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<taxonomicName id="B0EF6599FFA4FFBA31CDFB7EB609A18E" ID-CoL="5ZN7F" authority="(Steph.) E. W. Jones, Trans. Brit. Bryol. Soc." baseAuthorityName="Steph." box="[136,886,1199,1226]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Cololejeunea" kingdom="Plantae" order="Porellales" pageId="2" pageNumber="65" phylum="Marchantiophyta" rank="species" species="zenkeri">
<emphasis id="459BC208FFA4FFBA31CDFB7EB71AA18E" bold="true" box="[136,613,1199,1226]" pageId="2" pageNumber="65">
<emphasis id="459BC208FFA4FFBA31CDFB7EB403A18E" bold="true" box="[136,380,1199,1226]" italics="true" pageId="2" pageNumber="65">Cololejeunea zenkeri</emphasis>
(Steph.) E.W.Jones
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2: 420, 1954 (
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).
</heading>
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Basionym:—
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<emphasis id="459BC208FFA4FFBA304DFB2DB48DA055" box="[264,498,1276,1297]" italics="true" pageId="2" pageNumber="65">Aphanolejeunea zenkeri</emphasis>
Steph.,
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5: 858, 1916 (
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). Type:—
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. s.loc.,
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(G-00045059,
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<emphasis id="459BC208FFA4FFBA3376FAF1B7ACA071" bold="true" box="[563,723,1312,1333]" pageId="2" pageNumber="65">here designated</emphasis>
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=
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<emphasis id="459BC208FFA4FFBA31DEFA95B4EBA01D" box="[155,404,1348,1369]" italics="true" pageId="2" pageNumber="65">Cololejeunea duvignaudii</emphasis>
E.W.Jones,
<emphasis id="459BC208FFA4FFBA334FFA95B79EA01D" box="[522,737,1348,1369]" italics="true" pageId="2" pageNumber="65">Trans. Brit. Bryol. Soc</emphasis>
.
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2: 418, 1954 (
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),
<emphasis id="459BC208FFA4FFBA32B4FA92B13DA01C" bold="true" box="[1009,1090,1347,1368]" italics="true" pageId="2" pageNumber="65">syn. nov</emphasis>
. Type:—D. R.
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, Panzi (sousbois),
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(BR-5040032317160!,
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=
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<emphasis id="459BC208FFA4FFBA31DBFA5DB4E6A0E5" box="[158,409,1420,1441]" italics="true" pageId="2" pageNumber="65">Cololejeunea duvignaudii</emphasis>
var.
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Tixier,
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<emphasis id="459BC208FFA4FFBA32FFFA5DB698A0E5" box="[954,999,1420,1441]" italics="true" pageId="2" pageNumber="65">, n.s.</emphasis>
55: 223 1977 [1979] (
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),
<emphasis id="459BC208FFA4FFBA3417FA5AB0D4A0E5" box="[1362,1451,1419,1441]" italics="true" pageId="2" pageNumber="65">
<emphasis id="459BC208FFA4FFBA3417FA5AB0DAA0E4" bold="true" box="[1362,1445,1419,1440]" italics="true" pageId="2" pageNumber="65">syn. nov</emphasis>
.
</emphasis>
Type:—
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.
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: Manjakatompo,
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,
<emphasis id="459BC208FFA4FFBA3207FA61B6B2A081" box="[834,973,1456,1477]" italics="true" pageId="2" pageNumber="65">R. Benoist s.n.</emphasis>
(PC,
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<emphasis id="459BC208FFA4FFBA31CDFC5EB4ECA6EE" bold="true" box="[136,403,911,938]" italics="true" pageId="2" pageNumber="65">Cololejeunea vidaliana</emphasis>
Tixier,
<emphasis id="459BC208FFA4FFBA30AFFC40B7ECA6EE" box="[490,659,913,938]" italics="true" pageId="2" pageNumber="65">Nat. Hist. Bull.</emphasis>
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Soc.
