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<document id="0307AE620C5E90A4E8055764DAAFD65D" ID-DOI="10.11646/zootaxa.5026.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300630" ID-ZooBank="511C28F6-07E4-412C-B83E-91D2CAC5AA27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630038704990" checkinUser="plazi" docAuthor="Bennett, Robb, Copley, Claudia &amp; Copley, Darren" docDate="2021" docId="EE3CB72DFF8BFFB8BEA9A4AC3A64E78F" docLanguage="en" docName="zootaxa.5026.4.1.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybaeus topanga Bennett &amp; Copley &amp; Copley 2021, spec. nov." docType="treatment" docVersion="4" lastPageNumber="474" masterDocId="1205CF55FF9EFFAFBE3EA77E3E6CE446" masterDocTitle="Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade" masterLastPageNumber="479" masterPageNumber="451" pageNumber="472" updateTime="1699138999734" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="04799CAA27F5764EED61495292EDFDC2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9C5E9516BB251FB115E38E3FD282C7B">
<mods:title id="159285B20D1E16DBCAD0752E48D0D37A">Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade</mods:title>
</mods:titleInfo>
<mods:name id="A4FC2C45C35115175FC6E583925446EE" type="personal">
<mods:role id="6E3651D0410116197CAC7CF6A21C5C0F">
<mods:roleTerm id="C5930FBF9CE33A366E2388964F1CAAB3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F5439EAA4CB98C9DF5D35DBFB70E8C48">Bennett, Robb</mods:namePart>
<mods:affiliation id="0F0E677F44580CDDDC0E15345953FAA8">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2.</mods:affiliation>
</mods:name>
<mods:name id="1E8B4024DB51C588F37531DB06AE8D71" type="personal">
<mods:role id="59B07E7A150327E1FF0C4C45E55F38D6">
<mods:roleTerm id="1CB32B17C8424C84CE621E597E481D1F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5590C338ECE3E971EEBA94D2E522AA48">Copley, Claudia</mods:namePart>
<mods:nameIdentifier id="82C1A79C06C4C60F8358DE6592F9976D" type="ORCID">0000-0002-8184-5819</mods:nameIdentifier>
<mods:affiliation id="8FE28C5E3F3C35DD326CCAB97768B1F7">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2. &amp; ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
<mods:nameIdentifier id="AA63FC053EF1AB0A97322D911A5154C3" type="email">ccopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="9F78464F4389C226553F64C864FF6BA9" type="personal">
<mods:role id="73E601773DBF4089BC1D9202CF0A4420">
<mods:roleTerm id="AD6F9AFDFDBC5F50E5A39337384D0FE5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="34E2532894CD64E35CB54779AED1EAE7">Copley, Darren</mods:namePart>
<mods:nameIdentifier id="CF1E64D61CD4FF7AA553378DBC8869DA" type="ORCID">0000-0002-1944-4272</mods:nameIdentifier>
<mods:affiliation id="EA99094107040B11721D4C4E9E243330">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2. &amp; dcopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 1944 - 4272</mods:affiliation>
<mods:nameIdentifier id="E6277DDBECCC6DB9EEC6D6E3F17DE85B" type="email">dcopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="051030639F097D454A06D141FF43B1A0">text</mods:typeOfResource>
<mods:relatedItem id="FBCBBBD5D8D5F1B39512E78BC0EE12D1" type="host">
<mods:titleInfo id="5FD26ED48C8CBFD021AE8D3829374BBF">
<mods:title id="4C6DAAB43AED5804C3471B22027E6B37">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3D6F8849255FAD40BAD0B5DDEBEB67D8">
<mods:date id="7842D60D75C31867A03285FC9B37061C">2021</mods:date>
<mods:detail id="5AACB15BA767A4CF4F8B9E218E5B0F5B" type="pubDate">
<mods:number id="8E42B2BF95F2929CBBA4C4FDC5A261A6">2021-08-27</mods:number>
</mods:detail>
<mods:detail id="6E67E39D9866E40D8A3B3BA3F3D3ACC0" type="volume">
<mods:number id="E64579B9F7929B178A2282CFCF724496">5026</mods:number>
</mods:detail>
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<mods:number id="FF5184EAAD19D077740CFD81D9C5AE59">4</mods:number>
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<mods:start id="D726FF4DA49072B83D3C6D8968F52113">451</mods:start>
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<mods:classification id="6FF4F922C822B530CA0E6F24D103CEB7">journal article</mods:classification>
<mods:identifier id="960A489F8C67AABEB37ADDC89A3E5C68" type="DOI">10.11646/zootaxa.5026.4.1</mods:identifier>
<mods:identifier id="794E314096FAC44D86EBCF27CB93721A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="15B53732F9642421110898195573E252" type="Zenodo-Dep">5300630</mods:identifier>
<mods:identifier id="51442494F371F815F73D5C24C478DD10" type="ZooBank">511C28F6-07E4-412C-B83E-91D2CAC5AA27</mods:identifier>
</mods:mods>
<treatment id="EE3CB72DFF8BFFB8BEA9A4AC3A64E78F" ID-DOI="http://doi.org/10.5281/zenodo.5307738" ID-Zenodo-Dep="5307738" LSID="urn:lsid:plazi:treatment:EE3CB72DFF8BFFB8BEA9A4AC3A64E78F" httpUri="http://treatment.plazi.org/id/EE3CB72DFF8BFFB8BEA9A4AC3A64E78F" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472">
<subSubSection id="2E8F55B0FF8BFFBABEA9A4AC3C23E7AB" box="[151,591,978,1005]" pageId="21" pageNumber="472" type="nomenclature">
<paragraph id="662A063BFF8BFFBABEA9A4AC3C23E7AB" blockId="21.[151,591,978,1041]" box="[151,591,978,1005]" pageId="21" pageNumber="472">
<heading id="3D62B157FF8BFFBABEA9A4AC3C23E7AB" bold="true" box="[151,591,978,1005]" fontSize="11" level="1" pageId="21" pageNumber="472" reason="1">
<emphasis id="54E1DA29FF8BFFBABEA9A4AC3C23E7AB" bold="true" box="[151,591,978,1005]" pageId="21" pageNumber="472">
<taxonomicName id="A1957DB8FF8BFFBABEA9A4AC3FBCE7AB" ID-CoL="99BLK" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,464,978,1005]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABEA9A4AC3F0BE7AB" bold="true" box="[151,359,978,1005]" italics="true" pageId="21" pageNumber="472">Cybaeus topanga</emphasis>
Bennett
</taxonomicName>
<taxonomicNameLabel id="4FD26752FF8BFFBABFE9A4AD3C23E7AB" box="[471,591,979,1005]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8BFFBABEA9A4893F05E057" box="[151,361,1014,1041]" pageId="21" pageNumber="472" type="description">
<paragraph id="662A063BFF8BFFBABEA9A4893F05E057" blockId="21.[151,591,978,1041]" box="[151,361,1014,1041]" pageId="21" pageNumber="472">
<figureCitation id="FEAE1ABEFF8BFFBABEA9A4893F7BE057" box="[151,279,1014,1041]" captionStart-0="FIGURES 50–54" captionStart-1="FIGURES 55–58" captionStartId-0="19.[151,264,1939,1963]" captionStartId-1="20.[151,264,1891,1915]" captionTargetBox-0="[157,1429,689,1916]" captionTargetBox-1="[151,1417,191,1867]" captionTargetId-0="figure-163@19.[151,1436,684,1916]" captionTargetId-1="figure-31@20.[151,1436,181,1867]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." captionText-1="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi-0="http://doi.org/10.5281/zenodo.5300661" figureDoi-1="http://doi.org/10.5281/zenodo.5300663" httpUri-0="https://zenodo.org/record/5300661/files/figure.png" httpUri-1="https://zenodo.org/record/5300663/files/figure.png" pageId="21" pageNumber="472">Figs 52–58</figureCitation>
,
<figureCitation id="FEAE1ABEFF8BFFBABF1CA4893F05E057" box="[290,361,1015,1041]" captionStart-0="FIGURES 59–61" captionStart-1="FIGURE 62" captionStartId-0="21.[151,264,1830,1854]" captionStartId-1="22.[151,250,1909,1933]" captionTargetBox-0="[151,1436,1317,1807]" captionTargetBox-1="[186,1407,198,1875]" captionTargetId-0="figure-450@21.[151,1436,1317,1807]" captionTargetId-1="figure-31@22.[169,1419,181,1885]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." captionText-1="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi-0="http://doi.org/10.5281/zenodo.5300665" figureDoi-1="http://doi.org/10.5281/zenodo.5300667" httpUri-0="https://zenodo.org/record/5300665/files/figure.png" httpUri-1="https://zenodo.org/record/5300667/files/figure.png" pageId="21" pageNumber="472">61–62</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8BFFBABEA9A3413D88E0E7" pageId="21" pageNumber="472" type="materials_examined">
<paragraph id="662A063BFF8BFFBABEA9A3413D88E0E7" blockId="21.[151,1437,1086,1293]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEA9A3413F25E01E" bold="true" box="[151,329,1086,1113]" pageId="21" pageNumber="472">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF8BFFBABF6DA3403F21E03B" collectingDate="1957-06-30" collectorName="R. Schick" country="United States of America" county="Los Angeles County" location="Topanga Canyon" pageId="21" pageNumber="472" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF8BFFBABF6DA3403C94E01F" bold="true" box="[339,760,1086,1113]" pageId="21" pageNumber="472">
<collectingCountry id="1E8246ABFF8BFFBABF6DA3403FC8E01E" box="[339,420,1086,1112]" name="United States of America" pageId="21" pageNumber="472">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF8BFFBABF8AA3403C40E01F" bold="true" box="[436,556,1086,1113]" italics="true" pageId="21" pageNumber="472">
<collectingRegion id="A451C8D9FF8BFFBABF8AA3403C40E01F" box="[436,556,1086,1113]" country="United States of America" name="California" pageId="21" pageNumber="472">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF8BFFBABC7EA3413CC1E01F" box="[576,685,1087,1113]" pageId="21" pageNumber="472" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<collectingCounty id="8F4B7EB7FF8BFFBABD3DA3413D9DE01F" box="[771,1009,1087,1113]" pageId="21" pageNumber="472">Los Angeles County</collectingCounty>
,
<location id="634A50E0FF8BFFBABDC0A3413AAFE01F" LSID="urn:lsid:plazi:treatment:EE3CB72DFF8BFFB8BEA9A4AC3A64E78F:634A50E0FF8BFFBABDC0A3413AAFE01F" box="[1022,1219,1087,1113]" country="United States of America" county="Los Angeles County" name="Topanga Canyon" pageId="21" pageNumber="472" stateProvince="California">Topanga Canyon</location>
,
<date id="122B20FBFF8BFFBABAECA3413B03E01F" box="[1234,1391,1086,1113]" pageId="21" pageNumber="472" value="1957-06-30">
<collectingDate id="026FD913FF8BFFBABAECA3413B03E01F" box="[1234,1391,1086,1113]" pageId="21" pageNumber="472" value="1957-06-30">30 June 1957</collectingDate>
</date>
(
<collectorName id="CB6063EDFF8BFFBABBBCA3413E84E038" pageId="21" pageNumber="472">R. Schick</collectorName>
), in CAS
</materialsCitation>
.
<materialsCitation id="D6FD0C66FF8BFFBABF6BA31D3A35E03B" box="[341,1113,1123,1149]" collectionCode="AMNH" country="United States of America" location="San Ysidro" municipality="San Diego." pageId="21" pageNumber="472" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<emphasis id="54E1DA29FF8BFFBABF6BA31D3FB8E03B" bold="true" box="[341,468,1123,1149]" pageId="21" pageNumber="472">
<typeStatus id="B92EB899FF8BFFBABF6BA31D3FA3E03B" box="[341,463,1123,1149]" pageId="21" pageNumber="472" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingMunicipality id="864E9C41FF8BFFBABFE6A31D3C3EE03B" box="[472,594,1123,1149]" pageId="21" pageNumber="472">San Diego.</collectingMunicipality>
<specimenCount id="7093CDB2FF8BFFBABC69A31D3C16E03B" box="[599,634,1123,1149]" pageId="21" pageNumber="472" type="female">1♀</specimenCount>
,
<location id="634A50E0FF8BFFBABCBBA31D3D6EE03B" LSID="urn:lsid:plazi:treatment:EE3CB72DFF8BFFB8BEA9A4AC3A64E78F:634A50E0FF8BFFBABCBBA31D3D6EE03B" box="[645,770,1123,1149]" country="United States of America" municipality="San Diego." name="San Ysidro" pageId="21" pageNumber="472" stateProvince="California">San Ysidro</location>
, no date, no collector (
<collectionCode id="00849EFEFF8BFFBABDCBA31D3A3DE03B" box="[1013,1105,1123,1149]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="472">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF8BFFBABA5CA31D3FE2E0E7" collectingDate="1910-12-25" collectionCode="CAS" collectorName="W. M. Wheeler" country="United States of America" location="San Ysidro" pageId="21" pageNumber="472" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF8BFFBABA5CA31D3AE9E03B" box="[1122,1157,1123,1149]" pageId="21" pageNumber="472" type="female">1♀</specimenCount>
,
<location id="634A50E0FF8BFFBABAAEA31D3B61E03B" LSID="urn:lsid:plazi:treatment:EE3CB72DFF8BFFB8BEA9A4AC3A64E78F:634A50E0FF8BFFBABAAEA31D3B61E03B" box="[1168,1293,1123,1149]" country="United States of America" name="San Ysidro" pageId="21" pageNumber="472" stateProvince="California">San Ysidro</location>
,
<date id="122B20FBFF8BFFBABB28A31C3BF4E03B" box="[1302,1432,1122,1149]" pageId="21" pageNumber="472" value="1910-12-25">
<collectingDate id="026FD913FF8BFFBABB28A31C3BF4E03B" box="[1302,1432,1122,1149]" pageId="21" pageNumber="472" value="1910-12-25">25.xii.1910</collectingDate>
</date>
,
<collectorName id="CB6063EDFF8BFFBABEA9A3F93F57E0E7" box="[151,315,1159,1185]" pageId="21" pageNumber="472">W.M. Wheeler</collectorName>
(
<collectionCode id="00849EFEFF8BFFBABF75A3F93FEBE0E7" box="[331,391,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="21" pageNumber="472" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF8BFFBABFA4A3F93D8CE0E7" box="[410,992,1158,1185]" collectingDate="1910-12-25" collectionCode="MCZ" collectorName="W. M. Wheeler" country="United States of America" location="San Ysidro" pageId="21" pageNumber="472" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF8BFFBABFA4A3F93FD1E0E7" box="[410,445,1159,1185]" pageId="21" pageNumber="472" type="female">1♀</specimenCount>
,
<location id="634A50E0FF8BFFBABFF5A3F93C26E0E7" LSID="urn:lsid:plazi:treatment:EE3CB72DFF8BFFB8BEA9A4AC3A64E78F:634A50E0FF8BFFBABFF5A3F93C26E0E7" box="[459,586,1159,1185]" country="United States of America" name="San Ysidro" pageId="21" pageNumber="472" stateProvince="California">San Ysidro</location>
,
<date id="122B20FBFF8BFFBABC6BA3F83CB4E0E7" box="[597,728,1158,1185]" pageId="21" pageNumber="472" value="1910-12-25">
<collectingDate id="026FD913FF8BFFBABC6BA3F83CB4E0E7" box="[597,728,1158,1185]" pageId="21" pageNumber="472" value="1910-12-25">25.xii.1910</collectingDate>
</date>
,
<collectorName id="CB6063EDFF8BFFBABCDCA3F93DEBE0E7" box="[738,903,1159,1185]" pageId="21" pageNumber="472">W.M. Wheeler</collectorName>
(
<collectionCode id="00849EFEFF8BFFBABDABA3F93DB4E0E7" box="[917,984,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="472" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8BFFBABEF9A3D53A04E083" box="[199,1128,1195,1221]" pageId="21" pageNumber="472" type="etymology">
<paragraph id="662A063BFF8BFFBABEF9A3D53A04E083" blockId="21.[151,1437,1086,1293]" box="[199,1128,1195,1221]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A3D53F22E083" bold="true" box="[199,334,1195,1221]" pageId="21" pageNumber="472">Etymology.</emphasis>
The specific name is a noun in apposition taken from the
<typeStatus id="B92EB899FF8BFFBABDE9A3D53A6BE083" box="[983,1031,1195,1221]" pageId="21" pageNumber="472">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8BFFB8BEF9A3B03D3DE507" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472" type="diagnosis">
<paragraph id="662A063BFF8BFFBABEF9A3B03A3AE14B" blockId="21.[151,1437,1086,1293]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A3B03F2EE0AE" bold="true" box="[199,322,1230,1256]" pageId="21" pageNumber="472">Diagnosis.</emphasis>
The form of the distal and proximal arms of the tegular apophysis (
<figureCitation id="FEAE1ABEFF8BFFBABA05A3B13AFEE0AF" box="[1083,1170,1230,1257]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1891,1915]" captionTargetBox="[151,1417,191,1867]" captionTargetId="figure-31@20.[151,1436,181,1867]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="21" pageNumber="472">Figs 57</figureCitation>
,
<figureCitation id="FEAE1ABEFF8BFFBABAA0A3B13AD7E0AC" box="[1182,1211,1231,1258]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="21" pageNumber="472">61</figureCitation>
) diagnosis the male of
<taxonomicName id="A1957DB8FF8BFFBABE88A38D3F5EE14A" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[182,306,1267,1293]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABE88A38D3F5EE14A" box="[182,306,1267,1293]" italics="true" pageId="21" pageNumber="472">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABF07A38D3FC7E14B" bold="true" box="[313,427,1267,1293]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABF07A38D3FC7E14B" box="[313,427,1267,1293]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and is discussed under the diagnosis of
<taxonomicName id="A1957DB8FF8BFFBABD53A38D3DB0E14A" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[877,988,1267,1293]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABD53A38D3DB0E14A" box="[877,988,1267,1293]" italics="true" pageId="21" pageNumber="472">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABDDAA38D3A3AE14B" bold="true" box="[996,1110,1267,1293]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABDDAA38D3A3AE14B" box="[996,1110,1267,1293]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<caption id="32EA56B3FF8BFFBABEA9A0583DDAE3C0" ID-DOI="http://doi.org/10.5281/zenodo.5300665" ID-Zenodo-Dep="5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="21" pageNumber="472" startId="21.[151,264,1830,1854]" targetBox="[151,1436,1317,1807]" targetPageId="21">
<paragraph id="662A063BFF8BFFBABEA9A0583DDAE3C0" blockId="21.[151,1436,1830,1926]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEA9A0583F39E378" bold="true" box="[151,341,1830,1854]" pageId="21" pageNumber="472">FIGURES 59–61.</emphasis>
<taxonomicName id="A1957DB8FF8BFFBABF62A0583F88E378" box="[348,484,1830,1854]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="54E1DA29FF8BFFBABF62A0583FDFE378" box="[348,435,1830,1854]" italics="true" pageId="21" pageNumber="472">Cybaeus</emphasis>
spp.
</taxonomicName>
males from California, left palpus, proximal arm of tegular apophysis, ventral (59
<taxonomicName id="A1957DB8FF8BFFBABB08A0593BF0E378" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1334,1436,1831,1854]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABB08A0593BF0E378" box="[1334,1436,1831,1854]" italics="true" pageId="21" pageNumber="472">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABEA9A0353F6DE325" bold="true" box="[151,257,1867,1891]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABEA9A0353F6DE325" box="[151,257,1867,1891]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
from Santa Barbara, 60
<taxonomicName id="A1957DB8FF8BFFBABC39A0353C17E324" authorityName="Chamberlin" authorityYear="1924" box="[519,635,1866,1890]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8BFFBABC39A0353C17E324" box="[519,635,1866,1890]" italics="true" pageId="21" pageNumber="472">C. tardatus</emphasis>
</taxonomicName>
from Claremont, 61
<taxonomicName id="A1957DB8FF8BFFBABD64A0353DA1E324" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[858,973,1867,1890]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABD64A0353DA1E324" box="[858,973,1867,1890]" italics="true" pageId="21" pageNumber="472">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABDE8A0353A2CE325" bold="true" box="[982,1088,1867,1891]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABDE8A0353A2CE325" box="[982,1088,1867,1891]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm.
</paragraph>
</caption>
<caption id="32EA56B3FF88FFB9BEA9A00B3F60E393" ID-DOI="http://doi.org/10.5281/zenodo.5300667" ID-Zenodo-Dep="5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="22" pageNumber="473" startId="22.[151,250,1909,1933]" targetBox="[186,1407,198,1875]" targetPageId="22">
<paragraph id="662A063BFF88FFB9BEA9A00B3F60E393" blockId="22.[151,1437,1909,2005]" pageId="22" pageNumber="473">
<emphasis id="54E1DA29FF88FFB9BEA9A00B3F4DE3CB" bold="true" box="[151,289,1909,1933]" pageId="22" pageNumber="473">FIGURE 62.</emphasis>
Distribution of
<taxonomicName id="A1957DB8FF88FFB9BFF8A00B3C03E3CB" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[454,623,1909,1933]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="orarius" status="sp. nov.">
<emphasis id="54E1DA29FF88FFB9BFF8A00B3C03E3CB" box="[454,623,1909,1933]" italics="true" pageId="22" pageNumber="473">Cybaeus orarius</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF88FFB9BC48A00B3CB2E3CB" bold="true" box="[630,734,1909,1933]" pageId="22" pageNumber="473">
<taxonomicNameLabel id="4FD26752FF88FFB9BC48A00B3CB2E3CB" box="[630,734,1909,1933]" pageId="22" pageNumber="473" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="A1957DB8FF88FFB9BCD2A0083D3EE3CB" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[748,850,1910,1933]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF88FFB9BCD2A0083D3EE3CB" box="[748,850,1910,1933]" italics="true" pageId="22" pageNumber="473">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF88FFB9BD67A00B3DADE3CB" bold="true" box="[857,961,1909,1933]" pageId="22" pageNumber="473">
<taxonomicNameLabel id="4FD26752FF88FFB9BD67A00B3DADE3CB" box="[857,961,1909,1933]" pageId="22" pageNumber="473" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="A1957DB8FF88FFB9BDF1A0083A2EE3CB" authorityName="Chamberlin" authorityYear="1924" box="[975,1090,1909,1933]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF88FFB9BDF1A0083A2EE3CB" box="[975,1090,1909,1933]" italics="true" pageId="22" pageNumber="473">C. tardatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="A1957DB8FF88FFB9BA44A0083A80E3CB" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1146,1260,1910,1933]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF88FFB9BA44A0083A80E3CB" box="[1146,1260,1910,1933]" italics="true" pageId="22" pageNumber="473">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF88FFB9BACDA00B3B37E3CB" bold="true" box="[1267,1371,1909,1933]" pageId="22" pageNumber="473">
<taxonomicNameLabel id="4FD26752FF88FFB9BACDA00B3B37E3CB" box="[1267,1371,1909,1933]" pageId="22" pageNumber="473" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized.
</paragraph>
</caption>
<paragraph id="662A063BFF89FFB8BEF9A7E93D3DE507" blockId="23.[151,1437,151,970]" pageId="23" pageNumber="474">
Distinguishing the female
<taxonomicName id="A1957DB8FF89FFB8BFC1A7E93CEDE4F6" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[511,641,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF89FFB8BFC1A7E93CEDE4F6" box="[511,641,151,177]" italics="true" pageId="23" pageNumber="474">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF89FFB8BCB3A7E93D69E4F7" bold="true" box="[653,773,151,177]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="4FD26752FF89FFB8BCB3A7E93D69E4F7" box="[653,773,151,177]" pageId="23" pageNumber="474" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
from those of
<taxonomicName id="A1957DB8FF89FFB8BD80A7E93A5AE4F6" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[958,1078,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="orarius" status="sp. nov.">
<emphasis id="54E1DA29FF89FFB8BD80A7E93A5AE4F6" box="[958,1078,151,177]" italics="true" pageId="23" pageNumber="474">C. orarius</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF89FFB8BA7DA7E93AD6E4F7" bold="true" box="[1091,1210,151,177]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="4FD26752FF89FFB8BA7DA7E93AD6E4F7" box="[1091,1210,151,177]" pageId="23" pageNumber="474" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF89FFB8BAC5A7E93B11E4F7" authorityName="Chamberlin" authorityYear="1924" box="[1275,1405,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF89FFB8BAC5A7E93B11E4F7" box="[1275,1405,151,177]" italics="true" pageId="23" pageNumber="474">C. tardatus</emphasis>
</taxonomicName>
is discussed under the diagnoses of those two species. The females of
<taxonomicName id="A1957DB8FF89FFB8BD92A7C53A72E492" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[940,1054,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF89FFB8BD92A7C53A72E492" box="[940,1054,187,213]" italics="true" pageId="23" pageNumber="474">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF89FFB8BA16A7C53AF1E493" bold="true" box="[1064,1181,187,213]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="4FD26752FF89FFB8BA16A7C53AF1E493" box="[1064,1181,187,213]" pageId="23" pageNumber="474" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF89FFB8BAE7A7C53B34E492" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1241,1368,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF89FFB8BAE7A7C53B34E492" box="[1241,1368,187,213]" italics="true" pageId="23" pageNumber="474">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF89FFB8BB5CA7C53EA4E4BF" bold="true" pageId="23" pageNumber="474">
<taxonomicNameLabel id="4FD26752FF89FFB8BB5CA7C53EA4E4BF" pageId="23" pageNumber="474" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
are distinguished by the uniform diameter of the copulatory ducts throughout their length and the unobscured Bennett’s glands (
<figureCitation id="FEAE1ABEFF89FFB8BF61A67D3FB3E55B" box="[351,479,258,285]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="23" pageNumber="474">Figs 52–54</figureCitation>
) in
<taxonomicName id="A1957DB8FF89FFB8BC37A67D3CEAE55A" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[521,646,259,285]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="topanga">
<emphasis id="54E1DA29FF89FFB8BC37A67D3CEAE55A" box="[521,646,259,285]" italics="true" pageId="23" pageNumber="474">C. topanga</emphasis>
</taxonomicName>
(
<emphasis id="54E1DA29FF89FFB8BCABA67D3CB1E55A" box="[661,733,259,284]" italics="true" pageId="23" pageNumber="474">versus</emphasis>
copulatory ducts broadening posteriorly and usually obscuring the Bennett’s glands [
<figureCitation id="FEAE1ABEFF89FFB8BFB6A6593C58E507" box="[392,564,294,321]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="23" pageNumber="474">Figs 42–43, 45</figureCitation>
] in
<taxonomicName id="A1957DB8FF89FFB8BC61A6593CA3E506" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[607,719,295,321]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF89FFB8BC61A6593CA3E506" box="[607,719,295,321]" italics="true" pageId="23" pageNumber="474">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF89FFB8BCE8A6593D24E507" bold="true" box="[726,840,295,321]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="4FD26752FF89FFB8BCE8A6593D24E507" box="[726,840,295,321]" pageId="23" pageNumber="474" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
)
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF89FFB8BEF9A6343F39E7C7" pageId="23" pageNumber="474" type="description">
<paragraph id="662A063BFF89FFB8BEF9A6343A3FE523" blockId="23.[151,1437,151,970]" box="[199,1107,330,357]" pageId="23" pageNumber="474">
<emphasis id="54E1DA29FF89FFB8BEF9A6343F35E522" bold="true" box="[199,345,330,356]" pageId="23" pageNumber="474">Description.</emphasis>
Tibiae and femora banded. Ventral tibia I macrosetae 2-1p-2-1p-
<specimenCount id="7093CDB2FF89FFB8BA0AA6353A3FE523" box="[1076,1107,331,357]" pageId="23" pageNumber="474" type="generic">1p.</specimenCount>
</paragraph>
<paragraph id="662A063BFF89FFB8BEF9A6103A03E597" blockId="23.[151,1437,151,970]" pageId="23" pageNumber="474">
<emphasis id="54E1DA29FF89FFB8BEF9A6103F67E5CF" bold="true" box="[199,267,366,393]" pageId="23" pageNumber="474">
<emphasis id="54E1DA29FF89FFB8BEF9A6103F6EE5CF" bold="true" box="[199,258,366,393]" italics="true" pageId="23" pageNumber="474">Male</emphasis>
:
</emphasis>
(n=1). Patellar apophysis (
<figureCitation id="FEAE1ABEFF89FFB8BC01A6113CFEE5CF" box="[575,658,366,393]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1891,1915]" captionTargetBox="[151,1417,191,1867]" captionTargetId="figure-31@20.[151,1436,181,1867]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="23" pageNumber="474">Fig. 56</figureCitation>
) anteriorly directed with 25–30 peg setae. Carinate retrolateral tibial apophysis (
<figureCitation id="FEAE1ABEFF89FFB8BF29A6ED3F04E5EB" box="[279,360,402,429]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1891,1915]" captionTargetBox="[151,1417,191,1867]" captionTargetId="figure-31@20.[151,1436,181,1867]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="23" pageNumber="474">Fig. 55</figureCitation>
) nearly as long as tibia. Distal arm of tegular apophysis (
<figureCitation id="FEAE1ABEFF89FFB8BDD6A6ED3A56E5EB" box="[1000,1082,402,429]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1891,1915]" captionTargetBox="[151,1417,191,1867]" captionTargetId="figure-31@20.[151,1436,181,1867]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="23" pageNumber="474">Fig. 57</figureCitation>
) broad, elongate; proximal arm (
<figureCitation id="FEAE1ABEFF89FFB8BEA1A6C93E9AE597" box="[159,246,438,465]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1891,1915]" captionTargetBox="[151,1417,191,1867]" captionTargetId="figure-31@20.[151,1436,181,1867]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="23" pageNumber="474">Figs 57</figureCitation>
,
<figureCitation id="FEAE1ABEFF89FFB8BF3CA6C93F73E594" box="[258,287,439,466]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="23" pageNumber="474">61</figureCitation>
) with ventral and dorsal tips triangular, bluntly acuminate, basally directed.
</paragraph>
<paragraph id="662A063BFF89FFB8BEF9A6A53DD9E5B3" blockId="23.[151,1437,151,970]" box="[199,949,475,501]" pageId="23" pageNumber="474">
<emphasis id="54E1DA29FF89FFB8BEF9A6A53F6AE5B3" bold="true" box="[199,262,475,501]" pageId="23" pageNumber="474">Note.</emphasis>
<typeStatus id="B92EB899FF89FFB8BF33A6A53F19E5B3" box="[269,373,475,501]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
embolus apparently broken, lacking apical section.
</paragraph>
<paragraph id="662A063BFF89FFB8BEF9A6813D36E65F" blockId="23.[151,1437,151,970]" box="[199,858,510,537]" pageId="23" pageNumber="474">Measurements (n=1). CL 2.8, CW 2.03, SL 1.35, SW 1.23.</paragraph>
<paragraph id="662A063BFF89FFB8BEF9A55C3B3AE68B" blockId="23.[151,1437,151,970]" pageId="23" pageNumber="474">
<emphasis id="54E1DA29FF89FFB8BEF9A55C3F4BE67B" bold="true" box="[199,295,546,573]" pageId="23" pageNumber="474">
<emphasis id="54E1DA29FF89FFB8BEF9A55C3F72E67B" bold="true" box="[199,286,546,573]" italics="true" pageId="23" pageNumber="474">Female</emphasis>
:
</emphasis>
(n=3). Atrium (
<figureCitation id="FEAE1ABEFF89FFB8BFE9A55D3C3AE67B" box="[471,598,546,573]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="23" pageNumber="474">Figs 52–53</figureCitation>
) hooded anteriorly with area between atrial openings inverted vase-shaped, widest posteriorly, concave. Vulva (
<figureCitation id="FEAE1ABEFF89FFB8BC0FA5393CD8E627" box="[561,692,582,609]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="23" pageNumber="474">Figs 53–54</figureCitation>
) with copulatory ducts opening into atrium laterally from vulva, contiguous only at anterior end of atrium, diverging posteriorly then converging and turning anteriorly at posterior margin of vulva; spermathecal heads small dorsal lobes located medially on vulva, visible in dorsal view; spermathecal stalks narrow, relatively long. Bennett’s glands located dorso-laterally on small, rounded spermathecal bases.
</paragraph>
<paragraph id="662A063BFF89FFB8BEF9A5A93B54E6B7" blockId="23.[151,1437,151,970]" box="[199,1336,726,753]" pageId="23" pageNumber="474">Measurements (n=3). CL 3.1, 3.2, 3.3; CW 2.08, 2.25, 2.38; SL 1.43, 1.50, 1.58; SW 1.38, 1.48, 1.48.</paragraph>
<paragraph id="662A063BFF89FFB8BEF9A5853F39E7C7" blockId="23.[151,1437,151,970]" pageId="23" pageNumber="474">
The male came from the Los Angeles area, the females from near the Mexican border about
<quantity id="A16DABDEFF89FFB8BAC4A5843B3DE753" box="[1274,1361,762,789]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="474" unit="km" value="250.0">250 km</quantity>
to the south. Possibly they are representatives of two closely related species. In the absence of opposite sexes from both localities and in an effort to reduce the possibility of future synonymy this male and these females are described as one new species.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF89FFB8BEF9A4F43A64E78F" pageId="23" pageNumber="474" type="distribution">
<paragraph id="662A063BFF89FFB8BEF9A4F43A64E78F" blockId="23.[151,1437,151,970]" pageId="23" pageNumber="474">
<emphasis id="54E1DA29FF89FFB8BEF9A4F43C3FE7E2" bold="true" box="[199,595,906,933]" pageId="23" pageNumber="474">Distribution and natural history.</emphasis>
(
<figureCitation id="FEAE1ABEFF89FFB8BC5AA4F53CD6E7E0" box="[612,698,907,934]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="23" pageNumber="474">Fig. 62</figureCitation>
). Known only from the type locality and San Ysidro,
<collectingRegion id="A451C8D9FF89FFB8BB1AA4F53BF5E7E3" box="[1316,1433,907,933]" country="United States of America" name="California" pageId="23" pageNumber="474">California</collectingRegion>
. The single known male (the
<typeStatus id="B92EB899FF89FFB8BFE9A4D13C57E78C" box="[471,571,943,970]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
) was collected in June from under a rock
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="662A063BFF8DFFBCBEA9A7E93C77E4F7" blockId="19.[151,539,151,214]" box="[151,539,151,178]" pageId="19" pageNumber="470">
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<emphasis id="54E1DA29FF8DFFBCBEA9A7E93F04E4F4" bold="true" box="[151,360,151,178]" italics="true" pageId="19" pageNumber="470">Cybaeus tardatus</emphasis>
(Chamberlin)
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<subSubSection id="2E8F55B0FF8DFFBCBEA9A7C53F04E493" box="[151,360,187,214]" pageId="19" pageNumber="470" type="description">
<paragraph id="662A063BFF8DFFBCBEA9A7C53F04E493" blockId="19.[151,539,151,214]" box="[151,360,187,214]" pageId="19" pageNumber="470">
<figureCitation id="FEAE1ABEFF8DFFBCBEA9A7C53F7BE490" box="[151,279,187,214]" captionStart-0="FIGURES 46–49" captionStart-1="FIGURES 50–54" captionStartId-0="18.[151,264,1929,1953]" captionStartId-1="19.[151,264,1939,1963]" captionTargetBox-0="[164,1433,185,1891]" captionTargetBox-1="[157,1429,689,1916]" captionTargetId-0="figure-31@18.[151,1436,181,1905]" captionTargetId-1="figure-163@19.[151,1436,684,1916]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." captionText-1="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.5300659" figureDoi-1="http://doi.org/10.5281/zenodo.5300661" httpUri-0="https://zenodo.org/record/5300659/files/figure.png" httpUri-1="https://zenodo.org/record/5300661/files/figure.png" pageId="19" pageNumber="470">Figs 46–51</figureCitation>
,
<figureCitation id="FEAE1ABEFF8DFFBCBF1CA7C53F2CE493" box="[290,320,187,213]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="19" pageNumber="470">60</figureCitation>
,
<figureCitation id="FEAE1ABEFF8DFFBCBF72A7C53F04E493" box="[332,360,187,213]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="19" pageNumber="470">62</figureCitation>
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<paragraph id="662A063BFF8DFFBCBEA9A67E3CE9E55E" blockId="19.[151,1437,256,372]" box="[151,645,256,280]" pageId="19" pageNumber="470">
<treatmentCitationGroup id="46852115FF8DFFBCBEA9A67E3CE9E55E" box="[151,645,256,280]" pageId="19" pageNumber="470">
<treatmentCitation id="E734202AFF8DFFBCBEA9A67E3C52E55E" author="Chamberlin, R. V." box="[151,574,256,280]" page="3" pageId="19" pageNumber="470" year="1919">
<taxonomicName id="A1957DB8FF8DFFBCBEA9A67E3C52E55E" authority="Chamberlin 1919: 3" authorityName="Chamberlin" authorityPageNumber="3" authorityYear="1919" box="[151,574,256,280]" class="Arachnida" family="Cybaeidae" genus="Parauximus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8DFFBCBEA9A67E3F00E55E" box="[151,364,256,280]" italics="true" pageId="19" pageNumber="470">Parauximus tardatus</emphasis>
<bibRefCitation id="02047BCAFF8DFFBCBF4CA67E3C52E55E" author="Chamberlin, R. V." box="[370,574,256,280]" pageId="19" pageNumber="470" pagination="1 - 26" refId="ref14974" refString="Chamberlin, R. V. (1919) New Californian spiders. Journal of Entomology and Zoology, (Claremont) 12, 1 - 26. https: // doi. org / 10.1093 / aesa / 12.3.239" type="journal article" year="1919">Chamberlin 1919: 3</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="662A063BFF8DFFBCBEA9A6603C1BE570" blockId="19.[151,1437,256,372]" box="[151,631,286,310]" pageId="19" pageNumber="470">
<treatmentCitationGroup id="46852115FF8DFFBCBEA9A6603C1BE570" box="[151,631,286,310]" pageId="19" pageNumber="470">
<treatmentCitation id="E734202AFF8DFFBCBEA9A6603C5DE570" author="Chamberlin, R. V." box="[151,561,286,310]" page="14" pageId="19" pageNumber="470" year="1919">
<taxonomicName id="A1957DB8FF8DFFBCBEA9A6603C5DE570" authority="Chamberlin 1919: 14" authorityName="Chamberlin" authorityPageNumber="14" authorityYear="1919" box="[151,561,286,310]" class="Arachnida" family="Cybaeidae" genus="Namopsilus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="pletus">
<emphasis id="54E1DA29FF8DFFBCBEA9A6603F3EE570" box="[151,338,286,310]" italics="true" pageId="19" pageNumber="470">Namopsilus pletus</emphasis>
<bibRefCitation id="02047BCAFF8DFFBCBF66A6603C5DE570" author="Chamberlin, R. V." box="[344,561,286,310]" pageId="19" pageNumber="470" pagination="1 - 26" refId="ref14974" refString="Chamberlin, R. V. (1919) New Californian spiders. Journal of Entomology and Zoology, (Claremont) 12, 1 - 26. https: // doi. org / 10.1093 / aesa / 12.3.239" type="journal article" year="1919">Chamberlin 1919: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="662A063BFF8DFFBCBEA9A6433A69E532" blockId="19.[151,1437,256,372]" pageId="19" pageNumber="470">
<treatmentCitationGroup id="46852115FF8DFFBCBEA9A6433A69E532" pageId="19" pageNumber="470">
<treatmentCitation id="E734202AFF8DFFBCBEA9A6433C58E513" author="Chamberlin, R. V." box="[151,564,317,341]" page="669" pageId="19" pageNumber="470" year="1924">
<taxonomicName id="A1957DB8FF8DFFBCBEA9A6433C58E513" ID-CoL="32RHW" authority="Chamberlin 1924: 669" authorityName="Chamberlin" authorityPageNumber="669" authorityYear="1924" box="[151,564,317,341]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8DFFBCBEA9A6433F24E513" box="[151,328,317,341]" italics="true" pageId="19" pageNumber="470">Cybaeus tardatus</emphasis>
<bibRefCitation id="02047BCAFF8DFFBCBF70A6433C58E513" author="Chamberlin, R. V." box="[334,564,317,341]" pageId="19" pageNumber="470" pagination="561 - 694" refId="ref15014" refString="Chamberlin, R. V. (1924) The spider fauna of the shores and islands of the Gulf of California. Proceedings of the California Academy of Sciences, 12, 561 - 694." type="journal article" year="1924">Chamberlin 1924: 669</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Chamberlin and Ivie 1932: 23, fig. 56. Chamberlin and Ivie 1937: 225.
