B4C26A" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="montanus">
<emphasis id="265D7E4702428457DE671AB29F227BDE" italics="true">C. montanus</emphasis>
</taxonomicName>
by larger body size in male,
<abbrev id="ABBRID0E2GAK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="3E7383B11425DD0D4A6CBFD1717BD39E" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
(vs.
<abbrev id="ABBRID0E6GAK" xlink_title="snout to vent length">
<collectionCode id="FD8FC39311734741BFB7FE268047B011">SVL</collectionCode>
</abbrev>
<quantity id="EAC8EA2B54C6FDC93D6014FE548969B8" metricMagnitude="-2" metricUnit="m" metricValue="5.43" metricValueMax="5.5" metricValueMin="5.36" unit="mm" value="54.3" valueMax="55.0" valueMin="53.6">53.655 mm</quantity>
), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EDHAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EHHAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="6C6BB8BC88B80862944785D142330047">DTR</collectionCode>
</abbrev>
2123), fewer precloacal pores,
<abbrev id="ABBRID0ELHAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EPHAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
810); differs from
<taxonomicName id="E24F666814C7214B113447567990D854" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="myaleiktaung">
<emphasis id="23124DDA4648AE5273EA915A1B786EC7" italics="true">C. myaleiktaung</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0E5HAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="2557C43D7F6C169208CCA135A7812409">MVSR</collectionCode>
</abbrev>
35 or 36 (vs.
<abbrev id="ABBRID0ECIAK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="873686AF51BF94FECE71DC9964424FAF">MVSR</collectionCode>
</abbrev>
57); differs from
<taxonomicName id="A5F378F3E95D1EB37EEFBC3218809EE7" authorityName="Mahony &amp; Kamei" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namtiram">
<emphasis id="4FADDC8A8F4511001B8DF680DC0DB4A0" italics="true">C. namtiram</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ERIAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0EVIAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="5428C2C95167BE524094031A01233C3D">DTR</collectionCode>
</abbrev>
21), fewer number of paravertebral scales,
<abbrev id="ABBRID0EZIAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs.
<abbrev id="ABBRID0E4IAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="12818F958C3BCC912BA79C9AFE04F8CB">PVT</collectionCode>
</abbrev>
33), fewer precloacal pores,
<abbrev id="ABBRID0EBJAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EFJAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
12); differs from
<taxonomicName id="C0BB5068042CF66443927199D45495A6" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngopensis">
<emphasis id="C48BD89EE472DD2DD14A845C6714207A" italics="true">C. ngopensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EUJAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs. 1920), fewer paravertebral tubercles,
<abbrev id="ABBRID0EYJAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3236); differs from
<taxonomicName id="DC59B8CC6952F90B9C1DDD5E5F830A4A" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siahaensis">
<emphasis id="A21A0AA325BB937D425B2AFE2C4F2510" italics="true">C. siahaensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EHKAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs.
<abbrev id="ABBRID0ELKAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="02B3862A1AFE6AA75D6C45C7CAB563DC">DTR</collectionCode>
</abbrev>
2224), fewer paravertebral tubercles, 2629 (
<abbrev id="ABBRID0EPKAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="B4FD861B8270962FE02ACB9DA2EE90C1">PVT</collectionCode>
</abbrev>
3639); differs from
<taxonomicName id="20285FB448434EEFB1536D8C344EA8B9" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="vairengtensis">
<emphasis id="7A828F534FE7867BA35C5895456ACFCF" italics="true">C. vairengtensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0E5KAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
16 (vs. 2223), fewer paravertebral tubercles,
<abbrev id="ABBRID0ECLAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3439), fewer precloacal pores in male,
<abbrev id="ABBRID0EGLAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6 or 7 (vs.
