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Schmiedeknecht, 1910
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; stateProvince: South East England; locality:
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; Event: eventDate:
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; year: 2011; month: 2; day: 22; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 1; country:
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; Event: eventDate:
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; year: 2011; month: 2; day: 22; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 1; country:
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; stateProvince: South East England; locality:
<location LSID="urn:lsid:plazi:treatment:D64F3BF1A106A30F8DC4904CCDAAFC8B:F9AB3998EAA8740CBB5FAC48D706812C" country="Great Britain" name="Isle of Wight, coast at Bembridge, main area" pageId="0" pageNumber="5176">Isle of Wight, coast at Bembridge, main area</location>
; Event: eventDate:
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; year: 2011; month: 2; day: 22; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 1; country:
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; stateProvince: South East England; locality:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 1; country:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 1; country:
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; stateProvince: South East England; locality:
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; year: 2011; month: 2; day: 22; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 1; country:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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Type status:
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. Occurrence: catalogNumber:
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Type status:
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. Occurrence: catalogNumber:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 2; country:
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<location LSID="urn:lsid:plazi:treatment:D64F3BF1A106A30F8DC4904CCDAAFC8B:1172481FD55C7C7D58708B6A39BF7BF5" country="Great Britain" name="South of Berwick, Bynea, Llanelli, Carmarthenshire" pageId="0" pageNumber="5176">South of Berwick, Bynea, Llanelli, Carmarthenshire</location>
; Event: eventDate:
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</materialsCitation>
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 2; country:
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; stateProvince: Wales; locality:
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; Event: eventDate:
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; year: 2011; month: 3; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 2; country:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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Type status:
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. Occurrence: catalogNumber:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 2; country:
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; stateProvince: Wales; locality:
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; Event: eventDate:
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; year: 2011; month: 3; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 2; country:
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; Event: eventDate:
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; year: 2011; month: 3; Record Level: institutionID: ZFMK; collectionID: MYR
</materialsCitation>
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Type status:
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. Occurrence: catalogNumber:
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</materialsCitation>
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Type status:
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. Occurrence: catalogNumber:
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; year: 2011; month: 3; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Event: eventDate:
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; year: 2011; month: 3; Record Level: institutionID: ZFMK; collectionID: MYR
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Type status:
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. Occurrence: catalogNumber:
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. Dordogne,
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 3; country:
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; Event: eventDate:
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Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
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; Taxon: scientificName: Trachysphaera lobata; phylum: Arthropoda; class: Diplopoda; order: Glomerida; family: Glomeridae; genus: Trachysphaera; taxonRank: species; scientificNameAuthorship: (Ribaut 1954); Location: locationID: 3; country:
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<paragraph pageId="0" pageNumber="5176">Notes</paragraph>
<paragraph pageId="0" pageNumber="5176">Addtional individual information in Table 4.</paragraph>
</subSubSection>
</treatment>
</document>

View file

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Figs.
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i, 2j,
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<paragraph id="5E59E435FFD0FF8B1CD8F8DE4EC5F9E4" blockId="10.[151,1437,1266,1760]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1CD8F8DE4C24FAE8" bold="true" box="[151,306,1266,1291]" pageId="10" pageNumber="104">Third instar.</emphasis>
<emphasis id="6C923827FFD0FF8B1D73F8DE4C60FAE8" box="[316,374,1266,1291]" italics="true" pageId="10" pageNumber="104">Body</emphasis>
: length 32.0 mm, width (prothorax) 6.0 mm; with tiny, stout setae; prosternal setae absent. Primary setal pattern: 2 HPA, 0 PD, 3 PSL, 0 MSD, 2 MSL, 0 MTD, 1 MTL, 0 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFD0FF8B1977F93A4899FACC" box="[1336,1423,1302,1327]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="10" pageNumber="104" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
).
<emphasis id="6C923827FFD0FF8B1CD8F9164DC2FAB0" box="[151,212,1338,1363]" italics="true" pageId="10" pageNumber="104">Head</emphasis>
: without uniform pile of short, golden setae; epicranial suture short; frontal sutures extend to the base of the antennae; width of head capsule 4.0 mm (
<figureCitation id="C6DDF8B0FFD0FF8B1E3EF9724FA5FA94" box="[625,691,1374,1399]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="10" pageNumber="104">
Fig.
<date id="2A58C2F5FFD0FF8B1EE8F9724FA5FA94" box="[679,691,1374,1399]" pageId="10" pageNumber="104">1</date>
</figureCitation>
i). Mandibles with micro-asperities on the ventral base. Maxillary stipes with 19 micro-conical projections (
<figureCitation id="C6DDF8B0FFD0FF8B1E28F9AE4FBDFA78" box="[615,683,1410,1435]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="10" pageNumber="104">Fig. 2</figureCitation>
j).
<emphasis id="6C923827FFD0FF8B1E88F9A84FEAFA78" box="[711,764,1412,1435]" italics="true" pageId="10" pageNumber="104">Legs</emphasis>
: pars stridens (mesocoxae) length 676.23 µm, width 625.65 µm, distance into striate 12.67 µm, 44 striae; mesocoxae and metacoxae with two internal setae. Metathoracic legs with some short lateral setae (side of I tooth); anterior margin with 5 teeth, I and IIIV well developed, I and III are about same size and are larger than IVV, II is the smallest tooth (
<figureCitation id="C6DDF8B0FFD0FF8B1F36F9C24EABF9E4" box="[889,957,1518,1543]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="10" pageNumber="104">
Fig.
<date id="2A58C2F5FFD0FF8B1FE1F9C24EABF9E4" box="[942,957,1518,1543]" pageId="10" pageNumber="104">3</date>
</figureCitation>
i).
</paragraph>
<paragraph id="5E59E435FFD0FF8B1CD8FA3E4924F9C8" blockId="10.[151,1437,1266,1760]" box="[151,1074,1554,1579]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1CD8FA3E4C17F9C8" box="[151,257,1554,1579]" italics="true" pageId="10" pageNumber="104">Abdomen</emphasis>
: raster without setae; anal ring with 14 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFD0FF8B1F97FA3E490AF9C8" box="[984,1052,1554,1579]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="10" pageNumber="104">
Fig.
<date id="2A58C2F5FFD0FF8B1843FA3E490AF9C8" box="[1036,1052,1554,1579]" pageId="10" pageNumber="104">4</date>
</figureCitation>
i).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD0FF8B1C88FA1A492CF974" pageId="10" pageNumber="104" type="distribution">
<paragraph id="5E59E435FFD0FF8B1C88FA1A492CF974" blockId="10.[151,1437,1266,1760]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1C88FA1A4C76F9AC" bold="true" box="[199,352,1590,1615]" pageId="10" pageNumber="104">Distribution.</emphasis>
Known species of the central Amazon Basin (
<collectingCountry id="26F1A4A5FFD0FF8B1F2CFA1A4EBBF9AC" box="[867,941,1590,1615]" name="Brazil" pageId="10" pageNumber="104">Brazil</collectingCountry>
: Amazonas and Pará) and the upper Orinoco Basin (
<collectingCountry id="26F1A4A5FFD0FF8B1CA6FA764C74F990" box="[233,354,1626,1651]" name="Venezuela" pageId="10" pageNumber="104">Venezuela</collectingCountry>
: Amazonas) (
<bibRefCitation id="3A7799C4FFD0FF8B1E4EFA764FBFF990" author="Boucher" box="[513,681,1626,1651]" pageId="10" pageNumber="104" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
). The locality where was collected the material examined accord with the species distribution. This confirms a new record of
<taxonomicName id="99E69FB6FFD0FF8B1F76FA534E84F975" box="[825,914,1663,1686]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFD0FF8B1F76FA534E84F975" box="[825,914,1663,1686]" italics="true" pageId="10" pageNumber="104">Veturius</emphasis>
</taxonomicName>
for
<collectingCountry id="26F1A4A5FFD0FF8B1F8CFA524920F974" box="[963,1078,1662,1687]" name="Colombia" pageId="10" pageNumber="104">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD0FF8B1C88FA8F4EC0F903" pageId="10" pageNumber="104" type="materials_examined">
<paragraph id="5E59E435FFD0FF8B1C88FA8F4EC0F903" blockId="10.[151,1437,1266,1760]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1C88FA8F4CA5F95F" bold="true" box="[199,435,1699,1724]" pageId="10" pageNumber="104">Material examined.</emphasis>
<collectionCode id="38F77CF0FFD0FF8B1DF2FA884F43F95F" box="[445,597,1700,1724]" pageId="10" pageNumber="104">
<collectingCountry id="26F1A4A5FFD0FF8B1DF2FA884F43F95F" box="[445,597,1700,1724]" name="Colombia" pageId="10" pageNumber="104">COLOMBIA</collectingCountry>
</collectionCode>
, GUAINIA, Inirida, Com. La Ceiba,
<geoCoordinate id="3BD282F2FFD0FF8B1845FA884985F95F" box="[1034,1171,1698,1724]" direction="north" orientation="latitude" pageId="10" pageNumber="104" precision="1" value="3.6329722">3°37'58.7''N</geoCoordinate>
<geoCoordinate id="3BD282F2FFD0FF8B18D1FA8F482BF95F" box="[1182,1341,1698,1724]" direction="west" orientation="longitude" pageId="10" pageNumber="104" precision="1" value="-67.88947">67°53'22.1''W</geoCoordinate>
,
<quantity id="991E49D0FFD0FF8B1904FA884881F958" box="[1355,1431,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="10" pageNumber="104" unit="m" value="103.0">103 m</quantity>
,
<date id="2A58C2F5FFD0FF8B1CD8FAEB4C05F903" box="[151,275,1735,1760]" pageId="10" pageNumber="104" value="1998-03-18">18.iii.1998</date>
, Bosque. G. Amat, A. Garbanzo, &amp; C. Martínez (
<specimenCount id="48E02FBCFFD0FF8B1F75FAE44EDCF903" box="[826,970,1735,1760]" pageId="10" pageNumber="104" type="generic">2 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,114 @@
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<mods:title id="A93D8454C04018EFFEF4207AE1608FA3">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<treatment id="D64F5523FFD0FF8B1CD8FE084EA0FBD7" ID-DOI="http://doi.org/10.5281/zenodo.5618110" ID-GBIF-Taxon="119597921" ID-Zenodo-Dep="5618110" LSID="urn:lsid:plazi:treatment:D64F5523FFD0FF8B1CD8FE084EA0FBD7" httpUri="http://treatment.plazi.org/id/D64F5523FFD0FF8B1CD8FE084EA0FBD7" lastPageNumber="104" pageId="10" pageNumber="104">
<subSubSection id="16FCB7BEFFD0FF8B1CD8FE084DD5FC2B" pageId="10" pageNumber="104" type="nomenclature">
<paragraph id="5E59E435FFD0FF8B1CD8FE084FACFDDC" blockId="10.[151,698,548,608]" box="[151,698,548,575]" pageId="10" pageNumber="104">
<heading id="05115359FFD0FF8B1CD8FE084FACFDDC" bold="true" box="[151,698,548,575]" fontSize="11" level="1" pageId="10" pageNumber="104" reason="1">
<taxonomicName id="99E69FB6FFD0FF8B1CD8FE084FACFDDC" authority="Hincks, 1933" authorityName="Hincks" authorityYear="1933" box="[151,698,548,575]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="104" phylum="Arthropoda" rank="species" species="oberthuri" subGenus="Ouayana">
<emphasis id="6C923827FFD0FF8B1CD8FE084FACFDDC" bold="true" box="[151,698,548,575]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1CD8FE084DEAFDDD" bold="true" box="[151,252,548,574]" italics="true" pageId="10" pageNumber="104">Veturius</emphasis>
(
<emphasis id="6C923827FFD0FF8B1D42FE084C6AFDDD" bold="true" box="[269,380,548,574]" italics="true" pageId="10" pageNumber="104">Ouayana</emphasis>
)
<emphasis id="6C923827FFD0FF8B1DC3FE084CE8FDDD" bold="true" box="[396,510,548,574]" italics="true" pageId="10" pageNumber="104">oberthuri</emphasis>
(Hincks, 1933)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="5E59E435FFD0FF8B1CD8FE6B4C6AFD83" blockId="10.[151,698,548,608]" box="[151,380,583,608]" pageId="10" pageNumber="104">
(
<figureCitation id="C6DDF8B0FFD0FF8B1CD0FE6B4DFFFD83" box="[159,233,583,608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="10" pageNumber="104">Figs. 1</figureCitation>
h,
<date id="2A58C2F5FFD0FF8B1D47FE6B4C36FD83" box="[264,288,583,608]" pageId="10" pageNumber="104">2i</date>
, 3h, 4h)
</paragraph>
<paragraph id="5E59E435FFD0FF8B1CD8FEA34DD5FC2B" blockId="10.[151,1437,655,1077]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1CD8FEA34C32FD4B" bold="true" box="[151,292,655,680]" pageId="10" pageNumber="104">First instar.</emphasis>
<emphasis id="6C923827FFD0FF8B1D62FEA34C71FD4B" box="[301,359,655,680]" italics="true" pageId="10" pageNumber="104">Body</emphasis>
: length 10.0 mm, width (prothorax) 3.0 mm; with uniform pile of fine short setae; prosternal setae absent. Primary setal pattern: 10 HPA, 0 PD, 3 PSL, 0 MSD, 2 MSL, 0 MTD, 2 MTL, 1 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFD0FF8B1C93FEFB4C20FD13" box="[220,310,727,752]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="10" pageNumber="104" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
), with the addition of an extra pair of TD setae in all abdominal segments.
<emphasis id="6C923827FFD0FF8B18F8FEFB49E2FD13" box="[1207,1268,727,752]" italics="true" pageId="10" pageNumber="104">Head</emphasis>
: covered with uniform pile of short, golden setae; epicranial suture short; frontal sutures extend to the base of the antennae; width of head capsule 2.0 mm (
<figureCitation id="C6DDF8B0FFD0FF8B1DF3FF334CE8FCDB" box="[444,510,799,824]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="10" pageNumber="104">Fig. 1</figureCitation>
h). Mandibles with micro-asperities in the ventral base. Maxillary stipes with 17 micro-conical projections (
<figureCitation id="C6DDF8B0FFD0FF8B1D89FF684F1DFCBE" box="[454,523,836,861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="10" pageNumber="104">
Fig.
<date id="2A58C2F5FFD0FF8B1DB4FF684F1DFCBE" box="[507,523,836,861]" pageId="10" pageNumber="104">2</date>
</figureCitation>
i).
<emphasis id="6C923827FFD0FF8B1E66FF694F48FCBF" box="[553,606,837,860]" italics="true" pageId="10" pageNumber="104">Legs</emphasis>
: pars stridens (mesocoxae) length 417.35 µm, width 357.29 µm, distance into striate 7.38 µm, 40 striae. Metathoracic legs with some short lateral setae (side of I tooth); anterior margin with 6 teeth, all about same size (
<figureCitation id="C6DDF8B0FFD0FF8B1D95FFA04F0BFC46" box="[474,541,908,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="10" pageNumber="104">Fig. 3</figureCitation>
h).
<emphasis id="6C923827FFD0FF8B1E0CFFA04FBBFC46" box="[579,685,908,933]" italics="true" pageId="10" pageNumber="104">Abdomen</emphasis>
: raster with setae; anal ring with 18 setae; anal slit Y-shaped (
<figureCitation id="C6DDF8B0FFD0FF8B1920FFA04DB1FC2B" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="10" pageNumber="104">Fig. 4</figureCitation>
h).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD0FF8B1C88FFF84EE9FC0E" box="[199,1023,980,1005]" pageId="10" pageNumber="104" type="distribution">
<paragraph id="5E59E435FFD0FF8B1C88FFF84EE9FC0E" blockId="10.[151,1437,655,1077]" box="[199,1023,980,1005]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1C88FFF84C76FC0E" bold="true" box="[199,352,980,1005]" pageId="10" pageNumber="104">Distribution.</emphasis>
Colombian Central and Eastern Cordillera (
<bibRefCitation id="3A7799C4FFD0FF8B1F01FFF84EE5FC0E" author="Boucher" box="[846,1011,980,1005]" pageId="10" pageNumber="104" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD0FF8B1C88FFDB4EA0FBD7" pageId="10" pageNumber="104" type="materials_examined">
<paragraph id="5E59E435FFD0FF8B1C88FFDB4EA0FBD7" blockId="10.[151,1437,655,1077]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1C88FFDB4CA5FBF3" bold="true" box="[199,435,1015,1040]" pageId="10" pageNumber="104">Material examined.</emphasis>
<collectionCode id="38F77CF0FFD0FF8B1DF3FFD44F42FBF3" box="[444,596,1016,1040]" pageId="10" pageNumber="104">
<collectingCountry id="26F1A4A5FFD0FF8B1DF3FFD44F42FBF3" box="[444,596,1016,1040]" name="Colombia" pageId="10" pageNumber="104">COLOMBIA</collectingCountry>
</collectionCode>
, ANTIOQUIA, Gómez Plata, vereda Vega Botero,
<quantity id="991E49D0FFD0FF8B18E9FFD449E9FBEC" box="[1190,1279,1016,1040]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="10" pageNumber="104" unit="m" value="1070.0">1070 m</quantity>
,
<date id="2A58C2F5FFD0FF8B1941FFDB488FFBF3" box="[1294,1433,1015,1040]" pageId="10" pageNumber="104" value="2011-07-11">11.vii. 2011</date>
. Reyes-Castillo, Jiménez-Ferbans, Salazar-Niño, &amp; Amat-García (3 ex).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,115 @@
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<mods:title id="D4B4AF4F00F6D4D371E97F9578CDDF59">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<mods:name id="4399E2D973CB72042B150469CEC00BF3" type="personal">
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<mods:namePart id="868F1280CF000DA59A95982D66A9DE4C">Salazar-Niño, Karen</mods:namePart>
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<mods:namePart id="F82A1464912FBC79F29B45F540C239FE">Serrão, José Eduardo</mods:namePart>
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<mods:date id="E93D33374B5F54D1BE803C400EBE620C">2015</mods:date>
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<treatment id="D64F5523FFD2FF881CD8FAC94C96FDFB" ID-DOI="http://doi.org/10.5281/zenodo.5618106" ID-GBIF-Taxon="119597922" ID-Zenodo-Dep="5618106" LSID="urn:lsid:plazi:treatment:D64F5523FFD2FF881CD8FAC94C96FDFB" httpUri="http://treatment.plazi.org/id/D64F5523FFD2FF881CD8FAC94C96FDFB" lastPageId="9" lastPageNumber="103" pageId="8" pageNumber="102">
<subSubSection id="16FCB7BEFFD2FF881CD8FAC94FC5FEA3" lastPageId="9" lastPageNumber="103" pageId="8" pageNumber="102" type="nomenclature">
<paragraph id="5E59E435FFD2FF891CD8FAC94F95F8E3" blockId="8.[151,643,1765,1826]" box="[151,643,1765,1792]" pageId="8" pageNumber="102">
<heading id="05115359FFD2FF891CD8FAC94F95F8E3" bold="true" box="[151,643,1765,1792]" fontSize="11" level="1" pageId="8" pageNumber="102" reason="1">
<taxonomicName id="99E69FB6FFD2FF891CD8FAC94F95F8E3" authority="Smith, 1852" authorityName="Smith" authorityYear="1852" box="[151,643,1765,1792]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="102" phylum="Arthropoda" rank="species" species="crassus" subGenus="Publius">
<emphasis id="6C923827FFD2FF891CD8FAC94F95F8E3" bold="true" box="[151,643,1765,1792]" pageId="8" pageNumber="102">
<emphasis id="6C923827FFD2FF891CD8FAC94DEAF91C" bold="true" box="[151,252,1765,1791]" italics="true" pageId="8" pageNumber="102">Veturius</emphasis>
(
<emphasis id="6C923827FFD2FF891D42FAC94C7CF91C" bold="true" box="[269,362,1765,1791]" italics="true" pageId="8" pageNumber="102">Publius</emphasis>
)
<emphasis id="6C923827FFD2FF891D34FAC94CC2F91C" bold="true" box="[379,468,1765,1791]" italics="true" pageId="8" pageNumber="102">crassus</emphasis>
(Smith, 1852)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="5E59E435FFD2FF891CD8FB254C79F8C1" blockId="8.[151,643,1765,1826]" box="[151,367,1801,1826]" pageId="8" pageNumber="102">
(
<figureCitation id="C6DDF8B0FFD2FF891CD0FB254DFFF8C1" box="[159,233,1801,1826]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="8" pageNumber="102">Figs. 1</figureCitation>
f, 2f, 3f, 4f)
</paragraph>
<paragraph id="5E59E435FFD2FF881CD8FB7D4FC5FEA3" blockId="8.[151,1437,1873,2006]" lastBlockId="9.[151,1437,151,536]" lastPageId="9" lastPageNumber="103" pageId="8" pageNumber="102">
<emphasis id="6C923827FFD2FF891CD8FB7D4C24F889" bold="true" box="[151,306,1873,1898]" pageId="8" pageNumber="102">Third instar.</emphasis>
<emphasis id="6C923827FFD2FF891D73FB7D4C60F889" box="[316,374,1873,1898]" italics="true" pageId="8" pageNumber="102">Body</emphasis>
: length 38.0 mm, width (prothorax) 8.0 mm; with tiny, stout setae; prosternal setae absent. Primary setal pattern: 3 HPA, 0 PD, 3PSL, 0 MSD, 2 MSL, 0 MTD, 2 MTL, 0 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFD2FF891978FB594899F86D" box="[1335,1423,1909,1934]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="8" pageNumber="102" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
). TM setae present only on abdominal segment IX.
