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<document id="56985A3AA971966736E8C66FE0770CEE" ID-CLB-Dataset="300292" ID-DOI="10.3897/zookeys.1208.122126" ID-GBIF-Dataset="92f37e90-7f3a-42f4-987d-aeae1889d378" ID-ZooBank="56CC7AA9-4B0A-4EA0-86A5-B066E6A57FC3" ID-publisher-id="122126" URI-arpha="994AC1D6-2908-5674-A0CD-B8A92786C6CB" URI-zoobank="http://zoobank.org/56CC7AA9-4B0A-4EA0-86A5-B066E6A57FC3" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1721924801512" checkinUser="pensoft" docAuthor="Doménech, Carles" docDate="2024" docId="C03A6625891E551FB07136160948D577" docLanguage="en" docName="ZooKeys 1208: 49-80" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/122126/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Scolopendra oraniensis Lucas 1846" docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="80" masterDocId="994AC1D629085674A0CDB8A92786C6CB" masterDocTitle="Taxonomic and nomenclatural reassessment of the Iberian Peninsula’s nomina obscura, Scolopendra viridipes Dufour, 1820 and S. chlorotes L. Koch in Rosenhauer, 1856 (Chilopoda, Scolopendromorpha, Scolopendridae)" masterLastPageNumber="80" masterPageNumber="49" pageNumber="49" updateTime="1721938564851" updateUser="ExternalLinkService">
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<mods:title id="39A8647E0C737562FB8F67A64223F8B0">Taxonomic and nomenclatural reassessment of the Iberian Peninsula’s nomina obscura, Scolopendra viridipes Dufour, 1820 and S. chlorotes L. Koch in Rosenhauer, 1856 (Chilopoda, Scolopendromorpha, Scolopendridae)</mods:title>
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<mods:namePart id="072E1D92DFB377BD6B47D54C1DA455F5">Doménech, Carles</mods:namePart>
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<mods:affiliation id="F6642E457BCC0E645F71DA9031223275">Departament de Ciències Ambientals i Recursos Naturals, Universitat d’Alacant, Carretera de Sant Vicent del Raspeig s / n C. P. 03690, San Vicent del Raspeig, Alacant, Spain</mods:affiliation>
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<taxonomicName id="5F04CE0B82FDC0809953D5AF0C3A7F8F" ID-CoL="79XG2" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="E6F8CEB258B249A98039F58070D3CE21" italics="true">Scolopendra oraniensis</emphasis>
<bibRefCitation id="2D2045CF67B2CFF036BF86CBC5C00E14" author="Lucas PH" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du Nord de l'Afrique. Revue Zoologique 9: 283 - 289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas, 1846</bibRefCitation>
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<figureCitation id="290EF24CEF6D86EEBCF2E5D3B6305291" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">Types: Figs 7</figureCitation>
,
<figureCitation id="7CE2DC3F5F353554F67B2E619FCD6E4F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8</figureCitation>
;
<tableCitation id="6E754945672560F2D45E3945D6846044" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">Table 3</tableCitation>
.
<figureCitation id="81DCA04594E75865C12F3D06EE093C17" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Representatives of Scolopendromorpha from the Iberian Peninsula (uncollected) A family Plutoniumidae Bollman, 1893: Theatops erytrocephalus (C. L. Koch, 1847), Seoane do Caurel, Lugo (credit: J. Tizón) B family Cryptopidae Kohlrausch, 1881: Cryptops sp. Leach, 1814, Villena, Alacant (credit: D. Molina) C – I family Scolopendridae Leach, 1814 C Scolopendra cingulata Latreille, 1829, Benilloba, Alacant (credit: M. Huesca and CD) and D – I S. oraniensis Lucas, 1840. Observe that in this species the colour variants can occur sympatrically D, H, I Benilloba, Alacant and E – G Almeria Province, Andalucia (credit: F. Rodriguez Luque)." figureDoi="10.3897/zookeys.1208.122126.figure1" httpUri="https://binary.pensoft.net/fig/1101811">Non-types: Figs 1 C – I</figureCitation>
,
<figureCitation id="ECA555FA4CFA012DEBA0C64B755BF88D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A topotype collection localities in the Iberian Peninsula (southwestern Europe): S. viridipes topotypes (= S. oraniensis); V 1 = Moixent and V 2 = Xàtiva (previously San Felipe), Valencia province, Valencian Country / Community, Spain and S. chlorotes topotypes (= S. oraniensis); C 1 = Málaga municipality, C 2 = Alahurín de la Torre, C 3 = Casabermeja, C 4 = Totalán and C 5 = Estepona, Málaga province, Andalucía, Spain. The “ O ” points S. oraniensis type localities, O 1 = Santon’s mountains [corrected from “ Sauton ” (sic.); Lucas 1846] and O 2 = “ Between Oran and Mers-el-Kebir ”, Oran province (Wilayah), Algeria B “ S. viridipes ” topotype 2, female, (CEUAMr 22) from C Moixent, Alacant D “ S. chlorotes ” topotype 6, female, (CEUAMr 30) from E Alahurín de la Torre, Málaga." figureDoi="10.3897/zookeys.1208.122126.figure2" httpUri="https://binary.pensoft.net/fig/1101812">2 B, D</figureCitation>
,
<figureCitation id="052D1640411556F1924281FBEA2C8F75" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A cephalic plate, antennae, and TT 1–5 B forcipular segment and sternites 1–3 C tooth plates D tergites 13–18 E sternites 10–15. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure3" httpUri="https://binary.pensoft.net/fig/1101813">3</figureCitation>
,
<figureCitation id="4D084625B110C52921C5D808048534A7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A ULBS and UL ’ s dorsal view B ULBS ’ s and UL ’ s ventral view C ULBS ’ s, coxopleuron, and UL ’ s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure4" httpUri="https://binary.pensoft.net/fig/1101814">4</figureCitation>
,
<figureCitation id="A3B353F01B0CDDBE619F93C28EA54A5B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A cephalic plate, antennae, and TT 1–3 B forcipular segment and sternites 1, 2 C tooth plates D tergites 10–14 E sternites 8–12. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure5" httpUri="https://binary.pensoft.net/fig/1101815">5</figureCitation>
,
<figureCitation id="DDC3EEAE92D1CD28AF178DA862E404E2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A ULBS ’ s and UL ’ s dorsal view B ULBS ’ s and UL ’ s ventral view C ULBS ’ s, coxopleuron, and UL ’ s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure6" httpUri="https://binary.pensoft.net/fig/1101816">6</figureCitation>
,
<figureCitation id="9577DE4107D55349CE74D010E59BAF1E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7</figureCitation>
,
<figureCitation id="FB405B6E81F3409802DC0F920619AB18" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8</figureCitation>
,
<figureCitation id="EC563025DFC642C42ACA3C40A7D884F3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9</figureCitation>
;
<tableCitation id="DB57E8F2FA7A03227AE8ACC7F838A887" captionStart="Table 1" captionStartId="T1" captionText="Table 1. The original descriptive morphology of Scolopendra viridipes in comparison to other Scolopendromorpha species described in Peninsular Spain. Bold letters indicate compatible features in other species. * indicates common to the entire Scolopendromorpha, interpretable, or poorly differentiated feature. Nr pp = Not reported, presence possible." httpUri="http://table.plazi.org/id/0A770E22511DFC4FD10D48EC210F5DEA" tableUuid="0A770E22511DFC4FD10D48EC210F5DEA">Tables 1</tableCitation>
,
<tableCitation id="814696447C56652BF0B6C85AE9792BBE" captionStart="Table 2" captionStartId="T2" captionText="Table 2. The original descriptive morphology of Scolopendra chlorotes in comparison to other Scolopendromorpha species described in Peninsular Spain. Bold letters indicate compatible features in other species. * indicates common to the entire Scolopendromorpha, interpretable, or poorly differentiated feature. Nr pp = Not reported, presence possible. NA = Information not available." httpUri="http://table.plazi.org/id/83E668CADC992295344F956E110DB611" tableUuid="83E668CADC992295344F956E110DB611">2</tableCitation>
,
<tableCitation id="954DE872A9FFD63368B34E02AF495670" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
<xref id="077C1ED845C2061B5F163BB405B3D81D" ref-type="supplementary-material" rid="S1">; Suppl. material 1: files 1, 3</xref>
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<taxonomicName id="27255D32FAB818116899E5BBEFE812B4" ID-CoL="4VSD2" authority="Dufour, 1820" authorityName="Dufour" authorityYear="1820" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="viridipes">
<emphasis id="71B9AC5E1C0BD1CC7F3F60EFA175105E" italics="true">Scolopendra viridipes</emphasis>
<bibRefCitation id="1D34D980BC90B1990E743714D0702D18" author="Dufour JML" journalOrPublisher="Annales générales des sciences physiques" pagination="307-317" refId="B20" refString="Dufour JML (1820) Description de dix especes nouvelles ou peu connues d'Insectes recueillis en Espagne. Annales generales des sciences physiques 6: 307 - 317." title="Description de dix espèces nouvelles ou peu connues d’Insectes recueillis en Espagne." volume="6" year="1820">Dufour, 1820</bibRefCitation>
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; nomen oblitum; senior syn. nov.
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<taxonomicName id="5AD2BE242D9B2CD118013B377D181470" ID-CoL="79XG2" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="9653E285D5E415CD927395E21798F1D2" italics="true">S. oraniensis</emphasis>
<bibRefCitation id="0FC5128B64F0FA3FD0888B967822B984" author="Lucas PH" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du Nord de l'Afrique. Revue Zoologique 9: 283 - 289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas, 1846</bibRefCitation>
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; nomen protectum.
