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<document id="315551E38FDAD967D82736C07AAB77E1" ID-CLB-Dataset="7809" ID-DOI="10.5281/zenodo.4503519" ID-GBIF-Dataset="8713f3de-67ef-4823-929b-7b1b5d37e26a" ID-ISSN="0374-1036" ID-Zenodo-Dep="4503519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612461365170" checkinUser="felipe" docAuthor="Kočárek, Petr" docDate="2006" docId="A1441D06BF450305FEF6FA9CFBD8FA9F" docLanguage="en" docName="ActaEntMusNatPra.46.1-6.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 46" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Nala Zacher 1910" docType="key" docVersion="8" lastPageNumber="5" masterDocId="5D7D657EBF460301FFF2FFDEFFBFFFEA" masterDocTitle="A new species of Nala (Dermaptera: Labiduridae) from Cambodia" masterLastPageNumber="6" masterPageNumber="1" pageNumber="4" updateTime="1698927097416" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="382F85F95361F13BE3AD09DDB89813D0">A new species of Nala (Dermaptera: Labiduridae) from Cambodia</mods:title>
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<mods:date id="54DA59F63759DA89A58BE5A3FA43AC03">2006</mods:date>
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<mods:number id="289FAC8271D92026C0BFBDD2DEA18801">2006-11-06</mods:number>
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<treatment id="A1441D06BF450305FEF6FA9CFBD8FA9F" ID-DOI="http://doi.org/10.5281/zenodo.4503479" ID-GBIF-Taxon="180302135" ID-Zenodo-Dep="4503479" LSID="urn:lsid:plazi:treatment:A1441D06BF450305FEF6FA9CFBD8FA9F" httpUri="http://treatment.plazi.org/id/A1441D06BF450305FEF6FA9CFBD8FA9F" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="61F7FF9BBF450302FEF6FA9CFC11FAB6" box="[260,942,1346,1372]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="2952AC10BF450302FEF6FA9CFC11FAB6" blockId="3.[260,942,1346,1372]" box="[260,942,1346,1372]" pageId="3" pageNumber="4">
<heading id="721A1B7CBF450302FEF6FA9CFC11FAB6" bold="true" box="[260,942,1346,1372]" centered="true" fontSize="11" level="1" pageId="3" pageNumber="4" reason="2">
<emphasis id="1B997002BF450302FEF6FA9CFC11FAB6" bold="true" box="[260,942,1346,1372]" pageId="3" pageNumber="4">
Key to males of South-Eastern Asian species of
<taxonomicName id="EEEDD793BF450302FC86FA9CFC11FAB6" ID-CoL="8MSCY" authorityName="Zacher" authorityYear="1910" box="[884,942,1346,1372]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FC86FA9CFC11FAB6" bold="true" box="[884,942,1346,1372]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450305FF83FAA2FBD8FA9F" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="key">
<keyStep id="9219B4B5BF450302FF83FAA2FBFFFA3E" pageId="3" pageNumber="4">
<paragraph id="2952AC10BF450302FF83FAA2FBFFFA5E" blockId="3.[113,1090,1404,1556]" pageId="3" pageNumber="4">
<keyLead id="921C0F25BF450302FF83FAA2FBFFFA5E" pageId="3" pageNumber="4">
1 Forceps in basal third to basal half conspicuously dilated internally (
<figureCitation id="B1D6B095BF450302FC97FAA2FC1AFA7E" box="[869,933,1404,1428]" captionStart="Figs 6-13" captionStartId="4.[151,187,798,817]" captionTargetBox="[151,1058,220,764]" captionTargetId="figure-213@4.[151,1058,220,764]" captionTargetPageId="4" captionText="Figs 6-13. Right branches of forceps in South-East Asian species of Nala Zacher, 1910. 6 – N. tenuicornis (Bormans, 1900); 7 – N. timorensis Brindle, 1967; 8 – N. nepalensis (Burr, 1907); 9 – N. ornata Borelli, 1932; 10 – N. basalis Bey-Bienko, 1970; 11- N. lividipes (Dufour, 1829); 12 – N. alenae sp. nov. Scale bar = 1 mm. Adopted from BRINDLE (1966), BEY-BIENKO (1970), STEINMANN (1989b) and SRIVASTAVA (2003)." figureDoi="http://doi.org/10.5281/zenodo.4503525" httpUri="https://zenodo.org/record/4503525/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
). Distribution:
<collectingCountry id="51FAEC80BF450302FF62FA42FF52FA5E" box="[144,237,1436,1460]" name="Pakistan" pageId="3" pageNumber="4">Pakistan</collectingCountry>
,
<collectingCountry id="51FAEC80BF450302FF05FA42FE8FFA5E" box="[247,304,1436,1460]" name="India" pageId="3" pageNumber="4">India</collectingCountry>
,
<collectingCountry id="51FAEC80BF450302FEC9FA42FEC2FA5E" box="[315,381,1436,1460]" name="Nepal" pageId="3" pageNumber="4">Nepal</collectingCountry>
,
<collectingCountry id="51FAEC80BF450302FE7AFA42FE68FA5E" box="[392,471,1436,1460]" name="Bhutan" pageId="3" pageNumber="4">Bhutan</collectingCountry>
,
<collectingCountry id="51FAEC80BF450302FE11FA42FDF7FA5E" box="[483,584,1436,1460]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
. ..............................
<taxonomicName id="EEEDD793BF450302FCD3FA42FBFFFA5E" ID-CoL="6RSGW" authority="(Burr, 1907)" baseAuthorityName="Burr" baseAuthorityYear="1907" box="[801,1088,1435,1460]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="1B997002BF450302FCD3FA42FC0AFA59" bold="true" box="[801,949,1435,1459]" italics="true" pageId="3" pageNumber="4">N. nepalensis</emphasis>
(Burr, 1907)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2952AC10BF450302FF83FA62FBFFFA3E" blockId="3.[113,1090,1404,1556]" box="[113,1088,1468,1492]" pageId="3" pageNumber="4">
<keyLead id="921C0F25BF450302FF83FA62FBFFFA3E" box="[113,1088,1468,1492]" pageId="3" pageNumber="4">
– Forceps long and slender, not dilated in basal half ..........................................................
<emphasis id="1B997002BF450302FBC1FA62FBFFFA3E" bold="true" box="[1075,1088,1468,1492]" pageId="3" pageNumber="4">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9219B4B5BF450305FF83FA02FBD9FC3E" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<paragraph id="2952AC10BF450302FF83FA02FBFEF9FE" blockId="3.[113,1090,1404,1556]" pageId="3" pageNumber="4">
<keyLead id="921C0F25BF450302FF83FA02FBFEF9FE" pageId="3" pageNumber="4">
2 Each branch of forceps with large, distally oriented hooked tooth near inner base (
<figureCitation id="B1D6B095BF450302FC01FA02FB8CFA1E" box="[1011,1075,1500,1524]" captionStart="Figs 6-13" captionStartId="4.[151,187,798,817]" captionTargetBox="[151,1058,220,764]" captionTargetId="figure-213@4.[151,1058,220,764]" captionTargetPageId="4" captionText="Figs 6-13. Right branches of forceps in South-East Asian species of Nala Zacher, 1910. 6 – N. tenuicornis (Bormans, 1900); 7 – N. timorensis Brindle, 1967; 8 – N. nepalensis (Burr, 1907); 9 – N. ornata Borelli, 1932; 10 – N. basalis Bey-Bienko, 1970; 11- N. lividipes (Dufour, 1829); 12 – N. alenae sp. nov. Scale bar = 1 mm. Adopted from BRINDLE (1966), BEY-BIENKO (1970), STEINMANN (1989b) and SRIVASTAVA (2003)." figureDoi="http://doi.org/10.5281/zenodo.4503525" httpUri="https://zenodo.org/record/4503525/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
). Distribution: Timor. ...............................................................
