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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.144.62246" ID-GBIF-Dataset="3776e555-85f5-46ed-8ea1-98ca6a147ede" ID-PMC="PMC8222199" ID-Pensoft-Pub="1313-2970-1044-609" ID-Pensoft-UUID="C96ED07B505A52A8ADA354E1F70BB7C0" ID-PubMed="34183888" ID-ZooBank="66F01A49D32448A8AC2669BF3374894C" ModsDocID="1313-2970-1044-609" checkinTime="1623875354053" checkinUser="pensoft" docAuthor="Smith, Sarah M. & Cognato, Anthony I." docDate="2021" docId="FB800B123BA45B5FAB2239F345F20679" docLanguage="en" docName="ZooKeys 1044: 609-720" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.144.62246" docTitle="Coptoborus cuneatus" docType="treatment" docVersion="4" id="C96ED07B505A52A8ADA354E1F70BB7C0" lastPageNumber="609" masterDocId="C96ED07B505A52A8ADA354E1F70BB7C0" masterDocTitle="A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini)" masterLastPageNumber="720" masterPageNumber="609" pageNumber="609" updateTime="1668150492206" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Smith, Sarah M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
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||||
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cognato, Anthony I.</mods:namePart>
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<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-16</mods:number>
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<mods:detail type="volume">
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<mods:number>1044</mods:number>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.144.62246</mods:identifier>
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<treatment ID-GBIF-Taxon="182753844" LSID="urn:lsid:plazi:treatment:FB800B123BA45B5FAB2239F345F20679" httpUri="http://treatment.plazi.org/id/FB800B123BA45B5FAB2239F345F20679" lastPageNumber="609" pageId="0" pageNumber="609">
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<subSubSection pageId="0" pageNumber="609" type="nomenclature">
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<paragraph pageId="0" pageNumber="609">
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<taxonomicName LSID="FB800B12-3BA4-5B5F-AB22-39F345F20679" authority="(Eichhoff, 1878)" baseAuthorityName="Eichhoff" baseAuthorityYear="1878" class="Insecta" family="Curculionidae" genus="Coptoborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoborus cuneatus" order="Coleoptera" pageId="0" pageNumber="609" phylum="Arthropoda" rank="species" species="cuneatus">Coptoborus cuneatus (Eichhoff, 1878)</taxonomicName>
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal, lateral, frontal and declivital view of Coptoborus crassisororcula holotype, 1.8 - 1.9 mm (A-C, M), C. crinitulus holotype, 1.9 - 2.5 mm (D-F, N), C. cuneatus syntype, 2.1 mm (G-I, O), C. doliolum holotype, 1.7 mm (J-L, P). All photographs by SMS, D-F, N copyright National Museum of Natural History, Smithsonian Institution, Washington, D. C., published by permission." figureDoi="10.3897/zookeys.144.62246.figure5" httpUri="https://binary.pensoft.net/fig/555840" pageId="0" pageNumber="609">Figure 5G-I, O</figureCitation>
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</paragraph>
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="609" type="reference_group">
|
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<paragraph pageId="0" pageNumber="609">
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||||
<taxonomicName authorityName="Eichhoff" authorityYear="1878" class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus cuneatus" order="Coleoptera" pageId="0" pageNumber="609" phylum="Arthropoda" rank="species" species="cuneatus">Xyleborus cuneatus</taxonomicName>
|
||||
Eichhoff, 1878: 380.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="609">
|
||||
<taxonomicName baseAuthorityName="Eichhoff" baseAuthorityYear="1878" class="Insecta" family="Curculionidae" genus="Coptoborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoborus cuneatus" order="Coleoptera" pageId="0" pageNumber="609" phylum="Arthropoda" rank="species" species="cuneatus">Coptoborus cuneatus</taxonomicName>
|
||||
(Eichhoff):
|
||||
<bibRefCitation author="Wood, SL" journalOrPublisher="The Great Basin Naturalist Memoirs" pageId="0" pageNumber="609" pagination="1 - 1553" refId="B92" refString="Wood, SL, Bright, DE, 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), part 2: Taxonomic index. The Great Basin Naturalist Memoirs 13: 1 - 1553" title="A catalog of Scolytidae and Platypodidae (Coleoptera), part 2: Taxonomic index." volume="13" year="1992">Wood and Bright 1992</bibRefCitation>
|
||||
: 663.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="609" type="type material">
|
||||
<paragraph pageId="0" pageNumber="609">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="609">
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||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="609">Syntypes</emphasis>
|
||||
(NHMW, 1; examined) and (MIIZ, 1; not examined) (
|
||||
<bibRefCitation author="Wegrzynowicz, P" journalOrPublisher="Bulletin of the Museum and Institute of Zoology PAS" pageId="0" pageNumber="609" pagination="53 - 65" refId="B79" refString="Wegrzynowicz, P, Mokrzecki, T, 1996. The type material of families Scolytidae and Platypodidae (Coleoptera) in the Museum and Institute of Zoology, PAS, Warsaw. Bulletin of the Museum and Institute of Zoology PAS 1: 53 - 65" title="The type material of families Scolytidae and Platypodidae (Coleoptera) in the Museum and Institute of Zoology, PAS, Warsaw." volume="1" year="1996">
|
||||
<normalizedToken originalValue="Węgrzynowicz">Wegrzynowicz</normalizedToken>
|
||||
and Mokrzecki 1996
|
||||
</bibRefCitation>
|
||||
). See remarks.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="609" type="diagnosis">
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||||
<paragraph pageId="0" pageNumber="609">Diagnosis.</paragraph>
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||||
<paragraph pageId="0" pageNumber="609">
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2.1 mm (n = 1), 3.5
|
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<normalizedToken originalValue="×">x</normalizedToken>
|
||||
as long as wide. This species is distinguished by the elytral apex attenuate and weakly emarginate, declivital interstriae 2 convex, declivital interstriae 1-3 unarmed, elytral apex deeply emarginate, and declivity steep, occupying less than posterior 40% of declivity.
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="609" type="similar species">
|
||||
<paragraph pageId="0" pageNumber="609">Similar species.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="609">
|
||||
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. exutus" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="exutus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="609">C. exutus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName family="Combretaceae" kingdom="Animalia" lsidName="C. galacatosae" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="galacatosae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="609">C. galacatosae</emphasis>
|
||||
</taxonomicName>
|
||||
.
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="609" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="609">Distribution.</paragraph>
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||||
<paragraph pageId="0" pageNumber="609">
|
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Panama (
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<normalizedToken originalValue="Panamá">Panama</normalizedToken>
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), Peru (
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<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
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), Venezuela (Barinas).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="609" type="biology_ecology">
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<paragraph pageId="0" pageNumber="609">Biology.</paragraph>
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||||
<paragraph pageId="0" pageNumber="609">Unknown.</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="609" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="609">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="609">
|
||||
<bibRefCitation author="Eichhoff, WJ" journalOrPublisher="Arbeiten ueber Morphologische und Taxonomische Entomologie aus Berlin-Dahlem" pageId="0" pageNumber="609" refId="B31" refString="Eichhoff, WJ, 1878. Ratio, descriptio, emendatio eorum Tomicinorum qui sunt in Dr medic. Chapuisii et autoris ipsius collectionibus et quos praeterea recognovit scriptor. Memoires de la Societe Royale des Sciences de Liege, Serie 2e, 8: 1-531. [pls. I-V]" title="Ratio, descriptio, emendatio eorum Tomicinorum qui sunt in Dr medic. Chapuisii et autoris ipsius collectionibus et quos praeterea recognovit scriptor. Memoires de la Societe Royale des Sciences de Liege, Serie 2 e, 8: 1 - 531. [pls. I-V]" year="1878">Eichhoff (1878)</bibRefCitation>
|
||||
described the species from an unspecified number of specimens.
