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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.2.e1099" ID-PMC="PMC4107865" ID-Pensoft-Pub="1314-2828--1099" ID-PubMed="25057248" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1314-2828--e1099" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Guide to the Vascular Flora of the Savannas and Flatwoods of Shaken Creek Preserve and Vicinity (Pender &amp; Onslow Counties, North Carolina, U.S.A.)" checkinTime="1553126762101" checkinUser="pensoft" docAuthor="Thornhill, Robert, Krings, Alexander, Lindbo, David &amp; Stucky, Jon" docDate="2014" docId="A1191F24335A4D03553BECA3899681DE" docLanguage="en" docName="BiodivDatJour 2: e1099" docOrigin="Biodiversity Data Journal 2" docSource="http://dx.doi.org/10.3897/BDJ.2.e1099" docTitle="Dichanthelium acuminatum var. acuminatum var. acuminatum (Sw.) Gould &amp; C. A. Clark" docType="treatment" docVersion="5" lastPageNumber="1099" masterDocId="FF8EFFB0FF84FF8FFFDA5C15860F6657" masterDocTitle="Guide to the Vascular Flora of the Savannas and Flatwoods of Shaken Creek Preserve and Vicinity (Pender &amp; Onslow Counties, North Carolina, U. S. A.)" masterLastPageNumber="1099" masterPageNumber="1099" pageNumber="1099" updateTime="1668122592651" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Guide to the Vascular Flora of the Savannas and Flatwoods of Shaken Creek Preserve and Vicinity (Pender &amp; Onslow Counties, North Carolina, U. S. A.)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thornhill, Robert</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krings, Alexander</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lindbo, David</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stucky, Jon</mods:namePart>
</mods:name>
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<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1099</mods:start>
<mods:end>1099</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.2.e1099</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.2.e1099</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--1099</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:A1191F24335A4D03553BECA3899681DE" httpUri="http://treatment.plazi.org/id/A1191F24335A4D03553BECA3899681DE" lastPageNumber="1099" pageId="0" pageNumber="1099">
<subSubSection pageId="0" pageNumber="1099" type="nomenclature">
<paragraph pageId="0" pageNumber="1099">
<taxonomicName authority="(Sw.) Gould &amp; C. A. Clark" class="Liliopsida" family="Poaceae" genus="Dichanthelium" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Dichanthelium acuminatum" order="Poales" pageId="0" pageNumber="1099" phylum="Tracheophyta" rank="variety" species="acuminatum" variety="acuminatum">Dichanthelium acuminatum var. acuminatum (Sw.) Gould &amp; C.A. Clark</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1099" type="distribution">
<paragraph pageId="0" pageNumber="1099">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1099">Pine savannas.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1099" type="notes">
<paragraph pageId="0" pageNumber="1099">Notes</paragraph>
<paragraph pageId="0" pageNumber="1099">
<normalizedToken originalValue="May–Oct">May-Oct</normalizedToken>
. Not seen in Shaken Creek Preserve by the senior author. Specimens seen in the vicinity: Sandy Run [Hancock]: Taggart SARU 653 (WNC!). [&lt;
<taxonomicName class="Liliopsida" family="Poaceae" genus="Panicum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Panicum lanuginosum" order="Poales" pageId="0" pageNumber="1099" phylum="Tracheophyta" rank="species" species="lanuginosum">Panicum lanuginosum</taxonomicName>
Elliott sensu RAB; =
<taxonomicName basionym-authority="(Sw.) Gould &amp; C. A. Clark" class="Liliopsida" family="Poaceae" genus="Dichanthelium" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Dichanthelium acuminatum var. acuminatum" order="Poales" pageId="0" pageNumber="1099" phylum="Tracheophyta" rank="variety" species="acuminatum" variety="acuminatum">Dichanthelium acuminatum (Sw.) Gould &amp; C. A. Clark var. acuminatum</taxonomicName>
sensu FNA; = Weakley]
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="77D33729F1EE99DA5C8ABCFF7EADD38E" ID-CLB-Dataset="29764" ID-DOI="10.11646/zootaxa.4442.2.5" ID-GBIF-Dataset="0fea0081-d093-4611-89c2-316f9baf6ca4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303125" ID-ZooBank="8EB2105E-3EAA-444A-A20E-819AD215B3C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530533589605" checkinUser="plazi" docAuthor="Khaustov, Alexander A. &amp; Frolov, Andrey V." docDate="2018" docId="A1192263FE51BD3AC7E1038F426BFF1A" docLanguage="en" docName="zootaxa.4442.2.5.pdf" docOrigin="Zootaxa 4442 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nasutidania Khaustov &amp; Frolov 2018, gen. nov." docType="treatment" docVersion="7" lastPageNumber="285" masterDocId="5D205A1BFE57BD32C776016A461FFFCF" masterDocTitle="New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand" masterLastPageNumber="292" masterPageNumber="277" pageNumber="283" updateTime="1698552142957" updateUser="ExternalLinkService">
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<mods:title id="3117DED9D1A455F31694E970AAFF7BF6">New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand</mods:title>
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<mods:date id="CE0AC4739D4F49773F6A90E0C8DB11D8">2018</mods:date>
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<treatment id="A1192263FE51BD3AC7E1038F426BFF1A" ID-DOI="http://doi.org/10.5281/zenodo.5949611" ID-GBIF-Taxon="146044082" ID-Zenodo-Dep="5949611" LSID="urn:lsid:plazi:treatment:A1192263FE51BD3AC7E1038F426BFF1A" httpUri="http://treatment.plazi.org/id/A1192263FE51BD3AC7E1038F426BFF1A" lastPageId="8" lastPageNumber="285" pageId="6" pageNumber="283">
<subSubSection id="61AAC0FEFE51BD34C7E1038F47E8FD30" box="[151,503,741,767]" pageId="6" pageNumber="283" type="nomenclature">
<paragraph id="290F9375FE51BD34C7E1038F47E8FD30" blockId="6.[151,503,741,767]" box="[151,503,741,767]" pageId="6" pageNumber="283">
<heading id="72472419FE51BD34C7E1038F47E8FD30" bold="true" box="[151,503,741,767]" fontSize="11" level="1" pageId="6" pageNumber="283" reason="1">
<emphasis id="1BC44F67FE51BD34C7E1038F47E8FD30" bold="true" box="[151,503,741,767]" pageId="6" pageNumber="283">
Genus
<taxonomicName id="EEB0E8F6FE51BD34C798038F479CFD30" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[238,387,741,767]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BC44F67FE51BD34C798038F479CFD30" bold="true" box="[238,387,741,767]" italics="true" pageId="6" pageNumber="283">Nasutidania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE51BD34C6FF038F47E8FD30" box="[393,503,741,767]" pageId="6" pageNumber="283" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE51BD34C7E10246447FFC8F" box="[151,608,811,833]" pageId="6" pageNumber="283" type="materials_examined">
<paragraph id="290F9375FE51BD34C7E10246447FFC8F" blockId="6.[151,608,811,833]" box="[151,608,811,833]" pageId="6" pageNumber="283">
Type species:
<taxonomicName id="EEB0E8F6FE51BD34C65002414416FC8E" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[294,521,811,833]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="1BC44F67FE51BD34C65002414416FC8E" box="[294,521,811,833]" italics="true" pageId="6" pageNumber="283">Nasutidania orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE51BD34C5790241447FFC8F" box="[527,608,811,832]" pageId="6" pageNumber="283" rank="species">
<emphasis id="1BC44F67FE51BD34C5790241447FFC8F" bold="true" box="[527,608,811,832]" pageId="6" pageNumber="283">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE51BD35C7E102074263F86B" lastPageId="7" lastPageNumber="284" pageId="6" pageNumber="283" type="diagnosis">
<paragraph id="290F9375FE51BD34C7E102074737F8D7" blockId="6.[151,1437,877,2032]" pageId="6" pageNumber="283">
<emphasis id="1BC44F67FE51BD34C7E102074715FC49" bold="true" box="[151,266,877,902]" pageId="6" pageNumber="283">Diagnosis</emphasis>
. FEMALE. Body oval. Gnathosomal capsule elongate, beak-like, its length almost three times longer than width, dorsally with two pairs of cheliceral setae (
<emphasis id="1BC44F67FE51BD34C46402FB4525FC65" box="[786,826,913,938]" italics="true" pageId="6" pageNumber="283">сha</emphasis>
,
<emphasis id="1BC44F67FE51BD34C43C02FB456DFC65" box="[842,882,913,938]" italics="true" pageId="6" pageNumber="283">сhb</emphasis>
); postpalpal setae (
<emphasis id="1BC44F67FE51BD34C32202F9426FFC65" box="[1108,1136,915,938]" italics="true" pageId="6" pageNumber="283">pp</emphasis>
) present; palps very long, with two pairs of dorsal setae (
<emphasis id="1BC44F67FE51BD34C68D02DF4438FC01" box="[507,551,949,974]" italics="true" pageId="6" pageNumber="283">dFe</emphasis>
,
<emphasis id="1BC44F67FE51BD34C54002DF447AFC01" box="[566,613,949,974]" italics="true" pageId="6" pageNumber="283">dGe</emphasis>
). Palpal tibiotarsus without tibial claw and seta. Subcapitulum with one pair of setae
<emphasis id="1BC44F67FE51BD34C64C02B14751FC3D" box="[314,334,987,1010]" italics="true" pageId="6" pageNumber="283">m</emphasis>
and round pits in posterior part; palps ventrally without accessory setigenous structure and solenidion; ventrolaterally with unusual oval depressions (dep). Rostrum of subcapitulum with several lateral short spiniform projections in distal half. Pharyngeal pumps tripartite, striated, situated closely to each other; pharyngeal pump 1 small, butterfly-like, pharyngeal pump 2 large, subrectangular and about three times longer than oval pump 3. Posterior margin of prodorsum slightly covered by anterior part of tergite C. Prodorsum with two pairs of simple setae (
<date id="5D0EB5B5FE51BD34C7A905E546EBFB64" box="[223,244,1167,1195]" pageId="6" pageNumber="283">
<emphasis id="1BC44F67FE51BD34C7A905E546F4FB69" box="[223,235,1167,1190]" italics="true" pageId="6" pageNumber="283">v</emphasis>
2
</date>
,
<emphasis id="1BC44F67FE51BD34C67705E54707FB69" box="[257,280,1167,1190]" italics="true" pageId="6" pageNumber="283">sс</emphasis>
2), pair of clavate trichobothria (
<emphasis id="1BC44F67FE51BD34C5F605E54488FB69" box="[640,663,1167,1190]" italics="true" pageId="6" pageNumber="283">sс</emphasis>
1) and small, round stigmata located just anteriorly to setae
<date id="5D0EB5B5FE51BD34C25905E54355FB69" box="[1327,1354,1167,1195]" pageId="6" pageNumber="283">
<emphasis id="1BC44F67FE51BD34C25905E54324FB69" box="[1327,1339,1167,1190]" italics="true" pageId="6" pageNumber="283">v</emphasis>
2.
</date>
Dorsal idiosomal setae not modified. Tergite C with two pairs of setae (
<emphasis id="1BC44F67FE51BD34C4F405DC4591FB02" box="[898,910,1206,1229]" italics="true" pageId="6" pageNumber="283">с</emphasis>
1,
<emphasis id="1BC44F67FE51BD34C4D105DC45ACFB02" box="[935,947,1206,1229]" italics="true" pageId="6" pageNumber="283">с</emphasis>
2); tergite D with one pair of setae (
<emphasis id="1BC44F67FE51BD34C22505DE437EFB02" box="[1363,1377,1204,1229]" italics="true" pageId="6" pageNumber="283">d</emphasis>
) and round cupules
<emphasis id="1BC44F67FE51BD34C63505B64746FB3C" box="[323,345,1244,1267]" italics="true" pageId="6" pageNumber="283">ia</emphasis>
; tergite EF with two pairs of setae (
<emphasis id="1BC44F67FE51BD34C47005B6450DFB3C" box="[774,786,1244,1267]" italics="true" pageId="6" pageNumber="283">e</emphasis>
,
<emphasis id="1BC44F67FE51BD34C45205B64533FB3B" box="[804,812,1244,1268]" italics="true" pageId="6" pageNumber="283">f</emphasis>
); tergite H with two pairs of setae (
<emphasis id="1BC44F67FE51BD34C3A105B142FAFB3B" box="[1239,1253,1243,1268]" italics="true" pageId="6" pageNumber="283">h</emphasis>
1,
<emphasis id="1BC44F67FE51BD34C38805B14313FB3B" box="[1278,1292,1243,1268]" italics="true" pageId="6" pageNumber="283">h</emphasis>
2) and round cupules
<emphasis id="1BC44F67FE51BD34C78304684714FAD4" box="[245,267,1282,1307]" italics="true" pageId="6" pageNumber="283">ih</emphasis>
. Coxal fields I with two pairs of setae (1
<emphasis id="1BC44F67FE51BD34C591046944EAFAD5" box="[743,757,1283,1306]" italics="true" pageId="6" pageNumber="283">a</emphasis>
, 1
<emphasis id="1BC44F67FE51BD34C4620468453DFAD4" box="[788,802,1282,1307]" italics="true" pageId="6" pageNumber="283">b</emphasis>
); coxal fields II with two pairs of setae (2
<emphasis id="1BC44F67FE51BD34C27D04694306FAD5" box="[1291,1305,1283,1306]" italics="true" pageId="6" pageNumber="283">a</emphasis>
, 2
<emphasis id="1BC44F67FE51BD34C2410468435AFAD4" box="[1335,1349,1282,1307]" italics="true" pageId="6" pageNumber="283">b</emphasis>
); coxal fields III with three pairs of setae (3
<emphasis id="1BC44F67FE51BD34C54A044D4455FAF1" box="[572,586,1319,1342]" italics="true" pageId="6" pageNumber="283">a</emphasis>
, 3
<emphasis id="1BC44F67FE51BD34C51F044C4468FAF0" box="[617,631,1318,1343]" italics="true" pageId="6" pageNumber="283">b</emphasis>
, 3
<emphasis id="1BC44F67FE51BD34C5E0044D44BDFAF1" box="[662,674,1319,1342]" italics="true" pageId="6" pageNumber="283">с</emphasis>
); coxal fields IV with three pairs of setae (4
<emphasis id="1BC44F67FE51BD34C3DC044D42A7FAF1" box="[1194,1208,1319,1342]" italics="true" pageId="6" pageNumber="283">a</emphasis>
, 4
<emphasis id="1BC44F67FE51BD34C3A1044C42FAFAF0" box="[1239,1253,1318,1343]" italics="true" pageId="6" pageNumber="283">b</emphasis>
, 4
<emphasis id="1BC44F67FE51BD34C272044D430FFAF1" box="[1284,1296,1319,1342]" italics="true" pageId="6" pageNumber="283">с</emphasis>
). Pseudanal segment with three pairs of setae (
<emphasis id="1BC44F67FE51BD34C56E0421442FFAAD" box="[536,560,1355,1378]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
1-3). Ap1 and ap2 well developed and joined together and with appr, apsej well developed and joined with appr; ap3 very weak, linear, ap4 long, reaching distinctly beyond bases of setae 3
<emphasis id="1BC44F67FE51BD34C2F1041B438AFA45" box="[1415,1429,1393,1418]" italics="true" pageId="6" pageNumber="283">b</emphasis>
. Apodemes 5 vestigial, located near the bases of trochanters IV. Secondary transverse apodeme absent. Posterior margin of posterior sternal plate tripartite. Ags bell-like, pgs triangular, median genital sclerite (mgs) present. Ventral idiosomal setae not modified. Legs I and II subequal in length. Tibiotarsus I slightly thickened, with distinct claw directed laterally, situated on pretarsus; ventrodistal part of pretarsus with short and thick projection (fused unguinal setae); seta
<emphasis id="1BC44F67FE51BD34C6A8074F47F3F9F1" box="[478,492,1573,1598]" italics="true" pageId="6" pageNumber="283">d</emphasis>
of femur I hook-like, seta
<emphasis id="1BC44F67FE51BD34C458074F4525F9F1" box="[814,826,1573,1598]" italics="true" pageId="6" pageNumber="283">k</emphasis>
long, pointed, seta-like. Claws on tarsus IV simple, distinctly thinner than padded claws on tarsi II and III. Tarsus IV not strongly elongate. Femora III–IV divided into basi- and telofemur. Leg setation: leg I; Tr 1 (
<emphasis id="1BC44F67FE51BD34C5E3070444B4F94A" box="[661,683,1646,1669]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Fe 3 (
<emphasis id="1BC44F67FE51BD34C4720707450DF949" box="[772,786,1645,1670]" italics="true" pageId="6" pageNumber="283">d</emphasis>
,
<emphasis id="1BC44F67FE51BD34C4690707452FF949" box="[799,816,1645,1670]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C44807044545F94A" box="[830,858,1646,1669]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
), Ge 4 (
<emphasis id="1BC44F67FE51BD34C4CE070745D6F949" box="[952,969,1645,1670]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C4A1070745F1F949" box="[983,1006,1645,1670]" italics="true" pageId="6" pageNumber="283">l”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C48A0704420EF94A" box="[1020,1041,1646,1669]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C35607044224F94A" box="[1056,1083,1646,1669]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
), TiTa 16(4) (
<emphasis id="1BC44F67FE51BD34C3AC070742F7F949" box="[1242,1256,1645,1670]" italics="true" pageId="6" pageNumber="283">d</emphasis>
,
<emphasis id="1BC44F67FE51BD34C38007074318F949" box="[1270,1287,1645,1670]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C26307074332F949" box="[1301,1325,1645,1670]" italics="true" pageId="6" pageNumber="283">l”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C24D0704434FF94A" box="[1339,1360,1646,1669]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C22807044365F94A" box="[1374,1402,1646,1669]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C2FE0707438BF949" box="[1416,1428,1645,1670]" italics="true" pageId="6" pageNumber="283">k</emphasis>
,,
<emphasis id="1BC44F67FE51BD34C7D307F846DDF966" box="[165,194,1682,1705]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C7A607F846EBF966" box="[208,244,1682,1705]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C67407F84700F966" box="[258,287,1682,1705]" italics="true" pageId="6" pageNumber="283">p”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C65B07FB4759F966" box="[301,326,1681,1705]" italics="true" pageId="6" pageNumber="283">ft’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C62207FB476CF966" box="[340,371,1681,1705]" italics="true" pageId="6" pageNumber="283">ft”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C6F407F847B9F966" box="[386,422,1682,1705]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C6C507F847C2F966" box="[435,477,1682,1705]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C69D07FB4415F965" box="[491,522,1681,1706]" italics="true" pageId="6" pageNumber="283">pl’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C56E07FB4422F965" box="[536,573,1681,1706]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C53D07F84449F966" box="[587,598,1682,1705]" italics="true" pageId="6" pageNumber="283">s</emphasis>
,
<emphasis id="1BC44F67FE51BD34C51207F84467F966" box="[612,632,1682,1705]" italics="true" pageId="6" pageNumber="283">ω</emphasis>
1,
<emphasis id="1BC44F67FE51BD34C5F807F844BDF966" box="[654,674,1682,1705]" italics="true" pageId="6" pageNumber="283">ω</emphasis>
2,
<emphasis id="1BC44F67FE51BD34C5C107F844D9F966" box="[695,710,1682,1705]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
1,
<emphasis id="1BC44F67FE51BD34C5AA07F844F4F966" box="[732,747,1682,1705]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
2); leg II: Tr 1 (
<emphasis id="1BC44F67FE51BD34C4E307F845B4F966" box="[917,939,1682,1705]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Fe 3 (
<emphasis id="1BC44F67FE51BD34C37507FB422FF965" box="[1027,1072,1681,1706]" italics="true" pageId="6" pageNumber="283">d, l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C34807F84245F966" box="[1086,1114,1682,1705]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
), Ge 3 (
<emphasis id="1BC44F67FE51BD34C3CE07FB42D6F965" box="[1208,1225,1681,1706]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C3A107FB42F1F965" box="[1239,1262,1681,1706]" italics="true" pageId="6" pageNumber="283">l”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C38A07F8430EF966" box="[1276,1297,1682,1705]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Ti 4(1) (
<emphasis id="1BC44F67FE51BD34C2F107FB438AF965" box="[1415,1429,1681,1706]" italics="true" pageId="6" pageNumber="283">d</emphasis>
,
<emphasis id="1BC44F67FE51BD34C7E107DD46B7F91F" box="[151,168,1719,1744]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C7C007D246D3F900" box="[182,204,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C7AD07D246E9F900" box="[219,246,1720,1743]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C67307D3470BF91F" box="[261,276,1721,1744]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
), ta 6(1) (
<emphasis id="1BC44F67FE51BD34C6FE07D247BAF900" box="[392,421,1720,1743]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C6C207D247C8F900" box="[436,471,1720,1743]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C69007DD4415F91F" box="[486,522,1719,1744]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C56C07D24422F900" box="[538,573,1720,1743]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C53A07D2446AF900" box="[588,629,1720,1743]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C5F207D24484F900" box="[644,667,1720,1743]" italics="true" pageId="6" pageNumber="283">u’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C5DC07D344A1F91F" box="[682,702,1721,1744]" italics="true" pageId="6" pageNumber="283">ω</emphasis>
); leg III: Tr 1 (
<emphasis id="1BC44F67FE51BD34C41A07D2459DF900" box="[876,898,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Fe 2 (
<emphasis id="1BC44F67FE51BD34C4AD07DD45F6F91F" box="[987,1001,1719,1744]" italics="true" pageId="6" pageNumber="283">d</emphasis>
,
<emphasis id="1BC44F67FE51BD34C48E07D24211F900" box="[1016,1038,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Ge 2 (
<emphasis id="1BC44F67FE51BD34C31A07DD4262F91F" box="[1132,1149,1719,1744]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C3FA07D242BEF900" box="[1164,1185,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Ti 4(1) (
<emphasis id="1BC44F67FE51BD34C26E07DD4339F91F" box="[1304,1318,1719,1744]" italics="true" pageId="6" pageNumber="283">d</emphasis>
,
<emphasis id="1BC44F67FE51BD34C24307DD4359F91F" box="[1333,1350,1719,1744]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C22207D24375F900" box="[1364,1386,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C20F07D2438BF900" box="[1401,1428,1720,1743]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C7E107B746B9F93B" box="[151,166,1757,1780]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
), Ta 6 (
<emphasis id="1BC44F67FE51BD34C78A07B74706F93B" box="[252,281,1757,1780]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C65107B74754F93B" box="[295,331,1757,1780]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C62F07B64762F93A" box="[345,381,1756,1781]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C6FD07B747B0F93B" box="[395,431,1757,1780]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C6CA07B747FAF93B" box="[444,485,1757,1780]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C68207B74414F93B" box="[500,523,1757,1780]" italics="true" pageId="6" pageNumber="283">u’</emphasis>
); leg IV: Tr 1 (
<emphasis id="1BC44F67FE51BD34C5C407B744D8F93B" box="[690,711,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), F 2 (
<emphasis id="1BC44F67FE51BD34C46407B6453FF93A" box="[786,800,1756,1781]" italics="true" pageId="6" pageNumber="283">d</emphasis>
,
<emphasis id="1BC44F67FE51BD34C45B07B7455CF93B" box="[813,835,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Ge 1 (
<emphasis id="1BC44F67FE51BD34C4E807B745ABF93B" box="[926,948,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
), Ti 4(1) (
<emphasis id="1BC44F67FE51BD34C35107B6422AF93A" box="[1063,1077,1756,1781]" italics="true" pageId="6" pageNumber="283">d</emphasis>
,
<emphasis id="1BC44F67FE51BD34C33507B6424BF93A" box="[1091,1108,1756,1781]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C31707B74268F93B" box="[1121,1143,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C3F207B742BFF93B" box="[1156,1184,1757,1780]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C3D807B742A2F93B" box="[1198,1213,1757,1780]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
), Ta 6 (
<emphasis id="1BC44F67FE51BD34C26507B7432FF93B" box="[1299,1328,1757,1780]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C24807B7437DF93B" box="[1342,1378,1757,1780]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C21907B6438BF93A" box="[1391,1428,1756,1781]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
,
<emphasis id="1BC44F67FE51BD34C7E1066A46B1F8D8" box="[151,174,1792,1815]" italics="true" pageId="6" pageNumber="283">u’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C7CA066A46FFF8D8" box="[188,224,1792,1815]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
,
<emphasis id="1BC44F67FE51BD34C798066A4707F8D8" box="[238,280,1792,1815]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
).
