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<document id="AD622E0A682891C7851F8A65A11EBBEB" ID-DOI="10.11646/zootaxa.4039.4.1" ID-GBIF-Dataset="ca91ab86-49b9-4d90-ac6f-2051ca9d2f85" ID-ISSN="1175-5326" ID-Zenodo-Dep="232034" ID-ZooBank="4B12AA52-6A3F-44B9-BBEC-EBABF0E22DF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461332656152" checkinUser="plazi" docAuthor="Blake, James A. &amp; Göransson, Peter" docDate="2015" docId="EE2B2061FFC0027EFF2F0F1EFC03F9FA" docLanguage="en" docName="zt04039p515.pdf" docOrigin="Zootaxa 4039 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tharyx robustus Blake &amp; Göransson, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="510" masterDocId="12125819FFC70277FFB8095EFFE8FF8E" masterDocTitle="Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)" masterLastPageNumber="515" masterPageNumber="501" pageNumber="508" updateTime="1698618770507" updateUser="plazi">
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<mods:title id="FAAADEE19FF40D6B81C4864D707769EF">Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)</mods:title>
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<mods:namePart id="19F05EE666C043AAA8A3771C58EA827D">Blake, James A.</mods:namePart>
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<mods:namePart id="29B513314F5C99ECF394FF9DA09A63B8">Göransson, Peter</mods:namePart>
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<mods:date id="5529892E6F3B403E3A1FFE94E5245B40">2015</mods:date>
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<treatment id="EE2B2061FFC0027EFF2F0F1EFC03F9FA" ID-DOI="http://doi.org/10.5281/zenodo.5658081" ID-GBIF-Taxon="119644309" ID-Zenodo-Dep="5658081" LSID="urn:lsid:plazi:treatment:EE2B2061FFC0027EFF2F0F1EFC03F9FA" httpUri="http://treatment.plazi.org/id/EE2B2061FFC0027EFF2F0F1EFC03F9FA" lastPageId="9" lastPageNumber="510" pageId="7" pageNumber="508">
<subSubSection id="2E98C2FCFFC00270FF2F0F1EFED8F9F5" pageId="7" pageNumber="508" type="nomenclature">
<paragraph id="663D9177FFC00270FF2F0F1EFE1DF9D4" blockId="7.[151,501,1600,1660]" box="[151,501,1600,1626]" pageId="7" pageNumber="508">
<heading id="3D75261BFFC00270FF2F0F1EFE1DF9D4" bold="true" box="[151,501,1600,1626]" fontSize="11" level="1" pageId="7" pageNumber="508" reason="1">
<emphasis id="54F64D65FFC00270FF2F0F1EFE1DF9D4" bold="true" box="[151,501,1600,1626]" pageId="7" pageNumber="508">
<taxonomicName id="A182EAF4FFC00270FF2F0F1EFEB6F9D4" ID-CoL="564MP" box="[151,350,1600,1626]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFC00270FF2F0F1EFEB6F9D4" bold="true" box="[151,350,1600,1626]" italics="true" pageId="7" pageNumber="508">Tharyx robustus</emphasis>
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<taxonomicNameLabel id="4FC5F01EFFC00270FEDD0F1EFE1DF9D4" box="[357,501,1600,1626]" pageId="7" pageNumber="508" rank="species">new species</taxonomicNameLabel>
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<paragraph id="663D9177FFC00270FF2F0F3DFED8F9F5" blockId="7.[151,501,1600,1660]" box="[151,304,1635,1660]" pageId="7" pageNumber="508">
<figureCitation id="FEB98DF2FFC00270FF2F0F3DFF17F9F5" box="[151,255,1635,1660]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="7" pageNumber="508">Figures 3</figureCitation>
D, 4
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC00270FF2F0FF5FF08F882" pageId="7" pageNumber="508" type="materials_examined">
<paragraph id="663D9177FFC00270FF2F0FF5FF08F882" blockId="7.[151,1437,1707,2020]" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF2F0FF5FE62F94A" bold="true" box="[151,394,1707,1732]" pageId="7" pageNumber="508">Material examined.</emphasis>
<collectingCountry id="1E95D1E7FFC00270FE230FF5FDFBF94D" box="[411,531,1707,1731]" name="Sweden" pageId="7" pageNumber="508">SWEDEN</collectingCountry>
, Kattegat, near Varberg, 0
<date id="123CB7B7FFC00270FCE70FF5FC11F94D" box="[863,1017,1707,1731]" pageId="7" pageNumber="508" value="2012-08-03">3 Aug 2012</date>
, in sediments near eelgrass beds,
<geoCoordinate id="03B6F7B0FFC00270FF2F0F91FECAF966" box="[151,290,1743,1768]" direction="north" orientation="latitude" pageId="7" pageNumber="508" precision="1" value="57.11501">57.11501°N</geoCoordinate>
;
<geoCoordinate id="03B6F7B0FFC00270FE890F91FE50F966" box="[305,440,1743,1768]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="1" value="12.22611">12.22611°E</geoCoordinate>
,
<quantity id="A17A3C92FFC00270FE7F0F91FDD4F969" box="[455,572,1743,1767]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.7" metricValueMin="0.3" pageId="7" pageNumber="508" unit="m" value="1.0" valueMax="1.7" valueMin="0.3">0.31.7 m</quantity>
depth, coll. P. Göransson,
<typeStatus id="B9392FD5FFC00270FCC30F91FC35F966" box="[891,989,1743,1768]" pageId="7" pageNumber="508" type="holotype">holotype</typeStatus>
(
<collectionCode id="009309B2FFC00270FC570F91FBA9F969" box="[1007,1089,1743,1767]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="7" pageNumber="508">SMNH</collectionCode>
8756), six
<typeStatus id="B9392FD5FFC00270FB740F8EFADFF966" box="[1228,1335,1744,1768]" pageId="7" pageNumber="508" type="paratype">paratypes</typeStatus>
(
<collectionCode id="009309B2FFC00270FAF10F91FA74F969" box="[1353,1436,1743,1767]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="7" pageNumber="508">SMNH</collectionCode>
8757).
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC0027EFF7F0E49FBCAFC2B" lastPageId="9" lastPageNumber="510" pageId="7" pageNumber="508" type="description">
<paragraph id="663D9177FFC0027EFF7F0E49FB92FF76" blockId="7.[151,1437,1707,2020]" lastBlockId="9.[151,1436,151,1653]" lastPageId="9" lastPageNumber="510" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF7F0E49FEB1F8BE" bold="true" box="[199,345,1815,1840]" pageId="7" pageNumber="508">Description.</emphasis>
A moderate-sized species,
<typeStatus id="B9392FD5FFC00270FD270E49FCE9F8BE" box="[671,769,1815,1840]" pageId="7" pageNumber="508" type="holotype">holotype</typeStatus>
complete,
<quantity id="A17A3C92FFC00270FC300E49FC26F8A1" box="[904,974,1815,1839]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="508" unit="mm" value="8.0">8 mm</quantity>
long, 1.0 mm wide for 105 setigerous segments;
<typeStatus id="B9392FD5FFC00270FEAC0E62FE97F8DA" box="[276,383,1852,1876]" pageId="7" pageNumber="508" type="paratype">paratypes</typeStatus>
include complete and incomplete specimens,
<quantity id="A17A3C92FFC00270FC1C0E65FBEFF8DD" box="[932,1031,1851,1876]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="7" pageNumber="508" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
long,
<quantity id="A17A3C92FFC00270FBE10E65FB5DF8DD" box="[1113,1205,1851,1876]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="508" unit="mm" value="0.5">0.5 mm</quantity>
wide, with 80116 setigerous segments. Color in alcohol light tan to opaque white, with a cluster of black pigment on posterior lateral margins of peristomium (
<figureCitation id="FEB98DF2FFC00270FE0D0EDDFE11F815" box="[437,505,1923,1948]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="7" pageNumber="508">Fig. 4</figureCitation>
C); some specimens with brown pigment in lateral segmental grooves in middle of body. Body broad, dorsoventrally flattened throughout with numerous narrow segments (
<figureCitation id="FEB98DF2FFC00270FBCE0EF9FB29F831" box="[1142,1217,1959,1984]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="7" pageNumber="508">Figs. 3</figureCitation>
D, 4AB). Anterior segments becoming widest at about setiger 15, continuing posteriorly, narrowing at about setiger 60, then expanding again in posterior region, narrowing in far posterior segments to a simple pygidium consisting of a narrow, conical lobe; anal cirri absent (
<figureCitation id="FEB98DF2FFCE027EFDF609E2FD7BFF5A" box="[590,659,188,213]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
B). Venter flattened, with shallow ventral groove bearing mid-ventral ridge formed of two raised pads where each segment meets at ventral mid-line (
<figureCitation id="FEB98DF2FFCE027EFBAB0981FBBEFF76" box="[1043,1110,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="9" pageNumber="510">Fig. 3</figureCitation>
D).
</paragraph>
<caption id="32FDC1FFFFCF027FFF2F0E65FE7DF864" httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="8" pageNumber="509" targetBox="[187,1387,207,1825]" targetPageId="8">
<paragraph id="663D9177FFCF027FFF2F0E65FE7DF864" blockId="8.[151,1436,1851,2026]" pageId="8" pageNumber="509">
<emphasis id="54F64D65FFCF027FFF2F0E65FEFCF8DE" bold="true" box="[151,276,1851,1873]" pageId="8" pageNumber="509">FIGURE 4.</emphasis>
<taxonomicName id="A182EAF4FFCF027FFEA40E65FE2AF8DF" box="[284,450,1851,1873]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="8" pageNumber="509" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFCF027FFEA40E65FE2AF8DF" box="[284,450,1851,1873]" italics="true" pageId="8" pageNumber="509">Tharyx robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFCF027FFE730E65FDEEF8DE" box="[459,518,1851,1872]" pageId="8" pageNumber="509" rank="species">
<emphasis id="54F64D65FFCF027FFE730E65FDEEF8DE" bold="true" box="[459,518,1851,1872]" pageId="8" pageNumber="509">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle.
</paragraph>
</caption>
<paragraph id="663D9177FFCE027EFF7F085AFB0FFE5E" blockId="9.[151,1436,151,1653]" pageId="9" pageNumber="510">
Prostomium triangular, acutely pointed on anterior margin, broadening posteriorly (
<figureCitation id="FEB98DF2FFCE027EFB06085AFAFBFE92" box="[1214,1299,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="9" pageNumber="510">Figs. 3</figureCitation>
D, 4A, C); peristomium wider than long with weak dorsolateral groove forming two indistinct annuli (
<figureCitation id="FEB98DF2FFCE027EFB370879FB3BFECE" box="[1167,1235,295,320]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
C). Nuchal organs inconspicuous, located on posterior margin of prostomium (
<figureCitation id="FEB98DF2FFCE027EFC820812FC95FEEA" box="[826,893,332,357]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
C), not pigmented; eyes absent. Dorsal tentacles present on posterior margin of peristomium, with first pair of branchiae located immediately posterior to tentacles, also on peristomium (
<figureCitation id="FEB98DF2FFCE027EFE3708CAFE37FE22" box="[399,479,404,429]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Figs. 4</figureCitation>
A, C); second pair of branchiae on posterior margin of setiger 1, medial to notosetal fascicle; in posterior setigers branchiae arise in middle of segment rather than on posterior margin.
</paragraph>
<paragraph id="663D9177FFCE027EFF7F0882FEB3FDEE" blockId="9.[151,1436,151,1653]" pageId="9" pageNumber="510">
Parapodia on both sides of body separated from broad dorsum by a narrow groove extending from peristomium to end of body; individual notopodia slightly elevated above this groove, less obvious in posterior one-third of body (
<figureCitation id="FEB98DF2FFCE027EFECE0B7AFE55FDB2" box="[374,445,548,573]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
A). Broad dorsum of body raised above parapodia, rounded, low, without mid-dorsal groove (
<figureCitation id="FEB98DF2FFCE027EFF4C0B19FEDFFDEE" box="[244,311,583,608]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="663D9177FFCE027EFF7F0B33FE84FCD3" blockId="9.[151,1436,151,1653]" pageId="9" pageNumber="510">
Noto- and neuropodia with setal fascicles arising close together (
<figureCitation id="FEB98DF2FFCE027EFC640B32FBD9FD0A" box="[988,1073,620,645]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Figs. 4</figureCitation>
D, E), anterior parapodia with spreading fascicles of about 78 capillary setae in notopodia and 810 capillaries in neuropodia; in far posterior notopodia, 23 transitional pointed setae (
<figureCitation id="FEB98DF2FFCE027EFDC20BEAFD56FD42" box="[634,702,692,717]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
F) occur with capillaries from about setiger 85 replaced by stiff elongate spines from about setiger 8890 (
<figureCitation id="FEB98DF2FFCE027EFDCF0B89FD53FD7E" box="[631,699,727,752]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
H); neuropodial spines first present from setigers 8090 with 37 capillaries and 13 short, spines with variable lengths (
<figureCitation id="FEB98DF2FFCE027EFC980BA2FC9AFC9A" box="[800,882,764,789]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">Figs. 4</figureCitation>
HJ); all spines blunt-tipped, with apex weakly expanded, knob-like (
<figureCitation id="FEB98DF2FFCE027EFE290A41FE0AFCB6" box="[401,482,799,824]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[187,1387,207,1825]" captionTargetId="figure@8.[172,1408,193,1828]" captionTargetPageId="8" captionText="FIGURE 4. Tharyx robustus n. sp. Holotype (SMNH 8756): A, anterior end, dorsal view; B, posterior end, dorsal view; C, anterior end, right lateral view. Paratypes (SMNH 8757): D, posterior setiger 80, anterior view; E, far posterior setiger 90, anterior view; F, anteriormost notopodial aciculars transitional between capillaries and blunt-tipped spines (G); G, notopodial acicular spine from setiger 90 (E); H, neuroacicular spines and capillaries from setiger 80 (D); I, neuroacicular spines from E; J, detail of neuroacicular spines from another paratype. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232038/files/figure.png" pageId="9" pageNumber="510">
Figs.
<date id="123CB7B7FFCE027EFE6B0A7EFE0AFCB6" box="[467,482,800,824]" pageId="9" pageNumber="510">4</date>
</figureCitation>
IJ). Transitional notopodial spines with angled pointed tip, sometimes with fibrils visible along shaft.
