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<mods:title>The beetle fauna (Insecta, Coleoptera) of the Rawdhat Khorim National Park, Central Saudi Arabia</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Abdel-Dayem, Mahmoud S.</mods:namePart>
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<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
<mods:nameIdentifier type="email">mseleem@ksu.edu.sa</mods:nameIdentifier>
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</mods:role>
<mods:namePart>Fad, Hassan H.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>El-Torkey, Ashraf M.</mods:namePart>
<mods:affiliation>Plant Protection Research Institute, Agriculture Research Center, Giza, Egypt</mods:affiliation>
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<mods:namePart>Elgharbawy, Ali A.</mods:namePart>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia &amp; Zoology Department, Faculty of Science, Al Azhar University, Nasr City, Cairo, Egypt</mods:affiliation>
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<mods:namePart>Aldryhim, Yousif N.</mods:namePart>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
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</mods:role>
<mods:namePart>Kondratieff, Boris C.</mods:namePart>
<mods:affiliation>Department of Bioagricultural Sciences and Pest Management, Colorado State University, Campus Delivery 1177, Fort Collins, Colorado, U. S. A. 80523</mods:affiliation>
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</mods:role>
<mods:namePart>Ansi, Amin N. Al</mods:namePart>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aldhafer, Hathal M.</mods:namePart>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
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Phaeadoretus syriacus (C.
<normalizedToken originalValue="É">E</normalizedToken>
. Blanchard, 1851)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="distribution">
<paragraph pageId="26" pageNumber="27">World distribution.</paragraph>
<paragraph pageId="26" pageNumber="27">
<emphasis bold="true" pageId="26" pageNumber="27">Asia</emphasis>
: IQ, IR. New to Arabian Peninsula.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="distribution">
<paragraph pageId="26" pageNumber="27">General distribution.</paragraph>
<paragraph pageId="26" pageNumber="27">SAR.</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="collecting month and method">
<paragraph pageId="26" pageNumber="27">Collecting month and method.</paragraph>
<paragraph pageId="26" pageNumber="27">
Abundant species. The specimens were collected by PT under canopies of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia ehrenbergiana" order="Fabales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="ehrenbergiana">
<emphasis italics="true" pageId="26" pageNumber="27">Acacia ehrenbergiana</emphasis>
</taxonomicName>
,
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<emphasis italics="true" pageId="26" pageNumber="27">Acacia gerrardii</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Roem &amp; Schult" authorityYear="1819" class="Dicotyledoneae" family="Solanaceae" genus="Lycium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lycium shawii" order="Tubiflorae" pageId="26" pageNumber="27" phylum="Angiospermae" rank="species" species="shawii">
<emphasis italics="true" pageId="26" pageNumber="27">Lycium shawii</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" family="Rhamnaceae" genus="Ziziphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ziziphus nummularia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="nummularia">
<emphasis italics="true" pageId="26" pageNumber="27">Ziziphus nummularia</emphasis>
</taxonomicName>
; and by HP and LT through I-V.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,106 @@
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<taxonomicName authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Scrophulariaceae" genus="Chelone" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" phylum="Magnoliophyta" rank="species" species="glabra">
<emphasis bold="true" pageNumber="409">Chelone glabra</emphasis>
Linnaeus
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<subSubSection pageNumber="409" type="distribution">
<paragraph pageNumber="409">&quot;Habitat in Virginia, Canada.&quot; RCN: 4419.</paragraph>
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<paragraph pageNumber="409">
<materialsCitation ID-GBIF-Occurrence="2612073251" collectionCode="LINN UPS" specimenCode="765.1" typeStatus="lectotype">
<typeStatus>Lectotype</typeStatus>
(Reveal &amp; al. in
<emphasis italics="true" pageNumber="409">Huntia</emphasis>
7: 220. 1987):
<emphasis italics="true" pageNumber="409">Kalm</emphasis>
, Herb. Linn. No. 765.1 (
<collectionCode>LINN</collectionCode>
;
<typeStatus>iso-</typeStatus>
<collectionCode>UPS</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<paragraph pageNumber="409">
<typeStatus>Generitype</typeStatus>
of
<taxonomicName authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Scrophulariaceae" genus="Chelone" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageNumber="409">Chelone</emphasis>
Linnaeus
</taxonomicName>
(vide Pennell in
<emphasis italics="true" pageNumber="409">Proc. Acad. Nat. Sci. Philadelphia</emphasis>
82: 20. 1930).
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<subSubSection pageNumber="409" type="nomenclature">
<paragraph pageNumber="409">
Current name:
<taxonomicName authority="L." authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Scrophulariaceae" genus="Chelone" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" phylum="Magnoliophyta" rank="species" species="glabra">
<emphasis italics="true" pageNumber="409">
<emphasis bold="true" italics="true" pageNumber="409">Chelone glabra</emphasis>
</emphasis>
L.
</taxonomicName>
(
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<treatmentCitation id="876D7A01F51365FA0CAACF290ED354A7" author="Linnaeus" pageNumber="846" year="1753">
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<taxonomicName id="ED1BE83D4B712D4315AA9F0AF71F1A4D" ID-CoL="GW62" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="campestris">
<emphasis id="7334504793E1BF27691D7DEC28749806" bold="true" pageNumber="317">Artemisia campestris</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="FFDF2FB85A74A9F59A926E3869A2F6DF">
<emphasis id="D4F59720E66E2756F3DDA0958788F74D" italics="true" pageNumber="317">Species Plantarum</emphasis>
2
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: 846. 1753
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</treatmentCitation>
.
</paragraph>
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<paragraph id="E756202E38BA0D83263A7A145EE38E13" pageNumber="317">&quot;Habitat in Europae campis apricis, aridis.&quot; RCN: 6130.</paragraph>
</subSubSection>
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<paragraph id="A4DFB911B4B0C5F910CE9F439A0E54FE" pageNumber="317">
<materialsCitation id="549F50D387E201D4AE3817A2C458D04A" ID-GBIF-Occurrence="2612070413" collectionCode="LINN" specimenCode="988.16" typeStatus="lectotype">
<typeStatus id="574AB53B8B399140CA91B5D80E95CDD3">Lectotype</typeStatus>
(Alavi in Jafri &amp; El-Gadi,
<emphasis id="C5777853F8E9C5A13E64031D25E16377" italics="true" pageNumber="317">Fl. Libya</emphasis>
107: 184. 1983): Herb. Linn. No. 988.16 (
<collectionCode id="AB129218ADB2CF6BCE24C1913E56C8DF">LINN</collectionCode>
)
</materialsCitation>
.
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<paragraph id="7D02944868A7ECAD87867791FEC00FA8" pageNumber="317">
Current name:
<taxonomicName id="3F0355BB5733EBAE5D78E7A5ECC82208" ID-CoL="GW62" ID-ENA="72337" authority="L." authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="campestris">
<emphasis id="A529D26DE10F7E74EC21E8D38CFDD86B" italics="true" pageNumber="317">
<emphasis id="BD64E83FF1B7AA1ADA14AEFF970FFE5D" bold="true" italics="true" pageNumber="317">Artemisia campestris</emphasis>
</emphasis>
L.
