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data/BC/05/16/BC05165FF3C45F5A87646872BED82206.xml
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<document id="D2D9C54764D9B0345B6ED1C81B8953EC" ID-DOI="10.3897/zookeys.1224.142859" ID-ZooBank="5793D52C-81D9-40A7-B544-DEAED1E62F34" ID-publisher-id="142859" URI-arpha="E0EE9F26-F761-5F62-B769-57C28098847F" URI-zoobank="http://zoobank.org/5793D52C-81D9-40A7-B544-DEAED1E62F34" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1738077822721" checkinUser="pensoft" docAuthor="Byeon, Ui-Joung & Park, Jong-Seok" docDate="2025" docId="BC05165FF3C45F5A87646872BED82206" docLanguage="en" docName="ZooKeys 1224: 165-174" docOrigin="ZooKeys 1224" docSource="https://zookeys.pensoft.net/article/142859/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Veraphis Casey 1897" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="174" masterDocId="E0EE9F26F7615F62B76957C28098847F" masterDocTitle="Two new species of the genus Veraphis Casey (Coleoptera, Staphylinidae, Scydmaeninae) from Korea" masterLastPageNumber="174" masterPageNumber="165" pageNumber="165" updateTime="1738080875092" updateUser="admin">
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<mods:mods id="75E8B9F3ECA5D90FBF46F1243B905797" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="735A57BB7F5848601DB97BCCB0EB50A4">
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<mods:title id="C7CF4FD4AC9EFEDE1D990031DF4F3CD3">Two new species of the genus Veraphis Casey (Coleoptera, Staphylinidae, Scydmaeninae) from Korea</mods:title>
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<mods:name id="E3CB31E77D3E2DFBE71C118BC1F8E669" type="personal">
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<mods:roleTerm id="72B518DD9E4575FA3EA2805BDB1006CA">Author</mods:roleTerm>
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<mods:namePart id="B86BDA928B992C5560F11C4B7B00AC02">Byeon, Ui-Joung</mods:namePart>
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<mods:affiliation id="5C9F39ADFD4F2105AEDA9E287BA39A57">Department of Biological Sciences and Biotechnology, Chungbuk National University, 1 Chungdae-ro, Seowon-gu, Cheongju-si, Chungbuk-do, 28644, Republic of Korea</mods:affiliation>
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<mods:namePart id="9AB951608EF9D51B511C851E31032CED">Park, Jong-Seok</mods:namePart>
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<mods:nameIdentifier id="B91BD7A29D41B6C91A6A418DD2985E47" type="ORCID">0000-0003-2063-2534</mods:nameIdentifier>
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<mods:affiliation id="3F54385DD564CEEC9D17D7A261F0F6C8">Department of Biological Sciences and Biotechnology, Chungbuk National University, 1 Chungdae-ro, Seowon-gu, Cheongju-si, Chungbuk-do, 28644, Republic of Korea</mods:affiliation>
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<mods:title id="C77DFF05569ACF7F0B50E946B769525E">ZooKeys</mods:title>
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<mods:part id="27A7D86EE7EFCB34D0C096F935032AEF">
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<mods:date id="E63DBED908A3E7261975161FC1B90CAB">2025</mods:date>
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<mods:detail id="55EFBEFB32A40CDD7D5473C595D80E1F" type="pubDate">
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<mods:number id="B03963C6AC4871F78677C3144AACA29F">2025-01-28</mods:number>
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<mods:detail id="94AA6AF24327CB4F7D1B1F7D26B733BC" type="volume">
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<mods:number id="A8F706A13F4FB524FFA85A57B6F95F5C">1224</mods:number>
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<mods:identifier id="C277ABE5211A811718B9C349218B71F2" type="DOI">10.3897/zookeys.1224.142859</mods:identifier>
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<mods:identifier id="9089E482841D5D3F24C65F4735FDE058" type="ZooBank">5793D52C-81D9-40A7-B544-DEAED1E62F34</mods:identifier>
