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data/EE/BB/86/EEBB86ACFBB55F28A935634ECC2392B2.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.85399" ID-Pensoft-Pub="1313-2970-1110-39" ID-Pensoft-UUID="766F9A94793D5793A9BE1E70F575798B" ID-ZooBank="4681CC6DE5F347C6B1D052DEA78BE7C3" ModsDocID="1313-2970-1110-39" checkinTime="1657006630715" checkinUser="pensoft" docAuthor="Sian Man, Nem, Siriboon, Thanit, Lin, Aung, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="EEBB86ACFBB55F28A935634ECC2392B2" docLanguage="en" docName="ZooKeys 1110: 39-102" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.85399" docTitle="Haploptychius karenorum Man & Panha 2022, sp. nov." docType="treatment" docUuid="556A1CE1-7E73-4466-9C69-F5454EB4E5CC" docUuidSource="ZooBank" docVersion="1" id="766F9A94793D5793A9BE1E70F575798B" lastPageNumber="39" masterDocId="766F9A94793D5793A9BE1E70F575798B" masterDocTitle="Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species" masterLastPageNumber="102" masterPageNumber="39" pageNumber="39" updateTime="1657006630715" updateUser="pensoft">
|
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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||||
<mods:titleInfo>
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<mods:title>Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species</mods:title>
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||||
</mods:titleInfo>
|
||||
<mods:name type="personal">
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||||
<mods:role>
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||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart>Sian Man, Nem</mods:namePart>
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||||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4453-734X</mods:nameIdentifier>
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||||
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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||||
</mods:name>
|
||||
<mods:name type="personal">
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||||
<mods:role>
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||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart>Siriboon, Thanit</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok, 10110, Thailand</mods:affiliation>
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||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
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||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart>Lin, Aung</mods:namePart>
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||||
<mods:affiliation>Fauna and Flora International, No. 35, 3</mods:affiliation>
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||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
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||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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||||
<mods:name type="personal">
|
||||
<mods:role>
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||||
<mods:roleTerm>Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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||||
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>ZooKeys</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
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||||
<mods:date>2022</mods:date>
|
||||
<mods:detail type="pubDate">
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||||
<mods:number>2022-07-04</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="volume">
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||||
<mods:number>1110</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
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||||
<mods:start>39</mods:start>
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||||
<mods:end>102</mods:end>
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||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://dx.doi.org/10.3897/zookeys.1110.85399</mods:url>
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||||
</mods:location>
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||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1110.85399</mods:identifier>
|
||||
<mods:identifier type="Pensoft-Pub">1313-2970-1110-39</mods:identifier>
|
||||
<mods:identifier type="ZooBank">4681CC6DE5F347C6B1D052DEA78BE7C3</mods:identifier>
|
||||
<mods:identifier type="Pensoft-UUID">766F9A94793D5793A9BE1E70F575798B</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment LSID="urn:lsid:zoobank.org:act:556A1CE1-7E73-4466-9C69-F5454EB4E5CC" httpUri="http://treatment.plazi.org/id/EEBB86ACFBB55F28A935634ECC2392B2" lastPageNumber="39" pageId="0" pageNumber="39">
|
||||
<subSubSection pageId="0" pageNumber="39" type="nomenclature">
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<taxonomicName LSID="https://zoobank.org/556A1CE1-7E73-4466-9C69-F5454EB4E5CC" authority="Man & Panha" authorityName="Man & Panha" authorityYear="2022" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius karenorum" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="karenorum" status="sp. nov.">Haploptychius karenorum Man & Panha</taxonomicName>
|
||||
<taxonomicNameLabel pageId="0" pageNumber="39">sp. nov.</taxonomicNameLabel>
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="description">
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A, B Haploptychius heliakosus sp. nov. A paratype CUMZ 13014 from the type locality and B specimen CUMZ 13016 from Kyonknow Cave, Hpa-an, Kayin State C-E Haploptychius karenorum sp. nov. C holotype CUMZ 13017 with apertural dentition D paratype CUMZ 13018 from the type locality and E specimen CUMZ 13019 from Taung Lay Cave, Hpa-an, Kayin State." figureDoi="10.3897/zookeys.1110.85399.figure14" httpUri="https://binary.pensoft.net/fig/710479" pageId="0" pageNumber="39">Fig. 14C-E</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="type material">
