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<paragraph id="5B79EAC2FB6C4D18B5D0DB418AA1B649" pageNumber="9">Buchara (Schaar pres de Samarkand, 1 [[ worker ]], 24. IV. 187. Kaznakov!; Buchara orient. 2 [[ worker ]]. Barscevsku!); Region transcaspienne (St. Giaurs, 7 [[ worker ]]; St. Bairam-Ali, 2 [[ worker ]], 27. IX; St. Tedzen, 1 [[ worker ]], 17. VII. 1897. Ahnger!); Transcaucasie (Gouv. Baku, Col do Bum, 1 [[ worker ]], Schelkovnikov.).</paragraph>
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Key to puparia of the species of
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1. Most abdominal segments, with a submedian glandular area (fig. 7).....
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­ Abdominal segments without such glandular areas, but median tubercles or tuberclelike markings may present on abdominal segments ....................................................
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2. Abdominal segments without median tubercles or tubercle­like markings, apparent margin broadly crenate, each submarginal ridge correlated with two or three marginal teeth...............................................................................................................................
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­ Not with such combinations of characters ..................................................................
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1. Syntype, Belgian Congo: Barumbu, i.1926, (J. Ghesquière), on
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sp., B.M. 1933 (BMNH). 2. Nigeria: Ibadan, 20.i.1914, (W.A. Lambourn), on herbaceous plant, 204/61, B.M. 1916­190 (BMNH).
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3. Most abdominal segments with median tubercles, margin strongly recurved, apparent margin smooth, real margin weakly crenate ..............................................
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­ Most abdominal segments with tubercle­like markings, margin normally not or at most slightly recurved, real margin strongly crenate..............................................
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<mods:identifier id="348EEBCDD54BF1CB73B14899BD2FE54B" type="Zenodo-Dep">170652</mods:identifier>
</mods:mods>
<treatment id="DF3E1007F925FFCF9E79FAC17C40FAF8" ID-DOI="http://doi.org/10.5281/zenodo.6265768" ID-GBIF-Taxon="119334226" ID-Zenodo-Dep="6265768" LSID="urn:lsid:plazi:treatment:DF3E1007F925FFCF9E79FAC17C40FAF8" httpUri="http://treatment.plazi.org/id/DF3E1007F925FFCF9E79FAC17C40FAF8" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection id="1F8DF29AF925FFCD9E79FAC179F8FA60" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="5728A111F925FFCD9E79FAC17A0FFA85" blockId="1.[264,630,1353,1414]" box="[264,630,1353,1379]" pageId="1" pageNumber="2">
<heading id="0C60167DF925FFCD9E79FAC17A0FFA85" bold="true" box="[264,630,1353,1379]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="65E37D03F925FFCD9E79FAC17A0FFA85" bold="true" box="[264,630,1353,1379]" pageId="1" pageNumber="2">
<taxonomicName id="9097DA92F925FFCD9E79FAC17A68FA85" ID-CoL="93K7" box="[264,529,1353,1379]" class="Insecta" family="Aleyrodidae" genus="Acaudaleyrodes" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ebeni" status="sp. nov.">
<emphasis id="65E37D03F925FFCD9E79FAC17A68FA85" bold="true" box="[264,529,1353,1379]" italics="true" pageId="1" pageNumber="2">Acaudaleyrodes ebeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED0C078F925FFCD9D69FAC17A0FFA85" box="[536,630,1353,1379]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5728A111F925FFCD9E79FAE479F8FA60" blockId="1.[264,630,1353,1414]" box="[264,385,1388,1414]" pageId="1" pageNumber="2">
(
<figureCitation id="CFACBD94F925FFCD9E61FAE47901FA60" box="[272,376,1388,1414]" captionStart="FIGURES 1 9" captionStartId="2.[264,383,1128,1152]" captionTargetBox="[528,1052,283,1092]" captionTargetId="figure@2.[521,1065,272,1104]" captionTargetPageId="2" captionText="FIGURES 1 9. Acaudaleyrodes ebeni sp. nov. 1. Habitus photograph of puparia. 2. Puparium. 3. Margin and submargin. 4. Antenna. 5. Adhesive sac, mesal of pro / meso­thoracic legs. 6. Geminate pore / porettes and pronounced “ bright ” simple pores. 7. Submedian glandular area on an abdominal segment showing a very small polygonal reticulate pattern. 8. Posterior marginal seta. 9. Abdominal segmentation; ventral surface." httpUri="https://zenodo.org/record/170653/files/figure.png" pageId="1" pageNumber="2">Figs 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8DF29AF925FFCF9E79FA3479CFFC80" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="5728A111F925FFCD9E79FA347CC7F9A8" blockId="1.[264,1325,1468,1894]" pageId="1" pageNumber="2">DESCRIPTION. In life, puparia often occurring in aggregations, attached to the spine­like leaves of the host plant (fig. 1); cuticle shiny dark brown to black, with obvious peripheral and submedian dorsal white wax secretions, but subdorsum remains exposed; dorsal wax secretions relatively long and somewhat curly. Puparia often considerably raised.</paragraph>
<paragraph id="5728A111F925FFCD9E49F9D47A3CF880" blockId="1.[264,1325,1468,1894]" pageId="1" pageNumber="2">
<emphasis id="65E37D03F925FFCD9E49F9D479F7F993" box="[312,398,1628,1653]" italics="true" pageId="1" pageNumber="2">Margin</emphasis>
. Puparia oval (fig. 2), possibly sexually dimorphic, i.e. male puparia smaller than female ones, but size ranges overlap, 0.701.00 mm long,
<quantity id="906F0CF4F925FFCD9C99F90C7CEBF978" box="[1000,1170,1668,1694]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" metricValueMax="7.199999999999999" metricValueMin="5.0" pageId="1" pageNumber="2" unit="mm" value="0.61" valueMax="0.72" valueMin="0.5">0.500.72 mm</quantity>
wide; broadest across abdominal segment III/IV. Margin crenate (fig. 3), about 1113 crenations occupying
<quantity id="906F0CF4F925FFCD9E25F95C79C8F908" box="[340,433,1748,1774]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="2" unit="mm" value="0.1">0.1 mm</quantity>
of margin, not modified at thoracic and caudal tracheal openings; margin normally not, or only slightly recurved, depending on the mounting processes. Anterior and posterior marginal setae present, fine and apparently arising slightly ventrally, posterior pair the longer (fig. 8).
</paragraph>
<caption id="03E8F199F926FFCE9E79FBE07A08FAEC" httpUri="https://zenodo.org/record/170653/files/figure.png" pageId="2" pageNumber="3" targetBox="[528,1052,283,1092]" targetPageId="2">
<paragraph id="5728A111F926FFCE9E79FBE07A08FAEC" blockId="2.[264,1323,1128,1290]" pageId="2" pageNumber="3">
<emphasis id="65E37D03F926FFCE9E79FBE079CAFB66" bold="true" box="[264,435,1128,1152]" pageId="2" pageNumber="3">FIGURES 19.</emphasis>
<taxonomicName id="9097DA92F926FFCE9EC8FBE17ADDFB66" box="[441,676,1129,1152]" class="Insecta" family="Aleyrodidae" genus="Acaudaleyrodes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ebeni" status="sp. nov.">
<emphasis id="65E37D03F926FFCE9EC8FBE17ADDFB66" box="[441,676,1129,1152]" italics="true" pageId="2" pageNumber="3">Acaudaleyrodes ebeni</emphasis>
</taxonomicName>
<emphasis id="65E37D03F926FFCE9DDBFBE07B48FB66" bold="true" box="[682,817,1128,1152]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="7ED0C078F926FFCE9DDBFBE07A86FB66" box="[682,767,1128,1152]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
1.
</emphasis>
Habitus photograph of puparia.
<emphasis id="65E37D03F926FFCE9BF9FBE07CE5FB66" bold="true" box="[1160,1180,1128,1152]" pageId="2" pageNumber="3">2.</emphasis>
Puparium.
<emphasis id="65E37D03F926FFCE9A67FBE07D53FB66" bold="true" box="[1302,1322,1128,1152]" pageId="2" pageNumber="3">3.</emphasis>
Margin and submargin.
<emphasis id="65E37D03F926FFCE9D78FB037A65FB45" bold="true" box="[521,540,1163,1187]" pageId="2" pageNumber="3">4.</emphasis>
Antenna.
<emphasis id="65E37D03F926FFCE9DFDFB037AD8FB45" bold="true" box="[652,673,1163,1187]" pageId="2" pageNumber="3">5.</emphasis>
Adhesive sac, mesal of pro/meso­thoracic legs.
<emphasis id="65E37D03F926FFCE9BD9FB037CC5FB45" bold="true" box="[1192,1212,1163,1187]" pageId="2" pageNumber="3">6.</emphasis>
Geminate pore/porettes and pronounced “bright” simple pores.
<emphasis id="65E37D03F926FFCE9C31FB257B2AFB23" bold="true" box="[832,851,1197,1221]" pageId="2" pageNumber="3">7.</emphasis>
Submedian glandular area on an abdominal segment showing a very small polygonal reticulate pattern.
<emphasis id="65E37D03F926FFCE9CFCFB587BD8FB0E" bold="true" box="[909,929,1232,1256]" pageId="2" pageNumber="3">8.</emphasis>
Posterior marginal seta.
<emphasis id="65E37D03F926FFCE9BDFFB587CBAFB0E" bold="true" box="[1198,1219,1232,1256]" pageId="2" pageNumber="3">9.</emphasis>
Abdominal segmentation; ventral surface.
</paragraph>
</caption>
<paragraph id="5728A111F926FFCF9E49FAEE7C04FD90" blockId="2.[264,1323,1382,1894]" lastBlockId="3.[264,1323,284,1310]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="65E37D03F926FFCE9E49FAEE79EFFA99" box="[312,406,1382,1407]" italics="true" pageId="2" pageNumber="3">Dorsum</emphasis>
. Abdominal rhachis prominent, with lateral arms. First abdominal setae absent, eighth abdominal, caudal and cephalic setae present; submargin with seven pairs of setae (excluding the caudal setae), three pairs on cephalothorax and four pairs on abdomen (one pair on each abdominal segments four to seven), abdominal submarginal setae somewhat stout and slightly longer than submarginal setae on cephalothorax. A pair of faint small crescent­shaped folds postero­lateral to cephalic setae; a pair of subdorsal longitudinal furrow extending from the latter folds to 2nd abdominal segment; meso­metathoracic suture well defined. Transverse moulting suture reaching apically to anterior to thoracoabdominal suture but not reaching margin; longitudinal moulting suture almost reaching margin. Geminate pore/porettes (fig. 6) scattered in subdorsal area; outer subdorsum with a row of pronounced “bright” simple pores, somewhat thick­rimmed (fig. 6), which sometimes occur in pairs, but normally separated by a distance of more than the width of one pore. Length of 7th abdominal segment reduced medially; each abdominal segment with a median glandular area showing a very small polygonal reticulate pattern (fig. 7), but that of abdominal segment VII very small and in some specimens not visible; the same median reticular pattern also present on cephalothorax, extending anteriorly close to margin. Vasiform orifice typical for the genus, scutellate, longer than wide, elevated posteriorly, inset from posterior margin of pupal case by less than, rarely by about, its own length, floor with radiating lines of microtubercles; operculum nearly rectangular, width about twice as long as length, only occupying about a quarter of orifice; lingula extremely small, hardly visible, apparently not expanded apically, lingular terminal setae not discernible and/or difficult to see at least at 400 magnification. Caudal ridges and furrow absent.
</paragraph>
<paragraph id="5728A111F927FFCF9E49FD0C79CFFC80" blockId="3.[264,1323,284,1310]" pageId="3" pageNumber="4">
<emphasis id="65E37D03F927FFCF9E49FD0C7906FD7B" box="[312,383,644,669]" italics="true" pageId="3" pageNumber="4">Ven te r</emphasis>
. Thoracic and caudal tracheal folds at most poorly indicated only by faint pairs of boundary lines in some specimens. Adhesive sacs well developed, mesal of pro/mesothoracic legs (fig. 5). Each leg with an apical pad; one microseta present at the base of each middle and hind leg. Antennae short, mesal to prothoracic legs (fig. 4). Ventral abdominal setae hair­like, long enough to be easily recognisable. Abdominal segmentation plainly visible (fig. 9).
</paragraph>
</subSubSection>
<subSubSection id="1F8DF29AF927FFCF9E49FCFC7C76FBC8" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="5728A111F927FFCF9E49FCFC7C76FBC8" blockId="3.[264,1323,284,1310]" pageId="3" pageNumber="4">
MATERIAL EXAMINED.
<typeStatus id="882C1FB3F927FFCF9DF4FCFC7B5AFC68" box="[645,803,884,910]" pageId="3" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
, puparium,
<collectingCountry id="2F80E181F927FFCF9CC0FCFC7B91FC68" box="[945,1000,884,910]" name="Iran" pageId="3" pageNumber="4">
<emphasis id="65E37D03F927FFCF9CC0FCFC7B91FC68" bold="true" box="[945,1000,884,910]" pageId="3" pageNumber="4">Iran</emphasis>
</collectingCountry>
,
<collectingCountry id="2F80E181F927FFCF9C88FCFC7C55FC68" box="[1017,1068,884,910]" name="Iran" pageId="3" pageNumber="4">Fars</collectingCountry>
province, Lar, Pardi/ Pardis mountain, 2216 metres, 54° 27', 27° 28', on
<taxonomicName id="9097DA92F927FFCF9C00FC147C5EFC53" box="[881,1063,924,949]" class="Magnoliopsida" family="Fabaceae" genus="Ebenus" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="65E37D03F927FFCF9C00FC147C5EFC53" box="[881,1063,924,949]" italics="true" pageId="3" pageNumber="4">Ebenus stellata</emphasis>
</taxonomicName>
,
<date id="232987D1F927FFCF9B49FC147CC1FC50" box="[1080,1208,924,950]" pageId="3" pageNumber="4" value="2000-06-21">21.vi.2000</date>
, (H. Alemansoor) (
<collectionCode id="318639D4F927FFCF9EF4FC4C79A6FC38" box="[389,479,964,990]" httpUri="http://grbio.org/cool/gj99-xj2c" name="Jardí Botànic Marimurtra" pageId="3" pageNumber="4">HMIM</collectionCode>
).
