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<document id="E822CBF5A1657872CC7CFAF9110AD860" ID-DOI="10.11646/phytotaxa.217.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13634131" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725325955495" checkinUser="felipe" docAuthor="Jovanovska, Elena, Levkov, Zlatko &amp; Edlund, Mark B." docDate="2015" docId="4B0E6E2AFF98FF9DFF2AF8A1FDCEFF6F" docLanguage="en" docName="phytotaxa.217.3.1.pdf" docOrigin="Phytotaxa 217 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.217.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Diploneis peterseni Hustedt 1937" docType="treatment" docVersion="3" lastPageNumber="240" masterDocId="B7371652FFBDFFBAFFA2FFCFFFE4FFB2" masterDocTitle="The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia" masterLastPageNumber="248" masterPageNumber="201" pageNumber="238" updateTime="1725326923387" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="35E6CA5D0041D606143909E9DAA57B53">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<mods:namePart id="5B5F1BC099FD0430AFA61494246081EA">Jovanovska, Elena</mods:namePart>
<mods:affiliation id="5DF3B76D7F43EFD4B1A7EF0F46197860">Department of Animal Ecology and Systematics, Justus-Liebig University, Heinrich-Buff-Ring 26 - 32, 35392 Giessen, Germany</mods:affiliation>
<mods:nameIdentifier id="662AE4815DBA1E3531850A3D79BBD300" type="email">jovanovska.eci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C4F4FE87BA64A66B2942C4401499A1E7">Levkov, Zlatko</mods:namePart>
<mods:affiliation id="AA8543D393C89423940A3702F09A02FC">Institute of Biology, Faculty of Natural Sciences, Arhimedova 3, 1000 Skopje, R. of Macedonia</mods:affiliation>
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<mods:namePart id="1F74306259DFD1037A82BAD8206908C1">Edlund, Mark B.</mods:namePart>
<mods:affiliation id="A99EBDD96BBAFE84CE6CD3FB804192D7">St. Croix Watershed Research Station, Science Museum of Minnesota, Marine on St. Croix, Minnesota 55047, USA</mods:affiliation>
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<paragraph id="C318DF3CFF98FF9FFF2AF8A1FD93F83B" blockId="37.[136,631,1902,1929]" box="[136,631,1902,1929]" pageId="37" pageNumber="238">
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<taxonomicName id="04A7A4BFFF98FF9FFF2AF8A1FE20F83B" authority="Hustedt" authorityName="Hustedt" authorityYear="1937" box="[136,452,1902,1929]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="37" pageNumber="238" phylum="Ochrophyta" rank="species" species="peterseni">
<emphasis id="F1D3032EFF98FF9FFF2AF8A1FE82F83A" bold="true" box="[136,358,1902,1929]" italics="true" pageId="37" pageNumber="238">Diploneis peterseni</emphasis>
Hustedt
</taxonomicName>
(
<figureCitation id="5B9CC3B9FF98FF9FFE71F8BFFD94F83B" box="[467,624,1904,1929]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 246252</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFF98FF9CFF2AF87EFE33FE85" blockId="37.[136,1452,1969,2088]" lastBlockId="38.[136,1452,160,311]" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238">
The valves are lanceolate-elliptical with convex margins and bluntly round ends (
<figureCitation id="5B9CC3B9FF98FF9FFC58F87EFB2DF87A" box="[1018,1225,1969,1992]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 246249, 252</figureCitation>
). The valve length is 13.521.0 μm, and the breadth is 6.07.0 μm. The axial area is linear throughout the whole length. The central area is small and rectangular, ca. 0.51.0 μm wide. The longitudinal canal is lanceolate, slightly expanding at the valve apices (
<figureCitation id="5B9CC3B9FF98FF9FFAE5F83EFF50F79A" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 251, 252</figureCitation>
). The external openings of the canal are covered with volate occlusions (
<figureCitation id="5B9CC3B9FF98FF9FFC18F7DEFBFCF79A" box="[954,1048,2065,2088]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Fig. 252</figureCitation>
). Externally, the raphe is straight with simple and slightly expanded proximal and distal ends (
<figureCitation id="5B9CC3B9FF9BFF9CFD74FF6FFC8AFF05" box="[726,878,160,183]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="38" pageNumber="239">Figs 250252</figureCitation>
). Distally, the raphe branches terminate some distance from the valve margins (
<figureCitation id="5B9CC3B9FF9BFF9CFE3AFF0FFE1CFF65" box="[408,504,192,215]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="38" pageNumber="239">Fig. 251</figureCitation>
). The striae are parallel in the middle of the valve, becoming radiate toward the valve apices,
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10 μm. The single areola in each stria is covered with a volate occlusion that occupies less then half of the valve width (
<figureCitation id="5B9CC3B9FF9BFF9CFEB8FECFFE98FEA5" box="[282,380,256,279]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="38" pageNumber="239">Fig. 252</figureCitation>
). The hyaline area that covers part of the alveolate striae is lanceolate and broadly widens at the middle of the valve (
<figureCitation id="5B9CC3B9FF9BFF9CFEC9FEEFFE2FFE85" box="[363,459,288,311]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="38" pageNumber="239">Fig. 252</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="C318DF3CFF9BFF9CFF2AF8B6FAF6F7AC" blockId="38.[136,1452,1913,2078]" pageId="38" pageNumber="239">
<emphasis id="F1D3032EFF9BFF9CFF2AF8B6FEB5F83C" bold="true" box="[136,337,1913,1934]" pageId="38" pageNumber="239">FIGURES 236252.</emphasis>
LM and SEM external valve views. Figs 236245.
<taxonomicName id="04A7A4BFFF9BFF9CFC9AF8B6FC07F83C" box="[824,995,1913,1934]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="38" pageNumber="239" phylum="Ochrophyta" rank="species" species="oculata">
<emphasis id="F1D3032EFF9BFF9CFC9AF8B6FC07F83C" box="[824,995,1913,1934]" italics="true" pageId="38" pageNumber="239">Diploneis oculata</emphasis>
</taxonomicName>
. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252.
<taxonomicName id="04A7A4BFFF9BFF9CFD5CF80EFBADF864" authority=", LM and SEM" authorityName="LM and SEM" box="[766,1097,1985,2006]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="38" pageNumber="239" phylum="Ochrophyta" rank="species" species="peterseni">
<emphasis id="F1D3032EFF9BFF9CFD5CF80EFC5DF864" box="[766,953,1985,2006]" italics="true" pageId="38" pageNumber="239">Diploneis peterseni</emphasis>
, LM and SEM
</taxonomicName>
external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251).
</paragraph>
</caption>
<subSubSection id="8BBD8CB7FF9AFF9DFF2AFF6FFDCEFF6F" pageId="39" pageNumber="240" type="biology_ecology">
<paragraph id="C318DF3CFF9AFF9DFF2AFF6FFDCEFF6F" blockId="39.[136,1451,160,221]" pageId="39" pageNumber="240">
<emphasis id="F1D3032EFF9AFF9DFF2AFF6FFE24FF0B" bold="true" box="[136,448,160,185]" pageId="39" pageNumber="240">Ecology and Distribution:</emphasis>
—M166A;
<materialsCitation id="73CFD561FF9AFF9DFDE1FF6FFDC3FF6F" location="southern Lake" pageId="39" pageNumber="240" specimenCount="1" stateProvince="Hovsgol">
M068A: found on rocks in shallow habitats in
<location id="C67889E7FF9AFF9DFBF4FF6FFB12FF0B" LSID="urn:lsid:plazi:treatment:4B0E6E2AFF98FF9DFF2AF8A1FDCEFF6F:C67889E7FF9AFF9DFBF4FF6FFB12FF0B" box="[1110,1270,160,185]" name="southern Lake" pageId="39" pageNumber="240" stateProvince="Hovsgol">southern Lake</location>
<collectingRegion id="016311DEFF9AFF9DFB5CFF6FFAB9FF0B" box="[1278,1373,160,185]" country="Mongolia" name="Hovsgol" pageId="39" pageNumber="240">Hövsgöl</collectingRegion>
and in small streams in
<collectingRegion id="016311DEFF9AFF9DFEE6FF0BFDC3FF6F" box="[324,551,196,221]" country="Mongolia" name="Arhangay" pageId="39" pageNumber="240">Arkhangai province</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="DCC5AB654930057CE14AC3B879387B28">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<taxonomicName id="04A7A4BFFF98FF9FFF2AFAB5FDC1FA27" authority="(Brebisson) Cleve" authorityName="Cleve" baseAuthorityName="Brebisson" box="[136,549,1402,1429]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="37" pageNumber="238" phylum="Ochrophyta" rank="species" species="oculata">
<emphasis id="F1D3032EFF98FF9FFF2AFAB5FEB0FA27" bold="true" box="[136,340,1402,1429]" italics="true" pageId="37" pageNumber="238">Diploneis oculata</emphasis>
(Brébisson) Cleve
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(
<figureCitation id="5B9CC3B9FF98FF9FFD96FAB3FD34FA27" box="[564,720,1404,1429]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 236245</figureCitation>
)
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Valves are elongate-elliptical with parallel to slightly convex margins and bluntly round ends (
<figureCitation id="5B9CC3B9FF98FF9FFB30FA72FAC9FA66" box="[1170,1325,1469,1492]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 236244</figureCitation>
). The valve length is 13.024.5 μm, and the breadth is 5.07.5 μm. The axial area is linear, expanding into a small rectangular central area, ca. 0.51.0 μm wide. The longitudinal canal is linear, with a rectangular central nodule (
<figureCitation id="5B9CC3B9FF98FF9FFBDFFA32FAF2F9A6" box="[1149,1302,1533,1556]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 243245</figureCitation>
). At the valve apices the axial area expands slightly on the both sides of the valve (
<figureCitation id="5B9CC3B9FF98FF9FFCFBF9D2FC0BF986" box="[857,1007,1565,1588]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 243, 244</figureCitation>
). Externally, the canal is composed of one row of areolae throughout the whole length, covered with volate occlusions (
<figureCitation id="5B9CC3B9FF98FF9FFC0FF9F2FBA0F9E6" box="[941,1092,1597,1620]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 243245</figureCitation>
). The raphe is straight and slightly expanding into small drop-like central pores (
<figureCitation id="5B9CC3B9FF98FF9FFDC9F992FCE6F9C6" box="[619,770,1629,1652]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 244, 245</figureCitation>
). The distal raphe endings are simple and slightly expanded, terminating some distance from the valve margins (
<figureCitation id="5B9CC3B9FF98FF9FFD2DF9B2FCC2F926" box="[655,806,1661,1684]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 243, 244</figureCitation>
). The striae are parallel in the middle of the valve, becoming slightly radiate at the valve apices,
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10 μm. The areolae of the alveolate striae are not discernible with LM and SEM observations, due to the volate occlusions that occupy less then half of the valve width (
<figureCitation id="5B9CC3B9FF98FF9FFBEAF972FB00F966" box="[1096,1252,1725,1748]" captionStart="FIGURES 236252" captionStartId="38.[136,243,1913,1934]" captionTargetBox="[257,1328,339,1885]" captionTargetId="figure-107@38.[254,1334,336,1888]" captionTargetPageId="38" captionText="FIGURES 236252. LM and SEM external valve views. Figs 236245.Diploneis oculata. Figs 236242. LM valve views. Fig. 243. View of the whole valve overlaid by the girdle bands. Fig. 244. View of the entire valve. Fig. 245. Proximal raphe endings and detailed view of the external alveolate openings covered with vola. Fig. 246252. Diploneis peterseni, LM and SEM external valve views. Figs 246249. LM valve views. Fig. 250. Valve central area with simple proximal raphe endings. Fig. 251. Distal raphe endings. Fig. 252. View of the entire valve. Scale bars = 10 μm (Figs 236242, 246249); 5 μm (Fig. 252); 2 μm (Figs 243245, 250); 1 μm (Fig. 251)." figureDoi="http://doi.org/10.5281/zenodo.13634180" httpUri="https://zenodo.org/record/13634180/files/figure.png" pageId="37" pageNumber="238">Figs 244, 245</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF98FF9FFF2AF8CBFD25F8F3" pageId="37" pageNumber="238" type="biology_ecology">
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<emphasis id="F1D3032EFF98FF9FFF2AF8CBFE5FF8AF" bold="true" box="[136,443,1796,1821]" pageId="37" pageNumber="238">Ecology and Distribution:</emphasis>
—M068A; M077A; M248A; M273A;
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M287A: found in shallow to deep habitats including epilithon and sediments throughout
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<collectingRegion id="016311DEFF98FF9FFDF8F8E7FD59F8F3" box="[602,701,1832,1857]" country="Mongolia" name="Hovsgol" pageId="37" pageNumber="238">Hövsgöl</collectingRegion>
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.
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<mods:title id="3785CB60D038AA47E16BD3039AB10BE6">Phytotaxa</mods:title>
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<mods:part id="5437AC70F37FA88095D9D5C1B13B06F4">
<mods:date id="9F6B76A0BC82687B1E13E185FBA13A47">2015</mods:date>
<mods:detail id="5F26F05471C8546852958FFCE7E5F7B7" type="pubDate">
<mods:number id="AAFBC17FD8B8E15041B1F83B6F3AF73E">2015-06-26</mods:number>
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<mods:detail id="95F71E1B05C19AE8B2918C17DA0219B7" type="volume">
<mods:number id="1351E9243647BE55864670EB870FDC81">217</mods:number>
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<mods:number id="EA7E610184FC30C2EA6F4BF453F2C4E7">3</mods:number>
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<mods:location id="59AB6BDA7B5049BA4B6C8D47C1B1C5AB">
<mods:url id="AC4AE303307E98AA3DC8098EB03656AF">http://dx.doi.org/10.11646/phytotaxa.217.3.1</mods:url>
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<mods:classification id="805E17426A296F8CF21D1271EF5D36BC">journal article</mods:classification>
<mods:identifier id="3CFBF6BBB291784036D761106B9C34E4" type="DOI">10.11646/phytotaxa.217.3.1</mods:identifier>
<mods:identifier id="79AA7DB34DA4B0FA681633AB94353785" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="9748B6E6063344AE660E6AD8248E9C04" type="Zenodo-Dep">13634131</mods:identifier>
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<treatment id="4B0E6E2AFF98FF9FFF2AFF51FE8BFAFF" ID-DOI="http://doi.org/10.5281/zenodo.13634257" ID-Zenodo-Dep="13634257" LSID="urn:lsid:plazi:treatment:4B0E6E2AFF98FF9FFF2AFF51FE8BFAFF" httpUri="http://treatment.plazi.org/id/4B0E6E2AFF98FF9FFF2AFF51FE8BFAFF" lastPageNumber="238" pageId="37" pageNumber="238">
<subSubSection id="8BBD8CB7FF98FF9FFF2AFF51FE10FCCA" pageId="37" pageNumber="238" type="nomenclature">
<paragraph id="C318DF3CFF98FF9FFF2AFF51FD82FF0B" blockId="37.[136,614,158,185]" box="[136,614,158,185]" pageId="37" pageNumber="238">
<heading id="98506850FF98FF9FFF2AFF51FD82FF0B" box="[136,614,158,185]" fontSize="11" level="2" pageId="37" pageNumber="238" reason="3">
<taxonomicName id="04A7A4BFFF98FF9FFF2AFF51FE56FF0B" authority="Cleve" authorityName="Cleve" authorityYear="1891" box="[136,434,158,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="37" pageNumber="238" phylum="Ochrophyta" rank="species" species="boldtiana">
<emphasis id="F1D3032EFF98FF9FFF2AFF51FE8FFF0B" bold="true" box="[136,363,158,185]" italics="true" pageId="37" pageNumber="238">Diploneis boldtiana</emphasis>
Cleve
</taxonomicName>
(
<figureCitation id="5B9CC3B9FF98FF9FFE60FF6FFDBAFF0B" box="[450,606,160,185]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 220235</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFF98FF9FFF2AFF2EFE10FCCA" blockId="37.[136,1452,225,888]" pageId="37" pageNumber="238">
Valves are lanceolate-elliptic with slightly convex margins and bluntly rounded ends (
<figureCitation id="5B9CC3B9FF98FF9FFBE0FF2EFB39FF4A" box="[1090,1245,225,248]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 220231</figureCitation>
). The valve length is 16.526.0 μm, and the valve breadth is 8.09.0 μm. The axial area is very narrow, slightly expanding into a small to indistinguishable central area, 1.52.0 μm wide (
<figureCitation id="5B9CC3B9FF98FF9FFD3AFEEEFCD5FE8A" box="[664,817,289,312]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 222225</figureCitation>
). From inside, a thick and lanceolate silica plate closes the longitudinal canal throughout its whole length (
<figureCitation id="5B9CC3B9FF98FF9FFD2FFE8EFD09FEEA" box="[653,749,321,344]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Fig. 232</figureCitation>
). Externally, the longitudinal canal appears narrow, lanceolate to linear, slightly expanded in the middle of the valve with one to two rows of areolae becoming three towards the valve apices (
<figureCitation id="5B9CC3B9FF98FF9FFF32FE4EFECCFE2A" box="[144,296,385,408]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 231, 234</figureCitation>
). In LM it seems like the longitudinal canal is composed of one row of areolae throughout the whole length of the canal (
<figureCitation id="5B9CC3B9FF98FF9FFEB6FE6EFE49FE0A" box="[276,429,417,440]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 220230</figureCitation>
). From the exterior, the areolae of the canal are covered with simple cribra formed by irregular, small and round (310) pores, arranged in groups (
<figureCitation id="5B9CC3B9FF98FF9FFD03FE0EFCDCFE6A" box="[673,824,449,472]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 234, 235</figureCitation>
). The structure of areolar cribra is morphological similar to those from the striae, from which they are separated with a narrow hyaline area (
<figureCitation id="5B9CC3B9FF98FF9FFC40FE2EFB54FE4A" box="[994,1200,481,504]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 231, 233235</figureCitation>
). Internally the longitudinal canal is closed with a thick silica plate, forming a “depression” where the raphe is inserted (
<figureCitation id="5B9CC3B9FF98FF9FFB3EFDCEFB1EFDAA" box="[1180,1274,513,536]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Fig. 232</figureCitation>
). From the outer wall the raphe is straight with expanded proximal ends, bent to the same side of the valve (
<figureCitation id="5B9CC3B9FF98FF9FFBF7FDEEFB09FD8A" box="[1109,1261,545,568]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 231, 234</figureCitation>
). The distal raphe branches are bent into a short drop-like terminal fissure (
<figureCitation id="5B9CC3B9FF98FF9FFD5BFD8EFCBDFDEA" box="[761,857,577,600]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Fig. 235</figureCitation>
). Internally, the raphe is straight with simple proximal and distal ends, inserted in a slightly elevated sternum inside the “trench” formed by the longitudinal canal (
<figureCitation id="5B9CC3B9FF98FF9FFAB0FDAEFA94FDCA" box="[1298,1392,609,632]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Fig. 232</figureCitation>
). The proximal raphe endings reach the height of longitudinal canal and the valve itself (
<figureCitation id="5B9CC3B9FF98FF9FFC52FD4EFBA9FD2A" box="[1008,1101,641,664]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Fig. 232</figureCitation>
). Striae are parallel in the middle becoming radiate at the valve apices,
<quantity id="045F72D9FF98FF9FFDB6FD6EFD96FD0A" box="[532,626,673,696]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="37" pageNumber="238" unit="in" value="16.5" valueMax="18.0" valueMin="15.0">1518 in</quantity>
10 μm. Each alveolate stria is composed of two rows of areolae, covered with simple cribrate occlusions (
<figureCitation id="5B9CC3B9FF98FF9FFE0DFD0EFDACFD6A" box="[431,584,705,728]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 231, 235</figureCitation>
). The irregular and round cribrate pores (45), are increasing in number towards the valve margins (79). The perforations of the cribra are more regularly scattered then those of the longitudinal canal, where the perforations are more densely arranged (
<figureCitation id="5B9CC3B9FF98FF9FFD03FCCEFCD3FCAA" box="[673,823,769,792]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Figs 233, 235</figureCitation>
). At the valve apices the last two-three alveolate striae have a triangular shape (arrow on
<figureCitation id="5B9CC3B9FF98FF9FFE62FCEEFDF9FC8A" box="[448,541,801,824]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Fig. 235</figureCitation>
). Internally each alveolus opens through a single continuous opening covered with a fine silica layer (
<figureCitation id="5B9CC3B9FF98FF9FFE98FC8EFE73FCEA" box="[314,407,833,856]" captionStart="FIGURES 220235" captionStartId="36.[136,243,1959,1980]" captionTargetBox="[181,1407,356,1929]" captionTargetId="figure-86@36.[177,1410,353,1933]" captionTargetPageId="36" captionText="FIGURES 220235. Diploneis boldtiana, LM and SEM valve views. Figs 220230. LM valve views. Fig. 231. External valve view. Fig. 232. Internal view of the entire valve. Fig. 233.Detailed view of the external irregular and round areolar pores/openings.Fig. 234. External central area with proximal raphe endings, which are bent to the same side of the valve. Fig. 235. View of the distal raphe endings with a short terminal fissure. Scale bars = 10 μm (Figs 220232); 2 μm (Figs 233235)." figureDoi="http://doi.org/10.5281/zenodo.13634178" httpUri="https://zenodo.org/record/13634178/files/figure.png" pageId="37" pageNumber="238">Fig. 232</figureCitation>
). The single continuous opening has a clavate shape in the middle of the valve, becoming elongate in shape towards the valve apices.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF98FF9FFF2AFC67FB8EFB53" pageId="37" pageNumber="238" type="description">
<paragraph id="C318DF3CFF98FF9FFF2AFC67FB8EFB53" blockId="37.[136,1452,936,1357]" pageId="37" pageNumber="238">
<emphasis id="F1D3032EFF98FF9FFF2AFC67FED4FC73" bold="true" box="[136,304,936,961]" pageId="37" pageNumber="238">Observations:</emphasis>
<bibRefCitation id="A736A2CDFF98FF9FFEEFFC67FDE3FC73" author="Cleve, P. T." box="[333,519,936,961]" pageId="37" pageNumber="238" pagination="1 - 68" refId="ref25442" refString="Cleve, P. T. (1891) The Diatoms of Finland. Actas Societas Pro Fauna et Flora Fennica 8 (2): 1 - 68. http: // dx. doi. org / 10.5962 / bhl. title. 64355" type="journal article" year="1891">Cleve (1891: 43</bibRefCitation>
, pl. 2, fig. 12) in the original description gives only a single valve measurement for
<taxonomicName id="04A7A4BFFF98FF9FFF12FC03FED9FC57" authorityName="Cleve" authorityYear="1891" box="[176,317,972,997]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="37" pageNumber="238" phylum="Ochrophyta" rank="species" species="boldtiana">
<emphasis id="F1D3032EFF98FF9FFF12FC03FF20FC57" box="[176,196,972,997]" italics="true" pageId="37" pageNumber="238">D</emphasis>
.
<emphasis id="F1D3032EFF98FF9FFF70FC03FED9FC57" box="[210,317,972,997]" italics="true" pageId="37" pageNumber="238">boldtiana</emphasis>
</taxonomicName>
(length: 30.0 μm, breadth: 12.0 μm, and stria density:
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10 μm). Later,
<bibRefCitation id="A736A2CDFF98FF9FFB2EFC03FAD0FC57" author="Hustedt, F." box="[1164,1332,972,997]" pageId="37" pageNumber="238" pagination="577 - 736" refId="ref26709" refString="Hustedt, F. (1937) Die Kieselalgen Deutschlands, Osterreichs und der Schweiz unter Berucksichtigung der ubrigen Lander Europas sowie der angrenzenden Meeresgebiete. In: Rabenhorst, L. (Ed.) Kryptogamen Flora von Deutschland, Osterreich und der Schweiz. Akademische Verlagsgesellschaft m. b. h. Leipzig 7 (Teil 2, Lief. 5), pp. 577 - 736." type="journal article" year="1937">Hustedt (1937)</bibRefCitation>
illustrated a broader size range (length: 23.038.0 μm, breadth: 10.012.0 μm, and stria density: 1415 with 2730 areolae in 10 μm). According to the typification provided by Idei &amp; Kobayashi (1989b),
<taxonomicName id="04A7A4BFFF98FF9FFBACFBDBFB79FB9F" authorityName="Cleve" authorityYear="1891" box="[1038,1181,1044,1069]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="37" pageNumber="238" phylum="Ochrophyta" rank="species" species="boldtiana">
<emphasis id="F1D3032EFF98FF9FFBACFBDBFB79FB9F" box="[1038,1181,1044,1069]" italics="true" pageId="37" pageNumber="238">D. boldtiana</emphasis>
</taxonomicName>
unambiguously fits the measurements given by Cleve (length: 30.0 μm, breadth: 11.0 μm, and stria density:
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10 μm). In addition, Idei &amp; Kobayashi (1989b) observed a population of
<taxonomicName id="04A7A4BFFF98FF9FFD08FB93FCDCFBC7" authorityName="Cleve" authorityYear="1891" box="[682,824,1116,1141]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="37" pageNumber="238" phylum="Ochrophyta" rank="species" species="boldtiana">
<emphasis id="F1D3032EFF98FF9FFD08FB93FCDCFBC7" box="[682,824,1116,1141]" italics="true" pageId="37" pageNumber="238">D. boldtiana</emphasis>
</taxonomicName>
in the Finnish lakes, though with greater valve lengths (20.030.0 μm), but similar valve widths (11.012.0 μm) and stria densities (13/
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10 μm). The smaller size range observed in Lake
<collectingRegion id="016311DEFF98FF9FFEF8FB6BFE5DFB0F" box="[346,441,1188,1213]" country="Mongolia" name="Hovsgol" pageId="37" pageNumber="238">Hövsgöl</collectingRegion>
could be result of insufficient number of observed valves. The populations from Lake
<collectingRegion id="016311DEFF98FF9FFF2AFB07FF03FB53" box="[136,231,1224,1249]" country="Mongolia" name="Hovsgol" pageId="37" pageNumber="238">Hövsgöl</collectingRegion>
nicely fit the LM and SEM illustrations provided by Idei &amp; Kobayashi (1989b).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF98FF9FFF1FFB23FE8BFAFF" pageId="37" pageNumber="238" type="biology_ecology">
<paragraph id="C318DF3CFF98FF9FFF1FFB23FE8BFAFF" blockId="37.[136,1452,936,1357]" pageId="37" pageNumber="238">
<emphasis id="F1D3032EFF98FF9FFF1FFB23FE13FAB7" bold="true" box="[189,503,1260,1285]" pageId="37" pageNumber="238">Ecology and Distribution:</emphasis>
—M063A; M166A; M247A; M272A; M273A; M280A;
<materialsCitation id="73CFD561FF98FF9FFB21FB23FE8FFAFF" collectorName="Lake" location="Also" pageId="37" pageNumber="238" specimenCount="1" stateProvince="Hovsgol">
M330A: found in shallow to deep habitats throughout
<collectorName id="6E52BAEAFF98FF9FFE67FADFFE19FA9B" box="[453,509,1296,1321]" pageId="37" pageNumber="238">Lake</collectorName>
<collectingRegion id="016311DEFF98FF9FFDA7FADFFD80FA9B" box="[517,612,1296,1321]" country="Mongolia" name="Hovsgol" pageId="37" pageNumber="238">Hövsgöl</collectingRegion>
including epipelon,
<taxonomicName id="04A7A4BFFF98FF9FFCEFFADFFC71FA9B" box="[845,917,1296,1321]" class="Insecta" family="Noctuidae" genus="Chara" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="F1D3032EFF98FF9FFCEFFADFFC71FA9B" box="[845,917,1296,1321]" italics="true" pageId="37" pageNumber="238">Chara</emphasis>
</taxonomicName>
, marl, and sediments.
<location id="C67889E7FF98FF9FFB36FADFFB2DFA9B" LSID="urn:lsid:plazi:treatment:4B0E6E2AFF98FF9FFF2AFF51FE8BFAFF:C67889E7FF98FF9FFB36FADFFB2DFA9B" box="[1172,1225,1296,1321]" name="Also" pageId="37" pageNumber="238" stateProvince="Hovsgol">Also</location>
found in streams in
<collectingRegion id="016311DEFF98FF9FFF2AFAFBFE8FFAFF" box="[136,363,1332,1357]" country="Mongolia" name="Arhangay" pageId="37" pageNumber="238">Arkhangai province</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,135 @@
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<subSubSection id="8BBD8CB7FF9AFF9DFF2AFC9DFC9BFB7E" pageId="39" pageNumber="240" type="nomenclature">
<paragraph id="C318DF3CFF9AFF9DFF2AFC9DFD69FCDF" blockId="39.[136,653,850,877]" box="[136,653,850,877]" pageId="39" pageNumber="240">
<heading id="98506850FF9AFF9DFF2AFC9DFD69FCDF" box="[136,653,850,877]" fontSize="11" level="2" pageId="39" pageNumber="240" reason="3">
<taxonomicName id="04A7A4BFFF9AFF9DFF2AFC9DFE3EFCDF" authority="Petersen" authorityName="Petersen" box="[136,474,850,877]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="minuta">
<emphasis id="F1D3032EFF9AFF9DFF2AFC9DFF12FCDF" bold="true" box="[136,246,850,877]" italics="true" pageId="39" pageNumber="240">Diploneis</emphasis>
cf.
<emphasis id="F1D3032EFF9AFF9DFE83FC9DFE97FCDE" bold="true" box="[289,371,850,876]" italics="true" pageId="39" pageNumber="240">minuta</emphasis>
Petersen
</taxonomicName>
(
<figureCitation id="5B9CC3B9FF9AFF9DFE4AFC9BFD60FCDF" box="[488,644,852,877]" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Figs 254, 255</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFF9AFF9DFF2AFC5AFC9BFB7E" blockId="39.[136,1453,917,1228]" pageId="39" pageNumber="240">
The valves are linear-elliptical with parallel margins and round ends. The valve length is 18.0 μm and the valve breadth is 7.0 μm. The axial area is linear, slightly expanded into a small elongate central area, and also expanded at the valve apices. The central area is elongate, 1.4 μm wide. The longitudinal canal is linear, slightly expanded in the middle of the valve, and composed of one row of areolae throughout the whole length. Externally, the openings of the canal are covered with volate occlusions (
<figureCitation id="5B9CC3B9FF9AFF9DFEABFBDAFE41FB9E" box="[265,421,1045,1068]" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Figs 254, 255</figureCitation>
). At the valve apices the longitudinal canal is extended, circling the distal raphe endings (
<figureCitation id="5B9CC3B9FF9AFF9DFA22FBDAFF50FBFE" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Fig. 254</figureCitation>
). The raphe is straight with slightly expanded proximal ends, bent to the same side of the valve (
<figureCitation id="5B9CC3B9FF9AFF9DFB19FBFAFAFDFBFE" box="[1211,1305,1077,1100]" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Fig. 255</figureCitation>
). Distally, the raphe endings are simple and slightly expanded, terminating some distance from the valve mantle (
<figureCitation id="5B9CC3B9FF9AFF9DFB08FB9AFAECFBDE" box="[1194,1288,1109,1132]" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Fig. 254</figureCitation>
). The striae are parallel, becoming radiate towards the valve apices,
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10 μm. Externally, the alveoli open via a single areola covered with volate occlusions that occupy less than half of the valve width (
<figureCitation id="5B9CC3B9FF9AFF9DFCC5FB5AFC21FB1E" box="[871,965,1173,1196]" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Fig. 254</figureCitation>
). The other part of the alveoli is covered by a lanceolate hyaline area, merging with the longitudinal canal (
<figureCitation id="5B9CC3B9FF9AFF9DFCB7FB7AFC97FB7E" box="[789,883,1205,1228]" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Fig. 255</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9AFF9DFF2AFB33FC3FFA5F" pageId="39" pageNumber="240" type="description">
<paragraph id="C318DF3CFF9AFF9DFF2AFB33FC3FFA5F" blockId="39.[136,1452,1276,1553]" pageId="39" pageNumber="240">
<emphasis id="F1D3032EFF9AFF9DFF2AFB33FED4FAA7" bold="true" box="[136,304,1276,1301]" pageId="39" pageNumber="240">Observations:</emphasis>
—The population from Lake
<collectingRegion id="016311DEFF9AFF9DFDDCFB33FD39FAA7" box="[638,733,1276,1301]" country="Mongolia" name="Hovsgol" pageId="39" pageNumber="240">Hövsgöl</collectingRegion>
is characterized with narrow and linear longitudinal canal very similar to the longitudinal canal of
<taxonomicName id="04A7A4BFFF9AFF9DFDB0FAEFFC59FA8B" authority="Petersen (1928: 381" authorityName="Petersen" authorityYear="1928" box="[530,957,1312,1337]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="minuta">
<emphasis id="F1D3032EFF9AFF9DFDB0FAEFFD30FA8B" box="[530,724,1312,1337]" italics="true" pageId="39" pageNumber="240">Diploneis minuta</emphasis>
<bibRefCitation id="A736A2CDFF9AFF9DFD79FAEFFC59FA8B" author="Petersen, J. B." box="[731,957,1312,1337]" pageId="39" pageNumber="240" pagination="325 - 447" refId="ref28201" refString="Petersen, J. B. (1928) The aerial Algae of Iceland. In: Rosenvinge, L. K. &amp; Warming, E. (Eds.) The Botany of Iceland, vol. 2, part 2, no. 8. J. Frimodt, Kobenhavn, pp. 325 - 447." type="book chapter" year="1928">Petersen (1928: 381</bibRefCitation>
</taxonomicName>
, fig. 6). In the original description, Petersen gives a single valve measurement for
<taxonomicName id="04A7A4BFFF9AFF9DFD90FA8BFD47FAEF" box="[562,675,1348,1373]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="minuta">
<emphasis id="F1D3032EFF9AFF9DFD90FA8BFD47FAEF" box="[562,675,1348,1373]" italics="true" pageId="39" pageNumber="240">D. minuta</emphasis>
</taxonomicName>
(length: 13.0 μm and breadth: 4.4 μm). However, the valve found in Lake
<collectingRegion id="016311DEFF9AFF9DFF65FAA7FEC2FA33" box="[199,294,1384,1409]" country="Mongolia" name="Hovsgol" pageId="39" pageNumber="240">Hövsgöl</collectingRegion>
has larger dimensions (length: 18.0 μm and breadth: 7.0 μm), and as such is identified as
<taxonomicName id="04A7A4BFFF9AFF9DFAB3FAA7FA41FA33" box="[1297,1445,1384,1409]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="minuta">
<emphasis id="F1D3032EFF9AFF9DFAB3FAA7FAC1FA33" box="[1297,1317,1384,1409]" italics="true" pageId="39" pageNumber="240">D</emphasis>
. cf.
<emphasis id="F1D3032EFF9AFF9DFAF5FAA7FA41FA33" box="[1367,1445,1384,1409]" italics="true" pageId="39" pageNumber="240">minuta</emphasis>
</taxonomicName>
. Unfortunately the lack of additional LM observations makes more detailed comparison rather difficult. The SEM illustrations for
<taxonomicName id="04A7A4BFFF9AFF9DFEE2FA7FFE57FA7B" box="[320,435,1456,1481]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="minuta">
<emphasis id="F1D3032EFF9AFF9DFEE2FA7FFE57FA7B" box="[320,435,1456,1481]" italics="true" pageId="39" pageNumber="240">D. minuta</emphasis>
</taxonomicName>
given in
<bibRefCitation id="A736A2CDFF9AFF9DFD81FA7FFC78FA7B" author="Werum, M. &amp; Lange-Bertalot, H." box="[547,924,1456,1481]" pageId="39" pageNumber="240" pagination="3 - 417" refId="ref28669" refString="Werum, M. &amp; Lange-Bertalot, H. (2004) Diatoms in Springs from Central European. Iconographia Diatomologica 13: 3 - 417." type="journal article" year="2004">Werum &amp; Lange-Bertalot (2004)</bibRefCitation>
nicely fit the population from Lake
<collectingRegion id="016311DEFF9AFF9DFAE4FA7FFA4DFA7B" box="[1350,1449,1456,1481]" country="Mongolia" name="Hovsgol" pageId="39" pageNumber="240">Hövsgöl</collectingRegion>
. Further LM and SEM analyses are necessary to properly identify this taxon.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9AFF9DFF1FFA37FAF7F9A3" box="[189,1299,1528,1553]" pageId="39" pageNumber="240" type="biology_ecology">
<paragraph id="C318DF3CFF9AFF9DFF1FFA37FAF7F9A3" blockId="39.[136,1452,1276,1553]" box="[189,1299,1528,1553]" pageId="39" pageNumber="240">
<emphasis id="F1D3032EFF9AFF9DFF1FFA37FE17F9A3" bold="true" box="[189,499,1528,1553]" pageId="39" pageNumber="240">Ecology and Distribution:</emphasis>
—M276A: found only epilithic at
<quantity id="045F72D9FF9AFF9DFCD6FA37FC7BF9A3" box="[884,927,1528,1553]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="240" unit="m" value="5.0">5 m</quantity>
depth in northern Lake
<collectingRegion id="016311DEFF9AFF9DFB0FFA37FAF4F9A3" box="[1197,1296,1528,1553]" country="Mongolia" name="Hovsgol" pageId="39" pageNumber="240">Hövsgöl</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="C318DF3CFF9AFF9DFF2AFEC5FC65FE97" blockId="39.[136,897,266,293]" box="[136,897,266,293]" pageId="39" pageNumber="240">
<heading id="98506850FF9AFF9DFF2AFEC5FC65FE97" box="[136,897,266,293]" fontSize="11" level="2" pageId="39" pageNumber="240" reason="3">
<taxonomicName id="04A7A4BFFF9AFF9DFF2AFEC5FCEEFE97" authority="Metzeltin, Lange-Bertalot &amp; Nergui" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[136,778,266,293]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="eximia">
<emphasis id="F1D3032EFF9AFF9DFF2AFEC5FF12FE97" bold="true" box="[136,246,266,293]" italics="true" pageId="39" pageNumber="240">Diploneis</emphasis>
cf.
<emphasis id="F1D3032EFF9AFF9DFE83FEC5FE8BFE96" bold="true" box="[289,367,266,292]" italics="true" pageId="39" pageNumber="240">eximia</emphasis>
Metzeltin, Lange-Bertalot &amp; Nergui
</taxonomicName>
(
<figureCitation id="5B9CC3B9FF9AFF9DFCBBFEC3FC9DFE97" box="[793,889,268,293]" captionStart="FIGURES 253271" captionStartId="41.[136,243,1874,1895]" captionTargetBox="[252,1335,247,1846]" captionTargetId="figure-37@41.[250,1338,245,1848]" captionTargetPageId="41" captionText="FIGURES 253271. LM and SEM external valve views. Fig. 253. Diploneis cf. eximia, LM valve view. Figs 254, 255. Diploneis cf. minuta, SEM external valve views. Fig. 254. Valve view. Fig. 255. Proximal raphe endings. Figs 256259. Diploneis sp. 1. Fig. 256. LM valve view. Fig. 257. Central area with proximal raphe endings, SEM. Fig. 258. Distal raphe endings, SEM. Fig. 259. View of the entire valve, SEM. Figs 260269.Diploneis sp. 2, LM valve views. Fig. 270. Diploneis sp. 3, LM valve view.Fig. 271. Diploneis sp. 3, LM valve view. Scale bars = 10 μm (Figs 253, 256, 260271); 5 μm (Figs 254, 259); 2 μm (Figs 255, 257, 258)." figureDoi="http://doi.org/10.5281/zenodo.13634182" httpUri="https://zenodo.org/record/13634182/files/figure.png" pageId="39" pageNumber="240">Fig. 253</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFF9AFF9DFF2AFE82FF57FDB6" blockId="39.[136,1452,333,516]" pageId="39" pageNumber="240">
The valve is elliptical to linear-elliptic with slightly convex margins and bluntly rounded ends. The valve length is 67.5 μm, and the breadth 27.5 μm. The axial area is narrow and parallel, slightly expanded near the large transapically elliptical central area, which is 7.0 μm wide. The longitudinal canal is lanceolate to linear expanding in the middle with two to three rows of areolae that coalescence into one to two towards the valve apices. The raphe branches are straight and simple with distinctly expanded proximal ends. The striae are radiate,
<quantity id="045F72D9FF9AFF9DFD24FE02FD58FE56" box="[646,700,461,484]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="39" pageNumber="240" unit="in" value="10.0">10 in</quantity>
10 μm, composed of one row of round to rectangular areolae,
<quantity id="045F72D9FF9AFF9DFAF7FE02FA6FFE56" box="[1365,1419,461,484]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="39" pageNumber="240" unit="in" value="10.0">10 in</quantity>
10 μm.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9AFF9DFF2AFDFBFE03FD6F" pageId="39" pageNumber="240" type="description">
<paragraph id="C318DF3CFF9AFF9DFF2AFDFBFE03FD6F" blockId="39.[136,1452,564,805]" pageId="39" pageNumber="240">
<emphasis id="F1D3032EFF9AFF9DFF2AFDFBFEA8FDFF" bold="true" box="[136,332,564,589]" pageId="39" pageNumber="240">Observations:—</emphasis>
LM images of
<taxonomicName id="04A7A4BFFF9AFF9DFDA2FDFBFD27FDFF" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[512,707,564,589]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="eximia">
<emphasis id="F1D3032EFF9AFF9DFDA2FDFBFD27FDFF" box="[512,707,564,589]" italics="true" pageId="39" pageNumber="240">Diploneis eximia</emphasis>
</taxonomicName>
are not included in the original description, and therefore it is rather difficult to make comparisons (
<bibRefCitation id="A736A2CDFF9AFF9DFD90FD97FCC1FDC3" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[562,805,600,625]" pageId="39" pageNumber="240" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9AFF9DFD05FD97FD3DFDC3" box="[679,729,600,625]" italics="true" pageId="39" pageNumber="240">et al</emphasis>
. 2009
</bibRefCitation>
). Due to the similar valve outline and the structure of the longitudinal canal the population from Lake
<collectingRegion id="016311DEFF9AFF9DFDD9FDB3FD3EFD27" box="[635,730,636,661]" country="Mongolia" name="Hovsgol" pageId="39" pageNumber="240">Hövsgöl</collectingRegion>
is identified as
<taxonomicName id="04A7A4BFFF9AFF9DFC28FDB3FB89FD27" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[906,1133,636,661]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="eximia">
<emphasis id="F1D3032EFF9AFF9DFC28FDB3FC13FD27" box="[906,1015,636,661]" italics="true" pageId="39" pageNumber="240">Diploneis</emphasis>
cf.
<emphasis id="F1D3032EFF9AFF9DFB80FDB3FB89FD27" box="[1058,1133,636,661]" italics="true" pageId="39" pageNumber="240">eximia</emphasis>
</taxonomicName>
. Further analysis of the
<typeStatus id="1C1C619EFF9AFF9DFAD9FDB3FA4FFD27" box="[1403,1451,636,661]" pageId="39" pageNumber="240">type</typeStatus>
material is preferable in order to fully understand the identity of
<taxonomicName id="04A7A4BFFF9AFF9DFCF4FD6FFC27FD0B" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[854,963,672,697]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="eximia">
<emphasis id="F1D3032EFF9AFF9DFCF4FD6FFC8EFD0B" box="[854,874,672,697]" italics="true" pageId="39" pageNumber="240">D</emphasis>
.
<emphasis id="F1D3032EFF9AFF9DFCDAFD6FFC27FD0B" box="[888,963,672,697]" italics="true" pageId="39" pageNumber="240">eximia</emphasis>
</taxonomicName>
and its potential relationship to populations here identified as
<taxonomicName id="04A7A4BFFF9AFF9DFEF2FD0BFE05FD6F" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[336,481,708,733]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="39" pageNumber="240" phylum="Ochrophyta" rank="species" species="eximia">
<emphasis id="F1D3032EFF9AFF9DFEF2FD0BFE80FD6F" box="[336,356,708,733]" italics="true" pageId="39" pageNumber="240">D</emphasis>
. cf.
<emphasis id="F1D3032EFF9AFF9DFE34FD0BFE05FD6F" box="[406,481,708,733]" italics="true" pageId="39" pageNumber="240">eximia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9AFF9DFF1FFD27FD36FC97" pageId="39" pageNumber="240" type="biology_ecology">
<paragraph id="C318DF3CFF9AFF9DFF1FFD27FD36FC97" blockId="39.[136,1452,564,805]" pageId="39" pageNumber="240">
<emphasis id="F1D3032EFF9AFF9DFF1FFD27FE17FCB3" bold="true" box="[189,499,744,769]" pageId="39" pageNumber="240">Ecology and Distribution:</emphasis>
—M331A: found in shallow sediments of central Lake
<collectingRegion id="016311DEFF9AFF9DFBC3FD27FB24FCB3" box="[1121,1216,744,769]" country="Mongolia" name="Hovsgol" pageId="39" pageNumber="240">Hövsgöl</collectingRegion>
and tychoplanktonic in a lake in
<collectingRegion id="016311DEFF9AFF9DFEA9FCC3FE2CFC97" box="[267,456,780,805]" country="Mongolia" name="Hentiy" pageId="39" pageNumber="240">Khentii province</collectingRegion>
(
<bibRefCitation id="A736A2CDFF9AFF9DFE74FCC3FD22FC97" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[470,710,780,805]" pageId="39" pageNumber="240" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9AFF9DFDE8FCC3FD9FFC97" box="[586,635,780,805]" italics="true" pageId="39" pageNumber="240">et al</emphasis>
. 2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="FCB8B5DDB615F2B025903C7CFA10182B">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<mods:namePart id="AC4FEE81E753B8D8BF1FC57315F335B3">Jovanovska, Elena</mods:namePart>
<mods:affiliation id="60BCDAE808EFD4B067AE26357DC9ACEF">Department of Animal Ecology and Systematics, Justus-Liebig University, Heinrich-Buff-Ring 26 - 32, 35392 Giessen, Germany</mods:affiliation>
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<mods:namePart id="448AB1B8F42B2F5CB9080FB9D9B19F35">Edlund, Mark B.</mods:namePart>
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<taxonomicName id="04A7A4BFFF9DFF9AFF2AFDBDFCA4FD3F" authority="Metzeltin, Lange-Bertalot &amp; Nergui" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[136,832,626,653]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="linearielliptica">
<emphasis id="F1D3032EFF9DFF9AFF2AFDBDFE42FD3F" bold="true" box="[136,422,626,653]" italics="true" pageId="32" pageNumber="233">Diploneis linearielliptica</emphasis>
Metzeltin, Lange-Bertalot &amp; Nergui
</taxonomicName>
(
<figureCitation id="5B9CC3B9FF9DFF9AFCEDFDBBFC0FFD3F" box="[847,1003,628,653]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 184196</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFF9DFF9AFF2AFD7AFD0BFBFE" blockId="32.[136,1452,693,1100]" pageId="32" pageNumber="233">
The valves are linear-elliptical with slightly convex margins and round ends (
<figureCitation id="5B9CC3B9FF9DFF9AFC66FD7AFBB8FD7E" box="[964,1116,693,716]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 184191</figureCitation>
). The valve length is 21.034.5 μm, and the valve breadth is 7.010.5 μm. The axial area is linear to lanceolate, merging with and into a distinctly enlarged round central nodule. From inside, a silica plate covers the whole length of the lanceolate longitudinal canal (
<figureCitation id="5B9CC3B9FF9DFF9AFAA8FD3AFA44FCBE" box="[1290,1440,757,780]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 192, 194</figureCitation>
). The central area is 3.04.5 μm wide. From the internal side, the central nodule is raised and distinct (
<figureCitation id="5B9CC3B9FF9DFF9AFB74FCDAFA8BFC9E" box="[1238,1391,789,812]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 192194</figureCitation>
). The longitudinal canal is narrow and linear, expanded in the middle of the valve, with one row of areolae throughout the whole valve length. From inside the longitudinal canal is covered with a silica plate, forming a “depression” where the raphe is placed (
<figureCitation id="5B9CC3B9FF9DFF9AFF7EFCBAFE90FC3E" box="[220,372,885,908]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 192, 194</figureCitation>
). Externally, the raphe is straight and simple, positioned with an expanded depression. Internally, the raphe is straight, inserted in a slightly elevated sternum inside the “depression” formed by the longitudinal canal (
<figureCitation id="5B9CC3B9FF9DFF9AFAE5FC5AFF50FC7E" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 192, 194</figureCitation>
). The proximal raphe ends are simple and terminate in small drop-like terminal fissures at the margins of the central area, but are not raised onto the central area itself (
<figureCitation id="5B9CC3B9FF9DFF9AFDC8FC1AFCE5FC5E" box="[618,769,981,1004]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 193, 194</figureCitation>
). Distally, the raphe endings are raised into small helictoglossae (
<figureCitation id="5B9CC3B9FF9DFF9AFF32FC3AFEC3FBBE" box="[144,295,1013,1036]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 195, 196</figureCitation>
). The striae are radiate,
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10 μm, composed of round areolae,
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10 μm. Internally, the alveoli each open via a single oval and continuous opening (
<figureCitation id="5B9CC3B9FF9DFF9AFD65FBDAFC85FB9E" box="[711,865,1045,1068]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 192, 194</figureCitation>
). This species is characterized with small hyaline area near the longitudinal canal (see arrows on
<figureCitation id="5B9CC3B9FF9DFF9AFDEFFBFAFD07FBFE" box="[589,739,1077,1100]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 186, 188</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9DFF9AFF2AFBB3FF2FFADF" pageId="32" pageNumber="233" type="description">
<paragraph id="C318DF3CFF9DFF9AFF2AFBB3FF2FFADF" blockId="32.[136,1452,1148,1497]" pageId="32" pageNumber="233">
<emphasis id="F1D3032EFF9DFF9AFF2AFBB3FED4FB27" bold="true" box="[136,304,1148,1173]" pageId="32" pageNumber="233">Observations:</emphasis>
<taxonomicName id="04A7A4BFFF9DFF9AFEEEFBB3FD8DFB27" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[332,617,1148,1173]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="linearielliptica">
<emphasis id="F1D3032EFF9DFF9AFEEEFBB3FD8DFB27" box="[332,617,1148,1173]" italics="true" pageId="32" pageNumber="233">Diploneis linearielliptica</emphasis>
</taxonomicName>
is described from Mongolian lakes in
<collectingRegion id="016311DEFF9DFF9AFB88FBB3FB0DFB27" box="[1066,1257,1148,1173]" country="Mongolia" name="Hentiy" pageId="32" pageNumber="233">Khentii province</collectingRegion>
(
<bibRefCitation id="A736A2CDFF9DFF9AFB58FBB3FF21FB0B" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." pageId="32" pageNumber="233" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9DFF9AFAD3FBB3FA41FB27" box="[1393,1445,1148,1173]" italics="true" pageId="32" pageNumber="233">et al</emphasis>
. 2009
</bibRefCitation>
). The population from the
<typeStatus id="1C1C619EFF9DFF9AFE4CFB6FFDFAFB0B" box="[494,542,1184,1209]" pageId="32" pageNumber="233">type</typeStatus>
locality is characterized by a narrow size range in terms of that observed in Lake
<collectingRegion id="016311DEFF9DFF9AFF2AFB0BFF03FB6F" box="[136,231,1220,1245]" country="Mongolia" name="Hovsgol" pageId="32" pageNumber="233">Hövsgöl</collectingRegion>
and
<collectingRegion id="016311DEFF9DFF9AFE80FB0BFE7CFB6F" box="[290,408,1220,1245]" country="Mongolia" name="Arhangay" pageId="32" pageNumber="233">Arkhangai</collectingRegion>
populations (length: 32.046.0 μm
<emphasis id="F1D3032EFF9DFF9AFC9BFB0BFCB4FB6F" box="[825,848,1220,1245]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
. 21.034.5 μm, breadth: 10.014.0 μm
<emphasis id="F1D3032EFF9DFF9AFA83FB0BFADCFB6F" box="[1313,1336,1220,1245]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
. 7.010.5 μm). The stria density is an additional character that distinguishes these populations (
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10 μm
<emphasis id="F1D3032EFF9DFF9AFAA3FB27FAFCFAB3" box="[1281,1304,1256,1281]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
.
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10 μm). However, these differences may be the result of an insufficient number of observed valves in the material. SEM analyses of the valve exterior are needed to examine the potential narrow hyaline structure alongside the longitudinal canal.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9DFF9AFF1FFAB7FEF4FA6B" pageId="32" pageNumber="233" type="biology_ecology">
<paragraph id="C318DF3CFF9DFF9AFF1FFAB7FEF4FA6B" blockId="32.[136,1452,1148,1497]" pageId="32" pageNumber="233">
<emphasis id="F1D3032EFF9DFF9AFF1FFAB7FE1FFA23" bold="true" box="[189,507,1400,1425]" pageId="32" pageNumber="233">Ecology and Distribution:</emphasis>
—M052A; M166A; M167A;
<materialsCitation id="73CFD561FF9DFF9AFCF7FAB7FEE8FA6B" elevation="1" location="southern Lake" pageId="32" pageNumber="233" specimenCount="1" stateProvince="Hovsgol">
M262A: found in shallow habitats (&lt;
<elevation id="488A380FFF9DFF9AFAAFFAB7FA67FA23" box="[1293,1411,1400,1425]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="233" unit="m" value="1.0">
<quantity id="045F72D9FF9DFF9AFAAFFAB7FADCFA23" box="[1293,1336,1400,1425]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="233" unit="m" value="1.0">1 m</quantity>
depth
</elevation>
) in central and
<location id="C67889E7FF9DFF9AFEABFA53FE43FA07" LSID="urn:lsid:plazi:treatment:4B0E6E2AFF9DFF9AFF2AFDBDFEF4FA6B:C67889E7FF9DFF9AFEABFA53FE43FA07" box="[265,423,1436,1461]" name="southern Lake" pageId="32" pageNumber="233" stateProvince="Hovsgol">southern Lake</location>
<collectingRegion id="016311DEFF9DFF9AFE0FFA53FDE8FA07" box="[429,524,1436,1461]" country="Mongolia" name="Hovsgol" pageId="32" pageNumber="233">Hövsgöl</collectingRegion>
and also in streams in
<collectingRegion id="016311DEFF9DFF9AFCA7FA53FC00FA07" box="[773,996,1436,1461]" country="Mongolia" name="Arhangay" pageId="32" pageNumber="233">Arkhangai province</collectingRegion>
and lakes of
<collectingRegion id="016311DEFF9DFF9AFBD7FA53FAD5FA07" box="[1141,1329,1436,1461]" country="Mongolia" name="Hentiy" pageId="32" pageNumber="233">Khentii province</collectingRegion>
(
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Metzeltin
<emphasis id="F1D3032EFF9DFF9AFF2AFA0FFF5DFA6B" box="[136,185,1472,1497]" italics="true" pageId="32" pageNumber="233">et al</emphasis>
. 2009
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C7B8B15FE1642E4E4EA3A2B64006447D">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<subSubSection id="8BBD8CB7FF9DFF9AFF2AF9C9FBA3F952" pageId="32" pageNumber="233" type="nomenclature">
<paragraph id="C318DF3CFF9DFF9AFF2AF9C9FCE0F993" blockId="32.[136,772,1542,1569]" box="[136,772,1542,1569]" pageId="32" pageNumber="233">
<heading id="98506850FF9DFF9AFF2AF9C9FCE0F993" box="[136,772,1542,1569]" fontSize="11" level="2" pageId="32" pageNumber="233" reason="3">
<taxonomicName id="04A7A4BFFF9DFF9AFF2AF9C9FDB5F993" authority="Lange-Bertalot" authorityName="Lange-Bertalot" authorityYear="2004" box="[136,593,1542,1569]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="fontanella">
<emphasis id="F1D3032EFF9DFF9AFF2AF9C9FF12F993" bold="true" box="[136,246,1542,1569]" italics="true" pageId="32" pageNumber="233">Diploneis</emphasis>
aff.
<emphasis id="F1D3032EFF9DFF9AFE88F9C9FE44F993" bold="true" box="[298,416,1542,1569]" italics="true" pageId="32" pageNumber="233">fontanella</emphasis>
Lange-Bertalot
</taxonomicName>
(
<figureCitation id="5B9CC3B9FF9DFF9AFDC2F9C7FD18F993" box="[608,764,1544,1569]" captionStart="FIGURES 184205" captionStartId="33.[136,243,1961,1982]" captionTargetBox="[172,1416,248,1930]" captionTargetId="figure-38@33.[168,1420,245,1936]" captionTargetPageId="33" captionText="FIGURES 184205. LM and SEM internal valve views. Figs 184196. Diploneis linearielliptica. Figs 184191. LM valve views. Figs 192, 194. View of the whole valve. Fig. 193. Central area with proximal raphe endings. Figs 195, 196. Distal raphe endings. Figs 197205. Diploneis aff. fontanella, LM valve views. Scale bars = 10 μm (Figs 184192, 197205); 5 μm (Fig. 194); 2 μm (Figs 193, 195, 196)." figureDoi="http://doi.org/10.5281/zenodo.13634174" httpUri="https://zenodo.org/record/13634174/files/figure.png" pageId="32" pageNumber="233">Figs 197205</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFF9DFF9AFF2AF986FBA3F952" blockId="32.[136,1452,1609,1760]" pageId="32" pageNumber="233">
The valves are elliptical with slightly convex margins and broadly rounded ends. The valve length is 17.022.0 μm, and the valve breadth is 8.09.0 μm. The axial area is narrow, linear to lanceolate expanding toward the central area. The central area is round, 2.53.5 μm wide. The longitudinal canal is narrow, linear to lanceolate, and expanded in the middle of the valve, with one row of areolae throughout the whole length. The raphe is straight and simple, positioned with an expanded depression. The striae are radiate,
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10 μm, composed of round areolae,
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10 μm.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9DFF9AFF2AF8DFFE14F797" pageId="32" pageNumber="233" type="description">
<paragraph id="C318DF3CFF9DFF9AFF2AF8DFFE14F797" blockId="32.[136,1452,1808,2085]" pageId="32" pageNumber="233">
<emphasis id="F1D3032EFF9DFF9AFF2AF8DFFED4F89B" bold="true" box="[136,304,1808,1833]" pageId="32" pageNumber="233">Observations:</emphasis>
—The observed population is very similar to
<taxonomicName id="04A7A4BFFF9DFF9AFC8BF8DFFBEAF89B" authorityName="Lange-Bertalot" authorityYear="2004" box="[809,1038,1808,1833]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="fontanella">
<emphasis id="F1D3032EFF9DFF9AFC8BF8DFFBEAF89B" box="[809,1038,1808,1833]" italics="true" pageId="32" pageNumber="233">Diploneis fontanella</emphasis>
</taxonomicName>
, from which it differs in stria density (
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10 μm
<emphasis id="F1D3032EFF9DFF9AFEF3F8FBFE8CF8FF" box="[337,360,1844,1869]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
.
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10 μm) and areola density (
<quantity id="045F72D9FF9DFF9AFC82F8FBFC62F8FF" box="[800,902,1844,1869]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="7.62" metricValueMin="6.35" pageId="32" pageNumber="233" unit="in" value="27.5" valueMax="30.0" valueMin="25.0">2530 in</quantity>
10 μm
<emphasis id="F1D3032EFF9DFF9AFC43F8FBFC1CF8FF" box="[993,1016,1844,1869]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
.
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10 μm). The valve shape is an additional feature when distinguishing this
<collectingRegion id="016311DEFF9DFF9AFD3BF897FCEBF8C3" box="[665,783,1880,1905]" country="Mongolia" name="Arhangay" pageId="32" pageNumber="233">Arkhangai</collectingRegion>
population from the
<typeStatus id="1C1C619EFF9DFF9AFBA6F897FBD0F8C3" box="[1028,1076,1880,1905]" pageId="32" pageNumber="233">type</typeStatus>
(elliptical
<emphasis id="F1D3032EFF9DFF9AFB10F897FB2DF8C3" box="[1202,1225,1880,1905]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
. elliptical to linear-elliptic). In
<collectingRegion id="016311DEFF9DFF9AFEA9F8B3FE2CF827" box="[267,456,1916,1941]" country="Mongolia" name="Hentiy" pageId="32" pageNumber="233">Khentii province</collectingRegion>
<bibRefCitation id="A736A2CDFF9DFF9AFE6DF8B3FD2EF827" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[463,714,1916,1941]" pageId="32" pageNumber="233" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9DFF9AFDE3F8B3FD96F827" box="[577,626,1916,1941]" italics="true" pageId="32" pageNumber="233">et al</emphasis>
. (2009)
</bibRefCitation>
identified a population of
<taxonomicName id="04A7A4BFFF9DFF9AFC51F8B3FA48F827" authority="Reichardt &amp; Lange-Bertalot" authorityName="Reichardt &amp; Lange-Bertalot" authorityYear="2004" box="[1011,1452,1916,1941]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="fontium">
<emphasis id="F1D3032EFF9DFF9AFC51F8B3FB8FF827" box="[1011,1131,1916,1941]" italics="true" pageId="32" pageNumber="233">D. fontium</emphasis>
Reichardt &amp; Lange-Bertalot
</taxonomicName>
(2004: pl. 4, fig. 16, pl. 5, fig. 17), which morphologically is similar to
<taxonomicName id="04A7A4BFFF9DFF9AFC68F86FFB61F80B" authorityName="Lange-Bertalot" authorityYear="2004" box="[970,1157,1952,1977]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="fontinella">
<emphasis id="F1D3032EFF9DFF9AFC68F86FFC3AF80B" box="[970,990,1952,1977]" italics="true" pageId="32" pageNumber="233">D</emphasis>
. aff.
<emphasis id="F1D3032EFF9DFF9AFBB8F86FFB61F80B" box="[1050,1157,1952,1977]" italics="true" pageId="32" pageNumber="233">fontinella</emphasis>
</taxonomicName>
. The valve measurements of the
<collectingRegion id="016311DEFF9DFF9AFF6FF80BFEA7F86F" box="[205,323,1988,2013]" country="Mongolia" name="Arhangay" pageId="32" pageNumber="233">Arkhangai</collectingRegion>
population more closely resemble
<taxonomicName id="04A7A4BFFF9DFF9AFD6BF80BFCB8F86F" authorityName="Lange-Bertalot" authorityYear="2004" box="[713,860,1988,2013]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="fontanella">
<emphasis id="F1D3032EFF9DFF9AFD6BF80BFCB8F86F" box="[713,860,1988,2013]" italics="true" pageId="32" pageNumber="233">D. fontanella</emphasis>
</taxonomicName>
than
<taxonomicName id="04A7A4BFFF9DFF9AFC3BF80BFBF4F86F" authorityName="Reichardt &amp; Lange-Bertalot" authorityYear="2004" box="[921,1040,1988,2013]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="fontium">
<emphasis id="F1D3032EFF9DFF9AFC3BF80BFBF4F86F" box="[921,1040,1988,2013]" italics="true" pageId="32" pageNumber="233">D. fontium</emphasis>
</taxonomicName>
, therefore we have identified it as
<taxonomicName id="04A7A4BFFF9DFF9AFA32F80BFEC5F7B3" authorityName="Lange-Bertalot" authorityYear="2004" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="fontinella">
<emphasis id="F1D3032EFF9DFF9AFA32F80BFA40F86F" box="[1424,1444,1988,2013]" italics="true" pageId="32" pageNumber="233">D</emphasis>
. aff.
<emphasis id="F1D3032EFF9DFF9AFF14F827FEC5F7B3" box="[182,289,2024,2049]" italics="true" pageId="32" pageNumber="233">fontinella</emphasis>
</taxonomicName>
. In order to test the conspecificity between these two species, SEM analyses are necessary and should be the subject of further studies.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9CFF9BFF1FFF6FFCC4FF6F" pageId="33" pageNumber="234" type="biology_ecology">
<paragraph id="C318DF3CFF9CFF9BFF1FFF6FFA48FF0B" blockId="33.[136,1452,160,221]" box="[189,1452,160,185]" pageId="33" pageNumber="234">
<emphasis id="F1D3032EFF9CFF9BFF1FFF6FFE0AFF0B" bold="true" box="[189,494,160,185]" pageId="33" pageNumber="234">Ecology and Distribution:</emphasis>
—M166A; M167A: not found in Lake
<collectingRegion id="016311DEFF9CFF9BFC36FF6FFC17FF0B" box="[916,1011,160,185]" country="Mongolia" name="Hovsgol" pageId="33" pageNumber="234">Hövsgöl</collectingRegion>
nor previously reported from
<collectingCountry id="BBB09FACFF9CFF9BFA95FF6FFA4CFF0B" box="[1335,1448,160,185]" name="Mongolia" pageId="33" pageNumber="234">Mongolia</collectingCountry>
.
</paragraph>
<paragraph id="C318DF3CFF9CFF9BFF2AFF0BFCC4FF6F" blockId="33.[136,1452,160,221]" box="[136,800,196,221]" pageId="33" pageNumber="234">
Collections are from a small stream in
<collectingRegion id="016311DEFF9CFF9BFD9BFF0BFCF8FF6F" box="[569,796,196,221]" country="Mongolia" name="Arhangay" pageId="33" pageNumber="234">Arkhangai province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="E5DF4783D31912D84B5137434C4C7D65" ID-DOI="10.11646/phytotaxa.217.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13634131" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725325955495" checkinUser="felipe" docAuthor="Jovanovska, Elena, Levkov, Zlatko &amp; Edlund, Mark B." docDate="2015" docId="4B0E6E2AFF9EFF9EFF2AFF51FCEAFEFB" docLanguage="en" docName="phytotaxa.217.3.1.pdf" docOrigin="Phytotaxa 217 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.217.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Diploneis heteromorphiforma Metzeltin, Lange-Bertalot &amp; Nergui 2009" docType="treatment" docVersion="3" lastPageNumber="237" masterDocId="B7371652FFBDFFBAFFA2FFCFFFE4FFB2" masterDocTitle="The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia" masterLastPageNumber="248" masterPageNumber="201" pageNumber="236" updateTime="1725326923387" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B3C244C194E9F776947D1076EBD69DBC">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<mods:namePart id="2934BE009E9EA45540397905E8FB34D5">Jovanovska, Elena</mods:namePart>
<mods:affiliation id="8390EEA180E0660DC5C92D271FBF92B1">Department of Animal Ecology and Systematics, Justus-Liebig University, Heinrich-Buff-Ring 26 - 32, 35392 Giessen, Germany</mods:affiliation>
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<mods:namePart id="5CB779168012D59B8ACAA38BA2C2EDB6">Levkov, Zlatko</mods:namePart>
<mods:affiliation id="A33BD4F73C7CF58FA8363CF83154A4E0">Institute of Biology, Faculty of Natural Sciences, Arhimedova 3, 1000 Skopje, R. of Macedonia</mods:affiliation>
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<mods:namePart id="D214845FF6806F1B5A65EE2B7AF9CC8E">Edlund, Mark B.</mods:namePart>
<mods:affiliation id="91255741C809FF7F7931B6D3E01E5119">St. Croix Watershed Research Station, Science Museum of Minnesota, Marine on St. Croix, Minnesota 55047, USA</mods:affiliation>
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<subSubSection id="8BBD8CB7FF9EFF99FF2AFF51FC90FCCA" pageId="35" pageNumber="236" type="nomenclature">
<paragraph id="C318DF3CFF9EFF99FF2AFF51FB2FFF0B" blockId="35.[136,1227,158,185]" box="[136,1227,158,185]" pageId="35" pageNumber="236">
<heading id="98506850FF9EFF99FF2AFF51FB2FFF0B" box="[136,1227,158,185]" fontSize="11" level="2" pageId="35" pageNumber="236" reason="3">
<taxonomicName id="04A7A4BFFF9EFF99FF2AFF51FC4CFF0B" authority="Metzeltin, Lange-Bertalot &amp; Nergui" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[136,936,158,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FF2AFF51FF12FF0B" bold="true" box="[136,246,158,185]" italics="true" pageId="35" pageNumber="236">Diploneis</emphasis>
aff.
<emphasis id="F1D3032EFF9EFF99FE88FF51FDE9FF0B" bold="true" box="[298,525,158,185]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
Metzeltin, Lange-Bertalot &amp; Nergui
</taxonomicName>
(Figs 206210, 213219)
</heading>
</paragraph>
<paragraph id="C318DF3CFF9EFF99FF2AFF2EFC90FCCA" blockId="35.[136,1452,225,888]" pageId="35" pageNumber="236">
The valves are linear-elliptical with slightly convex margins and round ends (Figs 206210). The valve length is 16.029.0 μm, and the valve breadth is 8.511.5 μm. The axial area is narrow, lanceolate, and expands toward the round to elongate central area. From inside, a thick lanceolate silica plate covers the longitudinal canal (Fig. 216). The central area is round to elongate, 2.54.5 μm wide. The longitudinal canal is narrow, linear to lanceolate, expanded in mid-valve, composed in parts with two rows of areolae and in parts with one row of areolae (Fig. 214). The external openings of the canal areolae are covered with cribra, similar to those in the striae and from which they are separated by a thin hyaline area (Figs 214, 215). Internally, the longitudinal canal is closed with thick silica plate throughout its length, forming a “depression” where the raphe is located (Fig. 216). Externally, the raphe is simple and straight with slightly curved and expanded central pores (Figs 213, 214). The distal raphe branches are deflected, and the terminal fissures are slightly bent into a drop-like shape (Figs 214, 215). Internally, the raphe is straight with simple and slightly expanded proximal ends, inserted in a slightly elevated sternum inside the wide “depression” formed by the longitudinal canal (Fig. 216). The distal raphe endings are slightly curved (Fig. 218). Only the proximal raphe endings reach the height of the canal itself (Fig. 217). The striae are radiate and uniseriate becoming biseriate toward the valve margins,
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10 μm. The striae areolae are round and occluded externally with cribra,
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10 μm. The areolae are barely visible with LM observations, leaving the impression that each alveolate striae is composed of two rows of areolae (Fig. 206). In highly corroded valves it seems that from the valve exterior the alveolus opens though a single opening covered with cribrate occlusions (Fig. 219). The alveoli each open to the valve interior via a single continuous opening (Figs 216218). However, from the inside, it appears that each alveolus initially opens toward the valve exterior via three areolae aligned in staggered transapical rows (Figs 217, 218), then transition to the cell exterior through a layer of one or two cribrate occlusions that must each cover multiple internally staggered areolae (213215). The external openings of the areolae (uniseriate becoming biseriate) and the LM observations (biseriate throughout) is misleading as to the detailed structure of the alveolate striae and their areolae.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9EFF9EFF2AFC67FCCBFEB3" lastPageId="36" lastPageNumber="237" pageId="35" pageNumber="236" type="description">
<paragraph id="C318DF3CFF9EFF99FF2AFC67FEA2FBC7" blockId="35.[135,1452,936,2077]" pageId="35" pageNumber="236">
<emphasis id="F1D3032EFF9EFF99FF2AFC67FEA8FC73" bold="true" box="[136,332,936,961]" pageId="35" pageNumber="236">Observations:—</emphasis>
<taxonomicName id="04A7A4BFFF9EFF99FEEEFC67FD29FC73" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[332,717,936,961]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FEEEFC67FE5DFC73" box="[332,441,936,961]" italics="true" pageId="35" pageNumber="236">Diploneis</emphasis>
aff.
<emphasis id="F1D3032EFF9EFF99FE51FC67FD29FC73" box="[499,717,936,961]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="04A7A4BFFF9EFF99FC08FC67FB4DFC73" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[938,1193,936,961]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FC08FC67FB4DFC73" box="[938,1193,936,961]" italics="true" pageId="35" pageNumber="236">D. heteromorphiforma</emphasis>
</taxonomicName>
, described from Zyyn byrkh River in
<collectingRegion id="016311DEFF9EFF99FE94FC03FE11FC57" box="[310,501,972,997]" country="Mongolia" name="Hentiy" pageId="35" pageNumber="236">Khentii province</collectingRegion>
in
<collectingCountry id="BBB09FACFF9EFF99FDB9FC03FD6CFC57" box="[539,648,972,997]" name="Mongolia" pageId="35" pageNumber="236">Mongolia</collectingCountry>
(
<bibRefCitation id="A736A2CDFF9EFF99FD3AFC03FC68FC57" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[664,908,972,997]" pageId="35" pageNumber="236" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9EFF99FCACFC03FCA4FC57" box="[782,832,972,997]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. 2009
</bibRefCitation>
). Morphologically
<taxonomicName id="04A7A4BFFF9EFF99FBC6FC03FA74FC57" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[1124,1424,972,997]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FBC6FC03FB64FC57" box="[1124,1152,972,997]" italics="true" pageId="35" pageNumber="236">D.</emphasis>
aff.
<emphasis id="F1D3032EFF9EFF99FB14FC03FA74FC57" box="[1206,1424,972,997]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
is almost identical to
<taxonomicName id="04A7A4BFFF9EFF99FEF8FC3FFDB1FBBB" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[346,597,1008,1033]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FEF8FC3FFE8AFBBB" box="[346,366,1008,1033]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FED9FC3FFDB1FBBB" box="[379,597,1008,1033]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
, but differs in size range (length: 16.029.0 μm
<emphasis id="F1D3032EFF9EFF99FBCBFC3FFB64FBBB" box="[1129,1152,1008,1033]" italics="true" pageId="35" pageNumber="236">vs</emphasis>
. 22.043.0 μm; wide: 8.5 11.5 μm
<emphasis id="F1D3032EFF9EFF99FF4FFBDBFEE0FB9F" box="[237,260,1044,1069]" italics="true" pageId="35" pageNumber="236">vs</emphasis>
. 10.015.0 μm) and in the shape of the axial area (narrow-lanceolate
<emphasis id="F1D3032EFF9EFF99FBB6FBDBFBCFFB9F" box="[1044,1067,1044,1069]" italics="true" pageId="35" pageNumber="236">vs</emphasis>
. wide to narrow-lanceolate). Stria density is an additional character that separates
<taxonomicName id="04A7A4BFFF9EFF99FD30FBF7FC69FBE3" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[658,909,1080,1105]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FD30FBF7FD42FBE3" box="[658,678,1080,1105]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FD11FBF7FC69FBE3" box="[691,909,1080,1105]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
[“89 (not 10)”] from
<taxonomicName id="04A7A4BFFF9EFF99FB24FBF7FA4FFBE3" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[1158,1451,1080,1105]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FB24FBF7FB7EFBE3" box="[1158,1178,1080,1105]" italics="true" pageId="35" pageNumber="236">D</emphasis>
. aff.
<emphasis id="F1D3032EFF9EFF99FB73FBF7FA4FFBE3" box="[1233,1451,1080,1105]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
(
<quantity id="045F72D9FF9EFF99FF32FB93FF01FBC7" box="[144,229,1116,1141]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="35" pageNumber="236" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">911 in</quantity>
10 μm).
