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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.219.3406" ID-PMC="PMC3433696" ID-Pensoft-Pub="1313-2970-219-11" ID-Pensoft-UUID="FFE7FFDBAA3AFF8BF81AFFD7FFCDFF87" ID-PubMed="22977349" ID-Zenodo-Dep="577535" ID-ZooBank="35A0EBEF815740B5BE499DBD7B273918" ModsDocID="1313-2970-219-11" checkinTime="1451248767646" checkinUser="pensoft" docAuthor="Gloeer, Peter &amp; Pesic, Vladimir" docDate="2012" docId="A15C0206B600938C0A3B29714DC8303A" docLanguage="en" docName="ZooKeys 219: 11-61" docOrigin="ZooKeys 219" docPubDate="2012-09-04" docSource="http://dx.doi.org/10.3897/zookeys.219.3406" docTitle="Radix bactriana" docType="treatment" docVersion="7" id="FFE7FFDBAA3AFF8BF81AFFD7FFCDFF87" lastPageNumber="25" masterDocId="FFE7FFDBAA3AFF8BF81AFFD7FFCDFF87" masterDocTitle="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species" masterLastPageNumber="61" masterPageNumber="11" pageNumber="25" updateTime="1668154421448" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gloeer, Peter</mods:namePart>
<mods:affiliation>Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pesic, Vladimir</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Sciences, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>219</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>61</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.219.3406</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.219.3406</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-219-11</mods:identifier>
<mods:identifier type="ZooBank">35A0EBEF815740B5BE499DBD7B273918</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577535</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152036891" LSID="urn:lsid:plazi:treatment:A15C0206B600938C0A3B29714DC8303A" httpUri="http://treatment.plazi.org/id/A15C0206B600938C0A3B29714DC8303A" lastPageNumber="25" pageId="14" pageNumber="25">
<subSubSection pageId="14" pageNumber="25" type="nomenclature">
<paragraph pageId="14" pageNumber="25">
<taxonomicName LSID="A15C0206-B600-938C-0A3B-29714DC8303A" authority="(Annandale &amp; Prashad, 1919)" baseAuthorityName="Annandale &amp; Prashad" baseAuthorityYear="1919" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Radix bactriana" order="Basommatophora" pageId="14" pageNumber="25" phylum="Mollusca" rank="species" species="bactriana">Radix bactriana (Annandale &amp; Prashad, 1919)</taxonomicName>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. The Lymnaeidae of Iran. a Radix persica (IR 27 - 07) b-d Radix bactriana (b IR 03 - 05 c IR 87 - 05 d IR 88 - 05 e IR 91 - 05) f Radix persica (IR 107 - 05) g Radix iranica (IR 89 - 05) h Radix sp. i Radix sp. k Galba truncatula (IR 62 - 05) l Galba schirazensis." httpUri="https://binary.pensoft.net/fig/20827" pageId="14" pageNumber="25">Figs 16b-d</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="25" type="records from iran">
<paragraph pageId="14" pageNumber="25">Records from Iran.</paragraph>
<paragraph pageId="14" pageNumber="25">
Seistan and Baluchestan Province (
<bibRefCitation author="Annandale, N" journalOrPublisher="Records of the Indian Museum" pageId="19" pageNumber="30" pagination="18 - 62" refId="B3" refString="Annandale, N, Prashad, B, 1919. The Mollusca Fauna of the Inland Waters of Baluchistan and of Seistan. Records of the Indian Museum 18: 18 - 62" title="The Mollusca Fauna of the Inland Waters of Baluchistan and of Seistan." volume="18" year="1919">Annandale and Prashad 1919</bibRefCitation>
); Kerman Province (
<bibRefCitation author="Starmuehlner, F" journalOrPublisher="Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I," pageId="19" pageNumber="30" refId="B57" refString="Starmuehlner, F, Edlauer, A, 1957. Ergebnisse der Oesterreichischen Iran-Expedition 1949/50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I, 166 (9/10): 435-494." title="Ergebnisse der Oesterreichischen Iran-Expedition 1949 / 50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I, 166 (9 / 10): 435 - 494." year="1957">
<normalizedToken originalValue="Starmühlner">Starmuehlner</normalizedToken>
and Edlauer 1957
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="25" type="new records">
<paragraph pageId="14" pageNumber="25">New records.</paragraph>
<paragraph pageId="14" pageNumber="25">Markazi Province: IR03-05 [1 ex], IR87-05 [9 ex.], IR88-05 [3 ex.], IR89-05 [2 ex], IR91-05 [3 ex.]; Khorasan Province: IR67-05 [1 ex.], IR79-05 [1 ex.].</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="25" type="distribution">
<paragraph pageId="14" pageNumber="25">Distribution.</paragraph>
<paragraph pageId="14" pageNumber="25">Iran:Seistan and Baluchestan and Kerman Provinces.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="67C1DB3E975ECB43163E00BAD4506526">Seven new species of cestode parasites (Neodermata, Platyhelminthes) from Australian birds</mods:title>
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<mods:affiliation id="23047F95A8DED66873C4A09F618BF158">Natural History Museum of Geneva, CP 6434, 1211 Geneva 6, Switzerland. Department of Genetics and Evolution, University of Geneva, Boulevard d’Ivoy 4, 1205 Geneva, Switzerland. Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113 Sofia, Bulgaria.</mods:affiliation>
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<mods:namePart id="863FD77738D2B59AA1BB4C1AB8AADEFB">Georgiev, Boyko B.</mods:namePart>
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<treatment id="A15C4A58FFD5A562FE23FCA9FB6FFCD1" ID-DOI="http://doi.org/10.5281/zenodo.3846850" ID-GBIF-Taxon="164152615" ID-Zenodo-Dep="3846850" LSID="urn:lsid:plazi:treatment:A15C4A58FFD5A562FE23FCA9FB6FFCD1" httpUri="http://treatment.plazi.org/id/A15C4A58FFD5A562FE23FCA9FB6FFCD1" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="61EFA8C5FFD5A565FE23FCA9FBB8FCB9" box="[455,1132,843,870]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="294AFB4EFFD5A565FE23FCA9FBB8FCB9" blockId="35.[455,1132,843,905]" box="[455,1132,843,870]" pageId="35" pageNumber="36">
<heading id="72024C22FFD5A565FE23FCA9FBB8FCB9" box="[455,1132,843,870]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="36" reason="2">
<taxonomicName id="EEF580CDFFD5A565FE23FCA9FBB8FCB9" authority="(Maplestone &amp; Southwell, 1923)" authorityName="Hooks. Scale" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[455,1132,843,870]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD5A565FE23FCA9FD37FCBA" bold="true" box="[455,739,843,870]" italics="true" pageId="35" pageNumber="36">Cracticotaenia fieldingi</emphasis>
(
<bibRefCitation id="4D6486BFFFD5A565FD16FCA9FBB0FCB9" author="Maplestone P. &amp; Southwell T." box="[754,1124,843,869]" pageId="35" pageNumber="36" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell, 1923</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD5A565FD14FC8CFC97FC55" box="[752,835,878,905]" pageId="35" pageNumber="36" type="description">
<paragraph id="294AFB4EFFD5A565FD14FC8CFC97FC55" blockId="35.[455,1132,843,905]" box="[752,835,878,905]" pageId="35" pageNumber="36">
<figureCitation id="B1CEE7CBFFD5A565FD14FC8CFC97FC55" box="[752,835,878,905]" captionStart="Figs 39–41. 39" captionStartId="35.[189,241,1941,1967]" captionTargetBox="[201,1367,1516,1875]" captionTargetId="figure@35.[195,1393,1513,1883]" captionTargetPageId="35" captionText="Figs 39–41. 39. Dilepididae genus sp. Hooks. 40. Passerilepis sp. Hooks. 41. Cracticotaenia fieldingi (Maplestone &amp; Southwell, 1923) Hooks. Scale bars: 39–41 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827570" httpUri="https://zenodo.org/record/3827570/files/figure.png" pageId="35" pageNumber="36">Fig. 41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD5A565FF59FC52FD91FB5A" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="294AFB4EFFD5A565FF59FC52FE7EFC16" blockId="35.[189,426,944,970]" box="[189,426,944,970]" pageId="35" pageNumber="36">
<heading id="72024C22FFD5A565FF59FC52FE7EFC16" bold="true" box="[189,426,944,970]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="1B81275CFFD5A565FF59FC52FE7EFC16" bold="true" box="[189,426,944,970]" pageId="35" pageNumber="36">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD5A565FF59FC3DFD91FB5A" blockId="35.[189,1399,989,1158]" pageId="35" pageNumber="36">
<collectingCountry id="51E2BBDEFFD5A565FF59FC3DFEBCFC25" box="[189,360,991,1017]" name="Australia" pageId="35" pageNumber="36">AUSTRALIA</collectingCountry>
:
<specimenCount id="3FF330C7FFD5A565FE9CFC3DFE06FC25" box="[376,466,991,1017]" pageId="35" pageNumber="36" type="generic">1 spec.</specimenCount>
,
<collectingRegion id="EB3135ACFFD5A565FE07FC3DFD16FC24" box="[483,706,990,1017]" country="Australia" name="New South Wales" pageId="35" pageNumber="36">New South Wales</collectingRegion>
, coll. Tonks, Coluvale,
<geoCoordinate id="4CC19D89FFD5A565FC0FFC3FFB83FC25" box="[1003,1111,989,1017]" degrees="34" direction="south" minutes="24" orientation="latitude" pageId="35" pageNumber="36" precision="925" value="-34.4">34°24′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD5A565FB8DFC3FFB31FC25" box="[1129,1253,989,1017]" degrees="150" direction="east" minutes="30" orientation="longitude" pageId="35" pageNumber="36" precision="925" value="150.5">150°30′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD5A565FB12FC3DFAA6FC25" box="[1270,1394,991,1017]" pageId="35" pageNumber="36" value="1911-04">Apr. 1911</date>
,
<collectionCode id="4FE4638BFFD5A565FF59FBE0FF05FBC0" box="[189,209,1026,1052]" name="Tavera, Department of Geology and Geophysics" pageId="35" pageNumber="36">T</collectionCode>
.H. Johnston leg., host:
<taxonomicName id="EEF580CDFFD5A565FE17FBE0FC55FBC0" authority="(Shaw, 1790)" baseAuthorityName="Shaw" baseAuthorityYear="1790" box="[499,897,1026,1052]" class="Aves" family="Cracticidae" genus="Strepera" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="graculina">
<emphasis id="1B81275CFFD5A565FE17FBE0FD01FBC0" box="[499,725,1026,1052]" italics="true" pageId="35" pageNumber="36">Strepera graculina</emphasis>
(Shaw, 1790)
</taxonomicName>
(AHC 20393); 2 fragmented specs from small intestine,
<collectingRegion id="EB3135ACFFD5A565FE9EFBC7FD85FBE3" box="[378,593,1061,1087]" country="Australia" name="New South Wales" pageId="35" pageNumber="36">New South Wales</collectingRegion>
, Taronga Zoo (Sydney),
<date id="5D4BDD8EFFD5A565FC98FBC7FC2DFBE3" box="[892,1017,1061,1087]" pageId="35" pageNumber="36" value="2000-03">Mar. 2000</date>
, Karie Rose leg., host:
<taxonomicName id="EEF580CDFFD5A565FAF7FBC7FE27FBBE" authority="(Latham, 1801)" baseAuthorityName="Latham" baseAuthorityYear="1801" class="Aves" family="Cracticidae" genus="Strepera" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="1B81275CFFD5A565FAF7FBC7FEE0FBBE" italics="true" pageId="35" pageNumber="36">Strepera versicolor</emphasis>
(Latham, 1801)
</taxonomicName>
(AHC 28975); 3 specs,
<collectingRegion id="EB3135ACFFD5A565FCF4FBABFC31FBBE" box="[784,997,1096,1123]" country="Australia" name="New South Wales" pageId="35" pageNumber="36">New South Wales</collectingRegion>
, Rydal,
<geoCoordinate id="4CC19D89FFD5A565FBA0FBA4FB7FFBBE" box="[1092,1195,1094,1122]" degrees="33" direction="south" minutes="30" orientation="latitude" pageId="35" pageNumber="36" precision="925" value="-33.5">33°30′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD5A565FB5CFBA4FAF4FBBF" box="[1208,1312,1094,1123]" degrees="150" direction="east" minutes="0" orientation="longitude" pageId="35" pageNumber="36" precision="925" value="150.0">150°0′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD5A565FAC9FBAAFF2EFB59" pageId="35" pageNumber="36" value="1906-02-04">4 Feb. 1906</date>
, Moreau leg. (AHC 36526).
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD5A565FF59FB4FFABCFB29" box="[189,1384,1197,1270]" pageId="35" pageNumber="36" type="biology_ecology">
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<emphasis id="1B81275CFFD5A565FF59FB4FFED7FB1B" bold="true" box="[189,259,1197,1223]" pageId="35" pageNumber="36">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD5A565FF59FB3EFABCFB29" blockId="35.[189,1384,1243,1270]" box="[189,1384,1243,1270]" pageId="35" pageNumber="36">
<taxonomicName id="EEF580CDFFD5A565FF59FB3EFDEAFB29" authority="(Shaw, 1790)" baseAuthorityName="Shaw" baseAuthorityYear="1790" box="[189,574,1243,1270]" class="Aves" family="Cracticidae" genus="Strepera" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="graculina">
<emphasis id="1B81275CFFD5A565FF59FB3EFE4EFB29" box="[189,410,1243,1270]" italics="true" pageId="35" pageNumber="36">Strepera graculina</emphasis>
(Shaw, 1790)
</taxonomicName>
and
<taxonomicName id="EEF580CDFFD5A565FD93FB3EFBC3FB29" authority="(Latham, 1801)" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[631,1047,1243,1270]" class="Aves" family="Cracticidae" genus="Strepera" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="1B81275CFFD5A565FD93FB3EFC8DFB29" box="[631,857,1243,1270]" italics="true" pageId="35" pageNumber="36">Strepera versicolor</emphasis>
(Latham, 1801)
</taxonomicName>
(
<taxonomicName id="EEF580CDFFD5A565FBC2FB39FB73FB29" authorityName="Vigors" authorityYear="1825" box="[1062,1191,1243,1269]" class="Aves" family="Artamidae" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="family">Artamidae</taxonomicName>
,
<taxonomicName id="EEF580CDFFD5A565FB56FB39FA8DFB2A" box="[1202,1369,1243,1270]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD5A562FF59FAFFFB6FFCD1" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="294AFB4EFFD5A565FF59FAFFFE9AFAEB" blockId="35.[189,334,1309,1335]" box="[189,334,1309,1335]" pageId="35" pageNumber="36">
<heading id="72024C22FFD5A565FF59FAFFFE9AFAEB" bold="true" box="[189,334,1309,1335]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="1B81275CFFD5A565FF59FAFFFE9AFAEB" bold="true" box="[189,334,1309,1335]" pageId="35" pageNumber="36">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD5A562FF59FAA9FC1FFDC1" blockId="35.[189,1399,1355,1452]" lastBlockId="36.[189,1399,267,541]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
This material is very similar to
<taxonomicName id="EEF580CDFFD5A565FDB7FAAEFD24FAB9" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[595,752,1355,1382]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="species" species="adelaidae">
<emphasis id="1B81275CFFD5A565FDB7FAAEFD24FAB9" box="[595,752,1355,1382]" italics="true" pageId="35" pageNumber="36">C. adelaidae</emphasis>
</taxonomicName>
described above, in most of its morpho-anatomical characters. A clear difference lies however in the shape and size of copulatory organs. While the cirrussac of
<taxonomicName id="EEF580CDFFD5A565FEEDFA70FE4BFA70" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[265,415,1426,1452]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="species" species="adelaidae">
<emphasis id="1B81275CFFD5A565FEEDFA70FE4BFA70" box="[265,415,1426,1452]" italics="true" pageId="35" pageNumber="36">C. adelaidae</emphasis>
</taxonomicName>
is short and globular, it is much more elongated and cylindrical here, always clearly crossing the excretory canals and 70% longer, reaching 155–193 × 30–40 (176 × 36, n = 20). Another discriminating character is the presence of a massive terminal vaginal sphincter, which is absent in
<emphasis id="1B81275CFFD2A562FF59FEB0FE8AFEB0" box="[189,350,338,364]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="EEF580CDFFD2A562FF59FEB0FE8FFEB0" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[189,347,338,364]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="adelaidae">C. adelaidae</taxonomicName>
.
</emphasis>
Hooks are 23–25 (24, n = 9) and 22–24 (22.5, n = 9) long (
<figureCitation id="B1CEE7CBFFD2A562FBA9FEB0FB7CFEB0" box="[1101,1192,338,364]" captionStart="Figs 39–41. 39" captionStartId="35.[189,241,1941,1967]" captionTargetBox="[201,1367,1516,1875]" captionTargetId="figure@35.[195,1393,1513,1883]" captionTargetPageId="35" captionText="Figs 39–41. 39. Dilepididae genus sp. Hooks. 40. Passerilepis sp. Hooks. 41. Cracticotaenia fieldingi (Maplestone &amp; Southwell, 1923) Hooks. Scale bars: 39–41 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827570" httpUri="https://zenodo.org/record/3827570/files/figure.png" pageId="36" pageNumber="37">Fig. 41</figureCitation>
). The material is however macerated and difficult to interpret, and it is not possible to associate rostellar hook counts and other measurements to specimens for which anatomy is available. The few available specimens from AHC 36526 include some long ones (up to
<quantity id="EE0D56ABFFD2A562FD34FE59FCEBFE0A" box="[720,831,443,470]" metricMagnitude="-2" metricUnit="m" metricValue="3.92" pageId="36" pageNumber="37" unit="mm" value="39.2">39.2 mm</quantity>
), including one with a complete scolex (scolex diameter 525, suckers diameter 200–220, about 88 hooks). A vaginal sphincter is present and cirrus-sacs are of a similar shape as
<taxonomicName id="EEF580CDFFD2A562FE3BFDE0FDB5FDC0" authorityName="Hooks. Scale" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[479,609,514,540]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD2A562FE3BFDE0FDB5FDC0" box="[479,609,514,540]" italics="true" pageId="36" pageNumber="37">C. fieldingi</emphasis>
</taxonomicName>
but larger (235–300 × 38–65).
</paragraph>
<caption id="7D8AABC6FFD5A565FF59F877FC31F80F" ID-DOI="http://doi.org/10.5281/zenodo.3827570" ID-Zenodo-Dep="3827570" httpUri="https://zenodo.org/record/3827570/files/figure.png" pageId="35" pageNumber="36" startId="35.[189,241,1941,1967]" targetBox="[201,1367,1516,1875]" targetPageId="35">
<paragraph id="294AFB4EFFD5A565FF59F877FC31F80F" blockId="35.[189,1398,1941,2003]" pageId="35" pageNumber="36">
<emphasis id="1B81275CFFD5A565FF59F877FEA4F86C" bold="true" box="[189,368,1941,1968]" pageId="35" pageNumber="36">Figs 39–41. 39</emphasis>
.
<taxonomicName id="EEF580CDFFD5A565FE64F874FDDDF86C" authorityName="Fuhrmann" authorityYear="1907" box="[384,521,1942,1968]" class="Cestoda" family="Dilepididae" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="family">Dilepididae</taxonomicName>
genus sp. Hooks.
<emphasis id="1B81275CFFD5A565FD01F874FCD6F86C" bold="true" box="[741,770,1942,1968]" pageId="35" pageNumber="36">40</emphasis>
.
<taxonomicName id="EEF580CDFFD5A565FCF6F874FC75F86C" authorityName="Spasskii &amp; Spasskaya" authorityYear="1954" box="[786,929,1942,1968]" class="Cestoda" family="Hymenolepididae" genus="Passerilepis" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFD5A565FCF6F874FC75F86C" box="[786,929,1942,1968]" italics="true" pageId="35" pageNumber="36">Passerilepis</emphasis>
</taxonomicName>
sp. Hooks.
<emphasis id="1B81275CFFD5A565FBD5F874FB9AF86C" bold="true" box="[1073,1102,1942,1968]" pageId="35" pageNumber="36">41</emphasis>
.
<taxonomicName id="EEF580CDFFD5A565FBBAF874FD36F80F" authority="(Maplestone &amp; Southwell, 1923) Hooks. Scale" authorityName="Hooks. Scale" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD5A565FBBAF874FAA2F86C" box="[1118,1398,1942,1968]" italics="true" pageId="35" pageNumber="36">Cracticotaenia fieldingi</emphasis>
(
<bibRefCitation id="4D6486BFFFD5A565FF21F85BFDE3F80F" author="Maplestone P. &amp; Southwell T." box="[197,567,1977,2003]" pageId="35" pageNumber="36" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell, 1923</bibRefCitation>
) Hooks. Scale
</taxonomicName>
bars: 39–41 = 20 µm.
</paragraph>
</caption>
<paragraph id="294AFB4EFFD2A562FF59FDA1FB6FFCD1" blockId="36.[189,1399,579,782]" pageId="36" pageNumber="37">
Globally, this material corresponds to the description of
<taxonomicName id="EEF580CDFFD2A562FC73FDA6FBF6FD81" authorityName="Hooks. Scale" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[919,1058,579,606]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD2A562FC73FDA6FBF6FD81" box="[919,1058,579,606]" italics="true" pageId="36" pageNumber="37">C. fieldingi</emphasis>
</taxonomicName>
from
<taxonomicName id="EEF580CDFFD2A562FB98FDA6FAA3FD81" authorityName="Temminck" authorityYear="1824" box="[1148,1399,579,606]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="destructor">
<emphasis id="1B81275CFFD2A562FB98FDA6FAA3FD81" box="[1148,1399,579,606]" italics="true" pageId="36" pageNumber="37">Cracticus destructor</emphasis>
</taxonomicName>
[=
<taxonomicName id="EEF580CDFFD2A562FF39FD85FEB9FD5C" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[221,365,615,641]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="1B81275CFFD2A562FF39FD85FEB9FD5C" box="[221,365,615,641]" italics="true" pageId="36" pageNumber="37">C. torquatus</emphasis>
</taxonomicName>
] as described by
<bibRefCitation id="4D6486BFFFD2A562FDD1FD84FC78FD5C" author="Maplestone P. &amp; Southwell T." box="[565,940,614,640]" pageId="36" pageNumber="37" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell (1923)</bibRefCitation>
and observed by ourselves (see above). It should, however, be noted that
<taxonomicName id="EEF580CDFFD2A562FDB4FD68FD00FD78" authorityName="Hooks. Scale" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[592,724,650,676]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD2A562FDB4FD68FD00FD78" box="[592,724,650,676]" italics="true" pageId="36" pageNumber="37">C. fieldingi</emphasis>
</taxonomicName>
has not only been described in a different host, but also from a locality in
<collectingRegion id="EB3135ACFFD2A562FE6FFD4FFDC1FD1B" box="[395,533,685,711]" country="Australia" name="Queensland" pageId="36" pageNumber="37">Queensland</collectingRegion>
which is some
<quantity id="EE0D56ABFFD2A562FD2CFD4FFCFAFD1B" box="[712,814,685,712]" metricMagnitude="6" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="37" unit="km" value="1500.0">1500 km</quantity>
north of the
<collectingRegion id="EB3135ACFFD2A562FC26FD4FFB45FD1B" box="[962,1169,685,711]" country="Australia" name="New South Wales" pageId="36" pageNumber="37">New South Wales</collectingRegion>
stations considered here. The available specimens do not allow for a detailed study, but the variability observed in our different samples suggest a possible diversification of
<taxonomicName id="EEF580CDFFD2A562FCDDFD16FC3DFCD2" authorityName="Spasskii" authorityYear="1966" box="[825,1001,756,782]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFD2A562FCDDFD16FC3DFCD2" box="[825,1001,756,782]" italics="true" pageId="36" pageNumber="37">Cracticotaenia</emphasis>
</taxonomicName>
in the
<taxonomicName id="EEF580CDFFD2A562FBDCFD11FB6CFCD1" authorityName="Vigors" authorityYear="1825" box="[1080,1208,755,781]" class="Aves" family="Artamidae" kingdom="Animalia" order="Passeriformes" pageId="36" pageNumber="37" phylum="Chordata" rank="family">Artamidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

443
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<subSubSection id="61EFA8C5FFD6A566FDB7FE92FB6BFE72" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="294AFB4EFFD6A566FDB7FE92FC35FE56" blockId="32.[372,1215,368,464]" box="[595,993,368,395]" pageId="32" pageNumber="33">
<heading id="72024C22FFD6A566FDB7FE92FC35FE56" box="[595,993,368,395]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="33" reason="2">
<taxonomicName id="EEF580CDFFD6A566FDB7FE92FC54FE57" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[595,896,368,395]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="32" pageNumber="33" phylum="Platyhelminthes" rank="species" species="adelaidae" status="sp. nov.">
<emphasis id="1B81275CFFD6A566FDB7FE92FC54FE57" bold="true" box="[595,896,368,395]" italics="true" pageId="32" pageNumber="33">Cracticotaenia adelaidae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFD6A566FC63FE92FC35FE56" box="[903,993,368,394]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
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<uri id="5D64F74CFFD6A566FE90FE71FB6BFE72" box="[372,1215,403,430]" pageId="32" pageNumber="33">
urn:lsid:zoobank.org:act:
<uuid id="5D53C19BFFD6A566FD72FE71FB6BFE72" box="[662,1215,403,430]" pageId="32" pageNumber="33">5F32D506-1E05-421F-B891-DDB9E89F4195</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD6A566FD3CFE54FC8FFE0C" box="[728,859,438,464]" pageId="32" pageNumber="33" type="description">
<paragraph id="294AFB4EFFD6A566FD3CFE54FC8FFE0C" blockId="32.[372,1215,368,464]" box="[728,859,438,464]" pageId="32" pageNumber="33">
<figureCitation id="B1CEE7CBFFD6A566FD3CFE54FC8FFE0C" box="[728,859,438,464]" captionStart="Figs 34–38" captionStartId="33.[189,241,1911,1937]" captionTargetBox="[225,1380,276,1817]" captionTargetId="figure@33.[208,1380,265,1873]" captionTargetPageId="33" captionText="Figs 34–38. Cracticotaenia adelaidae sp. nov. 34. Scolex. 35. Detail of hooks crowns. 36. Hooks: anterior (A) and posterior (B). 37. Mature proglottis. 38. Eggs. Scale bars: 34 = 100 µm; 35 = 10 µm; 19 = 500 µm; 36, 38 = 20 µm; 37 = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3827568" httpUri="https://zenodo.org/record/3827568/files/figure.png" pageId="32" pageNumber="33">Figs 34–38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD6A566FF59FE1AFAA3FD9C" box="[189,1399,504,577]" pageId="32" pageNumber="33" type="etymology">
<paragraph id="294AFB4EFFD6A566FF59FE1AFE90FDCE" blockId="32.[189,324,504,530]" box="[189,324,504,530]" pageId="32" pageNumber="33">
<heading id="72024C22FFD6A566FF59FE1AFE90FDCE" bold="true" box="[189,324,504,530]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="1B81275CFFD6A566FF59FE1AFE90FDCE" bold="true" box="[189,324,504,530]" pageId="32" pageNumber="33">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59FDC4FAA3FD9C" blockId="32.[189,1399,550,577]" box="[189,1399,550,577]" pageId="32" pageNumber="33">
The species name is derived from the name of the City of Adelaide, where the
<typeStatus id="F64E45ECFFD6A566FBAAFDC5FB55FD9D" box="[1102,1153,551,577]" pageId="32" pageNumber="33">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD6A566FF59FD8AFEF3FA30" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="294AFB4EFFD6A566FF59FD8AFE7EFD5E" blockId="32.[189,426,616,642]" box="[189,426,616,642]" pageId="32" pageNumber="33">
<heading id="72024C22FFD6A566FF59FD8AFE7EFD5E" bold="true" box="[189,426,616,642]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="1B81275CFFD6A566FF59FD8AFE7EFD5E" bold="true" box="[189,426,616,642]" pageId="32" pageNumber="33">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="999DF113FFD6A566FF0EFD75FECCFD24" ID-GBIF-Occurrence="2619826302" collectingDate="1983-10-27" collectorName="David Shulz" country="Australia" latitude="-34.9" location="Adelaide" longLatPrecision="1197" longitude="138.6" pageId="32" pageNumber="33" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<paragraph id="294AFB4EFFD6A566FF0EFD75FE88FD6D" blockId="32.[189,1399,663,760]" box="[234,348,663,689]" pageId="32" pageNumber="33">
<emphasis id="1B81275CFFD6A566FF0EFD75FE88FD6D" bold="true" box="[234,348,663,689]" pageId="32" pageNumber="33">
<typeStatus id="F64E45ECFFD6A566FF0EFD75FE88FD6D" box="[234,348,663,689]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59FD58FECCFD24" blockId="32.[189,1399,663,760]" pageId="32" pageNumber="33">
<collectingCountry id="51E2BBDEFFD6A566FF59FD58FEBCFD08" box="[189,360,698,724]" name="Australia" pageId="32" pageNumber="33">AUSTRALIA</collectingCountry>
:
<collectingRegion id="EB3135ACFFD6A566FE90FD5BFDFBFD0F" box="[372,559,697,723]" country="Australia" name="South Australia" pageId="32" pageNumber="33">South Australia</collectingRegion>
,
<location id="2C2AAD95FFD6A566FDDEFD5BFD73FD0F" LSID="urn:lsid:plazi:treatment:A15C4A58FFD6A565FDB7FE92FEFAFCD1:2C2AAD95FFD6A566FDDEFD5BFD73FD0F" box="[570,679,697,723]" country="Australia" latitude="-34.9" longLatPrecision="1197" longitude="138.6" name="Adelaide" pageId="32" pageNumber="33" stateProvince="South Australia">Adelaide</location>
,
<geoCoordinate id="4CC19D89FFD6A566FD51FD5AFCC9FD08" box="[693,797,696,724]" degrees="34" direction="south" minutes="54" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-34.9">34°54′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD6A566FCCFFD5AFC70FD08" box="[811,932,696,724]" degrees="138" direction="east" minutes="36" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="138.6">138°36′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD6A566FC56FD58FB84FD08" box="[946,1104,698,724]" pageId="32" pageNumber="33" value="1983-10-27">
<collectingDate id="4D0F2466FFD6A566FC56FD58FB84FD08" box="[946,1104,698,724]" pageId="32" pageNumber="33" value="1983-10-27">27 Oct. 1983</collectingDate>
</date>
,
<collectorName id="84009E98FFD6A566FBBAFD5BFB26FD0F" box="[1118,1266,697,723]" pageId="32" pageNumber="33">David Shulz</collectorName>
leg. (AHC 36516).
</paragraph>
</materialsCitation>
<paragraph id="294AFB4EFFD6A566FF0EFCFCFEB3FCE4" blockId="32.[189,1036,798,860]" box="[234,359,798,824]" pageId="32" pageNumber="33">
<emphasis id="1B81275CFFD6A566FF0EFCFCFEB3FCE4" bold="true" box="[234,359,798,824]" pageId="32" pageNumber="33">
<typeStatus id="F64E45ECFFD6A566FF0EFCFCFEB3FCE4" box="[234,359,798,824]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59FCA0FBD8FC80" blockId="32.[189,1036,798,860]" box="[189,1036,833,860]" pageId="32" pageNumber="33">
<collectingCountry id="51E2BBDEFFD6A566FF59FCA0FEBCFC80" box="[189,360,834,860]" name="Australia" pageId="32" pageNumber="33">AUSTRALIA</collectingCountry>
: 5 specs, same data as for
<typeStatus id="F64E45ECFFD6A566FD78FCA3FCD7FC87" box="[668,771,833,859]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
(AHC 36517–36525).
</paragraph>
<paragraph id="294AFB4EFFD6A566FF0EFC61FD03FC41" blockId="32.[189,1399,899,1102]" box="[234,727,899,925]" pageId="32" pageNumber="33">
<emphasis id="1B81275CFFD6A566FF0EFC61FD03FC41" bold="true" box="[234,727,899,925]" pageId="32" pageNumber="33">Additional material from the same host</emphasis>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59FC44FBBEFB92" blockId="32.[189,1399,899,1102]" pageId="32" pageNumber="33">
<collectingCountry id="51E2BBDEFFD6A566FF59FC44FEBCFC1C" box="[189,360,934,960]" name="Australia" pageId="32" pageNumber="33">AUSTRALIA</collectingCountry>
:
<emphasis id="1B81275CFFD6A566FE90FC44FDEDFC1C" bold="true" box="[372,569,934,960]" pageId="32" pageNumber="33">
<collectingRegion id="EB3135ACFFD6A566FE90FC44FDEDFC1C" box="[372,569,934,960]" country="Australia" name="South Australia" pageId="32" pageNumber="33">South Australia</collectingRegion>
</emphasis>
3 specs,
<collectingRegion id="EB3135ACFFD6A566FD4CFC44FCB6FC1C" box="[680,866,934,960]" inferredBySuffix="true" pageId="32" pageNumber="33">South Australia</collectingRegion>
, Ashbourne,
<geoCoordinate id="4CC19D89FFD6A566FC1FFC46FBB6FC1C" box="[1019,1122,932,960]" degrees="35" direction="south" minutes="18" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-35.3">35°18′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD6A566FB94FC46FB3CFC1D" box="[1136,1256,932,961]" degrees="138" direction="east" minutes="36" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="138.6">138°36′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD6A566FB11FC45FAA5FC1C" box="[1269,1393,934,961]" pageId="32" pageNumber="33" value="1969-12">Dec. 1969</date>
, Michael Smyth and Lee leg. (AHC 29591); 2 specs, very fragmented, Coromandel Valley,
<geoCoordinate id="4CC19D89FFD6A566FAF2FC25FAA5FC3F" box="[1302,1393,967,995]" degrees="35" direction="south" minutes="6" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-35.1">35°6′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD6A566FF59FC09FEE7FBDB" box="[189,307,1003,1031]" degrees="138" direction="east" minutes="36" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="138.6">138°36′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD6A566FEDAFC0FFE1CFBDA" box="[318,456,1004,1031]" pageId="32" pageNumber="33" value="1969-04-04">4 Apr. 1969</date>
, Michael Smyth leg. (AHC 29592); about 30 specs, Coorong,
<geoCoordinate id="4CC19D89FFD6A566FB7DFC09FB24FBDB" box="[1177,1264,1003,1031]" degrees="36" direction="south" minutes="2" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-36.033333">36°2′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD6A566FB1FFC09FAA5FBDB" box="[1275,1393,1003,1031]" degrees="139" direction="east" minutes="33" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="139.55">139°33′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD6A566FF59FBF2FEE7FBF6" box="[189,307,1039,1066]" pageId="32" pageNumber="33" value="1935-10">Oct. 1935</date>
, D. Brunswick leg. (AHC 230). –
<emphasis id="1B81275CFFD6A566FD20FBEDFCFEFBF5" bold="true" box="[708,810,1039,1065]" pageId="32" pageNumber="33">
<collectingRegion id="EB3135ACFFD6A566FD20FBEDFCFEFBF5" box="[708,810,1039,1065]" country="Australia" name="Victoria" pageId="32" pageNumber="33">Victoria</collectingRegion>
</emphasis>
2 specs, Torquay,
<geoCoordinate id="4CC19D89FFD6A566FBE6FBECFBBCFBF6" box="[1026,1128,1038,1066]" degrees="38" direction="south" minutes="18" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-38.3">38°18′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD6A566FB91FBECFB3FFBF6" box="[1141,1259,1038,1066]" degrees="144" direction="east" minutes="18" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="144.3">144°18′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD6A566FB13FBF2FAA5FBF6" box="[1271,1393,1040,1066]" pageId="32" pageNumber="33" value="1979-05">May 1979</date>
, Karl Harrigan leg., from
<taxonomicName id="EEF580CDFFD6A566FE06FBD1FD92FB91" box="[482,582,1075,1101]" class="Aves" family="Cracticidae" genus="Strepera" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="1B81275CFFD6A566FE06FBD1FD92FB91" box="[482,582,1075,1101]" italics="true" pageId="32" pageNumber="33">Strepera</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="EEF580CDFFD6A566FD9AFBD1FD2BFB91" authorityName="Vigors" authorityYear="1825" box="[638,767,1075,1101]" class="Aves" family="Artamidae" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="family">Artamidae</taxonomicName>
,
<taxonomicName id="EEF580CDFFD6A566FCEEFBD1FC65FB92" box="[778,945,1075,1102]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
) (AHC 29596).
</paragraph>
<paragraph id="294AFB4EFFD6A566FF0EFB96FE72FB52" blockId="32.[189,1399,1140,1343]" box="[234,422,1140,1166]" pageId="32" pageNumber="33">
<emphasis id="1B81275CFFD6A566FF0EFB96FE72FB52" bold="true" box="[234,422,1140,1166]" pageId="32" pageNumber="33">Other material</emphasis>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59FB7AFB8BFAE3" blockId="32.[189,1399,1140,1343]" pageId="32" pageNumber="33">
<collectingCountry id="51E2BBDEFFD6A566FF59FB7AFEBCFB6E" box="[189,360,1176,1202]" name="Australia" pageId="32" pageNumber="33">AUSTRALIA</collectingCountry>
: from
<taxonomicName id="EEF580CDFFD6A566FE22FB7AFC53FB6E" authority="(Latham, 1801)" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[454,903,1175,1202]" class="Aves" family="Cracticidae" genus="Strepera" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="1B81275CFFD6A566FE22FB7AFD66FB6D" box="[454,690,1175,1202]" italics="true" pageId="32" pageNumber="33">Strepera versicolor</emphasis>
(Latham, 1801)
</taxonomicName>
: 6 specs and numerous fragments, no locality,
<date id="5D4BDD8EFFD6A566FEC7FB59FE16FB09" box="[291,450,1211,1237]" pageId="32" pageNumber="33" value="1910-12-12">12 Dec. 1910</date>
(AHC 533). – From
<taxonomicName id="EEF580CDFFD6A566FD5BFB59FB14FB09" authority="(Vieillot, 1817)" baseAuthorityName="Vieillot" baseAuthorityYear="1817" box="[703,1216,1211,1237]" class="Aves" family="Corcoracidae" genus="Corcorax" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="melanorhamphos">
<emphasis id="1B81275CFFD6A566FD5BFB59FBD6FB09" box="[703,1026,1211,1237]" italics="true" pageId="32" pageNumber="33">Corcorax melanorhamphos</emphasis>
(Vieillot, 1817)
</taxonomicName>
(
<taxonomicName id="EEF580CDFFD6A566FB35FB59FAA6FB09" box="[1233,1394,1211,1237]" class="Aves" family="Corcoracidae" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="family">Corcoracidae</taxonomicName>
,
<taxonomicName id="EEF580CDFFD6A566FF59FB3CFEBCFB25" box="[189,360,1246,1273]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
): 4 specs,
<collectingRegion id="EB3135ACFFD6A566FE0EFB3CFDACFB24" box="[490,632,1246,1272]" country="Australia" name="Queensland" pageId="32" pageNumber="33">Queensland</collectingRegion>
, Eidsvold,
<geoCoordinate id="4CC19D89FFD6A566FCE5FB3EFCBFFB24" box="[769,875,1244,1272]" degrees="25" direction="south" minutes="22" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-25.366667">25°22′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD6A566FC9FFB3EFC33FB24" box="[891,999,1244,1272]" degrees="151" direction="east" minutes="5" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="151.08333">151°5′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD6A566FC13FB3CFBA7FB24" box="[1015,1139,1246,1272]" pageId="32" pageNumber="33" value="1918-09">Sep. 1918</date>
, Thomas L. Bancroft leg. (AHC 531, AHC 29598, co-infection with a
<taxonomicName id="EEF580CDFFD6A566FCF5FAE3FC49FAC7" authorityName="Fuhrmann" authorityYear="1907" box="[785,925,1281,1307]" class="Cestoda" family="Dilepididae" kingdom="Animalia" order="Cyclophyllidea" pageId="32" pageNumber="33" phylum="Platyhelminthes" rank="family">Dilepididae</taxonomicName>
, see above); 12 specs,
<collectingRegion id="EB3135ACFFD6A566FB52FAE3FAA6FAC7" box="[1206,1394,1281,1307]" country="Australia" name="South Australia" pageId="32" pageNumber="33">South Australia</collectingRegion>
, Elwomple,
<geoCoordinate id="4CC19D89FFD6A566FEA0FAC0FE7EFAE2" box="[324,426,1314,1342]" degrees="35" direction="south" minutes="17" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-35.283333">35°17′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFD6A566FE53FAC0FDFAFAE3" box="[439,558,1314,1343]" degrees="139" direction="east" minutes="32" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="139.53334">139°32′ E</geoCoordinate>
,
<date id="5D4BDD8EFFD6A566FDDFFAC6FD01FAE2" box="[571,725,1316,1342]" pageId="32" pageNumber="33" value="1937-10-19">19 Oct. 1937</date>
, collector unknown (AHC 3368).
</paragraph>
<paragraph id="294AFB4EFFD6A566FF0EFA84FD94FA5C" blockId="32.[189,1399,1382,1516]" box="[234,576,1382,1408]" pageId="32" pageNumber="33">
<emphasis id="1B81275CFFD6A566FF0EFA84FD94FA5C" bold="true" box="[234,576,1382,1408]" pageId="32" pageNumber="33">Other comparison material</emphasis>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59FA6BFEF3FA30" blockId="32.[189,1399,1382,1516]" pageId="32" pageNumber="33">
Type material of
<taxonomicName id="EEF580CDFFD6A566FE6FFA6BFBE7FA7F" authority="(Maplestone &amp; Southwell, 1923)" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[395,1075,1417,1443]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="32" pageNumber="33" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD6A566FE6FFA6BFD4BFA7F" box="[395,671,1417,1443]" italics="true" pageId="32" pageNumber="33">Monopylidium fieldingi</emphasis>
(
<bibRefCitation id="4D6486BFFFD6A566FD54FA6BFBFEFA7F" author="Maplestone P. &amp; Southwell T." box="[688,1066,1417,1443]" pageId="32" pageNumber="33" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell, 1923</bibRefCitation>
)
</taxonomicName>
, from
<taxonomicName id="EEF580CDFFD6A566FB67FA6BFAA3FA7F" authorityName="Temminck" authorityYear="1824" box="[1155,1399,1417,1443]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="destructor">
<emphasis id="1B81275CFFD6A566FB67FA6BFAA3FA7F" box="[1155,1399,1417,1443]" italics="true" pageId="32" pageNumber="33">Cracticus destructor</emphasis>
</taxonomicName>
[now
<taxonomicName id="EEF580CDFFD6A566FEE7FA4DFE43FA14" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[259,407,1455,1481]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="1B81275CFFD6A566FEE7FA4DFE43FA14" box="[259,407,1455,1481]" italics="true" pageId="32" pageNumber="33">C. torquatus</emphasis>
</taxonomicName>
] in
<collectingRegion id="EB3135ACFFD6A566FE28FA4CFD8EFA14" box="[460,602,1454,1480]" country="Australia" name="Queensland" pageId="32" pageNumber="33">Queensland</collectingRegion>
, Townsville (The Natural History Museum, London 1977.11.3.34, 2 slides).
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD6A566FF59F9F1FEB9F917" pageId="32" pageNumber="33" type="biology_ecology">
<paragraph id="294AFB4EFFD6A566FF59F9F1FF2CF9F1" blockId="32.[189,248,1555,1581]" box="[189,248,1555,1581]" pageId="32" pageNumber="33">
<heading id="72024C22FFD6A566FF59F9F1FF2CF9F1" bold="true" box="[189,248,1555,1581]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="1B81275CFFD6A566FF59F9F1FF2CF9F1" bold="true" box="[189,248,1555,1581]" pageId="32" pageNumber="33">Host</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59F9A3FB07F980" blockId="32.[189,1235,1601,1628]" box="[189,1235,1601,1628]" pageId="32" pageNumber="33">
<taxonomicName id="EEF580CDFFD6A566FF59F9A3FDB2F980" authority="(Latham, 1801)" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[189,614,1601,1628]" class="Aves" family="Cracticidae" genus="Gymnorhina" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tibicen">
<emphasis id="1B81275CFFD6A566FF59F9A3FE7CF987" box="[189,424,1601,1627]" italics="true" pageId="32" pageNumber="33">Gymnorhina tibicen</emphasis>
(Latham, 1801)
</taxonomicName>
(syn.
<taxonomicName id="EEF580CDFFD6A566FD4AF9A0FCACF987" baseAuthorityName="Latham" baseAuthorityYear="1802" box="[686,888,1601,1628]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tibicen">
<emphasis id="1B81275CFFD6A566FD4AF9A0FCACF987" box="[686,888,1601,1628]" italics="true" pageId="32" pageNumber="33">Cracticus tibicen</emphasis>
</taxonomicName>
) (
<taxonomicName id="EEF580CDFFD6A566FC74F9A3FBC5F987" authorityName="Vigors" authorityYear="1825" box="[912,1041,1601,1627]" class="Aves" family="Artamidae" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="family">Artamidae</taxonomicName>
,
<taxonomicName id="EEF580CDFFD6A566FBF9F9A3FB13F980" box="[1053,1223,1601,1628]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="32" pageNumber="33" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
).
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59F961FEF8F941" blockId="32.[189,300,1667,1693]" box="[189,300,1667,1693]" pageId="32" pageNumber="33">
<heading id="72024C22FFD6A566FF59F961FEF8F941" bold="true" box="[189,300,1667,1693]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="1B81275CFFD6A566FF59F961FEF8F941" bold="true" box="[189,300,1667,1693]" pageId="32" pageNumber="33">Intensity</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD6A566FF59F953FEB9F917" blockId="32.[189,365,1713,1739]" box="[189,365,1713,1739]" pageId="32" pageNumber="33">Six specimens.</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD6A564FF59F911FC6BFA7C" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
<paragraph id="294AFB4EFFD6A566FF59F911FE9AF8D1" blockId="32.[189,334,1779,1805]" box="[189,334,1779,1805]" pageId="32" pageNumber="33">
<heading id="72024C22FFD6A566FF59F911FE9AF8D1" bold="true" box="[189,334,1779,1805]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="1B81275CFFD6A566FF59F911FE9AF8D1" bold="true" box="[189,334,1779,1805]" pageId="32" pageNumber="33">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD6A564FF59F8C3FD94FD59" blockId="32.[189,1399,1825,1993]" lastBlockId="34.[189,1399,267,646]" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
Body of medium size, with maximum length
<quantity id="EE0D56ABFFD6A566FD2BF8C3FCE3F8E0" box="[719,823,1825,1852]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="32" pageNumber="33" unit="mm" value="35.5">35.5 mm</quantity>
and maximum width
<quantity id="EE0D56ABFFD6A566FBD2F8C0FB06F8E0" box="[1078,1234,1826,1852]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" metricValueMax="2.12" metricValueMin="1.3" pageId="32" pageNumber="33" unit="mm" value="1.71" valueMax="2.12" valueMin="1.3">1.3–2.12 mm</quantity>
(1.7, n = 6) at level of pregravid or early gravid proglottides. Most complete specimen consisting of 240 proglottides. Proglottides craspedote, wider than long, progressively becoming longer, up to slightly longer than wide when gravid. Scolex rounded, not markedly delineated from neck, with diameter 350–475 (407, n = 8), Suckers unarmed, muscular, rounded, 142–198 (169, n = 35) in diameter.
<taxonomicName id="EEF580CDFFD6A566FB60F84CFB29F814" authorityName="Montfort" authorityYear="1810" box="[1156,1277,1966,1992]" family="Rostellum" pageId="32" pageNumber="18" rank="family">Rostellum</taxonomicName>
muscular, cushion-like 128–188 (149, n = 8) in diameter (
<figureCitation id="B1CEE7CBFFD4A564FD16FEE9FC9FFEF9" box="[754,843,267,293]" captionStart="Figs 34–38" captionStartId="33.[189,241,1911,1937]" captionTargetBox="[225,1380,276,1817]" captionTargetId="figure@33.[208,1380,265,1873]" captionTargetPageId="33" captionText="Figs 34–38. Cracticotaenia adelaidae sp. nov. 34. Scolex. 35. Detail of hooks crowns. 36. Hooks: anterior (A) and posterior (B). 37. Mature proglottis. 38. Eggs. Scale bars: 34 = 100 µm; 35 = 10 µm; 19 = 500 µm; 36, 38 = 20 µm; 37 = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3827568" httpUri="https://zenodo.org/record/3827568/files/figure.png" pageId="34" pageNumber="35">Fig. 34</figureCitation>
).
<taxonomicName id="EEF580CDFFD4A564FC85FEE9FC0EFEF9" authorityName="Montfort" authorityYear="1810" box="[865,986,267,293]" family="Rostellum" pageId="34" pageNumber="18" rank="family">Rostellum</taxonomicName>
bearing double crown of regularly arranged hooks, 52–61 (56, n = 8) in number. Hooks triangular, blade terminating with a short, curved tip. Handles and guards bearing well-developed epiphyses, more developed on guard. Epiphyses of hooks in posterior row clearly larger than those of hooks in anterior row (
<figureCitation id="B1CEE7CBFFD4A564FB94FE97FB29FE4C" box="[1136,1277,373,400]" captionStart="Figs 34–38" captionStartId="33.[189,241,1911,1937]" captionTargetBox="[225,1380,276,1817]" captionTargetId="figure@33.[208,1380,265,1873]" captionTargetPageId="33" captionText="Figs 34–38. Cracticotaenia adelaidae sp. nov. 34. Scolex. 35. Detail of hooks crowns. 36. Hooks: anterior (A) and posterior (B). 37. Mature proglottis. 38. Eggs. Scale bars: 34 = 100 µm; 35 = 10 µm; 19 = 500 µm; 36, 38 = 20 µm; 37 = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3827568" httpUri="https://zenodo.org/record/3827568/files/figure.png" pageId="34" pageNumber="35">Figs 35–36</figureCitation>
). Anterior hooks 23.5–25 (24, n = 4), posterior hooks 21–22 (21.5, n = 5) long. Neck 192–257 (216, n = 5) wide; proglottization starting shortly behind neck. Genital pores unilateral (a single side-switch observed), opening at border of first third of proglottis length margin. Ventral osmoregulatory canals up to 60 wide, connected posteriorly by anastomosis in each proglottis. Dorsal osmoregulatory canals up to 12 wide. Genital ducts passing between osmoregulatory canals. Genital atrium small. Internal organs closely packed in short and wide proglottides, their shape and relative position clearly visible only in very relaxed or extended proglottides.
</paragraph>
<caption id="7D8AABC6FFD7A567FF59F895FD1CF804" ID-DOI="http://doi.org/10.5281/zenodo.3827568" ID-Zenodo-Dep="3827568" httpUri="https://zenodo.org/record/3827568/files/figure.png" pageId="33" pageNumber="34" startId="33.[189,241,1911,1937]" targetBox="[225,1380,276,1817]" targetPageId="33">
<paragraph id="294AFB4EFFD7A567FF59F895FD1CF804" blockId="33.[189,1399,1911,2009]" pageId="33" pageNumber="34">
<emphasis id="1B81275CFFD7A567FF59F895FE99F84D" bold="true" box="[189,333,1911,1937]" pageId="33" pageNumber="34">Figs 34–38.</emphasis>
<taxonomicName id="EEF580CDFFD7A567FEBDF89AFD5CF84D" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[345,648,1911,1938]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="33" pageNumber="34" phylum="Platyhelminthes" rank="species" species="adelaidae" status="sp. nov.">
<emphasis id="1B81275CFFD7A567FEBDF89AFD5CF84D" box="[345,648,1911,1938]" italics="true" pageId="33" pageNumber="34">Cracticotaenia adelaidae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFD7A567FD70F89AFD27F84E" box="[660,755,1912,1938]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1B81275CFFD7A567FD1AF895FCCFF84D" bold="true" box="[766,795,1911,1937]" pageId="33" pageNumber="34">34</emphasis>
. Scolex.
<emphasis id="1B81275CFFD7A567FC75F895FC7AF84D" bold="true" box="[913,942,1911,1937]" pageId="33" pageNumber="34">35</emphasis>
. Detail of hooks crowns.
<emphasis id="1B81275CFFD7A567FB15F895FADAF84D" bold="true" box="[1265,1294,1911,1937]" pageId="33" pageNumber="34">36</emphasis>
. Hooks: anterior (A) and posterior (B).
<emphasis id="1B81275CFFD7A567FDC8F879FD9DF869" bold="true" box="[556,585,1947,1973]" pageId="33" pageNumber="34">37</emphasis>
. Mature proglottis.
<emphasis id="1B81275CFFD7A567FCD2F879FC87F869" bold="true" box="[822,851,1947,1973]" pageId="33" pageNumber="34">38</emphasis>
. Eggs. Scale bars: 34 = 100 µm; 35 = 10 µm; 19 = 500 µm; 36, 38 = 20 µm; 37 = 250 µm.
</paragraph>
</caption>
<paragraph id="294AFB4EFFD4A564FF59FD4FFEECFC46" blockId="34.[189,1399,685,922]" pageId="34" pageNumber="35">
Testes 12–18 (14.5, n = 38) in number, in two or, rarely, three layers, in one continuous posterior field, slightly extending laterally up to level of anterior margin of vitellarium (
<figureCitation id="B1CEE7CBFFD4A564FBEAFD32FBB1FD36" box="[1038,1125,720,746]" captionStart="Figs 34–38" captionStartId="33.[189,241,1911,1937]" captionTargetBox="[225,1380,276,1817]" captionTargetId="figure@33.[208,1380,265,1873]" captionTargetPageId="33" captionText="Figs 34–38. Cracticotaenia adelaidae sp. nov. 34. Scolex. 35. Detail of hooks crowns. 36. Hooks: anterior (A) and posterior (B). 37. Mature proglottis. 38. Eggs. Scale bars: 34 = 100 µm; 35 = 10 µm; 19 = 500 µm; 36, 38 = 20 µm; 37 = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3827568" httpUri="https://zenodo.org/record/3827568/files/figure.png" pageId="34" pageNumber="35">Fig. 37</figureCitation>
). External vas deferens discrete, convoluted, in antero-poral part of median field, at distance from cirrus-sac. Cirrus-sac short, with strong walls, most often pear-shaped, sometimes more oval, 90–118 × 43–58 (103 × 51, n = 57), not always reaching poral longitudinal osmoregulatory canals, sometimes slightly overlapping them. Internal vas deferens forming several coils. Cirrus straight with well marked walls, often appearing refractory.
</paragraph>
<paragraph id="294AFB4EFFD4A564FF59FC23FD00FB73" blockId="34.[189,1399,961,1199]" pageId="34" pageNumber="35">
Vitellarium central, compact, often heart-shaped or slightly transversely elongated, variable in shape. Ovary fanlike when not contracted, with deep lobes filling most of anterior part of proglottides. Mehlis’ gland central and anterior to vitellarium, connected to seminal receptacle by wide and often clearly visible canal that may become almost as wide as seminal receptacle. Seminal receptacle oval to fusiform, remaining rather small, reaching up to 150 × 50, dorsal and between ovary wings or over poral wing, closely to proximal extremity of cirrus-sac. Vagina opens dorsally and slightly posteriorly to male pore, mostly straight, with faint sphincter (
<figureCitation id="B1CEE7CBFFD4A564FD8BFB77FD13FB73" box="[623,711,1173,1199]" captionStart="Figs 34–38" captionStartId="33.[189,241,1911,1937]" captionTargetBox="[225,1380,276,1817]" captionTargetId="figure@33.[208,1380,265,1873]" captionTargetPageId="33" captionText="Figs 34–38. Cracticotaenia adelaidae sp. nov. 34. Scolex. 35. Detail of hooks crowns. 36. Hooks: anterior (A) and posterior (B). 37. Mature proglottis. 38. Eggs. Scale bars: 34 = 100 µm; 35 = 10 µm; 19 = 500 µm; 36, 38 = 20 µm; 37 = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3827568" httpUri="https://zenodo.org/record/3827568/files/figure.png" pageId="34" pageNumber="35">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="294AFB4EFFD4A564FF59FB35FC6BFA7C" blockId="34.[189,1399,1238,1441]" pageId="34" pageNumber="35">
Uterus starts its development in late mature proglottides as a postero-central and dorsal mass, developing into a lateraly-extended multisaccular and reticulate cavity with well marked wall, then becoming saccate. Outer shell of eggs delicate, oval 62–73 × 33–38 (68 × 35, n = 11), becoming more circular and irregular in broken proglottides. Embryophore thin and regular, oval, 30–41 (35, n = 23) in diameter. Oncosphere subspherical to slightly elongate, 30–40 (35.5, n = 18) in diameter. Embryonic hooks: central 13–15.5 long (14.5, n = 14) long, lateral 11–13 (12, n = 17) long (
<figureCitation id="B1CEE7CBFFD4A564FCBEFA64FC66FA7C" box="[858,946,1414,1440]" captionStart="Figs 34–38" captionStartId="33.[189,241,1911,1937]" captionTargetBox="[225,1380,276,1817]" captionTargetId="figure@33.[208,1380,265,1873]" captionTargetPageId="33" captionText="Figs 34–38. Cracticotaenia adelaidae sp. nov. 34. Scolex. 35. Detail of hooks crowns. 36. Hooks: anterior (A) and posterior (B). 37. Mature proglottis. 38. Eggs. Scale bars: 34 = 100 µm; 35 = 10 µm; 19 = 500 µm; 36, 38 = 20 µm; 37 = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3827568" httpUri="https://zenodo.org/record/3827568/files/figure.png" pageId="34" pageNumber="35">Fig. 38</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFD4A565FF59FA2AFEFAFCD1" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="discussion">
<paragraph id="294AFB4EFFD4A564FF59FA2AFEFBFA3E" blockId="34.[189,303,1480,1506]" box="[189,303,1480,1506]" pageId="34" pageNumber="35">
<heading id="72024C22FFD4A564FF59FA2AFEFBFA3E" bold="true" box="[189,303,1480,1506]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="1B81275CFFD4A564FF59FA2AFEFBFA3E" bold="true" box="[189,303,1480,1506]" pageId="34" pageNumber="35">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFD4A564FF59FA14FCB1F806" blockId="34.[189,1399,1526,2010]" pageId="34" pageNumber="35">
This material is very similar to
<emphasis id="1B81275CFFD4A564FDAAFA15FD25F9CD" box="[590,753,1527,1553]" italics="true" pageId="34" pageNumber="35">Cracticotania</emphasis>
<taxonomicName id="EEF580CDFFD4A564FD1AFA14FAD4F9CC" authority="(Maplestone &amp; Southwell, 1923)" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[766,1280,1526,1552]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="34" pageNumber="35" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD4A564FD1AFA14FC8AF9CC" box="[766,862,1526,1552]" italics="true" pageId="34" pageNumber="35">fieldingi</emphasis>
(
<bibRefCitation id="4D6486BFFFD4A564FC97FA14FB23F9CC" author="Maplestone P. &amp; Southwell T." box="[883,1271,1526,1552]" pageId="34" pageNumber="35" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell, 1923</bibRefCitation>
)
</taxonomicName>
, the only species know in the genus.
<emphasis id="1B81275CFFD4A564FDFEF9F8FCFEF9EF" box="[538,810,1561,1588]" italics="true" pageId="34" pageNumber="35">
Cracticotania
<taxonomicName id="EEF580CDFFD4A564FD2EF9FBFCFEF9EF" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[714,810,1561,1587]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="34" pageNumber="35" phylum="Platyhelminthes" rank="species" species="fieldingi">fieldingi</taxonomicName>
</emphasis>
was described from
<taxonomicName id="EEF580CDFFD4A564FBD1F9F8FAFAF9EF" authorityName="Temminck" authorityYear="1824" box="[1077,1326,1561,1588]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="destructor">
<emphasis id="1B81275CFFD4A564FBD1F9F8FAFAF9EF" box="[1077,1326,1561,1588]" italics="true" pageId="34" pageNumber="35">Cracticus destructor</emphasis>
</taxonomicName>
(now
<taxonomicName id="EEF580CDFFD4A564FF59F9DFFE85F98A" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[189,337,1597,1623]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="1B81275CFFD4A564FF59F9DFFE85F98A" box="[189,337,1597,1623]" italics="true" pageId="34" pageNumber="35">C. torquatus</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF580CDFFD4A564FE85F9DEFE0BF98B" authorityName="Vigors" authorityYear="1825" box="[353,479,1596,1623]" class="Aves" family="Artamidae" kingdom="Animalia" order="Passeriformes" pageId="34" pageNumber="35" phylum="Chordata" rank="family">Artamidae</taxonomicName>
) in
<collectingRegion id="EB3135ACFFD4A564FDF6F9DEFD48F98A" box="[530,668,1596,1622]" country="Australia" name="Queensland" pageId="34" pageNumber="35">Queensland</collectingRegion>
and attributed to
<emphasis id="1B81275CFFD4A564FC93F9DEFBCEF98A" box="[887,1050,1596,1622]" italics="true" pageId="34" pageNumber="35">Monopylidum</emphasis>
by
<bibRefCitation id="4D6486BFFFD4A564FBAFF9DEFEDBF9A6" author="Maplestone P. &amp; Southwell T." pageId="34" pageNumber="35" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell (1923)</bibRefCitation>
. It was later placed in a newly erected monotypic genus by
<bibRefCitation id="4D6486BFFFD4A564FC25F982FBACF9A7" author="Spasskii A. A." box="[961,1144,1632,1659]" pageId="34" pageNumber="35" pagination="1483 - 1485" refId="ref21581" refString="Spasskii A. A. 1966. On the heterogeneity of the genus Anomotaenia (Cestoda: Dilepididae). Doklady Akademii Nauk USSR 69: 1483 - 1485. [In Russian.]" type="journal article" year="1966">Spasskii (1966)</bibRefCitation>
and transferred to the
<taxonomicName id="EEF580CDFFD4A564FF59F961FEA8F941" authorityName="Spasskii" authorityYear="1959" box="[189,380,1667,1693]" class="Cestoda" family="Metadilepididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="34" pageNumber="35" phylum="Platyhelminthes" rank="family">Metadilepididae</taxonomicName>
by
<bibRefCitation id="4D6486BFFFD4A564FE43F961FCC9F941" author="Kornyushin V. V. &amp; Georgiev B. B." box="[423,797,1667,1693]" pageId="34" pageNumber="35" pagination="585 - 593" refId="ref20065" refString="Kornyushin V. V. &amp; Georgiev B. B. 1994. Family Metadilepididae Spasskii, 1959. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the cestode parasites of vertebrates: 585 - 593. CAB International, Wallingford." type="book chapter" year="1994">Kornyushin &amp; Georgiev (1994)</bibRefCitation>
. Our material fits the most recent generic diagnosis of these authors with the notable exception that only a very faint vaginal sphincter is visible. Besides this character, it differs from the single known species in the genus by a smaller number of rostellar hooks (52–61 vs about 80), a slightly fewer testes (12–18 vs 16–21) and the differently-shaped and shorter cirrus-sac (globular vs pipette-like and 90–118 vs 130). It should, however, be noted that we examined the
<typeStatus id="F64E45ECFFD4A564FF0CF8D1FECFF891" box="[232,283,1843,1869]" pageId="34" pageNumber="35">type</typeStatus>
material of
<taxonomicName id="EEF580CDFFD4A564FE4EF8D1FDF9F891" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[426,557,1843,1869]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="34" pageNumber="35" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD4A564FE4EF8D1FDF9F891" box="[426,557,1843,1869]" italics="true" pageId="34" pageNumber="35">C. fieldingi</emphasis>
</taxonomicName>
and the measurements provided by
<bibRefCitation id="4D6486BFFFD4A564FC32F8D1FA86F891" author="Maplestone P. &amp; Southwell T." box="[982,1362,1843,1869]" pageId="34" pageNumber="35" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell (1923)</bibRefCitation>
do not fully correspond to our observations. Although only two slides, mostly with gravid proglottides, are available, we noted a somewhat longer cirrus-sac (up to 180) and fewer testes (12–15). This, however, does not prevent the unambiguous separation of our material from
<taxonomicName id="EEF580CDFFD4A564FC2BF87FFB86F86A" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[975,1106,1948,1975]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="34" pageNumber="35" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFD4A564FC2BF87FFB86F86A" box="[975,1106,1948,1975]" italics="true" pageId="34" pageNumber="35">C. fieldingi</emphasis>
</taxonomicName>
and we place it in a new species which we name
<taxonomicName id="EEF580CDFFD4A564FE3CF822FCD0F806" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[472,772,1984,2010]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="34" pageNumber="35" phylum="Platyhelminthes" rank="species" species="adelaidae" status="sp. nov.">
<emphasis id="1B81275CFFD4A564FE3CF822FCD0F806" box="[472,772,1984,2010]" italics="true" pageId="34" pageNumber="35">Cracticotaenia adelaidae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFD4A564FCEFF822FCB1F806" box="[779,869,1984,2010]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="294AFB4EFFD5A565FF59FEE9FD30FE24" blockId="35.[189,1399,267,505]" pageId="35" pageNumber="36">
The specimens found in
<taxonomicName id="EEF580CDFFD5A565FE07FEEEFD2DFEF9" authority="Lesson, 1831" authorityName="Lesson" authorityYear="1831" box="[483,761,267,294]" class="Aves" family="Corcoracidae" genus="Corcorax" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
<emphasis id="1B81275CFFD5A565FE07FEEEFD86FEFA" box="[483,594,268,294]" italics="true" pageId="35" pageNumber="36">Corcorax</emphasis>
Lesson, 1831
</taxonomicName>
fit the above description of
<taxonomicName id="EEF580CDFFD5A565FBAEFEEEFAA3FEF9" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[1098,1399,267,294]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="species" species="adelaidae" status="sp. nov.">
<emphasis id="1B81275CFFD5A565FBAEFEEEFAA3FEF9" box="[1098,1399,267,294]" italics="true" pageId="35" pageNumber="36">Cracticotaenia adelaidae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFD5A565FF59FECDFECCFE95" box="[189,280,303,329]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
in
<taxonomicName id="EEF580CDFFD5A565FED9FECDFDFCFE95" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[317,552,303,329]" class="Aves" family="Cracticidae" genus="Gymnorhina" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="tibicen">
<emphasis id="1B81275CFFD5A565FED9FECDFDFCFE95" box="[317,552,303,329]" italics="true" pageId="35" pageNumber="36">Gymnorhina tibicen</emphasis>
</taxonomicName>
quite well for most metric and meristic characters that can be observed. The main observed differences are in slightly smaller suckers (123–147 vs 155–198) and possibly more rostellar hooks (62–66 vs 52–61), and the faint vaginal sphincter that could not be observed. Keeping in mind that these observations are based on a limited number of incomplete specimens, we consider for now that these small variations (which may be partially exaggerated due to the poor preservation state of our specimens) are likely to be intraspecific.
</paragraph>
<paragraph id="294AFB4EFFD5A565FF59FDC2FEFAFCD1" blockId="35.[189,1399,544,781]" pageId="35" pageNumber="36">
Although both belonging to the Corvides (see
<bibRefCitation id="4D6486BFFFD5A565FD0DFDC2FC07FDE7" author="Jonsson K. A. &amp; Fabre P. - H. &amp; Kennedy J. D. &amp; Holt B. G. &amp; Borregaard M. K. &amp; Rahbek C. &amp; Fjeldsa J." box="[745,979,544,571]" pageId="35" pageNumber="36" pagination="87 - 94" refId="ref19843" refString="Jonsson K. A., Fabre P. - H., Kennedy J. D., Holt B. G., Borregaard M. K., Rahbek C. &amp; Fjeldsa J. 2016. A supermatrix phylogeny of corvoid passerine birds (Aves: Corvides). Molecular Phylogenetics and Evolution 94: 87 - 94. https: // doi. org / 10.1016 / j. ympev. 2015.08.020" type="journal article" year="2016">
Jønsson
<emphasis id="1B81275CFFD5A565FCABFDC3FC5FFDE6" box="[847,907,544,570]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2016
</bibRefCitation>
), the hosts of
<taxonomicName id="EEF580CDFFD5A565FB9EFDC2FAC7FDE6" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[1146,1299,544,570]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="35" pageNumber="36" phylum="Platyhelminthes" rank="species" species="adelaidae" status="sp. nov.">
<emphasis id="1B81275CFFD5A565FB9EFDC2FAC7FDE6" box="[1146,1299,544,570]" italics="true" pageId="35" pageNumber="36">C. adelaidae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFD5A565FAFFFDC2FAA3FDE6" box="[1307,1399,544,570]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
are not closely related and belong to clades that have probably diverged some 20 to 30 Ma ago (
<bibRefCitation id="4D6486BFFFD5A565FF21FD84FE63FD5C" author="Jonsson K. A. &amp; Fabre P. - H. &amp; Kennedy J. D. &amp; Holt B. G. &amp; Borregaard M. K. &amp; Rahbek C. &amp; Fjeldsa J." box="[197,439,614,640]" pageId="35" pageNumber="36" pagination="87 - 94" refId="ref19843" refString="Jonsson K. A., Fabre P. - H., Kennedy J. D., Holt B. G., Borregaard M. K., Rahbek C. &amp; Fjeldsa J. 2016. A supermatrix phylogeny of corvoid passerine birds (Aves: Corvides). Molecular Phylogenetics and Evolution 94: 87 - 94. https: // doi. org / 10.1016 / j. ympev. 2015.08.020" type="journal article" year="2016">
Jønsson
<emphasis id="1B81275CFFD5A565FECBFD85FEBAFD5C" box="[303,366,614,640]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2016
</bibRefCitation>
;
<bibRefCitation id="4D6486BFFFD5A565FE21FD84FD71FD5D" author="Moyle R. G. &amp; Oliveros C. H. &amp; Andersen M. J. &amp; Hosner P. A. &amp; Benz B. W. &amp; Manthey J. D. &amp; Travers S. L. &amp; Brown R. M. &amp; Faircloth B. C." box="[453,677,614,641]" pageId="35" pageNumber="36" pagination="1 - 7" refId="ref20827" refString="Moyle R. G., Oliveros C. H., Andersen M. J., Hosner P. A., Benz B. W., Manthey J. D., Travers S. L., Brown R. M. &amp; Faircloth B. C. 2016. Tectonic collision and uplift of Wallacea triggered the global songbird radiation. Nature Communications 7: 1 - 7. https: // doi. org / 10.1038 / ncomms 12709" type="journal article" year="2016">
<collectingRegion id="EB3135ACFFD5A565FE21FD84FDC7FD5C" box="[453,531,614,640]" country="United Kingdom" name="Moyle" pageId="35" pageNumber="36">Moyle</collectingRegion>
<emphasis id="1B81275CFFD5A565FDF9FD85FD8FFD5C" box="[541,603,614,640]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2016
</bibRefCitation>
). The Australian magpie (
<taxonomicName id="EEF580CDFFD5A565FC00FD85FB88FD5C" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[996,1116,614,641]" class="Aves" family="Cracticidae" genus="Gymnorhina" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="tibicen">
<emphasis id="1B81275CFFD5A565FC00FD85FB88FD5C" box="[996,1116,614,641]" italics="true" pageId="35" pageNumber="36">G. tibicen</emphasis>
</taxonomicName>
) and the white winged chough (
<taxonomicName id="EEF580CDFFD5A565FECDFD68FDC9FD78" baseAuthorityName="Vieillot" baseAuthorityYear="1817" box="[297,541,650,676]" class="Aves" family="Corcoracidae" genus="Corcorax" kingdom="Animalia" order="Passeriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="melanorhamphos">
<emphasis id="1B81275CFFD5A565FECDFD68FDC9FD78" box="[297,541,650,676]" italics="true" pageId="35" pageNumber="36">C. melanorhamphos</emphasis>
</taxonomicName>
) have however similar diets and habitats (
<bibRefCitation id="4D6486BFFFD5A565FBC2FD6BFADBFD78" author="Russell E. &amp; Rowley I. &amp; Christie D. A." box="[1062,1295,649,676]" pageId="35" pageNumber="36" refId="ref21100" refString="Russell E., Rowley I. &amp; Christie D. A. 2017. Australian Magpie (Gymnorhina tibicen) and Whitewinged Chough (Corcorax melanorhamphos). In: del Hoyo J., Elliott A., Sargatal J., Christie D. A. &amp; de Juana E. (eds.) Handbook of the Birds of the World Alive. Lynx Edicions, Barcelona. Available from http: // www. hbw. com / node / 60621 [accessed 15 Sep. 2017]." type="url" year="2017">
Russell
<emphasis id="1B81275CFFD5A565FB6EFD68FB14FD78" box="[1162,1216,650,676]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2017
</bibRefCitation>
) as well as overlapping ranges (hosts reported in this study were captured a mere
<quantity id="EE0D56ABFFD5A565FBCDFD4FFB57FD1B" box="[1065,1155,685,711]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="36" unit="km" value="100.0">100 km</quantity>
apart). This relative phylogenetic proximity and quite similar ecology probably largely explain the hosting of shared parasites.
</paragraph>
</subSubSection>
</treatment>
</document>