<typeStatus id="882C1FB3F927FFCF9E83FC4C7AE4FC38" box="[498,669,964,990]" pageId="3" pageNumber="4" type="paratype">PARATYPES</typeStatus>
,
<emphasis id="65E37D03F927FFCF9DD9FC4C7AB3FC3B" box="[680,714,964,989]" italics="true" pageId="3" pageNumber="4">ca.</emphasis>
80 puparia, 1 third instar larva, 2?second instar larvae on slides, many puparia dry on leaves,
<collectingCountry id="2F80E181F927FFCF9C7BFC647B38FBE0" box="[778,833,1004,1030]" name="Iran" pageId="3" pageNumber="4">
<emphasis id="65E37D03F927FFCF9C7BFC647B38FBE0" bold="true" box="[778,833,1004,1030]" pageId="3" pageNumber="4">Iran</emphasis>
</collectingCountry>
, same data as
<typeStatus id="882C1FB3F927FFCF9C81FC647C2FFBE0" box="[1008,1110,1004,1030]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
(
<collectionCode id="318639D4F927FFCF9B17FC647CC7FBE0" box="[1126,1214,1004,1030]" httpUri="http://grbio.org/cool/gj99-xj2c" name="Jardí Botànic Marimurtra" pageId="3" pageNumber="4">HMIM</collectionCode>
,
<collectionCode id="318639D4F927FFCF9BBAFC647D5FFBE0" box="[1227,1318,1004,1030]" pageId="3" pageNumber="4">BMNH</collectionCode>
,
<collectionCode id="318639D4F927FFCF9E79FB9C791BFBC8" LSID="urn:lsid:biocol.org:col:34871" box="[264,354,1044,1070]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
); 3 puparia,
<date id="232987D1F927FFCF9E83FB9C7A24FBC8" box="[498,605,1044,1070]" pageId="3" pageNumber="4" value="1993-12-21">21.xii.93</date>
, other data as for
<typeStatus id="882C1FB3F927FFCF9C5FFB9C7BEDFBC8" box="[814,916,1044,1070]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
(
<collectionCode id="318639D4F927FFCF9CD5FB9C7C7AFBC8" box="[932,1027,1044,1070]" pageId="3" pageNumber="4">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="1F8DF29AF927FFCF9E49FBB47BDBFBB0" box="[312,930,1084,1110]" pageId="3" pageNumber="4" type="etymology">
<paragraph id="5728A111F927FFCF9E49FBB47BDBFBB0" blockId="3.[264,1323,284,1310]" box="[312,930,1084,1110]" pageId="3" pageNumber="4">
ETYMOLOGY. Named after its host genus,
<taxonomicName id="9097DA92F927FFCF9C34FBB47BE2FBB3" box="[837,923,1084,1109]" class="Magnoliopsida" family="Fabaceae" genus="Ebenus" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="65E37D03F927FFCF9C34FBB47BE2FBB3" box="[837,923,1084,1109]" italics="true" pageId="3" pageNumber="4">Ebenus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1F8DF29AF927FFCF9E49FBEC7B84FB40" pageId="3" pageNumber="4" type="distribution">
<paragraph id="5728A111F927FFCF9E49FBEC7A33FB98" blockId="3.[264,1323,284,1310]" box="[312,586,1124,1150]" pageId="3" pageNumber="4">
DISTRIBUTION.
<collectingCountry id="2F80E181F927FFCF9D62FBEC7A3FFB98" box="[531,582,1124,1150]" name="Iran" pageId="3" pageNumber="4">Iran</collectingCountry>
.
</paragraph>
<paragraph id="5728A111F927FFCF9E49FB047B84FB40" blockId="3.[264,1323,284,1310]" box="[312,1021,1164,1190]" pageId="3" pageNumber="4">
HOST PLANTS.
<taxonomicName id="9097DA92F927FFCF9D78FB047A02FB40" box="[521,635,1164,1190]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
:
<taxonomicName id="9097DA92F927FFCF9DF4FB047B4FFB40" box="[645,822,1164,1190]" class="Magnoliopsida" family="Fabaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="4" phylum="Magnoliophyta" rank="family">Papilionoideae</taxonomicName>
,
<taxonomicName id="9097DA92F927FFCF9C33FB047B8FFB43" box="[834,1014,1164,1189]" class="Magnoliopsida" family="Fabaceae" genus="Ebenus" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="65E37D03F927FFCF9C33FB047B8FFB43" box="[834,1014,1164,1189]" italics="true" pageId="3" pageNumber="4">Ebenus stellata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1F8DF29AF927FFCF9E49FB3C7C40FAF8" pageId="3" pageNumber="4" type="discussion">
<paragraph id="5728A111F927FFCF9E49FB3C7C40FAF8" blockId="3.[264,1323,284,1310]" pageId="3" pageNumber="4">COMMENTS. This species differs from the other described species of the genus in possessing a very distinctive glandular pattern along the median dorsal line of the puparia, which presumably associates with secreting the median dorsal waxes.</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1015.59019" ID-GBIF-Dataset="c25085a4-18a8-44a1-90a6-f24d2f27b272" ID-PMC="PMC7878466" ID-Pensoft-Pub="1313-2970-1015-87" ID-Pensoft-UUID="8129A24C3ED654AE88820BF4B447B42A" ID-PubMed="33613042" ID-ZooBank="C45FA29810C54869B4788E8502521940" ModsDocID="1313-2970-1015-87" checkinTime="1612497292298" checkinUser="pensoft" docAuthor="Chen, Juanjuan, Schmelz, Ruediger M. &amp; Xie, Zhicai" docDate="2021" docId="DF3E280FF93651FB8418CCB6BA1ACB36" docLanguage="en" docName="ZooKeys 1015: 87-97" docOrigin="ZooKeys 1015" docPubDate="2021-02-04" docSource="http://dx.doi.org/10.3897/zookeys.1015.59019" docTitle="Hemienchytraeus wuhanensis Chen &amp; Schmelz &amp; Xie 2021, sp. nov." docType="treatment" docUuid="D3137BCA-E1CC-4FC7-AA55-A88FA9ED06E6" docUuidSource="ZooBank" docVersion="4" id="8129A24C3ED654AE88820BF4B447B42A" lastPageNumber="87" masterDocId="8129A24C3ED654AE88820BF4B447B42A" masterDocTitle="Description of Hemienchytraeus wuhanensis sp. nov. (Annelida, Clitellata, Enchytraeidae) from central China, with comments on species records of Hemienchytraeus from China" masterLastPageNumber="97" masterPageNumber="87" pageNumber="87" updateTime="1668149778046" updateUser="ExternalLinkService">
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<mods:title>Description of Hemienchytraeus wuhanensis sp. nov. (Annelida, Clitellata, Enchytraeidae) from central China, with comments on species records of Hemienchytraeus from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Juanjuan</mods:namePart>
<mods:affiliation>The Key Laboratory of Aquatic Biodiversity and Conservation, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, 430072, China &amp; University of Chinese Academy of Sciences, Beijing, 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmelz, Ruediger M.</mods:namePart>
<mods:affiliation>IFAB, Institute for Applied Soil Biology, Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xie, Zhicai</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9658-4888</mods:nameIdentifier>
<mods:affiliation>The Key Laboratory of Aquatic Biodiversity and Conservation, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, 430072, China</mods:affiliation>
<mods:nameIdentifier type="email">zhcxie@ihb.ac.cn</mods:nameIdentifier>
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<mods:date>2021</mods:date>
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<mods:number>2021-02-04</mods:number>
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<mods:detail type="volume">
<mods:number>1015</mods:number>
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<mods:start>87</mods:start>
<mods:end>97</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1015.59019</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1015.59019</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1015-87</mods:identifier>
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<treatment ID-GBIF-Taxon="177212467" LSID="urn:lsid:zoobank.org:act:D3137BCA-E1CC-4FC7-AA55-A88FA9ED06E6" httpUri="http://treatment.plazi.org/id/DF3E280FF93651FB8418CCB6BA1ACB36" lastPageNumber="87" pageId="0" pageNumber="87">
<subSubSection pageId="0" pageNumber="87" type="nomenclature">
<paragraph pageId="0" pageNumber="87">
<taxonomicName LSID="http://zoobank.org/D3137BCA-E1CC-4FC7-AA55-A88FA9ED06E6" authority="Chen &amp; Schmelz &amp; Xie, 2021" authorityName="Chen &amp; Schmelz &amp; Xie" authorityYear="2021" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemienchytraeus wuhanensis" order="Enchytraeida" pageId="0" pageNumber="87" phylum="Annelida" rank="species" species="wuhanensis" status="sp. nov.">Hemienchytraeus wuhanensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="87">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="holotype">
<paragraph pageId="0" pageNumber="87">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="87">Fully mature, whole-mounted specimen, stained, HBO201904002.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="type locality">
<paragraph pageId="0" pageNumber="87">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="87">
Mount Shizi, litter layer of hardwood forest (
<geoCoordinate degrees="30" direction="north" minutes="28" orientation="latitude" precision="1" seconds="42.57" value="30.478493">30°28'42.57&quot;N</geoCoordinate>
,
<geoCoordinate degrees="114" direction="east" minutes="21" orientation="longitude" precision="1" seconds="10.48" value="114.35291">114°21'10.48&quot;E</geoCoordinate>
; 44 m a.s.l.), Huazhong Agricultural University (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitat of Hemienchytraeus wuhanensis sp. nov., Mount Shizi, Huazhong Agriculture University, Wuhan, Hubei Province, China." figureDoi="10.3897/zookeys.1015.59019.figure1" httpUri="https://binary.pensoft.net/fig/505240" pageId="0" pageNumber="87">1</figureCitation>
), Wuhan, Hubei Province, 6 April 2019, coll. Y. H. Ge.
</paragraph>
<caption doi="10.3897/zookeys.1015.59019.figure1" httpUri="https://binary.pensoft.net/fig/505240" pageId="0" pageNumber="87" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Figure 1.</emphasis>
Habitat of
<taxonomicName authorityName="Chen &amp; Schmelz &amp; Xie" authorityYear="2021" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" lsidName="Hemienchytraeus wuhanensis" order="Enchytraeida" pageId="0" pageNumber="87" phylum="Annelida" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">Hemienchytraeus wuhanensis</emphasis>
</taxonomicName>
sp. nov., Mount Shizi, Huazhong Agriculture University, Wuhan, Hubei Province, China.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="paratypes">
<paragraph pageId="0" pageNumber="87">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="87">
HBO201904003, HBO201904004 two whole-mounted fully mature specimens, HBO201904005-HBO201904007, three adult specimens, used entirely for DNA extraction; HBO201904008-HBO201904010 three adult specimens from the type locality maintained in 75% alcohol, same data as holotype. HBO201904001 one whole mounted fully mature specimen, HBO201904011-HBO201904012 two adult specimens used for extract DNA, and HBO201904013-HBO210904015 three adult specimens maintained in 75% alcohol from Mount Luojia, under a pine tree (
<geoCoordinate degrees="30" direction="north" minutes="32" orientation="latitude" precision="1" seconds="05.39" value="30.53483">30°32'05.39&quot;N</geoCoordinate>
,
<geoCoordinate degrees="114" direction="east" minutes="22" orientation="longitude" precision="1" seconds="10.95" value="114.36971">114°22'10.95&quot;E</geoCoordinate>
; 31 m a.s.l.), Wuhan University, Wuhan, Hubei Province, 2 April 2019, coll. X. K. Jiang &amp; J. J. Chen.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="etymology">
<paragraph pageId="0" pageNumber="87">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="87">Named after the city where the species was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="distribution">
<paragraph pageId="0" pageNumber="87">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="87">Mineral soil and organic layers under camphor trees near a narrow, tarred road at Mount Shizi, Huazhong Agricultural University; mineral soil and organic layers under pine trees at Mount Luojia, Wuhan University. The two hills are about 10 km apart, with little human disturbance.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="diagnosis">
<paragraph pageId="0" pageNumber="87">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="87">This new species can be recognized by the following combination of diagnostic traits: (1) chaetae anteriorly and posteriorly of about the same size, not enlarged in caudal segments; (2) oesophageal appendage with tertiary branches; (3) three pairs of secondary pharyngeal gland ventral lobes in V, VI, VII, small in VII; (4) five pairs of preclitellar nephridia in 5/6-9/10; (5) dorsal vessel originating in clitellum segments; (6) clitellum girdle-shaped; (7) seminal vesicle absent; (8) spermathecae extending to VI-VII, not enlarged.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">Description.</paragraph>
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Length</emphasis>
6.5-9.3 mm (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
),
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">diameter</emphasis>
0.3-0.4 mm (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
) at clitellum.
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Segment number</emphasis>
37-42. Two chaetae per bundle throughout, absent in XII in mature specimens.
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Chaetae</emphasis>
straight with slight proximal bend; in anterior segments, slight distal bend in opposite direction of proximal bend, i.e., chaetae faintly sigmoid; in proximal segments, chaetae distally straight. Chaetae in preclitellar bundles 37.5-42 mm long, diameter 5 mm, 27.5-32.5 mm in postclitellar segments, diameter 5 mm.