|
||||
<bibRefCitation author="Schedl, KE" journalOrPublisher="Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien, Entomologie, 3 (2)" pageId="0" pageNumber="609" refId="B69" refString="Schedl, KE, 1979. Die Typen der Sammlung Schedl. Familie Scolytidae (Coleoptera). Mit einer Würdigung des Autors von Max Fischer und Schriftenverzeichnis. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien, Entomologie, 3(2)" title="Die Typen der Sammlung Schedl. Familie Scolytidae (Coleoptera). Mit einer Wu ̈ rdigung des Autors von Max Fischer und Schriftenverzeichnis." year="1979">Schedl (1979)</bibRefCitation>
|
||||
apparently presumed that the specimen of
|
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<taxonomicName family="Curculionidae" kingdom="Animalia" lsidName="X. cuneatus" order="Coleoptera" pageId="0" pageNumber="609" rank="species" species="cuneatus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="609">X. cuneatus</emphasis>
|
||||
</taxonomicName>
|
||||
in his collection to be a holotype under the assumption that the species was described from a single specimen. This is not the case, as an additional syntype is in MIIZ along with several other Eichhoff types (
|
||||
<bibRefCitation author="Wegrzynowicz, P" journalOrPublisher="Bulletin of the Museum and Institute of Zoology PAS" pageId="0" pageNumber="609" pagination="53 - 65" refId="B79" refString="Wegrzynowicz, P, Mokrzecki, T, 1996. The type material of families Scolytidae and Platypodidae (Coleoptera) in the Museum and Institute of Zoology, PAS, Warsaw. Bulletin of the Museum and Institute of Zoology PAS 1: 53 - 65" title="The type material of families Scolytidae and Platypodidae (Coleoptera) in the Museum and Institute of Zoology, PAS, Warsaw." volume="1" year="1996">
|
||||
<normalizedToken originalValue="Węgrzynowicz">Wegrzynowicz</normalizedToken>
|
||||
and Mokrzecki 1996
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation DOI="https://doi.org/10.1093/jee/toz026" author="Cognato, AI" journalOrPublisher="Journal of Economic Entomology" pageId="0" pageNumber="609" pagination="1274 - 1284" refId="B19" refString="Cognato, AI, Smith, SM, Li, Y, Pham, TH, Hulcr, J, 2019. Genetic variability among native Xyleborus glabratus Eichhoff populations native to Southeast Asia and the description of two related species. Journal of Economic Entomology 112: 1274 - 1284, DOI: https://doi.org/10.1093/jee/toz026" title="Genetic variability among native Xyleborus glabratus Eichhoff populations native to Southeast Asia and the description of two related species." url="https://doi.org/10.1093/jee/toz026" volume="112" year="2019">Cognato et al. 2019</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation DOI="https://doi.org/10.3390/insects10090261" author="Smith, SM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="609" refId="B75" refString="Smith, SM, Gomez, DF, Beaver, RA, Hulcr, J, Cognato, AI, 2019b. Reassessment of the species in the Euwallacea fornicatus (Coleoptera: Curculionidae: Scolytinae) complex after the rediscovery of the 'lost' type specimen. Insects 10: e261. https://doi.org/10.3390/insects10090261" title="Reassessment of the species in the Euwallacea fornicatus (Coleoptera: Curculionidae: Scolytinae) complex after the rediscovery of the ' lost' type specimen. Insects 10: e 261." url="https://doi.org/10.3390/insects10090261" year="2019 b">Smith et al. 2019b</bibRefCitation>
|
||||
). Both specimens are therefore syntypes.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="609">
|
||||
The species was described from specimens collected in Varinas [sic, Barinas], Nova Grenada. The species was listed from Colombia (
|
||||
<bibRefCitation author="Wood, SL" journalOrPublisher="The Great Basin Naturalist Memoirs" pageId="0" pageNumber="609" pagination="1 - 1553" refId="B92" refString="Wood, SL, Bright, DE, 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), part 2: Taxonomic index. The Great Basin Naturalist Memoirs 13: 1 - 1553" title="A catalog of Scolytidae and Platypodidae (Coleoptera), part 2: Taxonomic index." volume="13" year="1992">Wood and Bright 1992</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Wood, SL" journalOrPublisher="Brigham Young University, M. L. Bean Life Science Museum; Provo" pageId="0" pageNumber="609" refId="B91" refString="Wood, SL, 2007. Bark and ambrosia beetles of South America (Coleoptera: Scolytidae). Brigham Young University, M.L. Bean Life Science Museum; Provo" title="Bark and ambrosia beetles of South America (Coleoptera: Scolytidae)." year="2007">Wood 2007</bibRefCitation>
|
||||
) but we were unable to find specimens definitively from this location (
|
||||
<bibRefCitation author="Wood, SL" journalOrPublisher="Brigham Young University, M. L. Bean Life Science Museum; Provo" pageId="0" pageNumber="609" refId="B91" refString="Wood, SL, 2007. Bark and ambrosia beetles of South America (Coleoptera: Scolytidae). Brigham Young University, M.L. Bean Life Science Museum; Provo" title="Bark and ambrosia beetles of South America (Coleoptera: Scolytidae)." year="2007">Wood 2007</bibRefCitation>
|
||||
). The type specimens were likely collected from Barinas, Venezuela.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
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||||
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data/FB/80/B6/FB80B6B01ADB502A8B2FB84ECB88CDE0.xml
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Branch, William R.</mods:namePart>
|
||||
<mods:affiliation>Port Elizabeth Museum, P. O. Box 13147, Humewood, 6013, Port Elizabeth, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela University, Port Elizabeth 6031, South Africa</mods:affiliation>
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<mods:namePart>Schmitz, Andreas</mods:namePart>
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<mods:affiliation>Natural History Museum of Geneva, UREC - Herpetology and Ichthyology, C. P. 6434, 1211 Geneva 6, Switzerland</mods:affiliation>
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<mods:namePart>Lobon-Rovira, Javier</mods:namePart>
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<mods:affiliation>CIBIO Centro de Investigacao em Biodiversidade e Recursos Geneticos, Universidade do Porto, Rua Padre Armando Quintas, Campus de Vairao, 4485 - 661 Vairao, Portugal & Faculdade de Ciencias, Universidade do Porto, 4169 - 007 Porto, Portugal</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Baptista, Ninda L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2859-6606</mods:nameIdentifier>
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<mods:affiliation>CIBIO Centro de Investigacao em Biodiversidade e Recursos Geneticos, Universidade do Porto, Rua Padre Armando Quintas, Campus de Vairao, 4485 - 661 Vairao, Portugal & Faculdade de Ciencias, Universidade do Porto, 4169 - 007 Porto, Portugal & Instituto Superior de Ciencias da Educacao da Huila (ISCED), Rua Sarmento Rodrigues s / n, Lubango, Angola</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Antonio, Telmo</mods:namePart>
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<mods:affiliation>Instituto Superior de Ciencias da Educacao da Huila (ISCED), Rua Sarmento Rodrigues s / n, Lubango, Angola & Faculty of Natural Resources and Spatial Sciences, Namibia University of Science and Technology, Private Bag 13388, Windhoek, Namibia</mods:affiliation>
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</mods:role>
|
||||
<mods:namePart>Conradie, Werner</mods:namePart>
|
||||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0805-9683</mods:nameIdentifier>
|
||||
<mods:affiliation>Port Elizabeth Museum, P. O. Box 13147, Humewood, 6013, Port Elizabeth, South Africa & School of Natural Resource Management, George Campus, Nelson Mandela University, George 6530, South Africa</mods:affiliation>
|
||||
<mods:nameIdentifier type="email">werner@bayworld.co.za</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Zoosystematics and Evolution</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2021</mods:date>
|
||||
<mods:detail type="pubDate">
|
||||
<mods:number>2021-01-12</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>97</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="issue">
|
||||
<mods:number>1</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>55</mods:start>
|
||||
<mods:end>82</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://dx.doi.org/10.3897/zse.97.57202</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.57202</mods:identifier>
|
||||
<mods:identifier type="Pensoft-Pub">1860-0743-1-55</mods:identifier>
|
||||
<mods:identifier type="ZooBank">A125EC81050E449A890427674F5265EF</mods:identifier>
|
||||
<mods:identifier type="Pensoft-UUID">8E2883E1FFEE5B6884EADB662AB420A1</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment ID-GBIF-Taxon="176584339" LSID="urn:lsid:zoobank.org:act:681544BA-AA5F-473B-90BD-B2790838BC81" httpUri="http://treatment.plazi.org/id/FB80B6B01ADB502A8B2FB84ECB88CDE0" lastPageNumber="55" pageId="0" pageNumber="55">