876
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<subSubSection id="61EFA8C5FFE3A553FDA9F9EBFB68F99B" pageId="21" pageNumber="22" type="nomenclature">
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<emphasis id="1B81275CFFE3A553FDA9F9EBFC50F9F8" bold="true" box="[589,900,1545,1572]" italics="true" pageId="21" pageNumber="22">Dictymetra longiuncinata</emphasis>
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<paragraph id="294AFB4EFFE3A553FD3CF9ADFC8FF9B5" blockId="21.[375,1212,1545,1641]" box="[728,859,1615,1641]" pageId="21" pageNumber="22">
<figureCitation id="B1CEE7CBFFE3A553FD3CF9ADFC8FF9B5" box="[728,859,1615,1641]" captionStart="Figs 23–28" captionStartId="25.[189,241,1915,1941]" captionTargetBox="[210,1386,286,1854]" captionTargetId="figure@25.[197,1390,265,1890]" captionTargetPageId="25" captionText="Figs 23–28. Dictymetra longiuncinata sp. nov. 23. Scolex. 24. Hooks. 25. Mature proglottis. 26. Terminal genitalia. 27. Cirrus armament. 28. Eggs. Scale bars: 23 = 100 µm; 24, 26 = 50 µm; 25 = 200 µm; 27 = 10 µm; 28 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827564" httpUri="https://zenodo.org/record/3827564/files/figure.png" pageId="21" pageNumber="22">Figs 23–28</figureCitation>
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</subSubSection>
<subSubSection id="61EFA8C5FFE3A553FF59F973FA83F905" box="[189,1367,1681,1754]" pageId="21" pageNumber="22" type="etymology">
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<emphasis id="1B81275CFFE3A553FF59F973FE90F977" bold="true" box="[189,324,1681,1707]" pageId="21" pageNumber="22">Etymology</emphasis>
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From Latin
<emphasis id="1B81275CFFE3A553FEADF95DFE43F905" box="[329,407,1727,1753]" italics="true" pageId="21" pageNumber="22">longus</emphasis>
(long) and
<emphasis id="1B81275CFFE3A553FDFBF922FDAEF905" box="[543,634,1728,1753]" italics="true" pageId="21" pageNumber="22">uncinus</emphasis>
(hook) in reference to the large rostellar hooks of this species.
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<emphasis id="1B81275CFFE3A553FF59F8E3FE7EF8C7" bold="true" box="[189,426,1793,1819]" pageId="21" pageNumber="22">Material examined</emphasis>
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</paragraph>
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:
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,
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,
<geoCoordinate id="4CC19D89FFE3A553FD3CF8B3FC98F8B1" box="[728,844,1873,1901]" degrees="150" direction="east" minutes="44" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="150.73334">150°44′ E</geoCoordinate>
,
<date id="5D4BDD8EFFE3A553FCB3F8B1FC3AF8B1" box="[855,1006,1875,1901]" pageId="21" pageNumber="22" value="1999-09-30">
<collectingDate id="4D0F2466FFE3A553FCB3F8B1FC3AF8B1" box="[855,1006,1875,1901]" pageId="21" pageNumber="22" value="1999-09-30">30 Sep. 1999</collectingDate>
</date>
,
<collectorName id="84009E98FFE3A553FC1CF8B1FB4EF8B0" box="[1016,1178,1874,1901]" pageId="21" pageNumber="22">Ian Beveridge</collectorName>
leg. (AHC 36496).
</paragraph>
</materialsCitation>
<paragraph id="294AFB4EFFE3A553FF0EF876FEB3F872" blockId="21.[189,1123,1940,2002]" box="[234,359,1940,1966]" pageId="21" pageNumber="22">
<emphasis id="1B81275CFFE3A553FF0EF876FEB3F872" bold="true" box="[234,359,1940,1966]" pageId="21" pageNumber="22">
<typeStatus id="F64E45ECFFE3A553FF0EF876FEB3F872" box="[234,359,1940,1966]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFE3A553FF59F855FBB7F80E" blockId="21.[189,1123,1940,2002]" box="[189,1123,1975,2002]" pageId="21" pageNumber="22">
<collectingCountry id="51E2BBDEFFE3A553FF59F855FEBCF80D" box="[189,360,1975,2001]" name="Australia" pageId="21" pageNumber="22">AUSTRALIA</collectingCountry>
: about 44 specs, same data as for
<typeStatus id="F64E45ECFFE3A553FD17F855FC8EF80D" box="[755,858,1975,2001]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
(AHC 36497–36515).
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFE0A550FF59FEE9FDB8FDEF" pageId="22" pageNumber="23" type="biology_ecology">
<paragraph id="294AFB4EFFE0A550FF59FEE9FF2CFEF9" blockId="22.[189,248,267,293]" box="[189,248,267,293]" pageId="22" pageNumber="23">
<heading id="72024C22FFE0A550FF59FEE9FF2CFEF9" bold="true" box="[189,248,267,293]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="1B81275CFFE0A550FF59FEE9FF2CFEF9" bold="true" box="[189,248,267,293]" pageId="22" pageNumber="23">Host</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFE0A550FF59FED8FAE2FE88" blockId="22.[189,1334,313,340]" box="[189,1334,313,340]" pageId="22" pageNumber="23">
<taxonomicName id="EEF580CDFFE0A550FF59FED8FDBCFE88" authority="(Vieillot, 1818)" baseAuthorityName="Vieillot" baseAuthorityYear="1818" box="[189,616,313,340]" class="Aves" family="Burhinidae" genus="Esacus" kingdom="Animalia" order="Charadriiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="magnirostris">
<emphasis id="1B81275CFFE0A550FF59FED8FE78FE8F" box="[189,428,314,339]" italics="true" pageId="22" pageNumber="23">Esacus magnirostris</emphasis>
(Vieillot, 1818)
</taxonomicName>
(syn.
<taxonomicName id="EEF580CDFFE0A550FD55FEDBFC6CFE8F" baseAuthorityName="Vieillot" baseAuthorityYear="1818" box="[689,952,313,339]" class="Aves" family="Burhinidae" genus="Burhinus" kingdom="Animalia" order="Charadriiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="magnirostris">
<emphasis id="1B81275CFFE0A550FD55FEDBFC6CFE8F" box="[689,952,313,339]" italics="true" pageId="22" pageNumber="23">Burhinus magnirostris</emphasis>
</taxonomicName>
) (
<taxonomicName id="EEF580CDFFE0A550FC35FEDBFB42FE8F" box="[977,1174,313,339]" class="Aves" kingdom="Animalia" order="Charadriiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="order">Charadriiformes</taxonomicName>
,
<taxonomicName id="EEF580CDFFE0A550FB46FEDBFAFEFE88" baseAuthorityName="Bona" baseAuthorityYear="1994" box="[1186,1322,313,340]" class="Aves" family="Burhinidae" kingdom="Animalia" order="Charadriiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="family">Burhinidae</taxonomicName>
).
</paragraph>
<paragraph id="294AFB4EFFE0A550FF59FE99FE54FE49" blockId="22.[189,384,379,405]" box="[189,384,379,405]" pageId="22" pageNumber="23">
<heading id="72024C22FFE0A550FF59FE99FE54FE49" bold="true" box="[189,384,379,405]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="1B81275CFFE0A550FF59FE99FE54FE49" bold="true" box="[189,384,379,405]" pageId="22" pageNumber="23">Site of infection</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFE0A550FF59FE4BFEA0FE1F" blockId="22.[189,372,425,451]" box="[189,372,425,451]" pageId="22" pageNumber="23">Small intestine.</paragraph>
<paragraph id="294AFB4EFFE0A550FF59FE09FEF8FDD9" blockId="22.[189,300,491,517]" box="[189,300,491,517]" pageId="22" pageNumber="23">
<heading id="72024C22FFE0A550FF59FE09FEF8FDD9" bold="true" box="[189,300,491,517]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="1B81275CFFE0A550FF59FE09FEF8FDD9" bold="true" box="[189,300,491,517]" pageId="22" pageNumber="23">Intensity</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFE0A550FF59FDFBFDB8FDEF" blockId="22.[189,620,537,564]" box="[189,620,537,564]" pageId="22" pageNumber="23">
About
<specimenCount id="3FF330C7FFE0A550FEEAFDFBFE78FDEF" box="[270,428,537,564]" pageId="22" pageNumber="23" type="generic">45 specimens</specimenCount>
in a single host.
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFE0A55CFF59FDB9FDBBFE94" lastPageId="26" lastPageNumber="27" pageId="22" pageNumber="23" type="description">
<paragraph id="294AFB4EFFE0A550FF59FDB9FE9AFDA9" blockId="22.[189,334,603,629]" box="[189,334,603,629]" pageId="22" pageNumber="23">
<heading id="72024C22FFE0A550FF59FDB9FE9AFDA9" bold="true" box="[189,334,603,629]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="1B81275CFFE0A550FF59FDB9FE9AFDA9" bold="true" box="[189,334,603,629]" pageId="22" pageNumber="23">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFE0A550FF59FD6BFE60FB26" blockId="22.[189,1399,649,1274]" pageId="22" pageNumber="23">
Body small,
<quantity id="EE0D56ABFFE0A550FEB7FD68FE2AFD78" box="[339,510,650,676]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.39" metricValueMin="1.01" pageId="22" pageNumber="23" unit="mm" value="12.0" valueMax="13.9" valueMin="10.1">10.1–13.9 mm</quantity>
(n = 2) long with maximum width 950–1400 (1144, n = 8) at level of early gravid proglottides. Most complete specimen consisting of 76–83 proglottides. Proglottides markedly craspedote, wider than long, progressively becoming longer, but mostly remaining wider than long except for a few terminal or exceptionally extended ones. Osmoregulatory canals only very partially visible. Scolex rounded, in continuity with strobila (without a marked neck); diameter 470–665 (551, n = 7) (
<figureCitation id="B1CEE7CBFFE0A550FF22FCDBFEF4FC8F" box="[198,288,825,851]" captionStart="Figs 23–28" captionStartId="25.[189,241,1915,1941]" captionTargetBox="[210,1386,286,1854]" captionTargetId="figure@25.[197,1390,265,1890]" captionTargetPageId="25" captionText="Figs 23–28. Dictymetra longiuncinata sp. nov. 23. Scolex. 24. Hooks. 25. Mature proglottis. 26. Terminal genitalia. 27. Cirrus armament. 28. Eggs. Scale bars: 23 = 100 µm; 24, 26 = 50 µm; 25 = 200 µm; 27 = 10 µm; 28 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827564" httpUri="https://zenodo.org/record/3827564/files/figure.png" pageId="22" pageNumber="23">Fig. 23</figureCitation>
). Proglottization starting precociously, often just behind suckers. Suckers rounded to slightly oval, muscular, unarmed, 230–300 (262, n = 32) in diameter. Rostellar apparatus mostly muscular, with a layer of glandular tissue at mid-length of rostellar sac. Rostellar sac well-delineated, with cylindrical anterior part and markedly tapering posterior part, extending well beyond level of posterior margin of suckers and sometimes into first proglottides, 475–690 × 150–210 (608 × 178, n = 7).
<taxonomicName id="EEF580CDFFE0A550FB7CFC24FAC5FC3C" authorityName="Montfort" authorityYear="1810" box="[1176,1297,966,992]" family="Rostellum" pageId="22" pageNumber="18" rank="family">Rostellum</taxonomicName>
strongly muscular, with expanded anterior part bearing rostellar hooks, 400–620 × 110–145 (537 × 133, n = 8). Rostellar hooks (
<figureCitation id="B1CEE7CBFFE0A550FE6CFBEEFE34FBFB" box="[392,480,1036,1063]" captionStart="Figs 23–28" captionStartId="25.[189,241,1915,1941]" captionTargetBox="[210,1386,286,1854]" captionTargetId="figure@25.[197,1390,265,1890]" captionTargetPageId="25" captionText="Figs 23–28. Dictymetra longiuncinata sp. nov. 23. Scolex. 24. Hooks. 25. Mature proglottis. 26. Terminal genitalia. 27. Cirrus armament. 28. Eggs. Scale bars: 23 = 100 µm; 24, 26 = 50 µm; 25 = 200 µm; 27 = 10 µm; 28 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827564" httpUri="https://zenodo.org/record/3827564/files/figure.png" pageId="22" pageNumber="23">Fig. 24</figureCitation>
) in two regular rows, 20 (n = 6) in number, long and robust, curved overall; blade longer than handle. Anterior hooks 119–125 (121.5, n = 13) long, posterior hooks 124–135 (129.5, n = 15) long. Genital pores situated at border of anterior 40% of lateral proglottis margin, alternating irregularly in very short series, e.g., 1, 1, 1, 1, 1, 1, 2, 1, 2, 1, 1, 1, 1 or 1, 1, 2, 1, 1, 3, 1, 1, 1, 1; no more than 3 consecutive pores observed on one side. Genital ducts passing between osmoregulatory canals. Genital atrium as simple depression without particular structures. Genital papilla occasionally expressed in some proglottides.
</paragraph>
<paragraph id="294AFB4EFFE0A550FF59FAC0FC33F9ED" blockId="22.[189,1399,1313,1586]" pageId="22" pageNumber="23">
Testes numerous, 18–28 (22, n = 19) in number, large, reaching 75–92 (84, n = 19) in diameter, situated in 2–3 layers, in one continuous and compact posterior field, extending in anterior direction to level of antiporal wing of ovary (
<figureCitation id="B1CEE7CBFFE0A550FDEEFA85FDB0FA5E" box="[522,612,1383,1410]" captionStart="Figs 23–28" captionStartId="25.[189,241,1915,1941]" captionTargetBox="[210,1386,286,1854]" captionTargetId="figure@25.[197,1390,265,1890]" captionTargetPageId="25" captionText="Figs 23–28. Dictymetra longiuncinata sp. nov. 23. Scolex. 24. Hooks. 25. Mature proglottis. 26. Terminal genitalia. 27. Cirrus armament. 28. Eggs. Scale bars: 23 = 100 µm; 24, 26 = 50 µm; 25 = 200 µm; 27 = 10 µm; 28 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827564" httpUri="https://zenodo.org/record/3827564/files/figure.png" pageId="22" pageNumber="23">Fig. 25</figureCitation>
). External vas deferens
<quantity id="EE0D56ABFFE0A550FC60FA8AFC39FA5D" box="[900,1005,1383,1410]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.572" metricValueMin="3.302" pageId="22" pageNumber="23" unit="in" value="15.5" valueMax="18.0" valueMin="13.0">13–18 in</quantity>
diameter, highly convoluted and particularly voluminous in antero-poral part of median field, sometimes slightly overlapping cirrussac, seminal receptacle or even testes. Cirrus-sac elongate, 137–180 × 33–45 (159 × 38, n = 44), delicate, slightly oblique, usually straight but with distal quarter sometimes bent anteriorly, crossing osmoregulatory canals, rarely not. Internal vas deferens forming several coils. Cirrus cylindrical,
<quantity id="EE0D56ABFFE0A550FAD0FA16FAA3F9D2" box="[1332,1399,1524,1551]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" pageId="22" pageNumber="23" unit="in" value="7.5">7.5 in</quantity>
diameter, ornamented with tuft of thin hair-like bristles (
<figureCitation id="B1CEE7CBFFE0A550FCB6F9F5FC0CF9EE" box="[850,984,1559,1586]" captionStart="Figs 23–28" captionStartId="25.[189,241,1915,1941]" captionTargetBox="[210,1386,286,1854]" captionTargetId="figure@25.[197,1390,265,1890]" captionTargetPageId="25" captionText="Figs 23–28. Dictymetra longiuncinata sp. nov. 23. Scolex. 24. Hooks. 25. Mature proglottis. 26. Terminal genitalia. 27. Cirrus armament. 28. Eggs. Scale bars: 23 = 100 µm; 24, 26 = 50 µm; 25 = 200 µm; 27 = 10 µm; 28 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827564" httpUri="https://zenodo.org/record/3827564/files/figure.png" pageId="22" pageNumber="23">Figs 26–27</figureCitation>
).
</paragraph>
<paragraph id="294AFB4EFFE0A550FF59F9BBFAA3F8DC" blockId="22.[189,1399,1625,1792]" pageId="22" pageNumber="23">Vitellarium central, compact, irregularly shaped and slightly lobated. Ovary transversely elongated, bi-alate, lobate, antiporal wing larger than poral wing; the latter mostly covered by vas deferens. Mehlis’ gland not observed. Empty seminal receptacle round, becoming oval when filled, reaching up to 135 × 68, close to proximal extremity of cirrus sac. Vagina, large, opened posteriorly to male pore, mostly straight and transverse, parallel to cirrus-sac, thin-walled. No sphincters or other ornamentations.</paragraph>
<paragraph id="294AFB4EFFE0A550FF59F8CAFAA3F877" blockId="22.[189,1399,1831,1964]" pageId="22" pageNumber="23">
Uterus starts its development in late mature proglottides as diffuse reticulum occupying the whole median field and part of lateral fields, and developing into dense network of tube and chambers. Eggs large, outer shell with polar extensions (
<figureCitation id="B1CEE7CBFFE0A550FD45F88CFD2FF854" box="[673,763,1902,1928]" captionStart="Figs 23–28" captionStartId="25.[189,241,1915,1941]" captionTargetBox="[210,1386,286,1854]" captionTargetId="figure@25.[197,1390,265,1890]" captionTargetPageId="25" captionText="Figs 23–28. Dictymetra longiuncinata sp. nov. 23. Scolex. 24. Hooks. 25. Mature proglottis. 26. Terminal genitalia. 27. Cirrus armament. 28. Eggs. Scale bars: 23 = 100 µm; 24, 26 = 50 µm; 25 = 200 µm; 27 = 10 µm; 28 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827564" httpUri="https://zenodo.org/record/3827564/files/figure.png" pageId="22" pageNumber="23">Fig. 28</figureCitation>
). Complete eggs with external membrane difficult to see
<emphasis id="1B81275CFFE0A550FF0FF870FE93F877" box="[235,327,1938,1963]" italics="true" pageId="22" pageNumber="23">in utero</emphasis>
but reaching at least 85 (without extensions). Embryophores oval, 53–65 (59.5, n = 15) in
</paragraph>
<caption id="7D8AABC6FFE1A551FF58FDC2F9CFFD15" ID-Table-UUID="7D8AABC6FFE1A551FF58FDC2F9CFFD15" httpUri="http://table.plazi.org/id/7D8AABC6FFE1A551FF58FDC2F9CFFD15" pageId="23" pageNumber="24" targetBox="[192,1945,762,1736]" targetIsTable="true" targetPageId="23">
<paragraph id="294AFB4EFFE1A551FF58FDC2F9CFFD15" blockId="23.[188,1942,544,713]" pageId="23" pageNumber="24">
<emphasis id="1B81275CFFE1A551FF58FDC2FEC3FDE0" bold="true" box="[188,279,544,572]" pageId="23" pageNumber="24">Table 4</emphasis>
(continued on next page). Comparison of two new
<taxonomicName id="EEF580CDFFE1A551FC89FDC2FB56FDE0" authority="Clark, 1952" authorityName="Clark" authorityYear="1952" box="[877,1154,544,572]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE1A551FC89FDC2FC25FDE0" box="[877,1009,544,572]" italics="true" pageId="23" pageNumber="24">Dictymetra</emphasis>
<bibRefCitation id="4D6486BFFFE1A551FC1CFDC2FB56FDE0" author="Clark D. T." box="[1016,1154,544,572]" pageId="23" pageNumber="24" pagination="18 - 27" refId="ref18941" refString="Clark D. T. 1952. Three new dilepidid cestodes, Dictymetra numenii n. gen. sp. nov.; Dictymetra paranumenii sp. nov. and Anomotaenia filovata sp. nov. Proceedings of the Helminthological Society of Washington 19: 18 - 27." type="journal article" year="1952">Clark, 1952</bibRefCitation>
</taxonomicName>
species with the presently recognized species in the genus and with “long”
<taxonomicName id="EEF580CDFFE1A551FEF6FDA1FDDDFD83" authority="Weinland, 1857" authorityName="Weinland" authorityYear="1857" box="[274,521,579,607]" class="Cestoda" family="Dilepididae" genus="Liga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE1A551FEF6FDA1FE9CFD83" box="[274,328,579,607]" italics="true" pageId="23" pageNumber="24">Liga</emphasis>
Weinland, 1857
</taxonomicName>
species (see text). All data as reported in
<bibRefCitation id="4D6486BFFFE1A551FC12FDA1FA9AFD83" author="Spasskaya L. P. &amp; Spasskii A." box="[1014,1358,579,607]" pageId="23" pageNumber="24" refId="ref21548" refString="Spasskaya L. P. &amp; Spasskii A. 1978. Cestodes of Birds in the USSR. Dilepididae of Aquatic Birds. Izdate'stvo Nauka, Moscow. [In Russian.]" type="book" year="1978">Spasskaya &amp; Spasskii (1978)</bibRefCitation>
for
<taxonomicName id="EEF580CDFFE1A551FA64FDA1F9D0FD83" authorityName="Clark" authorityYear="1952" box="[1408,1540,579,607]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE1A551FA64FDA1F9D0FD83" box="[1408,1540,579,607]" italics="true" pageId="23" pageNumber="24">Dictymetra</emphasis>
</taxonomicName>
and
<bibRefCitation id="4D6486BFFFE1A551F9DAFDA1F842FD83" author="Spasskaya L. P. &amp; Spasskii A." box="[1598,1942,579,607]" pageId="23" pageNumber="24" refId="ref21515" refString="Spasskaya L. P. &amp; Spasskii A. 1977. Cestodes of Birds in the USSR. Dilepididae of Terrestrial Birds. Izdate'stvo Nauka, Moscow. [In Russian.]" type="book" year="1977">Spasskaya &amp; Spasskii (1977)</bibRefCitation>
for
<taxonomicName id="EEF580CDFFE1A551FF02FD84FEC8FD5E" authorityName="Weinland" authorityYear="1857" box="[230,284,614,642]" class="Cestoda" family="Dilepididae" genus="Liga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE1A551FF02FD84FEC8FD5E" box="[230,284,614,642]" italics="true" pageId="23" pageNumber="24">Liga</emphasis>
</taxonomicName>
except
<bibRefCitation id="4D6486BFFFE1A551FE9CFD84FDD6FD5E" author="Burt D. R. R." box="[376,514,614,642]" pageId="23" pageNumber="24" pagination="1 - 63" refId="ref18802" refString="Burt D. R. R. 1940. New species of cestodes from Charadriiformes, Ardeiformes and Pelecaniformes in Ceylon. Ceylon Journal of Science. Section B. Zoology and Geology 22: 1 - 63." type="journal article" year="1940">Burt (1940)</bibRefCitation>
for
<taxonomicName id="EEF580CDFFE1A551FDD0FD84FC9DFD5E" authority=", Clark (1952)" authorityName="Clark" authorityYear="1952" box="[564,841,614,642]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="dispar">
<emphasis id="1B81275CFFE1A551FDD0FD84FD77FD5E" box="[564,675,614,642]" italics="true" pageId="23" pageNumber="24">D. dispar</emphasis>
,
<bibRefCitation id="4D6486BFFFE1A551FD56FD84FC9DFD5E" author="Clark D. T." box="[690,841,614,642]" pageId="23" pageNumber="24" pagination="18 - 27" refId="ref18941" refString="Clark D. T. 1952. Three new dilepidid cestodes, Dictymetra numenii n. gen. sp. nov.; Dictymetra paranumenii sp. nov. and Anomotaenia filovata sp. nov. Proceedings of the Helminthological Society of Washington 19: 18 - 27." type="journal article" year="1952">Clark (1952)</bibRefCitation>
</taxonomicName>
for
<taxonomicName id="EEF580CDFFE1A551FC9FFD84FAA4FD5E" authority=", Fuhrmann &amp; Baer (1943)" authorityName=", Fuhrmann &amp; Baer" authorityYear="1943" box="[891,1392,614,642]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="paranumenii">
<emphasis id="1B81275CFFE1A551FC9FFD84FBE1FD5E" box="[891,1077,614,642]" italics="true" pageId="23" pageNumber="24">D. paranumenii</emphasis>
,
<bibRefCitation id="4D6486BFFFE1A551FBA0FD84FAA4FD5E" author="Fuhrmann O. &amp; Baer J. G." box="[1092,1392,614,642]" pageId="23" pageNumber="24" pagination="113 - 140" refId="ref19370" refString="Fuhrmann O. &amp; Baer J. G. 1943. Mission biologique Sagan-Omo (Ethiopie meridionale), 1939, dirigee par le professeur Eduardo Zavattari. Cestodes. Bulletin de la Societe neuchateloise de Sciences naturelles 68: 113 - 140." type="journal article" year="1943">Fuhrmann &amp; Baer (1943)</bibRefCitation>
</taxonomicName>
for
<taxonomicName id="EEF580CDFFE1A551FA46FD84F9D0FD5E" baseAuthorityName="Fuhrmann &amp; Baer" baseAuthorityYear="1943" box="[1442,1540,614,642]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="riccii">
<emphasis id="1B81275CFFE1A551FA46FD84F9D0FD5E" box="[1442,1540,614,642]" italics="true" pageId="23" pageNumber="24">D. riccii</emphasis>
</taxonomicName>
and
<bibRefCitation id="4D6486BFFFE1A551F9DAFD84F842FD5E" author="Spasskaya L. P. &amp; Spasskii A." box="[1598,1942,614,642]" pageId="23" pageNumber="24" pagination="63 - 68" refId="ref21462" refString="Spasskaya L. P. &amp; Spasskii A. 1973. Dictymetra belopolskaje, sp. n. (Cestoda: Dilepididae) a new species of cestode in birds. Izvestiya Akademii Nauk Moldovskoi SSR (Biologicheskie i Khimicheskie Nauki) 4: 63 - 68. [In Russian.]" type="journal article" year="1973">Spasskaya &amp; Spasskii (1973)</bibRefCitation>
for
<taxonomicName id="EEF580CDFFE1A551FF02FD68FEB8FD7A" baseAuthorityName="von Linstow" baseAuthorityYear="1906" box="[230,364,650,678]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="volvulus">
<emphasis id="1B81275CFFE1A551FF02FD68FEB8FD7A" box="[230,364,650,678]" italics="true" pageId="23" pageNumber="24">D. volvulus</emphasis>
</taxonomicName>
. Note that values reported by various authors for
<taxonomicName id="EEF580CDFFE1A551FC57FD68FAC2FD7A" authority="(Rudolphi, 1919)" baseAuthorityName="Rudolphi" baseAuthorityYear="1919" box="[947,1302,650,678]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="laevigata">
<emphasis id="1B81275CFFE1A551FC57FD68FB90FD7A" box="[947,1092,650,678]" italics="true" pageId="23" pageNumber="24">D. laevigata</emphasis>
(Rudolphi, 1919)
</taxonomicName>
and
<taxonomicName id="EEF580CDFFE1A551FAABFD68F943FD7A" authority="(Linstow, 1903)" baseAuthorityName="Linstow" baseAuthorityYear="1903" box="[1359,1687,650,678]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="volvulus">
<emphasis id="1B81275CFFE1A551FAABFD68FA01FD7A" box="[1359,1493,650,678]" italics="true" pageId="23" pageNumber="24">D. volvulus</emphasis>
(Linstow, 1903)
</taxonomicName>
vary to a large extent (
<bibRefCitation id="4D6486BFFFE1A551FF20FD4FFDC2FD15" author="Spasskaya L. P. &amp; Spasskii A." box="[196,534,685,713]" pageId="23" pageNumber="24" refId="ref21548" refString="Spasskaya L. P. &amp; Spasskii A. 1978. Cestodes of Birds in the USSR. Dilepididae of Aquatic Birds. Izdate'stvo Nauka, Moscow. [In Russian.]" type="book" year="1978">Spasskaya &amp; Spasskii 1978</bibRefCitation>
).
<taxonomicName id="EEF580CDFFE1A551FDCEFD4FFC98FD15" authorityName=", Fuhrmann &amp; Baer" authorityYear="1943" box="[554,844,685,713]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="paranumenii">
<emphasis id="1B81275CFFE1A551FDCEFD4FFC98FD15" box="[554,844,685,713]" italics="true" pageId="23" pageNumber="24">Dictymetra paranumenii</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF580CDFFE1A551FC61FD4FFB95FD15" authorityName="Matevosyan" authorityYear="1953" box="[901,1089,685,713]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="radiaspinosa">
<emphasis id="1B81275CFFE1A551FC61FD4FFB95FD15" box="[901,1089,685,713]" italics="true" pageId="23" pageNumber="24">D. radiaspinosa</emphasis>
</taxonomicName>
have rows of small spines on
<taxonomicName id="EEF580CDFFE1A551FA40FD4FF9C3FD15" authorityName="Montfort" authorityYear="1810" box="[1444,1559,685,713]" family="Rostellum" pageId="23" pageNumber="18" rank="family">rostellum</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="294AFB4EFFE1A551FF24FCF7F842F914" pageId="23" pageNumber="24">
<table id="5BF509EEFFE15AB9FF24FD18F84DF914" box="[192,1945,762,1736]" gridcols="13" gridrows="12" pageId="23" pageNumber="24">
<tr id="97C5F90CFFE15AB9FF24FD18F84DFC9B" box="[192,1945,762,839]" gridrow="0" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FD18FE19FC9B" box="[192,461,762,839]" gridcol="0" gridrow="0" pageId="23" pageNumber="24">Species</th>
<th id="D4149070FFE15AB9FE38FD18FDADFC9B" box="[476,633,762,839]" gridcol="1" gridrow="0" pageId="23" pageNumber="24">Host family</th>
<th id="D4149070FFE15AB9FD71FD18FD2AFC9B" box="[661,766,762,839]" gridcol="2" gridrow="0" pageId="23" pageNumber="24">Type locality</th>
<th id="D4149070FFE15AB9FCF9FD18FC4FFC9B" box="[797,923,762,839]" gridcol="3" gridrow="0" pageId="23" pageNumber="24">Body maximum length (mm)</th>
<th id="D4149070FFE15AB9FC57FD18FBDBFC9B" box="[947,1039,762,839]" gridcol="4" gridrow="0" pageId="23" pageNumber="24">Scolex diameter</th>
<th id="D4149070FFE15AB9FBC2FD18FB56FC9B" box="[1062,1154,762,839]" gridcol="5" gridrow="0" pageId="23" pageNumber="24">Suckers diameter</th>
<th id="D4149070FFE15AB9FB7DFD18FAD5FC9B" box="[1177,1281,762,839]" gridcol="6" gridrow="0" pageId="23" pageNumber="24">Rostellar sac</th>
<th id="D4149070FFE15AB9FAF3FD18FAABFC9B" box="[1303,1407,762,839]" gridcol="7" gridrow="0" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FAF3FCF7FAABFCF0" authorityName="Montfort" authorityYear="1810" box="[1303,1407,789,812]" family="Rostellum" pageId="23" pageNumber="18" rank="family">Rostellum</taxonomicName>
</th>
<th id="D4149070FFE15AB9FA71FD18FA32FC9B" box="[1429,1510,762,839]" gridcol="8" gridrow="0" pageId="23" pageNumber="24">Hooks number</th>
<th id="D4149070FFE15AB9FA19FD18F982FC9B" box="[1533,1622,762,839]" gridcol="9" gridrow="0" pageId="23" pageNumber="24">Anterior hooks length</th>
<th id="D4149070FFE15AB9F989FD18F91FFC9B" box="[1645,1739,762,839]" gridcol="10" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="1B81275CFFE1A551F989FD18F91FFCCD" bold="true" box="[1645,1739,762,785]" pageId="23" pageNumber="24">Posterior</emphasis>
hooks
<emphasis id="1B81275CFFE1A551F998FCD2F968FC9B" bold="true" box="[1660,1724,816,839]" pageId="23" pageNumber="24">length</emphasis>
</th>
<th id="D4149070FFE15AB9F905FD18F8FBFC9B" box="[1761,1839,762,839]" gridcol="11" gridrow="0" pageId="23" pageNumber="24">Testes number</th>
<th id="D4149070FFE15AB9F8A1FD18F84DFC9B" box="[1861,1945,762,839]" gridcol="12" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="1B81275CFFE1A551F8A9FCEAF845FCC3" bold="true" box="[1869,1937,776,799]" pageId="23" pageNumber="24">Cirrus</emphasis>
<emphasis id="1B81275CFFE1A551F8BBFCC0F8ABFCE5" bold="true" box="[1887,1919,802,825]" pageId="23" pageNumber="24">sac</emphasis>
</th>
</tr>
<tr id="97C5F90CFFE15AB9FF24FC87F84DFC40" box="[192,1945,869,924]" gridrow="1" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FC87FE19FC40" box="[192,461,869,924]" gridcol="0" gridrow="1" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24FC97FEE3FC50" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[192,311,885,908]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="gerganae" status="sp. nov.">
<emphasis id="1B81275CFFE1A551FF24FC97FEE3FC50" box="[192,311,885,908]" italics="true" pageId="23" pageNumber="24">D. gerganae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFE1A551FED9FC97FE5CFC50" box="[317,392,885,908]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="D4149070FFE15AB9FE38FC87FDADFC40" box="[476,633,869,924]" gridcol="1" gridrow="1" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FE11FC97FDB5FC50" box="[501,609,885,908]" class="Aves" family="Podargidae" kingdom="Animalia" order="Strigiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Podargidae</taxonomicName>
</td>
<td id="D4149070FFE15AB9FD71FC87FD2AFC40" box="[661,766,869,924]" gridcol="2" gridrow="1" pageId="23" pageNumber="24">
<collectingCountry id="51E2BBDEFFE1A551FD7AFC97FD22FC50" box="[670,758,885,908]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
</td>
<td id="D4149070FFE15AB9FCF9FC87FC4FFC40" box="[797,923,869,924]" gridcol="3" gridrow="1" pageId="23" pageNumber="24">72</td>
<td id="D4149070FFE15AB9FC57FC87FBDBFC40" box="[947,1039,869,924]" gridcol="4" gridrow="1" pageId="23" pageNumber="24">305–360</td>
<td id="D4149070FFE15AB9FBC2FC87FB56FC40" box="[1062,1154,869,924]" gridcol="5" gridrow="1" pageId="23" pageNumber="24">145–175</td>
<td id="D4149070FFE15AB9FB7DFC87FAD5FC40" box="[1177,1281,869,924]" gridcol="6" gridrow="1" pageId="23" pageNumber="24">230–292 × 118–145</td>
<td id="D4149070FFE15AB9FAF3FC87FAABFC40" box="[1303,1407,869,924]" gridcol="7" gridrow="1" pageId="23" pageNumber="24">195–212 × 98–132</td>
<td id="D4149070FFE15AB9FA71FC87FA32FC40" box="[1429,1510,869,924]" gridcol="8" gridrow="1" pageId="23" pageNumber="24">24</td>
<td id="D4149070FFE15AB9FA19FC87F982FC40" box="[1533,1622,869,924]" gridcol="9" gridrow="1" pageId="23" pageNumber="24">46–53</td>
<td id="D4149070FFE15AB9F989FC87F91FFC40" box="[1645,1739,869,924]" gridcol="10" gridrow="1" pageId="23" pageNumber="24">50–56</td>
<td id="D4149070FFE15AB9F905FC87F8FBFC40" box="[1761,1839,869,924]" gridcol="11" gridrow="1" pageId="23" pageNumber="24">60–77</td>
<td id="D4149070FFE15AB9F8A1FC87F84DFC40" box="[1861,1945,869,924]" gridcol="12" gridrow="1" pageId="23" pageNumber="24">100–132 × 65–83</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FC56F84DFC37" box="[192,1945,948,1003]" gridrow="2" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FC56FE19FC37" box="[192,461,948,1003]" gridcol="0" gridrow="2" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24FC26FEB6FC07" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[192,354,964,987]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="longiuncinata" status="sp. nov.">
<emphasis id="1B81275CFFE1A551FF24FC26FEB6FC07" box="[192,354,964,987]" italics="true" pageId="23" pageNumber="24">D. longiuncinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFE1A551FE8DFC26FE67FC07" box="[361,435,964,987]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="D4149070FFE15AB9FE38FC56FDADFC37" box="[476,633,948,1003]" gridcol="1" gridrow="2" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FE11FC26FDB4FC07" baseAuthorityName="Bona" baseAuthorityYear="1994" box="[501,608,964,987]" class="Aves" family="Burhinidae" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Burhinidae</taxonomicName>
</td>
<td id="D4149070FFE15AB9FD71FC56FD2AFC37" box="[661,766,948,1003]" gridcol="2" gridrow="2" pageId="23" pageNumber="24">
<collectingCountry id="51E2BBDEFFE1A551FD7AFC26FD22FC07" box="[670,758,964,987]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
</td>
<td id="D4149070FFE15AB9FCF9FC56FC4FFC37" box="[797,923,948,1003]" gridcol="3" gridrow="2" pageId="23" pageNumber="24">13.2</td>
<td id="D4149070FFE15AB9FC57FC56FBDBFC37" box="[947,1039,948,1003]" gridcol="4" gridrow="2" pageId="23" pageNumber="24">470–665</td>
<td id="D4149070FFE15AB9FBC2FC56FB56FC37" box="[1062,1154,948,1003]" gridcol="5" gridrow="2" pageId="23" pageNumber="24">230–300</td>
<td id="D4149070FFE15AB9FB7DFC56FAD5FC37" box="[1177,1281,948,1003]" gridcol="6" gridrow="2" pageId="23" pageNumber="24">475–690 × 150–210</td>
<td id="D4149070FFE15AB9FAF3FC56FAABFC37" box="[1303,1407,948,1003]" gridcol="7" gridrow="2" pageId="23" pageNumber="24">400–620 × 110–145</td>
<td id="D4149070FFE15AB9FA71FC56FA32FC37" box="[1429,1510,948,1003]" gridcol="8" gridrow="2" pageId="23" pageNumber="24">20</td>
<td id="D4149070FFE15AB9FA19FC56F982FC37" box="[1533,1622,948,1003]" gridcol="9" gridrow="2" pageId="23" pageNumber="24">119–125</td>
<td id="D4149070FFE15AB9F989FC56F91FFC37" box="[1645,1739,948,1003]" gridcol="10" gridrow="2" pageId="23" pageNumber="24">124–135</td>
<td id="D4149070FFE15AB9F905FC56F8FBFC37" box="[1761,1839,948,1003]" gridcol="11" gridrow="2" pageId="23" pageNumber="24">18–28</td>
<td id="D4149070FFE15AB9F8A1FC56F84DFC37" box="[1861,1945,948,1003]" gridcol="12" gridrow="2" pageId="23" pageNumber="24">137–180 × 33–45</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FBE0F84DFBE6" box="[192,1945,1026,1082]" gridrow="3" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FBE0FE19FBE6" box="[192,461,1026,1082]" gridcol="0" gridrow="3" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24FBE1FE55FBE6" authority="Mathevossian, 1953" authorityName="Mathevossian" authorityYear="1953" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="radiaspinosa">
<emphasis id="1B81275CFFE1A551FF24FBE1FE8EFBC6" box="[192,346,1027,1050]" italics="true" pageId="23" pageNumber="24">D. radiaspinosa</emphasis>
Mathevossian, 1953
</taxonomicName>
</th>
<td id="D4149070FFE15AB9FE38FBE0FDADFBE6" box="[476,633,1026,1082]" gridcol="1" gridrow="3" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FE0FFBF0FDBFFBF5" box="[491,619,1042,1065]" class="Aves" family="Scolopacidae" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Scolopacidae</taxonomicName>
</td>
<td id="D4149070FFE15AB9FD71FBE0FD2AFBE6" box="[661,766,1026,1082]" gridcol="2" gridrow="3" pageId="23" pageNumber="24">
<collectingCountry id="51E2BBDEFFE1A551FD56FBF0FD36FBF5" box="[690,738,1042,1065]" name="United States of America" pageId="23" pageNumber="24">USA</collectingCountry>
</td>
<td id="D4149070FFE15AB9FCF9FBE0FC4FFBE6" box="[797,923,1026,1082]" gridcol="3" gridrow="3" pageId="23" pageNumber="24">15–25</td>
<td id="D4149070FFE15AB9FC57FBE0FBDBFBE6" box="[947,1039,1026,1082]" gridcol="4" gridrow="3" pageId="23" pageNumber="24">247–330</td>
<td id="D4149070FFE15AB9FBC2FBE0FB56FBE6" box="[1062,1154,1026,1082]" gridcol="5" gridrow="3" pageId="23" pageNumber="24">204–231</td>
<td id="D4149070FFE15AB9FB7DFBE0FAD5FBE6" box="[1177,1281,1026,1082]" gridcol="6" gridrow="3" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9FAF3FBE0FAABFBE6" box="[1303,1407,1026,1082]" gridcol="7" gridrow="3" pageId="23" pageNumber="24">83–115</td>
<td id="D4149070FFE15AB9FA71FBE0FA32FBE6" box="[1429,1510,1026,1082]" gridcol="8" gridrow="3" pageId="23" pageNumber="24">24</td>
<td id="D4149070FFE15AB9FA19FBE0F982FBE6" box="[1533,1622,1026,1082]" gridcol="9" gridrow="3" pageId="23" pageNumber="24">65–68</td>
<td id="D4149070FFE15AB9F989FBE0F91FFBE6" box="[1645,1739,1026,1082]" gridcol="10" gridrow="3" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9F905FBE0F8FBFBE6" box="[1761,1839,1026,1082]" gridcol="11" gridrow="3" pageId="23" pageNumber="24">15–17</td>
<td id="D4149070FFE15AB9F8A1FBE0F84DFBE6" box="[1861,1945,1026,1082]" gridcol="12" gridrow="3" pageId="23" pageNumber="24">146–198 × 23–25</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FBB3F84DFB54" box="[192,1945,1105,1160]" gridrow="4" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FBB3FE19FB54" box="[192,461,1105,1160]" gridcol="0" gridrow="4" pageId="23" pageNumber="24">
<emphasis id="1B81275CFFE1A551FF24FBB3FEB6FBB4" box="[192,354,1105,1128]" italics="true" pageId="23" pageNumber="24">D. belopolskajae</emphasis>
Spasskaya &amp;
<bibRefCitation id="4D6486BFFFE1A551FF3DFB93FEB1FB54" author="Spasskii A. A. &amp; Borgarenko L. F." box="[217,357,1137,1160]" pageId="23" pageNumber="24" pagination="87 - 93" refId="ref21661" refString="Spasskii A. A. &amp; Borgarenko L. F. 1973. Stenovaria n. g. (Cestoda: Dilepididae) - a new genus of cestodes from the stone curlew. Izvestiya Akademii Nauk Tadzhikskoi SSR, Biologicheskie Nauk 2: 87 - 93. [In Russian.]" type="journal article" year="1973">Spasskii, 1973</bibRefCitation>
</th>
<td id="D4149070FFE15AB9FE38FBB3FDADFB54" box="[476,633,1105,1160]" gridcol="1" gridrow="4" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FE1DFB83FD88FBA4" box="[505,604,1121,1144]" class="Aves" family="Alaudidae" kingdom="Animalia" order="Passeriformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Alaudidae</taxonomicName>
</td>
<td id="D4149070FFE15AB9FD71FBB3FD2AFB54" box="[661,766,1105,1160]" gridcol="2" gridrow="4" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FD47FBB3FD3BFBB4" authority="Spasskaya Russian" authorityName="Spasskaya Russian" box="[675,751,1105,1128]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="belopolskajae">Russian</taxonomicName>
Arctic
</td>
<td id="D4149070FFE15AB9FCF9FBB3FC4FFB54" box="[797,923,1105,1160]" gridcol="3" gridrow="4" pageId="23" pageNumber="24">80</td>
<td id="D4149070FFE15AB9FC57FBB3FBDBFB54" box="[947,1039,1105,1160]" gridcol="4" gridrow="4" pageId="23" pageNumber="24">313</td>
<td id="D4149070FFE15AB9FBC2FBB3FB56FB54" box="[1062,1154,1105,1160]" gridcol="5" gridrow="4" pageId="23" pageNumber="24">123</td>
<td id="D4149070FFE15AB9FB7DFBB3FAD5FB54" box="[1177,1281,1105,1160]" gridcol="6" gridrow="4" pageId="23" pageNumber="24">347–408 × 106–123</td>
<td id="D4149070FFE15AB9FAF3FBB3FAABFB54" box="[1303,1407,1105,1160]" gridcol="7" gridrow="4" pageId="23" pageNumber="24">252–308 × 95–112</td>
<td id="D4149070FFE15AB9FA71FBB3FA32FB54" box="[1429,1510,1105,1160]" gridcol="8" gridrow="4" pageId="23" pageNumber="24">20</td>
<td id="D4149070FFE15AB9FA19FBB3F982FB54" box="[1533,1622,1105,1160]" gridcol="9" gridrow="4" pageId="23" pageNumber="24">56–67</td>
<td id="D4149070FFE15AB9F989FBB3F91FFB54" box="[1645,1739,1105,1160]" gridcol="10" gridrow="4" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9F905FBB3F8FBFB54" box="[1761,1839,1105,1160]" gridcol="11" gridrow="4" pageId="23" pageNumber="24">18–30</td>
<td id="D4149070FFE15AB9F8A1FBB3F84DFB54" box="[1861,1945,1105,1160]" gridcol="12" gridrow="4" pageId="23" pageNumber="24">106 × 39</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FB42F84DFB0B" box="[192,1945,1184,1239]" gridrow="5" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FB42FE19FB0B" box="[192,461,1184,1239]" gridcol="0" gridrow="5" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24FB42FF2CFB0B" authority="(Beneden, 1868)" authorityName="Spasskaya &amp; Shumilo" authorityYear="1971" baseAuthorityName="Beneden" baseAuthorityYear="1868" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="discoidea">
<emphasis id="1B81275CFFE1A551FF24FB42FEEDFB6B" box="[192,313,1184,1207]" italics="true" pageId="23" pageNumber="24">D. discoidea</emphasis>
(Beneden, 1868)
</taxonomicName>
</th>
<td id="D4149070FFE15AB9FE38FB42FDADFB0B" box="[476,633,1184,1239]" gridcol="1" gridrow="5" pageId="23" pageNumber="24">Ciconiidae</td>
<td id="D4149070FFE15AB9FD71FB42FD2AFB0B" box="[661,766,1184,1239]" gridcol="2" gridrow="5" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FD71FB52FD2AFB1B" authority="Palaearctic" authorityName="Palaearctic" box="[661,766,1200,1223]" class="Aves" family="Ciconiidae" kingdom="Animalia" order="Ciconiiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Palaearctic</taxonomicName>
</td>
<td id="D4149070FFE15AB9FCF9FB42FC4FFB0B" box="[797,923,1184,1239]" gridcol="3" gridrow="5" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9FC57FB42FBDBFB0B" box="[947,1039,1184,1239]" gridcol="4" gridrow="5" pageId="23" pageNumber="24">370–400</td>
<td id="D4149070FFE15AB9FBC2FB42FB56FB0B" box="[1062,1154,1184,1239]" gridcol="5" gridrow="5" pageId="23" pageNumber="24">210</td>
<td id="D4149070FFE15AB9FB7DFB42FAD5FB0B" box="[1177,1281,1184,1239]" gridcol="6" gridrow="5" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9FAF3FB42FAABFB0B" box="[1303,1407,1184,1239]" gridcol="7" gridrow="5" pageId="23" pageNumber="24">100</td>
<td id="D4149070FFE15AB9FA71FB42FA32FB0B" box="[1429,1510,1184,1239]" gridcol="8" gridrow="5" pageId="23" pageNumber="24">22–23</td>
<td id="D4149070FFE15AB9FA19FB42F982FB0B" box="[1533,1622,1184,1239]" gridcol="9" gridrow="5" pageId="23" pageNumber="24">37–38</td>
<td id="D4149070FFE15AB9F989FB42F91FFB0B" box="[1645,1739,1184,1239]" gridcol="10" gridrow="5" pageId="23" pageNumber="24">33–34</td>
<td id="D4149070FFE15AB9F905FB42F8FBFB0B" box="[1761,1839,1184,1239]" gridcol="11" gridrow="5" pageId="23" pageNumber="24">40–45</td>
<td id="D4149070FFE15AB9F8A1FB42F84DFB0B" box="[1861,1945,1184,1239]" gridcol="12" gridrow="5" pageId="23" pageNumber="24">250–270 × 65–70</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FB18F84DFAED" box="[192,1945,1274,1329]" gridrow="6" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FB18FE19FAED" box="[192,461,1274,1329]" gridcol="0" gridrow="6" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24FAE8FE43FAFD" authority="(Burt, 1940)" baseAuthorityName="Burt" baseAuthorityYear="1940" box="[192,407,1290,1313]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="dispar">
<emphasis id="1B81275CFFE1A551FF24FAE8FECEFAFD" box="[192,282,1290,1313]" italics="true" pageId="23" pageNumber="24">D. dispar</emphasis>
(
<bibRefCitation id="4D6486BFFFE1A551FEC3FAE8FE44FAFD" author="Burt D. R. R." box="[295,400,1290,1313]" pageId="23" pageNumber="24" pagination="1 - 63" refId="ref18802" refString="Burt D. R. R. 1940. New species of cestodes from Charadriiformes, Ardeiformes and Pelecaniformes in Ceylon. Ceylon Journal of Science. Section B. Zoology and Geology 22: 1 - 63." type="journal article" year="1940">Burt, 1940</bibRefCitation>
)
</taxonomicName>
</th>
<td id="D4149070FFE15AB9FE38FB18FDADFAED" box="[476,633,1274,1329]" gridcol="1" gridrow="6" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FE08FAE8FDBCFAFD" box="[492,616,1290,1313]" class="Aves" family="Charadriidae" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Charadriidae</taxonomicName>
</td>
<td id="D4149070FFE15AB9FD71FB18FD2AFAED" box="[661,766,1274,1329]" gridcol="2" gridrow="6" pageId="23" pageNumber="24">
<collectingCountry id="51E2BBDEFFE1A551FD7EFAE8FD2CFAFD" box="[666,760,1290,1313]" name="Sri Lanka" pageId="23" pageNumber="24">Sri Lanka</collectingCountry>
</td>
<td id="D4149070FFE15AB9FCF9FB18FC4FFAED" box="[797,923,1274,1329]" gridcol="3" gridrow="6" pageId="23" pageNumber="24">40–50</td>
<td id="D4149070FFE15AB9FC57FB18FBDBFAED" box="[947,1039,1274,1329]" gridcol="4" gridrow="6" pageId="23" pageNumber="24">330–385</td>
<td id="D4149070FFE15AB9FBC2FB18FB56FAED" box="[1062,1154,1274,1329]" gridcol="5" gridrow="6" pageId="23" pageNumber="24">143–175</td>
<td id="D4149070FFE15AB9FB7DFB18FAD5FAED" box="[1177,1281,1274,1329]" gridcol="6" gridrow="6" pageId="23" pageNumber="24">325 × 110</td>
<td id="D4149070FFE15AB9FAF3FB18FAABFAED" box="[1303,1407,1274,1329]" gridcol="7" gridrow="6" pageId="23" pageNumber="24">175–229 × 65–88</td>
<td id="D4149070FFE15AB9FA71FB18FA32FAED" box="[1429,1510,1274,1329]" gridcol="8" gridrow="6" pageId="23" pageNumber="24">24</td>
<td id="D4149070FFE15AB9FA19FB18F982FAED" box="[1533,1622,1274,1329]" gridcol="9" gridrow="6" pageId="23" pageNumber="24">41–45</td>
<td id="D4149070FFE15AB9F989FB18F91FFAED" box="[1645,1739,1274,1329]" gridcol="10" gridrow="6" pageId="23" pageNumber="24">44–49</td>
<td id="D4149070FFE15AB9F905FB18F8FBFAED" box="[1761,1839,1274,1329]" gridcol="11" gridrow="6" pageId="23" pageNumber="24">18–28</td>
<td id="D4149070FFE15AB9F8A1FB18F84DFAED" box="[1861,1945,1274,1329]" gridcol="12" gridrow="6" pageId="23" pageNumber="24">170–200 × 38–41</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FAB6F84DFA57" box="[192,1945,1364,1419]" gridrow="7" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FAB6FE19FA57" box="[192,461,1364,1419]" gridcol="0" gridrow="7" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24FAB6FEADFAB7" authority="Spasskaya" authorityName="Spasskaya" box="[192,377,1364,1387]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="ganii">
<emphasis id="1B81275CFFE1A551FF24FAB6FEDAFAB7" box="[192,270,1364,1387]" italics="true" pageId="23" pageNumber="24">D. ganii</emphasis>
Spasskaya
</taxonomicName>
&amp; Shumilo, 1971
</th>
<td id="D4149070FFE15AB9FE38FAB6FDADFA57" box="[476,633,1364,1419]" gridcol="1" gridrow="7" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FE0EFA86FDBEFAA7" box="[490,618,1380,1403]" class="Aves" family="Scolopacidae" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Scolopacidae</taxonomicName>
</td>
<td id="D4149070FFE15AB9FD71FAB6FD2AFA57" box="[661,766,1364,1419]" gridcol="2" gridrow="7" pageId="23" pageNumber="24">
<collectingCountry id="51E2BBDEFFE1A551FD7FFA86FD23FAA7" box="[667,759,1380,1403]" name="Moldova" pageId="23" pageNumber="24">Moldavia</collectingCountry>
</td>
<td id="D4149070FFE15AB9FCF9FAB6FC4FFA57" box="[797,923,1364,1419]" gridcol="3" gridrow="7" pageId="23" pageNumber="24">70</td>
<td id="D4149070FFE15AB9FC57FAB6FBDBFA57" box="[947,1039,1364,1419]" gridcol="4" gridrow="7" pageId="23" pageNumber="24">695</td>
<td id="D4149070FFE15AB9FBC2FAB6FB56FA57" box="[1062,1154,1364,1419]" gridcol="5" gridrow="7" pageId="23" pageNumber="24">335</td>
<td id="D4149070FFE15AB9FB7DFAB6FAD5FA57" box="[1177,1281,1364,1419]" gridcol="6" gridrow="7" pageId="23" pageNumber="24">246 × 140</td>
<td id="D4149070FFE15AB9FAF3FAB6FAABFA57" box="[1303,1407,1364,1419]" gridcol="7" gridrow="7" pageId="23" pageNumber="24">162 × 67</td>
<td id="D4149070FFE15AB9FA71FAB6FA32FA57" box="[1429,1510,1364,1419]" gridcol="8" gridrow="7" pageId="23" pageNumber="24">22–24</td>
<td id="D4149070FFE15AB9FA19FAB6F982FA57" box="[1533,1622,1364,1419]" gridcol="9" gridrow="7" pageId="23" pageNumber="24">28</td>
<td id="D4149070FFE15AB9F989FAB6F91FFA57" box="[1645,1739,1364,1419]" gridcol="10" gridrow="7" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9F905FAB6F8FBFA57" box="[1761,1839,1364,1419]" gridcol="11" gridrow="7" pageId="23" pageNumber="24">(~40)</td>
<td id="D4149070FFE15AB9F8A1FAB6F84DFA57" box="[1861,1945,1364,1419]" gridcol="12" gridrow="7" pageId="23" pageNumber="24">94 × 67</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FA41F84DFA06" box="[192,1945,1443,1498]" gridrow="8" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FA41FE19FA06" box="[192,461,1443,1498]" gridcol="0" gridrow="8" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24FA41FF2CFA06" authority="(Rudolphi, 1819)" baseAuthorityName="Rudolphi" baseAuthorityYear="1819" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="laevigata">
<emphasis id="1B81275CFFE1A551FF24FA41FEE2FA66" box="[192,310,1443,1466]" italics="true" pageId="23" pageNumber="24">D. laevigata</emphasis>
(Rudolphi, 1819)
</taxonomicName>
</th>
<td id="D4149070FFE15AB9FE38FA41FDADFA06" box="[476,633,1443,1498]" gridcol="1" gridrow="8" pageId="23" pageNumber="24">Charadriiformes</td>
<td id="D4149070FFE15AB9FD71FA41FD2AFA06" box="[661,766,1443,1498]" gridcol="2" gridrow="8" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FD71FA51FD2AFA16" authority="Palaearctic" authorityName="Palaearctic" box="[661,766,1459,1482]" class="Aves" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="order">Palaearctic</taxonomicName>
</td>
<td id="D4149070FFE15AB9FCF9FA41FC4FFA06" box="[797,923,1443,1498]" gridcol="3" gridrow="8" pageId="23" pageNumber="24">75</td>
<td id="D4149070FFE15AB9FC57FA41FBDBFA06" box="[947,1039,1443,1498]" gridcol="4" gridrow="8" pageId="23" pageNumber="24">650</td>
<td id="D4149070FFE15AB9FBC2FA41FB56FA06" box="[1062,1154,1443,1498]" gridcol="5" gridrow="8" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9FB7DFA41FAD5FA06" box="[1177,1281,1443,1498]" gridcol="6" gridrow="8" pageId="23" pageNumber="24">428</td>
<td id="D4149070FFE15AB9FAF3FA41FAABFA06" box="[1303,1407,1443,1498]" gridcol="7" gridrow="8" pageId="23" pageNumber="24">314 × 141</td>
<td id="D4149070FFE15AB9FA71FA41FA32FA06" box="[1429,1510,1443,1498]" gridcol="8" gridrow="8" pageId="23" pageNumber="24">20</td>
<td id="D4149070FFE15AB9FA19FA41F982FA06" box="[1533,1622,1443,1498]" gridcol="9" gridrow="8" pageId="23" pageNumber="24">104–106</td>
<td id="D4149070FFE15AB9F989FA41F91FFA06" box="[1645,1739,1443,1498]" gridcol="10" gridrow="8" pageId="23" pageNumber="24">109–112</td>
<td id="D4149070FFE15AB9F905FA41F8FBFA06" box="[1761,1839,1443,1498]" gridcol="11" gridrow="8" pageId="23" pageNumber="24">50</td>
<td id="D4149070FFE15AB9F8A1FA41F84DFA06" box="[1861,1945,1443,1498]" gridcol="12" gridrow="8" pageId="23" pageNumber="24">98 × 48</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24FA10F84DF9F5" box="[192,1945,1522,1577]" gridrow="9" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24FA10FE19F9F5" box="[192,461,1522,1577]" gridcol="0" gridrow="9" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24F9E0FE62F9C5" authority="(Owen, 1946)" baseAuthorityName="Owen" baseAuthorityYear="1946" box="[192,438,1538,1561]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="numenii">
<emphasis id="1B81275CFFE1A551FF24F9E0FEFEF9C5" box="[192,298,1538,1561]" italics="true" pageId="23" pageNumber="24">D. numenii</emphasis>
(Owen, 1946)
</taxonomicName>
</th>
<td id="D4149070FFE15AB9FE38FA10FDADF9F5" box="[476,633,1522,1577]" gridcol="1" gridrow="9" pageId="23" pageNumber="24">Scolopacidae</td>
<td id="D4149070FFE15AB9FD71FA10FD2AF9F5" box="[661,766,1522,1577]" gridcol="2" gridrow="9" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FD79F9E0FD22F9C5" authority="Holarctic" authorityName="Holarctic" box="[669,758,1538,1561]" class="Aves" family="Scolopacidae" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Holarctic</taxonomicName>
</td>
<td id="D4149070FFE15AB9FCF9FA10FC4FF9F5" box="[797,923,1522,1577]" gridcol="3" gridrow="9" pageId="23" pageNumber="24">14–20</td>
<td id="D4149070FFE15AB9FC57FA10FBDBF9F5" box="[947,1039,1522,1577]" gridcol="4" gridrow="9" pageId="23" pageNumber="24">192–538</td>
<td id="D4149070FFE15AB9FBC2FA10FB56F9F5" box="[1062,1154,1522,1577]" gridcol="5" gridrow="9" pageId="23" pageNumber="24">133–221</td>
<td id="D4149070FFE15AB9FB7DFA10FAD5F9F5" box="[1177,1281,1522,1577]" gridcol="6" gridrow="9" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9FAF3FA10FAABF9F5" box="[1303,1407,1522,1577]" gridcol="7" gridrow="9" pageId="23" pageNumber="24">124–162</td>
<td id="D4149070FFE15AB9FA71FA10FA32F9F5" box="[1429,1510,1522,1577]" gridcol="8" gridrow="9" pageId="23" pageNumber="24">22</td>
<td id="D4149070FFE15AB9FA19FA10F982F9F5" box="[1533,1622,1522,1577]" gridcol="9" gridrow="9" pageId="23" pageNumber="24">99–103</td>
<td id="D4149070FFE15AB9F989FA10F91FF9F5" box="[1645,1739,1522,1577]" gridcol="10" gridrow="9" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9F905FA10F8FBF9F5" box="[1761,1839,1522,1577]" gridcol="11" gridrow="9" pageId="23" pageNumber="24">42–46</td>
<td id="D4149070FFE15AB9F8A1FA10F84DF9F5" box="[1861,1945,1522,1577]" gridcol="12" gridrow="9" pageId="23" pageNumber="24">111–140 × 44–58</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24F9A3F84DF9A4" box="[192,1945,1601,1656]" gridrow="10" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24F9A3FE19F9A4" box="[192,461,1601,1656]" gridcol="0" gridrow="10" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24F9A3FF2CF9A4" authority="(Schrank, 1790)" authorityName="Spassky &amp; Spasskaya" authorityYear="1971" baseAuthorityName="Schrank" baseAuthorityYear="1790" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="nymphaea">
<emphasis id="1B81275CFFE1A551FF24F9A3FE94F984" box="[192,320,1601,1624]" italics="true" pageId="23" pageNumber="24">D. nymphaea</emphasis>
(Schrank, 1790)
</taxonomicName>
</th>
<td id="D4149070FFE15AB9FE38F9A3FDADF9A4" box="[476,633,1601,1656]" gridcol="1" gridrow="10" pageId="23" pageNumber="24">Charadriiformes</td>
<td id="D4149070FFE15AB9FD71F9A3FD2AF9A4" box="[661,766,1601,1656]" gridcol="2" gridrow="10" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FD71F9B3FD2AF9B4" authority="Palaearctic" authorityName="Palaearctic" box="[661,766,1617,1640]" class="Aves" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="order">Palaearctic</taxonomicName>
</td>
<td id="D4149070FFE15AB9FCF9F9A3FC4FF9A4" box="[797,923,1601,1656]" gridcol="3" gridrow="10" pageId="23" pageNumber="24">25–30</td>
<td id="D4149070FFE15AB9FC57F9A3FBDBF9A4" box="[947,1039,1601,1656]" gridcol="4" gridrow="10" pageId="23" pageNumber="24">210–220</td>
<td id="D4149070FFE15AB9FBC2F9A3FB56F9A4" box="[1062,1154,1601,1656]" gridcol="5" gridrow="10" pageId="23" pageNumber="24">80–85</td>
<td id="D4149070FFE15AB9FB7DF9A3FAD5F9A4" box="[1177,1281,1601,1656]" gridcol="6" gridrow="10" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9FAF3F9A3FAABF9A4" box="[1303,1407,1601,1656]" gridcol="7" gridrow="10" pageId="23" pageNumber="24">80–85</td>
<td id="D4149070FFE15AB9FA71F9A3FA32F9A4" box="[1429,1510,1601,1656]" gridcol="8" gridrow="10" pageId="23" pageNumber="24">24</td>
<td id="D4149070FFE15AB9FA19F9A3F982F9A4" box="[1533,1622,1601,1656]" gridcol="9" gridrow="10" pageId="23" pageNumber="24">60–80</td>
<td id="D4149070FFE15AB9F989F9A3F91FF9A4" box="[1645,1739,1601,1656]" gridcol="10" gridrow="10" pageId="23" pageNumber="24">–</td>
<td id="D4149070FFE15AB9F905F9A3F8FBF9A4" box="[1761,1839,1601,1656]" gridcol="11" gridrow="10" pageId="23" pageNumber="24">30–36</td>
<td id="D4149070FFE15AB9F8A1F9A3F84DF9A4" box="[1861,1945,1601,1656]" gridcol="12" gridrow="10" pageId="23" pageNumber="24">100–130 × 38–55</td>
</tr>
<tr id="97C5F90CFFE15AB9FF24F973F84DF914" box="[192,1945,1681,1736]" gridrow="11" pageId="23" pageNumber="24">
<th id="D4149070FFE15AB9FF24F973FE19F914" box="[192,461,1681,1736]" gridcol="0" gridrow="11" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FF24F973FF2CF914" authority="(Clark, 1952)" authorityName=", Fuhrmann &amp; Baer" authorityYear="1943" baseAuthorityName="Clark" baseAuthorityYear="1952" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="paranumenii">
<emphasis id="1B81275CFFE1A551FF24F973FE8CF974" box="[192,344,1681,1704]" italics="true" pageId="23" pageNumber="24">D. paranumenii</emphasis>
(Clark, 1952)
</taxonomicName>
</th>
<td id="D4149070FFE15AB9FE38F973FDADF914" box="[476,633,1681,1736]" gridcol="1" gridrow="11" pageId="23" pageNumber="24">
<taxonomicName id="EEF580CDFFE1A551FE0EF943FDBEF964" box="[490,618,1697,1720]" class="Aves" family="Scolopacidae" kingdom="Animalia" order="Charadriiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="family">Scolopacidae</taxonomicName>
</td>
<td id="D4149070FFE15AB9FD71F973FD2AF914" box="[661,766,1681,1736]" gridcol="2" gridrow="11" pageId="23" pageNumber="24">
<collectingCountry id="51E2BBDEFFE1A551FD56F943FD36F964" box="[690,738,1697,1720]" name="United States of America" pageId="23" pageNumber="24">USA</collectingCountry>
</td>
<td id="D4149070FFE15AB9FCF9F973FC4FF914" box="[797,923,1681,1736]" gridcol="3" gridrow="11" pageId="23" pageNumber="24">150–300</td>
<td id="D4149070FFE15AB9FC57F973FBDBF914" box="[947,1039,1681,1736]" gridcol="4" gridrow="11" pageId="23" pageNumber="24">476–523</td>
<td id="D4149070FFE15AB9FBC2F973FB56F914" box="[1062,1154,1681,1736]" gridcol="5" gridrow="11" pageId="23" pageNumber="24">143–165</td>
<td id="D4149070FFE15AB9FB7DF973FAD5F914" box="[1177,1281,1681,1736]" gridcol="6" gridrow="11" pageId="23" pageNumber="24">408–620 × 176–204</td>
<td id="D4149070FFE15AB9FAF3F973FAABF914" box="[1303,1407,1681,1736]" gridcol="7" gridrow="11" pageId="23" pageNumber="24">442–516 × 136–176</td>
<td id="D4149070FFE15AB9FA71F973FA32F914" box="[1429,1510,1681,1736]" gridcol="8" gridrow="11" pageId="23" pageNumber="24">20</td>
<td id="D4149070FFE15AB9FA19F973F982F914" box="[1533,1622,1681,1736]" gridcol="9" gridrow="11" pageId="23" pageNumber="24">100–105</td>
<td id="D4149070FFE15AB9F989F973F91FF914" box="[1645,1739,1681,1736]" gridcol="10" gridrow="11" pageId="23" pageNumber="24">110–115</td>
<td id="D4149070FFE15AB9F905F973F8FBF914" box="[1761,1839,1681,1736]" gridcol="11" gridrow="11" pageId="23" pageNumber="24">35–40</td>
<td id="D4149070FFE15AB9F8A1F973F84DF914" box="[1861,1945,1681,1736]" gridcol="12" gridrow="11" pageId="23" pageNumber="24">132–165 × 33–60</td>
</tr>
</table>
</paragraph>
<caption id="7D8AABC6FFEEA55EFF58FD09FE79FCDB" box="[188,429,747,775]" pageId="24" pageNumber="25">
<paragraph id="294AFB4EFFEEA55EFF58FD09FE79FCDB" blockId="24.[188,429,747,775]" box="[188,429,747,775]" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFF58FD09FECCFCDB" bold="true" box="[188,280,747,775]" pageId="24" pageNumber="25">Table 4</emphasis>
(continued).
</paragraph>
</caption>
<paragraph id="294AFB4EFFEEA55EFF24FC48F842FA20" pageId="24" pageNumber="25">
<table id="5BF509EEFFEE5AB9FF24FC6DF84DFA20" box="[192,1945,911,1532]" gridcols="13" gridrows="8" pageId="24" pageNumber="25">
<tr id="97C5F90CFFEE5AB9FF24FC6DF84DFC00" box="[192,1945,911,988]" gridrow="0" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FC6DFE61FC00" box="[192,437,911,988]" gridcol="0" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFF24FC48FEDFFC1D" bold="true" box="[192,267,938,961]" pageId="24" pageNumber="25">Species</emphasis>
</th>
<th id="D4149070FFEE5AB9FE38FC6DFDADFC00" box="[476,633,911,988]" gridcol="1" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFE0BFC48FDB2FC1D" bold="true" box="[495,614,938,961]" pageId="24" pageNumber="25">Host family</emphasis>
</th>
<th id="D4149070FFEE5AB9FD74FC6DFCD7FC00" box="[656,771,911,988]" gridcol="2" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFD54FC7FFD3BFC12" bold="true" pageId="24" pageNumber="25">Type locality</emphasis>
</th>
<th id="D4149070FFEE5AB9FCF9FC6DFC4FFC00" box="[797,923,911,988]" gridcol="3" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFCA5FC6DFC4FFC00" bold="true" pageId="24" pageNumber="25">Body maximum length (mm)</emphasis>
</th>
<th id="D4149070FFEE5AB9FC57FC6DFBDBFC00" box="[947,1039,911,988]" gridcol="4" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFC25FC7FFBDBFC12" bold="true" pageId="24" pageNumber="25">Scolex diameter</emphasis>
</th>
<th id="D4149070FFEE5AB9FBC2FC6DFB56FC00" box="[1062,1154,911,988]" gridcol="5" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFBCFFC7FFB56FC12" bold="true" pageId="24" pageNumber="25">Suckers diameter</emphasis>
</th>
<th id="D4149070FFEE5AB9FB7DFC6DFAD5FC00" box="[1177,1281,911,988]" gridcol="6" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFB7AFC7FFB09FC12" bold="true" box="[1182,1275,925,974]" pageId="24" pageNumber="25">Rostellar sac</emphasis>
</th>
<th id="D4149070FFEE5AB9FAF3FC6DFAABFC00" box="[1303,1407,911,988]" gridcol="7" gridrow="0" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFAF3FC48FAABFC1D" authorityName="Montfort" authorityYear="1810" box="[1303,1407,938,961]" family="Rostellum" pageId="24" pageNumber="18" rank="family">
<emphasis id="1B81275CFFEEA55EFAF3FC48FAABFC1D" bold="true" box="[1303,1407,938,961]" pageId="24" pageNumber="25">Rostellum</emphasis>
</taxonomicName>
</th>
<th id="D4149070FFEE5AB9FA71FC6DFA32FC00" box="[1429,1510,911,988]" gridcol="8" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFA79FC7FFA32FC12" bold="true" pageId="24" pageNumber="25">Hooks number</emphasis>
</th>
<th id="D4149070FFEE5AB9FA19FC6DF982FC00" box="[1533,1622,911,988]" gridcol="9" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EFA19FC6DF99EFC00" bold="true" pageId="24" pageNumber="25">Anterior hooks length</emphasis>
</th>
<th id="D4149070FFEE5AB9F989FC6DF91FFC00" box="[1645,1739,911,988]" gridcol="10" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EF989FC6DF968FC00" bold="true" pageId="24" pageNumber="25">Posterior hooks length</emphasis>
</th>
<th id="D4149070FFEE5AB9F905FC6DF8FBFC00" box="[1761,1839,911,988]" gridcol="11" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EF90FFC7FF8FBFC12" bold="true" pageId="24" pageNumber="25">Testes number</emphasis>
</th>
<th id="D4149070FFEE5AB9F8A1FC6DF84DFC00" box="[1861,1945,911,988]" gridcol="12" gridrow="0" pageId="24" pageNumber="25">
<emphasis id="1B81275CFFEEA55EF8A9FC7FF8ABFC12" bold="true" box="[1869,1937,925,974]" pageId="24" pageNumber="25">Cirrus sac</emphasis>
</th>
</tr>
<tr id="97C5F90CFFEE5AB9FF24FC11F84DFBF6" box="[192,1945,1011,1066]" gridrow="1" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FC11FE61FBF6" box="[192,437,1011,1066]" gridcol="0" gridrow="1" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFF24FC11FEE4FBF6" authority="(Fuhrmann &amp; Baer, 1943)" baseAuthorityName="Fuhrmann &amp; Baer" baseAuthorityYear="1943" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="riccii">
<emphasis id="1B81275CFFEEA55EFF24FC11FEC4FBD6" box="[192,272,1011,1034]" italics="true" pageId="24" pageNumber="25">D. riccii</emphasis>
(
<bibRefCitation id="4D6486BFFFEEA55EFEF8FC11FEFDFBF6" author="Fuhrmann O. &amp; Baer J. G." pageId="24" pageNumber="25" pagination="113 - 140" refId="ref19370" refString="Fuhrmann O. &amp; Baer J. G. 1943. Mission biologique Sagan-Omo (Ethiopie meridionale), 1939, dirigee par le professeur Eduardo Zavattari. Cestodes. Bulletin de la Societe neuchateloise de Sciences naturelles 68: 113 - 140." type="journal article" year="1943">Fuhrmann &amp; Baer, 1943</bibRefCitation>
)
</taxonomicName>
</th>
<td id="D4149070FFEE5AB9FE38FC11FDADFBF6" box="[476,633,1011,1066]" gridcol="1" gridrow="1" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFE13FBE1FD8BFBC6" box="[503,607,1027,1050]" class="Aves" family="Ciconiidae" kingdom="Animalia" order="Ciconiiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="family">Ciconiidae</taxonomicName>
</td>
<td id="D4149070FFEE5AB9FD74FC11FCD7FBF6" box="[656,771,1011,1066]" gridcol="2" gridrow="1" pageId="24" pageNumber="25">Africa</td>
<td id="D4149070FFEE5AB9FCF9FC11FC4FFBF6" box="[797,923,1011,1066]" gridcol="3" gridrow="1" pageId="24" pageNumber="25">35–40</td>
<td id="D4149070FFEE5AB9FC57FC11FBDBFBF6" box="[947,1039,1011,1066]" gridcol="4" gridrow="1" pageId="24" pageNumber="25">400</td>
<td id="D4149070FFEE5AB9FBC2FC11FB56FBF6" box="[1062,1154,1011,1066]" gridcol="5" gridrow="1" pageId="24" pageNumber="25">240</td>
<td id="D4149070FFEE5AB9FB7DFC11FAD5FBF6" box="[1177,1281,1011,1066]" gridcol="6" gridrow="1" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9FAF3FC11FAABFBF6" box="[1303,1407,1011,1066]" gridcol="7" gridrow="1" pageId="24" pageNumber="25">400 × 72</td>
<td id="D4149070FFEE5AB9FA71FC11FA32FBF6" box="[1429,1510,1011,1066]" gridcol="8" gridrow="1" pageId="24" pageNumber="25">24</td>
<td id="D4149070FFEE5AB9FA19FC11F982FBF6" box="[1533,1622,1011,1066]" gridcol="9" gridrow="1" pageId="24" pageNumber="25">96–100</td>
<td id="D4149070FFEE5AB9F989FC11F91FFBF6" box="[1645,1739,1011,1066]" gridcol="10" gridrow="1" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9F905FC11F8FBFBF6" box="[1761,1839,1011,1066]" gridcol="11" gridrow="1" pageId="24" pageNumber="25">20–22</td>
<td id="D4149070FFEE5AB9F8A1FC11F84DFBF6" box="[1861,1945,1011,1066]" gridcol="12" gridrow="1" pageId="24" pageNumber="25">115–126 × 39</td>
</tr>
<tr id="97C5F90CFFEE5AB9FF24FBA0F84DFBA5" box="[192,1945,1090,1145]" gridrow="2" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FBA0FE61FBA5" box="[192,437,1090,1145]" gridcol="0" gridrow="2" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFF24FBA0FF2CFBA5" authority="(Linstow, 1906)" baseAuthorityName="von Linstow" baseAuthorityYear="1906" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="volvulus">
<emphasis id="1B81275CFFEEA55EFF24FBA0FEF9FB85" box="[192,301,1090,1113]" italics="true" pageId="24" pageNumber="25">D. volvulus</emphasis>
(Linstow, 1906)
</taxonomicName>
</th>
<td id="D4149070FFEE5AB9FE38FBA0FDADFBA5" box="[476,633,1090,1145]" gridcol="1" gridrow="2" pageId="24" pageNumber="25">Charadriiformes</td>
<td id="D4149070FFEE5AB9FD74FBA0FCD7FBA5" box="[656,771,1090,1145]" gridcol="2" gridrow="2" pageId="24" pageNumber="25">Palaearctic</td>
<td id="D4149070FFEE5AB9FCF9FBA0FC4FFBA5" box="[797,923,1090,1145]" gridcol="3" gridrow="2" pageId="24" pageNumber="25">60</td>
<td id="D4149070FFEE5AB9FC57FBA0FBDBFBA5" box="[947,1039,1090,1145]" gridcol="4" gridrow="2" pageId="24" pageNumber="25">275–26</td>
<td id="D4149070FFEE5AB9FBC2FBA0FB56FBA5" box="[1062,1154,1090,1145]" gridcol="5" gridrow="2" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9FB7DFBA0FAD5FBA5" box="[1177,1281,1090,1145]" gridcol="6" gridrow="2" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9FAF3FBA0FAABFBA5" box="[1303,1407,1090,1145]" gridcol="7" gridrow="2" pageId="24" pageNumber="25">125 × 67</td>
<td id="D4149070FFEE5AB9FA71FBA0FA32FBA5" box="[1429,1510,1090,1145]" gridcol="8" gridrow="2" pageId="24" pageNumber="25">24–26</td>
<td id="D4149070FFEE5AB9FA19FBA0F982FBA5" box="[1533,1622,1090,1145]" gridcol="9" gridrow="2" pageId="24" pageNumber="25">45</td>
<td id="D4149070FFEE5AB9F989FBA0F91FFBA5" box="[1645,1739,1090,1145]" gridcol="10" gridrow="2" pageId="24" pageNumber="25">48</td>
<td id="D4149070FFEE5AB9F905FBA0F8FBFBA5" box="[1761,1839,1090,1145]" gridcol="11" gridrow="2" pageId="24" pageNumber="25">40–60</td>
<td id="D4149070FFEE5AB9F8A1FBA0F84DFBA5" box="[1861,1945,1090,1145]" gridcol="12" gridrow="2" pageId="24" pageNumber="25">75–85 × 45</td>
</tr>
<tr id="97C5F90CFFEE5AB9FF24FB7FF84DFB68" box="[192,1945,1181,1204]" gridrow="3" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FB7FF84DFB68" box="[192,1945,1181,1204]" colspan="13" colspanRight="12" gridcol="0" gridrow="3" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFF24FB7FFF39FB68" authorityName="Weinland" authorityYear="1857" box="[192,237,1181,1204]" class="Cestoda" family="Dilepididae" genus="Liga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFEEA55EFF24FB7FFF39FB68" bold="true" box="[192,237,1181,1204]" italics="true" pageId="24" pageNumber="25">Liga</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="97C5F90CFFEE5AB9FF24FB3BF84DFACC" box="[192,1945,1241,1296]" gridrow="4" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FB3BFE61FACC" box="[192,437,1241,1296]" gridcol="0" gridrow="4" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFF24FB0BFE61FADC" authority="(Rysavy, 1966)" baseAuthorityName="Rysavy" baseAuthorityYear="1966" box="[192,437,1257,1280]" class="Cestoda" family="Dilepididae" genus="Liga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="centuri">
<emphasis id="1B81275CFFEEA55EFF24FB0BFEC9FADC" box="[192,285,1257,1280]" italics="true" pageId="24" pageNumber="25">L. centuri</emphasis>
(Rysavy, 1966)
</taxonomicName>
</th>
<td id="D4149070FFEE5AB9FE38FB3BFDADFACC" box="[476,633,1241,1296]" gridcol="1" gridrow="4" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFDE3FB0BFD9AFADC" box="[519,590,1257,1280]" class="Aves" family="Picidae" kingdom="Animalia" order="Piciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="family">Picidae</taxonomicName>
</td>
<td id="D4149070FFEE5AB9FD74FB3BFCD7FACC" box="[656,771,1241,1296]" gridcol="2" gridrow="4" pageId="24" pageNumber="25">Cuba</td>
<td id="D4149070FFEE5AB9FCF9FB3BFC4FFACC" box="[797,923,1241,1296]" gridcol="3" gridrow="4" pageId="24" pageNumber="25">22–47</td>
<td id="D4149070FFEE5AB9FC57FB3BFBDBFACC" box="[947,1039,1241,1296]" gridcol="4" gridrow="4" pageId="24" pageNumber="25">370–390</td>
<td id="D4149070FFEE5AB9FBC2FB3BFB56FACC" box="[1062,1154,1241,1296]" gridcol="5" gridrow="4" pageId="24" pageNumber="25">198–228</td>
<td id="D4149070FFEE5AB9FB7DFB3BFAD5FACC" box="[1177,1281,1241,1296]" gridcol="6" gridrow="4" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9FAF3FB3BFAABFACC" box="[1303,1407,1241,1296]" gridcol="7" gridrow="4" pageId="24" pageNumber="25">234–260 × 104–137</td>
<td id="D4149070FFEE5AB9FA71FB3BFA32FACC" box="[1429,1510,1241,1296]" gridcol="8" gridrow="4" pageId="24" pageNumber="25">18–20</td>
<td id="D4149070FFEE5AB9FA19FB3BF982FACC" box="[1533,1622,1241,1296]" gridcol="9" gridrow="4" pageId="24" pageNumber="25">62–64</td>
<td id="D4149070FFEE5AB9F989FB3BF91FFACC" box="[1645,1739,1241,1296]" gridcol="10" gridrow="4" pageId="24" pageNumber="25">one row</td>
<td id="D4149070FFEE5AB9F905FB3BF8FBFACC" box="[1761,1839,1241,1296]" gridcol="11" gridrow="4" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9F8A1FB3BF84DFACC" box="[1861,1945,1241,1296]" gridcol="12" gridrow="4" pageId="24" pageNumber="25">92–122 × 60–68</td>
</tr>
<tr id="97C5F90CFFEE5AB9FF24FACAF84DFA83" box="[192,1945,1320,1375]" gridrow="5" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FACAFE61FA83" box="[192,437,1320,1375]" gridcol="0" gridrow="5" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFF24FACAFF2CFA83" authority="(Goeze, 1782)" baseAuthorityName="Goeze" baseAuthorityYear="1782" class="Cestoda" family="Dilepididae" genus="Liga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="crateriformis">
<emphasis id="1B81275CFFEEA55EFF24FACAFE8CFAE3" box="[192,344,1320,1343]" italics="true" pageId="24" pageNumber="25">L. crateriformis</emphasis>
(Goeze, 1782)
</taxonomicName>
</th>
<td id="D4149070FFEE5AB9FE38FACAFDADFA83" box="[476,633,1320,1375]" gridcol="1" gridrow="5" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFDE3FADAFD9AFA93" box="[519,590,1336,1359]" class="Aves" family="Picidae" kingdom="Animalia" order="Piciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="family">Picidae</taxonomicName>
</td>
<td id="D4149070FFEE5AB9FD74FACAFCD7FA83" box="[656,771,1320,1375]" gridcol="2" gridrow="5" pageId="24" pageNumber="25">Palaearcic</td>
<td id="D4149070FFEE5AB9FCF9FACAFC4FFA83" box="[797,923,1320,1375]" gridcol="3" gridrow="5" pageId="24" pageNumber="25">18–25</td>
<td id="D4149070FFEE5AB9FC57FACAFBDBFA83" box="[947,1039,1320,1375]" gridcol="4" gridrow="5" pageId="24" pageNumber="25">275–430</td>
<td id="D4149070FFEE5AB9FBC2FACAFB56FA83" box="[1062,1154,1320,1375]" gridcol="5" gridrow="5" pageId="24" pageNumber="25">109–173</td>
<td id="D4149070FFEE5AB9FB7DFACAFAD5FA83" box="[1177,1281,1320,1375]" gridcol="6" gridrow="5" pageId="24" pageNumber="25">229–240</td>
<td id="D4149070FFEE5AB9FAF3FACAFAABFA83" box="[1303,1407,1320,1375]" gridcol="7" gridrow="5" pageId="24" pageNumber="25">136–229 × 70–73</td>
<td id="D4149070FFEE5AB9FA71FACAFA32FA83" box="[1429,1510,1320,1375]" gridcol="8" gridrow="5" pageId="24" pageNumber="25">28–35</td>
<td id="D4149070FFEE5AB9FA19FACAF982FA83" box="[1533,1622,1320,1375]" gridcol="9" gridrow="5" pageId="24" pageNumber="25">27–35</td>
<td id="D4149070FFEE5AB9F989FACAF91FFA83" box="[1645,1739,1320,1375]" gridcol="10" gridrow="5" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9F905FACAF8FBFA83" box="[1761,1839,1320,1375]" gridcol="11" gridrow="5" pageId="24" pageNumber="25">16–35</td>
<td id="D4149070FFEE5AB9F8A1FACAF84DFA83" box="[1861,1945,1320,1375]" gridcol="12" gridrow="5" pageId="24" pageNumber="25">64–145 × 40–55</td>
</tr>
<tr id="97C5F90CFFEE5AB9FF24FA95F84DFA72" box="[192,1945,1399,1454]" gridrow="6" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FA95FE61FA72" box="[192,437,1399,1454]" gridcol="0" gridrow="6" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFF24FA65FEA8FA42" authority="(Baer, 1959)" baseAuthorityName="Baer" baseAuthorityYear="1959" box="[192,380,1415,1438]" class="Cestoda" family="Dilepididae" genus="Liga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="pici">
<emphasis id="1B81275CFFEEA55EFF24FA65FF29FA42" box="[192,253,1415,1438]" italics="true" pageId="24" pageNumber="25">L. pici</emphasis>
(
<bibRefCitation id="4D6486BFFFEEA55EFEEEFA65FEA1FA42" author="Baer J. G." box="[266,373,1415,1438]" pageId="24" pageNumber="25" refId="ref18637" refString="Baer J. G. 1959. Exploration des Parcs Nationaux du Congo Belge. Mission J. G. Baer - W. Gerber (1958). Helminthes parasites. Institut des Parcs Nationaux du Congo Belge, Bruxelles." type="book" year="1959">Baer, 1959</bibRefCitation>
)
</taxonomicName>
</th>
<td id="D4149070FFEE5AB9FE38FA95FDADFA72" box="[476,633,1399,1454]" gridcol="1" gridrow="6" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFDE3FA65FD9AFA42" box="[519,590,1415,1438]" class="Aves" family="Picidae" kingdom="Animalia" order="Piciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="family">Picidae</taxonomicName>
</td>
<td id="D4149070FFEE5AB9FD74FA95FCD7FA72" box="[656,771,1399,1454]" gridcol="2" gridrow="6" pageId="24" pageNumber="25">D.R. Congo</td>
<td id="D4149070FFEE5AB9FCF9FA95FC4FFA72" box="[797,923,1399,1454]" gridcol="3" gridrow="6" pageId="24" pageNumber="25">15</td>
<td id="D4149070FFEE5AB9FC57FA95FBDBFA72" box="[947,1039,1399,1454]" gridcol="4" gridrow="6" pageId="24" pageNumber="25">160–190</td>
<td id="D4149070FFEE5AB9FBC2FA95FB56FA72" box="[1062,1154,1399,1454]" gridcol="5" gridrow="6" pageId="24" pageNumber="25">98–102</td>
<td id="D4149070FFEE5AB9FB7DFA95FAD5FA72" box="[1177,1281,1399,1454]" gridcol="6" gridrow="6" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9FAF3FA95FAABFA72" box="[1303,1407,1399,1454]" gridcol="7" gridrow="6" pageId="24" pageNumber="25">– × 45–50</td>
<td id="D4149070FFEE5AB9FA71FA95FA32FA72" box="[1429,1510,1399,1454]" gridcol="8" gridrow="6" pageId="24" pageNumber="25">26</td>
<td id="D4149070FFEE5AB9FA19FA95F982FA72" box="[1533,1622,1399,1454]" gridcol="9" gridrow="6" pageId="24" pageNumber="25">17–18</td>
<td id="D4149070FFEE5AB9F989FA95F91FFA72" box="[1645,1739,1399,1454]" gridcol="10" gridrow="6" pageId="24" pageNumber="25">–</td>
<td id="D4149070FFEE5AB9F905FA95F8FBFA72" box="[1761,1839,1399,1454]" gridcol="11" gridrow="6" pageId="24" pageNumber="25">38–40</td>
<td id="D4149070FFEE5AB9F8A1FA95F84DFA72" box="[1861,1945,1399,1454]" gridcol="12" gridrow="6" pageId="24" pageNumber="25">68–91 × 34–45</td>
</tr>
<tr id="97C5F90CFFEE5AB9FF24FA27F84DFA20" box="[192,1945,1477,1532]" gridrow="7" pageId="24" pageNumber="25">
<th id="D4149070FFEE5AB9FF24FA27FE61FA20" box="[192,437,1477,1532]" gridcol="0" gridrow="7" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFF24FA37FE49FA30" authority="(Singh, 1964)" baseAuthorityName="Singh" baseAuthorityYear="1964" box="[192,413,1493,1516]" class="Cestoda" family="Dilepididae" genus="Liga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="picusi">
<emphasis id="1B81275CFFEEA55EFF24FA37FEC6FA30" box="[192,274,1493,1516]" italics="true" pageId="24" pageNumber="25">L. picusi</emphasis>
(
<bibRefCitation id="4D6486BFFFEEA55EFEC4FA37FE42FA30" author="Singh K. S." box="[288,406,1493,1516]" pageId="24" pageNumber="25" pagination="177 - 194" refId="ref21316" refString="Singh K. S. 1964. On six new avian cestodes form India. Parasitology 54: 177 - 194. https: // doi. org / 10.1017 / s 0031182000074461" type="journal article" year="1964">Singh, 1964</bibRefCitation>
)
</taxonomicName>
</th>
<td id="D4149070FFEE5AB9FE38FA27FDADFA20" box="[476,633,1477,1532]" gridcol="1" gridrow="7" pageId="24" pageNumber="25">
<taxonomicName id="EEF580CDFFEEA55EFDE3FA37FD9AFA30" box="[519,590,1493,1516]" class="Aves" family="Picidae" kingdom="Animalia" order="Piciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="family">Picidae</taxonomicName>
</td>
<td id="D4149070FFEE5AB9FD74FA27FCD7FA20" box="[656,771,1477,1532]" gridcol="2" gridrow="7" pageId="24" pageNumber="25">India</td>
<td id="D4149070FFEE5AB9FCF9FA27FC4FFA20" box="[797,923,1477,1532]" gridcol="3" gridrow="7" pageId="24" pageNumber="25">19.5–26.2</td>
<td id="D4149070FFEE5AB9FC57FA27FBDBFA20" box="[947,1039,1477,1532]" gridcol="4" gridrow="7" pageId="24" pageNumber="25">370–490</td>
<td id="D4149070FFEE5AB9FBC2FA27FB56FA20" box="[1062,1154,1477,1532]" gridcol="5" gridrow="7" pageId="24" pageNumber="25">130–150</td>
<td id="D4149070FFEE5AB9FB7DFA27FAD5FA20" box="[1177,1281,1477,1532]" gridcol="6" gridrow="7" pageId="24" pageNumber="25">270–310 × 134–140</td>
<td id="D4149070FFEE5AB9FAF3FA27FAABFA20" box="[1303,1407,1477,1532]" gridcol="7" gridrow="7" pageId="24" pageNumber="25">– × 80–100</td>
<td id="D4149070FFEE5AB9FA71FA27FA32FA20" box="[1429,1510,1477,1532]" gridcol="8" gridrow="7" pageId="24" pageNumber="25">I30</td>
<td id="D4149070FFEE5AB9FA19FA27F982FA20" box="[1533,1622,1477,1532]" gridcol="9" gridrow="7" pageId="24" pageNumber="25">46–54</td>
<td id="D4149070FFEE5AB9F989FA27F91FFA20" box="[1645,1739,1477,1532]" gridcol="10" gridrow="7" pageId="24" pageNumber="25">one row</td>
<td id="D4149070FFEE5AB9F905FA27F8FBFA20" box="[1761,1839,1477,1532]" gridcol="11" gridrow="7" pageId="24" pageNumber="25">25–30</td>
<td id="D4149070FFEE5AB9F8A1FA27F84DFA20" box="[1861,1945,1477,1532]" gridcol="12" gridrow="7" pageId="24" pageNumber="25">120–200 × 32–40</td>
</tr>
</table>
</paragraph>
<caption id="7D8AABC6FFEFA55FFF59F899FE32F800" ID-DOI="http://doi.org/10.5281/zenodo.3827564" ID-Zenodo-Dep="3827564" httpUri="https://zenodo.org/record/3827564/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,241,1915,1941]" targetBox="[210,1386,286,1854]" targetPageId="25">
<paragraph id="294AFB4EFFEFA55FFF59F899FE32F800" blockId="25.[189,1399,1915,2012]" pageId="25" pageNumber="26">
<emphasis id="1B81275CFFEFA55FFF59F899FE93F849" bold="true" box="[189,327,1915,1941]" pageId="25" pageNumber="26">Figs 23–28.</emphasis>
<taxonomicName id="EEF580CDFFEFA55FFEA8F89EFDACF849" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[332,632,1915,1941]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="longiuncinata" status="sp. nov.">
<emphasis id="1B81275CFFEFA55FFEA8F89EFDACF849" box="[332,632,1915,1941]" italics="true" pageId="25" pageNumber="26">Dictymetra longiuncinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFEFA55FFD99F89EFD02F84A" box="[637,726,1916,1942]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1B81275CFFEFA55FFD3EF899FD23F849" bold="true" box="[730,759,1915,1941]" pageId="25" pageNumber="26">23</emphasis>
. Scolex.
<emphasis id="1B81275CFFEFA55FFC84F899FCA9F849" bold="true" box="[864,893,1915,1941]" pageId="25" pageNumber="26">24</emphasis>
. Hooks.
<emphasis id="1B81275CFFEFA55FFC07F899FBD4F849" bold="true" box="[995,1024,1915,1941]" pageId="25" pageNumber="26">25</emphasis>
. Mature proglottis.
<emphasis id="1B81275CFFEFA55FFB00F899FAD5F849" bold="true" box="[1252,1281,1915,1941]" pageId="25" pageNumber="26">26</emphasis>
. Terminal genitalia.
<emphasis id="1B81275CFFEFA55FFED0F87CFE85F864" bold="true" box="[308,337,1950,1976]" pageId="25" pageNumber="26">27</emphasis>
. Cirrus armament.
<emphasis id="1B81275CFFEFA55FFDDAF87CFD8FF864" bold="true" box="[574,603,1950,1976]" pageId="25" pageNumber="26">28</emphasis>
. Eggs. Scale bars: 23 = 100 µm; 24, 26 = 50 µm; 25 = 200 µm; 27 = 10 µm; 28 = 20 µm.
</paragraph>
</caption>
<paragraph id="294AFB4EFFECA55CFF59FEE9FDBBFE94" blockId="26.[189,1399,267,329]" pageId="26" pageNumber="27">diameter. Oncospheres 38–46 (42, n = 22) in diameter. Embryonic hooks: central 20–21.5 (21, n = 8) long, lateral 17–20 (19, n = 16) long.</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFECA55CFF59FE92FC99FB4B" pageId="26" pageNumber="27" type="discussion">
<paragraph id="294AFB4EFFECA55CFF59FE92FEFBFE56" blockId="26.[189,303,368,394]" box="[189,303,368,394]" pageId="26" pageNumber="27">
<heading id="72024C22FFECA55CFF59FE92FEFBFE56" bold="true" box="[189,303,368,394]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="1B81275CFFECA55CFF59FE92FEFBFE56" bold="true" box="[189,303,368,394]" pageId="26" pageNumber="27">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFECA55CFF59FE7CFC0FFCC5" blockId="26.[189,1399,414,793]" pageId="26" pageNumber="27">
This material is partly macerated, with many fragments contracted and overall in an average state of conservation. This explains the lack of data for several characters such as osmoregulatory canals or Mehlis’ gland. Our observations, nevertheless, allow for a description that fits the definition of
<taxonomicName id="EEF580CDFFECA55CFB0FFE04FABBFE23" authorityName="Clark" authorityYear="1952" box="[1259,1391,486,511]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFECA55CFB0FFE04FABBFE23" box="[1259,1391,486,511]" italics="true" pageId="26" pageNumber="27">Dictymetra</emphasis>
</taxonomicName>
, in particular because of its characteristic scolex structure and tuft of setae in the genital atrium and on the cirrus. The genus
<taxonomicName id="EEF580CDFFECA55CFE27FDCEFD93FD99" authorityName="Clark" authorityYear="1952" box="[451,583,556,581]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFECA55CFE27FDCEFD93FD99" box="[451,583,556,581]" italics="true" pageId="26" pageNumber="27">Dictymetra</emphasis>
</taxonomicName>
is typically found in various families of
<taxonomicName id="EEF580CDFFECA55CFBD3FDC9FB2EFD99" box="[1079,1274,555,581]" class="Aves" kingdom="Animalia" order="Charadriiformes" pageId="26" pageNumber="27" phylum="Chordata" rank="order">Charadriiformes</taxonomicName>
, including
<taxonomicName id="EEF580CDFFECA55CFF59FDACFE09FDB5" authority="(Bona 1994)" baseAuthorityName="Bona" baseAuthorityYear="1994" box="[189,477,590,617]" class="Aves" family="Burhinidae" kingdom="Animalia" order="Charadriiformes" pageId="26" pageNumber="27" phylum="Chordata" rank="family">
Burhinidae (
<bibRefCitation id="4D6486BFFFECA55CFEB4FDADFE01FDB5" author="Bona F. V." box="[336,469,591,617]" pageId="26" pageNumber="27" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">Bona 1994</bibRefCitation>
)
</taxonomicName>
(see remarks above for
<taxonomicName id="EEF580CDFFECA55CFCE5FDADFC40FDB4" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[769,916,591,616]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="gerganae" status="sp. nov.">
<emphasis id="1B81275CFFECA55CFCE5FDADFC40FDB4" box="[769,916,591,616]" italics="true" pageId="26" pageNumber="27">D. gerganae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFECA55CFC79FDADFC2FFDB5" box="[925,1019,591,617]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
) and is cosmopolitan (
<tableCitation id="6477CEF5FFECA55CFAEFFDACFABDFDB5" box="[1291,1385,590,617]" captionStart="Table 4" captionStartId="23.[188,257,544,572]" captionTargetBox="[192,1945,762,1736]" captionText="Table 4 (continued on next page). Comparison of two new Dictymetra Clark, 1952 species with the presently recognized species in the genus and with “long” Liga Weinland, 1857 species (see text). All data as reported in Spasskaya &amp; Spasskii (1978) for Dictymetra and Spasskaya &amp; Spasskii (1977) for Liga except Burt (1940) for D. dispar, Clark (1952) for D. paranumenii, Fuhrmann &amp; Baer (1943) for D. riccii and Spasskaya &amp; Spasskii (1973) for D. volvulus. Note that values reported by various authors for D. laevigata (Rudolphi, 1919) and D. volvulus (Linstow, 1903) vary to a large extent (Spasskaya &amp; Spasskii 1978). Dictymetra paranumenii and D. radiaspinosa have rows of small spines on rostellum." httpUri="http://table.plazi.org/id/7D8AABC6FFE1A551FF58FDC2F9CFFD15" pageId="26" pageNumber="27" tableUuid="7D8AABC6FFE1A551FF58FDC2F9CFFD15">Table 4</tableCitation>
). Our material can easily be differentiated from other species in the genus because of its very long hooks and large
<taxonomicName id="EEF580CDFFECA55CFECAFD77FE75FD73" authorityName="Montfort" authorityYear="1810" box="[302,417,661,687]" family="Rostellum" pageId="26" pageNumber="18" rank="family">rostellum</taxonomicName>
. Similar characters are only found in
<taxonomicName id="EEF580CDFFECA55CFCB4FD74FB66FD73" authority="(Rudolphi, 1919)" baseAuthorityName="Rudolphi" baseAuthorityYear="1919" box="[848,1202,661,687]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="laevigata">
<emphasis id="1B81275CFFECA55CFCB4FD74FC34FD73" box="[848,992,661,687]" italics="true" pageId="26" pageNumber="27">D. laevigata</emphasis>
(Rudolphi, 1919)
</taxonomicName>
, another parasite of thick-knees, and
<taxonomicName id="EEF580CDFFECA55CFE44FD5BFD2AFD0F" authority="(Clark, 1952)" baseAuthorityName="Clark" baseAuthorityYear="1952" box="[416,766,696,723]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="paranumenii">
<emphasis id="1B81275CFFECA55CFE44FD5BFD8DFD0E" box="[416,601,697,722]" italics="true" pageId="26" pageNumber="27">D. paranumenii</emphasis>
(
<bibRefCitation id="4D6486BFFFECA55CFD83FD5AFD22FD0F" author="Clark D. T." box="[615,758,696,723]" pageId="26" pageNumber="27" pagination="18 - 27" refId="ref18941" refString="Clark D. T. 1952. Three new dilepidid cestodes, Dictymetra numenii n. gen. sp. nov.; Dictymetra paranumenii sp. nov. and Anomotaenia filovata sp. nov. Proceedings of the Helminthological Society of Washington 19: 18 - 27." type="journal article" year="1952">Clark, 1952</bibRefCitation>
)
</taxonomicName>
, a parasite of curlews in North America. It differs from both by its smaller number of testes. Additionally, it can be distinguished from the former species by its larger cirrus-sac, and from the latter by its shorter strobila (
<tableCitation id="6477CEF5FFECA55CFC95FD1CFC19FCC5" box="[881,973,766,793]" captionStart="Table 4" captionStartId="23.[188,257,544,572]" captionTargetBox="[192,1945,762,1736]" captionText="Table 4 (continued on next page). Comparison of two new Dictymetra Clark, 1952 species with the presently recognized species in the genus and with “long” Liga Weinland, 1857 species (see text). All data as reported in Spasskaya &amp; Spasskii (1978) for Dictymetra and Spasskaya &amp; Spasskii (1977) for Liga except Burt (1940) for D. dispar, Clark (1952) for D. paranumenii, Fuhrmann &amp; Baer (1943) for D. riccii and Spasskaya &amp; Spasskii (1973) for D. volvulus. Note that values reported by various authors for D. laevigata (Rudolphi, 1919) and D. volvulus (Linstow, 1903) vary to a large extent (Spasskaya &amp; Spasskii 1978). Dictymetra paranumenii and D. radiaspinosa have rows of small spines on rostellum." httpUri="http://table.plazi.org/id/7D8AABC6FFE1A551FF58FDC2F9CFFD15" pageId="26" pageNumber="27" tableUuid="7D8AABC6FFE1A551FF58FDC2F9CFFD15">Table 4</tableCitation>
).
</paragraph>
<paragraph id="294AFB4EFFECA55CFF59FCA2FC99FB4B" blockId="26.[189,1399,832,1175]" pageId="26" pageNumber="27">
In addition, the genus
<taxonomicName id="EEF580CDFFECA55CFE2DFCA2FC64FC86" authority="Spasskii &amp; Borgarenko, 1973" authorityName="Spasskii &amp; Borgarenko" authorityYear="1973" box="[457,944,832,858]" class="Cestoda" family="Dilepididae" genus="Stenovaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFECA55CFE2DFCA2FD9CFC86" box="[457,584,832,858]" italics="true" pageId="26" pageNumber="27">Stenovaria</emphasis>
<bibRefCitation id="4D6486BFFFECA55CFDB5FCA2FC64FC86" author="Spasskii A. A. &amp; Borgarenko L. F." box="[593,944,832,858]" pageId="26" pageNumber="27" pagination="87 - 93" refId="ref21661" refString="Spasskii A. A. &amp; Borgarenko L. F. 1973. Stenovaria n. g. (Cestoda: Dilepididae) - a new genus of cestodes from the stone curlew. Izvestiya Akademii Nauk Tadzhikskoi SSR, Biologicheskie Nauk 2: 87 - 93. [In Russian.]" type="journal article" year="1973">Spasskii &amp; Borgarenko, 1973</bibRefCitation>
</taxonomicName>
also resembles our material. It differs from
<taxonomicName id="EEF580CDFFECA55CFF18FC86FE54FCA1" authorityName="Clark" authorityYear="1952" box="[252,384,868,893]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFECA55CFF18FC86FE54FCA1" box="[252,384,868,893]" italics="true" pageId="26" pageNumber="27">Dictymetra</emphasis>
</taxonomicName>
by relatively minor characters related to genital pore alternation, the position of genital ducts in relation to osmoregulatory canals and the absence of armament of long setae on the cirrus and (or) genital atrium. The genus was erected by
<bibRefCitation id="4D6486BFFFECA55CFD34FC4BFBE0FC18" author="Spasskii A. A. &amp; Borgarenko L. F." box="[720,1076,937,964]" pageId="26" pageNumber="27" pagination="87 - 93" refId="ref21661" refString="Spasskii A. A. &amp; Borgarenko L. F. 1973. Stenovaria n. g. (Cestoda: Dilepididae) - a new genus of cestodes from the stone curlew. Izvestiya Akademii Nauk Tadzhikskoi SSR, Biologicheskie Nauk 2: 87 - 93. [In Russian.]" type="journal article" year="1973">Spasskii &amp; Borgarenko (1973)</bibRefCitation>
and is presently recognized as valid (
<bibRefCitation id="4D6486BFFFECA55CFEC3FC2FFE7FFC3B" author="Bona F. V." box="[295,427,973,999]" pageId="26" pageNumber="27" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">
<collectingCountry id="51E2BBDEFFECA55CFEC3FC2FFEB3FC3B" box="[295,359,973,999]" name="Algeria" pageId="26" pageNumber="27">Bona</collectingCountry>
1994
</bibRefCitation>
,
<bibRefCitation id="4D6486BFFFECA55CFE5CFC2FFD71FC3B" author="Mariaux J. &amp; Tkach V. V. &amp; Vasileva G. P. &amp; Waeschenbach A. &amp; Beveridge I. &amp; Dimitrova Y. D. &amp; Haukisalmi V. &amp; Greiman S. E. &amp; Littlewood D. T. J. &amp; Makarikov A. A. &amp; Phillips A. J. &amp; Razafiarisolo T. &amp; Widmer V. &amp; Georgiev B. B." box="[440,677,973,999]" pageId="26" pageNumber="27" pagination="2008 - 2017" refId="ref20421" refString="Mariaux J., Tkach V. V., Vasileva G. P., Waeschenbach A., Beveridge I., Dimitrova Y. D., Haukisalmi V., Greiman S. E., Littlewood D. T. J., Makarikov A. A., Phillips A. J., Razafiarisolo T., Widmer V. &amp; Georgiev B. B. 2017. Cyclophyllidea van Beneden in Braun, 1900. In: Caira J. N. &amp; Jensen K. (eds) Planetary Biodiversity Inventory (2008 - 2017): Tapeworms from Vertebrate Bowels of the Earth: 77 - 148. University of Kansas, Lawrence." type="book chapter" year="2017">
Mariaux
<emphasis id="1B81275CFFECA55CFDC7FC2CFD82FC3B" box="[547,598,973,999]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. 2017
</bibRefCitation>
). It includes 2 species found in thick-knees:
<taxonomicName id="EEF580CDFFECA55CFB4BFC2FFD1EFBD6" authority="(Meggitt, 1927) Spasskaya &amp; Spasskii, 1978" authorityName="Spasskaya &amp; Spasskii" authorityYear="1978" baseAuthorityName="Meggitt" baseAuthorityYear="1927" class="Cestoda" family="Dilepididae" genus="Stenovaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="facile">
<emphasis id="1B81275CFFECA55CFB4BFC2FFAA2FC3B" box="[1199,1398,973,999]" italics="true" pageId="26" pageNumber="27">Stenovaria facile</emphasis>
(
<bibRefCitation id="4D6486BFFFECA55CFF21FC12FEBBFBD6" author="Meggitt F. J." box="[197,367,1008,1034]" pageId="26" pageNumber="27" pagination="420 - 450" refId="ref20655" refString="Meggitt F. J. 1927. Report on a collection of Cestoda, mainly from Egypt. Part II. Cyclophyllidea: Family Hymenolepididae. Parasitology 19: 420 - 450. https: // doi. org / 10.1017 / S 0031182000005904" type="journal article" year="1927">Meggitt, 1927</bibRefCitation>
)
<bibRefCitation id="4D6486BFFFECA55CFE9AFC12FD1EFBD6" author="Spasskaya L. P. &amp; Spasskii A." box="[382,714,1008,1034]" pageId="26" pageNumber="27" refId="ref21548" refString="Spasskaya L. P. &amp; Spasskii A. 1978. Cestodes of Birds in the USSR. Dilepididae of Aquatic Birds. Izdate'stvo Nauka, Moscow. [In Russian.]" type="book" year="1978">Spasskaya &amp; Spasskii, 1978</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="EEF580CDFFECA55CFCE7FC12FF2CFBF1" authority="(Meggitt, 1927) Spasskii &amp; Borgarenko, 1973" authorityName="Spasskii &amp; Borgarenko" authorityYear="1973" baseAuthorityName="Meggitt" baseAuthorityYear="1927" class="Cestoda" family="Dilepididae" genus="Stenovaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="falsificata">
<emphasis id="1B81275CFFECA55CFCE7FC12FC41FBD6" box="[771,917,1008,1034]" italics="true" pageId="26" pageNumber="27">S. falsificata</emphasis>
(
<bibRefCitation id="4D6486BFFFECA55CFC40FC12FB9AFBD6" author="Meggitt F. J." box="[932,1102,1008,1034]" pageId="26" pageNumber="27" pagination="420 - 450" refId="ref20655" refString="Meggitt F. J. 1927. Report on a collection of Cestoda, mainly from Egypt. Part II. Cyclophyllidea: Family Hymenolepididae. Parasitology 19: 420 - 450. https: // doi. org / 10.1017 / S 0031182000005904" type="journal article" year="1927">Meggitt, 1927</bibRefCitation>
)
<bibRefCitation id="4D6486BFFFECA55CFBBAFC12FF2CFBF1" author="Spasskii A. A. &amp; Borgarenko L. F." pageId="26" pageNumber="27" pagination="87 - 93" refId="ref21661" refString="Spasskii A. A. &amp; Borgarenko L. F. 1973. Stenovaria n. g. (Cestoda: Dilepididae) - a new genus of cestodes from the stone curlew. Izvestiya Akademii Nauk Tadzhikskoi SSR, Biologicheskie Nauk 2: 87 - 93. [In Russian.]" type="journal article" year="1973">Spasskii &amp; Borgarenko, 1973</bibRefCitation>
</taxonomicName>
that deserve to be compared to our material. Both differ from it by their much shorter hooks and smaller strobila. In addition, the former species also shows a smaller cirrus-sac and the latter has a larger cirrus-sac and a smaller number of testes. In consequence we consider the present material as a new species that we name
<taxonomicName id="EEF580CDFFECA55CFE59FB9CFD3FFB4B" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[445,747,1149,1175]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="longiuncinata" status="sp. nov.">
<emphasis id="1B81275CFFECA55CFE59FB9CFD3FFB4B" box="[445,747,1149,1175]" italics="true" pageId="26" pageNumber="27">Dictymetra longiuncinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFECA55CFD16FB9FFC99FB4B" box="[754,845,1149,1175]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="17E425163560446CD12C8EAEC66EDB26">Seven new species of cestode parasites (Neodermata, Platyhelminthes) from Australian birds</mods:title>
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<mods:affiliation id="B7D774AAC34206959380E235A362172C">Natural History Museum of Geneva, CP 6434, 1211 Geneva 6, Switzerland. Department of Genetics and Evolution, University of Geneva, Boulevard d’Ivoy 4, 1205 Geneva, Switzerland. Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113 Sofia, Bulgaria.</mods:affiliation>
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<treatment id="A15C4A58FFEAA558FDB7FD1BFD95F85E" ID-DOI="http://doi.org/10.5281/zenodo.3846842" ID-GBIF-Taxon="164152606" ID-Zenodo-Dep="3846842" LSID="urn:lsid:zoobank.org:act:99114137-DA7B-47EE-996F-49005C2B8A07" httpUri="http://treatment.plazi.org/id/A15C4A58FFEAA558FDB7FD1BFD95F85E" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="61EFA8C5FFEAA55AFDB7FD1BFB6BFCEB" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="294AFB4EFFEAA55AFDB7FD1BFC34FCC8" blockId="28.[372,1215,761,858]" box="[595,992,761,788]" pageId="28" pageNumber="29">
<heading id="72024C22FFEAA55AFDB7FD1BFC34FCC8" box="[595,992,761,788]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="EEF580CDFFEAA55AFDB7FD1BFCABFCC8" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[595,895,761,788]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="28" pageNumber="29" phylum="Platyhelminthes" rank="species" species="musealis" status="sp. nov.">
<emphasis id="1B81275CFFEAA55AFDB7FD1BFCABFCC8" bold="true" box="[595,895,761,788]" italics="true" pageId="28" pageNumber="29">Notopentorchis musealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFEAA55AFC62FD18FC34FCC8" box="[902,992,762,788]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<paragraph id="294AFB4EFFEAA55AFE90FCFEFB6BFCEB" blockId="28.[372,1215,761,858]" box="[372,1215,796,823]" pageId="28" pageNumber="29">
<uri id="5D64F74CFFEAA55AFE90FCFEFB6BFCEB" box="[372,1215,796,823]" pageId="28" pageNumber="29">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFEAA55AFD42FCDDFC5AFC86" box="[678,910,831,858]" pageId="28" pageNumber="29" type="description">
<paragraph id="294AFB4EFFEAA55AFD42FCDDFC5AFC86" blockId="28.[372,1215,761,858]" box="[678,910,831,858]" pageId="28" pageNumber="29">
<figureCitation id="B1CEE7CBFFEAA55AFD42FCDDFCFFFC86" box="[678,811,831,858]" captionStart="Figs 29–33" captionStartId="29.[189,242,1915,1941]" captionTargetBox="[228,1357,287,1851]" captionTargetId="figure@29.[211,1377,265,1890]" captionTargetPageId="29" captionText="Figs 29–33. Notopentorchis musealis sp. nov. 29. Scolex. 30. Hooks: A, C. Anterior hooks, B. Posterior hook. 31. Mature proglottis. 32. Postmature proglottis. 33. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="28" pageNumber="29">Figs 29–33</figureCitation>
,
<tableCitation id="6477CEF5FFEAA55AFCD3FCDDFC5AFC86" box="[823,910,831,858]" captionStart="Table 5" captionStartId="31.[188,257,654,682]" captionText="Table 5. Comparison of Notopentorchis musealis sp. nov., with the presently known species of Notopentorchis Burt, 1938 (only species with rostellar hooks of similar length in both rows are listed). Values for known species are summarized from various sources as reported in Dimitrova et al. (2017). All species are parasites of Apodidae." pageId="28" pageNumber="29">Table 5</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFEAA55AFF59FC63FB16FC15" box="[189,1218,897,970]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="294AFB4EFFEAA55AFF59FC63FE90FC47" blockId="28.[189,324,897,923]" box="[189,324,897,923]" pageId="28" pageNumber="29">
<heading id="72024C22FFEAA55AFF59FC63FE90FC47" bold="true" box="[189,324,897,923]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="1B81275CFFEAA55AFF59FC63FE90FC47" bold="true" box="[189,324,897,923]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEAA55AFF59FC4DFB16FC15" blockId="28.[189,1218,943,970]" box="[189,1218,943,970]" pageId="28" pageNumber="29">The species name refers to the fact that this species is described from museum material.</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFEAA55AFF59FC13FBCEFAEE" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="294AFB4EFFEAA55AFF59FC13FE7EFBD7" blockId="28.[189,426,1009,1035]" box="[189,426,1009,1035]" pageId="28" pageNumber="29">
<heading id="72024C22FFEAA55AFF59FC13FE7EFBD7" bold="true" box="[189,426,1009,1035]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="1B81275CFFEAA55AFF59FC13FE7EFBD7" bold="true" box="[189,426,1009,1035]" pageId="28" pageNumber="29">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="999DF113FFEAA55AFF0EFBC2FA9BFB82" ID-GBIF-Occurrence="2619826301" collectorName="Karie Rose" country="Australia" latitude="-38.133335" location="Sydney" longLatPrecision="1177" longitude="144.36667" pageId="28" pageNumber="29" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<paragraph id="294AFB4EFFEAA55AFF0EFBC2FE88FBE6" blockId="28.[189,1359,1056,1118]" box="[234,348,1056,1082]" pageId="28" pageNumber="29">
<emphasis id="1B81275CFFEAA55AFF0EFBC2FE88FBE6" bold="true" box="[234,348,1056,1082]" pageId="28" pageNumber="29">
<typeStatus id="F64E45ECFFEAA55AFF0EFBC2FE88FBE6" box="[234,348,1056,1082]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFEAA55AFF59FBA1FA9BFB82" blockId="28.[189,1359,1056,1118]" box="[189,1359,1089,1118]" pageId="28" pageNumber="29">
<collectingCountry id="51E2BBDEFFEAA55AFF59FBA1FEBCFB81" box="[189,360,1091,1117]" name="Australia" pageId="28" pageNumber="29">AUSTRALIA</collectingCountry>
:
<collectingRegion id="EB3135ACFFEAA55AFE97FBA1FD93FB81" box="[371,583,1091,1117]" country="Australia" name="New South Wales" pageId="28" pageNumber="29">New South Wales</collectingRegion>
,
<location id="2C2AAD95FFEAA55AFDB6FBA1FD7FFB81" LSID="urn:lsid:plazi:treatment:A15C4A58FFEAA558FDB7FD1BFD95F85E:2C2AAD95FFEAA55AFDB6FBA1FD7FFB81" box="[594,683,1091,1117]" country="Australia" latitude="-38.133335" longLatPrecision="1177" longitude="144.36667" name="Sydney" pageId="28" pageNumber="29" stateProvince="New South Wales">Sydney</location>
,
<geoCoordinate id="4CC19D89FFEAA55AFD53FBA3FCDAFB81" box="[695,782,1089,1117]" degrees="38" direction="south" minutes="8" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="-38.133335">38°8′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFEAA55AFCFFFBA3FC46FB81" box="[795,914,1089,1117]" degrees="144" direction="east" minutes="22" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="144.36667">144°22′ E</geoCoordinate>
, 2005,
<collectorName id="84009E98FFEAA55AFC0CFBA1FBB8FB81" box="[1000,1132,1091,1117]" pageId="28" pageNumber="29">Karie Rose</collectorName>
leg. (AHC 36478).
</paragraph>
</materialsCitation>
<paragraph id="294AFB4EFFEAA55AFF0EFB66FE88FB42" blockId="28.[189,848,1156,1218]" box="[234,348,1156,1182]" pageId="28" pageNumber="29">
<emphasis id="1B81275CFFEAA55AFF0EFB66FE88FB42" bold="true" box="[234,348,1156,1182]" pageId="28" pageNumber="29">
<typeStatus id="F64E45ECFFEAA55AFF0EFB66FE88FB42" box="[234,348,1156,1182]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFEAA55AFF59FB45FC84FB1E" blockId="28.[189,848,1156,1218]" box="[189,848,1191,1218]" pageId="28" pageNumber="29">
<collectingCountry id="51E2BBDEFFEAA55AFF59FB45FEBCFB1D" box="[189,360,1191,1217]" name="Australia" pageId="28" pageNumber="29">AUSTRALIA</collectingCountry>
: same data as for
<typeStatus id="F64E45ECFFEAA55AFDDCFB45FD4BFB1D" box="[568,671,1191,1217]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
(AHC 36478).
</paragraph>
<paragraph id="294AFB4EFFEAA55AFF59FB0BFEF8FADF" blockId="28.[189,300,1257,1283]" box="[189,300,1257,1283]" pageId="28" pageNumber="29">
<heading id="72024C22FFEAA55AFF59FB0BFEF8FADF" bold="true" box="[189,300,1257,1283]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="1B81275CFFEAA55AFF59FB0BFEF8FADF" bold="true" box="[189,300,1257,1283]" pageId="28" pageNumber="29">Intensity</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEAA55AFF59FAFAFBCEFAEE" blockId="28.[189,1050,1303,1330]" box="[189,1050,1303,1330]" pageId="28" pageNumber="29">
Two specimens and a few small fragments. Co-infection with
<taxonomicName id="EEF580CDFFEAA55AFC71FAF5FC26FAED" authorityName="Singh" authorityYear="1952" box="[917,1010,1303,1329]" class="Cestoda" family="Dilepididae" genus="Neoliga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="28" pageNumber="29" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFEAA55AFC71FAF5FC26FAED" box="[917,1010,1303,1329]" italics="true" pageId="28" pageNumber="29">Neoliga</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFEAA558FF59FABBFC41FC1E" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="294AFB4EFFEAA55AFF59FABBFE9AFAAF" blockId="28.[189,334,1369,1395]" box="[189,334,1369,1395]" pageId="28" pageNumber="29">
<heading id="72024C22FFEAA55AFF59FABBFE9AFAAF" bold="true" box="[189,334,1369,1395]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="1B81275CFFEAA55AFF59FABBFE9AFAAF" bold="true" box="[189,334,1369,1395]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEAA55AFF59FA65FE86F8B7" blockId="28.[189,1399,1415,1899]" pageId="28" pageNumber="29">
Body of medium size, with maximum length
<quantity id="EE0D56ABFFEAA55AFD09FA65FC97FA7E" box="[749,835,1415,1442]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="28" pageNumber="29" unit="mm" value="34.0">34 mm</quantity>
for 404 proglottides; maximum width 590, at level of gravid proglottides. Proglottides craspedote, all but the few most developed (pre-gravid) ones wider than long. Ventral osmoregulatory canals connected posteriorly in each proglottis with transverse anastomosis. Scolex slightly rounded, not distinctly separated from neck, with diameter 248–345 (n = 2). Suckers elongated
<quantity id="EE0D56ABFFEAA55AFD61F9F6FCC4F9F2" box="[645,784,1556,1583]" metricMagnitude="0" metricUnit="m" metricValue="4.8895" metricValueMax="5.461" metricValueMin="4.318" pageId="28" pageNumber="29" unit="in" value="192.5" valueMax="215.0" valueMin="170.0">170–215 in</quantity>
diameter (182, n = 7).
<taxonomicName id="EEF580CDFFEAA55AFBDFF9F6FB62F9F2" authorityName="Montfort" authorityYear="1810" box="[1083,1206,1556,1582]" family="Rostellum" pageId="28" pageNumber="18" rank="family">Rostellum</taxonomicName>
very muscular, sucker-like, slightly concave anteriorly and conical posteriorly,
<quantity id="EE0D56ABFFEAA55AFC34F9D5FB8DF98D" box="[976,1113,1591,1617]" metricMagnitude="0" metricUnit="m" metricValue="3.2131" metricValueMax="3.5305999999999997" metricValueMin="2.8956" pageId="28" pageNumber="29" unit="in" value="126.5" valueMax="139.0" valueMin="114.0">114–139 in</quantity>
diameter and 107–125 long (n = 2) (
<figureCitation id="B1CEE7CBFFEAA55AFE8AF9B8FE18F9A9" box="[366,460,1626,1653]" captionStart="Figs 29–33" captionStartId="29.[189,242,1915,1941]" captionTargetBox="[228,1357,287,1851]" captionTargetId="figure@29.[211,1377,265,1890]" captionTargetPageId="29" captionText="Figs 29–33. Notopentorchis musealis sp. nov. 29. Scolex. 30. Hooks: A, C. Anterior hooks, B. Posterior hook. 31. Mature proglottis. 32. Postmature proglottis. 33. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="28" pageNumber="29">Fig. 29</figureCitation>
). Rostellar hooks mostly lost (probably several tens, only 7 remaining), in two rows, subequal,
<quantity id="EE0D56ABFFEAA55AFE5FF99CFDF2F944" box="[443,550,1662,1688]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" metricValueMax="5.842" metricValueMin="5.334" pageId="28" pageNumber="29" unit="in" value="22.0" valueMax="23.0" valueMin="21.0">21–23 in</quantity>
length (22, n = 5). Posterior hooks with slightly more curved handle than anterior hooks (
<figureCitation id="B1CEE7CBFFEAA55AFE5EF943FDC2F967" box="[442,534,1697,1723]" captionStart="Figs 29–33" captionStartId="29.[189,242,1915,1941]" captionTargetBox="[228,1357,287,1851]" captionTargetId="figure@29.[211,1377,265,1890]" captionTargetPageId="29" captionText="Figs 29–33. Notopentorchis musealis sp. nov. 29. Scolex. 30. Hooks: A, C. Anterior hooks, B. Posterior hook. 31. Mature proglottis. 32. Postmature proglottis. 33. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="28" pageNumber="29">Fig. 30</figureCitation>
); guard bifid. Small epiphyses on both guards and handles. Neck long; proglottization distinct at 475 from posterior margin of suckers. Genital pores at mid-length of lateral proglottis margin, irregularly alternating in short series, maximum 5 consecutive pores observed on one side. Ventral osmoregulatory canals connected posteriorly in each proglottis by single transverse anastomosis. Genital ducts passing ventrally to osmoregulatory canals. Genital atrium cup-shaped, voluminous.
</paragraph>
<paragraph id="294AFB4EFFEAA558FF59F871FDCDFE09" blockId="28.[189,1399,1938,2000]" lastBlockId="30.[189,1399,267,469]" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
Testes large, spherical, 5–8 (6.5, n = 11) in number, in single posterior and dorsal layer, often overlapping vitellarium and posterior parts of ovary, occasionally overlapping longitudinal osmoregulatory canals. External vas deferens convoluted antero-porally, poral to developing uterus surrounded by intensely-stained cells and forming together with them dense mass in anteroporal part of median field. Cirrus-sac 91–112 × 54–64 (105 × 59, n = 15), consisting of narrower, almost cylindrical, distal part and spherical thick-walled proximal part, often overlapping poral osmoregulatory canals; internal vas deferens coiled in proximal part of cirrus-sac. Cirrus cylindrical, unarmed,
<quantity id="EE0D56ABFFE8A558FED2FE59FE76FE09" box="[310,418,443,469]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="30" pageNumber="31" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
diameter.
</paragraph>
<caption id="7D8AABC6FFEBA55BFF59F899FD81F800" ID-DOI="http://doi.org/10.5281/zenodo.3827566" ID-Zenodo-Dep="3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,242,1915,1941]" targetBox="[228,1357,287,1851]" targetPageId="29">
<paragraph id="294AFB4EFFEBA55BFF59F899FD81F800" blockId="29.[189,1399,1915,2012]" pageId="29" pageNumber="30">
<emphasis id="1B81275CFFEBA55BFF59F899FE86F849" bold="true" box="[189,338,1915,1941]" pageId="29" pageNumber="30">Figs 29–33.</emphasis>
<taxonomicName id="EEF580CDFFEBA55BFE84F899FD5AF849" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[352,654,1915,1941]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="29" pageNumber="30" phylum="Platyhelminthes" rank="species" species="musealis" status="sp. nov.">
<emphasis id="1B81275CFFEBA55BFE84F899FD5AF849" box="[352,654,1915,1941]" italics="true" pageId="29" pageNumber="30">Notopentorchis musealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFEBA55BFD7FF89EFD2AF84A" box="[667,766,1916,1942]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1B81275CFFEBA55BFCEFF899FCFDF849" bold="true" box="[779,809,1915,1941]" pageId="29" pageNumber="30">29</emphasis>
. Scolex.
<emphasis id="1B81275CFFEBA55BFC41F899FC17F849" bold="true" box="[933,963,1915,1941]" pageId="29" pageNumber="30">30</emphasis>
. Hooks: A, C. Anterior hooks, B. Posterior hook.
<emphasis id="1B81275CFFEBA55BFE98F87CFE4EF864" bold="true" box="[380,410,1950,1976]" pageId="29" pageNumber="30">31</emphasis>
. Mature proglottis.
<emphasis id="1B81275CFFEBA55BFD6EF87CFD7CF864" bold="true" box="[650,680,1950,1976]" pageId="29" pageNumber="30">32</emphasis>
. Postmature proglottis.
<emphasis id="1B81275CFFEBA55BFC2CF87CFC32F864" bold="true" box="[968,998,1950,1976]" pageId="29" pageNumber="30">33</emphasis>
. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm.
</paragraph>
</caption>
<paragraph id="294AFB4EFFE8A558FF59FE1FFD76FD2C" blockId="30.[189,1399,509,752]" pageId="30" pageNumber="31">
Vitellarium oval, massive, mostly irregular, 55–76 × 40–50 (68 × 46, n = 10), situated adjacent to posterior proglottis margin. Ovary central, transversely elongate, initially slightly bilobate but rapidly becoming more compact and irregular (
<figureCitation id="B1CEE7CBFFE8A558FD4CFDA7FCECFDBC" box="[680,824,581,608]" captionStart="Figs 29–33" captionStartId="29.[189,242,1915,1941]" captionTargetBox="[228,1357,287,1851]" captionTargetId="figure@29.[211,1377,265,1890]" captionTargetPageId="29" captionText="Figs 29–33. Notopentorchis musealis sp. nov. 29. Scolex. 30. Hooks: A, C. Anterior hooks, B. Posterior hook. 31. Mature proglottis. 32. Postmature proglottis. 33. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="30" pageNumber="31">Figs 31–32</figureCitation>
). Mehlis’ gland overlapping or just anterior to vitellarium. Seminal receptacle with rounded distal end and tubular and gradually tapering proximal end. Vaginal pore dorsal to pore of cirrus-sac; most of copulatory part of vagina passing posteriorly to cirrus-sac, straight; lumen encircled by thin cellular sleeve; connected with seminal receptacle by thin and short conductive part (
<figureCitation id="B1CEE7CBFFE8A558FDDDFD37FD40FD2C" box="[569,660,725,752]" captionStart="Figs 29–33" captionStartId="29.[189,242,1915,1941]" captionTargetBox="[228,1357,287,1851]" captionTargetId="figure@29.[211,1377,265,1890]" captionTargetPageId="29" captionText="Figs 29–33. Notopentorchis musealis sp. nov. 29. Scolex. 30. Hooks: A, C. Anterior hooks, B. Posterior hook. 31. Mature proglottis. 32. Postmature proglottis. 33. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="30" pageNumber="31">Fig. 32</figureCitation>
).
</paragraph>
<paragraph id="294AFB4EFFE8A558FF59FCFAFC41FC1E" blockId="30.[189,1399,792,962]" pageId="30" pageNumber="31">
Young uterus in mature proglottides well marked, with strong walls, developing antero-dorsally to ovary, initially globular (
<figureCitation id="B1CEE7CBFFE8A558FDE8FCDEFDBCFC8A" box="[524,616,828,854]" captionStart="Figs 29–33" captionStartId="29.[189,242,1915,1941]" captionTargetBox="[228,1357,287,1851]" captionTargetId="figure@29.[211,1377,265,1890]" captionTargetPageId="29" captionText="Figs 29–33. Notopentorchis musealis sp. nov. 29. Scolex. 30. Hooks: A, C. Anterior hooks, B. Posterior hook. 31. Mature proglottis. 32. Postmature proglottis. 33. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="30" pageNumber="31">Fig. 32</figureCitation>
), often with some irregular incomplete septa. With increasing in size, uterus becoming more irregular, with densification surrounding its anterior wall (
<figureCitation id="B1CEE7CBFFE8A558FB2BFC82FAFEFCA6" box="[1231,1322,864,890]" captionStart="Figs 29–33" captionStartId="29.[189,242,1915,1941]" captionTargetBox="[228,1357,287,1851]" captionTargetId="figure@29.[211,1377,265,1890]" captionTargetPageId="29" captionText="Figs 29–33. Notopentorchis musealis sp. nov. 29. Scolex. 30. Hooks: A, C. Anterior hooks, B. Posterior hook. 31. Mature proglottis. 32. Postmature proglottis. 33. Pregravid proglottis. Scale bars: 29, 31–33 = 100 µm; 30 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827566" httpUri="https://zenodo.org/record/3827566/files/figure.png" pageId="30" pageNumber="31">Fig. 33</figureCitation>
); with further development, uterus increasing in size; paruterine organ developing mostly anterior to it but also embracing uterus laterally. No fully developed eggs available.
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFE8A558FF59FC08FD95F85E" pageId="30" pageNumber="31" type="discussion">
<paragraph id="294AFB4EFFE8A558FF59FC08FEFBFBD8" blockId="30.[189,303,1002,1028]" box="[189,303,1002,1028]" pageId="30" pageNumber="31">
<heading id="72024C22FFE8A558FF59FC08FEFBFBD8" bold="true" box="[189,303,1002,1028]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="1B81275CFFE8A558FF59FC08FEFBFBD8" bold="true" box="[189,303,1002,1028]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFE8A558FF59FBFBFE06FB43" blockId="30.[189,1399,1049,1183]" pageId="30" pageNumber="31">
The present material unambiguously belongs to
<taxonomicName id="EEF580CDFFE8A558FCD9FBFBFB45FBEF" authority="Burt, 1938" authorityName="Burt" authorityYear="1938" box="[829,1169,1049,1076]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE8A558FCD9FBFBFC21FBEF" box="[829,1013,1049,1075]" italics="true" pageId="30" pageNumber="31">Notopentorchis</emphasis>
Burt, 1938
</taxonomicName>
as diagnosed by
<bibRefCitation id="4D6486BFFFE8A558FF59FBDFFD92FB8B" author="Kornyushin V. V. &amp; Georgiev B. B." box="[189,582,1085,1111]" pageId="30" pageNumber="31" pagination="585 - 593" refId="ref20065" refString="Kornyushin V. V. &amp; Georgiev B. B. 1994. Family Metadilepididae Spasskii, 1959. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the cestode parasites of vertebrates: 585 - 593. CAB International, Wallingford." type="book chapter" year="1994">Georgiev &amp; Kornyushin (1994)</bibRefCitation>
. This genus has recently been thoroughly reviewed and presently comprises eight species, all of them found in swifts from the Old World (
<bibRefCitation id="4D6486BFFFE8A558FBB4FB83FAA5FBA7" author="Georgiev B. B. &amp; Bray R. A." box="[1104,1393,1121,1148]" pageId="30" pageNumber="31" pagination="121 - 133" refId="ref19471" refString="Georgiev B. B. &amp; Bray R. A. 1991. Notopentorchis cyathiformis (Frolich, 1791) n. comb. and N. iduncula (Spasskii, 1946) (Cestoda: Paruterinidae) from Palaearctic swifts (Aves: Apodiformes), with a review of the genus Notopentorchis Burt, 1938. Systematic Parasitology 20: 121 - 133. https: // doi. org / 10.1007 / bf 00007388" type="journal article" year="1991">Georgiev &amp; Bray 1991</bibRefCitation>
;
<bibRefCitation id="4D6486BFFFE8A558FF59FB67FE11FB43" author="Dimitrova Y. &amp; Mariaux J. &amp; Georgiev B." box="[189,453,1157,1183]" pageId="30" pageNumber="31" pagination="61 - 82" refId="ref19193" refString="Dimitrova Y., Mariaux J. &amp; Georgiev B. 2017. Redescriptions of four Palaeotropical species of the cestode genus Notopentorchis Burt, 1938 (Cyclophyllidea: Paruterinidae). Zootaxa 4290: 61 - 82. https: // doi. org / 10.11646 / zootaxa. 4290.1.3" type="journal article" year="2017">
Dimitrova
<emphasis id="1B81275CFFE8A558FEA4FB64FEA8FB43" box="[320,380,1157,1183]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="294AFB4EFFE8A558FF59FB2AFC0CFA91" blockId="30.[189,1399,1223,1358]" pageId="30" pageNumber="31">Our material is unfortunately in a rather poor state of conservation and limited as neither a scolex with a full set of hooks nor fully developed gravid proglottides are available. It is, however, sufficient to allow examination of most diagnostic characters. These characters were proved to be sufficiently unique to distinguish it from congeners and to allow for a species description.</paragraph>
<paragraph id="294AFB4EFFE8A558FF59FA94FC68F998" blockId="30.[189,1399,1398,1604]" pageId="30" pageNumber="31">
Of the eight known species, three belong to a group with rostellar hooks of clearly distinct length in each of the two rostellar rows (
<bibRefCitation id="4D6486BFFFE8A558FE02FA78FD33FA68" author="Dimitrova Y. &amp; Mariaux J. &amp; Georgiev B." box="[486,743,1434,1460]" pageId="30" pageNumber="31" pagination="61 - 82" refId="ref19193" refString="Dimitrova Y., Mariaux J. &amp; Georgiev B. 2017. Redescriptions of four Palaeotropical species of the cestode genus Notopentorchis Burt, 1938 (Cyclophyllidea: Paruterinidae). Zootaxa 4290: 61 - 82. https: // doi. org / 10.11646 / zootaxa. 4290.1.3" type="journal article" year="2017">
Dimitrova
<emphasis id="1B81275CFFE8A558FD83FA79FD75FA68" box="[615,673,1434,1460]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2017
</bibRefCitation>
). The present material differs from all remaining species by a larger cirrus-sac and larger suckers (
<tableCitation id="6477CEF5FFE8A558FD44FA5CFD2FFA04" box="[672,763,1469,1496]" captionStart="Table 5" captionStartId="31.[188,257,654,682]" captionText="Table 5. Comparison of Notopentorchis musealis sp. nov., with the presently known species of Notopentorchis Burt, 1938 (only species with rostellar hooks of similar length in both rows are listed). Values for known species are summarized from various sources as reported in Dimitrova et al. (2017). All species are parasites of Apodidae." pageId="30" pageNumber="31">Table 5</tableCitation>
). In addition, it differs from
<taxonomicName id="EEF580CDFFE8A558FBA8FA5DFAA3FA04" authority="Burt, 1938" authorityName="Burt" authorityYear="1938" box="[1100,1399,1470,1496]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="collocaliae">
<emphasis id="1B81275CFFE8A558FBA8FA5DFB25FA04" box="[1100,1265,1470,1496]" italics="true" pageId="30" pageNumber="31">N. collocaliae</emphasis>
Burt, 1938
</taxonomicName>
by a larger scolex and
<taxonomicName id="EEF580CDFFE8A558FE2CFA00FDEFFA20" authorityName="Montfort" authorityYear="1810" box="[456,571,1506,1532]" family="Rostellum" pageId="30" pageNumber="18" rank="family">rostellum</taxonomicName>
; from
<taxonomicName id="EEF580CDFFE8A558FD63FA01FC37FA20" authority="(Hubscher, 1937)" baseAuthorityName="Hubscher" baseAuthorityYear="1937" box="[647,995,1506,1532]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="javanica">
<emphasis id="1B81275CFFE8A558FD63FA01FCDBFA20" box="[647,783,1507,1532]" italics="true" pageId="30" pageNumber="31">N. javanica</emphasis>
(Hübscher, 1937)
</taxonomicName>
by fewer testes; from
<taxonomicName id="EEF580CDFFE8A558FB09FA01FEAAF9FD" authority="(Spasskii, 1946)" baseAuthorityName="Spasskii" baseAuthorityYear="1946" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="iduncula">
<emphasis id="1B81275CFFE8A558FB09FA01FAA3FA20" box="[1261,1399,1506,1532]" italics="true" pageId="30" pageNumber="31">N. iduncula</emphasis>
(Spasskii, 1946)
</taxonomicName>
,
<taxonomicName id="EEF580CDFFE8A558FE6DF9E5FD30F9FC" authority="(Mokhehle, 1951)" baseAuthorityName="Mokhehle" baseAuthorityYear="1951" box="[393,740,1541,1568]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="caffrapi">
<emphasis id="1B81275CFFE8A558FE6DF9E5FDDCF9FC" box="[393,520,1542,1568]" italics="true" pageId="30" pageNumber="31">N. caffrapi</emphasis>
(
<bibRefCitation id="4D6486BFFFE8A558FDF2F9E4FD0FF9FC" author="Mokhehle" box="[534,731,1541,1568]" pageId="30" pageNumber="31" pagination="302 - 396" refId="ref20794" refString="Mokhehle 1951. Parasites of the swift Caffrapus caffer caffer (Roberts) with descriptions of one new genus and eight new species. Fort Hare Papers 1: 302 - 396." type="journal article" year="1951">Mokhehle, 1951</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="EEF580CDFFE8A558FCC4F9E5FB96F9FC" authority="Singh, 1952" authorityName="Singh" authorityYear="1952" box="[800,1090,1541,1568]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="micropus">
<emphasis id="1B81275CFFE8A558FCC4F9E5FC7AF9FC" box="[800,942,1543,1568]" italics="true" pageId="30" pageNumber="31">N. micropus</emphasis>
<bibRefCitation id="4D6486BFFFE8A558FC51F9E4FB96F9FC" author="Singh K. S." box="[949,1090,1541,1568]" pageId="30" pageNumber="31" pagination="1 - 72" refId="ref21294" refString="Singh K. S. 1952. Cestode parasites of birds. Indian Journal of Helminthology 4: 1 - 72." type="journal article" year="1952">Singh, 1952</bibRefCitation>
</taxonomicName>
by smaller rostellar hooks (
<bibRefCitation id="4D6486BFFFE8A558FF21F9C8FE57F998" author="Mokhehle" box="[197,387,1577,1604]" pageId="30" pageNumber="31" pagination="302 - 396" refId="ref20794" refString="Mokhehle 1951. Parasites of the swift Caffrapus caffer caffer (Roberts) with descriptions of one new genus and eight new species. Fort Hare Papers 1: 302 - 396." type="journal article" year="1951">Mokhehle 1951</bibRefCitation>
;
<bibRefCitation id="4D6486BFFFE8A558FE6BF9C8FDCDF998" author="Singh K. S." box="[399,537,1577,1604]" pageId="30" pageNumber="31" pagination="1 - 72" refId="ref21294" refString="Singh K. S. 1952. Cestode parasites of birds. Indian Journal of Helminthology 4: 1 - 72." type="journal article" year="1952">Singh 1952</bibRefCitation>
;
<bibRefCitation id="4D6486BFFFE8A558FDC1F9C8FD76F998" author="Baer J. G." box="[549,674,1577,1604]" pageId="30" pageNumber="31" refId="ref18637" refString="Baer J. G. 1959. Exploration des Parcs Nationaux du Congo Belge. Mission J. G. Baer - W. Gerber (1958). Helminthes parasites. Institut des Parcs Nationaux du Congo Belge, Bruxelles." type="book" year="1959">Baer 1959</bibRefCitation>
;
<bibRefCitation id="4D6486BFFFE8A558FD4AF9C8FC64F998" author="Dimitrova Y. &amp; Mariaux J. &amp; Georgiev B." box="[686,944,1578,1604]" pageId="30" pageNumber="31" pagination="61 - 82" refId="ref19193" refString="Dimitrova Y., Mariaux J. &amp; Georgiev B. 2017. Redescriptions of four Palaeotropical species of the cestode genus Notopentorchis Burt, 1938 (Cyclophyllidea: Paruterinidae). Zootaxa 4290: 61 - 82. https: // doi. org / 10.11646 / zootaxa. 4290.1.3" type="journal article" year="2017">
Dimitrova
<emphasis id="1B81275CFFE8A558FCCAF9C9FCBDF998" box="[814,873,1578,1604]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="294AFB4EFFE8A558FF59F98EFD95F85E" blockId="30.[189,1399,1644,1922]" pageId="30" pageNumber="31">
So far, no species of
<taxonomicName id="EEF580CDFFE8A558FE5BF98EFDA7F95A" authorityName="Burt" authorityYear="1938" box="[447,627,1644,1670]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE8A558FE5BF98EFDA7F95A" box="[447,627,1644,1670]" italics="true" pageId="30" pageNumber="31">Notopentorchis</emphasis>
</taxonomicName>
has been reported from
<collectingCountry id="51E2BBDEFFE8A558FC78F98EFBDFF95A" box="[924,1035,1644,1670]" name="Australia" pageId="30" pageNumber="31">Australia</collectingCountry>
. The white-throated needletail
<taxonomicName id="EEF580CDFFE8A558FF59F972FE01F976" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[189,469,1680,1706]" class="Aves" family="Apodidae" genus="Hirundapus" kingdom="Animalia" order="Apodiformes" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="caudacutus">
<emphasis id="1B81275CFFE8A558FF59F972FE01F976" box="[189,469,1680,1706]" italics="true" pageId="30" pageNumber="31">Hirundapus caudacutus</emphasis>
</taxonomicName>
, hosting the worms described here, however occurs in Central and
<collectingCountry id="51E2BBDEFFE8A558FB3EF972FA93F976" box="[1242,1351,1680,1706]" name="China" pageId="30" pageNumber="31">East Asia</collectingCountry>
and winters in Eastern
<collectingCountry id="51E2BBDEFFE8A558FE73F956FDD7F912" box="[407,515,1716,1742]" name="Australia" pageId="30" pageNumber="31">Australia</collectingCountry>
(
<bibRefCitation id="4D6486BFFFE8A558FDF5F956FD64F912" author="Lepage D." box="[529,688,1716,1742]" pageId="30" pageNumber="31" refId="ref20315" refString="Lepage D. 2017. Avibase - the World Bird Database. Available from http: // avibase. bsc-eoc. org [accessed 1 Nov. 2017]." type="url" year="2017">Lepage 2017</bibRefCitation>
), making this range extension logical. The single occurrence of
<taxonomicName id="EEF580CDFFE8A558FF05F93AFE41F92E" authorityName="Burt" authorityYear="1938" box="[225,405,1752,1778]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE8A558FF05F93AFE41F92E" box="[225,405,1752,1778]" italics="true" pageId="30" pageNumber="31">Notopentorchis</emphasis>
</taxonomicName>
in this host was reported as
<taxonomicName id="EEF580CDFFE8A558FD19F93BFBA1F92E" authority="(Krabbe, 1882)" baseAuthorityName="Krabbe" baseAuthorityYear="1882" box="[765,1141,1752,1778]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="vesiculigera">
<emphasis id="1B81275CFFE8A558FD19F93BFC67F92E" box="[765,947,1752,1778]" italics="true" pageId="30" pageNumber="31">N. vesiculigera</emphasis>
(Krabbe, 1882)
</taxonomicName>
by
<bibRefCitation id="4D6486BFFFE8A558FB4DF93AFAA3F92F" author="Oshmarin P." box="[1193,1399,1752,1779]" pageId="30" pageNumber="31" refId="ref21020" refString="Oshmarin P. 1963. Helminths of mammals and birds in the Primorsk region. Izdatelsvo Akademii Nauk SSSR, Moscow. [In Russian.]" type="book" year="1963">Oshmarin (1963)</bibRefCitation>
in 15 out 29 birds in the Primorye Region (South-Eastern
<collectingCountry id="51E2BBDEFFE8A558FC99F91EFC04F8CB" box="[893,976,1788,1815]" name="Russia" pageId="30" pageNumber="31">Russia</collectingCountry>
). Besides its European and African distribution,
<taxonomicName id="EEF580CDFFE8A558FEBCF8C3FDDAF8E6" baseAuthorityName="Krabbe" baseAuthorityYear="1882" box="[344,526,1824,1850]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="vesiculigera">
<emphasis id="1B81275CFFE8A558FEBCF8C3FDDAF8E6" box="[344,526,1824,1850]" italics="true" pageId="30" pageNumber="31">N. vesiculigera</emphasis>
</taxonomicName>
clearly differs from our material by its much larger hooks and higher number of testes. We conclude that the present material belongs to an undescribed species that we name
<taxonomicName id="EEF580CDFFE8A558FF59F88AFE0BF85E" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[189,479,1896,1922]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="30" pageNumber="31" phylum="Platyhelminthes" rank="species" species="musealis" status="sp. nov.">
<emphasis id="1B81275CFFE8A558FF59F88AFE0BF85E" box="[189,479,1896,1922]" italics="true" pageId="30" pageNumber="31">Notopentorchis musealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFE8A558FE02F88AFD95F85E" box="[486,577,1896,1922]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:affiliation id="A69BFC928FE183747A8C5AB25C3180E5">Natural History Museum of Geneva, CP 6434, 1211 Geneva 6, Switzerland. Department of Genetics and Evolution, University of Geneva, Boulevard d’Ivoy 4, 1205 Geneva, Switzerland. Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113 Sofia, Bulgaria.</mods:affiliation>
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<subSubSection id="61EFA8C5FFECA55CFD27FB5CFCA5FB05" box="[707,881,1214,1241]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="294AFB4EFFECA55CFD27FB5CFCA5FB05" blockId="26.[707,881,1214,1241]" box="[707,881,1214,1241]" pageId="26" pageNumber="27">
<heading id="72024C22FFECA55CFD27FB5CFCA5FB05" box="[707,881,1214,1241]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<taxonomicName id="EEF580CDFFECA55CFD27FB5CFCA5FB05" authorityName="Clark" authorityYear="1952" box="[707,881,1214,1241]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="sp.">
<emphasis id="1B81275CFFECA55CFD27FB5CFC9DFB04" bold="true" box="[707,841,1214,1240]" italics="true" pageId="26" pageNumber="27">Dictymetra</emphasis>
sp.
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="61EFA8C5FFECA55CFF59FB1DFCC8FAB0" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="294AFB4EFFECA55CFF59FB1DFE7EFAC5" blockId="26.[189,426,1279,1305]" box="[189,426,1279,1305]" pageId="26" pageNumber="27">
<heading id="72024C22FFECA55CFF59FB1DFE7EFAC5" bold="true" box="[189,426,1279,1305]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="1B81275CFFECA55CFF59FB1DFE7EFAC5" bold="true" box="[189,426,1279,1305]" pageId="26" pageNumber="27">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFECA55CFF59FACCFCC8FAB0" blockId="26.[189,1398,1324,1388]" pageId="26" pageNumber="27">
<collectingCountry id="51E2BBDEFFECA55CFF59FACCFEBCFA94" box="[189,360,1326,1352]" name="Australia" pageId="26" pageNumber="27">AUSTRALIA</collectingCountry>
:
<collectingRegion id="EB3135ACFFECA55CFE9DFACCFDD3FA94" box="[377,519,1326,1352]" country="Australia" name="Queensland" pageId="26" pageNumber="27">Queensland</collectingRegion>
, Redland Bay,
<geoCoordinate id="4CC19D89FFECA55CFD2CFACEFCE0FA94" box="[712,820,1324,1352]" degrees="27" direction="south" minutes="39" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="-27.65">27°39′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFECA55CFCA3FACEFC17FA95" box="[839,963,1324,1353]" degrees="153" direction="east" minutes="19" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="153.31667">153°19′ E</geoCoordinate>
, date unknown, Matthews (?) leg., fragments from 2 specs (AHC 35795, AHC 35796).
</paragraph>
</subSubSection>
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<emphasis id="1B81275CFFECA55CFF59FA71FF2CFA71" bold="true" box="[189,248,1427,1453]" pageId="26" pageNumber="27">Host</emphasis>
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<paragraph id="294AFB4EFFECA55CFF59FA23FC03FA00" blockId="26.[189,983,1473,1500]" box="[189,983,1473,1500]" pageId="26" pageNumber="27">
<taxonomicName id="EEF580CDFFECA55CFF59FA23FDB6FA00" authority="(Latham, 1802)" baseAuthorityName="Latham" baseAuthorityYear="1802" box="[189,610,1473,1500]" class="Aves" family="Burhinidae" genus="Burhinus" kingdom="Animalia" order="Charadriiformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="grallarius">
<emphasis id="1B81275CFFECA55CFF59FA23FE70FA07" box="[189,420,1473,1499]" italics="true" pageId="26" pageNumber="27">Burhinus grallarius</emphasis>
(Latham, 1802)
</taxonomicName>
(
<taxonomicName id="EEF580CDFFECA55CFD96FA23FCE3FA07" box="[626,823,1473,1499]" class="Aves" kingdom="Animalia" order="Charadriiformes" pageId="26" pageNumber="27" phylum="Chordata" rank="order">Charadriiformes</taxonomicName>
,
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).
</paragraph>
<paragraph id="294AFB4EFFECA55CFF59F9E1FE54F9C1" blockId="26.[189,384,1539,1565]" box="[189,384,1539,1565]" pageId="26" pageNumber="27">
<heading id="72024C22FFECA55CFF59F9E1FE54F9C1" bold="true" box="[189,384,1539,1565]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="1B81275CFFECA55CFF59F9E1FE54F9C1" bold="true" box="[189,384,1539,1565]" pageId="26" pageNumber="27">Site of infection</emphasis>
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</paragraph>
<paragraph id="294AFB4EFFECA55CFF59F9D3FEA0F997" blockId="26.[189,372,1585,1611]" box="[189,372,1585,1611]" pageId="26" pageNumber="27">Small intestine.</paragraph>
</subSubSection>
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<paragraph id="294AFB4EFFECA55CFF59F991FE9AF951" blockId="26.[189,334,1651,1677]" box="[189,334,1651,1677]" pageId="26" pageNumber="27">
<heading id="72024C22FFECA55CFF59F991FE9AF951" bold="true" box="[189,334,1651,1677]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="1B81275CFFECA55CFF59F991FE9AF951" bold="true" box="[189,334,1651,1677]" pageId="26" pageNumber="27">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFECA55CFF59F943FABEF853" blockId="26.[189,1399,1697,1935]" pageId="26" pageNumber="27">
Despite its similarity with the above described new species, this material probably belongs to another congeneric species. The available mounted specimens do not however allow for a detailed description. From the single scolex and few fragments available, the hook arrangement is similar to that of
<taxonomicName id="EEF580CDFFECA55CFF59F8EEFE52F8F9" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[189,390,1803,1829]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="longiuncinata">
<emphasis id="1B81275CFFECA55CFF59F8EEFE52F8F9" box="[189,390,1803,1829]" italics="true" pageId="26" pageNumber="27">D. longiuncinata</emphasis>
</taxonomicName>
but hooks are somewhat larger (about 145 long). The rostellar sac (320 × 140) and
<taxonomicName id="EEF580CDFFECA55CFF59F8CCFEF8F894" authorityName="Montfort" authorityYear="1810" box="[189,300,1838,1864]" family="Rostellum" pageId="26" pageNumber="18" rank="family">rostellum</taxonomicName>
(270 × 90) are smaller. It also has a cirrus armed with long setae but has a higher number of testes, i.e., 29–45 (37, n = 12) – and a smaller cirrus-sac, 107–113 × 25–28 (110 × 27, n = 4). It most probably belongs to
<taxonomicName id="EEF580CDFFECA55CFE49F897FDE5F852" authorityName="Clark" authorityYear="1952" box="[429,561,1909,1934]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFECA55CFE49F897FDE5F852" box="[429,561,1909,1934]" italics="true" pageId="26" pageNumber="27">Dictymetra</emphasis>
</taxonomicName>
but additional fresh material is obviously needed to confirm its status.
</paragraph>
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</treatment>
</document>

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sp.
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<collectingCountry id="51E2BBDEFFEDA55DFF58F94AFEB3F91E" box="[188,359,1704,1730]" name="Australia" pageId="27" pageNumber="28">AUSTRALIA</collectingCountry>
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, Kangaroo Grounds,
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,
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, 1979, Karl Harrigan leg. (AHC 36527).
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).
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<emphasis id="1B81275CFFEDA55DFF58F8BBFEFFF8AF" bold="true" box="[188,299,1881,1907]" pageId="27" pageNumber="28">Intensity</emphasis>
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About
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.
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<emphasis id="1B81275CFFEAA55AFF59FEE9FE9AFEF9" bold="true" box="[189,334,267,293]" pageId="28" pageNumber="29">Description</emphasis>
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Although several specimens of this species are available, including almost complete ones, just a single squashed scolex is present and the material is highly macerated, thus preventing a full description. Longest fragment
<quantity id="EE0D56ABFFEAA55AFE78FE62FE21FE46" box="[412,501,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="28" pageNumber="29" unit="mm" value="38.0">38 mm</quantity>
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at level of gravid proglottides. Ten hooks on a single crown, 24–25 long, with short blade, robust guard and slightly curved handle (
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). Three testes arranged in triangle, 2 antiporal. Cirrus-sac crossing osmoregulatory canals dorsally, about 165– 185 long. Genital openings unilateral, dextral. Both seminal vesicles and seminal receptacles inflating considerably. Uterus initially bisaccular, becoming unisaccular with deep lobes. Oncosphere 45–50. Embryonic hooks 20–22 (lateral), 22–24 (central). These few available characters hint toward the genus
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Spasskii &amp; Spasskaya, 1954 (Hymenolepididae)
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<taxonomicName id="EEF580CDFFEDA55DFD49FEE9FCE8FEF9" ID-CoL="7NH7H" authorityName="Fuhrmann" authorityYear="1907" box="[685,828,267,293]" class="Cestoda" family="Dilepididae" kingdom="Animalia" order="Cyclophyllidea" pageId="27" pageNumber="28" phylum="Platyhelminthes" rank="family">
<emphasis id="1B81275CFFEDA55DFD49FEE9FCE8FEF9" bold="true" box="[685,828,267,293]" pageId="27" pageNumber="28">Dilepididae</emphasis>
</taxonomicName>
indet.
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</paragraph>
<paragraph id="294AFB4EFFEDA55DFD0BFECCFC96FE95" blockId="27.[685,902,267,329]" box="[751,834,302,329]" pageId="27" pageNumber="28">
<figureCitation id="B1CEE7CBFFEDA55DFD0BFECCFC96FE95" box="[751,834,302,329]" captionStart="Figs 39–41. 39" captionStartId="35.[189,241,1941,1967]" captionTargetBox="[201,1367,1516,1875]" captionTargetId="figure@35.[195,1393,1513,1883]" captionTargetPageId="35" captionText="Figs 39–41. 39. Dilepididae genus sp. Hooks. 40. Passerilepis sp. Hooks. 41. Cracticotaenia fieldingi (Maplestone &amp; Southwell, 1923) Hooks. Scale bars: 39–41 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827570" httpUri="https://zenodo.org/record/3827570/files/figure.png" pageId="27" pageNumber="28">Fig. 39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFEDA55DFF58FE92FEC3FE00" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="294AFB4EFFEDA55DFF58FE92FE7EFE56" blockId="27.[188,426,368,394]" box="[188,426,368,394]" pageId="27" pageNumber="28">
<heading id="72024C22FFEDA55DFF58FE92FE7EFE56" bold="true" box="[188,426,368,394]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="1B81275CFFEDA55DFF58FE92FE7EFE56" bold="true" box="[188,426,368,394]" pageId="27" pageNumber="28">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEDA55DFF58FE7DFEC3FE00" blockId="27.[188,1398,413,476]" pageId="27" pageNumber="28">
<collectingCountry id="51E2BBDEFFEDA55DFF58FE7DFEB3FE65" box="[188,359,415,441]" name="Australia" pageId="27" pageNumber="28">AUSTRALIA</collectingCountry>
:
<collectingRegion id="EB3135ACFFEDA55DFE90FE7CFDD6FE64" box="[372,514,414,440]" country="Australia" name="Queensland" pageId="27" pageNumber="28">Queensland</collectingRegion>
, Eidsvold,
<geoCoordinate id="4CC19D89FFEDA55DFD6CFE7FFD24FE65" box="[648,752,413,441]" degrees="25" direction="south" minutes="22" orientation="latitude" pageId="27" pageNumber="28" precision="925" value="-25.366667">25°22′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFEDA55DFD1BFE7FFCBDFE65" box="[767,873,413,441]" degrees="151" direction="east" minutes="5" orientation="longitude" pageId="27" pageNumber="28" precision="925" value="151.08333">151°5′ E</geoCoordinate>
,
<date id="5D4BDD8EFFEDA55DFC9CFE7DFC25FE65" box="[888,1009,415,441]" pageId="27" pageNumber="28" value="1918-09">Sep. 1918</date>
, Thomas L. Bancroft leg. (AHC 36528).
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFEDA55DFF58FDE1FE71FCF0" pageId="27" pageNumber="28" type="biology_ecology">
<paragraph id="294AFB4EFFEDA55DFF58FDE1FF23FDC1" blockId="27.[188,247,515,541]" box="[188,247,515,541]" pageId="27" pageNumber="28">
<heading id="72024C22FFEDA55DFF58FDE1FF23FDC1" bold="true" box="[188,247,515,541]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="1B81275CFFEDA55DFF58FDE1FF23FDC1" bold="true" box="[188,247,515,541]" pageId="27" pageNumber="28">Host</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEDA55DFF58FDD0FBF7FD91" blockId="27.[188,1059,562,589]" box="[188,1059,562,589]" pageId="27" pageNumber="28">
<taxonomicName id="EEF580CDFFEDA55DFF58FDD0FD64FD90" authority="(Vieillot, 1817)" baseAuthorityName="Vieillot" baseAuthorityYear="1817" box="[188,688,562,588]" class="Aves" family="Corcoracidae" genus="Corcorax" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="melanorhynchos">
<emphasis id="1B81275CFFEDA55DFF58FDD0FE20FD90" box="[188,500,562,588]" italics="true" pageId="27" pageNumber="28">Corcorax melanorhynchos</emphasis>
(Vieillot, 1817)
</taxonomicName>
(
<taxonomicName id="EEF580CDFFEDA55DFD5BFDD0FCBDFD90" box="[703,873,562,588]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="28" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
,
<taxonomicName id="EEF580CDFFEDA55DFC91FDD0FBC3FD91" box="[885,1047,562,589]" class="Aves" family="Corcoracidae" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="28" phylum="Chordata" rank="family">Corcoracidae</taxonomicName>
).
</paragraph>
<paragraph id="294AFB4EFFEDA55DFF58FD91FE54FD51" blockId="27.[188,384,627,653]" box="[188,384,627,653]" pageId="27" pageNumber="28">
<heading id="72024C22FFEDA55DFF58FD91FE54FD51" bold="true" box="[188,384,627,653]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="1B81275CFFEDA55DFF58FD91FE54FD51" bold="true" box="[188,384,627,653]" pageId="27" pageNumber="28">Site of infection</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEDA55DFF58FD40FEA7FD60" blockId="27.[188,371,674,700]" box="[188,371,674,700]" pageId="27" pageNumber="28">Small intestine.</paragraph>
<paragraph id="294AFB4EFFEDA55DFF58FD01FEFFFD21" blockId="27.[188,299,739,765]" box="[188,299,739,765]" pageId="27" pageNumber="28">
<heading id="72024C22FFEDA55DFF58FD01FEFFFD21" bold="true" box="[188,299,739,765]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="1B81275CFFEDA55DFF58FD01FEFFFD21" bold="true" box="[188,299,739,765]" pageId="27" pageNumber="28">Intensity</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEDA55DFF58FCF0FE71FCF0" blockId="27.[188,421,786,812]" box="[188,421,786,812]" pageId="27" pageNumber="28">
About
<specimenCount id="3FF330C7FFEDA55DFEE9FCF0FE75FCF0" box="[269,417,786,812]" pageId="27" pageNumber="28" type="generic">8 specimens</specimenCount>
.
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<paragraph id="294AFB4EFFEDA55DFF58FCB1FE99FCB1" blockId="27.[188,333,851,877]" box="[188,333,851,877]" pageId="27" pageNumber="28">
<heading id="72024C22FFEDA55DFF58FCB1FE99FCB1" bold="true" box="[188,333,851,877]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="1B81275CFFEDA55DFF58FCB1FE99FCB1" bold="true" box="[188,333,851,877]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFEDA55DFF58FC60FEFAFA54" blockId="27.[188,1398,898,1416]" pageId="27" pageNumber="28">
A few fragmented specimens of this species are found as co-parasites of
<taxonomicName id="EEF580CDFFEDA55DFBA2FC60FAA2FC40" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[1094,1398,898,924]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="27" pageNumber="28" phylum="Platyhelminthes" rank="species" species="adelaidae" status="sp. nov.">
<emphasis id="1B81275CFFEDA55DFBA2FC60FAA2FC40" box="[1094,1398,898,924]" italics="true" pageId="27" pageNumber="28">Cracticotaenia adelaidae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFEDA55DFF58FC47FECAFC63" box="[188,286,933,959]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
in our material from
<collectingRegion id="EB3135ACFFEDA55DFDDBFC47FD19FC63" box="[575,717,933,959]" country="Australia" name="Queensland" pageId="27" pageNumber="28">Queensland</collectingRegion>
. Unfortunately, hooks are lost on the single available scolex. Longest fragment
<quantity id="EE0D56ABFFEDA55DFE12FC2AFD9AFC3F" box="[502,590,968,995]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="28" unit="mm" value="13.0">13 mm</quantity>
, maximum width
<quantity id="EE0D56ABFFEDA55DFCCEFC2AFC53FC3F" box="[810,903,968,995]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="28" unit="mm" value="1.4">1.4 mm</quantity>
at level of gravid proglottides. Up to 61 craspedote proglottides, wider than long except for the most gravid ones. Scolex not clearly separated from strobila,
<quantity id="EE0D56ABFFEDA55DFE8CFBEDFE61FBF4" box="[360,437,1038,1065]" metricMagnitude="0" metricUnit="m" metricValue="5.7658" pageId="27" pageNumber="28" unit="in" value="227.0">227 in</quantity>
diameter. Suckers rounded
<quantity id="EE0D56ABFFEDA55DFCE2FBECFCA9FBF4" box="[774,893,1038,1065]" metricMagnitude="0" metricUnit="m" metricValue="2.6924" metricValueMax="2.921" metricValueMin="2.4638" pageId="27" pageNumber="28" unit="in" value="106.0" valueMax="115.0" valueMin="97.0">97–115 in</quantity>
diameter. Rostellar pouch very glandular, 167 × 70, extending past posterior margin of suckers.
<taxonomicName id="EEF580CDFFEDA55DFCDBFBD0FC6CFB90" authorityName="Montfort" authorityYear="1810" box="[831,952,1074,1100]" family="Rostellum" pageId="27" pageNumber="18" rank="family">Rostellum</taxonomicName>
muscular, with well-marked terminal pad, 132 × 72. Neck not marked. Genital pores, opening laterally and very anteriorly, at about 15% of lateral proglottis margin, alternating irregularly in short series. Ventral osmoregulatory canals connected posteriorly in each proglottis by transverse anastomosis. Genital ducts passing between osmoregulatory canals. Testes 17–23 (19.7, n = 26) in number, in a single layer and one continuous posterior field, extending antero laterally up to level of vitellarium. Cirrus-sac short and weak, 76–92 × 30–42, not reaching poral osmoregulatory canals. Cirrus armed with a small tuft of setae, often lost in our material. Vitellarium central, compact. Ovary anterior transversely elongated. Seminal receptacle rounded, lateral to vitellarium. Vagina posterior to male pore. Uterus initially labyrinthic. No fully gravid proglottides observed.
</paragraph>
</subSubSection>
</treatment>
</document>

