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<document id="1CA15366A5B7B5A9FB3F05ADE760AC65" ID-DOI="10.11646/zootaxa.3701.1.5" ID-GBIF-Dataset="4990ca63-2241-4abb-bce6-df8099ff2de1" ID-ISSN="1175-5326" ID-Zenodo-Dep="284494" ID-ZooBank="27A3D064-8A6F-4FCE-B483-21BDF9BDDE8E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460666038872" checkinUser="plazi" docAuthor="Sabella, Giorgio, Nouira, Vera D’Urso Said, Souissi, Rhida &amp; Besuchet, Claude" docDate="2013" docId="24787B068C30B650FF7CF9CF741D2BA4" docLanguage="en" docName="zt03701p075.pdf" docOrigin="Zootaxa 3701 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tychobythinus peyerimhoffi Jeannel 1956" docType="treatment" docVersion="7" lastPageNumber="73" masterDocId="D841037E8C38B65AFFEBFF847676291C" masterDocTitle="A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="75" masterPageNumber="63" pageNumber="71" updateTime="1698347793766" updateUser="plazi">
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<mods:titleInfo id="246AC4EA1C6EE92C186A1C07F44B0DF8">
<mods:title id="AAD69730CA6F3AC3C3162F27B29FF314">A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="B31D2A68E80BB2A968DEEEEFE4634A7D">Sabella, Giorgio</mods:namePart>
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<mods:namePart id="46166D9036F2CC6FC2F11BBBC281C4A3">Nouira, Vera D’Urso Said</mods:namePart>
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<mods:namePart id="F7003C00A6885A7AEEED13646BA4DF3B">Souissi, Rhida</mods:namePart>
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<mods:namePart id="B78C00CCD9B51A0B67448C02C9F50DDF">Besuchet, Claude</mods:namePart>
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<mods:typeOfResource id="5EFACF8C55043808C3E67F88B423072F">text</mods:typeOfResource>
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<mods:title id="F24D655273A9ADD512CB610DAA02844C">Zootaxa</mods:title>
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<mods:part id="5F94B18F2BFCAB78743945F5A4DDDE95">
<mods:date id="69C38C84D6BD38AE6319C4D36E77A10E">2013</mods:date>
<mods:detail id="99DB0D69A608EEDBEFA2FA6147CC0116" type="volume">
<mods:number id="C6300CC4EDA0ED9B854D3D8DEA9CA598">3701</mods:number>
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<mods:identifier id="575E4F87EE4B4EF3C9931BF918306E2C" type="DOI">10.11646/zootaxa.3701.1.5</mods:identifier>
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<treatment id="24787B068C30B650FF7CF9CF741D2BA4" ID-DOI="http://doi.org/10.5281/zenodo.6145504" ID-GBIF-Taxon="127665327" ID-Zenodo-Dep="6145504" LSID="urn:lsid:plazi:treatment:24787B068C30B650FF7CF9CF741D2BA4" httpUri="http://treatment.plazi.org/id/24787B068C30B650FF7CF9CF741D2BA4" lastPageId="10" lastPageNumber="73" pageId="8" pageNumber="71">
<subSubSection id="E4CB999B8C30B652FF7CF9CF74A32FF6" pageId="8" pageNumber="71" type="nomenclature">
<paragraph id="AC6ECA108C30B652FF7CF9CF74D82F79" blockId="8.[151,686,1611,1670]" box="[151,686,1611,1637]" pageId="8" pageNumber="71">
<heading id="F7267D7C8C30B652FF7CF9CF74D82F79" bold="true" box="[151,686,1611,1637]" fontSize="11" level="1" pageId="8" pageNumber="71" reason="1">
<taxonomicName id="6BD1B1938C30B652FF7CF9CF74D82F79" ID-CoL="59SF8" authority="Jeannel, 1956" authorityName="Jeannel" authorityYear="1956" box="[151,686,1611,1637]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C30B652FF7CF9CF74D82F79" bold="true" box="[151,686,1611,1637]" pageId="8" pageNumber="71">
<emphasis id="9EA516028C30B652FF7CF9CF779C2F79" bold="true" box="[151,490,1611,1637]" italics="true" pageId="8" pageNumber="71">Tychobythinus peyerimhoffi</emphasis>
(Jeannel, 1956)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AC6ECA108C30B652FF7CF9E9775B2F9A" blockId="8.[151,686,1611,1670]" box="[151,301,1645,1670]" pageId="8" pageNumber="71">
(
<figureCitation id="34EAD6958C30B652FF74F9E977532F9A" box="[159,293,1645,1670]" captionStart="FIGURES 14 – 17" captionStartId="9.[151,264,1907,1929]" captionTargetBox="[171,1415,639,1875]" captionTargetId="figure@9.[151,1435,625,1897]" captionTargetPageId="9" captionText="FIGURES 14 – 17. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 14 — Head, schematic drawing. 15 — Left maxillary palpus, lectotype. 16 — Right antenna, lectotype. 17 — Aedeagus dorsal view, lectotype." httpUri="https://zenodo.org/record/284499/files/figure.png" pageId="8" pageNumber="71">Figs. 14–17</figureCitation>
)
</paragraph>
<paragraph id="AC6ECA108C30B652FF7CF93174A32FF6" blockId="8.[151,1180,1717,1770]" pageId="8" pageNumber="71">
<taxonomicName id="6BD1B1938C30B652FF7CF931742C2FD7" authority="Jeannel 1956: 61" authorityName="Jeannel" authorityPageNumber="61" authorityYear="1956" box="[151,602,1717,1739]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C30B652FF7CF93177D32FD7" box="[151,421,1717,1739]" italics="true" pageId="8" pageNumber="71">Anopsibythus peyerimhoffi</emphasis>
Jeannel 1956: 61
</taxonomicName>
; 63, figs 52 (habitus), 55 (aedeagus) and 56 (profemora);
<taxonomicName id="6BD1B1938C30B652FF7CF95074A72FF6" authority="Lobl &amp; Besuchet 2004: 315" authorityName="Lobl &amp; Besuchet" authorityPageNumber="315" authorityYear="2004" box="[151,721,1748,1770]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C30B652FF7CF95077D92FF6" box="[151,431,1748,1770]" italics="true" pageId="8" pageNumber="71">Tychobythinus peyerimhoffi</emphasis>
Löbl &amp; Besuchet 2004: 315
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C30B650FF7CF89274B62B4C" lastPageId="10" lastPageNumber="73" pageId="8" pageNumber="71" type="materials_examined">
<paragraph id="AC6ECA108C30B652FF7CF89272152E6B" blockId="8.[151,1436,1814,2019]" pageId="8" pageNumber="71">
<emphasis id="9EA516028C30B652FF7CF89277482E33" bold="true" box="[151,318,1814,1839]" pageId="8" pageNumber="71">
<typeStatus id="736A74B28C30B652FF7CF89276A42E33" box="[151,210,1814,1839]" pageId="8" pageNumber="71">Type</typeStatus>
material
</emphasis>
.
<emphasis id="9EA516028C30B652FEA6F89277B52E33" bold="true" box="[333,451,1814,1839]" pageId="8" pageNumber="71">
<typeStatus id="736A74B28C30B652FEA6F89277B52E33" box="[333,451,1814,1839]" pageId="8" pageNumber="71" type="lectotype">Lectotype</typeStatus>
</emphasis>
, male, present designation.
<emphasis id="9EA516028C30B652FD14F89272022E33" bold="true" box="[767,1140,1814,1839]" pageId="8" pageNumber="71">
NORTHEASTERN
<collectingCountry id="D4C68A808C30B652FC19F89272022E33" box="[1010,1140,1814,1839]" name="Algeria" pageId="8" pageNumber="71">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C30B652FB68F89273242E33" bold="true" box="[1155,1362,1814,1839]" pageId="8" pageNumber="71">Annaba province</emphasis>
: Bône (old name of Annaba),
<date id="D86FECD08C30B652FE4BF8BF77852E4F" box="[416,499,1851,1875]" pageId="8" pageNumber="71" value="1929-10">X.1929</date>
(H. Normand) (ex coll. Peyerimhoff, MNHN).
<emphasis id="9EA516028C30B652FBFAF8BE72B72E4F" bold="true" box="[1041,1217,1850,1875]" pageId="8" pageNumber="71">
<typeStatus id="736A74B28C30B652FBFAF8BE72B72E4F" box="[1041,1217,1850,1875]" pageId="8" pageNumber="71" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
,
<specimenCount id="BAD701998C30B652FB3BF8BF73692E4F" box="[1232,1311,1850,1875]" pageId="8" pageNumber="71" type="male">1 male</specimenCount>
, same data of
<typeStatus id="736A74B28C30B652FF5EF8DA776A2E6B" box="[181,284,1886,1911]" pageId="8" pageNumber="71" type="lectotype">lectotype</typeStatus>
(ex coll. Peyerimhoff, MNHN);
<specimenCount id="BAD701998C30B652FD67F8DB748D2E6B" box="[652,763,1886,1911]" pageId="8" pageNumber="71" type="female">2 females</specimenCount>
, same data of
<typeStatus id="736A74B28C30B652FC70F8DA72772E6B" box="[923,1025,1886,1911]" pageId="8" pageNumber="71" type="lectotype">lectotype</typeStatus>
(INAT).
</paragraph>
<paragraph id="AC6ECA108C30B652FF2CF80674CF2EFF" blockId="8.[151,1436,1814,2019]" pageId="8" pageNumber="71">
<emphasis id="9EA516028C30B652FF2CF806771B2E87" bold="true" box="[199,365,1922,1947]" pageId="8" pageNumber="71">Redescription</emphasis>
. Body length
<quantity id="6B2967F58C30B652FDF8F80674CC2E86" box="[531,698,1922,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="8" pageNumber="71" unit="mm" value="1.225" valueMax="1.25" valueMin="1.2">1.20–1.25 mm</quantity>
, pubescence consists of long and flattened setae (length:
<quantity id="6B2967F58C30B652FAA4F806777A2EA3" metricMagnitude="-4" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="8" pageNumber="71" unit="mm" value="0.1275" valueMax="0.13" valueMin="0.125">0.125– 0.130 mm</quantity>
) on head, pronotum, elytra and abdomen, other suberected setae shorter (length:
<quantity id="6B2967F58C30B652FB46F82373192EA3" box="[1197,1391,1959,1983]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="8" pageNumber="71" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.030–0.040 mm</quantity>
) on antennae and legs. Tegument smooth and glossy.
</paragraph>
<paragraph id="AC6ECA108C31B653FF2CFF1374C62B20" blockId="9.[151,1436,151,573]" pageId="9" pageNumber="72">
Head (
<figureCitation id="34EAD6958C31B653FEFAFF13771729AC" box="[273,353,151,176]" captionStart="FIGURES 14 – 17" captionStartId="9.[151,264,1907,1929]" captionTargetBox="[171,1415,639,1875]" captionTargetId="figure@9.[151,1435,625,1897]" captionTargetPageId="9" captionText="FIGURES 14 – 17. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 14 — Head, schematic drawing. 15 — Left maxillary palpus, lectotype. 16 — Right antenna, lectotype. 17 — Aedeagus dorsal view, lectotype." httpUri="https://zenodo.org/record/284499/files/figure.png" pageId="9" pageNumber="72">Fig. 14</figureCitation>
) slightly longer (
<quantity id="6B2967F58C31B653FDCBFF1C749429AC" box="[544,738,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="9" pageNumber="72" unit="mm" value="0.215" valueMax="0.22" valueMin="0.21">0.210–0.220 mm</quantity>
) than wide (0.200–
<quantity id="6B2967F58C31B653FC52FF1C725929AC" box="[953,1071,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="72" unit="mm" value="0.21">0.210 mm</quantity>
), frontal lobe width
<quantity id="6B2967F58C31B653FAFBFF1376B329C8" metricMagnitude="-4" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="9" pageNumber="72" unit="mm" value="0.1225" valueMax="0.125" valueMin="0.12">0.120–0.125 mm</quantity>
, its dorsal surface glossy with some punctures and well impressed median sulcus. Clypeal carina evident, both in dorsal and lateral view, and reaching ocular region. Occipital region with strong median longitudinal carina reaching posterior edge of vertexal foveae, tempora rounded. Gular region convex entirely traversed by evident median longitudinal carina, its surface glossy. Segment II of maxillary palpi elongated with 22–26 strong tubercles, segment III longer than wide with 4–6 tubercles, last segment of maxillary palpi (
<figureCitation id="34EAD6958C31B653FBCEFEC872032879" box="[1061,1141,332,357]" captionStart="FIGURES 14 – 17" captionStartId="9.[151,264,1907,1929]" captionTargetBox="[171,1415,639,1875]" captionTargetId="figure@9.[151,1435,625,1897]" captionTargetPageId="9" captionText="FIGURES 14 – 17. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 14 — Head, schematic drawing. 15 — Left maxillary palpus, lectotype. 16 — Right antenna, lectotype. 17 — Aedeagus dorsal view, lectotype." httpUri="https://zenodo.org/record/284499/files/figure.png" pageId="9" pageNumber="72">Fig. 15</figureCitation>
) more than 3 times longer (
<quantity id="6B2967F58C31B653FF75FEEB77152894" box="[158,355,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="2.625" metricValueMax="2.65" metricValueMin="2.6" pageId="9" pageNumber="72" unit="mm" value="0.2625" valueMax="0.265" valueMin="0.26">0.260–0.265 mm</quantity>
) than wide (
<quantity id="6B2967F58C31B653FE1DFEEB74072894" box="[502,625,367,392]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="72" unit="mm" value="0.075">0.075 mm</quantity>
), its lateral margin straight. Antenna (
<figureCitation id="34EAD6958C31B653FBC3FEEB72F02894" box="[1064,1158,367,392]" captionStart="FIGURES 14 – 17" captionStartId="9.[151,264,1907,1929]" captionTargetBox="[171,1415,639,1875]" captionTargetId="figure@9.[151,1435,625,1897]" captionTargetPageId="9" captionText="FIGURES 14 – 17. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 14 — Head, schematic drawing. 15 — Left maxillary palpus, lectotype. 16 — Right antenna, lectotype. 17 — Aedeagus dorsal view, lectotype." httpUri="https://zenodo.org/record/284499/files/figure.png" pageId="9" pageNumber="72">Fig. 16</figureCitation>
)
<quantity id="6B2967F58C31B653FB73FEEB732C289B" box="[1176,1370,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="6.8500000000000005" metricValueMax="6.8999999999999995" metricValueMin="6.800000000000001" pageId="9" pageNumber="72" unit="mm" value="0.685" valueMax="0.69" valueMin="0.68">0.680–0.690 mm</quantity>
long, scape more than 4.5 times longer (0.200–
<quantity id="6B2967F58C31B653FD62FE10757528B0" box="[649,771,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="72" unit="mm" value="0.21">0.210 mm</quantity>
) than wide (
<quantity id="6B2967F58C31B653FC4BFE10721D28B0" box="[928,1131,404,428]" metricMagnitude="-5" metricUnit="m" metricValue="4.299999999999999" metricValueMax="4.3999999999999995" metricValueMin="4.2" pageId="9" pageNumber="72" unit="mm" value="0.043" valueMax="0.044" valueMin="0.042">0.042–0.044 mm</quantity>
), antennomere II, slightly asymmetric, longer (
<quantity id="6B2967F58C31B653FE96FE33778728CC" box="[381,497,439,464]" metricMagnitude="-5" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="72" unit="mm" value="0.062">0.062 mm</quantity>
) than wide (
<quantity id="6B2967F58C31B653FD97FE3C748528CC" box="[636,755,440,464]" metricMagnitude="-5" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="72" unit="mm" value="0.037">0.037 mm</quantity>
), antennomeres III–V distinctly longer than wide, III slightly longer than IV and V, IV and V subequal in size, VI shorter than V and slightly longer than wide, VII and VIII as long as wide. Antennomeres IX–X distinctly wider than long, XI about 2 times longer than wide, and more than 1.5 times longer than combined length of IX and X.
</paragraph>
<caption id="F8AE9A988C31B653FF7CF8F775C12EBB" httpUri="https://zenodo.org/record/284499/files/figure.png" pageId="9" pageNumber="72" targetBox="[171,1415,639,1875]" targetPageId="9">
<paragraph id="AC6ECA108C31B653FF7CF8F775C12EBB" blockId="9.[151,1436,1907,1959]" pageId="9" pageNumber="72">
<emphasis id="9EA516028C31B653FF7CF8F777212E95" bold="true" box="[151,343,1907,1929]" pageId="9" pageNumber="72">FIGURES 14–17.</emphasis>
<taxonomicName id="6BD1B1938C31B653FE8AF8F7740B2E95" box="[353,637,1907,1929]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C31B653FE8AF8F7740B2E95" box="[353,637,1907,1929]" italics="true" pageId="9" pageNumber="72">Tychobythinus peyerimhoffi</emphasis>
</taxonomicName>
. Scale: 0,1 mm. 14—Head, schematic drawing. 15—Left maxillary palpus, lectotype. 16—Right antenna, lectotype. 17—Aedeagus dorsal view, lectotype.
</paragraph>
</caption>
<paragraph id="AC6ECA108C32B650FF2CFF1C747529CF" blockId="10.[151,1436,151,696]" pageId="10" pageNumber="73">
Pronotum wider (
<quantity id="6B2967F58C32B650FE67FF1C747729AC" box="[396,513,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="10" pageNumber="73" unit="mm" value="0.31">0.310 mm</quantity>
) than long (
<quantity id="6B2967F58C32B650FD61FF13757429AC" box="[650,770,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="2.8499999999999996" pageId="10" pageNumber="73" unit="mm" value="0.285">0.285 mm</quantity>
), widest in apical fourth, pleural carina difficult to see, and antebasal sulcus well impressed.
</paragraph>
<paragraph id="AC6ECA108C32B650FF2CFF59749A29E9" blockId="10.[151,1436,151,696]" box="[199,748,221,246]" pageId="10" pageNumber="73">
Elytra width 0.500 mm, length
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.
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<paragraph id="AC6ECA108C32B650FF2CFE8474BE2804" blockId="10.[151,1436,151,696]" box="[199,712,255,280]" pageId="10" pageNumber="73">Metasternum with surface smooth and glossy.</paragraph>
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Legs elongated, protrochanters with 4–6 tubercles; profemora with 12–14 small tubercles. Metatibiae
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long.
