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<mods:namePart id="7A296B410029E65344682E1F8B9D58DD">Xian, Chunlan</mods:namePart>
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<mods:namePart id="3E13F9F6B43E6439094558793CA73F48">Leong, Chi Man</mods:namePart>
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<bibRefCitation id="C567F3203EB4C57620898DB54B1C0FA2" author="Mai, Z Q" journalOrPublisher="Zootaxa" pageId="0" pageNumber="118110" refId="B10975682" refString="MBD Macao BioDatabase https://nature.iam.gov.mo/BioDatabase/c/catlist?category=Insecta 2023-12-19T00:00:00+02:00" year="2022">MBD (2022)</bibRefCitation>
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.80.12962" ID-PMC="PMC5543604" ID-Pensoft-Pub="1314-2003-80-33" ID-Pensoft-UUID="FFB61E2CFFBFFF8DFF93FFF3FFEFFFFA" ID-PubMed="28781556" ID-Zenodo-Dep="576387" ModsDocID="1314-2003-80-33" checkinTime="1493205412253" checkinUser="pensoft" docAuthor="Freeman, Craig C." docDate="2017" docId="A100A67E106C5F7CB2F1A835A3590EDC" docLanguage="en" docName="PhytoKeys 80: 33-39" docOrigin="PhytoKeys 80" docPubDate="2017-04-25" docSource="http://dx.doi.org/10.3897/phytokeys.80.12962" docTitle="Penstemon xylus A. Nelson, Bot. Gaz. 31 – 32. 1902." docType="treatment" docVersion="3" id="FFB61E2CFFBFFF8DFF93FFF3FFEFFFFA" lastPageNumber="37" masterDocId="FFB61E2CFFBFFF8DFF93FFF3FFEFFFFA" masterDocTitle="Nomenclatural novelties and notes in Penstemon (Plantaginaceae)" masterLastPageNumber="39" masterPageNumber="33" pageNumber="37" updateTime="1668141960729" updateUser="ExternalLinkService">
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<taxonomicName LSID="A100A67E-106C-5F7C-B2F1-A835A3590EDC" authority="A. Nelson, Bot. Gaz. 31 - 32. 1902." authorityName="A. Nelson, Bot. Gaz. 31 – 32. 1902." class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon xylus" order="Lamiales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="species" species="xylus">Penstemon xylus A. Nelson, Bot. Gaz. 31-32. 1902.</taxonomicName>
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<taxonomicName class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon caespitosus subsp. var. var. suffruticosus" order="Lamiales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="variety" species="caespitosus" subSpecies="var." variety="suffruticosus">Penstemon caespitosus var. suffruticosus</taxonomicName>
A. Gray in A. Gray et al., Syn. Fl. N. Amer. 2: 270. 1878 (non
<taxonomicName authorityName="Dougl. ex Benth" authorityYear="1846" class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon suffruticosus" order="Lamiales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="species" species="suffruticosus">Penstemon suffruticosus</taxonomicName>
Douglas ex Benth.);
<taxonomicName authorityName="N. H. Holmgren" authorityYear="1979" class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon tusharensis" order="Lamiales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="species" species="tusharensis">Penstemon tusharensis</taxonomicName>
N.H. Holmgren, Brittonia 31: 106. 1979, nom. illeg.
</paragraph>
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<typeStatus>Type</typeStatus>
.
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<paragraph pageId="4" pageNumber="37">
<materialsCitation collectingDate="2017-01-01" collectingDateMax="2017-12-31" collectingDateMin="2017-01-01" country="USA" location="Utah" specimenCount="1" stateProvince="Utah" typeStatus="holotype">
<collectingCountry name="United States of America">USA</collectingCountry>
.
<collectingRegion country="United States of America" name="Utah">Utah</collectingRegion>
. Beaver, 1877, Palmer s.n. (
<typeStatus>holotype</typeStatus>
: GH, GH00091198 [GH website image)]; probable isotype: ISC, ISC-v-0000867 [JSTOR image]
</materialsCitation>
.
