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<mods:title id="357919F4EA7444DF1616758F21112F59">A new species of Anadia (Reptilia, Squamata) from the Venezuelan ' Lost World', northern South America</mods:title>
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<mods:title id="21A71962AA2EE1A51BD47AFCCB519E97">A new species of Anadia (Reptilia, Squamata) from the Venezuelan ' Lost World', northern South America</mods:title>
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<mods:namePart id="09D1EAC66CE920BDFA9FE437053B27C4">Kok, Philippe J. R.</mods:namePart>
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<mods:namePart id="2985C6DC0F0A67A2DE7BAC7CCF523B78">Kok, Philippe J. R.</mods:namePart>
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@ -53,7 +55,7 @@
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<paragraph id="BA6D36F06162FFC7FD774405FCC7FAC9" blockId="2.[628,960,1323,1389]" box="[695,893,1363,1389]" pageId="2" pageNumber="3">
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<figureCitation id="22E92A756162FFC7FD774405FCA3FAC9" box="[695,793,1363,1389]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="3.[189,232,1895,1921]" captionStartId-1="4.[189,232,1965,1991]" captionStartId-2="5.[189,232,1930,1956]" captionStartId-3="6.[189,232,1824,1850]" captionStartId-4="8.[189,232,1930,1956]" captionStartId-5="9.[189,232,1930,1956]" captionTargetBox-0="[207,1380,265,1855]" captionTargetBox-1="[189,1372,1072,1925]" captionTargetBox-2="[189,1399,467,1890]" captionTargetBox-3="[189,1399,194,1787]" captionTargetBox-4="[248,1389,501,1830]" captionTargetBox-5="[219,1380,874,1814]" captionTargetId-0="figure@3.[207,1380,265,1855]" captionTargetId-1="figure@4.[189,1398,1072,1925]" captionTargetId-2="figure@5.[189,1399,467,1890]" captionTargetId-4="figure@8.[189,1398,462,1889]" captionTargetId-5="figure@9.[207,1382,855,1890]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." captionText-1="Fig. 2. Anadia mcdiarmidi sp. nov. Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Scale lines = 5 mm. Photographs by Philippe J.R. Kok." captionText-2="Fig. 3. Anadia mcdiarmidi sp. nov. Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok." captionText-3="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." captionText-4="Fig. 6. Anadia mcdiarmidi sp. nov. Intrapopulation variation in dorsal (above) and ventral (below) colour pattern in preserved specimens. From left to right: IRSNB 2677, IRSNB 2674, IRSNB 2676, IRSNB 2675. Photographs by Philippe J.R. Kok." captionText-5="Fig. 7. Anadia mcdiarmidi sp. nov. Intrapopulation variation in the condition of the parietals and the interparietal (frontoparietals in grey). Top, from left to right: IRSNB 2677, IRSNB 2676; bottom, from left to right: IRSNB 2675, IRSNB 2674. Drawings not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.3711437" figureDoi-1="http://doi.org/10.5281/zenodo.3711439" figureDoi-2="http://doi.org/10.5281/zenodo.3711441" figureDoi-3="http://doi.org/10.5281/zenodo.3711443" figureDoi-4="http://doi.org/10.5281/zenodo.3711447" figureDoi-5="http://doi.org/10.5281/zenodo.3711449" httpUri-0="https://zenodo.org/record/3711437/files/figure.png" httpUri-1="https://zenodo.org/record/3711439/files/figure.png" httpUri-2="https://zenodo.org/record/3711441/files/figure.png" httpUri-3="https://zenodo.org/record/3711443/files/figure.png" httpUri-4="https://zenodo.org/record/3711447/files/figure.png" httpUri-5="https://zenodo.org/record/3711449/files/figure.png" pageId="2" pageNumber="3">Figs 1-7</figureCitation>
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<figureCitation id="22E92A756162FFC7FD774405FCA3FAC9" box="[695,793,1363,1389]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="3.[189,232,1895,1921]" captionStartId-1="4.[189,232,1965,1991]" captionStartId-2="5.[189,232,1930,1956]" captionStartId-3="6.[189,232,1000,1026]" captionStartId-4="6.[189,232,1824,1850]" captionStartId-5="8.[189,232,1930,1956]" captionStartId-6="9.[189,232,1930,1956]" captionTargetBox-0="[207,1380,265,1855]" captionTargetBox-1="[189,1372,1072,1925]" captionTargetBox-2="[189,1399,467,1890]" captionTargetBox-3="[189,1399,1085,1787]" captionTargetBox-4="[189,1399,1085,1787]" captionTargetBox-5="[248,1389,501,1830]" captionTargetBox-6="[219,1380,874,1814]" captionTargetId-0="figure@3.[207,1380,265,1855]" captionTargetId-1="figure@4.[189,1398,1072,1925]" captionTargetId-2="figure@5.[189,1399,467,1890]" captionTargetId-3="figure@6.[189,1399,1099,1787]" captionTargetId-4="figure@6.[189,1399,1099,1787]" captionTargetId-5="figure@8.[189,1398,462,1889]" captionTargetId-6="figure@9.[207,1382,855,1890]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." captionText-1="Fig. 2. Anadia mcdiarmidi sp. nov. Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Scale lines = 5 mm. Photographs by Philippe J.R. Kok." captionText-2="Fig. 3. Anadia mcdiarmidi sp. nov. Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok." captionText-3="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." captionText-4="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." captionText-5="Fig. 6. Anadia mcdiarmidi sp. nov. Intrapopulation variation in dorsal (above) and ventral (below) colour pattern in preserved specimens. From left to right: IRSNB 2677, IRSNB 2674, IRSNB 2676, IRSNB 2675. Photographs by Philippe J.R. Kok." captionText-6="Fig. 7. Anadia mcdiarmidi sp. nov. Intrapopulation variation in the condition of the parietals and the interparietal (frontoparietals in grey). Top, from left to right: IRSNB 2677, IRSNB 2676; bottom, from left to right: IRSNB 2675, IRSNB 2674. Drawings not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.3711437" figureDoi-1="http://doi.org/10.5281/zenodo.3711439" figureDoi-2="http://doi.org/10.5281/zenodo.3711441" figureDoi-3="http://doi.org/10.5281/zenodo.3711443" figureDoi-4="http://doi.org/10.5281/zenodo.3711443" figureDoi-5="http://doi.org/10.5281/zenodo.3711447" figureDoi-6="http://doi.org/10.5281/zenodo.3711449" httpUri-0="https://zenodo.org/record/3711437/files/figure.png" httpUri-1="https://zenodo.org/record/3711439/files/figure.png" httpUri-2="https://zenodo.org/record/3711441/files/figure.png" httpUri-3="https://zenodo.org/record/3711443/files/figure.png" httpUri-4="https://zenodo.org/record/3711443/files/figure.png" httpUri-5="https://zenodo.org/record/3711447/files/figure.png" httpUri-6="https://zenodo.org/record/3711449/files/figure.png" pageId="2" pageNumber="3">Figs 1-7</figureCitation>
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||||
,
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<tableCitation id="F750034B6162FFC7FCE64405FCC7FAC9" box="[806,893,1363,1389]" captionStart="Table 1" captionStartId="13.[189,258,822,848]" captionText="Table 1. Sex, measurements (in mm) and scale counts of the type series of Anadia mcdiarmidi sp. nov. 1 = tail incomplete or obviously regenerated, 2 = data corresponding to right/left side, other abbreviations are defined in the text." pageId="2" pageNumber="3">Table 1</tableCitation>
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||||
</paragraph>
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@ -112,29 +114,24 @@ sp. A –
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<paragraph id="BA6D36F06162FFC7FF7D4705FE95F9C9" blockId="2.[189,1399,1619,1724]" box="[189,303,1619,1645]" pageId="2" pageNumber="3">
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<paragraph id="BA6D36F06162FFC7FF7D4705FA88F91F" blockId="2.[189,1399,1619,1724]" pageId="2" pageNumber="3">
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FF7D4705FA88F91F" ID-GBIF-Occurrence="2575931620" collectingDate="2011-05-03" collectionCode="R" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2201" latitude="5.1858335" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.29333" pageId="2" pageNumber="3" specimenCode="IRSNB 2677" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="holotype">
|
||||
<heading id="E125819C6162FFC7FF7D4705FE95F9C9" bold="true" box="[189,303,1619,1645]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||||
<emphasis id="88A6EAE26162FFC7FF7D4705FE95F9C9" bold="true" box="[189,303,1619,1645]" pageId="2" pageNumber="3">
|
||||
<typeStatus id="656988526162FFC7FF7D4705FE95F9C9" box="[189,303,1619,1645]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BA6D36F06162FFC7FF7D472CFA88F91F" blockId="2.[189,1399,1619,1724]" pageId="2" pageNumber="3">
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FF7D472CFA88F91F" ID-GBIF-Occurrence="2575931620" collectingDate="2011-05-03" collectionCode="R" collectorName="Philippe J." country="Venezuela" elevation="2201" latitude="5.1858335" location="Abakapa-tepui" longLatPrecision="21" longitude="-62.29333" pageId="2" pageNumber="3" specimenCode="IRSNB 2677, PK 3565" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" stateProvince="Bolivar" typeStatus="holotype">
|
||||
<specimenCode id="EA749E8B6162FFC7FF7D472CFEEDF930" box="[189,343,1658,1684]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="3">IRSNB 2677</specimenCode>
|
||||
(field number
|
||||
<specimenCode id="EA749E8B6162FFC7FDC6472CFDC9F931" box="[518,627,1658,1685]" collectionCode="PK" pageId="2" pageNumber="3">PK 3565</specimenCode>
|
||||
),
|
||||
<specimenCount id="ACD4FD796162FFC7FD48472CFD60F930" box="[648,730,1658,1684]" pageId="2" pageNumber="3" type="adult">1 adult</specimenCount>
|
||||
<specimenCount id="ACD4FD796162FFC7FD22472CFD42F930" box="[738,760,1658,1684]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