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<paragraph id="AC6ECA108C32B650FF2CFEE372052B17" blockId="10.[151,1436,151,696]" pageId="10" pageNumber="73">
Male.
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<emphasis id="9EA516028C32B650FEE4FEED77CE289C" box="[271,440,359,384]" italics="true" pageId="10" pageNumber="73">T. peyerimhoffi</emphasis>
</taxonomicName>
does not show a strong sexual dimorphism. Antennae are similar in both sexes; the only external character that distinguishes male from female is article II of protarsi which in male is slightly compressed laterally. Also pro- and mesotibiae of male are more sharply defined and sinuate in basal third. Aedeagus (
<figureCitation id="34EAD6958C32B650FAA9FE2973E528DA" box="[1346,1427,429,454]" captionStart="FIGURES 14 – 17" captionStartId="9.[151,264,1907,1929]" captionTargetBox="[171,1415,639,1875]" captionTargetId="figure@9.[151,1435,625,1897]" captionTargetPageId="9" captionText="FIGURES 14 – 17. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 14 — Head, schematic drawing. 15 — Left maxillary palpus, lectotype. 16 — Right antenna, lectotype. 17 — Aedeagus dorsal view, lectotype." httpUri="https://zenodo.org/record/284499/files/figure.png" pageId="10" pageNumber="73">Fig. 17</figureCitation>
)
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long, ovoid, with parameres relatively long, sinuate and convergent, apically rounded and narrowed, bearing two bristles each. Internal sac symmetrical with a large median tooth.
</paragraph>
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<emphasis id="9EA516028C32B650FF2CFD9177D82B32" bold="true" box="[199,430,533,558]" pageId="10" pageNumber="73">Comparative notes</emphasis>
. The characters which distinguish this species from
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<emphasis id="9EA516028C32B650FBF5FD9272CB2B32" box="[1054,1213,533,558]" italics="true" pageId="10" pageNumber="73">T. longipalpis</emphasis>
</taxonomicName>
have already been highlighted in the discussion of the latter species.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C32B650FF2CFDDE741D2BA4" pageId="10" pageNumber="73" type="discussion">
<paragraph id="AC6ECA108C32B650FF2CFDDE741D2BA4" blockId="10.[151,1436,151,696]" pageId="10" pageNumber="73">
<emphasis id="9EA516028C32B650FF2CFDDE77422B6F" bold="true" box="[199,308,602,627]" pageId="10" pageNumber="73">Remarks</emphasis>
. Original label on the glade preparation of aedeagus reports reads:
<taxonomicName id="6BD1B1938C32B650FBC0FDDE73FD2B6F" authority="Bordei Pey. Type" authorityName="Bordei Pey. Type" box="[1067,1419,602,627]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C32B650FBC0FDDE73632B6F" box="[1067,1301,602,627]" italics="true" pageId="10" pageNumber="73">Anopsibythus Bordei</emphasis>
Pey.
<typeStatus id="736A74B28C32B650FAB8FDDF73FD2B6F" box="[1363,1419,603,627]" pageId="10" pageNumber="73">Type</typeStatus>
</taxonomicName>
- Bône, but really it is the aedeagus of
<taxonomicName id="6BD1B1938C32B650FDD2FDFA75F62B8A" authority="Jeannel 1956" authorityName="Jeannel" authorityYear="1956" box="[569,896,637,662]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="73" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C32B650FDD2FDFA74942B8A" box="[569,738,637,662]" italics="true" pageId="10" pageNumber="73">T. peyerimhoffi</emphasis>
Jeannel 1956
</taxonomicName>
, as can be seen from the same in Jeannel (1956: 83 fig. 55); male of
<taxonomicName id="6BD1B1938C32B650FE9CFD2577AC2BA4" box="[375,474,671,696]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="73" phylum="Arthropoda" rank="species" species="bordei">
<emphasis id="9EA516028C32B650FE9CFD2577AC2BA4" box="[375,474,671,696]" italics="true" pageId="10" pageNumber="73">T. bordei</emphasis>
</taxonomicName>
is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

188
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<mods:title id="A4D32C5C03339E57D75E3F317A7CCFC8">A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<heading id="F7267D7C8C33B651FF7CFA0775072C81" bold="true" box="[151,881,1411,1437]" fontSize="11" level="1" pageId="11" pageNumber="74" reason="1">
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Key to identification of
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<emphasis id="9EA516028C33B651FE57FA0774CD2C81" bold="true" box="[444,699,1411,1437]" italics="true" pageId="11" pageNumber="74">Tychobythinus theryi</emphasis>
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species group
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1. Head frontal lobe with antennal lobes separated and with conspicuous mediodorsal conical horn-like process bearing a long seta.....................................................................................
<emphasis id="9EA516028C33B651FAEAFA6273E32CE7" box="[1281,1429,1510,1531]" italics="true" pageId="11" pageNumber="74">
..
<taxonomicName id="6BD1B1938C33B651FAE7FA6273E32CE7" ID-CoL="7CYDJ" box="[1292,1429,1510,1531]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="monoceros">T. monoceros</taxonomicName>
</emphasis>
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<keyLead id="172069258C33B651FF7CF98073E32F05" box="[151,1429,1540,1561]" pageId="11" pageNumber="74">- Head frontal lobe without mediodorsal conical horn-like process............................................... 2</keyLead>
</paragraph>
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<keyLead id="172069258C33B651FF7CF9A573ED2F2A" box="[151,1435,1569,1590]" pageId="11" pageNumber="74">
2. Tempora prolonged into a conspicuous acute spine..................................................
<taxonomicName id="6BD1B1938C33B651FACEF9A573ED2F2A" ID-CoL="59SD4" box="[1317,1435,1569,1590]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="araneipes">
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<keyLead id="172069258C33B651FF7CF9BA73ED2F4F" box="[151,1435,1598,1619]" pageId="11" pageNumber="74">- Tempora not prolonged into a conspicuous acute spine........................................................ 3</keyLead>
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<keyLead id="172069258C33B651FF7CF9D873ED2F6D" box="[151,1435,1628,1649]" pageId="11" pageNumber="74">
3. Tempora angulated, clypeal carina reaching temporal region...........................................
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<emphasis id="9EA516028C33B651FACCF9D873ED2F6D" box="[1319,1435,1628,1649]" italics="true" pageId="11" pageNumber="74">T. normandi</emphasis>
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<keyLead id="172069258C33B651FF7CF9FD73ED2F92" box="[151,1435,1657,1678]" pageId="11" pageNumber="74">- Tempora rounded, clypeal carina reaching ocular region....................................................... 4</keyLead>
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<paragraph id="AC6ECA108C33B651FF7CF91273EA2FB7" blockId="11.[151,1436,1481,2030]" box="[151,1436,1686,1707]" pageId="11" pageNumber="74">
<keyLead id="172069258C33B651FF7CF91273EA2FB7" box="[151,1436,1686,1707]" pageId="11" pageNumber="74">
4. Occipital carina very short reaching the middle of occipital region, antennal scape 5 times longer than wide....
<taxonomicName id="6BD1B1938C33B651FAF3F91273EA2FB7" ID-CoL="7CYDC" box="[1304,1436,1686,1707]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="9EA516028C33B651FAF3F91273EA2FB7" box="[1304,1436,1686,1707]" italics="true" pageId="11" pageNumber="74">T. longipalpis</emphasis>
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<paragraph id="AC6ECA108C33B651FF7CF93073EA2FD5" blockId="11.[151,1436,1481,2030]" box="[151,1436,1716,1737]" pageId="11" pageNumber="74">
<keyLead id="172069258C33B651FF7CF93073EA2FD5" box="[151,1436,1716,1737]" pageId="11" pageNumber="74">- Occipital carina reaching at least the posterior margin of vertexal foveae, antennal scape at most 4 times longer than wide... 5</keyLead>
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<keyLead id="172069258C33B651FF7CF95573EA2E1F" pageId="11" pageNumber="74">
5. Antennae
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long with antennomeres III–VI wider than long, palpomere III sligthly longer than wide, surface of metasternum strongly punctated....................................................................
<taxonomicName id="6BD1B1938C33B651FAA7F96A73EA2E1F" ID-CoL="59SFL" box="[1356,1436,1774,1795]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="theryi">
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- Antennae at least
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long with antennomeres III–VI longer than wide, palpomere III distinctly longer than wide, sur- face of metasternum unpunctated......................................................................... 6
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6. Lateral margin of last segment of maxillary palpi sligtlhy concave, occipital carina very long reaching posterior edge of median sulcus of frontal lobe, antennomeres VII–VIII longer than wide, mesofemora with 4–5 tubercles..........
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<emphasis id="9EA516028C33B651FAADF8E073EA2E65" box="[1350,1436,1892,1913]" italics="true" pageId="11" pageNumber="74">T. bordei</emphasis>
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<keyLead id="172069258C33B651FF7CF80573ED2EAF" pageId="11" pageNumber="74">- Lateral margin of last segment of maxillary palpi straigth, occipital carina reaching posterior edge of vertexal foveae, antenno- meres VII–VIII as long as wide, mesofemora without tubercles................................................. 7</keyLead>
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<paragraph id="AC6ECA108C33B651FF7CF83873EA2ECD" blockId="11.[151,1436,1481,2030]" box="[151,1436,1980,2001]" pageId="11" pageNumber="74">
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7. Head frontal lobe
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wide, antennae
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long, aedeagus as in fig.17..................
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<emphasis id="9EA516028C33B651FAE0F83873EA2ECD" box="[1291,1436,1980,2001]" italics="true" pageId="11" pageNumber="74">T. peyerimhoffi</emphasis>
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</paragraph>
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<keyLead id="172069258C33B651FF7CF85D73EA2EF2" box="[151,1436,2008,2030]" pageId="11" pageNumber="74">
<emphasis id="9EA516028C33B651FF7CF85D76E92EF2" box="[151,159,2009,2030]" italics="true" pageId="11" pageNumber="74">-</emphasis>
Head frontal lobe
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wide, antennae
<quantity id="6B2967F58C33B651FD94F85D75572EF2" box="[639,801,2009,2030]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="11" pageNumber="74" unit="mm" value="0.655" valueMax="0.66" valueMin="0.65">0.650–0.660 mm</quantity>
long, aedeagus as in fig. 21...............
<taxonomicName id="6BD1B1938C33B651FB06F85D73162EF2" ID-CoL="59SD9" box="[1261,1376,2009,2030]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="9EA516028C33B651FB06F85D73162EF2" box="[1261,1376,2009,2030]" italics="true" pageId="11" pageNumber="74">T. besucheti</emphasis>
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<taxonomicNameLabel id="8596AB798C33B651FA8DF85C73EA2EF2" box="[1382,1436,2008,2030]" pageId="11" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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</treatment>
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203
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<mods:title id="8658B75553B5EF81A74A7CCC82F5BF24">A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<taxonomicName id="6BD1B1938C33B651FF7CFF3877B429CA" ID-CoL="59SD9" box="[151,450,188,214]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="9EA516028C33B651FF7CFF3877B429CA" bold="true" box="[151,450,188,214]" italics="true" pageId="11" pageNumber="74">Tychobythinus besucheti</emphasis>
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<taxonomicNameLabel id="8596AB798C33B651FE22FF38747D29CA" box="[457,523,188,214]" pageId="11" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
Sabella
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<paragraph id="AC6ECA108C33B651FF7CFF5A775B29EB" blockId="11.[151,622,188,247]" box="[151,301,222,247]" pageId="11" pageNumber="74">
(
<figureCitation id="34EAD6958C33B651FF74FF5A775029EB" box="[159,294,222,247]" captionStart="FIGURES 18 – 21" captionStartId="10.[151,264,1981,2003]" captionTargetBox="[161,1414,749,1948]" captionTargetId="figure@10.[152,1435,726,1959]" captionTargetPageId="10" captionText="FIGURES 18 – 21. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 18 — Head, schematic drawing. 19 — Left maxillary palpus, holotype. 20 — Right antenna, holotype. 21 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284500/files/figure.png" pageId="11" pageNumber="74">Figs. 18–21</figureCitation>
)
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</subSubSection>
<subSubSection id="E4CB999B8C33B651FF7CFEA272C7287F" pageId="11" pageNumber="74" type="materials_examined">
<paragraph id="AC6ECA108C33B651FF7CFEA272C7287F" blockId="11.[151,1436,294,1327]" pageId="11" pageNumber="74">
<emphasis id="9EA516028C33B651FF7CFEA277302823" bold="true" box="[151,326,294,319]" pageId="11" pageNumber="74">
<typeStatus id="736A74B28C33B651FF7CFEA276A42823" box="[151,210,294,319]" pageId="11" pageNumber="74">Type</typeStatus>
material
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.
<emphasis id="9EA516028C33B651FEB6FEA277BC2823" bold="true" box="[349,458,294,319]" pageId="11" pageNumber="74">
<typeStatus id="736A74B28C33B651FEB6FEA277BC2823" box="[349,458,294,319]" pageId="11" pageNumber="74" type="holotype">Holotype</typeStatus>
</emphasis>
, male.
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NORTHEASTERN
<collectingCountry id="D4C68A808C33B651FCC2FEA275DD2823" box="[809,939,294,319]" name="Algeria" pageId="11" pageNumber="74">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C33B651FC29FEA272EF2823" bold="true" box="[962,1177,294,319]" pageId="11" pageNumber="74">Annaba province</emphasis>
: Bône (old name of Annaba),
<date id="D86FECD08C33B651FEEEFECF7714287F" box="[261,354,331,355]" pageId="11" pageNumber="74" value="1930-11">XI.1930</date>
(H. Normand) (MHNG).
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<typeStatus id="736A74B28C33B651FD6FFECE7487287F" box="[644,753,330,355]" pageId="11" pageNumber="74" type="paratype">Paratype</typeStatus>
</emphasis>
, male, same data of
<typeStatus id="736A74B28C33B651FC33FECE724C287F" box="[984,1082,330,355]" pageId="11" pageNumber="74" type="holotype">holotype</typeStatus>
(MHNG).
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C33B651FF2CFEEA76852C33" pageId="11" pageNumber="74" type="description">
<paragraph id="AC6ECA108C33B651FF2CFEEA746928D3" blockId="11.[151,1436,294,1327]" pageId="11" pageNumber="74">
<emphasis id="9EA516028C33B651FF2CFEEA7727289B" bold="true" box="[199,337,366,391]" pageId="11" pageNumber="74">Description</emphasis>
. Body length
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, pubescence consists of long and flattened setae (
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) on head, pronotum, elytra and abdomen, other suberected setae shorter (length:
<quantity id="6B2967F58C33B651FBFDFE1772A028B7" box="[1046,1238,403,427]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="11" pageNumber="74" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.030–0.040 mm</quantity>
) on antennae and legs. Tegument smooth and glossy.
</paragraph>
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Head (
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) longer (
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) than wide (0.200 mm), frontal lobe wide
<quantity id="6B2967F58C33B651FBF3FE5F72FF28EE" box="[1048,1161,475,499]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="74" unit="mm" value="0.11">0.110 mm</quantity>
its dorsal surface glossy with some punctures and well impressed median sulcus. Clypeal carina evident, reaching ocular region, well visibile median occipital carina reaching posterior edge of vertexal foveae, tempora rounded. Gular region convex entirely traversed by evident median longitudinal carina, its surface glossy. Segment II of maxillary palpi with 24– 26 tubercles, segment III longer than wide with 4–6 tubercles, last segment of maxillary palpi (
<figureCitation id="34EAD6958C33B651FB53FDEE737E2B9F" box="[1208,1288,618,643]" captionStart="FIGURES 18 – 21" captionStartId="10.[151,264,1981,2003]" captionTargetBox="[161,1414,749,1948]" captionTargetId="figure@10.[152,1435,726,1959]" captionTargetPageId="10" captionText="FIGURES 18 – 21. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 18 — Head, schematic drawing. 19 — Left maxillary palpus, holotype. 20 — Right antenna, holotype. 21 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284500/files/figure.png" pageId="11" pageNumber="74">Fig. 19</figureCitation>
) more than 3 times longer (
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) than wide (
<quantity id="6B2967F58C33B651FD6EFD0A748B2BBB" box="[645,765,654,679]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="74" unit="mm" value="0.075">0.075 mm</quantity>
), its lateral margin straight. Antenna (
<figureCitation id="34EAD6958C33B651FB47FD0A72892BBB" box="[1196,1279,654,679]" captionStart="FIGURES 18 – 21" captionStartId="10.[151,264,1981,2003]" captionTargetBox="[161,1414,749,1948]" captionTargetId="figure@10.[152,1435,726,1959]" captionTargetPageId="10" captionText="FIGURES 18 – 21. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 18 — Head, schematic drawing. 19 — Left maxillary palpus, holotype. 20 — Right antenna, holotype. 21 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284500/files/figure.png" pageId="11" pageNumber="74">Fig. 20</figureCitation>
)
<quantity id="6B2967F58C33B651FAFBFD0A76B52BD6" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="11" pageNumber="74" unit="mm" value="0.655" valueMax="0.66" valueMin="0.65">0.650–0.660 mm</quantity>
long with scape more than 4 times longer (
<quantity id="6B2967F58C33B651FD57FD3675F62BD7" box="[700,896,690,715]" metricMagnitude="-4" metricUnit="m" metricValue="1.775" metricValueMax="1.7999999999999998" metricValueMin="1.75" pageId="11" pageNumber="74" unit="mm" value="0.1775" valueMax="0.18" valueMin="0.175">0.175–0.180 mm</quantity>
) than wide (
<quantity id="6B2967F58C33B651FBF9FD3772AC2BD7" box="[1042,1242,691,715]" metricMagnitude="-5" metricUnit="m" metricValue="3.85" metricValueMax="3.9" metricValueMin="3.8" pageId="11" pageNumber="74" unit="mm" value="0.0385" valueMax="0.039" valueMin="0.038">0.038–0.039 mm</quantity>
), antennomere II slightly ovoid and asymmetric, longer (0.600 mm) than wide (
<quantity id="6B2967F58C33B651FCB9FD5275BC2BF3" box="[850,970,726,751]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="74" unit="mm" value="0.35">0.350 mm</quantity>
), antennomeres III–VII longer than wide, III distinctly longer than wide, IV and V subequal in size, slightly shorter than III, VI slightly longer than wide, VII longer than VI, VIII as long as wide. Antennomeres IX–X distinctly wider than long, XI about 2 times longer than wide, and more than 1.5 times longer than combined length of IX and X.