<treatmentCitation id="E734202AFF8DFFBCBB30A6433E8FE532" author="Roewer, C. F." page="92" pageId="19" pageNumber="470" year="1954">
<bibRefCitation id="02047BCAFF8DFFBCBB30A6433E8FE532" author="Roewer, C. F." pageId="19" pageNumber="470" refId="ref17427" refString="Roewer, C. F. (1954) Katalog der Araneae 2 a. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 923 pp." type="book" year="1954">Roewer 1954: 92</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="E734202AFF8DFFBCBED3A6223FDFE532" author="Bonnet, P." box="[237,435,347,372]" page="1304" pageId="19" pageNumber="470" year="1956">
<bibRefCitation id="02047BCAFF8DFFBCBED3A6223FDFE532" author="Bonnet, P." box="[237,435,347,372]" pageId="19" pageNumber="470" pagination="919 - 1926" refId="ref14738" refString="Bonnet, P. (1956) Bibliographia Araneorum. Douladoure, Toulouse, Tome 2, pp. 919 - 1926." type="journal article" year="1956">Bonnet 1956: 1304</bibRefCitation>
</treatmentCitation>
. Roth and Brown 1986: 4, 12.
<bibRefCitation id="02047BCAFF8DFFBCBCD7A6223A6DE532" author="World Spider Catalog" box="[745,1025,348,372]" pageId="19" pageNumber="470" refId="ref18189" refString="World Spider Catalog (2021) World Spider Catalog. Version 22.0. Natural History Museum, Bern. Available from: https: // wsc. nmbe. ch / (accessed 15 June 2021) https: // doi. org / 10.24436 / 2" type="book" year="2021">World Spider Catalog 2021</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8DFFBCBEA9A6DC3C02E60A" pageId="19" pageNumber="470" type="materials_examined">
<paragraph id="662A063BFF8DFFBCBEA9A6DC3F98E642" blockId="19.[151,1437,417,661]" pageId="19" pageNumber="470">
<emphasis id="54E1DA29FF8DFFBCBEA9A6DC3F2BE5FD" bold="true" box="[151,327,417,444]" pageId="19" pageNumber="470">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF8DFFBCBF6FA6DF3F01E5A6" collectionCode="MCZ" collectorName="Claremont &amp; R. V. Chamberlin" country="United States of America" county="Los Angeles County" location="Los Angeles County" pageId="19" pageNumber="470" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF8DFFBCBF6FA6DF3C9DE5FA" bold="true" box="[337,753,417,444]" pageId="19" pageNumber="470">
<collectingCountry id="1E8246ABFF8DFFBCBF6FA6DF3FCDE5FD" box="[337,417,417,443]" name="United States of America" pageId="19" pageNumber="470">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF8DFFBCBF8EA6DF3C44E5FA" bold="true" box="[432,552,417,444]" italics="true" pageId="19" pageNumber="470">
<collectingRegion id="A451C8D9FF8DFFBCBF8EA6DF3C44E5FA" box="[432,552,417,444]" country="United States of America" name="California" pageId="19" pageNumber="470">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF8DFFBCBC05A6DC3CC4E5FA" box="[571,680,418,444]" pageId="19" pageNumber="470" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<collectingCounty id="8F4B7EB7FF8DFFBCBCC5A6DC3D8AE5FA" box="[763,998,418,444]" pageId="19" pageNumber="470">Los Angeles County</collectingCounty>
,
<collectorName id="CB6063EDFF8DFFBCBDCDA6DC3A00E5FA" box="[1011,1132,418,444]" pageId="19" pageNumber="470">Claremont</collectorName>
, no date,
<collectorName id="CB6063EDFF8DFFBCBAE3A6DC3BF0E5FA" box="[1245,1436,418,444]" pageId="19" pageNumber="470">R.V. Chamberlin</collectorName>
(
<collectionCode id="00849EFEFF8DFFBCBEA1A6B83E8DE5A6" box="[159,225,454,480]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="19" pageNumber="470" type="Museum">MCZ</collectionCode>
), examined
</materialsCitation>
.
<materialsCitation id="D6FD0C66FF8DFFBCBF40A6B83F9DE642" collectingDate="1913-04" collectionCode="MCZ" collectorName="Claremont &amp; R. V. Chamberlin" country="United States of America" county="Los Angeles County" location="Los Angeles County" pageId="19" pageNumber="470" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF8DFFBCBF40A6B83C25E599" bold="true" box="[382,585,453,480]" pageId="19" pageNumber="470">
<typeStatus id="B92EB899FF8DFFBCBF40A6B83F87E5A6" box="[382,491,454,480]" pageId="19" pageNumber="470" type="holotype">Holotype</typeStatus>
female
</emphasis>
of
<emphasis id="54E1DA29FF8DFFBCBC40A6BB3D0DE5A6" bold="true" box="[638,865,453,480]" pageId="19" pageNumber="470">
<taxonomicName id="A1957DB8FF8DFFBCBC40A6BB3D35E5A6" authorityName="Chamberlin" authorityYear="1919" box="[638,857,453,480]" class="Arachnida" family="Cybaeidae" genus="Namopsilus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="pletus">
<emphasis id="54E1DA29FF8DFFBCBC40A6BB3D35E5A6" bold="true" box="[638,857,453,480]" italics="true" pageId="19" pageNumber="470">Namopsilus pletus</emphasis>
</taxonomicName>
.
</emphasis>
<collectingCounty id="8F4B7EB7FF8DFFBCBD51A6B83A08E5A6" box="[879,1124,454,480]" pageId="19" pageNumber="470">Los Angeles County</collectingCounty>
,
<collectorName id="CB6063EDFF8DFFBCBA49A6B83A9CE5A6" box="[1143,1264,454,480]" pageId="19" pageNumber="470">Claremont</collectorName>
,
<date id="122B20FBFF8DFFBCBB3CA6B83B34E5A6" box="[1282,1368,454,480]" pageId="19" pageNumber="470" value="1913-04">
<collectingDate id="026FD913FF8DFFBCBB3CA6B83B34E5A6" box="[1282,1368,454,480]" pageId="19" pageNumber="470" value="1913-04">iv.1913</collectingDate>
</date>
,
<collectorName id="CB6063EDFF8DFFBCBB55A6B83F77E642" pageId="19" pageNumber="470">R.V. Chamberlin</collectorName>
(
<collectionCode id="00849EFEFF8DFFBCBF14A6943F00E642" box="[298,364,490,516]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="19" pageNumber="470" type="Museum">MCZ</collectionCode>
), examined
</materialsCitation>
.
</paragraph>
<paragraph id="662A063BFF8DFFBCBEF9A5733C02E60A" blockId="19.[151,1437,417,661]" pageId="19" pageNumber="470">
<emphasis id="54E1DA29FF8DFFBCBEF9A5733F97E661" bold="true" box="[199,507,525,551]" pageId="19" pageNumber="470">Other material examined.</emphasis>
<materialsCitation id="D6FD0C66FF8DFFBCBC3FA5733C06E60A" collectingDate="1945-06" collectionCode="MCZ, AMNH" collectorName="S. C. Bishop" country="United States of America" location="Claremont" pageId="19" pageNumber="470" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="California">
<emphasis id="54E1DA29FF8DFFBCBC3FA5733CB3E66E" bold="true" box="[513,735,525,552]" pageId="19" pageNumber="470">
<collectingCountry id="1E8246ABFF8DFFBCBC3FA5733C3EE661" box="[513,594,525,551]" name="United States of America" pageId="19" pageNumber="470">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF8DFFBCBC60A5733CBAE66E" bold="true" box="[606,726,525,552]" italics="true" pageId="19" pageNumber="470">
<collectingRegion id="A451C8D9FF8DFFBCBC60A5733CBAE66E" box="[606,726,525,552]" country="United States of America" name="California" pageId="19" pageNumber="470">California</collectingRegion>
</emphasis>
:
</emphasis>
Los Angeles.
<specimenCount id="7093CDB2FF8DFFBCBD40A5703DCEE66E" box="[894,930,526,552]" pageId="19" pageNumber="470" type="male">1♂</specimenCount>
,
<location id="634A50E0FF8DFFBCBD90A5703A4BE66E" LSID="urn:lsid:plazi:treatment:EE3CB72DFF8DFFBABEA9A7E93D4BE7C7:634A50E0FF8DFFBCBD90A5703A4BE66E" box="[942,1063,526,552]" country="United States of America" name="Claremont" pageId="19" pageNumber="470" stateProvince="California">Claremont</location>
, no date, no collector (
<collectionCode id="00849EFEFF8DFFBCBB1AA5703B0BE66E" box="[1316,1383,526,552]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="19" pageNumber="470" type="Museum">MCZ</collectionCode>
);
<specimenCount id="7093CDB2FF8DFFBCBB47A5703BF0E66E" box="[1401,1436,526,552]" pageId="19" pageNumber="470" type="female">1♀</specimenCount>
Pasadena,
<date id="122B20FBFF8DFFBCBF33A54C3F09E60B" box="[269,357,562,589]" pageId="19" pageNumber="470" value="1945-06">
<collectingDate id="026FD913FF8DFFBCBF33A54C3F09E60B" box="[269,357,562,589]" pageId="19" pageNumber="470" value="1945-06">vi.1945</collectingDate>
</date>
,
<collectorName id="CB6063EDFF8DFFBCBF4EA54C3F9AE60A" box="[368,502,562,588]" pageId="19" pageNumber="470">S.C. Bishop</collectorName>
(
<collectionCode id="00849EFEFF8DFFBCBC38A54C3C0EE60A" box="[518,610,562,588]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="470">AMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8DFFBABEF9A52B3D83E5CF" lastPageId="21" lastPageNumber="472" pageId="19" pageNumber="470" type="diagnosis">
<paragraph id="662A063BFF8DFFBCBEF9A52B3B01E6D2" blockId="19.[151,1437,417,661]" pageId="19" pageNumber="470">
<emphasis id="54E1DA29FF8DFFBCBEF9A52B3F2EE629" bold="true" box="[199,322,597,623]" pageId="19" pageNumber="470">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF8DFFBCBFDEA5283C33E636" authorityName="Chamberlin" authorityYear="1924" box="[480,607,598,624]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8DFFBCBFDEA5283C33E636" box="[480,607,598,624]" italics="true" pageId="19" pageNumber="470">C. tardatus</emphasis>
</taxonomicName>
is readily diagnosed by the form of the proximal and distal arms of the tegular apophysis (
<figureCitation id="FEAE1ABEFF8DFFBCBF55A5043FAEE6D2" box="[363,450,634,660]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="19" pageNumber="470">Figs 48</figureCitation>
,
<figureCitation id="FEAE1ABEFF8DFFBCBFF0A5043F82E6D3" box="[462,494,634,661]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="19" pageNumber="470">60</figureCitation>
); these characteristics are discussed under the diagnosis of
<taxonomicName id="A1957DB8FF8DFFBCBABAA5043A9FE6D2" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1156,1267,634,660]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF8DFFBCBABAA5043A9FE6D2" box="[1156,1267,634,660]" italics="true" pageId="19" pageNumber="470">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8DFFBCBAC5A5043B01E6D2" bold="true" box="[1275,1389,634,660]" pageId="19" pageNumber="470">
<taxonomicNameLabel id="4FD26752FF8DFFBCBAC5A5043B01E6D2" box="[1275,1389,634,660]" pageId="19" pageNumber="470" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<caption id="32EA56B3FF8DFFBCBEA9A0ED3DAAE3B5" ID-DOI="http://doi.org/10.5281/zenodo.5300661" ID-Zenodo-Dep="5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="19" pageNumber="470" startId="19.[151,264,1939,1963]" targetBox="[157,1429,689,1916]" targetPageId="19">
<paragraph id="662A063BFF8DFFBCBEA9A0ED3DAAE3B5" blockId="19.[151,1436,1939,2036]" pageId="19" pageNumber="470">
<emphasis id="54E1DA29FF8DFFBCBEA9A0ED3F39E3ED" bold="true" box="[151,341,1939,1963]" pageId="19" pageNumber="470">FIGURES 50–54.</emphasis>
<taxonomicName id="A1957DB8FF8DFFBCBF63A0ED3F8BE3ED" box="[349,487,1939,1963]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="54E1DA29FF8DFFBCBF63A0ED3FD8E3ED" box="[349,436,1939,1963]" italics="true" pageId="19" pageNumber="470">Cybaeus</emphasis>
spp.
</taxonomicName>
, females, copulatory organ (50–51
<taxonomicName id="A1957DB8FF8DFFBCBD6DA0EA3DAAE3ED" authorityName="Chamberlin" authorityYear="1924" box="[851,966,1939,1963]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8DFFBCBD6DA0EA3DAAE3ED" box="[851,966,1939,1963]" italics="true" pageId="19" pageNumber="470">C. tardatus</emphasis>
</taxonomicName>
, holotype of
<taxonomicName id="A1957DB8FF8DFFBCBA6EA0ED3B60E3ED" authorityName="Chamberlin" authorityYear="1919" box="[1104,1292,1939,1963]" class="Arachnida" family="Cybaeidae" genus="Namopsilus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="pletus">
<emphasis id="54E1DA29FF8DFFBCBA6EA0ED3B60E3ED" box="[1104,1292,1939,1963]" italics="true" pageId="19" pageNumber="470">Namopsilus pletus</emphasis>
</taxonomicName>
; 52–54
<taxonomicName id="A1957DB8FF8DFFBCBB5FA0EA3EBAE389" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF8DFFBCBB5FA0EA3EBAE389" italics="true" pageId="19" pageNumber="470">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8DFFBCBEE2A0C63F2EE396" bold="true" box="[220,322,1976,2000]" pageId="19" pageNumber="470">
<taxonomicNameLabel id="4FD26752FF8DFFBCBEE2A0C63F2EE396" box="[220,322,1976,2000]" pageId="19" pageNumber="470" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca.
</paragraph>
</caption>
<caption id="32EA56B3FF8AFFBBBEA9A01D3AC0E382" ID-DOI="http://doi.org/10.5281/zenodo.5300663" ID-Zenodo-Dep="5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="20" pageNumber="471" startId="20.[151,264,1891,1915]" targetBox="[151,1417,191,1867]" targetPageId="20">
<paragraph id="662A063BFF8AFFBBBEA9A01D3AC0E382" blockId="20.[151,1437,1891,1988]" pageId="20" pageNumber="471">
<emphasis id="54E1DA29FF8AFFBBBEA9A01D3F38E33D" bold="true" box="[151,340,1891,1915]" pageId="20" pageNumber="471">FIGURES 55–58.</emphasis>
<taxonomicName id="A1957DB8FF8AFFBBBF65A01D3C67E33D" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[347,523,1891,1915]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF8AFFBBBF65A01D3C67E33D" box="[347,523,1891,1915]" italics="true" pageId="20" pageNumber="471">Cybaeus topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8AFFBBBC2CA01A3C15E33A" bold="true" box="[530,633,1892,1916]" pageId="20" pageNumber="471">
<taxonomicNameLabel id="4FD26752FF8AFFBBBC2CA01A3C15E33A" box="[530,633,1892,1916]" pageId="20" pageNumber="471" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral). E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA.
</paragraph>
</caption>
<paragraph id="662A063BFF8BFFBABEF9A7E93D83E5CF" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="472">
Distinguishing the females of
<taxonomicName id="A1957DB8FF8BFFBABC1AA7E93CF5E4F6" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[548,665,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="orarius" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABC1AA7E93CF5E4F6" box="[548,665,151,177]" italics="true" pageId="21" pageNumber="472">C. orarius</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABC9AA7E93D75E4F7" bold="true" box="[676,793,151,177]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABC9AA7E93D75E4F7" box="[676,793,151,177]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF8BFFBABD6BA7E93DB8E4F7" authorityName="Chamberlin" authorityYear="1924" box="[853,980,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8BFFBABD6BA7E93DB8E4F7" box="[853,980,151,177]" italics="true" pageId="21" pageNumber="472">C. tardatus</emphasis>
</taxonomicName>
is discussed under the diagnosis of the former. From the other females in the
<taxonomicName id="A1957DB8FF8BFFBABC6FA7C53CC1E493" authorityName="Chamberlin" authorityYear="1924" box="[593,685,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8BFFBABC6FA7C53CC1E493" box="[593,685,187,213]" italics="true" pageId="21" pageNumber="472">tardatus</emphasis>
</taxonomicName>
species group with rounded spermathecal bases (
<taxonomicName id="A1957DB8FF8BFFBABAD9A7C53B34E492" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1255,1368,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABAD9A7C53B34E492" box="[1255,1368,187,213]" italics="true" pageId="21" pageNumber="472">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABB5CA7C53EA4E4BF" bold="true" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABB5CA7C53EA4E4BF" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="A1957DB8FF8BFFBABEE9A7A13F39E4BE" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[215,341,223,249]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABEE9A7A13F39E4BE" box="[215,341,223,249]" italics="true" pageId="21" pageNumber="472">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABF63A7A13FBCE4BF" bold="true" box="[349,464,223,249]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABF63A7A13FBCE4BF" box="[349,464,223,249]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
) the female of
<taxonomicName id="A1957DB8FF8BFFBABCBEA7A13C91E4BF" authorityName="Chamberlin" authorityYear="1924" box="[640,765,223,249]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8BFFBABCBEA7A13C91E4BF" box="[640,765,223,249]" italics="true" pageId="21" pageNumber="472">C. tardatus</emphasis>
</taxonomicName>
is distinguished by the nearly parallel-sided atrial area, the copulatory ducts contiguous over much of their length, as well as the general simplicity of the vulval ducts (
<figureCitation id="FEAE1ABEFF8BFFBABB54A67D3EB3E507" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="21" pageNumber="472">Figs 50–51</figureCitation>
) (
<emphasis id="54E1DA29FF8BFFBABEC9A6593F53E506" box="[247,319,295,320]" italics="true" pageId="21" pageNumber="472">versus</emphasis>
atrial area inverted vase-shaped with sides diverging posteriorly [
<figureCitation id="FEAE1ABEFF8BFFBABA1EA6593A16E507" box="[1056,1146,295,321]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="21" pageNumber="472">Figs 42</figureCitation>
,
<figureCitation id="FEAE1ABEFF8BFFBABABBA6583ABCE507" box="[1157,1232,294,321]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="21" pageNumber="472">52–53</figureCitation>
], copulatory ducts non-contiguous [
<figureCitation id="FEAE1ABEFF8BFFBABF6BA6353FC6E523" box="[341,426,330,357]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="21" pageNumber="472">Fig. 43</figureCitation>
] or contiguous only along anterior-most section [
<figureCitation id="FEAE1ABEFF8BFFBABDE2A6353A5BE523" box="[988,1079,331,357]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="21" pageNumber="472">Figs 42</figureCitation>
,
<figureCitation id="FEAE1ABEFF8BFFBABA7DA6343A0FE523" box="[1091,1123,330,357]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="21" pageNumber="472">54</figureCitation>
], and vulval ducts complex [
<figureCitation id="FEAE1ABEFF8BFFBABEA0A6113F26E5CF" box="[158,330,366,393]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="21" pageNumber="472">Figs 42–43, 45</figureCitation>
,
<figureCitation id="FEAE1ABEFF8BFFBABF68A6103FF2E5CF" box="[342,414,366,393]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="21" pageNumber="472">53–54</figureCitation>
] in
<taxonomicName id="A1957DB8FF8BFFBABFF7A6113C55E5CE" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[457,569,367,393]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABFF7A6113C55E5CE" box="[457,569,367,393]" italics="true" pageId="21" pageNumber="472">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABC7EA6113CDEE5CF" bold="true" box="[576,690,367,393]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABC7EA6113CDEE5CF" box="[576,690,367,393]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF8BFFBABCD7A6113D09E5CE" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[745,869,367,393]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF8BFFBABCD7A6113D09E5CE" box="[745,869,367,393]" italics="true" pageId="21" pageNumber="472">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8BFFBABD52A6113DB3E5CF" bold="true" box="[876,991,367,393]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="4FD26752FF8BFFBABD52A6113DB3E5CF" box="[876,991,367,393]" pageId="21" pageNumber="472" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8BFFBABEF9A6EC3E81E753" pageId="21" pageNumber="472" type="description">
<paragraph id="662A063BFF8BFFBABEF9A6EC3AFCE5EB" blockId="21.[151,1437,151,897]" box="[199,1168,402,429]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A6EC3F35E5EA" bold="true" box="[199,345,402,428]" pageId="21" pageNumber="472">Description.</emphasis>
Ventral tibia I macrosetae: 2-1p-2-1p-2. Femora and tibia lightly banded.
</paragraph>
<paragraph id="662A063BFF8BFFBABEF9A6C83AC1E5B0" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A6C83F67E597" bold="true" box="[199,267,438,465]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A6C83F6EE597" bold="true" box="[199,258,438,465]" italics="true" pageId="21" pageNumber="472">Male</emphasis>
:
</emphasis>
(n=2). About 40 peg setae on strongly anteriorly directed patellar apophysis (
<figureCitation id="FEAE1ABEFF8BFFBABA4EA6C93AA8E597" box="[1136,1220,439,465]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="21" pageNumber="472">Fig. 47</figureCitation>
). Retrolateral tibial apophysis about 2/3 the length of the tibia, not extending to anterior margin of tibia (
<figureCitation id="FEAE1ABEFF8BFFBABA72A6A53ACCE5B0" box="[1100,1184,475,502]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="21" pageNumber="472">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="662A063BFF8BFFBABEF9A6813B54E65F" blockId="21.[151,1437,151,897]" box="[199,1336,510,537]" pageId="21" pageNumber="472">
Measurements (n=2). CL 3.2, 3.3; CW 2.50, 2.50; SL 1.55, 1.53; SW 1.48, 1.48.
<typeStatus id="B92EB899FF8BFFBABA6FA6813AD5E65F" box="[1105,1209,511,537]" pageId="21" pageNumber="472" type="holotype">Holotype</typeStatus>
listed first.
</paragraph>
<paragraph id="662A063BFF8BFFBABEF9A55C3B43E68B" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A55C3F4BE67B" bold="true" box="[199,295,546,573]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A55C3F72E67B" bold="true" box="[199,286,546,573]" italics="true" pageId="21" pageNumber="472">Female</emphasis>
:
</emphasis>
(n=2). Atrium (
<figureCitation id="FEAE1ABEFF8BFFBABFE7A55D3C47E67B" box="[473,555,546,573]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="21" pageNumber="472">Fig. 50</figureCitation>
) with area between atrial openings slightly raised and convex, largely parallelsided but slightly widening posteriorly. Vulva (
<figureCitation id="FEAE1ABEFF8BFFBABC95A5393D40E627" box="[683,812,582,609]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="21" pageNumber="472">Figs 50–51</figureCitation>
) with copulatory ducts opening into atria from midline of vulva, well sclerotized posteriorly, less so anteriorly; heads of spermathecae undifferentiated from adjacent ducts (
<emphasis id="54E1DA29FF8BFFBABE9FA5F13EAFE6EE" box="[161,195,655,680]" italics="true" pageId="21" pageNumber="472">i.e.</emphasis>
not distinct lobes), located near posterior margin of vulva, visible in dorsal view; spermathecal stalks short, thick walled; Bennett’s glands visible in dorsal view on small, thick walled, rounded spermathecal bases.
</paragraph>
<paragraph id="662A063BFF8BFFBABEF9A5A93E81E753" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="472">
Measurements (n=2). CL 2.9, 3.3; CW 2.03, 2.38; SL 1.33, 1.48; SW 1.30, 1.48.
<typeStatus id="B92EB899FF8BFFBABA5CA5A93AA6E6B7" box="[1122,1226,727,753]" pageId="21" pageNumber="472" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="A1957DB8FF8BFFBABACCA5A93B3BE6B7" authorityName="Chamberlin" authorityYear="1919" box="[1266,1367,727,753]" class="Arachnida" family="Cybaeidae" genus="Namopsilus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="pletus">
<emphasis id="54E1DA29FF8BFFBABACCA5A93B3BE6B7" box="[1266,1367,727,753]" italics="true" pageId="21" pageNumber="472">N. pletus</emphasis>
</taxonomicName>
listed second.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8BFFBABEF9A4603D4BE7C7" pageId="21" pageNumber="472" type="distribution">
<paragraph id="662A063BFF8BFFBABEF9A4603D4BE7C7" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="472">
<emphasis id="54E1DA29FF8BFFBABEF9A4603F0DE77E" bold="true" box="[199,353,798,824]" pageId="21" pageNumber="472">Distribution.</emphasis>
(
<figureCitation id="FEAE1ABEFF8BFFBABF4EA4613FA8E77C" box="[368,452,799,826]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="21" pageNumber="472">Fig. 62</figureCitation>
). Los Angeles County,
<collectingRegion id="A451C8D9FF8BFFBABCF6A4613D51E77F" box="[712,829,799,825]" country="United States of America" name="California" pageId="21" pageNumber="472">California</collectingRegion>
. Male specimens listed in Chamberlin and Ivie (1932) from Ensenada, Baja
<collectingRegion id="A451C8D9FF8BFFBABFB7A43D3F97E71B" box="[393,507,835,861]" country="United States of America" name="California" pageId="21" pageNumber="472">California</collectingRegion>
have not been found. We consider Ensenada to be outside the range of
<taxonomicName id="A1957DB8FF8BFFBABB21A43D3BF0E71B" authorityName="Chamberlin" authorityYear="1924" box="[1311,1436,835,861]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF8BFFBABB21A43D3BF0E71B" box="[1311,1436,835,861]" italics="true" pageId="21" pageNumber="472">C. tardatus</emphasis>
</taxonomicName>
and have not included that locality in the distribution map.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="E93AF7A5CFE47FDEF858078C22BD33C2" ID-DOI="10.11646/zootaxa.5026.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300630" ID-ZooBank="511C28F6-07E4-412C-B83E-91D2CAC5AA27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630038704990" checkinUser="plazi" docAuthor="Bennett, Robb, Copley, Claudia &amp; Copley, Darren" docDate="2021" docId="EE3CB72DFF91FFBEBEA9A5843C18E304" docLanguage="en" docName="zootaxa.5026.4.1.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybaeus piazzai Bennett &amp; Copley &amp; Copley 2021, spec. nov." docType="treatment" docVersion="4" lastPageNumber="468" masterDocId="1205CF55FF9EFFAFBE3EA77E3E6CE446" masterDocTitle="Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade" masterLastPageNumber="479" masterPageNumber="451" pageNumber="466" updateTime="1699138999734" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3A057C18A5588410205E1132145926D3">Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade</mods:title>
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<mods:namePart id="3D44E80FCB2248D81C81C7447F141415">Bennett, Robb</mods:namePart>
<mods:affiliation id="650ADC55F096127A31F073C245C9794A">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2.</mods:affiliation>
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<mods:namePart id="5F99F9AD5F64ABFF7367B68F42803338">Copley, Claudia</mods:namePart>
<mods:nameIdentifier id="6AA397C9D0ADDBA0326121E866894FFE" type="ORCID">0000-0002-8184-5819</mods:nameIdentifier>
<mods:affiliation id="4790AE8350E2242D2C2F2E5C957B49B1">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2. &amp; ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
<mods:nameIdentifier id="8DCD1E66560DB0C42A71B0DB8784A439" type="email">ccopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="63BF9C182204362E44B5D04FE96EB1FF">Copley, Darren</mods:namePart>
<mods:nameIdentifier id="A3ADEC20DA2564E900F19ECDB07B2D3C" type="ORCID">0000-0002-1944-4272</mods:nameIdentifier>
<mods:affiliation id="288EEA74703A3C5B2287C32873D31C56">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2. &amp; dcopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 1944 - 4272</mods:affiliation>
<mods:nameIdentifier id="02416EDC72A8CF88D2A4D12BBF1C0A20" type="email">dcopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:date id="BF0DB5B3A8F3584A91EF6C0314398BF1">2021</mods:date>
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<heading id="3D62B157FF91FFA0BEA9A5843C2CE753" bold="true" box="[151,576,762,789]" fontSize="11" level="1" pageId="15" pageNumber="466" reason="1">
<emphasis id="54E1DA29FF91FFA0BEA9A5843C2CE753" bold="true" box="[151,576,762,789]" pageId="15" pageNumber="466">
<taxonomicName id="A1957DB8FF91FFA0BEA9A5843FAEE753" ID-CoL="99CM9" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,450,762,789]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BEA9A5843F34E753" bold="true" box="[151,344,762,789]" italics="true" pageId="15" pageNumber="466">Cybaeus piazzai</emphasis>
Bennett
</taxonomicName>
<taxonomicNameLabel id="4FD26752FF91FFA0BFF7A5853C2CE753" box="[457,576,763,789]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
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<paragraph id="662A063BFF91FFA0BEA9A4613F04E77F" blockId="15.[151,576,762,825]" box="[151,360,798,825]" pageId="15" pageNumber="466">
<figureCitation id="FEAE1ABEFF91FFA0BEA9A4613F7BE77F" box="[151,279,798,825]" captionStart-0="FIGURES 39–41" captionStart-1="FIGURES 42–45" captionStartId-0="16.[151,264,1891,1915]" captionStartId-1="17.[151,264,1402,1426]" captionTargetBox-0="[231,1409,181,1839]" captionTargetBox-1="[151,1435,182,1378]" captionTargetId-0="figure-31@16.[178,1410,181,1868]" captionTargetId-1="figure-21@17.[151,1436,181,1378]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." captionText-1="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi-0="http://doi.org/10.5281/zenodo.5300655" figureDoi-1="http://doi.org/10.5281/zenodo.5300657" httpUri-0="https://zenodo.org/record/5300655/files/figure.png" httpUri-1="https://zenodo.org/record/5300657/files/figure.png" pageId="15" pageNumber="466">Figs 39–45</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BF1CA4603F2CE77F" box="[290,320,798,825]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="15" pageNumber="466">59</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BF72A4613F04E77F" box="[332,360,799,825]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="15" pageNumber="466">62</figureCitation>
</paragraph>
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<subSubSection id="2E8F55B0FF91FFA0BEA9A4193CF7E7AB" pageId="15" pageNumber="466" type="materials_examined">
<paragraph id="662A063BFF91FFA0BEA9A4193CF7E7AB" blockId="15.[151,1437,870,1977]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEA9A4193F25E7C6" bold="true" box="[151,329,870,897]" pageId="15" pageNumber="466">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF91FFA0BF6AA4183CE1E7E3" collectingDate="1967-02-07" collectionCode="CAS" collectorName="Roth" country="United States of America" county="Santa Barbara County" location="Santa Barbara Natural History Museum" pageId="15" pageNumber="466" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF91FFA0BF6AA4183C96E7C7" bold="true" box="[340,762,870,897]" pageId="15" pageNumber="466">
<collectingCountry id="1E8246ABFF91FFA0BF6AA4183FC9E7C6" box="[340,421,870,896]" name="United States of America" pageId="15" pageNumber="466">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF91FFA0BF8BA4183C41E7C7" bold="true" box="[437,557,870,897]" italics="true" pageId="15" pageNumber="466">
<collectingRegion id="A451C8D9FF91FFA0BF8BA4183C41E7C7" box="[437,557,870,897]" country="United States of America" name="California" pageId="15" pageNumber="466">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF91FFA0BC7FA4193CC2E7C7" box="[577,686,871,897]" pageId="15" pageNumber="466" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<collectingCounty id="8F4B7EB7FF91FFA0BD3BA4193A69E7C7" box="[773,1029,871,897]" pageId="15" pageNumber="466">Santa Barbara County</collectingCounty>
, grounds of
<location id="634A50E0FF91FFA0BAA5A4193F39E7E3" LSID="urn:lsid:plazi:treatment:EE3CB72DFF91FFBEBEA9A5843C18E304:634A50E0FF91FFA0BAA5A4193F39E7E3" country="United States of America" county="Santa Barbara County" name="Santa Barbara Natural History Museum" pageId="15" pageNumber="466" stateProvince="California">Santa Barbara Natural History Museum</location>
,
<date id="122B20FBFF91FFA0BF60A4F53FA8E7E3" box="[350,452,907,933]" pageId="15" pageNumber="466" value="1967-02-07">
<collectingDate id="026FD913FF91FFA0BF60A4F53FA8E7E3" box="[350,452,907,933]" pageId="15" pageNumber="466" value="1967-02-07">7.ii.1967</collectingDate>
</date>
, V.D.
<collectorName id="CB6063EDFF91FFA0BC3BA4F53C57E7E3" box="[517,571,907,933]" pageId="15" pageNumber="466">Roth</collectorName>
(
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)
</materialsCitation>
.
