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<figureCitation id="1319D388FFA54E5FFD45FDA0FCEF3AD1" box="[684,792,636,662]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="3.[136,229,1877,1899]" captionStartId-1="4.[136,229,1411,1433]" captionStartId-2="6.[136,229,1868,1890]" captionStartId-3="7.[136,229,2028,2050]" captionStartId-4="8.[136,229,633,655]" captionTargetBox-0="[227,1361,190,1853]" captionTargetBox-1="[227,1361,190,1386]" captionTargetBox-2="[231,1355,190,1844]" captionTargetBox-3="[201,1386,190,2004]" captionTargetBox-4="[227,1361,190,609]" captionTargetId-0="figure-19@3.[227,1361,190,1853]" captionTargetId-1="figure-20@4.[227,1361,190,1386]" captionTargetId-2="figure-20@6.[231,1355,190,1844]" captionTargetId-3="figure-19@7.[201,1386,190,2004]" captionTargetId-4="figure-20@8.[227,1361,190,609]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 2. Leopoldia oztasii; a: habit, b: inflorescence, c: infructescence." captionText-1="FIGURE 3. The habit of Leopoldia oztasii with its underground part in natural habitat (note: the size of the bulb is considerable)." captionText-2="FIGURE 4. Comparison of Leopoldia oztasii and L. tanuiflora; ac: L. oztasii, df: L. tenuiflora, (ad: habit, be: inflorescence, cf: infructescence) (Photo “b” was taken by Haydar Öztaş, photo “d” was taken by Abdurrahman Sefalı and photo “e” was taken by Hasan Şişman)." captionText-3="FIGURE 5. Comparison of infructescences and capsules of two closely related species; A1A2: L. oztasii, B1B2: L. tenuiflora." captionText-4="FIGURE 6. SEM photographs of seeds of two closely related species; ad: L. oztasii, eh: L. tenuiflora (a and e general view; bd and fh epidermal cells)." pageId="2" pageNumber="19">Figs. 26</figureCitation>
<figureCitation id="1319D388FFA54E5FFD45FDA0FCEF3AD1" box="[684,792,636,662]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="3.[136,229,1877,1899]" captionStartId-1="4.[136,229,1411,1433]" captionStartId-2="6.[136,229,1868,1890]" captionStartId-3="7.[136,229,2028,2050]" captionStartId-4="8.[136,229,633,655]" captionTargetBox-0="[227,1361,190,1853]" captionTargetBox-1="[227,1361,190,1386]" captionTargetBox-2="[231,1355,190,1844]" captionTargetBox-3="[201,1386,190,2004]" captionTargetBox-4="[227,1361,190,609]" captionTargetId-0="figure-19@3.[227,1361,190,1853]" captionTargetId-1="figure-20@4.[227,1361,190,1386]" captionTargetId-2="figure-20@6.[231,1355,190,1844]" captionTargetId-3="figure-19@7.[201,1386,190,2004]" captionTargetId-4="figure-20@8.[227,1361,190,609]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 2. Leopoldia oztasii; a: habit, b: inflorescence, c: infructescence." captionText-1="FIGURE 3. The habit of Leopoldia oztasii with its underground part in natural habitat (note: the size of the bulb is considerable)." captionText-2="FIGURE 4. Comparison of Leopoldia oztasii and L. tanuiflora; ac: L. oztasii, df: L. tenuiflora, (ad: habit, be: inflorescence, cf: infructescence) (Photo “b” was taken by Haydar Öztaş, photo “d” was taken by Abdurrahman Sefalı and photo “e” was taken by Hasan Şişman)." captionText-3="FIGURE 5. Comparison of infructescences and capsules of two closely related species; A1A2: L. oztasii, B1B2: L. tenuiflora." captionText-4="FIGURE 6. SEM photographs of seeds of two closely related species; ad: L. oztasii, eh: L. tenuiflora (a and e general view; bd and fh epidermal cells)." figureDoi-0="http://doi.org/10.5281/zenodo.13879295" figureDoi-1="http://doi.org/10.5281/zenodo.13879297" figureDoi-2="http://doi.org/10.5281/zenodo.13879299" figureDoi-3="http://doi.org/10.5281/zenodo.13879301" figureDoi-4="http://doi.org/10.5281/zenodo.13879303" httpUri-0="https://zenodo.org/record/13879295/files/figure.png" httpUri-1="https://zenodo.org/record/13879297/files/figure.png" httpUri-2="https://zenodo.org/record/13879299/files/figure.png" httpUri-3="https://zenodo.org/record/13879301/files/figure.png" httpUri-4="https://zenodo.org/record/13879303/files/figure.png" pageId="2" pageNumber="19">Figs. 26</figureCitation>
,
<tableCitation id="C6A0FAB6FFA54E5FFCCAFDA0FC683AD1" box="[803,927,636,662]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="5.[136,224,160,184]" captionStartId-1="5.[136,224,1698,1722]" captionText-0="TABLE 1. Comparison of the diagnostic characteristics between L. oztasii and L. tenuiflora." captionText-1="TABLE 2. Seed features of Leopoldia oztasii and L. tenuiflora." pageId="2" pageNumber="19">Tables 12</tableCitation>
).