|
||||
<paragraph pageId="0" pageNumber="39">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Holotype</emphasis>
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CUMZ 13017 (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A, B Haploptychius heliakosus sp. nov. A paratype CUMZ 13014 from the type locality and B specimen CUMZ 13016 from Kyonknow Cave, Hpa-an, Kayin State C-E Haploptychius karenorum sp. nov. C holotype CUMZ 13017 with apertural dentition D paratype CUMZ 13018 from the type locality and E specimen CUMZ 13019 from Taung Lay Cave, Hpa-an, Kayin State." figureDoi="10.3897/zookeys.1110.85399.figure14" httpUri="https://binary.pensoft.net/fig/710479" pageId="0" pageNumber="39">14C</figureCitation>
|
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). Measurements: shell height 9 mm, shell width 11.6 mm and 7 whorls.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="39">Paratypes</emphasis>
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CUMZ 13018 (6 shells; Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A, B Haploptychius heliakosus sp. nov. A paratype CUMZ 13014 from the type locality and B specimen CUMZ 13016 from Kyonknow Cave, Hpa-an, Kayin State C-E Haploptychius karenorum sp. nov. C holotype CUMZ 13017 with apertural dentition D paratype CUMZ 13018 from the type locality and E specimen CUMZ 13019 from Taung Lay Cave, Hpa-an, Kayin State." figureDoi="10.3897/zookeys.1110.85399.figure14" httpUri="https://binary.pensoft.net/fig/710479" pageId="0" pageNumber="39">14D</figureCitation>
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), NHMUK (2 shells).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="39">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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Limestone outcrops at Waiponla Hill, Hpa-an Township, Hpa-an District, Kayin State, Myanmar (
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<geoCoordinate degrees="16" direction="north" minutes="56" orientation="latitude" precision="1" seconds="7.4" value="16.935389">16°56'7.4"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="42" orientation="longitude" precision="1" seconds="56.8" value="97.715775">97°42'56.8"E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="materials_examined">
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<paragraph pageId="0" pageNumber="39">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Myanmar" county="Hpa-an District" latitude="17.194527" location="Taung Lay Cave" longLatPrecision="1" longitude="97.62972" municipality="Hpa-an Township" specimenCount="1" stateProvince="Kayin State">
|
||||
<location LSID="urn:lsid:plazi:treatment:EEBB86ACFBB55F28A935634ECC2392B2:DFC76CC3E5409DABBA2B7BD629B7C2D5" country="Myanmar" county="Hpa-an District" latitude="17.194527" longLatPrecision="1" longitude="97.62972" municipality="Hpa-an Township" name="Taung Lay Cave" stateProvince="Kayin State">Taung Lay Cave</location>
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,
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<collectingMunicipality>Hpa-an Township</collectingMunicipality>
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,
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<collectingCounty>Hpa-an District</collectingCounty>
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,
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<collectingRegion country="Myanmar" name="Kayin">Kayin State</collectingRegion>
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,
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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(
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<geoCoordinate degrees="17" direction="north" minutes="11" orientation="latitude" precision="1" seconds="40.3" value="17.194527">17°11'40.3"N</geoCoordinate>
|
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,
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<geoCoordinate degrees="97" direction="east" minutes="37" orientation="longitude" precision="1" seconds="47.0" value="97.62972">97°37'47.0"E</geoCoordinate>
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): CUMZ 13019 (2 shells; Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A, B Haploptychius heliakosus sp. nov. A paratype CUMZ 13014 from the type locality and B specimen CUMZ 13016 from Kyonknow Cave, Hpa-an, Kayin State C-E Haploptychius karenorum sp. nov. C holotype CUMZ 13017 with apertural dentition D paratype CUMZ 13018 from the type locality and E specimen CUMZ 13019 from Taung Lay Cave, Hpa-an, Kayin State." figureDoi="10.3897/zookeys.1110.85399.figure14" httpUri="https://binary.pensoft.net/fig/710479" pageId="0" pageNumber="39">14E</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="diagnosis">
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<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName authorityName="Man & Panha" authorityYear="2022" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius karenorum" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="karenorum">
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<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius karenorum</emphasis>
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</taxonomicName>
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sp. nov. can be differentiated from
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<taxonomicName genus="H." lsidName="H. heliakosus" pageId="0" pageNumber="39" rank="species" species="heliakosus">