</paragraph>
<paragraph id="C318DF3CFF9EFF99FF1FFB4FFD61F84B" blockId="35.[135,1452,936,2077]" pageId="35" pageNumber="236">
When describing
<taxonomicName id="04A7A4BFFF9EFF99FE24FB4FFC72FB2B" authority=", Metzeltin et al. (2009)" authorityName="Metzeltin" authorityYear="2009" box="[390,918,1152,1177]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FE24FB4FFE7EFB2B" box="[390,410,1152,1177]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FE08FB4FFD60FB2B" box="[426,644,1152,1177]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
,
<bibRefCitation id="A736A2CDFF9EFF99FD36FB4FFC72FB2B" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[660,918,1152,1177]" pageId="35" pageNumber="236" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9EFF99FCABFB4FFCDFFB2B" box="[777,827,1152,1177]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. (2009)
</bibRefCitation>
</taxonomicName>
illustrated two different morphotypes [narrow, lanceolate axial area (pl. 108, figs 13, 16)
<emphasis id="F1D3032EFF9EFF99FD31FB6BFD4EFB0F" box="[659,682,1188,1213]" italics="true" pageId="35" pageNumber="236">vs</emphasis>
. wide, lanceolate axial area (pl. 108, figs 415)], and designated the wide axial area form as the
<typeStatus id="1C1C619EFF9EFF99FE5CFB07FDCAFB53" box="[510,558,1224,1249]" pageId="35" pageNumber="236">type</typeStatus>
(pl. 108, fig. 4). However, when illustrating
<typeStatus id="1C1C619EFF9EFF99FB9DFB07FB8BFB53" box="[1087,1135,1224,1249]" pageId="35" pageNumber="236">type</typeStatus>
specimens they referred to both narrow and wide axial area specimens (pl. 108, figs 13, 6, 1012, 14). Due to this, it is rather difficult to characterize
<taxonomicName id="04A7A4BFFF9EFF99FEB7FADFFDF5FA9B" box="[277,529,1296,1321]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FEB7FADFFECDFA9B" box="[277,297,1296,1321]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FE95FADFFDF5FA9B" box="[311,529,1296,1321]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
, and an emended typification is in order to clarify the identity of the given morphs. Furthermore,
<bibRefCitation id="A736A2CDFF9EFF99FE80FAFBFDFAFAFF" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[290,542,1332,1357]" pageId="35" pageNumber="236" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9EFF99FE36FAFBFE21FAFF" box="[404,453,1332,1357]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. (2009)
</bibRefCitation>
were not consistent in designating the
<typeStatus id="1C1C619EFF9EFF99FC73FAFBFBE5FAFF" box="[977,1025,1332,1357]" pageId="35" pageNumber="236">type</typeStatus>
locality for
<taxonomicName id="04A7A4BFFF9EFF99FB28FAFBFA62FAFF" box="[1162,1414,1332,1357]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FB28FAFBFA62FAFF" box="[1162,1414,1332,1357]" italics="true" pageId="35" pageNumber="236">D. heteromorphiforma</emphasis>
</taxonomicName>
. In the description they gave Zyyn byrkh River as a
<typeStatus id="1C1C619EFF9EFF99FD16FA97FD00FAC3" box="[692,740,1368,1393]" pageId="35" pageNumber="236">type</typeStatus>
locality, while in the illustrations they point to Jargalant River (pl. 108, fig. 4). Additionally,
<bibRefCitation id="A736A2CDFF9EFF99FE77FAB3FD34FA27" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[469,720,1404,1429]" pageId="35" pageNumber="236" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9EFF99FDE5FAB3FD9CFA27" box="[583,632,1404,1429]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. (2009)
</bibRefCitation>
argue that the population from Zyyn byrkh River is characterized with morphological variation within the axial area, explaining the variation as possible for abnormal clones or maybe a separate species. Such morphological variations are observed neither in Lake
<collectingRegion id="016311DEFF9EFF99FBFEFA0BFB5FFA6F" box="[1116,1211,1476,1501]" country="Mongolia" name="Hovsgol" pageId="35" pageNumber="236">Hövsgöl</collectingRegion>
nor in the
<collectingRegion id="016311DEFF9EFF99FA97FA0BFF0AF9B3" country="Mongolia" name="Arhangay" pageId="35" pageNumber="236">Arkhangai province</collectingRegion>
. Furthermore, in the description
<bibRefCitation id="A736A2CDFF9EFF99FDC7FA27FC82F9B3" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[613,870,1512,1537]" pageId="35" pageNumber="236" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9EFF99FD7BFA27FCE8F9B3" box="[729,780,1512,1537]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. (2009)
</bibRefCitation>
compared
<taxonomicName id="04A7A4BFFF9EFF99FC44FA27FB00F9B3" box="[998,1252,1512,1537]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FC44FA27FC1EF9B3" box="[998,1018,1512,1537]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FBA8FA27FB00F9B3" box="[1034,1252,1512,1537]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
to a non-existent
<taxonomicName id="04A7A4BFFF9EFF99FF2AF9C3FE48F997" authority="Hustedt" authorityName="Hustedt" box="[136,428,1548,1573]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorpha">
<emphasis id="F1D3032EFF9EFF99FF2AF9C3FEAEF997" box="[136,330,1548,1573]" italics="true" pageId="35" pageNumber="236">D. heteromorpha</emphasis>
Hustedt
</taxonomicName>
, when they should have referred to
<taxonomicName id="04A7A4BFFF9EFF99FCE5F9C3FB33F997" authority="Hustedt (1937" authorityName="Hustedt" authorityYear="1937" box="[839,1239,1548,1573]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FCE5F9C3FBCDF997" box="[839,1065,1548,1573]" italics="true" pageId="35" pageNumber="236">Diploneis dimorpha</emphasis>
<bibRefCitation id="A736A2CDFF9EFF99FB90F9C3FB33F997" author="Hustedt, F." box="[1074,1239,1548,1573]" pageId="35" pageNumber="236" pagination="577 - 736" refId="ref26709" refString="Hustedt, F. (1937) Die Kieselalgen Deutschlands, Osterreichs und der Schweiz unter Berucksichtigung der ubrigen Lander Europas sowie der angrenzenden Meeresgebiete. In: Rabenhorst, L. (Ed.) Kryptogamen Flora von Deutschland, Osterreich und der Schweiz. Akademische Verlagsgesellschaft m. b. h. Leipzig 7 (Teil 2, Lief. 5), pp. 577 - 736." type="journal article" year="1937">Hustedt (1937</bibRefCitation>
</taxonomicName>
: pl. 1, fig. 1819). Keeping in mind that
<taxonomicName id="04A7A4BFFF9EFF99FED8F9FFFDECF9FB" authorityName="Hustedt" authorityYear="1937" box="[378,520,1584,1609]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FED8F9FFFDECF9FB" box="[378,520,1584,1609]" italics="true" pageId="35" pageNumber="236">D. dimorpha</emphasis>
</taxonomicName>
is characterized by heterovalvar frustules and has a distribution restricted to marine and brackish environments, it is rather unlikely that
<taxonomicName id="04A7A4BFFF9EFF99FD47F99BFC93F9DF" authorityName="Hustedt" authorityYear="1937" box="[741,887,1620,1645]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FD47F99BFD1DF9DF" box="[741,761,1620,1645]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FCA8F99BFC93F9DF" box="[778,887,1620,1645]" italics="true" pageId="35" pageNumber="236">dimorpha</emphasis>
</taxonomicName>
should be compared with
<taxonomicName id="04A7A4BFFF9EFF99FB0EF99BFA4FF9DF" box="[1196,1451,1620,1645]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FB0EF99BFB24F9DF" box="[1196,1216,1620,1645]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FB73F99BFA4FF9DF" box="[1233,1451,1620,1645]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
and
<taxonomicName id="04A7A4BFFF9EFF99FF1AF9B7FE05F923" box="[184,481,1656,1681]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FF1AF9B7FF28F923" box="[184,204,1656,1681]" italics="true" pageId="35" pageNumber="236">D</emphasis>
. aff.
<emphasis id="F1D3032EFF9EFF99FEA5F9B7FE05F923" box="[263,481,1656,1681]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
(
<bibRefCitation id="A736A2CDFF9EFF99FE52F9B7FD6FF923" author="Hustedt, F." box="[496,651,1656,1681]" pageId="35" pageNumber="236" pagination="577 - 736" refId="ref26709" refString="Hustedt, F. (1937) Die Kieselalgen Deutschlands, Osterreichs und der Schweiz unter Berucksichtigung der ubrigen Lander Europas sowie der angrenzenden Meeresgebiete. In: Rabenhorst, L. (Ed.) Kryptogamen Flora von Deutschland, Osterreich und der Schweiz. Akademische Verlagsgesellschaft m. b. h. Leipzig 7 (Teil 2, Lief. 5), pp. 577 - 736." type="journal article" year="1937">Hustedt 1937</bibRefCitation>
,
<bibRefCitation id="A736A2CDFF9EFF99FD34F9B7FC75F923" author="Witkowski, A. &amp; Lange-Bertalot, H. &amp; Metzeltin, D." box="[662,913,1656,1681]" pageId="35" pageNumber="236" pagination="1 - 925" refId="ref28750" refString="Witkowski, A., Lange-Bertalot, H. &amp; Metzeltin, D. (2000) Diatom Flora of Marine Coasts I. Iconographia Diatomologica 7: 1 - 925" type="journal article" year="2000">
Witkowski
<emphasis id="F1D3032EFF9EFF99FCB5F9B7FCACF923" box="[791,840,1656,1681]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. 2000
</bibRefCitation>
,
<bibRefCitation id="A736A2CDFF9EFF99FC3EF9B7FB80F923" author="Witon, E. &amp; Malmgren, B. &amp; Witkowski, A. &amp; Kuijpers, A." box="[924,1124,1656,1681]" pageId="35" pageNumber="236" pagination="487 - 509" refId="ref28781" refString="Witon, E., Malmgren, B., Witkowski, A. &amp; Kuijpers, A. (2006) Holocene marine diatoms from the Faeroe Islands and their paleoceanographic implications. Palaeogeography, Palaeoclimatology, Palaeoecology 239: 487 - 509 http: // dx. doi. org / 10.1016 / j. palaeo. 2006.02.006" type="journal article" year="2006">
Witon
<emphasis id="F1D3032EFF9EFF99FC4AF9B7FBFDF923" box="[1000,1049,1656,1681]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. 2006
</bibRefCitation>
). The heterovalvar frustule is nicely visible in Simonsens illustrations (1987, pl. 680, figs 1214). In order to support the comparisons and therefore to observe the heterovalvar feature, the Hustedt
<typeStatus id="1C1C619EFF9EFF99FD0BF90FFD3DF96B" box="[681,729,1728,1753]" pageId="35" pageNumber="236">type</typeStatus>
material of
<taxonomicName id="04A7A4BFFF9EFF99FCC4F90FFC12F96B" authorityName="Hustedt" authorityYear="1937" box="[870,1014,1728,1753]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FCC4F90FFC12F96B" box="[870,1014,1728,1753]" italics="true" pageId="35" pageNumber="236">D. dimorpha</emphasis>
</taxonomicName>
(O4/78) was analysed and included in this study (Figs 211, 212). According to the comparisons, no Mongolian population seems to be allied with the
<typeStatus id="1C1C619EFF9EFF99FAD9F92BFA4FF94F" box="[1403,1451,1764,1789]" pageId="35" pageNumber="236">type</typeStatus>
of
<taxonomicName id="04A7A4BFFF9EFF99FF0BF8C7FEDEF893" authorityName="Hustedt" authorityYear="1937" box="[169,314,1800,1825]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FF0BF8C7FEDEF893" box="[169,314,1800,1825]" italics="true" pageId="35" pageNumber="236">D. dimorpha</emphasis>
</taxonomicName>
. When describing
<taxonomicName id="04A7A4BFFF9EFF99FDB1F8C7FBC1F893" authority=", Metzeltin et al. (2009)" authorityName="Metzeltin" authorityYear="2009" box="[531,1061,1800,1825]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FDB1F8C7FDC3F893" box="[531,551,1800,1825]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FD9AF8C7FCFDF893" box="[568,793,1800,1825]" italics="true" pageId="35" pageNumber="236">heteromorphiforma,</emphasis>
<bibRefCitation id="A736A2CDFF9EFF99FC80F8C7FBC1F893" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[802,1061,1800,1825]" pageId="35" pageNumber="236" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9EFF99FC35F8C7FC2EF893" box="[919,970,1800,1825]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. (2009)
</bibRefCitation>
</taxonomicName>
mentioned Genkals report for
<taxonomicName id="04A7A4BFFF9EFF99FA32F8C7FF11F8F7" authorityName="Hustedt" authorityYear="1937" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FA32F8C7FF11F8F7" italics="true" pageId="35" pageNumber="236">D. dimorpha</emphasis>
</taxonomicName>
(
<bibRefCitation id="A736A2CDFF9EFF99FEA1F8E3FD79F8F7" author="Lange-Bertalot, H. &amp; Genkal, S. I." box="[259,669,1836,1861]" pageId="35" pageNumber="236" pagination="1 - 271" refId="ref27624" refString="Lange-Bertalot, H. &amp; Genkal, S. I. (1999) Diatomeen aus Sibirien I. Inseln im Arktischen Ozean (Yugorsky-Shar Strait). Iconographia Diatomologica 6: 1 - 271." type="journal article" year="1999">Lange-Bertalot &amp; Genkal 1999: 194</bibRefCitation>
, pl. 43, figs 12, 13), characterized with frustules that have isomorphic valves and suggested their separation into a new species. According to the comparisons,
<taxonomicName id="04A7A4BFFF9EFF99FBDDF89FFA4FF8DB" box="[1151,1451,1872,1897]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FBDDF89FFB7EF8DB" box="[1151,1178,1872,1897]" italics="true" pageId="35" pageNumber="236">D.</emphasis>
aff.
<emphasis id="F1D3032EFF9EFF99FB73F89FFA4FF8DB" box="[1233,1451,1872,1897]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
appears to be different from both
<taxonomicName id="04A7A4BFFF9EFF99FDB0F8BBFCF5F83F" box="[530,785,1908,1933]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FDB0F8BBFCF5F83F" box="[530,785,1908,1933]" italics="true" pageId="35" pageNumber="236">D. heteromorphiforma</emphasis>
</taxonomicName>
morphotypes (
<bibRefCitation id="A736A2CDFF9EFF99FC1DF8BBFB51F83F" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[959,1205,1908,1933]" pageId="35" pageNumber="236" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFF9EFF99FB94F8BBFB8EF83F" box="[1078,1130,1908,1933]" italics="true" pageId="35" pageNumber="236">et al</emphasis>
. 2009
</bibRefCitation>
) but closely allied to Genkals description of
<taxonomicName id="04A7A4BFFF9EFF99FE3FF857FC52F803" authority="(Lange-Bertalot &amp; Genkal 1999)" baseAuthorityName="Lange-Bertalot &amp; Genkal" baseAuthorityYear="1999" box="[413,950,1944,1969]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FE3FF857FDD4F803" box="[413,560,1944,1969]" italics="true" pageId="35" pageNumber="236">D. dimorpha</emphasis>
(
<bibRefCitation id="A736A2CDFF9EFF99FDE3F857FC49F803" author="Lange-Bertalot, H. &amp; Genkal, S. I." box="[577,941,1944,1969]" pageId="35" pageNumber="236" pagination="1 - 271" refId="ref27624" refString="Lange-Bertalot, H. &amp; Genkal, S. I. (1999) Diatomeen aus Sibirien I. Inseln im Arktischen Ozean (Yugorsky-Shar Strait). Iconographia Diatomologica 6: 1 - 271." type="journal article" year="1999">Lange-Bertalot &amp; Genkal 1999</bibRefCitation>
)
</taxonomicName>
. Further analysis is therefore preferable in order to clarify the morphological variation within the
<taxonomicName id="04A7A4BFFF9EFF99FD57F873FC17F867" box="[757,1011,1980,2005]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FD57F873FCEDF867" box="[757,777,1980,2005]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FCBBF873FC17F867" box="[793,1011,1980,2005]" italics="true" pageId="35" pageNumber="236">heteromorphiforma</emphasis>
</taxonomicName>
complex and possibly to ascertain any phylogenetic relationships with
<taxonomicName id="04A7A4BFFF9EFF99FE4DF82FFD9AF84B" baseAuthorityName="Lange-Bertalot &amp; Genkal" baseAuthorityYear="1999" box="[495,638,2016,2041]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="dimorpha">
<emphasis id="F1D3032EFF9EFF99FE4DF82FFD9AF84B" box="[495,638,2016,2041]" italics="true" pageId="35" pageNumber="236">D. dimorpha</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C318DF3CFF9EFF9EFF1FF7CBFCCBFEB3" blockId="35.[135,1452,936,2077]" lastBlockId="36.[136,1451,160,329]" lastPageId="36" lastPageNumber="237" pageId="35" pageNumber="236">
The original drawings (
<typeStatus id="1C1C619EFF9EFF99FE6DF7CBFDA5F7AF" box="[463,577,2052,2077]" pageId="35" pageNumber="236" type="iconotype">iconotype</typeStatus>
) of
<taxonomicName id="04A7A4BFFF9EFF99FDD4F7CBFC36F7AF" authority="Cleve-Euler (1953" authorityName="Cleve-Euler" authorityYear="1953" box="[630,978,2052,2077]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="robusta">
<emphasis id="F1D3032EFF9EFF99FDD4F7CBFD6EF7AF" box="[630,650,2052,2077]" italics="true" pageId="35" pageNumber="236">D</emphasis>
.
<emphasis id="F1D3032EFF9EFF99FD3EF7CBFD15F7AF" box="[668,753,2052,2077]" italics="true" pageId="35" pageNumber="236">robusta</emphasis>
<bibRefCitation id="A736A2CDFF9EFF99FD5EF7CBFC36F7AF" author="Cleve-Euler, A." box="[764,978,2052,2077]" pageId="35" pageNumber="236" pagination="1 - 255" refId="ref25541" refString="Cleve-Euler, A. (1953) Die Diatomeen von Schweden und Finnland. Teil III. Monoraphideae, Biraphideae 1. Kongiliga Svenska Vetenskaps-Akademiens Handlingar, Fjarde Serien 4: 1 - 255." type="journal article" year="1953">Cleve-Euler (1953</bibRefCitation>
</taxonomicName>
, fig 622 ac) are very similar to
<taxonomicName id="04A7A4BFFF9EFF9EFAC2F7CBFE86FF0B" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Chromista" lastPageId="36" lastPageNumber="237" order="Naviculales" pageId="35" pageNumber="236" phylum="Ochrophyta" rank="species" species="heteromorphiforma">
<emphasis id="F1D3032EFF9EFF99FAC2F7CBFA9FF7AF" box="[1376,1403,2052,2077]" italics="true" pageId="35" pageNumber="236">D.</emphasis>
aff.
<emphasis id="F1D3032EFF99FF9EFF2AFF6FFE86FF0B" box="[136,354,160,185]" italics="true" pageId="36" pageNumber="237">heteromorphiforma</emphasis>
</taxonomicName>
, whose numerical features unambiguously fit the measurements given in the original description of
<taxonomicName id="04A7A4BFFF99FF9EFF04FF0BFEF9FF6F" authorityName="Cleve-Euler" authorityYear="1953" box="[166,285,196,221]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="36" pageNumber="237" phylum="Ochrophyta" rank="species" species="robusta">
<emphasis id="F1D3032EFF99FF9EFF04FF0BFF5EFF6F" box="[166,186,196,221]" italics="true" pageId="36" pageNumber="237">D</emphasis>
.
<emphasis id="F1D3032EFF99FF9EFF6AFF0BFEF9FF6F" box="[200,285,196,221]" italics="true" pageId="36" pageNumber="237">robusta</emphasis>
</taxonomicName>
. However, additional analyses such as observation of
<taxonomicName id="04A7A4BFFF99FF9EFCD5FF0BFC09FF6F" authorityName="Cleve-Euler" authorityYear="1953" box="[887,1005,196,221]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="36" pageNumber="237" phylum="Ochrophyta" rank="species" species="robusta">
<emphasis id="F1D3032EFF99FF9EFCD5FF0BFC6FFF6F" box="[887,907,196,221]" italics="true" pageId="36" pageNumber="237">D</emphasis>
.
<emphasis id="F1D3032EFF99FF9EFC3AFF0BFC09FF6F" box="[920,1005,196,221]" italics="true" pageId="36" pageNumber="237">robusta</emphasis>
</taxonomicName>
type material are needed in order to test the similarity, or the possible conspecificity of these species.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF99FF9EFF1FFEC3FCEAFEFB" pageId="36" pageNumber="237" type="biology_ecology">
<paragraph id="C318DF3CFF99FF9EFF1FFEC3FCEAFEFB" blockId="36.[136,1451,160,329]" pageId="36" pageNumber="237">
<emphasis id="F1D3032EFF99FF9EFF1FFEC3FE1CFE97" bold="true" box="[189,504,268,293]" pageId="36" pageNumber="237">Ecology and Distribution:</emphasis>
—M068A; M166A;
<materialsCitation id="73CFD561FF99FF9EFD47FEC3FCEEFEFB" location="southern Lake" pageId="36" pageNumber="237" specimenCount="1" stateProvince="Hovsgol">
M167A: found in a shallow epilithic collection from
<location id="C67889E7FF99FF9EFAE9FEC3FF24FEFB" LSID="urn:lsid:plazi:treatment:4B0E6E2AFF9EFF9EFF2AFF51FCEAFEFB:C67889E7FF99FF9EFAE9FEC3FF24FEFB" name="southern Lake" pageId="36" pageNumber="237" stateProvince="Hovsgol">southern Lake</location>
<collectingRegion id="016311DEFF99FF9EFF65FEFFFEC2FEFB" box="[199,294,304,329]" country="Mongolia" name="Hovsgol" pageId="36" pageNumber="237">Hövsgöl</collectingRegion>
and also in streams of
<collectingRegion id="016311DEFF99FF9EFD85FEFFFCEEFEFB" box="[551,778,304,329]" country="Mongolia" name="Arhangay" pageId="36" pageNumber="237">Arkhangai province</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,283 @@
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<mods:title id="2B9CCA8297CA886067CF2EC2506C70D4">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<mods:namePart id="2CCB1C1F2704BC6040F6323CDC33DBCF">Jovanovska, Elena</mods:namePart>
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<treatment id="4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53" ID-DOI="http://doi.org/10.5281/zenodo.13634233" ID-Zenodo-Dep="13634233" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53" httpUri="http://treatment.plazi.org/id/4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53" lastPageNumber="230" pageId="29" pageNumber="230">
<subSubSection id="8BBD8CB7FFA0FFA7FF2AFF51FE4AFCCE" pageId="29" pageNumber="230" type="nomenclature">
<paragraph id="C318DF3CFFA0FFA7FF2AFF51FDB8FF0B" blockId="29.[136,604,158,185]" box="[136,604,158,185]" pageId="29" pageNumber="230">
<heading id="98506850FFA0FFA7FF2AFF51FDB8FF0B" box="[136,604,158,185]" fontSize="11" level="2" pageId="29" pageNumber="230" reason="3">
<taxonomicName id="04A7A4BFFFA0FFA7FF2AFF51FEA8FF0A" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[136,332,158,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="exigua" status="sp. nov.">Diploneis exigua</taxonomicName>
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<emphasis id="F1D3032EFFA0FFA7FEF0FF6FFE45FF0B" box="[338,417,160,185]" italics="true" pageId="29" pageNumber="230">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="5B9CC3B9FFA0FFA7FE15FF6FFDB7FF0B" box="[439,595,160,185]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 152165</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFFA0FFA7FF2AFF2AFE4AFCCE" blockId="29.[136,1452,229,892]" pageId="29" pageNumber="230">
Valves are elliptical with convex margins and round ends (
<figureCitation id="5B9CC3B9FFA0FFA7FD59FF2AFC77FF4E" box="[763,915,229,252]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 152160</figureCitation>
). The valve length is 14.517.0 μm, and the valve breadth is 9.010.5 μm. The axial area is narrow, linear, and expands into a small rectangular central area. From inside, a thin silica plate covers the whole length of the longitudinal canal (
<figureCitation id="5B9CC3B9FFA0FFA7FC82FEEAFC9AFE8E" box="[800,894,293,316]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 165</figureCitation>
). From outside, the central area is small and rectangular, ca. 1.5 μm wide. Externally, the longitudinal canal appears linear with one to two rows of small areolae in the middle of the valve continuing into one or two areolae covered with a large cribrum towards the valve apices (
<figureCitation id="5B9CC3B9FFA0FFA7FB7AFEAAFA8BFECE" box="[1240,1391,357,380]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 160, 161</figureCitation>
). The areolae of the longitudinal canal are covered with cribra morphologically identical to those on the striae, from which they are separated with a narrow hyaline area (
<figureCitation id="5B9CC3B9FFA0FFA7FDBAFE6AFD97FE0E" box="[536,627,421,444]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 161</figureCitation>
). In the middle of the valve the areolae of the canal are covered with small and simple cribra that become larger and irregularly polygonal towards the valve apices (
<figureCitation id="5B9CC3B9FFA0FFA7FB82FE0AFB9BFE6E" box="[1056,1151,453,476]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 160</figureCitation>
). Internally the longitudinal canal is closed with a thick linear silica plate forming a “depression” where the raphe is placed (
<figureCitation id="5B9CC3B9FFA0FFA7FB3FFE2AFB18FE4E" box="[1181,1276,485,508]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 165</figureCitation>
). Externally, the raphe is straight and simple with expanded drop-like proximal ends that are bent to the same side of the valve and positioned within an expanded depression (
<figureCitation id="5B9CC3B9FFA0FFA7FE7EFDEAFDDDFD8E" box="[476,569,549,572]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 160</figureCitation>
). Toward the central area the raphe branches drop into a “depression” below the rest of the non-porous silica, and are surrounded with a thick silica ridge (
<figureCitation id="5B9CC3B9FFA0FFA7FCCDFD8AFC29FDEE" box="[879,973,581,604]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 160</figureCitation>
). The depression is deepest and widest at the central area and gradually narrows toward the raphe distal ends. Distally, the raphe branches are bent into a short drop-like terminal fissures deflected to the same side of the valve (
<figureCitation id="5B9CC3B9FFA0FFA7FD7DFD4AFC97FD2E" box="[735,883,645,668]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 160, 161</figureCitation>
). Internally the raphe is straight with simple proximal ends, inserted in a slightly elevated sternum inside the “depression” formed by the longitudinal canal (
<figureCitation id="5B9CC3B9FFA0FFA7FB1CFD6AFAFFFD0E" box="[1214,1307,677,700]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 165</figureCitation>
). Distally, the raphe endings are slightly bent to the same side of the valve into very short terminal fissures (
<figureCitation id="5B9CC3B9FFA0FFA7FBDEFD0AFB3EFD6E" box="[1148,1242,709,732]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Fig. 163</figureCitation>
). Striae are parallel in the middle of the valve, becoming radiate at the distal ends of the valve,
<quantity id="045F72D9FFA0FFA7FC1CFD2AFBFAFD4E" box="[958,1054,741,764]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="29" pageNumber="230" unit="in" value="15.5" valueMax="16.0" valueMin="15.0">1516 in</quantity>
10 μm, and composed of round areolae,
<quantity id="045F72D9FFA0FFA7FF60FCCAFEC4FCAE" box="[194,288,773,796]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" metricValueMax="5.08" metricValueMin="4.064" pageId="29" pageNumber="230" unit="in" value="18.0" valueMax="20.0" valueMin="16.0">1620 in</quantity>
10 μm. Striae are uniseriate becoming biseriate and/or complex in structure towards the valve margins. From outside, the areolae are covered with cribrate occlusions, which increase in size towards the valve margins, becoming largest at the mantle (
<figureCitation id="5B9CC3B9FFA0FFA7FE86FC8AFE59FCEE" box="[292,445,837,860]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 160162</figureCitation>
). Internally the alveoli open via single continuous and elongate openings covered with a fine silica layer (
<figureCitation id="5B9CC3B9FFA0FFA7FEA8FCAAFE47FCCE" box="[266,419,869,892]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 162165</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA0FFA7FF2AFC61FBD4FB9D" pageId="29" pageNumber="230" type="materials_examined">
<paragraph id="C318DF3CFFA0FFA7FF2AFC61FBD4FB9D" blockId="29.[136,1452,942,1071]" pageId="29" pageNumber="230">
<emphasis id="F1D3032EFFA0FFA7FF2AFC61FF3EFC71" bold="true" box="[136,218,942,963]" pageId="29" pageNumber="230">Type:—</emphasis>
<materialsCitation id="73CFD561FFA0FFA7FF78FC61FA43FC71" box="[218,1447,942,963]" country="Mongolia" county="Lake" latitude="51.454468" location="Station" municipality="National Park" pageId="29" pageNumber="230" specimenCount="1" stateProvince="Hovsgol">
<collectingCountry id="BBB09FACFFA0FFA7FF78FC61FEBBFC71" box="[218,351,942,963]" name="Mongolia" pageId="29" pageNumber="230">MONGOLIA</collectingCountry>
,
<collectingCounty id="2A79A7B0FFA0FFA7FECEFC61FE78FC71" box="[364,412,942,963]" pageId="29" pageNumber="230">Lake</collectingCounty>
<collectingRegion id="016311DEFFA0FFA7FE04FC61FE13FC71" box="[422,503,942,963]" country="Mongolia" name="Hovsgol" pageId="29" pageNumber="230">Hövsgöl</collectingRegion>
(
<collectingRegion id="016311DEFFA0FFA7FDA5FC61FDBEFC71" box="[519,602,942,963]" country="Mongolia" name="Hovsgol" pageId="29" pageNumber="230">Hövsgöl</collectingRegion>
<collectingMunicipality id="237C4546FFA0FFA7FDC6FC61FD14FC71" box="[612,752,942,963]" pageId="29" pageNumber="230">National Park</collectingMunicipality>
),
<location id="C67889E7FFA0FFA7FCA1FC61FCACFC71" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53:C67889E7FFA0FFA7FCA1FC61FCACFC71" box="[771,840,942,963]" country="Mongolia" county="Lake" latitude="51.454468" municipality="National Park" name="Station" pageId="29" pageNumber="230" stateProvince="Hovsgol">Station</location>
anchored off
<location id="C67889E7FFA0FFA7FC7FFC61FBDBFC71" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53:C67889E7FFA0FFA7FC7FFC61FBDBFC71" box="[989,1087,942,963]" country="Mongolia" county="Lake" latitude="51.454468" municipality="National Park" name="Hanh Gol" pageId="29" pageNumber="230" stateProvince="Hovsgol">Hanh Gol</location>
mouth.
<location id="C67889E7FFA0FFA7FB3AFC61FAF6FC71" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53:C67889E7FFA0FFA7FB3AFC61FAF6FC71" box="[1176,1298,942,963]" country="Mongolia" county="Lake" latitude="51.454468" municipality="National Park" name="Coordinates" pageId="29" pageNumber="230" stateProvince="Hovsgol">Coordinates</location>
:
<geoCoordinate id="A693B9FBFFA0FFA7FA82FC61FA43FC71" box="[1312,1447,942,963]" degrees="51" direction="north" minutes="27.268" orientation="latitude" pageId="29" pageNumber="230" precision="1" value="51.454468">51°27.268 N</geoCoordinate>
</materialsCitation>
<materialsCitation id="73CFD561FFA0FFA7FA05FC61FF0EFB9D" collectingDate="1998-07-19" collectionCode="ANSP" collectorName="A, M. B. Edlund Collection &amp; Mark B. Edlund &amp; Nergui Soninkhishig" country="Mongolia" elevation="4" location="Slide" longitude="100.7185" municipality="Science Museum of Minnesota" pageId="29" pageNumber="230" specimenCode="GC-36355" specimenCount="1" typeStatus="holotype">
;
<geoCoordinate id="A693B9FBFFA0FFA7FF1FFC1DFEDDFC55" box="[189,313,978,999]" degrees="100" direction="east" minutes="43.11" orientation="longitude" pageId="29" pageNumber="230" precision="9" value="100.7185">100°43.11E</geoCoordinate>
,
<taxonomicName id="04A7A4BFFFA0FFA7FEE1FC1DFE64FC55" box="[323,384,978,999]" class="Insecta" family="Noctuidae" genus="Chara" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="F1D3032EFFA0FFA7FEE1FC1DFE64FC55" box="[323,384,978,999]" italics="true" pageId="29" pageNumber="230">Chara</emphasis>
</taxonomicName>
from
<elevation id="488A380FFFA0FFA7FE1BFC1DFDF3FC55" box="[441,535,978,999]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="230" unit="m" value="4.0">
<quantity id="045F72D9FFA0FFA7FE1BFC1DFE3AFC55" box="[441,478,978,999]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="230" unit="m" value="4.0">4 m</quantity>
depth
</elevation>
(accession number: M272
<collectorName id="6E52BAEAFFA0FFA7FCB5FC1DFBFEFC55" box="[791,1050,978,999]" pageId="29" pageNumber="230">A, M.B. Edlund Collection</collectorName>
,
<collectingMunicipality id="237C4546FFA0FFA7FB80FC1DFAA8FC55" box="[1058,1356,978,999]" pageId="29" pageNumber="230">Science Museum of Minnesota</collectingMunicipality>
, collected by
<collectorName id="6E52BAEAFFA0FFA7FF79FC39FE92FBB9" box="[219,374,1014,1035]" pageId="29" pageNumber="230">Mark B. Edlund</collectorName>
and
<collectorName id="6E52BAEAFFA0FFA7FE04FC39FD8BFBB9" box="[422,623,1014,1035]" pageId="29" pageNumber="230">Nergui Soninkhishig</collectorName>
,
<date id="B719F9FCFFA0FFA7FDDAFC39FD12FBB9" box="[632,758,1014,1035]" pageId="29" pageNumber="230" value="1998-07-19">
<collectingDate id="A75D0014FFA0FFA7FDDAFC39FD12FBB9" box="[632,758,1014,1035]" pageId="29" pageNumber="230" value="1998-07-19">19 July 1998</collectingDate>
</date>
) (
<location id="C67889E7FFA0FFA7FCABFC39FCD8FBB9" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53:C67889E7FFA0FFA7FCABFC39FCD8FBB9" box="[777,828,1014,1035]" country="Mongolia" longitude="100.7185" municipality="Science Museum of Minnesota" name="Slide" pageId="29" pageNumber="230">Slide</location>
M272A,
<collectionCode id="A5B647F9FFA0FFA7FC35FC39FC37FBB9" box="[919,979,1014,1035]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="29" pageNumber="230">ANSP</collectionCode>
<specimenCode id="93017747FFA0FFA7FC78FC39FBA6FBB9" box="[986,1090,1014,1035]" collectionCode="GC-" pageId="29" pageNumber="230">GC-36355</specimenCode>
, GCM-24057),
<emphasis id="F1D3032EFFA0FFA7FB78FC39FF0EFB9D" bold="true" pageId="29" pageNumber="230">
<typeStatus id="1C1C619EFFA0FFA7FB78FC39FAD7FBB9" box="[1242,1331,1014,1035]" pageId="29" pageNumber="230" type="holotype">holotype</typeStatus>
, designated here
</emphasis>
</materialsCitation>
; example specimens on
<figureCitation id="5B9CC3B9FFA0FFA7FE74FBD5FDB9FB9D" box="[470,605,1050,1071]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 153, 156</figureCitation>
;
<materialsCitation id="73CFD561FFA0FFA7FDC4FBD5FBC8FB9D" box="[614,1068,1050,1071]" collectionCode="CAS" country="Mongolia" location="Slide" pageId="29" pageNumber="230" specimenCount="1" typeStatus="isotype">
(
<location id="C67889E7FFA0FFA7FDCFFBD5FD44FB9D" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA0FFA7FF2AFF51FBE9FA53:C67889E7FFA0FFA7FDCFFBD5FD44FB9D" box="[621,672,1050,1071]" country="Mongolia" name="Slide" pageId="29" pageNumber="230">Slide</location>
919072,
<collectionCode id="A5B647F9FFA0FFA7FD58FBD5FCCFFB9D" box="[762,811,1050,1071]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="230" type="Museum">CAS</collectionCode>
,
<emphasis id="F1D3032EFFA0FFA7FC96FBD5FBC7FB9D" bold="true" box="[820,1059,1050,1071]" pageId="29" pageNumber="230">
<typeStatus id="1C1C619EFFA0FFA7FC96FBD5FC98FB9D" box="[820,892,1050,1071]" pageId="29" pageNumber="230" type="isotype">isotype</typeStatus>
designated here
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA0FFA7FF2AFBAFFD5AFA2B" pageId="29" pageNumber="230" type="etymology">
<paragraph id="C318DF3CFFA0FFA7FF2AFBAFFCBCFBCB" blockId="29.[136,1452,1120,1505]" box="[136,856,1120,1145]" pageId="29" pageNumber="230">
<emphasis id="F1D3032EFFA0FFA7FF2AFBAFFECAFBCB" bold="true" box="[136,302,1120,1145]" pageId="29" pageNumber="230">Etymology:—</emphasis>
The species name refers to the taxons small size.
</paragraph>
<paragraph id="C318DF3CFFA0FFA7FF1FFB4BFD5AFA2B" blockId="29.[136,1452,1120,1505]" pageId="29" pageNumber="230">
<emphasis id="F1D3032EFFA0FFA7FF1FFB4BFE81FB2F" bold="true" box="[189,357,1156,1181]" pageId="29" pageNumber="230">Observations:</emphasis>
—With respect to the valve size,
<taxonomicName id="04A7A4BFFFA0FFA7FD7AFB4BFC72FB2F" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[728,918,1156,1181]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="exigua">
<emphasis id="F1D3032EFFA0FFA7FD7AFB4BFC72FB2F" box="[728,918,1156,1181]" italics="true" pageId="29" pageNumber="230">Diploneis exigua</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="04A7A4BFFFA0FFA7FBC2FB4BFACEFB2F" box="[1120,1322,1156,1181]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA0FFA7FBC2FB4BFB90FB2F" box="[1120,1140,1156,1181]" italics="true" pageId="29" pageNumber="230">D</emphasis>
.
<emphasis id="F1D3032EFFA0FFA7FB20FB4BFACEFB2F" box="[1154,1322,1156,1181]" italics="true" pageId="29" pageNumber="230">soninkhishigae</emphasis>
</taxonomicName>
, but differs in: valve shape (elliptical
<emphasis id="F1D3032EFFA0FFA7FE0EFB67FE27FB73" box="[428,451,1192,1217]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
. elliptical-lanceolate); central area (rectangular
<emphasis id="F1D3032EFFA0FFA7FC42FB67FC13FB73" box="[992,1015,1192,1217]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
. elongate); striae density (
<quantity id="045F72D9FFA0FFA7FA81FB67FA6CFB73" box="[1315,1416,1192,1217]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="29" pageNumber="230" unit="in" value="15.5" valueMax="16.0" valueMin="15.0">1516 in</quantity>
10 μm
<emphasis id="F1D3032EFFA0FFA7FF15FB03FF32FB57" box="[183,214,1228,1253]" italics="true" pageId="29" pageNumber="230">vs.</emphasis>
<quantity id="045F72D9FFA0FFA7FF7DFB03FEA7FB57" box="[223,323,1228,1253]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.81" metricValueMin="2.794" pageId="29" pageNumber="230" unit="in" value="13.0" valueMax="15.0" valueMin="11.0">1115 in</quantity>
10 μm); and areola density (
<quantity id="045F72D9FFA0FFA7FD30FB03FD1DFB57" box="[658,761,1228,1253]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" metricValueMax="5.08" metricValueMin="4.064" pageId="29" pageNumber="230" unit="in" value="18.0" valueMax="20.0" valueMin="16.0">1620 in</quantity>
10 μm
<emphasis id="F1D3032EFFA0FFA7FCF4FB03FC89FB57" box="[854,877,1228,1253]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
.
<quantity id="045F72D9FFA0FFA7FCDFFB03FC06FB57" box="[893,994,1228,1253]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="29" pageNumber="230" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">1012 in</quantity>
10 μm).
<taxonomicName id="04A7A4BFFFA0FFA7FBF2FB03FAF5FB57" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[1104,1297,1228,1253]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="exigua">
<emphasis id="F1D3032EFFA0FFA7FBF2FB03FAF5FB57" box="[1104,1297,1228,1253]" italics="true" pageId="29" pageNumber="230">Diploneis exigua</emphasis>
</taxonomicName>
has a similar central area as
<taxonomicName id="04A7A4BFFFA0FFA7FE8CFB3FFE5FFABB" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[302,443,1264,1289]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFA0FFA7FE8CFB3FFE5FFABB" box="[302,443,1264,1289]" italics="true" pageId="29" pageNumber="230">D. stoermeri</emphasis>
</taxonomicName>
morphotype 1 (compare
<figureCitation id="5B9CC3B9FFA0FFA7FD77FB3FFC8AFABB" box="[725,878,1264,1289]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="29" pageNumber="230">Figs 152159</figureCitation>
with
<figureCitation id="5B9CC3B9FFA0FFA7FC0FFB3FFBCBFABB" box="[941,1071,1264,1289]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="29" pageNumber="230">Figs 8089</figureCitation>
). However, the size range (length: 14.517.0 μm in
<taxonomicName id="04A7A4BFFFA0FFA7FEEBFADBFE52FA9F" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[329,438,1300,1325]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="exigua">
<emphasis id="F1D3032EFFA0FFA7FEEBFADBFEB9FA9F" box="[329,349,1300,1325]" italics="true" pageId="29" pageNumber="230">D</emphasis>
.
<emphasis id="F1D3032EFFA0FFA7FEC9FADBFE52FA9F" box="[363,438,1300,1325]" italics="true" pageId="29" pageNumber="230">exigua</emphasis>
</taxonomicName>
<emphasis id="F1D3032EFFA0FFA7FE1EFADBFE37FA9F" box="[444,467,1300,1325]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
. 19.544.5 μm
<taxonomicName id="04A7A4BFFFA0FFA7FD27FADBFCF6FA9F" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[645,786,1300,1325]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFA0FFA7FD27FADBFD7DFA9F" box="[645,665,1300,1325]" italics="true" pageId="29" pageNumber="230">D</emphasis>
.
<emphasis id="F1D3032EFFA0FFA7FD05FADBFCF6FA9F" box="[679,786,1300,1325]" italics="true" pageId="29" pageNumber="230">stoermeri</emphasis>
</taxonomicName>
; breadth: 9.010.5 μm in
<taxonomicName id="04A7A4BFFFA0FFA7FB95FADBFB40FA9F" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[1079,1188,1300,1325]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="exigua">
<emphasis id="F1D3032EFFA0FFA7FB95FADBFBAFFA9F" box="[1079,1099,1300,1325]" italics="true" pageId="29" pageNumber="230">D</emphasis>
.
<emphasis id="F1D3032EFFA0FFA7FBFBFADBFB40FA9F" box="[1113,1188,1300,1325]" italics="true" pageId="29" pageNumber="230">exigua</emphasis>
</taxonomicName>
<emphasis id="F1D3032EFFA0FFA7FB08FADBFB25FA9F" box="[1194,1217,1300,1325]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
. 12.522.0 μm in
<taxonomicName id="04A7A4BFFFA0FFA7FA32FADBFF17FAE3" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFA0FFA7FA32FADBFA40FA9F" box="[1424,1444,1300,1325]" italics="true" pageId="29" pageNumber="230">D</emphasis>
.
<emphasis id="F1D3032EFFA0FFA7FF2AFAF7FF17FAE3" box="[136,243,1336,1361]" italics="true" pageId="29" pageNumber="230">stoermeri</emphasis>
</taxonomicName>
) separates these two species.
<taxonomicName id="04A7A4BFFFA0FFA7FDE0FAF7FCD0FAE3" box="[578,820,1336,1361]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFA0FFA7FDE0FAF7FCD0FAE3" box="[578,820,1336,1361]" italics="true" pageId="29" pageNumber="230">Diploneis paraparma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="04A7A4BFFFA0FFA7FC6AFAF7FBD1FAE3" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[968,1077,1336,1361]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="29" pageNumber="230" phylum="Ochrophyta" rank="species" species="exigua">
<emphasis id="F1D3032EFFA0FFA7FC6AFAF7FC38FAE3" box="[968,988,1336,1361]" italics="true" pageId="29" pageNumber="230">D</emphasis>
.
<emphasis id="F1D3032EFFA0FFA7FC48FAF7FBD1FAE3" box="[1002,1077,1336,1361]" italics="true" pageId="29" pageNumber="230">exigua</emphasis>
</taxonomicName>
by the greater size range (length: 19.542.0 μm
<emphasis id="F1D3032EFFA0FFA7FE88FA93FEA5FAC7" box="[298,321,1372,1397]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
. 14.517.0 μm; breadth: 12.516.5 μm
<emphasis id="F1D3032EFFA0FFA7FD5FFA93FCF0FAC7" box="[765,788,1372,1397]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
. 9.010.5 μm) and the striae structure (uniseriate becoming biseriate towards the valve margins
<emphasis id="F1D3032EFFA0FFA7FDB9FA4FFDD6FA2B" box="[539,562,1408,1433]" italics="true" pageId="29" pageNumber="230">vs</emphasis>
. uniseriate).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA0FFA7FF1FFA6BFBE9FA53" pageId="29" pageNumber="230" type="biology_ecology">
<paragraph id="C318DF3CFFA0FFA7FF1FFA6BFBE9FA53" blockId="29.[136,1452,1120,1505]" pageId="29" pageNumber="230">
<emphasis id="F1D3032EFFA0FFA7FF1FFA6BFE17FA0F" bold="true" box="[189,499,1444,1469]" pageId="29" pageNumber="230">Ecology and Distribution:</emphasis>
—M248A; M272A; M273A; M274A; M291A: found in depths from
<quantity id="045F72D9FFA0FFA7FB5EFA6BFA82FA0F" box="[1276,1382,1444,1469]" metricMagnitude="1" metricUnit="m" metricValue="2.2" metricValueMax="4.0" metricValueMin="0.4" pageId="29" pageNumber="230" unit="m" value="22.0" valueMax="40.0" valueMin="4.0">4 to 40 m</quantity>
in the central and northern areas of Lake
<collectingRegion id="016311DEFFA0FFA7FDACFA07FD89FA53" box="[526,621,1480,1505]" country="Mongolia" name="Hovsgol" pageId="29" pageNumber="230">Hövsgöl</collectingRegion>
in sediment and epiphytic on
<taxonomicName id="04A7A4BFFFA0FFA7FC1DFA07FBE3FA53" box="[959,1031,1480,1505]" class="Insecta" family="Noctuidae" genus="Chara" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="F1D3032EFFA0FFA7FC1DFA07FBE3FA53" box="[959,1031,1480,1505]" italics="true" pageId="29" pageNumber="230">Chara</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,216 @@
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<mods:title id="871283F4A1EF83F342D16C80961FE6D1">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<heading id="98506850FFA2FFA5FF2AF995FC27F9C7" box="[136,963,1626,1653]" fontSize="11" level="2" pageId="31" pageNumber="232" reason="3">
<taxonomicName id="04A7A4BFFFA2FFA5FF2AF995FCF4F9C7" authority="Metzeltin, Lange-Bertalot &amp; Nergui" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[136,784,1626,1653]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="31" pageNumber="232" phylum="Ochrophyta" rank="species" species="spectabilis">
<emphasis id="F1D3032EFFA2FFA5FF2AF995FE92F9C7" bold="true" box="[136,374,1626,1653]" italics="true" pageId="31" pageNumber="232">Diploneis spectabilis</emphasis>
Metzeltin, Lange-Bertalot &amp; Nergui
</taxonomicName>
(
<figureCitation id="5B9CC3B9FFA2FFA5FCBDF993FC5FF9C7" box="[799,955,1628,1653]" captionStart="FIGURES 166183" captionStartId="30.[136,243,2013,2034]" captionTargetBox="[182,1405,608,1983]" captionTargetId="figure-236@30.[178,1409,605,1988]" captionTargetPageId="30" captionText="FIGURES 166183. LM valve views. Figs 166169. Diploneis elliptica cf. var. ladogensis. Figs 170172. Diploneis pseudovalis. Figs 173178. Diploneis subovalis. Fig. 179. Diploneis mongolica. Figs 180183. Diploneis spectabilis. Scale bar = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13634172" httpUri="https://zenodo.org/record/13634172/files/figure.png" pageId="31" pageNumber="232">Figs 180183</figureCitation>
)
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</paragraph>
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Valves are elliptical with convex margins and broadly round apices becoming bluntly rounded with the valve size reduction. The valve length is 28.538.0 μm, and the valve breadth is 13.018.0 μm. The axial area is very narrow and linear, expanding towards the large almost circular to rectangular central area. The central area is 4.55.0 μm wide. The longitudinal canal is narrow, linear with a single row of areolae throughout the whole length. The central area is broadly expanded. The raphe is straight and simples, positioned with an expanded depression. Striae are uniseriate and radiate,
<quantity id="045F72D9FFA2FFA5FB37F8D2FB2FF886" box="[1173,1227,1821,1844]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="31" pageNumber="232" unit="in" value="18.0">18 in</quantity>
10 μm, composed of round areolae,
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10 μm.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA2FF9AFF2AF84BFB66FE4F" lastPageId="32" lastPageNumber="233" pageId="31" pageNumber="232" type="description">
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<emphasis id="F1D3032EFFA2FFA5FF2AF84BFED4F82F" bold="true" box="[136,304,1924,1949]" pageId="31" pageNumber="232">Observations:</emphasis>
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<emphasis id="F1D3032EFFA2FFA5FEEEF84BFDA5F82F" box="[332,577,1924,1949]" italics="true" pageId="31" pageNumber="232">Diploneis spectabilis</emphasis>
</taxonomicName>
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was described from
<collectingMunicipality id="237C4546FFA2FFA5FCF2F84BFC1FF82F" box="[848,1019,1924,1949]" pageId="31" pageNumber="232">Lake Khangal</collectingMunicipality>
,
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(
<bibRefCitation id="A736A2CDFFA2FFA5FB36F84BFA44F82F" author="Metzeltin, D. &amp; Lange-Bertalot, H. &amp; Nergui, S." box="[1172,1440,1924,1949]" pageId="31" pageNumber="232" pagination="3 - 686" refId="ref27855" refString="Metzeltin, D., Lange-Bertalot, H. &amp; Nergui, S. (2009) Diatom in Mongolia. Iconographia Diatomologica 20: 3 - 686." type="journal article" year="2009">
Metzeltin
<emphasis id="F1D3032EFFA2FFA5FAB3F84BFAA8F82F" box="[1297,1356,1924,1949]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
. 2009
</bibRefCitation>
).
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on the
<typeStatus id="1C1C619EFFA2FFA5FECDF867FE7BF873" box="[367,415,1960,1985]" pageId="31" pageNumber="232">type</typeStatus>
material were performed in order to ascertain the identity of
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population.
<collectorName id="6E52BAEAFFA2FFA5FAEEF867FF56F857" pageId="31" pageNumber="232">Detailed LM</collectorName>
comparisons of the
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population and
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population revealed morphological differences in: valve size (length: 36.037.0 μm
<emphasis id="F1D3032EFFA2FFA5FE28F83FFE45F7BB" box="[394,417,2032,2057]" italics="true" pageId="31" pageNumber="232">vs</emphasis>
. 28.538.0 μm; breadth: 19.031.0 μm
<emphasis id="F1D3032EFFA2FFA5FCC3F83FFC9CF7BB" box="[865,888,2032,2057]" italics="true" pageId="31" pageNumber="232">vs</emphasis>
. 13.018.0 μm)
</materialsCitation>
; stria density (
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10 μm
<emphasis id="F1D3032EFFA2FFA5FA2FF83FFA40F7BB" box="[1421,1444,2032,2057]" italics="true" pageId="31" pageNumber="232">vs</emphasis>
.
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10 μm);
<materialsCitation id="73CFD561FFA2FF9AFE81F7DBFD97FE97" country="Mongolia" lastPageId="32" lastPageNumber="233" location="Lake Khangal" pageId="31" pageNumber="232" specimenCount="1" stateProvince="Arhangay">
and areola density (
<quantity id="045F72D9FFA2FFA5FE59F7DBFDB9F79F" box="[507,605,2068,2093]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="31" pageNumber="232" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">1618 in</quantity>
10 μm
<emphasis id="F1D3032EFFA2FFA5FD12F7DBFD23F79F" box="[688,711,2068,2093]" italics="true" pageId="31" pageNumber="232">vs</emphasis>
.
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10 μm). In
<collectingRegion id="016311DEFFA2FFA5FC28F7DBFBE4F79F" box="[906,1024,2068,2093]" country="Mongolia" name="Arhangay" pageId="31" pageNumber="232">Arkhangai</collectingRegion>
materials
<taxonomicName id="04A7A4BFFFA2FFA5FBD0F7DBFAEFF79F" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[1138,1291,2068,2093]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="31" pageNumber="232" phylum="Ochrophyta" rank="species" species="spectabilis">
<emphasis id="F1D3032EFFA2FFA5FBD0F7DBFAEFF79F" box="[1138,1291,2068,2093]" italics="true" pageId="31" pageNumber="232">D. spectabilis</emphasis>
</taxonomicName>
was very rare, while in
<location id="C67889E7FF9DFF9AFF4EFF6FFE6FFF0B" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA2FF9AFF2AF995FB03FDF7:C67889E7FF9DFF9AFF4EFF6FFE6FFF0B" box="[236,395,160,185]" country="Mongolia" name="Lake Khangal" pageId="32" pageNumber="233" stateProvince="Arhangay">Lake Khangal</location>
was quite frequent. The differences in size dimensions might be a result of the small number of specimens in
<collectingRegion id="016311DEFF9DFF9AFE99FF0BFE55FF6F" box="[315,433,196,221]" country="Mongolia" name="Arhangay" pageId="32" pageNumber="233">Arkhangai</collectingRegion>
material or ecophenotypic variation. However, in most specimens (there is some variability in valve end shape; compare
<figureCitation id="5B9CC3B9FF9DFF9AFE6CFF27FDCFFEB3" box="[462,555,232,257]" captionStart="FIGURES 166183" captionStartId="30.[136,243,2013,2034]" captionTargetBox="[182,1405,608,1983]" captionTargetId="figure-236@30.[178,1409,605,1988]" captionTargetPageId="30" captionText="FIGURES 166183. LM valve views. Figs 166169. Diploneis elliptica cf. var. ladogensis. Figs 170172. Diploneis pseudovalis. Figs 173178. Diploneis subovalis. Fig. 179. Diploneis mongolica. Figs 180183. Diploneis spectabilis. Scale bar = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13634172" httpUri="https://zenodo.org/record/13634172/files/figure.png" pageId="32" pageNumber="233">Fig. 181</figureCitation>
with
<figureCitation id="5B9CC3B9FF9DFF9AFDC9FF27FCE1FEB3" box="[619,773,232,257]" captionStart="FIGURES 166183" captionStartId="30.[136,243,2013,2034]" captionTargetBox="[182,1405,608,1983]" captionTargetId="figure-236@30.[178,1409,605,1988]" captionTargetPageId="30" captionText="FIGURES 166183. LM valve views. Figs 166169. Diploneis elliptica cf. var. ladogensis. Figs 170172. Diploneis pseudovalis. Figs 173178. Diploneis subovalis. Fig. 179. Diploneis mongolica. Figs 180183. Diploneis spectabilis. Scale bar = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13634172" httpUri="https://zenodo.org/record/13634172/files/figure.png" pageId="32" pageNumber="233">Figs 180, 183</figureCitation>
) the valve outline in Arkhangais population fits the original description given by
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Metzeltin
<emphasis id="F1D3032EFF9DFF9AFE44FEC3FDF3FE97" box="[486,535,268,293]" italics="true" pageId="32" pageNumber="233">et al</emphasis>
. (2009)
</bibRefCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="C318DF3CFF9DFF9AFF1FFEFFFB66FE4F" blockId="32.[136,1452,160,581]" pageId="32" pageNumber="233">
The differences between
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<emphasis id="F1D3032EFF9DFF9AFE7BFEFFFE09FEFB" box="[473,493,304,329]" italics="true" pageId="32" pageNumber="233">D</emphasis>
.
<emphasis id="F1D3032EFF9DFF9AFE59FEFFFD95FEFB" box="[507,625,304,329]" italics="true" pageId="32" pageNumber="233">mongolica</emphasis>
</taxonomicName>
and
<taxonomicName id="04A7A4BFFF9DFF9AFD0BFEFFFCA0FEFB" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[681,836,304,329]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="spectabilis">
<emphasis id="F1D3032EFF9DFF9AFD0BFEFFFD59FEFB" box="[681,701,304,329]" italics="true" pageId="32" pageNumber="233">D</emphasis>
.
<emphasis id="F1D3032EFF9DFF9AFD6EFEFFFCA0FEFB" box="[716,836,304,329]" italics="true" pageId="32" pageNumber="233">spectabilis</emphasis>
</taxonomicName>
(see
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Metzeltin
<emphasis id="F1D3032EFF9DFF9AFC51FEFFFBC1FEFB" box="[1011,1061,304,329]" italics="true" pageId="32" pageNumber="233">et al</emphasis>
. 2009
</bibRefCitation>
) are based on the size range (38.058.0 μm
<emphasis id="F1D3032EFF9DFF9AFE97FE9BFEA8FEDF" box="[309,332,340,365]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
. 36.072.0 μm) and stria density (
<quantity id="045F72D9FF9DFF9AFD6EFE9BFCA1FEDF" box="[716,837,340,365]" metricMagnitude="-1" metricUnit="m" metricValue="3.3655" metricValueMax="3.556" metricValueMin="3.175" pageId="32" pageNumber="233" unit="in" value="13.25" valueMax="14.0" valueMin="12.5">12.514 in</quantity>
10 μm
<emphasis id="F1D3032EFF9DFF9AFC39FE9BFC56FEDF" box="[923,946,340,365]" italics="true" pageId="32" pageNumber="233">vs</emphasis>
.
<quantity id="045F72D9FF9DFF9AFC62FE9BFBC6FEDF" box="[960,1058,340,365]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="32" pageNumber="233" unit="in" value="15.5" valueMax="16.0" valueMin="15.0">1516 in</quantity>
10 μm). Additionally, the structure of the longitudinal canal (one row of areolae throughout the whole length in
<emphasis id="F1D3032EFF9DFF9AFC5AFEB7FB57FE23" box="[1016,1203,376,401]" italics="true" pageId="32" pageNumber="233">
<taxonomicName id="04A7A4BFFF9DFF9AFC5AFEB7FB70FE23" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[1016,1172,376,401]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="spectabilis">D. spectabilis</taxonomicName>
vs
</emphasis>
. two to three rows of areolae in the mid-valve narrowing into two to one at the valve apices in
<taxonomicName id="04A7A4BFFF9DFF9AFC67FE53FBBAFE07" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[965,1118,412,437]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="mongolica">
<emphasis id="F1D3032EFF9DFF9AFC67FE53FBBAFE07" box="[965,1118,412,437]" italics="true" pageId="32" pageNumber="233">D. mongolica</emphasis>
</taxonomicName>
) also separates these species. However, further SEM analyses are necessary to better distinguish differences between
<taxonomicName id="04A7A4BFFF9DFF9AFBE8FE0FFB06FE6B" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[1098,1250,448,473]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="spectabilis">
<emphasis id="F1D3032EFF9DFF9AFBE8FE0FFB06FE6B" box="[1098,1250,448,473]" italics="true" pageId="32" pageNumber="233">D. spectabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="04A7A4BFFF9DFF9AFAB7FE0FFA4FFE6B" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[1301,1451,448,473]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="32" pageNumber="233" phylum="Ochrophyta" rank="species" species="mongolica">
<emphasis id="F1D3032EFF9DFF9AFAB7FE0FFA4FFE6B" box="[1301,1451,448,473]" italics="true" pageId="32" pageNumber="233">D. mongolica</emphasis>
</taxonomicName>
and within populations of each, and should be supported by additional morphological data.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FF9DFF9AFF1FFDC7FB03FDF7" pageId="32" pageNumber="233" type="biology_ecology">
<paragraph id="C318DF3CFF9DFF9AFF1FFDC7FB03FDF7" blockId="32.[136,1452,160,581]" pageId="32" pageNumber="233">
<emphasis id="F1D3032EFF9DFF9AFF1FFDC7FE11FD93" bold="true" box="[189,501,520,545]" pageId="32" pageNumber="233">Ecology and Distribution:</emphasis>
—M166A; M167A: examined for comparison, not found in Lake
<collectingRegion id="016311DEFF9DFF9AFB7DFDC7FADAFD93" box="[1247,1342,520,545]" country="Mongolia" name="Hovsgol" pageId="32" pageNumber="233">Hövsgöl</collectingRegion>
but more broadly distributed in lakes and streams in
<collectingRegion id="016311DEFF9DFF9AFDCAFDE3FC8DFDF7" box="[616,873,556,581]" isEnumeration="true" pageId="32" pageNumber="233">Arkhangai and Khentii</collectingRegion>
provinces (
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Metzeltin
<emphasis id="F1D3032EFF9DFF9AFBFDFDE3FB74FDF7" box="[1119,1168,556,581]" italics="true" pageId="32" pageNumber="233">et al</emphasis>
. 2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,137 @@
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<paragraph id="C318DF3CFFA2FFA5FF2AFBFDFC6DFBFF" blockId="31.[136,905,1074,1101]" box="[136,905,1074,1101]" pageId="31" pageNumber="232">
<heading id="98506850FFA2FFA5FF2AFBFDFC6DFBFF" box="[136,905,1074,1101]" fontSize="11" level="2" pageId="31" pageNumber="232" reason="3">
<taxonomicName id="04A7A4BFFFA2FFA5FF2AFBFDFCF6FBFF" authority="Metzeltin, Lange-Bertalot &amp; Nergui" authorityName="Metzeltin, Lange-Bertalot &amp; Nergui" authorityYear="2009" box="[136,786,1074,1101]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="31" pageNumber="232" phylum="Ochrophyta" rank="species" species="mongolica">
<emphasis id="F1D3032EFFA2FFA5FF2AFBFDFE93FBFF" bold="true" box="[136,375,1074,1101]" italics="true" pageId="31" pageNumber="232">Diploneis mongolica</emphasis>
Metzeltin, Lange-Bertalot &amp; Nergui
</taxonomicName>
(
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)
</heading>
</paragraph>
<paragraph id="C318DF3CFFA2FFA5FF2AFBBAFB17FABE" blockId="31.[136,1452,1141,1292]" pageId="31" pageNumber="232">
The valve is elliptical with convex margins and bluntly rounded apices. The valve length is 34.5 μm and the valve breadth is 15.5 μm. The axial area is narrow and lanceolate, expanding into a large elongate central area. The central area is elongate-oval, 4.5 μm wide. The longitudinal canal is lanceolate with two rows of areolae in the middle of the valve, gradually narrowing into one row toward the valve apices. The raphe is straight and simples, positioned with an expanded depression. The striae are uniseriate and radiate,
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10 μm, composed of round to rectangular areolae,
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10 μm.
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<subSubSection id="8BBD8CB7FFA2FFA5FF2AFAF3FCCDFA57" pageId="31" pageNumber="232" type="description">
<paragraph id="C318DF3CFFA2FFA5FF2AFAF3FCCDFA57" blockId="31.[136,1452,1340,1581]" pageId="31" pageNumber="232">
<emphasis id="F1D3032EFFA2FFA5FF2AFAF3FEA8FAE7" bold="true" box="[136,332,1340,1365]" pageId="31" pageNumber="232">Observations:—</emphasis>
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<emphasis id="F1D3032EFFA2FFA5FEEEFAF3FDDCFAE7" box="[332,568,1340,1365]" italics="true" pageId="31" pageNumber="232">Diploneis mongolica</emphasis>
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was described by
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Metzeltin
<emphasis id="F1D3032EFFA2FFA5FC23FAF3FC57FAE7" box="[897,947,1340,1365]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
. (2009)
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from Lake Khangal,
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. The
<typeStatus id="1C1C619EFFA2FFA5FF2AFAAFFF5CFACB" box="[136,184,1376,1401]" pageId="31" pageNumber="232">type</typeStatus>
material of
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was observed and compared with the specimen from
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. In contrast to the
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population, where
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<emphasis id="F1D3032EFFA2FFA5FE1AFA4BFDB4FA2F" box="[440,592,1412,1437]" italics="true" pageId="31" pageNumber="232">D. mongolica</emphasis>
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is quite dominant, in
<collectingRegion id="016311DEFFA2FFA5FCE3FA4BFBE1FA2F" box="[833,1029,1412,1437]" country="Mongolia" name="Arhangay" pageId="31" pageNumber="232">Arkhangai region</collectingRegion>
this taxon is extremely rare with only
<specimenCount id="D5A114B5FFA2FFA5FF2AFA67FEC4FA73" box="[136,288,1448,1473]" count="1" pageId="31" pageNumber="232" type="generic">one specimen</specimenCount>
found. Even though larger valves of the given size range were not observed, the observed specimen fully fits the original description given by
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Metzeltin
<emphasis id="F1D3032EFFA2FFA5FD3AFA03FD2DFA57" box="[664,713,1484,1509]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
. (2009)
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.
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<emphasis id="F1D3032EFFA2FFA5FF1FFA3FFDE4F9BB" bold="true" box="[189,512,1520,1545]" pageId="31" pageNumber="232">Ecology and Distribution:</emphasis>
—M166A; M167A: examined for comparison, not found in Lake
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but distributed in lakes and streams in
<collectingRegion id="016311DEFFA2FFA5FDA9F9DBFCE8F99F" box="[523,780,1556,1581]" isEnumeration="true" pageId="31" pageNumber="232">Arkhangai and Khentii</collectingRegion>
provinces (
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Metzeltin
<emphasis id="F1D3032EFFA2FFA5FBA0F9DBFBD0F99F" box="[1026,1076,1556,1581]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
. 2009
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).
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<emphasis id="F1D3032EFFA2FFA5FF2AFD51FE8DFD0B" bold="true" box="[136,361,670,697]" italics="true" pageId="31" pageNumber="232">Diploneis subovalis</emphasis>
Cleve
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(
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Valves are broadly elliptic in larger specimens to linear-elliptic in smaller specimens with convex margins and rounded apices. The valve length is 20.033.0 μm and the breadth is 9.017.0 μm. The axial area is linear to lanceolate expanding into a large elongate central area. The central area is elongate, 2.04.0 μm wide. The longitudinal canal is narrow and expanding in the middle of the valve; linear to lanceolate with two rows of areolae throughout its whole length. The raphe is straight and simple, positioned with an expanded depression. Striae are radiate and biseriate,
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10 μm, composed of small round areolae,
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10 μm, counting along one row of the biseriate pattern.
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—M063A; M272A; M273A; M247A; M276A;
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M280A: found throughout
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<taxonomicName id="04A7A4BFFFA2FFA5FF2AFEC5FE05FE97" authority="Hustedt" authorityName="Hustedt" authorityYear="1930" box="[136,481,266,293]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="31" pageNumber="232" phylum="Ochrophyta" rank="species" species="pseudovalis">
<emphasis id="F1D3032EFFA2FFA5FF2AFEC5FE60FE97" bold="true" box="[136,388,266,293]" italics="true" pageId="31" pageNumber="232">Diploneis pseudovalis</emphasis>
Hustedt
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)
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</paragraph>
<paragraph id="C318DF3CFFA2FFA5FF2AFE82FCB1FDB6" blockId="31.[136,1452,333,516]" pageId="31" pageNumber="232">
Valves are linear-elliptic with slightly convex margins and round ends. The valve length is 29.536.5 μm and the valve breadth is 10.511.0 μm. The axial area is narrow, linear to linear-lanceolate expanding into a moderately large central area. The central area is elongate-ovoid and 3.03.5 μm wide. The longitudinal canal is linear to lanceolate with two rows of areolae throughout its whole length, expanding in the middle of the valve. The raphe branches are straight and simple, positioned with an expanded depression. Striae are biseriate, radiate throughout, 12(13) in 10 μm, consisting of small round areolae,
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10 μm along a single row of the biseriate pattern.
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<emphasis id="F1D3032EFFA2FFA5FF2AFDFBFE20FDFF" bold="true" box="[136,452,564,589]" pageId="31" pageNumber="232">Ecology and Distribution:</emphasis>
—M272A; M289A;
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M290A: found in
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on
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<emphasis id="F1D3032EFFA2FFA5FB60FDFBFAEEFDFF" box="[1218,1290,564,589]" italics="true" pageId="31" pageNumber="232">Chara</emphasis>
</taxonomicName>
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and sediment collections taken from
<elevation id="488A380FFFA2FFA5FE28FD97FDD9FDC3" box="[394,573,600,625]" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="3.0" metricValueMin="0.4" pageId="31" pageNumber="232" unit="m" value="17.0" valueMax="30.0" valueMin="4.0">
<quantity id="045F72D9FFA2FFA5FE28FD97FE10FDC3" box="[394,500,600,625]" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="3.0" metricValueMin="0.4" pageId="31" pageNumber="232" unit="m" value="17.0" valueMax="30.0" valueMin="4.0">4 to 30 m</quantity>
depth
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.