</paragraph>
<paragraph id="663D9177FFCE027EFF7F0A39FBCAFC2B" blockId="9.[151,1436,151,1653]" pageId="9" pageNumber="510">
<emphasis id="54F64D65FFCE027EFF7F0A39FE5CFC0E" bold="true" box="[199,436,871,896]" pageId="9" pageNumber="510">Methyl Green Stain</emphasis>
. No distinct stain apparent; most of the body stains darkly, but de-stains rapidly leaving no pattern after differentiation. The prostomium and pygidium never retain stain.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFCE027EFF7F0AF1FEF8FA6B" pageId="9" pageNumber="510" type="discussion">
<paragraph id="663D9177FFCE027EFF7F0AF1FEF8FA6B" blockId="9.[151,1436,151,1653]" pageId="9" pageNumber="510">
<emphasis id="54F64D65FFCE027EFF7F0AF1FED3FC46" bold="true" box="[199,315,943,968]" pageId="9" pageNumber="510">Remarks.</emphasis>
In having a robust dorsoventrally flattened body with crowded segments along its entire length,
<taxonomicName id="A182EAF4FFCE027EFA3F0AEFFF1EFC63" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFCE027EFA3F0AEFFF1EFC63" italics="true" pageId="9" pageNumber="510">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFCE027EFEBC0A8AFEA1FC63" box="[260,329,980,1005]" pageId="9" pageNumber="510" rank="species">
<emphasis id="54F64D65FFCE027EFEBC0A8AFEA1FC63" bold="true" box="[260,329,980,1005]" pageId="9" pageNumber="510">n. sp.</emphasis>
</taxonomicNameLabel>
most closely resembles
<taxonomicName id="A182EAF4FFCE027EFDCC0A8BFB87FC62" authority="Hutchings &amp; Murray, 1984" authorityName="Hutchings &amp; Murray" authorityYear="1984" box="[628,1135,980,1005]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="retusisetus">
<emphasis id="54F64D65FFCE027EFDCC0A8BFCE5FC62" box="[628,781,981,1004]" italics="true" pageId="9" pageNumber="510">T. retusisetus</emphasis>
(
<bibRefCitation id="0213EC86FFCE027EFC9A0A8AFB8FFC62" author="Hutchings" box="[802,1127,980,1005]" pageId="9" pageNumber="510" refString="Hutchings, P. &amp; Murray, M. (1984) Taxonomy of polychaetes from the Hawkesbury River and the southern estuaries of New South Wales, Australia. Records of the Australian Museum, 36, (Supplement 3), 1 - 118. http: // doi: 10.3853 / j. 0812 - 7387.3.1984.101." type="journal article" year="1984">Hutchings &amp; Murray, 1984</bibRefCitation>
)
</taxonomicName>
from New South
<collectingCountry id="1E95D1E7FFCE027EFAE90A8AFA70FC63" box="[1361,1432,980,1005]" name="United Kingdom" pageId="9" pageNumber="510">Wales</collectingCountry>
,
<collectingCountry id="1E95D1E7FFCE027EFF2F0AA9FEE9FB9E" box="[151,257,1015,1040]" name="Australia" pageId="9" pageNumber="510">Australia</collectingCountry>
. The latter species however, has posterior spines with sub-bidentate knob-like tips that are typical of many species of
<taxonomicName id="A182EAF4FFCE027EFEEF0D42FE4EFBBB" box="[343,422,1052,1077]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFEEF0D42FE4EFBBB" box="[343,422,1052,1077]" italics="true" pageId="9" pageNumber="510">Tharyx</emphasis>
</taxonomicName>
(
<bibRefCitation id="0213EC86FFCE027EFE0F0D42FDD4FBBA" author="Blake" box="[439,572,1052,1077]" pageId="9" pageNumber="510" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake 1991</bibRefCitation>
,
<bibRefCitation id="0213EC86FFCE027EFDF20D42FD6EFBBB" author="Blake" box="[586,646,1052,1077]" pageId="9" pageNumber="510" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">1996</bibRefCitation>
). In contrast, the posterior spines of
<taxonomicName id="A182EAF4FFCE027EFB920D43FB4EFBBB" box="[1066,1190,1052,1077]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFCE027EFB920D43FB4EFBBB" box="[1066,1190,1052,1077]" italics="true" pageId="9" pageNumber="510">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFCE027EFB170D42FB07FBBB" box="[1199,1263,1052,1077]" pageId="9" pageNumber="510" rank="species">
<emphasis id="54F64D65FFCE027EFB170D42FB07FBBB" bold="true" box="[1199,1263,1052,1077]" pageId="9" pageNumber="510">n. sp.</emphasis>
</taxonomicNameLabel>
are variable in length and have only a single knob on the end of the spine and are only weakly expanded. None of these spines resemble the sub-bidentate spines found in several other species. The kind of transitional notopodial spines found in
<taxonomicName id="A182EAF4FFCE027EFF0C0DD7FED8FB2E" box="[180,304,1159,1184]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="robustus">
<emphasis id="54F64D65FFCE027EFF0C0DD7FED8FB2E" box="[180,304,1159,1184]" italics="true" pageId="9" pageNumber="510">T. robustus</emphasis>
</taxonomicName>
has not been reported in other species, but may have been overlooked. The robust body and narrow crowded segments of
<taxonomicName id="A182EAF4FFCE027EFE280DF3FDE4FB4B" box="[400,524,1196,1221]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFCE027EFE280DF3FDE4FB4B" box="[400,524,1196,1221]" italics="true" pageId="9" pageNumber="510">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFCE027EFDAD0DF2FDBDFB4B" box="[533,597,1196,1221]" pageId="9" pageNumber="510" rank="species">
<emphasis id="54F64D65FFCE027EFDAD0DF2FDBDFB4B" bold="true" box="[533,597,1196,1221]" pageId="9" pageNumber="510">n. sp.</emphasis>
</taxonomicNameLabel>
are atypical for species of
<taxonomicName id="A182EAF4FFCE027EFC330DF2FC32FB4B" box="[907,986,1196,1221]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFC330DF2FC32FB4B" box="[907,986,1196,1221]" italics="true" pageId="9" pageNumber="510">Tharyx</emphasis>
</taxonomicName>
and are more reminiscent of species of
<taxonomicName id="A182EAF4FFCE027EFF2F0D91FED9FB66" box="[151,305,1231,1256]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFF2F0D91FED9FB66" box="[151,305,1231,1256]" italics="true" pageId="9" pageNumber="510">Aphelochaeta</emphasis>
</taxonomicName>
. However, another feature that allies this species with
<taxonomicName id="A182EAF4FFCE027EFC1A0D91FC1AFB66" box="[930,1010,1231,1256]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFC1A0D91FC1AFB66" box="[930,1010,1231,1256]" italics="true" pageId="9" pageNumber="510">Tharyx</emphasis>
</taxonomicName>
is the position of the dorsal tentacles and first pair of branchiae. In nearly all previously described species of
<taxonomicName id="A182EAF4FFCE027EFBBB0DAAFBBAFA83" box="[1027,1106,1268,1293]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFBBB0DAAFBBAFA83" box="[1027,1106,1268,1293]" italics="true" pageId="9" pageNumber="510">Tharyx</emphasis>
</taxonomicName>
, the tentacles occur on the peristomium with the first pair of branchiae occurring directly posterior to them, also on the peristomium (
<bibRefCitation id="0213EC86FFCE027EFAE10C49FF39FADA" author="Blake" pageId="9" pageNumber="510" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake 1991</bibRefCitation>
,
<bibRefCitation id="0213EC86FFCE027EFF650C62FEF0FADB" author="Blake" box="[221,280,1340,1365]" pageId="9" pageNumber="510" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">1996</bibRefCitation>
,
<bibRefCitation id="0213EC86FFCE027EFE9C0C62FE89FADB" author="Blake" box="[292,353,1340,1365]" pageId="9" pageNumber="510" refString="Blake, J. A. (2015) New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitzbergen in the Norwegian Arctic. Zootaxa, 3919 (3), 501 - 552. http: // dx. doi. org / 10.11646 / zootaxa. 3919.3.5" type="journal article" year="2015">2015</bibRefCitation>
). This is exactly how these two structures are positioned in
<taxonomicName id="A182EAF4FFCE027EFBBA0C63FB93FADB" box="[1026,1147,1340,1365]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFCE027EFBBA0C63FB93FADB" box="[1026,1147,1340,1365]" italics="true" pageId="9" pageNumber="510">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFCE027EFB3D0C62FB2DFADB" box="[1157,1221,1340,1365]" pageId="9" pageNumber="510" rank="species">
<emphasis id="54F64D65FFCE027EFB3D0C62FB2DFADB" bold="true" box="[1157,1221,1340,1365]" pageId="9" pageNumber="510">n. sp.</emphasis>
</taxonomicNameLabel>
In addition, lateral peristomial pigment spots are present on
<taxonomicName id="A182EAF4FFCE027EFDD80C3FFD30FAF6" box="[608,728,1375,1400]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFCE027EFDD80C3FFD30FAF6" box="[608,728,1375,1400]" italics="true" pageId="9" pageNumber="510">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFCE027EFD590C01FCF7FAF6" box="[737,799,1375,1400]" pageId="9" pageNumber="510" rank="species">
<emphasis id="54F64D65FFCE027EFD590C01FCF7FAF6" bold="true" box="[737,799,1375,1400]" pageId="9" pageNumber="510">n. sp.</emphasis>
</taxonomicNameLabel>
These are known from other species of
<taxonomicName id="A182EAF4FFCE027EFB640C01FAC3FAF6" box="[1244,1323,1375,1400]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFB640C01FAC3FAF6" box="[1244,1323,1375,1400]" italics="true" pageId="9" pageNumber="510">Tharyx</emphasis>
</taxonomicName>
including
<taxonomicName id="A182EAF4FFCE027EFF2F0CDBFE39FA12" authority="Blake, 1991" authorityName="Blake" authorityYear="1991" box="[151,465,1412,1437]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="kirkegaardi">
<emphasis id="54F64D65FFCE027EFF2F0CDBFED0FA13" box="[151,312,1412,1437]" italics="true" pageId="9" pageNumber="510">T. kirkegaardi</emphasis>
<bibRefCitation id="0213EC86FFCE027EFEFB0CDAFE39FA12" author="Blake" box="[323,465,1412,1437]" pageId="9" pageNumber="510" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A182EAF4FFCE027EFE590CDBFCE6FA13" authority="Blake, 2015" authorityName="Blake" authorityYear="2015" box="[481,782,1412,1437]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="circacutus">
<emphasis id="54F64D65FFCE027EFE590CDBFD9CFA12" box="[481,628,1413,1436]" italics="true" pageId="9" pageNumber="510">T. circacutus</emphasis>
<bibRefCitation id="0213EC86FFCE027EFDC70CDAFCE6FA13" author="Blake" box="[639,782,1412,1437]" pageId="9" pageNumber="510" refString="Blake, J. A. (2015) New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitzbergen in the Norwegian Arctic. Zootaxa, 3919 (3), 501 - 552. http: // dx. doi. org / 10.11646 / zootaxa. 3919.3.5" type="journal article" year="2015">Blake, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A182EAF4FFCE027EFCA40CDBFC67FA12" box="[796,911,1413,1436]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFCE027EFCA40CDBFC67FA12" box="[796,911,1413,1436]" italics="true" pageId="9" pageNumber="510">T. maryae</emphasis>
</taxonomicName>
<emphasis id="54F64D65FFCE027EFC230CDAFC0EFA13" bold="true" box="[923,998,1412,1437]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="4FC5F01EFFCE027EFC230CDAFC08FA13" box="[923,992,1412,1437]" pageId="9" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
and other species as yet undescribed (Blake, pers. obs.). Such pigment has not been reported for species of
<taxonomicName id="A182EAF4FFCE027EFC410CF9FB7BFA4E" box="[1017,1171,1447,1472]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFC410CF9FB7BFA4E" box="[1017,1171,1447,1472]" italics="true" pageId="9" pageNumber="510">Aphelochaeta</emphasis>
</taxonomicName>
or other bitentaculate cirratulids.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFCE027EFF7F0CB1FCF1F9A2" pageId="9" pageNumber="510" type="etymology">
<paragraph id="663D9177FFCE027EFF7F0CB1FCF1F9A2" blockId="9.[151,1436,151,1653]" pageId="9" pageNumber="510">
<emphasis id="54F64D65FFCE027EFF7F0CB1FEA6F986" bold="true" box="[199,334,1519,1544]" pageId="9" pageNumber="510">Etymology.</emphasis>
The name of this species is derived from the word “robust,” and refers to the nature of the overall body shape which is unlike most other species of
<taxonomicName id="A182EAF4FFCE027EFD7B0F4AFCFAF9A3" box="[707,786,1556,1581]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFCE027EFD7B0F4AFCFAF9A3" box="[707,786,1556,1581]" italics="true" pageId="9" pageNumber="510">Tharyx</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFCE027EFF7F0F69FD04F9DE" box="[199,748,1591,1616]" pageId="9" pageNumber="510" type="biology_ecology">
<paragraph id="663D9177FFCE027EFF7F0F69FD04F9DE" blockId="9.[151,1436,151,1653]" box="[199,748,1591,1616]" pageId="9" pageNumber="510">
<emphasis id="54F64D65FFCE027EFF7F0F69FEF7F9DE" bold="true" box="[199,287,1591,1616]" pageId="9" pageNumber="510">Biology</emphasis>
. Same data as for
<taxonomicName id="A182EAF4FFCE027EFE560F67FDB4F9DE" box="[494,604,1593,1616]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="510" phylum="Annelida" rank="species" species="maryae">
<emphasis id="54F64D65FFCE027EFE560F67FDB4F9DE" box="[494,604,1593,1616]" italics="true" pageId="9" pageNumber="510">T. maryae</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFCE027EFF7F0F02FC03F9FA" box="[199,1003,1628,1653]" pageId="9" pageNumber="510" type="distribution">
<paragraph id="663D9177FFCE027EFF7F0F02FC03F9FA" blockId="9.[151,1436,151,1653]" box="[199,1003,1628,1653]" pageId="9" pageNumber="510">
<emphasis id="54F64D65FFCE027EFF7F0F02FE88F9FB" bold="true" box="[199,352,1628,1653]" pageId="9" pageNumber="510">Distribution.</emphasis>
Known only from the west coast of
<collectingCountry id="1E95D1E7FFCE027EFD410F02FCBAF9FB" box="[761,850,1628,1653]" name="Sweden" pageId="9" pageNumber="510">Sweden</collectingCountry>
in low water.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,349 @@
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<mods:title id="1B58C10FE66C64E83748006B38030568">Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)</mods:title>
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<mods:namePart id="495DF2F864412500BC5FCAC4D20173D7">Blake, James A.</mods:namePart>
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<mods:part id="255E3E7C51620EEBB2EEE8EF80B6E3CB">
<mods:date id="9B474128EF5A4D3FCB8C4B558789754B">2015</mods:date>
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<treatment id="EE2B2061FFC30270FF2F0BCEFC03FA62" ID-DOI="http://doi.org/10.5281/zenodo.5658079" ID-GBIF-Taxon="119644308" ID-Zenodo-Dep="5658079" LSID="urn:lsid:plazi:treatment:EE2B2061FFC30270FF2F0BCEFC03FA62" httpUri="http://treatment.plazi.org/id/EE2B2061FFC30270FF2F0BCEFC03FA62" lastPageId="7" lastPageNumber="508" pageId="4" pageNumber="505">
<subSubSection id="2E98C2FCFFC30273FF2F0BCEFEB9FD45" pageId="4" pageNumber="505" type="nomenclature">
<paragraph id="663D9177FFC30273FF2F0BCEFE0FFD24" blockId="4.[151,487,656,715]" box="[151,487,656,682]" pageId="4" pageNumber="505">
<heading id="3D75261BFFC30273FF2F0BCEFE0FFD24" bold="true" box="[151,487,656,682]" fontSize="11" level="1" pageId="4" pageNumber="505" reason="1">
<emphasis id="54F64D65FFC30273FF2F0BCEFE0FFD24" bold="true" box="[151,487,656,682]" pageId="4" pageNumber="505">
<taxonomicName id="A182EAF4FFC30273FF2F0BCEFEA7FD24" ID-CoL="7C6Y6" box="[151,335,656,682]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="505" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC30273FF2F0BCEFEA7FD24" bold="true" box="[151,335,656,682]" italics="true" pageId="4" pageNumber="505">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC30273FEEF0BCEFE0FFD24" box="[343,487,656,682]" pageId="4" pageNumber="505" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="663D9177FFC30273FF2F0BECFEB9FD45" blockId="4.[151,487,656,715]" box="[151,337,690,715]" pageId="4" pageNumber="505">
<figureCitation id="FEB98DF2FFC30273FF2F0BECFEEAFD45" box="[151,258,690,715]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figures 2</figureCitation>
,
<figureCitation id="FEB98DF2FFC30273FEB60BEDFEF2FD45" box="[270,282,691,715]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="4" pageNumber="505">3</figureCitation>
AC
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30273FF2F0BA4FF37FCD5" pageId="4" pageNumber="505" type="materials_examined">
<paragraph id="663D9177FFC30273FF2F0BA4FF37FCD5" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF2F0BA4FE62FC9D" bold="true" box="[151,394,762,787]" pageId="4" pageNumber="505">Material examined.</emphasis>
<collectingCountry id="1E95D1E7FFC30273FE230BA5FDFBFC9D" box="[411,531,763,787]" name="Sweden" pageId="4" pageNumber="505">SWEDEN</collectingCountry>
, Kattegat, near Varberg, 0
<date id="123CB7B7FFC30273FCE70BA5FC11FC9D" box="[863,1017,763,787]" pageId="4" pageNumber="505" value="2012-08-03">3 Aug 2012</date>
, in sediments near eelgrass beds,
<geoCoordinate id="03B6F7B0FFC30273FF2F0A40FECAFCB9" box="[151,290,798,823]" direction="north" orientation="latitude" pageId="4" pageNumber="505" precision="1" value="57.11501">57.11501°N</geoCoordinate>
;
<geoCoordinate id="03B6F7B0FFC30273FE880A40FE5FFCB9" box="[304,439,798,823]" direction="east" orientation="longitude" pageId="4" pageNumber="505" precision="1" value="12.22611">12.22611°E</geoCoordinate>
,
<quantity id="A17A3C92FFC30273FE7D0A41FDD1FCB8" box="[453,569,799,823]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.7" metricValueMin="0.3" pageId="4" pageNumber="505" unit="m" value="1.0" valueMax="1.7" valueMin="0.3">0.31.7 m</quantity>
depth, coll. P. Göransson,
<typeStatus id="B9392FD5FFC30273FCCA0A40FC3CFCB9" box="[882,980,798,823]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
(
<collectionCode id="009309B2FFC30273FC5E0A41FBD1FCB9" box="[998,1081,799,823]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="4" pageNumber="505">SMNH</collectionCode>
8754), four
<typeStatus id="B9392FD5FFC30273FB750A41FAD0FCB9" box="[1229,1336,799,823]" pageId="4" pageNumber="505" type="paratype">paratypes</typeStatus>
(
<collectionCode id="009309B2FFC30273FAF10A41FA74FCB9" box="[1353,1436,799,823]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="4" pageNumber="505">SMNH</collectionCode>
8755).