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Head pore</emphasis>
mid-dorsally on prostomium.
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Epidermal gland cells</emphasis>
gray, three to four transverse rows per segment, the cells nearly rectangular and arranged in regular pattern (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3E</figureCitation>
).
<taxonomicName class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clitellum" order="Aplousobranchia" pageId="0" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Clitellum</emphasis>
</taxonomicName>
in XII-1/2XIII, inconspicuous thickening, cells ca 5-9 mm high, girdle-shaped (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3I, J</figureCitation>
), hyalocytes and granulocytes in reticulate arrangement with hyalocytes taking larger proportion dorsally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3I</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Body wall</emphasis>
25-37.5 mm thick.
</paragraph>
<paragraph pageId="0" pageNumber="87">
<taxonomicName genus="Brain" lsidName="Brain" pageId="0" pageNumber="87" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Brain</emphasis>
</taxonomicName>
about as long as wide (117 mm long, 93 mm wide,
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
), slightly indented anteriorly, deeply incised posteriorly (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hemienchytraeus wuhanensis sp. nov. A anterior body region, anterior 13 segments, lateral view, schematic B brain C spermatheca; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir D pharyngeal glands E oesophageal appendage F sperm funnel." figureDoi="10.3897/zookeys.1015.59019.figure2" httpUri="https://binary.pensoft.net/fig/505241" pageId="0" pageNumber="87">2B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3A</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Oesophageal appendage</emphasis>
arising from mid-dorsal region of pharynx in III as an unpaired root with large proximal chamber; following section longer than proximal chamber, with thick, meandering canal; two primary branches, longer than root, with smaller canal; each primary branch bifurcating into two short, secondary branches; each secondary branch bifurcating into four or more tertiary branches, the latter difficult to distinguish. Secondary and tertiary branches of same diameter, thinner than primary branches (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hemienchytraeus wuhanensis sp. nov. A anterior body region, anterior 13 segments, lateral view, schematic B brain C spermatheca; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir D pharyngeal glands E oesophageal appendage F sperm funnel." figureDoi="10.3897/zookeys.1015.59019.figure2" httpUri="https://binary.pensoft.net/fig/505241" pageId="0" pageNumber="87">2E</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3B, C</figureCitation>
). All three pairs of
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">pharyngeal glands</emphasis>
united dorsally, primary ventral lobes in V and VI. Three pairs of secondary pharyngeal gland lobes in V, VI and VII, small in VII (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hemienchytraeus wuhanensis sp. nov. A anterior body region, anterior 13 segments, lateral view, schematic B brain C spermatheca; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir D pharyngeal glands E oesophageal appendage F sperm funnel." figureDoi="10.3897/zookeys.1015.59019.figure2" httpUri="https://binary.pensoft.net/fig/505241" pageId="0" pageNumber="87">2D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3D</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Dorsal vessel</emphasis>
from XII-XIII, blood colorless.
</paragraph>
<caption doi="10.3897/zookeys.1015.59019.figure2" httpUri="https://binary.pensoft.net/fig/505241" pageId="0" pageNumber="87" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Figure 2.</emphasis>
<taxonomicName authorityName="Chen &amp; Schmelz &amp; Xie" authorityYear="2021" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" lsidName="Hemienchytraeus wuhanensis" order="Enchytraeida" pageId="0" pageNumber="87" phylum="Annelida" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">Hemienchytraeus wuhanensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="87">A</emphasis>
anterior body region, anterior 13 segments, lateral view, schematic
<emphasis bold="true" pageId="0" pageNumber="87">B</emphasis>
brain
<emphasis bold="true" pageId="0" pageNumber="87">C</emphasis>
spermatheca; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir
<emphasis bold="true" pageId="0" pageNumber="87">D</emphasis>
pharyngeal glands
<emphasis bold="true" pageId="0" pageNumber="87">E</emphasis>
oesophageal appendage
<emphasis bold="true" pageId="0" pageNumber="87">F</emphasis>
sperm funnel.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Figure 3.</emphasis>
Micrographs of
<taxonomicName authorityName="Chen &amp; Schmelz &amp; Xie" authorityYear="2021" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" lsidName="Hemienchytraeus wuhanensis" order="Enchytraeida" pageId="0" pageNumber="87" phylum="Annelida" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">Hemienchytraeus wuhanensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="87">A, B, E-I, K, L</emphasis>
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
<emphasis bold="true" pageId="0" pageNumber="87">C, D, J</emphasis>
fixed
<emphasis bold="true" pageId="0" pageNumber="87">A</emphasis>
brain
<emphasis bold="true" pageId="0" pageNumber="87">B</emphasis>
dorsal view of oesophageal appendage
<emphasis bold="true" pageId="0" pageNumber="87">C</emphasis>
lateral view of oesophageal appendage
<emphasis bold="true" pageId="0" pageNumber="87">D</emphasis>
pharyngeal glands
<emphasis bold="true" pageId="0" pageNumber="87">E e</emphasis>
pidermal gland cells in II-V ventrally
<emphasis bold="true" pageId="0" pageNumber="87">F</emphasis>
spermathecae and pharyngeal glands
<emphasis bold="true" pageId="0" pageNumber="87">G</emphasis>
nephridia in 7/8, anteseptale bottom-left
<emphasis bold="true" pageId="0" pageNumber="87">H</emphasis>
male glandular bulb, slightly everted
<emphasis bold="true" pageId="0" pageNumber="87">I</emphasis>
dorsal view of clitellum
<emphasis bold="true" pageId="0" pageNumber="87">J</emphasis>
ventral view of clitellum
<emphasis bold="true" pageId="0" pageNumber="87">K</emphasis>
sperm funnel
<emphasis bold="true" pageId="0" pageNumber="87">L</emphasis>
sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="87">
Five pairs of preclitellar
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">nephridia</emphasis>
from 5/6 to 9/10 (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hemienchytraeus wuhanensis sp. nov. A anterior body region, anterior 13 segments, lateral view, schematic B brain C spermatheca; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir D pharyngeal glands E oesophageal appendage F sperm funnel." figureDoi="10.3897/zookeys.1015.59019.figure2" httpUri="https://binary.pensoft.net/fig/505241" pageId="0" pageNumber="87">2A</figureCitation>
); each about 160 mm long and 60 mm wide (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
). Anteseptale globular, with minute and numerous brownish granules at periphery; funnel orientated obliquely ventrad, with small and narrow anterior projection; postseptale elongate, ca twice as long as anteseptale. Efferent duct originating from the middle of the postseptale (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Seminal vesicle</emphasis>
absent, cysts dorsally in XI.
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Sperm funnels</emphasis>
cylindrical, tapering distad, well developed, ca 150-250 mm long and 40 mm at collar (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
). Collar distinct, somewhat narrower than funnel body (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hemienchytraeus wuhanensis sp. nov. A anterior body region, anterior 13 segments, lateral view, schematic B brain C spermatheca; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir D pharyngeal glands E oesophageal appendage F sperm funnel." figureDoi="10.3897/zookeys.1015.59019.figure2" httpUri="https://binary.pensoft.net/fig/505241" pageId="0" pageNumber="87">2F</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3K</figureCitation>
). Spermatozoa ca 140 mm long, heads ca 20 mm long (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
). Sperm ducts elongate, diameter ca 6 mm, loose or tight coils in XII-XIII (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3L</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Male copulatory organs</emphasis>
with distinct musculature, male glandular body globular, ca 85
<normalizedToken originalValue="µm">µm</normalizedToken>
in diameter (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
). No accessory copulatory glands (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3H</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Spermathecae</emphasis>
free, not attached to oesophagus. Ectal pores laterally at 4/5, without ectal gland. Ectal ducts ca 400-500 mm long and 20-26 mm wide (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
), with distinct ampullar dilatation in V. Connecting tube between ampulla proper and ental reservoir thinner than ectal duct, extending into VI or VII, ending in a small, elongately ellipsoid ental reservoir of 88-128
<normalizedToken originalValue="µm">µm</normalizedToken>
length and 30-50
<normalizedToken originalValue="µm">µm</normalizedToken>
width (
<emphasis italics="true" pageId="0" pageNumber="87">in vivo</emphasis>
), empty or with spermatozoa (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hemienchytraeus wuhanensis sp. nov. A anterior body region, anterior 13 segments, lateral view, schematic B brain C spermatheca; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir D pharyngeal glands E oesophageal appendage F sperm funnel." figureDoi="10.3897/zookeys.1015.59019.figure2" httpUri="https://binary.pensoft.net/fig/505241" pageId="0" pageNumber="87">2C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micrographs of Hemienchytraeus wuhanensis sp. nov. A, B, E-I, K, L in vivo C, D, J fixed A brain B dorsal view of oesophageal appendage C lateral view of oesophageal appendage D pharyngeal glands E e pidermal gland cells in II-V ventrally F spermathecae and pharyngeal glands G nephridia in 7 / 8, anteseptale bottom-left H male glandular bulb, slightly everted I dorsal view of clitellum J ventral view of clitellum K sperm funnel L sperm duct and musculature of male copulatory organ Abbreviations: roa, root of oesophageal appendage; boa, branches of oesophageal appendage; oa, oesophageal appendage; sl, secondary pharyngeal gland lobes; dl, dorsal lobes of pharyngeal gland; am, ampulla; ct, connecting tube; ed, ectal duct; er, ental reservoir; hy, hyalocyte; gr, granulocyte; sd, sperm duct. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1015.59019.figure3" httpUri="https://binary.pensoft.net/fig/505242" pageId="0" pageNumber="87">3F</figureCitation>
). One mature egg or 3-4 immature eggs at a time.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="discussion">
<paragraph pageId="0" pageNumber="87">Molecular results</paragraph>
<paragraph pageId="0" pageNumber="87">
COI sequences of five paratype specimens of
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
sp. nov. were successfully acquired and submitted to GenBank with accession numbers. This is the fourth species of
<taxonomicName class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" lsidName="Hemienchytraeus" order="Enchytraeida" pageId="0" pageNumber="87" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="87">Hemienchytraeus</emphasis>
</taxonomicName>
of which DNA sequences are available (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. List of Hemienchytraeus specimens for molecular analyses with collection data and GenBank accession." httpUri="http://table.plazi.org/id/137AF86A35BD3FA3055395C20C9B646A" pageId="0" pageNumber="87" tableUuid="137AF86A35BD3FA3055395C20C9B646A">1</tableCitation>
), the other three being
<taxonomicName lsidName="H. quadratus" pageId="0" pageNumber="87" rank="species" species="quadratus">
<emphasis italics="true" pageId="0" pageNumber="87">H. quadratus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="H. jeojunensis" pageId="0" pageNumber="87" rank="species" species="jeojunensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. jeojunensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Dózsa-Farkas">Dozsa-Farkas</normalizedToken>
&amp; Hong, 2010, all from South Korea. Clear genetic gaps were observed among the four species with high interspecific distances (7.0-21.9%) and low intraspecific distances (0%) among
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
sp. nov. specimens based on the K2P distances of COI sequences (Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Genetic distances of four Hemienchytraeus species (K 2 P)." httpUri="http://table.plazi.org/id/3A3298FF181DB934ACD2C02E9683ECAA" pageId="0" pageNumber="87" tableUuid="3A3298FF181DB934ACD2C02E9683ECAA">2</tableCitation>
). Interestingly, among the three species from South Korea, the one with lowest genetical distance to
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
, is also the one which is most similar morphologically to the new species (see below).