</emphasis>
24 (3/4): 444, 1973 (
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Type:—
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. near Paksong (Bolovens), corticolous in clear forest,
<date id="035138DAFFA4FFBA325AFC0DB6A7A6B5" box="[799,984,988,1009]" pageId="2" pageNumber="65" value="1968-12-29">29 December 1968</date>
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<collectionCode id="11FE86DFFFA4FFBA32A7FC0DB68EA6B5" box="[994,1009,988,1009]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="65" type="Herbarium">P</collectionCode>
. Tixier 4139
</emphasis>
(
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,
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<paragraph id="77501E1AFFA4FFBA31CDFBD1B186A1C5" blockId="2.[136,1452,988,1153]" pageId="2" pageNumber="65">
=
<taxonomicName id="B0EF6599FFA4FFBA31DEFBD1B67BA151" ID-CoL="X7CY" authority="Tixier, Ann. Fac. Sci. Univ. Phnom Penh" box="[155,772,1024,1045]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Cololejeunea" kingdom="Plantae" order="Porellales" pageId="2" pageNumber="65" phylum="Marchantiophyta" rank="species" species="papulosa">
<emphasis id="459BC208FFA4FFBA31DEFBD1B406A151" box="[155,377,1024,1045]" italics="true" pageId="2" pageNumber="65">Cololejeunea papulosa</emphasis>
Tixier,
<emphasis id="459BC208FFA4FFBA3086FBD1B67BA151" box="[451,772,1024,1045]" italics="true" pageId="2" pageNumber="65">Ann. Fac. Sci. Univ. Phnom Penh</emphasis>
</taxonomicName>
1: 58, 1967 [1968] (
<bibRefCitation id="137E63EBFFA4FFBA328DFBD1B142A151" author="Tixier, P." box="[968,1085,1024,1045]" pageId="2" pageNumber="65" pagination="41 - 68" refId="ref5540" refString="Tixier, P. (1967) Bryophytae indosinicae. Inventaire bryologique du massif de Kiriom (Cambodge). Annales de la Faculte Sciences, Universite de Phnom Penh 1: 41 - 68." type="journal article" year="1967">Tixier 1967</bibRefCitation>
),
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<emphasis id="459BC208FFA4FFBA3090FBF5B7CAA17D" box="[469,693,1060,1081]" italics="true" pageId="2" pageNumber="65">Cololejeunea papulosa</emphasis>
Tixier,
<emphasis id="459BC208FFA4FFBA3247FBF5B6F0A17D" box="[770,911,1060,1081]" italics="true" pageId="2" pageNumber="65">Bryophyt. Bibl</emphasis>
.
</taxonomicName>
27: 41, 1985 (
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)
<emphasis id="459BC208FFA4FFBA35E2FBF5B076A17D" box="[1191,1289,1060,1081]" italics="true" pageId="2" pageNumber="65">nom. illeg</emphasis>
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<emphasis id="459BC208FFA4FFBA30E9FB96B77CA118" bold="true" box="[428,515,1095,1116]" italics="true" pageId="2" pageNumber="65">syn. nov.</emphasis>
Type:—
<collectingCountry id="0FF85E8AFFA4FFBA331DFB99B7B5A119" box="[600,714,1096,1117]" name="Vietnam" pageId="2" pageNumber="65">VIETNAM</collectingCountry>
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<collectingRegion id="B52BD0F8FFA4FFBA3391FB99B640A119" box="[724,831,1096,1117]" country="Vietnam" name="Lam Dong" pageId="2" pageNumber="65">Lam Dông</collectingRegion>
, forêt de la Dalao, épiphyllous,
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, 1960,
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(PC,
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). Blocking name:—
<taxonomicName id="B0EF6599FFA4FFBA3093FBBDB6D9A1C5" ID-CoL="X7CY" authority="R. M. Schust., Phytologia" authorityName="R. M. Schust., Phytologia" box="[470,934,1132,1153]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Cololejeunea" kingdom="Plantae" order="Porellales" pageId="2" pageNumber="65" phylum="Marchantiophyta" rank="species" species="papulosa">
<emphasis id="459BC208FFA4FFBA3093FBBDB7CBA1C5" box="[470,692,1132,1153]" italics="true" pageId="2" pageNumber="65">Cololejeunea papulosa</emphasis>
R.M.Schust.,
<emphasis id="459BC208FFA4FFBA3278FBBDB6D9A1C5" box="[829,934,1132,1153]" italics="true" pageId="2" pageNumber="65">Phytologia</emphasis>
</taxonomicName>
45 (5): 433, 1980 (
<bibRefCitation id="137E63EBFFA4FFBA3525FBBDB191A1C5" author="Schuster, R. M." box="[1120,1262,1132,1153]" pageId="2" pageNumber="65" pagination="415 - 437" refId="ref4944" refString="Schuster, R. M. (1980) New combinations and taxa of Hepaticae I. Phytologia 45: 415 - 437." type="journal article" year="1980">Schuster 1980</bibRefCitation>
).