|
||||
<subSubSection pageId="0" pageNumber="55" type="nomenclature">
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<taxonomicName LSID="http://zoobank.org/681544BA-AA5F-473B-90BD-B2790838BC81" authority="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie, 2021" authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="wulfhaackei" status="sp. nov.">Afroedura wulfhaackei</taxonomicName>
|
||||
<taxonomicNameLabel pageId="0" pageNumber="55">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Live photos of Angolan flat geckos: A. Afroedura bogerti (PEM R 24187); B. Afroedura wulfhaackei sp. nov. (paratype PEM R 24232); C. Afroedura praedicta sp. nov. (holotype NB 854); D. Afroedura donveae sp. nov. (holotype PEM R 17937); E. Afroedura vazpintorum sp. nov. (holotype PEM R 24118); F. Afroedura vazpintorum sp. nov. (NB 0745); Photos: A, B, D, E. William R. Branch; C, F. Pedro Vaz Pinto." figureDoi="10.3897/zse.97.57202.figure5" httpUri="https://binary.pensoft.net/fig/496674" pageId="0" pageNumber="55">Angolan Flat Gecko Osga-achatada de Angola Figures 5B</figureCitation>
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Afroedura wulfhaackei sp. nov. (PEM R 24234) from Maka-Mombolo, Benguela Province, Angola. Photos: Werner Conradie." figureDoi="10.3897/zse.97.57202.figure7" httpUri="https://binary.pensoft.net/fig/496676" pageId="0" pageNumber="55">, 7</figureCitation>
|
||||
;
|
||||
<figureCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Summary of morphological data for Afroedura bogerti, A. wulfhaackei sp. nov. (including the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades), A. donveae sp. nov., A. vazpintorum sp. nov. and A. praedicta sp. nov. Values are given as a range with mean values in parenthesis. M = male, F = female, n = sample size." pageId="0" pageNumber="55">Tables 3</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements (in mm) and scale counts for the type series of Afroedura wulfhaackei sp. nov." pageId="0" pageNumber="55">5</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="notes">
|
||||
<paragraph pageId="0" pageNumber="55">Note.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
The phylogenetic analysis identified four well-defined clades (
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 4-7) within the southern inland/highland major clade (see Results). Our data show that this southern major clade clearly represents at least one undescribed species of
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
. Despite comparatively-low genetic distances between the four clades recovered (Table
|
||||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of intra- and interclade uncorrected pairwise genetic distances for specimens of Afroedura clades compared to A. loveridgei for 16 S rRNA. Intraclade / intraspecific distances are marked in bold." httpUri="http://table.plazi.org/id/3F87ADCE25A20DA61B3A1568F02AD894" pageId="0" pageNumber="55" tableUuid="3F87ADCE25A20DA61B3A1568F02AD894">2</tableCitation>
|
||||
), they were all retrieved separately in our phylogenetic analyses and, as our morphological examination failed to separate the four clades, we cannot assign them as separate species. Therefore, we decided to take a conservative approach and to describe only one of the clades as a new species for now and tentatively assign
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 5-7 to this species pending further work. Thus, we grouped the material according to these clades and selected the clade called here
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 4 (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic tree topology for the 16 S gene (identical for Bayes and ML) using Afroedura loveridgei as outgroup. Support values for posterior probability and bootstraps are indicated by circles (ML:> 70 % / PP:> 0.95)." figureDoi="10.3897/zse.97.57202.figure2" httpUri="https://binary.pensoft.net/fig/496671" pageId="0" pageNumber="55">2</figureCitation>
|
||||
) as the name-bearing clade for our new species. Material from lower-altitude localities near Bocoio in Benguela Province groups morphologically with our new species, but needs to be tested in a phylogenetic framework.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="synonym">
|
||||
<paragraph pageId="0" pageNumber="55">Synonym.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
-
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/zse.93.10915" author="Branch, WR" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="55" pagination="157 - 166" refId="B13" refString="Branch, WR, Haacke, W, Vaz Pinto, P, Conradie, W, Baptista, N, Verburgt, L, Verissimo, L, 2017b. Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status. Zoosystematics and Evolution 93 (1): 157 - 166, DOI: https://doi.org/10.3897/zse.93.10915" title="Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status." url="https://doi.org/10.3897/zse.93.10915" volume="93" year="2017 b">Branch et al. 2017b</bibRefCitation>
|
||||
:157 (part);
|
||||
<bibRefCitation author="Marques, MP" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" refId="B49" refString="Marques, MP, Ceriaco, LMP, Blackburn, DC, Bauer, AM, 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola. Atlas of Historical and Bibliographic Records (1840-2017). Proceedings of the California Academy of Science, Series 4 65(Supplement II): 1-501." title="Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola. Atlas of Historical and Bibliographic Records (1840 - 2017). Proceedings of the California Academy of Science, Series 4 65 (Supplement II): 1 - 501." year="2018">Marques et al. 2018</bibRefCitation>
|
||||
: 177 (part);
|
||||
<bibRefCitation DOI="https://doi.org/10.1007/978-3-030-03083-4_13" author="Branch, WR" editor="Huntley, BJ" journalOrPublisher="Science and Conservation: A Modern Synthesis. Springer Open, Cham" pageId="0" pageNumber="55" pagination="283 - 326" refId="B9" refString="Branch, WR, Baptista, N, Vaz Pinto, P, Conradie, W, 2019a. The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research. In: Huntley, BJ, Ferrand, N, Eds., Biodiversity of Angola. Science and Conservation: A Modern Synthesis. Springer Open, Cham: 283 - 326, DOI: https://doi.org/10.1007/978-3-030-03083-4_13" title="The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research." url="https://doi.org/10.1007/978-3-030-03083-4_13" volumeTitle="Biodiversity of Angola." year="2019 a">Branch et al. 2019a</bibRefCitation>
|
||||
: 287 (part);
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
(clade 4) -
|
||||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4324.1.8" author="Branch, WR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="142 - 160" refId="B12" refString="Branch, WR, Guyton, JA, Schmitz, A, Barej, MF, Naskrecki, P, Farooq, H, Verburgt, L, Roedel, M-O, 2017a. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142 - 160, DOI: https://doi.org/10.11646/zootaxa.4324.1.8" title="Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique." url="https://doi.org/10.11646/zootaxa.4324.1.8" volume="4324" year="2017 a">Branch et al. 2017a</bibRefCitation>
|
||||
:147.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="holotype">
|
||||
<materialsCitation collectingDate="1944-01-01" collectingDateMax="1944-12-31" collectingDateMin="1944-01-01" collectionCode="PEM" collectorName="William R. Branch & Pedro Vaz Pinto" country="Angola" elevation="0" latitude="-12.17056" location="Maka-Mombolo" longLatPrecision="1" longitude="14.88167" specimenCode="PEM R24234" specimenCount="1" stateProvince="Benguela Province" typeStatus="holotype">
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<typeStatus>Holotype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<specimenCode>PEM R24234</specimenCode>
|
||||
,
|
||||
<specimenCount type="adult">adult male</specimenCount>
|
||||
, collected
|
||||
<locationDeviation>3 km north</locationDeviation>
|
||||
of
|
||||
<location LSID="urn:lsid:plazi:treatment:FB80B6B01ADB502A8B2FB84ECB88CDE0:A49F307E44A4A9D6E452283B24042C3F" country="Angola" latitude="-12.17056" longLatPrecision="1" longitude="14.88167" name="Maka-Mombolo" stateProvince="Benguela Province">Maka-Mombolo</location>
|
||||
(
|
||||
<geoCoordinate degrees="12.17056" direction="south" orientation="latitude" precision="1" value="-12.17056">-12.17056</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="14.88167" direction="east" orientation="longitude" precision="1" value="14.88167">14.88167</geoCoordinate>
|
||||
,
|
||||
<elevation>1756 m</elevation>
|
||||
a.s.l.),
|
||||
<collectingRegion>Benguela Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry>Angola</collectingCountry>
|
||||
, by
|
||||
<collectorName>William R. Branch</collectorName>
|
||||
and
|
||||
<collectorName>Pedro Vaz Pinto</collectorName>
|
||||
on
|
||||
<collectingDate>15 November 2016</collectingDate>
|
||||
.