370
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<mods:affiliation id="1794EEE788AC33AFCC6A6E990D3FD570">Natural History Museum of Geneva, CP 6434, 1211 Geneva 6, Switzerland. Department of Genetics and Evolution, University of Geneva, Boulevard d’Ivoy 4, 1205 Geneva, Switzerland. Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113 Sofia, Bulgaria.</mods:affiliation>
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<emphasis id="1B81275CFFF1A541FD83F93EFCBFF92B" bold="true" box="[615,875,1756,1783]" italics="true" pageId="7" pageNumber="8">Spiniglans beveridgei</emphasis>
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<paragraph id="294AFB4EFFF1A541FD4AF8C0FC52F8E1" blockId="7.[367,1220,1756,1853]" box="[686,902,1826,1853]" pageId="7" pageNumber="8">
<figureCitation id="B1CEE7CBFFF1A541FD4AF8C0FCF7F8E1" box="[686,803,1826,1853]" captionStart="Figs 7–11" captionStartId="9.[189,241,1915,1941]" captionTargetBox="[203,1391,278,1872]" captionTargetId="figure@9.[186,1391,261,1886]" captionTargetPageId="9" captionText="Figs 7–11. Spiniglans beveridgei sp. nov. 7. Scolex. 8. Hooks. 9. Mature proglottis. 10. Terminal genitalia. 11. Pregravid proglottis with young reticular uterus. Scale bars: 7 = 100 µm; 8 = 20 µm; 9 = 200 µm; 10 = 50 µm; 11 = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3827558" httpUri="https://zenodo.org/record/3827558/files/figure.png" pageId="7" pageNumber="8">Figs 7–11</figureCitation>
,
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</subSubSection>
<subSubSection id="61EFA8C5FFF1A541FF59F886FC1CF80C" pageId="7" pageNumber="8" type="etymology">
<paragraph id="294AFB4EFFF1A541FF59F886FE90F8A2" blockId="7.[189,324,1892,1918]" box="[189,324,1892,1918]" pageId="7" pageNumber="8">
<heading id="72024C22FFF1A541FF59F886FE90F8A2" bold="true" box="[189,324,1892,1918]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="1B81275CFFF1A541FF59F886FE90F8A2" bold="true" box="[189,324,1892,1918]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFF1A541FF59F870FC1CF80C" blockId="7.[189,1399,1938,2000]" pageId="7" pageNumber="8">The species is dedicated to Professor Ian Beveridge, a friend and colleague, collector of this material, in recognition of his major contribution to systematic helminthology.</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFEA54EFF59FEE9FBF7FD60" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="294AFB4EFFFEA54EFF59FEE9FE7EFEF9" blockId="8.[189,426,267,293]" box="[189,426,267,293]" pageId="8" pageNumber="9">
<heading id="72024C22FFFEA54EFF59FEE9FE7EFEF9" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="1B81275CFFFEA54EFF59FEE9FE7EFEF9" bold="true" box="[189,426,267,293]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="999DF113FFFEA54EFF0EFED8FABDFEAB" ID-GBIF-Occurrence="2619826304" collectingDate="2001-04-22" collectorName="Ian Beveridge" country="Australia" latitude="-38.133335" location="Geelong" longLatPrecision="1177" longitude="144.36667" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
<paragraph id="294AFB4EFFFEA54EFF0EFED8FE88FE88" blockId="8.[189,1385,314,375]" box="[234,348,314,340]" pageId="8" pageNumber="9">
<emphasis id="1B81275CFFFEA54EFF0EFED8FE88FE88" bold="true" box="[234,348,314,340]" pageId="8" pageNumber="9">
<typeStatus id="F64E45ECFFFEA54EFF0EFED8FE88FE88" box="[234,348,314,340]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FEBFFABDFEAB" blockId="8.[189,1385,314,375]" box="[189,1385,347,375]" pageId="8" pageNumber="9">
<collectingCountry id="51E2BBDEFFFEA54EFF59FEBFFEBCFEAB" box="[189,360,349,375]" name="Australia" pageId="8" pageNumber="9">AUSTRALIA</collectingCountry>
:
<collectingRegion id="EB3135ACFFFEA54EFE96FEBFFE00FEAB" box="[370,468,349,375]" country="Australia" name="Victoria" pageId="8" pageNumber="9">Victoria</collectingRegion>
,
<location id="2C2AAD95FFFEA54EFE3BFEBFFD91FEAB" LSID="urn:lsid:plazi:treatment:A15C4A58FFF1A54CFD83F93EFC31FAC3:2C2AAD95FFFEA54EFE3BFEBFFD91FEAB" box="[479,581,349,375]" country="Australia" latitude="-38.133335" longLatPrecision="1177" longitude="144.36667" name="Geelong" pageId="8" pageNumber="9" stateProvince="Victoria">Geelong</location>
,
<geoCoordinate id="4CC19D89FFFEA54EFDB5FEB9FD7DFEAB" box="[593,681,347,375]" degrees="38" direction="south" minutes="8" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-38.133335">38°8′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFFEA54EFD52FEB9FCF9FEAB" box="[694,813,347,375]" degrees="144" direction="east" minutes="22" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="144.36667">144°22′ E</geoCoordinate>
,
<date id="5D4BDD8EFFFEA54EFCDDFEBFFC00FEAB" box="[825,980,349,375]" pageId="8" pageNumber="9" value="2001-04-22">
<collectingDate id="4D0F2466FFFEA54EFCDDFEBFFC00FEAB" box="[825,980,349,375]" pageId="8" pageNumber="9" value="2001-04-22">22 Apr. 2001</collectingDate>
</date>
,
<collectorName id="84009E98FFFEA54EFC04FEBFFB52FEAB" box="[992,1158,349,375]" pageId="8" pageNumber="9">Ian Beveridge</collectorName>
leg. (AHC 36467).
</paragraph>
</materialsCitation>
<paragraph id="294AFB4EFFFEA54EFF0EFE7CFEB3FE64" blockId="8.[189,1051,414,476]" box="[234,359,414,440]" pageId="8" pageNumber="9">
<emphasis id="1B81275CFFFEA54EFF0EFE7CFEB3FE64" bold="true" box="[234,359,414,440]" pageId="8" pageNumber="9">
<typeStatus id="F64E45ECFFFEA54EFF0EFE7CFEB3FE64" box="[234,359,414,440]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FE23FBCFFE00" blockId="8.[189,1051,414,476]" box="[189,1051,449,476]" pageId="8" pageNumber="9">
<collectingCountry id="51E2BBDEFFFEA54EFF59FE23FEBCFE07" box="[189,360,449,475]" name="Australia" pageId="8" pageNumber="9">AUSTRALIA</collectingCountry>
: 10 specs, same data as for
<typeStatus id="F64E45ECFFFEA54EFD4FFE23FCC6FE07" box="[683,786,449,475]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(AHC 36468–36477).
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FDE1FEE2FDC1" blockId="8.[189,310,515,541]" box="[189,310,515,541]" pageId="8" pageNumber="9">
<heading id="72024C22FFFEA54EFF59FDE1FEE2FDC1" bold="true" box="[189,310,515,541]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="1B81275CFFFEA54EFF59FDE1FEE2FDC1" bold="true" box="[189,310,515,541]" pageId="8" pageNumber="9">Type host</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FDD0FCA0FD90" blockId="8.[189,884,561,588]" box="[189,884,561,588]" pageId="8" pageNumber="9">
<taxonomicName id="EEF580CDFFFEA54EFF59FDD0FDE6FD97" authority="Matthews, 1912" authorityName="Matthews" authorityYear="1912" box="[189,562,561,588]" class="Aves" family="Corvidae" genus="Corvus" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="mellori">
<emphasis id="1B81275CFFFEA54EFF59FDD0FEBAFD97" box="[189,366,561,588]" italics="true" pageId="8" pageNumber="9">Corvus mellori</emphasis>
Matthews, 1912
</taxonomicName>
(
<taxonomicName id="EEF580CDFFFEA54EFDA5FDD3FD3FFD97" box="[577,747,561,587]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
,
<taxonomicName id="EEF580CDFFFEA54EFD12FDD3FCB3FD90" box="[758,871,561,588]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="family">Corvidae</taxonomicName>
).
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FD90FEF8FD50" blockId="8.[189,300,626,652]" box="[189,300,626,652]" pageId="8" pageNumber="9">
<heading id="72024C22FFFEA54EFF59FD90FEF8FD50" bold="true" box="[189,300,626,652]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="1B81275CFFFEA54EFF59FD90FEF8FD50" bold="true" box="[189,300,626,652]" pageId="8" pageNumber="9">Intensity</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FD43FBF7FD60" blockId="8.[189,1059,673,700]" box="[189,1059,673,700]" pageId="8" pageNumber="9">
About
<specimenCount id="3FF330C7FFFEA54EFEEAFD43FE78FD67" box="[270,428,673,700]" pageId="8" pageNumber="9" type="generic">55 specimens</specimenCount>
in a single host, together with
<taxonomicName id="EEF580CDFFFEA54EFCF2FD40FC16FD67" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[790,962,673,700]" class="Cestoda" family="Dilepididae" genus="Sobolevitaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="whittingtoni" status="sp. nov.">
<emphasis id="1B81275CFFFEA54EFCF2FD40FC16FD67" box="[790,962,673,700]" italics="true" pageId="8" pageNumber="9">
<collectionCode id="4FE4638BFFFEA54EFCF2FD40FCFCFD60" box="[790,808,674,700]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="8" pageNumber="9" type="Herbarium">S</collectionCode>
. whittingtoni
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFFEA54EFC2DFD40FBF7FD60" box="[969,1059,674,700]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFEA54CFF59FD00FA96FE94" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="294AFB4EFFFEA54EFF59FD00FE9AFD20" blockId="8.[189,334,738,764]" box="[189,334,738,764]" pageId="8" pageNumber="9">
<heading id="72024C22FFFEA54EFF59FD00FE9AFD20" bold="true" box="[189,334,738,764]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="1B81275CFFFEA54EFF59FD00FE9AFD20" bold="true" box="[189,334,738,764]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FCF3FD19FA79" blockId="8.[189,1399,785,1445]" pageId="8" pageNumber="9">
Body of medium size with length
<quantity id="EE0D56ABFFFEA54EFDBFFCF3FD08FCF0" box="[603,732,785,812]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="6.3" metricValueMin="5.7" pageId="8" pageNumber="9" unit="mm" value="60.0" valueMax="63.0" valueMin="57.0">57–63 mm</quantity>
and maximum width 1050–1280 (1170, n = 6) at level of late mature proglottides. Most complete late mature to pregravid specimens consisting of 169–198 proglottides. Proglottides craspedote, initially wider than long, progressively becoming longer, square at mature stage and longer than wide at late mature stage. Ventral osmoregulatory canals connected posteriorly in each proglottis by transverse anastomosis. Longitudinal musculature with well-marked bundles. Scolex (
<figureCitation id="B1CEE7CBFFFEA54EFE66FC23FE12FC00" box="[386,454,961,988]" captionStart="Figs 7–11" captionStartId="9.[189,241,1915,1941]" captionTargetBox="[203,1391,278,1872]" captionTargetId="figure@9.[186,1391,261,1886]" captionTargetPageId="9" captionText="Figs 7–11. Spiniglans beveridgei sp. nov. 7. Scolex. 8. Hooks. 9. Mature proglottis. 10. Terminal genitalia. 11. Pregravid proglottis with young reticular uterus. Scale bars: 7 = 100 µm; 8 = 20 µm; 9 = 200 µm; 10 = 50 µm; 11 = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3827558" httpUri="https://zenodo.org/record/3827558/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
) rounded, well marked, sometimes showing a transverse fold behind suckers, with diameter 325–450 (397, n = 6). Suckers strong, rounded,
<quantity id="EE0D56ABFFFEA54EFC86FC06FC33FC22" box="[866,999,996,1023]" metricMagnitude="0" metricUnit="m" metricValue="4.1274999999999995" metricValueMax="4.572" metricValueMin="3.683" pageId="8" pageNumber="9" unit="in" value="162.5" valueMax="180.0" valueMin="145.0">145–180 in</quantity>
diameter (161, n = 24), unarmed. Rostellar apparatus musculo-glandular, with developed glandular tissue along whole rostellar sac length often slightly protruding apically even when retracted. Rostellar sac well-delineated, 374–450 × 132– 152 (383 × 142, n = 6) tapering posteriorly, extending beyond level of posterior margin of suckers, often with terminal bubble-like extension.
<taxonomicName id="EEF580CDFFFEA54EFD94FB93FD3DFB57" authorityName="Montfort" authorityYear="1810" box="[624,745,1137,1163]" family="Rostellum" pageId="8" pageNumber="18" rank="family">Rostellum</taxonomicName>
oval, strongly muscular, with well-marked constriction behind hooks, 224–270 × 85–105 (242 × 94, n = 6). Rostellar hooks (
<figureCitation id="B1CEE7CBFFFEA54EFC02FB76FBF8FB72" box="[998,1068,1172,1198]" captionStart="Figs 7–11" captionStartId="9.[189,241,1915,1941]" captionTargetBox="[203,1391,278,1872]" captionTargetId="figure@9.[186,1391,261,1886]" captionTargetPageId="9" captionText="Figs 7–11. Spiniglans beveridgei sp. nov. 7. Scolex. 8. Hooks. 9. Mature proglottis. 10. Terminal genitalia. 11. Pregravid proglottis with young reticular uterus. Scale bars: 7 = 100 µm; 8 = 20 µm; 9 = 200 µm; 10 = 50 µm; 11 = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3827558" httpUri="https://zenodo.org/record/3827558/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
) in two mostly regular rows (with occasional slight irregularities or indistinct row delineation), 20 (n = 1), 21 (n = 3) or 22 (n = 4) in number, robust, with strong guard and slightly bent tapering extremity of handle. Anterior hooks 40.5–44 (42, n = 8), posterior hooks 38.5–40.5 (39.5, n = 6). Proglottization distinct at 500–650 (568, n = 7) from posterior margin of suckers Genital pores anterior situated at about 20% of length of lateral proglottis margin, irregularly alternating in short series, e.g., …1, 1, 2, 2, 1, 1, 3, 1, 1, 2, 2, 3, 1, 4, 5, 3, 1, 1, 2... No more than five consecutive pores observed on single side. Genital ducts passing between osmoregulatory canals. Genital atrium small.
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59FA2FFCFAF901" blockId="8.[189,1399,1484,1757]" pageId="8" pageNumber="9">
Testes 40–58 (49, n = 54) in number (
<quantity id="EE0D56ABFFFEA54EFD9DFA2EFD37FA3A" box="[633,739,1484,1511]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" metricValueMax="1.3969999999999998" metricValueMin="1.0922" pageId="8" pageNumber="9" unit="in" value="49.0" valueMax="55.0" valueMin="43.0">43–55 in</quantity>
90% of counts), in one or two layers, in one continuous posterior file, denser posteriorly, mostly extending antero-laterally up to level of vitellarium but a few may be more anterior, especially at antiporal side, occasionally overlapping osmoregulatory canals (
<figureCitation id="B1CEE7CBFFFEA54EFF22F9D4FEDAF98C" box="[198,270,1590,1616]" captionStart="Figs 7–11" captionStartId="9.[189,241,1915,1941]" captionTargetBox="[203,1391,278,1872]" captionTargetId="figure@9.[186,1391,261,1886]" captionTargetPageId="9" captionText="Figs 7–11. Spiniglans beveridgei sp. nov. 7. Scolex. 8. Hooks. 9. Mature proglottis. 10. Terminal genitalia. 11. Pregravid proglottis with young reticular uterus. Scale bars: 7 = 100 µm; 8 = 20 µm; 9 = 200 µm; 10 = 50 µm; 11 = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3827558" httpUri="https://zenodo.org/record/3827558/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
). External vas deferens
<quantity id="EE0D56ABFFFEA54EFDC8F9D4FD7EF98C" box="[556,682,1590,1617]" metricMagnitude="-1" metricUnit="m" metricValue="4.3815" metricValueMax="5.334" metricValueMin="3.429" pageId="8" pageNumber="9" unit="in" value="17.25" valueMax="21.0" valueMin="13.5">13.5–21 in</quantity>
diameter (17, n = 15), highly convoluted in the antero-poral part of proglottis, not overlapping cirrus-sac. Cirrus-sac short, oval or bean-shaped, 87–118 × 47–68 (102 × 55, n = 44), transverse or slightly curved anteriorly, rarely reaching poral osmoregulatory canals. Internal vas deferens coiled, making several large loops within cirrus-sac. Cirrus armed with compact tuft of fine bristle-like spines about 35 long (
<figureCitation id="B1CEE7CBFFFEA54EFD2DF920FCF4F901" box="[713,800,1730,1757]" captionStart="Figs 7–11" captionStartId="9.[189,241,1915,1941]" captionTargetBox="[203,1391,278,1872]" captionTargetId="figure@9.[186,1391,261,1886]" captionTargetPageId="9" captionText="Figs 7–11. Spiniglans beveridgei sp. nov. 7. Scolex. 8. Hooks. 9. Mature proglottis. 10. Terminal genitalia. 11. Pregravid proglottis with young reticular uterus. Scale bars: 7 = 100 µm; 8 = 20 µm; 9 = 200 µm; 10 = 50 µm; 11 = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3827558" httpUri="https://zenodo.org/record/3827558/files/figure.png" pageId="8" pageNumber="9">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="294AFB4EFFFEA54EFF59F8E6FAA2F870" blockId="8.[189,1399,1796,1964]" pageId="8" pageNumber="9">
Vitellarium central, transversely elongated, lobate, variable in shape. Ovary transversely elongated, deeply and finely branched, lobulate, bi-alate, antiporal wing much larger than poral one. Mehlis’ gland 70–105 (84, n = 15) closely anterior to vitellarium. Seminal receptacle initially round, rapidly becoming oval, reaching up to 175 × 114, dorsal, posterior to external vas deferens, often over poral ovary wing. Vagina thick-walled, along posterior margin of cirrus-sac, mostly straight
<quantity id="EE0D56ABFFFEA54EFBA7F873FB1FF877" box="[1091,1227,1937,1964]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.921" metricValueMin="1.905" pageId="8" pageNumber="9" unit="in" value="9.5" valueMax="11.5" valueMin="7.5">7.5–11.5 in</quantity>
diameter (9.5
</paragraph>
<caption id="7D8AABC6FFFFA54FFF59F899FD58F800" ID-DOI="http://doi.org/10.5281/zenodo.3827558" ID-Zenodo-Dep="3827558" httpUri="https://zenodo.org/record/3827558/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,241,1915,1941]" targetBox="[203,1391,278,1872]" targetPageId="9">
<paragraph id="294AFB4EFFFFA54FFF59F899FD58F800" blockId="9.[189,1399,1915,2012]" pageId="9" pageNumber="10">
<emphasis id="1B81275CFFFFA54FFF59F899FEEAF849" bold="true" box="[189,318,1915,1941]" pageId="9" pageNumber="10">Figs 7–11.</emphasis>
<taxonomicName id="EEF580CDFFFFA54FFEAFF899FD84F849" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[331,592,1915,1941]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="beveridgei" status="sp. nov.">
<emphasis id="1B81275CFFFFA54FFEAFF899FD84F849" box="[331,592,1915,1941]" italics="true" pageId="9" pageNumber="10">Spiniglans beveridgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFFFA54FFDB8F89EFD68F84A" box="[604,700,1916,1942]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1B81275CFFFFA54FFD2DF899FD0CF849" bold="true" box="[713,728,1915,1941]" pageId="9" pageNumber="10">7</emphasis>
. Scolex.
<emphasis id="1B81275CFFFFA54FFCB4F899FC8BF849" bold="true" box="[848,863,1915,1941]" pageId="9" pageNumber="10">8</emphasis>
. Hooks.
<emphasis id="1B81275CFFFFA54FFC37F899FC36F849" bold="true" box="[979,994,1915,1941]" pageId="9" pageNumber="10">9</emphasis>
. Mature proglottis.
<emphasis id="1B81275CFFFFA54FFB38F899FB2DF849" bold="true" box="[1244,1273,1915,1941]" pageId="9" pageNumber="10">10</emphasis>
. Terminal genitalia.
<emphasis id="1B81275CFFFFA54FFED0F87CFE9AF864" bold="true" box="[308,334,1950,1976]" pageId="9" pageNumber="10">11</emphasis>
. Pregravid proglottis with young reticular uterus. Scale bars: 7 = 100 µm; 8 = 20 µm; 9 = 200 µm; 10 = 50 µm; 11 = 500 µm.
</paragraph>
</caption>
<paragraph id="294AFB4EFFFCA54CFF59FEEEFA96FE94" blockId="10.[189,1399,267,329]" pageId="10" pageNumber="11">
n = 12), externally gained with loose parenchymal cells. Vaginal pore posterior to pore of cirrus-sac. Young uterus reticular (
<figureCitation id="B1CEE7CBFFFCA54CFE36FECCFDF2FE95" box="[466,550,302,329]" captionStart="Figs 7–11" captionStartId="9.[189,241,1915,1941]" captionTargetBox="[203,1391,278,1872]" captionTargetId="figure@9.[186,1391,261,1886]" captionTargetPageId="9" captionText="Figs 7–11. Spiniglans beveridgei sp. nov. 7. Scolex. 8. Hooks. 9. Mature proglottis. 10. Terminal genitalia. 11. Pregravid proglottis with young reticular uterus. Scale bars: 7 = 100 µm; 8 = 20 µm; 9 = 200 µm; 10 = 50 µm; 11 = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3827558" httpUri="https://zenodo.org/record/3827558/files/figure.png" pageId="10" pageNumber="11">Fig. 11</figureCitation>
) in pregravid proglottides. Gravid segments and eggs not observed.
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFCA54CFF59FE92FC31FAC3" pageId="10" pageNumber="11" type="discussion">
<paragraph id="294AFB4EFFFCA54CFF59FE92FEFBFE56" blockId="10.[189,303,368,394]" box="[189,303,368,394]" pageId="10" pageNumber="11">
<heading id="72024C22FFFCA54CFF59FE92FEFBFE56" bold="true" box="[189,303,368,394]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="1B81275CFFFCA54CFF59FE92FEFBFE56" bold="true" box="[189,303,368,394]" pageId="10" pageNumber="11">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFCA54CFF59FE7CFE19FD99" blockId="10.[189,1399,414,581]" pageId="10" pageNumber="11">
The genus
<taxonomicName id="EEF580CDFFFCA54CFEAEFE7DFD46FE65" authority="Yamaguti, 1959" authorityName="Yamaguti" authorityYear="1959" box="[330,658,414,441]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFFCA54CFEAEFE7DFE12FE65" box="[330,454,415,441]" italics="true" pageId="10" pageNumber="11">Spiniglans</emphasis>
Yamaguti, 1959
</taxonomicName>
has been discussed by
<bibRefCitation id="4D6486BFFFFCA54CFC24FE7DFB88FE65" author="Bona F. V." box="[960,1116,415,441]" pageId="10" pageNumber="11" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">
<collectingCountry id="51E2BBDEFFFCA54CFC24FE7DFC2AFE65" box="[960,1022,415,441]" name="Algeria" pageId="10" pageNumber="11">Bona</collectingCountry>
(
<date id="5D4BDD8EFFFCA54CFBF0FE7DFB87FE65" box="[1044,1107,415,441]" pageId="10" pageNumber="11" value="1994">1994</date>
)
</bibRefCitation>
,
<bibRefCitation id="4D6486BFFFFCA54CFB8AFE7CFAEAFE65" author="Salamatin R. V." box="[1134,1342,414,441]" pageId="10" pageNumber="11" pagination="75 - 81" refId="ref21194" refString="Salamatin R. V. 1999. The cestodes of the genus Spiniglans (Dilepididae) from the corvid birds (Corvidae) from Ukraine. Vestnik Zoologii 33: 75 - 81. [In Russian.]" type="journal article" year="1999">
Salamatin (
<date id="5D4BDD8EFFFCA54CFB1DFE7DFAE2FE65" box="[1273,1334,415,441]" pageId="10" pageNumber="11" value="1999">1999</date>
)
</bibRefCitation>
and
<bibRefCitation id="4D6486BFFFFCA54CFF59FE20FE3DFE00" author="Kornyushin V. V. &amp; Salamatin R. V. &amp; Greben O. B. &amp; Georgiev B. B." box="[189,489,450,476]" pageId="10" pageNumber="11" pagination="85 - 93" refId="ref20121" refString="Kornyushin V. V., Salamatin R. V., Greben O. B. &amp; Georgiev B. B. 2009. Spiniglans sharpiloi sp. n. (Cestoda, Dilepididae), a parasite of the common magpie, Pica pica, in the Palaearctic. Vestnik Zoologii Supplement 23: 85 - 93." type="journal article" year="2009">
Kornyushin
<emphasis id="1B81275CFFFCA54CFEB5FE21FE52FE00" box="[337,390,450,476]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. (
<date id="5D4BDD8EFFFCA54CFE7BFE20FE34FE00" box="[415,480,450,476]" pageId="10" pageNumber="11" value="2009">2009</date>
)
</bibRefCitation>
. The latter authors briefly reviewed the generic characters and listed 4 valid species, from
<taxonomicName id="EEF580CDFFFCA54CFE85FE07FE19FE23" box="[353,461,485,511]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Corvidae</taxonomicName>
in the Old World, as well a few other closely related taxa possibly belonging to the genus. They also predicted that more taxa in this group would be found both in other European and tropical corvid species.
</paragraph>
<paragraph id="294AFB4EFFFCA54CFF59FD8FFE4FFC7D" blockId="10.[189,1399,621,929]" pageId="10" pageNumber="11">
Our material can easily be distinguished from most presently recognized species in the genus (
<tableCitation id="6477CEF5FFFCA54CFAE9FD8FFABCFD5B" box="[1293,1384,621,647]" captionStart="Table 2" captionStartId="11.[188,257,686,714]" captionText="Table 2. Comparison of Spiniglans beveridgei sp. nov. with the other species of the genus Spiniglans Yamaguti, 1959." pageId="10" pageNumber="11">Table 2</tableCitation>
). It differs from
<taxonomicName id="EEF580CDFFFCA54CFE8BFD72FD3AFD76" authority="(Southwell, 1922)" baseAuthorityName="Southwell" baseAuthorityYear="1922" box="[367,750,656,682]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="microsoma">
<emphasis id="1B81275CFFFCA54CFE8BFD72FDC4FD76" box="[367,528,656,682]" italics="true" pageId="10" pageNumber="11">S. microsoma</emphasis>
(
<bibRefCitation id="4D6486BFFFFCA54CFDC5FD72FD32FD76" author="Southwell T." box="[545,742,656,682]" pageId="10" pageNumber="11" pagination="355 - 382" refId="ref21349" refString="Southwell T. 1922. Cestodes from Indian birds with a note on Ligula intestinalis. Annals of Tropical Medicine &amp; Parasitology 16: 355 - 382. https: // doi. org / 10.1080 / 00034983.1922.11684330" type="journal article" year="1922">Southwell, 1922</bibRefCitation>
)
</taxonomicName>
by having more testes; from
<taxonomicName id="EEF580CDFFFCA54CFBB5FD72FAA2FD77" authority="(Krabbe, 1869)" baseAuthorityName="Krabbe" baseAuthorityYear="1869" box="[1105,1398,656,683]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="affinis">
<emphasis id="1B81275CFFFCA54CFBB5FD72FB6DFD76" box="[1105,1209,656,682]" italics="true" pageId="10" pageNumber="11">S. affinis</emphasis>
(
<bibRefCitation id="4D6486BFFFFCA54CFB2DFD72FABAFD77" author="Krabbe H." box="[1225,1390,656,683]" pageId="10" pageNumber="11" pagination="251 - 389" refId="ref20181" refString="Krabbe H. 1869. Bidrag til kundskab om fuglenes baendelorme. Videnskabelige Selskab Skrifter, 5. raekke, Naturvidenskabelig og Mathematisk Afdel 8: 251 - 389." type="journal article" year="1869">Krabbe, 1869</bibRefCitation>
)
</taxonomicName>
by smaller rostellar hooks, fewer testes and a smaller
<taxonomicName id="EEF580CDFFFCA54CFCA3FD51FC6EFD11" authorityName="Montfort" authorityYear="1810" box="[839,954,691,717]" family="Rostellum" pageId="10" pageNumber="18" rank="family">rostellum</taxonomicName>
; from
<taxonomicName id="EEF580CDFFFCA54CFBEEFD56FA9BFD12" authority="(Molin, 1858)" baseAuthorityName="Molin" baseAuthorityYear="1858" box="[1034,1359,691,718]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="constricta">
<emphasis id="1B81275CFFFCA54CFBEEFD56FB4BFD11" box="[1034,1183,692,718]" italics="true" pageId="10" pageNumber="11">S. constricta</emphasis>
(Molin, 1858)
</taxonomicName>
by a larger cirrus-sac, rostellar pouch and
<taxonomicName id="EEF580CDFFFCA54CFD7DFD34FCD8FD2C" authorityName="Montfort" authorityYear="1810" box="[665,780,726,752]" family="Rostellum" pageId="10" pageNumber="18" rank="family">rostellum</taxonomicName>
; from
<taxonomicName id="EEF580CDFFFCA54CFCBAFD35FED5FCC8" authority="(Georgiev, Kornyushin &amp; Genov, 1987)" baseAuthorityName="Georgiev, Kornyushin &amp; Genov" baseAuthorityYear="1987" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="pirinica">
<emphasis id="1B81275CFFFCA54CFCBAFD35FC0FFD2C" box="[862,987,727,753]" italics="true" pageId="10" pageNumber="11">S. pirinica</emphasis>
(
<bibRefCitation id="4D6486BFFFFCA54CFC0AFD34FF2DFCC8" author="Georgiev B. B. &amp; Kornyushin V. V. &amp; Genov T." pageId="10" pageNumber="11" pagination="3 - 7" refId="ref19658" refString="Georgiev B. B., Kornyushin V. V. &amp; Genov T. 1987. Choanotaenia pirinica sp. n. (Cestoda, Dilepididae) - a parasite of the Alpine Chough in Bulgaria. Vestnik Zoologii 1987: 3 - 7. [In Russian.]" type="journal article" year="1987">Georgiev, Kornyushin &amp; Genov, 1987</bibRefCitation>
)
</taxonomicName>
by much shorter hooks and a smaller cirrus-sac. Our material most closely resembles
<taxonomicName id="EEF580CDFFFCA54CFB0BFD18FE01FCEB" authority="Kornyushin et al., 2009" authorityName="Kornyushin" authorityYear="2009" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="sharpiloi">
<emphasis id="1B81275CFFFCA54CFB0BFD18FAA2FCC8" box="[1263,1398,762,788]" italics="true" pageId="10" pageNumber="11">S. sharpiloi</emphasis>
<bibRefCitation id="4D6486BFFFFCA54CFF59FCFFFE01FCEB" author="Kornyushin V. V. &amp; Salamatin R. V. &amp; Greben O. B. &amp; Georgiev B. B." box="[189,469,797,823]" pageId="10" pageNumber="11" pagination="85 - 93" refId="ref20121" refString="Kornyushin V. V., Salamatin R. V., Greben O. B. &amp; Georgiev B. B. 2009. Spiniglans sharpiloi sp. n. (Cestoda, Dilepididae), a parasite of the common magpie, Pica pica, in the Palaearctic. Vestnik Zoologii Supplement 23: 85 - 93." type="journal article" year="2009">
Kornyushin
<emphasis id="1B81275CFFFCA54CFEABFCFCFE50FCEB" box="[335,388,797,823]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
., 2009
</bibRefCitation>
</taxonomicName>
but can be distinguished from it by slightly larger rostellar hooks and a longer rostellar pouch and
<taxonomicName id="EEF580CDFFFCA54CFE46FCA2FDC1FC86" authorityName="Montfort" authorityYear="1810" box="[418,533,832,858]" family="Rostellum" pageId="10" pageNumber="18" rank="family">rostellum</taxonomicName>
. It is also similar to
<taxonomicName id="EEF580CDFFFCA54CFD18FCA2FB17FC86" authority="(Joyeux, Baer &amp; Martin, 1937)" baseAuthorityName="Joyeux, Baer &amp; Martin" baseAuthorityYear="1937" box="[764,1219,832,858]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="corvi">
<emphasis id="1B81275CFFFCA54CFD18FCA2FC81FC86" box="[764,853,832,858]" italics="true" pageId="10" pageNumber="11">S. corvi</emphasis>
(
<bibRefCitation id="4D6486BFFFFCA54CFC80FCA2FB6FFC86" author="Joyeux C. &amp; Baer J. G. &amp; Martin R." box="[868,1211,832,858]" pageId="10" pageNumber="11" pagination="416 - 423" refId="ref19954" refString="Joyeux C., Baer J. G. &amp; Martin R. 1937. Sur quelques helminthes de la Somalie-Nord (Deuxieme note). Bulletin de la Societe de Pathologie exotique 30: 416 - 423." type="journal article" year="1937">Joyeux, Baer &amp; Martin, 1937</bibRefCitation>
)
</taxonomicName>
but it has more testes and a shorter cirrus-sac. In consequence, we consider it to belong to a new species,
<taxonomicName id="EEF580CDFFFCA54CFB1FFC81FEEDFC7C" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="beveridgei" status="sp. nov.">
<emphasis id="1B81275CFFFCA54CFB1FFC81FEEDFC7C" italics="true" pageId="10" pageNumber="11">Spiniglans beveridgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFFCA54CFEA4FC65FE4FFC7D" box="[320,411,903,929]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="294AFB4EFFFCA54CFF59FC2AFC31FAC3" blockId="10.[189,1399,968,1311]" pageId="10" pageNumber="11">
Up to now,
<taxonomicName id="EEF580CDFFFCA54CFEACFC2AFE10FC3E" authorityName="Yamaguti" authorityYear="1959" box="[328,452,968,994]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFFCA54CFEACFC2AFE10FC3E" box="[328,452,968,994]" italics="true" pageId="10" pageNumber="11">Spiniglans</emphasis>
</taxonomicName>
was known from the Old World only and its host range essentially limited to the
<taxonomicName id="EEF580CDFFFCA54CFF59FC09FEFDFBD9" box="[189,297,1003,1029]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Corvidae</taxonomicName>
although the
<typeStatus id="F64E45ECFFFCA54CFE29FC0EFDD4FBDA" box="[461,512,1004,1030]" pageId="10" pageNumber="11">type</typeStatus>
species was reported in granivore passerines (
<taxonomicName id="EEF580CDFFFCA54CFBCCFC09FB6AFBD9" authorityName="Vigors" authorityYear="1831" box="[1064,1214,1003,1029]" class="Aves" family="Emberizidae" kingdom="Animalia" order="Passeriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Emberizidae</taxonomicName>
and
<taxonomicName id="EEF580CDFFFCA54CFB1FFC09FABBFBDA" authorityName="Sundevall" authorityYear="1836" box="[1275,1391,1003,1030]" class="Aves" family="Ploceidae" kingdom="Animalia" order="Passeriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Ploceidae</taxonomicName>
) in captivity (
<bibRefCitation id="4D6486BFFFFCA54CFEB0FBECFDC1FBF4" author="Southwell T." box="[340,533,1038,1064]" pageId="10" pageNumber="11" pagination="355 - 382" refId="ref21349" refString="Southwell T. 1922. Cestodes from Indian birds with a note on Ligula intestinalis. Annals of Tropical Medicine &amp; Parasitology 16: 355 - 382. https: // doi. org / 10.1080 / 00034983.1922.11684330" type="journal article" year="1922">Southwell 1922</bibRefCitation>
). Its presence in
<collectingCountry id="51E2BBDEFFFCA54CFD3BFBECFC9FFBF4" box="[735,843,1038,1064]" name="Australia" pageId="10" pageNumber="11">Australia</collectingCountry>
may be due to a recent importation as
<taxonomicName id="EEF580CDFFFCA54CFAC6FBEDFAA3FBF5" box="[1314,1399,1039,1065]" class="Aves" family="Corvidae" genus="Corvus" kingdom="Animalia" order="Passeriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="1B81275CFFFCA54CFAC6FBEDFAA3FBF5" box="[1314,1399,1039,1065]" italics="true" pageId="10" pageNumber="11">Corvus</emphasis>
</taxonomicName>
colonized (or re-colonized) this continent after diversifying in the Palaearctic (
<bibRefCitation id="4D6486BFFFFCA54CFB65FBD3FAA6FB90" author="Ericson P. G. &amp; Jansen A. - L. &amp; Johansson U. S. &amp; Ekman J." box="[1153,1394,1073,1100]" pageId="10" pageNumber="11" pagination="222 - 234" refId="ref19244" refString="Ericson P. G., Jansen A. - L., Johansson U. S. &amp; Ekman J. 2005. Inter-generic relationships of the crows, jays, magpies and allied groups (Aves: Corvidae) based on nucleotide sequence data. Journal of Avian Biology 36: 222 - 234. https: // doi. org / 10.1111 / j. 0908 - 8857.2001.03409. x" type="journal article" year="2005">
Ericson
<emphasis id="1B81275CFFFCA54CFB0DFBD0FAFCFB97" box="[1257,1320,1073,1099]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2005
</bibRefCitation>
;
<bibRefCitation id="4D6486BFFFFCA54CFF59FBB7FE70FBB3" author="Haring E. &amp; Daubl B. &amp; Pinsker W. &amp; Kryukov A. &amp; Gamauf A." box="[189,420,1109,1135]" pageId="10" pageNumber="11" pagination="230 - 246" refId="ref19709" refString="Haring E., Daubl B., Pinsker W., Kryukov A. &amp; Gamauf A. 2012. Genetic divergences and intraspecific variation in corvids of the genus Corvus (Aves: Passeriformes: Corvidae) - a first survey based on museum specimens. Journal of Zoological Systematics and Evolutionary Research 50: 230 - 246. https: // doi. org / 10.1111 / j. 1439 - 0469.2012.00664. x" type="journal article" year="2012">
Haring
<emphasis id="1B81275CFFFCA54CFEFEFBB7FE8DFBB3" box="[282,345,1109,1135]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2012
</bibRefCitation>
). The
<taxonomicName id="EEF580CDFFFCA54CFE17FBB7FD8BFBB3" box="[499,607,1109,1135]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Corvidae</taxonomicName>
includes around 130 species and is cosmopolitan. It is therefore expected that a larger diversity of parasites of this genus is still unknown, especially in Asia. It remains to be seen whether New World corvids are also hosts of these cestodes, as hinted by the brief report of
<bibRefCitation id="4D6486BFFFFCA54CFF04FB5CFD99FB04" author="Pfaffenberger G. S. &amp; Butler W. F." box="[224,589,1214,1240]" pageId="10" pageNumber="11" pagination="563 - 566" refId="ref21047" refString="Pfaffenberger G. S. &amp; Butler W. F. 1981. Helminths recovered from the white-necked raven (Corvus cryptoleucus Couch) in Eastern New Mexico. Journal of Wildlife Diseases 17: 563 - 566. https: // doi. org / 10.7589 / 0090 - 3558 - 17.4.563" type="journal article" year="1981">Pfaffenberger &amp; Butler (1981)</bibRefCitation>
of
<taxonomicName id="EEF580CDFFFCA54CFD9EFB5DFCDBFB04" baseAuthorityName="Molin" baseAuthorityYear="1858" box="[634,783,1215,1241]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="constricta">
<emphasis id="1B81275CFFFCA54CFD9EFB5DFCDBFB04" box="[634,783,1215,1241]" italics="true" pageId="10" pageNumber="11">S. constricta</emphasis>
</taxonomicName>
(“
<emphasis id="1B81275CFFFCA54CFCD4FB5DFBEBFB04" box="[816,1087,1215,1241]" italics="true" pageId="10" pageNumber="11">
Anomotania
<taxonomicName id="EEF580CDFFFCA54CFC2EFB5DFBEBFB04" baseAuthorityName="Molin" baseAuthorityYear="1858" box="[970,1087,1215,1240]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="constricta">constricta</taxonomicName>
</emphasis>
”) in
<taxonomicName id="EEF580CDFFFCA54CFB66FB5DFE81FB20" authority="Couch, 1854" authorityName="Couch" authorityYear="1854" class="Aves" family="Corvidae" genus="Corvus" kingdom="Animalia" order="Passeriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cryptoleucus">
<emphasis id="1B81275CFFFCA54CFB66FB5DFAA3FB04" box="[1154,1399,1214,1241]" italics="true" pageId="10" pageNumber="11">Corvus cryptoleucus</emphasis>
Couch,
<quantity id="EE0D56ABFFFCA54CFEFEFB03FE81FB20" box="[282,341,1249,1276]" metricMagnitude="1" metricUnit="m" metricValue="4.70916" pageId="10" pageNumber="11" unit="in" value="1854.0">1854</quantity>
</taxonomicName>
in New Mexico. If confirmed, many more
<taxonomicName id="EEF580CDFFFCA54CFCB8FB03FC0CFB27" authorityName="Yamaguti" authorityYear="1959" box="[860,984,1249,1275]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFFCA54CFCB8FB03FC0CFB27" box="[860,984,1249,1275]" italics="true" pageId="10" pageNumber="11">Spiniglans</emphasis>
</taxonomicName>
spp. are likely to be discovered in American corvids as the family is highly diversified in the Americas.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="294AFB4EFFF9A549FD49FA78FC53FA68" blockId="15.[685,903,1434,1460]" box="[685,903,1434,1460]" pageId="15" pageNumber="16">
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<taxonomicName id="EEF580CDFFF9A549FD49FA78FCE8FA68" ID-CoL="7NH7H" authorityName="Fuhrmann" authorityYear="1907" box="[685,828,1434,1460]" class="Cestoda" family="Dilepididae" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="family">
<emphasis id="1B81275CFFF9A549FD49FA78FCE8FA68" bold="true" box="[685,828,1434,1460]" pageId="15" pageNumber="16">Dilepididae</emphasis>
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indet.
</heading>
</paragraph>
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<subSubSection id="61EFA8C5FFF9A549FF59FA3FFDCFF992" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="294AFB4EFFF9A549FF59FA3FFE7EFA2B" blockId="15.[189,426,1501,1527]" box="[189,426,1501,1527]" pageId="15" pageNumber="16">
<heading id="72024C22FFF9A549FF59FA3FFE7EFA2B" bold="true" box="[189,426,1501,1527]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="1B81275CFFF9A549FF59FA3FFE7EFA2B" bold="true" box="[189,426,1501,1527]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
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<paragraph id="294AFB4EFFF9A549FF59F9ECFDCFF992" blockId="15.[189,1399,1548,1614]" pageId="15" pageNumber="16">
<collectingCountry id="51E2BBDEFFF9A549FF59F9ECFEBCF9F4" box="[189,360,1550,1576]" name="Australia" pageId="15" pageNumber="16">AUSTRALIA</collectingCountry>
: about 12 specs,
<collectingRegion id="EB3135ACFFF9A549FDACF9ECFD7EF9F4" box="[584,682,1550,1576]" country="Australia" name="Victoria" pageId="15" pageNumber="16">Victoria</collectingRegion>
, Healesville Sanctuary,
<geoCoordinate id="4CC19D89FFF9A549FC33F9EEFB90F9F4" box="[983,1092,1548,1576]" degrees="37" direction="south" minutes="41" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-37.683334">37°41′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFF9A549FBBCF9EEFB01F9F5" box="[1112,1237,1548,1577]" degrees="145" direction="east" minutes="32" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="145.53334">145°32′ E</geoCoordinate>
, 2002, Dave Middleton leg. (AHC 36488).
</paragraph>
</subSubSection>
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<heading id="72024C22FFF9A549FF59F995FF2CF94D" bold="true" box="[189,248,1655,1681]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="1B81275CFFF9A549FF59F995FF2CF94D" bold="true" box="[189,248,1655,1681]" pageId="15" pageNumber="16">Host</emphasis>
</heading>
</paragraph>
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<taxonomicName id="EEF580CDFFF9A549FF59F94BFD5BF91E" authority="Latham, 1801" authorityName="Latham" authorityYear="1801" box="[189,655,1704,1730]" class="Aves" family="Menuridae" genus="Menura" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="novaehollandiae">
<emphasis id="1B81275CFFF9A549FF59F94BFE31F91E" box="[189,485,1704,1730]" italics="true" pageId="15" pageNumber="16">Menura novaehollandiae</emphasis>
Latham, 1801
</taxonomicName>
(syn.
<taxonomicName id="EEF580CDFFF9A549FD33F94BFBD5F91E" authority="Davies, 1802" authorityName="Davies" authorityYear="1802" box="[727,1025,1704,1730]" class="Aves" family="Menuridae" genus="Menura" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="superba">
<emphasis id="1B81275CFFF9A549FD33F94BFC8AF91E" box="[727,862,1704,1730]" italics="true" pageId="15" pageNumber="16">M. superba</emphasis>
Davies, 1802
</taxonomicName>
) (
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,
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).
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<emphasis id="1B81275CFFF9A549FF59F909FE9AF8D9" bold="true" box="[189,334,1771,1797]" pageId="15" pageNumber="16">Description</emphasis>
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This material is in poor state of conservation with contracted and (or) macerated partial strobila and no intact scolex. Scolex width 300–370 (338, n = 3) in diameter; suckers muscular 125–208 (164, n = 12) in diameter.
<taxonomicName id="EEF580CDFFF9A549FEB0F885FE19F85D" authorityName="Montfort" authorityYear="1810" box="[340,461,1895,1921]" family="Rostellum" pageId="15" pageNumber="18" rank="family">Rostellum</taxonomicName>
225–255 × 85–137. A single scolex bears an incomplete series of hooks with length 21–25 (23.5, n = 8) (
<figureCitation id="B1CEE7CBFFF9A549FDF7F86EFDBAF87A" box="[531,622,1932,1958]" captionStart="Figs 39–41. 39" captionStartId="35.[189,241,1941,1967]" captionTargetBox="[201,1367,1516,1875]" captionTargetId="figure@35.[195,1393,1513,1883]" captionTargetPageId="35" captionText="Figs 39–41. 39. Dilepididae genus sp. Hooks. 40. Passerilepis sp. Hooks. 41. Cracticotaenia fieldingi (Maplestone &amp; Southwell, 1923) Hooks. Scale bars: 39–41 = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.3827570" httpUri="https://zenodo.org/record/3827570/files/figure.png" pageId="15" pageNumber="16">Fig. 39</figureCitation>
). Worms are much larger and wider than
<taxonomicName id="EEF580CDFFF9A549FB8EF86FFAF7F87A" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[1130,1315,1932,1958]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="species" species="australiense">
<emphasis id="1B81275CFFF9A549FB8EF86FFAF7F87A" box="[1130,1315,1932,1958]" italics="true" pageId="15" pageNumber="16">M. australiense</emphasis>
</taxonomicName>
(up to
<quantity id="EE0D56ABFFF9A549FF59F853FEC4F810" box="[189,272,1969,1996]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="16" unit="mm" value="26.0">26 mm</quantity>
long and
<quantity id="EE0D56ABFFF9A549FE62F853FE34F810" box="[390,480,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="16" unit="mm" value="2.6">2.6 mm</quantity>
wide observed, although complete worms should be even larger). The overall anatomy is similar, in particular the few observable rostellar hooks are of the same size and shape as in
<taxonomicName id="EEF580CDFFE6A556FF59FED0FEA6FE97" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[189,370,305,331]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="16" pageNumber="17" phylum="Platyhelminthes" rank="species" species="australiense">
<emphasis id="1B81275CFFE6A556FF59FED0FEA6FE97" box="[189,370,305,331]" italics="true" pageId="16" pageNumber="17">M. australiense</emphasis>
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, but, among the observable characters, it seems to have more testes (ca 35–40), a vagina lined with setae and a slightly larger cirrus-sac. The cirrus is armed with well-marked small spines. The available material does not allow for a description but hints at the possible diversity of
<taxonomicName id="EEF580CDFFE6A556FB0AFE99FAA3FE49" authorityName="Fuhrmann" authorityYear="1907" box="[1262,1399,379,405]" class="Cestoda" family="Dilepididae" kingdom="Animalia" order="Cyclophyllidea" pageId="16" pageNumber="17" phylum="Platyhelminthes" rank="family">Dilepididae</taxonomicName>
within lyrebirds.
</paragraph>
<paragraph id="294AFB4EFFE6A556FD82FE00FC1AFE21" blockId="16.[614,974,482,509]" box="[614,974,482,509]" pageId="16" pageNumber="17">
Genus
<taxonomicName id="EEF580CDFFE6A556FD5CFE01FC1AFE21" authority="Clark, 1952" authorityName="Clark" authorityYear="1952" box="[696,974,482,509]" class="Cestoda" family="Dilepididae" genus="Dictymetra" kingdom="Animalia" order="Cyclophyllidea" pageId="16" pageNumber="17" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFE6A556FD5CFE01FCE8FE20" box="[696,828,483,508]" italics="true" pageId="16" pageNumber="17">Dictymetra</emphasis>
<bibRefCitation id="4D6486BFFFE6A556FCA7FE00FC1AFE21" author="Clark D. T." box="[835,974,482,509]" pageId="16" pageNumber="17" pagination="18 - 27" refId="ref18941" refString="Clark D. T. 1952. Three new dilepidid cestodes, Dictymetra numenii n. gen. sp. nov.; Dictymetra paranumenii sp. nov. and Anomotaenia filovata sp. nov. Proceedings of the Helminthological Society of Washington 19: 18 - 27." type="journal article" year="1952">Clark, 1952</bibRefCitation>
</taxonomicName>
</paragraph>
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</treatment>
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399
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</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFAA54AFD42FC47FC5AFC63" box="[678,910,932,959]" pageId="12" pageNumber="13" type="description">
<paragraph id="294AFB4EFFFAA54AFD42FC47FC5AFC63" blockId="12.[366,1222,862,959]" box="[678,910,932,959]" pageId="12" pageNumber="13">
<figureCitation id="B1CEE7CBFFFAA54AFD42FC47FCFFFC62" box="[678,811,932,959]" captionStart="Figs 12–16" captionStartId="14.[189,241,1943,1969]" captionTargetBox="[218,1382,269,1894]" captionTargetId="figure@14.[203,1384,265,1915]" captionTargetPageId="14" captionText="Figs 12–16. Monopylidium australiense sp. nov. 12. Scolex. 13. Hooks. 14. Mature proglottis. 15. Terminal genitalia. 16. Eggs. Scale bars: 12, 14 = 100 µm; 13, 16 = 20 µm; 15 = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3827560" httpUri="https://zenodo.org/record/3827560/files/figure.png" pageId="12" pageNumber="13">Figs 12–16</figureCitation>
,
<tableCitation id="6477CEF5FFFAA54AFCD3FC47FC5AFC63" box="[823,910,933,959]" captionStart="Table 3" captionStartId="17.[188,257,542,570]" captionTargetBox="[199,1940,691,1741]" captionText="Table 3 (continued on next page). Comparison of Monopylidium australiense sp. nov. with the other species of the genera Monopylidium Fuhrmann, 1899 and Icterotaenia Railliet &amp; Henry, 1909. See text for the accepted generic concept. Note: * 20, 10 and 19 respectively in original reference, a printing error, as confirmed by the length of cirrus-sac on the original illustration." httpUri="http://table.plazi.org/id/7D8AABC6FFE7A557FF58FDFCFBACFD5C" pageId="12" pageNumber="13" tableUuid="7D8AABC6FFE7A557FF58FDFCFBACFD5C">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFAA54AFF59FC04FB97FBF3" box="[189,1091,998,1071]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="294AFB4EFFFAA54AFF59FC04FE90FBDC" blockId="12.[189,324,998,1024]" box="[189,324,998,1024]" pageId="12" pageNumber="13">
<heading id="72024C22FFFAA54AFF59FC04FE90FBDC" bold="true" box="[189,324,998,1024]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="1B81275CFFFAA54AFF59FC04FE90FBDC" bold="true" box="[189,324,998,1024]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59FBF7FB97FBF3" blockId="12.[189,1091,1045,1071]" box="[189,1091,1045,1071]" pageId="12" pageNumber="13">The specific name is derived from the country where the material was found.</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFAA54AFF59FBB4FDB8F899" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="294AFB4EFFFAA54AFF59FBB4FE7EFBAC" blockId="12.[189,426,1110,1136]" box="[189,426,1110,1136]" pageId="12" pageNumber="13">
<heading id="72024C22FFFAA54AFF59FBB4FE7EFBAC" bold="true" box="[189,426,1110,1136]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="1B81275CFFFAA54AFF59FBB4FE7EFBAC" bold="true" box="[189,426,1110,1136]" pageId="12" pageNumber="13">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="999DF113FFFAA54AFF0EFB67FDC0FB39" ID-GBIF-Occurrence="2619826307" collectingDate="1994-10-04" collectorName="Peter Holz" country="Australia" latitude="-37.683334" location="Healesville Sanctuary" longLatPrecision="1180" longitude="145.53334" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
<paragraph id="294AFB4EFFFAA54AFF0EFB67FE88FB43" blockId="12.[189,1399,1157,1253]" box="[234,348,1157,1183]" pageId="12" pageNumber="13">
<emphasis id="1B81275CFFFAA54AFF0EFB67FE88FB43" bold="true" box="[234,348,1157,1183]" pageId="12" pageNumber="13">
<typeStatus id="F64E45ECFFFAA54AFF0EFB67FE88FB43" box="[234,348,1157,1183]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59FB4AFDC0FB39" blockId="12.[189,1399,1157,1253]" pageId="12" pageNumber="13">
<collectingCountry id="51E2BBDEFFFAA54AFF59FB4AFEBCFB1E" box="[189,360,1192,1218]" name="Australia" pageId="12" pageNumber="13">AUSTRALIA</collectingCountry>
:
<collectingRegion id="EB3135ACFFFAA54AFE96FB4AFE00FB1E" box="[370,468,1192,1218]" country="Australia" name="Victoria" pageId="12" pageNumber="13">Victoria</collectingRegion>
,
<location id="2C2AAD95FFFAA54AFE3BFB4AFD34FB1E" LSID="urn:lsid:plazi:treatment:A15C4A58FFFAA549FDA7FCBCFF22FAAC:2C2AAD95FFFAA54AFE3BFB4AFD34FB1E" box="[479,736,1192,1218]" country="Australia" latitude="-37.683334" longLatPrecision="1180" longitude="145.53334" name="Healesville Sanctuary" pageId="12" pageNumber="13" stateProvince="Victoria">Healesville Sanctuary</location>
,
<geoCoordinate id="4CC19D89FFFAA54AFD08FB44FC86FB1E" box="[748,850,1190,1218]" degrees="37" direction="south" minutes="41" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="-37.683334">37°41′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFFAA54AFCBAFB44FC01FB1E" box="[862,981,1190,1218]" degrees="145" direction="east" minutes="32" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="145.53334">145°32′ E</geoCoordinate>
,
<date id="5D4BDD8EFFFAA54AFC05FB4AFBB9FB1E" box="[993,1133,1192,1218]" pageId="12" pageNumber="13" value="1994-10-04">
<collectingDate id="4D0F2466FFFAA54AFC05FB4AFBB9FB1E" box="[993,1133,1192,1218]" pageId="12" pageNumber="13" value="1994-10-04">4 Oct. 1994</collectingDate>
</date>
,
<collectorName id="84009E98FFFAA54AFB9DFB4AFB21FB1E" box="[1145,1269,1192,1218]" pageId="12" pageNumber="13">Peter Holz</collectorName>
leg. (AHC 36480, field number 94/773).
</paragraph>
</materialsCitation>
<paragraph id="294AFB4EFFFAA54AFF0EFAEFFEB3FAFB" blockId="12.[189,1135,1293,1355]" box="[234,359,1293,1319]" pageId="12" pageNumber="13">
<emphasis id="1B81275CFFFAA54AFF0EFAEFFEB3FAFB" bold="true" box="[234,359,1293,1319]" pageId="12" pageNumber="13">
<typeStatus id="F64E45ECFFFAA54AFF0EFAEFFEB3FAFB" box="[234,359,1293,1319]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59FAD2FBBBFA97" blockId="12.[189,1135,1293,1355]" box="[189,1135,1328,1355]" pageId="12" pageNumber="13">
<collectingCountry id="51E2BBDEFFFAA54AFF59FAD2FEBCFA96" box="[189,360,1328,1354]" name="Australia" pageId="12" pageNumber="13">AUSTRALIA</collectingCountry>
: about 24 specs, same datas as for
<typeStatus id="F64E45ECFFFAA54AFD1BFAD2FCB2FA96" box="[767,870,1328,1354]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(AHC 36481–36486).
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF0EFA93FE72FA57" blockId="12.[189,1399,1393,1525]" box="[234,422,1393,1419]" pageId="12" pageNumber="13">
<emphasis id="1B81275CFFFAA54AFF0EFA93FE72FA57" bold="true" box="[234,422,1393,1419]" pageId="12" pageNumber="13">Other material</emphasis>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59FA76FD2DFA29" blockId="12.[189,1399,1393,1525]" pageId="12" pageNumber="13">
<collectingCountry id="51E2BBDEFFFAA54AFF59FA76FEBCFA72" box="[189,360,1428,1454]" name="Australia" pageId="12" pageNumber="13">AUSTRALIA</collectingCountry>
: 2 specs,
<collectingRegion id="EB3135ACFFFAA54AFE31FA76FDE2FA72" box="[469,566,1428,1454]" country="Australia" name="Victoria" pageId="12" pageNumber="13">Victoria</collectingRegion>
, Healesville,
<date id="5D4BDD8EFFFAA54AFD31FA76FCB2FA72" box="[725,870,1428,1454]" pageId="12" pageNumber="13" value="1991-08-03">3 Aug. 1991</date>
, David Middleton leg. (AHC 22931, 36487);
<specimenCount id="3FF330C7FFFAA54AFF59FA5AFEC4FA0E" box="[189,272,1464,1490]" pageId="12" pageNumber="13" type="generic">1 spec.</specimenCount>
,
<collectingRegion id="EB3135ACFFFAA54AFEFFFA55FEA8FA0D" box="[283,380,1463,1489]" country="Australia" name="Victoria" pageId="12" pageNumber="13">Victoria</collectingRegion>
, Sherbrooke,
<geoCoordinate id="4CC19D89FFFAA54AFDF8FA54FD55FA0E" box="[540,641,1462,1490]" degrees="37" direction="south" minutes="53" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="-37.883335">37°53′ S</geoCoordinate>
,
<geoCoordinate id="4CC19D89FFFAA54AFD69FA54FCD7FA0E" box="[653,771,1462,1490]" degrees="145" direction="east" minutes="22" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="145.36667">145°22′ E</geoCoordinate>
, 2005, Peter Holz leg (AHC 29271);
<specimenCount id="3FF330C7FFFAA54AFB57FA5AFAD2FA0E" box="[1203,1286,1464,1490]" pageId="12" pageNumber="13" type="generic">1 spec.</specimenCount>
,
<collectingRegion id="EB3135ACFFFAA54AFAF5FA55FAA7FA0D" box="[1297,1395,1463,1489]" country="Australia" name="Victoria" pageId="12" pageNumber="13">Victoria</collectingRegion>
, Healesville, 2006, Peter Holz leg. (AHC 29281).
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59F9FEFEE2F9EA" blockId="12.[189,310,1564,1590]" box="[189,310,1564,1590]" pageId="12" pageNumber="13">
<heading id="72024C22FFFAA54AFF59F9FEFEE2F9EA" bold="true" box="[189,310,1564,1590]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="1B81275CFFFAA54AFF59F9FEFEE2F9EA" bold="true" box="[189,310,1564,1590]" pageId="12" pageNumber="13">Type host</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59F9AEFA88F9BA" blockId="12.[189,1372,1611,1638]" box="[189,1372,1611,1638]" pageId="12" pageNumber="13">
<taxonomicName id="EEF580CDFFFAA54AFF59F9AEFD5BF9B9" authority="Latham, 1801" authorityName="Latham" authorityYear="1801" box="[189,655,1611,1637]" class="Aves" family="Menuridae" genus="Menura" kingdom="Animalia" order="Passeriformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="novaehollandiae">
<emphasis id="1B81275CFFFAA54AFF59F9AEFE31F9B9" box="[189,485,1611,1637]" italics="true" pageId="12" pageNumber="13">Menura novaehollandiae</emphasis>
Latham, 1801
</taxonomicName>
(syn.
<taxonomicName id="EEF580CDFFFAA54AFD33F9AEFC2BF9B9" authority="Dacies, 1820" authorityName="Dacies" authorityYear="1820" box="[727,1023,1611,1637]" class="Aves" family="Menuridae" genus="Menura" kingdom="Animalia" order="Passeriformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="superba">
<emphasis id="1B81275CFFFAA54AFD33F9AEFC8AF9B9" box="[727,862,1611,1637]" italics="true" pageId="12" pageNumber="13">M. superba</emphasis>
Dacies, 1820
</taxonomicName>
) (
<taxonomicName id="EEF580CDFFFAA54AFBF2F9A9FB14F9B9" box="[1046,1216,1611,1637]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="12" pageNumber="13" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
,
<taxonomicName id="EEF580CDFFFAA54AFB28F9A9FA85F9BA" box="[1228,1361,1611,1638]" class="Aves" family="Menuridae" kingdom="Animalia" order="Passeriformes" pageId="12" pageNumber="13" phylum="Chordata" rank="family">Menuridae</taxonomicName>
).
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59F96EFE54F97A" blockId="12.[189,384,1676,1702]" box="[189,384,1676,1702]" pageId="12" pageNumber="13">
<heading id="72024C22FFFAA54AFF59F96EFE54F97A" bold="true" box="[189,384,1676,1702]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="1B81275CFFFAA54AFF59F96EFE54F97A" bold="true" box="[189,384,1676,1702]" pageId="12" pageNumber="13">Site of infection</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59F959FEA0F909" blockId="12.[189,372,1723,1749]" box="[189,372,1723,1749]" pageId="12" pageNumber="13">Small intestine.</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59F91EFEF8F8CA" blockId="12.[189,300,1788,1814]" box="[189,300,1788,1814]" pageId="12" pageNumber="13">
<heading id="72024C22FFFAA54AFF59F91EFEF8F8CA" bold="true" box="[189,300,1788,1814]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="1B81275CFFFAA54AFF59F91EFEF8F8CA" bold="true" box="[189,300,1788,1814]" pageId="12" pageNumber="13">Intensity</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59F8C9FDB8F899" blockId="12.[189,620,1834,1861]" box="[189,620,1834,1861]" pageId="12" pageNumber="13">
About
<specimenCount id="3FF330C7FFFAA54AFEEAF8C8FE78F899" box="[270,428,1834,1861]" pageId="12" pageNumber="13" type="generic">25 specimens</specimenCount>
in a single host.
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFAA54BFF59F88EFC9AF856" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="294AFB4EFFFAA54AFF59F88EFE9AF85A" blockId="12.[189,334,1900,1926]" box="[189,334,1900,1926]" pageId="12" pageNumber="13">
<heading id="72024C22FFFAA54AFF59F88EFE9AF85A" bold="true" box="[189,334,1900,1926]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="1B81275CFFFAA54AFF59F88EFE9AF85A" bold="true" box="[189,334,1900,1926]" pageId="12" pageNumber="13">Description</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFAA54BFF59F879FE65FCCE" blockId="12.[189,1399,1946,2009]" lastBlockId="13.[189,1399,267,786]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Body very small, up to 6.5–8.1 (7.3, n = 7) mm long, consisting of 39–53 proglottides; maximum width 600–725 (694, n = 9) at level of gravid proglottides. Proglottides craspedote, mostly wider than long, but shape depending on contraction. Ventral osmoregulatory canals with posterior transverse anastomoses in each proglottis. Strong longitudinal muscular bundles. Scolex
<quantity id="EE0D56ABFFFBA54BFAE9FECCFF01FEB0" metricMagnitude="0" metricUnit="m" metricValue="7.1501" metricValueMax="7.747" metricValueMin="6.5532" pageId="13" pageNumber="14" unit="in" value="281.5" valueMax="305.0" valueMin="258.0">258–305 in</quantity>
diameter (280, n = 6), slightly wider than neck, widest at level of suckers (
<figureCitation id="B1CEE7CBFFFBA54BFB8AFEB0FB1DFEB0" box="[1134,1225,338,364]" captionStart="Figs 12–16" captionStartId="14.[189,241,1943,1969]" captionTargetBox="[218,1382,269,1894]" captionTargetId="figure@14.[203,1384,265,1915]" captionTargetPageId="14" captionText="Figs 12–16. Monopylidium australiense sp. nov. 12. Scolex. 13. Hooks. 14. Mature proglottis. 15. Terminal genitalia. 16. Eggs. Scale bars: 12, 14 = 100 µm; 13, 16 = 20 µm; 15 = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3827560" httpUri="https://zenodo.org/record/3827560/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
). Neck poorly or not marked. Suckers elongated, very muscular, with irregular margins,
<quantity id="EE0D56ABFFFBA54BFBD3FE97FB14FE53" box="[1079,1216,373,400]" metricMagnitude="0" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.3019999999999996" pageId="13" pageNumber="14" unit="in" value="140.0" valueMax="150.0" valueMin="130.0">130–150 in</quantity>
diameter (132, n = 27). Rostellar pouch strong, musculo-glandular, usually with distinctive widening anterior to suckers, extending beyond level of posterior margin of suckers and becoming much narrower; often folded posteriorly, 270 × 112 (224–320 × 105–130, n = 6).
<taxonomicName id="EEF580CDFFFBA54BFC70FE3CFBC5FE24" authorityName="Montfort" authorityYear="1810" box="[916,1041,478,504]" family="Rostellum" pageId="13" pageNumber="18" rank="family">Rostellum</taxonomicName>
muscular, mushroom-shaped 135–150 × 55–83 (144 × 73, n = 3), with distinct apical pad. Rostellar hooks
<quantity id="EE0D56ABFFFBA54BFB9AFDE0FB3FFDC0" box="[1150,1259,514,540]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.334" metricValueMin="5.08" pageId="13" pageNumber="14" unit="in" value="20.5" valueMax="21.0" valueMin="20.0">20–21 in</quantity>
number, in two regular rows, sub-equal; anterior hooks 23–26 (24.2, n = 24) long, posterior hooks 22.5–25.0 (23.7, n = 26) long; all hooks of similar shape, with straight handle, strong short guard and very short blade; small epiphyseal-like outgrowth at tip of handle (
<figureCitation id="B1CEE7CBFFFBA54BFC85FD89FC6EFD59" box="[865,954,619,645]" captionStart="Figs 12–16" captionStartId="14.[189,241,1943,1969]" captionTargetBox="[218,1382,269,1894]" captionTargetId="figure@14.[203,1384,265,1915]" captionTargetPageId="14" captionText="Figs 12–16. Monopylidium australiense sp. nov. 12. Scolex. 13. Hooks. 14. Mature proglottis. 15. Terminal genitalia. 16. Eggs. Scale bars: 12, 14 = 100 µm; 13, 16 = 20 µm; 15 = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3827560" httpUri="https://zenodo.org/record/3827560/files/figure.png" pageId="13" pageNumber="14">Fig. 13</figureCitation>
). Neck short, proglottization distinct at 25–65 (46, n = 6) from posterior margin of suckers. Genital pores at border of first third of lateral proglottis margin, irregularly alternating in short series, rarely more than 2 (maximum observed 4) consecutive pores on a single side. Genital ducts passing between osmoregulatory canals. Genital atrium infundibular.
</paragraph>
<paragraph id="294AFB4EFFFBA54BFF59FCD8FB2DFBDF" blockId="13.[189,1399,825,1028]" pageId="13" pageNumber="14">
Testes large,
<quantity id="EE0D56ABFFFBA54BFEBEFCDBFE10FC8F" box="[346,452,825,851]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="7.366" metricValueMin="4.064" pageId="13" pageNumber="14" unit="in" value="22.5" valueMax="29.0" valueMin="16.0">16–29 in</quantity>
number (24, n = 52), in median field of proglottis, in double, sometime triple, compact posterior layer, often overlapping vitellarium and ovary. External vas deferens extremely developed, massive and highly convoluted, antero-central, often covering cirrus-sac. Cirrus-sac cylindrical, sometimes slightly wider distally, 86–138 × 26–38 (113 × 33, n = 43), crossing excretory canals and half of it might be situated in median field (
<figureCitation id="B1CEE7CBFFFBA54BFCAEFC24FC77FC3C" box="[842,931,966,992]" captionStart="Figs 12–16" captionStartId="14.[189,241,1943,1969]" captionTargetBox="[218,1382,269,1894]" captionTargetId="figure@14.[203,1384,265,1915]" captionTargetPageId="14" captionText="Figs 12–16. Monopylidium australiense sp. nov. 12. Scolex. 13. Hooks. 14. Mature proglottis. 15. Terminal genitalia. 16. Eggs. Scale bars: 12, 14 = 100 µm; 13, 16 = 20 µm; 15 = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3827560" httpUri="https://zenodo.org/record/3827560/files/figure.png" pageId="13" pageNumber="14">Fig. 14</figureCitation>
). Cirrus
<quantity id="EE0D56ABFFFBA54BFBEEFC24FBA7FC3C" box="[1034,1139,966,992]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="13" pageNumber="14" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
diameter, armed with well developed spines situated on part of it with length 50. Internal vas deferens convoluted.
</paragraph>
<paragraph id="294AFB4EFFFBA54BFF59FBC9FB4BFB2A" blockId="13.[189,1399,1067,1270]" pageId="13" pageNumber="14">
Vitellarium central, oval, mostly regular, composed of few compact lobes. Ovary central, massive, biwinged, multilobate, filling entire width of median field when fully developed, sometimes even slightly crossing lateral osmoregulatory canals. Mehlis’ gland central, overlapping seminal receptacle but not always distinct. Seminal receptacle elongated, anterior to vitellarium and between wings of ovary. Vagina thin-walled with discrete basal muscular sphincter; opening posteriorly to cirrus-sac, sometimes partly overlapping it; lined with thin spines in its distal copulatory part (
<figureCitation id="B1CEE7CBFFFBA54BFBEDFB39FB46FB2A" box="[1033,1170,1243,1270]" captionStart="Figs 12–16" captionStartId="14.[189,241,1943,1969]" captionTargetBox="[218,1382,269,1894]" captionTargetId="figure@14.[203,1384,265,1915]" captionTargetPageId="14" captionText="Figs 12–16. Monopylidium australiense sp. nov. 12. Scolex. 13. Hooks. 14. Mature proglottis. 15. Terminal genitalia. 16. Eggs. Scale bars: 12, 14 = 100 µm; 13, 16 = 20 µm; 15 = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3827560" httpUri="https://zenodo.org/record/3827560/files/figure.png" pageId="13" pageNumber="14">Figs 14–15</figureCitation>
).
</paragraph>
<paragraph id="294AFB4EFFFBA54BFF59FAFFFE24FA1F" blockId="13.[189,1399,1308,1475]" pageId="13" pageNumber="14">
Uterus initially broadly reticulate, developing quickly over whole median field width and extending in lateral field. With further development, uterus becoming compact, saccular, with few deep lobes. Oncospheres slightly oval, 19–24 (21, n = 20) in diameter. Eggs with thin outer shell provided with two big and distinct nuclei; embryophore spherical, thick (
<figureCitation id="B1CEE7CBFFFBA54BFCD5FA64FC5CFA7D" box="[817,904,1414,1441]" captionStart="Figs 12–16" captionStartId="14.[189,241,1943,1969]" captionTargetBox="[218,1382,269,1894]" captionTargetId="figure@14.[203,1384,265,1915]" captionTargetPageId="14" captionText="Figs 12–16. Monopylidium australiense sp. nov. 12. Scolex. 13. Hooks. 14. Mature proglottis. 15. Terminal genitalia. 16. Eggs. Scale bars: 12, 14 = 100 µm; 13, 16 = 20 µm; 15 = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3827560" httpUri="https://zenodo.org/record/3827560/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
). Central embryonic hooks 11 long, lateral embryonic hooks 10 long.
</paragraph>
<paragraph id="294AFB4EFFFBA54BFF59FA08FBDFF9DA" blockId="13.[189,1035,1514,1542]" box="[189,1035,1514,1542]" pageId="13" pageNumber="14">
<emphasis id="1B81275CFFFBA54BFF59FA08FD55F9D8" bold="true" box="[189,641,1514,1541]" pageId="13" pageNumber="14">Observations on additional material</emphasis>
(not belonging to the
<typeStatus id="F64E45ECFFFBA54BFC61FA09FC6CF9D9" box="[901,952,1515,1541]" pageId="13" pageNumber="14">type</typeStatus>
series)
</paragraph>
<paragraph id="294AFB4EFFFBA54BFF59F9FBFDC5F91D" blockId="13.[189,1399,1561,1729]" pageId="13" pageNumber="14">
AHC 22931: about 40, extremely small specimens, shorter than
<quantity id="EE0D56ABFFFBA54BFC48F9FBFC26F9E8" box="[940,1010,1561,1588]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="2.0">2 mm</quantity>
, maximum width 510, composed of fewer than 20 proglottides but with extremely rapid development. Some early gravid proglottides present. General characters in accordance with the above description to the exception of slightly wider rostella [85–105 (95, n = 10)] and slight variation in number of rostellar hooks (single specimens with 19, 22 or 23 rostellar hooks).
</paragraph>
<paragraph id="294AFB4EFFFBA54BFF59F90AFE93F8F9" blockId="13.[189,1398,1767,1829]" pageId="13" pageNumber="14">AHC 29271: two slides with fragments of generally poor quality. The observable characters fit the above description.</paragraph>
<paragraph id="294AFB4EFFFBA54BFF59F8AEFC9AF856" blockId="13.[189,1399,1868,1930]" pageId="13" pageNumber="14">
AHC 29281: a single slide composed of fragments of smaller worms, similar to material from host 22931. Slightly smaller scoleces (
<quantity id="EE0D56ABFFFBA54BFDACF88DFD1BF855" box="[584,719,1903,1930]" metricMagnitude="0" metricUnit="m" metricValue="5.3975" metricValueMax="5.715" metricValueMin="5.08" pageId="13" pageNumber="14" unit="in" value="212.5" valueMax="225.0" valueMin="200.0">200–225 in</quantity>
diameter).
</paragraph>
</subSubSection>
<caption id="7D8AABC6FFF8A548FF59F875FB25F809" ID-DOI="http://doi.org/10.5281/zenodo.3827560" ID-Zenodo-Dep="3827560" httpUri="https://zenodo.org/record/3827560/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,241,1943,1969]" targetBox="[218,1382,269,1894]" targetPageId="14">
<paragraph id="294AFB4EFFF8A548FF59F875FB25F809" blockId="14.[189,1399,1943,2005]" pageId="14" pageNumber="15">
<emphasis id="1B81275CFFF8A548FF59F875FE85F86D" bold="true" box="[189,337,1943,1969]" pageId="14" pageNumber="15">Figs 12–16.</emphasis>
<taxonomicName id="EEF580CDFFF8A548FE86F875FD78F86D" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[354,684,1943,1969]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="14" pageNumber="15" phylum="Platyhelminthes" rank="species" species="australiense" status="sp. nov.">
<emphasis id="1B81275CFFF8A548FE86F875FD78F86D" box="[354,684,1943,1969]" italics="true" pageId="14" pageNumber="15">Monopylidium australiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFF8A548FD59F875FCF4F86D" box="[701,800,1943,1969]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1B81275CFFF8A548FCD4F875FC99F86D" bold="true" box="[816,845,1943,1969]" pageId="14" pageNumber="15">12</emphasis>
. Scolex.
<emphasis id="1B81275CFFF8A548FC28F875FC3DF86D" bold="true" box="[972,1001,1943,1969]" pageId="14" pageNumber="15">13</emphasis>
. Hooks.
<emphasis id="1B81275CFFF8A548FB81F875FB56F86D" bold="true" box="[1125,1154,1943,1969]" pageId="14" pageNumber="15">14</emphasis>
. Mature proglottis.
<emphasis id="1B81275CFFF8A548FF59F858FF0EF808" bold="true" box="[189,218,1978,2004]" pageId="14" pageNumber="15">15</emphasis>
. Terminal genitalia.
<emphasis id="1B81275CFFF8A548FE29F858FE3EF808" bold="true" box="[461,490,1978,2004]" pageId="14" pageNumber="15">16</emphasis>
. Eggs. Scale bars: 12, 14 = 100 µm; 13, 16 = 20 µm; 15 = 50 µm.
</paragraph>
</caption>
<subSubSection id="61EFA8C5FFF9A549FF59FEE9FF22FAAC" pageId="15" pageNumber="16" type="discussion">
<paragraph id="294AFB4EFFF9A549FF59FEE9FEFBFEF9" blockId="15.[189,303,267,293]" box="[189,303,267,293]" pageId="15" pageNumber="16">
<heading id="72024C22FFF9A549FF59FEE9FEFBFEF9" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="1B81275CFFF9A549FF59FEE9FEFBFEF9" bold="true" box="[189,303,267,293]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFF9A549FF59FED8FBA9FE3C" blockId="15.[189,1399,313,481]" pageId="15" pageNumber="16">
In its report on helminth parasite of lyrebirds in
<collectingCountry id="51E2BBDEFFF9A549FCADFEDBFC6CFE8F" box="[841,952,313,339]" name="Australia" pageId="15" pageNumber="16">Australia</collectingCountry>
,
<bibRefCitation id="4D6486BFFFF9A549FC29FEDBFB41FE88" author="Johnston T. H." box="[973,1173,313,340]" pageId="15" pageNumber="16" pagination="63 - 91" refId="ref19818" refString="Johnston T. H. 1913. Notes on some Entozoa. Proceedings of the Royal Society of Queensland 24: 63 - 91." type="journal article" year="1913">Johnston (1913)</bibRefCitation>
described a new acanthocephalan but no cestode. Much later,
<bibRefCitation id="4D6486BFFFF9A549FD12FEBFFBF7FEAB" author="McOrist S. &amp; Scott P. C." box="[758,1059,349,375]" pageId="15" pageNumber="16" pagination="420 - 421" refId="ref20610" refString="McOrist S. &amp; Scott P. C. 1989. Parasitic enteritis in superb lyrebirds (Menura novaehollandiae). Journal of Wildlife Diseases 25: 420 - 421. https: // doi. org / 10.7589 / 0090 - 3558 - 25.3.420" type="journal article" year="1989">McOrist &amp; Scott (1989)</bibRefCitation>
examined superb lyrebirds suffering from enteritis that was caused by acanthocephalan infections in Southern
<collectingRegion id="EB3135ACFFF9A549FB7FFE62FB2DFE46" box="[1179,1273,384,410]" country="Australia" name="Victoria" pageId="15" pageNumber="16">Victoria</collectingRegion>
and noted the presence of “occasional cestodes of the genus
<taxonomicName id="EEF580CDFFF9A549FCEDFE41FC7BFE61" box="[777,943,419,445]" class="Cestoda" family="Dilepididae" genus="Choanotaenia" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFF9A549FCEDFE41FC7BFE61" box="[777,943,419,445]" italics="true" pageId="15" pageNumber="16">Choanotaenia</emphasis>
</taxonomicName>
” in the specimens they studied. To the best of our knowledge, these are the only helminthological reports for these birds.
</paragraph>
<paragraph id="294AFB4EFFF9A549FF59FDE8FC08FCF6" blockId="15.[189,1399,522,810]" pageId="15" pageNumber="16">
To our knowledge, there are only two reports of a
<taxonomicName id="EEF580CDFFF9A549FD12FDE8FC75FDF8" authorityName="Fuhrmann" authorityYear="1899" box="[758,929,522,548]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFF9A549FD12FDE8FC75FDF8" box="[758,929,522,548]" italics="true" pageId="15" pageNumber="16">Monopylidium</emphasis>
</taxonomicName>
in
<collectingCountry id="51E2BBDEFFF9A549FC25FDE8FBE4FDF8" box="[961,1072,522,548]" name="Australia" pageId="15" pageNumber="16">Australia</collectingCountry>
.
<bibRefCitation id="4D6486BFFFF9A549FBDDFDE8FB21FDF8" author="Johnston T. H." box="[1081,1269,522,548]" pageId="15" pageNumber="16" pagination="405 - 411" refId="ref19786" refString="Johnston T. H. 1909. On the anatomy of Monopylidium passerinum Fuhrmann. Journal and Proceedings of the Royal Society of New South Wales 43: 405 - 411." type="journal article" year="1909">Johnston (1909)</bibRefCitation>
mentioned the cosmopolitan species
<taxonomicName id="EEF580CDFFF9A549FE01FDD2FCB8FD95" authority="(Fuhrmann, 1907)" baseAuthorityName="Fuhrmann" baseAuthorityYear="1907" box="[485,876,559,585]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="species" species="passerinum">
<emphasis id="1B81275CFFF9A549FE01FDD2FD46FD95" box="[485,658,560,585]" italics="true" pageId="15" pageNumber="16">M. passerinum</emphasis>
(Fuhrmann, 1907)
</taxonomicName>
in
<taxonomicName id="EEF580CDFFF9A549FC6BFDD2FBB2FD95" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[911,1126,559,585]" class="Aves" family="Passeridae" genus="Passer" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="domesticus">
<emphasis id="1B81275CFFF9A549FC6BFDD2FBB2FD95" box="[911,1126,559,585]" italics="true" pageId="15" pageNumber="16">Passer domesticus</emphasis>
</taxonomicName>
L. in
<collectingRegion id="EB3135ACFFF9A549FB4CFDD2FAA3FD95" box="[1192,1399,559,586]" country="Australia" name="New South Wales" pageId="15" pageNumber="16">New South Wales</collectingRegion>
and gave a detailed description. This species differed from the present material by several characters including significantly shorter rostellar hooks (
<tableCitation id="6477CEF5FFF9A549FD07FD98FCEAFD48" box="[739,830,634,660]" captionStart="Table 3" captionStartId="17.[188,257,542,570]" captionTargetBox="[199,1940,691,1741]" captionText="Table 3 (continued on next page). Comparison of Monopylidium australiense sp. nov. with the other species of the genera Monopylidium Fuhrmann, 1899 and Icterotaenia Railliet &amp; Henry, 1909. See text for the accepted generic concept. Note: * 20, 10 and 19 respectively in original reference, a printing error, as confirmed by the length of cirrus-sac on the original illustration." httpUri="http://table.plazi.org/id/7D8AABC6FFE7A557FF58FDFCFBACFD5C" pageId="15" pageNumber="16" tableUuid="7D8AABC6FFE7A557FF58FDFCFBACFD5C">Table 3</tableCitation>
).
<taxonomicName id="EEF580CDFFF9A549FCB1FD98FBBCFD48" baseAuthorityName="Maplestone &amp; Southwell" baseAuthorityYear="1923" box="[853,1128,634,660]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="species" species="fieldingi">
<emphasis id="1B81275CFFF9A549FCB1FD98FBBCFD48" box="[853,1128,634,660]" italics="true" pageId="15" pageNumber="16">Monopylidium fieldingi</emphasis>
</taxonomicName>
has also been reported from
<collectingCountry id="51E2BBDEFFF9A549FF1BFD7DFEBFFD65" box="[255,363,671,697]" name="Australia" pageId="15" pageNumber="16">Australia</collectingCountry>
in the butcherbird
<taxonomicName id="EEF580CDFFF9A549FDB1FD42FBC9FD65" authority="Temminck, 1824" authorityName="Temminck" authorityYear="1824" box="[597,1053,671,698]" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="destructor">
<emphasis id="1B81275CFFF9A549FDB1FD42FC9EFD65" box="[597,842,671,698]" italics="true" pageId="15" pageNumber="16">Cracticus destructor</emphasis>
Temminck, 1824
</taxonomicName>
[now
<taxonomicName id="EEF580CDFFF9A549FB89FD42FED0FD03" authority="(Latham, 1801)" baseAuthorityName="Latham" baseAuthorityYear="1801" class="Aves" family="Cracticidae" genus="Cracticus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="1B81275CFFF9A549FB89FD42FAD5FD65" box="[1133,1281,672,698]" italics="true" pageId="15" pageNumber="16">C. torquatus</emphasis>
(Latham, 1801)
</taxonomicName>
] in North
<collectingRegion id="EB3135ACFFF9A549FE9CFD26FDD6FD02" box="[376,514,708,734]" country="Australia" name="Queensland" pageId="15" pageNumber="16">Queensland</collectingRegion>
(
<bibRefCitation id="4D6486BFFFF9A549FDF5FD26FCAEFD03" author="Maplestone P. &amp; Southwell T." box="[529,890,708,735]" pageId="15" pageNumber="16" pagination="317 - 331" refId="ref20383" refString="Maplestone P. &amp; Southwell T. 1923. Notes on Australian Cestodes. VII. Annals of Tropical Medicine and Parasitology 17: 317 - 331. https: // doi. org / 10.1080 / 00034983.1923.11684361" type="journal article" year="1923">Maplestone &amp; Southwell 1923</bibRefCitation>
). However, this taxon has a clearly different anatomy, in particular the presence of 80 rose-thorn shaped hooks in two rows, and is now placed in the genus
<taxonomicName id="EEF580CDFFF9A549FEEDFCF2FC91FCF6" authority="Spasskii, 1966" authorityName="Spasskii" authorityYear="1966" box="[265,837,783,810]" class="Cestoda" family="Metadilepididae" genus="Cracticotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFF9A549FEEDFCF2FE6DFCF6" box="[265,441,784,810]" italics="true" pageId="15" pageNumber="16">Cracticotaenia</emphasis>
<bibRefCitation id="4D6486BFFFF9A549FE24FCEDFDB8FCF5" author="Spasskii A. A." box="[448,620,783,809]" pageId="15" pageNumber="16" pagination="1483 - 1485" refId="ref21581" refString="Spasskii A. A. 1966. On the heterogeneity of the genus Anomotaenia (Cestoda: Dilepididae). Doklady Akademii Nauk USSR 69: 1483 - 1485. [In Russian.]" type="journal article" year="1966">Spasskii, 1966</bibRefCitation>
(Metadilepididae)
</taxonomicName>
(see below).
</paragraph>
<paragraph id="294AFB4EFFF9A549FF59FCB1FE62FAD4" blockId="15.[189,1399,851,1288]" pageId="15" pageNumber="16">
Here, we follow the concept of
<bibRefCitation id="4D6486BFFFF9A549FDB7FCB1FCC7FCB1" author="Spasskii A. A." box="[595,787,851,877]" pageId="15" pageNumber="16" pagination="42 - 49" refId="ref21616" refString="Spasskii A. A. 1993. On the structure of species of genus Monopylidium (Dilepididae) and its position in Cestoda system. Izvestiya Akademii Nauk Respubliki Moldova Biologicheskie i Khimicheskie Nauki 1: 42 - 49. [In Russian.]" type="journal article" year="1993">Spasskii (1993)</bibRefCitation>
as updated by
<bibRefCitation id="4D6486BFFFF9A549FC3CFCB1FBA0FCB1" author="Bona F. V." box="[984,1140,851,877]" pageId="15" pageNumber="16" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">
<collectingCountry id="51E2BBDEFFF9A549FC3CFCB1FBC3FCB1" box="[984,1047,851,877]" name="Algeria" pageId="15" pageNumber="16">Bona</collectingCountry>
(1994)
</bibRefCitation>
for the definition of
<taxonomicName id="EEF580CDFFF9A549FF59FC9AFEB8FC4E" authorityName="Fuhrmann" authorityYear="1899" box="[189,364,888,914]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFF9A549FF59FC9AFEB8FC4E" box="[189,364,888,914]" italics="true" pageId="15" pageNumber="16">Monopylidium</emphasis>
</taxonomicName>
. Thus,
<taxonomicName id="EEF580CDFFF9A549FE21FC9BFCA5FC4E" authority="Railliet &amp; Henry, 1909" authorityName="Railliet &amp; Henry" authorityYear="1909" box="[453,881,888,915]" class="Cestoda" family="Dilepididae" genus="Icterotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFF9A549FE21FC9BFD83FC4E" box="[453,599,889,914]" italics="true" pageId="15" pageNumber="16">Icterotaenia</emphasis>
Railliet &amp; Henry, 1909
</taxonomicName>
is considered a synonym and some species validated by
<bibRefCitation id="4D6486BFFFF9A549FE84FC7FFDF4FC64" author="Spasskii A. A." box="[352,544,925,952]" pageId="15" pageNumber="16" pagination="42 - 49" refId="ref21616" refString="Spasskii A. A. 1993. On the structure of species of genus Monopylidium (Dilepididae) and its position in Cestoda system. Izvestiya Akademii Nauk Respubliki Moldova Biologicheskie i Khimicheskie Nauki 1: 42 - 49. [In Russian.]" type="journal article" year="1993">Spasskii (1993)</bibRefCitation>
are now placed in different genera (like
<taxonomicName id="EEF580CDFFF9A549FBC2FC7FFA87FC64" authority="Bona, 1994" authorityName="Bona" authorityYear="1994" box="[1062,1363,925,952]" class="Cestoda" family="Dilepididae" genus="Cuculincola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFF9A549FBC2FC7FFB6DFC6B" box="[1062,1209,925,951]" italics="true" pageId="15" pageNumber="16">Cuculincola</emphasis>
<bibRefCitation id="4D6486BFFFF9A549FB21FC7CFA87FC64" author="Bona F. V." box="[1221,1363,926,952]" pageId="15" pageNumber="16" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">Bona, 1994</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="EEF580CDFFF9A549FF59FC21FDD4FC01" authority="Bona, 1994" authorityName="Bona" authorityYear="1994" box="[189,512,963,989]" class="Cestoda" family="Dilepididae" genus="Spasspasskya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFF9A549FF59FC21FEB4FC01" box="[189,352,963,989]" italics="true" pageId="15" pageNumber="16">Spasspasskya</emphasis>
<bibRefCitation id="4D6486BFFFF9A549FE88FC21FDD4FC01" author="Bona F. V." box="[364,512,963,989]" pageId="15" pageNumber="16" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">Bona, 1994</bibRefCitation>
</taxonomicName>
). Globally, about 25 species of
<emphasis id="1B81275CFFF9A549FC71FC21FBEFFC01" box="[917,1083,963,989]" italics="true" pageId="15" pageNumber="16">Monopylidum</emphasis>
are presently recognized (
<tableCitation id="6477CEF5FFF9A549FF21FC0AFEF3FBDE" box="[197,295,1000,1026]" captionStart="Table 3" captionStartId="17.[188,257,542,570]" captionTargetBox="[199,1940,691,1741]" captionText="Table 3 (continued on next page). Comparison of Monopylidium australiense sp. nov. with the other species of the genera Monopylidium Fuhrmann, 1899 and Icterotaenia Railliet &amp; Henry, 1909. See text for the accepted generic concept. Note: * 20, 10 and 19 respectively in original reference, a printing error, as confirmed by the length of cirrus-sac on the original illustration." httpUri="http://table.plazi.org/id/7D8AABC6FFE7A557FF58FDFCFBACFD5C" pageId="15" pageNumber="16" tableUuid="7D8AABC6FFE7A557FF58FDFCFBACFD5C">Table 3</tableCitation>
), although the status of some of them remains equivocal. Our material can easily be differentiated from nearly all of them, although
<taxonomicName id="EEF580CDFFF9A549FCF9FBECFB78FBF4" authority="(Dujardin, 1845)" baseAuthorityName="Dujardin" baseAuthorityYear="1845" box="[797,1196,1037,1064]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="species" species="attenuatum">
<emphasis id="1B81275CFFF9A549FCF9FBECFC05FBFB" box="[797,977,1038,1063]" italics="true" pageId="15" pageNumber="16">M. attenuatum</emphasis>
(Dujardin, 1845)
</taxonomicName>
and
<taxonomicName id="EEF580CDFFF9A549FB0BFBECFEBEFB92" authority="(Singh, 1964)" baseAuthorityName="Singh" baseAuthorityYear="1964" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="species" species="tandani">
<emphasis id="1B81275CFFF9A549FB0BFBECFAA3FBFB" box="[1263,1399,1037,1063]" italics="true" pageId="15" pageNumber="16">M. tandani</emphasis>
(
<bibRefCitation id="4D6486BFFFF9A549FF22FBD1FEB6FB92" author="Singh K. S." box="[198,354,1075,1102]" pageId="15" pageNumber="16" pagination="177 - 194" refId="ref21316" refString="Singh K. S. 1964. On six new avian cestodes form India. Parasitology 54: 177 - 194. https: // doi. org / 10.1017 / s 0031182000074461" type="journal article" year="1964">Singh, 1964</bibRefCitation>
)
</taxonomicName>
present relatively similar characteristics. However, the former shows a smaller
<taxonomicName id="EEF580CDFFF9A549FF59FBBAFEFAFBAE" authorityName="Montfort" authorityYear="1810" box="[189,302,1112,1138]" family="Rostellum" pageId="15" pageNumber="18" rank="family">rostellum</taxonomicName>
bearing hooks of different shape, larger oncospheres and embryonic hooks than those of our specimens. It may also have unarmed cirrus (
<bibRefCitation id="4D6486BFFFF9A549FD3FFB9FFBF9FB44" author="Spasskaya L. P. &amp; Spasskii A." box="[731,1069,1149,1176]" pageId="15" pageNumber="16" refId="ref21515" refString="Spasskaya L. P. &amp; Spasskii A. 1977. Cestodes of Birds in the USSR. Dilepididae of Terrestrial Birds. Izdate'stvo Nauka, Moscow. [In Russian.]" type="book" year="1977">Spasskaya &amp; Spasskii 1977</bibRefCitation>
). The latter is a short worm with few (&lt;20) proglottides, hooks with longer blades and shorter handles and an unarmed cirrus. It is only known from
<taxonomicName id="EEF580CDFFF9A549FE50FB2AFC24FB3E" authority="Vigors, 1831" authorityName="Vigors" authorityYear="1831" box="[436,1008,1224,1250]" class="Aves" family="Muscicapidae" genus="Myophonus" kingdom="Animalia" order="Passeriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="caeruleus" subSpecies="temminckii">
<emphasis id="1B81275CFFF9A549FE50FB2AFC9DFB3E" box="[436,841,1224,1250]" italics="true" pageId="15" pageNumber="16">Myophonus caeruleus temminckii</emphasis>
Vigors, 1831
</taxonomicName>
, a trush found in the Himalayan region (
<bibRefCitation id="4D6486BFFFF9A549FEFEFB0FFE7EFAD4" author="Singh K. S." box="[282,426,1261,1288]" pageId="15" pageNumber="16" pagination="177 - 194" refId="ref21316" refString="Singh K. S. 1964. On six new avian cestodes form India. Parasitology 54: 177 - 194. https: // doi. org / 10.1017 / s 0031182000074461" type="journal article" year="1964">Singh 1964</bibRefCitation>
).
</paragraph>
<paragraph id="294AFB4EFFF9A549FF59FAD3FF22FAAC" blockId="15.[189,1398,1329,1392]" pageId="15" pageNumber="16">
Consequently,
<taxonomicName id="EEF580CDFFF9A549FE8EFAD3FD7EFA97" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[362,682,1329,1355]" class="Cestoda" family="Dilepididae" genus="Monopylidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="15" pageNumber="16" phylum="Platyhelminthes" rank="species" species="australiense" status="sp. nov.">
<emphasis id="1B81275CFFF9A549FE8EFAD3FD7EFA97" box="[362,682,1329,1355]" italics="true" pageId="15" pageNumber="16">Monopylidium australiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFF9A549FD55FAD0FCDEFA90" box="[689,778,1330,1356]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
is the single known cestode parasitizing this endemic host.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DAF5F808791D68237D651ADAC4CCBE2F">Seven new species of cestode parasites (Neodermata, Platyhelminthes) from Australian birds</mods:title>
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<paragraph id="294AFB4EFFFCA54CFD32FA65FC89FA7E" blockId="10.[726,861,1415,1442]" box="[726,861,1415,1442]" pageId="10" pageNumber="11">
<heading id="72024C22FFFCA54CFD32FA65FC89FA7E" box="[726,861,1415,1442]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="EEF580CDFFFCA54CFD32FA65FCE1FA7E" authorityName="Singh" authorityYear="1952" box="[726,821,1415,1442]" class="Cestoda" family="Dilepididae" genus="Neoliga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFFCA54CFD32FA65FCE1FA7E" bold="true" box="[726,821,1415,1442]" italics="true" pageId="10" pageNumber="11">Neoliga</emphasis>
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sp.
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</subSubSection>
<subSubSection id="61EFA8C5FFFCA54CFF59FA2BFD25F9A7" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="294AFB4EFFFCA54CFF59FA2BFE7EFA3F" blockId="10.[189,426,1481,1507]" box="[189,426,1481,1507]" pageId="10" pageNumber="11">
<heading id="72024C22FFFCA54CFF59FA2BFE7EFA3F" bold="true" box="[189,426,1481,1507]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="1B81275CFFFCA54CFF59FA2BFE7EFA3F" bold="true" box="[189,426,1481,1507]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFCA54CFF59FA1AFD25F9A7" blockId="10.[189,1399,1526,1660]" pageId="10" pageNumber="11">
<collectingCountry id="51E2BBDEFFFCA54CFF59FA1AFEBCF9CE" box="[189,360,1528,1554]" name="Australia" pageId="10" pageNumber="11">AUSTRALIA</collectingCountry>
:
<collectingRegion id="EB3135ACFFFCA54CFE91FA1AFD9BF9CD" box="[373,591,1527,1554]" country="Australia" name="New South Wales" pageId="10" pageNumber="11">New South Wales</collectingRegion>
, Sydney, 33.48′ S,
<geoCoordinate id="4CC19D89FFFCA54CFCDDFA14FC70F9CE" box="[825,932,1526,1554]" degrees="151" direction="east" minutes="6" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="151.1">151°6′ E</geoCoordinate>
, 2005, Karie Rose leg.,
<specimenCount id="3FF330C7FFFCA54CFB2FFA1AFAF4F9CE" box="[1227,1312,1528,1554]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
in two parts, rather good condition but no gravid proglottides, on same slide as
<typeStatus id="F64E45ECFFFCA54CFBE0F9F9FBE3F9E9" box="[1028,1079,1563,1589]" pageId="10" pageNumber="11">type</typeStatus>
material of
<taxonomicName id="EEF580CDFFFCA54CFB27F9F9FEF0F984" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="musealis" status="sp. nov.">
<emphasis id="1B81275CFFFCA54CFB27F9F9FEF0F984" italics="true" pageId="10" pageNumber="11">Notopentorchis musealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFFCA54CFECFF9DCFE51F984" box="[299,389,1598,1624]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
(AHC 36479); multiple small to minute fragments in bad condition, co-infection with
<taxonomicName id="EEF580CDFFFCA54CFF59F983FE0BF9A7" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[189,479,1633,1659]" class="Cestoda" family="Paruterinidae" genus="Notopentorchis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="10" pageNumber="11" phylum="Platyhelminthes" rank="species" species="musealis" status="sp. nov.">
<emphasis id="1B81275CFFFCA54CFF59F983FE0BF9A7" box="[189,479,1633,1659]" italics="true" pageId="10" pageNumber="11">Notopentorchis musealis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFFCA54CFE02F980FD95F9A0" box="[486,577,1634,1660]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
(AHC 33108).
</paragraph>
</subSubSection>
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<heading id="72024C22FFFCA54CFF59F941FF2CF961" bold="true" box="[189,248,1699,1725]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="1B81275CFFFCA54CFF59F941FF2CF961" bold="true" box="[189,248,1699,1725]" pageId="10" pageNumber="11">Host</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFCA54CFF59F933FC05F930" blockId="10.[189,977,1745,1772]" box="[189,977,1745,1772]" pageId="10" pageNumber="11">
<taxonomicName id="EEF580CDFFFCA54CFF59F933FD40F937" authority="(Latham, 1801)" baseAuthorityName="Latham" baseAuthorityYear="1801" box="[189,660,1745,1771]" class="Aves" family="Apodidae" genus="Hirundapus" kingdom="Animalia" order="Apodiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="caudacutus">
<emphasis id="1B81275CFFFCA54CFF59F933FE02F937" box="[189,470,1745,1771]" italics="true" pageId="10" pageNumber="11">Hirundapus caudacutus</emphasis>
(Latham, 1801)
</taxonomicName>
(
<taxonomicName id="EEF580CDFFFCA54CFD47F933FC97F937" box="[675,835,1745,1771]" class="Aves" kingdom="Animalia" order="Apodiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="order">Apodiformes</taxonomicName>
,
<taxonomicName id="EEF580CDFFFCA54CFCA9F933FC11F930" box="[845,965,1745,1772]" class="Aves" family="Apodidae" kingdom="Animalia" order="Apodiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Apodidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="61EFA8C5FFFCA54AFF59F8F0FBEEFE6E" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="294AFB4EFFFCA54CFF59F8F0FE9AF8F0" blockId="10.[189,334,1810,1836]" box="[189,334,1810,1836]" pageId="10" pageNumber="11">
<heading id="72024C22FFFCA54CFF59F8F0FE9AF8F0" bold="true" box="[189,334,1810,1836]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="1B81275CFFFCA54CFF59F8F0FE9AF8F0" bold="true" box="[189,334,1810,1836]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
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The available material is not sufficient to allow for a full description. Scolex
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diameter. Suckers weak, oval,
<quantity id="EE0D56ABFFFCA54CFEA9F886FE07F8A2" box="[333,467,1892,1919]" metricMagnitude="0" metricUnit="m" metricValue="5.3975" metricValueMax="5.587999999999999" metricValueMin="5.207" pageId="10" pageNumber="11" unit="in" value="212.5" valueMax="220.0" valueMin="205.0">205–220 in</quantity>
diameter. Rostellar pouch 250 × 125, weakly muscular.
<taxonomicName id="EEF580CDFFFCA54CFB90F886FB39F8A2" authorityName="Montfort" authorityYear="1810" box="[1140,1261,1892,1918]" family="Rostellum" pageId="10" pageNumber="18" rank="family">Rostellum</taxonomicName>
250 × 100, bearing an incomplete (14 hooks observed, likely&gt; 20 when complete) double crown of subequal hooks, 46–49 long, with small epyphisis on guard. Genital pores alternating mostly regularly. Testes in single posterior field, in two layers,
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number. Cirrus-sac very large and strong, reaching antiporal excretory canals. Cirrus armed terminally. Vagina opening anteriorly (or dorsally) to cirrus-sac in a wellmarked ductus hermaphroditus connected to deep and wide atrium. Vagina dorsal to cirrus-sac strongly marked, terminating with characteristic sclerotized clamp [“dumb-bell organ”, see
<bibRefCitation id="4D6486BFFFFAA54AFB56FEB0FA99FEB0" author="Bona F. V." box="[1202,1357,338,364]" pageId="12" pageNumber="13" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">
<collectingCountry id="51E2BBDEFFFAA54AFB56FEB0FB24FEB0" box="[1202,1264,338,364]" name="Algeria" pageId="12" pageNumber="13">Bona</collectingCountry>
(1994)
</bibRefCitation>
, or “vaginal sclerites”, see
<bibRefCitation id="4D6486BFFFFAA54AFE37FE97FD39FE53" author="Dimitrova Y. D. &amp; Mariaux J. &amp; Georgiev B. B." box="[467,749,372,399]" pageId="12" pageNumber="13" pagination="215 - 233" refId="ref19097" refString="Dimitrova Y. D., Mariaux J. &amp; Georgiev B. B. 2013. Pseudangularia gonzalezi sp. nov. and Gibsonilepis swifti (Singh, 1952) n. g., n. comb. (Cestoda, Dilepididae) from the House Swift, Apus affinis (J. E. Gray) (Aves, Apodiformes) from Franceville, Republic of Gabon. Systematic Parasitology 86: 215 - 233. https: // doi. org / 10.1007 / s 11230 - 013 - 9445 - 4" type="journal article" year="2013">
Dimitrova
<emphasis id="1B81275CFFFAA54AFDB0FE94FD44FE53" box="[596,656,373,399]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2013)
</bibRefCitation>
]. Ovary central, deeply multilobulated, with digitiform lobes. Genital ducts dorsal to osmoregulatory canals. Young uterus saccular.
</paragraph>
</subSubSection>
<caption id="7D8AABC6FFFDA54DFF58FD4CF9F2FD16" box="[188,1574,686,714]" pageId="11" pageNumber="12">
<paragraph id="294AFB4EFFFDA54DFF58FD4CF9F2FD16" blockId="11.[188,1574,686,714]" box="[188,1574,686,714]" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFF58FD4CFECBFD16" bold="true" box="[188,287,686,714]" pageId="11" pageNumber="12">Table 2.</emphasis>
Comparison of
<taxonomicName id="EEF580CDFFFDA54DFE38FD4CFD08FD16" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[476,732,686,714]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="beveridgei" status="sp. nov.">
<emphasis id="1B81275CFFFDA54DFE38FD4CFD08FD16" box="[476,732,686,714]" italics="true" pageId="11" pageNumber="12">Spiniglans beveridgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFFDA54DFD07FD4CFCE9FD16" box="[739,829,686,714]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
with the other species of the genus
<taxonomicName id="EEF580CDFFFDA54DFB05FD4CF9F6FD16" authority="Yamaguti, 1959" authorityName="Yamaguti" authorityYear="1959" box="[1249,1570,686,714]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFFDA54DFB05FD4CFA89FD16" box="[1249,1373,686,714]" italics="true" pageId="11" pageNumber="12">Spiniglans</emphasis>
Yamaguti, 1959
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="294AFB4EFFFDA54DFF23FCA2F857F9E0" pageId="11" pageNumber="12">
<table id="5BF509EEFFFD5AB9FF23FCC2F852F990" box="[199,1926,800,1612]" gridcols="13" gridrows="8" pageId="11" pageNumber="12">
<tr id="97C5F90CFFFD5AB9FF23FCC2F852FCAB" box="[199,1926,800,887]" gridrow="0" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FCC2FE59FCAB" box="[199,397,800,887]" gridcol="0" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFF23FCA2FEC6FC8B" bold="true" box="[199,274,832,855]" pageId="11" pageNumber="12">Species</emphasis>
</th>
<th id="D4149070FFFD5AB9FE4DFCC2FDF6FCAB" box="[425,546,800,887]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFE2AFCA2FE2AFC8B" bold="true" box="[462,510,832,855]" pageId="11" pageNumber="12">Host</emphasis>
</th>
<th id="D4149070FFFD5AB9FDD9FCC2FD7AFCAB" box="[573,686,800,887]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFDB8FCD2FD4FFCBB" bold="true" pageId="11" pageNumber="12">Type locality</emphasis>
</th>
<th id="D4149070FFFD5AB9FD2CFCC2FCF0FCAB" box="[712,804,800,887]" gridcol="3" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFD31FCD2FCF0FCBB" bold="true" pageId="11" pageNumber="12">Scolex diameter</emphasis>
</th>
<th id="D4149070FFFD5AB9FCA5FCC2FC49FCAB" box="[833,925,800,887]" gridcol="4" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFCA2FCD2FC49FCBB" bold="true" pageId="11" pageNumber="12">Suckers diameter</emphasis>
</th>
<th id="D4149070FFFD5AB9FC5FFCC2FBF7FCAB" box="[955,1059,800,887]" gridcol="5" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFC24FCD2FC2AFCBB" bold="true" box="[960,1053,816,871]" pageId="11" pageNumber="12">Rostellar sac</emphasis>
</th>
<th id="D4149070FFFD5AB9FBA5FCC2FB7DFCAB" box="[1089,1193,800,887]" gridcol="6" gridrow="0" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFBA5FCA2FB7DFC8B" authorityName="Montfort" authorityYear="1810" box="[1089,1193,832,855]" family="Rostellum" pageId="11" pageNumber="18" rank="family">
<emphasis id="1B81275CFFFDA54DFBA5FCA2FB7DFC8B" bold="true" box="[1089,1193,832,855]" pageId="11" pageNumber="12">Rostellum</emphasis>
</taxonomicName>
</th>
<th id="D4149070FFFD5AB9FB2EFCC2FACFFCAB" box="[1226,1307,800,887]" gridcol="7" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFB36FCD2FACFFCBB" bold="true" pageId="11" pageNumber="12">Hooks number</emphasis>
</th>
<th id="D4149070FFFD5AB9FADCFCC2FA45FCAB" box="[1336,1425,800,887]" gridcol="8" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFADCFCC2FA51FCAB" bold="true" pageId="11" pageNumber="12">Anterior hooks length</emphasis>
</th>
<th id="D4149070FFFD5AB9FA49FCC2F9D8FCAB" box="[1453,1548,800,887]" gridcol="9" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DFA49FCC2FA29FCAB" bold="true" pageId="11" pageNumber="12">Posterior hooks length</emphasis>
</th>
<th id="D4149070FFFD5AB9F9CDFCC2F9AEFCAB" box="[1577,1658,800,887]" gridcol="10" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DF9D6FCD2F9AEFCBB" bold="true" pageId="11" pageNumber="12">Testes number</emphasis>
</th>
<th id="D4149070FFFD5AB9F976FCC2F92CFCAB" box="[1682,1784,800,887]" gridcol="11" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DF947FCD2F901FCBB" bold="true" box="[1699,1767,816,871]" pageId="11" pageNumber="12">Cirrus sac</emphasis>
</th>
<th id="D4149070FFFD5AB9F8E8FCC2F852FCAB" box="[1804,1926,800,887]" gridcol="12" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="1B81275CFFFDA54DF8E8FCA2F886FC8B" bold="true" box="[1804,1874,832,855]" pageId="11" pageNumber="12">Source</emphasis>
</th>
</tr>
<tr id="97C5F90CFFFD5AB9FF23FC7AF852FC13" box="[199,1926,920,975]" gridrow="1" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FC7AFE59FC13" box="[199,397,920,975]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFF23FC7AFE90FC73" authority="Mariaux &amp; Georgiev, 2018" authorityName="Mariaux &amp; Georgiev" authorityYear="2018" box="[199,324,920,943]" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="beveridgei" status="sp. nov.">
<emphasis id="1B81275CFFFDA54DFF23FC7AFE90FC73" box="[199,324,920,943]" italics="true" pageId="11" pageNumber="12">S. beveridgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B29A27FFFDA54DFF23FC5AFEC6FC13" box="[199,274,952,975]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="D4149070FFFD5AB9FE4DFC7AFDF6FC13" box="[425,546,920,975]" gridcol="1" gridrow="1" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFE5EFC4AFDC6FC63" box="[442,530,936,959]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Corvidae</taxonomicName>
</td>
<td id="D4149070FFFD5AB9FDD9FC7AFD7AFC13" box="[573,686,920,975]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">Australia</td>
<td id="D4149070FFFD5AB9FD2CFC7AFCF0FC13" box="[712,804,920,975]" gridcol="3" gridrow="1" pageId="11" pageNumber="12">325–450</td>
<td id="D4149070FFFD5AB9FCA5FC7AFC49FC13" box="[833,925,920,975]" gridcol="4" gridrow="1" pageId="11" pageNumber="12">145–180</td>
<td id="D4149070FFFD5AB9FC5FFC7AFBF7FC13" box="[955,1059,920,975]" gridcol="5" gridrow="1" pageId="11" pageNumber="12">374–450 × 132–152</td>
<td id="D4149070FFFD5AB9FBA5FC7AFB7DFC13" box="[1089,1193,920,975]" gridcol="6" gridrow="1" pageId="11" pageNumber="12">224–270 × 85–105</td>
<td id="D4149070FFFD5AB9FB2EFC7AFACFFC13" box="[1226,1307,920,975]" gridcol="7" gridrow="1" pageId="11" pageNumber="12">20–22</td>
<td id="D4149070FFFD5AB9FADCFC7AFA45FC13" box="[1336,1425,920,975]" gridcol="8" gridrow="1" pageId="11" pageNumber="12">40.5–44</td>
<td id="D4149070FFFD5AB9FA49FC7AF9D8FC13" box="[1453,1548,920,975]" gridcol="9" gridrow="1" pageId="11" pageNumber="12">38.5–40.5</td>
<td id="D4149070FFFD5AB9F9CDFC7AF9AEFC13" box="[1577,1658,920,975]" gridcol="10" gridrow="1" pageId="11" pageNumber="12">40–58</td>
<td id="D4149070FFFD5AB9F976FC7AF92CFC13" box="[1682,1784,920,975]" gridcol="11" gridrow="1" pageId="11" pageNumber="12">87–118 × 47–68</td>
<td id="D4149070FFFD5AB9F8E8FC7AF852FC13" box="[1804,1926,920,975]" gridcol="12" gridrow="1" pageId="11" pageNumber="12">New data</td>
</tr>
<tr id="97C5F90CFFFD5AB9FF23FC18F852FBED" box="[199,1926,1018,1073]" gridrow="2" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FC18FE59FBED" box="[199,397,1018,1073]" gridcol="0" gridrow="2" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFF23FC18FE8EFBED" authority="(Krabbe, 1869)" baseAuthorityName="Krabbe" baseAuthorityYear="1869" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="affinis">
<emphasis id="1B81275CFFFDA54DFF23FC18FECEFBCD" box="[199,282,1018,1041]" italics="true" pageId="11" pageNumber="12">S. affinis</emphasis>
(
<bibRefCitation id="4D6486BFFFFDA54DFF29FBF8FE86FBED" author="Krabbe H." box="[205,338,1050,1073]" pageId="11" pageNumber="12" pagination="251 - 389" refId="ref20181" refString="Krabbe H. 1869. Bidrag til kundskab om fuglenes baendelorme. Videnskabelige Selskab Skrifter, 5. raekke, Naturvidenskabelig og Mathematisk Afdel 8: 251 - 389." type="journal article" year="1869">Krabbe, 1869</bibRefCitation>
)
</taxonomicName>
</th>
<td id="D4149070FFFD5AB9FE4DFC18FDF6FBED" box="[425,546,1018,1073]" gridcol="1" gridrow="2" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFE5EFBE8FDC6FBFD" box="[442,530,1034,1057]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Corvidae</taxonomicName>
</td>
<td id="D4149070FFFD5AB9FDD9FC18FD7AFBED" box="[573,686,1018,1073]" gridcol="2" gridrow="2" pageId="11" pageNumber="12">Ukraine</td>
<td id="D4149070FFFD5AB9FD2CFC18FCF0FBED" box="[712,804,1018,1073]" gridcol="3" gridrow="2" pageId="11" pageNumber="12">550</td>
<td id="D4149070FFFD5AB9FCA5FC18FC49FBED" box="[833,925,1018,1073]" gridcol="4" gridrow="2" pageId="11" pageNumber="12">150</td>
<td id="D4149070FFFD5AB9FC5FFC18FBF7FBED" box="[955,1059,1018,1073]" gridcol="5" gridrow="2" pageId="11" pageNumber="12">420 × 200</td>
<td id="D4149070FFFD5AB9FBA5FC18FB7DFBED" box="[1089,1193,1018,1073]" gridcol="6" gridrow="2" pageId="11" pageNumber="12">350 × 120</td>
<td id="D4149070FFFD5AB9FB2EFC18FACFFBED" box="[1226,1307,1018,1073]" gridcol="7" gridrow="2" pageId="11" pageNumber="12">22</td>
<td id="D4149070FFFD5AB9FADCFC18FA45FBED" box="[1336,1425,1018,1073]" gridcol="8" gridrow="2" pageId="11" pageNumber="12">55–58</td>
<td id="D4149070FFFD5AB9FA49FC18F9D8FBED" box="[1453,1548,1018,1073]" gridcol="9" gridrow="2" pageId="11" pageNumber="12">50–55</td>
<td id="D4149070FFFD5AB9F9CDFC18F9AEFBED" box="[1577,1658,1018,1073]" gridcol="10" gridrow="2" pageId="11" pageNumber="12">78–86</td>
<td id="D4149070FFFD5AB9F976FC18F92CFBED" box="[1682,1784,1018,1073]" gridcol="11" gridrow="2" pageId="11" pageNumber="12">75–100 × 25–30</td>
<td id="D4149070FFFD5AB9F8E8FC18F852FBED" box="[1804,1926,1018,1073]" gridcol="12" gridrow="2" pageId="11" pageNumber="12">
<bibRefCitation id="4D6486BFFFFDA54DF8E8FC18F898FBED" author="Salamatin R. V." box="[1804,1900,1018,1073]" pageId="11" pageNumber="12" pagination="75 - 81" refId="ref21194" refString="Salamatin R. V. 1999. The cestodes of the genus Spiniglans (Dilepididae) from the corvid birds (Corvidae) from Ukraine. Vestnik Zoologii 33: 75 - 81. [In Russian.]" type="journal article" year="1999">Salamatin (1999)</bibRefCitation>
</td>
</tr>
<tr id="97C5F90CFFFD5AB9FF23FBBFF852FB48" box="[199,1926,1117,1172]" gridrow="3" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FBBFFE59FB48" box="[199,397,1117,1172]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFF23FBBFFE9AFB48" authority="(Molin, 1858)" baseAuthorityName="Molin" baseAuthorityYear="1858" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="constricta">
<emphasis id="1B81275CFFFDA54DFF23FBBFFEEBFBA8" box="[199,319,1117,1140]" italics="true" pageId="11" pageNumber="12">S. constricta</emphasis>
(Molin, 1858)
</taxonomicName>
</th>
<td id="D4149070FFFD5AB9FE4DFBBFFDF6FB48" box="[425,546,1117,1172]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFE5EFB8FFDC6FB58" box="[442,530,1133,1156]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Corvidae</taxonomicName>
</td>
<td id="D4149070FFFD5AB9FDD9FBBFFD7AFB48" box="[573,686,1117,1172]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">Ukraine</td>
<td id="D4149070FFFD5AB9FD2CFBBFFCF0FB48" box="[712,804,1117,1172]" gridcol="3" gridrow="3" pageId="11" pageNumber="12">330</td>
<td id="D4149070FFFD5AB9FCA5FBBFFC49FB48" box="[833,925,1117,1172]" gridcol="4" gridrow="3" pageId="11" pageNumber="12">120</td>
<td id="D4149070FFFD5AB9FC5FFBBFFBF7FB48" box="[955,1059,1117,1172]" gridcol="5" gridrow="3" pageId="11" pageNumber="12">250 × 134</td>
<td id="D4149070FFFD5AB9FBA5FBBFFB7DFB48" box="[1089,1193,1117,1172]" gridcol="6" gridrow="3" pageId="11" pageNumber="12">200 × 120</td>
<td id="D4149070FFFD5AB9FB2EFBBFFACFFB48" box="[1226,1307,1117,1172]" gridcol="7" gridrow="3" pageId="11" pageNumber="12">20</td>
<td id="D4149070FFFD5AB9FADCFBBFFA45FB48" box="[1336,1425,1117,1172]" gridcol="8" gridrow="3" pageId="11" pageNumber="12">40–41</td>
<td id="D4149070FFFD5AB9FA49FBBFF9D8FB48" box="[1453,1548,1117,1172]" gridcol="9" gridrow="3" pageId="11" pageNumber="12">36–37</td>
<td id="D4149070FFFD5AB9F9CDFBBFF9AEFB48" box="[1577,1658,1117,1172]" gridcol="10" gridrow="3" pageId="11" pageNumber="12">38–49</td>
<td id="D4149070FFFD5AB9F976FBBFF92CFB48" box="[1682,1784,1117,1172]" gridcol="11" gridrow="3" pageId="11" pageNumber="12">60–80 × 30–35</td>
<td id="D4149070FFFD5AB9F8E8FBBFF852FB48" box="[1804,1926,1117,1172]" gridcol="12" gridrow="3" pageId="11" pageNumber="12">
<bibRefCitation id="4D6486BFFFFDA54DF8E8FBBFF898FB48" author="Salamatin R. V." box="[1804,1900,1117,1172]" pageId="11" pageNumber="12" pagination="75 - 81" refId="ref21194" refString="Salamatin R. V. 1999. The cestodes of the genus Spiniglans (Dilepididae) from the corvid birds (Corvidae) from Ukraine. Vestnik Zoologii 33: 75 - 81. [In Russian.]" type="journal article" year="1999">Salamatin (1999)</bibRefCitation>
</td>
</tr>
<tr id="97C5F90CFFFD5AB9FF23FB5DF852FB2A" box="[199,1926,1215,1270]" gridrow="4" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FB5DFE59FB2A" box="[199,397,1215,1270]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFF23FB5DFE59FB2A" authority="(Joyeux et al., 1937)" baseAuthorityName="Joyeux" baseAuthorityYear="1937" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="corvi">
<emphasis id="1B81275CFFFDA54DFF23FB5DFEC4FB0A" box="[199,272,1215,1238]" italics="true" pageId="11" pageNumber="12">S. corvi</emphasis>
(
<bibRefCitation id="4D6486BFFFFDA54DFF29FB3DFE52FB2A" author="Joyeux C. &amp; Baer J. G. &amp; Martin R." box="[205,390,1247,1270]" pageId="11" pageNumber="12" pagination="416 - 423" refId="ref19954" refString="Joyeux C., Baer J. G. &amp; Martin R. 1937. Sur quelques helminthes de la Somalie-Nord (Deuxieme note). Bulletin de la Societe de Pathologie exotique 30: 416 - 423." type="journal article" year="1937">
Joyeux
<emphasis id="1B81275CFFFDA54DFEFDFB3DFE97FB2A" box="[281,323,1247,1270]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 1937
</bibRefCitation>
)
</taxonomicName>
</th>
<td id="D4149070FFFD5AB9FE4DFB5DFDF6FB2A" box="[425,546,1215,1270]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFE5EFB2DFDC6FB3A" box="[442,530,1231,1254]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Corvidae</taxonomicName>
</td>
<td id="D4149070FFFD5AB9FDD9FB5DFD7AFB2A" box="[573,686,1215,1270]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">Somalia</td>
<td id="D4149070FFFD5AB9FD2CFB5DFCF0FB2A" box="[712,804,1215,1270]" gridcol="3" gridrow="4" pageId="11" pageNumber="12">260</td>
<td id="D4149070FFFD5AB9FCA5FB5DFC49FB2A" box="[833,925,1215,1270]" gridcol="4" gridrow="4" pageId="11" pageNumber="12">170</td>
<td id="D4149070FFFD5AB9FC5FFB5DFBF7FB2A" box="[955,1059,1215,1270]" gridcol="5" gridrow="4" pageId="11" pageNumber="12">350–430 × 110</td>
<td id="D4149070FFFD5AB9FBA5FB5DFB7DFB2A" box="[1089,1193,1215,1270]" gridcol="6" gridrow="4" pageId="11" pageNumber="12">– × 160</td>
<td id="D4149070FFFD5AB9FB2EFB5DFACFFB2A" box="[1226,1307,1215,1270]" gridcol="7" gridrow="4" pageId="11" pageNumber="12">22</td>
<td id="D4149070FFFD5AB9FADCFB5DFA45FB2A" box="[1336,1425,1215,1270]" gridcol="8" gridrow="4" pageId="11" pageNumber="12">43–45</td>
<td id="D4149070FFFD5AB9FA49FB5DF9D8FB2A" box="[1453,1548,1215,1270]" gridcol="9" gridrow="4" pageId="11" pageNumber="12">–</td>
<td id="D4149070FFFD5AB9F9CDFB5DF9AEFB2A" box="[1577,1658,1215,1270]" gridcol="10" gridrow="4" pageId="11" pageNumber="12">32–35</td>
<td id="D4149070FFFD5AB9F976FB5DF92CFB2A" box="[1682,1784,1215,1270]" gridcol="11" gridrow="4" pageId="11" pageNumber="12">140 × 50</td>
<td id="D4149070FFFD5AB9F8E8FB5DF852FB2A" box="[1804,1926,1215,1270]" gridcol="12" gridrow="4" pageId="11" pageNumber="12">
<bibRefCitation id="4D6486BFFFFDA54DF8E8FB5DF898FB2A" author="Joyeux C. &amp; Baer J. G. &amp; Martin R." pageId="11" pageNumber="12" pagination="416 - 423" refId="ref19954" refString="Joyeux C., Baer J. G. &amp; Martin R. 1937. Sur quelques helminthes de la Somalie-Nord (Deuxieme note). Bulletin de la Societe de Pathologie exotique 30: 416 - 423." type="journal article" year="1937">
Joyeux
<emphasis id="1B81275CFFFDA54DF8B2FB5DF855FB0A" box="[1878,1921,1215,1238]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (1937)
</bibRefCitation>
</td>
</tr>
<tr id="97C5F90CFFFD5AB9FF23FAF5F852FAB2" box="[199,1926,1303,1390]" gridrow="5" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FAF5FE59FAB2" box="[199,397,1303,1390]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFF23FAF5FEA0FA92" authority="(Southwell, 1922)" baseAuthorityName="Southwell" baseAuthorityYear="1922" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="microsoma">
<emphasis id="1B81275CFFFDA54DFF23FAF5FE9DFAF2" box="[199,329,1303,1326]" italics="true" pageId="11" pageNumber="12">S. microsoma</emphasis>
(
<bibRefCitation id="4D6486BFFFFDA54DFF29FAD5FEB9FA92" author="Southwell T." box="[205,365,1335,1358]" pageId="11" pageNumber="12" pagination="355 - 382" refId="ref21349" refString="Southwell T. 1922. Cestodes from Indian birds with a note on Ligula intestinalis. Annals of Tropical Medicine &amp; Parasitology 16: 355 - 382. https: // doi. org / 10.1080 / 00034983.1922.11684330" type="journal article" year="1922">Southwell, 1922</bibRefCitation>
)
</taxonomicName>
(Type species)
</th>
<td id="D4149070FFFD5AB9FE4DFAF5FDF6FAB2" box="[425,546,1303,1390]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFE50FAC5FDC0FAE2" authorityName="Sundevall" authorityYear="1836" box="[436,532,1319,1342]" class="Aves" family="Ploceidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Ploceidae</taxonomicName>
,
<taxonomicName id="EEF580CDFFFDA54DFE4DFAA5FDF6FA82" authorityName="Vigors" authorityYear="1831" box="[425,546,1351,1374]" class="Aves" family="Emberizidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Emberizidae</taxonomicName>
</td>
<td id="D4149070FFFD5AB9FDD9FAF5FD7AFAB2" box="[573,686,1303,1390]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">India (Zoological Garden)</td>
<td id="D4149070FFFD5AB9FD2CFAF5FCF0FAB2" box="[712,804,1303,1390]" gridcol="3" gridrow="5" pageId="11" pageNumber="12">220</td>
<td id="D4149070FFFD5AB9FCA5FAF5FC49FAB2" box="[833,925,1303,1390]" gridcol="4" gridrow="5" pageId="11" pageNumber="12">140</td>
<td id="D4149070FFFD5AB9FC5FFAF5FBF7FAB2" box="[955,1059,1303,1390]" gridcol="5" gridrow="5" pageId="11" pageNumber="12">–</td>
<td id="D4149070FFFD5AB9FBA5FAF5FB7DFAB2" box="[1089,1193,1303,1390]" gridcol="6" gridrow="5" pageId="11" pageNumber="12">180 × 50</td>
<td id="D4149070FFFD5AB9FB2EFAF5FACFFAB2" box="[1226,1307,1303,1390]" gridcol="7" gridrow="5" pageId="11" pageNumber="12">16–20</td>
<td id="D4149070FFFD5AB9FADCFAF5FA45FAB2" box="[1336,1425,1303,1390]" gridcol="8" gridrow="5" pageId="11" pageNumber="12">35</td>
<td id="D4149070FFFD5AB9FA49FAF5F9D8FAB2" box="[1453,1548,1303,1390]" gridcol="9" gridrow="5" pageId="11" pageNumber="12">–</td>
<td id="D4149070FFFD5AB9F9CDFAF5F9AEFAB2" box="[1577,1658,1303,1390]" gridcol="10" gridrow="5" pageId="11" pageNumber="12">16–20</td>
<td id="D4149070FFFD5AB9F976FAF5F92CFAB2" box="[1682,1784,1303,1390]" gridcol="11" gridrow="5" pageId="11" pageNumber="12">short and narrow</td>
<td id="D4149070FFFD5AB9F8E8FAF5F852FAB2" box="[1804,1926,1303,1390]" gridcol="12" gridrow="5" pageId="11" pageNumber="12">
<bibRefCitation id="4D6486BFFFFDA54DF8E8FAC5F899FA82" author="Southwell T." box="[1804,1901,1319,1374]" pageId="11" pageNumber="12" pagination="355 - 382" refId="ref21349" refString="Southwell T. 1922. Cestodes from Indian birds with a note on Ligula intestinalis. Annals of Tropical Medicine &amp; Parasitology 16: 355 - 382. https: // doi. org / 10.1080 / 00034983.1922.11684330" type="journal article" year="1922">Southwell (1922)</bibRefCitation>
</td>
</tr>
<tr id="97C5F90CFFFD5AB9FF23FA64F852FA01" box="[199,1926,1414,1501]" gridrow="6" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FA64FE59FA01" box="[199,397,1414,1501]" gridcol="0" gridrow="6" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFF23FA64FF2BFA01" authority="(Georgiev et al., 1987)" baseAuthorityName="Georgiev" baseAuthorityYear="1987" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="pirinica">
<emphasis id="1B81275CFFFDA54DFF23FA64FEFFFA41" box="[199,299,1414,1437]" italics="true" pageId="11" pageNumber="12">S. pirinica</emphasis>
(
<bibRefCitation id="4D6486BFFFFDA54DFF29FA44FF2CFA01" author="Georgiev B. B. &amp; Kornyushin V. V. &amp; Genov T." pageId="11" pageNumber="12" pagination="3 - 7" refId="ref19658" refString="Georgiev B. B., Kornyushin V. V. &amp; Genov T. 1987. Choanotaenia pirinica sp. n. (Cestoda, Dilepididae) - a parasite of the Alpine Chough in Bulgaria. Vestnik Zoologii 1987: 3 - 7. [In Russian.]" type="journal article" year="1987">
Georgiev
<emphasis id="1B81275CFFFDA54DFECAFA44FE8CFA61" box="[302,344,1446,1469]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 1987
</bibRefCitation>
)
</taxonomicName>
</th>
<td id="D4149070FFFD5AB9FE4DFA64FDF6FA01" box="[425,546,1414,1501]" gridcol="1" gridrow="6" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFE5EFA44FDC6FA61" box="[442,530,1446,1469]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Corvidae</taxonomicName>
</td>
<td id="D4149070FFFD5AB9FDD9FA64FD7AFA01" box="[573,686,1414,1501]" gridcol="2" gridrow="6" pageId="11" pageNumber="12">Bulgaria</td>
<td id="D4149070FFFD5AB9FD2CFA64FCF0FA01" box="[712,804,1414,1501]" gridcol="3" gridrow="6" pageId="11" pageNumber="12">312–406</td>
<td id="D4149070FFFD5AB9FCA5FA64FC49FA01" box="[833,925,1414,1501]" gridcol="4" gridrow="6" pageId="11" pageNumber="12">114–175</td>
<td id="D4149070FFFD5AB9FC5FFA64FBF7FA01" box="[955,1059,1414,1501]" gridcol="5" gridrow="6" pageId="11" pageNumber="12">344–456 × 144–206</td>
<td id="D4149070FFFD5AB9FBA5FA64FB7DFA01" box="[1089,1193,1414,1501]" gridcol="6" gridrow="6" pageId="11" pageNumber="12">262–368 × 87–125</td>
<td id="D4149070FFFD5AB9FB2EFA64FACFFA01" box="[1226,1307,1414,1501]" gridcol="7" gridrow="6" pageId="11" pageNumber="12">20–22</td>
<td id="D4149070FFFD5AB9FADCFA64FA45FA01" box="[1336,1425,1414,1501]" gridcol="8" gridrow="6" pageId="11" pageNumber="12">67–73</td>
<td id="D4149070FFFD5AB9FA49FA64F9D8FA01" box="[1453,1548,1414,1501]" gridcol="9" gridrow="6" pageId="11" pageNumber="12">61–68</td>
<td id="D4149070FFFD5AB9F9CDFA64F9AEFA01" box="[1577,1658,1414,1501]" gridcol="10" gridrow="6" pageId="11" pageNumber="12">39–61</td>
<td id="D4149070FFFD5AB9F976FA64F92CFA01" box="[1682,1784,1414,1501]" gridcol="11" gridrow="6" pageId="11" pageNumber="12">85 × 110 × 32–45</td>
<td id="D4149070FFFD5AB9F8E8FA64F852FA01" box="[1804,1926,1414,1501]" gridcol="12" gridrow="6" pageId="11" pageNumber="12">
<bibRefCitation id="4D6486BFFFFDA54DF8E8FA74F856FA11" author="Georgiev B. B. &amp; Kornyushin V. V. &amp; Genov T." box="[1804,1922,1430,1485]" pageId="11" pageNumber="12" pagination="3 - 7" refId="ref19658" refString="Georgiev B. B., Kornyushin V. V. &amp; Genov T. 1987. Choanotaenia pirinica sp. n. (Cestoda, Dilepididae) - a parasite of the Alpine Chough in Bulgaria. Vestnik Zoologii 1987: 3 - 7. [In Russian.]" type="journal article" year="1987">
Georgiev
<emphasis id="1B81275CFFFDA54DF8E8FA54F8E9FA11" box="[1804,1853,1462,1485]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(1987)
</bibRefCitation>
</td>
</tr>
<tr id="97C5F90CFFFD5AB9FF23FA17F852F990" box="[199,1926,1525,1612]" gridrow="7" pageId="11" pageNumber="12">
<th id="D4149070FFFD5AB9FF23FA17FE59F990" box="[199,397,1525,1612]" gridcol="0" gridrow="7" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFF23FA17FF23F990" authority="Kornyushin et al., 2009" authorityName="Kornyushin" authorityYear="2009" class="Cestoda" family="Dilepididae" genus="Spiniglans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="11" pageNumber="12" phylum="Platyhelminthes" rank="species" species="sharpiloi">
<emphasis id="1B81275CFFFDA54DFF23FA17FEE2F9D0" box="[199,310,1525,1548]" italics="true" pageId="11" pageNumber="12">S. sharpiloi</emphasis>
<bibRefCitation id="4D6486BFFFFDA54DFF23F9F7FF23F990" author="Kornyushin V. V. &amp; Salamatin R. V. &amp; Greben O. B. &amp; Georgiev B. B." pageId="11" pageNumber="12" pagination="85 - 93" refId="ref20121" refString="Kornyushin V. V., Salamatin R. V., Greben O. B. &amp; Georgiev B. B. 2009. Spiniglans sharpiloi sp. n. (Cestoda, Dilepididae), a parasite of the common magpie, Pica pica, in the Palaearctic. Vestnik Zoologii Supplement 23: 85 - 93." type="journal article" year="2009">
Kornyushin
<emphasis id="1B81275CFFFDA54DFEDAF9F7FEBDF9F0" box="[318,361,1557,1580]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 2009
</bibRefCitation>
</taxonomicName>
</th>
<td id="D4149070FFFD5AB9FE4DFA17FDF6F990" box="[425,546,1525,1612]" gridcol="1" gridrow="7" pageId="11" pageNumber="12">
<taxonomicName id="EEF580CDFFFDA54DFE5EF9E7FDC6F9C0" box="[442,530,1541,1564]" class="Aves" family="Corvidae" kingdom="Animalia" order="Passeriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Corvidae</taxonomicName>
</td>
<td id="D4149070FFFD5AB9FDD9FA17FD7AF990" box="[573,686,1525,1612]" gridcol="2" gridrow="7" pageId="11" pageNumber="12">Ukraine</td>
<td id="D4149070FFFD5AB9FD2CFA17FCF0F990" box="[712,804,1525,1612]" gridcol="3" gridrow="7" pageId="11" pageNumber="12">276–530</td>
<td id="D4149070FFFD5AB9FCA5FA17FC49F990" box="[833,925,1525,1612]" gridcol="4" gridrow="7" pageId="11" pageNumber="12">130–210</td>
<td id="D4149070FFFD5AB9FC5FFA17FBF7F990" box="[955,1059,1525,1612]" gridcol="5" gridrow="7" pageId="11" pageNumber="12">301–350 × 112–220</td>
<td id="D4149070FFFD5AB9FBA5FA17FB7DF990" box="[1089,1193,1525,1612]" gridcol="6" gridrow="7" pageId="11" pageNumber="12">180–200 × 75–110</td>
<td id="D4149070FFFD5AB9FB2EFA17FACFF990" box="[1226,1307,1525,1612]" gridcol="7" gridrow="7" pageId="11" pageNumber="12">20–22</td>
<td id="D4149070FFFD5AB9FADCFA17FA45F990" box="[1336,1425,1525,1612]" gridcol="8" gridrow="7" pageId="11" pageNumber="12">31–38</td>
<td id="D4149070FFFD5AB9FA49FA17F9D8F990" box="[1453,1548,1525,1612]" gridcol="9" gridrow="7" pageId="11" pageNumber="12">28–36</td>
<td id="D4149070FFFD5AB9F9CDFA17F9AEF990" box="[1577,1658,1525,1612]" gridcol="10" gridrow="7" pageId="11" pageNumber="12">32–45</td>
<td id="D4149070FFFD5AB9F976FA17F92CF990" box="[1682,1784,1525,1612]" gridcol="11" gridrow="7" pageId="11" pageNumber="12">80–131 × 30–50</td>
<td id="D4149070FFFD5AB9F8E8FA17F852F990" box="[1804,1926,1525,1612]" gridcol="12" gridrow="7" pageId="11" pageNumber="12">
<bibRefCitation id="4D6486BFFFFDA54DF8E8F9E7F857F9E0" author="Kornyushin V. V. &amp; Salamatin R. V. &amp; Greben O. B. &amp; Georgiev B. B." box="[1804,1923,1541,1596]" pageId="11" pageNumber="12" pagination="85 - 93" refId="ref20121" refString="Kornyushin V. V., Salamatin R. V., Greben O. B. &amp; Georgiev B. B. 2009. Spiniglans sharpiloi sp. n. (Cestoda, Dilepididae), a parasite of the common magpie, Pica pica, in the Palaearctic. Vestnik Zoologii Supplement 23: 85 - 93." type="journal article" year="2009">
Kornyushin
<emphasis id="1B81275CFFFDA54DF8E8F9C7F8E3F9E0" box="[1804,1847,1573,1596]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (2009)
</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<subSubSection id="61EFA8C5FFFAA54AFF59FE38FEC1FD29" pageId="12" pageNumber="13" type="discussion">
<paragraph id="294AFB4EFFFAA54AFF59FE38FEFBFE28" blockId="12.[189,303,474,500]" box="[189,303,474,500]" pageId="12" pageNumber="13">
<heading id="72024C22FFFAA54AFF59FE38FEFBFE28" bold="true" box="[189,303,474,500]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="1B81275CFFFAA54AFF59FE38FEFBFE28" bold="true" box="[189,303,474,500]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="294AFB4EFFFAA54AFF59FDEAFEC1FD29" blockId="12.[189,1399,520,757]" pageId="12" pageNumber="13">
This material belongs to the genus
<taxonomicName id="EEF580CDFFFAA54AFDBFFDEAFC9AFDFF" authority="Singh, 1952" authorityName="Singh" authorityYear="1952" box="[603,846,520,547]" class="Cestoda" family="Dilepididae" genus="Neoliga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="12" pageNumber="13" phylum="Platyhelminthes" rank="genus">
<emphasis id="1B81275CFFFAA54AFDBFFDEAFD6CFDFE" box="[603,696,520,546]" italics="true" pageId="12" pageNumber="13">Neoliga</emphasis>
<bibRefCitation id="4D6486BFFFFAA54AFD24FDEAFC9AFDFF" author="Singh K. S." box="[704,846,520,547]" pageId="12" pageNumber="13" pagination="1 - 72" refId="ref21294" refString="Singh K. S. 1952. Cestode parasites of birds. Indian Journal of Helminthology 4: 1 - 72." type="journal article" year="1952">Singh, 1952</bibRefCitation>
</taxonomicName>
as defined by
<bibRefCitation id="4D6486BFFFFAA54AFC1EFDEBFB45FDFE" author="Bona F. V." box="[1018,1169,520,547]" pageId="12" pageNumber="13" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">
<collectingCountry id="51E2BBDEFFFAA54AFC1EFDEBFBECFDFF" box="[1018,1080,521,547]" name="Algeria" pageId="12" pageNumber="13">Bona</collectingCountry>
(1994)
</bibRefCitation>
, a genus specific to swifts and tree swifts (
<taxonomicName id="EEF580CDFFFAA54AFE20FDC9FDEEFD9A" box="[452,570,555,582]" class="Aves" family="Apodidae" kingdom="Animalia" order="Apodiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="family">Apodidae</taxonomicName>
) and known from
<collectingCountry id="51E2BBDEFFFAA54AFCF5FDCEFCAFFD9A" box="[785,891,556,582]" name="Morocco" pageId="12" pageNumber="13">Morocco</collectingCountry>
to
<collectingCountry id="51E2BBDEFFFAA54AFC44FDC9FC0BFD99" box="[928,991,555,581]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
, Java and Siberia. Four species are listed by
<bibRefCitation id="4D6486BFFFFAA54AFEC2FDACFDAFFDB5" author="Spasskaya L. P. &amp; Spasskii A." box="[294,635,590,617]" pageId="12" pageNumber="13" refId="ref21515" refString="Spasskaya L. P. &amp; Spasskii A. 1977. Cestodes of Birds in the USSR. Dilepididae of Terrestrial Birds. Izdate'stvo Nauka, Moscow. [In Russian.]" type="book" year="1977">Spasskaya &amp; Spasskii (1977)</bibRefCitation>
, but the present material does not allow for a detailed comparison. There is no known species of this genus in
<collectingCountry id="51E2BBDEFFFAA54AFD31FD90FC90FD50" box="[725,836,626,652]" name="Australia" pageId="12" pageNumber="13">Australia</collectingCountry>
, but given the migratory pattern of the whitethroated needletail it is quite possible that parasites from Siberian hosts could be found in
<collectingCountry id="51E2BBDEFFFAA54AFAE0FD77FAA7FD73" box="[1284,1395,661,687]" name="Australia" pageId="12" pageNumber="13">Australia</collectingCountry>
. This confirms the prediction of
<bibRefCitation id="4D6486BFFFFAA54AFDC9FD5BFD14FD0E" author="Bona F. V." box="[557,704,696,723]" pageId="12" pageNumber="13" pagination="443 - 554" refId="ref18725" refString="Bona F. V. 1994. Family Dilepididae Railliet &amp; Henry, 1909. In: Khalil L. F., Jones A. &amp; Bray R. A. (eds) Keys to the Cestode Parasites of Vertebrates: 443 - 554. CAB International, Wallingford." type="book chapter" year="1994">
<collectingCountry id="51E2BBDEFFFAA54AFDC9FD5BFDBFFD0F" box="[557,619,697,723]" name="Algeria" pageId="12" pageNumber="13">Bona</collectingCountry>
(1994)
</bibRefCitation>
that the genus distribution is probably wider than presently known.
</paragraph>
</subSubSection>
</treatment>
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<subSubSection id="61EF6510FFC06D71D6DDDE10BFFD0101" box="[151,506,1831,1858]" pageId="41" pageNumber="292" type="nomenclature">
<paragraph id="294A369BFFC06D71D6DDDE10BFFD0101" blockId="41.[151,506,1831,1894]" box="[151,506,1831,1858]" pageId="41" pageNumber="292">
<heading id="720281F7FFC06D71D6DDDE10BFFD0101" bold="true" box="[151,506,1831,1858]" fontSize="11" level="1" pageId="41" pageNumber="292" reason="1">
<emphasis id="1B81EA89FFC06D71D6DDDE10BFFD0101" bold="true" box="[151,506,1831,1858]" pageId="41" pageNumber="292">
<taxonomicName id="EEF54D18FFC06D71D6DDDE10BFB60101" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,433,1831,1858]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
<emphasis id="1B81EA89FFC06D71D6DDDE10BFB60101" bold="true" box="[151,433,1831,1858]" italics="true" pageId="41" pageNumber="292">Paraphloeostiba rufula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFC06D71D7F2DE1FBFFD0101" box="[440,506,1832,1858]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC06D71D6DDDE7BBF6A0125" box="[151,365,1868,1894]" pageId="41" pageNumber="292" type="description">
<paragraph id="294A369BFFC06D71D6DDDE7BBF6A0125" blockId="41.[151,506,1831,1894]" box="[151,365,1868,1894]" pageId="41" pageNumber="292">
(
<figureCitation id="B1CE2A1EFFC06D71D6D5DE7BBEF10125" box="[159,246,1868,1894]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="41" pageNumber="292">Figs 80</figureCitation>
,
<figureCitation id="B1CE2A1EFFC06D71D748DE7BBF620125" box="[258,357,1868,1894]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="41" pageNumber="292">126–132</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC06D72D6DDDEA3BAD606B4" lastPageId="42" lastPageNumber="293" pageId="41" pageNumber="292" type="materials_examined">
<paragraph id="294A369BFFC06D71D6DDDEA3BFAA01B5" blockId="41.[151,1436,1940,2038]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D6DDDEA3BF4101ED" bold="true" box="[151,326,1940,1966]" pageId="41" pageNumber="292">Type material.</emphasis>
<typeStatus id="F64E8839FFC06D71D704DEA3BFB101ED" box="[334,438,1940,1966]" pageId="41" pageNumber="292" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFC06D71D7F4DEA2BFD401EC" box="[446,467,1941,1967]" count="1" pageId="41" pageNumber="292" type="male">♂</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFC06D71D422DEA3BCAB01ED" box="[616,684,1940,1966]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="41" pageNumber="292">NEW</collectionCode>
<collectingCountry id="51E2760BFFC06D71D4FEDEA3BD1901ED" box="[692,798,1940,1966]" name="Guinea-Bissau" pageId="41" pageNumber="292">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFC06D71D5C6DEA3BA0901EC" box="[908,1038,1940,1967]" pageId="41" pageNumber="292" value="1965-06-16">16.VI.1965</date>
, | flying in daylight’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFC06D71D335DEA3BB9F01ED" box="[1407,1432,1940,1966]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="41" pageNumber="292">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC06D71D426DE8FBD020191" box="[620,773,1976,2002]" pageId="41" pageNumber="292" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC06D71D556DE8FBDD00191" authorityName="Steel" authorityYear="1960" box="[796,983,1976,2002]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC06D71D556DE8FBDD00191" box="[796,983,1976,2002]" italics="true" pageId="41" pageNumber="292">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC06D71D5A6DE8FBA370191" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1004,1072,1976,2002]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
<emphasis id="1B81EA89FFC06D71D5A6DE8FBA370191" box="[1004,1072,1976,2002]" italics="true" pageId="41" pageNumber="292">rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D273DE8FBA7D0191" bold="true" box="[1081,1146,1976,2002]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D273DE8FBA7D0191" box="[1081,1146,1976,2002]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC06D71D34DDE8FBB1A0191" box="[1287,1309,1976,2002]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="41" pageNumber="292" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC06D71D70DDEEBBFA601B5" box="[327,417,2012,2038]" pageId="41" pageNumber="292">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFC36D72D68DD9A0BAD606B4" blockId="42.[151,1437,151,632]" pageId="42" pageNumber="293">
<typeStatus id="F64E8839FFC36D72D68DD9A0BF3F06F2" box="[199,312,151,177]" pageId="42" pageNumber="293" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFC36D72D702D9A0BF8B06F2" box="[328,396,151,177]" count="3" pageId="42" pageNumber="293" type="female">3 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFC36D72D7D5D9A0BC3806F2" box="[415,575,151,177]" count="1" pageId="42" pageNumber="293" type="generic">one specimen</specimenCount>
dissected;
<specimenCount id="3FF3FD12FFC36D72D48FD9A0BD6406F2" box="[709,867,151,177]" count="1" pageId="42" pageNumber="293" type="generic">one specimen</specimenCount>
without right antennomeres 6–11); underside of the card of
<specimenCount id="3FF3FD12FFC36D72D751D98DBFC50690" box="[283,450,185,212]" count="2" pageId="42" pageNumber="293" type="generic">two specimens</specimenCount>
with handwritten ‘1/64’): same data as the holotype, with additional red printed label: ‘
<typeStatus id="F64E8839FFC36D72D6D1D9EABF2C06B4" box="[155,299,221,247]" pageId="42" pageNumber="293" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC36D72D775D9EBBFFD06B5" authorityName="Steel" authorityYear="1960" box="[319,506,220,246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC36D72D775D9EBBFFD06B5" box="[319,506,220,246]" italics="true" pageId="42" pageNumber="293">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC36D72D447D9EBBC5606B5" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[525,593,220,246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="293" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
<emphasis id="1B81EA89FFC36D72D447D9EBBC5606B5" box="[525,593,220,246]" italics="true" pageId="42" pageNumber="293">rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC36D72D413D9EABC9F06B4" bold="true" box="[601,664,221,247]" pageId="42" pageNumber="293">
<taxonomicNameLabel id="00B257F2FFC36D72D413D9EABC9F06B4" box="[601,664,221,247]" pageId="42" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC36D72D568D9EABD3F06B4" box="[802,824,221,247]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="42" pageNumber="293" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC36D72D221D9EABAC206B4" box="[1131,1221,221,247]" pageId="42" pageNumber="293">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC36D73D68DD9C8BF0202AA" lastPageId="43" lastPageNumber="294" pageId="42" pageNumber="293" type="description">
<paragraph id="294A369BFFC36D72D68DD9C8BB6E0723" blockId="42.[151,1437,151,632]" pageId="42" pageNumber="293">
<emphasis id="1B81EA89FFC36D72D68DD9C8BF50075A" bold="true" box="[199,343,255,281]" pageId="42" pageNumber="293">Description.</emphasis>
Measurements (n=4): HW: 0.45–0.46; HL: 0.23–0.24; OL: 0.15; TL: 0.04; AL (
<typeStatus id="F64E8839FFC36D72D2F4D9C8BB21075A" box="[1214,1318,255,281]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
): 0.61; PL: 0.37–0.41; PWmax: 0.55–0.61; PWmin: 0.48–0.53; ESL: 0.63–0.76; EW: 0.67–0.77; MTbL (
<typeStatus id="F64E8839FFC36D72D2E8D815BB0D077F" box="[1186,1290,290,316]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
): 0.35; MTrL (
<typeStatus id="F64E8839FFC36D72D6D5D872BF00071C" box="[159,263,325,351]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
): 0.18 (MTrL 1–4: 0.07; MTrL 5: 0.11); AW: 0.72–0.78; AedL: 0.55; BL: 2.00–2.32 (
<typeStatus id="F64E8839FFC36D72D2FDD872BB19071C" box="[1207,1310,325,351]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
: 2.15).
</paragraph>
<paragraph id="294A369BFFC36D72D68DD85FBF90043B" blockId="42.[151,1437,151,632]" pageId="42" pageNumber="293">
Habitus as in
<figureCitation id="B1CE2A1EFFC36D72D716D85FBFAB07C1" box="[348,428,360,386]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="42" pageNumber="293">Fig. 80</figureCitation>
. Head, pronotum and abdomen reddish brown; elytra yellow-brown, with slightly darkened latero-apical and apical portion of pronotum; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellowish. Middle portion of head with very fine and sparse punctation; neck without visible punctures; pronotum with fine and sparse punctation, slightly larger than that on middle portion of head, sparser in middle and indistinct in mediobasal and mediolateral portions; elytra with moderately sparse punctation about as that on pronotum, finer and sparser in parascutellar portion and along suture. Pronotum with dense and isodiametric microsculpture, distinctly coarser than that in middle portion of head; microsculpture of elytra dense, slightly larger and less coarser than that on pronotum.
</paragraph>
<caption id="7D8A6613FFC36D72D6DDDEACBD1B01BA" ID-DOI="http://doi.org/10.5281/zenodo.10821277" ID-Zenodo-Dep="10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="42" pageNumber="293" startId="42.[151,264,1947,1971]" targetBox="[159,1428,660,1914]" targetPageId="42" targetType="figure">
<paragraph id="294A369BFFC36D72D6DDDEACBD1B01BA" blockId="42.[151,1436,1947,2042]" pageId="42" pageNumber="293">
<emphasis id="1B81EA89FFC36D72D6DDDEACBF6B01F0" bold="true" box="[151,364,1947,1971]" pageId="42" pageNumber="293">FIGURES 126–132.</emphasis>
<taxonomicName id="EEF54D18FFC36D72D73ADEACBC5B01F0" authorityName="Shavrin" authorityYear="2024" box="[368,604,1947,1971]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="293" phylum="Arthropoda" rank="species" species="rufula">
<emphasis id="1B81EA89FFC36D72D73ADEACBC5B01F0" box="[368,604,1947,1971]" italics="true" pageId="42" pageNumber="293">Paraphloeostiba rufula</emphasis>
</taxonomicName>
: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFC26D73D68DD9A0BB55075E" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">Head 1.9 times as broad as long, with short and deep anteocellar foveae as long as diameter of ocellus. Ocelli large, located slightly below level of postocular carinae; distance between ocelli about as long as distance between ocellus and posterior margin of eyes. Antennomeres 7–10 distinctly transverse; antennomere 5 indisticntly broader than 4, 6 slightly longer and distinctly broader than 5, 7 slightly shorter than 6, 8–10 slightly broader than 7.</paragraph>
<paragraph id="294A369BFFC26D73D68DD810BA3807CA" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
Pronotum 1.4 times as broad as long, 1.2–1.3 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; middle portion without impressions, but
<specimenCount id="3FF3FD12FFC26D73D53ED87CBA160726" box="[884,1041,331,357]" count="2" pageId="43" pageNumber="294" type="generic" typeStatus="paratype">two paratypes</specimenCount>
with traces of very indistinct wide impressions in mediobasal third; lateral and laterobasal portions slightly impressed.
</paragraph>
<paragraph id="294A369BFFC26D73D68DD8A4BC9C07EE" blockId="43.[151,1437,151,1437]" box="[199,667,403,429]" pageId="43" pageNumber="294">Elytra 1.7–1.8 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFC26D73D68DD880BECF04C6" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC26D73D521D880BDC80792" box="[875,975,439,465]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 128</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFC26D73D70ED8ECBFAB07B6" box="[324,428,475,501]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 129</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward small rounded apex; parameres reaching apex of median lobe, distinctly broadened apically, with three moderately long apical and fourfive preapical setae, median parts of each paramere with three moderately long setae; internal sac moderately short and narrow, with elongate sclerotized structures in basal part (
<figureCitation id="B1CE2A1EFFC26D73D510DB70BDB90422" box="[858,958,583,609]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 126</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC26D73D33ADB70BEC304C6" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 127</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC26D73D68DDBB8BC9104B2" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC26D73D53FDBB8BDDF04EA" box="[885,984,655,681]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 130</figureCitation>
). Posterior margin of abdominal sternite VIII rounded (
<figureCitation id="B1CE2A1EFFC26D73D776DB84BF98048E" box="[316,415,691,717]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 131</figureCitation>
). Accessory sclerite short, from widest apical portion gradually narrowed toward rounded apex (
<figureCitation id="B1CE2A1EFFC26D73D691DBE0BF3904B2" box="[219,318,727,753]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 132</figureCitation>
). Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFC26D73D68DDBCDBA1E051E" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D68DDBCDBFB60556" bold="true" box="[199,433,762,789]" pageId="43" pageNumber="294">Comparative notes.</emphasis>
Based on the very broad median lobe and apical portions of the parameres,
<taxonomicName id="EEF54D18FFC26D73D345DBCCBB740556" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1295,1395,763,789]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFC26D73D345DBCCBB270557" box="[1295,1312,763,788]" italics="true" pageId="43" pageNumber="294">P</emphasis>
.
<emphasis id="1B81EA89FFC26D73D365DBCCBB740556" box="[1327,1395,763,789]" italics="true" pageId="43" pageNumber="294">rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D331DBCCBEA9057A" bold="true" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D331DBCCBEA9057A" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFC26D73D774DA28BFB2057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[318,437,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFC26D73D774DA28BFB2057A" box="[318,437,799,825]" italics="true" pageId="43" pageNumber="294">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D7F1DA28BFFD057A" bold="true" box="[443,506,799,825]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D7F1DA28BFFD057A" box="[443,506,799,825]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFC26D73D44DDA28BD1B057A" authority="Steel, 1960" authorityName="Steel" authorityYear="1960" box="[519,796,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFC26D73D44DDA28BC92057A" box="[519,661,799,825]" italics="true" pageId="43" pageNumber="294">P. celebensis</emphasis>
<bibRefCitation id="4D644B6AFFC26D73D4D1DA28BD1B057A" author="Steel, W. O." box="[667,796,799,825]" pageId="43" pageNumber="294" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel, 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFC26D73D56CDA28BDA4057B" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[806,931,799,824]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFC26D73D56CDA28BDA4057B" box="[806,931,799,824]" italics="true" pageId="43" pageNumber="294">P. papuana</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFC26D73D5FBDA28BA4B057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[945,1100,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFC26D73D5FBDA28BA4B057A" box="[945,1100,799,825]" italics="true" pageId="43" pageNumber="294">P. penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D218DA28BA96057A" bold="true" box="[1106,1169,799,825]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D218DA28BA96057A" box="[1106,1169,799,825]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFC26D73D28FDA28BB57057A" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[1221,1360,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFC26D73D28FDA28BB57057A" box="[1221,1360,799,825]" italics="true" pageId="43" pageNumber="294">P. tonkinesis</emphasis>
</taxonomicName>
. It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFC26D73D68DDA50BD5F05E6" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
from
<taxonomicName id="EEF54D18FFC26D73D74FDA50BF7B05C2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,871,897]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFC26D73D74FDA50BF7B05C2" box="[261,380,871,897]" italics="true" pageId="43" pageNumber="294">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D7C9DA50BFC405C2" bold="true" box="[387,451,871,897]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D7C9DA50BFC405C2" box="[387,451,871,897]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the larger body, darker head, pronotum and abdomen, the presence of the punctation on the head, the narrower pronotum and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFC26D73D68DDA98BA7F058A" blockId="43.[151,1437,151,1437]" box="[199,1144,943,969]" pageId="43" pageNumber="294">
from
<taxonomicName id="EEF54D18FFC26D73D74FDA98BF94058A" authorityName="Steel" authorityYear="1960" box="[261,403,943,969]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFC26D73D74FDA98BF94058A" box="[261,403,943,969]" italics="true" pageId="43" pageNumber="294">P. celebensis</emphasis>
</taxonomicName>
by the larger body and narrower apical portions of the parameres;
</paragraph>
<paragraph id="294A369BFFC26D73D68DDAE4BFB60252" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
from
<taxonomicName id="EEF54D18FFC26D73D74DDAE4BF8005AF" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[263,391,979,1004]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFC26D73D74DDAE4BF8005AF" box="[263,391,979,1004]" italics="true" pageId="43" pageNumber="294">P. papuana</emphasis>
</taxonomicName>
by the larger and darker body, the narrower pronotum, and slightly longer parameres with narrower apical portions;
</paragraph>
<paragraph id="294A369BFFC26D73D68DDD2CBAA9021A" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
from
<taxonomicName id="EEF54D18FFC26D73D74DDD2CBFA10276" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[263,422,1051,1077]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFC26D73D74DDD2CBFA10276" box="[263,422,1051,1077]" italics="true" pageId="43" pageNumber="294">P. penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D7E5DD2CBFF60276" bold="true" box="[431,497,1051,1077]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D7E5DD2CBFF60276" box="[431,497,1051,1077]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker body, the presence of the punctation on the head and the narrower pronotum, the longer elytra, smaller apical part of the median lobe and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFC26D73D68DDD54BAB102E2" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
from
<taxonomicName id="EEF54D18FFC26D73D74EDD54BF88023E" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[260,399,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFC26D73D74EDD54BF88023E" box="[260,399,1123,1149]" italics="true" pageId="43" pageNumber="294">P. tonkinesis</emphasis>
</taxonomicName>
by the distinctly larger body, the absence of the longitudinal impressions in the middle of the narrower pronotum, the narrower apical portion of the median lobe and the shorter parameres.
</paragraph>
<paragraph id="294A369BFFC26D73D68DDD9CBF0202AA" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
From all these species
<taxonomicName id="EEF54D18FFC26D73D791DD9CBC3A0286" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[475,573,1195,1221]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">P. rufula</taxonomicName>
<emphasis id="1B81EA89FFC26D73D403DD9CBC890286" bold="true" box="[585,654,1195,1221]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D403DD9CBC890286" box="[585,654,1195,1221]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs by the details of the external and internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC26D73D68DDDC5BEF70372" pageId="43" pageNumber="294" type="distribution">
<paragraph id="294A369BFFC26D73D68DDDC5BEF70372" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D68DDDC5BF66034F" bold="true" box="[199,353,1266,1292]" pageId="43" pageNumber="294">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFC26D73D721DDC4BC73034E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[363,628,1267,1293]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
<emphasis id="1B81EA89FFC26D73D721DDC4BC73034E" box="[363,628,1267,1293]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D4CADDC4BCC4034E" bold="true" box="[640,707,1267,1293]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D4CADDC4BCC4034E" box="[640,707,1267,1293]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from the
<typeStatus id="F64E8839FFC26D73D5E5DDC4BDD8034E" box="[943,991,1267,1293]" pageId="43" pageNumber="294">type</typeStatus>
locality in central part of
<collectingCountry id="51E2760BFFC26D73D352DDC4BEEB0372" name="Papua New Guinea" pageId="43" pageNumber="294">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC26D73D68DDC0CBD6D03DE" pageId="43" pageNumber="294" type="etymology">
<paragraph id="294A369BFFC26D73D68DDC0CBEF9033A" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D68DDC0CBF490316" bold="true" box="[199,334,1339,1365]" pageId="43" pageNumber="294">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFC26D73D48DDC0CBD110316" box="[711,790,1339,1365]" italics="true" pageId="43" pageNumber="294">rufulus</emphasis>
, -
<emphasis id="1B81EA89FFC26D73D57ADC0CBD390317" box="[816,830,1339,1364]" italics="true" pageId="43" pageNumber="294">a</emphasis>
, -
<emphasis id="1B81EA89FFC26D73D51CDC0CBD7F0317" box="[854,888,1339,1364]" italics="true" pageId="43" pageNumber="294">um</emphasis>
(reddish). It alludes to the general coloration of the body.
</paragraph>
<paragraph id="294A369BFFC26D73D68DDCB5BD6D03DE" blockId="43.[151,1437,151,1437]" box="[199,874,1410,1437]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D68DDCB5BF4C03DF" bold="true" box="[199,331,1410,1436]" pageId="43" pageNumber="294">Bionomics.</emphasis>
All specimens were collected flying in daylight.
</paragraph>
</subSubSection>
</treatment>
</document>