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Basionym:—
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R.M.Schust.,
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Basionym:—
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C.Massal.,
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17 (3): 248, 1885 (
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(VER, G). ≡
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<emphasis id="459BC208FFA7FFB931CDF91FB728A3AD" bold="true" box="[136,599,1742,1769]" italics="true" pageId="1" pageNumber="64">Cololejeunea obtusifolia var. madecassa</emphasis>
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Basionym:—
<taxonomicName id="B0EF6599FFA7FFB9304DF8CBB1DAA26B" authority="Tixier, Bull. Trimestriel Acad. Malgache" box="[264,1189,1818,1839]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Cololejeunea" kingdom="Plantae" order="Porellales" pageId="1" pageNumber="64" phylum="Marchantiophyta" rank="variety" species="androphylla" variety="madecassa">
<emphasis id="459BC208FFA7FFB9304DF8CBB77BA26B" box="[264,516,1818,1839]" italics="true" pageId="1" pageNumber="64">Cololejeunea androphylla</emphasis>
E.W.Jones var.
<emphasis id="459BC208FFA7FFB933E0F8CBB671A26B" box="[677,782,1818,1839]" italics="true" pageId="1" pageNumber="64">madecassa</emphasis>
Tixier,
<emphasis id="459BC208FFA7FFB93219F8CBB1DAA26B" box="[860,1189,1818,1839]" italics="true" pageId="1" pageNumber="64">Bull. Trimestriel Acad. Malgache</emphasis>
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<emphasis id="459BC208FFA7FFB935E0F8CBB1ACA26B" box="[1189,1235,1818,1839]" italics="true" pageId="1" pageNumber="64">, n.s.</emphasis>
55: 216, 1977 [1979] (
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.
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, Mandialaza,
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<date id="035138DAFFA7FFB932E9F8EFB11BA217" box="[940,1124,1854,1875]" pageId="1" pageNumber="64" value="1975-12-19">19 December 1975</date>
,
<emphasis id="459BC208FFA7FFB9352BF8EFB18BA217" box="[1134,1268,1854,1875]" italics="true" pageId="1" pageNumber="64">P. Tixier 7670</emphasis>
(PC).
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<emphasis id="459BC208FFA7FFB931CDFA93B4DCA019" bold="true" box="[136,419,1346,1373]" italics="true" pageId="1" pageNumber="64">Cololejeunea obtusifolia</emphasis>
(E.W.Jones) Tixier,
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11: 42, 1995 (
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Basionym:—
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var.
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E.W.Jones,
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=
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<emphasis id="459BC208FFA7FFB931DFFA07B4E7A0AF" box="[154,408,1494,1515]" italics="true" pageId="1" pageNumber="64">Cololejeunea heterolobula</emphasis>
Tixier,
<emphasis id="459BC208FFA7FFB9309AFA07B734A0AF" box="[479,587,1494,1515]" italics="true" pageId="1" pageNumber="64">Trop. Bryol</emphasis>
.
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11: 40, 1995 (
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),
<emphasis id="459BC208FFA7FFB93227FA07B6C8A0AF" bold="true" box="[866,951,1494,1515]" italics="true" pageId="1" pageNumber="64">syn. nov.</emphasis>
Type:—
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, Cyangugu: Forêt de Ngyungwé, Kamiranzovu,
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,
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=
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<emphasis id="459BC208FFA7FFB931D9F9CFB4D8A377" box="[156,423,1566,1587]" italics="true" pageId="1" pageNumber="64">Cololejeunea pseudopusilla</emphasis>
Tixier,
<emphasis id="459BC208FFA7FFB930B6F9CFB71EA377" box="[499,609,1566,1587]" italics="true" pageId="1" pageNumber="64">Trop. Bryol</emphasis>
.
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11: 50, 1995 (
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) “
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”,
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Type:—D. R.