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<treatment id="BC05165FF3C45F5A87646872BED82206" ID-DOI="http://doi.org/10.5281/zenodo.14755816" ID-Zenodo-Dep="14755816" ID-arpha="BC05165F-F3C4-5F5A-8764-6872BED82206" LSID="urn:lsid:plazi:treatment:BC05165FF3C45F5A87646872BED82206" httpUri="http://treatment.plazi.org/id/BC05165FF3C45F5A87646872BED82206" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Scydmaeninae">
|
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<subSubSection id="7494B56C72893C2EC53191C64CFFF450" type="nomenclature">
|
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<paragraph id="3C37EA4B7BECEDD752675AA6FC992511">
|
||||
<heading id="8FF88B34CD12815722116B82BDEA3A61" reason="label">Genus</heading>
|
||||
<taxonomicName id="9A61F7A4D23E29D969A070AB64DB2A7B" ID-CoL="8VS6J" authority="Casey, 1897" authorityName="Casey" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A39A2043EBBEC1D65EFC262AC4343A4B" italics="true">Veraphis</emphasis>
|
||||
<bibRefCitation id="4434347F73B2E21549DCECD5DFD8AC4E" DOI="10.1111/j.1749-6632.1896.tb55435.x" author="Casey TL" issue="1" journalOrPublisher="Annals of the New York Academy of Sciences" pagination="285-684" refId="B1" refString="Casey TL (1897) Coleopterological notices. VII. Annals of the New York Academy of Sciences 9 (1): 285 - 684. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" title="Coleopterological notices. VII." volume="9" year="1897">Casey, 1897</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="7256287267E703B3A1D36BC5EAF78F7E" type="reference_group">
|
||||
<paragraph id="4E99E5AEF487E4E31A0DB5482AF65E8A">
|
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<treatmentCitationGroup id="81DB063BCF086C69D8F5BA69C4C1D5C0">
|
||||
<treatmentCitation id="82FDC7626DF9B8E33AD6AF12D826D879" author="Casey TL" page="509" year="1897">
|
||||
<taxonomicName id="BAE1F33E21C472A8C3EB42D0EF5D2E6E" ID-CoL="8VS6J" authority="Casey, 1897: 509" authorityName="Casey" authorityPageNumber="509" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="5887EDE0A832747FC660DDAE9C6BA964" italics="true">Veraphis</emphasis>
|
||||
<comment id="F87E1A0454D5A3FCC814A60903AA6EA0">
|
||||
<bibRefCitation id="893CE6B7CA946B2C10FB2CF3BD96AFC1" DOI="10.1111/j.1749-6632.1896.tb55435.x" author="Casey TL" issue="1" journalOrPublisher="Annals of the New York Academy of Sciences" pagination="285-684" refId="B1" refString="Casey TL (1897) Coleopterological notices. VII. Annals of the New York Academy of Sciences 9 (1): 285 - 684. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" title="Coleopterological notices. VII." volume="9" year="1897">Casey, 1897: 509</bibRefCitation>
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||||
. Type species:
|
||||
<treatmentCitation id="26E80463C23549781A1D413A09F58576">
|
||||
<taxonomicName id="51901CCC0CA76BB2C444A794E4C20618" authority="LeConte, 1879" authorityName="LeConte" authorityYear="1879" class="Insecta" family="Staphylinidae" genus="Eutheia" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="impressa">
|
||||
<emphasis id="2D786C31644DE856756AEB9809DA55D6" italics="true">Eutheia impressa</emphasis>
|
||||
<bibRefCitation id="CDBE8B5CFD4EA5E1D8B85B4F1C308BF5" author="LeConte JL" issue="3" journalOrPublisher="Bulletin of the United States Geological and Geographical Survey of the Territories" pagination="331-520" refId="B14" refString="LeConte JL (1879) The Coleoptera of the alpine Rocky Mountain regions. Part II. Bulletin of the United States Geological and Geographical Survey of the Territories 5 (3): 331 - 520." title="The Coleoptera of the alpine Rocky Mountain regions. Part II." volume="5" year="1879">LeConte, 1879</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
(designated by Franz in
|
||||
<treatmentCitation id="90641575469866908CA9A2F62B075F13">
|
||||
<bibRefCitation id="00E0AC2C41670495CC25EBDE727391D5" author="Newton and Franz" firstAuthor="Newton" journalOrPublisher="Koleopterologische Rundschau" pagination="137-165" refId="B15" refString="Newton A, Franz H (1998) World catalog of the genera of Scydmaenidae. Koleopterologische Rundschau 68: 137–165." title="World catalog of the genera of Scydmaenidae." volume="68" year="1998">Newton and Franz 1998</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
).