295
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<paragraph id="294A369BFFC16D70D6DDD9A0BFFD06F2" blockId="40.[151,506,151,214]" box="[151,506,151,178]" pageId="40" pageNumber="291">
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<taxonomicName id="EEF54D18FFC16D70D6DDD9A0BFB606F1" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,433,151,178]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC16D70D6DDD9A0BFB606F1" bold="true" box="[151,433,151,178]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba riedeli</emphasis>
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<taxonomicNameLabel id="00B257F2FFC16D70D7F2D9A0BFFD06F2" box="[440,506,151,177]" pageId="40" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="61EF6510FFC16D70D6DDD98CBF6B0695" box="[151,364,187,214]" pageId="40" pageNumber="291" type="description">
<paragraph id="294A369BFFC16D70D6DDD98CBF6B0695" blockId="40.[151,506,151,214]" box="[151,364,187,214]" pageId="40" pageNumber="291">
(
<figureCitation id="B1CE2A1EFFC16D70D6D5D98CBEF10696" box="[159,246,187,213]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="40" pageNumber="291">Figs 78</figureCitation>
,
<figureCitation id="B1CE2A1EFFC16D70D748D98CBF630695" box="[258,356,187,214]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="40" pageNumber="291">119–125</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC16D70D6DDD834BEE707B6" pageId="40" pageNumber="291" type="materials_examined">
<paragraph id="294A369BFFC16D70D6DDD834BB480702" blockId="40.[151,1437,259,501]" pageId="40" pageNumber="291">
<emphasis id="1B81EA89FFC16D70D6DDD834BF4E075E" bold="true" box="[151,329,259,285]" pageId="40" pageNumber="291">Type material.</emphasis>
<typeStatus id="F64E8839FFC16D70D719D834BFBC075E" box="[339,443,259,285]" pageId="40" pageNumber="291" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFC16D70D78CD833BFD9075D" box="[454,478,260,286]" count="1" pageId="40" pageNumber="291" type="male">♂</specimenCount>
: ‘
<collectingRegion id="EB31F879FFC16D70D7BBD834BC91075E" box="[497,662,259,285]" country="Indonesia" name="Papua Barat" pageId="40" pageNumber="291">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Bilogay,
<date id="5D4B105BFFC16D70D5FBD834BA28075D" box="[945,1071,259,286]" pageId="40" pageNumber="291" value="1995-12-29">29.12.1995</date>
|
<quantity id="EE0D9B7EFFC16D70D200D834BADE075E" box="[1098,1241,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="40" pageNumber="291" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC16D70D752D810BFB60702" box="[280,433,295,321]" pageId="40" pageNumber="291" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC16D70D78FD810BC870702" authorityName="Steel" authorityYear="1960" box="[453,640,295,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC16D70D78FD810BC870702" box="[453,640,295,321]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC16D70D4D9D810BCDB0702" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[659,732,295,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC16D70D4D9D810BCDB0702" box="[659,732,295,321]" italics="true" pageId="40" pageNumber="291">riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC16D70D4A9D810BD240702" bold="true" box="[739,803,295,321]" pageId="40" pageNumber="291">
<taxonomicNameLabel id="00B257F2FFC16D70D4A9D810BD240702" box="[739,803,295,321]" pageId="40" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC16D70D5E6D810BDC50702" box="[940,962,295,321]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="40" pageNumber="291" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC16D70D2BFD810BB440702" box="[1269,1347,295,321]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="291">NMW</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFC16D70D68DD87BBEE707B6" blockId="40.[151,1437,259,501]" pageId="40" pageNumber="291">
<typeStatus id="F64E8839FFC16D70D68DD87BBF3F0725" box="[199,312,332,358]" pageId="40" pageNumber="291" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFC16D70D70FD87CBF930725" box="[325,404,331,358]" count="10" pageId="40" pageNumber="291" type="male">10 ♂♂</specimenCount>
(
<specimenCount id="3FF3FD12FFC16D70D7EFD87CBC480726" box="[421,591,331,357]" count="2" pageId="40" pageNumber="291" type="generic">two specimens</specimenCount>
dissected;
<specimenCount id="3FF3FD12FFC16D70D485D87BBD6C0726" box="[719,875,331,358]" count="1" pageId="40" pageNumber="291" type="generic">one specimen</specimenCount>
without apical part of the abdomen),
<specimenCount id="3FF3FD12FFC16D70D355D87CBB670725" box="[1311,1376,331,358]" count="5" pageId="40" pageNumber="291" type="female">5 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFC16D70D33BD87CBF0B07CA" count="2" pageId="40" pageNumber="291" type="generic">two specimens</specimenCount>
dissected): same data as the holotype (
<specimenCount id="3FF3FD12FFC16D70D486D858BD0807C9" box="[716,783,367,394]" count="2" pageId="40" pageNumber="291" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFC16D70D557D858BD4D07C9" box="[797,842,367,394]" count="1" pageId="40" pageNumber="291" type="female">1 ♀</specimenCount>
: cSh;
<specimenCount id="3FF3FD12FFC16D70D5D9D847BDD107C9" box="[915,982,368,394]" count="7" pageId="40" pageNumber="291" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFC16D70D5A9D858BA2107C9" box="[995,1062,367,394]" count="4" pageId="40" pageNumber="291" type="female">4 ♀♀</specimenCount>
:
<collectionCode id="4FE4AE5EFFC16D70D27ED858BA8607CA" box="[1076,1153,367,393]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="291">NMW</collectionCode>
);
<specimenCount id="3FF3FD12FFC16D70D2DFD858BAC607C9" box="[1173,1217,367,394]" count="1" pageId="40" pageNumber="291" type="male">1 ♂</specimenCount>
(left antennomeres 6–11 missing): ‘
<collectingRegion id="EB31F879FFC16D70D70DD8A4BFEF07EE" box="[327,488,403,429]" country="Indonesia" name="Papua Barat" pageId="40" pageNumber="291">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Bilogay,
<quantity id="EE0D9B7EFFC16D70D4A4D8A4BD7A07EE" box="[750,893,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="40" pageNumber="291" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
|
<date id="5D4B105BFFC16D70D5C5D8A4BA0A07ED" box="[911,1037,403,430]" pageId="40" pageNumber="291" value="1995-12-29">29.12.1995</date>
| leg. A. Riedel’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFC16D70D309D8A4BB9607EE" box="[1347,1425,403,429]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="291">NMW</collectionCode>
). All
<typeStatus id="F64E8839FFC16D70D689D88FBF290791" box="[195,302,440,466]" pageId="40" pageNumber="291" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFC16D70D4FDD880BD400792" box="[695,839,439,465]" pageId="40" pageNumber="291" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC16D70D514D880BA1E0792" authorityName="Steel" authorityYear="1960" box="[862,1049,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC16D70D514D880BA1E0792" box="[862,1049,439,465]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC16D70D265D880BA7F0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1071,1144,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC16D70D265D880BA7F0792" box="[1071,1144,439,465]" italics="true" pageId="40" pageNumber="291">riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC16D70D2CAD880BAC60792" bold="true" box="[1152,1217,439,465]" pageId="40" pageNumber="291">
<taxonomicNameLabel id="00B257F2FFC16D70D2CAD880BAC60792" box="[1152,1217,439,465]" pageId="40" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC16D70D304D880BB630792" box="[1358,1380,439,465]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="40" pageNumber="291" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<caption id="7D8A6613FFC16D70D6DDDE63BDDE01F7" ID-DOI="http://doi.org/10.5281/zenodo.10821275" ID-Zenodo-Dep="10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="40" pageNumber="291" startId="40.[151,264,1876,1900]" targetBox="[159,1428,533,1844]" targetPageId="40" targetType="figure">
<paragraph id="294A369BFFC16D70D6DDDE63BDDE01F7" blockId="40.[151,1437,1876,1973]" pageId="40" pageNumber="291">
<emphasis id="1B81EA89FFC16D70D6DDDE63BF77012F" bold="true" box="[151,368,1876,1900]" pageId="40" pageNumber="291">FIGURES 119–125.</emphasis>
<taxonomicName id="EEF54D18FFC16D70D733DE63BC69012F" authorityName="Shavrin" authorityYear="2024" box="[377,622,1876,1900]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="1B81EA89FFC16D70D733DE63BC69012F" box="[377,622,1876,1900]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba riedeli</emphasis>
</taxonomicName>
(paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="61EF6510FFC06D71D68DD9A1BC60006E" pageId="41" pageNumber="292" type="description">
<paragraph id="294A369BFFC06D71D68DD9A1BEDE075E" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DD9A1BF5E06F3" bold="true" box="[199,345,150,176]" pageId="41" pageNumber="292">Description.</emphasis>
Measurements (n=17): HW: 0.42–0.56; HL: 0.25–0.26; OL: 0.14; TL: 0.02; AL (
<typeStatus id="F64E8839FFC06D71D2A2D9A0BB5606F2" box="[1256,1361,151,177]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
): 0.62; PL: 0.29–0.34; PWmax: 0.54–0.56; PWmin: 0.46–0.47; ESL: 0.55–0.60; EW: 0.70–0.76; MTbL (
<typeStatus id="F64E8839FFC06D71D2ADD98CBB570696" box="[1255,1360,187,213]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFC06D71D6A1D9E8BF5306BA" box="[235,340,223,249]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
): 0.15 (MTrL 1–4: 0.05; MTrL 5: 0.10); AW: 0.61–0.79; AedL: 0.42; BL: 1.80–2.40 (
<typeStatus id="F64E8839FFC06D71D37BD9E8BB9E06BA" box="[1329,1433,223,249]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
: 1.87).
</paragraph>
<paragraph id="294A369BFFC06D71D68DD810BD720422" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
Habitus as in
<figureCitation id="B1CE2A1EFFC06D71D722D810BFBA0702" box="[360,445,295,321]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="41" pageNumber="292">Fig. 78</figureCitation>
. Body yellowish brown to reddish-brown, with slightly paler elytra (several
<typeStatus id="F64E8839FFC06D71D37BD810BB9B0702" box="[1329,1436,295,321]" pageId="41" pageNumber="292" type="paratype">paratypes</typeStatus>
with paler lateral and basal portion of pronotum); antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow (some
<typeStatus id="F64E8839FFC06D71D7E9D858BC0907CA" box="[419,526,367,393]" pageId="41" pageNumber="292" type="paratype">paratypes</typeStatus>
with distinctly darker preapical maxillary palpomere). Pronotum with fine and sparse punctation, slightly larger in lateral and finer and sparser in middle portions (some
<typeStatus id="F64E8839FFC06D71D2C9D8A4BAE907EE" box="[1155,1262,403,429]" pageId="41" pageNumber="292" type="paratype">paratypes</typeStatus>
without visible punctation in middle); elytra with moderately dense punctation, distinctly larger and deeper than that on pronotum, finer and sparser in middle; abdominal tergites with distinct, moderately fine and dense punctation. Middle portion of head with dense and coarse isodiametric microsculpture; pronotum with dense microreticulation somewhat similar to that in middle portion of head, finer in middle; microsculpture of elytra about as that in pronotum, slightly finer in middle; abdominal tergites with dense transverse meshes.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDB5CBFB0057A" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">Head 1.6 times to twice as broad as long, with narrow and moderately deep linear anteocellar foveae; postocular carinae acute. Ocelli large, located at level of postocular carinae; distance between ocelli 1.6–1.8 times as long as distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about 1.6 times as long as preapical segment. Antennomere 4 indistinctly longer than broad, twice shorter than 3, 5 slightly broader than 4, 6 slightly longer and about twice broader than 5, 7 slightly longer and broader than 5, 8–9 slightly shorter than 7, 10 indisitnctly longer than 9.</paragraph>
<paragraph id="294A369BFFC06D71D68DDA74BC7405E6" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">Pronotum 1.6–1.8 times as broad as long, 1.1–1.2 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; middle portion with indistinct wide longitudinal impressions; laterobasal impressions moderately wide and shallow.</paragraph>
<paragraph id="294A369BFFC06D71D68DDA98BDCB058A" blockId="41.[151,1437,150,1761]" box="[199,972,943,969]" pageId="41" pageNumber="292">Elytra 1.2 times as broad as long, 1.7–1.8 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFC06D71D68DDAE4BCFA05AE" blockId="41.[151,1437,150,1761]" box="[199,765,979,1005]" pageId="41" pageNumber="292">Abdomen slightly narrower or broader than elytra.</paragraph>
<paragraph id="294A369BFFC06D71D68DDAC0BD630286" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC06D71D506DAC0BDAB0252" box="[844,940,1015,1041]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 121</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFC06D71D749DD2CBF610276" box="[259,358,1051,1077]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 122</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward small rounded apex; parameres slightly not reaching apex of median lobe, slightly broadened in preapical and narrowed in apical portions, each with three apical and three preapical moderately short setae, inner margins of each paramere in middle with several moderately short setae; internal sac long and moderately narrow, with two elongate sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFC06D71D6BDDD9CBF5E0286" box="[247,345,1195,1221]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 119</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC06D71D54ADD9CBD670286" box="[768,864,1195,1221]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 120</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDDF8BD3B0372" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
Female. Posterior margin of abdominal tergite VIII rounded (
<figureCitation id="B1CE2A1EFFC06D71D53CDDF8BDDF02AA" box="[886,984,1231,1257]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 123</figureCitation>
). Posterior margin of abdominal sternite VIII truncate (
<figureCitation id="B1CE2A1EFFC06D71D770DDC4BF9A034E" box="[314,413,1267,1293]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 124</figureCitation>
). Accessory sclerite triangular, with wide basal portion, gradually narrowed apicad toward small rounded apex (
<figureCitation id="B1CE2A1EFFC06D71D7C8DC20BFE30372" box="[386,484,1302,1329]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 125</figureCitation>
). Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDC0DBC60006E" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDC0DBFA80316" bold="true" box="[199,431,1338,1365]" pageId="41" pageNumber="292">Comparative notes.</emphasis>
Based on the general structure of the aedeagus,
<taxonomicName id="EEF54D18FFC06D71D5F4DC0CBA260316" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[958,1057,1339,1365]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D5F4DC0CBA260316" box="[958,1057,1339,1365]" italics="true" pageId="41" pageNumber="292">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D26CDC0CBA630316" bold="true" box="[1062,1124,1339,1365]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D26CDC0CBA630316" box="[1062,1124,1339,1365]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFC06D71D2A5DC0CBB700317" authorityName="Shavrin" authorityYear="2023" box="[1263,1399,1339,1364]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFC06D71D2A5DC0CBB700317" box="[1263,1399,1339,1364]" italics="true" pageId="41" pageNumber="292">P. conjuncta</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFC06D71D3CEDC0CBF5C033A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D3CEDC0CBB920317" box="[1412,1429,1339,1364]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D6DDDC68BF5C033A" box="[151,347,1375,1401]" italics="true" pageId="41" pageNumber="292">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D728DC68BFA5033A" bold="true" box="[354,418,1375,1401]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D728DC68BFA5033A" box="[354,418,1375,1401]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="EEF54D18FFC06D71D795DC68BC45033B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[479,578,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D795DC68BFF7033B" box="[479,496,1375,1400]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D7B4DC68BC45033B" box="[510,578,1375,1400]" italics="true" pageId="41" pageNumber="292">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D403DC68BC8E033A" bold="true" box="[585,649,1375,1401]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D403DC68BC8E033A" box="[585,649,1375,1401]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from
<taxonomicName id="EEF54D18FFC06D71D59ADC68BA5D033B" authorityName="Shavrin" authorityYear="2023" box="[976,1114,1375,1400]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFC06D71D59ADC68BA5D033B" box="[976,1114,1375,1400]" italics="true" pageId="41" pageNumber="292">P. conjuncta</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF54D18FFC06D71D2DBDC68BB73033A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1169,1396,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D2DBDC68BAA5033B" box="[1169,1186,1375,1400]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D2FADC68BB73033A" box="[1200,1396,1375,1401]" italics="true" pageId="41" pageNumber="292">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D331DC68BEA903DE" bold="true" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D331DC68BEA903DE" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the less distinct longitudinal impressions in the middle of the pronotum. From
<taxonomicName id="EEF54D18FFC06D71D209DCB4BAA203DF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1091,1189,1411,1437]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D209DCB4BA5303DF" box="[1091,1108,1411,1436]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D22BDCB4BAA203DF" box="[1121,1189,1411,1436]" italics="true" pageId="41" pageNumber="292">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D2E7DCB4BAEB03DE" bold="true" box="[1197,1260,1411,1437]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D2E7DCB4BAEB03DE" box="[1197,1260,1411,1437]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
it differs by the slightly shorter parameres, the absence of the punctation of the head, and the less distinct longitudinal impressions in the middle of the pronotum. Additionally, it differs from
<taxonomicName id="EEF54D18FFC06D71D545DCFCBD9F03A7" authorityName="Shavrin" authorityYear="2023" box="[783,920,1483,1508]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFC06D71D545DCFCBD9F03A7" box="[783,920,1483,1508]" italics="true" pageId="41" pageNumber="292">P. conjuncta</emphasis>
</taxonomicName>
by the pronotum with not concave laterobasal margins. From all these species it differs by the narrower median lobe, the different structure of the internal sac and the shape of the female accessory sclerite
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC06D71D68DDF01BB930012" box="[199,1428,1590,1617]" pageId="41" pageNumber="292" type="distribution">
<paragraph id="294A369BFFC06D71D68DDF01BB930012" blockId="41.[151,1437,150,1761]" box="[199,1428,1590,1617]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDF01BF660013" bold="true" box="[199,353,1590,1616]" pageId="41" pageNumber="292">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFC06D71D72DDF00BC750012" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[359,626,1591,1617]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC06D71D72DDF00BC750012" box="[359,626,1591,1617]" italics="true" pageId="41" pageNumber="292">Paraphloeostiba riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D433DF00BCBE0012" bold="true" box="[633,697,1591,1617]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D433DF00BCBE0012" box="[633,697,1591,1617]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in Western New
<collectingCountry id="51E2760BFFC06D71D28BDF00BB120012" box="[1217,1301,1591,1617]" name="Guinea-Bissau" pageId="41" pageNumber="292">Guinea</collectingCountry>
,
<collectingCountry id="51E2760BFFC06D71D36ADF00BB960012" box="[1312,1425,1591,1617]" name="Indonesia" pageId="41" pageNumber="292">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC06D71D68DDF6CBF3700A2" pageId="41" pageNumber="292" type="etymology">
<paragraph id="294A369BFFC06D71D68DDF6CBF5C00DA" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDF6CBF490036" bold="true" box="[199,334,1627,1653]" pageId="41" pageNumber="292">Etymology.</emphasis>
Patronymic, the species is named after coleopterologist Alexander Riedel (Karlsruhe), collector of the
<typeStatus id="F64E8839FFC06D71D68ADF48BEF700DA" box="[192,240,1663,1689]" pageId="41" pageNumber="292">type</typeStatus>
material.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDF95BF3700A2" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDF95BF4C00FF" bold="true" box="[199,331,1698,1724]" pageId="41" pageNumber="292">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="EE0D9B7EFFC06D71D512DF94BA0A00FE" box="[856,1037,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="41" pageNumber="292" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
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314
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<mods:title id="7B157EA58B6498EAECD0165311915251">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<paragraph id="294A369BFFC26D73D6DDDCD9BFEC004A" blockId="43.[151,491,1518,1581]" box="[151,491,1518,1545]" pageId="43" pageNumber="294">
<heading id="720281F7FFC26D73D6DDDCD9BFEC004A" bold="true" box="[151,491,1518,1545]" fontSize="11" level="1" pageId="43" pageNumber="294" reason="1">
<emphasis id="1B81EA89FFC26D73D6DDDCD9BFEC004A" bold="true" box="[151,491,1518,1545]" pageId="43" pageNumber="294">
<taxonomicName id="EEF54D18FFC26D73D6DDDCD9BFA5004A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,418,1518,1545]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC26D73D6DDDCD9BFA5004A" bold="true" box="[151,418,1518,1545]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba steeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFC26D73D7E3DCD8BFEC004A" box="[425,491,1519,1545]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="61EF6510FFC26D73D6DDDF24BF6A006E" box="[151,365,1555,1581]" pageId="43" pageNumber="294" type="description">
<paragraph id="294A369BFFC26D73D6DDDF24BF6A006E" blockId="43.[151,491,1518,1581]" box="[151,365,1555,1581]" pageId="43" pageNumber="294">
(
<figureCitation id="B1CE2A1EFFC26D73D6D5DF24BEF0006E" box="[159,247,1555,1581]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="43" pageNumber="294">Figs 44</figureCitation>
,
<figureCitation id="B1CE2A1EFFC26D73D748DF24BF61006E" box="[258,358,1555,1581]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="43" pageNumber="294">133–140</figureCitation>
)
</paragraph>
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<subSubSection id="61EF6510FFC26D73D6DDDF6CBA27016A" pageId="43" pageNumber="294" type="materials_examined">
<paragraph id="294A369BFFC26D73D6DDDF6CBAD600FE" blockId="43.[151,1437,1626,2013]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D6DDDF6CBF430037" bold="true" box="[151,324,1626,1653]" pageId="43" pageNumber="294">Type material.</emphasis>
<typeStatus id="F64E8839FFC26D73D700DF6CBFB50036" box="[330,434,1627,1653]" pageId="43" pageNumber="294" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFC26D73D7F3DF6CBFC90036" box="[441,462,1627,1653]" count="1" pageId="43" pageNumber="294" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFC26D73D796DF6CBC280036" box="[476,559,1627,1653]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="43" pageNumber="294">Fig. 44</figureCitation>
; dissected): ‘
<collectionCode id="4FE4AE5EFFC26D73D4F6DF6CBD070036" box="[700,768,1627,1653]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="43" pageNumber="294">NEW</collectionCode>
<collectingCountry id="51E2760BFFC26D73D54CDF6CBD770036" box="[774,880,1627,1653]" name="Guinea-Bissau" pageId="43" pageNumber="294">GUINEA</collectingCountry>
| Kamin- | dimbit, Main River, Se- | pik, Angoram sub-div., |
<date id="5D4B105BFFC26D73D741DF49BF8C00DA" box="[267,395,1662,1689]" pageId="43" pageNumber="294" value="1965-03-12">12.III.1965</date>
, running on | swamp plants in marsh’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFC26D73D5FFDF48BDC900DA" box="[949,974,1663,1689]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="294">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC26D73D6D1DF94BF3300FE" box="[155,308,1699,1725]" pageId="43" pageNumber="294" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC26D73D702DF94BC0400FE" authorityName="Steel" authorityYear="1960" box="[328,515,1699,1725]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC26D73D702DF94BC0400FE" box="[328,515,1699,1725]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC26D73D45CDF94BC5600FE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[534,593,1699,1725]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC26D73D45CDF94BC5600FE" box="[534,593,1699,1725]" italics="true" pageId="43" pageNumber="294">steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D412DF94BC9F00FE" bold="true" box="[600,664,1699,1725]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D412DF94BC9F00FE" box="[600,664,1699,1725]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC26D73D56BDF94BD3000FE" box="[801,823,1699,1725]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="43" pageNumber="294" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC26D73D220DF94BAC300FE" box="[1130,1220,1699,1725]" pageId="43" pageNumber="294">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFC26D73D68DDFF0BA27016A" blockId="43.[151,1437,1626,2013]" pageId="43" pageNumber="294">
<typeStatus id="F64E8839FFC26D73D68DDFF0BF3F00A2" box="[199,312,1735,1761]" pageId="43" pageNumber="294" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFC26D73D702DFF0BF7100A2" box="[328,374,1735,1761]" count="1" pageId="43" pageNumber="294" type="female">1 ♀</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFC26D73D459DFF0BC5000A2" box="[531,599,1735,1761]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="43" pageNumber="294">NEW</collectionCode>
<collectingCountry id="51E2760BFFC26D73D429DFF0BCCA00A2" box="[611,717,1735,1761]" name="Guinea-Bissau" pageId="43" pageNumber="294">GUINEA</collectingCountry>
| Kamindim- | bit, Main River, Sepik, Ango- | ram sub-div.,
<date id="5D4B105BFFC26D73D6DDDFDDBF1F0146" box="[151,280,1770,1797]" pageId="43" pageNumber="294" value="1965-03-12">12.III.1965</date>
, | running on swamp plants in | marsh’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFC26D73D5D8DFDCBDAC0146" box="[914,939,1771,1797]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="294">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC26D73D6D1DE38BF2C016A" box="[155,299,1807,1833]" pageId="43" pageNumber="294" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC26D73D775DE38BFFD016A" authorityName="Steel" authorityYear="1960" box="[319,506,1807,1833]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC26D73D775DE38BFFD016A" box="[319,506,1807,1833]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC26D73D447DE38BC4F016A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[525,584,1807,1833]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC26D73D447DE38BC4F016A" box="[525,584,1807,1833]" italics="true" pageId="43" pageNumber="294">steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D405DE38BC88016A" bold="true" box="[591,655,1807,1833]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D405DE38BC88016A" box="[591,655,1807,1833]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC26D73D552DE38BD29016A" box="[792,814,1807,1833]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="43" pageNumber="294" type="Museum">V</collectionCode>
. des. 2024’ (
<collectionCode id="4FE4AE5EFFC26D73D5F0DE38BA13016A" box="[954,1044,1807,1833]" pageId="43" pageNumber="294">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC26D75D68DDE05BF2805C2" lastPageId="45" lastPageNumber="296" pageId="43" pageNumber="294" type="description">
<paragraph id="294A369BFFC26D73D68DDE05BB8601D6" blockId="43.[151,1437,1626,2013]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D68DDE05BF5E010F" bold="true" box="[199,345,1842,1868]" pageId="43" pageNumber="294">Description.</emphasis>
Measurements (n=2): HW: 0.25–0.26; HL: 0.21–0.22; OL: 0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFC26D73D2ADDE04BB57010E" box="[1255,1360,1843,1869]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
): 0.49; PL: 0.32–0.34; PWmax: 0.55–0.56; PWmin: 0.50–0.52; ESL: 0.52–0.53; EW: 0.64–0.66; MTbL (
<typeStatus id="F64E8839FFC26D73D2ADDE60BB570132" box="[1255,1360,1879,1905]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFC26D73D6A2DE4CBF5601D6" box="[232,337,1915,1941]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
): 0.14 (MTrL 1–4: 0.05; MTrL 5: 0.09); AW: 0.60–0.62; AedL: 0.37; BL: 1.78–1.90 (
<typeStatus id="F64E8839FFC26D73D346DE4CBB7201D6" box="[1292,1397,1915,1941]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="294A369BFFC26D74D68DDEA8BCE0075E" blockId="43.[151,1437,1626,2013]" lastBlockId="44.[151,1437,151,645]" lastPageId="44" lastPageNumber="295" pageId="43" pageNumber="294">
Habitus as in
<figureCitation id="B1CE2A1EFFC26D73D716DEA8BFAA01FA" box="[348,429,1951,1977]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="43" pageNumber="294">Fig. 44</figureCitation>
. Body reddish-brown (
<typeStatus id="F64E8839FFC26D73D4E0DEA8BD0901FA" box="[682,782,1951,1977]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
with slightly darker abdomen); antennomeres 6–11 brown; mouthparts, antennomeres 1–4 and legs yellow. Head and neck without visible punctures; pronotum with very sparse and fine indistinct punctation, invisible in middle; elytra with dense punctation, distinctly larger and deeper than that on pronotum, slightly finer and sparser in middle; abdominal tergites without visible punctation. Head with very dense and coarse isodiametric microsculpture; microsculpture of pronotum about as that in middle portion of head; elytra with fine isodiametric microreticulation.
</paragraph>
<paragraph id="294A369BFFC56D74D68DD810BCB40792" blockId="44.[151,1437,151,645]" pageId="44" pageNumber="295">Head with moderately deep and wide anteriomedian depressions, slightly broader than long, with short and deep narrow anteocellar foveae about as long as diameter of ocellus; postocular carina absent. Distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Antennomeres 6–10 distinctly transverse; antennomere 5 slightly broader than 4, 6 distinctly broader than 5, 7 longer and broader than 6, 8–9 slightly shorter than 7, 10 slightly longer than 9.</paragraph>
<paragraph id="294A369BFFC56D74D68DD8ECBF99045A" blockId="44.[151,1437,151,645]" pageId="44" pageNumber="295">Pronotum 1.4–1.6 times as broad as long, twice as broad as head, widest in middle, distinctly more narrowed posteriad than anteriad.</paragraph>
<paragraph id="294A369BFFC56D74D68DDB14BC7B0422" blockId="44.[151,1437,151,645]" pageId="44" pageNumber="295">Elytra 1.2 times as broad as long, 1.5–1.6 times as long as pronotum; surface between punctures slightly and longitudinally elevated, stronger in middle.</paragraph>
<paragraph id="294A369BFFC56D74D68DDB5CBC8504C6" blockId="44.[151,1437,151,645]" box="[199,642,619,645]" pageId="44" pageNumber="295">Abdomen slightly narrower than elytra.</paragraph>
<caption id="7D8A6613FFC56D74D6DDDEA0BDDD01B4" ID-DOI="http://doi.org/10.5281/zenodo.10821281" ID-Zenodo-Dep="10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="44" pageNumber="295" startId="44.[151,264,1943,1967]" targetBox="[185,1403,678,1912]" targetPageId="44" targetType="figure">
<paragraph id="294A369BFFC56D74D6DDDEA0BDDD01B4" blockId="44.[151,1436,1943,2040]" pageId="44" pageNumber="295">
<emphasis id="1B81EA89FFC56D74D6DDDEA0BF6A01EC" bold="true" box="[151,365,1943,1967]" pageId="44" pageNumber="295">FIGURES 133–140.</emphasis>
<taxonomicName id="EEF54D18FFC56D74D738DEA0BC5201EC" authorityName="Shavrin" authorityYear="2024" box="[370,597,1943,1967]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="295" phylum="Arthropoda" rank="species" species="steeli">
<emphasis id="1B81EA89FFC56D74D738DEA0BC5201EC" box="[370,597,1943,1967]" italics="true" pageId="44" pageNumber="295">Paraphloeostiba steeli</emphasis>
</taxonomicName>
: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFC46D75D68DD9A0BF300726" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC46D75D521D9A0BDC806F2" box="[875,975,150,177]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 135</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFC46D75D45CD98CBC7D0696" box="[534,634,187,213]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 136</figureCitation>
). Aedeagus with wide basal portion, strongly narrowed toward narrow elongate median lobe with very small rounded apex; parameres slightly exceeding apex of median lobe, moderately wide, distinctly broadened apically, with widely rounded apices, with two moderately short apical and preapical setae; internal sac long and moderately wide, without sclerotized structures (
<figureCitation id="B1CE2A1EFFC46D75D595D810BA470702" box="[991,1088,295,321]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 133</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC46D75D698D87CBF340726" box="[210,307,331,357]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 134</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC46D75D68DD858BC8F0792" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC46D75D53FD858BDDF07CA" box="[885,984,367,393]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 137</figureCitation>
). Posterior margin of abdominal sternite VIII rounded (
<figureCitation id="B1CE2A1EFFC46D75D774D8A4BFA507EE" box="[318,418,403,429]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 138</figureCitation>
). Accessory sclerite short, from widest basal portion gradually narrowed toward rounded apex (
<figureCitation id="B1CE2A1EFFC46D75D691D880BF390792" box="[219,318,439,465]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 139</figureCitation>
). Spermatheca as in
<figureCitation id="B1CE2A1EFFC46D75D46ED880BC830792" box="[548,644,439,465]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 140</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC46D75D68DD8EDBCFC047E" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DD8EDBFB607B6" bold="true" box="[199,433,474,501]" pageId="45" pageNumber="296">Comparative notes.</emphasis>
Based on the very narrow median lobe and the parameres,
<taxonomicName id="EEF54D18FFC46D75D21BD8ECBAAF07B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1105,1192,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D21BD8ECBAAF07B6" box="[1105,1192,475,501]" italics="true" pageId="45" pageNumber="296">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D2E5D8ECBAF707B6" bold="true" box="[1199,1264,475,501]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D2E5D8ECBAF707B6" box="[1199,1264,475,501]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFC46D75D3CED8ECBF2C045B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D3CED8ECBB9207B7" box="[1412,1429,475,500]" italics="true" pageId="45" pageNumber="296">P</emphasis>
.
<emphasis id="1B81EA89FFC46D75D6DDD8C8BF2C045B" box="[151,299,511,536]" italics="true" pageId="45" pageNumber="296">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D778D8C8BF75045A" bold="true" box="[306,370,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D778D8C8BF75045A" box="[306,370,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFC46D75D7CAD8C8BFFF045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[384,504,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D7CAD8C8BFFF045A" box="[384,504,511,537]" italics="true" pageId="45" pageNumber="296">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D44AD8C8BC47045A" bold="true" box="[512,576,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D44AD8C8BC47045A" box="[512,576,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFC46D75D404D8C8BCDD045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[590,730,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D404D8C8BCDD045A" box="[590,730,511,537]" italics="true" pageId="45" pageNumber="296">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D4A8D8C8BD26045A" bold="true" box="[738,801,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D4A8D8C8BD26045A" box="[738,801,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFC46D75D512D8C8BDE9045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[856,1006,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D512D8C8BDE9045A" box="[856,1006,511,537]" italics="true" pageId="45" pageNumber="296">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D5BCD8C8BA31045A" bold="true" box="[1014,1078,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D5BCD8C8BA31045A" box="[1014,1078,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFC46D75D68DDB70BB0C0422" blockId="45.[151,1437,150,1077]" box="[199,1291,583,609]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D74FDB70BFBF0423" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,440,583,609]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D74FDB70BF110423" box="[261,278,583,608]" italics="true" pageId="45" pageNumber="296">P</emphasis>
.
<emphasis id="1B81EA89FFC46D75D76EDB70BFBF0423" box="[292,440,583,608]" italics="true" pageId="45" pageNumber="296">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7F4DB70BFF90422" bold="true" box="[446,510,583,609]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7F4DB70BFF90422" box="[446,510,583,609]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller and slightly paler body, and slightly longer parameres;
</paragraph>
<paragraph id="294A369BFFC46D75D68DDB5CBDD404C6" blockId="45.[151,1437,150,1077]" box="[199,979,619,645]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D74FDB5CBF7B04C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D74FDB5CBF7B04C6" box="[261,380,619,645]" italics="true" pageId="45" pageNumber="296">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7C9DB5CBFC404C6" bold="true" box="[387,451,619,645]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7C9DB5CBFC404C6" box="[387,451,619,645]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker elytra and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFC46D75D68DDBB8BD47048E" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D749DBB8BF8B04EA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[259,396,655,681]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D749DBB8BF8B04EA" box="[259,396,655,681]" italics="true" pageId="45" pageNumber="296">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7DBDBB8BFC904EA" bold="true" box="[401,462,655,681]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7DBDBB8BFC904EA" box="[401,462,655,681]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the larger body, the presence of the punctation on the head and the stronger punctation of the pronotum, the shorter elytra and the longer parameres;
</paragraph>
<paragraph id="294A369BFFC46D75D68DDBE0BDC50556" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D749DBE0BF9F04B2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[259,408,727,753]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D749DBE0BF9F04B2" box="[259,408,727,753]" italics="true" pageId="45" pageNumber="296">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7D7DBE0BFDB04B2" bold="true" box="[413,476,727,753]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7D7DBE0BFDB04B2" box="[413,476,727,753]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the slightly darker body, the presence of the punctation on the head and the stronger punctation on the pronotum, the shorter elytra and the longer parameres.
</paragraph>
<paragraph id="294A369BFFC46D75D68DDA28BF2805C2" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
From all these species,
<taxonomicName id="EEF54D18FFC46D75D786DA28BC21057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[460,550,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D786DA28BFDA057B" box="[460,477,799,824]" italics="true" pageId="45" pageNumber="296">P</emphasis>
.
<emphasis id="1B81EA89FFC46D75D7A1DA28BC21057A" box="[491,550,799,825]" italics="true" pageId="45" pageNumber="296">steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D467DA28BC6B057A" bold="true" box="[557,620,799,825]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D467DA28BC6B057A" box="[557,620,799,825]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the presence of elevations between punctures on the elytra, distinctly broader apical portions of the parameres and details of the external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC46D75D68DDABDBB4B05E6" box="[199,1356,906,933]" pageId="45" pageNumber="296" type="distribution">
<paragraph id="294A369BFFC46D75D68DDABDBB4B05E6" blockId="45.[151,1437,150,1077]" box="[199,1356,906,933]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DDABDBF6605E7" bold="true" box="[199,353,906,932]" pageId="45" pageNumber="296">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFC46D75D72DDABCBC6305E6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[359,612,907,933]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC46D75D72DDABCBC6305E6" box="[359,612,907,933]" italics="true" pageId="45" pageNumber="296">Paraphloeostiba steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D421DABCBCAC05E6" bold="true" box="[619,683,907,933]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D421DABCBCAC05E6" box="[619,683,907,933]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFC46D75D5F4DABCBDE905E6" box="[958,1006,907,933]" pageId="45" pageNumber="296">type</typeStatus>
locality in
<collectingCountry id="51E2760BFFC46D75D227DABCBB4E05E6" box="[1133,1353,907,933]" name="Papua New Guinea" pageId="45" pageNumber="296">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC46D75D68DDA98BF390276" pageId="45" pageNumber="296" type="etymology">
<paragraph id="294A369BFFC46D75D68DDA98BD2B05AE" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DDA98BF49058A" bold="true" box="[199,334,943,969]" pageId="45" pageNumber="296">Etymology.</emphasis>
The species is named in the memory of the English coleopterologist William Owen Steel (1917– 1969) for his contributions to the knowledge of
<taxonomicName id="EEF54D18FFC46D75D4FADAE4BD2F05AE" box="[688,808,979,1005]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
.
</paragraph>
<paragraph id="294A369BFFC46D75D68DDAC1BF390276" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DDAC1BF4C0253" bold="true" box="[199,331,1014,1040]" pageId="45" pageNumber="296">Bionomics.</emphasis>
<specimenCount id="3FF3FD12FFC46D75D705DAC0BFFD0252" box="[335,506,1015,1041]" count="2" pageId="45" pageNumber="296" type="generic">Two specimens</specimenCount>
were collected in swamp plants in marsh, and
<specimenCount id="3FF3FD12FFC46D75D5B9DAC0BA8D0252" box="[1011,1162,1015,1041]" count="1" pageId="45" pageNumber="296" type="generic">one specimen</specimenCount>
(
<typeStatus id="F64E8839FFC46D75D2DCDAC0BAFD0252" box="[1174,1274,1015,1041]" pageId="45" pageNumber="296" type="paratype">paratype</typeStatus>
) was collected in rotting fruit.
</paragraph>
</subSubSection>
</treatment>
</document>

225
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<paragraph id="294A369BFFC46D75D6DDDDB1BFF802E2" blockId="45.[151,511,1158,1221]" box="[151,511,1158,1185]" pageId="45" pageNumber="296">
<heading id="720281F7FFC46D75D6DDDDB1BFF802E2" bold="true" box="[151,511,1158,1185]" fontSize="11" level="1" pageId="45" pageNumber="296" reason="1">
<emphasis id="1B81EA89FFC46D75D6DDDDB1BFF802E2" bold="true" box="[151,511,1158,1185]" pageId="45" pageNumber="296">
<taxonomicName id="EEF54D18FFC46D75D6DDDDB1BFB202E2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,437,1158,1185]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
<emphasis id="1B81EA89FFC46D75D6DDDDB1BFB202E2" bold="true" box="[151,437,1158,1185]" italics="true" pageId="45" pageNumber="296">Paraphloeostiba telnovi</emphasis>
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<subSubSection id="61EF6510FFC46D75D6DDDD9CBF7C0286" box="[151,379,1195,1221]" pageId="45" pageNumber="296" type="description">
<paragraph id="294A369BFFC46D75D6DDDD9CBF7C0286" blockId="45.[151,511,1158,1221]" box="[151,379,1195,1221]" pageId="45" pageNumber="296">
(
<figureCitation id="B1CE2A1EFFC46D75D6D5DD9CBF030286" box="[159,260,1195,1221]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="45" pageNumber="296">Figs 141</figureCitation>
,
<figureCitation id="B1CE2A1EFFC46D75D75ADD9CBF730286" box="[272,372,1195,1221]" captionStart="FIGURES 144–147" captionStartId="47.[151,264,1614,1638]" captionTargetBox="[171,1427,190,1582]" captionTargetId="figure-16@47.[151,1436,181,1591]" captionTargetPageId="47" captionText="FIGURES 144–147. Paraphloeostiba telnovi (holotype): 144–aedeagus, parameral view, 145–aedeagus, lateral view, 146–male abdominal tergite VIII, 147–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821291" httpUri="https://zenodo.org/record/10821291/files/figure.png" pageId="45" pageNumber="296">144–147</figureCitation>
)
</paragraph>
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<subSubSection id="61EF6510FFC46D75D6DDDDC4BFF40316" pageId="45" pageNumber="296" type="materials_examined">
<paragraph id="294A369BFFC46D75D6DDDDC4BFF40316" blockId="45.[151,1437,1266,1977]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D6DDDDC4BF43034F" bold="true" box="[151,324,1266,1293]" pageId="45" pageNumber="296">Type material.</emphasis>
<typeStatus id="F64E8839FFC46D75D700DDC4BFB5034E" box="[330,434,1267,1293]" pageId="45" pageNumber="296" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFC46D75D7F3DDC4BFC9034E" box="[441,462,1267,1293]" count="1" pageId="45" pageNumber="296" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFC46D75D796DDC4BC3A034E" box="[476,573,1267,1293]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="45" pageNumber="296">Fig. 141</figureCitation>
; dissected): ‘
<collectionCode id="4FE4AE5EFFC46D75D480DDC4BD09034E" box="[714,782,1267,1293]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="45" pageNumber="296">NEW</collectionCode>
<collectingCountry id="51E2760BFFC46D75D55EDDC4BD79034E" box="[788,894,1267,1293]" name="Guinea-Bissau" pageId="45" pageNumber="296">GUINEA</collectingCountry>
| Okapa, |
<date id="5D4B105BFFC46D75D5B8DDC5BA80034E" box="[1010,1159,1266,1293]" pageId="45" pageNumber="296" value="1965-08-27">27.VIII.1965</date>
, | forest litter’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFC46D75D6D7DC20BEB10372" box="[157,182,1303,1329]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="45" pageNumber="296">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC46D75D4D5DC20BD3F0372" box="[671,824,1303,1329]" pageId="45" pageNumber="296" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC46D75D51BDC20BA0B0372" authorityName="Steel" authorityYear="1960" box="[849,1036,1303,1329]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC46D75D51BDC20BA0B0372" box="[849,1036,1303,1329]" italics="true" pageId="45" pageNumber="296">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC46D75D26EDC20BA770372" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1060,1136,1303,1329]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
<emphasis id="1B81EA89FFC46D75D26EDC20BA770372" box="[1060,1136,1303,1329]" italics="true" pageId="45" pageNumber="296">telnovi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D230DC20BABB0372" bold="true" box="[1146,1212,1303,1329]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D230DC20BABB0372" box="[1146,1212,1303,1329]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC46D75D307DC20BB640372" box="[1357,1379,1303,1329]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="45" pageNumber="296" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC46D75D7C7DC0CBFE00316" box="[397,487,1339,1365]" pageId="45" pageNumber="296">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC46D76D68DDC69BB0D0161" lastPageId="46" lastPageNumber="297" pageId="45" pageNumber="296" type="description">
<paragraph id="294A369BFFC46D75D68DDC69BF450382" blockId="45.[151,1437,1266,1977]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DDC69BF5E033B" bold="true" box="[199,345,1374,1400]" pageId="45" pageNumber="296">Description.</emphasis>
Measurements: HW: 0.42; HL: 0.22; OL: 0.12; TL: 0.02; AL: 0.50; PL: 0.27; PWmax: 0.54; PWmin: 0.47; ESL: 0.50; EW: 0.65; MTbL: 0.30; MTrL: 0.16 (MTrL 1–4: 0.05; MTrL 5: 0.11); AW: 0.62; AedL: 0.37; BL: 1.82.
</paragraph>
<paragraph id="294A369BFFC46D75D68DDCFCBA6B00DA" blockId="45.[151,1437,1266,1977]" pageId="45" pageNumber="296">
Habitus as in
<figureCitation id="B1CE2A1EFFC46D75D714DCFCBFB903A6" box="[350,446,1483,1509]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="45" pageNumber="296">Fig. 141</figureCitation>
. Body reddish-brown, with paler laterobasal portions and lateral margin of pronotum and middle portion of elytra; antennomeres 6–11 brown; mouthparts, antennomeres 1–4 and legs yellow (two apical maxillary palpomere slightly darker). Head with indistinct fine punctation, denser and larger in middle portion; neck without visible punctures; pronotum with fine and sparse punctation, slightly finer than that in middle part of head, sparser and finer in middle. Pronotum with dense isodimaetric microsculpture, slightly finer than that in middle portion of head; microsculpture of elytra similar to that in pronotum, but slightly larger.
</paragraph>
<paragraph id="294A369BFFC46D75D68DDF94BA510146" blockId="45.[151,1437,1266,1977]" pageId="45" pageNumber="296">Head about twice as long as broad, with narrow and deep anteocellar foveae. Antennomere 6 slightly and antennomeres 7–10 distinctly transverse; antennomere 5 slightly broader than 4, 6 distinctly broader than 5, 7 slightly longer and distinctly broader than 6, 8–10 slightly shorter and broader than 7.</paragraph>
<paragraph id="294A369BFFC46D75D68DDE38BAB00132" blockId="45.[151,1437,1266,1977]" pageId="45" pageNumber="296">Pronotum twice as broad as long, 1.2 times as broad as head, widest in middle, strongly narrowed posteriad, with widely concave latero-basal margins in front of obtuse hind angles and widely rounded apical angles slightly protruded apicad; lateral margins distinctly impressed; laterobasal impressions wide and deep.</paragraph>
<paragraph id="294A369BFFC46D75D68DDE4CBC6D01D6" blockId="45.[151,1437,1266,1977]" box="[199,618,1915,1941]" pageId="45" pageNumber="296">Elytra 1.3 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFC46D75D68DDEA8BC8501FA" blockId="45.[151,1437,1266,1977]" box="[199,642,1951,1977]" pageId="45" pageNumber="296">Abdomen slightly narrower than elytra.</paragraph>
<caption id="7D8A6613FFC76D76D6DDDAC6BC84026E" ID-DOI="http://doi.org/10.5281/zenodo.10821285" ID-Zenodo-Dep="10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="46" pageNumber="297" startId="46.[151,264,1009,1033]" targetBox="[212,1369,193,974]" targetPageId="46" targetType="figure">
<paragraph id="294A369BFFC76D76D6DDDAC6BC84026E" blockId="46.[151,1436,1009,1070]" pageId="46" pageNumber="297">
<emphasis id="1B81EA89FFC76D76D6DDDAC6BF69024A" bold="true" box="[151,366,1009,1033]" pageId="46" pageNumber="297">FIGURES 141–143.</emphasis>
Habitus of
<taxonomicName id="EEF54D18FFC76D76D7ADDAC5BC420249" box="[487,581,1010,1034]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="tribe" tribe="Omaliini">Omaliini</taxonomicName>
: 141–
<taxonomicName id="EEF54D18FFC76D76D4C8DAC5BD700249" authorityName="Shavrin" authorityYear="2024" box="[642,887,1010,1034]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="telnovi">
<emphasis id="1B81EA89FFC76D76D4C8DAC5BD700249" box="[642,887,1010,1034]" italics="true" pageId="46" pageNumber="297">Paraphloeostiba telnovi</emphasis>
</taxonomicName>
(holotype), 142–
<taxonomicName id="EEF54D18FFC76D76D262DAC5BB22024A" authorityName="Shavrin" authorityYear="2024" box="[1064,1317,1010,1034]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="1B81EA89FFC76D76D262DAC5BB22024A" box="[1064,1317,1010,1034]" italics="true" pageId="46" pageNumber="297">Xanthonomus guineensis</emphasis>
</taxonomicName>
(holotype), 143–
<taxonomicName id="EEF54D18FFC76D76D680DD21BF38026D" authorityName="Shavrin" authorityYear="2024" box="[202,319,1046,1070]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="thayerae">
<emphasis id="1B81EA89FFC76D76D680DD21BF38026D" box="[202,319,1046,1070]" italics="true" pageId="46" pageNumber="297">X. thayerae</emphasis>
</taxonomicName>
(paratype). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFC76D76D68DDDB7BC88030C" blockId="46.[151,1437,1152,2006]" pageId="46" pageNumber="297">
Male. Posterior margin of abdominal tergite VIII rounded (
<figureCitation id="B1CE2A1EFFC76D76D521DDB7BDC802D9" box="[875,975,1152,1178]" captionStart="FIGURES 144–147" captionStartId="47.[151,264,1614,1638]" captionTargetBox="[171,1427,190,1582]" captionTargetId="figure-16@47.[151,1436,181,1591]" captionTargetPageId="47" captionText="FIGURES 144–147. Paraphloeostiba telnovi (holotype): 144–aedeagus, parameral view, 145–aedeagus, lateral view, 146–male abdominal tergite VIII, 147–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821291" httpUri="https://zenodo.org/record/10821291/files/figure.png" pageId="46" pageNumber="297">Fig. 146</figureCitation>
). Posterior margin of abdominal sternite VIII somewhat truncate (
<figureCitation id="B1CE2A1EFFC76D76D7E4DD93BC1702FD" box="[430,528,1188,1214]" captionStart="FIGURES 144–147" captionStartId="47.[151,264,1614,1638]" captionTargetBox="[171,1427,190,1582]" captionTargetId="figure-16@47.[151,1436,181,1591]" captionTargetPageId="47" captionText="FIGURES 144–147. Paraphloeostiba telnovi (holotype): 144–aedeagus, parameral view, 145–aedeagus, lateral view, 146–male abdominal tergite VIII, 147–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821291" httpUri="https://zenodo.org/record/10821291/files/figure.png" pageId="46" pageNumber="297">Fig. 147</figureCitation>
). Aedeagus with very wide basal portion, gradually narrowed toward median lobe with rounded apex; parameres narrow, short, distinctly not reaching apex of median lobe, with two moderately long apical and two preapical setae, inner margin of apical half of each paramere with several moderately long setae; internal sac long and moderately narrow, with several elongate sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFC76D76D361DC27BB970369" box="[1323,1424,1296,1322]" captionStart="FIGURES 144–147" captionStartId="47.[151,264,1614,1638]" captionTargetBox="[171,1427,190,1582]" captionTargetId="figure-16@47.[151,1436,181,1591]" captionTargetPageId="47" captionText="FIGURES 144–147. Paraphloeostiba telnovi (holotype): 144–aedeagus, parameral view, 145–aedeagus, lateral view, 146–male abdominal tergite VIII, 147–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821291" httpUri="https://zenodo.org/record/10821291/files/figure.png" pageId="46" pageNumber="297">Fig. 144</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC76D76D461DC03BC8C030C" box="[555,651,1332,1359]" captionStart="FIGURES 144–147" captionStartId="47.[151,264,1614,1638]" captionTargetBox="[171,1427,190,1582]" captionTargetId="figure-16@47.[151,1436,181,1591]" captionTargetPageId="47" captionText="FIGURES 144–147. Paraphloeostiba telnovi (holotype): 144–aedeagus, parameral view, 145–aedeagus, lateral view, 146–male abdominal tergite VIII, 147–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821291" httpUri="https://zenodo.org/record/10821291/files/figure.png" pageId="46" pageNumber="297">Fig. 145</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC76D76D68DDC6FBF960331" blockId="46.[151,1437,1152,2006]" box="[199,401,1368,1394]" pageId="46" pageNumber="297">Female unknown.</paragraph>
<paragraph id="294A369BFFC76D76D68DDC4BBB0D0161" blockId="46.[151,1437,1152,2006]" pageId="46" pageNumber="297">
<emphasis id="1B81EA89FFC76D76D68DDC4BBFA803D5" bold="true" box="[199,431,1404,1430]" pageId="46" pageNumber="297">Comparative notes.</emphasis>
Based on the general shape of the aedeagus with moderately broad median lobe and narrow parameres, distinctly shorter than apex of the median lobe,
<taxonomicName id="EEF54D18FFC76D76D570DC96BDA403F9" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[826,931,1440,1466]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="telnovi" status="sp. n.">
<emphasis id="1B81EA89FFC76D76D570DC96BDA403F9" box="[826,931,1440,1466]" italics="true" pageId="46" pageNumber="297">P. telnovi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC76D76D5E1DC97BDEB03F9" bold="true" box="[939,1004,1440,1466]" pageId="46" pageNumber="297">
<taxonomicNameLabel id="00B257F2FFC76D76D5E1DC97BDEB03F9" box="[939,1004,1440,1466]" pageId="46" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFC76D76D2C9DC96BB0A03F9" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1155,1293,1440,1466]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="okapensis" status="sp. n.">
<emphasis id="1B81EA89FFC76D76D2C9DC96BB0A03F9" box="[1155,1293,1440,1466]" italics="true" pageId="46" pageNumber="297">P. okapensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC76D76D35CDC97BB5003F9" bold="true" box="[1302,1367,1440,1466]" pageId="46" pageNumber="297">
<taxonomicNameLabel id="00B257F2FFC76D76D35CDC97BB5003F9" box="[1302,1367,1440,1466]" pageId="46" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the shape of the pronotum, more strongly narrowed posteriad and with wide apical portions slightly protruded apicad, the absence of the longitudinal impressions in the middle of the pronotum, the larger punctation of the forebody, broader apical part of the median lobe, slightly narrower parameres and different structure of the internal sac. Besides that, based on the absence of the longitudinal impressions in the middle of the pronotum, and the general shape of the apical portion of the median lobe,
<taxonomicName id="EEF54D18FFC76D76D24EDF62BA6B002D" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1028,1132,1620,1646]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="telnovi" status="sp. n.">
<emphasis id="1B81EA89FFC76D76D24EDF62BA6B002D" box="[1028,1132,1620,1646]" italics="true" pageId="46" pageNumber="297">P. telnovi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC76D76D23EDF63BAB3002D" bold="true" box="[1140,1204,1620,1646]" pageId="46" pageNumber="297">
<taxonomicNameLabel id="00B257F2FFC76D76D23EDF63BAB3002D" box="[1140,1204,1620,1646]" pageId="46" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is somewhat similar to
<taxonomicName id="EEF54D18FFC76D76D6FCDF4EBFF100D0" authority="(Kraatz, 1859)" baseAuthorityName="Kraatz" baseAuthorityYear="1859" box="[182,502,1656,1683]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="1B81EA89FFC76D76D6FCDF4EBF4300D1" box="[182,324,1656,1682]" italics="true" pageId="46" pageNumber="297">P. singularis</emphasis>
(Kraatz, 1859)
</taxonomicName>
, known from the Oriental and Australian regions (
<bibRefCitation id="4D644B6AFFC76D76D276DF4FBAD900D1" author="Herman, L. H." box="[1084,1246,1656,1682]" pageId="46" pageNumber="297" pagination="1 - 7" refId="ref35478" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 421. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001</bibRefCitation>
), and
<taxonomicName id="EEF54D18FFC76D76D36CDF4EBF5000F4" authority="(Cameron, 1925)" baseAuthorityName="Cameron" baseAuthorityYear="1925" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="1B81EA89FFC76D76D36CDF4EBB9A00D1" box="[1318,1437,1656,1682]" italics="true" pageId="46" pageNumber="297">P. apicalis</emphasis>
(Cameron, 1925)
</taxonomicName>
, known from
<collectingCountry id="51E2760BFFC76D76D7A5DFABBC6A00F5" box="[495,621,1692,1718]" name="Philippines" pageId="46" pageNumber="297">Philippines</collectingCountry>
and
<collectingCountry id="51E2760BFFC76D76D4EBDFABBD0900F5" box="[673,782,1692,1718]" name="Indonesia" pageId="46" pageNumber="297">Indonesia</collectingCountry>
(
<bibRefCitation id="4D644B6AFFC76D76D551DFABBDC300F5" author="Shavrin, A. V." box="[795,964,1692,1718]" pageId="46" pageNumber="297" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin 2023b</bibRefCitation>
). From both species it can be distinguished by the shape of the pronotum with distinctly impressed lateral margins and widely rounded apical angles, slightly protruded apicad, the shorter parameres, and the different morphology of the internal sac. From all these species
<taxonomicName id="EEF54D18FFC76D76D3C2DFD2BEE40161" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="telnovi" status="sp. n.">
<emphasis id="1B81EA89FFC76D76D3C2DFD2BEE40161" italics="true" pageId="46" pageNumber="297">P. telnovi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC76D76D6A0DE3FBF2D0161" bold="true" box="[234,298,1800,1826]" pageId="46" pageNumber="297">
<taxonomicNameLabel id="00B257F2FFC76D76D6A0DE3FBF2D0161" box="[234,298,1800,1826]" pageId="46" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the details of external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC76D76D68DDE1BBF2B0129" pageId="46" pageNumber="297" type="distribution">
<paragraph id="294A369BFFC76D76D68DDE1BBF2B0129" blockId="46.[151,1437,1152,2006]" pageId="46" pageNumber="297">
<emphasis id="1B81EA89FFC76D76D68DDE1BBF660105" bold="true" box="[199,353,1836,1862]" pageId="46" pageNumber="297">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFC76D76D722DE1BBC700105" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[360,631,1836,1862]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="297" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
<emphasis id="1B81EA89FFC76D76D722DE1BBC700105" box="[360,631,1836,1862]" italics="true" pageId="46" pageNumber="297">Paraphloeostiba telnovi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC76D76D435DE1BBCB80105" bold="true" box="[639,703,1836,1862]" pageId="46" pageNumber="297">
<taxonomicNameLabel id="00B257F2FFC76D76D435DE1BBCB80105" box="[639,703,1836,1862]" pageId="46" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFC76D76D59CDE1BBA010105" box="[982,1030,1836,1862]" pageId="46" pageNumber="297">type</typeStatus>
locality in the central part of
<collectingCountry id="51E2760BFFC76D76D312DE1BBF2F0129" name="Papua New Guinea" pageId="46" pageNumber="297">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC76D76D68DDE43BD540195" pageId="46" pageNumber="297" type="etymology">
<paragraph id="294A369BFFC76D76D68DDE43BD1001F1" blockId="46.[151,1437,1152,2006]" pageId="46" pageNumber="297">
<emphasis id="1B81EA89FFC76D76D68DDE43BF4901CD" bold="true" box="[199,334,1908,1934]" pageId="46" pageNumber="297">Etymology.</emphasis>
Patronymic, the species is named to honour my colleague, Dmitry Telnov (London), well known coleopterologist and explorer of the Papuan beetle fauna.
</paragraph>
<paragraph id="294A369BFFC76D76D68DDE8BBD540195" blockId="46.[151,1437,1152,2006]" box="[199,851,1980,2006]" pageId="46" pageNumber="297">
<emphasis id="1B81EA89FFC76D76D68DDE8BBF4C0195" bold="true" box="[199,331,1980,2006]" pageId="46" pageNumber="297">Bionomics.</emphasis>
The
<typeStatus id="F64E8839FFC76D76D7CEDE8BBFE10195" box="[388,486,1980,2006]" pageId="46" pageNumber="297" type="holotype">holotype</typeStatus>
was collected in the forest litter.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0FE26C22D837A22E320A0943F55EBACB">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<paragraph id="294A369BFFC66D77D6DDDFEBBFFC00B4" blockId="47.[151,507,1756,1820]" box="[151,507,1756,1783]" pageId="47" pageNumber="298">
<heading id="720281F7FFC66D77D6DDDFEBBFFC00B4" bold="true" box="[151,507,1756,1783]" fontSize="11" level="1" pageId="47" pageNumber="298" reason="1">
<emphasis id="1B81EA89FFC66D77D6DDDFEBBFFC00B4" bold="true" box="[151,507,1756,1783]" pageId="47" pageNumber="298">
<taxonomicName id="EEF54D18FFC66D77D6DDDFEBBFB500B4" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,434,1756,1783]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="298" phylum="Arthropoda" rank="species" species="vitiosa" status="sp. nov.">
<emphasis id="1B81EA89FFC66D77D6DDDFEBBFB500B4" bold="true" box="[151,434,1756,1783]" italics="true" pageId="47" pageNumber="298">Paraphloeostiba vitiosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFC66D77D7F3DFEABFFC00B4" box="[441,507,1757,1783]" pageId="47" pageNumber="298" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC66D77D6DDDE36BF44015F" box="[151,323,1793,1820]" pageId="47" pageNumber="298" type="description">
<paragraph id="294A369BFFC66D77D6DDDE36BF44015F" blockId="47.[151,507,1756,1820]" box="[151,323,1793,1820]" pageId="47" pageNumber="298">
(
<figureCitation id="B1CE2A1EFFC66D77D6D5DE36BF3C015F" box="[159,315,1793,1820]" captionStart="FIGURES 148–151" captionStartId="48.[151,264,1862,1886]" captionTargetBox="[168,1427,193,1830]" captionTargetId="figure-22@48.[151,1436,181,1838]" captionTargetPageId="48" captionText="FIGURES 148–151. Paraphloeostiba vitiosa (holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="47" pageNumber="298">Figs 148–151</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC66D77D6DDDE7EBB5601C4" pageId="47" pageNumber="298" type="materials_examined">
<paragraph id="294A369BFFC66D77D6DDDE7EBB5601C4" blockId="47.[151,1436,1865,2036]" pageId="47" pageNumber="298">
<emphasis id="1B81EA89FFC66D77D6DDDE7EBF4E0120" bold="true" box="[151,329,1865,1891]" pageId="47" pageNumber="298">Type material.</emphasis>
<typeStatus id="F64E8839FFC66D77D719DE7EBFBC0120" box="[339,443,1865,1891]" pageId="47" pageNumber="298" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFC66D77D78CDE7DBFD90127" box="[454,478,1866,1892]" count="1" pageId="47" pageNumber="298" type="male">♂</specimenCount>
: ‘
<collectingRegion id="EB31F879FFC66D77D7BBDE7EBC910120" box="[497,662,1865,1891]" country="Indonesia" name="Papua Barat" pageId="47" pageNumber="298">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Bilogay,
<date id="5D4B105BFFC66D77D5FBDE7EBA280127" box="[945,1071,1865,1892]" pageId="47" pageNumber="298" value="1995-12-29">29.12.1995</date>
|
<quantity id="EE0D9B7EFFC66D77D200DE7EBADE0120" box="[1098,1241,1865,1891]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="47" pageNumber="298" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC66D77D752DE5ABFB601C4" box="[280,433,1901,1927]" pageId="47" pageNumber="298" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC66D77D78FDE5ABC8701C4" authorityName="Steel" authorityYear="1960" box="[453,640,1901,1927]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="298" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC66D77D78FDE5ABC8701C4" box="[453,640,1901,1927]" italics="true" pageId="47" pageNumber="298">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC66D77D4D9DE59BCD901C4" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[659,734,1902,1927]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="298" phylum="Arthropoda" rank="species" species="vitiosa" status="sp. nov.">
<emphasis id="1B81EA89FFC66D77D4D9DE59BCD901C4" box="[659,734,1902,1927]" italics="true" pageId="47" pageNumber="298">vitiosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC66D77D4AFDE5ABD2301C4" bold="true" box="[741,804,1901,1927]" pageId="47" pageNumber="298">
<taxonomicNameLabel id="00B257F2FFC66D77D4AFDE5ABD2301C4" box="[741,804,1901,1927]" pageId="47" pageNumber="298" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC66D77D5E7DE5ABDC401C4" box="[941,963,1901,1927]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="47" pageNumber="298" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC66D77D2BDDE5ABB4201C4" box="[1271,1349,1901,1927]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="47" pageNumber="298">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC66D69D68DDEA6BB58021A" lastPageId="49" lastPageNumber="300" pageId="47" pageNumber="298" type="description">
<paragraph id="294A369BFFC66D77D68DDEA6BD2801B0" blockId="47.[151,1436,1865,2036]" pageId="47" pageNumber="298">
<emphasis id="1B81EA89FFC66D77D68DDEA6BF5E01E8" bold="true" box="[199,345,1937,1963]" pageId="47" pageNumber="298">Description.</emphasis>
Measurements (dissected; specimens with damaged abdomen): HW: 0.42; HL: 0.25; OL: 0.02; TL: 0.04; AL: 0.62; PL: 0.27; PWmax: 0.52; PWmin: 0.49; ESL: 0.57; EW: 0.67; MTbL: 0.31; MTrL: 0.15 (MTrL 1–4: 0.05; MTrL 5: 0.10); AW: 0.67; AedL: 0.42; BL: 2.10.
</paragraph>
<caption id="7D8A6613FFD96D68D6DDDE71BDAC01C2" ID-DOI="http://doi.org/10.5281/zenodo.10821293" ID-Zenodo-Dep="10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="48" pageNumber="299" startId="48.[151,264,1862,1886]" targetBox="[168,1427,193,1830]" targetPageId="48" targetType="figure">
<paragraph id="294A369BFFD96D68D6DDDE71BDAC01C2" blockId="48.[151,1436,1862,1922]" pageId="48" pageNumber="299">
<emphasis id="1B81EA89FFD96D68D6DDDE71BF6B011D" bold="true" box="[151,364,1862,1886]" pageId="48" pageNumber="299">FIGURES 148–151.</emphasis>
<taxonomicName id="EEF54D18FFD96D68D73BDE71BC64011D" authorityName="Shavrin" authorityYear="2024" box="[369,611,1862,1886]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="299" phylum="Arthropoda" rank="species" species="vitiosa">
<emphasis id="1B81EA89FFD96D68D73BDE71BC64011D" box="[369,611,1862,1886]" italics="true" pageId="48" pageNumber="299">Paraphloeostiba vitiosa</emphasis>
</taxonomicName>
(holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFD86D69D68DD9A0BA2A075E" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">Body yellow-brown; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow. Pronotum with dense, large and deep punctation, finer and sparser in parascutellar portion and along suture. Elytra with very dense fine punctation. Pronotum with very dense isodiametric microsculpture, slightly larger than that in middle portion of head; microsculpture of elytra dense, isodiametric, indistinct in middle.</paragraph>
<paragraph id="294A369BFFD86D69D68DD810BD0F0792" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">Head 1.6 times as broad as long, with narrow and deep anteocellar foveae. Distance between ocelli about as long as distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about 1.4 times as long as preapical segment. Antennomeres 8–10 slightly transverse; antennomere 4 slightly narrower and distinctly shorter than 3, 5 slightly longer and broader than 4, 6 broader than 5, 7 slightly longer and broader than 6, 8 slightly broader than 7, 9–10 slightly longer and broader than 8.</paragraph>
<paragraph id="294A369BFFD86D69D68DD8ECBCAA047E" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">Pronotum slightly narrower than long, 1.2 times as broad as head, widest in middle, more narrowed posteriad than anteriad; middle portion with distinct deep, wide longitudinal impressions, more broad and deep in mediobasal portion; laterobasal impressions wide and deep.</paragraph>
<paragraph id="294A369BFFD86D69D68DDB70BDC004C6" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">Elytra slightly more than twice as long as pronotum, with narrowly impressed lateral portions; surface of each elytron with distinct irregular longitudinal elevations between punctures.</paragraph>
<paragraph id="294A369BFFD86D69D68DDBB8BC0004EA" blockId="49.[151,1437,150,1329]" box="[199,519,655,681]" pageId="49" pageNumber="300">Abdomen as broad as elytra.</paragraph>
<paragraph id="294A369BFFD86D69D68DDB84BF1E05E6" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFD86D69D521DB84BDC8048E" box="[875,975,690,717]" captionStart="FIGURES 148–151" captionStartId="48.[151,264,1862,1886]" captionTargetBox="[168,1427,193,1830]" captionTargetId="figure-22@48.[151,1436,181,1838]" captionTargetPageId="48" captionText="FIGURES 148–151. Paraphloeostiba vitiosa (holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="49" pageNumber="300">Fig. 150</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFD86D69D775DBE0BFA504B2" box="[319,418,726,753]" captionStart="FIGURES 148–151" captionStartId="48.[151,264,1862,1886]" captionTargetBox="[168,1427,193,1830]" captionTargetId="figure-22@48.[151,1436,181,1838]" captionTargetPageId="48" captionText="FIGURES 148–151. Paraphloeostiba vitiosa (holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="49" pageNumber="300">Fig. 151</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward wide median lobe, with rounded apex; parameres significantly longer than apex of median lobe, with ditsinctly broadened apical portions, with several long apical and preapical setae on inner surface of each setiferous lobe, with several short setae along inner and outer margins of each paramere; internal sac complicated, moderately wide and long, with several large sclerotized structures: curved in middle and elongate in basal portion (
<figureCitation id="B1CE2A1EFFD86D69D5F8DA50BA1205C2" box="[946,1045,871,897]" captionStart="FIGURES 148–151" captionStartId="48.[151,264,1862,1886]" captionTargetBox="[168,1427,193,1830]" captionTargetId="figure-22@48.[151,1436,181,1838]" captionTargetPageId="48" captionText="FIGURES 148–151. Paraphloeostiba vitiosa (holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="49" pageNumber="300">Fig. 148</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFD86D69D6FEDABCBF1305E6" box="[180,276,907,933]" captionStart="FIGURES 148–151" captionStartId="48.[151,264,1862,1886]" captionTargetBox="[168,1427,193,1830]" captionTargetId="figure-22@48.[151,1436,181,1838]" captionTargetPageId="48" captionText="FIGURES 148–151. Paraphloeostiba vitiosa (holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="49" pageNumber="300">Fig. 149</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFD86D69D68DDA98BF96058A" blockId="49.[151,1437,150,1329]" box="[199,401,943,969]" pageId="49" pageNumber="300">Female unknown.</paragraph>
<paragraph id="294A369BFFD86D69D68DDAE5BB58021A" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">
<emphasis id="1B81EA89FFD86D69D68DDAE5BFB205AE" bold="true" box="[199,437,978,1005]" pageId="49" pageNumber="300">Comparative notes.</emphasis>
<taxonomicName id="EEF54D18FFD86D69D78BDAE4BCD505AF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[449,722,979,1005]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="vitiosa" status="sp. nov.">
<emphasis id="1B81EA89FFD86D69D78BDAE4BCD505AF" box="[449,722,979,1005]" italics="true" pageId="49" pageNumber="300">Paraphloeostiba vitiosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFD86D69D497DAE4BD2505AE" bold="true" box="[733,802,979,1005]" pageId="49" pageNumber="300">
<taxonomicNameLabel id="00B257F2FFD86D69D497DAE4BD2505AE" box="[733,802,979,1005]" pageId="49" pageNumber="300" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from other species of the genus by the slightly transverse antennomeres 8–10, elongate elytra, slightly more than twice as long as pronotum, the general shape of the aedeagus, with very wide middle part of the median lobe, gradually narrowed toward apex, the parameres, significantly exceeding apex of the median lobe, and the complicated structure of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFD86D69D68DDD55BF0B02E2" pageId="49" pageNumber="300" type="distribution">
<paragraph id="294A369BFFD86D69D68DDD55BF0B02E2" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">
<emphasis id="1B81EA89FFD86D69D68DDD55BF66023F" bold="true" box="[199,353,1122,1148]" pageId="49" pageNumber="300">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFD86D69D720DD54BC7D023F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[362,634,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="vitiosa" status="sp. nov.">
<emphasis id="1B81EA89FFD86D69D720DD54BC7D023F" box="[362,634,1123,1149]" italics="true" pageId="49" pageNumber="300">Paraphloeostiba vitiosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFD86D69D4C9DD54BCC1023E" bold="true" box="[643,710,1123,1149]" pageId="49" pageNumber="300">
<taxonomicNameLabel id="00B257F2FFD86D69D4C9DD54BCC1023E" box="[643,710,1123,1149]" pageId="49" pageNumber="300" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFD86D69D5A0DD54BA1D023E" box="[1002,1050,1123,1149]" pageId="49" pageNumber="300">type</typeStatus>
locality in Western New
<collectingCountry id="51E2760BFFD86D69D30EDD54BB9F023E" box="[1348,1432,1123,1149]" name="Guinea-Bissau" pageId="49" pageNumber="300">Guinea</collectingCountry>
,
<collectingCountry id="51E2760BFFD86D69D6DDDDB0BF0F02E2" box="[151,264,1159,1185]" name="Indonesia" pageId="49" pageNumber="300">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFD86D69D68DDD9CBF370372" pageId="49" pageNumber="300" type="etymology">
<paragraph id="294A369BFFD86D69D68DDD9CBF2D02AA" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">
<emphasis id="1B81EA89FFD86D69D68DDD9CBF490286" bold="true" box="[199,334,1195,1221]" pageId="49" pageNumber="300">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFD86D69D48ADD9CBD110287" box="[704,790,1195,1220]" italics="true" pageId="49" pageNumber="300">vitiosus</emphasis>
, -
<emphasis id="1B81EA89FFD86D69D564DD9CBD3B0287" box="[814,828,1195,1220]" italics="true" pageId="49" pageNumber="300">a</emphasis>
, -
<emphasis id="1B81EA89FFD86D69D51EDD9CBD710287" box="[852,886,1195,1220]" italics="true" pageId="49" pageNumber="300">um</emphasis>
(damaged). It alludes to the general condition of the
<typeStatus id="F64E8839FFD86D69D68ADDF8BF2102AA" box="[192,294,1231,1257]" pageId="49" pageNumber="300" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="294A369BFFD86D69D68DDDC5BF370372" blockId="49.[151,1437,150,1329]" pageId="49" pageNumber="300">
<emphasis id="1B81EA89FFD86D69D68DDDC5BF4C034F" bold="true" box="[199,331,1266,1292]" pageId="49" pageNumber="300">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="EE0D9B7EFFD86D69D512DDC4BA0A034E" box="[856,1037,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="49" pageNumber="300" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

286
data/A1/5C/87/A15C878DFFC96D78D6DDDAC1BD6A01C6.xml Normal file
View file

@ -0,0 +1,286 @@
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<paragraph id="294A369BFFC96D78D6DDDAC1BCA10253" blockId="32.[151,678,1014,1077]" box="[151,678,1014,1041]" pageId="32" pageNumber="283">
<heading id="720281F7FFC96D78D6DDDAC1BCA10253" bold="true" box="[151,678,1014,1041]" fontSize="11" level="1" pageId="32" pageNumber="283" reason="1">
<taxonomicName id="EEF54D18FFC96D78D6DDDAC1BCA10253" ID-CoL="762XP" authority="(Cameron, 1931)" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[151,678,1014,1041]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFC96D78D6DDDAC1BCA10253" bold="true" box="[151,678,1014,1041]" pageId="32" pageNumber="283">
<emphasis id="1B81EA89FFC96D78D6DDDAC1BFC90252" bold="true" box="[151,462,1014,1041]" italics="true" pageId="32" pageNumber="283">Paraphloeostiba papuana</emphasis>
(
<bibRefCitation id="4D644B6AFFC96D78D797DAC1BC990253" author="Cameron, M." box="[477,670,1014,1040]" pageId="32" pageNumber="283" pagination="355 - 364" refId="ref35403" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article" year="1931">Cameron, 1931</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC96D78D6DDDD2CBF200276" box="[151,295,1051,1077]" pageId="32" pageNumber="283" type="description">
<paragraph id="294A369BFFC96D78D6DDDD2CBF200276" blockId="32.[151,678,1014,1077]" box="[151,295,1051,1077]" pageId="32" pageNumber="283">
(
<figureCitation id="B1CE2A1EFFC96D78D6D5DD2CBF180276" box="[159,287,1051,1077]" captionStart-0="FIGURES 92–95" captionStart-1="FIGURE 96" captionStartId-0="33.[151,264,1780,1804]" captionStartId-1="34.[151,250,892,916]" captionTargetBox-0="[160,1427,189,1747]" captionTargetBox-1="[189,1398,193,868]" captionTargetId-0="figure-16@33.[151,1436,181,1756]" captionTargetId-1="figure-22@34.[189,1398,181,868]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="FIGURES 92–95. Paraphloeostiba papuana (Western New Guinea): 92–aedeagus, parameral view, 93–aedeagus, lateral view, 94–male abdominal tergite VIII, 95–male abdominal sternite VIII. Scale bar: 0.1 mm." captionText-1="FIGURE 96. Distribution of Paraphloeostiba papuana in New Guinea." figureDoi-0="http://doi.org/10.5281/zenodo.10821261" figureDoi-1="http://doi.org/10.5281/zenodo.10821263" httpUri-0="https://zenodo.org/record/10821261/files/figure.png" httpUri-1="https://zenodo.org/record/10821263/files/figure.png" pageId="32" pageNumber="283">Figs 92–96</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC96D78D6DDDD68BD2702F6" pageId="32" pageNumber="283" type="reference_group">
<paragraph id="294A369BFFC96D78D6DDDD68BC670234" blockId="32.[151,808,1119,1205]" box="[151,608,1119,1143]" pageId="32" pageNumber="283">
<treatmentCitationGroup id="09E511B5FFC96D78D6DDDD68BC670234" box="[151,608,1119,1143]" pageId="32" pageNumber="283">
<treatmentCitation id="A854108AFFC96D78D6DDDD68BC670234" author="Cameron, M." box="[151,608,1119,1143]" page="356" pageId="32" pageNumber="283" year="1931">
<taxonomicName id="EEF54D18FFC96D78D6DDDD68BC670234" authority="Cameron, 1931: 356" authorityName="Cameron" authorityPageNumber="356" authorityYear="1931" box="[151,608,1119,1143]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="1B81EA89FFC96D78D6DDDD68BF8D0234" box="[151,394,1119,1143]" italics="true" pageId="32" pageNumber="283">Phloeonomus papuanus</emphasis>
<bibRefCitation id="4D644B6AFFC96D78D7DADD68BC670234" author="Cameron, M." box="[400,608,1119,1143]" pageId="32" pageNumber="283" pagination="355 - 364" refId="ref35403" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article" year="1931">Cameron, 1931: 356</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="294A369BFFC96D78D6DDDD49BD2F02D5" blockId="32.[151,808,1119,1205]" box="[151,808,1150,1174]" pageId="32" pageNumber="283">
<treatmentCitationGroup id="09E511B5FFC96D78D6DDDD49BD2F02D5" box="[151,808,1150,1174]" pageId="32" pageNumber="283">
<taxonomicName id="EEF54D18FFC96D78D6DDDD49BC2B02D5" authorityName="Cameron" authorityYear="1931" box="[151,556,1150,1174]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="papuanus" subGenus="Phloeonomus">
<emphasis id="1B81EA89FFC96D78D6DDDD49BF2602D5" box="[151,289,1150,1174]" italics="true" pageId="32" pageNumber="283">Phloeonomus</emphasis>
(
<emphasis id="1B81EA89FFC96D78D77ADD49BFBD02D5" box="[304,442,1150,1174]" italics="true" pageId="32" pageNumber="283">Phloeonomus</emphasis>
)
<emphasis id="1B81EA89FFC96D78D783DD48BC2B02D5" box="[457,556,1151,1174]" italics="true" pageId="32" pageNumber="283">papuanus</emphasis>
</taxonomicName>
:
<treatmentCitation id="A854108AFFC96D78D472DD49BD2F02D5" author="Scheerpeltz, O. von" box="[568,808,1150,1174]" page="1050" pageId="32" pageNumber="283" year="1933">
<bibRefCitation id="4D644B6AFFC96D78D472DD49BD2F02D5" author="Scheerpeltz, O. von" box="[568,808,1150,1174]" pageId="32" pageNumber="283" pagination="989 - 1500" refId="ref35653" refString="Scheerpeltz, O. von (1933) Staphylinidae VII (pars 129). In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. VI. Staphylinidae II. W. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz 1933: 1050</bibRefCitation>
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</paragraph>
<paragraph id="294A369BFFC96D78D6DDDDABBD2702F6" blockId="32.[151,808,1119,1205]" box="[151,800,1180,1205]" pageId="32" pageNumber="283">
<treatmentCitationGroup id="09E511B5FFC96D78D6DDDDABBD2702F6" box="[151,800,1180,1205]" pageId="32" pageNumber="283">
<taxonomicName id="EEF54D18FFC96D78D6DDDDABBF9802F7" ID-CoL="762XP" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[151,415,1180,1204]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFC96D78D6DDDDABBF9802F7" box="[151,415,1180,1204]" italics="true" pageId="32" pageNumber="283">Paraphloeostiba papuana</emphasis>
</taxonomicName>
:
<treatmentCitation id="A854108AFFC96D78D7E6DDABBC4A02F7" author="Steel, W. O." box="[428,589,1180,1204]" page="153" pageId="32" pageNumber="283" year="1960">
<bibRefCitation id="4D644B6AFFC96D78D7E6DDABBC4A02F7" author="Steel, W. O." box="[428,589,1180,1204]" pageId="32" pageNumber="283" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960: 153</bibRefCitation>
</treatmentCitation>
,
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<bibRefCitation id="4D644B6AFFC96D78D412DDABBD2702F6" author="Shavrin, A. V." box="[600,800,1180,1205]" pageId="32" pageNumber="283" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin 2023b: 493</bibRefCitation>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC96D78D6DDDDD5BCFF03CE" pageId="32" pageNumber="283" type="materials_examined">
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<emphasis id="1B81EA89FFC96D78D6DDDDD5BCF602BE" bold="true" box="[151,753,1250,1277]" pageId="32" pageNumber="283">
Material examined.
<materialsCitation id="999D3CC6FFC96D78D7C4DDD5BCF602BE" box="[398,753,1250,1277]" collectionCode="WESTERN, NEW" country="Indonesia" location="INDONESIA" pageId="32" pageNumber="283" specimenCount="1">
<collectingCountry id="51E2760BFFC96D78D7C4DDD5BC3602BF" box="[398,561,1250,1276]" name="Indonesia" pageId="32" pageNumber="283">INDONESIA</collectingCountry>
:
<collectionCode id="4FE4AE5EFFC96D78D40ADDD5BCB702BF" box="[576,688,1250,1276]" pageId="32" pageNumber="283">WESTERN</collectionCode>
<collectionCode id="4FE4AE5EFFC96D78D4F0DDD4BCF602BE" box="[698,753,1251,1277]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="32" pageNumber="283">NEW</collectionCode>
</materialsCitation>
</emphasis>
<materialsCitation id="999D3CC6FFC96D78D4B1DDD5BA4902BE" box="[763,1102,1250,1277]" collectionCode="IRIAN, JAYA" country="Guinea-Bissau" location="GUINEA" pageId="32" pageNumber="283" specimenCount="1" specimenCount-female="1">
<emphasis id="1B81EA89FFC96D78D4B1DDD5BD5302BF" bold="true" box="[763,852,1250,1276]" pageId="32" pageNumber="283">
<collectingCountry id="51E2760BFFC96D78D4B1DDD5BD5302BF" box="[763,852,1250,1276]" name="Guinea-Bissau" pageId="32" pageNumber="283">GUINEA</collectingCountry>
</emphasis>
:
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: ‘
<collectionCode id="4FE4AE5EFFC96D78D5E0DDD4BA4902BE" box="[938,1102,1251,1277]" pageId="32" pageNumber="283">IRIAN JAYA</collectionCode>
</materialsCitation>
:
<materialsCitation id="999D3CC6FFC96D78D210DDD4BA590362" collectingDate="1997-10" collectionCode="NMW, NEW" collectorName="M. Balke" country="Guinea-Bissau" elevation="750" location="Nabire Prov." municipality="Ilaga" pageId="32" pageNumber="283" specimenCode="LUX 54" specimenCount="6" specimenCount-female="5" specimenCount-male="1">
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Nabire—
<collectingMunicipality id="C92EACE1FFC96D78D32BDDD4BB9F02BE" box="[1377,1432,1251,1277]" pageId="32" pageNumber="283">Ilaga</collectingMunicipality>
, km 54
<date id="5D4B105BFFC96D78D6A2DC30BF390362" box="[232,318,1287,1313]" pageId="32" pageNumber="283" value="1997-10">
<collectingDate id="4D0FE9B3FFC96D78D6A2DC30BF390362" box="[232,318,1287,1313]" pageId="32" pageNumber="283" value="1997-10">X.1997</collectingDate>
</date>
, ca.
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</quantity>
, light leg.
<collectorName id="8400534DFFC96D78D464DC30BC9F0362" box="[558,664,1287,1313]" pageId="32" pageNumber="283">M. Balke</collectorName>
(
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)’ (
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17.
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.1965 dead mouse’, ‘
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.’ (
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);
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: ‘
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, forest litter’, ‘
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.
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|
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.’ (
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28.
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.1965, dead branches’, ‘
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.’ (
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC96D78D68DDCA0BD6A01C6" pageId="32" pageNumber="283" type="discussion">
<paragraph id="294A369BFFC96D78D68DDCA0BA5D005E" blockId="32.[151,1437,1250,1925]" pageId="32" pageNumber="283">
<emphasis id="1B81EA89FFC96D78D68DDCA0BF3C03F2" bold="true" box="[199,315,1431,1457]" pageId="32" pageNumber="283">Remarks.</emphasis>
<taxonomicName id="EEF54D18FFC96D78D70BDCA0BC4A03F3" authorityName="Cameron" authorityYear="1931" box="[321,589,1431,1457]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="1B81EA89FFC96D78D70BDCA0BC4A03F3" box="[321,589,1431,1457]" italics="true" pageId="32" pageNumber="283">Phloeonomus papuanus</emphasis>
</taxonomicName>
was originally described from
<collectingCountry id="51E2760BFFC96D78D5EDDCA0BA8503F2" box="[935,1154,1431,1457]" name="Papua New Guinea" pageId="32" pageNumber="283">Papua New Guinea</collectingCountry>
.
<bibRefCitation id="4D644B6AFFC96D78D2C6DCA0BB1203F2" author="Steel, W. O." box="[1164,1301,1431,1457]" pageId="32" pageNumber="283" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
redescribed it and transferred to the genus
<taxonomicName id="EEF54D18FFC96D78D7A5DC8CBCAD0396" authorityName="Steel" authorityYear="1960" box="[495,682,1467,1493]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC96D78D7A5DC8CBCAD0396" box="[495,682,1467,1493]" italics="true" pageId="32" pageNumber="283">Paraphloeostiba</emphasis>
</taxonomicName>
.
<bibRefCitation id="4D644B6AFFC96D78D4F2DC8CBD770396" author="Shavrin, A. V." box="[696,880,1467,1493]" pageId="32" pageNumber="283" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
redescribed it and illustrated the spermatheca for the first time. The body length of this species can vary from
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(
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). Prior to the present study, the male of
<taxonomicName id="EEF54D18FFC96D78D756DF34BF9D005F" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[284,410,1539,1564]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFC96D78D756DF34BF9D005F" box="[284,410,1539,1564]" italics="true" pageId="32" pageNumber="283">P. papuana</emphasis>
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was unknown. The description of the male is presented below.
</paragraph>
<paragraph id="294A369BFFC96D78D68DDF10BA1D00B6" blockId="32.[151,1437,1250,1925]" pageId="32" pageNumber="283">
Male. Posterior margin of abdominal tergite VIII rounded (
<figureCitation id="B1CE2A1EFFC96D78D51FDF10BDAF0002" box="[853,936,1575,1601]" captionStart="FIGURES 92–95" captionStartId="33.[151,264,1780,1804]" captionTargetBox="[160,1427,189,1747]" captionTargetId="figure-16@33.[151,1436,181,1756]" captionTargetPageId="33" captionText="FIGURES 92–95. Paraphloeostiba papuana (Western New Guinea): 92–aedeagus, parameral view, 93–aedeagus, lateral view, 94–male abdominal tergite VIII, 95–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821261" httpUri="https://zenodo.org/record/10821261/files/figure.png" pageId="32" pageNumber="283">Fig. 94</figureCitation>
). Posterior margin of abdominal sternite VIII strongly and widely concave (
<figureCitation id="B1CE2A1EFFC96D78D7A4DF7CBC430026" box="[494,580,1610,1637]" captionStart="FIGURES 92–95" captionStartId="33.[151,264,1780,1804]" captionTargetBox="[160,1427,189,1747]" captionTargetId="figure-16@33.[151,1436,181,1756]" captionTargetPageId="33" captionText="FIGURES 92–95. Paraphloeostiba papuana (Western New Guinea): 92–aedeagus, parameral view, 93–aedeagus, lateral view, 94–male abdominal tergite VIII, 95–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821261" httpUri="https://zenodo.org/record/10821261/files/figure.png" pageId="32" pageNumber="283">Fig. 95</figureCitation>
). Aedeagus (AedL: 0.40) with very wide basal portion, gradually narrowed toward narrow median lobe with small rounded apex; parameres slightly not reaching apex of median lobe, with strongly broadened apical portions, with two long apical and several short preapical setae, middle portions of each paramere with several very short setae; internal sac moderately long and narrow, with three elongate sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFC96D78D783DFECBC1A00B6" box="[457,541,1755,1781]" captionStart="FIGURES 92–95" captionStartId="33.[151,264,1780,1804]" captionTargetBox="[160,1427,189,1747]" captionTargetId="figure-16@33.[151,1436,181,1756]" captionTargetPageId="33" captionText="FIGURES 92–95. Paraphloeostiba papuana (Western New Guinea): 92–aedeagus, parameral view, 93–aedeagus, lateral view, 94–male abdominal tergite VIII, 95–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821261" httpUri="https://zenodo.org/record/10821261/files/figure.png" pageId="32" pageNumber="283">Fig. 92</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC96D78D58EDFECBA1100B6" box="[964,1046,1755,1781]" captionStart="FIGURES 92–95" captionStartId="33.[151,264,1780,1804]" captionTargetBox="[160,1427,189,1747]" captionTargetId="figure-16@33.[151,1436,181,1756]" captionTargetPageId="33" captionText="FIGURES 92–95. Paraphloeostiba papuana (Western New Guinea): 92–aedeagus, parameral view, 93–aedeagus, lateral view, 94–male abdominal tergite VIII, 95–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821261" httpUri="https://zenodo.org/record/10821261/files/figure.png" pageId="32" pageNumber="283">Fig. 93</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC96D78D68DDFC8BFF9017E" blockId="32.[151,1437,1250,1925]" pageId="32" pageNumber="283">
<taxonomicName id="EEF54D18FFC96D78D68DDFC8BFED015B" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[199,490,1791,1817]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFC96D78D68DDFC8BFED015B" box="[199,490,1791,1817]" italics="true" pageId="32" pageNumber="283">Paraphloeostiba papuana</emphasis>
</taxonomicName>
is known from several localities in Western New
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(
<collectingCountry id="51E2760BFFC96D78D225DFC8BAD8015A" box="[1135,1247,1791,1817]" name="Indonesia" pageId="32" pageNumber="283">Indonesia</collectingCountry>
) and eastern part of
<collectingCountry id="51E2760BFFC96D78D6FCDE14BF89017E" box="[182,398,1827,1853]" name="Papua New Guinea" pageId="32" pageNumber="283">Papua New Guinea</collectingCountry>
(
<figureCitation id="B1CE2A1EFFC96D78D7D7DE14BFF6017E" box="[413,497,1827,1853]" captionStart="FIGURE 96" captionStartId="34.[151,250,892,916]" captionTargetBox="[189,1398,193,868]" captionTargetId="figure-22@34.[189,1398,181,868]" captionTargetPageId="34" captionText="FIGURE 96. Distribution of Paraphloeostiba papuana in New Guinea." figureDoi="http://doi.org/10.5281/zenodo.10821263" httpUri="https://zenodo.org/record/10821263/files/figure.png" pageId="32" pageNumber="283">Fig. 96</figureCitation>
).
</paragraph>
<paragraph id="294A369BFFC96D78D68DDE70BD6A01C6" blockId="32.[151,1437,1250,1925]" pageId="32" pageNumber="283">
The species was collected at altitudes from
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a.s.l. Some specimens from Western New
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were collected on dead mouse, in forest litter and dead branches.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart id="61F3E6214FFA43BE43C4889440F11874">Shavrin, Alexey V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="216EA5ED28C062F26F4944E1B99AE2BD">text</mods:typeOfResource>
<mods:relatedItem id="691452AA90337B9A7193F33626FE6C9A" type="host">
<mods:titleInfo id="6FA662368471ED0916C7940A467494A3">
<mods:title id="0C714ADD3C9C13AAFEC9A63B32EBDCE8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6EB0E93CC5AB07EB192C18951F43BA80">
<mods:date id="45076557C82646A87B0E0E804A57236A">2024</mods:date>
<mods:detail id="54CF61A04AE053F697AFE54DFED15F27" type="pubDate">
<mods:number id="052B0E0804E8EA606CE16AB3D33673CF">2024-03-14</mods:number>
</mods:detail>
<mods:detail id="2E0430501731B1F184A2E07AC8C4469C" type="volume">
<mods:number id="8F765233D4BCCBACC55F3C7F59FEAC21">5424</mods:number>
</mods:detail>
<mods:detail id="B2530D36963CB747EACF0F64041E31AC" type="issue">
<mods:number id="96ACDB14747E3A3E295D9FB358B35E8A">3</mods:number>
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<mods:extent id="8199E9262B10F8A4EE88124D2F218925" unit="page">
<mods:start id="6FAF2C4452EE6140E8BDEC511FDE2138">251</mods:start>
<mods:end id="2DD659EE9B3967DC8185D5E70D8E7D99">307</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location id="DE707AC9A43249CDBD10489C2261D148">
<mods:url id="BD0581AEA700422B0280BF325253E8C2">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
</mods:location>
<mods:classification id="46E7B1E62305A435A87010122534B5D4">journal article</mods:classification>
<mods:identifier id="8B77A6525FC14BCB419BD6D245957D69" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="4B0F7A8DB082B51C8DA934611401B004" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B7AFEE4F0D6F55D4A281B8E1482212EF" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="1EE95A9AC399036D5AA0FD467BADE544" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFCB6D7CD6DDDAD1BD300422" LSID="urn:lsid:plazi:treatment:A15C878DFFCB6D7CD6DDDAD1BD300422" httpUri="http://treatment.plazi.org/id/A15C878DFFCB6D7CD6DDDAD1BD300422" lastPageId="36" lastPageNumber="287" pageId="34" pageNumber="285">
<subSubSection id="61EF6510FFCB6D7AD6DDDAD1BC330242" box="[151,564,998,1025]" pageId="34" pageNumber="285" type="nomenclature">
<paragraph id="294A369BFFCB6D7AD6DDDAD1BC330242" blockId="34.[151,564,998,1061]" box="[151,564,998,1025]" pageId="34" pageNumber="285">
<heading id="720281F7FFCB6D7AD6DDDAD1BC330242" bold="true" box="[151,564,998,1025]" fontSize="11" level="1" pageId="34" pageNumber="285" reason="1">
<emphasis id="1B81EA89FFCB6D7AD6DDDAD1BC330242" bold="true" box="[151,564,998,1025]" pageId="34" pageNumber="285">
<taxonomicName id="EEF54D18FFCB6D7AD6DDDAD1BFEC0242" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,491,998,1025]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCB6D7AD6DDDAD1BFEC0242" bold="true" box="[151,491,998,1025]" italics="true" pageId="34" pageNumber="285">Paraphloeostiba penelopeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFCB6D7AD7B8DAD0BC330242" box="[498,564,999,1025]" pageId="34" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCB6D7AD6DDDD3CBF580266" box="[151,351,1035,1061]" pageId="34" pageNumber="285" type="description">
<paragraph id="294A369BFFCB6D7AD6DDDD3CBF580266" blockId="34.[151,564,998,1061]" box="[151,351,1035,1061]" pageId="34" pageNumber="285">
(
<figureCitation id="B1CE2A1EFFCB6D7AD6D5DD3CBEF00266" box="[159,247,1035,1061]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="34" pageNumber="285">Figs 76</figureCitation>
,
<figureCitation id="B1CE2A1EFFCB6D7AD748DD3CBF500266" box="[258,343,1035,1061]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="34" pageNumber="285">97–103</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCB6D7AD6DDDD64BCBA0362" pageId="34" pageNumber="285" type="materials_examined">
<paragraph id="294A369BFFCB6D7AD6DDDD64BD9102F6" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
<emphasis id="1B81EA89FFCB6D7AD6DDDD64BF43022F" bold="true" box="[151,324,1106,1133]" pageId="34" pageNumber="285">Type material.</emphasis>
<typeStatus id="F64E8839FFCB6D7AD703DD64BFB6022E" box="[329,433,1107,1133]" pageId="34" pageNumber="285" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFCB6D7AD7FDDD64BFC8022E" box="[439,463,1107,1133]" count="1" pageId="34" pageNumber="285" type="male">♂</specimenCount>
: ‘SOLOMON IS. |
<collectingRegion id="EB31F879FFCB6D7AD4E9DD64BD28022E" box="[675,815,1107,1133]" country="Solomon Islands" name="Guadalcanal" pageId="34" pageNumber="285">Guadalcanal</collectingRegion>
| Mt. Austen [handwritten] | Jan [handwritten] 1966 [the last number “6” is handwritten] | 19993 [handwritten] | P. Greenslade’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFCB6D7AD263DD40BAC502D2" box="[1065,1218,1143,1169]" pageId="34" pageNumber="285" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFCB6D7AD29FDD40BB9702D2" authorityName="Steel" authorityYear="1960" box="[1237,1424,1143,1169]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFCB6D7AD29FDD40BB9702D2" box="[1237,1424,1143,1169]" italics="true" pageId="34" pageNumber="285">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFCB6D7AD6DDDDACBF1002F6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,279,1179,1205]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCB6D7AD6DDDDACBF1002F6" box="[151,279,1179,1205]" italics="true" pageId="34" pageNumber="285">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCB6D7AD754DDACBF5A02F6" bold="true" box="[286,349,1179,1205]" pageId="34" pageNumber="285">
<taxonomicNameLabel id="00B257F2FFCB6D7AD754DDACBF5A02F6" box="[286,349,1179,1205]" pageId="34" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFCB6D7AD7ACDDACBFFB02F6" box="[486,508,1179,1205]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="34" pageNumber="285" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFCB6D7AD57ADDACBD8D02F6" box="[816,906,1179,1205]" pageId="34" pageNumber="285">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDD88BCBA0362" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
<typeStatus id="F64E8839FFCB6D7AD68DDD88BF3F029A" box="[199,312,1215,1241]" pageId="34" pageNumber="285" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFCB6D7AD709DD88BF96029A" box="[323,401,1215,1241]" count="14" pageId="34" pageNumber="285" type="male">14 ♂♂</specimenCount>
(
<specimenCount id="3FF3FD12FFCB6D7AD7EADD88BC3B029A" box="[416,572,1215,1241]" count="1" pageId="34" pageNumber="285" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="3FF3FD12FFCB6D7AD48BDD88BD08029A" box="[705,783,1215,1241]" count="18" pageId="34" pageNumber="285" type="female">18 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFCB6D7AD554DD88BDBD029A" box="[798,954,1215,1241]" count="1" pageId="34" pageNumber="285" type="generic">one specimen</specimenCount>
dissected): ‘SOLOMON IS. |
<collectingRegion id="EB31F879FFCB6D7AD35ADD88BB9B029A" box="[1296,1436,1215,1241]" country="Solomon Islands" name="Guadalcanal" pageId="34" pageNumber="285">Guadalcanal</collectingRegion>
Mt. | Austen
<date id="5D4B105BFFCB6D7AD77ADDD4BF9F02BE" box="[304,408,1251,1277]" pageId="34" pageNumber="285" value="1966-01">Jan 1966</date>
| 19993 P. | Greenslade’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFCB6D7AD508DDD4BDD502BE" box="[834,978,1251,1277]" pageId="34" pageNumber="285" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFCB6D7AD5A6DDD4BAA002BE" authorityName="Steel" authorityYear="1960" box="[1004,1191,1251,1277]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFCB6D7AD5A6DDD4BAA002BE" box="[1004,1191,1251,1277]" italics="true" pageId="34" pageNumber="285">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFCB6D7AD28BDDD4BB4602BE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1217,1345,1251,1277]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCB6D7AD28BDDD4BB4602BE" box="[1217,1345,1251,1277]" italics="true" pageId="34" pageNumber="285">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCB6D7AD300DDD4BB8A02BE" bold="true" box="[1354,1421,1251,1277]" pageId="34" pageNumber="285">
<taxonomicNameLabel id="00B257F2FFCB6D7AD300DDD4BB8A02BE" box="[1354,1421,1251,1277]" pageId="34" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFCB6D7AD747DC30BF240362" box="[269,291,1287,1313]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="34" pageNumber="285" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFCB6D7AD41DDC30BCB60362" box="[599,689,1287,1313]" pageId="34" pageNumber="285">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCB6D7CD68DDC1DBC1407EE" lastPageId="36" lastPageNumber="287" pageId="34" pageNumber="285" type="description">
<paragraph id="294A369BFFCB6D7AD68DDC1DBA7B03CE" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
<emphasis id="1B81EA89FFCB6D7AD68DDC1DBF5E0307" bold="true" box="[199,345,1322,1348]" pageId="34" pageNumber="285">Description.</emphasis>
Measurements (n=33): HW: 0.40; HL: 0.17–0.22; OL: 0.10–0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFCB6D7AD2A2DC1CBB560306" box="[1256,1361,1323,1349]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
): 0.45; PL: 0.18–0.25; PW: 0.43–0.52; ESL: 0.36–0.45; EW: 0.55–0.60; MTbL (
<typeStatus id="F64E8839FFCB6D7AD591DC78BA43032A" box="[987,1092,1359,1385]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFCB6D7AD2A4DC78BB50032A" box="[1262,1367,1359,1385]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
): 0.17 (MTrL 1–4: 0.05; MTrL 5: 0.12); AW: 0.58; AedL: 0.32; BL: 1.40–2.00 (
<typeStatus id="F64E8839FFCB6D7AD582DC44BA3703CE" box="[968,1072,1395,1421]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
: 1.85).
</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDCA0BFDF00CA" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
Habitus as in
<figureCitation id="B1CE2A1EFFCB6D7AD727DCA0BFC303F2" box="[365,452,1431,1457]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="34" pageNumber="285">Fig. 76</figureCitation>
. Body yellow-brown, with reddish-brown apical portions of elytra (some specimens with slightly darker neck, head (or only clypeus) and apical parts of abdominal tergites III–VI). Head and neck, pronotum and scutellum without punctation; elytra with moderately dense and large punctation, finer and denser around scutellum and finer along suture; abdominal tergites without visible punctation. Pronotum with very dense isodiametric microsculpture, slightly finer than that in middle portion of head; elytra with dense and coarse isodiametric microreticulation, slightly larger than that in pronotum; abdominal tergites with dense transverse or isodiametric microsculpture.
</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDFA4BEE40122" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">Head transverse, about twice as broad as long, with very short and moderately deep anteocellar foveae as long as diameter of ocellus. Ocelli moderately large, located at level of posterior margins of eyes; distance between ocelli 1.2–1.4 times as long as distance between ocellus and posterior margin of eyes. Antennomeres 6–10 distinctly transverse; antennomere 3 distinctly longer and slightly narrower than 2, 5 slightly broader than 4, 6 about as long as and about wice as broad as 5, 7–8 slightly longer and broader than 6, 9 slightly broader than 8, 10 slightly longer than 9.</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDE5CBF9901EA" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">Pronotum slightly more than twice as broad as long, broader than head, widest in middle, slightly more narrowed posteriad than anteriad.</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDE84BEFE01B2" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">Elytra 1.3–1.5 times as broad as long, 1.8 times to twice as long as pronotum, with narrowly impressed lateral margins.</paragraph>
<caption id="7D8A6613FFCA6D7BD6DDDC8CBD1B0058" ID-DOI="http://doi.org/10.5281/zenodo.10821267" ID-Zenodo-Dep="10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286" startId="35.[151,264,1467,1491]" targetBox="[160,1427,159,1442]" targetPageId="35" targetType="figure">
<paragraph id="294A369BFFCA6D7BD6DDDC8CBD1B0058" blockId="35.[151,1436,1467,1564]" pageId="35" pageNumber="286">
<emphasis id="1B81EA89FFCA6D7BD6DDDC8CBF630397" bold="true" box="[151,356,1467,1492]" pageId="35" pageNumber="286">FIGURES 97–103.</emphasis>
<taxonomicName id="EEF54D18FFCA6D7BD726DC8BBC960397" authorityName="Shavrin" authorityYear="2024" box="[364,657,1468,1492]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="penelopeae">
<emphasis id="1B81EA89FFCA6D7BD726DC8BBC960397" box="[364,657,1468,1492]" italics="true" pageId="35" pageNumber="286">Paraphloeostiba penelopeae</emphasis>
</taxonomicName>
(paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFCA6D7BD68DDF68BD23003A" blockId="35.[151,1437,1631,2042]" box="[199,804,1631,1658]" pageId="35" pageNumber="286">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFCA6D7BD68DDFB5BD23016B" blockId="35.[151,1437,1631,2042]" pageId="35" pageNumber="286">
Male. Posterior margin of abdominal tergite VIII straight (
<figureCitation id="B1CE2A1EFFCA6D7BD51BDFB5BDA200DE" box="[849,933,1666,1693]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 99</figureCitation>
). Posterior margin of abdominal sternite VIII widely concave (
<figureCitation id="B1CE2A1EFFCA6D7BD71DDF92BFBD0083" box="[343,442,1701,1728]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 100</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward wide median lobe, with widely rounded apex; parameres slightly exceeding apex of median lobe, with broadened apical portions, each with two long apical and four preapical setae; internal sac narrow and moderately long, with elongate structure in basal part (
<figureCitation id="B1CE2A1EFFCA6D7BD698DE39BF21016B" box="[210,294,1806,1832]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 97</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFCA6D7BD487DE39BD27016B" box="[717,800,1806,1832]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 98</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFCA6D7BD68DDE06BCB801D2" blockId="35.[151,1437,1631,2042]" pageId="35" pageNumber="286">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFCA6D7BD53FDE06BDD00108" box="[885,983,1841,1867]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 101</figureCitation>
). Posterior margin of abdominal sternite VIII widely rounded (
<figureCitation id="B1CE2A1EFFCA6D7BD7D8DE63BFF1012D" box="[402,502,1876,1902]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 102</figureCitation>
). Accessory sclerite with wide basal portion, gradually narrowed toward long and narrow apical part with subacute apex (
<figureCitation id="B1CE2A1EFFCA6D7BD41ADE40BCB501D2" box="[592,690,1911,1937]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 103</figureCitation>
).
</paragraph>
<paragraph id="294A369BFFCA6D7BD68DDEADBA1E01B9" blockId="35.[151,1437,1631,2042]" pageId="35" pageNumber="286">
<emphasis id="1B81EA89FFCA6D7BD68DDEADBFB701F7" bold="true" box="[199,432,1946,1972]" pageId="35" pageNumber="286">Comparative notes.</emphasis>
Based on the very broad median lobe and apical portions of the parameres,
<taxonomicName id="EEF54D18FFCA6D7BD2B4DEACBB9A01F7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1278,1437,1946,1972]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFCA6D7BD2B4DEACBB0801F7" box="[1278,1295,1947,1972]" italics="true" pageId="35" pageNumber="286">P</emphasis>
.
<emphasis id="1B81EA89FFCA6D7BD357DEADBB9A01F7" box="[1309,1437,1946,1972]" italics="true" pageId="35" pageNumber="286">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCA6D7BD6DDDE8ABED00194" bold="true" box="[151,215,1981,2007]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="00B257F2FFCA6D7BD6DDDE8ABED00194" box="[151,215,1981,2007]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFCA6D7BD720DE89BFE50194" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[362,482,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFCA6D7BD720DE89BFE50194" box="[362,482,1981,2007]" italics="true" pageId="35" pageNumber="286">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCA6D7BD7A3DE8ABC2F0194" bold="true" box="[489,552,1981,2007]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="00B257F2FFCA6D7BD7A3DE8ABC2F0194" box="[489,552,1981,2007]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFCA6D7BD47CDE89BD490194" authority="Steel, 1960" authorityName="Steel" authorityYear="1960" box="[566,846,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFCA6D7BD47CDE89BCC20194" box="[566,709,1981,2007]" italics="true" pageId="35" pageNumber="286">P. celebensis</emphasis>
<bibRefCitation id="4D644B6AFFCA6D7BD486DE8ABD490194" author="Steel, W. O." box="[716,846,1981,2007]" pageId="35" pageNumber="286" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel, 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFCA6D7BD513DE89BDD00194" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[857,983,1982,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFCA6D7BD513DE89BDD00194" box="[857,983,1982,2007]" italics="true" pageId="35" pageNumber="286">P. papuana</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFCA6D7BD5AFDE89BA420194" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[997,1093,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFCA6D7BD5AFDE89BA420194" box="[997,1093,1981,2007]" italics="true" pageId="35" pageNumber="286">P. rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCA6D7BD207DE8ABA8B0194" bold="true" box="[1101,1164,1981,2007]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="00B257F2FFCA6D7BD207DE8ABA8B0194" box="[1101,1164,1981,2007]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFCA6D7BD289DE89BB480194" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[1219,1359,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFCA6D7BD289DE89BB480194" box="[1219,1359,1981,2007]" italics="true" pageId="35" pageNumber="286">P. tonkinesis</emphasis>
</taxonomicName>
. It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD9A0BCE206F2" blockId="36.[151,1436,151,609]" box="[199,741,151,177]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD9A0BF7B06F2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,151,177]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFCD6D7CD74FD9A0BF7B06F2" box="[261,380,151,177]" italics="true" pageId="36" pageNumber="287">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD7C9D9A0BFC506F2" bold="true" box="[387,450,151,177]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD7C9D9A0BFC506F2" box="[387,450,151,177]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the shorter parameres;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD98CBA340696" blockId="36.[151,1436,151,609]" box="[199,1075,187,213]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD98CBF940696" authorityName="Steel" authorityYear="1960" box="[261,403,187,213]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFCD6D7CD74FD98CBF940696" box="[261,403,187,213]" italics="true" pageId="36" pageNumber="287">P. celebensis</emphasis>
</taxonomicName>
by slightly longer parameres with narrower apical portions;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD9E8BD4006BA" blockId="36.[151,1436,151,609]" box="[199,839,223,249]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD9E8BF8506BB" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[261,386,223,248]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFCD6D7CD74FD9E8BF8506BB" box="[261,386,223,248]" italics="true" pageId="36" pageNumber="287">P. papuana</emphasis>
</taxonomicName>
by slightly longer elytra and parameres;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD834BB60075E" blockId="36.[151,1436,151,609]" box="[199,1383,259,285]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD834BF63075E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,356,259,285]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFCD6D7CD74FD834BF63075E" box="[261,356,259,285]" italics="true" pageId="36" pageNumber="287">P. rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD726D834BFAC075E" bold="true" box="[364,427,259,285]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD726D834BFAC075E" box="[364,427,259,285]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller body, the darker head, pronotum and abdomen, and longer parameres;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD810BDCD0702" blockId="36.[151,1436,151,609]" box="[199,970,295,321]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD810BF970702" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[261,400,295,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFCD6D7CD74FD810BF970702" box="[261,400,295,321]" italics="true" pageId="36" pageNumber="287">P. tonkinesis</emphasis>
</taxonomicName>
by the narrower apical portion of the median lobe.
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD87CBC1407EE" blockId="36.[151,1436,151,609]" pageId="36" pageNumber="287">
From all these species
<taxonomicName id="EEF54D18FFCD6D7CD7F4D87CBC5F0726" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[446,600,331,357]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFCD6D7CD7F4D87CBC5F0726" box="[446,600,331,357]" italics="true" pageId="36" pageNumber="287">P. penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD417D87CBC9D0726" bold="true" box="[605,666,331,357]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD417D87CBC9D0726" box="[605,666,331,357]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs by the absence of the punctation on the head and the pronotum, the presence of distinct longitudinal impressions of the more transverse pronotum, and details of the external and internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCD6D7CD68DD881BF5D07B6" pageId="36" pageNumber="287" type="distribution">
<paragraph id="294A369BFFCD6D7CD68DD881BF5D07B6" blockId="36.[151,1436,151,609]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD68DD881BF660793" bold="true" box="[199,353,438,464]" pageId="36" pageNumber="287">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFCD6D7CD722D880BCAC0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[360,683,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCD6D7CD722D880BCAC0792" box="[360,683,439,465]" italics="true" pageId="36" pageNumber="287">Paraphloeostiba penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD4F8D880BCF40792" bold="true" box="[690,755,439,465]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD4F8D880BCF40792" box="[690,755,439,465]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFCD6D7CD241D880BA3C0792" box="[1035,1083,439,465]" pageId="36" pageNumber="287">type</typeStatus>
locality in
<collectingRegion id="EB31F879FFCD6D7CD2F7D880BB4E0792" box="[1213,1353,439,465]" country="Solomon Islands" name="Guadalcanal" pageId="36" pageNumber="287">Guadalcanal</collectingRegion>
Island,
<collectingCountry id="51E2760BFFCD6D7CD6DDD8ECBF5107B6" box="[151,342,475,501]" name="Solomon Islands" pageId="36" pageNumber="287">Solomon Islands</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCD6D7CD68DD8C8BD300422" pageId="36" pageNumber="287" type="etymology">
<paragraph id="294A369BFFCD6D7CD68DD8C8BB3F047E" blockId="36.[151,1436,151,609]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD68DD8C8BF49045A" bold="true" box="[199,334,511,537]" pageId="36" pageNumber="287">Etymology.</emphasis>
Patronymic, the species is named to honour of entomologist and naturalist Penelope Jean Macleod Greenslade, wife of P.J.M. Greenslade, who collected material on the
<collectingCountry id="51E2760BFFCD6D7CD5EFDB14BA67047E" box="[933,1120,547,573]" name="Solomon Islands" pageId="36" pageNumber="287">Solomon Islands</collectingCountry>
during 1960-1970.
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDB71BD300422" blockId="36.[151,1436,151,609]" box="[199,823,582,609]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD68DDB71BF4C0423" bold="true" box="[199,331,582,608]" pageId="36" pageNumber="287">Bionomics.</emphasis>
The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="1B81EA89FFCD6D7CD6DDDB9DBFF20486" bold="true" box="[151,501,682,709]" pageId="36" pageNumber="287">
<taxonomicName id="EEF54D18FFCD6D7CD6DDDB9DBFAB0486" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,428,682,709]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="pilosa" status="sp. nov.">
<emphasis id="1B81EA89FFCD6D7CD6DDDB9DBFAB0486" bold="true" box="[151,428,682,709]" italics="true" pageId="36" pageNumber="287">Paraphloeostiba pilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFCD6D7CD7F9DB9CBFF20486" box="[435,501,683,709]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCD6D7CD6DDDBF8BF6B04AA" box="[151,364,719,745]" pageId="36" pageNumber="287" type="description">
<paragraph id="294A369BFFCD6D7CD6DDDBF8BF6B04AA" blockId="36.[151,501,682,745]" box="[151,364,719,745]" pageId="36" pageNumber="287">
(
<figureCitation id="B1CE2A1EFFCD6D7CD6D5DBF8BEF104AA" box="[159,246,719,745]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="36" pageNumber="287">Figs 77</figureCitation>
,
<figureCitation id="B1CE2A1EFFCD6D7CD748DBF8BF6204AA" box="[258,357,719,745]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="36" pageNumber="287">104–110</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCD6D7CD6DDDA20BFAD024A" pageId="36" pageNumber="287" type="materials_examined">
<paragraph id="294A369BFFCD6D7CD6DDDA20BA1A053A" blockId="36.[151,1437,790,2041]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD6DDDA20BF420573" bold="true" box="[151,325,790,817]" pageId="36" pageNumber="287">Type material.</emphasis>
<typeStatus id="F64E8839FFCD6D7CD706DA20BFB30572" box="[332,436,791,817]" pageId="36" pageNumber="287" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFCD6D7CD7F1DA20BFD70572" box="[443,464,791,817]" count="1" pageId="36" pageNumber="287" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFCD6D7CD795DA20BC350572" box="[479,562,791,817]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="36" pageNumber="287">Fig. 77</figureCitation>
; left antennomeres 3–11 glued on the same card under the specimen): ‘
<collectingRegion id="EB31F879FFCD6D7CD303DA20BED90516" country="Indonesia" name="Papua Barat" pageId="36" pageNumber="287">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Sinak, trail to Ilaga |
<quantity id="EE0D9B7EFFCD6D7CD434DA0DBD170516" box="[638,784,826,853]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="36" pageNumber="287" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100-2500m</quantity>
,
<date id="5D4B105BFFCD6D7CD557DA0DBDAA0516" box="[797,941,826,853]" pageId="36" pageNumber="287" value="1995-12-15">15. 12. 1995</date>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFCD6D7CD2BFDA0CBB890516" box="[1269,1422,827,853]" pageId="36" pageNumber="287" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFCD6D7CD6DDDA68BF55053A" authorityName="Steel" authorityYear="1960" box="[151,338,863,889]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFCD6D7CD6DDDA68BF55053A" box="[151,338,863,889]" italics="true" pageId="36" pageNumber="287">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFCD6D7CD72FDA68BFAE053A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[357,425,863,889]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="pilosa" status="sp. nov.">
<emphasis id="1B81EA89FFCD6D7CD72FDA68BFAE053A" box="[357,425,863,889]" italics="true" pageId="36" pageNumber="287">pilosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD7FBDA68BFF7053A" bold="true" box="[433,496,863,889]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD7FBDA68BFF7053A" box="[433,496,863,889]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFCD6D7CD430DA68BC97053A" box="[634,656,863,889]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="36" pageNumber="287" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFCD6D7CD589DA68BA16053A" box="[963,1041,863,889]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="36" pageNumber="287">NMW</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDAB4BFAD024A" blockId="36.[151,1437,790,2041]" pageId="36" pageNumber="287">
<typeStatus id="F64E8839FFCD6D7CD68DDAB4BF3F05DE" box="[199,312,899,925]" pageId="36" pageNumber="287" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFCD6D7CD709DAB4BF6B05DE" box="[323,364,899,925]" count="1" pageId="36" pageNumber="287" type="male">1 ♂</specimenCount>
(dissected): same data as the holotype (
<collectionCode id="4FE4AE5EFFCD6D7CD563DAB4BD7005DE" box="[809,887,899,925]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="36" pageNumber="287">NMW</collectionCode>
);
<specimenCount id="3FF3FD12FFCD6D7CD5C0DAB4BDCF05DE" box="[906,968,899,925]" count="4" pageId="36" pageNumber="287" type="female">4 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFCD6D7CD59DDAB4BA7505DE" box="[983,1138,899,925]" count="1" pageId="36" pageNumber="287" type="generic">one specimen</specimenCount>
without left antennomeres 3–11;
<specimenCount id="3FF3FD12FFCD6D7CD694DA90BF7F0582" box="[222,376,935,961]" count="1" pageId="36" pageNumber="287" type="generic">one specimen</specimenCount>
dissected): ‘
<collectionCode id="4FE4AE5EFFCD6D7CD44EDA90BC4F0582" box="[516,584,935,961]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="36" pageNumber="287">NEW</collectionCode>
<collectingCountry id="51E2760BFFCD6D7CD41BDA90BCBC0582" box="[593,699,935,961]" name="Guinea-Bissau" pageId="36" pageNumber="287">GUINEA</collectingCountry>
| Okasa, |
<date id="5D4B105BFFCD6D7CD579DA90BD9B0582" box="[819,924,935,961]" pageId="36" pageNumber="287" value="1966-02-07">7.II.1966</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFCD6D7CD266DA90BA420582" box="[1068,1093,935,961]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="36" pageNumber="287">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFCD6D7CD75FDAFCBF6805A6" box="[277,367,971,997]" pageId="36" pageNumber="287">BMNH</collectionCode>
). All
<typeStatus id="F64E8839FFCD6D7CD7E0DAFCBC1205A6" box="[426,533,971,997]" pageId="36" pageNumber="287" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFCD6D7CD5DFDAFCBA2205A6" box="[917,1061,971,997]" pageId="36" pageNumber="287" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFCD6D7CD272DAFCBAF405A6" authorityName="Steel" authorityYear="1960" box="[1080,1267,971,997]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFCD6D7CD272DAFCBAF405A6" box="[1080,1267,971,997]" italics="true" pageId="36" pageNumber="287">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFCD6D7CD34CDAFCBB4D05A6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1286,1354,971,997]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="pilosa" status="sp. nov.">
<emphasis id="1B81EA89FFCD6D7CD34CDAFCBB4D05A6" box="[1286,1354,971,997]" italics="true" pageId="36" pageNumber="287">pilosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD31BDAFCBB9705A6" bold="true" box="[1361,1424,971,997]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD31BDAFCBB9705A6" box="[1361,1424,971,997]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFCD6D7CD747DAD8BF24024A" box="[269,291,1007,1033]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="36" pageNumber="287" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCD6D7DD68DDD25BFB0047E" lastPageId="37" lastPageNumber="288" pageId="36" pageNumber="287" type="description">
<paragraph id="294A369BFFCD6D7CD68DDD25BF1402DA" blockId="36.[151,1437,790,2041]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD68DDD25BF5E026F" bold="true" box="[199,345,1042,1068]" pageId="36" pageNumber="287">Description.</emphasis>
Measurements (n=6): HW: 0.42–0.49; HL: 0.25–0.27; OL: 0.12; TL: 0.02–0.03; AL (
<typeStatus id="F64E8839FFCD6D7CD362DD24BB96026E" box="[1320,1425,1043,1069]" pageId="36" pageNumber="287" type="holotype">holotype</typeStatus>
): 0.63; PL: 0.35–0.40; PWmax: 0.57–0.60; PWmin: 0.53–0.56; ESL: 0.66–0.74; EW: 0.73–0.88; MTbL (
<typeStatus id="F64E8839FFCD6D7CD362DD00BB960212" box="[1320,1425,1079,1105]" pageId="36" pageNumber="287" type="holotype">holotype</typeStatus>
): 0.40; MTrL (
<typeStatus id="F64E8839FFCD6D7CD77FDD6CBF990236" box="[309,414,1115,1141]" pageId="36" pageNumber="287" type="holotype">holotype</typeStatus>
): 0.24 (MTrL 1–4: 0.08; MTrL 5: 0.16); AW: 0.73–0.90; AedL: 0.47; BL: 1.95–2.65 (
<typeStatus id="F64E8839FFCD6D7CD6D5DD48BF0F02DA" box="[159,264,1151,1177]" pageId="36" pageNumber="287" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDD94BA4A03FA" blockId="36.[151,1437,790,2041]" pageId="36" pageNumber="287">
Habitus as in
<figureCitation id="B1CE2A1EFFCD6D7CD721DD94BFC702FE" box="[363,448,1187,1213]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="36" pageNumber="287">Fig. 77</figureCitation>
. Body dark-brown, with indistinctly paler elytra; antennomeres 6–11 or 7–11 brown; mouthparts and antennomeres 1–5 or 1–6 yellowish; legs reddish, with slightly paler tarsi. Head, neck, pronotum and scutellum without punctation; elytra with very dense, moderartely large and deep punctation, denser around scutellum and sparser along suture; abdominal tergites with indistinct, sparse and fine punctation, more visible on tergites III–V. Head with very dense, coarse, small microsculpture; pronotum with very dense isodiametric microsculpture, slightly coarser than that in middle portion of head; scutellum with dense and coarse transverse meshes; microsculpture of elytra moderartely fine, longitudinal or isodiametric, indistinct or absent in middle portions of each elytron, denser and caorser in mediobasal and medioapical portions.
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDCF4BDF200F6" blockId="36.[151,1437,790,2041]" pageId="36" pageNumber="287">Head slightly elevated in middle, with distinct, shallow or moderately deep oval anteriomedian depressions, 1.6–1.8 times as broad as long, with narrow and short anteocellar foveae, diagonally stretching anteriad; postocular carina distinct, acute. Ocelli large, located slightly below level of postocular carina; distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Apical maxillary palpomere 1.3–1.5 times as long as preapical segment. Antennomeres 7–10 slightly transverse; antennomere 3 disitnctly longer and narrower than 2, 5 slightly broader than 4, 6 slightly narrower than 5, 7 slightly longer and distinctly broader than 6, 8 distinctly longer and broader than 7, 9–10 slightly longer and broader than 8.</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDF88BCB900BE" blockId="36.[151,1437,790,2041]" pageId="36" pageNumber="287">Pronotum distinctly convex, 1.5–1.6 times as broad as long, 1.2–1.3 times as broad as head, from widest middle distinctly more narrowed posteriad than anteriad.</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDE30BC310106" blockId="36.[151,1437,790,2041]" pageId="36" pageNumber="287">Elytra 1.1 times as broad as long; surface of each elytron with longitudinal elevations between punctures and diagonal elevations in apical portion.</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDE78BD23012A" blockId="36.[151,1437,790,2041]" box="[199,804,1871,1897]" pageId="36" pageNumber="287">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDE44BC4A01CE" blockId="36.[151,1437,790,2041]" box="[199,589,1907,1933]" pageId="36" pageNumber="287">Abdomen about as broad as elytra.</paragraph>
<paragraph id="294A369BFFCD6D7DD68DDEA0BF3006BA" blockId="36.[151,1437,790,2041]" lastBlockId="37.[151,1437,151,573]" lastPageId="37" lastPageNumber="288" pageId="36" pageNumber="287">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFCD6D7CD521DEA0BDC801F2" box="[875,975,1943,1969]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="36" pageNumber="287">Fig. 106</figureCitation>
). Posterior margin of abdominal sternite VIII widely concave (
<figureCitation id="B1CE2A1EFFCD6D7CD7C5DE8CBFF50196" box="[399,498,1979,2005]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="36" pageNumber="287">Fig. 107</figureCitation>
). Aedeagus with very wide basal portion, gradually narrowed toward rounded apex of median lobe; parameres significantly exceeding apex of median lobe, from widest preapical portions slightly narrowed toward rounded apices, with numerous very long apical and preapical setae; internal sac moderately narrow and long, with two elongate and sclerotized structures in basal part (
<figureCitation id="B1CE2A1EFFCC6D7DD591D98CBA3A0696" box="[987,1085,187,213]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="37" pageNumber="288">Fig. 104</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFCC6D7DD698D9E8BF3406BA" box="[210,307,222,249]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="37" pageNumber="288">Fig. 105</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFCC6D7DD68DD834BFAB0726" blockId="37.[151,1437,151,573]" pageId="37" pageNumber="288">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFCC6D7DD53FD834BDDF075E" box="[885,984,259,285]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="37" pageNumber="288">Fig. 108</figureCitation>
). Posterior margin of abdominal sternite VIII rounded (
<figureCitation id="B1CE2A1EFFCC6D7DD70AD810BFA20702" box="[320,421,295,321]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="37" pageNumber="288">Fig. 109</figureCitation>
). Accessory sclerite short, from widest basal portion gradually narrowed toward widely rounded apex (
<figureCitation id="B1CE2A1EFFCC6D7DD774D87CBF980726" box="[318,415,331,357]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="37" pageNumber="288">Fig. 110</figureCitation>
).
</paragraph>
<paragraph id="294A369BFFCC6D7DD68DD859BFB0047E" blockId="37.[151,1437,151,573]" pageId="37" pageNumber="288">
<emphasis id="1B81EA89FFCC6D7DD68DD859BFB407CA" bold="true" box="[199,435,366,393]" pageId="37" pageNumber="288">Comparative notes.</emphasis>
Based on the general shape of the aedeagus, with the parameres distinctly longer than apex of the median lobe with very long apical and preapical setation,
<taxonomicName id="EEF54D18FFCC6D7DD588D8A4BA2307EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[962,1060,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="288" phylum="Arthropoda" rank="species" species="pilosa" status="sp. n.">
<emphasis id="1B81EA89FFCC6D7DD588D8A4BA2307EE" box="[962,1060,403,429]" italics="true" pageId="37" pageNumber="288">P. pilosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCC6D7DD264D8A4BA7707EE" bold="true" box="[1070,1136,403,429]" pageId="37" pageNumber="288">
<taxonomicNameLabel id="00B257F2FFCC6D7DD264D8A4BA7707EE" box="[1070,1136,403,429]" pageId="37" pageNumber="288" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFCC6D7DD347D8A4BF380792" authority="(Fauvel, 1904)" baseAuthorityName="Fauvel" baseAuthorityYear="1904" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="288" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="1B81EA89FFCC6D7DD347D8A4BB9A07EE" box="[1293,1437,403,429]" italics="true" pageId="37" pageNumber="288">P. sulcicollis</emphasis>
(Fauvel, 1904)
</taxonomicName>
from Sumatra and
<collectingCountry id="51E2760BFFCC6D7DD46BD880BCA00792" box="[545,679,439,465]" name="Hong Kong" pageId="37" pageNumber="288">Hong Kong</collectingCountry>
(see Shavrin 2020). It can be distinguished from
<taxonomicName id="EEF54D18FFCC6D7DD2ADD880BB700792" baseAuthorityName="Fauvel" baseAuthorityYear="1904" box="[1255,1399,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="288" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="1B81EA89FFCC6D7DD2ADD880BB700792" box="[1255,1399,439,465]" italics="true" pageId="37" pageNumber="288">P. sulcicollis</emphasis>
</taxonomicName>
by the presence of the punctation of the head, more convex pronotum without distinct longitudinal impressions, the broader aedeagus with narrower apical portions of the median lobe and parameres, and the different shape of the female accessory sclerite.
</paragraph>
</subSubSection>
<caption id="7D8A6613FFCC6D7DD6DDDEAEBD1C01BA" ID-DOI="http://doi.org/10.5281/zenodo.10821269" ID-Zenodo-Dep="10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="37" pageNumber="288" startId="37.[151,264,1945,1969]" targetBox="[159,1428,605,1914]" targetPageId="37" targetType="figure">
<paragraph id="294A369BFFCC6D7DD6DDDEAEBD1C01BA" blockId="37.[151,1436,1945,2042]" pageId="37" pageNumber="288">
<emphasis id="1B81EA89FFCC6D7DD6DDDEAEBF6C01F2" bold="true" box="[151,363,1945,1969]" pageId="37" pageNumber="288">FIGURES 104–110.</emphasis>
<taxonomicName id="EEF54D18FFCC6D7DD725DEADBC5C01F1" authorityName="Shavrin" authorityYear="2024" box="[367,603,1946,1970]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="288" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="1B81EA89FFCC6D7DD725DEADBC5C01F1" box="[367,603,1946,1970]" italics="true" pageId="37" pageNumber="288">Paraphloeostiba pilosa</emphasis>
</taxonomicName>
: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="61EF6510FFCF6D7ED68DD9A1BC4C0697" pageId="38" pageNumber="289" type="distribution">
<paragraph id="294A369BFFCF6D7ED68DD9A1BC4C0697" blockId="38.[151,1437,150,357]" pageId="38" pageNumber="289">
<emphasis id="1B81EA89FFCF6D7ED68DD9A1BF6606F3" bold="true" box="[199,353,150,176]" pageId="38" pageNumber="289">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFCF6D7ED722D9A1BC6906F3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[360,622,150,176]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="289" phylum="Arthropoda" rank="species" species="pilosa" status="sp. nov.">
<emphasis id="1B81EA89FFCF6D7ED722D9A1BC6906F3" box="[360,622,150,176]" italics="true" pageId="38" pageNumber="289">Paraphloeostiba pilosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCF6D7ED43CD9A1BCB106F3" bold="true" box="[630,694,150,176]" pageId="38" pageNumber="289">
<taxonomicNameLabel id="00B257F2FFCF6D7ED43CD9A1BCB106F3" box="[630,694,150,176]" pageId="38" pageNumber="289" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in Western New
<collectingCountry id="51E2760BFFCF6D7ED28ED9A1BB1206F3" box="[1220,1301,150,176]" name="Guinea-Bissau" pageId="38" pageNumber="289">Guinea</collectingCountry>
(
<collectingCountry id="51E2760BFFCF6D7ED36ED9A1BB9306F3" box="[1316,1428,150,176]" name="Indonesia" pageId="38" pageNumber="289">Indonesia</collectingCountry>
) and central part of
<collectingCountry id="51E2760BFFCF6D7ED721D98CBC400697" box="[363,583,186,213]" name="Papua New Guinea" pageId="38" pageNumber="289">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCF6D7ED68DD9E9BF370727" pageId="38" pageNumber="289" type="etymology">
<paragraph id="294A369BFFCF6D7ED68DD9E9BFBF075E" blockId="38.[151,1437,150,357]" pageId="38" pageNumber="289">
<emphasis id="1B81EA89FFCF6D7ED68DD9E9BF4906BB" bold="true" box="[199,334,222,248]" pageId="38" pageNumber="289">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFCF6D7ED487D9E9BD1B06BB" box="[717,796,222,248]" italics="true" pageId="38" pageNumber="289">pilosus</emphasis>
, -
<emphasis id="1B81EA89FFCF6D7ED57CD9E8BD4306BB" box="[822,836,223,248]" italics="true" pageId="38" pageNumber="289">a</emphasis>
, -
<emphasis id="1B81EA89FFCF6D7ED514D9E8BD8706BB" box="[862,896,223,248]" italics="true" pageId="38" pageNumber="289">um</emphasis>
(hairy). It alludes to the presence of very long setation of the parameres.
</paragraph>
<paragraph id="294A369BFFCF6D7ED68DD811BF370727" blockId="38.[151,1437,150,357]" pageId="38" pageNumber="289">
<emphasis id="1B81EA89FFCF6D7ED68DD811BF4C0703" bold="true" box="[199,331,294,320]" pageId="38" pageNumber="289">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="EE0D9B7EFFCF6D7ED512D811BA0A0702" box="[856,1037,294,321]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="38" pageNumber="289" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