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<mods:title id="078F222B8C0C0A3A017CAB2D28D1B5FA">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<taxonomicName id="04A7A4BFFFA7FFA0FF2AFF51FD54FF0B" authority="Lange-Bertalot &amp; Reichardt" authorityName="Lange-Bertalot &amp; Reichardt" authorityYear="2000" box="[136,688,158,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="26" pageNumber="227" phylum="Ochrophyta" rank="species" species="krammeri">
<emphasis id="F1D3032EFFA7FFA0FF2AFF51FE94FF0B" bold="true" box="[136,368,158,185]" italics="true" pageId="26" pageNumber="227">Diploneis krammeri</emphasis>
Lange-Bertalot &amp; Reichardt
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(
<figureCitation id="5B9CC3B9FFA7FFA0FD1DFF6FFCBEFF0B" box="[703,858,160,185]" captionStart-0="FIGURES 113128" captionStart-1="FIGURES 129134" captionStartId-0="24.[136,243,1990,2011]" captionStartId-1="25.[136,243,1884,1905]" captionTargetBox-0="[262,1326,566,1962]" captionTargetBox-1="[155,1433,194,1855]" captionTargetId-0="figure-192@24.[259,1328,563,1966]" captionTargetId-1="figure-14@25.[151,1436,190,1859]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." captionText-1="FIGURES 129134. Diploneis krammeri, SEM external valve views. Fig. 129. View of the whole valve. Fig. 130. Proximal raphe endings. Fig. 131. View of the whole frustule. Fig. 132. Showing the structure of the striae at the valve mantle. Fig. 133. Detailed view of the valve central area. Fig. 134. Distal raphe endings with short deflected terminal fissures. Scale bars = 5 μm (Figs 129, 131); 2 μm (Figs 130, 132134)." figureDoi-0="http://doi.org/10.5281/zenodo.13634164" figureDoi-1="http://doi.org/10.5281/zenodo.13634166" httpUri-0="https://zenodo.org/record/13634164/files/figure.png" httpUri-1="https://zenodo.org/record/13634166/files/figure.png" pageId="26" pageNumber="227">Figs 113134</figureCitation>
)
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Valves are elliptical-lanceolate with convex margins and round ends (
<figureCitation id="5B9CC3B9FFA7FFA0FCC8FF2EFBE4FF4A" box="[874,1024,225,248]" captionStart="FIGURES 113128" captionStartId="24.[136,243,1990,2011]" captionTargetBox="[262,1326,566,1962]" captionTargetId="figure-192@24.[259,1328,563,1966]" captionTargetPageId="24" captionText="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." figureDoi="http://doi.org/10.5281/zenodo.13634164" httpUri="https://zenodo.org/record/13634164/files/figure.png" pageId="26" pageNumber="227">Figs 113124</figureCitation>
). The valve length is 23.034.0 μm, and the valve breadth is 13.017.0 μm. The axial area is linear to lanceolate, expanding toward the round to slightly elongate central area. Internally a thick silica plate covers the longitudinal canal throughout (
<figureCitation id="5B9CC3B9FFA7FFA0FBABFEEEFB83FE8A" box="[1033,1127,289,312]" captionStart="FIGURES 113128" captionStartId="24.[136,243,1990,2011]" captionTargetBox="[262,1326,566,1962]" captionTargetId="figure-192@24.[259,1328,563,1966]" captionTargetPageId="24" captionText="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." figureDoi="http://doi.org/10.5281/zenodo.13634164" httpUri="https://zenodo.org/record/13634164/files/figure.png" pageId="26" pageNumber="227">Fig. 125</figureCitation>
). Externally the central area is elongate, 2.53.5 μm wide. From outside the longitudinal canal is linear to lanceolate, expanded in the middle of the valve with one to two rows of areolae. The external openings of the canal areolae are covered with cribra similar to those on the striae, from which the canal areolae are separated with a thick hyaline area (
<figureCitation id="5B9CC3B9FFA7FFA0FC12FE4EFB9FFE2A" box="[944,1147,385,408]" captionStart="FIGURES 129134" captionStartId="25.[136,243,1884,1905]" captionTargetBox="[155,1433,194,1855]" captionTargetId="figure-14@25.[151,1436,190,1859]" captionTargetPageId="25" captionText="FIGURES 129134. Diploneis krammeri, SEM external valve views. Fig. 129. View of the whole valve. Fig. 130. Proximal raphe endings. Fig. 131. View of the whole frustule. Fig. 132. Showing the structure of the striae at the valve mantle. Fig. 133. Detailed view of the valve central area. Fig. 134. Distal raphe endings with short deflected terminal fissures. Scale bars = 5 μm (Figs 129, 131); 2 μm (Figs 130, 132134)." figureDoi="http://doi.org/10.5281/zenodo.13634166" httpUri="https://zenodo.org/record/13634166/files/figure.png" pageId="26" pageNumber="227">Figs 129, 130, 133</figureCitation>
). From inside, the longitudinal canal is covered with thick silica forming a “depression” where the raphe is placed (
<figureCitation id="5B9CC3B9FFA7FFA0FB8AFE6EFB63FE0A" box="[1064,1159,417,440]" captionStart="FIGURES 113128" captionStartId="24.[136,243,1990,2011]" captionTargetBox="[262,1326,566,1962]" captionTargetId="figure-192@24.[259,1328,563,1966]" captionTargetPageId="24" captionText="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." figureDoi="http://doi.org/10.5281/zenodo.13634164" httpUri="https://zenodo.org/record/13634164/files/figure.png" pageId="26" pageNumber="227">Figs 125</figureCitation>
,
<figureCitation id="5B9CC3B9FFA7FFA0FB30FE6EFB5AFE0A" box="[1170,1214,417,440]" captionStart="FIGURES 152165" captionStartId="28.[136,243,1919,1940]" captionTargetBox="[233,1355,249,1891]" captionTargetId="figure-42@28.[229,1359,245,1894]" captionTargetPageId="28" captionText="FIGURES 152165. Diploneis exigua, LM and SEM valve views. Figs 152159. LM valve views. Figs 153, 156, 158. Specimens from the holotype slide, M272A. Fig. 160. External view of the entire valve. Fig. 161. External distal raphe endings with short terminal fissure. Fig. 162. Broken valve, showing the external and internal opening of the alveolate striae. Fig. 163. Internal view of the distal raphe endings. Fig. 164. Showing the simple proximal raphe endings from the interior. Fig. 165. Internal view of the whole valve. Scale bars = 10 μm (Figs 152160, 165); 3 μm (Fig. 161); 2 μm (Fig. 162); 1 μm (Figs 163, 164)." figureDoi="http://doi.org/10.5281/zenodo.13634170" httpUri="https://zenodo.org/record/13634170/files/figure.png" pageId="26" pageNumber="227">156</figureCitation>
). Externally, the raphe is straight with expanded proximal ends slightly bent to the same side of the valve and positioned within a small depression (
<figureCitation id="5B9CC3B9FFA7FFA0FF32FE2EFEBAFE4A" box="[144,350,481,504]" captionStart="FIGURES 129134" captionStartId="25.[136,243,1884,1905]" captionTargetBox="[155,1433,194,1855]" captionTargetId="figure-14@25.[151,1436,190,1859]" captionTargetPageId="25" captionText="FIGURES 129134. Diploneis krammeri, SEM external valve views. Fig. 129. View of the whole valve. Fig. 130. Proximal raphe endings. Fig. 131. View of the whole frustule. Fig. 132. Showing the structure of the striae at the valve mantle. Fig. 133. Detailed view of the valve central area. Fig. 134. Distal raphe endings with short deflected terminal fissures. Scale bars = 5 μm (Figs 129, 131); 2 μm (Figs 130, 132134)." figureDoi="http://doi.org/10.5281/zenodo.13634166" httpUri="https://zenodo.org/record/13634166/files/figure.png" pageId="26" pageNumber="227">Figs 129, 130, 133</figureCitation>
). From outside the distal raphe endings finish with short terminal fissures bent to the same side of the valve (
<figureCitation id="5B9CC3B9FFA7FFA0FF6DFDCEFE29FDAA" box="[207,461,513,536]" captionStart="FIGURES 129134" captionStartId="25.[136,243,1884,1905]" captionTargetBox="[155,1433,194,1855]" captionTargetId="figure-14@25.[151,1436,190,1859]" captionTargetPageId="25" captionText="FIGURES 129134. Diploneis krammeri, SEM external valve views. Fig. 129. View of the whole valve. Fig. 130. Proximal raphe endings. Fig. 131. View of the whole frustule. Fig. 132. Showing the structure of the striae at the valve mantle. Fig. 133. Detailed view of the valve central area. Fig. 134. Distal raphe endings with short deflected terminal fissures. Scale bars = 5 μm (Figs 129, 131); 2 μm (Figs 130, 132134)." figureDoi="http://doi.org/10.5281/zenodo.13634166" httpUri="https://zenodo.org/record/13634166/files/figure.png" pageId="26" pageNumber="227">Figs 129, 131, 132, 134</figureCitation>
). Internally, the raphe is straight and simple, placed in the “depression” formed by the longitudinal canal (
<figureCitation id="5B9CC3B9FFA7FFA0FEA8FDEEFE83FD8A" box="[266,359,545,568]" captionStart="FIGURES 113128" captionStartId="24.[136,243,1990,2011]" captionTargetBox="[262,1326,566,1962]" captionTargetId="figure-192@24.[259,1328,563,1966]" captionTargetPageId="24" captionText="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." figureDoi="http://doi.org/10.5281/zenodo.13634164" httpUri="https://zenodo.org/record/13634164/files/figure.png" pageId="26" pageNumber="227">Fig. 125</figureCitation>
). The proximal raphe endings are the only raphe section reaching the height of the longitudinal canal in contrast to the only slightly elevated distal ends (
<figureCitation id="5B9CC3B9FFA7FFA0FD30FD8EFCCDFDEA" box="[658,809,577,600]" captionStart="FIGURES 113128" captionStartId="24.[136,243,1990,2011]" captionTargetBox="[262,1326,566,1962]" captionTargetId="figure-192@24.[259,1328,563,1966]" captionTargetPageId="24" captionText="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." figureDoi="http://doi.org/10.5281/zenodo.13634164" httpUri="https://zenodo.org/record/13634164/files/figure.png" pageId="26" pageNumber="227">Figs 125127</figureCitation>
). The striae are radiate, 11(12) in 10 μm, with round areolae covered with cribra (
<figureCitation id="5B9CC3B9FFA7FFA0FEC6FDAEFE25FDCA" box="[356,449,609,632]" captionStart="FIGURES 113128" captionStartId="24.[136,243,1990,2011]" captionTargetBox="[262,1326,566,1962]" captionTargetId="figure-192@24.[259,1328,563,1966]" captionTargetPageId="24" captionText="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." figureDoi="http://doi.org/10.5281/zenodo.13634164" httpUri="https://zenodo.org/record/13634164/files/figure.png" pageId="26" pageNumber="227">Fig. 128</figureCitation>
),
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10 μm. The external areolar openings increase in size and become more complex in structure, or are biseriate toward the valve margins (
<figureCitation id="5B9CC3B9FFA7FFA0FD09FD4EFC4DFD2A" box="[683,937,641,664]" captionStart="FIGURES 129134" captionStartId="25.[136,243,1884,1905]" captionTargetBox="[155,1433,194,1855]" captionTargetId="figure-14@25.[151,1436,190,1859]" captionTargetPageId="25" captionText="FIGURES 129134. Diploneis krammeri, SEM external valve views. Fig. 129. View of the whole valve. Fig. 130. Proximal raphe endings. Fig. 131. View of the whole frustule. Fig. 132. Showing the structure of the striae at the valve mantle. Fig. 133. Detailed view of the valve central area. Fig. 134. Distal raphe endings with short deflected terminal fissures. Scale bars = 5 μm (Figs 129, 131); 2 μm (Figs 130, 132134)." figureDoi="http://doi.org/10.5281/zenodo.13634166" httpUri="https://zenodo.org/record/13634166/files/figure.png" pageId="26" pageNumber="227">Figs 129, 131, 132, 134</figureCitation>
). Internally the alveoli open via a single continuous and elongate opening covered with a fine silica layer (
<figureCitation id="5B9CC3B9FFA7FFA0FD57FD6EFC68FD0A" box="[757,908,673,696]" captionStart="FIGURES 113128" captionStartId="24.[136,243,1990,2011]" captionTargetBox="[262,1326,566,1962]" captionTargetId="figure-192@24.[259,1328,563,1966]" captionTargetPageId="24" captionText="FIGURES 113128. Diploneis krammeri, LM and SEM valve views. Figs 113124. LM valve views. Fig. 125. Internal view of the whole valve. Fig. 126. Detailed view of the internal proximal raphe ends. Fig. 127. Internal view of the distal raphe endings. Fig. 128. Close view of the external cribrate occlusions. Scale bars = 10 μm (Figs 113124); 5 μm (Figs 125, 127); 2 μm (Fig. 126); 1 μm (Fig. 128)." figureDoi="http://doi.org/10.5281/zenodo.13634164" httpUri="https://zenodo.org/record/13634164/files/figure.png" pageId="26" pageNumber="227">Figs 125, 126</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA7FFA0FF2AFD27FDFFFC07" pageId="26" pageNumber="227" type="description">
<paragraph id="C318DF3CFFA7FFA0FF2AFD27FDFFFC07" blockId="26.[136,1452,744,1021]" pageId="26" pageNumber="227">
<emphasis id="F1D3032EFFA7FFA0FF2AFD27FED4FCB3" bold="true" box="[136,304,744,769]" pageId="26" pageNumber="227">Observations:</emphasis>
—In the original description of
<taxonomicName id="04A7A4BFFFA7FFA0FD37FD27FCC1FCB3" authorityName="Lange-Bertalot &amp; Reichardt" authorityYear="2000" box="[661,805,744,769]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="26" pageNumber="227" phylum="Ochrophyta" rank="species" species="krammeri">
<emphasis id="F1D3032EFFA7FFA0FD37FD27FCC1FCB3" box="[661,805,744,769]" italics="true" pageId="26" pageNumber="227">D. krammeri</emphasis>
</taxonomicName>
the longitudinal canal is composed of one row of areolae throughout the whole length (
<bibRefCitation id="A736A2CDFFA7FFA0FE6EFCC3FCA1FC97" author="Lange-Bertalot, H. &amp; Reichardt, E." box="[460,837,780,805]" pageId="26" pageNumber="227" pagination="650 - 669" refId="ref27660" refString="Lange-Bertalot, H. &amp; Reichardt, E. (2000) Diploneis ovalis sensu stricto und Diploneis krammeri nov. spec. Revision des aktuellen Konzepts von Diploneis ovalis (Hilse) Cleve. Iconographia Diatomologica 9: 650 - 669." type="journal article" year="2000">Lange-Bertalot &amp; Reichardt 2000</bibRefCitation>
: pl. 4, figs 110, 12). However, on the SEM illustrations the external openings of the longitudinal canal are composed of two rows of areolae in the middle of the valve narrowing into one at the valve apices (
<bibRefCitation id="A736A2CDFFA7FFA0FE62FC9BFCD8FCDF" author="Lange-Bertalot, H. &amp; Reichardt, E." box="[448,828,852,877]" pageId="26" pageNumber="227" pagination="650 - 669" refId="ref27660" refString="Lange-Bertalot, H. &amp; Reichardt, E. (2000) Diploneis ovalis sensu stricto und Diploneis krammeri nov. spec. Revision des aktuellen Konzepts von Diploneis ovalis (Hilse) Cleve. Iconographia Diatomologica 9: 650 - 669." type="journal article" year="2000">Lange-Bertalot &amp; Reichardt 2000</bibRefCitation>
: pl. 5, figs 15). The longitudinal canal in Lake
<collectingRegion id="016311DEFFA7FFA0FAEFFC9BFA48FCDF" box="[1357,1452,852,877]" country="Mongolia" name="Hovsgol" pageId="26" pageNumber="227">Hövsgöl</collectingRegion>
populations are composed of one row of areolae throughout, which unambiguously fits the LM illustrations given by
<bibRefCitation id="A736A2CDFFA7FFA0FF2AFC53FDF3FC07" author="Lange-Bertalot, H. &amp; Reichardt, E." box="[136,535,924,949]" pageId="26" pageNumber="227" pagination="650 - 669" refId="ref27660" refString="Lange-Bertalot, H. &amp; Reichardt, E. (2000) Diploneis ovalis sensu stricto und Diploneis krammeri nov. spec. Revision des aktuellen Konzepts von Diploneis ovalis (Hilse) Cleve. Iconographia Diatomologica 9: 650 - 669." type="journal article" year="2000">Lange-Bertalot &amp; Reichardt (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA7FFA0FF1FFC0FFC2EFC4F" pageId="26" pageNumber="227" type="biology_ecology">
<paragraph id="C318DF3CFFA7FFA0FF1FFC0FFC2EFC4F" blockId="26.[136,1452,744,1021]" pageId="26" pageNumber="227">
<emphasis id="F1D3032EFFA7FFA0FF1FFC0FFE1FFC6B" bold="true" box="[189,507,960,985]" pageId="26" pageNumber="227">Ecology and Distribution:</emphasis>
—M063A; M248A; M273A; M274A; M280A;
<materialsCitation id="73CFD561FFA7FFA0FB85FC0FFC22FC4F" location="Lake" pageId="26" pageNumber="227" specimenCount="1" stateProvince="Hovsgol">
M329A: distributed in shallow to deep benthic habitats throughout
<location id="C67889E7FFA7FFA0FE5FFC2BFDD1FC4F" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA7FFA0FF2AFF51FC2EFC4F:C67889E7FFA7FFA0FE5FFC2BFDD1FC4F" box="[509,565,996,1021]" name="Lake" pageId="26" pageNumber="227" stateProvince="Hovsgol">Lake</location>
<collectingRegion id="016311DEFFA7FFA0FD9EFC2BFD7FFC4F" box="[572,667,996,1021]" country="Mongolia" name="Hovsgol" pageId="26" pageNumber="227">Hövsgöl</collectingRegion>
in the sediments and marl
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,301 @@
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<paragraph id="C318DF3CFFA7FFA0FF2AFBE5FD59FBF7" blockId="26.[136,701,1066,1093]" box="[136,701,1066,1093]" pageId="26" pageNumber="227">
<heading id="98506850FFA7FFA0FF2AFBE5FD59FBF7" box="[136,701,1066,1093]" fontSize="11" level="2" pageId="26" pageNumber="227" reason="3">
<taxonomicName id="04A7A4BFFFA7FFA0FF2AFBE5FE4AFBF7" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[136,430,1066,1093]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="26" pageNumber="227" phylum="Ochrophyta" rank="species" species="soninkhishigae" status="sp. nov.">Diploneis soninkhishigae</taxonomicName>
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<emphasis id="F1D3032EFFA7FFA0FE16FBE3FDE7FBF7" box="[436,515,1068,1093]" italics="true" pageId="26" pageNumber="227">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="5B9CC3B9FFA7FFA0FDB8FBE3FD52FBF7" box="[538,694,1068,1093]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Figs 135151</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFFA7FFA0FF2AFBA2FACEF9D6" blockId="26.[136,1452,1133,1636]" pageId="26" pageNumber="227">
The valves are elliptical-lanceolate with convex margins and round ends (
<figureCitation id="5B9CC3B9FFA7FFA0FC35FBA2FBCAFB36" box="[919,1070,1133,1156]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Figs 135148</figureCitation>
). The valve length is 14.524.5 μm, and the valve breadth is 8.012.0 μm. The axial area is narrow, linear to lanceolate expanding into a small elongate central area. From inside, a wide linear silica plate covers the longitudinal canal throughout the whole valve length (
<figureCitation id="5B9CC3B9FFA7FFA0FAB1FB62FA95FB76" box="[1299,1393,1197,1220]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 149</figureCitation>
). The central area is small and elongate, 1.52.5 μm wide. Externally, the longitudinal canal is narrow slightly expanded in the middle of the valve with one row of areolae throughout the whole valve length (
<figureCitation id="5B9CC3B9FFA7FFA0FC77FB22FB88FAB6" box="[981,1132,1261,1284]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Figs 135148</figureCitation>
). The areolae of the longitudinal canal are covered with cribra similar to the striae (
<figureCitation id="5B9CC3B9FFA7FFA0FD61FAC2FCC5FA96" box="[707,801,1293,1316]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 150</figureCitation>
). In the middle of the valve the cribra of the canal are simple, becoming wider and more complex in structure towards the valve apices (
<figureCitation id="5B9CC3B9FFA7FFA0FC05FAE2FBE2FAF6" box="[935,1030,1325,1348]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 148</figureCitation>
). From inside, the longitudinal canal is closed with thick silica plates, forming a “trench” where the raphe is inserted. Externally, the raphe is simple and straight with drop-like proximal ends that are slightly bent, positioned within an expanded depression (
<figureCitation id="5B9CC3B9FFA7FFA0FB22FAA2FAFCFA36" box="[1152,1304,1389,1412]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Figs 148, 150</figureCitation>
). Distally, the raphe branches finish with short drop-like terminal fissures, bent to the same side of the valve (
<figureCitation id="5B9CC3B9FFA7FFA0FB2FFA42FB0FFA16" box="[1165,1259,1421,1444]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 148</figureCitation>
). The distal raphe ends terminate some distance from the valve mantle. Internally, the raphe is straight with simple proximal raphe ends, placed in the “trench” formed by the longitudinal canal (
<figureCitation id="5B9CC3B9FFA7FFA0FD34FA02FD17FA56" box="[662,755,1485,1508]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 149</figureCitation>
). The proximal raphe endings are raised in the central area, while the distal raphe ends are also slightly raised but terminate into small helictoglossae (
<figureCitation id="5B9CC3B9FFA7FFA0FBAEFA22FB8EF9B6" box="[1036,1130,1517,1540]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 149</figureCitation>
). Striae are usually uniseriate, becoming biseriate toward the valve margins,
<quantity id="045F72D9FFA7FFA0FDD0F9C2FD2BF996" box="[626,719,1549,1572]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.81" metricValueMin="2.794" pageId="26" pageNumber="227" unit="in" value="13.0" valueMax="15.0" valueMin="11.0">1115 in</quantity>
10 μm, and composed of round areolae,
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10 μm. Externally, the areolae are covered with cribrate occlusions (
<figureCitation id="5B9CC3B9FFA7FFA0FD3EF9E2FD1EF9F6" box="[668,762,1581,1604]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 150</figureCitation>
), increasing in size towards the valve margins (
<figureCitation id="5B9CC3B9FFA7FFA0FB5DF9E2FAB9F9F6" box="[1279,1373,1581,1604]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 151</figureCitation>
). From inside, each alveolus opens through a single and continuous opening covered with a thin silica layer (
<figureCitation id="5B9CC3B9FFA7FFA0FB62F982FAFAF9D6" box="[1216,1310,1613,1636]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Fig. 149</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA7FFA0FF2AF959FCF3F8A5" pageId="26" pageNumber="227" type="materials_examined">
<paragraph id="C318DF3CFFA7FFA0FF2AF959FCF3F8A5" blockId="26.[136,1451,1686,1815]" pageId="26" pageNumber="227">
<emphasis id="F1D3032EFFA7FFA0FF2AF959FF3EF919" bold="true" box="[136,218,1686,1707]" pageId="26" pageNumber="227">Type:—</emphasis>
<materialsCitation id="73CFD561FFA7FFA0FF78F959FC1EF919" box="[218,1018,1686,1707]" country="Mongolia" county="Lake" latitude="50.9176" location="Coordinates" municipality="National Park" pageId="26" pageNumber="227" specimenCount="1" stateProvince="Hovsgol">
<collectingCountry id="BBB09FACFFA7FFA0FF78F959FEBBF919" box="[218,351,1686,1707]" name="Mongolia" pageId="26" pageNumber="227">MONGOLIA</collectingCountry>
,
<collectingCounty id="2A79A7B0FFA7FFA0FECBF959FE7DF919" box="[361,409,1686,1707]" pageId="26" pageNumber="227">Lake</collectingCounty>
<collectingRegion id="016311DEFFA7FFA0FE3DF959FE14F919" box="[415,496,1686,1707]" country="Mongolia" name="Hovsgol" pageId="26" pageNumber="227">Hövsgöl</collectingRegion>
(
<collectingRegion id="016311DEFFA7FFA0FE5FF959FDB4F919" box="[509,592,1686,1707]" country="Mongolia" name="Hovsgol" pageId="26" pageNumber="227">Hövsgöl</collectingRegion>
<collectingMunicipality id="237C4546FFA7FFA0FDF4F959FD3BF919" box="[598,735,1686,1707]" pageId="26" pageNumber="227">National Park</collectingMunicipality>
).
<location id="C67889E7FFA7FFA0FD52F959FC8EF919" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA7FFA6FF2AFBE5FB88FF6F:C67889E7FFA7FFA0FD52F959FC8EF919" box="[752,874,1686,1707]" country="Mongolia" county="Lake" latitude="50.9176" municipality="National Park" name="Coordinates" pageId="26" pageNumber="227" stateProvince="Hovsgol">Coordinates</location>
:
<geoCoordinate id="A693B9FBFFA7FFA0FCD6F959FC1EF919" box="[884,1018,1686,1707]" degrees="50" direction="north" minutes="55.056" orientation="latitude" pageId="26" pageNumber="227" precision="1" value="50.9176">50°55.056 N</geoCoordinate>
</materialsCitation>
<materialsCitation id="73CFD561FFA7FFA0FC59F959FB24F941" collectingDate="1998-07-18" collectionCode="ANSP" collectorName="Sediments &amp; Edlund Collection &amp; Mark B. Edlund &amp; Nergui Soninkhishig &amp; Slide M" country="Mongolia" elevation="40" location="Science Museum of Minnesota" longitude="100.63187" pageId="26" pageNumber="227" specimenCode="GC-36354" specimenCount="1" typeStatus="holotype">
;
<geoCoordinate id="A693B9FBFFA7FFA0FBA7F959FB71F919" box="[1029,1173,1686,1707]" degrees="100" direction="east" minutes="37.912" orientation="longitude" pageId="26" pageNumber="227" precision="1" value="100.63187">100°37.912 E</geoCoordinate>
,
<collectorName id="6E52BAEAFFA7FFA0FB02F959FAE0F919" box="[1184,1284,1686,1707]" pageId="26" pageNumber="227">Sediments</collectorName>
from
<elevation id="488A380FFFA7FFA0FA9DF959FA4FF919" box="[1343,1451,1686,1707]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="227" unit="m" value="40.0">
<quantity id="045F72D9FFA7FFA0FA9DF959FA94F919" box="[1343,1392,1686,1707]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="227" unit="m" value="40.0">40 m</quantity>
depth
</elevation>
(accession number: M248A, M.B.
<collectorName id="6E52BAEAFFA7FFA0FDB7F975FD2EF97D" box="[533,714,1722,1743]" pageId="26" pageNumber="227">Edlund Collection</collectorName>
,
<location id="C67889E7FFA7FFA0FD74F975FBEEF97D" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA7FFA6FF2AFBE5FB88FF6F:C67889E7FFA7FFA0FD74F975FBEEF97D" box="[726,1034,1722,1743]" country="Mongolia" longitude="100.63187" name="Science Museum of Minnesota" pageId="26" pageNumber="227">Science Museum of Minnesota</location>
, collected by
<collectorName id="6E52BAEAFFA7FFA0FB37F975FAD1F97D" box="[1173,1333,1722,1743]" pageId="26" pageNumber="227">Mark B. Edlund</collectorName>
and
<collectorName id="6E52BAEAFFA7FFA0FACBF975FEDAF941" pageId="26" pageNumber="227">Nergui Soninkhishig</collectorName>
,
<date id="B719F9FCFFA7FFA0FEE8F911FE2EF941" box="[330,458,1758,1779]" pageId="26" pageNumber="227" value="1998-07-18">
<collectingDate id="A75D0014FFA7FFA0FEE8F911FE2EF941" box="[330,458,1758,1779]" pageId="26" pageNumber="227" value="1998-07-18">18 July 1998</collectingDate>
</date>
) (
<collectorName id="6E52BAEAFFA7FFA0FE42F911FDD4F941" box="[480,560,1758,1779]" pageId="26" pageNumber="227">Slide M</collectorName>
248A,
<collectionCode id="A5B647F9FFA7FFA0FDD3F911FD49F941" box="[625,685,1758,1779]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="26" pageNumber="227">ANSP</collectionCode>
<specimenCode id="93017747FFA7FFA0FD17F911FCF9F941" box="[693,797,1758,1779]" collectionCode="GC-" pageId="26" pageNumber="227">GC-36354</specimenCode>
, GCM-24056),
<emphasis id="F1D3032EFFA7FFA0FC1BF911FB24F941" bold="true" box="[953,1216,1758,1779]" pageId="26" pageNumber="227">
<typeStatus id="1C1C619EFFA7FFA0FC1BF911FBF6F941" box="[953,1042,1758,1779]" pageId="26" pageNumber="227" type="holotype">holotype</typeStatus>
, designated here
</emphasis>
</materialsCitation>
; example specimens on
<figureCitation id="5B9CC3B9FFA7FFA0FF1FF8CDFEA0F8A5" box="[189,324,1794,1815]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Figs 136, 137</figureCitation>
;
<materialsCitation id="73CFD561FFA7FFA0FEEFF8CDFCF0F8A5" box="[333,788,1794,1815]" collectionCode="CAS" country="Mongolia" location="Slide" pageId="26" pageNumber="227" specimenCount="1" typeStatus="isotype">
(
<location id="C67889E7FFA7FFA0FEF6F8CDFE63F8A5" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFA7FFA6FF2AFBE5FB88FF6F:C67889E7FFA7FFA0FEF6F8CDFE63F8A5" box="[340,391,1794,1815]" country="Mongolia" name="Slide" pageId="26" pageNumber="227">Slide</location>
919048,
<collectionCode id="A5B647F9FFA7FFA0FE43F8CDFDF6F8A5" box="[481,530,1794,1815]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="26" pageNumber="227" type="Museum">CAS</collectionCode>
,
<emphasis id="F1D3032EFFA7FFA0FDB9F8CDFCEEF8A5" bold="true" box="[539,778,1794,1815]" pageId="26" pageNumber="227">
<typeStatus id="1C1C619EFFA7FFA0FDB9F8CDFD87F8A5" box="[539,611,1794,1815]" pageId="26" pageNumber="227" type="isotype">isotype</typeStatus>
designated here
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA7FFA1FF2AF887FC88FE6B" lastPageId="27" lastPageNumber="228" pageId="26" pageNumber="227" type="etymology">
<paragraph id="C318DF3CFFA7FFA0FF2AF887FB5EF837" blockId="26.[136,1452,1864,2069]" pageId="26" pageNumber="227">
<emphasis id="F1D3032EFFA7FFA0FF2AF887FECAF8D3" bold="true" box="[136,302,1864,1889]" pageId="26" pageNumber="227">Etymology:—</emphasis>
The species is named in honor of Dr. Nergui Soninkhishig, National University of
<collectingCountry id="BBB09FACFFA7FFA0FB7CF887FAAAF8D3" box="[1246,1358,1864,1889]" name="Mongolia" pageId="26" pageNumber="227">Mongolia</collectingCountry>
, Botany Faculty, for her pivotal role in developing diatom research in
<collectingCountry id="BBB09FACFFA7FFA0FC9AF8A3FC41F837" box="[824,933,1900,1925]" name="Mongolia" pageId="26" pageNumber="227">Mongolia</collectingCountry>
and our long friendship.
</paragraph>
<paragraph id="C318DF3CFFA7FFA1FF1FF85FFC88FE6B" blockId="26.[136,1452,1864,2069]" lastBlockId="27.[136,1452,160,473]" lastPageId="27" lastPageNumber="228" pageId="26" pageNumber="227">
<emphasis id="F1D3032EFFA7FFA0FF1FF85FFE81F81B" bold="true" box="[189,357,1936,1961]" pageId="26" pageNumber="227">Observations:</emphasis>
<taxonomicName id="04A7A4BFFFA7FFA0FE23F85FFD45F81B" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[385,673,1936,1961]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="26" pageNumber="227" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA7FFA0FE23F85FFD45F81B" box="[385,673,1936,1961]" italics="true" pageId="26" pageNumber="227">Diploneis soninkhishigae</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="04A7A4BFFFA7FFA0FC23F85FFF3EF87F" authority="Jovanovska, Nakov &amp; Levkov (2013 a" authorityName="Jovanovska, Nakov &amp; Levkov" authorityYear="2013" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="26" pageNumber="227" phylum="Ochrophyta" rank="species" species="vetusa">
<emphasis id="F1D3032EFFA7FFA0FC23F85FFBA5F81B" box="[897,1089,1936,1961]" italics="true" pageId="26" pageNumber="227">Diploneis vetusa</emphasis>
<bibRefCitation id="A736A2CDFFA7FFA0FBE9F85FFF3EF87F" author="Jovanovska, E. &amp; Nakov, T. &amp; Levkov, Z." pageId="26" pageNumber="227" pagination="237 - 262" refId="ref27009" refString="Jovanovska, E., Nakov, T. &amp; Levkov, Z. (2013 a) Observations of the genus Diploneis (Ehrenberg) Cleve from Lake Ohrid, Macedonia. Diatom Research 28: 237 - 262. http: // dx. doi. org / 10.1080 / 0269249 x. 2013.797219" type="journal article" year="2013">Jovanovska, Nakov &amp; Levkov (2013a</bibRefCitation>
</taxonomicName>
: figs 106121), which is characterized with biseriate striae throughout the whole length. In contrast,
<taxonomicName id="04A7A4BFFFA7FFA0FA9CF87BFED4F843" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="26" pageNumber="227" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA7FFA0FA9CF87BFED4F843" italics="true" pageId="26" pageNumber="227">Diploneis soninkhishigae</emphasis>
</taxonomicName>
has uniseriate striae becoming biseriate towards the valve margins. A population of
<taxonomicName id="04A7A4BFFFA7FFA0FB43F817FA4FF843" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[1249,1451,2008,2033]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="26" pageNumber="227" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA7FFA0FB43F817FA4FF843" box="[1249,1451,2008,2033]" italics="true" pageId="26" pageNumber="227">D. soninkhishigae</emphasis>
</taxonomicName>
was also observed in Lake
<collectingRegion id="016311DEFFA7FFA0FE61F833FDE3F7A7" box="[451,519,2044,2069]" country="Macedonia" name="Ohrid" pageId="26" pageNumber="227">Ohrid</collectingRegion>
,
<collectingCountry id="BBB09FACFFA7FFA0FDB6F833FD74F7A7" box="[532,656,2044,2069]" name="Macedonia" pageId="26" pageNumber="227">Macedonia</collectingCountry>
(
<figureCitation id="5B9CC3B9FFA7FFA0FD00F833FCA0F7A7" box="[674,836,2044,2069]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="26" pageNumber="227">Figs 142147</figureCitation>
). The striae pattern is more visible in the Lake
<collectingRegion id="016311DEFFA7FFA0FAC8F833FA4FF7A7" box="[1386,1451,2044,2069]" country="Macedonia" name="Ohrid" pageId="26" pageNumber="227">Ohrid</collectingRegion>
population (
<figureCitation id="5B9CC3B9FFA6FFA1FEB0FF6FFE5CFF0B" box="[274,440,160,185]" captionStart="FIGURES 135151" captionStartId="27.[136,243,1918,1939]" captionTargetBox="[205,1382,501,1888]" captionTargetId="figure-190@27.[202,1386,497,1893]" captionTargetPageId="27" captionText="FIGURES 135151. Diploneis soninkhishigae, LM and SEM valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151)." figureDoi="http://doi.org/10.5281/zenodo.13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="27" pageNumber="228">Figs 145, 147</figureCitation>
).
<taxonomicName id="04A7A4BFFFA6FFA1FE6DFF6FFD83FF0B" authorityName="Lange-Bertalot" authorityYear="2004" box="[463,615,160,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="fontanella">
<emphasis id="F1D3032EFFA6FFA1FE6DFF6FFD83FF0B" box="[463,615,160,185]" italics="true" pageId="27" pageNumber="228">D. fontanella</emphasis>
</taxonomicName>
has a linear-elliptical valve shape in contrast to the elliptical-lanceolate outline in
<taxonomicName id="04A7A4BFFFA6FFA1FF59FF0BFE22FF6F" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[251,454,196,221]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA6FFA1FF59FF0BFEEBFF6F" box="[251,271,196,221]" italics="true" pageId="27" pageNumber="228">D</emphasis>
.
<emphasis id="F1D3032EFFA6FFA1FEBCFF0BFE22FF6F" box="[286,454,196,221]" italics="true" pageId="27" pageNumber="228">soninkhishigae</emphasis>
</taxonomicName>
. In addition,
<taxonomicName id="04A7A4BFFFA6FFA1FDFDFF0BFD10FF6F" authorityName="Lange-Bertalot" authorityYear="2004" box="[607,756,196,221]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="fontanella">
<emphasis id="F1D3032EFFA6FFA1FDFDFF0BFD97FF6F" box="[607,627,196,221]" italics="true" pageId="27" pageNumber="228">D</emphasis>
.
<emphasis id="F1D3032EFFA6FFA1FD20FF0BFD10FF6F" box="[642,756,196,221]" italics="true" pageId="27" pageNumber="228">fontanella</emphasis>
</taxonomicName>
has a higher stria and areola density (striae:
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10 μm; areolae:
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10 μm), uniseriate striae and a round central area (
<bibRefCitation id="A736A2CDFFA6FFA1FC14FF27FACCFEB3" author="Werum, M. &amp; Lange-Bertalot, H." box="[950,1320,232,257]" pageId="27" pageNumber="228" pagination="3 - 417" refId="ref28669" refString="Werum, M. &amp; Lange-Bertalot, H. (2004) Diatoms in Springs from Central European. Iconographia Diatomologica 13: 3 - 417." type="journal article" year="2004">Werum &amp; Lange-Bertalot 2004</bibRefCitation>
).
<taxonomicName id="04A7A4BFFFA6FFA1FA9CFF27FE56FE97" authority="Lange-Bertalot" authorityName="Lange-Bertalot" authorityYear="2004" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="separanda">
<emphasis id="F1D3032EFFA6FFA1FA9CFF27FF1AFE97" italics="true" pageId="27" pageNumber="228">Diploneis separanda</emphasis>
Lange-Bertalot
</taxonomicName>
(2004: pl. 76, fig. 116; pl. 77, fig. 15) is another linear-elliptic taxon, with uniseriate striae becoming biseriate towards the valve margins. However, in addition to differences in valve shape, there are also differences in the stria density (
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10 μm in
<taxonomicName id="04A7A4BFFFA6FFA1FD72FE9BFC78FEDF" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[720,924,340,365]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA6FFA1FD72FE9BFD00FEDF" box="[720,740,340,365]" italics="true" pageId="27" pageNumber="228">D</emphasis>
.
<emphasis id="F1D3032EFFA6FFA1FD56FE9BFC78FEDF" box="[756,924,340,365]" italics="true" pageId="27" pageNumber="228">soninkhishigae</emphasis>
</taxonomicName>
<emphasis id="F1D3032EFFA6FFA1FC07FE9BFC58FEDF" box="[933,956,340,365]" italics="true" pageId="27" pageNumber="228">vs</emphasis>
.
<quantity id="045F72D9FFA6FFA1FC6EFE9BFBD5FEDF" box="[972,1073,340,365]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" metricValueMax="5.334" metricValueMin="4.826" pageId="27" pageNumber="228" unit="in" value="20.0" valueMax="21.0" valueMin="19.0">1921 in</quantity>
10 μm in
<taxonomicName id="04A7A4BFFFA6FFA1FB0EFE9BFAA2FEDF" authorityName="Lange-Bertalot" authorityYear="2004" box="[1196,1350,340,365]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="separanda">
<emphasis id="F1D3032EFFA6FFA1FB0EFE9BFAA2FEDF" box="[1196,1350,340,365]" italics="true" pageId="27" pageNumber="228">D. separanda</emphasis>
</taxonomicName>
) and the areola density (
<quantity id="045F72D9FFA6FFA1FE91FEB7FE73FE23" box="[307,407,376,401]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="27" pageNumber="228" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">1012 in</quantity>
10 μm in
<taxonomicName id="04A7A4BFFFA6FFA1FDABFEB7FD37FE23" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[521,723,376,401]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA6FFA1FDABFEB7FDF9FE23" box="[521,541,376,401]" italics="true" pageId="27" pageNumber="228">D</emphasis>
.
<emphasis id="F1D3032EFFA6FFA1FD89FEB7FD37FE23" box="[555,723,376,401]" italics="true" pageId="27" pageNumber="228">soninkhishigae</emphasis>
</taxonomicName>
<emphasis id="F1D3032EFFA6FFA1FD78FEB7FD15FE23" box="[730,753,376,401]" italics="true" pageId="27" pageNumber="228">vs</emphasis>
.
<quantity id="045F72D9FFA6FFA1FD5DFEB7FC86FE23" box="[767,866,376,401]" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="8.89" metricValueMin="7.62" pageId="27" pageNumber="228" unit="in" value="32.5" valueMax="35.0" valueMin="30.0">3035 in</quantity>
10 μm in
<taxonomicName id="04A7A4BFFFA6FFA1FC76FEB7FB88FE23" authorityName="Lange-Bertalot" authorityYear="2004" box="[980,1132,376,401]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="separanda">
<emphasis id="F1D3032EFFA6FFA1FC76FEB7FB88FE23" box="[980,1132,376,401]" italics="true" pageId="27" pageNumber="228">D. separanda</emphasis>
</taxonomicName>
). Additionally, SEM images show that the cribrate occlusions in
<taxonomicName id="04A7A4BFFFA6FFA1FDB5FE53FD04FE07" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[535,736,412,437]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA6FFA1FDB5FE53FDCFFE07" box="[535,555,412,437]" italics="true" pageId="27" pageNumber="228">D</emphasis>
.
<emphasis id="F1D3032EFFA6FFA1FD9AFE53FD04FE07" box="[568,736,412,437]" italics="true" pageId="27" pageNumber="228">soninkhishigae</emphasis>
</taxonomicName>
are more numerous and more complex in structure than those in
<taxonomicName id="04A7A4BFFFA6FFA1FF2AFE0FFEC4FE6B" authorityName="Lange-Bertalot" authorityYear="2004" box="[136,288,448,473]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="separanda">
<emphasis id="F1D3032EFFA6FFA1FF2AFE0FFEC4FE6B" box="[136,288,448,473]" italics="true" pageId="27" pageNumber="228">D. separanda</emphasis>
</taxonomicName>
(Werum &amp; Lange-Bertalot: 338, pl. 76, figs 16, 15).
</paragraph>
</subSubSection>
<caption id="97D88FB4FFA6FFA1FF2AF8B1FF0FF791" ID-DOI="http://doi.org/10.5281/zenodo.13634168" ID-Zenodo-Dep="13634168" httpUri="https://zenodo.org/record/13634168/files/figure.png" pageId="27" pageNumber="228" startId="27.[136,243,1918,1939]" targetBox="[205,1382,501,1888]" targetPageId="27" targetType="figure">
<paragraph id="C318DF3CFFA6FFA1FF2AF8B1FF0FF791" blockId="27.[136,1452,1918,2083]" pageId="27" pageNumber="228">
<emphasis id="F1D3032EFFA6FFA1FF2AF8B1FEB7F821" bold="true" box="[136,339,1918,1939]" pageId="27" pageNumber="228">FIGURES 135151.</emphasis>
<taxonomicName id="04A7A4BFFFA6FFA1FEFAF8B1FD3DF821" authority=", LM and SEM" authorityName="LM and SEM" box="[344,729,1918,1939]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="27" pageNumber="228" phylum="Ochrophyta" rank="species" species="soninkhishigae">
<emphasis id="F1D3032EFFA6FFA1FEFAF8B1FDAFF821" box="[344,587,1918,1939]" italics="true" pageId="27" pageNumber="228">Diploneis soninkhishigae</emphasis>
, LM and SEM
</taxonomicName>
valve views. Figs 135141. LM valve views, Lake Hövsgöl. Figs 142147. LM valve views, Lake Ohrid specimens. Figs 136, 137. Specimens from the holotype slide M248A. Fig. 148. External valve view. Fig. 149. Internal view of the whole valve. Fig. 150. View of the valve central area with proximal raphe ends. Fig. 151. Close-up view of the external openings of the alveolate striae, covered with cribrum. Scale bars = 10 μm (Figs 135147); 5 μm (Figs 148, 149); 2 μm (Figs 150, 151).
</paragraph>
</caption>
<subSubSection id="8BBD8CB7FFA1FFA6FF1FFF6FFB88FF6F" pageId="28" pageNumber="229" type="biology_ecology">
<paragraph id="C318DF3CFFA1FFA6FF1FFF6FFA48FF0B" blockId="28.[136,1452,160,221]" box="[189,1452,160,185]" pageId="28" pageNumber="229">
<emphasis id="F1D3032EFFA1FFA6FF1FFF6FFE12FF0B" bold="true" box="[189,502,160,185]" pageId="28" pageNumber="229">Ecology and Distribution:</emphasis>
—M248A; M272A; M273A; M274A; M287A; M289A; M291A: found throughout
</paragraph>
<paragraph id="C318DF3CFFA1FFA6FF2AFF0BFB88FF6F" blockId="28.[136,1452,160,221]" box="[136,1132,196,221]" pageId="28" pageNumber="229">
Lake
<collectingRegion id="016311DEFFA1FFA6FF65FF0BFEC2FF6F" box="[199,294,196,221]" country="Mongolia" name="Hovsgol" pageId="28" pageNumber="229">Hövsgöl</collectingRegion>
in shallow to deep water habitats including marl, sediments, and epipelon.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,383 @@
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<mods:title id="33A7A8446D1311B0A35D9EE035257656">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<treatment id="4B0E6E2AFFAAFFA2FF2AFF51FABCFDA8" ID-DOI="http://doi.org/10.5281/zenodo.13634229" ID-Zenodo-Dep="13634229" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAAFFA2FF2AFF51FABCFDA8" httpUri="http://treatment.plazi.org/id/4B0E6E2AFFAAFFA2FF2AFF51FABCFDA8" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224">
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<paragraph id="C318DF3CFFAAFFADFF2AFF51FD88FF0B" blockId="23.[136,620,158,185]" box="[136,620,158,185]" pageId="23" pageNumber="224">
<heading id="98506850FFAAFFADFF2AFF51FD88FF0B" box="[136,620,158,185]" fontSize="11" level="2" pageId="23" pageNumber="224" reason="3">
<taxonomicName id="04A7A4BFFFAAFFADFF2AFF51FE8FFF0A" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[136,363,158,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri" status="sp. nov.">Diploneis stoermeri</taxonomicName>
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<emphasis id="F1D3032EFFAAFFADFED3FF6FFE24FF0B" box="[369,448,160,185]" italics="true" pageId="23" pageNumber="224">sp. nov</emphasis>
.
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(
<figureCitation id="5B9CC3B9FFAAFFADFE75FF6FFD87FF0B" box="[471,611,160,185]" captionStart-0="FIGURES 8090" captionStart-1="FIGURES 9196" captionStart-2="FIGURES 97112" captionStartId-0="20.[136,243,1959,1980]" captionStartId-1="21.[136,243,1878,1899]" captionStartId-2="22.[136,243,1993,2014]" captionTargetBox-0="[174,1413,284,1930]" captionTargetBox-1="[155,1433,194,1849]" captionTargetBox-2="[155,1432,194,1964]" captionTargetId-0="figure-53@20.[170,1417,281,1934]" captionTargetId-1="figure-14@21.[151,1436,190,1853]" captionTargetId-2="figure-17@22.[151,1436,190,1967]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." captionText-1="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." captionText-2="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi-0="http://doi.org/10.5281/zenodo.13634158" figureDoi-1="http://doi.org/10.5281/zenodo.13634160" figureDoi-2="http://doi.org/10.5281/zenodo.13634162" httpUri-0="https://zenodo.org/record/13634158/files/figure.png" httpUri-1="https://zenodo.org/record/13634160/files/figure.png" httpUri-2="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">Figs 80112</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFFAAFFADFF2AFF2EFC6AFCCA" blockId="23.[136,1452,225,888]" pageId="23" pageNumber="224">
Valves are elliptical-lanceolate becoming almost circular with size reduction, with convex margins and broadly round ends (
<figureCitation id="5B9CC3B9FFAAFFADFF32FECEFEEEFEAA" box="[144,266,257,280]" captionStart-0="FIGURES 8090" captionStart-1="FIGURES 9196" captionStartId-0="20.[136,243,1959,1980]" captionStartId-1="21.[136,243,1878,1899]" captionTargetBox-0="[174,1413,284,1930]" captionTargetBox-1="[155,1433,194,1849]" captionTargetId-0="figure-53@20.[170,1417,281,1934]" captionTargetId-1="figure-14@21.[151,1436,190,1853]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." captionText-1="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi-0="http://doi.org/10.5281/zenodo.13634158" figureDoi-1="http://doi.org/10.5281/zenodo.13634160" httpUri-0="https://zenodo.org/record/13634158/files/figure.png" httpUri-1="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Figs 8091</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFEB6FECEFE7EFEAA" box="[276,410,257,280]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">97109, 111</figureCitation>
). The valve length is 19.544.5 μm, and the valve breadth is 12.522.0 μm. The axial area is linear to lanceolate, expanding towards the round-elongate to rectangular and slightly asymmetric central area. From inside, a thick linear silica plate covers the whole length of the longitudinal canal (
<figureCitation id="5B9CC3B9FFAAFFADFCECFE8EFC78FEEA" box="[846,924,321,344]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Fig. 93</figureCitation>
). The central area is round-elongate to rectangular and slightly asymmetric, 1.54.0 μm wide. Externally, the longitudinal canal is lanceolate to linear, expanded in the middle of the valve with two to three areolae that coalesce into one areola covered with large and complex cribra toward the valve apices (
<figureCitation id="5B9CC3B9FFAAFFADFF7AFE6EFECFFE0A" box="[216,299,417,440]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="23" pageNumber="224">Figs 90</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFE94FE6EFE19FE0A" box="[310,509,417,440]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">106, 107, 111, 112</figureCitation>
). The external openings of the canal are covered with cribrate occlusions similar to those occluding the striae from which they are separated by a thin hyaline area (
<figureCitation id="5B9CC3B9FFAAFFADFC1AFE0EFBE9FE6A" box="[952,1037,449,472]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="23" pageNumber="224">Figs 90</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFBB9FE0EFBDCFE6A" box="[1051,1080,449,472]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">91</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFBE7FE0EFB43FE6A" box="[1093,1191,449,472]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">109, 110</figureCitation>
); cribra are located in a depression slightly lower that the rest of the non-porous silica. Internally, the longitudinal canal is covered with silica plate throughout the whole length, forming a “trench” structure housing the raphe slits (
<figureCitation id="5B9CC3B9FFAAFFADFC58FDCEFBAEFDAA" box="[1018,1098,513,536]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Fig. 93</figureCitation>
). Externally, the raphe is straight and simple with drop-like proximal ends that are deflected to the same side of the valve and positioned within an expanded depression (
<figureCitation id="5B9CC3B9FFAAFFADFEABFD8EFEB8FDEA" box="[265,348,577,600]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="23" pageNumber="224">Figs 90</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFECAFD8EFE4AFDEA" box="[360,430,577,600]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">95, 96</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFE1BFD8EFDFEFDEA" box="[441,538,577,600]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">109, 110</figureCitation>
). Towards the central area the raphe branches are located in a slight “depression” below the surface of the rest of the non-porous silica (
<figureCitation id="5B9CC3B9FFAAFFADFD0CFDAEFCE5FDCA" box="[686,769,609,632]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="23" pageNumber="224">Figs 90</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFCAEFDAEFCCDFDCA" box="[780,809,609,632]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">95</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFC96FDAEFC74FDCA" box="[820,912,609,632]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">109, 111</figureCitation>
) and surrounded with a silica ridge (
<figureCitation id="5B9CC3B9FFAAFFADFAB5FDAEFA8EFDCA" box="[1303,1386,609,632]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Figs 96</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFAD7FDAEFA44FDCA" box="[1397,1440,609,632]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">110</figureCitation>
). Distally, the raphe branches are bent into a short drop-like terminal fissures (
<figureCitation id="5B9CC3B9FFAAFFADFC12FD4EFBCDFD2A" box="[944,1065,641,664]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Figs 91, 92</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFB96FD4EFB22FD2A" box="[1076,1222,641,664]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">109, 111, 112</figureCitation>
). Internally, the raphe is straight with simple proximal and distal ends, inserted in a slightly elevated sternum inside the “trench” formed by the longitudinal canal (
<figureCitation id="5B9CC3B9FFAAFFADFEF4FD0EFE42FD6A" box="[342,422,705,728]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Fig. 93</figureCitation>
). The proximal raphe ends are the only raphe structure that reaches the height of the longitudinal canal (
<figureCitation id="5B9CC3B9FFAAFFADFF72FD2EFEC5FD4A" box="[208,289,737,760]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Fig. 93</figureCitation>
). The striae are uniseriate, radiate,
<quantity id="045F72D9FFAAFFADFD3FFD2EFD19FD4A" box="[669,765,737,760]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="23" pageNumber="224" unit="in" value="13.5" valueMax="15.0" valueMin="12.0">1215 in</quantity>
10 μm, composed of round to rectangular areolae,
<quantity id="045F72D9FFAAFFADFA88FD2EFA6DFD4A" box="[1322,1417,737,760]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="23" pageNumber="224" unit="in" value="15.0" valueMax="20.0" valueMin="10.0">1020 in</quantity>
10 μm. In some valves there is a slight biseriate striae pattern toward the margins (
<figureCitation id="5B9CC3B9FFAAFFADFC72FCCEFBC6FCAA" box="[976,1058,769,792]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Figs 95</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFB8FFCCEFBBDFCAA" box="[1069,1113,769,792]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">102</figureCitation>
). Externally, the alveolate striae open through one row of areolae covered with cribrate occlusions. The cribrate occlusions increase in size and at the same time become more complex in structure toward the valve margins (
<figureCitation id="5B9CC3B9FFAAFFADFCC3FC8EFC51FCEA" box="[865,949,833,856]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="23" pageNumber="224">Figs 90</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFC60FC8EFBECFCEA" box="[962,1032,833,856]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">94, 95</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFBB7FC8EFB92FCEA" box="[1045,1142,833,856]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">109, 110</figureCitation>
). Internally, the alveoli open through a single linear opening covered with a fine silica layer (
<figureCitation id="5B9CC3B9FFAAFFADFC93FCAEFC65FCCA" box="[817,897,865,888]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Fig. 93</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFAAFFADFF2AFC65FE37FBFD" pageId="23" pageNumber="224" type="materials_examined">
<paragraph id="C318DF3CFFAAFFADFF2AFC65FE37FBFD" blockId="23.[136,1452,938,1103]" pageId="23" pageNumber="224">
<emphasis id="F1D3032EFFAAFFADFF2AFC65FF3EFC0D" bold="true" box="[136,218,938,959]" pageId="23" pageNumber="224">Type:—</emphasis>
<materialsCitation id="73CFD561FFAAFFADFF78FC65FBBFFC0D" box="[218,1115,938,959]" country="Mongolia" county="Lake" latitude="50.467533" location="Hatgal. Coordinates" municipality="National Park" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Hovsgol">
<collectingCountry id="BBB09FACFFAAFFADFF78FC65FEBBFC0D" box="[218,351,938,959]" name="Mongolia" pageId="23" pageNumber="224">MONGOLIA</collectingCountry>
,
<collectingCounty id="2A79A7B0FFAAFFADFEC9FC65FE7FFC0D" box="[363,411,938,959]" pageId="23" pageNumber="224">Lake</collectingCounty>
<collectingRegion id="016311DEFFAAFFADFE06FC65FE11FC0D" box="[420,501,938,959]" country="Mongolia" name="Hovsgol" pageId="23" pageNumber="224">Hövsgöl</collectingRegion>
(
<collectingRegion id="016311DEFFAAFFADFDA6FC65FDB3FC0D" box="[516,599,938,959]" country="Mongolia" name="Hovsgol" pageId="23" pageNumber="224">Hövsgöl</collectingRegion>
<collectingMunicipality id="237C4546FFAAFFADFDFDFC65FD0EFC0D" box="[607,746,938,959]" pageId="23" pageNumber="224">National Park</collectingMunicipality>
),
<location id="C67889E7FFAAFFADFD5EFC65FC22FC0D" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAAFFA2FF2AFF51FABCFDA8:C67889E7FFAAFFADFD5EFC65FC22FC0D" box="[764,966,938,959]" country="Mongolia" county="Lake" latitude="50.467533" municipality="National Park" name="Hatgal. Coordinates" pageId="23" pageNumber="224" stateProvince="Hovsgol">Hatgal. Coordinates</location>
:
<geoCoordinate id="A693B9FBFFAAFFADFC71FC65FBBFFC0D" box="[979,1115,938,959]" degrees="50" direction="north" minutes="28.052" orientation="latitude" pageId="23" pageNumber="224" precision="1" value="50.467533">50°28.052 N</geoCoordinate>
</materialsCitation>
<materialsCitation id="73CFD561FFAAFFADFBFEFC65FEBBFB99" collectingDate="1996-06-11" collectionCode="A, M, B, F, ANSP" collectorName="Mark &amp; Eugene &amp; Slide" country="Mongolia" county="Edlund Collection" elevation="1" location="Edlund" longitude="100.172264" municipality="Science Museum of Minnesota" pageId="23" pageNumber="224" specimenCode="M068, GC-36353" specimenCount="1" typeStatus="holotype">
;
<geoCoordinate id="A693B9FBFFAAFFADFBCBFC65FB1CFC0D" box="[1129,1272,938,959]" degrees="100" direction="east" minutes="10.336" orientation="longitude" pageId="23" pageNumber="224" precision="1" value="100.172264">100°10.336 E</geoCoordinate>
,
<taxonomicName id="04A7A4BFFFAAFFADFAA7FC65FAB2FC0D" authorityName="Kutzing" authorityYear="1833" box="[1285,1366,938,959]" class="Ulvophyceae" family="Ulotrichaceae" genus="Ulothrix" kingdom="Plantae" order="Ulotrichales" pageId="23" pageNumber="224" phylum="Chlorophyta" rank="genus">
<emphasis id="F1D3032EFFAAFFADFAA7FC65FAB2FC0D" box="[1285,1366,938,959]" italics="true" pageId="23" pageNumber="224">Ulothrix</emphasis>
</taxonomicName>
/epilithic from
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depth
</elevation>
(accession number:
<specimenCode id="93017747FFAAFFADFD9CFC01FD9DFC51" box="[574,633,974,995]" collectionCode="M" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="23" pageNumber="224" type="Herbarium">M068</specimenCode>
<collectionCode id="A5B647F9FFAAFFADFDDBFC01FD6FFC51" box="[633,651,974,995]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="23" pageNumber="224" type="Herbarium">A</collectionCode>
,
<collectionCode id="A5B647F9FFAAFFADFD35FC01FD54FC51" box="[663,688,974,995]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="23" pageNumber="224" type="Herbarium">M</collectionCode>
.
<collectionCode id="A5B647F9FFAAFFADFD16FC01FD21FC51" box="[692,709,974,995]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="23" pageNumber="224" type="Herbarium">B</collectionCode>
.
<collectingCounty id="2A79A7B0FFAAFFADFD70FC01FC6CFC51" box="[722,904,974,995]" pageId="23" pageNumber="224">Edlund Collection</collectingCounty>
,
<collectingMunicipality id="237C4546FFAAFFADFC36FC01FB2EFC51" box="[916,1226,974,995]" pageId="23" pageNumber="224">Science Museum of Minnesota</collectingMunicipality>
, collected by
<collectorName id="6E52BAEAFFAAFFADFAFAFC01FA68FC51" box="[1368,1420,974,995]" pageId="23" pageNumber="224">Mark</collectorName>
<collectionCode id="A5B647F9FFAAFFADFA37FC01FA43FC51" box="[1429,1447,974,995]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="23" pageNumber="224" type="Herbarium">B</collectionCode>
.
<location id="C67889E7FFAAFFADFF1FFC3DFEE1FBB5" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAAFFA2FF2AFF51FABCFDA8:C67889E7FFAAFFADFF1FFC3DFEE1FBB5" box="[189,261,1010,1031]" country="Mongolia" county="Edlund Collection" longitude="100.172264" municipality="Science Museum of Minnesota" name="Edlund" pageId="23" pageNumber="224">Edlund</location>
,
<collectorName id="6E52BAEAFFAAFFADFEB2FC3DFEBCFBB5" box="[272,344,1010,1031]" pageId="23" pageNumber="224">Eugene</collectorName>
<collectionCode id="A5B647F9FFAAFFADFEC2FC3DFE8AFBB5" box="[352,366,1010,1031]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="23" pageNumber="224" type="Herbarium">F</collectionCode>
.
<location id="C67889E7FFAAFFADFEDBFC3DFE37FBB5" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAAFFA2FF2AFF51FABCFDA8:C67889E7FFAAFFADFEDBFC3DFE37FBB5" box="[377,467,1010,1031]" country="Mongolia" county="Edlund Collection" longitude="100.172264" municipality="Science Museum of Minnesota" name="Stoermer" pageId="23" pageNumber="224">Stoermer</location>
, and
<location id="C67889E7FFAAFFADFDA5FC3DFD36FBB5" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAAFFA2FF2AFF51FABCFDA8:C67889E7FFAAFFADFDA5FC3DFD36FBB5" box="[519,722,1010,1031]" country="Mongolia" county="Edlund Collection" longitude="100.172264" municipality="Science Museum of Minnesota" name="Nergui Soninkhishig" pageId="23" pageNumber="224">Nergui Soninkhishig</location>
,
<date id="B719F9FCFFAAFFADFD7FFC3DFC84FBB5" box="[733,864,1010,1031]" pageId="23" pageNumber="224" value="1996-06-11">
<collectingDate id="A75D0014FFAAFFADFD7FFC3DFC84FBB5" box="[733,864,1010,1031]" pageId="23" pageNumber="224" value="1996-06-11">11 June 1996</collectingDate>
</date>
) (
<collectorName id="6E52BAEAFFAAFFADFCD7FC3DFC4CFBB5" box="[885,936,1010,1031]" pageId="23" pageNumber="224">Slide</collectorName>
<specimenCode id="93017747FFAAFFADFC0DFC3DFC0FFBB5" box="[943,1003,1010,1031]" collectionCode="M" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="23" pageNumber="224" type="Herbarium">M068</specimenCode>
<collectionCode id="A5B647F9FFAAFFADFC49FC3DFC19FBB5" box="[1003,1021,1010,1031]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="23" pageNumber="224" type="Herbarium">A</collectionCode>
,
<collectionCode id="A5B647F9FFAAFFADFBA4FC3DFBA6FBB5" box="[1030,1090,1010,1031]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="224">ANSP</collectionCode>
<specimenCode id="93017747FFAAFFADFBE8FC3DFB56FBB5" box="[1098,1202,1010,1031]" collectionCode="GC-" pageId="23" pageNumber="224">GC-36353</specimenCode>
, GCM-24055),
<emphasis id="F1D3032EFFAAFFADFAEFFC3DFEBBFB99" bold="true" pageId="23" pageNumber="224">
<typeStatus id="1C1C619EFFAAFFADFAEFFC3DFA42FBB5" box="[1357,1446,1010,1031]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
, designated here
</emphasis>
</materialsCitation>
; example specimens on
<figureCitation id="5B9CC3B9FFAAFFADFDEDFBD9FCEDFB99" box="[591,777,1046,1067]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="23" pageNumber="224">Figs 80, 84, 8789</figureCitation>
;
<materialsCitation id="73CFD561FFAAFFADFCB6FBD9FE34FBFD" collectionCode="CAS" collectorName="Slide" country="Mongolia" pageId="23" pageNumber="224" specimenCount="2" typeStatus="isotype">
(
<collectorName id="6E52BAEAFFAAFFADFCB8FBD9FCA9FB99" box="[794,845,1046,1067]" pageId="23" pageNumber="224">Slide</collectorName>
917068,
<collectionCode id="A5B647F9FFAAFFADFC0BFBD9FC3EFB99" box="[937,986,1046,1067]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="224" type="Museum">CAS</collectionCode>
,
<emphasis id="F1D3032EFFAAFFADFC47FBD9FB3FFB99" bold="true" box="[997,1243,1046,1067]" pageId="23" pageNumber="224">
<typeStatus id="1C1C619EFFAAFFADFC47FBD9FBCAFB99" box="[997,1070,1046,1067]" pageId="23" pageNumber="224" type="isotype">isotype</typeStatus>
, designated here
</emphasis>
;
<collectorName id="6E52BAEAFFAAFFADFB4BFBD9FAFEFB99" box="[1257,1306,1046,1067]" pageId="23" pageNumber="224">Slide</collectorName>
919072,
<collectionCode id="A5B647F9FFAAFFADFAD5FBD9FA4CFB99" box="[1399,1448,1046,1067]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="224" type="Museum">CAS</collectionCode>
,
<emphasis id="F1D3032EFFAAFFADFF1FFBF5FE22FBFD" bold="true" box="[189,454,1082,1103]" pageId="23" pageNumber="224">
<typeStatus id="1C1C619EFFAAFFADFF1FFBF5FEFEFBFD" box="[189,282,1082,1103]" pageId="23" pageNumber="224" type="paratype">paratype</typeStatus>
, designated here
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFAAFFA2FF2AFB4FFE04FE23" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224" type="etymology">
<paragraph id="C318DF3CFFAAFFADFF2AFB4FFB73FB0F" blockId="23.[136,1452,1152,2077]" pageId="23" pageNumber="224">
<emphasis id="F1D3032EFFAAFFADFF2AFB4FFECAFB2B" bold="true" box="[136,302,1152,1177]" pageId="23" pageNumber="224">Etymology:—</emphasis>
The species name honors Dr. Eugene Stoermer, University of
<collectingRegion id="016311DEFFAAFFADFBA6FB4FFB96FB2B" box="[1028,1138,1152,1177]" country="United States of America" name="Michigan" pageId="23" pageNumber="224">Michigan</collectingRegion>
, who initiated the research collaboration with the National University of Mongolia with a collecting expedition in 1996.
</paragraph>
<paragraph id="C318DF3CFFAAFFADFF1FFB07FD97F9DF" blockId="23.[136,1452,1152,2077]" pageId="23" pageNumber="224">
<emphasis id="F1D3032EFFAAFFADFF1FFB07FE81FB53" bold="true" box="[189,357,1224,1249]" pageId="23" pageNumber="224">Observations:</emphasis>
<taxonomicName id="04A7A4BFFFAAFFADFE23FB07FD8CFB53" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[385,616,1224,1249]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFAAFFADFE23FB07FD8CFB53" box="[385,616,1224,1249]" italics="true" pageId="23" pageNumber="224">Diploneis stoermeri</emphasis>
</taxonomicName>
appears to be morphologically variable in shape of the central area (rectangular and slightly asymmetric to round) and in the structure of the longitudinal canal (lanceolate, expanded in the middle of the valve to linear-lanceolate and slightly asymmetrical; compare
<figureCitation id="5B9CC3B9FFAAFFADFBE5FADFFB23FA9B" box="[1095,1223,1296,1321]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="23" pageNumber="224">Figs 8090</figureCitation>
with
<figureCitation id="5B9CC3B9FFAAFFADFAAFFADFFA44FA9B" box="[1293,1440,1296,1321]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">Figs 97109</figureCitation>
). Variability in other distinguishing features, such valve shape, the structure of the striae, and the density of striae and areolae was not observed, but based on the above noted variable structures, we consider the Lake
<collectingRegion id="016311DEFFAAFFADFB66FA97FAC7FAC3" box="[1220,1315,1368,1393]" country="Mongolia" name="Hovsgol" pageId="23" pageNumber="224">Hövsgöl</collectingRegion>
populations to be a morpholgical complex represented by at least two different phenodemes (
<figureCitation id="5B9CC3B9FFAAFFADFBBBFAB3FB7FFA27" box="[1049,1179,1404,1429]" captionStart-0="FIGURES 8090" captionStart-1="FIGURES 9196" captionStartId-0="20.[136,243,1959,1980]" captionStartId-1="21.[136,243,1878,1899]" captionTargetBox-0="[174,1413,284,1930]" captionTargetBox-1="[155,1433,194,1849]" captionTargetId-0="figure-53@20.[170,1417,281,1934]" captionTargetId-1="figure-14@21.[151,1436,190,1853]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." captionText-1="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi-0="http://doi.org/10.5281/zenodo.13634158" figureDoi-1="http://doi.org/10.5281/zenodo.13634160" httpUri-0="https://zenodo.org/record/13634158/files/figure.png" httpUri-1="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Figs 8096</figureCitation>
, morphotype 1 and
<figureCitation id="5B9CC3B9FFAAFFADFAD9FAB3FF3AFA0B" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="23" pageNumber="224">Figs 97112</figureCitation>
, morphotype 2). Sympatry among these phenodemes makes their separation even more difficult and opens further questions into their identity and phylogenetic relationship. Such morphological patterns might be correlated with /related to physiological, ecological and/or genetic traits. Therefore, additional analyses including paleolimnology, morphometry and genetic information could provide further support in revealing the patterns and mechanisms for such morphological variations within the
<taxonomicName id="04A7A4BFFFAAFFADFD80F9FFFD4BF9FB" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[546,687,1584,1609]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFAAFFADFD80F9FFFD4BF9FB" box="[546,687,1584,1609]" italics="true" pageId="23" pageNumber="224">D. stoermeri</emphasis>
</taxonomicName>
complex, or could simply explain the sympatric coexistence of two genotypically distinct entities (
<bibRefCitation id="A736A2CDFFAAFFADFE42F99BFD83F9DF" author="Mann, D. G." box="[480,615,1620,1645]" pageId="23" pageNumber="224" pagination="437 - 495" refId="ref27788" refString="Mann, D. G. (1999) The species concept in diatoms. Phycologia 38 (6): 437 - 495. http: // dx. doi. org / 10.2216 / i 0031 - 8884 - 38 - 6 - 437.1" type="journal article" year="1999">Mann 1999</bibRefCitation>
).
</paragraph>
<paragraph id="C318DF3CFFAAFFADFF1FF9B7FD9FF803" blockId="23.[136,1452,1152,2077]" pageId="23" pageNumber="224">
<taxonomicName id="04A7A4BFFFAAFFADFF1FF9B7FE79F923" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[189,413,1656,1681]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFAAFFADFF1FF9B7FE79F923" box="[189,413,1656,1681]" italics="true" pageId="23" pageNumber="224">Diploneis stoermeri</emphasis>
</taxonomicName>
can easily be confused with
<taxonomicName id="04A7A4BFFFAAFFADFD4AF9B7FC6EF923" box="[744,906,1656,1681]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAAFFADFD4AF9B7FD18F923" box="[744,764,1656,1681]" italics="true" pageId="23" pageNumber="224">D</emphasis>
.
<emphasis id="F1D3032EFFAAFFADFCAEF9B7FC6EF923" box="[780,906,1656,1681]" italics="true" pageId="23" pageNumber="224">paraparma</emphasis>
</taxonomicName>
. These two species share similar morphological features, such as the central area, the axial area, and the structure of the longitudinal canal, but differ in striae structure: uniseriate, rarely biseriate towards the valve margins in
<taxonomicName id="04A7A4BFFFAAFFADFCA8F90FFC7DF96B" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[778,921,1728,1753]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFAAFFADFCA8F90FFC7DF96B" box="[778,921,1728,1753]" italics="true" pageId="23" pageNumber="224">D. stoermeri</emphasis>
</taxonomicName>
and biseriate becoming uniseriate towards the axial area in
<taxonomicName id="04A7A4BFFFAAFFADFE83F92BFE21F94F" box="[289,453,1764,1789]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAAFFADFE83F92BFED1F94F" box="[289,309,1764,1789]" italics="true" pageId="23" pageNumber="224">D</emphasis>
.
<emphasis id="F1D3032EFFAAFFADFEE5F92BFE21F94F" box="[327,453,1764,1789]" italics="true" pageId="23" pageNumber="224">paraparma</emphasis>
</taxonomicName>
(compare
<figureCitation id="5B9CC3B9FFAAFFADFDE1F92BFD27F94F" box="[579,707,1764,1789]" captionStart-0="FIGURES 8090" captionStart-1="FIGURES 9196" captionStartId-0="20.[136,243,1959,1980]" captionStartId-1="21.[136,243,1878,1899]" captionTargetBox-0="[174,1413,284,1930]" captionTargetBox-1="[155,1433,194,1849]" captionTargetId-0="figure-53@20.[170,1417,281,1934]" captionTargetId-1="figure-14@21.[151,1436,190,1853]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." captionText-1="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi-0="http://doi.org/10.5281/zenodo.13634158" figureDoi-1="http://doi.org/10.5281/zenodo.13634160" httpUri-0="https://zenodo.org/record/13634158/files/figure.png" httpUri-1="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">Figs 8091</figureCitation>
with
<figureCitation id="5B9CC3B9FFAAFFADFCABF92BFC6BF94F" box="[777,911,1764,1789]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="23" pageNumber="224">Figs 7076</figureCitation>
). However, using the striae as a distinguishing feature might be a challenge due to the tendency of the cribrate occlusions to become complex in structure towards the valve margins (see
<figureCitation id="5B9CC3B9FFAAFFADFEFFF8E3FE3DF8F7" box="[349,473,1836,1861]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="23" pageNumber="224">Figs 76, 79</figureCitation>
,
<figureCitation id="5B9CC3B9FFAAFFADFE46F8E3FDE7F8F7" box="[484,515,1836,1861]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="23" pageNumber="224">95</figureCitation>
), which can leave the impression of a biseriate pattern. Hence the cribrate occlusions cover the alveolate striae, the number of areolae is difficult to observe, and sometimes can lead to misinterpretation. Therefore, when using the striae for identification, the uni- and biseriate pattern should be taken as a main feature in separating
<taxonomicName id="04A7A4BFFFAAFFADFEA0F857FE6BF803" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[258,399,1944,1969]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFAAFFADFEA0F857FEF2F803" box="[258,278,1944,1969]" italics="true" pageId="23" pageNumber="224">D</emphasis>
.
<emphasis id="F1D3032EFFAAFFADFE86F857FE6BF803" box="[292,399,1944,1969]" italics="true" pageId="23" pageNumber="224">stoermeri</emphasis>
</taxonomicName>
from
<taxonomicName id="04A7A4BFFFAAFFADFE76F857FD90F803" box="[468,628,1944,1969]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAAFFADFE76F857FE0CF803" box="[468,488,1944,1969]" italics="true" pageId="23" pageNumber="224">D</emphasis>
.
<emphasis id="F1D3032EFFAAFFADFE54F857FD90F803" box="[502,628,1944,1969]" italics="true" pageId="23" pageNumber="224">paraparma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C318DF3CFFAAFFA2FF1FF873FE04FE23" blockId="23.[136,1452,1152,2077]" lastBlockId="24.[136,1452,160,401]" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224">
<taxonomicName id="04A7A4BFFFAAFFADFF1FF873FE98F867" authorityName=", sensu Idei &amp; Kobayasi" authorityYear="1986" box="[189,380,1980,2005]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAAFFADFF1FF873FE98F867" box="[189,380,1980,2005]" italics="true" pageId="23" pageNumber="224">Diploneis parma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="04A7A4BFFFAAFFADFDB7F873FD40F867" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[533,676,1980,2005]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFAAFFADFDB7F873FDCDF867" box="[533,553,1980,2005]" italics="true" pageId="23" pageNumber="224">D</emphasis>
.
<emphasis id="F1D3032EFFAAFFADFD9BF873FD40F867" box="[569,676,1980,2005]" italics="true" pageId="23" pageNumber="224">stoermeri</emphasis>
</taxonomicName>
in: the valve outline (broadly elliptical
<emphasis id="F1D3032EFFAAFFADFBCEF873FB67F867" box="[1132,1155,1980,2005]" italics="true" pageId="23" pageNumber="224">vs</emphasis>
. elliptical-lanceolate); the central area (round
<emphasis id="F1D3032EFFAAFFADFEC8F82FFE65F84B" box="[362,385,2016,2041]" italics="true" pageId="23" pageNumber="224">vs</emphasis>
. rectangular and slightly asymmetric); and the striae structure (uniseriate becoming biseriate towards the valve margins
<emphasis id="F1D3032EFFAAFFADFE0CF7CBFE21F7AF" box="[430,453,2052,2077]" italics="true" pageId="23" pageNumber="224">vs</emphasis>
. uniseriate, rarely biseriate towards valve margins).
<taxonomicName id="04A7A4BFFFAAFFADFBA6F7CBFB05F7AF" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[1028,1249,2052,2077]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="23" pageNumber="224" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFAAFFADFBA6F7CBFB05F7AF" box="[1028,1249,2052,2077]" italics="true" pageId="23" pageNumber="224">Diploneis stoermeri</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="04A7A4BFFFA5FFA2FF2AFF6FFEF3FF0B" authorityName="Lange-Bertalot &amp; Reichardt" authorityYear="2000" box="[136,279,160,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="24" pageNumber="225" phylum="Ochrophyta" rank="species" species="krammeri">
<emphasis id="F1D3032EFFA5FFA2FF2AFF6FFEF3FF0B" box="[136,279,160,185]" italics="true" pageId="24" pageNumber="225">D. krammeri</emphasis>
</taxonomicName>
, though the stria density separates them [
<quantity id="045F72D9FFA5FFA2FD43FF6FFCA1FF0B" box="[737,837,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="24" pageNumber="225" unit="in" value="13.5" valueMax="15.0" valueMin="12.0">1215 in</quantity>
10 μm
<emphasis id="F1D3032EFFA5FFA2FC38FF6FFC55FF0B" box="[922,945,160,185]" italics="true" pageId="24" pageNumber="225">vs</emphasis>
. 11(12) in 10 μm]. Additionally, the structure of the longitudinal canal further supports the differences between
<taxonomicName id="04A7A4BFFFA5FFA2FCCEFF0BFC18FF6F" authorityName="Lange-Bertalot &amp; Reichardt" authorityYear="2000" box="[876,1020,196,221]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="24" pageNumber="225" phylum="Ochrophyta" rank="species" species="krammeri">
<emphasis id="F1D3032EFFA5FFA2FCCEFF0BFC18FF6F" box="[876,1020,196,221]" italics="true" pageId="24" pageNumber="225">D. krammeri</emphasis>
</taxonomicName>
and
<taxonomicName id="04A7A4BFFFA5FFA2FB90FF0BFB24FF6F" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[1074,1216,196,221]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="24" pageNumber="225" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFA5FFA2FB90FF0BFB24FF6F" box="[1074,1216,196,221]" italics="true" pageId="24" pageNumber="225">D. stoermeri</emphasis>
</taxonomicName>
(one to two rows of areolae in the middle of the valve narrowing into one at the valve apices
<emphasis id="F1D3032EFFA5FFA2FC60FF27FC3DFEB3" box="[962,985,232,257]" italics="true" pageId="24" pageNumber="225">vs</emphasis>
. two rows of areolae continuing into one that is covered with a large cribrum). Moreover, the extended depression where the proximal raphe ends are placed (
<figureCitation id="5B9CC3B9FFA5FFA2FF32FEFFFF03FEFB" box="[144,231,304,329]" captionStart="FIGURES 8090" captionStartId="20.[136,243,1959,1980]" captionTargetBox="[174,1413,284,1930]" captionTargetId="figure-53@20.[170,1417,281,1934]" captionTargetPageId="20" captionText="FIGURES 8090. Diploneis stoermeri morphotype 1, LMs and SEM external valve view. Figs 8089. LM valve views. Figs 80, 84, 8789. Specimens from the holotype slide, M068A. Fig. 90. External valve view. Scale bars = 10 μm (Figs 8089); 5 μm (Fig. 90)." figureDoi="http://doi.org/10.5281/zenodo.13634158" httpUri="https://zenodo.org/record/13634158/files/figure.png" pageId="24" pageNumber="225">Figs 90</figureCitation>
,
<figureCitation id="5B9CC3B9FFA5FFA2FF50FEFFFEF4FEFB" box="[242,272,304,329]" captionStart="FIGURES 9196" captionStartId="21.[136,243,1878,1899]" captionTargetBox="[155,1433,194,1849]" captionTargetId="figure-14@21.[151,1436,190,1853]" captionTargetPageId="21" captionText="FIGURES 9196. Diploneis stoermeri morphotype 1, SEM external and internal valve views. Fig. 91. External view of the whole valve. Fig. 92. External view of the distal raphe endings with short terminal fissure. Fig. 93. Internal view of the entire valve. Fig. 94. Close view of the external cribrate occlusions. Fig. 95. Detailed external view of the mid valve. Fig. 96. External view of the proximal raphe endings, bent to the same side of the valve. Scale bars = 5 μm (Figs 91, 93, 95); 2 μm (Figs 92, 94, 96)." figureDoi="http://doi.org/10.5281/zenodo.13634160" httpUri="https://zenodo.org/record/13634160/files/figure.png" pageId="24" pageNumber="225">96</figureCitation>
,
<figureCitation id="5B9CC3B9FFA5FFA2FEBEFEFFFEA1FEFB" box="[284,325,304,329]" captionStart="FIGURES 97112" captionStartId="22.[136,243,1993,2014]" captionTargetBox="[155,1432,194,1964]" captionTargetId="figure-17@22.[151,1436,190,1967]" captionTargetPageId="22" captionText="FIGURES 97112. Diploneis stoermeri morphotype 2, LM and SEM external valve views. Figs 97108. LM valve views. Figs 109, 111. External view of the whole valve. Fig. 110. External view of the proximal raphe endings. Fig. 112. Distal raphe endings. Scale bars = 10 μm (Figs 97108, 111); 5 μm (Figs 109, 112); 2 μm (Fig. 110)." figureDoi="http://doi.org/10.5281/zenodo.13634162" httpUri="https://zenodo.org/record/13634162/files/figure.png" pageId="24" pageNumber="225">111</figureCitation>
) is barely present in
<taxonomicName id="04A7A4BFFFA5FFA2FD8CFEFFFD59FEFB" authorityName="Lange-Bertalot &amp; Reichardt" authorityYear="2000" box="[558,701,304,329]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="24" pageNumber="225" phylum="Ochrophyta" rank="species" species="krammeri">
<emphasis id="F1D3032EFFA5FFA2FD8CFEFFFD59FEFB" box="[558,701,304,329]" italics="true" pageId="24" pageNumber="225">D. krammeri</emphasis>
</taxonomicName>
(
<bibRefCitation id="A736A2CDFFA5FFA2FD68FEFFFBADFEFB" author="Lange-Bertalot, H. &amp; Reichardt, E." box="[714,1097,304,329]" pageId="24" pageNumber="225" pagination="650 - 669" refId="ref27660" refString="Lange-Bertalot, H. &amp; Reichardt, E. (2000) Diploneis ovalis sensu stricto und Diploneis krammeri nov. spec. Revision des aktuellen Konzepts von Diploneis ovalis (Hilse) Cleve. Iconographia Diatomologica 9: 650 - 669." type="journal article" year="2000">Lange-Bertalot &amp; Reichardt 2000</bibRefCitation>
; this study,
<figureCitation id="5B9CC3B9FFA5FFA2FB69FEFFFA7BFEFB" box="[1227,1439,304,329]" captionStart="FIGURES 129134" captionStartId="25.[136,243,1884,1905]" captionTargetBox="[155,1433,194,1855]" captionTargetId="figure-14@25.[151,1436,190,1859]" captionTargetPageId="25" captionText="FIGURES 129134. Diploneis krammeri, SEM external valve views. Fig. 129. View of the whole valve. Fig. 130. Proximal raphe endings. Fig. 131. View of the whole frustule. Fig. 132. Showing the structure of the striae at the valve mantle. Fig. 133. Detailed view of the valve central area. Fig. 134. Distal raphe endings with short deflected terminal fissures. Scale bars = 5 μm (Figs 129, 131); 2 μm (Figs 130, 132134)." figureDoi="http://doi.org/10.5281/zenodo.13634166" httpUri="https://zenodo.org/record/13634166/files/figure.png" pageId="24" pageNumber="225">Figs 129, 130, 133</figureCitation>
).
<taxonomicName id="04A7A4BFFFA5FFA2FF2AFE9BFEBDFEDF" authorityName="Cleve" authorityYear="1894" baseAuthorityName="Kutzing" box="[136,345,340,365]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="24" pageNumber="225" phylum="Ochrophyta" rank="species" species="elliptica">
<emphasis id="F1D3032EFFA5FFA2FF2AFE9BFEBDFEDF" box="[136,345,340,365]" italics="true" pageId="24" pageNumber="225">Diploneis elliptica</emphasis>
</taxonomicName>
is another morphologically similar taxon, differing in stria density (811/10 μm in
<taxonomicName id="04A7A4BFFFA5FFA2FAA5FE9BFA62FEDF" authorityName="Cleve" authorityYear="1894" baseAuthorityName="Kutzing" box="[1287,1414,340,365]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="24" pageNumber="225" phylum="Ochrophyta" rank="species" species="elliptica">
<emphasis id="F1D3032EFFA5FFA2FAA5FE9BFAFFFEDF" box="[1287,1307,340,365]" italics="true" pageId="24" pageNumber="225">D</emphasis>
.
<emphasis id="F1D3032EFFA5FFA2FA88FE9BFA62FEDF" box="[1322,1414,340,365]" italics="true" pageId="24" pageNumber="225">elliptica</emphasis>
</taxonomicName>
<emphasis id="F1D3032EFFA5FFA2FA2FFE9BFA40FEDF" box="[1421,1444,340,365]" italics="true" pageId="24" pageNumber="225">vs</emphasis>
. 1215/10 μm in
<taxonomicName id="04A7A4BFFFA5FFA2FEE1FEB7FE34FE23" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[323,464,376,401]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="24" pageNumber="225" phylum="Ochrophyta" rank="species" species="stoermeri">
<emphasis id="F1D3032EFFA5FFA2FEE1FEB7FEB3FE23" box="[323,343,376,401]" italics="true" pageId="24" pageNumber="225">D</emphasis>
.