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30273FF7F0A38FD53F8E1" pageId="4" pageNumber="505" type="description">
<paragraph id="663D9177FFC30273FF7F0A38FBAEFB69" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0A38FEB1FCF1" bold="true" box="[199,345,870,895]" pageId="4" pageNumber="505">Description.</emphasis>
A moderate-sized species,
<typeStatus id="B9392FD5FFC30273FD360A38FD18FCF1" box="[654,752,870,895]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
complete,
<quantity id="A17A3C92FFC30273FCD50A39FC55FCF0" box="[877,957,871,895]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="505" unit="mm" value="10.0">10 mm</quantity>
long,
<quantity id="A17A3C92FFC30273FBBD0A38FBB4FCF0" box="[1029,1116,870,895]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="505" unit="mm" value="0.5">0.5 mm</quantity>
wide across thorax,
<quantity id="A17A3C92FFC30273FAFE0A39FA74FCF0" box="[1350,1436,871,895]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="505" unit="mm" value="0.8">0.8 mm</quantity>
wide across expanded posterior end for 115 setigerous segments;
<typeStatus id="B9392FD5FFC30273FC390AD5FC04FC2D" box="[897,1004,907,931]" pageId="4" pageNumber="505" type="paratype">paratypes</typeStatus>
all incomplete (lacking posteriormost segments), up to
<quantity id="A17A3C92FFC30273FEEE0AF1FE7FFC48" box="[342,407,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="505" unit="mm" value="9.0">9 mm</quantity>
long,
<quantity id="A17A3C92FFC30273FE660AF0FDDBFC48" box="[478,563,942,967]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="505" unit="mm" value="0.5">0.5 mm</quantity>
wide, with about 70 setigerous segments. Color in alcohol light tan to opaque white; lateral black pigment spots present on peristomium of some specimens (
<figureCitation id="FEB98DF2FFC30273FBAF0A8CFBB1FC65" box="[1047,1113,978,1003]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="4" pageNumber="505">Fig. 3</figureCitation>
A), faded or not apparent on others; some specimens with brown pigment spots in intersegmental grooves along body. Body elongate, laterally expanded through first 2025 thoracic setigers with narrow crowded segments (
<figureCitation id="FEB98DF2FFC30273FBA80D44FBB5FBBD" box="[1040,1117,1050,1075]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
A, 3C); abdominal segments narrower, longer and rounded, but not moniliform; posterior 2025 segments becoming enlarged, forming expanded posterior end with narrow segments, tapering to pygidium with dorsal anus and bulbous ventral lobe (
<figureCitation id="FEB98DF2FFC30273FF270DD8FF07FB11" box="[159,239,1158,1183]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
B, 3B). Dorsum of thoracic region broad between notopodia, with segmentation evident (
<figureCitation id="FEB98DF2FFC30273FB5E0DD8FAC3FB11" box="[1254,1323,1158,1183]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
A); dorsal groove absent. Venter generally flattened with mid-ventral line of ridges along entire body from setigers 23 (
<figureCitation id="FEB98DF2FFC30273FAD70DF4FF4DFB69" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="4" pageNumber="505">Fig. 3</figureCitation>
C); ridges formed from ventromedial bulges or pads associated with each segment.
</paragraph>
<paragraph id="663D9177FFC30273FF7F0DACFD26FA31" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
Prostomium triangular, tapering to pointed tip on anterior margin (
<figureCitation id="FEB98DF2FFC30273FC640DACFBC6FA85" box="[988,1070,1266,1291]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
A, 3A), broadening posteriorly, nuchal organs narrow slits on posterior margin of prostomium, inconspicuous, not pigmented; eyes absent. Peristomium as long as wide, with two annuli surmounted by longitudinal dorsal crest (
<figureCitation id="FEB98DF2FFC30273FB2A0C64FB31FADD" box="[1170,1241,1338,1363]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
A). Paired dorsal tentacles on posterior margin of peristomium, with first pair of branchiae located immediately posterior to tentacles on peristomium (
<figureCitation id="FEB98DF2FFC30273FEEE0CDCFE71FA15" box="[342,409,1410,1435]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
A); second pair of branchiae on posterior margin of setiger 1, dorsal to notosetae. Branchiae present on most segments through middle of body.
</paragraph>
<paragraph id="663D9177FFC30273FF7F0C95FE77F935" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
Noto- and neuropodia with setal fascicles arising close together; anterior parapodia with spreading fascicles of about 56 capillary setae in notopodia and 67 capillaries in neuropodia; middle body segments with about 910 capillaries in notopodia and
<quantity id="A17A3C92FFC30273FE670F4CFDC1F9A5" box="[479,553,1554,1579]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="4" pageNumber="505" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">56 in</quantity>
neuropodia; far posterior notopodia with 45 narrow capillaries; neuropodia with 34 capillaries and 13 short, curved, spines in posteriormost 45 setigers (
<figureCitation id="FEB98DF2FFC30273FB840F68FB6AF9C1" box="[1084,1154,1590,1615]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
C). Neuroacicular spines initially as straight pointed spine (
<figureCitation id="FEB98DF2FFC30273FDAA0F04FDBCF9FD" box="[530,596,1626,1651]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
D) transitioning over next four setigers to blunt-tipped spine (
<figureCitation id="FEB98DF2FFC30273FABA0F04FAADF9FD" box="[1282,1349,1626,1651]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
E), then spines with knob-like tips (
<figureCitation id="FEB98DF2FFC30273FE7F0F20FDFDF919" box="[455,533,1662,1687]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
FG). On
<typeStatus id="B9392FD5FFC30273FD3F0F20FD01F919" box="[647,745,1662,1687]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
of 115 setigers, neuroacicular spines limited to posteriormost five setigers (111115).
</paragraph>
<paragraph id="663D9177FFC30273FF7F0F98FD53F8E1" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0F98FE57F951" bold="true" box="[199,447,1734,1759]" pageId="4" pageNumber="505">Methyl Green Stain.</emphasis>
Most of body staining initially, tip of prostomium and dorsal peristomial crest clear; stain retained laterally in irregular patches on sides of peristomium and intersegmentally on dorsum, between parapodia, and mid-ventrally on thoracic region where ventral ridges are located sometimes producing a line of paired spots along the venter. The most conspicuous stain is present intersegmentally between the parapodia; no distinct pattern on prostomium and peristomium.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30270FF7F0E24FD7FFD7E" lastPageId="7" lastPageNumber="508" pageId="4" pageNumber="505" type="discussion">
<paragraph id="663D9177FFC30273FF7F0E24FA65F855" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0E24FED3F81D" bold="true" box="[199,315,1914,1939]" pageId="4" pageNumber="505">Remarks.</emphasis>
Because of the expanded posterior end on the
<typeStatus id="B9392FD5FFC30273FCD30E24FC25F81D" box="[875,973,1914,1939]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A182EAF4FFC30273FC420E22FB84F81D" box="[1018,1132,1916,1939]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="505" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC30273FC420E22FB84F81D" box="[1018,1132,1916,1939]" italics="true" pageId="4" pageNumber="505">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC30273FBC10E24FB53F81C" box="[1145,1211,1914,1939]" pageId="4" pageNumber="505" rank="species">
<emphasis id="54F64D65FFC30273FBC10E24FB5EF81D" bold="true" box="[1145,1206,1914,1939]" pageId="4" pageNumber="505">n. sp</emphasis>
.
</taxonomicNameLabel>
, the specimen was initially thought to be a species of
<taxonomicName id="A182EAF4FFC30273FD9A0EC0FD54F839" box="[546,700,1950,1975]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="505" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC30273FD9A0EC0FD54F839" box="[546,700,1950,1975]" italics="true" pageId="4" pageNumber="505">Aphelochaeta</emphasis>
</taxonomicName>
. However, careful inspection of the posterior parapodia revealed the presence of short, curved, blunt-tipped spines ventral to the capillaries in the posteriormost five neuropodia.
</paragraph>
<caption id="32FDC1FFFFC20272FF2F0E03FF05F860" httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="5" pageNumber="506" targetBox="[232,1354,201,1840]" targetPageId="5">
<paragraph id="663D9177FFC20272FF2F0E03FF05F860" blockId="5.[151,1436,1885,2030]" pageId="5" pageNumber="506">
<emphasis id="54F64D65FFC20272FF2F0E03FEFAF8FD" bold="true" box="[151,274,1885,1907]" pageId="5" pageNumber="506">FIGURE 2.</emphasis>
<taxonomicName id="A182EAF4FFC20272FEA00E03FE59F8FA" box="[280,433,1885,1908]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="506" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC20272FEA00E03FE59F8FA" box="[280,433,1885,1908]" italics="true" pageId="5" pageNumber="506">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC20272FE0F0E00FE18F8FD" box="[439,496,1886,1907]" pageId="5" pageNumber="506" rank="species">
<emphasis id="54F64D65FFC20272FE0F0E00FE18F8FD" bold="true" box="[439,496,1886,1907]" pageId="5" pageNumber="506">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; DG, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle.
</paragraph>
</caption>
<caption id="32FDC1FFFFC10271FF2F0E54FD6DF8F5" httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="6" pageNumber="507" targetBox="[157,1428,199,1777]" targetPageId="6">
<paragraph id="663D9177FFC10271FF2F0E54FD6DF8F5" blockId="6.[151,1436,1802,1915]" pageId="6" pageNumber="507">
<emphasis id="54F64D65FFC10271FF2F0E54FEE6F891" bold="true" box="[151,270,1802,1824]" pageId="6" pageNumber="507">FIGURE 3</emphasis>
.
<taxonomicName id="A182EAF4FFC10271FEA40E54FE5FF8AE" box="[284,439,1802,1824]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="507" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC10271FEA40E54FE5FF8AE" box="[284,439,1802,1824]" italics="true" pageId="6" pageNumber="507">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC10271FE070E54FE11F891" box="[447,505,1802,1823]" pageId="6" pageNumber="507" rank="species">
<emphasis id="54F64D65FFC10271FE070E54FE11F891" bold="true" box="[447,505,1802,1823]" pageId="6" pageNumber="507">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view.
<taxonomicName id="A182EAF4FFC10271FC4C0E76FB72F8B0" box="[1012,1178,1832,1854]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="507" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFC10271FC4C0E76FB72F8B0" box="[1012,1178,1832,1854]" italics="true" pageId="6" pageNumber="507">Tharyx robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC10271FB1A0E77FB35F8B0" box="[1186,1245,1833,1854]" pageId="6" pageNumber="507" rank="species">
<emphasis id="54F64D65FFC10271FB1A0E77FB35F8B0" bold="true" box="[1186,1245,1833,1854]" pageId="6" pageNumber="507">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype (SMNH 8756): D, entire animal in ventral view. AB stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium.
</paragraph>
</caption>
<paragraph id="663D9177FFC00270FF2F09C9FD30FE5E" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
These spines transition from curved pointed setae among capillaries to short curved blunt-tipped spines with knoblike tips in the very last setigers. No spines were observed in the notopodia. Because the neuropodial spines are limited to the very posteriormost setigers, they will not be observed in fragmented specimens. Therefore other characters will be required in order to identify the species. Other features of the body are similar to related species of
<taxonomicName id="A182EAF4FFC00270FF0E0879FEEDFECE" box="[182,261,295,320]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FF0E0879FEEDFECE" box="[182,261,295,320]" italics="true" pageId="7" pageNumber="508">Tharyx</emphasis>
</taxonomicName>
. These include the raised crest on the peristomium, the occurrence of lateral black pigment spots on the peristomium; the position of the paired dorsal tentacles on the posterior margin of the peristomium followed by the first pair of branchiae also on the peristomium and the generally elongated form of the body. In addition, the methyl green staining pattern in conjunction with the paired peristomial pigment spots readily differentiates this species from other cirratulids observed in these collections.
</paragraph>
<paragraph id="663D9177FFC00270FF7F0882FD7FFD7E" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
<taxonomicName id="A182EAF4FFC00270FF7F0882FE99FE7A" box="[199,369,476,501]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FF7F0882FE99FE7A" box="[199,369,476,501]" italics="true" pageId="7" pageNumber="508">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FEC10882FE50FE7B" box="[377,440,476,501]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FEC10882FE50FE7B" bold="true" box="[377,440,476,501]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
differs from all other species of the genus in having an expanded posterior end and with the neuropodial spines limited to the five posteriormost setigers. The expanded posterior end suggests a close relationship of
<taxonomicName id="A182EAF4FFC00270FEFC0B7BFE5CFDB2" box="[324,436,549,572]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FEFC0B7BFE5CFDB2" box="[324,436,549,572]" italics="true" pageId="7" pageNumber="508">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FE050B7AFE16FDB3" box="[445,510,548,573]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FE050B7AFE16FDB3" bold="true" box="[445,510,548,573]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
with species of
<taxonomicName id="A182EAF4FFC00270FD030B7AFCBDFDB3" box="[699,853,548,573]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FD030B7AFCBDFDB3" box="[699,853,548,573]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
also having this character. Among eight species of
<taxonomicName id="A182EAF4FFC00270FF2F0B19FEDAFDEE" box="[151,306,583,608]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FF2F0B19FEDAFDEE" box="[151,306,583,608]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
described by
<bibRefCitation id="0213EC86FFC00270FE6C0B19FD81FDEE" author="Blake" box="[468,617,583,608]" pageId="7" pageNumber="508" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">Blake (1996)</bibRefCitation>
from the eastern Pacific, six species had the posterior end expanded in some form. Of these, none had the first pair of branchiae positioned immediately posterior to the dorsal tentacle on the peristomium as is typical for species of
<taxonomicName id="A182EAF4FFC00270FD310BD1FD30FD26" box="[649,728,655,680]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FD310BD1FD30FD26" box="[649,728,655,680]" italics="true" pageId="7" pageNumber="508">Tharyx</emphasis>
</taxonomicName>
. When branchiae were present on the peristomium, they were positioned lateral to the tentacles. There are no described species of
<taxonomicName id="A182EAF4FFC00270FC2C0BEAFBC6FD43" box="[916,1070,692,717]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FC2C0BEAFBC6FD43" box="[916,1070,692,717]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
from European waters that have characters similar to those of
<taxonomicName id="A182EAF4FFC00270FE590B87FDA7FD7E" box="[481,591,729,752]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FE590B87FDA7FD7E" box="[481,591,729,752]" italics="true" pageId="7" pageNumber="508">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FDE00B89FD7FFD7E" box="[600,663,727,752]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FDE00B89FD7FFD7E" bold="true" box="[600,663,727,752]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC00270FF7F0BA2FC6FFCB6" pageId="7" pageNumber="508" type="etymology">
<paragraph id="663D9177FFC00270FF7F0BA2FC6FFCB6" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF7F0BA2FEA6FC9B" bold="true" box="[199,334,764,789]" pageId="7" pageNumber="508">Etymology.</emphasis>
This species is named for the late Dr. Mary Elizabeth Petersen, in recognition of her numerous contributions to polychaete systematics and research on cirratulids.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC00270FF7F0A1AFD3EFA47" pageId="7" pageNumber="508" type="biology_ecology">
<paragraph id="663D9177FFC00270FF7F0A1AFDD8FB43" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF7F0A1AFECFFCD3" bold="true" box="[199,295,836,861]" pageId="7" pageNumber="508">Biology.</emphasis>
The specimens of
<taxonomicName id="A182EAF4FFC00270FE460A1AFD42FCD2" box="[510,682,836,861]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FE460A1AFD42FCD2" box="[510,682,836,861]" italics="true" pageId="7" pageNumber="508">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FD0A0A1AFD1AFCD3" box="[690,754,836,861]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FD0A0A1AFD1AFCD3" bold="true" box="[690,754,836,861]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
were collected by one us (PG) in connection with a benthic macrofauna mapping project in sediments in and outside eelgrass (
<taxonomicName id="A182EAF4FFC00270FC310A37FBDFFC0E" box="[905,1079,873,896]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="508" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="54F64D65FFC00270FC310A37FBDFFC0E" box="[905,1079,873,896]" italics="true" pageId="7" pageNumber="508">Zostera marina</emphasis>
</taxonomicName>
) beds near Varberg on the west coast of
<collectingCountry id="1E95D1E7FFC00270FF4E0AD2FEBAFC2B" box="[246,338,908,933]" name="Sweden" pageId="7" pageNumber="508">Sweden</collectingCountry>
. Salinity and temperature variation in this shallow area is high (2030 PSU; &lt;0°C to&gt;25°C). The worms occurred in four of 30 quantitative haps-corer samples, calculated to each contain 160400 ind/m2 with a biomass of &lt;
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/m2 (variation in specimen length was
<quantity id="A17A3C92FFC00270FCEF0A87FC21FC7C" box="[855,969,985,1010]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="7" pageNumber="508" unit="mm" value="9.0" valueMax="12.0" valueMin="6.0">612 mm</quantity>
). These samples were taken at
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depth in coarse to fine sand with a low organic content (about 12 % loss on ignition) and almost no vegetation, outside or in the margin of the eelgrass bed. The associated fauna were relatively scarce in most samples (612 species and 3,093 ind/m2) and were dominated by
<taxonomicName id="A182EAF4FFC00270FD450D16FB34FBEF" authority="O.F. Muller, 1776" authorityName="O.F. Muller" authorityYear="1776" box="[765,1244,1096,1121]" class="Polychaeta" family="Nereididae" genus="Hediste" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="diversicolor">
<emphasis id="54F64D65FFC00270FD450D16FC0FFBEF" box="[765,999,1096,1121]" italics="true" pageId="7" pageNumber="508">Hediste diversicolor</emphasis>
(
<collectingCountry id="1E95D1E7FFC00270FC460D17FBFCFBEF" box="[1022,1044,1097,1121]" name="Iceland" pageId="7" pageNumber="508">O</collectingCountry>
.F. Müller, 1776)
</taxonomicName>
,
<taxonomicName id="A182EAF4FFC00270FB480D17FED1FB0B" authority="Pennant, 1777" authorityName="Pennant" authorityYear="1777" class="Gastropoda" family="Hydrobiidae" genus="Peringia" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="ulvae">
<emphasis id="54F64D65FFC00270FB480D17FA74FBEF" box="[1264,1436,1096,1121]" italics="true" pageId="7" pageNumber="508">Peringia ulvae</emphasis>
Pennant, 1777
</taxonomicName>
,
<taxonomicName id="A182EAF4FFC00270FEF00D32FD36FB0B" authority="Udekem, 1855" authorityName="Udekem" authorityYear="1855" box="[328,734,1132,1157]" class="Clitellata" family="Tubificidae" genus="Tubificoides" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="benedii">
<emphasis id="54F64D65FFC00270FEF00D32FDC3FB0B" box="[328,555,1132,1157]" italics="true" pageId="7" pageNumber="508">Tubificoides benedii</emphasis>
Udekem, 1855
</taxonomicName>
, and
<taxonomicName id="A182EAF4FFC00270FCA40D32FB3CFB0B" authority="O.F. Muller, 1776" authorityName="O.F. Muller" authorityYear="1776" box="[796,1236,1132,1157]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="armiger">
<emphasis id="54F64D65FFC00270FCA40D32FC03FB0A" box="[796,1003,1132,1157]" italics="true" pageId="7" pageNumber="508">Scoloplos armiger</emphasis>
(
<collectingCountry id="1E95D1E7FFC00270FC440D33FBFAFB0B" box="[1020,1042,1133,1157]" name="Iceland" pageId="7" pageNumber="508">O</collectingCountry>
.F. Müller, 1776)
</taxonomicName>
. The fauna in the eelgrass beds was richer and more diverse in most samples, with a representation of polychaetes and several species of crustaceans and molluscs.
