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<document id="4EF04697ABD275BF236DC94679226088" ID-DOI="10.4081/nhs.2018.358" ID-ISSN="2385-0922" ID-Zenodo-Dep="12523821" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1719276347457" checkinUser="felipe" docAuthor="Garassino, Alessandro &amp; Pasini, Giovanni" docDate="2018" docId="1F275648FFDDFFF1FC2A3A9C64E3FD7F" docLanguage="en" docName="NatHistSci.5.1.11-18.pdf" docOrigin="Natural History Sciences 5 (1)" docSource="http://dx.doi.org/10.4081/nhs.2018.358" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.3:NatHistSci.2014-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2014-.journal_article" docStyleVersion="3" docTitle="Amazighopsidae nov." docType="treatment" docVersion="5" lastPageNumber="13" masterDocId="E31E2E30FFDCFFF3FFF73F0B6146FFFC" masterDocTitle="Amazighopsidae, a new family of decapod macruran astacideans from the late Cretaceous (Cenomanian-Turonian) of Gara Sbaa, Southeastern Morocco" masterLastPageNumber="18" masterPageNumber="11" pageNumber="12" updateTime="1728074577425" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="18D72975CCF0FFBF2397DB46B3693932">Amazighopsidae, a new family of decapod macruran astacideans from the late Cretaceous (Cenomanian-Turonian) of Gara Sbaa, Southeastern Morocco</mods:title>
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<mods:namePart id="A600DBBBD228FF2389EB49F3415B2F48">Garassino, Alessandro</mods:namePart>
<mods:affiliation id="4C5F7970B8A08C29EB53A803467F0FF7">Sezione di Paleontologia degli Invertebrati, Museo di Storia Naturale, Corso Venezia 55, 20121 Milano, Italia.</mods:affiliation>
<mods:nameIdentifier id="5C2194B2D8DE87A75C6BC548A2C47747" type="email">alessandro.garassino@comune.milano.it</mods:nameIdentifier>
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<mods:namePart id="D0808E9C3660411EEC7262634C33105D">Pasini, Giovanni</mods:namePart>
<mods:affiliation id="6A0A10B6F0A7A8AD89B8D7167FCBFA84">Via Alessandro Volta 16, 22070 Appiano Gentile (Como), Italia. E-mail: juanaldopasini @ tiscali. it</mods:affiliation>
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<treatment id="1F275648FFDDFFF1FC2A3A9C64E3FD7F" ID-DOI="http://doi.org/10.5281/zenodo.13891966" ID-Zenodo-Dep="13891966" LSID="urn:lsid:plazi:treatment:1F275648FFDDFFF1FC2A3A9C64E3FD7F" httpUri="http://treatment.plazi.org/id/1F275648FFDDFFF1FC2A3A9C64E3FD7F" lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12">
<subSubSection id="DF94B4D5FFDDFFF2FC2A3A9C6463FA4D" box="[989,1317,1431,1457]" pageId="1" pageNumber="12" type="nomenclature">
<paragraph id="9731E75EFFDDFFF2FC2A3A9C6463FA4D" blockId="1.[952,1353,1370,1457]" box="[989,1317,1431,1457]" pageId="1" pageNumber="12">
<heading id="CC795032FFDDFFF2FC2A3A9C6463FA4D" box="[989,1317,1431,1457]" centered="true" fontSize="36" level="2" pageId="1" pageNumber="12" reason="7">
Family
<taxonomicName id="508E9CDDFFDDFFF2FBC53A9C6463FA4D" box="[1074,1317,1431,1457]" class="Malacostraca" family="Amazighopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="12" phylum="Arthropoda" rank="family">Amazighopsidae nov.</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDDFFF2FC923AD86453F8C4" pageId="1" pageNumber="12" type="diagnosis">
<paragraph id="9731E75EFFDDFFF2FC923AD86453F8C4" blockId="1.[832,1474,1491,1969]" pageId="1" pageNumber="12">
<emphasis id="A5FA3B4CFFDDFFF2FC923AD8629EFA11" bold="true" box="[869,984,1491,1517]" pageId="1" pageNumber="12">Diagnosis</emphasis>
: Carapace subcylindrical; triangular, short rostrum with serrate suprarostral margin, bearing 3-4 pairs of teeth; deep cervical groove strongly inclined forward, reaching the dorsal margin at the anterior third of the total carapace length (excluding rostrum); antennal groove short and weak; cephalic region shorter than branchial one; s1 short; s2 with subrounded pleura partially overlapping those of s1 and s3; s1, s3-s6 smooth with rounded pleurae; subrectangular telson without diaeresis and lateral margins with one median movable spine and two distal movable spines; chelate P1 with weak heterochely; uropodal exopod without diaeresis.