|
||||
</paragraph>
|
||||
</materialsCitation>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="paratypes">
|
||||
<materialsCitation collectingDate="1944-01-01" collectingDateMax="1944-12-31" collectingDateMin="1944-01-01" collectionCode="PEM" collectorName="William R. Branch & Pedro Vaz Pinto" country="Angola" elevation="0" latitude="-12.17056" location="Maka-Mombolo" longLatPrecision="1" longitude="14.88167" specimenCode="PEM R24232-3" specimenCount="1" stateProvince="Benguela Province" typeStatus="paratype">
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<typeStatus>Paratypes</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<specimenCode>PEM R24232-3</specimenCode>
|
||||
,
|
||||
<specimenCount type="adult">adult females</specimenCount>
|
||||
, collected
|
||||
<locationDeviation>3 km north</locationDeviation>
|
||||
of
|
||||
<location LSID="urn:lsid:plazi:treatment:FB80B6B01ADB502A8B2FB84ECB88CDE0:217E3B2B28CBABB33505FFC92C4BB1E6" country="Angola" latitude="-12.17056" longLatPrecision="1" longitude="14.88167" name="Maka-Mombolo" stateProvince="Benguela Province">Maka-Mombolo</location>
|
||||
(
|
||||
<geoCoordinate degrees="12.17056" direction="south" orientation="latitude" precision="1" value="-12.17056">-12.17056</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="14.88167" direction="east" orientation="longitude" precision="1" value="14.88167">14.88167</geoCoordinate>
|
||||
,
|
||||
<elevation>1756 m</elevation>
|
||||
a.s.l),
|
||||
<collectingRegion>Benguela Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry>Angola</collectingCountry>
|
||||
, by
|
||||
<collectorName>William R. Branch</collectorName>
|
||||
and
|
||||
<collectorName>Pedro Vaz Pinto</collectorName>
|
||||
on
|
||||
<collectingDate>15 November 2016</collectingDate>
|
||||
;
|
||||
<materialsCitation collectingDate="1944-01-01" collectingDateMax="1944-12-31" collectingDateMin="1944-01-01" collectionCode="PEM" collectorName="William R. Branch & Pedro Vaz Pinto" elevation="0" latitude="-12.19833" location="Maka-Mombolo" longLatPrecision="1" longitude="14.86833" specimenCode="PEM R24236" specimenCount="1" stateProvince="Benguela Province" typeStatus="paratype">
|
||||
<specimenCode>PEM R24236</specimenCode>
|
||||
,
|
||||
<specimenCount type="adult">adult male</specimenCount>
|
||||
, collected above
|
||||
<location LSID="urn:lsid:plazi:treatment:FB80B6B01ADB502A8B2FB84ECB88CDE0:3B3E0EC6BB5C8A70E906F23B4B9A765F" latitude="-12.19833" longLatPrecision="1" longitude="14.86833" name="Maka-Mombolo" stateProvince="Benguela Province">Maka-Mombolo</location>
|
||||
,
|
||||
<locationDeviation>north-east of Balombo</locationDeviation>
|
||||
(
|
||||
<geoCoordinate degrees="12.19833" direction="south" orientation="latitude" precision="1" value="-12.19833">-12.19833</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="14.86833" direction="east" orientation="longitude" precision="1" value="14.86833">14.86833</geoCoordinate>
|
||||
,
|
||||
<elevation>1857 m</elevation>
|
||||
a.s.l.),
|
||||
<collectingRegion>Benguela Province</collectingRegion>
|
||||
, Angola, by
|
||||
<collectorName>William R. Branch</collectorName>
|
||||
and
|
||||
<collectorName>Pedro Vaz Pinto</collectorName>
|
||||
on
|
||||
<collectingDate>15 November 2016</collectingDate>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</materialsCitation>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="55">Additional referred material</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<emphasis bold="true" pageId="0" pageNumber="55">(not examined).</emphasis>
|
||||
NB 817-9, collected at Morro do Moco, camp near Canjonde (
|
||||
<geoCoordinate degrees="12.42611" direction="south" orientation="latitude" precision="1" value="-12.42611">-12.42611</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.14778" direction="east" orientation="longitude" precision="1" value="15.14778">15.14778</geoCoordinate>
|
||||
, 1931 m a.s.l), Huambo Province, Angola, by Pedro Vaz Pinto on 13 November 2017 (genetic samples included in this study).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="55">Tentative referred additional material examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 5 (
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Males</emphasis>
|
||||
): PEM R24192-4, collected at William
|
||||
<normalizedToken originalValue="Chapman’s">Chapman's</normalizedToken>
|
||||
Farm Victoria-Verdun (Sandula), (
|
||||
<geoCoordinate degrees="12.17194" direction="south" orientation="latitude" precision="1" value="-12.17194">-12.17194</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.02667" direction="east" orientation="longitude" precision="1" value="15.02667">15.02667</geoCoordinate>
|
||||
, 1834 m a.s.l.), Cuanza-Sul Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 6 November 2016.
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 5 (
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Females</emphasis>
|
||||
): PEM R24190-1, PEM R24195-6, PEM R24199, collected at William
|
||||
<normalizedToken originalValue="Chapman’s">Chapman's</normalizedToken>
|
||||
Farm Victoria-Verdun (Sandula), (
|
||||
<geoCoordinate degrees="12.17194" direction="south" orientation="latitude" precision="1" value="-12.17194">-12.17194</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.02667" direction="east" orientation="longitude" precision="1" value="15.02667">15.02667</geoCoordinate>
|
||||
, 1834 m a.s.l.), Cuanza-Sul Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 6 November 2016.
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 5 (
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Juveniles</emphasis>
|
||||
): PEM R24197-8, collected at William
|
||||
<normalizedToken originalValue="Chapman’s">Chapman's</normalizedToken>
|
||||
Farm Victoria-Verdun (Sandula), (
|
||||
<geoCoordinate degrees="12.17194" direction="south" orientation="latitude" precision="1" value="-12.17194">-12.17194</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.02667" direction="east" orientation="longitude" precision="1" value="15.02667">15.02667</geoCoordinate>
|
||||
, 1834 m a.s.l.), Cuanza-Sul Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 6 November 2016.