|
||||
</comment>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EBJAC" type="diagnosis">
|
||||
<paragraph id="81BE8398171BDDB7337AB1C23CCD44EA">
|
||||
<heading id="51BA7062B1C42F5D6B7C70814B59F091" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="AD0CE1DE3F45BF9B45DCF8C54232DB98">
|
||||
This genus is distinguished from other genera of
|
||||
<taxonomicName id="8F852B714E880C57672859E13078845F" authorityName="Leach" authorityYear="1815" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
|
||||
by the following combination characteristics: body flattened and elongated (Figs
|
||||
<figureCitation id="C47A3AC353965156EB94C8BFF3B237AB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Veraphis odaesanensis sp. nov. A male habitus, dorsal view B male head and pronotum, dorsal view C male protibia D female protibia E meso-, metaventrite." figureDoi="10.3897/zookeys.1224.142859.figure1" httpUri="https://binary.pensoft.net/fig/1236352">1 A</figureCitation>
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||||
,
|
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<figureCitation id="52C56867B06CCE8E8132665BB83EE32F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Veraphis myeonggiensis sp. nov. A male habitus, dorsal view B male head and pronotum, dorsal view C male protibia D meso-, metaventrite." figureDoi="10.3897/zookeys.1224.142859.figure2" httpUri="https://binary.pensoft.net/fig/1236353">2 A</figureCitation>
|
||||
); procoxal cavities broadly open (Fig.
|
||||
<figureCitation id="DA25A023A4CA1717D0759982B49C95C3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Veraphis odaesanensis sp. nov. A male habitus, dorsal view B male head and pronotum, dorsal view C male protibia D female protibia E meso-, metaventrite." figureDoi="10.3897/zookeys.1224.142859.figure1" httpUri="https://binary.pensoft.net/fig/1236352">1 E</figureCitation>
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||||
); prosternum with a narrow intercoxal carina (Fig.
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||||
<figureCitation id="70708635CE5A20E0F48EDA688E81334A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Veraphis odaesanensis sp. nov. A male habitus, dorsal view B male head and pronotum, dorsal view C male protibia D female protibia E meso-, metaventrite." figureDoi="10.3897/zookeys.1224.142859.figure1" httpUri="https://binary.pensoft.net/fig/1236352">1 E</figureCitation>
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||||
); each elytron with a setose basal fovea (Figs
|
||||
<figureCitation id="D313054537042D37E408CA20A63C1E1D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Veraphis odaesanensis sp. nov. A male habitus, dorsal view B male head and pronotum, dorsal view C male protibia D female protibia E meso-, metaventrite." figureDoi="10.3897/zookeys.1224.142859.figure1" httpUri="https://binary.pensoft.net/fig/1236352">1 B</figureCitation>
|
||||
,
|
||||
<figureCitation id="CCB89BE272897748B7760D3850D1BA10" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Veraphis myeonggiensis sp. nov. A male habitus, dorsal view B male head and pronotum, dorsal view C male protibia D meso-, metaventrite." figureDoi="10.3897/zookeys.1224.142859.figure2" httpUri="https://binary.pensoft.net/fig/1236353">2 B</figureCitation>
|
||||
). For more detailed diagnostic characters and phylogenetic information on this genus, refer to
|
||||
<bibRefCitation id="9B957156A050FA716287BF278D22BB97" DOI="10.1111/syen.12044" author="Jałoszyński P" issue="1" journalOrPublisher="Systematic Entomology" pagination="159-189" refId="B8" refString="Jałoszyński P (2014 b) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology 39 (1): 159–189. https://doi.org/10.1111/syen.12044" title="Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae)." volume="39">Jałoszyński (2014 b</bibRefCitation>
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||||
).
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||||
</paragraph>
|
||||
<caption id="E07C7EC088C527965531F36E28C93877" ID-DOI="10.3897/zookeys.1224.142859.figure1" ID-arpha="890BE1CC-8453-5F76-8338-16B31D6E7F3E" httpUri="https://binary.pensoft.net/fig/1236352" startId="F1">
|
||||
<paragraph id="85C63EE944074FFE7714A8EEF6691BD6">
|
||||
<label id="BA47815464D64EC10ADE73F33A214AEE">Figure 1.</label>
|
||||
</paragraph>
|
||||
<paragraph id="48061B734F49222F373F03528C4D3CEB">
|
||||
<taxonomicName id="A229949DABB6C4A384DCE27563F546EC" authorityName="Byeon & Park" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="odaesanensis" status="sp. nov.">
|
||||
<emphasis id="0A4008DCD350474CCAD6AA24C0D730AA" italics="true">Veraphis odaesanensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="BC6D88B46DE818BEC99481362C2BD12C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="15A901C1ADED6C1A9EAD552CC8D90C93" bold="true">A</emphasis>
|
||||
male habitus, dorsal view
|
||||