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDDFFF2FC92383662F1F889" pageId="1" pageNumber="12" type="type_taxon">
<paragraph id="9731E75EFFDDFFF2FC92383662F1F889" blockId="1.[832,1474,1491,1969]" pageId="1" pageNumber="12">
<emphasis id="A5FA3B4CFFDDFFF2FC92383665D5F8AB" bold="true" box="[869,1171,1852,1879]" pageId="1" pageNumber="12">
<typeStatus id="483559FCFFDDFFF2FC92383662E6F8AB" box="[869,928,1853,1879]" pageId="1" pageNumber="12">Type</typeStatus>
and included genus
</emphasis>
:
<taxonomicName id="508E9CDDFFDDFFF2FB53383664DBF8AB" authority="Garassino &amp; Pasini, 2018" authorityName="Garassino &amp; Pasini" authorityYear="2018" box="[1188,1437,1853,1879]" class="Malacostraca" family="Amazighopsidae" genus="Amazighopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A5FA3B4CFFDDFFF2FB5338366406F8AB" box="[1188,1344,1853,1879]" italics="true" pageId="1" pageNumber="12">Amazighopsis</emphasis>
n. gen.,
</taxonomicName>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDDFFF2FC92387265ECF84D" pageId="1" pageNumber="12" type="etymology">
<paragraph id="9731E75EFFDDFFF2FC92387265ECF84D" blockId="1.[832,1474,1491,1969]" pageId="1" pageNumber="12">
<emphasis id="A5FA3B4CFFDDFFF2FC92387262A0F86F" bold="true" box="[869,998,1913,1939]" pageId="1" pageNumber="12">Etymology</emphasis>
: From Amazigh that means Berber in the original language of this people.
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDEFFF1FF603F8C61AEFF3D" pageId="2" pageNumber="13" type="description">
<paragraph id="9731E75EFFDEFFF1FF603F8C61AEFF3D" blockId="2.[113,756,135,1969]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FF603F8C602CFF5E" bold="true" box="[151,362,135,162]" pageId="2" pageNumber="13">Geological range</emphasis>
: Late Cretaceous (Cenomanian-Turonian).
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDEFFF1FF603FC864E3FD7F" pageId="2" pageNumber="13" type="discussion">
<paragraph id="9731E75EFFDEFFF1FF603FC860FBFDF7" blockId="2.[113,756,135,1969]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FF603FC86053FF21" bold="true" box="[151,277,195,221]" pageId="2" pageNumber="13">Discussion</emphasis>
. Some morphological characters of the studied specimens, such as the dorso-laterally compressed carapace, the cylindrical shape of the carapace, and the strong chelate P1 allow us to assign them tentatively to the infraorder
<taxonomicName id="508E9CDDFFDEFFF1FEE53E3760C5FEAA" authorityName="Latreille" authorityYear="1802" box="[274,387,316,342]" class="Malacostraca" higherTaxonomySource="GBIF" infraOrder="Astacidea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="infraOrder">Astacidea</taxonomicName>
, including all the marine chelate lobsters. Indeed the assignment to this infraorder is tentative because, according to
<bibRefCitation id="F31F9AAFFFDEFFF1FE5E3E7263A9FE6F" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M." box="[425,751,376,403]" pageId="2" pageNumber="13" pagination="78 - 123" refId="ref4659" refString="Karasawa H., Schweitzer C. E. &amp; Feldmann R. M. 2013 - Phylogeny and systematics of extant and extinct lobsters. Journal of Crustacean Biology, 33 (1): 78 - 123." type="journal article" year="2013">
Karasawa
<emphasis id="A5FA3B4CFFDEFFF1FDD63E72631BFE6E" box="[545,605,376,402]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
(2013: 110)
</bibRefCitation>
, some other diagnostic characters, such as the rostrum with supra- and subrostral teeth, s2 larger than s3-s6, and the uropodal exopod with diaeresis do not fit with the characters of the studied specimens.