</paragraph>
<paragraph id="AC6ECA108C33B651FF2CFCE3743D2ABF" blockId="11.[151,1436,294,1327]" pageId="11" pageNumber="74">
Pronotum slightly wider (0.300 mm) than long (
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), widest in forequater, pleural carina well visible, not very deep antebasal sulcus.
</paragraph>
<paragraph id="AC6ECA108C33B651FF2CFC2A754F2ADA" blockId="11.[151,1436,294,1327]" box="[199,825,942,967]" pageId="11" pageNumber="74">
Elytra width
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, lenght
<quantity id="6B2967F58C33B651FD99FC2B75432ADA" box="[626,821,943,967]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="3.9" metricValueMin="3.8" pageId="11" pageNumber="74" unit="mm" value="0.385" valueMax="0.39" valueMin="0.38">0.380–0.390 mm</quantity>
.
</paragraph>
<paragraph id="AC6ECA108C33B651FF2CFC5774BE2AF7" blockId="11.[151,1436,294,1327]" box="[199,712,978,1003]" pageId="11" pageNumber="74">Metasternum with surface smooth and glossy.</paragraph>
<paragraph id="AC6ECA108C33B651FF2CFC7375152D2F" blockId="11.[151,1436,294,1327]" pageId="11" pageNumber="74">
Legs elongated, protrochanters with 4–6 tubercles; profemora with 12–14 small tubercles, article II of protarsi slightly compressed laterally. Metatibiae
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long.
</paragraph>
<paragraph id="AC6ECA108C33B651FF2CFBBA72352D66" blockId="11.[151,1436,294,1327]" pageId="11" pageNumber="74">
Aedeagus (
<figureCitation id="34EAD6958C33B651FEA2FBBA77E92D4B" box="[329,415,1086,1111]" captionStart="FIGURES 18 – 21" captionStartId="10.[151,264,1981,2003]" captionTargetBox="[161,1414,749,1948]" captionTargetId="figure@10.[152,1435,726,1959]" captionTargetPageId="10" captionText="FIGURES 18 – 21. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 18 — Head, schematic drawing. 19 — Left maxillary palpus, holotype. 20 — Right antenna, holotype. 21 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284500/files/figure.png" pageId="11" pageNumber="74">Fig. 21</figureCitation>
)
<quantity id="6B2967F58C33B651FE58FBBB74002D4A" box="[435,630,1087,1111]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="11" pageNumber="74" unit="mm" value="0.325" valueMax="0.33" valueMin="0.32">0.320–0.330 mm</quantity>
long ovoid with parameres relatively long, sinuate and convergent, apically rounded and narrowed, bearing two bristles each. Symmetrical internal sac.
</paragraph>
<paragraph id="AC6ECA108C33B651FF2CFB0277EB2D83" blockId="11.[151,1436,294,1327]" box="[199,413,1158,1183]" pageId="11" pageNumber="74">Female. Unknown.</paragraph>
<paragraph id="AC6ECA108C33B651FF2CFB2E76852C33" blockId="11.[151,1436,294,1327]" pageId="11" pageNumber="74">
<emphasis id="9EA516028C33B651FF2CFB2E77DF2DDF" bold="true" box="[199,425,1194,1219]" pageId="11" pageNumber="74">Comparative notes</emphasis>
.
<taxonomicName id="6BD1B1938C33B651FE53FB2874482DDF" box="[440,574,1194,1219]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="9EA516028C33B651FE53FB2874482DDF" box="[440,574,1194,1219]" italics="true" pageId="11" pageNumber="74">T. besucheti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8596AB798C33B651FDAEFB2F74F72DDE" box="[581,641,1195,1218]" pageId="11" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
is very similar to
<taxonomicName id="6BD1B1938C33B651FCA5FB2875812DDF" box="[846,1015,1194,1219]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C33B651FCA5FB2875812DDF" box="[846,1015,1194,1219]" italics="true" pageId="11" pageNumber="74">T. peyerimhoffi</emphasis>
</taxonomicName>
from which it is distinguished by the narrower head frontal lobe (width
<quantity id="6B2967F58C33B651FDC0FB4B74E42DFA" box="[555,658,1231,1255]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="74" unit="mm" value="0.11">0.11 mm</quantity>
in n. sp.,
<quantity id="6B2967F58C33B651FCE4FB4A75F32DFA" box="[783,901,1230,1255]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="74" unit="mm" value="0.125">0.125 mm</quantity>
in
<taxonomicName id="6BD1B1938C33B651FC5BFB4A724B2DFB" box="[944,1085,1230,1255]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C33B651FC5BFB4A724B2DFB" box="[944,1085,1230,1255]" italics="true" pageId="11" pageNumber="74">peyerimhoffi</emphasis>
</taxonomicName>
), the shorter antennae (length
<quantity id="6B2967F58C33B651FF7CFB7677212C16" box="[151,343,1266,1291]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="11" pageNumber="74" unit="mm" value="0.655" valueMax="0.66" valueMin="0.65">0.650–0.660 mm</quantity>
in
<taxonomicName id="6BD1B1938C33B651FE96FB7074722C17" box="[381,516,1266,1291]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="9EA516028C33B651FE96FB7074722C17" box="[381,516,1266,1291]" italics="true" pageId="11" pageNumber="74">T. besucheti</emphasis>
</taxonomicName>
<quantity id="6B2967F58C33B651FDE7FB7674BA2C16" box="[524,716,1266,1291]" metricMagnitude="-4" metricUnit="m" metricValue="6.8500000000000005" metricValueMax="6.8999999999999995" metricValueMin="6.800000000000001" pageId="11" pageNumber="74" unit="mm" value="0.685" valueMax="0.69" valueMin="0.68">0.680–0.690 mm</quantity>
in
<taxonomicName id="6BD1B1938C33B651FD19FB7075EA2C17" box="[754,924,1266,1291]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="74" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C33B651FD19FB7075EA2C17" box="[754,924,1266,1291]" italics="true" pageId="11" pageNumber="74">T. peyerimhoffi</emphasis>
</taxonomicName>
) and by the features of adeagus (cfr.
<figureCitation id="34EAD6958C33B651FAACFB7673EA2C17" box="[1351,1436,1266,1291]" captionStart="FIGURES 14 – 17" captionStartId="9.[151,264,1907,1929]" captionTargetBox="[171,1415,639,1875]" captionTargetId="figure@9.[151,1435,625,1897]" captionTargetPageId="9" captionText="FIGURES 14 – 17. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 14 — Head, schematic drawing. 15 — Left maxillary palpus, lectotype. 16 — Right antenna, lectotype. 17 — Aedeagus dorsal view, lectotype." httpUri="https://zenodo.org/record/284499/files/figure.png" pageId="11" pageNumber="74">Figs 17</figureCitation>
and
<figureCitation id="34EAD6958C33B651FF2DFA9376932C33" box="[198,229,1303,1327]" captionStart="FIGURES 18 – 21" captionStartId="10.[151,264,1981,2003]" captionTargetBox="[161,1414,749,1948]" captionTargetId="figure@10.[152,1435,726,1959]" captionTargetPageId="10" captionText="FIGURES 18 – 21. Tychobythinus peyerimhoffi. Scale: 0, 1 mm. 18 — Head, schematic drawing. 19 — Left maxillary palpus, holotype. 20 — Right antenna, holotype. 21 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284500/files/figure.png" pageId="11" pageNumber="74">21</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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395
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<mods:title id="DFEF5B990CA5D0B7DC4A663EEF9FA5B3">A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<subSubSection id="E4CB999B8C39B65BFF7CF9B274822FAB" pageId="1" pageNumber="64" type="nomenclature">
<paragraph id="AC6ECA108C39B65BFF7CF9B274F52F4C" blockId="1.[151,643,1590,1650]" box="[151,643,1590,1616]" pageId="1" pageNumber="64">
<heading id="F7267D7C8C39B65BFF7CF9B274F52F4C" bold="true" box="[151,643,1590,1616]" fontSize="11" level="1" pageId="1" pageNumber="64" reason="1">
<taxonomicName id="6BD1B1938C39B65BFF7CF9B274F52F4C" ID-CoL="59SFL" authority="Guillebeau, 1894" authorityName="Guillebeau" authorityYear="1894" box="[151,643,1590,1616]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C39B65BFF7CF9B274F52F4C" bold="true" box="[151,643,1590,1616]" pageId="1" pageNumber="64">
<emphasis id="9EA516028C39B65BFF7CF9B277E02F4C" bold="true" box="[151,406,1590,1616]" italics="true" pageId="1" pageNumber="64">Tychobythinus theryi</emphasis>
(Guillebeau, 1894)
</emphasis>
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<paragraph id="AC6ECA108C39B65BFF7CF9DD77672F6E" blockId="1.[151,643,1590,1650]" box="[151,273,1625,1650]" pageId="1" pageNumber="64">
(
<figureCitation id="34EAD6958C39B65BFF74F9DD777E2F6E" box="[159,264,1625,1650]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1538,1560]" captionTargetBox="[393,1202,286,1496]" captionTargetId="figure@2.[369,1218,271,1517]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Tychobythinus theryi. Scale: 0, 1 mm. 1 — Head, schematic drawing. 2 — Left maxillary palpus, lectotype. 3 — Right antenna, lectotype." httpUri="https://zenodo.org/record/284495/files/figure.png" pageId="1" pageNumber="64">Figs. 1–3</figureCitation>
)
</paragraph>
<paragraph id="AC6ECA108C39B65BFF7CF92574822FAB" blockId="1.[151,1311,1697,1842]" box="[151,756,1697,1719]" pageId="1" pageNumber="64">
<taxonomicName id="6BD1B1938C39B65BFF7CF92574662FAB" authority="Theryi Guillebeau 1894" authorityName="Theryi Guillebeau" authorityYear="1894" box="[151,528,1697,1719]" class="Insecta" family="Staphylinidae" genus="Machaerites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C39B65BFF7CF925772B2FAB" box="[151,349,1697,1719]" italics="true" pageId="1" pageNumber="64">Machaerites Theryi</emphasis>
Guillebeau 1894
</taxonomicName>
a: CCXCII; 1894b: 47;
</paragraph>
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<paragraph id="AC6ECA108C39B65BFF7CF945741F2FCA" blockId="1.[151,1311,1697,1842]" box="[151,617,1728,1750]" pageId="1" pageNumber="64">
<taxonomicName id="6BD1B1938C39B65BFF7CF94576922FCA" box="[151,228,1729,1750]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C39B65BFF7CF94576922FCA" box="[151,228,1729,1750]" italics="true" pageId="1" pageNumber="64">Bryaxis</emphasis>
</taxonomicName>
(groupe V)
<taxonomicName id="6BD1B1938C39B65BFE8BF94474132FCA" authority="Raffray 1904: 276" authorityName="Raffray" authorityPageNumber="276" authorityYear="1904" box="[352,613,1728,1750]" class="Insecta" family="Staphylinidae" genus="Theryi" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C39B65BFE8BF94477D42FCA" box="[352,418,1728,1750]" italics="true" pageId="1" pageNumber="64">Theryi</emphasis>
Raffray 1904: 276
</taxonomicName>
;
</paragraph>
<paragraph id="AC6ECA108C39B65BFF7CF95A74822FE8" blockId="1.[151,1311,1697,1842]" box="[151,756,1758,1780]" pageId="1" pageNumber="64">
<taxonomicName id="6BD1B1938C39B65BFF7CF95A74612FE8" authority="Theryi Raffray 1908: 284" authorityName="Theryi Raffray" authorityPageNumber="284" authorityYear="1908" box="[151,535,1758,1780]" class="Insecta" family="Staphylinidae" genus="Bythinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C39B65BFF7CF95A77232FE8" box="[151,341,1758,1780]" italics="true" pageId="1" pageNumber="64">Bythinopsis Theryi</emphasis>
Raffray 1908: 284
</taxonomicName>
; 1911: 125; 1924: 85;
</paragraph>
<paragraph id="AC6ECA108C39B65BFF7CF97972D72E2E" blockId="1.[151,1311,1697,1842]" pageId="1" pageNumber="64">
<taxonomicName id="6BD1B1938C39B65BFF7CF979746A2E0F" authority="Theryi Jeannel 1956: 60" authorityName="Theryi Jeannel" authorityPageNumber="60" authorityYear="1956" box="[151,540,1789,1811]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C39B65BFF7CF97977112E0F" box="[151,359,1789,1811]" italics="true" pageId="1" pageNumber="64">Anopsibythus Theryi</emphasis>
Jeannel 1956: 60
</taxonomicName>
, 62, fig. 54 (habitus, the caption wrongly reports: male from Saint-Charles);
<taxonomicName id="6BD1B1938C39B65BFF7CF898744A2E2E" authority="Besuchet 1962: 349" authorityName="Besuchet" authorityPageNumber="349" authorityYear="1962" box="[151,572,1820,1842]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C39B65BFF7CF898771C2E2E" box="[151,362,1820,1842]" italics="true" pageId="1" pageNumber="64">Tychobythinus theryi</emphasis>
Besuchet 1962: 349
</taxonomicName>
; Newton &amp; Chandler 1989: 48; Löbl &amp; Besuchet 2004: 315.
</paragraph>
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<paragraph id="AC6ECA108C39B65BFF7CF8DA74112EA3" blockId="1.[151,1436,1886,1983]" pageId="1" pageNumber="64">
<emphasis id="9EA516028C39B65BFF7CF8DA774B2E6B" bold="true" box="[151,317,1886,1911]" pageId="1" pageNumber="64">
<typeStatus id="736A74B28C39B65BFF7CF8DA76A42E6B" box="[151,210,1886,1911]" pageId="1" pageNumber="64">Type</typeStatus>
material
</emphasis>
.
<emphasis id="9EA516028C39B65BFEA0F8DA77B72E6B" bold="true" box="[331,449,1886,1911]" pageId="1" pageNumber="64">
<typeStatus id="736A74B28C39B65BFEA0F8DA77B72E6B" box="[331,449,1886,1911]" pageId="1" pageNumber="64" type="lectotype">Lectotype</typeStatus>
</emphasis>
, female, present designation.
<emphasis id="9EA516028C39B65BFCE4F8DA72F22E6B" bold="true" box="[783,1156,1886,1911]" pageId="1" pageNumber="64">
NORTHEASTERN
<collectingCountry id="D4C68A808C39B65BFBEAF8DA72F22E6B" box="[1025,1156,1886,1911]" name="Algeria" pageId="1" pageNumber="64">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C39B65BFB7AF8DA73222E6B" bold="true" box="[1169,1364,1886,1911]" pageId="1" pageNumber="64">Skikda province</emphasis>
: Saint Charles, (A.
<taxonomicName id="6BD1B1938C39B65BFEC3F806771B2E87" box="[296,365,1922,1947]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="species" species="thery">Théry</taxonomicName>
) (ex coll. Peyerimhoff, MNHN).
<emphasis id="9EA516028C39B65BFD02F80675EF2E87" bold="true" box="[745,921,1922,1947]" pageId="1" pageNumber="64">
<typeStatus id="736A74B28C39B65BFD02F80675EF2E87" box="[745,921,1922,1947]" pageId="1" pageNumber="64" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
,
<specimenCount id="BAD701998C39B65BFC4CF80772612E87" box="[935,1047,1922,1947]" pageId="1" pageNumber="64" type="female">2 females</specimenCount>
, present designation, same data of
<typeStatus id="736A74B28C39B65BFF7CF82276882EA3" box="[151,254,1958,1983]" pageId="1" pageNumber="64" type="lectotype">lectotype</typeStatus>
(ex coll. Peyerimhoff, MNHN).
</paragraph>
<paragraph id="AC6ECA108C3AB658FF2CFF1375EB29EB" blockId="2.[151,1436,151,248]" pageId="2" pageNumber="65">
<emphasis id="9EA516028C3AB658FF2CFF13771B29AC" bold="true" box="[199,365,151,176]" pageId="2" pageNumber="65">Redescription</emphasis>
. Body length
<quantity id="6B2967F58C3AB658FDFFFF1374CA29B3" box="[532,700,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="2" pageNumber="65" unit="mm" value="1.2" valueMax="1.25" valueMin="1.15">1.15–1.25 mm</quantity>
, pubescence not very dense consists of long and flattened setae (length:
<quantity id="6B2967F58C3AB658FF1EFF3877ED29C8" box="[245,411,188,212]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7" pageId="2" pageNumber="65" unit="mm" value="0.075" valueMax="0.08" valueMin="0.07">0.07–0.08 mm</quantity>
) on head, pronotum, elytra and abdomen, other suberected setae shorter (length:
<quantity id="6B2967F58C3AB658FAC7FF3876B229E4" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="65" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.03–0.04 mm</quantity>
) on antennae and legs. Tegument slightly but distinctly punctate.
</paragraph>
<caption id="F8AE9A988C3AB658FF7CF98677E32F2B" httpUri="https://zenodo.org/record/284495/files/figure.png" pageId="2" pageNumber="65" targetBox="[393,1202,286,1496]" targetPageId="2">
<paragraph id="AC6ECA108C3AB658FF7CF98677E32F2B" blockId="2.[151,1435,1538,1591]" pageId="2" pageNumber="65">
<emphasis id="9EA516028C3AB658FF7CF986774D2F04" bold="true" box="[151,315,1538,1560]" pageId="2" pageNumber="65">FIGURES 1–3.</emphasis>
<taxonomicName id="6BD1B1938C3AB658FEA9F98674602F04" box="[322,534,1538,1560]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3AB658FEA9F98674602F04" box="[322,534,1538,1560]" italics="true" pageId="2" pageNumber="65">Tychobythinus theryi</emphasis>
</taxonomicName>
. Scale: 0,1 mm. 1—Head, schematic drawing. 2—Left maxillary palpus, lectotype. 3— Right antenna, lectotype.