<materialsCitation id="D6FD0C66FF91FFA0BCA8A4F53BF4E7E3" box="[662,1432,907,933]" collectionCode="AMNH" country="United States of America" location="Ventura." pageId="15" pageNumber="466" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<emphasis id="54E1DA29FF91FFA0BCA8A4F53D79E7E3" bold="true" box="[662,789,907,933]" pageId="15" pageNumber="466">
<typeStatus id="B92EB899FF91FFA0BCA8A4F53D7CE7E3" box="[662,784,907,933]" pageId="15" pageNumber="466" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<location id="634A50E0FF91FFA0BD24A4F53D14E7E3" LSID="urn:lsid:plazi:treatment:EE3CB72DFF91FFBEBEA9A5843C18E304:634A50E0FF91FFA0BD24A4F53D14E7E3" box="[794,888,907,933]" country="United States of America" name="Ventura." pageId="15" pageNumber="466" stateProvince="California">Ventura.</location>
<specimenCount id="7093CDB2FF91FFA0BD43A4F53DCCE7E3" box="[893,928,907,933]" pageId="15" pageNumber="466" type="female">1♀</specimenCount>
, “N34 W119” no date, no collector, (
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)
</materialsCitation>
;
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<location id="634A50E0FF91FFA0BEA9A4D13F50E78F" LSID="urn:lsid:plazi:treatment:EE3CB72DFF91FFBEBEA9A5843C18E304:634A50E0FF91FFA0BEA9A4D13F50E78F" box="[151,316,943,969]" country="United States of America" name="Santa Barbara." pageId="15" pageNumber="466" stateProvince="California">Santa Barbara.</location>
<specimenCount id="7093CDB2FF91FFA0BF7AA4D13F0BE78F" box="[324,359,943,969]" pageId="15" pageNumber="466" type="female">1♀</specimenCount>
, N end of
<collectorName id="CB6063EDFF91FFA0BFE1A4D13C16E78F" box="[479,634,943,969]" pageId="15" pageNumber="466">Lk. Cachuma</collectorName>
,
<date id="122B20FBFF91FFA0BCBBA4D13C87E78F" box="[645,747,943,969]" pageId="15" pageNumber="466" value="1967-02-08">
<collectingDate id="026FD913FF91FFA0BCBBA4D13C87E78F" box="[645,747,943,969]" pageId="15" pageNumber="466" value="1967-02-08">8.ii.1967</collectingDate>
</date>
,
<collectorName id="CB6063EDFF91FFA0BCC8A4D13D0BE78F" box="[758,871,943,969]" pageId="15" pageNumber="466">V.D. Roth</collectorName>
(
<collectionCode id="00849EFEFF91FFA0BD49A4D13DDFE78F" box="[887,947,943,969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="466" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF91FFA0BDF9A4D13CFBE7AB" collectingDate="1967-02-07" collectionCode="CAS" collectorName="V. D. Roth" country="United States of America" location="Santa Barbara Natural History Museum" pageId="15" pageNumber="466" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF91FFA0BDF9A4D13D86E78F" box="[967,1002,943,969]" pageId="15" pageNumber="466" type="male">1♂</specimenCount>
<specimenCount id="7093CDB2FF91FFA0BDCFA4D13A79E78F" box="[1009,1045,943,969]" pageId="15" pageNumber="466" type="female">7♀</specimenCount>
, grounds of
<location id="634A50E0FF91FFA0BA9DA4D13F3BE7AB" LSID="urn:lsid:plazi:treatment:EE3CB72DFF91FFBEBEA9A5843C18E304:634A50E0FF91FFA0BA9DA4D13F3BE7AB" country="United States of America" name="Santa Barbara Natural History Museum" pageId="15" pageNumber="466" stateProvince="California">Santa Barbara Natural History Museum</location>
,
<date id="122B20FBFF91FFA0BF5CA4AD3FA4E7AB" box="[354,456,979,1005]" pageId="15" pageNumber="466" value="1967-02-07">
<collectingDate id="026FD913FF91FFA0BF5CA4AD3FA4E7AB" box="[354,456,979,1005]" pageId="15" pageNumber="466" value="1967-02-07">7.ii.1967</collectingDate>
</date>
,
<collectorName id="CB6063EDFF91FFA0BFEDA4AD3C2FE7AB" box="[467,579,979,1005]" pageId="15" pageNumber="466">V.D. Roth</collectorName>
(
<collectionCode id="00849EFEFF91FFA0BC6DA4AD3CE3E7AB" box="[595,655,979,1005]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="466" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF91FFA0BEF9A4893F72E073" pageId="15" pageNumber="466" type="etymology">
<paragraph id="662A063BFF91FFA0BEF9A4893F72E073" blockId="15.[151,1437,870,1977]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A4893F22E057" bold="true" box="[199,334,1015,1041]" pageId="15" pageNumber="466">Etymology.</emphasis>
The specific epithet is a patronym honouring the prominent Los Angeles area blues harmonicist Rod Piazza.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF91FFA0BEF9A3403D6EE2FB" pageId="15" pageNumber="466" type="diagnosis">
<paragraph id="662A063BFF91FFA0BEF9A3403A4DE1A3" blockId="15.[151,1437,870,1977]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A3403F2EE01E" bold="true" box="[199,322,1086,1112]" pageId="15" pageNumber="466">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF91FFA0BFEDA3413C2EE01E" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[467,578,1087,1113]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BFEDA3413C2EE01E" box="[467,578,1087,1113]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BC77A3413CD6E01F" bold="true" box="[585,698,1087,1113]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BC77A3413CD6E01F" box="[585,698,1087,1113]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is distinguished by the form of the distal and proximal arms of the tegular apophysis; the female by the form of the copulatory ducts (
<figureCitation id="FEAE1ABEFF91FFA0BD42A31D3DBCE03B" box="[892,976,1122,1149]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="15" pageNumber="466">Fig. 45</figureCitation>
). The males of
<taxonomicName id="A1957DB8FF91FFA0BA45A31D3A9BE03B" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" box="[1147,1271,1123,1149]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF91FFA0BA45A31D3A9BE03B" box="[1147,1271,1123,1149]" italics="true" pageId="15" pageNumber="466">C. tardatus</emphasis>
</taxonomicName>
and
<taxonomicName id="A1957DB8FF91FFA0BB13A31D3BF0E03A" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1325,1436,1123,1149]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BB13A31D3BF0E03A" box="[1325,1436,1123,1149]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BEA9A3F93F66E0E7" bold="true" box="[151,266,1159,1185]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BEA9A3F93F66E0E7" box="[151,266,1159,1185]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="FEAE1ABEFF91FFA0BF24A3F93F1FE0E7" box="[282,371,1159,1185]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="15" pageNumber="466">Figs 40</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BF41A3F83FF0E0E7" box="[383,412,1158,1185]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="15" pageNumber="466">59</figureCitation>
) are distinguished by the concave distal prolateral margin of the elongate distal arm of the tegular apophysis (
<figureCitation id="FEAE1ABEFF91FFA0BF53A3D53FD3E083" box="[365,447,1195,1221]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="15" pageNumber="466">Fig. 40</figureCitation>
) and the acuminate ventral and dorsal tips of the proximal arm of the tegular apophysis (
<figureCitation id="FEAE1ABEFF91FFA0BEA1A3B13E9AE0AF" box="[159,246,1231,1257]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="15" pageNumber="466">Figs 40</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BF3EA3B03F71E0AF" box="[256,285,1230,1257]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="15" pageNumber="466">59</figureCitation>
) in
<taxonomicName id="A1957DB8FF91FFA0BF76A3B13FDBE0AE" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[328,439,1231,1257]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BF76A3B13FDBE0AE" box="[328,439,1231,1257]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BF83A3B13C43E0AF" bold="true" box="[445,559,1231,1257]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BF83A3B13C43E0AF" box="[445,559,1231,1257]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="54E1DA29FF91FFA0BC00A3B13CEAE0AE" box="[574,646,1231,1256]" italics="true" pageId="15" pageNumber="466">versus</emphasis>
a short, broad, and angular distal arm [
<figureCitation id="FEAE1ABEFF91FFA0BA0BA3B13AE4E0AF" box="[1077,1160,1230,1257]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="15" pageNumber="466">Fig. 48</figureCitation>
] and the rounded ventral tip and acuminate dorsal tip of the proximal arm [
<figureCitation id="FEAE1ABEFF91FFA0BCF8A38D3D73E14B" box="[710,799,1267,1293]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="15" pageNumber="466">Figs 48</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BD15A38C3D26E14B" box="[811,842,1266,1293]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="15" pageNumber="466">60</figureCitation>
] in
<taxonomicName id="A1957DB8FF91FFA0BD4BA38D3D9FE14B" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" box="[885,1011,1267,1293]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF91FFA0BD4BA38D3D9FE14B" box="[885,1011,1267,1293]" italics="true" pageId="15" pageNumber="466">C. tardatus</emphasis>
</taxonomicName>
). The males of
<taxonomicName id="A1957DB8FF91FFA0BA9BA38D3B4EE14A" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1189,1314,1267,1293]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BA9BA38D3B4EE14A" box="[1189,1314,1267,1293]" italics="true" pageId="15" pageNumber="466">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BB17A38D3BF0E14B" bold="true" box="[1321,1436,1267,1293]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BB17A38D3BF0E14B" box="[1321,1436,1267,1293]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF91FFA0BEF7A2693F57E176" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[201,315,1303,1329]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BEF7A2693F57E176" box="[201,315,1303,1329]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BF7BA2693FD6E177" bold="true" box="[325,442,1303,1329]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BF7BA2693FD6E177" box="[325,442,1303,1329]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by the ventral and dorsal tips of the proximal arm of the tegular apophysis being approximately at right angles to each other with the dorsal tip about twice as long as the ventral tip (
<figureCitation id="FEAE1ABEFF91FFA0BEFDA2213F77E13F" box="[195,283,1375,1401]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="15" pageNumber="466">Figs 40</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BF1BA2203F2EE13F" box="[293,322,1374,1401]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="15" pageNumber="466">59</figureCitation>
) in
<taxonomicName id="A1957DB8FF91FFA0BF50A2213FB1E13E" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[366,477,1375,1401]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BF50A2213FB1E13E" box="[366,477,1375,1401]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BFDAA2213C3AE13F" bold="true" box="[484,598,1375,1401]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BFDAA2213C3AE13F" box="[484,598,1375,1401]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="54E1DA29FF91FFA0BC58A2213CC2E13E" box="[614,686,1375,1400]" italics="true" pageId="15" pageNumber="466">versus</emphasis>
the ventral and dorsal tips are oriented in the same direction and of similar lengths (
<figureCitation id="FEAE1ABEFF91FFA0BF70A2FD3FC9E1DB" box="[334,421,1410,1437]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1891,1915]" captionTargetBox="[151,1417,191,1867]" captionTargetId="figure-31@20.[151,1436,181,1867]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="15" pageNumber="466">Figs 57</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BF8CA2FD3FA3E1D8" box="[434,463,1411,1438]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="15" pageNumber="466">61</figureCitation>
) in
<taxonomicName id="A1957DB8FF91FFA0BFC3A2FD3C17E1DA" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[509,635,1411,1437]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BFC3A2FD3C17E1DA" box="[509,635,1411,1437]" italics="true" pageId="15" pageNumber="466">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BCBDA2FD3C9AE1DB" bold="true" box="[643,758,1411,1437]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BCBDA2FD3C9AE1DB" box="[643,758,1411,1437]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
). Furthermore, the dorsal tip of the proximal arm is sharply acuminate in
<taxonomicName id="A1957DB8FF91FFA0BF10A2D93FF1E186" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[302,413,1447,1473]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BF10A2D93FF1E186" box="[302,413,1447,1473]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BF9AA2D93C7BE187" bold="true" box="[420,535,1447,1473]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BF9AA2D93C7BE187" box="[420,535,1447,1473]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="54E1DA29FF91FFA0BC19A2D93C03E186" box="[551,623,1447,1472]" italics="true" pageId="15" pageNumber="466">versus</emphasis>
bluntly acuminate in the male of
<taxonomicName id="A1957DB8FF91FFA0BDD9A2D93A0FE186" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[999,1123,1447,1473]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BDD9A2D93A0FE186" box="[999,1123,1447,1473]" italics="true" pageId="15" pageNumber="466">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BA54A2D93AB0E187" bold="true" box="[1130,1244,1447,1473]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BA54A2D93AB0E187" box="[1130,1244,1447,1473]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
). This difference can be difficult to resolve, however, because of the small size of these structures.
</paragraph>
<paragraph id="662A063BFF91FFA0BEF9A2913D6EE2FB" blockId="15.[151,1437,870,1977]" pageId="15" pageNumber="466">
Distinguishing the females of
<taxonomicName id="A1957DB8FF91FFA0BC25A2913CE7E24E" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[539,651,1519,1545]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BC25A2913CE7E24E" box="[539,651,1519,1545]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BCADA2913D6AE24F" bold="true" box="[659,774,1519,1545]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BCADA2913D6AE24F" box="[659,774,1519,1545]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF91FFA0BD00A2913DDDE24E" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[830,945,1519,1545]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="orarius" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BD00A2913DDDE24E" box="[830,945,1519,1545]" italics="true" pageId="15" pageNumber="466">C. orarius</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BD87A2913A40E24F" bold="true" box="[953,1068,1519,1545]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BD87A2913A40E24F" box="[953,1068,1519,1545]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is discussed under the diagnosis of the latter species. From the other females in the
<taxonomicName id="A1957DB8FF91FFA0BCD4A16D3D2AE26B" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" box="[746,838,1555,1581]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF91FFA0BCD4A16D3D2AE26B" box="[746,838,1555,1581]" italics="true" pageId="15" pageNumber="466">tardatus</emphasis>
</taxonomicName>
species group with rounded spermathecal bases (
<taxonomicName id="A1957DB8FF91FFA0BBBDA16D3E9FE217" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF91FFA0BBBDA16D3E9FE217" italics="true" pageId="15" pageNumber="466">C. tardatus</emphasis>
</taxonomicName>
and
<taxonomicName id="A1957DB8FF91FFA0BF19A1493FCFE216" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[295,419,1591,1617]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BF19A1493FCFE216" box="[295,419,1591,1617]" italics="true" pageId="15" pageNumber="466">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BF96A1493C76E217" bold="true" box="[424,538,1591,1617]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BF96A1493C76E217" box="[424,538,1591,1617]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
) the female of
<taxonomicName id="A1957DB8FF91FFA0BCFEA1493D42E216" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[704,814,1591,1617]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BCFEA1493D42E216" box="[704,814,1591,1617]" italics="true" pageId="15" pageNumber="466">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BD0AA1493DC9E217" bold="true" box="[820,933,1591,1617]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BD0AA1493DC9E217" box="[820,933,1591,1617]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is distinguished by the posteriorly broadened copulatory ducts which obscure the Bennett’s glands in cleared dorsal view of the vulva (
<figureCitation id="FEAE1ABEFF91FFA0BA51A1253AD3E233" box="[1135,1215,1626,1653]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="15" pageNumber="466">Fig. 45</figureCitation>
) (
<emphasis id="54E1DA29FF91FFA0BAE9A1253B73E232" box="[1239,1311,1627,1652]" italics="true" pageId="15" pageNumber="466">versus</emphasis>
copulatory ducts of similar narrow diameter throughout their length and not obscuring the Bennett’s glands in dorsal view in
<taxonomicName id="A1957DB8FF91FFA0BBBDA1013E9FE2FB" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF91FFA0BBBDA1013E9FE2FB" italics="true" pageId="15" pageNumber="466">C. tardatus</emphasis>
</taxonomicName>
[
<figureCitation id="FEAE1ABEFF91FFA0BF3CA1DD3F39E2FB" box="[258,341,1698,1725]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="15" pageNumber="466">Fig. 51</figureCitation>
] and
<taxonomicName id="A1957DB8FF91FFA0BFADA1DD3C7CE2FA" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[403,528,1699,1725]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF91FFA0BFADA1DD3C7CE2FA" box="[403,528,1699,1725]" italics="true" pageId="15" pageNumber="466">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF91FFA0BC28A1DD3CE5E2FB" bold="true" box="[534,649,1699,1725]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="4FD26752FF91FFA0BC28A1DD3CE5E2FB" box="[534,649,1699,1725]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
[
<figureCitation id="FEAE1ABEFF91FFA0BCA9A1DD3C81E2FB" box="[663,749,1698,1725]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="15" pageNumber="466">Fig. 54</figureCitation>
]).
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF91FFBEBEF9A1B83F37E2BD" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466" type="description">
<paragraph id="662A063BFF91FFA0BEF9A1B83A07E340" blockId="15.[151,1437,870,1977]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A1B83F35E2A6" bold="true" box="[199,345,1734,1760]" pageId="15" pageNumber="466">Description.</emphasis>
Ventral tibia I macrosetae variable: 2-1p-2-1p-1p(or 2 or 0). Femora and tibiae lightly banded or not (larger individuals are more likely to have banded legs than are smaller specimens).
</paragraph>
<paragraph id="662A063BFF91FFA0BEF9A0703C99E3D3" blockId="15.[151,1437,870,1977]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A0703F67E36F" bold="true" box="[199,267,1806,1833]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A0703F6EE36F" bold="true" box="[199,258,1806,1833]" italics="true" pageId="15" pageNumber="466">Male</emphasis>
:
</emphasis>
(n=2). About 30–35 peg setae on anteriorly directed patellar apophysis (
<figureCitation id="FEAE1ABEFF91FFA0BA70A0713AC8E36F" box="[1102,1188,1807,1833]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="15" pageNumber="466">Fig. 39</figureCitation>
). Carinate retrolateral tibial apophysis nearly as long as tibia. Distal arm of tegular apophysis (
<figureCitation id="FEAE1ABEFF91FFA0BD8BA04D3A69E30B" box="[949,1029,1843,1869]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="15" pageNumber="466">Fig. 40</figureCitation>
) very broad, elongate; proximal arm (
<figureCitation id="FEAE1ABEFF91FFA0BEA1A0293E95E337" box="[159,249,1879,1905]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="15" pageNumber="466">Figs 40</figureCitation>
,
<figureCitation id="FEAE1ABEFF91FFA0BF38A0283F4FE337" box="[262,291,1878,1905]" captionStart="FIGURES 59–61" captionStartId="21.[151,264,1830,1854]" captionTargetBox="[151,1436,1317,1807]" captionTargetId="figure-450@21.[151,1436,1317,1807]" captionTargetPageId="21" captionText="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi="http://doi.org/10.5281/zenodo.5300665" httpUri="https://zenodo.org/record/5300665/files/figure.png" pageId="15" pageNumber="466">59</figureCitation>
) appearing trifid in ventral view with ventral tip triangular, bluntly acuminate, prolaterally directed; dorsal tip triangular, sharply pointed, basally directed.
</paragraph>
<paragraph id="662A063BFF91FFA0BEF9A0E13B15E3FF" blockId="15.[151,1437,870,1977]" box="[199,1401,1951,1977]" pageId="15" pageNumber="466">
Measurements (n=2). CL 2.7, 2.9; CW 1.98, 2.10; SL 1.30, 1.33; SW 1.23, 1.33.
<typeStatus id="B92EB899FF91FFA0BA6FA0E13AD5E3FF" box="[1105,1209,1951,1977]" pageId="15" pageNumber="466" type="holotype">Holotype</typeStatus>
larger specimen.
</paragraph>
<caption id="32EA56B3FF8EFFBFBEA9A01D3F40E3AE" ID-DOI="http://doi.org/10.5281/zenodo.5300655" ID-Zenodo-Dep="5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="16" pageNumber="467" startId="16.[151,264,1891,1915]" targetBox="[231,1409,181,1839]" targetPageId="16">
<paragraph id="662A063BFF8EFFBFBEA9A01D3F40E3AE" blockId="16.[151,1437,1891,2024]" pageId="16" pageNumber="467">
<emphasis id="54E1DA29FF8EFFBFBEA9A01D3F3AE33D" bold="true" box="[151,342,1891,1915]" pageId="16" pageNumber="467">FIGURES 39–41.</emphasis>
<taxonomicName id="A1957DB8FF8EFFBFBF63A01D3C68E33D" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[349,516,1891,1915]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF8EFFBFBF63A01D3C68E33D" box="[349,516,1891,1915]" italics="true" pageId="16" pageNumber="467">Cybaeus piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8EFFBFBC32A01A3C18E33A" bold="true" box="[524,628,1892,1916]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="4FD26752FF8EFFBFBC32A01A3C18E33A" box="[524,628,1892,1916]" pageId="16" pageNumber="467" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA.
</paragraph>
</caption>
<caption id="32EA56B3FF8FFFBEBEA9A2043F51E19C" ID-DOI="http://doi.org/10.5281/zenodo.5300657" ID-Zenodo-Dep="5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="17" pageNumber="468" startId="17.[151,264,1402,1426]" targetBox="[151,1435,182,1378]" targetPageId="17">
<paragraph id="662A063BFF8FFFBEBEA9A2043F51E19C" blockId="17.[151,1436,1402,1498]" pageId="17" pageNumber="468">
<emphasis id="54E1DA29FF8FFFBEBEA9A2043F39E1D4" bold="true" box="[151,341,1402,1426]" pageId="17" pageNumber="468">FIGURES 42–45.</emphasis>
<taxonomicName id="A1957DB8FF8FFFBEBF65A2043C6EE1D4" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[347,514,1402,1426]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF8FFFBEBF65A2043C6EE1D4" box="[347,514,1402,1426]" italics="true" pageId="17" pageNumber="468">Cybaeus piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF8FFFBEBC36A2043C1CE1D4" bold="true" box="[520,624,1402,1426]" pageId="17" pageNumber="468">
<taxonomicNameLabel id="4FD26752FF8FFFBEBC36A2043C1CE1D4" box="[520,624,1402,1426]" pageId="17" pageNumber="468" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct.
</paragraph>
</caption>
<paragraph id="662A063BFF8FFFBEBEF9A1763A7AE2F4" blockId="17.[151,1437,1544,1859]" pageId="17" pageNumber="468">
<emphasis id="54E1DA29FF8FFFBEBEF9A1763F72E265" bold="true" box="[199,286,1544,1571]" italics="true" pageId="17" pageNumber="468">Female</emphasis>
: (n=9). Atrium (
<figureCitation id="FEAE1ABEFF8FFFBEBFDCA1763C05E265" box="[482,617,1544,1571]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="17" pageNumber="468">Figs 42, 44</figureCitation>
) slightly hooded anteriorly with area between atrial openings inverted vase-shaped, widest posteriorly, concave. Copulatory ducts (
<figureCitation id="FEAE1ABEFF8FFFBEBD7EA1523DADE201" box="[832,961,1580,1607]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="17" pageNumber="468">Figs 42–45</figureCitation>
) opening into atrium laterally from vulva, contiguous or separated anteriorly, gradually broadening posteriorly, turning anteriorly and narrowing at posterior margin of vulva. Spermathecal heads small dorsal lobes medially on vulva, visible in dorsal view. Stalks narrow, relatively long. Bennett’s glands located dorsolaterally on small, rounded bases.
</paragraph>
<paragraph id="662A063BFF8FFFBEBEF9A1C33F37E2BD" blockId="17.[151,1437,1544,1859]" pageId="17" pageNumber="468">Measurements (n=9). CL 2.13–3.4 (2.6+0.4), CW 1.40–2.30 (1.80+0.30), SL 1.04–1.55 (1.26+0.17), SW 0.99– 1.45 (1.20+0.16).</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF8FFFBEBEF9A07A3C18E304" pageId="17" pageNumber="468" type="distribution">
<paragraph id="662A063BFF8FFFBEBEF9A07A3C18E304" blockId="17.[151,1437,1544,1859]" pageId="17" pageNumber="468">
<emphasis id="54E1DA29FF8FFFBEBEF9A07A3C26E358" bold="true" box="[199,586,1796,1823]" pageId="17" pageNumber="468">Distribution and natural history.</emphasis>
(
<figureCitation id="FEAE1ABEFF8FFFBEBC69A07A3CC6E359" box="[599,682,1796,1823]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="17" pageNumber="468">Fig. 62</figureCitation>
). Known only from Ventura and Santa Barbara Counties,
<collectingRegion id="A451C8D9FF8FFFBEBB1AA07A3BF5E358" box="[1316,1433,1796,1822]" country="United States of America" name="California" pageId="17" pageNumber="468">California</collectingRegion>
. The
<specimenCount id="7093CDB2FF8FFFBEBEF4A0573F50E304" box="[202,316,1832,1859]" pageId="17" pageNumber="468" type="male">two males</specimenCount>
were collected in February.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="28A071B5EB5A87C2F69C47296F03B63F">Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade</mods:title>
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<mods:name id="D5C76E53CDBB95F7A7ACCAD7A068E607" type="personal">
<mods:role id="305CD0778A3F2590008F3143F61B76B5">
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<mods:namePart id="6FF29764A4F384ED05D1885DA2F96025">Bennett, Robb</mods:namePart>
<mods:affiliation id="2482FA1D8FBD17DF075C98820024F5D2">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2.</mods:affiliation>
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<mods:name id="4165261074C8DE574679E16734741325" type="personal">
<mods:role id="52D73E6271E75D606712698228A77B92">
<mods:roleTerm id="DBB2101F70C9951931217E48EF36B28F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C56CC97FBFCE1969450598EED6E6753A">Copley, Claudia</mods:namePart>
<mods:nameIdentifier id="35142DA78873029B48BCC2BC22904B2B" type="ORCID">0000-0002-8184-5819</mods:nameIdentifier>
<mods:affiliation id="2DEFC3AA67EA476FB529E1D11A5A071C">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2. &amp; ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
<mods:nameIdentifier id="3170D5C0A7ADF27365DFD18BB7D2B458" type="email">ccopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="7B96F402B5354CBA310EABD823B55A2B" type="personal">
<mods:role id="9202142F8126883C187B3330994632BC">
<mods:roleTerm id="9B4456295F8AAC7B17BADF0608846F0A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B34EB42DBFC88312F09A6D9FDB065F8">Copley, Darren</mods:namePart>
<mods:nameIdentifier id="C0589C8DB5D7332A0D724A4AE9DA18A2" type="ORCID">0000-0002-1944-4272</mods:nameIdentifier>
<mods:affiliation id="A5AAF2BFE1BF3B8CA8337ED2E5724A61">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2. &amp; dcopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 1944 - 4272</mods:affiliation>
<mods:nameIdentifier id="CB3A1054FF2AF22720761DFE6B1C073E" type="email">dcopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0F8AE50929BC69A693E72954563657B9">text</mods:typeOfResource>
<mods:relatedItem id="A2967FE66F8CA49CD7B23B41A5D60D23" type="host">
<mods:titleInfo id="42EDEB587EAF1EE1A8E6C6B2223305EF">
<mods:title id="944A2D224DB056A99064593C07AF05B7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="468DBA229BB33A5093BFEDF2480840E4">
<mods:date id="130EEC33267C67CCCFB7414CC7620825">2021</mods:date>
<mods:detail id="8DA1CEA25D98FB27444500C438239432" type="pubDate">
<mods:number id="65C9E40A2B5D41966D4DA307DDDDCE84">2021-08-27</mods:number>
</mods:detail>
<mods:detail id="FBEF5E84258206EFBE76AFDE8CC417F7" type="volume">
<mods:number id="1DA0A17E18795BA0BBC24131B0637A88">5026</mods:number>
</mods:detail>
<mods:detail id="5317CAB469E2A17633D790C1019067A5" type="issue">
<mods:number id="67B36E0B3FAB65BD1423788BD8D120B3">4</mods:number>
</mods:detail>
<mods:extent id="BE06A653B2BF0F936A42E5DBB7B92BD5" unit="page">
<mods:start id="8F981B389559B17E6953618DB8E9C309">451</mods:start>
<mods:end id="6FA9A2D17E575D9CB528FA0E13BA0021">479</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="4778B3A6DC2133F0C86D0DFFC5BDA449">journal article</mods:classification>
<mods:identifier id="89D0F223CB0F02B0064CB0A3C1A4BDFB" type="DOI">10.11646/zootaxa.5026.4.1</mods:identifier>
<mods:identifier id="8AF9C2B74C50B89DEB13A378B8261CAD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C142F02B43BE19B6DAED2A97AD077591" type="Zenodo-Dep">5300630</mods:identifier>
<mods:identifier id="31C123E754351EDB40B353C30F52592C" type="ZooBank">511C28F6-07E4-412C-B83E-91D2CAC5AA27</mods:identifier>
</mods:mods>
<treatment id="EE3CB72DFF93FFA0BEA9A10D3A9CE6EC" ID-DOI="http://doi.org/10.5281/zenodo.5307732" ID-Zenodo-Dep="5307732" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC" httpUri="http://treatment.plazi.org/id/EE3CB72DFF93FFA0BEA9A10D3A9CE6EC" lastPageId="15" lastPageNumber="466" pageId="13" pageNumber="464">
<subSubSection id="2E8F55B0FF93FFA2BEA9A10D3C29E2CB" box="[151,581,1651,1678]" pageId="13" pageNumber="464" type="nomenclature">
<paragraph id="662A063BFF93FFA2BEA9A10D3C29E2CB" blockId="13.[151,581,1651,1713]" box="[151,581,1651,1678]" pageId="13" pageNumber="464">
<heading id="3D62B157FF93FFA2BEA9A10D3C29E2CB" bold="true" box="[151,581,1651,1678]" fontSize="11" level="1" pageId="13" pageNumber="464" reason="1">
<emphasis id="54E1DA29FF93FFA2BEA9A10D3C29E2CB" bold="true" box="[151,581,1651,1678]" pageId="13" pageNumber="464">
<taxonomicName id="A1957DB8FF93FFA2BEA9A10D3FABE2CB" ID-CoL="99BYV" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,455,1651,1678]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="orarius" status="sp. nov.">
<emphasis id="54E1DA29FF93FFA2BEA9A10D3F31E2CB" bold="true" box="[151,349,1651,1678]" italics="true" pageId="13" pageNumber="464">Cybaeus orarius</emphasis>
Bennett
</taxonomicName>
<taxonomicNameLabel id="4FD26752FF93FFA2BFF0A10D3C29E2CB" box="[462,581,1651,1677]" pageId="13" pageNumber="464" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF93FFA2BEA9A1E93F52E2F7" box="[151,318,1687,1713]" pageId="13" pageNumber="464" type="description">
<paragraph id="662A063BFF93FFA2BEA9A1E93F52E2F7" blockId="13.[151,581,1651,1713]" box="[151,318,1687,1713]" pageId="13" pageNumber="464">
<figureCitation id="FEAE1ABEFF93FFA2BEA9A1E93F7BE2F7" box="[151,279,1687,1713]" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="13" pageNumber="464">Figs 33–38</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BF1CA1E93F52E2F7" box="[290,318,1687,1713]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="13" pageNumber="464">62</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF93FFA2BEA9A1A13DEBE397" pageId="13" pageNumber="464" type="materials_examined">
<paragraph id="662A063BFF93FFA2BEA9A1A13DEBE397" blockId="13.[151,1436,1759,2038]" pageId="13" pageNumber="464">
<emphasis id="54E1DA29FF93FFA2BEA9A1A13F29E2BF" bold="true" box="[151,325,1759,1785]" pageId="13" pageNumber="464">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF93FFA2BF72A1A13C64E35B" collectingDate="1960-04-02" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; W. Ivie &amp; Schrammel" country="United States of America" county="San Luis Obispo County" location="Cambria" pageId="13" pageNumber="464" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF93FFA2BF72A1A13C90E2BF" bold="true" box="[332,764,1759,1786]" pageId="13" pageNumber="464">
<collectingCountry id="1E8246ABFF93FFA2BF72A1A13FF1E2BF" box="[332,413,1759,1785]" name="United States of America" pageId="13" pageNumber="464">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF93FFA2BF97A1A13C4DE2BC" bold="true" box="[425,545,1759,1786]" italics="true" pageId="13" pageNumber="464">
<collectingRegion id="A451C8D9FF93FFA2BF97A1A13C4DE2BC" box="[425,545,1759,1786]" country="United States of America" name="California" pageId="13" pageNumber="464">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF93FFA2BC0FA1A13CF2E2BF" box="[561,670,1759,1785]" pageId="13" pageNumber="464" type="holotype">Holotype</typeStatus>
female.
</emphasis>
<collectingCounty id="8F4B7EB7FF93FFA2BD3CA1A13A74E2BF" box="[770,1048,1759,1785]" pageId="13" pageNumber="464">San Luis Obispo County</collectingCounty>
,
<location id="634A50E0FF93FFA2BA1DA1A13AEAE2BF" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BA1DA1A13AEAE2BF" box="[1059,1158,1759,1785]" country="United States of America" county="San Luis Obispo County" name="Cambria" pageId="13" pageNumber="464" stateProvince="California">Cambria</location>
,
<date id="122B20FBFF93FFA2BAAFA1A13A97E2BF" box="[1169,1275,1759,1785]" pageId="13" pageNumber="464" value="1960-04-02">
<collectingDate id="026FD913FF93FFA2BAAFA1A13A97E2BF" box="[1169,1275,1759,1785]" pageId="13" pageNumber="464" value="1960-04-02">2.iv.1960</collectingDate>
</date>
,
<collectorName id="CB6063EDFF93FFA2BB38A19E3BF4E2BF" box="[1286,1432,1759,1786]" pageId="13" pageNumber="464">W.J. Gertsch</collectorName>
,
<collectorName id="CB6063EDFF93FFA2BEA9A07A3E80E35B" box="[151,236,1795,1822]" pageId="13" pageNumber="464">W. Ivie</collectorName>
, &amp;
<collectorName id="CB6063EDFF93FFA2BF2BA07D3FF9E35B" box="[277,405,1795,1821]" pageId="13" pageNumber="464">Schrammel</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BF9AA07D3C6CE35B" box="[420,512,1795,1821]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="464">AMNH</collectionCode>
)
</materialsCitation>
.
<emphasis id="54E1DA29FF93FFA2BC2DA07D3CFEE35B" bold="true" box="[531,658,1795,1821]" pageId="13" pageNumber="464">
<typeStatus id="B92EB899FF93FFA2BC2DA07D3CE1E35B" box="[531,653,1795,1821]" pageId="13" pageNumber="464" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Monterey.