</paragraph>
<caption ID-Table-UUID="137AF86A35BD3FA3055395C20C9B646A" httpUri="http://table.plazi.org/id/137AF86A35BD3FA3055395C20C9B646A" pageId="0" pageNumber="87" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Table 1.</emphasis>
List of
<taxonomicName class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" lsidName="Hemienchytraeus" order="Enchytraeida" pageId="0" pageNumber="87" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="87">Hemienchytraeus</emphasis>
</taxonomicName>
specimens for molecular analyses with collection data and GenBank accession.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="87">
<table pageId="0" pageNumber="87">
<tr pageId="0" pageNumber="87">
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">Species</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">Collection information</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">Specimen ID</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">Accession number</th>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-1
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Luojia, China</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">HBO201904011</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MW000758</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-2
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Luojia, China</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">HBO201904012</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MW000759</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-3
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Shizi, China</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">HBO201904005</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MW000760</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-4
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Shizi, China</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">HBO201904006</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MW000761</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-5
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Shizi, China</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">HBO201904007</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MW000762</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. quadratus" pageId="0" pageNumber="87" rank="species" species="quadratus">
<emphasis italics="true" pageId="0" pageNumber="87">H. quadratus</emphasis>
</taxonomicName>
-1
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252159</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. quadratus" pageId="0" pageNumber="87" rank="species" species="quadratus">
<emphasis italics="true" pageId="0" pageNumber="87">H. quadratus</emphasis>
</taxonomicName>
-2
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">991</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252158</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-1
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1131</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252157</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-2
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1005</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252156</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-3
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1004</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252155</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-4
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1003</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252154</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-5
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1002</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252153</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. jeonjuensis" pageId="0" pageNumber="87" rank="species" species="jeonjuensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. jeonjuensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">Mt Hallasan, Korea</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1115</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">MG252152</td>
</tr>
</table>
</paragraph>
<caption ID-Table-UUID="3A3298FF181DB934ACD2C02E9683ECAA" httpUri="http://table.plazi.org/id/3A3298FF181DB934ACD2C02E9683ECAA" pageId="0" pageNumber="87" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Table 2.</emphasis>
Genetic distances of four
<taxonomicName class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" lsidName="Hemienchytraeus" order="Enchytraeida" pageId="0" pageNumber="87" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="87">Hemienchytraeus</emphasis>
</taxonomicName>
species (K2P).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="87">
<table pageId="0" pageNumber="87">
<tr pageId="0" pageNumber="87">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="1">-</th>
<th colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">1</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">2</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">3</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">4</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">5</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">6</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">7</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">8</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">9</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">10</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">11</th>
<th colspan="1" pageId="0" pageNumber="87" rowspan="1">12</th>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-1
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="2">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="3">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="4">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="5">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="6">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="7">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="8">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="9">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="10">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="11">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="87" rowspan="12">-</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-2
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-3
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-4
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. wuhanensis" pageId="0" pageNumber="87" rank="species" species="wuhanensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. wuhanensis</emphasis>
</taxonomicName>
-5
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. quadratus" pageId="0" pageNumber="87" rank="species" species="quadratus">
<emphasis italics="true" pageId="0" pageNumber="87">H. quadratus</emphasis>
</taxonomicName>
-1
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.200</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.200</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.200</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.200</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.200</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. quadratus" pageId="0" pageNumber="87" rank="species" species="quadratus">
<emphasis italics="true" pageId="0" pageNumber="87">H. quadratus</emphasis>
</taxonomicName>
-2
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.198</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.198</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.198</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.198</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.198</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.007</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
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-1
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-2
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-3
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.000</td>
</tr>
<tr pageId="0" pageNumber="87">
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-4
</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.070</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.216</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.003</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.003</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.003</td>
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<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. koreanus" pageId="0" pageNumber="87" rank="species" species="koreanus">
<emphasis italics="true" pageId="0" pageNumber="87">H. koreanus</emphasis>
</taxonomicName>
-5
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<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.072</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.072</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.072</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.072</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.072</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.213</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.213</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.002</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.002</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.002</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.005</td>
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<td colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName lsidName="H. jeonjuensis" pageId="0" pageNumber="87" rank="species" species="jeonjuensis">
<emphasis italics="true" pageId="0" pageNumber="87">H. jeonjuensis</emphasis>
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<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.219</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.219</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.219</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.219</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.219</td>
<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.189</td>
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<td colspan="1" pageId="0" pageNumber="87" rowspan="1">0.214</td>
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</treatment>
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@ -0,0 +1,82 @@
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Reported from Greece by
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who considers the species closer to
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than to the Pacific form of
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described by
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from the Galapagos Islands. It is likely that the specimens from the Mediterranean belong to neither of the two species but to an undescribed species (
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).
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<paragraph id="EA694B8C5C1A274281B5C0D9D0062DD6" pageId="0" pageNumber="99555">New record for Nigeria</paragraph>
<paragraph id="000C266E5DE0D06864A5F61FC1CB0C1D" pageId="0" pageNumber="99555">New Records: 2, 7</paragraph>
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View file

@ -0,0 +1,125 @@
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(
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[OLM].
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, Pacholatko,
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;
<collectingCountry id="2F807671FFC1B904BDC9E0B3A911FB5D" box="[1260,1387,1220,1243]" name="Vietnam" pageId="21" pageNumber="22">VIETNAM</collectingCountry>
: 1 Ƥ (
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), Pa Co, Mai Chau, Hoa Binh, alt.
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,
<date id="23291021FFC1B904BBC2E090AF19FB78" box="[743,867,1255,1278]" pageId="21" pageNumber="22" value="2001-06-27">27.VI.2001</date>
, ISD-c.
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<emphasis id="65E3EAF3FFC1B904B9E2E17EAD1AFAA4" bold="true" box="[199,352,1289,1314]" pageId="21" pageNumber="22">Distribution.</emphasis>
<collectingCountry id="2F807671FFC1B904B842E17DADDBFAA7" box="[359,417,1290,1313]" name="Laos" pageId="21" pageNumber="22">Laos</collectingCountry>
; northern
<collectingCountry id="2F807671FFC1B904BB37E17EAE0BFAA7" box="[530,625,1289,1313]" name="Vietnam" pageId="21" pageNumber="22">Vietnam</collectingCountry>
(new record).
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<emphasis id="65E3EAF3FFC1B904B9B2E102AC8EFA09" bold="true" box="[151,244,1397,1423]" italics="true" pageId="21" pageNumber="22">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC1B904B821E102AD0EFA09" bold="true" box="[260,372,1397,1423]" italics="true" pageId="21" pageNumber="22">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC1B904B8A3E102AE62FA09" bold="true" box="[390,536,1397,1423]" italics="true" pageId="21" pageNumber="22">mandarinus</emphasis>
</taxonomicName>
de Saussure 1853
</emphasis>
</heading>
</paragraph>
<paragraph id="572836E1FFC1B904B9B2E1EFAD2BFA36" blockId="21.[151,758,1397,1456]" box="[151,337,1432,1456]" pageId="21" pageNumber="22">
(
<figureCitation id="CFAC2A64FFC1B904B9BAE1EFAD32FA36" box="[159,328,1432,1456]" captionStart="FIGURES 75 84" captionStartId="22.[151,269,1885,1909]" captionTargetBox="[191,1377,189,1861]" captionTargetId="figure@22.[191,1396,149,1861]" captionTargetPageId="22" captionText="FIGURES 75 84. Characters of Polistes species of subgenus Polistella occurring in northern Vietnam. 75 80. 3. 81 84. Ƥ. 75, 77, 79. P. mandarinus. 76, 78, 80. P. nipponensis. 81, 83. P. adustus. 82, 84. P. nigerrimus. 75 76, 81. Head, frontal view. 77 78. terminal sterna, lateral view. 79 80. Aedeagus, lateral view. 82. Head, lateral view. 83 84. Metasomal segments 1 and 2 in lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201102/files/figure.png" pageId="21" pageNumber="22">Figs 75, 77, 79</figureCitation>
)
</paragraph>
<paragraph id="572836E1FFC1B904B9B2E195A953FA71" blockId="21.[151,1321,1504,1527]" box="[151,1321,1504,1527]" pageId="21" pageNumber="22">
<taxonomicName id="90974D62FFC1B904B9B2E195AD26FA71" box="[151,348,1506,1527]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mandarinus">
<emphasis id="65E3EAF3FFC1B904B9B2E195AD26FA71" box="[151,348,1506,1527]" italics="true" pageId="21" pageNumber="22">Polistes mandarinus</emphasis>
</taxonomicName>
de Saussure 1853: 58, Ƥ,
<typeStatus id="882C8843FFC1B904BB7CE195AEC9FA71" box="[601,691,1506,1527]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
, &quot;Le norde de la
<collectingCountry id="2F807671FFC1B904BA72E195AFE8FA71" box="[855,914,1506,1527]" name="China" pageId="21" pageNumber="22">Chine</collectingCountry>
&quot; [The Natural History Museum, London].
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC1B904B9B2E251A910F90C" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="572836E1FFC1B904B9B2E251A910F90C" blockId="21.[151,1436,1574,1710]" pageId="21" pageNumber="22">
<emphasis id="65E3EAF3FFC1B904B9B2E251ADFAF9B9" bold="true" box="[151,384,1574,1599]" pageId="21" pageNumber="22">Material examined.</emphasis>
<collectingCountry id="2F807671FFC1B904B8A2E250AD9EF9B9" box="[391,484,1575,1599]" name="China" pageId="21" pageNumber="22">CHINA</collectingCountry>
: 2 Ƥ (
<collectionCode id="3186AE24FFC1B904BB06E250AE04F9B9" LSID="urn:lsid:biocol.org:col:34925" box="[547,638,1575,1599]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH</collectionCode>
), Yen-Ping [1 Ƥ,
<date id="23291021FFC1B904BA1AE250AFAFF9B9" box="[831,981,1575,1599]" pageId="21" pageNumber="22" value="1917-08-19">19.VIII.1917</date>
; 1 Ƥ,
<date id="23291021FFC1B904BD30E250A8E0F9B9" box="[1045,1178,1575,1599]" pageId="21" pageNumber="22" value="1917-09-10">10.IX.1917</date>
];
<collectingCountry id="2F807671FFC1B904BD88E25FA949F9C6" box="[1197,1331,1576,1600]" name="Vietnam" pageId="21" pageNumber="22">VIETNAM</collectingCountry>
:
<date id="23291021FFC1B904BC19E250A9E6F9B9" box="[1340,1436,1575,1599]" pageId="21" pageNumber="22" value="2010-11-03">11 3, 10</date>
Ƥ (
<collectionCode id="3186AE24FFC1B904B99DE23AAC86F9E3" box="[184,252,1613,1637]" pageId="21" pageNumber="22">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFC1B904B82DE23AAD28F9E3" box="[264,338,1613,1637]" pageId="21" pageNumber="22">IUNH</collectionCode>
), Xuan Son NP, Phu Tho, alt.
<quantity id="906F9B04FFC1B904BBB2E239AF68F9E3" box="[663,786,1614,1637]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="21" pageNumber="22" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400500 m</quantity>
,
<date id="23291021FFC1B904BA3BE239AFC7F9E3" box="[798,957,1614,1637]" pageId="21" pageNumber="22" value="2004-04-11" valueMax="2004-04-12" valueMin="2004-04-11">1112.IV.2004</date>
, LN; 3 3 (
<collectionCode id="3186AE24FFC1B904BD1CE239A801F9E3" box="[1081,1147,1614,1637]" pageId="21" pageNumber="22">IEBR</collectionCode>
), Tam Dao NP, Vinh Phuc, alt.
<quantity id="906F9B04FFC1B904B99AE205AD7EF90F" box="[191,260,1650,1673]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="22" unit="m" value="800.0">800 m</quantity>
,
<date id="23291021FFC1B904B82AE205ADFDF90F" box="[271,391,1650,1673]" pageId="21" pageNumber="22" value="2003-07-04">4.VII.2003</date>
, LN; 1 Ƥ (
<collectionCode id="3186AE24FFC1B904B8DFE206AE45F90F" box="[506,575,1649,1673]" pageId="21" pageNumber="22">IEBR</collectionCode>
), Bach Ma NP, Thua Thien Hue, alt.