<taxonomicName id="EEEDD793BF450302FCE6FA22FBFEF9FE" ID-CoL="45L5Y" authority="Brindle, 1967" authorityName="Brindle" authorityYear="1967" box="[788,1089,1532,1556]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="timorensis">
<emphasis id="1B997002BF450302FCE6FA22FC18F9FE" bold="true" box="[788,935,1532,1556]" italics="true" pageId="3" pageNumber="4">N. timorensis</emphasis>
Brindle, 1967
</taxonomicName>
</keyLead>
</paragraph>
<caption id="7D92FC98BF420305FF65FCC0FCC6FC6B" ID-DOI="http://doi.org/10.5281/zenodo.4503525" ID-Zenodo-Dep="4503525" httpUri="https://zenodo.org/record/4503525/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,187,798,817]" targetBox="[151,1058,220,764]" targetPageId="4">
<paragraph id="2952AC10BF420305FF65FCC0FCC6FC6B" blockId="4.[151,1128,798,897]" pageId="4" pageNumber="5">
Figs 6-13. Right branches of forceps in South-East Asian species of
<taxonomicName id="EEEDD793BF420305FD1BFCC0FC33FCDB" ID-CoL="8MSCY" authority="Zacher, 1910" authorityName="Zacher" authorityYear="1910" box="[745,908,798,817]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF420305FD1BFCC0FCADFCDB" box="[745,786,798,817]" italics="true" pageId="4" pageNumber="5">Nala</emphasis>
Zacher, 1910
</taxonomicName>
. 6 –
<taxonomicName id="EEEDD793BF420305FC4BFCC0FEBCFCA6" ID-CoL="45L5X" authority="(Bormans, 1900)" baseAuthorityName="Bormans" baseAuthorityYear="1900" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tenuicornis">
<emphasis id="1B997002BF420305FC4BFCC0FB8DFCDA" box="[953,1074,798,816]" italics="true" pageId="4" pageNumber="5">N. tenuicornis</emphasis>
(Bormans, 1900)
</taxonomicName>
; 7 –
<taxonomicName id="EEEDD793BF420305FEDFFCE7FDA5FCA6" ID-CoL="45L5Y" authority="Brindle, 1967" authorityName="Brindle" authorityYear="1967" box="[301,538,825,844]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="timorensis">
<emphasis id="1B997002BF420305FEDFFCE7FE22FCA1" box="[301,413,825,843]" italics="true" pageId="4" pageNumber="5">N. timorensis</emphasis>
Brindle, 1967
</taxonomicName>
; 8 –
<taxonomicName id="EEEDD793BF420305FDB7FCE7FC96FCA6" ID-CoL="6RSGW" authority="(Burr, 1907)" baseAuthorityName="Burr" baseAuthorityYear="1907" box="[581,809,825,844]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="1B997002BF420305FDB7FCE7FD08FCA6" box="[581,695,825,844]" italics="true" pageId="4" pageNumber="5">N. nepalensis</emphasis>
(Burr, 1907)
</taxonomicName>
; 9 –
<taxonomicName id="EEEDD793BF420305FCA6FCE7FBA1FCA6" ID-CoL="45L5V" authority="Borelli, 1932" authorityName="Borelli" authorityYear="1932" box="[852,1054,825,844]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="1B997002BF420305FCA6FCE7FC1AFCA1" box="[852,933,825,843]" italics="true" pageId="4" pageNumber="5">N. ornata</emphasis>
Borelli, 1932
</taxonomicName>
; 10 –
<taxonomicName id="EEEDD793BF420305FBA1FCE7FEC5FC8C" ID-CoL="45L5N" authority="Bey-Bienko, 1970" authorityName="Bey-Bienko" authorityYear="1970" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="1B997002BF420305FBA1FCE7FF6CFC8C" italics="true" pageId="4" pageNumber="5">N. basalis</emphasis>
<bibRefCitation id="4D7CD1E1BF420305FF28FC8DFEC5FC8C" author="BEY-BIENKO G. J." box="[218,378,851,870]" pageId="4" pageNumber="5" pagination="1710 - 1820" refId="ref2238" refString="BEY-BIENKO G. J. 1970: Novye i interesnye kozhistokrylye (Dermaptera) iz tropicheskoy i subtropicheskoy Azii. [New and interesting earwigs (Dermaptera) from tropical and subtropical Asia]. Zoologichesky Zhurnal 69: 1710 - 1820 (in Russian)." type="journal article" year="1970">Bey-Bienko, 1970</bibRefCitation>
</taxonomicName>
; 11-
<taxonomicName id="EEEDD793BF420305FE5AFC8AFD2AFC8C" ID-CoL="45L5S" authority="(Dufour, 1829)" baseAuthorityName="Dufour" baseAuthorityYear="1829" box="[424,661,851,870]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lividipes">
<emphasis id="1B997002BF420305FE5AFC8AFDB5FC8C" box="[424,522,851,870]" italics="true" pageId="4" pageNumber="5">N. lividipes</emphasis>
(Dufour, 1829)
</taxonomicName>
; 12 –
<taxonomicName id="EEEDD793BF420305FD3DFC8AFC9DFC8C" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[719,802,851,870]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF420305FD3DFC8AFC9DFC8C" box="[719,802,851,870]" italics="true" pageId="4" pageNumber="5">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF420305FCD8FC8AFCD1FC8D" box="[810,878,852,871]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
Scale bar = 1 mm. Adopted from
<bibRefCitation id="4D7CD1E1BF420305FF35FCB0FEF2FC6B" author="BRINDLE A." box="[199,333,878,897]" pageId="4" pageNumber="5" pagination="239 - 269" refId="ref2357" refString="BRINDLE A. 1966: A revision of the subfamily Labidurinae (Dermaptera, Labiduridae). Annals and Magazine of Natural History 13: 239 - 269." type="journal article" year="1966">BRINDLE (1966)</bibRefCitation>
, BEY- BIENKO (1970),
<bibRefCitation id="4D7CD1E1BF420305FDFAFCB0FD12FC6B" author="STEINMANN H." box="[520,685,878,897]" pageId="4" pageNumber="5" refId="ref2480" refString="STEINMANN H. 1989 b: Dermaptera Catadermaptera 2. Tierreich 105. Walter de Gruyter, Berlin - New York, 504 pp." type="book" year="1989">STEINMANN (1989b)</bibRefCitation>
and
<bibRefCitation id="4D7CD1E1BF420305FD2BFCB0FCCAFC6B" author="SRIVASTAVA G. K." box="[729,885,878,897]" pageId="4" pageNumber="5" refId="ref2425" refString="SRIVASTAVA G. K. 2003: Dermaptera. Part II. Superfamily: Anisolaboidea. Fauna of India and adjacent countries. Zoological Survey of India, Kolkata, 235 pp." type="book" year="2003">SRIVASTAVA (2003)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="2952AC10BF420305FF65FC63FBD9FC3E" blockId="4.[151,1128,956,1397]" box="[151,1126,956,981]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FC63FBD9FC3E" box="[151,1126,956,981]" pageId="4" pageNumber="5">
– Forceps without prominent hooked teeth (
<figureCitation id="B1D6B095BF420305FD81FC62FD45FC3F" box="[627,762,956,981]" captionStart-0="Figs 6-13" captionStart-1="Figs 1-5" captionStartId-0="4.[151,187,798,817]" captionStartId-1="2.[151,187,1563,1582]" captionTargetBox-0="[151,1058,220,764]" captionTargetBox-1="[170,1103,390,1539]" captionTargetId-0="figure-213@4.[151,1058,220,764]" captionTargetId-1="figure-10@2.[170,1103,390,1539]" captionTargetPageId-0="4" captionTargetPageId-1="2" captionText-0="Figs 6-13. Right branches of forceps in South-East Asian species of Nala Zacher, 1910. 6 – N. tenuicornis (Bormans, 1900); 7 – N. timorensis Brindle, 1967; 8 – N. nepalensis (Burr, 1907); 9 – N. ornata Borelli, 1932; 10 – N. basalis Bey-Bienko, 1970; 11- N. lividipes (Dufour, 1829); 12 – N. alenae sp. nov. Scale bar = 1 mm. Adopted from BRINDLE (1966), BEY-BIENKO (1970), STEINMANN (1989b) and SRIVASTAVA (2003)." captionText-1="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4503525" figureDoi-1="http://doi.org/10.5281/zenodo.4503521" httpUri-0="https://zenodo.org/record/4503525/files/figure.png" httpUri-1="https://zenodo.org/record/4503521/files/figure.png" pageId="4" pageNumber="5">Figs. 6, 9-12</figureCitation>
) .................................................
<emphasis id="1B997002BF420305FBABFC62FBD9FC3E" bold="true" box="[1113,1126,956,980]" pageId="4" pageNumber="5">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9219B4B5BF420305FF65FC03FBD8FBDE" pageId="4" pageNumber="5">
<paragraph id="2952AC10BF420305FF65FC03FBD9FC1E" blockId="4.[151,1128,956,1397]" box="[151,1126,988,1013]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FC03FBD9FC1E" box="[151,1126,988,1013]" pageId="4" pageNumber="5">
3 Tegmina long, 1.5-2 times as long as pronotum. Wings visible or concealed ..................
<emphasis id="1B997002BF420305FBABFC02FBD9FC1E" bold="true" box="[1113,1126,988,1012]" pageId="4" pageNumber="5">4</emphasis>
</keyLead>
</paragraph>
<paragraph id="2952AC10BF420305FF65FC23FBD8FBDE" blockId="4.[151,1128,956,1397]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FC23FBD8FBDE" pageId="4" pageNumber="5">
– Tegmina short, approximately as long as pronotum. Wings entirely concealed. Distribution:
<collectingCountry id="51FAEC80BF420305FF1CFBC2FEE0FBDE" box="[238,351,1052,1076]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
. .....................................................................................