<emphasis id="F1D3032EFFA5FFA2FEC7FEB7FE34FE23" box="[357,464,376,401]" italics="true" pageId="24" pageNumber="225">stoermeri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFA5FFA2FF1FFE76FABCFDA8" pageId="24" pageNumber="225" type="biology_ecology">
<paragraph id="C318DF3CFFA5FFA2FF1FFE76FABCFDA8" blockId="24.[136,1452,441,538]" pageId="24" pageNumber="225">
<emphasis id="F1D3032EFFA5FFA2FF1FFE76FE12FE60" bold="true" box="[189,502,441,466]" pageId="24" pageNumber="225">Ecology and Distribution:</emphasis>
—M063A, M068A; M077A; M247A; M248A; M272A; M273A; M274A; M276A; M280A; M284A; M287A; M289A; M291A; M329A; M351A: distributed in shallow to deep waters in the northern and southern areas of Lake
<collectingRegion id="016311DEFFA5FFA2FE1FFDCEFDF8FDA8" box="[445,540,513,538]" country="Mongolia" name="Hovsgol" pageId="24" pageNumber="225">Hövsgöl</collectingRegion>
on epipelon, epiphyton, epilithon, marl accretions, and sediment habitats.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,125 @@
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<mods:title id="15D5EFA80ECE5FF9BDB43B270F81AB24">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<paragraph id="C318DF3CFFACFFABFF2AF8FEFD49F8FE" blockId="17.[136,685,1841,1868]" box="[136,685,1841,1868]" pageId="17" pageNumber="218">
<heading id="98506850FFACFFABFF2AF8FEFD49F8FE" box="[136,685,1841,1868]" fontSize="11" level="2" pageId="17" pageNumber="218" reason="3">
<taxonomicName id="04A7A4BFFFACFFABFF2AF8FEFDF1F8FE" authority="(Kutzing) Cleve" authorityName="Cleve" authorityYear="1894" baseAuthorityName="Kutzing" box="[136,533,1841,1868]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="17" pageNumber="218" phylum="Ochrophyta" rank="species" species="elliptica">
<emphasis id="F1D3032EFFACFFABFF2AF8FEFEBEF8FE" bold="true" box="[136,346,1841,1868]" italics="true" pageId="17" pageNumber="218">Diploneis elliptica</emphasis>
(Kützing) Cleve
</taxonomicName>
(
<figureCitation id="5B9CC3B9FFACFFABFD86F8FCFD41F8FE" box="[548,677,1843,1868]" captionStart-0="FIGURES 5464" captionStart-1="FIGURES 6569" captionStartId-0="16.[136,243,1993,2014]" captionStartId-1="17.[136,243,1702,1723]" captionTargetBox-0="[208,1380,500,1964]" captionTargetBox-1="[154,1434,191,1676]" captionTargetId-0="figure-156@16.[204,1383,497,1967]" captionTargetId-1="figure-14@17.[153,1434,190,1677]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 5464. Diploneis elliptica, LM and SEM internal valve views. Figs 5461. LM valve views. Figs 62, 63. Internal views of the entire valve. Fig. 64. Internal view of the central area with the proximal raphe endings. Scale bars = 10 μm (Figs 5463); 2 μm (Fig. 64)." captionText-1="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi-0="http://doi.org/10.5281/zenodo.13634152" figureDoi-1="http://doi.org/10.5281/zenodo.13634154" httpUri-0="https://zenodo.org/record/13634152/files/figure.png" httpUri-1="https://zenodo.org/record/13634154/files/figure.png" pageId="17" pageNumber="218">Figs 5469</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFFACFFA8FF2AF8BBFDB1FE65" blockId="17.[136,1452,1908,2091]" lastBlockId="18.[136,1452,160,471]" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218">
Valves are elliptical with convex margins and rounded ends (
<figureCitation id="5B9CC3B9FFACFFABFCBAF8BBFC72F839" box="[792,918,1908,1931]" captionStart="FIGURES 5464" captionStartId="16.[136,243,1993,2014]" captionTargetBox="[208,1380,500,1964]" captionTargetId="figure-156@16.[204,1383,497,1967]" captionTargetPageId="16" captionText="FIGURES 5464. Diploneis elliptica, LM and SEM internal valve views. Figs 5461. LM valve views. Figs 62, 63. Internal views of the entire valve. Fig. 64. Internal view of the central area with the proximal raphe endings. Scale bars = 10 μm (Figs 5463); 2 μm (Fig. 64)." figureDoi="http://doi.org/10.5281/zenodo.13634152" httpUri="https://zenodo.org/record/13634152/files/figure.png" pageId="17" pageNumber="218">Figs 5461</figureCitation>
). The valve length is 21.040.0 μm and the valve breadth is 13.021.0 μm. The axial area is linear, slightly expanding towards the central area. The central area is round to slightly transapically elongate, 2.55.0 μm wide. From inside a thick lanceolate to rhombic silica plate covers the whole length of the longitudinal canal (
<figureCitation id="5B9CC3B9FFACFFABFE43F81BFDB8F859" box="[481,604,2004,2027]" captionStart="FIGURES 5464" captionStartId="16.[136,243,1993,2014]" captionTargetBox="[208,1380,500,1964]" captionTargetId="figure-156@16.[204,1383,497,1967]" captionTargetPageId="16" captionText="FIGURES 5464. Diploneis elliptica, LM and SEM internal valve views. Figs 5461. LM valve views. Figs 62, 63. Internal views of the entire valve. Fig. 64. Internal view of the central area with the proximal raphe endings. Scale bars = 10 μm (Figs 5463); 2 μm (Fig. 64)." figureDoi="http://doi.org/10.5281/zenodo.13634152" httpUri="https://zenodo.org/record/13634152/files/figure.png" pageId="17" pageNumber="218">Figs 63, 62</figureCitation>
). Externally, the longitudinal canal is linear and slightly expanded in the middle with one, rarely two rows of areolae (
<figureCitation id="5B9CC3B9FFACFFABFD8AF83BFD37F7B9" box="[552,723,2036,2059]" captionStart="FIGURES 6569" captionStartId="17.[136,243,1702,1723]" captionTargetBox="[154,1434,191,1676]" captionTargetId="figure-14@17.[153,1434,190,1677]" captionTargetPageId="17" captionText="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi="http://doi.org/10.5281/zenodo.13634154" httpUri="https://zenodo.org/record/13634154/files/figure.png" pageId="17" pageNumber="218">Figs 65, 66, 68</figureCitation>
). The external openings of the longitudinal canal are covered with cribra similar to those of the striae, from which they are separated with a thick hyaline area (
<figureCitation id="5B9CC3B9FFACFFABFBFDF7DBFAE7F799" box="[1119,1283,2068,2091]" captionStart="FIGURES 6569" captionStartId="17.[136,243,1702,1723]" captionTargetBox="[154,1434,191,1676]" captionTargetId="figure-14@17.[153,1434,190,1677]" captionTargetPageId="17" captionText="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi="http://doi.org/10.5281/zenodo.13634154" httpUri="https://zenodo.org/record/13634154/files/figure.png" pageId="17" pageNumber="218">Figs 65, 66, 68</figureCitation>
). Externally the raphe is straight with expanded proximal ends, bent to the same side of the valve and positioned within a small depression (
<figureCitation id="5B9CC3B9FFAFFFA8FF32FF0FFEEEFF65" box="[144,266,192,215]" captionStart="FIGURES 6569" captionStartId="17.[136,243,1702,1723]" captionTargetBox="[154,1434,191,1676]" captionTargetId="figure-14@17.[153,1434,190,1677]" captionTargetPageId="17" captionText="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi="http://doi.org/10.5281/zenodo.13634154" httpUri="https://zenodo.org/record/13634154/files/figure.png" pageId="18" pageNumber="219">Figs 67, 68</figureCitation>
). Distally, the raphe branches end with short terminal fissure some distance from the valve mantle (
<figureCitation id="5B9CC3B9FFAFFFA8FA86FF0FFA7BFF65" box="[1316,1439,192,215]" captionStart="FIGURES 6569" captionStartId="17.[136,243,1702,1723]" captionTargetBox="[154,1434,191,1676]" captionTargetId="figure-14@17.[153,1434,190,1677]" captionTargetPageId="17" captionText="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi="http://doi.org/10.5281/zenodo.13634154" httpUri="https://zenodo.org/record/13634154/files/figure.png" pageId="18" pageNumber="219">Figs 65, 69</figureCitation>
). From inside the raphe is straight, placed in the “depression” formed by the longitudinal canal (
<figureCitation id="5B9CC3B9FFAFFFA8FB2AFF2FFAE3FF45" box="[1160,1287,224,247]" captionStart="FIGURES 5464" captionStartId="16.[136,243,1993,2014]" captionTargetBox="[208,1380,500,1964]" captionTargetId="figure-156@16.[204,1383,497,1967]" captionTargetPageId="16" captionText="FIGURES 5464. Diploneis elliptica, LM and SEM internal valve views. Figs 5461. LM valve views. Figs 62, 63. Internal views of the entire valve. Fig. 64. Internal view of the central area with the proximal raphe endings. Scale bars = 10 μm (Figs 5463); 2 μm (Fig. 64)." figureDoi="http://doi.org/10.5281/zenodo.13634152" httpUri="https://zenodo.org/record/13634152/files/figure.png" pageId="18" pageNumber="219">Figs 62, 63</figureCitation>
). The proximal ends reach the height of the canal itself (
<figureCitation id="5B9CC3B9FFAFFFA8FD97FECFFD54FEA5" box="[565,688,256,279]" captionStart="FIGURES 5464" captionStartId="16.[136,243,1993,2014]" captionTargetBox="[208,1380,500,1964]" captionTargetId="figure-156@16.[204,1383,497,1967]" captionTargetPageId="16" captionText="FIGURES 5464. Diploneis elliptica, LM and SEM internal valve views. Figs 5461. LM valve views. Figs 62, 63. Internal views of the entire valve. Fig. 64. Internal view of the central area with the proximal raphe endings. Scale bars = 10 μm (Figs 5463); 2 μm (Fig. 64)." figureDoi="http://doi.org/10.5281/zenodo.13634152" httpUri="https://zenodo.org/record/13634152/files/figure.png" pageId="18" pageNumber="219">Figs 62, 64</figureCitation>
), whereas the distal raphe endings are slightly elevated but not reaching the level of the canal (
<figureCitation id="5B9CC3B9FFAFFFA8FED5FEEFFE10FE85" box="[375,500,288,311]" captionStart="FIGURES 5464" captionStartId="16.[136,243,1993,2014]" captionTargetBox="[208,1380,500,1964]" captionTargetId="figure-156@16.[204,1383,497,1967]" captionTargetPageId="16" captionText="FIGURES 5464. Diploneis elliptica, LM and SEM internal valve views. Figs 5461. LM valve views. Figs 62, 63. Internal views of the entire valve. Fig. 64. Internal view of the central area with the proximal raphe endings. Scale bars = 10 μm (Figs 5463); 2 μm (Fig. 64)." figureDoi="http://doi.org/10.5281/zenodo.13634152" httpUri="https://zenodo.org/record/13634152/files/figure.png" pageId="18" pageNumber="219">Figs 62, 63</figureCitation>
). The striae are radiate in mid-valve becoming strongly radiate towards the valve apices,
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10 μm. Externally the alveolate striae are composed of large rectangular to round areolae,
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10 μm. Striae are uniseriate, becoming biseriate towards the valve apices and the valve mantle (black arrow on
<figureCitation id="5B9CC3B9FFAFFFA8FB00FEAFFB16FEC5" box="[1186,1266,352,375]" captionStart="FIGURES 6569" captionStartId="17.[136,243,1702,1723]" captionTargetBox="[154,1434,191,1676]" captionTargetId="figure-14@17.[153,1434,190,1677]" captionTargetPageId="17" captionText="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi="http://doi.org/10.5281/zenodo.13634154" httpUri="https://zenodo.org/record/13634154/files/figure.png" pageId="18" pageNumber="219">Fig. 69</figureCitation>
). The areolae are occluded by cribra, which are slightly lower than the rest of the non-porous valve surface (
<figureCitation id="5B9CC3B9FFAFFFA8FBC3FE4FFAEDFE25" box="[1121,1289,384,407]" captionStart="FIGURES 6569" captionStartId="17.[136,243,1702,1723]" captionTargetBox="[154,1434,191,1676]" captionTargetId="figure-14@17.[153,1434,190,1677]" captionTargetPageId="17" captionText="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi="http://doi.org/10.5281/zenodo.13634154" httpUri="https://zenodo.org/record/13634154/files/figure.png" pageId="18" pageNumber="219">Figs 65, 67, 68</figureCitation>
). Such cribrate depressions are not present at the valve apices (see arrow on
<figureCitation id="5B9CC3B9FFAFFFA8FCB1FE6FFC86FE05" box="[787,866,416,439]" captionStart="FIGURES 6569" captionStartId="17.[136,243,1702,1723]" captionTargetBox="[154,1434,191,1676]" captionTargetId="figure-14@17.[153,1434,190,1677]" captionTargetPageId="17" captionText="FIGURES 6569. Diploneis elliptica, SEM external valve views. Figs 65, 67. View of the whole valve. Fig. 66. Close view of the cribrate occlusions. Fig. 68. Valve central area with proximal raphe endings. Fig. 69. Distal raphe endings with terminal raphe fissures. Areolae biseriate formation (black arrow on the cribra structure at the valve apices). Scale bars = 10 μm (Figs 65, 67); 2 μm (Figs 66, 68, 69)." figureDoi="http://doi.org/10.5281/zenodo.13634154" httpUri="https://zenodo.org/record/13634154/files/figure.png" pageId="18" pageNumber="219">Fig. 69</figureCitation>
). Internally the alveoli open via a wide single opening, covered with fine silica layer (
<figureCitation id="5B9CC3B9FFAFFFA8FE6EFE0FFDADFE65" box="[460,585,448,471]" captionStart="FIGURES 5464" captionStartId="16.[136,243,1993,2014]" captionTargetBox="[208,1380,500,1964]" captionTargetId="figure-156@16.[204,1383,497,1967]" captionTargetPageId="16" captionText="FIGURES 5464. Diploneis elliptica, LM and SEM internal valve views. Figs 5461. LM valve views. Figs 62, 63. Internal views of the entire valve. Fig. 64. Internal view of the central area with the proximal raphe endings. Scale bars = 10 μm (Figs 5463); 2 μm (Fig. 64)." figureDoi="http://doi.org/10.5281/zenodo.13634152" httpUri="https://zenodo.org/record/13634152/files/figure.png" pageId="18" pageNumber="219">Figs 6264</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFAFFFA8FF2AFDC8FCA1FDDA" pageId="18" pageNumber="219" type="biology_ecology">
<paragraph id="C318DF3CFFAFFFA8FF2AFDC8FCA1FDDA" blockId="18.[136,1452,519,616]" pageId="18" pageNumber="219">
<emphasis id="F1D3032EFFAFFFA8FF2AFDC8FE5CFD92" bold="true" box="[136,440,519,544]" pageId="18" pageNumber="219">Ecology and Distribution:</emphasis>
—M063A; M247A; M248A; M262A; M272A; M273A; M274A; M280A; M284A; M287A; M288A; M289A; M290A; M291A; M329A;
<materialsCitation id="73CFD561FFAFFFA8FD22FDE4FCDAFDDA" location="Lake" pageId="18" pageNumber="219" specimenCount="1" stateProvince="Hovsgol">
M330A: widely distributed throughout
<location id="C67889E7FFAFFFA8FB97FDE4FB89FDF6" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFACFFA8FF2AF8FEFCA1FDDA:C67889E7FFAFFFA8FB97FDE4FB89FDF6" box="[1077,1133,555,580]" name="Lake" pageId="18" pageNumber="219" stateProvince="Hovsgol">Lake</location>
<collectingRegion id="016311DEFFAFFFA8FBD0FDE4FB35FDF6" box="[1138,1233,555,580]" country="Mongolia" name="Hovsgol" pageId="18" pageNumber="219">Hövsgöl</collectingRegion>
on habitats ranging from sand, epipelon, sediments, marl, and epiphytic
<collectedFrom id="70BB9CC1FFAFFFA8FD71FD80FCDAFDDA" box="[723,830,591,616]" pageId="18" pageNumber="219">
on
<taxonomicName id="04A7A4BFFFAFFFA8FD54FD80FCDAFDDA" box="[758,830,591,616]" class="Insecta" family="Noctuidae" genus="Chara" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="F1D3032EFFAFFFA8FD54FD80FCDAFDDA" box="[758,830,591,616]" italics="true" pageId="18" pageNumber="219">Chara</emphasis>
</taxonomicName>
</collectedFrom>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,332 @@
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<mods:title id="11935759CF69A17B1C384E9648835ACF">The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia</mods:title>
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<paragraph id="C318DF3CFFAFFFA8FF2AFD5AFD96FD02" blockId="18.[136,626,661,688]" box="[136,626,661,688]" pageId="18" pageNumber="219">
<heading id="98506850FFAFFFA8FF2AFD5AFD96FD02" box="[136,626,661,688]" fontSize="11" level="2" pageId="18" pageNumber="219" reason="3">
<taxonomicName id="04A7A4BFFFAFFFA8FF2AFD5AFE9AFD02" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[136,382,661,688]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="paraparma" status="sp. nov.">Diploneis paraparma</taxonomicName>
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<emphasis id="F1D3032EFFAFFFA8FE26FD58FE37FD02" box="[388,467,663,688]" italics="true" pageId="18" pageNumber="219">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="5B9CC3B9FFAFFFA8FE4BFD58FD8EFD02" box="[489,618,663,688]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Figs 7079</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFFAFFFA8FF2AFD17FB79FB5D" blockId="18.[136,1452,728,1263]" pageId="18" pageNumber="219">
Valves are elliptic to linear-elliptic with convex margins and rounded ends (
<figureCitation id="5B9CC3B9FFAFFFA8FC17FD17FBBEFD5D" box="[949,1114,728,751]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Figs 7076, 79</figureCitation>
). The valve length is 19.542.0 μm and the valve breadth 12.516.5 μm. The axial area is narrow, linear to lanceolate and slightly expanding into a lanceolate and weakly asymmetric central area. From inside a thick silica plate covers the whole length of the longitudinal canal (
<figureCitation id="5B9CC3B9FFAFFFA8FF32FCF7FF05FCFD" box="[144,225,824,847]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 78</figureCitation>
). The central area is lanceolate and slightly asymmetrical, 1.53.0 μm wide. Externally, the longitudinal canal appears narrow, lanceolate to linear and slightly expanded in the middle of the valve. Two rows of areolae open externally, slightly widening into one row of larger areolae towards the valve apices (
<figureCitation id="5B9CC3B9FFAFFFA8FC05FCB7FBC1FC3D" box="[935,1061,888,911]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Figs 76, 79</figureCitation>
). The external openings of the canal are occluded with cribra (
<figureCitation id="5B9CC3B9FFAFFFA8FE37FC57FDD2FC1D" box="[405,566,920,943]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Figs 76, 77, 79</figureCitation>
), which are open in a depression slightly lower that the rest of the non-porous silica. From inside the longitudinal canal is covered by a thick silica plate (
<figureCitation id="5B9CC3B9FFAFFFA8FCCEFC77FC59FC7D" box="[876,957,952,975]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 78</figureCitation>
). The heavily siliceous plate forms a “trench” along the whole length of the valve in which the raphe is situated (
<figureCitation id="5B9CC3B9FFAFFFA8FCFFFC17FC4AFC5D" box="[861,942,984,1007]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 78</figureCitation>
). From outside the raphe is straight and simple with drop-like proximal ends that are bent to the same side of the valve, positioned with an expanded depression (
<figureCitation id="5B9CC3B9FFAFFFA8FAEDFC37FA7BFBBD" box="[1359,1439,1016,1039]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 77</figureCitation>
). Distally, the raphe branches are bent to the same side of the valve into drop-like short terminal fissures (
<figureCitation id="5B9CC3B9FFAFFFA8FB52FBD7FA8BFB9D" box="[1264,1391,1048,1071]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Figs 76, 79</figureCitation>
). The striae are parallel in mid-valve, becoming radiate towards the distal ends of the valve,
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10 μm. Striae are biseriate on the mantle and valve margins, alternately positioned (arrow on
<figureCitation id="5B9CC3B9FFAFFFA8FCE8FB97FC72FBDD" box="[842,918,1112,1135]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 78</figureCitation>
) and becoming uniseriate toward the axial area. In some valves the biseriate pattern is present throughout the whole striae length (
<figureCitation id="5B9CC3B9FFAFFFA8FC6CFBB7FBADFB3D" box="[974,1097,1144,1167]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Figs 71, 75</figureCitation>
). Each stria is composed of small round to elliptical areolae,
<quantity id="045F72D9FFAFFFA8FE08FB57FDEEFB1D" box="[426,522,1176,1199]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="5.08" metricValueMin="3.81" pageId="18" pageNumber="219" unit="in" value="17.5" valueMax="20.0" valueMin="15.0">1520 in</quantity>
10 μm. Externally, the areolae are covered with cribrate occlusions, increasing in size towards the valve margins (
<figureCitation id="5B9CC3B9FFAFFFA8FE0CFB77FDCEFB7D" box="[430,554,1208,1231]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Figs 76, 79</figureCitation>
). Internally, the alveoli open via a single elongate opening covered with a fine silica layer (
<figureCitation id="5B9CC3B9FFAFFFA8FF6EFB17FEF8FB5D" box="[204,284,1240,1263]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 78</figureCitation>
). The structure of the alveolus can be seen through the fine siliceous layer (
<figureCitation id="5B9CC3B9FFAFFFA8FBE3FB17FB75FB5D" box="[1089,1169,1240,1263]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFAFFFA8FF2AFAEEFBBFFA10" pageId="18" pageNumber="219" type="materials_examined">
<paragraph id="C318DF3CFFAFFFA8FF2AFAEEFBBFFA10" blockId="18.[136,1452,1313,1442]" pageId="18" pageNumber="219">
<emphasis id="F1D3032EFFAFFFA8FF2AFAEEFF3EFA84" bold="true" box="[136,218,1313,1334]" pageId="18" pageNumber="219">Type:—</emphasis>
<materialsCitation id="73CFD561FFAFFFA8FF78FAEEFAF9FA84" box="[218,1309,1313,1334]" country="Mongolia" county="Lake" latitude="50.4284" location="Hatgal. Coordinates" municipality="National Park" pageId="18" pageNumber="219" specimenCount="1" stateProvince="Hovsgol">
<collectingCountry id="BBB09FACFFAFFFA8FF78FAEEFEBBFA84" box="[218,351,1313,1334]" name="Mongolia" pageId="18" pageNumber="219">MONGOLIA</collectingCountry>
,
<collectingCounty id="2A79A7B0FFAFFFA8FECBFAEEFE7DFA84" box="[361,409,1313,1334]" pageId="18" pageNumber="219">Lake</collectingCounty>
<collectingRegion id="016311DEFFAFFFA8FE3CFAEEFE0BFA84" box="[414,495,1313,1334]" country="Mongolia" name="Hovsgol" pageId="18" pageNumber="219">Hövsgöl</collectingRegion>
(
<collectingRegion id="016311DEFFAFFFA8FE59FAEEFDAAFA84" box="[507,590,1313,1334]" country="Mongolia" name="Hovsgol" pageId="18" pageNumber="219">Hövsgöl</collectingRegion>
<collectingMunicipality id="237C4546FFAFFFA8FDF6FAEEFD38FA84" box="[596,732,1313,1334]" pageId="18" pageNumber="219">National Park</collectingMunicipality>
), south end of lake near
<location id="C67889E7FFAFFFA8FC6BFAEEFB74FA84" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAFFFAEFF2AFD5AFE5BFEB3:C67889E7FFAFFFA8FC6BFAEEFB74FA84" box="[969,1168,1313,1334]" country="Mongolia" county="Lake" latitude="50.4284" municipality="National Park" name="Hatgal. Coordinates" pageId="18" pageNumber="219" stateProvince="Hovsgol">Hatgal. Coordinates</location>
:
<geoCoordinate id="A693B9FBFFAFFFA8FB3BFAEEFAF9FA84" box="[1177,1309,1313,1334]" degrees="50" direction="north" minutes="25.704" orientation="latitude" pageId="18" pageNumber="219" precision="1" value="50.4284">50°25.704 N</geoCoordinate>
</materialsCitation>
<materialsCitation id="73CFD561FFAFFFA8FABFFAEEFEBAFA10" collectingDate="1996-06-13" collectionCode="ANSP" collectorName="A, M. B. Edlund Collection &amp; Mark B. Edlund &amp; Eugene F. Stoermer &amp; Nergui Soninkhishig" country="Mongolia" elevation="1" location="Slide" longitude="100.15228" municipality="Science Museum of Minnesota" pageId="18" pageNumber="219" specimenCode="GC-36352" specimenCount="1" typeStatus="holotype">
;
<geoCoordinate id="A693B9FBFFAFFFA8FA85FAEEFA4CFA84" box="[1319,1448,1313,1334]" degrees="100" direction="east" minutes="9.137" orientation="longitude" pageId="18" pageNumber="219" precision="1" value="100.15228">100°9.137 E</geoCoordinate>
, epipelon from
<elevation id="488A380FFFAFFFA8FEE8FA8AFE5FFAE8" box="[330,443,1349,1370]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="219" unit="m" value="0.1">
<quantity id="045F72D9FFAFFFA8FEE8FA8AFE65FAE8" box="[330,385,1349,1370]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="219" unit="m" value="0.1">0.1 m</quantity>
depth
</elevation>
(accession number: M129
<collectorName id="6E52BAEAFFAFFFA8FD19FA8AFC24FAE8" box="[699,960,1349,1370]" pageId="18" pageNumber="219">A, M.B. Edlund Collection</collectorName>
,
<collectingMunicipality id="237C4546FFAFFFA8FC6BFA8AFB10FAE8" box="[969,1268,1349,1370]" pageId="18" pageNumber="219">Science Museum of Minnesota</collectingMunicipality>
, collected by
<collectorName id="6E52BAEAFFAFFFA8FADAFA8AFEC5FACC" pageId="18" pageNumber="219">Mark B. Edlund</collectorName>
,
<collectorName id="6E52BAEAFFAFFFA8FE89FAA6FE0CFACC" box="[299,488,1385,1406]" pageId="18" pageNumber="219">Eugene F. Stoermer</collectorName>
and
<collectorName id="6E52BAEAFFAFFFA8FDB4FAA6FD3BFACC" box="[534,735,1385,1406]" pageId="18" pageNumber="219">Nergui Soninkhishig</collectorName>
,
<date id="B719F9FCFFAFFFA8FD4BFAA6FC8DFACC" box="[745,873,1385,1406]" pageId="18" pageNumber="219" value="1996-06-13">
<collectingDate id="A75D0014FFAFFFA8FD4BFAA6FC8DFACC" box="[745,873,1385,1406]" pageId="18" pageNumber="219" value="1996-06-13">13 June 1996</collectingDate>
</date>
) (
<location id="C67889E7FFAFFFA8FCDFFAA6FC54FACC" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAFFFAEFF2AFD5AFE5BFEB3:C67889E7FFAFFFA8FCDFFAA6FC54FACC" box="[893,944,1385,1406]" country="Mongolia" longitude="100.15228" municipality="Science Museum of Minnesota" name="Slide" pageId="18" pageNumber="219">Slide</location>
M129A,
<collectionCode id="A5B647F9FFAFFFA8FBA9FAA6FBA3FACC" box="[1035,1095,1385,1406]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="18" pageNumber="219">ANSP</collectionCode>
<specimenCode id="93017747FFAFFFA8FBEFFAA6FB51FACC" box="[1101,1205,1385,1406]" collectionCode="GC-" pageId="18" pageNumber="219">GC-36352</specimenCode>
, GCM-24054),
<emphasis id="F1D3032EFFAFFFA8FAEFFAA6FEBAFA10" bold="true" pageId="18" pageNumber="219">
<typeStatus id="1C1C619EFFAFFFA8FAEFFAA6FA42FACC" box="[1357,1446,1385,1406]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
, designated here
</emphasis>
</materialsCitation>
; example specimen on
<figureCitation id="5B9CC3B9FFAFFFA8FDE2FA42FD63FA10" box="[576,647,1421,1442]" captionStart="FIGURES 7079" captionStartId="19.[136,243,1986,2007]" captionTargetBox="[205,1381,429,1956]" captionTargetId="figure-149@19.[202,1385,425,1960]" captionTargetPageId="19" captionText="FIGURES 7079. Diploneis paraparma, LM and SEM valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77)." figureDoi="http://doi.org/10.5281/zenodo.13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="18" pageNumber="219">Fig. 70</figureCitation>
;
<materialsCitation id="73CFD561FFAFFFA8FD33FA42FBB3FA10" box="[657,1111,1421,1442]" collectionCode="CAS" country="Mongolia" location="Slide" pageId="18" pageNumber="219" specimenCount="1" typeStatus="isotype">
(
<location id="C67889E7FFAFFFA8FD3AFA42FD2FFA10" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFAFFFAEFF2AFD5AFE5BFEB3:C67889E7FFAFFFA8FD3AFA42FD2FFA10" box="[664,715,1421,1442]" country="Mongolia" name="Slide" pageId="18" pageNumber="219">Slide</location>
918029,
<collectionCode id="A5B647F9FFAFFFA8FC86FA42FCB1FA10" box="[804,853,1421,1442]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="219" type="Museum">CAS</collectionCode>
,
<emphasis id="F1D3032EFFAFFFA8FCFDFA42FBA9FA10" bold="true" box="[863,1101,1421,1442]" pageId="18" pageNumber="219">
<typeStatus id="1C1C619EFFAFFFA8FCFDFA42FC43FA10" box="[863,935,1421,1442]" pageId="18" pageNumber="219" type="isotype">isotype</typeStatus>
designated here
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFAFFFA9FF2AFA1CFC88FE23" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219" type="etymology">
<paragraph id="C318DF3CFFAFFFA8FF2AFA1CFAB1FA5E" blockId="18.[136,1452,1491,2092]" box="[136,1365,1491,1516]" pageId="18" pageNumber="219">
<emphasis id="F1D3032EFFAFFFA8FF2AFA1CFECAFA5E" bold="true" box="[136,302,1491,1516]" pageId="18" pageNumber="219">Etymology:—</emphasis>
The species name refers to this taxons close relationship and confusion with
<taxonomicName id="04A7A4BFFFAFFFA8FB33FA1CFAAAFA5E" authorityName="Cleve" authorityYear="1891" box="[1169,1358,1491,1516]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAFFFA8FB33FA1CFAAAFA5E" box="[1169,1358,1491,1516]" italics="true" pageId="18" pageNumber="219">Diploneis parma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C318DF3CFFAFFFA9FF1FFA38FC88FE23" blockId="18.[136,1452,1491,2092]" lastBlockId="19.[136,1452,160,401]" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
<emphasis id="F1D3032EFFAFFFA8FF1FFA38FE81F9A2" bold="true" box="[189,357,1527,1552]" pageId="18" pageNumber="219">Observations:</emphasis>
<taxonomicName id="04A7A4BFFFAFFFA8FE23FA38FD91F9A2" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[385,629,1527,1552]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAFFFA8FE23FA38FD91F9A2" box="[385,629,1527,1552]" italics="true" pageId="18" pageNumber="219">Diploneis paraparma</emphasis>
</taxonomicName>
can easily be associated with
<taxonomicName id="04A7A4BFFFAFFFA8FC71FA38FBA4F9A2" authorityName="Cleve" authorityYear="1891" box="[979,1088,1527,1552]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAFFFA8FC71FA38FBA4F9A2" box="[979,1088,1527,1552]" italics="true" pageId="18" pageNumber="219">D. parma</emphasis>
</taxonomicName>
. However, the population from Lake Hövsgöl is characterised with an elongated outline compared to the elliptical outline in the
<taxonomicName id="04A7A4BFFFAFFFA8FB73F9D4FAD8F986" authorityName="Cleve" authorityYear="1891" box="[1233,1340,1563,1588]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAFFFA8FB73F9D4FAD8F986" box="[1233,1340,1563,1588]" italics="true" pageId="18" pageNumber="219">D. parma</emphasis>
</taxonomicName>
<typeStatus id="1C1C619EFFAFFFA8FAE1F9D4FA4FF986" box="[1347,1451,1563,1588]" pageId="18" pageNumber="219" type="lectotype">lectotype</typeStatus>
given by
<bibRefCitation id="A736A2CDFFAFFFA8FF56F9F0FDEAF9EA" author="Idei, M. &amp; Kobayasi, H." box="[244,526,1599,1624]" pageId="18" pageNumber="219" pagination="397 - 403" refId="ref26822" refString="Idei, M. &amp; Kobayasi, H. (1986 a) Examination of the type specimens of Diploneis parma Cl. In: Round, F. E. (Ed.) Proceedings of the Ninth International Diatom Symposium, Biopress Ltd., Bristol &amp; Koeltz Scientific Book, Koenigstein, pp. 397 - 403." type="book chapter" year="1986">Idei &amp; Kobayasi (1986a</bibRefCitation>
: figs 1, 2).
<bibRefCitation id="A736A2CDFFAFFFA8FD31F9F0FCDBF9EA" author="Hustedt, F." box="[659,831,1599,1624]" pageId="18" pageNumber="219" pagination="577 - 736" refId="ref26709" refString="Hustedt, F. (1937) Die Kieselalgen Deutschlands, Osterreichs und der Schweiz unter Berucksichtigung der ubrigen Lander Europas sowie der angrenzenden Meeresgebiete. In: Rabenhorst, L. (Ed.) Kryptogamen Flora von Deutschland, Osterreich und der Schweiz. Akademische Verlagsgesellschaft m. b. h. Leipzig 7 (Teil 2, Lief. 5), pp. 577 - 736." type="journal article" year="1937">Hustedt (1937)</bibRefCitation>
depicts two different shapes for
<emphasis id="F1D3032EFFAFFFA8FB19F9F0FAD4F9EA" box="[1211,1328,1599,1624]" italics="true" pageId="18" pageNumber="219">
<taxonomicName id="04A7A4BFFFAFFFA8FB19F9F0FAC8F9EA" authorityName="Cleve" authorityYear="1891" box="[1211,1324,1599,1624]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">D. parma</taxonomicName>
,
</emphasis>
an elliptic and an elongate-elliptic, arguing that these differences in outline do not merit taxonomic separation. However, the observations of the type material (
<bibRefCitation id="A736A2CDFFAFFFA8FDACF948FCFFF912" author="Idei, M. &amp; Kobayasi, H." box="[526,795,1671,1696]" pageId="18" pageNumber="219" pagination="397 - 403" refId="ref26822" refString="Idei, M. &amp; Kobayasi, H. (1986 a) Examination of the type specimens of Diploneis parma Cl. In: Round, F. E. (Ed.) Proceedings of the Ninth International Diatom Symposium, Biopress Ltd., Bristol &amp; Koeltz Scientific Book, Koenigstein, pp. 397 - 403." type="book chapter" year="1986">Idei &amp; Kobayasi 1986a</bibRefCitation>
) did not show specimens with a linear-elliptic form; their analysis of Cleves type material showed no relation to later reports for
<taxonomicName id="04A7A4BFFFAFFFA8FC67F964FBD7F976" authorityName="Cleve" authorityYear="1891" box="[965,1075,1707,1732]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAFFFA8FC67F964FC3DF976" box="[965,985,1707,1732]" italics="true" pageId="18" pageNumber="219">D</emphasis>
.
<emphasis id="F1D3032EFFAFFFA8FC4BF964FBD7F976" box="[1001,1075,1707,1732]" italics="true" pageId="18" pageNumber="219">parma</emphasis>
</taxonomicName>
. It appears that a later
<taxonomicName id="04A7A4BFFFAFFFA8FA9CF964FA4FF976" authorityName="Cleve" authorityYear="1891" box="[1342,1451,1707,1732]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAFFFA8FA9CF964FAB6F976" box="[1342,1362,1707,1732]" italics="true" pageId="18" pageNumber="219">D</emphasis>
.
<emphasis id="F1D3032EFFAFFFA8FAC3F964FA4FF976" box="[1377,1451,1707,1732]" italics="true" pageId="18" pageNumber="219">parma</emphasis>
</taxonomicName>
concept has been erroneously perpetuated. Therefore, Hustedts concept for two shapes within
<taxonomicName id="04A7A4BFFFAFFFA8FB77F900FAA0F95A" authorityName="Cleve" authorityYear="1891" box="[1237,1348,1743,1768]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAFFFA8FB77F900FB0DF95A" box="[1237,1257,1743,1768]" italics="true" pageId="18" pageNumber="219">D</emphasis>
.
<emphasis id="F1D3032EFFAFFFA8FB58F900FAA0F95A" box="[1274,1348,1743,1768]" italics="true" pageId="18" pageNumber="219">parma</emphasis>
</taxonomicName>
is rather questionable because the type material has only a broadly elliptic coarse form. There are no reports for the two different shapes in one locality, and the same is true for Lake Hövsgöl population (this study). Based on historical documentation and our analysis, the concept for two different shapes is poorly supported and therefore we recognize the linear-elliptic form as a new species,
<taxonomicName id="04A7A4BFFFAFFFA8FDC8F890FC84F8CA" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[618,864,1887,1912]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAFFFA8FDC8F890FC84F8CA" box="[618,864,1887,1912]" italics="true" pageId="18" pageNumber="219">Diploneis paraparma</emphasis>
</taxonomicName>
. In addition to Hustedts misguided and expanded concept, the striae structure is usually reported as uniseriate becoming biseriate towards the valve margins, which was a key feature for further widening the concept of this species. In addition to valve shape differences between
<taxonomicName id="04A7A4BFFFAFFFA8FAE3F868FA4FF872" authorityName="Cleve" authorityYear="1891" box="[1345,1451,1959,1984]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAFFFA8FAE3F868FA4FF872" box="[1345,1451,1959,1984]" italics="true" pageId="18" pageNumber="219">D. parma</emphasis>
</taxonomicName>
and
<taxonomicName id="04A7A4BFFFAFFFA8FF18F804FEB9F856" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[186,349,1995,2020]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAFFFA8FF18F804FEB9F856" box="[186,349,1995,2020]" italics="true" pageId="18" pageNumber="219">D. paraparma</emphasis>
</taxonomicName>
, morphological differences can be observed in: the structure of the external raphe branches-i) proximal ends (straight without central pores in
<emphasis id="F1D3032EFFAFFFA8FD1AF820FCA3F7BA" box="[696,839,2031,2056]" italics="true" pageId="18" pageNumber="219">
<taxonomicName id="04A7A4BFFFAFFFA8FD1AF820FCC2F7BA" authorityName="Cleve" authorityYear="1891" box="[696,806,2031,2056]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">D. parma</taxonomicName>
vs
</emphasis>
. straight with drop-like central pores, slightly bent to the same side of the valve in
<taxonomicName id="04A7A4BFFFAFFFA8FE71F7DCFD90F79E" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[467,628,2067,2092]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAFFFA8FE71F7DCFE03F79E" box="[467,487,2067,2092]" italics="true" pageId="18" pageNumber="219">D</emphasis>
.
<emphasis id="F1D3032EFFAFFFA8FE54F7DCFD90F79E" box="[502,628,2067,2092]" italics="true" pageId="18" pageNumber="219">paraparma</emphasis>
</taxonomicName>
) and ii) distal ends (deflect and straight in
<emphasis id="F1D3032EFFAFFFA8FBC4F7DCFB17F79E" box="[1126,1267,2067,2092]" italics="true" pageId="18" pageNumber="219">
<taxonomicName id="04A7A4BFFFAFFFA8FBC4F7DCFB37F79E" authorityName="Cleve" authorityYear="1891" box="[1126,1235,2067,2092]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="18" pageNumber="219" phylum="Ochrophyta" rank="species" species="parma">D. parma</taxonomicName>
vs
</emphasis>
. bent into drop-like short terminal fissures in
<taxonomicName id="04A7A4BFFFAEFFA9FE76FF6FFD90FF0B" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[468,628,160,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAEFFA9FE76FF6FFE0CFF0B" box="[468,488,160,185]" italics="true" pageId="19" pageNumber="220">D</emphasis>
.
<emphasis id="F1D3032EFFAEFFA9FE54FF6FFD90FF0B" box="[502,628,160,185]" italics="true" pageId="19" pageNumber="220">paraparma</emphasis>
</taxonomicName>
); the longitudinal canal (lanceolate expanded in the middle of the valve, composed of one to three areolae in
<emphasis id="F1D3032EFFAEFFA9FD92FF0BFD5BFF6F" box="[560,703,196,221]" italics="true" pageId="19" pageNumber="220">
<taxonomicName id="04A7A4BFFFAEFFA9FD92FF0BFD7AFF6F" authorityName="Cleve" authorityYear="1891" box="[560,670,196,221]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="parma">D. parma</taxonomicName>
vs
</emphasis>
. lanceolate to linear slightly expanded in the middle of the valve, composed of one or two areolae in
<taxonomicName id="04A7A4BFFFAEFFA9FD80FF27FD20FEB3" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[546,708,232,257]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAEFFA9FD80FF27FDD2FEB3" box="[546,566,232,257]" italics="true" pageId="19" pageNumber="220">D</emphasis>
.
<emphasis id="F1D3032EFFAEFFA9FDE4FF27FD20FEB3" box="[582,708,232,257]" italics="true" pageId="19" pageNumber="220">paraparma</emphasis>
</taxonomicName>
); and the valve width (16.027.5 μm in
<taxonomicName id="04A7A4BFFFAEFFA9FB3EFF27FED9FE97" authority=", sensu Idei &amp; Kobayasi 1986 a" authorityName=", sensu Idei &amp; Kobayasi" authorityYear="1986" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAEFFA9FB3EFF27FAF4FEB3" box="[1180,1296,232,257]" italics="true" pageId="19" pageNumber="220">D. parma,</emphasis>
sensu
<bibRefCitation id="A736A2CDFFAEFFA9FAC3FF27FED9FE97" author="Idei, M. &amp; Kobayasi, H." pageId="19" pageNumber="220" pagination="397 - 403" refId="ref26822" refString="Idei, M. &amp; Kobayasi, H. (1986 a) Examination of the type specimens of Diploneis parma Cl. In: Round, F. E. (Ed.) Proceedings of the Ninth International Diatom Symposium, Biopress Ltd., Bristol &amp; Koeltz Scientific Book, Koenigstein, pp. 397 - 403." type="book chapter" year="1986">Idei &amp; Kobayasi 1986a</bibRefCitation>
</taxonomicName>
<emphasis id="F1D3032EFFAEFFA9FEE7FEC3FE80FE97" box="[325,356,268,293]" italics="true" pageId="19" pageNumber="220">vs.</emphasis>
12.516.5 μm in
<taxonomicName id="04A7A4BFFFAEFFA9FD89FEC3FD2FFE97" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[555,715,268,293]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAEFFA9FD89FEC3FD2FFE97" box="[555,715,268,293]" italics="true" pageId="19" pageNumber="220">D. paraparma</emphasis>
</taxonomicName>
). Even though
<taxonomicName id="04A7A4BFFFAEFFA9FCD5FEC3FC06FE97" authorityName=", sensu Idei &amp; Kobayasi" authorityYear="1986" box="[887,994,268,293]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAEFFA9FCD5FEC3FC06FE97" box="[887,994,268,293]" italics="true" pageId="19" pageNumber="220">D. parma</emphasis>
</taxonomicName>
was described from lakes in
<collectingCountry id="BBB09FACFFAEFFA9FA85FEC3FA99FE97" box="[1319,1405,268,293]" name="Finland" pageId="19" pageNumber="220">Finland</collectingCountry>
and
<collectingCountry id="BBB09FACFFAEFFA9FF2AFEFFFF05FEFB" box="[136,225,304,329]" name="Sweden" pageId="19" pageNumber="220">Sweden</collectingCountry>
(
<bibRefCitation id="A736A2CDFFAEFFA9FF53FEFFFE93FEFB" author="Cleve, P. T." box="[241,375,304,329]" pageId="19" pageNumber="220" pagination="1 - 68" refId="ref25442" refString="Cleve, P. T. (1891) The Diatoms of Finland. Actas Societas Pro Fauna et Flora Fennica 8 (2): 1 - 68. http: // dx. doi. org / 10.5962 / bhl. title. 64355" type="journal article" year="1891">Cleve 1891</bibRefCitation>
,
<bibRefCitation id="A736A2CDFFAEFFA9FE26FEFFFD6AFEFB" author="Idei, M. &amp; Kobayasi, H." box="[388,654,304,329]" pageId="19" pageNumber="220" pagination="397 - 403" refId="ref26822" refString="Idei, M. &amp; Kobayasi, H. (1986 a) Examination of the type specimens of Diploneis parma Cl. In: Round, F. E. (Ed.) Proceedings of the Ninth International Diatom Symposium, Biopress Ltd., Bristol &amp; Koeltz Scientific Book, Koenigstein, pp. 397 - 403." type="book chapter" year="1986">Idei &amp; Kobayasi 1986a</bibRefCitation>
), reports of
<taxonomicName id="04A7A4BFFFAEFFA9FCBEFEFFFC6DFEFB" authorityName=", sensu Idei &amp; Kobayasi" authorityYear="1986" box="[796,905,304,329]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAEFFA9FCBEFEFFFC6DFEFB" box="[796,905,304,329]" italics="true" pageId="19" pageNumber="220">D. parma</emphasis>
</taxonomicName>
exist from different localities worldwide. Some reports likely belong to the now newly described
<taxonomicName id="04A7A4BFFFAEFFA9FD1BFE9BFCBEFEDF" authorityName="Jovanovska &amp; Levkov &amp; Edlund" authorityYear="2015" box="[697,858,340,365]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAEFFA9FD1BFE9BFCBEFEDF" box="[697,858,340,365]" italics="true" pageId="19" pageNumber="220">D. paraparma</emphasis>
</taxonomicName>
. Further detailed analyses are necessary to ascertain the identity of the many populations widely reported as
<taxonomicName id="04A7A4BFFFAEFFA9FD58FEB7FC81FE23" authorityName=", sensu Idei &amp; Kobayasi" authorityYear="1986" box="[762,869,376,401]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="parma">
<emphasis id="F1D3032EFFAEFFA9FD58FEB7FC81FE23" box="[762,869,376,401]" italics="true" pageId="19" pageNumber="220">D. parma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="97D88FB4FFAEFFA9FF2AF80DFB33F7AD" ID-DOI="http://doi.org/10.5281/zenodo.13634156" ID-Zenodo-Dep="13634156" httpUri="https://zenodo.org/record/13634156/files/figure.png" pageId="19" pageNumber="220" startId="19.[136,243,1986,2007]" targetBox="[205,1381,429,1956]" targetPageId="19" targetType="figure">
<paragraph id="C318DF3CFFAEFFA9FF2AF80DFB33F7AD" blockId="19.[136,1452,1986,2079]" pageId="19" pageNumber="220">
<emphasis id="F1D3032EFFAEFFA9FF2AF80DFEDFF865" bold="true" box="[136,315,1986,2007]" pageId="19" pageNumber="220">FIGURES 7079.</emphasis>
<taxonomicName id="04A7A4BFFFAEFFA9FEE0F80DFD46F865" authority=", LM and SEM" authorityName="LM and SEM" box="[322,674,1986,2007]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="19" pageNumber="220" phylum="Ochrophyta" rank="species" species="paraparma">
<emphasis id="F1D3032EFFAEFFA9FEE0F80DFDF5F865" box="[322,529,1986,2007]" italics="true" pageId="19" pageNumber="220">Diploneis paraparma</emphasis>
, LM and SEM
</taxonomicName>
valve views. Figs 7075. LM valve views. Fig. 70. Specimen from the holotype slide, M129A. Figs 76, 79. External view of the whole valve. Fig. 77. External view of the central area with deflected proximal raphe ends. Fig. 78. Internal view of the entire valve. Scale bars = 10 μm (Figs 7075); 5 μm (Figs 76, 78, 79); 2 μm (Fig. 77).
</paragraph>
</caption>
<subSubSection id="8BBD8CB7FFA9FFAEFF1FFF6FFE5BFEB3" pageId="20" pageNumber="221" type="biology_ecology">
<paragraph id="C318DF3CFFA9FFAEFF1FFF6FFE5BFEB3" blockId="20.[136,1452,160,257]" pageId="20" pageNumber="221">
<emphasis id="F1D3032EFFA9FFAEFF1FFF6FFE12FF0B" bold="true" box="[189,502,160,185]" pageId="20" pageNumber="221">Ecology and Distribution:</emphasis>
—M068A; M077A; M121A; M129A; M248A; M262A; M329A; M331A; M351A: distributed in central and southern Lake
<collectingRegion id="016311DEFFA9FFAEFD98FF0BFD7DFF6F" box="[570,665,196,221]" country="Mongolia" name="Hovsgol" pageId="20" pageNumber="221">Hövsgöl</collectingRegion>
in the sediments, marl, and in epipelic, epilithic, periphytic communities from
<quantity id="045F72D9FFA9FFAEFF64FF27FEEFFEB3" box="[198,267,232,257]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="221" unit="cm" value="10.0">10 cm</quantity>
to
<quantity id="045F72D9FFA9FFAEFE8DFF27FE8CFEB3" box="[303,360,232,257]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="221" unit="m" value="40.0">40 m</quantity>
depths.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,230 @@
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<taxonomicName id="04A7A4BFFFB0FFB7FF2AF8DCFD61F89C" authority="(Pantocsek) Cleve-Euler" authorityName="Cleve-Euler" authorityYear="1934" baseAuthorityName="Pantocsek" box="[136,645,1811,1838]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="13" pageNumber="214" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFB0FFB7FF2AF8DCFE88F89C" bold="true" box="[136,364,1811,1838]" italics="true" pageId="13" pageNumber="214">Diploneis praeclara</emphasis>
(Pantocsek) Cleve-Euler
</taxonomicName>
(
<figureCitation id="5B9CC3B9FFB0FFB7FD36F8DAFCF0F89C" box="[660,788,1813,1838]" captionStart-0="FIGURES 4148" captionStart-1="FIGURES 4953" captionStartId-0="14.[136,243,1782,1803]" captionStartId-1="15.[136,243,1982,2003]" captionTargetBox-0="[191,1396,308,1755]" captionTargetBox-1="[199,1389,321,1953]" captionTargetId-0="figure-103@14.[187,1400,304,1757]" captionTargetId-1="figure-98@15.[197,1391,317,1956]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 4148. Diploneis praeclara, LM valve views. Scale bar = 10 μm." captionText-1="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi-0="http://doi.org/10.5281/zenodo.13634148" figureDoi-1="http://doi.org/10.5281/zenodo.13634150" httpUri-0="https://zenodo.org/record/13634148/files/figure.png" httpUri-1="https://zenodo.org/record/13634150/files/figure.png" pageId="13" pageNumber="214">Figs 4153</figureCitation>
)
</heading>
</paragraph>
<paragraph id="C318DF3CFFB0FFB4FF2AF895FAC7FEA5" blockId="13.[136,1452,1882,2065]" lastBlockId="14.[136,1451,160,279]" lastPageId="14" lastPageNumber="215" pageId="13" pageNumber="214">
Valves are elliptical-lanceolate to elliptic with convex margins and bluntly rounded ends (
<figureCitation id="5B9CC3B9FFB0FFB7FBECF895FB28F8C3" box="[1102,1228,1882,1905]" captionStart-0="FIGURES 4148" captionStart-1="FIGURES 4953" captionStartId-0="14.[136,243,1782,1803]" captionStartId-1="15.[136,243,1982,2003]" captionTargetBox-0="[191,1396,308,1755]" captionTargetBox-1="[199,1389,321,1953]" captionTargetId-0="figure-103@14.[187,1400,304,1757]" captionTargetId-1="figure-98@15.[197,1391,317,1956]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 4148. Diploneis praeclara, LM valve views. Scale bar = 10 μm." captionText-1="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi-0="http://doi.org/10.5281/zenodo.13634148" figureDoi-1="http://doi.org/10.5281/zenodo.13634150" httpUri-0="https://zenodo.org/record/13634148/files/figure.png" httpUri-1="https://zenodo.org/record/13634150/files/figure.png" pageId="13" pageNumber="214">Figs 4149</figureCitation>
). The valve length is 39.569.5 μm, and the valve breadth is 18.026.0 μm. The axial area is narrow, linear to lanceolate, expanding toward the central area. The central area is transapically expanded, 4.06.0 μm wide. The longitudinal canal is lanceolate, expanded in the middle with two rows of areolae, that coalescence into one at the valve apices (
<figureCitation id="5B9CC3B9FFB0FFB7FC4DF875FB8FF863" box="[1007,1131,1978,2001]" captionStart-0="FIGURES 4148" captionStart-1="FIGURES 4953" captionStartId-0="14.[136,243,1782,1803]" captionStartId-1="15.[136,243,1982,2003]" captionTargetBox-0="[191,1396,308,1755]" captionTargetBox-1="[199,1389,321,1953]" captionTargetId-0="figure-103@14.[187,1400,304,1757]" captionTargetId-1="figure-98@15.[197,1391,317,1956]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 4148. Diploneis praeclara, LM valve views. Scale bar = 10 μm." captionText-1="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi-0="http://doi.org/10.5281/zenodo.13634148" figureDoi-1="http://doi.org/10.5281/zenodo.13634150" httpUri-0="https://zenodo.org/record/13634148/files/figure.png" httpUri-1="https://zenodo.org/record/13634150/files/figure.png" pageId="13" pageNumber="214">Figs 4149</figureCitation>
). Externally, the areolae of the longitudinal canal open in depressions slightly lower then the rest of the non-porous valve surface (
<figureCitation id="5B9CC3B9FFB0FFB7FB0BF815FAC0F843" box="[1193,1316,2010,2033]" captionStart="FIGURES 4953" captionStartId="15.[136,243,1982,2003]" captionTargetBox="[199,1389,321,1953]" captionTargetId="figure-98@15.[197,1391,317,1956]" captionTargetPageId="15" captionText="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi="http://doi.org/10.5281/zenodo.13634150" httpUri="https://zenodo.org/record/13634150/files/figure.png" pageId="13" pageNumber="214">Figs 49, 51</figureCitation>
). Externally, the areolae of the canal are each covered with a cribrum similar to those of the striae (
<figureCitation id="5B9CC3B9FFB0FFB7FBBAF835FB58F7A3" box="[1048,1212,2042,2065]" captionStart="FIGURES 4953" captionStartId="15.[136,243,1982,2003]" captionTargetBox="[199,1389,321,1953]" captionTargetId="figure-98@15.[197,1391,317,1956]" captionTargetPageId="15" captionText="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi="http://doi.org/10.5281/zenodo.13634150" httpUri="https://zenodo.org/record/13634150/files/figure.png" pageId="13" pageNumber="214">Figs 49, 50, 53</figureCitation>
). Externally, the raphe is straight with expanded proximal ends, which are positioned within a small depression (
<figureCitation id="5B9CC3B9FFB3FFB4FBF6FF6FFB36FF05" box="[1108,1234,160,183]" captionStart="FIGURES 4953" captionStartId="15.[136,243,1982,2003]" captionTargetBox="[199,1389,321,1953]" captionTargetId="figure-98@15.[197,1391,317,1956]" captionTargetPageId="15" captionText="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi="http://doi.org/10.5281/zenodo.13634150" httpUri="https://zenodo.org/record/13634150/files/figure.png" pageId="14" pageNumber="215">Figs 49, 53</figureCitation>
). Distally, the raphe branches terminate with short terminal fissures that are bent to the same side of the valve (
<figureCitation id="5B9CC3B9FFB3FFB4FB9FFF0FFB3AFF65" box="[1085,1246,192,215]" captionStart="FIGURES 4953" captionStartId="15.[136,243,1982,2003]" captionTargetBox="[199,1389,321,1953]" captionTargetId="figure-98@15.[197,1391,317,1956]" captionTargetPageId="15" captionText="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi="http://doi.org/10.5281/zenodo.13634150" httpUri="https://zenodo.org/record/13634150/files/figure.png" pageId="14" pageNumber="215">Figs 49, 51, 52</figureCitation>
). The stria are radiate,
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10 μm, composed of round to rectangular areolae,
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10 μm. Externally the alveolate striae are covered with cribrate occlusions that are placed in depressions slightly lower then the rest of the non-porous silica (
<figureCitation id="5B9CC3B9FFB3FFB4FB65FECFFAF2FEA5" box="[1223,1302,256,279]" captionStart="FIGURES 4953" captionStartId="15.[136,243,1982,2003]" captionTargetBox="[199,1389,321,1953]" captionTargetId="figure-98@15.[197,1391,317,1956]" captionTargetPageId="15" captionText="FIGURES 4953. Diploneis praeclara, SEM external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50)." figureDoi="http://doi.org/10.5281/zenodo.13634150" httpUri="https://zenodo.org/record/13634150/files/figure.png" pageId="14" pageNumber="215">Fig. 50</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="97D88FB4FFB3FFB4FF2AF939FC8AF8B9" ID-DOI="http://doi.org/10.5281/zenodo.13634148" ID-Zenodo-Dep="13634148" box="[136,878,1782,1803]" httpUri="https://zenodo.org/record/13634148/files/figure.png" pageId="14" pageNumber="215" startId="14.[136,243,1782,1803]" targetBox="[191,1396,308,1755]" targetPageId="14" targetType="figure">
<paragraph id="C318DF3CFFB3FFB4FF2AF939FC8AF8B9" blockId="14.[136,878,1782,1803]" box="[136,878,1782,1803]" pageId="14" pageNumber="215">
<emphasis id="F1D3032EFFB3FFB4FF2AF939FEDFF8B9" bold="true" box="[136,315,1782,1803]" pageId="14" pageNumber="215">FIGURES 4148.</emphasis>
<taxonomicName id="04A7A4BFFFB3FFB4FEE3F939FDD7F8B9" authority=", LM" authorityName="LM" box="[321,563,1782,1803]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="14" pageNumber="215" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFB3FFB4FEE3F939FDE7F8B9" box="[321,515,1782,1803]" italics="true" pageId="14" pageNumber="215">Diploneis praeclara</emphasis>
, LM
</taxonomicName>
valve views. Scale bar = 10 μm.
</paragraph>
</caption>
<subSubSection id="8BBD8CB7FFB3FFAAFF2AF8F3FE7DFE23" lastPageId="16" lastPageNumber="217" pageId="14" pageNumber="215" type="description">
<paragraph id="C318DF3CFFB3FFB5FF2AF8F3FB73FE97" blockId="14.[136,1452,1852,2093]" lastBlockId="15.[136,1451,160,293]" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
<emphasis id="F1D3032EFFB3FFB4FF2AF8F3FEA8F8E7" bold="true" box="[136,332,1852,1877]" pageId="14" pageNumber="215">Observations:—</emphasis>
The measurements of Lake Hövsgöls
<taxonomicName id="04A7A4BFFFB3FFB4FD5AF8F3FAD9F8E7" authority="(Pantocsek) Cleve-Euler (1934" authorityName="Cleve-Euler" authorityYear="1934" baseAuthorityName="Pantocsek" box="[760,1341,1852,1877]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="14" pageNumber="215" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFB3FFB4FD5AF8F3FC3DF8E7" box="[760,985,1852,1877]" italics="true" pageId="14" pageNumber="215">Diploneis praeclara</emphasis>
(Pantocsek)
<bibRefCitation id="A736A2CDFFB3FFB4FBC8F8F3FAD9F8E7" author="Cleve-Euler, A." box="[1130,1341,1852,1877]" pageId="14" pageNumber="215" pagination="1 - 154" refId="ref25516" refString="Cleve-Euler, A. (1934) The diatoms of Finnish Lapland. Societas Scientiarum Fennica. Commentationes Biologicae 4: 1 - 154." type="journal article" year="1934">Cleve-Euler (1934</bibRefCitation>
</taxonomicName>
: pl. 5, fig. 151), fully fit the original description given by
<bibRefCitation id="A736A2CDFFB3FFB4FD06F8AFFC86F8CB" author="Pantocsek, J." box="[676,866,1888,1913]" pageId="14" pageNumber="215" refId="ref28024" refString="Pantocsek, J. (1892) Beitrage Zur Kenntniss der Fossilen Bacillarien Ungarns. III. Susswasser Bacillarien. Anhang: Analysen 15 neuer Depotsvon Bulgarien, Japan, Mahren, Rublandsund Ungarn, 42 Taf. Nagytapolcsany, Julius Platzko." type="book" year="1892">Pantocsek (1892</bibRefCitation>
,
<bibRefCitation id="A736A2CDFFB3FFB4FCCCF8AFFC4EF8CB" author="Pantocsek, J." box="[878,938,1888,1913]" pageId="14" pageNumber="215" refId="ref28116" refString="Pantocsek, J. (1905) Beitragezur Kenntnissder fossilen Bacillarien Ungarns III. Beschreibungderauf Tafel 1 - 4 abgebilden Arten, Pozsoni." type="book" year="1905">1905</bibRefCitation>
).
<taxonomicName id="04A7A4BFFFB3FFB4FC1DF8AFFB46F8CB" authorityName="Cleve-Euler" authorityYear="1934" baseAuthorityName="Pantocsek" box="[959,1186,1888,1913]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="14" pageNumber="215" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFB3FFB4FC1DF8AFFB46F8CB" box="[959,1186,1888,1913]" italics="true" pageId="14" pageNumber="215">Diploneis praeclara</emphasis>
</taxonomicName>
is described as a fossil from Köpecz, the Neogene fossil deposits in
<collectingCountry id="BBB09FACFFB3FFB4FDD7F84BFD3EF82F" box="[629,730,1924,1949]" name="Romania" pageId="14" pageNumber="215">Romania</collectingCountry>
(
<bibRefCitation id="A736A2CDFFB3FFB4FD45F84BFC7FF82F" author="Pantocsek, J." box="[743,923,1924,1949]" pageId="14" pageNumber="215" refId="ref28024" refString="Pantocsek, J. (1892) Beitrage Zur Kenntniss der Fossilen Bacillarien Ungarns. III. Susswasser Bacillarien. Anhang: Analysen 15 neuer Depotsvon Bulgarien, Japan, Mahren, Rublandsund Ungarn, 42 Taf. Nagytapolcsany, Julius Platzko." type="book" year="1892">Pantocsek 1892</bibRefCitation>
,
<bibRefCitation id="A736A2CDFFB3FFB4FC07F84BFC05F82F" author="Pantocsek, J." box="[933,993,1924,1949]" pageId="14" pageNumber="215" refId="ref28116" refString="Pantocsek, J. (1905) Beitragezur Kenntnissder fossilen Bacillarien Ungarns III. Beschreibungderauf Tafel 1 - 4 abgebilden Arten, Pozsoni." type="book" year="1905">1905</bibRefCitation>
). In the modern flora it was reported from Lake Ohrid, reassigning its status from fossil to relic (
<bibRefCitation id="A736A2CDFFB3FFB4FD52F867FC94F873" author="Jurilj, A." box="[752,880,1960,1985]" pageId="14" pageNumber="215" pagination="99 - 190" refId="ref27204" refString="Jurilj, A. (1954) Flora i vegetacija Dijatomeja Ohridskog Jezera. Flora and vegetation of Diatoms from Ochrida Lake in Yugoslavia. J Prirodnoslovnih Istrazivanja 26: 99 - 190." type="journal article" year="1954">Jurilj 1954</bibRefCitation>
). Recently,
<bibRefCitation id="A736A2CDFFB3FFB4FC54F867FAFEF873" author="Jovanovska, E. &amp; Nakov, T. &amp; Levkov, Z." box="[1014,1306,1960,1985]" pageId="14" pageNumber="215" pagination="237 - 262" refId="ref27009" refString="Jovanovska, E., Nakov, T. &amp; Levkov, Z. (2013 a) Observations of the genus Diploneis (Ehrenberg) Cleve from Lake Ohrid, Macedonia. Diatom Research 28: 237 - 262. http: // dx. doi. org / 10.1080 / 0269249 x. 2013.797219" type="journal article" year="2013">
Jovanovska
<emphasis id="F1D3032EFFB3FFB4FB23F867FB50F873" box="[1153,1204,1960,1985]" italics="true" pageId="14" pageNumber="215">et al</emphasis>
. (2013a)
</bibRefCitation>
documented a population with a smaller size range in both modern and fossil sediments in Lake Ohrid (length: 31.039.7 μm; breadth: 16.318.3 μm). The illustrations given by
<bibRefCitation id="A736A2CDFFB3FFB4FD76F83FFC1EF7BB" author="Jovanovska, E. &amp; Nakov, T. &amp; Levkov, Z." box="[724,1018,2032,2057]" pageId="14" pageNumber="215" pagination="237 - 262" refId="ref27009" refString="Jovanovska, E., Nakov, T. &amp; Levkov, Z. (2013 a) Observations of the genus Diploneis (Ehrenberg) Cleve from Lake Ohrid, Macedonia. Diatom Research 28: 237 - 262. http: // dx. doi. org / 10.1080 / 0269249 x. 2013.797219" type="journal article" year="2013">
Jovanovska
<emphasis id="F1D3032EFFB3FFB4FCFDF83FFC77F7BB" box="[863,915,2032,2057]" italics="true" pageId="14" pageNumber="215">et al</emphasis>
. (2013a)
</bibRefCitation>
show differences in the valve outline between Lake Ohrid and Köpecz populations (elliptical with acute apices
<emphasis id="F1D3032EFFB3FFB4FC6DF7DBFC02F79F" box="[975,998,2068,2093]" italics="true" pageId="14" pageNumber="215">vs</emphasis>
. elliptical with bluntly rounded apices). The areola density is slightly different between lakes Hövsgöl and Ohrid populations (
<quantity id="045F72D9FFB2FFB5FBF8FF6FFB40FF0B" box="[1114,1188,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="15" pageNumber="216" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">89 in</quantity>
10 μm
<emphasis id="F1D3032EFFB2FFB5FB5FFF6FFAF0FF0B" box="[1277,1300,160,185]" italics="true" pageId="15" pageNumber="216">vs</emphasis>
.
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10 μm). In the original description
<bibRefCitation id="A736A2CDFFB2FFB5FE4CFF0BFD55FF6F" author="Pantocsek, J." box="[494,689,196,221]" pageId="15" pageNumber="216" refId="ref28116" refString="Pantocsek, J. (1905) Beitragezur Kenntnissder fossilen Bacillarien Ungarns III. Beschreibungderauf Tafel 1 - 4 abgebilden Arten, Pozsoni." type="book" year="1905">Pantocsek (1905)</bibRefCitation>
gives a stria density of
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10 μm, which fully fits the Lake Ohrid population. On the other hand,
<bibRefCitation id="A736A2CDFFB2FFB5FE4EFF27FD9CFEB3" author="Jurilj, A." box="[492,632,232,257]" pageId="15" pageNumber="216" pagination="99 - 190" refId="ref27204" refString="Jurilj, A. (1954) Flora i vegetacija Dijatomeja Ohridskog Jezera. Flora and vegetation of Diatoms from Ochrida Lake in Yugoslavia. J Prirodnoslovnih Istrazivanja 26: 99 - 190." type="journal article" year="1954">Jurilj (1954)</bibRefCitation>
in his description gives a stria density of
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10 μm, similar to Lake Hövsgöl population. Due to this, using the stria density as a distinguishing feature is unclear.
</paragraph>
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<paragraph id="C318DF3CFFB2FFB5FF2AF871FBF8F7A9" blockId="15.[136,1451,1982,2075]" pageId="15" pageNumber="216">
<emphasis id="F1D3032EFFB2FFB5FF2AF871FED8F861" bold="true" box="[136,316,1982,2003]" pageId="15" pageNumber="216">FIGURES 4953.</emphasis>
<taxonomicName id="04A7A4BFFFB2FFB5FEE0F871FDA6F861" authority=", SEM" authorityName="SEM" box="[322,578,1982,2003]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="15" pageNumber="216" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFB2FFB5FEE0F871FDE1F861" box="[322,517,1982,2003]" italics="true" pageId="15" pageNumber="216">Diploneis praeclara</emphasis>
, SEM
</taxonomicName>
external valve views. Fig. 49. View of the entire valve. Fig. 50. Close view of the cribrate occlusions. Fig. 51. Distal raphe endings. Fig. 52. One row of areolae of the longitudinal canal at the valve apices. Fig. 53. Central area with proximal raphe endings. Scale bars = 10 μm (Fig. 49); 5 μm (Figs 5153); 2 μm (Fig. 50).
</paragraph>
</caption>
<paragraph id="C318DF3CFFADFFAAFF1FFF6FFAC3FE97" blockId="16.[136,1452,160,473]" pageId="16" pageNumber="217">
Followed by the recent typification of
<taxonomicName id="04A7A4BFFFADFFAAFDD3FF6FFBCAFF0B" authority="(Jovanovska et al. 2013 b)" baseAuthorityName="Jovanovska" baseAuthorityYear="2013" box="[625,1070,160,185]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFADFFAAFDD3FF6FFCE6FF0B" box="[625,770,160,185]" italics="true" pageId="16" pageNumber="217">D. praeclara</emphasis>
(
<bibRefCitation id="A736A2CDFFADFFAAFCB0FF6FFBC2FF0B" author="Jovanovska, E. &amp; Buczko, K. &amp; Ognjanova-Rumenova, N. &amp; Nakov, T. &amp; Levkov, Z." box="[786,1062,160,185]" pageId="16" pageNumber="217" pagination="15 - 26" refId="ref27065" refString="Jovanovska, E., Buczko, K., Ognjanova-Rumenova, N., Nakov, T. &amp; Levkov, Z. (2013 b) Identity and typification of Diploneis ostracodarum, Diploneis budayana and Diploneis praeclara (Bacillariophyta). Phytotaxa 137 (1): 15 - 26. http: // dx. doi. org / 10.11646 / phytotaxa. 137.1.2" type="journal article" year="2013">
Jovanovska
<emphasis id="F1D3032EFFADFFAAFC3CFF6FFC34FF0B" box="[926,976,160,185]" italics="true" pageId="16" pageNumber="217">et al</emphasis>
. 2013b
</bibRefCitation>
)
</taxonomicName>
it seems like the population from Lake
<collectingRegion id="016311DEFFADFFAAFF64FF0BFEC1FF6F" box="[198,293,196,221]" country="Mongolia" name="Hovsgol" pageId="16" pageNumber="217">Hövsgöl</collectingRegion>
more closely resembles the
<typeStatus id="1C1C619EFFADFFAAFDFCFF0BFD6AFF6F" box="[606,654,196,221]" pageId="16" pageNumber="217">type</typeStatus>
population rather then the one from Lake
<collectingRegion id="016311DEFFADFFAAFBC3FF0BFB42FF6F" box="[1121,1190,196,221]" country="Macedonia" name="Ohrid" pageId="16" pageNumber="217">Ohrid</collectingRegion>
. However, the presence of the biseriate pattern in the middle of the longitudinal canal makes the separation of Lake Ohrids population rather difficult, and we currently consider this morphological variation to be within the concept of
<taxonomicName id="04A7A4BFFFADFFAAFB2DFEC3FAC4FE97" box="[1167,1312,268,293]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFADFFAAFB2DFEC3FB47FE97" box="[1167,1187,268,293]" italics="true" pageId="16" pageNumber="217">D</emphasis>
.
<emphasis id="F1D3032EFFADFFAAFB10FEC3FAC4FE97" box="[1202,1312,268,293]" italics="true" pageId="16" pageNumber="217">praeclara</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C318DF3CFFADFFAAFF2AFEFFFE7DFE23" blockId="16.[136,1452,160,473]" pageId="16" pageNumber="217">
This study expands the distribution of
<taxonomicName id="04A7A4BFFFADFFAAFDEAFEFFFD3FFEFB" box="[584,731,304,329]" class="Bacillariophyceae" family="Diploneidaceae" genus="Diploneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="praeclara">
<emphasis id="F1D3032EFFADFFAAFDEAFEFFFD3FFEFB" box="[584,731,304,329]" italics="true" pageId="16" pageNumber="217">D. praeclara</emphasis>
</taxonomicName>
, previously reported only from the Neogene fossil deposits in
<collectingCountry id="BBB09FACFFADFFAAFF2AFE9BFF09FEDF" box="[136,237,340,365]" name="Romania" pageId="16" pageNumber="217">Romania</collectingCountry>
(
<bibRefCitation id="A736A2CDFFADFFAAFF59FE9BFE54FEDF" author="Pantocsek, J." box="[251,432,340,365]" pageId="16" pageNumber="217" refId="ref28024" refString="Pantocsek, J. (1892) Beitrage Zur Kenntniss der Fossilen Bacillarien Ungarns. III. Susswasser Bacillarien. Anhang: Analysen 15 neuer Depotsvon Bulgarien, Japan, Mahren, Rublandsund Ungarn, 42 Taf. Nagytapolcsany, Julius Platzko." type="book" year="1892">Pantocsek 1892</bibRefCitation>
,
<bibRefCitation id="A736A2CDFFADFFAAFE18FE9BFE11FEDF" author="Pantocsek, J." box="[442,501,340,365]" pageId="16" pageNumber="217" refId="ref28116" refString="Pantocsek, J. (1905) Beitragezur Kenntnissder fossilen Bacillarien Ungarns III. Beschreibungderauf Tafel 1 - 4 abgebilden Arten, Pozsoni." type="book" year="1905">1905</bibRefCitation>
), Scandinavian fossil deposits (
<bibRefCitation id="A736A2CDFFADFFAAFCF0FE9BFBF9FEDF" author="Cleve-Euler, A." box="[850,1053,340,365]" pageId="16" pageNumber="217" pagination="1 - 154" refId="ref25516" refString="Cleve-Euler, A. (1934) The diatoms of Finnish Lapland. Societas Scientiarum Fennica. Commentationes Biologicae 4: 1 - 154." type="journal article" year="1934">Cleve-Euler 1934</bibRefCitation>
,
<bibRefCitation id="A736A2CDFFADFFAAFB85FE9BFB84FEDF" author="Cleve-Euler, A." box="[1063,1120,340,365]" pageId="16" pageNumber="217" pagination="1 - 255" refId="ref25541" refString="Cleve-Euler, A. (1953) Die Diatomeen von Schweden und Finnland. Teil III. Monoraphideae, Biraphideae 1. Kongiliga Svenska Vetenskaps-Akademiens Handlingar, Fjarde Serien 4: 1 - 255." type="journal article" year="1953">1953</bibRefCitation>
) and Lake Ohrid (
<bibRefCitation id="A736A2CDFFADFFAAFA89FE9BFA4CFEDF" author="Jurilj, A." box="[1323,1448,340,365]" pageId="16" pageNumber="217" pagination="99 - 190" refId="ref27204" refString="Jurilj, A. (1954) Flora i vegetacija Dijatomeja Ohridskog Jezera. Flora and vegetation of Diatoms from Ochrida Lake in Yugoslavia. J Prirodnoslovnih Istrazivanja 26: 99 - 190." type="journal article" year="1954">Jurilj 1954</bibRefCitation>
, Jovanovska
<emphasis id="F1D3032EFFADFFAAFEB0FEB7FEA7FE23" box="[274,323,376,401]" italics="true" pageId="16" pageNumber="217">et al</emphasis>
. 2013).
</paragraph>
</subSubSection>
<subSubSection id="8BBD8CB7FFADFFAAFF1FFE53FBD0FE6B" pageId="16" pageNumber="217" type="biology_ecology">
<paragraph id="C318DF3CFFADFFAAFF1FFE53FBD0FE6B" blockId="16.[136,1452,160,473]" pageId="16" pageNumber="217">
<emphasis id="F1D3032EFFADFFAAFF1FFE53FE16FE07" bold="true" box="[189,498,412,437]" pageId="16" pageNumber="217">Ecology and Distribution:</emphasis>
—M274A; M247A; M248A; M284A; M288A; M329A;
<materialsCitation id="73CFD561FFADFFAAFBCCFE53FBCBFE6B" location="central Lake" pageId="16" pageNumber="217" specimenCount="1" stateProvince="Hovsgol">
M330A: found in shallow to deep habitats in northern and
<location id="C67889E7FFADFFAAFE76FE0FFDBBFE6B" LSID="urn:lsid:plazi:treatment:4B0E6E2AFFB0FFAAFF2AF8DCFBD0FE6B:C67889E7FFADFFAAFE76FE0FFDBBFE6B" box="[468,607,448,473]" name="central Lake" pageId="16" pageNumber="217" stateProvince="Hovsgol">central Lake</location>
<collectingRegion id="016311DEFFADFFAAFDC4FE0FFD21FE6B" box="[614,709,448,473]" country="Mongolia" name="Hovsgol" pageId="16" pageNumber="217">Hövsgöl</collectingRegion>
on sediments and marl deposits
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,311 @@
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11.
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<emphasis id="F1CA3E20C941A501D183F9FCFE20AD15" bold="true" box="[177,453,1598,1625]" italics="true" pageId="18" pageNumber="231">Leptochilus wusugongii</emphasis>
Liang Zhang &amp; Li Bing Zhang
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,
<emphasis id="F1CA3E20C941A501D201F9FDFC6DAD15" box="[819,904,1599,1624]" italics="true" pageId="18" pageNumber="231">
<taxonomicNameLabel id="EAF9835BC941A501D201F9FDFC6DAD15" box="[819,904,1599,1624]" pageId="18" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="5B85FEB7C941A501D2A4F9FCFC0FAD15" box="[918,1002,1598,1624]" captionStart="FIGURE 13" captionStartId="19.[136,229,1785,1807]" captionTargetBox="[195,1392,329,1748]" captionTargetId="figure-91@19.[183,1404,317,1760]" captionTargetPageId="19" captionText="FIGURE 13. Leptochilus wusugongii Liang Zhang &amp; Li Bing Zhang,—A. Habit.—B. Rhizome.—C. Rhizome scales.—D. Portion of abaxial lobe showing sori.—E. Portion of adaxial lobe. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217920" httpUri="https://zenodo.org/record/13217920/files/figure.png" pageId="18" pageNumber="231">Fig. 13</figureCitation>
).
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<emphasis id="F1CA3E20C941A501D1BAF944FF26ADED" bold="true" box="[136,195,1670,1696]" pageId="18" pageNumber="231">Type</emphasis>
:
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.