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<taxonomicName id="1C66D437D58550BF686E40264604C4BC" ID-CoL="4VS6H" authority="L. Koch" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="927EC078113866C3359794454131F815" italics="true">S. chlorotes</emphasis>
L. Koch
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in
<bibRefCitation id="62988CA33BFE8B569005B767A8A33281" DOI="10.5962/bhl.title.66016" author="Rosenhauer WG" journalOrPublisher="Verlag von Theodor Blaesing, Erlangen" refId="B65" refString="Rosenhauer WG (1856) Die Thiere Andalusiens nach dem resulte einer Reise. Ed. Verlag von Theodor Blaesing, Erlangen, 429 pp. https: // doi. org / 10.5962 / bhl. title. 66016" title="Die Thiere Andalusiens nach dem resulte einer Reise. Ed." year="1856">Rosenhauer, 1856</bibRefCitation>
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; junior syn. nov.
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</paragraph>
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<taxonomicName id="231AEB2AAC226C372B7C7C5836565032" ID-CoL="4VS9J" authority="Verhoeff, 1893: 318" authorityName="Verhoeff" authorityPageNumber="318" authorityYear="1893" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="mediterranea">
<emphasis id="B31D1A9598F2CEF6347CE7FC3B8A2A45" italics="true">S. mediterranea</emphasis>
Verhoeff, 1893: 318
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.
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<taxonomicName id="774E3695F13ADFCAE607EDB0850FB491" ID-CoL="5L3YJ" authority="Verhoeff, 1893: 318" authorityName="Verhoeff" authorityPageNumber="318" authorityYear="1893" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="subSpecies" species="mediterranea" subSpecies="lusitanica">
<emphasis id="E7F2A1FB0B0B3ED5417A06C73C9FF2E4" italics="true">S. mediterranea lusitanica</emphasis>
Verhoeff, 1893: 318
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.
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<paragraph id="47D7F0635DE6ADA857D139F4AFBC6A84">
<treatmentCitationGroup id="A99C10DE6B07AEEFF17009237938D741">
<treatmentCitation id="F1F6D648C385D884986A2CA107E05B46">
<taxonomicName id="D769C67C11C4F8BF023DEA3C45BC8AF9" ID-CoL="4VS6L" authority="Silvestri, 1897: 7" authorityName="Silvestri" authorityPageNumber="7" authorityYear="1897" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="9B88EF523F6C19F5819A04FCD04D473E" italics="true">S. clavipes</emphasis>
Silvestri, 1897: 7
</taxonomicName>
(sic.).
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8828A8211682938F425DE825F9F22561">
<treatmentCitationGroup id="1430A8D5ED2A4EDBBFEBC17D48368A49">
<treatmentCitation id="AC920E3EE121BAE7B15FBFFF4F56E645">
<taxonomicName id="98CAB925604DD2E89A575A9DC0BA8AE0" ID-CoL="5L3YD" authority="Attems, 1902: 556" authorityName="Attems" authorityPageNumber="556" authorityYear="1902" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="subSpecies" species="oraniensis" subSpecies="africana">
<emphasis id="7055F57294A9ABDFABE6C012D36B45EA" italics="true">S. oraniensis africana</emphasis>
Attems, 1902: 556
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="961F73E1B14A9500F91623E75E34F8BE">
<treatmentCitationGroup id="9FD42A9721742C963D8B072B9FCF39BB">
<treatmentCitation id="8794303BDE350E0269717C8F2AB005E6">
<taxonomicName id="3642073CE316BBA25E102E9958D1412A" authorityName="Attems" authorityYear="1926" class="Chilopoda" family="Scolopendridae" genus="Rhadinoscytalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="subSpecies" species="canidens" subSpecies="oraniensis">
<emphasis id="C07B7513A362D4DDEDA02D075B5C7227" italics="true">Rhadinoscytalis canidens oraniensis</emphasis>
</taxonomicName>
: Attems 1926: 246.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="7E8B112BDD509B8702157960180813F8">
<treatmentCitationGroup id="422A660DEF1B90429D16041B4539E2F5">
<taxonomicName id="B19A8BAB34E5FAD90A7F216E7AB47739" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Rhadinoscytalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="subSpecies" species="canidens" subSpecies="oraniensis">
<emphasis id="58B231A0F590B49068B0F64ABA02F98B" italics="true">S. canidens oraniensis</emphasis>
</taxonomicName>
<comment id="CCDDA5B384E6E3E3CBE0BB3B0F81CE0F">
:
<treatmentCitation id="AABD21E8E199648B75FEC3BB4F7A159A" author="Attems C" page="19" year="1930">
<bibRefCitation id="AE9756BB3437AD4FFD798F73A8AF370F" DOI="10.1515/9783112373002" author="Attems C" firstAuthor="Attems" journalOrPublisher="De Gruyter, Berlin and Leipzig 54" page="19" refId="B1" refString="Attems C (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich. W. De Gruyter, Berlin and Leipzig 54, 308 pp. https://doi.org/10.1515/9783112373002" title="Myriopoda. 2. Scolopendromorpha. Das Tierreich. W." year="1930">Attems 1930: 19</bibRefCitation>
</treatmentCitation>
, 36, 37, fig. 50.
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="345C63332D5483FF9595EBFF2267B5EB">
<treatmentCitationGroup id="C5D4D030D207157EA0117D67845C2693">
<treatmentCitation id="2720CAD1DE9BA0F3AEB46B59145C3741" author="Verhoeff KW" page="309" year="1931">
<taxonomicName id="A5983BFE03D781FFA0E6103931FD8A54" authority="Verhoeff, 1931: 309" authorityName="Verhoeff" authorityPageNumber="309" authorityYear="1931" class="Chilopoda" family="Scolopendridae" genus="Rhadinoscytalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="subSpecies" species="canidens" subSpecies="lusitanica">
<emphasis id="697ABDFB0DA400FE40998218053BBCDA" italics="true">S. canidens lusitanica</emphasis>
<comment id="E48CC06F560C29A536D102A2B581E798">
<bibRefCitation id="90421D648E36B9771B411766CB430331" author="Verhoeff KW" journalOrPublisher="Abteilung fur Systematik, Ökologie und Geographie der Tiere" pagination="263-288" refId="B76" refString="Verhoeff KW (1931) Uber europaische Cryptops - Arten. Zoologische Jahrbucher. Abteilung fur Systematik, Okologie und Geographie der Tiere 62: 263 - 288." title="Über europäische Cryptops - Arten. Zoologische Jahrbucher." volume="62" year="1931">Verhoeff, 1931: 309</bibRefCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8CFFA8C436B38FE0450F12B87328E80B">
<treatmentCitationGroup id="097183D578AA4DAE6B43D633D81B3E1F">
<taxonomicName id="E152E6EBCD9121F7BF5A15FBF693B30F" ID-CoL="79XG2" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="B894587EA83256353BEF327E3128F22C" italics="true">S. oraniensis</emphasis>
</taxonomicName>
<comment id="B6604755D633085D6E8C698B4B3ED953">
:
<treatmentCitation id="30E86AFAE06030A15191C1A0AF8858EF" author="Würmli M" page="348 - 350" year="1980">
<bibRefCitation id="4B638BE6FE1076AF3F04D832EC65626A" author="Würmli M" firstAuthor="Wurmli" journalOrPublisher="Sitzungsberichte der Österreichischen Akademie der Wissenschaften" page="348 - 350" pagination="315-353" refId="B80" refString="Würmli M (1980) Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha). Sitzungsberichte der Österreichischen Akademie der Wissenschaften 189: 315–353." title="Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha)." volume="189" year="1980">Würmli 1980: 348–350</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVZDI" type="Morphological diagnosis">
<paragraph id="1D90D4FFC58D24C68A26CFE910385EB8">
<heading id="D3B901BC1921FEB641D3E376743D29D9" reason="title">Morphological diagnosis</heading>
</paragraph>
<paragraph id="1F61D928CDE7BE89D35A0BD05F8C3BE9">
[
<emphasis id="58B6665F96EB1E9FDFC9B03A20B62A76" bold="true">
based on
<taxonomicName id="7049BDDFECA97F84776809C3F277D41B" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="01CEAEF10C4C299F986F6F2EBE3F4630" bold="true" italics="true">S. oraniensis</emphasis>
</taxonomicName>
type series].
</emphasis>
Body length up to
<quantity id="993A65B6AF7D9B58AE0AA0F96405C32F" metricMagnitude="-2" metricUnit="m" metricValue="6.4" unit="mm" value="64.0">64 mm</quantity>
. 18 or 19 antennal articles; 5 ½ basal ones glabrous in their entire surface. Antennae / cephalic plate length ratio ≈ 3.30. Forcipular trochanteroprefemoral process clearly defined, with two or three rather inconspicuous denticles. Tooth plate with 4 + 4 (rarely 3 + 3) teeth, divided into two groups. T 1 without sutures or sulci. Paramedian sutures complete on tergites TT 2–20. T 21 with a complete median longitudinal suture. Margination starting at T 14, complete on TT 19–21. Sternite paramedian sutures on TT 2–20. Coxopleuron basally pore-field, with a single medio-distal spine. Coxopleural process with a small pore-field, sub-cylindrical and quite elongated, with 9–14 spines altogether, disposed in sub / apical, dorsal, or lateral positions. Legs 3–20 with few setae. Two tarsal spurs on legs 1; a single tarsal spur on legs 2–18 or 19. Ultimate leg elongated, sometimes with a sinusoid transverse sulcus on the ventro-distal part; prefemoral spines (usually between 18–28) arranged in five frequently miss-aligned or duplicated rows with the
<abbrev id="ABBRID0EJ1DI" xlink_title="ventro-lateral spines">VL</abbrev>
: 1-5, V: 2-6,
<abbrev id="ABBRID0EN1DI" xlink_title="ventro-median spines">VM</abbrev>
: 2-6, M: 4-8 and
<abbrev id="ABBRID0ER1DI" xlink_title="dorso-median spines">DM</abbrev>
: 2-4 formula. Prefemoral process inconspicuous ending with two or three (rarely four) spines. Prefemur and femur of
<abbrev id="ABBRID0EV1DI" xlink_title="ultimate legs">UL</abbrev>
glabrous; tibia distally covered by sparse setae; tarsi 1 and 2 covered by small setae.