</paragraph>
<paragraph id="663D9177FFC00270FF7F0D86FD3EFA47" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
Both
<taxonomicName id="A182EAF4FFC00270FEBE0D86FE5AFB7E" box="[262,434,1240,1265]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae">
<emphasis id="54F64D65FFC00270FEBE0D86FE5AFB7E" box="[262,434,1240,1265]" italics="true" pageId="7" pageNumber="508">Tharyx maryae</emphasis>
</taxonomicName>
<emphasis id="54F64D65FFC00270FE030D86FE1CFB7F" bold="true" box="[443,500,1240,1265]" pageId="7" pageNumber="508">n. sp</emphasis>
and
<taxonomicName id="A182EAF4FFC00270FD970D87FD43FB7F" box="[559,683,1240,1265]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFC00270FD970D87FD43FB7F" box="[559,683,1240,1265]" italics="true" pageId="7" pageNumber="508">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FD0C0D86FD1DFB7F" box="[692,757,1240,1265]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FD0C0D86FD1DFB7F" bold="true" box="[692,757,1240,1265]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
(described below) were found in the same sample together with an anterior fragment of
<taxonomicName id="A182EAF4FFC00270FE6F0DA2FD99FA9B" box="[471,625,1276,1301]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FE6F0DA2FD99FA9B" box="[471,625,1276,1301]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
sp., also likely new to science. These specimens were taken in relatively fine sediment with higher organic content outside the eel grass beds where small patches of the algae
<taxonomicName id="A182EAF4FFC00270FAAD0C7FFA8EFAB6" box="[1301,1382,1313,1336]" class="Liliopsida" family="Ruppiaceae" genus="Ruppia" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="508" phylum="Tracheophyta" rank="genus">
<emphasis id="54F64D65FFC00270FAAD0C7FFA8EFAB6" box="[1301,1382,1313,1336]" italics="true" pageId="7" pageNumber="508">Ruppia</emphasis>
</taxonomicName>
spp. and
<taxonomicName id="A182EAF4FFC00270FF740C1BFD85FAD3" authority="Linnaeus, 1753" authorityName="Linnaeus" authorityYear="1753" box="[204,621,1348,1373]" class="Phaeophyceae" family="Fucaceae" genus="Fucus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fucales" pageId="7" pageNumber="508" phylum="Heterokontophyta" rank="species" species="vesiculosus">
<emphasis id="54F64D65FFC00270FF740C1BFE74FAD3" box="[204,412,1348,1373]" italics="true" pageId="7" pageNumber="508">Fucus vesiculosus</emphasis>
(Linnaeus, 1753)
</taxonomicName>
were found; however, cirratulids were recorded in sediments almost entirely free of vegetation in three of the four samples. These results suggest that
<taxonomicName id="A182EAF4FFC00270FB890C37FB49FA0E" box="[1073,1185,1385,1408]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FB890C37FB49FA0E" box="[1073,1185,1385,1408]" italics="true" pageId="7" pageNumber="508">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FB110C36FB00FA0F" box="[1193,1256,1384,1409]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FB110C36FB00FA0F" bold="true" box="[1193,1256,1384,1409]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="A182EAF4FFC00270FA980C37FA73FA0F" box="[1312,1435,1384,1409]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFC00270FA980C37FA73FA0F" box="[1312,1435,1384,1409]" italics="true" pageId="7" pageNumber="508">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FF2F0CD2FF30FA2B" box="[151,216,1420,1445]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FF2F0CD2FF30FA2B" bold="true" box="[151,216,1420,1445]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
are very tolerant of variations in salinity and temperature and prefer relatively coarse sand with almost no vegetation and low competition from other species.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC00270FF7F0C8AFC03FA62" box="[199,1003,1492,1517]" pageId="7" pageNumber="508" type="distribution">
<paragraph id="663D9177FFC00270FF7F0C8AFC03FA62" blockId="7.[151,1437,151,1517]" box="[199,1003,1492,1517]" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF7F0C8AFE88FA63" bold="true" box="[199,352,1492,1517]" pageId="7" pageNumber="508">Distribution.</emphasis>
Known only from the west coast of
<collectingCountry id="1E95D1E7FFC00270FD410C8AFCBAFA63" box="[761,850,1492,1517]" name="Sweden" pageId="7" pageNumber="508">Sweden</collectingCountry>
in low water.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,370 @@
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<mods:title id="75C11C3F39A39523AE959CBC396913E5">Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)</mods:title>
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<heading id="3D75261BFFC50275FF2F09C6FDBBFF3C" bold="true" box="[151,595,152,178]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<taxonomicName id="A182EAF4FFC50275FF2F09C6FDBBFF3C" ID-CoL="564M4" authority="Southern, 1914" authorityName="Southern" authorityYear="1914" box="[151,595,152,178]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FF2F09C6FDBBFF3C" bold="true" box="[151,595,152,178]" pageId="2" pageNumber="503">
<emphasis id="54F64D65FFC50275FF2F09C6FE93FF3C" bold="true" box="[151,379,152,178]" italics="true" pageId="2" pageNumber="503">Tharyx killariensis</emphasis>
(
<bibRefCitation id="0213EC86FFC50275FE3309C6FDA3FF3C" author="Southern" box="[395,587,152,178]" pageId="2" pageNumber="503" refString="Southern, R. (1914) Archiannelida and Polychaeta. In Clare Island Survey Part 47. Proceedings of the Royal Irish Academy, 31, 47 - 160, pls. 1 - 15." type="journal article" year="1914">Southern, 1914</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="663D9177FFC50275FF2F09E4FF1DFF5D" blockId="2.[151,595,152,211]" box="[151,245,186,211]" pageId="2" pageNumber="503">
<figureCitation id="FEB98DF2FFC50275FF2F09E4FF1DFF5D" box="[151,245,186,211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Figure 1</figureCitation>
</paragraph>
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<subSubSection id="2E98C2FCFFC50275FF2F085CFC3CFE1D" pageId="2" pageNumber="503" type="reference_group">
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<taxonomicName id="A182EAF4FFC50275FF2F085CFDB2FE96" ID-CoL="TF26" authority="Southern, 1914: 113" authorityName="Southern" authorityPageNumber="113" authorityYear="1914" box="[151,602,258,280]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FF2F085CFE6EFE96" box="[151,390,258,280]" italics="true" pageId="2" pageNumber="503">Chaetozone killariensis</emphasis>
<treatmentCitation id="E723B766FFC50275FE34085CFDB2FE96" author="Southern" box="[396,602,258,280]" page="113" pageId="2" pageNumber="503" year="1914">
<bibRefCitation id="0213EC86FFC50275FE34085CFDC3FE96" author="Southern" box="[396,555,258,280]" pageId="2" pageNumber="503" refString="Southern, R. (1914) Archiannelida and Polychaeta. In Clare Island Survey Part 47. Proceedings of the Royal Irish Academy, 31, 47 - 160, pls. 1 - 15." type="journal article" year="1914">Southern, 1914</bibRefCitation>
: 113
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115, Plate XII, figs. 27 AD;
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<bibRefCitation id="0213EC86FFC50275FC2D085CFBDBFE96" author="McIntosh" box="[917,1075,258,280]" pageId="2" pageNumber="503" refString="McIntosh, C. C. A. (1923) A monograph of the British annelids. Vol. 4. Part 2. Polychaeta: Sabellidae to Serpulidae. With additions to the British marine Polychaeta during the publication of the monograph. Ray Society, London, xii + 278 pp. [pp. i - xii &amp; pp. 251 - 528, pls. 115 - 138, text figs. 151 - 174.]" type="book chapter" year="1923">McIntosh 1923</bibRefCitation>
: 491
</treatmentCitation>
492, Plate CXXXVI, Fig. 13;
<treatmentCitation id="E723B766FFC50275FF7F087FFE17FEB9" author="Hartmann-Schroder" box="[199,511,289,311]" page="355" pageId="2" pageNumber="503" year="1971">
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: 355
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.
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</paragraph>
<paragraph id="663D9177FFC50275FF2F081FFD07FED8" blockId="2.[151,1436,258,403]" box="[151,751,320,342]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="4692B659FFC50275FF2F081FFD07FED8" box="[151,751,320,342]" pageId="2" pageNumber="503">
<taxonomicName id="A182EAF4FFC50275FF2F081FFDAAFED8" authority="Fauvel 1927: 97" authorityName="Fauvel" authorityPageNumber="97" authorityYear="1927" box="[151,578,320,342]" class="Polychaeta" family="Cirratulidae" genus="Heterocirrus" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FF2F081FFE67FED8" box="[151,399,320,342]" italics="true" pageId="2" pageNumber="503">Heterocirrus killariensis</emphasis>
:
<treatmentCitation id="E723B766FFC50275FE25081EFDAAFED8" author="Fauvel" box="[413,578,320,342]" page="97" pageId="2" pageNumber="503" year="1927">
<bibRefCitation id="0213EC86FFC50275FE25081EFDF6FED8" author="Fauvel" box="[413,542,320,342]" pageId="2" pageNumber="503" refString="Fauvel, P. (1927) Polychetes sedentaires. Addenda aux errantes, archiannelides, myzostomaires. Faune de France, 16, 1 - 494." type="journal article" year="1927">Fauvel 1927</bibRefCitation>
: 97
</treatmentCitation>
</taxonomicName>
98, Fig. 86 dh.
</treatmentCitationGroup>
</paragraph>
<paragraph id="663D9177FFC50275FF2F0800FD89FEFA" blockId="2.[151,1436,258,403]" box="[151,609,350,372]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="4692B659FFC50275FF2F0800FD89FEFA" box="[151,609,350,372]" pageId="2" pageNumber="503">
<taxonomicName id="A182EAF4FFC50275FF2F0800FDB5FEFA" ID-CoL="RX7X" authority="Hartman 1959: 401" authorityName="Hartman" authorityPageNumber="401" authorityYear="1959" box="[151,605,350,372]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FF2F0800FE60FEFA" box="[151,392,350,372]" italics="true" pageId="2" pageNumber="503">Caulleriella killariensis</emphasis>
:
<treatmentCitation id="E723B766FFC50275FE2D0801FDB5FEFA" author="Hartman" box="[405,605,350,372]" page="401" pageId="2" pageNumber="503" year="1959">
<bibRefCitation id="0213EC86FFC50275FE2D0801FDC2FEFA" author="Hartman" box="[405,554,350,372]" pageId="2" pageNumber="503" refString="Hartman, O. (1959) Catalogue of the Polychaetous Annelids of the World. Allan Hancock Foundation Occasional Paper, 23, 1 - 628." type="journal article" year="1959">Hartman 1959</bibRefCitation>
: 401
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="663D9177FFC50275FF2F0823FC3CFE1D" blockId="2.[151,1436,258,403]" box="[151,980,381,403]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="4692B659FFC50275FF2F0823FC3CFE1D" box="[151,980,381,403]" pageId="2" pageNumber="503">
<taxonomicName id="A182EAF4FFC50275FF2F0823FE11FE1D" ID-CoL="564M4" authority="Blake 1991:19" authorityName="Blake" authorityPageNumber="19" authorityYear="1991" box="[151,505,381,403]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FF2F0823FEBDFE1D" box="[151,341,381,403]" italics="true" pageId="2" pageNumber="503">Tharyx killariensis</emphasis>
:
<treatmentCitation id="E723B766FFC50275FEDA0823FE11FE1D" author="Blake" box="[354,505,381,403]" page="19" pageId="2" pageNumber="503" year="1991">
<bibRefCitation id="0213EC86FFC50275FEDA0823FE34FE1D" author="Blake" box="[354,476,381,403]" pageId="2" pageNumber="503" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake 1991</bibRefCitation>
:19
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="E723B766FFC50275FDBC0823FCD1FE1D" author="Hartmann-Schroder" box="[516,825,381,403]" page="399" pageId="2" pageNumber="503" year="1996">
<bibRefCitation id="0213EC86FFC50275FDBC0823FCE1FE1D" author="Hartmann-Schroder" box="[516,777,381,403]" pageId="2" pageNumber="503" refString="Hartmann-Schroder, G. (1996) Annelida, Borstenwurmer, Polychaeta. Second Edition. Die Tierwelt Deutschlands 2, neubearbeitete Auflage, 58, 1 - 648." type="journal article" year="1996">Hartmann-Schröder 1996</bibRefCitation>
: 399
</treatmentCitation>
400, Fig. 192.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC50275FF2F08E1FBCDFDE6" pageId="2" pageNumber="503" type="materials_examined">
<paragraph id="663D9177FFC50275FF2F08E1FC2FFDAE" blockId="2.[151,1437,447,2021]" pageId="2" pageNumber="503">
<emphasis id="54F64D65FFC50275FF2F08E1FE68FE56" bold="true" box="[151,384,447,472]" pageId="2" pageNumber="503">Material examined.</emphasis>
<collectingCountry id="1E95D1E7FFC50275FE3E089EFDECFE56" box="[390,516,448,472]" name="Ireland" pageId="2" pageNumber="503">IRELAND</collectingCountry>
, Killary Harbor, Station W. 86, 1914, dredged in rich black mud at 7 fathoms (
<quantity id="A17A3C92FFC50275FAD2089EFF58FE72" metricMagnitude="1" metricUnit="m" metricValue="1.28" pageId="2" pageNumber="503" unit="m" value="12.8">12.8 m</quantity>
),
<typeStatus id="B9392FD5FFC50275FF7208BAFEDDFE73" box="[202,309,484,509]" pageId="2" pageNumber="503" type="lectotype">
<emphasis id="54F64D65FFC50275FF7208BAFEDDFE73" bold="true" box="[202,309,484,509]" pageId="2" pageNumber="503">lectotype</emphasis>
</typeStatus>
(
<collectionCode id="009309B2FFC50275FEF208BAFE43FE72" box="[330,427,484,508]" httpUri="http://grbio.org/cool/hz2c-tkpx" name="National Museum of Ireland" pageId="2" pageNumber="503">NMINH</collectionCode>
1914.348). Note: the label suggests two specimens (
<typeStatus id="B9392FD5FFC50275FB9008BAFB65FE72" box="[1064,1165,484,508]" pageId="2" pageNumber="503" type="syntype">syntypes</typeStatus>
) in vial, but these are fragmented parts of the same specimen, here designated as the
<typeStatus id="B9392FD5FFC50275FCE00B59FC2BFDAE" box="[856,963,519,544]" pageId="2" pageNumber="503" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="663D9177FFC50275FF7F0B72FBCDFDE6" blockId="2.[151,1437,447,2021]" pageId="2" pageNumber="503">
<emphasis id="54F64D65FFC50275FF7F0B72FE55FDCB" bold="true" box="[199,445,556,581]" pageId="2" pageNumber="503">Additional material.</emphasis>
Same station, collected in 1909 from Killary Harbor. Two specimens in vial labeled
<typeStatus id="B9392FD5FFC50275FF2F0B0EFF11FDE6" box="[151,249,592,616]" pageId="2" pageNumber="503" type="syntype">syntypes</typeStatus>
are fragments. No head or posterior ends available (
<collectionCode id="009309B2FFC50275FCFB0B0EFC4CFDE6" box="[835,932,592,616]" httpUri="http://grbio.org/cool/hz2c-tkpx" name="National Museum of Ireland" pageId="2" pageNumber="503">NMINH</collectionCode>
1909.151).