</paragraph>
<paragraph id="9731E75EFFDEFFF1FF603D0461FCFBD5" blockId="2.[113,756,135,1969]" pageId="2" pageNumber="13">
According to
<bibRefCitation id="F31F9AAFFFDEFFF1FEC63D1B631CFDD5" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M." box="[305,602,527,554]" pageId="2" pageNumber="13" pagination="78 - 123" refId="ref4659" refString="Karasawa H., Schweitzer C. E. &amp; Feldmann R. M. 2013 - Phylogeny and systematics of extant and extinct lobsters. Journal of Crustacean Biology, 33 (1): 78 - 123." type="journal article" year="2013">
Karasawa
<emphasis id="A5FA3B4CFFDEFFF1FE513D1B6098FDD5" box="[422,478,527,553]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
(2013: 94)
</bibRefCitation>
the new classification for lobsters, after the phylogenetic systematics proposed by
<bibRefCitation id="F31F9AAFFFDEFFF1FEFF3D40636AFD9A" author="Scholtz G. &amp; Richter S." box="[264,556,587,614]" pageId="2" pageNumber="13" pagination="289 - 328" refId="ref4842" refString="Scholtz G. &amp; Richter S., 1995 - Phylogenetic systematics of the reptantian Decapoda (Crustacea, Malacostraca). Zoological Journal of the Linnean Society, 113: 289 - 328." type="journal article" year="1995">Scholtz &amp; Richter (1995)</bibRefCitation>
and
<bibRefCitation id="F31F9AAFFFDEFFF1FD9F3D4061F9FD78" author="Wahle R. A. &amp; Tshudy D. &amp; Cobb J. S. &amp; Factor J. &amp; Jaini M." pageId="2" pageNumber="13" pagination="3 - 108" refId="ref4999" refString="Wahle R. A., Tshudy D., Cobb J. S., Factor J. &amp; Jaini M., 2012 - Infraorder Astacidea Latreille, 1802 p. p.: the marine clawed lobsters. In: Treatise on Zoology, Anatomy, Taxonomy, Biology. Schram F. R. &amp; von Vaupel Klein J. C. (eds.). The Crustacea, 9 (B): 3 - 108." type="book chapter" year="2012">
Wahle
<emphasis id="A5FA3B4CFFDEFFF1FD4E3D4763B2FD99" box="[697,756,587,613]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
(2012)
</bibRefCitation>
, includes two sections for the infraorder
<taxonomicName id="508E9CDDFFDEFFF1FD723D62605CFD5E" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" class="Malacostraca" higherTaxonomySource="GBIF" infraOrder="Astacidea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="infraOrder">Astacidea Latreille, 1802</taxonomicName>
, as follows: Homarida
<bibRefCitation id="F31F9AAFFFDEFFF1FDD53D8361EFFD3D" author="Scholtz G. &amp; Richter S." pageId="2" pageNumber="13" pagination="289 - 328" refId="ref4842" refString="Scholtz G. &amp; Richter S., 1995 - Phylogenetic systematics of the reptantian Decapoda (Crustacea, Malacostraca). Zoological Journal of the Linnean Society, 113: 289 - 328." type="journal article" year="1995">Scholtz &amp; Richter, 1995</bibRefCitation>
and Astacida
<bibRefCitation id="F31F9AAFFFDEFFF1FEB93DAD6320FD3D" author="Scholtz G. &amp; Richter S." box="[334,614,678,705]" pageId="2" pageNumber="13" pagination="289 - 328" refId="ref4842" refString="Scholtz G. &amp; Richter S., 1995 - Phylogenetic systematics of the reptantian Decapoda (Crustacea, Malacostraca). Zoological Journal of the Linnean Society, 113: 289 - 328." type="journal article" year="1995">Scholtz &amp; Richter, 1995</bibRefCitation>
. The former includes three superfamilies, Enoplometopoidea de Saint Laurent, 1988 (
<taxonomicName id="508E9CDDFFDEFFF1FEEA3DE963A9FD00" authority="de Saint Laurent, 1988" authorityName="de Saint Laurent" authorityYear="1988" box="[285,751,738,765]" class="Malacostraca" family="Enoplometopidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Enoplometopidae de Saint Laurent, 1988</taxonomicName>
;
<taxonomicName id="508E9CDDFFDEFFF1FF863C0B60D6FCE6" authority="Beurlen, 1930" authorityName="Beurlen" authorityYear="1930" box="[113,400,768,794]" class="Malacostraca" family="Uncinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Uncinidae Beurlen, 1930</taxonomicName>
); Stenochiroidea Beurlen, 1928 (
<taxonomicName id="508E9CDDFFDEFFF1FF8E3C156084FCC4" authority="Beurlen, 1928" authorityName="Beurlen" authorityYear="1928" box="[121,450,798,824]" class="Malacostraca" family="Stenochiridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Stenochiridae Beurlen, 1928</taxonomicName>
); and Nephropoidea Dana, 1852 (
<taxonomicName id="508E9CDDFFDEFFF1FF403C376092FCAB" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[183,468,828,855]" class="Malacostraca" family="Nephropidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Nephropidae Dana, 1852</taxonomicName>
). The latter includes three superfamilies, Protastacoidea Albrecht, 1983 (
<taxonomicName id="508E9CDDFFDEFFF1FD8D3C506008FC6F" authority="Albrecht, 1983" authorityName="Albrecht" authorityYear="1983" class="Malacostraca" family="Protastacidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Protastacidae Albrecht, 1983</taxonomicName>
); Astacoidea Latreille, 1802 (