<abbrev id="ABBRID0EZ1DI" xlink_title="ultimate legs">UL</abbrev>
/ T 21 length ratio ≈ 5.15.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E41DI" type="etymology">
<paragraph id="C2A5B1AA9276EF4F96BAE8FC258DEC9E">
<heading id="46085FFB8C30784C2F79B5662CFAF9B5" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="1ECE8DC54EB82A55A7F4B418090B9C93">
from the toponym “
<collectingCountry id="2A724F0FC83A678AEAACBB844A4CF99C" name="Algeria">Oran</collectingCountry>
”,
<collectingCountry id="FFBC5D881AC18FB2DB1372A2A9C21FD0" name="Algeria">Algeria</collectingCountry>
and the feminine (or masculine) suffix -
<emphasis id="DAB15B39B0DA567C200D1F3915C68F26" italics="true">ensis</emphasis>
(from) meaning “ from
<collectingCountry id="F0589B50D3136C714A30B58DA5EA485D" name="Algeria">Oran</collectingCountry>
”,
<collectingCountry id="7226885B44BD6D1FB648D06E45D7DB04" name="Algeria">Algeria</collectingCountry>
, in reference to the type locality.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF2DI" type="Type series and type depository">
<paragraph id="7E0185FB7A551691239BC8F2BF400BAF">
<heading id="971535C2F6AC6C48927538C8B0AEBB05" reason="title">Type series and type depository.</heading>
</paragraph>
<paragraph id="DE32EDAF4809D40356D7FCEA1A0C935A">
<bibRefCitation id="29F15A10651FA83432F70BE88702E748" author="Lucas PH" firstAuthor="Lucas" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique. Revue Zoologique 9: 283–289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas (1846)</bibRefCitation>
did not declare on which specimens he based the nominal taxon of
<taxonomicName id="E9FAC4188F641D8F8897C99CFA41A972" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="091C63CA32BC006C8F2F5F88FFA2FA62" italics="true">S. oraniensis</emphasis>
</taxonomicName>
(type series) (
<bibRefCitation id="16E31E0C3AE2C172209717119B41A7C7" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 72.4. 1), neither exposing the specimens’ depository nor designating a
<typeStatus id="3F261740635E9F1A1C7A65B82E75D0C9" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="1481FC930DE59B8B3F2191EC7F98006A" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 72.1. 1). Otherwise,
<bibRefCitation id="0B0D66E798FC487F72A92E409AEB2ADA" author="Kraepelin K" firstAuthor="Kraepelin" issue="10" journalOrPublisher="Bulletin du Muséum National d’Histoire Naturelle" pagination="316-324" refId="B41" refString="Kraepelin K (1904) Catalogue des scolopendrides des collections du muséum d’histroire naturelle de Paris. Bulletin du Muséum National d’Histoire Naturelle 26 (10): 316–324." title="Catalogue des scolopendrides des collections du muséum d’histroire naturelle de Paris." volume="26" year="1904">Kraepelin (1904)</bibRefCitation>
indicated the presence of the types [
<typeStatus id="4D00F4B92B7572D4C595377081AEBB68" type="syntype">syntypes</typeStatus>
] at the
<named-content id="73C64AF92883F14EF84F63A2902FEE7F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire Naturelle, Paris">
<collectionCode id="5996882E65353EA50914D16629FAAFCE" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
in writing “ – [
<taxonomicName id="19494000E82A927CE6F7AD2971C75F63" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="genus">
<emphasis id="52C1F3F3D8CB77F65A7F3FAC8EA022F6" italics="true">Scolopendra</emphasis>
</taxonomicName>
]
<taxonomicName id="ED64DD64EF16791DDB4E66FFAAFCFCA1" authority="Lucas." authorityName="Lucas." class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="FD9EF7F3A4DB7E55F9C6E490CF444611" italics="true">oraniensis</emphasis>
Lucas.
</taxonomicName>
–
<collectingCountry id="34AD3F5CAEB0AD72F7549774CB263BDB" name="Algeria">Algérie</collectingCountry>
: envir. d’Oran [surroundings of
<collectingCountry id="4804F89568B4135095A9A76C0C10C275" name="Algeria">Oran</collectingCountry>
] (H. Lucas, 1849 [sic.]. Types) ”, without further data. Therefore, after the examination of the
<named-content id="7B02EDDB484242F8461A96EE0051FDFC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire Naturelle, Paris">
<collectionCode id="9D8A99C0D566D1DB4A5A60ACE9B1FD07" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
specimens and their labels (Figs
<figureCitation id="BEA475E1D366E90096B24DA11137A706" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7</figureCitation>
,
<figureCitation id="CCAC40C64D0F76CE7A78CBA77FE24708" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8</figureCitation>
) (
<bibRefCitation id="EC651AA2B937E7BEBB7912C2D1A321C1" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 72.4. 2), I conclude that the type series for the
<taxonomicName id="C4EC77FE26937BE989107192DD5C64F7" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="3428C8DD017EB3526B689EA2DFF936B2" italics="true">S. oraniensis</emphasis>
</taxonomicName>
nominal taxon was erected on the basis of
<specimenCount id="CFA3D372AF3F8344D0FD603377127428" count="7" type="generic" typeStatus="syntype">seven syntypes</specimenCount>
, all of them unsexed adults (Table
<tableCitation id="B592A2FC19C458684D61078866AA5014" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
). Depository
<named-content id="11F09595CA07640100D617427DF3FBAC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire Naturelle, Paris">
<collectionCode id="452782EF9768BD400FDB9EC1A1FFE59F" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
, Paris,
<collectingCountry id="5516F0D86C5933344CF16D6036EC5356" name="France">France</collectingCountry>
. Jar Number 282. Samples (Figs
<figureCitation id="EE4B34B071F6041835EF09FEA0FD26CE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7</figureCitation>
,
<figureCitation id="14AAB66623906EB82F1B3D41FEB022D8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 A – C</figureCitation>
) and labels (Fig.