|
||||
(field number PK 3565),
|
||||
<specimenCount id="ACD4FD796162FFC7FD48472CFD42F930" box="[648,760,1658,1684]" pageId="2" pageNumber="3">1 adult ♂</specimenCount>
|
||||
collected by
|
||||
<collectorName id="172753266162FFC7FC58472CFBA9F930" box="[920,1043,1658,1684]" pageId="2" pageNumber="3">Philippe J.</collectorName>
|
||||
R. Kok,
|
||||
<collectorName id="172753266162FFC7FC58472CFBD6F930" box="[920,1132,1658,1684]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||||
,
|
||||
<date id="CE6C10306162FFC7FBBA472CFAB8F931" box="[1146,1282,1658,1685]" pageId="2" pageNumber="3" value="2011-05-03">
|
||||
<collectingDate id="DE28E9D86162FFC7FBBA472CFAB8F931" box="[1146,1282,1658,1685]" pageId="2" pageNumber="3" value="2011-05-03">03-05-2011</collectingDate>
|
||||
</date>
|
||||
at 11h39, summit of
|
||||
<location id="BF0D602B6162FFC7FEFA47F7FE50F91F" LSID="urn:lsid:plazi:treatment:327B87E66162FFCBFDB4447DFC5CFCEB:BF0D602B6162FFC7FEFA47F7FE50F91F" box="[314,490,1697,1723]" country="Venezuela" latitude="5.1858335" longLatPrecision="21" longitude="-62.29333" name="Abakapa-tepui" pageId="2" pageNumber="3" stateProvince="Bolivar">Abakapá-tepui</location>
|
||||
at 11h39,
|
||||
<location id="BF0D602B6162FFC7FF7D47F7FE50F91F" LSID="urn:lsid:plazi:treatment:327B87E66162FFCBFDB4447DFC5CFCEB:BF0D602B6162FFC7FF7D47F7FE50F91F" box="[189,490,1697,1723]" country="Venezuela" latitude="5.1858335" longLatPrecision="21" longitude="-62.29333" name="summit of Abakapa-tepui" pageId="2" pageNumber="3" stateProvince="Bolivar State">summit of Abakapá-tepui</location>
|
||||
,
|
||||
<collectingRegion id="7816F8126162FFC7FE3647F7FD29F91F" box="[502,659,1697,1723]" country="Venezuela" name="Bolivar" pageId="2" pageNumber="3">Bolívar State</collectingRegion>
|
||||
,
|
||||
|
|
@ -149,21 +146,19 @@ at 11h39, summit of
|
|||
elevation.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="BA6D36F06162FFC7FF7D47B0FEC8F8A4" blockId="2.[189,1399,1766,2027]" box="[189,370,1766,1792]" pageId="2" pageNumber="3">
|
||||
<paragraph id="BA6D36F06162FFC7FF7D47B0FACDF84F" blockId="2.[189,1399,1766,2027]" pageId="2" pageNumber="3">
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FF7D47B0FACCF8EB" ID-GBIF-Occurrence="2575931621" collectingDate="2011-05-06" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2242" latitude="5.1805553" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.296944" pageId="2" pageNumber="3" specimenCode="IRSNB 2674" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="paratopotypes">
|
||||
<heading id="E125819C6162FFC7FF7D47B0FEC8F8A4" bold="true" box="[189,370,1766,1792]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||||
<emphasis id="88A6EAE26162FFC7FF7D47B0FEC8F8A4" bold="true" box="[189,370,1766,1792]" pageId="2" pageNumber="3">Paratopotypes</emphasis>
|
||||
<emphasis id="88A6EAE26162FFC7FF7D47B0FEC8F8A4" bold="true" box="[189,370,1766,1792]" pageId="2" pageNumber="3">
|
||||
<typeStatus id="656988526162FFC7FF7D47B0FEC8F8A4" box="[189,370,1766,1792]" pageId="2" pageNumber="3">Paratopotypes</typeStatus>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BA6D36F06162FFC7FF7D465BFACDF84F" blockId="2.[189,1399,1766,2027]" pageId="2" pageNumber="3">
|
||||
(n=3)
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FEC7465BFACCF8EB" ID-GBIF-Occurrence="2575931621" collectingDate="2011-05-06" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="0" latitude="5.1805553" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.296944" pageId="2" pageNumber="3" specimenCode="IRSNB 2674, PK 3578" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="paratopotype">
|
||||
<specimenCount id="ACD4FD796162FFC7FEC7465BFEE0F883" box="[263,346,1805,1831]" pageId="2" pageNumber="3" type="adult">1 adult</specimenCount>
|
||||
<specimenCount id="ACD4FD796162FFC7FEA34658FEC3F88C" box="[355,377,1806,1832]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
|
||||
(
|
||||
<specimenCode id="EA749E8B6162FFC7FE4A465BFD93F883" box="[394,553,1805,1831]" pageId="2" pageNumber="3">IRSNB 2674</specimenCode>
|
||||
, field number
|
||||
<specimenCode id="EA749E8B6162FFC7FD164658FCFBF88C" box="[726,833,1805,1832]" pageId="2" pageNumber="3">PK 3578</specimenCode>
|
||||
) collected by
|
||||
, field number PK 3578) collected by
|
||||
<collectorName id="172753266162FFC7FC2C465BFB78F883" box="[1004,1218,1805,1832]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||||
,
|
||||
<date id="CE6C10306162FFC7FB10465BFAE2F88C" box="[1232,1368,1805,1832]" pageId="2" pageNumber="3" value="2011-05-06">
|
||||
|
|
@ -180,20 +175,17 @@ at 9h40,
|
|||
<geoCoordinate id="DFE650376162FFC7FC334662FB2BF8EA" box="[1011,1169,1844,1870]" degrees="062" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="49" value="-62.296944">062°17'49''W</geoCoordinate>