<taxonomicName id="EEEDD793BF420305FC57FBC3FBB1FBDE" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[933,1038,1052,1076]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF420305FC57FBC3FBB1FBDE" bold="true" box="[933,1038,1052,1076]" italics="true" pageId="4" pageNumber="5">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF420305FBE7FBC3FBD8FBDE" box="[1045,1127,1053,1076]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9219B4B5BF420305FF65FBE3FBD9FB9E" pageId="4" pageNumber="5">
<paragraph id="2952AC10BF420305FF65FBE3FBD9FBBF" blockId="4.[151,1128,956,1397]" box="[151,1126,1084,1109]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FBE3FBD9FBBF" box="[151,1126,1084,1109]" pageId="4" pageNumber="5">
4 Dark brown species, head black or blackish brown. ........................................................
<emphasis id="1B997002BF420305FBABFBE3FBD9FBBF" bold="true" box="[1113,1126,1085,1109]" pageId="4" pageNumber="5">5</emphasis>
</keyLead>
</paragraph>
<paragraph id="2952AC10BF420305FF65FB83FBD9FB9E" blockId="4.[151,1128,956,1397]" box="[151,1126,1116,1140]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FB83FBD9FB9E" box="[151,1126,1116,1140]" pageId="4" pageNumber="5">
– Yellowish-brown to pale reddish-brown species (including head). ..................................
<emphasis id="1B997002BF420305FBABFB82FBD9FB9E" bold="true" box="[1113,1126,1116,1140]" pageId="4" pageNumber="5">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9219B4B5BF420305FF65FBA2FBD8FB1F" pageId="4" pageNumber="5">
<paragraph id="2952AC10BF420305FF65FBA2FBD9FB5F" blockId="4.[151,1128,956,1397]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FBA2FBD9FB5F" pageId="4" pageNumber="5">
5 Slender species, forceps twice as long as width of ultimate tergite (
<figureCitation id="B1D6B095BF420305FC7AFBA2FC74FB7E" box="[904,971,1148,1172]" captionStart="Figs 6-13" captionStartId="4.[151,187,798,817]" captionTargetBox="[151,1058,220,764]" captionTargetId="figure-213@4.[151,1058,220,764]" captionTargetPageId="4" captionText="Figs 6-13. Right branches of forceps in South-East Asian species of Nala Zacher, 1910. 6 – N. tenuicornis (Bormans, 1900); 7 – N. timorensis Brindle, 1967; 8 – N. nepalensis (Burr, 1907); 9 – N. ornata Borelli, 1932; 10 – N. basalis Bey-Bienko, 1970; 11- N. lividipes (Dufour, 1829); 12 – N. alenae sp. nov. Scale bar = 1 mm. Adopted from BRINDLE (1966), BEY-BIENKO (1970), STEINMANN (1989b) and SRIVASTAVA (2003)." figureDoi="http://doi.org/10.5281/zenodo.4503525" httpUri="https://zenodo.org/record/4503525/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
). Distribution:
<collectingCountry id="51FAEC80BF420305FF44FB42FE9EFB5E" box="[182,289,1180,1204]" name="Indonesia" pageId="4" pageNumber="5">Indonesia</collectingCountry>
. ........................................................................
<taxonomicName id="EEEDD793BF420305FCFDFB43FBD9FB5F" ID-CoL="45L5X" authority="(Bormans, 1900)" baseAuthorityName="Bormans" baseAuthorityYear="1900" box="[783,1126,1180,1205]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tenuicornis">
<emphasis id="1B997002BF420305FCFDFB43FC14FB5E" bold="true" box="[783,939,1180,1204]" italics="true" pageId="4" pageNumber="5">N. tenuicornis</emphasis>
(Bormans, 1900)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2952AC10BF420305FF65FB63FBD8FB1F" blockId="4.[151,1128,956,1397]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FB63FBD8FB1F" pageId="4" pageNumber="5">
– Stout species, forceps 1.1 to 1.4 times as long as width of ultimate tergite (
<figureCitation id="B1D6B095BF420305FC30FB62FBB3FB3F" box="[962,1036,1212,1237]" captionStart="Figs 6-13" captionStartId="4.[151,187,798,817]" captionTargetBox="[151,1058,220,764]" captionTargetId="figure-213@4.[151,1058,220,764]" captionTargetPageId="4" captionText="Figs 6-13. Right branches of forceps in South-East Asian species of Nala Zacher, 1910. 6 – N. tenuicornis (Bormans, 1900); 7 – N. timorensis Brindle, 1967; 8 – N. nepalensis (Burr, 1907); 9 – N. ornata Borelli, 1932; 10 – N. basalis Bey-Bienko, 1970; 11- N. lividipes (Dufour, 1829); 12 – N. alenae sp. nov. Scale bar = 1 mm. Adopted from BRINDLE (1966), BEY-BIENKO (1970), STEINMANN (1989b) and SRIVASTAVA (2003)." figureDoi="http://doi.org/10.5281/zenodo.4503525" httpUri="https://zenodo.org/record/4503525/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
). Distribution: Europe, Africa, Asia. ...................................................
<taxonomicName id="EEEDD793BF420305FCB7FB03FBD8FB1F" ID-CoL="45L5S" authority="(Dufour, 1829)" baseAuthorityName="Dufour" baseAuthorityYear="1829" box="[837,1127,1244,1269]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lividipes">
<emphasis id="1B997002BF420305FCB7FB03FC7FFB1E" bold="true" box="[837,960,1244,1268]" italics="true" pageId="4" pageNumber="5">N. lividipes</emphasis>
(Dufour, 1829)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9219B4B5BF420305FF65FB22FBD8FA9F" pageId="4" pageNumber="5">
<paragraph id="2952AC10BF420305FF65FB22FBD8FAFF" blockId="4.[151,1128,956,1397]" box="[151,1127,1276,1301]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FB22FBD8FAFF" box="[151,1127,1276,1301]" pageId="4" pageNumber="5">
6 Forceps with two pairs of small teeth on inner margin (
<figureCitation id="B1D6B095BF420305FD0EFB22FC84FAFF" box="[764,827,1276,1301]" captionStart="Figs 6-13" captionStartId="4.[151,187,798,817]" captionTargetBox="[151,1058,220,764]" captionTargetId="figure-213@4.[151,1058,220,764]" captionTargetPageId="4" captionText="Figs 6-13. Right branches of forceps in South-East Asian species of Nala Zacher, 1910. 6 – N. tenuicornis (Bormans, 1900); 7 – N. timorensis Brindle, 1967; 8 – N. nepalensis (Burr, 1907); 9 – N. ornata Borelli, 1932; 10 – N. basalis Bey-Bienko, 1970; 11- N. lividipes (Dufour, 1829); 12 – N. alenae sp. nov. Scale bar = 1 mm. Adopted from BRINDLE (1966), BEY-BIENKO (1970), STEINMANN (1989b) and SRIVASTAVA (2003)." figureDoi="http://doi.org/10.5281/zenodo.4503525" httpUri="https://zenodo.org/record/4503525/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
)....
<taxonomicName id="EEEDD793BF420305FC9BFB23FBD8FAFF" ID-CoL="45L5V" authority="Borelli, 1932" authorityName="Borelli" authorityYear="1932" box="[873,1127,1276,1301]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="1B997002BF420305FC9BFB23FC6DFAFE" bold="true" box="[873,978,1277,1300]" italics="true" pageId="4" pageNumber="5">N. ornata</emphasis>
Borelli, 1932
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2952AC10BF420305FF65FAC3FBD8FA9F" blockId="4.[151,1128,956,1397]" pageId="4" pageNumber="5">
<keyLead id="921C0F25BF420305FF65FAC3FBD8FA9F" pageId="4" pageNumber="5">
– Each branch of forceps with cylindrical tooth (not hooked) on inner margin close to base (
<figureCitation id="B1D6B095BF420305FF4FFAE2FEB4FABF" box="[189,267,1340,1365]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
), entire inner margin sparsely serrated. Distribution:
<collectingCountry id="51FAEC80BF420305FCCBFAE2FC02FABE" box="[825,957,1340,1364]" name="Afghanistan" pageId="4" pageNumber="5">Afghanistan</collectingCountry>
,
<collectingCountry id="51FAEC80BF420305FC35FAE2FB9CFABE" box="[967,1059,1340,1364]" name="Pakistan" pageId="4" pageNumber="5">Pakistan</collectingCountry>
,
<collectingCountry id="51FAEC80BF420305FBDFFAE2FF58FA9E" name="Thailand" pageId="4" pageNumber="5">Thailand</collectingCountry>
. ......................................................................................