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC50275FF7F0B2AFD96FA46" pageId="2" pageNumber="503" type="description">
<paragraph id="663D9177FFC50275FF7F0B2AFA7DFC06" blockId="2.[151,1437,447,2021]" pageId="2" pageNumber="503">
<emphasis id="54F64D65FFC50275FF7F0B2AFEB1FD03" bold="true" box="[199,345,628,653]" pageId="2" pageNumber="503">Description.</emphasis>
<typeStatus id="B9392FD5FFC50275FEDB0B2BFE3BFD03" box="[355,467,629,653]" pageId="2" pageNumber="503" type="lectotype">Lectotype</typeStatus>
composed of three large fragments and two smaller fragments clearly all from the same specimen and with evidence of prior dissection; collectively about
<quantity id="A17A3C92FFC50275FC700BC6FBFFFD21" box="[968,1047,664,688]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="503" unit="mm" value="10.0">10 mm</quantity>
long with 77 setigerous segments; anterior fragment
<quantity id="A17A3C92FFC50275FED90BE2FE4BFD5A" box="[353,419,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="503" unit="mm" value="5.0">5 mm</quantity>
long with 47 setigers. Largest specimen recorded by
<bibRefCitation id="0213EC86FFC50275FC430BE2FB5AFD5A" author="Southern" box="[1019,1202,700,725]" pageId="2" pageNumber="503" refString="Southern, R. (1914) Archiannelida and Polychaeta. In Clare Island Survey Part 47. Proceedings of the Royal Irish Academy, 31, 47 - 160, pls. 1 - 15." type="journal article" year="1914">Southern (1914)</bibRefCitation>
<quantity id="A17A3C92FFC50275FB010BE2FAEFFD5A" box="[1209,1287,700,724]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="503" unit="mm" value="11.0">11 mm</quantity>
long with 84 setigerous segments. Body not pigmented, light tan in alcohol. Body with dorsum of anterior and middle segments elevated above parapodia, forming distinct ridge along body (
<figureCitation id="FEB98DF2FFC50275FCE70A5AFC48FC92" box="[863,928,772,797]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
AB); first 30 segments narrower and more crowded than middle segments, middle segments not moniliform; far posterior segments again becoming narrow. Venter somewhat flattened throughout body, with narrow, weakly developed mid-ventral ridge formed of raised pads located in middle of each segmental annulus. Pygidium with dorsal anus and flattened ventral lobe (
<figureCitation id="FEB98DF2FFC50275FA880A31FA98FC06" box="[1328,1392,879,904]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
C).
</paragraph>
<paragraph id="663D9177FFC50275FF7F0ACAFEECFBEE" blockId="2.[151,1437,447,2021]" pageId="2" pageNumber="503">
Prostomium triangular, pointed on anterior margin; eyes absent (
<figureCitation id="FEB98DF2FFC50275FC090ACAFC1AFC22" box="[945,1010,916,941]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
AB); nuchal organ a narrow slit on posterior margin. Peristomium about as wide as long, annulations not visible dorsally, but with 23 weak divisions ventrally (
<figureCitation id="FEB98DF2FFC50275FEB40A82FEA6FC7A" box="[268,334,988,1013]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
A). Paired dorsal tentacles arising from posterior margin of peristomium, first pair of branchiae arising on peristomium posterior and slightly lateral to tentacles, but in line with subsequent segmental branchiae (
<figureCitation id="FEB98DF2FFC50275FF270D7AFF35FBB2" box="[159,221,1060,1085]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
AB); second pair of branchiae on setiger 1, subsequent branchiae present on at least another 2025 setigers (
<figureCitation id="FEB98DF2FFC50275FF270D19FF37FBEE" box="[159,223,1095,1120]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
B).
</paragraph>
<paragraph id="663D9177FFC50275FF7F0D32FCECFA2B" blockId="2.[151,1437,447,2021]" pageId="2" pageNumber="503">
Parapodia low mounds from which setal fascicles arise; noto- and neuropodial setal fascicles positioned close to one another throughout. Setae all capillaries for first 50 or so setigers, with 57 capillaries in noto- and neuropodia.
<bibRefCitation id="0213EC86FFC50275FE9B0DEAFE31FB42" author="Southern" box="[291,473,1204,1229]" pageId="2" pageNumber="503" refString="Southern, R. (1914) Archiannelida and Polychaeta. In Clare Island Survey Part 47. Proceedings of the Royal Irish Academy, 31, 47 - 160, pls. 1 - 15." type="journal article" year="1914">Southern (1914)</bibRefCitation>
reported spines (crotchets) from setiger
<quantity id="A17A3C92FFC50275FC270DEAFC30FB43" box="[927,984,1204,1229]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="2" pageNumber="503" unit="in" value="56.0">56 in</quantity>
neuropodia and setiger
<quantity id="A17A3C92FFC50275FB5D0DEAFAF6FB43" box="[1253,1310,1204,1229]" metricMagnitude="0" metricUnit="m" metricValue="1.5493999999999999" pageId="2" pageNumber="503" unit="in" value="61.0">61 in</quantity>
notopodia;
<typeStatus id="B9392FD5FFC50275FF2F0D89FF16FB7E" box="[151,254,1239,1264]" pageId="2" pageNumber="503" type="lectotype">lectotype</typeStatus>
similar with neuropodial spines present from setiger 51; notopodial spines first appearing with a single elongate spine among capillaries on setiger 56; posterior setigers with up to two long, narrow blunted spines and 2 3 capillaries in notopodia and 34 short curved spines and 23 long capillaries in neuropodia (
<figureCitation id="FEB98DF2FFC50275FA950C41FA9DFAB6" box="[1325,1397,1311,1336]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
D). Notoacicular spines long, thin, with narrow rounded tip bearing only a single knob (
<figureCitation id="FEB98DF2FFC50275FBFD0C1AFB6CFAD2" box="[1093,1156,1348,1373]" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
E); Neuroacicular spines curved, somewhat geniculate with blunt tip bearing two weakly developed knobs; shortest spines ventralmost (
<figureCitation id="FEB98DF2FFC50275FAD70C39FF49FA2A" captionStart="FIGURE 1" captionStartId="3.[151,250,1723,1745]" captionTargetBox="[162,1411,284,1695]" captionTargetId="figure@3.[151,1436,265,1701]" captionTargetPageId="3" captionText="FIGURE 1. Tharyx killariensis (Southern, 1914). Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="2" pageNumber="503">Fig. 1</figureCitation>
F) grading dorsally into longer and thinner capillaries.
</paragraph>
<paragraph id="663D9177FFC50275FF7F0CF1FD96FA46" blockId="2.[151,1437,447,2021]" box="[199,638,1455,1480]" pageId="2" pageNumber="503">
<emphasis id="54F64D65FFC50275FF7F0CF1FE59FA46" bold="true" box="[199,433,1455,1480]" pageId="2" pageNumber="503">Methyl green Stain.</emphasis>
No stain retained.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC50275FF7F0C8AFA74F94B" pageId="2" pageNumber="503" type="discussion">
<paragraph id="663D9177FFC50275FF7F0C8AFA74F94B" blockId="2.[151,1437,447,2021]" pageId="2" pageNumber="503">
<emphasis id="54F64D65FFC50275FF7F0C8AFED3FA63" bold="true" box="[199,315,1492,1517]" pageId="2" pageNumber="503">Remarks.</emphasis>
The general features originally described by
<bibRefCitation id="0213EC86FFC50275FCDD0C8AFBCAFA62" author="Southern" box="[869,1058,1492,1517]" pageId="2" pageNumber="503" refString="Southern, R. (1914) Archiannelida and Polychaeta. In Clare Island Survey Part 47. Proceedings of the Royal Irish Academy, 31, 47 - 160, pls. 1 - 15." type="journal article" year="1914">Southern (1914)</bibRefCitation>
for
<taxonomicName id="A182EAF4FFC50275FBE60C8AFA84FA63" box="[1118,1388,1492,1517]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FBE60C8AFA84FA63" box="[1118,1388,1492,1517]" italics="true" pageId="2" pageNumber="503">Chaetozone killariensis</emphasis>
</taxonomicName>
are confirmed in the present description, including the nature of body segmentation, prostomium/peristomium, position of the dorsal tentacles, branchiae, and nature of the pygidium. However, Southerns depiction of the spines or crotchets as having two small pointed teeth is not confirmed. The
<typeStatus id="B9392FD5FFC50275FC0C0F61FBF3F9D6" box="[948,1051,1599,1624]" pageId="2" pageNumber="503" type="lectotype">lectotype</typeStatus>
clearly shows that the tips of the noto- and neuropodial spines are blunted with, at best, two low knobs, not teeth. In this respect, this species belongs in the genus
<taxonomicName id="A182EAF4FFC50275FE9E0FD9FE9EF92E" box="[294,374,1671,1696]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC50275FE9E0FD9FE9EF92E" box="[294,374,1671,1696]" italics="true" pageId="2" pageNumber="503">Tharyx</emphasis>
</taxonomicName>
as defined by
<bibRefCitation id="0213EC86FFC50275FDA20FD9FD59F92E" author="Blake" box="[538,689,1671,1696]" pageId="2" pageNumber="503" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
. Further, the setal fascicles of noto- and neuropodia are positioned close to one another instead of being widely separated as in most reported species of
<taxonomicName id="A182EAF4FFC50275FBF20FF2FB39F94B" box="[1098,1233,1708,1733]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC50275FBF20FF2FB39F94B" box="[1098,1233,1708,1733]" italics="true" pageId="2" pageNumber="503">Caulleriella</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0213EC86FFC50275FAB40FF2FA78F94B" author="Blake" box="[1292,1424,1708,1733]" pageId="2" pageNumber="503" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">Blake 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC50274FF7F0F91FCA1FF5A" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="reference_group">
<paragraph id="663D9177FFC50274FF7F0F91FCA1FF5A" blockId="2.[151,1437,447,2021]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<taxonomicName id="A182EAF4FFC50275FF7F0F91FE74F966" ID-CoL="564M4" box="[199,412,1743,1768]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FF7F0F91FE74F966" box="[199,412,1743,1768]" italics="true" pageId="2" pageNumber="503">Tharyx killariensis</emphasis>
</taxonomicName>
is similar to most of the known species of
<taxonomicName id="A182EAF4FFC50275FC280F91FC08F966" ID-CoL="84V7M" box="[912,992,1743,1768]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC50275FC280F91FC08F966" box="[912,992,1743,1768]" italics="true" pageId="2" pageNumber="503">Tharyx</emphasis>
</taxonomicName>
in having spines in both the noto- and neuropodia of posterior setigers. However, the notoacicular spines of
<taxonomicName id="A182EAF4FFC50275FC180FABFBD1F883" ID-CoL="564M4" box="[928,1081,1780,1805]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FC180FABFBD1F883" box="[928,1081,1780,1805]" italics="true" pageId="2" pageNumber="503">T. killariensis</emphasis>
</taxonomicName>
are few in number, narrow, and blunted, with heavier recurved spines limited to the neuropodia; the spines are also accompanied by capillaries. By having different
<typeStatus id="B9392FD5FFC50275FEED0E63FE78F8DB" box="[341,400,1853,1877]" pageId="2" pageNumber="503">types</typeStatus>
of spines in the noto- and neuropodia,
<taxonomicName id="A182EAF4FFC50275FCEF0E63FC19F8DB" ID-CoL="564M4" box="[855,1009,1852,1877]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FCEF0E63FC19F8DB" box="[855,1009,1852,1877]" italics="true" pageId="2" pageNumber="503">T. killariensis</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="A182EAF4FFC50275FAB30E63FEC8F8F6" ID-CoL="564LR" authority="Blake, 2015" authorityName="Blake" authorityYear="2015" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="alaskensis">
<emphasis id="54F64D65FFC50275FAB30E63FA74F8DB" box="[1291,1436,1852,1877]" italics="true" pageId="2" pageNumber="503">T. alaskensis</emphasis>
<bibRefCitation id="0213EC86FFC50275FF2F0E01FEC8F8F6" author="Blake" box="[151,288,1887,1912]" pageId="2" pageNumber="503" refString="Blake, J. A. (2015) New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitzbergen in the Norwegian Arctic. Zootaxa, 3919 (3), 501 - 552. http: // dx. doi. org / 10.11646 / zootaxa. 3919.3.5" type="journal article" year="2015">Blake, 2015</bibRefCitation>
</taxonomicName>
from the Alaskan Arctic. Further, the paired knobs on the tips of the spines are poorly developed in both
<taxonomicName id="A182EAF4FFC50275FF770EDBFE80F813" ID-CoL="564M4" box="[207,360,1924,1949]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FF770EDBFE80F813" box="[207,360,1924,1949]" italics="true" pageId="2" pageNumber="503">T. killariensis</emphasis>
</taxonomicName>
and
<taxonomicName id="A182EAF4FFC50275FE250EDBFDC3F813" ID-CoL="564LR" box="[413,555,1924,1949]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="alaskensis">
<emphasis id="54F64D65FFC50275FE250EDBFDC3F813" box="[413,555,1924,1949]" italics="true" pageId="2" pageNumber="503">T. alaskensis</emphasis>
</taxonomicName>
compared with other species (
<bibRefCitation id="0213EC86FFC50275FCC70EDAFBEBF812" author="Blake" box="[895,1027,1924,1949]" pageId="2" pageNumber="503" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake 1991</bibRefCitation>
,
<bibRefCitation id="0213EC86FFC50275FBB70EDAFBA2F813" author="Blake" box="[1039,1098,1924,1949]" pageId="2" pageNumber="503" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">1996</bibRefCitation>
,
<bibRefCitation id="0213EC86FFC50275FBED0EDAFB7AF813" author="Blake" box="[1109,1170,1924,1949]" pageId="2" pageNumber="503" refString="Blake, J. A. (2015) New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitzbergen in the Norwegian Arctic. Zootaxa, 3919 (3), 501 - 552. http: // dx. doi. org / 10.11646 / zootaxa. 3919.3.5" type="journal article" year="2015">2015</bibRefCitation>
). The two species differ in that capillaries accompany the spines in the posterior noto- and neuropodia of
<taxonomicName id="A182EAF4FFC50275FB8C0EF7FB25F84F" ID-CoL="564M4" box="[1076,1229,1960,1985]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC50275FB8C0EF7FB25F84F" box="[1076,1229,1960,1985]" italics="true" pageId="2" pageNumber="503">T. killariensis</emphasis>
</taxonomicName>
, whereas they are absent in
<taxonomicName id="A182EAF4FFC50275FEBC0E93FE7BF86B" ID-CoL="564LR" box="[260,403,1996,2021]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="alaskensis">
<emphasis id="54F64D65FFC50275FEBC0E93FE7BF86B" box="[260,403,1996,2021]" italics="true" pageId="2" pageNumber="503">T. alaskensis</emphasis>
</taxonomicName>
. Further,
<taxonomicName id="A182EAF4FFC50275FDBA0E93FD7AF86B" ID-CoL="564LR" box="[514,658,1996,2021]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="503" phylum="Annelida" rank="species" species="alaskensis">
<emphasis id="54F64D65FFC50275FDBA0E93FD7AF86B" box="[514,658,1996,2021]" italics="true" pageId="2" pageNumber="503">T. alaskensis</emphasis>
</taxonomicName>
has both the first and second pairs of branchiae located on setiger 1, whereas
<taxonomicName id="A182EAF4FFC40274FF4109C7FE78FF3E" ID-CoL="564M4" box="[249,400,151,176]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="504" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC40274FF4109C7FE78FF3E" box="[249,400,151,176]" italics="true" pageId="3" pageNumber="504">T. killariensis</emphasis>
</taxonomicName>
and other
<taxonomicName id="A182EAF4FFC40274FDBD09C9FDBDFF3E" ID-CoL="84V7M" box="[517,597,151,176]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="504" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC40274FDBD09C9FDBDFF3E" box="[517,597,151,176]" italics="true" pageId="3" pageNumber="504">Tharyx</emphasis>
</taxonomicName>
species have the first pair of branchiae either on the posterior margin of the peristomium or on an achaetous segment anterior to setiger 1.