</paragraph>
<paragraph id="290F9375FE51BD34C7B1064E45D7F83F" blockId="6.[151,1437,877,2032]" pageId="6" pageNumber="283">
<emphasis id="1BC44F67FE51BD34C7B1064E47D8F8F2" bold="true" box="[199,455,1828,1853]" pageId="6" pageNumber="283">Differential diagnosis</emphasis>
. The new genus is most similar to
<taxonomicName id="EEB0E8F6FE51BD34C42D064F426FF8F2" authority="Mahunka" authorityName="Mahunka" authorityYear="1968" box="[859,1136,1828,1853]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE51BD34C42D064F45E4F8F3" box="[859,1019,1829,1852]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
Mahunka
</taxonomicName>
, 19681 by the presence of extremely long palps. The new genus differs from
<taxonomicName id="EEB0E8F6FE51BD34C59E06234598F8AF" authorityName="Mahunka" authorityYear="1968" box="[744,903,1865,1888]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE51BD34C59E06234598F8AF" box="[744,903,1865,1888]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
</taxonomicName>
by the tripartite posterior margin of posterior sternal plate (vs. entire in
<taxonomicName id="EEB0E8F6FE51BD34C6B906074471F84B" authorityName="Mahunka" authorityYear="1968" box="[463,622,1901,1924]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE51BD34C6B906074471F84B" box="[463,622,1901,1924]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
</taxonomicName>
), presence of a well developed median genital sclerite (vs. absent in
<taxonomicName id="EEB0E8F6FE51BD34C7E106FB4728F867" authorityName="Mahunka" authorityYear="1968" box="[151,311,1937,1960]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE51BD34C7E106FB4728F867" box="[151,311,1937,1960]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
</taxonomicName>
), absence of the secondary transverse apodeme (vs. present in
<taxonomicName id="EEB0E8F6FE51BD34C36606FB42AFF867" authorityName="Mahunka" authorityYear="1968" box="[1040,1200,1937,1960]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE51BD34C36606FB42AFF867" box="[1040,1200,1937,1960]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
</taxonomicName>
), presence of lateral cuticular projections on the rostrum of the subcapitulum (vs. absent in
<taxonomicName id="EEB0E8F6FE51BD34C4DD06DF4254F803" authorityName="Mahunka" authorityYear="1968" box="[939,1099,1973,1996]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE51BD34C4DD06DF4254F803" box="[939,1099,1973,1996]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
</taxonomicName>
), and absence of an accessory setigenous structure and palpal solenidion (vs. present in
<taxonomicName id="EEB0E8F6FE51BD34C46E06B345A7F83F" authorityName="Mahunka" authorityYear="1968" box="[792,952,2009,2032]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE51BD34C46E06B345A7F83F" box="[792,952,2009,2032]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="7DCFC3FDFE50BD35C7E106664297F890" httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="7" pageNumber="284" startId="7.[151,250,1804,1826]" targetBox="[151,1436,180,1782]" targetPageId="7">
<paragraph id="290F9375FE50BD35C7E106664297F890" blockId="7.[151,1436,1804,1887]" pageId="7" pageNumber="284">
<emphasis id="1BC44F67FE50BD35C7E106664713F8EE" bold="true" box="[151,268,1804,1826]" pageId="7" pageNumber="284">FIGURE 5</emphasis>
. DIC micrographs of
<taxonomicName id="EEB0E8F6FE50BD35C69D06664535F8ED" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[491,810,1804,1826]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
<emphasis id="1BC44F67FE50BD35C69D06664535F8ED" box="[491,810,1804,1826]" italics="true" pageId="7" pageNumber="284">Spatulaphorus enoplotruporum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE50BD35C4460666459FF8EE" box="[816,896,1804,1825]" pageId="7" pageNumber="284" rank="species">
<emphasis id="1BC44F67FE50BD35C4460666459FF8EE" bold="true" box="[816,896,1804,1825]" pageId="7" pageNumber="284">sp. nov.</emphasis>
</taxonomicNameLabel>
, female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites.
</paragraph>
</caption>
<paragraph id="290F9375FE50BD35C7B106E64263F86B" blockId="7.[151,1436,1932,2029]" box="[199,1148,1932,1957]" pageId="7" pageNumber="284">
<emphasis id="1BC44F67FE50BD35C7B106E64793F86A" bold="true" box="[199,396,1932,1957]" pageId="7" pageNumber="284">Species included</emphasis>
. The genus
<taxonomicName id="EEB0E8F6FE50BD35C56206E64482F86A" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[532,669,1932,1957]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="290" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BC44F67FE50BD35C56206E64482F86A" box="[532,669,1932,1957]" italics="true" pageId="7" pageNumber="284">Nasutidania</emphasis>
</taxonomicName>
includes one species
<taxonomicName id="EEB0E8F6FE50BD35C4E606E74203F86A" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[912,1052,1932,1957]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="1BC44F67FE50BD35C4E606E745BCF86B" box="[912,931,1933,1956]" italics="true" pageId="7" pageNumber="284">N</emphasis>
.
<emphasis id="1BC44F67FE50BD35C4C706E64203F86A" box="[945,1052,1932,1957]" italics="true" pageId="7" pageNumber="284">orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE50BD35C35506E64263F86B" box="[1059,1148,1932,1957]" pageId="7" pageNumber="284" rank="species">
<emphasis id="1BC44F67FE50BD35C35506E64269F86A" bold="true" box="[1059,1142,1932,1957]" pageId="7" pageNumber="284">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE50BD35C7B106DA47A2F822" pageId="7" pageNumber="284" type="distribution">
<paragraph id="290F9375FE50BD35C7B106DA47A2F822" blockId="7.[151,1436,1932,2029]" pageId="7" pageNumber="284">
<emphasis id="1BC44F67FE50BD35C7B106DA4418F806" bold="true" box="[199,519,1968,1993]" pageId="7" pageNumber="284">Distribution and habitats</emphasis>
.
<taxonomicName id="EEB0E8F6FE50BD35C56806DA453CF806" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[542,803,1968,1993]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="1BC44F67FE50BD35C56806DA453CF806" box="[542,803,1968,1993]" italics="true" pageId="7" pageNumber="284">Nasutidania orientalis</emphasis>
</taxonomicName>
known only from the scarab beetles of the genus
<taxonomicName id="EEB0E8F6FE50BD35C7E106BE4731F822" authorityName="Lucas" authorityYear="1869" box="[151,302,2004,2029]" class="Insecta" family="Geotrupidae" genus="Enoplotrupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE50BD35C7E106BE4731F822" box="[151,302,2004,2029]" italics="true" pageId="7" pageNumber="284">Enoplotrupes</emphasis>
</taxonomicName>
in
<collectingCountry id="51A7D3E5FE50BD35C62706BE47A6F822" box="[337,441,2004,2029]" name="Thailand" pageId="7" pageNumber="284">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE5FBD3AC7B101FD426BFF1A" pageId="8" pageNumber="285" type="etymology">
<paragraph id="290F9375FE5FBD3AC7B101FD426BFF1A" blockId="8.[151,1436,151,213]" pageId="8" pageNumber="285">
<emphasis id="1BC44F67FE5FBD3AC7B101FD4759FF7F" bold="true" box="[199,326,151,176]" pageId="8" pageNumber="285">Etymology</emphasis>
. The generic name is a combination of two words: Latin
<emphasis id="1BC44F67FE5FBD3AC49201F34226FF7F" box="[996,1081,153,176]" italics="true" pageId="8" pageNumber="285">nasutus</emphasis>
meaning
<emphasis id="1BC44F67FE5FBD3AC3DA01FD42F2FF7F" box="[1196,1261,151,176]" italics="true" pageId="8" pageNumber="285">nosed</emphasis>
and
<emphasis id="1BC44F67FE5FBD3AC25F01FD436FFF7F" box="[1321,1392,151,176]" italics="true" pageId="8" pageNumber="285">dania,</emphasis>
the second part of the generic name
<taxonomicName id="EEB0E8F6FE5FBD3AC57501D6449AFF1A" authorityName="Sasa" authorityYear="1961" box="[515,645,188,213]" class="Arachnida" family="Pygmephoridae" genus="Bakerdania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE5FBD3AC57501D6449AFF1A" box="[515,645,188,213]" italics="true" pageId="8" pageNumber="285">Bakerdania</emphasis>
</taxonomicName>
, the most abundant neopygmephorid genus.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="1BC44F67FE52BD37C7E107064234F949" bold="true" box="[151,1067,1643,1670]" pageId="5" pageNumber="282">
Key to world species of
<taxonomicName id="EEB0E8F6FE52BD37C6C807014470F94A" authorityName="Rack" authorityYear="1993" box="[446,623,1643,1669]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="282" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE52BD37C6C807014470F94A" bold="true" box="[446,623,1643,1669]" italics="true" pageId="5" pageNumber="282">Spatulaphorus</emphasis>
</taxonomicName>
(based on
<bibRefCitation id="4D21EE84FE52BD37C5800706423CF949" box="[758,1059,1644,1670]" pageId="5" pageNumber="282" refId="ref8366" refString="Khaustov, A. A. &amp; Trach, V. A. (2018) New species of Spatulaphorus (Acari: Heterostigmata: Pygmephoridae) associated with Geotrupes baicalicus (Coleoptera: Geotrupidae) from Altai Republic, Russia. Systematic &amp; Applied Acarology, 23, 123 - 131. https: // doi. org / 10.11158 / saa. 23.1.10" type="journal article">Khaustov &amp; Trach 2018</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="61AAC0FEFE52BD34C7E107D84383FD97" lastPageId="6" lastPageNumber="283" pageId="5" pageNumber="282" type="key">
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<keyLead id="92413040FE52BD34C7E107D84383FD97" lastPageId="6" lastPageNumber="283" pageId="5" pageNumber="282">
1. Genu I with two setae.................................................................................. 2 - Genu I with one seta.................................................
<taxonomicName id="EEB0E8F6FE52BD37C48907A54313F92B" authority="Khaustov, 2007 a" authorityName="Khaustov" authorityYear="2007" box="[1023,1292,1743,1764]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="copridis">
<emphasis id="1BC44F67FE52BD37C48907A54279F92B" box="[1023,1126,1743,1764]" italics="true" pageId="5" pageNumber="282">S. copridis</emphasis>
<bibRefCitation id="4D21EE84FE52BD37C31A07A54313F92B" box="[1132,1292,1743,1764]" pageId="5" pageNumber="282" refId="ref7693" refString="Khaustov, A. A. (2007 a) A new species of the genus Spatulaphorus (Acari: Heterostigmata: Pygmephoridae) associated with Copris lunaris (Coleoptera: Scarabaeidae) from Crimea. Acarina, 15, 155 - 157." type="journal article">Khaustov, 2007a</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="EB745D97FE52BD37C26F07A5437CF92B" box="[1305,1379,1743,1764]" country="Ukraine" name="Crimea" pageId="5" pageNumber="282">Crimea</collectingRegion>
,
<collectingCountry id="51A7D3E5FE52BD37C21B07A5438AF92B" box="[1389,1429,1743,1764]" name="Iran" pageId="5" pageNumber="282">Iran</collectingCountry>
) 2. Tarsi II and III each with spiniform setae
<emphasis id="1BC44F67FE52BD37C51807864490F8CE" box="[622,655,1772,1793]" italics="true" pageId="5" pageNumber="282">pl”</emphasis>
and
<emphasis id="1BC44F67FE52BD37C5CB078644C9F8CE" box="[701,726,1772,1793]" italics="true" pageId="5" pageNumber="282">tc’</emphasis>
......................................................... 3 - Tarsi II and III without spiniform setae, or only
<emphasis id="1BC44F67FE52BD37C5DA066044D4F8D0" box="[684,715,1802,1823]" italics="true" pageId="5" pageNumber="282">pl”</emphasis>
or
<emphasis id="1BC44F67FE52BD37C59A06604519F8D0" box="[748,774,1802,1823]" italics="true" pageId="5" pageNumber="282">tc’</emphasis>
spiniform.............................................. 5 3. Setae
<emphasis id="1BC44F67FE52BD37C66D064D4730F8F3" box="[283,303,1831,1852]" italics="true" pageId="5" pageNumber="282">sc</emphasis>
2 foliate..............................................
<taxonomicName id="EEB0E8F6FE52BD37C4D1064D42ACF8F3" authority="Khaustov, 2005" authorityName="Khaustov" authorityYear="2005" box="[935,1203,1831,1852]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="vladimiri">
<emphasis id="1BC44F67FE52BD37C4D1064D4208F8F3" box="[935,1047,1831,1852]" italics="true" pageId="5" pageNumber="282">S. vladimiri</emphasis>
<bibRefCitation id="4D21EE84FE52BD37C36B064D42ACF8F3" box="[1053,1203,1831,1852]" pageId="5" pageNumber="282" refId="ref7652" refString="Khaustov, A. A. (2005) Three new species of mite family Pygmephoridae (Acari: Heterostigmata) associated with scarab beetles (Coleoptera: Scarabaeidae) from Ukraine. Acarina, 13, 35 - 42." type="journal article">Khaustov, 2005</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="51A7D3E5FE52BD37C3B6064D430EF8F3" box="[1216,1297,1831,1852]" name="Ukraine" pageId="5" pageNumber="282">Ukraine</collectingCountry>
,
<collectingRegion id="EB745D97FE52BD37C26C064D437CF8F3" box="[1306,1379,1831,1852]" country="Ukraine" name="Crimea" pageId="5" pageNumber="282">Crimea</collectingRegion>
,
<collectingCountry id="51A7D3E5FE52BD37C21B064D438AF8F3" box="[1389,1429,1831,1852]" name="Iran" pageId="5" pageNumber="282">Iran</collectingCountry>
) - Setae
<emphasis id="1BC44F67FE52BD37C66D062D4730F893" box="[283,303,1863,1884]" italics="true" pageId="5" pageNumber="282">sc</emphasis>
2 not modified, setiform.......................................................................... 4 4. Setae
<emphasis id="1BC44F67FE52BD37C66D060D4739F8B3" box="[283,294,1895,1916]" italics="true" pageId="5" pageNumber="282">c</emphasis>
2 foliate, blunt-ended......................................