<emphasis id="6C923827FFD2FF891E87FBB54E12F851" box="[712,772,1945,1970]" italics="true" pageId="8" pageNumber="102">Head</emphasis>
: without uniform pile of short, golden setae; width of head capsule 6.0 mm. Epicranial suture short; frontal sutures extend to base of antennae (
<figureCitation id="C6DDF8B0FFD2FF89182FFB9149B7F835" box="[1120,1185,1981,2006]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="8" pageNumber="102">Fig. 1</figureCitation>
f). Mandibles without micro-asperities on the ventral base. Maxillary stipes with 20 micro-conical projections (
<figureCitation id="C6DDF8B0FFD3FF8818F9FCBB49E9FF53" box="[1206,1279,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="9" pageNumber="103">Fig. 2</figureCitation>
f).
<emphasis id="6C923827FFD3FF88196BFCB5484FFF53" box="[1316,1369,153,176]" italics="true" pageId="9" pageNumber="103">Legs</emphasis>
: pars stridens (mesocoxae) length 636.02 µm, width 569.30 µm, distance into striate 12.72 µm, 42 striae; mesocoxae and metacoxae with 2 internal setae. Metathoracic legs with some very short setae; anterior margin with 6 teeth, I and IIIVI well developed, I and III are larger than IVVI, II is the smallest tooth (
<figureCitation id="C6DDF8B0FFD3FF88184BFD284950FEFE" box="[1028,1094,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="9" pageNumber="103">Fig. 3</figureCitation>
f).
<emphasis id="6C923827FFD3FF881829FD2849C6FEFE" box="[1126,1232,260,285]" italics="true" pageId="9" pageNumber="103">Abdomen</emphasis>
: raster with setae; anal ring with 18 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFD3FF881E38FD0B4FACFEA3" box="[631,698,295,320]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="9" pageNumber="103">Fig. 4</figureCitation>
f).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD3FF881C88FD604EEEFE33" pageId="9" pageNumber="103" type="distribution">
<paragraph id="5E59E435FFD3FF881C88FD604EEEFE33" blockId="9.[151,1437,151,536]" pageId="9" pageNumber="103">
<emphasis id="6C923827FFD3FF881C88FD604C76FE86" bold="true" box="[199,352,332,357]" pageId="9" pageNumber="103">Distribution.</emphasis>
Endemic species from center of the Colombian Eastern Cordillera (
<bibRefCitation id="3A7799C4FFD3FF88181EFD6049E3FE86" author="Boucher" box="[1105,1269,332,357]" pageId="9" pageNumber="103" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
). However, the records cited in the Colombian Central Cordillera are considered erroneous because the altitude less than the average for the species, also the climate and forests are different (
<bibRefCitation id="3A7799C4FFD3FF881F34FDB8490BFE4E" author="Boucher" box="[891,1053,404,429]" pageId="9" pageNumber="103" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
) or the records need confirmation because the collection localities are doubtful (K.S.-N., personal observation).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD3FF881C88FDF04C96FDFB" pageId="9" pageNumber="103" type="materials_examined">
<paragraph id="5E59E435FFD3FF881C88FDF04C96FDFB" blockId="9.[151,1437,151,536]" pageId="9" pageNumber="103">
<emphasis id="6C923827FFD3FF881C88FDF04CA7FE16" bold="true" box="[199,433,476,501]" pageId="9" pageNumber="103">Material examined.</emphasis>
<collectionCode id="38F77CF0FFD3FF881DF7FDF04F46FE17" box="[440,592,476,500]" pageId="9" pageNumber="103">
<collectingCountry id="26F1A4A5FFD3FF881DF7FDF04F46FE17" box="[440,592,476,500]" name="Colombia" pageId="9" pageNumber="103">COLOMBIA</collectingCountry>
</collectionCode>
, CUNDINAMARCA,
<collectingCountry id="26F1A4A5FFD3FF881F1DFDF04E8FFE16" box="[850,921,476,501]" name="United Kingdom" pageId="9" pageNumber="103">Albán</collectingCountry>
, Granja del padre Luna,
<quantity id="991E49D0FFD3FF8818E0FDF04810FE17" box="[1199,1286,476,500]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="103" unit="m" value="2200.0">2200 m</quantity>
,
<date id="2A58C2F5FFD3FF88195DFDF0486FFE16" box="[1298,1401,476,501]" pageId="9" pageNumber="103" value="2000-02-09">9.ii.2000</date>
. G. Amat (
<specimenCount id="48E02FBCFFD3FF881CABFE2C4C62FDFB" box="[228,372,511,536]" pageId="9" pageNumber="103" type="generic">4 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D9FFA5567C8F9599C6BB2726310EFCAB">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<mods:namePart id="90E2F8154D0AE1A8E97D1329BC295F3D">Serrão, José Eduardo</mods:namePart>
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<treatment id="D64F5523FFD3FF8B1CD8FE47497DFE32" ID-DOI="http://doi.org/10.5281/zenodo.5618108" ID-GBIF-Taxon="119597919" ID-Zenodo-Dep="5618108" LSID="urn:lsid:plazi:treatment:D64F5523FFD3FF8B1CD8FE47497DFE32" httpUri="http://treatment.plazi.org/id/D64F5523FFD3FF8B1CD8FE47497DFE32" lastPageId="10" lastPageNumber="104" pageId="9" pageNumber="103">
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<paragraph id="5E59E435FFD3FF881CD8FE474F8AFD65" blockId="9.[151,668,619,679]" box="[151,668,619,646]" pageId="9" pageNumber="103">
<heading id="05115359FFD3FF881CD8FE474F8AFD65" bold="true" box="[151,668,619,646]" fontSize="11" level="1" pageId="9" pageNumber="103" reason="1">
<taxonomicName id="99E69FB6FFD3FF881CD8FE474F8AFD65" authority="Hincks, 1934" authorityName="Hincks" authorityYear="1934" box="[151,668,619,646]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="103" phylum="Arthropoda" rank="species" species="impressus" subGenus="Publius">
<emphasis id="6C923827FFD3FF881CD8FE474F8AFD65" bold="true" box="[151,668,619,646]" pageId="9" pageNumber="103">
<emphasis id="6C923827FFD3FF881CD8FE474DEAFD66" bold="true" box="[151,252,619,645]" italics="true" pageId="9" pageNumber="103">Veturius</emphasis>
(
<emphasis id="6C923827FFD3FF881D42FE474C7CFD66" bold="true" box="[269,362,619,645]" italics="true" pageId="9" pageNumber="103">Publius</emphasis>
)
<emphasis id="6C923827FFD3FF881D34FE474CE5FD66" bold="true" box="[379,499,619,645]" italics="true" pageId="9" pageNumber="103">impressus</emphasis>
Hincks, 1934
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="5E59E435FFD3FF881CD8FEA24CE7FD44" blockId="9.[151,668,619,679]" box="[151,497,654,679]" pageId="9" pageNumber="103">
(
<figureCitation id="C6DDF8B0FFD3FF881CD0FEA24DFFFD44" box="[159,233,654,679]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="9" pageNumber="103">
Figs.
<quantity id="991E49D0FFD3FF881C91FEA34DFFFD44" box="[222,233,655,679]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="9" pageNumber="103" unit="g" value="1.0">1</quantity>
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g,
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, 2h,
<quantity id="991E49D0FFD3FF881D12FEA34C6CFD44" box="[349,378,655,679]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="9" pageNumber="103" unit="g" value="3.0">3g</quantity>
,
<quantity id="991E49D0FFD3FF881DC9FEA34CB3FD44" box="[390,421,655,679]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="9" pageNumber="103" unit="g" value="4.0">4g</quantity>
, 6, 9c)
</paragraph>
<paragraph id="5E59E435FFD3FF881CD8FEFA49E6FBEC" blockId="9.[151,1437,726,1039]" pageId="9" pageNumber="103">
<emphasis id="6C923827FFD3FF881CD8FEFA4C24FD0C" bold="true" box="[151,306,726,751]" pageId="9" pageNumber="103">Third instar.</emphasis>
<emphasis id="6C923827FFD3FF881D73FEFA4C60FD0C" box="[316,374,726,751]" italics="true" pageId="9" pageNumber="103">Body</emphasis>
: length 50.055.0 mm, width (prothorax) 9.0 mm; covered with tiny, stout setae (
<figureCitation id="C6DDF8B0FFD3FF881976FEFA4896FD0C" box="[1337,1408,726,751]" captionStart="FIGURE 9" captionStartId="12.[151,250,1377,1399]" captionTargetBox="[151,1436,193,1355]" captionTargetId="figure@12.[151,1436,193,1356]" captionTargetPageId="12" captionText="FIGURE 9. Light micrographs of the abdominal segment V (laterodorsal view), showing the setae of the body. (a) Uniform pile of fine short setae (Veturius aspina), with detail of the setae. (b) Tiny, stout setae (V. simillimus), with detail of the setae. (c) Tiny, stout setae (V. impressus), with detail of the setae. Scale 0.1 mm." httpUri="https://zenodo.org/record/232519/files/figure.png" pageId="9" pageNumber="103">Fig. 9</figureCitation>
c); prosternal setae absent. Primary setal pattern: 3 HPA, 0 PD, 3 PSL, 0 MSD, 2 MSL, 0 MTD, 12 MTL, 0 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFD3FF881DCCFF324CCBFCD4" box="[387,477,798,823]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="9" pageNumber="103" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
).
<emphasis id="6C923827FFD3FF881DBCFF324F39FCD4" box="[499,559,798,823]" italics="true" pageId="9" pageNumber="103">Head</emphasis>
: without uniform pile of short, golden setae; width of head capsule 7.0 mm. Epicranial suture short; frontal sutures extend to base of antennae (
<figureCitation id="C6DDF8B0FFD3FF881FCBFF6E4ED2FCB8" box="[900,964,834,859]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="9" pageNumber="103">
Fig.
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g). Mandibles with micro-asperities in the ventral base. Maxillary stipes with 22 micro-conical projections (
<figureCitation id="C6DDF8B0FFD3FF881FDFFF4A4EF4FC9C" box="[912,994,870,895]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="9" pageNumber="103">
Figs.
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g, h).
<emphasis id="6C923827FFD3FF881865FF4B4949FC9D" box="[1066,1119,871,894]" italics="true" pageId="9" pageNumber="103">Legs</emphasis>
: pars stridens (mesocoxae) length 776.29 µm, width 498.24 µm, distance into striate 13.97 µm, 55 striae; mesocoxae and metacoxae with 2 internal setae. Metathoracic legs with some short lateral setae (side of I tooth) and micro-asperities in the posterior areas; anterior margin with 6 teeth, I and IIIVI well developed and gradually decreasing in size, II is the smallest tooth (
<figureCitation id="C6DDF8B0FFD3FF881CAFFFDA4C34FBEC" box="[224,290,1014,1039]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="9" pageNumber="103">
Fig.
<quantity id="991E49D0FFD3FF881D5BFFDB4C34FBEC" box="[276,290,1015,1039]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="9" pageNumber="103" unit="g" value="3.0">3</quantity>
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g).
<emphasis id="6C923827FFD3FF881D08FFDA4CA7FBEC" box="[327,433,1014,1039]" italics="true" pageId="9" pageNumber="103">Abdomen</emphasis>
: raster with setae; anal ring with 1516 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFD3FF8818C0FFDA49C5FBEC" box="[1167,1235,1014,1039]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="9" pageNumber="103">
Fig.
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</figureCitation>
g).
</paragraph>
<caption id="0A99B4BDFFD3FF881CD8FB0D4C65F8B5" httpUri="https://zenodo.org/record/232516/files/figure.png" pageId="9" pageNumber="103" targetBox="[163,1429,1101,1803]" targetPageId="9">
<paragraph id="5E59E435FFD3FF881CD8FB0D4C65F8B5" blockId="9.[151,1436,1825,1878]" pageId="9" pageNumber="103">
<emphasis id="6C923827FFD3FF881CD8FB0D4C02F8D4" bold="true" box="[151,276,1825,1847]" pageId="9" pageNumber="103">FIGURE 6.</emphasis>
Light micrographs of pupa of
<taxonomicName id="99E69FB6FFD3FF881E19FB0E4E03F8D4" box="[598,789,1826,1847]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="103" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="6C923827FFD3FF881E19FB0E4E03F8D4" box="[598,789,1826,1847]" italics="true" pageId="9" pageNumber="103">Veturius impressus</emphasis>
</taxonomicName>
.
<emphasis id="6C923827FFD3FF881F6BFB0D4E57F8D4" bold="true" box="[804,833,1825,1847]" pageId="9" pageNumber="103">(a)</emphasis>
Dorsal view,
<emphasis id="6C923827FFD3FF881F9CFB0D4EE4F8D4" bold="true" box="[979,1010,1825,1847]" pageId="9" pageNumber="103">(b)</emphasis>
lateral view,
<emphasis id="6C923827FFD3FF881830FB0D498AF8D4" bold="true" box="[1151,1180,1825,1847]" pageId="9" pageNumber="103">(c)</emphasis>
ventral view,
<emphasis id="6C923827FFD3FF88197FFB0D4859F8D4" bold="true" box="[1328,1359,1825,1847]" pageId="9" pageNumber="103">(d)</emphasis>
genital ampulla. Scale 2 mm.
</paragraph>
</caption>
<paragraph id="5E59E435FFD3FF8B1C88FB514CADFE86" blockId="9.[151,1436,1917,2014]" lastBlockId="10.[151,1437,151,465]" lastPageId="10" lastPageNumber="104" pageId="9" pageNumber="103">
<emphasis id="6C923827FFD3FF881C88FB514C1AF875" bold="true" box="[199,268,1917,1942]" pageId="9" pageNumber="103">Pupa.</emphasis>
<emphasis id="6C923827FFD3FF881D5AFB514C59F875" box="[277,335,1917,1942]" italics="true" pageId="9" pageNumber="103">Body</emphasis>
: length 40.0 mm, width (prothorax) 17.0 mm. Body shape elongate, oval, stout, exarate, creamwhite (
<figureCitation id="C6DDF8B0FFD3FF881CABFB8D4C3EF859" box="[228,296,1953,1978]" captionStart="FIGURE 6" captionStartId="9.[151,250,1825,1847]" captionTargetBox="[163,1429,1101,1803]" captionTargetId="figure@9.[151,1437,1092,1804]" captionTargetPageId="9" captionText="FIGURE 6. Light micrographs of pupa of Veturius impressus. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) genital ampulla. Scale 2 mm." httpUri="https://zenodo.org/record/232516/files/figure.png" pageId="9" pageNumber="103">Fig. 6</figureCitation>
). Head visible in dorsal view. Antennae, labrum, mandibles, maxillae, maxillary palpi, and compound eyes discernible (
<figureCitation id="C6DDF8B0FFD3FF881D13FBE94CBAF83D" box="[348,428,1989,2014]" captionStart="FIGURE 6" captionStartId="9.[151,250,1825,1847]" captionTargetBox="[163,1429,1101,1803]" captionTargetId="figure@9.[151,1437,1092,1804]" captionTargetPageId="9" captionText="FIGURE 6. Light micrographs of pupa of Veturius impressus. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) genital ampulla. Scale 2 mm." httpUri="https://zenodo.org/record/232516/files/figure.png" pageId="9" pageNumber="103">Figs. 6</figureCitation>
a, c).
<emphasis id="6C923827FFD3FF881DA3FBE94F2BF83D" box="[492,573,1989,2014]" italics="true" pageId="9" pageNumber="103">Thorax</emphasis>
: elytral and posterior wing pads closely juxtaposed curved ventrally around body, extending to the middle of abdominal segment V (
<figureCitation id="C6DDF8B0FFD0FF8B1F58FCBB4E70FF53" box="[791,870,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1825,1847]" captionTargetBox="[163,1429,1101,1803]" captionTargetId="figure@9.[151,1437,1092,1804]" captionTargetPageId="9" captionText="FIGURE 6. Light micrographs of pupa of Veturius impressus. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) genital ampulla. Scale 2 mm." httpUri="https://zenodo.org/record/232516/files/figure.png" pageId="10" pageNumber="104">Figs. 6</figureCitation>
a, b).
<emphasis id="6C923827FFD0FF8B1FE9FCBB4906FF53" box="[934,1040,151,176]" italics="true" pageId="10" pageNumber="104">Abdomen</emphasis>
: segment VIII visible under dorsal view and well defined (
<figureCitation id="C6DDF8B0FFD0FF8B1DECFC904CF1FF36" box="[419,487,188,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1825,1847]" captionTargetBox="[163,1429,1101,1803]" captionTargetId="figure@9.[151,1437,1092,1804]" captionTargetPageId="9" captionText="FIGURE 6. Light micrographs of pupa of Veturius impressus. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) genital ampulla. Scale 2 mm." httpUri="https://zenodo.org/record/232516/files/figure.png" pageId="10" pageNumber="104">Fig. 6</figureCitation>
a), but 10 are considered to exist, abdominal segments IX and X related to genitalia formation; segments IVVIII in ventral view well defined (
<figureCitation id="C6DDF8B0FFD0FF8B1F70FCF34E90FF1B" box="[831,902,223,248]" captionStart="FIGURE 6" captionStartId="9.[151,250,1825,1847]" captionTargetBox="[163,1429,1101,1803]" captionTargetId="figure@9.[151,1437,1092,1804]" captionTargetPageId="9" captionText="FIGURE 6. Light micrographs of pupa of Veturius impressus. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) genital ampulla. Scale 2 mm." httpUri="https://zenodo.org/record/232516/files/figure.png" pageId="10" pageNumber="104">Fig. 6</figureCitation>
c); segment X with small genital ampulla (
<figureCitation id="C6DDF8B0FFD0FF8B1920FCF34DB0FEFE" captionStart="FIGURE 6" captionStartId="9.[151,250,1825,1847]" captionTargetBox="[163,1429,1101,1803]" captionTargetId="figure@9.[151,1437,1092,1804]" captionTargetPageId="9" captionText="FIGURE 6. Light micrographs of pupa of Veturius impressus. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) genital ampulla. Scale 2 mm." httpUri="https://zenodo.org/record/232516/files/figure.png" pageId="10" pageNumber="104">Fig. 6</figureCitation>
d); spiracle I elongate, with fine peritreme and covered by wing pads; spiracles IIIII oval, prominent, with strongly sclerotized peritreme; spiracle IVVII with fine peritreme. Spiracle VII smaller than the others. Abdominal apex rounded.
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD0FF8B1C88FD43497DFE32" pageId="10" pageNumber="104" type="distribution">
<paragraph id="5E59E435FFD0FF8B1C88FD43497DFE32" blockId="10.[151,1437,151,465]" pageId="10" pageNumber="104">
<emphasis id="6C923827FFD0FF8B1C88FD434C76FE6B" bold="true" box="[199,352,367,392]" pageId="10" pageNumber="104">Distribution.</emphasis>
Endemic species of the Sierra Nevada de Santa
<collectingCountry id="26F1A4A5FFD0FF8B1F31FD5C4ED2FE64" box="[894,964,368,391]" name="Malta" pageId="10" pageNumber="104">Marta</collectingCountry>
,
<collectingCountry id="26F1A4A5FFD0FF8B1F80FD43492BFE6B" box="[975,1085,367,392]" name="Colombia" pageId="10" pageNumber="104">Colombia</collectingCountry>
(
<bibRefCitation id="3A7799C4FFD0FF8B1803FD4349E7FE6B" author="Boucher" box="[1100,1265,367,392]" pageId="10" pageNumber="104" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
).