<abbrev id="ABBRID0EKLAK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
911). The new species is morphologically close to
<taxonomicName id="C009C4A6459C6C7CD534EFFA47E288E2" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="9D2944D6A6B129E15FDABCBD0EAC25F5" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
; however, the new species differs from it by fewer paravertebral tubercles,
<abbrev id="ABBRID0EZLAK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
2629 (vs. 3537). We could not compare with male specimens as data for
<taxonomicName id="7C3115DFAE2094D186DC431946823B54" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="555283909F04F22D8469249D1AF9C3D0" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
is not available. Morphological differences with other members of
<taxonomicName id="E76E555048D38A4BF8FFA52D0EF42141" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="AB826410026D8845148E391049619862" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="65D36B8A1573A4A7E1DF28E6849D6876" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWMAK" type="Sequence divergence">
<paragraph id="00D27784C02B5DBD4640554A5340E1B5">
<heading id="50B96D17333767F0EEBBAB991C145818" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="3DAE78EEC9C60D3588C9C78880764ED5">
<taxonomicName id="0145A8271EEDA2740AE1F61A152A733A" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="8DB3010276D460D35CF3DB37E1429240" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="C225477C71BD1D3DAB2543C3B0B03640" bold="true">
<taxonomicNameLabel id="EF42E0BFCC3A9E7B27A66FC520E16001" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a high genetic divergence of 12 % from its closely related
<taxonomicName id="F4336A89CCC1B7D5EA647EFD868118EB" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="7AEE5DFE91D749032B0F98E46F7678D4" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
, 16.3 % from
<taxonomicName id="946E5542B83C957C7B500704F20CE095" authorityName="Liu &amp; Rao" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="dianxiensis">
<emphasis id="F30A858F5375B637E1469BD48BE821D1" italics="true">C. dianxiensis</emphasis>
</taxonomicName>
, 16.9 % from
<taxonomicName id="3552A44864D633235EAA44B5960E1004" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="gansi">
<emphasis id="2A53D95B121DF67D4A2DB7C9C317ECC2" italics="true">C. gansi</emphasis>
</taxonomicName>
and 17.9 % from
<taxonomicName id="B7B217A50470968FEE9AD27845ADCF29" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="jaintiaensis">
<emphasis id="942A55421EED5AFEB2CCA950C7836404" italics="true">C. jaintiaensis</emphasis>
</taxonomicName>
. With other members of the clade,
<taxonomicName id="843A4541148A462981061EFA886207B5" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="B96878DC57BFB2FF18C14194F8C2C0FD" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="A422B104D0B147FF893F8EDDE3C86D5E" bold="true">
<taxonomicNameLabel id="E02E684DE170BE19661400FE32A79188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a genetic divergence of 16.4 % and 22.5 % in the ND 2 gene (Table S 3).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECPAK" type="etymology">
<paragraph id="6440FBEF1773FC21C532113FF3397EC3">
<heading id="AC90CCA57EDF93F4B04A24DC7B4414D7" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="BBD8AA84E78FC59E9E9EB53AAFA48068">
The specific epithet is derived from Kiphire, a District of
<collectingRegion id="29DB84EBDCAE1128B03F310817038E58" country="India" name="Nagaland">Nagaland</collectingRegion>
from where the type series of this species were collected.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHPAK" type="Suggested common name">
<paragraph id="1C913A888AD0ED31825423228A16D270">
<heading id="DFB9630D18077DBA15AFD3BB46546018" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="59934BC653AC2E24692DC90BE886FC1B">Kiphire bent-toed gecko.</paragraph>
</subSubSection>
<subSubSection id="SECID0EMPAK" type="distribution">
<paragraph id="E7A508F09E1F52E41B9CEAC706C52802">
<heading id="467689291B31F11CF6474D0DB24C9688" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="DF4706F50A302C0C7F4AE22AF8B96C48">
We recorded
<taxonomicName id="325E165EBFD0FC824983E41BC4A2400A" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="300A91A478B7F1FE1E3AA9A8CD22AB19" italics="true">Cyrtodactylus kiphire</emphasis>
</taxonomicName>
<emphasis id="0701829309F9FB9E9C5FD2A447246C2A" bold="true">
<taxonomicNameLabel id="A1309266CAB55C67D3D47535B6C784C9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Kiphire forest colony, Kiphire District,
<collectingRegion id="1A731610357C0B82ACAE4CF190D61671" country="India" name="Nagaland">Nagaland</collectingRegion>
,
<collectingCountry id="4089BF9DF2F4ABABD1AF4DCE69C038EE" name="India">India</collectingCountry>
. The area is characterised as subtropical forest with regenerating jhum forest dominated by
<taxonomicName id="B4671D4638210AFE089E0EBD0B371B9F" authority="(Roxb. ex Lindl.) A. DC." authorityName="A. DC." baseAuthorityName="Lindl." class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="8C0717B0D2AB95845AF66A4BF87A828F" italics="true">Castanopsis indica</emphasis>
(Roxb. ex Lindl.) A. DC.