<quantity id="906F9B04FFC1B904BAF8E206A84EF90F" box="[989,1076,1649,1673]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="1000.0">1000 m</quantity>
,
<date id="23291021FFC1B904BD65E206A8B1F90F" box="[1088,1227,1649,1673]" pageId="21" pageNumber="22" value="2003-07-27">27.VII.2003</date>
, X.L. Truong.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC1B904B9E2E2E2AEC1F92B" box="[199,699,1685,1710]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="572836E1FFC1B904B9E2E2E2AEC1F92B" blockId="21.[151,1436,1574,1710]" box="[199,699,1685,1710]" pageId="21" pageNumber="22">
<emphasis id="65E3EAF3FFC1B904B9E2E2E2AD1AF928" bold="true" box="[199,352,1685,1710]" pageId="21" pageNumber="22">Distribution.</emphasis>
North
<collectingCountry id="2F807671FFC1B904B88AE2E2AD8FF92B" box="[431,501,1685,1709]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
;
<collectingCountry id="2F807671FFC1B904BB24E2E1AE33F92B" box="[513,585,1686,1709]" name="South Korea" pageId="21" pageNumber="22">Korea</collectingCountry>
;
<collectingCountry id="2F807671FFC1B904BB71E2E2AECCF92B" box="[596,694,1685,1709]" name="Vietnam" pageId="21" pageNumber="22">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,120 @@
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<mods:title id="11E346BF22667E40384CEDB4800E6374">Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species</mods:title>
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<mods:namePart id="6535537299A88F3D8D2CBC15DA7D883F">Nguyen, Lien Thi Phuong</mods:namePart>
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<mods:namePart id="D505ACB1B2E495779FEFFA6BCFC85F0E">Kojima, Jun-Ichi</mods:namePart>
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<mods:namePart id="49767B8ED8A508079C95C89A7E557CAC">Saito, Fuki</mods:namePart>
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<treatment id="DF3E87F7FFC1B904B9B2E550ADE9FDDF" ID-DOI="http://doi.org/10.5281/zenodo.5489180" ID-GBIF-Taxon="119419242" ID-Zenodo-Dep="5489180" LSID="urn:lsid:plazi:treatment:DF3E87F7FFC1B904B9B2E550ADE9FDDF" httpUri="http://treatment.plazi.org/id/DF3E87F7FFC1B904B9B2E550ADE9FDDF" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="1F8D656AFFC1B904B9B2E550AD5CFEE4" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="572836E1FFC1B904B9B2E550AEE4FEC7" blockId="21.[151,670,295,354]" box="[151,670,295,321]" pageId="21" pageNumber="22">
<heading id="0C60818DFFC1B904B9B2E550AEE4FEC7" bold="true" box="[151,670,295,321]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="90974D62FFC1B904B9B2E550AEE4FEC7" ID-CoL="6W4Q3" authority="Bingham 1897" authorityName="Bingham" authorityYear="1897" box="[151,670,295,321]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="adustus" subGenus="Polistella">
<emphasis id="65E3EAF3FFC1B904B9B2E550AEE4FEC7" bold="true" box="[151,670,295,321]" pageId="21" pageNumber="22">
<emphasis id="65E3EAF3FFC1B904B9B2E550AC8EFEC7" bold="true" box="[151,244,295,321]" italics="true" pageId="21" pageNumber="22">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC1B904B821E550AD0EFEC7" bold="true" box="[260,372,295,321]" italics="true" pageId="21" pageNumber="22">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC1B904B8A3E550AD99FEC7" bold="true" box="[390,483,295,321]" italics="true" pageId="21" pageNumber="22">adustus</emphasis>
<bibRefCitation id="33064B10FFC1B904B8CFE550AEE4FEC7" author="Bingham" box="[490,670,295,321]" pageId="21" pageNumber="22" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma. Hymenoptera. Vol. I. Wasps and Bees. Taylor and Francis, London. xxix + 579 pp. + 4 pls." type="book" year="1897">Bingham 1897</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="572836E1FFC1B904B9B2E53DAD5CFEE4" blockId="21.[151,670,295,354]" box="[151,294,330,354]" pageId="21" pageNumber="22">
(
<figureCitation id="CFAC2A64FFC1B904B9BAE53DAD64FEE4" box="[159,286,330,354]" captionStart="FIGURES 75 84" captionStartId="22.[151,269,1885,1909]" captionTargetBox="[191,1377,189,1861]" captionTargetId="figure@22.[191,1396,149,1861]" captionTargetPageId="22" captionText="FIGURES 75 84. Characters of Polistes species of subgenus Polistella occurring in northern Vietnam. 75 80. 3. 81 84. Ƥ. 75, 77, 79. P. mandarinus. 76, 78, 80. P. nipponensis. 81, 83. P. adustus. 82, 84. P. nigerrimus. 75 76, 81. Head, frontal view. 77 78. terminal sterna, lateral view. 79 80. Aedeagus, lateral view. 82. Head, lateral view. 83 84. Metasomal segments 1 and 2 in lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201102/files/figure.png" pageId="21" pageNumber="22">Figs 81, 83</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC1B904B9B2E5E5A938FE2E" box="[151,1346,402,424]" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="572836E1FFC1B904B9B2E5E5A938FE2E" blockId="21.[151,1346,402,424]" box="[151,1346,402,424]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="778711CFFFC1B904B9B2E5E5A938FE2E" box="[151,1346,402,424]" pageId="21" pageNumber="22">
<taxonomicName id="90974D62FFC1B904B9B2E5E5AE70FE2E" ID-CoL="6W4Q3" authority="Bingham 1897: 397" authorityName="Bingham" authorityPageNumber="397" authorityYear="1897" box="[151,522,402,424]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="adustus">
<emphasis id="65E3EAF3FFC1B904B9B2E5E5AD40FE2E" box="[151,314,402,424]" italics="true" pageId="21" pageNumber="22">Polistes adustus</emphasis>
<treatmentCitation id="D63610F0FFC1B904B864E5E5AE70FE2E" author="Bingham" box="[321,522,402,424]" page="397" pageId="21" pageNumber="22" year="1897">
<bibRefCitation id="33064B10FFC1B904B864E5E5ADACFE2E" author="Bingham" box="[321,470,402,424]" pageId="21" pageNumber="22" refString="Bingham, C. T. (1897) The Fauna of British India, including Ceylon and Burma. Hymenoptera. Vol. I. Wasps and Bees. Taylor and Francis, London. xxix + 579 pp. + 4 pls." type="book" year="1897">Bingham 1897</bibRefCitation>
: 397
</treatmentCitation>
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFC1B904BB10E5E5AEEAFE2E" box="[565,656,402,424]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
, &quot;Sikkim, up to
<quantity id="906F9B04FFC1B904BA11E5E5AFEFFE2E" box="[820,917,402,424]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="21" pageNumber="22" unit="ft" value="6000.0">6000 feet</quantity>
&quot; [The Natural History Museum, London].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC1B904B9B2E5A2AF7FFD97" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="572836E1FFC1B904B9B2E5A2AF7FFD97" blockId="21.[151,1436,469,601]" pageId="21" pageNumber="22">
<emphasis id="65E3EAF3FFC1B904B9B2E5A2ADFAFE68" bold="true" box="[151,384,469,494]" pageId="21" pageNumber="22">Material examined.</emphasis>
<collectingCountry id="2F807671FFC1B904B8A3E5A1AD87FE68" box="[390,509,470,494]" name="Bhutan" pageId="21" pageNumber="22">BHUTAN</collectingCountry>
: 1 Ƥ (
<collectionCode id="3186AE24FFC1B904BB1EE5A2AEEFFE6B" LSID="urn:lsid:biocol.org:col:34992" box="[571,661,469,493]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22">RMNH</collectionCode>
), Lungtenphu, alt.
<quantity id="906F9B04FFC1B904BA43E5A2AFC6FE6B" box="[870,956,469,493]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="22" unit="m" value="2300.0">2300 m</quantity>
,
<date id="23291021FFC1B904BAEDE5A2A806FE6B" box="[968,1148,469,493]" pageId="21" pageNumber="22" value="1990-07-22" valueMax="1990-07-27" valueMin="1990-07-22">2227.VII.1990</date>
, H.R. Feijen;
<collectingCountry id="2F807671FFC1B904BC3DE5A1A916FE68" box="[1304,1388,470,494]" name="India" pageId="21" pageNumber="22">INDIA</collectingCountry>
: 1 Ƥ (
<collectionCode id="3186AE24FFC1B904B9BBE58DAC88FD97" box="[158,242,506,529]" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="21" pageNumber="22">NIAES</collectionCode>
), Mussoorie, Uttar Pradesh,
<date id="23291021FFC1B904BB04E58DAEEBFD97" box="[545,657,506,529]" pageId="21" pageNumber="22" value="1970-05-24">24.V.1970</date>
, I. Hattori.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC1B904B9E2E66AADE9FDDF" pageId="21" pageNumber="22" type="distribution">
<paragraph id="572836E1FFC1B904B9E2E66AADE9FDDF" blockId="21.[151,1436,469,601]" pageId="21" pageNumber="22">
<emphasis id="65E3EAF3FFC1B904B9E2E66AAD1AFDB0" bold="true" box="[199,352,541,566]" pageId="21" pageNumber="22">Distribution.</emphasis>
<collectingCountry id="2F807671FFC1B904B84FE66AADD2FDB3" box="[362,424,541,565]" name="India" pageId="21" pageNumber="22">India</collectingCountry>
: Himachal Pradesh, Delhi, Uttaranchal, Sikkim, Meghalaya, West Bengal;
<collectingCountry id="2F807671FFC1B904BC2CE66AA920FDB3" box="[1289,1370,541,565]" name="Bhutan" pageId="21" pageNumber="22">Bhutan</collectingCountry>
(new record);
<collectingCountry id="2F807671FFC1B904B9D3E636AD46FDDF" box="[246,316,577,601]" name="Nepal" pageId="21" pageNumber="22">Nepal</collectingCountry>
;
<collectingCountry id="2F807671FFC1B904B86DE636ADF4FDDF" box="[328,398,577,601]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,124 @@
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(
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, Ƥ,
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<emphasis id="65E3EAF3FFC1B904B9B2E741ADF8FCC9" bold="true" box="[151,386,822,847]" pageId="21" pageNumber="22">Material examined.</emphasis>
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,
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, ISD-c; 1 Ƥ (
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), Tam Dao NP, Vinh Phuc, alt.
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,
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, LN.
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<emphasis id="65E3EAF3FFC1B904B9E2E7D2AD1AFC38" bold="true" box="[199,352,933,958]" pageId="21" pageNumber="22">Distribution.</emphasis>
<collectingCountry id="2F807671FFC1B904B842E7D1ADDBFC3B" box="[359,417,934,957]" name="Laos" pageId="21" pageNumber="22">Laos</collectingCountry>
; northern
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(new record).
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(
<figureCitation id="CFAC2A64FFC1B904B9BAE353AD64F8BA" box="[159,286,1828,1852]" captionStart="FIGURES 75 84" captionStartId="22.[151,269,1885,1909]" captionTargetBox="[191,1377,189,1861]" captionTargetId="figure@22.[191,1396,149,1861]" captionTargetPageId="22" captionText="FIGURES 75 84. Characters of Polistes species of subgenus Polistella occurring in northern Vietnam. 75 80. 3. 81 84. Ƥ. 75, 77, 79. P. mandarinus. 76, 78, 80. P. nipponensis. 81, 83. P. adustus. 82, 84. P. nigerrimus. 75 76, 81. Head, frontal view. 77 78. terminal sterna, lateral view. 79 80. Aedeagus, lateral view. 82. Head, lateral view. 83 84. Metasomal segments 1 and 2 in lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201102/files/figure.png" pageId="21" pageNumber="22">Figs 82, 84</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC1B904B9B2E31AA837F825" pageId="21" pageNumber="22" type="reference_group">
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<treatmentCitationGroup id="778711CFFFC1B904B9B2E31AA837F825" infered="true" pageId="21" pageNumber="22">
<taxonomicName id="90974D62FFC1B904B9B2E31AAE26F805" ID-CoL="6VSPW" authority="Gusenleitner 2006: 674" authorityName="Gusenleitner" authorityPageNumber="674" authorityYear="2006" box="[151,604,1901,1923]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="65E3EAF3FFC1B904B9B2E31AAD25F805" box="[151,351,1901,1923]" italics="true" pageId="21" pageNumber="22">Polistes nigerrimus</emphasis>
<treatmentCitation id="D63610F0FFC1B904B84CE31AAE26F805" author="Gusenleitner" box="[361,604,1901,1923]" httpUri="http://treatment.plazi.org/id/72298783FFD3710EFF48FF321CC3FA07" page="674" pageId="21" pageNumber="22" year="2006">
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: 674
</treatmentCitation>
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFC1B904BBABE31AAE93F805" box="[654,745,1901,1923]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
, &quot;
<collectingCountry id="2F807671FFC1B904BBDBE31BAF24F805" box="[766,862,1900,1923]" name="Vietnam" pageId="21" pageNumber="22">Vietnam</collectingCountry>
, Phongsaly [not mentioned on the label], Tam Dao, Vinh Phu [= part of current Vinh Phuc] Province, 27.5-
<date id="23291021FFC1B904BA28E3FAAF13F825" box="[781,873,1933,1955]" pageId="21" pageNumber="22" value="1986-06-02">2.6.1986</date>
, M. Hradsky&quot; [OLM].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="03E86669FFC2B907B9B2E32AAE86F85A" httpUri="https://zenodo.org/record/201102/files/figure.png" pageId="22" pageNumber="23" targetBox="[191,1377,189,1861]" targetPageId="22">
<paragraph id="572836E1FFC2B907B9B2E32AAE86F85A" blockId="22.[151,1436,1885,2012]" pageId="22" pageNumber="23">
<emphasis id="65E3EAF3FFC2B907B9B2E32AAD22F8F3" bold="true" box="[151,344,1885,1909]" pageId="22" pageNumber="23">FIGURES 7584</emphasis>
. Characters of
<taxonomicName id="90974D62FFC2B907B8DAE329AE29F8F3" box="[511,595,1886,1909]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="65E3EAF3FFC2B907B8DAE329AE29F8F3" box="[511,595,1886,1909]" italics="true" pageId="22" pageNumber="23">Polistes</emphasis>
</taxonomicName>
species of subgenus
<emphasis id="65E3EAF3FFC2B907BA1EE329AFDBF8F3" box="[827,929,1886,1909]" italics="true" pageId="22" pageNumber="23">Polistella</emphasis>
occurring in northern Vietnam. 7580. 3. 81 84. Ƥ. 75, 77, 79.
<taxonomicName id="90974D62FFC2B907B87EE3F7AD8FF811" box="[347,501,1920,1943]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mandarinus">
<emphasis id="65E3EAF3FFC2B907B87EE3F7AD8FF811" box="[347,501,1920,1943]" italics="true" pageId="22" pageNumber="23">P. mandarinus</emphasis>
</taxonomicName>
. 76, 78, 80.
<taxonomicName id="90974D62FFC2B907BB5BE3F7AF62F811" box="[638,792,1920,1943]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="65E3EAF3FFC2B907BB5BE3F7AF62F811" box="[638,792,1920,1943]" italics="true" pageId="22" pageNumber="23">P. nipponensis</emphasis>
</taxonomicName>
. 81, 83.
<taxonomicName id="90974D62FFC2B907BA5DE3F7AF9FF811" box="[888,997,1920,1943]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="adustus">
<emphasis id="65E3EAF3FFC2B907BA5DE3F7AF9FF811" box="[888,997,1920,1943]" italics="true" pageId="22" pageNumber="23">P. adustus</emphasis>
</taxonomicName>
. 82, 84.
<taxonomicName id="90974D62FFC2B907BD60E3F7A8AFF811" box="[1093,1237,1920,1943]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="65E3EAF3FFC2B907BD60E3F7A8AFF811" box="[1093,1237,1920,1943]" italics="true" pageId="22" pageNumber="23">P. nigerrimus</emphasis>
</taxonomicName>
. 7576, 81. Head, frontal view. 7778. terminal sterna, lateral view. 7980. Aedeagus, lateral view. 82. Head, lateral view. 8384. Metasomal segments 1 and 2 in lateral view. Scale 1 mm.
</paragraph>
</caption>
<subSubSection id="1F8D656AFFC3B906B9B2E4ECADB6FF7D" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="572836E1FFC3B906B9B2E4ECADB6FF7D" blockId="23.[151,1436,155,252]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E4ECADFBFF32" bold="true" box="[151,385,155,180]" pageId="23" pageNumber="24">Material examined.</emphasis>
<collectingCountry id="2F807671FFC3B906B8ACE4EBAE75FF32" box="[393,527,156,180]" name="Vietnam" pageId="23" pageNumber="24">VIETNAM</collectingCountry>
: 1 Ƥ (
<typeStatus id="882C8843FFC3B906BB79E4ECAEB8FF35" box="[604,706,155,179]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, OLM), Tam Dao, Vinh Phu,
<date id="23291021FFC3B906BD30E4ECA8B4FF35" box="[1045,1230,155,179]" pageId="23" pageNumber="24" value="1986-05-27" valueMax="1986-06-02" valueMin="1986-05-27">27.V2.VI.1986</date>
, M. Hradsky; 2 Ƥ (
<collectionCode id="3186AE24FFC3B906B9BAE4C8AC9EFF51" box="[159,228,191,215]" pageId="23" pageNumber="24">IEBR</collectionCode>
), Lung Cao, Ba Thuoc, Thanh Hoa, alt.