<emphasis id="6C923827FFD0FF8B1C88FDB84CAEFE4E" bold="true" box="[199,440,404,429]" pageId="10" pageNumber="104">Material examined.</emphasis>
<collectingCountry id="26F1A4A5FFD0FF8B1D8AFDB84F4BFE4F" box="[453,605,404,428]" name="Colombia" pageId="10" pageNumber="104">COLOMBIA</collectingCountry>
, MAGDALENA, Sierra Nevada de Santa
<collectingCountry id="26F1A4A5FFD0FF8B182BFDB949BCFE4F" box="[1124,1194,405,428]" name="Malta" pageId="10" pageNumber="104">Marta</collectingCountry>
, Sector Bella Vista,
<geoCoordinate id="3BD282F2FFD0FF8B1CD8FD954C0FFE32" box="[151,281,439,465]" direction="north" orientation="latitude" pageId="10" pageNumber="104" precision="15" value="11.09611">11°05'46''N</geoCoordinate>
<geoCoordinate id="3BD282F2FFD0FF8B1D6FFD954CA8FE32" box="[288,446,439,465]" direction="west" orientation="longitude" pageId="10" pageNumber="104" precision="1" value="-74.081024">74°04'51.7''W</geoCoordinate>
,
<quantity id="991E49D0FFD0FF8B1D85FD944F36FE33" box="[458,544,440,465]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="10" pageNumber="104" unit="m" value="1560.0">1560 m</quantity>
,
<date id="2A58C2F5FFD0FF8B1E63FD944FB0FE32" box="[556,678,440,465]" pageId="10" pageNumber="104" value="2007-04-28">28.iv.2007</date>
. L. Jiménez &amp; A. Mejía (
<specimenCount id="48E02FBCFFD0FF8B1F8DFD954976FE32" box="[962,1120,440,465]" pageId="10" pageNumber="104" type="generic">12 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,128 @@
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<mods:title id="4847C74E3B6ECD092024263088760FFE">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<mods:date id="DE0D0797394D76CBE40F9B1C08AC489B">2015</mods:date>
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<treatment id="D64F5523FFD8FF831CD8F9384863F870" ID-DOI="http://doi.org/10.5281/zenodo.5618096" ID-GBIF-Taxon="119597874" ID-Zenodo-Dep="5618096" LSID="urn:lsid:plazi:treatment:D64F5523FFD8FF831CD8F9384863F870" httpUri="http://treatment.plazi.org/id/D64F5523FFD8FF831CD8F9384863F870" lastPageNumber="96" pageId="2" pageNumber="96">
<subSubSection id="16FCB7BEFFD8FF831CD8F938483EF93E" pageId="2" pageNumber="96" type="nomenclature">
<paragraph id="5E59E435FFD8FF831CD8F9384F89FACC" blockId="2.[151,671,1300,1360]" box="[151,671,1300,1327]" pageId="2" pageNumber="96">
<heading id="05115359FFD8FF831CD8F9384F89FACC" bold="true" box="[151,671,1300,1327]" fontSize="11" level="1" pageId="2" pageNumber="96" reason="1">
<emphasis id="6C923827FFD8FF831CD8F9384F89FACC" bold="true" box="[151,671,1300,1327]" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD8F9384DEAFACD" box="[151,252,1300,1326]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFD8FF831CD8F9384DEAFACD" bold="true" box="[151,252,1300,1326]" italics="true" pageId="2" pageNumber="96">Veturius</emphasis>
</taxonomicName>
(
<emphasis id="6C923827FFD8FF831D42F9384C79FACD" bold="true" box="[269,367,1300,1326]" italics="true" pageId="2" pageNumber="96">Vetur ius</emphasis>
)
<taxonomicName id="99E69FB6FFD8FF831DCCF9384F89FACC" authority="Weber, 1801" authorityName="Weber" authorityYear="1801" box="[387,671,1300,1327]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="6C923827FFD8FF831DCCF9384CFDFACD" bold="true" box="[387,491,1300,1326]" italics="true" pageId="2" pageNumber="96">assimilis</emphasis>
(Weber, 1801)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="5E59E435FFD8FF831CD8F91B4C88FAB3" blockId="2.[151,671,1300,1360]" box="[151,414,1335,1360]" pageId="2" pageNumber="96">
(
<figureCitation id="C6DDF8B0FFD8FF831CD0F91B4DFFFAB3" box="[159,233,1335,1360]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
b, 2b, 3b, 4b, 7)
</paragraph>
<paragraph id="5E59E435FFD8FF831CD8F953483EF93E" blockId="2.[151,1436,1407,1939]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831CD8F9534C24FA7B" bold="true" box="[151,306,1407,1432]" pageId="2" pageNumber="96">Third instar.</emphasis>
<emphasis id="6C923827FFD8FF831D74F9534C63FA7B" box="[315,373,1407,1432]" italics="true" pageId="2" pageNumber="96">Body</emphasis>
: length 35.0 mm, width (prothorax) 7.0 mm; with tiny, stout setae; prosternal setae present; presence of other setae in the lateroventral part of the abdominal segments IIX. Primary setal pattern: 3 HPA, 0 PD, 3 PSL, 0 MSD, 2 MSL, 0 MTD, 2 MTL, 1 TM, 0 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFD8FF831F96F9EB4939FA03" box="[985,1071,1479,1504]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="2" pageNumber="96" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
). The TSM setae present only in the abdominal segments VIIIIX.
<emphasis id="6C923827FFD8FF831E5AF9C04F47F9E6" box="[533,593,1516,1541]" italics="true" pageId="2" pageNumber="96">Head</emphasis>
: without uniform pile of short, golden setae; epicranial suture short; frontal sutures extend to the base of the antennae; width of head capsule 4.0 mm (
<figureCitation id="C6DDF8B0FFD8FF83186FFA234972F9CB" box="[1056,1124,1551,1576]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="2" pageNumber="96">Fig. 1</figureCitation>
b). Mandibles with microasperities in the ventral base. Maxillary stipes with 18 micro-conical projections (
<figureCitation id="C6DDF8B0FFD8FF83181BFA18498DF9AE" box="[1108,1179,1588,1613]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="2" pageNumber="96">Fig. 2</figureCitation>
b).
<emphasis id="6C923827FFD8FF83188BFA1949EFF9AF" box="[1220,1273,1589,1612]" italics="true" pageId="2" pageNumber="96">Legs</emphasis>
: pars stridens (mesocoxae) length 793.04 µm, width 495.63 µm, distance into striate 10.34 µm, 51 striae (
<figureCitation id="C6DDF8B0FFD8FF8318D8FA7B49CDF993" box="[1175,1243,1623,1648]" captionStart="FIGURE 7" captionStartId="11.[151,250,913,935]" captionTargetBox="[151,1436,267,891]" captionTargetId="figure@11.[151,1436,265,892]" captionTargetPageId="11" captionText="FIGURE 7. Scanning electronic micrograph of the mesocoxae of Veturius assimilis showing the pars stridens (ps). (a) View general of the mesocoxa. (b) High magnification of the ps in the lateral region. (c) High magnification of the ps in the central region." httpUri="https://zenodo.org/record/232517/files/figure.png" pageId="2" pageNumber="96">Fig. 7</figureCitation>
); mesocoxae and metacoxae with 2 internal setae. Metathoracic legs with some short medial and posterior setae and micro-asperities in the posterior areas; anterior margin with 6 teeth, I and IIIVI well developed and almost the same size, II is the smallest tooth (
<figureCitation id="C6DDF8B0FFD8FF831D0CFAE84C93F93E" box="[323,389,1732,1757]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="2" pageNumber="96">Fig. 3</figureCitation>
b).
<emphasis id="6C923827FFD8FF831DE5FAE84F02F93E" box="[426,532,1732,1757]" italics="true" pageId="2" pageNumber="96">Abdomen</emphasis>
: raster with setae; anal ring with 18 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFD8FF831887FAE8481AF93E" box="[1224,1292,1732,1757]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="2" pageNumber="96">Fig. 4</figureCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD8FF831C88FACB4F75F8AB" pageId="2" pageNumber="96" type="distribution">
<paragraph id="5E59E435FFD8FF831C88FACB4F75F8AB" blockId="2.[151,1436,1407,1939]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831C88FACB4C76F8E3" bold="true" box="[199,352,1767,1792]" pageId="2" pageNumber="96">Distribution.</emphasis>
States of Minas Gerais, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, Rio Grande do Sul,
<collectingCountry id="26F1A4A5FFD8FF831CD8FB204DCDF8C6" box="[151,219,1804,1829]" name="Brazil" pageId="2" pageNumber="96">Brazil</collectingCountry>
and Alto Paraná,
<collectingCountry id="26F1A4A5FFD8FF831DFFFB214F0EF8C6" box="[432,536,1805,1829]" name="Paraguay" pageId="2" pageNumber="96">Paraguay</collectingCountry>
(
<bibRefCitation id="3A7799C4FFD8FF831E65FB204FC7F8C6" author="Boucher" box="[554,721,1804,1829]" pageId="2" pageNumber="96" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
). The material examined confirms their presence in Misiones,
<collectingCountry id="26F1A4A5FFD8FF831CD8FB034C10F8AB" box="[151,262,1839,1864]" name="Argentina" pageId="2" pageNumber="96">Argentina</collectingCountry>
(
<bibRefCitation id="3A7799C4FFD8FF831D5AFB034F41F8AB" author="Jimenez-Ferbans" box="[277,599,1839,1864]" pageId="2" pageNumber="96" refString="Jimenez-Ferbans, L., Reyes-Castillo, P., Schuster, J. C. &amp; Salazar-Nino, K. (2013) A checklist and key for the identification of bess beetles (Coleoptera: Passalidae) of Argentina. Zootaxa, 3701 (2), 192 - 206. http: // dx. doi. org / 10.11646 / zootaxa. 3701.2.4" type="journal article" year="2013">
Jiménez-Ferbans
<emphasis id="6C923827FFD8FF831D94FB1D4F02F8AB" box="[475,532,1839,1864]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD8FF831C88FB794863F870" pageId="2" pageNumber="96" type="materials_examined">
<paragraph id="5E59E435FFD8FF831C88FB794863F870" blockId="2.[151,1436,1407,1939]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831C88FB794CD3F88D" bold="true" box="[199,453,1877,1902]" pageId="2" pageNumber="96">Material examined.</emphasis>
<collectingCountry id="26F1A4A5FFD8FF831DAFFB794F90F88E" box="[480,646,1877,1901]" name="Argentina" pageId="2" pageNumber="96">ARGENTINA</collectingCountry>
, MISIONES, Andrecito, Bio-Reserva Karadyá,
<geoCoordinate id="3BD282F2FFD8FF831947FB79488AF88E" box="[1288,1436,1875,1902]" direction="south" orientation="latitude" pageId="2" pageNumber="96" precision="1" value="-25.861834">25°51'42.6''S</geoCoordinate>
<geoCoordinate id="3BD282F2FFD8FF831CD8FB564C23F870" box="[151,309,1912,1939]" direction="west" orientation="longitude" pageId="2" pageNumber="96" precision="1" value="-53.96164">53°57'41.9''W</geoCoordinate>
,
<quantity id="991E49D0FFD8FF831D0FFB564C9CF871" box="[320,394,1914,1938]" metricMagnitude="2" metricUnit="m" metricValue="3.88" pageId="2" pageNumber="96" unit="m" value="388.0">388 m</quantity>
,
<date id="2A58C2F5FFD8FF831DDAFB564F07F870" box="[405,529,1914,1939]" pageId="2" pageNumber="96" value="2012-03-11">11.iii.2012</date>
. Reyes-Castillo, Jiménez-Ferbans, Salazar-Niño, &amp; J. Moreno (
<specimenCount id="48E02FBCFFD8FF831896FB56487FF870" box="[1241,1385,1914,1939]" pageId="2" pageNumber="96" type="generic">4 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="341C12A3B408A02E4547404092C125F6" ID-DOI="10.11646/zootaxa.3925.1.6" ID-GBIF-Dataset="08e0d80e-3aae-411c-acb9-450bbb4ae133" ID-ISSN="1175-5326" ID-Zenodo-Dep="232510" ID-ZooBank="2249FA36-09F3-4A85-A79B-7564A187CCEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461000639597" checkinUser="plazi" docAuthor="Salazar-Niño, Karen &amp; Serrão, José Eduardo" docDate="2015" docId="D64F5523FFDBFF831CD8FAFF4F21FB21" docLanguage="en" docName="zt03925p108.pdf" docOrigin="Zootaxa 3925 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Veturius" docType="treatment" docVersion="8" lastPageNumber="96" masterDocId="2A762D5BFFDAFF811C4FFC2C4D16FFE3" masterDocTitle="Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)" masterLastPageNumber="108" masterPageNumber="94" pageNumber="95" updateTime="1698564453905" updateUser="plazi">
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<mods:title id="7DB6906B82B6A4CC361C180BED739D3F">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<mods:namePart id="A6073DB521A8EC537DB842C40D6EA638">Serrão, José Eduardo</mods:namePart>
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<treatment id="D64F5523FFDBFF831CD8FAFF4F21FB21" ID-DOI="http://doi.org/10.5281/zenodo.5618094" ID-GBIF-Taxon="126095364" ID-Zenodo-Dep="5618094" LSID="urn:lsid:plazi:treatment:D64F5523FFDBFF831CD8FAFF4F21FB21" httpUri="http://treatment.plazi.org/id/D64F5523FFDBFF831CD8FAFF4F21FB21" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="95">
<subSubSection id="16FCB7BEFFDBFF831CD8FAFF497BFBE8" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="95" type="nomenclature">
<paragraph id="5E59E435FFDBFF801CD8FAFF4F92F90E" blockId="1.[151,644,1747,1807]" box="[151,644,1747,1773]" pageId="1" pageNumber="95">
<heading id="05115359FFDBFF801CD8FAFF4F92F90E" bold="true" box="[151,644,1747,1773]" fontSize="11" level="1" pageId="1" pageNumber="95" reason="1">
<emphasis id="6C923827FFDBFF801CD8FAFF4F92F90E" bold="true" box="[151,644,1747,1773]" pageId="1" pageNumber="95">
<taxonomicName id="99E69FB6FFDBFF801CD8FAFF4DEAF90E" box="[151,252,1747,1773]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFDBFF801CD8FAFF4DEAF90E" bold="true" box="[151,252,1747,1773]" italics="true" pageId="1" pageNumber="95">Veturius</emphasis>
</taxonomicName>
(
<emphasis id="6C923827FFDBFF801D42FAFF4C79F90E" bold="true" box="[269,367,1747,1773]" italics="true" pageId="1" pageNumber="95">Vetur ius</emphasis>
)
<taxonomicName id="99E69FB6FFDBFF801DCCFAFF4F92F90E" authority="Kuwert, 1898" authorityName="Kuwert" authorityYear="1898" box="[387,644,1747,1773]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="aspina">
<emphasis id="6C923827FFDBFF801DCCFAFF4CC5F90E" bold="true" box="[387,467,1747,1773]" italics="true" pageId="1" pageNumber="95">aspina</emphasis>
Kuwert, 1898
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="5E59E435FFDBFF801CD8FADA4CB2F8EC" blockId="1.[151,644,1747,1807]" box="[151,420,1782,1807]" pageId="1" pageNumber="95">
(
<figureCitation id="C6DDF8B0FFDBFF801CD0FADA4DFFF8EC" box="[159,233,1782,1807]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
a, 2a, 3a, 4a, 9a)
</paragraph>
<paragraph id="5E59E435FFDBFF801CD8FB12488AF804" blockId="1.[151,1436,1854,2023]" pageId="1" pageNumber="95">
<emphasis id="6C923827FFDBFF801CD8FB124C27F8B4" bold="true" box="[151,305,1854,1879]" pageId="1" pageNumber="95">Third instar.</emphasis>
<emphasis id="6C923827FFDBFF801D75FB124C62F8B4" box="[314,372,1854,1879]" italics="true" pageId="1" pageNumber="95">Body</emphasis>
: length 60.063.0 mm, width (prothorax) 12.0 mm; covered with uniform pile of fine, short setae (
<figureCitation id="C6DDF8B0FFDBFF801C90FB4E4C35F898" box="[223,291,1890,1915]" captionStart="FIGURE 9" captionStartId="12.[151,250,1377,1399]" captionTargetBox="[151,1436,193,1355]" captionTargetId="figure@12.[151,1436,193,1356]" captionTargetPageId="12" captionText="FIGURE 9. Light micrographs of the abdominal segment V (laterodorsal view), showing the setae of the body. (a) Uniform pile of fine short setae (Veturius aspina), with detail of the setae. (b) Tiny, stout setae (V. simillimus), with detail of the setae. (c) Tiny, stout setae (V. impressus), with detail of the setae. Scale 0.1 mm." httpUri="https://zenodo.org/record/232519/files/figure.png" pageId="1" pageNumber="95">Fig. 9</figureCitation>
a); prosternal setae absent. Primary setal pattern: 45 HPA, 1 PD, 4 PSL, 1 MSD, 23 MSL, 1 MTD, 23 MTL, 1 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFDBFF801E1DFBAA4FBFF87C" box="[594,681,1926,1951]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="1" pageNumber="95" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
).
<emphasis id="6C923827FFDBFF801EF2FBAA4FECF87C" box="[701,762,1926,1951]" italics="true" pageId="1" pageNumber="95">Head</emphasis>
: covered with uniform pile of short, golden setae; epicranial suture short; frontal sutures extended to the base of the antennae; width of head capsule 7.08.0 mm (
<figureCitation id="C6DDF8B0FFDBFF801975FB86486AF820" box="[1338,1404,1962,1987]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="1" pageNumber="95">Fig. 1</figureCitation>
a). Mandibles without micro-asperities on the ventral base. Maxillary stipes with 13 micro-conical projections (
<figureCitation id="C6DDF8B0FFDBFF801920FBE2488AF804" box="[1391,1436,1998,2023]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="1" pageNumber="95">Fig.</figureCitation>
</paragraph>
<paragraph id="5E59E435FFD8FF831CD8FCB44F2BFEFF" blockId="2.[151,1436,151,285]" pageId="2" pageNumber="96">
2a).
<emphasis id="6C923827FFD8FF831C83FCB54C17FF53" box="[204,257,153,176]" italics="true" pageId="2" pageNumber="96">Legs</emphasis>
: pars stridens (mesocoxae) length 1097.80 µm, width 737.89 µm, distance into striate 20.12 µm, 41 striae; mesocoxae and metacoxae with 2 internal setae. Metathoracic legs covered with short setae and microasperities in medial and posterior areas; anterior margin with 5 well developed teeth, I is the largest tooth, IIV gradually decreasing in size (
<figureCitation id="C6DDF8B0FFD8FF831D91FD284F36FEFF" box="[478,544,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="2" pageNumber="96">Fig. 3</figureCitation>
a).
</paragraph>
<caption id="0A99B4BDFFD8FF831CD8FD604CFDFE60" ID-Table-UUID="0A99B4BDFFD8FF831CD8FD604CFDFE60" httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="2" pageNumber="96" targetBox="[159,1407,414,850]" targetIsTable="true" targetPageId="2">
<paragraph id="5E59E435FFD8FF831CD8FD604CFDFE60" blockId="2.[151,1435,332,387]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831CD8FD604C1FFE80" bold="true" box="[151,265,332,355]" pageId="2" pageNumber="96">TABLE 2.</emphasis>
Number of primary setae of larvae of
<taxonomicName id="99E69FB6FFD8FF831EEDFD604FEFFE80" box="[674,761,332,355]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFD8FF831EEDFD604FEFFE80" box="[674,761,332,355]" italics="true" pageId="2" pageNumber="96">Veturius</emphasis>
</taxonomicName>
species (
<taxonomicName id="99E69FB6FFD8FF831F15FD604EC3FE80" box="[858,981,332,355]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="99E69FB6FFD8FF831F91FD604944FE80" box="[990,1106,332,355]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
). Numbers in parentheses refer to character states less frequent.