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 6 (
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">
|
||||
<normalizedToken originalValue="Мales">Males</normalizedToken>
|
||||
</emphasis>
|
||||
): PEM R24201, collected near Rio Chicanda, 5 km southwest of Lepi (
|
||||
<geoCoordinate degrees="12.90861" direction="south" orientation="latitude" precision="1" value="-12.90861">-12.90861</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.36472" direction="east" orientation="longitude" precision="1" value="15.36472">15.36472</geoCoordinate>
|
||||
, 1534 m a.s.l.), Huambo Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 8 November 2016.
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 7 (
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">
|
||||
<normalizedToken originalValue="Мales">Males</normalizedToken>
|
||||
</emphasis>
|
||||
): TM 45382, TM 45387-8, TM 45392, TM 45396, collected at Candumbo Rocks, 16 km west of Vila Nova = Huambo (
|
||||
<geoCoordinate degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
|
||||
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971.
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 7 (
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Females</emphasis>
|
||||
): PEM R22490-1, collected 1 km west of Candumbo on road to Boas
|
||||
<normalizedToken originalValue="Águas">Aguas</normalizedToken>
|
||||
(
|
||||
<geoCoordinate degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
|
||||
, 1760 m a.s.l), Huambo Province, Angola, by Luke Verburgt on 11 March 2016; TM 45383, TM 45390, TM 45393-4, TM 45397, collected at Candumbo Rocks, 16 km west of Vila Nova = Huambo (
|
||||
<geoCoordinate degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
|
||||
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971.
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 7 (
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Juveniles</emphasis>
|
||||
): PEM R24200, collected at Candumbo Rocks Memorial, 20 km east Humana to Cuito (
|
||||
<geoCoordinate degrees="12.73722" direction="south" orientation="latitude" precision="1" value="-12.73722">-12.73722</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.97333" direction="east" orientation="longitude" precision="1" value="15.97333">15.97333</geoCoordinate>
|
||||
, 1750 m a.s.l), Huambo Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 7 November 2016; TM 45386, TM 45389, TM 45391, TM 45395, collected at Candumbo Rocks, 16 km west of Vila Nova = Huambo (
|
||||
<geoCoordinate degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
|
||||
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971.
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Unassigned clades></emphasis>
|
||||
(
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Females</emphasis>
|
||||
): TM 45374, collected 1 km south of Luimbale (
|
||||
<geoCoordinate degrees="12.25367" direction="south" orientation="latitude" precision="1" value="-12.25367">-12.25367</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.31694" direction="east" orientation="longitude" precision="1" value="15.31694">15.31694</geoCoordinate>
|
||||
, 1591 m a.s.l.), Huambo Province, Angola, collected by Wulf Haacke on 10 May 1971; TM 45367-8, collected 10 km west of Soque (
|
||||
<geoCoordinate degrees="12.34590" direction="south" orientation="latitude" precision="1" value="-12.3459">-12.34590</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.01180" direction="east" orientation="longitude" precision="1" value="15.0118">15.01180</geoCoordinate>
|
||||
, 1974 m a.s.l), Benguela Province, Angola, collected by Wulf Haacke on 10 May 1971; PEM R24743, collected at Morro do Pundo (
|
||||
<geoCoordinate degrees="12.44389" direction="south" orientation="latitude" precision="1" value="-12.44389">-12.44389</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="13.92250" direction="east" orientation="longitude" precision="1" value="13.9225">13.92250</geoCoordinate>
|
||||
, 939 m a.s.l.), Benguela Province, Angola, by Pedro Vaz Pinto on 6 June 2018; TM 46587-8, TM 465890, collected 3 km west of Bocoio (
|
||||
<geoCoordinate degrees="12.46605" direction="south" orientation="latitude" precision="1" value="-12.46605">-12.46605</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="14.10694" direction="east" orientation="longitude" precision="1" value="14.10694">14.10694</geoCoordinate>
|
||||
, 926 m a.s.l.), Benguela Province, Angola by Wulf Haacke on 25 May 1971.
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Unassigned clades</emphasis>
|
||||
(
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Males</emphasis>
|
||||
): TM 45366 collected 10 km west of Soque (
|
||||
<geoCoordinate degrees="12.34590" direction="south" orientation="latitude" precision="1" value="-12.3459">-12.34590</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.01180" direction="east" orientation="longitude" precision="1" value="15.0118">15.01180</geoCoordinate>
|
||||
, 1974 m a.s.l), Benguela Province, Angola, by Wulf Haacke on 10 May 1971, TM 46589, adult male, collected 3 km west of Bocoio (
|
||||
<geoCoordinate degrees="12.46605" direction="south" orientation="latitude" precision="1" value="-12.46605">-12.46605</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="14.10694" direction="east" orientation="longitude" precision="1" value="14.10694">14.10694</geoCoordinate>
|
||||
, 926 m a.s.l.), Benguela Province, Angola, by Wulf Haacke on 25 May 1971.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="55">Tentative referred additional material not examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
TM 45381, 45384-5, TM 45398, collected at Candumbo Rocks, about 25 km east of the city of Huambo (
|
||||
<geoCoordinate degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
|
||||
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971 (placed in
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 7, based on same geographical area as genetically-assigned material); FKH 0239, collected at Monte Verde-Sandula (
|
||||
<geoCoordinate degrees="12.17924" direction="south" orientation="latitude" precision="1" value="-12.17924">-12.17924</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="15.03086" direction="east" orientation="longitude" precision="1" value="15.03086">15.03086</geoCoordinate>
|
||||
, 2055 m a.s.l.), Cuanza-Sul Province, Angola, by Pedro Vaz Pinto on 29 May 2019 (genetic sample included in this study placed it in
|
||||
<taxonomicName authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
|
||||
</taxonomicName>
|
||||
sp. 5).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="55">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">The new species is named in honour of Wulf Haacke, retired curator of the herpetology collection at the former Transvaal Museum (now Ditsong National Museum of Natural History). His herpetological expeditions to Angola in the early 1970s paved the way for this study and much of the material used in this study resulted from his expeditions. The name is constructed in the masculine singular genitive.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="55">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
A member of the greater '
|
||||
<taxonomicName baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="transvaalica">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura transvaalica</emphasis>
|
||||
</taxonomicName>
|
||||
' group in possessing two pairs of enlarged scansors per digit and a strongly verticillate and flattened tail (
|
||||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3846.4.1" author="Jacobsen, NHG" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="451 - 501" refId="B41" refString="Jacobsen, NHG, Kuhn, AL, Jackman, TR, Bauer, AM, 2014. A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa. Zootaxa 3846 (4): 451 - 501, DOI: https://doi.org/10.11646/zootaxa.3846.4.1" title="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa." url="https://doi.org/10.11646/zootaxa.3846.4.1" volume="3846" year="2014">Jacobsen et al. 2014</bibRefCitation>
|
||||
). It is part of the
|
||||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
-group which differs from other members of the '
|
||||
<taxonomicName baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="transvaalica">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura transvaalica</emphasis>
|
||||
</taxonomicName>
|
||||
' group by having less than 88 mid-body scale rows (vs. 97-102 in
|
||||
<taxonomicName genus="A." lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
|
||||
</taxonomicName>
|
||||
, 113-120 in
|
||||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||||
</taxonomicName>
|
||||
, 102-119 in
|
||||
<taxonomicName genus="A." lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
|
||||
</taxonomicName>
|
||||
); by the rostral bordering the nostril (nostril excluded from rostral in
|
||||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||||