|
||||
,
|
||||
<quantity id="7D2A9B156162FFC7FB5C4662FB49F8EB" box="[1180,1267,1844,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.242" pageId="2" pageNumber="3" unit="m" value="2242.0">
|
||||
<elevation id="31FFD1C36162FFC7FB5C4662FB6DF8EA" box="[1180,1239,1844,1870]" pageId="2" pageNumber="3">2242</elevation>
|
||||
m
|
||||
<elevation id="31FFD1C36162FFC7FB5C4662FB49F8EB" box="[1180,1267,1844,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.242" pageId="2" pageNumber="3" unit="m" value="2242.0">2242 m</elevation>
|
||||
</quantity>
|
||||
elevation);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FF7D460AFA84F838" ID-GBIF-Occurrence="2575931622" collectingDate="2011-05-06" collectionCode="IRSNB, PK" collectorName="by Philippe J. R. Kok" country="Venezuela" elevation="2200" latitude="5.1833334" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.292778" pageId="2" pageNumber="3" specimenCode="IRSNB 2675, PK 3577" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="paratopotype">
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FF7D460AFA84F838" ID-GBIF-Occurrence="2575931622" collectingDate="2011-05-06" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2200" latitude="5.1833334" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.292778" pageId="2" pageNumber="3" specimenCode="IRSNB 2675" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="paratopotype">
|
||||
<specimenCount id="ACD4FD796162FFC7FF7D460AFE8BF8D1" box="[189,305,1883,1910]" pageId="2" pageNumber="3" type="juvenile">1 juvenile</specimenCount>
|
||||
<specimenCount id="ACD4FD796162FFC7FEF8460AFEF4F8D2" box="[312,334,1884,1910]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
|
||||
(
|
||||
<specimenCode id="EA749E8B6162FFC7FE9D460AFE40F8D2" box="[349,506,1883,1910]" pageId="2" pageNumber="3">IRSNB 2675</specimenCode>
|
||||
, field number
|
||||
<specimenCode id="EA749E8B6162FFC7FD60460AFCB2F8D2" box="[672,776,1883,1910]" pageId="2" pageNumber="3">PK 3577</specimenCode>
|
||||
) collected
|
||||
<collectorName id="172753266162FFC7FC48460DFBC4F8D1" box="[904,1150,1883,1910]" pageId="2" pageNumber="3">by Philippe J.R. Kok</collectorName>
|
||||
, field number PK 3577) collected by
|
||||
<collectorName id="172753266162FFC7FC6C460DFBC4F8D1" box="[940,1150,1883,1910]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||||
,
|
||||
<date id="CE6C10306162FFC7FB4A460DFAA8F8D2" box="[1162,1298,1883,1910]" pageId="2" pageNumber="3" value="2011-05-06">
|
||||
<collectingDate id="DE28E9D86162FFC7FB4A460DFAA8F8D2" box="[1162,1298,1883,1910]" pageId="2" pageNumber="3" value="2011-05-06">06-05-2011</collectingDate>
|
||||
|
|
@ -214,15 +206,11 @@ at 9h10,
|
|||
elevation
|
||||
</materialsCitation>
|
||||
; and
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FF7D46FCFACDF84F" ID-GBIF-Occurrence="2575931619" collectingDate="2011-05-09" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2204" latitude="5.185278" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.288612" pageId="2" pageNumber="3" specimenCode="IRSNB 2676, PK 3589" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Bolivar State">
|
||||
1 other
|
||||
<specimenCount id="ACD4FD796162FFC7FED646FCFECEF860" box="[278,372,1962,1988]" pageId="2" pageNumber="3" type="juvenile">juvenile</specimenCount>
|
||||
<specimenCount id="ACD4FD796162FFC7FEBC46FCFE28F860" box="[380,402,1962,1988]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
|
||||
<materialsCitation id="0ABA3CAD6162FFC7FF7D46FCFACDF84F" ID-GBIF-Occurrence="2575931619" collectingDate="2011-05-09" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2204" latitude="5.185278" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.288612" pageId="2" pageNumber="3" specimenCode="IRSNB 2676" specimenCount="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Bolivar State">
|
||||
<specimenCount id="ACD4FD796162FFC7FF7D46FCFE28F860" box="[189,402,1962,1988]" pageId="2" pageNumber="3">1 other juvenile ♂</specimenCount>
|
||||
(
|
||||
<specimenCode id="EA749E8B6162FFC7FE6146FCFD84F867" box="[417,574,1961,1988]" pageId="2" pageNumber="3">IRSNB 2676</specimenCode>
|
||||
, field number
|
||||
<specimenCode id="EA749E8B6162FFC7FD2646FCFCF4F860" box="[742,846,1961,1988]" pageId="2" pageNumber="3">PK 3589</specimenCode>
|
||||
) collected by
|
||||
, field number PK 3589) collected by
|
||||
<collectorName id="172753266162FFC7FC3346FCFB7CF860" box="[1011,1222,1962,1988]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||||
,
|
||||
<date id="CE6C10306162FFC7FB1246FFFAE0F860" box="[1234,1370,1961,1988]" pageId="2" pageNumber="3" value="2011-05-09">
|
||||
|
|
@ -262,7 +250,7 @@ elevation.