</paragraph>
</caption>
<paragraph id="5E59E435FFD8FF831D20FDB24843FCB1" pageId="2" pageNumber="96">
<table id="2CE61695FFD8007E1CD0FDB24869FCB1" box="[159,1407,414,850]" gridcols="11" gridrows="11" pageId="2" pageNumber="96">
<tr id="E0D6E677FFD8007E1CD0FDB24869FE50" box="[159,1407,414,435]" gridrow="0" pageId="2" pageNumber="96" rowspan-0="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="A3078F0BFFD8007E1D20FDB24F55FE50" box="[367,579,414,435]" gridcol="1" gridrow="0" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831D20FDB24F0EFE50" bold="true" box="[367,536,414,435]" pageId="2" pageNumber="96">SETAL GROUP</emphasis>
*
</th>
</tr>
<tr id="E0D6E677FFD8007E1CD0FDE44869FE3D" box="[159,1407,456,478]" gridrow="1" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FDE44C2FFE3D" box="[159,313,456,478]" gridcol="0" gridrow="1" pageId="2" pageNumber="96">SPECIES</th>
<td id="A3078F0BFFD8007E1D20FDE44F55FE3D" box="[367,579,456,478]" gridcol="1" gridrow="1" pageId="2" pageNumber="96">HPA PD PSL</td>
<td id="A3078F0BFFD8007E1E24FDE44FB4FE3D" box="[619,674,456,478]" gridcol="2" gridrow="1" pageId="2" pageNumber="96">MSD</td>
<td id="A3078F0BFFD8007E1E86FDE44E1AFE3D" box="[713,780,456,478]" gridcol="3" gridrow="1" pageId="2" pageNumber="96">MSL</td>
<td id="A3078F0BFFD8007E1F7BFDE44E7AFE3D" box="[820,876,456,478]" gridcol="4" gridrow="1" pageId="2" pageNumber="96">MTD</td>
<td id="A3078F0BFFD8007E1FDAFDE44EDCFE3D" box="[917,970,456,478]" gridcol="5" gridrow="1" pageId="2" pageNumber="96">MTL</td>
<td id="A3078F0BFFD8007E1FBAFDE44909FE3D" box="[1013,1055,456,478]" gridcol="6" gridrow="1" pageId="2" pageNumber="96">TM</td>
<td id="A3078F0BFFD8007E180CFDE44961FE3D" box="[1091,1143,456,478]" gridcol="7" gridrow="1" pageId="2" pageNumber="96">TSM</td>
<td id="A3078F0BFFD8007E18EFFDE449A8FE3D" box="[1184,1214,456,478]" gridcol="8" gridrow="1" pageId="2" pageNumber="96">TL</td>
<td id="A3078F0BFFD8007E18AAFDE44805FE3D" box="[1253,1299,456,478]" gridcol="9" gridrow="1" pageId="2" pageNumber="96">AV9</td>
<td id="A3078F0BFFD8007E1974FDE44869FE3D" box="[1339,1407,456,478]" gridcol="10" gridrow="1" pageId="2" pageNumber="96">AR</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FDDD4869FDE4" box="[159,1407,497,519]" gridrow="2" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FDDD4C2FFDE4" box="[159,313,497,519]" gridcol="0" gridrow="2" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FDDE4DEAFDE4" box="[159,252,498,519]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="aspina">
<emphasis id="6C923827FFD8FF831CD0FDDE4DEAFDE4" box="[159,252,498,519]" italics="true" pageId="2" pageNumber="96">V. aspina</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FDDD4F55FDE4" box="[367,579,497,519]" gridcol="1" gridrow="2" pageId="2" pageNumber="96">45 1(2) 4</td>
<td id="A3078F0BFFD8007E1E24FDDD4FB4FDE4" box="[619,674,497,519]" gridcol="2" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1E86FDDD4E1AFDE4" box="[713,780,497,519]" gridcol="3" gridrow="2" pageId="2" pageNumber="96">23(4)</td>
<td id="A3078F0BFFD8007E1F7BFDDD4E7AFDE4" box="[820,876,497,519]" gridcol="4" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1FDAFDDD4EDCFDE4" box="[917,970,497,519]" gridcol="5" gridrow="2" pageId="2" pageNumber="96">23</td>
<td id="A3078F0BFFD8007E1FBAFDDD4909FDE4" box="[1013,1055,497,519]" gridcol="6" gridrow="2" pageId="2" pageNumber="96">1(2)</td>
<td id="A3078F0BFFD8007E180CFDDD4961FDE4" box="[1091,1143,497,519]" gridcol="7" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFDDD49A8FDE4" box="[1184,1214,497,519]" gridcol="8" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFDDD4805FDE4" box="[1253,1299,497,519]" gridcol="9" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FDDD4869FDE4" box="[1339,1407,497,519]" gridcol="10" gridrow="2" pageId="2" pageNumber="96">2022</td>
</tr>
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<th id="A3078F0BFFD8007E1CD0FE364C2FFDD3" box="[159,313,538,560]" gridcol="0" gridrow="3" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FE374C07FDD3" box="[159,273,538,560]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="6C923827FFD8FF831CD0FE374C07FDD3" box="[159,273,538,560]" italics="true" pageId="2" pageNumber="96">V. assimilis</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FE364F55FDD3" box="[367,579,538,560]" gridcol="1" gridrow="3" pageId="2" pageNumber="96">3 0 3</td>
<td id="A3078F0BFFD8007E1E24FE364FB4FDD3" box="[619,674,538,560]" gridcol="2" gridrow="3" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FE364E1AFDD3" box="[713,780,538,560]" gridcol="3" gridrow="3" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFE364E7AFDD3" box="[820,876,538,560]" gridcol="4" gridrow="3" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFE364EDCFDD3" box="[917,970,538,560]" gridcol="5" gridrow="3" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFE364909FDD3" box="[1013,1055,538,560]" gridcol="6" gridrow="3" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFE364961FDD3" box="[1091,1143,538,560]" gridcol="7" gridrow="3" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E18EFFE3649A8FDD3" box="[1184,1214,538,560]" gridcol="8" gridrow="3" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFE364805FDD3" box="[1253,1299,538,560]" gridcol="9" gridrow="3" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FE364869FDD3" box="[1339,1407,538,560]" gridcol="10" gridrow="3" pageId="2" pageNumber="96">18</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FE684869FDB9" box="[159,1407,580,602]" gridrow="4" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FE684C2FFDB9" box="[159,313,580,602]" gridcol="0" gridrow="4" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FE694C36FDB9" box="[159,288,580,602]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="simillimus">
<emphasis id="6C923827FFD8FF831CD0FE694C36FDB9" box="[159,288,580,602]" italics="true" pageId="2" pageNumber="96">V. simillimus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FE684F55FDB9" box="[367,579,580,602]" gridcol="1" gridrow="4" pageId="2" pageNumber="96">2 1 3</td>
<td id="A3078F0BFFD8007E1E24FE684FB4FDB9" box="[619,674,580,602]" gridcol="2" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1E86FE684E1AFDB9" box="[713,780,580,602]" gridcol="3" gridrow="4" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFE684E7AFDB9" box="[820,876,580,602]" gridcol="4" gridrow="4" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFE684EDCFDB9" box="[917,970,580,602]" gridcol="5" gridrow="4" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFE684909FDB9" box="[1013,1055,580,602]" gridcol="6" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFE684961FDB9" box="[1091,1143,580,602]" gridcol="7" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFE6849A8FDB9" box="[1184,1214,580,602]" gridcol="8" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFE684805FDB9" box="[1253,1299,580,602]" gridcol="9" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FE684869FDB9" box="[1339,1407,580,602]" gridcol="10" gridrow="4" pageId="2" pageNumber="96">19</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FE414869FD60" box="[159,1407,621,643]" gridrow="5" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FE414C2FFD60" box="[159,313,621,643]" gridcol="0" gridrow="5" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FE424C2FFD60" box="[159,313,621,643]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="sinuatocollis">
<emphasis id="6C923827FFD8FF831CD0FE424C2FFD60" box="[159,313,621,643]" italics="true" pageId="2" pageNumber="96">V. sinuatocollis</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FE414F55FD60" box="[367,579,621,643]" gridcol="1" gridrow="5" pageId="2" pageNumber="96">45 1 3</td>
<td id="A3078F0BFFD8007E1E24FE414FB4FD60" box="[619,674,621,643]" gridcol="2" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1E86FE414E1AFD60" box="[713,780,621,643]" gridcol="3" gridrow="5" pageId="2" pageNumber="96">23</td>
<td id="A3078F0BFFD8007E1F7BFE414E7AFD60" box="[820,876,621,643]" gridcol="4" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1FDAFE414EDCFD60" box="[917,970,621,643]" gridcol="5" gridrow="5" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFE414909FD60" box="[1013,1055,621,643]" gridcol="6" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFE414961FD60" box="[1091,1143,621,643]" gridcol="7" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFE4149A8FD60" box="[1184,1214,621,643]" gridcol="8" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFE414805FD60" box="[1253,1299,621,643]" gridcol="9" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FE414869FD60" box="[1339,1407,621,643]" gridcol="10" gridrow="5" pageId="2" pageNumber="96">1721</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FEBA4869FD4E" box="[159,1407,662,685]" gridrow="6" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FEBA4C2FFD4E" box="[159,313,662,685]" gridcol="0" gridrow="6" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FEB44C1AFD4E" box="[159,268,664,685]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="6C923827FFD8FF831CD0FEB44C1AFD4E" box="[159,268,664,685]" italics="true" pageId="2" pageNumber="96">V. sinuatus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FEBA4F55FD4E" box="[367,579,662,685]" gridcol="1" gridrow="6" pageId="2" pageNumber="96">3 0 3</td>
<td id="A3078F0BFFD8007E1E24FEBA4FB4FD4E" box="[619,674,662,685]" gridcol="2" gridrow="6" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FEBA4E1AFD4E" box="[713,780,662,685]" gridcol="3" gridrow="6" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFEBA4E7AFD4E" box="[820,876,662,685]" gridcol="4" gridrow="6" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFEBA4EDCFD4E" box="[917,970,662,685]" gridcol="5" gridrow="6" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFEBA4909FD4E" box="[1013,1055,662,685]" gridcol="6" gridrow="6" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFEBA4961FD4E" box="[1091,1143,662,685]" gridcol="7" gridrow="6" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFEBA49A8FD4E" box="[1184,1214,662,685]" gridcol="8" gridrow="6" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFEBA4805FD4E" box="[1253,1299,662,685]" gridcol="9" gridrow="6" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FEBA4869FD4E" box="[1339,1407,662,685]" gridcol="10" gridrow="6" pageId="2" pageNumber="96">20</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FEEC4869FD35" box="[159,1407,704,726]" gridrow="7" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FEEC4C2FFD35" box="[159,313,704,726]" gridcol="0" gridrow="7" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FEED4C12FD35" box="[159,260,705,726]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="6C923827FFD8FF831CD0FEED4C12FD35" box="[159,260,705,726]" italics="true" pageId="2" pageNumber="96">V. crassus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FEEC4F55FD35" box="[367,579,704,726]" gridcol="1" gridrow="7" pageId="2" pageNumber="96">3(4) 0 3</td>
<td id="A3078F0BFFD8007E1E24FEEC4FB4FD35" box="[619,674,704,726]" gridcol="2" gridrow="7" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FEEC4E1AFD35" box="[713,780,704,726]" gridcol="3" gridrow="7" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFEEC4E7AFD35" box="[820,876,704,726]" gridcol="4" gridrow="7" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFEEC4EDCFD35" box="[917,970,704,726]" gridcol="5" gridrow="7" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFEEC4909FD35" box="[1013,1055,704,726]" gridcol="6" gridrow="7" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFEEC4961FD35" box="[1091,1143,704,726]" gridcol="7" gridrow="7" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFEEC49A8FD35" box="[1184,1214,704,726]" gridcol="8" gridrow="7" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFEEC4805FD35" box="[1253,1299,704,726]" gridcol="9" gridrow="7" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FEEC4869FD35" box="[1339,1407,704,726]" gridcol="10" gridrow="7" pageId="2" pageNumber="96">18(16)</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FEC54869FD1C" box="[159,1407,745,767]" gridrow="8" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FEC54C2FFD1C" box="[159,313,745,767]" gridcol="0" gridrow="8" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FEC64C0BFD1C" box="[159,285,746,767]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="6C923827FFD8FF831CD0FEC64C0BFD1C" box="[159,285,746,767]" italics="true" pageId="2" pageNumber="96">V. impressus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FEC54F55FD1C" box="[367,579,745,767]" gridcol="1" gridrow="8" pageId="2" pageNumber="96">3 0 3</td>
<td id="A3078F0BFFD8007E1E24FEC54FB4FD1C" box="[619,674,745,767]" gridcol="2" gridrow="8" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FEC54E1AFD1C" box="[713,780,745,767]" gridcol="3" gridrow="8" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFEC54E7AFD1C" box="[820,876,745,767]" gridcol="4" gridrow="8" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFEC54EDCFD1C" box="[917,970,745,767]" gridcol="5" gridrow="8" pageId="2" pageNumber="96">12</td>
<td id="A3078F0BFFD8007E1FBAFEC54909FD1C" box="[1013,1055,745,767]" gridcol="6" gridrow="8" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFEC54961FD1C" box="[1091,1143,745,767]" gridcol="7" gridrow="8" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFEC549A8FD1C" box="[1184,1214,745,767]" gridcol="8" gridrow="8" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFEC54805FD1C" box="[1253,1299,745,767]" gridcol="9" gridrow="8" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FEC54869FD1C" box="[1339,1407,745,767]" gridcol="10" gridrow="8" pageId="2" pageNumber="96">1516</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FF3E4869FCCB" box="[159,1407,786,808]" gridrow="9" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FF3E4C2FFCCB" box="[159,313,786,808]" gridcol="0" gridrow="9" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FF3F4C0EFCCB" box="[159,280,786,808]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="6C923827FFD8FF831CD0FF3F4C0EFCCB" box="[159,280,786,808]" italics="true" pageId="2" pageNumber="96">V. oberthuri</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FF3E4F55FCCB" box="[367,579,786,808]" gridcol="1" gridrow="9" pageId="2" pageNumber="96">10 0 3</td>
<td id="A3078F0BFFD8007E1E24FF3E4FB4FCCB" box="[619,674,786,808]" gridcol="2" gridrow="9" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FF3E4E1AFCCB" box="[713,780,786,808]" gridcol="3" gridrow="9" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFF3E4E7AFCCB" box="[820,876,786,808]" gridcol="4" gridrow="9" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFF3E4EDCFCCB" box="[917,970,786,808]" gridcol="5" gridrow="9" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFF3E4909FCCB" box="[1013,1055,786,808]" gridcol="6" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFF3E4961FCCB" box="[1091,1143,786,808]" gridcol="7" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFF3E49A8FCCB" box="[1184,1214,786,808]" gridcol="8" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFF3E4805FCCB" box="[1253,1299,786,808]" gridcol="9" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FF3E4869FCCB" box="[1339,1407,786,808]" gridcol="10" gridrow="9" pageId="2" pageNumber="96">18</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FF104869FCB1" box="[159,1407,828,850]" gridrow="10" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FF104C2FFCB1" box="[159,313,828,850]" gridcol="0" gridrow="10" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FF114C33FCB1" box="[159,293,829,850]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="negroensis">
<emphasis id="6C923827FFD8FF831CD0FF114C33FCB1" box="[159,293,829,850]" italics="true" pageId="2" pageNumber="96">V. negroensis</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FF104F55FCB1" box="[367,579,828,850]" gridcol="1" gridrow="10" pageId="2" pageNumber="96">2 0 3</td>
<td id="A3078F0BFFD8007E1E24FF104FB4FCB1" box="[619,674,828,850]" gridcol="2" gridrow="10" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FF104E1AFCB1" box="[713,780,828,850]" gridcol="3" gridrow="10" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFF104E7AFCB1" box="[820,876,828,850]" gridcol="4" gridrow="10" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFF104EDCFCB1" box="[917,970,828,850]" gridcol="5" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1FBAFF104909FCB1" box="[1013,1055,828,850]" gridcol="6" gridrow="10" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFF104961FCB1" box="[1091,1143,828,850]" gridcol="7" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFF1049A8FCB1" box="[1184,1214,828,850]" gridcol="8" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFF104805FCB1" box="[1253,1299,828,850]" gridcol="9" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FF104869FCB1" box="[1339,1407,828,850]" gridcol="10" gridrow="10" pageId="2" pageNumber="96">14</td>
</tr>
</table>
</paragraph>
<paragraph id="5E59E435FFD8FF831CD8FF5E4898FC2A" blockId="2.[151,1435,882,969]" pageId="2" pageNumber="96">* Post-antennal head setae (HPA), dorsal pronotal setae (PD), lateral pronotal shield setae (PSL), dorsal mesonotal setae (MSD), lateral mesonotal setae (MSL), dorsal metanotal setae (MTD), lateral metanotal setae (MTL), median tergal setae (TM), submedial tergal setae (TSM), lateral tergal setae (TL), ventral abdominal setae (AV), anal ring setae (AR).</paragraph>
<paragraph id="5E59E435FFD8FF831C88FFDE497BFBE8" blockId="2.[151,1436,1010,1218]" box="[199,1133,1010,1035]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831C88FFDE4C27FBE8" box="[199,305,1010,1035]" italics="true" pageId="2" pageNumber="96">Abdomen</emphasis>
: raster with setae; anal ring with 2022 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFD8FF831841FFDE4944FBE8" box="[1038,1106,1010,1035]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="2" pageNumber="96">Fig. 4</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD8FF831C88F83A4F21FB21" pageId="2" pageNumber="96" type="distribution">
<paragraph id="5E59E435FFD8FF831C88F83A4F21FB21" blockId="2.[151,1436,1010,1218]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831C88F83A4C76FBCC" bold="true" box="[199,352,1046,1071]" pageId="2" pageNumber="96">Distribution.</emphasis>
Species with geographical distribution from
<collectingCountry id="26F1A4A5FFD8FF831F15F83A4ED1FBCC" box="[858,967,1046,1071]" name="Honduras" pageId="2" pageNumber="96">Honduras</collectingCountry>
to
<collectingCountry id="26F1A4A5FFD8FF831FA5F83A4951FBCC" box="[1002,1095,1046,1071]" name="Ecuador" pageId="2" pageNumber="96">Ecuador</collectingCountry>
(
<bibRefCitation id="3A7799C4FFD8FF831819F83A49EDFBCC" author="Boucher" box="[1110,1275,1046,1071]" pageId="2" pageNumber="96" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
).
<emphasis id="6C923827FFD8FF831C88F8174CA7FBB7" bold="true" box="[199,433,1083,1108]" pageId="2" pageNumber="96">Material examined.</emphasis>
<collectingCountry id="26F1A4A5FFD8FF831DF7F8104F46FBB7" box="[440,592,1084,1108]" name="Colombia" pageId="2" pageNumber="96">COLOMBIA</collectingCountry>
, TOLIMA, Municipio Ibagué, vereda Potrerillo, Sur el Totumo,
<geoCoordinate id="3BD282F2FFD8FF831968F810488AFBB7" box="[1319,1436,1082,1108]" direction="north" orientation="latitude" pageId="2" pageNumber="96" precision="15" value="4.3891664">4°23'21''N</geoCoordinate>
<geoCoordinate id="3BD282F2FFD8FF831CD8F84C4C23FB9A" box="[151,309,1119,1145]" direction="west" orientation="longitude" pageId="2" pageNumber="96" precision="1" value="-75.21547">75°12'55.7''W</geoCoordinate>
,
<quantity id="991E49D0FFD8FF831D07F84D4CB1FB9B" box="[328,423,1121,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="96" unit="m" value="1040.0">1040 m</quantity>
,
<date id="2A58C2F5FFD8FF831DF5F84C4F31FB9A" box="[442,551,1120,1145]" pageId="2" pageNumber="96" value="2008-06-05">5.vi.2008</date>
. G. Amat, P. Reyes-Castillo, &amp; L. Jiménez (
<specimenCount id="48E02FBCFFD8FF831800F84C49F0FB9A" box="[1103,1254,1120,1145]" pageId="2" pageNumber="96" type="generic">5 specimens</specimenCount>
).