</paragraph>
</subSubSection>
<caption id="32FDC1FFFFC40274FF2F0FE5FD87F8A2" httpUri="https://zenodo.org/record/232035/files/figure.png" pageId="3" pageNumber="504" targetBox="[162,1411,284,1695]" targetPageId="3">
<paragraph id="663D9177FFC40274FF2F0FE5FD87F8A2" blockId="3.[151,1436,1723,1836]" pageId="3" pageNumber="504">
<emphasis id="54F64D65FFC40274FF2F0FE5FEF0F95E" bold="true" box="[151,280,1723,1745]" pageId="3" pageNumber="504">FIGURE 1.</emphasis>
<taxonomicName id="A182EAF4FFC40274FE9B0FE5FD4AF95F" authority="Southern, 1914" authorityName="Southern" authorityYear="1914" box="[291,674,1723,1745]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="504" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC40274FE9B0FE5FE0FF95F" box="[291,487,1723,1745]" italics="true" pageId="3" pageNumber="504">Tharyx killariensis</emphasis>
(Southern, 1914)
</taxonomicName>
. Lectotype (NMINH 1914.348): A, anterior end, right lateral view; B, anterior end, dorsal view; C, posterior end, dorsal view; D, posterior parapodium; E, capillary and notoacicular spine from posterior notopodium; F, neuroacicular spines from posterior neuropodium. Abbreviations: br, branchiae; nuO, nuchal organ; per, peristomium; pr, prostomium; tn, tentacle.
</paragraph>
</caption>
<subSubSection id="2E98C2FCFFC40273FF7F0E0BFD30FE93" lastPageId="4" lastPageNumber="505" pageId="3" pageNumber="504" type="materials_examined">
<paragraph id="663D9177FFC40273FF7F0E0BFD30FE93" blockId="3.[151,1436,1876,2009]" lastBlockId="4.[151,1437,151,536]" lastPageId="4" lastPageNumber="505" pageId="3" pageNumber="504">
Most published accounts of
<taxonomicName id="A182EAF4FFC40274FDA10E0AFD19F8E3" box="[537,753,1876,1901]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="504" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC40274FDA10E0AFD19F8E3" box="[537,753,1876,1901]" italics="true" pageId="3" pageNumber="504">Tharyx killariensis</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation id="0213EC86FFC40274FC850E0AFC39F8E2" author="Fauvel" box="[829,977,1876,1901]" pageId="3" pageNumber="504" refString="Fauvel, P. (1927) Polychetes sedentaires. Addenda aux errantes, archiannelides, myzostomaires. Faune de France, 16, 1 - 494." type="journal article" year="1927">Fauvel 1927</bibRefCitation>
;
<bibRefCitation id="0213EC86FFC40274FC5A0E0AFAE0F8E2" author="Hartmann-Schroder" box="[994,1288,1876,1901]" pageId="3" pageNumber="504" refString="Hartmann-Schroder, G. (1971) Annelida, Borstenwurmer, Polychaeta. Die Tierwelt Deutschlands, 58, 1 - 594." type="journal article" year="1971">Hartmann-Schröder 1971</bibRefCitation>
,
<bibRefCitation id="0213EC86FFC40274FAA10E0AFABAF8E3" author="Hartmann-Schroder" box="[1305,1362,1876,1901]" pageId="3" pageNumber="504" refString="Hartmann-Schroder, G. (1996) Annelida, Borstenwurmer, Polychaeta. Second Edition. Die Tierwelt Deutschlands 2, neubearbeitete Auflage, 58, 1 - 648." type="journal article" year="1996">1996</bibRefCitation>
) have largely repeated the descriptions and figures of
<bibRefCitation id="0213EC86FFC40274FD040E26FC9DF81E" author="Southern" box="[700,885,1912,1937]" pageId="3" pageNumber="504" refString="Southern, R. (1914) Archiannelida and Polychaeta. In Clare Island Survey Part 47. Proceedings of the Royal Irish Academy, 31, 47 - 160, pls. 1 - 15." type="journal article" year="1914">Southern (1914)</bibRefCitation>
and as such the actual nature of any specimens subsequently collected and reported as this species in the intervening years from different localities cannot be confirmed.
<bibRefCitation id="0213EC86FFC40274FEA30E9EFE33F856" author="McIntosh" box="[283,475,1984,2009]" pageId="3" pageNumber="504" refString="McIntosh, C. C. A. (1923) A monograph of the British annelids. Vol. 4. Part 2. Polychaeta: Sabellidae to Serpulidae. With additions to the British marine Polychaeta during the publication of the monograph. Ray Society, London, xii + 278 pp. [pp. i - xii &amp; pp. 251 - 528, pls. 115 - 138, text figs. 151 - 174.]" type="book chapter" year="1923">McIntosh (1923)</bibRefCitation>
appears to have examined some of Southerns material, but he did not change the description of the spines; his only illustration was of a notopodial capillary. The brief description by
<bibRefCitation id="0213EC86FFC30273FB4F09C9FA74FF3E" author="Eliason" box="[1271,1436,151,176]" pageId="4" pageNumber="505" refString="Eliason, A. (1962) Die Polychaeten der Skagerak-Expedition 1933. Zoologiska Bidrag fran Uppsala, 33, 207 - 293." type="journal article" year="1962">Eliason (1962)</bibRefCitation>
from the Skagerrak appears to be a different species because distinct pigment is present on the dorsolateral margins of the peristomium, neuropodial hooks are first present in the anterior one-third of the body instead of the posterior one-third, and notopodial hooks are entirely absent.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30273FF7F0879FD8DFE5E" pageId="4" pageNumber="505" type="biology_ecology">
<paragraph id="663D9177FFC30273FF7F0879FD8DFE5E" blockId="4.[151,1437,151,536]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0879FEF7FECE" bold="true" box="[199,287,295,320]" pageId="4" pageNumber="505">Biology</emphasis>
. Since the original report from Killary Harbor,
<collectingCountry id="1E95D1E7FFC30273FC860879FC78FECE" box="[830,912,295,320]" name="Ireland" pageId="4" pageNumber="505">Ireland</collectingCountry>
, in shallow subtidal muds,
<taxonomicName id="A182EAF4FFC30273FB700879FA74FECE" box="[1224,1436,295,320]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="505" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC30273FB700879FA74FECE" box="[1224,1436,295,320]" italics="true" pageId="4" pageNumber="505">Tharyx killariensis</emphasis>
</taxonomicName>
has been reported from SW
<collectingCountry id="1E95D1E7FFC30273FE690812FDD9FEEB" box="[465,561,332,357]" name="United Kingdom" pageId="4" pageNumber="505">England</collectingCountry>
, the west coast of
<collectingCountry id="1E95D1E7FFC30273FD470813FCA6FEEA" box="[767,846,333,356]" name="France" pageId="4" pageNumber="505">France</collectingCountry>
, the southern North Sea, and the Wadden Sea as part of various marine benthic community studies. In some recent accounts the species is among the dominant species in low intertidal and shallow subtidal benthic assemblages (
<bibRefCitation id="0213EC86FFC30273FCF608CAFBB5FE22" author="Reiss" box="[846,1117,404,429]" pageId="4" pageNumber="505" refString="Reiss, H. &amp; Kroncke, I. (2001) Spatial and temporal distribution of macrofauna in the Otzumer Balje (East Frisian Wadden Sea, Germany). Marine Biodiversity, 31 (2), 283 - 298." type="journal article" year="2001">Reiss &amp; Kröncke 2001</bibRefCitation>
;
<bibRefCitation id="0213EC86FFC30273FBD408CAFA7FFE22" author="Nehmer" box="[1132,1431,404,429]" pageId="4" pageNumber="505" refString="Nehmer, P. &amp; Kroncke, I. (2003) Macrofaunal communities in the Wichter Ee, a channel system in the East Frisian Wadden Sea. Senckenbergiana maritime, 32, 1 - 10." type="journal article" year="2003">Nehmer &amp; Kröncke 2003</bibRefCitation>
;
<bibRefCitation id="0213EC86FFC30273FF2F08E9FE8CFE5E" author="Callier" box="[151,356,439,464]" pageId="4" pageNumber="505" refString="Callier, M. D., Fletcher, R. L., Thorp, C. H. &amp; Ficher, D. (2009) Macrofaunal community responses to marine-related pollution on the south coast of England and west coast of France. Journal of the Marine Biological Association of the United Kingdom, 89, 19 - 29." type="journal article" year="2009">
Callier
<emphasis id="54F64D65FFC30273FF5208E7FECAFE5E" box="[234,290,439,464]" italics="true" pageId="4" pageNumber="505">et al.</emphasis>
2009
</bibRefCitation>
;
<bibRefCitation id="0213EC86FFC30273FEC808E9FDB1FE5E" author="Schuckel" box="[368,601,439,464]" pageId="4" pageNumber="505" refString="Schuckel, U., Beck, M. &amp; Kroncke, I. (2013) Spatial variability in structural and functional aspects of macrofauna communities and their environmental parameters in the Jade Bay (Wadden Sea Lower Saxony, southern North Sea). Helgoland Marine Research, 67 (1), 121 - 136." type="journal article" year="2013">
Schückel
<emphasis id="54F64D65FFC30273FE6508E7FDFEFE5E" box="[477,534,439,464]" italics="true" pageId="4" pageNumber="505">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30273FF7F0882FE36FD96" pageId="4" pageNumber="505" type="distribution">
<paragraph id="663D9177FFC30273FF7F0882FE36FD96" blockId="4.[151,1437,151,536]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0882FE88FE7B" bold="true" box="[199,352,476,501]" pageId="4" pageNumber="505">Distribution.</emphasis>
Northern Europe:
<collectingCountry id="1E95D1E7FFC30273FD8E0882FD60FE7B" box="[566,648,476,501]" name="Ireland" pageId="4" pageNumber="505">Ireland</collectingCountry>
, SW
<collectingCountry id="1E95D1E7FFC30273FD7E0882FCCEFE7B" box="[710,806,476,501]" name="United Kingdom" pageId="4" pageNumber="505">England</collectingCountry>
, West coast of
<collectingCountry id="1E95D1E7FFC30273FC6A0883FBC9FE7A" box="[978,1057,477,500]" name="France" pageId="4" pageNumber="505">France</collectingCountry>
, North Sea and Wadden Sea, low intertidal to shallow subtidal.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="EE2B2061FFC60276FF2F0ADCFC4AF8E8" ID-DOI="http://doi.org/10.5281/zenodo.5658075" ID-GBIF-Taxon="119644305" ID-Zenodo-Dep="5658075" LSID="urn:lsid:plazi:treatment:EE2B2061FFC60276FF2F0ADCFC4AF8E8" httpUri="http://treatment.plazi.org/id/EE2B2061FFC60276FF2F0ADCFC4AF8E8" lastPageNumber="502" pageId="1" pageNumber="502">
<subSubSection id="2E98C2FCFFC60276FF2F0ADCFD7BFC12" box="[151,659,898,924]" pageId="1" pageNumber="502" type="nomenclature">
<paragraph id="663D9177FFC60276FF2F0ADCFD7BFC12" blockId="1.[151,659,898,924]" box="[151,659,898,924]" pageId="1" pageNumber="502">
<heading id="3D75261BFFC60276FF2F0ADCFD7BFC12" bold="true" box="[151,659,898,924]" fontSize="11" level="1" pageId="1" pageNumber="502" reason="1">
<emphasis id="54F64D65FFC60276FF2F0ADCFD7BFC12" bold="true" box="[151,659,898,924]" pageId="1" pageNumber="502">
Genus
<taxonomicName id="A182EAF4FFC60276FF560ADCFD7BFC12" ID-CoL="84V7M" authority="Webster &amp; Benedict, 1887" authorityName="Webster &amp; Benedict" authorityYear="1887" box="[238,659,898,924]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FF560ADCFEAEFC12" bold="true" box="[238,326,898,924]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
<bibRefCitation id="0213EC86FFC60276FEF50ADCFD7BFC12" author="Webster" box="[333,659,898,924]" pageId="1" pageNumber="502" refString="Webster, H. E &amp; Benedict, J. E. (1887) The Annelida Chaetopoda from Eastport, Maine. Reports of the U. S. Fish Commission for 1885, 707 - 755, pls. 1 - 8." type="journal article" year="1887">Webster &amp; Benedict, 1887</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC60276FF2F0A97FA7BFC6C" box="[151,1427,969,994]" pageId="1" pageNumber="502" type="reference_group">
<paragraph id="663D9177FFC60276FF2F0A97FA7BFC6C" blockId="1.[151,1437,969,1894]" box="[151,1427,969,994]" pageId="1" pageNumber="502">
<emphasis id="54F64D65FFC60276FF2F0A97FEC5FC6C" bold="true" box="[151,301,969,994]" pageId="1" pageNumber="502">Type-species</emphasis>
<taxonomicName id="A182EAF4FFC60276FE8C0A97FCEEFC6C" ID-CoL="564LP" authority="Webster &amp; Benedict, 1887" authorityName="Webster &amp; Benedict" authorityYear="1887" box="[308,774,969,994]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="species" species="acutus">
<emphasis id="54F64D65FFC60276FE8C0A97FE3CFC6F" box="[308,468,969,994]" italics="true" pageId="1" pageNumber="502">Tharyx acutus</emphasis>
<bibRefCitation id="0213EC86FFC60276FE630A97FCEEFC6C" author="Webster" box="[475,774,969,994]" pageId="1" pageNumber="502" refString="Webster, H. E &amp; Benedict, J. E. (1887) The Annelida Chaetopoda from Eastport, Maine. Reports of the U. S. Fish Commission for 1885, 707 - 755, pls. 1 - 8." type="journal article" year="1887">Webster &amp; Benedict, 1887</bibRefCitation>
</taxonomicName>
, designated by
<bibRefCitation id="0213EC86FFC60276FC0B0A94FB81FC6C" author="Hartman" box="[947,1129,969,994]" pageId="1" pageNumber="502" refString="Hartman, O. (1959) Catalogue of the Polychaetous Annelids of the World. Allan Hancock Foundation Occasional Paper, 23, 1 - 628." type="journal article" year="1959">Hartman (1959)</bibRefCitation>
. Revised by
<bibRefCitation id="0213EC86FFC60276FB410A97FA67FC6C" author="Blake" box="[1273,1423,969,994]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC60276FF7F0AB3FF24FA8C" pageId="1" pageNumber="502" type="diagnosis">
<paragraph id="663D9177FFC60276FF7F0AB3FF24FA8C" blockId="1.[151,1437,969,1894]" pageId="1" pageNumber="502">
<emphasis id="54F64D65FFC60276FF7F0AB3FEA9FB88" bold="true" box="[199,321,1005,1030]" pageId="1" pageNumber="502">Diagnosis.</emphasis>
Prostomium conical; peristomium elongate, with pair of grooved dorsal tentacles arising on posterior margin anterior to setiger 1; first pair of branchiae typically arising immediately posterior to dorsal tentacles either on posterior margin of peristomium or rarely on anterior margin of setiger 1; abdominal segments sometimes beadlike. Noto- and neuropodia located close to one another, not widely separated. Setae including simple capillaries in anterior and middle setigers, acicular spines in posterior setigers with irregular notched tips, sometimes appearing weakly bidentate, with pair of stunted or rounded knobs but never with distinct, sharply pointed teeth; spines present either in both posterior noto- and neuropodia or only in neuropodia. Pygidium with terminal anus and small ventral lobe or disk.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC60276FF7F0C53FC4AF8E8" pageId="1" pageNumber="502" type="discussion">
<paragraph id="663D9177FFC60276FF7F0C53FD9AF9E4" blockId="1.[151,1437,969,1894]" pageId="1" pageNumber="502">
<emphasis id="54F64D65FFC60276FF7F0C53FEDCFAA8" bold="true" box="[199,308,1293,1318]" pageId="1" pageNumber="502">Remarks</emphasis>
.