<taxonomicName id="EEB0E8F6FE52BD37C4B6060D4384F8B3" authority="Khaustov and Trach, 2012 a" authorityName="Khaustov and Trach" authorityYear="2012" box="[960,1435,1895,1916]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="1BC44F67FE52BD37C4B6060D45D3F8B3" box="[960,972,1895,1916]" italics="true" pageId="5" pageNumber="282">S</emphasis>
.
<emphasis id="1BC44F67FE52BD37C4AE060D4234F8B3" box="[984,1067,1895,1916]" italics="true" pageId="5" pageNumber="282">venustus</emphasis>
<bibRefCitation id="4D21EE84FE52BD37C346060D4327F8B3" box="[1072,1336,1895,1916]" pageId="5" pageNumber="282" refId="ref8182" refString="Khaustov, A. A. &amp; Trach, V. A. (2012 a) A new species and new records of mites of the genus Spatulaphorus Rack (Acari: Heterostigmata: Pygmephoridae) from Ukraine. International Journal of Acarology, 38, 480 - 485." type="journal article">Khaustov and Trach, 2012a</bibRefCitation>
(
<collectingCountry id="51A7D3E5FE52BD37C233060D438AF8B3" box="[1349,1429,1895,1916]" name="Ukraine" pageId="5" pageNumber="282">Ukraine</collectingCountry>
)
</taxonomicName>
- Setae
<emphasis id="1BC44F67FE52BD37C66D06EC4739F854" box="[283,294,1926,1947]" italics="true" pageId="5" pageNumber="282">c</emphasis>
2 not modified, pointed...................................
<taxonomicName id="EEB0E8F6FE52BD37C4D406EC4316F854" authority="Khaustov and Trach, 2018" authorityName="Khaustov and Trach" authorityYear="2018" box="[930,1289,1926,1947]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="altaicus">
<emphasis id="1BC44F67FE52BD37C4D406EC4219F854" box="[930,1030,1926,1947]" italics="true" pageId="5" pageNumber="282">S. altaicus</emphasis>
<bibRefCitation id="4D21EE84FE52BD37C37A06EC4316F854" box="[1036,1289,1926,1947]" pageId="5" pageNumber="282" refId="ref8366" refString="Khaustov, A. A. &amp; Trach, V. A. (2018) New species of Spatulaphorus (Acari: Heterostigmata: Pygmephoridae) associated with Geotrupes baicalicus (Coleoptera: Geotrupidae) from Altai Republic, Russia. Systematic &amp; Applied Acarology, 23, 123 - 131. https: // doi. org / 10.11158 / saa. 23.1.10" type="journal article">Khaustov and Trach, 2018</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="51A7D3E5FE52BD37C26006EC4344F854" box="[1302,1371,1926,1947]" name="Russia" pageId="5" pageNumber="282">Russia</collectingCountry>
:
<collectingRegion id="EB745D97FE52BD37C21206EC438AF854" box="[1380,1429,1926,1947]" country="Russia" name="Altay" pageId="5" pageNumber="282">Altai</collectingRegion>
) 5. At least setae (
<emphasis id="1BC44F67FE52BD37C61806CC479AF874" box="[366,389,1958,1979]" italics="true" pageId="5" pageNumber="282">pv</emphasis>
) on tarsus II foliate distally................................................................ 6 - Setae (
<emphasis id="1BC44F67FE52BD37C65506AE4725F816" box="[291,314,1988,2009]" italics="true" pageId="5" pageNumber="282">pv</emphasis>
) on tarsus II not foliate distally.................................................................. 10 6. Tibiotarsus I with large, hook-like claw and a distinct counterpart opposite to the claw base........................... 7 - Tibiotarsus I with a small, almost straight claw, and no counterpart.........
<taxonomicName id="EEB0E8F6FE51BD34C4A101FD4383FF63" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[983,1436,151,172]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="1BC44F67FE51BD34C4A101FD4228FF63" box="[983,1079,151,172]" italics="true" pageId="6" pageNumber="283">S. foliatus</emphasis>
<bibRefCitation id="4D21EE84FE51BD34C34A01FD4338FF63" box="[1084,1319,151,172]" pageId="6" pageNumber="283" refId="ref7015" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych and Rack, 1993</bibRefCitation>
(
<collectingCountry id="51A7D3E5FE51BD34C24501FD438AFF63" box="[1331,1429,151,172]" name="Botswana" pageId="6" pageNumber="283">Botswana</collectingCountry>
)
</taxonomicName>
7. Tarsi II and III with setae
<emphasis id="1BC44F67FE51BD34C6AF01DF47EDFF05" box="[473,498,181,202]" italics="true" pageId="6" pageNumber="283">tc’</emphasis>
not modified.................................................................. 8 - Tarsi II and III with setae
<emphasis id="1BC44F67FE51BD34C6AF01B847EDFF28" box="[473,498,210,231]" italics="true" pageId="6" pageNumber="283">tc’</emphasis>
spiniform...................................
<taxonomicName id="EEB0E8F6FE51BD34C48D01B84384FF28" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[1019,1435,210,231]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="1BC44F67FE51BD34C48D01B8425BFF28" box="[1019,1092,210,231]" italics="true" pageId="6" pageNumber="283">S. langi</emphasis>
<bibRefCitation id="4D21EE84FE51BD34C33C01B8432AFF28" box="[1098,1333,210,231]" pageId="6" pageNumber="283" refId="ref7015" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych and Rack, 1993</bibRefCitation>
(
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)
</taxonomicName>
8. Setae
<emphasis id="1BC44F67FE51BD34C66D0185472FFECB" box="[283,304,239,260]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
3 subequal to or shorter than
<emphasis id="1BC44F67FE51BD34C53401854448FECB" box="[578,599,239,260]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
1, 2.................................................................. 9 - Setae
<emphasis id="1BC44F67FE51BD34C66D0065472FFEEB" box="[283,304,271,292]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
3 distinctly longer than
<emphasis id="1BC44F67FE51BD34C5780065443CFEEB" box="[526,547,271,292]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
1, 2..........................
<taxonomicName id="EEB0E8F6FE51BD34C4F600654315FEEB" authority="Dastych, Rack &amp; Camerik, 1997" authorityName="Dastych, Rack &amp; Camerik" authorityYear="1997" box="[896,1290,271,292]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="luriei">
<emphasis id="1BC44F67FE51BD34C4F6006545D3FEEB" box="[896,972,271,292]" italics="true" pageId="6" pageNumber="283">S. luriei</emphasis>
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</taxonomicName>
(
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) 9. Setae (
<emphasis id="1BC44F67FE51BD34C65500454725FE8B" box="[291,314,303,324]" italics="true" pageId="6" pageNumber="283">pv</emphasis>
) on tarsus III foliate, setae
<emphasis id="1BC44F67FE51BD34C5360045444AFE8B" box="[576,597,303,324]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
1, 2 distinctly barbed.......................
<taxonomicName id="EEB0E8F6FE51BD34C35C004542C1FE8B" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[1066,1246,303,324]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
<emphasis id="1BC44F67FE51BD34C35C004542C1FE8B" box="[1066,1246,303,324]" italics="true" pageId="6" pageNumber="283">S. enoplotruporum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE51BD34C3920044432DFE8B" box="[1252,1330,302,324]" pageId="6" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(
<collectingCountry id="51A7D3E5FE51BD34C24B0045438AFE8B" box="[1341,1429,303,324]" name="Thailand" pageId="6" pageNumber="283">Thailand</collectingCountry>
) - Setae (
<emphasis id="1BC44F67FE51BD34C6550024475DFEAC" box="[291,322,334,355]" italics="true" pageId="6" pageNumber="283">pv)</emphasis>
on tarsus III not foliate, setae
<emphasis id="1BC44F67FE51BD34C51300244465FEAC" box="[613,634,334,355]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
1, 2 smooth..................
<taxonomicName id="EEB0E8F6FE51BD34C4CD00244383FEAC" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[955,1436,334,355]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="camerikae">
<emphasis id="1BC44F67FE51BD34C4CD00244229FEAC" box="[955,1078,334,355]" italics="true" pageId="6" pageNumber="283">S. camerikae</emphasis>
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(
<collectingCountry id="51A7D3E5FE51BD34C24200244389FEAC" box="[1332,1430,334,355]" name="Botswana" pageId="6" pageNumber="283">Botswana</collectingCountry>
)
</taxonomicName>
10. Tarsi II and III with setae
<emphasis id="1BC44F67FE51BD34C6AF000447E5FE4C" box="[473,506,366,387]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
not modified................................................................. 11 - Tarsi II and III with setae
<emphasis id="1BC44F67FE51BD34C6AF00E647E5FE6E" box="[473,506,396,417]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
spiniform..................
<taxonomicName id="EEB0E8F6FE51BD34C44900E64313FE6E" authority="Rahiminejad and Hajiqanbar, 2011" authorityName="Rahiminejad and Hajiqanbar" authorityYear="2011" box="[831,1292,396,417]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="gorganica">
<emphasis id="1BC44F67FE51BD34C44900E645A6FE6E" box="[831,953,396,417]" italics="true" pageId="6" pageNumber="283">S. gorganica</emphasis>
<bibRefCitation id="4D21EE84FE51BD34C4C900E64313FE6E" box="[959,1292,396,417]" pageId="6" pageNumber="283" refId="ref8646" refString="Rahiminejad, V., Hajiqanbar, H. &amp; Fathipour, Y. (2011) A new species of the genus Spatulaphorus (Acari: Heterostigmatina: Pygmephoridae) phoretic on Geotrupes spiniger (Coleoptera: Geotrupidae) from Iran. Annales Zoologici, 61, 547 551. https: // doi. org / 10.3161 / 000345411 X 603391" type="journal article">Rahiminejad and Hajiqanbar, 2011</bibRefCitation>
</taxonomicName>
(
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,
<collectingRegion id="EB745D97FE51BD34C23B00E6438AFE6E" box="[1357,1429,396,417]" country="Ukraine" name="Crimea" pageId="6" pageNumber="283">Crimea</collectingRegion>
) 11. Apodemes 4 long, exceeding beyond setae 3
<emphasis id="1BC44F67FE51BD34C5FA00C34487FE71" box="[652,664,425,446]" italics="true" pageId="6" pageNumber="283">b</emphasis>
.............................................................. 12 - Apodemes 4 vestigial.................................................
<taxonomicName id="EEB0E8F6FE51BD34C37100AC4384FE14" authority="Khaustov, 2005" authorityName="Khaustov" authorityYear="2005" box="[1031,1435,454,475]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="geotruporum">
<emphasis id="1BC44F67FE51BD34C37100AC4283FE14" box="[1031,1180,454,475]" italics="true" pageId="6" pageNumber="283">S. geotruporum</emphasis>
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)
</taxonomicName>
12. Setae
<emphasis id="1BC44F67FE51BD34C66D008E4739FE36" box="[283,294,484,505]" italics="true" pageId="6" pageNumber="283">e</emphasis>
and
<emphasis id="1BC44F67FE51BD34C622008E477FFE36" box="[340,352,484,505]" italics="true" pageId="6" pageNumber="283">h</emphasis>
2 subequal................................................................................ 13 - Setae
<emphasis id="1BC44F67FE51BD34C66D03694739FDD7" box="[283,294,515,536]" italics="true" pageId="6" pageNumber="283">e</emphasis>
more than two times longer than
<emphasis id="1BC44F67FE51BD34C51E0369446BFDD7" box="[616,628,515,536]" italics="true" pageId="6" pageNumber="283">h</emphasis>
2..................................
<taxonomicName id="EEB0E8F6FE51BD34C36A03694383FDD7" authority="(Mahunka, 1970) (Hungary)" authorityName="Mahunka" authorityYear="1970" baseAuthorityName="Mahunka" baseAuthorityYear="1970" box="[1052,1436,515,536]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="geotrupi">
<emphasis id="1BC44F67FE51BD34C36A0369429BFDD7" box="[1052,1156,515,536]" italics="true" pageId="6" pageNumber="283">S. geotrupi</emphasis>
(
<bibRefCitation id="4D21EE84FE51BD34C3E403694334FDD7" box="[1170,1323,515,536]" pageId="6" pageNumber="283" refId="ref8585" refString="Mahunka, S. (1970) Considerations on the systematics of the Tarsonemina and the description of new European taxa (Acari: Trombidiformes). Acta Zoologica Academiae Scientiarum Hungaricae, 16, 137 - 174." type="journal article">Mahunka, 1970</bibRefCitation>
) (
<collectingCountry id="51A7D3E5FE51BD34C2480369438AFDD7" box="[1342,1429,515,536]" name="Hungary" pageId="6" pageNumber="283">Hungary</collectingCountry>
)
</taxonomicName>
13. Setae
<emphasis id="1BC44F67FE51BD34C66D03494739FDF7" box="[283,294,547,568]" italics="true" pageId="6" pageNumber="283">e</emphasis>
and
<emphasis id="1BC44F67FE51BD34C6220349477FFDF7" box="[340,352,547,568]" italics="true" pageId="6" pageNumber="283">h</emphasis>
2 barbed; setae
<emphasis id="1BC44F67FE51BD34C6860349441AFDF7" box="[496,517,547,568]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
3 more than two times longer than
<emphasis id="1BC44F67FE51BD34C4390349457BFDF7" box="[847,868,547,568]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
1, 2.....
<taxonomicName id="EEB0E8F6FE51BD34C4CA03494383FDF7" authority="Sobhi and Hajiqanbar, 2017" authorityName="Sobhi and Hajiqanbar" authorityYear="2017" box="[956,1436,547,568]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="gymnopleuri">
<emphasis id="1BC44F67FE51BD34C4CA03494252FDF7" box="[956,1101,547,568]" italics="true" pageId="6" pageNumber="283">S. gymnopleuri</emphasis>
Sobhi and Hajiqanbar, 2017 (
<collectingCountry id="51A7D3E5FE51BD34C21A0349438AFDF7" box="[1388,1429,547,568]" name="Iran" pageId="6" pageNumber="283">Iran</collectingCountry>
)
</taxonomicName>
- Setae
<emphasis id="1BC44F67FE51BD34C66D03294739FD97" box="[283,294,579,600]" italics="true" pageId="6" pageNumber="283">e</emphasis>
and
<emphasis id="1BC44F67FE51BD34C6220329477FFD97" box="[340,352,579,600]" italics="true" pageId="6" pageNumber="283">h</emphasis>
2 smooth; setae
<emphasis id="1BC44F67FE51BD34C68303294415FD97" box="[501,522,579,600]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
3 subequal to
<emphasis id="1BC44F67FE51BD34C5FD032944BFFD97" box="[651,672,579,600]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
1, 2.....................
<taxonomicName id="EEB0E8F6FE51BD34C4CC03294383FD97" authority="Khaustov and Frolov, 2017" authorityName="Khaustov and Frolov" authorityYear="2017" box="[954,1436,579,600]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="braziliensis">
<emphasis id="1BC44F67FE51BD34C4CC032945D9FD97" box="[954,966,579,600]" italics="true" pageId="6" pageNumber="283">S</emphasis>
.
<emphasis id="1BC44F67FE51BD34C4A40329425FFD97" box="[978,1088,579,600]" italics="true" pageId="6" pageNumber="283">braziliensis</emphasis>
<bibRefCitation id="4D21EE84FE51BD34C33003294354FD97" box="[1094,1355,579,600]" pageId="6" pageNumber="283" refId="ref7997" refString="Khaustov, A. A. &amp; Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article">Khaustov and Frolov, 2017</bibRefCitation>
(
<collectingCountry id="51A7D3E5FE51BD34C22E0329438AFD97" box="[1368,1429,579,600]" name="Brazil" pageId="6" pageNumber="283">Brazil</collectingCountry>
)
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
</subSubSection>
</treatment>
</document>

73
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Genus
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<paragraph id="290F9375FE56BD33C7E10459423BFA87" blockId="1.[151,1060,1330,1352]" box="[151,1060,1330,1352]" pageId="1" pageNumber="278">
Type species:
<taxonomicName id="EEB0E8F6FE56BD33C65004584536FA87" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[294,809,1330,1352]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="278" phylum="Arthropoda" rank="species" species="camerikae">
<emphasis id="1BC44F67FE56BD33C65004584435FA87" box="[294,554,1330,1352]" italics="true" pageId="1" pageNumber="278">Spatulaphorus camerikae</emphasis>
<bibRefCitation id="4D21EE84FE56BD33C55904584536FA87" box="[559,809,1330,1352]" pageId="1" pageNumber="278" refId="ref7015" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych and Rack, 1993</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="A1192263FE56BD36C7E104F3437CF81A" ID-DOI="http://doi.org/10.5281/zenodo.5949607" ID-GBIF-Taxon="146044081" ID-Zenodo-Dep="5949607" LSID="urn:lsid:plazi:treatment:A1192263FE56BD36C7E104F3437CF81A" httpUri="http://treatment.plazi.org/id/A1192263FE56BD36C7E104F3437CF81A" lastPageId="4" lastPageNumber="281" pageId="1" pageNumber="278">
<subSubSection id="61AAC0FEFE56BD33C7E104F34715FA1A" pageId="1" pageNumber="278" type="nomenclature">
<paragraph id="290F9375FE56BD33C7E104F34465FA7C" blockId="1.[151,634,1433,1493]" box="[151,634,1433,1459]" pageId="1" pageNumber="278">
<heading id="72472419FE56BD33C7E104F34465FA7C" bold="true" box="[151,634,1433,1459]" fontSize="11" level="1" pageId="1" pageNumber="278" reason="1">
<emphasis id="1BC44F67FE56BD33C7E104F34465FA7C" bold="true" box="[151,634,1433,1459]" pageId="1" pageNumber="278">
<taxonomicName id="EEB0E8F6FE56BD33C7E104F3440AFA7C" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[151,533,1433,1459]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="278" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
<emphasis id="1BC44F67FE56BD33C7E104F3440AFA7C" bold="true" box="[151,533,1433,1459]" italics="true" pageId="1" pageNumber="278">Spatulaphorus enoplotruporum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE56BD33C56A04F34465FA7C" box="[540,634,1433,1459]" pageId="1" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="290F9375FE56BD33C7E104D64715FA1A" blockId="1.[151,634,1433,1493]" box="[151,266,1468,1493]" pageId="1" pageNumber="278">
(
<figureCitation id="B18B8FF0FE56BD33C7E904D6471EFA1A" box="[159,257,1468,1493]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1393,1415]" captionStartId-1="3.[151,250,1954,1976]" captionStartId-2="4.[151,250,1330,1352]" captionStartId-3="5.[151,250,1546,1568]" captionStartId-4="7.[151,250,1804,1826]" captionTargetBox-0="[179,1414,369,1343]" captionTargetBox-1="[209,1332,690,1862]" captionTargetBox-2="[160,1411,246,1289]" captionTargetBox-3="[191,1406,208,1529]" captionTargetBox-4="[151,1436,180,1782]" captionTargetId-0="figure@2.[151,1436,312,1369]" captionTargetId-1="figure@3.[181,1405,643,1923]" captionTargetId-2="figure@4.[151,1436,193,1309]" captionTargetId-3="figure@5.[167,1421,193,1533]" captionTargetId-4="figure@7.[151,1436,180,1782]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionText-0="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." captionText-2="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." captionText-3="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." captionText-4="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri-0="https://zenodo.org/record/1303127/files/figure.png" httpUri-1="https://zenodo.org/record/1303129/files/figure.png" httpUri-2="https://zenodo.org/record/1303131/files/figure.png" httpUri-3="https://zenodo.org/record/1303133/files/figure.png" httpUri-4="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Figs 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE56BD31C7E1076E4792FDF0" lastPageId="3" lastPageNumber="280" pageId="1" pageNumber="278" type="description">
<paragraph id="290F9375FE56BD33C7E1076E45BEF8F3" blockId="1.[151,1437,1540,2039]" pageId="1" pageNumber="278">
<emphasis id="1BC44F67FE56BD33C7E1076E473EF9D2" bold="true" box="[151,289,1540,1565]" pageId="1" pageNumber="278">Description</emphasis>
. FEMALE (
<figureCitation id="B18B8FF0FE56BD33C6C9076E4433F9D2" box="[447,556,1540,1565]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1393,1415]" captionStartId-1="3.[151,250,1954,1976]" captionStartId-2="4.[151,250,1330,1352]" captionStartId-3="5.[151,250,1546,1568]" captionStartId-4="7.[151,250,1804,1826]" captionTargetBox-0="[179,1414,369,1343]" captionTargetBox-1="[209,1332,690,1862]" captionTargetBox-2="[160,1411,246,1289]" captionTargetBox-3="[191,1406,208,1529]" captionTargetBox-4="[151,1436,180,1782]" captionTargetId-0="figure@2.[151,1436,312,1369]" captionTargetId-1="figure@3.[181,1405,643,1923]" captionTargetId-2="figure@4.[151,1436,193,1309]" captionTargetId-3="figure@5.[167,1421,193,1533]" captionTargetId-4="figure@7.[151,1436,180,1782]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionText-0="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." captionText-2="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." captionText-3="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." captionText-4="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri-0="https://zenodo.org/record/1303127/files/figure.png" httpUri-1="https://zenodo.org/record/1303129/files/figure.png" httpUri-2="https://zenodo.org/record/1303131/files/figure.png" httpUri-3="https://zenodo.org/record/1303133/files/figure.png" httpUri-4="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Figs 1–5</figureCitation>
). Body well sclerotized. Length of idiosoma 205 (175–240), width 120 (100–140).