</taxonomicName>
(
<taxonomicName id="8A17C4559541CEA360E743C00A4F4BBA" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
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<taxonomicName authority="Prout 1935" authorityYear="1935" class="Insecta" family="Geometridae" genus="Ardonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ardonis (Chloroclystis) thaumasta" order="Lepidoptera" pageId="0" pageNumber="5447" phylum="Arthropoda" rank="species" species="thaumasta" subGenus="Chloroclystis">Ardonis (Chloroclystis) thaumasta (Prout 1935)</taxonomicName>
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<taxonomicName class="Insecta" family="Geometridae" genus="Ardonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ardonis (Chloroclystis) thaumasta" order="Lepidoptera" pageId="0" pageNumber="5447" phylum="Arthropoda" rank="species" species="thaumasta" subGenus="Chloroclystis">Ardonis (Chloroclystis) thaumasta</taxonomicName>
<bibRefCitation author="Prout, L. B." journalOrPublisher="Novitates Zoologicae Tring" pageId="0" pageNumber="5447" pagination="221 - 238" title="New Geometridae from East Java." volume="39" year="1935">Prout 1935</bibRefCitation>
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Type status:
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. Occurrence: sex:
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; Record Level: ownerInstitutionCode: NHM
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<paragraph pageId="0" pageNumber="5447">Distribution</paragraph>
<paragraph pageId="0" pageNumber="5447">Type locality: Java (east), Tengger, Kletak, 6000 ft.</paragraph>
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<paragraph id="05B4D161F654F9C782B549CAFE81CF22" pageId="129" pageNumber="130">Original source.</paragraph>
<paragraph id="1CEFEC0FE7B08150FEC8028D90BA3568" pageId="129" pageNumber="130">
<bibRefCitation id="60C908B8B19441F18CDF92EE74C5BB66" author="Neubauer, TA" journalOrPublisher="Zootaxa" pageId="320" pageNumber="321" pagination="453 - 468" publicationUrl="10.11646/zootaxa.3785.3.7" refId="B335" refString="Neubauer, TA, Harzhauser, M, Georgopoulou, E, Mandic, O, Kroh, A, 2014a. Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda. Zootaxa 3785 (3): 453 - 468, DOI: 10.11646/zootaxa.3785.3.7" title="Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda." url="10.11646/zootaxa.3785.3.7" volume="3785" year="2014 a">Neubauer et al. 2014a</bibRefCitation>
: 456.
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<paragraph id="6BE96766C4DAE9415D70B7D73E49645B" pageId="129" pageNumber="130">Type horizon.</paragraph>
<paragraph id="F739B40D307DA9735581F66DC0A0BBBB" pageId="129" pageNumber="130">Early Pleistocene (?).</paragraph>
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<subSubSection id="1DF0C056D43A287AE45DCDD3B6F40833" pageId="129" pageNumber="130" type="type locality">
<paragraph id="E6A0DD03723850C8DA69085DB69DB51D" pageId="129" pageNumber="130">Type locality.</paragraph>
<paragraph id="F10A67F520E329D6C9ABE1E98F80D6CC" pageId="129" pageNumber="130">&quot;Weganrisse und Regenrisse 3 Km N Antirrion in Akarnanien&quot; (Symeonidis et al. 1986: 342) [sections 3 km N of Antirrio, Aetolia-Acarnania], Greece.</paragraph>
</subSubSection>
<subSubSection id="7EA7E4CB982831CE0707EAFA08E77F82" pageId="129" pageNumber="130" type="types">
<paragraph id="C5B6033DB45E5B56D5014C33AEF2DCED" pageId="129" pageNumber="130">Types.</paragraph>
<paragraph id="AAD6E536192FB988611592C0BCF66FD6" pageId="129" pageNumber="130">Senckenberg Forschungsinstitut und Naturmuseum Frankfurt; no number indicated.</paragraph>
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<paragraph id="C67282C63366B9FF951BF778DA687BC3" pageId="129" pageNumber="130">Remarks.</paragraph>
<paragraph id="4903ED1E8450DB46870351A50B7B92BD" pageId="129" pageNumber="130">
Replacement name for
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, 1986, non Papp, 1953.
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<taxonomicName id="74B9B72223D2E74D04F1C52FB62B7202" LSID="http://zoobank.org/B648B8AE-BE21-44B9-A5A5-D2BD5CEF9EE9" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="2" pageNumber="125" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
<taxonomicNameLabel id="29D20D2D3E6ED55C97E77B48C103BB4E" pageId="2" pageNumber="125">sp. n.</taxonomicNameLabel>
Figures 1, 2, 3, 4
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<paragraph id="6D69C603F84AA42DB518FD344D26398F" pageId="2" pageNumber="125">Holotype.</paragraph>
<paragraph id="972872296C951C34DA80EFB6D81294B6" pageId="2" pageNumber="125">
UARC-IC 539, 23.1 mm SL, male, Colombia, Meta,
<normalizedToken id="D874681504355BCA9E562C50CF221A2B" originalValue="Mapiripán">Mapiripan</normalizedToken>
County, upper Guaviare River drainage,
<normalizedToken id="E8D484F28D2B894B89495A0EDE092BD1" originalValue="Caño">Cano</normalizedToken>
Claro,
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;
<geoCoordinate id="3D00C5AF66299692F82EF3DD7675FB65" direction="west" orientation="longitude" precision="1" value="-72.50411">72°30'14.8&quot;W</geoCoordinate>
; 209 masl; Paratypes. UARC-IC 540, five, 22.1-24.2 mm SL, collected with holotype; UARC-IC 541, two CS, 20.2-22.3 mm SL, collected with holotype; UARC-IC 542, eight, 20.1-23.3 mm SL,
<normalizedToken id="DBA60F123CF2579AC14AA2A4C35C1D8A" originalValue="Mapiripán">Mapiripan</normalizedToken>
county, upper Guaviare River drainage,
<normalizedToken id="29F30FFFC12CE0944C804B7233AF822F" originalValue="Caño">Cano</normalizedToken>
La
<normalizedToken id="3A284B02A63B00F05BE2E7D339CD5D18" originalValue="División">Division</normalizedToken>
,
<geoCoordinate id="FCC7BFDD6246142BE109FFE59E5FDE3A" direction="north" orientation="latitude" precision="1" value="3.1182775">03°07'05.8&quot;N</geoCoordinate>
;
<geoCoordinate id="A55EB0B8B89AE080E07DACA59E186A14" direction="west" orientation="longitude" precision="1" value="-72.543526">72°32'36.7&quot;W</geoCoordinate>
, 209 masl; UARC-IC 543, two, 25.1-28.4 mm SL, El Castillo County, upper Guaviare River drainage,
<normalizedToken id="0FC21AD975671617FB01BF1929B0F350" originalValue="Caño">Cano</normalizedToken>
Hondo,
<geoCoordinate id="06B2B2BFDF04C4B9B63DD96C64E7C6AB" direction="north" orientation="latitude" precision="1" value="3.552389">03°33'08.6&quot;N</geoCoordinate>
;
<geoCoordinate id="9ABD09A8FCAC952CEA17AAAB49E15798" direction="west" orientation="longitude" precision="1" value="-73.78831">73°47'17.9&quot;W</geoCoordinate>
, 209 masl; UARC-IC 544, six, 22.1-23.4 mm SL,
<normalizedToken id="29289FE3EC01CBA4CDA3BF94A698E0FB" originalValue="Mapiripán">Mapiripan</normalizedToken>
County, upper Guaviare River drainage,
<normalizedToken id="CF0B9963E76F8F1C7C27426D966D5B55" originalValue="Caño">Cano</normalizedToken>
<normalizedToken id="0561DDA0A196148522F10F03393B1837" originalValue="División">Division</normalizedToken>
2,
<geoCoordinate id="B55537CA34CE10F6485214B5A95FD460" direction="north" orientation="latitude" precision="1" value="3.1175833">03°07'03.3&quot;N</geoCoordinate>
;
<geoCoordinate id="9B51967989160EAB69C7E1E90671AE1E" direction="west" orientation="longitude" precision="1" value="-72.54236">72°32'32.5&quot;W</geoCoordinate>
, 221 masl; MPUJ 7857, eight, 25.4-31.4 mm SL, El Castillo County,
<normalizedToken id="2F97C4E768A1FD486C56EF30B0A73DFA" originalValue="Caño">Cano</normalizedToken>
Hondo, upper drainage of Guaviare River,
<geoCoordinate id="D9E6F2423C76873AC0CAD544B95E73CF" direction="north" orientation="latitude" precision="1" value="3.552389">03°33'08.6&quot;N</geoCoordinate>
;
<geoCoordinate id="774AEC07DF854FE8E3CBCA4A74202646" direction="west" orientation="longitude" precision="1" value="-73.78831">73°47'17.9&quot;W</geoCoordinate>
.