<emphasis id="743C0C99457815585595FBB15C01D3EC" bold="true">B</emphasis>
|
||||
male head and pronotum, dorsal view
|
||||
<emphasis id="37B7E0F664C45BEB320B34106A01254D" bold="true">C</emphasis>
|
||||
male protibia
|
||||
<emphasis id="75F59930686E1783FC832473313220C6" bold="true">D</emphasis>
|
||||
female protibia
|
||||
<emphasis id="C74471062F167816EC42B23C3645DE49" bold="true">E</emphasis>
|
||||
meso-, metaventrite.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="3FEF14CCDF1D9CB5A59E20891DB0D5F0" ID-DOI="10.3897/zookeys.1224.142859.figure2" ID-arpha="A6D9BC8B-4857-5A07-9A06-358D7E971AC8" httpUri="https://binary.pensoft.net/fig/1236353" startId="F2">
|
||||
<paragraph id="7254B39FE29C64501EB33DD60F61B180">
|
||||
<label id="498FF3C6D7F409B5B2F9A6A5B581D2A9">Figure 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="4FD155AD9E94923AE27E34F4CE0C2CCB">
|
||||
<taxonomicName id="E385D193E2F3CDDC223A08AB01234E5C" authorityName="Byeon & Park" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="myeonggiensis" status="sp. nov.">
|
||||
<emphasis id="70F145586E5BBB6BA773A74B9B694A1F" italics="true">Veraphis myeonggiensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="AC027DA34D7F939BCF0E995D34CD0AF0" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="B0343951E0BADD0C21673DA706F66107" bold="true">A</emphasis>
|
||||
male habitus, dorsal view
|
||||
<emphasis id="E394E2687BEE99F4E8687612CEAB1C1B" bold="true">B</emphasis>
|
||||
male head and pronotum, dorsal view
|
||||
<emphasis id="70C0BD703B5C7092815F0486729DE5D7" bold="true">C</emphasis>
|
||||
male protibia
|
||||
<emphasis id="CE480EA132329349ED044FAE81E8BF85" bold="true">D</emphasis>
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meso-, metaventrite.
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</paragraph>
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||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EKMAC" type="distribution">
|
||||
<paragraph id="C1BB09484B01F5DB3417B6B48C43BE5C">
|
||||
<heading id="105E79ECC9F7B23F65E6418A19BB3507" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="627CD6BDA0D6E3AEB5748289F119DFE6">
|
||||
Northern Hemisphere (
|
||||
<collectingCountry id="DD41657FFE3AD7C005D5009AADE1AAF9" name="China">China</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="27F45F38921E012F4191AD93323FFF73" name="Japan">Japan</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="ACD92F6F19C9CC699B7F189EAF4BCF7E" name="South Korea">Korea</collectingCountry>
|
||||
,
|
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<collectingCountry id="4A44788C04394547E50B2105403EA0A9" name="United States of America">USA</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="A443B01E8B289193B0074749569E0498" name="Finland">Finland</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="1FA06FBACD54D360DC6E324319750106" name="Sweden">Sweden</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="3BEAF6A1AB345C4F48B94226CF34C980" name="Mongolia">Mongolia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="96E7B9DE25A327DC65958D7DB2690D4B" name="Russia">Russia</collectingCountry>
|
||||
(Far East, East Siberia )).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPMAC" type="remarks">
|
||||
<paragraph id="D8AE1C237D5F1D78318817D75266835F">
|
||||
<heading id="580ECD67A2F625A2F9A1CD644C2FE073" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B52AF75ECC658D13926D57EFDD0B1ACA">
|
||||
According to
|
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<bibRefCitation id="C31EA661B02A0195097000E40C764EA3" DOI="10.11646/zootaxa.5507.1.11" author="Jaloszynski" firstAuthor="Jaloszynski" issue="1" journalOrPublisher="Zootaxa" pagination="194-200" refId="B10" refString="Jałoszyński P (2024) Two new Veraphis species in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5507 (1): 194–200. https://doi.org/10.11646/zootaxa.5507.1.11" title="Two new Veraphis species in China (Coleoptera, Staphylinidae, Scydmaeninae)." volume="5507" year="2024">Jałoszyński 2024</bibRefCitation>
|
||||
, this genus is primarily found in leaf litter and soil, and all species discovered in
|
||||
<collectingCountry id="8DD1D89E314F038DEF16F33E20645748" name="China">China</collectingCountry>
|
||||
inhabit alpine regions at altitudes above
|
||||
<quantity id="CC0E817E112449C67DFE3A503EC7A619" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</quantity>
|
||||
. Additionally, species found in
|
||||
<collectingCountry id="AB8E451F1CC065DADD02630436DA9812" name="Japan">Japan</collectingCountry>
|
||||
are also thought to be distributed in cooler climates. These species are very similar in external morphology, making examination of the aedeagus crucial for species identification. In some species, modifications of the male legs are present, which can be useful for distinguishing species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
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