<figureCitation id="58DE6B415462B537618C125AA148A480" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 D</figureCitation>
) separated in three different assay tubes (containing 2 / 2 /
<specimenCount id="FE034EC01739033BA8A589AD34434CB4" count="3" type="generic">3 specimens</specimenCount>
, respectively).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK5DI" type="Collector and collection date">
<paragraph id="CCDDDE080256D4A35148B3124E4E41E4">
<heading id="6C08D376D1DC0FF5EAC47D52A23375AE" reason="title">Collector and collection date.</heading>
</paragraph>
<paragraph id="43357DA07EE8B6D63C3644ABEE6CCBD9">
P. H. Lucas, during winter, between 1839 and 1842 (
<bibRefCitation id="B22A2CBCD4AD4209A35128A30EF62D63" author="Lucas" firstAuthor="Lucas" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique. Revue Zoologique 9: 283–289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas 1846</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU5DI" type="type locality">
<paragraph id="59825A2A4BF9202A6377ED09AEBC2E91">
<heading id="8209300D68FCC47AC47BEFD67FA02C65" reason="title">Type locality and habitat</heading>
</paragraph>
<paragraph id="661269279B3E22847A384A10D3E5E5FB">
from
<bibRefCitation id="33BF1A00819BEF3D37177A6397AD5CEC" author="Lucas" firstAuthor="Lucas" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique. Revue Zoologique 9: 283–289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas 1846</bibRefCitation>
: Ravines of the “ Sauton’s ” (sic.) [Santon] mountains and ravines between
<collectingCountry id="AFA5824B38B9AA49FD525F036A7ED25B" name="Algeria">Oran</collectingCountry>
and Mers-el-Kebir,
<collectingCountry id="ECE72AB997B72D89BF83F6549E98D4B4" name="Algeria">Oran</collectingCountry>
wilayah;
<collectingCountry id="5289FD93629A9D8A70EB8802E15183CE" name="Algeria">Algeria</collectingCountry>
(
<bibRefCitation id="0D62559663ED29842FE38EF393785083" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Recommendation 76 A. 2). Lapidicolous.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC6DI" type="distribution">
<paragraph id="0DEDD4E54E117872E383D38FE2469CEA">
<heading id="AE6F34AA20E983E7A127BD51792CE6B3" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="8DF8AB44EAE1453F1F238E2305F9776B">
Known from southern
<collectingCountry id="BE9D8F61C270322B5914DAD85175343C" name="France">France</collectingCountry>
(including
<collectingCountry id="251906F4BEF86ED40DBC1F31F27B2D16" name="France">Corsica</collectingCountry>
), southern
<collectingCountry id="B842CC98A8C55B5C2DC94C73ACF59B89" name="Italy">Italy</collectingCountry>
(including
<collectingRegion id="7151F642B26B0CF0142A302899F2A662" country="Italy" name="Sardegna">Sardinia</collectingRegion>
and
<collectingRegion id="04C37F794BEF1AC9DF146FA41E884EF2" country="Italy" name="Sicilia">Sicilia</collectingRegion>
),
<collectingCountry id="A672E9B86F2CEA88B1F8C50418BCCCAB" name="Malta">Malta</collectingCountry>
,
<collectingCountry id="A271C65EF4D09622FE83AF5D6B122495" name="Spain">Spain</collectingCountry>
(including
<collectingRegion id="0D1084FED328C0012B22264CB48ED7BC" country="Spain" name="Illes Baleares">Balearic Islands</collectingRegion>
),
<collectingCountry id="7781012E96D4B92810A825F56927D89B" name="Portugal">Portugal</collectingCountry>
,
<collectingCountry id="D4787B6BA94F348833EA029E2CF60B62" name="Morocco">Morocco</collectingCountry>
, and
<collectingCountry id="FBB5C5C85F2A84314168587A0BBD4DAC" name="Algeria">Algeria</collectingCountry>
. Introduced in
<collectingCountry id="CBCD94754A52E5A1AA976EDCD6ED0B0A" name="Japan">Japan</collectingCountry>
(
<bibRefCitation id="03197BD4C44D5B9A2F9C402A7C4BC8B3" DOI="10.1515/9783112373002" author="Attems" firstAuthor="Attems" journalOrPublisher="De Gruyter, Berlin and Leipzig 54" refId="B1" refString="Attems C (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich. W. De Gruyter, Berlin and Leipzig 54, 308 pp. https://doi.org/10.1515/9783112373002" title="Myriopoda. 2. Scolopendromorpha. Das Tierreich. W." year="1930">Attems 1930</bibRefCitation>
;
<bibRefCitation id="95A233C6EE2104E9F3EE86ACDDC758B0" author="Wurmli" firstAuthor="Wurmli" journalOrPublisher="Sitzungsberichte der Österreichischen Akademie der Wissenschaften" pagination="315-353" refId="B80" refString="Würmli M (1980) Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha). Sitzungsberichte der Österreichischen Akademie der Wissenschaften 189: 315–353." title="Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha)." volume="189" year="1980">Würmli 1980</bibRefCitation>
;
<bibRefCitation id="8D118501F1F2F7B3DD4BE28277A993EE" author="Bonato L &amp; Chagas A Junior &amp; Edgecombe GD &amp; Lewis JGE &amp; Minelli A &amp; Pereira LA &amp; Shelley RM &amp; Stoev P &amp; Zapparoli M" refId="B5" refString="Bonato L, Chagas A Junior, Edgecombe GD, Lewis JGE, Minelli A, Pereira LA, Shelley RM, Stoev P, Zapparoli M (2017 a) ChiloBase 2.0. A World Catalogue of Centipedes (Chilopoda). http://chilobase.biologia.unipd.it [Accessed 10 Jan. 2020]" url="http://chilobase.biologia.unipd.it">Bonato et al. 2017 a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU6DI" type="Proposed new nomenclatural status">
<paragraph id="0F4375CD5FDFAC5DCBE4C962DB0FB048">
<heading id="52AF96C7B9B3E77B221C49E12C12195E" reason="title">Proposed new nomenclatural status.</heading>
</paragraph>
<paragraph id="4A181021D51B5B0C7072EE54475B6963">Nomen protectum.</paragraph>
</subSubSection>
<subSubSection id="SECID0EZ6DI" type="description">
<paragraph id="BFE07BEB30314F5DC9F8C50367766BB2">
<heading id="11A0CDD7D355EFB39E21C87D78C96084" reason="title">Type series composite redescription</heading>
</paragraph>
<paragraph id="C48D16C34023A8976AD7AEA0F2BBE9EE">
<emphasis id="9D46EF3CC7643DDAD1B7C8BABEAEC083" bold="true">
(Table
<tableCitation id="D4A28CBAD94D951BCD7EBF2EA71A991A" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
).
<emphasis id="72CFFA2FED175692922352DA247F3577" bold="true" italics="true">Colouration</emphasis>
</emphasis>
: Colouration of specimens preserved in ethanol is toasted to pale yellow (Figs
<figureCitation id="D81D8ED4CDB17FFE415404639D80B3AD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7</figureCitation>
,
<figureCitation id="71434FB52A75A19D68633572FE45C573" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 A – C</figureCitation>
). According to
<bibRefCitation id="2E76C9EF57C9A5C8DA25ABEF2BFEE674" author="Lucas PH" firstAuthor="Lucas" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique. Revue Zoologique 9: 283–289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas (1846)</bibRefCitation>
, colouration of living specimen was as follows [morphological traits are interpreted in brackets]:
</paragraph>
<paragraph id="2431A7C7675F66C63D3049DECFF1F428">
(Original colour description available from:
<ext-link id="4B207DEE4FA7A6EAE05DAE1FE50DD309" ext-link-type="uri" xlink_href="https://www.biodiversitylibrary.org/page/2362627#page/301/mode/1up" xlink_type="simple">https://www.biodiversitylibrary.org/page/2362627#page/301/mode/1up</ext-link>
)
</paragraph>
<paragraph id="7B2D01B423221A70C851E95F00834ED0">
Upper body part [anterior tergites] coppery black, lower [posterior tergites] green, in the middle ornamented with a yellowish green longitudinal stripe [probably, the translucence of the Malpighian tubule], [...] jaw [forcipules] reddish [...] palps greenish [second maxillae]; base of antenna green, in the middle greenish and in front stained dark red; feet [locomotory legs] green with dark red nails [unguis proper]; last pair of legs [
<abbrev id="ABBRID0E4AAK" xlink_title="ultimate legs">UL</abbrev>
] dark green [...].
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBBAK" type="description">
<paragraph id="2C12F42C570DCA0D542C5F92E15C3CD7">
<heading id="09F402B12AC70B53C1E5DC1152543D1F" reason="title">Morphological description of the type series.</heading>
</paragraph>
<paragraph id="B23DAAF4FFC4143E3F4F8E7F9F0B47F7">
[notes on brackets are comprehensive annotations from this text author]: Body length up to
<quantity id="CDC08E429DEDD51FAB794E1033DD4FF2" metricMagnitude="-2" metricUnit="m" metricValue="6.4" unit="mm" value="64.0">64 mm</quantity>
. Antennae reaching up to T 2 [maybe up to T 3; actual length shrivelled by ethanol retraction]; with 18 or 19 articles, the basal 5 ½ ones dorsally and ventrally glabrous (Fig.
<figureCitation id="9601C1910BF0B6BAE5CEC56540BA4C70" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 A, B</figureCitation>
). Antennae / cephalic plate length ratio: 2.01 [reduced because of the antennae articles retraction caused by the preservation in ethanol; estimated to be ~ 3.30].
</paragraph>
<paragraph id="B91302152E853F49F97D85642AE26950">
Cephalic plate with disperse puncta and a short anterior longitudinal depression; disto-median or / and paramedian sutures are absent. Posterior edge of cephalic plate overlapping the T 1 (Fig.
<figureCitation id="C40FCCFDA6451B9BCE22BC5FD7E14145" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 A</figureCitation>
). Coxosternal surface with disperse puncta, without sutures (Fig.
<figureCitation id="CC99AAEDE8FB0731E391BC25D706A801" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 B</figureCitation>
). Article 2 of second maxillary telopodite with spur. Forcipular trochanteroprefemoral process with one apical and two (rarely one) medial poorly differentiable teeth (Fig.
<figureCitation id="6E0A3705AE18E21C4B3B154FB8DD0952" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 C</figureCitation>
). Tooth plates slightly wider than tall, forming an obtuse angle (&gt; 120 °) with respect to the coxosternite; sensillae present. Each tooth plate with 4 + 4, rarely 3 + 3 teeth: the external one separated from the other three and inner ones sometimes fused. (Fig.
<figureCitation id="41C4CA162864EE1A5EA77F5455A54921" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 C</figureCitation>
).
</paragraph>
<paragraph id="C51124FC57E3F227B9AA041605FBF1C9">Spiracles triangular with three valves, present on body segments 3, 5, 8, 10, 12, 14, 16, 18, and 20.</paragraph>
<paragraph id="127B609706A9E1B4B9AED505F34B07E4">
T 1 without sutures or sulcus (Fig.
<figureCitation id="499D6F82C7BF5E4859D4C3E35F39E637" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 A</figureCitation>
); TT 2–20 with complete paramedian sutures (Fig.
<figureCitation id="C765CBA8E7D747A9BB3FEDEC6EAF00E9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 D</figureCitation>
); T 21 with a longitudinal median suture, surface of all tergites smooth, without depressions (Fig.
<figureCitation id="0F6668EDD87105343C4B350E0F410820" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 A</figureCitation>
). T 21 width / length ratio ≈ 1.35. Margination starting at T 14, being complete in TT 19–21.
</paragraph>
<paragraph id="F55A5E20C65D90C8AB210225D9506FE7">
Sternites with complete paramedian sutures from TT 2–20 (Fig.
<figureCitation id="D9AD15C2BDEF63D82EFB0DDC472F4F78" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 3–6 E sternites 6–8. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7 E</figureCitation>
). Sternite of ultimate leg-bearing segment (Fig.
<figureCitation id="62E4E3316543B525470D905460B83F24" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 B</figureCitation>
) with sides converging posteriorly; surface smooth, without depressions.