<taxonomicName id="508E9CDDFFDEFFF1FD563C72600DFC4D" authority=", Latreille, 1802" authorityName="Latreille" authorityYear="1802" class="Malacostraca" family="Astacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Astacidae, Latreille, 1802</taxonomicName>
;
<taxonomicName id="508E9CDDFFDEFFF1FEA13C9C6335FC4D" authority="Hobbs, 1942" authorityName="Hobbs" authorityYear="1942" box="[342,627,919,945]" class="Malacostraca" family="Cambaridae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Cambaridae Hobbs, 1942</taxonomicName>
;
<taxonomicName id="508E9CDDFFDEFFF1FD893C9C6310FC33" authority="Taylor, Schram &amp; Yan-Bin, 1999" authorityName="Taylor, Schram &amp; Yan-Bin" authorityYear="1999" class="Malacostraca" family="Cricoidoscelosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Cricoidoscelosidae Taylor, Schram &amp; Yan-Bin, 1999</taxonomicName>
); and Parastacoidea Huxley, 1879 (
<taxonomicName id="508E9CDDFFDEFFF1FE863CD863EEFC11" authority="Huxley, 1879" authorityName="Huxley" authorityYear="1879" box="[369,680,979,1005]" class="Malacostraca" family="Parastacidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Parastacidae Huxley, 1879</taxonomicName>
), truly “ …
<emphasis id="A5FA3B4CFFDEFFF1FF553CFA6366FBF7" box="[162,544,1009,1035]" italics="true" pageId="2" pageNumber="13">poorly known in the fossil record.</emphasis>
” (
<bibRefCitation id="F31F9AAFFFDEFFF1FDCB3CFA61E8FBD5" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M." pageId="2" pageNumber="13" pagination="78 - 123" refId="ref4659" refString="Karasawa H., Schweitzer C. E. &amp; Feldmann R. M. 2013 - Phylogeny and systematics of extant and extinct lobsters. Journal of Crustacean Biology, 33 (1): 78 - 123." type="journal article" year="2013">
Karasawa
<emphasis id="A5FA3B4CFFDEFFF1FD433CF963ABFBF7" box="[692,749,1009,1035]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
, 2013
</bibRefCitation>
).
</paragraph>
<paragraph id="9731E75EFFDEFFF1FF603B26603EFAC5" blockId="2.[113,756,135,1969]" pageId="2" pageNumber="13">
Though the two sections have substantial morphological differences concerning the rostrum (short or elongate with supra- or subrostral teeth), the carina and groove system of the carapace, the pleon (pleurae more or less pointed or rounded), and the telson (with or without diaeresis), they share one constant and consistent diagnostic character, the uropodal exopod with diaeresis, according to the most recent phylogenetic systematics proposed by
<bibRefCitation id="F31F9AAFFFDEFFF1FF863A146032FAC5" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M." box="[113,372,1310,1337]" pageId="2" pageNumber="13" pagination="78 - 123" refId="ref4659" refString="Karasawa H., Schweitzer C. E. &amp; Feldmann R. M. 2013 - Phylogeny and systematics of extant and extinct lobsters. Journal of Crustacean Biology, 33 (1): 78 - 123." type="journal article" year="2013">
Karasawa
<emphasis id="A5FA3B4CFFDEFFF1FF103A146059FAC4" box="[231,287,1310,1336]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
(2013)
</bibRefCitation>
.
</paragraph>
<paragraph id="9731E75EFFDEFFF1FF603A36605BF84D" blockId="2.[113,756,135,1969]" pageId="2" pageNumber="13">
Based upon the diagnoses proposed by
<bibRefCitation id="F31F9AAFFFDEFFF1FD743A366041FA89" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M." pageId="2" pageNumber="13" pagination="78 - 123" refId="ref4659" refString="Karasawa H., Schweitzer C. E. &amp; Feldmann R. M. 2013 - Phylogeny and systematics of extant and extinct lobsters. Journal of Crustacean Biology, 33 (1): 78 - 123." type="journal article" year="2013">
Karasawa
<emphasis id="A5FA3B4CFFDEFFF1FF863A5761E9FA89" box="[113,175,1371,1397]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
(2013)
</bibRefCitation>
for each family within the two sections, we point out that some morphological characters of the studied specimens, such as the strongly tuberculate carapace and P1; a short rostrum with serrate suprarostral margins, bearing 3-4 pairs of teeth; a deep cervical groove and a weak antennal groove not forming the characteristic “W” shape; s2 with subrounded pleura partially overlapping those of s1 and s3; smooth s1, s3- s6 with rounded pleurae; a subtrapezoidal telson without diaeresis and lateral margins with one median movable spine and two distal movable spines; chelate P1 slightly heterochelous (peculiar shape of the P1 chela: P1 index occlusal margin with a row of strong rounded teeth inclined forward and arranged randomly; P1 dactylus occlusal margin with a row of pointed, elongate, and slender teeth strongly inclined distally); and mainly the uropodal exopod without diaeresis, are so unique and peculiar that the studied specimens do not fit any diagnosis for known clawed lobster families, provided by the authors.