<collectingCountry id="26F1A4A5FFD8FF83194FF84D488EFB9A" box="[1280,1432,1121,1145]" name="Colombia" pageId="2" pageNumber="96">COLOMBIA</collectingCountry>
, ANTIOQUIA, Gómez Plata, vereda Vega Botero,
<quantity id="991E49D0FFD8FF831E98F8AA4E34FB7E" box="[727,802,1158,1182]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="2" pageNumber="96" unit="m" value="940.0">940 m</quantity>
,
<date id="2A58C2F5FFD8FF831F7FF8A94EA7FB7D" box="[816,945,1157,1182]" pageId="2" pageNumber="96" value="2011-07-12">12.vii.2011</date>
. Reyes-Castillo, Jiménez-Ferbans, Salazar- Niño, &amp; Amat-García (
<specimenCount id="48E02FBCFFD8FF831DD3F8854F3DFB21" box="[412,555,1193,1218]" pageId="2" pageNumber="96" type="generic">5 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,136 @@
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<mods:title id="332045797F953796BBC1F8C38E2D0149">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<treatment id="D64F5523FFDDFF861CD8FCBB4EC5FC46" ID-DOI="http://doi.org/10.5281/zenodo.5618098" ID-GBIF-Taxon="126095363" ID-Zenodo-Dep="5618098" LSID="urn:lsid:plazi:treatment:D64F5523FFDDFF861CD8FCBB4EC5FC46" httpUri="http://treatment.plazi.org/id/D64F5523FFDDFF861CD8FCBB4EC5FC46" lastPageNumber="101" pageId="7" pageNumber="101">
<subSubSection id="16FCB7BEFFDDFF861CD8FCBB496AFD0C" pageId="7" pageNumber="101" type="nomenclature">
<paragraph id="5E59E435FFDDFF861CD8FCBB4FA7FF51" blockId="7.[151,689,151,211]" box="[151,689,151,178]" pageId="7" pageNumber="101">
<heading id="05115359FFDDFF861CD8FCBB4FA7FF51" bold="true" box="[151,689,151,178]" fontSize="11" level="1" pageId="7" pageNumber="101" reason="1">
<emphasis id="6C923827FFDDFF861CD8FCBB4FA7FF51" bold="true" box="[151,689,151,178]" pageId="7" pageNumber="101">
<taxonomicName id="99E69FB6FFDDFF861CD8FCBB4DEAFF52" box="[151,252,151,177]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFDDFF861CD8FCBB4DEAFF52" bold="true" box="[151,252,151,177]" italics="true" pageId="7" pageNumber="101">Veturius</emphasis>
</taxonomicName>
(
<emphasis id="6C923827FFDDFF861D42FCBB4C79FF52" bold="true" box="[269,367,151,177]" italics="true" pageId="7" pageNumber="101">Vetur ius</emphasis>
)
<taxonomicName id="99E69FB6FFDDFF861DCCFCBB4FA7FF51" authority="Kuwert, 1891" authorityName="Kuwert" authorityYear="1891" box="[387,689,151,178]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="101" phylum="Arthropoda" rank="species" species="simillimus">
<emphasis id="6C923827FFDDFF861DCCFCBB4F17FF52" bold="true" box="[387,513,151,177]" italics="true" pageId="7" pageNumber="101">simillimus</emphasis>
Kuwert, 1891
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="5E59E435FFDDFF861CD8FC964CB3FF30" blockId="7.[151,689,151,211]" box="[151,421,186,211]" pageId="7" pageNumber="101">
(
<figureCitation id="C6DDF8B0FFDDFF861CD0FC964DFCFF30" box="[159,234,186,211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="7" pageNumber="101">Figs. 1</figureCitation>
c, 2c, 3c, 4c, 9b)
</paragraph>
<paragraph id="5E59E435FFDDFF861CD8FD2E4833FDBC" blockId="7.[151,1436,258,933]" pageId="7" pageNumber="101">
<emphasis id="6C923827FFDDFF861CD8FD2E4C3FFEF8" bold="true" box="[151,297,258,283]" pageId="7" pageNumber="101">Third instar</emphasis>
.
<emphasis id="6C923827FFDDFF861D76FD2E4C65FEF8" box="[313,371,258,283]" italics="true" pageId="7" pageNumber="101">Body</emphasis>
: length 46.0 mm, width (prothorax) 8.0 mm; with tiny, stout setae (
<figureCitation id="C6DDF8B0FFDDFF861830FD2E49D2FEF8" box="[1151,1220,258,283]" captionStart="FIGURE 9" captionStartId="12.[151,250,1377,1399]" captionTargetBox="[151,1436,193,1355]" captionTargetId="figure@12.[151,1436,193,1356]" captionTargetPageId="12" captionText="FIGURE 9. Light micrographs of the abdominal segment V (laterodorsal view), showing the setae of the body. (a) Uniform pile of fine short setae (Veturius aspina), with detail of the setae. (b) Tiny, stout setae (V. simillimus), with detail of the setae. (c) Tiny, stout setae (V. impressus), with detail of the setae. Scale 0.1 mm." httpUri="https://zenodo.org/record/232519/files/figure.png" pageId="7" pageNumber="101">Fig. 9</figureCitation>
b); prosternal setae absent. Primary setal pattern: 2 HPA, 1 PD, 3 PSL, 1 MSD, 2 MSL, 0 MTD, 2 MTL, 1 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFDDFF861CD1FD664DE3FE80" box="[158,245,330,355]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="7" pageNumber="101" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
). The TM is evident in the segments VIIIX, but in the others seems have a small and thicker setae in this position.
<emphasis id="6C923827FFDDFF861D4FFD424C2AFE64" box="[256,316,366,391]" italics="true" pageId="7" pageNumber="101">Head</emphasis>
: without uniform pile of short, golden setae; epicranial suture short; frontal sutures extended to the base of the antennae; width of head capsule 5.0 mm (
<figureCitation id="C6DDF8B0FFDDFF861F4CFDBE4E50FE48" box="[771,838,402,427]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="7" pageNumber="101">Fig. 1</figureCitation>
c). Mandibles with micro-asperities and setae in the ventral base. Maxillary stipes with 12 micro-conical projections (
<figureCitation id="C6DDF8B0FFDDFF861F34FD9A4ED7FE2C" box="[891,961,438,463]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="7" pageNumber="101">Fig. 2</figureCitation>
c).
<emphasis id="6C923827FFDDFF861FABFD9B490FFE2D" box="[996,1049,439,462]" italics="true" pageId="7" pageNumber="101">Legs</emphasis>
: pars stridens (mesocoxae) length 940.87 µm, width 741.91 µm, distance into striate 13.55 µm, 56 striae; mesocoxae and metacoxae with one internal seta. Metathoracic legs with some short medial and lateral setae (side of I tooth), and micro-asperities in the posterior areas; anterior margin with 6 teeth, I and IIIVI well developed and gradually decreasing in size, II is the smallest tooth (
<figureCitation id="C6DDF8B0FFDDFF861D0CFE6A4C90FDBC" box="[323,390,582,607]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="7" pageNumber="101">Fig. 3</figureCitation>
c).
<emphasis id="6C923827FFDDFF861DE7FE6A4F04FDBC" box="[424,530,582,607]" italics="true" pageId="7" pageNumber="101">Abdomen</emphasis>
: raster with setae; anal ring with 19 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFDDFF861889FE6A481CFDBC" box="[1222,1290,582,607]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="7" pageNumber="101">Fig. 4</figureCitation>
c).
</paragraph>
<paragraph id="5E59E435FFDDFF861C88FE46496AFD0C" blockId="7.[151,1436,258,933]" pageId="7" pageNumber="101">
<emphasis id="6C923827FFDDFF861C88FE464C69FD60" bold="true" box="[199,383,618,643]" pageId="7" pageNumber="101">Natural history</emphasis>
. Larvae were collected in third instar and reared in laboratory conditions (25+2 °C; 54+2% relative humidity; photoperiod 12 light: 8 dark hours) using pieces of decaying logs as food. The third instars to pre-pupal phase took an average of 60 days, pre-pupal phase took an average of three days, pupal phase took an average of 22 days. The complete sclerotization of the adults took an average of 42 days.
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFDDFF861C88FED649E1FCD4" pageId="7" pageNumber="101" type="distribution">
<paragraph id="5E59E435FFDDFF861C88FED649E1FCD4" blockId="7.[151,1436,258,933]" pageId="7" pageNumber="101">
<emphasis id="6C923827FFDDFF861C88FED64C76FCF0" bold="true" box="[199,352,762,787]" pageId="7" pageNumber="101">Distribution.</emphasis>
Endemic species from Brazilian Atlantic rainforest in the states of Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, and northern Rio Grande do Sul (
<bibRefCitation id="3A7799C4FFDDFF861809FF3249FDFCD4" author="Boucher" box="[1094,1259,798,823]" pageId="7" pageNumber="101" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFDDFF861C88FF6F4EC5FC46" pageId="7" pageNumber="101" type="materials_examined">
<paragraph id="5E59E435FFDDFF861C88FF6F4EC5FC46" blockId="7.[151,1436,258,933]" pageId="7" pageNumber="101">
<emphasis id="6C923827FFDDFF861C88FF6F4CADFCBF" bold="true" box="[199,443,835,860]" pageId="7" pageNumber="101">Material examined.</emphasis>
<collectingCountry id="26F1A4A5FFDDFF861D82FF684F23FCBF" box="[461,565,836,860]" name="Brazil" pageId="7" pageNumber="101">BRAZIL</collectingCountry>
,
<collectionCode id="38F77CF0FFDDFF861E04FF684FB3FCBF" box="[587,677,836,860]" pageId="7" pageNumber="101">MINAS</collectionCode>
GERAIS, Viçosa,
<collectingCountry id="26F1A4A5FFDDFF861FD5FF684EC2FCB8" box="[922,980,836,859]" name="Malta" pageId="7" pageNumber="101">Mata</collectingCountry>
do Paraíso,
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<geoCoordinate id="3BD282F2FFDDFF861941FF684881FCBF" box="[1294,1431,834,860]" direction="west" orientation="longitude" pageId="7" pageNumber="101" precision="15" value="-42.85861">42°51'31''W</geoCoordinate>
,
<date id="2A58C2F5FFDDFF861CD8FF444C10FC63" box="[151,262,872,896]" pageId="7" pageNumber="101" value="2013-10-11">11.x.2013</date>
. I. Salvatore, C. Ferraz, &amp; Salazar-Niño (
<specimenCount id="48E02FBCFFDDFF861EA5FF444E6AFC63" box="[746,892,871,896]" pageId="7" pageNumber="101" type="generic">4 specimens</specimenCount>
).
<collectingCountry id="26F1A4A5FFDDFF861FDEFF444EEFFC63" box="[913,1017,872,896]" name="Brazil" pageId="7" pageNumber="101">BRAZIL</collectingCountry>
,
<collectionCode id="38F77CF0FFDDFF861849FF444976FC63" box="[1030,1120,872,896]" pageId="7" pageNumber="101">MINAS</collectionCode>
GERAIS, Viçosa,
<collectingCountry id="26F1A4A5FFDDFF861972FF444861FC9C" box="[1341,1399,872,895]" name="Malta" pageId="7" pageNumber="101">Mata</collectingCountry>
do Paraíso,
<geoCoordinate id="3BD282F2FFDDFF861CB8FFA14C60FC46" box="[247,374,907,933]" direction="south" orientation="latitude" pageId="7" pageNumber="101" precision="15" value="-20.801943">20°48'07''S</geoCoordinate>
<geoCoordinate id="3BD282F2FFDDFF861D33FFA14F10FC46" box="[380,518,907,933]" direction="west" orientation="longitude" pageId="7" pageNumber="101" precision="15" value="-42.85861">42°51'31''W</geoCoordinate>
,
<date id="2A58C2F5FFDDFF861E5EFFA04F9DFC46" box="[529,651,908,933]" pageId="7" pageNumber="101" value="2014-04-25">25.iv.2014</date>
. Salazar-Niño (
<specimenCount id="48E02FBCFFDDFF861F77FFA14EDEFC46" box="[824,968,908,933]" pageId="7" pageNumber="101" type="generic">2 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="B63515F5E3B3897D8F2A1F44D99CD1BA">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<taxonomicName id="99E69FB6FFDDFF861CD8FFD44DEAFBF1" box="[151,252,1016,1042]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="101" phylum="Arthropoda" rank="genus">
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(
<emphasis id="6C923827FFDDFF861D42FFD44C79FBF1" bold="true" box="[269,367,1016,1042]" italics="true" pageId="7" pageNumber="101">Vetur ius</emphasis>
)
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<emphasis id="6C923827FFDDFF861DCCFFD44F0AFBF1" bold="true" box="[387,540,1016,1042]" italics="true" pageId="7" pageNumber="101">sinuatocollis</emphasis>
Kuwert, 1890
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(
<figureCitation id="C6DDF8B0FFDDFF861CD0F8304DFFFBD6" box="[159,233,1052,1077]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="7" pageNumber="101">Figs. 1</figureCitation>
d, 2d, 3d, 4d, 5)
</paragraph>
<paragraph id="5E59E435FFDDFF861CD8F8484929FA23" blockId="7.[151,1437,1124,1581]" pageId="7" pageNumber="101">
<emphasis id="6C923827FFDDFF861CD8F8484C3FFB9E" bold="true" box="[151,297,1124,1149]" pageId="7" pageNumber="101">Third instar</emphasis>
.
<emphasis id="6C923827FFDDFF861D75F8484C65FB9E" box="[314,371,1124,1149]" italics="true" pageId="7" pageNumber="101">Body</emphasis>
: length 40.0 mm, width (prothorax)
<quantity id="991E49D0FFDDFF861F51F8484E6CFB9F" box="[798,890,1124,1149]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="101" unit="mm" value="8.5">8.5 mm</quantity>
; covered with tiny, stout setae; prosternal setae present; presence of other setae in the lateroventral part of the abdominal segments IIX. Primary setal pattern: 45 HPA, 1 PD, 3 PSL, 1 MSD, 23 MSL, 1 MTD, 2 MTL, 1 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFDDFF8618C8F88049F7FB27" box="[1159,1249,1196,1221]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="7" pageNumber="101" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
).
<emphasis id="6C923827FFDDFF8618B7F8804822FB26" box="[1272,1332,1196,1221]" italics="true" pageId="7" pageNumber="101">Head</emphasis>
: without uniform pile of short, golden setae; epicranial suture short; frontal sutures extended to the base of the antennae; width of head capsule 6.0 mm (
<figureCitation id="C6DDF8B0FFDDFF861E48F8D84F5FFAEE" box="[519,585,1268,1293]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="7" pageNumber="101">Fig. 1</figureCitation>
d). Mandibles with micro-asperities and setae in the ventral base (
<figureCitation id="C6DDF8B0FFDDFF861908F8D84898FAEE" box="[1351,1422,1268,1293]" captionStart="FIGURE 5" captionStartId="8.[151,250,1607,1629]" captionTargetBox="[151,1436,774,1585]" captionTargetId="figure@8.[151,1436,774,1586]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electronic micrographs of the mandible of Veturius sinuatocollis. (a) Ventral view with micro-asperities and setae (circle) in the base of the mandible. (b) High magnification of encircled area of Fig. 5 a, showing detail of microasperities (white arrow) and setae (black arrow)." httpUri="https://zenodo.org/record/232515/files/figure.png" pageId="7" pageNumber="101">Fig. 5</figureCitation>
). Maxillary stipes with 16 micro-conical projections (
<figureCitation id="C6DDF8B0FFDDFF861EACF93B4E31FAD3" box="[739,807,1303,1328]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="7" pageNumber="101">Fig. 2</figureCitation>
d).
<emphasis id="6C923827FFDDFF861F03F9354E97FAD3" box="[844,897,1305,1328]" italics="true" pageId="7" pageNumber="101">Legs</emphasis>
: pars stridens (mesocoxae) length 1040.56 µm, width 951.52 µm, distance into striate 20.00 µm, 43 striae; mesocoxae and metacoxae with 2 internal setae. Metathoracic legs with some short medial and lateral setae and with tiny stout setae in the posterior areas; anterior margin with 6 teeth, I and IIIVI well developed and gradually decreasing in size, II is the smallest tooth (
<figureCitation id="C6DDF8B0FFDDFF861973F9A84868FA7E" box="[1340,1406,1412,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="7" pageNumber="101">Fig. 3</figureCitation>
d).
<emphasis id="6C923827FFDDFF861CD8F98B4C17FA23" box="[151,257,1447,1472]" italics="true" pageId="7" pageNumber="101">Abdomen</emphasis>
: raster with setae; anal ring with 1721 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFDDFF861F91F98B4934FA23" box="[990,1058,1447,1472]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="7" pageNumber="101">Fig. 4</figureCitation>
d).
</paragraph>
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<emphasis id="6C923827FFDDFF861C88F9E04C76FA06" bold="true" box="[199,352,1484,1509]" pageId="7" pageNumber="101">Distribution.</emphasis>
Species is distributed from northwest Chiapas,
<collectingCountry id="26F1A4A5FFDDFF861F38F9E04EDAFA06" box="[887,972,1484,1509]" name="Mexico" pageId="7" pageNumber="101">Mexico</collectingCountry>
to southern
<collectingCountry id="26F1A4A5FFDDFF861817F9E049A3FA06" box="[1112,1205,1484,1509]" name="Ecuador" pageId="7" pageNumber="101">Ecuador</collectingCountry>
(
<bibRefCitation id="3A7799C4FFDDFF86188BF9E0487EFA06" author="Boucher" box="[1220,1384,1484,1509]" pageId="7" pageNumber="101" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
).
<emphasis id="6C923827FFDDFF861C88F9C34CA2F9EB" bold="true" box="[199,436,1519,1544]" pageId="7" pageNumber="101">Material examined.</emphasis>
<collectingCountry id="26F1A4A5FFDDFF861D8FF9DC4F4EF9EB" box="[448,600,1520,1544]" name="Colombia" pageId="7" pageNumber="101">COLOMBIA</collectingCountry>
, CHOCO, Quibdó, Tutunendó, Corregimiento Pacorita,
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,
<date id="2A58C2F5FFDDFF861975F9C34883F9EB" box="[1338,1429,1519,1544]" pageId="7" pageNumber="101" value="2011-07">vii.2011</date>
. Reyes-Castillo &amp; Jiménez-Ferbans (
<specimenCount id="48E02FBCFFDDFF861E61FA384FABF9CE" box="[558,701,1556,1581]" pageId="7" pageNumber="101" type="generic">3 specimens</specimenCount>
).
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<paragraph id="5E59E435FFDDFF861CD8FA534FF6F979" blockId="7.[151,736,1663,1723]" box="[151,736,1663,1690]" pageId="7" pageNumber="101">
<heading id="05115359FFDDFF861CD8FA534FF6F979" bold="true" box="[151,736,1663,1690]" fontSize="11" level="1" pageId="7" pageNumber="101" reason="1">
<emphasis id="6C923827FFDDFF861CD8FA534FF6F979" bold="true" box="[151,736,1663,1690]" pageId="7" pageNumber="101">
<taxonomicName id="99E69FB6FFDDFF861CD8FA534DEAF97A" box="[151,252,1663,1689]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFDDFF861CD8FA534DEAF97A" bold="true" box="[151,252,1663,1689]" italics="true" pageId="7" pageNumber="101">Veturius</emphasis>
</taxonomicName>
(
<emphasis id="6C923827FFDDFF861D42FA534C79F97A" bold="true" box="[269,367,1663,1689]" italics="true" pageId="7" pageNumber="101">Vetur ius</emphasis>
)
<taxonomicName id="99E69FB6FFDDFF861DCCFA534FF6F979" authority="Eschscholtz, 1829" authorityName="Eschscholtz" authorityYear="1829" box="[387,736,1663,1690]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="101" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="6C923827FFDDFF861DCCFA534CFCF97A" bold="true" box="[387,490,1663,1689]" italics="true" pageId="7" pageNumber="101">sinuatus</emphasis>
(Eschscholtz, 1829)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="5E59E435FFDDFF861CD8FA8E4C6DF958" blockId="7.[151,736,1663,1723]" box="[151,379,1698,1723]" pageId="7" pageNumber="101">
(
<figureCitation id="C6DDF8B0FFDDFF861CD0FA8E4DFFF958" box="[159,233,1698,1723]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="7" pageNumber="101">Figs. 1</figureCitation>
e, 2e, 3e, 4e)
</paragraph>
<paragraph id="5E59E435FFDDFF891CD8FAC64DE9FEFF" blockId="7.[151,1437,1770,2011]" lastBlockId="8.[151,1437,151,719]" lastPageId="8" lastPageNumber="102" pageId="7" pageNumber="101">
<emphasis id="6C923827FFDDFF861CD8FAC64C3FF8E0" bold="true" box="[151,297,1770,1795]" pageId="7" pageNumber="101">Third instar</emphasis>
.
<emphasis id="6C923827FFDDFF861D75FAC64C65F8E0" box="[314,371,1770,1795]" italics="true" pageId="7" pageNumber="101">Body</emphasis>
: length 54.0 mm, width (prothorax) 9.0 mm; with tiny, stout setae; prosternal setae present; presence of other setae in the lateroventral part of the abdominal segments IIX and thoracic segments. Primary setal pattern: 3 HPA, 0 PD, 3 PSL, 0 MSD, 2 MSL, 0 MTD, 2 MTL, 0 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFDDFF86189AFB1E483AF8A8" box="[1237,1324,1842,1867]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="7" pageNumber="101" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
). The TM setae present only in the abdominal segments VIIIIX.
<emphasis id="6C923827FFDDFF861F75FB7A4E60F88C" box="[826,886,1878,1903]" italics="true" pageId="7" pageNumber="101">Head</emphasis>
: without uniform pile of short, golden setae; epicranial suture short; frontal sutures extend to the base of the antennae; width of head capsule 7.0 mm (
<figureCitation id="C6DDF8B0FFDDFF861973FB56486BF870" box="[1340,1405,1914,1939]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="7" pageNumber="101">Fig. 1</figureCitation>
e). Mandibles with micro-asperities in the ventral base. Maxillary stipes with 15 micro-conical projections (
<figureCitation id="C6DDF8B0FFDDFF861974FBB24894F854" box="[1339,1410,1950,1975]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="7" pageNumber="101">Fig. 2</figureCitation>
e).