<materialsCitation id="D6FD0C66FF93FFA2BD2AA07D3FE6E307" collectingDate="1960-04-02" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; W. Ivie &amp; Schrammel" country="United States of America" location="4 mi. S of Gorda" pageId="13" pageNumber="464" specimenCount="2" specimenCount-female="2" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF93FFA2BD2AA07D3D54E35B" box="[788,824,1795,1821]" pageId="13" pageNumber="464" type="female">2♀</specimenCount>
,
<location id="634A50E0FF93FFA2BD7BA07D3A6AE35B" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BD7BA07D3A6AE35B" box="[837,1030,1795,1821]" country="United States of America" name="4 mi. S of Gorda" pageId="13" pageNumber="464" stateProvince="California">
<quantity id="A16DABDEFF93FFA2BD7BA07D3DECE35B" box="[837,896,1795,1821]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="13" pageNumber="464" unit="mi" value="4.0">4 mi.</quantity>
S of Gorda
</location>
,
<date id="122B20FBFF93FFA2BA2CA07D3A10E35B" box="[1042,1148,1795,1821]" pageId="13" pageNumber="464" value="1960-04-02">
<collectingDate id="026FD913FF93FFA2BA2CA07D3A10E35B" box="[1042,1148,1795,1821]" pageId="13" pageNumber="464" value="1960-04-02">2.iv.1960</collectingDate>
</date>
,
<collectorName id="CB6063EDFF93FFA2BAB6A07A3B77E35B" box="[1160,1307,1795,1822]" pageId="13" pageNumber="464">W.J. Gertsch</collectorName>
,
<collectorName id="CB6063EDFF93FFA2BB18A07A3B16E35B" box="[1318,1402,1795,1822]" pageId="13" pageNumber="464">W. Ivie</collectorName>
, &amp;
<collectorName id="CB6063EDFF93FFA2BEA9A0593F7BE307" box="[151,279,1831,1857]" pageId="13" pageNumber="464">Schrammel</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BF18A0593FEEE307" box="[294,386,1831,1857]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="464">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF93FFA2BFA8A0593D85E307" box="[406,1001,1831,1858]" collectingDate="1991-05-30" collectingDateMax="1991-05-31" collectingDateMin="1991-05-30" collectionCode="CAS" collectorName="D. Ubick" country="United States of America" location="Limekiln Creek" pageId="13" pageNumber="464" specimenCount="3" specimenCount-female="3" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF93FFA2BFA8A0593FD6E307" box="[406,442,1831,1857]" pageId="13" pageNumber="464" type="female">3♀</specimenCount>
,
<location id="634A50E0FF93FFA2BFF6A0593C10E307" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BFF6A0593C10E307" box="[456,636,1831,1857]" country="United States of America" name="Limekiln Creek" pageId="13" pageNumber="464" stateProvince="California">Limekiln Creek</location>
,
<date id="122B20FBFF93FFA2BCB6A0593D4EE307" box="[648,802,1831,1857]" pageId="13" pageNumber="464" value="1991-05-30" valueMax="1991-05-31" valueMin="1991-05-30">
<collectingDate id="026FD913FF93FFA2BCB6A0593D4EE307" box="[648,802,1831,1857]" pageId="13" pageNumber="464" value="1991-05-30" valueMax="1991-05-31" valueMin="1991-05-30">30–31.v.1991</collectingDate>
</date>
,
<collectorName id="CB6063EDFF93FFA2BD10A0563DF9E307" box="[814,917,1831,1858]" pageId="13" pageNumber="464">D. Ubick</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BD98A0593D8EE307" box="[934,994,1831,1857]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="464" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF93FFA2BDC8A0593D7AE323" collectingDate="1967-02-10" collectionCode="CAS" collectorName="V. D. Roth" country="United States of America" county="Monterey Co." location="Big Sur Hwy." pageId="13" pageNumber="464" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF93FFA2BDC8A0593A75E307" box="[1014,1049,1831,1857]" pageId="13" pageNumber="464" type="female">1♀</specimenCount>
,
<quantity id="A16DABDEFF93FFA2BA16A0593A0EE307" box="[1064,1122,1831,1857]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="13" pageNumber="464" unit="mi" value="4.0">4 mi.</quantity>
N of the
<collectingCounty id="8F4B7EB7FF93FFA2BAF0A0563B07E307" box="[1230,1387,1831,1858]" pageId="13" pageNumber="464">Monterey Co.</collectingCounty>
line on
<location id="634A50E0FF93FFA2BE85A0353F3AE320" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BE85A0353F3AE320" box="[187,342,1867,1894]" country="United States of America" county="Monterey Co." name="Big Sur Hwy." pageId="13" pageNumber="464" stateProvince="California">Big Sur Hwy.</location>
[Hwy. 1],
<date id="122B20FBFF93FFA2BFEFA0353C29E323" box="[465,581,1867,1893]" pageId="13" pageNumber="464" value="1967-02-10">
<collectingDate id="026FD913FF93FFA2BFEFA0353C29E323" box="[465,581,1867,1893]" pageId="13" pageNumber="464" value="1967-02-10">10.ii.1967</collectingDate>
</date>
,
<collectorName id="CB6063EDFF93FFA2BC6FA0323CADE323" box="[593,705,1867,1894]" pageId="13" pageNumber="464">V.D. Roth</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BCECA0353D62E323" box="[722,782,1867,1893]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="464" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF93FFA2BD1DA0353DB3E323" box="[803,991,1867,1893]" country="United States of America" location="San Luis Obispo" pageId="13" pageNumber="464" specimenCount="1" stateProvince="California" typeStatus="paratype">
<location id="634A50E0FF93FFA2BD1DA0353DB3E323" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BD1DA0353DB3E323" box="[803,991,1867,1893]" country="United States of America" name="San Luis Obispo" pageId="13" pageNumber="464" stateProvince="California">San Luis Obispo</location>
</materialsCitation>
.
<materialsCitation id="D6FD0C66FF93FFA2BDD0A0353C6EE3CF" collectingDate="1960-04-02" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; W. Ivie &amp; Schrammel" country="United States of America" location="Cambria" pageId="13" pageNumber="464" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF93FFA2BDD0A0353A7DE323" box="[1006,1041,1867,1893]" pageId="13" pageNumber="464" type="female">1♀</specimenCount>
,
<location id="634A50E0FF93FFA2BA1EA0353AEFE323" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BA1EA0353AEFE323" box="[1056,1155,1867,1893]" country="United States of America" name="Cambria" pageId="13" pageNumber="464" stateProvince="California">Cambria</location>
,
<date id="122B20FBFF93FFA2BAB1A0353A96E323" box="[1167,1274,1867,1893]" pageId="13" pageNumber="464" value="1960-04-02">
<collectingDate id="026FD913FF93FFA2BAB1A0353A96E323" box="[1167,1274,1867,1893]" pageId="13" pageNumber="464" value="1960-04-02">2.iv.1960</collectingDate>
</date>
,
<collectorName id="CB6063EDFF93FFA2BB38A0323BF4E323" box="[1286,1432,1867,1894]" pageId="13" pageNumber="464">W.J. Gertsch</collectorName>
,
<collectorName id="CB6063EDFF93FFA2BEA9A00E3E86E3CF" box="[151,234,1903,1930]" pageId="13" pageNumber="464">W. Ivie</collectorName>
, &amp;
<collectorName id="CB6063EDFF93FFA2BF2FA0113FFDE3CF" box="[273,401,1903,1929]" pageId="13" pageNumber="464">Schrammel</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BFA0A0113F96E3CF" box="[414,506,1903,1929]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="464">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF93FFA2BC32A0113B3FE3CF" box="[524,1363,1903,1930]" collectingDate="1987-02-13" collectionCode="CAS" collectorName="T. Briggs &amp; V. Lee" country="United States of America" location="Hwy" pageId="13" pageNumber="464" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF93FFA2BC32A0113C43E3CF" box="[524,559,1903,1929]" pageId="13" pageNumber="464" type="female">1♀</specimenCount>
,
<locationDeviation id="77981F19FF93FFA2BC02A0113C99E3CC" box="[572,757,1903,1930]" location="Hwy" pageId="13" pageNumber="464">
<quantity id="A16DABDEFF93FFA2BC02A0113C10E3CF" box="[572,636,1903,1929]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="13" pageNumber="464" unit="mi" value="12.0">12 mi</quantity>
W of Hwy
</locationDeviation>
101 on
<location id="634A50E0FF93FFA2BD70A00E3DE8E3CC" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BD70A00E3DE8E3CC" box="[846,900,1904,1930]" country="United States of America" name="Hwy" pageId="13" pageNumber="464" stateProvince="California">Hwy</location>
46,
<date id="122B20FBFF93FFA2BD8DA0113A4BE3CF" box="[947,1063,1903,1929]" pageId="13" pageNumber="464" value="1987-02-13">
<collectingDate id="026FD913FF93FFA2BD8DA0113A4BE3CF" box="[947,1063,1903,1929]" pageId="13" pageNumber="464" value="1987-02-13">13.ii.1987</collectingDate>
</date>
,
<collectorName id="CB6063EDFF93FFA2BA0FA00E3AF4E3CF" box="[1073,1176,1903,1930]" pageId="13" pageNumber="464">T. Briggs</collectorName>
&amp;
<collectorName id="CB6063EDFF93FFA2BA87A00E3B6DE3CC" box="[1209,1281,1904,1930]" pageId="13" pageNumber="464">V. Lee</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BB31A0113B27E3CF" box="[1295,1355,1903,1929]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="464" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF93FFA2BB60A0113AFCE3EB" collectingDate="1987-02-14" collectionCode="CAS" collectorName="T. Briggs &amp; Lee" country="United States of America" location="1 mi W of Los Osos Valley Rd on Prefumo Canyon Rd" pageId="13" pageNumber="464" specimenCount="2" specimenCount-female="2" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF93FFA2BB60A0113BEFE3CF" box="[1374,1411,1903,1929]" pageId="13" pageNumber="464" type="female">2♀</specimenCount>
,
<location id="634A50E0FF93FFA2BBB0A0113C88E3EB" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BBB0A0113C88E3EB" country="United States of America" name="1 mi W of Los Osos Valley Rd on Prefumo Canyon Rd" pageId="13" pageNumber="464" stateProvince="California">
<quantity id="A16DABDEFF93FFA2BBB0A0113ED9E3EB" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="13" pageNumber="464" unit="mi" value="1.0">1 mi</quantity>
W of Los Osos Valley Rd on Prefumo Canyon Rd
</location>
,
<date id="122B20FBFF93FFA2BCD1A0ED3D0FE3EB" box="[751,867,1939,1965]" pageId="13" pageNumber="464" value="1987-02-14">
<collectingDate id="026FD913FF93FFA2BCD1A0ED3D0FE3EB" box="[751,867,1939,1965]" pageId="13" pageNumber="464" value="1987-02-14">14.ii.1987</collectingDate>
</date>
,
<collectorName id="CB6063EDFF93FFA2BD53A0EA3DB8E3EB" box="[877,980,1939,1966]" pageId="13" pageNumber="464">T. Briggs</collectorName>
&amp; V.
<collectorName id="CB6063EDFF93FFA2BA2AA0EA3A52E3E8" box="[1044,1086,1940,1966]" pageId="13" pageNumber="464">Lee</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BA72A0ED3AE4E3EB" box="[1100,1160,1939,1965]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="464" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF93FFA2BAA5A0ED3DEFE397" collectingDate="1959-08-15" collectionCode="AMNH" collectorName="V. D. Roth &amp; W. J. Gertsch" country="United States of America" location="San Luis Obispo" municipality="Reservoir Canyon" pageId="13" pageNumber="464" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF93FFA2BAA5A0ED3AD2E3EB" box="[1179,1214,1939,1965]" pageId="13" pageNumber="464" type="female">1♀</specimenCount>
,
<collectingMunicipality id="864E9C41FF93FFA2BAF5A0ED3BF4E3EB" box="[1227,1432,1939,1965]" pageId="13" pageNumber="464">Reservoir Canyon</collectingMunicipality>
,
<location id="634A50E0FF93FFA2BEA9A0C93F39E397" LSID="urn:lsid:plazi:treatment:EE3CB72DFF93FFA0BEA9A10D3A9CE6EC:634A50E0FF93FFA2BEA9A0C93F39E397" box="[151,341,1975,2001]" country="United States of America" municipality="Reservoir Canyon" name="San Luis Obispo" pageId="13" pageNumber="464" stateProvince="California">San Luis Obispo</location>
,
<date id="122B20FBFF93FFA2BF5EA0C93F8BE394" box="[352,487,1975,2002]" pageId="13" pageNumber="464" value="1959-08-15">
<collectingDate id="026FD913FF93FFA2BF5EA0C93F8BE394" box="[352,487,1975,2002]" pageId="13" pageNumber="464" value="1959-08-15">15.viii.1959</collectingDate>
</date>
<collectorName id="CB6063EDFF93FFA2BFD0A0C63C31E397" box="[494,605,1975,2002]" pageId="13" pageNumber="464">V.D. Roth</collectorName>
&amp;
<collectorName id="CB6063EDFF93FFA2BCBFA0C63D7CE397" box="[641,784,1975,2002]" pageId="13" pageNumber="464">W.J. Gertsch</collectorName>
(
<collectionCode id="00849EFEFF93FFA2BD21A0C93D17E397" box="[799,891,1975,2001]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="464">AMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF93FFA2BEF9A0A53A62E3B3" box="[199,1038,2011,2038]" pageId="13" pageNumber="464" type="etymology">
<paragraph id="662A063BFF93FFA2BEF9A0A53A62E3B3" blockId="13.[151,1436,1759,2038]" box="[199,1038,2011,2038]" pageId="13" pageNumber="464">
<emphasis id="54E1DA29FF93FFA2BEF9A0A53F24E3B3" bold="true" box="[199,328,2011,2037]" pageId="13" pageNumber="464">Etymology</emphasis>
. The specific name is a Latin adjective meaning “of the coast”.
</paragraph>
</subSubSection>
<caption id="32EA56B3FF90FFA1BEA9A1883DCEE310" ID-DOI="http://doi.org/10.5281/zenodo.5300653" ID-Zenodo-Dep="5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="14" pageNumber="465" startId="14.[151,264,1782,1806]" targetBox="[151,1436,181,1758]" targetPageId="14">
<paragraph id="662A063BFF90FFA1BEA9A1883DCEE310" blockId="14.[151,1437,1782,1878]" pageId="14" pageNumber="465">
<emphasis id="54E1DA29FF90FFA1BEA9A1883F34E348" bold="true" box="[151,344,1782,1806]" pageId="14" pageNumber="465">FIGURES 33–38.</emphasis>
<taxonomicName id="A1957DB8FF90FFA1BF5FA1883C61E348" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[353,525,1782,1806]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="orarius" status="sp. nov.">
<emphasis id="54E1DA29FF90FFA1BF5FA1883C61E348" box="[353,525,1782,1806]" italics="true" pageId="14" pageNumber="465">Cybaeus orarius</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF90FFA1BC28A1883CECE348" bold="true" box="[534,640,1782,1806]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="4FD26752FF90FFA1BC28A1883CECE348" box="[534,640,1782,1806]" pageId="14" pageNumber="465" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca.
</paragraph>
</caption>
<subSubSection id="2E8F55B0FF90FFA1BEF9A0FA3AABE3A0" pageId="14" pageNumber="465" type="diagnosis">
<paragraph id="662A063BFF90FFA1BEF9A0FA3AABE3A0" blockId="14.[151,1436,1924,2023]" pageId="14" pageNumber="465">
<emphasis id="54E1DA29FF90FFA1BEF9A0FA3F2EE3D8" bold="true" box="[199,322,1924,1950]" pageId="14" pageNumber="465">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF90FFA1BFEEA0FB3C2DE3D8" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[464,577,1925,1951]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="orarius">
<emphasis id="54E1DA29FF90FFA1BFEEA0FB3C2DE3D8" box="[464,577,1925,1951]" italics="true" pageId="14" pageNumber="465">C. orarius</emphasis>
</taxonomicName>
is unknown. The female of
<taxonomicName id="A1957DB8FF90FFA1BD4DA0FB3D88E3D8" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[883,996,1925,1951]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="orarius">
<emphasis id="54E1DA29FF90FFA1BD4DA0FB3D88E3D8" box="[883,996,1925,1951]" italics="true" pageId="14" pageNumber="465">C. orarius</emphasis>
</taxonomicName>
is distinguished from the other
<taxonomicName id="A1957DB8FF90FFA1BB7EA0FA3BF0E3D8" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" box="[1344,1436,1924,1950]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF90FFA1BB7EA0FA3BF0E3D8" box="[1344,1436,1924,1950]" italics="true" pageId="14" pageNumber="465">tardatus</emphasis>
</taxonomicName>
species group females by its elongated spermathecal bases (
<figureCitation id="FEAE1ABEFF90FFA1BD0FA0D63A69E384" box="[817,1029,1960,1987]" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="14" pageNumber="465">Figs 34–35, 37–38</figureCitation>
) (
<emphasis id="54E1DA29FF90FFA1BA22A0D73A08E384" box="[1052,1124,1961,1986]" italics="true" pageId="14" pageNumber="465">versus</emphasis>
bases rounded in
<taxonomicName id="A1957DB8FF90FFA1BB13A0D73BF0E384" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1325,1436,1961,1987]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF90FFA1BB13A0D73BF0E384" box="[1325,1436,1961,1987]" italics="true" pageId="14" pageNumber="465">C. piazzai</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF90FFA1BEA9A0B23F66E3A0" bold="true" box="[151,266,1996,2022]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="4FD26752FF90FFA1BEA9A0B23F66E3A0" box="[151,266,1996,2022]" pageId="14" pageNumber="465" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
[
<figureCitation id="FEAE1ABEFF90FFA1BF29A0B23FA9E3A0" box="[279,453,1995,2022]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="14" pageNumber="465">Figs 42–43, 45</figureCitation>
],
<taxonomicName id="A1957DB8FF90FFA1BFE6A0B33C39E3A0" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" box="[472,597,1996,2023]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF90FFA1BFE6A0B33C39E3A0" box="[472,597,1996,2023]" italics="true" pageId="14" pageNumber="465">C. tardatus</emphasis>
</taxonomicName>
[
<figureCitation id="FEAE1ABEFF90FFA1BC5DA0B23C84E3A0" box="[611,744,1995,2022]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="14" pageNumber="465">Figs 50–51</figureCitation>
], and
<taxonomicName id="A1957DB8FF90FFA1BD14A0B33DCAE3A0" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[810,934,1997,2023]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF90FFA1BD14A0B33DCAE3A0" box="[810,934,1997,2023]" italics="true" pageId="14" pageNumber="465">C. topanga</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF90FFA1BD93A0B23A4CE3A0" bold="true" box="[941,1056,1996,2022]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="4FD26752FF90FFA1BD93A0B23A4CE3A0" box="[941,1056,1996,2022]" pageId="14" pageNumber="465" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
[
<figureCitation id="FEAE1ABEFF90FFA1BA13A0B23AD8E3A0" box="[1069,1204,1995,2022]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="14" pageNumber="465">Figs 53–54</figureCitation>
]).
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF91FFA0BEF9A7E83D17E67B" pageId="15" pageNumber="466" type="description">
<paragraph id="662A063BFF91FFA0BEF9A7E83C7BE493" blockId="15.[151,1436,150,682]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A7E83F35E4F6" bold="true" box="[199,345,150,176]" pageId="15" pageNumber="466">Description.</emphasis>
Ventral tibia I macrosetae 2-1p-2-1p-1p(or 0). Legs with femora and tibiae banded (
<collectingRegion id="A451C8D9FF91FFA0BB0BA7E93BF0E4F7" box="[1333,1436,151,177]" country="Argentina" name="San Luis" pageId="15" pageNumber="466">San Luis</collectingRegion>
Obispo County specimens) or not.
</paragraph>
<paragraph id="662A063BFF91FFA0BEF9A7A03F3EE5B3" blockId="15.[151,1436,150,682]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A7A03F4BE4BF" bold="true" box="[199,295,222,249]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A7A03F72E4BF" bold="true" box="[199,286,222,249]" italics="true" pageId="15" pageNumber="466">Female</emphasis>
:
</emphasis>
(n=12). Atrium (
<figureCitation id="FEAE1ABEFF91FFA0BFDBA7A13C08E4BC" box="[485,612,223,250]" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="15" pageNumber="466">Figs 33, 36</figureCitation>
) difficult to discern, a concave area on epigynum with paired longitudinal atrial openings diverging posteriorly and area between openings slightly raised, convex. Vulva (
<figureCitation id="FEAE1ABEFF91FFA0BADFA67D3ED8E507" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="15" pageNumber="466">Figs 34–35, 37– 38</figureCitation>
) with copulatory ducts joined to atrial openings dorsally or dorso-laterally, contiguous anteriorly, diverging posteriorly then converging and turning anteriorly at posterior margin of vulva; heads of spermathecae (
<figureCitation id="FEAE1ABEFF91FFA0BB28A6353BF8E523" box="[1302,1428,330,357]" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="15" pageNumber="466">Figs 35, 38</figureCitation>
) small dorsal lobes; spermathecal stalks proceeding anteriorly from heads for short distance then turning posteriorly before angling laterally to cross and join spermathecal bases; bases longitudinally elongated, about twice as long as wide, rounded anteriorly, gradually tapering into fertilization ducts posteriorly; Bennett’s glands located on dorsal surface of bases.
</paragraph>
<paragraph id="662A063BFF91FFA0BEF9A6813D17E67B" blockId="15.[151,1436,150,682]" pageId="15" pageNumber="466">
Measurements (n=6). CL 2.10–3.1 (2.6+0.4), CW 1.43–2.13 (1.76+0.27), SL 1.05–1.43 (1.24+0.14), SW 1.03– 1.38 (1.18+0.14).
<typeStatus id="B92EB899FF91FFA0BF5CA55D3FA6E67B" box="[354,458,547,573]" pageId="15" pageNumber="466" type="holotype">Holotype</typeStatus>
CL 2.23, CW 1.60, SL 1.17, SW 1.07.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF91FFA0BEF9A5383A9CE6EC" pageId="15" pageNumber="466" type="distribution">
<paragraph id="662A063BFF91FFA0BEF9A5383A9CE6EC" blockId="15.[151,1436,150,682]" pageId="15" pageNumber="466">
<emphasis id="54E1DA29FF91FFA0BEF9A5383C35E626" bold="true" box="[199,601,582,609]" pageId="15" pageNumber="466">Distribution and natural history.</emphasis>
(
<figureCitation id="FEAE1ABEFF91FFA0BC52A5393CA8E624" box="[620,708,583,610]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="15" pageNumber="466">Fig. 62</figureCitation>
). Known only from an approximately
<quantity id="A16DABDEFF91FFA0BAB9A5393ABDE627" box="[1159,1233,583,609]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="466" unit="km" value="70.0">70 km</quantity>
length of coastal
<collectingRegion id="A451C8D9FF91FFA0BEA9A5153F65E6C3" box="[151,265,619,645]" country="United States of America" name="California" pageId="15" pageNumber="466">California</collectingRegion>
from southwestern Monterey County to central
<collectingRegion id="A451C8D9FF91FFA0BD23A5153D13E6C3" box="[797,895,619,645]" country="Argentina" name="San Luis" pageId="15" pageNumber="466">San Luis</collectingRegion>
Obispo County. Specimens have been recovered from oak leaf litter (Prefumo Canyon Road) and under rocks in a redwood forest (Limekiln Creek).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7AFFB86820B70E54A65AE56CD7CFAC7C">Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade</mods:title>
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<subSubSection id="2E8F55B0FF93FFA2BEA9A7E93F8EE4F7" box="[151,482,151,178]" pageId="13" pageNumber="464" type="nomenclature">
<paragraph id="662A063BFF93FFA2BEA9A7E93F8EE4F7" blockId="13.[151,482,151,178]" box="[151,482,151,178]" pageId="13" pageNumber="464">
<heading id="3D62B157FF93FFA2BEA9A7E93F8EE4F7" bold="true" box="[151,482,151,178]" fontSize="11" level="1" pageId="13" pageNumber="464" reason="1">
<emphasis id="54E1DA29FF93FFA2BEA9A7E93F8EE4F7" bold="true" box="[151,482,151,178]" pageId="13" pageNumber="464">
The
<taxonomicName id="A1957DB8FF93FFA2BEF1A7E93F5EE4F4" ID-CoL="32RHW" authorityName="Chamberlin" authorityYear="1924" box="[207,306,151,178]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF93FFA2BEF1A7E93F5EE4F4" bold="true" box="[207,306,151,178]" italics="true" pageId="13" pageNumber="464">tardatus</emphasis>
</taxonomicName>
species group
</emphasis>
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</subSubSection>
<subSubSection id="2E8F55B0FF93FFA2BEA9A7A13A99E558" pageId="13" pageNumber="464" type="diagnosis">
<paragraph id="662A063BFF93FFA2BEA9A7A13A99E558" blockId="13.[151,1437,223,861]" pageId="13" pageNumber="464">
<emphasis id="54E1DA29FF93FFA2BEA9A7A13F66E4BF" bold="true" box="[151,266,223,249]" pageId="13" pageNumber="464">Diagnosis</emphasis>
. The
<taxonomicName id="A1957DB8FF93FFA2BF69A7A13FDFE4BF" authorityName="Chamberlin" authorityYear="1924" box="[343,435,223,249]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF93FFA2BF69A7A13FDFE4BF" box="[343,435,223,249]" italics="true" pageId="13" pageNumber="464">tardatus</emphasis>
</taxonomicName>
group is diagnosed by the somewhat triangular form of the vulva and the simple, longitudinally elongated, centrally located copulatory ducts (
<figureCitation id="FEAE1ABEFF93FFA2BD7EA67D3A79E55B" box="[832,1045,259,286]" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="13" pageNumber="464">Figs 34–35, 37–38</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BA1EA67D3AFEE558" box="[1056,1170,259,286]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="13" pageNumber="464">42–43, 45</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BAA0A67D3A85E558" box="[1182,1257,259,286]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="13" pageNumber="464">50–51</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF93FFA2BEF9A6593F23E71B" pageId="13" pageNumber="464" type="description">
<paragraph id="662A063BFF93FFA2BEF9A6593C08E5CF" blockId="13.[151,1437,223,861]" pageId="13" pageNumber="464">
<emphasis id="54E1DA29FF93FFA2BEF9A6593F3DE507" bold="true" box="[199,337,295,321]" pageId="13" pageNumber="464">Description</emphasis>
. Medium- to large-sized spiders: carapace lengths averaging
<quantity id="A16DABDEFF93FFA2BDD2A6593A1CE504" box="[1004,1136,295,322]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="13" pageNumber="464" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
(females). Males subequal. Abdomen patterned. Femora and tibiae banded or not. Two or three complete pairs of ventral tibia I macrosetae; distal pair present, absent, or incomplete.
</paragraph>
<paragraph id="662A063BFF93FFA2BEF9A6ED3CB6E65F" blockId="13.[151,1437,223,861]" pageId="13" pageNumber="464">
<emphasis id="54E1DA29FF93FFA2BEF9A6ED3F6EE5E8" bold="true" box="[199,258,403,430]" italics="true" pageId="13" pageNumber="464">Male</emphasis>
: Patellar apophysis (
<figureCitation id="FEAE1ABEFF93FFA2BFCEA6ED3C25E5EB" box="[496,585,403,429]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="13" pageNumber="464">Figs 39</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BC68A6ED3C18E5EB" box="[598,628,403,429]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="13" pageNumber="464">47</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BCBFA6ED3CF2E5E8" box="[641,670,403,430]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1891,1915]" captionTargetBox="[151,1417,191,1867]" captionTargetId="figure-31@20.[151,1436,181,1867]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300663" httpUri="https://zenodo.org/record/5300663/files/figure.png" pageId="13" pageNumber="464">56</figureCitation>
) about as long as tibial width, anteriorly directed, about 25–40 peg setae. Retrolateral tibial apophysis carinate, nearly as long as tibia or slightly truncated anteriorly. Embolus (
<figureCitation id="FEAE1ABEFF93FFA2BB54A6C93EDAE5B3" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="13" pageNumber="464">Figs 40</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BEFEA6A53EB1E5B3" box="[192,221,475,501]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="13" pageNumber="464">48</figureCitation>
) long, thin. Distal and proximal arms of tegular apophysis species specific (
<figureCitation id="FEAE1ABEFF93FFA2BA1BA6A53AC9E5B3" box="[1061,1189,475,501]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1891,1915]" captionTargetBox="[231,1409,181,1839]" captionTargetId="figure-31@16.[178,1410,181,1868]" captionTargetPageId="16" captionText="FIGURES 39–41. Cybaeus piazzai spec. nov., male from Santa Barbara, California, left palpus. 39 Patella and tibia, dorsal. 40–41 Genital bulb (40 ventral, 41 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA— proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." figureDoi="http://doi.org/10.5281/zenodo.5300655" httpUri="https://zenodo.org/record/5300655/files/figure.png" pageId="13" pageNumber="464">Figs 40–41</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BA91A6A53A94E5B3" box="[1199,1272,475,501]" captionStart="FIGURES 46–49" captionStartId="18.[151,264,1929,1953]" captionTargetBox="[164,1433,185,1891]" captionTargetId="figure-31@18.[151,1436,181,1905]" captionTargetPageId="18" captionText="FIGURES 46–49. Cybaeus tardatus, male from Claremont, California, left palpus. 46–47 Patella and tibia (46 retrolateral, 47 dorsal). 48–49 Genital bulb (48 ventral, 49 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300659" httpUri="https://zenodo.org/record/5300659/files/figure.png" pageId="13" pageNumber="464">48–49</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BB3CA6A53B21E5B0" box="[1282,1357,475,502]" captionStart-0="FIGURES 55–58" captionStart-1="FIGURES 59–61" captionStartId-0="20.[151,264,1891,1915]" captionStartId-1="21.[151,264,1830,1854]" captionTargetBox-0="[151,1417,191,1867]" captionTargetBox-1="[151,1436,1317,1807]" captionTargetId-0="figure-31@20.[151,1436,181,1867]" captionTargetId-1="figure-450@21.[151,1436,1317,1807]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURES 55–58. Cybaeus topanga spec. nov., holotype male, left palpus. 55–56 Patella and tibia (55 retrolateral, 56 dorsal). 57–58 Genital bulb (57 ventral, 58 retrolateral).E—embolus, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates concave distal prolateral margin of DA." captionText-1="FIGURES 59–61. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis, ventral (59 C. piazzai spec. nov. from Santa Barbara, 60 C. tardatus from Claremont, 61 C. topanga spec. nov., holotype). Solid arrows indicate ventral tip of proximal arm, dashed arrows indicate dorsal tip of proximal arm." figureDoi-0="http://doi.org/10.5281/zenodo.5300663" figureDoi-1="http://doi.org/10.5281/zenodo.5300665" httpUri-0="https://zenodo.org/record/5300663/files/figure.png" httpUri-1="https://zenodo.org/record/5300665/files/figure.png" pageId="13" pageNumber="464">57–61</figureCitation>
): distal arm elongated or short, proximal arm with bifid tip.
</paragraph>
<paragraph id="662A063BFF93FFA2BEF9A55D3EB2E68B" blockId="13.[151,1437,223,861]" pageId="13" pageNumber="464">
<emphasis id="54E1DA29FF93FFA2BEF9A55D3F72E678" bold="true" box="[199,286,547,574]" italics="true" pageId="13" pageNumber="464">Female</emphasis>
: Atrium (
<figureCitation id="FEAE1ABEFF93FFA2BFB6A55D3C6AE67B" box="[392,518,547,573]" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="13" pageNumber="464">Figs 33, 36</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BC2EA55D3C43E67B" box="[528,559,547,573]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="13" pageNumber="464">44</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BC07A55D3C3AE678" box="[569,598,547,574]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="13" pageNumber="464">50</figureCitation>
) with paired longitudinal atrial openings, atrium hooded anteriorly (
<figureCitation id="FEAE1ABEFF93FFA2BB7AA55D3BF8E678" box="[1348,1428,547,574]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="13" pageNumber="464">Fig. 52</figureCitation>
) or not. Copulatory ducts (
<figureCitation id="FEAE1ABEFF93FFA2BF8AA5393C5FE627" box="[436,563,583,610]" captionStart="FIGURES 33–38" captionStartId="14.[151,264,1782,1806]" captionTargetBox="[151,1436,181,1758]" captionTargetId="figure-31@14.[151,1436,181,1758]" captionTargetPageId="14" captionText="FIGURES 33–38. Cybaeus orarius spec. nov., females from California, copulatory organ (33–35 from Big Sur Highway, 36–38 from Cambria). 33, 36 Epigynum, ventral. 34–35, 37–38 Vulva (34, 37 ventral, 35, 38 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300653" httpUri="https://zenodo.org/record/5300653/files/figure.png" pageId="13" pageNumber="464">Figs 35, 38</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BC01A5393CEAE624" box="[575,646,583,610]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1402,1426]" captionTargetBox="[151,1435,182,1378]" captionTargetId="figure-21@17.[151,1436,181,1378]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus piazzai spec. nov., females from California, copulatory organ (42–43 from Santa Barbara Natural History Museum grounds, 44–45 from “W119/N34”). 42–43 Vulva, ventral. 44 Epigynum, ventral. 45 Vulva, dorsal). CD— copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.5300657" httpUri="https://zenodo.org/record/5300657/files/figure.png" pageId="13" pageNumber="464">43, 45</figureCitation>
,
<figureCitation id="FEAE1ABEFF93FFA2BCAFA5393CB4E624" box="[657,728,583,610]" captionStart="FIGURES 50–54" captionStartId="19.[151,264,1939,1963]" captionTargetBox="[157,1429,689,1916]" captionTargetId="figure-163@19.[151,1436,684,1916]" captionTargetPageId="19" captionText="FIGURES 50–54. Cybaeus spp., females, copulatory organ (50–51 C. tardatus, holotype of Namopsilus pletus; 52–54 C. topanga spec. nov., from San Ysidro, California). 50–51, 53–54 Vulva (50, 53 ventral, 51, 54 dorsal). 52 Atrium (anterior portion), ventral. BG—Bennett’s gland, CD—copulatory duct, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300661" httpUri="https://zenodo.org/record/5300661/files/figure.png" pageId="13" pageNumber="464">51, 54</figureCitation>
) long, relatively simple, contiguous or separated. Spermathecal heads simple, small lobes or undifferentiated from rest of vulval ducts. Spermathecal stalks simple, of variable length. Spermathecal bases rounded or elongated. Bennett’s glands located on surface of bases near junction with stalks.
</paragraph>
<paragraph id="662A063BFF93FFA2BEF9A5A93F23E71B" blockId="13.[151,1437,223,861]" pageId="13" pageNumber="464">
<emphasis id="54E1DA29FF93FFA2BEF9A5A93C46E6B7" bold="true" box="[199,554,727,753]" pageId="13" pageNumber="464">Composition and distribution</emphasis>
. (
<figureCitation id="FEAE1ABEFF93FFA2BC7EA5A93CF9E6B4" box="[576,661,727,754]" captionStart="FIGURE 62" captionStartId="22.[151,250,1909,1933]" captionTargetBox="[186,1407,198,1875]" captionTargetId="figure-31@22.[169,1419,181,1885]" captionTargetPageId="22" captionText="FIGURE 62. Distribution of Cybaeus orarius spec. nov., C. piazzai spec. nov., C. tardatus, and C. topanga spec. nov. AZ— Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300667" httpUri="https://zenodo.org/record/5300667/files/figure.png" pageId="13" pageNumber="464">Fig. 62</figureCitation>
).
<taxonomicName id="A1957DB8FF93FFA2BC97A5A93DAEE6B4" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[681,962,727,754]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="orarius" status="sp. nov.">
<emphasis id="54E1DA29FF93FFA2BC97A5A93D0FE6B7" box="[681,867,727,753]" italics="true" pageId="13" pageNumber="464">Cybaeus orarius</emphasis>
Bennett
</taxonomicName>
<emphasis id="54E1DA29FF93FFA2BDF4A5A93A51E6B7" bold="true" box="[970,1085,727,753]" pageId="13" pageNumber="464">
<taxonomicNameLabel id="4FD26752FF93FFA2BDF4A5A93A51E6B7" box="[970,1085,727,753]" pageId="13" pageNumber="464" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="A1957DB8FF93FFA2BA75A5A63B77E6B4" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1099,1307,728,754]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="piazzai" status="sp. nov.">
<emphasis id="54E1DA29FF93FFA2BA75A5A63AD0E6B7" box="[1099,1212,728,754]" italics="true" pageId="13" pageNumber="464">C. piazzai</emphasis>
Bennett
</taxonomicName>
<emphasis id="54E1DA29FF93FFA2BB1CA5A93BF9E6B7" bold="true" box="[1314,1429,727,753]" pageId="13" pageNumber="464">
<taxonomicNameLabel id="4FD26752FF93FFA2BB1CA5A93BF9E6B7" box="[1314,1429,727,753]" pageId="13" pageNumber="464" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="A1957DB8FF93FFA2BEA9A5823FD0E750" authority="(Chamberlin)" baseAuthorityName="Chamberlin" baseAuthorityYear="1919" box="[151,444,763,790]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="tardatus">
<emphasis id="54E1DA29FF93FFA2BEA9A5823F7BE753" box="[151,279,763,790]" italics="true" pageId="13" pageNumber="464">C. tardatus</emphasis>
(Chamberlin)
</taxonomicName>
, and
<taxonomicName id="A1957DB8FF93FFA2BFC3A5823CB2E750" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[509,734,764,790]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="topanga" status="sp. nov.">
<emphasis id="54E1DA29FF93FFA2BFC3A5823C11E753" box="[509,637,764,790]" italics="true" pageId="13" pageNumber="464">C. topanga</emphasis>
Bennett
</taxonomicName>
<emphasis id="54E1DA29FF93FFA2BCD6A5853D32E753" bold="true" box="[744,862,763,789]" pageId="13" pageNumber="464">
<taxonomicNameLabel id="4FD26752FF93FFA2BCD6A5853D32E753" box="[744,862,763,789]" pageId="13" pageNumber="464" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
All four species are rarely encountered and have small ranges scattered along the southern coast of western
<collectingRegion id="A451C8D9FF93FFA2BD12A4613DF2E77F" box="[812,926,799,825]" country="United States of America" name="California" pageId="13" pageNumber="464">California</collectingRegion>
from southern Monterey County south to the Mexican border.