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: Mont Kahuzi, près de la route Bukawu-Irangi,
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(G-00113768!,
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=
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<emphasis id="459BC208FFA7FFB931D9F9B7B4D8A33F" box="[156,423,1638,1659]" italics="true" pageId="1" pageNumber="64">Cololejeunea sphaerocarpa</emphasis>
Tixier,
<emphasis id="459BC208FFA7FFB930B6F9B7B71EA33F" box="[499,609,1638,1659]" italics="true" pageId="1" pageNumber="64">Trop. Bryol</emphasis>
.
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11: 42, 1995 (
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),
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Type:—D. R.
<collectingCountry id="0FF85E8AFFA7FFB93529F9B7B1BAA33F" box="[1132,1221,1638,1659]" name="Democratic Republic of the Congo" pageId="1" pageNumber="64">CONGO</collectingCountry>
.
<collectingRegion id="B52BD0F8FFA7FFB9358BF9B7B07FA33F" box="[1230,1280,1638,1659]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="1" pageNumber="64">Kivu</collectingRegion>
, station forestière d’Irangi, au S.W. Du Mt. Ilimo,
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,
<date id="035138DAFFA7FFB93302F95BB79EA3DB" box="[583,737,1674,1695]" pageId="1" pageNumber="64" value="1991-08-23">23 August 1991</date>
,
<emphasis id="459BC208FFA7FFB933AEF95BB6E9A3DB" box="[747,918,1674,1695]" italics="true" pageId="1" pageNumber="64">E. Fischer 8511/4</emphasis>
(G-00113767,
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).
</paragraph>
</subSubSection>
</treatment>
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128
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<emphasis id="459BC208FFA7FFB931CDFEA7B4CDA4D5" bold="true" box="[136,434,374,401]" italics="true" pageId="1" pageNumber="64">Cololejeunea cardiocarpa</emphasis>
(Mont.) A.Evans,
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<emphasis id="459BC208FFA7FFB93208FEA9B6FCA4D5" box="[845,899,376,401]" italics="true" pageId="1" pageNumber="64">Club</emphasis>
8: 172, 1902 (
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Basionym:—
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<emphasis id="459BC208FFA7FFB9304DFE13B4A3A493" box="[264,476,450,471]" italics="true" pageId="1" pageNumber="64">Lejeunea cardiocarpa</emphasis>
Mont.,
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repentem in Cuba insula legerunt hanc speciem clarr.
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=
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<emphasis id="459BC208FFA7FFB931DBFDDBB4B6A75B" box="[158,457,522,543]" italics="true" pageId="1" pageNumber="64">Cololejeunea cardiocarpioides</emphasis>
Tixier,
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11: 38, 1995 (
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Type:—
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<mods:affiliation id="C22977E11B0A7B598FF0D11AD1F8E100">The Herbarium, Royal Botanic Gardens, Kew, Richmond TW 9 3 AB, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D7696BD430FDAA85EC24ECBBF7B2FDBA">text</mods:typeOfResource>
<mods:relatedItem id="D0E9D33E815E4C1A260A6BEC83760716" type="host">
<mods:titleInfo id="3343A529C89C31144911E27CB778F78C">
<mods:title id="E9F817A1F13E57DC9883E10E3EEBE743">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="A9F41C75067C51D398F4A68C86D41BA8">
<mods:date id="6E30654E9213EE32785E04159EDF008A">2024</mods:date>
<mods:detail id="A81CB29909E5687708C7968BBDE5C18E" type="pubDate">
<mods:number id="73E94A31323B33C25B93E63D8A0D8A08">2024-09-02</mods:number>
</mods:detail>