</taxonomicName>
,
<taxonomicName id="338DF10302A2896AE567D4BAE22FF9EB" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="F19DA128BB4D8C3E4E526698ABA96C9D" italics="true">Quercus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="C8E9C5A714998638011E0EB16503A3B1" authority="Wall. ex Roxb." authorityName="Roxb." class="Magnoliopsida" family="Iteaceae" genus="Itea" kingdom="Plantae" order="Saxifragales" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="AEBA572ED33B69487D97D8211A139E6E" italics="true">Itea macrophylla</emphasis>
Wall. ex Roxb.
</taxonomicName>
,
<taxonomicName id="ECB461268F53803A7C372CC05CAF258B" authority="(Osbeck) Merr" authorityName="Merr" baseAuthorityName="Osbeck" class="Magnoliopsida" family="Fabaceae" genus="Albizia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis id="57DC8F1543B2DFA132A4A202844401E3" italics="true">Albizia chinensis</emphasis>
(Osbeck) Merr
</taxonomicName>
.. The
<typeStatus id="496B55122EA9BC41F0F20B5A364D0DE7" type="holotype">holotype</typeStatus>
was collected from on a forest trail shrub (approximately 2.0 m from the ground) at approximately 23: 00 hrs. The
<typeStatus id="DEB7DC6D6974F3F8A01F48A53DFD7342" type="paratype">paratype</typeStatus>
was collected from a rocky slope along the edge of a stream.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,78 +1,80 @@
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<mods:title id="1C56D9FEABF457A0838F2CA480CB9FC7">Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India</mods:title>
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<mods:title id="B63FD8F5CC890DD77418EC746CAF2898">Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India</mods:title>
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<mods:name id="11204E9A14EC5890187EDA5EE8C920BE" type="personal">
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<mods:namePart id="D2E8709D493F387373348A81011D8D23">Boruah, Bitupan</mods:namePart>
<mods:nameIdentifier id="A58200510CEE49F25DC22F735EDFCFAC" type="ORCID">0000-0001-8829-6069</mods:nameIdentifier>
<mods:affiliation id="4E8034651808796DD463008E105E9657">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
<mods:namePart id="DD77B9CF2D6A8322EA439211BDA8E508">Boruah, Bitupan</mods:namePart>
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<mods:affiliation id="6DDA877E9DFE1ABBCB073B7D8817A0A1">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
</mods:name>
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<mods:roleTerm id="AC90CCAB5F0976766E6BC9325860133D">Author</mods:roleTerm>
<mods:name id="E881DB09A2563952BF20F1157A896BD3" type="personal">
<mods:role id="6E2018FC41E6A7A32A782513FB2C14B7">
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<mods:namePart id="6A6121635C01A5C81270DD4B18AD87DC">Narayanan, Surya</mods:namePart>
<mods:nameIdentifier id="312D7FDAB395F17F048E73A6ACD691E8" type="ORCID">0000-0001-9359-2815</mods:nameIdentifier>
<mods:affiliation id="9142F448CDFAA0EEACE5232C89900F10">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
<mods:namePart id="17A0D17A445C010094882084BD32B7C9">Narayanan, Surya</mods:namePart>
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<mods:affiliation id="19FDB1D07E3151696E1EC8A18C7C42B4">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