<taxonomicName id="EEEDD793BF420305FCC3FA83FBD8FA9F" ID-CoL="45L5N" authority="Bey-Bienko, 1970" authorityName="Bey-Bienko" authorityYear="1970" box="[817,1127,1372,1397]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="1B997002BF420305FCC3FA83FC23FA9E" bold="true" box="[817,924,1372,1396]" italics="true" pageId="4" pageNumber="5">N. basalis</emphasis>
<bibRefCitation id="4D7CD1E1BF420305FC56FA82FBD8FA9F" author="BEY-BIENKO G. J." box="[932,1127,1372,1397]" pageId="4" pageNumber="5" pagination="1710 - 1820" refId="ref2238" refString="BEY-BIENKO G. J. 1970: Novye i interesnye kozhistokrylye (Dermaptera) iz tropicheskoy i subtropicheskoy Azii. [New and interesting earwigs (Dermaptera) from tropical and subtropical Asia]. Zoologichesky Zhurnal 69: 1710 - 1820 (in Russian)." type="journal article" year="1970">Bey-Bienko, 1970</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

317
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<emphasis id="1B997002BF470300FE1BFD64FDD2FD38" bold="true" box="[489,621,698,722]" italics="true" pageId="1" pageNumber="2">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF470300FD86FD64FD75FD38" box="[628,714,698,722]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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(
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)
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locality.
</emphasis>
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,
<collectingRegion id="EB2962F2BF470300FEA6FCCEFE0CFCC9" box="[340,435,784,803]" country="Cambodia" name="Siem Reap" pageId="1" pageNumber="2">Siem Reap</collectingRegion>
, Angkor Wat temple.
</paragraph>
<paragraph id="2952AC10BF470300FF83FCF4FD12FC99" blockId="1.[113,1089,784,883]" pageId="1" pageNumber="2">
<emphasis id="1B997002BF470300FF83FCF4FF4BFCD7" bold="true" box="[113,244,810,829]" pageId="1" pageNumber="2">Type material.</emphasis>
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<typeStatus id="F65612B2BF470300FF0BFCF5FEF4FCD6" box="[249,331,811,830]" pageId="1" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
: ♁, ‘NW
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,
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env. /
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temple;
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a.s.l.
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,
<geoCoordinate id="4CD9CAD7BF470300FF83FC9BFF45FCB3" box="[113,250,836,857]" degrees="103" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="2" precision="1" seconds="49.84" value="103.863846">103°51′49.84″E</geoCoordinate>
/
<collectorName id="8418C9C6BF470300FEFDFC98FED0FCB2" box="[271,367,837,857]" pageId="1" pageNumber="2">P. Kočárek</collectorName>
leg.;
<date id="5D538AD0BF470300FE51FC98FDA8FCB2" box="[419,535,837,857]" pageId="1" pageNumber="2" value="2002-09-07" valueMax="2002-09-11" valueMin="2002-09-07">
<collectingDate id="4D177338BF470300FE51FC98FDA8FCB2" box="[419,535,837,857]" pageId="1" pageNumber="2" value="2002-09-07" valueMax="2002-09-11" valueMin="2002-09-07">7.-11.ix.2002</collectingDate>
</date>
’ (
<collectionCode id="4FFC34D5BF470300FDDAFC9BFDD0FCB2" box="[552,623,837,856]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="2">MMBC</collectionCode>
)
</materialsCitation>
;
<typeStatus id="F65612B2BF470300FD72FC98FD69FCB3" box="[640,726,838,857]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FEB6799BF470300FD12FC9BFCB4FCB3" box="[736,779,833,857]" pageId="1" pageNumber="2" type="female">5 ♀♀</specimenCount>
, same label data as
<typeStatus id="F65612B2BF470300FC3BFC9BFBACFCB2" box="[969,1043,837,856]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
(one specimen
<collectionCode id="4FFC34D5BF470300FF3AFCBEFEB3FC99" box="[200,268,864,883]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="2">MMBC</collectionCode>
, one specimen
<collectionCode id="4FFC34D5BF470300FE63FCBEFE71FC99" box="[401,462,864,883]" pageId="1" pageNumber="2">ZSMG</collectionCode>
, three specimens
<collectionCode id="4FFC34D5BF470300FD95FCBEFD1CFC99" box="[615,675,864,883]" pageId="1" pageNumber="2">PKOC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF470302FF83FC57FBABFE5F" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="2952AC10BF470300FF83FC57FD31FBCB" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
<emphasis id="1B997002BF470300FF83FC57FEE8FC48" bold="true" box="[113,343,905,930]" pageId="1" pageNumber="2">Description of male.</emphasis>
Body dark reddish brown; head entirely black, antennae dark brown; pronotum and elytra reddish brown to brown with paler sides; legs yellowish brown, femora and tibiae with dark brown basal parts; ultimate abdominal tergite dark brown; forceps dark brown to reddish brown at apex. Cuticle punctured, dull, pubescent; tegmina rugose. Length without forceps
<quantity id="EE1501F5BF470300FEEFFBD4FECEFBCB" box="[285,369,1034,1058]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="2" unit="mm" value="7.3">7.3 mm</quantity>
, length of forceps
<quantity id="EE1501F5BF470300FDC7FBD4FD35FBCB" box="[565,650,1034,1058]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="2" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
<paragraph id="2952AC10BF470300FF62FBF7FC02FB0B" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
Head globose, as wide as long, postfrontal and coronal sutures indistinct, posterior margin straight, with short and dense pubescence. Eyes small, less than 0.5 times as long as length of head behind eyes. Antennae of
<typeStatus id="F65612B2BF470300FE4DFBB7FDA3FB6B" box="[447,540,1129,1153]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
male with 17 antennomeres (incomplete); antennomere 1 short, narrowed basally, widened terminally, half as long as the distance between antennal bases; antennomere 2 transverse, only little longer than wide; antennomere 3 longer than antennomere 4. Antennomeres 1-3 cylindrical, antennomeres 4-17 conical.
</paragraph>
<paragraph id="2952AC10BF470300FF62FB34FE3AFA4B" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
Pronotum transverse, widened posteriorly.Anterior margin straight, posterior margin broadly rounded. Pronotal disc rugose, with smoother anterior half, median suture distinct. Tegmina as long as pronotum, rugose, parallel-sided; each with well marked lateral longitudinal ridge, tegminal disc flattened; posterior margin convex, with broadly rounded interior angle. Wings entirely concealed under tegmina, reduced to very small, longitudinally oriented flaps (
<quantity id="EE1501F5BF470300FBE3FAB7FF25FA4B" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="2" unit="mm" value="0.25">0.25 mm</quantity>
long,
<quantity id="EE1501F5BF470300FF2FFA57FE84FA4B" box="[221,315,1417,1441]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="2" unit="mm" value="0.05">0.05 mm</quantity>
wide).
</paragraph>
<paragraph id="2952AC10BF470300FF62FA77FED6F98B" blockId="1.[113,1089,905,1633]" pageId="1" pageNumber="2">
Abdomen convex, somewhat widened posteriorly, densely punctured, shortly pubescent with long setae along hind margin of each tergite except the ultimate one, lateral glandular folds invisible. Penultimate sternite punctulate, sides scarcely concave, hind margin obtusely produced in middle (
<figureCitation id="B1D6B095BF470300FEA4F9D7FE25F9C8" box="[342,410,1545,1570]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
). Ultimate tergite transverse, slightly depressed medially, with invisible median line, little oblique laterally, hind margin straight with protuberance at each base of forceps branch.
</paragraph>
<paragraph id="2952AC10BF440303FF44FF01FBF2FEBD" blockId="2.[151,1127,223,343]" pageId="2" pageNumber="3">
Pygidium invisible in dorsal view. Forceps (
<figureCitation id="B1D6B095BF440303FD66FF01FD44FF1D" box="[660,763,223,247]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 2</figureCitation>
) with branches remote, gradually incurved, each tapering to pointed and gently hooked apex, triangular in basal half and dorsoventrally flattened in distal half. Inner and upper margins of forceps serrated in proximal third, middle of forceps with distinct tapering, distal third widened into flat protuberance.
</paragraph>
<caption id="7D92FC98BF440303FF65F9C5FED6F98E" ID-DOI="http://doi.org/10.5281/zenodo.4503521" ID-Zenodo-Dep="4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="2" pageNumber="3" startId="2.[151,187,1563,1582]" targetBox="[170,1103,390,1539]" targetPageId="2">
<paragraph id="2952AC10BF440303FF65F9C5FED6F98E" blockId="2.[151,1127,1563,1636]" pageId="2" pageNumber="3">
Figs 1-5.
<taxonomicName id="EEEDD793BF440303FF1BF9C5FEEFF9C4" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[233,336,1563,1582]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF440303FF1BF9C5FEEFF9C4" box="[233,336,1563,1582]" italics="true" pageId="2" pageNumber="3">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF440303FEA5F9C5FE25F9C4" box="[343,410,1563,1582]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="2952AC10BF450302FF62FF03FC5EFEDF" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
Genitalia (
<figureCitation id="B1D6B095BF450302FF0DFF03FE80FF1E" box="[255,319,220,245]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
) typical for the genus
<taxonomicName id="EEEDD793BF450302FDDCFF03FDDEFF1F" authorityName="Zacher" authorityYear="1910" box="[558,609,221,245]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FDDCFF03FDDEFF1F" box="[558,609,221,245]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
, with parameres a little over six times longer than broad, narrowed apically, tip acuminate; genital lobes widening to the apex, with straight apical part; virga tubular with undulated central part and sclerotized basal vesicles.