</paragraph>
</subSubSection>
</treatment>
</document>

294
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<heading id="3D62B157FF97FFA6BEA9A12E3C3EE22D" bold="true" box="[151,594,1616,1643]" fontSize="11" level="1" pageId="9" pageNumber="460" reason="1">
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<taxonomicName id="A1957DB8FF97FFA6BEA9A12E3FB8E22D" ID-CoL="999XZ" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,468,1616,1643]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF97FFA6BEA9A12E3F06E22D" bold="true" box="[151,362,1616,1643]" italics="true" pageId="9" pageNumber="460">Cybaeus wilsonia</emphasis>
Bennett
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<paragraph id="662A063BFF97FFA6BEA9A10B3F8BE2C9" blockId="9.[151,594,1616,1680]" box="[151,487,1653,1680]" pageId="9" pageNumber="460">
<figureCitation id="FEAE1ABEFF97FFA6BEA9A10B3F7BE2C9" box="[151,279,1653,1679]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="9" pageNumber="460">Figs 17–19</figureCitation>
,
<figureCitation id="FEAE1ABEFF97FFA6BF1CA10B3F00E2D6" box="[290,364,1653,1680]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="9" pageNumber="460">23–25</figureCitation>
,
<figureCitation id="FEAE1ABEFF97FFA6BF48A10B3FF8E2C9" box="[374,404,1653,1679]" captionStart="FIGURES 26–28" captionStartId="11.[151,264,1216,1240]" captionTargetBox="[151,1436,705,1193]" captionTargetId="figure-226@11.[151,1436,705,1193]" captionTargetPageId="11" captionText="FIGURES 26–28. Cybaeus spp. males, left palpus, patellar apophysis, dorsal (26 C. bilectus spec. nov., from Double Bunk Meadows, California, 27 C. bryoncavus spec. nov., holotype, 28 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (26 concave distal margin; 28 concave dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.5300646" httpUri="https://zenodo.org/record/5300646/files/figure.png" pageId="9" pageNumber="460">28</figureCitation>
,
<figureCitation id="FEAE1ABEFF97FFA6BF9EA10B3F8BE2C9" box="[416,487,1653,1679]" captionStart-0="FIGURES 29–31" captionStart-1="FIGURE 32" captionStartId-0="11.[151,264,1858,1882]" captionStartId-1="12.[151,250,1918,1942]" captionTargetBox-0="[151,1436,1337,1834]" captionTargetBox-1="[184,1406,198,1877]" captionTargetId-0="figure-280@11.[151,1436,1337,1834]" captionTargetId-1="figure-31@12.[163,1423,181,1894]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 29–31. Cybaeus spp. males, left palpus, proximal arm of tegular apophysis, ventral (29 C. bilectus spec. nov., from Double Bunk Meadows, California, 30 C. bryoncavus spec. nov., holotype, 31 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (29 bluntly hooked tip, 30 inconspicuous basal projection)." captionText-1="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi-0="http://doi.org/10.5281/zenodo.5300649" figureDoi-1="http://doi.org/10.5281/zenodo.5300651" httpUri-0="https://zenodo.org/record/5300649/files/figure.png" httpUri-1="https://zenodo.org/record/5300651/files/figure.png" pageId="9" pageNumber="460">31–32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF97FFA6BEA9A1C33CDAE359" pageId="9" pageNumber="460" type="materials_examined">
<paragraph id="662A063BFF97FFA6BEA9A1C33CDAE359" blockId="9.[151,1437,1724,2040]" pageId="9" pageNumber="460">
<emphasis id="54E1DA29FF97FFA6BEA9A1C33F27E291" bold="true" box="[151,331,1725,1751]" pageId="9" pageNumber="460">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF97FFA6BF66A1C33CD0E2BD" collectingDate="1959-09-13" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; V. D. Roth" country="United States of America" location="California" pageId="9" pageNumber="460" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF97FFA6BF66A1C33D6FE291" bold="true" box="[344,771,1724,1751]" pageId="9" pageNumber="460">
<collectingCountry id="1E8246ABFF97FFA6BF66A1C33FC4E291" box="[344,424,1725,1751]" name="United States of America" pageId="9" pageNumber="460">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF97FFA6BF85A1C23C5FE291" bold="true" box="[443,563,1724,1751]" italics="true" pageId="9" pageNumber="460">
<collectingRegion id="A451C8D9FF97FFA6BF85A1C23C5FE291" box="[443,563,1724,1751]" country="United States of America" name="California" pageId="9" pageNumber="460">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF97FFA6BC77A1C33CDAE291" box="[585,694,1725,1751]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<taxonomicName id="A1957DB8FF97FFA6BD2EA1C33BF4E291" authority="County, Wilsonia, Kings Canyon National Park" authorityName="County, Wilsonia, Kings Canyon National Park" box="[784,1432,1725,1751]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="tulare">Tulare County, Wilsonia, Kings Canyon National Park</taxonomicName>
,
<date id="122B20FBFF97FFA6BEA9A19F3F7DE2BA" box="[151,273,1761,1788]" pageId="9" pageNumber="460" value="1959-09-13">
<collectingDate id="026FD913FF97FFA6BEA9A19F3F7DE2BA" box="[151,273,1761,1788]" pageId="9" pageNumber="460" value="1959-09-13">13.ix.1959</collectingDate>
</date>
,
<collectorName id="CB6063EDFF97FFA6BF20A19F3FC3E2BD" box="[286,431,1761,1787]" pageId="9" pageNumber="460">W.J. Gertsch</collectorName>
&amp;
<collectorName id="CB6063EDFF97FFA6BFE8A19F3C2BE2BD" box="[470,583,1761,1787]" pageId="9" pageNumber="460">V.D. Roth</collectorName>
(
<collectionCode id="00849EFEFF97FFA6BC66A19F3CD8E2BD" box="[600,692,1761,1787]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="460">AMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="D6FD0C66FF97FFA6BCF6A19F3CDEE359" collectingDate="1959-09-13" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; V. D. Roth" country="United States of America" location="Kings Canyon National Park" municipality="Wilsonia" pageId="9" pageNumber="460" specimenCount="3" specimenCount-female="3" stateProvince="California" typeStatus="paratype">
<emphasis id="54E1DA29FF97FFA6BCF6A19F3D2BE2BD" bold="true" box="[712,839,1761,1787]" pageId="9" pageNumber="460">
<typeStatus id="B92EB899FF97FFA6BCF6A19F3D2EE2BD" box="[712,834,1761,1787]" pageId="9" pageNumber="460" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Tulare.
<specimenCount id="7093CDB2FF97FFA6BD99A19F3DA7E2BD" box="[935,971,1761,1787]" pageId="9" pageNumber="460" type="female">3♀</specimenCount>
,
<collectingMunicipality id="864E9C41FF97FFA6BDE7A19F3A2CE2BD" box="[985,1088,1761,1787]" pageId="9" pageNumber="460">Wilsonia</collectingMunicipality>
,
<location id="634A50E0FF97FFA6BA73A19F3BF4E2BD" LSID="urn:lsid:plazi:treatment:EE3CB72DFF97FFA4BEA9A12E3F79E6EF:634A50E0FF97FFA6BA73A19F3BF4E2BD" box="[1101,1432,1761,1787]" country="United States of America" municipality="Wilsonia" name="Kings Canyon National Park" pageId="9" pageNumber="460" stateProvince="California">Kings Canyon National Park</location>
,
<date id="122B20FBFF97FFA6BEA9A07B3F7DE366" box="[151,273,1797,1824]" pageId="9" pageNumber="460" value="1959-09-13">
<collectingDate id="026FD913FF97FFA6BEA9A07B3F7DE366" box="[151,273,1797,1824]" pageId="9" pageNumber="460" value="1959-09-13">13.ix.1959</collectingDate>
</date>
,
<collectorName id="CB6063EDFF97FFA6BF22A07B3FC7E359" box="[284,427,1797,1823]" pageId="9" pageNumber="460">W.J. Gertsch</collectorName>
&amp;
<collectorName id="CB6063EDFF97FFA6BFF1A07B3C52E359" box="[463,574,1797,1823]" pageId="9" pageNumber="460">V.D. Roth</collectorName>
(
<collectionCode id="00849EFEFF97FFA6BC70A07B3CC6E359" box="[590,682,1797,1823]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="460">AMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF97FFA6BEF9A0573A05E305" box="[199,1129,1833,1859]" pageId="9" pageNumber="460" type="etymology">
<paragraph id="662A063BFF97FFA6BEF9A0573A05E305" blockId="9.[151,1437,1724,2040]" box="[199,1129,1833,1859]" pageId="9" pageNumber="460">
<emphasis id="54E1DA29FF97FFA6BEF9A0573F22E305" bold="true" box="[199,334,1833,1859]" pageId="9" pageNumber="460">Etymology.</emphasis>
The specific name is a noun in apposition taken from the
<typeStatus id="B92EB899FF97FFA6BDE6A0573A64E305" box="[984,1032,1833,1859]" pageId="9" pageNumber="460">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF97FFA4BEF9A0333D41E4BF" lastPageId="11" lastPageNumber="462" pageId="9" pageNumber="460" type="diagnosis">
<paragraph id="662A063BFF97FFA6BEF9A0333B3FE3B1" blockId="9.[151,1437,1724,2040]" pageId="9" pageNumber="460">
<emphasis id="54E1DA29FF97FFA6BEF9A0333F2EE321" bold="true" box="[199,322,1869,1895]" pageId="9" pageNumber="460">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF97FFA6BFE3A0333C32E321" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[477,606,1869,1895]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF97FFA6BFE3A0333C32E321" box="[477,606,1869,1895]" italics="true" pageId="9" pageNumber="460">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF97FFA6BC58A0333CB6E321" bold="true" box="[614,730,1869,1895]" pageId="9" pageNumber="460">
<taxonomicNameLabel id="4FD26752FF97FFA6BC58A0333CB6E321" box="[614,730,1869,1895]" pageId="9" pageNumber="460" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is unlikely to be confused with the males of
<taxonomicName id="A1957DB8FF97FFA6BADEA0333B35E321" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1248,1369,1869,1895]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF97FFA6BADEA0333B35E321" box="[1248,1369,1869,1895]" italics="true" pageId="9" pageNumber="460">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF97FFA6BB5CA0333EA4E3CD" bold="true" pageId="9" pageNumber="460">
<taxonomicNameLabel id="4FD26752FF97FFA6BB5CA0333EA4E3CD" pageId="9" pageNumber="460" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
or
<taxonomicName id="A1957DB8FF97FFA6BED2A00F3FE2E3CD" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[236,398,1905,1931]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF97FFA6BED2A00F3FE2E3CD" box="[236,398,1905,1931]" italics="true" pageId="9" pageNumber="460">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF97FFA6BFAAA00F3C6AE3CD" bold="true" box="[404,518,1905,1931]" pageId="9" pageNumber="460">
<taxonomicNameLabel id="4FD26752FF97FFA6BFAAA00F3C6AE3CD" box="[404,518,1905,1931]" pageId="9" pageNumber="460" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, the other members of the
<taxonomicName id="A1957DB8FF97FFA6BD0EA00F3D14E3CD" box="[816,888,1905,1931]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF97FFA6BD0EA00F3D14E3CD" box="[816,888,1905,1931]" italics="true" pageId="9" pageNumber="460">devius</emphasis>
</taxonomicName>
species group with known males. The male of
<taxonomicName id="A1957DB8FF97FFA6BBBDA00F3E9AE3E9" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF97FFA6BBBDA00F3E9AE3E9" italics="true" pageId="9" pageNumber="460">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF97FFA6BEC3A0EB3F03E3E9" bold="true" box="[253,367,1941,1967]" pageId="9" pageNumber="460">
<taxonomicNameLabel id="4FD26752FF97FFA6BEC3A0EB3F03E3E9" box="[253,367,1941,1967]" pageId="9" pageNumber="460" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is diagnosed by the narrow elongate distal arm of the tegular apophysis (length about 2.5 times width) (
<figureCitation id="FEAE1ABEFF97FFA6BECFA0C73F28E395" box="[241,324,1977,2003]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="9" pageNumber="460">Fig. 24</figureCitation>
) and the laterally projecting patellar apophysis which is about as long as the width of the patella, somewhat concave dorsally, and has about 15 peg setae scattered dorsally about the distal half (
<figureCitation id="FEAE1ABEFF97FFA6BAFDA0A33B76E3B1" box="[1219,1306,2013,2039]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="9" pageNumber="460">Figs 23</figureCitation>
,
<figureCitation id="FEAE1ABEFF97FFA6BB18A0A33B2AE3B1" box="[1318,1350,2013,2039]" captionStart="FIGURES 26–28" captionStartId="11.[151,264,1216,1240]" captionTargetBox="[151,1436,705,1193]" captionTargetId="figure-226@11.[151,1436,705,1193]" captionTargetPageId="11" captionText="FIGURES 26–28. Cybaeus spp. males, left palpus, patellar apophysis, dorsal (26 C. bilectus spec. nov., from Double Bunk Meadows, California, 27 C. bryoncavus spec. nov., holotype, 28 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (26 concave distal margin; 28 concave dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.5300646" httpUri="https://zenodo.org/record/5300646/files/figure.png" pageId="9" pageNumber="460">28</figureCitation>
).
</paragraph>
<caption id="32EA56B3FF94FFA5BEA9A01D3A54E385" ID-DOI="http://doi.org/10.5281/zenodo.5300644" ID-Zenodo-Dep="5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="10" pageNumber="461" startId="10.[151,264,1891,1915]" targetBox="[151,1424,183,1827]" targetPageId="10">
<paragraph id="662A063BFF94FFA5BEA9A01D3A54E385" blockId="10.[151,1437,1891,1987]" pageId="10" pageNumber="461">
<emphasis id="54E1DA29FF94FFA5BEA9A01D3F38E33D" bold="true" box="[151,340,1891,1915]" pageId="10" pageNumber="461">FIGURES 23–25.</emphasis>
<taxonomicName id="A1957DB8FF94FFA5BF64A01D3C61E33D" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[346,525,1891,1915]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF94FFA5BF64A01D3C61E33D" box="[346,525,1891,1915]" italics="true" pageId="10" pageNumber="461">Cybaeus wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF94FFA5BC2DA01D3C15E33D" bold="true" box="[531,633,1891,1915]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="4FD26752FF94FFA5BC2DA01D3C15E33D" box="[531,633,1891,1915]" pageId="10" pageNumber="461" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25).
</paragraph>
</caption>
<paragraph id="662A063BFF95FFA4BEF9A7E93D41E4BF" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="462">
The female of
<taxonomicName id="A1957DB8FF95FFA4BF56A7E93F8AE4F7" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[360,486,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF95FFA4BF56A7E93F8AE4F7" box="[360,486,151,177]" italics="true" pageId="11" pageNumber="462">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF95FFA4BFD2A7E93C31E4F7" bold="true" box="[492,605,151,177]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="4FD26752FF95FFA4BFD2A7E93C31E4F7" box="[492,605,151,177]" pageId="11" pageNumber="462" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is diagnosed by the form and configuration of the vulval components (
<figureCitation id="FEAE1ABEFF95FFA4BB54A7E93EB3E493" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="11" pageNumber="462">Figs 18–19</figureCitation>
) and is only likely to be confused with the females of
<taxonomicName id="A1957DB8FF95FFA4BD67A7C53DA8E493" box="[857,964,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF95FFA4BD67A7C53DA8E493" box="[857,964,187,213]" italics="true" pageId="11" pageNumber="462">C. devius</emphasis>
</taxonomicName>
or
<taxonomicName id="A1957DB8FF95FFA4BDD3A7C53A29E493" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1005,1093,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF95FFA4BDD3A7C53A29E493" box="[1005,1093,187,213]" italics="true" pageId="11" pageNumber="462">C. echo</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF95FFA4BA70A7C53AAFE493" bold="true" box="[1102,1219,187,213]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="4FD26752FF95FFA4BA70A7C53AAFE493" box="[1102,1219,187,213]" pageId="11" pageNumber="462" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
Distinguishing the females of these three species is discussed under
<taxonomicName id="A1957DB8FF95FFA4BC80A7A13D4AE4BF" box="[702,806,223,249]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF95FFA4BC80A7A13D4AE4BF" box="[702,806,223,249]" italics="true" pageId="11" pageNumber="462">C. devius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF95FFA4BEF9A67C3B0DE67B" pageId="11" pageNumber="462" type="description">
<paragraph id="662A063BFF95FFA4BEF9A67C3A9CE55B" blockId="11.[151,1436,151,681]" box="[199,1264,258,285]" pageId="11" pageNumber="462">
<emphasis id="54E1DA29FF95FFA4BEF9A67C3F3DE55A" bold="true" box="[199,337,258,284]" pageId="11" pageNumber="462">Description</emphasis>
. Tibia I ventral macrosetae 2-1p-2-1p-0. Femora very lightly banded or unbanded.
</paragraph>
<paragraph id="662A063BFF95FFA4BEF9A6583CFFE5CF" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="462">
<emphasis id="54E1DA29FF95FFA4BEF9A6583F67E507" bold="true" box="[199,267,294,321]" pageId="11" pageNumber="462">
<emphasis id="54E1DA29FF95FFA4BEF9A6583F6EE507" bold="true" box="[199,258,294,321]" italics="true" pageId="11" pageNumber="462">Male</emphasis>
:
</emphasis>
(n=1). Embolus (
<figureCitation id="FEAE1ABEFF95FFA4BFE7A6593C40E507" box="[473,556,295,321]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="11" pageNumber="462">Fig. 24</figureCitation>
) relatively broad and ribbon-like, about twice the thickness of the emboli in males of
<taxonomicName id="A1957DB8FF95FFA4BF3EA6353F14E523" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[256,376,331,357]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF95FFA4BF3EA6353F14E523" box="[256,376,331,357]" italics="true" pageId="11" pageNumber="462">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF95FFA4BFBEA6353F98E523" bold="true" box="[384,500,331,357]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="4FD26752FF95FFA4BFBEA6353F98E523" box="[384,500,331,357]" pageId="11" pageNumber="462" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF95FFA4BC12A6353CA2E523" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[556,718,331,357]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF95FFA4BC12A6353CA2E523" box="[556,718,331,357]" italics="true" pageId="11" pageNumber="462">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF95FFA4BCE8A6353D26E523" bold="true" box="[726,842,331,357]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="4FD26752FF95FFA4BCE8A6353D26E523" box="[726,842,331,357]" pageId="11" pageNumber="462" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
Proximal arm of tegular apophysis (
<figureCitation id="FEAE1ABEFF95FFA4BAD6A6353B07E523" box="[1256,1387,330,357]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="11" pageNumber="462">Figs 24–25</figureCitation>
,
<figureCitation id="FEAE1ABEFF95FFA4BB49A6353BF8E523" box="[1399,1428,331,357]" captionStart="FIGURES 29–31" captionStartId="11.[151,264,1858,1882]" captionTargetBox="[151,1436,1337,1834]" captionTargetId="figure-280@11.[151,1436,1337,1834]" captionTargetPageId="11" captionText="FIGURES 29–31. Cybaeus spp. males, left palpus, proximal arm of tegular apophysis, ventral (29 C. bilectus spec. nov., from Double Bunk Meadows, California, 30 C. bryoncavus spec. nov., holotype, 31 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (29 bluntly hooked tip, 30 inconspicuous basal projection)." figureDoi="http://doi.org/10.5281/zenodo.5300649" httpUri="https://zenodo.org/record/5300649/files/figure.png" pageId="11" pageNumber="462">30</figureCitation>
) simple, elongate, acuminate, basally swollen.
</paragraph>
<paragraph id="662A063BFF95FFA4BEF9A6ED3D04E5EB" blockId="11.[151,1436,151,681]" box="[199,872,403,429]" pageId="11" pageNumber="462">Measurements (n=1). CL 2.13, CW 1.68, SL 1.12, SW 1.08.</paragraph>
<paragraph id="662A063BFF95FFA4BEF9A6C83FF0E65F" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="462">
<emphasis id="54E1DA29FF95FFA4BEF9A6C83F4BE597" bold="true" box="[199,295,438,465]" pageId="11" pageNumber="462">
<emphasis id="54E1DA29FF95FFA4BEF9A6C83F72E597" bold="true" box="[199,286,438,465]" italics="true" pageId="11" pageNumber="462">Female</emphasis>
:
</emphasis>
(n=3). See note under description of
<taxonomicName id="A1957DB8FF95FFA4BCE8A6C93D2DE597" box="[726,833,439,465]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF95FFA4BCE8A6C93D2DE597" box="[726,833,439,465]" italics="true" pageId="11" pageNumber="462">C. devius</emphasis>
</taxonomicName>
. Atrium (
<figureCitation id="FEAE1ABEFF95FFA4BD8EA6C93A5DE597" box="[944,1073,439,465]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="11" pageNumber="462">Figs 17–18</figureCitation>
) very small, about 1/4 width of vulva. Spermathecal heads exit vulval ducts ventrally posterior to region enclosed by copulatory ducts, dorsolaterally deflected (
<figureCitation id="FEAE1ABEFF95FFA4BF33A6813FFCE65F" box="[269,400,511,537]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="11" pageNumber="462">Figs 18–19</figureCitation>
).
</paragraph>
<paragraph id="662A063BFF95FFA4BEF9A55D3B0DE67B" blockId="11.[151,1436,151,681]" box="[199,1377,546,573]" pageId="11" pageNumber="462">Measurements (n=3). CL 1.95, 2.08, 2.30; CW 1.38, 1.43, 1.63; SL 1.01, 1.07, 1.18; SW 1.00, 1.04, 1.11.</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF95FFA4BEF9A5383F79E6EF" pageId="11" pageNumber="462" type="distribution">
<paragraph id="662A063BFF95FFA4BEF9A5383F79E6EF" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="462">
<emphasis id="54E1DA29FF95FFA4BEF9A5383C20E626" bold="true" box="[199,588,582,609]" pageId="11" pageNumber="462">Distribution and natural history</emphasis>
. (
<figureCitation id="FEAE1ABEFF95FFA4BC5DA5393CD5E627" box="[611,697,583,609]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="11" pageNumber="462">Fig. 32</figureCitation>
). Known only from the
<typeStatus id="B92EB899FF95FFA4BDF4A5393D96E627" box="[970,1018,583,609]" pageId="11" pageNumber="462">type</typeStatus>
locality on the western slopes of the southern Sierra
<collectingRegion id="A451C8D9FF95FFA4BF74A5153FCCE6C3" box="[330,416,619,645]" country="United States of America" name="Nevada" pageId="11" pageNumber="462">Nevada</collectingRegion>
in Kings Canyon National Park in
<taxonomicName id="A1957DB8FF95FFA4BD0EA5153A6FE6C3" authority="County. All" authorityName="County. All" box="[816,1027,619,645]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="tulare">Tulare County. All</taxonomicName>
known specimens were collected in September.
</paragraph>
</subSubSection>
</treatment>
</document>

200
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<mods:title id="E101A0D7791C436119ABEA4724ED02B5">Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade</mods:title>
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<mods:namePart id="EC347495A3CFABC4CD0590AEA71F5E44">Copley, Claudia</mods:namePart>
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<mods:affiliation id="673CC919707035ACFD9B5EB0FEA88E62">Department of Entomology, Natural History Section, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V 8 W 9 W 2. &amp; ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
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<mods:namePart id="467B46CA7BD06F38110FC3201906DCDE">Copley, Darren</mods:namePart>
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<subSubSection id="2E8F55B0FF99FFA8BEA9A10D3C4AE2C8" box="[151,550,1651,1678]" pageId="7" pageNumber="458" type="nomenclature">
<paragraph id="662A063BFF99FFA8BEA9A10D3C4AE2C8" blockId="7.[151,550,1651,1714]" box="[151,550,1651,1678]" pageId="7" pageNumber="458">
<heading id="3D62B157FF99FFA8BEA9A10D3C4AE2C8" bold="true" box="[151,550,1651,1678]" fontSize="11" level="1" pageId="7" pageNumber="458" reason="1">
<emphasis id="54E1DA29FF99FFA8BEA9A10D3C4AE2C8" bold="true" box="[151,550,1651,1678]" pageId="7" pageNumber="458">
<taxonomicName id="A1957DB8FF99FFA8BEA9A10D3FC4E2C8" ID-CoL="99BCF" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,424,1651,1678]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BEA9A10D3F52E2C8" bold="true" box="[151,318,1651,1678]" italics="true" pageId="7" pageNumber="458">Cybaeus echo</emphasis>
Bennett
</taxonomicName>
<taxonomicNameLabel id="4FD26752FF99FFA8BF91A10A3C4AE2C8" box="[431,550,1652,1678]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEA9A1E63F52E2F4" box="[151,318,1688,1714]" pageId="7" pageNumber="458" type="description">
<paragraph id="662A063BFF99FFA8BEA9A1E63F52E2F4" blockId="7.[151,550,1651,1714]" box="[151,318,1688,1714]" pageId="7" pageNumber="458">
<figureCitation id="FEAE1ABEFF99FFA8BEA9A1E63F7BE2F4" box="[151,279,1688,1714]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="7" pageNumber="458">Figs 20–22</figureCitation>
,
<figureCitation id="FEAE1ABEFF99FFA8BF1CA1E63F52E2F4" box="[290,318,1688,1714]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="7" pageNumber="458">32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEA9A19E3F28E358" pageId="7" pageNumber="458" type="materials_examined">
<paragraph id="662A063BFF99FFA8BEA9A19E3F28E358" blockId="7.[151,1437,1759,2003]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEA9A19E3F2BE2BC" bold="true" box="[151,327,1760,1786]" pageId="7" pageNumber="458">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF99FFA8BF6EA19E3F28E358" collectingDate="1958-07-09" collectionCode="AMNH" collectorName="Echo Summit &amp; V. D. Roth" country="United States of America" county="El Dorado County" location="El Dorado County" pageId="7" pageNumber="458" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF99FFA8BF6EA19E3D6AE2BC" bold="true" box="[336,774,1759,1786]" pageId="7" pageNumber="458">
<collectingCountry id="1E8246ABFF99FFA8BF6EA19E3FCDE2BC" box="[336,417,1760,1786]" name="United States of America" pageId="7" pageNumber="458">U.S.A.</collectingCountry>
:
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<collectingRegion id="A451C8D9FF99FFA8BF8EA1A13C44E2BC" box="[432,552,1759,1786]" country="United States of America" name="California" pageId="7" pageNumber="458">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF99FFA8BC04A19E3CCBE2BC" box="[570,679,1760,1786]" pageId="7" pageNumber="458" type="holotype">Holotype</typeStatus>
female.
</emphasis>
<collectingCounty id="8F4B7EB7FF99FFA8BD31A19E3D8EE2BC" box="[783,994,1760,1786]" pageId="7" pageNumber="458">El Dorado County</collectingCounty>
, below
<collectorName id="CB6063EDFF99FFA8BA03A19E3AB7E2BC" box="[1085,1243,1760,1786]" pageId="7" pageNumber="458">Echo Summit</collectorName>
,
<date id="122B20FBFF99FFA8BAD6A19E3B30E2BD" box="[1256,1372,1760,1787]" pageId="7" pageNumber="458" value="1958-07-09">
<collectingDate id="026FD913FF99FFA8BAD6A19E3B30E2BD" box="[1256,1372,1760,1787]" pageId="7" pageNumber="458" value="1958-07-09">9.vii.1958</collectingDate>
</date>
,
<collectorName id="CB6063EDFF99FFA8BB57A19F3EA1E358" pageId="7" pageNumber="458">V.D. Roth</collectorName>
(
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).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEF9A0563A04E304" box="[199,1128,1832,1859]" pageId="7" pageNumber="458" type="etymology">
<paragraph id="662A063BFF99FFA8BEF9A0563A04E304" blockId="7.[151,1437,1759,2003]" box="[199,1128,1832,1859]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A0563F22E304" bold="true" box="[199,334,1832,1858]" pageId="7" pageNumber="458">Etymology.</emphasis>
The specific name is a noun in apposition taken from the
<typeStatus id="B92EB899FF99FFA8BDE9A0573A6BE305" box="[983,1031,1833,1859]" pageId="7" pageNumber="458">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEF9A0323A56E3E9" pageId="7" pageNumber="458" type="diagnosis">
<paragraph id="662A063BFF99FFA8BEF9A0323A56E3E9" blockId="7.[151,1437,1759,2003]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A0323F2EE320" bold="true" box="[199,322,1868,1894]" pageId="7" pageNumber="458">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF99FFA8BFDEA0323C54E320" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[480,568,1868,1894]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo">
<emphasis id="54E1DA29FF99FFA8BFDEA0323C54E320" box="[480,568,1868,1894]" italics="true" pageId="7" pageNumber="458">C. echo</emphasis>
</taxonomicName>
is unknown. The female is diagnosed by the form and configuration of the vulval components (
<figureCitation id="FEAE1ABEFF99FFA8BF47A00E3F94E3CC" box="[377,504,1904,1930]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="7" pageNumber="458">Figs 21–22</figureCitation>
) and is only likely to be confused with the females of
<taxonomicName id="A1957DB8FF99FFA8BA6FA00E3AD5E3CC" box="[1105,1209,1904,1930]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BA6FA00E3AD5E3CC" box="[1105,1209,1904,1930]" italics="true" pageId="7" pageNumber="458">C. devius</emphasis>
</taxonomicName>
or
<taxonomicName id="A1957DB8FF99FFA8BAE3A00E3B30E3CC" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1245,1372,1904,1930]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BAE3A00E3B30E3CC" box="[1245,1372,1904,1930]" italics="true" pageId="7" pageNumber="458">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BB5CA00E3EA4E3E8" bold="true" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BB5CA00E3EA4E3E8" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
Distinguishing the females of these three species is discussed under
<taxonomicName id="A1957DB8FF99FFA8BDF5A0EA3A5FE3E8" box="[971,1075,1940,1966]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BDF5A0EA3A5FE3E8" box="[971,1075,1940,1966]" italics="true" pageId="7" pageNumber="458">C. devius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA6BEF9A0C63D36E55B" lastPageId="9" lastPageNumber="460" pageId="7" pageNumber="458" type="description">
<paragraph id="662A063BFF99FFA8BEF9A0C63A4CE394" blockId="7.[151,1437,1759,2003]" box="[199,1056,1976,2003]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A0C63F35E394" bold="true" box="[199,345,1976,2002]" pageId="7" pageNumber="458">Description.</emphasis>
Ventral tibia I macrosetae 2-1p-2-1p-0. Femora lightly banded.
</paragraph>
<caption id="32EA56B3FF96FFA7BEA9A1A83DA2E31C" ID-DOI="http://doi.org/10.5281/zenodo.5300638" ID-Zenodo-Dep="5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="8" pageNumber="459" startId="8.[151,264,1750,1774]" targetBox="[153,1434,181,1726]" targetPageId="8">
<paragraph id="662A063BFF96FFA7BEA9A1A83DA2E31C" blockId="8.[151,1436,1750,1882]" pageId="8" pageNumber="459">
<emphasis id="54E1DA29FF96FFA7BEA9A1A83F3AE2A8" bold="true" box="[151,342,1750,1774]" pageId="8" pageNumber="459">FIGURES 14–19.</emphasis>
<taxonomicName id="A1957DB8FF96FFA7BF60A1A83F85E2AB" box="[350,489,1750,1774]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="54E1DA29FF96FFA7BF60A1A83FD9E2A8" box="[350,437,1750,1774]" italics="true" pageId="8" pageNumber="459">Cybaeus</emphasis>
spp.
</taxonomicName>
, females, copulatory organ (14–16
<taxonomicName id="A1957DB8FF96FFA7BD66A1A83DD4E2A8" box="[856,952,1750,1774]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF96FFA7BD66A1A83DD4E2A8" box="[856,952,1750,1774]" italics="true" pageId="8" pageNumber="459">C. devius</emphasis>
</taxonomicName>
, holotype; 17–19
<taxonomicName id="A1957DB8FF96FFA7BA48A1A83A87E2A8" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1142,1259,1750,1774]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF96FFA7BA48A1A83A87E2A8" box="[1142,1259,1750,1774]" italics="true" pageId="8" pageNumber="459">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF96FFA7BACDA1A83B30E2A8" bold="true" box="[1267,1372,1750,1774]" pageId="8" pageNumber="459">
<taxonomicNameLabel id="4FD26752FF96FFA7BACDA1A83B30E2A8" box="[1267,1372,1750,1774]" pageId="8" pageNumber="459" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows).
</paragraph>
</caption>
<paragraph id="662A063BFF97FFA6BEF9A7E83FC2E4BF" blockId="9.[151,1437,150,321]" pageId="9" pageNumber="460">
<emphasis id="54E1DA29FF97FFA6BEF9A7E83F4BE4F7" bold="true" box="[199,295,150,177]" pageId="9" pageNumber="460">
<emphasis id="54E1DA29FF97FFA6BEF9A7E83F72E4F7" bold="true" box="[199,286,150,177]" italics="true" pageId="9" pageNumber="460">Female</emphasis>
:
</emphasis>
(n=1). See note under description of
<taxonomicName id="A1957DB8FF97FFA6BCE8A7E93D2DE4F7" box="[726,833,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF97FFA6BCE8A7E93D2DE4F7" box="[726,833,151,177]" italics="true" pageId="9" pageNumber="460">C. devius</emphasis>
</taxonomicName>
. Atrium (
<figureCitation id="FEAE1ABEFF97FFA6BD8EA7E93A5EE4F7" box="[944,1074,151,177]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="9" pageNumber="460">Figs 20–21</figureCitation>
) very small, about 1/4 width of vulva. Spermathecal heads (
<figureCitation id="FEAE1ABEFF97FFA6BFE8A7C53C34E493" box="[470,600,187,213]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="9" pageNumber="460">Figs 21–22</figureCitation>
) exit vulval ducts ventrally basal to region enclosed by copulatory ducts, deflected dorso-laterally.
</paragraph>
<paragraph id="662A063BFF97FFA6BEF9A67D3D36E55B" blockId="9.[151,1437,150,321]" box="[199,858,259,285]" pageId="9" pageNumber="460">Measurements (n=1). CL 2.7, CW 2.00, SL 1.33, SW 1.27.</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF97FFA6BEF9A6583BF1E507" box="[199,1437,294,321]" pageId="9" pageNumber="460" type="distribution">
<paragraph id="662A063BFF97FFA6BEF9A6583BF1E507" blockId="9.[151,1437,150,321]" box="[199,1437,294,321]" pageId="9" pageNumber="460">
<emphasis id="54E1DA29FF97FFA6BEF9A6583F35E506" bold="true" box="[199,345,294,320]" pageId="9" pageNumber="460">Distribution</emphasis>
. (
<figureCitation id="FEAE1ABEFF97FFA6BF51A6593FAFE507" box="[367,451,295,321]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="9" pageNumber="460">Fig. 32</figureCitation>
). Known only from the
<typeStatus id="B92EB899FF97FFA6BCF2A6593C90E507" box="[716,764,295,321]" pageId="9" pageNumber="460">type</typeStatus>
locality in El Dorado County in the northern Sierra
<collectingRegion id="A451C8D9FF97FFA6BB7EA6593BF5E507" box="[1344,1433,295,321]" country="United States of America" name="Nevada" pageId="9" pageNumber="460">Nevada</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="662A063BFF99FFA8BEA9A6343C32E522" blockId="7.[151,606,330,393]" box="[151,606,330,357]" pageId="7" pageNumber="458">
<heading id="3D62B157FF99FFA8BEA9A6343C32E522" bold="true" box="[151,606,330,357]" fontSize="11" level="1" pageId="7" pageNumber="458" reason="1">
<taxonomicName id="A1957DB8FF99FFA8BEA9A6343C32E522" ID-CoL="6BSTR" authority="Chamberlin and Ivie" authorityName="Chamberlin and Ivie" box="[151,606,330,357]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BEA9A6343C32E522" bold="true" box="[151,606,330,357]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEA9A6343F3EE523" bold="true" box="[151,338,330,357]" italics="true" pageId="7" pageNumber="458">Cybaeus devius</emphasis>
Chamberlin and Ivie
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEA9A6113F52E5CF" box="[151,318,367,393]" pageId="7" pageNumber="458" type="description">
<paragraph id="662A063BFF99FFA8BEA9A6113F52E5CF" blockId="7.[151,606,330,393]" box="[151,318,367,393]" pageId="7" pageNumber="458">
<figureCitation id="FEAE1ABEFF99FFA8BEA9A6113F7BE5CF" box="[151,279,367,393]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Figs 14–16</figureCitation>
,
<figureCitation id="FEAE1ABEFF99FFA8BF1CA6113F52E5CF" box="[290,318,367,393]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="7" pageNumber="458">32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEA9A6CD3CF4E5AC" pageId="7" pageNumber="458" type="reference_group">
<paragraph id="662A063BFF99FFA8BEA9A6CD3CF4E5AC" blockId="7.[151,1437,435,490]" pageId="7" pageNumber="458">
<treatmentCitationGroup id="46852115FF99FFA8BEA9A6CD3CF4E5AC" pageId="7" pageNumber="458">
<taxonomicName id="A1957DB8FF99FFA8BEA9A6CD3C1BE58D" ID-CoL="6BSTR" authority="Chamberlin and Ivie 1942: 18" authorityName="Chamberlin and Ivie" authorityPageNumber="18" authorityYear="1942" box="[151,631,435,459]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BEA9A6CD3F5BE58D" box="[151,311,435,459]" italics="true" pageId="7" pageNumber="458">Cybaeus devius</emphasis>
Chamberlin and Ivie 1942: 18
</taxonomicName>
, fig. 48.