</taxonomicName>
|
||||
); by the anterior nasals being mostly in contact ~ 68% (separated by 1-3 granules in
|
||||
<taxonomicName genus="A." lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
|
||||
</taxonomicName>
|
||||
; always in broad contact in
|
||||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||||
</taxonomicName>
|
||||
; usually in broad contact in
|
||||
<taxonomicName genus="A." lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
|
||||
</taxonomicName>
|
||||
~ 3-18%); and in having 11-16 scales between the anterior borders of the eyes (19-22 in
|
||||
<taxonomicName genus="A." lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
|
||||
</taxonomicName>
|
||||
; 15-19 in
|
||||
<taxonomicName genus="A." lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
|
||||
</taxonomicName>
|
||||
; 15-20 in
|
||||
<taxonomicName genus="A." lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
|
||||
</taxonomicName>
|
||||
) (comparative data
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">fide</emphasis>
|
||||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4324.1.8" author="Branch, WR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="142 - 160" refId="B12" refString="Branch, WR, Guyton, JA, Schmitz, A, Barej, MF, Naskrecki, P, Farooq, H, Verburgt, L, Roedel, M-O, 2017a. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142 - 160, DOI: https://doi.org/10.11646/zootaxa.4324.1.8" title="Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique." url="https://doi.org/10.11646/zootaxa.4324.1.8" volume="4324" year="2017 a">Branch et al. 2017a</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="wulfhaackei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. differs from other members of the
|
||||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
-group by a combination of the following characters (see Tables
|
||||
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Summary of morphological data for Afroedura bogerti, A. wulfhaackei sp. nov. (including the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades), A. donveae sp. nov., A. vazpintorum sp. nov. and A. praedicta sp. nov. Values are given as a range with mean values in parenthesis. M = male, F = female, n = sample size." httpUri="http://table.plazi.org/id/742172291B68E9AFACEE2244550D3FEF" pageId="0" pageNumber="55" tableUuid="742172291B68E9AFACEE2244550D3FEF">3</tableCitation>
|
||||
,
|
||||
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Tukey multiple comparisons of means of the four species combinations of Angolan Afroedura (values for A. wulfhaackei sp. nov. include the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades). Bold values indicate significant pairwise comparisons. SVL - snout-vent length; HL - head length; HW - head width; ED - eye-diameter; EE - ear-eye length; IN - internostril distance; MSR - mid-body scale rows." httpUri="http://table.plazi.org/id/C30887A74A39988C01C45C883AEB5E57" pageId="0" pageNumber="55" tableUuid="C30887A74A39988C01C45C883AEB5E57">4</tableCitation>
|
||||
): 76-88 (mean 79.3) mid-body scale rows (69-77 [mean 73.5] in
|
||||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
, 64-78 [mean 72.8] in
|
||||
<taxonomicName genus="A." lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., 73-86 [mean 80.3] in
|
||||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., 73-78 [mean 74.8] in
|
||||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.); by the anterior nasals being mostly (~ 68% of the time) in contact (~ 33% of the time in contact in
|
||||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
; always in contact in
|
||||
<taxonomicName genus="A." lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.,
|
||||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. and
|
||||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.); each verticil comprising 4-5 (mean 4.0) ventral and 5-6 (mean 5.1) dorsal rows of scales (4 and 5 in
|
||||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.; 5-6 [mean 5.5] and 6-7 [mean 6.6] in
|
||||
<taxonomicName genus="A." lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.; 5-6 [mean 5.0] and 6-7 [mean 6.1]
|
||||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.); ventral surfaces greyish with scattered small black spots (similar to
|
||||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., immaculate in
|
||||
<taxonomicName genus="A." lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. and
|
||||
<taxonomicName genus="A." lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.).
|
||||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="wulfhaackei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. differs more specifically from its sister highland species
|
||||
<taxonomicName genus="A." lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
|
||||
</taxonomicName>
|
||||
in having a higher number of mid-body scale counts (76-88 [mean 79.3] versus 69-77 [mean 73.5]) and differs from
|
||||
<taxonomicName genus="A." lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. in that the nasals are separated by smaller granules (versus always in contact).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="description">
|
||||
<paragraph pageId="0" pageNumber="55">Holotype description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
Adult male; SVL 51.4 mm; tail 47.1 mm (full original tail), with a small mid-ventral incision for the removal of liver sample. Measurements and meristic characters of holotype presented in Table
|
||||
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements (in mm) and scale counts for the type series of Afroedura wulfhaackei sp. nov." httpUri="http://table.plazi.org/id/8580963F746364997A5BD1D5F11DD033" pageId="0" pageNumber="55" tableUuid="8580963F746364997A5BD1D5F11DD033">5</tableCitation>
|
||||
. Head and body dorsoventrally compressed; HL 11.9 mm, HW 9.0 mm, broadest at posterior level of eye and 1.32 times longer than wide. Eye large (2.3 mm wide), pupil vertical with indented margins; circumorbital scales small and smooth, elongate at upper anterior margin, upper three posterior scales with small upward pointing spines. Snout rounded, 4.9 mm long, slightly longer than distance between eye and ear openings (4.0 mm). Scales on top of snout smooth, rounded, scales to the edge larger than central ones, with no intervening minute granules. Scales on snout slightly subequal in size to those on the back of head or the nape. Scales on eyelids larger than those on the crown, five scales deep from circumorbital scale to crown. Circumorbital scales are separated by a row of smaller scales from the larger scales on eyelids. Nostril pierced between rostral and three nasal scales; 1st supralabial narrowly excluded from nostril; the supranasal being much larger than the subequal postnasals and separated from each other by two smaller scales. Nostrils slightly elevated. Rostral roughly rectangular, but with its upper edges elongated due to extensions into the nostril. Eight supralabials on each side, the labial margin flexing upwards at the rictus (approx. mid-orbital position), with 3-4 minute scales proximal to the flexure. Ten infralabials on either side, with a small scale proximal to the flexure. At the lip, mental slightly narrower than adjacent infralabial, only three quarters the width of rostral and in contact with two distinctly elongate postmental scales. Scales on throat much smaller than those on belly; scales touching infralabials larger. Fifteen scales across the crown at level of front of eyes; 16 scales from ear to eye; 83 scales around mid-body. Ear opening deep, oblique and roughly oval, only half as high as wide (0.3
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
0.5 mm). Scales on dorsum smooth, non-overlapping, largest at mid-body, smaller on nape and tail base. Scales on ventrum flattened, not overlapping, more-or-less ovate at mid-ventrum, twice the size of lateral granules and 1.5 times those along the backbone. Original tail slightly dorsoventrally flattened and distinctly verticillate (11 verticils in total), with obvious lateral constrictions that are not that distinct to tip of tail; each verticil comprising 5 imbricate rows of scales dorsally and 4 imbricate scale rows ventrally and with ventral scales approximately twice the size of those on the dorsal surface. Limbs well developed, hindlimbs slightly longer than forelimbs, mite pockets (dermal crevices inhabited by small ectoparasitic mites) only present at anterior margin of hindlimbs. All digits with a large pair of distal scansors, separated by a large, curved claw and followed by a large gap (twice the length of terminal scansor) by a smaller pair of scansors; infero-median row of digital scales enlarged transversely, particularly towards the scansors, where the terminal scale adjoining the first pair of scansors may be medially constricted, swollen and scansor-like; enlarged subdigital lamellae on 4th toe 9. Precloacal pores 9.