</paragraph>
<paragraph id="2F4DDDF459C13ECB5448EB5180FA60F6">
Coxopleuron not surpassing the posterior border of the tergite of the
<abbrev id="ABBRID0EYCAK" xlink_title="ultimate leg-bearing segment">
<collectionCode id="ACA79DD61DD0D389F76CF07ABBA58BE9">ULBS</collectionCode>
</abbrev>
(Fig.
<figureCitation id="A9223645E02361B079501897979F66D4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 B</figureCitation>
); with a dense pore-field area at the base and a single medio-distal spine. Coxopleural process sub-cylindrical and distinctly elongated, reaching up to the first 1 / 4 of the
<abbrev id="ABBRID0EADAK" xlink_title="ultimate legs">UL</abbrev>
prefemoral length. The complete surface is covered by a loose pore-field with some small setae (Fig.
<figureCitation id="32974AECB4B44192EB573328A2A4996D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 C</figureCitation>
) and a total of 9–14 usually asymmetrically disposed spines; 4–6 of them in an apical / subapical position;
<quantity id="868A55AC03906B15FF2C6F8BE3FD73E7" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.524" metricValueMin="0.762" unit="in" value="4.5" valueMax="6.0" valueMin="3.0">3–6 in</quantity>
a lateral position, and
<quantity id="E0A18A15816BC620759F05C3F1BA82E5" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
a dorsal position.
</paragraph>
<paragraph id="7A65F2C788C24DE8B1AD0B2063AEAACD">
Leg 1 with two distal tarsal spurs on tarsus 1, one lateral anterior and one ventral; legs 1–18 or 19 (mode 19, Table
<tableCitation id="E2C733D444E815ECF15059638FD85723" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
) with only one ventral tarsal spur. Tibial spurs absent. Legs 1, 2, or 3 with or without isolated setae. Legs 3 or 4–20 with scarce setae.
</paragraph>
<paragraph id="CA5AFF60D7ABD9918F7A1742F3CBBDA6">
<abbrev id="ABBRID0EQDAK" xlink_title="ultimate legs">UL</abbrev>
moderately long and slender with ratios of lengths of prefemur and femur = 1.15, femur and tibia = 1.25, tibia and tarsus 2 = 1.80; tarsus 1 and tarsus 2 = 1.20 (Fig.
<figureCitation id="3217C54B8A7BEF70494BCC962D7E9A71" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 C</figureCitation>
). Prefemora flattened dorsally, sometimes with a visible sinusoid transverse sulcus on the ventro-distal side (Fig.
<figureCitation id="527837CCC3169920F18E51EEB4F123EC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 A, B</figureCitation>
). Spines are variable in number (total: 17–26) and size, usually arranged in five frequently asymmetrical, duplicate, and non-ordered rows, with a formula:
<abbrev id="ABBRID0E3DAK" xlink_title="ventro-lateral spines">VL</abbrev>
: 3-6, V: 2-6,
<abbrev id="ABBRID0EAEAK" xlink_title="ventro-median spines">VM</abbrev>
: 2-6, M: 3-8 and
<abbrev id="ABBRID0EEEAK" xlink_title="dorso-median spines">DM</abbrev>
: 2-4.
<abbrev id="ABBRID0EIEAK" xlink_title="ventro-median spines">VM</abbrev>
and M spine rows usually converging proximally; between them a flattened surface is present, regularly rounded proximally and without spines. Prefemoral process inconspicuously prominent, sometimes scarcely setose, ending with two, or rarely three, spines (Figs
<figureCitation id="C05B58E7EF04734D39F061EF274E72A8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8 A – C</figureCitation>
). Setae on prefemur and femur almost absent, tibia distally scarce setose and tarsi 1 and 2 densely covered with short setae.
<abbrev id="ABBRID0EQEAK" xlink_title="ultimate legs">UL</abbrev>
/ T 21 length ratio ≈ 4.39 [reduced because of the retraction of the articles caused by the preservation in ethanol; ~ 5.15].
</paragraph>
<paragraph id="4F7DFDD3ED7B43AB5E4666A1F4ACE522">Genitalia retracted in the whole type series.</paragraph>
</subSubSection>
<subSubSection id="SECID0EVEAK" type="description">
<paragraph id="3EC8B5B1FABFDB0D6F3435062F6800CA">
<heading id="87B9FA09F0CC8BE06A2EEFB4D09DE599" reason="title">Genitalia description.</heading>
</paragraph>
<paragraph id="673B915A3DE0CFB83BF26175F789D16A">
(Based on
<taxonomicName id="11ED7C12C99BAD7B109ACA4B5ED18199" authorityName="Dufour" authorityYear="1820" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="viridipes">
<emphasis id="179E8F743EC79780406013427C060188" italics="true">S. viridipes</emphasis>
</taxonomicName>
and
<taxonomicName id="A72773513C154FF99659B2A6A05B887C" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="B5D54653A736AAE3971DF1E365E4FFDF" italics="true">S. chlorotes</emphasis>
</taxonomicName>
“ topotypical ” material; Fig.
<figureCitation id="FF8F3C73F93804244FA41937BB0325BE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9 A – D</figureCitation>
).
</paragraph>
<caption id="2A813C4F70F8361820F80BF77DAB547F" ID-DOI="10.3897/zookeys.1208.122126.figure9" ID-arpha="48954DD4-7472-57E6-B68B-11EF59BBA8F2" httpUri="https://binary.pensoft.net/fig/1101819" startId="F9">
<paragraph id="3DB595B4C58186DBE7E4F3D63CB25F5B">
<label id="0A43E8213C2F8721B74093882685B289">Figure 9.</label>
</paragraph>
<paragraph id="FD0F8303E03692219D2FA7EA42A039A2">
<taxonomicName id="07758465CAE6DE54D3047808B457392F" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="BF1518A7604C167EBD5C12292C3BBCEB" italics="true">Scolopendra oraniensis</emphasis>
</taxonomicName>
genitalia
<emphasis id="34665837D85810FC258DD188E071F9BD" bold="true">A</emphasis>
male (CEUAMr 24); and
<emphasis id="06BDD1441686C6236D91299A1D1AB21E" bold="true">B</emphasis>
female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (
<emphasis id="B3E5FF61F0DACC94BDD7ACF50FE53BAB" bold="true">C, D)</emphasis>
on the same specimens
<emphasis id="1B4DC3163EAEDA38DFA63FE191408835" bold="true">C</emphasis>
male and
<emphasis id="6DECC5713FCBD13A138EBE70D3A2B10E" bold="true">D</emphasis>
female. Observe that on the contrary that
<bibRefCitation id="16D2FC0AB2DA76A7EC971BAEE0705737" author="Würmli M" firstAuthor="Wurmli" journalOrPublisher="Sitzungsberichte der Österreichischen Akademie der Wissenschaften" pagination="315-353" refId="B80" refString="Würmli M (1980) Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha). Sitzungsberichte der Österreichischen Akademie der Wissenschaften 189: 315–353." title="Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha)." volume="189" year="1980">Würmli (1980)</bibRefCitation>
indicated, the numbers of setae are actually similar in both genders. Legend:
<abbrev id="ABBRID0EWGAK" xlink_title="anal valve">AV</abbrev>
, anal valve; G. gonopod (folded back);
<abbrev id="ABBRID0E1GAK" xlink_title="lamina adanalis">LA</abbrev>
, lamina adanalis;
<abbrev id="ABBRID0E5GAK" xlink_title="lateral spines">LS</abbrev>
, lamina subanalis; P, penis;
<abbrev id="ABBRID0ECHAK" xlink_title="sternite of genital segment 1">SGS I</abbrev>
, sternite of genital segment 1;
<abbrev id="ABBRID0EGHAK" xlink_title="sternite of genital segment 2">SGS II</abbrev>
, sternite of genital segment 2. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="0CC96C656B2E2BE291491C3E1DCA2985">
Genitalia well developed, reaching the apical part of coxopleural process when extended. Sternite and tergite of genital segment 1 (TGS and
<abbrev id="ABBRID0ERHAK" xlink_title="sternite of genital segment 1">SGS I</abbrev>
) convex and distally round, with a proximal median suture – distally attenuated – forming the vertex of a poorly angulated keel (Fig.
<figureCitation id="B48D8CD22776702C4B7FC30BDBEA4D07" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9 A, B</figureCitation>
). In males (Fig.