</paragraph>
<caption id="9815B32EF9946A041C7E742E6DD8F1E3" pageId="2" pageNumber="125">
<paragraph id="895C16CBBA2592B4C7D936F000C0D0C0" pageId="2" pageNumber="125">
Figure 1.
<taxonomicName id="16CFC2170245859860D8CBAB28626AB0" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="2" pageNumber="125" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n., UARC-IC 539, holotype, male, 23.1 mm SL,
<normalizedToken id="1C51E0D18C0249C7BD51C5146C1CB75E" originalValue="Mapiripán">Mapiripan</normalizedToken>
County, upper Guaviare River drainage,
<normalizedToken id="E9ED78089483D056B091A44A746F548D" originalValue="Caño">Cano</normalizedToken>
Claro, Meta, Colombia.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3EFD5037A1933E2D6D6AC6DA218FBA2" lastPageId="3" lastPageNumber="126" pageId="2" pageNumber="125" type="diagnosis">
<paragraph id="4EF0F78F430EE62109BC118F25A92188" pageId="2" pageNumber="125">Diagnosis.</paragraph>
<paragraph id="0B1E7641D6D390AC1499932EA0F996CF" lastPageId="3" lastPageNumber="126" pageId="2" pageNumber="125">
<taxonomicName id="A9981405968AFA4F4A1871207A1B2D1B" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="2" pageNumber="125" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n. can be diagnosed from all other species of
<taxonomicName id="71DEC75980BB159A95158B447B35C196" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon" order="Characiformes" pageId="2" pageNumber="125" phylum="Chordata" rank="genus">Hyphessobrycon</taxonomicName>
, except of
<taxonomicName id="16689E68B59C77FD54974EB4F73DE109" lsidName="H. cyanotaenia" pageId="2" pageNumber="125" rank="species" species="cyanotaenia">H. cyanotaenia</taxonomicName>
,
<taxonomicName id="D719838ABF22512600485B2DC93BC4F9" lsidName="H. loretoensis" pageId="2" pageNumber="125" rank="species" species="loretoensis">H. loretoensis</taxonomicName>
,
<taxonomicName id="2F94715485ACF04759CF8FAC194373DF" lsidName="H. melanostichos" pageId="2" pageNumber="125" rank="species" species="melanostichos">H. melanostichos</taxonomicName>
,
<taxonomicName id="6EEB29BBC337E51E6F3C634CA33D2239" lsidName="H. nigricinctus" pageId="2" pageNumber="125" rank="species" species="nigricinctus">H. nigricinctus</taxonomicName>
,
<taxonomicName id="AEF2CAF0DEA3861FD2F7BBAE05C42F53" lsidName="H. herbertaxelrodi" pageId="2" pageNumber="125" rank="species" species="herbertaxelrodi">H. herbertaxelrodi</taxonomicName>
,
<taxonomicName id="44A3ADEF836D8152958B225553AFE457" lsidName="H. eschwartzae" pageId="2" pageNumber="125" rank="species" species="eschwartzae">H. eschwartzae</taxonomicName>
,
<taxonomicName id="2E1CB545ED4D1DB1CD91AECB71EFF063" lsidName="H. montogoi" pageId="2" pageNumber="125" rank="species" species="montogoi">H. montogoi</taxonomicName>
,
<taxonomicName id="514F9728EABA6809E0BA9086A4DA3CA1" lsidName="H. psittacus" pageId="2" pageNumber="125" rank="species" species="psittacus">H. psittacus</taxonomicName>
,
<taxonomicName id="9E4BEDC1D954C66601076391EFDDF367" lsidName="H. metae" pageId="2" pageNumber="125" rank="species" species="metae">H. metae</taxonomicName>
,
<taxonomicName id="682C18CF1F2DACF9911CBAFB2A8BD556" lsidName="H. margitae" pageId="2" pageNumber="125" rank="species" species="margitae">H. margitae</taxonomicName>
,
<taxonomicName id="87B03B7BE3FB6F8C62BD8C0CEE10377F" lsidName="H. vanzolinii" pageId="2" pageNumber="125" rank="species" species="vanzolinii">H. vanzolinii</taxonomicName>
and
<taxonomicName id="1A73E17C0D286A71026A6537BE0F458E" lsidName="H. peruvianus" pageId="2" pageNumber="125" rank="species" species="peruvianus">H. peruvianus</taxonomicName>
, by having a wide, conspicuous, dark lateral stripe extending from the anterior margin of the eye across the head and body and continuing through the middle caudal-fin rays to their tips. It differs from the species excepted above in having a wider lateral stripe that covers three or four horizontal scale rows (vs. stripe covering just one or two scales rows); in having an orange-yellow stripe extending from the anterodorsal margin of the eye to the caudal peduncle above the dark lateral stripe in life (vs. red lateral stripe extending from the anterodorsal margin of the eye to the caudal peduncle above the dark lateral stripe in live
<taxonomicName id="F1C7167ABFCFFDFC83269BD60FDE62F5" lsidName="H. heterorhabdus" pageId="2" pageNumber="125" rank="species" species="heterorhabdus">H. heterorhabdus</taxonomicName>
,
<taxonomicName id="0438FAB7EB93177D7D59DBA81327C61F" lsidName="H. amapaensis" pageId="3" pageNumber="126" rank="species" species="amapaensis">
<pageBreakToken id="43EF75A5C1289763E98D589F755A6CF5" pageId="3" pageNumber="126" start="start">H</pageBreakToken>
. amapaensis
</taxonomicName>
,
<taxonomicName id="69DF8A41B1D6ECC1B9A88F17AFDBFAF1" lsidName="H. eschwartzae" pageId="3" pageNumber="126" rank="species" species="eschwartzae">H. eschwartzae</taxonomicName>
and
<taxonomicName id="2B1B7EDF18CBD51D21A34FC75BC7520A" lsidName="H. montagi" pageId="3" pageNumber="126" rank="species" species="montagi">H. montagi</taxonomicName>
); by having fewer pored lateral line scales (three to four vs. five to10); fewer lateral scales (21 to 24 vs. 29 to 34); and in having more teeth on the inner premaxillary row (six vs. five). It differs from
<taxonomicName id="E5058288497C13D456FC3D3649A6A4AE" lsidName="H. loretoensis" pageId="3" pageNumber="126" rank="species" species="loretoensis">H. loretoensis</taxonomicName>
in having seven to eight maxillary teeth (vs. three to four) and in having a longer caudal peduncle (12.4-17.0% SL vs. 4.6-8.0% SL). Additionally
<taxonomicName id="443D33E531E7C8EA7195FB6C78DAEF07" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="3" pageNumber="126" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
can be distinguished from the other species of
<taxonomicName id="46596F1818DBD751989288D65182A345" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon" order="Characiformes" pageId="3" pageNumber="126" phylum="Chordata" rank="genus">Hyphessobrycon</taxonomicName>
with a dark lateral stripe from the Orinoco River Basin (