373
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<paragraph id="294A369BFFCF6D7ED6DDD881BCC80793" blockId="38.[151,719,438,500]" box="[151,719,438,465]" pageId="38" pageNumber="289">
<heading id="720281F7FFCF6D7ED6DDD881BCC80793" bold="true" box="[151,719,438,465]" fontSize="11" level="1" pageId="38" pageNumber="289" reason="1">
<taxonomicName id="EEF54D18FFCF6D7ED6DDD881BCC80793" ID-CoL="7639P" authority="(Cameron, 1928)" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[151,719,438,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="289" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFCF6D7ED6DDD881BCC80793" bold="true" box="[151,719,438,465]" pageId="38" pageNumber="289">
<emphasis id="1B81EA89FFCF6D7ED6DDD881BFF00792" bold="true" box="[151,503,438,465]" italics="true" pageId="38" pageNumber="289">Paraphloeostiba rhopalocera</emphasis>
(
<bibRefCitation id="4D644B6AFFCF6D7ED44CD881BCC00793" author="Cameron, M." box="[518,711,438,464]" pageId="38" pageNumber="289" pagination="90 - 110" refId="ref35375" refString="Cameron, M. (1928) Fauna sumatrensis. Staphylinidae (Col.). Entomologische Mitteilungen, 17 (2), 90 - 110." type="journal article" year="1928">Cameron, 1928</bibRefCitation>
)
</emphasis>
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<subSubSection id="61EF6510FFCF6D7ED6DDD8EDBF6D07B7" box="[151,362,474,500]" pageId="38" pageNumber="289" type="description">
<paragraph id="294A369BFFCF6D7ED6DDD8EDBF6D07B7" blockId="38.[151,719,438,500]" box="[151,362,474,500]" pageId="38" pageNumber="289">
(
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,
<figureCitation id="B1CE2A1EFFCF6D7ED748D8EDBF6507B7" box="[258,354,474,500]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">111–118</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCF6D7ED6DDDB28BC770437" pageId="38" pageNumber="289" type="reference_group">
<paragraph id="294A369BFFCF6D7ED6DDDB28BC700474" blockId="38.[151,844,543,628]" box="[151,631,543,567]" pageId="38" pageNumber="289">
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<treatmentCitation id="A854108AFFCF6D7ED6DDDB28BC700474" author="Cameron, M." box="[151,631,543,567]" page="99" pageId="38" pageNumber="289" year="1928">
<taxonomicName id="EEF54D18FFCF6D7ED6DDDB28BC700474" ID-CoL="4GGF2" authority="Cameron, 1928: 99" authorityName="Cameron" authorityPageNumber="99" authorityYear="1928" box="[151,631,543,567]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="289" phylum="Arthropoda" rank="species" species="rhopalocerus">
<emphasis id="1B81EA89FFCF6D7ED6DDDB28BFA90474" box="[151,430,543,567]" italics="true" pageId="38" pageNumber="289">Phloeonomus rhopalocerus</emphasis>
<bibRefCitation id="4D644B6AFFCF6D7ED7FEDB28BC700474" author="Cameron, M." box="[436,631,543,567]" pageId="38" pageNumber="289" pagination="90 - 110" refId="ref35375" refString="Cameron, M. (1928) Fauna sumatrensis. Staphylinidae (Col.). Entomologische Mitteilungen, 17 (2), 90 - 110." type="journal article" year="1928">Cameron, 1928: 99</bibRefCitation>
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<paragraph id="294A369BFFCF6D7ED6DDDB0ABD4B0416" blockId="38.[151,844,543,628]" box="[151,844,573,597]" pageId="38" pageNumber="289">
<treatmentCitationGroup id="09E511B5FFCF6D7ED6DDDB0ABD4B0416" box="[151,844,573,597]" pageId="38" pageNumber="289">
<taxonomicName id="EEF54D18FFCF6D7ED6DDDB0ABC570416" ID-CoL="4GGF2" authorityName="Cameron" authorityYear="1928" box="[151,592,573,597]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="289" phylum="Arthropoda" rank="species" species="rhopalocerus" subGenus="Phloeonomus">
<emphasis id="1B81EA89FFCF6D7ED6DDDB0ABF260416" box="[151,289,573,597]" italics="true" pageId="38" pageNumber="289">Phloeonomus</emphasis>
(
<emphasis id="1B81EA89FFCF6D7ED77ADB0ABFBD0416" box="[304,442,573,597]" italics="true" pageId="38" pageNumber="289">Phloeonomus</emphasis>
)
<emphasis id="1B81EA89FFCF6D7ED783DB0ABC570416" box="[457,592,573,597]" italics="true" pageId="38" pageNumber="289">rhopalocerus</emphasis>
</taxonomicName>
:
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<bibRefCitation id="4D644B6AFFCF6D7ED417DB0ABD4B0416" author="Scheerpeltz, O. von" box="[605,844,573,597]" pageId="38" pageNumber="289" pagination="989 - 1500" refId="ref35653" refString="Scheerpeltz, O. von (1933) Staphylinidae VII (pars 129). In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. VI. Staphylinidae II. W. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz 1933: 1050</bibRefCitation>
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</paragraph>
<paragraph id="294A369BFFCF6D7ED6DDDB6BBC770437" blockId="38.[151,844,543,628]" box="[151,624,604,628]" pageId="38" pageNumber="289">
<treatmentCitationGroup id="09E511B5FFCF6D7ED6DDDB6BBC770437" box="[151,624,604,628]" pageId="38" pageNumber="289">
<taxonomicName id="EEF54D18FFCF6D7ED6DDDB6BBFC40437" ID-CoL="7639P" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[151,451,604,628]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="289" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFCF6D7ED6DDDB6BBFC40437" box="[151,451,604,628]" italics="true" pageId="38" pageNumber="289">Paraphloeostiba rhopalocera</emphasis>
</taxonomicName>
:
<treatmentCitation id="A854108AFFCF6D7ED79BDB6BBC770437" author="Steel, W. O." box="[465,624,604,628]" page="146" pageId="38" pageNumber="289" year="1960">
<bibRefCitation id="4D644B6AFFCF6D7ED79BDB6BBC770437" author="Steel, W. O." box="[465,624,604,628]" pageId="38" pageNumber="289" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960: 146</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCF6D7ED6DDDB95BC76052C" pageId="38" pageNumber="289" type="materials_examined">
<paragraph id="294A369BFFCF6D7ED6DDDB95BC76052C" blockId="38.[151,1437,674,1987]" pageId="38" pageNumber="289">
<emphasis id="1B81EA89FFCF6D7ED6DDDB95BF8704FF" bold="true" box="[151,384,674,700]" pageId="38" pageNumber="289">Material examined.</emphasis>
<materialsCitation id="999D3CC6FFCF6D7ED7CFDB95BDEF04FF" box="[389,1000,674,700]" collectionCode="NEW, GUINEA" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="38" pageNumber="289" specimenCount="5" specimenCount-female="4" specimenCount-male="1">
<emphasis id="1B81EA89FFCF6D7ED7CFDB95BCAF04FF" bold="true" box="[389,680,674,700]" pageId="38" pageNumber="289">
<collectingCountry id="51E2760BFFCF6D7ED7CFDB95BCA404FF" box="[389,675,674,700]" name="Papua New Guinea" pageId="38" pageNumber="289">PAPUA NEW GUINEA</collectingCountry>
:
</emphasis>
<specimenCount id="3FF3FD12FFCF6D7ED4E4DB95BCDE04FF" box="[686,729,674,700]" count="1" pageId="38" pageNumber="289" type="male">1 ♂</specimenCount>
,
<specimenCount id="3FF3FD12FFCF6D7ED4AEDB95BD2304FF" box="[740,804,674,700]" count="4" pageId="38" pageNumber="289" type="female">4 ♀♀</specimenCount>
: ‘
<collectionCode id="4FE4AE5EFFCF6D7ED57EDB95BD7F04FF" box="[820,888,674,700]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="38" pageNumber="289">NEW</collectionCode>
<collectionCode id="4FE4AE5EFFCF6D7ED534DB95BDEF04FF" box="[894,1000,674,700]" pageId="38" pageNumber="289">GUINEA</collectionCode>
</materialsCitation>
<materialsCitation id="999D3CC6FFCF6D7ED5A4DB95BB0D04FF" box="[1006,1290,674,700]" collectingDate="1964-08-23" collectionCode="R" country="Papua New Guinea" location="Okapa" pageId="38" pageNumber="289" specimenCount="1">
<location id="2C2A6040FFCF6D7ED5A4DB95BA3D04FF" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED5A4DB95BA3D04FF" box="[1006,1082,674,700]" country="Papua New Guinea" name="Okapa" pageId="38" pageNumber="289">Okapa</location>
,
<date id="5D4B105BFFCF6D7ED20EDB95BADB04FF" box="[1092,1244,674,700]" pageId="38" pageNumber="289" value="1964-08-23">
<collectingDate id="4D0FE9B3FFCF6D7ED20EDB95BADB04FF" box="[1092,1244,674,700]" pageId="38" pageNumber="289" value="1964-08-23">23.VIII.1964</collectingDate>
</date>
’, ‘
<collectionCode id="4FE4AE5EFFCF6D7ED2BBDB95BB0D04FF" box="[1265,1290,674,700]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="289">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="999D3CC6FFCF6D7ED35FDB95BF7B04A3" collectionCode="R" country="Papua New Guinea" location="Hornabrook" pageId="38" pageNumber="289" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFCF6D7ED35FDB95BB9B04FF" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED35FDB95BB9B04FF" box="[1301,1436,674,700]" country="Papua New Guinea" name="Hornabrook" pageId="38" pageNumber="289">Hornabrook</location>
<specimenCode id="79539EE0FFCF6D7ED6DDDBF1BF4204A3" box="[151,325,710,736]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="38" pageNumber="289">BM 1970-232</specimenCode>
.’, ‘
<collectionCode id="4FE4AE5EFFCF6D7ED729DBF1BF7B04A3" box="[355,380,710,736]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="289">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="999D3CC6FFCF6D7ED7C7DBF1BD4E04A3" box="[397,841,710,736]" collectionCode="BMNH" country="Papua New Guinea" location="Hornabrook" pageId="38" pageNumber="289" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFCF6D7ED7C7DBF1BC1304A3" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED7C7DBF1BC1304A3" box="[397,532,710,736]" country="Papua New Guinea" name="Hornabrook" pageId="38" pageNumber="289">Hornabrook</location>
<specimenCode id="79539EE0FFCF6D7ED46BDBF1BCCD04A3" box="[545,714,710,736]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="38" pageNumber="289">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFCF6D7ED4ADDBF1BD4604A3" box="[743,833,710,736]" pageId="38" pageNumber="289">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="999D3CC6FFCF6D7ED513DBF1BA5B04A3" box="[857,1116,710,736]" collectionCode="NEW, GUINEA" country="Papua New Guinea" pageId="38" pageNumber="289" specimenCount="1" specimenCount-male="1">
<specimenCount id="3FF3FD12FFCF6D7ED513DBF1BD8B04A3" box="[857,908,710,736]" count="1" pageId="38" pageNumber="289" type="male">1 ♂</specimenCount>
: ‘
<collectionCode id="4FE4AE5EFFCF6D7ED5EBDBF1BDE204A3" box="[929,997,710,736]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="38" pageNumber="289">NEW</collectionCode>
<collectionCode id="4FE4AE5EFFCF6D7ED5B8DBF1BA5B04A3" box="[1010,1116,710,736]" pageId="38" pageNumber="289">GUINEA</collectionCode>
</materialsCitation>
<materialsCitation id="999D3CC6FFCF6D7ED222DBF1BFC00547" collectingDate="1965-07-18" collectionCode="R" country="Papua New Guinea" elevation="1829" location="Lufa" pageId="38" pageNumber="289" specimenCount="1">
<location id="2C2A6040FFCF6D7ED222DBF1BAA704A3" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED222DBF1BAA704A3" box="[1128,1184,710,736]" country="Papua New Guinea" name="Lufa" pageId="38" pageNumber="289">Lufa</location>
,
<quantity id="EE0D9B7EFFCF6D7ED2FADBF1BAFB04A3" box="[1200,1276,710,736]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="38" pageNumber="289" unit="ft" value="6000.0">
<elevation id="A2D8D1A8FFCF6D7ED2FADBF1BAFB04A3" box="[1200,1276,710,736]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="38" pageNumber="289" unit="ft" value="6000.0">6000ft</elevation>
</quantity>
,
<date id="5D4B105BFFCF6D7ED346DBF2BB9004A3" box="[1292,1431,709,736]" pageId="38" pageNumber="289" value="1965-07-18">
<collectingDate id="4D0FE9B3FFCF6D7ED346DBF2BB9004A3" box="[1292,1431,709,736]" pageId="38" pageNumber="289" value="1965-07-18">18.VII.1965</collectingDate>
</date>
, under bark of wet log’, ‘
<collectionCode id="4FE4AE5EFFCF6D7ED7E4DBDDBFC00547" box="[430,455,746,772]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="289">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="999D3CC6FFCF6D7ED79EDBDEBD820547" box="[468,901,745,772]" collectionCode="BMNH" country="Papua New Guinea" location="Hornabrook" pageId="38" pageNumber="289" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFCF6D7ED79EDBDEBC5C0540" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED79EDBDEBC5C0540" box="[468,603,745,771]" country="Papua New Guinea" name="Hornabrook" pageId="38" pageNumber="289">Hornabrook</location>
<specimenCode id="79539EE0FFCF6D7ED42EDBDDBD0D0547" box="[612,778,746,772]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="38" pageNumber="289">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFCF6D7ED569DBDDBD7A0547" box="[803,893,746,772]" pageId="38" pageNumber="289">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="999D3CC6FFCF6D7ED5D8DBDDBA8D0547" box="[914,1162,746,772]" collectionCode="NEW, GUINEA" country="Papua New Guinea" pageId="38" pageNumber="289" specimenCount="1" specimenCount-male="1">
<specimenCount id="3FF3FD12FFCF6D7ED5D8DBDDBDC60547" box="[914,961,746,772]" count="1" pageId="38" pageNumber="289" type="male">1 ♂</specimenCount>
: ‘
<collectionCode id="4FE4AE5EFFCF6D7ED599DBDDBA100547" box="[979,1047,746,772]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="38" pageNumber="289">NEW</collectionCode>
<collectionCode id="4FE4AE5EFFCF6D7ED26ADBDDBA8D0547" box="[1056,1162,746,772]" pageId="38" pageNumber="289">GUINEA</collectionCode>
</materialsCitation>
<materialsCitation id="999D3CC6FFCF6D7ED2D9DBDEBB9F0547" box="[1171,1432,745,772]" collectingDate="1964-07-18" collectionCode="R" country="Papua New Guinea" location="Lufa" pageId="38" pageNumber="289" specimenCount="1">
<location id="2C2A6040FFCF6D7ED2D9DBDEBACC0540" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED2D9DBDEBACC0540" box="[1171,1227,745,771]" country="Papua New Guinea" name="Lufa" pageId="38" pageNumber="289">Lufa</location>
,
<date id="5D4B105BFFCF6D7ED29DDBDEBB600540" box="[1239,1383,745,771]" pageId="38" pageNumber="289" value="1964-07-18">
<collectingDate id="4D0FE9B3FFCF6D7ED29DDBDEBB600540" box="[1239,1383,745,771]" pageId="38" pageNumber="289" value="1964-07-18">18.VII.1964</collectingDate>
</date>
’, ‘
<collectionCode id="4FE4AE5EFFCF6D7ED335DBDDBB9F0547" box="[1407,1432,746,772]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="289">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="999D3CC6FFCF6D7ED6DDDA3ABB9F0564" box="[151,1432,781,808]" collectionCode="BMNH, NEW, GUINEA" country="Papua New Guinea" location="Hornabrook" pageId="38" pageNumber="289" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFCF6D7ED6DDDA3ABF190564" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED6DDDA3ABF190564" box="[151,286,781,807]" country="Papua New Guinea" name="Hornabrook" pageId="38" pageNumber="289">Hornabrook</location>
<specimenCode id="79539EE0FFCF6D7ED762DA39BFC80564" box="[296,463,781,808]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="38" pageNumber="289">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFCF6D7ED7A3DA3ABC440564" box="[489,579,781,807]" pageId="38" pageNumber="289">BMNH</collectionCode>
); 1 unsexed specimen (apical part of abdomen missing): ‘
<collectionCode id="4FE4AE5EFFCF6D7ED296DA3ABB270564" box="[1244,1312,781,807]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="38" pageNumber="289">NEW</collectionCode>
<collectionCode id="4FE4AE5EFFCF6D7ED360DA3ABB9F0564" box="[1322,1432,781,807]" pageId="38" pageNumber="289">GUINEA</collectionCode>
</materialsCitation>
,
<materialsCitation id="999D3CC6FFCF6D7ED6DDDA06BFBE0508" box="[151,441,817,844]" collectingDate="1964-09-15" collectionCode="R" country="Papua New Guinea" location="Amused" pageId="38" pageNumber="289" specimenCount="1">
<location id="2C2A6040FFCF6D7ED6DDDA06BEF00508" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED6DDDA06BEF00508" box="[151,247,817,843]" country="Papua New Guinea" name="Amused" pageId="38" pageNumber="289">Amused</location>
,
<date id="5D4B105BFFCF6D7ED749DA06BF8D050F" box="[259,394,817,844]" pageId="38" pageNumber="289" value="1964-09-15">
<collectingDate id="4D0FE9B3FFCF6D7ED749DA06BF8D050F" box="[259,394,817,844]" pageId="38" pageNumber="289" value="1964-09-15">15.IX.1964</collectingDate>
</date>
’, ‘
<collectionCode id="4FE4AE5EFFCF6D7ED7EADA06BFBE0508" box="[416,441,817,843]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="289">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="999D3CC6FFCF6D7ED78CDA06BD730508" box="[454,884,817,843]" collectionCode="BMNH" country="Papua New Guinea" location="Hornabrook" pageId="38" pageNumber="289" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFCF6D7ED78CDA06BC4A0508" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED78CDA06BC4A0508" box="[454,589,817,843]" country="Papua New Guinea" name="Hornabrook" pageId="38" pageNumber="289">Hornabrook</location>
<specimenCode id="79539EE0FFCF6D7ED41FDA06BCFE0508" box="[597,761,817,843]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="38" pageNumber="289">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFCF6D7ED55BDA06BD6C0508" box="[785,875,817,843]" pageId="38" pageNumber="289">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="999D3CC6FFCF6D7ED5CADA06BAC40508" box="[896,1219,817,843]" collectionCode="NEW, GUINEA" country="Papua New Guinea" pageId="38" pageNumber="289" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="3FF3FD12FFCF6D7ED5CADA06BDAB0508" box="[896,940,817,843]" count="1" pageId="38" pageNumber="289" type="male">1 ♂</specimenCount>
,
<specimenCount id="3FF3FD12FFCF6D7ED5F3DA06BDFC0508" box="[953,1019,817,843]" count="2" pageId="38" pageNumber="289" type="female">2 ♀♀</specimenCount>
: ‘
<collectionCode id="4FE4AE5EFFCF6D7ED247DA06BA560508" box="[1037,1105,817,843]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="38" pageNumber="289">NEW</collectionCode>
<collectionCode id="4FE4AE5EFFCF6D7ED213DA06BAC40508" box="[1113,1219,817,843]" pageId="38" pageNumber="289">GUINEA</collectionCode>
</materialsCitation>
<materialsCitation id="999D3CC6FFCF6D7ED280DA06BEB1052C" collectingDate="1964-09" collectionCode="R" country="Papua New Guinea" location="Okapa" pageId="38" pageNumber="289" specimenCount="1">
<location id="2C2A6040FFCF6D7ED280DA06BB110508" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED280DA06BB110508" box="[1226,1302,817,843]" country="Papua New Guinea" name="Okapa" pageId="38" pageNumber="289">Okapa</location>
, |
<date id="5D4B105BFFCF6D7ED365DA06BB940508" box="[1327,1427,817,843]" pageId="38" pageNumber="289" value="1964-09">
<collectingDate id="4D0FE9B3FFCF6D7ED365DA06BB940508" box="[1327,1427,817,843]" pageId="38" pageNumber="289" value="1964-09">IX.1964</collectingDate>
</date>
’, ‘
<collectionCode id="4FE4AE5EFFCF6D7ED6D7DA62BEB1052C" box="[157,182,853,879]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="289">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="999D3CC6FFCF6D7ED68BDA62BC6A052C" box="[193,621,853,879]" collectionCode="BMNH" country="Papua New Guinea" location="Hornabrook" pageId="38" pageNumber="289" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFCF6D7ED68BDA62BF4F052C" LSID="urn:lsid:plazi:treatment:A15C878DFFCF6D7FD6DDD881BF4201B9:2C2A6040FFCF6D7ED68BDA62BF4F052C" box="[193,328,853,879]" country="Papua New Guinea" name="Hornabrook" pageId="38" pageNumber="289">Hornabrook</location>
<specimenCode id="79539EE0FFCF6D7ED705DA62BFF3052C" box="[335,500,853,879]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="38" pageNumber="289">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFCF6D7ED441DA62BC62052C" box="[523,613,853,879]" pageId="38" pageNumber="289">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCF6D7FD68DDA4FBD010147" lastPageId="39" lastPageNumber="290" pageId="38" pageNumber="289" type="description">
<paragraph id="294A369BFFCF6D7ED68DDA4FBB420598" blockId="38.[151,1437,674,1987]" pageId="38" pageNumber="289">
<emphasis id="1B81EA89FFCF6D7ED68DDA4FBF5E05D1" bold="true" box="[199,345,888,914]" pageId="38" pageNumber="289">Description.</emphasis>
Measurements (n=11): HW: 0.35–0.36; HL: 0.20–0.22; OL: 0.11–0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFCF6D7ED362DA4FBB9605D1" box="[1320,1425,888,914]" pageId="38" pageNumber="289" type="holotype">holotype</typeStatus>
): 0.45; PL: 0.22–0.23; PWmax: 0.40–0.42; PWmin: 0.37–0.38; ESL: 0.45–0.47; EW: 0.60–0.62; MTbL (
<typeStatus id="F64E8839FFCF6D7ED362DAABBB9605F5" box="[1320,1425,924,950]" pageId="38" pageNumber="289" type="holotype">holotype</typeStatus>
): 0.27; MTrL (
<typeStatus id="F64E8839FFCF6D7ED762DAF7BF960599" box="[296,401,960,986]" pageId="38" pageNumber="289" type="holotype">holotype</typeStatus>
): 0.16 (MTrL 1–4: 0.06; MTrL 5: 0.10); AW: 0.62–0.63; AedL: 0.27; BL: 1.53–1.87.
</paragraph>
<paragraph id="294A369BFFCF6D7ED68DDAD4BC11037C" blockId="38.[151,1437,674,1987]" pageId="38" pageNumber="289">
Habitus as in
<figureCitation id="B1CE2A1EFFCF6D7ED72DDAD4BFBB05BD" box="[359,444,995,1022]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="38" pageNumber="289">Fig. 79</figureCitation>
. Head, antennomeres 6–11, latero-apical and basal portions of elytra reddish or darker (some specimens with paler middle portion of head); pronotum, elytra and abdomen yellow-brown (abdominal tergites VI–VII slightly darker); mouthparts, antennomeres 1–5 and legs yellow. Head with very sparse and fine punctation, sometimes indistinct in middle (some specimens only with several fine punctures in middle); neck without visible punctures; pronotum with very fine and sparse punctation, invisible in middle portion; elytra with moderatrely dense punctation, distinctly larger and deeper than that on pronotum, finer and sparser around scutellum and along suture; abdominal tergites with fine and moderately dense punctation. Body glossy; pronotum with dense isodiametric microsculpture, slightly finer than that in middle portion of head; scutellum without meshes; microsculpture of elytra dense, but distinctly finer than that in middle portion of pronotum; abdominal tergites without distinct microreticulation.
</paragraph>
<paragraph id="294A369BFFCF6D7ED68DDC7EBD460056" blockId="38.[151,1437,674,1987]" pageId="38" pageNumber="289">Head with shallow narrow anteriomedian depressions, 1.6–1.7 times as broad as long, with very short and deep anteocellar foveae and about as long as diameter of ocellus; postocular carina absent. Ocelli moderately large, located about at level of posterior margins of eyes; distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Antennomeres 6–10 strongly transverse; antennomere 3 distinctly shorter and narrower than 2, 4 distinctly shorter than 3, 5 slightly broader than 4, 6 about twice as broad as 5, 7–10 slightly broader than 6, antennomere 10 three times as broad as long.</paragraph>
<paragraph id="294A369BFFCF6D7ED68DDF17BFE1001E" blockId="38.[151,1437,674,1987]" pageId="38" pageNumber="289">Pronotum slightly convex, distinctly transverse and slightly broader than head; laterobasal impressions moderately wide and shallow.</paragraph>
<paragraph id="294A369BFFCF6D7ED68DDF50BD7600C2" blockId="38.[151,1437,674,1987]" box="[199,881,1639,1665]" pageId="38" pageNumber="289">Elytra 1.3 times as broad as long, twice as long as pronotum.</paragraph>
<paragraph id="294A369BFFCF6D7ED68DDFBDBCE500E7" blockId="38.[151,1437,674,1987]" box="[199,738,1674,1701]" pageId="38" pageNumber="289">Metatrsi 1.6 times as about as long as metatibia.</paragraph>
<paragraph id="294A369BFFCF6D7ED68DDF99BB270114" blockId="38.[151,1437,674,1987]" pageId="38" pageNumber="289">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFCF6D7ED521DF99BDC8008B" box="[875,975,1710,1736]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 113</figureCitation>
). Posterior margin of abdominal sternite VIII deeply concave (
<figureCitation id="B1CE2A1EFFCF6D7ED7C7DFE5BFE800AF" box="[397,495,1746,1772]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 114</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward moderately narrow median lobe with small rounded apex; parameres significantly exceeding apex of median lobe, with disitnctly broadened apical portion, each with two moderately long apical and five preapical setae; internal sac narrow and moderately long, without sclerotized structures (
<figureCitation id="B1CE2A1EFFCF6D7ED4FFDE0ABD120114" box="[693,789,1853,1879]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 111</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFCF6D7ED2F6DE0ABB1C0114" box="[1212,1307,1853,1879]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 112</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFCF6D7ED68DDE56BFC90180" blockId="38.[151,1437,674,1987]" pageId="38" pageNumber="289">
Female. Posterior margins of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFCF6D7ED557DE56BD7B013F" box="[797,892,1889,1916]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 115</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFCF6D7ED219DE56BAB50138" box="[1107,1202,1889,1915]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 116</figureCitation>
) truncate. Accessory sclerite wide and short, from widest basal part gradually narrowed toward somewhat truncate apex (
<figureCitation id="B1CE2A1EFFCF6D7ED361DEB2BB9701DC" box="[1323,1424,1925,1951]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 117</figureCitation>
). Spermatheca as in
<figureCitation id="B1CE2A1EFFCF6D7ED720DE9EBFCE0180" box="[362,457,1961,1987]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="38" pageNumber="289">Fig. 118</figureCitation>
.
</paragraph>
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<paragraph id="294A369BFFCE6D7FD6DDDC8FBA4E005B" blockId="39.[151,1436,1464,1561]" pageId="39" pageNumber="290">
<emphasis id="1B81EA89FFCE6D7FD6DDDC8FBF6A0392" bold="true" box="[151,365,1464,1489]" pageId="39" pageNumber="290">FIGURES 111–118.</emphasis>
<taxonomicName id="EEF54D18FFCE6D7FD732DC8EBCAE0392" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[376,681,1465,1489]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFCE6D7FD732DC8EBCAE0392" box="[376,681,1465,1489]" italics="true" pageId="39" pageNumber="290">Paraphloeostiba rhopalocera</emphasis>
</taxonomicName>
: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFCE6D7FD68DDF6BBD010147" blockId="39.[151,1437,1628,2042]" pageId="39" pageNumber="290">
<emphasis id="1B81EA89FFCE6D7FD68DDF6BBFB50034" bold="true" box="[199,434,1628,1655]" pageId="39" pageNumber="290">Comparative notes.</emphasis>
Based on the general shape of the body and the aedeagus, and the shape of the strongly transverse preapical antennomere three times as broad as long,
<taxonomicName id="EEF54D18FFCE6D7FD51ADFB6BDFF00D9" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[848,1016,1664,1691]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFCE6D7FD51ADFB6BD6600D9" box="[848,865,1665,1690]" italics="true" pageId="39" pageNumber="290">P</emphasis>
.
<emphasis id="1B81EA89FFCE6D7FD524DFB7BDFF00D9" box="[878,1016,1664,1690]" italics="true" pageId="39" pageNumber="290">rhopalocera</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="EEF54D18FFCE6D7FD2CFDFB6BEDC00FD" authority="(Bernhauer, 1915)" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="specularis">
<emphasis id="1B81EA89FFCE6D7FD2CFDFB6BB1500D9" box="[1157,1298,1664,1690]" italics="true" pageId="39" pageNumber="290">P. specularis</emphasis>
(
<bibRefCitation id="4D644B6AFFCE6D7FD36ADFB7BED500FD" author="Bernhauer, M" pageId="39" pageNumber="290" pagination="179 - 202" refId="ref35314" refString="Bernhauer, M (1915) Beitrag zur Staphylinidenfauna von Neu-Guinea. Deutsche Entomologische Zeitschrift, 1915, 179 - 202. https: // doi. org / 10.1002 / mmnd. 48019150207" type="journal article" year="1915">Bernhauer, 1915</bibRefCitation>
)
</taxonomicName>
, known from
<collectingRegion id="EB31F879FFCE6D7FD73BDF93BFFB00FE" box="[369,508,1699,1726]" country="Papua New Guinea" name="West New Britain" pageId="39" pageNumber="290">New Britain</collectingRegion>
,
<collectingCountry id="51E2760BFFCE6D7FD44EDF94BCFA00FE" box="[516,765,1699,1725]" name="Papua New Guinea" pageId="39" pageNumber="290">Bismarck Archipelago</collectingCountry>
(
<bibRefCitation id="4D644B6AFFCE6D7FD543DF94BDC700FD" author="Bernhauer, M" box="[777,960,1699,1726]" pageId="39" pageNumber="290" pagination="179 - 202" refId="ref35314" refString="Bernhauer, M (1915) Beitrag zur Staphylinidenfauna von Neu-Guinea. Deutsche Entomologische Zeitschrift, 1915, 179 - 202. https: // doi. org / 10.1002 / mmnd. 48019150207" type="journal article" year="1915">Bernhauer 1915</bibRefCitation>
,
<bibRefCitation id="4D644B6AFFCE6D7FD583DF94BA4500FE" author="Steel, W. O." box="[969,1090,1699,1725]" pageId="39" pageNumber="290" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960</bibRefCitation>
).
<taxonomicName id="EEF54D18FFCE6D7FD219DF94BB9B00FE" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[1107,1436,1699,1725]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFCE6D7FD219DF94BB9B00FE" box="[1107,1436,1699,1725]" italics="true" pageId="39" pageNumber="290">Paraphloeostiba rhopalocera</emphasis>
</taxonomicName>
can be distinguished from it by the slightly shorter body, the presence of the distinct microsculpture on the pronotum, and distinctly narrower apical portion of the parameres.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCE6D7FD68DDE3BBEEE01D3" pageId="39" pageNumber="290" type="distribution">
<paragraph id="294A369BFFCE6D7FD68DDE3BBEF70109" blockId="39.[151,1437,1628,2042]" pageId="39" pageNumber="290">
<emphasis id="1B81EA89FFCE6D7FD68DDE3BBF660165" bold="true" box="[199,353,1804,1830]" pageId="39" pageNumber="290">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFCE6D7FD727DE3ABCB80164" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[365,703,1805,1831]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFCE6D7FD727DE3ABCB80164" box="[365,703,1805,1831]" italics="true" pageId="39" pageNumber="290">Paraphloeostiba rhopalocera</emphasis>
</taxonomicName>
is known from several localities in Sumatra and
<collectingCountry id="51E2760BFFCE6D7FD35CDE3ABEEB0109" name="Papua New Guinea" pageId="39" pageNumber="290">Papua New Guinea</collectingCountry>
.
</paragraph>
<paragraph id="294A369BFFCE6D7FD68DDE64BEEE01D3" blockId="39.[151,1437,1628,2042]" pageId="39" pageNumber="290">
<emphasis id="1B81EA89FFCE6D7FD68DDE64BF4C012E" bold="true" box="[199,331,1875,1901]" pageId="39" pageNumber="290">Bionomics.</emphasis>
<materialsCitation id="999D3CC6FFCE6D7FD71BDE64BEEE01D3" country="Papua New Guinea" elevation="1828" location="Papua New Guinea" pageId="39" pageNumber="290" specimenCount="5">
<specimenCount id="3FF3FD12FFCE6D7FD71BDE64BFF9012E" box="[337,510,1875,1901]" count="5" pageId="39" pageNumber="290" type="generic">Five specimens</specimenCount>
from
<collectingCountry id="51E2760BFFCE6D7FD40BDE63BD1F012E" box="[577,792,1875,1902]" name="Papua New Guinea" pageId="39" pageNumber="290">Papua New Guinea</collectingCountry>
were collected under bark of wet log at an
<elevation id="A2D8D1A8FFCE6D7FD2BDDE64BEEE01D3" metricMagnitude="3" metricUnit="m" metricValue="1.828" pageId="39" pageNumber="290" unit="m" value="1828.0">
elevation
<quantity id="EE0D9B7EFFCE6D7FD32EDE64BEAA01D2" metricMagnitude="3" metricUnit="m" metricValue="1.828" pageId="39" pageNumber="290" unit="m" value="1828.0">1828 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCE6D7FD68DDEADBF4201B9" pageId="39" pageNumber="290" type="discussion">
<paragraph id="294A369BFFCE6D7FD68DDEADBF4201B9" blockId="39.[151,1437,1628,2042]" pageId="39" pageNumber="290">
<emphasis id="1B81EA89FFCE6D7FD68DDEADBF3C01F7" bold="true" box="[199,315,1946,1972]" pageId="39" pageNumber="290">Remarks.</emphasis>
<taxonomicName id="EEF54D18FFCE6D7FD70FDEAEBC7A01F0" authorityName="Cameron" authorityYear="1928" box="[325,637,1945,1971]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="rhopalocerus">
<emphasis id="1B81EA89FFCE6D7FD70FDEAEBC7A01F0" box="[325,637,1945,1971]" italics="true" pageId="39" pageNumber="290">Phloeonomus rhopalocerus</emphasis>
</taxonomicName>
was originally described from
<collectingRegion id="EB31F879FFCE6D7FD5A0DEADBA4D01F7" box="[1002,1098,1946,1972]" country="Indonesia" name="Sumatera Utara" pageId="39" pageNumber="290">Sumatra</collectingRegion>
,
<collectingCountry id="51E2760BFFCE6D7FD212DEAEBAC201F0" box="[1112,1221,1945,1971]" name="Indonesia" pageId="39" pageNumber="290">Indonesia</collectingCountry>
(“Fort de Kock”).
<bibRefCitation id="4D644B6AFFCE6D7FD6DDDE8ABF210194" author="Steel, W. O." box="[151,294,1981,2007]" pageId="39" pageNumber="290" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
redescribed it, transferred to the genus
<taxonomicName id="EEF54D18FFCE6D7FD54CDE8ABDC60194" authorityName="Steel" authorityYear="1960" box="[774,961,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFCE6D7FD54CDE8ABDC60194" box="[774,961,1981,2007]" italics="true" pageId="39" pageNumber="290">Paraphloeostiba</emphasis>
</taxonomicName>
, and recorded from
<collectingCountry id="51E2760BFFCE6D7FD2F0DE8ABB9B0194" box="[1210,1436,1981,2007]" name="Papua New Guinea" pageId="39" pageNumber="290">Papua New Guinea</collectingCountry>
(“Finschafen”).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:part id="3D44C9CF2766FA2B9D401B2191C93BC5">
<mods:date id="3AC27146DB46BD6397F970A5CFD9209D">2024</mods:date>
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<mods:number id="0A30868A0B922CFA96266EC82C9AE3A7">2024-03-14</mods:number>
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<mods:number id="7B312D51192F134327241A348BF72F52">5424</mods:number>
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<mods:classification id="019F61EA6AF22A3021DBFE12C0CDA606">journal article</mods:classification>
<mods:identifier id="633FA2F48B93C9B5F8CB77A4960BC330" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="E6F1329443B9FBC56887B303573F18BB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="173D57ABB0E2C722A6CCF774A3B7B973" type="Zenodo-Dep">10821214</mods:identifier>
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<subSubSection id="61EF6510FFD86D69D6DDDCB4BA7803DE" box="[151,1151,1410,1437]" pageId="49" pageNumber="300" type="nomenclature">
<paragraph id="294A369BFFD86D69D6DDDCB4BA7803DE" blockId="49.[151,1151,1410,1437]" box="[151,1151,1410,1437]" pageId="49" pageNumber="300">
<heading id="720281F7FFD86D69D6DDDCB4BA7803DE" bold="true" box="[151,1151,1410,1437]" fontSize="11" level="1" pageId="49" pageNumber="300" reason="1">
<emphasis id="1B81EA89FFD86D69D6DDDCB4BA7803DE" bold="true" box="[151,1151,1410,1437]" pageId="49" pageNumber="300">
Key to species of the genus
<taxonomicName id="EEF54D18FFD86D69D7A6DCB5BCB603DE" ID-CoL="8W25K" authorityName="Steel" authorityYear="1960" box="[492,689,1410,1437]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFD86D69D7A6DCB5BCB603DE" bold="true" box="[492,689,1410,1437]" italics="true" pageId="49" pageNumber="300">Paraphloeostiba</emphasis>
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of New
<collectingCountry id="51E2760BFFD86D69D55DDCB5BD7503DF" box="[791,882,1410,1436]" name="Guinea-Bissau" pageId="49" pageNumber="300">Guinea</collectingCountry>
and
<collectingCountry id="51E2760BFFD86D69D5FADCB5BA7803DE" box="[944,1151,1410,1437]" name="Solomon Islands" pageId="49" pageNumber="300">Solomon Islands</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFD86D6BD6DDDCFFBB9B052E" lastPageId="51" lastPageNumber="302" pageId="49" pageNumber="300" type="key">
<paragraph id="294A369BFFD86D69D6DDDCFFBB9B039E" blockId="49.[151,1437,1480,2030]" box="[151,1436,1480,1502]" pageId="49" pageNumber="300">1 Middle portion of pronotum with two distinct longitudinal impressions.......................................... 2</paragraph>
<paragraph id="294A369BFFD86D69D6DDDCD2BB9B03B9" blockId="49.[151,1437,1480,2030]" box="[151,1436,1509,1531]" pageId="49" pageNumber="300">- Middle portion of pronotum without or with indistinct longitudinal impressions................................... 8</paragraph>
<paragraph id="294A369BFFD86D69D6DDDF35BB9B0054" blockId="49.[151,1437,1480,2030]" box="[151,1436,1538,1560]" pageId="49" pageNumber="300">2 Body more than 2.00 mm.............................................................................. 4</paragraph>
<paragraph id="294A369BFFD86D69D6DDDF17BB9B0076" blockId="49.[151,1437,1480,2030]" box="[151,1436,1568,1590]" pageId="49" pageNumber="300">- Body distinctly less than 2.00 mm....................................................................... 7</paragraph>
<paragraph id="294A369BFFD86D69D6DDDF0ABB9B00CE" blockId="49.[151,1437,1480,2030]" pageId="49" pageNumber="300">
4 Antennomeres 8–10 slightly transverse. Elytra slightly more than twice as long as pronotum. Median lobe wide, gradually narrowed toward apex; parameres significantly longer than apex of median lobe, with wide apical portions (
<figureCitation id="B1CE2A1EFFD86D69D34BDF6DBB51002C" box="[1281,1366,1626,1648]" captionStart="FIGURES 148–151" captionStartId="48.[151,264,1862,1886]" captionTargetBox="[168,1427,193,1830]" captionTargetId="figure-22@48.[151,1436,181,1838]" captionTargetPageId="48" captionText="FIGURES 148–151. Paraphloeostiba vitiosa (holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="49" pageNumber="300">Fig. 148</figureCitation>
). Body yellow-brown. Body length:
<quantity id="EE0D9B7EFFD86D69D7BDDF4FBC4800CE" box="[503,591,1656,1677]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="49" pageNumber="300" unit="mm" value="2.1">2.10 mm</quantity>
.
<collectingCountry id="51E2760BFFD86D69D413DF4FBCB100CD" box="[601,694,1656,1678]" name="Indonesia" pageId="49" pageNumber="300">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFD86D69D501DF4FBD9500CD" box="[843,914,1656,1678]" name="Guinea-Bissau" pageId="49" pageNumber="300">Guinea</collectingCountry>
)..............................
<taxonomicName id="EEF54D18FFD86D69D343DF4FBB6700CE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1289,1376,1656,1677]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="vitiosa" status="sp. n.">
<emphasis id="1B81EA89FFD86D69D343DF4FBB6700CE" box="[1289,1376,1656,1677]" italics="true" pageId="49" pageNumber="300">P. vitiosa</emphasis>
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<emphasis id="1B81EA89FFD86D69D32CDF4FBB9B00CE" bold="true" box="[1382,1436,1656,1677]" pageId="49" pageNumber="300">
<taxonomicNameLabel id="00B257F2FFD86D69D32CDF4FBB9B00CE" box="[1382,1436,1656,1677]" pageId="49" pageNumber="300" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFD86D69D6DDDFA2BB9B008B" blockId="49.[151,1437,1480,2030]" pageId="49" pageNumber="300">- Antennomeres 8–10 distinctly transverse. Elytra less than twice as long as pronotum. Median lobe different, parameres shorter............................................................................................. 5</paragraph>
<paragraph id="294A369BFFD86D69D6DDDFF8BB9B0140" blockId="49.[151,1437,1480,2030]" pageId="49" pageNumber="300">
5 Head and pronotum without punctation. Aedeagus as in
<figureCitation id="B1CE2A1EFFD86D69D54ADFE7BD4300A6" box="[768,836,1744,1766]" captionStart="FIGURE 11" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[151,1435,525,1610]" captionTargetId="figure-143@5.[151,1436,525,1610]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of Nipponophloeostiba: N. verrucifera (circles), N. thayeri (triangle), N. longilobata (square), N. explanata (diamond), N. hornabrooki (star)." figureDoi="http://doi.org/10.5281/zenodo.10821220" httpUri="https://zenodo.org/record/10821220/files/figure.png" pageId="49" pageNumber="300">
Fig.
<quantity id="EE0D9B7EFFD86D69D567DFE7BD4300A6" box="[813,836,1744,1765]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="49" pageNumber="300" unit="in" value="11.0">11</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFD86D69D529DFE7BA0400A5" author="Shavrin, A. V." box="[867,1027,1744,1766]" pageId="49" pageNumber="300" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Body brown. Habitus as in
<figureCitation id="B1CE2A1EFFD86D69D35CDFE7BB5D00A6" box="[1302,1370,1744,1766]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="49" pageNumber="300">Fig. 39</figureCitation>
and in
<figureCitation id="B1CE2A1EFFD86D69D6A8DFDABF1D0141" box="[226,282,1773,1795]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="49" pageNumber="300">
Fig.
<quantity id="EE0D9B7EFFD86D69D744DFDABF1D0141" box="[270,282,1773,1794]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="49" pageNumber="300" unit="in" value="2.0">2</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFD86D69D773DFDABFDF0140" author="Shavrin, A. V." box="[313,472,1773,1795]" pageId="49" pageNumber="300" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFD86D69D545DFDABD510140" box="[783,854,1773,1795]" captionStart="FIGURES 53–56" captionStartId="21.[151,264,1187,1211]" captionTargetBox="[159,1428,187,1164]" captionTargetId="figure-16@21.[151,1436,181,1164]" captionTargetPageId="21" captionText="FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="49" pageNumber="300">Fig. 56</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFD86D69D5A8DFDABA6A0141" box="[994,1133,1773,1794]" metricMagnitude="-3" metricUnit="m" metricValue="2.58" metricValueMax="3.08" metricValueMin="2.08" pageId="49" pageNumber="300" unit="mm" value="2.58" valueMax="3.08" valueMin="2.08">2.08–3.08 mm</quantity>
<taxonomicName id="EEF54D18FFD86D69D2CEDFD9BB9B0140" authority="(Cameron, 1952)" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[1156,1436,1773,1795]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="coriacea">
<emphasis id="1B81EA89FFD86D69D2CEDFD9BA940140" box="[1156,1171,1774,1795]" italics="true" pageId="49" pageNumber="300">P</emphasis>
.
<emphasis id="1B81EA89FFD86D69D2D4DFD9BAF50140" box="[1182,1266,1774,1795]" italics="true" pageId="49" pageNumber="300">coriacea</emphasis>
(
<bibRefCitation id="4D644B6AFFD86D69D2B5DFDABB920140" author="Cameron, M." box="[1279,1429,1773,1795]" pageId="49" pageNumber="300" pagination="241 - 256" refId="ref35440" refString="Cameron, M. (1952) Results of the Archibold Expeditions. New species of Staphylinidae (Col.) from New Guinea and Misool. Treubia, 21 (2), 241 - 256." type="journal article" year="1952">Cameron, 1952</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFD86D69D6DDDE3CBB9B0163" blockId="49.[151,1437,1480,2030]" box="[151,1436,1802,1824]" pageId="49" pageNumber="300">- Head and pronotum with distinct punctation................................................................ 6</paragraph>
<paragraph id="294A369BFFD86D69D6DDDE1FBB9B01D6" blockId="49.[151,1437,1480,2030]" pageId="49" pageNumber="300">
6 Pronotum widest slightly above middle, strongly narrowed posteriad, with slightly concave laterobasal margins. Apical part of median lobe moderately narrow; parameres slightly not reaching apex of median lobe, with distinctly broadened apical portions (
<figureCitation id="B1CE2A1EFFD86D69D6A3DE55BF35013B" box="[233,306,1890,1912]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="49" pageNumber="300">Fig. 57</figureCitation>
). Body yellow-brown to reddish-brown. Habitus as in
<figureCitation id="B1CE2A1EFFD86D69D50CDE55BD880134" box="[838,911,1890,1912]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="49" pageNumber="300">Fig. 42</figureCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFD86D69D286DE55BB13013B" box="[1228,1300,1890,1912]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="49" pageNumber="300">Fig. 63</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFD86D69D6A8DEB7BF7701D6" box="[226,368,1920,1941]" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="3.07" metricValueMin="2.07" pageId="49" pageNumber="300" unit="mm" value="2.57" valueMax="3.07" valueMin="2.07">2.07–3.07 mm</quantity>
.
<collectingCountry id="51E2760BFFD86D69D733DEB7BC1D01D5" box="[377,538,1920,1942]" name="Solomon Islands" pageId="49" pageNumber="300">Solomon Islands</collectingCountry>
.....................................................
<taxonomicName id="EEF54D18FFD86D69D2EBDEB7BB6701D6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1185,1376,1920,1941]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFD86D69D2EBDEB7BB6701D6" box="[1185,1376,1920,1941]" italics="true" pageId="49" pageNumber="300">P. guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFD86D69D32CDEB7BB9B01D6" bold="true" box="[1382,1436,1920,1941]" pageId="49" pageNumber="300">
<taxonomicNameLabel id="00B257F2FFD86D69D32CDEB7BB9B01D6" box="[1382,1436,1920,1941]" pageId="49" pageNumber="300" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFD86D6AD6DDDEAABB9B068A" blockId="49.[151,1437,1480,2030]" lastBlockId="50.[151,1437,151,2017]" lastPageId="50" lastPageNumber="301" pageId="49" pageNumber="300">
- Pronotum widest in middle, relatively gradually narowed toward hind angles, without distinctly concave laterobasal margins. Apical part of median lobe moderately broad; parameres reaching or slightly exceeding apex of median lobe, with slightly broadened apical portions (
<figureCitation id="B1CE2A1EFFD86D69D7ADDEEFBC2B01AE" box="[487,556,2007,2030]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="49" pageNumber="300">
Fig.
<quantity id="EE0D9B7EFFD86D69D45EDEE0BC2B01AE" box="[532,556,2007,2029]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="49" pageNumber="300" unit="in" value="25.0">25</quantity>
</figureCitation>
in Shavrin (2020)). Body brown, usually with paler middle portions of elytra. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD6A8D9A0BF1206EE" box="[226,277,151,173]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">
Fig
<quantity id="EE0D9B7EFFDB6D6AD743D9AFBF1206EE" box="[265,277,152,173]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="50" pageNumber="301" unit="in" value="4.0">4</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD77FD9A0BFD206EE" author="Shavrin, A. V." box="[309,469,151,173]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDB6D6AD55AD9A0BD5206EE" box="[784,853,151,173]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD577D9AFBD5206EE" box="[829,853,152,173]" metricMagnitude="0" metricUnit="m" metricValue="2.5146" pageId="50" pageNumber="301" unit="in" value="99.0">99</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD53FD9A0BDE806EE" author="Steel, W. O." box="[885,1007,151,173]" pageId="50" pageNumber="301" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD237D9A0BB0C06EE" box="[1149,1291,151,173]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.5" metricValueMin="2.1" pageId="50" pageNumber="301" unit="mm" value="2.8" valueMax="3.5" valueMin="2.1">2.10–3.50 mm</quantity>
. Cosmopolitan species...................................................................
<taxonomicName id="EEF54D18FFDB6D6AD21DD982BB9B068A" authority="(MacLeay, 1873)" baseAuthorityName="MacLeay" baseAuthorityYear="1873" box="[1111,1436,180,202]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="gayndahensis">
<emphasis id="1B81EA89FFDB6D6AD21DD982BAF5068A" box="[1111,1266,180,202]" italics="true" pageId="50" pageNumber="301">P. gayndahensis</emphasis>
(
<bibRefCitation id="4D644B6AFFDB6D6AD2B4D983BB92068A" author="MacLeay, W. J." box="[1278,1429,180,201]" pageId="50" pageNumber="301" pagination="79 - 205" refId="ref35618" refString="MacLeay, W. J. (1873) Notes on a collection of insects from Gayndah. The Transactions of the Entomological Society of New South Wales, 2, 79 - 205." type="journal article" year="1873">MacLeay, 1873</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD9E5BB9B0763" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
7 Elytra about twice as long as pronotum. Apical part of median lobe moderately wide; parameres slightly exceeding apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD72FD9D9BFAA0747" box="[357,429,238,260]" captionStart="FIGURES 64–67" captionStartId="25.[151,264,1719,1743]" captionTargetBox="[159,1426,190,1687]" captionTargetId="figure-16@25.[151,1436,181,1695]" captionTargetPageId="25" captionText="FIGURES 64–67. Paraphloeostiba iriana (holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821247" httpUri="https://zenodo.org/record/10821247/files/figure.png" pageId="50" pageNumber="301">Fig. 64</figureCitation>
). Body brown. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD48ED9D9BD0B0740" box="[708,780,238,260]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="50" pageNumber="301">Fig. 40</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD5DDD9D9BA220740" box="[919,1061,238,260]" metricMagnitude="-3" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.96" metricValueMin="1.72" pageId="50" pageNumber="301" unit="mm" value="1.8399999999999999" valueMax="1.96" valueMin="1.72">1.72–1.96 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD265D9D9BA8B0747" box="[1071,1164,238,260]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFDB6D6AD36BD9D9BB6F0747" box="[1313,1384,238,260]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
)...........................................................................................
<taxonomicName id="EEF54D18FFDB6D6AD344D83CBB660763" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1294,1377,267,288]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD344D83CBB660763" box="[1294,1377,267,288]" italics="true" pageId="50" pageNumber="301">P. iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CD83CBB9B0763" bold="true" box="[1382,1436,267,288]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CD83CBB9B0763" box="[1382,1436,267,288]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD81FBB9B0735" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Elytra distinctly less than twice as long as pronotum. Apex of median lobe narrow; parameres distinctly not exceeding apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD72FD873BFAA0719" box="[357,429,324,346]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="50" pageNumber="301">Fig. 85</figureCitation>
). Body reddish-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD559D873BD5D0719" box="[787,858,324,346]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 75</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD5ACD873BA730719" box="[998,1140,324,346]" metricMagnitude="-3" metricUnit="m" metricValue="1.745" metricValueMax="1.86" metricValueMin="1.63" pageId="50" pageNumber="301" unit="mm" value="1.745" valueMax="1.86" valueMin="1.63">1.63–1.86 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD237D872BB310719" box="[1149,1334,324,346]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
............................................................................................
<taxonomicName id="EEF54D18FFDB6D6AD2A0D855BB660735" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1258,1377,353,375]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="okapensis" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2A0D855BB660735" box="[1258,1377,353,375]" italics="true" pageId="50" pageNumber="301">P. okapensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CD856BB9B0735" bold="true" box="[1382,1436,353,374]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CD856BB9B0735" box="[1382,1436,353,374]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD849BB9B07D0" blockId="50.[151,1437,151,2017]" box="[151,1436,382,404]" pageId="50" pageNumber="301">8 Antennomeres 6–10 strongly transverse, preapical antennomere three times as broad as long......................... 9</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD8ACBB9B07F3" blockId="50.[151,1437,151,2017]" box="[151,1436,411,433]" pageId="50" pageNumber="301">- Antennomeres 6–10 less transverse, preapical antennomere about twice as broad as long........................... 10</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD88FBB9B0460" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
9 Elytra about twice as long as pronotum. Parameres significantly longer than apex of median lobe, with wide apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD6A3D8E3BF3B07AA" box="[233,316,468,490]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="50" pageNumber="301">Fig. 111</figureCitation>
). Head and elytra reddish or darker; pronotum and abdomen yellow-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD2EBD8E3BAEF07AA" box="[1185,1256,468,490]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 79</figureCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDB6D6AD72AD8C6BFB50445" box="[352,434,497,519]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="50" pageNumber="301">Fig. 117</figureCitation>
. Body smaller:
<quantity id="EE0D9B7EFFDB6D6AD402D8C6BCD10444" box="[584,726,497,519]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.87" metricValueMin="1.53" pageId="50" pageNumber="301" unit="mm" value="1.7" valueMax="1.87" valueMin="1.53">1.53–1.87 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD4AAD8C6BD3A0444" box="[736,829,497,519]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(
<collectingRegion id="EB31F879FFDB6D6AD500D8C6BD980445" box="[842,927,497,518]" country="Indonesia" name="Sumatera Utara" pageId="50" pageNumber="301">Sumatra</collectingRegion>
),
<collectingCountry id="51E2760BFFDB6D6AD5E5D8C5BA6F0444" box="[943,1128,497,519]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
...................................................................................................
<taxonomicName id="EEF54D18FFDB6D6AD22EDB38BB9B0460" authority="(Cameron, 1928)" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[1124,1436,526,548]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFDB6D6AD22EDB38BAF50460" box="[1124,1266,526,548]" italics="true" pageId="50" pageNumber="301">P. rhopalocera</emphasis>
(
<bibRefCitation id="4D644B6AFFDB6D6AD2B5DB39BB920460" author="Cameron, M." box="[1279,1429,526,547]" pageId="50" pageNumber="301" pagination="90 - 110" refId="ref35375" refString="Cameron, M. (1928) Fauna sumatrensis. Staphylinidae (Col.). Entomologische Mitteilungen, 17 (2), 90 - 110." type="journal article" year="1928">Cameron, 1928</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDB1CBB9B0439" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Elytra distinctly less than twice as long as pronotum. Parameres distinctly exceeding apex of median lobe, with very wide apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD777DB70BF86041E" box="[317,385,583,605]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD723DB7FBF86041E" box="[361,385,584,605]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="50" pageNumber="301" unit="in" value="32.0">32</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD7D5DB70BC0F041E" author="Steel, W. O." box="[415,520,583,605]" pageId="50" pageNumber="301" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960</bibRefCitation>
). Body reddish-brown. Forebody as in
<figureCitation id="B1CE2A1EFFDB6D6AD536DB70BDB4041E" box="[892,947,583,605]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD5EDDB7FBDB4041E" box="[935,947,584,605]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="50" pageNumber="301" unit="in" value="9.0">9</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD59BDB70BA4E041E" author="Steel, W. O." box="[977,1097,583,605]" pageId="50" pageNumber="301" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
. Body larger: 2.00 mm.
<collectingCountry id="51E2760BFFDB6D6AD365DB7FBF200439" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
(
<collectingRegion id="EB31F879FFDB6D6AD77EDB53BFAB0439" box="[308,428,612,634]" country="Papua New Guinea" name="West New Britain" pageId="50" pageNumber="301">New Britain</collectingRegion>
).........................................................
<taxonomicName id="EEF54D18FFDB6D6AD227DB52BB9B0439" authority="(Bernhauer, 1915)" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[1133,1436,612,634]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="specularis">
<emphasis id="1B81EA89FFDB6D6AD227DB52BAEF043A" box="[1133,1256,612,634]" italics="true" pageId="50" pageNumber="301">P. specularis</emphasis>
(
<bibRefCitation id="4D644B6AFFDB6D6AD2BEDB53BB920439" author="Bernhauer, M" box="[1268,1429,612,634]" pageId="50" pageNumber="301" pagination="179 - 202" refId="ref35314" refString="Bernhauer, M (1915) Beitrag zur Staphylinidenfauna von Neu-Guinea. Deutsche Entomologische Zeitschrift, 1915, 179 - 202. https: // doi. org / 10.1002 / mmnd. 48019150207" type="journal article" year="1915">Bernhauer, 1915</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDBB6BB9B04D5" blockId="50.[151,1437,151,2017]" box="[151,1436,641,663]" pageId="50" pageNumber="301">10 Laterobasal margins of pronotum distinctly concave in front of hind angles...................................... 11</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDBA9BB9B04F0" blockId="50.[151,1437,151,2017]" box="[151,1436,670,692]" pageId="50" pageNumber="301">- Laterobasal margins of pronotum gradually narrowed toward hind angles....................................... 13</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDB8CBB9B054A" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
11 Pronotum twice as broad as long, with widely rounded apical angles slighlty protruded apicad. Parameres very narrow, significantly shorter than apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD4E2DBE0BCFB04AE" box="[680,764,727,749]" captionStart="FIGURES 144–147" captionStartId="47.[151,264,1614,1638]" captionTargetBox="[171,1427,190,1582]" captionTargetId="figure-16@47.[151,1436,181,1591]" captionTargetPageId="47" captionText="FIGURES 144–147. Paraphloeostiba telnovi (holotype): 144–aedeagus, parameral view, 145–aedeagus, lateral view, 146–male abdominal tergite VIII, 147–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821291" httpUri="https://zenodo.org/record/10821291/files/figure.png" pageId="50" pageNumber="301">Fig. 144</figureCitation>
). Body reddish-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD228DBE0BAB204AE" box="[1122,1205,727,749]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="50" pageNumber="301">Fig. 141</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD30BDBEFBB9E04AE" box="[1345,1433,728,749]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="50" pageNumber="301" unit="mm" value="1.82">1.82 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD6A8DBC2BF9C0549" box="[226,411,756,778]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
........................................................................
<taxonomicName id="EEF54D18FFDB6D6AD34DDBC2BB67054A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1287,1376,756,778]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="telnovi" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD34DDBC2BB67054A" box="[1287,1376,756,778]" italics="true" pageId="50" pageNumber="301">P. telnovi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDBC3BB9B054A" bold="true" box="[1382,1436,756,777]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDBC3BB9B054A" box="[1382,1436,756,777]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDA25BB9B0500" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">- Pronotum distinctly less than twice as broad as long, with narrower apical angles not protruded apicad. Parameres wider and longer............................................................................................. 12</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDA7CBB9B05DA" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
12 Head without or with indistinct fine punctation. Median lobe narrow; parameres narrow, reaching apex of median lobe, with narrow apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD78DDA50BC09053E" box="[455,526,871,893]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="50" pageNumber="301">Fig. 20</figureCitation>
). Body yellowish-brown to reddish brown, with paler elytra. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD296DA50BB11053E" box="[1244,1302,871,893]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">Fig. 4</figureCitation>
. Body length: 1.58–2.00 mm.
<collectingCountry id="51E2760BFFDB6D6AD733DAB2BC3505D9" box="[377,562,900,922]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
.........................................................
<taxonomicName id="EEF54D18FFDB6D6AD2A4DAB2BB6605D9" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1262,1377,901,922]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="attenuata" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2A4DAB2BB6605D9" box="[1262,1377,901,922]" italics="true" pageId="50" pageNumber="301">P. attenuata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDAB3BB9B05DA" bold="true" box="[1382,1436,900,921]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDAB3BB9B05DA" box="[1382,1436,900,921]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDA95BB9B024E" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Head without punctation. Median lobe moderately wide; parameres slightly exceeding apex of median lobe, with distinctly broadened apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD7A0DA89BC230590" box="[490,548,958,980]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD452DA89BC230590" box="[536,548,958,979]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="50" pageNumber="301" unit="in" value="7.0">7</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD40EDA89BCEA0597" author="Shavrin, A. V." box="[580,749,958,980]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b))</bibRefCitation>
. Body reddish-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD21BDA89BA8D0590" box="[1105,1162,958,980]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD234DA89BA8D0590" box="[1150,1162,958,979]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="50" pageNumber="301" unit="in" value="1.0">1</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD2E1DA89BB4C0597" author="Shavrin, A. V." box="[1195,1355,958,980]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDB6D6AD789DAECBC0D05B3" box="[451,522,987,1009]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="50" pageNumber="301">Fig. 38</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD4DCDAECBD2305B3" box="[662,804,987,1009]" metricMagnitude="-3" metricUnit="m" metricValue="2.065" metricValueMax="2.35" metricValueMin="1.78" pageId="50" pageNumber="301" unit="mm" value="2.065" valueMax="2.35" valueMin="1.78">1.78–2.35 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD567DAECBD8D05B2" box="[813,906,987,1009]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFDB6D6AD26ADAECBA6E05B2" box="[1056,1129,987,1009]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
),
<collectingCountry id="51E2760BFFDB6D6AD233DAECBB3505B2" box="[1145,1330,987,1009]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
......................................................................................
<taxonomicName id="EEF54D18FFDB6D6AD2DCDACFBB9B024E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1174,1436,1015,1037]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFDB6D6AD2DCDACFBAA2024E" box="[1174,1189,1016,1037]" italics="true" pageId="50" pageNumber="301">P</emphasis>
.
<emphasis id="1B81EA89FFDB6D6AD2FBDACFBB17024E" box="[1201,1296,1016,1037]" italics="true" pageId="50" pageNumber="301">conjuncta</emphasis>
Shavrin, 2023
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDD23BB9B026A" blockId="50.[151,1437,151,2017]" box="[151,1436,1044,1066]" pageId="50" pageNumber="301">13 Head and pronotum without punctation.................................................................. 14</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDD05BB9B0204" blockId="50.[151,1437,151,2017]" box="[151,1436,1073,1095]" pageId="50" pageNumber="301">- Head without or with distinct punctation, but pronotum always with punctures................................... 15</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDD79BB9B02FA" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
14 Pronotum transverse, slightly more than twice as broad as long; surface of elytra without elevations between punctures. Parameres slightly exceeding apex of median lobe, with wide apical portions, with several short apical and preapical setae (
<figureCitation id="B1CE2A1EFFDB6D6AD33FDD5CBEFA02DE" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="50" pageNumber="301">Fig. 97</figureCitation>
). Body yellow-brown, with reddish-brown apical portions of elytra. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD254DDB0BA6102DE" box="[1054,1126,1159,1181]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 76</figureCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDB6D6AD6A8DD93BF3202FA" box="[226,309,1188,1210]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="50" pageNumber="301">Fig. 103</figureCitation>
. Body length: 1.40–2.00 mm.
<collectingCountry id="51E2760BFFDB6D6AD412DD93BCFE02F9" box="[600,761,1188,1210]" name="Solomon Islands" pageId="50" pageNumber="301">Solomon Islands</collectingCountry>
.......................................
<taxonomicName id="EEF54D18FFDB6D6AD291DD92BB6702FA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1243,1376,1188,1210]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD291DD92BB6702FA" box="[1243,1376,1188,1210]" italics="true" pageId="50" pageNumber="301">P. penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDD93BB9B02FA" bold="true" box="[1382,1436,1188,1209]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDD93BB9B02FA" box="[1382,1436,1188,1209]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDDF5BB9B036E" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Pronotum less transverse, about one and a half times as broad as long; surface of elytra with distinct elevations between punctures. Parameres significantly exceeding apes of median lobe, with numerous very long apical and preapical setae (
<figureCitation id="B1CE2A1EFFDB6D6AD33FDDE9BF0E0353" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="50" pageNumber="301">Fig. 104</figureCitation>
). Body dark-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD41BDDCCBC900353" box="[593,663,1275,1297]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 77</figureCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDB6D6AD586DDCCBA190353" box="[972,1054,1275,1297]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="50" pageNumber="301">Fig. 110</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD2E2DDCCBB310353" box="[1192,1334,1275,1297]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.65" metricValueMin="1.95" pageId="50" pageNumber="301" unit="mm" value="2.3" valueMax="2.65" valueMin="1.95">1.95–2.65 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD375DDCCBB9B0352" box="[1343,1436,1275,1297]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFDB6D6AD73BDC20BFBF036E" box="[369,440,1303,1325]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
).....................................................................
<taxonomicName id="EEF54D18FFDB6D6AD344DC2FBB66036F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1294,1377,1303,1325]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="pilosa" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD344DC2FBB66036F" box="[1294,1377,1303,1325]" italics="true" pageId="50" pageNumber="301">P. pilosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDC2FBB9B036E" bold="true" box="[1382,1436,1304,1325]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDC2FBB9B036E" box="[1382,1436,1304,1325]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDC03BB9B03C0" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
15 Antennomeres 6–7 about as broad as long. Aedeagus long, with moderately wide and elongate median lobe with rounded apex (
<figureCitation id="B1CE2A1EFFDB6D6AD6A3DC66BF360325" box="[233,305,1361,1383]" captionStart="FIGURES 81–84" captionStartId="29.[151,264,1649,1673]" captionTargetBox="[168,1428,710,1617]" captionTargetId="figure-145@29.[151,1436,704,1625]" captionTargetPageId="29" captionText="FIGURES 81–84. Paraphloeostiba margineguttata (holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="50" pageNumber="301">Fig. 81</figureCitation>
). Body reddish-brown, with paler elytra and abdomen. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD59ADC66BA1F0325" box="[976,1048,1361,1383]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="50" pageNumber="301">Fig. 43</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD2ECDC66BB320324" box="[1190,1333,1361,1383]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="2.77" metricValueMin="2.57" pageId="50" pageNumber="301" unit="mm" value="2.67" valueMax="2.77" valueMin="2.57">2.57–2.77 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD375DC66BB9B0324" box="[1343,1436,1361,1383]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFDB6D6AD73BDC59BFBF03C7" box="[369,440,1390,1412]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
).............................................................
<taxonomicName id="EEF54D18FFDB6D6AD2FCDC58BB6703C7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1206,1376,1391,1412]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="margineguttata" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2FCDC58BB6703C7" box="[1206,1376,1391,1412]" italics="true" pageId="50" pageNumber="301">P. margineguttata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDC59BB9B03C0" bold="true" box="[1382,1436,1390,1411]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDC59BB9B03C0" box="[1382,1436,1390,1411]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDCBCBB9B03E2" blockId="50.[151,1437,151,2017]" box="[151,1436,1419,1441]" pageId="50" pageNumber="301">- Antennomeres 6–10 distinctly transverse. Aedeagus and coloration of body different.............................. 16</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDC90BB9B03FE" blockId="50.[151,1437,151,2017]" box="[151,1436,1447,1469]" pageId="50" pageNumber="301">16 Head without punctation.............................................................................. 17</paragraph>
<paragraph id="294A369BFFDB6D6AD6D7DCF2BB9B039A" blockId="50.[151,1437,151,2017]" box="[157,1436,1476,1498]" pageId="50" pageNumber="301">- Head with distinct punctation.......................................................................... 22</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDCD6BB9B0073" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
17 Elytra with large and deep punctation and with distinct elevations between punctures. Apical portion of median lobe very narrow; parameres with very wide apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD49BDCC9BD210050" box="[721,806,1534,1556]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="50" pageNumber="301">Fig. 133</figureCitation>
). Body reddish-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD2DBDCC9BADF0050" box="[1169,1240,1534,1556]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="50" pageNumber="301">Fig. 44</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD32CDCC9BF3D0073" metricMagnitude="-3" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.9" metricValueMin="1.78" pageId="50" pageNumber="301" unit="mm" value="1.8399999999999999" valueMax="1.9" valueMin="1.78">1.78– 1.90 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD709DF2CBFFB0072" box="[323,508,1563,1585]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
................................................................
<taxonomicName id="EEF54D18FFDB6D6AD35CDF2CBB660073" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1302,1377,1563,1584]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD35CDF2CBB660073" box="[1302,1377,1563,1584]" italics="true" pageId="50" pageNumber="301">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDF2CBB9B0073" bold="true" box="[1382,1436,1563,1584]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDF2CBB9B0073" box="[1382,1436,1563,1584]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDF0FBB9B002A" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">- Elytra with distinctly finer punctation, without elevations between punctures.Apical portion of median lobe narrower; parameres different. Coloration of body different................................................................... 18</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDF46BB9B0083" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
18 Elytra longer, twice as long as pronotum. Aedeagus as in
<figureCitation id="B1CE2A1EFFDB6D6AD546DF46BD5300C4" box="[780,852,1649,1671]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="50" pageNumber="301">Fig. 46</figureCitation>
. Body yellow-brown to brown, with slightly paler elytra and darkened apical portions. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDB6D6AD542DFB9BD4800E7" box="[776,847,1678,1700]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="50" pageNumber="301">Fig. 52</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFDB6D6AD590DFB9BA6F00E0" box="[986,1128,1678,1699]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" metricValueMax="2.07" metricValueMin="1.47" pageId="50" pageNumber="301" unit="mm" value="1.77" valueMax="2.07" valueMin="1.47">1.47–2.07 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD238DFB9BB2C00E7" box="[1138,1323,1678,1700]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
............................................................................................
<taxonomicName id="EEF54D18FFDB6D6AD2AADF9CBB670083" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1248,1376,1707,1728]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2AADF9CBB670083" box="[1248,1376,1707,1728]" italics="true" pageId="50" pageNumber="301">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDF9CBB9B0083" bold="true" box="[1382,1436,1707,1728]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDF9CBB9B0083" box="[1382,1436,1707,1728]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDFFFBB9B009E" blockId="50.[151,1437,151,2017]" box="[151,1436,1735,1757]" pageId="50" pageNumber="301">- Elytra shorter, distinctly less than twice as long as pronotum. Coloration of body different.......................... 19</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDFD3BB9B00BA" blockId="50.[151,1437,151,2017]" box="[151,1436,1764,1786]" pageId="50" pageNumber="301">19 Body yellow-brown. Median lobe and apical portions of parameres wide........................................ 20</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDE35BB9B0155" blockId="50.[151,1437,151,2017]" box="[151,1436,1793,1815]" pageId="50" pageNumber="301">- Body darker. Median lobe and apical portions of parameres narrow............................................ 21</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDE29BB9B012E" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
20 Lateral sides of pronotum gradually narrowed both anteriad and posteriad. Body yellow-brown, with reddish-brown apical portions of elytra. Median lobe slightly broader; parameres significantly longer than apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD295DE0CBB200113" box="[1247,1319,1851,1873]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="50" pageNumber="301">Fig. 12</figureCitation>
). Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD6B1DE60BF28012E" box="[251,303,1879,1901]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">Fig 2</figureCitation>
. Body smaller:
<quantity id="EE0D9B7EFFDB6D6AD78CDE60BC19012E" box="[454,542,1879,1901]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="50" pageNumber="301" unit="mm" value="1.76">1.76 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD46DDE6FBCE7012E" box="[551,736,1879,1901]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
...........................................
<taxonomicName id="EEF54D18FFDB6D6AD2B0DE6FBB66012F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1274,1377,1879,1901]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2B0DE6FBB66012F" box="[1274,1377,1879,1901]" italics="true" pageId="50" pageNumber="301">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDE6FBB9B012E" bold="true" box="[1382,1436,1880,1901]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDE6FBB9B012E" box="[1382,1436,1880,1901]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDE42BB9B01A3" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Lateral sides of pronotum slightly more narrowed posteriad than anteriad. Body yellow-brown. Median lobe relatively narrower; parameres slightly exceeding apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD480DEA6BD0901E4" box="[714,782,1937,1959]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD4BCDEA6BD0901E4" box="[758,782,1937,1959]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="50" pageNumber="301" unit="in" value="15.0">15</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD561DEA6BDD501E4" author="Shavrin, A. V." box="[811,978,1937,1959]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b))</bibRefCitation>
. Habitus as in
<figureCitation id="B1CE2A1EFFDB6D6AD211DEA6BA9501E5" box="[1115,1170,1937,1959]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD2CCDEA6BA9501E5" box="[1158,1170,1937,1958]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="50" pageNumber="301" unit="in" value="3.0">3</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD2E5DEA6BB4A01E4" author="Shavrin, A. V." box="[1199,1357,1937,1959]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite asi n
<figureCitation id="B1CE2A1EFFDB6D6AD789DE99BC000180" box="[451,519,1966,1988]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="50" pageNumber="301">
Fig.
<quantity id="EE0D9B7EFFDB6D6AD7A5DE99BC000180" box="[495,519,1966,1987]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="50" pageNumber="301" unit="in" value="21.0">21</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDB6D6AD46CDE99BCC20187" author="Shavrin, A. V." box="[550,709,1966,1988]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Body larger: 2.00–
<quantity id="EE0D9B7EFFDB6D6AD5C9DE99BDDC0180" box="[899,987,1966,1988]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="50" pageNumber="301" unit="mm" value="2.35">2.35 mm</quantity>
.
<collectingCountry id="51E2760BFFDB6D6AD5AEDE99BA9A0187" box="[996,1181,1966,1988]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
...................................................................................................
<taxonomicName id="EEF54D18FFDB6D6AD2E2DEFBBB9B01A3" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1192,1436,1995,2017]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="1B81EA89FFDB6D6AD2E2DEFBBAB001A2" box="[1192,1207,1996,2017]" italics="true" pageId="50" pageNumber="301">P</emphasis>
.
<emphasis id="1B81EA89FFDB6D6AD288DEFCBB0801A3" box="[1218,1295,1995,2016]" italics="true" pageId="50" pageNumber="301">flaveola</emphasis>
Shavrin, 2023
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD9AFBB9B0689" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
21 Disc of pronotum without longitudinal impressions. Apical parts of median lobe and parameres narrow (
<figureCitation id="B1CE2A1EFFDA6D6BD28DD9A0BB0906EE" box="[1223,1294,151,173]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="51" pageNumber="302">Fig. 32</figureCitation>
). Body brown. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD72FD982BF980688" box="[357,415,181,203]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">Fig. 6</figureCitation>
. Body smaller:
<quantity id="EE0D9B7EFFDA6D6BD47DD982BC880689" box="[567,655,181,202]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="51" pageNumber="302" unit="mm" value="1.42">1.42 mm</quantity>
.
<collectingCountry id="51E2760BFFDA6D6BD4D2D982BD560688" box="[664,849,181,203]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
.................................
<taxonomicName id="EEF54D18FFDA6D6BD2A3D982BB670689" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1257,1376,181,202]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD2A3D982BB670689" box="[1257,1376,181,202]" italics="true" pageId="51" pageNumber="302">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD982BB9B0689" bold="true" box="[1382,1436,181,202]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD982BB9B0689" box="[1382,1436,181,202]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD9E4BB9B0761" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
- Disc of pronotum with indistinct longitudinal impressions. Apical parts of median lobe and parameres distinctly broader (
<figureCitation id="B1CE2A1EFFDA6D6BD33FD9E5BF0E0746" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="51" pageNumber="302">Fig. 119</figureCitation>
). Body yellowish brown to reddish-brown, with slightly paler elytra. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD26FD9D8BA6B0746" box="[1061,1132,239,261]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="51" pageNumber="302">Fig. 78</figureCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDA6D6BD6A8D83ABF320760" box="[226,309,269,291]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="51" pageNumber="302">Fig. 125</figureCitation>
. Body larger:
<quantity id="EE0D9B7EFFDA6D6BD7F6D83ABC4D0761" box="[444,586,269,290]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="51" pageNumber="302" unit="mm" value="2.1" valueMax="2.4" valueMin="1.8">1.80–2.40 mm</quantity>
.
<collectingCountry id="51E2760BFFDA6D6BD419D83ABCB70760" box="[595,688,269,291]" name="Indonesia" pageId="51" pageNumber="302">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFDA6D6BD50CD83ABD8A0760" box="[838,909,269,291]" name="Guinea-Bissau" pageId="51" pageNumber="302">Guinea</collectingCountry>
).............................
<taxonomicName id="EEF54D18FFDA6D6BD340D83ABB660761" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1290,1377,269,290]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD340D83ABB660761" box="[1290,1377,269,290]" italics="true" pageId="51" pageNumber="302">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD83ABB9B0761" bold="true" box="[1382,1436,269,290]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD83ABB9B0761" box="[1382,1436,269,290]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD81DBB9B071E" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
22 Body dark brown. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD46DD81DBC64077C" box="[551,611,298,320]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">Fig. 3</figureCitation>
. Body distinctly larger:
<quantity id="EE0D9B7EFFDA6D6BD51ED81DBDA90703" box="[852,942,298,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="51" pageNumber="302" unit="mm" value="2.75">2.75 mm</quantity>
. Aedeagus as in
<figureCitation id="B1CE2A1EFFDA6D6BD21CD81DBAA70703" box="[1110,1184,298,320]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="51" pageNumber="302">Fig. 16</figureCitation>
.
<collectingCountry id="51E2760BFFDA6D6BD2E1D81DBB0F0703" box="[1195,1288,298,320]" name="Indonesia" pageId="51" pageNumber="302">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFDA6D6BD6A8D870BF2E071E" box="[226,297,327,349]" name="Guinea-Bissau" pageId="51" pageNumber="302">Guinea</collectingCountry>
)...........................................................................
<taxonomicName id="EEF54D18FFDA6D6BD280D87FBB66071E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1226,1377,328,349]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD280D87FBB66071E" box="[1226,1377,328,349]" italics="true" pageId="51" pageNumber="302">P. atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD87FBB9B071E" bold="true" box="[1382,1436,328,349]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD87FBB9B071E" box="[1382,1436,328,349]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD852BB9B0739" blockId="51.[151,1437,151,877]" box="[151,1436,357,379]" pageId="51" pageNumber="302">- Body paler and smaller............................................................................... 23</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD8B5BB9B07F6" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
23 Median lobe wide, with widely rounded apex (
<figureCitation id="B1CE2A1EFFDA6D6BD4EDD8B5BCF707DB" box="[679,752,386,408]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="51" pageNumber="302">Fig. 68</figureCitation>
). Body reddish brown, with paler elytra. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD34ED8B5BB4B07D4" box="[1284,1356,386,408]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="51" pageNumber="302">Fig. 41</figureCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDA6D6BD789D8A8BC0D07F6" box="[451,522,415,437]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="51" pageNumber="302">Fig. 74</figureCitation>
. Body length:
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.
<collectingCountry id="51E2760BFFDA6D6BD567D897BDE107F6" box="[813,998,415,437]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
....................
<taxonomicName id="EEF54D18FFDA6D6BD2B8D897BB6607F7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1266,1377,415,437]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="latissima" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD2B8D897BB6607F7" box="[1266,1377,415,437]" italics="true" pageId="51" pageNumber="302">P. latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD897BB9B07F6" bold="true" box="[1382,1436,416,437]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD897BB9B07F6" box="[1382,1436,416,437]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD88ABB9B0791" blockId="51.[151,1437,151,877]" box="[151,1436,445,467]" pageId="51" pageNumber="302">- Median lobe distinctly narrower, with narrower apex. Coloration different....................................... 24</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD8EDBB9B07B3" blockId="51.[151,1437,151,877]" box="[151,1436,474,496]" pageId="51" pageNumber="302">24 Median lobe wide; parameres with distinctly broadened apical portions......................................... 25</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD8CFBB9B044E" blockId="51.[151,1437,151,877]" box="[151,1436,503,525]" pageId="51" pageNumber="302">- Median lobe and apical portion of parameres narrower...................................................... 26</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDDB22BB9B0426" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
25 Pronotum strongly transverse. Apical portions of parameres very wide (
<figureCitation id="B1CE2A1EFFDA6D6BD5D0DB22BDE40469" box="[922,995,533,555]" captionStart="FIGURES 92–95" captionStartId="33.[151,264,1780,1804]" captionTargetBox="[160,1427,189,1747]" captionTargetId="figure-16@33.[151,1436,181,1756]" captionTargetPageId="33" captionText="FIGURES 92–95. Paraphloeostiba papuana (Western New Guinea): 92–aedeagus, parameral view, 93–aedeagus, lateral view, 94–male abdominal tergite VIII, 95–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821261" httpUri="https://zenodo.org/record/10821261/files/figure.png" pageId="51" pageNumber="302">Fig. 92</figureCitation>
). Body yellow-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD303DB22BB850468" box="[1353,1410,533,555]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">
Fig.
<quantity id="EE0D9B7EFFDA6D6BD33CDB22BB850468" box="[1398,1410,533,555]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="51" pageNumber="302" unit="in" value="5.0">5</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDA6D6BD6A8DB05BF86040B" author="Shavrin, A. V." box="[226,385,562,584]" pageId="51" pageNumber="302" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDA6D6BD4FCDB05BCFD0404" box="[694,762,562,584]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="51" pageNumber="302">
Fig.
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</figureCitation>
in
<bibRefCitation id="4D644B6AFFDA6D6BD552DB05BDB0040B" author="Shavrin, A. V." box="[792,951,562,584]" pageId="51" pageNumber="302" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Body smaller:
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.
<collectingCountry id="51E2760BFFDA6D6BD2A9DB05BB47040B" box="[1251,1344,562,584]" name="Indonesia" pageId="51" pageNumber="302">Indonesia</collectingCountry>
(Western New
<collectingCountry id="51E2760BFFDA6D6BD75FDB78BF590426" box="[277,350,591,613]" name="Guinea-Bissau" pageId="51" pageNumber="302">Guinea</collectingCountry>
),
<collectingCountry id="51E2760BFFDA6D6BD724DB67BC200426" box="[366,551,591,613]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
.................................................
<taxonomicName id="EEF54D18FFDA6D6BD2CEDB67BB9B0426" authority="(Cameron, 1931)" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[1156,1436,592,613]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFDA6D6BD2CEDB67BA940426" box="[1156,1171,592,613]" italics="true" pageId="51" pageNumber="302">P</emphasis>
.
<emphasis id="1B81EA89FFDA6D6BD2D4DB67BAF50426" box="[1182,1266,592,613]" italics="true" pageId="51" pageNumber="302">papuana</emphasis>
(
<bibRefCitation id="4D644B6AFFDA6D6BD2B5DB67BB920426" author="Cameron, M." box="[1279,1429,592,613]" pageId="51" pageNumber="302" pagination="355 - 364" refId="ref35403" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article" year="1931">Cameron, 1931</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDDB5ABB9B04FE" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
- Pronotum slightly transverse. Apical portions of parameres distinctly narrower (
<figureCitation id="B1CE2A1EFFDA6D6BD597DB5ABA3604C0" box="[989,1073,621,643]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="51" pageNumber="302">Fig. 126</figureCitation>
). Body reddish-brown, with distinctly paler elytra. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD794DBBDBC2204DC" box="[478,549,650,672]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="51" pageNumber="302">Fig. 80</figureCitation>
. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFDA6D6BD516DBBDBDA804DC" box="[860,943,650,672]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="51" pageNumber="302">Fig. 132</figureCitation>
. Body larger: 2.00–
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.
<collectingCountry id="51E2760BFFDA6D6BD287DBBCBB8104E3" box="[1229,1414,650,672]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
.........................................................................................
<taxonomicName id="EEF54D18FFDA6D6BD344DB9FBB6604FF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1294,1377,679,701]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD344DB9FBB6604FF" box="[1294,1377,679,701]" italics="true" pageId="51" pageNumber="302">P. rufula</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFDA6D6BD32CDB9FBB9B04FE" box="[1382,1436,680,701]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDDBF2BB9B0556" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
26 Basal portion of aedeagus narrow; median lobe moderately wide and long; parameres narrow, distinctly not reaching apex of median lobe (
<figureCitation id="B1CE2A1EFFDA6D6BD729DBD5BFAF04BB" box="[355,424,738,760]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="51" pageNumber="302">
Fig.
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</figureCitation>
in
<bibRefCitation id="4D644B6AFFDA6D6BD78FDBD5BC6B04BB" author="Shavrin, A. V." box="[453,620,738,760]" pageId="51" pageNumber="302" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b))</bibRefCitation>
. Body reddish-brown, with paler elytra. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD227DBD5BAA304BB" box="[1133,1188,738,760]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">
Fig.
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</figureCitation>
in
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. Body length:
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..............................................
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Shavrin, 2023
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- Basal portion of aedeagus wide; median lobe very narrow, moderately short; parameres distinctly exceeding apex of median lobe, with broadened apical portions (
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). Body yellow-brown to reddish-brown. Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD29BDA0DBB080513" box="[1233,1295,826,848]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">Fig. 5</figureCitation>
. Body length:
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..........................................................
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The genus
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<bibRefCitation id="4D644B6AFFDA6D6BD79BDAF7BC980599" author="Bernhauer, M." box="[465,671,960,986]" pageId="51" pageNumber="302" pagination="311 - 327" refId="ref35348" refString="Bernhauer, M. (1926) Fauna Buruana Staphylinidae. Zur Staphylinidenfauna der Molukken. Treubia, 7 (3), 311 - 327." type="journal article" year="1926">Bernhauer, 1926</bibRefCitation>
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The genus
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<emphasis id="1B81EA89FFDA6D6BD75ADD3FBFAD0261" box="[272,426,1032,1058]" italics="true" pageId="51" pageNumber="302">Xanthonomus</emphasis>
<bibRefCitation id="4D644B6AFFDA6D6BD7E5DD3FBC6F0261" author="Bernhauer, M." box="[431,616,1032,1058]" pageId="51" pageNumber="302" pagination="311 - 327" refId="ref35348" refString="Bernhauer, M. (1926) Fauna Buruana Staphylinidae. Zur Staphylinidenfauna der Molukken. Treubia, 7 (3), 311 - 327." type="journal article" year="1926">Bernhauer, 1926</bibRefCitation>
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was originally described as the subgenus of the genus
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<emphasis id="1B81EA89FFDA6D6BD288DD3FBB5D0261" box="[1218,1370,1032,1058]" italics="true" pageId="51" pageNumber="302">Phloeonomus</emphasis>
Heer, 1839
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with
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. (
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)
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(Indionesia: Buru), fixed by the monotypy (
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). Steel (1995) raised this taxon to the generic status, redescribed it and
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, and described
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.
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<bibRefCitation id="4D644B6AFFDA6D6BD6DDDD43BF1102CC" author="Steel, W. O." box="[151,278,1140,1167]" pageId="51" pageNumber="302" pagination="275 - 278" refId="ref36054" refString="Steel, W. O. (1955) Notes on the Omaliinae (Col., Staphylinidae). 7. The genus Xanthonomus Bernhauer. The Entomologist's Monthly Magazine, 91, 275 - 278." type="journal article" year="1955">Steel, 1955</bibRefCitation>
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(
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: Gunong Tahan Padang). Additional two closely related species, shorter in the length of the body, are described below. Species of this genus are characterized by the small body, sometimes with paler elytra, distinctly transverse apical four antennomeres forming loose club, transverse pronotum with obtuse or widely rounded hind angles, short elytra, the absence of the distinct paratergites of the abdomen, and the narrow median lobe and parameres.
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<mods:identifier id="E8D63846D6B6B2F7A6DE5DEA0A2E8B98" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="8B921ACFEA41FBA0BBAC3E3DFB9DC257" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFDA6D6ED6DDDC58BF37078D" ID-DOI="http://doi.org/10.5281/zenodo.10821321" ID-Zenodo-Dep="10821321" LSID="urn:lsid:plazi:treatment:A15C878DFFDA6D6ED6DDDC58BF37078D" httpUri="http://treatment.plazi.org/id/A15C878DFFDA6D6ED6DDDC58BF37078D" lastPageId="54" lastPageNumber="305" pageId="51" pageNumber="302">
<subSubSection id="61EF6510FFDA6D6BD6DDDC58BC1303C9" box="[151,532,1391,1418]" pageId="51" pageNumber="302" type="nomenclature">
<paragraph id="294A369BFFDA6D6BD6DDDC58BC1303C9" blockId="51.[151,532,1391,1455]" box="[151,532,1391,1418]" pageId="51" pageNumber="302">
<heading id="720281F7FFDA6D6BD6DDDC58BC1303C9" bold="true" box="[151,532,1391,1418]" fontSize="11" level="1" pageId="51" pageNumber="302" reason="1">
<emphasis id="1B81EA89FFDA6D6BD6DDDC58BC1303C9" bold="true" box="[151,532,1391,1418]" pageId="51" pageNumber="302">
<taxonomicName id="EEF54D18FFDA6D6BD6DDDC58BFCC03C9" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,459,1391,1418]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
<emphasis id="1B81EA89FFDA6D6BD6DDDC58BFCC03C9" bold="true" box="[151,459,1391,1418]" italics="true" pageId="51" pageNumber="302">Xanthonomus guineensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFDA6D6BD798DC47BC1303C9" box="[466,532,1392,1418]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDA6D6BD6DDDCA3BF7C03EC" box="[151,379,1428,1455]" pageId="51" pageNumber="302" type="description">
<paragraph id="294A369BFFDA6D6BD6DDDCA3BF7C03EC" blockId="51.[151,532,1391,1455]" box="[151,379,1428,1455]" pageId="51" pageNumber="302">
(
<figureCitation id="B1CE2A1EFFDA6D6BD6D5DCA3BF0303ED" box="[159,260,1428,1454]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="51" pageNumber="302">Figs 142</figureCitation>
,
<figureCitation id="B1CE2A1EFFDA6D6BD75ADCA3BF7403EC" box="[272,371,1428,1455]" captionStart-0="FIGURES 152–157" captionStart-1="FIGURE 158" captionStartId-0="52.[151,264,1589,1613]" captionStartId-1="53.[151,250,1956,1980]" captionTargetBox-0="[185,1403,415,1547]" captionTargetBox-1="[151,1436,1165,1932]" captionTargetId-0="figure-117@52.[177,1411,415,1566]" captionTargetId-1="figure-421@53.[151,1436,1165,1932]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionText-0="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." captionText-1="FIGURE 158. Distribution of Xanthonomus: X. toxopeanus (circle), X. latus (triangle), X. guineensis (squares), X. thayerae (diamonds)." figureDoi-0="http://doi.org/10.5281/zenodo.10821297" figureDoi-1="http://doi.org/10.5281/zenodo.10821299" httpUri-0="https://zenodo.org/record/10821297/files/figure.png" httpUri-1="https://zenodo.org/record/10821299/files/figure.png" pageId="51" pageNumber="302">152–158</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDA6D6BD6DDDCEBBD20008D" pageId="51" pageNumber="302" type="materials_examined">
<paragraph id="294A369BFFDA6D6BD6DDDCEBBEFE007D" blockId="51.[151,1437,1500,2030]" pageId="51" pageNumber="302">
<emphasis id="1B81EA89FFDA6D6BD6DDDCEBBF4003B5" bold="true" box="[151,327,1500,1526]" pageId="51" pageNumber="302">Type material.</emphasis>
<typeStatus id="F64E8839FFDA6D6BD71BDCEBBFBE03B5" box="[337,441,1500,1526]" pageId="51" pageNumber="302" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFDA6D6BD788DCEBBFD003B5" box="[450,471,1500,1526]" count="1" pageId="51" pageNumber="302" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFDA6D6BD7A3DCEBBC4803B5" box="[489,591,1500,1526]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="51" pageNumber="302">Fig. 142</figureCitation>
): ‘
<collectingRegion id="EB31F879FFDA6D6BD423DCEBBD0B03B5" box="[617,780,1500,1526]" country="Indonesia" name="Papua Barat" pageId="51" pageNumber="302">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Bilogay,
<date id="5D4B105BFFDA6D6BD268DCEBBAB403B4" box="[1058,1203,1500,1527]" pageId="51" pageNumber="302" value="1995-12-29">29. 12. 1995</date>
|
<quantity id="EE0D9B7EFFDA6D6BD281DCEBBB5D03B5" box="[1227,1370,1500,1526]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="51" pageNumber="302" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFDA6D6BD7D1DF37BC330059" box="[411,564,1536,1562]" pageId="51" pageNumber="302" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFDA6D6BD407DF37BCE00059" authorityName="Bernhauer" authorityYear="1926" box="[589,743,1536,1562]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFDA6D6BD407DF37BCE00059" box="[589,743,1536,1562]" italics="true" pageId="51" pageNumber="302">Xanthonomus</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFDA6D6BD54BDF36BD700059" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[769,887,1537,1562]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
<emphasis id="1B81EA89FFDA6D6BD54BDF36BD700059" box="[769,887,1537,1562]" italics="true" pageId="51" pageNumber="302">guineensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD5CBDF37BDC30059" bold="true" box="[897,964,1536,1562]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD5CBDF37BDC30059" box="[897,964,1536,1562]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFDA6D6BD21CDF37BA6B0059" box="[1110,1132,1536,1562]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="51" pageNumber="302" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFDA6D6BD6D5DF13BEEA007D" box="[159,237,1572,1598]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="302">NMW</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFDA6D6BD68DDF7FBD20008D" blockId="51.[151,1437,1500,2030]" pageId="51" pageNumber="302">
<typeStatus id="F64E8839FFDA6D6BD68DDF7FBF3F0021" box="[199,312,1608,1634]" pageId="51" pageNumber="302" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFDA6D6BD70BDF7FBF780021" box="[321,383,1608,1635]" count="5" pageId="51" pageNumber="302" type="male">5 ♂♂</specimenCount>
(
<specimenCount id="3FF3FD12FFDA6D6BD7C6DF7FBC210021" box="[396,550,1608,1634]" count="1" pageId="51" pageNumber="302" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="3FF3FD12FFDA6D6BD4EDDF7FBCE20021" box="[679,741,1608,1634]" count="3" pageId="51" pageNumber="302" type="female">3 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFDA6D6BD4B8DF7FBD8B0021" box="[754,908,1608,1634]" count="1" pageId="51" pageNumber="302" type="generic">one specimen</specimenCount>
dissected): same data as the holotype (
<specimenCount id="3FF3FD12FFDA6D6BD373DF7FBB660021" box="[1337,1377,1608,1634]" count="1" pageId="51" pageNumber="302" type="male">1 ♂</specimenCount>
,
<specimenCount id="3FF3FD12FFDA6D6BD326DF7FBB910021" box="[1388,1430,1608,1634]" count="1" pageId="51" pageNumber="302" type="female">1 ♀</specimenCount>
: cSh;
<specimenCount id="3FF3FD12FFDA6D6BD69ADF5BBF1600C5" box="[208,273,1644,1670]" count="4" pageId="51" pageNumber="302" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFDA6D6BD757DF5BBF5900C5" box="[285,350,1644,1670]" count="2" pageId="51" pageNumber="302" type="female">2 ♀♀</specimenCount>
:
<collectionCode id="4FE4AE5EFFDA6D6BD721DF5BBFBF00C5" box="[363,440,1644,1670]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="302">NMW</collectionCode>
);
<specimenCount id="3FF3FD12FFDA6D6BD781DF5BBC0B00C5" box="[459,524,1644,1670]" count="3" pageId="51" pageNumber="302" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFDA6D6BD452DF5BBC4500C5" box="[536,578,1644,1670]" count="1" pageId="51" pageNumber="302" type="female">1 ♀</specimenCount>
(dissected): ‘
<collectingRegion id="EB31F879FFDA6D6BD49CDF5BBD7000C5" box="[726,887,1644,1670]" country="Indonesia" name="Papua Barat" pageId="51" pageNumber="302">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Sinak, trail to Ilaga |
<quantity id="EE0D9B7EFFDA6D6BD34CDF5BBB9F00C4" box="[1286,1432,1644,1671]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="51" pageNumber="302" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100-2500m</quantity>
,
<date id="5D4B105BFFDA6D6BD6DDDFA7BF2F00E8" box="[151,296,1680,1707]" pageId="51" pageNumber="302" value="1995-12-15">15. 12. 1995</date>
| leg. A. Riedel’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFDA6D6BD438DFA7BCC700E9" box="[626,704,1680,1706]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="302">NMW</collectionCode>
). All
<typeStatus id="F64E8839FFDA6D6BD54BDFA7BD6B00E9" box="[769,876,1680,1706]" pageId="51" pageNumber="302" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFDA6D6BD2B7DFA7BB8A00E9" box="[1277,1421,1680,1706]" pageId="51" pageNumber="302" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFDA6D6BD6DDDF83BF36008D" authorityName="Bernhauer" authorityYear="1926" box="[151,305,1716,1742]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFDA6D6BD6DDDF83BF36008D" box="[151,305,1716,1742]" italics="true" pageId="51" pageNumber="302">Xanthonomus</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFDA6D6BD70FDF82BFBC008D" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[325,443,1717,1742]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
<emphasis id="1B81EA89FFDA6D6BD70FDF82BFBC008D" box="[325,443,1717,1742]" italics="true" pageId="51" pageNumber="302">guineensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD788DF83BC06008D" bold="true" box="[450,513,1716,1742]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD788DF83BC06008D" box="[450,513,1716,1742]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFDA6D6BD4C1DF83BCA6008D" box="[651,673,1716,1742]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="51" pageNumber="302" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDA6D6ED68DDFEFBF1506BB" lastPageId="54" lastPageNumber="305" pageId="51" pageNumber="302" type="description">
<paragraph id="294A369BFFDA6D6BD68DDFEFBF8C011D" blockId="51.[151,1437,1500,2030]" pageId="51" pageNumber="302">
<emphasis id="1B81EA89FFDA6D6BD68DDFEFBF5E00B1" bold="true" box="[199,345,1752,1778]" pageId="51" pageNumber="302">Description.</emphasis>
Measurements (n=13): HW: 0.40–0.46; HL: 0.20–0.25; OL: 0.12–0.14; TL: 0.02–0.03; AL (
<typeStatus id="F64E8839FFDA6D6BD6D5DFCBBF0F0155" box="[159,264,1788,1814]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
): 0.52; PL: 0.29–0.38; PWmax: 0.53–0.57; PWmin: 0.49–0.52; ESL: 0.45–0.52; EW: 0.66–0.73; MTbL (
<typeStatus id="F64E8839FFDA6D6BD6D5DE17BF0F0179" box="[159,264,1824,1850]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
): 0.35; MTrL (
<typeStatus id="F64E8839FFDA6D6BD7E4DE17BC100179" box="[430,535,1824,1850]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
): 0.24 (MTrL 1–4: 0.07; MTrL 5: 0.17); AW: 0.75–0.80; AedL: 0.45; BL: 1.87– 2.25 (
<typeStatus id="F64E8839FFDA6D6BD69DDE73BF38011D" box="[215,319,1860,1886]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
: 2.04).
</paragraph>
<paragraph id="294A369BFFDA6D6CD68DDE5FBF9007C4" blockId="51.[151,1437,1500,2030]" lastBlockId="52.[151,1437,151,391]" lastPageId="52" lastPageNumber="303" pageId="51" pageNumber="302">
Habitus as in
<figureCitation id="B1CE2A1EFFDA6D6BD714DE5FBFB901C1" box="[350,446,1896,1922]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="51" pageNumber="302">Fig. 142</figureCitation>
. Body dark brown, with slightly paler elytra and sometimes abdominal tergites VII–VIII; antennomeres 7–11 and femora brown; tibia yellow-brown, with paler apical portion; mouthparts, antennomeres 1–6 and tarsi yellow. Head with very fine, indistinct and dense punctation, finer and more indistinct on infraorbital portions; neck without visible punctures; pronotum with very dense and fine punctation, missing in mediobasal portion; scutellum without punctures; elytra with very fine and dense punctation similar to that in pronotum, usually finer and sparser around scutellum and along suture; abdomen with very dense and fine punctation. Head with dense and coarse microreticulation, transverse on clypeus and isodiametric in middle; neck with moderately fine transverse microreticulation; pronotum with very dense and fine isodiametric meshes about as that in middle portion of head; scutellum with dense and coarse transverse microsculpture; elytra with fine and sometimes indistinct isodiametric microreticulation, sometimes indistinct along suture; abdomen with very dense and moderately coarse isodiametric sculpture.
</paragraph>
<caption id="7D8A6613FFDD6D6CD6DDDF02BFD800D6" ID-DOI="http://doi.org/10.5281/zenodo.10821297" ID-Zenodo-Dep="10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="52" pageNumber="303" startId="52.[151,264,1589,1613]" targetBox="[185,1403,415,1547]" targetPageId="52" targetType="figure">
<paragraph id="294A369BFFDD6D6CD6DDDF02BFD800D6" blockId="52.[151,1437,1589,1686]" pageId="52" pageNumber="303">
<emphasis id="1B81EA89FFDD6D6CD6DDDF02BF68000E" bold="true" box="[151,367,1589,1613]" pageId="52" pageNumber="303">FIGURES 152–157.</emphasis>
<taxonomicName id="EEF54D18FFDD6D6CD73DDF02BC72000E" authorityName="Shavrin" authorityYear="2024" box="[375,629,1589,1613]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="303" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="1B81EA89FFDD6D6CD73DDF02BC72000E" box="[375,629,1589,1613]" italics="true" pageId="52" pageNumber="303">Xanthonomus guineensis</emphasis>
</taxonomicName>
(paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFDD6D6DD68DDFD0BFE907CB" blockId="52.[151,1436,1767,2044]" lastBlockId="53.[151,1437,151,1142]" lastPageId="53" lastPageNumber="304" pageId="52" pageNumber="303">
Head distinctly transverse, 1.8 times to twice as broad as long, with slightly elevated infraorbital portions and shallow oval anteriomedian depressions, indistinct in some
<typeStatus id="F64E8839FFDD6D6CD524DE3CBDDA0166" box="[878,989,1803,1829]" pageId="52" pageNumber="303" type="paratype">paratypes</typeStatus>
; anteocellar foveae very narrow, linear, reaching level of apical third or about middle length of eyes; postocular parts very short, from indistinct and obtuse postocular carina strongly narrowed posteriad. Eyes very large, convex. Ocelli moderately large, located slightly below level of postocular carina; distance between ocelli slightly longer or about as long as distance between ocellus and posterior margin of eye. Maxillary palpi long; preapical palpomere about twice as long as wide; apical palpomere narrow, slightly longer than preapical segment, from preapical portion gradually narrowed apicad toward rounded or subacute apex. Antennomeres short, reaching basal third of pronotum when reclined, with distinctly transverse antennomere 7 and strongly transverse antennomeres 8–10; antennomeres 6–10 gradually widened apicad; four apical antennomere forming distinct loose club; basal antennomere wide, slightly more than two and a half times as long as wide, antennomere 2 about twice shorter than basal antennomere, 3 slightly shorter and about twice narrower than 2, 4 about twice shorter than 3, 5 slightly longer and distinctly broader than 4, 6 distinctly shorter and broader than 5, 7 distinctly shorter and broader than 6, 8 three times broader than 6, 9–10 about twice longer and distinctly broader than 8, apical antennomere about 1.3–1.5 times as long as 10, from middle gradually rounded toward rounded apex.
</paragraph>
<paragraph id="294A369BFFDC6D6DD68DD8A6BCB90455" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Pronotum slightly convex, 1.5–1.8 times as broad as long, distinctly broader than head, widest in middle, slightly more narrowed posteriad than anteriad; apical and hind angles widely rounded; anterior margin straight, about as long as rounded posterior margin; median part without longitudinal impressions; lateral portions narrowly bordered; laterobasal portion without depression.</paragraph>
<paragraph id="294A369BFFDC6D6DD68DDB17BC8F04C2" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Elytra 1.4 times as long as broad, short, reaching basal or middle portion of abdominal tergite III, 1.3–1.5 times as long as pronotum, slightly broadened posteriad, with slightly concave lateral margins and striagth or rounded apical margins. Hind wings fully developed.</paragraph>
<paragraph id="294A369BFFDC6D6DD68DDBBCBF89048A" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Metatarsus about 1.4 times as long as metatibia; ventral surface of protarsomeres 1–4 with several rows of modified tenent setae.</paragraph>
<paragraph id="294A369BFFDC6D6DD68DDBE5BCB40577" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Abdomen slightly broadened toward andominal tergite IV or V, distinctly broadened than elytra, with two very small and sometimes indistinct oval wing-folding patches in middle of abdominal tergite IV and narrow palisade fringe at apical margin of abdominal tergite VII.</paragraph>
<paragraph id="294A369BFFDC6D6DD68DDA0ABD43026C" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">
Male. Femora very wide; tibia significantly broadened in middle; tarsomeres 1–4 slightly broadened, with very long tenent setae. Posterior margin of abdominal tergite VIII slightly sinuate (
<figureCitation id="B1CE2A1EFFDC6D6DD20DDA56BAAC053F" box="[1095,1195,865,892]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 154</figureCitation>
). Posterior margin of abdominal sternite VIII slighly sinuate, with row of very short setae on apical margin (
<figureCitation id="B1CE2A1EFFDC6D6DD23FDAB2BADE05E3" box="[1141,1241,901,928]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 155</figureCitation>
). Aedeagus long, with very wide basal portion, gradually narrowed toward small and rounded apex; parameres narrow, distinctly not reaching apex of median lobe, with two long apical and two preapical setae; internal sac long and moderately wide, with two very long, narrow and curved sclerotized structure, stretching from basal to about middle part of median lobe (
<figureCitation id="B1CE2A1EFFDC6D6DD69DDD23BF3E026C" box="[215,313,1044,1071]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 152</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFDC6D6DD4AADD23BD47026C" box="[736,832,1044,1071]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 153</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFDC6D6DD68DDD0FBABD0236" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">
Female. Femora, tibia and tarsomeres 1–4 moderately narrow. Posterior margin of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFDC6D6DD325DD0FBEC50235" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 156</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFDC6D6DD7D3DD6CBFFE0235" box="[409,505,1115,1142]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 157</figureCitation>
) trunctate. Accessory sclerite and spermatheca not recognized.
</paragraph>
<caption id="7D8A6613FFDC6D6DD6DDDE93BF1501A2" ID-DOI="http://doi.org/10.5281/zenodo.10821299" ID-Zenodo-Dep="10821299" httpUri="https://zenodo.org/record/10821299/files/figure.png" pageId="53" pageNumber="304" startId="53.[151,250,1956,1980]" targetBox="[151,1436,1165,1932]" targetPageId="53" targetType="figure">
<paragraph id="294A369BFFDC6D6DD6DDDE93BF1501A2" blockId="53.[151,1437,1956,2017]" pageId="53" pageNumber="304">
<emphasis id="1B81EA89FFDC6D6DD6DDDE93BF2801FF" bold="true" box="[151,303,1956,1980]" pageId="53" pageNumber="304">FIGURE 158.</emphasis>
Distribution of
<taxonomicName id="EEF54D18FFDC6D6DD79DDE92BC6501FE" authorityName="Bernhauer" authorityYear="1926" box="[471,610,1957,1981]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFDC6D6DD79DDE92BC6501FE" box="[471,610,1957,1981]" italics="true" pageId="53" pageNumber="304">Xanthonomus</emphasis>
</taxonomicName>
:
<taxonomicName id="EEF54D18FFDC6D6DD438DE92BD0401FF" authorityName="Bernhauer" authorityYear="1926" box="[626,771,1957,1980]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="toxopeanus">
<emphasis id="1B81EA89FFDC6D6DD438DE92BD0401FF" box="[626,771,1957,1980]" italics="true" pageId="53" pageNumber="304">X. toxopeanus</emphasis>
</taxonomicName>
(circle),
<taxonomicName id="EEF54D18FFDC6D6DD52EDE92BDB401FE" authorityName="Steel" authorityYear="1955" box="[868,947,1957,1981]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="1B81EA89FFDC6D6DD52EDE92BDB401FE" box="[868,947,1957,1981]" italics="true" pageId="53" pageNumber="304">X. latus</emphasis>
</taxonomicName>
(triangle),
<taxonomicName id="EEF54D18FFDC6D6DD262DE92BAB501FF" authorityName="Shavrin" authorityYear="2024" box="[1064,1202,1957,1980]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="guineensis">
<emphasis id="1B81EA89FFDC6D6DD262DE92BAB501FF" box="[1064,1202,1957,1980]" italics="true" pageId="53" pageNumber="304">X. guineensis</emphasis>
</taxonomicName>
(squares),
<taxonomicName id="EEF54D18FFDC6D6DD36FDE92BB9A01FE" authorityName="Shavrin" authorityYear="2024" box="[1317,1437,1957,1981]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="thayerae">
<emphasis id="1B81EA89FFDC6D6DD36FDE92BB9A01FE" box="[1317,1437,1957,1981]" italics="true" pageId="53" pageNumber="304">X. thayerae</emphasis>
</taxonomicName>
(diamonds).
</paragraph>
</caption>
<paragraph id="294A369BFFDF6D6ED68DD9A1BF1506BB" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DD9A1BFB506F3" bold="true" box="[199,434,150,176]" pageId="54" pageNumber="305">Comparative notes.</emphasis>
Based on the general shape and sculpture of the body
<taxonomicName id="EEF54D18FFDF6D6ED260D9A0BAC606F3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1066,1217,151,176]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="guineensis" status="sp. n.">
<emphasis id="1B81EA89FFDF6D6ED260D9A0BAC606F3" box="[1066,1217,151,176]" italics="true" pageId="54" pageNumber="305">X. guineensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED280D9A1BB0B06F3" bold="true" box="[1226,1292,150,176]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED280D9A1BB0B06F3" box="[1226,1292,150,176]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFDF6D6ED6DDD98CBF1D0697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,282,186,212]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="thayerae" status="sp. n.">
<emphasis id="1B81EA89FFDF6D6ED6DDD98CBF1D0697" box="[151,282,186,212]" italics="true" pageId="54" pageNumber="305">X. thayerae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED768D98DBF640697" bold="true" box="[290,355,186,212]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED768D98DBF640697" box="[290,355,186,212]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the slightly larger body, wider median lobe and shorter parameres.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDF6D6ED68DD836BF850703" pageId="54" pageNumber="305" type="distribution">
<paragraph id="294A369BFFDF6D6ED68DD836BF850703" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DD836BF660758" bold="true" box="[199,353,257,283]" pageId="54" pageNumber="305">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFDF6D6ED720D836BC840758" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[362,643,257,283]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
<emphasis id="1B81EA89FFDF6D6ED720D836BC840758" box="[362,643,257,283]" italics="true" pageId="54" pageNumber="305">Xanthonomus guineensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED4C6D835BCC9075F" bold="true" box="[652,718,258,284]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED4C6D835BCC9075F" box="[652,718,258,284]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFDF6D6ED5A7D835BA1A075F" box="[1005,1053,258,284]" pageId="54" pageNumber="305">type</typeStatus>
locality in Western New
<collectingCountry id="51E2760BFFDF6D6ED30ED836BB9F0758" box="[1348,1432,257,283]" name="Guinea-Bissau" pageId="54" pageNumber="305">Guinea</collectingCountry>
,
<collectingCountry id="51E2760BFFDF6D6ED6DDD812BF03077C" box="[151,260,293,319]" name="Indonesia" pageId="54" pageNumber="305">Indonesia</collectingCountry>
(
<figureCitation id="B1CE2A1EFFDF6D6ED759D812BF710703" box="[275,374,293,320]" captionStart="FIGURE 158" captionStartId="53.[151,250,1956,1980]" captionTargetBox="[151,1436,1165,1932]" captionTargetId="figure-421@53.[151,1436,1165,1932]" captionTargetPageId="53" captionText="FIGURE 158. Distribution of Xanthonomus: X. toxopeanus (circle), X. latus (triangle), X. guineensis (squares), X. thayerae (diamonds)." figureDoi="http://doi.org/10.5281/zenodo.10821299" httpUri="https://zenodo.org/record/10821299/files/figure.png" pageId="54" pageNumber="305">Fig. 158</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDF6D6ED68DD87EBF37078D" pageId="54" pageNumber="305" type="etymology">
<paragraph id="294A369BFFDF6D6ED68DD87EBED207C4" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DD87EBF490720" bold="true" box="[199,334,329,355]" pageId="54" pageNumber="305">Etymology.</emphasis>
The specific epithet is the latinized adjective derived from the area the specimens were collected from.
</paragraph>
<paragraph id="294A369BFFDF6D6ED68DD8A7BF37078D" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DD8A7BF4C07E9" bold="true" box="[199,331,400,426]" pageId="54" pageNumber="305">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="EE0D9B7EFFDF6D6ED512D8A7BA0A07E8" box="[856,1037,400,427]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="54" pageNumber="305" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
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167
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Key to species of the genus
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<emphasis id="1B81EA89FFDE6D6FD7A6DF55BC90003E" bold="true" box="[492,663,1634,1661]" italics="true" pageId="55" pageNumber="306">Xanthonomus</emphasis>
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1 Pronotum with distinct obtuse hind angles. Abdomen very wide. Habitus as in
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Fig.
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. Body length:
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<emphasis id="1B81EA89FFDE6D6FD2B6DFF2BB2C0099" box="[1276,1323,1733,1754]" italics="true" pageId="55" pageNumber="306">latus</emphasis>
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<keyLead id="920495AEFFDE6D6FD6DDDFD4BB9B00B4" box="[151,1436,1762,1784]" pageId="55" pageNumber="306">- Pronotum with widely rounded hind angles. Abdomen distinctly narrower........................................ 2</keyLead>
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2 Pronotum slightly transverse, with two median, shallow and oval impressions. Elytra yellow. Habitus as in
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Fig.
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. Body larger:
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. Aedeagus as in
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Fig.
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(Buru)...................................................................................................
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<emphasis id="1B81EA89FFDE6D6FD230DE0CBAE00113" box="[1146,1255,1851,1872]" italics="true" pageId="55" pageNumber="306">toxopeanus</emphasis>
(
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)
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<keyLead id="920495AEFFDE6D6FD6DDDE6FBB9B012E" box="[151,1436,1880,1902]" pageId="55" pageNumber="306">- Pronotum distinctly transverse, without median impressions. Elytra darker. Body smaller............................ 3</keyLead>
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<paragraph id="294A369BFFDE6D6FD6DDDE42BB9B01E4" blockId="55.[151,1437,1703,2019]" pageId="55" pageNumber="306">
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3 Median lobe moderately wide; parameres distinctly not exceeding apex of median lobe (
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). Habitus asi n
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. Body larger:
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.
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(New
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)..............................................
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X.
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<keyLead id="920495AEFFDE6D6FD6DDDE87BB9B01A1" pageId="55" pageNumber="306">
- Median lobe narrow; parameres reaching apex of median lobe (
<figureCitation id="B1CE2A1EFFDE6D6FD574DE87BD950186" box="[830,914,1967,1990]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="55" pageNumber="306">Fig. 159</figureCitation>
). Habitus as in
<figureCitation id="B1CE2A1EFFDE6D6FD26EDE87BA710186" box="[1060,1142,1968,1990]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="55" pageNumber="306">Fig. 143</figureCitation>
. Body smaller:
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.
<collectingCountry id="51E2760BFFDE6D6FD6A8DEFABF9C01A0" box="[226,411,1997,2019]" name="Papua New Guinea" pageId="55" pageNumber="306">Papua New Guinea</collectingCountry>
......................................................................
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<emphasis id="1B81EA89FFDE6D6FD2B8DEF9BB0601A0" box="[1266,1281,1998,2019]" italics="true" pageId="55" pageNumber="306">X</emphasis>
.
<emphasis id="1B81EA89FFDE6D6FD346DEFABB6701A1" box="[1292,1376,1997,2018]" italics="true" pageId="55" pageNumber="306">thayerae</emphasis>
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<emphasis id="1B81EA89FFDF6D6ED6DDDB2DBFFB0476" bold="true" box="[151,508,538,565]" pageId="54" pageNumber="305">
<taxonomicName id="EEF54D18FFDF6D6ED6DDDB2DBFB50476" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,434,538,565]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="thayerae" status="sp. nov.">
<emphasis id="1B81EA89FFDF6D6ED6DDDB2DBFB50476" bold="true" box="[151,434,538,565]" italics="true" pageId="54" pageNumber="305">Xanthonomus thayerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFDF6D6ED7F3DB2CBFFB0476" box="[441,508,539,565]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDF6D6ED6DDDB08BF7C0419" box="[151,379,575,602]" pageId="54" pageNumber="305" type="description">
<paragraph id="294A369BFFDF6D6ED6DDDB08BF7C0419" blockId="54.[151,508,538,602]" box="[151,379,575,602]" pageId="54" pageNumber="305">
(
<figureCitation id="B1CE2A1EFFDF6D6ED6D5DB08BF03041A" box="[159,260,575,601]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="54" pageNumber="305">Figs 143</figureCitation>
,
<figureCitation id="B1CE2A1EFFDF6D6ED75ADB08BF740419" box="[272,371,575,602]" captionStart-0="FIGURE 158" captionStart-1="FIGURES 159–165" captionStartId-0="53.[151,250,1956,1980]" captionStartId-1="55.[151,264,1456,1480]" captionTargetBox-0="[151,1436,1165,1932]" captionTargetBox-1="[228,1379,199,1423]" captionTargetId-0="figure-421@53.[151,1436,1165,1932]" captionTargetId-1="figure-16@55.[201,1386,181,1432]" captionTargetPageId-0="53" captionTargetPageId-1="55" captionText-0="FIGURE 158. Distribution of Xanthonomus: X. toxopeanus (circle), X. latus (triangle), X. guineensis (squares), X. thayerae (diamonds)." captionText-1="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10821299" figureDoi-1="http://doi.org/10.5281/zenodo.10821301" httpUri-0="https://zenodo.org/record/10821299/files/figure.png" httpUri-1="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">158–165</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDF6D6ED6DDDBB0BFAA0516" pageId="54" pageNumber="305" type="materials_examined">
<paragraph id="294A369BFFDF6D6ED6DDDBB0BAD004AA" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED6DDDBB0BF4204E3" bold="true" box="[151,325,646,673]" pageId="54" pageNumber="305">Type material.</emphasis>
<typeStatus id="F64E8839FFDF6D6ED706DBB0BFB304E2" box="[332,436,647,673]" pageId="54" pageNumber="305" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFDF6D6ED7F1DBB0BFD704E2" box="[443,464,647,673]" count="1" pageId="54" pageNumber="305" type="male">♂</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFDF6D6ED429DBB0BCA004E2" box="[611,679,647,673]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="54" pageNumber="305">NEW</collectionCode>
<collectingCountry id="51E2760BFFDF6D6ED4E5DBB0BD1E04E2" box="[687,793,647,673]" name="Guinea-Bissau" pageId="54" pageNumber="305">GUINEA</collectingCountry>
| Mt. Michael |
<date id="5D4B105BFFDF6D6ED580DBB0BA5104E2" box="[970,1110,647,673]" pageId="54" pageNumber="305" value="1964-07-18">18.VII.1964</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFDF6D6ED2AEDBB0BAFA04E2" box="[1252,1277,647,673]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="54" pageNumber="305">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<taxonomicName id="EEF54D18FFDF6D6ED787DB9CBC680486" authorityName="Bernhauer" authorityYear="1926" box="[461,623,683,709]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="307" phylum="Arthropoda" rank="genus">Xanthonomus</taxonomicName>
| prob. | n.sp. | det. M.K. Thayer 198[printed]9’ &lt;handwritten in black&gt;, ‘
<typeStatus id="F64E8839FFDF6D6ED6D1DBF8BF3304AA" box="[155,308,719,745]" pageId="54" pageNumber="305" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFDF6D6ED702DBF8BFE504AA" authorityName="Bernhauer" authorityYear="1926" box="[328,482,719,745]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFDF6D6ED702DBF8BFE504AA" box="[328,482,719,745]" italics="true" pageId="54" pageNumber="305">Xanthonomus</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFDF6D6ED7BFDBF8BC5004AA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[501,599,719,745]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="thayerae" status="sp. nov.">
<emphasis id="1B81EA89FFDF6D6ED7BFDBF8BC5004AA" box="[501,599,719,745]" italics="true" pageId="54" pageNumber="305">thayerae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED414DBF8BC9904AA" bold="true" box="[606,670,719,745]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED414DBF8BC9904AA" box="[606,670,719,745]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFDF6D6ED56DDBF8BD3A04AA" box="[807,829,719,745]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="54" pageNumber="305" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFDF6D6ED23BDBF8BACC04AA" box="[1137,1227,719,745]" pageId="54" pageNumber="305">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDBC4BFAA0516" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
<typeStatus id="F64E8839FFDF6D6ED68DDBC4BF2E054E" box="[199,297,755,781]" pageId="54" pageNumber="305" type="paratype">Paratype</typeStatus>
<specimenCount id="3FF3FD12FFDF6D6ED778DBC4BF40054E" box="[306,327,755,781]" count="1" pageId="54" pageNumber="305" type="female">♀</specimenCount>
(
<figureCitation id="B1CE2A1EFFDF6D6ED71DDBC4BFBC054E" box="[343,443,755,781]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="54" pageNumber="305">Fig. 143</figureCitation>
): ‘
<collectionCode id="4FE4AE5EFFDF6D6ED79FDBC4BC1E054E" box="[469,537,755,781]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="54" pageNumber="305">NEW</collectionCode>
<collectingCountry id="51E2760BFFDF6D6ED468DBC4BC8B054E" box="[546,652,755,781]" name="Guinea-Bissau" pageId="54" pageNumber="305">GUINEA</collectingCountry>
| Okasa
<quantity id="EE0D9B7EFFDF6D6ED4BBDBC4BD3A054E" box="[753,829,755,781]" metricMagnitude="3" metricUnit="m" metricValue="1.09728" pageId="54" pageNumber="305" unit="ft" value="3600.0">3600ft</quantity>
, |
<date id="5D4B105BFFDF6D6ED512DBC4BDE5054D" box="[856,994,755,782]" pageId="54" pageNumber="305" value="1965-07-11">11.VII.1965</date>
, leaf | mould in pine forest’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFDF6D6ED6D7DA20BEB10572" box="[157,182,791,817]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="54" pageNumber="305">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFDF6D6ED4C6DA20BD1B0572" box="[652,796,791,817]" pageId="54" pageNumber="305" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFDF6D6ED565DA20BDCE0572" authorityName="Bernhauer" authorityYear="1926" box="[815,969,791,817]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFDF6D6ED565DA20BDCE0572" box="[815,969,791,817]" italics="true" pageId="54" pageNumber="305">Xanthonomus</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFDF6D6ED596DA20BA390572" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[988,1086,791,817]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="thayerae" status="sp. nov.">
<emphasis id="1B81EA89FFDF6D6ED596DA20BA390572" box="[988,1086,791,817]" italics="true" pageId="54" pageNumber="305">thayerae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED20EDA20BA840572" bold="true" box="[1092,1155,791,817]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED20EDA20BA840572" box="[1092,1155,791,817]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFDF6D6ED341DA20BB260572" box="[1291,1313,791,817]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="54" pageNumber="305" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFDF6D6ED70DDA0CBFA60516" box="[327,417,827,853]" pageId="54" pageNumber="305">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDF6D6ED68DDA69BF660106" pageId="54" pageNumber="305" type="description">
<paragraph id="294A369BFFDF6D6ED68DDA69BF1405A6" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DDA69BF5E053B" bold="true" box="[199,345,862,888]" pageId="54" pageNumber="305">Description.</emphasis>
Measurements (n=2): HW: 0.41–0.45; HL: 0.21–0.25; OL: 0.11–0.13; TL: 0.02; AL (
<typeStatus id="F64E8839FFDF6D6ED362DA68BB96053A" box="[1320,1425,863,889]" pageId="54" pageNumber="305" type="holotype">holotype</typeStatus>
): 0.45; PL: 0.31–0.35; PWmax: 0.53–0.60; PWmin: 0.44–0.50; ESL: 0.47–0.49; EW: 0.68–0.71; MTbL (
<typeStatus id="F64E8839FFDF6D6ED362DAB4BB9605DE" box="[1320,1425,899,925]" pageId="54" pageNumber="305" type="holotype">holotype</typeStatus>
): 0.33; MTrL (
<typeStatus id="F64E8839FFDF6D6ED77FDA90BF990582" box="[309,414,935,961]" pageId="54" pageNumber="305" type="holotype">holotype</typeStatus>
): 0.18 (MTrL 1–4: 0.07; MTrL 5: 0.11); AW: 0.70–0.72; AedL: 0.45; BL: 1.61–1.80 (
<typeStatus id="F64E8839FFDF6D6ED6D5DAFCBF0F05A6" box="[159,264,971,997]" pageId="54" pageNumber="305" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDAD8BCEB02DA" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
Habitus as in
<figureCitation id="B1CE2A1EFFDF6D6ED728DAD8BFC4024A" box="[354,451,1007,1033]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="54" pageNumber="305">Fig. 143</figureCitation>
. Body and antennomeres 7–11 dark brown, with slightly paler middle portion of elytra; femora and basal half of tibia reddish brown; mouthparts, antennomeres 1–6, apical half of tibia and tarsi yellowish. Head with very fine and dense, regular and fine punctation except of clypeus; pronotum with very dense indistinct and fine punctation; elytra with fine and very dense punctation, somewhat similar to that in pronotum, more distinct in basal portion; abdomen without distinct punctures.
</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDD94BF74030E" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">Head 1.8–1.9 times as broad as long, with very indistinct anteriomedian depressions; anteocellar foveae reaching level of apical third length of eyes; postocular carina absent. Distance between ocelli about as long as distance between ocellus and posterior margin of eye. Antennomere 2 slightly more than twice shorter than basal antennomere, 4–5 twice shorter than 3, 8 slightly shorter and distintly broader than 7, apical antennomere about 1.2 times as long as 10.</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDC60BA8A0332" blockId="54.[151,1437,646,2041]" box="[199,1165,1367,1393]" pageId="54" pageNumber="305">Pronotum 1.7 times as broad as long; anterior margin indistinctly and widely concave.</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDC4CBAC403D6" blockId="54.[151,1437,646,2041]" box="[199,1219,1403,1430]" pageId="54" pageNumber="305">Elytra reaching middle portion of abdominal tergite III, 1.4–1.5 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDCA8BC9B03FA" blockId="54.[151,1437,646,2041]" box="[199,668,1439,1465]" pageId="54" pageNumber="305">Metatarsus 1.8 times as long as metatibia.</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDCF4BFA200D2" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
Male. Femora and tibia moderately wide. Posterior margin of abdominal tergite VIII straight (
<figureCitation id="B1CE2A1EFFDF6D6ED362DCF4BB88039E" box="[1320,1423,1475,1501]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">Fig. 161</figureCitation>
). Posterior margin of abdominal sternite VIII indistinctly sinuate (
<figureCitation id="B1CE2A1EFFDF6D6ED5C0DCD0BDF60042" box="[906,1009,1511,1537]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">Fig. 162</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward moderately narrow median lobe, in preapical portion narrowed toward small rounded apex; parameres very narrow, reaching apex of median lobe, with two long apical and two preapical setae; internal sac long and moderately wide, with two narrow elongate structures in basal portion (
<figureCitation id="B1CE2A1EFFDF6D6ED20FDF64BAA0002D" box="[1093,1191,1619,1646]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">Fig. 159</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFDF6D6ED70ADF40BFA700D2" box="[320,416,1655,1681]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">Fig. 160</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDFACBCAC0099" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
Female. Femora and tibia narrow. Posterior margin of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFDF6D6ED27CDFACBA9100F6" box="[1078,1174,1691,1717]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">Fig. 163</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFDF6D6ED325DFACBEC5009A" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">Fig. 164</figureCitation>
) truncate. Accessory sclerite as in
<figureCitation id="B1CE2A1EFFDF6D6ED40DDF88BCA00099" box="[583,679,1727,1754]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="54" pageNumber="305">Fig. 165</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDFD5BF660106" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DDFD5BFB500BE" bold="true" box="[199,434,1762,1789]" pageId="54" pageNumber="305">Comparative notes.</emphasis>
Based on the general shape and sculpture of the body
<taxonomicName id="EEF54D18FFDF6D6ED268DFD3BAA300BE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1058,1188,1763,1789]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="thayerae" status="sp. n.">
<emphasis id="1B81EA89FFDF6D6ED268DFD3BAA300BE" box="[1058,1188,1763,1789]" italics="true" pageId="54" pageNumber="305">X. thayerae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED2E7DFD4BAEA00BE" bold="true" box="[1197,1261,1763,1789]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED2E7DFD4BAEA00BE" box="[1197,1261,1763,1789]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFDF6D6ED3CEDFD3BF0A0162" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="guineensis" status="sp. n.">
<emphasis id="1B81EA89FFDF6D6ED3CEDFD3BF0A0162" italics="true" pageId="54" pageNumber="305">X. guineensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED75EDE30BF550162" bold="true" box="[276,338,1799,1825]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED75EDE30BF550162" box="[276,338,1799,1825]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which can be distinguished by the slightly shorter body, narrower median lobe and distinctly longer parameres.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDF6D6ED68DDE79BF0901CD" pageId="54" pageNumber="305" type="distribution">
<paragraph id="294A369BFFDF6D6ED68DDE79BF0901CD" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DDE79BF66012B" bold="true" box="[199,353,1870,1896]" pageId="54" pageNumber="305">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFDF6D6ED722DE78BC6C012A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[360,619,1871,1897]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="thayerae" status="sp. nov.">
<emphasis id="1B81EA89FFDF6D6ED722DE78BC6C012A" box="[360,619,1871,1897]" italics="true" pageId="54" pageNumber="305">Xanthonomus thayerae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDF6D6ED438DE78BCB5012A" bold="true" box="[626,690,1871,1897]" pageId="54" pageNumber="305">
<taxonomicNameLabel id="00B257F2FFDF6D6ED438DE78BCB5012A" box="[626,690,1871,1897]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in central part of
<collectingCountry id="51E2760BFFDF6D6ED28EDE78BB9B012A" box="[1220,1436,1871,1897]" name="Papua New Guinea" pageId="54" pageNumber="305">Papua New Guinea</collectingCountry>
(
<figureCitation id="B1CE2A1EFFDF6D6ED6D5DE44BF0501CD" box="[159,258,1907,1934]" captionStart="FIGURE 158" captionStartId="53.[151,250,1956,1980]" captionTargetBox="[151,1436,1165,1932]" captionTargetId="figure-421@53.[151,1436,1165,1932]" captionTargetPageId="53" captionText="FIGURE 158. Distribution of Xanthonomus: X. toxopeanus (circle), X. latus (triangle), X. guineensis (squares), X. thayerae (diamonds)." figureDoi="http://doi.org/10.5281/zenodo.10821299" httpUri="https://zenodo.org/record/10821299/files/figure.png" pageId="54" pageNumber="305">Fig. 158</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDF6D6ED68DDEA0BDC501BA" pageId="54" pageNumber="305" type="etymology">
<paragraph id="294A369BFFDF6D6ED68DDEA0BF130196" blockId="54.[151,1437,646,2041]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DDEA0BF4901F2" bold="true" box="[199,334,1943,1969]" pageId="54" pageNumber="305">Etymology.</emphasis>
Patronymic, the species is named after my colleague Margaret K. Thayer (Chicago), specialist in
<taxonomicName id="EEF54D18FFDF6D6ED6DDDE8CBF080196" box="[151,271,1979,2005]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
.
</paragraph>
<paragraph id="294A369BFFDF6D6ED68DDEE9BDC501BA" blockId="54.[151,1437,646,2041]" box="[199,962,2014,2041]" pageId="54" pageNumber="305">
<emphasis id="1B81EA89FFDF6D6ED68DDEE9BF4C01BB" bold="true" box="[199,331,2014,2040]" pageId="54" pageNumber="305">Bionomics.</emphasis>
The
<typeStatus id="F64E8839FFDF6D6ED7CEDEE8BFE301BA" box="[388,484,2015,2041]" pageId="54" pageNumber="305" type="paratype">paratype</typeStatus>
was collected in leaf mould in pine forest.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5616B831D84C90DDBCAA74C8FDC06E34">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<subSubSection id="61EF6510FFE16D50D6DDDD09BC4C021A" box="[151,587,1086,1113]" pageId="8" pageNumber="259" type="nomenclature">
<paragraph id="294A369BFFE16D50D6DDDD09BC4C021A" blockId="8.[151,587,1086,1149]" box="[151,587,1086,1113]" pageId="8" pageNumber="259">
<heading id="720281F7FFE16D50D6DDDD09BC4C021A" bold="true" box="[151,587,1086,1113]" fontSize="11" level="1" pageId="8" pageNumber="259" reason="1">
<emphasis id="1B81EA89FFE16D50D6DDDD09BC4C021A" bold="true" box="[151,587,1086,1113]" pageId="8" pageNumber="259">
<taxonomicName id="EEF54D18FFE16D50D6DDDD09BC05021A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,514,1086,1113]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. nov.">
<emphasis id="1B81EA89FFE16D50D6DDDD09BC05021A" bold="true" box="[151,514,1086,1113]" italics="true" pageId="8" pageNumber="259">Paraphloeostiba atramentaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFE16D50D443DD08BC4C021A" box="[521,587,1087,1113]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE16D50D6DDDD54BF44023E" box="[151,323,1123,1149]" pageId="8" pageNumber="259" type="description">
<paragraph id="294A369BFFE16D50D6DDDD54BF44023E" blockId="8.[151,587,1086,1149]" box="[151,323,1123,1149]" pageId="8" pageNumber="259">
(
<figureCitation id="B1CE2A1EFFE16D50D6D5DD54BEE0023E" box="[159,231,1123,1149]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="8" pageNumber="259">Figs 3</figureCitation>
,
<figureCitation id="B1CE2A1EFFE16D50D6BEDD54BF3D023E" box="[244,314,1123,1149]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="8" pageNumber="259">16–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE16D50D6DDDD9CBF58034E" pageId="8" pageNumber="259" type="materials_examined">
<paragraph id="294A369BFFE16D50D6DDDD9CBF58034E" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">
<emphasis id="1B81EA89FFE16D50D6DDDD9CBF430287" bold="true" box="[151,324,1194,1221]" pageId="8" pageNumber="259">Type material.</emphasis>
<typeStatus id="F64E8839FFE16D50D700DD9CBFB50286" box="[330,434,1195,1221]" pageId="8" pageNumber="259" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFE16D50D7F3DD9CBFC90286" box="[441,462,1195,1221]" count="1" pageId="8" pageNumber="259" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFE16D50D797DD9CBC270286" box="[477,544,1195,1221]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="8" pageNumber="259">Fig. 3</figureCitation>
; dissected): ‘
<collectingRegion id="EB31F879FFE16D50D4E5DD9CBD480286" box="[687,847,1195,1221]" country="Indonesia" name="Papua Barat" pageId="8" pageNumber="259">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Sinak,
<quantity id="EE0D9B7EFFE16D50D274DD9CBACA0286" box="[1086,1229,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="8" pageNumber="259" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000-2200m</quantity>
|
<date id="5D4B105BFFE16D50D295DD9CBB9B0286" box="[1247,1436,1194,1221]" pageId="8" pageNumber="259" value="1995-12-14" valueMax="1995-12-17" valueMin="1995-12-14">14. -17. 12. 1995</date>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFE16D50D79FDDF8BC6902AA" box="[469,622,1231,1257]" pageId="8" pageNumber="259" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE16D50D4CFDDF8BD4702AA" authorityName="Steel" authorityYear="1960" box="[645,832,1231,1257]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE16D50D4CFDDF8BD4702AA" box="[645,832,1231,1257]" italics="true" pageId="8" pageNumber="259">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE16D50D51DDDF8BDEC02AB" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[855,1003,1231,1256]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. nov.">
<emphasis id="1B81EA89FFE16D50D51DDDF8BDEC02AB" box="[855,1003,1231,1256]" italics="true" pageId="8" pageNumber="259">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D5B9DDF8BA3202AA" bold="true" box="[1011,1077,1231,1257]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D5B9DDF8BA3202AA" box="[1011,1077,1231,1257]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE16D50D289DDF8BADE02AA" box="[1219,1241,1231,1257]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="259" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFE16D50D74FDDC4BF54034E" box="[261,339,1267,1293]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="259">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE16D52D68DDC21BC6E0556" lastPageId="10" lastPageNumber="261" pageId="8" pageNumber="259" type="description">
<paragraph id="294A369BFFE16D50D68DDC21BB9A0316" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">
<emphasis id="1B81EA89FFE16D50D68DDC21BF5E0373" bold="true" box="[199,345,1302,1328]" pageId="8" pageNumber="259">Description.</emphasis>
Measurements: HW: 0.47; HL: 0.32; OL: 0.17; AL: 0.63; PL: 0.42; PWmax: 0.62; PWmin: 0.51; ESL: 0.68; EW: 0.80; MTbL: 0.40; MTrL: 0.22 (MTrL 1–4: 0.07; MTrL 5: 0.15); AW: 0.82; AedL: 0.51; BL: 2.75.
</paragraph>
<paragraph id="294A369BFFE16D50D68DDC68BFE300FE" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">
Habitus as in
<figureCitation id="B1CE2A1EFFE16D50D712DC68BF9F033A" box="[344,408,1375,1401]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="8" pageNumber="259">Fig.3</figureCitation>
. Body and antennomeres 6–11 dark brown; mouthparts, antennomeres 1–5 and legs yellowish. Head with sparse and fine punctation, more indistinct in middle and slightly denser in infraorbital portions; neck without visible punctation; pronotum with fine and dense punctation, slightly larger than that in head, sparser in middle; scutellum without punctures; elytra with dense and fine punctation, slightly larger than that in pronotum, finer and sparser in middle, with narrow impunctate area along suture in basal half; abdominal tergites with very indistinct, fine and sparse punctation. Head with very dense and coarse microsculpture, transverse in frontal portion and distinctly isodiametric in middle; pronotum with very dense, coarse, isodiametric microsculpture; scutellum with very fine transverse meshes; elytra with dense isodiametric microreticulation, slightly larger than that in pronotum; abdominal tergites with dense microsculpture, transverse in abdominal tergites III–IV and isodiametric in abdominal tergites V–VIII.
</paragraph>
<paragraph id="294A369BFFE16D50D68DDFF0BD5A019E" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">Head slightly elevated in middle, with elongate anteriomedian depressions, 1.4 times as broad as long, with short and deep anteocellar foveae, slightly subdiagonally stretching anteriad and reaching basal third of length of eyes; postocular carina indistinct, acute. Ocelli small, located slightly below level of posterior margins of eyes; distance between ocelli about 1.6 times as long as distince between ocellus and posterior margin of eyes. Apical antennomere about 1.5 times as long as preapical segment. Antennomere 3 about as long as and distinctly narrower than 2, 4 about as wide as and about twice shorter than 3, 5 distinctly longer and broader than 4, 6–9 slightly longer and distinctly broader than 5, 10 indistinctly longer than 9, apical antennomere about 1.5 times as long as 10, from about middle gradually narrowed apicad toward subacute apex.</paragraph>
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<paragraph id="294A369BFFE06D51D6DDDF8FBD9900B0" blockId="9.[151,1436,1720,1780]" pageId="9" pageNumber="260">
<emphasis id="1B81EA89FFE06D51D6DDDF8FBF540093" bold="true" box="[151,339,1720,1744]" pageId="9" pageNumber="260">FIGURES 16–19.</emphasis>
<taxonomicName id="EEF54D18FFE06D51D71DDF8FBC8C0093" authorityName="Shavrin" authorityYear="2024" box="[343,651,1720,1744]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="atramentaria">
<emphasis id="1B81EA89FFE06D51D71DDF8FBC8C0093" box="[343,651,1720,1744]" italics="true" pageId="9" pageNumber="260">Paraphloeostiba atramentaria</emphasis>
</taxonomicName>
(holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFE06D51D68DDE70BD7D01C7" blockId="9.[151,1437,1862,2033]" pageId="9" pageNumber="260">Pronotum 1.4 times as broad as long, slightly wider than head, widest in middle, distinctly more narrowed posteriad than anteriad; posteriolateral impressions wide, shallow.</paragraph>
<paragraph id="294A369BFFE06D51D68DDEB9BC6D01EA" blockId="9.[151,1437,1862,2033]" box="[199,618,1934,1961]" pageId="9" pageNumber="260">Elytra 1.6 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFE06D51D68DDE85BD23018F" blockId="9.[151,1437,1862,2033]" box="[199,804,1970,1997]" pageId="9" pageNumber="260">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFE06D52D68DDEE1BEEA0702" blockId="9.[151,1437,1862,2033]" lastBlockId="10.[151,1437,151,1005]" lastPageId="10" lastPageNumber="261" pageId="9" pageNumber="260">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFE06D51D51EDEE1BDA101B3" box="[852,934,2006,2032]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="9" pageNumber="260">Fig. 18</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFE36D52D793D9A0BC2A06F2" box="[473,557,151,177]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="10" pageNumber="261">Fig. 19</figureCitation>
). Aedeagus with wide basal portion, strongly narrowed toward narrow middle portion of elongate median lobe with small rounded apex; parameres moderately narrow, reaching apex of median lobe, each with three moderately short apical and six setae on inner margin; internal sac narrow and moderately short, with several long curved sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFE36D52D5E3D834BDFB075E" box="[937,1020,259,285]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="10" pageNumber="261">Fig. 16</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFE36D52D6DDD810BEED0702" box="[151,234,295,321]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="10" pageNumber="261">Fig. 17</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFE36D52D68DD87CBF960726" blockId="10.[151,1437,151,1005]" box="[199,401,331,357]" pageId="10" pageNumber="261">Female unknown.</paragraph>
<paragraph id="294A369BFFE36D52D68DD859BCBC0792" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
<emphasis id="1B81EA89FFE36D52D68DD859BFA807CA" bold="true" box="[199,431,366,393]" pageId="10" pageNumber="261">Comparative notes.</emphasis>
Based on the very narrow median lobe and the parameres,
<taxonomicName id="EEF54D18FFE36D52D271D858BAEA07CB" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1083,1261,367,393]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D271D858BA4B07CB" box="[1083,1100,367,392]" italics="true" pageId="10" pageNumber="261">P</emphasis>
.
<emphasis id="1B81EA89FFE36D52D213D858BAEA07CB" box="[1113,1261,367,392]" italics="true" pageId="10" pageNumber="261">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D2B8D858BB3607CA" bold="true" box="[1266,1329,367,393]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D2B8D858BB3607CA" box="[1266,1329,367,393]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFE36D52D6F9D8A4BF2B07EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[179,300,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D6F9D8A4BEC307EF" box="[179,196,403,428]" italics="true" pageId="10" pageNumber="261">P</emphasis>
.
<emphasis id="1B81EA89FFE36D52D69AD8A4BF2B07EE" box="[208,300,403,429]" italics="true" pageId="10" pageNumber="261">barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D778D8A4BF7707EE" bold="true" box="[306,368,403,429]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D778D8A4BF7707EE" box="[306,368,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFE36D52D737D8A4BC0007EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[381,519,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D737D8A4BC0007EE" box="[381,519,403,429]" italics="true" pageId="10" pageNumber="261">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D446D8A4BC4D07EE" bold="true" box="[524,586,403,429]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D446D8A4BC4D07EE" box="[524,586,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFE36D52D41DD8A4BCEC07EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[599,747,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D41DD8A4BCEC07EE" box="[599,747,403,429]" italics="true" pageId="10" pageNumber="261">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D4BAD8A4BD2907EE" bold="true" box="[752,814,403,429]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D4BAD8A4BD2907EE" box="[752,814,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="EEF54D18FFE36D52D523D8A4BDB907EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[873,958,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D523D8A4BDB907EE" box="[873,958,403,429]" italics="true" pageId="10" pageNumber="261">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D589D8A4BA0507EE" bold="true" box="[963,1026,403,429]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D589D8A4BA0507EE" box="[963,1026,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFE36D52D68DD8ECBB1207B6" blockId="10.[151,1437,151,1005]" box="[199,1301,475,501]" pageId="10" pageNumber="261">
from
<taxonomicName id="EEF54D18FFE36D52D74FD8ECBF7B07B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D74FD8ECBF7B07B6" box="[261,380,475,501]" italics="true" pageId="10" pageNumber="261">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D7C9D8ECBFC407B6" bold="true" box="[387,451,475,501]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D7C9D8ECBFC407B6" box="[387,451,475,501]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker elytra, slightly narrower pronotum and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFE36D52D68DD8C8BD9A047E" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
from
<taxonomicName id="EEF54D18FFE36D52D74ED8C8BF97045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,400,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D74ED8C8BF97045A" box="[260,400,511,537]" italics="true" pageId="10" pageNumber="261">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D7DDD8C8BFD1045A" bold="true" box="[407,470,511,537]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D7DDD8C8BFD1045A" box="[407,470,511,537]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the presence of the punctation on the head, coarser punctation of the pronotum, the narrower pronotum, the shorter elytra and slightly shorter parameres;
</paragraph>
<paragraph id="294A369BFFE36D52D68DDB70BD2E04C6" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
from
<taxonomicName id="EEF54D18FFE36D52D74CDB70BF9A0422" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[262,413,583,609]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D74CDB70BF9A0422" box="[262,413,583,609]" italics="true" pageId="10" pageNumber="261">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D7ECDB70BFE00422" bold="true" box="[422,487,583,609]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D7ECDB70BFE00422" box="[422,487,583,609]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the presence of the punctation on the head, coarser punctation of the pronotum, shorter elytra, and slightly narrower and longer parameres;
</paragraph>
<paragraph id="294A369BFFE36D52D68DDBB8BD4C048E" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
from
<taxonomicName id="EEF54D18FFE36D52D74DDBB8BF5804EA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[263,351,655,681]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D74DDBB8BF5804EA" box="[263,351,655,681]" italics="true" pageId="10" pageNumber="261">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D722DBB8BFAD04EA" bold="true" box="[360,426,655,681]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D722DBB8BFAD04EA" box="[360,426,655,681]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker coloration, the absence of elevations between punctures on the elytra, the shorter parameres with significantly narrower apical portions.
</paragraph>
<paragraph id="294A369BFFE36D52D68DDBE0BC6E0556" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
From all these species,
<taxonomicName id="EEF54D18FFE36D52D780DBE0BC7B04B3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[458,636,727,753]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE36D52D780DBE0BFDC04B3" box="[458,475,727,752]" italics="true" pageId="10" pageNumber="261">P</emphasis>
.
<emphasis id="1B81EA89FFE36D52D7A2DBE0BC7B04B3" box="[488,636,727,752]" italics="true" pageId="10" pageNumber="261">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D4C8DBE0BCC704B2" bold="true" box="[642,704,727,753]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D4C8DBE0BCC704B2" box="[642,704,727,753]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the larger body and details of the external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE36D52D68DDA29BF6C051E" pageId="10" pageNumber="261" type="distribution">
<paragraph id="294A369BFFE36D52D68DDA29BF6C051E" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
<emphasis id="1B81EA89FFE36D52D68DDA29BF66057B" bold="true" box="[199,353,798,824]" pageId="10" pageNumber="261">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFE36D52D726DA28BCC1057B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[364,710,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. nov.">
<emphasis id="1B81EA89FFE36D52D726DA28BCC1057B" box="[364,710,799,825]" italics="true" pageId="10" pageNumber="261">Paraphloeostiba atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D498DA28BD11057A" bold="true" box="[722,790,799,825]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D498DA28BD11057A" box="[722,790,799,825]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFE36D52D20EDA28BA73057A" box="[1092,1140,799,825]" pageId="10" pageNumber="261">type</typeStatus>
locality in Western New
<collectingCountry id="51E2760BFFE36D52D6DDDA74BEEC051E" box="[151,235,835,861]" name="Guinea-Bissau" pageId="10" pageNumber="261">Guinea</collectingCountry>
,
<collectingCountry id="51E2760BFFE36D52D6BCDA74BF60051E" box="[246,359,835,861]" name="Indonesia" pageId="10" pageNumber="261">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE36D52D68DDA50BF3705AE" pageId="10" pageNumber="261" type="etymology">
<paragraph id="294A369BFFE36D52D68DDA50BEF905E6" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
<emphasis id="1B81EA89FFE36D52D68DDA50BF4905C2" bold="true" box="[199,334,871,897]" pageId="10" pageNumber="261">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFE36D52D482DA50BD6005C3" box="[712,871,871,896]" italics="true" pageId="10" pageNumber="261">atramentarius</emphasis>
, -
<emphasis id="1B81EA89FFE36D52D535DA50BDCF05C3" box="[895,968,871,896]" italics="true" pageId="10" pageNumber="261">a, -um</emphasis>
(blackish). It alludes to the coloration of the body.
</paragraph>
<paragraph id="294A369BFFE36D52D68DDA99BF3705AE" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
<emphasis id="1B81EA89FFE36D52D68DDA99BF4C058B" bold="true" box="[199,331,942,968]" pageId="10" pageNumber="261">Bionomics.</emphasis>
The
<typeStatus id="F64E8839FFE36D52D7CCDA98BFEF058A" box="[390,488,943,969]" pageId="10" pageNumber="261" type="holotype">holotype</typeStatus>
was collected at elevations from
<quantity id="EE0D9B7EFFE36D52D52DDA98BA1C058A" box="[871,1051,943,969]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="10" pageNumber="261" unit="m" value="2200.0" valueMax="2400.0" valueMin="2000.0">2000 to 2400 m</quantity>
a.s.l. The detailed bionomial data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="A9A5C7A5E338850E61013C9F493F4D74">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<mods:title id="386F8128653E01107CE98FFCE07FBC44">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="75964C7FE53634035BFB9191A659306F">
<mods:date id="26E2DA567AE62CBB67BD303CAB284409">2024</mods:date>
<mods:detail id="B3388994B9EE30702840CBA5D8130475" type="pubDate">
<mods:number id="E7AEAF1F39F62543284A341C98233952">2024-03-14</mods:number>
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<mods:detail id="AAE092B21C505D19519AC97EDC6FF329" type="volume">
<mods:number id="700F2BEE34F3D7F87FA2E9C05C2F00BF">5424</mods:number>
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<mods:number id="75CBE135075546A0A2C80A17A6C4DF71">3</mods:number>
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<mods:start id="571D7F906A50DF02915D368C81488AC0">251</mods:start>
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<mods:location id="071818134BBA13CDB5118A2E2EAD3A74">
<mods:url id="ABCAC4C72A564733E4FF5C8360324799">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
</mods:location>
<mods:classification id="279A55D1B82133CE82DB034B6C750A9C">journal article</mods:classification>
<mods:identifier id="6633EB45242E1ACE077FE3BC640F0392" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="B9B96596A8FDBC7716F3CA7F3611B9F2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="760D0A1EC4FF060217289FE5BA7BBB4C" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="0DBD80D90808A4C304929C396BB111DE" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFE36D54D6DDDD04BA0D051E" ID-DOI="http://doi.org/10.5281/zenodo.10821257" ID-Zenodo-Dep="10821257" LSID="urn:lsid:plazi:treatment:A15C878DFFE36D54D6DDDD04BA0D051E" httpUri="http://treatment.plazi.org/id/A15C878DFFE36D54D6DDDD04BA0D051E" lastPageId="12" lastPageNumber="263" pageId="10" pageNumber="261">
<subSubSection id="61EF6510FFE36D52D6DDDD04BC18020D" box="[151,543,1075,1102]" pageId="10" pageNumber="261" type="nomenclature">
<paragraph id="294A369BFFE36D52D6DDDD04BC18020D" blockId="10.[151,543,1075,1139]" box="[151,543,1075,1102]" pageId="10" pageNumber="261">
<heading id="720281F7FFE36D52D6DDDD04BC18020D" bold="true" box="[151,543,1075,1102]" fontSize="11" level="1" pageId="10" pageNumber="261" reason="1">
<emphasis id="1B81EA89FFE36D52D6DDDD04BC18020D" bold="true" box="[151,543,1075,1102]" pageId="10" pageNumber="261">
<taxonomicName id="EEF54D18FFE36D52D6DDDD04BFD1020D" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,470,1075,1102]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="attenuata" status="sp. nov.">
<emphasis id="1B81EA89FFE36D52D6DDDD04BFD1020D" bold="true" box="[151,470,1075,1102]" italics="true" pageId="10" pageNumber="261">Paraphloeostiba attenuata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFE36D52D797DD03BC18020D" box="[477,543,1076,1102]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE36D52D6DDDD6FBF440230" box="[151,323,1112,1139]" pageId="10" pageNumber="261" type="description">
<paragraph id="294A369BFFE36D52D6DDDD6FBF440230" blockId="10.[151,543,1075,1139]" box="[151,323,1112,1139]" pageId="10" pageNumber="261">
(
<figureCitation id="B1CE2A1EFFE36D52D6D5DD6FBEEF0231" box="[159,232,1112,1138]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="10" pageNumber="261">Figs 4</figureCitation>
,
<figureCitation id="B1CE2A1EFFE36D52D6BEDD6FBF3B0230" box="[244,316,1112,1139]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="10" pageNumber="261">20–25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE36D52D6DDDD97BFD300CD" pageId="10" pageNumber="261" type="materials_examined">
<paragraph id="294A369BFFE36D52D6DDDD97BD170341" blockId="10.[151,1437,1184,2039]" pageId="10" pageNumber="261">
<emphasis id="1B81EA89FFE36D52D6DDDD97BF4302F9" bold="true" box="[151,324,1184,1210]" pageId="10" pageNumber="261">Type material.</emphasis>
<typeStatus id="F64E8839FFE36D52D700DD97BFB502F9" box="[330,434,1184,1210]" pageId="10" pageNumber="261" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFE36D52D7F3DD96BFC902F8" box="[441,462,1185,1211]" count="1" pageId="10" pageNumber="261" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFE36D52D796DD97BC2602F9" box="[476,545,1184,1210]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="10" pageNumber="261">Fig. 4</figureCitation>
; underside of the card with handwritten number ‘65’): ‘
<collectionCode id="4FE4AE5EFFE36D52D2CCDD97BACD02F9" box="[1158,1226,1184,1210]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="10" pageNumber="261">NEW</collectionCode>
<collectingCountry id="51E2760BFFE36D52D29ADD97BB3D02F9" box="[1232,1338,1184,1210]" name="Guinea-Bissau" pageId="10" pageNumber="261">GUINEA</collectingCountry>
| Okapa, |
<date id="5D4B105BFFE36D52D6EFDDF3BF20029C" box="[165,295,1220,1247]" pageId="10" pageNumber="261" value="1965-06-27">27.VI.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE36D52D7F2DDF3BFD6029D" box="[440,465,1220,1246]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="261">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFE36D52D5F9DDF3BA4B029D" box="[947,1100,1220,1246]" pageId="10" pageNumber="261" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE36D52D228DDF3BB1A029D" authorityName="Steel" authorityYear="1960" box="[1122,1309,1220,1246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE36D52D228DDF3BB1A029D" box="[1122,1309,1220,1246]" italics="true" pageId="10" pageNumber="261">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE36D52D378DDF2BB9B029D" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1330,1436,1221,1246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="attenuata" status="sp. nov.">
<emphasis id="1B81EA89FFE36D52D378DDF2BB9B029D" box="[1330,1436,1221,1246]" italics="true" pageId="10" pageNumber="261">attenuata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D6DDDDDFBED00341" bold="true" box="[151,215,1256,1282]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D6DDDDDFBED00341" box="[151,215,1256,1282]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE36D52D72ADDDFBF710341" box="[352,374,1256,1282]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="261" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFE36D52D4E3DDDFBD040341" box="[681,771,1256,1282]" pageId="10" pageNumber="261">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFE36D52D68DDC3ABFD300CD" blockId="10.[151,1437,1184,2039]" pageId="10" pageNumber="261">
<typeStatus id="F64E8839FFE36D52D68DDC3ABF3F0364" box="[199,312,1293,1319]" pageId="10" pageNumber="261" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFE36D52D70CDC3BBF740364" box="[326,371,1292,1319]" count="1" pageId="10" pageNumber="261" type="female">1 ♀</specimenCount>
(dissected): same data as the holotype (
<collectionCode id="4FE4AE5EFFE36D52D50EDC3BBD990365" box="[836,926,1292,1318]" pageId="10" pageNumber="261">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFE36D52D5FFDC3BBDE50364" box="[949,994,1292,1319]" count="1" pageId="10" pageNumber="261" type="male">1 ♂</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFE36D52D231DC3BBAB80365" box="[1147,1215,1292,1318]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="10" pageNumber="261">NEW</collectionCode>
<collectingCountry id="51E2760BFFE36D52D283DC3BBB340365" box="[1225,1331,1292,1318]" name="Guinea-Bissau" pageId="10" pageNumber="261">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFE36D52D6DDDC07BF0B0308" box="[151,268,1328,1355]" pageId="10" pageNumber="261" value="1965-05-23">
23.
<collectionCode id="4FE4AE5EFFE36D52D6F2DC07BEC90308" box="[184,206,1328,1355]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="261" type="Museum">V</collectionCode>
.1965
</date>
, | cow dung and be- | ating’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE36D52D4BBDC07BD0D0309" box="[753,778,1328,1354]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="261">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFE36D52D2B6DC07BB510309" box="[1276,1366,1328,1354]" pageId="10" pageNumber="261">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFE36D52D324DC07BB9B0308" box="[1390,1436,1328,1355]" count="1" pageId="10" pageNumber="261" type="female">1 ♀</specimenCount>
(underside of the card with handwritten number ‘64’): ‘
<collectionCode id="4FE4AE5EFFE36D52D546DC63BD57032D" box="[780,848,1364,1390]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="10" pageNumber="261">NEW</collectionCode>
<collectingCountry id="51E2760BFFE36D52D513DC63BDC4032D" box="[857,963,1364,1390]" name="Guinea-Bissau" pageId="10" pageNumber="261">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFE36D52D278DC63BAB3032C" box="[1074,1204,1364,1391]" pageId="10" pageNumber="261" value="1965-06-16">
16.
<collectionCode id="4FE4AE5EFFE36D52D218DC63BA74032C" box="[1106,1139,1364,1391]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="10" pageNumber="261">VI</collectionCode>
.1965
</date>
, | flying in daylight’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE36D52D753DC4FBF3503D1" box="[281,306,1400,1426]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="261">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFE36D52D542DC4FBD6503D1" box="[776,866,1400,1426]" pageId="10" pageNumber="261">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFE36D52D53FDC4FBD9903D0" box="[885,926,1400,1427]" count="1" pageId="10" pageNumber="261" type="male">1 ♂</specimenCount>
(right antennomeres 3–11 missing; underside of the card with handwritten number ‘85’),
<specimenCount id="3FF3FD12FFE36D52D4C3DCABBCCE03F4" box="[649,713,1436,1463]" count="2" pageId="10" pageNumber="261" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFE36D52D490DCABBD7103F5" box="[730,886,1436,1462]" count="1" pageId="10" pageNumber="261" type="generic">one specimen</specimenCount>
without left antenna; underside of the card with handwritten number ‘85’): ‘
<collectionCode id="4FE4AE5EFFE36D52D781DCF7BC080399" box="[459,527,1472,1498]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="10" pageNumber="261">NEW</collectionCode>
<collectingCountry id="51E2760BFFE36D52D45DDCF7BC860399" box="[535,641,1472,1498]" name="Guinea-Bissau" pageId="10" pageNumber="261">GUINEA</collectingCountry>
| Okapa, |
<date id="5D4B105BFFE36D52D4BDDCF7BD8B0398" box="[759,908,1472,1499]" pageId="10" pageNumber="261" value="1965-08-27">27.VIII.1965</date>
, | forest litter’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE36D52D2FBDCF7BACD0399" box="[1201,1226,1472,1498]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="261">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFE36D52D7DFDCD3BFE803BD" box="[405,495,1508,1534]" pageId="10" pageNumber="261">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFE36D52D449DCD3BC2B03BC" box="[515,556,1508,1535]" count="1" pageId="10" pageNumber="261" type="female">1 ♀</specimenCount>
(underside of the card with handwritten number ‘170’): ‘
<collectionCode id="4FE4AE5EFFE36D52D2E3DCD3BAEA03BD" box="[1193,1261,1508,1534]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="10" pageNumber="261">NEW</collectionCode>
<collectingCountry id="51E2760BFFE36D52D2BEDCD3BB5903BD" box="[1268,1374,1508,1534]" name="Guinea-Bissau" pageId="10" pageNumber="261">GUINEA</collectingCountry>
Go- | roka, |
<date id="5D4B105BFFE36D52D6A2DF3FBF6E0061" box="[232,361,1544,1570]" pageId="10" pageNumber="261" value="1971-03-28">28.III.1971</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE36D52D7B6DF3FBC120061" box="[508,533,1544,1570]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="261">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFE36D52D5B0DF3FBA530061" box="[1018,1108,1544,1570]" pageId="10" pageNumber="261">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFE36D52D220DF3FBA900060" box="[1130,1175,1544,1571]" count="1" pageId="10" pageNumber="261" type="male">1 ♂</specimenCount>
(underside of the card with handwritten number ‘86’): ‘
<collectionCode id="4FE4AE5EFFE36D52D449DF1BBC400005" box="[515,583,1580,1606]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="10" pageNumber="261">NEW</collectionCode>
<collectingCountry id="51E2760BFFE36D52D404DF1BBCBF0005" box="[590,696,1580,1606]" name="Guinea-Bissau" pageId="10" pageNumber="261">GUINEA</collectingCountry>
| Lae, |
<date id="5D4B105BFFE36D52D545DF1BBDA30004" box="[783,932,1580,1607]" pageId="10" pageNumber="261" value="1965-08-25">25.VIII.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE36D52D278DF1BBA4C0005" box="[1074,1099,1580,1606]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="261">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFE36D52D75FDF67BF680029" box="[277,367,1616,1642]" pageId="10" pageNumber="261">BMNH</collectionCode>
). All
<typeStatus id="F64E8839FFE36D52D7E1DF66BC110028" box="[427,534,1617,1643]" pageId="10" pageNumber="261" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFE36D52D5DDDF67BA200029" box="[919,1063,1616,1642]" pageId="10" pageNumber="261" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE36D52D276DF67BAF00029" authorityName="Steel" authorityYear="1960" box="[1084,1271,1616,1642]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE36D52D276DF67BAF00029" box="[1084,1271,1616,1642]" italics="true" pageId="10" pageNumber="261">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE36D52D340DF66BB730029" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1290,1396,1617,1642]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="attenuata" status="sp. nov.">
<emphasis id="1B81EA89FFE36D52D340DF66BB730029" box="[1290,1396,1617,1642]" italics="true" pageId="10" pageNumber="261">attenuata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE36D52D331DF67BEA900CD" bold="true" pageId="10" pageNumber="261">
<taxonomicNameLabel id="00B257F2FFE36D52D331DF67BEA900CD" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE36D52D77DDF43BF4A00CD" box="[311,333,1652,1678]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="261" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE36D54D68DDFAFBFBB04EA" lastPageId="12" lastPageNumber="263" pageId="10" pageNumber="261" type="description">
<paragraph id="294A369BFFE36D52D68DDFAFBB7000B9" blockId="10.[151,1437,1184,2039]" pageId="10" pageNumber="261">
<emphasis id="1B81EA89FFE36D52D68DDFAFBF5E00F1" bold="true" box="[199,345,1688,1714]" pageId="10" pageNumber="261">Description.</emphasis>
Measurements (n=9): HW: 0.38–0.42; HL: 0.22–0.35; OL: 0.11–0.15; AL (
<typeStatus id="F64E8839FFE36D52D2FCDFAFBB1800F1" box="[1206,1311,1688,1714]" pageId="10" pageNumber="261" type="holotype">holotype</typeStatus>
): 0.51; PL: 0.27–0.36; PWmax: 0.48–0.62; PWmin: 0.43–0.57; ESL: 0.37–0.47; EW: 0.72–0.76; MTbL (
<typeStatus id="F64E8839FFE36D52D2EBDF8BBB0D0095" box="[1185,1290,1724,1750]" pageId="10" pageNumber="261" type="holotype">holotype</typeStatus>
): 0.27; MTrL (
<typeStatus id="F64E8839FFE36D52D6D5DFD7BF0F00B9" box="[159,264,1760,1786]" pageId="10" pageNumber="261" type="holotype">holotype</typeStatus>
): 0.18 (MTrL 1–4: 0.04; MTrL 5: 0.14); AW: 0.66–0.75; AedL: 0.30; BL: 1.58–2.00 (
<typeStatus id="F64E8839FFE36D52D289DFD7BB2C00B9" box="[1219,1323,1760,1786]" pageId="10" pageNumber="261" type="holotype">holotype</typeStatus>
: 1.92).
</paragraph>
<paragraph id="294A369BFFE36D53D68DDE33BC9D06BA" blockId="10.[151,1437,1184,2039]" lastBlockId="11.[151,1437,151,681]" lastPageId="11" lastPageNumber="262" pageId="10" pageNumber="261">
Habitus as in
<figureCitation id="B1CE2A1EFFE36D52D72EDE33BFAD015D" box="[356,426,1796,1822]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="10" pageNumber="261">Fig. 4</figureCitation>
. Body glossy, yellowish-brown to reddish brown, with paler elytra and slightly darkened lateroapical and posterior portions of elytra; antennomeres 6–11 or 7–11 brown; mouthparts, antennomeres 1–5 or 1–6 and legs yellow. Head without or with indistinct, very fine and sparse punctation, slightly denser and larger in middle in
<specimenCount id="3FF3FD12FFE36D52D742DE46BFAF01C8" box="[264,424,1905,1931]" count="2" pageId="10" pageNumber="261" type="generic" typeStatus="paratype">two paratypes</specimenCount>
; neck without visible punctures; pronotum with fine and sparse punctation, finer and more indistinct in middle (
<specimenCount id="3FF3FD12FFE36D52D7C9DEA3BC1101EC" box="[387,534,1940,1967]" count="1" pageId="10" pageNumber="261" type="generic" typeStatus="paratype">one paratype</specimenCount>
without visible punctation in middle); scutellum without punctures; elytra with moderately dense punctation, distinctly larger than that in pronotum, sparser and finer around scutellum and along suture; abdominal tergites with indistinct or distinct sparse and fine punctation, finer and sparser in middle. Head with dense microsculpture, usually coarser in middle; pronotum with dense isodiametric microsculpture similar to that in middle portion of head; microreticulation of elytra dense, similar to that in pronotum; abdominal tergites with fine and dense isodimaetric microreticulation.
</paragraph>
<paragraph id="294A369BFFE26D53D68DD834BB9B045A" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="262">
Head with distinct, shallow or moderately deep and wide anteriomedian depressions, 1.2–1.7 times as broad as long, with very short and deep anteocellar foveae about as long as broad of each ocellus; middle portion between ocelli in
<specimenCount id="3FF3FD12FFE26D53D6BCD87CBF830726" box="[246,388,331,357]" count="1" pageId="11" pageNumber="262" type="generic" typeStatus="paratype">one paratype</specimenCount>
semicircularly depressed; postocular carina distinct, acute. Ocelli small, located slightly above or about at level of posterior margins of eyes; distance between ocelli distinctly larger than distance between ocellus and posterior margin of eyes. Apical maxillary palpomere 1.5–1.7 times as long as preapical segment. Antennomere 3 about as long as or slightly shorter and distinctly narrower than 2, 4 about twice shorter than 3, 5 slightly shorter than 4, 6 slightly longer and distinctly broader than 5, 7–8 distinctly broader than 6, 9–10 slightly longer than 8, apical antennomere slightly more than twice as long as 10, from middle gradually narrowed toward subacute apex.
</paragraph>
<paragraph id="294A369BFFE26D53D68DDB14BAD404EA" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="262">Pronotum 1.7 times as broad as long, 1.2–1.4 times as broad as head, widest in or slightly behind middle, significantly more narrowed posteriad than anteriad; laterobasal portions widely concave, sometimes with subparallel laterobasal margins in front of obtuse hind angles; middle portion without or with very indistinct wide and long median impressons; lateral impressions wide and deep, distinctly broader in laterobasal portions.</paragraph>
<caption id="7D8A6613FFE26D53D6DDDEBFBF5001AB" ID-DOI="http://doi.org/10.5281/zenodo.10821227" ID-Zenodo-Dep="10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="11" pageNumber="262" startId="11.[151,264,1928,1952]" targetBox="[159,1428,711,1895]" targetPageId="11" targetType="figure">
<paragraph id="294A369BFFE26D53D6DDDEBFBF5001AB" blockId="11.[151,1437,1928,2024]" pageId="11" pageNumber="262">
<emphasis id="1B81EA89FFE26D53D6DDDEBFBF5201E3" bold="true" box="[151,341,1928,1952]" pageId="11" pageNumber="262">FIGURES 20–25.</emphasis>
<taxonomicName id="EEF54D18FFE26D53D717DEBFBC6A01E3" authorityName="Shavrin" authorityYear="2024" box="[349,621,1928,1952]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="262" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="1B81EA89FFE26D53D717DEBFBC6A01E3" box="[349,621,1928,1952]" italics="true" pageId="11" pageNumber="262">Paraphloeostiba attenuata</emphasis>
</taxonomicName>
(paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFE56D54D68DD9A0BB9A06F2" blockId="12.[151,1437,151,861]" box="[199,1437,151,177]" pageId="12" pageNumber="263">Elytra 1.6–1.9 times as broad as long, 1.3 times as long as pronotum, with narrowly impressed lateral margins.</paragraph>
<paragraph id="294A369BFFE56D54D68DD98CBC8C0696" blockId="12.[151,1437,151,861]" box="[199,651,186,213]" pageId="12" pageNumber="263">Metatarsi 1.5 times as long as metatibia.</paragraph>
<paragraph id="294A369BFFE56D54D68DD9E8BB7F07CA" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="263">
Male. Posterior margin of abdominal tergite VIII straight (
<figureCitation id="B1CE2A1EFFE56D54D53AD9E8BDCF06BA" box="[880,968,223,249]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="12" pageNumber="263">Fig. 22</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFE56D54D774D834BF93075E" box="[318,404,259,285]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="12" pageNumber="263">Fig. 23</figureCitation>
). Aedeagus with very wide basal portion, gradually narrowed toward small rounded apex; parameres moderately narrow, reaching apex of median lobe, slightly broadened apically, with rounded apices, each with four moderately long apical setae and two short setae in middle of inner margin; internal sac wide and short, with three elongate sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFE56D54D56CD858BD7C07CA" box="[806,891,367,393]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="12" pageNumber="263">Fig. 20</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFE56D54D368D858BB7307CA" box="[1314,1396,367,393]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="12" pageNumber="263">Fig. 21</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFE56D54D68DD8A4BD2B0792" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="263">
Female. Posterior margins of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFE56D54D56DD8A4BD7D07EE" box="[807,890,403,429]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="12" pageNumber="263">Fig. 24</figureCitation>
) and abdominal sternite VIII (
<figureCitation id="B1CE2A1EFFE56D54D29CD8A4BB2E07EE" box="[1238,1321,402,429]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="12" pageNumber="263">Fig. 25</figureCitation>
) rounded. Female accessory sclerite and spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFE56D54D68DD8EDBFBB04EA" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D68DD8EDBFB207B6" bold="true" box="[199,437,474,501]" pageId="12" pageNumber="263">Comparative notes.</emphasis>
Based on the general shape of the body with the presence of indistinct longitudinal impressions, the length and the coloration of the body, and the general shape of the aedeagus,
<taxonomicName id="EEF54D18FFE56D54D2FCD8C8BB3B045B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1206,1340,511,536]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="attenuata" status="sp. n.">
<emphasis id="1B81EA89FFE56D54D2FCD8C8BB3B045B" box="[1206,1340,511,536]" italics="true" pageId="12" pageNumber="263">P. attenuata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE56D54D309D8C8BB84045A" bold="true" box="[1347,1411,511,537]" pageId="12" pageNumber="263">
<taxonomicNameLabel id="00B257F2FFE56D54D309D8C8BB84045A" box="[1347,1411,511,537]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFE56D54D741DB14BC4D047E" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[267,586,547,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="gracillima">
<emphasis id="1B81EA89FFE56D54D741DB14BF1B047F" box="[267,284,547,572]" italics="true" pageId="12" pageNumber="263">P</emphasis>
.
<emphasis id="1B81EA89FFE56D54D766DB14BFA6047E" box="[300,417,547,573]" italics="true" pageId="12" pageNumber="263">gracillima</emphasis>
<bibRefCitation id="4D644B6AFFE56D54D7E3DB14BC4D047E" author="Shavrin, A. V." box="[425,586,547,573]" pageId="12" pageNumber="263" pagination="457 - 471" refId="ref35824" refString="Shavrin, A. V. (2022) New species and records of Omaliini McLeay, 1825 from Eastern Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Omaliinae). Zootaxa, 5169 (5), 457 - 471. https: // doi. org / 10.11646 / zootaxa. 5169.5.4" type="journal article" year="2022">Shavrin, 2022</bibRefCitation>
</taxonomicName>
, recently described from
<collectingCountry id="51E2760BFFE56D54D520DB14BDC9047E" box="[874,974,547,573]" name="Thailand" pageId="12" pageNumber="263">Thailand</collectingCountry>
(
<bibRefCitation id="4D644B6AFFE56D54D594DB14BA7B047E" author="Shavrin, A. V." box="[990,1148,547,573]" pageId="12" pageNumber="263" pagination="457 - 471" refId="ref35824" refString="Shavrin, A. V. (2022) New species and records of Omaliini McLeay, 1825 from Eastern Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Omaliinae). Zootaxa, 5169 (5), 457 - 471. https: // doi. org / 10.11646 / zootaxa. 5169.5.4" type="journal article" year="2022">Shavrin 2022</bibRefCitation>
). The new species can be distinguished from it by the concave laterobasal margins of the pronotum in front of hind angles, finer punctation of the head and the pronotum, the longer elytra, slightly broader apical portion of the median lobe and narrower apical portions of the parameres.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D54D68DDB85BF4D04B2" pageId="12" pageNumber="263" type="distribution">
<paragraph id="294A369BFFE56D54D68DDB85BF4D04B2" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D68DDB85BF66048F" bold="true" box="[199,353,690,716]" pageId="12" pageNumber="263">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFE56D54D721DB84BC9D048F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[363,666,691,717]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="attenuata" status="sp. nov.">
<emphasis id="1B81EA89FFE56D54D721DB84BC9D048F" box="[363,666,691,717]" italics="true" pageId="12" pageNumber="263">Paraphloeostiba attenuata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE56D54D4EEDB84BCE0048E" bold="true" box="[676,743,691,717]" pageId="12" pageNumber="263">
<taxonomicNameLabel id="00B257F2FFE56D54D4EEDB84BCE0048E" box="[676,743,691,717]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in central part of
<collectingCountry id="51E2760BFFE56D54D352DB84BEEF04B2" name="Papua New Guinea" pageId="12" pageNumber="263">Papua New Guinea</collectingCountry>
(Fig. x).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D54D68DDBCCBA0D051E" pageId="12" pageNumber="263" type="etymology">
<paragraph id="294A369BFFE56D54D68DDBCCBF93057A" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D68DDBCCBF490556" bold="true" box="[199,334,763,789]" pageId="12" pageNumber="263">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFE56D54D4F4DBCCBD340557" box="[702,819,763,788]" italics="true" pageId="12" pageNumber="263">attenuatus</emphasis>
, -
<emphasis id="1B81EA89FFE56D54D500DBCCBD5F0557" box="[842,856,763,788]" italics="true" pageId="12" pageNumber="263">a</emphasis>
, -
<emphasis id="1B81EA89FFE56D54D53ADBCCBD950557" box="[880,914,763,788]" italics="true" pageId="12" pageNumber="263">um</emphasis>
(narrowed to a point). It alludes to the general shape of the aedeagus.
</paragraph>
<paragraph id="294A369BFFE56D54D68DDA75BA0D051E" blockId="12.[151,1437,151,861]" box="[199,1034,834,861]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D68DDA75BF4C051F" bold="true" box="[199,331,834,860]" pageId="12" pageNumber="263">Bionomics.</emphasis>
<typeStatus id="F64E8839FFE56D54D718DA74BFB8051E" box="[338,447,835,861]" pageId="12" pageNumber="263" type="paratype">Paratypes</typeStatus>
were collected in cow dung, forest litter or in flight.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="EBED71EA82B1B354FD96E2DFF4B7C98C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AE72C5FD10C147F3C9D24DE9D3048EAD" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="FAB6C643E35F578CE1B24BC4D449127D" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFE56D56D6DDDA99BF00021A" LSID="urn:lsid:plazi:treatment:A15C878DFFE56D56D6DDDA99BF00021A" httpUri="http://treatment.plazi.org/id/A15C878DFFE56D56D6DDDA99BF00021A" lastPageId="14" lastPageNumber="265" pageId="12" pageNumber="263">
<subSubSection id="61EF6510FFE56D54D6DDDA99BC08058A" box="[151,527,942,969]" pageId="12" pageNumber="263" type="nomenclature">
<paragraph id="294A369BFFE56D54D6DDDA99BC08058A" blockId="12.[151,527,942,1005]" box="[151,527,942,969]" pageId="12" pageNumber="263">
<heading id="720281F7FFE56D54D6DDDA99BC08058A" bold="true" box="[151,527,942,969]" fontSize="11" level="1" pageId="12" pageNumber="263" reason="1">
<emphasis id="1B81EA89FFE56D54D6DDDA99BC08058A" bold="true" box="[151,527,942,969]" pageId="12" pageNumber="263">
<taxonomicName id="EEF54D18FFE56D54D6DDDA99BFC1058A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,454,942,969]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE56D54D6DDDA99BFC1058A" bold="true" box="[151,454,942,969]" italics="true" pageId="12" pageNumber="263">Paraphloeostiba barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFE56D54D787DA98BC08058A" box="[461,527,943,969]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D54D6DDDAE4BF4405AE" box="[151,323,978,1005]" pageId="12" pageNumber="263" type="description">
<paragraph id="294A369BFFE56D54D6DDDAE4BF4405AE" blockId="12.[151,527,942,1005]" box="[151,323,978,1005]" pageId="12" pageNumber="263">
(
<figureCitation id="B1CE2A1EFFE56D54D6D5DAE4BEEF05AE" box="[159,232,978,1005]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="12" pageNumber="263">Figs 5</figureCitation>
,
<figureCitation id="B1CE2A1EFFE56D54D6BEDAE4BF3B05AE" box="[244,316,979,1005]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="12" pageNumber="263">26–31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D54D6DDDD2CBCCD02AA" pageId="12" pageNumber="263" type="materials_examined">
<paragraph id="294A369BFFE56D54D6DDDD2CBA46023E" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D6DDDD2CBF4D0277" bold="true" box="[151,330,1050,1077]" pageId="12" pageNumber="263">Type material.</emphasis>
<typeStatus id="F64E8839FFE56D54D71CDD2CBFB90276" box="[342,446,1051,1077]" pageId="12" pageNumber="263" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFE56D54D780DD2CBFD80276" box="[458,479,1051,1077]" count="1" pageId="12" pageNumber="263" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFE56D54D7B9DD2CBC3B0276" box="[499,572,1050,1077]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="12" pageNumber="263">Fig. 5</figureCitation>
; right antennomeres 3–11 missing; dissected): ‘
<collectionCode id="4FE4AE5EFFE56D54D226DD2CBAB70276" box="[1132,1200,1051,1077]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="263">NEW</collectionCode>
<collectingCountry id="51E2760BFFE56D54D2F6DD2CBB210276" box="[1212,1318,1051,1077]" name="Guinea-Bissau" pageId="12" pageNumber="263">GUINEA</collectingCountry>
| Awande (ca.
<geoCoordinate id="4CC1505CFFE56D54D684DD08BF1B021A" box="[206,284,1087,1113]" degrees="6" direction="south" minutes="31" orientation="latitude" pageId="12" pageNumber="263" precision="925" value="-6.5166664">6°31’S</geoCoordinate>
<geoCoordinate id="4CC1505CFFE56D54D760DD09BF9E021A" box="[298,409,1086,1113]" degrees="145" direction="east" minutes="34" orientation="longitude" pageId="12" pageNumber="263" precision="925" value="145.56667">145°34’E</geoCoordinate>
), |
<date id="5D4B105BFFE56D54D782DD09BC4D021A" box="[456,586,1086,1113]" pageId="12" pageNumber="263" value="1965-11-21">21.XI.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE56D54D4ACDD08BCF8021A" box="[742,767,1087,1113]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="263">R</collectionCode>
. Hornabrook BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFE56D54D2BADD08BB8E021A" box="[1264,1417,1087,1113]" pageId="12" pageNumber="263" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE56D54D6DDDD54BF55023E" authorityName="Steel" authorityYear="1960" box="[151,338,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE56D54D6DDDD54BF55023E" box="[151,338,1123,1149]" italics="true" pageId="12" pageNumber="263">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE56D54D72FDD54BFC6023E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[357,449,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE56D54D72FDD54BFC6023E" box="[357,449,1123,1149]" italics="true" pageId="12" pageNumber="263">barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE56D54D783DD54BC0F023E" bold="true" box="[457,520,1123,1149]" pageId="12" pageNumber="263">
<taxonomicNameLabel id="00B257F2FFE56D54D783DD54BC0F023E" box="[457,520,1123,1149]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE56D54D4DBDD54BCA0023E" box="[657,679,1123,1149]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="263" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFE56D54D591DD54BA32023E" box="[987,1077,1123,1149]" pageId="12" pageNumber="263">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFE56D54D68DDDB0BCCD02AA" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
<typeStatus id="F64E8839FFE56D54D68DDDB0BF2E02E2" box="[199,297,1159,1185]" pageId="12" pageNumber="263" type="paratype">Paratype</typeStatus>
<specimenCount id="3FF3FD12FFE56D54D778DDB0BF4002E2" box="[306,327,1159,1185]" count="1" pageId="12" pageNumber="263" type="female">♀</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFE56D54D797DDB0BC2602E2" box="[477,545,1159,1185]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="263">NEW</collectionCode>
<collectingCountry id="51E2760BFFE56D54D460DDB0BC9302E2" box="[554,660,1159,1185]" name="Guinea-Bissau" pageId="12" pageNumber="263">GUINEA</collectingCountry>
| Border Ranges, Miarosa | (
<quantity id="EE0D9B7EFFE56D54D597DDB0BA2802E2" box="[989,1071,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="263" unit="m" value="1900.0">1900m</quantity>
, ca.
<geoCoordinate id="4CC1505CFFE56D54D22EDDB0BAD802E2" box="[1124,1247,1159,1185]" degrees="6" direction="south" minutes="31" orientation="latitude" pageId="12" pageNumber="263" precision="15" seconds="10" value="-6.519444">6°31’10’’S</geoCoordinate>
|
<geoCoordinate id="4CC1505CFFE56D54D2BCDDB1BB8902E2" box="[1270,1422,1158,1185]" degrees="145" direction="east" minutes="33" orientation="longitude" pageId="12" pageNumber="263" precision="15" seconds="00" value="145.55">145°33’00’’E</geoCoordinate>
|
<date id="5D4B105BFFE56D54D6DDDD9DBF1D0286" box="[151,282,1194,1221]" pageId="12" pageNumber="263" value="1965-11-16">16.XI.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE56D54D7ECDD9CBFB80286" box="[422,447,1195,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="263">R</collectionCode>
. Hornabrook
<collectionCode id="4FE4AE5EFFE56D54D412DD9CBC830286" box="[600,644,1195,1221]" country="United Kingdom" name="Bristol Museum" pageId="12" pageNumber="263">BM</collectionCode>
1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFE56D54D5C1DD9CBA1C0286" box="[907,1051,1195,1221]" pageId="12" pageNumber="263" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE56D54D264DD9CBAEE0286" authorityName="Steel" authorityYear="1960" box="[1070,1257,1195,1221]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE56D54D264DD9CBAEE0286" box="[1070,1257,1195,1221]" italics="true" pageId="12" pageNumber="263">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE56D54D2B1DD9CBB500286" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1275,1367,1195,1221]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE56D54D2B1DD9CBB500286" box="[1275,1367,1195,1221]" italics="true" pageId="12" pageNumber="263">barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE56D54D314DD9CBB9A0286" bold="true" box="[1374,1437,1195,1221]" pageId="12" pageNumber="263">
<taxonomicNameLabel id="00B257F2FFE56D54D314DD9CBB9A0286" box="[1374,1437,1195,1221]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE56D54D750DDF8BF3702AA" box="[282,304,1231,1257]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="263" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFE56D54D429DDF8BCBA02AA" box="[611,701,1231,1257]" pageId="12" pageNumber="263">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D56D68DDDC5BFDA05C2" lastPageId="14" lastPageNumber="265" pageId="12" pageNumber="263" type="description">
<paragraph id="294A369BFFE56D54D68DDDC5BA720316" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D68DDDC5BF5E034F" bold="true" box="[199,345,1266,1292]" pageId="12" pageNumber="263">Description.</emphasis>
Measurements (n=2): HW: 0.35; HL: 0.20–0.21; OL: 0.11–0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFE56D54D2ADDDC4BB57034E" box="[1255,1360,1267,1293]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
): 0.52; PL: 0.25–0.27; PW: 0.42–0.45; ESL: 0.40–0.42; EW: 0.52–0.55; MTbL (
<typeStatus id="F64E8839FFE56D54D591DC20BA430372" box="[987,1092,1303,1329]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
): 0.27; MTrL (
<typeStatus id="F64E8839FFE56D54D2A4DC20BB500372" box="[1262,1367,1303,1329]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
): 0.12 (MTrL 1–4: 0.05; MTrL 5: 0.07); AW: 0.50–0.52; AedL: 0.35; BL: 1.62(
<typeStatus id="F64E8839FFE56D54D58BDC0CBA2E0316" box="[961,1065,1339,1365]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
)–1.79.
</paragraph>
<paragraph id="294A369BFFE56D54D68DDC68BB5B00DA" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
Habitus as in
<figureCitation id="B1CE2A1EFFE56D54D722DC68BFA9033A" box="[360,430,1374,1401]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="12" pageNumber="263">Fig. 5</figureCitation>
. Head, pronotum and abdomen reddish-brown (pronotum slightly paler, with yellowish lateroapical portions in
<typeStatus id="F64E8839FFE56D54D7F8DCB4BC1E03DE" box="[434,537,1411,1437]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
); elytra yellowish-brown, with darkened mediobasal, lateroapical and apical portions; antennomeres 5–11 brown; mouthparts, antennomeres 1–4 and legs yellow. Head with irregular sparse and relatively large punctation, denser in middle; neck without visible punctation; pronotum with relatively large and dense punctation, similar to that in middle portion of head, larger and deeper in lateral and finer in mediobasal portions; punctation of elytra similar to that in pronotum, but distinctly finer and sparser in middle portions of each elytron along suture. Head with distinct dense microsculpture: transverse on vertex, subdiagonal in middle and longitudinal in infraorbital portions; pronotum with dense transverse microreticulation, finer in middle; elytra with dense subdiagonal microreticulation, finer along suture; abdominal tergites with very fine transverse meshes.
</paragraph>
<paragraph id="294A369BFFE56D54D68DDF94BF3A01FA" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">Head with distinct narrow and moderately deep anteriomedian depression, 1.6 times as broad as long, with short, narrow and relatively deep anteocellar foveae, subdiagonally stretching posteriad and reaching level of of anterior third length of eyes; postocular carina absent. Ocelli moderately large, located at level of posterior margins of eyes; distance between ocelli slightly larger than distance between ocellus and posterior margin of eyes. Antennomere 3 distinctly shorter and narrower than 2, 4 about twice as long as 3, 5 slightly broader than 4, 6 slightly longer and distinctly broader than 5, 7–9 slightly longer and broader than 6, 10 indistinctly longer than 9, apical antennomere about as long combined length of two preceeding segments, from about middle strongly narrowed apicad toward subacute apex.</paragraph>
<caption id="7D8A6613FFE46D55D6DDDF18BF5000CC" ID-DOI="http://doi.org/10.5281/zenodo.10821229" ID-Zenodo-Dep="10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="13" pageNumber="264" startId="13.[151,264,1583,1607]" targetBox="[187,1428,185,1557]" targetPageId="13" targetType="figure">
<paragraph id="294A369BFFE46D55D6DDDF18BF5000CC" blockId="13.[151,1437,1583,1680]" pageId="13" pageNumber="264">
<emphasis id="1B81EA89FFE46D55D6DDDF18BF50000B" bold="true" box="[151,343,1583,1608]" pageId="13" pageNumber="264">FIGURES 26–31.</emphasis>
<taxonomicName id="EEF54D18FFE46D55D72ADF07BC61000B" authorityName="Shavrin" authorityYear="2024" box="[352,614,1584,1608]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="species" species="barclayi">
<emphasis id="1B81EA89FFE46D55D72ADF07BC61000B" box="[352,614,1584,1608]" italics="true" pageId="13" pageNumber="264">Paraphloeostiba barclayi</emphasis>
</taxonomicName>
(holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFE46D55D68DDFD5BFD80107" blockId="13.[151,1437,1762,2040]" pageId="13" pageNumber="264">Pronotum 1.6 times as broad as long, 1.2 times as broad as head, widest in middle, more narrowed posteriad than anteriad; lateral impressions moderately deep, distinctly wider and deeper in laterobasal portions; lateral margins with fine regular crenulation.</paragraph>
<paragraph id="294A369BFFE46D55D68DDE79BB9A012B" blockId="13.[151,1437,1762,2040]" box="[199,1437,1870,1897]" pageId="13" pageNumber="264">Elytra 1.3 times as broad as long, 1.5–1.6 times as long as pronotum, with narrowly impressed lateral margins.</paragraph>
<paragraph id="294A369BFFE46D55D68DDE45BD3301CF" blockId="13.[151,1437,1762,2040]" box="[199,820,1906,1932]" pageId="13" pageNumber="264">Metatarsi slightly more than twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFE46D55D68DDEA1BC8501F3" blockId="13.[151,1437,1762,2040]" box="[199,642,1942,1968]" pageId="13" pageNumber="264">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="294A369BFFE46D56D68DDE8DBC86075E" blockId="13.[151,1437,1762,2040]" lastBlockId="14.[151,1437,151,1113]" lastPageId="14" lastPageNumber="265" pageId="13" pageNumber="264">
Male. Posterior margin of abdominal tergite VIII rounded (
<figureCitation id="B1CE2A1EFFE46D55D51FDE8DBDAF0197" box="[853,936,1978,2004]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="13" pageNumber="264">Fig. 28</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFE46D55D748DEE9BF5001BB" box="[258,343,2014,2040]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="13" pageNumber="264">Fig. 29</figureCitation>
). Aedeagus with wide basal portion, strongly narrowed toward narrow and long median lobe with small rounded apex; parameres narrow, distinctly exceeding apex of median lobe and distinctly broadened apically, each with two long apical and two preapical setae and with additional three short setae in about middle of inner margin; internal sac moderately long and narrow, with large elongate sclerotized structure in basal portion (
<figureCitation id="B1CE2A1EFFE76D56D376D9E8BB9706BA" box="[1340,1424,223,249]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 26</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFE76D56D461D834BC7A075E" box="[555,637,259,285]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 27</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFE76D56D68DD810BD2B0726" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
Female. Posterior margins of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFE76D56D562D810BD7C0702" box="[808,891,295,321]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 30</figureCitation>
) and abdominal sternite VIII (
<figureCitation id="B1CE2A1EFFE76D56D29DD810BB2D0702" box="[1239,1322,295,321]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 31</figureCitation>
) truncate. Female accessory sclerite and spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFE76D56D68DD858BA3107EE" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFE76D56D537D858BDD407CA" box="[893,979,367,393]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 30</figureCitation>
). Posterior margin of abdominal sternite VIII widely rounded (
<figureCitation id="B1CE2A1EFFE76D56D7C7D8A4BFE707EE" box="[397,480,403,429]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 31</figureCitation>
). Accessory sclerite and spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFE76D56D68DD881BCFC045A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DD881BFB60792" bold="true" box="[199,433,438,465]" pageId="14" pageNumber="265">Comparative notes.</emphasis>
Based on the very narrow median lobe and the parameres,
<taxonomicName id="EEF54D18FFE76D56D21AD880BACF0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1104,1224,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D21AD880BACF0792" box="[1104,1224,439,465]" italics="true" pageId="14" pageNumber="265">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D29AD880BB170792" bold="true" box="[1232,1296,439,465]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D29AD880BB170792" box="[1232,1296,439,465]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFE76D56D6DDD8ECBF4C07B7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,331,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D6DDD8ECBEAF07B7" box="[151,168,475,500]" italics="true" pageId="14" pageNumber="265">P</emphasis>
.
<emphasis id="1B81EA89FFE76D56D6FDD8ECBF4C07B7" box="[183,331,475,500]" italics="true" pageId="14" pageNumber="265">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D718D8ECBF9607B6" bold="true" box="[338,401,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D718D8ECBF9607B6" box="[338,401,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFE76D56D7D5D8ECBC2C07B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[415,555,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D7D5D8ECBC2C07B6" box="[415,555,475,501]" italics="true" pageId="14" pageNumber="265">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D478D8ECBC7507B6" bold="true" box="[562,626,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D478D8ECBC7507B6" box="[562,626,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFE76D56D435D8ECBD1207B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[639,789,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D435D8ECBD1207B6" box="[639,789,475,501]" italics="true" pageId="14" pageNumber="265">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D556D8ECBD5B07B6" bold="true" box="[796,860,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D556D8ECBD5B07B6" box="[796,860,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="EEF54D18FFE76D56D5D3D8ECBDF707B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[921,1008,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D5D3D8ECBDF707B6" box="[921,1008,475,501]" italics="true" pageId="14" pageNumber="265">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D5BDD8ECBA3007B6" bold="true" box="[1015,1079,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D5BDD8ECBA3007B6" box="[1015,1079,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFE76D56D68DDB14BF150422" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D74DDB14BFBA047F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[263,445,547,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D74DDB14BF1F047F" box="[263,280,547,572]" italics="true" pageId="14" pageNumber="265">P</emphasis>
.
<emphasis id="1B81EA89FFE76D56D763DB14BFBA047F" box="[297,445,547,572]" italics="true" pageId="14" pageNumber="265">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D78CDB14BC0F047E" bold="true" box="[454,520,547,573]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D78CDB14BC0F047E" box="[454,520,547,573]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller body, the paler elytra, slightly broader pronotum and the longer parameres;
</paragraph>
<paragraph id="294A369BFFE76D56D68DDB5CBDC504EA" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D74EDB5CBF8904C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,398,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D74EDB5CBF8904C6" box="[260,398,619,645]" italics="true" pageId="14" pageNumber="265">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D7DEDB5CBFD404C6" bold="true" box="[404,467,619,645]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D7DEDB5CBFD404C6" box="[404,467,619,645]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the larger body, the paler elytra, the presence of the punctation on the head, the finer punctation on the pronotum, the shorter elytra and the longer parameres;
</paragraph>
<paragraph id="294A369BFFE76D56D68DDB84BA8C04B2" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D749DB84BF90048E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[259,407,691,717]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D749DB84BF90048E" box="[259,407,691,717]" italics="true" pageId="14" pageNumber="265">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D7D7DB84BFDC048E" bold="true" box="[413,475,691,717]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D7D7DB84BFDC048E" box="[413,475,691,717]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker head, pronotum and abdomen, the presence of the punctation of the head, sparser and larger punctation of the pronotum, the shorter elytra and the longer parameres;
</paragraph>
<paragraph id="294A369BFFE76D56D68DDBCCBC01057A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D74EDBCCBF5C0556" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,347,763,789]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D74EDBCCBF5C0556" box="[260,347,763,789]" italics="true" pageId="14" pageNumber="265">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D728DBCCBFA60556" bold="true" box="[354,417,763,789]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D728DBCCBFA60556" box="[354,417,763,789]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the paler elytra, the absence of elevations between punctures on the elytra and narrower apical portions of the parameres.
</paragraph>
<paragraph id="294A369BFFE76D56D68DDA74BFDA05C2" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
From all these species,
<taxonomicName id="EEF54D18FFE76D56D799DA74BC80051F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[467,647,835,861]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D799DA74BFE3051F" box="[467,484,835,860]" italics="true" pageId="14" pageNumber="265">P</emphasis>
.
<emphasis id="1B81EA89FFE76D56D7B9DA74BC80051F" box="[499,647,835,860]" italics="true" pageId="14" pageNumber="265">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D4DADA74BCD6051E" bold="true" box="[656,721,835,861]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D4DADA74BCD6051E" box="[656,721,835,861]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the details of the external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE76D56D68DDABDBEF7058A" pageId="14" pageNumber="265" type="distribution">
<paragraph id="294A369BFFE76D56D68DDABDBEF7058A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DDABDBF6605E7" bold="true" box="[199,353,906,932]" pageId="14" pageNumber="265">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFE76D56D726DABCBC8905E6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[364,654,907,933]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE76D56D726DABCBC8905E6" box="[364,654,907,933]" italics="true" pageId="14" pageNumber="265">Paraphloeostiba barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D4D3DABCBCDA05E6" bold="true" box="[665,733,907,933]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D4D3DABCBCDA05E6" box="[665,733,907,933]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in central part of
<collectingCountry id="51E2760BFFE76D56D35DDABCBEEB058A" name="Papua New Guinea" pageId="14" pageNumber="265">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE76D56D68DDAE4BF00021A" pageId="14" pageNumber="265" type="etymology">
<paragraph id="294A369BFFE76D56D68DDAE4BF0E0252" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DDAE4BF4905AE" bold="true" box="[199,334,979,1005]" pageId="14" pageNumber="265">Etymology.</emphasis>
Patronymic, the species is named to honour of well-known coleopterologist Maxwell V.L. Barclay (London).
</paragraph>
<paragraph id="294A369BFFE76D56D68DDD2DBF00021A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DDD2DBF4C0277" bold="true" box="[199,331,1050,1076]" pageId="14" pageNumber="265">Bionomics.</emphasis>
The
<typeStatus id="F64E8839FFE76D56D7DBDD2CBFF60276" box="[401,497,1051,1077]" pageId="14" pageNumber="265" type="paratype">paratype</typeStatus>
was collected at an elevation
<quantity id="EE0D9B7EFFE76D56D523DD2CBDC30276" box="[873,964,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="265" unit="m" value="1900.0">1900 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