|
|||
. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="F2C8657B6164FFC2FF7D405DFA82FAA8" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="etymology">
|
||||
<subSubSection id="F2C8657B6164FFC1FF7D405DFD44FD6D" pageId="4" pageNumber="5" type="etymology">
|
||||
<paragraph id="BA6D36F06164FFC1FF7D405DFEFEFE81" blockId="4.[189,1399,267,399]" box="[189,324,267,293]" pageId="4" pageNumber="5">
|
||||
<heading id="E125819C6164FFC1FF7D405DFEFEFE81" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||||
<emphasis id="88A6EAE26164FFC1FF7D405DFEFEFE81" bold="true" box="[189,324,267,293]" pageId="4" pageNumber="5">Etymology</emphasis>
|
||||
|
|
@ -286,19 +274,21 @@ because it fits the 11 diagnostic traits for the genus proposed by
|
|||
<bibRefCitation id="DE434B016164FFC1FDF143DAFD5CFD02" author="Oftedal O. T." box="[561,742,651,678]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="4" pageNumber="5" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||||
]; femoral pores conspicuous in adult males; and limbs well developed, pentadactyl with all toes clawed.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="F2C8657B6164FFC2FF7D43A7FA82FAA8" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="diagnosis">
|
||||
<paragraph id="BA6D36F06164FFC1FF7D43A7FE51FCAF" blockId="4.[189,1399,753,1028]" box="[189,491,753,779]" pageId="4" pageNumber="5">
|
||||
<heading id="E125819C6164FFC1FF7D43A7FE51FCAF" bold="true" box="[189,491,753,779]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||||
<emphasis id="88A6EAE26164FFC1FF7D43A7FE51FCAF" bold="true" box="[189,491,753,779]" pageId="4" pageNumber="5">Definition and diagnosis</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BA6D36F06164FFC0FF7D4243FACDFE34" blockId="4.[189,1399,753,1028]" lastBlockId="5.[189,1399,267,400]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||||
<paragraph id="BA6D36F06164FFC2FF7D4243FBBFFEED" blockId="4.[189,1399,753,1028]" lastBlockId="7.[189,1398,267,330]" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
|
||||
A fairly robust species currently not assigned to any species group. The new species is characterized by the following combination of characters: (1) size small, body robust (maximum known SVL
|
||||
<quantity id="7D2A9B156164FFC1FF7D420AFE96FCD2" box="[189,300,860,886]" metricMagnitude="-2" metricUnit="m" metricValue="6.22" pageId="4" pageNumber="5" unit="mm" value="62.2">62.2 mm</quantity>
|
||||
); (2) tail longer than SVL; (3) dorsal scales small, quadrangular; (4) middorsal scales 53-57; (5) suboculars large, unequal in size, with sometimes one scale slightly protruding downward between 4
|
||||
<superScript id="4DA79BB86164FFC1FF0C42F7FF63FC0B" attach="left" box="[204,217,929,943]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
||||
and 5
|
||||
<superScript id="4DA79BB86164FFC1FEE342F7FE8AFC0B" attach="left" box="[291,304,929,943]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
||||
supralabial, suboculars separated from palpebrals by 2-3 rows of mostly inconspicuous tiny scales, a few of them distinctly enlarged with angular downward protrusion between suboculars; (6) nasal entire, without sub-nostril groove; (7) 3-4 large pigmented palpebrals; (8) body uniform beige or greyish to bluish brown in life, devoid of any conspicuous pattern in males; (9) absence of any dark line on side of head in adult males; (10) venter immaculate golden grey in life; (11) femoral pores 9-10 on each side in males, unknown in females; (12) preanal pores absent; (13) hemipenis globose, weakly bilobed, bordered by numerous flounces (>20, including about eight medioproximal asulcate flounces)
|
||||
supralabial, suboculars separated from palpebrals by 2-3 rows of mostly inconspicuous tiny scales, a few of them distinctly enlarged with angular downward protrusion between suboculars; (6) nasal entire, without sub-nostril groove; (7) 3-4 large pigmented palpebrals; (8) body uniform beige or greyish to bluish brown in life, devoid of any conspicuous pattern in males; (9) absence of any dark line on side of head in adult males; (10) venter immaculate golden grey in life; (11) femoral pores 9-10 on each side in males, unknown in females; (12) preanal pores absent; (13) hemipenis globose, weakly bilobed, bordered by numerous flounces (>20, including about eight medioproximal asulcate flounces) bearing comblike rows of minute weakly mineralized spinules, sulcus spermaticus divided in the crotch by a small fleshy bumplike structure, sulcus branches barely detectable.