</paragraph>
<paragraph id="2952AC10BF450302FF83FEE2FBABFE5F" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FEE2FED3FEBE" bold="true" box="[113,364,316,340]" pageId="3" pageNumber="4">Description of female.</emphasis>
Body length without forceps
<quantity id="EE1501F5BF450302FD5FFEE3FC93FEBE" box="[685,812,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.9" metricValueMin="6.8" pageId="3" pageNumber="4" unit="mm" value="7.35" valueMax="7.9" valueMin="6.8">6.8-7.9 mm</quantity>
, length of forceps
<quantity id="EE1501F5BF450302FC04FEE3FF21FE9E" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="3" pageNumber="4" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4-1.5 mm</quantity>
. Body colouration similar to
<typeStatus id="F65612B2BF450302FE12FE83FD82FE9F" box="[480,573,349,373]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
male, abdomen wider in middle part. Ultimate tergite with tapering sides, hind margin sinuate in middle. Forceps straight, simple, tapering, inner margin serrated, triangular in basal third, the rest dorsoventrally flattened (
<figureCitation id="B1D6B095BF450302FC34FE43FBB9FE5F" box="[966,1030,413,437]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FE62FC7AFCDF" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="2952AC10BF450302FF83FE62FC7AFCDF" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FE62FED2FE3E" bold="true" box="[113,365,444,468]" pageId="3" pageNumber="4">Differential diagnosis.</emphasis>
<taxonomicName id="EEEDD793BF450302FE84FE63FE45FE3F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[374,506,445,469]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FE84FE63FE45FE3F" box="[374,506,445,469]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FDF6FE63FDE6FE3E" box="[516,601,445,468]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="EEEDD793BF450302FD61FE63FC04FE3F" authority="(Dufour, 1829)" baseAuthorityName="Dufour" baseAuthorityYear="1829" box="[659,955,445,469]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lividipes">
<emphasis id="1B997002BF450302FD61FE63FCB0FE3F" box="[659,783,445,469]" italics="true" pageId="3" pageNumber="4">N. lividipes</emphasis>
(Dufour, 1829)
</taxonomicName>
are the two smallest species of this genus known from South-East Asia, with total body length including forceps ranging between
<quantity id="EE1501F5BF450302FE74FE22FDBBFDFE" box="[390,516,508,532]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="3" pageNumber="4" unit="mm" value="8.0" valueMax="8.5" valueMin="7.5">7.5-8.5 mm</quantity>
.
<taxonomicName id="EEEDD793BF450302FDE7FE23FD26FDFF" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[533,665,509,533]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FDE7FE23FD26FDFF" box="[533,665,509,533]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FD51FE23FD46FDFE" box="[675,761,509,532]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="EEEDD793BF450302FBDAFE23FF75FDDF" baseAuthorityName="Dufour" baseAuthorityYear="1829" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lividipes">
<emphasis id="1B997002BF450302FBDAFE23FF75FDDF" italics="true" pageId="3" pageNumber="4">N. lividipes</emphasis>
</taxonomicName>
by the very short tegmina, which are only as long as the pronotum, and the wings entirely concealed under the tegmina.
<taxonomicName id="EEEDD793BF450302FDE0FDE3FCE4FDBF" authority="Brindle, 1967" authorityName="Brindle" authorityYear="1967" box="[530,859,573,597]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="timorensis">
<emphasis id="1B997002BF450302FDE0FDE3FD04FDBE" box="[530,699,573,597]" italics="true" pageId="3" pageNumber="4">Nala timorensis</emphasis>
Brindle, 1967
</taxonomicName>
, and
<taxonomicName id="EEEDD793BF450302FC65FDE3FBBBFDBF" authorityName="Bey-Bienko" authorityYear="1970" box="[919,1028,573,597]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="1B997002BF450302FC65FDE3FBBBFDBF" box="[919,1028,573,597]" italics="true" pageId="3" pageNumber="4">N. basalis</emphasis>
</taxonomicName>
Bey-Bienko, 1970, also have concealed wings; however, winged specimens were described for the latter species (
<bibRefCitation id="4D7CD1E1BF450302FEE2FDA3FE7BFD7F" author="SRIVASTAVA G. K." box="[272,452,637,661]" pageId="3" pageNumber="4" refId="ref2425" refString="SRIVASTAVA G. K. 2003: Dermaptera. Part II. Superfamily: Anisolaboidea. Fauna of India and adjacent countries. Zoological Survey of India, Kolkata, 235 pp." type="book" year="2003">SRIVASTAVA 2003</bibRefCitation>
). The forceps of
<taxonomicName id="EEEDD793BF450302FD70FDA3FD54FD7F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[642,747,637,661]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FD70FDA3FD54FD7F" box="[642,747,637,661]" italics="true" pageId="3" pageNumber="4">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FD06FDA3FCF5FD7E" box="[756,842,637,660]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
are unusually short in males, being only as long as the width of the ultimate tergite.
<taxonomicName id="EEEDD793BF450302FCF2FD43FC3EFD5F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[768,897,669,693]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FCF2FD43FC3EFD5F" box="[768,897,669,693]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FC7AFD43FC65FD5E" box="[904,986,669,692]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is further characterized by the entirely black head. Although the females of
<taxonomicName id="EEEDD793BF450302FCDDFD63FCDDFD3F" authorityName="Zacher" authorityYear="1910" box="[815,866,701,725]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FCDDFD63FCDDFD3F" box="[815,866,701,725]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
are morphologically very similar and cannot be reliably identified, the females of
<taxonomicName id="EEEDD793BF450302FCE2FD03FCC8FD1F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[784,887,733,757]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FCE2FD03FCC8FD1F" box="[784,887,733,757]" italics="true" pageId="3" pageNumber="4">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FC73FD03FC69FD1E" box="[897,982,733,756]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
are fairly easily recognized by their exceptionally short tegmina and the absence of wings (see above). The males of South-East Asia
<taxonomicName id="EEEDD793BF450302FE41FCC3FE59FCDF" authorityName="Zacher" authorityYear="1910" box="[435,486,797,821]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FE41FCC3FE59FCDF" box="[435,486,797,821]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
can be separated by the key provided below.
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FCE3FDA0FC3F" pageId="3" pageNumber="4" type="etymology">
<paragraph id="2952AC10BF450302FF83FCE3FF58FC9F" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FCE3FF4EFCBF" bold="true" box="[113,241,829,853]" pageId="3" pageNumber="4">Etymology.</emphasis>
The new species is dedicated to my wife Alena, who helped me collect the
<typeStatus id="F65612B2BF450302FBE1FCE3FBFEFCBF" box="[1043,1089,829,853]" pageId="3" pageNumber="4">type</typeStatus>
specimens.
</paragraph>
<paragraph id="2952AC10BF450302FF83FCA2FDA0FC3F" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FCA2FF51FC7E" bold="true" box="[113,238,892,916]" pageId="3" pageNumber="4">Bionomics.</emphasis>
Unknown.
<typeStatus id="F65612B2BF450302FE9FFCA3FE1DFC7F" box="[365,418,893,917]" pageId="3" pageNumber="4">Type</typeStatus>
specimens have been collected in the surroundings of Angkor Wat temples in a stony habitat surrounded by lowland rain forest. The earwigs were observed to run on stony ground in late afternoon.