<emphasis id="6C923827FFDDFF861CD8FBE84DDAF838" box="[151,204,1988,2011]" italics="true" pageId="7" pageNumber="101">Legs</emphasis>
: pars stridens (mesocoxae) length 1143.14 µm, width 849.42 µm, distance into striate 14.76 µm, 62 striae; mesocoxae and metacoxae with 3 internal setae. Metathoracic legs with some short lateral setae (side of I tooth) and micro-asperities in the posterior areas; anterior margin with 6 teeth, I and IIIVI well developed and almost the same size, II is the smallest tooth (
<figureCitation id="C6DDF8B0FFD2FF891E51FCF34F77FF1B" box="[542,609,223,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="8" pageNumber="102">Fig. 3</figureCitation>
e).
<emphasis id="6C923827FFD2FF891EC9FCF34FE6FF1B" box="[646,752,223,248]" italics="true" pageId="8" pageNumber="102">Abdomen</emphasis>
: raster with setae; anal ring with 20 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFD2FF891CD0FD284DF2FEFF" box="[159,228,260,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="8" pageNumber="102">Fig. 4</figureCitation>
e).
</paragraph>
<paragraph id="5E59E435FFD2FF891C88FD0B4944FE4E" blockId="8.[151,1437,151,719]" pageId="8" pageNumber="102">
<emphasis id="6C923827FFD2FF891C88FD0B4C69FEA3" bold="true" box="[199,383,295,320]" pageId="8" pageNumber="102">Natural history</emphasis>
. Larvae were collected in third instar and reared in laboratory conditions (25+2 °C; 54+2% relative humidity; photoperiod 12 light: 8 dark hours) using pieces of decaying logs as food. The third instars to pre-pupal phase took an average of 46 days, pre-pupal phase took an average of four days, pupal phase took an average of 20 days. The complete sclerotization of adults took an average of 40 days.
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD2FF891C88FD9B4C59FDDE" pageId="8" pageNumber="102" type="distribution">
<paragraph id="5E59E435FFD2FF891C88FD9B4C59FDDE" blockId="8.[151,1437,151,719]" pageId="8" pageNumber="102">
<emphasis id="6C923827FFD2FF891C88FD9B4C76FE33" bold="true" box="[199,352,439,464]" pageId="8" pageNumber="102">Distribution.</emphasis>
Endemic species from Brazilian Atlantic. States of Bahia, Espírito Santo, Goias, Mato Grosso do Sul, Mato Grosso, Minas Gerais, Paraná, Santa Catarina, Rio de Janeiro (included
<collectingCountry id="26F1A4A5FFD2FF891864FDF04941FE16" box="[1067,1111,476,501]" name="Iceland" pageId="8" pageNumber="102">Ilha</collectingCountry>
Grande), Rio Grande do Sul, São Paulo (included
<collectingCountry id="26F1A4A5FFD2FF891DCAFDD34CA7FDFB" box="[389,433,511,536]" name="Iceland" pageId="8" pageNumber="102">Ilha</collectingCountry>
de São Sebastião),
<collectingCountry id="26F1A4A5FFD2FF891EDAFDD34FCDFDFB" box="[661,731,511,536]" name="Brazil" pageId="8" pageNumber="102">Brasil</collectingCountry>
; Eastern Region,
<collectingCountry id="26F1A4A5FFD2FF891FE8FE2C4919FDFB" box="[935,1039,512,536]" name="Paraguay" pageId="8" pageNumber="102">Paraguay</collectingCountry>
and Misiones Province,
<collectingCountry id="26F1A4A5FFD2FF891962FDD3488AFDFB" box="[1325,1436,511,536]" name="Argentina" pageId="8" pageNumber="102">Argentina</collectingCountry>
(
<bibRefCitation id="3A7799C4FFD2FF891CD1FE084C55FDDE" author="Boucher" box="[158,323,548,573]" pageId="8" pageNumber="102" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD2FF891C88FE6449B7FD2C" pageId="8" pageNumber="102" type="materials_examined">
<paragraph id="5E59E435FFD2FF891C88FE6449B7FD2C" blockId="8.[151,1437,151,719]" pageId="8" pageNumber="102">
<emphasis id="6C923827FFD2FF891C88FE644CADFD82" bold="true" box="[199,443,584,609]" pageId="8" pageNumber="102">Material examined.</emphasis>
<collectingCountry id="26F1A4A5FFD2FF891D82FE654F23FD82" box="[461,565,585,609]" name="Brazil" pageId="8" pageNumber="102">BRAZIL</collectingCountry>
,
<collectionCode id="38F77CF0FFD2FF891E04FE654FB3FD82" box="[587,677,585,609]" pageId="8" pageNumber="102">MINAS</collectionCode>
GERAIS, Viçosa,
<collectingCountry id="26F1A4A5FFD2FF891FD5FE654EC2FD83" box="[922,980,585,608]" name="Malta" pageId="8" pageNumber="102">Mata</collectingCountry>
do Paraíso,
<geoCoordinate id="3BD282F2FFD2FF891831FE6549EBFD82" box="[1150,1277,583,609]" direction="south" orientation="latitude" pageId="8" pageNumber="102" precision="15" value="-20.801943">20°48'07''S</geoCoordinate>
<geoCoordinate id="3BD282F2FFD2FF891941FE654881FD82" box="[1294,1431,583,609]" direction="west" orientation="longitude" pageId="8" pageNumber="102" precision="15" value="-42.85861">42°51'31''W</geoCoordinate>
,
<date id="2A58C2F5FFD2FF891CD8FE424C10FD65" box="[151,262,622,646]" pageId="8" pageNumber="102" value="2013-10-11">11.x.2013</date>
. H. Vacacela, H. Cirqueira, &amp; Salazar-Niño (
<specimenCount id="48E02FBCFFD2FF891EB1FE414E98FD65" box="[766,910,621,646]" pageId="8" pageNumber="102" type="generic">6 specimens</specimenCount>
).
<collectingCountry id="26F1A4A5FFD2FF891FD0FE424911FD65" box="[927,1031,622,646]" name="Brazil" pageId="8" pageNumber="102">BRAZIL</collectingCountry>
,
<collectionCode id="38F77CF0FFD2FF89185DFE42497AFD65" box="[1042,1132,622,646]" pageId="8" pageNumber="102">MINAS</collectionCode>
GERAIS, Viçosa,
<collectingCountry id="26F1A4A5FFD2FF89190FFE42486CFD66" box="[1344,1402,622,645]" name="Malta" pageId="8" pageNumber="102">Mata</collectingCountry>
do Paraíso,
<geoCoordinate id="3BD282F2FFD2FF891CB7FEBE4C61FD49" box="[248,375,656,682]" direction="south" orientation="latitude" pageId="8" pageNumber="102" precision="15" value="-20.801943">20°48'07''S</geoCoordinate>
<geoCoordinate id="3BD282F2FFD2FF891D31FEBE4F11FD48" box="[382,519,656,683]" direction="west" orientation="longitude" pageId="8" pageNumber="102" precision="15" value="-42.85861">42°51'31''W</geoCoordinate>
,
<date id="2A58C2F5FFD2FF891E5BFEBE4F89FD48" box="[532,671,658,683]" pageId="8" pageNumber="102" value="2014-08-27">27.viii.2014</date>
. Salazar-Niño (
<specimenCount id="48E02FBCFFD2FF891F02FEBE4EFAFD48" box="[845,1004,658,683]" pageId="8" pageNumber="102" type="generic">12 specimens</specimenCount>
).
<collectingCountry id="26F1A4A5FFD2FF891FB0FEBE4971FD49" box="[1023,1127,658,682]" name="Brazil" pageId="8" pageNumber="102">BRAZIL</collectingCountry>
,
<collectionCode id="38F77CF0FFD2FF89183CFEBE49DBFD49" box="[1139,1229,658,682]" pageId="8" pageNumber="102">MINAS</collectionCode>
GERAIS, Viçosa, Federal University of Viçosa, Recanto das Cigarras,
<date id="2A58C2F5FFD2FF891EACFE9B4E41FD2C" box="[739,855,695,719]" pageId="8" pageNumber="102" value="2012-10-17">17.x.2012</date>
. L. Monteiro (
<specimenCount id="48E02FBCFFD2FF891FB7FE9B4980FD2C" box="[1016,1174,694,719]" pageId="8" pageNumber="102" type="generic">10 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,92 @@
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<mods:title id="2511F4026D50A29960CA025F6F4D91E9">An annotated checklist of millipede fauna from Slovakia, with ecological and biogeographic characteristics</mods:title>
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<mods:namePart id="2BD015C958D5067C91375F2CC59F9ECE">Haľkova, Beata</mods:namePart>
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<treatment id="D64F6E1F0B9A58809006DBADB01E7E7C" LSID="urn:lsid:plazi:treatment:D64F6E1F0B9A58809006DBADB01E7E7C" httpUri="http://treatment.plazi.org/id/D64F6E1F0B9A58809006DBADB01E7E7C" lastPageNumber="71495" pageId="0" pageNumber="71495">
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<paragraph id="BDD4964F4C50256624CE37513FAB44F7" pageId="0" pageNumber="71495">
<taxonomicName id="EA0092AD5C152DC773DBBED4E9E3FEAB" ID-CoL="3GCTK" ID-ENA="1392625" LSID="D64F6E1F-0B9A-5880-9006-DBADB01E7E7C" authority="Brandt, 1833" authorityName="Brandt" authorityYear="1833" class="Diplopoda" family="Glomeridae" genus="Glomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glomeris tetrasticha" order="Glomerida" pageId="0" pageNumber="71495" phylum="Arthropoda" rank="species" species="tetrasticha">Glomeris tetrasticha Brandt, 1833</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F4D8A67E4C22889BC48E2D491F28FDB8" pageId="0" pageNumber="71495" type="distribution">
<paragraph id="E1491342DBE0197DF28455FA112518BD" pageId="0" pageNumber="71495">Distribution</paragraph>
<paragraph id="0B34665923D1122B22CFC2C6EF6CD0E7" pageId="0" pageNumber="71495">Central and East European</paragraph>
</subSubSection>
<subSubSection id="C5B8F424B97372E8DB707F4332B4DFA3" pageId="0" pageNumber="71495" type="notes">
<paragraph id="F9380236F473AFB928C224B8D00540F9" pageId="0" pageNumber="71495">Notes</paragraph>
<paragraph id="8D7E1D5E62ECEC7C7590F6871E3E8589" pageId="0" pageNumber="71495">A, e</paragraph>
<paragraph id="46521E137190F1994D1B0E9A2706322C" pageId="0" pageNumber="71495">
Some authors mentioned the occurrence of
<taxonomicName id="1131E6BD6288D438D29C5CC687053CE8" class="Diplopoda" family="Glomeridae" genus="Glomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glomeris connexa" order="Glomerida" pageId="0" pageNumber="71495" phylum="Arthropoda" rank="species" species="connexa">
<emphasis id="D4D377A582BB2AD6550F695922DFB6D8" italics="true" pageId="0" pageNumber="71495">Glomeris connexa</emphasis>
</taxonomicName>
C. L. Koch, 1847 in Slovakia. This species is characteristic for its south-western European distribution (
<bibRefCitation id="7E7F1C9D50716E5535461A6F52200C35" author="Kime, R. D." journalOrPublisher="Atlas of European millipedes (Class Diplopoda)" pageId="0" pageNumber="71495" refId="B7164722" refString="Kime, R. D., Enghoff, H., 2011. . Atlas of European millipedes (Class Diplopoda) Vol. 1" volume="Vol. 1" year="2011">Kime and Enghoff 2011</bibRefCitation>
). In the literature from the last century, there is a lot of data on the findings and its occurrence. However, it should be noted that, in most cases, under this name, there were referred other species, especially
<taxonomicName id="6E7ADEC178E5DD7DCE200D29CE823286" lsidName="G. tetrasticha" pageId="0" pageNumber="71495" rank="species" species="tetrasticha">
<emphasis id="383AB59FD44C27AB237C0512DF6815C4" italics="true" pageId="0" pageNumber="71495">G. tetrasticha</emphasis>
</taxonomicName>
(
<bibRefCitation id="AA60097E6734976B17AF941EB5704EDB" author="Kime, R. D." journalOrPublisher="Atlas of European millipedes (Class Diplopoda)" pageId="0" pageNumber="71495" refId="B7164722" refString="Kime, R. D., Enghoff, H., 2011. . Atlas of European millipedes (Class Diplopoda) Vol. 1" volume="Vol. 1" year="2011">Kime and Enghoff 2011</bibRefCitation>
,
<bibRefCitation id="D1F1E29245C264564E2358935A873C11" author="Kocourek, P" journalOrPublisher="Cesky svaz ochrancu prirody Vlasim" pageId="0" pageNumber="71495" refId="B7329953" refString="Kocourek, P, Tajovsky, K, Dolejs, P, 2017. Mnohonozky Ceske republiky. Prirucka pro urcovani nasich druhu. Cesky svaz ochrancu prirody Vlasim" title="Mnohonozky Ceske republiky. Prirucka pro urcovani nasich druhu" year="2017">Kocourek et al. 2017</bibRefCitation>
). From Slovakia, no recent reliable records of its occurrence are available; therefore, the older data on this species name are preliminarily all included under
<taxonomicName id="22E5535847564148442F50A9CD045B81" lsidName="G. tetrasticha" pageId="0" pageNumber="71495" rank="species" species="tetrasticha">
<emphasis id="4E197093C2E594FEF9F976477EC17D46" italics="true" pageId="0" pageNumber="71495">G. tetrasticha</emphasis>
</taxonomicName>
. However, due to the findings from the Czech Republic, very close to the border with Slovakia, the occurrence of this species cannot be ruled out.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,415 @@
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<mods:title id="D45734436108F52B1C238526DFA2990D">Telosticta fugispinosa sp. nov. from Sabah (Odonata: Zygoptera: Platystictidae)</mods:title>
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<treatment id="D64F87A53E424C3782A0F94DFF1DF8CA" ID-DOI="http://doi.org/10.5281/zenodo.6088186" ID-GBIF-Taxon="119568799" ID-Zenodo-Dep="6088186" LSID="urn:lsid:plazi:treatment:D64F87A53E424C3782A0F94DFF1DF8CA" httpUri="http://treatment.plazi.org/id/D64F87A53E424C3782A0F94DFF1DF8CA" lastPageId="4" lastPageNumber="394" pageId="0" pageNumber="390">
<subSubSection id="16FC65383E424C3382A0F94DFB71F888" pageId="0" pageNumber="390" type="nomenclature">
<paragraph id="5E5936B33E424C3382A0F94DFDFAF92F" blockId="0.[151,513,1734,1793]" box="[151,513,1734,1760]" pageId="0" pageNumber="390">
<heading id="051181DF3E424C3382A0F94DFDFAF92F" bold="true" box="[151,513,1734,1760]" fontSize="11" level="1" pageId="0" pageNumber="390" reason="1">
<emphasis id="6C92EAA13E424C3382A0F94DFDFAF92F" bold="true" box="[151,513,1734,1760]" pageId="0" pageNumber="390">
<taxonomicName id="99E64D303E424C3382A0F94DFE67F92F" ID-CoL="9X4GR" box="[151,412,1734,1760]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="390" phylum="Arthropoda" rank="species" species="fugispinosa" status="sp. nov.">
<emphasis id="6C92EAA13E424C3382A0F94DFE67F92F" bold="true" box="[151,412,1734,1760]" italics="true" pageId="0" pageNumber="390">Telosticta fugispinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77A157DA3E424C338393F94DFDFAF92F" box="[420,513,1734,1760]" pageId="0" pageNumber="390" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5E5936B33E424C3382A0F963FEDBF8CE" blockId="0.[151,513,1734,1793]" box="[151,288,1768,1793]" pageId="0" pageNumber="390">
(
<figureCitation id="C6DD2A363E424C3382A8F963FEECF8CE" box="[159,279,1768,1793]" captionStart-0="FIGURES 1 2" captionStart-1="FIGURES 3 4" captionStart-2="FIGURES 5 6" captionStart-3="FIGURE 7 9" captionStart-4="FIGURES 10 12" captionStartId-0="1.[151,264,1116,1139]" captionStartId-1="1.[151,264,1964,1987]" captionStartId-2="2.[151,264,706,729]" captionStartId-3="2.[151,250,1934,1956]" captionStartId-4="3.[151,264,1014,1037]" captionTargetBox-0="[152,1434,807,1091]" captionTargetBox-1="[286,1296,1302,1929]" captionTargetBox-2="[151,1435,193,685]" captionTargetBox-3="[453,1133,778,1911]" captionTargetBox-4="[182,1405,200,989]" captionTargetId-0="figure@1.[152,1435,807,1091]" captionTargetId-1="figure@1.[264,1323,1188,1944]" captionTargetId-2="figure@2.[151,1435,193,685]" captionTargetId-3="figure@2.[453,1133,778,1912]" captionTargetId-4="figure@3.[173,1413,193,993]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="2" captionTargetPageId-3="2" captionTargetPageId-4="3" captionText-0="FIGURES 1 2. Habitus of Telosticta fugispinosa: (1) holotype male; (2) paratype female (SAB 12 _ PST 27)." captionText-1="FIGURES 3 4. Telosticta fugispinosa posterior pronotal lobe: (3) holotype male, lateral view; (4) paratype female (SAB 12 _ PST 27), lateral view." captionText-2="FIGURES 5 6. Telosticta fugispinosa holotype male synthorax: (5) dorsal view; (6) lateral view." captionText-3="FIGURE 7 9. Telosticta fugispinosa abdominal markings: (7) terminal abdominal segments of holotype male, dorsal view; (8) terminal abdominal segments of paratype male (SAB 12 _ 49), dorsal view; (9) terminal abdominal segments of paratype female (SAB 12 _ PST 27), lateral view." captionText-4="FIGURES 10 12. Telosticta fugispinosa holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm." httpUri-0="https://zenodo.org/record/255771/files/figure.png" httpUri-1="https://zenodo.org/record/255772/files/figure.png" httpUri-2="https://zenodo.org/record/255773/files/figure.png" httpUri-3="https://zenodo.org/record/255774/files/figure.png" httpUri-4="https://zenodo.org/record/255775/files/figure.png" pageId="0" pageNumber="390">Figs. 112</figureCitation>
)
</paragraph>
<paragraph id="5E5936B33E424C3382A0F8BAFB71F888" blockId="0.[151,1162,1841,1863]" box="[151,1162,1841,1863]" pageId="0" pageNumber="390">
<taxonomicName id="99E64D303E424C3382A0F8BAFF02F888" box="[151,249,1841,1863]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E424C3382A0F8BAFF02F888" box="[151,249,1841,1863]" italics="true" pageId="0" pageNumber="390">Telosticta</emphasis>
</taxonomicName>
undetermined D;—
<bibRefCitation id="3A774B423E424C3383F6F8B9FB7DF888" author="Dow" box="[449,1158,1841,1863]" pageId="0" pageNumber="390" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012">Dow &amp; Orr (2012: 396, record of female, Poring Hot Springs, Sabah)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="16FC65383E424C3282A0F8F8FDDAFE3B" lastPageId="1" lastPageNumber="391" pageId="0" pageNumber="390" type="materials_examined">
<paragraph id="5E5936B33E424C3282A0F8F8FDDAFE3B" blockId="0.[151,1436,1907,2004]" lastBlockId="1.[151,1437,151,752]" lastPageId="1" lastPageNumber="391" pageId="0" pageNumber="390">
<emphasis id="6C92EAA13E424C3382A0F8F8FE34F843" bold="true" box="[151,463,1907,1932]" pageId="0" pageNumber="390">
<typeStatus id="815D88113E424C3382A0F8F8FF29F843" box="[151,210,1907,1932]" pageId="0" pageNumber="390">Type</typeStatus>
material:
<typeStatus id="815D88113E424C33836EF8F8FE31F843" box="[345,458,1907,1932]" pageId="0" pageNumber="390" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="48E0FD3A3E424C3383ECF8FFFDF0F844" box="[475,523,1908,1932]" pageId="0" pageNumber="390" type="generic">1 ♂</specimenCount>
(SAB12_PST1,
<collectionCode id="38F7AE763E424C3380E5F8F8FCD7F843" LSID="urn:lsid:biocol.org:col:34992" box="[722,812,1907,1932]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="0" pageNumber="390">RMNH</collectionCode>
.INS.507772),
<collectingCountry id="26F176233E424C3381E5F8F8FBC0F843" box="[978,1083,1907,1932]" name="Malaysia" pageId="0" pageNumber="390">Malaysia</collectingCountry>
, Sabah, West Coast Division, Crocker Range National Park, Inobong, Kimamabang waterfall stream system (below waterfall),
<date id="2A5810733E424C3386DEF813FA9DF87F" box="[1257,1382,1943,1968]" pageId="0" pageNumber="390" value="2012-09-21">21 ix 2012</date>
, leg. R. A. Dow, in
<collectionCode id="38F7AE763E424C338362F837FE56F81C" LSID="urn:lsid:biocol.org:col:34992" box="[341,429,1980,2003]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="0" pageNumber="390">RMNH</collectionCode>
.