<collectingRegion id="017A2CD0C941A501D01DF945FDAAADED" box="[303,591,1670,1697]" country="Laos" name="Louangphabang" pageId="18" pageNumber="231">Luang Prabang Province</collectingRegion>
:
<collectingRegion id="017A2CD0C941A501D36FF945FC89ADED" box="[605,876,1670,1697]" country="Laos" name="Louangphabang" pageId="18" pageNumber="231">Luang Prabang District</collectingRegion>
, state water supply protected area, outside
<collectingRegion id="017A2CD0C941A501D456F945FF01AD89" country="Laos" name="Louangphabang" pageId="18" pageNumber="231">Luang Prabang</collectingRegion>
City,
<quantity id="04464FD7C941A501D01BF968FE4EAD88" box="[297,427,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="18" pageNumber="231" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500700 m</quantity>
, on limestone rock in primary forest,
<date id="B700C4F2C941A501D26AF968FBFDAD89" box="[856,1048,1706,1732]" pageId="18" pageNumber="231" value="2011-10-31">31 October 2011</date>
,
<emphasis id="F1CA3E20C941A501D516F969FD1EADA5" italics="true" pageId="18" pageNumber="231">Su-Gong Wu, Xun Gong, Bo Ding, Onevilay Souliya &amp; Khamphanh Thepkaysone WS-2591</emphasis>
(
<typeStatus id="1C055C90C941A501D238F90CFC8BADA5" box="[778,878,1742,1768]" pageId="18" pageNumber="231" type="holotype">holotype</typeStatus>
KUN-1229050!,
<typeStatus id="1C055C90C941A501D506F90CFB60ADA5" box="[1076,1157,1742,1768]" pageId="18" pageNumber="231" type="isotype">isotype</typeStatus>
MO-6351014!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C941A500D18FF930FC68AA69" lastPageId="19" lastPageNumber="232" pageId="18" pageNumber="231" type="diagnosis">
<paragraph id="C301E232C941A501D18FF930FF2AAC7D" blockId="18.[136,1452,1670,2093]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FF930FED5AC41" bold="true" box="[189,304,1778,1804]" pageId="18" pageNumber="231">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C941A501D070F930FDB5AC41" box="[322,592,1778,1804]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="wusugongii">
<emphasis id="F1CA3E20C941A501D070F930FDB5AC41" box="[322,592,1778,1804]" italics="true" pageId="18" pageNumber="231">Leptochilus wusugongii</emphasis>
</taxonomicName>
is characterized by lamina thickly chartaceous, lobes large,
<quantity id="04464FD7C941A501D43EF930FA6EAC40" box="[1292,1419,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.9" metricValueMin="3.5" pageId="18" pageNumber="231" unit="cm" value="4.2" valueMax="4.9" valueMin="3.5">3.54.9 cm</quantity>
in width.
</paragraph>
<paragraph id="C301E232C941A500D18FF8F8FC68AA69" blockId="18.[136,1452,1670,2093]" lastBlockId="19.[136,1451,158,293]" lastPageId="19" lastPageNumber="232" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FF8F8FEEDAC19" bold="true" box="[189,264,1850,1876]" pageId="18" pageNumber="231">Plants</emphasis>
perennial, evergreen.
<emphasis id="F1CA3E20C941A501D337F8F8FD8AAC19" bold="true" box="[517,623,1850,1876]" pageId="18" pageNumber="231">Rhizome</emphasis>
creeping, ca.
<quantity id="04464FD7C941A501D23CF8F8FC73AC18" box="[782,918,1850,1877]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="18" pageNumber="231" unit="mm" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.61.2 mm</quantity>
in diam., dorsiventrally flattened, scaly; scales appressed, slightly spreading at rhizome apex, membranous, margins entire, brown, (1.6)2.13.5 (4.6) × (0.4)
<quantity id="04464FD7C941A501D44BF89CFF04ACD0" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="18" pageNumber="231" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8 1.2 mm</quantity>
.
<emphasis id="F1CA3E20C941A501D1DEF840FEA4ACD1" bold="true" box="[236,321,1922,1948]" pageId="18" pageNumber="231">Fronds</emphasis>
monomorphic, (43)
<quantity id="04464FD7C941A501D31FF840FD49ACD0" box="[557,684,1922,1949]" metricMagnitude="-1" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="18" pageNumber="231" unit="cm" value="95.0" valueMax="120.0" valueMin="70.0">70120 cm</quantity>
long, petiole slightly longer than lamina;
<emphasis id="F1CA3E20C941A501D5B0F840FB39ACD1" bold="true" box="[1154,1244,1922,1948]" pageId="18" pageNumber="231">petioles</emphasis>
stramineous, (26)
<quantity id="04464FD7C941A501D1A2F864FEE7AC8C" box="[144,258,1958,1985]" metricMagnitude="-1" metricUnit="m" metricValue="5.2" metricValueMax="6.6" metricValueMin="3.8" pageId="18" pageNumber="231" unit="cm" value="52.0" valueMax="66.0" valueMin="38.0">3866 cm</quantity>
long, ca.
<quantity id="04464FD7C941A501D046F864FE19AC8C" box="[372,508,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.1" metricValueMin="2.5" pageId="18" pageNumber="231" unit="mm" value="3.3" valueMax="4.1" valueMin="2.5">2.54.1 mm</quantity>
in diam. at middle, wingless;
<emphasis id="F1CA3E20C941A501D264F864FC53AC8D" bold="true" box="[854,950,1958,1984]" pageId="18" pageNumber="231">rachises</emphasis>
wingless, and only slightly winged at distal rachis and the connecting part between rachis and lobe.
<emphasis id="F1CA3E20C941A501D23CF808FC9DACA9" bold="true" box="[782,888,1994,2020]" pageId="18" pageNumber="231">Laminae</emphasis>
broad elliptic, deeply 1-pinnatifid, (15)3355 ×
<quantity id="04464FD7C941A501D1BAF82CFF18A344" box="[136,253,2030,2057]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" metricValueMax="4.0" metricValueMin="2.4" pageId="18" pageNumber="231" unit="cm" value="32.0" valueMax="40.0" valueMin="24.0">2440 cm</quantity>
, widest at middle or below middle of laminae, thickly chartaceous, brown when dry, glabrous, margins of lobes entire, slightly repand.
<emphasis id="F1CA3E20C941A501D0C0F7D0FDDFA361" bold="true" box="[498,570,2066,2092]" pageId="18" pageNumber="231">Lobes</emphasis>
26 pairs, lanceolate, apex acuminate, 1418 ×
<quantity id="04464FD7C941A501D552F7D0FB07A360" box="[1120,1250,2066,2093]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.9" metricValueMin="3.5" pageId="18" pageNumber="231" unit="cm" value="4.2" valueMax="4.9" valueMin="3.5">3.54.9 cm</quantity>
, basal portions of lobes narrowed, ascending upward, midribs of lobes at angles of 3979 degree with rachis, distal pair of lobes slightly narrowed, alternate or nearly opposite, lobe base ca.
<quantity id="04464FD7C940A500D3DFFF00FC88AB90" box="[749,877,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="19" pageNumber="232" unit="cm" value="3.95" valueMax="4.2" valueMin="3.7">3.74.2 cm</quantity>
distant with each other. Midribs distinctly raised, veins slightly visible.
<emphasis id="F1CA3E20C940A500D04FFF24FE4AAA4D" bold="true" box="[381,431,230,256]" pageId="19" pageNumber="232">Sori</emphasis>
linear, 1825 on each side of midrib, centers ca.
<quantity id="04464FD7C940A500D2E0FF24FBCAAA4C" box="[978,1071,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="19" pageNumber="232" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
apart from one another, 1.72.2 ×
<quantity id="04464FD7C940A500D1BAFEC8FEF7AA68" box="[136,274,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="19" pageNumber="232" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
. close to midrib, ca.
<quantity id="04464FD7C940A500D0CBFEC8FDB3AA68" box="[505,598,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="19" pageNumber="232" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">37 mm</quantity>
distant to the lobe margins.
</paragraph>
</subSubSection>
<caption id="97C1B2BAC940A500D1BAF93BFC47AC7E" ID-DOI="http://doi.org/10.5281/zenodo.13217920" ID-Zenodo-Dep="13217920" httpUri="https://zenodo.org/record/13217920/files/figure.png" pageId="19" pageNumber="232" startId="19.[136,229,1785,1807]" targetBox="[195,1392,329,1748]" targetPageId="19" targetType="figure">
<paragraph id="C301E232C940A500D1BAF93BFC47AC7E" blockId="19.[136,1451,1785,1843]" pageId="19" pageNumber="232">
<emphasis id="F1CA3E20C940A500D1BAF93BFEE0AC43" bold="true" box="[136,261,1785,1807]" pageId="19" pageNumber="232">FIGURE 13</emphasis>
.
<taxonomicName id="04BE99B1C940A500D021F93BFCD1AC42" authority="Liang Zhang &amp; Li Bing Zhang" authorityName="Liang Zhang &amp; Li Bing Zhang" box="[275,820,1785,1807]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="19" pageNumber="232" phylum="Tracheophyta" rank="species" species="wusugongii">
<emphasis id="F1CA3E20C940A500D021F93BFE1FAC42" box="[275,506,1785,1807]" italics="true" pageId="19" pageNumber="232">Leptochilus wusugongii</emphasis>
Liang Zhang &amp; Li Bing Zhang
</taxonomicName>
,—A. Habit.—B. Rhizome.—C. Rhizome scales.—D. Portion of abaxial lobe showing sori.—E. Portion of adaxial lobe. Photo credit: Liang Zhang.
</paragraph>
</caption>
<subSubSection id="8BA4B1B9C940A500D18FF8A3FD23A341" pageId="19" pageNumber="232" type="materials_examined">
<paragraph id="C301E232C940A500D18FF8A3FD23A341" blockId="19.[136,1452,1889,2060]" pageId="19" pageNumber="232">
<emphasis id="F1CA3E20C940A500D18FF8A3FDDFAC36" bold="true" box="[189,570,1889,1915]" pageId="19" pageNumber="232">Additional specimens examined</emphasis>
:
<collectingCountry id="BBA9A2A2C940A500D37BF8A0FD73AC31" box="[585,662,1890,1916]" name="Laos" pageId="19" pageNumber="232">LAOS</collectingCountry>
.
<collectingRegion id="017A2CD0C940A500D390F8A0FC58AC36" box="[674,957,1889,1916]" country="Laos" name="Louangphabang" pageId="19" pageNumber="232">Luang Prabang Province</collectingRegion>
:
<collectingRegion id="017A2CD0C940A500D2FAF8A0FB36AC36" box="[968,1235,1889,1916]" country="Laos" name="Louangphabang" pageId="19" pageNumber="232">Luang Prabang District</collectingRegion>
, state water supply protected area, outside
<collectingRegion id="017A2CD0C940A500D0A9F844FDAEACD2" box="[411,587,1925,1952]" country="Laos" name="Louangphabang" pageId="19" pageNumber="232">Luang Prabang</collectingRegion>
City,
<quantity id="04464FD7C940A500D3AAF847FD03ACED" box="[664,742,1925,1952]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="19" pageNumber="232" unit="m" value="610.0">610 m</quantity>
,
<geoCoordinate id="A68A84F5C940A500D3C5F847FCB1ACED" box="[759,852,1925,1952]" degrees="19.83" direction="north" orientation="latitude" pageId="19" pageNumber="232" precision="555" value="19.83">19.83°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C940A500D26FF847FC23ACED" box="[861,966,1925,1952]" degrees="102.15" direction="east" orientation="longitude" pageId="19" pageNumber="232" precision="555" value="102.15">102.15°E</geoCoordinate>
, on limestone rock,
<date id="B700C4F2C940A500D58DF844FA87ACED" box="[1215,1378,1926,1952]" pageId="19" pageNumber="232" value="2019-05-13">13 May 2019</date>
,
<emphasis id="F1CA3E20C940A500D440F847FE2BAC89" italics="true" pageId="19" pageNumber="232">Zhuo Zhou &amp; Liang Zhang LZ403</emphasis>
(KUN!).
<collectingRegion id="017A2CD0C940A500D373F86BFCF8AC8E" box="[577,797,1961,1987]" country="Laos" name="Viangchan" pageId="19" pageNumber="232">Vientiane Province</collectingRegion>
:
<collectingRegion id="017A2CD0C940A500D21BF868FBEAAC8E" box="[809,1039,1961,1988]" inferredBySuffix="true" pageId="19" pageNumber="232">Vang Vieng District</collectingRegion>
, Phou Houat Pha Tang Village,
<quantity id="04464FD7C940A500D4B3F86BFF45ACA5" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="232" unit="m" value="600.0">600 m</quantity>
,
<geoCoordinate id="A68A84F5C940A500D19CF80CFEE9ACA5" box="[174,268,1998,2024]" degrees="19.07" direction="north" orientation="latitude" pageId="19" pageNumber="232" precision="555" value="19.07">19.07°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C940A500D027F80CFE98ACA5" box="[277,381,1998,2024]" degrees="102.41" direction="east" orientation="longitude" pageId="19" pageNumber="232" precision="555" value="102.41">102.41°E</geoCoordinate>
, in primary forest,
<date id="B700C4F2C940A500D36FF80FFCD9ACA5" box="[605,828,1997,2024]" pageId="19" pageNumber="232" value="2007-11-15">15 November 2007</date>
,
<emphasis id="F1CA3E20C940A500D27BF80CFDBAA341" italics="true" pageId="19" pageNumber="232">Su-Gong Wu, Jian-Ying Xiang, En-De Liu, Somsanith Bouamanivong, Onevilay Souliya WS-424</emphasis>
(KUN!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C947A507D18FFF5FFBC7AA4D" pageId="20" pageNumber="233" type="distribution">
<paragraph id="C301E232C947A507D18FFF5FFBC7AA4D" blockId="20.[136,1452,157,580]" pageId="20" pageNumber="233">
<emphasis id="F1CA3E20C947A507D18FFF5FFE0FABFA" bold="true" box="[189,490,157,184]" pageId="20" pageNumber="233">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C947A507D0C3FF5CFD1EABF5" box="[497,763,158,184]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="species" species="wusugongii">
<emphasis id="F1CA3E20C947A507D0C3FF5CFD1EABF5" box="[497,763,158,184]" italics="true" pageId="20" pageNumber="233">Leptochilus wusugongii</emphasis>
</taxonomicName>
was discovered in two provinces in northern
<collectingCountry id="BBA9A2A2C947A507D5CBFF5CFAD6ABF5" box="[1273,1331,158,184]" name="Laos" pageId="20" pageNumber="233">Laos</collectingCountry>
. There is a possibility that this species is distributed in neighboring areas, such as northern
<collectingCountry id="BBA9A2A2C947A507D531FF00FB87AB91" box="[1027,1122,194,220]" name="Vietnam" pageId="20" pageNumber="233">Vietnam</collectingCountry>
and southern
<collectingRegion id="017A2CD0C947A507D5CCFF00FAB2AB91" box="[1278,1367,194,220]" country="China" name="Yunnan" pageId="20" pageNumber="233">Yunnan</collectingRegion>
,
<collectingCountry id="BBA9A2A2C947A507D453FF00FA4DAB91" box="[1377,1448,194,220]" name="China" pageId="20" pageNumber="233">China</collectingCountry>
. This species was exclusively observed thriving on limestone in the primary forest.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C947A507D18FFEC8FAE8A909" pageId="20" pageNumber="233" type="etymology">
<paragraph id="C301E232C947A507D18FFEC8FB6AAA21" blockId="20.[136,1452,157,580]" pageId="20" pageNumber="233">
<emphasis id="F1CA3E20C947A507D18FFEC8FEDBAA69" bold="true" box="[189,318,266,292]" pageId="20" pageNumber="233">Etymology</emphasis>
:
<taxonomicName id="04BE99B1C947A507D07EFEC8FDB3AA69" box="[332,598,266,292]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="species" species="wusugongii">
<emphasis id="F1CA3E20C947A507D07EFEC8FDB3AA69" box="[332,598,266,292]" italics="true" pageId="20" pageNumber="233">Leptochilus wusugongii</emphasis>
</taxonomicName>
is named in honor of Professor Su-Gong Wu, distinguished pteridologist and one of the collectors of the new species. Professor Wu diligently collected a significant number of fern specimens in Southeast Asia, contributing greatly to our understanding of the fern diversity in this region.
</paragraph>
<paragraph id="C301E232C947A507D18FFEB4FAE8A909" blockId="20.[136,1452,157,580]" pageId="20" pageNumber="233">
<emphasis id="F1CA3E20C947A507D18FFEB4FF10AADD" bold="true" box="[189,245,374,400]" pageId="20" pageNumber="233">Note</emphasis>
:
<taxonomicName id="04BE99B1C947A507D037FEB4FDF5AADD" box="[261,528,374,400]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="species" species="wusugongii">
<emphasis id="F1CA3E20C947A507D037FEB4FDF5AADD" box="[261,528,374,400]" italics="true" pageId="20" pageNumber="233">Leptochilus wusugongii</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="04BE99B1C947A507D297FEB5FB88AADD" box="[933,1133,375,400]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="species" species="sinovietnamica">
<emphasis id="F1CA3E20C947A507D297FEB5FB88AADD" box="[933,1133,375,400]" italics="true" pageId="20" pageNumber="233">L. sinovietnamica</emphasis>
</taxonomicName>
in having large habit, broad lobes, and typically 34 pairs of lobes. However, it differs from
<taxonomicName id="04BE99B1C947A507D249FE59FBA3AAF9" box="[891,1094,411,436]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="species" species="sinovietnamica">
<emphasis id="F1CA3E20C947A507D249FE59FBA3AAF9" box="[891,1094,411,436]" italics="true" pageId="20" pageNumber="233">L. sinovietnamica</emphasis>
</taxonomicName>
by having a thicker texture of laminae. To better define the morphological boundaries between these two species, additional field observations and specimen examinations are necessary. Our phylogenetical study resolved
<taxonomicName id="04BE99B1C947A507D28EFE21FBBEAAB1" box="[956,1115,483,508]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="species" species="wusugongii">
<emphasis id="F1CA3E20C947A507D28EFE21FBBEAAB1" box="[956,1115,483,508]" italics="true" pageId="20" pageNumber="233">L. wusugongii</emphasis>
</taxonomicName>
as sister to
<taxonomicName id="04BE99B1C947A507D5ECFE21FA40AAB1" box="[1246,1445,483,508]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="species" species="sinovietnamica">
<emphasis id="F1CA3E20C947A507D5ECFE21FA40AAB1" box="[1246,1445,483,508]" italics="true" pageId="20" pageNumber="233">L. sinovietnamica</emphasis>
</taxonomicName>
, displaying distinct genetic divergence (
<figureCitation id="5B85FEB7C947A507D37EFDC4FD75A96D" box="[588,656,518,544]" captionStart="FIGURE 1" captionStartId="3.[136,229,1956,1978]" captionTargetBox="[153,1430,353,1922]" captionTargetId="figure-76@3.[151,1435,347,1928]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of Leptochilus based on six plastid markers (modified from Zhang et al. 2024). Orange color indicates the new species described here." figureDoi="http://doi.org/10.5281/zenodo.13217896" httpUri="https://zenodo.org/record/13217896/files/figure.png" pageId="20" pageNumber="233">Fig. 1</figureCitation>
). This discovery adds a new species to the
<collectingCountry id="BBA9A2A2C947A507D5A6FDC4FB16A96D" box="[1172,1267,518,544]" name="Vietnam" pageId="20" pageNumber="233">Vietnam</collectingCountry>
clade, which is recognized as one of the earliest-diverging clades within
<taxonomicName id="04BE99B1C947A507D235FDE8FC6FA909" authorityName="Kaulfuss" authorityYear="1824" box="[775,906,554,580]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="20" pageNumber="233" phylum="Tracheophyta" rank="genus">
<emphasis id="F1CA3E20C947A507D235FDE8FC6FA909" box="[775,906,554,580]" italics="true" pageId="20" pageNumber="233">Leptochilus</emphasis>
</taxonomicName>
, as defined in
<bibRefCitation id="A72F9FC3C947A507D51FFDE8FAECA909" author="Zhang, L. &amp; Lu, N. T. &amp; Zhou, X. - M. &amp; Chen, D. - K. &amp; Knapp, R. &amp; Zhou, L. &amp; Guo, L. &amp; Luong, T. T. &amp; Sun, H. &amp; Gao, X. - F. &amp; Zhang, L. - B." box="[1069,1289,554,580]" pageId="20" pageNumber="233" pagination="311 - 322" refId="ref12520" refString="Zhang, L., Lu, N. T., Zhou, X. - M., Chen, D. - K., Knapp, R., Zhou, L., Guo, L., Luong, T. T., Sun, H., Gao, X. - F. &amp; Zhang, L. - B. (2019) A plastid phylogeny of the Old World fern genus Leptochilus (Polypodiaceae): Implications for cryptic speciation and progressive colonization from lower to higher latitudes. Molecular Phylogenetics and Evolution 134: 311 - 322. https: // doi. org / 10.1016 / j. ympev. 2019.01.013" type="journal article" year="2019">
Zhang
<emphasis id="F1CA3E20C947A507D54EFDE9FB51A909" box="[1148,1204,554,580]" italics="true" pageId="20" pageNumber="233">et al.</emphasis>
(2019)
</bibRefCitation>
.
</paragraph>
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</treatment>
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@ -0,0 +1,481 @@
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10.
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<emphasis id="F1CA3E20C942A502D180FF5CFE30ABF5" bold="true" box="[178,469,158,185]" italics="true" pageId="17" pageNumber="230">Leptochilus vietnamensis</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
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,
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(
<figureCitation id="5B85FEB7C942A502D53EFF5CFB85ABF5" box="[1036,1120,158,184]" captionStart="FIGURE 12" captionStartId="17.[136,229,1989,2011]" captionTargetBox="[195,1392,365,1952]" captionTargetId="figure-85@17.[183,1404,353,1964]" captionTargetPageId="17" captionText="FIGURE 12. Leptochilus vietnamensis Liang Zhang, N.T. Lu &amp; Li Bing Zhang,—A. Habit.—B. Abaxial laminae.—C. Abaxial view of portion of sterile lamina.—D. Rhizomes and petioles.—E. Rhizome scales. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217918" httpUri="https://zenodo.org/record/13217918/files/figure.png" pageId="17" pageNumber="230">Fig. 12</figureCitation>
).
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: Lac Son District, Ngoc Son Ngo Luong Nature Reserve, elev.
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,
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, on a tree trunk in limestone mountains,
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,
<emphasis id="F1CA3E20C942A502D228FEC9FADEAA69" box="[794,1339,266,293]" italics="true" pageId="17" pageNumber="230">Li Bing Zhang, Liang Zhang &amp; Ngan Thi Lu 6334</emphasis>
(
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VNMN,
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!).
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<emphasis id="F1CA3E20C942A502D1BAF807FEE1AC97" bold="true" box="[136,260,1989,2011]" pageId="17" pageNumber="230">FIGURE 12</emphasis>
.
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<emphasis id="F1CA3E20C942A502D023F807FDE6AC97" box="[273,515,1989,2010]" italics="true" pageId="17" pageNumber="230">Leptochilus vietnamensis</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
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,—A. Habit.—B. Abaxial laminae.—C. Abaxial view of portion of sterile lamina.—D. Rhizomes and petioles.—E. Rhizome scales. Photo credit: Liang Zhang.
</paragraph>
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<emphasis id="F1CA3E20C941A501D18FFF5CFED5ABF5" bold="true" box="[189,304,158,184]" pageId="18" pageNumber="231">Diagnosis</emphasis>
:
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<emphasis id="F1CA3E20C941A501D073FF5CFDB8ABF5" box="[321,605,158,184]" italics="true" pageId="18" pageNumber="231">Leptochilus vietnamensis</emphasis>
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differs from other species in the genus by the combination of having long petioles in both fertile and sterile fronds, sterile laminae that can reach up to
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in length, and fertile laminae that are less than half the length of the sterile laminae.
</paragraph>
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<emphasis id="F1CA3E20C941A501D18FFEC8FEEDAA69" bold="true" box="[189,264,266,292]" pageId="18" pageNumber="231">Plants</emphasis>
perennial, evergreen, epiphytic. Rhizome long creeping, robust, ca. 2.55.0 mm in diam., dorsiventrally flattened, scaled, with a few short phyllopodia and roots.Apex of rhizome and basal petiole densely scaly, scales subulatelanceolate, 1.82.9 ×
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, brown, basifixed, concolorous, clathrate, margins entire or sparely denticulate, apex acuminate.
<emphasis id="F1CA3E20C941A501D03EFEB4FE84AADD" bold="true" box="[268,353,374,400]" pageId="18" pageNumber="231">Fronds</emphasis>
dimorphic, fertile fronds longer than sterile fronds.
<emphasis id="F1CA3E20C941A501D2F0FEB4FB82AADD" bold="true" box="[962,1127,374,400]" pageId="18" pageNumber="231">Sterile fronds</emphasis>
(15)
<quantity id="04464FD7C941A501D598FEB4FAC5AADC" box="[1194,1312,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="18" pageNumber="231" unit="cm" value="35.0" valueMax="40.0" valueMin="30.0">3040 cm</quantity>
tall, petiole shorter than lamina;
<emphasis id="F1CA3E20C941A501D05DFE58FE2BAAF9" bold="true" box="[367,462,410,436]" pageId="18" pageNumber="231">laminae</emphasis>
ribbon-lanceolate, margins entire, slightly repand, base cuneate, apex acuminate,
<quantity id="04464FD7C941A501D457FE58FF4FAA94" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="18" pageNumber="231" unit="cm" value="25.5" valueMax="30.0" valueMin="21.0">2130 cm</quantity>
long, (2.5)
<quantity id="04464FD7C941A501D00EFE7CFE76AA94" box="[316,403,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="18" pageNumber="231" unit="cm" value="6.5" valueMax="7.0" valueMin="6.0">67 cm</quantity>
wide at middle, chartaceous, glabrous;
<emphasis id="F1CA3E20C941A501D25AFE7CFC27AA95" bold="true" box="[872,962,446,472]" pageId="18" pageNumber="231">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
<quantity id="04464FD7C941A501D1BAFE20FEF4AAB0" box="[136,273,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="18" pageNumber="231" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.52.2 mm</quantity>
in diam., (3)
<quantity id="04464FD7C941A501D084FE20FDE9AAB0" box="[438,524,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="18" pageNumber="231" unit="cm" value="7.0" valueMax="9.0" valueMin="5.0">59 cm</quantity>
long.
<emphasis id="F1CA3E20C941A501D36AFE20FD18AAB1" bold="true" box="[600,765,482,508]" pageId="18" pageNumber="231">Fertile fronds</emphasis>
<quantity id="04464FD7C941A501D235FE20FC9DAAB0" box="[775,888,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="18" pageNumber="231" unit="cm" value="25.5" valueMax="30.0" valueMin="21.0">2130 cm</quantity>
tall, petiole usually longer or the same length as lamina;
<emphasis id="F1CA3E20C941A501D1D1FDC4FEA7A96D" bold="true" box="[227,322,518,544]" pageId="18" pageNumber="231">laminae</emphasis>
elliptic-lanceolate to ovate-lanceolate, margins entire, base cuneate, apex long acuminate, (8)
<quantity id="04464FD7C941A501D456FDC4FF4FA908" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="18" pageNumber="231" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">1015 cm</quantity>
long,
<quantity id="04464FD7C941A501D1C6FDE8FEB8A908" box="[244,349,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.5" metricValueMin="3.0" pageId="18" pageNumber="231" unit="cm" value="4.25" valueMax="5.5" valueMin="3.0">35.5 cm</quantity>
wide at middle, chartaceous, glabrous;
<emphasis id="F1CA3E20C941A501D211FDE8FC98A909" bold="true" box="[803,893,554,580]" pageId="18" pageNumber="231">petioles</emphasis>
<quantity id="04464FD7C941A501D2B7FDE8FBEBA908" box="[901,1038,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="18" pageNumber="231" unit="mm" value="1.75" valueMax="2.1" valueMin="1.4">1.42.1 mm</quantity>
in diam., (8)
<quantity id="04464FD7C941A501D582FDE8FAC4A908" box="[1200,1313,554,581]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="18" pageNumber="231" unit="cm" value="13.5" valueMax="15.0" valueMin="12.0">1215 cm</quantity>
long.
<emphasis id="F1CA3E20C941A501D459FDE8FA4EA909" bold="true" box="[1387,1451,554,580]" pageId="18" pageNumber="231">Veins</emphasis>
anastomosing, lateral vein distinct, veinlets visible.
<emphasis id="F1CA3E20C941A501D3EAFD8CFCEFA925" bold="true" box="[728,778,590,616]" pageId="18" pageNumber="231">Sori</emphasis>
linear, reach lamina margins, 2542 (50) on each side of midrib,
<quantity id="04464FD7C941A501D1D3FDB0FEBAA9C0" box="[225,351,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.2" metricValueMin="1.8" pageId="18" pageNumber="231" unit="cm" value="2.5" valueMax="3.2" valueMin="1.8">1.83.2 cm</quantity>
long,
<quantity id="04464FD7C941A501D095FDB0FDCBA9C0" box="[423,558,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="18" pageNumber="231" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
wide when mature, at angles of 4462 degree with midribs, centers ca.
<quantity id="04464FD7C941A501D465FDB0FF51A9FC" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="18" pageNumber="231" unit="mm" value="1.8" valueMax="2.1" valueMin="1.5">1.52.1 mm</quantity>
apart from one another. Paraphyses absent.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C941A501D18FFD78FD4DAF80" pageId="18" pageNumber="231" type="materials_examined">
<paragraph id="C301E232C941A501D18FFD78FD4DAF80" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFD78FDD3A999" bold="true" box="[189,566,698,724]" pageId="18" pageNumber="231">Additional specimens examined</emphasis>
:
<materialsCitation id="73D6E86FC941A501D376FD78FB90A850" ID-GBIF-Occurrence="4928466308" collectingDate="1998-04-06" collectingDateMax="2003-11-29" collectingDateMin="1998-04-06" collectorName="Daniel K. Harder, P. K &amp; Loc &amp; T. T. Tai &amp; Su-Gong Wu, L. K &amp; Phan, J. Y &amp; Xiang, X &amp; Gong, V. T &amp; Nguyen &amp; K. S. Nguyen" country="Vietnam" county="Tra Tinh District" elevation="749" latitude="20.54" location="Trang Hen" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Cao Bang">
<collectingCountry id="BBA9A2A2C941A501D376FD78FD2CA999" box="[580,713,698,724]" name="Vietnam" pageId="18" pageNumber="231">VIETNAM</collectingCountry>
.
<collectingRegion id="017A2CD0C941A501D3E1FD78FCA3A998" box="[723,838,698,725]" country="Vietnam" name="Cao Bang" pageId="18" pageNumber="231">Cao Bang</collectingRegion>
:
<collectingCounty id="2A609ABEC941A501D262FD79FBE8A999" box="[848,1037,698,725]" pageId="18" pageNumber="231">Tra Tinh District</collectingCounty>
,
<collectingMunicipality id="23657848C941A501D52AFD78FACFA999" box="[1048,1322,698,725]" pageId="18" pageNumber="231">Quoc Toan Municipality</collectingMunicipality>
,
<location id="C661B4E9C941A501D406FD79FA49A998" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D406FD79FA49A998" box="[1332,1452,699,725]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Trang Hen" pageId="18" pageNumber="231" stateProvince="Cao Bang">Trang Hen</location>
(
<location id="C661B4E9C941A501D1A2FD1CFED3A9B5" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D1A2FD1CFED3A9B5" box="[144,310,734,760]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Lakes District" pageId="18" pageNumber="231" stateProvince="Cao Bang">Lakes District</location>
), elev. 666831 m,
<geoCoordinate id="A68A84F5C941A501D325FD1CFD93A9B4" box="[535,630,734,761]" degrees="22.75" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="22.75">22.75°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C941A501D34DFD1CFD02A9B4" box="[639,743,734,761]" degrees="106.28" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="106.28">106.28°E</geoCoordinate>
, on tree branches in disturbed forests in mountains with solid limestone rocks,
<date id="B700C4F2C941A501D076FCC0FE34A851" box="[324,465,770,796]" pageId="18" pageNumber="231" value="1998-04-06">
<collectingDate id="A7443D1AC941A501D076FCC0FE34A851" box="[324,465,770,796]" pageId="18" pageNumber="231" value="1998-04-06">6 April 1998</collectingDate>
</date>
,
<emphasis id="F1CA3E20C941A501D0E9FCC0FC27A850" box="[475,962,770,797]" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D0E9FCC0FD34A851" box="[475,721,770,796]" pageId="18" pageNumber="231">Daniel K. Harder, P.K</collectorName>
.
<collectorName id="6E4B87E4C941A501D3E8FCC1FCE1A851" box="[730,772,771,796]" pageId="18" pageNumber="231">Loc</collectorName>
&amp;
<collectorName id="6E4B87E4C941A501D217FCC1FC92A851" box="[805,887,771,796]" pageId="18" pageNumber="231">T.T. Tai</collectorName>
4254 (
</emphasis>
MO-3200946!
<emphasis id="F1CA3E20C941A501D556FCC1FB88A850" box="[1124,1133,771,797]" italics="true" pageId="18" pageNumber="231">)</emphasis>
.
</materialsCitation>
<materialsCitation id="73D6E86FC941A501D548FCC1FE7CA8C4" ID-GBIF-Occurrence="4928466306" collectingDate="1998-04-06" collectingDateMax="2003-11-29" collectingDateMin="1998-04-06" collectorName="Daniel K. Harder, P. K &amp; Loc &amp; T. T. Tai &amp; Su-Gong Wu, L. K &amp; Phan, J. Y &amp; Xiang, X &amp; Gong, V. T &amp; Nguyen &amp; K. S. Nguyen" country="Vietnam" county="Tra Tinh District" elevation="749" latitude="20.54" location="Trang Hen" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Cao Bang">
<collectingRegion id="017A2CD0C941A501D548FCC1FB0CA851" box="[1146,1257,770,797]" country="Vietnam" name="Hoa Binh" pageId="18" pageNumber="231">Hoa Binh</collectingRegion>
:
<location id="C661B4E9C941A501D5C0FCC1FA42A851" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D5C0FCC1FA42A851" box="[1266,1447,770,797]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Tan Lac District" pageId="18" pageNumber="231" stateProvince="Cao Bang">Tan Lac District</location>
,
<location id="C661B4E9C941A501D1BAFCE5FE77A80D" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D1BAFCE5FE77A80D" box="[136,402,806,833]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Lung Van Municipality" pageId="18" pageNumber="231" stateProvince="Cao Bang">Lung Van Municipality</location>
,
<location id="C661B4E9C941A501D0ADFCE5FDD0A80D" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D0ADFCE5FDD0A80D" box="[415,565,806,833]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Huop village" pageId="18" pageNumber="231" stateProvince="Cao Bang">Huop village</location>
, elev. 10951100 m,
<geoCoordinate id="A68A84F5C941A501D219FCE4FC6EA80C" box="[811,907,806,833]" degrees="20.54" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.54">20.54°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C941A501D2A6FCE4FC1EA80C" box="[916,1019,806,833]" degrees="105.18" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="105.18">105.18°E</geoCoordinate>
, epiphytic, in heavily logged primary evergreen broad-leaved forest,
<date id="B700C4F2C941A501D0EDFC88FD94A829" box="[479,625,842,869]" pageId="18" pageNumber="231" value="2003-11-29">
<collectingDate id="A7443D1AC941A501D0EDFC88FD94A829" box="[479,625,842,869]" pageId="18" pageNumber="231" value="2003-11-29">29 Nov 2003</collectingDate>
</date>
,
<emphasis id="F1CA3E20C941A501D349FC89FF01A8C5" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D349FC89FCA4A829" box="[635,833,843,869]" pageId="18" pageNumber="231">Su-Gong Wu, L.K</collectorName>
.
<collectorName id="6E4B87E4C941A501D278FC88FC50A829" box="[842,949,842,868]" pageId="18" pageNumber="231">Phan, J.Y</collectorName>
.
<collectorName id="6E4B87E4C941A501D28CFC89FBC6A829" box="[958,1059,843,868]" pageId="18" pageNumber="231">Xiang, X</collectorName>
.
<collectorName id="6E4B87E4C941A501D519FC89FB79A829" box="[1067,1180,843,869]" pageId="18" pageNumber="231">Gong, V.T</collectorName>
.
<collectorName id="6E4B87E4C941A501D596FC89FB1FA829" box="[1188,1274,843,868]" pageId="18" pageNumber="231">Nguyen</collectorName>
&amp;
<collectorName id="6E4B87E4C941A501D429FC89FA42A829" box="[1307,1447,843,869]" pageId="18" pageNumber="231">K.S. Nguyen</collectorName>
, WP-303
</emphasis>
(MO-5980968)
</materialsCitation>
;
<materialsCitation id="73D6E86FC941A501D091FCADFC78A8E1" ID-GBIF-Occurrence="4928466310" collectingDate="2013-11-22" collectionCode="VNMN, CDBI" collectorName="Li Bing Zhang &amp; Liang Zhang &amp; Ngan Thi Lu" country="Vietnam" elevation="840" latitude="20.53" location="Tam Son Community" longLatPrecision="761" longitude="105.18" municipality="Tan Lac District" pageId="18" pageNumber="231" specimenCount="1">
<collectingMunicipality id="23657848C941A501D091FCADFDBEA8C5" box="[419,603,878,905]" pageId="18" pageNumber="231">Tan Lac District</collectingMunicipality>
,
<location id="C661B4E9C941A501D357FCADFCB2A8C5" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D357FCADFCB2A8C5" box="[613,855,878,905]" country="Vietnam" latitude="20.53" longLatPrecision="761" longitude="105.18" municipality="Tan Lac District" name="Tam Son Community" pageId="18" pageNumber="231">Tam Son Community</location>
, elev. 840 m,
<geoCoordinate id="A68A84F5C941A501D2C2FCACFBABA8C4" box="[1008,1102,878,905]" degrees="20.53" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.53">20.53°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C941A501D565FCACFB5AA8C4" box="[1111,1215,878,905]" degrees="105.18" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="105.18">105.18°E</geoCoordinate>
,
<date id="B700C4F2C941A501D5FFFCACFA42A8C5" box="[1229,1447,878,904]" pageId="18" pageNumber="231" value="2013-11-22">
<collectingDate id="A7443D1AC941A501D5FFFCACFA42A8C5" box="[1229,1447,878,904]" pageId="18" pageNumber="231" value="2013-11-22">22 November 2013</collectingDate>
</date>
,
<emphasis id="F1CA3E20C941A501D1BAFC51FD29A8E1" box="[136,716,914,941]" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D1BAFC51FED4A8E1" box="[136,305,914,940]" pageId="18" pageNumber="231">Li Bing Zhang</collectorName>
,
<collectorName id="6E4B87E4C941A501D00CFC51FE34A8E1" box="[318,465,914,940]" pageId="18" pageNumber="231">Liang Zhang</collectorName>
&amp;
<collectorName id="6E4B87E4C941A501D0CBFC51FD6EA8E1" box="[505,651,914,940]" pageId="18" pageNumber="231">Ngan Thi Lu</collectorName>
6378
</emphasis>
(
<collectionCode id="A5AF7AF7C941A501D3EFFC50FCD2A8E1" box="[733,823,914,940]" country="Viet Nam" name="Vietnam National Museum of Nature" pageId="18" pageNumber="231">VNMN</collectionCode>
,
<collectionCode id="A5AF7AF7C941A501D276FC50FC74A8E1" box="[836,913,914,940]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="18" pageNumber="231" type="Herbarium">CDBI</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC941A501D299FC50FCD9A8B8" ID-GBIF-Occurrence="4928466313" collectingDate="2002-12-03" collectorName="Daniel &amp; Harder &amp; P. K. Loc &amp; B. Hall &amp; N. Q. Hung" country="Vietnam" elevation="1368" latitude="20.74" location="Pa Co Municipality" longLatPrecision="760" longitude="104.89" municipality="Mai Chau District" pageId="18" pageNumber="231" specimenCount="1">
<collectingMunicipality id="23657848C941A501D299FC50FB99A8E1" box="[939,1148,914,940]" pageId="18" pageNumber="231">Mai Chau District</collectingMunicipality>
,
<location id="C661B4E9C941A501D5BBFC51FA8FA8E1" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D5BBFC51FA8FA8E1" box="[1161,1386,914,941]" country="Vietnam" latitude="20.74" longLatPrecision="760" longitude="104.89" municipality="Mai Chau District" name="Pa Co Municipality" pageId="18" pageNumber="231">Pa Co Municipality</location>
, elev. 1368 m,
<geoCoordinate id="A68A84F5C941A501D1C2FC74FEAAA89D" box="[240,335,950,976]" degrees="20.74" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.74">20.74°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C941A501D06AFC74FE5AA89C" box="[344,447,950,977]" degrees="104.89" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="104.89">104.89°E</geoCoordinate>
, epiphytic, in wet, semi-deciduous and evergreen forest,
<date id="B700C4F2C941A501D560FC74FACAA89D" box="[1106,1327,950,976]" pageId="18" pageNumber="231" value="2002-12-03">
<collectingDate id="A7443D1AC941A501D560FC74FACAA89D" box="[1106,1327,950,976]" pageId="18" pageNumber="231" value="2002-12-03">03 December 2002</collectingDate>
</date>
,
<emphasis id="F1CA3E20C941A501D40EFC74FD98A8B9" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D40EFC74FA6DA89D" box="[1340,1416,950,976]" pageId="18" pageNumber="231">Daniel</collectorName>
K.
<collectorName id="6E4B87E4C941A501D1BAFC18FF3CA8B9" box="[136,217,986,1012]" pageId="18" pageNumber="231">Harder</collectorName>
,
<collectorName id="6E4B87E4C941A501D1D6FC19FEA3A8B9" box="[228,326,986,1012]" pageId="18" pageNumber="231">P.K. Lôc</collectorName>
,
<collectorName id="6E4B87E4C941A501D063FC19FE47A8B9" box="[337,418,986,1012]" pageId="18" pageNumber="231">B. Hall</collectorName>
&amp;
<collectorName id="6E4B87E4C941A501D0F7FC19FDDBA8B9" box="[453,574,987,1013]" pageId="18" pageNumber="231">N.Q. Hung</collectorName>
7701
</emphasis>
(MO-3321598!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC941A501D275FC18FDC9AF71" ID-GBIF-Occurrence="4928466303" collectingDate="2002-12-05" collectorName="Daniel &amp; Harder &amp; B. Hall &amp; N. Q. Hung &amp; P. H. Hoang" country="Vietnam" county="Mai Chau District" elevation="1198" latitude="20.72" location="Thung Ang Village" longLatPrecision="760" longitude="104.89" municipality="Hang Kia Municipality" pageId="18" pageNumber="231" specimenCount="1">
<collectingCounty id="2A609ABEC941A501D275FC18FBF6A8B9" box="[839,1043,986,1012]" pageId="18" pageNumber="231">Mai Chau District</collectingCounty>
,
<collectingMunicipality id="23657848C941A501D52CFC19FAC6A8B9" box="[1054,1315,986,1013]" pageId="18" pageNumber="231">Hang Kia Municipality</collectingMunicipality>
,
<location id="C661B4E9C941A501D41FFC18FF39AF55" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D41FFC18FF39AF55" country="Vietnam" county="Mai Chau District" latitude="20.72" longLatPrecision="760" longitude="104.89" municipality="Hang Kia Municipality" name="Thung Ang Village" pageId="18" pageNumber="231">Thung Ang Village</location>
, elev. 10951300 m,
<geoCoordinate id="A68A84F5C941A501D0F9FC3CFDCFAF55" box="[459,554,1022,1048]" degrees="20.72" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.72">20.72°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C941A501D301FC3CFD7FAF54" box="[563,666,1022,1049]" degrees="104.89" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="104.89">104.89°E</geoCoordinate>
, epiphytic,
<date id="B700C4F2C941A501D22CFC3CFC12AF55" box="[798,1015,1022,1049]" pageId="18" pageNumber="231" value="2002-12-05">
<collectingDate id="A7443D1AC941A501D22CFC3CFC12AF55" box="[798,1015,1022,1049]" pageId="18" pageNumber="231" value="2002-12-05">05 December 2002</collectingDate>
</date>
,
<emphasis id="F1CA3E20C941A501D530FC3CFE89AF71" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D530FC3CFBABAF55" box="[1026,1102,1022,1048]" pageId="18" pageNumber="231">Daniel</collectorName>
K.
<collectorName id="6E4B87E4C941A501D544FC3CFB22AF55" box="[1142,1223,1022,1048]" pageId="18" pageNumber="231">Harder</collectorName>
,
<collectorName id="6E4B87E4C941A501D5E0FC3DFAC3AF55" box="[1234,1318,1022,1048]" pageId="18" pageNumber="231">B. Hall</collectorName>
,
<collectorName id="6E4B87E4C941A501D403FC3DFA4EAF55" box="[1329,1451,1023,1049]" pageId="18" pageNumber="231">N.Q. Hung</collectorName>
&amp;
<collectorName id="6E4B87E4C941A501D194FBE1FECEAF71" box="[166,299,1059,1084]" pageId="18" pageNumber="231">P.H. Hoang</collectorName>
7782
</emphasis>
(MO-6082983!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC941A501D30BFBE0FE18AFC8" ID-GBIF-Occurrence="4928466304" collectingDate="1994-03-08" collectingDateMax="2002-12-09" collectingDateMin="1994-03-08" collectorName="Daniel K. Harder, T. H &amp; Nguyen &amp; L. V. Averyanov &amp; D. K. Harder &amp; N. T. Hiep &amp; H. H. Schmidt &amp; N. Q. Binh" country="Vietnam" county="Mai Chau District" elevation="1147" latitude="22.33" location="Na Pai Village" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Tuyen Quang">
<collectingCounty id="2A609ABEC941A501D30BFBE0FCEFAF71" box="[569,778,1058,1084]" pageId="18" pageNumber="231">Mai Chau District</collectingCounty>
,
<collectingMunicipality id="23657848C941A501D224FBE1FC13AF71" box="[790,1014,1058,1085]" pageId="18" pageNumber="231">Pa Co Municipality</collectingMunicipality>
, near
<location id="C661B4E9C941A501D509FBE1FB00AF71" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D509FBE1FB00AF71" box="[1083,1253,1058,1085]" country="Vietnam" county="Mai Chau District" latitude="22.33" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" name="Na Pai Village" pageId="18" pageNumber="231" stateProvince="Tuyen Quang">Na Pai Village</location>
, elev. 10941200 m,
<geoCoordinate id="A68A84F5C941A501D19EFB84FEEEAF2D" box="[172,267,1094,1120]" degrees="20.74" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.74">20.74°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C941A501D026FB84FE9EAF2C" box="[276,379,1094,1121]" degrees="104.94" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="104.94">104.94°E</geoCoordinate>
, lithophytic and epiphytic on rocks,
<date id="B700C4F2C941A501D224FB84FC0BAF2D" box="[790,1006,1094,1120]" pageId="18" pageNumber="231" value="2002-12-09">
<collectingDate id="A7443D1AC941A501D224FB84FC0BAF2D" box="[790,1006,1094,1120]" pageId="18" pageNumber="231" value="2002-12-09">09 December 2002</collectingDate>
</date>
,
<emphasis id="F1CA3E20C941A501D2CBFB84FED9AFC9" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D2CBFB84FB13AF2D" box="[1017,1270,1094,1120]" pageId="18" pageNumber="231">Daniel K. Harder, T.H</collectorName>
.
<collectorName id="6E4B87E4C941A501D432FB84FAB3AF2D" box="[1280,1366,1094,1120]" pageId="18" pageNumber="231">Nguyên</collectorName>
&amp;
<collectorName id="6E4B87E4C941A501D44BFB85FF1BAFC8" pageId="18" pageNumber="231">L. V. Averyanov</collectorName>
8020
</emphasis>
(MO-3321599!).
</materialsCitation>
<materialsCitation id="73D6E86FC941A501D333FBA9FD41AF80" ID-GBIF-Occurrence="4928466320" collectingDate="1994-03-08" collectingDateMax="2002-12-09" collectingDateMin="1994-03-08" collectorName="Daniel K. Harder, T. H &amp; Nguyen &amp; L. V. Averyanov &amp; D. K. Harder &amp; N. T. Hiep &amp; H. H. Schmidt &amp; N. Q. Binh" country="Vietnam" county="Mai Chau District" elevation="1147" latitude="22.33" location="Na Pai Village" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Tuyen Quang">
<collectingRegion id="017A2CD0C941A501D333FBA9FD79AFC9" box="[513,668,1130,1157]" country="Vietnam" name="Tuyen Quang" pageId="18" pageNumber="231">Tuyen Quang</collectingRegion>
:
<location id="C661B4E9C941A501D397FBA9FC81AFC9" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D397FBA9FC81AFC9" box="[677,868,1130,1157]" country="Vietnam" county="Mai Chau District" latitude="22.33" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" name="Na Hang District" pageId="18" pageNumber="231" stateProvince="Tuyen Quang">Na Hang District</location>
,
<location id="C661B4E9C941A501D25FFBA9FBA4AFC8" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D25FFBA9FBA4AFC8" box="[877,1089,1130,1157]" country="Vietnam" county="Mai Chau District" latitude="22.33" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" name="Nam Chang Forest" pageId="18" pageNumber="231" stateProvince="Tuyen Quang">Nam Chang Forest</location>
, elev. 210 m,
<geoCoordinate id="A68A84F5C941A501D5E7FBA8FAD6AFC8" box="[1237,1331,1130,1157]" degrees="22.33" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="22.33">22.33°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C941A501D40EFBA8FA41AFC8" box="[1340,1444,1130,1157]" degrees="105.42" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="105.42">105.42°E</geoCoordinate>
, on tree branches, 1 m from ground, in primary forests on sandy quartz soils,
<date id="B700C4F2C941A501D2C2FB4CFB77AFE5" box="[1008,1170,1166,1192]" pageId="18" pageNumber="231" value="1994-03-08">
<collectingDate id="A7443D1AC941A501D2C2FB4CFB77AFE5" box="[1008,1170,1166,1192]" pageId="18" pageNumber="231" value="1994-03-08">8 March 1994</collectingDate>
</date>
,
<emphasis id="F1CA3E20C941A501D5ACFB4DFE16AF81" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D5ACFB4DFACEAFE5" box="[1182,1323,1166,1192]" pageId="18" pageNumber="231">D.K. Harder</collectorName>
,
<collectorName id="6E4B87E4C941A501D40AFB4DFA42AFE5" box="[1336,1447,1166,1192]" pageId="18" pageNumber="231">N.T. Hiêp</collectorName>
,
<collectorName id="6E4B87E4C941A501D1BAFB71FEFAAF81" box="[136,287,1202,1228]" pageId="18" pageNumber="231">H.H. Schmidt</collectorName>
&amp;
<collectorName id="6E4B87E4C941A501D071FB71FE51AF81" box="[323,436,1202,1229]" pageId="18" pageNumber="231">N.Q. Biñh</collectorName>
2412
</emphasis>
(MO-320056!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C941A501D18FFB14FB83AE59" pageId="18" pageNumber="231" type="distribution">
<paragraph id="C301E232C941A501D18FFB14FB83AE59" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFB14FE09AFBD" bold="true" box="[189,492,1238,1264]" pageId="18" pageNumber="231">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C941A501D0C7FB14FCF4AFBD" box="[501,785,1238,1264]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D0C7FB14FCF4AFBD" box="[501,785,1238,1264]" italics="true" pageId="18" pageNumber="231">Leptochilus vietnamensis</emphasis>
</taxonomicName>
was discovered in four localities within three provinces in northern
<collectingCountry id="BBA9A2A2C941A501D1DFFB38FEB5AE59" box="[237,336,1274,1300]" name="Vietnam" pageId="18" pageNumber="231">Vietnam</collectingCountry>
. The species is observed growing on tree branches, trunks or on rocks.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C941A501D18FFADCFEDDAD5D" pageId="18" pageNumber="231" type="etymology">
<paragraph id="C301E232C941A501D18FFADCFF0AAE11" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFADCFEDBAE75" bold="true" box="[189,318,1310,1336]" pageId="18" pageNumber="231">Etymology</emphasis>
: The specific epithet is derived from the name of the country where the new species was discovered,
<collectingCountry id="BBA9A2A2C941A501D1BAFA80FF0EAE11" box="[136,235,1346,1372]" name="Vietnam" pageId="18" pageNumber="231">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="C301E232C941A501D18FFAA4FEDDAD5D" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFAA4FF1AAECD" bold="true" box="[189,255,1382,1408]" pageId="18" pageNumber="231">Note:</emphasis>
<taxonomicName id="04BE99B1C941A501D03AFAA4FDC0AECD" box="[264,549,1382,1408]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D03AFAA4FDC0AECD" box="[264,549,1382,1408]" italics="true" pageId="18" pageNumber="231">Leptochilus vietnamensis</emphasis>
</taxonomicName>
shares similarities with both
<taxonomicName id="04BE99B1C941A501D24BFAA5FC25AECD" authorityName="Liang Zhang &amp; Li Bing Zhang" baseAuthorityName="C. Christensen" baseAuthorityYear="1933" box="[889,960,1383,1408]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="wui">
<emphasis id="F1CA3E20C941A501D24BFAA5FC25AECD" box="[889,960,1383,1408]" italics="true" pageId="18" pageNumber="231">L. wui</emphasis>
</taxonomicName>
and
<taxonomicName id="04BE99B1C941A501D2CEFAA5FB8FAECD" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" box="[1020,1130,1382,1408]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C941A501D2CEFAA5FB8FAECD" box="[1020,1130,1382,1408]" italics="true" pageId="18" pageNumber="231">L. chingii</emphasis>
</taxonomicName>
, as the latter two both have slightly dimorphic fronds and linear sori. However,
<taxonomicName id="04BE99B1C941A501D3D4FA49FC7CAEE9" box="[742,921,1419,1444]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D3D4FA49FC7CAEE9" box="[742,921,1419,1444]" italics="true" pageId="18" pageNumber="231">L. vietnamensis</emphasis>
</taxonomicName>
can be distinguished from them by its fertile fronds, which are shorter than its sterile fronds, and the long, ribbon-like lamina of its sterile fronds. The phylogenetic analysis resolved
<taxonomicName id="04BE99B1C941A501D066FA11FDE3AEA1" box="[340,518,1491,1516]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D066FA11FDE3AEA1" box="[340,518,1491,1516]" italics="true" pageId="18" pageNumber="231">L. vietnamensis</emphasis>
</taxonomicName>
as sister to
<taxonomicName id="04BE99B1C941A501D3A6FA11FC6EAEA1" box="[660,907,1490,1516]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C941A501D3A6FA11FC6EAEA1" box="[660,907,1490,1516]" italics="true" pageId="18" pageNumber="231">L. luangprabangensis</emphasis>
</taxonomicName>
, and these two species together are sister to
<taxonomicName id="04BE99B1C941A501D4A7FA11FF33AD5D" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C941A501D4A7FA11FF33AD5D" italics="true" pageId="18" pageNumber="231">L. chingii</emphasis>
</taxonomicName>
(
<figureCitation id="5B85FEB7C941A501D1D7FA34FEC2AD5D" box="[229,295,1526,1552]" captionStart="FIGURE 1" captionStartId="3.[136,229,1956,1978]" captionTargetBox="[153,1430,353,1922]" captionTargetId="figure-76@3.[151,1435,347,1928]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of Leptochilus based on six plastid markers (modified from Zhang et al. 2024). Orange color indicates the new species described here." figureDoi="http://doi.org/10.5281/zenodo.13217896" httpUri="https://zenodo.org/record/13217896/files/figure.png" pageId="18" pageNumber="231">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,866 @@
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<mods:title id="2B7B80513ACA59B58A5521F93DE61588">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:namePart id="7792D0439D473D1B49B5762492F0B5C6">Zhang, Liang</mods:namePart>
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<mods:affiliation id="CD306139882AB492B7B2210C732DC946">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
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<mods:namePart id="25EC9F2DCD350D8ED4485DABDB0F79EA">Lu, Ngan Thi</mods:namePart>
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<mods:affiliation id="623AF01637A18648BD22C9D7A4BEBD07">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="2E74BDE4FB1C33BFF0CDE02361383913" type="email">vnmngan@gmail.com</mods:nameIdentifier>
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<mods:affiliation id="24BCF08E9663923699670C7D626395F6">School of Ecology and Environmental Science, Yunnan University, Kunming, Yunnan 650091, China</mods:affiliation>
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<mods:namePart id="7045F6BBDE0665B0989CD7EBAAD4AFE7">Zhou, Zhuo</mods:namePart>
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<mods:affiliation id="72CBE54E1EB101B27287478EE2F64CCD">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier id="85E969391D6135934FAEADC5D8DECAAA" type="email">zhouzhuo@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:namePart id="437C76B9EBFCA140062253596776D072">Thepkayson, Khamphanh</mods:namePart>
<mods:nameIdentifier id="BA6F18F1D4DC5A092CB6243A49E63F56" type="ORCID">0009-0001-1757-9907</mods:nameIdentifier>
<mods:affiliation id="505E6D82E55EB6CD88F861F118F835D3">Institute of Traditional Medicine, Vientiane, Lao PDR</mods:affiliation>
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<mods:namePart id="3661F7A0C2AB6CF1F2B3808E3434AEE2">Luong, Thien Tam</mods:namePart>
<mods:nameIdentifier id="D597A6D7DB40F2C984A3E962D508369D" type="ORCID">0000-0001-8354-6321</mods:nameIdentifier>
<mods:affiliation id="40E8C3B81E2C36A8FA8F205787F8B449">Department of Ecology - Evolutionary Biology, Vietnam National University Ho Chi Minh City (VNUHCM) - University of Science, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</mods:affiliation>
<mods:nameIdentifier id="6DF454B7EC5FF9EA6079E3CD939A65FB" type="email">lttam1511@gmail.com</mods:nameIdentifier>
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<mods:namePart id="75A565BA4F5B39C67AC9229A34F996BF">Zhang, Li-Bing</mods:namePart>
<mods:nameIdentifier id="C342AA684E678CB9A5286BF54AF44E36" type="ORCID">0000-0002-4905-040X</mods:nameIdentifier>
<mods:affiliation id="F2AB3387382FB3F407C5B1D37421C2D7">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:nameIdentifier id="3C8EDED051D742946D6C9A9C1A80763F" type="email">Libing.Zhang@mobot.org</mods:nameIdentifier>
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<mods:title id="682432FA36AB2F206A81D02E19E971D7">Phytotaxa</mods:title>
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<mods:part id="C178D1C3D4F59CBCA57F9515824B8E4F">
<mods:date id="378A8321188EE382A3C96C033A48D550">2024</mods:date>
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<mods:number id="AEBB2D197F7623B6EA275CB4C4914593">659</mods:number>
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<mods:number id="6143DB6AD8D6E0760E337388C6B1B470">3</mods:number>
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<treatment id="4B175324C952A510D1BAFB76FA49AA08" ID-DOI="http://doi.org/10.5281/zenodo.13652213" ID-GBIF-Taxon="240245546" ID-Zenodo-Dep="13652213" LSID="urn:lsid:plazi:treatment:4B175324C952A510D1BAFB76FA49AA08" httpUri="http://treatment.plazi.org/id/4B175324C952A510D1BAFB76FA49AA08" lastPageId="3" lastPageNumber="216" pageId="1" pageNumber="214">
<subSubSection id="8BA4B1B9C952A512D1BAFB76FE39AF82" box="[136,476,1204,1231]" pageId="1" pageNumber="214" type="nomenclature">
<paragraph id="C301E232C952A512D1BAFB76FE39AF82" blockId="1.[136,476,1204,1231]" box="[136,476,1204,1231]" pageId="1" pageNumber="214">
<heading id="9849555EC952A512D1BAFB76FE39AF82" bold="true" box="[136,476,1204,1231]" fontSize="11" level="1" pageId="1" pageNumber="214" reason="1">
<emphasis id="F1CA3E20C952A512D1BAFB76FE39AF82" bold="true" box="[136,476,1204,1231]" pageId="1" pageNumber="214">
Key to species of
<taxonomicName id="04BE99B1C952A512D067FB76FE39AF82" ID-CoL="5CWQ" authorityName="Kaulfuss" authorityYear="1824" box="[341,476,1204,1231]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="genus">
<emphasis id="F1CA3E20C952A512D067FB76FE39AF82" bold="true" box="[341,476,1204,1231]" italics="true" pageId="1" pageNumber="214">Leptochilus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="8BA4B1B9C952A510D1BAFB3BFA49AA08" lastPageId="3" lastPageNumber="216" pageId="1" pageNumber="214" type="key">
<keyStep id="784AFA97C952A512D1BAFB3BFA49AE67" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAFB3BFA49AE43" blockId="1.[136,1452,1272,2072]" box="[136,1452,1272,1294]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFB3BFA49AE43" box="[136,1452,1272,1294]" pageId="1" pageNumber="214">1. Laminae simple or sometime lamina base lobed...............................................................................................................................2.</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAFAD4FA49AE67" blockId="1.[136,1452,1272,2072]" box="[136,1452,1301,1323]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFAD4FA49AE67" box="[136,1452,1301,1323]" pageId="1" pageNumber="214">- Laminae lobed, pinnatifid, or rarely 2-pinnate to 4-pinnatifid........................................................................................................32.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAFAF0FA49AE29" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAFAF0FA4EAE0A" blockId="1.[136,1452,1272,2072]" box="[136,1451,1330,1352]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFAF0FA4EAE0A" box="[136,1451,1330,1352]" pageId="1" pageNumber="214">2. Sori orbicular or elongated, irregularly scattered ..............................................................................................................................3.</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAFA8DFA49AE29" blockId="1.[136,1452,1272,2072]" box="[136,1452,1359,1381]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFA8DFA49AE29" box="[136,1452,1359,1381]" pageId="1" pageNumber="214">- Sori linear, acrostichoid, or elongated then regularly in one row along each side of midrib............................................................4.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAFAAEFA4EAED3" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAFAAEFA49AECC" blockId="1.[136,1452,1272,2072]" box="[136,1452,1387,1410]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFAAEFA49AECC" box="[136,1452,1387,1410]" pageId="1" pageNumber="214">
3. Petioles and rachises sparely scaly or nearly glabrous, laminae 69 ×
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...........................................
<taxonomicName id="04BE99B1C952A512D5D3FAAEFA49AECC" box="[1249,1452,1387,1409]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="khammouanensis">
<emphasis id="F1CA3E20C952A512D5D3FAAEFA49AECC" bold="true" box="[1249,1452,1387,1409]" italics="true" pageId="1" pageNumber="214">L. khammouanensis</emphasis>
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<keyLead id="784F4107C952A512D1BAFA4BFA4EAED3" box="[136,1451,1416,1438]" pageId="1" pageNumber="214">
- Petioles and rachises distinctively scaly, laminae (3.5)830 × (0.2)
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...............................................................
<taxonomicName id="04BE99B1C952A512D409FA4BFA4EAED3" ID-CoL="3TLKY" authorityName="Fraser-Jenkins" authorityYear="2008" baseAuthorityName="Blume" baseAuthorityYear="1828" box="[1339,1451,1417,1438]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="pteropus">
<emphasis id="F1CA3E20C952A512D409FA4BFA4EAED3" bold="true" box="[1339,1451,1417,1438]" italics="true" pageId="1" pageNumber="214">L. pteropus</emphasis>
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<paragraph id="C301E232C952A512D1BAFA67FA4EAEF6" blockId="1.[136,1452,1272,2072]" box="[136,1451,1445,1467]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFA67FA4EAEF6" box="[136,1451,1445,1467]" pageId="1" pageNumber="214">4. Fronds distinctly dimorphic...............................................................................................................................................................5.</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAFA01FA49AE95" blockId="1.[136,1452,1272,2072]" box="[136,1452,1474,1496]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFA01FA49AE95" box="[136,1452,1474,1496]" pageId="1" pageNumber="214">- Frond monomorphic or slightly dimorphic .....................................................................................................................................25.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAFA1DFA49AD5C" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAFA1DFA49AEB8" blockId="1.[136,1452,1272,2072]" box="[136,1452,1503,1525]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFA1DFA49AEB8" box="[136,1452,1503,1525]" pageId="1" pageNumber="214">5. Fronds strongly dimorphic, fertile fronds linear, sori fully cover the lower surface.........................................................................6.</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAFA3EFA49AD5C" blockId="1.[136,1452,1272,2072]" box="[136,1452,1532,1554]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAFA3EFA49AD5C" box="[136,1452,1532,1554]" pageId="1" pageNumber="214">- Fronds dimorphic, fertile fronds lanceolate, broadly lanceolate, or ovate, sori shortly linear or long linear .................................17.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAF9DAFA49AD06" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF9DAFA49AD63" blockId="1.[136,1452,1272,2072]" box="[136,1452,1560,1582]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF9DAFA49AD63" box="[136,1452,1560,1582]" pageId="1" pageNumber="214">6. Sori acrostichoid................................................................................................................................................................................7.</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAF9F4FA49AD06" blockId="1.[136,1452,1272,2072]" box="[136,1452,1589,1611]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF9F4FA49AD06" box="[136,1452,1589,1611]" pageId="1" pageNumber="214">- Sori shortly linear ............................................................................................................................................................................16.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAF991FA49ADC9" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF991FA4EAD2A" blockId="1.[136,1452,1272,2072]" box="[136,1451,1618,1640]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF991FA4EAD2A" box="[136,1451,1618,1640]" pageId="1" pageNumber="214">7. Sterile fronds shortly petiolated or sessile.........................................................................................................................................8.</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAF9ADFA49ADC9" blockId="1.[136,1452,1272,2072]" box="[136,1452,1647,1669]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF9ADFA49ADC9" box="[136,1452,1647,1669]" pageId="1" pageNumber="214">- Sterile fronds distinctly petiolated...................................................................................................................................................13.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAF94EFA49ADF3" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF94EFA4EADEC" blockId="1.[136,1452,1272,2072]" box="[136,1451,1676,1698]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF94EFA4EADEC" box="[136,1451,1676,1698]" pageId="1" pageNumber="214">
8. Sterile fronds less than
<quantity id="04464FD7C952A512D086F94EFE0AADEC" box="[436,495,1676,1697]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="214" unit="cm" value="10.0">10 cm</quantity>
long....................................................................................................................................................9.
</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAF96BFA49ADF3" blockId="1.[136,1452,1272,2072]" box="[136,1452,1704,1726]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF96BFA49ADF3" box="[136,1452,1704,1726]" pageId="1" pageNumber="214">
- Sterile fronds more than
<quantity id="04464FD7C952A512D0F3F96BFE19ADF3" box="[449,508,1705,1726]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="214" unit="cm" value="10.0">10 cm</quantity>
long ...............................................................................................................................................11.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAF907FA49ADBA" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF907FA4EAD96" blockId="1.[136,1452,1272,2072]" box="[136,1451,1733,1755]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF907FA4EAD96" box="[136,1451,1733,1755]" pageId="1" pageNumber="214">
9. Apexes of sterile laminae cuneate ..................................................................................................................................
<taxonomicName id="04BE99B1C952A512D419F904FA4EAD96" ID-CoL="3TLHN" box="[1323,1451,1733,1755]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="metallicus">
<emphasis id="F1CA3E20C952A512D419F904FA4EAD96" bold="true" box="[1323,1451,1733,1755]" pageId="1" pageNumber="214">
<emphasis id="F1CA3E20C952A512D419F904FADFAD96" bold="true" box="[1323,1338,1734,1755]" italics="true" pageId="1" pageNumber="214">L</emphasis>
.
<emphasis id="F1CA3E20C952A512D474F907FA4EAD96" bold="true" box="[1350,1451,1733,1755]" italics="true" pageId="1" pageNumber="214">metallicus</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAF921FA49ADBA" blockId="1.[136,1452,1272,2072]" box="[136,1452,1762,1784]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF921FA49ADBA" box="[136,1452,1762,1784]" pageId="1" pageNumber="214">- Apexes of sterile laminae rounded ..................................................................................................................................................10.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAF93DFA4EAC7C" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF93DFA4EAC59" blockId="1.[136,1452,1272,2072]" box="[136,1451,1791,1813]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF93DFA4EAC59" box="[136,1451,1791,1813]" pageId="1" pageNumber="214">
10. Length of sterile laminae 3-6 times than lamina width .........................................................................................................
<taxonomicName id="04BE99B1C952A512D466F93DFA4EAC59" ID-CoL="6PFBG" box="[1364,1451,1791,1812]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="minor">
<emphasis id="F1CA3E20C952A512D466F93DFA4EAC59" bold="true" box="[1364,1451,1791,1812]" italics="true" pageId="1" pageNumber="214">L. minor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAF8DEFA4EAC7C" blockId="1.[136,1452,1272,2072]" box="[136,1451,1819,1842]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF8DEFA4EAC7C" box="[136,1451,1819,1842]" pageId="1" pageNumber="214">
- Length of sterile laminae 2-3 times than lamina width ..................................................................................................
<taxonomicName id="04BE99B1C952A512D41AF8DEFA4EAC7C" box="[1320,1451,1819,1841]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="ovatifolius">
<emphasis id="F1CA3E20C952A512D41AF8DEFA4EAC7C" bold="true" box="[1320,1451,1819,1841]" pageId="1" pageNumber="214">
<emphasis id="F1CA3E20C952A512D41AF8DEFAD2AC7C" bold="true" box="[1320,1335,1820,1841]" italics="true" pageId="1" pageNumber="214">L</emphasis>
.
<emphasis id="F1CA3E20C952A512D471F8D9FA4EAC7C" bold="true" box="[1347,1451,1819,1841]" italics="true" pageId="1" pageNumber="214">ovatifolius</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAF8FBFA49AC27" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF8FBFA49AC03" blockId="1.[136,1452,1272,2072]" box="[136,1452,1848,1870]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF8FBFA49AC03" box="[136,1452,1848,1870]" pageId="1" pageNumber="214">
11. Sterile laminae ca.
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wide..............................................................................................................................................
<taxonomicName id="04BE99B1C952A512D46BF8FBFA49AC03" ID-CoL="3HTWY" box="[1369,1452,1848,1870]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="wallii">
<emphasis id="F1CA3E20C952A512D46BF8FBFA49AC03" bold="true" box="[1369,1452,1848,1870]" italics="true" pageId="1" pageNumber="214">L. wallii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="784F4107C952A512D1BAF894FA49AC27" box="[136,1452,1877,1899]" pageId="1" pageNumber="214">
- Sterile laminae more than
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wide ..............................................................................................................................................12.
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</keyStep>
<keyStep id="784AFA97C952A512D1BAF8B0FA4EACE9" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF8B0FA4EACC5" blockId="1.[136,1452,1272,2072]" box="[136,1451,1906,1928]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF8B0FA4EACC5" box="[136,1451,1906,1928]" pageId="1" pageNumber="214">
12. Rhizome sparely scaly with few roots................................................................................................................................
<taxonomicName id="04BE99B1C952A512D40DF8B0FA4EACC5" ID-CoL="3TLBT" box="[1343,1451,1906,1928]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis id="F1CA3E20C952A512D40DF8B0FA4EACC5" bold="true" box="[1343,1451,1906,1928]" italics="true" pageId="1" pageNumber="214">L. axillaris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAF84DFA4EACE9" blockId="1.[136,1452,1272,2072]" box="[136,1451,1934,1957]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF84DFA4EACE9" box="[136,1451,1934,1957]" pageId="1" pageNumber="214">
- Rhizome scaly with many roots .........................................................................................................................................
<taxonomicName id="04BE99B1C952A512D40DF84DFA4EACE9" ID-CoL="C2XKC" box="[1343,1451,1934,1956]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="brevipes">
<emphasis id="F1CA3E20C952A512D40DF84DFA4EACE9" bold="true" box="[1343,1451,1934,1956]" italics="true" pageId="1" pageNumber="214">L. brevipes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C952A512D1BAF86EFA49AC93" pageId="1" pageNumber="214">
<paragraph id="C301E232C952A512D1BAF86EFA4EAC8C" blockId="1.[136,1452,1272,2072]" box="[136,1451,1963,1986]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF86EFA4EAC8C" box="[136,1451,1963,1986]" pageId="1" pageNumber="214">
13. Laminae lobed ....................................................................................................................................................................
<taxonomicName id="04BE99B1C952A512D472F86EFA4EAC8C" ID-CoL="3TLKR" box="[1344,1451,1963,1985]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="poilanei">
<emphasis id="F1CA3E20C952A512D472F86EFA4EAC8C" box="[1344,1451,1963,1985]" italics="true" pageId="1" pageNumber="214">
<emphasis id="F1CA3E20C952A512D472F86EFAAAAC8C" bold="true" box="[1344,1359,1964,1985]" italics="true" pageId="1" pageNumber="214">L</emphasis>
.
<emphasis id="F1CA3E20C952A512D469F869FA4EAC8C" bold="true" box="[1371,1451,1963,1985]" italics="true" pageId="1" pageNumber="214">poilanei</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C952A512D1BAF80BFA49AC93" blockId="1.[136,1452,1272,2072]" box="[136,1452,1992,2014]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF80BFA49AC93" box="[136,1452,1992,2014]" pageId="1" pageNumber="214">- Laminae entire.................................................................................................................................................................................14.</keyLead>
</paragraph>
</keyStep>
<paragraph id="C301E232C952A512D1BAF827FA4EACB7" blockId="1.[136,1452,1272,2072]" box="[136,1451,2021,2043]" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A512D1BAF827FA4EACB7" box="[136,1451,2021,2043]" pageId="1" pageNumber="214">
<keyStep id="784AFA97C952A512D1BAF827FA4EACB7" box="[136,1451,2021,2043]" pageId="1" pageNumber="214">
14. Sterile fronds less than
<quantity id="04464FD7C952A512D086F827FE0AACB6" box="[436,495,2021,2043]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="214" unit="cm" value="10.0">10 cm</quantity>
long..............................................................................................................................
<taxonomicName id="04BE99B1C952A512D426F824FA4EACB7" ID-CoL="3TLCJ" box="[1300,1451,2021,2043]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="214" phylum="Tracheophyta" rank="species" species="cantoniensis">
<emphasis id="F1CA3E20C952A512D426F824FA4EACB7" bold="true" box="[1300,1451,2021,2043]" pageId="1" pageNumber="214">
<emphasis id="F1CA3E20C952A512D426F824FAC6ACB6" bold="true" box="[1300,1315,2022,2043]" italics="true" pageId="1" pageNumber="214">L</emphasis>
.
<emphasis id="F1CA3E20C952A512D41DF827FA4EACB7" bold="true" box="[1327,1451,2021,2042]" italics="true" pageId="1" pageNumber="214">cantoniensis</emphasis>
</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="C301E232C952A511D1BAF7C1FA49ABF8" blockId="1.[136,1452,1272,2072]" lastBlockId="2.[136,1452,159,2053]" lastPageId="2" lastPageNumber="215" pageId="1" pageNumber="214">
<keyLead id="784F4107C952A511D1BAF7C1FA49ABF8" lastPageId="2" lastPageNumber="215" pageId="1" pageNumber="214">
- Sterile fronds more than
<quantity id="04464FD7C952A512D0F3F7C0FE19A355" box="[449,508,2050,2072]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="214" unit="cm" value="10.0">10 cm</quantity>
long ...............................................................................................................................................15.
<keyStep id="784AFA97C951A511D1BAFF5DFA49ABF8" box="[136,1452,159,181]" pageId="2" pageNumber="215">
15. Sterile laminae elliptic or narrowly ovate .......................................................................................................................
<taxonomicName id="04BE99B1C951A511D41FFF5DFA49ABF8" ID-CoL="6PFJT" box="[1325,1452,159,181]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="decurrens">
<emphasis id="F1CA3E20C951A511D41FFF5DFA49ABF8" bold="true" box="[1325,1452,159,181]" italics="true" pageId="2" pageNumber="215">L. decurrens</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFF7EFA4EAB9C" blockId="2.[136,1452,159,2053]" box="[136,1451,187,210]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFF7EFA4EAB9C" box="[136,1451,187,210]" pageId="2" pageNumber="215">
- Sterile laminae lanceolate.............................................................................................................................................
<taxonomicName id="04BE99B1C951A511D429FF7EFA4EAB9C" box="[1307,1451,187,209]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis id="F1CA3E20C951A511D429FF7EFA4EAB9C" bold="true" box="[1307,1451,187,209]" italics="true" pageId="2" pageNumber="215">L. kachinensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<keyStep id="784AFA97C951A511D1BAFF1BFA49AA47" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFF1BFA4EABA3" blockId="2.[136,1452,159,2053]" box="[136,1451,216,239]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFF1BFA4EABA3" box="[136,1451,216,239]" pageId="2" pageNumber="215">
16. Fertile fronds longer than sterile frond..............................................................................................................................
<taxonomicName id="04BE99B1C951A511D471FF1BFA4EABA3" ID-CoL="6PFK4" box="[1347,1451,216,238]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="evrardii">
<emphasis id="F1CA3E20C951A511D471FF1BFA4EABA3" bold="true" box="[1347,1451,216,238]" italics="true" pageId="2" pageNumber="215">L. evrardii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFF34FA49AA47" blockId="2.[136,1452,159,2053]" box="[136,1452,245,267]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFF34FA49AA47" box="[136,1452,245,267]" pageId="2" pageNumber="215">
- Fertile fronds the same length as, or shorter than sterile frond ..............................................................................