273
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<mods:title id="E6B5E3948E504492FFD835138BD3FAFA">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<mods:title id="3FFEE6231B6FD9ECFDFD0154406D7F6D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="50E68D34EC3ABA503E35AA9CF1BF210F">
<mods:date id="4902B28CF2E9855A3F16FB541E870987">2024</mods:date>
<mods:detail id="425D8656F2E66F8E9F3891B6B85E76D0" type="pubDate">
<mods:number id="DD748ED29189D764C974147B7E628568">2024-03-14</mods:number>
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<mods:number id="CD77C06DFB98B43637FB083C03A13886">5424</mods:number>
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<mods:number id="1A0823B3650D4E6FC2B8CDAA5FB5413E">3</mods:number>
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<mods:identifier id="1CA5AB0BF21C1E66923C62A0ACD610F0" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="001C9A2735BC991FBB78D0378FEB54EB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A15C878DFFE76D48D6DDDD95BAEB0557" LSID="urn:lsid:plazi:treatment:A15C878DFFE76D48D6DDDD95BAEB0557" httpUri="http://treatment.plazi.org/id/A15C878DFFE76D48D6DDDD95BAEB0557" lastPageId="16" lastPageNumber="267" pageId="14" pageNumber="265">
<subSubSection id="61EF6510FFE76D56D6DDDD95BC2302FE" box="[151,548,1186,1213]" pageId="14" pageNumber="265" type="nomenclature">
<paragraph id="294A369BFFE76D56D6DDDD95BC2302FE" blockId="14.[151,548,1186,1249]" box="[151,548,1186,1213]" pageId="14" pageNumber="265">
<heading id="720281F7FFE76D56D6DDDD95BC2302FE" bold="true" box="[151,548,1186,1213]" fontSize="11" level="1" pageId="14" pageNumber="265" reason="1">
<emphasis id="1B81EA89FFE76D56D6DDDD95BC2302FE" bold="true" box="[151,548,1186,1213]" pageId="14" pageNumber="265">
<taxonomicName id="EEF54D18FFE76D56D6DDDD95BFDC02FE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,475,1186,1213]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. nov.">
<emphasis id="1B81EA89FFE76D56D6DDDD95BFDC02FE" bold="true" box="[151,475,1186,1213]" italics="true" pageId="14" pageNumber="265">Paraphloeostiba betlephila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFE76D56D7A8DD94BC2302FE" box="[482,548,1187,1213]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE76D56D6DDDDF0BF4402A2" box="[151,323,1222,1249]" pageId="14" pageNumber="265" type="description">
<paragraph id="294A369BFFE76D56D6DDDDF0BF4402A2" blockId="14.[151,548,1186,1249]" box="[151,323,1222,1249]" pageId="14" pageNumber="265">
(
<figureCitation id="B1CE2A1EFFE76D56D6D5DDF0BEEF02A2" box="[159,232,1223,1249]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="14" pageNumber="265">Figs 6</figureCitation>
,
<figureCitation id="B1CE2A1EFFE76D56D6BEDDF1BF3C02A2" box="[244,315,1222,1249]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="14" pageNumber="265">32–35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE76D56D6DDDC38BFA60332" pageId="14" pageNumber="265" type="materials_examined">
<paragraph id="294A369BFFE76D56D6DDDC38BFA60332" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D6DDDC38BF42036B" bold="true" box="[151,325,1294,1321]" pageId="14" pageNumber="265">Type material.</emphasis>
<typeStatus id="F64E8839FFE76D56D706DC38BFB3036A" box="[332,436,1295,1321]" pageId="14" pageNumber="265" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFE76D56D7F6DC38BFD6036A" box="[444,465,1295,1321]" count="1" pageId="14" pageNumber="265" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFE76D56D7AADC38BC22036A" box="[480,549,1295,1321]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="14" pageNumber="265">Fig. 6</figureCitation>
; dissected): ‘
<collectingRegion id="EB31F879FFE76D56D4FFDC38BD15036A" box="[693,786,1295,1321]" country="Indonesia" name="Papua Barat" pageId="14" pageNumber="265">PAPUA</collectingRegion>
<taxonomicNameLabel id="00B257F2FFE76D56D553DC38BD54036A" box="[793,851,1295,1321]" pageId="14" pageNumber="265" rank="genus">N.G.</taxonomicNameLabel>
:
<collectingRegion id="EB31F879FFE76D56D515DC38BDFE036A" box="[863,1017,1295,1321]" country="Papua New Guinea" name="Morobe" pageId="14" pageNumber="265">Morobe Prov.</collectingRegion>
| Aseki,
<quantity id="EE0D9B7EFFE76D56D211DC38BAB7036A" box="[1115,1200,1295,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="265" unit="m" value="1700.0">1700 m</quantity>
|
<date id="5D4B105BFFE76D56D28EDC38BB30036A" box="[1220,1335,1295,1321]" pageId="14" pageNumber="265" value="1998-02-22">22.2.1998</date>
, betelnut - | flower, leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFE76D56D474DC04BCD0030E" box="[574,727,1331,1357]" pageId="14" pageNumber="265" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE76D56D4A4DC04BDAE030E" authorityName="Steel" authorityYear="1960" box="[750,937,1331,1357]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE76D56D4A4DC04BDAE030E" box="[750,937,1331,1357]" italics="true" pageId="14" pageNumber="265">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE76D56D5F5DC04BA28030E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[959,1071,1331,1357]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. nov.">
<emphasis id="1B81EA89FFE76D56D5F5DC04BA28030E" box="[959,1071,1331,1357]" italics="true" pageId="14" pageNumber="265">betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D27DDC04BA7F030E" bold="true" box="[1079,1144,1331,1357]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D27DDC04BA7F030E" box="[1079,1144,1331,1357]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE76D56D34CDC04BB1B030E" box="[1286,1308,1331,1357]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="265" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFE76D56D70DDC60BF920332" box="[327,405,1367,1393]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="265">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE76D48D68DDC4DBCCD0423" lastPageId="16" lastPageNumber="267" pageId="14" pageNumber="265" type="description">
<paragraph id="294A369BFFE76D56D68DDC4DBF45039E" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DDC4DBF5E03D7" bold="true" box="[199,345,1402,1428]" pageId="14" pageNumber="265">Description.</emphasis>
Measurements: HW: 0.30; HL: 0.23; OL: 0.10; TL: 0.02; AL: 0.57; PL: 0.25; PWmax: 0.45; PWmin: 0.38; ESL: 0.45; EW: 0.56; MTbL: 0.30; MTrL: 0.14 (MTrL 1–4: 0.04; MTrL 5: 0.10); AW: 0.57; AedL: 0.30; BL: 1.42.
</paragraph>
<paragraph id="294A369BFFE76D56D68DDCD0BEE4009A" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">
Habitus as in
<figureCitation id="B1CE2A1EFFE76D56D726DCD0BFB30042" box="[364,436,1511,1537]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="14" pageNumber="265">Fig. 6</figureCitation>
. Body and antennomeres 6–11 brown; mouthparts, antennomere 1–5 and legs yellow. Head and neck without punctation; pronotum with indistinct, fine and moderately dense punctation, invisible in mediobasal portion; elytra with distinct, very dense, moderately large and deep punctation, finer and sparser around scutellum and along suture. Head with dense coarse isodiametric microsculpture; pronotum with very dense isodiametric microsculpture, slightly finer than that in middle portion of head; scutellum with coarse isodiametric microreticulation; microsculpture of elytra very dense, transverse, finer around scutellum and longitudinal along suture.
</paragraph>
<paragraph id="294A369BFFE76D56D68DDFD4BF7401F2" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">Head 1.3 times as broad as long, with short and deep narrow anteocellar foveae almost reaching level of middle length of eyes. Ocelli very small, located about at level of posterior margins of eyes; distance between ocelli slightly larger than distance between ocellus and posterior margin of eyes. Antennae with slightly transverse antennomeres 6–8 and distinctly transverse antennomeres 9–10; antennomere 5 slightly broader than 4, 6 slightly longer and broader than 5, 7–8 distinctly longer and broader than 6, 9–10 slightly longer than 8, apical antennomere about 1.8 times as long as 10.</paragraph>
<paragraph id="294A369BFFE76D56D69ADE8CBDD801BA" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">Pronotum 1.8 times as broad as long, 1.5 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; laterobasal portions slightly and widely impressed.</paragraph>
<caption id="7D8A6613FFE66D57D6DDDF27BA2D0033" ID-DOI="http://doi.org/10.5281/zenodo.10821231" ID-Zenodo-Dep="10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266" startId="15.[151,264,1552,1576]" targetBox="[176,1428,191,1520]" targetPageId="15" targetType="figure">
<paragraph id="294A369BFFE66D57D6DDDF27BA2D0033" blockId="15.[151,1436,1552,1649]" pageId="15" pageNumber="266">
<emphasis id="1B81EA89FFE66D57D6DDDF27BF53006B" bold="true" box="[151,340,1552,1576]" pageId="15" pageNumber="266">FIGURES 32–38.</emphasis>
<taxonomicName id="EEF54D18FFE66D57D713DF27BC6A006B" authorityName="Shavrin" authorityYear="2024" box="[345,621,1552,1576]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="betlephila">
<emphasis id="1B81EA89FFE66D57D713DF27BC6A006B" box="[345,621,1552,1576]" italics="true" pageId="15" pageNumber="266">Paraphloeostiba betlephila</emphasis>
</taxonomicName>
(Figs. 32
<emphasis id="1B81EA89FFE66D57D487DF27BCDD006B" bold="true" box="[717,730,1552,1576]" pageId="15" pageNumber="266">–</emphasis>
35: holotype) and
<taxonomicName id="EEF54D18FFE66D57D5DBDF26BA0A006B" authorityName="Shavrin" authorityYear="2023" box="[913,1037,1553,1576]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFE66D57D5DBDF26BA0A006B" box="[913,1037,1553,1576]" italics="true" pageId="15" pageNumber="266">P. conjuncta</emphasis>
</taxonomicName>
(Figs. 36
<emphasis id="1B81EA89FFE66D57D227DF27BA7D006B" bold="true" box="[1133,1146,1552,1576]" pageId="15" pageNumber="266">–</emphasis>
38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFE66D57D68DDF89BDCF009A" blockId="15.[151,1437,1726,2040]" box="[199,968,1726,1753]" pageId="15" pageNumber="266">Elytra 1.2 times as broad as long and slightly shorter than pronotum.</paragraph>
<paragraph id="294A369BFFE66D57D68DDFD5BD2300BF" blockId="15.[151,1437,1726,2040]" box="[199,804,1762,1789]" pageId="15" pageNumber="266">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFE66D57D68DDE31BEE90197" blockId="15.[151,1437,1726,2040]" pageId="15" pageNumber="266">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFE66D57D51EDE31BDA00163" box="[852,935,1798,1824]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 34</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFE66D57D749DE1DBF500106" box="[259,343,1834,1861]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 35</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward very narrow apical portion, with small rounded apex; parameres slightly not reaching apex of median lobe, moderately wide, with rounded apices, each with three long apical and one preapical setae, apical half of inner margin with four to five short setae; internal sac moderately short and wide, without large sclerotized structures (
<figureCitation id="B1CE2A1EFFE66D57D5D7DEA1BDF401F3" box="[925,1011,1942,1968]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 32</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFE66D57D6DDDE8DBEED0197" box="[151,234,1978,2004]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 33</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFE66D57D68DDEE9BF9601BB" blockId="15.[151,1437,1726,2040]" box="[199,401,2014,2040]" pageId="15" pageNumber="266">Female unknown.</paragraph>
<paragraph id="294A369BFFF96D48D68DD9A1BCFC06BB" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
<emphasis id="1B81EA89FFF96D48D68DD9A1BFB506F3" bold="true" box="[199,434,150,176]" pageId="16" pageNumber="267">Comparative notes.</emphasis>
Based on the very narrow median lobe and the parameres,
<taxonomicName id="EEF54D18FFF96D48D21CD9A0BAE406F3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1110,1251,150,176]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D21CD9A0BAE406F3" box="[1110,1251,150,176]" italics="true" pageId="16" pageNumber="267">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D2A1D9A1BB2B06F3" bold="true" box="[1259,1324,150,176]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D2A1D9A1BB2B06F3" box="[1259,1324,150,176]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFF96D48D6FED98CBF600697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[180,359,187,213]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D6FED98CBEC20697" box="[180,197,187,212]" italics="true" pageId="16" pageNumber="267">P</emphasis>
.
<emphasis id="1B81EA89FFF96D48D699D98CBF600697" box="[211,359,187,212]" italics="true" pageId="16" pageNumber="267">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D727D98DBFAA0697" bold="true" box="[365,429,186,212]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D727D98DBFAA0697" box="[365,429,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFF96D48D7F0D98CBC360697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[442,561,186,212]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D7F0D98CBC360697" box="[442,561,186,212]" italics="true" pageId="16" pageNumber="267">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D472D98DBC7F0697" bold="true" box="[568,632,186,212]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D472D98DBC7F0697" box="[568,632,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFF96D48D4CFD98CBD1D0697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[645,794,186,212]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D4CFD98CBD1D0697" box="[645,794,186,212]" italics="true" pageId="16" pageNumber="267">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D56BD98DBD670697" bold="true" box="[801,864,186,212]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D56BD98DBD670697" box="[801,864,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="EEF54D18FFF96D48D5D7D98CBDF40697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[925,1011,186,212]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D5D7D98CBDF40697" box="[925,1011,186,212]" italics="true" pageId="16" pageNumber="267">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D5B0D98DBA3D0697" bold="true" box="[1018,1082,186,212]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D5B0D98DBA3D0697" box="[1018,1082,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFF96D48D68DD835BD760702" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
from
<taxonomicName id="EEF54D18FFF96D48D742D834BFB9075F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[264,446,259,285]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D742D834BF1E075F" box="[264,281,259,284]" italics="true" pageId="16" pageNumber="267">P</emphasis>
.
<emphasis id="1B81EA89FFF96D48D760D834BFB9075F" box="[298,446,259,284]" italics="true" pageId="16" pageNumber="267">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D782D835BC0D075F" bold="true" box="[456,522,258,284]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D782D835BC0D075F" box="[456,522,258,284]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the absence of the punctation of the head, the finer punctation of the more transverse pronotum, the longer elytra and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFF96D48D68DD87DBCFE07CA" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
from
<taxonomicName id="EEF54D18FFF96D48D74ED87CBF7C0727" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,379,330,356]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D74ED87CBF7C0727" box="[260,379,330,356]" italics="true" pageId="16" pageNumber="267">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D7C8D87DBFC60727" bold="true" box="[386,449,330,356]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D7C8D87DBFC60727" box="[386,449,330,356]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker body, the absence of the punctation of the head, the finer punctation on the pornotum, the longer elytra and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFF96D48D68DD8A5BAB207EE" blockId="16.[151,1437,150,789]" box="[199,1205,402,429]" pageId="16" pageNumber="267">
from
<taxonomicName id="EEF54D18FFF96D48D74FD8A4BF9D07EF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,410,402,428]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D74FD8A4BF9D07EF" box="[261,410,402,428]" italics="true" pageId="16" pageNumber="267">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D7EBD8A5BFE707EF" bold="true" box="[417,480,402,428]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D7EBD8A5BFE707EF" box="[417,480,402,428]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the slightly shorter elytra and moderately broader parameres;
</paragraph>
<paragraph id="294A369BFFF96D48D68DD881BF42045B" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
from
<taxonomicName id="EEF54D18FFF96D48D74FD880BF5C0793" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,347,438,464]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D74FD880BF5C0793" box="[261,347,438,464]" italics="true" pageId="16" pageNumber="267">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D729D881BFA50793" bold="true" box="[355,418,438,464]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D729D881BFA50793" box="[355,418,438,464]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the paler body, the absence of the punctation of the head and the presence of elevations between punctures on the elytra, the finer punctation of the pronotum, and slightly shorter parameres with narrower apical portions.
</paragraph>
<paragraph id="294A369BFFF96D48D68DDB14BCCD0423" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
From all these species,
<taxonomicName id="EEF54D18FFF96D48D793DB14BC6C047F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[473,619,546,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFF96D48D793DB14BFED047F" box="[473,490,547,572]" italics="true" pageId="16" pageNumber="267">P</emphasis>
.
<emphasis id="1B81EA89FFF96D48D7B1DB15BC6C047F" box="[507,619,546,572]" italics="true" pageId="16" pageNumber="267">betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D43FDB15BCBF047F" bold="true" box="[629,696,546,572]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D43FDB15BCBF047F" box="[629,696,546,572]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the smaller body and the details of the external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF96D48D68DDB5DBF2B04EB" pageId="16" pageNumber="267" type="distribution">
<paragraph id="294A369BFFF96D48D68DDB5DBF2B04EB" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
<emphasis id="1B81EA89FFF96D48D68DDB5DBF6604C7" bold="true" box="[199,353,618,644]" pageId="16" pageNumber="267">Distribution.</emphasis>
The new species is known only from the
<typeStatus id="F64E8839FFF96D48D563DB5CBD5E04C6" box="[809,857,619,645]" pageId="16" pageNumber="267">type</typeStatus>
locality in Morobi Province of eastern part of
<collectingCountry id="51E2760BFFF96D48D312DB5CBF2F04EB" name="Papua New Guinea" pageId="16" pageNumber="267">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF96D48D68DDB85BAEB0557" pageId="16" pageNumber="267" type="etymology">
<paragraph id="294A369BFFF96D48D68DDB85BB0B04B3" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
<emphasis id="1B81EA89FFF96D48D68DDB85BF49048F" bold="true" box="[199,334,690,716]" pageId="16" pageNumber="267">Etymology.</emphasis>
The name of the new species is derived from the fact that the
<typeStatus id="F64E8839FFF96D48D252DB85BA7D048F" box="[1048,1146,690,716]" pageId="16" pageNumber="267" type="holotype">holotype</typeStatus>
was collected in betelnut flower (
<taxonomicName id="EEF54D18FFF96D48D6BBDBE0BF6B04B3" box="[241,364,726,752]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="16" pageNumber="267" phylum="Tracheophyta" rank="species" species="betle">
<emphasis id="1B81EA89FFF96D48D6BBDBE0BF6B04B3" box="[241,364,726,752]" italics="true" pageId="16" pageNumber="267">Piper betle</emphasis>
</taxonomicName>
;
<taxonomicName id="EEF54D18FFF96D48D731DBE1BFFE04B3" box="[379,505,726,752]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="16" pageNumber="267" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
), the most commonly used as flavoring in chewing in Asian countries.
</paragraph>
<paragraph id="294A369BFFF96D48D68DDBCDBAEB0557" blockId="16.[151,1437,150,789]" box="[199,1260,762,789]" pageId="16" pageNumber="267">
<emphasis id="1B81EA89FFF96D48D68DDBCDBF4C0557" bold="true" box="[199,331,762,788]" pageId="16" pageNumber="267">Bionomics.</emphasis>
The
<typeStatus id="F64E8839FFF96D48D7CEDBCDBFE10557" box="[388,486,762,788]" pageId="16" pageNumber="267" type="holotype">holotype</typeStatus>
was collected in the flower of betelnut at an elevation of
<quantity id="EE0D9B7EFFF96D48D220DBCDBAB60556" box="[1130,1201,762,789]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="267" unit="m" value="170.0">170 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
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142
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<paragraph id="294A369BFFE86D59D6DDDFE9BCE600BB" blockId="1.[151,737,1758,1785]" box="[151,737,1758,1785]" pageId="1" pageNumber="252">
<heading id="720281F7FFE86D59D6DDDFE9BCE600BB" bold="true" box="[151,737,1758,1785]" fontSize="11" level="1" pageId="1" pageNumber="252" reason="1">
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The genus
<taxonomicName id="EEF54D18FFE86D59D754DFE9BCE600BB" ID-CoL="8XN7R" authority="Watanabe, 1962" authorityName="Watanabe" authorityYear="1962" box="[286,737,1758,1785]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE86D59D754DFE9BC1600BA" bold="true" box="[286,529,1758,1785]" italics="true" pageId="1" pageNumber="252">Nipponophloeostiba</emphasis>
<bibRefCitation id="4D644B6AFFE86D59D452DFE9BCE600BB" author="Watanabe, Y." box="[536,737,1758,1784]" pageId="1" pageNumber="252" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe, 1962</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
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<subSubSection id="61EF6510FFE86D5AD6DDDE10BB530696" lastPageId="2" lastPageNumber="253" pageId="1" pageNumber="252" type="description">
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The genus
<taxonomicName id="EEF54D18FFE86D59D75ADE10BFF40102" authorityName="Watanabe" authorityYear="1962" box="[272,499,1831,1857]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE86D59D75ADE10BFF40102" box="[272,499,1831,1857]" italics="true" pageId="1" pageNumber="252">Nipponophloeostiba</emphasis>
</taxonomicName>
was originally described for
<taxonomicName id="EEF54D18FFE86D59D57DDE1FBA890102" authority="Watanabe, 1962" authorityName="Watanabe" authorityYear="1962" box="[823,1166,1831,1858]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="verrucifera">
<emphasis id="1B81EA89FFE86D59D57DDE1FBD4D0102" box="[823,842,1832,1857]" italics="true" pageId="1" pageNumber="252">N</emphasis>
.
<emphasis id="1B81EA89FFE86D59D51DDE10BDD20102" box="[855,981,1831,1857]" italics="true" pageId="1" pageNumber="252">verrucifera</emphasis>
<bibRefCitation id="4D644B6AFFE86D59D591DE10BA890102" author="Watanabe, Y." box="[987,1166,1831,1857]" pageId="1" pageNumber="252" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe, 1962</bibRefCitation>
</taxonomicName>
from Izu islands,
<collectingCountry id="51E2760BFFE86D59D31FDE1FBB9F0101" box="[1365,1432,1832,1858]" name="Japan" pageId="1" pageNumber="252">Japan</collectingCountry>
. Based on the published (
<bibRefCitation id="4D644B6AFFE86D59D7F8DE7BBC610125" author="Watanabe, Y." box="[434,614,1868,1894]" pageId="1" pageNumber="252" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe 1962</bibRefCitation>
,
<bibRefCitation id="4D644B6AFFE86D59D439DE7BBD1D0125" author="Shavrin, A. V." box="[627,794,1868,1894]" pageId="1" pageNumber="252" pagination="196 - 200" refId="ref35701" refString="Shavrin, A. V. (2020 a) Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4802 (1), 196 - 200. https: // doi. org / 10.11646 / zootaxa. 4802.1.14" type="journal article" year="2020">
Shavrin 2020
<bibRefCitation id="4D644B6AFFE86D59D544DE7BBD1D0125" author="Shavrin, A. V." box="[782,794,1868,1894]" pageId="1" pageNumber="252" pagination="301 - 329" refId="ref35758" refString="Shavrin, A. V. (2020 b) New species and records of Paraphloeostiba Steel, 1960 from China and Laos, and descriptions of four news species of related genera (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4890 (3), 301 - 329. https: // doi. org / 10.11646 / zootaxa. 4890.3.1" type="journal article" year="2020">a</bibRefCitation>
</bibRefCitation>
, 2020b,
<bibRefCitation id="4D644B6AFFE86D59D537DE7BBDC40125" author="Shavrin, A. V." box="[893,963,1868,1894]" pageId="1" pageNumber="252" pagination="245 - 250" refId="ref35879" refString="Shavrin, A. V. (2023 a) Two new species and a new record of Omaliini McLeay, 1825 (Coleoptera: Staphylinidae: Omaliinae) from Indonesia. Zootaxa, 5258 (2), 245 - 250. https: // doi. org / 10.11646 / zootaxa. 5258.2.7" type="journal article" year="2023">2023a</bibRefCitation>
) and the present data, the genus contains five species from south-eastern part of the Palaearctic and Oriental regions:
<taxonomicName id="EEF54D18FFE86D59D5ACDE46BB1F01C9" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[998,1304,1904,1930]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="explanata">
<emphasis id="1B81EA89FFE86D59D5ACDE46BA7301C9" box="[998,1140,1904,1930]" italics="true" pageId="1" pageNumber="252">N. explanata</emphasis>
Shavrin, 2023
</taxonomicName>
(
<collectingCountry id="51E2760BFFE86D59D36CDE47BB9E01C9" box="[1318,1433,1904,1930]" name="Indonesia" pageId="1" pageNumber="252">Indonesia</collectingCountry>
:
<collectingRegion id="EB31F879FFE86D59D6DDDEA2BECE01EC" box="[151,201,1941,1967]" country="Indonesia" name="Bali" pageId="1" pageNumber="252">Bali</collectingRegion>
),
<taxonomicName id="EEF54D18FFE86D59D694DEA1BC3C01EC" authority="Watanabe, 1962" authorityName="Watanabe" authorityYear="1962" box="[222,571,1941,1967]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="verrucifera">
<emphasis id="1B81EA89FFE86D59D694DEA1BF7901EC" box="[222,382,1941,1967]" italics="true" pageId="1" pageNumber="252">N. verrucifera</emphasis>
<bibRefCitation id="4D644B6AFFE86D59D7CCDEA2BC3C01EC" author="Watanabe, Y." box="[390,571,1941,1967]" pageId="1" pageNumber="252" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe, 1962</bibRefCitation>
</taxonomicName>
(south
<collectingCountry id="51E2760BFFE86D59D4D8DEA2BCD101EC" box="[658,726,1941,1967]" name="Japan" pageId="1" pageNumber="252">Japan</collectingCountry>
),
<taxonomicName id="EEF54D18FFE86D59D4A0DEA1BA0501EC" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[746,1026,1941,1967]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thayeri">
<emphasis id="1B81EA89FFE86D59D4A0DEA1BD5C01EC" box="[746,859,1941,1967]" italics="true" pageId="1" pageNumber="252">N. thayeri</emphasis>
Shavrin, 2020
</taxonomicName>
(
<collectingCountry id="51E2760BFFE86D59D258DEA2BA7901EC" box="[1042,1150,1941,1967]" name="Malaysia" pageId="1" pageNumber="252">Malaysia</collectingCountry>
),
<taxonomicName id="EEF54D18FFE86D59D2D9DEA1BEC80190" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="longilobata">
<emphasis id="1B81EA89FFE86D59D2D9DEA1BB3101EC" box="[1171,1334,1941,1967]" italics="true" pageId="1" pageNumber="252">N. longilobata</emphasis>
Shavrin, 2020
</taxonomicName>
(
<collectingCountry id="51E2760BFFE86D59D6A8DE8EBF1A0190" box="[226,285,1977,2003]" name="Laos" pageId="1" pageNumber="252">Laos</collectingCountry>
), and
<taxonomicName id="EEF54D18FFE86D59D723DE8DBC1F0190" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[361,536,1977,2003]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="hornabrooki" status="sp. n.">
<emphasis id="1B81EA89FFE86D59D723DE8DBC1F0190" box="[361,536,1977,2003]" italics="true" pageId="1" pageNumber="252">N. hornabrooki</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE86D59D469DE8EBC600190" bold="true" box="[547,615,1977,2003]" pageId="1" pageNumber="252">
<taxonomicNameLabel id="00B257F2FFE86D59D469DE8EBC600190" box="[547,615,1977,2003]" pageId="1" pageNumber="252" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="51E2760BFFE86D59D433DE8EBD5A0190" box="[633,861,1977,2004]" name="Papua New Guinea" pageId="1" pageNumber="252">Papua New Guinea</collectingCountry>
) (
<bibRefCitation id="4D644B6AFFE86D59D53CDE8EBA2B0190" author="Watanabe, Y." box="[886,1068,1977,2003]" pageId="1" pageNumber="252" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe 1962</bibRefCitation>
,
<bibRefCitation id="4D644B6AFFE86D59D271DE8EBAE30190" author="Shavrin, A. V." box="[1083,1252,1977,2003]" pageId="1" pageNumber="252" pagination="196 - 200" refId="ref35701" refString="Shavrin, A. V. (2020 a) Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4802 (1), 196 - 200. https: // doi. org / 10.11646 / zootaxa. 4802.1.14" type="journal article" year="2020">
Shavrin 2020
<bibRefCitation id="4D644B6AFFE86D59D292DE8EBAE30190" author="Shavrin, A. V." box="[1240,1252,1977,2003]" pageId="1" pageNumber="252" pagination="301 - 329" refId="ref35758" refString="Shavrin, A. V. (2020 b) New species and records of Paraphloeostiba Steel, 1960 from China and Laos, and descriptions of four news species of related genera (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4890 (3), 301 - 329. https: // doi. org / 10.11646 / zootaxa. 4890.3.1" type="journal article" year="2020">a</bibRefCitation>
</bibRefCitation>
, 2020b, present study). The distribution of the genus is shown in the
<figureCitation id="B1CE2A1EFFE86D59D54BDEE9BD5201BB" box="[769,853,2014,2040]" captionStart="FIGURE 11" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[151,1435,525,1610]" captionTargetId="figure-143@5.[151,1436,525,1610]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of Nipponophloeostiba: N. verrucifera (circles), N. thayeri (triangle), N. longilobata (square), N. explanata (diamond), N. hornabrooki (star)." figureDoi="http://doi.org/10.5281/zenodo.10821220" httpUri="https://zenodo.org/record/10821220/files/figure.png" pageId="1" pageNumber="252">Fig. 11</figureCitation>
. Species of this genus have flattened and elongate body, well-developed postocular parts of the head, two elongate carinae in basal portion of the head and two small tubercles near apical margins of the pronotum (for details see
<bibRefCitation id="4D644B6AFFEB6D5AD506D98CBDFC0696" author="Watanabe, Y." box="[844,1019,187,213]" pageId="2" pageNumber="253" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe 1962</bibRefCitation>
,
<bibRefCitation id="4D644B6AFFEB6D5AD24CD98CBA460696" author="Watanabe, Y." box="[1030,1089,187,213]" pageId="2" pageNumber="253" pagination="59 - 391" refId="ref36201" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">1990</bibRefCitation>
,
<bibRefCitation id="4D644B6AFFEB6D5AD206D98CBAF60696" author="Shavrin, A. V." box="[1100,1265,187,213]" pageId="2" pageNumber="253" pagination="196 - 200" refId="ref35701" refString="Shavrin, A. V. (2020 a) Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4802 (1), 196 - 200. https: // doi. org / 10.11646 / zootaxa. 4802.1.14" type="journal article" year="2020">
Shavrin 2020
<bibRefCitation id="4D644B6AFFEB6D5AD2AFD98CBAF60696" author="Shavrin, A. V." box="[1253,1265,187,213]" pageId="2" pageNumber="253" pagination="301 - 329" refId="ref35758" refString="Shavrin, A. V. (2020 b) New species and records of Paraphloeostiba Steel, 1960 from China and Laos, and descriptions of four news species of related genera (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4890 (3), 301 - 329. https: // doi. org / 10.11646 / zootaxa. 4890.3.1" type="journal article" year="2020">a</bibRefCitation>
</bibRefCitation>
, 2020b).
</paragraph>
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223
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(
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,
<figureCitation id="B1CE2A1EFFEB6D5AD6BED877BF2B0719" box="[244,300,320,346]" captionStart-0="FIGURES 7–10" captionStart-1="FIGURE 11" captionStartId-0="4.[151,264,1664,1688]" captionStartId-1="5.[151,250,1633,1657]" captionTargetBox-0="[171,1427,192,1632]" captionTargetBox-1="[151,1435,525,1610]" captionTargetId-0="figure-22@4.[151,1435,181,1641]" captionTargetId-1="figure-143@5.[151,1436,525,1610]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." captionText-1="FIGURE 11. Distribution of Nipponophloeostiba: N. verrucifera (circles), N. thayeri (triangle), N. longilobata (square), N. explanata (diamond), N. hornabrooki (star)." figureDoi-0="http://doi.org/10.5281/zenodo.10821218" figureDoi-1="http://doi.org/10.5281/zenodo.10821220" httpUri-0="https://zenodo.org/record/10821218/files/figure.png" httpUri-1="https://zenodo.org/record/10821220/files/figure.png" pageId="2" pageNumber="253">7–11</figureCitation>
)
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<emphasis id="1B81EA89FFEB6D5AD6DDD8BFBF4307E1" bold="true" box="[151,324,392,418]" pageId="2" pageNumber="253">Type material.</emphasis>
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<specimenCount id="3FF3FD12FFEB6D5AD7F3D8BEBFC907E0" box="[441,462,393,419]" count="1" pageId="2" pageNumber="253" type="male">♂</specimenCount>
(
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; specimen without left middle leg and right elytron; dissected): ‘
<collectionCode id="4FE4AE5EFFEB6D5AD2A2D8BFBB2B07E1" box="[1256,1324,392,418]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="2" pageNumber="253">NEW</collectionCode>
<collectingCountry id="51E2760BFFEB6D5AD378D8BFBB9B07E1" box="[1330,1436,392,418]" name="Guinea-Bissau" pageId="2" pageNumber="253">GUINEA</collectingCountry>
| Okapa, |
<date id="5D4B105BFFEB6D5AD742D89BBF7A0784" box="[264,381,428,455]" pageId="2" pageNumber="253" value="1965-05-28">28.V.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFEB6D5AD441D89BBC230785" box="[523,548,428,454]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="253">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFEB6D5AD5B5D89BBA9F0785" box="[1023,1176,428,454]" pageId="2" pageNumber="253" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFEB6D5AD2E6D89BBB880785" authorityName="Watanabe" authorityYear="1962" box="[1196,1423,428,454]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFEB6D5AD2E6D89BBB880785" box="[1196,1423,428,454]" italics="true" pageId="2" pageNumber="253">Nipponophloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFEB6D5AD6DDD8E7BF2507A9" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,290,464,490]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="253" phylum="Arthropoda" rank="species" species="hornabrooki" status="sp. nov.">
<emphasis id="1B81EA89FFEB6D5AD6DDD8E7BF2507A9" box="[151,290,464,490]" italics="true" pageId="2" pageNumber="253">hornabrooki</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFEB6D5AD763D8E7BF6E07A9" bold="true" box="[297,361,464,490]" pageId="2" pageNumber="253">
<taxonomicNameLabel id="00B257F2FFEB6D5AD763D8E7BF6E07A9" box="[297,361,464,490]" pageId="2" pageNumber="253" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFEB6D5AD7B8D8E7BC0F07A9" box="[498,520,464,490]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="253" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFEB6D5AD571D8E7BD9207A9" box="[827,917,464,490]" pageId="2" pageNumber="253">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFEB6D5DD68DD8C3BD280726" lastPageId="5" lastPageNumber="256" pageId="2" pageNumber="253" type="description">
<paragraph id="294A369BFFEB6D5AD68DD8C3BF450415" blockId="2.[151,1437,392,1931]" pageId="2" pageNumber="253">
<emphasis id="1B81EA89FFEB6D5AD68DD8C3BF5E044D" bold="true" box="[199,345,500,526]" pageId="2" pageNumber="253">Description.</emphasis>
Measurements: HW: 0.47; HL: 0.30; OL: 0.15; TL: 0.05; AL: 0.61; PL: 0.40; PWmax: 0.57; PWmin: 0.40; ESL: 0.62; EW: 0.60; MTbL: 0.30; MTrL: 0.15 (MTrL 1–4: 0.05; MTrL 5: 0.10); AW: 0.60; AedL: 0.27; BL: 2.34.
</paragraph>
<paragraph id="294A369BFFEB6D5AD68DDB57BC1B05A0" blockId="2.[151,1437,392,1931]" pageId="2" pageNumber="253">
Habitus as in
<figureCitation id="B1CE2A1EFFEB6D5AD72CDB57BFAD0439" box="[358,426,608,634]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="2" pageNumber="253">Fig. 1</figureCitation>
. Head, pronotum and abdomen reddish-brown; elytra yellow-brown, with reddish apical portion; antennomeres 6–11 brown; mouthparts, antennomeres 1–5, intersegmental membranes between abdominal tergites and legs yellowish. Head with sparse and fine punctation, slightly denser in middle; neck with several very fine punctures in middle; pronotum with very fine punctation, similar to that in head, but distinctly denser, finer in lateral and sparser and finer in mediobasal portions; scutellum without punctures; elytra with moderately dense punctation, slightly deeper than that in pronotum, finer and sparser around scutellum and along suture; abdomen without visible punctation. Head with dense and coarse microsculpture, transverse in apical, longitudinal in middle and isodiametric in lateral portions; neck with very fine transverse microreticulation; pronotum with very dense isodiametric microsculpture, longitudinal in mediobasal and transverse in laterobasal portions; scutellum with dense and coarse transverse meshes; abdomen with dense and coarse microreticulation, transverse in abdominal tergites III–V and isodiametric in VI–VIII.
</paragraph>
<paragraph id="294A369BFFEB6D5AD68DDADBBB0A03BC" blockId="2.[151,1437,392,1931]" pageId="2" pageNumber="253">Head 1.5 times as broad as long, with moderately wide and shallow anteriomedian depressions with distinct narrow elevated carina along outer margin of each depression, reaching level of apical third of eyes; anteocellar foveae (grooves in front of ocelli) deep and long, linear, extending about level of middle length of eyes; basal margin straight; posterior portions between ocelli and temples with narrow, distinctly elevated straight carina, reaching laterobasal margin of head; each basal margin of infraorbital portion near carina with small oval depression, with setae on bottom of it; temples (postocular parts) three times shorter than longitudinal length of eyes, convex, not expanded laterad. Eyes moderately large, convex. Ocelli moderately large, located very close to neck constriction, distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Maxillary palpi long; preapical palpomere indistinctly longer than wide; apical palpomere narrow, about 1.3 times as long as preapical segment, from apical third gradually narrowed toward acute apex. Antennae almost reaching basal margin of pronotum when reclined; antennomeres 6–10 distinctly transverse; basal antennomere moderately wide, slightly more than twice as long as broad, antennomere 2 distinctly shorter and slightly narrower than basal antennomere, 3 slightly longer and narrower than 2, 4 twice shorter than 3, 5 about as long as wide, slightly broader than 4, 6 distinctly broader than 5, 7–9 slightly longer and distinctly broader than 6, 10 indistinctly longer than 9, apical antennomere about twice longer than 10, from about middle gradually narrowed towar subacute apex.</paragraph>
<paragraph id="294A369BFFEB6D5AD68DDF3EBAFC00F1" blockId="2.[151,1437,392,1931]" pageId="2" pageNumber="253">Pronotum transverse, 1.4 times as broad as long, slightly broader than head, widest in preapical portion, strongly narrowed posteriad toward obtuse hind angles; apical angles widely rounded; anterior margin slightly concave in middle, with two distinct, relatively small elevated tubercles at level of small oval depression near carina of head; posterior margin rounded; median part without longitudinal band, only with short and narrow trace of it in mediobasal portion; lateral portions narrowly bordered, with slightly impressed laterobasal portions.</paragraph>
<paragraph id="294A369BFFEB6D5AD68DDF8BBB6F00B9" blockId="2.[151,1437,392,1931]" pageId="2" pageNumber="253">Elytra slightly longer than broad, reaching basal margin of abdominal tergite III, 1.5 times as long as pronotum, somewhat parallelsided, slightly concave in middle, with straight apical margins. Hind wings fully developed.</paragraph>
<paragraph id="294A369BFFEB6D5AD68DDE32BC75015D" blockId="2.[151,1437,392,1931]" box="[199,626,1796,1823]" pageId="2" pageNumber="253">Metatarsus twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFEB6D5AD68DDE1FBC0301C8" blockId="2.[151,1437,392,1931]" pageId="2" pageNumber="253">Abdomen indistinctly widened toward abdominal tergite V, about as broad as elytra, with two small oval wing-folding patches (tomentose spots) in middle of abdominal tergite IV and distinct narrow palisade fringe at apical margin of abdominal tergite VII.</paragraph>
<caption id="7D8A6613FFEA6D5BD6DDDE75BEC101E1" ID-DOI="http://doi.org/10.5281/zenodo.10821216" ID-Zenodo-Dep="10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="3" pageNumber="254" startId="3.[151,264,1858,1882]" targetBox="[164,1423,192,1823]" targetPageId="3" targetType="figure">
<paragraph id="294A369BFFEA6D5BD6DDDE75BEC101E1" blockId="3.[151,1437,1858,1954]" pageId="3" pageNumber="254">
<emphasis id="1B81EA89FFEA6D5BD6DDDE75BF3C0119" bold="true" box="[151,315,1858,1882]" pageId="3" pageNumber="254">FIGURES 1–6.</emphasis>
Habitus of
<taxonomicName id="EEF54D18FFEA6D5BD7FFDE75BC080119" box="[437,527,1858,1882]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="tribe" tribe="Omaliini">
<emphasis id="1B81EA89FFEA6D5BD7FFDE75BC080119" box="[437,527,1858,1882]" italics="true" pageId="3" pageNumber="254">Omaliini</emphasis>
</taxonomicName>
: 1–
<taxonomicName id="EEF54D18FFEA6D5BD47DDE75BD8D0119" authorityName="Shavrin" authorityYear="2024" box="[567,906,1858,1882]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="hornabrooki">
<emphasis id="1B81EA89FFEA6D5BD47DDE75BD8D0119" box="[567,906,1858,1882]" italics="true" pageId="3" pageNumber="254">Nipponophloeostiba hornabrooki</emphasis>
</taxonomicName>
(holotype), 2–
<taxonomicName id="EEF54D18FFEA6D5BD268DE75BB220119" authorityName="Shavrin" authorityYear="2024" box="[1058,1317,1858,1882]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="assimile">
<emphasis id="1B81EA89FFEA6D5BD268DE75BB220119" box="[1058,1317,1858,1882]" italics="true" pageId="3" pageNumber="254">Paraphloeostiba assimile</emphasis>
</taxonomicName>
(holotype), 3–
<taxonomicName id="EEF54D18FFEA6D5BD6FBDE50BF56013D" authorityName="Shavrin" authorityYear="2024" box="[177,337,1895,1918]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="atramentaria">
<emphasis id="1B81EA89FFEA6D5BD6FBDE50BF56013D" box="[177,337,1895,1918]" italics="true" pageId="3" pageNumber="254">P. atramentaria</emphasis>
</taxonomicName>
(holotype), 4–
<taxonomicName id="EEF54D18FFEA6D5BD7A1DE50BC62013D" authorityName="Shavrin" authorityYear="2024" box="[491,613,1895,1918]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="1B81EA89FFEA6D5BD7A1DE50BC62013D" box="[491,613,1895,1918]" italics="true" pageId="3" pageNumber="254">P. attenuata</emphasis>
</taxonomicName>
(paratype), 5–
<taxonomicName id="EEF54D18FFEA6D5BD4B7DE50BD6C013D" authorityName="Shavrin" authorityYear="2024" box="[765,875,1894,1918]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="barclayi">
<emphasis id="1B81EA89FFEA6D5BD4B7DE50BD6C013D" box="[765,875,1894,1918]" italics="true" pageId="3" pageNumber="254">P. barclayi</emphasis>
</taxonomicName>
(paratype), 6–
<taxonomicName id="EEF54D18FFEA6D5BD24EDE50BA84013D" authorityName="Shavrin" authorityYear="2024" box="[1028,1155,1894,1918]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="betlephila">
<emphasis id="1B81EA89FFEA6D5BD24EDE50BA84013D" box="[1028,1155,1894,1918]" italics="true" pageId="3" pageNumber="254">P. betlephila</emphasis>
</taxonomicName>
(holotype). Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="7D8A6613FFED6D5CD6DDDFB7BD9900FF" ID-DOI="http://doi.org/10.5281/zenodo.10821218" ID-Zenodo-Dep="10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="4" pageNumber="255" startId="4.[151,264,1664,1688]" targetBox="[171,1427,192,1632]" targetPageId="4" targetType="figure">
<paragraph id="294A369BFFED6D5CD6DDDFB7BD9900FF" blockId="4.[151,1436,1664,1725]" pageId="4" pageNumber="255">
<emphasis id="1B81EA89FFED6D5CD6DDDFB7BF4F00DB" bold="true" box="[151,328,1664,1688]" pageId="4" pageNumber="255">FIGURES 7–10.</emphasis>
<taxonomicName id="EEF54D18FFED6D5CD705DFB7BCA600DB" authorityName="Shavrin" authorityYear="2024" box="[335,673,1664,1688]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="hornabrooki">
<emphasis id="1B81EA89FFED6D5CD705DFB7BCA600DB" box="[335,673,1664,1688]" italics="true" pageId="4" pageNumber="255">Nipponophloeostiba hornabrooki</emphasis>
</taxonomicName>
(holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFED6D5CD68DDE38BAFE019E" blockId="4.[151,1437,1807,2013]" pageId="4" pageNumber="255">
Male. Posterior margin of abdominal tergite VIII somewhat straight (
<figureCitation id="B1CE2A1EFFED6D5CD5ABDE38BA20016A" box="[993,1063,1807,1833]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="4" pageNumber="255">Fig. 9</figureCitation>
). Posterior margin of abdominal sternite VIII widely sinuate (
<figureCitation id="B1CE2A1EFFED6D5CD79BDE04BC25010E" box="[465,546,1843,1869]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="4" pageNumber="255">Fig. 10</figureCitation>
).Aedeagus short, with wide median lobe and very wide truncate apex; parameres moderately narrow, reaching apex of median lobe, slightly widened in apical portions, each with lateral obtuse tooth on inner margin of preapical portion, with three to four moderarely short apical setae and three short setae along inner margin of each paramere; internal sac short and relatively wide, with long flagellum, starting from basal portion and reaching apical third of median lobe (
<figureCitation id="B1CE2A1EFFED6D5CD489DEF4BD00019E" box="[707,775,1987,2013]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="4" pageNumber="255">Fig. 7</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFED6D5CD2FADEF4BAF3019E" box="[1200,1268,1987,2013]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="4" pageNumber="255">Fig. 8</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFEC6D5DD68DD9A0BF9606F2" blockId="5.[151,1437,151,501]" box="[199,401,151,177]" pageId="5" pageNumber="256">Female unknown.</paragraph>
<paragraph id="294A369BFFEC6D5DD68DD98DBD280726" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="256">
<emphasis id="1B81EA89FFEC6D5DD68DD98DBFB30696" bold="true" box="[199,436,186,213]" pageId="5" pageNumber="256">Comparative notes.</emphasis>
Based on the body size, the shape of postocular portions of the head slightly broader than pronotum and relatively short elytra, slightly broader than pronotum,
<taxonomicName id="EEF54D18FFEC6D5DD5A5D9E8BA9A06BA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1007,1181,223,249]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="hornabrooki" status="sp. n.">
<emphasis id="1B81EA89FFEC6D5DD5A5D9E8BA9A06BA" box="[1007,1181,223,249]" italics="true" pageId="5" pageNumber="256">N. hornabrooki</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFEC6D5DD2ECD9E8BAEF06BA" bold="true" box="[1190,1256,223,249]" pageId="5" pageNumber="256">
<taxonomicNameLabel id="00B257F2FFEC6D5DD2ECD9E8BAEF06BA" box="[1190,1256,223,249]" pageId="5" pageNumber="256" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFEC6D5DD3C9D9E8BFAC075E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="explanata">
<emphasis id="1B81EA89FFEC6D5DD3C9D9E8BB9106BB" box="[1411,1430,223,248]" italics="true" pageId="5" pageNumber="256">N</emphasis>
.
<emphasis id="1B81EA89FFEC6D5DD6DDD834BF02075E" box="[151,261,259,285]" italics="true" pageId="5" pageNumber="256">explanata</emphasis>
Shavrin, 2023
</taxonomicName>
, recently described from
<collectingRegion id="EB31F879FFEC6D5DD489D834BCF5075E" box="[707,754,259,285]" country="Indonesia" name="Bali" pageId="5" pageNumber="256">Bali</collectingRegion>
(
<bibRefCitation id="4D644B6AFFEC6D5DD4B5D834BDA0075E" author="Shavrin, A. V." box="[767,935,259,285]" pageId="5" pageNumber="256" pagination="245 - 250" refId="ref35879" refString="Shavrin, A. V. (2023 a) Two new species and a new record of Omaliini McLeay, 1825 (Coleoptera: Staphylinidae: Omaliinae) from Indonesia. Zootaxa, 5258 (2), 245 - 250. https: // doi. org / 10.11646 / zootaxa. 5258.2.7" type="journal article" year="2023">Shavrin 2023a</bibRefCitation>
). It can be distinguished from it by the longer temples, paler elytra and the different morphology of the aedeagus. From all species of the genus it differs by the broader median lobe and internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFEC6D5DD68DD859BFD907EE" pageId="5" pageNumber="256" type="distribution">
<paragraph id="294A369BFFEC6D5DD68DD859BFD907EE" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="256">
<emphasis id="1B81EA89FFEC6D5DD68DD859BF6607CB" bold="true" box="[199,353,366,392]" pageId="5" pageNumber="256">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFEC6D5DD723D858BCE707CA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[361,736,367,393]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="hornabrooki" status="sp. nov.">
<emphasis id="1B81EA89FFEC6D5DD723D858BCE707CA" box="[361,736,367,393]" italics="true" pageId="5" pageNumber="256">Nipponophloeostiba hornabrooki</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFEC6D5DD4A3D858BD2D07CA" bold="true" box="[745,810,367,393]" pageId="5" pageNumber="256">
<taxonomicNameLabel id="00B257F2FFEC6D5DD4A3D858BD2D07CA" box="[745,810,367,393]" pageId="5" pageNumber="256" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFEC6D5DD20DD858BA7007CA" box="[1095,1143,367,393]" pageId="5" pageNumber="256">type</typeStatus>
locality in central part of
<collectingCountry id="51E2760BFFEC6D5DD6DDD8A4BF6807EE" box="[151,367,403,429]" name="Papua New Guinea" pageId="5" pageNumber="256">Papua New Guinea</collectingCountry>
(
<figureCitation id="B1CE2A1EFFEC6D5DD735D8A4BFD707EE" box="[383,464,403,429]" captionStart="FIGURE 11" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[151,1435,525,1610]" captionTargetId="figure-143@5.[151,1436,525,1610]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of Nipponophloeostiba: N. verrucifera (circles), N. thayeri (triangle), N. longilobata (square), N. explanata (diamond), N. hornabrooki (star)." figureDoi="http://doi.org/10.5281/zenodo.10821220" httpUri="https://zenodo.org/record/10821220/files/figure.png" pageId="5" pageNumber="256">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFEC6D5DD68DD880BD3007B6" pageId="5" pageNumber="256" type="etymology">
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<emphasis id="1B81EA89FFEC6D5DD68DD880BF490792" bold="true" box="[199,334,439,465]" pageId="5" pageNumber="256">Etymology.</emphasis>
The species is named after R. Hornabrook, the collector ot the
<typeStatus id="F64E8839FFEC6D5DD258D880BA450792" box="[1042,1090,439,465]" pageId="5" pageNumber="256">type</typeStatus>
material.
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<emphasis id="1B81EA89FFEC6D5DD68DD8EDBF4C07B7" bold="true" box="[199,331,474,500]" pageId="5" pageNumber="256">Bionomics.</emphasis>
The detailed bionomical data are unknown.
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Key to species of the genus
<taxonomicName id="EEF54D18FFEC6D5DD7A6DFD8BCD80149" ID-CoL="8XN7R" authorityName="Watanabe" authorityYear="1962" box="[492,735,1775,1802]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFEC6D5DD7A6DFD8BCD80149" bold="true" box="[492,735,1775,1802]" italics="true" pageId="5" pageNumber="256">Nipponophloeostiba</emphasis>
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(modified from Shavrin 2020)
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1 Postocular portions of head distinctly widened lateroposteriad, with short posterior carina not reaching posterior margin of eye. Aedeagus as in
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Figs.
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. Head and pronotum reddish-brown, elytra and abdomen yellow-brown. Body length:
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. South
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..............................................
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<bibRefCitation id="4D644B6AFFEC6D5DD348DE58BB9B01C6" author="Watanabe, Y." box="[1282,1436,1903,1925]" pageId="5" pageNumber="256" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe, 1962</bibRefCitation>
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<keyLead id="920495AEFFEC6D5DD6DDDEBABB9B01FC" pageId="5" pageNumber="256">- Postocular portions of head not or indistinctly widened lateroposteriad, with longer posterior carina reaching posterior margin of eye or basal margin of ptonotum. Coloration different...................................................... 2</keyLead>
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2 Head as broad as pronotum. Elytra moderately long, significantly longer than pronotum. Median lobe and parameres narrow as in
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Fig.
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in Shavrin (2020). Head and abdomen brown, pronotum reddish-brown. Body length:
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....................................................................................
<taxonomicName id="EEF54D18FFEF6D5ED2FAD9AFBB9B06EE" ID-CoL="8Y6P9" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[1200,1436,151,173]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="thayeri">
<emphasis id="1B81EA89FFEF6D5ED2FAD9AFBB1706EF" box="[1200,1296,151,173]" italics="true" pageId="6" pageNumber="257">N. thayeri</emphasis>
Shavrin, 2020
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<keyLead id="920495AEFFEF6D5ED6DDD982BB9B06A4" pageId="6" pageNumber="257">- Head slightly narrower or broader than pronotum. Elytra shorter, slightly broader than pronotum. Median lobes and parameres different. Coloration different........................................................................... 3</keyLead>
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3 Head slightly narrower than pronotum.Aedeagus as in
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Fig.
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in Shavrin (2020). Head, pronotum and abdomen reddish-brown, elytra yellow-brown, with darker parascutellar portion. Body length:
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.
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,
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(
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)............................................................................................
<taxonomicName id="EEF54D18FFEF6D5ED2CFD81CBB9B077C" ID-CoL="8Y6P8" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[1157,1436,298,320]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="longilobata">
<emphasis id="1B81EA89FFEF6D5ED2CFD81CBB17077C" box="[1157,1296,298,320]" italics="true" pageId="6" pageNumber="257">N. longilobata</emphasis>
Shavrin, 2020
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<keyLead id="920495AEFFEF6D5ED6DDD87FBB9B071E" box="[151,1436,327,349]" pageId="6" pageNumber="257">- Head slightly broader than pronotum. Aedeagus and coloration different. Body larger............................... 4</keyLead>
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<keyLead id="920495AEFFEF6D5ED6DDD852BB9B07F6" pageId="6" pageNumber="257">
4 Temples slightly shorter than longitudinal length of eyes. Elytra as broad as long. Body reddish-brown. Median lobe narrow; parameres strongly broadened in apical portions as in
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Fig.
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in
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. Habitus as in
<figureCitation id="B1CE2A1EFFEF6D5ED220D8B5BAA507D4" box="[1130,1186,386,408]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">
Fig.
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in
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. Body length:
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.
<collectingCountry id="51E2760BFFEF6D5ED7C6D8A8BFEE07F6" box="[396,489,415,437]" name="Indonesia" pageId="6" pageNumber="257">Indonesia</collectingCountry>
(
<collectingRegion id="EB31F879FFEF6D5ED7BCD8A8BC2707F6" box="[502,544,415,437]" country="Indonesia" name="Bali" pageId="6" pageNumber="257">Bali</collectingRegion>
)...................................................
<taxonomicName id="EEF54D18FFEF6D5ED2DFD897BB9B07F6" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1173,1436,415,437]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="explanata">
<emphasis id="1B81EA89FFEF6D5ED2DFD897BAA207F6" box="[1173,1189,416,437]" italics="true" pageId="6" pageNumber="257">N</emphasis>
.
<emphasis id="1B81EA89FFEF6D5ED2FBD8A8BB1707F7" box="[1201,1296,415,436]" italics="true" pageId="6" pageNumber="257">explanata</emphasis>
Shavrin, 2023
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="294A369BFFEF6D5ED6DDD88ABB9B044E" blockId="6.[151,1437,151,525]" pageId="6" pageNumber="257">
<keyLead id="920495AEFFEF6D5ED6DDD88ABB9B044E" pageId="6" pageNumber="257">
- Temples three times shorter than longitudinal length of eyes. Elytra slightly longer than broad. Head, pronotum and abdomen reddish-brown; elytra yellow-brown, with reddish apical portion. Median lobe wide; parameres slightly broadened in apical portions (
<figureCitation id="B1CE2A1EFFEF6D5ED774D8C0BF7E044E" box="[318,377,503,525]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="6" pageNumber="257">Fig. 7</figureCitation>
). Habitus as in
<figureCitation id="B1CE2A1EFFEF6D5ED444D8C0BC40044E" box="[526,583,503,525]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">Fig. 1</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFEF6D5ED49FD8CFBD2A044E" box="[725,813,504,525]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" pageId="6" pageNumber="257" unit="mm" value="2.34">2.34 mm</quantity>
.
<collectingCountry id="51E2760BFFEF6D5ED57CD8CFBDE8044E" box="[822,1007,503,525]" name="Papua New Guinea" pageId="6" pageNumber="257">Papua New Guinea</collectingCountry>
.................
<taxonomicName id="EEF54D18FFEF6D5ED287D8CFBB67044F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1229,1376,503,525]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="hornabrooki" status="sp. n.">
<emphasis id="1B81EA89FFEF6D5ED287D8CFBADA044E" box="[1229,1245,504,525]" italics="true" pageId="6" pageNumber="257">N</emphasis>
.
<emphasis id="1B81EA89FFEF6D5ED2A3D8C0BB67044F" box="[1257,1376,503,524]" italics="true" pageId="6" pageNumber="257">hornabrooki</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFEF6D5ED32CD8CFBB9B044E" bold="true" box="[1382,1436,504,525]" pageId="6" pageNumber="257">
<taxonomicNameLabel id="00B257F2FFEF6D5ED32CD8CFBB9B044E" box="[1382,1436,504,525]" pageId="6" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