<emphasis id="6C92EAA13E424C33838AF830FDC5F81B" bold="true" box="[445,574,1979,2004]" pageId="0" pageNumber="390">
<typeStatus id="815D88113E424C33838AF830FDC2F81B" box="[445,569,1979,2004]" pageId="0" pageNumber="390" type="paratype">Paratypes</typeStatus>
:
</emphasis>
All from
<collectingCountry id="26F176233E424C3380F5F830FCD7F81B" box="[706,812,1979,2004]" name="Malaysia" pageId="0" pageNumber="390">Malaysia</collectingCountry>
, Sabah, West Coast Division:
<specimenCount id="48E0FD3A3E424C3386A8F837FB2BF81C" box="[1183,1232,1980,2004]" pageId="0" pageNumber="390" type="generic">3 ♂</specimenCount>
(SAB12_PST49; SAB12_PST72,
<collectionCode id="38F7AE763E434C328363FF1CFE55FF7F" LSID="urn:lsid:biocol.org:col:34992" box="[340,430,151,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.507677; SAB12_PST77,
<collectionCode id="38F7AE763E434C328132FF1CFCA4FF7F" LSID="urn:lsid:biocol.org:col:34992" box="[773,863,151,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.507668), Kinabalu National Park, Poring Hot Springs, small streams crossed by or near to trail to Langanan waterfall, above Kipungit stream,
<date id="2A5810733E434C3286DEFF37FA9DFF1B" box="[1257,1382,188,213]" pageId="1" pageNumber="391" value="2012-09-11">11 ix 2012</date>
, leg. R.A. Dow;
<specimenCount id="48E0FD3A3E434C32832FFF6BFEB8FF38" box="[280,323,224,248]" pageId="1" pageNumber="391" type="generic">1 ♂</specimenCount>
(SAB12_PST48), same location and collector,
<date id="2A5810733E434C32816EFF6BFC2FFF37" box="[857,980,223,248]" pageId="1" pageNumber="391" value="2012-09-12">12 ix 2012</date>
; 1 ♀, same area, trailside in forest,
<date id="2A5810733E434C328754FF6BFF29FED2" pageId="1" pageNumber="391" value="2005-04-21">21 iv 2005</date>
, leg. unknown, in
<collectionCode id="38F7AE763E434C32839DFE8EFDF9FED3" LSID="urn:lsid:biocol.org:col:34992" box="[426,514,261,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
;
<specimenCount id="48E0FD3A3E434C328038FE8FFDB1FED3" box="[527,586,260,284]" pageId="1" pageNumber="391" type="generic">10 ♂</specimenCount>
(SAB12_PST3,
<collectionCode id="38F7AE763E434C32813BFE8FFC9DFED2" LSID="urn:lsid:biocol.org:col:34992" box="[780,870,260,285]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.507747; SAB12_PST3237; SAB12_PST62,
<collectionCode id="38F7AE763E434C3282A0FEACFF0AFE8F" LSID="urn:lsid:biocol.org:col:34992" box="[151,241,295,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.597748; SAB12_PST66,
<collectionCode id="38F7AE763E434C328073FEACFD65FE8F" LSID="urn:lsid:biocol.org:col:34992" box="[580,670,295,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.507755), 1 ♀ (SAB12_PST4;
<collectionCode id="38F7AE763E434C32861FFEACFB79FE8F" LSID="urn:lsid:biocol.org:col:34992" box="[1064,1154,295,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.507749), location as
<typeStatus id="815D88113E434C3282A0FEC7FF07FEAA" box="[151,252,332,357]" pageId="1" pageNumber="391" type="holotype">holotype</typeStatus>
,
<date id="2A5810733E434C32833FFEC7FE7DFEAB" box="[264,390,332,357]" pageId="1" pageNumber="391" value="2012-09-18">18 ix 2012</date>
, leg. R.A. Dow;
<specimenCount id="48E0FD3A3E434C32807FFEC7FD88FEAB" box="[584,627,332,356]" pageId="1" pageNumber="391" type="generic">4 ♂</specimenCount>
(SAB12_PST2931, 87), same location and collector,
<date id="2A5810733E434C3286D1FEC7FA9FFEAB" box="[1254,1380,332,357]" pageId="1" pageNumber="391" value="2012-09-19">19 ix 2012</date>
;
<specimenCount id="48E0FD3A3E434C328746FEC7FA67FEAB" box="[1393,1436,332,357]" pageId="1" pageNumber="391" type="generic">6 ♂</specimenCount>
(SAB126_PST69, 26; SAB12_PST61,
<collectionCode id="38F7AE763E434C3280A2FEE4FD15FE47" LSID="urn:lsid:biocol.org:col:34992" box="[661,750,367,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.507770), 1 ♀ (SAB12_PST27, in tandem with SAB12_PST26), data as
<typeStatus id="815D88113E434C32838BFE1FFDD9FE62" box="[444,546,404,429]" pageId="1" pageNumber="391" type="holotype">holotype</typeStatus>
;
<specimenCount id="48E0FD3A3E434C328005FE1FFD9BFE63" box="[562,608,404,428]" pageId="1" pageNumber="391" type="generic">3 ♂</specimenCount>
(AA044, 49, 57; ODO 01378-80), same location,
<date id="2A5810733E434C32868EFE1FFAC6FE63" box="[1209,1341,404,429]" pageId="1" pageNumber="391" value="2012-09-22">22 ix 2012</date>
, leg. A. Afendy, in
<collectionCode id="38F7AE763E434C328320FE33FEA1FE1F" box="[279,346,440,464]" pageId="1" pageNumber="391">ITCB</collectionCode>
;
<specimenCount id="48E0FD3A3E434C328350FE33FE69FE00" box="[359,402,440,464]" pageId="1" pageNumber="391" type="generic">1 ♂</specimenCount>
(SAB12_PST84,
<collectionCode id="38F7AE763E434C328056FE3CFD40FE1F" LSID="urn:lsid:biocol.org:col:34992" box="[609,699,439,464]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="391">RMNH</collectionCode>
.INS.507766), same national park, Inobong, Batu Dinding stream system,
<date id="2A5810733E434C3282C4FE57FE96FE3B" box="[243,365,476,501]" pageId="1" pageNumber="391" value="2012-09-20">20 ix 2012</date>
, leg. R.A. Dow.
</paragraph>
</subSubSection>
<subSubSection id="16FC65383E434C3282F0FE74FEA4FDF3" pageId="1" pageNumber="391" type="etymology">
<paragraph id="5E5936B33E434C3282F0FE74FEA4FDF3" blockId="1.[151,1437,151,752]" pageId="1" pageNumber="391">
<emphasis id="6C92EAA13E434C3282F0FE74FEB5FDD7" bold="true" box="[199,334,511,536]" pageId="1" pageNumber="391">Etymology.</emphasis>
<taxonomicName id="99E64D303E434C328362FD8BFE2DFDD7" box="[341,470,512,536]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E434C328362FD8BFE2DFDD7" box="[341,470,512,536]" italics="true" pageId="1" pageNumber="391">fugispinosa</emphasis>
</taxonomicName>
, an adjective suggesting a fleeting spine, in reference to the absent or vestigial spine on the paraprocts.
</paragraph>
</subSubSection>
<subSubSection id="16FC65383E434C3782F0FDCCFEFCFC93" lastPageId="4" lastPageNumber="394" pageId="1" pageNumber="391" type="description">
<paragraph id="5E5936B33E434C3282F0FDCCFAB3FD3F" blockId="1.[151,1437,151,752]" pageId="1" pageNumber="391">
<emphasis id="6C92EAA13E434C3282F0FDCCFDD0FDAF" bold="true" box="[199,555,583,608]" pageId="1" pageNumber="391">
Description of
<typeStatus id="815D88113E434C32834CFDCCFE1AFDAF" box="[379,481,583,608]" pageId="1" pageNumber="391" type="holotype">holotype</typeStatus>
male.
</emphasis>
Head: Labium pale. Basal 2/3 of labrum pale blue, black along free margin. Anteclypeus blue, postclypeus shining black. Mandible bases blue in corner by clypeus, black below. Vertex and frons bronzy black, occiput shining black. Ratio of width of compound eye to width of vertex measured at level of lateral ocelli slightly more than 9/10. Transverse occipital carina with lateral extremities angulated and prominent. Ocelli whitish. Antenna with scape and pedicel pale yellowish, brown at top of pedicel, remainder missing.
</paragraph>
<caption id="0A99663B3E434C3282A0FBD7FB26FBBD" box="[151,1245,1116,1139]" httpUri="https://zenodo.org/record/255771/files/figure.png" pageId="1" pageNumber="391" targetBox="[152,1434,807,1091]" targetPageId="1">
<paragraph id="5E5936B33E434C3282A0FBD7FB26FBBD" blockId="1.[151,1245,1116,1139]" box="[151,1245,1116,1139]" pageId="1" pageNumber="391">
<emphasis id="6C92EAA13E434C3282A0FBD7FEC1FBBE" bold="true" box="[151,314,1116,1139]" pageId="1" pageNumber="391">FIGURES 12.</emphasis>
Habitus of
<taxonomicName id="99E64D303E434C328386FBD7FD76FBBD" box="[433,653,1116,1138]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E434C328386FBD7FD76FBBD" box="[433,653,1116,1138]" italics="true" pageId="1" pageNumber="391">Telosticta fugispinosa</emphasis>
</taxonomicName>
: (1) holotype male; (2) paratype female (SAB12_PST27).
</paragraph>
</caption>
<caption id="0A99663B3E434C3282A0F827FE36F82D" httpUri="https://zenodo.org/record/255772/files/figure.png" pageId="1" pageNumber="391" targetBox="[286,1296,1302,1929]" targetPageId="1">
<paragraph id="5E5936B33E434C3282A0F827FE36F82D" blockId="1.[151,1436,1964,2018]" pageId="1" pageNumber="391">
<emphasis id="6C92EAA13E434C3282A0F827FEB8F80D" bold="true" box="[151,323,1964,1987]" pageId="1" pageNumber="391">FIGURES 34.</emphasis>
<taxonomicName id="99E64D303E434C328365F826FDCCF80C" box="[338,567,1965,1987]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E434C328365F826FDCCF80C" box="[338,567,1965,1987]" italics="true" pageId="1" pageNumber="391">Telosticta fugispinosa</emphasis>
</taxonomicName>
posterior pronotal lobe: (3) holotype male, lateral view; (4) paratype female (SAB12_PST27), lateral view.
</paragraph>
</caption>
<caption id="0A99663B3E404C3182A0FD49FB89FD17" box="[151,1138,706,729]" httpUri="https://zenodo.org/record/255773/files/figure.png" pageId="2" pageNumber="392" targetBox="[151,1435,193,685]" targetPageId="2">
<paragraph id="5E5936B33E404C3182A0FD49FB89FD17" blockId="2.[151,1138,706,729]" box="[151,1138,706,729]" pageId="2" pageNumber="392">
<emphasis id="6C92EAA13E404C3182A0FD49FEC1FD18" bold="true" box="[151,314,706,729]" pageId="2" pageNumber="392">FIGURES 56.</emphasis>
<taxonomicName id="99E64D303E404C318377FD49FDE6FD17" box="[320,541,706,728]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="392" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E404C318377FD49FDE6FD17" box="[320,541,706,728]" italics="true" pageId="2" pageNumber="392">Telosticta fugispinosa</emphasis>
</taxonomicName>
holotype male synthorax: (5) dorsal view; (6) lateral view.
</paragraph>
</caption>
<caption id="0A99663B3E404C3182A0F805FE36F82E" httpUri="https://zenodo.org/record/255774/files/figure.png" pageId="2" pageNumber="392" targetBox="[453,1133,778,1911]" targetPageId="2">
<paragraph id="5E5936B33E404C3182A0F805FE36F82E" blockId="2.[151,1436,1934,2017]" pageId="2" pageNumber="392">
<emphasis id="6C92EAA13E404C3182A0F805FED7F86C" bold="true" box="[151,300,1934,1956]" pageId="2" pageNumber="392">FIGURE 79.</emphasis>
<taxonomicName id="99E64D303E404C318305F805FDF4F86B" box="[306,527,1934,1956]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="392" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E404C318305F805FDF4F86B" box="[306,527,1934,1956]" italics="true" pageId="2" pageNumber="392">Telosticta fugispinosa</emphasis>
</taxonomicName>
abdominal markings: (7) terminal abdominal segments of holotype male, dorsal view; (8) terminal abdominal segments of paratype male (SAB12_49), dorsal view; (9) terminal abdominal segments of paratype female (SAB12_PST27), lateral view.
</paragraph>
</caption>
<caption id="0A99663B3E414C3082A0FC7DFC29FBE4" httpUri="https://zenodo.org/record/255775/files/figure.png" pageId="3" pageNumber="393" targetBox="[182,1405,200,989]" targetPageId="3">
<paragraph id="5E5936B33E414C3082A0FC7DFC29FBE4" blockId="3.[151,1436,1014,1067]" pageId="3" pageNumber="393">
<emphasis id="6C92EAA13E414C3082A0FC7DFEAFFBC3" bold="true" box="[151,340,1014,1037]" pageId="3" pageNumber="393">FIGURES 1012.</emphasis>
<taxonomicName id="99E64D303E414C30836BFC7CFDC2FBC2" box="[348,569,1015,1037]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="393" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E414C30836BFC7CFDC2FBC2" box="[348,569,1015,1037]" italics="true" pageId="3" pageNumber="393">Telosticta fugispinosa</emphasis>
</taxonomicName>
holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="5E5936B33E414C3082F0FBD3FB66F9A1" blockId="3.[151,1437,1112,2006]" pageId="3" pageNumber="393">
Thorax: Prothorax whitish with blue tint to central parts anterior and middle pronotal lobes, except to rear of propleuron where there are irregular dark markings; small black central marking on anterior pronotal lobe, irregular black markings to rear of middle pronotal lobe; entire posterior lobe black, becoming grey apically on long lateral process (
<figureCitation id="C6DD2A363E414C3082CBFB4FFEBBFB12" box="[252,320,1220,1245]" captionStart="FIGURES 3 4" captionStartId="1.[151,264,1964,1987]" captionTargetBox="[286,1296,1302,1929]" captionTargetId="figure@1.[264,1323,1188,1944]" captionTargetPageId="1" captionText="FIGURES 3 4. Telosticta fugispinosa posterior pronotal lobe: (3) holotype male, lateral view; (4) paratype female (SAB 12 _ PST 27), lateral view." httpUri="https://zenodo.org/record/255772/files/figure.png" pageId="3" pageNumber="393">Fig. 3</figureCitation>
) with tip reaching level of lower margin of propleuron. Synthorax: Mesepisternum bronzy black, with pair of blue antehumeral stripes extending ca two-thirds of distance to wing bases (
<figureCitation id="C6DD2A363E414C30864EFB63FB45FACE" box="[1145,1214,1256,1281]" captionStart="FIGURES 5 6" captionStartId="2.[151,264,706,729]" captionTargetBox="[151,1435,193,685]" captionTargetId="figure@2.[151,1435,193,685]" captionTargetPageId="2" captionText="FIGURES 5 6. Telosticta fugispinosa holotype male synthorax: (5) dorsal view; (6) lateral view." httpUri="https://zenodo.org/record/255773/files/figure.png" pageId="3" pageNumber="393">Fig. 5</figureCitation>
). Antealar triangles pale blue along mid-dorsal carina and in half at wing bases, rest black. Mesepimeron black. Metepisternum largely occupied by pale band, becoming blue towards wing bases, with short brown triangle based at antealar carina below, this extended as brownish band along metapleural suture to level of spiracle (
<figureCitation id="C6DD2A363E414C308660FADFFB67FAA2" box="[1111,1180,1364,1389]" captionStart="FIGURES 5 6" captionStartId="2.[151,264,706,729]" captionTargetBox="[151,1435,193,685]" captionTargetId="figure@2.[151,1435,193,685]" captionTargetPageId="2" captionText="FIGURES 5 6. Telosticta fugispinosa holotype male synthorax: (5) dorsal view; (6) lateral view." httpUri="https://zenodo.org/record/255773/files/figure.png" pageId="3" pageNumber="393">Fig. 6</figureCitation>
). Metepimeron almost entirely pale. Venter of synthorax pale. Legs (only right anterior and right middle legs still present beyond trochanter, left posterior leg entirely missing, however all legs agreeing with the description below in
<typeStatus id="815D88113E414C308717FA16FA6AFA7A" box="[1312,1425,1437,1461]" pageId="3" pageNumber="393" type="paratype">paratypes</typeStatus>
): each with coxa and trochanter pale, femur and tibia pale except around joint and black stripe along extensor surface of femur. Tarsus pale with some darker areas, very dark brown at apex and brown claws. Wings: 13 (left) to 14 (right) Px in Fw, 12 (left) to 13 (right) Px in Hw. Vein ab absent. Arculus slightly distal to Ax2. R4 arising distal to subnodus, IR3 joined to it by short stalk. Pterostigma trapezoidal with costal side slightly shorter than anal side, very dark brown with narrow white border, covering slightly more than one underlying cell.