@ -288,13 +289,13 @@ is distributed in
in
<collectingCountry id="F3358F9DFFA54E5FFB28F968FAED3E89" box="[1217,1306,1716,1742]" name="Turkey" pageId="2" pageNumber="19">Türkiye</collectingCountry>
(
<figureCitation id="1319D388FFA54E5FFAC1F968FA9E3E89" box="[1320,1385,1716,1742]" captionStart="FIGURE 1" captionStartId="1.[136,229,1720,1742]" captionTargetBox="[170,1417,826,1677]" captionTargetId="figure-247@1.[151,1436,784,1693]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Leopoldia oztasii and L. tenuiflora: red circle = L. oztasii; blue squares = L. tenuiflora (The map was prepared and colored by Büşra ARIK)." pageId="2" pageNumber="19">Fig. 1</figureCitation>
<figureCitation id="1319D388FFA54E5FFAC1F968FA9E3E89" box="[1320,1385,1716,1742]" captionStart="FIGURE 1" captionStartId="1.[136,229,1720,1742]" captionTargetBox="[170,1417,826,1677]" captionTargetId="figure-247@1.[151,1436,784,1693]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Leopoldia oztasii and L. tenuiflora: red circle = L. oztasii; blue squares = L. tenuiflora (The map was prepared and colored by Büşra ARIK)." figureDoi="http://doi.org/10.5281/zenodo.13879293" httpUri="https://zenodo.org/record/13879293/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
). The habitat of the
<taxonomicName id="4C22B48EFFA54E5FFECCF905FE7B3EB5" authorityName="Eker, Eroglu &amp; Pinar" authorityYear="2024" box="[293,396,1753,1778]" class="Liliopsida" family="Asparagaceae" genus="Leopoldia" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="19" phylum="Tracheophyta" rank="species" species="oztasii">
<emphasis id="B956131FFFA54E5FFECCF905FE7B3EB5" box="[293,396,1753,1778]" italics="true" pageId="2" pageNumber="19">L. oztasii</emphasis>
</taxonomicName>
is stony steppes (
<figureCitation id="1319D388FFA54E5FFDBCF904FD6D3EB5" box="[597,666,1752,1778]" captionStart="FIGURE 3" captionStartId="4.[136,229,1411,1433]" captionTargetBox="[227,1361,190,1386]" captionTargetId="figure-20@4.[227,1361,190,1386]" captionTargetPageId="4" captionText="FIGURE 3. The habit of Leopoldia oztasii with its underground part in natural habitat (note: the size of the bulb is considerable)." pageId="2" pageNumber="19">Fig. 3</figureCitation>
<figureCitation id="1319D388FFA54E5FFDBCF904FD6D3EB5" box="[597,666,1752,1778]" captionStart="FIGURE 3" captionStartId="4.[136,229,1411,1433]" captionTargetBox="[227,1361,190,1386]" captionTargetId="figure-20@4.[227,1361,190,1386]" captionTargetPageId="4" captionText="FIGURE 3. The habit of Leopoldia oztasii with its underground part in natural habitat (note: the size of the bulb is considerable)." figureDoi="http://doi.org/10.5281/zenodo.13879297" httpUri="https://zenodo.org/record/13879297/files/figure.png" pageId="2" pageNumber="19">Fig. 3</figureCitation>
). The flowering season of the species is late May and early June, and fruiting season is late June.