<quantity id="906F9B04FFC3B906BB80E4C8AE94FF51" box="[677,750,191,215]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="24" unit="m" value="500.0">500 m</quantity>
,
<date id="23291021FFC3B906BBDFE4C8AF06FF51" box="[762,892,191,215]" pageId="23" pageNumber="24" value="2003-06-12">12.VI.2003</date>
, X.H. Le.
<emphasis id="65E3EAF3FFC3B906B9E2E494AD1AFF7A" bold="true" box="[199,352,227,252]" pageId="23" pageNumber="24">Distribution.</emphasis>
<collectingCountry id="2F807671FFC3B906B842E494ADB2FF7D" box="[359,456,227,251]" name="Vietnam" pageId="23" pageNumber="24">Vietnam</collectingCountry>
.
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<emphasis id="65E3EAF3FFC3B906B9B2E042AC8EFBC9" bold="true" box="[151,244,1077,1103]" italics="true" pageId="23" pageNumber="24">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC3B906B821E042AD0EFBC9" bold="true" box="[260,372,1077,1103]" italics="true" pageId="23" pageNumber="24">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC3B906B8A3E042AE2FFBC9" bold="true" box="[390,597,1077,1103]" italics="true" pageId="23" pageNumber="24">quadricingulatus</emphasis>
Guisenleitner 2006
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<emphasis id="65E3EAF3FFC3B906B9B2E00CADE0FB17" box="[151,410,1147,1169]" italics="true" pageId="23" pageNumber="24">Polistes quadricingulatus</emphasis>
Guisenleitner 2006: 683
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, Ƥ,
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, N-India [OLM].
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<emphasis id="65E3EAF3FFC3B906B9B2E0CAAD4AFB50" bold="true" box="[151,304,1213,1238]" pageId="23" pageNumber="24">Distribution.</emphasis>
<collectingCountry id="2F807671FFC3B906B812E0C9AD0FFB50" box="[311,373,1214,1238]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
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<treatment id="DF3E87F7FFC3B906B9B2E15DAE50FA4D" ID-DOI="http://doi.org/10.5281/zenodo.5489196" ID-GBIF-Taxon="119419238" ID-Zenodo-Dep="5489196" LSID="urn:lsid:plazi:treatment:DF3E87F7FFC3B906B9B2E15DAE50FA4D" httpUri="http://treatment.plazi.org/id/DF3E87F7FFC3B906B9B2E15DAE50FA4D" lastPageNumber="24" pageId="23" pageNumber="24">
<subSubSection id="1F8D656AFFC3B906B9B2E15DAEA6FAC2" box="[151,732,1322,1348]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="572836E1FFC3B906B9B2E15DAEA6FAC2" blockId="23.[151,732,1322,1348]" box="[151,732,1322,1348]" pageId="23" pageNumber="24">
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<emphasis id="65E3EAF3FFC3B906B9B2E15DAEA6FAC2" bold="true" box="[151,732,1322,1348]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E15DAC8EFAC2" bold="true" box="[151,244,1322,1348]" italics="true" pageId="23" pageNumber="24">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC3B906B821E15DAD0EFAC2" bold="true" box="[260,372,1322,1348]" italics="true" pageId="23" pageNumber="24">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC3B906B8A3E15DAD93FAC2" bold="true" box="[390,489,1322,1348]" italics="true" pageId="23" pageNumber="24">rubellus</emphasis>
Guisenleitner 2006
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9B2E106A80DFA01" box="[151,1143,1393,1415]" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="572836E1FFC3B906B9B2E106A80DFA01" blockId="23.[151,1143,1393,1415]" box="[151,1143,1393,1415]" pageId="23" pageNumber="24">
<taxonomicName id="90974D62FFC3B906B9B2E106AE41FA01" ID-CoL="4KYH4" authority="Guisenleitner 2006: 684" authorityName="Guisenleitner" authorityPageNumber="684" authorityYear="2006" box="[151,571,1393,1415]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="rubellus">
<emphasis id="65E3EAF3FFC3B906B9B2E106AD3AFA01" box="[151,320,1393,1415]" italics="true" pageId="23" pageNumber="24">Polistes rubellus</emphasis>
Guisenleitner 2006: 684
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFC3B906BB43E106AEBBFA01" box="[614,705,1393,1415]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, Yuksam
<quantity id="906F9B04FFC3B906BA01E106AF2DFA01" box="[804,855,1393,1415]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="24" unit="km" value="8.0">8 km</quantity>
S, W-Sikkim,
<collectingCountry id="2F807671FFC3B906BAC8E106A85BFA01" box="[1005,1057,1393,1415]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
[OLM].
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9B2E1C4AE50FA4D" box="[151,554,1459,1484]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="572836E1FFC3B906B9B2E1C4AE50FA4D" blockId="23.[151,554,1459,1484]" box="[151,554,1459,1484]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E1C4AD4AFA4A" bold="true" box="[151,304,1459,1484]" pageId="23" pageNumber="24">Distribution.</emphasis>
<collectingCountry id="2F807671FFC3B906B812E1C4AD0FFA4D" box="[311,373,1459,1483]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
: Sikkim;
<collectingCountry id="2F807671FFC3B906B8FAE1C4AE5CFA4D" box="[479,550,1459,1483]" name="Nepal" pageId="23" pageNumber="24">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,110 @@
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<emphasis id="65E3EAF3FFC3B906B9B2E268AE91F9BF" bold="true" box="[151,747,1567,1593]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E268AC8EF9BF" bold="true" box="[151,244,1567,1593]" italics="true" pageId="23" pageNumber="24">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC3B906B821E268AD0EF9BF" bold="true" box="[260,372,1567,1593]" italics="true" pageId="23" pageNumber="24">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC3B906B8A3E268AD85F9BF" bold="true" box="[390,511,1567,1593]" italics="true" pageId="23" pageNumber="24">santoshae</emphasis>
<bibRefCitation id="33064B10FFC3B906BB23E268AE91F9BF" author="Das" box="[518,747,1567,1593]" pageId="23" pageNumber="24" refString="Das, B. P. &amp; Gupta, V. K. (1989) The Social Wasps of India and the Adjacent Countries (Hymenoptera: Vespidae). Oriental Insects Monograph 11. The Association for the Study of Oriental Insects, Gainesville, 292 pp." type="book" year="1989">
Das
<emphasis id="65E3EAF3FFC3B906BB19E268AE2BF9BF" bold="true" box="[572,593,1567,1593]" italics="true" pageId="23" pageNumber="24">et</emphasis>
Gupta 1989
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9B2E211A9EFF9FA" box="[151,1429,1638,1660]" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="572836E1FFC3B906B9B2E211A9EFF9FA" blockId="23.[151,1429,1638,1660]" box="[151,1429,1638,1660]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="778711CFFFC3B906B9B2E211A9EFF9FA" box="[151,1429,1638,1660]" pageId="23" pageNumber="24">
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<emphasis id="65E3EAF3FFC3B906B9B2E211AD25F9FA" box="[151,351,1638,1660]" italics="true" pageId="23" pageNumber="24">Polistes shantoshae</emphasis>
<treatmentCitation id="D63610F0FFC3B906B840E210AE33F9FA" author="Das" box="[357,585,1639,1660]" page="72" pageId="23" pageNumber="24" year="1989">
<bibRefCitation id="33064B10FFC3B906B840E210AE59F9FA" author="Das" box="[357,547,1639,1660]" pageId="23" pageNumber="24" refString="Das, B. P. &amp; Gupta, V. K. (1989) The Social Wasps of India and the Adjacent Countries (Hymenoptera: Vespidae). Oriental Insects Monograph 11. The Association for the Study of Oriental Insects, Gainesville, 292 pp." type="book" year="1989">Das &amp; Gupta 1989</bibRefCitation>
: 72
</treatmentCitation>
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFC3B906BB51E211AEB5F9FA" box="[628,719,1638,1660]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, Shillong,
<quantity id="906F9B04FFC3B906BA1EE211AFF0F9FA" box="[827,906,1638,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="23" pageNumber="24" unit="m" value="1524.0">1524 m</quantity>
, Meghalaya [Zoological Survey of
<collectingCountry id="2F807671FFC3B906BDCAE211A95CF9FA" box="[1263,1318,1638,1660]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
, Kolikata].
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="1F8D656AFFC3B906B9B2E2DBA841F961" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="572836E1FFC3B906B9B2E2DBA841F961" blockId="23.[151,1338,1708,1768]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E2DBAD0FF942" bold="true" box="[151,373,1708,1732]" pageId="23" pageNumber="24">Material examined.</emphasis>
<collectingCountry id="2F807671FFC3B906B859E2DBAD81F945" box="[380,507,1708,1731]" name="Vietnam" pageId="23" pageNumber="24">VIETNAM</collectingCountry>
: 1 Ƥ (
<collectionCode id="3186AE24FFC3B906BB64E2DBAEF9F945" box="[577,643,1708,1731]" pageId="23" pageNumber="24">IEBR</collectionCode>
), Ngoc Linh Mt., Quang Nam, alt.
<quantity id="906F9B04FFC3B906BADCE2DBA836F945" box="[1017,1100,1708,1731]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="24" unit="m" value="1500.0">1500 m</quantity>
, X.H. Le,
<date id="23291021FFC3B906BD9EE2DBA94CF945" box="[1211,1334,1708,1731]" pageId="23" pageNumber="24" value="2004-03-31">31.III.2004</date>
.
<emphasis id="65E3EAF3FFC3B906B9E2E2B8AD23F96E" bold="true" box="[199,345,1743,1768]" pageId="23" pageNumber="24">Distribution</emphasis>
.
<collectingCountry id="2F807671FFC3B906B842E2B8ADDFF961" box="[359,421,1743,1767]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
: Sikkim, Meghalaya;
<collectingCountry id="2F807671FFC3B906BBBCE2B8AE9AF961" box="[665,736,1743,1767]" name="Nepal" pageId="23" pageNumber="24">Nepal</collectingCountry>
; central
<collectingCountry id="2F807671FFC3B906BA1BE2B8AFE7F961" box="[830,925,1743,1767]" name="Vietnam" pageId="23" pageNumber="24">Vietnam</collectingCountry>
(new record).
</paragraph>
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</treatment>
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View file

@ -0,0 +1,166 @@
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<treatment id="DF3E87F7FFC3B906B9B2E538AF7EFD6D" ID-DOI="http://doi.org/10.5281/zenodo.5489190" ID-GBIF-Taxon="119419236" ID-Zenodo-Dep="5489190" LSID="urn:lsid:plazi:treatment:DF3E87F7FFC3B906B9B2E538AF7EFD6D" httpUri="http://treatment.plazi.org/id/DF3E87F7FFC3B906B9B2E538AF7EFD6D" lastPageNumber="24" pageId="23" pageNumber="24">
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<heading id="0C60818DFFC3B906B9B2E538AEDCFEEF" bold="true" box="[151,678,335,361]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<taxonomicName id="90974D62FFC3B906B9B2E538AEDCFEEF" ID-CoL="4KYF9" authority="Perez 1905" authorityName="Perez" authorityYear="1905" box="[151,678,335,361]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nipponensis" subGenus="Polistella">
<emphasis id="65E3EAF3FFC3B906B9B2E538AEDCFEEF" bold="true" box="[151,678,335,361]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E538AC8EFEEF" bold="true" box="[151,244,335,361]" italics="true" pageId="23" pageNumber="24">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC3B906B821E538AD0EFEEF" bold="true" box="[260,372,335,361]" italics="true" pageId="23" pageNumber="24">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC3B906B8A3E538AE6DFEEF" bold="true" box="[390,535,335,361]" italics="true" pageId="23" pageNumber="24">nipponensis</emphasis>
<bibRefCitation id="33064B10FFC3B906BB3BE538AEDCFEEF" author="Perez" box="[542,678,335,361]" pageId="23" pageNumber="24" refString="Perez, J. (1905) Hymenopteres recueillis dans le Japon central par M. Harmand, Ministre Plenipotentiaire de France a Tokio. Bulletin du Museum National d'Histoire Naturelle, Paris, 1905, 23 - 41, 79 - 87, 148 - 158." type="journal article" year="1905">Pérez 1905</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="572836E1FFC3B906B9B2E505AD2BFE0C" blockId="23.[151,678,335,394]" box="[151,337,370,394]" pageId="23" pageNumber="24">
(
<figureCitation id="CFAC2A64FFC3B906B9BAE505AD32FE0C" box="[159,328,370,394]" captionStart="FIGURES 75 84" captionStartId="22.[151,269,1885,1909]" captionTargetBox="[191,1377,189,1861]" captionTargetId="figure@22.[191,1396,149,1861]" captionTargetPageId="22" captionText="FIGURES 75 84. Characters of Polistes species of subgenus Polistella occurring in northern Vietnam. 75 80. 3. 81 84. Ƥ. 75, 77, 79. P. mandarinus. 76, 78, 80. P. nipponensis. 81, 83. P. adustus. 82, 84. P. nigerrimus. 75 76, 81. Head, frontal view. 77 78. terminal sterna, lateral view. 79 80. Aedeagus, lateral view. 82. Head, lateral view. 83 84. Metasomal segments 1 and 2 in lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201102/files/figure.png" pageId="23" pageNumber="24">Figs 76, 78, 80</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9B2E5CDA814FD8B" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="572836E1FFC3B906B9B2E5CDA814FD8B" blockId="23.[151,1436,442,526]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="778711CFFFC3B906B9B2E5CDA814FD8B" pageId="23" pageNumber="24">
<taxonomicName id="90974D62FFC3B906B9B2E5CDAE7EFE56" ID-CoL="4KYF9" authority="Perez 1905: 82" authorityName="Perez" authorityPageNumber="82" authorityYear="1905" box="[151,516,442,464]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="65E3EAF3FFC3B906B9B2E5CDAD1CFE56" box="[151,358,442,464]" italics="true" pageId="23" pageNumber="24">Polistes nipponensis</emphasis>
<treatmentCitation id="D63610F0FFC3B906B849E5CDAE7EFE56" author="Perez" box="[364,516,442,464]" page="82" pageId="23" pageNumber="24" year="1905">
<bibRefCitation id="33064B10FFC3B906B849E5CDAD9BFE56" author="Perez" box="[364,481,442,464]" pageId="23" pageNumber="24" refString="Perez, J. (1905) Hymenopteres recueillis dans le Japon central par M. Harmand, Ministre Plenipotentiaire de France a Tokio. Bulletin du Museum National d'Histoire Naturelle, Paris, 1905, 23 - 41, 79 - 87, 148 - 158." type="journal article" year="1905">Pérez 1905</bibRefCitation>
: 82
</treatmentCitation>
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFC3B906BB15E5CDAEF1FE56" box="[560,651,442,464]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, Yokohama,
<collectingCountry id="2F807671FFC3B906BA2BE5CCAF32FE56" box="[782,840,443,464]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
[Musésum National dHistoire Naturelle, Paris].