<emphasis id="1BC44F67FE56BD33C66C074D47B6F98F" box="[282,425,1575,1600]" italics="true" pageId="1" pageNumber="278">Gnathosoma</emphasis>
(
<figureCitation id="B18B8FF0FE56BD33C6CF074D47E1F98F" box="[441,510,1575,1600]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[209,1332,690,1862]" captionTargetId="figure@3.[181,1405,643,1923]" captionTargetPageId="3" captionText="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." httpUri="https://zenodo.org/record/1303129/files/figure.png" pageId="1" pageNumber="278">Fig. 2</figureCitation>
). Gnathosomal capsule, excluding palps, subrectangular, length 22 (19–24), width 22 (19–24). Dorsum with one pair of needle-like cheliceral setae (
<emphasis id="1BC44F67FE56BD33C4E6072645A6F9AA" box="[912,953,1612,1637]" italics="true" pageId="1" pageNumber="278">сha</emphasis>
) 7 (5–7). Dorsal median apodeme absent. Postpalpal setae (
<emphasis id="1BC44F67FE56BD33C613071B479EF947" box="[357,385,1649,1672]" italics="true" pageId="1" pageNumber="278">pp</emphasis>
) spine-like, situated laterally to setae
<emphasis id="1BC44F67FE56BD33C43F0705456EF947" box="[841,881,1647,1672]" italics="true" pageId="1" pageNumber="278">сha</emphasis>
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
<emphasis id="1BC44F67FE56BD33C6B507FF47C8F963" box="[451,471,1685,1708]" italics="true" pageId="1" pageNumber="278">m</emphasis>
6 (5–7) and round pits situated posteriad to
<emphasis id="1BC44F67FE56BD33C4B807FF45FDF963" box="[974,994,1685,1708]" italics="true" pageId="1" pageNumber="278">m</emphasis>
. Palps with smooth setae
<emphasis id="1BC44F67FE56BD33C27C07FE4329F962" box="[1290,1334,1684,1709]" italics="true" pageId="1" pageNumber="278">dFe</emphasis>
and
<emphasis id="1BC44F67FE56BD33C21B07FE4383F962" box="[1389,1436,1684,1709]" italics="true" pageId="1" pageNumber="278">dGe</emphasis>
dorsolaterally, setae
<emphasis id="1BC44F67FE56BD33C6F607DD47B0F91F" box="[384,431,1719,1744]" italics="true" pageId="1" pageNumber="278">dGe</emphasis>
pointed, about twice longer than needle-like
<emphasis id="1BC44F67FE56BD33C4C007DD45FDF91F" box="[950,994,1719,1744]" italics="true" pageId="1" pageNumber="278">dFe</emphasis>
. Palps ventrally each with mushroomlike accessory setigenous structure (
<emphasis id="1BC44F67FE56BD33C54A07B7447FF93B" box="[572,608,1757,1780]" italics="true" pageId="1" pageNumber="278">ass</emphasis>
) and tiny solenidion. Palps terminated with small tibial claw and tiny eupathidium-like seta. Pharyngeal pump 2 with weak transverse striation; pharyngeal pump 1 small, butterfly-like; pharyngeal pump 3 oval, slightly smaller than oval pump 2 (
<figureCitation id="B18B8FF0FE56BD33C44D064E458BF8F3" box="[827,916,1828,1853]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[209,1332,690,1862]" captionTargetId="figure@3.[181,1405,643,1923]" captionTargetPageId="3" captionText="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." httpUri="https://zenodo.org/record/1303129/files/figure.png" pageId="1" pageNumber="278">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="290F9375FE56BD30C7B1062D4319FF31" blockId="1.[151,1437,1540,2039]" lastBlockId="2.[151,1435,151,258]" lastPageId="2" lastPageNumber="279" pageId="1" pageNumber="278">
<emphasis id="1BC44F67FE56BD33C7B1062D478CF8AF" box="[199,403,1863,1888]" italics="true" pageId="1" pageNumber="278">Idiosomal dorsum</emphasis>
(
<figureCitation id="B18B8FF0FE56BD33C6D5062D441DF8AF" box="[419,514,1863,1888]" captionStart="FIGURE 1" captionStartId="2.[151,250,1393,1415]" captionTargetBox="[179,1414,369,1343]" captionTargetId="figure@2.[151,1436,312,1369]" captionTargetPageId="2" captionText="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/1303127/files/figure.png" pageId="1" pageNumber="278">Figs 1A</figureCitation>
,
<figureCitation id="B18B8FF0FE56BD33C566062D4483F8AF" box="[528,668,1863,1888]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">5A, C, E, G</figureCitation>
). All dorsal shields with numerous tiny, barely discernable dimples (
<figureCitation id="B18B8FF0FE56BD33C7E90606476FF84B" box="[159,368,1900,1925]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Figs 5A, C, E, G</figureCitation>
). Prodorsal shield with three pairs of setae (
<date id="5D0EB5B5FE56BD33C40B0607458DF846" box="[893,914,1901,1929]" pageId="1" pageNumber="278">
<emphasis id="1BC44F67FE56BD33C40B06074596F84B" box="[893,905,1901,1924]" italics="true" pageId="1" pageNumber="278">v</emphasis>
1
</date>
,
<date id="5D0EB5B5FE56BD33C4D2060745A7F846" box="[932,952,1901,1929]" pageId="1" pageNumber="278">
<emphasis id="1BC44F67FE56BD33C4D2060745AFF84B" box="[932,944,1901,1924]" italics="true" pageId="1" pageNumber="278">v</emphasis>
2
</date>
,
<emphasis id="1BC44F67FE56BD33C4BC060745FEF84B" box="[970,993,1901,1924]" italics="true" pageId="1" pageNumber="278">sс</emphasis>
2), one pair of weakly barbed capitate trichobothria and one pair of round stigmata (
<figureCitation id="B18B8FF0FE56BD33C5D606F8451DF864" box="[672,770,1938,1963]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Fig. 5 C</figureCitation>
). Stigmata associated with well developed tracheal trunks. All dorsal idiosomal setae blunt-ended and barbed. Cupules
<emphasis id="1BC44F67FE56BD33C43F06D24540F800" box="[841,863,1976,1999]" italics="true" pageId="1" pageNumber="278">ia</emphasis>
on tergite D,
<emphasis id="1BC44F67FE56BD33C37606D24203F800" box="[1024,1052,1976,1999]" italics="true" pageId="1" pageNumber="278">im</emphasis>
on tergite EF and
<emphasis id="1BC44F67FE56BD33C38206DC4315F800" box="[1268,1290,1974,1999]" italics="true" pageId="1" pageNumber="278">ih</emphasis>
on tergite H large, round. Tergites D, EF and H each with one pair of oval porous areas. Lengths of dorsal setae:
<date id="5D0EB5B5FE56BD33C38B06B6432AF83C" box="[1277,1333,2011,2039]" pageId="1" pageNumber="278" value="1923-05-01">
<emphasis id="1BC44F67FE56BD33C38B06B64316F83C" box="[1277,1289,2012,2035]" italics="true" pageId="1" pageNumber="278">v</emphasis>
1 23
</date>
(21–23),
<date id="5D0EB5B5FE55BD30C7E101F346D1FF7F" box="[151,206,152,180]" pageId="2" pageNumber="279" value="2013-05-02">
<emphasis id="1BC44F67FE55BD30C7E101F346BCFF7F" box="[151,163,153,176]" italics="true" pageId="2" pageNumber="279">v</emphasis>
2 13
</date>
(10–13),
<emphasis id="1BC44F67FE55BD30C64C01F3474EFF7F" box="[314,337,153,176]" italics="true" pageId="2" pageNumber="279">sс</emphasis>
2 49 (45–54),
<emphasis id="1BC44F67FE55BD30C69F01F347EAFF7F" box="[489,501,153,176]" italics="true" pageId="2" pageNumber="279">с</emphasis>
1 37 (30–38),
<emphasis id="1BC44F67FE55BD30C5FF01F3448AFF7F" box="[649,661,153,176]" italics="true" pageId="2" pageNumber="279">с</emphasis>
2 49 (44–54),
<emphasis id="1BC44F67FE55BD30C45B01FD4524FF7F" box="[813,827,151,176]" italics="true" pageId="2" pageNumber="279">d</emphasis>
42 (39–48),
<emphasis id="1BC44F67FE55BD30C4BC01F345C9FF7F" box="[970,982,153,176]" italics="true" pageId="2" pageNumber="279">e</emphasis>
25 (23–31),
<emphasis id="1BC44F67FE55BD30C31001F24271FF7F" box="[1126,1134,152,176]" italics="true" pageId="2" pageNumber="279">f</emphasis>
54 (44–58),
<emphasis id="1BC44F67FE55BD30C38B01FD4314FF7F" box="[1277,1291,151,176]" italics="true" pageId="2" pageNumber="279">h</emphasis>
1 51 (45–57),
<emphasis id="1BC44F67FE55BD30C7E101D446BAFF18" box="[151,165,190,215]" italics="true" pageId="2" pageNumber="279">h</emphasis>
2 16 (16–21). Distances between setae:
<date id="5D0EB5B5FE55BD30C51201AA44A0FF18" box="[612,703,191,219]" pageId="2" pageNumber="279" value="1922-05-01" valueMax="1922-05-01" valueMin="1922-05-01">
<emphasis id="1BC44F67FE55BD30C51201AA446FFF18" box="[612,624,192,215]" italics="true" pageId="2" pageNumber="279">v</emphasis>
1–
<emphasis id="1BC44F67FE55BD30C5F001AA448DFF18" box="[646,658,192,215]" italics="true" pageId="2" pageNumber="279">v</emphasis>
1 22
</date>
(20–23),
<date id="5D0EB5B5FE55BD30C45901AA4595FF18" box="[815,906,191,219]" pageId="2" pageNumber="279" value="1939-05-02" valueMax="1939-05-02" valueMin="1939-05-02">
<emphasis id="1BC44F67FE55BD30C45901AA4524FF18" box="[815,827,192,215]" italics="true" pageId="2" pageNumber="279">v</emphasis>
2
<emphasis id="1BC44F67FE55BD30C43501AA4541FF18" box="[835,862,192,215]" italics="true" pageId="2" pageNumber="279">–v</emphasis>
2 39
</date>
(34–42),
<emphasis id="1BC44F67FE55BD30C48C01AA420EFF18" box="[1018,1041,192,215]" italics="true" pageId="2" pageNumber="279">sс</emphasis>
2
<emphasis id="1BC44F67FE55BD30C36F01AA4221FF18" box="[1049,1086,192,215]" italics="true" pageId="2" pageNumber="279">–sс</emphasis>
2 47 (39–49),
<emphasis id="1BC44F67FE55BD30C3AD01AA42F8FF18" box="[1243,1255,192,215]" italics="true" pageId="2" pageNumber="279">с</emphasis>
1–
<emphasis id="1BC44F67FE55BD30C38B01AA4316FF18" box="[1277,1289,192,215]" italics="true" pageId="2" pageNumber="279">с</emphasis>
1 46 (39–47),
<emphasis id="1BC44F67FE55BD30C7E1018C46BCFF32" box="[151,163,230,253]" italics="true" pageId="2" pageNumber="279">с</emphasis>
1–
<emphasis id="1BC44F67FE55BD30C7CF018C46DAFF32" box="[185,197,230,253]" italics="true" pageId="2" pageNumber="279">с</emphasis>
2 27 (23–29),
<emphasis id="1BC44F67FE55BD30C628018F4797FF31" box="[350,392,229,254]" italics="true" pageId="2" pageNumber="279">d–d</emphasis>
68 (55–74),
<emphasis id="1BC44F67FE55BD30C56E018C4424FF31" box="[536,571,230,254]" italics="true" pageId="2" pageNumber="279">e–f</emphasis>
17 (14–17),
<emphasis id="1BC44F67FE55BD30C5BD018C44F6FF31" box="[715,745,230,254]" italics="true" pageId="2" pageNumber="279">f–f</emphasis>
40 (31–46),
<emphasis id="1BC44F67FE55BD30C40F018F4598FF31" box="[889,903,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
1–
<emphasis id="1BC44F67FE55BD30C4EB018F45B4FF31" box="[925,939,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
1 26 (22–32),
<emphasis id="1BC44F67FE55BD30C335018F424EFF31" box="[1091,1105,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
1–
<emphasis id="1BC44F67FE55BD30C311018F426AFF31" box="[1127,1141,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
2 18 (14–20).
</paragraph>
<caption id="7DCFC3FDFE55BD30C7E1041B439CFA48" box="[151,1411,1393,1415]" httpUri="https://zenodo.org/record/1303127/files/figure.png" pageId="2" pageNumber="279" startId="2.[151,250,1393,1415]" targetBox="[179,1414,369,1343]" targetPageId="2">
<paragraph id="290F9375FE55BD30C7E1041B439CFA48" blockId="2.[151,1411,1393,1415]" box="[151,1411,1393,1415]" pageId="2" pageNumber="279">
<emphasis id="1BC44F67FE55BD30C7E1041B4712FA49" bold="true" box="[151,269,1393,1415]" pageId="2" pageNumber="279">FIGURE 1</emphasis>
.
<taxonomicName id="EEB0E8F6FE55BD30C66F041B4447FA48" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[281,600,1393,1415]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
<emphasis id="1BC44F67FE55BD30C66F041B4447FA48" box="[281,600,1393,1415]" italics="true" pageId="2" pageNumber="279">Spatulaphorus enoplotruporum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE55BD30C528041B44B1FA49" box="[606,686,1393,1414]" pageId="2" pageNumber="279" rank="species">
<emphasis id="1BC44F67FE55BD30C528041B44B1FA49" bold="true" box="[606,686,1393,1414]" pageId="2" pageNumber="279">sp. nov.</emphasis>
</taxonomicNameLabel>
, female: A—dorsum of the body, B—venter of the body. Legs omitted.
</paragraph>
</caption>
<paragraph id="290F9375FE55BD30C7B104C44515F8FB" blockId="2.[151,1437,1454,2027]" pageId="2" pageNumber="279">
<emphasis id="1BC44F67FE55BD30C7B104C44799FA09" box="[199,390,1454,1479]" italics="true" pageId="2" pageNumber="279">Idiosomal venter</emphasis>
(
<figureCitation id="B18B8FF0FE55BD30C6EE04C447E9FA08" box="[408,502,1454,1479]" captionStart="FIGURE 1" captionStartId="2.[151,250,1393,1415]" captionTargetBox="[179,1414,369,1343]" captionTargetId="figure@2.[151,1436,312,1369]" captionTargetPageId="2" captionText="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/1303127/files/figure.png" pageId="2" pageNumber="279">Figs 1B</figureCitation>
,
<figureCitation id="B18B8FF0FE55BD30C57304C444B5FA08" box="[517,682,1454,1479]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="2" pageNumber="279">5B, D, F, H, I</figureCitation>
). All ventral plates with numerous tiny dimples. Setae 1
<emphasis id="1BC44F67FE55BD30C24E04C44359FA08" box="[1336,1350,1454,1479]" italics="true" pageId="2" pageNumber="279">b</emphasis>
and 2
<emphasis id="1BC44F67FE55BD30C2F904C54384FA09" box="[1423,1435,1455,1478]" italics="true" pageId="2" pageNumber="279">с</emphasis>
pointed; other ventral setae with slightly rounded tips. Setae 1
<emphasis id="1BC44F67FE55BD30C40604B84561FA24" box="[880,894,1490,1515]" italics="true" pageId="2" pageNumber="279">b</emphasis>
and
<emphasis id="1BC44F67FE55BD30C4CA04B945CAFA25" box="[956,981,1491,1514]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
3 with a few barbs; setae
<emphasis id="1BC44F67FE55BD30C27504B94303FA25" box="[1283,1308,1491,1514]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
1, 2 strongly barbed; other ventral setae smooth. Bases of setae
<emphasis id="1BC44F67FE55BD30C5A2049044F2F9DE" box="[724,749,1530,1553]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
3 situated distinctly anteriad bases of setae
<emphasis id="1BC44F67FE55BD30C3A4049042F5F9DE" box="[1234,1258,1530,1553]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
1-2. Apodemes 1 (ap1) and apodemes 2 (ap2) well developed and joined with thick prosternal apodeme (appr); sejugal apodeme (apsej) developed only laterally; apodemes 3 (ap3) weak, not reaching to poststernal apodeme (appo); apodemes 4 (ap4) well developed, exceeding beyond bases of setae 3
<emphasis id="1BC44F67FE55BD30C47A070D4505F94F" box="[780,794,1639,1664]" italics="true" pageId="2" pageNumber="279">b</emphasis>
. Apodemes 5 (ap5) well developed, joined with appo and reaching beyond setae 4
<emphasis id="1BC44F67FE55BD30C6D207E747ADF96B" box="[420,434,1677,1700]" italics="true" pageId="2" pageNumber="279">a</emphasis>
. Anterior genital sclerite (ags) small, cup-like; posterior genital sclerite (pgs) small, oval, with lateral processes (
<figureCitation id="B18B8FF0FE55BD30C6D507C5440BF907" box="[419,532,1711,1736]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="2" pageNumber="279">
Figure
<date id="5D0EB5B5FE55BD30C68C07C5440BF907" box="[506,532,1711,1736]" pageId="2" pageNumber="279">5I</date>
</figureCitation>
). Posterior margins of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae: 1
<emphasis id="1BC44F67FE55BD30C4B607BF45D1F923" box="[960,974,1749,1772]" italics="true" pageId="2" pageNumber="279">a</emphasis>
9 (9–10), 1
<emphasis id="1BC44F67FE55BD30C32207BE427DF922" box="[1108,1122,1748,1773]" italics="true" pageId="2" pageNumber="279">b</emphasis>
17 (15–21), 1
<emphasis id="1BC44F67FE55BD30C27207BF430FF923" box="[1284,1296,1749,1772]" italics="true" pageId="2" pageNumber="279">с</emphasis>
11 (10–12), 2
<emphasis id="1BC44F67FE55BD30C7D3079346ACF8DF" box="[165,179,1785,1808]" italics="true" pageId="2" pageNumber="279">a</emphasis>
10 (10–11), 2
<emphasis id="1BC44F67FE55BD30C6230793477EF8DF" box="[341,353,1785,1808]" italics="true" pageId="2" pageNumber="279">с</emphasis>
23 (18–23), 3
<emphasis id="1BC44F67FE55BD30C5730793440CF8DF" box="[517,531,1785,1808]" italics="true" pageId="2" pageNumber="279">a</emphasis>
14 (12–16), 3
<emphasis id="1BC44F67FE55BD30C5C1079D44DAF8DF" box="[695,709,1783,1808]" italics="true" pageId="2" pageNumber="279">b</emphasis>
10 (9–11), 3
<emphasis id="1BC44F67FE55BD30C42C07934579F8DF" box="[858,870,1785,1808]" italics="true" pageId="2" pageNumber="279">с</emphasis>
12 (11–14), 4
<emphasis id="1BC44F67FE55BD30C37F07934208F8DF" box="[1033,1047,1785,1808]" italics="true" pageId="2" pageNumber="279">a</emphasis>
11 (10–13), 4
<emphasis id="1BC44F67FE55BD30C3CF079D42D8F8DF" box="[1209,1223,1783,1808]" italics="true" pageId="2" pageNumber="279">b</emphasis>
15 (14–17), 4
<emphasis id="1BC44F67FE55BD30C21D07934368F8DF" box="[1387,1399,1785,1808]" italics="true" pageId="2" pageNumber="279">с</emphasis>
12 (12–14),
<emphasis id="1BC44F67FE55BD30C78B06774709F8FB" box="[253,278,1821,1844]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
1 22 (21–25),
<emphasis id="1BC44F67FE55BD30C6D9067747D8F8FB" box="[431,455,1821,1844]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
2 23 (21–25),
<emphasis id="1BC44F67FE55BD30C51606774466F8FB" box="[608,633,1821,1844]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
3 18 (17–20).