<bibRefCitation id="0213EC86FFC60276FEFB0C53FE3EFAA8" author="Blake" box="[323,470,1293,1318]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
determined that the
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species of the genus
<emphasis id="54F64D65FFC60276FC5C0C53FB40FAAB" box="[996,1192,1293,1318]" italics="true" pageId="1" pageNumber="502">
<taxonomicName id="A182EAF4FFC60276FC5C0C53FBDEFAA8" box="[996,1078,1293,1318]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">Tharyx</taxonomicName>
,
<taxonomicName id="A182EAF4FFC60276FBFB0C50FB40FAAB" authority="Webster &amp; Benedict, 1887" authorityName="Webster &amp; Benedict" authorityYear="1887" box="[1091,1192,1294,1317]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="species" species="acutus">T. acutus</taxonomicName>
</emphasis>
<bibRefCitation id="0213EC86FFC60276FB080C53FF3AFAC4" author="Webster" pageId="1" pageNumber="502" refString="Webster, H. E &amp; Benedict, J. E. (1887) The Annelida Chaetopoda from Eastport, Maine. Reports of the U. S. Fish Commission for 1885, 707 - 755, pls. 1 - 8." type="journal article" year="1887">Webster &amp; Benedict, 1887</bibRefCitation>
, had knob-tipped hooks or spines in addition to capillaries. This discovery differed from the traditional concept of
<taxonomicName id="A182EAF4FFC60276FEA10C0BFE80FAE0" box="[281,360,1365,1390]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FEA10C0BFE80FAE0" box="[281,360,1365,1390]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
, which had been based upon specimens having all capillaries (
<bibRefCitation id="0213EC86FFC60276FBFE0C08FB18FAE0" author="Hartman" box="[1094,1264,1365,1390]" pageId="1" pageNumber="502" refString="Hartman, O. (1961) Polychaetous annelids from California. Allan Hancock Pacific Expeditions, 22, 1 - 226." type="journal article" year="1961">Hartman 1961</bibRefCitation>
).
<bibRefCitation id="0213EC86FFC60276FABE0C0BFA74FAE0" author="Blake" box="[1286,1436,1365,1390]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
restricted species of
<taxonomicName id="A182EAF4FFC60276FE3E0C27FE3EFA1C" box="[390,470,1401,1426]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FE3E0C27FE3EFA1C" box="[390,470,1401,1426]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
to those having blunt or knob-tipped spines and moved species having simple and denticulated capillaries to the genera
<taxonomicName id="A182EAF4FFC60276FD800CC3FD3BFA38" box="[568,723,1437,1462]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FD800CC3FD3BFA38" box="[568,723,1437,1462]" italics="true" pageId="1" pageNumber="502">Aphelochaeta</emphasis>
</taxonomicName>
and
<taxonomicName id="A182EAF4FFC60276FCB00CC3FC7FFA38" box="[776,919,1437,1462]" class="Polychaeta" family="Cirratulidae" genus="Monticellina" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FCB00CC3FC7FFA38" box="[776,919,1437,1462]" italics="true" pageId="1" pageNumber="502">Monticellina</emphasis>
</taxonomicName>
, respectively. The genus
<taxonomicName id="A182EAF4FFC60276FB0B0CC3FAEBFA38" box="[1203,1283,1437,1462]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FB0B0CC3FAEBFA38" box="[1203,1283,1437,1462]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
appears most closely related to
<taxonomicName id="A182EAF4FFC60276FEE40C9FFE0BFA54" box="[348,483,1473,1498]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FEE40C9FFE0BFA54" box="[348,483,1473,1498]" italics="true" pageId="1" pageNumber="502">Caulleriella</emphasis>
</taxonomicName>
in that both genera have species with modified spines that are more or less bidentate. In addition to the nature of the posterior spines,
<taxonomicName id="A182EAF4FFC60276FD700CBBFCA7FA70" box="[712,847,1509,1534]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FD700CBBFCA7FA70" box="[712,847,1509,1534]" italics="true" pageId="1" pageNumber="502">Caulleriella</emphasis>
</taxonomicName>
and
<taxonomicName id="A182EAF4FFC60276FC330CBBFC32FA70" box="[907,986,1509,1534]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FC330CBBFC32FA70" box="[907,986,1509,1534]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
species are easily distinguished by the position of the noto- and neuropodia. In
<taxonomicName id="A182EAF4FFC60276FDD90F57FD00F9AC" box="[609,744,1545,1570]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FDD90F57FD00F9AC" box="[609,744,1545,1570]" italics="true" pageId="1" pageNumber="502">Caulleriella</emphasis>
</taxonomicName>
, the setal fascicles are widely separated from one another, so much so, that in cross section of some species, they appear at four corners of the body. In
<taxonomicName id="A182EAF4FFC60276FB090F73FAE8F9C8" box="[1201,1280,1581,1606]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FB090F73FAE8F9C8" box="[1201,1280,1581,1606]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
, on the other hand, the setal fascicles are close together.
</paragraph>
<paragraph id="663D9177FFC60276FF7F0F2BFC4AF8E8" blockId="1.[151,1437,969,1894]" pageId="1" pageNumber="502">
<bibRefCitation id="0213EC86FFC60276FF7F0F2BFEB3F900" author="Blake" box="[199,347,1653,1678]" pageId="1" pageNumber="502" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. In: Petersen, M. E. &amp; Kirkegaard, J. B. (Eds.), Proceedings of the Second International Polychaete Conference, Copenhagen. Ophelia, Supplement, No. 5, pp. 17 - 30." type="book chapter" year="1991">Blake (1991)</bibRefCitation>
originally included five species in the genus
<taxonomicName id="A182EAF4FFC60276FCE60F2BFC45F900" box="[862,941,1653,1678]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FCE60F2BFC45F900" box="[862,941,1653,1678]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
, one of which,
<taxonomicName id="A182EAF4FFC60276FBE70F28FB10F900" box="[1119,1272,1653,1678]" class="Polychaeta" family="Cirratulidae" genus="Caulleriella" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="species" species="killariensis">
<emphasis id="54F64D65FFC60276FBE70F28FB10F900" box="[1119,1272,1653,1678]" italics="true" pageId="1" pageNumber="502">T. killariensis</emphasis>
</taxonomicName>
is redescribed in the present paper. Four additional species were added by
<bibRefCitation id="0213EC86FFC60276FCF30FC7FC32F93C" author="Blake" box="[843,986,1689,1714]" pageId="1" pageNumber="502" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">Blake (1996</bibRefCitation>
,
<bibRefCitation id="0213EC86FFC60276FC5F0FC7FBC9F93C" author="Blake" box="[999,1057,1689,1714]" pageId="1" pageNumber="502" refString="Blake, J. A. (2015) New species of Chaetozone and Tharyx (Polychaeta: Cirratulidae) from the Alaskan and Canadian Arctic and the Northeastern Pacific, including a description of the lectotype of Chaetozone setosa Malmgren from Spitzbergen in the Norwegian Arctic. Zootaxa, 3919 (3), 501 - 552. http: // dx. doi. org / 10.11646 / zootaxa. 3919.3.5" type="journal article" year="2015">2015</bibRefCitation>
) and
<bibRefCitation id="0213EC86FFC60276FBDC0FC7FF0CF958" author="Magalhaes" pageId="1" pageNumber="502" refString="Magalhaes, W. F. &amp; Bailey-Brock, J. H. (2013) Bitentaculate Cirratulidae (Annelida: Polychaeta) from the northwestern Pacific Islands with a description of nine new species. Zootaxa, 3630 (1), 80 - 116. http: // dx. doi. org / 10.11646 / zootaxa. 3630.1.3." type="journal article" year="2013">Magalhães &amp; Bailey-Brock (2013)</bibRefCitation>
. Two new species discovered in shallow-water sediments in
<collectingCountry id="1E95D1E7FFC60276FC210FE3FC1AF958" box="[921,1010,1725,1750]" name="Sweden" pageId="1" pageNumber="502">Sweden</collectingCountry>
are here described, bringing the total number of species in the genus to eleven. In addition, another eight species of
<taxonomicName id="A182EAF4FFC60276FBBE0FBFFBBEF974" box="[1030,1110,1761,1786]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FBBE0FBFFBBEF974" box="[1030,1110,1761,1786]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
are known to the first author from North
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and
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. With the number of species thus nearly tripled over the past 24 years, there is now a larger suite of characters available to define the genus than was available in the past. The morphological characters that best define the known species of
<taxonomicName id="A182EAF4FFC60276FD0C0E13FCECF8E8" box="[692,772,1869,1894]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="502" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC60276FD0C0E13FCECF8E8" box="[692,772,1869,1894]" italics="true" pageId="1" pageNumber="502">Tharyx</emphasis>
</taxonomicName>
are reviewed.
</paragraph>
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</treatment>
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<taxonomicName id="B531F3332598DFCB736A55FB6A4DF3FE" ID-CoL="68ZPY" authority="(Peters 1872)" baseAuthorityName="Peters" baseAuthorityYear="1872" class="Mammalia" family="Megadermatidae" genus="Cardioderma" kingdom="Animalia" order="Chiroptera" pageNumber="379" phylum="Chordata" rank="species" species="cor">
<emphasis id="BB187A93FFFA6DDE58C3192AAEF9F01A" italics="true" pageNumber="379">Cardioderma cor</emphasis>
(Peters 1872)
</taxonomicName>
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<treatmentCitation id="8B62F4141F2F74DBFA5C56EEE6510414" author="Peters" authority="Peters 1872" authorityName="Peters" authorityYear="1872" class="Mammalia" family="Megadermatidae" genus="Megaderma" journal="Monatsb. K. Preuss. Akad. Wiss. Berlin" kingdom="Animalia" order="Chiroptera" page="194" pageNumber="379" phylum="Chordata" rank="species" species="cor" title="Megaderma cor" volume="1872" year="1872">
<taxonomicName id="B62B603975BC5CE5E75C7DF8F2C2AC8F" ID-CoL="3YTFP" authority="Peters 1872" authorityName="Peters" authorityYear="1872" class="Mammalia" family="Megadermatidae" genus="Megaderma" kingdom="Animalia" order="Chiroptera" pageNumber="379" phylum="Chordata" rank="species" species="cor">
<emphasis id="7B56BB9211913E9BD07D57A825D876CD" italics="true" pageNumber="379">[Megaderma] cor</emphasis>
Peters 1872
</taxonomicName>
,
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</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F2EA6B7804E023891BD947D8F9E91A41" pageNumber="379" type="materials_examined">
<paragraph id="24EA16EF7B1EC1C45508A3333EC1D043" pageNumber="379">
<emphasis id="7795CE67D767D03FBF2DED3D0C8B4CEE" inLineHeading="true" pageNumber="379">Type Locality:</emphasis>
<materialsCitation id="1ABECF8C08BE7C5B86502C4DEF0D4C3E" country="Ethiopia" pageNumber="379">
<collectingCountry id="99244892D4501B56BCF15224670862C2" name="Ethiopia">Ethiopia</collectingCountry>
.
</materialsCitation>
</paragraph>
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<subSubSection id="67E4DE336B52973CF2410265C90BB285" pageNumber="379" type="vernacular_names">
<paragraph id="33861984045C85E207F7B38B7F399A7B" pageNumber="379">
<emphasis id="CB3F93F78BF2751DBF5F3486BC28AFEB" inLineHeading="true" pageNumber="379">Vernacular Names:</emphasis>
<vernacularName id="A8FDE514E25E7B41C907BD8662E4E6A7" language="eng" pageNumber="379">Heart-nosed Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="DBF01F19E0DFD6ED1B5883877C4C3696" pageNumber="379" type="distribution">
<paragraph id="E7EFEDF5D007674903550A6E295E86FA" pageNumber="379">
<emphasis id="647528442FFBCF84410AF93C6AFE5C3D" inLineHeading="true" pageNumber="379">Distribution:</emphasis>
<collectingCountry id="449216E8B015E79E894A3BC1F1308BD2" name="Ethiopia">Ethiopia</collectingCountry>
,
<collectingCountry id="3A09E00D3346C642831DE6B969D3B502" name="Djibouti">Djibouti</collectingCountry>
,
<collectingCountry id="B272F03A065756D3BB1D9704D49A18FB" name="Somalia">Somalia</collectingCountry>
,
<collectingCountry id="F652036E163B4B0A9EA05526DC32136F" name="Kenya">Kenya</collectingCountry>
,
<collectingCountry id="AB9A11DE51CC629F14D51AAEB44C68D0" name="Uganda">Uganda</collectingCountry>
, E
<collectingCountry id="CB85264ED73BD89C5DCB65D885C8F8EF" name="Sudan">Sudan</collectingCountry>
,
<collectingCountry id="AF157E879F4C376A38FA0D122EFDAB6E" name="Tanzania">Tanzania</collectingCountry>
,
<collectingCountry id="2101514ECEA1360B1D39BA70A073A619" name="Tanzania">Zanzibar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F488C62CE9517482CC2578C75DFF2CF5" pageNumber="379" type="conservation">
<paragraph id="490B05556F61A8FF3967ECDEE0657BDA" pageNumber="379">
<emphasis id="BF667C53D72AE98E5DD946D07328F5EC" inLineHeading="true" pageNumber="379">Conservation:</emphasis>
<collectionCode id="C5E1370E9156CFC59A7BBA9E6B4BC744">IUCN</collectionCode>
2003 and
<collectionCode id="36FCAFD0FCC5FF555FF560BDADE1447C">IUCN</collectionCode>
/
<collectionCode id="4FF687DA86DF65FE407BFBF3418367C2" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) Lower Risk (nt).
</paragraph>
</subSubSection>
</treatment>
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Haliday, 1834
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<taxonomicName id="23AF2C253CF92FCA63DDE739C760DAC2" genus="Ichneumonoidea" lsidName="Ichneumonoidea carbonarius" pageId="0" pageNumber="8151" rank="species" species="carbonarius">carbonarius</taxonomicName>
(Ratzeburg, 1848) preocc.
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<taxonomicName id="1123E5E2B1F22D1F45FA918BEA6E97EB" genus="Ichneumonoidea" lsidName="Ichneumonoidea ater" pageId="0" pageNumber="8151" rank="species" species="ater">ater</taxonomicName>
(Ratzeburg, 1852); synonymy by
<bibRefCitation id="E8761A35CA51D4F1A9DFD57B74A502C5" author="Achterberg, C. van" journalOrPublisher="Zoologische Verhandeligen" pageId="0" pageNumber="8151" pagination="1 - 115" title="Revision of the Haliday collection of Braconidae (Hymenoptera)" volume="314" year="1997">Achterberg (1997)</bibRefCitation>
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<taxonomicName id="BEC52ADB6535F92E883E048F0803A6DA" genus="Ichneumonoidea" lsidName="Ichneumonoidea lugens" pageId="0" pageNumber="8151" rank="species" species="lugens">lugens</taxonomicName>
(Ratzeburg, 1852)
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<taxonomicName id="73997711D921C5CD1A85A0EC7B8ABFAB" genus="Ichneumonoidea" lsidName="Ichneumonoidea lindbergi" pageId="0" pageNumber="8151" rank="species" species="lindbergi">lindbergi</taxonomicName>
Hedqvist, 1965
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<paragraph id="9A094D1B81B32F10DF0FF26DFD69C04C" pageId="0" pageNumber="8151">Distribution</paragraph>
<paragraph id="B8AD6BC3ADBC1F9D6CB3D13A27A15C03" pageId="0" pageNumber="8151">England</paragraph>
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<taxonomicName id="30FC91288DF9FD0517871F5C6AC16C39" ID-CoL="4FYFS" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="megacephala">Pheidole megacephala (Fabricius)</taxonomicName>
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<paragraph id="08CD2E2C820766CBABCD8398D322F3F6" pageNumber="55">Figs. 15a-g</paragraph>
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<taxonomicName id="603D3AAA21434F8DD9396C72E178F110" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:147791" class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Formica megacephala Fabricius" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="megacephala">Formica megacephala Fabricius</taxonomicName>
, 1793: 36. Roger 1863b: 30 (combination in
<taxonomicName id="0A4CC7B7C0EFB03528FEB6DFEBBFF79B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
). Syntype(s): major, no locality given, not examined.
</paragraph>
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<taxonomicName id="22499B2DAB1E1A8AE6D9FABA9D21FEDE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142417" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica trinodis Losana" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="trinodis">Myrmica trinodis Losana</taxonomicName>
, 1834: 327. Roger 1863b: 30 (junior synonym of
<taxonomicName id="E24946F75445A5E5EB1B8EC07DDF0BEA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:189116" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica megacephala (Fabricius)" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
). Syntype(s): &quot;worker&quot;, Italy, not examined.