|
||||
</paragraph>
|
||||
<caption id="EEAD66786164FFC1FF7D46FBFC2DF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711439" ID-Zenodo-Dep="3711439" httpUri="https://zenodo.org/record/3711439/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1965,1991]" targetBox="[189,1372,1072,1925]" targetPageId="4">
|
||||
<paragraph id="BA6D36F06164FFC1FF7D46FBFC2DF84F" blockId="4.[189,1398,1965,2028]" pageId="4" pageNumber="5">
|
||||
|
|
@ -320,7 +310,14 @@ Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Sca
|
|||
Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="EEAD66786166FFC3FF7D4676FACDF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711443" ID-Zenodo-Dep="3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1824,1850]" targetBox="[189,1399,194,1787]" targetPageId="6">
|
||||
<caption id="EEAD66786166FFC3FF7D42BEFD48FB82" pageId="6" pageNumber="7" startId="6.[189,232,1000,1026]" targetBox="[189,1399,1085,1787]" targetPageId="6">
|
||||
<paragraph id="BA6D36F06166FFC3FF7D42BEFD48FB82" pageId="6" pageNumber="7">
|
||||
<emphasis id="88A6EAE26166FFC3FF7D42BEFEB0FBA6" bold="true" box="[189,266,1000,1026]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
|
||||
<emphasis id="88A6EAE26166FFC3FED142BEFE4EFBA6" box="[273,500,1000,1026]" italics="true" pageId="6" pageNumber="7">Anadia mcdiarmidi</emphasis>
|
||||
sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="EEAD66786166FFC3FF7D4676FACDF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711443" ID-Zenodo-Dep="3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1824,1850]" targetBox="[189,1399,1085,1787]" targetPageId="6">
|
||||
<paragraph id="BA6D36F06166FFC3FF7D4676FACDF84F" blockId="6.[189,1399,1824,2027]" pageId="6" pageNumber="7">
|
||||
<emphasis id="88A6EAE26166FFC3FF7D4676FEB0F89F" bold="true" box="[189,266,1824,1851]" pageId="6" pageNumber="7">Fig. 5.</emphasis>
|
||||
<taxonomicName id="7DD24D736166FFC3FED24677FE4FF89F" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[274,501,1825,1851]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||||
|
|
@ -330,7 +327,6 @@ Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal sca
|
|||
The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="BA6D36F06167FFC2FF7D405DFBBFFEED" blockId="7.[189,1398,267,330]" pageId="7" pageNumber="8">bearing comblike rows of minute weakly mineralized spinules, sulcus spermaticus divided in the crotch by a small fleshy bumplike structure, sulcus branches barely detectable.</paragraph>
|
||||
<paragraph id="BA6D36F06167FFC2FF7D4027FA82FAA8" blockId="7.[189,1399,369,1292]" pageId="7" pageNumber="8">
|
||||
The new taxon is immediately distinguished from
|
||||
<taxonomicName id="7DD24D736167FFC2FD364027FC14FE2F" authorityName="Gray" authorityYear="1845" box="[758,942,369,395]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ocellata">
|
||||
|
|
@ -612,7 +608,9 @@ An adult male in good condition (
|
|||
<figureCitation id="22E92A756167FFC2FD2D4401FD44FAD5" box="[749,766,1367,1393]" captionStart="Fig" captionStartId="4.[189,232,1965,1991]" captionTargetBox="[189,1372,1072,1925]" captionTargetId="figure@4.[189,1398,1072,1925]" captionTargetPageId="4" captionText="Fig. 2. Anadia mcdiarmidi sp. nov. Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Scale lines = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711439" httpUri="https://zenodo.org/record/3711439/files/figure.png" pageId="7" pageNumber="8">2</figureCitation>
|
||||
,
|
||||
<figureCitation id="22E92A756167FFC2FCCC4401FCA6FAD5" box="[780,796,1367,1393]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,467,1890]" captionTargetId="figure@5.[189,1399,467,1890]" captionTargetPageId="5" captionText="Fig. 3. Anadia mcdiarmidi sp. nov. Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711441" httpUri="https://zenodo.org/record/3711441/files/figure.png" pageId="7" pageNumber="8">3</figureCitation>