</paragraph>
<paragraph id="8B9DCF0DFFA54E5FFF54F8FCFE0C3F19" blockId="2.[136,1452,780,1994]" pageId="2" pageNumber="19">

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@ -0,0 +1,292 @@
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<mods:title id="E3F76AA29C35B445958EEBA98CB252BD">Didymoplexiella laosensis (Orchidaceae: Epidendroideae: Gastrodieae), a new holomycotrophic orchid from Lao PDR</mods:title>
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<paragraph id="8BB036DCFFA5D730FF3CF843FC5CF80C" blockId="1.[136,906,1940,1967]" box="[136,906,1940,1967]" pageId="1" pageNumber="2">
<heading id="D0F881B0FFA5D730FF3CF843FC5CF80C" box="[136,906,1940,1967]" fontSize="11" level="2" pageId="1" pageNumber="2" reason="3">
<taxonomicName id="4C0F4D5FFFA5D730FF3CF843FD41F80D" ID-CoL="9V6PT" authority="S.W.Gale &amp; Kumar" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[136,663,1940,1967]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="B97BEACEFFA5D730FF3CF843FE65F80D" bold="true" box="[136,435,1940,1967]" italics="true" pageId="1" pageNumber="2">Didymoplexiella laosensis</emphasis>
S.W.Gale &amp; Kumar
</taxonomicName>
<emphasis id="B97BEACEFFA5D730FD29F842FD24F80C" box="[669,754,1941,1966]" italics="true" pageId="1" pageNumber="2">
<taxonomicNameLabel id="A24857B5FFA5D730FD29F842FD24F80C" box="[669,754,1941,1966]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13342A59FFA5D730FCB5F843FC98F80C" box="[769,846,1940,1966]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[216,1373,197,1919]" captionTargetId="figure-29@2.[203,1385,187,1925]" captionTargetPageId="2" captionText="FIGURE 1. Didymoplexiella laosensis. A. Plants in habitat. B. Tuber and roots with orange nodules. C. Inflorescence. D. Side view of flower. E. Front view of flower. F. Fused dorsal sepal and petals. G. Fused lateral sepals. H. Labellum. I. Side view of column. J. Front view of column. K. Operculum. L. Pollinarium (one pollinium missing on either side, cf. Fig. 2-K). M. Gynostemium with ovary, pedicel and pollinia." figureDoi="http://doi.org/10.5281/zenodo.13879375" httpUri="https://zenodo.org/record/13879375/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
&amp;
<figureCitation id="13342A59FFA5D730FCC6F843FC57F80C" box="[882,897,1940,1966]" captionStart="FIGURE 2" captionStartId="3.[136,229,1953,1975]" captionTargetBox="[229,1359,199,1888]" captionTargetId="figure-18@3.[203,1385,190,1928]" captionTargetPageId="3" captionText="FIGURE 2. Didymoplexiella laosensis. A. Inflorescence. B. Tuber and roots. C. Fused dorsal sepal and petals. D. Fused lateral sepals. E. Labellum. F. Gynostemium with ovary and pedicel. G. Front view of column. H. Side view of column. I. Oblique view of column. J. Operculum. K. Pollinarium. (Drawn by Pankaj Kumar from specimen HNL-KFBG 0007.)" figureDoi="http://doi.org/10.5281/zenodo.13879377" httpUri="https://zenodo.org/record/13879377/files/figure.png" pageId="1" pageNumber="2">2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3156557FFA5D730FF3CF86BFCB7F857" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BB036DCFFA5D730FF3CF86BFCB7F857" blockId="1.[136,1451,1980,2038]" pageId="1" pageNumber="2">
Type:―
<collectingCountry id="F318764CFFA5D730FF6CF86BFECFF873" box="[216,281,1980,2001]" name="Laos" pageId="1" pageNumber="2">LAOS</collectingCountry>
.
<collectingRegion id="49CBF83EFFA5D730FE96F86BFE08F870" box="[290,478,1980,2002]" country="Laos" name="Viangchan" pageId="1" pageNumber="2">Vientiane Province</collectingRegion>
: Vang Vieng District, Phone Ngueng Area,
<quantity id="4CF79B39FFA5D730FC32F86BFC12F873" box="[902,964,1980,2001]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="2" unit="m" value="280.0">280 m</quantity>
elevation,
<date id="FFB1101CFFA5D730FB84F86BFB6CF873" box="[1072,1210,1980,2002]" pageId="1" pageNumber="2" value="2012-04-26">26 April 2012</date>
,
<emphasis id="B97BEACEFFA5D730FB71F86BFDCFF857" italics="true" pageId="1" pageNumber="2">Gale, Kumar, Santainsy and Phunthavong HNL-KFBG 0007</emphasis>
(
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!,
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<collectionCode id="ED1EAE19FFA5D730FCA5F837FC81F857" box="[785,855,2016,2037]" country="China" httpUri="http://grbio.org/cool/300g-kc4n" name="Kadoorie Farm and Botanic Garden" pageId="1" pageNumber="2" type="Herbarium">KFBG</collectionCode>
!)