<taxonomicName id="93641494C0ED7EBE0F0DAB4862644F36" lsidName="H. metae" pageId="3" pageNumber="126" rank="species" species="metae">H. metae</taxonomicName>
and
<taxonomicName id="E512C3D0AF4D2FB94502A6D085813D60" lsidName="H. acaciae" pageId="3" pageNumber="126" rank="species" species="acaciae">H. acaciae</taxonomicName>
), in having: two teeth in the outer premaxillary row (vs. three to four see Fig. 2) and 10 branched pectoral-fin rays (vs. 11 to 12). It further differs from
<taxonomicName id="C5F23A70E42BCE85AABF818BAA03DFC1" lsidName="H. metae" pageId="3" pageNumber="126" rank="species" species="metae">H. metae</taxonomicName>
by the length of the snout (17.6-22.8% HL vs. 9.9-15.2% HL) and by the length of the caudal peduncle (12.4-17.0% SL vs. 7.3-11.8% SL).
</paragraph>
<caption id="5FE135819E96DC8D2B0DD247236E78B6" pageId="3" pageNumber="126">
<paragraph id="D04FD9D34FE7579F3135ACF71140E46D" pageId="3" pageNumber="126">
Figure 2. Upper and lower jaws of A
<taxonomicName id="1454FB508B7619A65ADF56A0ADC30A43" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="3" pageNumber="126" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n., UARC-IC 541, 20.2 mm SLB
<taxonomicName id="26830212192D9B578E6FFD55FE864149" lsidName="H. metae" pageId="3" pageNumber="126" rank="species" species="metae">H. metae</taxonomicName>
MCNG 32469, 19.9 mm SL and C
<taxonomicName id="106B4598CB350634B3FC7F594C88E0F8" lsidName="H. acaciae" pageId="3" pageNumber="126" rank="species" species="acaciae">H. acaciae</taxonomicName>
, IUQ 2795 Paratypes, 28.6 mm SL. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A5C26BD6039436BE198844507DCFBA50" lastPageId="5" lastPageNumber="128" pageId="3" pageNumber="126" type="description">
<paragraph id="14864A6A3187A6FF554DF93EB54D5EB6" pageId="3" pageNumber="126">Description.</paragraph>
<paragraph id="5808C92CA6A69CB540A480A3AED9D5AC" pageId="3" pageNumber="126">Morphometric data shown in Table 1. Body compressed, moderately thick, greatest depth between pelvic-fin insertions and dorsal-fin origin. Dorsal profile of head straight from upper lip to vertical through middle of orbit; then convex to dorsal-fin origin. Dorsal-fin base convex, then slightly concave to adipose-fin origin, then straight to base of upper caudal-fin lobe. Ventral profile of head convex from lower lip to insertion of anal fin, anal-fin base straight and then slightly concave to base of lower caudal-fin lobe.</paragraph>
<caption id="3B616A81C9D1BE81E2189D6FD1457A6A" pageId="3" pageNumber="126">
<paragraph id="61C2992C8FD780DEEC3D45D87045479A" pageId="3" pageNumber="126">
Table 1. Morphometric and meristic data of
<taxonomicName id="15AE1F4CEC0508CF52EF8F8BCAF79223" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="3" pageNumber="126" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n. Standard length given in mm. Mean given in parentheses. SD = Standard deviation.
</paragraph>
</caption>
<paragraph id="579D8E678DE8D7272E8EABBA6AC0A4DC" pageId="3" pageNumber="126">
<table id="6745264BCF452A4F5D6C840FBA6A9EFF" pageId="3" pageNumber="126">
<tr id="A8B34CB7840BAD9A6899EE017262EE16" pageId="3" pageNumber="126">
<th id="E01FF63D26EB3EF11252D72FFD95D8E3" colspan="1" pageId="3" pageNumber="126" rowspan="1">Holotype</th>
<th id="3EFDB6DF7EF7C3F6696BF90D35E6F661" colspan="1" pageId="3" pageNumber="126" rowspan="1">Paratypes</th>
<th id="31BF658D53BE43EFFA2C62391FD11C9E" colspan="1" pageId="3" pageNumber="126" rowspan="1">
<abbrev id="ABBRID0EDTAG" pageId="3" pageNumber="126">SD</abbrev>
</th>
</tr>
<tr id="068FEF578FAF0A19D496976FFD20F541" pageId="3" pageNumber="126">
<td id="1F1932293C19604B21E94D77CB4636F5" colspan="4" pageId="3" pageNumber="126" rowspan="1">Percentages of SL</td>
</tr>
<tr id="BFF354BFE5661A40852E343817F56A3A" pageId="3" pageNumber="126">
<td id="BB8710BE7C3CFD19A6B0695DD1B7C8DF" colspan="4" pageId="3" pageNumber="126" rowspan="1">Percentages of HL</td>
</tr>
<tr id="E2D7B9DEE50F6B1D92937FFE230EA87B" pageId="3" pageNumber="126">
<td id="80AC5736F342B4239A7BB9337A424D40" colspan="4" pageId="3" pageNumber="126" rowspan="1">MERISTICS</td>
</tr>
</table>
</paragraph>
<paragraph id="2FFFD35CAFCC871536FDCC0F49225E76" pageId="3" pageNumber="126">Head and snout long, jaws equal, mouth terminal, lips soft and flexible, outer row of premaxillary teeth not exposed. Premaxilla with long, sharp lateral process over ethmoids, with two rows of teeth: the external row with one (2) or two* (23), all tricuspid; inner row with six* (25) tricuspid teeth the gradually diminish in size away from symphysis. Maxillary long and narrow, posterior margin straight, anterior margin convex, ventral margin reaching anterior border of third infraorbital, with seven* (10) or eight (15) tricuspid and conical teeth. Dentary with convex ventral margin, four (25) frontal multicuspid teeth followed by six* (15) or eight (10) smaller conical teeth (Fig. 2).</paragraph>
<paragraph id="66ABC473D464159C61D1DDFE85D1AD9B" lastPageId="5" lastPageNumber="128" pageId="3" pageNumber="126">
Scales cycloid. Lateral line with three (10) or four* (15) pored scales. Lateral scale series with 21 (10), 22 (6) or 24* (9) scales, including those with pores. Transverse
<pageBreakToken id="4D5F8D19A429BAF621CB8C8135964121" pageId="4" pageNumber="127" start="start">scales</pageBreakToken>
rows five* (25) between dorsal-fin origin and lateral line, not including the predorsal series just in front of first dorsal-fin ray. Four (25) horizontal scale rows from anal-fin origin to lateral line. Three (25) horizontal scale rows between pelvic-fin origins and lateral line. Predorsal scales nine*(18) or 10 (7). Five scales in single row along anterior anal-fin base. Fin rays: Dorsal ii, 9 (25). Anal iii, 19 (18) or 20* (7). Pelvic ii,
<pageBreakToken id="BDD9D73C8ADB36E2418F29598AB0D3DA" pageId="5" pageNumber="128" start="start">7</pageBreakToken>
(25). Pectoral ii, 10 (25). Caudal 10+10 (2) principal and 10 (2) procurrent. Caudal fin bifurcate, upper and lower lobes similar in size, pointed. Total vertebrae 32-33.