</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3F8C65B3FF22" blockId="2.[903,1474,135,644]" pageId="2" pageNumber="13">
Therefore, these differences warrant a distinct family within the
<taxonomicName id="508E9CDDFFDEFFF1FBAB3FAE658CFF43" authorityName="Latreille" authorityYear="1802" box="[1116,1226,165,191]" class="Malacostraca" higherTaxonomySource="GBIF" infraOrder="Astacidea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="18" phylum="Arthropoda" rank="infraOrder">Astacidea</taxonomicName>
to accommodate the
<typeStatus id="483559FCFFDEFFF1FC703FCF62F1FF22" box="[903,951,196,222]" pageId="2" pageNumber="13">type</typeStatus>
genus
<taxonomicName id="508E9CDDFFDEFFF1FBF13FC865E4FF21" authority="Garassino &amp; Pasini, 2018" authorityName="Garassino &amp; Pasini" authorityYear="2018" box="[1030,1186,195,221]" class="Malacostraca" family="Amazighopsidae" genus="Amazighopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A5FA3B4CFFDEFFF1FBF13FC865E4FF21" box="[1030,1186,195,221]" italics="true" pageId="2" pageNumber="13">Amazighopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEC98637FFDEFFF1FB5E3FCF65B3FF22" box="[1193,1269,196,222]" pageId="2" pageNumber="13" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3FE964E3FD7F" blockId="2.[903,1474,135,644]" pageId="2" pageNumber="13">
The new family cannot be assigned to both sections of the infraorder
<taxonomicName id="508E9CDDFFDEFFF1FB623E0B6445FEE6" authorityName="Latreille" authorityYear="1802" box="[1173,1283,256,282]" class="Malacostraca" higherTaxonomySource="GBIF" infraOrder="Astacidea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="infraOrder">Astacidea</taxonomicName>
based upon the characters proposed by
<bibRefCitation id="F31F9AAFFFDEFFF1FB783E1564F8FEC4" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M." box="[1167,1470,286,312]" pageId="2" pageNumber="13" pagination="78 - 123" refId="ref4659" refString="Karasawa H., Schweitzer C. E. &amp; Feldmann R. M. 2013 - Phylogeny and systematics of extant and extinct lobsters. Journal of Crustacean Biology, 33 (1): 78 - 123." type="journal article" year="2013">
Karasawa
<emphasis id="A5FA3B4CFFDEFFF1FAF33E14647AFEC4" box="[1284,1340,286,312]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
(2013: 110
</bibRefCitation>
, 113). According to
<bibRefCitation id="F31F9AAFFFDEFFF1FB963E376418FEAA" author="Holthuis L. B." box="[1121,1374,316,342]" pageId="2" pageNumber="13" refId="ref4628" refString="Holthuis L. B., 1991 - FAO species catalogue. 13. Marine lobsters of the world. Food and Agriculture Organization of the United Nations, Rome." type="book" year="1991">Holthuis (1991: 31-47</bibRefCitation>
; fig. 61), the new family may be, however, tentatively assigned to the section Homarida based upon
<taxonomicName id="508E9CDDFFDEFFF1FA893E7365FFFE4C" authority="Wood-Mason, 1873" authorityName="Wood-Mason" authorityYear="1873" class="Malacostraca" family="Nephropidae" genus="Nephropsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="A5FA3B4CFFDEFFF1FA893E73628CFE4C" italics="true" pageId="2" pageNumber="13">Nephropsis</emphasis>
Wood-Mason, 1873
</taxonomicName>
, the sole genus within this section, including the extant species with or without a diaeresis on the uropodal exopod. Moreover, the lack of the diaeresis on the telson in the new family, a peculiar character of the Homarida, as proposed by
<bibRefCitation id="F31F9AAFFFDEFFF1FBE13D25640FFDB4" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M." box="[1046,1353,557,584]" pageId="2" pageNumber="13" pagination="78 - 123" refId="ref4659" refString="Karasawa H., Schweitzer C. E. &amp; Feldmann R. M. 2013 - Phylogeny and systematics of extant and extinct lobsters. Journal of Crustacean Biology, 33 (1): 78 - 123." type="journal article" year="2013">
Karasawa
<emphasis id="A5FA3B4CFFDEFFF1FB7D3D256584FDBB" box="[1162,1218,557,583]" italics="true" pageId="2" pageNumber="13">et al.</emphasis>
(2013: 116)
</bibRefCitation>
(vs. telson with diaeresis in Astacida), would further support the assignment of the new family to this section.