<figureCitation id="775BD33233FB01003E9B867255E5E137" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9 A</figureCitation>
) tergite of genital segment 2 (
<abbrev id="ABBRID0E4HAK" xlink_title="sternite of genital segment 2">SGS II</abbrev>
) small and horseshoe-shaped, with a poorly visible longitudinal suture. Gonopods short and small. Penis present, with fee endpoint, ventally serrated. TGS,
<abbrev id="ABBRID0EBIAK" xlink_title="sternite of genital segment 1">SGS I</abbrev>
,
<abbrev id="ABBRID0EFIAK" xlink_title="sternite of genital segment 2">SGS II</abbrev>
, lamina adanalis, gonopods and penis carrying scatted small setae, the remaining part of the genital structures glabrous. Sexual dimorphism or secondary sexual characteristics indistinct (Figs
<figureCitation id="3835E6C92D664BA3E432095C8FBCF487" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9 C, D</figureCitation>
);
<abbrev id="ABBRID0ENIAK" xlink_title="ultimate legs">UL</abbrev>
tibia and tarsi weakly and inconstantly more hirsute in females, with gender differences frequently indistinguishable.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERIAK" type="Species variability">
<paragraph id="B398C2716F005DEE5637187A1A83A144">
<heading id="13261E8E82D1195A115D7E0EDE1DF547" reason="title">Species variability.</heading>
</paragraph>
<paragraph id="2C74AF044F77C0EBE2D898130FA77C37">
<taxonomicName id="CC8CAABC4CB0848C333852CF2CFAAB40" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="66A95AB6E1A20CDA658C4979001F3C82" italics="true">Scolopendra oraniensis</emphasis>
</taxonomicName>
has been observed to be somewhat variable in some few morphological features (Figs
<figureCitation id="22C3BEB3C0A906B3C08E8FA6A58E8639" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A cephalic plate, antennae, and TT 1–5 B forcipular segment and sternites 1–3 C tooth plates D tergites 13–18 E sternites 10–15. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure3" httpUri="https://binary.pensoft.net/fig/1101813">3</figureCitation>
–
<figureCitation id="2CDFDBD76F7FEF1F24B06B6D30C5AAC8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9 B</figureCitation>
; Table
<tableCitation id="352581B3D34EB6287DDFA2354B7D60AC" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
, Suppl. material
<xref id="608D742AD954AB6C2F9783086C4E1619" ref-type="supplementary-material" rid="S1">1</xref>
: files 1, 3): number of antennal articles (17–20; mode 18–19), number of hirsute basal antennal articles (5–5 ½), number of teeth on tooth plates and on forcipular trochanteroprefemoral processes (respectively 3 + 3 vs 4 + 4; mode 4 + 4 and 3 vs 2; mode 3), beginning of complete margination of tergites (on TT 17, 19, or 20; mode 17), tarsal spurs on leg 1 (0-2; mode 2), single tarsal spurs on legs (2–18 or 19; mode 2–19), coxopleural process morphology and
<abbrev id="ABBRID0ESJAK" xlink_title="ultimate legs">UL</abbrev>
prefemur number of spines (respectively 9–14 and 18–32), visibility of
<abbrev id="ABBRID0EWJAK" xlink_title="ultimate legs">UL</abbrev>
prefemoral ventro-distal horizontal suture, coxopleural and prefemoral process lengths or number of setae on the
<abbrev id="ABBRID0E1JAK" xlink_title="ultimate legs">UL</abbrev>
tarsi.
</paragraph>
<paragraph id="90A5475BA3DD6118304BA1817A950537">
Additionally, the colouration has proven to be variable sympatrically (Figs
<figureCitation id="B832FF340A0543F55A84DF842C745C1A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Representatives of Scolopendromorpha from the Iberian Peninsula (uncollected) A family Plutoniumidae Bollman, 1893: Theatops erytrocephalus (C. L. Koch, 1847), Seoane do Caurel, Lugo (credit: J. Tizón) B family Cryptopidae Kohlrausch, 1881: Cryptops sp. Leach, 1814, Villena, Alacant (credit: D. Molina) C – I family Scolopendridae Leach, 1814 C Scolopendra cingulata Latreille, 1829, Benilloba, Alacant (credit: M. Huesca and CD) and D – I S. oraniensis Lucas, 1840. Observe that in this species the colour variants can occur sympatrically D, H, I Benilloba, Alacant and E – G Almeria Province, Andalucia (credit: F. Rodriguez Luque)." figureDoi="10.3897/zookeys.1208.122126.figure1" httpUri="https://binary.pensoft.net/fig/1101811">1 D – I</figureCitation>
,
<figureCitation id="ADA3468D16886BDD84C16D666A11533F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A cephalic plate, antennae, and TT 1–5 B forcipular segment and sternites 1–3 C tooth plates D tergites 13–18 E sternites 10–15. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure3" httpUri="https://binary.pensoft.net/fig/1101813">3</figureCitation>
–
<figureCitation id="8E62E775FB8D2AA2D9AEB5F3CEA20F23" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9</figureCitation>
). The cephalic plate and tergites can be monochromatic pale yellow to dark green / brown, sometimes reddish, sometimes with a darker / lighter longitudinal stripe. The forcipules can be yellowish to reddish, while the legs and antennae can be pale green, yellowish, bluish, reddish, or bicoloured, with occasional metallic reflections (Figs
<figureCitation id="EB2FCD4A45CDE61ED28908F61EB9AA57" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Representatives of Scolopendromorpha from the Iberian Peninsula (uncollected) A family Plutoniumidae Bollman, 1893: Theatops erytrocephalus (C. L. Koch, 1847), Seoane do Caurel, Lugo (credit: J. Tizón) B family Cryptopidae Kohlrausch, 1881: Cryptops sp. Leach, 1814, Villena, Alacant (credit: D. Molina) C – I family Scolopendridae Leach, 1814 C Scolopendra cingulata Latreille, 1829, Benilloba, Alacant (credit: M. Huesca and CD) and D – I S. oraniensis Lucas, 1840. Observe that in this species the colour variants can occur sympatrically D, H, I Benilloba, Alacant and E – G Almeria Province, Andalucia (credit: F. Rodriguez Luque)." figureDoi="10.3897/zookeys.1208.122126.figure1" httpUri="https://binary.pensoft.net/fig/1101811">1 D – I</figureCitation>
,
<figureCitation id="CE7120A2939DF058A27720A57F8655C3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A cephalic plate, antennae, and TT 1–5 B forcipular segment and sternites 1–3 C tooth plates D tergites 13–18 E sternites 10–15. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure3" httpUri="https://binary.pensoft.net/fig/1101813">3</figureCitation>
,
<figureCitation id="43C598BE84B89C6FD68BF5906B825564" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9</figureCitation>
).
</paragraph>
<paragraph id="2A25DE2C492BB123B325B5D669358980">
These morphological and colouration characteristics, although tending towards a certain geographic distribution, often turn out to be variable within those populations (Table
<tableCitation id="B8269F3AE3F55AFA496A58D8017BEAE7" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
, Suppl. material
<xref id="3473BFBC99184870EF88883CF0843713" ref-type="supplementary-material" rid="S1">1</xref>
: files 1, 3). Therefore, the separation of the species by their old names are clearly unjustified, since all the features can be explained by the known intraspecific variability of the taxon (
<bibRefCitation id="D32E97C2EF7D3C78371E97574A7B5055" author="Iorio and Geoffroy" firstAuthor="Iorio" journalOrPublisher="Le bulletin d’ Arthropoda" pagination="48-51" refId="B34" refString="Iorio É, Geoffroy JJ (2006) Contribution à la connaissance de Scolopendra oraniensis H. Lucas, 1846 (Chilopoda, Scolopendromorpha, Scolopendridae). Le bulletin d’ Arthropoda 27: 48–51." title="Contribution à la connaissance de Scolopendra oraniensis H. Lucas, 1846 (Chilopoda, Scolopendromorpha, Scolopendridae)." volume="27" year="2006">Iorio and Geoffroy 2006</bibRefCitation>
;
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), which is also shared with many other species in the genus (
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;
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;
<bibRefCitation id="EB28D49FE29808C760CC004A6568372E" DOI="10.11646/zootaxa.4483.3.1" author="Domenech" etAl="et al." firstAuthor="Domenech" issue="3" journalOrPublisher="Zootaxa" pagination="401-427" refId="B18" refString="Doménech C, Barberá VM, Larriba E (2018) A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species. Zootaxa 4483 (3): 401–427. https://doi.org/10.11646/zootaxa.4483.3.1" title="A phylogenetic approach to the Philippines endemic centipedes of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae), with the description of a new species." volume="4483" year="2018">Doménech et al. 2018</bibRefCitation>
;
<bibRefCitation id="1BC5E72BA0287C9FE269C83515D4CC6B" DOI="10.37828/em.2021.41.6" author="Dyachkov and Nedoev" firstAuthor="Dyachkov" journalOrPublisher="Ecologica Montenegrina" pagination="41-50" refId="B22" refString="Dyachkov YV, Nedoev KK (2021) A contribution to the centipede (Chilopoda: Geophilomorpha, Scolopendromorpha) fauna of Uzbekistan and Turkmenistan. Ecologica Montenegrina 41: 41–50. https://doi.org/10.37828/em.2021.41.6" title="A contribution to the centipede (Chilopoda: Geophilomorpha, Scolopendromorpha) fauna of Uzbekistan and Turkmenistan." volume="41" year="2021">Dyachkov and Nedoev 2021</bibRefCitation>
;
<bibRefCitation id="05761CEC5A156DC896B725E641FCA918" DOI="10.11646/zootaxa.4952.3.3" author="Tsukamoto" etAl="et al." firstAuthor="Tsukamoto" issue="3" journalOrPublisher="Zootaxa" pagination="465-494" refId="B73" refString="Tsukamoto S, Hiruta SF, Eguchi K, Liao JR, Shimano S (2021) A new amphibious species of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae) from the Ryukyu Archipelago and Taiwan. Zootaxa 4952 (3): 465–494. https://doi.org/10.11646/zootaxa.4952.3.3" title="A new amphibious species of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae) from the Ryukyu Archipelago and Taiwan." volume="4952" year="2021">Tsukamoto et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5LAK" type="Differential diagnosis">
<paragraph id="9BCE576AA826B5E80F0D1963B569F72F">
<heading id="97E8A65BC42BD571DA5FCDD64BB02674" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="AFD6B1DBE9661DC5B00C76083D1E5899">
Morphologically,
<taxonomicName id="2614BBBD306DD0F9E33712B2E29114C3" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="A77A2B93F849A78BADB28AFADDC6F93E" italics="true">S. oraniensis</emphasis>
</taxonomicName>
is related to five other species of the
<taxonomicName id="D0A20DD0F86EC5C2353C6A7CA54E21F8" class="Chilopoda" family="Scolopendridae" genus="Rhadinoscytalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="canidens">
<emphasis id="E2F307A72F27F9D8668EDF4F3DD89445" italics="true">S. canidens</emphasis>
</taxonomicName>
group distributed around the Mediterranean Sea and Middle Asia (
<bibRefCitation id="9148862F794F6134A4870E8CE8BE4DF7" author="Wurmli" firstAuthor="Wurmli" journalOrPublisher="Sitzungsberichte der Österreichischen Akademie der Wissenschaften" pagination="315-353" refId="B80" refString="Würmli M (1980) Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha). Sitzungsberichte der Österreichischen Akademie der Wissenschaften 189: 315–353." title="Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha)." volume="189" year="1980">Würmli 1980</bibRefCitation>
;
<bibRefCitation id="771D8BF5056EB9C6025D2B0BA44C6DA4" DOI="10.11646/zootaxa.1792.1.3" author="Simaiakis and Mylonas" firstAuthor="Simaiakis" issue="1" journalOrPublisher="Zootaxa" pagination="39-53" refId="B70" refString="Simaiakis S, Mylonas M (2008) The Scolopendra species (Chilopoda: Scolopendromorpha: Scolopendridae) of Greece (E-Mediterranean): a theoretical approach on the effect of geography and palaeogeography on their distribution. Zootaxa 1792 (1): 39–53. https://doi.org/10.11646/zootaxa.1792.1.3" title="The Scolopendra species (Chilopoda: Scolopendromorpha: Scolopendridae) of Greece (E-Mediterranean): a theoretical approach on the effect of geography and palaeogeography on their distribution." volume="1792" year="2008">Simaiakis and Mylonas 2008</bibRefCitation>
).