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FC02FEC1FC1F" box="[113,382,988,1013]" pageId="3" pageNumber="4" type="distribution">
<paragraph id="2952AC10BF450302FF83FC02FEC1FC1F" blockId="3.[113,1090,220,1301]" box="[113,382,988,1013]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FC02FEBDFC1E" bold="true" box="[113,258,988,1012]" pageId="3" pageNumber="4">Distribution.</emphasis>
<collectingCountry id="51FAEC80BF450302FEF8FC03FEC5FC1F" box="[266,378,989,1013]" name="Cambodia" pageId="3" pageNumber="4">Cambodia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61F7FF9BBF450302FF83FC22FD3CFAFF" pageId="3" pageNumber="4" type="discussion">
<paragraph id="2952AC10BF450302FF83FC22FD3CFAFF" blockId="3.[113,1090,220,1301]" pageId="3" pageNumber="4">
<emphasis id="1B997002BF450302FF83FC22FF4DFBFE" bold="true" box="[113,242,1020,1044]" pageId="3" pageNumber="4">Comments.</emphasis>
<taxonomicName id="EEEDD793BF450302FF09FC23FE3FFBFF" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[251,384,1021,1045]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alanae" status="sp. nov.">
<emphasis id="1B997002BF450302FF09FC23FE3FFBFF" box="[251,384,1021,1045]" italics="true" pageId="3" pageNumber="4">Nala alanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FE78FC23FE60FBFE" box="[394,479,1021,1044]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is a very characteristic species, differing from all other Asian species in the shape of the reduced tegmina with well marked longitudinal ridges (see
<figureCitation id="B1D6B095BF450302FF83FBE3FF0FFBBF" box="[113,176,1085,1109]" captionStart="Figs 1-5" captionStartId="2.[151,187,1563,1582]" captionTargetBox="[170,1103,390,1539]" captionTargetId="figure-10@2.[170,1103,390,1539]" captionTargetPageId="2" captionText="Figs 1-5. Nala alenae sp. nov. 1 – habitus of male, dorsal view; 2 – right branch of male forceps, dorsal view; 3 – right branch of female forceps, dorsal view; 4 – subgenital plate of male, dorsal view; 5 – male genitalia, dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503521" httpUri="https://zenodo.org/record/4503521/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
). Other
<taxonomicName id="EEEDD793BF450302FEF9FBE3FE81FBBF" authorityName="Zacher" authorityYear="1910" box="[267,318,1085,1109]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="1B997002BF450302FEF9FBE3FE81FBBF" box="[267,318,1085,1109]" italics="true" pageId="3" pageNumber="4">Nala</emphasis>
</taxonomicName>
species with similar tegmina, reduced wings and short stout forceps are known only from Central and
<collectingCountry id="51FAEC80BF450302FE41FB83FD84FB9F" box="[435,571,1117,1141]" name="South Africa" pageId="3" pageNumber="4">South Africa</collectingCountry>
(
<taxonomicName id="EEEDD793BF450302FDB8FB83FCE8FB9F" authority="Menozzi, 1929" authorityName="Menozzi" authorityYear="1929" box="[586,855,1117,1141]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caprai">
<emphasis id="1B997002BF450302FDB8FB83FD11FB9E" box="[586,686,1117,1140]" italics="true" pageId="3" pageNumber="4">N. caprai</emphasis>
Menozzi, 1929
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FC90FB83FB83FB9F" authority="Burr, 1908" authorityName="Burr" authorityYear="1908" box="[866,1084,1117,1141]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="figinii">
<emphasis id="1B997002BF450302FC90FB83FC7EFB9F" box="[866,961,1117,1141]" italics="true" pageId="3" pageNumber="4">N. figinii</emphasis>
Burr, 1908
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FF83FBA3FE14FB7F" authority="Menozzi, 1937" authorityName="Menozzi" authorityYear="1937" box="[113,427,1149,1173]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="1B997002BF450302FF83FBA3FEBCFB7F" box="[113,259,1149,1173]" italics="true" pageId="3" pageNumber="4">N. intermedia</emphasis>
Menozzi, 1937
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FE44FBA3FD02FB7F" authority="Brindle, 1968" authorityName="Brindle" authorityYear="1968" box="[438,701,1149,1173]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="1B997002BF450302FE44FBA3FD9CFB7E" box="[438,547,1149,1172]" italics="true" pageId="3" pageNumber="4">N. saegeri</emphasis>
Brindle, 1968
</taxonomicName>
,
<taxonomicName id="EEEDD793BF450302FD3AFBA3FBB0FB7F" authority="Vigna Taglianti, 1994" authorityName="Vigna Taglianti" authorityYear="1994" box="[712,1039,1149,1173]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rossii">
<emphasis id="1B997002BF450302FD3AFBA3FC9EFB7E" box="[712,801,1149,1172]" italics="true" pageId="3" pageNumber="4">N. rossii</emphasis>
<bibRefCitation id="4D7CD1E1BF450302FCDAFBA3FBB0FB7F" author="VIGNA TAGLIANTI A." box="[808,1039,1149,1173]" pageId="3" pageNumber="4" pagination="199 - 212" refId="ref2505" refString="VIGNA TAGLIANTI A. 1994: Further notes on Dermaptera from Sierra Leone. Quaderni dell'Accademia Nazionale dei Lincei 267: 199 - 212." type="journal article" year="1994">Vigna Taglianti, 1994</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="EEEDD793BF450302FF83FB43FE6BFB5F" authority="Bivar de Sousa, 1999" authorityName="Bivar de Sousa" authorityYear="1999" box="[113,468,1181,1205]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mendesi">
<emphasis id="1B997002BF450302FF83FB43FF58FB5F" box="[113,231,1181,1205]" italics="true" pageId="3" pageNumber="4">N. mendesi</emphasis>
<bibRefCitation id="4D7CD1E1BF450302FF1FFB43FE6BFB5F" author="BIVAR DE SOUSA A." box="[237,468,1181,1205]" pageId="3" pageNumber="4" pagination="141 - 146" refId="ref2285" refString="BIVAR DE SOUSA A. 1999: Descricao de uma nova especie de dermaptero de Guine Bissau. Nala mendesi sp. nov. (Insecta, Dermaptera). (Description of a new species of Dermaptera from Guinea-Bissau. Nala mendesi n. sp. (Insecta, Dermaptera )). Boletim da Sociedade Portuguesa de Entomologia 194: 141 - 146 (in Portugese, English summary)." type="journal article" year="1999">Bivar de Sousa, 1999</bibRefCitation>
</taxonomicName>
).
<taxonomicName id="EEEDD793BF450302FE14FB43FDD9FB5F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[486,614,1181,1205]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FE14FB43FDD9FB5F" box="[486,614,1181,1205]" italics="true" pageId="3" pageNumber="4">Nala alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FD9FFB43FD00FB5E" box="[621,703,1181,1204]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
is the only known Asian representative with a noticeable morphological similarity to this African group of species, which may indicate some affinity to this group. From the known African species,
<taxonomicName id="EEEDD793BF450302FC88FB03FC5CFB1F" authority="Kočárek, 2006" authorityName="Kočárek" authorityYear="2006" box="[890,995,1245,1269]" class="Insecta" family="Labiduridae" genus="Nala" kingdom="Animalia" order="Dermaptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alenae" status="sp. nov.">
<emphasis id="1B997002BF450302FC88FB03FC5CFB1F" box="[890,995,1245,1269]" italics="true" pageId="3" pageNumber="4">N. alenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AACD79BF450302FC1EFB03FBFEFB1E" box="[1004,1089,1245,1268]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
differs in the characteristic shape of male forceps.
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70
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, 1869
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<paragraph id="0BF737D1196369A9F8456D2287ED6628" pageId="0" pageNumber="9042">Notes</paragraph>
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Some distribution data from
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and
<bibRefCitation id="9B7E38FA543D0A33B9BD8632C4150206" author="Horstmann, K." journalOrPublisher="Entomofauna" pageId="0" pageNumber="9042" pagination="85 - 148" title="Revision der brachypteren Weibchen der wespalaearktischen Cryptinae (Hymenoptera, Ichneumonidae)" volume="14" year="1993">Horstmann (1993c)</bibRefCitation>
.
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173
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<taxonomicName id="D9CF582BD140B5C4E2D98E608AAFC186" ID-CoL="4659H" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:32851" class="Insecta" family="Formicidae" genus="Neivamyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Neivamyrmex pauxillus (Wheeler)" order="Hymenoptera" pageNumber="487" phylum="Arthropoda" rank="species" species="pauxillus">Neivamyrmex pauxillus (W. M. Wheeler)</taxonomicName>
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<paragraph id="8679D50FD269E295E3973D29E69D5DCC" pageNumber="487">Figures 12, 24, 43, 58, 71, 85</paragraph>
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<paragraph id="4AFAE1ED904651802E8061587F3EA701" pageNumber="487">
<taxonomicName id="8794D388C38A11F7A587B2B2A823E936" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:140805" class="Insecta" family="Formicidae" genus="Eciton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Eciton (Acamatus) pauxillum Wheeler" order="Hymenoptera" pageNumber="487" phylum="Arthropoda" rank="species" species="pauxillum" subGenus="Acamatus">Eciton (Acamatus) pauxillum Wheeler</taxonomicName>
, 1903: 93 (w).
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<collectingCountry id="9C9269C3E3E408CB5D0E328394D92F14">U. S.</collectingCountry>
A,
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,
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,
<location id="5FFBC1C4E6112A60262CD2A3BA9D7721" LSID="plazi:21B04C20D6886114A7E0EDB794B86009:295C99ED6171612866548B416AC69593" country="U. S. A," name="Paisano Pass" stateProvince="Texas">Paisano Pass</location>
(
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<collectionCode id="29AB3ABFB012F35416AD1C114B6F24D2" collectionName="USA, New York, New York, American Museum of Natural History">AMNH</collectionCode>
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,
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.