</paragraph>
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<paragraph id="8BB036DCFFA6D733FF3CF876FED7F781" blockId="2.[136,1451,1953,2083]" pageId="2" pageNumber="3">
<emphasis id="B97BEACEFFA6D733FF3CF876FF28F814" bold="true" box="[136,254,1953,1975]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
<taxonomicName id="4C0F4D5FFFA6D733FEB2F876FDD5F814" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[262,515,1953,1974]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="laosensis">
<emphasis id="B97BEACEFFA6D733FEB2F876FDD5F814" box="[262,515,1953,1974]" italics="true" pageId="2" pageNumber="3">Didymoplexiella laosensis</emphasis>
</taxonomicName>
. A. Plants in habitat. B. Tuber and roots with orange nodules. C. Inflorescence. D. Side view of flower. E. Front view of flower. F. Fused dorsal sepal and petals. G. Fused lateral sepals. H. Labellum. I. Side view of column. J. Front view of column. K. Operculum. L. Pollinarium (one pollinium missing on either side,
<emphasis id="B97BEACEFFA6D733FC77F83EFC0AF85C" box="[963,988,2025,2046]" italics="true" pageId="2" pageNumber="3">cf.</emphasis>
Fig. 2-K). M. Gynostemium with ovary, pedicel and pollinia.
</paragraph>
</caption>
<caption id="DF706654FFA7D732FF3CF876FC31F85C" ID-DOI="http://doi.org/10.5281/zenodo.13879377" ID-Zenodo-Dep="13879377" httpUri="https://zenodo.org/record/13879377/files/figure.png" pageId="3" pageNumber="4" startId="3.[136,229,1953,1975]" targetBox="[229,1359,199,1888]" targetPageId="3" targetType="figure">
<paragraph id="8BB036DCFFA7D732FF3CF876FC31F85C" blockId="3.[136,1452,1953,2047]" pageId="3" pageNumber="4">
<emphasis id="B97BEACEFFA7D732FF3CF876FF29F814" bold="true" box="[136,255,1953,1975]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
<taxonomicName id="4C0F4D5FFFA7D732FEB2F876FDD2F814" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[262,516,1953,1974]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="laosensis">
<emphasis id="B97BEACEFFA7D732FEB2F876FDD2F814" box="[262,516,1953,1974]" italics="true" pageId="3" pageNumber="4">Didymoplexiella laosensis</emphasis>
</taxonomicName>
. A. Inflorescence. B. Tuber and roots. C. Fused dorsal sepal and petals. D. Fused lateral sepals. E. Labellum. F. Gynostemium with ovary and pedicel. G. Front view of column. H. Side view of column. I. Oblique view of column. J. Operculum. K. Pollinarium. (Drawn by Pankaj Kumar from specimen
<emphasis id="B97BEACEFFA7D732FC99F83EFC0FF85C" box="[813,985,2025,2046]" italics="true" pageId="3" pageNumber="4">HNL-KFBG 0007</emphasis>
.)
</paragraph>
</caption>
<subSubSection id="C3156557FFA0D735FF3CFF49FA44FB0A" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BB036DCFFA0D735FF3CFF49FC34FE87" blockId="4.[136,1452,158,1841]" pageId="4" pageNumber="5">
<emphasis id="B97BEACEFFA0D735FF3CFF49FEF7FF1A" bold="true" box="[136,289,158,184]" pageId="4" pageNumber="5">Diagnosis:—</emphasis>
<taxonomicName id="4C0F4D5FFFA0D735FE95FF49FD9FFF1A" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[289,585,158,184]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="laosensis">
<emphasis id="B97BEACEFFA0D735FE95FF49FD9FFF1A" box="[289,585,158,184]" italics="true" pageId="4" pageNumber="5">Didymoplexiella laosensis</emphasis>
</taxonomicName>
is similar in morphology and colouration to
<taxonomicName id="4C0F4D5FFFA0D735FBFDFF48FB61FF1A" authorityName="Ridley" authorityYear="1891" box="[1097,1207,159,184]" class="Liliopsida" family="Orchidaceae" genus="Leucolena" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="ornata">
<emphasis id="B97BEACEFFA0D735FBFDFF48FB61FF1A" box="[1097,1207,159,184]" italics="true" pageId="4" pageNumber="5">D. ornata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0F4D5FFFA0D735FB72FF48FA81FF1A" authorityName="Seidenfaden" authorityYear="1972" baseAuthorityName="Rolfe ex Downie" baseAuthorityYear="1925" box="[1222,1367,159,184]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="siamensis">
<emphasis id="B97BEACEFFA0D735FB72FF48FA81FF1A" box="[1222,1367,159,184]" italics="true" pageId="4" pageNumber="5">D. siamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D5FFFA0D735FA24FF48FF2BFF7E" authorityName="Smith" authorityYear="1920" class="Liliopsida" family="Orchidaceae" genus="Didymoplexis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="trichechus">
<emphasis id="B97BEACEFFA0D735FA24FF48FF2BFF7E" italics="true" pageId="4" pageNumber="5">D. trichechus</emphasis>
</taxonomicName>
. However, the lateral sepals of
<taxonomicName id="4C0F4D5FFFA0D735FDC4FF14FD2AFF7E" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[624,764,194,220]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="laosensis">
<emphasis id="B97BEACEFFA0D735FDC4FF14FD2AFF7E" box="[624,764,194,220]" italics="true" pageId="4" pageNumber="5">D. laosensis</emphasis>
</taxonomicName>
have a rounded apex; the fleshy, longitudinal callus on the labellum terminates more than
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before the apex of the midlobe; and the projecting labellum side lobes exceed the midlobe and are truncate-rounded and irregularly denticulate at the apex.