</paragraph>
</subSubSection>
<subSubSection id="8064A9CD2B8ACDBE6A178A0F5E2D39FA" pageId="5" pageNumber="128" type="sexual dimorphism">
<paragraph id="21959A60C7D3DEC119F10BDB0EC55CCE" pageId="5" pageNumber="128">Sexual dimorphism.</paragraph>
<paragraph id="F950DBA828A0C15A3247AD000DBBBE06" pageId="5" pageNumber="128">
Males have hooks on anal-fin and
<normalizedToken id="94EB483C77E76BCBBF200BABC043C83C" originalValue="pelvicfin">pelvic-fin</normalizedToken>
rays. Anal fin with pair of rows of eight to ten small hooks along third simple ray followed by two to eight pairs of hooks on first to fifth branched rays. Pelvic fins with two to ten pairs of hooks on branched rays, located above internal branch of ray, each segment of branched rays with pair of hooks.
</paragraph>
</subSubSection>
<subSubSection id="3C1529DB7161FAB812AB5C74A2BFB3B2" pageId="5" pageNumber="128" type="color in alcohol">
<paragraph id="2F84D5F886DA1C6B7CA1E106AFD8CDD6" pageId="5" pageNumber="128">Color in alcohol.</paragraph>
<paragraph id="DB3970F3F26BDD37B7F8F8D27E4A16AC" pageId="5" pageNumber="128">Opercular and humeral spots absent. Dorsal part of head and body to dorsal fin dark brown, then yellow on sides and light yellow ventrally. Base color divided by conspicuous, wide (three to four horizontal scale rows), dark lateral stripe from anterior margin of eye through middle caudal-fin rays. Pectoral, pelvic and anal fins hyaline. Dorsal, anal and caudal fins with dark margins. Anal fin with dark pigment concentrated on interradial membranes.</paragraph>
</subSubSection>
<subSubSection id="CED6D7C56ADA3D01B0DB9224DD8E1E85" lastPageId="6" lastPageNumber="129" pageId="5" pageNumber="128" type="color in life">
<paragraph id="D661A15E6BA6C4E508AD3EC11D3E044F" pageId="5" pageNumber="128">Color in life.</paragraph>
<paragraph id="B5E12F7B83E0346852AB428AAB10EBBF" lastPageId="6" lastPageNumber="129" pageId="5" pageNumber="128">
Body greenish-yellow, predorsal area orange-yellow, preventral area silvery-yellow, upper and lower margins of eye red and black respectively, dorsal area of head orange-yellow, ventral area greenish-yellow with great concentration of mel
<pageBreakToken id="453BE363457F2CDDF54ADE8F5DD3921E" pageId="6" pageNumber="129" start="start">anophores</pageBreakToken>
on infraorbitals, preopercle and opercle. Wide, black, lateral stripe from anterior part of eye along sides through middle caudal-fin rays, covering at least half of body height near midbody. Iridescent orange stripe present above black stripe from eye to upper caudal-fin lobe. Lower lobe of caudal fin with orange iridophores at bases of rays. Bases of dorsal-fin rays orange, base of caudal and pelvic fins
<normalizedToken id="2E0CE254A0576B0D059652BD8DB0EEED" originalValue="greenishyellow">greenish-yellow</normalizedToken>
. Adipose fin light orange (Fig. 3).
</paragraph>
<caption id="85593785FE42ABDF6960B97BB66AF2BE" pageId="6" pageNumber="129">
<paragraph id="8794BEFBA31165F6D00D866D9FD42C0D" pageId="6" pageNumber="129">
Figure 3. Live colours of
<taxonomicName id="405C4BF296DF5261186B4B2F77311651" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n. Paratype,
<normalizedToken id="97166C16FBD27C1ECE11097BE5C7C5C6" originalValue="UARCIC">UARC-IC</normalizedToken>
540, 22.4 mm SL.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5490C711F3DC5209B9188FDBF147405E" pageId="6" pageNumber="129" type="distribution">
<paragraph id="AA78A32F448156A044802ECDDC40714D" pageId="6" pageNumber="129">Distribution.</paragraph>
<paragraph id="BBD7B2302B2F349C733127AE6EE9BC72" pageId="6" pageNumber="129">
<taxonomicName id="DCC1E70222E188C6A052CEDC0AAC2C40" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
is known only from the type locality in the upper Guaviare River Orinoco Basin in Colombia (Fig. 4).