<taxonomicName id="04BE99B1C951A511D5CBFF34FA49AA47" ID-CoL="3TLHK" box="[1273,1452,245,267]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="mengsongensis">
<emphasis id="F1CA3E20C951A511D5CBFF34FA49AA47" bold="true" box="[1273,1452,245,267]" italics="true" pageId="2" pageNumber="215">L. mengsongensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFED0FA49AA09" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFED0FA49AA6A" blockId="2.[136,1452,159,2053]" box="[136,1452,274,296]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFED0FA49AA6A" box="[136,1452,274,296]" pageId="2" pageNumber="215">17. Fertile fronds shorter than, the same length as, or slight longer than sterile fronds........................................................................18.</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFEF2FA49AA09" blockId="2.[136,1452,159,2053]" box="[136,1452,303,325]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFEF2FA49AA09" box="[136,1452,303,325]" pageId="2" pageNumber="215">- Fertile fronds at least 1.5 times longer than sterile fronds ..............................................................................................................20.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFE8EFA49AA33" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFE8EFA4EAA2C" blockId="2.[136,1452,159,2053]" box="[136,1451,332,354]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFE8EFA4EAA2C" box="[136,1451,332,354]" pageId="2" pageNumber="215">
18. Fertile fronds shorter than sterile frond ......................................................................................................................
<taxonomicName id="04BE99B1C951A511D422FE8EFA4EAA2C" box="[1296,1451,332,353]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C951A511D422FE8EFA4EAA2C" bold="true" box="[1296,1451,332,353]" italics="true" pageId="2" pageNumber="215">L. vietnamensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFEABFA49AA33" blockId="2.[136,1452,159,2053]" box="[136,1452,361,383]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFEABFA49AA33" box="[136,1452,361,383]" pageId="2" pageNumber="215">- Fertile fronds the same length as or slightly longer than sterile frond ............................................................................................19.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFE44FA4EAAF5" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFE44FA4EAAD6" blockId="2.[136,1452,159,2053]" box="[136,1451,389,411]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFE44FA4EAAD6" box="[136,1451,389,411]" pageId="2" pageNumber="215">
19. Laminae thickly chartaceous, petioles of sterile fronds (0.5)
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long...........................................................................
<taxonomicName id="04BE99B1C951A511D479FE44FA4EAAD6" ID-CoL="3TLCS" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" box="[1355,1451,389,411]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C951A511D479FE44FA4EAAD6" bold="true" box="[1355,1451,389,411]" italics="true" pageId="2" pageNumber="215">L. chingii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFE61FA4EAAF5" blockId="2.[136,1452,159,2053]" box="[136,1451,418,440]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFE61FA4EAAF5" box="[136,1451,418,440]" pageId="2" pageNumber="215">
- Laminae thinly chartaceous, petioles of sterile fronds (0.6)
<quantity id="04464FD7C951A511D220FE60FCB7AAF5" box="[786,850,418,440]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="215" unit="cm" value="1.7">1.7 cm</quantity>
long .......................................................
<taxonomicName id="04BE99B1C951A511D5E1FE61FA4EAAF5" box="[1235,1451,418,440]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C951A511D5E1FE61FA4EAAF5" bold="true" box="[1235,1451,418,440]" italics="true" pageId="2" pageNumber="215">L. luangprabangensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFE7DFA49AABC" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFE7DFA49AA99" blockId="2.[136,1452,159,2053]" box="[136,1452,447,469]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFE7DFA49AA99" box="[136,1452,447,469]" pageId="2" pageNumber="215">
20. Fertile fronds 1.5
<emphasis id="F1CA3E20C951A511D04BFE7DFE60AA99" bold="true" box="[377,389,447,468]" pageId="2" pageNumber="215"></emphasis>
2 times longer than sterile fronds.......................................................................................................................21.
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFE1EFA49AABC" blockId="2.[136,1452,159,2053]" box="[136,1452,476,498]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFE1EFA49AABC" box="[136,1452,476,498]" pageId="2" pageNumber="215">- Fertile fronds more than two times longer than sterile fronds.........................................................................................................22.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFE3BFA4EA966" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFE3BFA4EA943" blockId="2.[136,1452,159,2053]" box="[136,1451,504,527]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFE3BFA4EA943" box="[136,1451,504,527]" pageId="2" pageNumber="215">
21. Fertile laminae oblong to broadly oblong .................................................................................................................
<taxonomicName id="04BE99B1C951A511D439FE3BFA4EA943" ID-CoL="6PFB9" box="[1291,1451,504,526]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="pedunculatus">
<emphasis id="F1CA3E20C951A511D439FE3BFA4EA943" bold="true" box="[1291,1451,504,526]" italics="true" pageId="2" pageNumber="215">L. pedunculatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFDD4FA4EA966" blockId="2.[136,1452,159,2053]" box="[136,1451,533,555]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFDD4FA4EA966" box="[136,1451,533,555]" pageId="2" pageNumber="215">
- Fertile laminae ovate to narrowly ovate ................................................................................................................................
<taxonomicName id="04BE99B1C951A511D462FDD4FA4EA966" ID-CoL="4CYKS" authorityName="Copeland" authorityYear="1914" box="[1360,1451,534,555]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C951A511D462FDD4FA4EA966" bold="true" box="[1360,1451,534,555]" italics="true" pageId="2" pageNumber="215">L. ovatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFDF0FA49A929" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFDF0FA4EA905" blockId="2.[136,1452,159,2053]" box="[136,1451,562,584]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFDF0FA4EA905" box="[136,1451,562,584]" pageId="2" pageNumber="215">
22. Fertile laminae narrowly lanceolate ..........................................................................................................................
<taxonomicName id="04BE99B1C951A511D43EFDF1FA4EA905" ID-CoL="7WX8W" box="[1292,1451,562,584]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="sarawakensis">
<emphasis id="F1CA3E20C951A511D43EFDF1FA4EA905" bold="true" box="[1292,1451,562,584]" italics="true" pageId="2" pageNumber="215">L. sarawakensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFD92FA49A929" blockId="2.[136,1452,159,2053]" box="[136,1452,591,613]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFD92FA49A929" box="[136,1452,591,613]" pageId="2" pageNumber="215">- Fertile laminae broadly lanceolate to ovate.....................................................................................................................................23.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFDAEFA49A9D3" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFDAEFA4EA9CC" blockId="2.[136,1452,159,2053]" box="[136,1451,619,642]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFDAEFA4EA9CC" box="[136,1451,619,642]" pageId="2" pageNumber="215">
23. Sterile laminae elliptic-lanceolate, base gradually narrowed.......................................................................................
<taxonomicName id="04BE99B1C951A511D42EFDAEFA4EA9CC" box="[1308,1451,619,641]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C951A511D42EFDAEFA4EA9CC" bold="true" box="[1308,1451,619,641]" italics="true" pageId="2" pageNumber="215">L. daklakensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFD4BFA49A9D3" blockId="2.[136,1452,159,2053]" box="[136,1452,649,671]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFD4BFA49A9D3" box="[136,1452,649,671]" pageId="2" pageNumber="215">- Sterile laminae ovate-lanceolate to ovate, base truncate .................................................................................................................24.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFD64FA4EA99A" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFD64FA49A9F6" blockId="2.[136,1452,159,2053]" box="[136,1452,677,699]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFD64FA49A9F6" box="[136,1452,677,699]" pageId="2" pageNumber="215">
24. Fertile laminae
<quantity id="04464FD7C951A511D05BFD67FE31A9F6" box="[361,468,677,699]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="2" pageNumber="215" unit="cm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 cm</quantity>
wide, venation indistinct...............................................................................................................
<taxonomicName id="04BE99B1C951A511D457FD64FA49A9F6" box="[1381,1452,677,699]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C951A511D457FD64FA49A9F6" bold="true" box="[1381,1452,677,699]" italics="true" pageId="2" pageNumber="215">L. locii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFD01FA4EA99A" blockId="2.[136,1452,159,2053]" box="[136,1451,706,728]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFD01FA4EA99A" box="[136,1451,706,728]" pageId="2" pageNumber="215">
- Fertile laminae
<quantity id="04464FD7C951A511D05BFD00FE55A995" box="[361,432,706,728]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="2" pageNumber="215" unit="cm" value="5.0" valueMax="7.0" valueMin="3.0">37 cm</quantity>
wide, venation prominent ....................................................................................................................
<taxonomicName id="04BE99B1C951A511D45EFD01FA4EA99A" ID-CoL="6W6ZV" authorityName="Liang Zhang &amp; Li Bing Zhang" baseAuthorityName="C. Christensen" baseAuthorityYear="1933" box="[1388,1451,706,728]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="wui">
<emphasis id="F1CA3E20C951A511D45EFD01FA4EA99A" bold="true" box="[1388,1451,706,728]" italics="true" pageId="2" pageNumber="215">L. wui</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFD1DFA49A85C" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFD1DFA49A9B8" blockId="2.[136,1452,159,2053]" box="[136,1452,735,757]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFD1DFA49A9B8" box="[136,1452,735,757]" pageId="2" pageNumber="215">25. Sori interrupted................................................................................................................................................................................26.</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFD3EFA49A85C" blockId="2.[136,1452,159,2053]" box="[136,1452,764,786]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFD3EFA49A85C" box="[136,1452,764,786]" pageId="2" pageNumber="215">- Sori continuous................................................................................................................................................................................27.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFCDBFA49A806" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFCDBFA4EA863" blockId="2.[136,1452,159,2053]" box="[136,1451,792,815]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFCDBFA4EA863" box="[136,1451,792,815]" pageId="2" pageNumber="215">
26. Circumvascular sheaths present in rhizome, interrupted sori mostly oblong, irregular in shape.............................
<taxonomicName id="04BE99B1C951A511D436FCDBFA4EA863" ID-CoL="3TLF7" authorityName="Nooteboom" authorityYear="1997" baseAuthorityName="C. Presl" baseAuthorityYear="1836" box="[1284,1451,792,814]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="hemionitideus">
<emphasis id="F1CA3E20C951A511D436FCDBFA4EA863" bold="true" box="[1284,1451,792,814]" italics="true" pageId="2" pageNumber="215">L. hemionitideus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFCF4FA49A806" blockId="2.[136,1452,159,2053]" box="[136,1452,821,843]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFCF4FA49A806" box="[136,1452,821,843]" pageId="2" pageNumber="215">
- Circumvascular sheaths absent in rhizome, interrupted sori shortly linear to linear, regular in shape .....................
<taxonomicName id="04BE99B1C951A511D43BFCF4FA49A806" ID-CoL="6PFBH" box="[1289,1452,821,843]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="macrophyllus">
<emphasis id="F1CA3E20C951A511D43BFCF4FA49A806" bold="true" box="[1289,1452,821,843]" italics="true" pageId="2" pageNumber="215">L. macrophyllus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFC90FA49A8C9" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFC90FA4EA825" blockId="2.[136,1452,159,2053]" box="[136,1451,850,872]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFC90FA4EA825" box="[136,1451,850,872]" pageId="2" pageNumber="215">
27. Venation indistinct .................................................................................................................................................................
<taxonomicName id="04BE99B1C951A511D462FC91FA4EA825" ID-CoL="3TLF9" box="[1360,1451,850,872]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="henryi">
<emphasis id="F1CA3E20C951A511D462FC91FA4EA825" bold="true" box="[1360,1451,850,872]" italics="true" pageId="2" pageNumber="215">L. henryi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFCB2FA49A8C9" blockId="2.[136,1452,159,2053]" box="[136,1452,879,901]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFCB2FA49A8C9" box="[136,1452,879,901]" pageId="2" pageNumber="215">- Venation distinct ..............................................................................................................................................................................28.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFC4EFA49A8F3" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFC4EFA49A8EC" blockId="2.[136,1452,159,2053]" box="[136,1452,908,930]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFC4EFA49A8EC" box="[136,1452,908,930]" pageId="2" pageNumber="215">28. Lamina base with outgrows or lobed...............................................................................................................................................29.</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFC6BFA49A8F3" blockId="2.[136,1452,159,2053]" box="[136,1452,937,959]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFC6BFA49A8F3" box="[136,1452,937,959]" pageId="2" pageNumber="215">- Lamina base entire...........................................................................................................................................................................31.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFC04FA49A8BA" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFC04FA4EA896" blockId="2.[136,1452,159,2053]" box="[136,1451,965,987]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFC04FA4EA896" box="[136,1451,965,987]" pageId="2" pageNumber="215">
29. Petioles distinctively winged.........................................................................................................................................
<taxonomicName id="04BE99B1C951A511D411FC04FA4EA896" ID-CoL="3TLNR" box="[1315,1451,965,987]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="hemitomus">
<emphasis id="F1CA3E20C951A511D411FC04FA4EA896" bold="true" box="[1315,1451,965,987]" italics="true" pageId="2" pageNumber="215">L. hemitomus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFC21FA49A8BA" blockId="2.[136,1452,159,2053]" box="[136,1452,994,1016]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFC21FA49A8BA" box="[136,1452,994,1016]" pageId="2" pageNumber="215">- Petioles wingless or slightly winged ...............................................................................................................................................30.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFC3DFA49AF7C" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFC3DFA4EAF58" blockId="2.[136,1452,159,2053]" box="[136,1451,1023,1045]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFC3DFA4EAF58" box="[136,1451,1023,1045]" pageId="2" pageNumber="215">
30. Lamina base slightly lobed.................................................................................................................................................
<taxonomicName id="04BE99B1C951A511D40DFC3DFA4EAF58" ID-CoL="3TLLV" box="[1343,1451,1023,1045]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="saxicola">
<emphasis id="F1CA3E20C951A511D40DFC3DFAABAF59" bold="true" box="[1343,1358,1023,1044]" italics="true" pageId="2" pageNumber="215">L</emphasis>
.
<emphasis id="F1CA3E20C951A511D468FC3DFA4EAF58" bold="true" box="[1370,1451,1023,1045]" italics="true" pageId="2" pageNumber="215">saxicola</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFBDEFA49AF7C" blockId="2.[136,1452,159,2053]" box="[136,1452,1051,1074]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFBDEFA49AF7C" box="[136,1452,1051,1074]" pageId="2" pageNumber="215">
- Laminae distinctly lobed, lobes irregular ....................................................................................................................
<taxonomicName id="04BE99B1C951A511D427FBDEFA49AF7C" ID-CoL="3TLNV" box="[1301,1452,1051,1073]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="shintenensis">
<emphasis id="F1CA3E20C951A511D427FBDEFA49AF7C" bold="true" box="[1301,1452,1051,1073]" italics="true" pageId="2" pageNumber="215">L. shintenensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFBFBFA4EAF26" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFBFBFA4EAF03" blockId="2.[136,1452,159,2053]" box="[136,1451,1080,1103]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFBFBFA4EAF03" box="[136,1451,1080,1103]" pageId="2" pageNumber="215">
31. Laminae widest at middle or above middle part .................................................................................................................
<taxonomicName id="04BE99B1C951A511D476FBFBFA4EAF03" ID-CoL="3TLGP" box="[1348,1451,1080,1102]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="leveillei">
<emphasis id="F1CA3E20C951A511D476FBFBFA4EAF03" bold="true" box="[1348,1451,1080,1102]" italics="true" pageId="2" pageNumber="215">L. leveillei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFB94FA4EAF26" blockId="2.[136,1452,159,2053]" box="[136,1451,1109,1131]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFB94FA4EAF26" box="[136,1451,1109,1131]" pageId="2" pageNumber="215">
- Laminae widest bellow middle part ...................................................................................................................................
<taxonomicName id="04BE99B1C951A511D471FB94FA4EAF26" ID-CoL="3TLNJ" box="[1347,1451,1109,1131]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="wrightii">
<emphasis id="F1CA3E20C951A511D471FB94FA4EAF26" bold="true" box="[1347,1451,1109,1131]" italics="true" pageId="2" pageNumber="215">L. wrightii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFBB0FA49AFE9" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFBB0FA49AFC5" blockId="2.[136,1452,159,2053]" box="[136,1452,1138,1160]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFBB0FA49AFC5" box="[136,1452,1138,1160]" pageId="2" pageNumber="215">
32. Laminae 2-pinnate to 4-pinnatifid .............................................................................................................................
<taxonomicName id="04BE99B1C951A511D43BFBB1FA49AFC5" ID-CoL="3TLFD" box="[1289,1452,1138,1160]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="F1CA3E20C951A511D43BFBB1FA49AFC5" bold="true" box="[1289,1452,1138,1160]" italics="true" pageId="2" pageNumber="215">L. heterophyllus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFB52FA49AFE9" blockId="2.[136,1452,159,2053]" box="[136,1452,1167,1189]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFB52FA49AFE9" box="[136,1452,1167,1189]" pageId="2" pageNumber="215">- Laminae lobed, pinnatifid................................................................................................................................................................33.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFB6EFA49AF93" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFB6EFA49AF8C" blockId="2.[136,1452,159,2053]" box="[136,1452,1196,1218]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFB6EFA49AF8C" box="[136,1452,1196,1218]" pageId="2" pageNumber="215">33. Laminae lobed in different extent, from 2- or 3-forked, pedately lobed, to digitately lobed..........................................................34.</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFB0BFA49AF93" blockId="2.[136,1452,159,2053]" box="[136,1452,1224,1247]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFB0BFA49AF93" box="[136,1452,1224,1247]" pageId="2" pageNumber="215">- Laminae pinnatifid...........................................................................................................................................................................35.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFB24FA4EAE55" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFB24FA4EAFB6" blockId="2.[136,1452,159,2053]" box="[136,1451,1253,1275]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFB24FA4EAFB6" box="[136,1451,1253,1275]" pageId="2" pageNumber="215">
34. Laminae trilobed, lobe base not narrowed ...................................................................................................................
<taxonomicName id="04BE99B1C951A511D416FB24FA4EAFB6" ID-CoL="BDM7R" box="[1316,1451,1253,1275]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="ornithopus">
<emphasis id="F1CA3E20C951A511D416FB24FA4EAFB6" bold="true" box="[1316,1451,1253,1275]" italics="true" pageId="2" pageNumber="215">L. ornithopus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFAC1FA4EAE55" blockId="2.[136,1452,159,2053]" box="[136,1451,1282,1304]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFAC1FA4EAE55" box="[136,1451,1282,1304]" pageId="2" pageNumber="215">
- Laminae often digitately lobed, or if trilobed then lobe base distinctly narrowed............................................................
<taxonomicName id="04BE99B1C951A511D409FAC1FA4EAE55" ID-CoL="3TLDG" box="[1339,1451,1282,1304]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="digitatus">
<emphasis id="F1CA3E20C951A511D409FAC1FA4EAE55" bold="true" box="[1339,1451,1282,1304]" italics="true" pageId="2" pageNumber="215">L. digitatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFADDFA49AE1C" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFADDFA49AE78" blockId="2.[136,1452,159,2053]" box="[136,1452,1311,1333]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFADDFA49AE78" box="[136,1452,1311,1333]" pageId="2" pageNumber="215">
35. Fronds large,
<quantity id="04464FD7C951A511D065FADDFE27AE78" box="[343,450,1311,1333]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="2" pageNumber="215" unit="cm" value="90.0" valueMax="120.0" valueMin="60.0">60120 cm</quantity>
.................................................................................................................................................................36.
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFAFEFA49AE1C" blockId="2.[136,1452,159,2053]" box="[136,1452,1340,1362]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFAFEFA49AE1C" box="[136,1452,1340,1362]" pageId="2" pageNumber="215">
- Fronds less than
<quantity id="04464FD7C951A511D047FAFEFE55AE1C" box="[373,432,1340,1362]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="215" unit="cm" value="60.0">60 cm</quantity>
....................................................................................................................................................................38.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFA9BFA49AEC6" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFA9BFA49AE23" blockId="2.[136,1452,159,2053]" box="[136,1452,1368,1391]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFA9BFA49AE23" box="[136,1452,1368,1391]" pageId="2" pageNumber="215">
36. Lobes often more than 6 pairs, occurs in mountains with acid soils...................................................................................
<taxonomicName id="04BE99B1C951A511D477FA9BFA49AE23" ID-CoL="3TLDV" box="[1349,1452,1368,1390]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="elegans">
<emphasis id="F1CA3E20C951A511D477FA9BFA49AE23" bold="true" box="[1349,1452,1368,1390]" italics="true" pageId="2" pageNumber="215">L. elegans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFAB4FA49AEC6" blockId="2.[136,1452,159,2053]" box="[136,1452,1397,1419]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFAB4FA49AEC6" box="[136,1452,1397,1419]" pageId="2" pageNumber="215">
- Lobes 2
<emphasis id="F1CA3E20C951A511D02DFAB4FECEAEC6" bold="true" box="[287,299,1398,1419]" pageId="2" pageNumber="215"></emphasis>
6 pairs, occurs in limestone mountains.............................................................................................................................37.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFA50FA49AE89" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFA50FA4EAEEA" blockId="2.[136,1452,159,2053]" box="[136,1451,1426,1448]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFA50FA4EAEEA" box="[136,1451,1426,1448]" pageId="2" pageNumber="215">
37. Laminae thinly chartaceous....................................................................................................................................
<taxonomicName id="04BE99B1C951A511D5C9FA51FA4EAEEA" box="[1275,1451,1426,1448]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="sinovietnamica">
<emphasis id="F1CA3E20C951A511D5C9FA51FAEFAEE5" bold="true" box="[1275,1290,1427,1448]" italics="true" pageId="2" pageNumber="215">L</emphasis>
.
<emphasis id="F1CA3E20C951A511D424FA50FA4EAEEA" bold="true" box="[1302,1451,1426,1447]" italics="true" pageId="2" pageNumber="215">sinovietnamica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFA72FA49AE89" blockId="2.[136,1452,159,2053]" box="[136,1452,1455,1477]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFA72FA49AE89" box="[136,1452,1455,1477]" pageId="2" pageNumber="215">
- Laminae thickly chartaceous ........................................................................................................................................
<taxonomicName id="04BE99B1C951A511D42FFA6DFA49AE89" box="[1309,1452,1455,1477]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="wusugongii">
<emphasis id="F1CA3E20C951A511D42FFA6DFAC9AE89" bold="true" box="[1309,1324,1455,1476]" italics="true" pageId="2" pageNumber="215">L</emphasis>
.
<emphasis id="F1CA3E20C951A511D40BFA6DFA49AE89" bold="true" box="[1337,1452,1455,1476]" italics="true" pageId="2" pageNumber="215">wusugongii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAFA0EFA49AEB3" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAFA0EFA49AEAC" blockId="2.[136,1452,159,2053]" box="[136,1452,1483,1506]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFA0EFA49AEAC" box="[136,1452,1483,1506]" pageId="2" pageNumber="215">
38. Sori oblong, close to the lobe margins .............................................................................................................................
<taxonomicName id="04BE99B1C951A511D407FA0EFA49AEAC" ID-CoL="3TLJR" box="[1333,1452,1483,1505]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="oblongus">
<emphasis id="F1CA3E20C951A511D407FA0EFA49AEAC" bold="true" box="[1333,1452,1483,1505]" italics="true" pageId="2" pageNumber="215">L. oblongus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAFA2BFA49AEB3" blockId="2.[136,1452,159,2053]" box="[136,1452,1513,1535]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAFA2BFA49AEB3" box="[136,1452,1513,1535]" pageId="2" pageNumber="215">- Sori linear, rarely oblong, located on main lateral veins between midribs and lobe margins .........................................................39.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF9C4FA49AD7A" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF9C4FA49AD56" blockId="2.[136,1452,159,2053]" box="[136,1452,1541,1563]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF9C4FA49AD56" box="[136,1452,1541,1563]" pageId="2" pageNumber="215">39. Rachis broadly winged ....................................................................................................................................................................40.</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF9E1FA49AD7A" blockId="2.[136,1452,159,2053]" box="[136,1452,1570,1592]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF9E1FA49AD7A" box="[136,1452,1570,1592]" pageId="2" pageNumber="215">- Rachis narrowly winged, partial rachis wingless ............................................................................................................................44.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF9FDFA49AD3C" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF9FDFA49AD18" blockId="2.[136,1452,159,2053]" box="[136,1452,1599,1621]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF9FDFA49AD18" box="[136,1452,1599,1621]" pageId="2" pageNumber="215">
40. Widest of lobes
<quantity id="04464FD7C951A511D05CF9FDFE22AD18" box="[366,455,1599,1621]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="2" pageNumber="215" unit="cm" value="3.25" valueMax="4.0" valueMin="2.5">2.54 cm</quantity>
..............................................................................................................................................
<taxonomicName id="04BE99B1C951A511D417F9FDFA49AD18" ID-CoL="3TLKS" box="[1317,1452,1599,1621]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="pothifolius">
<emphasis id="F1CA3E20C951A511D417F9FDFA49AD18" bold="true" box="[1317,1452,1599,1621]" italics="true" pageId="2" pageNumber="215">L. pothifolius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF99EFA49AD3C" blockId="2.[136,1452,159,2053]" box="[136,1452,1628,1650]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF99EFA49AD3C" box="[136,1452,1628,1650]" pageId="2" pageNumber="215">
- Widest of lobes less than
<quantity id="04464FD7C951A511D0FAF99EFE12AD3C" box="[456,503,1628,1649]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="215" unit="cm" value="2.0">2 cm</quantity>
........................................................................................................................................................41.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF9BBFA49ADE6" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF9BBFA49ADC3" blockId="2.[136,1452,159,2053]" box="[136,1452,1656,1679]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF9BBFA49ADC3" box="[136,1452,1656,1679]" pageId="2" pageNumber="215">
41. Lobes 10
<emphasis id="F1CA3E20C951A511D019F9BBFED2ADC3" bold="true" box="[299,311,1657,1678]" pageId="2" pageNumber="215"></emphasis>
12 pairs....................................................................................................................................................
<taxonomicName id="04BE99B1C951A511D433F9BBFA49ADC3" ID-CoL="3TLDJ" box="[1281,1452,1656,1678]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="dissimilialatus">
<emphasis id="F1CA3E20C951A511D433F9BBFAF5ADC3" bold="true" box="[1281,1296,1657,1678]" italics="true" pageId="2" pageNumber="215">L</emphasis>
.
<emphasis id="F1CA3E20C951A511D42FF9BAFA49ADC3" bold="true" box="[1309,1452,1656,1678]" italics="true" pageId="2" pageNumber="215">dissimilialatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF954FA49ADE6" blockId="2.[136,1452,159,2053]" box="[136,1452,1685,1707]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF954FA49ADE6" box="[136,1452,1685,1707]" pageId="2" pageNumber="215">
- Lobes 6
<emphasis id="F1CA3E20C951A511D02DF954FECEADE6" bold="true" box="[287,299,1686,1707]" pageId="2" pageNumber="215"></emphasis>
9 pairs.................................................................................................................................................................................42
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF970FA49ADA9" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF970FA49AD8A" blockId="2.[136,1452,159,2053]" box="[136,1452,1714,1736]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF970FA49AD8A" box="[136,1452,1714,1736]" pageId="2" pageNumber="215">
42. Basal most pair of lobes forked.................................................................................................................................
<taxonomicName id="04BE99B1C951A511D43BF971FA49AD8A" ID-CoL="XDGT" authorityName="Liang Zhang &amp; Li Bing Zhang" baseAuthorityName="H. G. Zhou &amp; H. Li" baseAuthorityYear="1993" box="[1289,1452,1714,1736]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="sanjiangensis">
<emphasis id="F1CA3E20C951A511D43BF971FA49AD8A" bold="true" box="[1289,1452,1714,1736]" italics="true" pageId="2" pageNumber="215">L. sanjiangensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF912FA49ADA9" blockId="2.[136,1452,159,2053]" box="[136,1452,1743,1765]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF912FA49ADA9" box="[136,1452,1743,1765]" pageId="2" pageNumber="215">- Basal most pair of lobes not forked.................................................................................................................................................43.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF92EFA4EAC53" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF92EFA49AC4C" blockId="2.[136,1452,159,2053]" box="[136,1452,1771,1794]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF92EFA49AC4C" box="[136,1452,1771,1794]" pageId="2" pageNumber="215">
43. Lamia margins flat or sometimes slightly undulate, lobes 1215 ×
<quantity id="04464FD7C951A511D264F92EFC24AC4C" box="[854,961,1772,1793]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="2" pageNumber="215" unit="cm" value="2.15" valueMax="2.4" valueMin="1.9">1.92.4 cm</quantity>
........................................................
<taxonomicName id="04BE99B1C951A511D42BF92EFA49AC4C" box="[1305,1452,1771,1793]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="neolongipes">
<emphasis id="F1CA3E20C951A511D42BF92EFA49AC4C" bold="true" box="[1305,1452,1771,1793]" italics="true" pageId="2" pageNumber="215">L. neolongipes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF8CBFA4EAC53" blockId="2.[136,1452,159,2053]" box="[136,1451,1800,1823]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF8CBFA4EAC53" box="[136,1451,1800,1823]" pageId="2" pageNumber="215">
- Lamina margins distinctly undulate-repand, lobes 512 ×
<quantity id="04464FD7C951A511D3CCF8CBFC8CAC53" box="[766,873,1801,1823]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="2" pageNumber="215" unit="cm" value="1.25" valueMax="1.6" valueMin="0.9">0.91.6 cm</quantity>
...........................................................................
<taxonomicName id="04BE99B1C951A511D41DF8CBFA4EAC53" ID-CoL="6PFJF" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" baseAuthorityName="Christ" baseAuthorityYear="1904" box="[1327,1451,1800,1822]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="flexilobus">
<emphasis id="F1CA3E20C951A511D41DF8CBFA4EAC53" bold="true" box="[1327,1451,1800,1822]" italics="true" pageId="2" pageNumber="215">L. flexilobus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF8E4FA49AC15" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF8E4FA49AC76" blockId="2.[136,1452,159,2053]" box="[136,1452,1829,1851]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF8E4FA49AC76" box="[136,1452,1829,1851]" pageId="2" pageNumber="215">44. Fronds oblong, lobes 58(14) pairs..............................................................................................................................................45.</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF881FA49AC15" blockId="2.[136,1452,159,2053]" box="[136,1452,1858,1880]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF881FA49AC15" box="[136,1452,1858,1880]" pageId="2" pageNumber="215">
- Fronds broad oblong to ovate-deltoid, lobes 1
<emphasis id="F1CA3E20C951A511D3BBF880FD70AC1A" bold="true" box="[649,661,1858,1879]" pageId="2" pageNumber="215"></emphasis>
4 pairs ...................................................................................................................46.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF89DFA4EACDC" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF89DFA4EAC38" blockId="2.[136,1452,159,2053]" box="[136,1451,1887,1909]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF89DFA4EAC38" box="[136,1451,1887,1909]" pageId="2" pageNumber="215">
45. Sori oblong to shortly linear...........................................................................................................................................
<taxonomicName id="04BE99B1C951A511D415F89DFA4EAC38" box="[1319,1451,1887,1909]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="multilobus">
<emphasis id="F1CA3E20C951A511D415F89DFA4EAC38" bold="true" box="[1319,1451,1887,1909]" italics="true" pageId="2" pageNumber="215">L. multilobus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF8BEFA4EACDC" blockId="2.[136,1452,159,2053]" box="[136,1451,1915,1938]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF8BEFA4EACDC" box="[136,1451,1915,1938]" pageId="2" pageNumber="215">
- Sori linear .........................................................................................................................................................................
<taxonomicName id="04BE99B1C951A511D405F8BEFA4EACDC" ID-CoL="3TLDX" box="[1335,1451,1915,1937]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="ellipticus">
<emphasis id="F1CA3E20C951A511D405F8BEFA4EACDC" bold="true" box="[1335,1451,1915,1937]" italics="true" pageId="2" pageNumber="215">L. ellipticus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C951A511D1BAF85BFA49AC86" pageId="2" pageNumber="215">
<paragraph id="C301E232C951A511D1BAF85BFA49ACE3" blockId="2.[136,1452,159,2053]" box="[136,1452,1945,1967]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF85BFA49ACE3" box="[136,1452,1945,1967]" pageId="2" pageNumber="215">
46. Fronds less than
<quantity id="04464FD7C951A511D047F85BFE55ACE3" box="[373,432,1945,1966]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="215" unit="cm" value="18.0">18 cm</quantity>
long ............................................................................................................................................................47.
</keyLead>
</paragraph>
<paragraph id="C301E232C951A511D1BAF874FA49AC86" blockId="2.[136,1452,159,2053]" box="[136,1452,1973,1995]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF874FA49AC86" box="[136,1452,1973,1995]" pageId="2" pageNumber="215">
- Fronds more than
<quantity id="04464FD7C951A511D0B0F874FE58AC86" box="[386,445,1974,1995]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="215" unit="cm" value="20.0">20 cm</quantity>
long..........................................................................................................................................................49.
</keyLead>
</paragraph>
</keyStep>
<paragraph id="C301E232C951A511D1BAF810FA49ACA5" blockId="2.[136,1452,159,2053]" box="[136,1452,2002,2024]" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A511D1BAF810FA49ACA5" box="[136,1452,2002,2024]" pageId="2" pageNumber="215">
<keyStep id="784AFA97C951A511D1BAF810FA49ACA5" box="[136,1452,2002,2024]" pageId="2" pageNumber="215">
47. Sori oblong ..........................................................................................................................................................
<taxonomicName id="04BE99B1C951A511D5D7F811FA49ACA5" box="[1253,1452,2002,2024]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="215" phylum="Tracheophyta" rank="species" species="bolikhamsaiensis">
<emphasis id="F1CA3E20C951A511D5D7F811FA49ACA5" bold="true" box="[1253,1452,2002,2024]" italics="true" pageId="2" pageNumber="215">L. bolikhamsaiensis</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="C301E232C951A510D1BAF832FA4EABF8" blockId="2.[136,1452,159,2053]" lastBlockId="3.[136,1452,159,325]" lastPageId="3" lastPageNumber="216" pageId="2" pageNumber="215">
<keyLead id="784F4107C951A510D1BAF832FA4EABF8" lastPageId="3" lastPageNumber="216" pageId="2" pageNumber="215">
- Sori shortly linear ............................................................................................................................................................................48.
<keyStep id="784AFA97C950A510D1BAFF5DFA4EABF8" box="[136,1451,159,181]" pageId="3" pageNumber="216">
48. Lobes 12 pairs....................................................................................................................................................................
<taxonomicName id="04BE99B1C950A510D475FF5DFA4EABF8" ID-CoL="7WX8R" box="[1351,1451,159,181]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="216" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="F1CA3E20C950A510D475FF5DFA4EABF8" bold="true" box="[1351,1451,159,181]" italics="true" pageId="3" pageNumber="216">L. gracilis</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="C301E232C950A510D1BAFF7EFA4EAB9C" blockId="3.[136,1452,159,325]" box="[136,1451,187,210]" pageId="3" pageNumber="216">
<keyLead id="784F4107C950A510D1BAFF7EFA4EAB9C" box="[136,1451,187,210]" pageId="3" pageNumber="216">
- Lobes 46 pairs............................................................................................................................................................
<taxonomicName id="04BE99B1C950A510D426FF7EFA4EAB9C" ID-CoL="3TLCP" box="[1300,1451,187,209]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="216" phylum="Tracheophyta" rank="species" species="chilangensis">
<emphasis id="F1CA3E20C950A510D426FF7EFA4EAB9C" bold="true" box="[1300,1451,187,209]" italics="true" pageId="3" pageNumber="216">L. chilangensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<keyStep id="784AFA97C950A510D1BAFF1BFA49AA46" pageId="3" pageNumber="216">
<paragraph id="C301E232C950A510D1BAFF1BFA49ABA3" blockId="3.[136,1452,159,325]" box="[136,1452,216,239]" pageId="3" pageNumber="216">
<keyLead id="784F4107C950A510D1BAFF1BFA49ABA3" box="[136,1452,216,239]" pageId="3" pageNumber="216">
49. Laminae chartaceous to thickly chartaceous .....................................................................................................................
<taxonomicName id="04BE99B1C950A510D40FFF1BFA49ABA3" ID-CoL="3TLH4" authorityName="X. C. Zhang" authorityYear="2012" baseAuthorityName="Ching" baseAuthorityYear="1933" box="[1341,1452,216,238]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="216" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="F1CA3E20C950A510D40FFF1BFA49ABA3" bold="true" box="[1341,1452,216,238]" italics="true" pageId="3" pageNumber="216">L. longipes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C950A510D1BAFF34FA49AA46" blockId="3.[136,1452,159,325]" box="[136,1452,245,267]" pageId="3" pageNumber="216">
<keyLead id="784F4107C950A510D1BAFF34FA49AA46" box="[136,1452,245,267]" pageId="3" pageNumber="216">- Laminae herbaceous to thinly chartaceous......................................................................................................................................50.</keyLead>
</paragraph>
</keyStep>
<keyStep id="784AFA97C950A510D1BAFED0FA49AA08" pageId="3" pageNumber="216">
<paragraph id="C301E232C950A510D1BAFED0FA49AA65" blockId="3.[136,1452,159,325]" box="[136,1452,274,296]" pageId="3" pageNumber="216">
<keyLead id="784F4107C950A510D1BAFED0FA49AA65" box="[136,1452,274,296]" pageId="3" pageNumber="216">
50. Petioles the same length or slightly longer than laminae .................................................................................................
<taxonomicName id="04BE99B1C950A510D407FED1FA49AA65" ID-CoL="BTZG3" box="[1333,1452,274,296]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="216" phylum="Tracheophyta" rank="species" species="scandens">
<emphasis id="F1CA3E20C950A510D407FED1FA49AA65" bold="true" box="[1333,1452,274,296]" italics="true" pageId="3" pageNumber="216">L. scandens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C301E232C950A510D1BAFEF2FA49AA08" blockId="3.[136,1452,159,325]" box="[136,1452,303,325]" pageId="3" pageNumber="216">
<keyLead id="784F4107C950A510D1BAFEF2FA49AA08" box="[136,1452,303,325]" pageId="3" pageNumber="216">
- Petioles almost as two times as laminae.....................................................................................................................
<taxonomicName id="04BE99B1C950A510D423FEEDFA49AA08" ID-CoL="3TLKG" box="[1297,1452,303,325]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="216" phylum="Tracheophyta" rank="species" species="pentaphyllus">
<emphasis id="F1CA3E20C950A510D423FEEDFA49AA08" bold="true" box="[1297,1452,303,325]" italics="true" pageId="3" pageNumber="216">L. pentaphyllus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

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<mods:title id="E532A9906C97C9C27F45D3E6AA8DF8DB">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:namePart id="FFA5F1E90ADB23DD71D3699C0A6C3C3F">Zhang, Liang</mods:namePart>
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<mods:namePart id="73416860A651CAF62E5F7FE02E716163">Lu, Ngan Thi</mods:namePart>
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<mods:affiliation id="187E73D41A8F38217958B886E83EA199">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="F6AE8D81CC6AC84EB19652CA41D63A9D">Zhou, Xin-Mao</mods:namePart>
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<mods:affiliation id="8A7059D6E05AE2065B0256A9553285AA">School of Ecology and Environmental Science, Yunnan University, Kunming, Yunnan 650091, China</mods:affiliation>
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<mods:namePart id="0787436300DCE1CD3BC49B9E16C60C69">Zhou, Zhuo</mods:namePart>
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<mods:affiliation id="B5C127CDC3E9BA194620171176118BDF">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier id="039CEB759616E20FEF532386FC61DDCB" type="email">zhouzhuo@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:namePart id="0F10B1525E1C954E49BD752BC9DC0674">Thepkayson, Khamphanh</mods:namePart>
<mods:nameIdentifier id="BE44C373C2C962DC485108A6CBE93B30" type="ORCID">0009-0001-1757-9907</mods:nameIdentifier>
<mods:affiliation id="3173932619837AA31C9F54D9CBAC5129">Institute of Traditional Medicine, Vientiane, Lao PDR</mods:affiliation>
<mods:nameIdentifier id="E6B30BE1D4BB1D5687F8B256667F66EA" type="email">khamphanh@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="3B4C541ECE6F1A31BF5A3E3F7B939374">Luong, Thien Tam</mods:namePart>
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<mods:affiliation id="AEC42E6261FF8C4BC33287E17DE93536">Department of Ecology - Evolutionary Biology, Vietnam National University Ho Chi Minh City (VNUHCM) - University of Science, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</mods:affiliation>
<mods:nameIdentifier id="7AE0B86939B075363B7B9C878467F33E" type="email">lttam1511@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F17B2AE63C25C363704BC5C3DE5EA635">Zhang, Li-Bing</mods:namePart>
<mods:nameIdentifier id="BFC0AE6C97B6D79CC5C444D91F408CB4" type="ORCID">0000-0002-4905-040X</mods:nameIdentifier>
<mods:affiliation id="7CDE9DA651806D11544AD0D96C9F4FFB">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:nameIdentifier id="37DCF0DF9E176F428E8F29258CC9FEBE" type="email">Libing.Zhang@mobot.org</mods:nameIdentifier>
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<mods:title id="B4EE0BB4D078C429E3086B3CCCF1247C">Phytotaxa</mods:title>
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<mods:date id="8154C3514319E75991F89922E4F7F3BA">2024</mods:date>
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<mods:number id="AB9F77A1F4D7A838E43AB54484A27CE0">659</mods:number>
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<mods:number id="88F8ADD738D05B7B84013E78762600C1">3</mods:number>
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<treatment id="4B175324C954A51BD1BAFD78FABAA80D" ID-DOI="http://doi.org/10.5281/zenodo.13652229" ID-GBIF-Taxon="240245556" ID-Zenodo-Dep="13652229" LSID="urn:lsid:plazi:treatment:4B175324C954A51BD1BAFD78FABAA80D" httpUri="http://treatment.plazi.org/id/4B175324C954A51BD1BAFD78FABAA80D" lastPageId="8" lastPageNumber="221" pageId="7" pageNumber="220">
<subSubSection id="8BA4B1B9C954A514D1BAFD78FC3AA998" box="[136,991,698,725]" pageId="7" pageNumber="220" type="nomenclature">
<paragraph id="C301E232C954A514D1BAFD78FC3AA998" blockId="7.[136,991,698,725]" box="[136,991,698,725]" pageId="7" pageNumber="220">
<heading id="9849555EC954A514D1BAFD78FC3AA998" box="[136,991,698,725]" fontSize="11" level="2" pageId="7" pageNumber="220" reason="4">
3.
<taxonomicName id="04BE99B1C954A514D196FD78FCF8A999" authority="Liang Zhang &amp; Li Bing Zhang" authorityName="Liang Zhang &amp; Li Bing Zhang" box="[164,797,698,725]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="kachinensis" status="sp. nov.">
<emphasis id="F1CA3E20C954A514D196FD78FE5EA998" bold="true" box="[164,443,698,725]" italics="true" pageId="7" pageNumber="220">Leptochilus kachinensis</emphasis>
Liang Zhang &amp; Li Bing Zhang
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,
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<taxonomicNameLabel id="EAF9835BC954A514D21BFD79FC9BA999" box="[809,894,699,724]" pageId="7" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5B85FEB7C954A514D2BEFD78FC34A998" box="[908,977,698,725]" captionStart="FIGURE 5" captionStartId="7.[136,229,2019,2041]" captionTargetBox="[163,1424,905,1982]" captionTargetId="figure-290@7.[151,1436,893,1994]" captionTargetPageId="7" captionText="FIGURE 5. Leptochilus kachinensis Liang Zhang &amp; Li Bing Zhang,—A. Habit.—B. Venation.—C. Rhizome." figureDoi="http://doi.org/10.5281/zenodo.13217904" httpUri="https://zenodo.org/record/13217904/files/figure.png" pageId="7" pageNumber="220">Fig. 5</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C954A514D1BAFCC0FD54A829" pageId="7" pageNumber="220" type="materials_examined">
<paragraph id="C301E232C954A514D1BAFCC0FD54A829" blockId="7.[136,1451,770,868]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D1BAFCC0FF26A851" bold="true" box="[136,195,770,796]" pageId="7" pageNumber="220">Type</emphasis>
:
<collectingCountry id="BBA9A2A2C954A514D1E5FCC0FE88A851" box="[215,365,770,796]" name="Myanmar" pageId="7" pageNumber="220">MYANMAR</collectingCountry>
.
<collectingRegion id="017A2CD0C954A514D04EFCC0FDF0A851" box="[380,533,770,796]" country="Myanmar" name="Kachin" pageId="7" pageNumber="220">Kachin State</collectingRegion>
: Putero District, Putao Town, Hpaw Wang Daru, N of Sum Pi Yang,
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,
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, on the tree trunk,
<date id="B700C4F2C954A514D31FFCE4FD37A80D" box="[557,722,806,832]" pageId="7" pageNumber="220" value="2009-04-12">12 April 2009</date>
,
<emphasis id="F1CA3E20C954A514D3D2FCE5FE8AA829" italics="true" pageId="7" pageNumber="220">
Yun-Fei Deng, Chang-Chun Ding &amp; Lin-Fang Wu,
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Expedition No. 3200
</emphasis>
(
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CDBI-0193822!).
</paragraph>
</subSubSection>
<caption id="97C1B2BAC954A514D1BAF821FB4BACB4" ID-DOI="http://doi.org/10.5281/zenodo.13217904" ID-Zenodo-Dep="13217904" box="[136,1198,2019,2041]" httpUri="https://zenodo.org/record/13217904/files/figure.png" pageId="7" pageNumber="220" startId="7.[136,229,2019,2041]" targetBox="[163,1424,905,1982]" targetPageId="7" targetType="figure">
<paragraph id="C301E232C954A514D1BAF821FB4BACB4" blockId="7.[136,1198,2019,2041]" box="[136,1198,2019,2041]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D1BAF821FF12ACB5" bold="true" box="[136,247,2019,2041]" pageId="7" pageNumber="220">FIGURE 5</emphasis>
.
<taxonomicName id="04BE99B1C954A514D031F821FCFDACB4" authority="Liang Zhang &amp; Li Bing Zhang" authorityName="Liang Zhang &amp; Li Bing Zhang" box="[259,792,2019,2041]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis id="F1CA3E20C954A514D031F821FE0CACB5" box="[259,489,2019,2040]" italics="true" pageId="7" pageNumber="220">Leptochilus kachinensis</emphasis>
Liang Zhang &amp; Li Bing Zhang
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,—A. Habit.—B. Venation.—C. Rhizome.
</paragraph>
</caption>
<subSubSection id="8BA4B1B9C95BA51BD18FFF5CFDDDA925" pageId="8" pageNumber="221" type="diagnosis">
<paragraph id="C301E232C95BA51BD18FFF5CFF1FAA4D" blockId="8.[136,1452,158,833]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFF5CFED5ABF5" bold="true" box="[189,304,158,184]" pageId="8" pageNumber="221">Diagnosis</emphasis>
:
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<emphasis id="F1CA3E20C95BA51BD00DFF5CFDAEABF5" box="[319,587,158,184]" italics="true" pageId="8" pageNumber="221">Leptochilus kachinensis</emphasis>
</taxonomicName>
is distinguished by its dimorphic fronds, fertile fronds nearly the same length as sterile fronds, sterile laminae lanceolate, up to
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long, thinly chartaceous, and venation distinct on both sides of lamina.
</paragraph>
<paragraph id="C301E232C95BA51BD18FFEC8FDDDA925" blockId="8.[136,1452,158,833]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFEC8FEEDAA69" bold="true" box="[189,264,266,292]" pageId="8" pageNumber="221">Plants</emphasis>
perennial, evergreen, epiphytic. Rhizome long creeping, robust, ca.
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in diam., dorsiventrally flattened, with a few short phyllopodia and roots, sparsely scaly, scales lanceolate, 1.94.5 ×
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, brown, basifixed, concolorous, clathrate, margin entire or sparely denticulate, apex acuminate.
<emphasis id="F1CA3E20C95BA51BD55CFE90FB26AA21" bold="true" box="[1134,1219,338,364]" pageId="8" pageNumber="221">Fronds</emphasis>
strongly dimorphic, fertile fronds shorter or nearly the same length as sterile frond.
<emphasis id="F1CA3E20C95BA51BD25FFEB4FBF7AADD" bold="true" box="[877,1042,374,400]" pageId="8" pageNumber="221">Sterile fronds</emphasis>
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tall, petiole shorter than lamina;
<emphasis id="F1CA3E20C95BA51BD1D7FE58FEA1AAF9" bold="true" box="[229,324,410,436]" pageId="8" pageNumber="221">laminae</emphasis>
ribbon-lanceolate, margins entire, slightly repand, base cuneate, decurrent, apex acuminate,
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long,
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wide at middle, thinly chartaceous, glabrous;
<emphasis id="F1CA3E20C95BA51BD245FE7CFC34AA95" bold="true" box="[887,977,446,472]" pageId="8" pageNumber="221">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
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in diam.,
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long;
<emphasis id="F1CA3E20C95BA51BD372FE20FD42AAB1" bold="true" box="[576,679,482,508]" pageId="8" pageNumber="221">venation</emphasis>
anastomosing, lateral veins distinct, veinlets visible.
<emphasis id="F1CA3E20C95BA51BD434FE20FA4EAAB1" bold="true" box="[1286,1451,482,508]" pageId="8" pageNumber="221">Fertile fronds</emphasis>
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tall, petioles the same length or longer than laminae;
<emphasis id="F1CA3E20C95BA51BD259FDC4FC2FA96D" bold="true" box="[875,970,518,544]" pageId="8" pageNumber="221">laminae</emphasis>
linear, margins entire, base cuneate, apex long acuminate,
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long, 0.81.0 mm wide at middle;
<emphasis id="F1CA3E20C95BA51BD264FDE8FC55A909" bold="true" box="[854,944,554,580]" pageId="8" pageNumber="221">petioles</emphasis>
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in diam.,
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long.
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acrostichoid, continuous along midrib.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95BA51BD18FFDB0FBA0A9FD" pageId="8" pageNumber="221" type="distribution">
<paragraph id="C301E232C95BA51BD18FFDB0FBA0A9FD" blockId="8.[136,1452,158,833]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFDB0FE15A9C1" bold="true" box="[189,496,626,652]" pageId="8" pageNumber="221">Distribution and habitat:</emphasis>
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<emphasis id="F1CA3E20C95BA51BD0C9FDB0FCEEA9C1" box="[507,779,626,652]" italics="true" pageId="8" pageNumber="221">Leptochilus kachinensis</emphasis>
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is currently only known from the
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locality in
<collectingRegion id="017A2CD0C95BA51BD468FDB0FF26A9FD" country="Myanmar" name="Kachin" pageId="8" pageNumber="221">Kachin State</collectingRegion>
, located in northern
<collectingCountry id="BBA9A2A2C95BA51BD09BFD55FDFDA9FC" box="[425,536,663,689]" name="Myanmar" pageId="8" pageNumber="221">Myanmar</collectingCountry>
. The species was observed growing on tree trunk.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95BA51BD18FFD78FABAA80D" pageId="8" pageNumber="221" type="etymology">
<paragraph id="C301E232C95BA51BD18FFD78FB4FA999" blockId="8.[136,1452,158,833]" box="[189,1194,698,725]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFD78FEDBA999" bold="true" box="[189,318,698,724]" pageId="8" pageNumber="221">Etymology</emphasis>
: The species epithet is derived from
<collectingRegion id="017A2CD0C95BA51BD3EEFD78FC8AA999" box="[732,879,698,724]" country="Myanmar" name="Kachin" pageId="8" pageNumber="221">Kachin State</collectingRegion>
, where the species is found.
</paragraph>
<paragraph id="C301E232C95BA51BD18FFD1CFABAA80D" blockId="8.[136,1452,158,833]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFD1CFF10A9B5" bold="true" box="[189,245,734,760]" pageId="8" pageNumber="221">Note</emphasis>
: The phylogenetic analysis resolved
<taxonomicName id="04BE99B1C95BA51BD3A4FD1CFC46A9B5" box="[662,931,734,760]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="kachinensis">
<emphasis id="F1CA3E20C95BA51BD3A4FD1CFC46A9B5" box="[662,931,734,760]" italics="true" pageId="8" pageNumber="221">Leptochilus kachinensis</emphasis>
</taxonomicName>
as the sister species to the
<taxonomicName id="04BE99B1C95BA51BD5E1FD1CFAC0A9B5" authorityName="K.B.Presl" authorityYear="1851" box="[1235,1317,734,760]" class="Polypodiopsida" family="Polypodiaceae" genus="Colysis" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="genus">
<emphasis id="F1CA3E20C95BA51BD5E1FD1CFAC0A9B5" box="[1235,1317,734,760]" italics="true" pageId="8" pageNumber="221">Colysis</emphasis>
</taxonomicName>
subclade in the
<taxonomicName id="04BE99B1C95BA51BD180FCC0FEE1A851" authorityName="K.B.Presl" authorityYear="1851" box="[178,260,770,796]" class="Polypodiopsida" family="Polypodiaceae" genus="Colysis" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="genus">
<emphasis id="F1CA3E20C95BA51BD180FCC0FEE1A851" box="[178,260,770,796]" italics="true" pageId="8" pageNumber="221">Colysis</emphasis>
</taxonomicName>
clade (
<figureCitation id="5B85FEB7C95BA51BD06AFCC0FE7FA851" box="[344,410,770,796]" captionStart="FIGURE 1" captionStartId="3.[136,229,1956,1978]" captionTargetBox="[153,1430,353,1922]" captionTargetId="figure-76@3.[151,1435,347,1928]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of Leptochilus based on six plastid markers (modified from Zhang et al. 2024). Orange color indicates the new species described here." figureDoi="http://doi.org/10.5281/zenodo.13217896" httpUri="https://zenodo.org/record/13217896/files/figure.png" pageId="8" pageNumber="221">Fig. 1</figureCitation>
). The close relationship between the new species and
<taxonomicName id="04BE99B1C95BA51BD530FCC1FE72A80C" authority="(C. Presl 1836: 216) Nooteboom (1997: 285)" authorityName="Nooteboom" authorityPageNumber="285" authorityYear="1997" baseAuthorityName="C. Presl" baseAuthorityPageNumber="216" baseAuthorityYear="1836" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="hemionitideus">
<emphasis id="F1CA3E20C95BA51BD530FCC1FB5BA851" box="[1026,1214,770,796]" italics="true" pageId="8" pageNumber="221">L. hemionitideus</emphasis>
(C.
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)
<bibRefCitation id="A72F9FC3C95BA51BD1BAFCE4FE72A80C" author="Nooteboom, H. P." box="[136,407,806,833]" pageId="8" pageNumber="221" pagination="261 - 395" refId="ref11978" refString="Nooteboom, H. P. (1997) The microsoroid ferns (Polypodiaceae). Blumea 42: 261 - 395." type="journal article" year="1997">Nooteboom (1997: 285)</bibRefCitation>
</taxonomicName>
is confirmed by both of them having herbaceous sterile laminae and distinct venation.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,272 @@
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<mods:title id="8E53B057EFFBE6902FD6DE344341F90F">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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2.
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<emphasis id="F1CA3E20C955A515D196FF5CFE5CABF4" bold="true" box="[164,441,158,185]" italics="true" pageId="6" pageNumber="219">Leptochilus daklakensis</emphasis>
Liang Zhang, X.M. Zhou, T.T. Luong &amp; Li Bing Zhang
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,
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(
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. Đắk Lắk: Chu Yang Sin National Park, Sub-area 1351,
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,
<geoCoordinate id="A68A84F5C955A515D556FF24FB27AA4D" box="[1124,1218,230,256]" degrees="12.41" direction="north" orientation="latitude" pageId="6" pageNumber="219" precision="555" value="12.41">12.41°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C955A515D5F9FF24FAD3AA4D" box="[1227,1334,230,256]" degrees="108.37" direction="east" orientation="longitude" pageId="6" pageNumber="219" precision="555" value="108.37">108.37°E</geoCoordinate>
, on rocks, small branch of Krông Kmar River,
<date id="B700C4F2C955A515D325FEC8FD31AA68" box="[535,724,266,293]" pageId="6" pageNumber="219" value="2015-10-30">30 October 2015</date>
,
<emphasis id="F1CA3E20C955A515D3ECFEC9FA4EAA69" box="[734,1451,266,292]" italics="true" pageId="6" pageNumber="219">Li Bing Zhang, Xin Mao Zhou, Thien Tam Luong, Minh Tri Dang</emphasis>
&amp;
<emphasis id="F1CA3E20C955A515D197FEECFE71AA05" box="[165,404,302,328]" italics="true" pageId="6" pageNumber="219">Bich Ha Duong 8944</emphasis>
(
<typeStatus id="1C055C90C955A515D090FEECFDE3AA05" box="[418,518,302,328]" pageId="6" pageNumber="219" type="holotype">holotype</typeStatus>
CDBI!,
<typeStatus id="1C055C90C955A515D355FEECFD26AA05" box="[615,707,302,328]" pageId="6" pageNumber="219" type="isotype">isotypes</typeStatus>
PHH, MO!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C955A514D18FFE90FE6BAA21" lastPageId="7" lastPageNumber="220" pageId="6" pageNumber="219" type="diagnosis">
<paragraph id="C301E232C955A515D18FFE90FC9CAAF9" blockId="6.[136,1452,230,437]" pageId="6" pageNumber="219">
<emphasis id="F1CA3E20C955A515D18FFE90FED5AA21" bold="true" box="[189,304,338,364]" pageId="6" pageNumber="219">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C955A515D00DFE90FDAEAA21" box="[319,587,338,364]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="6" pageNumber="219" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C955A515D00DFE90FDAEAA21" box="[319,587,338,364]" italics="true" pageId="6" pageNumber="219">Leptochilus daklakensis</emphasis>
</taxonomicName>
is distinguished from other species by its dimorphic fronds, fertile fronds two times longer than sterile fronds, sterile fronds with broadly winged petioles, petioles of fertile fronds four times longer than fertile laminae, and apex of fertile laminae abruptly narrowed.
</paragraph>
<caption id="97C1B2BAC955A515D1BAF979FDBBADBB" ID-DOI="http://doi.org/10.5281/zenodo.13217902" ID-Zenodo-Dep="13217902" httpUri="https://zenodo.org/record/13217902/files/figure.png" pageId="6" pageNumber="219" startId="6.[136,229,1723,1745]" targetBox="[163,1424,473,1687]" targetPageId="6" targetType="figure">
<paragraph id="C301E232C955A515D1BAF979FDBBADBB" blockId="6.[136,1451,1723,1782]" pageId="6" pageNumber="219">
<emphasis id="F1CA3E20C955A515D1BAF979FF12AD9D" bold="true" box="[136,247,1723,1745]" pageId="6" pageNumber="219">FIGURE 4</emphasis>
.
<taxonomicName id="04BE99B1C955A515D031F97EFC68AD9F" authority="Liang Zhang, X. M. Zhou &amp; Li Bing Zhang" authorityName="Liang Zhang, X. M. Zhou &amp; Li Bing Zhang" box="[259,909,1724,1746]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="6" pageNumber="219" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C955A515D031F97EFE0CAD9C" box="[259,489,1724,1745]" italics="true" pageId="6" pageNumber="219">Leptochilus daklakensis</emphasis>
Liang Zhang, X.M. Zhou &amp; Li Bing Zhang
</taxonomicName>
,—A, B, C. Habit.—D. Abaxial view of fertile lamina.— E. Rhizome scales. Photo credit: Xin-Mao Zhou.
</paragraph>
</caption>
<paragraph id="C301E232C955A514D18FF8E6FE6BAA21" blockId="6.[136,1452,1828,2071]" lastBlockId="7.[136,1452,158,652]" lastPageId="7" lastPageNumber="220" pageId="6" pageNumber="219">
<emphasis id="F1CA3E20C955A515D18FF8E6FEEDAC73" bold="true" box="[189,264,1828,1854]" pageId="6" pageNumber="219">Plants</emphasis>
perennial, evergreen, lithophytic.
<emphasis id="F1CA3E20C955A515D34DF8E6FD0CAC73" bold="true" box="[639,745,1828,1854]" pageId="6" pageNumber="219">Rhizome</emphasis>
long creeping, ca.
<quantity id="04464FD7C955A515D286F8E6FBDCAC73" box="[948,1081,1828,1855]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="6" pageNumber="219" unit="mm" value="3.05" valueMax="3.5" valueMin="2.6">2.63.5 mm</quantity>
in diam., dorsiventrally flattened, densely scaly, scales lanceolate, 1.23.5 ×
<quantity id="04464FD7C955A515D35CF88AFD1DAC2F" box="[622,760,1864,1890]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="6" pageNumber="219" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.40.7 mm</quantity>
, dark brown, basifixed, concolorous, clathrate, margin entire or sparely denticulate, apex acuminate.
<emphasis id="F1CA3E20C955A515D365F8AEFD49ACCB" bold="true" box="[599,684,1900,1926]" pageId="6" pageNumber="219">Fronds</emphasis>
dimorphic, fertile fronds usually much longer than sterile fronds, or rarely the same length as sterile fronds.
<emphasis id="F1CA3E20C955A515D345F852FCFEACE7" bold="true" box="[631,795,1936,1962]" pageId="6" pageNumber="219">Sterile fronds</emphasis>
(6)
<quantity id="04464FD7C955A515D260F852FC20ACE7" box="[850,965,1936,1962]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="6" pageNumber="219" unit="cm" value="20.0" valueMax="22.0" valueMin="18.0">1822 cm</quantity>
tall, sessile or shortly petiolated;
<emphasis id="F1CA3E20C955A515D47EF852FA4EACE7" bold="true" box="[1356,1451,1936,1962]" pageId="6" pageNumber="219">laminae</emphasis>
elliptic-lanceolate, base gradually narrowed, apex acuminate or cuspidate, (6)
<quantity id="04464FD7C955A515D2CCF876FB8AAC83" box="[1022,1135,1972,1998]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="6" pageNumber="219" unit="cm" value="20.0" valueMax="22.0" valueMin="18.0">1822 cm</quantity>
long,
<quantity id="04464FD7C955A515D58AF876FAD0AC83" box="[1208,1333,1972,1998]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="6" pageNumber="219" unit="cm" value="3.05" valueMax="3.2" valueMin="2.9">2.93.2 cm</quantity>
wide near middle, chartaceous, glabrous;
<emphasis id="F1CA3E20C955A515D0D7F81AFDDAACBF" bold="true" box="[485,575,2008,2034]" pageId="6" pageNumber="219">petioles</emphasis>
short, stramineous, grooved adaxially,
<quantity id="04464FD7C955A515D2C5F81AFB9BACBF" box="[1015,1150,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="6" pageNumber="219" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
in diam.,
<quantity id="04464FD7C955A515D5DCF81AFA8EACBF" box="[1262,1387,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="6" pageNumber="219" unit="cm" value="0.5" valueMax="0.7" valueMin="0.3">0.30.7 cm</quantity>
long, petiole base scaly; scales similar to rhizome scales but smaller, 1.31.5 ×
<quantity id="04464FD7C955A515D2F4F83EFBB6A35B" box="[966,1107,2044,2071]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="219" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.40.5 mm</quantity>
.
<emphasis id="F1CA3E20C955A515D56DF83EFAE1A35B" bold="true" box="[1119,1284,2044,2070]" pageId="6" pageNumber="219">Fertile fronds</emphasis>
(16)
<quantity id="04464FD7C955A515D40AF83EFA4EA35B" box="[1336,1451,2044,2071]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="6" pageNumber="219" unit="cm" value="38.0" valueMax="41.0" valueMin="35.0">3541 cm</quantity>
tall, petiole much longer than lamina;
<emphasis id="F1CA3E20C954A514D307FF5CFD71ABF5" bold="true" box="[565,660,158,184]" pageId="7" pageNumber="220">laminae</emphasis>
deltate-lanceolate, margins entire, base cuneate, apex long acuminate, 5.49.0 cm long,
<quantity id="04464FD7C954A514D078FF00FE23AB90" box="[330,454,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="4.1" metricValueMin="2.2" pageId="7" pageNumber="220" unit="cm" value="3.15" valueMax="4.1" valueMin="2.2">2.24.1 cm</quantity>
wide bellow middle part, chartaceous, glabrous;
<emphasis id="F1CA3E20C954A514D2DAFF00FBA7AB91" bold="true" box="[1000,1090,194,220]" pageId="7" pageNumber="220">petioles</emphasis>
stramineous, grooved adaxially,
<quantity id="04464FD7C954A514D1BAFF24FEEAAA4C" box="[136,271,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="7" pageNumber="220" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.91.1 mm</quantity>
in diam., (11)
<quantity id="04464FD7C954A514D084FF24FDCDAA4C" box="[438,552,230,257]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="7" pageNumber="220" unit="cm" value="29.5" valueMax="32.0" valueMin="27.0">2732 cm</quantity>
long.
<emphasis id="F1CA3E20C954A514D35CFF24FD4BAA4D" bold="true" box="[622,686,230,256]" pageId="7" pageNumber="220">Veins</emphasis>
anastomosing, distinct adaxially and abaxially, veinlets visible.
<emphasis id="F1CA3E20C954A514D448FF24FA49AA4D" bold="true" box="[1402,1452,230,256]" pageId="7" pageNumber="220">Sori</emphasis>
linear, reach lamina margins, ca.
<quantity id="04464FD7C954A514D337FEC8FDB8AA68" box="[517,605,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="220" unit="mm" value="0.8">0.8 mm</quantity>
from margins and
<quantity id="04464FD7C954A514D208FEC8FC4AAA68" box="[826,943,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="7" pageNumber="220" unit="mm" value="1.75" valueMax="2.5" valueMin="1.0">12.5 mm</quantity>
from midrib, 1523 on each side of midrib,
<quantity id="04464FD7C954A514D1BAFEECFEE0AA04" box="[136,261,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.9" metricValueMin="1.2" pageId="7" pageNumber="220" unit="cm" value="2.05" valueMax="2.9" valueMin="1.2">1.22.9 cm</quantity>
long,
<quantity id="04464FD7C954A514D07EFEECFE36AA04" box="[332,467,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="7" pageNumber="220" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.40.6 mm</quantity>
wide when mature, at angles of 4247 degree with midribs, centers ca. (1)1.53.0 mm apart from one another.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C954A514D18FFEB4FCB5AAF9" pageId="7" pageNumber="220" type="distribution">
<paragraph id="C301E232C954A514D18FFEB4FCB5AAF9" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D18FFEB4FE09AADD" bold="true" box="[189,492,374,400]" pageId="7" pageNumber="220">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C954A514D0C6FEB4FCE7AADD" box="[500,770,374,400]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C954A514D0C6FEB4FCE7AADD" box="[500,770,374,400]" italics="true" pageId="7" pageNumber="220">Leptochilus daklakensis</emphasis>
</taxonomicName>
is currently known from south Đắk Lắk Province,
<collectingCountry id="BBA9A2A2C954A514D474FEB4FA4CAADD" box="[1350,1449,374,400]" name="Vietnam" pageId="7" pageNumber="220">Vietnam</collectingCountry>
. The species was observed only growing on rocks in rainforests.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C954A514D18FFE7CFD26A9C1" pageId="7" pageNumber="220" type="etymology">
<paragraph id="C301E232C954A514D18FFE7CFC30AAB1" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D18FFE7CFEDBAA95" bold="true" box="[189,318,446,472]" pageId="7" pageNumber="220">Etymology</emphasis>
: The species epithet is from the Vietnamese name,
<emphasis id="F1CA3E20C954A514D24AFE7CFC26AA95" box="[888,963,446,472]" italics="true" pageId="7" pageNumber="220">daklak</emphasis>
, the name of the province in South
<collectingCountry id="BBA9A2A2C954A514D47FFE7CFA49AA95" box="[1357,1452,446,472]" name="Vietnam" pageId="7" pageNumber="220">Vietnam</collectingCountry>
where the
<typeStatus id="1C055C90C954A514D1CFFE21FEC8AAB0" box="[253,301,483,509]" pageId="7" pageNumber="220">type</typeStatus>
was collected, and the Latin suffix
<emphasis id="F1CA3E20C954A514D38EFE21FD18AAB1" box="[700,765,483,508]" italics="true" pageId="7" pageNumber="220">-ensis</emphasis>
, of origin or place.
</paragraph>
<paragraph id="C301E232C954A514D18FFDC4FD26A9C1" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D18FFDC4FEE5A96D" bold="true" box="[189,256,518,544]" pageId="7" pageNumber="220">Notes</emphasis>
:
<taxonomicName id="04BE99B1C954A514D03DFDC4FDFFA96D" box="[271,538,518,544]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C954A514D03DFDC4FDFFA96D" box="[271,538,518,544]" italics="true" pageId="7" pageNumber="220">Leptochilus daklakensis</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="04BE99B1C954A514D3D8FDC5FCAAA96D" authorityName="Copeland" authorityYear="1914" box="[746,847,519,544]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C954A514D3D8FDC5FCAAA96D" box="[746,847,519,544]" italics="true" pageId="7" pageNumber="220">L. ovatus</emphasis>
</taxonomicName>
but differs the latter in having broadly winged petioles (vs. narrowly wringed petioles, or sometimes petioles wingless), apex of fertile laminae abruptly narrowed (vs. apex of fertile laminae rounded or gradually narrowed). Phylogenetically,
<taxonomicName id="04BE99B1C954A514D2B6FD8DFBC6A925" box="[900,1059,590,616]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C954A514D2B6FD8DFBC6A925" box="[900,1059,590,616]" italics="true" pageId="7" pageNumber="220">L. daklakensis</emphasis>
</taxonomicName>
was resolved as an isolated lineage quite distant from
<taxonomicName id="04BE99B1C954A514D064FDB1FE58A9C1" authorityName="Copeland" authorityYear="1914" box="[342,445,627,652]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C954A514D064FDB1FE58A9C1" box="[342,445,627,652]" italics="true" pageId="7" pageNumber="220">L. ovatus</emphasis>
</taxonomicName>
in the
<emphasis id="F1CA3E20C954A514D338FDB1FD9CA9C1" box="[522,633,626,652]" italics="true" pageId="7" pageNumber="220">
L.
<taxonomicName id="04BE99B1C954A514D315FDB0FD9CA9C1" authorityName="K.B.Presl" authorityYear="1851" box="[551,633,626,652]" class="Polypodiopsida" family="Polypodiaceae" genus="Colysis" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="genus">Colysis</taxonomicName>
</emphasis>
clade.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,258 @@
<document id="E43D4E652ED4D683CB4911AD43E841C8" ID-CLB-Dataset="302193" ID-DOI="10.11646/phytotaxa.659.3.1" ID-GBIF-Dataset="52524fe3-dfb8-4afc-829a-31089bb4e306" ID-ISSN="1179-3163" ID-Zenodo-Dep="13217894" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1722613780751" checkinUser="felipe" docAuthor="Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, Zhou, Zhuo, Thepkayson, Khamphanh, Luong, Thien Tam &amp; Zhang, Li-Bing" docDate="2024" docId="4B175324C957A516D1BAFB6AFB3EAC93" docLanguage="en" docName="phytotaxa.659.3.1.pdf" docOrigin="Phytotaxa 659 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.659.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Leptochilus bolikhamsaiensis Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="218" masterDocId="B72E2B5CC953A513D132FFC2FFE5AB4D" masterDocTitle="Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot" masterLastPageNumber="235" masterPageNumber="213" pageNumber="217" updateTime="1725399130512" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8849B7374B6D2D746BC658D5CF958011">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
</mods:titleInfo>
<mods:name id="493B016EF14B9193A5DA0346ACAB9A2E" type="personal">
<mods:role id="7F940F5B3B4429BA560F2001E449739D">
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<mods:namePart id="36776C029602C585BD26DAD1FCDD50E7">Zhang, Liang</mods:namePart>
<mods:nameIdentifier id="B58DC3ECC6AAE60DF0BEE10AC7615DE0" type="ORCID">0000-0003-3784-3135</mods:nameIdentifier>
<mods:affiliation id="710D06CD6976A37F09A761A42FA132CC">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier id="CFB61E9F8440231953BCB8CB13997305" type="email">zhangliang@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:namePart id="D452FC26D46A82176220AD42424490B4">Lu, Ngan Thi</mods:namePart>
<mods:nameIdentifier id="759A8D1FD78246DE30DC23D5366D6D55" type="ORCID">0000-0001-9823-2018</mods:nameIdentifier>
<mods:affiliation id="C30BAAE4B80872912C3B9BEEA50B1BF9">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="A46B71002A19A2F0449DB3EF857CC5B5" type="email">vnmngan@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="CCADA348E871D8E21EC6D45F7F4A58A1" type="personal">
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</mods:role>
<mods:namePart id="95AFC2238FD35569D61A06CD7442E738">Zhou, Xin-Mao</mods:namePart>
<mods:nameIdentifier id="5C5897721907865B4C2962803D6E9962" type="ORCID">0000-0003-3555-7784</mods:nameIdentifier>
<mods:affiliation id="E891D1E7E671EB0E7C48EE935D4B083D">School of Ecology and Environmental Science, Yunnan University, Kunming, Yunnan 650091, China</mods:affiliation>
<mods:nameIdentifier id="3D55403885A6377DA3D57927D5B3FB22" type="email">xinmao.zhou@ynu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="B5830E80C6B5E67D76B5028FADAF6DD1" type="personal">
<mods:role id="C58DE3BC19E5C289BE64AA54E65638FB">
<mods:roleTerm id="AFEB1CEC6C28CDFC03E17277CA9A47E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25245C6C1DD8F3FFC76C9F391CC7ED0B">Zhou, Zhuo</mods:namePart>
<mods:nameIdentifier id="5E5C1D5ECF8D266E831D46CF8892969F" type="ORCID">0000-0001-9682-5717</mods:nameIdentifier>
<mods:affiliation id="5FB75C44A91AF08477D1D9D4ECFE7B6C">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier id="B614AF206F0E111E0DFB0A668577D774" type="email">zhouzhuo@mail.kib.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="31CF58B9333A62C966457D1CB5E1F722" type="personal">
<mods:role id="D6A7FBD4AD24330490231C7441889F26">
<mods:roleTerm id="76F594874F622D62BEB2BC4001275636">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36BF46FDFF442A71BE8EF842C06BB54A">Thepkayson, Khamphanh</mods:namePart>
<mods:nameIdentifier id="816C643FEC4B8595E180DE00C7C02A0C" type="ORCID">0009-0001-1757-9907</mods:nameIdentifier>
<mods:affiliation id="D30EBEE56554183D8F1EFF6EBB35869E">Institute of Traditional Medicine, Vientiane, Lao PDR</mods:affiliation>
<mods:nameIdentifier id="14940ABDB1729C1CABCA0115E3C950E7" type="email">khamphanh@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="305B8BD3E2E0015C11A793B9AC7E0306" type="personal">
<mods:role id="F2E5CA63528E38D9696F0EAF18F555A3">
<mods:roleTerm id="41B75A98F3180DF2CB4C8D713F713C3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="309B0831F24EEE7EB20888095C8972A5">Luong, Thien Tam</mods:namePart>
<mods:nameIdentifier id="83D5DF683087EAD6F54D5B71EAE65A45" type="ORCID">0000-0001-8354-6321</mods:nameIdentifier>
<mods:affiliation id="BBFC9DABED5CCB2DB1C21C0A45D26CFB">Department of Ecology - Evolutionary Biology, Vietnam National University Ho Chi Minh City (VNUHCM) - University of Science, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</mods:affiliation>
<mods:nameIdentifier id="DE860E13A99FD286403359ABC17CBA7D" type="email">lttam1511@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="CC47B69AA69CAB0A9F223205D026BCB5" type="personal">
<mods:role id="1D5A04899AF70296FC612B4FBFB94E74">
<mods:roleTerm id="2848D7992DB991BB5EA1D681D761B76D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1062E4B6885CA28CC9072582212F8F6A">Zhang, Li-Bing</mods:namePart>
<mods:nameIdentifier id="758DEA79D8F0AFE89539B71EF7EEB209" type="ORCID">0000-0002-4905-040X</mods:nameIdentifier>
<mods:affiliation id="A6191AAEED9D61352A7E17A985A19B90">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:nameIdentifier id="013F6D802B23F9A3AACF1A8DBB514EA6" type="email">Libing.Zhang@mobot.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8F71CC33001D4D70A905FE4AEF3051EA">text</mods:typeOfResource>
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<mods:titleInfo id="3F046E8C3A29AF6058E06D81E28C81F8">
<mods:title id="1F2FDA673164966AC6D1DE2D9CEC69A8">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="5463BFABA915E726ECDDC00532248407">
<mods:date id="DC6E3B9AC0AF7DE873D18DBF23AEBAD3">2024</mods:date>
<mods:detail id="C4949FB3371A95FBB44057C43D5FCF36" type="pubDate">
<mods:number id="A3489DD89F68AD5F710A835C9F06F4DC">2024-07-11</mods:number>
</mods:detail>
<mods:detail id="50AB87C469F696D0250E7D16ACDC6239" type="volume">
<mods:number id="D3255AEBEDE2A9C4AB624F1C820AD65C">659</mods:number>
</mods:detail>
<mods:detail id="1B8A9B274F558B50D3BDA97DF4A913C5" type="issue">
<mods:number id="5A982FC4970ABE0A46E2EF8EF6F1F719">3</mods:number>
</mods:detail>
<mods:extent id="6FC68FC68FFD0269F94A4AFF258E1FB3" unit="page">
<mods:start id="2FB58B850852943E4C579A379F401764">213</mods:start>
<mods:end id="06B0EAFAE2B77D37E8B64D13D65185DE">235</mods:end>
</mods:extent>
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<mods:location id="CA10E956FDE687934EE3E829E86F43AC">
<mods:url id="617D4FB35809D77114DBF41C2E020FA8">http://dx.doi.org/10.11646/phytotaxa.659.3.1</mods:url>
</mods:location>
<mods:classification id="4F529213FC1DD465CA6993AC1E154147">journal article</mods:classification>
<mods:identifier id="059CDAE737A4D84087B3D883584C1141" type="CLB-Dataset">302193</mods:identifier>
<mods:identifier id="638562CF4F310B2C3BE033FDA2A0F966" type="DOI">10.11646/phytotaxa.659.3.1</mods:identifier>
<mods:identifier id="AD25CB6468A8F092EBFE0E5AECC73B98" type="GBIF-Dataset">52524fe3-dfb8-4afc-829a-31089bb4e306</mods:identifier>
<mods:identifier id="CDF47526AA406B69348BA420F785D74B" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="BC72F1EA9D9B6BDD8A44C69A12501402" type="Zenodo-Dep">13217894</mods:identifier>
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<treatment id="4B175324C957A516D1BAFB6AFB3EAC93" ID-DOI="http://doi.org/10.5281/zenodo.13652219" ID-GBIF-Taxon="240245554" ID-Zenodo-Dep="13652219" LSID="urn:lsid:plazi:treatment:4B175324C957A516D1BAFB6AFB3EAC93" httpUri="http://treatment.plazi.org/id/4B175324C957A516D1BAFB6AFB3EAC93" lastPageId="5" lastPageNumber="218" pageId="4" pageNumber="217">
<subSubSection id="8BA4B1B9C957A517D1BAFB6AFACDAF8F" box="[136,1320,1191,1218]" pageId="4" pageNumber="217" type="nomenclature">
<paragraph id="C301E232C957A517D1BAFB6AFACDAF8F" blockId="4.[136,1320,1191,1218]" box="[136,1320,1191,1218]" pageId="4" pageNumber="217">
<heading id="9849555EC957A517D1BAFB6AFACDAF8F" box="[136,1320,1191,1218]" fontSize="11" level="2" pageId="4" pageNumber="217" reason="4">
1.