</paragraph>
<paragraph id="8BB036DCFFA0D735FF09FEF9FEEDFBBA" blockId="4.[136,1452,158,1841]" pageId="4" pageNumber="5">
Leafless,holomycotrophic terrestrial herb,
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tall.Rhizome horizontal or ascending, tuberous,irregularly fusiform, 5.87.5 long,
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in diameter, slightly fleshy, several-noded, beige. Roots 47, arising from base of subterranean stem immediately above rhizome apex, slender, wiry, up to about
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long, sometimes bearing 1 or more globose, orange root nodules. Peduncle erect, slender, 4- or 5-noded, glabrous,
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long, 2.03.0 mm in diameter, beige to dark brown. Floral rachis 6.510.0 cm long, elongating during anthesis, laxly to sub-densely 1620- flowered; floral bracts ovate-triangular, 1.02.0 mm long, acute, dark brown. Flowers resupinate, opening acropetally one or two at a time, white flushed pale lilac, the inner surface of the lateral sepals streaked orange-rose at base, the lip lilac, the column with a blue-violet tinge; pedicel and ovary 6.0
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long, brown; dorsal sepal connate with the petals for 2/3 of its length, forming a hood over the column, elliptic-obovate,
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long, 3.0
<quantity id="4CF79B39FFA0D735FAA5FD99FAB1FDCB" box="[1297,1383,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="5" unit="mm" value="4.2">4.2 mm</quantity>
wide, 3-veined, free apical portion ovate, obtuse, outer surface slightly verrucose towards the apex; lateral sepals connate for 2/3 of their length, longitudinally deflexed from their common margin, broadly elliptic-obovate,
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long,
<quantity id="4CF79B39FFA0D735FF3CFD6DFEC6FD77" box="[136,272,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="4" pageNumber="5" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.83.1 mm</quantity>
wide, 3-veined, free apical portion broadly ovate, rounded, outer surface slightly verrucose towards the apex; petals broadly oblong-ovate, slightly oblique, 8.09.0 mm long, 4.0
<quantity id="4CF79B39FFA0D735FC73FD09FBCAFD5B" box="[967,1052,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="5" unit="mm" value="4.4">4.4 mm</quantity>
wide, 2-veined, free apical portions broadly ovate, apical margins erose; labellum projecting forwards, obtriangular when flattened,
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long,
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wide, distinctly 3-lobed above the middle, bearing a long, narrow, raised, fleshy callus extending from a pair of small gland-like domes at the base to midway along the mid-lobe; mid-lobe quadrate, minutely erose along the apical margin; side lobes oblong, 2.0
<quantity id="4CF79B39FFA0D735FD8DFCB9FD59FC2B" box="[569,655,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="5" unit="mm" value="2.7">2.7 mm</quantity>
long,
<quantity id="4CF79B39FFA0D735FD6CFCB9FCB6FC2B" box="[728,864,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="4" pageNumber="5" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.21.4 mm</quantity>
wide, distinctly longer than the mid-lobe, deflexed, truncate-rounded, apical margin irregularly denticulate. Column erect, slightly arched, ca.
<quantity id="4CF79B39FFA0D735FB33FC45FB1FFC0F" box="[1159,1225,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="mm" value="5.0">5 mm</quantity>
long, with a pair of prominent, decurved, fang-like stelidia ca.
<quantity id="4CF79B39FFA0D735FDC5FC61FD1FFC73" box="[625,713,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="5" unit="mm" value="2.2">2.2 mm</quantity>
long at the apex, foot absent; anther hemi-spherical, ca.
<quantity id="4CF79B39FFA0D735FAE0FC61FA7AFC73" box="[1364,1452,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="5" unit="mm" value="1.1">1.1 mm</quantity>
across; pollinia 4, in 2 pairs; stigma ventral, slightly raised, with a prominent rostellum along apex. Capsule fusiform, ca.
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long.