</paragraph>
</caption>
<paragraph id="AC6ECA108C3AB658FF2CF9DA73142EC3" blockId="2.[151,1436,1630,2015]" pageId="2" pageNumber="65">
Head (
<figureCitation id="34EAD6958C3AB658FEF8F9DA77232F6B" box="[275,341,1630,1655]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1538,1560]" captionTargetBox="[393,1202,286,1496]" captionTargetId="figure@2.[369,1218,271,1517]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Tychobythinus theryi. Scale: 0, 1 mm. 1 — Head, schematic drawing. 2 — Left maxillary palpus, lectotype. 3 — Right antenna, lectotype." httpUri="https://zenodo.org/record/284495/files/figure.png" pageId="2" pageNumber="65">Fig. 1</figureCitation>
) wider (
<quantity id="6B2967F58C3AB658FE52F9DA740B2F6B" box="[441,637,1630,1655]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.25" metricValueMin="2.15" pageId="2" pageNumber="65" unit="mm" value="0.22" valueMax="0.225" valueMin="0.215">0.215–0.225 mm</quantity>
) than long (
<quantity id="6B2967F58C3AB658FCE3F9DA75B92F6B" box="[776,975,1630,1655]" metricMagnitude="-4" metricUnit="m" metricValue="1.825" metricValueMax="1.9" metricValueMin="1.75" pageId="2" pageNumber="65" unit="mm" value="0.1825" valueMax="0.19" valueMin="0.175">0.175–0.190 mm</quantity>
), frontal lobe width
<quantity id="6B2967F58C3AB658FB53F9DA730F2F6B" box="[1208,1401,1630,1655]" metricMagnitude="-4" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="2" pageNumber="65" unit="mm" value="0.1275" valueMax="0.13" valueMin="0.125">0.125–0.130 mm</quantity>
its dorsal surface glossy with some punctures concentrated on sides and shallow median sulcus. Clypeal carina reaches ocular region, occipital carina reaches over posterior edge of vertexal foveae, tempora rounded. Gular region flattened, anteriorly bounded by a deep transverse depression and lacking median longitudinal carina, its surface glossy. Segment II of maxillary palpi with 24–30 tubercles, segment III slightly longer than wide with 4–6 tubercles, apical segment of maxillary palpi (
<figureCitation id="34EAD6958C3AB658FD7CF89674AA2E37" box="[663,732,1810,1835]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1538,1560]" captionTargetBox="[393,1202,286,1496]" captionTargetId="figure@2.[369,1218,271,1517]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Tychobythinus theryi. Scale: 0, 1 mm. 1 — Head, schematic drawing. 2 — Left maxillary palpus, lectotype. 3 — Right antenna, lectotype." httpUri="https://zenodo.org/record/284495/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
) more than 3 times longer (
<quantity id="6B2967F58C3AB658FBF2F89672F92E37" box="[1049,1167,1810,1835]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" pageId="2" pageNumber="65" unit="mm" value="0.225">0.225 mm</quantity>
) than wide (
<quantity id="6B2967F58C3AB658FAF5F89673E22E37" box="[1310,1428,1810,1835]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="65" unit="mm" value="0.065">0.065 mm</quantity>
) with lateral margin straight. Antenna (
<figureCitation id="34EAD6958C3AB658FDB9F8B274EE2E53" box="[594,664,1846,1871]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1538,1560]" captionTargetBox="[393,1202,286,1496]" captionTargetId="figure@2.[369,1218,271,1517]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Tychobythinus theryi. Scale: 0, 1 mm. 1 — Head, schematic drawing. 2 — Left maxillary palpus, lectotype. 3 — Right antenna, lectotype." httpUri="https://zenodo.org/record/284495/files/figure.png" pageId="2" pageNumber="65">Fig. 3</figureCitation>
)
<quantity id="6B2967F58C3AB658FD40F8B275182E53" box="[683,878,1846,1871]" metricMagnitude="-4" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="2" pageNumber="65" unit="mm" value="0.555" valueMax="0.56" valueMin="0.55">0.550–0.560 mm</quantity>
long with scape about 3.5 times longer (
<quantity id="6B2967F58C3AB658FAA6F8B2777D2E6F" metricMagnitude="-4" metricUnit="m" metricValue="1.575" metricValueMax="1.6" metricValueMin="1.55" pageId="2" pageNumber="65" unit="mm" value="0.1575" valueMax="0.16" valueMin="0.155">0.155– 0.160 mm</quantity>
) than wide (
<quantity id="6B2967F58C3AB658FE77F8DE74142E6F" box="[412,610,1882,1907]" metricMagnitude="-5" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.8" metricValueMin="4.6" pageId="2" pageNumber="65" unit="mm" value="0.047" valueMax="0.048" valueMin="0.046">0.046–0.048 mm</quantity>
), antennomere II slightly ovoid and asymmetric longer (
<quantity id="6B2967F58C3AB658FB0EF8DF732D2E6F" box="[1253,1371,1883,1907]" metricMagnitude="-5" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="65" unit="mm" value="0.049">0.049 mm</quantity>
) than wide (
<quantity id="6B2967F58C3AB658FF30F8FB77D62E8B" box="[219,416,1919,1943]" metricMagnitude="-5" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="2" pageNumber="65" unit="mm" value="0.0395" valueMax="0.04" valueMin="0.039">0.039–0.040 mm</quantity>
), antennomere III longer than wide, antennomeres IV and V subequal in size and as long as wide, VI–VIII shorter than previous subequal in size and wider than long, antennomeres IX and X distinctly transverse, XI about two times longer than wide, and about 2 times longer than combined length of IX and X.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CFF1C744329C9" blockId="3.[151,1436,151,645]" pageId="3" pageNumber="66">
Pronotum wider (0.300 mm) than long (
<quantity id="6B2967F58C3BB659FD71FF1C751529AC" box="[666,867,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="3" pageNumber="66" unit="mm" value="0.235" valueMax="0.24" valueMin="0.23">0.230–0.240 mm</quantity>
), widest a little before the middle, pleural carina barely visible, deep antebasal sulcus.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CFF5B749A29EB" blockId="3.[151,1436,151,645]" box="[199,748,223,248]" pageId="3" pageNumber="66">
Elytra width 0.500 mm, lenght
<quantity id="6B2967F58C3BB659FDCEFF5B749E29EB" box="[549,744,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="4.875" metricValueMax="4.95" metricValueMin="4.8" pageId="3" pageNumber="66" unit="mm" value="0.4875" valueMax="0.495" valueMin="0.48">0.480–0.495 mm</quantity>
.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CFE8175D92800" blockId="3.[151,1436,151,645]" box="[199,943,260,285]" pageId="3" pageNumber="66">Metasternum with surface impressed by large and thick punctures.</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CFEAC75E42879" blockId="3.[151,1436,151,645]" pageId="3" pageNumber="66">
Legs not particularly elongated, protrochanters with 4–6 larges and prominent tubercles; profemora with 14– 16 large and prominent tubercles. Metatibiae 0.400–
<quantity id="6B2967F58C3BB659FD0AFEC875242878" box="[737,850,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="66" unit="mm" value="0.41">0.410 mm</quantity>
long.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CFEEB77F32894" blockId="3.[151,1436,151,645]" box="[199,389,367,392]" pageId="3" pageNumber="66">Male. Unknown.</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CFE1077B92B98" blockId="3.[151,1436,151,645]" pageId="3" pageNumber="66">
<emphasis id="9EA516028C3BB659FF2CFE1077DC28B1" bold="true" box="[199,426,404,429]" pageId="3" pageNumber="66">Comparative notes</emphasis>
.
<taxonomicName id="6BD1B1938C3BB659FE51FE11746E28B1" box="[442,536,404,429]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3BB659FE51FE11746E28B1" box="[442,536,404,429]" italics="true" pageId="3" pageNumber="66">T. theryi</emphasis>
</taxonomicName>
is distinguished from other species of the
<taxonomicName id="6BD1B1938C3BB659FC14FE10723628B1" box="[1023,1088,404,429]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3BB659FC14FE10723628B1" box="[1023,1088,404,429]" italics="true" pageId="3" pageNumber="66">theryi</emphasis>
</taxonomicName>
group by the shorter antennae (
<quantity id="6B2967F58C3BB659FF75FE33771628D3" box="[158,352,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="3" pageNumber="66" unit="mm" value="0.555" valueMax="0.56" valueMin="0.55">0.550–0.560 mm</quantity>
long, at least
<quantity id="6B2967F58C3BB659FE16FE33740628D3" box="[509,624,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="66" unit="mm" value="0.66">0.660 mm</quantity>
in other species of this group) and by the segment III of maxillary palpi slightly longer than wide (distinctly longer than wide in other species of
<taxonomicName id="6BD1B1938C3BB659FC2BFE58727728E9" box="[960,1025,476,501]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3BB659FC2BFE58727728E9" box="[960,1025,476,501]" italics="true" pageId="3" pageNumber="66">theryi</emphasis>
</taxonomicName>
group). It is similar to
<taxonomicName id="6BD1B1938C3BB659FAE8FE5973EA28E9" box="[1283,1436,476,501]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="9EA516028C3BB659FAE8FE5973EA28E9" box="[1283,1436,476,501]" italics="true" pageId="3" pageNumber="66">T. longipalpis</emphasis>
</taxonomicName>
by having short and flattened setae (
<quantity id="6B2967F58C3BB659FDC5FD8474992B0B" box="[558,751,512,536]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7" pageId="3" pageNumber="66" unit="mm" value="0.075" valueMax="0.08" valueMin="0.07">0.070–0.080 mm</quantity>
against
<quantity id="6B2967F58C3BB659FCA5FE7B727B2B0B" box="[846,1037,511,536]" metricMagnitude="-4" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="3" pageNumber="66" unit="mm" value="0.1275" valueMax="0.13" valueMin="0.125">0.125–0.130 mm</quantity>
in other species of this group), and the gular region not modified (with median longitudinal carina, tubercles or excavations in other species of
<taxonomicName id="6BD1B1938C3BB659FAB0FDA073EA2B21" box="[1371,1436,548,573]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3BB659FAB0FDA073EA2B21" box="[1371,1436,548,573]" italics="true" pageId="3" pageNumber="66">theryi</emphasis>
</taxonomicName>
group). It is also similar to
<taxonomicName id="6BD1B1938C3BB659FE20FDCD74232B7C" box="[459,597,583,608]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3BB659FE20FDCD74232B7C" box="[459,597,583,608]" italics="true" pageId="3" pageNumber="66">T. normandi</emphasis>
</taxonomicName>
for the surface of metasternum distinctly punctated (smooth and glossy in other species of this group).
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3BB659FF7CFD5674E12AAC" pageId="3" pageNumber="66" type="description">
<paragraph id="AC6ECA108C3BB659FF7CFD5677642A13" blockId="3.[151,644,722,783]" pageId="3" pageNumber="66">
<taxonomicName id="6BD1B1938C3BB659FF7CFD5674F22BF7" authority="Peyerimhoff, 1919" authorityName="Peyerimhoff" authorityYear="1919" box="[151,644,722,747]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="bordei">
<emphasis id="9EA516028C3BB659FF7CFD56740D2BF7" bold="true" box="[151,635,722,747]" pageId="3" pageNumber="66">
<emphasis id="9EA516028C3BB659FF7CFD5677E72BF7" bold="true" box="[151,401,722,747]" italics="true" pageId="3" pageNumber="66">Tychobythinus bordei</emphasis>
(Peyerimhoff, 1919
</emphasis>
)
</taxonomicName>
(
<figureCitation id="34EAD6958C3BB659FF74FD72777F2A13" box="[159,265,758,783]" captionStart="FIGURES 4 – 6" captionStartId="4.[151,264,1807,1829]" captionTargetBox="[397,1208,182,1757]" captionTargetId="figure@4.[355,1232,167,1774]" captionTargetPageId="4" captionText="FIGURES 4 – 6. Tychobythinus bordei. Scale: 0, 1 mm. 4 — Head, schematic drawing. 5 — Left maxillary palpus, lectotype. 6 — Right antenna, lectotype." httpUri="https://zenodo.org/record/284496/files/figure.png" pageId="3" pageNumber="66">Figs. 4–6</figureCitation>
)
</paragraph>
<paragraph id="AC6ECA108C3BB659FF7CFCBA746B2A6F" blockId="3.[151,994,830,944]" pageId="3" pageNumber="66">
<taxonomicName id="6BD1B1938C3BB659FF7CFCBA77602A48" box="[151,278,830,852]" class="Insecta" family="Staphylinidae" genus="Machaerites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3BB659FF7CFCBA77602A48" box="[151,278,830,852]" italics="true" pageId="3" pageNumber="66">Machaerites</emphasis>
</taxonomicName>
(
<taxonomicName id="6BD1B1938C3BB659FECFFCBA77EF2A48" box="[292,409,830,852]" class="Insecta" family="Staphylinidae" genus="Bythoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3BB659FECFFCBA77EF2A48" box="[292,409,830,852]" italics="true" pageId="3" pageNumber="66">Bythoxenus</emphasis>
</taxonomicName>
s. Raffray)
<taxonomicName id="6BD1B1938C3BB659FDFAFCBA754D2A48" authority="Peyerimhoff 1919: 72" authorityName="Peyerimhoff" authorityPageNumber="72" authorityYear="1919" box="[529,827,830,852]" class="Insecta" family="Staphylinidae" genus="Bordei" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3BB659FDFAFCBA74232A48" box="[529,597,830,852]" italics="true" pageId="3" pageNumber="66">Bordei</emphasis>
Peyerimhoff 1919: 72
</taxonomicName>
, fig. 1 (habitus);
<taxonomicName id="6BD1B1938C3BB659FF7CFCD9746F2A6F" authority="Bordei Raffray, 1924: 85" authorityName="Bordei Raffray" authorityPageNumber="85" authorityYear="1924" box="[151,537,861,883]" class="Insecta" family="Staphylinidae" genus="Machaerites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3BB659FF7CFCD977162A6F" box="[151,352,861,883]" italics="true" pageId="3" pageNumber="66">Machaerites Bordei</emphasis>
Raffray, 1924: 85
</taxonomicName>
;
</paragraph>
<paragraph id="AC6ECA108C3BB659FF7CFCF874E12AAC" blockId="3.[151,994,830,944]" pageId="3" pageNumber="66">
<taxonomicName id="6BD1B1938C3BB659FF7CFCF874682A8E" authority="Bordei Jeannel 1956: 61" authorityName="Bordei Jeannel" authorityPageNumber="61" authorityYear="1956" box="[151,542,892,914]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3BB659FF7CFCF8771F2A8E" box="[151,361,892,914]" italics="true" pageId="3" pageNumber="66">Anopsibythus Bordei</emphasis>
Jeannel 1956: 61
</taxonomicName>
; 64, fig. 53 (habitus);
<taxonomicName id="6BD1B1938C3BB659FF7CFC1E74E42AAC" authority="Lobl &amp; Besuchet 2004: 314" authorityName="Lobl &amp; Besuchet" authorityPageNumber="314" authorityYear="2004" box="[151,658,922,944]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="bordei">
<emphasis id="9EA516028C3BB659FF7CFC1E77062AAC" box="[151,368,922,944]" italics="true" pageId="3" pageNumber="66">Tychobythinus bordei</emphasis>
Löbl &amp; Besuchet 2004: 314
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3BB659FF7CFC59776A2EDC" pageId="3" pageNumber="66" type="materials_examined">
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<typeStatus id="736A74B28C3BB659FF7CFC5976A42AEA" box="[151,210,989,1014]" pageId="3" pageNumber="66">Type</typeStatus>
material
</emphasis>
.
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<emphasis id="9EA516028C3BB659FEBCFC5977BB2AEA" bold="true" box="[343,461,989,1014]" pageId="3" pageNumber="66">Lectotype</emphasis>
</typeStatus>
, female, present designation.
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NORTHEASTERN
<collectingCountry id="D4C68A808C3BB659FBC0FC5972DB2AEA" box="[1067,1197,989,1014]" name="Algeria" pageId="3" pageNumber="66">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C3BB659FB2AFC5973E22AEA" bold="true" box="[1217,1428,989,1014]" pageId="3" pageNumber="66">Annaba province</emphasis>
: Bône (old name of Annaba),
<date id="D86FECD08C3BB659FE36FB8474242D04" box="[477,594,1024,1048]" pageId="3" pageNumber="66" value="1918-02-19">19.II.1918</date>
(R. de Bordé) (ex coll. Peyerimhoff, MNHN).