|
||||
, 4, 6), with
|
||||
,
|
||||
<figureCitation id="22E92A756167FFC2FCEC4401FC87FAD5" box="[812,829,1367,1393]" captionStart="Fig" captionStartId="6.[189,232,1000,1026]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionText="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." pageId="7" pageNumber="8">4</figureCitation>
|
||||
, 6), with
|
||||
<quantity id="7D2A9B156167FFC2FC724401FBA7FAD5" box="[946,1053,1367,1393]" metricMagnitude="-2" metricUnit="m" metricValue="6.22" pageId="7" pageNumber="8" unit="mm" value="62.2">62.2 mm</quantity>
|
||||
SVL and 110.0 mm TL (see also
|
||||
<tableCitation id="F750034B6167FFC2FF32442DFEF2FA31" box="[242,328,1403,1429]" captionStart="Table 1" captionStartId="13.[189,258,822,848]" captionText="Table 1. Sex, measurements (in mm) and scale counts of the type series of Anadia mcdiarmidi sp. nov. 1 = tail incomplete or obviously regenerated, 2 = data corresponding to right/left side, other abbreviations are defined in the text." pageId="7" pageNumber="8">Table 1</tableCitation>
|
||||
|
|
@ -670,12 +668,16 @@ Eight marginal preanal scales anterior to vent, the middle two narrower than the
|
|||
</paragraph>
|
||||
<paragraph id="BA6D36F0616AFFCFFF7D42E8FBDEFC7C" blockId="10.[189,1124,958,985]" box="[189,1124,958,985]" pageId="10" pageNumber="11">Caudal scales smooth, subimbricate, rectangular, in transverse rows around tail.</paragraph>
|
||||
<paragraph id="BA6D36F0616AFFCFFF7D4556FBD0FB21" blockId="10.[189,1398,1024,1157]" pageId="10" pageNumber="11">Scales on dorsal surfaces of arm large, smooth, imbricate; ventral surfaces of arm with similar, but smaller and less imbricate scales. Hind limbs with large, smooth subimbricate to imbricate scales on anterior face of thighs and on lower legs; scales similar but smaller on ventral side of thigh. Dorsal and posterior sides of thigh with much smaller, irregularly shaped juxtaposed scales.</paragraph>
|
||||
<paragraph id="BA6D36F0616AFFCFFF7D45FBFBF6FADE" blockId="10.[189,1399,1197,1402]" pageId="10" pageNumber="11">Moderate-size, imbricate scales atop hands and feet. Supradigital scales single; upper and lower ungualsheath scales covering base of claws, leaving tips well exposed. Palms and soles with small slightly raised juxtaposed scales (Fig. 4). Two enlarged smooth thenar scales at base of pollex. Subdigital lamellae mostly single. Lamellae under first (I) through fifth (V) finger (right/left side): I: 7/6, II: 9/9, III: 11/11, IV: 13/14, V: 10/9. Lamellae under first (I) through fifth (V) toe (right/left side): I: 6/6, II: 10/10, III: 14/12, IV: 17/15, V: 15/6 (large part of Toe V missing on left side).</paragraph>
|
||||
<paragraph id="BA6D36F0616AFFCFFF7D45FBFBF6FADE" blockId="10.[189,1399,1197,1402]" pageId="10" pageNumber="11">
|
||||
Moderate-size, imbricate scales atop hands and feet. Supradigital scales single; upper and lower ungualsheath scales covering base of claws, leaving tips well exposed. Palms and soles with small slightly raised juxtaposed scales (
|
||||
<figureCitation id="22E92A75616AFFCFFE3645A2FDFEFAAB" box="[502,580,1268,1295]" captionStart="Fig" captionStartId="6.[189,232,1000,1026]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionText="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||||
). Two enlarged smooth thenar scales at base of pollex. Subdigital lamellae mostly single. Lamellae under first (I) through fifth (V) finger (right/left side): I: 7/6, II: 9/9, III: 11/11, IV: 13/14, V: 10/9. Lamellae under first (I) through fifth (V) toe (right/left side): I: 6/6, II: 10/10, III: 14/12, IV: 17/15, V: 15/6 (large part of Toe V missing on left side).