</paragraph>
<paragraph id="8BB036DCFFA0D735FF09FBF5FA9AFBC2" blockId="4.[136,1452,158,1841]" pageId="4" pageNumber="5">
<emphasis id="B97BEACEFFA0D735FF09FBF5FDF7FB9E" bold="true" box="[189,545,1058,1084]" pageId="4" pageNumber="5">Other specimens examined:―</emphasis>
<collectingCountry id="F318764CFFA0D735FD95FBF5FDB8FB9E" box="[545,622,1058,1084]" name="Laos" pageId="4" pageNumber="5">LAOS</collectingCountry>
.
<collectingRegion id="49CBF83EFFA0D735FDC2FBF5FC98FB9E" box="[630,846,1058,1084]" country="Laos" name="Viangchan" pageId="4" pageNumber="5">Vientiane Province</collectingRegion>
: Vang Vieng District, Phou Din Deng,
<quantity id="4CF79B39FFA0D735FB47FBF5FAEFFB9F" box="[1267,1337,1058,1085]" metricMagnitude="2" metricUnit="m" metricValue="3.53" pageId="4" pageNumber="5" unit="m" value="353.0">353 m</quantity>
elevation,
<date id="FFB1101CFFA0D735FF3CFB91FEF1FBC2" box="[136,295,1094,1120]" pageId="4" pageNumber="5" value="2012-04-30">30 April 2012</date>
,
<emphasis id="B97BEACEFFA0D735FE86FB91FB26FBC2" box="[306,1264,1094,1121]" italics="true" pageId="4" pageNumber="5">Fischer, Gale, Kumar, Bouamanivong, Santainsy and Phunthavong HNL-KFBG 0282</emphasis>
(HNL!)
</paragraph>
<paragraph id="8BB036DCFFA0D735FF09FBBDFE08FB26" blockId="4.[136,1452,158,1841]" box="[189,478,1130,1156]" pageId="4" pageNumber="5">
<emphasis id="B97BEACEFFA0D735FF09FBBDFE8AFB26" bold="true" box="[189,348,1130,1156]" pageId="4" pageNumber="5">Flowering:—</emphasis>
AprilMay.
</paragraph>
<paragraph id="8BB036DCFFA0D735FF09FB59FA44FB0A" blockId="4.[136,1452,158,1841]" box="[189,1426,1166,1193]" pageId="4" pageNumber="5">
<emphasis id="B97BEACEFFA0D735FF09FB59FEE8FB0A" bold="true" box="[189,318,1166,1192]" pageId="4" pageNumber="5">Habitat:—</emphasis>
Plants growing in organic debris between rocky crevices under broadleaf forest on limestone karst.
</paragraph>
</subSubSection>
<subSubSection id="C3156557FFA0D735FF09FB65FB49F892" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BB036DCFFA0D735FF09FB65FB75FB6E" blockId="4.[136,1452,158,1841]" box="[189,1187,1202,1229]" pageId="4" pageNumber="5">
<emphasis id="B97BEACEFFA0D735FF09FB65FEB2FB6E" bold="true" box="[189,356,1202,1228]" pageId="4" pageNumber="5">Etymology:—</emphasis>
The species is named in honour of the country in which it was discovered.
</paragraph>
<paragraph id="8BB036DCFFA0D735FF09FB01FA90FAFE" blockId="4.[136,1452,158,1841]" pageId="4" pageNumber="5">
<emphasis id="B97BEACEFFA0D735FF09FB01FE7CFB52" bold="true" box="[189,426,1238,1264]" pageId="4" pageNumber="5">Taxonomic notes:—</emphasis>
Recognition of
<taxonomicName id="4C0F4D5FFFA0D735FDE2FB00FD08FB52" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[598,734,1238,1264]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="laosensis">
<emphasis id="B97BEACEFFA0D735FDE2FB00FD08FB52" box="[598,734,1238,1264]" italics="true" pageId="4" pageNumber="5">D. laosensis</emphasis>
</taxonomicName>
brings the total number of species in the genus to eight. All share a similar column and column-foot structure, on the basis of which they can be readily distinguished from members of both
<taxonomicName id="4C0F4D5FFFA0D735FF77FAC9FE8BFA9A" authorityName="Griffith" authorityYear="1844" box="[195,349,1310,1336]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
<emphasis id="B97BEACEFFA0D735FF77FAC9FE8BFA9A" box="[195,349,1310,1336]" italics="true" pageId="4" pageNumber="5">Didymoplexis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F4D5FFFA0D735FE22FAC9FD89FA9A" authorityName="Seidenfaden" authorityYear="1997" box="[406,607,1310,1336]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiopsis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
<emphasis id="B97BEACEFFA0D735FE22FAC9FD89FA9A" box="[406,607,1310,1336]" italics="true" pageId="4" pageNumber="5">Didymoplexiopsis</emphasis>
</taxonomicName>
. Flower colour is variable and so cannot be used for species delimitation. The structure and outline of the labellum are therefore the most important features for identification (