</paragraph>
<caption id="556C4B2F2BFA929ADD24DFC7B24AACF4" pageId="6" pageNumber="129">
<paragraph id="E4FB9C6B5243393045DDA6AA3D8817EA" pageId="6" pageNumber="129">
Figure 4. Distribution of
<taxonomicName id="C549B17666215EE9D22681FA86E9F173" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n. in the upper tributaries of Guaviare River drainage. Type locality indicated by an asterisk.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="915DC94FCBAAB20F78166CB2B4D8648E" pageId="6" pageNumber="129" type="etymology">
<paragraph id="615FA298FE7473F5DA40A0A8067006CA" pageId="6" pageNumber="129">Etymology.</paragraph>
<paragraph id="7C4EF27ECCFA6F665C4BC009BE9F30D4" pageId="6" pageNumber="129">
Research leading to the discovery and recognition of this species was partially funded by Mr. Klaus-Peter Lang from Oberhausen, Germany. To commemorate the 80th birthday of his mother, this species is dedicated to and named for his father
<normalizedToken id="5C96B44F458EDBD624AFC04339C93453" originalValue="“Klaus”">&quot;Klaus&quot;</normalizedToken>
and his mother
<normalizedToken id="F4622DFE766724F80A1BDB71C135647D" originalValue="“Anni”">&quot;Anni&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="BDD8510463EBD287521BA49E876C6A35" pageId="6" pageNumber="129" type="remarks">
<paragraph id="6F52F6C7CF57A9509FC116AB7C37A9ED" pageId="6" pageNumber="129">Remarks.</paragraph>
<paragraph id="9E728456FA5CAD0258B3659338057A5C" pageId="6" pageNumber="129">
Principal component analysis (PCA) detected morphological differences among
<taxonomicName id="D34B846A3E5D18B9E2CCB3E45FDF22CC" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
and
<taxonomicName id="2D79DAA309F85BB1C2C813D6D5B930CB" lsidName="H. acaciae" pageId="6" pageNumber="129" rank="species" species="acaciae">H. acaciae</taxonomicName>
,
<taxonomicName id="AFA5CED0475AE69D7365697AC7BFC979" lsidName="H. mavro" pageId="6" pageNumber="129" rank="species" species="mavro">H. mavro</taxonomicName>
,
<taxonomicName id="547726FDECC90CD85D8144D3B73C30A3" lsidName="H. metae" pageId="6" pageNumber="129" rank="species" species="metae">H. metae</taxonomicName>
and
<taxonomicName id="EC93C461FF241774AF7BCDF050B5BA63" lsidName="H. niger" pageId="6" pageNumber="129" rank="species" species="niger">H. niger</taxonomicName>
. For the first component, upper jaw length, postorbital length and dorsal-pectoral fin distance were the most important variables. For the second component, orbital diameter, caudal peduncle length and snout length were most important. The first component explained 43.71% of total variation, and combined with the second this rose to 85.78% (Fig. 5, Tables 2 and 3).
</paragraph>
<caption id="9ECA47D8E2AE87490D4C0ABF44FDADE2" pageId="6" pageNumber="129">
<paragraph id="A3A9916C57336CEA83EB4537BD62DC68" pageId="6" pageNumber="129">
Figure 5. Principal component analysis for morphological data of
<taxonomicName id="210261D3B74B10C41C7B8495B425EF1B" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n. (□),
<taxonomicName id="AF61B12E54BAFF4D31D985F030BF5DA3" lsidName="H. acaciae" pageId="6" pageNumber="129" rank="species" species="acaciae">H. acaciae</taxonomicName>
(+),
<taxonomicName id="C6DBED3D78E922E668A3CCF29D6E4513" lsidName="H. mavro" pageId="6" pageNumber="129" rank="species" species="mavro">H. mavro</taxonomicName>
(▲),
<taxonomicName id="610283ECF1518B1F2CDEB6B196304B7F" lsidName="H. metae" pageId="6" pageNumber="129" rank="species" species="metae">H. metae</taxonomicName>
(○) and
<taxonomicName id="F2E924234761150F69D8413265FAC149" lsidName="H. niger" pageId="6" pageNumber="129" rank="species" species="niger">H. niger</taxonomicName>
(x); component 1 on X axis and component 2 plotted on Y axis. Abbreviations same as Table 1.
</paragraph>
</caption>
<caption id="0ACE840EB600EBDB18AAE8F60CB6DA0E" ID-Table-UUID="0ACE840EB600EBDB18AAE8F60CB6DA0E" httpUri="http://table.plazi.org/id/0ACE840EB600EBDB18AAE8F60CB6DA0E" pageId="6" pageNumber="129">
<paragraph id="E4A1C868B15BCA524F0CD0E6F561D256" pageId="6" pageNumber="129">
Table 2. Eigenvalue for principal components (PC) between
<taxonomicName id="57A6AD996DE7187001616F1E67C41D87" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n.,
<taxonomicName id="B849E0EC5985FD9375EA3F3B6B73BB7C" lsidName="H. acaciae" pageId="6" pageNumber="129" rank="species" species="acaciae">H. acaciae</taxonomicName>
,
<taxonomicName id="65FD16FF3512E6FC311D13299A998CB1" lsidName="H. mavro" pageId="6" pageNumber="129" rank="species" species="mavro">H. mavro</taxonomicName>
,
<taxonomicName id="B995661463F2357E62750D6A13F4FED3" lsidName="H. metae" pageId="6" pageNumber="129" rank="species" species="metae">H. metae</taxonomicName>
and
<taxonomicName id="0FEE8FD942275A2B31FFAF77DCC62748" lsidName="H. niger" pageId="6" pageNumber="129" rank="species" species="niger">H. niger</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="53C085408032953B151191E70FFAE5CA" pageId="6" pageNumber="129">
<table id="B86B54DEE88EC67D136F9ECF6EA4FF6B" pageId="6" pageNumber="129">
<tr id="5E065B969BE47678AA21F26262C0F509" pageId="6" pageNumber="129">
<th id="C6C5D026F58D95C70778905D79F7A3BB" colspan="1" pageId="6" pageNumber="129" rowspan="1">
<abbrev id="ABBRID0EGMBG" pageId="6" pageNumber="129">PC</abbrev>
</th>
<th id="C01CFDDF39071A2C0EBCC24F2179444E" colspan="1" pageId="6" pageNumber="129" rowspan="1">Eigenvalue</th>
<th id="D56146F5BAE299C3B6C7B3D1223D80A4" colspan="1" pageId="6" pageNumber="129" rowspan="1">Percentage variance</th>
</tr>
</table>
</paragraph>
<caption id="CA7225765CF75D95E2A8214D08193526" pageId="6" pageNumber="129">
<paragraph id="E88AC631805039D9A07FBCA562620FF4" pageId="6" pageNumber="129">
Table 3. Eigenvector for first four principal components (PC) between
<taxonomicName id="49AD7DE720C029F0A641E6F59F6AF8C3" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
sp. n.,
<taxonomicName id="4C5A8D65E1BEDB7B60B95A08F9DE2100" lsidName="H. acaciae" pageId="6" pageNumber="129" rank="species" species="acaciae">H. acaciae</taxonomicName>
,
<taxonomicName id="C3EA243DA37D2AE994621EAECE4E65E6" lsidName="H. mavro" pageId="6" pageNumber="129" rank="species" species="mavro">H. mavro</taxonomicName>
,
<taxonomicName id="35DECB783C327740C7691C977719AF08" lsidName="H. metae" pageId="6" pageNumber="129" rank="species" species="metae">H. metae</taxonomicName>
and
<taxonomicName id="218F7678E81C49BA96A916F7BDFC5B63" lsidName="H. niger" pageId="6" pageNumber="129" rank="species" species="niger">H. niger</taxonomicName>
. Abbreviations same as Table 1.