</paragraph>
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<emphasis id="9EA516028C3BB659FF2CFBA6771B2D27" bold="true" box="[199,365,1058,1083]" pageId="3" pageNumber="66">Redescription</emphasis>
. Body length
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, pubescence not very dense, consists of long and flattened setae (length:
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) on head, pronotum, elytra and abdomen, other suberected setae shorter (length:
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) on antennae and legs. Tegument smooth and glossy.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CFB0E754C2F3C" blockId="3.[151,1436,989,2019]" pageId="3" pageNumber="66">
Head (
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) longer (
<quantity id="6B2967F58C3BB659FE50FB0E74462DBF" box="[443,560,1162,1187]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="66" unit="mm" value="0.25">0.250 mm</quantity>
) than wide (
<quantity id="6B2967F58C3BB659FD50FB0E75452DBF" box="[699,819,1162,1187]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" pageId="3" pageNumber="66" unit="mm" value="0.225">0.225 mm</quantity>
), frontal lobe width
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its dorsal surface glossy with some punctures and well impressed median sulcus. Very evident clypeal carina reaches ocular region, very marked and well visible median occipital carina reaching to the posterior edge of median sulcus of frontal lobe, tempora rounded. Gular region convex entirely traversed by evident median longitudinal carina, its surface glossy. Segment II of maxillary palpi with 28–30 tubercles, segment III distinctly longer than wide with 4–6 tubercles, last segment of maxillary palpi (
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) about 3.5 times longer (0.300 mm) than wide (
<quantity id="6B2967F58C3BB659FBD4FAB372C32C4C" box="[1087,1205,1335,1360]" metricMagnitude="-5" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="66" unit="mm" value="0.085">0.085 mm</quantity>
) with lateral margin slightly concave. Antenna (
<figureCitation id="34EAD6958C3BB659FE23FADE747D2C6F" box="[456,523,1370,1395]" captionStart="FIGURES 4 – 6" captionStartId="4.[151,264,1807,1829]" captionTargetBox="[397,1208,182,1757]" captionTargetId="figure@4.[355,1232,167,1774]" captionTargetPageId="4" captionText="FIGURES 4 – 6. Tychobythinus bordei. Scale: 0, 1 mm. 4 — Head, schematic drawing. 5 — Left maxillary palpus, lectotype. 6 — Right antenna, lectotype." httpUri="https://zenodo.org/record/284496/files/figure.png" pageId="3" pageNumber="66">Fig. 6</figureCitation>
)
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long, scape about 4.5 times longer (
<quantity id="6B2967F58C3BB659FBF0FADF72E62C6F" box="[1051,1168,1371,1395]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="66" unit="mm" value="0.21">0.210 mm</quantity>
) than wide (
<quantity id="6B2967F58C3BB659FAF2FADF73E62C6F" box="[1305,1424,1371,1395]" metricMagnitude="-5" metricUnit="m" metricValue="4.699999999999999" pageId="3" pageNumber="66" unit="mm" value="0.047">0.047 mm</quantity>
), antennomere II, slightly asymmetric, longer (
<quantity id="6B2967F58C3BB659FD77FAF975652C8A" box="[668,787,1405,1430]" metricMagnitude="-5" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="66" unit="mm" value="0.062">0.062 mm</quantity>
) than wide (
<quantity id="6B2967F58C3BB659FC4EFAFA72692C8A" box="[933,1055,1406,1430]" metricMagnitude="-5" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="66" unit="mm" value="0.037">0.037 mm</quantity>
), antennomeres III–VII distinctly longer than wide, III slightly longer than other funicular antennomeres, IV and V subequal in length, VI shorter than previous, VII slightly longer than VI, VIII slightly longer than wide and shorter than previous funicular antennomeres. Antennomere IX as long as wide, X slightly wider than long, XI more than 2 times longer than wide, and 2 times longer than length of IX and X combined.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CF9AF75732F7A" blockId="3.[151,1436,989,2019]" pageId="3" pageNumber="66">
Pronotum slightly wider (
<quantity id="6B2967F58C3BB659FE0EF9AE742D2F5F" box="[485,603,1578,1603]" metricMagnitude="-4" metricUnit="m" metricValue="3.35" pageId="3" pageNumber="66" unit="mm" value="0.335">0.335 mm</quantity>
) than long (
<quantity id="6B2967F58C3BB659FD0BF9AE75212F5F" box="[736,855,1578,1603]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="66" unit="mm" value="0.325">0.325 mm</quantity>
), widest at forequarter, pleural carina barely visible, very deep antebasal sulcus with strongly marked edges.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CF9EB74E92F9B" blockId="3.[151,1436,989,2019]" box="[199,671,1647,1672]" pageId="3" pageNumber="66">
Elytra width
<quantity id="6B2967F58C3BB659FEB2F9EB77BB2F9B" box="[345,461,1647,1672]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="66" unit="mm" value="0.55">0.550 mm</quantity>
, length 0.500 mm.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CF91774BE2FB7" blockId="3.[151,1436,989,2019]" box="[199,712,1682,1707]" pageId="3" pageNumber="66">Metasternum with surface smooth and glossy.</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CF93272E62FEC" blockId="3.[151,1436,989,2019]" pageId="3" pageNumber="66">
Legs elongated, protrochanters with 3–4 small tubercles; profemora with 14–16 small tubercles, base of mesofemora with 4–5 small protuberances on its anterior edge. Metatibiae
<quantity id="6B2967F58C3BB659FC35F95C72262FF3" box="[990,1104,1752,1776]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="66" unit="mm" value="0.49">0.490 mm</quantity>
long.
</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CF97E77F32E0F" blockId="3.[151,1436,989,2019]" box="[199,389,1786,1811]" pageId="3" pageNumber="66">Male. Unknown.</paragraph>
<paragraph id="AC6ECA108C3BB659FF2CF899776A2EDC" blockId="3.[151,1436,989,2019]" pageId="3" pageNumber="66">
<emphasis id="9EA516028C3BB659FF2CF89977DE2E2A" bold="true" box="[199,424,1821,1846]" pageId="3" pageNumber="66">Comparative notes</emphasis>
.
<taxonomicName id="6BD1B1938C3BB659FE5DF89A746E2E2A" box="[438,536,1821,1846]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="bordei">
<emphasis id="9EA516028C3BB659FE5DF89A746E2E2A" box="[438,536,1821,1846]" italics="true" pageId="3" pageNumber="66">T. bordei</emphasis>
</taxonomicName>
is distinguished from other species of the
<taxonomicName id="6BD1B1938C3BB659FC1BF89972472E2A" box="[1008,1073,1821,1846]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3BB659FC1BF89972472E2A" box="[1008,1073,1821,1846]" italics="true" pageId="3" pageNumber="66">theryi</emphasis>
</taxonomicName>
group by the very long occipital median carina reaching the posterior edge of median sulcus of frontal lobe (at most reaching the anterior edge of vertexal foveae in other species). It is similar to
<taxonomicName id="6BD1B1938C3BB659FD5BF8E0754E2E67" box="[688,824,1890,1915]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3BB659FD5BF8E0754E2E67" box="[688,824,1890,1915]" italics="true" pageId="3" pageNumber="66">T. normandi</emphasis>
</taxonomicName>
in having the anterior margin of mesofemora with 4–5 small tubercles, while its lateral margin of the last article of the maxillary palpi is slightly concave and similar to
<taxonomicName id="6BD1B1938C3BB659FA6CF80277632EDD" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="9EA516028C3BB659FA6CF80277632EDD" italics="true" pageId="3" pageNumber="66">T. longipalpis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3BB659FF2CF84E73152EFF" box="[199,1379,1994,2019]" pageId="3" pageNumber="66" type="discussion">
<paragraph id="AC6ECA108C3BB659FF2CF84E73152EFF" blockId="3.[151,1436,989,2019]" box="[199,1379,1994,2019]" pageId="3" pageNumber="66">
<emphasis id="9EA516028C3BB659FF2CF84E77422EFF" bold="true" box="[199,308,1994,2019]" pageId="3" pageNumber="66">Remarks</emphasis>
. In the original description Peyerimhoff (1919: 72) mentioned two females, one of them is lost.
</paragraph>
</subSubSection>
</treatment>
</document>

200
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<heading id="F7267D7C8C3DB65FFF7CFCA974C72A5B" bold="true" box="[151,689,813,839]" fontSize="11" level="1" pageId="5" pageNumber="68" reason="1">
<taxonomicName id="6BD1B1938C3DB65FFF7CFCA974C72A5B" ID-CoL="7CYDC" authority="Normand, 1930" authorityName="Normand" authorityYear="1930" box="[151,689,813,839]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="longipalpis">
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(Normand, 1930)
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<paragraph id="AC6ECA108C3DB65FFF7CFCCB77672A74" blockId="5.[151,689,813,872]" box="[151,273,847,872]" pageId="5" pageNumber="68">
(
<figureCitation id="34EAD6958C3DB65FFF74FCCB777C2A74" box="[159,266,847,872]" captionStart="FIGURES 7 – 9" captionStartId="6.[151,264,1710,1732]" captionTargetBox="[379,1211,284,1671]" captionTargetId="figure@6.[348,1239,267,1688]" captionTargetPageId="6" captionText="FIGURES 7 – 9. Tychobythinus longipalpis. Scale: 0, 1 mm. 7 — Head, schematic drawing. 8 — Left maxillary palpus, holotype. 9 — Right antenna, holotype." httpUri="https://zenodo.org/record/284497/files/figure.png" pageId="5" pageNumber="68">Figs. 7–9</figureCitation>
)
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF7CFC1C742A2AB2" blockId="5.[151,805,920,1003]" box="[151,604,920,942]" pageId="5" pageNumber="68">
<taxonomicName id="6BD1B1938C3DB65FFF7CFC1C742E2AB2" authority="Normand 1930: 165" authorityName="Normand" authorityPageNumber="165" authorityYear="1930" box="[151,600,920,942]" class="Insecta" family="Staphylinidae" genus="Bythoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="9EA516028C3DB65FFF7CFC1C77F22AB2" box="[151,388,920,942]" italics="true" pageId="5" pageNumber="68">Bythoxenus longipalpis</emphasis>
Normand 1930: 165
</taxonomicName>
;
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3DB65FFF7CFC3274B12AF7" pageId="5" pageNumber="68" type="description">
<paragraph id="AC6ECA108C3DB65FFF7CFC3274B12AF7" blockId="5.[151,805,920,1003]" pageId="5" pageNumber="68">
<taxonomicName id="6BD1B1938C3DB65FFF7CFC32743D2AD0" authority="Jeannel 1956: 61" authorityName="Jeannel" authorityPageNumber="61" authorityYear="1956" box="[151,587,950,972]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="9EA516028C3DB65FFF7CFC3277E12AD0" box="[151,407,950,972]" italics="true" pageId="5" pageNumber="68">Anopsibythus longipalpis</emphasis>
Jeannel 1956: 61
</taxonomicName>
, 63, fig. 51 (habitus);
<taxonomicName id="6BD1B1938C3DB65FFF7CFC5174B52AF7" authority="Lobl &amp; Besuchet 2004: 315" authorityName="Lobl &amp; Besuchet" authorityPageNumber="315" authorityYear="2004" box="[151,707,981,1003]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="9EA516028C3DB65FFF7CFC5177D72AF7" box="[151,417,981,1003]" italics="true" pageId="5" pageNumber="68">Tychobythinus longipalpis</emphasis>
Löbl &amp; Besuchet 2004: 315
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3DB65CFF7CFB93741029C8" lastPageId="6" lastPageNumber="69" pageId="5" pageNumber="68" type="materials_examined">
<paragraph id="AC6ECA108C3DB65FFF7CFB93771D2D48" blockId="5.[151,1436,1047,2009]" pageId="5" pageNumber="68">
<emphasis id="9EA516028C3DB65FFF7CFB9377312D2C" bold="true" box="[151,327,1047,1072]" pageId="5" pageNumber="68">
<typeStatus id="736A74B28C3DB65FFF7CFB9376A42D2C" box="[151,210,1047,1072]" pageId="5" pageNumber="68">Type</typeStatus>
material
</emphasis>
.
<emphasis id="9EA516028C3DB65FFEB5FB9377BD2D2C" bold="true" box="[350,459,1047,1072]" pageId="5" pageNumber="68">
<typeStatus id="736A74B28C3DB65FFEB5FB9377BD2D2C" box="[350,459,1047,1072]" pageId="5" pageNumber="68" type="holotype">Holotype</typeStatus>
</emphasis>
, female.
<emphasis id="9EA516028C3DB65FFDAEFB9375B52D2C" bold="true" box="[581,963,1047,1072]" pageId="5" pageNumber="68">
NORTHEASTERN
<collectingCountry id="D4C68A808C3DB65FFCAAFB9375B52D2C" box="[833,963,1047,1072]" name="Algeria" pageId="5" pageNumber="68">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C3DB65FFC30FB9372D12D2C" bold="true" box="[987,1191,1047,1072]" pageId="5" pageNumber="68">Skikda province</emphasis>
: Collo,
<date id="D86FECD08C3DB65FFAFFFB9C73112D2C" box="[1300,1383,1048,1072]" pageId="5" pageNumber="68" value="1929-10">X.1929</date>
(H. Normand) (INAT).
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CFBDB75282DDC" blockId="5.[151,1436,1047,2009]" pageId="5" pageNumber="68">
<emphasis id="9EA516028C3DB65FFF2CFBDB771B2D64" bold="true" box="[199,365,1119,1144]" pageId="5" pageNumber="68">Redescription</emphasis>
. Body length
<quantity id="6B2967F58C3DB65FFDF7FBE474FE2D6B" box="[540,648,1120,1144]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="68" unit="mm" value="1.2">1.20 mm</quantity>
, pubescence not very dense consisting of long and flattened setae (length:
<quantity id="6B2967F58C3DB65FFF1DFB0077C12D80" box="[246,439,1156,1180]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7" pageId="5" pageNumber="68" unit="mm" value="0.075" valueMax="0.08" valueMin="0.07">0.070–0.080 mm</quantity>
) on head, pronotum, elytra and abdomen, other suberected setae shorter (length:
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) on antennae and legs. Tegument smooth and glossy.
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CFB4872722F50" blockId="5.[151,1436,1047,2009]" pageId="5" pageNumber="68">
Head (
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) slightly wider (0.210) than long (0.200 mm) frontal lobe width
<quantity id="6B2967F58C3DB65FFBB3FB4872BB2DF8" box="[1112,1229,1228,1252]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="68" unit="mm" value="0.11">0.110 mm</quantity>
its dorsal surface glossy with some punctures concentrated on sides and well impressed median sulcus. Clypeal carina reaches ocular region, occipital region strongly convex with short median occipital carina reaching its middle, tempora rounded and slightly prominent downward. Gular region slightly convex, separated from the anterior portion of the head by a deep transverse groove, its surface glossy. Segment II of maxillary palpi elongated with 20 strong tubercles, segment III longer than wide with 6 tubercles, last segment of maxillary palpi (
<figureCitation id="34EAD6958C3DB65FFBF9FAFB72232C84" box="[1042,1109,1407,1432]" captionStart="FIGURES 7 – 9" captionStartId="6.[151,264,1710,1732]" captionTargetBox="[379,1211,284,1671]" captionTargetId="figure@6.[348,1239,267,1688]" captionTargetPageId="6" captionText="FIGURES 7 – 9. Tychobythinus longipalpis. Scale: 0, 1 mm. 7 — Head, schematic drawing. 8 — Left maxillary palpus, holotype. 9 — Right antenna, holotype." httpUri="https://zenodo.org/record/284497/files/figure.png" pageId="5" pageNumber="68">Fig. 8</figureCitation>
) about 3 times longer (
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) than wide (
<quantity id="6B2967F58C3DB65FFEB8FA2077BA2CA0" box="[339,460,1444,1468]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="68" unit="mm" value="0.08">0.080 mm</quantity>
), its lateral margin slightly concave. Antenna (
<figureCitation id="34EAD6958C3DB65FFC35FA2072552CA0" box="[990,1059,1444,1469]" captionStart="FIGURES 7 – 9" captionStartId="6.[151,264,1710,1732]" captionTargetBox="[379,1211,284,1671]" captionTargetId="figure@6.[348,1239,267,1688]" captionTargetPageId="6" captionText="FIGURES 7 – 9. Tychobythinus longipalpis. Scale: 0, 1 mm. 7 — Head, schematic drawing. 8 — Left maxillary palpus, holotype. 9 — Right antenna, holotype." httpUri="https://zenodo.org/record/284497/files/figure.png" pageId="5" pageNumber="68">Fig. 9</figureCitation>
)
<quantity id="6B2967F58C3DB65FFBDFFA2072D12CA0" box="[1076,1191,1444,1469]" metricMagnitude="-4" metricUnit="m" metricValue="6.85" pageId="5" pageNumber="68" unit="mm" value="0.685">0.685 mm</quantity>
long very thin, scape about 5 times longer (
<quantity id="6B2967F58C3DB65FFE64FA4C74732CFC" box="[399,517,1480,1504]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="68" unit="mm" value="0.21">0.210 mm</quantity>
) than wide (
<quantity id="6B2967F58C3DB65FFD78FA4C757D2CFC" box="[659,779,1480,1504]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="68" unit="mm" value="0.04">0.040 mm</quantity>
), antennomere II, slightly asymmetric, longer (
<quantity id="6B2967F58C3DB65FFAF5FA4373E22CFC" box="[1310,1428,1479,1504]" metricMagnitude="-5" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="68" unit="mm" value="0.055">0.055 mm</quantity>
) than wide (
<quantity id="6B2967F58C3DB65FFEF8FA6877FD2F18" box="[275,395,1516,1540]" metricMagnitude="-5" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="68" unit="mm" value="0.037">0.037 mm</quantity>
), antennomeres III–V distinctly longer than wide and subequal in size, VI shorter than V and longer than wide, VII and VIII as long than as wide. Antennomeres IX–X wider than long, XI more than 2 times longer than wide, and about 2 times longer than combined length of IX and X.
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CF9DC75AD2F89" blockId="5.[151,1436,1047,2009]" pageId="5" pageNumber="68">
Pronotum distinctly wider (0.300 mm) than long (
<quantity id="6B2967F58C3DB65FFD16F9D375002F6C" box="[765,886,1623,1648]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="68" unit="mm" value="0.26">0.260 mm</quantity>
), widest a little before the middle, pleural carina barely visibile, deep antebasal sulcus with its posterior edge very detected.
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CF91B75352FA4" blockId="5.[151,1436,1047,2009]" box="[199,835,1695,1720]" pageId="5" pageNumber="68">
Elytra strongly wider (
<quantity id="6B2967F58C3DB65FFE28F91B744F2FA4" box="[451,569,1695,1720]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" pageId="5" pageNumber="68" unit="mm" value="0.525">0.525 mm</quantity>
) than long (
<quantity id="6B2967F58C3DB65FFD54F92475412FA4" box="[703,823,1696,1720]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="68" unit="mm" value="0.44">0.440 mm</quantity>
).
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CF94174BE2FC1" blockId="5.[151,1436,1047,2009]" box="[199,712,1732,1757]" pageId="5" pageNumber="68">Metasternum with surface smooth and glossy.</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CF96D755C2E39" blockId="5.[151,1436,1047,2009]" pageId="5" pageNumber="68">
Legs elongated, protrochanters with 4 tubercles; profemora with 8 small tubercles. All tibiae slightly flattened and enlarged in the distal third. Metatibiae
<quantity id="6B2967F58C3DB65FFD93F888749C2E38" box="[632,746,1804,1828]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="5" pageNumber="68" unit="mm" value="0.47">0.470 mm</quantity>
long.