271
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@ -0,0 +1,271 @@
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<mods:title id="DE37064EC42A7C435FE427F19EC196D6">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<subSubSection id="61EF6510FFEF6D5ED6DDDADCBC080245" box="[151,527,1003,1030]" pageId="6" pageNumber="257" type="nomenclature">
<paragraph id="294A369BFFEF6D5ED6DDDADCBC080245" blockId="6.[151,527,1003,1067]" box="[151,527,1003,1030]" pageId="6" pageNumber="257">
<heading id="720281F7FFEF6D5ED6DDDADCBC080245" bold="true" box="[151,527,1003,1030]" fontSize="11" level="1" pageId="6" pageNumber="257" reason="1">
<emphasis id="1B81EA89FFEF6D5ED6DDDADCBC080245" bold="true" box="[151,527,1003,1030]" pageId="6" pageNumber="257">
<taxonomicName id="EEF54D18FFEF6D5ED6DDDADCBFC10245" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,454,1003,1030]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="assimile" status="sp. nov.">
<emphasis id="1B81EA89FFEF6D5ED6DDDADCBFC10245" bold="true" box="[151,454,1003,1030]" italics="true" pageId="6" pageNumber="257">Paraphloeostiba assimile</emphasis>
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<taxonomicNameLabel id="00B257F2FFEF6D5ED787DADBBC080245" box="[461,527,1004,1030]" pageId="6" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="61EF6510FFEF6D5ED6DDDD27BF440268" box="[151,323,1040,1067]" pageId="6" pageNumber="257" type="description">
<paragraph id="294A369BFFEF6D5ED6DDDD27BF440268" blockId="6.[151,527,1003,1067]" box="[151,323,1040,1067]" pageId="6" pageNumber="257">
(
<figureCitation id="B1CE2A1EFFEF6D5ED6D5DD27BEEF0269" box="[159,232,1040,1066]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">Figs 2</figureCitation>
,
<figureCitation id="B1CE2A1EFFEF6D5ED6BEDD27BF3C0268" box="[244,315,1040,1067]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="6" pageNumber="257">12–15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFEF6D5ED6DDDD6FBF0202F9" pageId="6" pageNumber="257" type="materials_examined">
<paragraph id="294A369BFFEF6D5ED6DDDD6FBF0202F9" blockId="6.[151,1437,1112,2038]" pageId="6" pageNumber="257">
<emphasis id="1B81EA89FFEF6D5ED6DDDD6FBF440231" bold="true" box="[151,323,1112,1138]" pageId="6" pageNumber="257">Type material.</emphasis>
<typeStatus id="F64E8839FFEF6D5ED703DD6FBFB60231" box="[329,433,1112,1138]" pageId="6" pageNumber="257" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFEF6D5ED7FCDD6FBFCC0231" box="[438,459,1112,1138]" count="1" pageId="6" pageNumber="257" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFEF6D5ED793DD6FBC1A0231" box="[473,541,1112,1138]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">Fig. 2</figureCitation>
; dissected): ‘
<collectionCode id="4FE4AE5EFFEF6D5ED4E2DD6FBCEB0231" box="[680,748,1112,1138]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="257">NEW</collectionCode>
<collectingCountry id="51E2760BFFEF6D5ED4B8DD6FBD5B0231" box="[754,860,1112,1138]" name="Guinea-Bissau" pageId="6" pageNumber="257">GUINEA</collectingCountry>
| Karimui, |
<date id="5D4B105BFFEF6D5ED5ABDD6FBA650231" box="[993,1122,1112,1138]" pageId="6" pageNumber="257" value="1966-03-13">13.III.1966</date>
’ &lt;printed&gt;,‘
<collectionCode id="4FE4AE5EFFEF6D5ED2A7DD6FBB060231" box="[1261,1281,1112,1138]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="257">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFEF6D5ED782DD4BBC6602D5" box="[456,609,1148,1174]" pageId="6" pageNumber="257" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFEF6D5ED439DD4BBD2902D5" authorityName="Steel" authorityYear="1960" box="[627,814,1148,1174]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFEF6D5ED439DD4BBD2902D5" box="[627,814,1148,1174]" italics="true" pageId="6" pageNumber="257">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFEF6D5ED575DD4BBD9C02D5" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[831,923,1148,1174]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="assimile" status="sp. nov.">
<emphasis id="1B81EA89FFEF6D5ED575DD4BBD9C02D5" box="[831,923,1148,1174]" italics="true" pageId="6" pageNumber="257">assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFEF6D5ED5EBDD4BBDD802D5" bold="true" box="[929,991,1148,1174]" pageId="6" pageNumber="257">
<taxonomicNameLabel id="00B257F2FFEF6D5ED5EBDD4BBDD802D5" box="[929,991,1148,1174]" pageId="6" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFEF6D5ED22CDD4BBA7B02D5" box="[1126,1148,1148,1174]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="257" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFEF6D5ED6D5DD97BEFE02F9" box="[159,249,1184,1210]" pageId="6" pageNumber="257">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFEF6D50D68DDDF3BCA4057A" lastPageId="8" lastPageNumber="259" pageId="6" pageNumber="257" type="description">
<paragraph id="294A369BFFEF6D5ED68DDDF3BB9A0341" blockId="6.[151,1437,1112,2038]" pageId="6" pageNumber="257">
<emphasis id="1B81EA89FFEF6D5ED68DDDF3BF5E029D" bold="true" box="[199,345,1220,1246]" pageId="6" pageNumber="257">Description.</emphasis>
Measurements: HW: 0.36; HL: 0.21; OL: 0.11; AL: 0.55; PL: 0.27; PWmax: 0.50; PWmin: 0.43; ESL: 0.50; EW: 0.56; MTbL: 0.26; MTrL: 0.13 (MTrL 1–4: 0.03; MTrL 5: 0.10); AW: 0.64; AedL: 0.25; BL: 1.76.
</paragraph>
<paragraph id="294A369BFFEF6D5ED68DDC3BBD890005" blockId="6.[151,1437,1112,2038]" pageId="6" pageNumber="257">
Habitus as in
<figureCitation id="B1CE2A1EFFEF6D5ED723DC3BBFB70365" box="[361,432,1292,1318]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">Fig. 2</figureCitation>
. Body yellow-brown, with reddish-brown apical portions of elytra; antennomeres 5–11 brown; mouthparts, antennomeres 1–4 and legs yellow. Head and neck without visible punctation; pronotum with indistinct and sparse fine punctation, slightly more visible in lateral portions; scutellum without punctures; elytra with fine indistinct punctation, slightly larger than that on pronotum, more indistinct in middle; abdominal tergites with indistinct fine and sparse punctation. Head with very dense and coarse microsculpture, transverse on vertex and isodiametric in middle portion; neck with fine transverse meshes; pronotum with very dense isodiametric microreticulation similar to that in middle portion of head; scutellum with fine transverse microreticulation; microsculpture of elytra very dense, isodiametric, similar to that in pronotum, slightly transverse in mediobasal portion; abdominal tergites with dense isodiametric microsculpture.
</paragraph>
<paragraph id="294A369BFFEF6D5ED68DDF67BA7001ED" blockId="6.[151,1437,1112,2038]" pageId="6" pageNumber="257">Head indistinctly elevated in middle, with shallow wide anteriomedian depressions, 1.7 times as broad as long, with short and deep anteocellar foveae, reaching level of posterior third length of eyes; postocular carina indistinct, acute, located very close to posterior margin of eyes. Ocelli moderately large, located slightly below level of posterior margins of eyes; distance between ocelli 1.4 times larger than distance between ocellus and posterior margin of eyes. Apical maxillary palpomere very narrow, about 1.7 times as long as preapical segment. Antenna reaching basal margin of pronotum when reclined, with distinctly transverse antennomeres 6–10; basal antennomere about twice as long as broad, antennomere 2 distinctly shorter and slightly narrower than basal antennomere, 3 distinctly narrower than 2, 4 about as long as wide, about twice shorter than 3, 5 slightly shorter than 4, 6 slightly longer and distinctly broader than 5, 7–9 slightly longer and distinctly broader than 6, 10 slightly longer than 9, apical antennomere about twice as long as 10, from about middle gradually narrowed apicad toward rounded apex.</paragraph>
<paragraph id="294A369BFFEF6D5FD68DDE8FBA1B0696" blockId="6.[151,1437,1112,2038]" lastBlockId="7.[151,1437,151,393]" lastPageId="7" lastPageNumber="258" pageId="6" pageNumber="257">Pronotum slightly convex, transverse, 1.8 times as broad as long, 1.3 times as broad as head, widest in middle, gradually narrowed both anteriad and posteriad; anterior angles widely rounded, not protruded anteriad; posterior angles obtuse; anterior margin slightly concave in middle, slightly shorter than rounded posterior margin; middle portion without impressions; laterobasal impressions moderately wide and deep.</paragraph>
<paragraph id="294A369BFFEE6D5FD68DD9E8BEF4075E" blockId="7.[151,1437,151,393]" pageId="7" pageNumber="258">Elytra slightly broader than long, 1.8 times as long as pronotum, slightly broadened posteriad; posterior margin straight.</paragraph>
<paragraph id="294A369BFFEE6D5FD68DD810BC7F0702" blockId="7.[151,1437,151,393]" box="[199,632,295,321]" pageId="7" pageNumber="258">Metatarsi twice shorter than metatibia.</paragraph>
<paragraph id="294A369BFFEE6D5FD68DD87CBA8707CA" blockId="7.[151,1437,151,393]" pageId="7" pageNumber="258">Abdomen slightly broader than elytra, with two very small and round wing-folding patches in middle of abdominal tergite IV; apical margin of abdominal tergite VII with narrow palisade fringe.</paragraph>
<caption id="7D8A6613FFEE6D5FD6DDDE8ABD9901BB" ID-DOI="http://doi.org/10.5281/zenodo.10821223" ID-Zenodo-Dep="10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="7" pageNumber="258" startId="7.[151,264,1981,2005]" targetBox="[160,1427,418,1948]" targetPageId="7" targetType="figure">
<paragraph id="294A369BFFEE6D5FD6DDDE8ABD9901BB" blockId="7.[151,1437,1981,2041]" pageId="7" pageNumber="258">
<emphasis id="1B81EA89FFEE6D5FD6DDDE8ABF500196" bold="true" box="[151,343,1981,2005]" pageId="7" pageNumber="258">FIGURES 12–15.</emphasis>
<taxonomicName id="EEF54D18FFEE6D5FD72ADE8ABC610196" authorityName="Shavrin" authorityYear="2024" box="[352,614,1981,2005]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="assimile">
<emphasis id="1B81EA89FFEE6D5FD72ADE8ABC610196" box="[352,614,1981,2005]" italics="true" pageId="7" pageNumber="258">Paraphloeostiba assimile</emphasis>
</taxonomicName>
(holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFE16D50D68DD9A0BA500726" blockId="8.[151,1436,151,1005]" pageId="8" pageNumber="259">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFE16D50D539D9A0BDCE06F2" box="[883,969,151,177]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="8" pageNumber="259">Fig. 13</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFE16D50D455D98CBC710696" box="[543,630,187,213]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="8" pageNumber="259">Fig. 14</figureCitation>
). Aedeagus with very wide basal portion, gradually narrowed toward relatively wide median lobe with rounded apex; parameres significantly longer than apex of median lobe, from middle significantly widened apically toward rounded apices, each with four to five moderately short setae, with three short setae on inner margin in about middle; internal sac wide and moderately long, with several elongate sclerotized structures in middle (
<figureCitation id="B1CE2A1EFFE16D50D44FD87CBC5E0726" box="[517,601,331,357]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="8" pageNumber="259">Fig. 12</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFE16D50D24AD87CBA540726" box="[1024,1107,331,357]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="8" pageNumber="259">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFE16D50D68DD858BF9607CA" blockId="8.[151,1436,151,1005]" box="[199,401,367,393]" pageId="8" pageNumber="259">Female unknown.</paragraph>
<paragraph id="294A369BFFE16D50D68DD8A5BD19045A" blockId="8.[151,1436,151,1005]" pageId="8" pageNumber="259">
<emphasis id="1B81EA89FFE16D50D68DD8A5BFB707EE" bold="true" box="[199,432,402,429]" pageId="8" pageNumber="259">Comparative notes.</emphasis>
Based on the very broad median lobe and apical portions of the parameres,
<taxonomicName id="EEF54D18FFE16D50D2B7D8A4BB7307EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1277,1396,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFE16D50D2B7D8A4BB7307EE" box="[1277,1396,403,429]" italics="true" pageId="8" pageNumber="259">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D331D8A4BEA90792" bold="true" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D331D8A4BEA90792" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFE16D50D70ED880BC5C0792" authority="Steel, 1960" authorityName="Steel" authorityYear="1960" box="[324,603,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFE16D50D70ED880BFD30792" box="[324,468,439,465]" italics="true" pageId="8" pageNumber="259">P. celebensis</emphasis>
<bibRefCitation id="4D644B6AFFE16D50D796D880BC5C0792" author="Steel, W. O." box="[476,603,439,465]" pageId="8" pageNumber="259" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel, 1960</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="51E2760BFFE16D50D421D880BCD90792" box="[619,734,439,465]" name="Indonesia" pageId="8" pageNumber="259">Indonesia</collectingCountry>
:
<collectingRegion id="EB31F879FFE16D50D4A0D880BD540792" box="[746,851,439,465]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="259">Sulawesi</collectingRegion>
),
<taxonomicName id="EEF54D18FFE16D50D52DD880BDE10793" authorityName="Cameron" authorityYear="1931" box="[871,998,439,464]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFE16D50D52DD880BDE10793" box="[871,998,439,464]" italics="true" pageId="8" pageNumber="259">P. papuana</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFE16D50D5BFD880BA920792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1013,1173,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFE16D50D5BFD880BA010793" box="[1013,1030,439,464]" italics="true" pageId="8" pageNumber="259">P</emphasis>
.
<emphasis id="1B81EA89FFE16D50D25FD880BA920792" box="[1045,1173,439,465]" italics="true" pageId="8" pageNumber="259">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D2D7D880BADA0792" bold="true" box="[1181,1245,439,465]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D2D7D880BADA0792" box="[1181,1245,439,465]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFE16D50D2A6D880BB4B0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1260,1356,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFE16D50D2A6D880BB4B0792" box="[1260,1356,439,465]" italics="true" pageId="8" pageNumber="259">P. rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D31FD880BB910792" bold="true" box="[1365,1430,439,465]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D31FD880BB910792" box="[1365,1430,439,465]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="EEF54D18FFE16D50D68DD8ECBC1B07B6" authority="(Cameron, 1940)" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[199,540,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFE16D50D68DD8ECBF5407B6" box="[199,339,475,501]" italics="true" pageId="8" pageNumber="259">P. tonkinesis</emphasis>
(Cameron, 1940)
</taxonomicName>
, known from
<collectingCountry id="51E2760BFFE16D50D4F3D8ECBD1F07B6" box="[697,792,475,501]" name="Vietnam" pageId="8" pageNumber="259">Vietnam</collectingCountry>
and
<collectingCountry id="51E2760BFFE16D50D505D8ECBDC607B6" box="[847,961,475,501]" name="Singapore" pageId="8" pageNumber="259">Singapore</collectingCountry>
(
<bibRefCitation id="4D644B6AFFE16D50D59AD8ECBA4A07B6" author="Steel, W. O." box="[976,1101,475,501]" pageId="8" pageNumber="259" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960</bibRefCitation>
). It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFE16D50D68DDB14BDBB047E" blockId="8.[151,1436,151,1005]" box="[199,956,547,573]" pageId="8" pageNumber="259">
from
<taxonomicName id="EEF54D18FFE16D50D74FDB14BF94047E" authorityName="Steel" authorityYear="1960" box="[261,403,547,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFE16D50D74FDB14BF94047E" box="[261,403,547,573]" italics="true" pageId="8" pageNumber="259">P. celebensis</emphasis>
</taxonomicName>
by the absence of visible punctation on the head;
</paragraph>
<paragraph id="294A369BFFE16D50D68DDB70BA910422" blockId="8.[151,1436,151,1005]" box="[199,1174,583,609]" pageId="8" pageNumber="259">
from
<taxonomicName id="EEF54D18FFE16D50D74FDB70BF850423" authorityName="Cameron" authorityYear="1931" box="[261,386,583,608]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFE16D50D74FDB70BF850423" box="[261,386,583,608]" italics="true" pageId="8" pageNumber="259">P. papuana</emphasis>
</taxonomicName>
by the absence of punctation on the head and slightly narrower elytra;
</paragraph>
<paragraph id="294A369BFFE16D50D68DDB5CBD7F04C6" blockId="8.[151,1436,151,1005]" box="[199,888,619,645]" pageId="8" pageNumber="259">
from
<taxonomicName id="EEF54D18FFE16D50D74FDB5CBFA304C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,420,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFE16D50D74FDB5CBF1104C7" box="[261,278,619,644]" italics="true" pageId="8" pageNumber="259">P</emphasis>
.
<emphasis id="1B81EA89FFE16D50D76EDB5CBFA304C6" box="[292,420,619,645]" italics="true" pageId="8" pageNumber="259">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D7E0DB5CBFED04C6" bold="true" box="[426,490,619,645]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D7E0DB5CBFED04C6" box="[426,490,619,645]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the slightly narrower pronotum;
</paragraph>
<paragraph id="294A369BFFE16D50D68DDBB8BAB004EA" blockId="8.[151,1436,151,1005]" box="[199,1207,655,681]" pageId="8" pageNumber="259">
from
<taxonomicName id="EEF54D18FFE16D50D74FDBB8BF6304EA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,356,655,681]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFE16D50D74FDBB8BF6304EA" box="[261,356,655,681]" italics="true" pageId="8" pageNumber="259">P. rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D726DBB8BFAC04EA" bold="true" box="[364,427,655,681]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D726DBB8BFAC04EA" box="[364,427,655,681]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller body, the darker elytra and more transverse pronotum;
</paragraph>
<paragraph id="294A369BFFE16D50D68DDB84BD0E04B2" blockId="8.[151,1436,151,1005]" pageId="8" pageNumber="259">
from
<taxonomicName id="EEF54D18FFE16D50D742DB84BF91048E" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[264,406,691,717]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFE16D50D742DB84BF91048E" box="[264,406,691,717]" italics="true" pageId="8" pageNumber="259">P. tonkinesis</emphasis>
</taxonomicName>
by the longer body, the absence of punctation on the head and the presence of traces of longitudinal impressions in the middle of the pronotum.
</paragraph>
<paragraph id="294A369BFFE16D50D68DDBCCBCA4057A" blockId="8.[151,1436,151,1005]" pageId="8" pageNumber="259">
From all these species,
<taxonomicName id="EEF54D18FFE16D50D784DBCCBC420556" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[462,581,763,789]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFE16D50D784DBCCBC420556" box="[462,581,763,789]" italics="true" pageId="8" pageNumber="259">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D406DBCCBC8B0556" bold="true" box="[588,652,763,789]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D406DBCCBC8B0556" box="[588,652,763,789]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the longer parameres, and other details of the external and internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE16D50D68DDA75BEF705C2" pageId="8" pageNumber="259" type="distribution">
<paragraph id="294A369BFFE16D50D68DDA75BEF705C2" blockId="8.[151,1436,151,1005]" pageId="8" pageNumber="259">
<emphasis id="1B81EA89FFE16D50D68DDA75BF66051F" bold="true" box="[199,353,834,860]" pageId="8" pageNumber="259">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFE16D50D72CDA74BC84051E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[358,643,835,861]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="assimile" status="sp. nov.">
<emphasis id="1B81EA89FFE16D50D72CDA74BC84051E" box="[358,643,835,861]" italics="true" pageId="8" pageNumber="259">Paraphloeostiba assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE16D50D4C2DA74BCC1051E" bold="true" box="[648,710,835,861]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="00B257F2FFE16D50D4C2DA74BCC1051E" box="[648,710,835,861]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFE16D50D598DA74BA05051E" box="[978,1026,835,861]" pageId="8" pageNumber="259">type</typeStatus>
locality in central part of
<collectingCountry id="51E2760BFFE16D50D357DA74BEEB05C2" name="Papua New Guinea" pageId="8" pageNumber="259">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE16D50D68DDABCBD3005AE" pageId="8" pageNumber="259" type="etymology">
<paragraph id="294A369BFFE16D50D68DDABCBC01058A" blockId="8.[151,1436,151,1005]" pageId="8" pageNumber="259">
<emphasis id="1B81EA89FFE16D50D68DDABCBF4905E6" bold="true" box="[199,334,907,933]" pageId="8" pageNumber="259">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFE16D50D4FCDABCBD1F05E6" box="[694,792,907,933]" italics="true" pageId="8" pageNumber="259">assimilis</emphasis>
, -
<emphasis id="1B81EA89FFE16D50D565DABCBD3C05E7" box="[815,827,907,932]" italics="true" pageId="8" pageNumber="259">e</emphasis>
(similar). It alludes to the similarity of the new species with some other relative species.
</paragraph>
<paragraph id="294A369BFFE16D50D68DDAE5BD3005AE" blockId="8.[151,1436,151,1005]" box="[199,823,978,1005]" pageId="8" pageNumber="259">
<emphasis id="1B81EA89FFE16D50D68DDAE5BF4C05AF" bold="true" box="[199,331,978,1004]" pageId="8" pageNumber="259">Bionomics.</emphasis>
The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

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The genus
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Based on the previously published data (
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) and new data, the genus
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containing 59 extant species from the Palaearctic, Nearctic, Oriental, Australian and Oceanic regions (see the map of the distribution of the genus in
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Fig.
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in
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. Three extinct species were described from the Baltic (
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) and
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(
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) amber. The morphological features of the genus can be found in
<bibRefCitation id="4D644B6AFFEF6D5ED4A5DA0FBD7B0511" author="Steel, W. O." box="[751,892,824,850]" pageId="6" pageNumber="257" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
,
<bibRefCitation id="4D644B6AFFEF6D5ED5C3DA0FBA2B0511" author="Zanetti, A." box="[905,1068,824,850]" pageId="6" pageNumber="257" pagination="49 - 117" refId="ref36237" refString="Zanetti, A. (2011) Unterfamilie Omaliinae. In: Assing, V. &amp; Schulke, M. (Eds.), Die Kafer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49 - 117." type="book chapter" year="2011">Zanetti (2011)</bibRefCitation>
,
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, etc. Members of the genus can be found in leaf litter, decaying plant remains, dung and carrion, in fermenting fruits, under the bark of trees, on flowers, etc.
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(dissected): ‘
<collectingRegion id="EB31F879FFF16D40D43EDD97BD1F02F9" box="[628,792,1184,1210]" country="Indonesia" name="Papua Barat" pageId="24" pageNumber="275">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Bilogay,
<quantity id="EE0D9B7EFFF16D40D27BDD97BAC702F9" box="[1073,1216,1184,1210]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="24" pageNumber="275" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
|
<date id="5D4B105BFFF16D40D291DD97BB5E02F8" box="[1243,1369,1184,1211]" pageId="24" pageNumber="275" value="1995-12-29">29.12.1995</date>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFF16D40D7D4DDF3BC30029D" box="[414,567,1220,1246]" pageId="24" pageNumber="275" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF16D40D419DDF3BD09029D" authorityName="Steel" authorityYear="1960" box="[595,782,1220,1246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF16D40D419DDF3BD09029D" box="[595,782,1220,1246]" italics="true" pageId="24" pageNumber="275">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF16D40D563DDF2BD6A029D" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[809,877,1221,1246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="iriana" status="sp. nov.">
<emphasis id="1B81EA89FFF16D40D563DDF2BD6A029D" box="[809,877,1221,1246]" italics="true" pageId="24" pageNumber="275">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D533DDF3BDBB029D" bold="true" box="[889,956,1220,1246]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D533DDF3BDBB029D" box="[889,956,1220,1246]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF16D40D218DDF3BA6F029D" box="[1106,1128,1220,1246]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="24" pageNumber="275" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF16D40D6D5DDDFBEEA0341" box="[159,237,1256,1282]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="275">NMW</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFF16D40D68DDC3ABD5C0309" blockId="24.[151,1437,1184,2039]" pageId="24" pageNumber="275">
<typeStatus id="F64E8839FFF16D40D68DDC3ABF3F0364" box="[199,312,1293,1319]" pageId="24" pageNumber="275" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFF16D40D709DC3BBF800364" box="[323,391,1292,1319]" count="2" pageId="24" pageNumber="275" type="male">2 ♂♂</specimenCount>
: same data as the holotype, with additional red printed label: ‘
<typeStatus id="F64E8839FFF16D40D277DC3BBACA0365" box="[1085,1229,1292,1318]" pageId="24" pageNumber="275" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF16D40D2A8DC3BBB9A0365" authorityName="Steel" authorityYear="1960" box="[1250,1437,1292,1318]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF16D40D2A8DC3BBB9A0365" box="[1250,1437,1292,1318]" italics="true" pageId="24" pageNumber="275">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF16D40D6EEDC06BEEF0309" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[164,232,1329,1354]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="iriana" status="sp. nov.">
<emphasis id="1B81EA89FFF16D40D6EEDC06BEEF0309" box="[164,232,1329,1354]" italics="true" pageId="24" pageNumber="275">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D6A5DC07BF280309" bold="true" box="[239,303,1328,1354]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D6A5DC07BF280309" box="[239,303,1328,1354]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF16D40D7F2DC07BFC90309" box="[440,462,1328,1354]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="24" pageNumber="275" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF16D40D54BDC07BD480309" box="[769,847,1328,1354]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="275">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF16D42D68DDC63BABF048E" lastPageId="26" lastPageNumber="277" pageId="24" pageNumber="275" type="description">
<paragraph id="294A369BFFF16D40D68DDC63BEDE0399" blockId="24.[151,1437,1184,2039]" pageId="24" pageNumber="275">
<emphasis id="1B81EA89FFF16D40D68DDC63BF5E032D" bold="true" box="[199,345,1364,1390]" pageId="24" pageNumber="275">Description.</emphasis>
Measurements (n=3): HW: 0.35–0.37; HL: 0.20–0.21; OL: 0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFF16D40D2ADDC63BB57032D" box="[1255,1360,1364,1390]" pageId="24" pageNumber="275" type="holotype">holotype</typeStatus>
): 0.57; PL: 0.24–0.27; PWmax: 0.41–0.47; PWmin: 0.38–0.43; ESL: 0.48–0.50; EW: 0.56–0.57; MTbL (
<typeStatus id="F64E8839FFF16D40D2ADDC4FBB5703D1" box="[1255,1360,1400,1426]" pageId="24" pageNumber="275" type="holotype">holotype</typeStatus>
): 0.27; MTrL (
<typeStatus id="F64E8839FFF16D40D6A1DCABBF5303F5" box="[235,340,1436,1462]" pageId="24" pageNumber="275" type="holotype">holotype</typeStatus>
): 0.17 (MTrL 1–4: 0.07; MTrL 5: 0.10); AW: 0.55–0.57; AedL: 0.37; BL: 1.72–1.96 (
<typeStatus id="F64E8839FFF16D40D37BDCABBB9E03F5" box="[1329,1433,1436,1462]" pageId="24" pageNumber="275" type="holotype">holotype</typeStatus>
: 1.92).
</paragraph>
<paragraph id="294A369BFFF16D40D68DDCD3BF6700B9" blockId="24.[151,1437,1184,2039]" pageId="24" pageNumber="275">
Habitus as in
<figureCitation id="B1CE2A1EFFF16D40D72FDCD3BFBE03BD" box="[357,441,1508,1534]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="24" pageNumber="275">Fig. 40</figureCitation>
. Body and antennomeres 6–11 brown, with slightly paler laterobasal and basal margins of pronotum and middle portions of each elytron; mouthparts, antennomeres 1–5 and legs yellow. Head, neck, pronotum and scutellum without visible punctation; elytra with very fine and sparse indistinct punctation, slightly denser in basal portion and invisible along suture; abdominal tergites with very fine and sparse punctation, indistinct in middle portions of abdominal tergites IV–VI. Head with dense and coarse microsculpture, transverse on vertex and isodiametric in middle; pronotum with very dense isodiametric microsculpture; microsculpture similar to that in pronotum, but slightly finer and longer; abdominal tergites with very dense, moderately coarse and isodiametric microreticulation.
</paragraph>
<paragraph id="294A369BFFF16D40D68DDE33BCBA01B4" blockId="24.[151,1437,1184,2039]" pageId="24" pageNumber="275">Head with distinct moderately deep, wide and oval anteriomedian depressions, with short anteocellar foveae, about twice as long as diameter of each ocellus; poctocular carina very indistinct. Ocelli moderately large, located slightly below level of posterior margins of eyes; distance between ocelli about 1.4 times as long as distance between ocellus and posterior margin of eyes. Antennomere 4 about as long as wide, disitnctly shorter than 3, 5 slightly shorter and indistinctly broader than 4, 6 significantly broader than 5, 7–9 indistinctly longer than 6, 10 slightly longer and indistinctly broader than 9, apical antennomere about 1.6 times as long preapical segment, from middle gradually narrowed apicad toward rounded apex.</paragraph>
<caption id="7D8A6613FFF06D41D6DDDF80BD9900B0" ID-DOI="http://doi.org/10.5281/zenodo.10821247" ID-Zenodo-Dep="10821247" httpUri="https://zenodo.org/record/10821247/files/figure.png" pageId="25" pageNumber="276" startId="25.[151,264,1719,1743]" targetBox="[159,1426,190,1687]" targetPageId="25" targetType="figure">
<paragraph id="294A369BFFF06D41D6DDDF80BD9900B0" blockId="25.[151,1437,1719,1779]" pageId="25" pageNumber="276">
<emphasis id="1B81EA89FFF06D41D6DDDF80BF5E008C" bold="true" box="[151,345,1719,1743]" pageId="25" pageNumber="276">FIGURES 64–67.</emphasis>
<taxonomicName id="EEF54D18FFF06D41D72EDF80BC50008C" authorityName="Shavrin" authorityYear="2024" box="[356,599,1719,1743]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="276" phylum="Arthropoda" rank="species" species="iriana">
<emphasis id="1B81EA89FFF06D41D72EDF80BC50008C" box="[356,599,1719,1743]" italics="true" pageId="25" pageNumber="276">Paraphloeostiba iriana</emphasis>
</taxonomicName>
(holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFF06D41D68DDE71BD5501E4" blockId="25.[151,1437,1861,2032]" pageId="25" pageNumber="276">Pronotum 1.7 times as broad as long, 1.1–1.2 times as broad as head, widest in about middle, distinctly more narrowed posteriad than anteriad; middle surface with moderately wide shallow impressions, broadened toward mediobasal margin; laterobasal impressions wide and shallow.</paragraph>
<paragraph id="294A369BFFF06D41D68DDE86BC81018F" blockId="25.[151,1437,1861,2032]" box="[199,646,1969,1996]" pageId="25" pageNumber="276">Elytra about twice as long as pronotum.</paragraph>
<paragraph id="294A369BFFF06D41D68DDEE2BC8C01AC" blockId="25.[151,1437,1861,2032]" box="[199,651,2005,2032]" pageId="25" pageNumber="276">Metatarsi 1.5 times as long as metatibia.</paragraph>
<paragraph id="294A369BFFF36D42D68DD9A0BC4A06F2" blockId="26.[151,1437,151,933]" box="[199,589,151,177]" pageId="26" pageNumber="277">Abdomen about as broad as elytra.</paragraph>
<paragraph id="294A369BFFF36D42D68DD98CBB370726" blockId="26.[151,1437,151,933]" pageId="26" pageNumber="277">
Male. Posterior margin of abdominal tergite VIII indistinctly sinuate (
<figureCitation id="B1CE2A1EFFF36D42D591D98CBA370696" box="[987,1072,187,213]" captionStart="FIGURES 64–67" captionStartId="25.[151,264,1719,1743]" captionTargetBox="[159,1426,190,1687]" captionTargetId="figure-16@25.[151,1436,181,1695]" captionTargetPageId="25" captionText="FIGURES 64–67. Paraphloeostiba iriana (holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821247" httpUri="https://zenodo.org/record/10821247/files/figure.png" pageId="26" pageNumber="277">Fig. 66</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFF36D42D425D9E8BCC206BA" box="[623,709,223,249]" captionStart="FIGURES 64–67" captionStartId="25.[151,264,1719,1743]" captionTargetBox="[159,1426,190,1687]" captionTargetId="figure-16@25.[151,1436,181,1695]" captionTargetPageId="25" captionText="FIGURES 64–67. Paraphloeostiba iriana (holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821247" httpUri="https://zenodo.org/record/10821247/files/figure.png" pageId="26" pageNumber="277">Fig. 67</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward rounded apex; parameres slightly exceeding apex of median lobe, moderately narrow, with two long apical and numerous long setae along inner margin of each paramere; internal sac wide and moderately long, with several short and elongate sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFF36D42D494D87CBD340726" box="[734,819,331,357]" captionStart="FIGURES 64–67" captionStartId="25.[151,264,1719,1743]" captionTargetBox="[159,1426,190,1687]" captionTargetId="figure-16@25.[151,1436,181,1695]" captionTargetPageId="25" captionText="FIGURES 64–67. Paraphloeostiba iriana (holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821247" httpUri="https://zenodo.org/record/10821247/files/figure.png" pageId="26" pageNumber="277">Fig. 64</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFF36D42D293D87CBB2B0726" box="[1241,1324,330,357]" captionStart="FIGURES 64–67" captionStartId="25.[151,264,1719,1743]" captionTargetBox="[159,1426,190,1687]" captionTargetId="figure-16@25.[151,1436,181,1695]" captionTargetPageId="25" captionText="FIGURES 64–67. Paraphloeostiba iriana (holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821247" httpUri="https://zenodo.org/record/10821247/files/figure.png" pageId="26" pageNumber="277">Fig. 65</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFF36D42D68DD858BF9607CA" blockId="26.[151,1437,151,933]" box="[199,401,367,393]" pageId="26" pageNumber="277">Female unknown.</paragraph>
<paragraph id="294A369BFFF36D42D68DD8A5BABF048E" blockId="26.[151,1437,151,933]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D68DD8A5BFBF07EE" bold="true" box="[199,440,402,429]" pageId="26" pageNumber="277">Comparative notes.</emphasis>
Based on the general structure of the aedeagus,
<taxonomicName id="EEF54D18FFF36D42D25CD8A4BA8707EF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1046,1152,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFF36D42D25CD8A4BA2007EF" box="[1046,1063,403,428]" italics="true" pageId="26" pageNumber="277">P</emphasis>
.
<emphasis id="1B81EA89FFF36D42D276D8A4BA8707EF" box="[1084,1152,403,428]" italics="true" pageId="26" pageNumber="277">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D2C5D8A4BAD207EE" bold="true" box="[1167,1237,403,429]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D2C5D8A4BAD207EE" box="[1167,1237,403,429]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFF36D42D3CED8A4BF5C0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFF36D42D3CED8A4BB9207EF" box="[1412,1429,403,428]" italics="true" pageId="26" pageNumber="277">P</emphasis>
.
<emphasis id="1B81EA89FFF36D42D6DDD880BF5C0792" box="[151,347,439,465]" italics="true" pageId="26" pageNumber="277">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D72CD880BFAD0792" bold="true" box="[358,426,439,465]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D72CD880BFAD0792" box="[358,426,439,465]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFF36D42D7F1D880BC230792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[443,548,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFF36D42D7F1D880BC230792" box="[443,548,439,465]" italics="true" pageId="26" pageNumber="277">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D465D880BC750792" bold="true" box="[559,626,439,465]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D465D880BC750792" box="[559,626,439,465]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFF36D42D4FAD880BD3A0793" authorityName="Shavrin" authorityYear="2023" box="[688,829,439,464]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF36D42D4FAD880BD3A0793" box="[688,829,439,464]" italics="true" pageId="26" pageNumber="277">P. conjuncta</emphasis>
</taxonomicName>
. Based on the presence of two distinct longitudinal impressions on the pronotum, it is more similar to
<taxonomicName id="EEF54D18FFF36D42D48CD8ECBDAF07B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[710,936,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFF36D42D48CD8ECBCD007B7" box="[710,727,475,500]" italics="true" pageId="26" pageNumber="277">P</emphasis>
.
<emphasis id="1B81EA89FFF36D42D4AED8ECBDAF07B6" box="[740,936,475,501]" italics="true" pageId="26" pageNumber="277">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D5E5D8ECBDEA07B6" bold="true" box="[943,1005,475,501]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D5E5D8ECBDEA07B6" box="[943,1005,475,501]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the smaller body, slightly longer parameres with distinctly narrower apical portions. It can be distinguished from
<taxonomicName id="EEF54D18FFF36D42D3C2D8C8BF02047F" authorityName="Shavrin" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF36D42D3C2D8C8BF02047F" italics="true" pageId="26" pageNumber="277">P. conjuncta</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF54D18FFF36D42D774DB14BFA4047E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[318,419,547,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFF36D42D774DB14BFA4047E" box="[318,419,547,573]" italics="true" pageId="26" pageNumber="277">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D7E1DB14BFEC047E" bold="true" box="[427,491,547,573]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D7E1DB14BFEC047E" box="[427,491,547,573]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the broader apical portion of the median lobe. From both these species it can be distinguished by the slightly longer parameres. Additionally, it can be distinguished from
<taxonomicName id="EEF54D18FFF36D42D230DB70BB040423" authorityName="Shavrin" authorityYear="2023" box="[1146,1283,583,608]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF36D42D230DB70BB040423" box="[1146,1283,583,608]" italics="true" pageId="26" pageNumber="277">P. conjuncta</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF54D18FFF36D42D372DB70BB9B0422" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1336,1436,583,609]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFF36D42D372DB70BB9B0422" box="[1336,1436,583,609]" italics="true" pageId="26" pageNumber="277">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D6DDDB5CBED004C6" bold="true" box="[151,215,619,645]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D6DDDB5CBED004C6" box="[151,215,619,645]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the finer punctation of the head, the presence of longitudinal impressions in the middle of the pronotum, and details of the morphology of the aedeagus. From all these species it differs by the details of the external and internal morphology of the aedeagus, and the different shape of the female accesssory sclerite.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D42D68DDBE1BFA10556" pageId="26" pageNumber="277" type="distribution">
<paragraph id="294A369BFFF36D42D68DDBE1BFA10556" blockId="26.[151,1437,151,933]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D68DDBE1BF6604B3" bold="true" box="[199,353,726,752]" pageId="26" pageNumber="277">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFF36D42D72DDBE0BC6B04B3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[359,620,727,753]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="iriana" status="sp. nov.">
<emphasis id="1B81EA89FFF36D42D72DDBE0BC6B04B3" box="[359,620,727,753]" italics="true" pageId="26" pageNumber="277">Paraphloeostiba iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D439DBE0BCB504B2" bold="true" box="[627,690,727,753]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D439DBE0BCB504B2" box="[627,690,727,753]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFF36D42D58BDBE0BDF604B2" box="[961,1009,727,753]" pageId="26" pageNumber="277">type</typeStatus>
locality in Paniai Province of Western New
<collectingCountry id="51E2760BFFF36D42D699DBCCBF200556" box="[211,295,763,789]" name="Guinea-Bissau" pageId="26" pageNumber="277">Guinea</collectingCountry>
,
<collectingCountry id="51E2760BFFF36D42D778DBCCBFA40556" box="[306,419,763,789]" name="Indonesia" pageId="26" pageNumber="277">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D42D68DDA28BF3705E6" pageId="26" pageNumber="277" type="etymology">
<paragraph id="294A369BFFF36D42D68DDA28BD31051E" blockId="26.[151,1437,151,933]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D68DDA28BF49057A" bold="true" box="[199,334,799,825]" pageId="26" pageNumber="277">Etymology.</emphasis>
The specific epithet is the Latinized adjective derived from the name of the province of Irian Barat before being renamed
<collectingRegion id="EB31F879FFF36D42D7D8DA74BC07051E" box="[402,512,835,861]" country="Indonesia" name="Papua Barat" pageId="26" pageNumber="277">Irian Jaya</collectingRegion>
(literally “Glorious Irian”).
</paragraph>
<paragraph id="294A369BFFF36D42D68DDA51BF3705E6" blockId="26.[151,1437,151,933]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D68DDA51BF4C05C3" bold="true" box="[199,331,870,896]" pageId="26" pageNumber="277">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="EE0D9B7EFFF36D42D512DA50BA0A05C2" box="[856,1037,871,897]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="26" pageNumber="277" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

293
data/A1/5C/87/A15C878DFFF36D45D6DDDAC1BA550703.xml Normal file
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@ -0,0 +1,293 @@
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<mods:title id="34BB81A7FCC014AFBC39FDA2450E2684">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<treatment id="A15C878DFFF36D45D6DDDAC1BA550703" ID-DOI="http://doi.org/10.5281/zenodo.10821279" ID-Zenodo-Dep="10821279" LSID="urn:lsid:plazi:treatment:A15C878DFFF36D45D6DDDAC1BA550703" httpUri="http://treatment.plazi.org/id/A15C878DFFF36D45D6DDDAC1BA550703" lastPageId="29" lastPageNumber="280" pageId="26" pageNumber="277">
<subSubSection id="61EF6510FFF36D42D6DDDAC1BC1E0252" box="[151,537,1014,1041]" pageId="26" pageNumber="277" type="nomenclature">
<paragraph id="294A369BFFF36D42D6DDDAC1BC1E0252" blockId="26.[151,537,1014,1077]" box="[151,537,1014,1041]" pageId="26" pageNumber="277">
<heading id="720281F7FFF36D42D6DDDAC1BC1E0252" bold="true" box="[151,537,1014,1041]" fontSize="11" level="1" pageId="26" pageNumber="277" reason="1">
<emphasis id="1B81EA89FFF36D42D6DDDAC1BC1E0252" bold="true" box="[151,537,1014,1041]" pageId="26" pageNumber="277">
<taxonomicName id="EEF54D18FFF36D42D6DDDAC1BFD70252" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,464,1014,1041]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF36D42D6DDDAC1BFD70252" bold="true" box="[151,464,1014,1041]" italics="true" pageId="26" pageNumber="277">Paraphloeostiba latissima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFF36D42D79DDAC0BC1E0252" box="[471,537,1015,1041]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D42D6DDDD2CBF560276" box="[151,337,1051,1077]" pageId="26" pageNumber="277" type="description">
<paragraph id="294A369BFFF36D42D6DDDD2CBF560276" blockId="26.[151,537,1014,1077]" box="[151,337,1051,1077]" pageId="26" pageNumber="277">
(
<figureCitation id="B1CE2A1EFFF36D42D6D5DD2CBEF10276" box="[159,246,1051,1077]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="26" pageNumber="277">Figs 41</figureCitation>
,
<figureCitation id="B1CE2A1EFFF36D42D748DD2CBF4E0276" box="[258,329,1051,1077]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="26" pageNumber="277">68–74</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D42D6DDDD54BB1B033A" pageId="26" pageNumber="277" type="materials_examined">
<paragraph id="294A369BFFF36D42D6DDDD54BFAA0286" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D6DDDD54BF42023F" bold="true" box="[151,325,1122,1149]" pageId="26" pageNumber="277">Type material.</emphasis>
<typeStatus id="F64E8839FFF36D42D707DD54BFB2023E" box="[333,437,1123,1149]" pageId="26" pageNumber="277" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFF36D42D7F6DD54BFD6023E" box="[444,465,1123,1149]" count="1" pageId="26" pageNumber="277" type="male">♂</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFF36D42D42CDD54BCAD023E" box="[614,682,1123,1149]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="26" pageNumber="277">NEW</collectionCode>
<collectingCountry id="51E2760BFFF36D42D4FBDD54BD1C023E" box="[689,795,1123,1149]" name="Guinea-Bissau" pageId="26" pageNumber="277">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFF36D42D5CDDD54BA12023E" box="[903,1045,1122,1149]" pageId="26" pageNumber="277" value="1965-06-16">16. VI. 1965</date>
| flying in daylight’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF36D42D335DD54BB9F023E" box="[1407,1432,1123,1149]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="277">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFF36D42D428DDB0BCFC02E2" box="[610,763,1159,1185]" pageId="26" pageNumber="277" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF36D42D547DDB0BDCF02E2" authorityName="Steel" authorityYear="1960" box="[781,968,1159,1185]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF36D42D547DDB0BDCF02E2" box="[781,968,1159,1185]" italics="true" pageId="26" pageNumber="277">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF36D42D590DDB0BA3802E2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[986,1087,1159,1185]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF36D42D590DDB0BA3802E2" box="[986,1087,1159,1185]" italics="true" pageId="26" pageNumber="277">latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D20FDDB0BA8302E2" bold="true" box="[1093,1156,1159,1185]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D20FDDB0BA8302E2" box="[1093,1156,1159,1185]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF36D42D341DDB0BB2602E2" box="[1291,1313,1159,1185]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="277" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF36D42D70DDD9CBFA60286" box="[327,417,1195,1221]" pageId="26" pageNumber="277">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFF36D42D68DDDF8BB1B033A" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<typeStatus id="F64E8839FFF36D42D68DDDF8BF3F02AA" box="[199,312,1231,1257]" pageId="26" pageNumber="277" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFF36D42D70CDDF8BF7402AA" box="[326,371,1231,1257]" count="1" pageId="26" pageNumber="277" type="male">1 ♂</specimenCount>
(underside of the card with handwritten ‘Okapo 1164’),
<specimenCount id="3FF3FD12FFF36D42D241DDF8BA4A02AA" box="[1035,1101,1231,1257]" count="2" pageId="26" pageNumber="277" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFF36D42D215DDF8BAF902AA" box="[1119,1278,1231,1257]" count="1" pageId="26" pageNumber="277" type="generic">one specimen</specimenCount>
without right antennomeres 3–11 dissected;
<specimenCount id="3FF3FD12FFF36D42D7A6DDC4BC82034E" box="[492,645,1267,1293]" count="1" pageId="26" pageNumber="277" type="generic">one specimen</specimenCount>
with underside of the card with handwritten ‘Okapo 1164’): same data as the holotype (
<collectionCode id="4FE4AE5EFFF36D42D719DC20BFAA0372" box="[339,429,1303,1329]" pageId="26" pageNumber="277">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFF36D42D78BDC20BFEA0372" box="[449,493,1303,1329]" count="1" pageId="26" pageNumber="277" type="male">1 ♂</specimenCount>
(dissected),
<specimenCount id="3FF3FD12FFF36D42D437DC20BCAF0372" box="[637,680,1303,1329]" count="1" pageId="26" pageNumber="277" type="female">1 ♀</specimenCount>
(underside of the card with handwritten ‘90H’): ‘
<collectionCode id="4FE4AE5EFFF36D42D292DC20BB1B0372" box="[1240,1308,1303,1329]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="26" pageNumber="277">NEW</collectionCode>
<collectingCountry id="51E2760BFFF36D42D36FDC20BB880372" box="[1317,1423,1303,1329]" name="Guinea-Bissau" pageId="26" pageNumber="277">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFF36D42D6A5DC0DBF870316" box="[239,384,1338,1365]" pageId="26" pageNumber="277" value="1965-09-25">25. IX. 1965</date>
, | refuse’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF36D42D424DC0CBC800316" box="[622,647,1339,1365]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="277">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFF36D42D22BDC0CBABC0316" box="[1121,1211,1339,1365]" pageId="26" pageNumber="277">BMNH</collectionCode>
). All
<typeStatus id="F64E8839FFF36D42D2B2DC0CBB640316" box="[1272,1379,1339,1365]" pageId="26" pageNumber="277" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFF36D42D792DC68BC6F033A" box="[472,616,1375,1401]" pageId="26" pageNumber="277" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF36D42D436DC68BD30033A" authorityName="Steel" authorityYear="1960" box="[636,823,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF36D42D436DC68BD30033A" box="[636,823,1375,1401]" italics="true" pageId="26" pageNumber="277">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF36D42D501DC68BDB7033A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[843,944,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF36D42D501DC68BDB7033A" box="[843,944,1375,1401]" italics="true" pageId="26" pageNumber="277">latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D5FDDC68BDF1033A" bold="true" box="[951,1014,1375,1401]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D5FDDC68BDF1033A" box="[951,1014,1375,1401]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF36D42D2CADC68BA91033A" box="[1152,1174,1375,1401]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="277" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D44D68DDCB5BDD701FC" lastPageId="28" lastPageNumber="279" pageId="26" pageNumber="277" type="description">
<paragraph id="294A369BFFF36D42D68DDCB5BEDE004A" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D68DDCB5BF5E03DF" bold="true" box="[199,345,1410,1436]" pageId="26" pageNumber="277">Description.</emphasis>
Measurements (n=6): HW: 0.37–0.43; HL: 0.25–0.26;OL: 0.12–0.14; TL: 0.02; AL (
<typeStatus id="F64E8839FFF36D42D362DCB4BB9603DE" box="[1320,1425,1411,1437]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
): 0.60; PL: 0.30–0.36; PWmax: 0.46–0.53; PWmin: 0.42–0.48; ESL: 0.47–0.53; EW: 0.60–0.62; MTbL (
<typeStatus id="F64E8839FFF36D42D362DC90BB960382" box="[1320,1425,1447,1473]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFF36D42D76FDCFCBF8903A6" box="[293,398,1483,1509]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
): 0.16 (MTrL 1–4: 0.05; MTrL 5: 0.11); AW: 0.57–0.60; AedL: 0.30; BL: 1.67–2.10 (
<typeStatus id="F64E8839FFF36D42D37BDCFCBB9E03A6" box="[1329,1433,1483,1509]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
: 1.82).
</paragraph>
<paragraph id="294A369BFFF36D42D68DDF24BA3C0132" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<materialsCitation id="999D3CC6FFF36D42D68DDF24BA300132" collectionCode="V" pageId="26" pageNumber="277" specimenCount="9" typeStatus="holotype">
Habitus as in
<figureCitation id="B1CE2A1EFFF36D42D72FDF24BFBD006E" box="[357,442,1555,1581]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="26" pageNumber="277">Fig. 41</figureCitation>
. Body reddish brown, with yellowish to yellow-brown elytra, with darker medioapical, laterobasal and basal portions (laterobasal portions of pronotum of
<typeStatus id="F64E8839FFF36D42D5CBDF00BDE40012" box="[897,995,1591,1617]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
and
<specimenCount id="3FF3FD12FFF36D42D252DF00BAB30012" box="[1048,1204,1591,1617]" count="2" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">two paratypes</specimenCount>
yellowish; abdomen slightly paler in
<specimenCount id="3FF3FD12FFF36D42D717DF6CBC150036" box="[349,530,1627,1653]" count="3" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">three paratypes</specimenCount>
); antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow. Head with fine punctation, denser in middle and sparser on infraorbital portions (
<specimenCount id="3FF3FD12FFF36D42D262DF48BABD00DA" box="[1064,1210,1663,1689]" count="1" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">one paratype</specimenCount>
with very indistinct punctation); neck without visible punctures; pronotum with fine and moderately dense punctation, similar to that in middle portion of head, finer and sparser in middle; elytra with fine punctation, slightly denser and deeper than that in pronotum, finer around scutellum and along suture; abdominal tergites with very fine and moderately dense punctation, indistinct in abdominal tergites IV–
<collectionCode id="4FE4AE5EFFF36D42D4FBDE38BCC2016A" box="[689,709,1807,1833]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="277" type="Museum">V</collectionCode>
in
<specimenCount id="3FF3FD12FFF36D42D4A1DE38BD9D016A" box="[747,922,1807,1833]" count="3" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">three paratypes</specimenCount>
. Head with dense and coarse microsculpture, transverse on vertex, coarser and distinctly isodiametric in middle, isodiametric and elongate on infraorbital portions; microsculpture of elytra very dense, about as that in pronotum, but slightly coarser
</materialsCitation>
.
</paragraph>
<paragraph id="294A369BFFF36D43D68DDE4CBA1E06BA" blockId="26.[151,1437,1122,2013]" lastBlockId="27.[151,1437,151,609]" lastPageId="27" lastPageNumber="278" pageId="26" pageNumber="277">Head with slightly elevated middle and infraorbital portions, with deep and moderately wide anteriomedian depressions, 1.4–1.5 times as broad as head, with narrow nad deep linear anteocellar foveae, almost reaching level of middle length of eyes; postocular carina absent. Ocelli moderately large, located at level of posterior margins of eyes; distance between ocelli about as long as distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about 1.4 times as long as preceding segment. Antennomere 6 slightly longer and significantly broader than 5, 7–9 slightly longer and broader than 6, 10 slightly longer than 9.</paragraph>
<paragraph id="294A369BFFF26D43D68DD834BD140702" blockId="27.[151,1437,151,609]" pageId="27" pageNumber="278">Pronotum 1.4–1.5 times as broad as long, 1.2 times as broad as head, widest in or slightly above middle, strongly narrowed posteriad toward subacute hing angle.</paragraph>
<paragraph id="294A369BFFF26D43D68DD87CBDFA0726" blockId="27.[151,1437,151,609]" box="[199,1021,330,357]" pageId="27" pageNumber="278">Elytra 1.1–1.2 times as broad as long, 1.4–1.5 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFF26D43D68DD858BC8507CA" blockId="27.[151,1437,151,609]" box="[199,642,367,393]" pageId="27" pageNumber="278">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="294A369BFFF26D43D68DD8A4BDB00422" blockId="27.[151,1437,151,609]" pageId="27" pageNumber="278">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFF26D43D51ED8A4BDA007EE" box="[852,935,403,429]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 70</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFF26D43D794D880BC340792" box="[478,563,439,465]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 71</figureCitation>
). Aedeagus with wide median lobe, indistinctly narrowed to widely rounded apex; parameres moderately short, significantly not reaching apex of median lobe, with narrow apical portion, with two long apical and two to three preapical setae, and two small setae in middle part of inner margin of each paramere; internal sac wide and moderately short, with two narrow and moderately long sclerotized structures in preapical portion (
<figureCitation id="B1CE2A1EFFF26D43D72FDB70BFBE0422" box="[357,441,583,609]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 68</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFF26D43D52ADB70BDB40422" box="[864,947,583,609]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 69</figureCitation>
.
</paragraph>
<caption id="7D8A6613FFF26D43D6DDDE4EBC80019A" ID-DOI="http://doi.org/10.5281/zenodo.10821251" ID-Zenodo-Dep="10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278" startId="27.[151,264,1913,1937]" targetBox="[172,1428,642,1882]" targetPageId="27" targetType="figure">
<paragraph id="294A369BFFF26D43D6DDDE4EBC80019A" blockId="27.[151,1436,1913,2010]" pageId="27" pageNumber="278">
<emphasis id="1B81EA89FFF26D43D6DDDE4EBF5101D2" bold="true" box="[151,342,1913,1937]" pageId="27" pageNumber="278">FIGURES 68–74.</emphasis>
<taxonomicName id="EEF54D18FFF26D43D717DE4DBC6D01D1" authorityName="Shavrin" authorityYear="2024" box="[349,618,1914,1938]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="latissima">
<emphasis id="1B81EA89FFF26D43D717DE4DBC6D01D1" box="[349,618,1914,1938]" italics="true" pageId="27" pageNumber="278">Paraphloeostiba latissima</emphasis>
</taxonomicName>
(paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="7D8A6613FFF56D44D6DDDF09BA920039" ID-DOI="http://doi.org/10.5281/zenodo.10821253" ID-Zenodo-Dep="10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="28" pageNumber="279" startId="28.[151,264,1598,1622]" targetBox="[212,1375,188,1560]" targetPageId="28" targetType="figure">
<paragraph id="294A369BFFF56D44D6DDDF09BA920039" blockId="28.[151,1436,1598,1659]" pageId="28" pageNumber="279">
<emphasis id="1B81EA89FFF56D44D6DDDF09BF5C0015" bold="true" box="[151,347,1598,1622]" pageId="28" pageNumber="279">FIGURES 75–80.</emphasis>
Habitus of
<taxonomicName id="EEF54D18FFF56D44D7ACDF09BC880015" authorityName="Steel" authorityYear="1960" box="[486,655,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF56D44D7ACDF09BC880015" box="[486,655,1598,1622]" italics="true" pageId="28" pageNumber="279">Paraphloeostiba</emphasis>
</taxonomicName>
: 75–
<taxonomicName id="EEF54D18FFF56D44D480DF08BD4A0015" authorityName="Shavrin" authorityYear="2024" box="[714,845,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="okapensis">
<emphasis id="1B81EA89FFF56D44D480DF08BD4A0015" box="[714,845,1598,1622]" italics="true" pageId="28" pageNumber="279">P. okapensis</emphasis>
</taxonomicName>
(paratype), 76–
<taxonomicName id="EEF54D18FFF56D44D5B6DF08BA880015" authorityName="Shavrin" authorityYear="2024" box="[1020,1167,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="penelopeae">
<emphasis id="1B81EA89FFF56D44D5B6DF08BA880015" box="[1020,1167,1598,1622]" italics="true" pageId="28" pageNumber="279">P. penelopeae</emphasis>
</taxonomicName>
(paratype), 77–
<taxonomicName id="EEF54D18FFF56D44D374DF08BB9B0015" authorityName="Shavrin" authorityYear="2024" box="[1342,1436,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="1B81EA89FFF56D44D374DF08BB9B0015" box="[1342,1436,1598,1622]" italics="true" pageId="28" pageNumber="279">P. pilosa</emphasis>
</taxonomicName>
(holotype), 78–
<taxonomicName id="EEF54D18FFF56D44D779DF54BF890039" authorityName="Shavrin" authorityYear="2024" box="[307,398,1634,1658]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="1B81EA89FFF56D44D779DF54BF890039" box="[307,398,1634,1658]" italics="true" pageId="28" pageNumber="279">P. riedeli</emphasis>
</taxonomicName>
(paratype), 79–
<taxonomicName id="EEF54D18FFF56D44D465DF54BCC20039" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[559,709,1634,1658]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFF56D44D465DF54BCC20039" box="[559,709,1634,1658]" italics="true" pageId="28" pageNumber="279">P. rhopalocera</emphasis>
</taxonomicName>
, 80–
<taxonomicName id="EEF54D18FFF56D44D4B2DF54BD480039" authorityName="Shavrin" authorityYear="2024" box="[760,847,1634,1658]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="rufula">
<emphasis id="1B81EA89FFF56D44D4B2DF54BD480039" box="[760,847,1634,1658]" italics="true" pageId="28" pageNumber="279">P. rufula</emphasis>
</taxonomicName>
(holotype). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFF56D44D68DDFFABCCB0148" blockId="28.[151,1437,1741,1983]" pageId="28" pageNumber="279">
Female. Posterior margin of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFF56D44D56EDFFABD7F00A4" box="[804,888,1741,1767]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="28" pageNumber="279">Fig. 72</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFF56D44D213DFFABAAA00A4" box="[1113,1197,1741,1767]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="28" pageNumber="279">Fig. 73</figureCitation>
) truncate. Accessory sclerite as in
<figureCitation id="B1CE2A1EFFF56D44D760DFC6BF7A0148" box="[298,381,1777,1803]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="28" pageNumber="279">Fig. 74</figureCitation>
. Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFF56D44D68DDE23BDD701FC" blockId="28.[151,1437,1741,1983]" pageId="28" pageNumber="279">
<emphasis id="1B81EA89FFF56D44D68DDE23BFB7016C" bold="true" box="[199,432,1812,1839]" pageId="28" pageNumber="279">Comparative notes.</emphasis>
Based on the presence of longitudinal impressions on the pronotum and the general shape of the aedeagus with narrow and short parameres,
<taxonomicName id="EEF54D18FFF56D44D4AADE0EBD640110" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[736,867,1849,1875]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="latissima" status="sp. n.">
<emphasis id="1B81EA89FFF56D44D4AADE0EBD640110" box="[736,867,1849,1875]" italics="true" pageId="28" pageNumber="279">P. latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF56D44D527DE0EBDA80110" bold="true" box="[877,943,1849,1875]" pageId="28" pageNumber="279">
<taxonomicNameLabel id="00B257F2FFF56D44D527DE0EBDA80110" box="[877,943,1849,1875]" pageId="28" pageNumber="279" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is somewhat similar to
<taxonomicName id="EEF54D18FFF56D44D28DDE0EBED50134" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="cooteri">
<emphasis id="1B81EA89FFF56D44D28DDE0EBB330111" box="[1223,1332,1849,1874]" italics="true" pageId="28" pageNumber="279">P. cooteri</emphasis>
Shavrin, 2020
</taxonomicName>
, recently described from
<collectingRegion id="EB31F879FFF56D44D7A6DE6ABC540134" box="[492,595,1885,1911]" country="China" name="Zhejiang" pageId="28" pageNumber="279">Zhejiang</collectingRegion>
,
<collectingCountry id="51E2760BFFF56D44D417DE6ABCA70134" box="[605,672,1885,1911]" name="China" pageId="28" pageNumber="279">China</collectingCountry>
(Shavrin 2020). The new species can be distinguished from it by the less transverse pronotum, finer punctation of the head and the pronotum, broader apical portion of the median lobe, narrower and shorter parameres and different structure of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF46D45D68DD9A1BF2B0697" pageId="29" pageNumber="280" type="distribution">
<paragraph id="294A369BFFF46D45D68DD9A1BF2B0697" blockId="29.[151,1436,150,321]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D68DD9A1BF6606F3" bold="true" box="[199,353,150,176]" pageId="29" pageNumber="280">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFF46D45D723D9A1BC9506F3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[361,658,150,176]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF46D45D723D9A1BC9506F3" box="[361,658,150,176]" italics="true" pageId="29" pageNumber="280">Paraphloeostiba latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF46D45D4D1D9A1BCDB06F3" bold="true" box="[667,732,150,176]" pageId="29" pageNumber="280">
<taxonomicNameLabel id="00B257F2FFF46D45D4D1D9A1BCDB06F3" box="[667,732,150,176]" pageId="29" pageNumber="280" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFF46D45D5B0D9A0BA2D06F2" box="[1018,1066,151,177]" pageId="29" pageNumber="280">type</typeStatus>
locality in central part of
<collectingCountry id="51E2760BFFF46D45D312D9A0BF2F0697" name="Papua New Guinea" pageId="29" pageNumber="280">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF46D45D68DD9E9BA550703" pageId="29" pageNumber="280" type="etymology">
<paragraph id="294A369BFFF46D45D68DD9E9BF4A075F" blockId="29.[151,1436,150,321]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D68DD9E9BF4906BB" bold="true" box="[199,334,222,248]" pageId="29" pageNumber="280">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFF46D45D489D9E9BD3406BB" box="[707,819,222,248]" italics="true" pageId="29" pageNumber="280">latissimus</emphasis>
, -
<emphasis id="1B81EA89FFF46D45D501D9E8BD9406BB" box="[843,915,223,248]" italics="true" pageId="29" pageNumber="280">a, -um</emphasis>
(broadest). It alludes to the wide median lobe of the aedeagus.
</paragraph>
<paragraph id="294A369BFFF46D45D68DD811BA550703" blockId="29.[151,1436,150,321]" box="[199,1106,294,321]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D68DD811BF4C0703" bold="true" box="[199,331,294,320]" pageId="29" pageNumber="280">Bionomics.</emphasis>
Specimens were collected flying in daylight or were found in refuse.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="5EB06DF04AD82CB5695D70191EFCE965" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFF46D46D6DDD8A5BF3703A6" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba margineguttata Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="281" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="280" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7F5E875EB5BD4F990D2DA2EA035220AD">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<mods:namePart id="A684E0CE8599840F214DBBEB574A2F4A">Shavrin, Alexey V.</mods:namePart>
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<subSubSection id="61EF6510FFF46D45D6DDD8A5BC6007EF" box="[151,615,402,429]" pageId="29" pageNumber="280" type="nomenclature">
<paragraph id="294A369BFFF46D45D6DDD8A5BC6007EF" blockId="29.[151,615,402,464]" box="[151,615,402,429]" pageId="29" pageNumber="280">
<heading id="720281F7FFF46D45D6DDD8A5BC6007EF" bold="true" box="[151,615,402,429]" fontSize="11" level="1" pageId="29" pageNumber="280" reason="1">
<emphasis id="1B81EA89FFF46D45D6DDD8A5BC6007EF" bold="true" box="[151,615,402,429]" pageId="29" pageNumber="280">
<taxonomicName id="EEF54D18FFF46D45D6DDD8A5BC1907EF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,542,402,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="margineguttata" status="sp. nov.">
<emphasis id="1B81EA89FFF46D45D6DDD8A5BC1907EF" bold="true" box="[151,542,402,429]" italics="true" pageId="29" pageNumber="280">Paraphloeostiba margineguttata</emphasis>
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<taxonomicNameLabel id="00B257F2FFF46D45D46FD8A5BC6007EF" box="[549,615,402,428]" pageId="29" pageNumber="280" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="61EF6510FFF46D45D6DDD881BF560793" box="[151,337,438,464]" pageId="29" pageNumber="280" type="description">
<paragraph id="294A369BFFF46D45D6DDD881BF560793" blockId="29.[151,615,402,464]" box="[151,337,438,464]" pageId="29" pageNumber="280">
(
<figureCitation id="B1CE2A1EFFF46D45D6D5D881BEF10793" box="[159,246,438,464]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="29" pageNumber="280">Figs 43</figureCitation>
,
<figureCitation id="B1CE2A1EFFF46D45D748D881BF4E0793" box="[258,329,438,464]" captionStart="FIGURES 81–84" captionStartId="29.[151,264,1649,1673]" captionTargetBox="[168,1428,710,1617]" captionTargetId="figure-145@29.[151,1436,704,1625]" captionTargetPageId="29" captionText="FIGURES 81–84. Paraphloeostiba margineguttata (holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="29" pageNumber="280">81–84</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF46D45D6DDD8C9BDB104EB" pageId="29" pageNumber="280" type="materials_examined">
<paragraph id="294A369BFFF46D45D6DDD8C9BFE00423" blockId="29.[151,1437,510,680]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D6DDD8C9BF41045B" bold="true" box="[151,326,510,536]" pageId="29" pageNumber="280">Type material.</emphasis>
<typeStatus id="F64E8839FFF46D45D704D8C9BFB1045B" box="[334,438,510,536]" pageId="29" pageNumber="280" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFF46D45D7F5D8C8BFD3045A" box="[447,468,511,537]" count="1" pageId="29" pageNumber="280" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFF46D45D7AED8C9BC3E045B" box="[484,569,510,536]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="29" pageNumber="280">Fig. 43</figureCitation>
; dissected): ‘
<collectingRegion id="EB31F879FFF46D45D480D8C9BD6B045B" box="[714,876,510,536]" country="Indonesia" name="Papua Barat" pageId="29" pageNumber="280">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Sinak, trail to Ilaga |
<quantity id="EE0D9B7EFFF46D45D34CD8C9BB9F045A" box="[1286,1432,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="29" pageNumber="280" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100-2500m</quantity>
,
<date id="5D4B105BFFF46D45D6DDDB15BF25047E" box="[151,290,546,573]" pageId="29" pageNumber="280" value="1995-12-15">15. 12. 1995</date>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFF46D45D417DB15BCF1047F" box="[605,758,546,572]" pageId="29" pageNumber="280" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF46D45D543DB15BDC3047F" authorityName="Steel" authorityYear="1960" box="[777,964,546,572]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF46D45D543DB15BDC3047F" box="[777,964,546,572]" italics="true" pageId="29" pageNumber="280">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF46D45D59CDB14BA85047F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[982,1154,547,572]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="margineguttata" status="sp. nov.">
<emphasis id="1B81EA89FFF46D45D59CDB14BA85047F" box="[982,1154,547,572]" italics="true" pageId="29" pageNumber="280">margineguttata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF46D45D2C3DB15BACF047F" bold="true" box="[1161,1224,546,572]" pageId="29" pageNumber="280">
<taxonomicNameLabel id="00B257F2FFF46D45D2C3DB15BACF047F" box="[1161,1224,546,572]" pageId="29" pageNumber="280" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF46D45D31ADB15BB61047F" box="[1360,1382,546,572]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="29" pageNumber="280" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF46D45D7C7DB71BFDC0423" box="[397,475,582,608]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="29" pageNumber="280">NMW</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFF46D45D68DDB5CBDB104EB" blockId="29.[151,1437,510,680]" pageId="29" pageNumber="280">
<typeStatus id="F64E8839FFF46D45D68DDB5CBF2E04C6" box="[199,297,619,645]" pageId="29" pageNumber="280" type="paratype">Paratype</typeStatus>
<specimenCount id="3FF3FD12FFF46D45D77EDB5CBF4B04C6" box="[308,332,619,645]" count="1" pageId="29" pageNumber="280" type="male">♂</specimenCount>
: same data as the holotype, with additional red printed label: ‘
<typeStatus id="F64E8839FFF46D45D26CDB5DBAB104C7" box="[1062,1206,618,644]" pageId="29" pageNumber="280" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF46D45D29BDB5DBB8B04C7" authorityName="Steel" authorityYear="1960" box="[1233,1420,618,644]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF46D45D29BDB5DBB8B04C7" box="[1233,1420,618,644]" italics="true" pageId="29" pageNumber="280">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF46D45D6DDDBB8BF4404EB" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,323,655,680]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="margineguttata" status="sp. nov.">
<emphasis id="1B81EA89FFF46D45D6DDDBB8BF4404EB" box="[151,323,655,680]" italics="true" pageId="29" pageNumber="280">margineguttata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF46D45D700DBB9BF8E04EB" bold="true" box="[330,393,654,680]" pageId="29" pageNumber="280">
<taxonomicNameLabel id="00B257F2FFF46D45D700DBB9BF8E04EB" box="[330,393,654,680]" pageId="29" pageNumber="280" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF46D45D459DBB9BC2E04EB" box="[531,553,654,680]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="29" pageNumber="280" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF46D45D516DBB9BDAD04EB" box="[860,938,654,680]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="29" pageNumber="280">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="7D8A6613FFF46D45D6DDDF46BDD100EE" ID-DOI="http://doi.org/10.5281/zenodo.10821255" ID-Zenodo-Dep="10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="29" pageNumber="280" startId="29.[151,264,1649,1673]" targetBox="[168,1428,710,1617]" targetPageId="29" targetType="figure">
<paragraph id="294A369BFFF46D45D6DDDF46BDD100EE" blockId="29.[151,1436,1649,1710]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D6DDDF46BF5200CA" bold="true" box="[151,341,1649,1673]" pageId="29" pageNumber="280">FIGURES 81–84.</emphasis>
<taxonomicName id="EEF54D18FFF46D45D716DF46BCAF00CA" authorityName="Shavrin" authorityYear="2024" box="[348,680,1649,1673]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="margineguttata">
<emphasis id="1B81EA89FFF46D45D716DF46BCAF00CA" box="[348,680,1649,1673]" italics="true" pageId="29" pageNumber="280">Paraphloeostiba margineguttata</emphasis>
</taxonomicName>
(holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="61EF6510FFF46D46D68DDFC8BF61034E" lastPageId="30" lastPageNumber="281" pageId="29" pageNumber="280" type="description">
<paragraph id="294A369BFFF46D45D68DDFC8BA3A0121" blockId="29.[151,1437,1791,2034]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D68DDFC8BF5E015A" bold="true" box="[199,345,1791,1817]" pageId="29" pageNumber="280">Description.</emphasis>
Measurements (n=2): HW: 0.55–0.56; HL: 0.32; OL: 0.17; TL: 0.02; AL (
<typeStatus id="F64E8839FFF46D45D2FEDE37BB1A0159" box="[1204,1309,1792,1818]" pageId="29" pageNumber="280" type="holotype">holotype</typeStatus>
): 0.82; PL: 0.42; PWmax: 0.66; PWmin: 0.62; ESL: 0.80–0.82; EW: 0.90; MTbL (
<typeStatus id="F64E8839FFF46D45D59FDE13BA39017D" box="[981,1086,1828,1854]" pageId="29" pageNumber="280" type="holotype">holotype</typeStatus>
): 0.40; MTrL (
<typeStatus id="F64E8839FFF46D45D2A6DE13BB52017D" box="[1260,1365,1828,1854]" pageId="29" pageNumber="280" type="holotype">holotype</typeStatus>
): 0.17 (MTrL 1–4: 0.07; MTrL 5: 0.10); AW: 0.80; AedL: 0.45; BL: 2.57 (
<typeStatus id="F64E8839FFF46D45D5C3DE7FBDF70121" box="[905,1008,1864,1890]" pageId="29" pageNumber="280" type="holotype">holotype</typeStatus>
)–2.77.
</paragraph>
<paragraph id="294A369BFFF46D46D68DDE5BBA030726" blockId="29.[151,1437,1791,2034]" lastBlockId="30.[151,1437,151,1509]" lastPageId="30" lastPageNumber="281" pageId="29" pageNumber="280">
Habitus as in
<figureCitation id="B1CE2A1EFFF46D45D729DE5BBFB101C5" box="[355,438,1900,1926]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="29" pageNumber="280">Fig. 43</figureCitation>
. Head and pronotum reddish-brown; elytra yellowish-brown, with brown apical portion; abdominal tergites bicoloured: abdominal tergites IV–V yelllowish and VI–VIII brownish; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow. Head with very fine and dense punctation, finer and invisible on infraorbital portions; pronotum with moderately dense and fine punctation, slightly larger and sparser than that in middle portion of head, finer and invisible in midline; elytra with fine and moderately dense punctation, slightly deeper than that on pronotum, finer and sparser around scutellum and along suture; abdominal tergites with distinct fine and dense punctation. Head with dense microsculpture, finer and transverse on vertex and isodiametric and slightly coarser in middle; pronotum with dense isodiametric microreticulation, distinctly coarser and larger than that in middle portion of head, slightly finer in middle; microsculpture of elytra very dense, isodiametric, distinctly coarser than that in pronotum; abdominal tergites with fine transverse meshes.
</paragraph>
<paragraph id="294A369BFFF76D46D68DD858BAE60422" blockId="30.[151,1437,151,1509]" pageId="30" pageNumber="281">Head with short narrow and moderately deep anteocellar foveae, slightly divergent posteriad toward about level of posterior third of eyes; postocular carina absent. Ocelli large, located slightly below level of posterior margins of eyes; distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Antenna reaching apical margin of elytra when reclined, with antennomeres 6–7 about as broad as long; antennomere 3 disitnctly narrower and longer than 2, 4 about twice shorter than 3, 5 distinctly longer than 4, 6 slightly shorter and broader than 5, 7–9 slightly longer and broader than 6, 10 slightly shorter than 9, about as long as wide, apical antennomere about 1.4 times as long as 10, from middle strongly narrowed toward subacute apex.</paragraph>
<paragraph id="294A369BFFF76D46D68DDB5CBD8704EA" blockId="30.[151,1437,151,1509]" pageId="30" pageNumber="281">Pronotum 1.5 times as broad as long and 1.2 times as broad as head, widest slightly above middle, more narrowed posteriad than anteriad; laterobasal impressions shallow.</paragraph>
<paragraph id="294A369BFFF76D46D68DDB84BB30048E" blockId="30.[151,1437,151,1509]" box="[199,1335,691,717]" pageId="30" pageNumber="281">Elytra 1.9 times as long as pronotum, distinctly broadened posteriad, with rounded posterior margins.</paragraph>
<paragraph id="294A369BFFF76D46D68DDBE0BD3C04B2" blockId="30.[151,1437,151,1509]" box="[199,827,727,753]" pageId="30" pageNumber="281">Metatarsi slightly less than twice shorter than metatibia.</paragraph>
<paragraph id="294A369BFFF76D46D68DDBCCBC910556" blockId="30.[151,1437,151,1509]" box="[199,662,763,789]" pageId="30" pageNumber="281">Abdomen distinctly narrower than elytra.</paragraph>
<paragraph id="294A369BFFF76D46D68DDA28BAF6058A" blockId="30.[151,1437,151,1509]" pageId="30" pageNumber="281">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFF76D46D51EDA28BDA1057A" box="[852,934,799,825]" captionStart="FIGURES 81–84" captionStartId="29.[151,264,1649,1673]" captionTargetBox="[168,1428,710,1617]" captionTargetId="figure-145@29.[151,1436,704,1625]" captionTargetPageId="29" captionText="FIGURES 81–84. Paraphloeostiba margineguttata (holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="30" pageNumber="281">Fig. 83</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFF76D46D7AADA74BC31051E" box="[480,566,835,861]" captionStart="FIGURES 81–84" captionStartId="29.[151,264,1649,1673]" captionTargetBox="[168,1428,710,1617]" captionTargetId="figure-145@29.[151,1436,704,1625]" captionTargetPageId="29" captionText="FIGURES 81–84. Paraphloeostiba margineguttata (holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="30" pageNumber="281">Fig. 84</figureCitation>
). Aedeagus with moderately wide basal portion, gradually narrowed toward relatively wide median lobe with rounded apex; parameres relatively narrow, indistinctly broadened apicad, slightly not exceeding apex of median lobe, with three long apical setae; internal sac narrow, short, with three long and narrow sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFF76D46D4D5DA98BCF4058A" box="[671,755,943,969]" captionStart="FIGURES 81–84" captionStartId="29.[151,264,1649,1673]" captionTargetBox="[168,1428,710,1617]" captionTargetId="figure-145@29.[151,1436,704,1625]" captionTargetPageId="29" captionText="FIGURES 81–84. Paraphloeostiba margineguttata (holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="30" pageNumber="281">Fig. 81</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFF76D46D2D0DA98BAEA058A" box="[1178,1261,943,969]" captionStart="FIGURES 81–84" captionStartId="29.[151,264,1649,1673]" captionTargetBox="[168,1428,710,1617]" captionTargetId="figure-145@29.[151,1436,704,1625]" captionTargetPageId="29" captionText="FIGURES 81–84. Paraphloeostiba margineguttata (holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="30" pageNumber="281">Fig. 82</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFF76D46D68DDAE4BF9605AE" blockId="30.[151,1437,151,1509]" box="[199,401,979,1005]" pageId="30" pageNumber="281">Female unknown.</paragraph>
<paragraph id="294A369BFFF76D46D68DDAC1BF61034E" blockId="30.[151,1437,151,1509]" pageId="30" pageNumber="281">
<emphasis id="1B81EA89FFF76D46D68DDAC1BFB60252" bold="true" box="[199,433,1014,1041]" pageId="30" pageNumber="281">Comparative notes.</emphasis>
Based on the glossy body with moderately fine sculpture of the forebody and the general shape of the aedeagus,
<taxonomicName id="EEF54D18FFF76D46D7FADD2CBC7A0277" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[432,637,1051,1076]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="margineguttata" status="sp. n.">
<emphasis id="1B81EA89FFF76D46D7FADD2CBC7A0277" box="[432,637,1051,1076]" italics="true" pageId="30" pageNumber="281">P. margineguttata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF76D46D4C3DD2CBCC90276" bold="true" box="[649,718,1051,1077]" pageId="30" pageNumber="281">
<taxonomicNameLabel id="00B257F2FFF76D46D4C3DD2CBCC90276" box="[649,718,1051,1077]" pageId="30" pageNumber="281" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFF76D46D53CDD2CBAB90276" authority="Steel, 1960" authorityName="Steel" authorityYear="1960" box="[886,1214,1051,1077]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="solomonensis">
<emphasis id="1B81EA89FFF76D46D53CDD2CBA290276" box="[886,1070,1051,1077]" italics="true" pageId="30" pageNumber="281">P. solomonensis</emphasis>
<bibRefCitation id="4D644B6AFFF76D46D270DD2CBAB90276" author="Steel, W. O." box="[1082,1214,1051,1077]" pageId="30" pageNumber="281" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel, 1960</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="51E2760BFFF76D46D298DD2CBB9E0276" box="[1234,1433,1051,1077]" name="Solomon Islands" pageId="30" pageNumber="281">Solomon Islands</collectingCountry>
:
<collectingRegion id="EB31F879FFF76D46D6DDDD08BF22021A" box="[151,293,1087,1113]" country="Solomon Islands" name="Guadalcanal" pageId="30" pageNumber="281">Guadalcanal</collectingRegion>
) and
<taxonomicName id="EEF54D18FFF76D46D723DD08BC81021A" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[361,646,1087,1113]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="ullrichi">
<emphasis id="1B81EA89FFF76D46D723DD08BFDE021A" box="[361,473,1087,1113]" italics="true" pageId="30" pageNumber="281">P. ullrichi</emphasis>
Shavrin, 2023
</taxonomicName>
, recently described from
<collectingCountry id="51E2760BFFF76D46D5E7DD08BA8C021A" box="[941,1163,1087,1113]" name="Papua New Guinea" pageId="30" pageNumber="281">Papua New Guinea</collectingCountry>
(
<bibRefCitation id="4D644B6AFFF76D46D2D7DD08BB4D021A" author="Shavrin, A. V." box="[1181,1354,1087,1113]" pageId="30" pageNumber="281" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin 2023b</bibRefCitation>
). From
<taxonomicName id="EEF54D18FFF76D46D6DDDD54BF4A023E" authorityName="Steel" authorityYear="1960" box="[151,333,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="solomonensis">
<emphasis id="1B81EA89FFF76D46D6DDDD54BF4A023E" box="[151,333,1123,1149]" italics="true" pageId="30" pageNumber="281">P. solomonensis</emphasis>
</taxonomicName>
it differs by the paler and significantly larger body, with the absence of traces of longitudinal impressions in the middle of the pronotum, and distinctly broader apical portion of the median lobe. From
<taxonomicName id="EEF54D18FFF76D46D365DDB0BB9B02E2" authorityName="Shavrin" authorityYear="2023" box="[1327,1436,1159,1185]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="ullrichi">
<emphasis id="1B81EA89FFF76D46D365DDB0BB9B02E2" box="[1327,1436,1159,1185]" italics="true" pageId="30" pageNumber="281">P. ullrichi</emphasis>
</taxonomicName>
it differs by the paler and slightly larger body, broader median lobe and longer parameres. From all species of the genus it can be distinguished by the shape of the antennomeres 6–7 about as broad as long, and different structure of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF76D46D68DDC21BC210316" pageId="30" pageNumber="281" type="distribution">
<paragraph id="294A369BFFF76D46D68DDC21BC210316" blockId="30.[151,1437,151,1509]" pageId="30" pageNumber="281">
<emphasis id="1B81EA89FFF76D46D68DDC21BF660373" bold="true" box="[199,353,1302,1328]" pageId="30" pageNumber="281">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFF76D46D722DC20BCD00373" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[360,727,1303,1329]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="margineguttata" status="sp. nov.">
<emphasis id="1B81EA89FFF76D46D722DC20BCD00373" box="[360,727,1303,1329]" italics="true" pageId="30" pageNumber="281">Paraphloeostiba margineguttata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF76D46D495DC20BD180372" bold="true" box="[735,799,1303,1329]" pageId="30" pageNumber="281">
<taxonomicNameLabel id="00B257F2FFF76D46D495DC20BD180372" box="[735,799,1303,1329]" pageId="30" pageNumber="281" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFF76D46D273DC20BA6E0372" box="[1081,1129,1303,1329]" pageId="30" pageNumber="281">type</typeStatus>
locality in Paniai Province of Western New
<collectingCountry id="51E2760BFFF76D46D719DC0CBFA00316" box="[339,423,1339,1365]" name="Guinea-Bissau" pageId="30" pageNumber="281">Guinea</collectingCountry>
,
<collectingCountry id="51E2760BFFF76D46D7F8DC0CBC240316" box="[434,547,1339,1365]" name="Indonesia" pageId="30" pageNumber="281">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF76D46D68DDC68BF3703A6" pageId="30" pageNumber="281" type="etymology">
<paragraph id="294A369BFFF76D46D68DDC68BCFB03DE" blockId="30.[151,1437,151,1509]" pageId="30" pageNumber="281">
<emphasis id="1B81EA89FFF76D46D68DDC68BF49033A" bold="true" box="[199,334,1375,1401]" pageId="30" pageNumber="281">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFF76D46D48ADC68BD70033B" box="[704,887,1375,1400]" italics="true" pageId="30" pageNumber="281">margineguttatus</emphasis>
, -
<emphasis id="1B81EA89FFF76D46D5C4DC68BDD1033B" box="[910,982,1375,1400]" italics="true" pageId="30" pageNumber="281">a, -um</emphasis>
(with spots at margins). It alludes to the wide darkened spots in apical portions of each elytron.
</paragraph>
<paragraph id="294A369BFFF76D46D68DDC91BF3703A6" blockId="30.[151,1437,151,1509]" pageId="30" pageNumber="281">
<emphasis id="1B81EA89FFF76D46D68DDC91BF4C0383" bold="true" box="[199,331,1446,1472]" pageId="30" pageNumber="281">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="EE0D9B7EFFF76D46D512DC90BA0A0382" box="[856,1037,1446,1473]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="30" pageNumber="281" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100 to 2500 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

231
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<paragraph id="294A369BFFF76D46D6DDDF19BC23000A" blockId="30.[151,548,1582,1645]" box="[151,548,1582,1609]" pageId="30" pageNumber="281">
<heading id="720281F7FFF76D46D6DDDF19BC23000A" bold="true" box="[151,548,1582,1609]" fontSize="11" level="1" pageId="30" pageNumber="281" reason="1">
<emphasis id="1B81EA89FFF76D46D6DDDF19BC23000A" bold="true" box="[151,548,1582,1609]" pageId="30" pageNumber="281">
<taxonomicName id="EEF54D18FFF76D46D6DDDF19BFDC000A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,475,1582,1609]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="okapensis" status="sp. nov.">
<emphasis id="1B81EA89FFF76D46D6DDDF19BFDC000A" bold="true" box="[151,475,1582,1609]" italics="true" pageId="30" pageNumber="281">Paraphloeostiba okapensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFF76D46D7A8DF18BC23000A" box="[482,548,1583,1609]" pageId="30" pageNumber="281" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF76D46D6DDDF64BF56002E" box="[151,337,1618,1645]" pageId="30" pageNumber="281" type="description">
<paragraph id="294A369BFFF76D46D6DDDF64BF56002E" blockId="30.[151,548,1582,1645]" box="[151,337,1618,1645]" pageId="30" pageNumber="281">
(
<figureCitation id="B1CE2A1EFFF76D46D6D5DF64BEF1002E" box="[159,246,1618,1645]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="30" pageNumber="281">Figs 75</figureCitation>
,
<figureCitation id="B1CE2A1EFFF76D46D748DF65BF4E002E" box="[258,329,1618,1645]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="30" pageNumber="281">85–91</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF76D46D6DDDFACBA8401CE" pageId="30" pageNumber="281" type="materials_examined">
<paragraph id="294A369BFFF76D46D6DDDFACBFAA00BE" blockId="30.[151,1437,1690,2041]" pageId="30" pageNumber="281">
<emphasis id="1B81EA89FFF76D46D6DDDFACBF4100F7" bold="true" box="[151,326,1690,1717]" pageId="30" pageNumber="281">Type material.</emphasis>
<typeStatus id="F64E8839FFF76D46D704DFACBFB100F6" box="[334,438,1691,1717]" pageId="30" pageNumber="281" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFF76D46D7F4DFACBFD400F6" box="[446,467,1691,1717]" count="1" pageId="30" pageNumber="281" type="male">♂</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFF76D46D422DFACBCAB00F6" box="[616,684,1691,1717]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="30" pageNumber="281">NEW</collectionCode>
<collectingCountry id="51E2760BFFF76D46D4FEDFACBD1900F6" box="[692,798,1691,1717]" name="Guinea-Bissau" pageId="30" pageNumber="281">GUINEA</collectingCountry>
Okapa, |
<date id="5D4B105BFFF76D46D5C6DFADBA0900F6" box="[908,1038,1690,1717]" pageId="30" pageNumber="281" value="1965-06-16">16.VI.1965</date>
, flying in | daylight’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF76D46D335DFACBB9F00F6" box="[1407,1432,1691,1717]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="30" pageNumber="281">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFF76D46D415DF88BCFF009A" box="[607,760,1727,1753]" pageId="30" pageNumber="281" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF76D46D543DF88BDC3009A" authorityName="Steel" authorityYear="1960" box="[777,964,1727,1753]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF76D46D543DF88BDC3009A" box="[777,964,1727,1753]" italics="true" pageId="30" pageNumber="281">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF76D46D59FDF88BA44009A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[981,1091,1727,1753]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="okapensis" status="sp. nov.">
<emphasis id="1B81EA89FFF76D46D59FDF88BA44009A" box="[981,1091,1727,1753]" italics="true" pageId="30" pageNumber="281">okapensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF76D46D203DF88BA80009A" bold="true" box="[1097,1159,1727,1753]" pageId="30" pageNumber="281">
<taxonomicNameLabel id="00B257F2FFF76D46D203DF88BA80009A" box="[1097,1159,1727,1753]" pageId="30" pageNumber="281" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF76D46D346DF88BB25009A" box="[1292,1314,1727,1753]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="281" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF76D46D70DDFD4BFA600BE" box="[327,417,1763,1789]" pageId="30" pageNumber="281">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFF76D46D68DDE30BA8401CE" blockId="30.[151,1437,1690,2041]" pageId="30" pageNumber="281">
<typeStatus id="F64E8839FFF76D46D68DDE30BF3F0162" box="[199,312,1799,1825]" pageId="30" pageNumber="281" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFF76D46D70DDE30BF8D0162" box="[327,394,1799,1825]" count="1" pageId="30" pageNumber="281" type="female">1 ♀♀</specimenCount>
(left antennomeres 7–11 missing; dissected): same data as the holotype (
<collectionCode id="4FE4AE5EFFF76D46D2ADDE30BB460162" box="[1255,1345,1799,1825]" pageId="30" pageNumber="281">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFF76D46D313DE30BB9B0162" box="[1369,1436,1799,1825]" count="2" pageId="30" pageNumber="281" type="female">2 ♀♀</specimenCount>
(underside of the card of
<specimenCount id="3FF3FD12FFF76D46D7FDDE1CBC6B0106" box="[439,620,1835,1861]" count="3" pageId="30" pageNumber="281" type="generic">three specimens</specimenCount>
with handwritten ‘H 28’): ‘
<collectionCode id="4FE4AE5EFFF76D46D5E9DE1CBDE00106" box="[931,999,1835,1861]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="30" pageNumber="281">NEW</collectionCode>
<collectingCountry id="51E2760BFFF76D46D5BADE1CBA5D0106" box="[1008,1114,1835,1861]" name="Guinea-Bissau" pageId="30" pageNumber="281">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFF76D46D28DDE1CBB410106" box="[1223,1350,1835,1861]" pageId="30" pageNumber="281" value="1964-11-23">23.XI.1964</date>
| under bark of Xamata’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF76D46D79EDE78BFEA012A" box="[468,493,1871,1897]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="30" pageNumber="281">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFF76D46D588DE78BA1B012A" box="[962,1052,1871,1897]" pageId="30" pageNumber="281">BMNH</collectionCode>
). All
<typeStatus id="F64E8839FFF76D46D212DE78BAC4012A" box="[1112,1219,1871,1897]" pageId="30" pageNumber="281" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFF76D46D77DDE44BFC001CE" box="[311,455,1907,1933]" pageId="30" pageNumber="281" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF76D46D791DE44BC9101CE" authorityName="Steel" authorityYear="1960" box="[475,662,1907,1933]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF76D46D791DE44BC9101CE" box="[475,662,1907,1933]" italics="true" pageId="30" pageNumber="281">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF76D46D4E3DE44BD1001CE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[681,791,1907,1933]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="281" phylum="Arthropoda" rank="species" species="okapensis" status="sp. nov.">
<emphasis id="1B81EA89FFF76D46D4E3DE44BD1001CE" box="[681,791,1907,1933]" italics="true" pageId="30" pageNumber="281">okapensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF76D46D554DE44BD5901CE" bold="true" box="[798,862,1907,1933]" pageId="30" pageNumber="281">
<taxonomicNameLabel id="00B257F2FFF76D46D554DE44BD5901CE" box="[798,862,1907,1933]" pageId="30" pageNumber="281" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF76D46D5ADDE44BDFA01CE" box="[999,1021,1907,1933]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="281" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF76D78D68DDEA1BD3B0556" lastPageId="32" lastPageNumber="283" pageId="30" pageNumber="281" type="description">
<paragraph id="294A369BFFF76D46D68DDEA1BAB301BA" blockId="30.[151,1437,1690,2041]" pageId="30" pageNumber="281">
<emphasis id="1B81EA89FFF76D46D68DDEA1BF5E01F3" bold="true" box="[199,345,1942,1968]" pageId="30" pageNumber="281">Description.</emphasis>
Measurements (n=4): HW: 0.39–0.41; HL: 0.21–0.24; OL: 0.13–0.15; TL: 0.02; AL (
<typeStatus id="F64E8839FFF76D46D362DEA0BB9601F2" box="[1320,1425,1943,1969]" pageId="30" pageNumber="281" type="holotype">holotype</typeStatus>
): 0.62; PL: 0.31–0.32; PW: 0.46–0.50; ESL: 0.54–0.57; EW: 0.64–0.67; MTbL (
<typeStatus id="F64E8839FFF76D46D25DDE8CBA870196" box="[1047,1152,1979,2005]" pageId="30" pageNumber="281" type="holotype">holotype</typeStatus>
): 0.36; MTrL (
<typeStatus id="F64E8839FFF76D46D362DE8CBB960196" box="[1320,1425,1979,2005]" pageId="30" pageNumber="281" type="holotype">holotype</typeStatus>
): 0.16 (MTrL 1–4: 0.06; MTrL 5: 0.10); AW: 0.56–0.65; AedL: 0.32; BL: 1.63–1.86 (
<typeStatus id="F64E8839FFF76D46D275DEE8BAAF01BA" box="[1087,1192,2015,2041]" pageId="30" pageNumber="281" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="294A369BFFF66D47D68DD9A0BF420792" blockId="31.[151,1437,150,645]" pageId="31" pageNumber="282">
Habitus as in
<figureCitation id="B1CE2A1EFFF66D47D723D9A0BFB906F2" box="[361,446,150,177]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="31" pageNumber="282">Fig. 75</figureCitation>
. Body reddish-brown, with slightly darker head and abdomen and slightly paler basal and middle portions of elytra (latero-apical and basal margins of elytra slightly darker); antennomeres 6–11 brown; mouthparts, antennomeres 1–4 and legs yellow. Head, neck, pronotum and scutellum without visible punctation; elytra with dense, moderately large and deep punctation, slightly denser and finer around scutellum, finer and sparser along suture; abdominal tergites with very fine and dense punctation, finer and indistinct on tergites VII–VIII. Head with very dense, microsculpture, slightly transverse in vertex and isodiametric in middle; neck with dense transverse microreticulation; pronotum with very dense and coarse microsculpture similar to that in middle portion of head, transverse in mediobasal portion; elytra with distinct, dense and isodiametric microreticulation, coarser in basal and apical portions.
</paragraph>
<paragraph id="294A369BFFF66D47D68DD8ECBD6E04C6" blockId="31.[151,1437,150,645]" pageId="31" pageNumber="282">Head 1.7–1.8 times as broad as long, with short and narrow linear anteocellar foveae, reaching level of posterior third length of eyes. Ocelli large, located at level of posterior margins of eyes; distance between ocelli slightly larger than distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about 1.5 times as long as preapical segment. Antennomere 5 slightly broader than 4, 6 slightly longer and distinctly broader than 5, 7–8 distinctly longer and broader than 6, 9–10 slightly longer than 8.</paragraph>
<caption id="7D8A6613FFF66D47D6DDDEA6BC8001B2" ID-DOI="http://doi.org/10.5281/zenodo.10821259" ID-Zenodo-Dep="10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="31" pageNumber="282" startId="31.[151,264,1937,1961]" targetBox="[164,1428,677,1906]" targetPageId="31" targetType="figure">
<paragraph id="294A369BFFF66D47D6DDDEA6BC8001B2" blockId="31.[151,1436,1937,2034]" pageId="31" pageNumber="282">
<emphasis id="1B81EA89FFF66D47D6DDDEA6BF5201EA" bold="true" box="[151,341,1937,1961]" pageId="31" pageNumber="282">FIGURES 85–91.</emphasis>
<taxonomicName id="EEF54D18FFF66D47D716DEA6BC7701EA" authorityName="Shavrin" authorityYear="2024" box="[348,624,1937,1961]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="282" phylum="Arthropoda" rank="species" species="okapensis">
<emphasis id="1B81EA89FFF66D47D716DEA6BC7701EA" box="[348,624,1937,1961]" italics="true" pageId="31" pageNumber="282">Paraphloeostiba okapensis</emphasis>
</taxonomicName>
(paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFC96D78D68DD9A0BD9606BA" blockId="32.[151,1437,150,933]" pageId="32" pageNumber="283">Pronotum 1.4–1.5 times as broad as long, 1.1–1.2 times as broad as head, widest in middle, gradually narrrowed both anteriad and posteriad; posterior angles subacute; middle portion with distinct longitudinal impressions, widest in about middle; laterobasal impressions wide and moderately deep.</paragraph>
<paragraph id="294A369BFFC96D78D68DD834BC6D075E" blockId="32.[151,1437,150,933]" box="[199,618,259,285]" pageId="32" pageNumber="283">Elytra 1.7 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFC96D78D68DD810BB5E0792" blockId="32.[151,1437,150,933]" pageId="32" pageNumber="283">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC96D78D51ED810BDA10702" box="[852,934,295,321]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="32" pageNumber="283">Fig. 87</figureCitation>
). Posterior margin of abdominal sternite VIII strongly and widely concave (
<figureCitation id="B1CE2A1EFFC96D78D7A9D87CBC320726" box="[483,565,331,357]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="32" pageNumber="283">Fig. 88</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward small rounded apex; parameres not reaching apex of median lobe, slightly narrowed apically, with three long apical setae and four elongate setae on preapical inner margin of each paramere; internal sac moderately short and narrow, with two very long and narrow sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFC96D78D54DD880BD5C0792" box="[775,859,438,465]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="32" pageNumber="283">Fig. 85</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC96D78D349D880BB520792" box="[1283,1365,439,465]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="32" pageNumber="283">Fig. 86</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC96D78D68DD8ECBF1D047E" blockId="32.[151,1437,150,933]" pageId="32" pageNumber="283">
Female. Posterior margin of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFC96D78D56ED8ECBD7F07B6" box="[804,888,475,501]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="32" pageNumber="283">Fig. 89</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFC96D78D213D8ECBAAA07B6" box="[1113,1197,475,501]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="32" pageNumber="283">Fig. 90</figureCitation>
) truncate. Accessory sclerite with wide basal portion, slightly narrowed toward somewhat truncate apex (
<figureCitation id="B1CE2A1EFFC96D78D224D8C8BAC2045A" box="[1134,1221,511,537]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="32" pageNumber="283">Fig. 91</figureCitation>
). Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFC96D78D68DDB71BD3B0556" blockId="32.[151,1437,150,933]" pageId="32" pageNumber="283">
<emphasis id="1B81EA89FFC96D78D68DDB71BFB30422" bold="true" box="[199,436,582,609]" pageId="32" pageNumber="283">Comparative notes.</emphasis>
Based on the general shape of the aedeagus with moderately broad median lobe and narrow parameres, distinctly shorter than apex of the median lobe,
<taxonomicName id="EEF54D18FFC96D78D5CFDB5CBA0804C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[901,1039,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="okapensis" status="sp. n.">
<emphasis id="1B81EA89FFC96D78D5CFDB5CBA0804C6" box="[901,1039,619,645]" italics="true" pageId="32" pageNumber="283">P. okapensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC96D78D25CDB5CBA5204C6" bold="true" box="[1046,1109,619,645]" pageId="32" pageNumber="283">
<taxonomicNameLabel id="00B257F2FFC96D78D25CDB5CBA5204C6" box="[1046,1109,619,645]" pageId="32" pageNumber="283" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFC96D78D2ADDB5CBB4804C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1255,1359,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="telnovi" status="sp. n.">
<emphasis id="1B81EA89FFC96D78D2ADDB5CBB4804C6" box="[1255,1359,619,645]" italics="true" pageId="32" pageNumber="283">P. telnovi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC96D78D31CDB5CBB9104C6" bold="true" box="[1366,1430,619,645]" pageId="32" pageNumber="283">
<taxonomicNameLabel id="00B257F2FFC96D78D31CDB5CBB9104C6" box="[1366,1430,619,645]" pageId="32" pageNumber="283" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the shape of the pronotum, from the middle gradually narrowed both anteriad and posteriad, with narrower apical portions not protuded apicad, the finer punctation of the forebody, the presence of the distinct longitudinal impressions in the middle of the pronotum, narrower apex of the median lobe, slightly broader parameres and different structure of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC96D78D68DDA29BB87057A" box="[199,1408,798,825]" pageId="32" pageNumber="283" type="distribution">
<paragraph id="294A369BFFC96D78D68DDA29BB87057A" blockId="32.[151,1437,150,933]" box="[199,1408,798,825]" pageId="32" pageNumber="283">
<emphasis id="1B81EA89FFC96D78D68DDA29BF66057B" bold="true" box="[199,353,798,824]" pageId="32" pageNumber="283">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFC96D78D72DDA28BC90057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[359,663,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="283" phylum="Arthropoda" rank="species" species="okapensis" status="sp. nov.">
<emphasis id="1B81EA89FFC96D78D72DDA28BC90057A" box="[359,663,799,825]" italics="true" pageId="32" pageNumber="283">Paraphloeostiba okapensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC96D78D4D5DA28BCD9057A" bold="true" box="[671,734,799,825]" pageId="32" pageNumber="283">
<taxonomicNameLabel id="00B257F2FFC96D78D4D5DA28BCD9057A" box="[671,734,799,825]" pageId="32" pageNumber="283" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFC96D78D5BBDA28BA26057A" box="[1009,1057,799,825]" pageId="32" pageNumber="283">type</typeStatus>
locality in
<collectingCountry id="51E2760BFFC96D78D2EADA28BB7B057A" box="[1184,1404,799,825]" name="Papua New Guinea" pageId="32" pageNumber="283">Papua New Guinea</collectingCountry>
.
</paragraph>
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<subSubSection id="61EF6510FFC96D78D68DDA74BA6E05E6" pageId="32" pageNumber="283" type="etymology">
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<emphasis id="1B81EA89FFC96D78D68DDA74BF49051E" bold="true" box="[199,334,835,861]" pageId="32" pageNumber="283">Etymology.</emphasis>
The specific epithet is the latinized adjective derived from the area the specimens were collected from.
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<emphasis id="1B81EA89FFC96D78D68DDABDBF4C05E7" bold="true" box="[199,331,906,932]" pageId="32" pageNumber="283">Bionomics.</emphasis>
Specimens were collected flying in daylight or were found under bark.
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<emphasis id="1B81EA89FFF96D48D6DDDA51BFDA05C3" bold="true" box="[151,477,870,897]" italics="true" pageId="16" pageNumber="267">Paraphloeostiba conjuncta</emphasis>
Shavrin, 2023
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Material examined.
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(ca.
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<geoCoordinate id="4CC1505CFFF96D48D56EDDB6BDC702DF" box="[804,960,1153,1180]" degrees="145" direction="east" minutes="32" orientation="longitude" pageId="16" pageNumber="267" precision="15" seconds="40" value="145.54445">145°32’40’’E</geoCoordinate>
),
<date id="5D4B105BFFF96D48D59EDDB6BA4002DF" box="[980,1095,1153,1180]" pageId="16" pageNumber="267" value="1965-11-08">
<collectingDate id="4D0FE9B3FFF96D48D59EDDB6BA4002DF" box="[980,1095,1153,1180]" pageId="16" pageNumber="267" value="1965-11-08">8.XI.1965</collectingDate>
</date>
, rotting fruit’, ‘
<collectionCode id="4FE4AE5EFFF96D48D2BBDDB6BB0D02D8" box="[1265,1290,1153,1179]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="267">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="999D3CC6FFF96D48D35FDDB6BFB202FC" collectionCode="BMNH" country="Guinea-Bissau" location="Hornabrook" pageId="16" pageNumber="267" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFF96D48D35FDDB6BB9B02D8" LSID="urn:lsid:plazi:treatment:A15C878DFFF96D48D6DDDA51BB1F03F8:2C2A6040FFF96D48D35FDDB6BB9B02D8" box="[1301,1436,1153,1179]" country="Guinea-Bissau" name="Hornabrook" pageId="16" pageNumber="267">Hornabrook</location>
<specimenCode id="79539EE0FFF96D48D6DDDD92BF3C02FC" box="[151,315,1189,1215]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="16" pageNumber="267">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFF96D48D718DD92BFAB02FC" box="[338,428,1189,1215]" pageId="16" pageNumber="267">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF96D48D68DDDFEBB1F03F8" pageId="16" pageNumber="267" type="discussion">
<paragraph id="294A369BFFF96D48D68DDDFEBA6B0368" blockId="16.[151,1437,1045,1467]" pageId="16" pageNumber="267">
<emphasis id="1B81EA89FFF96D48D68DDDFEBF3C02A0" bold="true" box="[199,315,1225,1251]" pageId="16" pageNumber="267">Remarks.</emphasis>
<taxonomicName id="EEF54D18FFF96D48D70EDDFEBC7202A0" authorityName="Shavrin" authorityYear="2023" box="[324,629,1225,1251]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF96D48D70EDDFEBC7202A0" box="[324,629,1225,1251]" italics="true" pageId="16" pageNumber="267">Paraphloeostiba conjuncta</emphasis>
</taxonomicName>
was recently described from north-eastern part of
<collectingCountry id="51E2760BFFF96D48D2F0DDFEBB9E02A0" box="[1210,1433,1225,1251]" name="Papua New Guinea" pageId="16" pageNumber="267">Papua New Guinea</collectingCountry>
. The body length of this species can vary from
<quantity id="EE0D9B7EFFF96D48D4EADDDABD5E0344" box="[672,857,1261,1288]" metricMagnitude="-3" metricUnit="m" metricValue="2.065" metricValueMax="2.35" metricValueMin="1.78" pageId="16" pageNumber="267" unit="mm" value="2.065" valueMax="2.35" valueMin="1.78">1.78 to 2.35 mm</quantity>
(
<typeStatus id="F64E8839FFF96D48D52DDDDABDC80344" box="[871,975,1261,1287]" pageId="16" pageNumber="267" type="holotype">holotype</typeStatus>
:
<quantity id="EE0D9B7EFFF96D48D593DDDABA460344" box="[985,1089,1261,1287]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="16" pageNumber="267" unit="mm" value="1.8">1.80 mm</quantity>
). Prior to the present study, the female of
<taxonomicName id="EEF54D18FFF96D48D743DC25BF910368" authorityName="Shavrin" authorityYear="2023" box="[265,406,1297,1323]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF96D48D743DC25BF1D0368" box="[265,282,1298,1323]" italics="true" pageId="16" pageNumber="267">P</emphasis>
.
<emphasis id="1B81EA89FFF96D48D762DC25BF910368" box="[296,406,1298,1323]" italics="true" pageId="16" pageNumber="267">conjuncta</emphasis>
</taxonomicName>
was unknown. The description of the female is presented below.
</paragraph>
<paragraph id="294A369BFFF96D48D68DDC02BABA0330" blockId="16.[151,1437,1045,1467]" pageId="16" pageNumber="267">
Female. Posterior margin of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFF96D48D569DC02BD70030C" box="[803,887,1333,1359]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="16" pageNumber="267">Fig. 36</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFF96D48D21DDC02BAAC030C" box="[1111,1195,1333,1359]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="16" pageNumber="267">Fig. 37</figureCitation>
) rounded. Accessory sclerite wide, gradually narrowed toward rounded apex (
<figureCitation id="B1CE2A1EFFF96D48D55BDC6EBD620330" box="[785,869,1369,1395]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="16" pageNumber="267">Fig. 38</figureCitation>
). Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFF96D48D68DDC4ABB1F03F8" blockId="16.[151,1437,1045,1467]" pageId="16" pageNumber="267">
<taxonomicName id="EEF54D18FFF96D48D68DDC4ABFFE03D4" authorityName="Shavrin" authorityYear="2023" box="[199,505,1405,1431]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF96D48D68DDC4ABFFE03D4" box="[199,505,1405,1431]" italics="true" pageId="16" pageNumber="267">Paraphloeostiba conjuncta</emphasis>
</taxonomicName>
is known from several localities in Western New
<collectingCountry id="51E2760BFFF96D48D273DC4ABA8D03D4" box="[1081,1162,1405,1431]" name="Guinea-Bissau" pageId="16" pageNumber="267">Guinea</collectingCountry>
(
<collectingCountry id="51E2760BFFF96D48D2D1DC4ABB0C03D4" box="[1179,1291,1405,1431]" name="Indonesia" pageId="16" pageNumber="267">Indonesia</collectingCountry>
) and eastern part of
<collectingCountry id="51E2760BFFF96D48D6A2DC96BFC303F8" box="[232,452,1441,1467]" name="Papua New Guinea" pageId="16" pageNumber="267">Papua New Guinea</collectingCountry>
. It is here recorded from Western New
<collectingCountry id="51E2760BFFF96D48D531DC96BDCB03F8" box="[891,972,1441,1467]" name="Guinea-Bissau" pageId="16" pageNumber="267">Guinea</collectingCountry>
(
<collectingCountry id="51E2760BFFF96D48D590DC96BA4D03F8" box="[986,1098,1441,1467]" name="Indonesia" pageId="16" pageNumber="267">Indonesia</collectingCountry>
) for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