</paragraph>
</caption>
<paragraph id="A7A4EA5771CF24F5B191ADB7792EB1CA" pageId="6" pageNumber="129">
<table id="734AC73EC7EF3CB0AB780E66C981359F" pageId="6" pageNumber="129">
<tr id="912EC78B907356AA06228574B6D8A053" pageId="6" pageNumber="129">
<th id="EE4C9C6C120C2C43C51226E2B8E5E46D" colspan="1" pageId="6" pageNumber="129" rowspan="1">
<abbrev id="ABBRID0EKWBG" pageId="6" pageNumber="129">PC</abbrev>
1
</th>
<th id="096C40CEFEDC86EAD018013A5A00E5C5" colspan="1" pageId="6" pageNumber="129" rowspan="1">
<abbrev id="ABBRID0ESWBG" pageId="6" pageNumber="129">PC</abbrev>
2
</th>
<th id="8ACC1B77E22D7AF09E060FB4A98E60E9" colspan="1" pageId="6" pageNumber="129" rowspan="1">
<abbrev id="ABBRID0E1WBG" pageId="6" pageNumber="129">PC</abbrev>
3
</th>
<th id="107553B863C633567007FAA3AF78B128" colspan="1" pageId="6" pageNumber="129" rowspan="1">
<abbrev id="ABBRID0ECXBG" pageId="6" pageNumber="129">PC</abbrev>
4
</th>
</tr>
<tr id="52D54C10B98EE901A0A51817A40E7ACA" pageId="6" pageNumber="129">
<td id="51599FBCAF0E96A271CF1A02E9AE659C" colspan="1" pageId="6" pageNumber="129" rowspan="1">SL</td>
</tr>
<tr id="DB139DCC30FEB67C125D79C275B47518" pageId="6" pageNumber="129">
<td id="EE15A2BD7651F0FB53AACDDF6C97789F" colspan="1" pageId="6" pageNumber="129" rowspan="1">HL</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="4A598DAB599274BA4F3193D60F985D5D" pageId="6" pageNumber="129" type="ecological note">
<paragraph id="4EE20B95D60298EAEA36DAAC867F9323" pageId="6" pageNumber="129">Ecological note.</paragraph>
<paragraph id="B22F7981503F20D7D938FCEAE16492E2" pageId="6" pageNumber="129">
<taxonomicName id="6EE5DE8D936CB1989C1E0AF85B7E37C6" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
lives in shallow (0.30-1.5 m) well-oxygenated (6.39-7.68 mg/l) streams with transparent waters flowing (0.063 m/s) over different types of substrates (rocks, sand, gravel and decomposing organic material). The temperature range was narrow, 25.5-26°C but pH varied from moderately acidic to basic (6.47-8.7).
<taxonomicName id="6C539A8B8AAA0D7FF01DF0B476E7D3F7" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphessobrycon klausanni" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="klausanni">Hyphessobrycon klausanni</taxonomicName>
was found near shore among aquatic vegetation, tree roots and fallen logs. Other species found at the sites included:
<taxonomicName id="FA05DEE51B13B778F452B207A0A32207" family="Characidae" genus="Hemigrammus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemigrammus barrigonae" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="barrigonae">Hemigrammus barrigonae</taxonomicName>
,
<taxonomicName id="D22DBF170CC563588AAC9CA9DA30E442" family="Characidae" genus="Ctenobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenobrycon spilurus" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="spilurus">Ctenobrycon spilurus</taxonomicName>
,
<taxonomicName id="E63B045824BF5E45813BA6D5BF8A3B63" family="Characidae" genus="Tyttocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyttocharax metae" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="metae">Tyttocharax metae</taxonomicName>
,
<taxonomicName id="21F1D64C8A4E5CC284A384D9EC5D18D6" family="Characidae" genus="Moenkhausia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Moenkhausia oligolepis" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="oligolepis">Moenkhausia oligolepis</taxonomicName>
,
<taxonomicName id="501498004458FD3E44BCB764119AF1FD" lsidName="M. chrysargyrea" pageId="6" pageNumber="129" rank="species" species="chrysargyrea">M. chrysargyrea</taxonomicName>
,
<taxonomicName id="F3C6842CD1973A28FE487DA31E714F7A" lsidName="M. lepidura" pageId="6" pageNumber="129" rank="species" species="lepidura">M. lepidura</taxonomicName>
,
<taxonomicName id="2982D3B91EAA4684DCE1D6151FAC137A" family="Characidae" genus="Chrysobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysobrycon guahibo" order="Characiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="guahibo">Chrysobrycon guahibo</taxonomicName>
,
<taxonomicName id="E2BF79855D15900AE7F4F92CB8779209" family="Trichomycteridae" genus="Ochmacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochmacanthus orinoco" order="Siluriformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="orinoco">Ochmacanthus orinoco</taxonomicName>
,
<taxonomicName id="0F31BE727E1DE1C26DF4F8B0577FD666" family="Loricariidae" genus="Farlowella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Farlowella vittata" order="Siluriformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="vittata">Farlowella vittata</taxonomicName>
,
<taxonomicName id="86B648B65490E22520A76EE18B80B05C" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancistrus triradiatus" order="Siluriformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="triradiatus">Ancistrus triradiatus</taxonomicName>
,
<taxonomicName id="C6DE596CA06967CD559668681A30649F" family="Auchenipteridae" genus="Centromochlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centromochlus reticulatus" order="Siluriformes" pageId="6" pageNumber="129" phylum="Chordata" rank="species" species="reticulatus">Centromochlus reticulatus</taxonomicName>
and
<taxonomicName id="D7214B72875E5F961DA6F3FFBC921B2D" family="Rivulidae" genus="Anablepsoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anablepsoides" order="Cyprinodontiformes" pageId="6" pageNumber="129" phylum="Chordata" rank="genus">Anablepsoides</taxonomicName>
sp. Stomach content analysis (n = 4) revealed a diet of aquatic and terrestrial invertebrates:
<taxonomicName id="D7A3DF1482BCBA2B5A70C22AD7580FA6" lsidName="" order="Coleoptera" pageId="6" pageNumber="129" rank="order">Coleoptera</taxonomicName>
(
<taxonomicName id="AD8D2A08BE4FC04C8D0E3A1D712B0808" family="Dytiscidae" lsidName="" pageId="6" pageNumber="129" rank="family">Dytiscidae</taxonomicName>
),
<taxonomicName id="76E5198DDF9340009B06851628A37FFD" lsidName="" order="Ephemeroptera" pageId="6" pageNumber="129" rank="order">Ephemeroptera</taxonomicName>
,
<taxonomicName id="F5B9C726D82941AA5BA337E905834163" lsidName="" order="Hymenoptera" pageId="6" pageNumber="129" rank="order">Hymenoptera</taxonomicName>
(
<taxonomicName id="1174B73AEA4B4EC8318A03DED96C014A" family="Formicidae" lsidName="" pageId="6" pageNumber="129" rank="family">Formicidae</taxonomicName>
), larvae of
<taxonomicName id="693010BACE3E41B5D54C8096E18BA5FA" family="Chironomidae" lsidName="" pageId="6" pageNumber="129" rank="family">Chironomidae</taxonomicName>
and others not identified due to the degree of fragmentation.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C2CC69FCBEF11002E3BD5B51834C9CEF">Species of Elasmogorgia and Euplexaura (Cnidaria, Octocorallia) from Japan with a discussion about the genus Filigella</mods:title>
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<paragraph id="AD08932B99F2E1FFB2A29A868D8327E4" pageId="6" pageNumber="7">Taxon classification Animalia Alcyonacea Plexauridae</paragraph>
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<taxonomicName id="696534C230C50F733EC3DD7F9F3E6B1B" ID-CoL="3CVGJ" authority="Aurivillius, 1931" authorityName="Aurivillius" authorityYear="1931" class="Anthozoa" family="Plexauridae" genus="Euplexaura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euplexaura boninensis" order="Alcyonacea" pageId="6" pageNumber="7" phylum="Cnidaria" rank="species" species="boninensis">Euplexaura boninensis (Aurivillius, 1931)</taxonomicName>
Figures 1, 2c, 6, 7
</paragraph>
</subSubSection>
<subSubSection id="4A37A45A4F38B327C2E2728D96EE5181" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="6A63EBD21B910AF1940482CDE2101C5B" pageId="6" pageNumber="7">
<taxonomicName id="D7FDB653249BC7B9EAD970B6B9F609A4" class="Anthozoa" family="Plexauridae" genus="Filigella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Filigella boninensis" order="Alcyonacea" pageId="6" pageNumber="7" phylum="Cnidaria" rank="species" species="boninensis">Filigella boninensis</taxonomicName>
Aurivillius, 1931: 139 (Bonin Islands).