|
||||
</paragraph>
|
||||
<caption ID-Table-UUID="8580963F746364997A5BD1D5F11DD033" httpUri="http://table.plazi.org/id/8580963F746364997A5BD1D5F11DD033" pageId="0" pageNumber="55" start="Table 5" startId="T5">
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<emphasis bold="true" pageId="0" pageNumber="55">Table 5.</emphasis>
|
||||
Measurements (in mm) and scale counts for the type series of
|
||||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="wulfhaackei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<table pageId="0" pageNumber="55">
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<th colspan="1" pageId="0" pageNumber="55" rowspan="1">Catalogue number</th>
|
||||
<th colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R24234</th>
|
||||
<th colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R24232</th>
|
||||
<th colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R24233</th>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Type Status</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Holotype</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Paratype</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Paratype</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Sex</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Male</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Female</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Female</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Snout vent length</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">51.4</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">49.1</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">55.7</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Tail length</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">47.1</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">35.4</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">-57.5</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Tail condition</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Original</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Regenerated</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Regenerated</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Head length</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">11.9</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">10.9</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">12.3</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Head width</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">9.0</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">7.5</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">8.5</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Snout length</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4.9</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4.5</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">5.1</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Eye distance</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">2.3</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">2.2</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">2.6</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Eye-Ear distance</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4.0</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">3.5</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4.3</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Precloacal pores (males)</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Dorsal rows per tail verticil</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">5</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">5</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Ventral rows per tail verticil</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">4</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales below 4th toe</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Mid-body scale rows</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">83</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">82</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">76</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales between eyes</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">15</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">13</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">14</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales: nostril to eye</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales: ear to eye</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">16</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">14</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">17</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Anterior nasals in contact</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">No</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">No</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">No</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Supralabials</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="55">
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">Infralabials</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
|
||||
<td colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="paratype variation">
|
||||
<paragraph pageId="0" pageNumber="55">Paratype variation</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<emphasis bold="true" pageId="0" pageNumber="55">
|
||||
(see Table
|
||||
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements (in mm) and scale counts for the type series of Afroedura wulfhaackei sp. nov." httpUri="http://table.plazi.org/id/8580963F746364997A5BD1D5F11DD033" pageId="0" pageNumber="55" tableUuid="8580963F746364997A5BD1D5F11DD033">5</tableCitation>
|
||||
for more measurements and scale counts of type series).
|
||||
</emphasis>
|
||||
SVL varied from 49.1-55.7 mm; head length 1.34-1.45 times the head width; snout 1.99 times the diameter of eye. Supranasals always separated by smaller granules, usually with a single large granule in contact with the rostral between the supranasals, followed by 1-2 smaller granules in lateral contact; the first upper labial and rostral always enters the nostril and the width of the rostral at the lip margin is always wider than that of the mental; 2-3 postmental scales; supralabials 7-9, infralabials 8-9; scales between anterior edges of eyes 13-14; scales between nostril and anterior edge of orbit 7-9; scales between anterior edge of ear and rear margin of orbit 14-17; scales around mid-body 76-82; subdigital lamellae on 4th toe 7-8; dorsal scales per tail verticil 5; ventral scales per tail verticil 4. Precloacal pores 9.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="55">Additional material variation.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">SVL 36.4-59.6 mm; original tail length 36.4-57.8 mm, 0.94 times SVL; head length 1.19-1.75 times head width; snout 1.91 times diameter of eye. The supranasals in contact in 22 specimens and separated by granules in eight specimens, usually with a single large granule in contact with the rostral between the supranasals, followed by 1-2 smaller granules in lateral contact; the first upper labial and rostral always enters the nostril and the width of the rostral at the lip margin is always wider than that of the mental; 2-4 postmental scales; supralabials 8-9; infralabials 8-9; scales between anterior edge of eye 12-16; scales between nostril and anterior edge of orbit 7-11; scales between anterior edge of ear and rear margin of orbit 15-18; scales around mid-body 75-88; subdigital lamellae on 4th toe 7-9; dorsal scales per tail verticil 5 (TM 45366, TM 46588 and PEM R24743 with 6); ventral scales per tail verticil 4 (TM 46588 with 5). Precloacal pores 9-12.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="colouration">
|
||||
<paragraph pageId="0" pageNumber="55">Colouration.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">In life</emphasis>
|
||||
(paratype PEM R24232, Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Live photos of Angolan flat geckos: A. Afroedura bogerti (PEM R 24187); B. Afroedura wulfhaackei sp. nov. (paratype PEM R 24232); C. Afroedura praedicta sp. nov. (holotype NB 854); D. Afroedura donveae sp. nov. (holotype PEM R 17937); E. Afroedura vazpintorum sp. nov. (holotype PEM R 24118); F. Afroedura vazpintorum sp. nov. (NB 0745); Photos: A, B, D, E. William R. Branch; C, F. Pedro Vaz Pinto." figureDoi="10.3897/zse.97.57202.figure5" httpUri="https://binary.pensoft.net/fig/496674" pageId="0" pageNumber="55">5B</figureCitation>
|
||||
): Greyish above with five irregularly-spaced darker crossbars from the occiput to the sacrum, each crossbar consisting posteriorly out of three to four black scales wide forming a W-shape; anterior to W-shape are 8-10 scales deep with a mix of dark grey and mustard colours; each dark crossbar separated by light grey to beige blotches; head with irregular dark grey-mustard blotches on the crown with intervening light grey colouration; dark mustard to dark grey bar from nostril to the anterior margins of the ear opening; a vague, thin pale grey canthal stripe, extends on both sides from the nasal region to anterior margins of eye; upper and lower labials grey with diffuse mustard edges; lateral sides of the body with a mix of dark grey and yellow-mustard colouration; limbs greyish above with scattered darker grey markings with intervening yellow-mustard colouration; tail (regenerated) with irregular grey-mustard mottling; iris golden with a black narrow elliptic pupil with crenulated edge and black reticulation with light grey intervening blotches; ventrum uniform greyish with scattered black specks; ventral limbs with scattered black specks, more prominent than on the ventrum.
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">In preservative</emphasis>
|
||||
(holotype PEM R24234, Fig.
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Afroedura wulfhaackei sp. nov. (PEM R 24234) from Maka-Mombolo, Benguela Province, Angola. Photos: Werner Conradie." figureDoi="10.3897/zse.97.57202.figure7" httpUri="https://binary.pensoft.net/fig/496676" pageId="0" pageNumber="55">7</figureCitation>
|
||||
): Dorsum with five irregularly-spaced dark grey W-shaped crossbars from the occiput to the sacrum with beige intervening blotches; ventrum is beige with numerous small scattered black specks on each scale, more prominent posteriorly.
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Variation</emphasis>
|
||||
: Greyish to brownish above with five to six irregularly-spaced darker grey-brown W-shaped crossbars from the occiput to the sacrum, limbs and tail with grey blotches; ventrum uniform greyish with scattered black specks. Juveniles have sharper patterns and colours.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zse.97.57202.figure7" httpUri="https://binary.pensoft.net/fig/496676" pageId="0" pageNumber="55" start="Figure 7" startId="F7">
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<emphasis bold="true" pageId="0" pageNumber="55">Figure 7.</emphasis>
|
||||
Holotype of
|
||||
<taxonomicName authorityName="Branch & Schmitz & Lobón-Rovira & Baptista & António & Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="wulfhaackei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. (PEM R24234) from Maka-Mombolo, Benguela Province, Angola. Photos: Werner Conradie.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="natural history">
|
||||
<paragraph pageId="0" pageNumber="55">Natural history and habitat</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
<emphasis bold="true" pageId="0" pageNumber="55">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Typical habitat of Angolan Afroedura. A. Namba area (A. bogerti); B. Mt Sandula (Afroedura sp. 5); C. Serra da Neve (A. praedicta sp. nov.); D. Omauha Lodge (A. donveae sp. nov.); E. Farm Mucungo (A. vazpintorum sp. nov.); F. Bimbe (A. vazpintorum sp. nov.). Photos: A. Javier Lobon-Rovira; B, D, E. William R. Branch; C. Pedro Vaz Pinto; F. Ninda L. Baptista." figureDoi="10.3897/zse.97.57202.figure4" httpUri="https://binary.pensoft.net/fig/496673" pageId="0" pageNumber="55">4B</figureCitation>
|
||||
).
|
||||
</emphasis>
|
||||
An exclusively rupicolous species living in crevices between rocks or under flakes of exfoliating rocks in boulders at elevations of 920-2,055 m a.s.l. Specimens were collected in cracks of relatively small- to medium-sized boulders of carbonatitic origin surrounded by montane grassland. Some individuals were also found under flakes in large granite boulders and often in steep vertical rock faces and overhangs.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="55" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="55">Distribution and conservation.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="55">
|
||||
This species is currently known from southern Cuanza-Sul, central Huambo and northern Benguela Provinces, Angola (Fig.