<taxonomicName id="04BE99B1C957A517D196FB65FB82AF8F" authority="Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou" authorityName="Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou" authorityYear="2024" box="[164,1127,1191,1218]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="217" phylum="Tracheophyta" rank="species" species="bolikhamsaiensis" status="sp. nov.">
<emphasis id="F1CA3E20C957A517D196FB65FE1CAF8F" bold="true" box="[164,505,1191,1218]" italics="true" pageId="4" pageNumber="217">Leptochilus bolikhamsaiensis</emphasis>
Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou
</taxonomicName>
,
<emphasis id="F1CA3E20C957A517D540FB6BFB22AF8F" box="[1138,1223,1193,1218]" italics="true" pageId="4" pageNumber="217">
<taxonomicNameLabel id="EAF9835BC957A517D540FB6BFB22AF8F" box="[1138,1223,1193,1218]" pageId="4" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5B85FEB7C957A517D5E4FB6AFAFFAF8F" box="[1238,1306,1192,1218]" captionStart="FIGURE 3" captionStartId="5.[136,229,1703,1725]" captionTargetBox="[208,1380,203,1667]" captionTargetId="figure-21@5.[195,1392,190,1679]" captionTargetPageId="5" captionText="FIGURE 3. Leptochilus bolikhamsaiensis Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou,—A. Habitat.—B.Adaxial lamina.—C. Abaxial lamina with five pairs of lobes.—D.Abaxial lamina with three pairs of lobes.—E. Rhizome with a fiddlehead. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217900" httpUri="https://zenodo.org/record/13217900/files/figure.png" pageId="4" pageNumber="217">Fig. 3</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C957A517D1BAFB32FB8CAE1E" pageId="4" pageNumber="217" type="materials_examined">
<paragraph id="C301E232C957A517D1BAFB32FB8CAE1E" blockId="4.[136,1452,1264,2046]" pageId="4" pageNumber="217">
<emphasis id="F1CA3E20C957A517D1BAFB32FF29AE47" bold="true" box="[136,204,1264,1290]" pageId="4" pageNumber="217">Type:</emphasis>
<collectingCountry id="BBA9A2A2C957A517D1E5FB32FEC1AE47" box="[215,292,1264,1290]" name="Laos" pageId="4" pageNumber="217">LAOS</collectingCountry>
.
<collectingRegion id="017A2CD0C957A517D003FB32FE26AE47" box="[305,451,1264,1290]" country="Laos" name="Bolikhamxai" pageId="4" pageNumber="217">Bolikhamsai</collectingRegion>
: Viengthong District, near Samsok Noi, elev.
<quantity id="04464FD7C957A517D2E9FB32FBCDAE47" box="[987,1064,1264,1290]" metricMagnitude="2" metricUnit="m" metricValue="9.77" pageId="4" pageNumber="217" unit="m" value="977.0">977 m</quantity>
,
<geoCoordinate id="A68A84F5C957A517D505FB32FB71AE46" box="[1079,1172,1264,1291]" degrees="18.57" direction="north" orientation="latitude" pageId="4" pageNumber="217" precision="555" value="18.57">18.57°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C957A517D5AFFB32FAE3AE47" box="[1181,1286,1264,1291]" degrees="104.62" direction="east" orientation="longitude" pageId="4" pageNumber="217" precision="555" value="104.62">104.62°E</geoCoordinate>
, on limestone rocks in disturbed rainforest,
<date id="B700C4F2C957A517D0D0FAD6FD6FAE63" box="[482,650,1300,1326]" pageId="4" pageNumber="217" value="2019-04-30">30 April 2019</date>
,
<emphasis id="F1CA3E20C957A517D3ABFAD6FEC0AE1F" italics="true" pageId="4" pageNumber="217">Zhuo Zhou, Khamphanh Thepkaysone, Mouachanh Xayvue, Kosorn Sorsoulithanh</emphasis>
,
<emphasis id="F1CA3E20C957A517D001FAFBFE21AE1F" box="[307,452,1336,1362]" italics="true" pageId="4" pageNumber="217">Liang Zhang</emphasis>
LZ156 (
<typeStatus id="1C055C90C957A517D317FAFAFD6CAE1F" box="[549,649,1336,1362]" pageId="4" pageNumber="217" type="holotype">holotype</typeStatus>
KUN-1576320!,
<typeStatus id="1C055C90C957A517D27DFAFAFC45AE1F" box="[847,928,1336,1362]" pageId="4" pageNumber="217" type="isotype">isotype</typeStatus>
KUN-1576319!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C957A517D18FFA99FE24AC6B" pageId="4" pageNumber="217" type="diagnosis">
<paragraph id="C301E232C957A517D18FFA99FE8BAED7" blockId="4.[136,1452,1264,2046]" pageId="4" pageNumber="217">
<emphasis id="F1CA3E20C957A517D18FFA99FED5AE38" bold="true" box="[189,304,1371,1397]" pageId="4" pageNumber="217">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C957A517D00DFA9EFD6CAE3B" box="[319,649,1372,1398]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="217" phylum="Tracheophyta" rank="species" species="bolikhamsaiensis">
<emphasis id="F1CA3E20C957A517D00DFA9EFD6CAE3B" box="[319,649,1372,1398]" italics="true" pageId="4" pageNumber="217">Leptochilus bolikhamsaiensis</emphasis>
</taxonomicName>
is distinguished from other species by its small habit with laminae
<quantity id="04464FD7C957A517D441FA9EFF48AED7" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="4" pageNumber="217" unit="cm" value="10.5" valueMax="15.0" valueMin="6.0">615 cm</quantity>
, and oblong sori.
</paragraph>
<paragraph id="C301E232C957A517D18FFA66FE24AC6B" blockId="4.[136,1452,1264,2046]" pageId="4" pageNumber="217">
<emphasis id="F1CA3E20C957A517D18FFA66FEEDAEF3" bold="true" box="[189,264,1444,1470]" pageId="4" pageNumber="217">Plants</emphasis>
perennial, evergreen.
<emphasis id="F1CA3E20C957A517D0CCFA61FD8DAEF0" bold="true" box="[510,616,1443,1469]" pageId="4" pageNumber="217">Rhizome</emphasis>
creeping, ca.
<quantity id="04464FD7C957A517D232FA66FC63AEF3" box="[768,902,1444,1470]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="4" pageNumber="217" unit="mm" value="2.5" valueMax="2.9" valueMin="2.1">2.12.9 mm</quantity>
in diam., dorsiventrally flattened, densely scaled, with many short roots; scales brown, clathrate, membranous, margins entire or sparely denticulate, slightly spreading, 0.61.6 ×
<quantity id="04464FD7C957A517D033FA2EFE6AAD4B" box="[257,399,1516,1542]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="4" pageNumber="217" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.40.7 mm</quantity>
.
<emphasis id="F1CA3E20C957A517D0AFFA2EFE17AD4B" bold="true" box="[413,498,1516,1542]" pageId="4" pageNumber="217">Fronds</emphasis>
monomorphic or slightly dimorphic, (0.3)
<quantity id="04464FD7C957A517D2DFFA2EFB68AD4B" box="[1005,1165,1516,1542]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.8" metricValueMin="0.9" pageId="4" pageNumber="217" unit="cm" value="1.35" valueMax="1.8" valueMin="0.9">0.9 to 1.8 cm</quantity>
distant,
<quantity id="04464FD7C957A517D5C6FA2EFA82AD4B" box="[1268,1383,1516,1542]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="4" pageNumber="217" unit="cm" value="19.5" valueMax="23.0" valueMin="16.0">1623 cm</quantity>
long; petioles stramineous,
<quantity id="04464FD7C957A517D049F9D2FE3EAD67" box="[379,475,1552,1578]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.4" metricValueMin="0.6" pageId="4" pageNumber="217" unit="cm" value="10.0" valueMax="14.0" valueMin="6.0">614 cm</quantity>
long, ca.
<quantity id="04464FD7C957A517D37AF9D2FD2AAD67" box="[584,719,1552,1579]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="4" pageNumber="217" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
in diam. at middle, slightly winged at distal part; rachis distinctly winged,
<quantity id="04464FD7C957A517D1DBF9F6FE94AD03" box="[233,369,1588,1614]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="4" pageNumber="217" unit="mm" value="2.35" valueMax="2.6" valueMin="2.1">2.12.6 mm</quantity>
wide at the distal part,
<quantity id="04464FD7C957A517D34BF9F6FCE4AD03" box="[633,769,1588,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="4" pageNumber="217" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.71.9 mm</quantity>
at base.
<emphasis id="F1CA3E20C957A517D256F9F1FC2BAD00" bold="true" box="[868,974,1587,1613]" pageId="4" pageNumber="217">Laminae</emphasis>
oblong-ovate, deeply 1-pinnatifid, 615 ×
<quantity id="04464FD7C957A517D1BAF99AFF0EAD3F" box="[136,235,1624,1650]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="4" pageNumber="217" unit="cm" value="8.5" valueMax="10.0" valueMin="7.0">710 cm</quantity>
, widest at middle or below middle of lamina, chartaceous, brown when dry, glabrous, margins of lobes entire, slightly repand.
<emphasis id="F1CA3E20C957A517D073F9BEFE6CADDB" bold="true" box="[321,393,1660,1686]" pageId="4" pageNumber="217">Lobes</emphasis>
(1)56 pairs, narrowly lanceolate, apex acuminate, 4.35.2 ×
<quantity id="04464FD7C957A517D56CF9BEFB05ADDB" box="[1118,1248,1660,1686]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="4" pageNumber="217" unit="cm" value="0.8" valueMax="0.9" valueMin="0.7">0.70.9 cm</quantity>
, basal portions of lobes narrowed, ascending upward, midribs of lobes at angles of 4772(90) degree with rachis, distal pair of lobes slightly narrowed, opposite or slightly alternate, lobe base ca.
<quantity id="04464FD7C957A517D264F906FC33AD93" box="[854,982,1732,1758]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="4" pageNumber="217" unit="cm" value="0.65" valueMax="0.9" valueMin="0.4">0.40.9 cm</quantity>
distant with each other. Midribs slightly raised, lateral veins invisible.
<emphasis id="F1CA3E20C957A517D0EEF925FDEBAC4C" bold="true" box="[476,526,1767,1793]" pageId="4" pageNumber="217">Sori</emphasis>
oblong, (4)911 on each side of midrib, 13 × 0.71.0 mm, centers ca. 35(9) mm apart from one another.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C957A517D18FF8EDFCBFAC22" pageId="4" pageNumber="217" type="description">
<paragraph id="C301E232C957A517D18FF8EDFCBFAC22" blockId="4.[136,1452,1264,2046]" pageId="4" pageNumber="217">
<emphasis id="F1CA3E20C957A517D18FF8EDFDD2AC04" bold="true" box="[189,567,1839,1865]" pageId="4" pageNumber="217">Additional specimen examined</emphasis>
:
<collectingCountry id="BBA9A2A2C957A517D378F8F2FD72AC07" box="[586,663,1840,1866]" name="Laos" pageId="4" pageNumber="217">LAOS</collectingCountry>
. Na be, Provincia,
<date id="B700C4F2C957A517D24DF8F2FBDCAC07" box="[895,1081,1840,1866]" pageId="4" pageNumber="217" value="1930-12">December 1930</date>
, collectors unknown, a number labeled as “4.107” may be the collecting number (
<collectingCountry id="BBA9A2A2C957A517D388F896FD05AC22" box="[698,736,1876,1903]" name="United States of America" pageId="4" pageNumber="217">US</collectingCountry>
1595356!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C957A517D18FF8B5FCC9ACB3" pageId="4" pageNumber="217" type="distribution">
<paragraph id="C301E232C957A517D18FF8B5FCC9ACB3" blockId="4.[136,1452,1264,2046]" pageId="4" pageNumber="217">
<emphasis id="F1CA3E20C957A517D18FF8B5FE0AACDC" bold="true" box="[189,495,1911,1938]" pageId="4" pageNumber="217">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C957A517D0CBF8BAFCA2ACDF" box="[505,839,1912,1938]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="217" phylum="Tracheophyta" rank="species" species="bolikhamsaiensis">
<emphasis id="F1CA3E20C957A517D0CBF8BAFCA2ACDF" box="[505,839,1912,1938]" italics="true" pageId="4" pageNumber="217">Leptochilus bolikhamsaiensis</emphasis>
</taxonomicName>
was initially collected in
<collectingCountry id="BBA9A2A2C957A517D547F8BAFB4EACDF" box="[1141,1195,1912,1938]" name="Laos" pageId="4" pageNumber="217">Laos</collectingCountry>
in 1930, but its exact location was not specified. Currently, only one population in Viengthong District of
<collectingRegion id="017A2CD0C957A517D51FF85EFAC0ACFB" box="[1069,1317,1948,1974]" country="Laos" name="Bolikhamxai" pageId="4" pageNumber="217">Bolikhamsai Province</collectingRegion>
is known to us. The individuals of the species were observed growing exclusively on limestone rocks in the rainforest. It is likely that the species occurs in other karst regions in central
<collectingCountry id="BBA9A2A2C957A517D3DCF826FCCDACB3" box="[750,808,2020,2046]" name="Laos" pageId="4" pageNumber="217">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="97C1B2BAC956A516D1BAF965FF29AC4B" ID-DOI="http://doi.org/10.5281/zenodo.13217900" ID-Zenodo-Dep="13217900" httpUri="https://zenodo.org/record/13217900/files/figure.png" pageId="5" pageNumber="218" startId="5.[136,229,1703,1725]" targetBox="[208,1380,203,1667]" targetPageId="5" targetType="figure">
<paragraph id="C301E232C956A516D1BAF965FF29AC4B" blockId="5.[136,1452,1703,1798]" pageId="5" pageNumber="218">
<emphasis id="F1CA3E20C956A516D1BAF965FF18ADF0" bold="true" box="[136,253,1703,1725]" pageId="5" pageNumber="218">FIGURE 3.</emphasis>
<taxonomicName id="04BE99B1C956A516D030F96AFBC9ADF3" authority="Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou" authorityName="Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou" box="[258,1068,1704,1726]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="218" phylum="Tracheophyta" rank="species" species="bolikhamsaiensis">
<emphasis id="F1CA3E20C956A516D030F96AFDF8ADF0" box="[258,541,1704,1725]" italics="true" pageId="5" pageNumber="218">Leptochilus bolikhamsaiensis</emphasis>
Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou
</taxonomicName>
,—A. Habitat.—B. Adaxial lamina.—C. Abaxial lamina with five pairs of lobes.—D. Abaxial lamina with three pairs of lobes.—E. Rhizome with a fiddlehead. Photo credit: Liang Zhang.
</paragraph>
</caption>
<subSubSection id="8BA4B1B9C956A516D18FF8F6FB3EAC93" pageId="5" pageNumber="218" type="etymology">
<paragraph id="C301E232C956A516D18FF8F6FF34AC3F" blockId="5.[136,1452,1844,2015]" pageId="5" pageNumber="218">
<emphasis id="F1CA3E20C956A516D18FF8F6FEDBAC03" bold="true" box="[189,318,1844,1870]" pageId="5" pageNumber="218">Etymology</emphasis>
: The species epithet is derived from the name of
<collectingRegion id="017A2CD0C956A516D24AF8F6FB91AC03" box="[888,1140,1844,1870]" country="Laos" name="Bolikhamxai" pageId="5" pageNumber="218">Bolikhamsai Province</collectingRegion>
,
<collectingCountry id="BBA9A2A2C956A516D5B3F8F7FB5FAC02" box="[1153,1210,1845,1871]" name="Laos" pageId="5" pageNumber="218">Laos</collectingCountry>
, where the species is found.
</paragraph>
<paragraph id="C301E232C956A516D18FF8BEFB3EAC93" blockId="5.[136,1452,1844,2015]" pageId="5" pageNumber="218">
<emphasis id="F1CA3E20C956A516D18FF8BEFF10ACDB" bold="true" box="[189,245,1916,1942]" pageId="5" pageNumber="218">Note</emphasis>
:
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<emphasis id="F1CA3E20C956A516D038F8BEFDBFACDB" box="[266,602,1916,1942]" italics="true" pageId="5" pageNumber="218">Leptochilus bolikhamsaiensis</emphasis>
</taxonomicName>
is morphologically the smallest species, regarding to the
<emphasis id="F1CA3E20C956A516D420F8BEFA49ACDB" box="[1298,1452,1916,1942]" italics="true" pageId="5" pageNumber="218">
<taxonomicName id="04BE99B1C956A516D42CF8BEFA49ACDB" box="[1310,1452,1916,1942]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="218" phylum="Tracheophyta" rank="species" species="ellipticus">L. ellipticus</taxonomicName>
</emphasis>
complex”. However, according to the phylogenetic analysis, this species forms an isolated clade within the genus, and its relationships to other species of the genus, particularly to the
<taxonomicName id="04BE99B1C956A516D268F807FC3BAC93" box="[858,990,1988,2014]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="218" phylum="Tracheophyta" rank="species" species="ellipticus">
<emphasis id="F1CA3E20C956A516D268F807FC3BAC93" box="[858,990,1988,2014]" italics="true" pageId="5" pageNumber="218">L. ellipticus</emphasis>
</taxonomicName>
clade, remain unclear.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,307 @@
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6.
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<emphasis id="F1CA3E20C958A518D196FCACFDEAA8C4" bold="true" box="[164,527,878,905]" italics="true" pageId="11" pageNumber="224">Leptochilus luangprabangensis</emphasis>
Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou
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(
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).
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<emphasis id="F1CA3E20C958A518D1BAFC74FF26A89D" bold="true" box="[136,195,950,976]" pageId="11" pageNumber="224">Type</emphasis>
:
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, on a small tree trunk in limestone mountain,
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&amp;
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415
</emphasis>
</materialsCitation>
<materialsCitation id="73D6E86FC958A518D2AEFC18FB22A8B8" ID-GBIF-Occurrence="4928466301" box="[924,1223,986,1013]" collectingDate="2019-05-13" collectionCode="KUN" collectorName="Zhuo Zhou &amp; Liang Zhang LZ" country="Laos" elevation="586" latitude="19.83" location="City" longLatPrecision="762" longitude="102.15" pageId="11" pageNumber="224" specimenCount="2" stateProvince="Louangphabang" typeStatus="holotype">
(
<typeStatus id="1C055C90C958A518D296FC18FBEDA8B9" box="[932,1032,986,1012]" pageId="11" pageNumber="224" type="holotype">holotype</typeStatus>
KUN-1576324!;
</materialsCitation>
<materialsCitation id="73D6E86FC958A518D5FCFC18FA66A8B9" ID-GBIF-Occurrence="4928466316" box="[1230,1411,986,1012]" collectingDate="2019-05-13" collectionCode="KUN" collectorName="Zhuo Zhou &amp; Liang Zhang LZ" country="Laos" elevation="586" latitude="19.83" location="City" longLatPrecision="762" longitude="102.15" pageId="11" pageNumber="224" specimenCount="2" stateProvince="Louangphabang" typeStatus="holotype">
<typeStatus id="1C055C90C958A518D5FCFC18FACFA8B9" box="[1230,1322,986,1012]" pageId="11" pageNumber="224" type="isotype">isotypes</typeStatus>
<collectionCode id="A5AF7AF7C958A518D403FC18FA9DA8B9" box="[1329,1400,986,1012]" country="China" lsid="urn:lsid:biocol.org:col:15842" name="Kunming Institute of Botany, Chinese Academy of Sciences" pageId="11" pageNumber="224" type="Herbarium">KUN</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFC3CFD1EAEA1" pageId="11" pageNumber="224" type="diagnosis">
<paragraph id="C301E232C958A518D18FFC3CFBEDAF71" blockId="11.[136,1452,950,1985]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFC3CFED5AF55" bold="true" box="[189,304,1022,1048]" pageId="11" pageNumber="224">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C958A518D072FC3CFD44AF55" box="[320,673,1022,1048]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C958A518D072FC3CFD44AF55" box="[320,673,1022,1048]" italics="true" pageId="11" pageNumber="224">Leptochilus luangprabangensis</emphasis>
</taxonomicName>
is characterized by its climbing habit, the length of fertile and sterile fronds nearly identical, thin lamina texture, and sterile fronds shortly petiolated.
</paragraph>
<paragraph id="C301E232C958A518D18FFB84FD1EAEA1" blockId="11.[136,1452,950,1985]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFB84FEEDAF2D" bold="true" box="[189,264,1094,1120]" pageId="11" pageNumber="224">Plants</emphasis>
perennial, evergreen, epiphytic.
<emphasis id="F1CA3E20C958A518D341FB84FD38AF2D" bold="true" box="[627,733,1094,1120]" pageId="11" pageNumber="224">Rhizome</emphasis>
long creeping, ca. 2.53.0 mm in diam., dorsiventrally flattened, scaled, with a few short phyllopodia and roots. Apex of rhizome and basal petiole densely scaly, scales subulatelanceolate, 1.42.4 ×
<quantity id="04464FD7C958A518D04BFB4CFDE7AFE4" box="[377,514,1166,1193]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="11" pageNumber="224" unit="mm" value="0.5" valueMax="0.7" valueMin="0.3">0.30.7 mm</quantity>
, brown, basifixed, concolorous, clathrate, margin entire or sparely denticulate, apex acuminate.
<emphasis id="F1CA3E20C958A518D039FB70FE85AF81" bold="true" box="[267,352,1202,1228]" pageId="11" pageNumber="224">Fronds</emphasis>
dimorphic, sterile fronds larger than fertile fronds.
<emphasis id="F1CA3E20C958A518D283FB70FBB1AF81" bold="true" box="[945,1108,1202,1228]" pageId="11" pageNumber="224">Sterile fronds</emphasis>
<quantity id="04464FD7C958A518D56CFB70FB2AAF80" box="[1118,1231,1202,1229]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="11" pageNumber="224" unit="cm" value="15.0" valueMax="18.0" valueMin="12.0">1218 cm</quantity>
tall, laminae much longer than petioles;
<emphasis id="F1CA3E20C958A518D04BFB14FE3DAFBD" bold="true" box="[377,472,1238,1264]" pageId="11" pageNumber="224">laminae</emphasis>
ribbon-lanceolate, base cuneate, apex acuminate,
<quantity id="04464FD7C958A518D527FB14FB63AFBC" box="[1045,1158,1238,1265]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="11" pageNumber="224" unit="cm" value="13.0" valueMax="16.0" valueMin="10.0">1016 cm</quantity>
long, 2.0
<quantity id="04464FD7C958A518D431FB14FAAAAFBC" box="[1283,1359,1238,1265]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="224" unit="cm" value="3.1">3.1 cm</quantity>
wide at middle, chartaceous, glabrous;
<emphasis id="F1CA3E20C958A518D0D9FB38FDA0AE59" bold="true" box="[491,581,1274,1300]" pageId="11" pageNumber="224">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
<quantity id="04464FD7C958A518D51CFB38FB52AE58" box="[1070,1207,1274,1301]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="11" pageNumber="224" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 mm</quantity>
in diam.,
<quantity id="04464FD7C958A518D41EFB38FA4EAE58" box="[1324,1451,1274,1301]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.7" metricValueMin="0.6" pageId="11" pageNumber="224" unit="cm" value="1.15" valueMax="1.7" valueMin="0.6">0.61.7 cm</quantity>
long.
<emphasis id="F1CA3E20C958A518D1F8FADCFE95AE75" bold="true" box="[202,368,1310,1336]" pageId="11" pageNumber="224">Fertile fronds</emphasis>
<quantity id="04464FD7C958A518D048FADCFE0EAE74" box="[378,491,1310,1337]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="11" pageNumber="224" unit="cm" value="15.5" valueMax="17.0" valueMin="14.0">1417 cm</quantity>
tall, petioles shorter than laminae;
<emphasis id="F1CA3E20C958A518D2B7FADCFC01AE75" bold="true" box="[901,996,1310,1336]" pageId="11" pageNumber="224">laminae</emphasis>
elliptic-lanceolate, margins entire, base cuneate, apex long acuminate,
<quantity id="04464FD7C958A518D0C2FA80FD87AE10" box="[496,610,1346,1373]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="11" pageNumber="224" unit="cm" value="12.0" valueMax="13.0" valueMin="11.0">1113 cm</quantity>
long,
<quantity id="04464FD7C958A518D383FA80FCD4AE10" box="[689,817,1346,1373]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="11" pageNumber="224" unit="cm" value="1.4" valueMax="1.5" valueMin="1.3">1.31.5 cm</quantity>
wide at middle, thinly chartaceous, glabrous;
<emphasis id="F1CA3E20C958A518D463FA80FA4EAE11" bold="true" box="[1361,1451,1346,1372]" pageId="11" pageNumber="224">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
<quantity id="04464FD7C958A518D369FAA4FD04AECC" box="[603,737,1382,1409]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="11" pageNumber="224" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.50.7 mm</quantity>
in diam.,
<quantity id="04464FD7C958A518D27EFAA4FC7BAECC" box="[844,926,1382,1409]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="11" pageNumber="224" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">34 cm</quantity>
long.
<emphasis id="F1CA3E20C958A518D2D0FAA4FBC7AECD" bold="true" box="[994,1058,1382,1408]" pageId="11" pageNumber="224">Veins</emphasis>
anastomosing, lateral veins visible, veinlets invisible.
<emphasis id="F1CA3E20C958A518D060FA48FE61AEE9" bold="true" box="[338,388,1418,1444]" pageId="11" pageNumber="224">Sori</emphasis>
linear, reach lamina margins, ca.
<quantity id="04464FD7C958A518D3C5FA48FCA9AEE8" box="[759,844,1418,1445]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="224" unit="mm" value="0.5">0.5 mm</quantity>
from margins and
<quantity id="04464FD7C958A518D52EFA48FB94AEE8" box="[1052,1137,1418,1445]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="224" unit="mm" value="0.4">0.4 mm</quantity>
from midrib, 2528 on each side of midrib,
<quantity id="04464FD7C958A518D004FA6CFE51AE84" box="[310,436,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="11" pageNumber="224" unit="cm" value="0.8" valueMax="0.9" valueMin="0.7">0.70.9 cm</quantity>
long,
<quantity id="04464FD7C958A518D0CEFA6CFD61AE84" box="[508,644,1454,1481]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="11" pageNumber="224" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.40.6 mm</quantity>
wide when mature, at angles of 3645 degree with midribs, centers ca. (1)
<quantity id="04464FD7C958A518D181FA10FEF1AEA0" box="[179,276,1490,1517]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="11" pageNumber="224" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
apart from one another. Paraphyses absent.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFA34FA98AD15" pageId="11" pageNumber="224" type="distribution">
<paragraph id="C301E232C958A518D18FFA34FA98AD15" blockId="11.[136,1452,950,1985]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFA34FE00AD5D" bold="true" box="[189,485,1526,1552]" pageId="11" pageNumber="224">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C958A518D0D8FA34FCADAD5D" box="[490,840,1526,1552]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C958A518D0D8FA34FCADAD5D" box="[490,840,1526,1552]" italics="true" pageId="11" pageNumber="224">Leptochilus luangprabangensis</emphasis>
</taxonomicName>
is known from a single collection made in the limestone forest near the capital city of
<collectingRegion id="017A2CD0C958A518D0FCF9D9FD00AD79" box="[462,741,1562,1589]" country="Laos" name="Louangphabang" pageId="11" pageNumber="224">Luang Prabang Province</collectingRegion>
,
<collectingCountry id="BBA9A2A2C958A518D3C2F9D9FCCFAD78" box="[752,810,1563,1589]" name="Laos" pageId="11" pageNumber="224">Laos</collectingCountry>
. The species was observed growing as a hemiepiphyte on a small tree trunk. It was anchored to the tree with rhizomes and roots, and the roots extended into the soil below.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FF9A0FE16AC8C" pageId="11" pageNumber="224" type="etymology">
<paragraph id="C301E232C958A518D18FF9A0FB1FAD31" blockId="11.[136,1452,950,1985]" box="[189,1274,1634,1661]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FF9A0FEDBAD31" bold="true" box="[189,318,1634,1660]" pageId="11" pageNumber="224">Etymology</emphasis>
: The species is named after
<collectingRegion id="017A2CD0C958A518D34FF9A1FC71AD31" box="[637,916,1634,1661]" country="Laos" name="Louangphabang" pageId="11" pageNumber="224">Luang Prabang Province</collectingRegion>
,
<collectingCountry id="BBA9A2A2C958A518D292F9A1FC3CAD30" box="[928,985,1635,1661]" name="Laos" pageId="11" pageNumber="224">Laos</collectingCountry>
, where it was discovered.
</paragraph>
<paragraph id="C301E232C958A518D18FF944FE16AC8C" blockId="11.[136,1452,950,1985]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FF944FF10ADED" bold="true" box="[189,245,1670,1696]" pageId="11" pageNumber="224">Note</emphasis>
: Morphologically,
<taxonomicName id="04BE99B1C958A518D0FBF944FCCDADED" box="[457,808,1670,1696]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C958A518D0FBF944FCCDADED" box="[457,808,1670,1696]" italics="true" pageId="11" pageNumber="224">Leptochilus luangprabangensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="04BE99B1C958A518D2DCF945FEE8AD89" authority="Liang Zhang &amp; Li Bing Zhang (2018: 173)" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityPageNumber="173" authorityYear="2018" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C958A518D2DCF945FBB2ADED" box="[1006,1111,1670,1696]" italics="true" pageId="11" pageNumber="224">L. chingii</emphasis>
Liang Zhang &amp; Li Bing
<bibRefCitation id="A72F9FC3C958A518D456F944FEE8AD89" author="Zhang, L. &amp; Guo, L. &amp; Zhang, L. - B." pageId="11" pageNumber="224" pagination="172 - 176" refId="ref12331" refString="Zhang, L., Guo, L. &amp; Zhang, L. - B. (2018) New combinations in the fern genus Leptochilus (Polypodiaceae). Phytotaxa 374: 172 - 176. https: // doi. org / 10.11646 / phytotaxa. 374.2.10" type="journal article" year="2018">Zhang (2018: 173)</bibRefCitation>
</taxonomicName>
due to their simple and slightly dimorphic fronds, lanceolate laminae, and laminae gradually narrowed towards base. However, they differ in the lamina texture (thinly chartaceous vs. chartaceous or thickly chartaceous in
<taxonomicName id="04BE99B1C958A518D1BAF931FF11AC41" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" box="[136,244,1778,1804]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C958A518D1BAF931FF11AC41" box="[136,244,1778,1804]" italics="true" pageId="11" pageNumber="224">L. chingii</emphasis>
</taxonomicName>
and petiole length of sterile fronds (
<quantity id="04464FD7C958A518D3A3F930FCF5AC40" box="[657,784,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.7" metricValueMin="0.6" pageId="11" pageNumber="224" unit="cm" value="1.15" valueMax="1.7" valueMin="0.6">0.61.7 cm</quantity>
vs. (0.5)
<quantity id="04464FD7C958A518D2B6F930FC38AC41" box="[900,989,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="11" pageNumber="224" unit="cm" value="3.5" valueMax="5.0" valueMin="2.0">25 cm</quantity>
). Additionally,
<taxonomicName id="04BE99B1C958A518D5BBF931FA9BAC41" box="[1161,1406,1778,1804]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C958A518D5BBF931FA9BAC41" box="[1161,1406,1778,1804]" italics="true" pageId="11" pageNumber="224">L. luangprabangensis</emphasis>
</taxonomicName>
has only been observed growing as a hemiepiphyte on tree trunks, while
<taxonomicName id="04BE99B1C958A518D297F8D5FBF7AC7D" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" box="[933,1042,1814,1840]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C958A518D297F8D5FBF7AC7D" box="[933,1042,1814,1840]" italics="true" pageId="11" pageNumber="224">L. chingii</emphasis>
</taxonomicName>
was found on moist rocks. Though
<taxonomicName id="04BE99B1C958A518D1BAF8F9FE9BAC19" box="[136,382,1850,1876]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C958A518D1BAF8F9FE9BAC19" box="[136,382,1850,1876]" italics="true" pageId="11" pageNumber="224">L. luangprabangensis</emphasis>
</taxonomicName>
is phylogenetically the closest to
<taxonomicName id="04BE99B1C958A518D231F8F9FC56AC19" box="[771,947,1851,1876]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C958A518D231F8F9FC56AC19" box="[771,947,1851,1876]" italics="true" pageId="11" pageNumber="224">L. vietnamensis</emphasis>
</taxonomicName>
, they have 12 bp differences in 3074 bps of aligned three plastid markers (
<bibRefCitation id="A72F9FC3C958A518D0EBF89CFD40AC35" author="Zhang, L. &amp; Liang, Z. - L. &amp; Fan, X. - P. &amp; Lu, N. T. &amp; Zhou, X. - M. &amp; Wei, H. - J. &amp; Zhang, L. - B. The &amp; Biodiversity Hotspot" box="[473,677,1886,1912]" pageId="11" pageNumber="224" refId="ref12381" refString="Zhang, L., Liang, Z. - L., Fan, X. - P., Lu, N. T., Zhou, X. - M., Wei, H. - J. &amp; Zhang, L. - B. The Indo-Burma Biodiversity Hotspot for ferns: updated phylogeny helps discover hidden diversity in the Java fern genus Leptochilus (Polypodiaceae). (2024) Plant Diversity. [in press]" type="book" year="2024">
Zhang
<emphasis id="F1CA3E20C958A518D31BF89DFD84AC35" box="[553,609,1886,1912]" italics="true" pageId="11" pageNumber="224">et al.</emphasis>
2024
</bibRefCitation>
). Morphologically,
<taxonomicName id="04BE99B1C958A518D24CF89DFB96AC35" box="[894,1139,1886,1912]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C958A518D24CF89DFB96AC35" box="[894,1139,1886,1912]" italics="true" pageId="11" pageNumber="224">L. luangprabangensis</emphasis>
</taxonomicName>
has fertile fronds nearly the same length as sterile fronds that are
<quantity id="04464FD7C958A518D300F840FD47ACD0" box="[562,674,1922,1949]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="11" pageNumber="224" unit="cm" value="15.0" valueMax="18.0" valueMin="12.0">1218 cm</quantity>
long, while
<taxonomicName id="04BE99B1C958A518D201F841FC06ACD1" box="[819,995,1923,1948]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C958A518D201F841FC06ACD1" box="[819,995,1923,1948]" italics="true" pageId="11" pageNumber="224">L. vietnamensis</emphasis>
</taxonomicName>
has fertile fronds much shorter than the sterile fronds that are
<quantity id="04464FD7C958A518D04EF864FE0AAC8C" box="[380,495,1958,1985]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="11" pageNumber="224" unit="cm" value="35.0" valueMax="40.0" valueMin="30.0">3040 cm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="001326A6F7A61FE7921A7A995760DE5D" ID-CLB-Dataset="302193" ID-DOI="10.11646/phytotaxa.659.3.1" ID-GBIF-Dataset="52524fe3-dfb8-4afc-829a-31089bb4e306" ID-ISSN="1179-3163" ID-Zenodo-Dep="13217894" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1722613780751" checkinUser="felipe" docAuthor="Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, Zhou, Zhuo, Thepkayson, Khamphanh, Luong, Thien Tam &amp; Zhang, Li-Bing" docDate="2024" docId="4B175324C95AA518D1BAF8E5FEE3A80D" docLanguage="en" docName="phytotaxa.659.3.1.pdf" docOrigin="Phytotaxa 659 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.659.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Leptochilus locii Liang Zhang, N. T. Lu &amp; Li Bing Zhang, sp. nov." docType="treatment" docVersion="5" lastPageNumber="224" masterDocId="B72E2B5CC953A513D132FFC2FFE5AB4D" masterDocTitle="Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot" masterLastPageNumber="235" masterPageNumber="213" pageNumber="222" updateTime="1725399130512" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8F6E48E5E9324DF8900AB7373FE6FDE8">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:namePart id="29BA4DFF4B0E6DE2B18F0747963A0F10">Zhang, Liang</mods:namePart>
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<mods:affiliation id="615129A5219FE9FA8DE5C15500E85CB3">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
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<mods:namePart id="F3CB5055ECAB49D6DCABF0FA6682E604">Lu, Ngan Thi</mods:namePart>
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<mods:affiliation id="C936997C4B99B81D3B0F65BC22536D7D">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="B4BC4D591A2454E023B8D9F2505DC2DB">Zhou, Xin-Mao</mods:namePart>
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<mods:namePart id="010DFE724E16E89F2444FF086D22FECA">Zhou, Zhuo</mods:namePart>
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<mods:namePart id="50390A7E3222F206E1ECD8E0B5E6F2BD">Thepkayson, Khamphanh</mods:namePart>
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<mods:affiliation id="7746039B96591B2563CFA1ED43F6CE36">Institute of Traditional Medicine, Vientiane, Lao PDR</mods:affiliation>
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<mods:namePart id="CD1342F7FFB65F71568D6A10E9F93D8A">Luong, Thien Tam</mods:namePart>
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<mods:affiliation id="E405480BC771D3AA04944C65B104FE8D">Department of Ecology - Evolutionary Biology, Vietnam National University Ho Chi Minh City (VNUHCM) - University of Science, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</mods:affiliation>
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<mods:namePart id="EAC5E7CE15398365FFF125221B609F9B">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="497E985CD84FC7BE18CC756B542CD3FF">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:nameIdentifier id="E62D39B9D1184ACE9B0DADC039630CEB" type="email">Libing.Zhang@mobot.org</mods:nameIdentifier>
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<mods:date id="0F52E7B66660D776367329332CD50816">2024</mods:date>
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<treatment id="4B175324C95AA518D1BAF8E5FEE3A80D" ID-DOI="http://doi.org/10.5281/zenodo.13652242" ID-GBIF-Taxon="240245552" ID-Zenodo-Dep="13652242" LSID="urn:lsid:plazi:treatment:4B175324C95AA518D1BAF8E5FEE3A80D" httpUri="http://treatment.plazi.org/id/4B175324C95AA518D1BAF8E5FEE3A80D" lastPageId="11" lastPageNumber="224" pageId="9" pageNumber="222">
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5.
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<emphasis id="F1CA3E20C95AA51AD196F8E4FE81AC0C" bold="true" box="[164,356,1830,1857]" italics="true" pageId="9" pageNumber="222">Leptochilus locii</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
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,
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<taxonomicNameLabel id="EAF9835BC95AA51AD205F8EAFC69AC0C" box="[823,908,1832,1857]" pageId="9" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5B85FEB7C95AA51AD2A9F8E5FC3AAC0C" box="[923,991,1831,1857]" captionStart="FIGURE 7" captionStartId="10.[136,229,1804,1826]" captionTargetBox="[163,1424,202,1767]" captionTargetId="figure-25@10.[151,1436,190,1780]" captionTargetPageId="10" captionText="FIGURE 7. Leptochilus locii Liang Zhang, N.T. Lu &amp; Li Bing Zhang,—A. Habitat.—B. Adaxial view of sterile frond.—C. Habit.—D. Abaxial view of sterile frond.—E. Abaxial view of fertile lamina.—F. Adaxial view of fertile lamina. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217908" httpUri="https://zenodo.org/record/13217908/files/figure.png" pageId="9" pageNumber="222">Fig. 7</figureCitation>
).
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</paragraph>
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<emphasis id="F1CA3E20C95AA51AD1BAF8ADFF26ACC4" bold="true" box="[136,195,1903,1929]" pageId="9" pageNumber="222">Type</emphasis>
:
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.
<collectingRegion id="017A2CD0C95AA51AD054F8ADFE10ACC4" box="[358,501,1903,1929]" country="Vietnam" name="Quang Binh" pageId="9" pageNumber="222">Quang Binh</collectingRegion>
: Phong Nha Ke Bang National Park, Tan Traih, elev.
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,
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/106.20°, on rocks in limestone mountains,
<date id="B700C4F2C95AA51AD0E9F851FD43ACE0" box="[475,678,1939,1965]" pageId="9" pageNumber="222" value="2014-11-06">6 November 2014</date>
,
<emphasis id="F1CA3E20C95AA51AD382F856FB3DACE0" box="[688,1240,1939,1965]" italics="true" pageId="9" pageNumber="222">Li Bing Zhang, Liang Zhang &amp; Ngan Thi Lu 7352</emphasis>
(
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VNMN,
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!).
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<emphasis id="F1CA3E20C95AA51AD18FF819FED5ACB8" bold="true" box="[189,304,2011,2037]" pageId="9" pageNumber="222">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C95AA51AD071F819FDE7ACB8" box="[323,514,2011,2037]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="9" pageNumber="222" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C95AA51AD071F819FDE7ACB8" box="[323,514,2011,2037]" italics="true" pageId="9" pageNumber="222">Leptochilus locii</emphasis>
</taxonomicName>
is characterized by its dimorphic fronds, fertile fronds approximately four times longer or more than sterile fronds, fertile fronds long-petiolated, and laminae thickly chartaceous.
</paragraph>
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<emphasis id="F1CA3E20C959A519D1BAF8CEFF1DAC6F" bold="true" box="[136,248,1804,1826]" pageId="10" pageNumber="223">FIGURE 7</emphasis>
.
<taxonomicName id="04BE99B1C959A519D037F8CFFCD6AC6E" authority="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" authorityName="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" box="[261,819,1805,1827]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="10" pageNumber="223" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C959A519D037F8CFFE42AC6F" box="[261,423,1805,1826]" italics="true" pageId="10" pageNumber="223">Leptochilus locii</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
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,—A. Habitat.—B. Adaxial view of sterile frond.—C. Habit.—D. Abaxial view of sterile frond.—E. Abaxial view of fertile lamina.—F. Adaxial view of fertile lamina. Photo credit: Liang Zhang.
</paragraph>
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<emphasis id="F1CA3E20C959A519D18FF8B7FEEDACC2" bold="true" box="[189,264,1909,1935]" pageId="10" pageNumber="223">Plants</emphasis>
perennial, evergreen, lithophytic.
<emphasis id="F1CA3E20C959A519D34DF8B7FD0CACC2" bold="true" box="[639,745,1909,1935]" pageId="10" pageNumber="223">Rhizome</emphasis>
long creeping, ca.
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in diam., dorsiventrally flattened, scaled, with a few short phyllopodia and roots. Apex of rhizome and basal petiole densely scaly, scales lanceolate, base enlarged, (0.7)1.23.3 ×
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, brown, basifixed, concolorous, margin entire, apex long acuminate.
<emphasis id="F1CA3E20C959A519D464F87FFA4EAC9A" bold="true" box="[1366,1451,1981,2007]" pageId="10" pageNumber="223">Fronds</emphasis>
dimorphic, fertile fronds much longer than sterile fronds.
<emphasis id="F1CA3E20C959A519D22CF823FC27ACB6" bold="true" box="[798,962,2017,2043]" pageId="10" pageNumber="223">Sterile fronds</emphasis>
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tall, laminae slightly longer than or the same length as petioles; laminae triangular-ovate, base truncate, apex obtuse-acuminate,
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long,
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wide near base, thickly chartaceous, glabrous;
<emphasis id="F1CA3E20C958A518D3F7FF5CFCFAABF5" bold="true" box="[709,799,158,184]" pageId="11" pageNumber="224">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
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in diam.,
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long.
<emphasis id="F1CA3E20C958A518D05DFF00FDF6AB91" bold="true" box="[367,531,194,220]" pageId="11" pageNumber="224">Fertile fronds</emphasis>
<quantity id="04464FD7C958A518D328FF00FD6CAB90" box="[538,649,194,221]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="11" pageNumber="224" unit="cm" value="22.0" valueMax="25.0" valueMin="19.0">1925 cm</quantity>
tall, petioles much longer than laminae;
<emphasis id="F1CA3E20C958A518D561FF00FB57AB91" bold="true" box="[1107,1202,194,220]" pageId="11" pageNumber="224">laminae</emphasis>
ovate, margins entire, base cuneate, apex long acuminate, (1.6)
<quantity id="04464FD7C958A518D36BFF24FD3CAA4C" box="[601,729,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.5" metricValueMin="3.9" pageId="11" pageNumber="224" unit="cm" value="4.2" valueMax="4.5" valueMin="3.9">3.94.5 cm</quantity>
long, (0.8)
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near base, thickly chartaceous, glabrous;
<emphasis id="F1CA3E20C958A518D1BAFEC8FF07AA69" bold="true" box="[136,226,266,292]" pageId="11" pageNumber="224">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
<quantity id="04464FD7C958A518D3F7FEC8FCA9AA68" box="[709,844,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="11" pageNumber="224" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.91.1 mm</quantity>
in diam.,
<quantity id="04464FD7C958A518D28CFEC8FBC8AA68" box="[958,1069,266,293]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="11" pageNumber="224" unit="cm" value="19.0" valueMax="20.0" valueMin="18.0">1820 cm</quantity>
long.
<emphasis id="F1CA3E20C958A518D547FEC8FB50AA69" bold="true" box="[1141,1205,266,292]" pageId="11" pageNumber="224">Veins</emphasis>
anastomosing, lateral vein distinct, veinlets visible.
<emphasis id="F1CA3E20C958A518D0E6FEECFDE3AA05" bold="true" box="[468,518,302,328]" pageId="11" pageNumber="224">Sori</emphasis>
linear, ca.
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from margins and
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from midrib, 912 on each side of midrib,
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long,
<quantity id="04464FD7C958A518D0FAFE90FDB5AA20" box="[456,592,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="11" pageNumber="224" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.91.1 mm</quantity>
wide when mature, at angles of 4250 degree with midribs, centers ca.
<quantity id="04464FD7C958A518D4B3FE90FF51AADC" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="11" pageNumber="224" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
apart from one another. Paraphyses absent.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFE58FB58AAB1" pageId="11" pageNumber="224" type="materials_examined">
<paragraph id="C301E232C958A518D18FFE58FB58AAB1" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFE58FDDBAAF9" bold="true" box="[189,574,410,436]" pageId="11" pageNumber="224">Additional specimens examined:</emphasis>
<collectingCountry id="BBA9A2A2C958A518D376FE58FD2CAAF9" box="[580,713,410,436]" name="Vietnam" pageId="11" pageNumber="224">VIETNAM</collectingCountry>
.
<collectingRegion id="017A2CD0C958A518D3E0FE58FCBBAAF9" box="[722,862,410,436]" country="Vietnam" name="Quang Binh" pageId="11" pageNumber="224">Quang Binh</collectingRegion>
: Bo Trach District, Hung Trach, Phong Nha-Ke Bang National Park,
<quantity id="04464FD7C958A518D002FE7CFE54AA94" box="[304,433,446,473]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="11" pageNumber="224" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150200 m</quantity>
,
<geoCoordinate id="A68A84F5C958A518D08FFE7CFDF9AA94" box="[445,540,446,473]" degrees="17.50" direction="north" orientation="latitude" pageId="11" pageNumber="224" precision="555" value="17.5">17.50°N</geoCoordinate>
/106.28°, on limestone rock in slightly logged primary forest,
<date id="B700C4F2C958A518D5E2FE7CFA42AA95" box="[1232,1447,446,472]" pageId="11" pageNumber="224" value="2004-12-12">12 December 2004</date>
,
<emphasis id="F1CA3E20C958A518D1BAFE21FBF7AAB1" box="[136,1042,482,509]" italics="true" pageId="11" pageNumber="224">S.-K. Wu, L.K. Phan, X. Gong, J.-Y. Xiang, V.T. Nguyen &amp; K.S. Nguyen WP-1077</emphasis>
(KUN!, MO!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFDC4FCDFA909" pageId="11" pageNumber="224" type="distribution">
<paragraph id="C301E232C958A518D18FFDC4FCDFA909" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFDC4FE0BA96D" bold="true" box="[189,494,518,544]" pageId="11" pageNumber="224">Distribution and habitat:</emphasis>
Three populations of
<taxonomicName id="04BE99B1C958A518D3DFFDC4FC4EA96D" box="[749,939,518,544]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C958A518D3DFFDC4FC4EA96D" box="[749,939,518,544]" italics="true" pageId="11" pageNumber="224">Leptochilus locii</emphasis>
</taxonomicName>
were discovered in central
<collectingCountry id="BBA9A2A2C958A518D5DBFDC4FAAEA96D" box="[1257,1355,518,544]" name="Vietnam" pageId="11" pageNumber="224">Vietnam</collectingCountry>
, and the species was exclusively observed growing on limestone rock.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFD8CFEE3A80D" pageId="11" pageNumber="224" type="etymology">
<paragraph id="C301E232C958A518D18FFD8CFD3FA9C1" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFD8CFEDBA925" bold="true" box="[189,318,590,616]" pageId="11" pageNumber="224">Etymology</emphasis>
: Named in honor of Professor Phan Ke Loc of
<collectingRegion id="017A2CD0C958A518D259FD8CFC4AA925" box="[875,943,590,616]" country="Vietnam" name="Ha Noi" pageId="11" pageNumber="224">Hanoi</collectingRegion>
University of Science, in recognition of his significant contributions to the fern flora of
<collectingCountry id="BBA9A2A2C958A518D346FDB0FD32A9C1" box="[628,727,626,652]" name="Vietnam" pageId="11" pageNumber="224">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="C301E232C958A518D18FFD54FEE3A80D" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFD54FF1AA9FD" bold="true" box="[189,255,662,688]" pageId="11" pageNumber="224">Note:</emphasis>
<taxonomicName id="04BE99B1C958A518D037FD54FE5AA9FD" box="[261,447,662,688]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C958A518D037FD54FE5AA9FD" box="[261,447,662,688]" italics="true" pageId="11" pageNumber="224">Leptochilus locii</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="04BE99B1C958A518D3B8FD55FD15A9FD" authorityName="Copeland" authorityYear="1914" box="[650,752,663,688]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C958A518D3B8FD55FD15A9FD" box="[650,752,663,688]" italics="true" pageId="11" pageNumber="224">L. ovatus</emphasis>
</taxonomicName>
by sharing features such as dimorphic fronds, sterile fronds ca. 46 times larger than fertile fronds, and petioles of fertile fronds ca. 810 times longer than fertile laminae. However, the new species differs from the latter in having triangular-ovate sterile lamina, and lamina base slightly cordate to truncate. In contrast,
<taxonomicName id="04BE99B1C958A518D049FCC1FE06A851" authorityName="Copeland" authorityYear="1914" box="[379,483,771,796]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C958A518D049FCC1FE06A851" box="[379,483,771,796]" italics="true" pageId="11" pageNumber="224">L. ovatus</emphasis>
</taxonomicName>
has elliptic-ovate to broadly elliptic sterile lamina, and sterile lamina base gradually decrescent.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,289 @@
<document id="B87EEE108C9F3800BA440A11C0C7D4DE" ID-CLB-Dataset="302193" ID-DOI="10.11646/phytotaxa.659.3.1" ID-GBIF-Dataset="52524fe3-dfb8-4afc-829a-31089bb4e306" ID-ISSN="1179-3163" ID-Zenodo-Dep="13217894" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1722613780751" checkinUser="felipe" docAuthor="Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, Zhou, Zhuo, Thepkayson, Khamphanh, Luong, Thien Tam &amp; Zhang, Li-Bing" docDate="2024" docId="4B175324C95BA51BD1BAFCACFB50AC8C" docLanguage="en" docName="phytotaxa.659.3.1.pdf" docOrigin="Phytotaxa 659 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.659.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Leptochilus khammouanensis Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="221" masterDocId="B72E2B5CC953A513D132FFC2FFE5AB4D" masterDocTitle="Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot" masterLastPageNumber="235" masterPageNumber="213" pageNumber="221" updateTime="1725399130512" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7B3EF241ECF346BD15795818800DFD9D">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:namePart id="02899696459485966E0C3CAEEF86ABD7">Zhang, Liang</mods:namePart>
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<mods:affiliation id="4F185BAE23C699ECA98F7FA2512E1A32">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="D015F7775B6EBF27F422C374AC94244F" type="email">vnmngan@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9B1DE8BAF2E47FCBC75EFCF8ED1028BC">Zhou, Zhuo</mods:namePart>
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<mods:affiliation id="A1C806F4C6E23B7C5AA63754471C7CF6">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
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<mods:identifier id="04525CA8DE67DE8AA565913B8F78C5CA" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="C6DF3DD0263A80B16642179811955874" type="Zenodo-Dep">13217894</mods:identifier>
</mods:mods>
<treatment id="4B175324C95BA51BD1BAFCACFB50AC8C" ID-DOI="http://doi.org/10.5281/zenodo.13652236" ID-GBIF-Taxon="240245551" ID-Zenodo-Dep="13652236" LSID="urn:lsid:plazi:treatment:4B175324C95BA51BD1BAFCACFB50AC8C" httpUri="http://treatment.plazi.org/id/4B175324C95BA51BD1BAFCACFB50AC8C" lastPageNumber="221" pageId="8" pageNumber="221">
<subSubSection id="8BA4B1B9C95BA51BD1BAFCACFAC8A8C5" box="[136,1325,878,905]" pageId="8" pageNumber="221" type="nomenclature">
<paragraph id="C301E232C95BA51BD1BAFCACFAC8A8C5" blockId="8.[136,1325,878,905]" box="[136,1325,878,905]" pageId="8" pageNumber="221">
<heading id="9849555EC95BA51BD1BAFCACFAC8A8C5" box="[136,1325,878,905]" fontSize="11" level="2" pageId="8" pageNumber="221" reason="4">
4.
<taxonomicName id="04BE99B1C95BA51BD196FCACFB89A8C5" authority="Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou" authorityName="Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou" authorityYear="2024" box="[164,1132,878,905]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="khammouanensis" status="sp. nov.">
<emphasis id="F1CA3E20C95BA51BD196FCACFE1BA8C4" bold="true" box="[164,510,878,905]" italics="true" pageId="8" pageNumber="221">Leptochilus khammouanensis</emphasis>
Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou
</taxonomicName>
,
<emphasis id="F1CA3E20C95BA51BD545FCADFB29A8C5" box="[1143,1228,879,904]" italics="true" pageId="8" pageNumber="221">
<taxonomicNameLabel id="EAF9835BC95BA51BD545FCADFB29A8C5" box="[1143,1228,879,904]" pageId="8" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5B85FEB7C95BA51BD5E8FCACFAFAA8C5" box="[1242,1311,878,904]" captionStart="FIGURE 6" captionStartId="9.[136,229,1690,1712]" captionTargetBox="[163,1424,202,1653]" captionTargetId="figure-21@9.[151,1436,190,1666]" captionTargetPageId="9" captionText="FIGURE 6. Leptochilus khammouanensis Liang Zhang, Khamphanh Thepkaysone &amp; Zhuo Zhou,—A. Habitat.—B. Habit.—C. Portion of abaxial lamina showing sori.—D.Adaxial lamina.—E. Abaxial lamina lacking sori.—F.Abaxial lamina with sori.—G. Rhizome. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217906" httpUri="https://zenodo.org/record/13217906/files/figure.png" pageId="8" pageNumber="221">Fig. 6</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95BA51BD1BAFC74FB99AF54" pageId="8" pageNumber="221" type="materials_examined">
<paragraph id="C301E232C95BA51BD1BAFC74FB99AF54" blockId="8.[136,1452,950,1985]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD1BAFC74FF26A89D" bold="true" box="[136,195,950,976]" pageId="8" pageNumber="221">Type</emphasis>
:
<collectingCountry id="BBA9A2A2C95BA51BD1EAFC74FEC0A89D" box="[216,293,950,976]" name="Laos" pageId="8" pageNumber="221">LAOS</collectingCountry>
.
<collectingRegion id="017A2CD0C95BA51BD007FC74FE32A89D" box="[309,471,950,976]" country="Laos" name="Khammouan" pageId="8" pageNumber="221">Khammouane</collectingRegion>
: Kounkham District, Na Hin, on the way to Nam Sanam Waterfall, elev.
<quantity id="04464FD7C95BA51BD465FC74FA42A89C" box="[1367,1447,950,977]" metricMagnitude="2" metricUnit="m" metricValue="1.88" pageId="8" pageNumber="221" unit="m" value="188.0">188 m</quantity>
,
<geoCoordinate id="A68A84F5C95BA51BD1BAFC18FF03A8B8" box="[136,230,986,1013]" degrees="18.21" direction="north" orientation="latitude" pageId="8" pageNumber="221" precision="555" value="18.21">18.21°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C95BA51BD1DDFC18FEB2A8B8" box="[239,343,986,1013]" degrees="104.51" direction="east" orientation="longitude" pageId="8" pageNumber="221" precision="555" value="104.51">104.51°E</geoCoordinate>
, on rock in forest,
<date id="B700C4F2C95BA51BD315FC18FD4AA8B9" box="[551,687,986,1013]" pageId="8" pageNumber="221" value="2019-05-02">2 May 2019</date>
,
<emphasis id="F1CA3E20C95BA51BD38BFC18FDCCAF55" italics="true" pageId="8" pageNumber="221">Zhuo Zhou, Khamphanh Thepkaysone, Mouachanh Xayvue, Kosorn Sorsoulithanh &amp; Liang Zhang LZ202</emphasis>
(
<typeStatus id="1C055C90C95BA51BD30AFC3CFD79AF55" box="[568,668,1022,1048]" pageId="8" pageNumber="221" type="holotype">holotype</typeStatus>
KUN-1576321!,
<typeStatus id="1C055C90C95BA51BD250FC3CFC56AF55" box="[866,947,1022,1048]" pageId="8" pageNumber="221" type="isotype">isotype</typeStatus>
KUN-1576322!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95BA51BD18FFBE0FF3AAEE9" pageId="8" pageNumber="221" type="diagnosis">
<paragraph id="C301E232C95BA51BD18FFBE0FC6CAF2D" blockId="8.[136,1452,950,1985]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFBE0FED5AF71" bold="true" box="[189,304,1058,1084]" pageId="8" pageNumber="221">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C95BA51BD076FBE0FD71AF71" box="[324,660,1058,1084]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="khammouanensis">
<emphasis id="F1CA3E20C95BA51BD076FBE0FD71AF71" box="[324,660,1058,1084]" italics="true" pageId="8" pageNumber="221">Leptochilus khammouanensis</emphasis>
</taxonomicName>
is distinguished in the genus by its combination of characteristics, including small habit, simple fronds, and orbicular or elongated sori.
</paragraph>
<paragraph id="C301E232C95BA51BD18FFBA8FF3AAEE9" blockId="8.[136,1452,950,1985]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFBA8FEEDAFC9" bold="true" box="[189,264,1130,1156]" pageId="8" pageNumber="221">Plants</emphasis>
perennial, evergreen.
<emphasis id="F1CA3E20C95BA51BD330FBA8FD89AFC9" bold="true" box="[514,620,1130,1156]" pageId="8" pageNumber="221">Rhizome</emphasis>
creeping,
<quantity id="04464FD7C95BA51BD3D0FBA8FC8FAFC8" box="[738,874,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.1" metricValueMin="2.5" pageId="8" pageNumber="221" unit="mm" value="3.3" valueMax="4.1" valueMin="2.5">2.54.1 mm</quantity>
in diam., dorsiventrally flattened, scaly, with a few short roots; scales narrowly ovate or lanceolate, brown, 1.33.7 ×
<quantity id="04464FD7C95BA51BD24FFB4CFBEDAFE4" box="[893,1032,1166,1193]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="8" pageNumber="221" unit="mm" value="0.75" valueMax="1.1" valueMin="0.4">0.41.1 mm</quantity>
, margin entire, apex acute, clathrate.
<emphasis id="F1CA3E20C95BA51BD1BAFB70FF38AF81" bold="true" box="[136,221,1202,1228]" pageId="8" pageNumber="221">Fronds</emphasis>
monomorphic, clustered,
<quantity id="04464FD7C95BA51BD33AFB70FD8EAF80" box="[520,619,1202,1229]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="8" pageNumber="221" unit="cm" value="11.0" valueMax="14.0" valueMin="8.0">814 cm</quantity>
long, winged stipes shorted or nearly the same length as lamina petiole.
<emphasis id="F1CA3E20C95BA51BD1BAFB14FF17AFBD" bold="true" box="[136,242,1238,1264]" pageId="8" pageNumber="221">Laminae</emphasis>
simple, margins entire, rarely lobed, elliptic-lanceolate, abaxially not scaly, 69 ×
<quantity id="04464FD7C95BA51BD5A8FB14FAFEAFBC" box="[1178,1307,1238,1265]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="8" pageNumber="221" unit="cm" value="1.6" valueMax="1.9" valueMin="1.3">1.31.9 cm</quantity>
, chartaceous or thickly herbaceous, base narrowly decrescent;
<emphasis id="F1CA3E20C95BA51BD380FB38FCE9AE59" bold="true" box="[690,780,1274,1300]" pageId="8" pageNumber="221">petioles</emphasis>
(0.5)
<quantity id="04464FD7C95BA51BD26AFB38FC4BAE58" box="[856,942,1274,1301]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="8" pageNumber="221" unit="cm" value="5.0" valueMax="8.0" valueMin="2.0">28 cm</quantity>
, winged, wings towards petiole base, slightly scaly abaxially, scales similar to but smaller than rhizome scales; midribs distinctly raised.
<emphasis id="F1CA3E20C95BA51BD5A5FADCFAE4AE75" bold="true" box="[1175,1281,1310,1336]" pageId="8" pageNumber="221">Venation</emphasis>
anastomosing, main veins distinct, veinlets slightly visible.
<emphasis id="F1CA3E20C95BA51BD3B3FA80FD56AE11" bold="true" box="[641,691,1346,1372]" pageId="8" pageNumber="221">Sori</emphasis>
separate, irregularly scattered, 0.41.9 ×
<quantity id="04464FD7C95BA51BD5BBFA80FAF6AE10" box="[1161,1299,1346,1373]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="8" pageNumber="221" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.30.5 mm</quantity>
, forming 26 irregular rows between lateral veins, orbicular or elongate, superficial, absent in marginal areoles, present in costal areoles.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95BA51BD18FFA6CFC2EAD5D" pageId="8" pageNumber="221" type="materials_examined">
<paragraph id="C301E232C95BA51BD18FFA6CFC2EAD5D" blockId="8.[136,1452,950,1985]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FFA6CFDD6AE85" bold="true" box="[189,563,1454,1480]" pageId="8" pageNumber="221">Additional specimens examined</emphasis>
:
<collectingCountry id="BBA9A2A2C95BA51BD372FA6DFD9FAE84" box="[576,634,1455,1481]" name="Laos" pageId="8" pageNumber="221">Laos</collectingCountry>
.
<collectingRegion id="017A2CD0C95BA51BD3B0FA6CFC69AE85" box="[642,908,1454,1480]" country="Laos" name="Khammouan" pageId="8" pageNumber="221">Khammouane Province</collectingRegion>
: Ban Xoang (Hin Nammo National Bio-Diversity Conservation Area), elev.
<quantity id="04464FD7C95BA51BD09EFA10FE10AEA0" box="[428,501,1490,1517]" metricMagnitude="2" metricUnit="m" metricValue="1.56" pageId="8" pageNumber="221" unit="m" value="156.0">156 m</quantity>
,
<geoCoordinate id="A68A84F5C95BA51BD333FA10FDBAAEA0" box="[513,607,1490,1517]" degrees="17.46" direction="north" orientation="latitude" pageId="8" pageNumber="221" precision="555" value="17.46">17.46°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C95BA51BD35AFA10FD35AEA0" box="[616,720,1490,1517]" degrees="105.58" direction="east" orientation="longitude" pageId="8" pageNumber="221" precision="555" value="105.58">105.58°E</geoCoordinate>
, on limestone rock in primary forests,
<date id="B700C4F2C95BA51BD5B6FA10FAAAAEA1" box="[1156,1359,1490,1516]" pageId="8" pageNumber="221" value="2017-02-16">16 February 2017</date>
,
<emphasis id="F1CA3E20C95BA51BD468FA11FF5FAD5D" italics="true" pageId="8" pageNumber="221">Yu-Min Shui</emphasis>
,
<emphasis id="F1CA3E20C95BA51BD1FAFA35FE64AD5D" box="[200,385,1526,1552]" italics="true" pageId="8" pageNumber="221">Wen-Hong Chen</emphasis>
,
<emphasis id="F1CA3E20C95BA51BD0BDFA34FC88AD5D" box="[399,877,1526,1553]" italics="true" pageId="8" pageNumber="221">Phetlasy Souladeth &amp; Liang Zhang LK136</emphasis>
(KUN).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95BA51BD18FF9D8FB95AD31" pageId="8" pageNumber="221" type="distribution">
<paragraph id="C301E232C95BA51BD18FF9D8FB95AD31" blockId="8.[136,1452,950,1985]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FF9D8FE0BAD79" bold="true" box="[189,494,1562,1588]" pageId="8" pageNumber="221">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C95BA51BD0C5F9D8FCA1AD79" box="[503,836,1562,1588]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="khammouanensis">
<emphasis id="F1CA3E20C95BA51BD0C5F9D8FCA1AD79" box="[503,836,1562,1588]" italics="true" pageId="8" pageNumber="221">Leptochilus khammouanensis</emphasis>
</taxonomicName>
is currently only documented in two locations within
<collectingRegion id="017A2CD0C95BA51BD1BAF9FCFE71AD15" box="[136,404,1598,1624]" country="Laos" name="Khammouan" pageId="8" pageNumber="221">Khammouane Province</collectingRegion>
, located in central
<collectingCountry id="BBA9A2A2C95BA51BD342F9FDFD4FAD14" box="[624,682,1599,1625]" name="Laos" pageId="8" pageNumber="221">Laos</collectingCountry>
. The species was observed growing on limestone rock in disturbed primary forests. During the extended dry season, the fronds of this species tend to shrink.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95BA51BD18FF944FB50AC8C" pageId="8" pageNumber="221" type="etymology">
<paragraph id="C301E232C95BA51BD18FF944FBF7AD89" blockId="8.[136,1452,950,1985]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FF944FEDBADED" bold="true" box="[189,318,1670,1696]" pageId="8" pageNumber="221">Etymology</emphasis>
: The species epithet is from the Laotian spelling,
<emphasis id="F1CA3E20C95BA51BD24BF944FBEBADED" box="[889,1038,1670,1696]" italics="true" pageId="8" pageNumber="221">khammouane</emphasis>
, the name of the province in central
<collectingCountry id="BBA9A2A2C95BA51BD1BAF969FF5BAD88" box="[136,190,1707,1733]" name="Laos" pageId="8" pageNumber="221">Laos</collectingCountry>
where the
<typeStatus id="1C055C90C95BA51BD008F969FE8FAD88" box="[314,362,1707,1733]" pageId="8" pageNumber="221">type</typeStatus>
was collected, and the Latin suffix -
<emphasis id="F1CA3E20C95BA51BD231F969FCDEAD89" box="[771,827,1707,1732]" italics="true" pageId="8" pageNumber="221">ensis</emphasis>
, of origin or place.
</paragraph>
<paragraph id="C301E232C95BA51BD18FF90CFB50AC8C" blockId="8.[136,1452,950,1985]" pageId="8" pageNumber="221">
<emphasis id="F1CA3E20C95BA51BD18FF90CFF10ADA5" bold="true" box="[189,245,1742,1768]" pageId="8" pageNumber="221">Note</emphasis>
:
<taxonomicName id="04BE99B1C95BA51BD037F90CFDB4ADA5" box="[261,593,1742,1768]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="khammouanensis">
<emphasis id="F1CA3E20C95BA51BD037F90CFDB4ADA5" box="[261,593,1742,1768]" italics="true" pageId="8" pageNumber="221">Leptochilus khammouanensis</emphasis>
</taxonomicName>
is morphologically most similar to some individuals of
<taxonomicName id="04BE99B1C95BA51BD5FDF90DFDDEAC41" authority="(Blume 1828: 125) Fraser-Jenkins (2008: 62)" authorityName="Fraser-Jenkins" authorityPageNumber="62" authorityYear="2008" baseAuthorityName="Blume" baseAuthorityPageNumber="125" baseAuthorityYear="1828" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="pteropus">
<emphasis id="F1CA3E20C95BA51BD5FDF90DFAAAADA5" box="[1231,1359,1743,1768]" italics="true" pageId="8" pageNumber="221">L. pteropus</emphasis>
(
<bibRefCitation id="A72F9FC3C95BA51BD46DF90CFF1BAC40" author="Blume, C." pageId="8" pageNumber="221" refId="ref11334" refString="Blume, C. (1828) Enumeratio plantarum Javae et insularum adjacentium: minus cognitarum vel novarum ex herbariis Reinwardtii, Kuhlii, hasseltii et Blumii fasc. 2. J. W. van Leeuwen, Leiden, 274 pp. https: // doi. org / 10.5962 / bhl. title. 44901" type="book" year="1828">Blume 1828: 125</bibRefCitation>
)
<bibRefCitation id="A72F9FC3C95BA51BD020F930FDDEAC41" author="Fraser-Jenkins, C. R." box="[274,571,1778,1804]" pageId="8" pageNumber="221" refId="ref11690" refString="Fraser-Jenkins, C. R. (2008) Taxonomic revision of three hundred Indian subcontinental Pteridophytes: with a revised census list; a new picture of fern-taxonomy and nomenclature in the Indian subcontinent. Bishen Singh Mahendra Pal Singh, Dehra Dun, 685 pp." type="book" year="2008">Fraser-Jenkins (2008: 62)</bibRefCitation>
</taxonomicName>
, which have simple fronds. However,
<taxonomicName id="04BE99B1C95BA51BD531F931FB00AC41" box="[1027,1253,1778,1804]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="khammouanensis">
<emphasis id="F1CA3E20C95BA51BD531F931FB00AC41" box="[1027,1253,1778,1804]" italics="true" pageId="8" pageNumber="221">L. khammouanensis</emphasis>
</taxonomicName>
differs the latter by having sparely scaly to nearly glabrous petioles and rachises.
<bibRefCitation id="A72F9FC3C95BA51BD241F8D4FBB7AC7D" author="Zhang, L. &amp; Lu, N. T. &amp; Zhou, X. - M. &amp; Chen, D. - K. &amp; Knapp, R. &amp; Zhou, L. &amp; Guo, L. &amp; Luong, T. T. &amp; Sun, H. &amp; Gao, X. - F. &amp; Zhang, L. - B." box="[883,1106,1814,1840]" pageId="8" pageNumber="221" pagination="311 - 322" refId="ref12520" refString="Zhang, L., Lu, N. T., Zhou, X. - M., Chen, D. - K., Knapp, R., Zhou, L., Guo, L., Luong, T. T., Sun, H., Gao, X. - F. &amp; Zhang, L. - B. (2019) A plastid phylogeny of the Old World fern genus Leptochilus (Polypodiaceae): Implications for cryptic speciation and progressive colonization from lower to higher latitudes. Molecular Phylogenetics and Evolution 134: 311 - 322. https: // doi. org / 10.1016 / j. ympev. 2019.01.013" type="journal article" year="2019">
Zhang
<emphasis id="F1CA3E20C95BA51BD2F6F8D5FC1BAC7D" box="[964,1022,1814,1840]" italics="true" pageId="8" pageNumber="221">et al.</emphasis>
(2019)
</bibRefCitation>
recognized a monospecific
<taxonomicName id="04BE99B1C95BA51BD4A7F8D5FF0CAC19" authorityName="Fraser-Jenkins" authorityYear="2008" baseAuthorityName="Blume" baseAuthorityYear="1828" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="pteropus">
<emphasis id="F1CA3E20C95BA51BD4A7F8D5FF0CAC19" italics="true" pageId="8" pageNumber="221">L. pteropus</emphasis>
</taxonomicName>
clade, which is one of the earliest-diverging lineages within
<taxonomicName id="04BE99B1C95BA51BD29EF8F8FBCAAC19" authorityName="Kaulfuss" authorityYear="1824" box="[940,1071,1850,1876]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="genus">
<emphasis id="F1CA3E20C95BA51BD29EF8F8FBCAAC19" box="[940,1071,1850,1876]" italics="true" pageId="8" pageNumber="221">Leptochilus</emphasis>
</taxonomicName>
. Our new phylogeny resolved
<taxonomicName id="04BE99B1C95BA51BD4A7F8F9FEACAC35" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="khammouanensis">
<emphasis id="F1CA3E20C95BA51BD4A7F8F9FEACAC35" italics="true" pageId="8" pageNumber="221">L. khammouanensis</emphasis>
</taxonomicName>
as sister to
<emphasis id="F1CA3E20C95BA51BD0F8F89DFDABAC35" box="[458,590,1887,1912]" italics="true" pageId="8" pageNumber="221">
<taxonomicName id="04BE99B1C95BA51BD0F8F89DFDAEAC35" authorityName="Fraser-Jenkins" authorityYear="2008" baseAuthorityName="Blume" baseAuthorityYear="1828" box="[458,587,1887,1912]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="pteropus">L. pteropus</taxonomicName>
,
</emphasis>
thus adding one more species to the
<taxonomicName id="04BE99B1C95BA51BD2D4F89DFB86AC35" authorityName="Fraser-Jenkins" authorityYear="2008" baseAuthorityName="Blume" baseAuthorityYear="1828" box="[998,1123,1887,1912]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="pteropus">
<emphasis id="F1CA3E20C95BA51BD2D4F89DFB86AC35" box="[998,1123,1887,1912]" italics="true" pageId="8" pageNumber="221">L. pteropus</emphasis>
</taxonomicName>
clade. Our observations of
<taxonomicName id="04BE99B1C95BA51BD4A7F89DFF0CACD1" authorityName="Fraser-Jenkins" authorityYear="2008" baseAuthorityName="Blume" baseAuthorityYear="1828" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="pteropus">
<emphasis id="F1CA3E20C95BA51BD4A7F89DFF0CACD1" italics="true" pageId="8" pageNumber="221">L. pteropus</emphasis>
</taxonomicName>
from both basic and acidic mountains showed similar morphological similarities: variable leaf shape but with dense scales on the petioles and rachises, which can be differentiated from
<taxonomicName id="04BE99B1C95BA51BD2E2F865FB4AAC8D" box="[976,1199,1958,1984]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="8" pageNumber="221" phylum="Tracheophyta" rank="species" species="khammouanensis">
<emphasis id="F1CA3E20C95BA51BD2E2F865FB4AAC8D" box="[976,1199,1958,1984]" italics="true" pageId="8" pageNumber="221">L. khammouanensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,381 @@
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<mods:title id="D1B261997D9D55595CD3A87E99020ACE">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:affiliation id="8719F31BBB2F5CCE53847DD214E27B14">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
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9.