</paragraph>
<paragraph id="5E5936B33E414C3082F0F9F2FA67F819" blockId="3.[151,1437,1112,2006]" pageId="3" pageNumber="393">
Abdomen: Largely dark brown and black. S1 pale with narrow darker apical annulus. S2 with yellowish cream basal annulus, divided dorsally, laterally this extending to posterior carina as pale wedge, otherwise dark brown. S37 dark brown with narrow pale basal annulus, faint centrally dorsally. S8 black with pale band along lower margin joined to large blue lateral dorsal marking, narrowly and irregularly divided dorsally (
<figureCitation id="C6DD2A363E414C308687F96EFB0FF931" box="[1200,1268,1765,1790]" captionStart="FIGURE 7 9" captionStartId="2.[151,250,1934,1956]" captionTargetBox="[453,1133,778,1911]" captionTargetId="figure@2.[453,1133,778,1912]" captionTargetPageId="2" captionText="FIGURE 7 9. Telosticta fugispinosa abdominal markings: (7) terminal abdominal segments of holotype male, dorsal view; (8) terminal abdominal segments of paratype male (SAB 12 _ 49), dorsal view; (9) terminal abdominal segments of paratype female (SAB 12 _ PST 27), lateral view." httpUri="https://zenodo.org/record/255774/files/figure.png" pageId="3" pageNumber="393">Fig. 7</figureCitation>
). S910 black. Anal appendages (
<figureCitation id="C6DD2A363E414C308352F882FE1EF8ED" box="[357,485,1801,1826]" captionStart="FIGURES 10 12" captionStartId="3.[151,264,1014,1037]" captionTargetBox="[182,1405,200,989]" captionTargetId="figure@3.[173,1413,193,993]" captionTargetPageId="3" captionText="FIGURES 10 12. Telosticta fugispinosa holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm." httpUri="https://zenodo.org/record/255775/files/figure.png" pageId="3" pageNumber="393">Figs.1012</figureCitation>
) largely black, with pale areas interior apically on cerci and scoop of paraprocts pale, also ventrally at interior base of paraprocts. Cerci ca 2.5 times length of S10, interior projection well developed as spur at ca one-third length from base, directed first inwards then expanded slightly upwards and strongly downwards directed, spur clearly visible in dorsal (
<figureCitation id="C6DD2A363E414C3080EFF8FEFCD1F841" box="[728,810,1909,1934]" captionStart="FIGURES 10 12" captionStartId="3.[151,264,1014,1037]" captionTargetBox="[182,1405,200,989]" captionTargetId="figure@3.[173,1413,193,993]" captionTargetPageId="3" captionText="FIGURES 10 12. Telosticta fugispinosa holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm." httpUri="https://zenodo.org/record/255775/files/figure.png" pageId="3" pageNumber="393">Fig. 10</figureCitation>
) and lateral (
<figureCitation id="C6DD2A363E414C3081F7F8FEFBEBF841" box="[960,1040,1909,1934]" captionStart="FIGURES 10 12" captionStartId="3.[151,264,1014,1037]" captionTargetBox="[182,1405,200,989]" captionTargetId="figure@3.[173,1413,193,993]" captionTargetPageId="3" captionText="FIGURES 10 12. Telosticta fugispinosa holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm." httpUri="https://zenodo.org/record/255775/files/figure.png" pageId="3" pageNumber="393">Fig. 11</figureCitation>
) view, with peg-like appearance in lateral view. Dorsal projection weakly developed, visible in dorsal view as small bump (
<figureCitation id="C6DD2A363E414C3086B5F812FB2CF87D" box="[1154,1239,1945,1970]" captionStart="FIGURES 10 12" captionStartId="3.[151,264,1014,1037]" captionTargetBox="[182,1405,200,989]" captionTargetId="figure@3.[173,1413,193,993]" captionTargetPageId="3" captionText="FIGURES 10 12. Telosticta fugispinosa holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm." httpUri="https://zenodo.org/record/255775/files/figure.png" pageId="3" pageNumber="393">Fig. 10</figureCitation>
). Cerci expanded dorsoventrally and interiorly shortly after half length, lower margin irregular, small subapical cleft (
<figureCitation id="C6DD2A363E414C308700F836FA75F819" box="[1335,1422,1981,2006]" captionStart="FIGURES 10 12" captionStartId="3.[151,264,1014,1037]" captionTargetBox="[182,1405,200,989]" captionTargetId="figure@3.[173,1413,193,993]" captionTargetPageId="3" captionText="FIGURES 10 12. Telosticta fugispinosa holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm." httpUri="https://zenodo.org/record/255775/files/figure.png" pageId="3" pageNumber="393">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="5E5936B33E464C3782A0FF13FB5EFF1A" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
Paraprocts just shorter than cerci, scoop spoon-like, turned inwards (
<figureCitation id="C6DD2A363E464C37819EFF1CFBF2FF7F" box="[937,1033,151,176]" captionStart="FIGURES 10 12" captionStartId="3.[151,264,1014,1037]" captionTargetBox="[182,1405,200,989]" captionTargetId="figure@3.[173,1413,193,993]" captionTargetPageId="3" captionText="FIGURES 10 12. Telosticta fugispinosa holotype male anal appendages: (10) dorsal view, scale bar 1 mm; (11) lateral view, scale bar 1 mm; (12) apical part right paraproct in ventral view, scale bar 0.5 mm." httpUri="https://zenodo.org/record/255775/files/figure.png" pageId="4" pageNumber="394">Figs. 12</figureCitation>
), spine vestigial, not easily visible. Genital ligula of typical form for genus, with tongue-like structure of terminal segment long.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FF54FB36FF37" blockId="4.[151,1437,151,1797]" box="[199,1229,223,248]" pageId="4" pageNumber="394">
<emphasis id="6C92EAA13E464C3782F0FF54FE8DFF37" bold="true" box="[199,374,223,248]" pageId="4" pageNumber="394">Measurements</emphasis>
(mm): abdomen without anal appendages 38.5, cercus just over 1, Hw 20.5.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FE8FFBDFFED2" blockId="4.[151,1437,151,1797]" box="[199,1060,260,285]" pageId="4" pageNumber="394">
<emphasis id="6C92EAA13E464C3782F0FE8FFDC2FED2" bold="true" box="[199,569,260,285]" pageId="4" pageNumber="394">
Description of
<typeStatus id="815D88113E464C378341FE8FFE19FED2" box="[374,482,260,285]" pageId="4" pageNumber="394" type="paratype">paratype</typeStatus>
female
</emphasis>
(SAB12_PST27).
<collectingCountry id="26F176233E464C37813BFE8FFCD0FED3" box="[780,811,260,284]" name="American Samoa" pageId="4" pageNumber="394">As</collectingCountry>
male except as noted.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FEACFD71FE1F" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
Thorax: Posterior pronotal lobe with only short lateral processes (
<figureCitation id="C6DD2A363E464C378191FEACFC17FE8F" box="[934,1004,295,320]" captionStart="FIGURES 3 4" captionStartId="1.[151,264,1964,1987]" captionTargetBox="[286,1296,1302,1929]" captionTargetId="figure@1.[264,1323,1188,1944]" captionTargetPageId="1" captionText="FIGURES 3 4. Telosticta fugispinosa posterior pronotal lobe: (3) holotype male, lateral view; (4) paratype female (SAB 12 _ PST 27), lateral view." httpUri="https://zenodo.org/record/255772/files/figure.png" pageId="4" pageNumber="394">Fig. 4</figureCitation>
). Wings: 14 Px in Fw, 13 Px in Hw. Abdomen: S35 with pale basal annulus not faded dorsally. S6 with no basal annulus but small white basal dorsal mark present. S7 with complete broad white basal annulus. Blue mark covering much of dorsum of 8 (
<figureCitation id="C6DD2A363E464C378721FEE4FAA0FE47" box="[1302,1371,367,392]" captionStart="FIGURE 7 9" captionStartId="2.[151,250,1934,1956]" captionTargetBox="[453,1133,778,1911]" captionTargetId="figure@2.[453,1133,778,1912]" captionTargetPageId="2" captionText="FIGURE 7 9. Telosticta fugispinosa abdominal markings: (7) terminal abdominal segments of holotype male, dorsal view; (8) terminal abdominal segments of paratype male (SAB 12 _ 49), dorsal view; (9) terminal abdominal segments of paratype female (SAB 12 _ PST 27), lateral view." httpUri="https://zenodo.org/record/255774/files/figure.png" pageId="4" pageNumber="394">Fig. 9</figureCitation>
). S10 short. Cerci just shorter than S10, approximately triangular. Ovipositor extending just beyond cerci, mostly black and dark brown with obscure pale markings.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FE57FBCAFE3B" blockId="4.[151,1437,151,1797]" box="[199,1073,476,501]" pageId="4" pageNumber="394">
<emphasis id="6C92EAA13E464C3782F0FE57FE8DFE3A" bold="true" box="[199,374,476,501]" pageId="4" pageNumber="394">Measurements</emphasis>
(mm): abdomen without appendages or ovipositor 33, Hw 21.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FE74FB50FD3F" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
<emphasis id="6C92EAA13E464C3782F0FE74FE29FDD7" bold="true" box="[199,466,511,536]" pageId="4" pageNumber="394">
Variation in
<typeStatus id="815D88113E464C37836BFE74FE29FDD7" box="[348,466,511,536]" pageId="4" pageNumber="394" type="paratype">paratypes</typeStatus>
</emphasis>
. The black marks on the anterior pronotal lobe and the rear part of the middle pronotal lobes are variable in size, almost entirely absent in a few individuals and more extensive in others; occasionally the anterior lobe marking is joined to those on the rear of the posterior pronotal lobe via the central pit. The shape of the blue dorsal marking on S8 of the male
<typeStatus id="815D88113E464C378045FDE6FD26FD4A" box="[626,733,621,645]" pageId="4" pageNumber="394" type="paratype">paratypes</typeStatus>
is variable, and the marking is frequently not divided centrally (e.g.,
<figureCitation id="C6DD2A363E464C3782EFFD04FEE0FD67" box="[216,283,655,680]" captionStart="FIGURE 7 9" captionStartId="2.[151,250,1934,1956]" captionTargetBox="[453,1133,778,1911]" captionTargetId="figure@2.[453,1133,778,1912]" captionTargetPageId="2" captionText="FIGURE 7 9. Telosticta fugispinosa abdominal markings: (7) terminal abdominal segments of holotype male, dorsal view; (8) terminal abdominal segments of paratype male (SAB 12 _ 49), dorsal view; (9) terminal abdominal segments of paratype female (SAB 12 _ PST 27), lateral view." httpUri="https://zenodo.org/record/255774/files/figure.png" pageId="4" pageNumber="394">Fig. 8</figureCitation>
); this is the case in all individuals from Poring. The spine of the paraprocts is entirely absent in some males. In males the only other significant variation is in size, with all individuals from Poring at the upper end of size range given below. In the two other female
<typeStatus id="815D88113E464C378086FD53FCE7FD3F" box="[689,796,728,752]" pageId="4" pageNumber="394" type="paratype">paratypes</typeStatus>
there is a basal pale annulus on S6.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FD77FEFCFC93" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
<emphasis id="6C92EAA13E464C3782F0FD77FE8DFCDA" bold="true" box="[199,374,764,789]" pageId="4" pageNumber="394">Measurements</emphasis>
(mm): Males: abdomen without anal appendages 3646, Hw 1924.5, 1214 Px in Fw, 1113 Px in Hw. Females: abdomen without anal appendages or ovipositor 3437, Hw 21.524, 1314 Px in Fw, 1213 Px in Hw.
</paragraph>
</subSubSection>
<subSubSection id="16FC65383E464C3782F0FCECFDACFC23" pageId="4" pageNumber="394" type="diagnosis">
<paragraph id="5E5936B33E464C3782F0FCECFDACFC23" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
<emphasis id="6C92EAA13E464C3782F0FCECFEBAFC4F" bold="true" box="[199,321,871,896]" pageId="4" pageNumber="394">Diagnosis.</emphasis>
A small
<taxonomicName id="99E64D303E464C378391FCECFDE9FC4F" box="[422,530,871,896]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C378391FCECFDE9FC4F" box="[422,530,871,896]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
with long blue antehumeral stripes. Males are easily distinguished from all other species of
<taxonomicName id="99E64D303E464C37833BFC07FE83FC6A" box="[268,376,908,933]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C37833BFC07FE83FC6A" box="[268,376,908,933]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
by the form of the paraprocts, with the scoop turned sharply inwards and with a vestigial or no spine; the shape of the terminal half of the cerci is also distinctive, and dorsal blue markings on the terminal abdominal segments are confined to S8.
</paragraph>
</subSubSection>
<subSubSection id="16FC65383E464C3782F0FC7CFF1DF8CA" pageId="4" pageNumber="394" type="discussion">
<paragraph id="5E5936B33E464C3782F0FC7CFD8AFBFA" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
<emphasis id="6C92EAA13E464C3782F0FC7CFECFFBDF" bold="true" box="[199,308,1015,1040]" pageId="4" pageNumber="394">Remarks</emphasis>
. Individuals of both sexes were found at moderate- to high-gradient forest streams from approximately
<quantity id="991E9B563E464C378308FB97FE4CFBFA" box="[319,439,1052,1077]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="4" pageNumber="394" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800m</quantity>
above sea level.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FBCBFC9EFBB2" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
In
<figureCitation id="C6DD2A363E464C3782D1FBB4FED3FB97" box="[230,296,1087,1112]" captionStart="FIGURES 5 6" captionStartId="2.[151,264,706,729]" captionTargetBox="[151,1435,193,685]" captionTargetId="figure@2.[151,1435,193,685]" captionTargetPageId="2" captionText="FIGURES 5 6. Telosticta fugispinosa holotype male synthorax: (5) dorsal view; (6) lateral view." httpUri="https://zenodo.org/record/255773/files/figure.png" pageId="4" pageNumber="394">Fig. 6</figureCitation>
there appears to be a small pale mark on the mesopleural suture near antealar carina of the
<typeStatus id="815D88113E464C378704FBB4FA63FB97" box="[1331,1432,1087,1112]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
, this is actually an artefact in the image and does not really exist.
</paragraph>
<paragraph id="5E5936B33E464C3782F0FB0CFEDBF9C7" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
The female reported as
<taxonomicName id="99E64D303E464C3783E9FB0CFDB1FB6F" box="[478,586,1159,1184]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C3783E9FB0CFDB1FB6F" box="[478,586,1159,1184]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
undetermined D from Poring Hot Springs in
<bibRefCitation id="3A774B423E464C37866DFB02FA90FB6F" author="Dow" box="[1114,1387,1159,1184]" pageId="4" pageNumber="394" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012">Dow &amp; Orr (2012: 396)</bibRefCitation>
and listed as a
<typeStatus id="815D88113E464C37833BFB26FE97FB0A" box="[268,364,1197,1221]" pageId="4" pageNumber="394" type="paratype">paratype</typeStatus>
here, along with the other female not collected in tandem with a male, are considered to belong to
<taxonomicName id="99E64D303E464C378284FB5AFEB4FB27" box="[179,335,1232,1256]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E464C378284FB5AFEB4FB27" box="[179,335,1232,1256]" italics="true" pageId="4" pageNumber="394">T. fugispinosa</emphasis>
</taxonomicName>
, as males were found in the same area and no other species of
<taxonomicName id="99E64D303E464C37863BFB44FB83FB27" box="[1036,1144,1231,1256]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C37863BFB44FB83FB27" box="[1036,1144,1231,1256]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
has been found at Poring. However, in general the female may prove difficult to separate from those of other species of
<taxonomicName id="99E64D303E464C3786C4FB7FFAA4FAC2" box="[1267,1375,1268,1293]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C3786C4FB7FFAA4FAC2" box="[1267,1375,1268,1293]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
with similar-length antehumeral markings; the female of the only other species known to occur in Sabah,
<taxonomicName id="99E64D303E464C3786CDFA92FED3FA9B" authority="Dow &amp; Orr, 2012" authorityName="Dow &amp; Orr" authorityYear="2012" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="janeus">
<emphasis id="6C92EAA13E464C3786CDFA92FAA5FAFF" box="[1274,1374,1305,1328]" italics="true" pageId="4" pageNumber="394">T. janeus</emphasis>
<bibRefCitation id="3A774B423E464C378752FA92FED3FA9B" author="Dow" pageId="4" pageNumber="394" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012">Dow &amp; Orr, 2012</bibRefCitation>
</taxonomicName>
, is not yet known. The known distributions of
<taxonomicName id="99E64D303E464C37817CFAB6FC11FA9B" box="[843,1002,1340,1364]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E464C37817CFAB6FC11FA9B" box="[843,1002,1340,1364]" italics="true" pageId="4" pageNumber="394">T. fugispinosa</emphasis>
</taxonomicName>
and
<taxonomicName id="99E64D303E464C378611FAB6FB77FA9B" box="[1062,1164,1341,1364]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="janeus">
<emphasis id="6C92EAA13E464C378611FAB6FB77FA9B" box="[1062,1164,1341,1364]" italics="true" pageId="4" pageNumber="394">T. janeus</emphasis>
</taxonomicName>
are allopatric, with the former known from Mount Kinabalu and the Crocker range and the latter from the south and east of Sabah, with published records from the Danum Valley and Imbak Canyon (
<bibRefCitation id="3A774B423E464C378157FA0FFB98FA53" author="Dow" box="[864,1123,1412,1436]" pageId="4" pageNumber="394" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012">Dow &amp; Orr 2012: 394</bibRefCitation>
). The identity of
<taxonomicName id="99E64D303E464C378707FA0FFA67FA52" box="[1328,1436,1412,1437]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C378707FA0FFA67FA52" box="[1328,1436,1412,1437]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
undetermined C, reported from the Silau Silau stream near to the Kinabalu National Park headquarters (
<bibRefCitation id="3A774B423E464C378772FA23FD8AFA2A" author="Dow" pageId="4" pageNumber="394" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012" yearSuffix="a">Dow &amp; Orr 2012: 396; a large-sized teneral male</bibRefCitation>
) remains open, but it is likely to represent a third species of
<taxonomicName id="99E64D303E464C378707FA47FA67FA2A" box="[1328,1436,1484,1509]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C378707FA47FA67FA2A" box="[1328,1436,1484,1509]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
from Sabah.
</paragraph>
<paragraph id="5E5936B33E464C3782F0F99FFF1DF8CA" blockId="4.[151,1437,151,1797]" pageId="4" pageNumber="394">
Of the provisional species groups defined in
<bibRefCitation id="3A774B423E464C37808BF99EFC6AF9E3" author="Dow" box="[700,913,1556,1580]" pageId="4" pageNumber="394" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012">Dow &amp; Orr (2012)</bibRefCitation>
,
<taxonomicName id="99E64D303E464C3781ABF99EFBC2F9E3" box="[924,1081,1556,1580]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E464C3781ABF99EFBC2F9E3" box="[924,1081,1556,1580]" italics="true" pageId="4" pageNumber="394">T. fugispinosa</emphasis>
</taxonomicName>
fits best into the
<emphasis id="6C92EAA13E464C3786CDF99FFAB1F9E3" box="[1274,1354,1556,1580]" italics="true" pageId="4" pageNumber="394">feronia</emphasis>
-group. Using the key in
<bibRefCitation id="3A774B423E464C37836BF9B2FDCDF99F" author="Dow" box="[348,566,1592,1616]" pageId="4" pageNumber="394" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012">Dow &amp; Orr (2012)</bibRefCitation>
, exactly as in the case of
<taxonomicName id="99E64D303E464C37816AF9B2FB0AF99F" authority="Dow, 2014 (Dow 2014: 78)" authorityName="Dow, 2014 (Dow" authorityPageNumber="78" authorityYear="2014" box="[861,1265,1591,1616]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="iban">
<emphasis id="6C92EAA13E464C37816AF9B2FC57F99F" box="[861,940,1591,1616]" italics="true" pageId="4" pageNumber="394">T. iban</emphasis>
<bibRefCitation id="3A774B423E464C378183F9B2FBCBF99F" author="Dow" box="[948,1072,1592,1616]" pageId="4" pageNumber="394" refString="Dow, R. A. (2014) Telosticta iban sp. nov. from Sarawak (Odonata: Zygoptera: Platystictidae). Zootaxa, 3784 (1), 74 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3784.1.4" type="journal article" year="2014">Dow, 2014</bibRefCitation>
(
<bibRefCitation id="3A774B423E464C378677F9B3FB13F99F" author="Dow" box="[1088,1256,1592,1616]" pageId="4" pageNumber="394" refString="Dow, R. A. (2014) Telosticta iban sp. nov. from Sarawak (Odonata: Zygoptera: Platystictidae). Zootaxa, 3784 (1), 74 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3784.1.4" type="journal article" year="2014">Dow 2014: 78</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="99E64D303E464C3786C9F9B2FA67F99F" box="[1278,1436,1592,1616]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E464C3786C9F9B2FA67F99F" box="[1278,1436,1592,1616]" italics="true" pageId="4" pageNumber="394">T. fugispinosa</emphasis>
</taxonomicName>
would key out as
<taxonomicName id="99E64D303E464C378344F9D6FD46F9BB" authority="Dow &amp; Orr, 2012" authorityName="Dow &amp; Orr" authorityYear="2012" box="[371,701,1628,1653]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="dayak">
<emphasis id="6C92EAA13E464C378344F9D6FE2CF9BA" box="[371,471,1628,1653]" italics="true" pageId="4" pageNumber="394">T. dayak</emphasis>
<bibRefCitation id="3A774B423E464C3783D3F9D6FD46F9BB" author="Dow" box="[484,701,1628,1652]" pageId="4" pageNumber="394" refString="Dow, R. A. &amp; Orr, A. G. (2012) Telosticta, a new damselfly genus from Borneo and Palawan (Odonata: Zygoptera: Platystictidae). The Raffles Bulletin of Zoology, 60 (2), 361 - 397." type="journal article" year="2012">Dow &amp; Orr, 2012</bibRefCitation>
</taxonomicName>
, or break the key, depending on how liberally couplet 7 was interpreted. The key could most easily be modified to accommodate
<taxonomicName id="99E64D303E464C3781ACF90AFBCCF957" box="[923,1079,1664,1688]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E464C3781ACF90AFC53F957" box="[923,936,1665,1688]" italics="true" pageId="4" pageNumber="394">T</emphasis>
.
<emphasis id="6C92EAA13E464C378181F90BFBCCF957" box="[950,1079,1664,1688]" italics="true" pageId="4" pageNumber="394">fugispinosa</emphasis>
</taxonomicName>
by introducing a new couplet 6 distinguishing between species with the scoop of the paraproct sharply inturned, with vestigial spine (e.g.,
<taxonomicName id="99E64D303E464C3787B0F92EFEE3F92F" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="fugispinosa">
<emphasis id="6C92EAA13E464C3787B0F92EFEE3F92F" italics="true" pageId="4" pageNumber="394">T. fugispinosa</emphasis>
</taxonomicName>
) from all others. No other species included in
<taxonomicName id="99E64D303E464C37810FF94CFC5FF92F" box="[824,932,1735,1760]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="6C92EAA13E464C37810FF94CFC5FF92F" box="[824,932,1735,1760]" italics="true" pageId="4" pageNumber="394">Telosticta</emphasis>
</taxonomicName>
has the spine on the paraprocts vestigial or absent.
</paragraph>
</subSubSection>
</treatment>
</document>