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CF8AB77F32E54" blockId="5.[151,1436,1047,2009]" box="[199,389,1839,1864]" pageId="5" pageNumber="68">Male. Unknown.</paragraph>
<paragraph id="AC6ECA108C3DB65CFF2CF8D0741029C8" blockId="5.[151,1436,1047,2009]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="69" pageId="5" pageNumber="68">
<emphasis id="9EA516028C3DB65FFF2CF8D077DF2E71" bold="true" box="[199,425,1876,1901]" pageId="5" pageNumber="68">Comparative notes</emphasis>
. This species is similar to
<taxonomicName id="6BD1B1938C3DB65FFD3AF8D1750F2E71" box="[721,889,1876,1901]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C3DB65FFD3AF8D1750F2E71" box="[721,889,1876,1901]" italics="true" pageId="5" pageNumber="68">T. peyerimhoffi</emphasis>
</taxonomicName>
from which it is distinguished by the head wider than long (longer than wide in
<taxonomicName id="6BD1B1938C3DB65FFE1CF8FD74D72E8C" box="[503,673,1911,1936]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C3DB65FFE1CF8FD74D72E8C" box="[503,673,1911,1936]" italics="true" pageId="5" pageNumber="68">T. peyerimhoffi</emphasis>
</taxonomicName>
), by the median occipital carina short and reaching approximately half of the occipital length (reaching the posterior edge of vertexal foveae in
<taxonomicName id="6BD1B1938C3DB65FFC1FF81972EA2EA9" box="[1012,1180,1948,1973]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C3DB65FFC1FF81972EA2EA9" box="[1012,1180,1948,1973]" italics="true" pageId="5" pageNumber="68">T. peyerimhoffi</emphasis>
</taxonomicName>
), by the lateral margin of last segment of maxillary palpi slightly concave (straight in
<taxonomicName id="6BD1B1938C3DB65FFC9FF84572692EC5" box="[884,1055,1984,2009]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C3DB65FFC9FF84572692EC5" box="[884,1055,1984,2009]" italics="true" pageId="5" pageNumber="68">T. peyerimhoffi</emphasis>
</taxonomicName>
), by the antennal scape 5 times longer than wide (about 4 times longer than wide in
<taxonomicName id="6BD1B1938C3EB65CFD07FF1D75E029AC" box="[748,918,151,176]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C3EB65CFD07FF1D75E029AC" box="[748,918,151,176]" italics="true" pageId="6" pageNumber="69">T. peyerimhoffi</emphasis>
</taxonomicName>
), and by the absence of median carina on the metasternum (present in
<taxonomicName id="6BD1B1938C3EB65CFE47FF39742329C9" box="[428,597,188,213]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="9EA516028C3EB65CFE47FF39742329C9" box="[428,597,188,213]" italics="true" pageId="6" pageNumber="69">T. peyerimhoffi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

170
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<mods:title id="644CEF9F6135469AE776180442D97B86">A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<subSubSection id="E4CB999B8C3DB65FFF7CFF1C72DC29E8" pageId="5" pageNumber="68" type="nomenclature">
<paragraph id="AC6ECA108C3DB65FFF7CFF1C74D729AE" blockId="5.[151,673,152,178]" box="[151,673,152,178]" pageId="5" pageNumber="68">
<heading id="F7267D7C8C3DB65FFF7CFF1C74D729AE" bold="true" box="[151,673,152,178]" fontSize="11" level="1" pageId="5" pageNumber="68" reason="1">
<taxonomicName id="6BD1B1938C3DB65FFF7CFF1C74D729AE" ID-CoL="59SD4" authority="Normand, 1930" authorityName="Normand" authorityYear="1930" box="[151,673,152,178]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="araneipes">
<emphasis id="9EA516028C3DB65FFF7CFF1C74D729AE" bold="true" box="[151,673,152,178]" pageId="5" pageNumber="68">
<emphasis id="9EA516028C3DB65FFF7CFF1C77B329AE" bold="true" box="[151,453,152,178]" italics="true" pageId="5" pageNumber="68">Tychobythinus araneipes</emphasis>
(Normand, 1930)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF7CFF5A72DC29E8" blockId="5.[151,1435,222,336]" box="[151,1194,222,245]" pageId="5" pageNumber="68">
<taxonomicName id="6BD1B1938C3DB65FFF7CFF5A748129E8" authority="Normand 1930: 163" authorityName="Normand" authorityPageNumber="163" authorityYear="1930" box="[151,759,222,245]" class="Insecta" family="Staphylinidae" genus="Bythoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="araneipes" subGenus="Acanthobythus">
<emphasis id="9EA516028C3DB65FFF7CFF5A777A29E8" box="[151,268,222,244]" italics="true" pageId="5" pageNumber="68">Bythoxenus</emphasis>
(
<emphasis id="9EA516028C3DB65FFEF1FF5A77C629E8" box="[282,432,222,244]" italics="true" pageId="5" pageNumber="68">Acanthobythus</emphasis>
)
<emphasis id="9EA516028C3DB65FFE54FF64745529E9" box="[447,547,224,245]" italics="true" pageId="5" pageNumber="68">araneipes</emphasis>
Normand 1930: 163
</taxonomicName>
, figs 1 (habitus) and 2 (head, lateral view);
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3DB65FFF7CFF7972E9284C" pageId="5" pageNumber="68" type="description">
<paragraph id="AC6ECA108C3DB65FFF7CFF797278280F" blockId="5.[151,1435,222,336]" box="[151,1038,253,275]" pageId="5" pageNumber="68">
<taxonomicName id="6BD1B1938C3DB65FFF7CFF79743D280F" authority="Jeannel 1956: 65" authorityName="Jeannel" authorityPageNumber="65" authorityYear="1956" box="[151,587,253,275]" class="Insecta" family="Staphylinidae" genus="Acanthobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="araneipes">
<emphasis id="9EA516028C3DB65FFF7CFF7977E1280F" box="[151,407,253,275]" italics="true" pageId="5" pageNumber="68">Acanthobythus araneipes</emphasis>
Jeannel 1956: 65
</taxonomicName>
, figs 59 (habitus) and 60 (maxillary palpus);
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF7CFE9872E9284C" blockId="5.[151,1435,222,336]" pageId="5" pageNumber="68">
<taxonomicName id="6BD1B1938C3DB65FFF7CFE9874BE282E" authority="Newton &amp; Chandler 1989: 48" authorityName="Newton &amp; Chandler" authorityPageNumber="48" authorityYear="1989" box="[151,712,284,306]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="araneipes">
<emphasis id="9EA516028C3DB65FFF7CFE9877E5282E" box="[151,403,284,306]" italics="true" pageId="5" pageNumber="68">Tychobythinus araneipes</emphasis>
Newton &amp; Chandler 1989: 48
</taxonomicName>
; Löbl &amp; Besuchet 2004: 314; Besuchet, Cuccodoro, Sabella 2013: 272, figs 1 (habitus), 2 (head and palpi), 3 (base of antenna), 4 (aedeagus) and 8 (head and pronotum).
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3DB65FFF7CFEF9749F2BA9" pageId="5" pageNumber="68" type="materials_examined">
<paragraph id="AC6ECA108C3DB65FFF7CFEF9736F28A5" blockId="5.[151,1436,381,730]" pageId="5" pageNumber="68">
<emphasis id="9EA516028C3DB65FFF7CFEF97736288A" bold="true" box="[151,320,381,406]" pageId="5" pageNumber="68">
<typeStatus id="736A74B28C3DB65FFF7CFEF976A4288A" box="[151,210,381,406]" pageId="5" pageNumber="68">Type</typeStatus>
material
</emphasis>
.
<emphasis id="9EA516028C3DB65FFEBBFEF977B0288A" bold="true" box="[336,454,381,406]" pageId="5" pageNumber="68">
<typeStatus id="736A74B28C3DB65FFEBBFEF977B0288A" box="[336,454,381,406]" pageId="5" pageNumber="68" type="lectotype">Lectotype</typeStatus>
</emphasis>
, male.
<emphasis id="9EA516028C3DB65FFDF5FEF975E3288A" bold="true" box="[542,917,381,406]" pageId="5" pageNumber="68">
NORTHEASTERN
<collectingCountry id="D4C68A808C3DB65FFCF9FEF975E3288A" box="[786,917,381,406]" name="Algeria" pageId="5" pageNumber="68">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C3DB65FFC4EFEF9721D288A" bold="true" box="[933,1131,381,406]" pageId="5" pageNumber="68">Skikda province</emphasis>
: male, Philippeville, route de Collo,
<date id="D86FECD08C3DB65FFEEFFE25772E28A5" box="[260,344,417,441]" pageId="5" pageNumber="68" value="1929-10">X.1929</date>
(H. Normand) (MHNG).
<emphasis id="9EA516028C3DB65FFD91FE25755C28A6" bold="true" box="[634,810,417,442]" pageId="5" pageNumber="68">
<typeStatus id="736A74B28C3DB65FFD91FE25755C28A6" box="[634,810,417,442]" pageId="5" pageNumber="68" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
,
<specimenCount id="BAD701998C3DB65FFCD3FE2575EA28A6" box="[824,924,417,442]" pageId="5" pageNumber="68" type="female">2 female</specimenCount>
, same data as
<typeStatus id="736A74B28C3DB65FFBD0FE2572D428A6" box="[1083,1186,417,442]" pageId="5" pageNumber="68" type="lectotype">lectotype</typeStatus>
(MHNG).
</paragraph>
<paragraph id="AC6ECA108C3DB65FFF2CFE41749F2BA9" blockId="5.[151,1436,381,730]" pageId="5" pageNumber="68">
<emphasis id="9EA516028C3DB65FFF2CFE4177DC28C2" bold="true" box="[199,426,453,478]" pageId="5" pageNumber="68">Comparative notes</emphasis>
.
<taxonomicName id="6BD1B1938C3DB65FFE52FE4174A628C1" box="[441,720,453,478]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="araneipes">
<emphasis id="9EA516028C3DB65FFE52FE4174A628C1" box="[441,720,453,478]" italics="true" pageId="5" pageNumber="68">Tychobythinus araneipes</emphasis>
</taxonomicName>
is the only member of the genus to have conspicuous temporal spines. This species is similar to
<taxonomicName id="6BD1B1938C3DB65FFDF0FE6E74CE2B1E" box="[539,696,489,514]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="9EA516028C3DB65FFDF0FE6E74CE2B1E" box="[539,696,489,514]" italics="true" pageId="5" pageNumber="68">T. longipalpis</emphasis>
</taxonomicName>
in the short occipital median carina reaching the middle of the occipital region (occipital median carina at least reaching the anterior edge of vertexal foveae in other species) and in antennal scape about 5 times longer than wide (at most 4 times longer than wide in other species), it has the aedeagus similar to
<taxonomicName id="6BD1B1938C3DB65FFE9DFDD277892B72" box="[374,511,597,622]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3DB65FFE9DFDD277892B72" box="[374,511,597,622]" italics="true" pageId="5" pageNumber="68">T. normandi</emphasis>
</taxonomicName>
and
<taxonomicName id="6BD1B1938C3DB65FFDDDFDD274BA2B71" box="[566,716,598,621]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="monoceros">
<emphasis id="9EA516028C3DB65FFDDDFDD274BA2B71" box="[566,716,598,621]" italics="true" pageId="5" pageNumber="68">T. monoceros</emphasis>
</taxonomicName>
. Also for the deep and broad depression of gular region of male and the clypeal carina reaching the temporal region (reaching the ocular region in all other species of the
<taxonomicName id="6BD1B1938C3DB65FFAB0FDFD73EA2B8E" box="[1371,1436,633,658]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3DB65FFAB0FDFD73EA2B8E" box="[1371,1436,633,658]" italics="true" pageId="5" pageNumber="68">theryi</emphasis>
</taxonomicName>
group)
<taxonomicName id="6BD1B1938C3DB65FFF03FD1A77022BA9" box="[232,372,670,693]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="araneipes">
<emphasis id="9EA516028C3DB65FFF03FD1A77022BA9" box="[232,372,670,693]" italics="true" pageId="5" pageNumber="68">T. araneipes</emphasis>
</taxonomicName>
is comparable with
<taxonomicName id="6BD1B1938C3DB65FFDBCFD1A74942BAA" box="[599,738,669,694]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3DB65FFDBCFD1A74942BAA" box="[599,738,669,694]" italics="true" pageId="5" pageNumber="68">T. normandi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3DB65FFF2CFD45720A2BC6" box="[199,1148,705,730]" pageId="5" pageNumber="68" type="discussion">
<paragraph id="AC6ECA108C3DB65FFF2CFD45720A2BC6" blockId="5.[151,1436,381,730]" box="[199,1148,705,730]" pageId="5" pageNumber="68">
<emphasis id="9EA516028C3DB65FFF2CFD4577422BC6" bold="true" box="[199,308,705,730]" pageId="5" pageNumber="68">Remarks</emphasis>
. Species recently attributed to genus
<taxonomicName id="6BD1B1938C3DB65FFD3CFD45720E2BC6" authority="Besuchet et al. 2013" authorityName="Besuchet et al." authorityYear="2013" box="[727,1144,705,730]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3DB65FFD3CFD45750E2BC6" box="[727,888,705,730]" italics="true" pageId="5" pageNumber="68">Tychobythinus</emphasis>
(Besuchet et al. 2013)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

137
data/24/78/7B/24787B068C3EB65DFF7CF8AA720A2878.xml Normal file
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<mods:title id="B7C22646AAA50F375BDD39CFB7E81F9D">A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<subSubSection id="E4CB999B8C3EB65CFF7CF8AA774F2EFA" pageId="6" pageNumber="69" type="nomenclature">
<paragraph id="AC6ECA108C3EB65CFF7CF8AA74C62E54" blockId="6.[151,688,1838,1864]" box="[151,688,1838,1864]" pageId="6" pageNumber="69">
<heading id="F7267D7C8C3EB65CFF7CF8AA74C62E54" bold="true" box="[151,688,1838,1864]" fontSize="11" level="1" pageId="6" pageNumber="69" reason="1">
<taxonomicName id="6BD1B1938C3EB65CFF7CF8AA74C62E54" ID-CoL="7CYDJ" authority="Normand, 1932" authorityName="Normand" authorityYear="1932" box="[151,688,1838,1864]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="monoceros">
<emphasis id="9EA516028C3EB65CFF7CF8AA74C62E54" bold="true" box="[151,688,1838,1864]" pageId="6" pageNumber="69">
<emphasis id="9EA516028C3EB65CFF7CF8AA77A22E54" bold="true" box="[151,468,1838,1864]" italics="true" pageId="6" pageNumber="69">Tychobythinus monoceros</emphasis>
(Normand, 1932)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AC6ECA108C3EB65CFF7CF8F073EA2ED4" blockId="6.[151,1436,1908,2022]" pageId="6" pageNumber="69">
<taxonomicName id="6BD1B1938C3EB65CFF7CF8F074912E96" authority="Normand 1932: 16" authorityName="Normand" authorityPageNumber="16" authorityYear="1932" box="[151,743,1908,1931]" class="Insecta" family="Staphylinidae" genus="Bythoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="monoceros" subGenus="Ceratobythus">
<emphasis id="9EA516028C3EB65CFF7CF8F0777A2E96" box="[151,268,1908,1930]" italics="true" pageId="6" pageNumber="69">Bythoxenus</emphasis>
(
<emphasis id="9EA516028C3EB65CFEF1F8F077D42E96" box="[282,418,1908,1930]" italics="true" pageId="6" pageNumber="69">Ceratobythus</emphasis>
)
<emphasis id="9EA516028C3EB65CFE5AF8F274692E97" box="[433,543,1910,1931]" italics="true" pageId="6" pageNumber="69">monoceros</emphasis>
Normand 1932: 16
</taxonomicName>
, figs 2–3 (head dorsal and lateral view);
<taxonomicName id="6BD1B1938C3EB65CFF7CF817743E2EB5" authority="Jeannel 1956: 64" authorityName="Jeannel" authorityPageNumber="64" authorityYear="1956" box="[151,584,1939,1961]" class="Insecta" family="Staphylinidae" genus="Ceratobythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="monoceros">
<emphasis id="9EA516028C3EB65CFF7CF81777E52EB5" box="[151,403,1939,1961]" italics="true" pageId="6" pageNumber="69">Ceratobythus monoceros</emphasis>
Jeannel 1956: 64
</taxonomicName>
, fig. 58 (habitus), Newton &amp; Chandler 1989: 48; Löbl &amp; Besuchet 2004: 312;
<taxonomicName id="6BD1B1938C3EB65CFF7CF83675332ED4" authority="Besuchet, Cuccodoro, Sabella 2013: 27" authorityName="Besuchet, Cuccodoro, Sabella" authorityPageNumber="27" authorityYear="2013" box="[151,837,1970,1992]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="monoceros">
<emphasis id="9EA516028C3EB65CFF7CF83677D62ED4" box="[151,416,1970,1992]" italics="true" pageId="6" pageNumber="69">Tychobythinus monoceros</emphasis>
Besuchet, Cuccodoro, Sabella 2013: 27
</taxonomicName>
, figs 5 (habitus), 6 (head), 7 (aedeagus) and 9 (head and
</paragraph>
<paragraph id="AC6ECA108C3EB65CFF2CF855774F2EFA" blockId="6.[151,1436,1908,2022]" box="[199,313,2001,2022]" pageId="6" pageNumber="69">pronotum).</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3FB65DFF7CFF3877D52823" pageId="7" pageNumber="70" type="materials_examined">
<paragraph id="AC6ECA108C3FB65DFF7CFF3877E029E4" blockId="7.[151,1436,188,357]" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFF7CFF38774B29C9" bold="true" box="[151,317,188,213]" pageId="7" pageNumber="70">
<typeStatus id="736A74B28C3FB65DFF7CFF3876A429C9" box="[151,210,188,213]" pageId="7" pageNumber="70">Type</typeStatus>
material
</emphasis>
.
<emphasis id="9EA516028C3FB65DFEA0FF3877CE29C9" bold="true" box="[331,440,188,213]" pageId="7" pageNumber="70">
<typeStatus id="736A74B28C3FB65DFEA0FF3877CE29C9" box="[331,440,188,213]" pageId="7" pageNumber="70" type="holotype">Holotype</typeStatus>
</emphasis>
, male.