<treatmentCitation id="E734202AFF99FFA8BCEBA6CA3DE6E58D" author="Roewer, C. F." box="[725,906,435,459]" page="90" pageId="7" pageNumber="458" year="1954">
<bibRefCitation id="02047BCAFF99FFA8BCEBA6CA3DE6E58D" author="Roewer, C. F." box="[725,906,435,459]" pageId="7" pageNumber="458" refId="ref17427" refString="Roewer, C. F. (1954) Katalog der Araneae 2 a. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 923 pp." type="book" year="1954">Roewer 1954: 90</bibRefCitation>
</treatmentCitation>
. Roth and Brown 1986: 3. Bennett in
<treatmentCitation id="E734202AFF99FFA8BB29A6CD3F43E5AC" author="Copley, C. R. &amp; Bennett, R. &amp; Perlman, S. J." page="373" pageId="7" pageNumber="458" year="2009">
<bibRefCitation id="02047BCAFF99FFA8BB29A6CD3F43E5AC" author="Copley, C. R. &amp; Bennett, R. &amp; Perlman, S. J." pageId="7" pageNumber="458" pagination="367 - 401" refId="ref15419" refString="Copley, C. R., Bennett, R. &amp; Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
Copley
<emphasis id="54E1DA29FF99FFA8BB56A6CA3BF1E58D" box="[1384,1437,435,459]" italics="true" pageId="7" pageNumber="458">et al.</emphasis>
2009: 373
</bibRefCitation>
</treatmentCitation>
, fig. 9.
<bibRefCitation id="02047BCAFF99FFA8BF42A6AC3CF8E5AC" author="World Spider Catalog" box="[380,660,466,490]" pageId="7" pageNumber="458" refId="ref18189" refString="World Spider Catalog (2021) World Spider Catalog. Version 22.0. Natural History Museum, Bern. Available from: https: // wsc. nmbe. ch / (accessed 15 June 2021) https: // doi. org / 10.24436 / 2" type="book" year="2021">World Spider Catalog 2021</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEA9A5663F9FE610" pageId="7" pageNumber="458" type="materials_examined">
<paragraph id="662A063BFF99FFA8BEA9A5663F9FE610" blockId="7.[151,1437,535,1570]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEA9A5663FD3E674" bold="true" box="[151,447,536,562]" pageId="7" pageNumber="458">Type material examined.</emphasis>
<materialsCitation id="D6FD0C66FF99FFA8BFF8A5663F9FE610" collectionCode="AMNH" country="United States of America" county="Madera County" latitude="37.25" location="North Fork" longLatPrecision="1182" longitude="-119.5" pageId="7" pageNumber="458" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF99FFA8BFF8A5663D1AE674" bold="true" box="[454,886,535,562]" pageId="7" pageNumber="458">
<collectingCountry id="1E8246ABFF99FFA8BFF8A5663C7BE674" box="[454,535,536,562]" name="United States of America" pageId="7" pageNumber="458">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF99FFA8BC1AA5693CF0E674" bold="true" box="[548,668,535,562]" italics="true" pageId="7" pageNumber="458">
<collectingRegion id="A451C8D9FF99FFA8BC1AA5693CF0E674" box="[548,668,535,562]" country="United States of America" name="California" pageId="7" pageNumber="458">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF99FFA8BC92A5663D75E674" box="[684,793,536,562]" pageId="7" pageNumber="458" type="holotype">Holotype</typeStatus>
female.
</emphasis>
<collectingCounty id="8F4B7EB7FF99FFA8BD43A5663A42E674" box="[893,1070,536,562]" pageId="7" pageNumber="458">Madera County</collectingCounty>
,
<location id="634A50E0FF99FFA8BA07A5663AD5E674" LSID="urn:lsid:plazi:treatment:EE3CB72DFF99FFA8BEA9A6343FFDE264:634A50E0FF99FFA8BA07A5663AD5E674" box="[1081,1209,536,562]" country="United States of America" county="Madera County" latitude="37.25" longLatPrecision="1182" longitude="-119.5" name="North Fork" pageId="7" pageNumber="458" stateProvince="California">North Fork</location>
,
<geoCoordinate id="03A160FCFF99FFA8BAFAA5663B71E675" box="[1220,1309,536,563]" degrees="37" direction="north" minutes="15" orientation="latitude" pageId="7" pageNumber="458" precision="925" value="37.25">37°15'N</geoCoordinate>
/
<geoCoordinate id="03A160FCFF99FFA8BB18A5663BFBE675" box="[1318,1431,536,563]" degrees="119" direction="west" minutes="30" orientation="longitude" pageId="7" pageNumber="458" precision="925" value="-119.5">119°30'W</geoCoordinate>
, no date, no collector (
<collectionCode id="00849EFEFF99FFA8BFB5A5423F8BE610" box="[395,487,572,598]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="7" pageNumber="458">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEF9A51E3BFFE008" pageId="7" pageNumber="458" type="diagnosis">
<paragraph id="662A063BFF99FFA8BEF9A51E3BFFE008" blockId="7.[151,1437,535,1570]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A51E3F2EE63C" bold="true" box="[199,322,608,634]" pageId="7" pageNumber="458">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF99FFA8BFE4A51E3C28E63C" box="[474,580,608,634]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BFE4A51E3C28E63C" box="[474,580,608,634]" italics="true" pageId="7" pageNumber="458">C. devius</emphasis>
</taxonomicName>
is unknown. The female of
<taxonomicName id="A1957DB8FF99FFA8BDBBA51E3D82E63C" box="[901,1006,608,634]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BDBBA51E3D82E63C" box="[901,1006,608,634]" italics="true" pageId="7" pageNumber="458">C. devius</emphasis>
</taxonomicName>
is only likely to be confused with the females of
<taxonomicName id="A1957DB8FF99FFA8BF26A5FA3F1CE6D8" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[280,368,644,670]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BF26A5FA3F1CE6D8" box="[280,368,644,670]" italics="true" pageId="7" pageNumber="458">C. echo</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BF47A5FA3F82E6D8" bold="true" box="[377,494,644,670]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BF47A5FA3F82E6D8" box="[377,494,644,670]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
or
<taxonomicName id="A1957DB8FF99FFA8BC26A5FA3CF6E6D8" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[536,666,644,670]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BC26A5FA3CF6E6D8" box="[536,666,644,670]" italics="true" pageId="7" pageNumber="458">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BC9DA5FA3D7BE6D8" bold="true" box="[675,791,644,670]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BC9DA5FA3D7BE6D8" box="[675,791,644,670]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
with which it shares very short spermathecal stalks and the unique copulatory duct morphology of a heavily sclerotized apical section of each copulatory duct enveloped by a very lightly sclerotized, almost diaphanous, basal section (
<figureCitation id="FEAE1ABEFF99FFA8BD4BA5B23A23E6A0" box="[885,1103,716,743]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Figs 15–16, 18–19</figureCitation>
,
<figureCitation id="FEAE1ABEFF99FFA8BA60A5B23AC4E6A0" box="[1118,1192,716,742]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="7" pageNumber="458">21–22</figureCitation>
). In the female of
<taxonomicName id="A1957DB8FF99FFA8BBBDA5B23EB3E74C" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BBBDA5B23EB3E74C" italics="true" pageId="7" pageNumber="458">C. devius</emphasis>
</taxonomicName>
(
<figureCitation id="FEAE1ABEFF99FFA8BED1A58E3F1DE74D" box="[239,369,752,779]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Figs 15–16</figureCitation>
) the basal section of each copulatory duct is voluminous, makes three loops around the more heavily sclerotized apical section, and occupies a greater area than (and, in ventral view, partially obscures) each relatively small spermathecal base. In the female of
<taxonomicName id="A1957DB8FF99FFA8BCD6A4463D05E714" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[744,873,824,850]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BCD6A4463D05E714" box="[744,873,824,850]" italics="true" pageId="7" pageNumber="458">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BD4FA4463D89E714" bold="true" box="[881,997,824,850]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BD4FA4463D89E714" box="[881,997,824,850]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="FEAE1ABEFF99FFA8BDCBA4463A1AE714" box="[1013,1142,824,850]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Figs 18–19</figureCitation>
) the basal section of each copulatory duct is also voluminous but is looped only twice around the apical section. In the female of
<taxonomicName id="A1957DB8FF99FFA8BB39A4223B30E730" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1287,1372,860,886]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BB39A4223B30E730" box="[1287,1372,860,886]" italics="true" pageId="7" pageNumber="458">C. echo</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BB5CA4223EA4E7DC" bold="true" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BB5CA4223EA4E7DC" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="FEAE1ABEFF99FFA8BEE6A4FE3F36E7DC" box="[216,346,896,922]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="7" pageNumber="458">Figs 21–22</figureCitation>
) the basal section of each copulatory duct loops three times around the apical section but is not voluminous (the basal and apical sections are of similar diameter) and occupies a much smaller area. Furthermore, the spermathecal bases are large and unobscured in females of
<taxonomicName id="A1957DB8FF99FFA8BD74A4B63DF3E7A4" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[842,927,968,994]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BD74A4B63DF3E7A4" box="[842,927,968,994]" italics="true" pageId="7" pageNumber="458">C. echo</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BD9AA4B63A7AE7A4" bold="true" box="[932,1046,968,994]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BD9AA4B63A7AE7A4" box="[932,1046,968,994]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="FEAE1ABEFF99FFA8BA1DA4B63ACEE7A4" box="[1059,1186,968,994]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="7" pageNumber="458">Figs 21–22</figureCitation>
) and
<taxonomicName id="A1957DB8FF99FFA8BAE0A4B63B30E7A4" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1246,1372,968,994]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BAE0A4B63B30E7A4" box="[1246,1372,968,994]" italics="true" pageId="7" pageNumber="458">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BB5CA4B63EA4E040" bold="true" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BB5CA4B63EA4E040" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="FEAE1ABEFF99FFA8BEE6A4923F36E040" box="[216,346,1004,1030]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Figs 18–19</figureCitation>
) bases. Finally, the basal section of each copulatory duct in
<taxonomicName id="A1957DB8FF99FFA8BA36A4923A1DE040" box="[1032,1137,1004,1030]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BA36A4923A1DE040" box="[1032,1137,1004,1030]" italics="true" pageId="7" pageNumber="458">C. devius</emphasis>
</taxonomicName>
travels from the atrium to the anterior margin of the vulva before beginning to loop around the apical section (
<figureCitation id="FEAE1ABEFF99FFA8BA7EA36E3AFDE06D" box="[1088,1169,1040,1067]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Fig. 15</figureCitation>
) (
<emphasis id="54E1DA29FF99FFA8BA94A36F3A9EE06C" box="[1194,1266,1041,1066]" italics="true" pageId="7" pageNumber="458">versus</emphasis>
travelling only partway to the anterior margin before looping in
<taxonomicName id="A1957DB8FF99FFA8BC84A34A3D65E008" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[698,777,1076,1102]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BC84A34A3D65E008" box="[698,777,1076,1102]" italics="true" pageId="7" pageNumber="458">C echo</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BD2EA34A3DEEE008" bold="true" box="[784,898,1076,1102]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BD2EA34A3DEEE008" box="[784,898,1076,1102]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
[
<figureCitation id="FEAE1ABEFF99FFA8BDAEA34A3D88E008" box="[912,996,1075,1102]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="7" pageNumber="458">Fig. 21</figureCitation>
] and
<taxonomicName id="A1957DB8FF99FFA8BA1FA34A3ACDE008" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1057,1185,1076,1102]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BA1FA34A3ACDE008" box="[1057,1185,1076,1102]" italics="true" pageId="7" pageNumber="458">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BA96A34A3B76E008" bold="true" box="[1192,1306,1076,1102]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BA96A34A3B76E008" box="[1192,1306,1076,1102]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
[
<figureCitation id="FEAE1ABEFF99FFA8BB16A34A3B12E008" box="[1320,1406,1075,1102]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Fig. 18</figureCitation>
]).
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEF9A3263B71E19C" pageId="7" pageNumber="458" type="description">
<paragraph id="662A063BFF99FFA8BEF9A3263D7FE034" blockId="7.[151,1437,535,1570]" box="[199,787,1112,1139]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A3263F35E034" bold="true" box="[199,345,1112,1138]" pageId="7" pageNumber="458">Description.</emphasis>
Ventral tibia I macrosetae 2-1p-2-1p-0.
</paragraph>
<paragraph id="662A063BFF99FFA8BEF9A3053ABFE0FD" blockId="7.[151,1437,535,1570]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A3053F4BE0D0" bold="true" box="[199,295,1147,1174]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A3053F72E0D0" bold="true" box="[199,286,1147,1174]" italics="true" pageId="7" pageNumber="458">Female</emphasis>
:
</emphasis>
(n=1). Atrium (
<figureCitation id="FEAE1ABEFF99FFA8BFDAA3023C04E0D1" box="[484,616,1148,1175]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Figs 14–15</figureCitation>
) very small, about 1/4 width of vulva. Spermathecal heads apparently protruding dorsally from vulval ducts from within region enclosed by copulatory ducts (
<figureCitation id="FEAE1ABEFF99FFA8BA4CA3DE3AA9E0FD" box="[1138,1221,1184,1211]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="7" pageNumber="458">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="662A063BFF99FFA8BEF9A3BB3D0BE098" blockId="7.[151,1437,535,1570]" box="[199,871,1220,1247]" pageId="7" pageNumber="458">Measurements (n=1). CL 2.11, CW 1.44, SL 1.04, SW 1.01.</paragraph>
<paragraph id="662A063BFF99FFA8BEF9A3963A5AE160" blockId="7.[151,1437,535,1570]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A3963F64E144" bold="true" box="[199,264,1256,1282]" pageId="7" pageNumber="458">Note:</emphasis>
<taxonomicName id="A1957DB8FF99FFA8BF31A3963FC7E144" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[271,427,1256,1282]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BF31A3963FC7E144" box="[271,427,1256,1282]" italics="true" pageId="7" pageNumber="458">Cybaeus echo</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BF8CA3963C48E144" bold="true" box="[434,548,1256,1282]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BF8CA3963C48E144" box="[434,548,1256,1282]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and perhaps
<taxonomicName id="A1957DB8FF99FFA8BC86A3963D54E144" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[696,824,1256,1282]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF99FFA8BC86A3963D54E144" box="[696,824,1256,1282]" italics="true" pageId="7" pageNumber="458">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF99FFA8BD00A3963DDCE144" bold="true" box="[830,944,1256,1282]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="4FD26752FF99FFA8BD00A3963DDCE144" box="[830,944,1256,1282]" pageId="7" pageNumber="458" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
may be synonyms of
<taxonomicName id="A1957DB8FF99FFA8BA98A3963B62E144" box="[1190,1294,1256,1282]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF99FFA8BA98A3963B62E144" box="[1190,1294,1256,1282]" italics="true" pageId="7" pageNumber="458">C. devius</emphasis>
</taxonomicName>
. Insufficient specimens are available to make a strongly supported conclusion of conspecificity.
</paragraph>
<paragraph id="662A063BFF99FFA8BEF9A24E3B71E19C" blockId="7.[151,1437,535,1570]" pageId="7" pageNumber="458">
Although its coloration indicates the single specimen of
<taxonomicName id="A1957DB8FF99FFA8BD79A24E3DF2E10C" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[839,926,1328,1354]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="echo">
<emphasis id="54E1DA29FF99FFA8BD79A24E3DF2E10C" box="[839,926,1328,1354]" italics="true" pageId="7" pageNumber="458">C. echo</emphasis>
</taxonomicName>
is fully matured the vulva at least appears to be teneral with thin-walled ducts and weakly developed Bennett’s glands. Likewise the three known females of
<taxonomicName id="A1957DB8FF99FFA8BBBDA22A3E9AE1D4" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wilsonia">
<emphasis id="54E1DA29FF99FFA8BBBDA22A3E9AE1D4" italics="true" pageId="7" pageNumber="458">C. wilsonia</emphasis>
</taxonomicName>
all have apparently teneral vulvae. Clearly more specimens (especially males) from across the entire Sierra
<collectingRegion id="A451C8D9FF99FFA8BEA9A2E23E81E1F0" box="[151,237,1436,1462]" country="United States of America" name="Nevada" pageId="7" pageNumber="458">Nevada</collectingRegion>
range of these three putative species need to be examined to analyze variation. At this time we consider the specimens examined here to be sufficiently distinct to warrant the recognition of three separate species.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF99FFA8BEF9A29A3FFDE264" pageId="7" pageNumber="458" type="distribution">
<paragraph id="662A063BFF99FFA8BEF9A29A3FFDE264" blockId="7.[151,1437,535,1570]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A29A3F35E1B8" bold="true" box="[199,345,1508,1534]" pageId="7" pageNumber="458">Distribution</emphasis>
. (
<figureCitation id="FEAE1ABEFF99FFA8BF4DA29A3FA6E1B8" box="[371,458,1508,1534]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="7" pageNumber="458">Fig. 32</figureCitation>
). Known only from the
<typeStatus id="B92EB899FF99FFA8BCDAA29B3D78E1B9" box="[740,788,1509,1535]" pageId="7" pageNumber="458">type</typeStatus>
locality in Madera County on the western slopes of the central Sierra
<collectingRegion id="A451C8D9FF99FFA8BF0AA1763FE1E264" box="[308,397,1544,1570]" country="United States of America" name="Nevada" pageId="7" pageNumber="458">Nevada</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="662A063BFF9AFFABBEA9A3073C14E0D5" blockId="4.[151,632,1145,1208]" box="[151,632,1145,1172]" pageId="4" pageNumber="455">
<heading id="3D62B157FF9AFFABBEA9A3073C14E0D5" bold="true" box="[151,632,1145,1172]" fontSize="11" level="1" pageId="4" pageNumber="455" reason="1">
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<taxonomicName id="A1957DB8FF9AFFABBEA9A3073F95E0D5" ID-CoL="99CCW" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,505,1145,1172]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBEA9A3073FFCE0D2" bold="true" box="[151,400,1145,1172]" italics="true" pageId="4" pageNumber="455">Cybaeus bryoncavus</emphasis>
Bennett
</taxonomicName>
<taxonomicNameLabel id="4FD26752FF9AFFABBC3EA3073C14E0D5" box="[512,632,1145,1171]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9AFFABBEA9A3E33FA5E0F1" box="[151,457,1181,1208]" pageId="4" pageNumber="455" type="description">
<paragraph id="662A063BFF9AFFABBEA9A3E33FA5E0F1" blockId="4.[151,632,1145,1208]" box="[151,457,1181,1208]" pageId="4" pageNumber="455">
<figureCitation id="FEAE1ABEFF9AFFABBEA9A3E33E97E0FE" box="[151,251,1181,1208]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="4" pageNumber="455">Figs 5–7</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBF38A3E33F22E0F1" box="[262,334,1181,1207]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">11–13</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBF67A3E33F1BE0F1" box="[345,375,1181,1207]" captionStart="FIGURES 26–28" captionStartId="11.[151,264,1216,1240]" captionTargetBox="[151,1436,705,1193]" captionTargetId="figure-226@11.[151,1436,705,1193]" captionTargetPageId="11" captionText="FIGURES 26–28. Cybaeus spp. males, left palpus, patellar apophysis, dorsal (26 C. bilectus spec. nov., from Double Bunk Meadows, California, 27 C. bryoncavus spec. nov., holotype, 28 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (26 concave distal margin; 28 concave dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.5300646" httpUri="https://zenodo.org/record/5300646/files/figure.png" pageId="4" pageNumber="455">27</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBFBDA3E33FCDE0F1" box="[387,417,1181,1207]" captionStart="FIGURES 29–31" captionStartId="11.[151,264,1858,1882]" captionTargetBox="[151,1436,1337,1834]" captionTargetId="figure-280@11.[151,1436,1337,1834]" captionTargetPageId="11" captionText="FIGURES 29–31. Cybaeus spp. males, left palpus, proximal arm of tegular apophysis, ventral (29 C. bilectus spec. nov., from Double Bunk Meadows, California, 30 C. bryoncavus spec. nov., holotype, 31 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (29 bluntly hooked tip, 30 inconspicuous basal projection)." figureDoi="http://doi.org/10.5281/zenodo.5300649" httpUri="https://zenodo.org/record/5300649/files/figure.png" pageId="4" pageNumber="455">30</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBF93A3E33FA5E0F1" box="[429,457,1181,1207]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="4" pageNumber="455">32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9AFFABBEA9A3A13D25E107" pageId="4" pageNumber="455" type="materials_examined">
<paragraph id="662A063BFF9AFFABBEA9A3A13D25E107" blockId="4.[151,1437,1246,2029]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEA9A3A13F27E0BE" bold="true" box="[151,331,1246,1273]" pageId="4" pageNumber="455">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF9AFFABBF67A3A03FB8E15B" collectingDate="1987-10-17" collectionCode="CAS" collectorName="D. Ubick" country="United States of America" county="Amador County" location="Volcano" municipality="Moss Cave" pageId="4" pageNumber="455" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF9AFFABBF67A3A03D69E0BF" bold="true" box="[345,773,1246,1273]" pageId="4" pageNumber="455">
<collectingCountry id="1E8246ABFF9AFFABBF67A3A03FC5E0BE" box="[345,425,1246,1272]" name="United States of America" pageId="4" pageNumber="455">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF9AFFABBF82A3A03C58E0BF" bold="true" box="[444,564,1246,1273]" italics="true" pageId="4" pageNumber="455">
<collectingRegion id="A451C8D9FF9AFFABBF82A3A03C58E0BF" box="[444,564,1246,1273]" country="United States of America" name="California" pageId="4" pageNumber="455">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF9AFFABBC75A3A13CD4E0BF" box="[587,696,1247,1273]" pageId="4" pageNumber="455" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<collectingCounty id="8F4B7EB7FF9AFFABBD2FA3A13DA2E0BF" box="[785,974,1247,1273]" pageId="4" pageNumber="455">Amador County</collectingCounty>
,
<collectingMunicipality id="864E9C41FF9AFFABBDE1A3A13A09E0BF" box="[991,1125,1247,1273]" pageId="4" pageNumber="455">Moss Cave</collectingMunicipality>
, near
<location id="634A50E0FF9AFFABBA8AA3A13B7EE0BF" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9AFFA8BEA9A3073C8CE4BF:634A50E0FF9AFFABBA8AA3A13B7EE0BF" box="[1204,1298,1247,1273]" country="United States of America" county="Amador County" municipality="Moss Cave" name="Volcano" pageId="4" pageNumber="455" stateProvince="California">Volcano</location>
,
<location id="634A50E0FF9AFFABBB1AA3A13BF4E0BF" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9AFFA8BEA9A3073C8CE4BF:634A50E0FF9AFFABBB1AA3A13BF4E0BF" box="[1316,1432,1247,1273]" country="United States of America" county="Amador County" municipality="Moss Cave" name="California" pageId="4" pageNumber="455" stateProvince="California">California</location>
,
<date id="122B20FBFF9AFFABBEA9A27D3F66E15B" box="[151,266,1283,1309]" pageId="4" pageNumber="455" value="1987-10-17">
<collectingDate id="026FD913FF9AFFABBEA9A27D3F66E15B" box="[151,266,1283,1309]" pageId="4" pageNumber="455" value="1987-10-17">17.x.1987</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9AFFABBF2BA27D3F11E15B" box="[277,381,1283,1309]" pageId="4" pageNumber="455">D. Ubick</collectorName>
, (
<collectionCode id="00849EFEFF9AFFABBFAEA27D3FA0E15B" box="[400,460,1283,1309]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="4" pageNumber="455" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="D6FD0C66FF9AFFABBFE1A27D3BF4E15B" box="[479,1432,1282,1309]" collectingDate="1958-07-07" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; V. D. Roth" country="United States of America" location="Pine Grove" pageId="4" pageNumber="455" specimenCount="4" specimenCount-female="4" stateProvince="California" typeStatus="paratype">
<emphasis id="54E1DA29FF9AFFABBFE1A27D3C31E15B" bold="true" box="[479,605,1283,1309]" pageId="4" pageNumber="455">
<typeStatus id="B92EB899FF9AFFABBFE1A27D3C35E15B" box="[479,601,1283,1309]" pageId="4" pageNumber="455" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Amador.
<specimenCount id="7093CDB2FF9AFFABBCF5A27D3C9CE15B" box="[715,752,1283,1309]" pageId="4" pageNumber="455" type="female">4♀</specimenCount>
,
<location id="634A50E0FF9AFFABBCC2A27D3D11E15B" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9AFFA8BEA9A3073C8CE4BF:634A50E0FF9AFFABBCC2A27D3D11E15B" box="[764,893,1283,1309]" country="United States of America" name="Pine Grove" pageId="4" pageNumber="455" stateProvince="California">Pine Grove</location>
,
<date id="122B20FBFF9AFFABBDB9A27C3D97E15B" box="[903,1019,1282,1309]" pageId="4" pageNumber="455" value="1958-07-07">
<collectingDate id="026FD913FF9AFFABBDB9A27C3D97E15B" box="[903,1019,1282,1309]" pageId="4" pageNumber="455" value="1958-07-07">7.vii.1958</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9AFFABBA38A27D3AF8E15B" box="[1030,1172,1283,1309]" pageId="4" pageNumber="455">W.J. Gertsch</collectorName>
&amp;
<collectorName id="CB6063EDFF9AFFABBA88A27D3B49E15B" box="[1206,1317,1283,1309]" pageId="4" pageNumber="455">V.D. Roth</collectorName>
(
<collectionCode id="00849EFEFF9AFFABBB0AA27D3BFCE15B" box="[1332,1424,1283,1309]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="455">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF9AFFABBEA9A2593D29E107" box="[151,837,1318,1345]" collectingDate="1995-03-25" collectionCode="CAS" collectorName="D. Ubick" country="United States of America" location="Volcano" pageId="4" pageNumber="455" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF9AFFABBEA9A2593ED6E107" box="[151,186,1319,1345]" pageId="4" pageNumber="455" type="male">1♂</specimenCount>
<specimenCount id="7093CDB2FF9AFFABBEFFA2593E88E107" box="[193,228,1319,1345]" pageId="4" pageNumber="455" type="female">1♀</specimenCount>
,
<quantity id="A16DABDEFF9AFFABBECCA2583F2DE107" box="[242,321,1318,1345]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="4" pageNumber="455" unit="mi" value="0.5">0.5 mi.</quantity>
NW of
<location id="634A50E0FF9AFFABBFA5A2593F95E107" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9AFFA8BEA9A3073C8CE4BF:634A50E0FF9AFFABBFA5A2593F95E107" box="[411,505,1319,1345]" country="United States of America" name="Volcano" pageId="4" pageNumber="455" stateProvince="California">Volcano</location>
,
<date id="122B20FBFF9AFFABBC3AA2583C13E107" box="[516,639,1318,1345]" pageId="4" pageNumber="455" value="1995-03-25">
<collectingDate id="026FD913FF9AFFABBC3AA2583C13E107" box="[516,639,1318,1345]" pageId="4" pageNumber="455" value="1995-03-25">25.iii.1995</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9AFFABBCB5A2593C9EE107" box="[651,754,1319,1345]" pageId="4" pageNumber="455">D. Ubick</collectorName>
(
<collectionCode id="00849EFEFF9AFFABBD3FA2593D51E107" box="[769,829,1319,1345]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="4" pageNumber="455" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9AFFABBEF9A2353DBEE1CF" pageId="4" pageNumber="455" type="etymology">
<paragraph id="662A063BFF9AFFABBEF9A2353DBEE1CF" blockId="4.[151,1437,1246,2029]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A2353F22E123" bold="true" box="[199,334,1355,1381]" pageId="4" pageNumber="455">Etymology.</emphasis>
The specific epithet, derived from a combination of the Greek (
<emphasis id="54E1DA29FF9AFFABBA2EA2353A3DE123" box="[1040,1105,1355,1381]" italics="true" pageId="4" pageNumber="455">bryon</emphasis>
) and Latin (
<emphasis id="54E1DA29FF9AFFABBAE4A2353B4FE122" box="[1242,1315,1355,1380]" italics="true" pageId="4" pageNumber="455">cavum</emphasis>
) nouns for “moss” and “hole” respectively, refers to the
<typeStatus id="B92EB899FF9AFFABBCAEA2113CACE1CF" box="[656,704,1391,1417]" pageId="4" pageNumber="455">type</typeStatus>
locality for this species.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9AFFABBEF9A2EC3A9DE28B" pageId="4" pageNumber="455" type="diagnosis">
<paragraph id="662A063BFF9AFFABBEF9A2EC3D4AE2C3" blockId="4.[151,1437,1246,2029]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A2EC3F2EE1EA" bold="true" box="[199,322,1426,1452]" pageId="4" pageNumber="455">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF9AFFABBFEBA2ED3C1BE1EB" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[469,631,1427,1453]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBFEBA2ED3C1BE1EB" box="[469,631,1427,1453]" italics="true" pageId="4" pageNumber="455">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBC43A2ED3C9CE1EB" bold="true" box="[637,752,1427,1453]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBC43A2ED3C9CE1EB" box="[637,752,1427,1453]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is unlikely to be confused with the males of
<taxonomicName id="A1957DB8FF9AFFABBADAA2ED3B37E1EB" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1252,1371,1427,1453]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBADAA2ED3B37E1EB" box="[1252,1371,1427,1453]" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBB5CA2ED3EA4E197" bold="true" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBB5CA2ED3EA4E197" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
or
<taxonomicName id="A1957DB8FF9AFFABBECEA2C93F1DE197" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[240,369,1463,1489]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBECEA2C93F1DE197" box="[240,369,1463,1489]" italics="true" pageId="4" pageNumber="455">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBF47A2C93F80E197" bold="true" box="[377,492,1463,1489]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBF47A2C93F80E197" box="[377,492,1463,1489]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, the other members of the
<taxonomicName id="A1957DB8FF9AFFABBD1EA2C93D04E197" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[800,872,1463,1489]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9AFFABBD1EA2C93D04E197" box="[800,872,1463,1489]" italics="true" pageId="4" pageNumber="455">devius</emphasis>
</taxonomicName>
species group with known males. The male of
<taxonomicName id="A1957DB8FF9AFFABBBBDA2C93F74E1B3" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBBBDA2C93F74E1B3" italics="true" pageId="4" pageNumber="455">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBF1EA2A53FFFE1B3" bold="true" box="[288,403,1499,1525]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBF1EA2A53FFFE1B3" box="[288,403,1499,1525]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is diagnosed by the form of the patellar apophysis (
<figureCitation id="FEAE1ABEFF9AFFABBDE0A2A53A4BE1B3" box="[990,1063,1498,1525]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="4" pageNumber="455">Figs 5</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBA0AA2A53A3DE1B3" box="[1076,1105,1499,1525]" captionStart="FIGURES 26–28" captionStartId="11.[151,264,1216,1240]" captionTargetBox="[151,1436,705,1193]" captionTargetId="figure-226@11.[151,1436,705,1193]" captionTargetPageId="11" captionText="FIGURES 26–28. Cybaeus spp. males, left palpus, patellar apophysis, dorsal (26 C. bilectus spec. nov., from Double Bunk Meadows, California, 27 C. bryoncavus spec. nov., holotype, 28 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (26 concave distal margin; 28 concave dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.5300646" httpUri="https://zenodo.org/record/5300646/files/figure.png" pageId="4" pageNumber="455">27</figureCitation>
) and the proximal arm of the tegular apophysis (
<figureCitation id="FEAE1ABEFF9AFFABBF51A2813FBBE25F" box="[367,471,1535,1561]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="4" pageNumber="455">Figs 6–7</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBFDBA2813C69E25F" box="[485,517,1535,1561]" captionStart="FIGURES 29–31" captionStartId="11.[151,264,1858,1882]" captionTargetBox="[151,1436,1337,1834]" captionTargetId="figure-280@11.[151,1436,1337,1834]" captionTargetPageId="11" captionText="FIGURES 29–31. Cybaeus spp. males, left palpus, proximal arm of tegular apophysis, ventral (29 C. bilectus spec. nov., from Double Bunk Meadows, California, 30 C. bryoncavus spec. nov., holotype, 31 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (29 bluntly hooked tip, 30 inconspicuous basal projection)." figureDoi="http://doi.org/10.5281/zenodo.5300649" httpUri="https://zenodo.org/record/5300649/files/figure.png" pageId="4" pageNumber="455">30</figureCitation>
): patellar apophysis moderately long (length about 2/3 the width of the patella), antero-laterally directed, lacking concave areas, and with about 25 to 30 peg setae dorsally along the length of the distal margin; the proximal arm of the tegular apophysis has a small, inconspicuous, posteriorly directed basal projection and an elongate, acuminate, slightly twisted tip.
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A1F13A9DE28B" blockId="4.[151,1437,1246,2029]" pageId="4" pageNumber="455">
The female of
<taxonomicName id="A1957DB8FF9AFFABBF56A1F13C65E2EF" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[360,521,1679,1705]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBF56A1F13C65E2EF" box="[360,521,1679,1705]" italics="true" pageId="4" pageNumber="455">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBC31A1F13CECE2EF" bold="true" box="[527,640,1679,1705]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBC31A1F13CECE2EF" box="[527,640,1679,1705]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="FEAE1ABEFF9AFFABBCB0A1F13D67E2EF" box="[654,779,1679,1705]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">Figs 11–13</figureCitation>
) is only likely to be confused with the female of
<taxonomicName id="A1957DB8FF9AFFABBB18A1F13BF0E2EF" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1318,1436,1679,1705]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBB18A1F13BF0E2EF" box="[1318,1436,1679,1705]" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBEA9A1CD3F66E28B" bold="true" box="[151,266,1715,1741]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBEA9A1CD3F66E28B" box="[151,266,1715,1741]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
; distinguishing the females of these two species is discussed under
<taxonomicName id="A1957DB8FF9AFFABBA3EA1CD3A1BE28B" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1024,1143,1715,1741]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBA3EA1CD3A1BE28B" box="[1024,1143,1715,1741]" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBA40A1CD3A9DE28B" bold="true" box="[1150,1265,1715,1741]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBA40A1CD3A9DE28B" box="[1150,1265,1715,1741]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9AFFABBEF9A1A83BF0E3AB" pageId="4" pageNumber="455" type="description">
<paragraph id="662A063BFF9AFFABBEF9A1A83BEEE2B7" blockId="4.[151,1437,1246,2029]" box="[199,1410,1750,1778]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A1A83F3AE2B6" bold="true" box="[199,342,1750,1776]" pageId="4" pageNumber="455">Description.</emphasis>
Femora unbanded (
<typeStatus id="B92EB899FF9AFFABBC11A1A93CFEE2B4" box="[559,658,1751,1778]" pageId="4" pageNumber="455" type="holotype">holotype</typeStatus>
) or banded (
<typeStatus id="B92EB899FF9AFFABBD25A1A93DE6E2B7" box="[795,906,1751,1777]" pageId="4" pageNumber="455" type="paratype">paratypes</typeStatus>
). Ventral tibia I macrosetae 2-1p-2-1p-0(or 2).
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A1843AC4E37F" blockId="4.[151,1437,1246,2029]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A1843F67E353" bold="true" box="[199,267,1786,1813]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A1843F6EE353" bold="true" box="[199,258,1786,1813]" italics="true" pageId="4" pageNumber="455">Male</emphasis>
:
</emphasis>
(n=2). Distal arm of tegular apophysis (
<figureCitation id="FEAE1ABEFF9AFFABBCF4A1853D61E350" box="[714,781,1787,1814]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="4" pageNumber="455">Fig. 6</figureCitation>
) broad basally, elongate (but length less than twice width), gradually narrowing to tip. Embolus narrow; tip apparently broken off in
<typeStatus id="B92EB899FF9AFFABBDF3A0613A43E37F" box="[973,1071,1823,1849]" pageId="4" pageNumber="455" type="holotype">holotype</typeStatus>
specimen.