</paragraph>
<paragraph id="290F9375FE55BD31C7B1062E4735FDD4" blockId="2.[151,1437,1454,2027]" lastBlockId="3.[151,1436,151,575]" lastPageId="3" lastPageNumber="280" pageId="2" pageNumber="279">
<emphasis id="1BC44F67FE55BD30C7B1062E46E3F894" box="[199,252,1860,1883]" italics="true" pageId="2" pageNumber="279">Legs</emphasis>
(
<figureCitation id="B18B8FF0FE55BD30C67D0628474CF894" box="[267,339,1858,1883]" captionStart="FIGURE 3" captionStartId="4.[151,250,1330,1352]" captionTargetBox="[160,1411,246,1289]" captionTargetId="figure@4.[151,1436,193,1309]" captionTargetPageId="4" captionText="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="2" pageNumber="279">Figs 3</figureCitation>
,
<figureCitation id="B18B8FF0FE55BD30C6160629476EF894" box="[352,369,1859,1883]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1568]" captionTargetBox="[191,1406,208,1529]" captionTargetId="figure@5.[167,1421,193,1533]" captionTargetPageId="5" captionText="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303133/files/figure.png" pageId="2" pageNumber="279">4</figureCitation>
). Leg I (
<figureCitation id="B18B8FF0FE55BD30C6A706284435F894" box="[465,554,1858,1883]" captionStart="FIGURE 3" captionStartId="4.[151,250,1330,1352]" captionTargetBox="[160,1411,246,1289]" captionTargetId="figure@4.[151,1436,193,1309]" captionTargetPageId="4" captionText="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="2" pageNumber="279">Fig. 3A</figureCitation>
) slightly thicker than leg II. Setal formula:
<date id="5D0EB5B5FE55BD30C36406294298F894" box="[1042,1159,1859,1883]" pageId="2" pageNumber="279" value="2017-01-04" valueMax="2017-01-02" valueMin="2017-01-04">1–4–2–17</date>
(4). Tibiotarsus enlarged, with large claw. Setae (
<emphasis id="1BC44F67FE55BD30C6EB060247B4F8B0" box="[413,427,1896,1919]" italics="true" pageId="2" pageNumber="279">u</emphasis>
) consolidated into a horn-like structure opposing to claw. Tarsal claw blunt-tipped. Seta
<emphasis id="1BC44F67FE55BD30C2E706024383F8B0" box="[1425,1436,1896,1919]" italics="true" pageId="2" pageNumber="279">s</emphasis>
distinctly foliate distally; seta
<emphasis id="1BC44F67FE55BD30C57606E14436F86D" box="[512,553,1931,1954]" italics="true" pageId="2" pageNumber="279">pv”</emphasis>
slightly foliate distally. Seta
<emphasis id="1BC44F67FE55BD30C4F806E04583F86C" box="[910,924,1930,1955]" italics="true" pageId="2" pageNumber="279">d</emphasis>
of femur spatulate, with short subterminal projection. Setae
<emphasis id="1BC44F67FE55BD30C62806C44770F808" box="[350,367,1966,1991]" italics="true" pageId="2" pageNumber="279">l’</emphasis>
of femur
<emphasis id="1BC44F67FE55BD30C69206C4441CF808" box="[484,515,1966,1991]" italics="true" pageId="2" pageNumber="279">pl’</emphasis>
and
<emphasis id="1BC44F67FE55BD30C54B06C44456F808" box="[573,585,1966,1991]" italics="true" pageId="2" pageNumber="279">k</emphasis>
of tibiotarsus smooth; other leg setae (except eupathidia) sparsely barbed. Setae
<emphasis id="1BC44F67FE55BD30C7AD06B846F2F824" box="[219,237,2002,2027]" italics="true" pageId="2" pageNumber="279">l’</emphasis>
of femur, (
<emphasis id="1BC44F67FE55BD30C61A06B8476BF824" box="[364,372,2002,2027]" italics="true" pageId="2" pageNumber="279">l</emphasis>
) of genu and
<emphasis id="1BC44F67FE55BD30C56706B84402F824" box="[529,541,2002,2027]" italics="true" pageId="2" pageNumber="279">k</emphasis>
of tibiotarsus blunt-ended; other leg setae (except eupathidia) pointed. Lengths of solenidia
<emphasis id="1BC44F67FE54BD31C65101F34724FF7F" box="[295,315,153,176]" italics="true" pageId="3" pageNumber="280">ω</emphasis>
1 8 (7–9),
<emphasis id="1BC44F67FE54BD31C6C401F347D9FF7F" box="[434,454,153,176]" italics="true" pageId="3" pageNumber="280">ω</emphasis>
2 6 (5–7),
<emphasis id="1BC44F67FE54BD31C54B01F34453FF7F" box="[573,588,153,176]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
1 8 (8–9),
<emphasis id="1BC44F67FE54BD31C5B201F344CCFF7F" box="[708,723,153,176]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
2 6 (5–6); Solenidion
<emphasis id="1BC44F67FE54BD31C4B801F345C2FF7F" box="[974,989,153,176]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
1 thick, clavate, other solenidia weakly clavate. Leg II (
<figureCitation id="B18B8FF0FE54BD31C63C01D447BEFF18" box="[330,417,190,215]" captionStart="FIGURE 3" captionStartId="4.[151,250,1330,1352]" captionTargetBox="[160,1411,246,1289]" captionTargetId="figure@4.[151,1436,193,1309]" captionTargetPageId="4" captionText="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="3" pageNumber="280">Fig. 3B</figureCitation>
). Setal formula: 1–3–1–4–6(1). Tarsus with padded claws and empodium with three lobes, one distal and two lateral. Solenidion
<emphasis id="1BC44F67FE54BD31C548018E444DFF34" box="[574,594,228,251]" italics="true" pageId="3" pageNumber="280">ω</emphasis>
6 (5–7), weakly clavate, solenidion
<emphasis id="1BC44F67FE54BD31C49F018E45E7FF34" box="[1001,1016,228,251]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
absent. Setae (
<emphasis id="1BC44F67FE54BD31C3D4018942D9FF35" box="[1186,1222,227,250]" italics="true" pageId="3" pageNumber="280">pv)</emphasis>
of tarsus distinctly foliate distally; setae
<emphasis id="1BC44F67FE54BD31C6F2006C478DFED0" box="[388,402,262,287]" italics="true" pageId="3" pageNumber="280">d</emphasis>
and
<emphasis id="1BC44F67FE54BD31C6BE006C47C6FED0" box="[456,473,262,287]" italics="true" pageId="3" pageNumber="280">l’</emphasis>
of femur blunt-ended; other leg setae pointed. Seta
<emphasis id="1BC44F67FE54BD31C36A006D422CFED1" box="[1052,1075,263,286]" italics="true" pageId="3" pageNumber="280">u’</emphasis>
of tarsus smooth and thin; other leg setae barbed. Leg III (
<figureCitation id="B18B8FF0FE54BD31C6CC00404409FE8C" box="[442,534,298,323]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1568]" captionTargetBox="[191,1406,208,1529]" captionTargetId="figure@5.[167,1421,193,1533]" captionTargetPageId="5" captionText="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303133/files/figure.png" pageId="3" pageNumber="280">Fig. 4A</figureCitation>
). Femur divided into basi- and telofemur. Setal formula: 1–2–1–4–6. Claws and empodium of same shape as on tarsus II. Setae (
<emphasis id="1BC44F67FE54BD31C5CF002544C2FEA9" box="[697,733,335,358]" italics="true" pageId="3" pageNumber="280">pv)</emphasis>
of tarsus distinctly foliate distally; setae
<emphasis id="1BC44F67FE54BD31C3D8002542DBFEA9" box="[1198,1220,335,358]" italics="true" pageId="3" pageNumber="280">v’</emphasis>
of trochanter,
<emphasis id="1BC44F67FE54BD31C21F00244368FEA8" box="[1385,1399,334,359]" italics="true" pageId="3" pageNumber="280">d</emphasis>
,
<emphasis id="1BC44F67FE54BD31C2F000254383FEA9" box="[1414,1436,335,358]" italics="true" pageId="3" pageNumber="280">v’</emphasis>
of femur and
<emphasis id="1BC44F67FE54BD31C6470018475DFE44" box="[305,322,370,395]" italics="true" pageId="3" pageNumber="280">l‘</emphasis>
of genu blunt-ended; other leg setae pointed. Seta
<emphasis id="1BC44F67FE54BD31C4F700194587FE45" box="[897,920,371,394]" italics="true" pageId="3" pageNumber="280">u’</emphasis>
of tarsus smooth; other leg setae barbed. Leg IV (
<figureCitation id="B18B8FF0FE54BD31C7B200FC4708FE60" box="[196,279,406,431]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1568]" captionTargetBox="[191,1406,208,1529]" captionTargetId="figure@5.[167,1421,193,1533]" captionTargetPageId="5" captionText="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303133/files/figure.png" pageId="3" pageNumber="280">Fig 4B</figureCitation>
). Femur divided into basi- and telofemur. Setal formula: 1–2–0–4–6. Claws simple, empodium round, stalked. Setae
<emphasis id="1BC44F67FE54BD31C64F00D14742FE1D" box="[313,349,443,466]" italics="true" pageId="3" pageNumber="280">tс”</emphasis>
,
<emphasis id="1BC44F67FE54BD31C61A00D1479CFE1D" box="[364,387,443,466]" italics="true" pageId="3" pageNumber="280">u’</emphasis>
, and (
<emphasis id="1BC44F67FE54BD31C6BD00D647FAFE1C" box="[459,485,444,467]" italics="true" pageId="3" pageNumber="280">pv</emphasis>
) slightly foliate distally (flattened tips difficult to discern in some specimens); setae
<emphasis id="1BC44F67FE54BD31C7E100B446B7FE38" box="[151,168,478,503]" italics="true" pageId="3" pageNumber="280">l’</emphasis>
,
<emphasis id="1BC44F67FE54BD31C7C100B546D2FE39" box="[183,205,479,502]" italics="true" pageId="3" pageNumber="280">v’</emphasis>
of tibia and t
<emphasis id="1BC44F67FE54BD31C61300B54764FE39" box="[357,379,479,502]" italics="true" pageId="3" pageNumber="280">с’</emphasis>
of tarsus pointed; other leg setae blunt-ended. Seta
<emphasis id="1BC44F67FE54BD31C4B300B445F5FE38" box="[965,1002,478,503]" italics="true" pageId="3" pageNumber="280">pl”</emphasis>
of tarsus smooth, spiniform; other leg setae barbed.
</paragraph>
<paragraph id="290F9375FE54BD31C7B1034D4792FDF0" blockId="3.[151,1436,151,575]" box="[199,397,550,575]" pageId="3" pageNumber="280">MALE unknown.</paragraph>
</subSubSection>
<caption id="7DCFC3FDFE54BD31C7E106C8479CF818" httpUri="https://zenodo.org/record/1303129/files/figure.png" pageId="3" pageNumber="280" startId="3.[151,250,1954,1976]" targetBox="[209,1332,690,1862]" targetPageId="3">
<paragraph id="290F9375FE54BD31C7E106C8479CF818" blockId="3.[151,1436,1954,2007]" pageId="3" pageNumber="280">
<emphasis id="1BC44F67FE54BD31C7E106C84712F878" bold="true" box="[151,269,1954,1976]" pageId="3" pageNumber="280">FIGURE 2</emphasis>
.
<taxonomicName id="EEB0E8F6FE54BD31C66F06C84447F877" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[281,600,1954,1976]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
<emphasis id="1BC44F67FE54BD31C66F06C84447F877" box="[281,600,1954,1976]" italics="true" pageId="3" pageNumber="280">Spatulaphorus enoplotruporum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE54BD31C52806C944B1F877" box="[606,686,1955,1976]" pageId="3" pageNumber="280" rank="species">
<emphasis id="1BC44F67FE54BD31C52806C944B1F877" bold="true" box="[606,686,1955,1976]" pageId="3" pageNumber="280">sp. nov.</emphasis>
</taxonomicNameLabel>
, female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps.
</paragraph>
</caption>
<caption id="7DCFC3FDFE53BD36C7E104584370FA87" box="[151,1391,1330,1352]" httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="4" pageNumber="281" startId="4.[151,250,1330,1352]" targetBox="[160,1411,246,1289]" targetPageId="4">
<paragraph id="290F9375FE53BD36C7E104584370FA87" blockId="4.[151,1391,1330,1352]" box="[151,1391,1330,1352]" pageId="4" pageNumber="281">
<emphasis id="1BC44F67FE53BD36C7E104584712FA88" bold="true" box="[151,269,1330,1352]" pageId="4" pageNumber="281">FIGURE 3</emphasis>
.
<taxonomicName id="EEB0E8F6FE53BD36C66F04584447FA87" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[281,600,1330,1352]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
<emphasis id="1BC44F67FE53BD36C66F04584447FA87" box="[281,600,1330,1352]" italics="true" pageId="4" pageNumber="281">Spatulaphorus enoplotruporum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE53BD36C528045844B1FA88" box="[606,686,1330,1351]" pageId="4" pageNumber="281" rank="species">
<emphasis id="1BC44F67FE53BD36C528045844B1FA88" bold="true" box="[606,686,1330,1351]" pageId="4" pageNumber="281">sp. nov.</emphasis>
</taxonomicNameLabel>
, female: A—right leg I in dorsal view, B—right leg II in dorsal view.