<emphasis id="9EA516028C3FB65DFDE2FF38750829C9" bold="true" box="[521,894,188,213]" pageId="7" pageNumber="70">
NORTHEASTERN
<collectingCountry id="D4C68A808C3FB65DFD10FF38750829C9" box="[763,894,188,213]" name="Algeria" pageId="7" pageNumber="70">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C3FB65DFC60FF38723929C9" bold="true" box="[907,1103,188,213]" pageId="7" pageNumber="70">Bejaia province:</emphasis>
Adekar (Soummam),
<date id="D86FECD08C3FB65DFAA2FF3873ED29C8" box="[1353,1435,188,212]" pageId="7" pageNumber="70" value="1931-10">X.1931</date>
(H. Normand) (INAT).
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CFE8077D52823" blockId="7.[151,1436,188,357]" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFF2CFE8077DD2801" bold="true" box="[199,427,260,285]" pageId="7" pageNumber="70">Comparative notes</emphasis>
. Within
<taxonomicName id="6BD1B1938C3FB65DFDFBFE8074C72801" box="[528,689,260,285]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3FB65DFDFBFE8074C72801" box="[528,689,260,285]" italics="true" pageId="7" pageNumber="70">Tychobythinus</emphasis>
</taxonomicName>
the presence of a conspicuous frontal horn bearing a long seta is unique to
<taxonomicName id="6BD1B1938C3FB65DFEECFEAD77EA285C" box="[263,412,297,320]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="monoceros">
<emphasis id="9EA516028C3FB65DFEECFEAD77EA285C" box="[263,412,297,320]" italics="true" pageId="7" pageNumber="70">T. monoceros</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3FB65DFF2CFEC8720A2878" box="[199,1148,332,357]" pageId="7" pageNumber="70" type="discussion">
<paragraph id="AC6ECA108C3FB65DFF2CFEC8720A2878" blockId="7.[151,1436,188,357]" box="[199,1148,332,357]" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFF2CFEC877422879" bold="true" box="[199,308,332,357]" pageId="7" pageNumber="70">Remarks</emphasis>
. Species recently attributed to genus
<taxonomicName id="6BD1B1938C3FB65DFD3CFEC8720E2878" authority="Besuchet et al. 2013" authorityName="Besuchet et al." authorityYear="2013" box="[727,1144,332,357]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="9EA516028C3FB65DFD3CFEC8750E2879" box="[727,888,332,357]" italics="true" pageId="7" pageNumber="70">Tychobythinus</emphasis>
(Besuchet et al. 2013)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

263
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<mods:title id="1CA0289C06DCA3FB5E2C14AD8487B2DD">A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<paragraph id="AC6ECA108C3FB65DFF7CFE3C74FD28CE" blockId="7.[151,651,440,499]" box="[151,651,440,466]" pageId="7" pageNumber="70">
<heading id="F7267D7C8C3FB65DFF7CFE3C74FD28CE" bold="true" box="[151,651,440,466]" fontSize="11" level="1" pageId="7" pageNumber="70" reason="1">
<taxonomicName id="6BD1B1938C3FB65DFF7CFE3C74FD28CE" ID-CoL="7CYDT" authority="Jeannel, 1956" authorityName="Jeannel" authorityYear="1956" box="[151,651,440,466]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3FB65DFF7CFE3C74FD28CE" bold="true" box="[151,651,440,466]" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFF7CFE3C77B128CE" bold="true" box="[151,455,440,466]" italics="true" pageId="7" pageNumber="70">Tychobythinus normandi</emphasis>
(Jeannel, 1956)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF7CFE5E775B28EF" blockId="7.[151,651,440,499]" box="[151,301,474,499]" pageId="7" pageNumber="70">
(
<figureCitation id="34EAD6958C3FB65DFF74FE5E775028EF" box="[159,294,474,499]" captionStart="FIGURES 10 – 13" captionStartId="8.[151,264,1483,1505]" captionTargetBox="[157,1436,267,1450]" captionTargetId="figure@8.[150,1436,245,1460]" captionTargetPageId="8" captionText="FIGURES 10 – 13. Tychobythinus normandi. Scale: 0, 1 mm. 10 — Head, schematic drawing. 11 — Left maxillary palpus, holotype. 12 — Right antenna, holotype. 13 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284498/files/figure.png" pageId="7" pageNumber="70">Figs. 10–13</figureCitation>
)
</paragraph>
</subSubSection>
<paragraph id="AC6ECA108C3FB65DFF7CFDA674CE2B6A" pageId="7" pageNumber="70">
<table id="DED138B08C3F49A5FF7CFDA673ED2B6A" box="[151,1435,546,630]" gridcols="12" gridrows="3" pageId="7" pageNumber="70">
<tr id="12E1C8528C3F49A5FF7CFDA673ED2B24" box="[151,1435,546,568]" gridrow="0" pageId="7" pageNumber="70">
<th id="5130A12E8C3F49A5FF7CFDA674CD2B24" box="[151,699,546,568]" gridcol="0" gridrow="0" pageId="7" pageNumber="70">
<taxonomicName id="6BD1B1938C3FB65DFF7CFDA674372B24" authority="Jeannel 1956: 60" authorityName="Jeannel" authorityPageNumber="60" authorityYear="1956" box="[151,577,546,568]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3FB65DFF7CFDA677FC2B24" box="[151,394,546,568]" italics="true" pageId="7" pageNumber="70">Anopsibythus normandi</emphasis>
Jeannel 1956: 60
</taxonomicName>
, 61, figs 50
</th>
<th id="5130A12E8C3F49A5FD28FDA6756D2B24" box="[707,795,546,568]" gridcol="1" gridrow="0" pageId="7" pageNumber="70">(habitus,</th>
<th id="5130A12E8C3F49A5FCC8FDA675342B24" box="[803,834,546,568]" gridcol="2" gridrow="0" pageId="7" pageNumber="70">the</th>
<th id="5130A12E8C3F49A5FCA2FDA675E52B24" box="[841,915,546,568]" gridcol="3" gridrow="0" pageId="7" pageNumber="70">caption</th>
<th id="5130A12E8C3F49A5FC70FDA675992B24" box="[923,1007,546,568]" gridcol="4" gridrow="0" pageId="7" pageNumber="70">wrongly</th>
<th id="5130A12E8C3F49A5FC1CFDA672302B24" box="[1015,1094,546,568]" gridcol="5" gridrow="0" pageId="7" pageNumber="70">reports:</th>
<th id="5130A12E8C3F49A5FBA6FDA672152B24" box="[1101,1123,546,568]" gridcol="6" gridrow="0" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFBA6FDA772152B24" box="[1101,1123,547,568]" italics="true" pageId="7" pageNumber="70">A.</emphasis>
</th>
<th id="5130A12E8C3F49A5FB81FDA672A72B24" box="[1130,1233,546,568]" gridcol="7" gridrow="0" pageId="7" pageNumber="70">
<taxonomicName id="6BD1B1938C3FB65DFB81FDA672A72B24" box="[1130,1233,546,568]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3FB65DFB81FDA672A72B24" box="[1130,1233,546,568]" italics="true" pageId="7" pageNumber="70">Normandi</emphasis>
</taxonomicName>
</th>
<th id="5130A12E8C3F49A5FB32FDA6729A2B24" box="[1241,1260,546,568]" gridcol="8" gridrow="0" pageId="7" pageNumber="70">n.</th>
<th id="5130A12E8C3F49A5FB18FDA673612B24" box="[1267,1303,546,568]" gridcol="9" gridrow="0" pageId="7" pageNumber="70">sp.,</th>
<th id="5130A12E8C3F49A5FAF5FDA673152B24" box="[1310,1379,546,568]" gridcol="10" gridrow="0" pageId="7" pageNumber="70">female</th>
<th id="5130A12E8C3F49A5FA81FDA673ED2B24" box="[1386,1435,546,568]" gridcol="11" gridrow="0" pageId="7" pageNumber="70">from</th>
</tr>
<tr id="12E1C8528C3F49A5FF7CFDC573ED2B4B" box="[151,1435,577,599]" gridrow="1" pageId="7" pageNumber="70" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="5130A12E8C3F49A5FF7CFDC574CD2B4B" box="[151,699,577,599]" gridcol="0" gridrow="1" pageId="7" pageNumber="70">Philippeville), 57 (aedeagus);</th>
</tr>
<tr id="12E1C8528C3F49A5FF7CFDE473ED2B6A" box="[151,1435,608,630]" gridrow="2" pageId="7" pageNumber="70" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="5130A12E8C3F49A5FF7CFDE474CD2B6A" box="[151,699,608,630]" gridcol="0" gridrow="2" pageId="7" pageNumber="70">
<taxonomicName id="6BD1B1938C3FB65DFF7CFDE474C22B6A" authority="Lobl &amp; Besuchet 2004: 315" authorityName="Lobl &amp; Besuchet" authorityPageNumber="315" authorityYear="2004" box="[151,692,608,630]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3FB65DFF7CFDE477E42B6A" box="[151,402,608,630]" italics="true" pageId="7" pageNumber="70">Tychobythinus normandi</emphasis>
Löbl &amp; Besuchet 2004: 315
</taxonomicName>
.
</th>
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</paragraph>
<subSubSection id="E4CB999B8C3FB65DFF7CFD2677722E9F" pageId="7" pageNumber="70" type="materials_examined">
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<emphasis id="9EA516028C3FB65DFF7CFD2677362BA7" bold="true" box="[151,320,674,699]" pageId="7" pageNumber="70">
<typeStatus id="736A74B28C3FB65DFF7CFD2676A42BA7" box="[151,210,674,699]" pageId="7" pageNumber="70">Type</typeStatus>
material
</emphasis>
.
<emphasis id="9EA516028C3FB65DFEBBFD2677CB2BA7" bold="true" box="[336,445,674,699]" pageId="7" pageNumber="70">
<typeStatus id="736A74B28C3FB65DFEBBFD2677CB2BA7" box="[336,445,674,699]" pageId="7" pageNumber="70" type="holotype">Holotype</typeStatus>
</emphasis>
, male.
<emphasis id="9EA516028C3FB65DFDFFFD2675FC2BA7" bold="true" box="[532,906,674,699]" pageId="7" pageNumber="70">
NORTHEASTERN
<collectingCountry id="D4C68A808C3FB65DFCECFD2675FC2BA7" box="[775,906,674,699]" name="Algeria" pageId="7" pageNumber="70">ALGERIA</collectingCountry>
</emphasis>
.
<emphasis id="9EA516028C3FB65DFC71FD2672292BA7" bold="true" box="[922,1119,674,699]" pageId="7" pageNumber="70">Skikda province</emphasis>
: Philippeville (old name of Skikda),
<date id="D86FECD08C3FB65DFF17FD4377392BC3" box="[252,335,711,735]" pageId="7" pageNumber="70" value="1929-10">X.1929</date>
(H. Normand) (INAT).
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CFD6E74382A57" blockId="7.[151,1436,674,2031]" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFF2CFD6E771B2A1F" bold="true" box="[199,365,746,771]" pageId="7" pageNumber="70">Redescription</emphasis>
. Body length
<quantity id="6B2967F58C3FB65DFDF8FD6E740D2A1E" box="[531,635,746,771]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="70" unit="mm" value="1.5">1.50 mm</quantity>
, pubescence consists of long and flattened setae (length:
<quantity id="6B2967F58C3FB65DFAFBFD6E76B22A3B" metricMagnitude="-4" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="7" pageNumber="70" unit="mm" value="0.1275" valueMax="0.13" valueMin="0.125">0.125–0.130 mm</quantity>
) on head, pronotum, elytra and abdomen, other suberected setae short (length:
<quantity id="6B2967F58C3FB65DFBAFFC8B73702A3B" box="[1092,1286,783,807]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="7" pageNumber="70" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.030–0.040 mm</quantity>
) on antennae and legs. Tegument smooth and glossy.
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CFCD276C42C5E" blockId="7.[151,1436,674,2031]" pageId="7" pageNumber="70">
Head (
<figureCitation id="34EAD6958C3FB65DFEFAFCD277142A73" box="[273,354,854,879]" captionStart="FIGURES 10 – 13" captionStartId="8.[151,264,1483,1505]" captionTargetBox="[157,1436,267,1450]" captionTargetId="figure@8.[150,1436,245,1460]" captionTargetPageId="8" captionText="FIGURES 10 – 13. Tychobythinus normandi. Scale: 0, 1 mm. 10 — Head, schematic drawing. 11 — Left maxillary palpus, holotype. 12 — Right antenna, holotype. 13 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284498/files/figure.png" pageId="7" pageNumber="70">Fig. 10</figureCitation>
) strongly wider (
<quantity id="6B2967F58C3FB65DFDC9FCD374E12A73" box="[546,663,855,879]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="70" unit="mm" value="0.32">0.320 mm</quantity>
) than long (
<quantity id="6B2967F58C3FB65DFCF7FCD275E22A73" box="[796,916,854,879]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="70" unit="mm" value="0.25">0.250 mm</quantity>
), frontal lobe width
<quantity id="6B2967F58C3FB65DFB9DFCD3729E2A72" box="[1142,1256,855,879]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="70" unit="mm" value="0.13">0.130 mm</quantity>
with very sharp lateral sides, its dorsal surface glossy with large median sulcus. Clypeal carina evident, very long, and reaching temporal region. Occipital region convex with strong median longitudinal carina reaching posterior edge of vertexal foveae, tempora angulated with deep groove between dorsal surface of occipital region and clypeal carina, sides of tempora prominent ventrally. Gular region deeply excavated in the middle with strongly pubescent lateral edges. From the middle of the posterior edge of this excavation starts a long spiniform median process slightly raised and curved, bearing a short thick tuft of yellow hairs on each side of the apex. II segment of maxillary palpi elongated with 24 tubercles, III segment distinctly longer than wide with 6 tubercles, last segment of maxillary palpi (
<figureCitation id="34EAD6958C3FB65DFF0BFBF277472D93" box="[224,305,1142,1167]" captionStart="FIGURES 10 – 13" captionStartId="8.[151,264,1483,1505]" captionTargetBox="[157,1436,267,1450]" captionTargetId="figure@8.[150,1436,245,1460]" captionTargetPageId="8" captionText="FIGURES 10 – 13. Tychobythinus normandi. Scale: 0, 1 mm. 10 — Head, schematic drawing. 11 — Left maxillary palpus, holotype. 12 — Right antenna, holotype. 13 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284498/files/figure.png" pageId="7" pageNumber="70">Fig. 11</figureCitation>
) about 3 times longer (
<quantity id="6B2967F58C3FB65DFDD6FBF274C22D93" box="[573,692,1142,1167]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="70" unit="mm" value="0.25">0.250 mm</quantity>
) than wide (
<quantity id="6B2967F58C3FB65DFCAFFBF375CB2D93" box="[836,957,1143,1167]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="70" unit="mm" value="0.08">0.080 mm</quantity>
), its lateral margin straight. Antenna (
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)
<quantity id="6B2967F58C3FB65DFF2FFB1E774E2DAE" box="[196,312,1178,1203]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="7" pageNumber="70" unit="mm" value="0.66">0.660 mm</quantity>
long, scape about 3 times longer (
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) than wide (
<quantity id="6B2967F58C3FB65DFC28FB1E724A2DAF" box="[963,1084,1178,1203]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="70" unit="mm" value="0.06">0.060 mm</quantity>
), antennomere II longer (
<quantity id="6B2967F58C3FB65DFAB0FB1E76B22DCB" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="70" unit="mm" value="0.06">0.060 mm</quantity>
) than wide (
<quantity id="6B2967F58C3FB65DFEBBFB3A77BF2DCB" box="[336,457,1214,1239]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="70" unit="mm" value="0.045">0.045 mm</quantity>
), antennomere III longer than wide, antennomeres IV–VIII subequal in size wider than long, V only a little longer and wider than other funicular antennomeres. Antennomeres IX and X short, strongly wider than long, XI about 1.5 times longer than wide, and about 1.25 times longer than combined length of IX and X.
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CFACB75D72C97" blockId="7.[151,1436,674,2031]" pageId="7" pageNumber="70">
Pronotum strongly wider (
<quantity id="6B2967F58C3FB65DFE18FACA741C2C7B" box="[499,618,1358,1383]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="70" unit="mm" value="0.35">0.350 mm</quantity>
) than long (0.300 mm), widest a little before the middle, pleural carina well visible, very deep and close antebasal sulcus with detected edge.
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CFA1274E92CB2" blockId="7.[151,1436,674,2031]" box="[199,671,1430,1455]" pageId="7" pageNumber="70">Elytra width 0.700 mm, length 0.600 mm.</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CFA3F73FE2CCF" blockId="7.[151,1436,674,2031]" box="[199,1416,1466,1491]" pageId="7" pageNumber="70">Metasternum with surface strongly punctated. Laterally of each coxal cavity is present a small pubescent pit.</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CFA5B75E22F07" blockId="7.[151,1436,674,2031]" pageId="7" pageNumber="70">
Legs stout, protrocanthers with 4 tubercles, profemora with 14 small tubercles, base of mesofemora with 4–5 small protuberances on its anterior edge. Metatibiae
<quantity id="6B2967F58C3FB65DFD08F98675232F06" box="[739,853,1538,1563]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="70" unit="mm" value="0.45">0.450 mm</quantity>
long.
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CF9A275582F9B" blockId="7.[151,1436,674,2031]" pageId="7" pageNumber="70">
Aedeagus (
<figureCitation id="34EAD6958C3FB65DFEADF9A277EF2F23" box="[326,409,1574,1599]" captionStart="FIGURES 10 – 13" captionStartId="8.[151,264,1483,1505]" captionTargetBox="[157,1436,267,1450]" captionTargetId="figure@8.[150,1436,245,1460]" captionTargetPageId="8" captionText="FIGURES 10 – 13. Tychobythinus normandi. Scale: 0, 1 mm. 10 — Head, schematic drawing. 11 — Left maxillary palpus, holotype. 12 — Right antenna, holotype. 13 — Aedeagus dorsal view, holotype." httpUri="https://zenodo.org/record/284498/files/figure.png" pageId="7" pageNumber="70">Fig. 13</figureCitation>
)
<quantity id="6B2967F58C3FB65DFE42F9A274792F22" box="[425,527,1574,1599]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="70" unit="mm" value="0.26">0.26 mm</quantity>
length, ovoid with parameres relatively long, sinuates and convergent, apically rounded and narrowed, bearing two bristles for each. Symmetric internal sac armure with numerous large teeth at base and numerous longer and thiner teeth in apical region.