</paragraph>
<paragraph id="E0D3B9F42D69E6CB1F2276A103530760" pageId="6" pageNumber="7">
<taxonomicName id="31CBBE883073B733C0996064472101AB" class="Anthozoa" family="Plexauridae" genus="Thesea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thesea boninensis" order="Alcyonacea" pageId="6" pageNumber="7" phylum="Cnidaria" rank="species" species="boninensis">Thesea boninensis</taxonomicName>
;
<bibRefCitation id="BA35118ED951B1BD05086F627D2DA84F" author="Matsumoto, AK" journalOrPublisher="Journal for the Comparative Study of Civilizations (JCSC)" pageId="10" pageNumber="11" pagination="153 - 186" title="The Relation between Telegraph Cables of Great Nordic Ltd. (Store Nordiske) and Japanese Octocoral Collection in Copenhagen, Denmark and UUZM, Uppsala, Sweden." volume="19" year="2014">Matsumoto 2014</bibRefCitation>
: 158 (Table 1, listed only).
</paragraph>
</subSubSection>
<subSubSection id="9AC924927F47A77C2B183849F0EBB25D" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="D9E2A326A291A3220F5871C0B4763D8C" pageId="6" pageNumber="7">Material examined.</paragraph>
<paragraph id="2977D53164967F1987A9F651513E7120" pageId="6" pageNumber="7">Holotype UPSZTY2165 (UUZM 68), East of Chichijima I., Ogasawara Is. (= Bonin Is.), Japan, depth 100 m, in formalin, Dr. Sixten Bock's, Japan Expedition, coll. Dr. Sixten Bock, 1 August 1914.</paragraph>
</subSubSection>
<subSubSection id="303D996F62924E51BECAC1A9B0C4C25C" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="61CD8C3F07CD67B26CF56D741CEF6C9B" pageId="6" pageNumber="7">Diagnosis.</paragraph>
<paragraph id="6D04D1D8254D7623B936E81A40596C85" pageId="6" pageNumber="7">Branches thread-like, 6 cm in length. Calyces dome-shaped, arranged all around the branches (Figure 2c). The polyps have points with flattened spindles, up to 0.15 mm long (Figure 6b), with simple tubercles and spiny distal end. Collaret present, with slightly bent, flattened spindles, up to 0.25 mm long, with simple tubercles (Figure 6c). Tentacles with small scales, up to 0.10 mm long (Figure 6a).</paragraph>
<caption id="358B00BD13F7DA295871823F0B84EB53" pageId="6" pageNumber="7">
<paragraph id="8D9DE999E628EA14BF21B027A4174729" pageId="6" pageNumber="7">
Figure 6.
<taxonomicName id="24FC395F6DC0DDC47AC51BE1C2F92583" class="Anthozoa" family="Plexauridae" genus="Euplexaura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euplexaura boninensis" order="Alcyonacea" pageId="6" pageNumber="7" phylum="Cnidaria" rank="species" species="boninensis">Euplexaura boninensis</taxonomicName>
(Aurivillius, 1931), holotype UPSZTY2165 (UUZM 68), a tentacle scales b point spindles c collaret spindles d spindles from surface layer of coenenchyme e blunt ellipsoids from surface layer of coenenchyme.
</paragraph>
</caption>
<paragraph id="5F493E7E6000A4DA8BAC46E76F43A603" pageId="6" pageNumber="7">The surface layer of the coenenchyme has spindles (Figure 6d) and blunt ellipsoids (Figure 6e), up to 0.65 mm long, with complex tubercles. Several of them with one side less tuberculate. The interior has small spindles and rods, up to 0.25 mm long (Figure 7), with simple tubercles.</paragraph>
<caption id="D30A1301D71DFDBAE45331B3AA3B15B1" pageId="6" pageNumber="7">
<paragraph id="3DB27BDEB6F11567B85A305714D8DE0C" pageId="6" pageNumber="7">
Figure 7.
<taxonomicName id="53C3E5593CBB328F86FAA219900BAE08" class="Anthozoa" family="Plexauridae" genus="Euplexaura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euplexaura boninensis" order="Alcyonacea" pageId="6" pageNumber="7" phylum="Cnidaria" rank="species" species="boninensis">Euplexaura boninensis</taxonomicName>
(Aurivillius, 1931), holotype UPSZTY2165 (UUZM 68), sclerites of interior of coenenchyme.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2872EC4AA76D1E2B577FFA5387B3CCE0" pageId="6" pageNumber="7" type="remarks">
<paragraph id="473D5A5EC671006C2791F7EE715BF50C" pageId="6" pageNumber="7">Remarks.</paragraph>
<paragraph id="C60511FB60745D9355E86F9B9799BEFC" pageId="6" pageNumber="7">
Because the sclerites of this species are spindles and ellipsoids with complex tubercles it actually represents an
<taxonomicName id="04DFEBD996DBDD5549722A6195D34B40" class="Anthozoa" family="Plexauridae" genus="Euplexaura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euplexaura" order="Alcyonacea" pageId="6" pageNumber="7" phylum="Cnidaria" rank="genus">Euplexaura</taxonomicName>
species. It is the only species of
<taxonomicName id="667F9CA25F6545EFA0ADFB49EED024B3" class="Anthozoa" family="Plexauridae" genus="Euplexaura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euplexaura" order="Alcyonacea" pageId="6" pageNumber="7" phylum="Cnidaria" rank="genus">Euplexaura</taxonomicName>
with thread-like colony shape which has many sclerites with one side that is less tuberculate.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName authority="Graenicher, 1911" authorityName="Graenicher" authorityYear="1911" class="Insecta" family="Halictidae" genus="Lasioglossum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasioglossum (Dialictus) nigroviride" order="Hymenoptera" pageId="0" pageNumber="27161" phylum="Arthropoda" rank="species" species="nigroviride" subGenus="Dialictus">Lasioglossum (Dialictus) nigroviride (Graenicher, 1911)</taxonomicName>
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<paragraph pageId="0" pageNumber="27161">Notes</paragraph>
<paragraph pageId="0" pageNumber="27161">Collected from the Lewis and Clark County, Park County and Flathead County sites (Table 1, Suppl. material 1)</paragraph>
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