<figureCitation id="13342A59FFA0D735FB40FA95FAEEFAFE" box="[1268,1336,1346,1372]" captionStart="FIGURE 3" captionStartId="5.[136,229,1103,1125]" captionTargetBox="[156,1433,193,1053]" captionTargetId="figure-18@5.[151,1436,190,1079]" captionTargetPageId="5" captionText="FIGURE 3. Comparison of flower and labellum of Didymoplexiella laosensis with those of morphologically similar members of the genus.AB. Didymoplexiella laosensis. CD.Didymoplexiella siamensis. EF.Didymoplexiella ornata. GH.Didymoplexiella trichechus. Photo credits:A. Stephan Gale; C. Pankaj Kumar; E. Ong Poh Teck; G. Tsukaya Hirokazu.All line drawings drawn by P. Kumar (D drawn from fresh specimen collected in Hong Kong; F drawn from colour photographs of plants in Malaysia provided by Ong Poh Teck and from original description (Ridley 1891); H redrawn from Tsukaya et al. (2014) from Borneo)." figureDoi="http://doi.org/10.5281/zenodo.13879379" httpUri="https://zenodo.org/record/13879379/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BB036DCFFA0D735FF09FAB1FB49F892" blockId="4.[136,1452,158,1841]" pageId="4" pageNumber="5">
<emphasis id="B97BEACEFFA0D735FF09FAB1FDDAFA22" bold="true" box="[189,524,1382,1408]" pageId="4" pageNumber="5">Conservation assessment:—</emphasis>
Three scattered populations of
<taxonomicName id="4C0F4D5FFFA0D735FCD5FAB1FB50FA22" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[865,1158,1382,1408]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="laosensis">
<emphasis id="B97BEACEFFA0D735FCD5FAB1FB50FA22" box="[865,1158,1382,1408]" italics="true" pageId="4" pageNumber="5">Didymoplexiella laosensis</emphasis>
</taxonomicName>
were found in Vang Vieng District of
<collectingRegion id="49CBF83EFFA0D735FEB1FA5DFE09FA06" box="[261,479,1418,1444]" country="Laos" name="Viangchan" pageId="4" pageNumber="5">Vientiane Province</collectingRegion>
, central
<collectingCountry id="F318764CFFA0D735FDF6FA5CFDAAFA07" box="[578,636,1419,1445]" name="Laos" pageId="4" pageNumber="5">Laos</collectingCountry>
. The extensive limestone karst landscape of the region offers hope that additional populations may yet be found but further surveys are needed to confirm this. The Extent of Occurrence and Area of Occupancy (AOO) are estimated to be &lt;
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and
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, respectively (
<bibRefCitation id="EF9E4B2DFFA0D735FBF2FA05FB1FFA4E" author="Moat, J." box="[1094,1225,1490,1516]" pageId="4" pageNumber="5" refId="ref4310" refString="Moat, J. (2007) Conservation assessment tools extension for Arc View 3. x, version 1.2. GIS Unit, Royal Botanic Gardens, Kew. Available from: http: // www. rbgkew. org. uk / gis / cats (accessed 10 October 2019)" type="url" year="2007">Moat 2007</bibRefCitation>
). On the strength of current knowledge, the total population size is estimated to amount to fewer than 500 mature individuals. Being a holomycotroph, species of
<taxonomicName id="4C0F4D5FFFA0D735FE76F9CDFDADF996" authorityName="Garay" authorityYear="1954" box="[450,635,1562,1588]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
<emphasis id="B97BEACEFFA0D735FE76F9CDFDADF996" box="[450,635,1562,1588]" italics="true" pageId="4" pageNumber="5">Didymoplexiella</emphasis>
</taxonomicName>
are expected to be highly specific in their mycorrhizal symbiosis and therefore also their habitat (
<bibRefCitation id="EF9E4B2DFFA0D735FE70F9E9FD1BF9FA" author="Jacquemyn, H. &amp; Waud, M. &amp; Merckx, V. S. F. T. &amp; Brys, R. &amp; Tyteca, D. &amp; Hedren, M. &amp; Lievens, B." box="[452,717,1598,1625]" pageId="4" pageNumber="5" refId="ref3794" refString="Jacquemyn, H., Waud, M., Merckx, V. S. F. T., Brys, R., Tyteca, D., Hedren, M. &amp; Lievens, B. (2016) Habitat-driven variation in mycorrhizal communities in the terrestrial orchid genus Dactylorhiza. Scientific Reports 6: 37182. https: // doi. org / 10.1038 / srep 37182" type="journal volume" year="2016">