<taxonomicName id="986C2B2F10E5EA2DB95A6FD39D0E1076" authorityName="Silvestri" authorityYear="1897" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="5D593DB0DAB293D9A0818607389ABEA1" italics="true">Scolopendra clavipes</emphasis>
</taxonomicName>
(Middle Asia) can easily be distinguished from
<taxonomicName id="A372AF71AE140B29184C57F9BB78510F" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="C0DDD3401D3A4C190D114039858FE45B" italics="true">S. oraniensis</emphasis>
</taxonomicName>
by the clavate morphology of the
<abbrev id="ABBRID0EYNAK" xlink_title="ultimate legs">UL</abbrev>
, while the two subspecies of
<taxonomicName id="DE469E35AE0D006DEC430BE475430941" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="dalmatica">
<emphasis id="9703B159A59A3876DA591787C0A7537A" italics="true">S. dalmatica</emphasis>
C. L.
<bibRefCitation id="C4BC434999A6D9D4FE10B59C4C7E26CE" author="Koch CL" journalOrPublisher="Pustet, Regensburg" refId="B38" refString="Koch CL (1847) System der Myriapoden. In: Herrich-Schaffer L (Ed.) Kritische Revision der Insectenfauna Deutschlands. Pustet, Regensburg, 270 pp." title="System der Myriapoden. In: Herrich-Schäffer L (Ed.) Kritische Revision der Insectenfauna Deutschlands." year="1847">Koch, 1847</bibRefCitation>
</taxonomicName>
(northeastern Mediterranean) can be differentiated by their larger size (up to
<quantity id="7C53F20A4777EAC5BF6C5864DA2EB3F0" metricMagnitude="-2" metricUnit="m" metricValue="8.0" unit="mm" value="80.0">80 mm</quantity>
), the incomplete median suture of T 21, and the four spines of the
<abbrev id="ABBRID0EHOAK" xlink_title="ultimate legs">UL</abbrev>
prefemoral process. However, the morphologically closest relatives to
<taxonomicName id="1054D52040C0993E08EBECD6286C01E0" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="0D248A4B83248BF446B2EB2D7E664524" italics="true">S. oraniensis</emphasis>
</taxonomicName>
are
<taxonomicName id="CF9CFB756DB91DE1D2B52E5E645CE1DA" class="Chilopoda" family="Scolopendridae" genus="Rhadinoscytalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="canidens">
<emphasis id="C2E9536E228F8DDC82C1BFE110865C93" italics="true">S. canidens</emphasis>
</taxonomicName>
(southeastern and eastern Mediterranean) and
<taxonomicName id="5134F4944263E81D2A4A92049223A8BC" authority="Lucas, 1853" authorityName="Lucas" authorityYear="1853" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="cretica">
<emphasis id="73D1ED94918389CDC8CEEB52FCA94388" italics="true">S. cretica</emphasis>
<bibRefCitation id="50A434AABC4C4EF7D4B078C7E60E60BD" author="Lucas PH" refId="B53" refString="Lucas PH (1853) Essai sur les animaux articules qui habitent l'ile de Crete - Revue et Magasin de Zoologie Pure et Appliquee 5 (2): 514 - 531." year="1853">Lucas, 1853</bibRefCitation>
</taxonomicName>
(eastern Mediterranean, probably endemic to
<collectingRegion id="2FBAD82F1AA658181FB730BD03C2EBE2" country="Greece" name="Kriti">Crete</collectingRegion>
,
<collectingCountry id="DF0C11681A59DECDD0D2DC38B7A26B4C" name="Greece">Greece</collectingCountry>
). These two latter species can be unequivocally differentiated from
<taxonomicName id="A3D027F87D6A237FE6697D0BF9EDCB3E" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="664C06C6A3EBFCA9FB5BEC9FFB5E0B80" italics="true">S. oraniensis</emphasis>
</taxonomicName>
by the number of glabrous articles in the antennae (10–12 vs 5–5 ½, respectively), the type of transition between the glabrous / hirsute articles (gradual vs abrupt), and the number of spines in the prefemoral process (generally 3 vs 2 or 3). Between
<taxonomicName id="ED2E379A9748C43592F57323479D86F1" class="Chilopoda" family="Scolopendridae" genus="Rhadinoscytalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="canidens">
<emphasis id="0107DC8C8FF11FDB6B1D541A8C2344AF" italics="true">S. canidens</emphasis>
</taxonomicName>
and
<taxonomicName id="29A36DC1957BC3A76D7A03533F495AD6" authorityName="Lucas" authorityYear="1853" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="cretica">
<emphasis id="97A534CF0F47CDF0C0CCFCCB0857A0D1" italics="true">S. cretica</emphasis>
</taxonomicName>
, the only visible difference is the presence of brush-like setae in tarsi 1 and 2 of the
<abbrev id="ABBRID0ENQAK" xlink_title="ultimate legs">UL</abbrev>
in the
<taxonomicName id="40C50316546C8D1B3AA3C5B279724F58" authorityName="Lucas" authorityYear="1853" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="cretica">
<emphasis id="59D7B0F02350C8FCE005F29B43B121E2" italics="true">S. cretica</emphasis>
</taxonomicName>
females (
<bibRefCitation id="9882F2DDCE18AD3046BE6E487913ACEB" author="Wurmli" firstAuthor="Wurmli" journalOrPublisher="Sitzungsberichte der Österreichischen Akademie der Wissenschaften" pagination="315-353" refId="B80" refString="Würmli M (1980) Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha). Sitzungsberichte der Österreichischen Akademie der Wissenschaften 189: 315–353." title="Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha)." volume="189" year="1980">Würmli 1980</bibRefCitation>
;
<bibRefCitation id="524EBEA10DF3D481C66E692D6B540FBF" DOI="10.1163/187525410X12578602960380" author="Lewis" firstAuthor="Lewis" issue="1" journalOrPublisher="International Journal of Myriapodology" pagination="83-122" refId="B47" refString="Lewis JGE (2010) A key and annotated list of the Scolopendra species of the Old World with a reappraisal of Arthrorhabdus (Chilopoda: Scolopendromorpha: Scolopendridae). International Journal of Myriapodology 3 (1): 83–122. https://doi.org/10.1163/187525410X12578602960380" title="A key and annotated list of the Scolopendra species of the Old World with a reappraisal of Arthrorhabdus (Chilopoda: Scolopendromorpha: Scolopendridae)." volume="3" year="2010">Lewis 2010</bibRefCitation>
; Dyachkov (
<named-content id="D719A6E02DD0943254D6831F4E24B49E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/altai-state-university" xlink_title="Altai State University">
<collectionCode id="72B9CB0BC907BC9ECB215BF25728D778" country="USA" lsid="urn:lsid:biocol.org:col:15600" name="Arizona State University" type="Herbarium">ASU</collectionCode>
</named-content>
) pers. comm. 2022; Huesca pers. comm. 2018).