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<emphasis italics="true" pageId="0" pageNumber="39">H. heliakosus</emphasis>
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</taxonomicName>
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sp. nov. by having a convex spire, penultimate whorl angular and extended well beyond the diameter of the last whorl, and more axially deflected last whorl. In contrast,
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<taxonomicName genus="H." lsidName="H. heliakosus" pageId="0" pageNumber="39" rank="species" species="heliakosus">
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<emphasis italics="true" pageId="0" pageNumber="39">H. heliakosus</emphasis>
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</taxonomicName>
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sp. nov. possesses an elevated spire, penultimate whorl rounded, less extended beyond the diameter of the last whorl, and less axially deflected last whorl. This new species differs from
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<taxonomicName genus="H." lsidName="H. bombax" pageId="0" pageNumber="39" rank="species" species="bombax">
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<emphasis italics="true" pageId="0" pageNumber="39">H. bombax</emphasis>
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</taxonomicName>
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by having an angular penultimate whorl, more axially deflected last whorl, subcircular aperture, and broadly expanded lip. In comparison,
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<taxonomicName genus="H." lsidName="H. bombax" pageId="0" pageNumber="39" rank="species" species="bombax">
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<emphasis italics="true" pageId="0" pageNumber="39">H. bombax</emphasis>
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</taxonomicName>
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has a rounded penultimate whorl, less axially deflected last whorl, semi-ovate aperture, with thickened and slightly expanded lip. For further comparison,
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<taxonomicName genus="H." lsidName="H. karenorum" pageId="0" pageNumber="39" rank="species" species="karenorum">
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<emphasis italics="true" pageId="0" pageNumber="39">H. karenorum</emphasis>
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</taxonomicName>
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sp. nov. differs from
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<taxonomicName genus="H." lsidName="H. burmanicus" pageId="0" pageNumber="39" rank="species" species="burmanicus">
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<emphasis italics="true" pageId="0" pageNumber="39">H. burmanicus</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="H." lsidName="H. blanfordi" pageId="0" pageNumber="39" rank="species" species="blanfordi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. blanfordi</emphasis>
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</taxonomicName>
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, and
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<taxonomicName genus="H." lsidName="H. thebawi" pageId="0" pageNumber="39" rank="species" species="thebawi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. thebawi</emphasis>
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</taxonomicName>
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in having a more axially deflected last whorl, lower spire, penultimate whorl angular and strongly extended beyond the diameter of the last whorl, and without sinulus. The three latter species have an elevated spire, penultimate whorl rounded and slightly extended beyond the diameter of the last whorl, and less axially deflected last whorl.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="description">
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<paragraph pageId="0" pageNumber="39">Description.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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Shell oblique-ovate, solid and translucent; whorls 6-
|
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<normalizedToken originalValue="7½">71/2</normalizedToken>
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; spire depressed convex and with distinct suture. Embryonic shell ~
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<normalizedToken originalValue="2½">21/2</normalizedToken>
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whorls with smooth surface; following whorls growing regularly and last whorl intermediately expanded. Shell surface has moderately strong radial ridges that diminish below periphery of last whorl and around umbilicus. Penultimate whorl bluntly angular and extended beyond last whorl. Last whorl compressed to flattened, axially deflected from columellar axis. Aperture subcircular; peristome thickened, expanded, and slightly reflected. Apertural dentition with one strong parietal lamella. Umbilicus widely open and deep (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A, B Haploptychius heliakosus sp. nov. A paratype CUMZ 13014 from the type locality and B specimen CUMZ 13016 from Kyonknow Cave, Hpa-an, Kayin State C-E Haploptychius karenorum sp. nov. C holotype CUMZ 13017 with apertural dentition D paratype CUMZ 13018 from the type locality and E specimen CUMZ 13019 from Taung Lay Cave, Hpa-an, Kayin State." figureDoi="10.3897/zookeys.1110.85399.figure14" httpUri="https://binary.pensoft.net/fig/710479" pageId="0" pageNumber="39">14C-E</figureCitation>
|