</paragraph>
</caption>
<subSubSection id="61AAC0FEFE53BD36C7B10405445AF9D7" pageId="4" pageNumber="281" type="materials_examined">
<paragraph id="290F9375FE53BD36C7B104054258FA63" blockId="4.[151,1436,1391,2006]" pageId="4" pageNumber="281">
<materialsCitation id="99D89928FE53BD36C7B1040542F0FA47" ID-GBIF-Occurrence="1852084072" box="[199,1263,1391,1416]" collectingDate="1986-10" collectionCode="ZISP" country="Thailand" location="Chiang Mai" pageId="4" pageNumber="281" specimenCount="2" stateProvince="Chiang Mai" typeStatus="holotype">
<emphasis id="1BC44F67FE53BD36C7B10405476CFA47" bold="true" box="[199,371,1391,1416]" pageId="4" pageNumber="281">
<typeStatus id="F60B2DD7FE53BD36C7B10405471DFA47" box="[199,258,1391,1416]" pageId="4" pageNumber="281">Type</typeStatus>
material
</emphasis>
. Female
<typeStatus id="F60B2DD7FE53BD36C69304054455FA47" box="[485,586,1391,1416]" pageId="4" pageNumber="281" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="4FA10BB0FE53BD36C5EB041A44C8FA47" box="[669,727,1392,1416]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="281">ZISP</collectionCode>
T-Pygm-001,
<collectingCountry id="51A7D3E5FE53BD36C4F5040545F6FA47" box="[899,1001,1391,1416]" name="Thailand" pageId="4" pageNumber="281">Thailand</collectingCountry>
,
<collectingRegion id="EB745D97FE53BD36C48C04054297FA47" box="[1018,1160,1391,1416]" country="Thailand" name="Chiang Mai" pageId="4" pageNumber="281">Chiang Mai</collectingRegion>
,
<collectingDate id="4D4A4C5DFE53BD36C3EF040542F0FA47" box="[1177,1263,1391,1416]" pageId="4" pageNumber="281" value="1986-10">
<date id="5D0EB5B5FE53BD36C3EF040542F0FA47" box="[1177,1263,1391,1416]" pageId="4" pageNumber="281" value="1986-10">X.1986</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="99D89928FE53BD36C276041A4771FA63" ID-GBIF-Occurrence="1852084069" collectingDate="1986-10" collectionCode="ZISP" country="Thailand" location="Chiang Mai" pageId="4" pageNumber="281" specimenCount="19" specimenCount-female="19" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus id="F60B2DD7FE53BD36C276041A436FFA47" box="[1280,1392,1392,1416]" pageId="4" pageNumber="281" type="paratype">paratypes</typeStatus>
:
<specimenCount id="3FB658FCFE53BD36C2F6041A46EEFA63" pageId="4" pageNumber="281" type="female">19 females</specimenCount>
, same data
</materialsCitation>
,
<materialsCitation id="99D89928FE53BD36C60F04FE4258FA63" ID-GBIF-Occurrence="1852084070" box="[377,1095,1427,1452]" collectingDate="1992-07" collectorName="N. Kagana." country="Thailand" location="Mae Ai" pageId="4" pageNumber="281" specimenCount="4" specimenCount-female="4" stateProvince="Chiang Mai" typeStatus="paratype">
<specimenCount id="3FB658FCFE53BD36C60F04FE47F7FA63" box="[377,488,1427,1452]" pageId="4" pageNumber="281" type="female">4 females</specimenCount>
,
<collectingCountry id="51A7D3E5FE53BD36C68504F94446FA63" box="[499,601,1427,1452]" name="Thailand" pageId="4" pageNumber="281">Thailand</collectingCountry>
,
<collectingRegion id="EB745D97FE53BD36C51304F944F2FA63" box="[613,749,1427,1452]" country="Thailand" name="Chiang Mai" pageId="4" pageNumber="281">Chiang Mai</collectingRegion>
,
<location id="2C6FC5AEFE53BD36C58E04FE4550FA63" LSID="urn:lsid:plazi:treatment:A1192263FE56BD36C7E104F3437CF81A:2C6FC5AEFE53BD36C58E04FE4550FA63" box="[760,847,1427,1452]" country="Thailand" name="Mae Ai" pageId="4" pageNumber="281" stateProvince="Chiang Mai">Mae Ai</location>
,
<collectingDate id="4D4A4C5DFE53BD36C42C04FE45DCFA63" box="[858,963,1428,1452]" pageId="4" pageNumber="281" value="1992-07">
<date id="5D0EB5B5FE53BD36C42C04FE45DCFA63" box="[858,963,1428,1452]" pageId="4" pageNumber="281" value="1992-07">VII.1992</date>
</collectingDate>
,
<collectorName id="8445F6A3FE53BD36C4B804FE4258FA63" box="[974,1095,1428,1452]" pageId="4" pageNumber="281">N.Kagana.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="290F9375FE53BD36C7B104DD445AF9D7" blockId="4.[151,1436,1391,2006]" pageId="4" pageNumber="281">
<materialsCitation id="99D89928FE53BD36C7B104DD441FFA3B" ID-GBIF-Occurrence="1852084154" collectionCode="RAS" country="Russia" location="Saint Petersburg" pageId="4" pageNumber="281" specimenCount="3" stateProvince="Saint Petersburg" typeStatus="holotype">
<emphasis id="1BC44F67FE53BD36C7B104DD4799FA1F" bold="true" box="[199,390,1463,1488]" pageId="4" pageNumber="281">
<typeStatus id="F60B2DD7FE53BD36C7B104DD471DFA1F" box="[199,258,1463,1488]" pageId="4" pageNumber="281">Type</typeStatus>
deposition
</emphasis>
. The
<typeStatus id="F60B2DD7FE53BD36C6BA04DD4431FA1F" box="[460,558,1463,1488]" pageId="4" pageNumber="281" type="holotype">holotype</typeStatus>
and four
<typeStatus id="F60B2DD7FE53BD36C5D404D24512FA1F" box="[674,781,1464,1488]" pageId="4" pageNumber="281" type="paratype">paratypes</typeStatus>
are deposited in the collection of Zoological Institute of
<collectionCode id="4FA10BB0FE53BD36C7E104B646CFFA3B" box="[151,208,1500,1524]" country="Myanmar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14256" name="Union of Burma Applied Research Institute" pageId="4" pageNumber="281">RAS</collectionCode>
,
<collectingRegion id="EB745D97FE53BD36C79704B147BEFA3B" box="[225,417,1499,1524]" country="Russia" name="Saint Petersburg" pageId="4" pageNumber="281">Saint Petersburg</collectingRegion>
,
<collectingCountry id="51A7D3E5FE53BD36C6C404B1441FFA3B" box="[434,512,1499,1524]" name="Russia" pageId="4" pageNumber="281">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="99D89928FE53BD36C56404B6445AF9D7" ID-GBIF-Occurrence="1852084071" country="Russia" location="Tyumen State University Museum of Zoology" pageId="4" pageNumber="281" specimenCount="19" stateProvince="Tyumen" typeStatus="paratype">
<specimenCount id="3FB658FCFE53BD36C56404B644B9FA3B" box="[530,678,1500,1524]" pageId="4" pageNumber="281" type="generic" typeStatus="paratypes">19 paratypes</specimenCount>
are deposited in the collection of the
<location id="2C6FC5AEFE53BD36C30F04B64761F9D7" LSID="urn:lsid:plazi:treatment:A1192263FE56BD36C7E104F3437CF81A:2C6FC5AEFE53BD36C30F04B64761F9D7" country="Russia" name="Tyumen State University Museum of Zoology" pageId="4" pageNumber="281" stateProvince="Tyumen">Tyumen State University Museum of Zoology</location>
,
<collectingRegion id="EB745D97FE53BD36C6FF076A47F8F9D7" box="[393,487,1536,1560]" country="Russia" name="Tyumen'" pageId="4" pageNumber="281">Tyumen</collectingRegion>
,
<collectingCountry id="51A7D3E5FE53BD36C6850495445EF9D7" box="[499,577,1535,1560]" name="Russia" pageId="4" pageNumber="281">Russia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE53BD36C7B10749476DF87D" pageId="4" pageNumber="281" type="diagnosis">
<paragraph id="290F9375FE53BD36C7B10749476DF87D" blockId="4.[151,1436,1391,2006]" pageId="4" pageNumber="281">
<emphasis id="1BC44F67FE53BD36C7B1074947D7F9F3" bold="true" box="[199,456,1571,1596]" pageId="4" pageNumber="281">Differential diagnosis</emphasis>
. The new species is most similar to
<emphasis id="1BC44F67FE53BD36C418074E4563F9F4" box="[878,892,1572,1595]" italics="true" pageId="4" pageNumber="281">S</emphasis>
.
<emphasis id="1BC44F67FE53BD36C4FA0749421EF9F3" box="[908,1025,1571,1596]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
<bibRefCitation id="4D21EE84FE53BD36C37F0749433BF9F3" box="[1033,1316,1571,1596]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych and Rack, 1993</bibRefCitation>
, described from
<collectingCountry id="51A7D3E5FE53BD36C7AC07224757F990" box="[218,328,1608,1631]" name="Botswana" pageId="4" pageNumber="281">Botswana</collectingCountry>
(
<bibRefCitation id="4D21EE84FE53BD36C62A072D4476F9AF" box="[348,617,1607,1632]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych &amp; Rack 1993</bibRefCitation>
), by the similar modifications of setae on tibiotarsus I and tarsus II, presence of apodemes 5, and characteristic position of setae
<emphasis id="1BC44F67FE53BD36C44907064547F94C" box="[831,856,1644,1667]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
3 anteriorly to
<emphasis id="1BC44F67FE53BD36C48F0706420DF94C" box="[1017,1042,1644,1667]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
1, 2. The new species differs from
<emphasis id="1BC44F67FE53BD36C2F10706438AF94C" box="[1415,1429,1644,1667]" italics="true" pageId="4" pageNumber="281">S</emphasis>
.
<emphasis id="1BC44F67FE53BD36C7E107F84713F964" box="[151,268,1682,1707]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
by distinctly barbed setae
<emphasis id="1BC44F67FE53BD36C54907F94447F965" box="[575,600,1683,1706]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
1, 2 (vs. smooth in
<emphasis id="1BC44F67FE53BD36C45707F94530F965" box="[801,815,1683,1706]" italics="true" pageId="4" pageNumber="281">S</emphasis>
.
<emphasis id="1BC44F67FE53BD36C44807F845ADF964" box="[830,946,1682,1707]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
), foliate setae on tarsi III and IV (vs. nonmodified in
<emphasis id="1BC44F67FE53BD36C66907D047B0F91D" box="[287,431,1721,1746]" italics="true" pageId="4" pageNumber="281">S. сamerikae</emphasis>
) and by distinctly longer dorsal setae
<emphasis id="1BC44F67FE53BD36C42E07D34579F91D" box="[856,870,1721,1746]" italics="true" pageId="4" pageNumber="281">d</emphasis>
39–48,
<emphasis id="1BC44F67FE53BD36C4B607D345D7F91E" box="[960,968,1721,1745]" italics="true" pageId="4" pageNumber="281">f</emphasis>
44–58,
<emphasis id="1BC44F67FE53BD36C35507D3422EF91D" box="[1059,1073,1721,1746]" italics="true" pageId="4" pageNumber="281">h</emphasis>
1 45–57 (vs.
<emphasis id="1BC44F67FE53BD36C3CA07D342D5F91D" box="[1212,1226,1721,1746]" italics="true" pageId="4" pageNumber="281">d</emphasis>
25–30,
<emphasis id="1BC44F67FE53BD36C25207D34333F91E" box="[1316,1324,1721,1745]" italics="true" pageId="4" pageNumber="281">f</emphasis>
24–31,
<emphasis id="1BC44F67FE53BD36C2F007D3438BF91D" box="[1414,1428,1721,1746]" italics="true" pageId="4" pageNumber="281">h</emphasis>
<date id="5D0EB5B5FE53BD36C2E207A246C2F937" pageId="4" pageNumber="281" value="1924-01-31">
1
<quantity id="EE483E90FE53BD36C7E1078A46C2F937" box="[151,221,1760,1784]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="7.874" metricValueMin="6.096" pageId="4" pageNumber="281" unit="in" value="27.5" valueMax="31.0" valueMin="24.0">24–31</quantity>
</date>
in
<emphasis id="1BC44F67FE53BD36C677078A4710F938" box="[257,271,1760,1783]" italics="true" pageId="4" pageNumber="281">S</emphasis>
.
<emphasis id="1BC44F67FE53BD36C66B07B5478EF937" box="[285,401,1759,1784]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
). The new species also similar to S.
<taxonomicName id="EEB0E8F6FE53BD36C45A07B5429DF937" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[812,1154,1759,1784]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="1BC44F67FE53BD36C45A07B54579F937" box="[812,870,1759,1784]" italics="true" pageId="4" pageNumber="281">langi</emphasis>
<bibRefCitation id="4D21EE84FE53BD36C41B07B5429DF937" box="[877,1154,1759,1784]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych and Rack, 1993</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="51A7D3E5FE53BD36C24B07B54383F937" box="[1341,1436,1759,1784]" name="Vietnam" pageId="4" pageNumber="281">Vietnam</collectingCountry>
(
<bibRefCitation id="4D21EE84FE53BD36C7E8066E47B7F8D3" box="[158,424,1796,1821]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych &amp; Rack 1993</bibRefCitation>
), by the similar relative lengths of dorsal idiosomal setae and by the same number of modified foliate setae on tibiotarsus I and tarsi I and II. The new species differs from
<taxonomicName id="EEB0E8F6FE53BD36C30A064242CBF88F" authorityName="Dastych and Rack" authorityYear="1993" box="[1148,1236,1831,1856]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="1BC44F67FE53BD36C30A064242CBF88F" box="[1148,1236,1831,1856]" italics="true" pageId="4" pageNumber="281">S. langi</emphasis>
</taxonomicName>
by non-modified setae
<emphasis id="1BC44F67FE53BD36C7A0062746ECF8AB" box="[214,243,1869,1892]" italics="true" pageId="4" pageNumber="281">tс’</emphasis>
on tarsi II and III (vs. spiniform in
<taxonomicName id="EEB0E8F6FE53BD36C5FC062744FEF8AA" authorityName="Dastych and Rack" authorityYear="1993" box="[650,737,1868,1893]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="1BC44F67FE53BD36C5FC062744FEF8AA" box="[650,737,1868,1893]" italics="true" pageId="4" pageNumber="281">S. langi</emphasis>
</taxonomicName>
), by setae
<emphasis id="1BC44F67FE53BD36C42A0627456AF8AB" box="[860,885,1869,1892]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
1, 2 distinctly barbed and subequal to
<emphasis id="1BC44F67FE53BD36C26406274334F8AB" box="[1298,1323,1869,1892]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
3 (vs.
<emphasis id="1BC44F67FE53BD36C21A0627439AF8AB" box="[1388,1413,1869,1892]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
1, 2 smooth and distinctly shorter than
<emphasis id="1BC44F67FE53BD36C56E061E442EF844" box="[536,561,1908,1931]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
<quantity id="EE483E90FE53BD36C54706EB444AF844" box="[561,597,1906,1935]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="281" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="EEB0E8F6FE53BD36C52D061944AEF844" authorityName="Dastych and Rack" authorityYear="1993" box="[603,689,1906,1931]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="1BC44F67FE53BD36C52D061944AEF844" box="[603,689,1906,1931]" italics="true" pageId="4" pageNumber="281">S. langi</emphasis>
</taxonomicName>
), and by setae 2
<emphasis id="1BC44F67FE53BD36C413061E456CF844" box="[869,883,1908,1931]" italics="true" pageId="4" pageNumber="281">a</emphasis>
distinctly shorter than 2
<emphasis id="1BC44F67FE53BD36C3F7061E4292F844" box="[1153,1165,1908,1931]" italics="true" pageId="4" pageNumber="281">с</emphasis>
(vs. 2
<emphasis id="1BC44F67FE53BD36C3A7061E42C0F844" box="[1233,1247,1908,1931]" italics="true" pageId="4" pageNumber="281">a</emphasis>
distinctly longer than 2
<emphasis id="1BC44F67FE53BD36C7AA06F046F7F87E" box="[220,232,1946,1969]" italics="true" pageId="4" pageNumber="281">с</emphasis>
in
<taxonomicName id="EEB0E8F6FE53BD36C67A06F0477EF87D" authorityName="Dastych and Rack" authorityYear="1993" box="[268,353,1945,1970]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="1BC44F67FE53BD36C67A06F04705F87E" box="[268,282,1946,1969]" italics="true" pageId="4" pageNumber="281">S</emphasis>
.
<emphasis id="1BC44F67FE53BD36C65E06F3477EF87D" box="[296,353,1945,1970]" italics="true" pageId="4" pageNumber="281">langi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE53BD36C7B106D7437CF81A" box="[199,1379,1981,2006]" pageId="4" pageNumber="281" type="etymology">
<paragraph id="290F9375FE53BD36C7B106D7437CF81A" blockId="4.[151,1436,1391,2006]" box="[199,1379,1981,2006]" pageId="4" pageNumber="281">
<emphasis id="1BC44F67FE53BD36C7B106D74759F819" bold="true" box="[199,326,1981,2006]" pageId="4" pageNumber="281">Etymology</emphasis>
. The name of the new species refers to its association with beetles of the genus
<taxonomicName id="EEB0E8F6FE53BD36C3B306D74343F819" authorityName="Lucas" authorityYear="1869" box="[1221,1372,1981,2006]" class="Insecta" family="Geotrupidae" genus="Enoplotrupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="1BC44F67FE53BD36C3B306D74343F819" box="[1221,1372,1981,2006]" italics="true" pageId="4" pageNumber="281">Enoplotrupes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5CF1C9495940F3E643C2CFB071D3BCA5">New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand</mods:title>
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<treatment id="A1192263FE5FBD38C7E1004D42BBFD67" ID-DOI="http://doi.org/10.5281/zenodo.5949613" ID-GBIF-Taxon="146044083" ID-Zenodo-Dep="5949613" LSID="urn:lsid:plazi:treatment:A1192263FE5FBD38C7E1004D42BBFD67" httpUri="http://treatment.plazi.org/id/A1192263FE5FBD38C7E1004D42BBFD67" lastPageId="10" lastPageNumber="287" pageId="8" pageNumber="285">
<subSubSection id="61AAC0FEFE5FBD3AC7E1004D473FFEAC" pageId="8" pageNumber="285" type="nomenclature">
<paragraph id="290F9375FE5FBD3AC7E1004D4416FE8D" blockId="8.[151,521,295,355]" box="[151,521,295,322]" pageId="8" pageNumber="285">
<heading id="72472419FE5FBD3AC7E1004D4416FE8D" bold="true" box="[151,521,295,322]" fontSize="11" level="1" pageId="8" pageNumber="285" reason="1">
<emphasis id="1BC44F67FE5FBD3AC7E1004D4416FE8D" bold="true" box="[151,521,295,322]" pageId="8" pageNumber="285">
<taxonomicName id="EEB0E8F6FE5FBD3AC7E1004D47BBFE8E" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[151,420,295,321]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="1BC44F67FE5FBD3AC7E1004D47BBFE8E" bold="true" box="[151,420,295,321]" italics="true" pageId="8" pageNumber="285">Nasutidania orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE5FBD3AC6DD00424416FE8D" box="[427,521,296,322]" pageId="8" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="290F9375FE5FBD3AC7E10020473FFEAC" blockId="8.[151,521,295,355]" box="[151,288,330,355]" pageId="8" pageNumber="285">
(
<figureCitation id="B18B8FF0FE5FBD3AC7E900204708FEAC" box="[159,279,330,355]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="8.[151,250,1961,1983]" captionStartId-1="9.[151,250,1405,1427]" captionStartId-2="10.[151,250,1947,1969]" captionStartId-3="11.[151,250,1978,2000]" captionStartId-4="12.[151,250,1913,1935]" captionTargetBox-0="[170,1408,631,1907]" captionTargetBox-1="[238,1358,247,1311]" captionTargetBox-2="[180,1407,792,1873]" captionTargetBox-3="[182,1395,312,1859]" captionTargetBox-4="[151,1436,240,1841]" captionTargetId-0="figure@8.[151,1436,605,1929]" captionTargetId-1="figure@9.[192,1395,220,1348]" captionTargetId-2="figure@10.[167,1419,760,1908]" captionTargetId-3="figure@11.[151,1436,255,1883]" captionTargetId-4="figure@12.[151,1436,240,1841]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 6. Nasutidania orientalis sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 7. Nasutidania orientalis sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view." captionText-2="FIGURE 8. Nasutidania orientalis sp. nov., female: A—right leg I in dorsal view, B—pretarsus and tarsal claw I in lateral view, C—right leg II in dorsal view." captionText-3="FIGURE 9. Nasutidania orientalis sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." captionText-4="FIGURE 10. DIC micrographs of Nasutidania orientalis sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri-0="https://zenodo.org/record/1303137/files/figure.png" httpUri-1="https://zenodo.org/record/1303139/files/figure.png" httpUri-2="https://zenodo.org/record/1303141/files/figure.png" httpUri-3="https://zenodo.org/record/1303143/files/figure.png" httpUri-4="https://zenodo.org/record/1303147/files/figure.png" pageId="8" pageNumber="285">Figs. 6–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE5FBD38C7E100F8443BFE1F" lastPageId="10" lastPageNumber="287" pageId="8" pageNumber="285" type="description">
<paragraph id="290F9375FE5FBD3AC7E100F842A0FE64" blockId="8.[151,1436,402,571]" box="[151,1215,402,427]" pageId="8" pageNumber="285">
<emphasis id="1BC44F67FE5FBD3AC7E100F8473EFE64" bold="true" box="[151,289,402,427]" pageId="8" pageNumber="285">Description</emphasis>
. FEMALE (
<figureCitation id="B18B8FF0FE5FBD3AC6D800F8443CFE64" box="[430,547,402,427]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="8.[151,250,1961,1983]" captionStartId-1="9.[151,250,1405,1427]" captionStartId-2="10.[151,250,1947,1969]" captionStartId-3="11.[151,250,1978,2000]" captionStartId-4="12.[151,250,1913,1935]" captionTargetBox-0="[170,1408,631,1907]" captionTargetBox-1="[238,1358,247,1311]" captionTargetBox-2="[180,1407,792,1873]" captionTargetBox-3="[182,1395,312,1859]" captionTargetBox-4="[151,1436,240,1841]" captionTargetId-0="figure@8.[151,1436,605,1929]" captionTargetId-1="figure@9.[192,1395,220,1348]" captionTargetId-2="figure@10.[167,1419,760,1908]" captionTargetId-3="figure@11.[151,1436,255,1883]" captionTargetId-4="figure@12.[151,1436,240,1841]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 6. Nasutidania orientalis sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 7. Nasutidania orientalis sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view." captionText-2="FIGURE 8. Nasutidania orientalis sp. nov., female: A—right leg I in dorsal view, B—pretarsus and tarsal claw I in lateral view, C—right leg II in dorsal view." captionText-3="FIGURE 9. Nasutidania orientalis sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." captionText-4="FIGURE 10. DIC micrographs of Nasutidania orientalis sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri-0="https://zenodo.org/record/1303137/files/figure.png" httpUri-1="https://zenodo.org/record/1303139/files/figure.png" httpUri-2="https://zenodo.org/record/1303141/files/figure.png" httpUri-3="https://zenodo.org/record/1303143/files/figure.png" httpUri-4="https://zenodo.org/record/1303147/files/figure.png" pageId="8" pageNumber="285">Figs 6–10</figureCitation>
). Length of idiosoma 185 (190–220), width 105 (110–125).