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<emphasis id="F1CA3E20C95CA51CD196FB38FE05AE59" bold="true" box="[164,480,1274,1301]" italics="true" pageId="15" pageNumber="228">Leptochilus sinovietnamica</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
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,
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<taxonomicNameLabel id="EAF9835BC95CA51CD281FB39FBEDAE59" box="[947,1032,1275,1300]" pageId="15" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5B85FEB7C95CA51CD525FB38FB8DAE59" box="[1047,1128,1274,1300]" captionStart="FIGURE 11" captionStartId="16.[136,229,1586,1608]" captionTargetBox="[239,1348,256,1551]" captionTargetId="figure-54@16.[228,1359,245,1562]" captionTargetPageId="16" captionText="FIGURE 11. Leptochilus sinovietnamica Liang Zhang, N.T. Lu &amp; Li Bing Zhang,—A&amp; B.Abnormal individuals with one pair of lobes.— C. Portion of adaxial lamina.—D. Upper portion of abaxial lamina showing sori.—E. Portion of abaxial lamina lobe.—F. Rhizome.—G. Rhizome scales. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217916" httpUri="https://zenodo.org/record/13217916/files/figure.png" pageId="15" pageNumber="228">Fig. 11</figureCitation>
).
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<emphasis id="F1CA3E20C95CA51CD1BAFA80FF26AE11" bold="true" box="[136,195,1346,1372]" pageId="15" pageNumber="228">Type</emphasis>
:
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.
<collectingRegion id="017A2CD0C95CA51CD008FA81FE71AE10" box="[314,404,1347,1373]" country="China" name="Yunnan" pageId="15" pageNumber="228">Yunnan</collectingRegion>
: Hekou, Nanxi Town, Anjiahe, elev.
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,
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, on limestone rocks beside a stream in forest,
<date id="B700C4F2C95CA51CD079FAA4FDE1AECD" box="[331,516,1382,1408]" pageId="15" pageNumber="228" value="2019-08-31">31 August 2019</date>
,
<emphasis id="F1CA3E20C95CA51CD320FAA5FB02AECD" box="[530,1255,1382,1409]" italics="true" pageId="15" pageNumber="228">Liang Zhang, Zhen-Long Liang, Xiang-Li Yue &amp; Zi-Yu Ye 3040</emphasis>
(
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KUN-1576330!,
<typeStatus id="1C055C90C95CA51CD033FA48FEB7AEE9" box="[257,338,1418,1444]" pageId="15" pageNumber="228" type="isotype">isotype</typeStatus>
KUN-1576329!).
</paragraph>
</subSubSection>
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<emphasis id="F1CA3E20C95CA51CD18FFA6CFED5AE85" bold="true" box="[189,304,1454,1480]" pageId="15" pageNumber="228">Diagnosis</emphasis>
:
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<emphasis id="F1CA3E20C95CA51CD072FA6CFD90AE85" box="[320,629,1454,1480]" italics="true" pageId="15" pageNumber="228">Leptochilus sinovietnamica</emphasis>
</taxonomicName>
is characterized by its large habit, fronds
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long, lamina thinly chartaceous, and lobes large,
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in width.
</paragraph>
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<emphasis id="F1CA3E20C95CA51CD18FFA34FEEDAD5D" bold="true" box="[189,264,1526,1552]" pageId="15" pageNumber="228">Plants</emphasis>
perennial, evergreen.
<emphasis id="F1CA3E20C95CA51CD0CEFA34FD83AD5D" bold="true" box="[508,614,1526,1552]" pageId="15" pageNumber="228">Rhizome</emphasis>
creeping, ca.
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in diam., dorsiventrally flattened, scaly, with many short roots; scales, appressed, slightly spreading at rhizome apex, irregularly ovate, clathrate, thickly membranous, margins entire or sparely denticulate, brown, (0.8)1.83.5 (4.6) ×
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.
<emphasis id="F1CA3E20C95CA51CD521F9FCFB8DAD15" bold="true" box="[1043,1128,1598,1624]" pageId="15" pageNumber="228">Fronds</emphasis>
monomorphic or sometimes slightly dimorphic,
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distant, (30)
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long;
<emphasis id="F1CA3E20C95CA51CD27CF9A0FC4DAD31" bold="true" box="[846,936,1634,1660]" pageId="15" pageNumber="228">petioles</emphasis>
stramineous, (16)
<quantity id="04464FD7C95CA51CD54EF9A0FB08AD30" box="[1148,1261,1634,1661]" metricMagnitude="-1" metricUnit="m" metricValue="3.95" metricValueMax="4.4" metricValueMin="3.5" pageId="15" pageNumber="228" unit="cm" value="39.5" valueMax="44.0" valueMin="35.0">3544 cm</quantity>
long, ca.
<quantity id="04464FD7C95CA51CD465F9A0FF51ADEC" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="15" pageNumber="228" unit="mm" value="2.75" valueMax="3.1" valueMin="2.4">2.43.1 mm</quantity>
in diam. at middle, wingless;
<emphasis id="F1CA3E20C95CA51CD33FF944FD88ADED" bold="true" box="[525,621,1670,1696]" pageId="15" pageNumber="228">rachises</emphasis>
wingless, and only slightly winged at distal rachis at the connecting part between rachis and lobe.
<emphasis id="F1CA3E20C95CA51CD097F968FDEAAD89" bold="true" box="[421,527,1706,1732]" pageId="15" pageNumber="228">Laminae</emphasis>
broadly elliptic, deeply 1-pinnatifid, (18)3156 ×
<quantity id="04464FD7C95CA51CD567F968FB22AD88" box="[1109,1223,1706,1733]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="15" pageNumber="228" unit="cm" value="40.0" valueMax="50.0" valueMin="30.0">3050 cm</quantity>
, widest at middle or below middle of laminae, thinly chartaceous, brown when dry, glabrous, margins of lobes entire, slightly repand.
<emphasis id="F1CA3E20C95CA51CD456F90CFA49ADA5" bold="true" box="[1380,1452,1742,1768]" pageId="15" pageNumber="228">Lobes</emphasis>
(1)26 pairs, lanceolate, apex acuminate, 1526 × (2.1)
<quantity id="04464FD7C95CA51CD212F930FC40AC40" box="[800,933,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="15" pageNumber="228" unit="cm" value="4.5" valueMax="5.5" valueMin="3.5">3.55.5 cm</quantity>
, basal portions of lobes narrowed, ascending upward, midribs of lobes at angles of 4154 degree with rachis, distal pair of lobes slightly narrowed, alternate or nearly opposite, lobe base ca.
<quantity id="04464FD7C95CA51CD0EEF8F8FDBCAC18" box="[476,601,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="4.9" metricValueMin="2.5" pageId="15" pageNumber="228" unit="cm" value="3.7" valueMax="4.9" valueMin="2.5">2.54.9 cm</quantity>
distant with each other. Midribs distinctly raised, veins visible.
<emphasis id="F1CA3E20C95CA51CD41EF8F8FABBAC19" bold="true" box="[1324,1374,1850,1876]" pageId="15" pageNumber="228">Sori</emphasis>
linear, 1825 on each side of midrib, centers ca.
<quantity id="04464FD7C95CA51CD369F89CFD5DAC34" box="[603,696,1886,1913]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="15" pageNumber="228" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
apart from one another, 1.32.8 ×
<quantity id="04464FD7C95CA51CD50FF89CFB22AC34" box="[1085,1223,1886,1913]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="15" pageNumber="228" unit="mm" value="0.7" valueMax="0.9" valueMin="0.5">0.50.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95CA503D18FF840FD3CAB91" lastPageId="16" lastPageNumber="229" pageId="15" pageNumber="228" type="materials_examined">
<paragraph id="C301E232C95CA503D18FF840FD3CAB91" blockId="15.[136,1452,1346,2093]" lastBlockId="16.[136,1451,158,221]" lastPageId="16" lastPageNumber="229" pageId="15" pageNumber="228">
<emphasis id="F1CA3E20C95CA51CD18FF840FDA6ACD1" bold="true" box="[189,579,1922,1948]" pageId="15" pageNumber="228">Additional specimens examined</emphasis>
:
<materialsCitation id="73D6E86FC95CA51CD36AF840FC72AC8D" ID-GBIF-Occurrence="4928466318" collectingDate="1962-08-25" collectionCode="KUN" collectorName="Sugong Wu" country="China" county="Jinping" elevation="650" location="Jiushiliang" municipality="Mengla" pageId="15" pageNumber="228" specimenCount="1" stateProvince="Yunnan">
<collectingCountry id="BBA9A2A2C95CA51CD36AF840FD54ACD1" box="[600,689,1922,1948]" name="China" pageId="15" pageNumber="228">CHINA</collectingCountry>
.
<collectingRegion id="017A2CD0C95CA51CD3F6F841FCFBACD0" box="[708,798,1923,1949]" country="China" name="Yunnan" pageId="15" pageNumber="228">Yunnan</collectingRegion>
:
<collectingCounty id="2A609ABEC95CA51CD21CF840FC66ACD1" box="[814,899,1922,1948]" pageId="15" pageNumber="228">Jinping</collectingCounty>
,
<collectingMunicipality id="23657848C95CA51CD2A7F840FC08ACD1" box="[917,1005,1922,1948]" pageId="15" pageNumber="228">Mengla</collectingMunicipality>
,
<location id="C661B4E9C95CA51CD2CDF840FB9EACD1" LSID="urn:lsid:plazi:treatment:4B175324C95CA503D1BAFB38FC11ACB4:C661B4E9C95CA51CD2CDF840FB9EACD1" box="[1023,1147,1922,1948]" country="China" county="Jinping" municipality="Mengla" name="Jiushiliang" pageId="15" pageNumber="228" stateProvince="Yunnan">Jiushiliang</location>
, elev.
<quantity id="04464FD7C95CA51CD5FFF840FAF9ACD0" box="[1229,1308,1922,1949]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="228" unit="m" value="650.0">
<elevation id="48930501C95CA51CD5FFF840FAF9ACD0" box="[1229,1308,1922,1949]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="228" unit="m" value="650.0">650 m</elevation>
</quantity>
, on rock in limestone mountains,
<date id="B700C4F2C95CA51CD04DF864FDD1AC8D" box="[383,564,1958,1985]" pageId="15" pageNumber="228" value="1962-08-25">
<collectingDate id="A7443D1AC95CA51CD04DF864FDD1AC8D" box="[383,564,1958,1985]" pageId="15" pageNumber="228" value="1962-08-25">25 August 1962</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD30DF865FD1BAC8D" box="[575,766,1958,1985]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD30DF865FD5AAC8D" box="[575,703,1959,1985]" pageId="15" pageNumber="228">Sugong Wu</collectorName>
4044
</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD23CF864FCB1AC8D" box="[782,852,1958,1984]" country="China" lsid="urn:lsid:biocol.org:col:15842" name="Kunming Institute of Botany, Chinese Academy of Sciences" pageId="15" pageNumber="228" type="Herbarium">KUN</collectionCode>
!, PE!)
</materialsCitation>
.
<materialsCitation id="73D6E86FC95CA51CD296F864FA42ACA9" ID-GBIF-Occurrence="4928466311" collectingDate="2013-12-05" collectionCode="CDBI" collectorName="Li-Bing Zhang &amp; Liang Zhang &amp; Ngan Thi Lu" country="Vietnam" county="Na Ri District" elevation="400" location="Tham Mu" municipality="An Tinh Community" pageId="15" pageNumber="228" specimenCount="1" stateProvince="Bac Kan">
<collectingCountry id="BBA9A2A2C95CA51CD296F864FBCCAC8D" box="[932,1065,1958,1984]" name="Vietnam" pageId="15" pageNumber="228">VIETNAM</collectingCountry>
.
<collectingRegion id="017A2CD0C95CA51CD501F865FB7CAC8C" box="[1075,1177,1959,1985]" country="Vietnam" name="Bac Kan" pageId="15" pageNumber="228">Bac Kan</collectingRegion>
:
<collectingCounty id="2A609ABEC95CA51CD596F865FAA0AC8D" box="[1188,1349,1958,1985]" pageId="15" pageNumber="228">Na Ri District</collectingCounty>
,
<collectingMunicipality id="23657848C95CA51CD47CF864FEEAACA9" pageId="15" pageNumber="228">An Tinh Community</collectingMunicipality>
,
<location id="C661B4E9C95CA51CD029F808FE6BACA8" LSID="urn:lsid:plazi:treatment:4B175324C95CA503D1BAFB38FC11ACB4:C661B4E9C95CA51CD029F808FE6BACA8" box="[283,398,1994,2021]" country="Vietnam" county="Na Ri District" municipality="An Tinh Community" name="Tham Mu" pageId="15" pageNumber="228" stateProvince="Bac Kan">Tham Mu</location>
,
<quantity id="04464FD7C95CA51CD0A8F808FDF8ACA8" box="[410,541,1994,2021]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="15" pageNumber="228" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">
<elevation id="48930501C95CA51CD0A8F808FDF8ACA8" box="[410,541,1994,2021]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="15" pageNumber="228" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300500 m</elevation>
</quantity>
,
<date id="B700C4F2C95CA51CD318F808FD13ACA9" box="[554,758,1994,2021]" pageId="15" pageNumber="228" value="2013-12-05">
<collectingDate id="A7443D1AC95CA51CD318F808FD13ACA9" box="[554,758,1994,2021]" pageId="15" pageNumber="228" value="2013-12-05">5 December 2013</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD230F809FAD8ACA9" box="[770,1341,1994,2021]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD230F809FC4FACA9" box="[770,938,1994,2020]" pageId="15" pageNumber="228">Li-Bing Zhang</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD285F809FBADACA9" box="[951,1096,1994,2020]" pageId="15" pageNumber="228">Liang Zhang</collectorName>
&amp;
<collectorName id="6E4B87E4C95CA51CD55CF809FB1AACA9" box="[1134,1279,1994,2020]" pageId="15" pageNumber="228">Ngan Thi Lu</collectorName>
6711
</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD47FF808FA7FACA9" box="[1357,1434,1994,2020]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="15" pageNumber="228" type="Herbarium">CDBI</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC95CA51CD1BAF82DFBFFA361" ID-GBIF-Occurrence="4928466314" collectingDate="2013-11-22" collectionCode="CDBI" collectorName="Li-Bing Zhang &amp; Liang Zhang &amp; Ngan Thi Lu" country="Vietnam" county="Tan Lac District" elevation="840" latitude="20.53" location="Ngoc Son Ngo Luong Nature Reserve" longLatPrecision="761" longitude="105.18" municipality="Tam Son Community" pageId="15" pageNumber="228" specimenCount="1" stateProvince="Hoa Binh">
<collectingRegion id="017A2CD0C95CA51CD1BAF82DFF1CA345" box="[136,249,2030,2057]" country="Vietnam" name="Hoa Binh" pageId="15" pageNumber="228">Hoa Binh</collectingRegion>
:
<collectingCounty id="2A609ABEC95CA51CD031F82DFE59A345" box="[259,444,2030,2057]" pageId="15" pageNumber="228">Tan Lac District</collectingCounty>
,
<collectingMunicipality id="23657848C95CA51CD0F5F82DFD5CA345" box="[455,697,2030,2057]" pageId="15" pageNumber="228">Tam Son Community</collectingMunicipality>
,
<location id="C661B4E9C95CA51CD3F7F82DFB91A344" LSID="urn:lsid:plazi:treatment:4B175324C95CA503D1BAFB38FC11ACB4:C661B4E9C95CA51CD3F7F82DFB91A344" box="[709,1140,2030,2057]" country="Vietnam" county="Tan Lac District" latitude="20.53" longLatPrecision="761" longitude="105.18" municipality="Tam Son Community" name="Ngoc Son Ngo Luong Nature Reserve" pageId="15" pageNumber="228" stateProvince="Hoa Binh">Ngoc Son Ngo Luong Nature Reserve</location>
,
<quantity id="04464FD7C95CA51CD54DF82CFB2CA344" box="[1151,1225,2030,2057]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="15" pageNumber="228" unit="m" value="840.0">
<elevation id="48930501C95CA51CD54DF82CFB2CA344" box="[1151,1225,2030,2057]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="15" pageNumber="228" unit="m" value="840.0">840 m</elevation>
</quantity>
,
<geoCoordinate id="A68A84F5C95CA51CD5E7F82CFAD1A344" box="[1237,1332,2030,2057]" degrees="20.53" direction="north" orientation="latitude" pageId="15" pageNumber="228" precision="555" value="20.53">20.53°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C95CA51CD40FF82CFA41A344" box="[1341,1444,2030,2057]" degrees="105.18" direction="east" orientation="longitude" pageId="15" pageNumber="228" precision="555" value="105.18">105.18°E</geoCoordinate>
,
<date id="B700C4F2C95CA51CD1BAF7D0FE83A361" box="[136,358,2066,2092]" pageId="15" pageNumber="228" value="2013-11-22">
<collectingDate id="A7443D1AC95CA51CD1BAF7D0FE83A361" box="[136,358,2066,2092]" pageId="15" pageNumber="228" value="2013-11-22">22 November 2013</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD040F7D1FC4AA361" box="[370,943,2066,2093]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD040F7D1FDFFA361" box="[370,538,2066,2092]" pageId="15" pageNumber="228">Li-Bing Zhang</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD315F7D1FD5CA361" box="[551,697,2066,2092]" pageId="15" pageNumber="228">Liang Zhang</collectorName>
&amp;
<collectorName id="6E4B87E4C95CA51CD3EDF7D1FC95A361" box="[735,880,2066,2092]" pageId="15" pageNumber="228">Ngan Thi Lu</collectorName>
6377
</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD2F2F7D0FBE8A361" box="[960,1037,2066,2092]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="15" pageNumber="228" type="Herbarium">CDBI</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC95CA503D515F7D0FD30AB91" ID-GBIF-Occurrence="4928466319" collectingDate="2014-11-07" collectionCode="CDBI" collectorName="Li-Bing Zhang &amp; Liang Zhang &amp; Ngan Thi Lu" country="Vietnam" elevation="400" lastPageId="16" lastPageNumber="229" latitude="17.44" location="Road" longLatPrecision="767" longitude="106.24" municipality="Phong Nha Ke Bang National Park" pageId="15" pageNumber="228" specimenCount="1" stateProvince="Quang Binh">
<collectingRegion id="017A2CD0C95CA51CD515F7D0FB50A361" box="[1063,1205,2066,2092]" country="Vietnam" name="Quang Binh" pageId="15" pageNumber="228">Quang Binh</collectingRegion>
:
<collectingMunicipality id="23657848C95CA503D5F2F7D0FEC0ABF5" lastPageId="16" lastPageNumber="229" pageId="15" pageNumber="228">Phong Nha Ke Bang National Park</collectingMunicipality>
,
<quantity id="04464FD7C943A503D002FF5CFE95ABF5" box="[304,368,158,184]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="229" unit="km" value="27.0">27km</quantity>
of #20
<location id="C661B4E9C943A503D0F4FF5CFDE1ABF5" LSID="urn:lsid:plazi:treatment:4B175324C95CA503D1BAFB38FC11ACB4:C661B4E9C943A503D0F4FF5CFDE1ABF5" box="[454,516,158,184]" country="Vietnam" latitude="17.44" longLatPrecision="767" longitude="106.24" municipality="Phong Nha Ke Bang National Park" name="Road" pageId="16" pageNumber="229" stateProvince="Quang Binh">Road</location>
,
<quantity id="04464FD7C943A503D33DFF5CFDBDABF4" box="[527,600,158,185]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="229" unit="m" value="400.0">
<elevation id="48930501C943A503D33DFF5CFDBDABF4" box="[527,600,158,185]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="229" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<geoCoordinate id="A68A84F5C943A503D351FF5CFD24ABF5" box="[611,705,158,184]" degrees="17.44" direction="north" orientation="latitude" pageId="16" pageNumber="229" precision="555" value="17.44">17.44°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C943A503D3F8FF5CFCD6ABF4" box="[714,819,158,185]" degrees="106.24" direction="east" orientation="longitude" pageId="16" pageNumber="229" precision="555" value="106.24">106.24°E</geoCoordinate>
, near rock in forest,
<date id="B700C4F2C943A503D529FF5CFB10ABF5" box="[1051,1269,158,184]" pageId="16" pageNumber="229" value="2014-11-07">
<collectingDate id="A7443D1AC943A503D529FF5CFB10ABF5" box="[1051,1269,158,184]" pageId="16" pageNumber="229" value="2014-11-07">07 November 2014</collectingDate>
</date>
,
<emphasis id="F1CA3E20C943A503D432FF5DFDEDAB91" italics="true" pageId="16" pageNumber="229">
<collectorName id="6E4B87E4C943A503D432FF5DFA42ABF5" box="[1280,1447,158,184]" pageId="16" pageNumber="229">Li-Bing Zhang</collectorName>
,
<collectorName id="6E4B87E4C943A503D1BAFF01FEFDAB91" box="[136,280,194,220]" pageId="16" pageNumber="229">Liang Zhang</collectorName>
&amp;
<collectorName id="6E4B87E4C943A503D00EFF01FE2FAB91" box="[316,458,194,220]" pageId="16" pageNumber="229">Ngan Thi Lu</collectorName>
7365
</emphasis>
(
<collectionCode id="A5AF7AF7C943A503D32AFF00FD81AB91" box="[536,612,194,220]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="16" pageNumber="229" type="Herbarium">CDBI</collectionCode>
!, VNMN)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="97C1B2BAC943A503D1BAF9F0FDCBADDD" ID-DOI="http://doi.org/10.5281/zenodo.13217916" ID-Zenodo-Dep="13217916" httpUri="https://zenodo.org/record/13217916/files/figure.png" pageId="16" pageNumber="229" startId="16.[136,229,1586,1608]" targetBox="[239,1348,256,1551]" targetPageId="16" targetType="figure">
<paragraph id="C301E232C943A503D1BAF9F0FDCBADDD" blockId="16.[136,1452,1586,1680]" pageId="16" pageNumber="229">
<emphasis id="F1CA3E20C943A503D1BAF9F0FEE4AD0A" bold="true" box="[136,257,1586,1608]" pageId="16" pageNumber="229">FIGURE 11</emphasis>
.
<taxonomicName id="04BE99B1C943A503D039F9F0FC6EAD05" authority="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" authorityName="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" box="[267,907,1586,1608]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="16" pageNumber="229" phylum="Tracheophyta" rank="species" species="sinovietnamica">
<emphasis id="F1CA3E20C943A503D039F9F0FDF4AD05" box="[267,529,1586,1608]" italics="true" pageId="16" pageNumber="229">Leptochilus sinovietnamica</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
</taxonomicName>
,—A &amp; B. Abnormal individuals with one pair of lobes.— C. Portion of adaxial lamina.—D. Upper portion of abaxial lamina showing sori.—E. Portion of abaxial lamina lobe.—F. Rhizome.—G. Rhizome scales. Photo credit: Liang Zhang.
</paragraph>
</caption>
<subSubSection id="8BA4B1B9C943A503D18FF97DFF1FAC6C" pageId="16" pageNumber="229" type="distribution">
<paragraph id="C301E232C943A503D18FF97DFF1FAC6C" blockId="16.[136,1452,1727,2041]" pageId="16" pageNumber="229">
<emphasis id="F1CA3E20C943A503D18FF97DFE17AD94" bold="true" box="[189,498,1727,1753]" pageId="16" pageNumber="229">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C943A503D0CFF97DFCD0AD94" box="[509,821,1727,1753]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="16" pageNumber="229" phylum="Tracheophyta" rank="species" species="sinovietnamica">
<emphasis id="F1CA3E20C943A503D0CFF97DFCD0AD94" box="[509,821,1727,1753]" italics="true" pageId="16" pageNumber="229">Leptochilus sinovietnamica</emphasis>
</taxonomicName>
is found in the southern region of
<collectingRegion id="017A2CD0C943A503D5E5F97DFAD5AD94" box="[1239,1328,1727,1753]" country="China" name="Yunnan" pageId="16" pageNumber="229">Yunnan</collectingRegion>
,
<collectingCountry id="BBA9A2A2C943A503D40DF97DFA60AD94" box="[1343,1413,1727,1753]" name="China" pageId="16" pageNumber="229">China</collectingCountry>
, as well as in central and northern
<collectingCountry id="BBA9A2A2C943A503D0D5F921FDAFADB0" box="[487,586,1763,1789]" name="Vietnam" pageId="16" pageNumber="229">Vietnam</collectingCountry>
. This species is observed growing on or in proximity to rocks in a limestone mountain.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C943A503D18FF8E9FC11ACB4" pageId="16" pageNumber="229" type="etymology">
<paragraph id="C301E232C943A503D18FF8E9FE82AC24" blockId="16.[136,1452,1727,2041]" pageId="16" pageNumber="229">
<emphasis id="F1CA3E20C943A503D18FF8E9FEDBAC08" bold="true" box="[189,318,1835,1861]" pageId="16" pageNumber="229">Etymology</emphasis>
: The specific epithet, sinovietnamica, indicates that the species is currently known distributed in
<collectingCountry id="BBA9A2A2C943A503D1BAF88DFF2EAC24" box="[136,203,1871,1897]" name="China" pageId="16" pageNumber="229">China</collectingCountry>
and
<collectingCountry id="BBA9A2A2C943A503D033F88DFE81AC24" box="[257,356,1871,1897]" name="Vietnam" pageId="16" pageNumber="229">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="C301E232C943A503D18FF8B1FC11ACB4" blockId="16.[136,1452,1727,2041]" pageId="16" pageNumber="229">
<emphasis id="F1CA3E20C943A503D18FF8B1FF10ACC0" bold="true" box="[189,245,1907,1933]" pageId="16" pageNumber="229">Note</emphasis>
:
<taxonomicName id="04BE99B1C943A503D037F8B1FDDFACC0" box="[261,570,1907,1933]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="16" pageNumber="229" phylum="Tracheophyta" rank="species" species="sinovietnamica">
<emphasis id="F1CA3E20C943A503D037F8B1FDDFACC0" box="[261,570,1907,1933]" italics="true" pageId="16" pageNumber="229">Leptochilus sinovietnamica</emphasis>
</taxonomicName>
is distinguished by having the large fronds up to
<quantity id="04464FD7C943A503D55EF8B1FB57ACC0" box="[1132,1202,1907,1933]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="229" unit="cm" value="90.0">90 cm</quantity>
long and 34 pairs of lateral lobes. The sample labeled as “Li-Bing Zhang
<emphasis id="F1CA3E20C943A503D3EDF85AFCFCACFC" box="[735,793,1943,1969]" italics="true" pageId="16" pageNumber="229">et al.</emphasis>
6711”, featuring one pairs of lobes, presents an exception within the species. To fully characterize the species in terms of its morphology, additional morphological studies should be conducted by incorporating a broader range of sequenced materials.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,402 @@
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8.
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<emphasis id="F1CA3E20C95DA51DD196FDC4FE59A96C" bold="true" box="[164,444,518,545]" italics="true" pageId="14" pageNumber="227">Leptochilus neolongipes</emphasis>
Liang Zhang, X.M. Zhou, T.T. Luong &amp; Li Bing Zhang
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,
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</emphasis>
(
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).
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</paragraph>
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<emphasis id="F1CA3E20C95DA51DD1BAFD8CFF26A925" bold="true" box="[136,195,590,616]" pageId="14" pageNumber="227">Type</emphasis>
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.
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: Bidoup-Nui Ba National Park, behind Hòn Giao station,
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, besides rocks near stream,
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,
<emphasis id="F1CA3E20C95DA51DD0D1FDB1FB32A9C1" box="[483,1239,626,653]" italics="true" pageId="14" pageNumber="227">Li Bing Zhang, Xin-Mao Zhou &amp; Thien Tam Luong, Minh Tri Dang</emphasis>
&amp;
<emphasis id="F1CA3E20C95DA51DD5C9FDB0FF25A9FD" italics="true" pageId="14" pageNumber="227">Bich Ha Duong 8596</emphasis>
(
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MO!,
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!).
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<emphasis id="F1CA3E20C95DA51DD1BAF829FEE0A34D" bold="true" box="[136,261,2027,2049]" pageId="14" pageNumber="227">FIGURE 10</emphasis>
.
<taxonomicName id="04BE99B1C95DA51DD020F82EFBC2A34F" authority="Liang Zhang, X. M. Zhou, T. T. Luong &amp; Li Bing Zhang" authorityName="Liang Zhang, X. M. Zhou, T. T. Luong &amp; Li Bing Zhang" box="[274,1063,2028,2050]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="14" pageNumber="227" phylum="Tracheophyta" rank="species" species="neolongipes">
<emphasis id="F1CA3E20C95DA51DD020F82EFE18A34C" box="[274,509,2028,2049]" italics="true" pageId="14" pageNumber="227">Leptochilus neolongipes</emphasis>
Liang Zhang, X.M. Zhou, T.T. Luong &amp; Li Bing Zhang
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,—A. Habitat.—B. Abaxial lamina.—C. Adaxial lamina.—D. Portion of abaxial lamina showing sori.—E. Portion of adaxial lamina. Photo credit: Xin Mao Zhou.
</paragraph>
</caption>
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<emphasis id="F1CA3E20C95CA51CD18FFF5CFED5ABF5" bold="true" box="[189,304,158,184]" pageId="15" pageNumber="228">Diagnosis</emphasis>
:
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<emphasis id="F1CA3E20C95CA51CD072FF5CFDB7ABF5" box="[320,594,158,184]" italics="true" pageId="15" pageNumber="228">Leptochilus neolongipes</emphasis>
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is characterized by having medium size of plant, fronds
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, petioles longer or nearly the same length as laminae, distal rachis broadly winged, wings gradually narrowed towards rachis base, lamina thickly chartaceous, and linear sori.
</paragraph>
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<emphasis id="F1CA3E20C95CA51CD18FFEC8FEEDAA69" bold="true" box="[189,264,266,292]" pageId="15" pageNumber="228">Plants</emphasis>
perennial, evergreen. Rhizome creeping, ca.
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in diam., dorsiventrally flattened, densely scaled, with many short roots; scales brown, clathrate, membranous, margins entire or sparely denticulate, appressed on rhizome or slightly spreading, 0.82.7 ×
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.
<emphasis id="F1CA3E20C95CA51CD356FE90FD5CAA21" bold="true" box="[612,697,338,364]" pageId="15" pageNumber="228">Fronds</emphasis>
monomorphic,
<quantity id="04464FD7C95CA51CD259FE90FC1BAA20" box="[875,1022,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.8" metricValueMin="0.9" pageId="15" pageNumber="228" unit="cm" value="2.35" valueMax="3.8" valueMin="0.9">0.9 to 3.8 cm</quantity>
distant, 2546(62) cm long;
<emphasis id="F1CA3E20C95CA51CD463FE90FA4EAA21" bold="true" box="[1361,1451,338,364]" pageId="15" pageNumber="228">petioles</emphasis>
stramineous,
<quantity id="04464FD7C95CA51CD02EFEB4FE6FAADC" box="[284,394,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.7" metricValueMin="1.7" pageId="15" pageNumber="228" unit="cm" value="22.0" valueMax="27.0" valueMin="17.0">1727 cm</quantity>
long, ca.
<quantity id="04464FD7C95CA51CD0C6FEB4FD9FAADC" box="[500,634,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="15" pageNumber="228" unit="mm" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.61.2 mm</quantity>
in diam. at middle, slightly winged at distal part; rachis distinctly winged, 3.09.0 mm wide at the distal part,
<quantity id="04464FD7C95CA51CD33FFE58FD76AAF8" box="[525,659,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="15" pageNumber="228" unit="mm" value="3.5" valueMax="4.5" valueMin="2.5">2.54.5 mm</quantity>
at base.
<emphasis id="F1CA3E20C95CA51CD3C3FE58FCBEAAF9" bold="true" box="[753,859,410,436]" pageId="15" pageNumber="228">Laminae</emphasis>
oblong-ovate, deeply 1-pinnatifid, 820 ×
<quantity id="04464FD7C95CA51CD404FE58FA42AAF8" box="[1334,1447,410,437]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="15" pageNumber="228" unit="cm" value="23.0" valueMax="30.0" valueMin="16.0">1630 cm</quantity>
, widest below middle of laminae, chartaceous, brown when dry, glabrous, margins of lobes entire.
<emphasis id="F1CA3E20C95CA51CD5F0FE7CFAEFAA95" bold="true" box="[1218,1290,446,472]" pageId="15" pageNumber="228">Lobes</emphasis>
(1)35 pairs, lanceolate, apex acuminate, 1215 ×
<quantity id="04464FD7C95CA51CD315FE20FD42AAB0" box="[551,679,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="15" pageNumber="228" unit="cm" value="2.15" valueMax="2.4" valueMin="1.9">1.92.4 cm</quantity>
, basal portions of lobes obviously narrowed, ca. 1/3 to 1/4 as wide as middle portions, slightly ascending upward, midribs of lobes at angles of 5573(90) degree with rachis, distal pair of lobes slightly narrowed or the same length as lower pairs, opposite or slightly alternate, lobe base ca.
<quantity id="04464FD7C95CA51CD5EFFDE8FABEA908" box="[1245,1371,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="15" pageNumber="228" unit="cm" value="2.15" valueMax="2.4" valueMin="1.9">1.92.4 cm</quantity>
distant with each other. Midribs slightly raised, lateral veins invisible.
<emphasis id="F1CA3E20C95CA51CD27DFD8CFC64A925" bold="true" box="[847,897,590,616]" pageId="15" pageNumber="228">Sori</emphasis>
linear, (9)1619 on each side of midrib, 812 ×
<quantity id="04464FD7C95CA51CD1BAFDB0FEF4A9C0" box="[136,273,626,653]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="15" pageNumber="228" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.60.9 mm</quantity>
, centers ca.
<quantity id="04464FD7C95CA51CD0A8FDB0FE12A9C0" box="[410,503,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="15" pageNumber="228" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
apart from one another.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95CA51CD18FFD54FB2CA999" pageId="15" pageNumber="228" type="distribution">
<paragraph id="C301E232C95CA51CD18FFD54FB2CA999" blockId="15.[136,1452,158,1229]" pageId="15" pageNumber="228">
<emphasis id="F1CA3E20C95CA51CD18FFD54FE0BA9FD" bold="true" box="[189,494,662,688]" pageId="15" pageNumber="228">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C95CA51CD0C5FD54FCEFA9FD" box="[503,778,662,688]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="neolongipes">
<emphasis id="F1CA3E20C95CA51CD0C5FD54FCEFA9FD" box="[503,778,662,688]" italics="true" pageId="15" pageNumber="228">Leptochilus neolongipes</emphasis>
</taxonomicName>
was discovered in three provinces in southern
<collectingCountry id="BBA9A2A2C95CA51CD41AFD54FA6FA9FD" box="[1320,1418,662,688]" name="Vietnam" pageId="15" pageNumber="228">Vietnam</collectingCountry>
, at elevations between
<quantity id="04464FD7C95CA51CD057FD78FDE6A998" box="[357,515,698,725]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="15" pageNumber="228" unit="m" value="1500.0" valueMax="1700.0" valueMin="1300.0">13001700 m</quantity>
. It typically grows on rock sides or near the base of tree trunks.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95CA51CD18FFD1CFB48A850" pageId="15" pageNumber="228" type="etymology">
<paragraph id="C301E232C95CA51CD18FFD1CFB48A850" blockId="15.[136,1452,158,1229]" pageId="15" pageNumber="228">
<emphasis id="F1CA3E20C95CA51CD18FFD1CFEDBA9B5" bold="true" box="[189,318,734,760]" pageId="15" pageNumber="228">Etymology</emphasis>
: From the Latin prefix,
<emphasis id="F1CA3E20C95CA51CD378FD1DFD97A9B5" box="[586,626,735,760]" italics="true" pageId="15" pageNumber="228">neo</emphasis>
-, new, and the Latin epithet,
<taxonomicName id="04BE99B1C95CA51CD287FD1CFBF1A9B5" authorityName="X. C. Zhang" authorityYear="2012" baseAuthorityName="Ching" baseAuthorityYear="1933" box="[949,1044,734,760]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="F1CA3E20C95CA51CD287FD1CFBF1A9B5" box="[949,1044,734,760]" italics="true" pageId="15" pageNumber="228">longipes</emphasis>
</taxonomicName>
, referring to the similar morphology between the new species and
<taxonomicName id="04BE99B1C95CA51CD0E0FCC0FB4CA850" authority="(Ching 1933: 332) X. C. Zhang (2012: 655)" authorityName="X. C. Zhang" authorityPageNumber="655" authorityYear="2012" baseAuthorityName="Ching" baseAuthorityPageNumber="332" baseAuthorityYear="1933" box="[466,1193,770,797]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="F1CA3E20C95CA51CD0E0FCC0FD5EA851" box="[466,699,770,796]" italics="true" pageId="15" pageNumber="228">Leptochilus longipes</emphasis>
(
<bibRefCitation id="A72F9FC3C95CA51CD3F8FCC0FC6FA851" author="Ching, R. - C." box="[714,906,770,796]" pageId="15" pageNumber="228" pagination="293 - 362" refId="ref11548" refString="Ching, R. - C. (1933) Studies of Chinese ferns X. Bulletin of the Fan Memorial Institute of Biology 4: 293 - 362." type="journal article" year="1933">Ching 1933: 332</bibRefCitation>
) X.C.
<bibRefCitation id="A72F9FC3C95CA51CD2E6FCC0FB4CA850" author="Zhang, X. - C." box="[980,1193,770,797]" pageId="15" pageNumber="228" refId="ref12644" refString="Zhang, X. - C. (2012) Lycophytes and ferns of China. Peking University Press, Beijing, 711 pp." type="book" year="2012">Zhang (2012: 655)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95CA51CD18FFCE4FCD5AF81" pageId="15" pageNumber="228" type="materials_examined">
<paragraph id="C301E232C95CA51CD18FFCE4FACDA8B9" blockId="15.[136,1452,158,1229]" pageId="15" pageNumber="228">
<emphasis id="F1CA3E20C95CA51CD18FFCE4FDDFA80D" bold="true" box="[189,570,806,832]" pageId="15" pageNumber="228">Additional specimens examined</emphasis>
:
<materialsCitation id="73D6E86FC95CA51CD37BFCE4FDE3A8C5" ID-GBIF-Occurrence="4928466312" collectingDate="2015-10-23" collectionCode="CDBI" collectorName="Li Bing Zhang &amp; Xinmao Zhou &amp; Thien Tam Luong &amp; Minh Tri Dang &amp; Bich Ha Duong" country="Vietnam" elevation="1540" location="Hon Ba forest" municipality="Hon Ba Nature Reserve" pageId="15" pageNumber="228" specimenCount="1" stateProvince="Khanh Hoa">
<collectingCountry id="BBA9A2A2C95CA51CD37BFCE4FD2BA80D" box="[585,718,806,832]" name="Vietnam" pageId="15" pageNumber="228">VIETNAM</collectingCountry>
.
<collectingRegion id="017A2CD0C95CA51CD3E8FCE4FC2EA80D" box="[730,971,806,832]" country="Vietnam" name="Khanh Hoa" pageId="15" pageNumber="228">Khánh Hòa Province</collectingRegion>
:
<collectingMunicipality id="23657848C95CA51CD2E5FCE4FB0DA80C" box="[983,1256,806,833]" pageId="15" pageNumber="228">Hòn Bà Nature Reserve</collectingMunicipality>
, trails along
<location id="C661B4E9C95CA51CD449FCE4FF0AA829" LSID="urn:lsid:plazi:treatment:4B175324C95DA51CD1BAFDC4FCD5AF81:C661B4E9C95CA51CD449FCE4FF0AA829" country="Vietnam" municipality="Hon Ba Nature Reserve" name="Hon Ba forest" pageId="15" pageNumber="228" stateProvince="Khanh Hoa">Hòn Bà forest</location>
station,
<quantity id="04464FD7C95CA51CD07DFC88FE42A828" box="[335,423,842,869]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="15" pageNumber="228" unit="m" value="1540.0">
<elevation id="48930501C95CA51CD07DFC88FE42A828" box="[335,423,842,869]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="15" pageNumber="228" unit="m" value="1540.0">1540 m</elevation>
</quantity>
,
<date id="B700C4F2C95CA51CD081FC88FD96A828" box="[435,627,842,869]" pageId="15" pageNumber="228" value="2015-10-23">
<collectingDate id="A7443D1AC95CA51CD081FC88FD96A828" box="[435,627,842,869]" pageId="15" pageNumber="228" value="2015-10-23">23 October 2015</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD34CFC89FAB6A829" box="[638,1363,842,868]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD34CFC89FCC1A829" box="[638,804,842,868]" pageId="15" pageNumber="228">Li Bing Zhang</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD202FC89FC2FA829" box="[816,970,842,868]" pageId="15" pageNumber="228">Xinmao Zhou</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD2E4FC88FB7BA829" box="[982,1182,842,868]" pageId="15" pageNumber="228">Thien Tam Luong</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD598FC88FAB6A829" box="[1194,1363,842,868]" pageId="15" pageNumber="228">Minh Tri Dang</collectorName>
</emphasis>
&amp;
<emphasis id="F1CA3E20C95CA51CD44AFC88FEDBA8C5" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD44AFC88FF1BA8C5" pageId="15" pageNumber="228">Bich Ha Duong</collectorName>
8677
</emphasis>
&amp;
<emphasis id="F1CA3E20C95CA51CD056FCACFE79A8C5" box="[356,412,878,904]" italics="true" pageId="15" pageNumber="228">8727</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD09EFCACFE1CA8C5" box="[428,505,878,904]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="15" pageNumber="228" type="Herbarium">CDBI</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC95CA51CD320FCACFC05A89D" ID-GBIF-Occurrence="4928466317" collectingDate="2011-06-20" collectingDateMax="2015-10-24" collectingDateMin="2011-06-20" collectionCode="CDBI, TAIF" collectorName="Li Bing Zhang &amp; Xinmao Zhou &amp; Thien Tam Luong &amp; Minh Tri Dang &amp; Bich Ha Duong &amp; Cheng-Wei Chen Wade" country="Vietnam" county="Hon Ba Nature Reserve" elevation="1350" location="Da Lat" municipality="Yersin's" pageId="15" pageNumber="228" specimenCount="1" stateProvince="Lam Dong">
<collectingCounty id="2A609ABEC95CA51CD320FCACFCC7A8C4" box="[530,802,878,905]" pageId="15" pageNumber="228">Hòn Bà Nature Reserve</collectingCounty>
, stream behind
<collectingMunicipality id="23657848C95CA51CD2E1FCACFBCEA8C5" box="[979,1067,878,904]" pageId="15" pageNumber="228">Yersin's</collectingMunicipality>
house,
<quantity id="04464FD7C95CA51CD5B1FCACFAC4A8C4" box="[1155,1313,878,905]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="15" pageNumber="228" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">
<elevation id="48930501C95CA51CD5B1FCACFAC4A8C4" box="[1155,1313,878,905]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="15" pageNumber="228" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">13001400 m</elevation>
</quantity>
,
<date id="B700C4F2C95CA51CD41FFCACFF26A8E0" pageId="15" pageNumber="228" value="2015-10-24">
<collectingDate id="A7443D1AC95CA51CD41FFCACFF26A8E0" pageId="15" pageNumber="228" value="2015-10-24">24 October 2015</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD1FDFC51FC42A8E1" box="[207,935,914,940]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD1FDFC51FE90A8E1" box="[207,373,914,940]" pageId="15" pageNumber="228">Li Bing Zhang</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD0B3FC51FDF9A8E1" box="[385,540,914,940]" pageId="15" pageNumber="228">Xinmao Zhou</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD31AFC50FD14A8E1" box="[552,753,914,940]" pageId="15" pageNumber="228">Thien Tam Luong</collectorName>
,
<collectorName id="6E4B87E4C95CA51CD3CFFC50FC42A8E1" box="[765,935,914,940]" pageId="15" pageNumber="228">Minh Tri Dang</collectorName>
</emphasis>
&amp;
<emphasis id="F1CA3E20C95CA51CD2FEFC50FB58A8E1" box="[972,1213,914,940]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD2FEFC50FB98A8E1" box="[972,1149,914,940]" pageId="15" pageNumber="228">Bich Ha Duong</collectorName>
8768
</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD5FCFC50FAFFA8E1" box="[1230,1306,914,940]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="15" pageNumber="228" type="Herbarium">CDBI</collectionCode>
!, MO!).
<collectingRegion id="017A2CD0C95CA51CD44AFC51FED3A89D" country="Vietnam" name="Lam Dong" pageId="15" pageNumber="228">Lam Dong Province</collectingRegion>
:
<location id="C661B4E9C95CA51CD073FC75FE74A89C" LSID="urn:lsid:plazi:treatment:4B175324C95DA51CD1BAFDC4FCD5AF81:C661B4E9C95CA51CD073FC75FE74A89C" box="[321,401,951,977]" country="Vietnam" county="Hon Ba Nature Reserve" municipality="Yersin's" name="Da Lat" pageId="15" pageNumber="228" stateProvince="Lam Dong">Da Lat</location>
,
<date id="B700C4F2C95CA51CD0ACFC74FDD3A89D" box="[414,566,950,977]" pageId="15" pageNumber="228" value="2011-06-20">
<collectingDate id="A7443D1AC95CA51CD0ACFC74FDD3A89D" box="[414,566,950,977]" pageId="15" pageNumber="228" value="2011-06-20">20 June 2011</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD370FC74FC98A89D" box="[578,893,950,976]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD370FC74FCA3A89D" box="[578,838,950,976]" pageId="15" pageNumber="228">Cheng-Wei Chen Wade</collectorName>
1594
</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD2BFFC74FC31A89D" box="[909,980,950,976]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15478" lsid="urn:lsid:biocol.org:col:15478" name="Taiwan Forestry Research Institute" pageId="15" pageNumber="228" type="Herbarium">TAIF</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC95CA51CD2DFFC74FDC1A8B9" ID-GBIF-Occurrence="4928466309" collectingDate="2013-06-17" collectionCode="TAIF" collectorName="Cheng-Wei Chen Wade" country="Vietnam" elevation="1650" location="Giang Ly" pageId="15" pageNumber="228" specimenCount="1">
<location id="C661B4E9C95CA51CD2DFFC74FBBDA89C" LSID="urn:lsid:plazi:treatment:4B175324C95DA51CD1BAFDC4FCD5AF81:C661B4E9C95CA51CD2DFFC74FBBDA89C" box="[1005,1112,950,977]" country="Vietnam" name="Giang Ly" pageId="15" pageNumber="228">Giang Ly</location>
,
<quantity id="04464FD7C95CA51CD551FC74FAE4A89C" box="[1123,1281,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="15" pageNumber="228" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<elevation id="48930501C95CA51CD551FC74FAE4A89C" box="[1123,1281,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="15" pageNumber="228" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">16001700 m</elevation>
</quantity>
,
<date id="B700C4F2C95CA51CD43CFC74FA42A89D" box="[1294,1447,950,977]" pageId="15" pageNumber="228" value="2013-06-17">
<collectingDate id="A7443D1AC95CA51CD43CFC74FA42A89D" box="[1294,1447,950,977]" pageId="15" pageNumber="228" value="2013-06-17">17 June 2013</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD1BAFC18FE27A8B9" box="[136,450,986,1012]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD1BAFC18FE6FA8B9" box="[136,394,986,1012]" pageId="15" pageNumber="228">Cheng-Wei Chen Wade</collectorName>
2615
</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD0E3FC18FDFDA8B9" box="[465,536,986,1012]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15478" lsid="urn:lsid:biocol.org:col:15478" name="Taiwan Forestry Research Institute" pageId="15" pageNumber="228" type="Herbarium">TAIF</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="73D6E86FC95CA51CD31DFC18FAC6A8B9" ID-GBIF-Occurrence="4928466302" box="[559,1315,986,1013]" collectingDate="2014-12-24" collectionCode="TAIF" collectorName="Cheng-Wei Chen Wade" country="Vietnam" location="Giang Ly" pageId="15" pageNumber="228" specimenCount="1">
<location id="C661B4E9C95CA51CD31DFC18FD7FA8B8" LSID="urn:lsid:plazi:treatment:4B175324C95DA51CD1BAFDC4FCD5AF81:C661B4E9C95CA51CD31DFC18FD7FA8B8" box="[559,666,986,1013]" country="Vietnam" name="Giang Ly" pageId="15" pageNumber="228">Giang Ly</location>
,
<date id="B700C4F2C95CA51CD397FC18FC98A8B9" box="[677,893,986,1012]" pageId="15" pageNumber="228" value="2014-12-24">
<collectingDate id="A7443D1AC95CA51CD397FC18FC98A8B9" box="[677,893,986,1012]" pageId="15" pageNumber="228" value="2014-12-24">24 December 2014</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95CA51CD2BAFC18FB27A8B9" box="[904,1218,986,1012]" italics="true" pageId="15" pageNumber="228">
<collectorName id="6E4B87E4C95CA51CD2BAFC18FB6FA8B9" box="[904,1162,986,1012]" pageId="15" pageNumber="228">Cheng-Wei Chen Wade</collectorName>
4102
</emphasis>
(
<collectionCode id="A5AF7AF7C95CA51CD5E3FC18FAFDA8B9" box="[1233,1304,986,1012]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15478" lsid="urn:lsid:biocol.org:col:15478" name="Taiwan Forestry Research Institute" pageId="15" pageNumber="228" type="Herbarium">TAIF</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="C301E232C95CA51CD18FFC3CFCD5AF81" blockId="15.[136,1452,158,1229]" pageId="15" pageNumber="228">
<emphasis id="F1CA3E20C95CA51CD18FFC3CFF10AF55" bold="true" box="[189,245,1022,1048]" pageId="15" pageNumber="228">Note</emphasis>
:
<taxonomicName id="04BE99B1C95CA51CD034FC3CFDFCAF55" box="[262,537,1022,1048]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="neolongipes">
<emphasis id="F1CA3E20C95CA51CD034FC3CFDFCAF55" box="[262,537,1022,1048]" italics="true" pageId="15" pageNumber="228">Leptochilus neolongipes</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="04BE99B1C95CA51CD285FC3DFBD3AF55" authorityName="X. C. Zhang" authorityYear="2012" baseAuthorityName="Ching" baseAuthorityYear="1933" box="[951,1078,1022,1048]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="F1CA3E20C95CA51CD285FC3DFBD3AF55" box="[951,1078,1022,1048]" italics="true" pageId="15" pageNumber="228">L. longipes</emphasis>
</taxonomicName>
. However, it differs the latter in having larger habit, fronds
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long, and lobes
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long. In contrast,
<taxonomicName id="04BE99B1C95CA51CD535FBE1FB66AF71" authorityName="X. C. Zhang" authorityYear="2012" baseAuthorityName="Ching" baseAuthorityYear="1933" box="[1031,1155,1058,1084]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="F1CA3E20C95CA51CD535FBE1FB66AF71" box="[1031,1155,1058,1084]" italics="true" pageId="15" pageNumber="228">L. longipes</emphasis>
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has fronds
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long and lobes
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long. Phylogenetically, the new species was resolved as isolated within the
<taxonomicName id="04BE99B1C95CA51CD59EFB85FAD4AF2D" box="[1196,1329,1094,1120]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="ellipticus">
<emphasis id="F1CA3E20C95CA51CD59EFB85FAD4AF2D" box="[1196,1329,1094,1120]" italics="true" pageId="15" pageNumber="228">L. ellipticus</emphasis>
</taxonomicName>
clade, and its sister relationship with
<taxonomicName id="04BE99B1C95CA51CD083FBA9FD35AFC9" authority="T. Fujiwara" authorityName="T. Fujiwara" box="[433,720,1130,1157]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="228" phylum="Tracheophyta" rank="species" species="ornithopus">
<emphasis id="F1CA3E20C95CA51CD083FBA9FDADAFC9" box="[433,584,1130,1156]" italics="true" pageId="15" pageNumber="228">L. ornithopus</emphasis>
T. Fujiwara
</taxonomicName>
&amp; B.H. Quang (2023: 153154) was not well supported (
<figureCitation id="5B85FEB7C95CA51CD46AFBA8FA7FAFC9" box="[1368,1434,1130,1156]" captionStart="FIGURE 1" captionStartId="3.[136,229,1956,1978]" captionTargetBox="[153,1430,353,1922]" captionTargetId="figure-76@3.[151,1435,347,1928]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of Leptochilus based on six plastid markers (modified from Zhang et al. 2024). Orange color indicates the new species described here." figureDoi="http://doi.org/10.5281/zenodo.13217896" httpUri="https://zenodo.org/record/13217896/files/figure.png" pageId="15" pageNumber="228">Fig. 1</figureCitation>
). Notably, a specimen with a smaller habit,
<emphasis id="F1CA3E20C95CA51CD36CFB4CFC7DAFE5" box="[606,920,1166,1192]" italics="true" pageId="15" pageNumber="228">Cheng-Wei Chen Wade3697</emphasis>
(TAIF), collected from
<collectingRegion id="017A2CD0C95CA51CD597FB4CFACDAFE4" box="[1189,1320,1166,1193]" country="Vietnam" name="Khanh Hoa" pageId="15" pageNumber="228">Khanh Hoa</collectingRegion>
, potentially represents an underdeveloped individual of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,318 @@
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7.
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<emphasis id="F1CA3E20C95FA51FD196F94EFE48ADEA" bold="true" box="[164,429,1676,1703]" italics="true" pageId="12" pageNumber="225">Leptochilus multilobus</emphasis>
Liang Zhang &amp; X.M. Zhou
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,
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<emphasis id="F1CA3E20C95FA51FD3C1F94FFCA7ADEB" box="[755,834,1677,1702]" italics="true" pageId="12" pageNumber="225">sp. nov</emphasis>
.
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(
<figureCitation id="5B85FEB7C95FA51FD26BF94FFC78ADEA" box="[857,925,1677,1703]" captionStart="FIGURE 9" captionStartId="13.[136,229,1773,1795]" captionTargetBox="[220,1368,402,1735]" captionTargetId="figure-120@13.[207,1380,389,1748]" captionTargetPageId="13" captionText="FIGURE 9. Leptochilus multilobus Liang Zhang &amp; X.M. Zhou,—A. Habitat.—B. Abaxial lamina and fiddlehead.—C. Portion of abaxial lamina showing sori.—D. Rhizome.—E. Portion of adaxial lamina. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217912" httpUri="https://zenodo.org/record/13217912/files/figure.png" pageId="12" pageNumber="225">Fig. 9</figureCitation>
).
</heading>
</paragraph>
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<emphasis id="F1CA3E20C95FA51FD1BAF917FF26ADA2" bold="true" box="[136,195,1749,1775]" pageId="12" pageNumber="225">Type</emphasis>
:
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.
<collectingRegion id="017A2CD0C95FA51FD072F917FE70ADA2" box="[320,405,1749,1775]" country="China" name="Xizang" pageId="12" pageNumber="225">Xizang</collectingRegion>
: Medog County, Beibengxiang, Aniqiao to #3 bridge, elev.
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,
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/
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, on wet rocks at forest margins,
<date id="B700C4F2C95FA51FD0FFF93BFD6AAC5E" box="[461,655,1785,1811]" pageId="12" pageNumber="225" value="2017-10-17">17 October 2017</date>
,
<emphasis id="F1CA3E20C95FA51FD3AEF93BFB0CAC5E" box="[668,1257,1785,1811]" italics="true" pageId="12" pageNumber="225">Liang Zhang, Wen-Bin Ju &amp; Heng-Ning Deng 2193</emphasis>
(
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KUN-1576328!,
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KUN-1576327!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95FA51ED18FF882FCB5AA21" lastPageId="13" lastPageNumber="226" pageId="12" pageNumber="225" type="diagnosis">
<paragraph id="C301E232C95FA51FD18FF882FABAAC32" blockId="12.[136,1452,1748,2063]" pageId="12" pageNumber="225">
<emphasis id="F1CA3E20C95FA51FD18FF882FED5AC17" bold="true" box="[189,304,1856,1882]" pageId="12" pageNumber="225">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C95FA51FD00DF883FDDBAC16" box="[319,574,1857,1883]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="225" phylum="Tracheophyta" rank="species" species="multilobus">
<emphasis id="F1CA3E20C95FA51FD00DF883FDDBAC16" box="[319,574,1857,1883]" italics="true" pageId="12" pageNumber="225">Leptochilus multilobus</emphasis>
</taxonomicName>
can be distinguished from other species by its lobes up to 15 pairs, sori oblong to shortly linear, and length of sori approximately half that of the distance from the midrib to the lobe margins.
</paragraph>
<paragraph id="C301E232C95FA51ED18FF84BFCB5AA21" blockId="12.[136,1452,1748,2063]" lastBlockId="13.[136,1452,158,365]" lastPageId="13" lastPageNumber="226" pageId="12" pageNumber="225">
<emphasis id="F1CA3E20C95FA51FD18FF84BFEEDACEE" bold="true" box="[189,264,1929,1955]" pageId="12" pageNumber="225">Plants</emphasis>
perennial, evergreen.
<emphasis id="F1CA3E20C95FA51FD0CCF84AFD8DACEF" bold="true" box="[510,616,1928,1954]" pageId="12" pageNumber="225">Rhizome</emphasis>
creeping, ca.
<quantity id="04464FD7C95FA51FD232F84BFC63ACEE" box="[768,902,1929,1955]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.2" metricValueMin="2.9" pageId="12" pageNumber="225" unit="mm" value="3.55" valueMax="4.2" valueMin="2.9">2.94.2 mm</quantity>
in diam., dorsiventrally flattened, densely scaled, with many short roots; scales brown, clathrate, membranous, margins entire or sparely denticulate, 0.83.2 ×
<quantity id="04464FD7C95FA51FD465F86EFF52ACA6" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="12" pageNumber="225" unit="mm" value="1.05" valueMax="1.5" valueMin="0.6">0.61.5 mm</quantity>
.
<emphasis id="F1CA3E20C95FA51FD1F3F813FEF3ACA6" bold="true" box="[193,278,2001,2027]" pageId="12" pageNumber="225">Fronds</emphasis>
monomorphic, 0.4 to 0.6 (2.5) cm distant,
<quantity id="04464FD7C95FA51FD3C6F812FC86ACA6" box="[756,867,2000,2027]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" metricValueMax="5.2" metricValueMin="3.2" pageId="12" pageNumber="225" unit="cm" value="42.0" valueMax="52.0" valueMin="32.0">3252 cm</quantity>
long;
<emphasis id="F1CA3E20C95FA51FD298F812FBE1ACA7" bold="true" box="[938,1028,2000,2026]" pageId="12" pageNumber="225">petioles</emphasis>
stramineous,
<quantity id="04464FD7C95FA51FD5ADF813FAEBACA6" box="[1183,1294,2001,2027]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" metricValueMax="2.6" metricValueMin="1.3" pageId="12" pageNumber="225" unit="cm" value="19.5" valueMax="26.0" valueMin="13.0">1326 cm</quantity>
long, ca.
<quantity id="04464FD7C95FA51FD448F813FF3BA342" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="12" pageNumber="225" unit="mm" value="1.75" valueMax="2.1" valueMin="1.4">1.4 2.1 mm</quantity>
in diam. at middle, slightly winged at distal part; rachis narrowly winged,
<quantity id="04464FD7C95FA51FD517F836FB49A342" box="[1061,1196,2036,2063]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="12" pageNumber="225" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.12.5 mm</quantity>
wide at the distal part,
<quantity id="04464FD7C95EA51ED1BAFF5CFEEBABF4" box="[136,270,158,185]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="13" pageNumber="226" unit="mm" value="0.9" valueMax="1.1" valueMin="0.7">0.71.1 mm</quantity>
at base.
<emphasis id="F1CA3E20C95EA51ED059FF5CFE30ABF5" bold="true" box="[363,469,158,184]" pageId="13" pageNumber="226">Laminae</emphasis>
elliptic, deeply 1-pinnatifid, 2027 ×
<quantity id="04464FD7C95EA51ED244FF5CFC3CABF4" box="[886,985,158,185]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="13" pageNumber="226" unit="cm" value="10.5" valueMax="12.0" valueMin="9.0">912 cm</quantity>
, widest at middle of laminae, chartaceous, brown when dry, glabrous, margins of lobes entire.
<emphasis id="F1CA3E20C95EA51ED3F6FF00FCE9AB91" bold="true" box="[708,780,194,220]" pageId="13" pageNumber="226">Lobes</emphasis>
1215 pairs, narrowly lanceolate, apex acuminate, 6.57.0×
<quantity id="04464FD7C95EA51ED1BAFF24FEECAA4C" box="[136,265,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="13" pageNumber="226" unit="cm" value="0.8" valueMax="0.9" valueMin="0.7">0.70.9 cm</quantity>
, basal portions of lobes distinctly narrowed, ascending upward, midribs of lobes at angles of 4268(77) degree with rachis, distal pairs of lobes narrowed, mostly opposite, lobe base ca.
<quantity id="04464FD7C95EA51ED52EFEC8FB7EAA68" box="[1052,1179,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.7" metricValueMin="1.1" pageId="13" pageNumber="226" unit="cm" value="1.4" valueMax="1.7" valueMin="1.1">1.11.7 cm</quantity>
distant with each other. Midribs slightly raised, lateral veins invisible.
<emphasis id="F1CA3E20C95EA51ED3A1FEECFD20AA05" bold="true" box="[659,709,302,328]" pageId="13" pageNumber="226">Sori</emphasis>
oblong or shortly linear, (4)711 on each side of midrib, 1.84.0 ×
<quantity id="04464FD7C95EA51ED1ADFE90FECDAA20" box="[159,296,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="13" pageNumber="226" unit="mm" value="0.8" valueMax="1.1" valueMin="0.5">0.51.1 mm</quantity>
, centers ca. 36 (9) mm apart from one another.
</paragraph>
</subSubSection>
<caption id="97C1B2BAC95EA51ED1BAF92FFBFDAC6A" ID-DOI="http://doi.org/10.5281/zenodo.13217912" ID-Zenodo-Dep="13217912" httpUri="https://zenodo.org/record/13217912/files/figure.png" pageId="13" pageNumber="226" startId="13.[136,229,1773,1795]" targetBox="[220,1368,402,1735]" targetPageId="13" targetType="figure">
<paragraph id="C301E232C95EA51ED1BAF92FFBFDAC6A" blockId="13.[136,1451,1773,1831]" pageId="13" pageNumber="226">
<emphasis id="F1CA3E20C95EA51ED1BAF92FFF12AC4F" bold="true" box="[136,247,1773,1795]" pageId="13" pageNumber="226">FIGURE 9</emphasis>
.
<taxonomicName id="04BE99B1C95EA51ED031F92FFD0DAC4E" authority="Liang Zhang &amp; X. M. Zhou" authorityName="Liang Zhang &amp; X. M. Zhou" box="[259,744,1773,1795]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="13" pageNumber="226" phylum="Tracheophyta" rank="species" species="multilobus">
<emphasis id="F1CA3E20C95EA51ED031F92FFE38AC4F" box="[259,477,1773,1794]" italics="true" pageId="13" pageNumber="226">Leptochilus multilobus</emphasis>
Liang Zhang &amp; X.M. Zhou
</taxonomicName>
,—A. Habitat.—B. Abaxial lamina and fiddlehead.—C. Portion of abaxial lamina showing sori.—D. Rhizome.—E. Portion of adaxial lamina. Photo credit: Liang Zhang.
</paragraph>
</caption>
<subSubSection id="8BA4B1B9C95EA51ED18FF897FCD8AC96" pageId="13" pageNumber="226" type="materials_examined">
<paragraph id="C301E232C95EA51ED18FF897FCD8AC96" blockId="13.[136,1452,1877,2084]" pageId="13" pageNumber="226">
<emphasis id="F1CA3E20C95EA51ED18FF897FDDDAC22" bold="true" box="[189,568,1877,1903]" pageId="13" pageNumber="226">Additional specimens examined</emphasis>
:
<materialsCitation id="73D6E86FC95EA51ED37AF897FE29ACFA" ID-GBIF-Occurrence="4928466315" collectingDate="2021-06-23" collectionCode="KUN" country="China" county="Medog County" elevation="935" latitude="29.36" location="Deguo village" longLatPrecision="736" longitude="95.34" municipality="Dexingxiang" pageId="13" pageNumber="226" specimenCount="1" stateProvince="Xizang">
<collectingCountry id="BBA9A2A2C95EA51ED37AF897FD44AC22" box="[584,673,1877,1903]" name="China" pageId="13" pageNumber="226">CHINA</collectingCountry>
.
<collectingRegion id="017A2CD0C95EA51ED39DF897FCE1AC22" box="[687,772,1877,1903]" country="China" name="Xizang" pageId="13" pageNumber="226">Xizang</collectingRegion>
:
<collectingCounty id="2A609ABEC95EA51ED222F897FC5FAC22" box="[784,954,1877,1903]" pageId="13" pageNumber="226">Medog County</collectingCounty>
,
<collectingMunicipality id="23657848C95EA51ED2F4F897FBBFAC22" box="[966,1114,1877,1903]" pageId="13" pageNumber="226">Dexingxiang</collectingMunicipality>
, on the way to
<location id="C661B4E9C95EA51ED434F894FA42AC22" LSID="urn:lsid:plazi:treatment:4B175324C95FA51DD1BAF94FFCD7AA94:C661B4E9C95EA51ED434F894FA42AC22" box="[1286,1447,1877,1904]" country="China" county="Medog County" latitude="29.36" longLatPrecision="736" longitude="95.34" municipality="Dexingxiang" name="Deguo village" pageId="13" pageNumber="226" stateProvince="Xizang">Deguo village</location>
, elev.
<quantity id="04464FD7C95EA51ED1F7F8BBFEF4ACD9" box="[197,273,1913,1940]" metricMagnitude="2" metricUnit="m" metricValue="9.35" pageId="13" pageNumber="226" unit="m" value="935.0">
<elevation id="48930501C95EA51ED1F7F8BBFEF4ACD9" box="[197,273,1913,1940]" metricMagnitude="2" metricUnit="m" metricValue="9.35" pageId="13" pageNumber="226" unit="m" value="935.0">935 m</elevation>
</quantity>
,
<geoCoordinate id="A68A84F5C95EA51ED02DF8BBFE98ACD9" box="[287,381,1913,1940]" degrees="29.36" direction="north" orientation="latitude" pageId="13" pageNumber="226" precision="555" value="29.36">29.36°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C95EA51ED0B4F8BBFE05ACD9" box="[390,480,1913,1940]" degrees="95.34" direction="east" orientation="longitude" pageId="13" pageNumber="226" precision="555" value="95.34">95.34°E</geoCoordinate>
, on a wet cliff near waterfall,
<date id="B700C4F2C95EA51ED271F8BBFC05ACDE" box="[835,992,1913,1940]" pageId="13" pageNumber="226" value="2021-06-23">
<collectingDate id="A7443D1AC95EA51ED271F8BBFC05ACDE" box="[835,992,1913,1940]" pageId="13" pageNumber="226" value="2021-06-23">23 June 2021</collectingDate>
</date>
,
<emphasis id="F1CA3E20C95EA51ED2DCF8B8FE91ACFA" italics="true" pageId="13" pageNumber="226">Liang Zhang, Xia Wan, Jian-Jun Yang, Wang-Lin Zhao 4455</emphasis>
(
<collectionCode id="A5AF7AF7C95EA51ED0B0F85FFE20ACFA" box="[386,453,1949,1975]" country="China" lsid="urn:lsid:biocol.org:col:15842" name="Kunming Institute of Botany, Chinese Academy of Sciences" pageId="13" pageNumber="226" type="Herbarium">KUN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73D6E86FC95EA51ED0E5F85FFCDFAC96" ID-GBIF-Occurrence="4928466305" collectionCode="CDBI" country="China" county="Medog County" elevation="1085" latitude="29.32" location="Aniqiao" longLatPrecision="736" longitude="95.16" municipality="Beibengxiang" pageId="13" pageNumber="226" specimenCount="1">
<collectingCounty id="2A609ABEC95EA51ED0E5F85FFD64ACFA" box="[471,641,1949,1975]" pageId="13" pageNumber="226">Medog County</collectingCounty>
,
<collectingMunicipality id="23657848C95EA51ED3B9F85FFCCCACFA" box="[651,809,1949,1975]" pageId="13" pageNumber="226">Beibengxiang</collectingMunicipality>
,
<location id="C661B4E9C95EA51ED200F85FFC69ACFA" LSID="urn:lsid:plazi:treatment:4B175324C95FA51DD1BAF94FFCD7AA94:C661B4E9C95EA51ED200F85FFC69ACFA" box="[818,908,1949,1975]" country="China" county="Medog County" latitude="29.32" longLatPrecision="736" longitude="95.16" municipality="Beibengxiang" name="Aniqiao" pageId="13" pageNumber="226">Aniqiao</location>
to #3 bridge, elev.
<quantity id="04464FD7C95EA51ED56DF85FFB53ACF5" box="[1119,1206,1949,1976]" metricMagnitude="3" metricUnit="m" metricValue="1.085" pageId="13" pageNumber="226" unit="m" value="1085.0">
<elevation id="48930501C95EA51ED56DF85FFB53ACF5" box="[1119,1206,1949,1976]" metricMagnitude="3" metricUnit="m" metricValue="1.085" pageId="13" pageNumber="226" unit="m" value="1085.0">1085 m</elevation>
</quantity>
,
<geoCoordinate id="A68A84F5C95EA51ED5F3F85FFAFAACF5" box="[1217,1311,1949,1976]" degrees="29.32" direction="north" orientation="latitude" pageId="13" pageNumber="226" precision="555" value="29.32">29.32°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C95EA51ED41AF85FFA67ACF5" box="[1320,1410,1949,1976]" degrees="95.16" direction="east" orientation="longitude" pageId="13" pageNumber="226" precision="555" value="95.16">95.16°E</geoCoordinate>
, on a wet rock at forest margins,
<emphasis id="F1CA3E20C95EA51ED0FFF800FD35AC96" box="[461,720,1985,2011]" italics="true" pageId="13" pageNumber="226">Bo Xu et al. YLZB2004</emphasis>
(
<collectionCode id="A5AF7AF7C95EA51ED3D2F803FCC8AC96" box="[736,813,1985,2011]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="13" pageNumber="226" type="Herbarium">CDBI</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95EA51DD18FF827FD6EAA4D" lastPageId="14" lastPageNumber="227" pageId="13" pageNumber="226" type="distribution">
<paragraph id="C301E232C95EA51DD18FF827FD6EAA4D" blockId="13.[136,1452,1877,2084]" lastBlockId="14.[136,1452,158,473]" lastPageId="14" lastPageNumber="227" pageId="13" pageNumber="226">
<emphasis id="F1CA3E20C95EA51ED18FF827FE02ACB2" bold="true" box="[189,487,2021,2047]" pageId="13" pageNumber="226">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C95EA51ED0DFF827FD09ACB2" box="[493,748,2021,2047]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="13" pageNumber="226" phylum="Tracheophyta" rank="species" species="multilobus">
<emphasis id="F1CA3E20C95EA51ED0DFF827FD09ACB2" box="[493,748,2021,2047]" italics="true" pageId="13" pageNumber="226">Leptochilus multilobus</emphasis>
</taxonomicName>
is currently only known from Medog, a remote county in
<collectingRegion id="017A2CD0C95EA51ED458F827FA42ACB2" box="[1386,1447,2021,2047]" country="China" name="Xizang" pageId="13" pageNumber="226">Tibet</collectingRegion>
,
<collectingCountry id="BBA9A2A2C95EA51ED1BAF7CBFF2AA36E" box="[136,207,2057,2083]" name="China" pageId="13" pageNumber="226">China</collectingCountry>
. Two large populations, approximately
<quantity id="04464FD7C95EA51ED3A0F7CBFD3EA36E" box="[658,731,2057,2083]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="226" unit="km" value="20.0">20 km</quantity>
apart in an air distance, have been discovered. One population found in Beibengxiang has been destroyed due to road construction, while the other population, which has about 30 individuals, is in good condition. The species is restricted in rocky areas near streams or waterfalls, and high humidity is crucial to maintaining a healthy population.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95DA51DD18FFEC8FCD7AA94" pageId="14" pageNumber="227" type="etymology">
<paragraph id="C301E232C95DA51DD18FFEC8FCD4AA05" blockId="14.[136,1452,158,473]" pageId="14" pageNumber="227">
<emphasis id="F1CA3E20C95DA51DD18FFEC8FEDBAA69" bold="true" box="[189,318,266,292]" pageId="14" pageNumber="227">Etymology</emphasis>
: The species epithet is derived from the Latin words
<emphasis id="F1CA3E20C95DA51DD2A8FEC8FC38AA69" box="[922,989,266,292]" italics="true" pageId="14" pageNumber="227">multi-</emphasis>
, multiple, and
<emphasis id="F1CA3E20C95DA51DD5BAFEC8FB2BAA69" box="[1160,1230,266,292]" italics="true" pageId="14" pageNumber="227">-lobus</emphasis>
, lobes. This epithet refers to the numerous pairs of lobes present on each lamina.
</paragraph>
<paragraph id="C301E232C95DA51DD18FFE90FCD7AA94" blockId="14.[136,1452,158,473]" pageId="14" pageNumber="227">
<emphasis id="F1CA3E20C95DA51DD18FFE90FF10AA21" bold="true" box="[189,245,338,364]" pageId="14" pageNumber="227">Note</emphasis>
:
<taxonomicName id="04BE99B1C95DA51DD037FE90FDE0AA21" box="[261,517,338,364]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="14" pageNumber="227" phylum="Tracheophyta" rank="species" species="multilobus">
<emphasis id="F1CA3E20C95DA51DD037FE90FDE0AA21" box="[261,517,338,364]" italics="true" pageId="14" pageNumber="227">Leptochilus multilobus</emphasis>
</taxonomicName>
resembles species in the
<taxonomicName id="04BE99B1C95DA51DD216FE91FC4CAA21" box="[804,937,338,364]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="14" pageNumber="227" phylum="Tracheophyta" rank="species" species="ellipticus">
<emphasis id="F1CA3E20C95DA51DD216FE91FC4CAA21" box="[804,937,338,364]" italics="true" pageId="14" pageNumber="227">L. ellipticus</emphasis>
</taxonomicName>
clade by having pinnatifid laminae (
<bibRefCitation id="A72F9FC3C95DA51DD474FE90FF0DAADD" author="Zhang, L. &amp; Lu, N. T. &amp; Zhou, X. - M. &amp; Chen, D. - K. &amp; Knapp, R. &amp; Zhou, L. &amp; Guo, L. &amp; Luong, T. T. &amp; Sun, H. &amp; Gao, X. - F. &amp; Zhang, L. - B." pageId="14" pageNumber="227" pagination="311 - 322" refId="ref12520" refString="Zhang, L., Lu, N. T., Zhou, X. - M., Chen, D. - K., Knapp, R., Zhou, L., Guo, L., Luong, T. T., Sun, H., Gao, X. - F. &amp; Zhang, L. - B. (2019) A plastid phylogeny of the Old World fern genus Leptochilus (Polypodiaceae): Implications for cryptic speciation and progressive colonization from lower to higher latitudes. Molecular Phylogenetics and Evolution 134: 311 - 322. https: // doi. org / 10.1016 / j. ympev. 2019.01.013" type="journal article" year="2019">
Zhang
<emphasis id="F1CA3E20C95DA51DD4A5FE91FF40AADD" italics="true" pageId="14" pageNumber="227">et al.</emphasis>
2019
</bibRefCitation>
), while differs them in having a great number of lobes and oblong to shortly linear sori. Phylogenetically,
<taxonomicName id="04BE99B1C95DA51DD4A7FEB5FF1BAAF9" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="14" pageNumber="227" phylum="Tracheophyta" rank="species" species="multilobus">
<emphasis id="F1CA3E20C95DA51DD4A7FEB5FF1BAAF9" italics="true" pageId="14" pageNumber="227">L. multilobus</emphasis>
</taxonomicName>
was resolved as a member in the
<taxonomicName id="04BE99B1C95DA51DD345FE58FD2CAAF9" authorityName="K.B.Presl" authorityYear="1851" box="[631,713,410,436]" class="Polypodiopsida" family="Polypodiaceae" genus="Colysis" kingdom="Plantae" order="Polypodiales" pageId="14" pageNumber="227" phylum="Tracheophyta" rank="genus">
<emphasis id="F1CA3E20C95DA51DD345FE58FD2CAAF9" box="[631,713,410,436]" italics="true" pageId="14" pageNumber="227">Colysis</emphasis>
</taxonomicName>
clade (
<figureCitation id="5B85FEB7C95DA51DD229FE58FCB9AAF9" box="[795,860,410,436]" captionStart="FIGURE 1" captionStartId="3.[136,229,1956,1978]" captionTargetBox="[153,1430,353,1922]" captionTargetId="figure-76@3.[151,1435,347,1928]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of Leptochilus based on six plastid markers (modified from Zhang et al. 2024). Orange color indicates the new species described here." figureDoi="http://doi.org/10.5281/zenodo.13217896" httpUri="https://zenodo.org/record/13217896/files/figure.png" pageId="14" pageNumber="227">Fig. 1</figureCitation>
), which is composed of those with simple and entire laminae, making
<taxonomicName id="04BE99B1C95DA51DD078FE7DFE38AA95" box="[330,477,446,472]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="14" pageNumber="227" phylum="Tracheophyta" rank="species" species="multilobus">
<emphasis id="F1CA3E20C95DA51DD078FE7DFE38AA95" box="[330,477,446,472]" italics="true" pageId="14" pageNumber="227">L. multilobus</emphasis>
</taxonomicName>
a unique species in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>