<mods:detail id="5EE773E661BF7FDCEDF5D27590CE6879" type="volume">
<mods:number id="147FC3256472BFB44103EBD1EF287249">246</mods:number>
</mods:detail>
<mods:extent id="9488C6785344CC9CAA6905B206E00B76" unit="page">
<mods:start id="6BFE6E9C1609BC9C24D5542E41A52556">71</mods:start>
<mods:end id="C75520606DF42588E975E1454F49DEA7">178</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1BF3ED9356D951D1A63A015600912E77">journal article</mods:classification>
<mods:identifier id="948BE2A745CEDF2036ADB405DC422ABF" type="CLB-Dataset">302074</mods:identifier>
<mods:identifier id="6261AFDF4C015A43DF918C17D05E0AA3" type="DOI">10.3897/phytokeys.246.129476</mods:identifier>
<mods:identifier id="93B436216CCBAEFE2F99CDF3EDC83BB2" type="GBIF-Dataset">b381880a-e0bb-4b02-ad0f-2f90a0b6c995</mods:identifier>
</mods:mods>
<treatment id="FFD2F625AD9A504AB0300285B338EB01" ID-DOI="http://doi.org/10.5281/zenodo.13629795" ID-GBIF-Taxon="240061330" ID-Zenodo-Dep="13629795" ID-arpha="FFD2F625-AD9A-504A-B030-0285B338EB01" LSID="urn:lsid:plazi:treatment:FFD2F625AD9A504AB0300285B338EB01" httpUri="http://treatment.plazi.org/id/FFD2F625AD9A504AB0300285B338EB01">
<subSubSection id="DEBBFAE05318DD65902DBF7196368185" type="nomenclature">
<paragraph id="A883B31C4D9C4B96CDFF53082CBA389C">
<treatmentCitation id="54175E78230BE569D73497BFDFFEE59B" author="A. J. Paton" httpUri="http://treatment.plazi.org/id/CB3171494ACD57189A62EA00435DA2FF" journal="Phytokeys" page="46" score="9.729594737742866" volume="129" year="2019">
<taxonomicName id="CD619F8EF7EEE243161F1492A3AFEF0D" ID-CoL="8XY5L" authority="(A. J. Paton) A. J. Paton" authorityName="A. J. Paton" baseAuthorityName="A. J. Paton" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="foliatus">
<emphasis id="67445697E839B7A68FD1F99F82CE15DE" italics="true">Coleus foliatus</emphasis>
(A. J. Paton) A. J. Paton
</taxonomicName>
, Phytokeys 129: 46. 2019
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AE9140389C95E4E69B16F8EB8B74F69" type="reference_group">
<paragraph id="59954B94029971FF57AF6C288C77B114">
<treatmentCitationGroup id="6E2CB3ACBEA1E846833461EC57EB8A61">
≡
<treatmentCitation id="C3AA7575F74F09457B3BC1D423FE268F" author="E. Afr." page="284" volumeTitle="Lamiac." year="2009">
<taxonomicName id="DA169653C4AC32CAA20191062A4E9866" ID-CoL="4K48Z" authority="A. J. Paton" authorityName="A. J. Paton" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="foliatus">
<emphasis id="0A2CE088392FF3BF1419B69D5B51BB2F" italics="true">Plectranthus foliatus</emphasis>
A. J. Paton
</taxonomicName>
, Fl. Trop. E. Afr., Lamiac.: 284. 2009
</treatmentCitation>
. Type:
<collectingCountry id="090C3E2DA466520DC5CE58D0E5C91ACB" name="Tanzania">Tanzania</collectingCountry>
, Sumbawanga District, Tatanda, Mbaa Hill,
<date id="67B5036403B5F088FD0433BB813976B4" value="1997-04-25">25 Apr 1997</date>
,
<emphasis id="6D731E44B7B0201197858661799B4B7E" italics="true">S. Bidgood et al. 3459</emphasis>
(
<typeStatus id="90F78B3C02012F08919F619EDFC1F7CD" type="holotype">holotype</typeStatus>
<collectionCode id="4313D0806E31E2E711B519E3D801D32E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
[
<specimenCode id="CB6A83BD3F1FFDF0F320BD980D41F399">
<collectionCode id="D162C12AC0C356BFDCE940560071D230" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
000194843
</specimenCode>
], [
<specimenCode id="9E9926630B9155AC4C10B87389FDD9DD">
<collectionCode id="66046AA53425C6B8DF43840D9FA51446" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
000194844
</specimenCode>
];
<typeStatus id="C307E0792136E06EF6712BA2EB424F6B" type="isotype">isotype</typeStatus>