</mods:name>
<mods:name id="3CA4F7BEC06C5842FFA8F951D09F1878" type="personal">
<mods:role id="59CC1051669F8D559F39138CE856780E">
<mods:roleTerm id="8D551097C60F7262DCF02B4835117341">Author</mods:roleTerm>
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<mods:namePart id="252782A1250C77C5DA3BF91CDABBA5D7">Aravind, Neelavar Ananthram</mods:namePart>
<mods:nameIdentifier id="CFF065D7D9A7B354D01963576A97690E" type="ORCID">0000-0002-4515-8421</mods:nameIdentifier>
<mods:affiliation id="140B2A6076DE11008F192322E7DCEC5D">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India &amp; Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
<mods:namePart id="65D2BD919AEAEA78ADFB6F335656A696">Aravind, Neelavar Ananthram</mods:namePart>
<mods:nameIdentifier id="2C972D808CA20071BBDCE28DAF10353E" type="ORCID">0000-0002-4515-8421</mods:nameIdentifier>
<mods:affiliation id="C8690AEE44647BC04100C1A3CFFCA95B">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India &amp; Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
</mods:name>
<mods:name id="490DC007A8B12DB69E44EACB9BD5B66A" type="personal">
<mods:role id="2FA7A1BAC1F222C7FBC01EFCC895BE16">
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<mods:role id="CE7CE8227D7F6E85C65C2FD9876F03D8">
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</mods:role>
<mods:namePart id="CBC0A0D312FD91FA40BA96D5C6FC23E0">Lalronunga, Samuel</mods:namePart>
<mods:nameIdentifier id="DCE05B5206605C4D9D3DCB3109A8122F" type="ORCID">0000-0002-7542-4815</mods:nameIdentifier>
<mods:affiliation id="FB1E3FA7F2175ABCA9888A933E0D3C3F">Holy Child Society, Nalkata, Tripura 799263, India</mods:affiliation>
<mods:namePart id="5CAA2C951742C653D3E2B26F767B75CD">Lalronunga, Samuel</mods:namePart>
<mods:nameIdentifier id="5B664EA3246CD1495A96705C865B21A8" type="ORCID">0000-0002-7542-4815</mods:nameIdentifier>
<mods:affiliation id="DF03E1A7F98BCDB09C50088ACDFD3E2C">Holy Child Society, Nalkata, Tripura 799263, India</mods:affiliation>
</mods:name>
<mods:name id="87ABC24B1E0939EACE67E520061F1F90" type="personal">
<mods:role id="2F38BBE89E1D33BD1091D7F08A6A3EEA">
<mods:roleTerm id="DEBEFEE11A37A588DD562DE9C3E1B8CA">Author</mods:roleTerm>
<mods:name id="0EDD26B4BCEA2E3AA8B1491B6C4909DC" type="personal">
<mods:role id="85E6E2D1CBFB25B3518934DC6A46B69D">
<mods:roleTerm id="DD817AF2B5BBA528D409220FD8F2DFA7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C1FA7CA2FFDC58B478346D2FD4EC1AF">Deepak, V.</mods:namePart>
<mods:nameIdentifier id="E061C7411D9C953ADD689FB4CB9E3D64" type="ORCID">0000-0002-8826-9367</mods:nameIdentifier>
<mods:affiliation id="39BAA6C466E5A300AD488ECCA2973BA7">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
<mods:namePart id="B7A80D38EB3C343CC3D18D6A763222DE">Deepak, V.</mods:namePart>
<mods:nameIdentifier id="D48F57284DB47480330E819FD451BFF5" type="ORCID">0000-0002-8826-9367</mods:nameIdentifier>
<mods:affiliation id="6FF4C3C08BBABDAD870C72F12DD250F2">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
</mods:name>
<mods:name id="EB93B58530650B89C18AE49B99AE8DBA" type="personal">
<mods:role id="7B3C1D451E596FBA5A720ADCEB0003C5">
<mods:roleTerm id="98041AC69FA441DC6F4DC96734BC2219">Author</mods:roleTerm>