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A03D3D36E31B" blockId="4.[151,1437,1246,2029]" box="[199,858,1859,1885]" pageId="4" pageNumber="455">Measurements (n=1). CL 2.7, CW 2.03, SL 1.31, SW 1.29.</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A0183BF1E38F" blockId="4.[151,1437,1246,2029]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A0183F4BE3C7" bold="true" box="[199,295,1894,1921]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A0183F72E3C7" bold="true" box="[199,286,1894,1921]" italics="true" pageId="4" pageNumber="455">Female</emphasis>
:
</emphasis>
(n=5).Atrium (
<figureCitation id="FEAE1ABEFF9AFFABBFECA0193C22E3C7" box="[466,590,1895,1921]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">Figs 11–12</figureCitation>
) deep but small, about 1/2 width of vulva.Copulatory ducts and spermathecal ducts (
<figureCitation id="FEAE1ABEFF9AFFABBEE1A0F53F31E3E3" box="[223,349,1931,1957]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">Figs 12–13</figureCitation>
) simple but elongate, convoluted. Copulatory ducts proceed anteriorly along mid-line of vulva then diverge and intertwine with spermathecal stalks.Stalks with one coil looped around medial section of copulatory ducts.
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A0AD3BF0E3AB" blockId="4.[151,1437,1246,2029]" box="[199,1436,2002,2029]" pageId="4" pageNumber="455">Measurements (n=4). CL 2.30–3.0 (2.5), CW 1.63–2.15 (1.80), SL 1.13–1.40 (1.22), SW 1.09–1.38 (1.18).</paragraph>
</subSubSection>
<caption id="32EA56B3FF9BFFAABEA9A0133A6DE3B4" ID-DOI="http://doi.org/10.5281/zenodo.5300634" ID-Zenodo-Dep="5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="5" pageNumber="456" startId="5.[151,264,1901,1925]" targetBox="[215,1372,181,1842]" targetPageId="5">
<paragraph id="662A063BFF9BFFAABEA9A0133A6DE3B4" blockId="5.[151,1436,1901,2034]" pageId="5" pageNumber="456">
<emphasis id="54E1DA29FF9BFFAABEA9A0133F56E3C3" bold="true" box="[151,314,1901,1925]" pageId="5" pageNumber="456">FIGURES 5–7.</emphasis>
<taxonomicName id="A1957DB8FF9BFFAABF7EA0133C7EE3C3" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[320,530,1901,1925]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9BFFAABF7EA0133C7EE3C3" box="[320,530,1901,1925]" italics="true" pageId="5" pageNumber="456">Cybaeus bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9BFFAABC26A0133C12E3C3" bold="true" box="[536,638,1901,1925]" pageId="5" pageNumber="456">
<taxonomicNameLabel id="4FD26752FF9BFFAABC26A0133C12E3C3" box="[536,638,1901,1925]" pageId="5" pageNumber="456" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA.
</paragraph>
</caption>
<caption id="32EA56B3FF98FFA9BEA9A0103A91E388" ID-DOI="http://doi.org/10.5281/zenodo.5300636" ID-Zenodo-Dep="5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="6" pageNumber="457" startId="6.[151,264,1902,1926]" targetBox="[152,1436,190,1878]" targetPageId="6">
<paragraph id="662A063BFF98FFA9BEA9A0103A91E388" blockId="6.[151,1436,1902,1998]" pageId="6" pageNumber="457">
<emphasis id="54E1DA29FF98FFA9BEA9A0103F25E3C0" bold="true" box="[151,329,1902,1926]" pageId="6" pageNumber="457">FIGURES 8–13.</emphasis>
<taxonomicName id="A1957DB8FF98FFA9BF6CA0103FB1E3C3" box="[338,477,1902,1926]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="54E1DA29FF98FFA9BF6CA0103FC5E3C0" box="[338,425,1902,1926]" italics="true" pageId="6" pageNumber="457">Cybaeus</emphasis>
spp.
</taxonomicName>
, females from California, copulatory organ (8–10
<taxonomicName id="A1957DB8FF98FFA9BDD4A0103A34E3C0" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1002,1112,1902,1926]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF98FFA9BDD4A0103A34E3C0" box="[1002,1112,1902,1926]" italics="true" pageId="6" pageNumber="457">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF98FFA9BA5EA0103AA6E3C0" bold="true" box="[1120,1226,1902,1926]" pageId="6" pageNumber="457">
<taxonomicNameLabel id="4FD26752FF98FFA9BA5EA0103AA6E3C0" box="[1120,1226,1902,1926]" pageId="6" pageNumber="457" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, from Double Bunk Meadows; 11–13
<taxonomicName id="A1957DB8FF98FFA9BF73A0EC3F8CE3EC" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[333,480,1938,1962]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF98FFA9BF73A0EC3F8CE3EC" box="[333,480,1938,1962]" italics="true" pageId="6" pageNumber="457">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF98FFA9BFD8A0EC3C21E3EC" bold="true" box="[486,589,1938,1962]" pageId="6" pageNumber="457">
<taxonomicNameLabel id="4FD26752FF98FFA9BFD8A0EC3C21E3EC" box="[486,589,1938,1962]" pageId="6" pageNumber="457" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca.
</paragraph>
</caption>
<subSubSection id="2E8F55B0FF99FFA8BEF9A7E83C8CE4BF" pageId="7" pageNumber="458" type="distribution">
<paragraph id="662A063BFF99FFA8BEF9A7E83C8CE4BF" blockId="7.[151,1436,150,249]" pageId="7" pageNumber="458">
<emphasis id="54E1DA29FF99FFA8BEF9A7E83C20E4F6" bold="true" box="[199,588,150,177]" pageId="7" pageNumber="458">Distribution and natural history.</emphasis>
(
<figureCitation id="FEAE1ABEFF99FFA8BC65A7E93CC2E4F7" box="[603,686,151,177]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="7" pageNumber="458">Fig. 32</figureCitation>
). Known only from the area around the
<typeStatus id="B92EB899FF99FFA8BA57A7E93AF5E4F7" box="[1129,1177,151,177]" pageId="7" pageNumber="458">type</typeStatus>
locality on the western slopes of the central Sierra
<collectingRegion id="A451C8D9FF99FFA8BFEFA7C53C46E493" box="[465,554,187,213]" country="United States of America" name="Nevada" pageId="7" pageNumber="458">Nevada</collectingRegion>
. The
<specimenCount id="7093CDB2FF99FFA8BC54A7C53CB2E493" box="[618,734,187,213]" pageId="7" pageNumber="458" type="male">two males</specimenCount>
were collected in March and October. The Volcano male and female were found under limestone in an oak forest.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,415 @@
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<mods:title id="3F9306F7361A387A4EF46094A8A52EBC">Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade</mods:title>
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<mods:number id="07C57F3AD7603B9059EAAAFE3A9D7A19">2021-08-27</mods:number>
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<mods:detail id="A4FE442EAE186FFC56473971AB89904B" type="volume">
<mods:number id="063263A9B81CA9A0B985FBC3D1521AE1">5026</mods:number>
</mods:detail>
<mods:detail id="320D3936FD48B2AE5712550BC07E28FD" type="issue">
<mods:number id="5CC965E21D29E34FB76C14A42A2AC217">4</mods:number>
</mods:detail>
<mods:extent id="4A470D816A9C215A0FB5784A5843D432" unit="page">
<mods:start id="8C3B3BAD1C178A0F4FCF8B211DB3DA65">451</mods:start>
<mods:end id="1C37558CDE75C791B974D4DACBF348B2">479</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="40A15B87395FBC4295FBCE5134C0B657">journal article</mods:classification>
<mods:identifier id="C2D624983DA99E4BEBA51B461BEBF40D" type="DOI">10.11646/zootaxa.5026.4.1</mods:identifier>
<mods:identifier id="3C15AAC7E747D5DA86AB1B124283B7EA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A5E3F6FE75A6F1024566FC8161CE4CD0" type="Zenodo-Dep">5300630</mods:identifier>
<mods:identifier id="6A0D385FBE91C82E5A51232301C5BFA5" type="ZooBank">511C28F6-07E4-412C-B83E-91D2CAC5AA27</mods:identifier>
</mods:mods>
<treatment id="EE3CB72DFF9CFFABBEA9A19D3D90E073" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073" httpUri="http://treatment.plazi.org/id/EE3CB72DFF9CFFABBEA9A19D3D90E073" lastPageId="4" lastPageNumber="455" pageId="2" pageNumber="453">
<subSubSection id="2E8F55B0FF9CFFADBEA9A19D3C26E2B8" box="[151,586,1763,1790]" pageId="2" pageNumber="453" type="nomenclature">
<paragraph id="662A063BFF9CFFADBEA9A19D3C26E2B8" blockId="2.[151,586,1763,1826]" box="[151,586,1763,1790]" pageId="2" pageNumber="453">
<heading id="3D62B157FF9CFFADBEA9A19D3C26E2B8" bold="true" box="[151,586,1763,1790]" fontSize="11" level="1" pageId="2" pageNumber="453" reason="1">
<emphasis id="54E1DA29FF9CFFADBEA9A19D3C26E2B8" bold="true" box="[151,586,1763,1790]" pageId="2" pageNumber="453">
<taxonomicName id="A1957DB8FF9CFFADBEA9A19D3FA7E2B8" ID-CoL="99C7V" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,459,1763,1790]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9CFFADBEA9A19D3F0EE2B8" bold="true" box="[151,354,1763,1790]" italics="true" pageId="2" pageNumber="453">Cybaeus bilectus</emphasis>
Bennett
</taxonomicName>
<taxonomicNameLabel id="4FD26752FF9CFFADBFECA19A3C26E2B8" box="[466,586,1764,1790]" pageId="2" pageNumber="453" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9CFFADBEA9A0763FD0E364" box="[151,444,1800,1826]" pageId="2" pageNumber="453" type="description">
<paragraph id="662A063BFF9CFFADBEA9A0763FD0E364" blockId="2.[151,586,1763,1826]" box="[151,444,1800,1826]" pageId="2" pageNumber="453">
<figureCitation id="FEAE1ABEFF9CFFADBEA9A0763E96E364" box="[151,250,1800,1826]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="2" pageNumber="453">Figs 1–4</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBF38A0763F2DE364" box="[262,321,1800,1826]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="2" pageNumber="453">8–10</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBF72A0763F06E364" box="[332,362,1800,1826]" captionStart="FIGURES 26–28" captionStartId="11.[151,264,1216,1240]" captionTargetBox="[151,1436,705,1193]" captionTargetId="figure-226@11.[151,1436,705,1193]" captionTargetPageId="11" captionText="FIGURES 26–28. Cybaeus spp. males, left palpus, patellar apophysis, dorsal (26 C. bilectus spec. nov., from Double Bunk Meadows, California, 27 C. bryoncavus spec. nov., holotype, 28 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (26 concave distal margin; 28 concave dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.5300646" httpUri="https://zenodo.org/record/5300646/files/figure.png" pageId="2" pageNumber="453">26</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBF48A0763FF8E364" box="[374,404,1800,1826]" captionStart="FIGURES 29–31" captionStartId="11.[151,264,1858,1882]" captionTargetBox="[151,1436,1337,1834]" captionTargetId="figure-280@11.[151,1436,1337,1834]" captionTargetPageId="11" captionText="FIGURES 29–31. Cybaeus spp. males, left palpus, proximal arm of tegular apophysis, ventral (29 C. bilectus spec. nov., from Double Bunk Meadows, California, 30 C. bryoncavus spec. nov., holotype, 31 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (29 bluntly hooked tip, 30 inconspicuous basal projection)." figureDoi="http://doi.org/10.5281/zenodo.5300649" httpUri="https://zenodo.org/record/5300649/files/figure.png" pageId="2" pageNumber="453">29</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBF9EA0763FD0E364" box="[416,444,1800,1826]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="2" pageNumber="453">32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9CFFACBEA9A02E3CD0E493" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453" type="materials_examined">
<paragraph id="662A063BFF9CFFADBEA9A02E3BF0E3BC" blockId="2.[151,1436,1871,2043]" pageId="2" pageNumber="453">
<emphasis id="54E1DA29FF9CFFADBEA9A02E3F24E32C" bold="true" box="[151,328,1872,1898]" pageId="2" pageNumber="453">Type material.</emphasis>
<materialsCitation id="D6FD0C66FF9CFFADBF6CA02E3D2AE3C8" collectingDate="1959-09-15" collectionCode="AMNH" collectorName="V. D. Roth &amp; W. J. Gertsch" country="United States of America" location="Johnsondale" pageId="2" pageNumber="453" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="54E1DA29FF9CFFADBF6CA02E3C9AE32C" bold="true" box="[338,758,1871,1898]" pageId="2" pageNumber="453">
<collectingCountry id="1E8246ABFF9CFFADBF6CA02E3FCFE32C" box="[338,419,1872,1898]" name="United States of America" pageId="2" pageNumber="453">U.S.A.</collectingCountry>
:
<emphasis id="54E1DA29FF9CFFADBF8DA0313C47E32C" bold="true" box="[435,555,1871,1898]" italics="true" pageId="2" pageNumber="453">
<collectingRegion id="A451C8D9FF9CFFADBF8DA0313C47E32C" box="[435,555,1871,1898]" country="United States of America" name="California" pageId="2" pageNumber="453">California</collectingRegion>
</emphasis>
:
<typeStatus id="B92EB899FF9CFFADBC00A02E3CC7E32C" box="[574,683,1872,1898]" pageId="2" pageNumber="453" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<taxonomicName id="A1957DB8FF9CFFADBD3EA02E3AABE32C" authority="County, Double Bunk Meadows" authorityName="County, Double Bunk Meadows" box="[768,1223,1872,1898]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="tulare">Tulare County, Double Bunk Meadows</taxonomicName>
, six miles west of
<location id="634A50E0FF9CFFADBEA9A00A3F48E3C8" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9CFFADBEA9A00A3F48E3C8" box="[151,292,1908,1934]" country="United States of America" name="Johnsondale" pageId="2" pageNumber="453" stateProvince="California">Johnsondale</location>
,
<date id="122B20FBFF9CFFADBF11A00A3FC5E3C9" box="[303,425,1908,1935]" pageId="2" pageNumber="453" value="1959-09-15">
<collectingDate id="026FD913FF9CFFADBF11A00A3FC5E3C9" box="[303,425,1908,1935]" pageId="2" pageNumber="453" value="1959-09-15">15.ix.1959</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9CFFADBF8AA00B3C4FE3C8" box="[436,547,1908,1935]" pageId="2" pageNumber="453">V.D. Roth</collectorName>
&amp;
<collectorName id="CB6063EDFF9CFFADBC78A00B3CB8E3C8" box="[582,724,1908,1935]" pageId="2" pageNumber="453">W.J. Gertsch</collectorName>
(
<collectionCode id="00849EFEFF9CFFADBCDCA00A3D52E3C8" box="[738,830,1908,1934]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="453">AMNH</collectionCode>
)
</materialsCitation>
.
<emphasis id="54E1DA29FF9CFFADBD6EA00A3DA3E3C8" bold="true" box="[848,975,1908,1934]" pageId="2" pageNumber="453">
<typeStatus id="B92EB899FF9CFFADBD6EA00A3DA6E3C8" box="[848,970,1908,1934]" pageId="2" pageNumber="453" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Tulare.
<materialsCitation id="D6FD0C66FF9CFFADBA14A00A3C65E3F4" collectingDate="1961-06-18" collectionCode="CAS" collectorName="Maier" country="United States of America" location="10 mi. N of Hwy" pageId="2" pageNumber="453" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF9CFFADBA14A00A3A21E3C8" box="[1066,1101,1908,1934]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
, “dirt road”,
<location id="634A50E0FF9CFFADBADEA00A3BF0E3C9" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9CFFADBADEA00A3BF0E3C9" box="[1248,1436,1908,1935]" country="United States of America" name="10 mi. N of Hwy" pageId="2" pageNumber="453" stateProvince="California">
<quantity id="A16DABDEFF9CFFADBADEA00A3B4BE3C8" box="[1248,1319,1908,1934]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="2" pageNumber="453" unit="mi" value="10.0">10 mi.</quantity>
N of Hwy
</location>
155,
<date id="122B20FBFF9CFFADBEF1A0E63F24E3F4" box="[207,328,1944,1970]" pageId="2" pageNumber="453" value="1961-06-18">
<collectingDate id="026FD913FF9CFFADBEF1A0E63F24E3F4" box="[207,328,1944,1970]" pageId="2" pageNumber="453" value="1961-06-18">18.vi.1961</collectingDate>
</date>
, A.
<collectorName id="CB6063EDFF9CFFADBF4AA0E63FDBE3F4" box="[372,439,1944,1970]" pageId="2" pageNumber="453">Maier</collectorName>
(
<collectionCode id="00849EFEFF9CFFADBFF8A0E63C6EE3F4" box="[454,514,1944,1970]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="2" pageNumber="453" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF9CFFADBC2BA0E63FDEE390" collectingDate="1959-09-15" collectionCode="AMNH" collectorName="V. D. Roth &amp; W. J. Gertsch" country="United States of America" location="Double Bunk Meadows" pageId="2" pageNumber="453" specimenCount="22" specimenCount-female="9" specimenCount-male="13" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF9CFFADBC2BA0E63C29E3F4" box="[533,581,1944,1970]" pageId="2" pageNumber="453" type="male">13♂</specimenCount>
<specimenCount id="7093CDB2FF9CFFADBC72A0E63C1CE3F4" box="[588,624,1944,1970]" pageId="2" pageNumber="453" type="female">9♀</specimenCount>
,
<location id="634A50E0FF9CFFADBC42A0E63DE4E3F4" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9CFFADBC42A0E63DE4E3F4" box="[636,904,1944,1970]" country="United States of America" name="Double Bunk Meadows" pageId="2" pageNumber="453" stateProvince="California">Double Bunk Meadows</location>
,
<locationDeviation id="77981F19FF9CFFADBDADA0E63AF1E3F4" box="[915,1181,1944,1971]" location="Johnsondale" pageId="2" pageNumber="453">
<quantity id="A16DABDEFF9CFFADBDADA0E63DA0E3F4" box="[915,972,1944,1970]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="2" pageNumber="453" unit="mi" value="6.0">6 mi.</quantity>
W of Johnsondale
</locationDeviation>
,
<date id="122B20FBFF9CFFADBA96A0E63B4EE3F5" box="[1192,1314,1944,1971]" pageId="2" pageNumber="453" value="1959-09-15">
<collectingDate id="026FD913FF9CFFADBA96A0E63B4EE3F5" box="[1192,1314,1944,1971]" pageId="2" pageNumber="453" value="1959-09-15">15.ix.1959</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9CFFADBB12A0E73BF0E3F4" box="[1324,1436,1944,1971]" pageId="2" pageNumber="453">V.D. Roth</collectorName>
&amp;
<collectorName id="CB6063EDFF9CFFADBE8DA0C33F2CE390" box="[179,320,1980,2007]" pageId="2" pageNumber="453">W.J. Gertsch</collectorName>
(
<collectionCode id="00849EFEFF9CFFADBF70A0C23FC6E390" box="[334,426,1980,2006]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="453">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF9CFFADBF83A0C23AB4E390" box="[445,1240,1980,2007]" collectingDate="1956-07-04" collectionCode="CAS" collectorName="Kern R. &amp; V. D. Roth &amp; W. J. Gertsch" country="United States of America" location="Johnsondale" pageId="2" pageNumber="453" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF9CFFADBF83A0C23FB3E390" box="[445,479,1980,2006]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
,
<location id="634A50E0FF9CFFADBFD3A0C23C16E390" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9CFFADBFD3A0C23C16E390" box="[493,634,1980,2006]" country="United States of America" name="Johnsondale" pageId="2" pageNumber="453" stateProvince="California">Johnsondale</location>
,
<collectorName id="CB6063EDFF9CFFADBCBAA0C33CB2E391" box="[644,734,1981,2007]" pageId="2" pageNumber="453">Kern R.</collectorName>
,
<date id="122B20FBFF9CFFADBCD7A0C23D31E391" box="[745,861,1980,2007]" pageId="2" pageNumber="453" value="1956-07-04">
<collectingDate id="026FD913FF9CFFADBCD7A0C23D31E391" box="[745,861,1980,2007]" pageId="2" pageNumber="453" value="1956-07-04">4.vii.1956</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9CFFADBD59A0C33DBAE390" box="[871,982,1980,2007]" pageId="2" pageNumber="453">V.D. Roth</collectorName>
&amp;
<collectorName id="CB6063EDFF9CFFADBDC6A0C33AEAE390" box="[1016,1158,1980,2007]" pageId="2" pageNumber="453">W.J. Gertsch</collectorName>
(
<collectionCode id="00849EFEFF9CFFADBAAAA0C23ABCE390" box="[1172,1232,1980,2006]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="2" pageNumber="453" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF9CFFADBADDA0C23D36E3BC" collectingDate="1959-09-09" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; V. D. Roth" country="United States of America" location="Camp" pageId="2" pageNumber="453" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF9CFFADBADDA0C23B6AE390" box="[1251,1286,1980,2006]" pageId="2" pageNumber="453" type="male">3♂</specimenCount>
<specimenCount id="7093CDB2FF9CFFADBB32A0C23B5CE390" box="[1292,1328,1980,2006]" pageId="2" pageNumber="453" type="female">3♀</specimenCount>
, Quaking Aspen
<location id="634A50E0FF9CFFADBED7A09E3F43E3BC" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9CFFADBED7A09E3F43E3BC" box="[233,303,2016,2042]" country="United States of America" name="Camp" pageId="2" pageNumber="453" stateProvince="California">Camp</location>
,
<date id="122B20FBFF9CFFADBF02A09E3FC5E3BD" box="[316,425,2016,2043]" pageId="2" pageNumber="453" value="1959-09-09">
<collectingDate id="026FD913FF9CFFADBF02A09E3FC5E3BD" box="[316,425,2016,2043]" pageId="2" pageNumber="453" value="1959-09-09">9.ix.1959</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9CFFADBF88A09F3C24E3BC" box="[438,584,2016,2043]" pageId="2" pageNumber="453">W.J. Gertsch</collectorName>
&amp;
<collectorName id="CB6063EDFF9CFFADBC4FA09F3C88E3BC" box="[625,740,2016,2043]" pageId="2" pageNumber="453">V.D. Roth</collectorName>
(
<collectionCode id="00849EFEFF9CFFADBCC8A09E3D3EE3BC" box="[758,850,2016,2042]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="453">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D6FD0C66FF9CFFADBD56A09E3BF0E3BC" box="[872,1436,2016,2042]" country="United States of America" location="Quaking Aspen Meadow" pageId="2" pageNumber="453" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="7093CDB2FF9CFFADBD56A09E3DE7E3BC" box="[872,907,2016,2042]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
,
<location id="634A50E0FF9CFFADBDA2A09E3AD5E3BC" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9CFFADBDA2A09E3AD5E3BC" box="[924,1209,2016,2042]" country="United States of America" name="Quaking Aspen Meadow" pageId="2" pageNumber="453" stateProvince="California">Quaking Aspen Meadow</location>
,
<locationDeviation id="77981F19FF9CFFADBAF9A09E3BF0E3BC" box="[1223,1436,2016,2042]" location="Camp" pageId="2" pageNumber="453">
<quantity id="A16DABDEFF9CFFADBAF9A09E3B68E3BC" box="[1223,1284,2016,2042]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="2" pageNumber="453" unit="mi" value="3.0">3 mi.</quantity>
SE of Camp
</locationDeviation>
</materialsCitation>
</paragraph>
<paragraph id="662A063BFF9DFFACBEA9A7E93CD0E493" blockId="3.[151,1436,151,249]" pageId="3" pageNumber="454">
<materialsCitation id="D6FD0C66FF9DFFACBEA9A7E93CCBE4F7" box="[151,679,151,177]" collectingDate="1988-07-03" collectionCode="CAS" collectorName="D. Ubick" location="Nelson" pageId="3" pageNumber="454" specimenCount="1" specimenCount-female="1">
<location id="634A50E0FF9DFFACBEA9A7E93E85E4F7" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9DFFACBEA9A7E93E85E4F7" box="[151,233,151,177]" name="Nelson" pageId="3" pageNumber="454">Nelson</location>
, 6900’,
<date id="122B20FBFF9DFFACBF7CA7E93FDAE4F7" box="[322,438,151,177]" pageId="3" pageNumber="454" value="1988-07-03">
<collectingDate id="026FD913FF9DFFACBF7CA7E93FDAE4F7" box="[322,438,151,177]" pageId="3" pageNumber="454" value="1988-07-03">3.vii.1988</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9DFFACBFFFA7E93C4BE4F7" box="[449,551,151,177]" pageId="3" pageNumber="454">D. Ubick</collectorName>
(
<collectionCode id="00849EFEFF9DFFACBC08A7E93C1EE4F7" box="[566,626,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="454" type="Museum">CAS</collectionCode>
);
<specimenCount id="7093CDB2FF9DFFACBCBAA7E93CCBE4F7" box="[644,679,151,177]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="D6FD0C66FF9DFFACBC8BA7E93CD4E493" collectingDate="1968-09-15" collectionCode="CAS" collectorName="T. Briggs" latitude="36.4" location="Sequoia National Park" longLatPrecision="1188" longitude="-118.583336" municipality="Cirque Cave" pageId="3" pageNumber="454" specimenCount="1">
<location id="634A50E0FF9DFFACBC8BA7E93DDEE4F7" LSID="urn:lsid:plazi:treatment:EE3CB72DFF9CFFABBEA9A19D3D90E073:634A50E0FF9DFFACBC8BA7E93DDEE4F7" box="[693,946,151,177]" latitude="36.4" longLatPrecision="1188" longitude="-118.583336" municipality="Cirque Cave" name="Sequoia National Park" pageId="3" pageNumber="454">Sequoia National Park</location>
, near
<collectingMunicipality id="864E9C41FF9DFFACBDCDA7E93AEDE4F7" box="[1011,1153,151,177]" pageId="3" pageNumber="454">Cirque Cave</collectingMunicipality>
,
<locationDeviation id="77981F19FF9DFFACBAB2A7E93BF4E4F7" box="[1164,1432,151,177]" location="Mineral King" pageId="3" pageNumber="454">
<quantity id="A16DABDEFF9DFFACBAB2A7E93AA9E4F7" box="[1164,1221,151,177]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="3" pageNumber="454" unit="mi" value="3.0">3 mi.</quantity>
S of Mineral King
</locationDeviation>
,
<geoCoordinate id="03A160FCFF9DFFACBEA9A7C23E98E491" box="[151,244,188,215]" degrees="36" direction="north" minutes="24" orientation="latitude" pageId="3" pageNumber="454" precision="925" value="36.4">36°24′N</geoCoordinate>
<geoCoordinate id="03A160FCFF9DFFACBEC2A7C23F01E491" box="[252,365,188,215]" degrees="118" direction="west" minutes="35" orientation="longitude" pageId="3" pageNumber="454" precision="925" value="-118.583336">118°35′W</geoCoordinate>
,
<date id="122B20FBFF9DFFACBF46A7C43F9EE493" box="[376,498,186,213]" pageId="3" pageNumber="454" value="1968-09-15">
<collectingDate id="026FD913FF9DFFACBF46A7C43F9EE493" box="[376,498,186,213]" pageId="3" pageNumber="454" value="1968-09-15">15.ix.1968</collectingDate>
</date>
,
<collectorName id="CB6063EDFF9DFFACBFC3A7C53C09E493" box="[509,613,187,213]" pageId="3" pageNumber="454">T. Briggs</collectorName>
(
<collectionCode id="00849EFEFF9DFFACBC4AA7C53CDCE493" box="[628,688,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="454" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9DFFACBEF9A7A13B46E4BF" box="[199,1322,223,249]" pageId="3" pageNumber="454" type="etymology">
<paragraph id="662A063BFF9DFFACBEF9A7A13B46E4BF" blockId="3.[151,1436,151,249]" box="[199,1322,223,249]" pageId="3" pageNumber="454">
<emphasis id="54E1DA29FF9DFFACBEF9A7A13F22E4BF" bold="true" box="[199,334,223,249]" pageId="3" pageNumber="454">Etymology.</emphasis>
The specific epithet, derived from the Latin for “double bed”, refers to the
<typeStatus id="B92EB899FF9DFFACBAA4A7A13AA6E4BF" box="[1178,1226,223,249]" pageId="3" pageNumber="454">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="32EA56B3FF9DFFACBEA9A02C3F41E390" ID-DOI="http://doi.org/10.5281/zenodo.5300632" ID-Zenodo-Dep="5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="3" pageNumber="454" startId="3.[151,264,1874,1898]" targetBox="[151,1377,270,1847]" targetPageId="3">
<paragraph id="662A063BFF9DFFACBEA9A02C3F41E390" blockId="3.[151,1436,1874,2007]" pageId="3" pageNumber="454">
<emphasis id="54E1DA29FF9DFFACBEA9A02C3F57E32C" bold="true" box="[151,315,1874,1898]" pageId="3" pageNumber="454">FIGURES 1–4.</emphasis>
<taxonomicName id="A1957DB8FF9DFFACBF7CA02C3F9CE32C" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[322,496,1874,1898]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9DFFACBF7CA02C3F9CE32C" box="[322,496,1874,1898]" italics="true" pageId="3" pageNumber="454">Cybaeus bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9DFFACBFC9A02D3C33E32D" bold="true" box="[503,607,1875,1899]" pageId="3" pageNumber="454">
<taxonomicNameLabel id="4FD26752FF9DFFACBFC9A02D3C33E32D" box="[503,607,1875,1899]" pageId="3" pageNumber="454" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum.
</paragraph>
</caption>
<subSubSection id="2E8F55B0FF9AFFABBEF9A7E83AF6E6EE" pageId="4" pageNumber="455" type="diagnosis">
<paragraph id="662A063BFF9AFFABBEF9A7E83A8AE523" blockId="4.[151,1437,150,1077]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A7E83F2EE4F6" bold="true" box="[199,322,150,176]" pageId="4" pageNumber="455">Diagnosis.</emphasis>
The male of
<taxonomicName id="A1957DB8FF9AFFABBFEBA7E93C21E4F7" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[469,589,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBFEBA7E93C21E4F7" box="[469,589,151,177]" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBC6DA7E93CAAE4F7" bold="true" box="[595,710,151,177]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBC6DA7E93CAAE4F7" box="[595,710,151,177]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is unlikely to be confused with the males of
<taxonomicName id="A1957DB8FF9AFFABBA84A7E93B37E4F7" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1210,1371,151,177]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBA84A7E93B37E4F7" box="[1210,1371,151,177]" italics="true" pageId="4" pageNumber="455">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBB5CA7E93EA4E493" bold="true" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBB5CA7E93EA4E493" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF9AFFABBEC1A7C53F13E493" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[255,383,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBEC1A7C53F13E493" box="[255,383,187,213]" italics="true" pageId="4" pageNumber="455">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBFB8A7C53F95E493" bold="true" box="[390,505,187,213]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBFB8A7C53F95E493" box="[390,505,187,213]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, the other members of the
<taxonomicName id="A1957DB8FF9AFFABBD16A7C53D1CE493" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[808,880,187,213]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9AFFABBD16A7C53D1CE493" box="[808,880,187,213]" italics="true" pageId="4" pageNumber="455">devius</emphasis>
</taxonomicName>
species group with known males. The male of
<taxonomicName id="A1957DB8FF9AFFABBBBDA7C53E82E4BF" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus">
<emphasis id="54E1DA29FF9AFFABBBBDA7C53E82E4BF" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
is diagnosed by the morphology of the patellar apophysis (
<figureCitation id="FEAE1ABEFF9AFFABBD45A7A13DB3E4BF" box="[891,991,223,249]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="4" pageNumber="455">Figs 1–2</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBDD4A7A13A6BE4BC" box="[1002,1031,223,250]" captionStart="FIGURES 26–28" captionStartId="11.[151,264,1216,1240]" captionTargetBox="[151,1436,705,1193]" captionTargetId="figure-226@11.[151,1436,705,1193]" captionTargetPageId="11" captionText="FIGURES 26–28. Cybaeus spp. males, left palpus, patellar apophysis, dorsal (26 C. bilectus spec. nov., from Double Bunk Meadows, California, 27 C. bryoncavus spec. nov., holotype, 28 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (26 concave distal margin; 28 concave dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.5300646" httpUri="https://zenodo.org/record/5300646/files/figure.png" pageId="4" pageNumber="455">26</figureCitation>
) and the proximal arm of the tegular apophysis (
<figureCitation id="FEAE1ABEFF9AFFABBF28A67D3F17E55B" box="[278,379,259,285]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="4" pageNumber="455">Figs 3–4</figureCitation>
,
<figureCitation id="FEAE1ABEFF9AFFABBFB9A67D3FCBE55B" box="[391,423,259,285]" captionStart="FIGURES 29–31" captionStartId="11.[151,264,1858,1882]" captionTargetBox="[151,1436,1337,1834]" captionTargetId="figure-280@11.[151,1436,1337,1834]" captionTargetPageId="11" captionText="FIGURES 29–31. Cybaeus spp. males, left palpus, proximal arm of tegular apophysis, ventral (29 C. bilectus spec. nov., from Double Bunk Meadows, California, 30 C. bryoncavus spec. nov., holotype, 31 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (29 bluntly hooked tip, 30 inconspicuous basal projection)." figureDoi="http://doi.org/10.5281/zenodo.5300649" httpUri="https://zenodo.org/record/5300649/files/figure.png" pageId="4" pageNumber="455">29</figureCitation>
): the patellar apophysis is short (length about 1/2 the width of the patella) and prolaterally directed with a slightly concave distal margin and approximately 20 to 30 peg setae dorsally along the length of the distal margin; the proximal arm of the tegular apophysis is simple with a short, bluntly hooked tip.
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A6113AF6E6EE" blockId="4.[151,1437,150,1077]" pageId="4" pageNumber="455">
The female of
<taxonomicName id="A1957DB8FF9AFFABBF50A6113F8AE5CF" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[366,486,367,393]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBF50A6113F8AE5CF" box="[366,486,367,393]" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBFD0A6113C0DE5CF" bold="true" box="[494,609,367,393]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBFD0A6113C0DE5CF" box="[494,609,367,393]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is diagnosed by the form and configuration of the copulatory ducts (
<figureCitation id="FEAE1ABEFF9AFFABBB54A6113EBDE5EB" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">Figs 9–10</figureCitation>
) and is only likely to be confused with the female of
<taxonomicName id="A1957DB8FF9AFFABBD74A6ED3D82E5EB" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[842,1006,403,429]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBD74A6ED3D82E5EB" box="[842,1006,403,429]" italics="true" pageId="4" pageNumber="455">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBDC6A6ED3A02E5EB" bold="true" box="[1016,1134,403,429]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBDC6A6ED3A02E5EB" box="[1016,1134,403,429]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, the only other species of the
<taxonomicName id="A1957DB8FF9AFFABBEFCA6C93F66E597" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[194,266,439,465]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9AFFABBEFCA6C93F66E597" box="[194,266,439,465]" italics="true" pageId="4" pageNumber="455">devius</emphasis>
</taxonomicName>
group with elongate convoluted and undifferentiated spermathecal stalks and copulatory ducts. (
<emphasis id="54E1DA29FF9AFFABBB65A6C93BF0E597" bold="true" box="[1371,1436,439,465]" pageId="4" pageNumber="455">Note:</emphasis>
because of their relatively light sclerotization and similar morphology, it is difficult to trace accurately the pathways of the spermathecal stalks and copulatory ducts in the females of
<taxonomicName id="A1957DB8FF9AFFABBDB8A6813D93E65F" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[902,1023,511,537]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBDB8A6813D93E65F" box="[902,1023,511,537]" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBA36A6813A10E65F" bold="true" box="[1032,1148,511,537]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBA36A6813A10E65F" box="[1032,1148,511,537]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF9AFFABBA88A6813B35E65F" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1206,1369,511,537]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBA88A6813B35E65F" box="[1206,1369,511,537]" italics="true" pageId="4" pageNumber="455">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBB5CA6813EBCE67B" bold="true" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBB5CA6813EA6E67B" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
,
</emphasis>
even in well cleared preparations of the vulva.) In
<taxonomicName id="A1957DB8FF9AFFABBD31A55D3DE4E67B" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[783,904,547,573]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBD31A55D3DE4E67B" box="[783,904,547,573]" italics="true" pageId="4" pageNumber="455">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBDB1A55D3A6EE67B" bold="true" box="[911,1026,547,573]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBDB1A55D3A6EE67B" box="[911,1026,547,573]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
the basal section of each copulatory duct travels antero-laterally from the atrium and traverses the ventral surface of the medial section of the copulatory duct (
<emphasis id="54E1DA29FF9AFFABBEE7A5153F4DE6C2" box="[217,289,619,644]" italics="true" pageId="4" pageNumber="455">versus</emphasis>
basal section travels anteriorly from the atrium along the midline of the vulva prior to diverging and intertwining with loops of the spermathecal stalk [
<figureCitation id="FEAE1ABEFF9AFFABBCF6A5F13D26E6EF" box="[712,842,654,681]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">Figs 12–13</figureCitation>
] in
<taxonomicName id="A1957DB8FF9AFFABBD4BA5F13A7BE6EF" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[885,1047,655,681]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9AFFABBD4BA5F13A7BE6EF" box="[885,1047,655,681]" italics="true" pageId="4" pageNumber="455">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9AFFABBA20A5F13AF6E6EE" bold="true" box="[1054,1178,654,681]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="4FD26752FF9AFFABBA20A5F13AFEE6EE" box="[1054,1170,654,681]" pageId="4" pageNumber="455" rank="species">spec. nov.</taxonomicNameLabel>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9AFFABBEF9A5CC3FF6E7AB" pageId="4" pageNumber="455" type="description">
<paragraph id="662A063BFF9AFFABBEF9A5CC3D04E68B" blockId="4.[151,1437,150,1077]" box="[199,872,690,717]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A5CC3F35E68A" bold="true" box="[199,345,690,716]" pageId="4" pageNumber="455">Description.</emphasis>
Ventral tibia I macrosetae: 2-1p-2-1p-2(or 1p).