<collectionCode id="0FFC74FCF74BCC117426CE30BBE52B0D" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
,
<collectionCode id="30938A690FAD1AF417368F399EF64808" country="Germany" httpUri="http://grbio.org/cool/4644-42cg" name="Deutsche Sammlung von Mikroorganismen und Zellkulturen GmbH" type="">DSM</collectionCode>
,
<collectionCode id="A809EC8732A495904696B5E69D149ACC" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
,
<collectionCode id="24EDB2C478EA8CEE36617D593F416692" country="Tanzania" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14235" lsid="urn:lsid:biocol.org:col:14235" name="Tropical Pesticides Research Institute" type="Herbarium">NHT</collectionCode>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3HGK" type="description">
<paragraph id="49A2C7708B6185AA01E6947CAC5EF744">
<heading id="BF20291B107F24693FBC765DBDA7AF11" reason="title">Description.</heading>
</paragraph>
<paragraph id="A3DF02FA5C5D48D2AAD8FA3EB63C5BFA">
<bibRefCitation id="5EDE7E40F875EC55261CE377D7484BF0" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Beentje HJ &amp; Ghazanfar SA &amp; Polhill RM" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="284" pagination="1-430" refId="B20" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2009) Lamiaceae (Labiatae). In: Beentje HJ, Ghazanfar SA, Polhill RM (Eds) Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond, 1–430." title="Lamiaceae (Labiatae)." volumeTitle="Flora of Tropical East Africa." year="2009">Paton et al. (2009: 284)</bibRefCitation>
,
<bibRefCitation id="91B4745C83F54794604096AC000D34D2" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Timberlake J" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="234" pagination="1-346" refId="B21" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2013) Lamiaceae. In: Timberlake J (Ed.) Flora Zambesiaca 8 (8). Royal Botanic Gardens, Kew, Richmond, 1–346." title="Lamiaceae." volumeTitle="Flora Zambesiaca 8 (8)." year="2013">Paton et al. (2013: 234)</bibRefCitation>
, as
<taxonomicName id="3A0586A782685F9E6BD4FE02E675D59E" authority="A. J. Paton." authorityName="A. J. Paton." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="foliatus">
<emphasis id="64701799455F0558D630615695D075D9" italics="true">Plectranthus foliatus</emphasis>
A. J. Paton.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVIGK" type="distribution">
<paragraph id="50747A4CD099B6663D22C64894F58A54">
<heading id="6CD55FC98152835ACFED0931C2D21585" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="B4F34949F0E0673B926C1283DB20F2E4">
<collectionCode id="766529A5ECDEDE8AD32B3AA734376B7F">SW</collectionCode>
<collectingCountry id="C6C88402AD4ECD91F656486578245F5A" name="Tanzania">Tanzania</collectingCountry>
, N
<collectingCountry id="40B706DF1C26B7DD76860FBC25CBF0DB" name="Zambia">Zambia</collectingCountry>
,
<collectingCountry id="7AE6E1DA26C6D4C498EC4483BAB4064F" name="Burundi">Burundi</collectingCountry>
, SE
<collectingCountry id="E8D7745EF267A5AB3FEEDD740FFC13A6">DR. Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBJGK" type="habitat">
<paragraph id="FDAF988AB9C8026E7571D0E32703F9E5">
<heading id="095B6D09560B721FBE11295A0B4D558A" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="90E86A70025645FBDAF734666E79D04A">
Steppic savannah, scrub, mostly on highlands, often on shallow rocky soil, occasionally on copper-rich soil;
<quantity id="E22A3B7A24EA3299B011E99290F6F84E" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">1500–2000 m</quantity>
elev.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGJGK" type="Additional specimens">
<paragraph id="5388962D02ACF952247E2F22BC2ACC57">
<heading id="8E6F3D3093BDA34294A7420EA27DF9BF" reason="title">Additional specimens.</heading>