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<mods:roleTerm id="BB679127D4B3E63D95210996B55B35C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15DD785CAF25E1B38D3510588B7347B4">Das, Abhijit</mods:namePart>
<mods:nameIdentifier id="C1D3F49F40137C74FDFF05F0FA1050A0" type="ORCID">0000-0002-5851-8457</mods:nameIdentifier>
<mods:affiliation id="EFE2F259BC3C8AE23DDD1E0F2289B804">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
<mods:namePart id="1260CCCDAEAE9F26339E68321C347CEF">Das, Abhijit</mods:namePart>
<mods:nameIdentifier id="C541CA496181202CD241F186617C4BE6" type="ORCID">0000-0002-5851-8457</mods:nameIdentifier>
<mods:affiliation id="6DECE87BD861E6982E4E71EB486EDB8A">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="42138A24354914FB2AC6303F50791F02">text</mods:typeOfResource>
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<mods:titleInfo id="0D0F754F1E4388A74ABFEF5B32E43D15">
<mods:title id="AC269CDA522FF12FBAFC091053539524">Vertebrate Zoology</mods:title>
<mods:typeOfResource id="D7463223A2FFE9DCE407BD7AF4B51654">text</mods:typeOfResource>
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<mods:title id="63DA96FE8277C330E9B30858F16BD42E">Vertebrate Zoology</mods:title>
</mods:titleInfo>
<mods:part id="AF551DDAC85F96F54421C2D0C38D0538">
<mods:date id="9ED3A71781D86A237E09126A75F36E0B">2024</mods:date>
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<mods:date id="73869972784D82187E67C97B48117177">2024</mods:date>
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<mods:number id="77AB9D83A3399B2BBADE921E9601C638">2024-07-29</mods:number>
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<mods:detail id="32E8AE3EA3C9A11AC174858BB4EAA4F8" type="volume">
<mods:number id="703A736B3262D3A750EF2B9CC4BD5535">74</mods:number>
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<mods:number id="6F0D4DE1275D15F378CA931B80F7BEE1">74</mods:number>
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<mods:start id="C71860B0959FCA897C7B90A670D244A9">453</mods:start>
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<treatment id="CB20AD95DD3E53A58D7FC3B88F1BCC17" ID-DOI="http://doi.org/10.5281/zenodo.13124037" ID-Zenodo-Dep="13124037" ID-arpha="CB20AD95-DD3E-53A5-8D7F-C3B88F1BCC17" ID-zoobank="https://zoobank.org/B53EA935-E7F2-480B-8771-240B3A6E9939" LSID="urn:lsid:plazi:treatment:CB20AD95DD3E53A58D7FC3B88F1BCC17" LSID-ZBK="urn:lsid:zoobank.org:act:B53EA935-E7F2-480B-8771-240B3A6E9939" httpUri="http://treatment.plazi.org/id/CB20AD95DD3E53A58D7FC3B88F1BCC17">
<subSubSection id="3269139D434245584C4351BA11B30001" type="nomenclature">
@ -100,7 +102,10 @@ Boruah, Narayanan, Aravind, Deepak &amp; Das
</paragraph>
<paragraph id="E095E7B39059F4D06C45837085032524">
<materialsCitation id="BEFE5CDC72BEC4D6B7A02F12E50D8A4A" collectingDate="2021-10-23" collectorName="Bitupan Boruah" country="India" elevation="210" latitude="28.1711" location="Bodak village" longLatPrecision="6" longitude="95.242" municipality="East Siang District" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
Adult female (WII-ADR 1177; Fig.
<specimenCount id="7D11E1E5092CB4DE1FB2CF32DBDA14EF" count="1" type="female">Adult female</specimenCount>
(
<specimenCode id="FED268BDB56D9EE98006876065E6F6EB">WII-ADR 1177</specimenCode>
; Fig.