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Digital elevation map showing the predicted distribution of the five newly-recognised species of Afroedura occurring in Angola (data for A. wulfhaackei sp. nov. also contain the morphologically-indistinguishable Afroedura sp. 5 - 7 clades). Confirmed locality records are indicated by solid circles, with type localities indicated by stars. Coloured polygons indicate potential distribution areas with more than 90 % of climatic habitat suitability." figureDoi="10.3897/zse.97.57202.figure1" httpUri="https://binary.pensoft.net/fig/496670" pageId="0" pageNumber="55">1</figureCitation>
|
||||
). It appears to have a relatively large, but patchy, distribution on the Angolan highlands and may extend its range into neighbouring provinces. Although sometimes locally common, it appears to be absent from vast areas in-between, where the species would be expected to occur. Populations in isolated granite outcrops may be threatened by removal of rock flakes for construction of homes and other buildings.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
86
data/FB/80/F9/FB80F98FCCEF5862B5FFEBC743EA9665.xml
Normal file
86
data/FB/80/F9/FB80F98FCCEF5862B5FFEBC743EA9665.xml
Normal file
|
|
@ -0,0 +1,86 @@
|
|||
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.934.49157" ID-GBIF-Dataset="cf3fd874-3850-4a1b-ad76-0cb80a894cee" ID-PMC="PMC7250956" ID-Pensoft-Pub="1313-2970-934-93" ID-Pensoft-UUID="E2CB96C5CE7E51178249BE8A83235383" ID-PubMed="32508496" ID-ZooBank="B6D4D8BAB7D041B781716E08759B3933" ModsDocID="1313-2970-934-93" checkinTime="1589939926823" checkinUser="pensoft" docAuthor="Banamar, Ouarda, Chandler, Peter J., Driauach, Ouafaa & Belqat, Boutaina" docDate="2020" docId="FB80F98FCCEF5862B5FFEBC743EA9665" docLanguage="en" docName="ZooKeys 934: 93-110" docOrigin="ZooKeys 934" docSource="http://dx.doi.org/10.3897/zookeys.934.49157" docTitle="Brevicornu verralli" docType="treatment" docVersion="4" id="E2CB96C5CE7E51178249BE8A83235383" lastPageNumber="93" masterDocId="E2CB96C5CE7E51178249BE8A83235383" masterDocTitle="New faunistic records of the family Mycetophilidae (Insecta, Diptera) from Morocco" masterLastPageNumber="110" masterPageNumber="93" pageNumber="93" updateTime="1668168863374" updateUser="ExternalLinkService">
|
||||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>New faunistic records of the family Mycetophilidae (Insecta, Diptera) from Morocco</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Banamar, Ouarda</mods:namePart>
|
||||
<mods:affiliation>Department of Biology, Faculty of Sciences, University Abdelmalek Essaadi, BP 2121, Tetouan, Morocco</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Chandler, Peter J.</mods:namePart>
|
||||
<mods:affiliation>606 B Berryfield Lane, Melksham, Wilts SN 12 6 EL, UK</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Driauach, Ouafaa</mods:namePart>
|
||||
<mods:affiliation>Department of Biology, Faculty of Sciences, University Abdelmalek Essaadi, BP 2121, Tetouan, Morocco</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Belqat, Boutaina</mods:namePart>
|
||||
<mods:affiliation>Department of Biology, Faculty of Sciences, University Abdelmalek Essaadi, BP 2121, Tetouan, Morocco</mods:affiliation>
|
||||
<mods:nameIdentifier type="email">b_belqat@hotmail.com</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>ZooKeys</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2020</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>934</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>93</mods:start>
|
||||
<mods:end>110</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://dx.doi.org/10.3897/zookeys.934.49157</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.934.49157</mods:identifier>
|
||||
<mods:identifier type="Pensoft-Pub">1313-2970-934-93</mods:identifier>
|
||||
<mods:identifier type="ZooBank">B6D4D8BAB7D041B781716E08759B3933</mods:identifier>
|
||||
<mods:identifier type="Pensoft-UUID">E2CB96C5CE7E51178249BE8A83235383</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment ID-GBIF-Taxon="164241298" LSID="urn:lsid:plazi:treatment:FB80F98FCCEF5862B5FFEBC743EA9665" httpUri="http://treatment.plazi.org/id/FB80F98FCCEF5862B5FFEBC743EA9665" lastPageNumber="93" pageId="0" pageNumber="93">
|
||||
<subSubSection pageId="0" pageNumber="93" type="nomenclature">
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
*
|
||||
<taxonomicName LSID="FB80F98F-CCEF-5862-B5FF-EBC743EA9665" authority="(Edwards, 1925)" baseAuthorityName="Edwards" baseAuthorityYear="1925" class="Insecta" family="Mycetophilidae" genus="Brevicornu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brevicornu verralli" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="verralli">Brevicornu verralli (Edwards, 1925)</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="literature records">
|
||||
<paragraph pageId="0" pageNumber="93">Literature records.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
Recorded from North Africa (Tunisia) (
|
||||
<bibRefCitation author="Chandler, PJ" journalOrPublisher="Israel Journal of Entomology" pageId="0" pageNumber="93" pagination="1 - 100" refId="B3" refString="Chandler, PJ, 1994. The fungus gnats of Israel (Diptera: Sciaroidea, excluding Sciaridae). Israel Journal of Entomology 28: 1 - 100" title="The fungus gnats of Israel (Diptera: Sciaroidea, excluding Sciaridae)." volume="28" year="1994">Chandler 1994</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Chandler, PJ" journalOrPublisher="Boletim do Museu municipal do Funchal (Historia Natural), Suplemento" pageId="0" pageNumber="93" pagination="1 - 170" refId="B5" refString="Chandler, PJ, Ribeiro, E, 1995. The Sciaroidea (Diptera) (excluding Sciaridae) of the Atlantic Islands (Canary Islands, Madeira and the Azores). Boletim do Museu municipal do Funchal (Historia Natural), Suplemento 3: 1 - 170" title="The Sciaroidea (Diptera) (excluding Sciaridae) of the Atlantic Islands (Canary Islands, Madeira and the Azores)." volume="3" year="1995">Chandler and Ribeiro 1995</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="93" type="new records">
|
||||
<paragraph pageId="0" pageNumber="93">New records.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="93">
|
||||
Rif: Daya Afersiw, 1♂, 11/VI/2013;
|
||||
<normalizedToken originalValue="Forêt-Jbel">Foret-Jbel</normalizedToken>
|
||||
Lekraa, 1♂, 12/VI/2013, coll. Ebejer.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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