</paragraph>
<paragraph pageId="4" pageNumber="37">
The correct name for this taxon at the rank of species has been the source of confusion. With the earlier name
<taxonomicName authorityName="Dougl. ex Benth" authorityYear="1846" class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon suffruticosus" order="Lamiales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="species" species="suffruticosus">
<emphasis italics="true" pageId="4" pageNumber="37">Penstemon suffruticosus</emphasis>
</taxonomicName>
Douglas ex Benth. blocking transfer of A.
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
varietal epithet
<taxonomicName authorityName="Dougl. ex Benth" authorityYear="1846" class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="suffruticosus" order="Lamiales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="species" species="suffruticosus">
<emphasis italics="true" pageId="4" pageNumber="37">suffruticosus</emphasis>
</taxonomicName>
at the rank of species,
<bibRefCitation DOI="https://doi.org/10.2307/2478562" author="Rydberg, PA" journalOrPublisher="Club" pageId="5" pageNumber="38" pagination="499 - 513" refId="B7" refString="Rydberg, PA, 1901. Studies on the Rocky Mountain flora.-VI. Bull. Torrey Bot. Club 28: 499 - 513, DOI: https://doi.org/10.2307/2478562" title="Studies on the Rocky Mountain flora. - VI. Bull. Torrey Bot." url="https://doi.org/10.2307/2478562" volume="28" year="1901">Rydberg (1901)</bibRefCitation>
published
<taxonomicName lsidName="P. suffrutescens" pageId="4" pageNumber="37" rank="species" species="suffrutescens">
<emphasis italics="true" pageId="4" pageNumber="37">P. suffrutescens</emphasis>
</taxonomicName>
in an attempt to provide a substitute. However, in doing so, he flagged
<taxonomicName lsidName="P. suffrutescens" pageId="4" pageNumber="37" rank="species" species="suffrutescens">
<emphasis italics="true" pageId="4" pageNumber="37">P. suffrutescens</emphasis>
</taxonomicName>
as a new species (sp. nov.). He also provided a brief diagnosis and cited a specimen (
<collectingCountry name="United States of America">USA</collectingCountry>
.
<collectingRegion country="United States of America" name="Colorado">Colorado</collectingRegion>
. Ridgway, woods, altitude
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.286" unit="ft" value="7500.0">7500 ft</quantity>
,
<date value="1895-06-20">20 Jun 1895</date>
, F. Tweedy 170;
<typeStatus>holotype</typeStatus>
: NY, NY00130526 [NY website image]) different from that of
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
type for
<taxonomicName lsidName="P. caespitosus subsp. var. var. suffruticosus" pageId="4" pageNumber="37" rank="variety" species="caespitosus" subSpecies="var." variety="suffruticosus">P. caespitosus var. suffruticosus</taxonomicName>
.
<normalizedToken originalValue="Rydberg’s">Rydberg's</normalizedToken>
name must be considered a validly published species name-not an avowed substitute for
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
name. As noted by
<bibRefCitation DOI="https://doi.org/10.2307/2806680" author="Holmgren, NH" journalOrPublisher="Brittonia" pageId="5" pageNumber="38" pagination="104 - 107" refId="B2" refString="Holmgren, NH, 1979. Nomenclatural changes in some Intermountain penstemons. Brittonia 31: 104 - 107, DOI: https://doi.org/10.2307/2806680" title="Nomenclatural changes in some Intermountain penstemons." url="https://doi.org/10.2307/2806680" volume="31" year="1979">Holmgren (1979)</bibRefCitation>
, the specimen cited by Rydberg is referable to
<taxonomicName lsidName="P. crandallii" pageId="4" pageNumber="37" rank="species" species="crandallii">
<emphasis italics="true" pageId="4" pageNumber="37">P. crandallii</emphasis>
</taxonomicName>
, where
<taxonomicName lsidName="P. suffrutescens" pageId="4" pageNumber="37" rank="species" species="suffrutescens">
<emphasis italics="true" pageId="4" pageNumber="37">P. suffrutescens</emphasis>
</taxonomicName>
must be placed as a synonym.