</paragraph>
</subSubSection>
</treatment>
</document>

152
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<subSubSection id="DF94B4D5FFDEFFF1FC263C9C6431FC4D" box="[977,1399,919,945]" pageId="2" pageNumber="13" type="nomenclature">
<paragraph id="9731E75EFFDEFFF1FC263C9C6431FC4D" blockId="2.[977,1399,919,975]" box="[977,1399,919,945]" pageId="2" pageNumber="13">
<heading id="CC795032FFDEFFF1FC263C9C6431FC4D" box="[977,1399,919,945]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="13" reason="3">
<taxonomicName id="508E9CDDFFDEFFF1FC263C9C6431FC4D" authority="Garassino &amp; Pasini, 2018" authorityName="Garassino &amp; Pasini" authorityYear="2018" box="[977,1399,919,945]" class="Malacostraca" family="Amazighopsidae" genus="Amazighopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="cretacica" status="n. gen., n. sp.">
<emphasis id="A5FA3B4CFFDEFFF1FC263C9C659DFC4C" box="[977,1243,919,945]" italics="true" pageId="2" pageNumber="13">Amazighopsis cretacica</emphasis>
n. gen., n. sp.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDEFFF1FB853CBE6590FC33" box="[1138,1238,949,975]" pageId="2" pageNumber="13" type="description">
<paragraph id="9731E75EFFDEFFF1FB853CBE6590FC33" blockId="2.[977,1399,919,975]" box="[1138,1238,949,975]" pageId="2" pageNumber="13">
<figureCitation id="0FB5FBDBFFDEFFF1FB853CBE6590FC33" box="[1138,1238,949,975]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="3.[189,222,1918,1940]" captionStartId-1="4.[189,222,1919,1941]" captionStartId-2="5.[189,222,1917,1939]" captionStartId-3="6.[113,146,796,818]" captionStartId-4="6.[113,146,1840,1862]" captionStartId-5="6.[831,864,1916,1938]" captionTargetBox-0="[189,1399,135,1856]" captionTargetBox-1="[189,1398,442,1894]" captionTargetBox-2="[189,1398,387,1893]" captionTargetBox-3="[113,783,136,787]" captionTargetBox-4="[113,771,1066,1834]" captionTargetBox-5="[843,1469,136,1888]" captionTargetId-1="figure-165@4.[186,1400,438,1234]" captionTargetId-2="figure-97@5.[189,1398,387,1193]" captionTargetId-3="figure-36@6.[110,758,136,772]" captionTargetId-5="figure-117@6.[831,1469,136,1584]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="6" captionText-0="Fig. 1 - Amazighopsis cretacica n. gen., n. sp., MSNM i27545, Holotype. A) General view in natural light (x 1.51). B) General view in UV light (x 1.44)." captionText-1="Fig. 2 - Amazighopsis cretacica n. gen., n. sp., MSNM i27546, Paratype. A) General view in natural light (x 1.23). B) General view in UV light (x 1.12)." captionText-2="Fig. 3 - Amazighopsis cretacica n. gen., n. sp., MSNM i27646, Paratype. A) General view in natural light (x 2). B) General view in UV light (x 1.71)." captionText-3="Fig. 4 - Amazighopsis cretacica n. gen., n. sp., MSNM i27545, close-up of the telson without diaeresis and with movable spines on lateral margin (arrows) and uropodal exopod without diaeresis (x 6.81)." captionText-4="Fig. 5 - Amazighopsis cretacica n. gen., n. sp., MSNM i27546. A) Close-up of the carapace with the simple system grooves [cervical, (ee 1) and antennal (b) grooves] and short rostrum with serrate suprarostral margin (x 1.36). B) Close-up of P1 chela with the occlusal margins of dactylus and index (x 1.78)." captionText-5="Fig. 6 - Amazighopsis cretacica n. gen., n. sp. A) reconstruction. B) close-up of the occlusal margins of P1 chela." figureDoi-0="http://doi.org/10.5281/zenodo.12523823" figureDoi-1="http://doi.org/10.5281/zenodo.12523825" figureDoi-2="http://doi.org/10.5281/zenodo.12523827" figureDoi-3="http://doi.org/10.5281/zenodo.12523829" figureDoi-4="http://doi.org/10.5281/zenodo.13891953" figureDoi-5="http://doi.org/10.5281/zenodo.12523831" httpUri-0="https://zenodo.org/record/12523823/files/figure.png" httpUri-1="https://zenodo.org/record/12523825/files/figure.png" httpUri-2="https://zenodo.org/record/12523827/files/figure.png" httpUri-3="https://zenodo.org/record/12523829/files/figure.png" httpUri-4="https://zenodo.org/record/13891953/files/figure.png" httpUri-5="https://zenodo.org/record/12523831/files/figure.png" pageId="2" pageNumber="13">Figs. 1-6</figureCitation>
</paragraph>
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<subSubSection id="DF94B4D5FFDEFFF1FC5B3CFA6585FBF7" box="[940,1219,1009,1036]" pageId="2" pageNumber="13" type="diagnosis">
<paragraph id="9731E75EFFDEFFF1FC5B3CFA6585FBF7" blockId="2.[903,1475,1009,1969]" box="[940,1219,1009,1036]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FC5B3CFA6559FBF7" bold="true" box="[940,1055,1009,1035]" pageId="2" pageNumber="13">Diagnosis</emphasis>
: as for family.