</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CF91677EB2FB7" blockId="7.[151,1436,674,2031]" box="[199,413,1682,1707]" pageId="7" pageNumber="70">Female. Unknown.</paragraph>
<paragraph id="AC6ECA108C3FB65DFF2CF93277722E9F" blockId="7.[151,1436,674,2031]" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFF2CF93277DC2FD3" bold="true" box="[199,426,1718,1743]" pageId="7" pageNumber="70">Comparative notes</emphasis>
.
<taxonomicName id="6BD1B1938C3FB65DFE53F93374372FD3" box="[440,577,1718,1743]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3FB65DFE53F93374372FD3" box="[440,577,1718,1743]" italics="true" pageId="7" pageNumber="70">T. normandi</emphasis>
</taxonomicName>
is distinguished from other species of the
<taxonomicName id="6BD1B1938C3FB65DFBCBF93272172FD3" box="[1056,1121,1718,1743]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3FB65DFBCBF93272172FD3" box="[1056,1121,1718,1743]" italics="true" pageId="7" pageNumber="70">theryi</emphasis>
</taxonomicName>
group by the angulated and excavated tempora. It is comparable with
<taxonomicName id="6BD1B1938C3FB65DFD9DF95875782FEF" box="[630,782,1756,1779]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="monoceros">
<emphasis id="9EA516028C3FB65DFD9DF95875782FEF" box="[630,782,1756,1779]" italics="true" pageId="7" pageNumber="70">T. monoceros</emphasis>
</taxonomicName>
and
<taxonomicName id="6BD1B1938C3FB65DFCA3F95875E92FEF" box="[840,927,1754,1779]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="thery">
<emphasis id="9EA516028C3FB65DFCA3F95875E92FEF" box="[840,927,1754,1779]" italics="true" pageId="7" pageNumber="70">T. thery</emphasis>
</taxonomicName>
in the features of IV–VIII antennal segment which are at most as long as wide (longer than wide in all other species of
<taxonomicName id="6BD1B1938C3FB65DFC34F97A72562E0B" box="[991,1056,1790,1815]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3FB65DFC34F97A72562E0B" box="[991,1056,1790,1815]" italics="true" pageId="7" pageNumber="70">theryi</emphasis>
</taxonomicName>
group) and also with
<taxonomicName id="6BD1B1938C3FB65DFAFCF97B73022E0B" box="[1303,1396,1790,1815]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3FB65DFAFCF97B73022E0B" box="[1303,1396,1790,1815]" italics="true" pageId="7" pageNumber="70">T. theryi</emphasis>
</taxonomicName>
for the punctated surface of metasternum (unpunctated in all other species of
<taxonomicName id="6BD1B1938C3FB65DFC0BF8A672572E27" box="[992,1057,1826,1851]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3FB65DFC0BF8A672572E27" box="[992,1057,1826,1851]" italics="true" pageId="7" pageNumber="70">theryi</emphasis>
</taxonomicName>
group), with
<taxonomicName id="6BD1B1938C3FB65DFB56F8A0733E2E27" box="[1213,1352,1828,1851]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="araneipes">
<emphasis id="9EA516028C3FB65DFB56F8A0733E2E27" box="[1213,1352,1828,1851]" italics="true" pageId="7" pageNumber="70">T. araneipes</emphasis>
</taxonomicName>
for the clypeal carina reaching temporal region and with
<taxonomicName id="6BD1B1938C3FB65DFD20F8C375592E43" box="[715,815,1862,1887]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="bordei">
<emphasis id="9EA516028C3FB65DFD20F8C375592E43" box="[715,815,1862,1887]" italics="true" pageId="7" pageNumber="70">T. bordei</emphasis>
</taxonomicName>
for the anterior margin of mesofemora with 4–5 small tubercles.
</paragraph>
</subSubSection>
<subSubSection id="E4CB999B8C3FB652FF2CF80A768229C8" lastPageId="8" lastPageNumber="71" pageId="7" pageNumber="70" type="discussion">
<paragraph id="AC6ECA108C3FB65DFF2CF80A74142EF2" blockId="7.[151,1436,674,2031]" pageId="7" pageNumber="70">
<emphasis id="9EA516028C3FB65DFF2CF80A77422EBB" bold="true" box="[199,308,1934,1959]" pageId="7" pageNumber="70">Remarks</emphasis>
. Label on the slide preparation of aedeagus reads:
<taxonomicName id="6BD1B1938C3FB65DFC9BF80A73312EBB" authority="Guill., Philippeville. The" authorityName="Guill., Philippeville. The" box="[880,1351,1934,1959]" class="Insecta" family="Staphylinidae" genus="Euplectus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3FB65DFC9BF80A72512EBB" box="[880,1063,1934,1959]" italics="true" pageId="7" pageNumber="70">Euplectus theryi</emphasis>
Guill., Philippeville. The
</taxonomicName>
second label reads:
<taxonomicName id="6BD1B1938C3FB65DFEF5F83674102ED7" authority="Guill. Finally" authorityName="Guill. Finally" box="[286,614,1970,1995]" class="Insecta" family="Staphylinidae" genus="Bythinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="9EA516028C3FB65DFEF5F83677BE2ED7" box="[286,456,1970,1995]" italics="true" pageId="7" pageNumber="70">Bythinus theryi</emphasis>
Guill. Finally
</taxonomicName>
, and the third label written by Besuchet and glued on the opposite side of the slide reads:
<taxonomicName id="6BD1B1938C3FB65DFEADF852742D2EF3" box="[326,603,2006,2031]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="normandi">
<emphasis id="9EA516028C3FB65DFEADF852742D2EF3" box="[326,603,2006,2031]" italics="true" pageId="7" pageNumber="70">Tychobythinus normandi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="AC6ECA108C30B652FF2CFF13768229C8" blockId="8.[151,1436,151,212]" pageId="8" pageNumber="71">
The
<typeStatus id="736A74B28C30B652FF12FF13772D29AC" box="[249,347,151,176]" pageId="8" pageNumber="71" type="holotype">holotype</typeStatus>
specimen is housed in the INAT, while the slide with the preparation of aedeagus is housed in the MNHN.
</paragraph>
</subSubSection>
</treatment>
</document>

214
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<mods:namePart>Lin, Xiao-Bin</mods:namePart>
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<mods:namePart>Peng, Zhong</mods:namePart>
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<subSection pageId="0" pageNumber="15" type="results">
<treatment ID-GBIF-Taxon="183909178" LSID="urn:lsid:zoobank.org:act:A47F4DF3-D990-4DEB-A7C9-0F2F3891E52D" httpUri="http://treatment.plazi.org/id/2478B7584B355CE58C26B029AF1F493E" lastPageNumber="15" pageId="0" pageNumber="15">
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<taxonomicName LSID="http://zoobank.org/A47F4DF3-D990-4DEB-A7C9-0F2F3891E52D" authority="Peng &amp; Lin" authorityName="Peng &amp; Lin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lathrobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lathrobium wangxingmini" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="wangxingmini" status="sp. nov.">Lathrobium wangxingmini Peng &amp; Lin</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="15">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="description">
<paragraph pageId="0" pageNumber="15">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus of Lathrobium spp. A L. yangyihani B L. jiaxingyangi C L. wangxingmini. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1054.68991.figure1" httpUri="https://binary.pensoft.net/fig/572868" pageId="0" pageNumber="15">Figures 1C</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">, 4</figureCitation>
<figureCitation captionStart="Figures 5–7" captionStartId="F5" captionText="Figures 5 - 7. Collecting sites on the Nanling National Nature Reserve 5 type localities of Lathrobium yangyihani (1,260 m) 6 type locality of Lathrobium jiaxingyangi (1,850 m) 7 type locality of Lathrobium wangxingmini (1,090 m)." figureDoi="10.3897/zookeys.1054.68991.figures5-7" httpUri="https://binary.pensoft.net/fig/572872" pageId="0" pageNumber="15">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="type material">
<paragraph pageId="0" pageNumber="15">Type material.</paragraph>
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">
<emphasis italics="true" pageId="0" pageNumber="15">Holotype</emphasis>
:
</emphasis>
♂, labeled 'China: Guangdong Prov., Ruyuan County, Nanling National Nature Reserve, Walkway,
<geoCoordinate degrees="24" direction="north" minutes="55" orientation="latitude" precision="15" seconds="57" value="24.932499">24°55'57&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="00" orientation="longitude" precision="15" seconds="18" value="113.005">113°00'18&quot;E</geoCoordinate>
, 1,220 m, 28.IV.2015, Peng, Tu &amp; Zhou leg.' (SNUC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="15">Paratypes</emphasis>
: 1 ♀, labeled 'China: Guangdong Prov., Ruyuan County, Nanling National Nature Reserve, Shikengkong,
<geoCoordinate degrees="24" direction="north" minutes="55" orientation="latitude" precision="15" seconds="33" value="24.925833">24°55'33&quot;N</geoCoordinate>
,
<geoCoordinate degrees="112" direction="east" minutes="59" orientation="longitude" precision="15" seconds="29" value="112.99139">112°59'29&quot;E</geoCoordinate>
, 1,820 m, 30.IV.2015, Peng, Tu &amp; Zhou leg.' (SNUC); 1 ♀, labeled 'China: Guangdong Prov., Ruyuan County, Nanling National Nature Reserve, Shikengkong,
<geoCoordinate degrees="24" direction="north" minutes="55" orientation="latitude" precision="15" seconds="38" value="24.927221">24°55'38&quot;N</geoCoordinate>
,
<geoCoordinate degrees="112" direction="east" minutes="59" orientation="longitude" precision="15" seconds="30" value="112.99166">112°59'30&quot;E</geoCoordinate>
, 1,850 m, 27.IV.2015, Peng, Tu &amp; Zhou
<normalizedToken originalValue="leg’">leg'</normalizedToken>
(SNUC); 1 ♀, labeled 'China: Hunan Prov., Yizhang County, Mangshan Nature reserve,
<geoCoordinate degrees="24" direction="north" minutes="56" orientation="latitude" precision="15" seconds="26" value="24.940556">24°56'26&quot;N</geoCoordinate>
,
<geoCoordinate degrees="112" direction="east" minutes="59" orientation="longitude" precision="15" seconds="18" value="112.98833">112°59'18&quot;E</geoCoordinate>
, 1,400 m, 26.IV.2015, Peng, Tu &amp; Zhou leg.' (SNUC); 1 ♂, 2 ♀♀, labeled 'China: Guangdong Prov., Ruyuan County, Nanling National Nature Reserve, 1,090 m, 18.VI.2007, Huang &amp; Xu leg.' (SNUC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="description">
<paragraph pageId="0" pageNumber="15">Description.</paragraph>
<paragraph pageId="0" pageNumber="15">Measurements (in mm) and ratios: BL 8.62-10.56, FL 3.48-4.27, HL 1.30-1.67, HW 1.30-1.52, AnL 2.32-2.70, PL 1.57-1.85, PW 1.33-1.57, EL 0.61-0.83, AL 2.25, HL/HW 1.00-1.15, HW/PW 0.96-0.98, HL/PL 0.82-0.95, PL/PW 1.18-1.19, EL/PL 0.39-0.45.</paragraph>
<paragraph pageId="0" pageNumber="15">
Habitus as in Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4C</figureCitation>
. Body blackish brown, legs brown, antennae dark to light brown.
</paragraph>
<caption doi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">Figure 4.</emphasis>
<taxonomicName authorityName="Peng &amp; Lin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lathrobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lathrobium wangxingmini" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="wangxingmini">
<emphasis italics="true" pageId="0" pageNumber="15">Lathrobium wangxingmini</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="15">A</emphasis>
female tergite VIII
<emphasis bold="true" pageId="0" pageNumber="15">B</emphasis>
female sternite VIII
<emphasis bold="true" pageId="0" pageNumber="15">C</emphasis>
female tergites IX-X.
<emphasis bold="true" pageId="0" pageNumber="15">D</emphasis>
male sternite VII
<emphasis bold="true" pageId="0" pageNumber="15">E</emphasis>
male sternite VIII
<emphasis bold="true" pageId="0" pageNumber="15">F</emphasis>
aedeagus in ventral view
<emphasis bold="true" pageId="0" pageNumber="15">G</emphasis>
aedeagus in lateral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="15">Head transverse; punctation coarse and dense, sparser in median dorsal portion; interstices with distinct microsculpture.</paragraph>
<paragraph pageId="0" pageNumber="15">Pronotum nearly parallel-sided; punctation somewhat sparser than that of head; impunctate midline broad; interstices without microsculpture. Eyes moderately big and composed of approximately 80 ommatidia.</paragraph>
<paragraph pageId="0" pageNumber="15">Elytral punctation dense and defined. Hind wings completely reduced. Protarsi without appreciable sexual dimorphism, distinctly dilated.</paragraph>
<paragraph pageId="0" pageNumber="15">Abdomen with fine and dense punctation, punctation of tergite VII slightly less dense than that of anterior tergites; interstices with shallow microsculpture; posterior margin of tergite VII without palisade fringe.</paragraph>
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">Male.</emphasis>
Sternites III-VI unmodified; sternite VII (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4D</figureCitation>
) strongly transverse, with shallow median impression posteriorly with numerous distinctly modified, short black setae, posterior margin nearly truncate; sternite VIII (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4E</figureCitation>
) strongly modified and of distinctive shape and chaetotaxy, with deep and extensive median impression, middle of this impression with unmodified pubescence, laterally with dense short and very stout black setae, posterior margin deep and weakly asymmetric; aedeagus as in Figure
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4F and G</figureCitation>
; ventral process asymmetric and apically acute; dorsal plate with long apical portion and very short basal portion; internal sac with one long sclerotized spine.
</paragraph>
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">Female.</emphasis>
Posterior margin of tergite VIII (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4A</figureCitation>
) strongly convex. Posterior margin of sternite VIII (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4B</figureCitation>
) strongly convex and with moderately dense micropubescence; tergite IX (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4C</figureCitation>
) with short antero-median portion and slender postero-lateral processes; tergite X (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lathrobium wangxingmini. A female tergite VIII B female sternite VIII C female tergites IX-X. D male sternite VII E male sternite VIII F aedeagus in ventral view G aedeagus in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1054.68991.figure4" httpUri="https://binary.pensoft.net/fig/572871" pageId="0" pageNumber="15">4C</figureCitation>
) 2.5 times as long as antero-median portion of tergite IX.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="comparisons">
<paragraph pageId="0" pageNumber="15">Comparative notes.</paragraph>
<paragraph pageId="0" pageNumber="15">
<taxonomicName authorityName="Peng &amp; Lin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lathrobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lathrobium wangxingmini" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="wangxingmini">
<emphasis italics="true" pageId="0" pageNumber="15">Lathrobium wangxingmini</emphasis>
</taxonomicName>
resembles
<taxonomicName lsidName="L. jiaxingyangi" pageId="0" pageNumber="15" rank="species" species="jiaxingyangi">
<emphasis italics="true" pageId="0" pageNumber="15">L. jiaxingyangi</emphasis>
</taxonomicName>
sp. nov. in having the similarly derived chaetotaxy of the asymmetric male sternite VIII and the long dorsal plate of the aedeagus. It is distinguished from
<taxonomicName lsidName="L. jiaxingyangi" pageId="0" pageNumber="15" rank="species" species="jiaxingyangi">
<emphasis italics="true" pageId="0" pageNumber="15">L. jiaxingyangi</emphasis>
</taxonomicName>
by the darker coloration, the larger body size, more dense short black setae in the impression of the male sternite VII, the deeper posterior excision of the male sternite VIII and the morphology of the aedeagus (shape of ventral process; internal sac with one longer sclerotized spine).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="etymology">
<paragraph pageId="0" pageNumber="15">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="15">The species is dedicated to Xing-Min Wang, who supported us on our field trips.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="distribution">
<paragraph pageId="0" pageNumber="15">Distribution and natural history.</paragraph>
<paragraph pageId="0" pageNumber="15">
The species was found in five adjacent localities in western Ruyuan County to southern Yizhang County. Some specimens were sifted from the leaf litter of mixed deciduous forests at an altitude of 1,090 m (Fig.
<figureCitation captionStart="Figures 5–7" captionStartId="F5" captionText="Figures 5 - 7. Collecting sites on the Nanling National Nature Reserve 5 type localities of Lathrobium yangyihani (1,260 m) 6 type locality of Lathrobium jiaxingyangi (1,850 m) 7 type locality of Lathrobium wangxingmini (1,090 m)." figureDoi="10.3897/zookeys.1054.68991.figures5-7" httpUri="https://binary.pensoft.net/fig/572872" pageId="0" pageNumber="15">7</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1054.68991.figures5-7" httpUri="https://binary.pensoft.net/fig/572872" pageId="0" pageNumber="15" start="Figures 5–7" startId="F5">
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">Figures 5-7.</emphasis>
Collecting sites on the Nanling National Nature Reserve
<emphasis bold="true" pageId="0" pageNumber="15">5</emphasis>
type localities of
<taxonomicName authorityName="Lin &amp; Peng" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lathrobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lathrobium yangyihani" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="yangyihani">
<emphasis italics="true" pageId="0" pageNumber="15">Lathrobium yangyihani</emphasis>
</taxonomicName>
(1,260 m)
<emphasis bold="true" pageId="0" pageNumber="15">6</emphasis>
type locality of
<taxonomicName authorityName="Lin &amp; Peng" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lathrobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lathrobium jiaxingyangi" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="jiaxingyangi">
<emphasis italics="true" pageId="0" pageNumber="15">Lathrobium jiaxingyangi</emphasis>
</taxonomicName>
(1,850 m)
<emphasis bold="true" pageId="0" pageNumber="15">7</emphasis>
type locality of
<taxonomicName authorityName="Peng &amp; Lin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lathrobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lathrobium wangxingmini" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="wangxingmini">
<emphasis italics="true" pageId="0" pageNumber="15">Lathrobium wangxingmini</emphasis>
</taxonomicName>
(1,090 m).
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>