</paragraph>
<paragraph pageId="4" pageNumber="37">
<bibRefCitation DOI="https://doi.org/10.1086/328257" author="Nelson, A" journalOrPublisher="Contributions from the Rocky Mountain Herbarium" pageId="5" pageNumber="38" pagination="21 - 35" refId="B5" refString="Nelson, A, 1902. . Contributions from the Rocky Mountain Herbarium III: 21 - 35, DOI: https://doi.org/10.1086/328257" url="https://doi.org/10.1086/328257" volume="III" year="1902">Nelson (1902)</bibRefCitation>
published
<taxonomicName class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon xylus" order="Lamiales" pageId="4" pageNumber="37" phylum="Tracheophyta" rank="species" species="xylus">
<emphasis italics="true" pageId="4" pageNumber="37">Penstemon xylus</emphasis>
</taxonomicName>
and clearly stated it was a new name (nom. nov.) for
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
<taxonomicName lsidName="P. caespitosus subsp. var. var. suffruticosus" pageId="4" pageNumber="37" rank="variety" species="caespitosus" subSpecies="var." variety="suffruticosus">P. caespitosus var. suffruticosus</taxonomicName>
. Nelson cited a specimen (
<collectingCountry name="United States of America">USA</collectingCountry>
.
<collectingRegion country="United States of America" name="Colorado">Colorado</collectingRegion>
. Sapinero, 1898, H.N. Wheeler 446 [COLO, COLO00396499 [JSTOR image], NY, NY00130556 [NY website image], RM, RM0004372 [JSTOR image]) that he believed was referable to A.
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
concept of
<taxonomicName lsidName="P. caespitosus subsp. var. var. suffruticosus" pageId="4" pageNumber="37" rank="variety" species="caespitosus" subSpecies="var." variety="suffruticosus">P. caespitosus var. suffruticosus</taxonomicName>
. However, it too is referable to
<taxonomicName lsidName="P. crandallii" pageId="4" pageNumber="37" rank="species" species="crandallii">
<emphasis italics="true" pageId="4" pageNumber="37">P. crandallii</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.2307/2806680" author="Holmgren, NH" journalOrPublisher="Brittonia" pageId="5" pageNumber="38" pagination="104 - 107" refId="B2" refString="Holmgren, NH, 1979. Nomenclatural changes in some Intermountain penstemons. Brittonia 31: 104 - 107, DOI: https://doi.org/10.2307/2806680" title="Nomenclatural changes in some Intermountain penstemons." url="https://doi.org/10.2307/2806680" volume="31" year="1979">Holmgren (1979)</bibRefCitation>
, believing that neither
<normalizedToken originalValue="Rydberg’s">Rydberg's</normalizedToken>
nor
<normalizedToken originalValue="Nelson’s">Nelson's</normalizedToken>
name could be applied to
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
<taxonomicName lsidName="P. caespitosus subsp. var. var. suffruticosus" pageId="4" pageNumber="37" rank="variety" species="caespitosus" subSpecies="var." variety="suffruticosus">P. caespitosus var. suffruticosus</taxonomicName>
because of the specimens cited by each author, published
<taxonomicName lsidName="P. tusharensis" pageId="4" pageNumber="37" rank="species" species="tusharensis">
<emphasis italics="true" pageId="4" pageNumber="37">P. tusharensis</emphasis>
</taxonomicName>
as an avowed substitute. In spite of Nelson having cited a specimen that is not referable to
<taxonomicName lsidName="P. caespitosus subsp. var. var. suffruticosus" pageId="4" pageNumber="37" rank="variety" species="caespitosus" subSpecies="var." variety="suffruticosus">P. caespitosus var. suffruticosus</taxonomicName>
,
<taxonomicName lsidName="P. xylus" pageId="4" pageNumber="37" rank="species" species="xylus">
<emphasis italics="true" pageId="4" pageNumber="37">P. xylus</emphasis>
</taxonomicName>
must stand as the avowed substitute for
<taxonomicName lsidName="P. caespitosus subsp. var. var. suffruticosus" pageId="4" pageNumber="37" rank="variety" species="caespitosus" subSpecies="var." variety="suffruticosus">P. caespitosus var. suffruticosus</taxonomicName>
at the rank of species, making
<taxonomicName lsidName="P. tusharensis" pageId="4" pageNumber="37" rank="species" species="tusharensis">
<emphasis italics="true" pageId="4" pageNumber="37">P. tusharensis</emphasis>
</taxonomicName>
illegitimate and superfluous.
</paragraph>
</subSubSection>
</treatment>
</document>