<figureCitation id="B8FEFDE37A633B6AA2534EF4F53E2FA7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyrtodactylus siangensis sp. nov. A, B, E I holotype (WII-ADR 1177), A dorsal, and B ventral view of whole body; C paratype (WII-ADR 1582); D paratype (WII-ADR 1581); E dorsal, F ventral, and G lateral view of the head (WII-ADR 1177); H ventral view of left manus, and I left pes (WII-ADR 1177); J an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure9" httpUri="https://binary.pensoft.net/fig/1102275">9 A, B, E I</figureCitation>
), from Kalek stream (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHODK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.242000,28.171100]}">
@ -153,25 +158,35 @@ A
<emphasis id="1E8D5621599E17D5F187579C3339166B" bold="true">E</emphasis>
<emphasis id="814F943CC876D417723FAB59D3DCB73A" bold="true">I</emphasis>
holotype (WII-ADR 1177),
holotype (
<specimenCode id="1A93EF4A99A5571D4D16E963BC9DC50F">WII-ADR 1177</specimenCode>
),
<emphasis id="69631AD9EE12C8396A8DCA38406CC9B9" bold="true">A</emphasis>
dorsal, and
<emphasis id="303C83588077C86070CE3B2EF9AE4C3C" bold="true">B</emphasis>
ventral view of whole body;
<emphasis id="DC82E60793E6506D85F2B6339824CEA9" bold="true">C</emphasis>
paratype (WII-ADR 1582);
paratype (
<specimenCode id="FE03676EA09F28D944EE2F794A07E0E6">WII-ADR 1582</specimenCode>
);
<emphasis id="D51E513EDEB2CEE35D85927F7BED29F8" bold="true">D</emphasis>
paratype (WII-ADR 1581);
paratype (
<specimenCode id="8512200D7F4440261658C3AA38C31C44">WII-ADR 1581</specimenCode>
);
<emphasis id="3D722352F7AB86280F219A2375B2461E" bold="true">E</emphasis>
dorsal,
<emphasis id="5665FCDBE382F0F3E6C1E91DBA75A9F0" bold="true">F</emphasis>
ventral, and
<emphasis id="E2B7906F4B7B8D44E8368DE66198D696" bold="true">G</emphasis>
lateral view of the head (WII-ADR 1177);
lateral view of the head (
<specimenCode id="F105B9ECBC6F32BE4A535DEC6F589856">WII-ADR 1177</specimenCode>
);
<emphasis id="AE63576E38317412672F3FD2828E6657" bold="true">H</emphasis>
ventral view of left manus, and
<emphasis id="616F70A1A7445AE4EB39D135628DC8BA" bold="true">I</emphasis>
left pes (WII-ADR 1177);
left pes (
<specimenCode id="EDEE9548DAA307DFD3FC6B536474E80D">WII-ADR 1177</specimenCode>
);
<emphasis id="0AA545FE0820680272CEF63241944B10" bold="true">J</emphasis>
an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm.
<emphasis id="DECBFBEEC8BB3D7BAAFECA957F12EA71" bold="true">J</emphasis>
@ -187,15 +202,31 @@ not to scale.
</heading>
</paragraph>
<paragraph id="E1D7ECE38496E2DAFDE251A840CADF3D">
<specimenCount id="821D08F5249F402AA0B97869E0B30C96" count="2" type="adult">Two adult</specimenCount>
females (WII-ADR 1581 and WII-ADR 1582; Fig.
<materialsCitation id="F622F6D8870C98EEAB121AE37D5446F5" collectedFrom="from a hill slope" collectingDate="2022-06-15" collectorName="Bitupan Boruah" country="India" county="East Siang District" location="Kalek stream" specimenCode="WII-ADR 1581, WII-ADR 1582" specimenCount="2" specimenCount-female="2" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="9C50FE0BEFA625E022F82E940F26B0B3" count="2" type="female">Two adult females</specimenCount>
(
<specimenCode id="10FE3FBEC38528FABC84BF211052F9A7">WII-ADR 1581</specimenCode>
and
<specimenCode id="A24F3873B229A99BEF42910B60C7D583">WII-ADR 1582</specimenCode>
; Fig.