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="etymology">
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<paragraph pageId="0" pageNumber="39">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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The specific name
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<emphasis italics="true" pageId="0" pageNumber="39">karenorum</emphasis>
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refers to the Karen people, the major ethnicity in Kayin State, Myanmar.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="distribution">
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<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="39">This species is known from two localities in the limestone karts of Kayin State, southern Myanmar.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="remarks">
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<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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Comparing populations from the two localities, shells of
|
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<taxonomicName genus="H." lsidName="H. karenorum" pageId="0" pageNumber="39" rank="species" species="karenorum">
|
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<emphasis italics="true" pageId="0" pageNumber="39">H. karenorum</emphasis>
|
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</taxonomicName>
|
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sp. nov. from the Taung Lay population (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A, B Haploptychius heliakosus sp. nov. A paratype CUMZ 13014 from the type locality and B specimen CUMZ 13016 from Kyonknow Cave, Hpa-an, Kayin State C-E Haploptychius karenorum sp. nov. C holotype CUMZ 13017 with apertural dentition D paratype CUMZ 13018 from the type locality and E specimen CUMZ 13019 from Taung Lay Cave, Hpa-an, Kayin State." figureDoi="10.3897/zookeys.1110.85399.figure14" httpUri="https://binary.pensoft.net/fig/710479" pageId="0" pageNumber="39">14E</figureCitation>
|
||||
) have a more elevated spire and weaker parietal lamella. However, the bluntly angular penultimate whorl and oblique ovate shells are identical to the type specimens, which we consider an intraspecific shell variation. However, these two localities are quite distant from each other, and on opposite sides of the Attaran River (Fig.
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||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Approximate sampling locations used in this study. The location name and coordinates are listed in Table 1. Historical type localities of a broad scale, such as Tavoy, Burma, Moulmein, Mergui, and Tenasserim, are not indicated on this map." figureDoi="10.3897/zookeys.1110.85399.figure1" httpUri="https://binary.pensoft.net/fig/710466" pageId="0" pageNumber="39">1</figureCitation>
|
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). Thus further, genitalia information will help determine whether they are just geographical variations.
|
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</paragraph>
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</subSubSection>
|
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</treatment>
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</document>
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data/EE/BB/90/EEBB90E16B2D7F2E75E28A014E8660B4.xml
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data/EE/BB/90/EEBB90E16B2D7F2E75E28A014E8660B4.xml
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@ -0,0 +1,130 @@
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|||
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<mods:date id="BB721200296A1CCACBC0A17FD6705E99">2016</mods:date>
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<mods:number id="71442C9439418B81F66D3BFD1FDF3966">4</mods:number>
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<mods:identifier id="506693FFD07A2BA1C7B6765E2058F4FB" type="Pensoft-Pub">1314-2828-4-8354</mods:identifier>
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<treatment id="EEBB90E16B2D7F2E75E28A014E8660B4" LSID="urn:lsid:plazi:treatment:EEBB90E16B2D7F2E75E28A014E8660B4" httpUri="http://treatment.plazi.org/id/EEBB90E16B2D7F2E75E28A014E8660B4" lastPageNumber="8354" pageId="0" pageNumber="8354">
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<subSubSection id="F48F83619911AC956504D2594DC5E1CE" pageId="0" pageNumber="8354" type="nomenclature">
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<paragraph id="6D45FE200DBF2488DC76D7DD71F016FD" pageId="0" pageNumber="8354">
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<taxonomicName id="DF90A6F28FEC64DF74FFDFCF072F82E8" ID-CoL="3CXZ" authority="Kirby, 1889" authorityName="Kirby" authorityYear="1889" class="Insecta" family="Libellulidae" genus="Brechmorhoga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brechmorhoga" order="Odonata" pageId="0" pageNumber="8354" phylum="Arthropoda" rank="genus">Brechmorhoga Kirby, 1889</taxonomicName>
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</paragraph>
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<subSubSection id="7D05E3C1076F4CC2B6EEDDCB3B1574E9" pageId="0" pageNumber="8354" type="notes">
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<paragraph id="34ED89A5410A2546CAF15E709953AB70" pageId="0" pageNumber="8354">Notes</paragraph>
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<paragraph id="D0564D37BA1DAD7FE7A760AB7B882823" pageId="0" pageNumber="8354">New genus record for PI.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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||||
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