</paragraph>
<paragraph id="290F9375FE5FBD3AC7B100DC45F9FDF4" blockId="8.[151,1436,402,571]" pageId="8" pageNumber="285">
<emphasis id="1BC44F67FE5FBD3AC7B100DC4749FE00" box="[199,342,438,463]" italics="true" pageId="8" pageNumber="285">Gnathosoma</emphasis>
(
<figureCitation id="B18B8FF0FE5FBD3AC61F00DC47ADFE00" box="[361,434,438,463]" captionStart="FIGURE 7" captionStartId="9.[151,250,1405,1427]" captionTargetBox="[238,1358,247,1311]" captionTargetId="figure@9.[192,1395,220,1348]" captionTargetPageId="9" captionText="FIGURE 7. Nasutidania orientalis sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view." httpUri="https://zenodo.org/record/1303139/files/figure.png" pageId="8" pageNumber="285">Fig. 7</figureCitation>
). Length of gnathosoma 57 (57–59), width 21 (21–23). Dorsal median apodeme well developed. Cheliceral setae smooth and pointed. Setae
<emphasis id="1BC44F67FE5FBD3AC45F00B0454DFE3C" box="[809,850,474,499]" italics="true" pageId="8" pageNumber="285">сha</emphasis>
8 (8–13) distinctly shorter than
<emphasis id="1BC44F67FE5FBD3AC3A800B04319FE3C" box="[1246,1286,474,499]" italics="true" pageId="8" pageNumber="285">сhb</emphasis>
14 (14–18). Postpalpal setae (
<emphasis id="1BC44F67FE5FBD3AC62B00954766FDD9" box="[349,377,511,534]" italics="true" pageId="8" pageNumber="285">pp</emphasis>
) short, needle-like, situated anterolaterally to bases of
<emphasis id="1BC44F67FE5FBD3AC49100944210FDD8" box="[999,1039,510,535]" italics="true" pageId="8" pageNumber="285">сha</emphasis>
. Setae
<emphasis id="1BC44F67FE5FBD3AC31500944290FDD8" box="[1123,1167,510,535]" italics="true" pageId="8" pageNumber="285">dFe</emphasis>
very short, needle-like; setae
<emphasis id="1BC44F67FE5FBD3AC7A00348471AFDF4" box="[214,261,546,571]" italics="true" pageId="8" pageNumber="285">dGe</emphasis>
, smooth and pointed. Subcapitular setae
<emphasis id="1BC44F67FE5FBD3AC5A7034E44FAFDF4" box="[721,741,548,571]" italics="true" pageId="8" pageNumber="285">m</emphasis>
9 smooth and pointed.
</paragraph>
<caption id="7DCFC3FDFE5FBD3AC7E106C34336F870" box="[151,1321,1961,1983]" httpUri="https://zenodo.org/record/1303137/files/figure.png" pageId="8" pageNumber="285" startId="8.[151,250,1961,1983]" targetBox="[170,1408,631,1907]" targetPageId="8">
<paragraph id="290F9375FE5FBD3AC7E106C34336F870" blockId="8.[151,1321,1961,1983]" box="[151,1321,1961,1983]" pageId="8" pageNumber="285">
<emphasis id="1BC44F67FE5FBD3AC7E106C34712F871" bold="true" box="[151,269,1961,1983]" pageId="8" pageNumber="285">FIGURE 6</emphasis>
.
<taxonomicName id="EEB0E8F6FE5FBD3AC66F06C347E3F870" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[281,508,1961,1983]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="1BC44F67FE5FBD3AC66F06C347E3F870" box="[281,508,1961,1983]" italics="true" pageId="8" pageNumber="285">Nasutidania orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE5FBD3AC57406C3444CF871" box="[514,595,1961,1982]" pageId="8" pageNumber="285" rank="species">
<emphasis id="1BC44F67FE5FBD3AC57406C3444CF871" bold="true" box="[514,595,1961,1982]" pageId="8" pageNumber="285">sp. nov.</emphasis>
</taxonomicNameLabel>
, female: A—dorsum of the body, B—venter of the body. Legs omitted.
</paragraph>
</caption>
<caption id="7DCFC3FDFE5EBD3BC7E10417435EFA5C" box="[151,1345,1405,1427]" httpUri="https://zenodo.org/record/1303139/files/figure.png" pageId="9" pageNumber="286" startId="9.[151,250,1405,1427]" targetBox="[238,1358,247,1311]" targetPageId="9">
<paragraph id="290F9375FE5EBD3BC7E10417435EFA5C" blockId="9.[151,1345,1405,1427]" box="[151,1345,1405,1427]" pageId="9" pageNumber="286">
<emphasis id="1BC44F67FE5EBD3BC7E104174712FA5C" bold="true" box="[151,269,1405,1427]" pageId="9" pageNumber="286">FIGURE 7</emphasis>
.
<taxonomicName id="EEB0E8F6FE5EBD3BC66F041747E3FA5C" authority="Khaustov &amp; Frolov, 2018" authorityName="Khaustov &amp; Frolov" authorityYear="2018" box="[281,508,1405,1427]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="1BC44F67FE5EBD3BC66F041747E3FA5C" box="[281,508,1405,1427]" italics="true" pageId="9" pageNumber="286">Nasutidania orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00F7F21CFE5EBD3BC5740414444CFA5C" box="[514,595,1406,1427]" pageId="9" pageNumber="286" rank="species">
<emphasis id="1BC44F67FE5EBD3BC5740414444CFA5C" bold="true" box="[514,595,1406,1427]" pageId="9" pageNumber="286">sp. nov.</emphasis>
</taxonomicNameLabel>
, female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view.
</paragraph>
</caption>
<paragraph id="290F9375FE5EBD3BC7B104D0438AF915" blockId="9.[151,1436,1466,2019]" pageId="9" pageNumber="286">
<emphasis id="1BC44F67FE5EBD3BC7B104D0478EFA1C" box="[199,401,1466,1491]" italics="true" pageId="9" pageNumber="286">Idiosomal dorsum</emphasis>
(
<figureCitation id="B18B8FF0FE5EBD3BC6D604D047E3FA1C" box="[416,508,1466,1491]" captionStart="FIGURE 6" captionStartId="8.[151,250,1961,1983]" captionTargetBox="[170,1408,631,1907]" captionTargetId="figure@8.[151,1436,605,1929]" captionTargetPageId="8" captionText="FIGURE 6. Nasutidania orientalis sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/1303137/files/figure.png" pageId="9" pageNumber="286">Figs 6A</figureCitation>
,
<figureCitation id="B18B8FF0FE5EBD3BC57E04D14482FA1C" box="[520,669,1467,1491]" captionStart="FIGURE 10" captionStartId="12.[151,250,1913,1935]" captionTargetBox="[151,1436,240,1841]" captionTargetId="figure@12.[151,1436,240,1841]" captionTargetPageId="12" captionText="FIGURE 10. DIC micrographs of Nasutidania orientalis sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303147/files/figure.png" pageId="9" pageNumber="286">10A, C, E, G</figureCitation>
). All dorsal shields with numerous very small dimples (
<figureCitation id="B18B8FF0FE5EBD3BC27A04D046CFFA38" captionStart="FIGURE 10" captionStartId="12.[151,250,1913,1935]" captionTargetBox="[151,1436,240,1841]" captionTargetId="figure@12.[151,1436,240,1841]" captionTargetPageId="12" captionText="FIGURE 10. DIC micrographs of Nasutidania orientalis sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303147/files/figure.png" pageId="9" pageNumber="286">Figs 10A, C, E, G</figureCitation>
). Setae
<date id="5D0EB5B5FE5EBD3BC65A048A475EFA33" box="[300,321,1504,1532]" pageId="9" pageNumber="286">
<emphasis id="1BC44F67FE5EBD3BC65A048A4727FA38" box="[300,312,1504,1527]" italics="true" pageId="9" pageNumber="286">v</emphasis>
2
</date>
smooth, other dorsal setae barbed. Setae
<date id="5D0EB5B5FE5EBD3BC454048A4529FA33" box="[802,822,1504,1532]" pageId="9" pageNumber="286">
<emphasis id="1BC44F67FE5EBD3BC454048A4531FA38" box="[802,814,1504,1527]" italics="true" pageId="9" pageNumber="286">v</emphasis>
2
</date>
,
<emphasis id="1BC44F67FE5EBD3BC430048A4542FA38" box="[838,861,1504,1527]" italics="true" pageId="9" pageNumber="286">sс</emphasis>
2,
<emphasis id="1BC44F67FE5EBD3BC400048A459DFA38" box="[886,898,1504,1527]" italics="true" pageId="9" pageNumber="286">с</emphasis>
2, and
<emphasis id="1BC44F67FE5EBD3BC4BA04B445C5FA38" box="[972,986,1502,1527]" italics="true" pageId="9" pageNumber="286">h</emphasis>
2 pointed, other dorsal setae moderately blunt-ended. Trichobothria with long stem, clavate, sparsely barbed, with pointed apex. Tergite C with two pairs of porous areas, one of which located between setae
<emphasis id="1BC44F67FE5EBD3BC5BC074144C9F98D" box="[714,726,1579,1602]" italics="true" pageId="9" pageNumber="286">с</emphasis>
1 and
<emphasis id="1BC44F67FE5EBD3BC4600741453DF98D" box="[790,802,1579,1602]" italics="true" pageId="9" pageNumber="286">с</emphasis>
2, and another pair located anteromedially to bases of
<emphasis id="1BC44F67FE5EBD3BC2F607414393F98D" box="[1408,1420,1579,1602]" italics="true" pageId="9" pageNumber="286">с</emphasis>
1; tergite H with a pair of round porous areas in anterior part covered by tergite EF. Posterior margin of tergite H evenly rounded. Lengths of dorsal setae:
<date id="5D0EB5B5FE5EBD3BC514071F4469F95E" box="[610,630,1653,1681]" pageId="9" pageNumber="286">
<emphasis id="1BC44F67FE5EBD3BC514071F4471F943" box="[610,622,1653,1676]" italics="true" pageId="9" pageNumber="286">v</emphasis>
2
</date>
8 (7–8),
<emphasis id="1BC44F67FE5EBD3BC5AA071F44ECF943" box="[732,755,1653,1676]" italics="true" pageId="9" pageNumber="286">sс</emphasis>
2 48 (51–53),
<emphasis id="1BC44F67FE5EBD3BC4FD071F4588F943" box="[907,919,1653,1676]" italics="true" pageId="9" pageNumber="286">с</emphasis>
1 46 (45–49),
<emphasis id="1BC44F67FE5EBD3BC35D071F4228F943" box="[1067,1079,1653,1676]" italics="true" pageId="9" pageNumber="286">с</emphasis>
2 58 (55–59),
<emphasis id="1BC44F67FE5EBD3BC3B9071E42C2F942" box="[1231,1245,1652,1677]" italics="true" pageId="9" pageNumber="286">d</emphasis>
43 (45–47),
<emphasis id="1BC44F67FE5EBD3BC21B071F4366F943" box="[1389,1401,1653,1676]" italics="true" pageId="9" pageNumber="286">e</emphasis>
24 (24–26),
<emphasis id="1BC44F67FE5EBD3BC78B07F6471AF97B" box="[253,261,1692,1716]" italics="true" pageId="9" pageNumber="286">f</emphasis>
58 (60–62),
<emphasis id="1BC44F67FE5EBD3BC6E307F147BCF97B" box="[405,419,1691,1716]" italics="true" pageId="9" pageNumber="286">h</emphasis>
1 58 (60–61),
<emphasis id="1BC44F67FE5EBD3BC54A07F14455F97B" box="[572,586,1691,1716]" italics="true" pageId="9" pageNumber="286">h</emphasis>
2 57 (57–60). Distances between setae:
<date id="5D0EB5B5FE5EBD3BC37607F64246F97B" box="[1024,1113,1691,1720]" pageId="9" pageNumber="286" value="1926-05-02" valueMax="1926-05-02" valueMin="1926-05-02">
<emphasis id="1BC44F67FE5EBD3BC37607F64213F97C" box="[1024,1036,1692,1715]" italics="true" pageId="9" pageNumber="286">v</emphasis>
2
<emphasis id="1BC44F67FE5EBD3BC36207F64231F97C" box="[1044,1070,1692,1715]" italics="true" pageId="9" pageNumber="286">–v</emphasis>
2 26
</date>
(26–28),
<emphasis id="1BC44F67FE5EBD3BC3B007F642C2F97C" box="[1222,1245,1692,1715]" italics="true" pageId="9" pageNumber="286">sс</emphasis>
2
<emphasis id="1BC44F67FE5EBD3BC39307F64315F97C" box="[1253,1290,1692,1715]" italics="true" pageId="9" pageNumber="286">–sс</emphasis>
2 30 (28–31),
<emphasis id="1BC44F67FE5EBD3BC7E107A946BCF915" box="[151,163,1731,1754]" italics="true" pageId="9" pageNumber="286">с</emphasis>
1–
<emphasis id="1BC44F67FE5EBD3BC7CF07A946DAF915" box="[185,197,1731,1754]" italics="true" pageId="9" pageNumber="286">с</emphasis>
1 50 (49–52),
<emphasis id="1BC44F67FE5EBD3BC62807A94775F915" box="[350,362,1731,1754]" italics="true" pageId="9" pageNumber="286">с</emphasis>
1–
<emphasis id="1BC44F67FE5EBD3BC6F607A94793F915" box="[384,396,1731,1754]" italics="true" pageId="9" pageNumber="286">с</emphasis>
2 19 (19–21),
<emphasis id="1BC44F67FE5EBD3BC55207A84451F914" box="[548,590,1730,1755]" italics="true" pageId="9" pageNumber="286">d–d</emphasis>
48 (50–52),
<emphasis id="1BC44F67FE5EBD3BC5A907A9451DF915" box="[735,770,1730,1754]" italics="true" pageId="9" pageNumber="286">e–f</emphasis>
9 (9–10),
<emphasis id="1BC44F67FE5EBD3BC40007A8458BF915" box="[886,916,1730,1754]" italics="true" pageId="9" pageNumber="286">f–f</emphasis>
48 (51–53),
<emphasis id="1BC44F67FE5EBD3BC35207A8422DF914" box="[1060,1074,1730,1755]" italics="true" pageId="9" pageNumber="286">h</emphasis>
1–
<emphasis id="1BC44F67FE5EBD3BC33E07A84249F914" box="[1096,1110,1730,1755]" italics="true" pageId="9" pageNumber="286">h</emphasis>
1 29 (30–31),
<emphasis id="1BC44F67FE5EBD3BC39807A842E3F914" box="[1262,1276,1730,1755]" italics="true" pageId="9" pageNumber="286">h</emphasis>
1–
<emphasis id="1BC44F67FE5EBD3BC26407A8433FF914" box="[1298,1312,1730,1755]" italics="true" pageId="9" pageNumber="286">h</emphasis>
2 9 (9–10).
</paragraph>
<paragraph id="290F9375FE5EBD3BC7B1078245DFF85E" blockId="9.[151,1436,1466,2019]" pageId="9" pageNumber="286">
<emphasis id="1BC44F67FE5EBD3BC7B10782479BF8CE" box="[199,388,1768,1793]" italics="true" pageId="9" pageNumber="286">Idiosomal venter</emphasis>
(
<figureCitation id="B18B8FF0FE5EBD3BC6E5078247F0F8CE" box="[403,495,1768,1793]" captionStart="FIGURE 6" captionStartId="8.[151,250,1961,1983]" captionTargetBox="[170,1408,631,1907]" captionTargetId="figure@8.[151,1436,605,1929]" captionTargetPageId="8" captionText="FIGURE 6. Nasutidania orientalis sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/1303137/files/figure.png" pageId="9" pageNumber="286">Figs 6B</figureCitation>
,
<figureCitation id="B18B8FF0FE5EBD3BC68A078344BBF8CE" box="[508,676,1769,1793]" captionStart="FIGURE 10" captionStartId="12.[151,250,1913,1935]" captionTargetBox="[151,1436,240,1841]" captionTargetId="figure@12.[151,1436,240,1841]" captionTargetPageId="12" captionText="FIGURE 10. DIC micrographs of Nasutidania orientalis sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303147/files/figure.png" pageId="9" pageNumber="286">10B, D, F, H, I</figureCitation>
). Ventral plates with numerous very small dimples (
<figureCitation id="B18B8FF0FE5EBD3BC399078246B0F8EA" captionStart="FIGURE 10" captionStartId="12.[151,250,1913,1935]" captionTargetBox="[151,1436,240,1841]" captionTargetId="figure@12.[151,1436,240,1841]" captionTargetPageId="12" captionText="FIGURE 10. DIC micrographs of Nasutidania orientalis sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303147/files/figure.png" pageId="9" pageNumber="286">Figs 10B, D, F, H</figureCitation>
). All ventral setae pointed. Setae 1
<emphasis id="1BC44F67FE5EBD3BC5440664445FF8EA" box="[562,576,1806,1829]" italics="true" pageId="9" pageNumber="286">a</emphasis>
, 1
<emphasis id="1BC44F67FE5EBD3BC52A06664475F8EA" box="[604,618,1804,1829]" italics="true" pageId="9" pageNumber="286">b</emphasis>
, and 2
<emphasis id="1BC44F67FE5EBD3BC5C2066444DDF8EA" box="[692,706,1806,1829]" italics="true" pageId="9" pageNumber="286">a</emphasis>
barbed, other ventral setae smooth. Posterior margin of aggenital plate straight. Mgs very large, round, weaker sclerotized than ags and pgs (
<figureCitation id="B18B8FF0FE5EBD3BC4A0065A422DF886" box="[982,1074,1840,1865]" captionStart="FIGURE 10" captionStartId="12.[151,250,1913,1935]" captionTargetBox="[151,1436,240,1841]" captionTargetId="figure@12.[151,1436,240,1841]" captionTargetPageId="12" captionText="FIGURE 10. DIC micrographs of Nasutidania orientalis sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303147/files/figure.png" pageId="9" pageNumber="286">
Fig.