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A5A83D7BE753" blockId="4.[151,1437,150,1077]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A5A83F67E6B7" bold="true" box="[199,267,726,753]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A5A83F6EE6B7" bold="true" box="[199,258,726,753]" italics="true" pageId="4" pageNumber="455">Male</emphasis>
:
</emphasis>
(n=17). Distal arm of tegular apophysis (
<figureCitation id="FEAE1ABEFF9AFFABBCE3A5A93D73E6B7" box="[733,799,727,753]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="4" pageNumber="455">Fig. 3</figureCitation>
) prominent, broad basally, elongate (but length less than twice width), abruptly tapering distally. Embolus narrow.
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A4613DC0E71B" blockId="4.[151,1437,150,1077]" pageId="4" pageNumber="455">
Measurements (n=17). CL 2.33–3.4 (2.8+0.3), CW 1.73–2.50 (2.06+0.20), SL 1.20–1.65 (1.37+0.11), SW 1.12–1.50 (1.29+0.09).
<typeStatus id="B92EB899FF9AFFABBF9FA43D3C65E71B" box="[417,521,835,861]" pageId="4" pageNumber="455" type="holotype">Holotype</typeStatus>
CL 2.9, CW 2.08, SL 1.43, SW 1.30.
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A4183C92E7E3" blockId="4.[151,1437,150,1077]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A4183F72E7C7" bold="true" box="[199,286,870,897]" italics="true" pageId="4" pageNumber="455">Female</emphasis>
: (n=16). Atrium (
<figureCitation id="FEAE1ABEFF9AFFABBFD9A4193C27E7C7" box="[487,587,871,897]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">Figs 8–9</figureCitation>
) shallow, small, about 1/2 width of vulva. Spermathecal stalks looped once around medial section of copulatory ducts (
<figureCitation id="FEAE1ABEFF9AFFABBC42A4F53C9DE7E3" box="[636,753,907,933]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="4" pageNumber="455">Figs 9–10</figureCitation>
).
</paragraph>
<paragraph id="662A063BFF9AFFABBEF9A4D13FF6E7AB" blockId="4.[151,1437,150,1077]" pageId="4" pageNumber="455">Measurements (n=14). CL 2.18–3.3 (2.8+0.3), CW 1.50–2.28 (1.95+0.24), SL 1.14–1.58 (1.38+0.13), SW 1.04–1.48 (1.29+0.12).</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9AFFABBEF9A4883D90E073" pageId="4" pageNumber="455" type="distribution">
<paragraph id="662A063BFF9AFFABBEF9A4883D90E073" blockId="4.[151,1437,150,1077]" pageId="4" pageNumber="455">
<emphasis id="54E1DA29FF9AFFABBEF9A4883C3CE056" bold="true" box="[199,592,1014,1041]" pageId="4" pageNumber="455">Distribution and natural history.</emphasis>
(
<figureCitation id="FEAE1ABEFF9AFFABBC5EA4893CDAE057" box="[608,694,1015,1041]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="4" pageNumber="455">Fig. 32</figureCitation>
). Known only from a small area of
<taxonomicName id="A1957DB8FF9AFFABBA74A4893A80E057" authority="County" authorityName="County" box="[1098,1260,1015,1041]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="tulare">Tulare County</taxonomicName>
in the southern Sierra
<collectingRegion id="A451C8D9FF9AFFABBEDFA3653F5BE073" box="[225,311,1051,1077]" country="United States of America" name="Nevada" pageId="4" pageNumber="455">Nevada</collectingRegion>
of eastern
<collectingRegion id="A451C8D9FF9AFFABBF8CA3653C4BE073" box="[434,551,1051,1077]" country="United States of America" name="California" pageId="4" pageNumber="455">California</collectingRegion>
. Males have been collected in September.
</paragraph>
</subSubSection>
</treatment>
</document>

248
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<paragraph id="662A063BFF9FFFAEBEA9A2973FA7E245" blockId="1.[151,459,1513,1540]" box="[151,459,1513,1540]" pageId="1" pageNumber="452">
<heading id="3D62B157FF9FFFAEBEA9A2973FA7E245" bold="true" box="[151,459,1513,1540]" fontSize="11" level="1" pageId="1" pageNumber="452" reason="1">
<emphasis id="54E1DA29FF9FFFAEBEA9A2973FA7E245" bold="true" box="[151,459,1513,1540]" pageId="1" pageNumber="452">
The
<taxonomicName id="A1957DB8FF9FFFAEBEF1A2973F70E242" ID-CoL="6BSTR" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[207,284,1513,1540]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9FFFAEBEF1A2973F70E242" bold="true" box="[207,284,1513,1540]" italics="true" pageId="1" pageNumber="452">devius</emphasis>
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species group
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9FFFAEBEA9A14F3AFFE301" pageId="1" pageNumber="452" type="diagnosis">
<paragraph id="662A063BFF9FFFAEBEA9A14F3AFFE301" blockId="1.[151,1437,1585,2008]" pageId="1" pageNumber="452">
<emphasis id="54E1DA29FF9FFFAEBEA9A14F3F66E20D" bold="true" box="[151,266,1585,1611]" pageId="1" pageNumber="452">Diagnosis</emphasis>
. The
<taxonomicName id="A1957DB8FF9FFFAEBF73A14F3FF9E20D" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[333,405,1585,1611]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9FFFAEBF73A14F3FF9E20D" box="[333,405,1585,1611]" italics="true" pageId="1" pageNumber="452">devius</emphasis>
</taxonomicName>
group is diagnosed through the females by the long, convoluted copulatory ducts with the basal sections at least partially enveloping or looped about the more apical sections (
<figureCitation id="FEAE1ABEFF9FFFAEBA4DA12B3B2CE229" box="[1139,1344,1621,1647]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="1" pageNumber="452">Figs 9–10, 12–13</figureCitation>
,
<figureCitation id="FEAE1ABEFF9FFFAEBB71A12B3EB3E2D5" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="1" pageNumber="452">15–16, 18–19</figureCitation>
,
<figureCitation id="FEAE1ABEFF9FFFAEBED0A1073F54E2D5" box="[238,312,1657,1683]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="1" pageNumber="452">21–22</figureCitation>
). Within this group two subgroups are definable: one comprised of
<taxonomicName id="A1957DB8FF9FFFAEBA7AA1043AC3E2D5" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[1092,1199,1657,1684]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9FFFAEBA7AA1043AC3E2D5" box="[1092,1199,1657,1684]" italics="true" pageId="1" pageNumber="452">C. devius</emphasis>
</taxonomicName>
,
<taxonomicName id="A1957DB8FF9FFFAEBA81A1043B7BE2D5" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1215,1303,1657,1684]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF9FFFAEBA81A1043B7BE2D5" box="[1215,1303,1657,1684]" italics="true" pageId="1" pageNumber="452">C. echo</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9FFFAEBB1FA1073BF9E2D5" bold="true" box="[1313,1429,1657,1683]" pageId="1" pageNumber="452">
<taxonomicNameLabel id="4FD26752FF9FFFAEBB1FA1073BF9E2D5" box="[1313,1429,1657,1683]" pageId="1" pageNumber="452" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="A1957DB8FF9FFFAEBEF9A1E03F2AE2F1" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[199,326,1693,1720]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF9FFFAEBEF9A1E03F2AE2F1" box="[199,326,1693,1720]" italics="true" pageId="1" pageNumber="452">C. wilsonia</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9FFFAEBF73A1E33FACE2F1" bold="true" box="[333,448,1693,1719]" pageId="1" pageNumber="452">
<taxonomicNameLabel id="4FD26752FF9FFFAEBF73A1E33FACE2F1" box="[333,448,1693,1719]" pageId="1" pageNumber="452" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, and the other including
<taxonomicName id="A1957DB8FF9FFFAEBCE6A1E03D3CE2F1" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[728,848,1693,1720]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9FFFAEBCE6A1E03D3CE2F1" box="[728,848,1693,1720]" italics="true" pageId="1" pageNumber="452">C. bilectus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9FFFAEBD69A1E33DA6E2F1" bold="true" box="[855,970,1693,1719]" pageId="1" pageNumber="452">
<taxonomicNameLabel id="4FD26752FF9FFFAEBD69A1E33DA6E2F1" box="[855,970,1693,1719]" pageId="1" pageNumber="452" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="A1957DB8FF9FFFAEBA3EA1E03ACEE2F1" authority="Bennett &amp; Copley &amp; Copley, 2021" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1024,1186,1693,1720]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9FFFAEBA3EA1E03ACEE2F1" box="[1024,1186,1693,1720]" italics="true" pageId="1" pageNumber="452">C. bryoncavus</emphasis>
</taxonomicName>
<emphasis id="54E1DA29FF9FFFAEBA97A1E33B70E2F1" bold="true" box="[1193,1308,1693,1719]" pageId="1" pageNumber="452">
<taxonomicNameLabel id="4FD26752FF9FFFAEBA97A1E33B70E2F1" box="[1193,1308,1693,1719]" pageId="1" pageNumber="452" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
The
<taxonomicName id="A1957DB8FF9FFFAEBB6BA1E33BF1E2F1" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[1365,1437,1693,1719]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9FFFAEBB6BA1E33BF1E2F1" box="[1365,1437,1693,1719]" italics="true" pageId="1" pageNumber="452">devius</emphasis>
</taxonomicName>
subgroup is characterized by distinctly bipartite copulatory ducts (a basal, very lightly sclerotized region enveloping an apical, relatively straight, heavily sclerotized section) and very short stalks (
<figureCitation id="FEAE1ABEFF9FFFAEBA17A19B3B6FE2B9" box="[1065,1283,1765,1792]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="1" pageNumber="452">Figs 15–16, 18–19</figureCitation>
,
<figureCitation id="FEAE1ABEFF9FFFAEBB2FA19B3B37E2B9" box="[1297,1371,1765,1791]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="1" pageNumber="452">21–22</figureCitation>
). The
<taxonomicName id="A1957DB8FF9FFFAEBEA9A0773E82E365" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[151,238,1801,1827]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="bilectus">
<emphasis id="54E1DA29FF9FFFAEBEA9A0773E82E365" box="[151,238,1801,1827]" italics="true" pageId="1" pageNumber="452">bilectus</emphasis>
</taxonomicName>
subgroup has the copulatory ducts gradually becoming more heavily sclerotized apically and longer stalks which take a single turn around the medial part of the copulatory ducts (
<figureCitation id="FEAE1ABEFF9FFFAEBD80A0533AEBE301" box="[958,1159,1837,1863]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="1" pageNumber="452">Figs 9–10, 12–13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9FFFADBEF9A02F3D51E6EF" lastPageId="2" lastPageNumber="453" pageId="1" pageNumber="452" type="description">
<paragraph id="662A063BFF9FFFAEBEF9A02F3AE9E3F2" blockId="1.[151,1437,1585,2008]" pageId="1" pageNumber="452">
<emphasis id="54E1DA29FF9FFFAEBEF9A02F3F3DE32D" bold="true" box="[199,337,1873,1899]" pageId="1" pageNumber="452">Description</emphasis>
. Medium-sized spiders: carapace lengths ranging 2.0–
<quantity id="A16DABDEFF9FFFAEBD92A02F3D93E32A" box="[940,1023,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="452" unit="mm" value="2.8">2.8 mm</quantity>
(females). Males subequal to slightly larger. Abdomen patterned. Legs usually unbanded (femora occasionally lightly banded). Two or occasionally three complete pairs of ventral tibia I macrosetae (distal pair sometimes present or incomplete).
</paragraph>
<paragraph id="662A063BFF9FFFADBEF9A0C33C86E507" blockId="1.[151,1437,1585,2008]" lastBlockId="2.[151,1437,150,681]" lastPageId="2" lastPageNumber="453" pageId="1" pageNumber="452">
<emphasis id="54E1DA29FF9FFFAEBEF9A0C33F6EE39E" bold="true" box="[199,258,1981,2008]" italics="true" pageId="1" pageNumber="452">Male</emphasis>
: Patellar apophysis (
<figureCitation id="FEAE1ABEFF9FFFAEBFD6A0C33C27E391" box="[488,587,1981,2007]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="1" pageNumber="452">Figs 1–2</figureCitation>
,
<figureCitation id="FEAE1ABEFF9FFFAEBC68A0C33C0AE39E" box="[598,614,1981,2008]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="1" pageNumber="452">5</figureCitation>
,
<figureCitation id="FEAE1ABEFF9FFFAEBC4EA0C33CE2E391" box="[624,654,1981,2007]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="1" pageNumber="452">23</figureCitation>
,
<figureCitation id="FEAE1ABEFF9FFFAEBCA7A0C33C8DE39E" box="[665,737,1981,2008]" captionStart="FIGURES 26–28" captionStartId="11.[151,264,1216,1240]" captionTargetBox="[151,1436,705,1193]" captionTargetId="figure-226@11.[151,1436,705,1193]" captionTargetPageId="11" captionText="FIGURES 26–28. Cybaeus spp. males, left palpus, patellar apophysis, dorsal (26 C. bilectus spec. nov., from Double Bunk Meadows, California, 27 C. bryoncavus spec. nov., holotype, 28 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (26 concave distal margin; 28 concave dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.5300646" httpUri="https://zenodo.org/record/5300646/files/figure.png" pageId="1" pageNumber="452">26–28</figureCitation>
) laterally to disto-laterally directed, length 1/2 to nearly as long as width of tibia, 12–30 peg setae, number is species specific. Retrolateral tibial apophysis (
<figureCitation id="FEAE1ABEFF9CFFADBA94A7E93B7CE4F7" box="[1194,1296,151,177]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="2" pageNumber="453">Figs 1–2</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBB23A7E83B47E4F7" box="[1309,1323,150,177]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="2" pageNumber="453">5</figureCitation>
) carinate, nearly as long as tibia. Embolus long and narrow (
<figureCitation id="FEAE1ABEFF9CFFADBCF9A7C53D62E493" box="[711,782,187,213]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="2" pageNumber="453">Figs 3</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBD25A7C53D2BE490" box="[795,839,187,214]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="2" pageNumber="453">6–7</figureCitation>
) or long and slightly ribbon-like and broadened (
<figureCitation id="FEAE1ABEFF9CFFADBB54A7C53E8EE4BF" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="2" pageNumber="453">Figs 24–25</figureCitation>
). Tegulum (
<figureCitation id="FEAE1ABEFF9CFFADBF58A7A13FC2E4BF" box="[358,430,223,249]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="2" pageNumber="453">Figs 3</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBF82A7A13FB5E4BF" box="[444,473,223,249]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="2" pageNumber="453">24</figureCitation>
) conspicuous in ventral view, partially obscuring basal part of embolus. Distal arm of tegular apophysis prominent, elongate and either narrow (
<figureCitation id="FEAE1ABEFF9CFFADBD23A67D3D02E55B" box="[797,878,259,285]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="2" pageNumber="453">Fig. 24</figureCitation>
) or broad (
<figureCitation id="FEAE1ABEFF9CFFADBDD7A67D3A5DE55B" box="[1001,1073,259,285]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="2" pageNumber="453">Figs 3</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBA00A67D3A22E558" box="[1086,1102,259,286]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="2" pageNumber="453">6</figureCitation>
); proximal arm (
<figureCitation id="FEAE1ABEFF9CFFADBB37A67D3B3DE55B" box="[1289,1361,259,285]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1874,1898]" captionTargetBox="[151,1377,270,1847]" captionTargetId="figure-59@3.[151,1436,270,1848]" captionTargetPageId="3" captionText="FIGURES 1–4. Cybaeus bilectus spec. nov., male from Double Bunk Meadows, California, left palpus. 1–2 Patella and tibia (1 retrolateral, 2 dorsal). 3–4 Genital bulb (3 ventral, 4 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E— embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis, TG—tegulum." figureDoi="http://doi.org/10.5281/zenodo.5300632" httpUri="https://zenodo.org/record/5300632/files/figure.png" pageId="2" pageNumber="453">Figs 3</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBB63A67D3B01E55B" box="[1373,1389,259,285]" captionStart="FIGURES 5–7" captionStartId="5.[151,264,1901,1925]" captionTargetBox="[215,1372,181,1842]" captionTargetId="figure-17@5.[185,1402,181,1877]" captionTargetPageId="5" captionText="FIGURES 5–7. Cybaeus bryoncavus spec. nov., holotype male, left palpus. 5 Patella and tibia, dorsal. 6 Genital bulb (embolus tip apparently missing; base of embolus, part of tegulum obscured in specimen, ventral. 7 Tegular apophysis and embolus (part), retrolateral. CY—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrow indicates basal projection of PA." figureDoi="http://doi.org/10.5281/zenodo.5300634" httpUri="https://zenodo.org/record/5300634/files/figure.png" pageId="2" pageNumber="453">6</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBB47A67C3BFBE55B" box="[1401,1431,258,285]" captionStart="FIGURES 23–25" captionStartId="10.[151,264,1891,1915]" captionTargetBox="[151,1424,183,1827]" captionTargetId="figure-31@10.[151,1436,181,1867]" captionTargetPageId="10" captionText="FIGURES 23–25. Cybaeus wilsonia spec. nov., holotype male, left palpus. 23 Patella and tibia, dorsal. 24–25 Genital bulb (24 ventral, 25 retrolateral). CY—cymbium, DA—distal arm of tegular apophysis, E—embolus. Unlabelled arrows indicate length and width of DA (Fig. 24) and swollen base of proximal arm of tegular apophysis (Fig. 25)." figureDoi="http://doi.org/10.5281/zenodo.5300644" httpUri="https://zenodo.org/record/5300644/files/figure.png" pageId="2" pageNumber="453">25</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBEA9A6593EB3E507" box="[151,223,295,321]" captionStart="FIGURES 29–31" captionStartId="11.[151,264,1858,1882]" captionTargetBox="[151,1436,1337,1834]" captionTargetId="figure-280@11.[151,1436,1337,1834]" captionTargetPageId="11" captionText="FIGURES 29–31. Cybaeus spp. males, left palpus, proximal arm of tegular apophysis, ventral (29 C. bilectus spec. nov., from Double Bunk Meadows, California, 30 C. bryoncavus spec. nov., holotype, 31 C. wilsonia spec. nov., holotype). Arrows indicate diagnostic characters (29 bluntly hooked tip, 30 inconspicuous basal projection)." figureDoi="http://doi.org/10.5281/zenodo.5300649" httpUri="https://zenodo.org/record/5300649/files/figure.png" pageId="2" pageNumber="453">29–31</figureCitation>
) simple, prolaterally directed, slightly twisted.
</paragraph>
<paragraph id="662A063BFF9CFFADBEF9A6343F0FE65F" blockId="2.[151,1437,150,681]" pageId="2" pageNumber="453">
<emphasis id="54E1DA29FF9CFFADBEF9A6343F72E523" bold="true" box="[199,286,330,357]" italics="true" pageId="2" pageNumber="453">Female</emphasis>
: Simple undivided transverse atrium (
<figureCitation id="FEAE1ABEFF9CFFADBCE5A6353D3FE523" box="[731,851,331,357]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="2" pageNumber="453">Figs 8, 11</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBD5CA6353DC1E523" box="[866,941,331,357]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="2" pageNumber="453">14, 17</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBD82A6353DB5E523" box="[956,985,331,357]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="2" pageNumber="453">20</figureCitation>
) medially located on epigynum. Vulva (
<figureCitation id="FEAE1ABEFF9CFFADBEA1A6113F08E5CF" box="[159,356,367,393]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[152,1436,190,1878]" captionTargetId="figure-31@6.[151,1436,181,1878]" captionTargetPageId="6" captionText="FIGURES 8–13. Cybaeus spp., females from California, copulatory organ (8–10 C. bilectus spec. nov., from Double Bunk Meadows; 11–13 C. bryoncavus spec. nov., from Pine Grove). 8, 11 Epigynum, ventral. 9–10, 12–13 Vulva (9, 12 ventral, 10, 13 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5300636" httpUri="https://zenodo.org/record/5300636/files/figure.png" pageId="2" pageNumber="453">Figs 9–10, 12–13</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBF51A6103C66E5CF" box="[367,522,366,393]" captionStart="FIGURES 14–19" captionStartId="8.[151,264,1750,1774]" captionTargetBox="[153,1434,181,1726]" captionTargetId="figure-31@8.[151,1436,181,1726]" captionTargetPageId="8" captionText="FIGURES 14–19. Cybaeus spp., females, copulatory organ (14–16 C. devius, holotype; 17–19 C. wilsonia spec. nov., from Wilsonia, California). 14, 17 Epigynum, ventral. 15–16, 18–19 Vulva (15, 18 ventral, 16, 19 dorsal). BG—Bennett’s gland, HS—head of spermatheca. Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300638" httpUri="https://zenodo.org/record/5300638/files/figure.png" pageId="2" pageNumber="453">15–16, 18–19</figureCitation>
,
<figureCitation id="FEAE1ABEFF9CFFADBC2AA6113C30E5CF" box="[532,604,367,393]" captionStart="FIGURES 20–22" captionStartId="9.[151,264,1438,1462]" captionTargetBox="[155,1431,349,1415]" captionTargetId="figure-85@9.[151,1436,345,1415]" captionTargetPageId="9" captionText="FIGURES 20–22. Cybaeus echo spec. nov., holotype female, copulatory organ. 20 Epigynum, ventral. 21–22 Vulva (21 ventral, 22 dorsal). Unlabelled solid arrows indicate basal lightly sclerotized sections of copulatory ducts which envelop apical more heavily sclerotized sections (indicated by unlabelled dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.5300642" httpUri="https://zenodo.org/record/5300642/files/figure.png" pageId="2" pageNumber="453">21–22</figureCitation>
) with basal sections of copulatory ducts completely or partially enveloping apical sections of the ducts or spermathecal stalks; spermathecal heads small, dorsolateral lobes; spermathecal stalks undifferentiated, relatively long or very short; lumina narrow throughout heads and stalks; Bennett’s glands large, prominent, near junction of stalks and spermathecal bases; bases large, rounded; fertilization ducts exit bases near entrance of stalks.
</paragraph>
<paragraph id="662A063BFF9CFFADBEF9A55C3D51E6EF" blockId="2.[151,1437,150,681]" pageId="2" pageNumber="453">
<emphasis id="54E1DA29FF9CFFADBEF9A55C3C47E67A" bold="true" box="[199,555,546,573]" pageId="2" pageNumber="453">Composition and distribution</emphasis>
. (
<figureCitation id="FEAE1ABEFF9CFFADBC7DA55D3CF4E67B" box="[579,664,547,573]" captionStart="FIGURE 32" captionStartId="12.[151,250,1918,1942]" captionTargetBox="[184,1406,198,1877]" captionTargetId="figure-31@12.[163,1423,181,1894]" captionTargetPageId="12" captionText="FIGURE 32. Distributions of Cybaeus bilectus spec. nov., C. bryoncavus spec. nov., C. devius, C. echo spec. nov., and C. wilsonia spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.5300651" httpUri="https://zenodo.org/record/5300651/files/figure.png" pageId="2" pageNumber="453">Fig. 32</figureCitation>
).
<taxonomicName id="A1957DB8FF9CFFADBC93A55D3DA0E67B" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[685,972,547,573]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="bilectus" status="sp. nov.">
<emphasis id="54E1DA29FF9CFFADBC93A55D3D01E67B" box="[685,877,547,573]" italics="true" pageId="2" pageNumber="453">Cybaeus bilectus</emphasis>
Bennett
</taxonomicName>
<emphasis id="54E1DA29FF9CFFADBDEAA55D3A24E67B" bold="true" box="[980,1096,547,573]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="4FD26752FF9CFFADBDEAA55D3A24E67B" box="[980,1096,547,573]" pageId="2" pageNumber="453" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="A1957DB8FF9CFFADBA69A55D3B36E67B" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1111,1370,547,573]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="bryoncavus" status="sp. nov.">
<emphasis id="54E1DA29FF9CFFADBA69A55D3A96E67B" box="[1111,1274,547,573]" italics="true" pageId="2" pageNumber="453">C. bryoncavus</emphasis>
Bennett
</taxonomicName>
<emphasis id="54E1DA29FF9CFFADBB5CA55D3EA4E627" bold="true" pageId="2" pageNumber="453">
<taxonomicNameLabel id="4FD26752FF9CFFADBB5CA55D3EA4E627" pageId="2" pageNumber="453" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="A1957DB8FF9CFFADBEE6A5393C4EE627" authority="Chamberlin &amp; Ivie" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[216,546,583,609]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="devius">
<emphasis id="54E1DA29FF9CFFADBEE6A5393F2EE627" box="[216,322,583,609]" italics="true" pageId="2" pageNumber="453">C. devius</emphasis>
Chamberlin &amp; Ivie
</taxonomicName>
,
<taxonomicName id="A1957DB8FF9CFFADBC11A5393C89E627" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[559,741,583,609]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="echo" status="sp. nov.">
<emphasis id="54E1DA29FF9CFFADBC11A5393CEAE627" box="[559,646,583,609]" italics="true" pageId="2" pageNumber="453">C. echo</emphasis>
Bennett
</taxonomicName>
<emphasis id="54E1DA29FF9CFFADBCD3A5393D0DE627" bold="true" box="[749,865,583,609]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="4FD26752FF9CFFADBCD3A5393D0DE627" box="[749,865,583,609]" pageId="2" pageNumber="453" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="A1957DB8FF9CFFADBD9FA5393AEDE627" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[929,1153,583,609]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="wilsonia" status="sp. nov.">
<emphasis id="54E1DA29FF9CFFADBD9FA5393A4EE627" box="[929,1058,583,609]" italics="true" pageId="2" pageNumber="453">C. wilsonia</emphasis>
Bennett
</taxonomicName>
<emphasis id="54E1DA29FF9CFFADBAB4A5393A92E627" bold="true" box="[1162,1278,583,609]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="4FD26752FF9CFFADBAB4A5393A92E627" box="[1162,1278,583,609]" pageId="2" pageNumber="453" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
All are rarely encountered species with restricted ranges within the central and southern Sierra
<collectingRegion id="A451C8D9FF9CFFADBA77A5153AF3E6C3" box="[1097,1183,619,645]" country="United States of America" name="Nevada" pageId="2" pageNumber="453">Nevada</collectingRegion>
Mountains of eastern
<collectingRegion id="A451C8D9FF9CFFADBEA9A5F13F65E6EF" box="[151,265,655,681]" country="United States of America" name="California" pageId="2" pageNumber="453">California</collectingRegion>
from the Lake Tahoe area south to
<taxonomicName id="A1957DB8FF9CFFADBCABA5F13D51E6EF" authority="County." authorityName="County." box="[661,829,655,681]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="tulare">Tulare County.</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
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137
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Genus
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<emphasis id="54E1DA29FF9FFFAEBED0A31C3F39E03B" bold="true" box="[238,341,1122,1149]" italics="true" pageId="1" pageNumber="452">Cybaeus</emphasis>
L.
<bibRefCitation id="02047BCAFF9FFFAEBF41A31D3C68E03A" author="Koch, L." box="[383,516,1122,1149]" pageId="1" pageNumber="452" pagination="1 - 52" refId="ref16783" refString="Koch, L. (1868) Die Arachnidengattungen Amaurobius, Coelotes und Cybaeus. Abhandlungen der Naturhistorischen Gesellschaft zu Nurnberg, 4, 1 - 52." type="journal article" year="1868">Koch 1868</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8F55B0FF9FFFAEBEA9A3D93A80E1D1" pageId="1" pageNumber="452" type="type_taxon">
<paragraph id="662A063BFF9FFFAEBEA9A3D93CC4E0F9" blockId="1.[151,680,1191,1215]" box="[151,680,1191,1215]" pageId="1" pageNumber="452">
<emphasis id="54E1DA29FF9FFFAEBEA9A3D93F4FE0F9" bold="true" box="[151,291,1191,1215]" pageId="1" pageNumber="452">
<typeStatus id="B92EB899FF9FFFAEBEA9A3D93EA0E0F9" box="[151,204,1191,1215]" pageId="1" pageNumber="452">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="A1957DB8FF9FFFAEBF14A3D93CC8E0F9" authority="C.L. Koch 1839" authorityName="C. L. Koch" authorityYear="1839" box="[298,676,1191,1215]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="tetricus">
<emphasis id="54E1DA29FF9FFFAEBF14A3D93F9BE0F9" box="[298,503,1191,1215]" italics="true" pageId="1" pageNumber="452">Amaurobius tetricus</emphasis>
C.L.
<bibRefCitation id="02047BCAFF9FFFAEBC0FA3D93CC8E0F9" author="Koch, C. L." box="[561,676,1191,1215]" pageId="1" pageNumber="452" refId="ref16760" refString="Koch, C. L. (1839) Die Arachniden. Sechster Band. Zeh'sche Buchhandlung, Nurnberg, 156 pp." type="book" year="1839">Koch 1839</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="662A063BFF9FFFAEBEA9A3933A80E1D1" blockId="1.[151,1437,1261,1432]" pageId="1" pageNumber="452">
<emphasis id="54E1DA29FF9FFFAEBEA9A3933EBAE141" bold="true" box="[151,214,1261,1287]" pageId="1" pageNumber="452">Note.</emphasis>
<bibRefCitation id="02047BCAFF9FFFAEBEE1A3903FE8E14E" author="Bennett, R. G." box="[223,388,1261,1288]" pageId="1" pageNumber="452" pagination="85 - 90" refId="ref14056" refString="Bennett, R. G. (2005) Cybaeidae. In: Ubick, D., Paquin, P., Cushing, P. E. &amp; Roth, V. D. (Eds.), Spiders of North America: an Identification Manual. American Arachnological Society, pp. 85 - 90." type="book chapter" year="2005">Bennett (2005</bibRefCitation>
,
<bibRefCitation id="02047BCAFF9FFFAEBFAFA3933FA7E141" author="Bennett, R. G." box="[401,459,1261,1287]" pageId="1" pageNumber="452" pagination="96 - 101" refId="ref14165" refString="Bennett, R. G. (2017) Cybaeidae. In: Ubick, D., Paquin, P., Cushing, P. E. &amp; Roth, V. D. (Eds), Spiders of North America: An Identification Manual, 2 nd ed. American Arachnological Society, pp. 96 - 101." type="book chapter" year="2017">2017</bibRefCitation>
) provided keys to all Nearctic genera then classified in
<taxonomicName id="A1957DB8FF9FFFAEBA6EA3933AA6E141" ID-CoL="8TS" authorityName="Banks" authorityYear="1892" box="[1104,1226,1261,1287]" class="Arachnida" family="Cybaeidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="family">Cybaeidae</taxonomicName>
. See
<bibRefCitation id="02047BCAFF9FFFAEBB36A3933E8EE16D" author="Copley, C. R. &amp; Bennett, R. &amp; Perlman, S. J." pageId="1" pageNumber="452" pagination="367 - 401" refId="ref15419" refString="Copley, C. R., Bennett, R. &amp; Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
Copley
<emphasis id="54E1DA29FF9FFFAEBB5CA3903BF1E141" box="[1378,1437,1261,1287]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
(2009)
</bibRefCitation>
for genus diagnosis and descriptive details as well as a key to the species groups (females only) of the Holarctic and Californian clades of
<taxonomicName id="A1957DB8FF9FFFAEBC0CA24B3CFEE109" ID-CoL="3XSF" box="[562,658,1333,1359]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="genus">
<emphasis id="54E1DA29FF9FFFAEBC0CA24B3CFEE109" box="[562,658,1333,1359]" italics="true" pageId="1" pageNumber="452">Cybaeus</emphasis>
</taxonomicName>
. Males of
<taxonomicName id="A1957DB8FF9FFFAEBD2EA24B3DA7E109" box="[784,971,1333,1359]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="54E1DA29FF9FFFAEBD2EA24B3D1CE109" box="[784,880,1333,1359]" italics="true" pageId="1" pageNumber="452">Cybaeus</emphasis>
species
</taxonomicName>
often are difficult to assign to particular species groups. See
<bibRefCitation id="02047BCAFF9FFFAEBF46A2243C35E135" author="Bennett, R. &amp; Copley, C. &amp; Copley, D." box="[376,601,1369,1396]" pageId="1" pageNumber="452" pagination="1 - 67" refId="ref14228" refString="Bennett, R., Copley, C. &amp; Copley, D. (2016) Cybaeus (Araneae: Cybaeidae): the Nearctic species of the Holarctic clade. Zootaxa, 4164 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 4164.1.1" type="journal article" year="2016">
Bennett
<emphasis id="54E1DA29FF9FFFAEBFE8A2243C63E135" box="[470,527,1369,1395]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
(2016
</bibRefCitation>
,
<bibRefCitation id="02047BCAFF9FFFAEBC5BA2273CCCE135" author="Bennett, R. &amp; Copley, C. &amp; Copley, D." box="[613,672,1369,1395]" pageId="1" pageNumber="452" pagination="245 - 274" refId="ref14283" refString="Bennett, R., Copley, C. &amp; Copley, D. (2019) Cybaeus (Araneae: Cybaeidae): the adenes species group of the Californian clade. Zootaxa, 4711 (2), 245 - 274. https: // doi. org / 10.11646 / zootaxa. 4711.2.2" type="journal article" year="2019">2019</bibRefCitation>
,
<bibRefCitation id="02047BCAFF9FFFAEBC94A2273C9EE135" author="Bennett, R. &amp; Copley, C. &amp; Copley, D." box="[682,754,1369,1395]" pageId="1" pageNumber="452" pagination="224 - 244" refId="ref14397" refString="Bennett, R., Copley, C. &amp; Copley, D. (2021 a) Cybaeus (Araneae: Cybaeidae): the aspenicolens species group of the Californian clade. Zootaxa, 4926 (2), 224 - 244. https: // doi. org / 10.11646 / zootaxa. 4926.2.4" type="journal article" year="2021">2021a</bibRefCitation>
,
<bibRefCitation id="02047BCAFF9FFFAEBCC3A2273D28E132" author="Bennett, R. &amp; Copley, C. &amp; Copley, D." box="[765,836,1369,1396]" pageId="1" pageNumber="452" pagination="401 - 436" refId="ref14454" refString="Bennett, R., Copley, C. &amp; Copley, D. (2021 b) Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade. Zootaxa, 4965 (3), 401 - 436. https: // doi. org / 10.11646 / zootaxa. 4965.3.1" type="journal article" year="2021">2021b</bibRefCitation>
) for, respectively, revisions of the Nearctic species of the Holarctic clade and of the
<taxonomicName id="A1957DB8FF9FFFAEBFD7A2033C5BE1D1" ID-CoL="32RD8" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[489,567,1405,1431]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="adenes">
<emphasis id="54E1DA29FF9FFFAEBFD7A2033C5BE1D1" box="[489,567,1405,1431]" italics="true" pageId="1" pageNumber="452">adenes</emphasis>
</taxonomicName>
,
<taxonomicName id="A1957DB8FF9FFFAEBC7AA2033CB9E1D1" ID-CoL="32RDN" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[580,725,1405,1431]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="aspenicolens">
<emphasis id="54E1DA29FF9FFFAEBC7AA2033CB9E1D1" box="[580,725,1405,1431]" italics="true" pageId="1" pageNumber="452">aspenicolens</emphasis>
</taxonomicName>
, and
<taxonomicName id="A1957DB8FF9FFFAEBD2CA2003DECE1D1" ID-CoL="6C55P" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[786,896,1406,1431]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="consocius">
<emphasis id="54E1DA29FF9FFFAEBD2CA2003DECE1D1" box="[786,896,1406,1431]" italics="true" pageId="1" pageNumber="452">consocius</emphasis>
</taxonomicName>
groups of the Californian clade.
</paragraph>
</subSubSection>
</treatment>
</document>