<taxonomicName id="90974D62FFC3B906B9B2E5AEAE61FE69" ID-CoL="4KYKW" authority="Sonan 1937: 169" authorityName="Sonan" authorityPageNumber="169" authorityYear="1937" box="[151,539,473,495]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="yamanakai">
<emphasis id="65E3EAF3FFC3B906B9B2E5AEAD1AFE69" box="[151,352,473,495]" italics="true" pageId="23" pageNumber="24">Polistes yamanakai</emphasis>
<treatmentCitation id="D63610F0FFC3B906B84CE5AEAE61FE69" author="Sonan" box="[361,539,473,495]" page="169" pageId="23" pageNumber="24" year="1937">
<bibRefCitation id="33064B10FFC3B906B84CE5AEAD93FE68" author="Sonan" box="[361,489,473,494]" pageId="23" pageNumber="24" refString="Sonan J. (1937) Two new species and one new genus of Hymenoptera. Transactions of the Natural History Society of Formosa, 27, 169 - 174." type="journal article" year="1937">Sonan 1937</bibRefCitation>
: 169
</treatmentCitation>
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFC3B906BB68E5AEAED2FE69" box="[589,680,473,495]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, Tokyo,
<collectingCountry id="2F807671FFC3B906BA23E5ADAF3AFE69" box="[774,832,474,495]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
[
<collectingCountry id="2F807671FFC3B906BA6BE5AEAFE1FE69" box="[846,923,473,495]" name="Taiwan" pageId="23" pageNumber="24">Taiwan</collectingCountry>
Agricultural Research Institute, Wufeng]. Junior subjective synonym of
<taxonomicName id="90974D62FFC3B906B897E580AE80FD8B" ID-CoL="4KYF9" authority="Perez 1905" authorityName="Perez" authorityYear="1905" box="[434,762,503,526]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="65E3EAF3FFC3B906B897E580AEF8FD88" box="[434,642,503,526]" italics="true" pageId="23" pageNumber="24">Polistes nipponensis</emphasis>
<bibRefCitation id="33064B10FFC3B906BBADE580AE80FD8B" author="Perez" box="[648,762,503,525]" pageId="23" pageNumber="24" refString="Perez, J. (1905) Hymenopteres recueillis dans le Japon central par M. Harmand, Ministre Plenipotentiaire de France a Tokio. Bulletin du Museum National d'Histoire Naturelle, Paris, 1905, 23 - 41, 79 - 87, 148 - 158." type="journal article" year="1905">Pérez 1905</bibRefCitation>
</taxonomicName>
according to
<treatmentCitation id="D63610F0FFC3B906BAA1E580A810FD8B" author="Kojima" box="[900,1130,503,526]" page="60" pageId="23" pageNumber="24" year="2011">
<bibRefCitation id="33064B10FFC3B906BAA1E580A83EFD8B" author="Kojima" box="[900,1092,503,526]" pageId="23" pageNumber="24" refString="Kojima, J., Saito, F. &amp; Nguyen, L. T. P. (2011) On the species-group taxa of Taiwanese social wasps (Hymenoptera: Vespidae) described and / or treated by J. Sonan. Zootaxa, 2920, 42 - 64." type="journal article" year="2011">
Kojima
<emphasis id="65E3EAF3FFC3B906BAF3E58EA878FD8B" box="[982,1026,503,526]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
. 2011
</bibRefCitation>
: 60
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9B2E64AAF95FD41" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="572836E1FFC3B906B9B2E64AAF95FD41" blockId="23.[151,1436,573,748]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E64AAD06FDD3" bold="true" box="[151,380,573,597]" pageId="23" pageNumber="24">Material examined.</emphasis>
<collectingCountry id="2F807671FFC3B906B8ACE64AADA4FDD2" box="[393,478,573,596]" name="Japan" pageId="23" pageNumber="24">JAPAN</collectingCountry>
: Daigo, Ibaraki, [1 3 (
<collectionCode id="3186AE24FFC3B906BBD8E64AAF3FFDD2" box="[765,837,573,596]" pageId="23" pageNumber="24">IUNH</collectionCode>
),
<date id="23291021FFC3B906BA7BE64AAF96FDD2" box="[862,1004,573,596]" pageId="23" pageNumber="24" value="1978-08-14">14.VIII.1978</date>
, JK; 2 Ƥ (
<collectionCode id="3186AE24FFC3B906BDA5E64AA8B2FDD2" box="[1152,1224,573,596]" pageId="23" pageNumber="24">IUNH</collectionCode>
),
<date id="23291021FFC3B906BDC4E64AA927FDD2" box="[1249,1373,573,596]" pageId="23" pageNumber="24" value="2000-09-10">10.IX.2000</date>
, JK].
<collectingCountry id="2F807671FFC3B906B9B2E613AD6CFDFD" box="[151,278,612,635]" name="Vietnam" pageId="23" pageNumber="24">VIETNAM</collectingCountry>
: 1 3, 2 Ƥ (
<collectionCode id="3186AE24FFC3B906B8B6E613ADAFFDFD" box="[403,469,612,635]" pageId="23" pageNumber="24">IEBR</collectionCode>
), Tam Dao NP, Vinh Phuc, alt.
<quantity id="906F9B04FFC3B906BA01E613AF12FDFD" box="[804,872,612,635]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="24" unit="m" value="800.0">800 m</quantity>
[1 3, 1 Ƥ,
<date id="23291021FFC3B906BAC6E613A821FDFD" box="[995,1115,612,635]" pageId="23" pageNumber="24" value="2003-07-04">4.VII.2003</date>
; 1 Ƥ,
<date id="23291021FFC3B906BDB9E613A96FFDFD" box="[1180,1301,612,635]" pageId="23" pageNumber="24" value="2003-05-12">12.V. 2003</date>
], LN; 1 3, 1 Ƥ (
<collectionCode id="3186AE24FFC3B906B99AE6FDAD7BFD27" box="[191,257,650,673]" pageId="23" pageNumber="24">IEBR</collectionCode>
), Cat Ba NP, Hai Phong,
<date id="23291021FFC3B906BB28E6FDAEC3FD27" box="[525,697,650,673]" pageId="23" pageNumber="24" value="2003-07-15" valueMax="2003-07-18" valueMin="2003-07-15">1518.VII.2003</date>
, LN; 1 Ƥ (
<collectionCode id="3186AE24FFC3B906BA17E6FDAF0EFD27" box="[818,884,650,673]" pageId="23" pageNumber="24">IEBR</collectionCode>
), Lac Thinh, Yen Thuy, Hoa Binh,
<date id="23291021FFC3B906BDCDE6FDA91AFD27" box="[1256,1376,650,673]" pageId="23" pageNumber="24" value="2002-04-30">30.IV.2002</date>
, D.L. Khuat; 2 Ƥ (
<collectionCode id="3186AE24FFC3B906B806E6C7AD1EFD41" box="[291,356,688,711]" pageId="23" pageNumber="24">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFC3B906B84BE6C7ADCFFD41" box="[366,437,688,711]" pageId="23" pageNumber="24">IUNH</collectionCode>
), Cuc Phuong NP, Ninh Binh,
<date id="23291021FFC3B906BBDFE6C7AF19FD41" box="[762,867,688,711]" pageId="23" pageNumber="24" value="2002-05-07">7.V. 2002</date>
, V.T. Hoang.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9E2E6A4AF7EFD6D" box="[199,772,723,748]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="572836E1FFC3B906B9E2E6A4AF7EFD6D" blockId="23.[151,1436,573,748]" box="[199,772,723,748]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9E2E6A4AD1AFD6A" bold="true" box="[199,352,723,748]" pageId="23" pageNumber="24">Distribution.</emphasis>
<collectingCountry id="2F807671FFC3B906B842E6A4ADD7FD6D" box="[359,429,723,747]" name="China" pageId="23" pageNumber="24">China</collectingCountry>
;
<collectingCountry id="2F807671FFC3B906B89CE6A3AD86FD6D" box="[441,508,724,747]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
;
<collectingCountry id="2F807671FFC3B906BB22E6A4AE1CFD6D" box="[519,614,723,747]" name="Vietnam" pageId="23" pageNumber="24">Vietnam</collectingCountry>
(new record).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,85 @@
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<heading id="0C60818DFFC3B906B9B2E748AEB7FCDF" bold="true" box="[151,717,831,857]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="65E3EAF3FFC3B906B9B2E748AEB7FCDF" bold="true" box="[151,717,831,857]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E748AC8EFCDF" bold="true" box="[151,244,831,857]" italics="true" pageId="23" pageNumber="24">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC3B906B821E748AD0EFCDF" bold="true" box="[260,372,831,857]" italics="true" pageId="23" pageNumber="24">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC3B906B8A3E748ADA1FCDF" bold="true" box="[390,475,831,857]" italics="true" pageId="23" pageNumber="24">opacus</emphasis>
Guisenleitner 2006
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9B2E7F1A85EFC1A" box="[151,1060,902,924]" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="572836E1FFC3B906B9B2E7F1A85EFC1A" blockId="23.[151,1060,902,924]" box="[151,1060,902,924]" pageId="23" pageNumber="24">
<taxonomicName id="90974D62FFC3B906B9B2E7F1AE4AFC1A" ID-CoL="4KYFR" authority="Guisenleitner 2006: 681" authorityName="Guisenleitner" authorityPageNumber="681" authorityYear="2006" box="[151,560,902,924]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="65E3EAF3FFC3B906B9B2E7F1AD4FFC1A" box="[151,309,902,924]" italics="true" pageId="23" pageNumber="24">Polistes opacus</emphasis>
Guisenleitner 2006: 681
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFC3B906BB7EE7F1AECCFC1A" box="[603,694,902,924]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, Paykong, W-Sikkim,
<collectingCountry id="2F807671FFC3B906BABFE7F1AFB4FC1A" box="[922,974,902,924]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
[OLM].