Jacquemyn
<emphasis id="B97BEACEFFA0D735FDF9F9E8FD56F9FA" box="[589,640,1598,1624]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2016
</bibRefCitation>
). Clearance of broadleaf forest across the seasonal Asian tropics continues to endanger the unique biota of the region (
<bibRefCitation id="EF9E4B2DFFA0D735FD54F9B5FC19F9DE" author="Clements, R. &amp; Sodhi, N. S. &amp; Schilthuizen, M. &amp; Peter, K. L. N." box="[736,975,1634,1661]" pageId="4" pageNumber="5" pagination="733 - 742" refId="ref3172" refString="Clements, R., Sodhi, N. S., Schilthuizen, M. &amp; Peter, K. L. N. (2006) Limestone karsts of Southeast Asia: Imperiled arks of biodiversity. BioScience 56 (9): 733 - 742. https: // doi. org / 10.1641 / 0006 - 3568 (2006) 56 [733: LKOSAI] 2.0. CO; 2" type="journal article" year="2006">
Clements
<emphasis id="B97BEACEFFA0D735FCE7F9B4FC53F9DE" box="[851,901,1634,1660]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2006
</bibRefCitation>
), with natural regeneration and restoration of limestone floras being regarded as especially difficult (
<bibRefCitation id="EF9E4B2DFFA0D735FCAFF951FBCEF902" author="Urich, P. B. &amp; Reeder, P." box="[795,1048,1670,1696]" pageId="4" pageNumber="5" pagination="493 - 506" refId="ref5074" refString="Urich, P. B. &amp; Reeder, P. (1999) Plantation Forestry in Tropical Limestone Uplands: Environmental Constraints and Opportunities. The Professional Geographer 51 (4): 493 - 506. https: // doi. org / 10.1111 / 0033 - 0124.00186" type="journal article" year="1999">Urich &amp; Reeder 1999</bibRefCitation>
). However, the immediate threat to this species is habitat destruction due to mining (
<bibRefCitation id="EF9E4B2DFFA0D735FD0AF97DFC4BF966" author="Day, M. J. &amp; Urich, P. B." box="[702,925,1706,1732]" pageId="4" pageNumber="5" pagination="61 - 70" refId="ref3247" refString="Day, M. J. &amp; Urich, P. B. (2000) An assessment of protected karst landscapes in Southeast Asia. Cave and Karst Science 27: 61 - 70." type="journal article" year="2000">Day &amp; Urich 2000</bibRefCitation>
). Part of the limestone mountain range where
<taxonomicName id="4C0F4D5FFFA0D735FF3CF918FEC4F94A" authorityName="S. W. Gale &amp; Kumar" authorityYear="2020" box="[136,274,1742,1768]" class="Liliopsida" family="Orchidaceae" genus="Didymoplexiella" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="laosensis">
<emphasis id="B97BEACEFFA0D735FF3CF918FEC4F94A" box="[136,274,1742,1768]" italics="true" pageId="4" pageNumber="5">D. laosensis</emphasis>
</taxonomicName>
was discovered was leased to a cement production company by the Lao government in 2015. Presently, only 10% of the limestone karst area of central and northern
<collectingCountry id="F318764CFFA0D735FC8FF924FCA7F8AF" box="[827,881,1779,1805]" name="Laos" pageId="4" pageNumber="5">Laos</collectingCountry>
is under statutory protection (
<bibRefCitation id="EF9E4B2DFFA0D735FB73F925FA76F8AE" author="Delang, C. O. &amp; Toro, M. &amp; Phommachanh, M. C." box="[1223,1440,1778,1804]" pageId="4" pageNumber="5" pagination="150 - 164" refId="ref3283" refString="Delang, C. O., Toro, M. &amp; Phommachanh, M. C. (2013) Coffee, mines and dams: conflicts over land in the Bolaven Plateau, southern Lao PDR. The Geographical Journal 179: 150 - 164. https: // doi. org / 10.1111 / j. 1475 - 4959.2012.00481. x" type="journal article" year="2013">
Delang
<emphasis id="B97BEACEFFA0D735FA96F924FA8DF8AE" box="[1314,1371,1778,1804]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2013
</bibRefCitation>
). Applying
<bibRefCitation id="EF9E4B2DFFA0D735FF4DF8C1FE59F892" author="IUCN Standards &amp; Petitions Committee" box="[249,399,1814,1840]" pageId="4" pageNumber="5" refId="ref3740" refString="IUCN Standards and Petitions Committee (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed on 6 October 2019)" type="book" year="2019">IUCN (2019)</bibRefCitation>
guidelines, the species is assessed as Endangered (EN B1+B2ab(iii)).
</paragraph>
</subSubSection>
</treatment>
</document>