<figureCitation id="C23D8FCF326552F39CD8ECEDAD3869B8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyrtodactylus siangensis sp. nov. A, B, E I holotype (WII-ADR 1177), A dorsal, and B ventral view of whole body; C paratype (WII-ADR 1582); D paratype (WII-ADR 1581); E dorsal, F ventral, and G lateral view of the head (WII-ADR 1177); H ventral view of left manus, and I left pes (WII-ADR 1177); J an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure9" httpUri="https://binary.pensoft.net/fig/1102275">9 C, D</figureCitation>
) from a hill slope near Kalek stream, East Siang District,
)
<collectedFrom id="F17B8DA2F5FBC6748203CAC26A3D0A95">from a hill slope</collectedFrom>
near
<location id="A0BAF2D7AF1348CB30884B30DCCF7C18" LSID="urn:lsid:plazi:treatment:CB20AD95DD3E53A58D7FC3B88F1BCC17:A0BAF2D7AF1348CB30884B30DCCF7C18" country="India" county="East Siang District" name="Kalek stream" stateProvince="Arunachal Pradesh">Kalek stream</location>
,
<collectingCounty id="9A6A64A9A4617A0C9157B0311541446D">East Siang District</collectingCounty>
,
<collectingRegion id="18536F4F9E01708FCFC64337D1E8289C" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="0DF3FA7A8DCA8039D8D4B25E17A1FB47" name="India">India</collectingCountry>
collected by Bitupan Boruah on
<date id="1BC5B8AB66F8E7F627F99424C7D0919A" value="2022-06-15">15 June 2022</date>
collected by
<collectorName id="29D86A2165335050B25E0428CF7B83FD">Bitupan Boruah</collectorName>
on
<date id="1BC5B8AB66F8E7F627F99424C7D0919A" value="2022-06-15">
<collectingDate id="A8381F34FD3219BFB2D0434312528D47" value="2022-06-15">15 June 2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -293,7 +324,9 @@ are variable in the form of irregular zig-zagged bands to reticulation; reticula
<specimenCount id="A1B9E6929F0D9E0190F860205A87D672" count="2" type="generic" typeStatus="paratype">two paratypes</specimenCount>
distinct than the
<typeStatus id="82BE053DCEFAE93EDCB7E8966D1C87A9" type="holotype">holotype</typeStatus>
; cross bands on tail of WII-ADR 1581 darker posteriorly (Fig.
; cross bands on tail of
<specimenCode id="8D9790E549245F669F8F4A24F7B2521A">WII-ADR 1581</specimenCode>
darker posteriorly (Fig.
<figureCitation id="F9E77C93362C422507286D4086577A14" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyrtodactylus siangensis sp. nov. A, B, E I holotype (WII-ADR 1177), A dorsal, and B ventral view of whole body; C paratype (WII-ADR 1582); D paratype (WII-ADR 1581); E dorsal, F ventral, and G lateral view of the head (WII-ADR 1177); H ventral view of left manus, and I left pes (WII-ADR 1177); J an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure9" httpUri="https://binary.pensoft.net/fig/1102275">9</figureCitation>
).
</paragraph>
@ -316,9 +349,7 @@ is close to
</taxonomicName>
, however, phylogenetically it is distinct from the later (Fig.
<figureCitation id="E493ADCCD8DC4BDD209B5132B74295CE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. ML phylogeny showing the relationship of the khasiensis group. New species described here are marked in blue, other samples generated in this study are in bold. Values in the node represent PP and UFB on the left and right side, respectively. See Fig. S 1 for a full tree with outgroups." figureDoi="10.3897/vz.74.e124752.figure2" httpUri="https://binary.pensoft.net/fig/1102268">2</figureCitation>
; Table S 3), also the
<typeStatus id="DBD0E909526F0BFEB24AD2015A63052B">type</typeStatus>
locality of the new species is separated by Siang River from the
; Table S 3), also the type locality of the new species is separated by Siang River from the
<taxonomicName id="A71D22EDDB0C8781C5FE30901522709A" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="79B4D84F12B7C4F376A7C3A3ED7C05EC" italics="true">C. cayuensis</emphasis>
</taxonomicName>
@ -356,9 +387,7 @@ has a high genetic divergence of 11.717.0 % from its closely related and the
<heading id="7E9DEA0816DA5C12EFD99A8E0B4267CC" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="ADF92A7891E5B7337EBF73F0C33EAC49">
The specific epithet is a toponym derived from the name of the river “ Siang ”. The
<typeStatus id="FCEAF392A1B25D5387698503822F96CD">type</typeStatus>
locality of this species lies in the Siang valley of
The specific epithet is a toponym derived from the name of the river “ Siang ”. The type locality of this species lies in the Siang valley of
<collectingRegion id="0F215FDB60C2462F59AAB4544F16D66E" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
.
</paragraph>