</paragraph>
<paragraph id="4557C675181FC40FE23EE27CCB1EBA85">
<collectingCountry id="FEFE07EDA9F5CB2487B812F52FFB7777">DR. Congo</collectingCountry>
,
<emphasis id="07594AB81E21FB4120DBF3D37A189E3C" bold="true" italics="true">
<collectingRegion id="613D620BC92E8D70AC081FD74ED9E287">Haut-Katanga</collectingRegion>
</emphasis>
, Marungu, Kasiki,
<date id="840E9890106AF1FDB79836845EB936DC" value="1931-06-20">20 Jun 1931</date>
,
<emphasis id="8359D6665525DBEDFD677D788CF12E78" italics="true">
<collectionCode id="D053A05B0993E43A59D1C304235AF1AE" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. F. de Witte 470
</emphasis>
(
<collectionCode id="14611EC444E2CBAD71AFF6BF41D07EAB" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Tenke,
<emphasis id="F995CD12ED88579A79996DFECFB57C1E" italics="true">
P. Duvigneaud &amp; J. Timperman 2624
<collectionCode id="1C88D1405C43F5CAB1E1A0031F1E3846" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
</emphasis>
(
<collectionCode id="8BC2FFB1C7ED197244ECF9E472E17254" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
); Mitwaba-hôtel, steppe,
<date id="2EEA3A1E0B512A12513CE9671DAE00AA" value="1956-09-09">9 Sep 1956</date>
,
<emphasis id="CFD3736A1D6DFA983448902AF78220D8" italics="true">P. Duvigneaud &amp; J. Timperman 2691 Co</emphasis>
(
<collectionCode id="F4BED71ED82B6680EBE4450E09161982" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
); Marungu,
<quantity id="DBC7836058025875EFB5A38EE0CBDFE7" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
S Mont Lusale,
<date id="55D5A8E0CA8F589CB345AED8C42CDD9B" value="1957-06-27">27 Jun 1957</date>
,
<emphasis id="0AFA896FC169AE9F35A49043FB30F981" italics="true">
P. Duvigneaud 3738
<collectionCode id="54A4553E30CD51888B8546F85A9B47CD" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
</emphasis>
(
<collectionCode id="A78EC4CDF82FD1669F309F1BD657B0B6" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
); Kundelungu,
<quantity id="F11227F16E98976C3C69959387423999" metricMagnitude="3" metricUnit="m" metricValue="6.4" unit="km" value="6.4">6.4 km</quantity>
NNW Katshupa,
<date id="8455BA7399B8ABDB0D85AC0A68024C84" value="1966-10-17">17 Oct 1966</date>
,
<emphasis id="464F6880A783D51C23E898BDCF8130CA" italics="true">F. Malaisse 4660</emphasis>
(
<collectionCode id="0BEBABD0566109A48F57EFF26FDA4F41" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
,
<collectionCode id="1CCF3B6873A6553EA191B8368F2E7C97" country="Democratic Republic of the Congo" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14451" lsid="urn:lsid:biocol.org:col:14451" name="Université Nationale du Zaïre" type="Herbarium">LSHI</collectionCode>
); Poste de Mitwaba,
<date id="81AFC4C2C09F8210FE1BD33F9230AA31" value="1945-08">Aug 1945</date>
,
<emphasis id="99D10013E80786797CD4213F9511C029" italics="true">
<collectionCode id="5D3027E839234BDD23551E769911AEB3" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Mortelmans 96
</emphasis>
(
<collectionCode id="1EF9D82B82F904DAB583E390A703B679" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Domaine de Muhila, près de Kansimba,
<quantity id="669BE5E3F5AF59B5E96D2C6A8FCA6AB2" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</quantity>
elev., steppe,
<date id="DF43A36B5B21ECA892B13E3CC9AF775E" value="1970-11-07">7 Nov 1970</date>
,
<emphasis id="5229D3CF1BC49C8B4CCDBA7EC01797D7" italics="true">S. Lisowski 23488</emphasis>
(
<collectionCode id="E60E1DCCEDA389CDD94BF4C97A36DB3B" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15257" lsid="urn:lsid:biocol.org:col:15257" name="Adam Mickiewicz University" type="Herbarium">POZG</collectionCode>
).