</paragraph>
<paragraph id="0060F1F350C8CDCC7F0EE971CD8101FC" pageNumber="55">
<taxonomicName id="38D4D6AC4ABBCF96F2F36D3E7350A5D9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142412" class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Formica edax Forskal" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="edax">Formica edax Forskal</taxonomicName>
, 1775: 84. Emery 1892: 160 (junior synonym of
<taxonomicName id="5AE48282963C437EADDAB0320C177297" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:147791" class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Formica megacephala Fabricius" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
), Dalla Torre 1892: 90 (same). Syntype(s): &quot;worker&quot;, Egypt, not examined.
</paragraph>
<paragraph id="2762006E811F7B05CAB58AC525E5D2AC" pageNumber="55">
<taxonomicName id="417C831C240C7FEA5A1ABC8237CCF80D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142413" class="Insecta" genus="Oecophthora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Oecophthora perniciosa Gerstaecker" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="perniciosa">Oecophthora perniciosa Gerstacker</taxonomicName>
, 1859: 263. Roger 1863b: 31 (combination in
<taxonomicName id="26BA0B5DFA1400B5DDBA437A3A89406A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
), Emery, 1915c: 235 (junior synonym of
<taxonomicName id="632FC7E5C3B6B7D9D293C39907295272" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="55" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
). Syntype(s): &quot;worker&quot;, Mozambique, not examined.
</paragraph>
<paragraph id="A15B339F2385083783839654B384F281" pageNumber="56">
<taxonomicName id="A77A5686DFD7726EE81C122883C66659" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142414" class="Insecta" genus="Oecophthora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Oecophthora pusilla Heer" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="pusilla">
<pageStartToken id="3F15D942A9778A3669C88FA8744386BD" pageNumber="56">Oecophthora</pageStartToken>
pusilla Heer
</taxonomicName>
, 1852: 15. F. Smith 1858: 173 (combination in
<taxonomicName id="387E38DC8DB3A96357FB8E500BFAE654" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
), Roger 1859: 259 (senior synonym of
<taxonomicName id="A958A1E8CCF97D9CD1FD509D3DEF8B1C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142696" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole laevigata (Smith)" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="laevigata">laevigata Fr. Smith</taxonomicName>
, 1855: 130), Mayr 1870: 981 (senior synonym of
<taxonomicName id="9D902F973F9735C421826EFADAB1F836" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142696" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole laevigata (Smith)" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="laevigata">laevigata Mayr</taxonomicName>
, 1862: 747), Mayr 1886: 360 (senior synonym of
<taxonomicName id="434D5DD819E17EE293763952D1820D67" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142694" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole janus Smith" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="janus">janus</taxonomicName>
), Emery 1915: 235 (subspecies of
<taxonomicName id="E6975A4AC96B1A21350526438FC4F427" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
), Wheeler 1922: 812 (junior synonym of
<taxonomicName id="10C8613D374DD534710A9FB366DDB62E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
). Syntypes: major, minor, queen &amp; male, Madeira, not examined.
</paragraph>
<paragraph id="A930A10D6ADD9A4A212F4C296D0DCA0D" pageNumber="56">
<taxonomicName id="CBF2E0C991E1A49367C1B2087E3542E5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142132" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica agilis Smith" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="agilis">Myrmica agilis F. Smith</taxonomicName>
, 1857: 71. Donisthorpe 1932: 449 (combination in
<taxonomicName id="4C126733B332265BE73651944197902A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
).
<taxonomicNameLabel id="15B7DF1590D1EDBF2C6F92CF5983FA7D">Syn.n.</taxonomicNameLabel>
Syntypes: 3 minors, &quot;MALAC&quot; [= Malacca, S. Malay Peninsula], OXUM TYPE HYM: 988 1-3/3, examined.
</paragraph>
<paragraph id="E5A135E37E74D4ED60A2242D22014A99" pageNumber="56">
<taxonomicName id="0AD3B1BB31870AFC9803A715A3694757" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142415" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica suspiciosa Smith" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="suspiciosa">Myrmica suspiciosa F. Smith</taxonomicName>
, 1859: 148. Donisthorpe 1932: 455 (junior synonym of
<taxonomicName id="2773BBB1CF9D82575219C73B9DEDF7C4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:189116" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica megacephala (Fabricius)" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
). Syntype (s): &quot;worker&quot;, Aru I. (Indonesia), not examined.
</paragraph>
<paragraph id="25D91B72C05511C100CD19CFD31FD648" pageNumber="56">
<taxonomicName id="D6DAA13B07BB14E5AB29617BF26561E6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142416" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Atta testacea Smith" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="testacea">Atta testacea F. Smith</taxonomicName>
, 1858: 168. Mayr 1886: 360 (combination in
<taxonomicName id="EE3E7B252E15997D00CB1439CD5FBA8B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
), Brown, 1981: 530 (junior synonym of
<taxonomicName id="5271B2C0FA24661173103FFBB94A357A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
). Syntypes: major &amp; minor, Brazil, not examined.
</paragraph>
<paragraph id="F462900C82DD674528EF2B29424B16BE" pageNumber="56">
Subspecies enumerated in Bolton, 1995: nominal plus
<taxonomicName id="A4A0DEBA5D9650BFE2894DF295BADCBF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151403" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala costauriensis Santschi" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="costauriensis">costauriensis Santschi</taxonomicName>
, 1914: 443, syntype(s): major, Ghana, not examined;
<taxonomicName id="9A9D52CBEDA17E9B216E81704857CADD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151405" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala duplex Santschi" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="duplex">duplex Santschi</taxonomicName>
, 1937a: 220, syntypes: major, minor &amp; queen, Angola, not examined;
<taxonomicName id="A32F897B1A6564782A94819E5D7CA02C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151406" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala ilgi Forel" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="ilgi">ilgi Forel</taxonomicName>
, 1907: 82, syntypes: major &amp; minor, Ethiopia, not examined.;
<taxonomicName id="2B456388F0E1171BB7051F2DB439A73A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151407" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala impressifrons Wasmann" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="impressifrons">impressifrons Wasmann</taxonomicName>
, 1905: 110 (replacement name for
<taxonomicName id="9BC8B5C5F5A9BD9F6840BE1A3F6C8BED" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33794" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole impressiceps Mayr" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="impressiceps">impressiceps Wasmann</taxonomicName>
, 1904: 72), syntypes: major, minor &amp; queen, South Africa, not examined;
<taxonomicName id="BBBA8D062C53F7405549FE2DADCBA477" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:229941" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole melancholica Santschi" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="melancholica">melancholica Santschi</taxonomicName>
, 1912: 164, syntypes: major &amp; minor, Ivory Coast, not examined;
<taxonomicName id="0D838262195F82B24575AA3A66DF9A5A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151409" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala nkomoana Forel" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="nkomoana">nkomoana Forel</taxonomicName>
, 1916: 415, syntypes: major, minor, queen &amp; male, Zaire, not examined;
<taxonomicName id="5C72DCBF438BB4176DE94B4301FAA082" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142408" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole rotundata Forel" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="rotundata">rotundata Forel</taxonomicName>
, 1894: 92, syntypes: major &amp; minor, Mozambique, not examined;
<taxonomicName id="EF747CC2F99E867EC6C1FC659C14A374" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151411" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala scabrior Forel" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="scabrior">scabrior Forel</taxonomicName>
, 1891: 178, syntypes: major &amp; minor, Madagascar, not examined;
<taxonomicName id="15FFCFFF61A755FC4CFF5C9FB1AE208F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:189134" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole speculifrons Emery" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="speculifrons">speculifrons Stitz</taxonomicName>
, 1911: 386, syntypes: major &amp; minor, Tanzania, not examined;
<taxonomicName id="0430489216B68F5BC61CBB6D6199915F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142411" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole talpa Gerstaecker" order="Hymenoptera" pageNumber="56" phylum="Arthropoda" rank="species" species="talpa">talpa Gerstacker</taxonomicName>
, 1871: 356, syntypes: &quot;worker&quot; &amp; queen, Kenya, not examined. For these forms type material not examined.
</paragraph>
</subSubSection>
<subSubSection id="91A7794AB73E4B01C34CD3F969F6D48C" type="materials_examined">
<paragraph id="BA591C591C62C3679AB8F46C0EB659D9" pageNumber="56">Other material examined: S. China: Hong Kong: Victoria Park, Hong Kong I. [K. Eguchi]; Macau: Mong Ha [K. Eguchi]. N. Vietnam: Ha Noi: Hanoi Agric. Univ. (Gia Lam) [K. Ogata: 15-min TUS #2]; Quang Ninh: Hoanh Bo [K. Eguchi]. S. Vietnam: Vinh Long (misspelled as &quot;Vinlong&quot;): Vinh Long (10°15'N, 105°58'N) [S. Kawaguchi]. Thailand: Trang: Khao Chong Waterfall [Eg01-VN-761]. W. Malaysia: Penang: beside a building of Univ. Sains Malaysia [C.Y. Lee]. E. Malaysia: Sabah: Kota Kinabalu [Eg97-BOR-376], Tambunan Village [H. Okido], Danam Valley [Eg96-BOR-108]. Indonesia: Kalimantan Timur: Tandjung Isuy [Seyfert &amp; Graindl]; Irian Jaya: Wamena, 1600 m alt. [Eg98-IRI-674, -675, -676, -703]. Australia: Queensland: S. Mission Beach near Tully [AU01-SKY-12]. Tonga: Tongatapu: Vaini [J.K. Wetterer].</paragraph>
<paragraph id="57B566F065AE4A44C7BCACED2FB3B4A9" pageNumber="56">Worker measurements &amp; indices: Major (n=5). - HL 1.28-1.45 mm; HW 1.25-1.45 mm; CI 98-100; SL 0.71-0.76 mm; SI 52-57; FL 0.94-0.98 mm; FI 68-77.</paragraph>
<paragraph id="1A28930DDDEEB9C16DC7659C66DA3CB6" pageNumber="56">Minor (n=5). - HL 0.62-0.72 mm; HW 0.55-0.65 mm; CI 88-91; SL 0.67-0.73 mm; SI 111-121; FL 0.68-0.77 mm; FI 118-123. Worker description</paragraph>
<paragraph id="E4F3F0377E7B3E70AC3BF6485FF82F3E" pageNumber="56">Major. - Head in lateral view roundly convex dorsally, not impressed on vertex, in full-face view shallowly concave posteriorly; frons longitudinally rugose (or rarely almost smooth, only sparsely with short interrupted longitudinal rugulae); vertex and dorsum of vertexal lobe smooth and shining or shagreened; frontal carina absent or present just as weak rugula(e); antennal scrobe absent; median longitudinal carina of clypeus weak or absent; hypostoma at most with a pair of very small or inconspicuous submedian processes in addition to a pair of conspicuous lateral processes; antenna with a 3-segmented club; maximal diameter of eye almost as long as or longer than antennal segment X. Promesonotal dome in dorsal view smooth and shining or shagreened, sometimes with several weak transverse rugulae, in lateral view at most with an inconspicuous mound on its posterior slope; humerus not or weakly produced laterad; the dome at the humeri narrower than at the bottom; mesopleuron, metapleuron and lateral face of propodeum weakly or very weakly punctured. Petiole a little longer than postpetiole (excluding helcium); postpetiole not massive; its anteroventral part weakly swollen. First gastral tergite smooth and shining entirely, or very weakly punctured around its articulation with postpetiole and smooth or shagreened in the remainder.</paragraph>
</subSubSection>
<subSubSection id="2AF22C81752C7263E1EDF4B340E32B7E" type="multiple">
<caption id="9CF8FB5CC3F34F70298E68A07F1E8CE4">
<paragraph id="36A69418F165CCA1CDF9A856859D60F5" pageNumber="57">
<pageStartToken id="49CA2B5555296B3D77B418608E5DBFDC" pageNumber="57">
<pageStartToken id="7A788E1FE70A56CC0D8E4B95CA46FD49" pageNumber="57">FIGURE</pageStartToken>
</pageStartToken>
15a-d,
<taxonomicName id="F250C8B492B2D76B4C34C82D1B9B2B52" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="57" phylum="Arthropoda" rank="species" species="megacephala">Pheidole megacephala</taxonomicName>
, major, Thailand [Eg01-TH-761] - a, head in full-face view; b, head in lateral view; c, mesosoma and waist in dorsal view; d, mesosoma and waist in lateral view.
</paragraph>
</caption>
<caption id="A929F29516D26ADECE2E748649A5D2BD">
<paragraph id="CDF0325B91C28C3E8CD9F027C87E1188" pageNumber="58">
<pageStartToken id="E97EAF85C0ECD61D9E0E58876C935FC3" pageNumber="58">
<pageStartToken id="551CCE7CC4291D1BF80BD8CF0585A688" pageNumber="58">FIGURE</pageStartToken>
</pageStartToken>
15e-g,
<taxonomicName id="94FEFA1C693C05CE3ABC41E1502CCB5E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="58" phylum="Arthropoda" rank="species" species="megacephala">Pheidole megacephala</taxonomicName>
, minor, Thailand [Eg01-TH-761] - e, head in full-face view; f, mesosoma and waist in dorsal view; g, mesosoma and waist in lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DCEAE8D0AD8C3BECAB4BF01E1154AAF6" type="description">
<paragraph id="62EC50499EA9B8156A21A70EFB3DCF05" lastPageNumber="59" pageNumber="58">Minor. - Head smooth and shining; preoccipital carina weak but present dorsally and laterally; median part of clypeus smooth and shining, without a median longitudinal carina; antenna with a 3-segmented club; scape extending beyond posterolateral margin of head by the double length of antennal segment II or more; maximal diameter of eye almost as long as, or sometimes a little shorter than antennal segment X. Promesonotal dome smooth and shining, in lateral view lacking a mound on its posterior slope; humerus in dorso-oblique view not or hardly produced; mesopleuron, metapleuron and lateral face of propodeum punctured weakly; metanotal groove inconspicuous. Petiole almost as long as or a little longer than postpetiole (excluding helcium); postpetiole relatively long but not massive; its anteroventral part weakly swollen.</paragraph>
<paragraph id="56A289FFE178C75D6243B6474AC09012" pageNumber="59">
Recognition: The syntype minors of &quot;
<taxonomicName id="CD46A2D54FCFFD324820FAE52A8FFD51" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142132" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica agilis Smith" order="Hymenoptera" pageNumber="59" phylum="Arthropoda" rank="species" species="agilis">Myrmica agilis</taxonomicName>
&quot; agree well with minors of Bornean populations (e.g., Eg96-BOR-108) of P
<taxonomicName id="9A306786627FD2AE7FA89FDEDDF4A432" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:189116" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica megacephala (Fabricius)" order="Hymenoptera" pageNumber="59" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
. I conclude that P
<taxonomicName id="BA84D50C1A4511AC7F107E2672053F76" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142132" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica agilis Smith" order="Hymenoptera" pageNumber="59" phylum="Arthropoda" rank="species" species="agilis">agilis</taxonomicName>
is a juninor synonym of P
<taxonomicName id="2BCFC24653B75B04ABCEDFAA6EB18F13" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:189116" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmica megacephala (Fabricius)" order="Hymenoptera" pageNumber="59" phylum="Arthropoda" rank="species" species="megacephala">megacephala</taxonomicName>
.
</paragraph>
<paragraph id="F9A6F8522700BB79F8BC9DAD149609CA" pageNumber="59">
<taxonomicName id="41874E34ED0CA5478CE9D904FAC471E2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33860" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole megacephala (Fabricius)" order="Hymenoptera" pageNumber="59" phylum="Arthropoda" rank="species" species="megacephala">P. megacephala</taxonomicName>
is well distinguished from Indo-Chinese species by the combination of the following characteristics: in the major head in full-face view only shallowly concave posteriorly; in the major dorsum of vertexal lobe smooth and shining or shagreened; in the major hypostoma in the middle at most with a pair of very small or inconspicuous submedian processes; in the minor preoccipital carina weak but present dorsally and laterally; posterior slope of promesonotal dome at most with an inconspicuous mound in the major, and without any mound in the minor; in the major and minor anteroventral part of postpetiole weakly swollen.
</paragraph>
</subSubSection>
<subSubSection id="5E02E8BF816ECF6705F20A358680818E" type="distribution">
<paragraph id="C963E06D06371A663A3F5EBC4EFBE68B" pageNumber="59">Distribution &amp; bionomics: Widely distributed in the world tropics and subtropics. For detailed information on biology and ecological and economic impacts of this species see Reimer et al. (1993), Campbell (1994), Hoffmann (1998), Wetterer (1998), Hoffmann et al. (1999), Vanderwoude et al. (2000), etc.</paragraph>
</subSubSection>
</treatment>
</document>

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