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<taxonomicName id="3778C87A614BA80BE559E9D61E7F9003" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:140805" class="Insecta" family="Formicidae" genus="Eciton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Eciton (Acamatus) pauxillum Wheeler" order="Hymenoptera" pageNumber="487" phylum="Arthropoda" rank="species" species="pauxillum" subGenus="Neivamyrmex">Eciton (Neivamyrmex) pauxillum</taxonomicName>
: M. R. Smith, 1942: 569
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<paragraph id="3FBC854E78E17EC177E305D55EFA95C0" pageNumber="487">
<taxonomicName id="16EBDD2428430C918D2CE4917B1B07AE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:32851" class="Insecta" family="Formicidae" genus="Neivamyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Neivamyrmex pauxillus (Wheeler)" order="Hymenoptera" pageNumber="487" phylum="Arthropoda" rank="species" species="pauxillus">Neivamyrmex pauxillus</taxonomicName>
: Borgmeier, 1953: 570 - 572. Watkins, 1982: 197 - 247. Watkins, 1985: 479 - 502.
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<paragraph id="344C5570653A3FFD1C67980911D702E2" pageNumber="487">DISTRIBUTION (Map 5)</paragraph>
<paragraph id="909B3F4BF678C8D3114F4019EA31F0D7" pageNumber="487">UNITED STATES: Louisiana and Texas; MEXICO: Hidalgo (Watkins, 1982).</paragraph>
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<paragraph id="D6B598CDE8FCB1FD01DA817F1D4D68D5" pageNumber="487">SPECIMENS EXAMINED</paragraph>
<paragraph id="2553760C75CA5CA7B5F9B9D701AF6C6E" pageNumber="487">
<materialsCitation id="F894B206830A1EFBE8512D4815489E34" ID-GBIF-Occurrence="923900064" collectionCode="LACM" country="United States" location="Kisatchue National Forest nr. Alexandria" municipality="Rapides Parish" stateProvince="LOUISIANA">
<collectingCountry id="905D688F2870F4D18BE6A3E4D6F6B64E">UNITED STATES</collectingCountry>
,
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,
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:
<location id="9D096EBB2BAFB1C497FEBF6983BD81D9" LSID="plazi:21B04C20D6886114A7E0EDB794B86009:08D055DFAF513A8E272BCF6D21D1A7C2" country="United States" municipality="Rapides Parish" name="Kisatchue National Forest nr. Alexandria" stateProvince="LOUISIANA">Kisatchue National Forest nr. Alexandria</location>
(
<collectionCode id="BB640A8C61DDA00F7D84263B797435E9" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
)
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.
<materialsCitation id="2B99D99DE589FFD40E12BAC5463AE49F" ID-GBIF-Occurrence="923900079" collectionCode="LACM, WPMC" country="UNITED STATES" county="Bell Co." location="Bowmer Ranch" municipality="Bell Co." stateProvince="TEXAS">
<collectingRegion id="DFBCBC90D40DA5863B8DF30CEA58882E">TEXAS</collectingRegion>
,
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:
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(
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<collectionCode id="12821CDE5DB6D7549B8641EBADE2537F">WPMC</collectionCode>
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.
<materialsCitation id="0DF2F5E5FFCAE97E31759A1657AC7B1E" ID-GBIF-Occurrence="923900082" collectionCode="MCZC" country="UNITED STATES" location="Paisano Pass" municipality="Brewster Co." stateProvince="Brewster Co.">
<collectingMunicipality id="28D46033DA6B4A007A9BE84F4B5014C1">Brewster Co.</collectingMunicipality>
:
<location id="C188132BEBDF52D5BA44E4210D0E76BB" LSID="plazi:21B04C20D6886114A7E0EDB794B86009:18FC25CA2C7197C44EB5C3BACBD5C3CC" country="United States" municipality="Brewster Co." name="Paisano Pass" stateProvince="TEXAS">Paisano Pass</location>
, nr.
<location id="75BAB9B2DE6C58811D175E6AFCE10CD9" LSID="plazi:21B04C20D6886114A7E0EDB794B86009:239CB385652132F36858539CDBA45182" country="United States" municipality="Brewster Co." name="Alpine" stateProvince="TEXAS">Alpine</location>
(
<collectionCode id="E8BABB6AC0315F8F13D9D0DAECD09BCD">MCZC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="65CBC918AEDB0114D2112B0F88792002" ID-GBIF-Occurrence="923900094" collectionCode="AMNH, MCZC" country="UNITED STATES" county="Travis Co." location="Austin" municipality="Travis Co." stateProvince="Travis Co">
<collectingMunicipality id="B31D736CC5BB6A66007936D80A7E9CCB">Travis Co.</collectingMunicipality>
:
<location id="F00C339003AD8DA31CCEA8AA2548BDD2" LSID="plazi:21B04C20D6886114A7E0EDB794B86009:E8FFE36948EB037E526DC3334EBC63E4" country="United States" county="Travis Co." municipality="Travis Co." name="Austin" stateProvince="TEXAS">Austin</location>
(
<collectionCode id="9BAB43E9C235A0F0BE77E027F13D4D41" collectionName="USA, New York, New York, American Museum of Natural History">AMNH</collectionCode>
,
<collectionCode id="206FD084ABC2A96B1252F9567FF9A410">MCZC</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="5D22FCF75B0FCD24E95FB11E52874D62" type="discussion">
<paragraph id="4DDDD0F9CD760A49413F1F8E91C9F1E9" pageNumber="487">DISCUSSION</paragraph>
<paragraph id="DE3CDA6E51D776E1E9D5774323C855E2" pageNumber="488">
This is one of several small, nondescript yellow
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species. Because they are rarely encountered, little is known of their habits. Some success has been had locating these minute species using underground baiting, a practice that should be frequently employed. They will likely prove to be subterranean predators of either other ants or of termites.
</paragraph>
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</treatment>
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</taxonomicName>
(L) Rich.
</paragraph>
</subSubSection>
<subSubSection id="0EB62079953ED05A934B26336C19FFE7" pageId="null" pageNumber="613" type="reference_group">
<paragraph id="607229180A1E4CC6AFDA94C30C0631D3" pageId="null" pageNumber="613">
(
<taxonomicName id="1F902195922FF363253286628763ADFB" class="Insecta" family="Tachinidae" genus="Sturmia" kingdom="Animalia" order="Diptera" pageId="null" pageNumber="613" phylum="Arthropoda" rank="species" species="loeselii">
<emphasis id="E382950ACE6A759DAA12013774ACCF44" italics="true" pageId="null" pageNumber="613">Sturmia Loeselii</emphasis>
</taxonomicName>
[
<authorityName id="5E94F900A73F3BE31D2667408498A9EF" pageId="null" pageNumber="613">L.</authorityName>
] Rchb.)
</paragraph>
</subSubSection>
<subSubSection id="F4FBC10DC497ECB32F3D19346C57CF08" pageId="null" pageNumber="613" type="vernacular_names">
<paragraph id="A31778688832BE8E4A2424BDF5E12CC4" pageId="null" pageNumber="613">Loesels Glanzkraut</paragraph>
</subSubSection>
</title>
<subSubSection id="579239CD3824A8F639633A8AC5BBF129" pageId="null" pageNumber="613" type="description">
<paragraph id="114AC84C44611CFFB97F770236BFEC88" pageId="null" pageNumber="613">
Stengel 10-20 cm hoch, am Grunde mit 1-3
<normalizedToken id="642A7DFC379ECA0D59655E9165CF51A4" originalValue="Blättern" pageId="null" pageNumber="613">Blaettern</normalizedToken>
, sonst ohne
<normalizedToken id="83613F96F0F517C30933CCD5287BD98F" originalValue="Blätter" pageId="null" pageNumber="613">Blaetter</normalizedToken>
.
<normalizedToken id="929AA679390D6A4948847C353308419B" originalValue="Blätter" pageId="null" pageNumber="613">Blaetter</normalizedToken>
lanzettlich, 4-10 cm lang, 4-6mal so lang wie breit,
<normalizedToken id="CB2E9C9DA2BEFA6725E239942364D1E5" originalValue="glänzend" pageId="null" pageNumber="613">glaenzend</normalizedToken>
.
<normalizedToken id="F563B86AEC6E0715281ACE2E7FE41B52" originalValue="Blütenstand" pageId="null" pageNumber="613">Bluetenstand</normalizedToken>
3-7
<normalizedToken id="D136C08CC49A6A8BC98627E0F9E96411" originalValue="blütig" pageId="null" pageNumber="613">bluetig</normalizedToken>
.
<normalizedToken id="4D54E22CE94A28EA648A4E556DC749ED" originalValue="Tragblätter" pageId="null" pageNumber="613">Tragblaetter</normalizedToken>
unscheinbar,
<normalizedToken id="D62A4E28176F62FF4FDE41E62A7DFA10" originalValue="schuppenförmig" pageId="null" pageNumber="613">schuppenfoermig</normalizedToken>
, etwa 1 mm lang.