422
data/A1/5C/87/A15C878DFFF96D4CD6DDDF3BBF12045A.xml Normal file
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@ -0,0 +1,422 @@
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<mods:title id="9EE2737B6F48062CD337C84C26A60526">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<treatment id="A15C878DFFF96D4CD6DDDF3BBF12045A" LSID="urn:lsid:plazi:treatment:A15C878DFFF96D4CD6DDDF3BBF12045A" httpUri="http://treatment.plazi.org/id/A15C878DFFF96D4CD6DDDF3BBF12045A" lastPageId="20" lastPageNumber="271" pageId="16" pageNumber="267">
<subSubSection id="61EF6510FFF96D48D6DDDF3BBC370064" box="[151,560,1548,1575]" pageId="16" pageNumber="267" type="nomenclature">
<paragraph id="294A369BFFF96D48D6DDDF3BBC370064" blockId="16.[151,560,1548,1612]" box="[151,560,1548,1575]" pageId="16" pageNumber="267">
<heading id="720281F7FFF96D48D6DDDF3BBC370064" bold="true" box="[151,560,1548,1575]" fontSize="11" level="1" pageId="16" pageNumber="267" reason="1">
<emphasis id="1B81EA89FFF96D48D6DDDF3BBC370064" bold="true" box="[151,560,1548,1575]" pageId="16" pageNumber="267">
<taxonomicName id="EEF54D18FFF96D48D6DDDF3BBFE10064" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,486,1548,1575]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="coprophila" status="sp. nov.">
<emphasis id="1B81EA89FFF96D48D6DDDF3BBFE10064" bold="true" box="[151,486,1548,1575]" italics="true" pageId="16" pageNumber="267">Paraphloeostiba coprophila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFF96D48D7A7DF3ABC370064" box="[493,560,1549,1575]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF96D48D6DDDF06BF20000F" box="[151,295,1585,1612]" pageId="16" pageNumber="267" type="description">
<paragraph id="294A369BFFF96D48D6DDDF06BF20000F" blockId="16.[151,560,1548,1612]" box="[151,295,1585,1612]" pageId="16" pageNumber="267">
(
<figureCitation id="B1CE2A1EFFF96D48D6D5DF06BF18000F" box="[159,287,1585,1612]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="16" pageNumber="267">Figs 46–52</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF96D4AD6DDDF4EBB6E021B" lastPageId="18" lastPageNumber="269" pageId="16" pageNumber="267" type="materials_examined">
<paragraph id="294A369BFFF96D48D6DDDF4EBD170098" blockId="16.[151,1437,1657,2044]" pageId="16" pageNumber="267">
<emphasis id="1B81EA89FFF96D48D6DDDF4EBF4300D0" bold="true" box="[151,324,1657,1683]" pageId="16" pageNumber="267">Type material.</emphasis>
<typeStatus id="F64E8839FFF96D48D701DF4EBFB400D0" box="[331,435,1657,1683]" pageId="16" pageNumber="267" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFF96D48D7F3DF4EBFC900D0" box="[441,462,1657,1683]" count="1" pageId="16" pageNumber="267" type="male">♂</specimenCount>
(disseced; left antennomeres 3–11 missing): ‘
<collectionCode id="4FE4AE5EFFF96D48D583DF4EBA0A00D0" box="[969,1037,1657,1683]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="16" pageNumber="267">NEW</collectionCode>
<collectingCountry id="51E2760BFFF96D48D25EDF4EBA7900D0" box="[1044,1150,1657,1683]" name="Guinea-Bissau" pageId="16" pageNumber="267">GUINEA</collectingCountry>
| Okasa
<date id="5D4B105BFFF96D48D296DF4EBB5F00D7" box="[1244,1368,1657,1684]" pageId="16" pageNumber="267" value="1965-04-20">20.IV.1965</date>
| wild sinses fruit’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF96D48D7ECDFAABFB800F4" box="[422,447,1693,1719]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="267">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFF96D48D5E8DFAABA3C00F4" box="[930,1083,1693,1719]" pageId="16" pageNumber="267" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF96D48D21BDFAABB0B00F4" authorityName="Steel" authorityYear="1960" box="[1105,1292,1693,1719]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF96D48D21BDFAABB0B00F4" box="[1105,1292,1693,1719]" italics="true" pageId="16" pageNumber="267">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF96D48D368DFAABB9B00F4" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1314,1436,1693,1719]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="coprophila" status="sp. nov.">
<emphasis id="1B81EA89FFF96D48D368DFAABB9B00F4" box="[1314,1436,1693,1719]" italics="true" pageId="16" pageNumber="267">coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF96D48D6DDDFF6BED00098" bold="true" box="[151,215,1729,1755]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="00B257F2FFF96D48D6DDDFF6BED00098" box="[151,215,1729,1755]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF96D48D72ADFF6BF710098" box="[352,374,1729,1755]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="267" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF96D48D4E3DFF6BD040098" box="[681,771,1729,1755]" pageId="16" pageNumber="267">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFF96D4AD68DDFD2BB6E021B" blockId="16.[151,1437,1657,2044]" lastBlockId="18.[151,1437,978,2042]" lastPageId="18" lastPageNumber="269" pageId="16" pageNumber="267">
<typeStatus id="F64E8839FFF96D48D68DDFD2BF3F00BC" box="[199,312,1765,1791]" pageId="16" pageNumber="267" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFF96D48D70DDFD2BF7200BC" box="[327,373,1765,1791]" count="1" pageId="16" pageNumber="267" type="male">1 ♂</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFF96D48D45ADFD2BC5300BC" box="[528,596,1765,1791]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="16" pageNumber="267">NEW</collectionCode>
<collectingCountry id="51E2760BFFF96D48D415DFD2BCCE00BC" box="[607,713,1765,1791]" name="Guinea-Bissau" pageId="16" pageNumber="267">GUINEA</collectingCountry>
| Okapa, |
<date id="5D4B105BFFF96D48D505DFD2BDC30143" box="[847,964,1765,1792]" pageId="16" pageNumber="267" value="1965-05-23">
23.
<collectionCode id="4FE4AE5EFFF96D48D53ADFD2BD810143" box="[880,902,1765,1792]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="267" type="Museum">V</collectionCode>
.1965
</date>
, cow | dung and beating’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF96D48D335DFD2BB9F00BC" box="[1407,1432,1765,1791]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="267">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFF96D48D432DE3EBCD50160" box="[632,722,1801,1827]" pageId="16" pageNumber="267">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFF96D48D4A0DE3EBD1F0160" box="[746,792,1801,1827]" count="1" pageId="16" pageNumber="267" type="male">1 ♂</specimenCount>
(antennomeres missing),
<specimenCount id="3FF3FD12FFF96D48D20CDE3EBA8E0160" box="[1094,1161,1801,1827]" count="2" pageId="16" pageNumber="267" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFF96D48D2D6DE3EBB3B0160" box="[1180,1340,1801,1827]" count="1" pageId="16" pageNumber="267" type="generic">one specimen</specimenCount>
without right antenna; underside of the card with handwritten number ‘H13’): ‘
<collectionCode id="4FE4AE5EFFF96D48D5F3DE1ABDFA0104" box="[953,1021,1837,1863]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="16" pageNumber="267">NEW</collectionCode>
<collectingCountry id="51E2760BFFF96D48D24CDE1ABA770104" box="[1030,1136,1837,1863]" name="Guinea-Bissau" pageId="16" pageNumber="267">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFF96D48D294DE1ABB54010B" box="[1246,1363,1837,1864]" pageId="16" pageNumber="267" value="1965-05-23">
23.
<collectionCode id="4FE4AE5EFFF96D48D2B4DE1ABB13010B" box="[1278,1300,1837,1864]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="267" type="Museum">V</collectionCode>
.1965
</date>
, | cow dung’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF96D48D728DE66BF7C0128" box="[354,379,1873,1899]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="267">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFF96D48D512DE66BDB50128" box="[856,946,1873,1899]" pageId="16" pageNumber="267">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFF96D48D58DDE66BA000128" box="[967,1031,1873,1899]" count="8" pageId="16" pageNumber="267" type="female">8 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFF96D48D25DDE66BAB40128" box="[1047,1203,1873,1899]" count="1" pageId="16" pageNumber="267" type="generic">one specimen</specimenCount>
without left elytron; underside of the card with handwritten number ‘71H’): ‘
<collectionCode id="4FE4AE5EFFF96D48D552DE42BD5B01CC" box="[792,860,1909,1935]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="16" pageNumber="267">NEW</collectionCode>
<collectingCountry id="51E2760BFFF96D48D52FDE42BDC801CC" box="[869,975,1909,1935]" name="Guinea-Bissau" pageId="16" pageNumber="267">GUINEA</collectingCountry>
Okapa |
<date id="5D4B105BFFF96D48D272DE42BAAA01D3" box="[1080,1197,1909,1936]" pageId="16" pageNumber="267" value="1965-05-24">
24.
<collectionCode id="4FE4AE5EFFF96D48D213DE42BA6801D3" box="[1113,1135,1909,1936]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="267" type="Museum">V</collectionCode>
.1965
</date>
, small white | mould on Kamata stump’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF96D48D7BFDEAEBC0901F0" box="[501,526,1945,1971]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="267">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFF96D48D5B8DEAEBA4B01F0" box="[1010,1100,1945,1971]" pageId="16" pageNumber="267">BMNH</collectionCode>
); 1 unsexed specimen (apical part of the abdomen missing; underside of the card with handwritten number ‘H62’): ‘
<collectionCode id="4FE4AE5EFFF96D48D231DE8ABAB80194" box="[1147,1215,1981,2007]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="16" pageNumber="267">NEW</collectionCode>
<collectingCountry id="51E2760BFFF96D48D283DE8ABB340194" box="[1225,1331,1981,2007]" name="Guinea-Bissau" pageId="16" pageNumber="267">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFF96D48D6DDDED6BF0B01BF" box="[151,268,2017,2044]" pageId="16" pageNumber="267" value="1965-05-22">
22.
<collectionCode id="4FE4AE5EFFF96D48D6F3DED6BEC801BF" box="[185,207,2017,2044]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="267" type="Museum">V</collectionCode>
.1965
</date>
, | dead leaves’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF96D48D475DED6BC5F01B8" box="[575,600,2017,2043]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="267">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFF96D48D272DED6BA9501B8" box="[1080,1170,2017,2043]" pageId="16" pageNumber="267">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFF96D48D2EDDED6BAD401B8" box="[1191,1235,2017,2043]" count="1" pageId="16" pageNumber="267" type="female">1 ♀</specimenCount>
(underside of the card with handwritten number ‘117H’): ‘
<collectionCode id="4FE4AE5EFFFB6D4AD42ADAE5BCA305AF" box="[608,676,978,1004]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="18" pageNumber="269">NEW</collectionCode>
<collectingCountry id="51E2760BFFFB6D4AD4E6DAE5BD1105AF" box="[684,790,978,1004]" name="Guinea-Bissau" pageId="18" pageNumber="269">GUINEA</collectingCountry>
In-| dona, |
<date id="5D4B105BFFFB6D4AD5D4DAE5BA2705AE" box="[926,1056,978,1005]" pageId="18" pageNumber="269" value="1965-11-17">17.XI.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFFB6D4AD2E5DAE5BACF05AF" box="[1199,1224,978,1004]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="269">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFFB6D4AD7DFDAC1BFE80253" box="[405,495,1014,1040]" pageId="18" pageNumber="269">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFFB6D4AD448DAC1BC2B0252" box="[514,556,1014,1041]" count="1" pageId="18" pageNumber="269" type="female">1 ♀</specimenCount>
(underside of the card with handwritten number ‘H41’): ‘
<collectionCode id="4FE4AE5EFFFB6D4AD2E7DAC1BAF60253" box="[1197,1265,1014,1040]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="18" pageNumber="269">NEW</collectionCode>
<collectingCountry id="51E2760BFFFB6D4AD2B3DAC1BB640253" box="[1273,1379,1014,1040]" name="Guinea-Bissau" pageId="18" pageNumber="269">GUINEA</collectingCountry>
Pu- | rosa,
<date id="5D4B105BFFFB6D4AD69EDD2DBF580276" box="[212,351,1050,1077]" pageId="18" pageNumber="269" value="1965-07-21">21.VII.1965</date>
, | wet leaves etc.’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFFB6D4AD4E1DD2DBCC30277" box="[683,708,1050,1076]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="269">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFFB6D4AD2D1DD2DBAF20277" box="[1179,1269,1050,1076]" pageId="18" pageNumber="269">BMNH</collectionCode>
). All
<typeStatus id="F64E8839FFFB6D4AD37BDD2DBB9B0277" box="[1329,1436,1050,1076]" pageId="18" pageNumber="269" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFFB6D4AD45BDD09BCA6021B" box="[529,673,1086,1112]" pageId="18" pageNumber="269" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFFB6D4AD4FFDD0ABD770214" authorityName="Steel" authorityYear="1960" box="[693,880,1085,1111]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFFB6D4AD4FFDD0ABD770214" box="[693,880,1085,1111]" italics="true" pageId="18" pageNumber="269">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFFB6D4AD5C9DD0ABDFA0214" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[899,1021,1085,1111]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="coprophila" status="sp. nov.">
<emphasis id="1B81EA89FFFB6D4AD5C9DD0ABDFA0214" box="[899,1021,1085,1111]" italics="true" pageId="18" pageNumber="269">coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFB6D4AD24EDD09BA43021B" bold="true" box="[1028,1092,1086,1112]" pageId="18" pageNumber="269">
<taxonomicNameLabel id="00B257F2FFFB6D4AD24EDD09BA43021B" box="[1028,1092,1086,1112]" pageId="18" pageNumber="269" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFFB6D4AD287DD09BAE4021B" box="[1229,1251,1086,1112]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="18" pageNumber="269" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<caption id="7D8A6613FFF86D49D6DDDE6EBB8F01D6" ID-DOI="http://doi.org/10.5281/zenodo.10821233" ID-Zenodo-Dep="10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="17" pageNumber="268" startId="17.[151,264,1881,1905]" targetBox="[177,1411,194,1844]" targetPageId="17" targetType="figure">
<paragraph id="294A369BFFF86D49D6DDDE6EBB8F01D6" blockId="17.[151,1437,1881,1941]" pageId="17" pageNumber="268">
<emphasis id="1B81EA89FFF86D49D6DDDE6EBF530132" bold="true" box="[151,340,1881,1905]" pageId="17" pageNumber="268">FIGURES 39–44.</emphasis>
Habitus of
<taxonomicName id="EEF54D18FFF86D49D782DE6EBC760132" authorityName="Steel" authorityYear="1960" box="[456,625,1881,1905]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF86D49D782DE6EBC760132" box="[456,625,1881,1905]" italics="true" pageId="17" pageNumber="268">Paraphloeostiba</emphasis>
</taxonomicName>
: 39–
<taxonomicName id="EEF54D18FFF86D49D4E9DE6DBD130132" authorityName="Cameron" authorityYear="1952" box="[675,788,1882,1905]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="coriacea">
<emphasis id="1B81EA89FFF86D49D4E9DE6DBD130132" box="[675,788,1882,1905]" italics="true" pageId="17" pageNumber="268">P. coriacea</emphasis>
</taxonomicName>
(Western New Guinea), 40–
<taxonomicName id="EEF54D18FFF86D49D279DE6DBA8E0132" authorityName="Shavrin" authorityYear="2024" box="[1075,1161,1882,1905]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="iriana">
<emphasis id="1B81EA89FFF86D49D279DE6DBA8E0132" box="[1075,1161,1882,1905]" italics="true" pageId="17" pageNumber="268">P. iriana</emphasis>
</taxonomicName>
(paratype), 41–
<taxonomicName id="EEF54D18FFF86D49D363DE6DBB9A0132" authorityName="Shavrin" authorityYear="2024" box="[1321,1437,1881,1905]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="latissima">
<emphasis id="1B81EA89FFF86D49D363DE6DBB9A0132" box="[1321,1437,1881,1905]" italics="true" pageId="17" pageNumber="268">P. latissima</emphasis>
</taxonomicName>
(paratype), 42–
<taxonomicName id="EEF54D18FFF86D49D778DE49BFFB01D6" authorityName="Shavrin" authorityYear="2024" box="[306,508,1917,1941]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="guadalcanalensis">
<emphasis id="1B81EA89FFF86D49D778DE49BFFB01D6" box="[306,508,1917,1941]" italics="true" pageId="17" pageNumber="268">P. guadalcanalensis</emphasis>
</taxonomicName>
(paratype), 43–
<taxonomicName id="EEF54D18FFF86D49D4D7DE49BD5601D6" authorityName="Shavrin" authorityYear="2024" box="[669,849,1918,1941]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="margineguttata">
<emphasis id="1B81EA89FFF86D49D4D7DE49BD5601D6" box="[669,849,1918,1941]" italics="true" pageId="17" pageNumber="268">P. margineguttata</emphasis>
</taxonomicName>
(holotype), 44–
<taxonomicName id="EEF54D18FFF86D49D5BEDE49BA4501D6" authorityName="Shavrin" authorityYear="2024" box="[1012,1090,1917,1941]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="steeli">
<emphasis id="1B81EA89FFF86D49D5BEDE49BA4501D6" box="[1012,1090,1917,1941]" italics="true" pageId="17" pageNumber="268">P. steeli</emphasis>
</taxonomicName>
(holotype). Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="7D8A6613FFFB6D4AD6DDDA42BB1405CD" ID-DOI="http://doi.org/10.5281/zenodo.10821237" ID-Zenodo-Dep="10821237" box="[151,1299,885,910]" httpUri="https://zenodo.org/record/10821237/files/figure.png" pageId="18" pageNumber="269" startId="18.[151,264,885,909]" targetBox="[151,1436,193,862]" targetPageId="18" targetType="figure">
<paragraph id="294A369BFFFB6D4AD6DDDA42BB1405CD" blockId="18.[151,1299,885,910]" box="[151,1299,885,910]" pageId="18" pageNumber="269">
<emphasis id="1B81EA89FFFB6D4AD6DDDA42BF2905CD" bold="true" box="[151,302,885,910]" pageId="18" pageNumber="269">FIGURES 45.</emphasis>
Distribution of
<taxonomicName id="EEF54D18FFFB6D4AD79ADA41BCDE05CE" authorityName="Cameron" authorityYear="1952" box="[464,729,886,910]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="coriacea">
<emphasis id="1B81EA89FFFB6D4AD79ADA41BCDE05CE" box="[464,729,886,910]" italics="true" pageId="18" pageNumber="269">Paraphloeostiba coriacea</emphasis>
</taxonomicName>
(circles) and
<taxonomicName id="EEF54D18FFFB6D4AD529DA41BA0E05CD" baseAuthorityName="MacLeay" baseAuthorityYear="1873" box="[867,1033,886,910]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="269" phylum="Arthropoda" rank="species" species="gayndahensis">
<emphasis id="1B81EA89FFFB6D4AD529DA41BD7505CE" box="[867,882,886,909]" italics="true" pageId="18" pageNumber="269">P</emphasis>
.
<emphasis id="1B81EA89FFFB6D4AD535DA41BA0E05CD" box="[895,1033,886,910]" italics="true" pageId="18" pageNumber="269">gayndahensis</emphasis>
</taxonomicName>
(squares) in New Guinea.
</paragraph>
</caption>
<subSubSection id="61EF6510FFFB6D4CD68DDD56BFDA0726" lastPageId="20" lastPageNumber="271" pageId="18" pageNumber="269" type="description">
<paragraph id="294A369BFFFB6D4AD68DDD56BF1402A5" blockId="18.[151,1437,978,2042]" pageId="18" pageNumber="269">
<emphasis id="1B81EA89FFFB6D4AD68DDD56BF5E0238" bold="true" box="[199,345,1121,1147]" pageId="18" pageNumber="269">Description.</emphasis>
Measurements (n=17): HW: 0.25–0.37; HL: 0.20–0.27; OL: 0.11–0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFFB6D4AD362DD56BB960238" box="[1320,1425,1121,1147]" pageId="18" pageNumber="269" type="holotype">holotype</typeStatus>
): 0.57; PL: 0.26–0.30; PWmax: 0.46–0.52; PWmin: 0.41–0.47; ESL: 0.55–0.62; EW: 0.62–0.70; MTbL (
<typeStatus id="F64E8839FFFB6D4AD362DDB2BB9602DC" box="[1320,1425,1157,1183]" pageId="18" pageNumber="269" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFFB6D4AD77FDD9EBF990280" box="[309,414,1193,1219]" pageId="18" pageNumber="269" type="holotype">holotype</typeStatus>
): 0.14 (MTrL 1–4: 0.04; MTrL 5: 0.10); AW: 0.57–0.60; AedL: 0.32; BL: 1.47–2.07 (
<typeStatus id="F64E8839FFFB6D4AD6D5DDFBBF0F02A5" box="[159,264,1228,1254]" pageId="18" pageNumber="269" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="294A369BFFFB6D4AD68DDDC7BCC803A3" blockId="18.[151,1437,978,2042]" pageId="18" pageNumber="269">Body yellow-brown to brown, with slightly paler elytra and slightly darkened apical portions of each elytron; antennomeres 7–11 brown; mouthparts, antennomeres 1–6 and legs yellow. Head without visible punctation; middle portion of neck without or with several very fine punctures; pronotum with very fine and moderately dense punctation, indistinct or absent in middle; elytra with fine and dense punctation, slightly larger than that in pronotum, distinctly finer and sparser around scutellum and along suture. Head with dense and coarse microsculpture; pronotum with dense and coarse microreticulation, transverse in apical portion and isodiametric in middle; abdominal tergites with dense transverse or isodiametric microreticulation.</paragraph>
<paragraph id="294A369BFFFB6D4AD68DDCDDBB5500D0" blockId="18.[151,1437,978,2042]" pageId="18" pageNumber="269">Head 1.2–1.3 times as broad as long, with narrow and moderately deep anteocellar foveae reaching level of basal third of eyes; postocular carina absent. Ocelli moderately large, located at level of posterior margins of eyes; distance between ocelli slightly larger than distance between ocellus and posterior margin of eyes. Antennomere 4 slightly longer than wide and about twice shorter than 3, 5 slightly shorter than 4, 6 disitnctly broader than 5, 7–9 slightly broader than 6, 10 slightly longer than 9, apical antennomere slightly more than twice as long as 10.</paragraph>
<paragraph id="294A369BFFFB6D4AD68DDFA9BCC300BD" blockId="18.[151,1437,978,2042]" pageId="18" pageNumber="269">Pronotum 1.7 times as broad as long, 1.4–1.8 times as broad as head, widest in middle, distinctly more narrowed posteriad than anteriad; hind angles obtuse or subacute; disc without or with traces of very shallow and indistinct wide impressions in middle, broadened posteriad.</paragraph>
<paragraph id="294A369BFFFB6D4AD68DDE3FBC470160" blockId="18.[151,1437,978,2042]" box="[199,576,1800,1827]" pageId="18" pageNumber="269">Elytra twice as long as pronotum.</paragraph>
<paragraph id="294A369BFFFB6D4AD68DDE1BBC850105" blockId="18.[151,1437,978,2042]" box="[199,642,1836,1862]" pageId="18" pageNumber="269">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="294A369BFFFB6D4AD68DDE67BAA201BA" blockId="18.[151,1437,978,2042]" pageId="18" pageNumber="269">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFFB6D4AD51EDE67BDA00129" box="[852,935,1872,1898]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="18" pageNumber="269">Fig. 48</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFFB6D4AD748DE44BF5101CE" box="[258,342,1907,1933]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="18" pageNumber="269">Fig. 49</figureCitation>
). Aedeagus with very wide basal portion, significantly narrowed toward very narrow median lobe with small rounded apex; parameres moderately wide, slightly not reaching apex of median lobe, with three long apical and several relatively short setae on inner margin of each paramere; internal sac moderately short and witde, without strongly sclerotized structures (
<figureCitation id="B1CE2A1EFFFB6D4AD419DEE8BCA001BA" box="[595,679,2015,2041]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="18" pageNumber="269">Fig. 46</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFFB6D4AD204DEE8BAA601BA" box="[1102,1185,2015,2041]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="18" pageNumber="269">Fig. 47</figureCitation>
.
</paragraph>
<caption id="7D8A6613FFFA6D4BD6DDDCC3BC800017" ID-DOI="http://doi.org/10.5281/zenodo.10821239" ID-Zenodo-Dep="10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="19" pageNumber="270" startId="19.[151,264,1524,1548]" targetBox="[172,1415,189,1492]" targetPageId="19" targetType="figure">
<paragraph id="294A369BFFFA6D4BD6DDDCC3BC800017" blockId="19.[151,1437,1524,1621]" pageId="19" pageNumber="270">
<emphasis id="1B81EA89FFFA6D4BD6DDDCC3BF53004F" bold="true" box="[151,340,1524,1548]" pageId="19" pageNumber="270">FIGURES 46–52.</emphasis>
<taxonomicName id="EEF54D18FFFA6D4BD710DCC3BC70004F" authorityName="Shavrin" authorityYear="2024" box="[346,631,1524,1548]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="coprophila">
<emphasis id="1B81EA89FFFA6D4BD710DCC3BC70004F" box="[346,631,1524,1548]" italics="true" pageId="19" pageNumber="270">Paraphloeostiba coprophila</emphasis>
</taxonomicName>
(paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFFA6D4BD68DDFAEBDAC009B" blockId="19.[151,1436,1689,2040]" pageId="19" pageNumber="270">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFFA6D4BD537DFAEBDD400F7" box="[893,979,1689,1716]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="19" pageNumber="270">Fig. 50</figureCitation>
). Posterior margin of abdominal sternite VIII widely rounded (
<figureCitation id="B1CE2A1EFFFA6D4BD7C7DF8ABFE7009B" box="[397,480,1725,1752]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="19" pageNumber="270">Fig. 51</figureCitation>
). Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFFA6D4BD51FDF8ABDA0009B" box="[853,935,1725,1752]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="19" pageNumber="270">Fig. 52</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFFA6D4BD68DDFD6BCFC0100" blockId="19.[151,1436,1689,2040]" pageId="19" pageNumber="270">
<emphasis id="1B81EA89FFFA6D4BD68DDFD6BFB600B8" bold="true" box="[199,433,1761,1787]" pageId="19" pageNumber="270">Comparative notes.</emphasis>
Based on the very narrow median lobe and the parameres,
<taxonomicName id="EEF54D18FFFA6D4BD205DFD5BAE100B8" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1103,1254,1761,1787]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFFA6D4BD205DFD5BAE100B8" box="[1103,1254,1761,1787]" italics="true" pageId="19" pageNumber="270">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFA6D4BD2A7DFD6BB2A00B8" bold="true" box="[1261,1325,1761,1787]" pageId="19" pageNumber="270">
<taxonomicNameLabel id="00B257F2FFFA6D4BD2A7DFD6BB2A00B8" box="[1261,1325,1761,1787]" pageId="19" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFFA6D4BD6FFDE31BF6F015C" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[181,360,1798,1824]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFFA6D4BD6FFDE31BEC1015C" box="[181,198,1798,1823]" italics="true" pageId="19" pageNumber="270">P</emphasis>
.
<emphasis id="1B81EA89FFFA6D4BD69EDE31BF6F015C" box="[212,360,1798,1823]" italics="true" pageId="19" pageNumber="270">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFA6D4BD73ADE32BFB7015C" bold="true" box="[368,432,1797,1823]" pageId="19" pageNumber="270">
<taxonomicNameLabel id="00B257F2FFFA6D4BD73ADE32BFB7015C" box="[368,432,1797,1823]" pageId="19" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFFA6D4BD7F4DE31BC31015C" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[446,566,1797,1823]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFFA6D4BD7F4DE31BC31015C" box="[446,566,1797,1823]" italics="true" pageId="19" pageNumber="270">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFA6D4BD474DE32BC79015C" bold="true" box="[574,638,1797,1823]" pageId="19" pageNumber="270">
<taxonomicNameLabel id="00B257F2FFFA6D4BD474DE32BC79015C" box="[574,638,1797,1823]" pageId="19" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFFA6D4BD4C6DE31BD1F015C" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[652,792,1797,1823]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFFA6D4BD4C6DE31BD1F015C" box="[652,792,1797,1823]" italics="true" pageId="19" pageNumber="270">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFA6D4BD555DE32BD58015C" bold="true" box="[799,863,1797,1823]" pageId="19" pageNumber="270">
<taxonomicNameLabel id="00B257F2FFFA6D4BD555DE32BD58015C" box="[799,863,1797,1823]" pageId="19" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFFA6D4BD5DCDE31BDEA015C" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[918,1005,1797,1823]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFFA6D4BD5DCDE31BDEA015C" box="[918,1005,1797,1823]" italics="true" pageId="19" pageNumber="270">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFA6D4BD5BFDE32BA32015C" bold="true" box="[1013,1077,1797,1823]" pageId="19" pageNumber="270">
<taxonomicNameLabel id="00B257F2FFFA6D4BD5BFDE32BA32015C" box="[1013,1077,1797,1823]" pageId="19" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFFA6D4BD68DDE7ABF1501F3" blockId="19.[151,1436,1689,2040]" pageId="19" pageNumber="270">
from
<taxonomicName id="EEF54D18FFFA6D4BD742DE79BFB80124" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[264,447,1870,1896]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFFA6D4BD742DE79BF1E0124" box="[264,281,1870,1895]" italics="true" pageId="19" pageNumber="270">P</emphasis>
.
<emphasis id="1B81EA89FFFA6D4BD761DE79BFB80124" box="[299,447,1870,1895]" italics="true" pageId="19" pageNumber="270">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFA6D4BD783DE7ABC0B0124" bold="true" box="[457,524,1869,1895]" pageId="19" pageNumber="270">
<taxonomicNameLabel id="00B257F2FFFA6D4BD783DE7ABC0B0124" box="[457,524,1869,1895]" pageId="19" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller and paler body, the absence of the punctation on the head, the finer punctation on the pronotum, slightly more transverse pronotum, slightly longer elytra and slightly shorter parameres;
</paragraph>
<paragraph id="294A369BFFFA6D4BD68DDE8EBD2501BB" blockId="19.[151,1436,1689,2040]" pageId="19" pageNumber="270">
from
<taxonomicName id="EEF54D18FFFA6D4BD74FDE8DBF790190" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,382,1977,2003]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFFA6D4BD74FDE8DBF790190" box="[261,382,1977,2003]" italics="true" pageId="19" pageNumber="270">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFA6D4BD7CCDE8EBFC10190" bold="true" box="[390,454,1977,2003]" pageId="19" pageNumber="270">
<taxonomicNameLabel id="00B257F2FFFA6D4BD7CCDE8EBFC10190" box="[390,454,1977,2003]" pageId="19" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker elytra, the absence of the punctation on the head, the finer punctation on the pronotum, the longer elytra and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFFD6D4CD68DD9A0BAC806F2" blockId="20.[151,1437,151,537]" box="[199,1231,151,177]" pageId="20" pageNumber="271">
from
<taxonomicName id="EEF54D18FFFD6D4CD74FD9A0BF9706F2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,400,151,177]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFFD6D4CD74FD9A0BF9706F2" box="[261,400,151,177]" italics="true" pageId="20" pageNumber="271">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFD6D4CD7DDD9A0BFD006F2" bold="true" box="[407,471,151,177]" pageId="20" pageNumber="271">
<taxonomicNameLabel id="00B257F2FFFD6D4CD7DDD9A0BFD006F2" box="[407,471,151,177]" pageId="20" pageNumber="271" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the longer body and the elytra, and slightly narrower parameres;
</paragraph>
<paragraph id="294A369BFFFD6D4CD68DD98CBEFC075E" blockId="20.[151,1437,151,537]" pageId="20" pageNumber="271">
from
<taxonomicName id="EEF54D18FFFD6D4CD742D98CBF640696" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[264,355,187,213]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFFD6D4CD742D98CBF640696" box="[264,355,187,213]" italics="true" pageId="20" pageNumber="271">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFD6D4CD724D98CBFB60696" bold="true" box="[366,433,187,213]" pageId="20" pageNumber="271">
<taxonomicNameLabel id="00B257F2FFFD6D4CD724D98CBFB60696" box="[366,433,187,213]" pageId="20" pageNumber="271" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the paler body, the absence of the punctation on the head and elevations between punctures on the longer elytra, the finer punctation of the pronotum, and the shorter parameres with narrower apical portions.
</paragraph>
<paragraph id="294A369BFFFD6D4CD68DD810BFDA0726" blockId="20.[151,1437,151,537]" pageId="20" pageNumber="271">
From all these species,
<taxonomicName id="EEF54D18FFFD6D4CD79BD810BC650702" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[465,610,295,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFFD6D4CD79BD810BFE50703" box="[465,482,295,320]" italics="true" pageId="20" pageNumber="271">P</emphasis>
.
<emphasis id="1B81EA89FFFD6D4CD7B8D810BC650702" box="[498,610,295,321]" italics="true" pageId="20" pageNumber="271">betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFD6D4CD420D810BCAD0702" bold="true" box="[618,682,295,321]" pageId="20" pageNumber="271">
<taxonomicNameLabel id="00B257F2FFFD6D4CD420D810BCAD0702" box="[618,682,295,321]" pageId="20" pageNumber="271" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the the details of the external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFD6D4CD68DD859BB5E07CA" box="[199,1369,366,393]" pageId="20" pageNumber="271" type="distribution">
<paragraph id="294A369BFFFD6D4CD68DD859BB5E07CA" blockId="20.[151,1437,151,537]" box="[199,1369,366,393]" pageId="20" pageNumber="271">
<emphasis id="1B81EA89FFFD6D4CD68DD859BF6607CB" bold="true" box="[199,353,366,392]" pageId="20" pageNumber="271">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFFD6D4CD72DD858BCA407CA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[359,675,367,393]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="coprophila" status="sp. nov.">
<emphasis id="1B81EA89FFFD6D4CD72DD858BCA407CA" box="[359,675,367,393]" italics="true" pageId="20" pageNumber="271">Paraphloeostiba coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFD6D4CD4E0D858BCEE07CA" bold="true" box="[682,745,367,393]" pageId="20" pageNumber="271">
<taxonomicNameLabel id="00B257F2FFFD6D4CD4E0D858BCEE07CA" box="[682,745,367,393]" pageId="20" pageNumber="271" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from several localities in
<collectingCountry id="51E2760BFFFD6D4CD233D858BB5107CA" box="[1145,1366,367,393]" name="Papua New Guinea" pageId="20" pageNumber="271">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFD6D4CD68DD8A4BF12045A" pageId="20" pageNumber="271" type="etymology">
<paragraph id="294A369BFFFD6D4CD68DD8A4BD820792" blockId="20.[151,1437,151,537]" pageId="20" pageNumber="271">
<emphasis id="1B81EA89FFFD6D4CD68DD8A4BF4907EE" bold="true" box="[199,334,403,429]" pageId="20" pageNumber="271">Etymology.</emphasis>
The specific epithet is derived from two Greek words “κόπρος” (dung) and φίλος (loving). It alludes to the fact that some
<typeStatus id="F64E8839FFFD6D4CD792D880BC440792" box="[472,579,439,465]" pageId="20" pageNumber="271" type="paratype">paratypes</typeStatus>
were collected in cow dung.
</paragraph>
<paragraph id="294A369BFFFD6D4CD68DD8EDBF12045A" blockId="20.[151,1437,151,537]" pageId="20" pageNumber="271">
<emphasis id="1B81EA89FFFD6D4CD68DD8EDBF4C07B7" bold="true" box="[199,331,474,500]" pageId="20" pageNumber="271">Bionomics.</emphasis>
Specimens were collected by sifting leaves or were found in decaying organic material (fruits, cow dung, etc.).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="6FC5263003B16C5F2987C4395D3C09C3">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<emphasis id="1B81EA89FFFD6D4CD6DDDB68BFCC0439" bold="true" box="[151,459,607,634]" italics="true" pageId="20" pageNumber="271">Paraphloeostiba coriacea</emphasis>
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,
<figureCitation id="B1CE2A1EFFFD6D4CD748DBB3BF2704DC" box="[258,288,644,671]" captionStart="FIGURES 45" captionStartId="18.[151,264,885,909]" captionTargetBox="[151,1436,193,862]" captionTargetId="figure-22@18.[151,1436,181,862]" captionTargetPageId="18" captionText="FIGURES 45. Distribution of Paraphloeostiba coriacea (circles) and P. gayndahensis (squares) in New Guinea." figureDoi="http://doi.org/10.5281/zenodo.10821237" httpUri="https://zenodo.org/record/10821237/files/figure.png" pageId="20" pageNumber="271">45</figureCitation>
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<figureCitation id="B1CE2A1EFFFD6D4CD766DBB3BF7304DC" box="[300,372,644,671]" captionStart="FIGURES 53–56" captionStartId="21.[151,264,1187,1211]" captionTargetBox="[159,1428,187,1164]" captionTargetId="figure-16@21.[151,1436,181,1164]" captionTargetPageId="21" captionText="FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="20" pageNumber="271">53–56</figureCitation>
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<emphasis id="1B81EA89FFFD6D4CD6DDDBF4BF8E0498" box="[151,393,707,731]" italics="true" pageId="20" pageNumber="271">Phloeonomus coriaceus</emphasis>
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<taxonomicName id="EEF54D18FFFD6D4CD6DDDBD5BF9804BA" ID-CoL="6TR9M" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[151,415,738,762]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="coriacea">
<emphasis id="1B81EA89FFFD6D4CD6DDDBD5BF9804BA" box="[151,415,738,762]" italics="true" pageId="20" pageNumber="271">Paraphloeostiba coriacea</emphasis>
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<bibRefCitation id="4D644B6AFFFD6D4CD7E6DBD5BC4A04B9" author="Steel, W. O." box="[428,589,738,762]" pageId="20" pageNumber="271" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960: 146</bibRefCitation>
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Material examined.
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Sinak,
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</quantity>
14.-
<date id="5D4B105BFFFD6D4CD6F7DABBBF4505E4" box="[189,322,908,935]" pageId="20" pageNumber="271" value="1995-12-17">
<collectingDate id="4D0FE9B3FFFD6D4CD6F7DABBBF4505E4" box="[189,322,908,935]" pageId="20" pageNumber="271" value="1995-12-17">17.12.1995</collectingDate>
</date>
leg.
<collectorName id="8400534DFFFD6D4CD732DABABFEC05E5" box="[376,491,908,935]" pageId="20" pageNumber="271">A. Riedel’</collectorName>
(
<specimenCount id="3FF3FD12FFFD6D4CD7B1DABABC3905E4" box="[507,574,909,935]" count="2" pageId="20" pageNumber="271" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFFD6D4CD400DABABC8B05E4" box="[586,652,909,935]" count="2" pageId="20" pageNumber="271" type="female">2 ♀♀</specimenCount>
: cSh;
<specimenCount id="3FF3FD12FFFD6D4CD498DABBBD1305E4" box="[722,788,908,935]" count="6" pageId="20" pageNumber="271" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFFD6D4CD56ADABABD6505E4" box="[800,866,909,935]" count="8" pageId="20" pageNumber="271" type="female">8 ♀♀</specimenCount>
:
<collectionCode id="4FE4AE5EFFFD6D4CD525DABABDBB05E4" box="[879,956,909,935]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="271">NMW</collectionCode>
);
<specimenCount id="3FF3FD12FFFD6D4CD585DABABDFB05E4" box="[975,1020,909,935]" count="1" pageId="20" pageNumber="271" type="female">1 ♀</specimenCount>
: ‘
<collectionCode id="4FE4AE5EFFFD6D4CD244DABABAA805E4" box="[1038,1199,909,935]" pageId="20" pageNumber="271">IRIAN JAYA</collectionCode>
</materialsCitation>
:
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<location id="2C2A6040FFFD6D4CD2F3DABBBB4605E4" LSID="urn:lsid:plazi:treatment:A15C878DFFFD6D4DD6DDDB68BC3301BB:2C2A6040FFFD6D4CD2F3DABBBB4605E4" box="[1209,1345,908,935]" country="Guinea-Bissau" county="Ilaga" municipality="Sinak" name="Paniai Prov." pageId="20" pageNumber="271">Paniai Prov.</location>
|
<collectingMunicipality id="C92EACE1FFFD6D4CD31FDABBBB9F05E5" box="[1365,1432,908,934]" pageId="20" pageNumber="271">Sinak</collectingMunicipality>
, trail to
<collectingCounty id="C02B4E17FFFD6D4CD6ADDA87BF180589" box="[231,287,944,970]" pageId="20" pageNumber="271">Ilaga</collectingCounty>
|
<quantity id="EE0D9B7EFFFD6D4CD778DA87BFC30588" box="[306,452,944,971]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="20" pageNumber="271" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">
<elevation id="A2D8D1A8FFFD6D4CD778DA87BFC30588" box="[306,452,944,971]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="20" pageNumber="271" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100-2500m</elevation>
</quantity>
,
<date id="5D4B105BFFFD6D4CD784DA87BC5E0588" box="[462,601,944,971]" pageId="20" pageNumber="271" value="1995-12-15">
<collectingDate id="4D0FE9B3FFFD6D4CD784DA87BC5E0588" box="[462,601,944,971]" pageId="20" pageNumber="271" value="1995-12-15">15. 12. 1995</collectingDate>
</date>
| leg.
<collectorName id="8400534DFFFD6D4CD4D0DA87BD0B0589" box="[666,780,944,970]" pageId="20" pageNumber="271">A. Riedel’</collectorName>
&lt;printed&gt; (
<collectionCode id="4FE4AE5EFFFD6D4CD5DADA87BDD90589" box="[912,990,944,970]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="271">NMW</collectionCode>
);
<specimenCount id="3FF3FD12FFFD6D4CD5BADA87BA1B0588" box="[1008,1052,944,971]" count="1" pageId="20" pageNumber="271" type="female">1 ♀</specimenCount>
: ‘
<collectionCode id="4FE4AE5EFFFD6D4CD266DA87BA770589" box="[1068,1136,944,970]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="20" pageNumber="271">NEW</collectionCode>
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<materialsCitation id="999D3CC6FFFD6D4CD23DDA87BF4A05AD" collectingDate="1965-11-17" collectionCode="R" country="Guinea-Bissau" location="Indona" pageId="20" pageNumber="271" specimenCount="1">
<collectingCountry id="51E2760BFFFD6D4CD23DDA87BAE60589" box="[1143,1249,944,970]" name="Guinea-Bissau" pageId="20" pageNumber="271">GUINEA</collectingCountry>
<location id="2C2A6040FFFD6D4CD2A2DA87BB310589" LSID="urn:lsid:plazi:treatment:A15C878DFFFD6D4DD6DDDB68BC3301BB:2C2A6040FFFD6D4CD2A2DA87BB310589" box="[1256,1334,944,970]" country="Guinea-Bissau" name="Indona" pageId="20" pageNumber="271">Indona</location>
[Sefuna]
<date id="5D4B105BFFFD6D4CD6DDDAE4BF1905AD" box="[151,286,979,1006]" pageId="20" pageNumber="271" value="1965-11-17">
<collectingDate id="4D0FE9B3FFFD6D4CD6DDDAE4BF1905AD" box="[151,286,979,1006]" pageId="20" pageNumber="271" value="1965-11-17">17.XI.1965</collectingDate>
</date>
’, ‘
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</materialsCitation>
.
<materialsCitation id="999D3CC6FFFD6D4CD712DAE4BD1705AD" box="[344,784,979,1006]" collectionCode="BMNH" country="Guinea-Bissau" location="Hornabrook" pageId="20" pageNumber="271" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFFD6D4CD712DAE4BFD805AE" LSID="urn:lsid:plazi:treatment:A15C878DFFFD6D4DD6DDDB68BC3301BB:2C2A6040FFFD6D4CD712DAE4BFD805AE" box="[344,479,979,1005]" country="Guinea-Bissau" name="Hornabrook" pageId="20" pageNumber="271">Hornabrook</location>
|
<specimenCode id="79539EE0FFFD6D4CD7B9DAE3BC9005AD" box="[499,663,980,1006]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="20" pageNumber="271">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFFD6D4CD4E4DAE3BD0F05AD" box="[686,776,980,1006]" pageId="20" pageNumber="271">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFD6D4DD68DDAC0BD1500BF" lastPageId="21" lastPageNumber="272" pageId="20" pageNumber="271" type="description">
<paragraph id="294A369BFFFD6D4CD68DDAC0BD86023F" blockId="20.[151,1437,802,2034]" pageId="20" pageNumber="271">
<emphasis id="1B81EA89FFFD6D4CD68DDAC0BF720252" bold="true" box="[199,373,1015,1041]" pageId="20" pageNumber="271">Redescription.</emphasis>
Measurements (n=133, including the
<typeStatus id="F64E8839FFFD6D4CD563DAC0BD8C0252" box="[809,907,1015,1041]" pageId="20" pageNumber="271" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="4D644B6AFFFD6D4CD5D6DAC0BA4E0252" author="Shavrin, A. V." box="[924,1097,1015,1041]" pageId="20" pageNumber="271" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin 2023b</bibRefCitation>
)): HW: 0.43–0.52; HL: 0.24– 0.36; OL: 0.14–0.17; TL: 0.02–0.06; AL (
<typeStatus id="F64E8839FFFD6D4CD427DD2DBCD10277" box="[621,726,1050,1076]" pageId="20" pageNumber="271" type="holotype">holotype</typeStatus>
): 0.85; PL: 0.32–0.42; PWmax: 0.52–0.69; PWmin: 0.47–0.66; ESL: 0.63–0.76; EW: 0.76–0.91; MTbL (
<typeStatus id="F64E8839FFFD6D4CD42DDD09BCD7021B" box="[615,720,1086,1112]" pageId="20" pageNumber="271" type="holotype">holotype</typeStatus>
): 0.34; MTrL (
<typeStatus id="F64E8839FFFD6D4CD53EDD09BDDA021B" box="[884,989,1086,1112]" pageId="20" pageNumber="271" type="holotype">holotype</typeStatus>
): 0.16 (MTrL 1–4: 0.05; MTrL 5: 0.11); AW: 0.69–0.87; AedL: 0.42–0.50; BL: 2.08–3.08 (
<typeStatus id="F64E8839FFFD6D4CD487DD56BD320238" box="[717,821,1121,1147]" pageId="20" pageNumber="271" type="holotype">holotype</typeStatus>
: 2.56).
</paragraph>
<paragraph id="294A369BFFFD6D4CD68DDDB2BA62000B" blockId="20.[151,1437,802,2034]" pageId="20" pageNumber="271">
Habitus as in
<figureCitation id="B1CE2A1EFFFD6D4CD72CDDB2BFAE02DC" box="[358,425,1157,1183]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="20" pageNumber="271">
Fig.
<quantity id="EE0D9B7EFFFD6D4CD7D1DDB2BFAE02DC" box="[411,425,1157,1183]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="271" unit="in" value="2.0">2</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFFD6D4CD79ADDB2BC8E02DC" author="Shavrin, A. V." box="[464,649,1157,1183]" pageId="20" pageNumber="271" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
and in
<figureCitation id="B1CE2A1EFFFD6D4CD4A8DDB2BD3102DC" box="[738,822,1157,1183]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="20" pageNumber="271">Fig. 39</figureCitation>
. Body brown, with paler lateroapical portion of head, lateral and basal portion of elytra (some specimens with uniformly paler elytra in contrast to the body), paratergites and apical tergites of abdomen in some specimens (some specimens with slightly darker abdominal tergites V–VII in middle); antennomeres 6–11 yellow-brown to brown; mouthparts, antennomeres 1–5 and legs yellowish (two preapical segments of maxillary palpomeres distintly darker in some specimens). Head (including neck), pronotum and scutellum without punctures; elytra with moderately fine and dense punctation, sometimes finer along suture and indistinct in apical portions; abdomen with very indistinct dense punctation. Head with regular and very strong coarse microsculpture, sometimes slightly finer in middle and coarser on infraorbital portions; neck with fine or relatively coarse and dense isodiametric or transverse meshes; microsculpture of pronotum about as that in middle portion of head or slightly stronger and coarser; scutellum with dense and coarse transverse or isodiametric meshes; microsculpture of elytra isodiametric and very dense, slightly finer and sometimes larger than that in pronotum, usually denser and finer in parascutellar portion and along suture; abdomen with very dense isodiametric microsculpture, distinctly transverse in abdominal tergites VI–VIII in some specimens.
</paragraph>
<paragraph id="294A369BFFFD6D4CD68DDF65BF370127" blockId="20.[151,1437,802,2034]" pageId="20" pageNumber="271">Head 1.4–1.7 times as broad as long, with moderately wide and deep anteocellar foveae, sublinear or slightly diverging posteriad, about as long as diameter of ocellus or slightly longer, reaching level of posterior third or about middle length of eyes; postocular carina absent or very indistinct, obtuse. Ocelli moderately large, located about at level of postocular ridges; distance between ocelli about as long as or slightly greater than distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about twice as long as preapical segment, or slightly shorter. Antennomere 4 distinctly longer than broad, 5 slightly broader than 4, 6 distinctly broader than 5, 7 slightly broader and longer than 6, 8–9 slightly shorter than 7, 10 slightly longer than 9, apical antennomere about 1.2 times as long as 10.</paragraph>
<paragraph id="294A369BFFFD6D4CD68DDE59BF2201B1" blockId="20.[151,1437,802,2034]" pageId="20" pageNumber="271">Pronotum 1.6–1.7 times as broad as long, 1.2–1.3 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; posterior angles subacute or obtuse; middle portion with slightly impressed medioapical portion in some specimens and with two indistinct to distinct wide and moderately long paramedian impressions.</paragraph>
<caption id="7D8A6613FFFC6D4DD6DDDD94BA77029C" ID-DOI="http://doi.org/10.5281/zenodo.10821243" ID-Zenodo-Dep="10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="21" pageNumber="272" startId="21.[151,264,1187,1211]" targetBox="[159,1428,187,1164]" targetPageId="21" targetType="figure">
<paragraph id="294A369BFFFC6D4DD6DDDD94BA77029C" blockId="21.[151,1436,1187,1248]" pageId="21" pageNumber="272">
<emphasis id="1B81EA89FFFC6D4DD6DDDD94BF5F02F8" bold="true" box="[151,344,1187,1211]" pageId="21" pageNumber="272">FIGURES 53–56.</emphasis>
<taxonomicName id="EEF54D18FFFC6D4DD72BDD93BC6A02F8" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[353,621,1188,1212]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="272" phylum="Arthropoda" rank="species" species="coriacea">
<emphasis id="1B81EA89FFFC6D4DD72BDD93BC6A02F8" box="[353,621,1188,1212]" italics="true" pageId="21" pageNumber="272">Paraphloeostiba coriacea</emphasis>
</taxonomicName>
: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFFC6D4DD68DDC05BEFE0333" blockId="21.[151,1437,1330,2040]" pageId="21" pageNumber="272">Elytra 1.1–1.3 times as broad as long, 1.8 times to twice as long as pronotum, with narrowly impressed lateral margins.</paragraph>
<paragraph id="294A369BFFFC6D4DD68DDC4DBC9103D7" blockId="21.[151,1437,1330,2040]" box="[199,662,1402,1428]" pageId="21" pageNumber="272">Abdomen distinctly narrower than elytra.</paragraph>
<paragraph id="294A369BFFFC6D4DD68DDCA9BD0100D3" blockId="21.[151,1437,1330,2040]" pageId="21" pageNumber="272">
Male. Posterior margin of abdominal tergite VIII truncate, as in
<figureCitation id="B1CE2A1EFFFC6D4DD5C5DCA9BDD903FB" box="[911,990,1438,1464]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="21" pageNumber="272">
Fig.
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in
<bibRefCitation id="4D644B6AFFFC6D4DD24ADCA9BABD03FB" author="Shavrin, A. V." box="[1024,1210,1438,1464]" pageId="21" pageNumber="272" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Posterior margin of abdominal sternite VIII slightly concave, as in
<figureCitation id="B1CE2A1EFFFC6D4DD4E8DCF5BCF6039F" box="[674,753,1474,1500]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="21" pageNumber="272">
Fig.
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in
<bibRefCitation id="4D644B6AFFFC6D4DD55EDCF5BDCA039F" author="Shavrin, A. V." box="[788,973,1474,1500]" pageId="21" pageNumber="272" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Aedeagus with wide basal bulb, strongly narrowed toward moderately small truncate apex; parameres distinctly exceeding apex of median lobe, gradually widened apically, with two long and two moderately short apical setae, and with a row of 7–8 of moderately short setae on inner margins and one short seta on outer margins of each paramere; internal sac narrow and moderately short, with several sclerotized elongate stuctures in basal portion (see
<figureCitation id="B1CE2A1EFFFC6D4DD5D5DF65BDEA002F" box="[927,1005,1618,1644]" captionStart="FIGURE 11" captionStartId="5.[151,250,1633,1657]" captionTargetBox="[151,1435,525,1610]" captionTargetId="figure-143@5.[151,1436,525,1610]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of Nipponophloeostiba: N. verrucifera (circles), N. thayeri (triangle), N. longilobata (square), N. explanata (diamond), N. hornabrooki (star)." figureDoi="http://doi.org/10.5281/zenodo.10821220" httpUri="https://zenodo.org/record/10821220/files/figure.png" pageId="21" pageNumber="272">
Fig.
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</figureCitation>
in
<bibRefCitation id="4D644B6AFFFC6D4DD245DF65BAB4002F" author="Shavrin, A. V." box="[1039,1203,1618,1644]" pageId="21" pageNumber="272" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin 2023b</bibRefCitation>
and
<figureCitation id="B1CE2A1EFFFC6D4DD2A2DF65BB3E002E" box="[1256,1337,1618,1645]" captionStart="FIGURES 53–56" captionStartId="21.[151,264,1187,1211]" captionTargetBox="[159,1428,187,1164]" captionTargetId="figure-16@21.[151,1436,181,1164]" captionTargetPageId="21" captionText="FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="21" pageNumber="272">Fig. 53</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFFC6D4DD79EDF41BC2300D3" box="[468,548,1654,1680]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="21" pageNumber="272">
Fig.
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</figureCitation>
in
<bibRefCitation id="4D644B6AFFFC6D4DD402DF41BD0500D3" author="Shavrin, A. V." box="[584,770,1654,1680]" pageId="21" pageNumber="272" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
.
</paragraph>
<paragraph id="294A369BFFFC6D4DD68DDFADBD9D009B" blockId="21.[151,1437,1330,2040]" pageId="21" pageNumber="272">
Female. Posterior margins of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFFC6D4DD562DFADBD7C00F6" box="[808,891,1690,1717]" captionStart="FIGURES 53–56" captionStartId="21.[151,264,1187,1211]" captionTargetBox="[159,1428,187,1164]" captionTargetId="figure-16@21.[151,1436,181,1164]" captionTargetPageId="21" captionText="FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="21" pageNumber="272">Fig. 54</figureCitation>
) and abdominal sternite VIII (
<figureCitation id="B1CE2A1EFFFC6D4DD29DDFADBB2D00F6" box="[1239,1322,1690,1717]" captionStart="FIGURES 53–56" captionStartId="21.[151,264,1187,1211]" captionTargetBox="[159,1428,187,1164]" captionTargetId="figure-16@21.[151,1436,181,1164]" captionTargetPageId="21" captionText="FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="21" pageNumber="272">Fig. 55</figureCitation>
) truncate. Female accessory sclerite as in
<figureCitation id="B1CE2A1EFFFC6D4DD7BDDF89BC4D009A" box="[503,586,1726,1753]" captionStart="FIGURES 53–56" captionStartId="21.[151,264,1187,1211]" captionTargetBox="[159,1428,187,1164]" captionTargetId="figure-16@21.[151,1436,181,1164]" captionTargetPageId="21" captionText="FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="21" pageNumber="272">Fig. 56</figureCitation>
. Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFFC6D4DD68DDFD6BD1500BF" blockId="21.[151,1437,1330,2040]" box="[199,786,1761,1788]" pageId="21" pageNumber="272">
<emphasis id="1B81EA89FFFC6D4DD68DDFD6BFB600BF" bold="true" box="[199,433,1761,1788]" pageId="21" pageNumber="272">Comparative notes.</emphasis>
See details in
<bibRefCitation id="4D644B6AFFFC6D4DD41EDFD5BD0900BF" author="Shavrin, A. V." box="[596,782,1762,1788]" pageId="21" pageNumber="272" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFC6D4DD68DDE32BFEC01CF" pageId="21" pageNumber="272" type="distribution">
<paragraph id="294A369BFFFC6D4DD68DDE32BFA10107" blockId="21.[151,1437,1330,2040]" pageId="21" pageNumber="272">
<emphasis id="1B81EA89FFFC6D4DD68DDE32BF66015C" bold="true" box="[199,353,1797,1823]" pageId="21" pageNumber="272">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFFC6D4DD722DE31BC8A0163" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[360,653,1798,1824]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="272" phylum="Arthropoda" rank="species" species="coriacea">
<emphasis id="1B81EA89FFFC6D4DD722DE31BC8A0163" box="[360,653,1798,1824]" italics="true" pageId="21" pageNumber="272">Paraphloeostiba coriacea</emphasis>
</taxonomicName>
is known from several localities in Western New
<collectingCountry id="51E2760BFFFC6D4DD28EDE31BB120163" box="[1220,1301,1798,1824]" name="Guinea-Bissau" pageId="21" pageNumber="272">Guinea</collectingCountry>
(
<collectingCountry id="51E2760BFFFC6D4DD36EDE31BB930163" box="[1316,1428,1798,1824]" name="Indonesia" pageId="21" pageNumber="272">Indonesia</collectingCountry>
) and
<collectingCountry id="51E2760BFFFC6D4DD68DDE1DBFA40107" box="[199,419,1834,1860]" name="Papua New Guinea" pageId="21" pageNumber="272">Papua New Guinea</collectingCountry>
.
</paragraph>
<paragraph id="294A369BFFFC6D4DD68DDE7ABFEC01CF" blockId="21.[151,1437,1330,2040]" pageId="21" pageNumber="272">
<emphasis id="1B81EA89FFFC6D4DD68DDE7ABF4C0124" bold="true" box="[199,331,1869,1895]" pageId="21" pageNumber="272">Bionomics.</emphasis>
The species was collected at elevations from 1500 (
<typeStatus id="F64E8839FFFC6D4DD58DDE79BA2B012B" box="[967,1068,1870,1896]" pageId="21" pageNumber="272" type="holotype">holotype</typeStatus>
) to
<quantity id="EE0D9B7EFFFC6D4DD229DE79BAB9012B" box="[1123,1214,1870,1897]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="272" unit="m" value="2500.0">2500 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFC6D4DD68DDEA1BC3301BB" pageId="21" pageNumber="272" type="discussion">
<paragraph id="294A369BFFFC6D4DD68DDEA1BC3301BB" blockId="21.[151,1437,1330,2040]" pageId="21" pageNumber="272">
<emphasis id="1B81EA89FFFC6D4DD68DDEA1BF3C01F3" bold="true" box="[199,315,1942,1968]" pageId="21" pageNumber="272">Remarks.</emphasis>
Based on the study of the
<typeStatus id="F64E8839FFFC6D4DD415DEA1BCC301F3" box="[607,708,1942,1968]" pageId="21" pageNumber="272" type="holotype">holotype</typeStatus>
, this species was redescribed by
<bibRefCitation id="4D644B6AFFFC6D4DD27ADEA1BAEE01F3" author="Shavrin, A. V." box="[1072,1257,1942,1968]" pageId="21" pageNumber="272" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. I provided here the redescription this species, including the description of the female. It is here recorded from the Western New
<collectingCountry id="51E2760BFFFC6D4DD6DDDEE9BEEF01BB" box="[151,232,2014,2040]" name="Guinea-Bissau" pageId="21" pageNumber="272">Guinea</collectingCountry>
(
<collectingCountry id="51E2760BFFFC6D4DD6BDDEE9BF6001BB" box="[247,359,2014,2040]" name="Indonesia" pageId="21" pageNumber="272">Indonesia</collectingCountry>
) for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

276
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<emphasis id="1B81EA89FFFF6D4ED6DDDA0FBC310510" bold="true" box="[151,566,824,851]" italics="true" pageId="22" pageNumber="273">Paraphloeostiba guadalcanalensis</emphasis>
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(
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,
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)
</paragraph>
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<paragraph id="294A369BFFFF6D4ED6DDDA92BB5D0244" blockId="22.[151,1437,932,2003]" pageId="22" pageNumber="273">
<emphasis id="1B81EA89FFFF6D4ED6DDDA92BF4005FD" bold="true" box="[151,327,932,959]" pageId="22" pageNumber="273">Type material.</emphasis>
<typeStatus id="F64E8839FFFF6D4ED71ADA93BFBF05FD" box="[336,440,932,958]" pageId="22" pageNumber="273" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFFF6D4ED78BDA92BFDE05FC" box="[449,473,933,959]" count="1" pageId="22" pageNumber="273" type="male">♂</specimenCount>
: ‘SOLOMON IS. |
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| Popamanasiu [handwritten] | 5500`[handwritten] | 25 11 [handwritten] 1965 [the last number “5” is handwritten] | 20217 [handwritten] | P. Greenslade’ &lt;printed&gt;, ‘
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|
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<emphasis id="1B81EA89FFFF6D4ED702DADBBC040245" box="[328,515,1004,1030]" italics="true" pageId="22" pageNumber="273">Paraphloeostiba</emphasis>
</taxonomicName>
|
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<emphasis id="1B81EA89FFFF6D4ED45CDADBBCDD0245" box="[534,730,1004,1030]" italics="true" pageId="22" pageNumber="273">guadalcanalensis</emphasis>
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<taxonomicNameLabel id="00B257F2FFFF6D4ED4ABDADABD260244" box="[737,801,1005,1031]" pageId="22" pageNumber="273" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFFF6D4ED5E0DADABDC70244" box="[938,960,1005,1031]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="273" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFFF6D4ED2B9DADABB4A0244" box="[1267,1357,1005,1031]" pageId="22" pageNumber="273">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFFF6D4ED68DDD26BD170230" blockId="22.[151,1437,932,2003]" pageId="22" pageNumber="273">
<typeStatus id="F64E8839FFFF6D4ED68DDD26BF3F0268" box="[199,312,1041,1067]" pageId="22" pageNumber="273" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFFF6D4ED775DD26BF7C0268" box="[319,379,1041,1067]" count="7" pageId="22" pageNumber="273" type="male">7 ♂♂</specimenCount>
(
<specimenCount id="3FF3FD12FFFF6D4ED7CDDD26BC190269" box="[391,542,1040,1067]" count="1" pageId="22" pageNumber="273" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="3FF3FD12FFFF6D4ED4D6DD26BCE20268" box="[668,741,1041,1067]" count="11" pageId="22" pageNumber="273" type="female">11 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFFF6D4ED4BADD26BD8F0269" box="[752,904,1040,1067]" count="1" pageId="22" pageNumber="273" type="generic">one specimen</specimenCount>
dissected):‘SOLOMON IS. |
<collectingRegion id="EB31F879FFFF6D4ED281DD27BB500269" box="[1227,1367,1040,1066]" country="Solomon Islands" name="Guadalcanal" pageId="22" pageNumber="273">Guadalcanal</collectingRegion>
Popa- | manasiu 5500`|
<date id="5D4B105BFFFF6D4ED714DD03BFDC020C" box="[350,475,1076,1103]" pageId="22" pageNumber="273" value="1965-11-25">25.11.1965</date>
20217 | P. Greenslade’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFFF6D4ED52FDD02BDF2020C" box="[869,1013,1077,1103]" pageId="22" pageNumber="273" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFFF6D4ED243DD03BAC3020D" authorityName="Steel" authorityYear="1960" box="[1033,1220,1076,1102]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFFF6D4ED243DD03BAC3020D" box="[1033,1220,1076,1102]" italics="true" pageId="22" pageNumber="273">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFFF6D4ED292DD03BB9B020D" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1240,1436,1076,1102]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. nov.">
<emphasis id="1B81EA89FFFF6D4ED292DD03BB9B020D" box="[1240,1436,1076,1102]" italics="true" pageId="22" pageNumber="273">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFFF6D4ED6DDDD6EBED00230" bold="true" box="[151,215,1113,1139]" pageId="22" pageNumber="273">
<taxonomicNameLabel id="00B257F2FFFF6D4ED6DDDD6EBED00230" box="[151,215,1113,1139]" pageId="22" pageNumber="273" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFFF6D4ED72ADD6EBF710230" box="[352,374,1113,1139]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="273" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFFF6D4ED4E3DD6EBD040230" box="[681,771,1113,1139]" pageId="22" pageNumber="273">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFF6D40D68DDD4BBFF90556" lastPageId="24" lastPageNumber="275" pageId="22" pageNumber="273" type="description">
<paragraph id="294A369BFFFF6D4ED68DDD4BBF8C0340" blockId="22.[151,1437,932,2003]" pageId="22" pageNumber="273">
<emphasis id="1B81EA89FFFF6D4ED68DDD4BBF5E02D5" bold="true" box="[199,345,1148,1174]" pageId="22" pageNumber="273">Description.</emphasis>
Measurements (n=19): HW: 0.44–0.53; HL: 0.27–0.35; OL: 0.15–0.17; TL: 0.05–0.06; AL (
<typeStatus id="F64E8839FFFF6D4ED6D5DD97BF0F02F9" box="[159,264,1184,1210]" pageId="22" pageNumber="273" type="holotype">holotype</typeStatus>
): 0.71; PL: 0.33–0.40; PWmax: 0.54–0.63; PWmin: 0.43–0.53; ESL: 0.64–0.77; EW: 0.78–0.89; MTbL (
<typeStatus id="F64E8839FFFF6D4ED6D5DDF3BF0F029D" box="[159,264,1220,1246]" pageId="22" pageNumber="273" type="holotype">holotype</typeStatus>
): 0.42; MTrL (
<typeStatus id="F64E8839FFFF6D4ED7E4DDF3BC10029D" box="[430,535,1220,1246]" pageId="22" pageNumber="273" type="holotype">holotype</typeStatus>
): 0.20 (MTrL 1–4: 0.07; MTrL 5: 0.13); AW: 0.76–0.88; AedL: 0.45; BL: 2.07– 3.07 (
<typeStatus id="F64E8839FFFF6D4ED69DDDDFBF380341" box="[215,319,1256,1282]" pageId="22" pageNumber="273" type="holotype">holotype</typeStatus>
: 2.32).
</paragraph>
<paragraph id="294A369BFFFF6D4ED68DDC3BBB1F00CD" blockId="22.[151,1437,932,2003]" pageId="22" pageNumber="273">
Habitus as in
<figureCitation id="B1CE2A1EFFFF6D4ED722DC3BBFBB0364" box="[360,444,1292,1319]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="22" pageNumber="273">Fig. 42</figureCitation>
. Body yellow-brown to reddish-brown, with slightly darker apical and lateral portions of head and abdominal tergites (lateral and basal portions of pronotum slightly paler); antennomeres 6–11 brown; femora and tibia yellow-brown; mouthparts, antennomeres 1–5 and tarsi yellowish. Head with very sparse fine punctation, but some
<typeStatus id="F64E8839FFFF6D4ED7DBDC4EBFFB03D0" box="[401,508,1401,1427]" pageId="22" pageNumber="273" type="paratype">paratypes</typeStatus>
without visible punctation in middle portion; neck without punctures; pronotum with moderately dense, large and deep punctures, with moderately wide impunctate band in middle; elytra with dense punctation, distinctly denser, larger and deeper than that in pronotum, finer around scutellum and along suture; abdominal tergites with distinct, fine and dense punctation, finer and sparser in middle (abdominal tergites of some specimens without visible punctures in middle). Head with very dense and coarse microsculpture, transverse in vertex and isodiametric in middle; pronotum with very dense isodiametric microreticulation, similar to that in middle portion of pronotum, sometimes finer in middle; microsculpture of elytra similar to that in pronotum, sometimes finer along suture; abdominal tergites with dense and coarse isodiametric microreticulation.
</paragraph>
<paragraph id="294A369BFFFF6D4ED68DDFAFBEE40190" blockId="22.[151,1437,932,2003]" pageId="22" pageNumber="273">Head with flattened middle and slightly elevated infraorbital portions, with moderately wide and deep anteriomedian depressions, 1.4–1.5 times as broad as long, with narrow and moderately deep anteocellar foveae, reaching about level of anterior third of eyes; postocular carina absent. Ocelli moderately small, located slightly below level of posterior margins of eyes; distance between ocellus 1.3–1.4 times as long as distance between ocellus and posterior margin of eyes. Apical maxillary palpomere very long, about three times as long as preapical segment, from basal portion gradually narrowed toward acute apex. Antenna reaching apical margin of elytra when reclined; antennomere 3 distinctly narrower and longer than 2, 4 small, slightly more than twice as long as 3, 5 indisitnctly broader than 4, 6 distinctly longer and significantly broader than 5, 7–9 slightly broader than 6, 10 slightly longer than 9.</paragraph>
<caption id="7D8A6613FFFE6D4FD6DDDF59BD08008E" ID-DOI="http://doi.org/10.5281/zenodo.10821245" ID-Zenodo-Dep="10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="23" pageNumber="274" startId="23.[151,264,1646,1670]" targetBox="[159,1428,189,1614]" targetPageId="23" targetType="figure">
<paragraph id="294A369BFFFE6D4FD6DDDF59BD08008E" blockId="23.[151,1436,1646,1742]" pageId="23" pageNumber="274">
<emphasis id="1B81EA89FFFE6D4FD6DDDF59BF5000C5" bold="true" box="[151,343,1646,1670]" pageId="23" pageNumber="274">FIGURES 57–63.</emphasis>
<taxonomicName id="EEF54D18FFFE6D4FD715DF59BCC500C5" authorityName="Shavrin" authorityYear="2024" box="[351,706,1646,1670]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="guadalcanalensis">
<emphasis id="1B81EA89FFFE6D4FD715DF59BCC500C5" box="[351,706,1646,1670]" italics="true" pageId="23" pageNumber="274">Paraphloeostiba guadalcanalensis</emphasis>
</taxonomicName>
(paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFFE6D4FD68DDE3ABEFC01F2" blockId="23.[151,1436,1804,2038]" pageId="23" pageNumber="274">Pronotum 1.5–1.6 times as broad as long and 1.1–1.2 times as broad as head, widest slightly above middle, strongly narrowed posteriad, with slightly concave laterobasal margins in front of obtuse hind angles; middle portion with wide and shallow or moderately deep longitudinal impressions, widened basally, with narrow, rounded and slightly elevated ridges in lateroapical portions; lateral portions distinctly depressed, broader in laterobasal portions.</paragraph>
<paragraph id="294A369BFFFE6D4FD68DDE8EBC0401B5" blockId="23.[151,1436,1804,2038]" pageId="23" pageNumber="274">Elytra 1.1–1.2 times as broad as long, 1.9 times as long as pronotum, distinctly broadened posteriad and narrowly impressed along lateral margins.</paragraph>
<paragraph id="294A369BFFF16D40D68DD9A0BC8506F2" blockId="24.[151,1437,151,1005]" box="[199,642,151,177]" pageId="24" pageNumber="275">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="294A369BFFF16D40D68DD98CBC2C07CA" blockId="24.[151,1437,151,1005]" pageId="24" pageNumber="275">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFF16D40D51ED98CBDA00696" box="[852,935,186,213]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="24" pageNumber="275">Fig. 59</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFF16D40D7AAD9E8BC3106BA" box="[480,566,223,249]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="24" pageNumber="275">Fig. 60</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward moderately narrow median lobe with small rounded apex; parameres slightly not reaching apex of median lobe, distinctly broadened apicad, with three long apical setae and several moderately long setae on inner margin of each paramere; internal sac long, narrow, with several strongly sclerotized elongate structures in basal portion (
<figureCitation id="B1CE2A1EFFF16D40D2ABD87CBB300726" box="[1249,1335,330,357]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="24" pageNumber="275">Fig. 57</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFF16D40D79ED858BC2007CA" box="[468,551,366,393]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="24" pageNumber="275">Fig. 58</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFF16D40D68DD8A4BAC10792" blockId="24.[151,1437,151,1005]" pageId="24" pageNumber="275">
Female. Posterior margin of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFF16D40D569D8A4BD7007EE" box="[803,887,403,429]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="24" pageNumber="275">Fig. 61</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFF16D40D21DD8A4BAAC07EE" box="[1111,1195,403,429]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="24" pageNumber="275">Fig. 62</figureCitation>
) rounded. Accessory sclerite with wide basal and narrow long apical portions (
<figureCitation id="B1CE2A1EFFF16D40D553D880BD6A0792" box="[793,877,439,465]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="24" pageNumber="275">Fig. 63</figureCitation>
). Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFF16D40D68DD8EDBFF90556" blockId="24.[151,1437,151,1005]" pageId="24" pageNumber="275">
<emphasis id="1B81EA89FFF16D40D68DD8EDBFB707B6" bold="true" box="[199,432,474,501]" pageId="24" pageNumber="275">Comparative notes.</emphasis>
Based on the general structure of the aedeagus,
<taxonomicName id="EEF54D18FFF16D40D581D8ECBAA907B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[971,1198,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFF16D40D581D8ECBDDB07B7" box="[971,988,475,500]" italics="true" pageId="24" pageNumber="275">P</emphasis>
.
<emphasis id="1B81EA89FFF16D40D5A0D8ECBAA907B6" box="[1002,1198,475,501]" italics="true" pageId="24" pageNumber="275">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D2FED8ECBAF407B6" bold="true" box="[1204,1267,475,501]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D2FED8ECBAF407B6" box="[1204,1267,475,501]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFF16D40D3CED8ECBEDC045B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFF16D40D3CED8ECBB9207B7" box="[1412,1429,475,500]" italics="true" pageId="24" pageNumber="275">P</emphasis>
.
<emphasis id="1B81EA89FFF16D40D6DDD8C8BEDC045B" box="[151,219,511,536]" italics="true" pageId="24" pageNumber="275">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D6A8D8C8BF27045A" bold="true" box="[226,288,511,537]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D6A8D8C8BF27045A" box="[226,288,511,537]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFF16D40D766D8C8BF97045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[300,400,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFF16D40D766D8C8BF97045A" box="[300,400,511,537]" italics="true" pageId="24" pageNumber="275">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D7DCD8C8BFD3045A" bold="true" box="[406,468,511,537]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D7DCD8C8BFD3045A" box="[406,468,511,537]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFF16D40D442D8C8BC97045B" authorityName="Shavrin" authorityYear="2023" box="[520,656,511,536]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF16D40D442D8C8BC97045B" box="[520,656,511,536]" italics="true" pageId="24" pageNumber="275">P. conjuncta</emphasis>
</taxonomicName>
. Based on the presence of two distinct longitudinal impressions on the pronotum, it is more similar to
<taxonomicName id="EEF54D18FFF16D40D7A6DB14BC49047F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[492,590,547,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFF16D40D7A6DB14BFFA047F" box="[492,509,547,572]" italics="true" pageId="24" pageNumber="275">P</emphasis>
.
<emphasis id="1B81EA89FFF16D40D440DB14BC49047F" box="[522,590,547,572]" italics="true" pageId="24" pageNumber="275">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D41EDB14BC94047E" bold="true" box="[596,659,547,573]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D41EDB14BC94047E" box="[596,659,547,573]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the larger body, slightly shorter parameres with distinctly broader apical portions. It can be distinguished from
<taxonomicName id="EEF54D18FFF16D40D24EDB70BA8A0423" authorityName="Shavrin" authorityYear="2023" box="[1028,1165,583,608]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF16D40D24EDB70BA8A0423" box="[1028,1165,583,608]" italics="true" pageId="24" pageNumber="275">P. conjuncta</emphasis>
</taxonomicName>
by the narrower median lobe and broader apical portions of the shorter parameres, and from
<taxonomicName id="EEF54D18FFF16D40D5D2DB5CBDFA04C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[920,1021,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFF16D40D5D2DB5CBDFA04C6" box="[920,1021,619,645]" italics="true" pageId="24" pageNumber="275">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D24EDB5CBA4304C6" bold="true" box="[1028,1092,619,645]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D24EDB5CBA4304C6" box="[1028,1092,619,645]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the broader apical portions of the aedeagus. Additionally, it can be distinguished from
<taxonomicName id="EEF54D18FFF16D40D509DBB8BDC804EB" authorityName="Shavrin" authorityYear="2023" box="[835,975,655,680]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFF16D40D509DBB8BDC804EB" box="[835,975,655,680]" italics="true" pageId="24" pageNumber="275">P. conjuncta</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF54D18FFF16D40D241DBB8BA7404EA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1035,1139,655,681]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFF16D40D241DBB8BA7404EA" box="[1035,1139,655,681]" italics="true" pageId="24" pageNumber="275">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D236DBB8BAB804EA" bold="true" box="[1148,1215,655,681]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D236DBB8BAB804EA" box="[1148,1215,655,681]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the presence of longitudinal impressions and fine punctation of the head. From all these species it differs by the concave laterobasal margins of the pronotum in front of hind angles, details of the morphology of the aedeagus, and the different shape of the female accessory sclerite.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF16D40D68DDA29BFAA051E" pageId="24" pageNumber="275" type="distribution">
<paragraph id="294A369BFFF16D40D68DDA29BFAA051E" blockId="24.[151,1437,151,1005]" pageId="24" pageNumber="275">
<emphasis id="1B81EA89FFF16D40D68DDA29BF66057B" bold="true" box="[199,353,798,824]" pageId="24" pageNumber="275">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFF16D40D723DA28BCF6057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[361,753,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. nov.">
<emphasis id="1B81EA89FFF16D40D723DA28BCF6057A" box="[361,753,799,825]" italics="true" pageId="24" pageNumber="275">Paraphloeostiba guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF16D40D4B0DA28BD3B057A" bold="true" box="[762,828,799,825]" pageId="24" pageNumber="275">
<taxonomicNameLabel id="00B257F2FFF16D40D4B0DA28BD3B057A" box="[762,828,799,825]" pageId="24" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFF16D40D211DA28BA8C057A" box="[1115,1163,799,825]" pageId="24" pageNumber="275">type</typeStatus>
locality in
<collectingRegion id="EB31F879FFF16D40D35ADA28BB9B057A" box="[1296,1436,799,825]" country="Solomon Islands" name="Guadalcanal" pageId="24" pageNumber="275">Guadalcanal</collectingRegion>
Island,
<collectingCountry id="51E2760BFFF16D40D6A0DA74BFAE051E" box="[234,425,835,861]" name="Solomon Islands" pageId="24" pageNumber="275">Solomon Islands</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF16D40D68DDA50BF0005AE" pageId="24" pageNumber="275" type="etymology">
<paragraph id="294A369BFFF16D40D68DDA50BF0305E6" blockId="24.[151,1437,151,1005]" pageId="24" pageNumber="275">
<emphasis id="1B81EA89FFF16D40D68DDA50BF4905C2" bold="true" box="[199,334,871,897]" pageId="24" pageNumber="275">Etymology.</emphasis>
The specific epithet derived from the name of the island where the
<typeStatus id="F64E8839FFF16D40D2EEDA50BAD305C2" box="[1188,1236,871,897]" pageId="24" pageNumber="275">type</typeStatus>
specimens were collected.
</paragraph>
<paragraph id="294A369BFFF16D40D68DDA99BF0005AE" blockId="24.[151,1437,151,1005]" pageId="24" pageNumber="275">
<emphasis id="1B81EA89FFF16D40D68DDA99BF4C058B" bold="true" box="[199,331,942,968]" pageId="24" pageNumber="275">Bionomics.</emphasis>
Specimens were collected at an elevation
<quantity id="EE0D9B7EFFF16D40D514DA98BDBC058A" box="[862,955,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.6759999999999997" pageId="24" pageNumber="275" unit="m" value="1676.0">1676 m</quantity>
a.s.l. The detailed bionomical data ara unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

283
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<mods:title id="F48892D8571B779E10D372A2CAEAE610">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<paragraph id="294A369BFFFF6D4ED6DDD9A0BCE706F2" blockId="22.[151,736,151,214]" box="[151,736,151,178]" pageId="22" pageNumber="273">
<heading id="720281F7FFFF6D4ED6DDD9A0BCE706F2" bold="true" box="[151,736,151,178]" fontSize="11" level="1" pageId="22" pageNumber="273" reason="1">
<taxonomicName id="EEF54D18FFFF6D4ED6DDD9A0BCE706F2" ID-CoL="6TQXN" authority="(MacLeay, 1873)" baseAuthorityName="MacLeay" baseAuthorityYear="1873" box="[151,736,151,178]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="gayndahensis">
<emphasis id="1B81EA89FFFF6D4ED6DDD9A0BCE706F2" bold="true" box="[151,736,151,178]" pageId="22" pageNumber="273">
<emphasis id="1B81EA89FFFF6D4ED6DDD9A0BC0E06F1" bold="true" box="[151,521,151,178]" italics="true" pageId="22" pageNumber="273">Paraphloeostiba gayndahensis</emphasis>
(
<bibRefCitation id="4D644B6AFFFF6D4ED452D9A0BCDF06F2" author="MacLeay, W. J." box="[536,728,151,177]" pageId="22" pageNumber="273" pagination="79 - 205" refId="ref35618" refString="MacLeay, W. J. (1873) Notes on a collection of insects from Gayndah. The Transactions of the Entomological Society of New South Wales, 2, 79 - 205." type="journal article" year="1873">MacLeay, 1873</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFF6D4ED6DDD98CBEFE0695" box="[151,249,187,214]" pageId="22" pageNumber="273" type="description">
<paragraph id="294A369BFFFF6D4ED6DDD98CBEFE0695" blockId="22.[151,736,151,214]" box="[151,249,187,214]" pageId="22" pageNumber="273">
(
<figureCitation id="B1CE2A1EFFFF6D4ED6D5D98CBEF60695" box="[159,241,187,214]" captionStart="FIGURES 45" captionStartId="18.[151,264,885,909]" captionTargetBox="[151,1436,193,862]" captionTargetId="figure-22@18.[151,1436,181,862]" captionTargetPageId="18" captionText="FIGURES 45. Distribution of Paraphloeostiba coriacea (circles) and P. gayndahensis (squares) in New Guinea." figureDoi="http://doi.org/10.5281/zenodo.10821237" httpUri="https://zenodo.org/record/10821237/files/figure.png" pageId="22" pageNumber="273">Fig. 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFF6D4ED6DDD837BCDA0775" pageId="22" pageNumber="273" type="reference_group">
<paragraph id="294A369BFFFF6D4ED6DDD837BC8D0754" blockId="22.[151,733,255,310]" box="[151,650,255,280]" pageId="22" pageNumber="273">
<treatmentCitationGroup id="09E511B5FFFF6D4ED6DDD837BC8D0754" box="[151,650,255,280]" pageId="22" pageNumber="273">
<treatmentCitation id="A854108AFFFF6D4ED6DDD837BC8D0754" author="MacLeay, W. J." box="[151,650,255,280]" page="151" pageId="22" pageNumber="273" year="1873">
<taxonomicName id="EEF54D18FFFF6D4ED6DDD837BC8D0754" ID-CoL="74KKF" authority="W. J. MacLeay, 1873: 151" authorityName="W. J. MacLeay" authorityPageNumber="151" authorityYear="1873" box="[151,650,255,280]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="gayndahense">
<emphasis id="1B81EA89FFFF6D4ED6DDD837BF87075B" box="[151,384,256,280]" italics="true" pageId="22" pageNumber="273">Omalium gayndahense</emphasis>
W.J.
<bibRefCitation id="4D644B6AFFFF6D4ED7F3D837BC8D0754" author="MacLeay, W. J." box="[441,650,255,280]" pageId="22" pageNumber="273" pagination="79 - 205" refId="ref35618" refString="MacLeay, W. J. (1873) Notes on a collection of insects from Gayndah. The Transactions of the Entomological Society of New South Wales, 2, 79 - 205." type="journal article" year="1873">MacLeay, 1873: 151</bibRefCitation>
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<paragraph id="294A369BFFFF6D4ED6DDD829BCDA0775" blockId="22.[151,733,255,310]" box="[151,733,286,310]" pageId="22" pageNumber="273">Additional references see in Shavrin (2020, 2022, 2023b)</paragraph>
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<emphasis id="1B81EA89FFFF6D4ED6DDD853BCE0073C" bold="true" box="[151,743,356,383]" pageId="22" pageNumber="273">
Material examined.
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<collectingCountry id="51E2760BFFFF6D4ED7C3D853BC2B073D" box="[393,556,356,382]" name="Indonesia" pageId="22" pageNumber="273">INDONESIA</collectingCountry>
:
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<collectionCode id="4FE4AE5EFFFF6D4ED4FAD852BCE0073C" box="[688,743,357,383]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="22" pageNumber="273">NEW</collectionCode>
</materialsCitation>
</emphasis>
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<emphasis id="1B81EA89FFFF6D4ED4A4D853BD40073D" bold="true" box="[750,839,356,382]" pageId="22" pageNumber="273">
<collectingCountry id="51E2760BFFFF6D4ED4A4D853BD40073D" box="[750,839,356,382]" name="Guinea-Bissau" pageId="22" pageNumber="273">GUINEA</collectingCountry>
</emphasis>
:
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,
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: ‘
<collectionCode id="4FE4AE5EFFFF6D4ED584D852BA15073C" box="[974,1042,357,383]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="22" pageNumber="273">NEW</collectionCode>
</materialsCitation>
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<collectingCountry id="51E2760BFFFF6D4ED250D852BA83073C" box="[1050,1156,357,383]" name="Guinea-Bissau" pageId="22" pageNumber="273">GUINEA</collectingCountry>
<location id="2C2A6040FFFF6D4ED2C6D853BADF073D" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED2C6D853BADF073D" box="[1164,1240,356,382]" country="Guinea-Bissau" name="Okapa" pageId="22" pageNumber="273">Okapa</location>
,
<date id="5D4B105BFFFF6D4ED2A9D853BB6E073C" box="[1251,1385,356,383]" pageId="22" pageNumber="273" value="1965-03-26">
<collectingDate id="4D0FE9B3FFFF6D4ED2A9D853BB6E073C" box="[1251,1385,356,383]" pageId="22" pageNumber="273" value="1965-03-26">26.III.1965</collectingDate>
</date>
’, ‘
<collectionCode id="4FE4AE5EFFFF6D4ED335D852BB9F073C" box="[1407,1432,357,383]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="273">R</collectionCode>
</materialsCitation>
.
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<location id="2C2A6040FFFF6D4ED6DDD8BFBF1907E1" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED6DDD8BFBF1907E1" box="[151,286,392,418]" country="Guinea-Bissau" name="Hornabrook" pageId="22" pageNumber="273">Hornabrook</location>
<specimenCode id="79539EE0FFFF6D4ED76CD8BEBFCC07E0" box="[294,459,393,419]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="22" pageNumber="273">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFFF6D4ED7A9D8BEBC3A07E0" box="[483,573,393,419]" pageId="22" pageNumber="273">BMNH</collectionCode>
);
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,
<specimenCount id="3FF3FD12FFFF6D4ED4C6D8BEBCC907E0" box="[652,718,393,419]" count="3" pageId="22" pageNumber="273" type="female">3 ♀♀</specimenCount>
: ‘
<collectionCode id="4FE4AE5EFFFF6D4ED495D8BEBD2407E0" box="[735,803,393,419]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="22" pageNumber="273">NEW</collectionCode>
</materialsCitation>
<materialsCitation id="999D3CC6FFFF6D4ED566D8BEBB9F07E0" box="[812,1432,392,419]" collectingDate="1965-03-28" collectionCode="R" country="Guinea-Bissau" location="Okapa" pageId="22" pageNumber="273" specimenCount="1">
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<location id="2C2A6040FFFF6D4ED5D7D8BFBDEE07E1" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED5D7D8BFBDEE07E1" box="[925,1001,392,418]" country="Guinea-Bissau" name="Okapa" pageId="22" pageNumber="273">Okapa</location>
,
<date id="5D4B105BFFFF6D4ED5BFD8BFBA7107E0" box="[1013,1142,392,419]" pageId="22" pageNumber="273" value="1965-03-28">
<collectingDate id="4D0FE9B3FFFF6D4ED5BFD8BFBA7107E0" box="[1013,1142,392,419]" pageId="22" pageNumber="273" value="1965-03-28">28.III.1965</collectingDate>
</date>
, from dead branches’, ‘
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</materialsCitation>
.
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16.
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.1965, inside husk of nut (?of
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.
<materialsCitation id="999D3CC6FFFF6D4ED6DDD8E7BCE507A8" box="[151,738,464,491]" collectionCode="BMNH, NEW" country="Guinea-Bissau" location="Hornabrook" pageId="22" pageNumber="273" specimenCode="BM 1970-232" specimenCount="6" specimenCount-female="6">
<location id="2C2A6040FFFF6D4ED6DDD8E7BF1907A9" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED6DDD8E7BF1907A9" box="[151,286,464,490]" country="Guinea-Bissau" name="Hornabrook" pageId="22" pageNumber="273">Hornabrook</location>
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.’ (
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);
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: ‘
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<location id="2C2A6040FFFF6D4ED513D8E7BDA207A9" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED513D8E7BDA207A9" box="[857,933,464,490]" country="Guinea-Bissau" name="Okapa" pageId="22" pageNumber="273">Okapa</location>
,
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<collectingDate id="4D0FE9B3FFFF6D4ED5FAD8E7BA2E07A8" box="[944,1065,464,491]" pageId="22" pageNumber="273" value="1965-11-08">8.XI.1965</collectingDate>
</date>
’, ‘
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</materialsCitation>
.
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<location id="2C2A6040FFFF6D4ED228D8E7BAEE07A9" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED228D8E7BAEE07A9" box="[1122,1257,464,490]" country="Guinea-Bissau" name="Hornabrook" pageId="22" pageNumber="273">Hornabrook</location>
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.’ (
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);
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: ‘
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<location id="2C2A6040FFFF6D4ED420D8C3BCB1044D" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED420D8C3BCB1044D" box="[618,694,500,526]" country="Guinea-Bissau" municipality="Awande" name="Okapa" pageId="22" pageNumber="273">Okapa</location>
,
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<collectingDate id="4D0FE9B3FFFF6D4ED560D8C3BDB7044C" box="[810,944,500,527]" pageId="22" pageNumber="273" value="1965-09-27">27.IX.1965</collectingDate>
</date>
’, ‘
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</materialsCitation>
.
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<location id="2C2A6040FFFF6D4ED5A1D8C3BA75044D" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED5A1D8C3BA75044D" box="[1003,1138,500,526]" country="Guinea-Bissau" name="Hornabrook" pageId="22" pageNumber="273">Hornabrook</location>
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.’ (
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);
<specimenCount id="3FF3FD12FFFF6D4ED6DDDB2EBEDB0470" box="[151,220,537,563]" count="3" pageId="22" pageNumber="273" type="female">3 ♀♀</specimenCount>
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<location id="2C2A6040FFFF6D4ED7FDDB2FBFFD0471" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED7FDDB2FBFFD0471" box="[439,506,536,562]" country="Guinea-Bissau" name="Oiana" pageId="22" pageNumber="273">Oiana</location>
[Auyana],
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<collectingDate id="4D0FE9B3FFFF6D4ED4CBDB2FBD000470" box="[641,775,536,563]" pageId="22" pageNumber="273" value="1965-11-12">12.XI.1965</collectingDate>
</date>
’, ‘
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</materialsCitation>
.
<materialsCitation id="999D3CC6FFFF6D4ED500DB2FBB9B0470" box="[842,1436,536,563]" collectionCode="BMNH, NEW" country="Guinea-Bissau" location="Hornabrook" pageId="22" pageNumber="273" specimenCode="BM 1970-232" specimenCount="1" specimenCount-female="1">
<location id="2C2A6040FFFF6D4ED500DB2FBDD60471" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED500DB2FBDD60471" box="[842,977,536,562]" country="Guinea-Bissau" name="Hornabrook" pageId="22" pageNumber="273">Hornabrook</location>
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.’ (
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);
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<collectingCountry id="51E2760BFFFF6D4ED6DDDB0ABF060414" box="[151,257,573,599]" name="Guinea-Bissau" pageId="22" pageNumber="273">GUINEA</collectingCountry>
<location id="2C2A6040FFFF6D4ED742DB0ABF500414" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED742DB0ABF500414" box="[264,343,573,599]" country="Guinea-Bissau" name="Purosa" pageId="22" pageNumber="273">Purosa</location>
,
<date id="5D4B105BFFFF6D4ED728DB0BBFEA0414" box="[354,493,572,599]" pageId="22" pageNumber="273" value="1965-07-21">
<collectingDate id="4D0FE9B3FFFF6D4ED728DB0BBFEA0414" box="[354,493,572,599]" pageId="22" pageNumber="273" value="1965-07-21">21.VII.1965</collectingDate>
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, wet leaves etc.’, ‘
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</materialsCitation>
.
<materialsCitation id="999D3CC6FFFF6D4ED494DB0BBA8D0414" box="[734,1162,572,599]" collectionCode="BMNH" country="Guinea-Bissau" location="Hornabrook" pageId="22" pageNumber="273" specimenCode="BM 1970-232" specimenCount="1">
<location id="2C2A6040FFFF6D4ED494DB0BBD620415" LSID="urn:lsid:plazi:treatment:A15C878DFFFF6D4ED6DDD9A0BFEC04A5:2C2A6040FFFF6D4ED494DB0BBD620415" box="[734,869,572,598]" country="Guinea-Bissau" name="Hornabrook" pageId="22" pageNumber="273">Hornabrook</location>
<specimenCode id="79539EE0FFFF6D4ED527DB0ABA160414" box="[877,1041,573,599]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="22" pageNumber="273">BM 1970-232</specimenCode>
.’ (
<collectionCode id="4FE4AE5EFFFF6D4ED262DB0ABA850414" box="[1064,1154,573,599]" pageId="22" pageNumber="273">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFFF6D4ED68DDB56BFEC04A5" pageId="22" pageNumber="273" type="discussion">
<paragraph id="294A369BFFFF6D4ED68DDB56BFEC04A5" blockId="22.[151,1437,356,743]" pageId="22" pageNumber="273">
<emphasis id="1B81EA89FFFF6D4ED68DDB56BF3C0438" bold="true" box="[199,315,609,635]" pageId="22" pageNumber="273">Remarks.</emphasis>
<taxonomicName id="EEF54D18FFFF6D4ED708DB57BC440439" authorityName="W. J. MacLeay" authorityYear="1873" box="[322,579,608,634]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="gayndahense">
<emphasis id="1B81EA89FFFF6D4ED708DB57BC440439" box="[322,579,608,634]" italics="true" pageId="22" pageNumber="273">Omalium gayndahense</emphasis>
</taxonomicName>
, originally described from
<collectingCountry id="51E2760BFFFF6D4ED53EDB57BDD90439" box="[884,990,608,634]" name="Australia" pageId="22" pageNumber="273">Australia</collectingCountry>
, is widely distributed in the world as an introduced species (e.g. Shavrin 2020, 2022, 2023b). It was recorded by
<bibRefCitation id="4D644B6AFFFF6D4ED589DBB3BA7E04DD" author="Shavrin, A. V." box="[963,1145,644,670]" pageId="22" pageNumber="273" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
from
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. The distribution of the species in New
<collectingCountry id="51E2760BFFFF6D4ED407DB9FBC990481" box="[589,670,680,706]" name="Guinea-Bissau" pageId="22" pageNumber="273">Guinea</collectingCountry>
as in
<figureCitation id="B1CE2A1EFFFF6D4ED4ABDB9FBD330480" box="[737,820,680,707]" captionStart="FIGURES 45" captionStartId="18.[151,264,885,909]" captionTargetBox="[151,1436,193,862]" captionTargetId="figure-22@18.[151,1436,181,862]" captionTargetPageId="18" captionText="FIGURES 45. Distribution of Paraphloeostiba coriacea (circles) and P. gayndahensis (squares) in New Guinea." figureDoi="http://doi.org/10.5281/zenodo.10821237" httpUri="https://zenodo.org/record/10821237/files/figure.png" pageId="22" pageNumber="273">Fig. 45</figureCitation>
. The species is here recorded from
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(Western New
<collectingCountry id="51E2760BFFFF6D4ED699DBFBBF2104A5" box="[211,294,716,742]" name="Guinea-Bissau" pageId="22" pageNumber="273">Guinea</collectingCountry>
) for the frst time.
</paragraph>
</subSubSection>
</treatment>
</document>

352
data/A1/5C/93/A15C9394F16ED8E2FB96B843907397C0.xml Normal file
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<mods:namePart id="F0F413331AB7C66ED7F2437016509825">Couderc, Marie</mods:namePart>
<mods:nameIdentifier id="16FA647FFDC3A2EA3761CD05021B91DB" type="ORCID">https://orcid.org/0000-0001-7297-155X</mods:nameIdentifier>
<mods:affiliation id="60E4158B5B10F7F379B9855E088C5D4B">DIADE, Universite de Montpellier, IRD, CIRAD, Montpellier, France</mods:affiliation>
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<mods:namePart id="368F28CF9C5AE2B67802982BB1EE21F3">Gaudeul, Myriam</mods:namePart>
<mods:affiliation id="5665C95C0E38B4F78012F897699138FC">Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum National d'Histoire Naturelle-CNRS-SU-EPHE-UA, 57 rue Cuvier, CP 39, 75231 Paris, Cedex 05, France</mods:affiliation>
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<mods:namePart id="9AB0B6EA40BD9E3A785E2CEE5D44A334">Grall, Aurelie</mods:namePart>
<mods:nameIdentifier id="9AE99052E43EF3BBADD49FD0640A5E87" type="ORCID">https://orcid.org/0000-0002-1613-4703</mods:nameIdentifier>
<mods:affiliation id="6091BA7466260E4FE1F3FE8D8087CF82">Herbaria Basel, Department of Environmental Sciences, University of Basel, Basel, Switzerland &amp; Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
</mods:name>
<mods:name id="65D0B6A5756547ACFDF0D4DAEC04D198" type="personal">
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<mods:namePart id="4BB6F96E880536D85A8D22BFA741CC4E">Loup, Caroline</mods:namePart>
<mods:affiliation id="6003099D09FF7E0C14948C794E140DC2">DIADE, Universite de Montpellier, IRD, CIRAD, Montpellier, France</mods:affiliation>
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</mods:role>
<mods:namePart id="CD1920E647ABD4E958916A1BA08484BF">Wieringa, Jan J.</mods:namePart>
<mods:nameIdentifier id="1C6B19C269F58BF63C42BB87BEE2F87A" type="ORCID">https://orcid.org/0000-0003-0566-372X</mods:nameIdentifier>
<mods:affiliation id="874F850DD4EDAC99D7E36191DBEF7A1A">Herbier MPU, DCSPH - CC 99010, Universite de Montpellier, 163 rue A. Broussonnet, F- 34090 Montpellier, France</mods:affiliation>
</mods:name>
<mods:name id="01E14DD2AD3BBA3FA3C262E10FCF2559" type="personal">
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</mods:role>
<mods:namePart id="E1DF46B48BF55F3FA5C8325C91290652">Sonke, Bonaventure</mods:namePart>
<mods:nameIdentifier id="1C86DF49C6E0A52352F2AA79F9E2126E" type="ORCID">https://orcid.org/0000-0002-4310-3603</mods:nameIdentifier>
<mods:affiliation id="D9C0DD9FE09820705A1674E222DDE318">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name id="1E6F3C9EA2CABD3AFFCB14B1F50820F4" type="personal">
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</mods:role>
<mods:namePart id="0C6DB8A37C74C6D5F1A96F1B6FC07867">Couvreur, Thomas L. P.</mods:namePart>
<mods:nameIdentifier id="2D255AD52C47EEDE208E15D13C70DF8B" type="ORCID">https://orcid.org/0000-0002-8509-6587</mods:nameIdentifier>
<mods:affiliation id="47E9A4C903090B4D2EACE896784C8BDF">DIADE, Universite de Montpellier, IRD, CIRAD, Montpellier, France</mods:affiliation>
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<mods:title id="00DC4FA4073B8F0E35D230A83B7AA0A6">PhytoKeys</mods:title>
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<mods:part id="A031BAB73E04CF49942CA66404E244F5">
<mods:date id="7D163F0AC7946BFAF4104D2B438FB4D6">2023</mods:date>
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<mods:number id="56335DC4417C14513C034199F109D852">233</mods:number>
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<mods:start id="C06ED028115B8388D7A1DA594D700759">1</mods:start>
<mods:end id="69B0C35CBD8BAA9D8ECCBEC7230E1A18">200</mods:end>
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<mods:url id="A0332E16273B48D7FB46D7ADF4C1F4CF">http://dx.doi.org/10.3897/phytokeys.233.103096</mods:url>
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<mods:classification id="FFDF87E8F61A1770E7F7ACA96FD3B9B7">journal article</mods:classification>
<mods:identifier id="CB3B57F721FB107132DD7B5982481A93" type="DOI">http://dx.doi.org/10.3897/phytokeys.233.103096</mods:identifier>
<mods:identifier id="7F1AB626386CDA59CEF4EDF410B03A82" type="Pensoft-Pub">1314-2003-233-1</mods:identifier>
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<treatment id="A15C9394F16ED8E2FB96B843907397C0" LSID="urn:lsid:plazi:treatment:A15C9394F16ED8E2FB96B843907397C0" httpUri="http://treatment.plazi.org/id/A15C9394F16ED8E2FB96B843907397C0" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="BAC911380EE9503469913308454DE07A" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="E0A9657C5B99780A19CD6356D4C23286" pageId="0" pageNumber="1">
<taxonomicName id="5CE9760F5FE64C8B7343298E7C007F7A" ID-CoL="7F65R" authority="Verdc., Kew Bull. 41 (2): 289 (1986)" authorityName="Verdc., Kew Bull. 41 (2): 289" authorityYear="1986" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis bisexualis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bisexualis">Uvariopsis bisexualis Verdc., Kew Bull. 41(2): 289 (1986)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5F5E80853A134056F86C70A44C2FF77E" pageId="0" pageNumber="1" type="description">
<paragraph id="CF7571E13203BDCB39DEBFDDDAC45273" pageId="0" pageNumber="1">
<figureCitation id="6847BB0B99FFA4C8AE1BD3CDF35AE3BA" captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Uvariopsis bisexualis Verdc A flowering branch B flower bud, side view C sepal, outer view D flower, two petals removed, side view E detail of receptacle, some stamens fallen, side view F stamen, upper view G stamen, lower view H carpel, outer view J carpel, inner view K stigma L longitudinal section of carpel M transversal section of ovary. A-M from Lovett 233 (type). Drawings by Maureen Church, from Verdcourt (1986; fig. 3, p. 292), Kew Bulletin 1969 Royal Botanic Gardens, Kew." figureDoi="10.3897/phytokeys.233.103096.figure51" httpUri="https://binary.pensoft.net/fig/909910" pageId="0" pageNumber="1">Figs 51</figureCitation>
<figureCitation id="951A7A6CD9E41E0BE933ACBFA97D2DDC" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Distribution map of Uvariopsis bisexualis. Shades of grey represent elevation, from white (sea level) to darker grey (higher elevation). The inset shows the extent of the map over Africa." figureDoi="10.3897/phytokeys.233.103096.figure52" httpUri="https://binary.pensoft.net/fig/909911" pageId="0" pageNumber="1">, 52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2FC818E5D6554C24F1FE74B8838E3BC3" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="BA667A8856680259854F930F659CD031" pageId="0" pageNumber="1">
<typeStatus id="C4C88E95B4306A5ACA111F2B957569E6">Type</typeStatus>
.
</paragraph>
<paragraph id="9575227D012BA23FF70FEDC476A4A8D2" pageId="0" pageNumber="1">
<materialsCitation id="DF9540D095D31E93357E56FA045975EB" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="J. C. Lovett" country="Tanzania" location="Iringa" specimenCount="1" stateProvince="Iringa" typeStatus="holotype">
<collectingCountry id="B5C9151FB8A6CBA28F131D393EF39FB7" name="Tanzania">Tanzania</collectingCountry>
-
<collectingRegion id="7ACE99907F8D7CD22B44DD5038773DFC" country="Tanzania" name="Iringa">Iringa</collectingRegion>
•
<collectorName id="02E4884F794863C5E7AC6B1BD6E41E71">J.C. Lovett</collectorName>
233 (
<typeStatus id="0C346AE880E0A30254CBD04C2078DD97">holotype</typeStatus>
: K! (K000199040))
</materialsCitation>
;
<materialsCitation id="0ED379159B9A5F024C0EDA435FDAD216" collectingDate="1983-12-02" country="Tanzania" county="Rural District" elevation="1480" latitude="-7.7666664" location="Sanje" longLatPrecision="1303" longitude="36.9" municipality="Udzungwa Mts." specimenCount="1" stateProvince="Iringa" typeStatus="holotype">
<collectingRegion id="C2A0EA6C066DE9B22C28F63E48EAF4BC" country="Tanzania" name="Iringa">Iringa</collectingRegion>
<collectingCounty id="1867443CA02C953141A6852D5B77B13B">Rural District</collectingCounty>
,
<collectingMunicipality id="A50261B6D39024F5CEBEB56B92630FE0">Udzungwa Mts.</collectingMunicipality>
,
<location id="8A4F7FE74BD22B3949BCD861137A06DF" LSID="urn:lsid:plazi:treatment:A15C9394F16ED8E2FB96B843907397C0:8A4F7FE74BD22B3949BCD861137A06DF" country="Tanzania" county="Rural District" latitude="-7.7666664" longLatPrecision="1303" longitude="36.9" municipality="Udzungwa Mts." name="Sanje" stateProvince="Iringa">Sanje</location>
;
<geoCoordinate id="C20BF2F42FD4B4B93774D5DD0F8FCAA6" degrees="7" direction="south" minutes="46" orientation="latitude" precision="925" value="-7.7666664">7°46'S</geoCoordinate>
,
<geoCoordinate id="5097B0CF0B4BB549C7D0D00605F84952" degrees="36" direction="east" minutes="54" orientation="longitude" precision="925" value="36.9">36°54'E</geoCoordinate>
; alt.
<quantity id="88B1F1910ACD70272FD41349E1794CF0" metricMagnitude="3" metricUnit="m" metricValue="1.48" unit="m" value="1480.0">
<elevation id="3B8A61B41F222E55BA33B11C18B12777" metricMagnitude="3" metricUnit="m" metricValue="1.48" unit="m" value="1480.0">1480 m</elevation>
</quantity>
;
<collectingDate id="4A656184000E44BE9865C1C124698C1D" value="1983-12-02">02 Dec. 1983</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="391451E65760B55C53C5638FBBCE5248" pageId="0" pageNumber="1" type="description">
<paragraph id="42D9427B5BEF5B61E77B4336E70E52EB" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="0B02CD7D939DC9C97B7E2BCCD82EABCD" pageId="0" pageNumber="1">Tree ca. 10 m tall, D.B.H unknown; young branches pubescent, old branches glabrous. Petiole 3.5-5 mm long, ca. 2 mm wide, sparsely pubescent to glabrous. Leaf lamina 135-152 mm long, 37-48 mm wide, length:width ratio ca. 3, elliptic to oblong, coriaceous, base decurrent, apex attenuate, surface above glabrous, surface below sparsely pubescent to glabrous when young, glabrous when old; midrib impressed above, raised below, glabrous above, sparsely pubescent to glabrous below; secondary veins ca. 12 pairs, weakly brochidodromous, impressed above, raised below; tertiary veins reticulate. Flowers bisexual. Flower buds ovoid to conical. Inflorescences borne on branches, composed of 1 flower. Peduncle inconspicuous. Flower pedicel 12-20 mm long, ca. 1 mm in diameter, pubescent. Bracts 1 at base to towards the lower half of the pedicel, ca. 1 mm long, ca. 1 mm wide. Sepals 2, 1.6-2.5 mm long, 2.2-2.5 mm wide, broadly ovate, free, pubescent outside, glabrous inside, color unknown. Petals 4, 12-14 mm long, 4-8 mm wide, length:width ratio 3-3.5, narrowly ovate, sparsely pubescent outside, glabrous inside, cream outside, cream with red base inside. Stamens numerous (exact number unknown), 0.5-0.7 mm long, ca. 0.5 mm wide, anthers linear, connective prolongation truncate or absent. Carpels 3 to 6, 2-4 mm long, 1-1.4 mm wide, glabrate at base to glabrous, free, ovules 22-24 per ovary, biseriate. Fruits unknown.</paragraph>
<caption id="09E373E3ABCB0AB0FBB43C964D7F35F8" doi="10.3897/phytokeys.233.103096.figure51" httpUri="https://binary.pensoft.net/fig/909910" pageId="0" pageNumber="1" start="Figure 51" startId="F51">
<paragraph id="638292383C5EC6F9CAAEE336B2CDA196" pageId="0" pageNumber="1">
<emphasis id="D460CD28377FA7BBDC31099BCFBE8D55" bold="true" pageId="0" pageNumber="1">Figure 51.</emphasis>
<taxonomicName id="211766852C766BB7770B9BCAFDA44520" authorityName="Verdc., Kew Bull. 41 (2): 289" authorityYear="1986" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis bisexualis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bisexualis">
<emphasis id="0FD71B2913A74330BCFC4E35341DF88B" italics="true" pageId="0" pageNumber="1">Uvariopsis bisexualis</emphasis>
</taxonomicName>
Verdc
<emphasis id="A6C9634E122CA057150533A0F3C18235" bold="true" pageId="0" pageNumber="1">A</emphasis>
flowering branch
<emphasis id="99DDE0BF3C957D6EA1F07C2DACE72BE1" bold="true" pageId="0" pageNumber="1">B</emphasis>
flower bud, side view
<emphasis id="39916EFEB8D6F5C4F62E59057C6B32D8" bold="true" pageId="0" pageNumber="1">C</emphasis>
sepal, outer view
<emphasis id="639A354D4C3C9360EFB5B976E2DD7C26" bold="true" pageId="0" pageNumber="1">D</emphasis>
flower, two petals removed, side view
<emphasis id="EE0A05F78860A0DB599617210176467A" bold="true" pageId="0" pageNumber="1">E</emphasis>
detail of receptacle, some stamens fallen, side view
<emphasis id="4956D8E32549451B74A0DD1EDE6D036E" bold="true" pageId="0" pageNumber="1">F</emphasis>
stamen, upper view
<emphasis id="C2F305512A9036DF5818AF609F967F71" bold="true" pageId="0" pageNumber="1">G</emphasis>
stamen, lower view
<emphasis id="3D7D51AEB2C4DF1F1C7AD006B8C54716" bold="true" pageId="0" pageNumber="1">H</emphasis>
carpel, outer view
<emphasis id="1F9D07F07A8BCAA632BB2B6E958795C4" bold="true" pageId="0" pageNumber="1">J</emphasis>
carpel, inner view
<emphasis id="6F17FBDC5B4B3C7D8278074ED81FDEF2" bold="true" pageId="0" pageNumber="1">K</emphasis>
stigma
<emphasis id="89BFD81F47608C04EA06EC2AE1D8307B" bold="true" pageId="0" pageNumber="1">L</emphasis>
longitudinal section of carpel
<emphasis id="B23A16315C0FBACA835823D9FB2141BD" bold="true" pageId="0" pageNumber="1">M</emphasis>
transversal section of ovary.
<emphasis id="B2C8BC5923CB578DA12413061C0E0075" bold="true" pageId="0" pageNumber="1">A-M</emphasis>
from Lovett 233 (type). Drawings by Maureen Church, from
<bibRefCitation id="2C805C4E318554E9CFC4C6CD0814D1A2" DOI="https://doi.org/10.2307/4102932" author="Verdcourt, B" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="1" pagination="287 - 297" refId="B180" refString="Verdcourt, B, 1986. New Taxa of East African Annonaceae. Kew Bulletin 41 (2): 287 - 297, DOI: https://doi.org/10.2307/4102932" title="New Taxa of East African Annonaceae." url="https://doi.org/10.2307/4102932" volume="41" year="1986">Verdcourt (1986</bibRefCitation>
; fig. 3, p. 292), Kew Bulletin 1969 Royal Botanic Gardens, Kew.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C5B8D451D408F3A545C04AE52A3789DA" pageId="0" pageNumber="1" type="distribution">
<paragraph id="4E8503B2BEAD41A617F3C004610055D4" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="1E83FB262AD7F7E935B4B55DB3D76920" pageId="0" pageNumber="1">Endemic to Somalia-Masai Region. Known from only one locality in Tanzania: Sanje in the Udzungwa mountains.</paragraph>
</subSubSection>
<subSubSection id="63663062240DB327A993FB70BDB54E3D" pageId="0" pageNumber="1" type="habitat">
<paragraph id="D3ECE3CD66A29BE8DB6B017F733E209F" pageId="0" pageNumber="1">Habitat and ecology.</paragraph>
<paragraph id="3F433656773F943598F6D276CCB0CE3E" pageId="0" pageNumber="1">Montane rain forests. Altitude ca. 1480 m asl.</paragraph>
</subSubSection>
<subSubSection id="BCEF64B05FBFEA327A17F105C643BBC3" pageId="0" pageNumber="1" type="phenology">
<paragraph id="D5CA3C8BCC2F604DCE84A63C1BE3CB85" pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph id="DAA9278218E414BDA29CD0FA1F5DD4C5" pageId="0" pageNumber="1">Flowers collected in December.</paragraph>
</subSubSection>
<subSubSection id="C1BE16D540F97394916CE7E70A680DF8" pageId="0" pageNumber="1" type="notes">
<paragraph id="4FC5323F82AFE7980A4EAAD83829CA03" pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph id="068A6B59FD6A6F95689FF3D3C4132DB4" pageId="0" pageNumber="1">
This species resembles
<taxonomicName id="94D578C4572791D46B0B64BC87E64C1B" class="Magnoliopsida" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Up. congensis" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="congensis">
<emphasis id="4E6EFE6F836096F8BC673313264A2FE7" italics="true" pageId="0" pageNumber="1">Up. congensis</emphasis>
</taxonomicName>
,
<taxonomicName id="EB2EA5DEBFA91753118ABAF48511D4D2" class="Magnoliopsida" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Up. oligocarpa" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="oligocarpa">
<emphasis id="D7DC59B181C9E06F8677A4F09B65A8FF" italics="true" pageId="0" pageNumber="1">Up. oligocarpa</emphasis>
</taxonomicName>
and
<taxonomicName id="5E6BD64DFE7ADD16A6F3DD143D9C4F53" class="Magnoliopsida" family="Annonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Up. zenkeri" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="zenkeri">
<emphasis id="1AACDD8B4030250E9F1B8EC60D184D1B" italics="true" pageId="0" pageNumber="1">Up. zenkeri</emphasis>
</taxonomicName>
in having elliptic leaves generally less than 16 cm long with decurrent base. It differs from all the other
<taxonomicName id="635795F8B230DC26C8F36E0B8BA788DF" authorityName="Engl., Notizbl. Koenigl. Bot. Gart. Berlin 2: 298" authorityYear="1899" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="DD9B4F6E114B94F65A5848873D3C4D1B" italics="true" pageId="0" pageNumber="1">Uvariopsis</emphasis>
</taxonomicName>
in having bisexual flowers and less than 6 carpels (vs. more than 10 carpels) (Table
<tableCitation id="4B9F015464DB11A895D543FE581B2AD6" captionStart="Table 6" captionStartId="T6" captionText="Table 6. Morphological comparison between Uvariopsis bisexualis, Uvariopsis congensis, Uvariopsis oligocarpa and Uvariopsis zenkeri. In bold: character useful to differentiate the species." httpUri="http://table.plazi.org/id/DAE5546502DDDCC540255EAAB62B22A0" pageId="0" pageNumber="1" tableUuid="DAE5546502DDDCC540255EAAB62B22A0">6</tableCitation>
).
</paragraph>
<caption id="BF1D96C8998151E3E72C2E22836444B4" doi="10.3897/phytokeys.233.103096.figure52" httpUri="https://binary.pensoft.net/fig/909911" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
<paragraph id="1363D1068DB4F79327FFBB8BB278F068" pageId="0" pageNumber="1">
<emphasis id="91FD058A46EC8ED06D5072B6A6AE429B" bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
Distribution map of
<taxonomicName id="44A668447C2CDBBC6ABB2137A2A46CB0" authorityName="Verdc., Kew Bull. 41 (2): 289" authorityYear="1986" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis bisexualis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bisexualis">
<emphasis id="AC097804B9BD7EFACC4EB217101C2A61" italics="true" pageId="0" pageNumber="1">Uvariopsis bisexualis</emphasis>
</taxonomicName>
. Shades of grey represent elevation, from white (sea level) to darker grey (higher elevation). The inset shows the extent of the map over Africa.
</paragraph>
</caption>
<caption id="DAE5546502DDDCC540255EAAB62B22A0" ID-Table-UUID="DAE5546502DDDCC540255EAAB62B22A0" httpUri="http://table.plazi.org/id/DAE5546502DDDCC540255EAAB62B22A0" pageId="0" pageNumber="1" start="Table 6" startId="T6">
<paragraph id="D3100AC0D96F74AEC74D2FA0BAAC6C16" pageId="0" pageNumber="1">
<emphasis id="E7E0CA549E5BB9DB6D92080757CA2116" bold="true" pageId="0" pageNumber="1">Table 6.</emphasis>
Morphological comparison between
<taxonomicName id="71130CD3CF18D4E48791420CDB17FFEC" authorityName="Verdc., Kew Bull. 41 (2): 289" authorityYear="1986" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis bisexualis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bisexualis">
<emphasis id="BC1B43F6B37544E18622D4E69800DAF9" italics="true" pageId="0" pageNumber="1">Uvariopsis bisexualis</emphasis>
</taxonomicName>
,
<taxonomicName id="20DD09EB8C212EB5ABD281BAD0506327" authorityName="Robyns &amp; Ghesq., Ann. Soc. Sci. Bruxelles, Ser. B liii. 317" authorityYear="1933" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis congensis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="congensis">
<emphasis id="926EF873E7F5229C8D1AE4B96E9D416F" italics="true" pageId="0" pageNumber="1">Uvariopsis congensis</emphasis>
</taxonomicName>
,
<taxonomicName id="915972EDF63C8109438B095D811256D0" authorityName="Dagallier &amp; Couvreur" authorityYear="2023" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis oligocarpa" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="oligocarpa">
<emphasis id="8EE0820202F8FABBCFA9B2253CAE1CB6" italics="true" pageId="0" pageNumber="1">Uvariopsis oligocarpa</emphasis>
</taxonomicName>
and
<taxonomicName id="704377B41E60727472D3E266C1047B2E" authorityName="Engl., Notizbl. Koenigl. Bot. Gart. Berlin 2: 298" authorityYear="1899" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis zenkeri" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="zenkeri">
<emphasis id="CF9A8CD31570B2B3BBA60EDC3637D35F" italics="true" pageId="0" pageNumber="1">Uvariopsis zenkeri</emphasis>
</taxonomicName>
. In bold: character useful to differentiate the species.
</paragraph>
</caption>
<paragraph id="0D43ADECFCC0BF528C8573A79665B21A" pageId="0" pageNumber="1">
<table id="95DA8DE320D37E0D672BEE71EED73045" pageId="0" pageNumber="1">
<tr id="7C543A9B6925A31D525E7A65B3E9E621" pageId="0" pageNumber="1">
<th id="AB73CFBD957D0D52B0EC6B8EFC7683FA" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</th>
<th id="F19024AD2E24954B1D51CDA94B6E3B4A" colspan="1" pageId="0" pageNumber="1" rowspan="1">Young branches indumentum</th>
<th id="3E48899A31714F38A302B80640417CF8" colspan="1" pageId="0" pageNumber="1" rowspan="1">Flowers</th>
<th id="1F3D66FFBDE04F244E5B9C78FE1E09CD" colspan="1" pageId="0" pageNumber="1" rowspan="1">Carpels number</th>
<th id="EA22ABA077343EFB6142DC04E195CFA4" colspan="1" pageId="0" pageNumber="1" rowspan="1">Fruits</th>
</tr>
<tr id="B7BF059E3D30B6DC7811423989F9B122" pageId="0" pageNumber="1">
<td id="7298388382D6C76F1A9ECB8E33B68B5E" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="B820D06CB92BBEB8E743AA10A7C13991" authorityName="Verdc., Kew Bull. 41 (2): 289" authorityYear="1986" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis bisexualis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bisexualis">
<emphasis id="2030913EB37A876D1D677E7522E10242" italics="true" pageId="0" pageNumber="1">Uvariopsis bisexualis</emphasis>
</taxonomicName>
</td>
<td id="F1C0415534BB10460A5F585866CBA97F" colspan="1" pageId="0" pageNumber="1" rowspan="1">pubescent</td>
<td id="A67FB5F9401E26AF16923C77C1AE8B45" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="86A08F9EE68A91EDE4CE5D76A517211B" bold="true" pageId="0" pageNumber="1">bisexual</emphasis>
</td>
<td id="50B3E27F0BBC81F86BD83097F97AAD73" colspan="1" pageId="0" pageNumber="1" rowspan="1">3 to 6</td>
<td id="A20248729B1D2C33499CE7842A7D0940" colspan="1" pageId="0" pageNumber="1" rowspan="1">unknown</td>
</tr>
<tr id="7D0C5DAB15087D6376495AE673A4DF89" pageId="0" pageNumber="1">
<td id="79C96C9ECE2E3E8F7611871E52D2D7B4" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="5BC1AF42A0513C501B45B7A9323F66FB" authorityName="Robyns &amp; Ghesq., Ann. Soc. Sci. Bruxelles, Ser. B liii. 317" authorityYear="1933" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis congensis" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="congensis">
<emphasis id="257421C0AB227BB3F850F67D48D35104" italics="true" pageId="0" pageNumber="1">Uvariopsis congensis</emphasis>
</taxonomicName>
</td>
<td id="E41E49D31856484EEB627F2CCA92B000" colspan="1" pageId="0" pageNumber="1" rowspan="1">sparsely pubescent to glabrous</td>
<td id="68B1913738C095A27EA228B166A2D4D6" colspan="1" pageId="0" pageNumber="1" rowspan="1">unisexual</td>
<td id="F5274715C9626D56352605197D0A9D9C" colspan="1" pageId="0" pageNumber="1" rowspan="1">20 to 40</td>
<td id="A2CA9293DB41A24B7A64983374AEE774" colspan="1" pageId="0" pageNumber="1" rowspan="1">
1 to 15 monocarps, cylindrical, smooth to lumpy,
<emphasis id="C24D2EA94087831A4744663F19E2CBAE" bold="true" pageId="0" pageNumber="1">strongly constricted between the seeds</emphasis>
,
<emphasis id="349D64D2D7090B7AF66CB1AC8E25C7A8" bold="true" pageId="0" pageNumber="1">glabrate to glabrous</emphasis>
, green to
<emphasis id="3DE99632FAC4F8B947FEF33A9AEE6763" bold="true" pageId="0" pageNumber="1">orange to red</emphasis>
when ripe (
<emphasis id="BF496BDF8F1335D9963CE3B6C0725705" bold="true" pageId="0" pageNumber="1">black when dry</emphasis>
), sessile to stipitate with stipe 0-7 mm long
</td>
</tr>
<tr id="1FC48E0563A94D371D04E9AAD37F4CC2" pageId="0" pageNumber="1">
<td id="D1A030F2988AAEC7DE21E0B57FD131EC" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="8E5BA035BFEFB63BA2D5F422628204AF" authorityName="Dagallier &amp; Couvreur" authorityYear="2023" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis oligocarpa" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="oligocarpa">
<emphasis id="4D25DE530A84A3531CC87B738E844F1C" italics="true" pageId="0" pageNumber="1">Uvariopsis oligocarpa</emphasis>
</taxonomicName>
</td>
<td id="B1EE7FB607273D635E891470EC43AC51" colspan="1" pageId="0" pageNumber="1" rowspan="1">glabrous</td>
<td id="509A8053D7709A8DE0FF2E6F9D5ED3EF" colspan="1" pageId="0" pageNumber="1" rowspan="1">unisexual</td>
<td id="A108D39D1CEAC8A5F9FDB0202193DC79" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="A6EF5AB1BA5AE4C89E000723EF61EFD7" bold="true" pageId="0" pageNumber="1">13 to 20</emphasis>
</td>
<td id="E1BEB85EE8F7E81CEA2871C22C823A2D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="13C6076E7C8F45CE01F5114ABC08D737" bold="true" pageId="0" pageNumber="1">1-3 monocarps</emphasis>
, cylindrical, smooth,
<emphasis id="9CA02AC567100644492AAAD760B2D342" bold="true" pageId="0" pageNumber="1">pubescent to sparsely pubescent</emphasis>
, green to
<emphasis id="F6AA6D68F99D6DAE56DBE340D3D1D581" bold="true" pageId="0" pageNumber="1">yellow to orange</emphasis>
when ripe (
<emphasis id="33E11FA427CA2540CF04EE089B5E46B7" bold="true" pageId="0" pageNumber="1">orange-brown when dry</emphasis>
), sessile to very shortly stipitate with stipe 0-1 mm long
</td>
</tr>
<tr id="EA7B7DF45FA8AFB7E83BFA825C266C6B" pageId="0" pageNumber="1">
<td id="DDFD75605FBAAB01C34425915F33FE2F" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="7DEEAEBD59C2BBB800845219E198070A" authorityName="Engl., Notizbl. Koenigl. Bot. Gart. Berlin 2: 298" authorityYear="1899" class="Magnoliopsida" family="Annonaceae" genus="Uvariopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uvariopsis zenkeri" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="zenkeri">
<emphasis id="D5BBC868D2040224E2A559AD03C72F99" italics="true" pageId="0" pageNumber="1">Uvariopsis zenkeri</emphasis>
</taxonomicName>
</td>
<td id="F7A3CAA62D325C86AB8041812B8F2882" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="D136D6445B0DF12F38AB792DA67B9091" bold="true" pageId="0" pageNumber="1">densely pubescent to pubescent</emphasis>
</td>
<td id="4A5D68F437312E3A40F2BE66805ACB3E" colspan="1" pageId="0" pageNumber="1" rowspan="1">unisexual</td>
<td id="93AF7594EC69906213CBF533D0177E6E" colspan="1" pageId="0" pageNumber="1" rowspan="1">13 to 22</td>
<td id="645C1433C79349F3D939095C24D1F322" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="D93520C77EAB98AC8BA5520BCABBF742" bold="true" pageId="0" pageNumber="1">1-3 monocarps</emphasis>
, cylindrical, slightly veined and constricted between the seed when dried,
<emphasis id="944A1F9C25576FA3A417770E96269029" bold="true" pageId="0" pageNumber="1">tomentose</emphasis>
, brown, subsessile with stipe ca. 1 mm long
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="ACD9822AB2DD55FB11B8953016DB5278" pageId="0" pageNumber="1" type="conservation status">
<paragraph id="8389367ABBD9DC9B9DA3E253782E88DD" pageId="0" pageNumber="1">Conservation status.</paragraph>
<paragraph id="5B08E24E482D762D2C61BD2EBC5A10D3" pageId="0" pageNumber="1">This species has been assessed as Endangered EN under criteria B1ab(iii) (Eastern Arc Mountains &amp; Coastal Forests CEPF Plant Assessment Project 2009e). It is known from a single specimen collected more than 30 years ago in the Udzungwa Mountains National Park in Tanzania.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart>de Montpellier d'Annevoie, Clotilde</mods:namePart>
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<mods:namePart>Serteyn, Laurent</mods:namePart>
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<taxonomicName LSID="A15CB08A-5268-5E21-A817-536C1D11A458" authority="(Fabricius 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" class="Hexapoda" family="Apidae" genus="Bombus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bombus (Bombus) cryptarum" order="Hymenoptera" pageId="0" pageNumber="60665" phylum="Arthropoda" rank="species" species="cryptarum" subGenus="Bombus">Bombus (Bombus) cryptarum (Fabricius 1775)</taxonomicName>
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<paragraph pageId="0" pageNumber="60665">Ecological interactions</paragraph>
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<paragraph pageId="0" pageNumber="60665">Feeds on</paragraph>
<paragraph pageId="0" pageNumber="60665">Polylectic</paragraph>
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<paragraph pageId="0" pageNumber="60665">Conservation status</paragraph>
<paragraph pageId="0" pageNumber="60665">Endangered</paragraph>
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<paragraph pageId="0" pageNumber="60665">Notes</paragraph>
<paragraph pageId="0" pageNumber="60665">Table S1 (Historical data)</paragraph>
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