</paragraph>
<paragraph id="A9ED319F056AB03E976DFF2E54D2FC5B">
<collectingCountry id="82D04F5C6244347BC64592568D0FB3C6" name="Burundi">Burundi</collectingCountry>
, Route Matana-Rutovu, km 15,
<date id="923B6C202B270950705BC2A341A25871" value="1966-02-26">26 Feb 1966</date>
,
<emphasis id="19572EAA9D1ECD83BC7B7CEBCA19453C" italics="true">J. Lewalle 469</emphasis>
(
<collectionCode id="502C12A06794C0002EA92932274039C8" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
);
<collectingRegion id="A0E6144129A0E1BB9F56E6B6DE24277F" country="Burundi" name="Bururi">Bururi</collectingRegion>
,
<date id="3D0D43F8799C391E307A7F6BD5524110" value="1968-02-03">3 Feb 1968</date>
,
<emphasis id="28430201EA88DFA3818234921C466ECD" italics="true">J. Lewalle 2760</emphasis>
(
<collectionCode id="838DC0C63F09EB42A9D22F3BA8B63A65" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
, P); Luvironza, Bututsi,
<date id="A2132255717D1A26F2445DDC12486E27" value="1955-03-03">3 Mar 1955</date>
,
<emphasis id="B3B82655F392C2B8ACF72BEF7D6D9245" italics="true">
<collectionCode id="588EFA15A822DAA7E3B031DE965708C9" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Michel 4702
</emphasis>
(
<collectionCode id="EF0EBBAB27F6B9815132CEE7A2613C71" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELKGK" type="notes">
<paragraph id="8EF2CD9282465548D2D418AEE32FBF45">
<heading id="47885CDF286F66EA6BE974D17F208F90" reason="title">Notes.</heading>
</paragraph>
<paragraph id="B7060A93677CE81A0842556DD399A9F4">
1. New species record for
<collectingCountry id="22B04531BA34044A05298CF47DDF1A9B">DR. Congo</collectingCountry>
. Widespread on the Mitwaba and the Marungu Plateau (
<collectingRegion id="9CCCAEA6E3B936CC281272B0689B37BE">Haut-Katanga</collectingRegion>
).
</paragraph>
<paragraph id="721EF5A72C1DD1B11006F0F9DAFD62E2">
2.
<taxonomicName id="D400BC4DB0F0012E53FA41DFA095A8F6" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="foliatus">
<emphasis id="43903226CE38293CA7DF190A4D8E7AA3" italics="true">Coleus foliatus</emphasis>
</taxonomicName>
is more variable in leaf shape in
<collectingCountry id="7821C51E0E9E82CB768308DAC7E82D73">DR. Congo</collectingCountry>
and
<collectingCountry id="385557B2EB9775814D653CC9EC178070" name="Burundi">Burundi</collectingCountry>
compared to neighbouring regions; specimens with unusually narrow leaves (ca. 45 ×
<quantity id="C35701BBF42AD873A0FF811DEE4A948C" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
) are found in
<collectingCountry id="465A822C0EDF594EB883F47E94A113A0" name="Burundi">Burundi</collectingCountry>
(e. g.
<emphasis id="620CD9FC807940E589CDB0C0FF210164" italics="true">J. Lewalle 469</emphasis>
,
<emphasis id="50CCB4FB9CC3EF86BF6982A2A2C3BAE3" italics="true">
<collectionCode id="96AF05334BB504931A2A8580C2037009" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Reekmans 8659
</emphasis>
); they differ from
<taxonomicName id="7EF7DCB1CF504ABF12E684E931B25273" authorityName="Meerts &amp; A. J. Paton A" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="duvigneaudii">
<emphasis id="9B5B086DA3BDABB473E59C5D99C3B1EE" italics="true">C. duvigneaudii</emphasis>
</taxonomicName>
in having opposite leaves, relatively short leaf blade and blunt leaf apex. In Upper Katanga, where
<taxonomicName id="94B2FC316AFAC89E0D6C516A3477AE49" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="foliatus">
<emphasis id="3D2506C538AB95F1FFE0A3742C0FBCC0" italics="true">C. foliatus</emphasis>
</taxonomicName>
and
<taxonomicName id="B5DEC6E8179A9F74293908A640EBE4AD" authorityName="Meerts &amp; A. J. Paton A" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="duvigneaudii">
<emphasis id="9DC36FB81684C4DC08A1946B5A21BCC1" italics="true">C. duvigneaudii</emphasis>
</taxonomicName>
are sympatric, more or less intermediate specimens occur (e. g.
<emphasis id="5C7962C9681E1A3B88249D9B1597E102" italics="true">S. Lisowski, F. Malaisse &amp; J. - J. Symoens 1247</emphasis>
,
<emphasis id="47B44B3E0760493AB6F2E4D418A7253D" italics="true">S. Lisowski, F. Malaisse &amp; J. - J. Symoens 11295; A. Schmitz 3150</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>