<emphasis id="9A2F2434245833CE7061F722052EFDF2" italics="true" pageId="null" pageNumber="613">
<normalizedToken id="5BE8CD01E8C063D7B587404B9D56059E" originalValue="Blüten" pageId="null" pageNumber="613">Blueten</normalizedToken>
<normalizedToken id="4D3426CC5A368949CCAC9C480037478F" originalValue="gelbgrün" pageId="null" pageNumber="613">gelbgruen</normalizedToken>
, klein;
<normalizedToken id="7AE131FE4CC8208FD6CE90C28AFB06BC" originalValue="Perigonblätter" pageId="null" pageNumber="613">Perigonblaetter</normalizedToken>
4-5 mm lang.
</emphasis>
Lippe oval, rinnig gefaltet,
<normalizedToken id="82A83DB3DC3B64F013846B80847B8FA7" originalValue="abwärts" pageId="null" pageNumber="613">abwaerts</normalizedToken>
gebogen, etwa so lang wie die
<normalizedToken id="A8FAEE57693C35304332053C68B0857D" originalValue="Perigonblätter" pageId="null" pageNumber="613">Perigonblaetter</normalizedToken>
.
<emphasis id="E750DAAB6C551C64F9CA666C493EA89E" italics="true" pageId="null" pageNumber="613">
<normalizedToken id="36DFE5EB1D8EAB713AA0FCCBF7A45818" originalValue="Säule" pageId="null" pageNumber="613">Saeule</normalizedToken>
</emphasis>
(
<normalizedToken id="2EDEEFDC1501ABDD7D1D66A3E6EFEFED" originalValue="Staubblätter" pageId="null" pageNumber="613">Staubblaetter</normalizedToken>
und Griffel)
<emphasis id="08CAEDFD95D458EBA95CB715D408CE39" italics="true" pageId="null" pageNumber="613">
frei stehend, 2-2,5 mm lang, nicht durch
<normalizedToken id="404F03292A55B839B2FB7CE44D023978" originalValue="Perigonblätter" pageId="null" pageNumber="613">Perigonblaetter</normalizedToken>
verdeckt
</emphasis>
(im Gebiet nur bei dieser Art so!),
<normalizedToken id="4FBFB55995365F4237A4C38120EB9A9E" originalValue="schräg" pageId="null" pageNumber="613">schraeg</normalizedToken>
<normalizedToken id="7D45F361E11A6F8CEE21BC679B197DAA" originalValue="aufwärts" pageId="null" pageNumber="613">aufwaerts</normalizedToken>
gerichtet. -
<normalizedToken id="483B36D384F8575DC201BCF52DED93F0" originalValue="Blüte" pageId="null" pageNumber="613">Bluete</normalizedToken>
:
<normalizedToken id="C94DB0562903F1382F14C358DA83834F" originalValue="Früher" pageId="null" pageNumber="613">Frueher</normalizedToken>
Sommer.
</paragraph>
<paragraph id="435617349519F098CF52C34333CDA5E8" pageId="null" pageNumber="613">
<emphasis id="361E66D8B6F0268314E89170550FFE3D" italics="true" pageId="null" pageNumber="613">Zytologische Angaben. 2n = 32:</emphasis>
Material aus
<normalizedToken id="7F5E8F877B7C0D07D988633984C18BAD" originalValue="Dänemark" pageId="null" pageNumber="613">Daenemark</normalizedToken>
(Hagerup 1941).
<emphasis id="79C8E1936C4734B8C3337DB98635DBB6" italics="true" pageId="null" pageNumber="613">2n</emphasis>
=
<emphasis id="94DFC4C89EB3A3B07E15DE4FCDA92C82" italics="true" pageId="null" pageNumber="613">26:</emphasis>
Material aus Holland (Kliphuis 1963, Gadella und Kliphuis 1963).
</paragraph>
<paragraph id="93DF1FD68866F1C472385C6DE38BB595" pageId="null" pageNumber="613">
<emphasis id="7D20ADFCB953F1D0EB1303024515B3B0" italics="true" pageId="null" pageNumber="613">Standort.</emphasis>
Kollin und montan. Kalkhaltige (Tuffbildungen), an der
<normalizedToken id="468C6335511F04629E482602F99AA53A" originalValue="Oberfläche" pageId="null" pageNumber="613">Oberflaeche</normalizedToken>
nie austrocknender
<normalizedToken id="771FE9A642DEC4231E1806C7E51BBCB9" originalValue="Böden" pageId="null" pageNumber="613">Boeden</normalizedToken>
. Flachmoore, besonders
<emphasis id="48A135171F5398BF0B50B05499FEEF40" italics="true" pageId="null" pageNumber="613">Schoenetum nigricantis</emphasis>
W. Koch 1926.
</paragraph>
<paragraph id="4FF023A92B47E130C9C612B20D052CCA" pageId="null" pageNumber="613">
<emphasis id="55982526FC97D2AB11A24AF817DC40EF" italics="true" pageId="null" pageNumber="613">Verbreitung. Eurosibirisch-nordamerikanische Pflanze:</emphasis>
Nordgrenze durch
<taxonomicName id="D399EC86B604DD96F70A73B027D59272" class="Insecta" family="Cerambycidae" genus="Europa" kingdom="Animalia" order="Coleoptera" pageId="null" pageNumber="613" phylum="Arthropoda" rank="genus">Europa</taxonomicName>
entlang 60° NB, vom Baltikum und Polen an
<normalizedToken id="8086419D3EF76647E8EF4FABCFB1787D" originalValue="ostwärts" pageId="null" pageNumber="613">ostwaerts</normalizedToken>
vereinzelt bis ins Obgebiet;
<normalizedToken id="6B6822FF81B6E365B7119F73349224A5" originalValue="Südgrenze" pageId="null" pageNumber="613">Suedgrenze</normalizedToken>
durch
<normalizedToken id="6F0B8A87CFC4003FE676F7F077693105" originalValue="Pyrenäen" pageId="null" pageNumber="613">Pyrenaeen</normalizedToken>
, Alpen,
<normalizedToken id="2EAE51260963356E7AEE7EF74D7AB9F6" originalValue="nördlichen" pageId="null" pageNumber="613">noerdlichen</normalizedToken>
Apennin,
<normalizedToken id="6126E0D30A7A6681E36208C6F3DA8B4F" originalValue="nördliche" pageId="null" pageNumber="613">noerdliche</normalizedToken>
Balkanhalbinsel; isoliert in Jakutsk; im
<normalizedToken id="EAFC9DC52C4977A5E5FAAFB48AF7511F" originalValue="östlichen" pageId="null" pageNumber="613">oestlichen</normalizedToken>
Nordamerika zwischen 30 und 55° NB. Verbreitungskarte von Meusel (1964). - Im Gebiet zerstreut, selten und schwierig zu finden. Zahlreiche neue Angaben aus der Schweiz von Becherer (1966), aus dem
<normalizedToken id="2B0F325660BC7A7A4069258EB087EC55" originalValue="Französischen" pageId="null" pageNumber="613">Franzoesischen</normalizedToken>
Jura von Moreau (1965).
</paragraph>
</subSubSection>
</treatment>
</document>

86
data/A1/44/D2/A144D2D733F0CF5ACD6AD7AD75689E72.xml Normal file
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<mods:title>Chapter 7: Linnaean Plant Names and their Types (part P)</mods:title>
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<bibCitation author="Linnaeus" pagination="704" title="Polygala spinosa" volumeTitle="Species Plantarum 2" year="1753">
<treatmentCitation author="Linnaeus" pageNumber="704" year="1753">
<taxonomicName authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Polygalaceae" genus="Polygala" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="spinosa">
<emphasis bold="true" pageNumber="756">Polygala spinosa</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle>
<emphasis italics="true" pageNumber="756">Species Plantarum</emphasis>
2
</volumeTitle>
: 704. 1753
</treatmentCitation>
</bibCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageNumber="756" type="distribution">
<paragraph pageNumber="756">&quot;Habitat in Aethiopia.&quot; RCN: 5145.</paragraph>
</subSubSection>
<subSubSection pageNumber="756" type="materials_examined">
<paragraph pageNumber="756">
<materialsCitation ID-GBIF-Occurrence="2611900368" collectionCode="G" typeStatus="neotype">
<typeStatus>Neotype</typeStatus>
(Johnson &amp; Weitz in
<emphasis italics="true" pageNumber="756">S. African J. Bot</emphasis>
. 57: 230. 1991): Herb. Burman (
<collectionCode>G</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageNumber="756" type="nomenclature">
<paragraph pageNumber="756">
Current name:
<taxonomicName authority="(L.) F. Forrest &amp; J. C. Manning" authorityName="F. Forrest &amp; J. C. Manning" baseAuthorityName="Linnaeus" baseAuthorityYear="1753" class="Magnoliopsida" family="Polygalaceae" genus="Muraltia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="spinosa">
<emphasis bold="true" italics="true" pageNumber="756">Muraltia spinosa</emphasis>
(L.) F. Forrest &amp; J.C. Manning
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Polygalaceae" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="family">Polygalaceae</taxonomicName>
).
</paragraph>
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