|
||||
</paragraph>
|
||||
<paragraph id="BA6D36F0616AFFCFFF7D44F4FC6CF9EE" blockId="10.[189,1399,1441,1610]" pageId="10" pageNumber="11">Tongue lanceolate; proximal two-third unpigmented, distal one-third heavily pigmented, including fork. Upper surface behind fork covered with imbricate scalelike papillae that extend onto ventrolateral sides of tongue. Raised medioventral side of tongue with numerous (>10) of thin, oblique, anteriorly converging and pointed (chevronlike) infralingual plicae; anterior pair of plicae swollen, bluntly pointed and larger than those following. Very shallow medioventral groove.</paragraph>
|
||||
<paragraph id="BA6D36F0616AFFCFFF7D4724FE37F90B" blockId="10.[189,1399,1650,1711]" pageId="10" pageNumber="11">Anterior maxillary and dentary teeth conical, unicuspid, with no or very feeble recurvature, becoming larger posteriorly.</paragraph>
|
||||
<paragraph id="BA6D36F0616AFFC9FF7D4781FB4DFEEF" blockId="10.[189,1399,1751,2027]" lastBlockId="12.[189,1399,267,331]" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
|
||||
The left hemipenis (
|
||||
<figureCitation id="22E92A75616AFFCFFE6B4781FE4BF956" box="[427,497,1751,1778]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,194,1787]" captionTargetPageId="6" captionText="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||||
<figureCitation id="22E92A75616AFFCFFE6B4781FE4BF956" box="[427,497,1751,1778]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionTargetPageId="6" captionText="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||||
) extends about four subcaudal rows when adpressed to the tail, and is about 7.0 mm in length and 4.0 mm across its widest point when fully everted (organ not maximally expanded because one of the lobes was slightly damaged during the eversion process). Hemipenial body globose, tapered near the base, weakly and symmetrically bilobed. Sulcus spermaticus broader near crotch, relatively shallow, curving about halfway around the base of the organ, then running medially towards the lobes. Sulcus flanked by a narrow nude area on each side, and divided in the crotch by a small fleshy bumplike structure, from where two narrow, ill-defined and barely detectable sulcus branches seem to extend laterally into the folded thickened 'ear-like' tissues of each lobe towards the apices of the lobes. Lobes are complexly folded, not flattened terminally, and sulcus branches seem to end in small depressions (from which residual seminal fluid was removed before hemipenis preparation).
|
||||
</paragraph>
|
||||
<caption id="EEAD6678616BFFCEFF7D46DCFEFFF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711451" ID-Zenodo-Dep="3711451" httpUri="https://zenodo.org/record/3711451/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1930,1956]" targetBox="[271,1316,265,1890]" targetPageId="11">
|
||||
|
|
@ -697,7 +699,7 @@ Myers
|
|||
2009
|
||||
</bibRefCitation>
|
||||
); fifteen remaining rows chevron-shaped, also circling the organ, but separated by a broad median nude space on the asulcate face and by the nude areas edging each side of the sulcus spermaticus. The plicae bear comblike rows of minute weakly mineralized spinules that project slightly from the edges of the plicae and usually increase in size towards the lateral faces and the basal part of the organ (staining of the spinules by Alizarin Red is well visible under the microscope although not observable in
|
||||
<figureCitation id="22E92A75616CFFC9FD4B4207FD68FCC8" box="[651,722,849,876]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,194,1787]" captionTargetPageId="6" captionText="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||||
<figureCitation id="22E92A75616CFFC9FD4B4207FD68FCC8" box="[651,722,849,876]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionTargetPageId="6" captionText="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="BA6D36F0616CFFC9FF7D42C2FD90FC0A" blockId="12.[189,1399,916,1127]" box="[189,554,916,943]" pageId="12" pageNumber="13">
|
||||
|
|
@ -712,7 +714,9 @@ in life
|
|||
<paragraph id="BA6D36F0616CFFC9FF7D42ECFA8DFBC3" blockId="12.[189,1399,916,1127]" pageId="12" pageNumber="13">
|
||||
Dorsum greyish to bluish brown depending on light intensity, devoid of any conspicuous pattern, but with a few scattered black scales on flanks. Arms and legs, including digits, greyish brown with a few scattered black scales; rear of thighs with a few ill-defined dark markings. Ventral surfaces of head and body golden grey, ventral surfaces of legs and tail bluish grey, palms and soles lemon yellow speckled with brown and yellowish orange scales. Tympanum dark brown, iris pale yellow (
|
||||
<figureCitation id="22E92A75616CFFC9FB47451AFAB1FBC3" box="[1159,1291,1100,1127]" captionStart="Fig" captionStartId="3.[189,232,1895,1921]" captionTargetBox="[207,1380,265,1855]" captionTargetId="figure@3.[207,1380,265,1855]" captionTargetPageId="3" captionText="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711437" httpUri="https://zenodo.org/record/3711437/files/figure.png" pageId="12" pageNumber="13">Figs 1A, C</figureCitation>
|
||||
, 4).
|
||||
,
|
||||
<figureCitation id="22E92A75616CFFC9FAD7451BFA93FBC3" box="[1303,1321,1101,1127]" captionStart="Fig" captionStartId="6.[189,232,1000,1026]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionText="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." pageId="12" pageNumber="13">4</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="BA6D36F0616CFFC9FF7D45D9FD27FB0D" blockId="12.[189,1399,1167,1378]" box="[189,669,1167,1194]" pageId="12" pageNumber="13">
|
||||
<heading id="E125819C616CFFC9FF7D45D9FD27FB0D" bold="true" box="[189,669,1167,1194]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||||
|
|
|
|||
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Reference in a new issue