<date id="5D0EB5B5FE5EBD3BC37F065B422DF886" box="[1033,1074,1841,1865]" pageId="9" pageNumber="286" value="2010-01">10I</date>
</figureCitation>
). Lengths of ventral setae: 1
<emphasis id="1BC44F67FE5EBD3BC21A06584365F886" box="[1388,1402,1842,1865]" italics="true" pageId="9" pageNumber="286">a</emphasis>
24 (27–29), 1
<emphasis id="1BC44F67FE5EBD3BC666063F4701F8A1" box="[272,286,1877,1902]" italics="true" pageId="9" pageNumber="286">b</emphasis>
23 (27–29), 2
<emphasis id="1BC44F67FE5EBD3BC6BF063C47C8F8A2" box="[457,471,1878,1901]" italics="true" pageId="9" pageNumber="286">a</emphasis>
26 (30–31), 2
<emphasis id="1BC44F67FE5EBD3BC5F5063F448EF8A1" box="[643,657,1877,1902]" italics="true" pageId="9" pageNumber="286">b</emphasis>
17 (19–21), 3
<emphasis id="1BC44F67FE5EBD3BC44A063C4555F8A2" box="[828,842,1878,1901]" italics="true" pageId="9" pageNumber="286">a</emphasis>
26 (20–22), 3
<emphasis id="1BC44F67FE5EBD3BC483063F421CF8A1" box="[1013,1027,1877,1902]" italics="true" pageId="9" pageNumber="286">b</emphasis>
23 (22–25), 3
<emphasis id="1BC44F67FE5EBD3BC3D9063C42A4F8A2" box="[1199,1211,1878,1901]" italics="true" pageId="9" pageNumber="286">с</emphasis>
20 (20–22), 4
<emphasis id="1BC44F67FE5EBD3BC211063C436AF8A2" box="[1383,1397,1878,1901]" italics="true" pageId="9" pageNumber="286">a</emphasis>
17 (19–22), 4
<emphasis id="1BC44F67FE5EBD3BC67A06124705F85E" box="[268,282,1912,1937]" italics="true" pageId="9" pageNumber="286">b</emphasis>
29 (30–33), 4
<emphasis id="1BC44F67FE5EBD3BC6CE061047DBF85E" box="[440,452,1914,1937]" italics="true" pageId="9" pageNumber="286">с</emphasis>
22 (23–26),
<emphasis id="1BC44F67FE5EBD3BC52306104471F85E" box="[597,622,1914,1937]" italics="true" pageId="9" pageNumber="286">ps</emphasis>
1, 2 26 (25–29),
<emphasis id="1BC44F67FE5EBD3BC46006104530F85E" box="[790,815,1914,1937]" italics="true" pageId="9" pageNumber="286">ps</emphasis>
3 19 (21–22).
</paragraph>
<paragraph id="290F9375FE5EBD38C7B106CA45B8FE62" blockId="9.[151,1436,1466,2019]" lastBlockId="10.[151,1437,151,680]" lastPageId="10" lastPageNumber="287" pageId="9" pageNumber="286">
<emphasis id="1BC44F67FE5EBD3BC7B106CA46E3F878" box="[199,252,1952,1975]" italics="true" pageId="9" pageNumber="286">Legs</emphasis>
(
<figureCitation id="B18B8FF0FE5EBD3BC67A06F54743F877" box="[268,348,1951,1976]" captionStart="FIGURE 8" captionStartId="10.[151,250,1947,1969]" captionTargetBox="[180,1407,792,1873]" captionTargetId="figure@10.[167,1419,760,1908]" captionTargetPageId="10" captionText="FIGURE 8. Nasutidania orientalis sp. nov., female: A—right leg I in dorsal view, B—pretarsus and tarsal claw I in lateral view, C—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303141/files/figure.png" pageId="9" pageNumber="286">Figs. 8</figureCitation>
,
<figureCitation id="B18B8FF0FE5EBD3BC61C06CA4765F877" box="[362,378,1952,1976]" captionStart="FIGURE 9" captionStartId="11.[151,250,1978,2000]" captionTargetBox="[182,1395,312,1859]" captionTargetId="figure@11.[151,1436,255,1883]" captionTargetPageId="11" captionText="FIGURE 9. Nasutidania orientalis sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303143/files/figure.png" pageId="9" pageNumber="286">9</figureCitation>
). Leg I (
<figureCitation id="B18B8FF0FE5EBD3BC6A806F54426F877" box="[478,569,1951,1976]" captionStart="FIGURE 8" captionStartId="10.[151,250,1947,1969]" captionTargetBox="[180,1407,792,1873]" captionTargetId="figure@10.[167,1419,760,1908]" captionTargetPageId="10" captionText="FIGURE 8. Nasutidania orientalis sp. nov., female: A—right leg I in dorsal view, B—pretarsus and tarsal claw I in lateral view, C—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303141/files/figure.png" pageId="9" pageNumber="286">Fig. 8A</figureCitation>
). Lengths of solenidia
<emphasis id="1BC44F67FE5EBD3BC44B06CA454EF878" box="[829,849,1952,1975]" italics="true" pageId="9" pageNumber="286">ω</emphasis>
1 9 (9–10),
<emphasis id="1BC44F67FE5EBD3BC4A706CA45FAF878" box="[977,997,1952,1975]" italics="true" pageId="9" pageNumber="286">ω</emphasis>
2 4 (3–4),
<emphasis id="1BC44F67FE5EBD3BC32006CA427AF878" box="[1110,1125,1952,1975]" italics="true" pageId="9" pageNumber="286">φ</emphasis>
1 9 (9),
<emphasis id="1BC44F67FE5EBD3BC3CA06CA42D4F878" box="[1212,1227,1952,1975]" italics="true" pageId="9" pageNumber="286">φ</emphasis>
2 4 (4–5);
<emphasis id="1BC44F67FE5EBD3BC24B06CA4353F878" box="[1341,1356,1952,1975]" italics="true" pageId="9" pageNumber="286">φ</emphasis>
1 thick, clavate,
<emphasis id="1BC44F67FE5EBD3BC78306AD4716F811" box="[245,265,1991,2014]" italics="true" pageId="9" pageNumber="286">ω</emphasis>
1 finger-shaped,
<emphasis id="1BC44F67FE5EBD3BC6B406AD47C9F811" box="[450,470,1991,2014]" italics="true" pageId="9" pageNumber="286">ω</emphasis>
2 and
<emphasis id="1BC44F67FE5EBD3BC56106AD4439F811" box="[535,550,1991,2014]" italics="true" pageId="9" pageNumber="286">φ</emphasis>
2 weakly clavate;
<emphasis id="1BC44F67FE5EBD3BC59A06AD451FF811" box="[748,768,1991,2014]" italics="true" pageId="9" pageNumber="286">ω</emphasis>
2 not striated. Seta
<emphasis id="1BC44F67FE5EBD3BC4A006AC45F8F810" box="[982,999,1990,2015]" italics="true" pageId="9" pageNumber="286">l’</emphasis>
of femur short and blunt-ended; other leg setae (except eupathidia) pointed. Setae
<emphasis id="1BC44F67FE5DBD38C5FA01FD4481FF7F" box="[652,670,151,176]" italics="true" pageId="10" pageNumber="287">l’</emphasis>
of femur and
<emphasis id="1BC44F67FE5DBD38C43401FD4551FF7F" box="[834,846,151,176]" italics="true" pageId="10" pageNumber="287">k</emphasis>
smooth; other leg setae (except eupathidia) barbed. Leg II (
<figureCitation id="B18B8FF0FE5DBD38C79801D64758FF1B" box="[238,327,188,213]" captionStart="FIGURE 8" captionStartId="10.[151,250,1947,1969]" captionTargetBox="[180,1407,792,1873]" captionTargetId="figure@10.[167,1419,760,1908]" captionTargetPageId="10" captionText="FIGURE 8. Nasutidania orientalis sp. nov., female: A—right leg I in dorsal view, B—pretarsus and tarsal claw I in lateral view, C—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303141/files/figure.png" pageId="10" pageNumber="287">Fig. 8B</figureCitation>
). Tarsus with symmetrical padded claws and flipper-like empodium. Solenidia
<emphasis id="1BC44F67FE5DBD38C3BB01D742FEFF1B" box="[1229,1249,189,212]" italics="true" pageId="10" pageNumber="287">ω</emphasis>
7 (6–7) and
<emphasis id="1BC44F67FE5DBD38C20001D7439AFF1B" box="[1398,1413,189,212]" italics="true" pageId="10" pageNumber="287">φ</emphasis>
4 (4) weakly clavate. Seta
<emphasis id="1BC44F67FE5DBD38C6DF01B547A8FF37" box="[425,439,223,248]" italics="true" pageId="10" pageNumber="287">d</emphasis>
of femur blunt-ended; other leg setae pointed. Setae
<emphasis id="1BC44F67FE5DBD38C37201B5420AFF37" box="[1028,1045,223,248]" italics="true" pageId="10" pageNumber="287">l’</emphasis>
of femur and
<emphasis id="1BC44F67FE5DBD38C3C4018A42CAFF38" box="[1202,1237,224,247]" italics="true" pageId="10" pageNumber="287">tс”</emphasis>
of tarsus smooth; other leg setae barbed. Trochanter with short spine-like projection near base of seta
<emphasis id="1BC44F67FE5DBD38C34C006F424FFED3" box="[1082,1104,261,284]" italics="true" pageId="10" pageNumber="287">v’</emphasis>
. Leg III (
<figureCitation id="B18B8FF0FE5DBD38C3CC006E430BFED3" box="[1210,1300,260,285]" captionStart="FIGURE 9" captionStartId="11.[151,250,1978,2000]" captionTargetBox="[182,1395,312,1859]" captionTargetId="figure@11.[151,1436,255,1883]" captionTargetPageId="11" captionText="FIGURE 9. Nasutidania orientalis sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303143/files/figure.png" pageId="10" pageNumber="287">Fig. 9A</figureCitation>
). Claws and empodium as on tarsus II. Solenidion
<emphasis id="1BC44F67FE5DBD38C53A00434444FE8F" box="[588,603,297,320]" italics="true" pageId="10" pageNumber="287">φ</emphasis>
4 (4) weakly clavate. All leg setae barbed. Seta
<emphasis id="1BC44F67FE5DBD38C3FA004D4285FE8F" box="[1164,1178,295,320]" italics="true" pageId="10" pageNumber="287">d</emphasis>
of femur blunt-ended; other leg setae pointed. Leg IV (
<figureCitation id="B18B8FF0FE5DBD38C5620026446FFEAB" box="[532,624,332,357]" captionStart="FIGURE 9" captionStartId="11.[151,250,1978,2000]" captionTargetBox="[182,1395,312,1859]" captionTargetId="figure@11.[151,1436,255,1883]" captionTargetPageId="11" captionText="FIGURE 9. Nasutidania orientalis sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303143/files/figure.png" pageId="10" pageNumber="287">Fig. 9B</figureCitation>
). Tarsus with symmetrical simple hooked claws and small flipper-like empodium. Solenidion
<emphasis id="1BC44F67FE5DBD38C6E9001B47B1FE47" box="[415,430,369,392]" italics="true" pageId="10" pageNumber="287">φ</emphasis>
4 (4) weakly clavate. All leg setae barbed. Setae
<emphasis id="1BC44F67FE5DBD38C497000545F0FE47" box="[993,1007,367,392]" italics="true" pageId="10" pageNumber="287">d</emphasis>
of femur and
<emphasis id="1BC44F67FE5DBD38C3E7001A42B3FE48" box="[1169,1196,368,391]" italics="true" pageId="10" pageNumber="287">v”</emphasis>
of tibia blunt-ended; other leg setae pointed. Trochanter with short dorsal u-like projection.
</paragraph>
<paragraph id="290F9375FE5DBD38C7B100D2443BFE1F" blockId="10.[151,1437,151,680]" box="[199,548,439,464]" pageId="10" pageNumber="287">MALE and LARVA unknown.</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE5DBD38C7B100B6445AFD4A" pageId="10" pageNumber="287" type="materials_examined">
<paragraph id="290F9375FE5DBD38C7B100B6448FFDD7" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="287">
<materialsCitation id="99D89928FE5DBD38C7B100B64714FDD7" ID-GBIF-Occurrence="1852084152" collectingDate="1992-07" collectionCode="ZISP" collectorName="N. Kagana" country="Thailand" location="Mae Ai" pageId="10" pageNumber="287" specimenCount="2" stateProvince="Chiang Mai" typeStatus="holotype">
<emphasis id="1BC44F67FE5DBD38C7B100B64768FE3A" bold="true" box="[199,375,476,501]" pageId="10" pageNumber="287">
<typeStatus id="F60B2DD7FE5DBD38C7B100B6471DFE3A" box="[199,258,476,501]" pageId="10" pageNumber="287">Type</typeStatus>
material
</emphasis>
. Female
<typeStatus id="F60B2DD7FE5DBD38C68700B64449FE3A" box="[497,598,476,501]" pageId="10" pageNumber="287" type="holotype">holotype</typeStatus>
, slide
<collectionCode id="4FA10BB0FE5DBD38C5C600B644F5FE3B" box="[688,746,476,500]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="287">ZISP</collectionCode>
T-Neop-001,
<collectingCountry id="51A7D3E5FE5DBD38C4EF00B645E0FE3A" box="[921,1023,476,501]" name="Thailand" pageId="10" pageNumber="287">Thailand</collectingCountry>
,
<collectingRegion id="EB745D97FE5DBD38C36200B642BAFE3A" box="[1044,1189,476,501]" country="Thailand" name="Chiang Mai" pageId="10" pageNumber="287">Chiang Mai</collectingRegion>
,
<location id="2C6FC5AEFE5DBD38C3CF00B74305FE3A" LSID="urn:lsid:plazi:treatment:A1192263FE5FBD38C7E1004D42BBFD67:2C6FC5AEFE5DBD38C3CF00B74305FE3A" box="[1209,1306,476,501]" country="Thailand" name="Mae Ai" pageId="10" pageNumber="287" stateProvince="Chiang Mai">Mae Ai</location>
,
<collectingDate id="4D4A4C5DFE5DBD38C25900B64388FE3B" box="[1327,1431,476,500]" pageId="10" pageNumber="287" value="1992-07">
<date id="5D0EB5B5FE5DBD38C25900B64388FE3B" box="[1327,1431,476,500]" pageId="10" pageNumber="287" value="1992-07">VII.1992</date>
</collectingDate>
,
<collectorName id="8445F6A3FE5DBD38C7E1036A4714FDD7" box="[151,267,512,536]" pageId="10" pageNumber="287">N.Kagana</collectorName>
</materialsCitation>
;
<materialsCitation id="99D89928FE5DBD38C660036A4493FDD7" ID-GBIF-Occurrence="1852084155" box="[278,652,511,536]" collectingDate="1992-07" collectionCode="ZISP" collectorName="N. Kagana" country="Thailand" location="Mae Ai" pageId="10" pageNumber="287" specimenCount="24" specimenCount-female="24" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus id="F60B2DD7FE5DBD38C660036A4799FDD7" box="[278,390,512,536]" pageId="10" pageNumber="287" type="paratype">paratypes</typeStatus>
:
<specimenCount id="3FB658FCFE5DBD38C6E6036A4412FDD7" box="[400,525,511,536]" pageId="10" pageNumber="287" type="female">24 females</specimenCount>
, same data
</materialsCitation>
.
</paragraph>
<paragraph id="290F9375FE5DBD38C7B1034E445AFD4A" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="287">
<materialsCitation id="99D89928FE5DBD38C7B1034E441FFDAF" ID-GBIF-Occurrence="1852084153" collectionCode="RAS" country="Russia" location="Saint Petersburg" pageId="10" pageNumber="287" specimenCount="3" stateProvince="Saint Petersburg" typeStatus="holotype">
<emphasis id="1BC44F67FE5DBD38C7B1034E479CFDF2" bold="true" box="[199,387,548,573]" pageId="10" pageNumber="287">
<typeStatus id="F60B2DD7FE5DBD38C7B1034E471DFDF2" box="[199,258,548,573]" pageId="10" pageNumber="287">Type</typeStatus>
deposition
</emphasis>
. The
<typeStatus id="F60B2DD7FE5DBD38C6B2034E4439FDF2" box="[452,550,548,573]" pageId="10" pageNumber="287" type="holotype">holotype</typeStatus>
and four
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are deposited in the collection of the Zoological Institute of
<collectionCode id="4FA10BB0FE5DBD38C7E1032246CFFDAF" box="[151,208,584,608]" country="Myanmar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14256" name="Union of Burma Applied Research Institute" pageId="10" pageNumber="287">RAS</collectionCode>
,
<collectingRegion id="EB745D97FE5DBD38C797032D47BEFDAF" box="[225,417,583,608]" country="Russia" name="Saint Petersburg" pageId="10" pageNumber="287">Saint Petersburg</collectingRegion>
,
<collectingCountry id="51A7D3E5FE5DBD38C6C4032D441FFDAF" box="[434,512,583,608]" name="Russia" pageId="10" pageNumber="287">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="99D89928FE5DBD38C5640322445AFD4A" ID-GBIF-Occurrence="1852084151" country="Russia" location="Tyumen State University Museum of Zoology" pageId="10" pageNumber="287" specimenCount="20" stateProvince="Tyumen" typeStatus="paratype">
<specimenCount id="3FB658FCFE5DBD38C564032244B9FDAF" box="[530,678,584,608]" pageId="10" pageNumber="287" type="generic" typeStatus="paratypes">20 paratypes</specimenCount>
are deposited in the collection of the
<location id="2C6FC5AEFE5DBD38C30F03224761FD4A" LSID="urn:lsid:plazi:treatment:A1192263FE5FBD38C7E1004D42BBFD67:2C6FC5AEFE5DBD38C30F03224761FD4A" country="Russia" name="Tyumen State University Museum of Zoology" pageId="10" pageNumber="287" stateProvince="Tyumen">Tyumen State University Museum of Zoology</location>
,
<collectingRegion id="EB745D97FE5DBD38C6FF030747F8FD4A" box="[393,487,621,645]" country="Russia" name="Tyumen'" pageId="10" pageNumber="287">Tyumen</collectingRegion>
,
<collectingCountry id="51A7D3E5FE5DBD38C6850306445EFD4A" box="[499,577,620,645]" name="Russia" pageId="10" pageNumber="287">Russia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="61AAC0FEFE5DBD38C7B103E542BBFD67" box="[199,1188,655,680]" pageId="10" pageNumber="287" type="etymology">
<paragraph id="290F9375FE5DBD38C7B103E542BBFD67" blockId="10.[151,1437,151,680]" box="[199,1188,655,680]" pageId="10" pageNumber="287">
<emphasis id="1BC44F67FE5DBD38C7B103E54759FD67" bold="true" box="[199,326,655,680]" pageId="10" pageNumber="287">Etymology</emphasis>
. The name of the new species refers to its distribution in the Oriental region.
</paragraph>
</subSubSection>
</treatment>
</document>

55
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<taxonomicName id="A385A55281C754E12394254A79CE4019" ID-CoL="3VHW9" authority="Verh." authorityName="Verh." class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Lithobiomorpha" pageId="0" pageNumber="100" phylum="Arthropoda" rank="subSpecies" species="acuminatus" subSpecies="cassinensis">
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acuminatus cassinensis
<authority id="D436A7128F15C0740055F723567C884A" pageId="0" pageNumber="100">Verh.</authority>
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;
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<specimenCount id="86D4B1DBE72DC18EF32B7D85BFC1EFFF" pageId="0" pageNumber="100" type="generic">un ♂</specimenCount>
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. Questa specie era gia nota per la penisola sorrentina e per le isole di
<location id="065661177CFE286493A06B4343A9C8A4" LSID="urn:lsid:plazi:treatment:A11970EE916844710AD5EF3EA7BACFA3:065661177CFE286493A06B4343A9C8A4" name="Ischia" pageId="0" pageNumber="100">Ischia</location>
e
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.
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