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDEFFF1FC5B3B04628FFB9A" pageId="2" pageNumber="13" type="etymology">
<paragraph id="9731E75EFFDEFFF1FC5B3B04628FFB9A" blockId="2.[903,1475,1009,1969]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FC5B3B04656BFBD5" bold="true" box="[940,1069,1039,1065]" pageId="2" pageNumber="13">Etymology</emphasis>
: the trivial name alludes to the Cretaceous, the geological age of the studied specimens.
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDEFFF1FC5B3B6162F5FA33" pageId="2" pageNumber="13" type="materials_examined">
<paragraph id="9731E75EFFDEFFF1FC5B3B61659CFB78" blockId="2.[903,1475,1009,1969]" box="[940,1242,1129,1156]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FC5B3B61655FFB78" bold="true" box="[940,1049,1130,1156]" pageId="2" pageNumber="13">
<typeStatus id="483559FCFFDEFFF1FC5B3B61655FFB78" box="[940,1049,1130,1156]" pageId="2" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
:
<materialsCitation id="27E6ED03FFDEFFF1FBDF3B616590FB78" box="[1064,1238,1129,1156]" collectionCode="MSNM" location="Kem Kem" municipality="Gara Sbaa" pageId="2" pageNumber="13" specimenCode="MSNM i27545" specimenCount="1" type="holotype">
<specimenCode id="C7284F25FFDEFFF1FBDF3B616590FB78" box="[1064,1238,1129,1156]" pageId="2" pageNumber="13">MSNM i27545</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3B8364DBFB5D" blockId="2.[903,1475,1009,1969]" box="[940,1437,1159,1186]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FC5B3B836562FB5E" bold="true" box="[940,1060,1160,1186]" pageId="2" pageNumber="13">
<typeStatus id="483559FCFFDEFFF1FC5B3B836562FB5E" box="[940,1060,1160,1186]" pageId="2" pageNumber="13" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<materialsCitation id="27E6ED03FFDEFFF1FBC43B8365A2FB5E" box="[1075,1252,1159,1186]" collectionCode="MSNM" location="Kem Kem" municipality="Gara Sbaa" pageId="2" pageNumber="13" specimenCode="MSNM i27546" specimenCount="1" type="paratype">
<specimenCode id="C7284F25FFDEFFF1FBC43B8365A6FB5E" box="[1075,1248,1159,1186]" pageId="2" pageNumber="13">MSNM i27546</specimenCode>
,
</materialsCitation>
<materialsCitation id="27E6ED03FFDEFFF1FB1C3B8364DFFB5D" box="[1259,1433,1159,1186]" collectionCode="MSNM" location="Kem Kem" municipality="Gara Sbaa" pageId="2" pageNumber="13" specimenCode="MSNM i27646" specimenCount="1" type="paratype">
<specimenCode id="C7284F25FFDEFFF1FB1C3B8364DFFB5D" box="[1259,1433,1159,1186]" pageId="2" pageNumber="13">MSNM i27646</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3BAD641BFB3C" blockId="2.[903,1475,1009,1969]" box="[940,1373,1189,1216]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FC5B3BAD6503FB43" bold="true" box="[940,1093,1189,1216]" pageId="2" pageNumber="13">Type locality</emphasis>
: Gara Sbaa (Kem Kem).
</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3BCF65C8FAE7" blockId="2.[903,1475,1009,1969]" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FC5B3BCF6443FB22" bold="true" box="[940,1285,1220,1246]" pageId="2" pageNumber="13">Material and measurements</emphasis>
: three complete specimens, one of these (MSNM i27546) preserved as part and counterpart.