</paragraph>
<paragraph id="02BB4A670E0A3A7C24B4BC84F9D26F15">
In these last two species closest to
<taxonomicName id="ABD1FEF03769ECCBF1D1E3283B259EB3" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="152BBCF6E6753340009944E847B4F725" italics="true">S. oraniensis</emphasis>
</taxonomicName>
, comparative analyses such as antennae / cephalic plates and T 21 /
<abbrev id="ABBRID0EVRAK" xlink_title="ultimate legs">UL</abbrev>
length ratios and their genitalia descriptions have not been performed yet.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZRAK" type="Nomenclatural considerations">
<paragraph id="93456C5539626B343C62DE048723BFD8">
<heading id="6C0DAA3A6A164B59C8DEEA4442B8D106" reason="title">Nomenclatural considerations.</heading>
</paragraph>
<paragraph id="DA1C6FBDF5FBB327D5F2702205D55FB0">
To verify the actual identities of
<taxonomicName id="65759590EC3AB92FDF95FA961D65B26D" authorityName="Dufour" authorityYear="1820" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="viridipes">
<emphasis id="321193921E7610BB8016DB78A0F795CE" italics="true">S. viridipes</emphasis>
</taxonomicName>
and
<taxonomicName id="7E56E87954E87BD1E7F072AD72A8F3E9" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="E05DE855D4E988BA5DDB581BA2CA4209" italics="true">S. chlorotes</emphasis>
</taxonomicName>
, establishing the
<taxonomicName id="A98B0FB26C801F15ACBD0CC66560C20F" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="B820CA05019EBD1B6B473E6B22CA892B" italics="true">S. oraniensis</emphasis>
</taxonomicName>
specimens
<bibRefCitation id="CD8C3D5C7E15AE2D019A2257445E4120" author="Lucas PH" firstAuthor="Lucas" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique. Revue Zoologique 9: 283–289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas (1846)</bibRefCitation>
on which he based the type series was necessary, to have reference material with which to unambiguously compare new material (
<bibRefCitation id="5F8D30265882ED93FF87F90154BF3D31" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Preamble, Art. 13, Recomm. 13 A). The
<taxonomicName id="8428F8DBCAF195F89DF776B80D284348" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="67F15926F1AA56B1790324D8A4DA9CFA" italics="true">S. oraniensis</emphasis>
</taxonomicName>
<typeStatus id="037FABEB8489782EE1AFEA5494341D27" type="lectotype">lectotype</typeStatus>
designation was found to be not nomenclaturally required (
<bibRefCitation id="ADA918218A78B5647E252E845B6F5887" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: 11; point
<quantity id="2FD354981F24EF3DB2656BE1339DAE10" metricMagnitude="-1" metricUnit="m" metricValue="1.524" unit="in" value="6.0">6 in</quantity>
introduction; Art. 72.1. 1; 74.7). However,
<typeStatus id="C0B56336325A6ED3167BB31284A39AFE" type="neotype">neotypes</typeStatus>
designations for
<taxonomicName id="6EDBB585CD4F0C98E1DD818EA65B03CE" authorityName="Dufour" authorityYear="1820" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="viridipes">
<emphasis id="F33543936320E0A37D2A939C551F5047" italics="true">S. viridipes</emphasis>
</taxonomicName>
and
<taxonomicName id="B371C74DC7728ECD0C9274C56DBD4CE6" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="08E10DFCE68B0786CB4CE001BA2A0C5F" italics="true">S. chlorotes</emphasis>
</taxonomicName>
were mandatory to clarify their type localities and taxonomic situations in the absence of the original types (
<bibRefCitation id="249645D269884CAB948B0D1CC8EAE004" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 75, 76). Finally, the identities of
<taxonomicName id="32C901927400CDE1B46104DCD81C4247" authorityName="Dufour" authorityYear="1820" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="viridipes">
<emphasis id="38C70168D4F87CB198B79BBAB354F6E3" italics="true">S. viridipes</emphasis>
</taxonomicName>
,
<taxonomicName id="519C73C168463AD17AD01DDC5537EEAE" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="FE66D4B8BF10A41205FB76D5697D5EBA" italics="true">S. chlorotes</emphasis>
</taxonomicName>
, and
<taxonomicName id="8BC18737343DE65ADB75D299AA9FD778" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="365EB4A932903B207D8F0FC1FB36A9D2" italics="true">S. oraniensis</emphasis>
</taxonomicName>
have been solved, and they are all conspecific (Figs
<figureCitation id="D534771378EDA02E26F2C577909C148D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A cephalic plate, antennae, and TT 1–5 B forcipular segment and sternites 1–3 C tooth plates D tergites 13–18 E sternites 10–15. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure3" httpUri="https://binary.pensoft.net/fig/1101813">3</figureCitation>
–
<figureCitation id="9C8466F3EB1EF6AA8AA10C8D77CE6D75" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelin’s 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8</figureCitation>
; Table
<tableCitation id="927843D5C45E9DA6C1DA9A11C83CE88D" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes ’ and S. chlorotes ’ topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
) leaving the name
<taxonomicName id="CF9FBAB96C22C52113FA89AEA26A53BA" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="84F10F6DAC894CFDDF9F3F4EBE56D617" italics="true">S. oraniensis</emphasis>
<bibRefCitation id="B2D17D6387434300C83FFED98202F138" author="Lucas PH" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du Nord de l'Afrique. Revue Zoologique 9: 283 - 289." title="Note sur quelques nouvelles espèces d’insectes qui habitent les possessions francaises du Nord de l’Afrique." volume="9" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
as nomen protectum with which to unambiguously refer to “ all ” of these species (
<bibRefCitation id="BF91FBDA820E713296830BDBFDFB930E" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 23.3 and 23.9. 2).
</paragraph>
<paragraph id="DE1125AFF445CEEF13A76BDFF3B18226">
At a nomenclatural level, the specific epithet of
<taxonomicName id="560CE350627FFF5E5FCBD1DA3F2F8C2B" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="30347E30577E99BDE8AE3B74DC6AF036" italics="true">S. oraniensis</emphasis>
</taxonomicName>
obviously conformed to the reference of the type locality (
<bibRefCitation id="B64C8AC00B8F48E1A50C267FF3123CA6" author="Gervais" firstAuthor="Gervais" journalOrPublisher="Librairie encyclopédique de Roret, Paris" refId="B28" refString="Gervais MP (1847) Histoire naturelle des insectes. Aptères. Librairie encyclopédique de Roret, Paris, 623 pp." title="Histoire naturelle des insectes. Aptères." year="1847">Gervais 1847</bibRefCitation>
). However, that noun is composed by the feminine (or masculine) suffix “ -
<emphasis id="A7C00BC05E3457B64E046584338939CA" italics="true">ensis</emphasis>
” (coming from) and the prefix (a toponym) “ Orani ”, in the place of “
<collectingCountry id="ED400CFD724C6AACA05E26E8C0781D13" name="Algeria">Oran</collectingCountry>
”. Coincidentally, “ Orani ” is the name of two other localities located in
<collectingCountry id="863F8984364D4141BD6ACDFE5B670E11" name="Italy">Italy</collectingCountry>
and the
<collectingCountry id="76D74D1BCF125838104F7377050D0E92" name="Philippines">Philippines</collectingCountry>
. This can lead to etymological confusion, especially for the Italian Orani, on Sardinia, a region where
<taxonomicName id="9C1518D1011120829C0C08A4EFDADD45" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="274EE672381F081304FBFAA109C581B9" italics="true">S. oraniensis</emphasis>
</taxonomicName>
also lives (
<bibRefCitation id="EA6DA8706866E13D20E0095998964760" author="Wurmli" firstAuthor="Wurmli" journalOrPublisher="Sitzungsberichte der Österreichischen Akademie der Wissenschaften" pagination="315-353" refId="B80" refString="Würmli M (1980) Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha). Sitzungsberichte der Österreichischen Akademie der Wissenschaften 189: 315–353." title="Statistische Unterschungen zur Systematik und postembryonalen Entwicklung der Scolopendra canidens Gruppe (Chilopoda: Scolopendromorpha)." volume="189" year="1980">Würmli 1980</bibRefCitation>
;
<bibRefCitation id="BBDA3AFE07DDE165F47509FE497C831D" author="Bonato L &amp; Chagas A Junior &amp; Edgecombe GD &amp; Lewis JGE &amp; Minelli A &amp; Pereira LA &amp; Shelley RM &amp; Stoev P &amp; Zapparoli M" refId="B5" refString="Bonato L, Chagas A Junior, Edgecombe GD, Lewis JGE, Minelli A, Pereira LA, Shelley RM, Stoev P, Zapparoli M (2017 a) ChiloBase 2.0. A World Catalogue of Centipedes (Chilopoda). http://chilobase.biologia.unipd.it [Accessed 10 Jan. 2020]" url="http://chilobase.biologia.unipd.it">Bonato et al. 2017 a</bibRefCitation>
). Therefore, it is thought to be an incorrect original spelling (
<bibRefCitation id="62C576A1EA65C1D7D58F7C0BE63602BF" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 32.4) and according to the author’s intention to honour
<collectingCountry id="FA9C50EE6B25D911BD8BFC8C53709EB6" name="Algeria">Oran</collectingCountry>
,
<collectingCountry id="8C12FC30D8AE0A54B06CD83E5AC73782" name="Algeria">Algeria</collectingCountry>
, rather than Orani,
<collectingCountry id="C8E1DA3816F1698A7EB8AEE2C96CDB52" name="Italy">Italy</collectingCountry>
, the proper name for
<taxonomicName id="32F97E8F246BAFA2B07CEE29FFDDCBAB" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="2DBE1AB621F773B4F280418598790021" italics="true">S. oraniensis</emphasis>
</taxonomicName>
should be amended as “
<taxonomicName id="96B81C5C46AD9926A2AB14B341555F90" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="64662748BC7ED74A8F2D6ADC1B066780" italics="true">S. oraniensis</emphasis>
</taxonomicName>
”. However,
<bibRefCitation id="63B82E9A3D654B078833080E455C0FAA" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN (1999</bibRefCitation>
: Art. 32.5. 1) clearly states that this supposition should not be adhered to since, in the original publication, “ The […] use of an inappropriate connecting vowel is not to be considered inadvertent errors ”. Therefore, despite being confusing or misspelled, this name must not be corrected as it is valid in its original spelling;
<taxonomicName id="F90D8C5F959BC214B605E1C91AE936DA" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="879892FC19556E248A2E86F85D78B2A3" italics="true">S. oraniensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>