</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3A156415FAAB" blockId="2.[903,1475,1009,1969]" pageId="2" pageNumber="13">
MSNM i27545 – tl:
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; lcpx:
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; wcpx:
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; lp:
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; wp:
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; ldi:
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</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3A506568FA6F" blockId="2.[903,1475,1009,1969]" pageId="2" pageNumber="13">
MSNM i27546 – tl:
<quantity id="50764ABBFFDEFFF1FB6F3A5165AAFA89" box="[1176,1260,1370,1397]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="13" unit="mm" value="65.0">65 mm</quantity>
; lp:
<quantity id="50764ABBFFDEFFF1FAD73A506432FA89" box="[1312,1396,1371,1397]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="13" unit="mm" value="10.0">10 mm</quantity>
; wp:
<quantity id="50764ABBFFDEFFF1FA433A5062F0FA6F" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="13" unit="mm" value="7.0">7 mm</quantity>
; ldi:
<quantity id="50764ABBFFDEFFF1FC1A3A726568FA6F" box="[1005,1070,1401,1427]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="13" unit="mm" value="9.0">9 mm</quantity>
</paragraph>
<paragraph id="9731E75EFFDEFFF1FC5B3A9C62F5FA33" blockId="2.[903,1475,1009,1969]" pageId="2" pageNumber="13">
MSNM i27646 – lp:
<quantity id="50764ABBFFDEFFF1FB683A9C65B5FA4D" box="[1183,1267,1431,1457]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="13" unit="mm" value="10.0">10 mm</quantity>
; wp:
<quantity id="50764ABBFFDEFFF1FAC43A9C643FFA4D" box="[1331,1401,1431,1457]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="13" unit="mm" value="7.0">7 mm</quantity>
; ldi:
<quantity id="50764ABBFFDEFFF1FA433A9C62F5FA33" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="13" unit="mm" value="9.0">9 mm</quantity>
</paragraph>
</subSubSection>
<subSubSection id="DF94B4D5FFDEFFF6FC5B3AD8658CFEAA" lastPageId="5" lastPageNumber="16" pageId="2" pageNumber="13" type="description">
<paragraph id="9731E75EFFDEFFF6FC5B3AD8658CFEAA" blockId="2.[903,1475,1009,1969]" lastBlockId="5.[831,1474,135,342]" lastPageId="5" lastPageNumber="16" pageId="2" pageNumber="13">
<emphasis id="A5FA3B4CFFDEFFF1FC5B3AD86570FA11" bold="true" box="[940,1078,1491,1517]" pageId="2" pageNumber="13">Description</emphasis>
. Carapace – Subcylindrical carapace strongly granular, laterally compressed; dorsal margin slightly convex; posterior margin nearly straight; triangular, short rostrum with serrate suprarostral margin, bearing 4-5 pairs of teeth; dorsal rostral surface concave without median carina; subrostral margin spineless; deep cervical groove nearly straight, strongly inclined forward, reaching the dorsal margin at the anterior third of the total carapace length (excluding rostrum), making an angle of about 45° to the dorsal margin; antennal groove short and weak; cephalic region shorter than branchial one; branchiocardiac, postcervical, and hepatic grooves absent; carapace surface without accessory spines. Pleon – s1 short, narrow; s2 with subrounded pleura partially overlapping those of s1 and s3; convex pleurae, slightly rounded ventrally, ending in a short point; tergites and pleurae of s1-s6 smooth; subtrapezoidal telson without diaeresis, and lateral margins with one median movable spine and two distal movable spines. Cephalic appendages – Cephalic appendages poorly preserved, flagella of antennulae and antennae partially preserved; antennae flagella stronger than those of the antennulae. Thoracic appendages – mpx3 and P2-P5 not preserved; P1 slightly heterochelous, with ischium, merus, carpus, and propodus strongly granulate; right P1 with squared ischium, subrectangular elongate merus, convex and serrate dorsally; stout subtriangular carpus with serrate dorsal margin; elongate subrectangular chela serrate dorsally, outer surface with two parallel longitudinal rows of strongly aligned tuberculate spines; right P1 chela with elongate dactylus and index, equal in length and curved distally; index slightly stronger than dactylus; index occlusal margin with a row of strong, rounded molariform teeth inclined distally and arranged randomly; dactylus occlusal margin, with a row of pointed, elongate, and slender teeth strongly inclined distally; left P1 propodus, merus, and carpus similar in shape and ornamentation of right P1; left chela poorly preserved, looks slightly stouter than the right with stronger index and dactylus, occlusal margins bearing a rim of similar rounded molariform teeth. Pleonal appendages – Pleopods poorly preserved; uropodal endopod and exopod as long as telson; uropodal endopod and exopod rounded posteriorly with one thin longitudinal median carina and spiny margins; uropodal exopod without diaeresis.
</paragraph>
</subSubSection>
</treatment>
</document>