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC3B906B9B2E7BFAE09FC67" box="[151,627,968,993]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="572836E1FFC3B906B9B2E7BFAE09FC67" blockId="23.[151,627,968,993]" box="[151,627,968,993]" pageId="23" pageNumber="24">
<emphasis id="65E3EAF3FFC3B906B9B2E7BFAD4AFC67" bold="true" box="[151,304,968,993]" pageId="23" pageNumber="24">Distribution.</emphasis>
<collectingCountry id="2F807671FFC3B906B812E7BEAD0FFC67" box="[311,373,969,993]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
: Sikkim, West Bengal.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,277 @@
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<mods:title id="A499477A2F8E756A6ABDF2DAA491A962">Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species</mods:title>
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<mods:namePart id="58A052C9AC6B0655B0A9B3086AB072C1">Nguyen, Lien Thi Phuong</mods:namePart>
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<mods:namePart id="4F0AC776ABE6E8602DA90599599FCA91">Kojima, Jun-Ichi</mods:namePart>
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<mods:date id="2093F4542BADF39854C9B29D14248E84">2011</mods:date>
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<treatment id="DF3E87F7FFC4B903B9B2E0B8AF79FEE2" ID-DOI="http://doi.org/10.5281/zenodo.5489176" ID-GBIF-Taxon="119419235" ID-Zenodo-Dep="5489176" LSID="urn:lsid:plazi:treatment:DF3E87F7FFC4B903B9B2E0B8AF79FEE2" httpUri="http://treatment.plazi.org/id/DF3E87F7FFC4B903B9B2E0B8AF79FEE2" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="1F8D656AFFC4B901B9B2E0B8AD2AFA8C" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="572836E1FFC4B901B9B2E0B8AF0FFB6F" blockId="16.[151,885,1231,1290]" box="[151,885,1231,1257]" pageId="16" pageNumber="17">
<heading id="0C60818DFFC4B901B9B2E0B8AF0FFB6F" bold="true" box="[151,885,1231,1257]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="65E3EAF3FFC4B901B9B2E0B8AF0FFB6F" bold="true" box="[151,885,1231,1257]" pageId="16" pageNumber="17">
<taxonomicName id="90974D62FFC4B901B9B2E0B8AF70FB6F" authority="Nguyen, Kojima &amp; Saito" authorityName="Nguyen, Kojima &amp; Saito" box="[151,778,1231,1257]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gilvus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFC4B901B9B2E0B8AC8EFB6F" bold="true" box="[151,244,1231,1257]" italics="true" pageId="16" pageNumber="17">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC4B901B821E0B8AD0EFB6F" bold="true" box="[260,372,1231,1257]" italics="true" pageId="16" pageNumber="17">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC4B901B8A3E0B8ADB4FB6F" bold="true" box="[390,462,1231,1257]" italics="true" pageId="16" pageNumber="17">gilvus</emphasis>
Nguyen, Kojima &amp; Saito
</taxonomicName>
,
<taxonomicNameLabel id="7ED05788FFC4B901BA32E0B8AF0FFB6F" box="[791,885,1231,1257]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="572836E1FFC4B901B9B2E085AD2AFA8C" blockId="16.[151,885,1231,1290]" box="[151,336,1266,1290]" pageId="16" pageNumber="17">
(
<figureCitation id="CFAC2A64FFC4B901B9BAE085AD5AFA8C" box="[159,288,1266,1290]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Figs 5460</figureCitation>
,
<figureCitation id="CFAC2A64FFC4B901B80EE085AD32FA8C" box="[299,328,1266,1290]" captionStart="FIGURES 85 88" captionStartId="26.[151,269,1501,1525]" captionTargetBox="[346,1242,198,1469]" captionTargetId="figure@26.[346,1243,194,1469]" captionTargetPageId="26" captionText="FIGURES 85 88. Nests of Polistes species of subgenus Polistella occurring in northern Vietnam. 85. P. dawnae (Nest # 2009 - DB-P- 01). The thick and thin arrows indicate the position of the primary petiole and the subsidiary petioles, respectively. 86. P. l e p c h a (Nest # VN-P- 2006 - 3). 87. P. reliciniclypeus sp. nov. (Nest # VN-P- 2006 - 13). 88. P. gilvus sp. nov. (Nest # 2008 - SL-P- 1)." httpUri="https://zenodo.org/record/201103/files/figure.png" pageId="16" pageNumber="17">88</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC4B901B9B2E14CAD68FA64" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="572836E1FFC4B901B9B2E14CAD68FA64" blockId="16.[151,1436,1338,2011]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9B2E14CAD3CFAD5" bold="true" box="[151,326,1338,1363]" pageId="16" pageNumber="17">
<typeStatus id="882C8843FFC4B901B9B2E14CACA8FAD5" box="[151,210,1339,1363]" pageId="16" pageNumber="17">Type</typeStatus>
material.
</emphasis>
<typeStatus id="882C8843FFC4B901B86BE14DAD9CFAD4" box="[334,486,1338,1362]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
: Ƥ (
<collectionCode id="3186AE24FFC4B901BB31E14DAE24FAD4" box="[532,606,1338,1362]" pageId="16" pageNumber="17">IUNH</collectionCode>
, on long-term loan from the
<collectionCode id="3186AE24FFC4B901BA8DE14DAF96FAD4" box="[936,1004,1338,1362]" pageId="16" pageNumber="17">IEBR</collectionCode>
), labeled “
<collectingCountry id="2F807671FFC4B901BD41E14DA891FAD4" box="[1124,1259,1338,1362]" name="Vietnam" pageId="16" pageNumber="17">VIETNAM</collectingCountry>
, Co Ma, Thuan Chau, Son La,
<quantity id="906F9B04FFC4B901B818E129ADEEFAF0" box="[317,404,1374,1398]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="17" unit="m" value="1600.0">1600 m</quantity>
, Nest #: 2008-SL-P-1, L.P.T. Nguyen, H.P. Pham,
<date id="23291021FFC4B901BAE2E129A839FAF0" box="[967,1091,1374,1398]" pageId="16" pageNumber="17" value="2008-06-07">07.vi.2008</date>
”.
<typeStatus id="882C8843FFC4B901BD7CE129A882FAF0" box="[1113,1272,1374,1398]" pageId="16" pageNumber="17" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="2F807671FFC4B901BC27E129A91FFAF0" box="[1282,1381,1374,1398]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
:
<emphasis id="65E3EAF3FFC4B901BC4AE129ACC2FA1D" bold="true" pageId="16" pageNumber="17">Son La</emphasis>
: 8 Ƥ (
<collectionCode id="3186AE24FFC4B901B823E1F5AD30FA1C" box="[262,330,1410,1434]" pageId="16" pageNumber="17">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFC4B901B87CE1F4ADD8FA1C" box="[345,418,1411,1434]" pageId="16" pageNumber="17">IUNH</collectionCode>
,
<collectionCode id="3186AE24FFC4B901B895E1F5AE70FA1C" LSID="urn:lsid:biocol.org:col:34925" box="[432,522,1410,1434]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="17">AMNH</collectionCode>
), same data as
<typeStatus id="882C8843FFC4B901BB9AE1F5AF5FFA1C" box="[703,805,1410,1434]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
;
<emphasis id="65E3EAF3FFC4B901BA16E1F4AFECFA1D" bold="true" box="[819,918,1410,1435]" pageId="16" pageNumber="17">Lao Cai</emphasis>
: 1 Ƥ (
<collectionCode id="3186AE24FFC4B901BAC0E1F5A850FA1C" box="[997,1066,1410,1434]" pageId="16" pageNumber="17">IEBR</collectionCode>
), Sa Pa, alt.
<quantity id="906F9B04FFC4B901BDE7E1F5A919FA1C" box="[1218,1379,1410,1434]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="16" pageNumber="17" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">15002000 m</quantity>
,
<date id="23291021FFC4B901BC57E1F5AD76FA38" pageId="16" pageNumber="17" value="2003-05-17" valueMax="2003-05-18" valueMin="2003-05-17">17 18.V.2003</date>
, ISD-c; 1 Ƥ (
<collectionCode id="3186AE24FFC4B901B894E1D1AD89FA38" box="[433,499,1446,1470]" httpUri="http://grbio.org/cool/zeys-ppzt" name="NSM-PV, National Science Museum" pageId="16" pageNumber="17">NSM</collectionCode>
), Mt. Phan Xi Pang, alt.
<quantity id="906F9B04FFC4B901BA07E1D1AFBEFA38" box="[802,964,1446,1470]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.95" metricValueMin="1.75" pageId="16" pageNumber="17" unit="m" value="1850.0" valueMax="1950.0" valueMin="1750.0">17501950 m</quantity>
, Hoang Lien Son Mts.,
<date id="23291021FFC4B901BDC0E1D1A916FA38" box="[1253,1388,1446,1470]" pageId="16" pageNumber="17" value="1995-10-02" valueMax="1995-10-05" valueMin="1995-10-02">25.X.1995</date>
, M. Tomokuni.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC4B901B9E2E199AF6FF910" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="572836E1FFC4B901B9E2E199AF6FF910" blockId="16.[151,1436,1338,2011]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9E2E199AD3BF981" bold="true" box="[199,321,1518,1543]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
This species can be distinguished from other known species of subgenus
<emphasis id="65E3EAF3FFC4B901BD56E199A8A4F980" box="[1139,1246,1518,1542]" italics="true" pageId="16" pageNumber="17">Polistella</emphasis>
by the following combination of characters: female clypeus weakly convex, in lateral view anterior margin smoothly passing into supraclypeal area; propodeum with strongly and sharply raised transverse striae; border between posterior and lateral faces of propodeum marked by blunt angulation; metasomal sternum
<quantity id="906F9B04FFC4B901BAF8E22DA873F9F4" box="[989,1033,1626,1650]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
lateral view square in outline, with corner between anterior and ventral faces bluntly angled.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC4B900B9E2E2D5AE34F8F8" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="572836E1FFC4B901B9E2E2D5AD16F958" blockId="16.[151,1436,1338,2011]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9E2E2D5AD23F93D" bold="true" box="[199,345,1698,1723]" pageId="16" pageNumber="17">Description.</emphasis>
FEMALE. Body length
<quantity id="906F9B04FFC4B901BB48E2D5AE8EF93C" box="[621,756,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="16" pageNumber="17" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.59.5 mm</quantity>
(
<typeStatus id="882C8843FFC4B901BA27E2D5AF10F93C" box="[770,874,1698,1722]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFC4B901BA9CE2D5A869F93C" box="[953,1043,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="16" pageNumber="17" unit="mm" value="8.5">8.5 mm</quantity>
); fore wing length
<quantity id="906F9B04FFC4B901BDC3E2D5A92BF93C" box="[1254,1361,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="16" pageNumber="17" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910 mm</quantity>
(
<typeStatus id="882C8843FFC4B901BC7AE2D5ACB1F959" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFC4B901B83EE2B1AD1AF958" box="[283,352,1734,1758]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="17" unit="mm" value="9.0">9 mm</quantity>
).
</paragraph>
<paragraph id="572836E1FFC4B901B9E2E29DAF55F814" blockId="16.[151,1436,1338,2011]" pageId="16" pageNumber="17">
Structure similar to that of
<taxonomicName id="90974D62FFC4B901B8DAE29CAED6F885" box="[511,684,1771,1795]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFC4B901B8DAE29CAED6F885" box="[511,684,1771,1795]" italics="true" pageId="16" pageNumber="17">Polistes lepcha</emphasis>
</taxonomicName>
, but different as follows: POD:OOD=1:1.9; POD about 1.6 × Od (
<figureCitation id="CFAC2A64FFC4B901B9BAE379AC8FF8A0" box="[159,245,1806,1830]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Fig. 54</figureCitation>
); clypeus in frontal view about 1.1 × as wide as long (
<figureCitation id="CFAC2A64FFC4B901BA43E379AFC6F8A0" box="[870,956,1806,1830]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
), in lateral view weakly convex (
<figureCitation id="CFAC2A64FFC4B901BC1CE379A9F5F8A0" box="[1337,1423,1806,1830]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Fig. 56</figureCitation>
); antennal flagellomere 1 slightly greater than 2.5 × as long as its maximum width (
<figureCitation id="CFAC2A64FFC4B901BD15E345A8FEF8CC" box="[1072,1156,1842,1866]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Fig. 57</figureCitation>
); scutellum convex (
<figureCitation id="CFAC2A64FFC4B901BC4BE345ACCCF8E8" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Fig. 58</figureCitation>
); jugal lobe of hind wing rounded (
<figureCitation id="CFAC2A64FFC4B901BB60E321AEE0F8E8" box="[581,666,1878,1902]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Fig. 59</figureCitation>
); metasomal sternum
<quantity id="906F9B04FFC4B901BAB7E321AFC4F8E8" box="[914,958,1878,1902]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
lateral view square in outline, with corner between anterior and ventral faces bluntly angled (
<figureCitation id="CFAC2A64FFC4B901BBEBE30DAF58F814" box="[718,802,1914,1938]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="16" pageNumber="17">Fig. 60</figureCitation>
).
</paragraph>
<paragraph id="572836E1FFC4B901B9E2E3E9AFC6F830" blockId="16.[151,1436,1338,2011]" box="[199,956,1950,1975]" pageId="16" pageNumber="17">
Body surface sculpture as in
<taxonomicName id="90974D62FFC4B901BB2EE3D7AE0BF830" box="[523,625,1950,1975]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFC4B901BB2EE3D7AE0BF830" box="[523,625,1950,1975]" italics="true" pageId="16" pageNumber="17">P. lepcha</emphasis>
</taxonomicName>
, but propodeal striae weaker.
</paragraph>
<paragraph id="572836E1FFC4B901B9E2E3B4A848F85C" blockId="16.[151,1436,1338,2011]" box="[199,1074,1986,2011]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9E2E3B4AD77F85D" box="[199,269,1987,2011]" italics="true" pageId="16" pageNumber="17">Color.</emphasis>
As in
<emphasis id="65E3EAF3FFC4B901B873E3B3ADC3F85D" box="[342,441,1987,2011]" italics="true" pageId="16" pageNumber="17">P. l e pc h a</emphasis>
except paired small lateral spots on metanotum present.
</paragraph>
<caption id="03E86669FFC5B900B9B2E145AE92FA0F" httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="17" pageNumber="18" targetBox="[209,1396,191,1305]" targetPageId="17">
<paragraph id="572836E1FFC5B900B9B2E145AE92FA0F" blockId="17.[151,1437,1330,1417]" pageId="17" pageNumber="18">
<emphasis id="65E3EAF3FFC5B900B9B2E145AD25FACC" bold="true" box="[151,351,1330,1354]" pageId="17" pageNumber="18">FIGURES 5460.</emphasis>
<taxonomicName id="90974D62FFC5B900B843E145AEF8FACF" box="[358,642,1330,1353]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gilvus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFC5B900B843E145ADC0FACF" box="[358,442,1330,1353]" italics="true" pageId="17" pageNumber="18">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC5B900B8EEE145AE4BFACF" box="[459,561,1330,1353]" italics="true" pageId="17" pageNumber="18">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC5B900BB67E145AEF8FACF" box="[578,642,1330,1353]" italics="true" pageId="17" pageNumber="18">gilvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFC5B900BBAFE145AE9AFACC" box="[650,736,1330,1354]" pageId="17" pageNumber="18" rank="species">
<emphasis id="65E3EAF3FFC5B900BBAFE145AE9AFACC" bold="true" box="[650,736,1330,1354]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
, Ƥ. 54. Ocelli, eye and posterior margin of vertex. 5556. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="572836E1FFC5B900B9E2E1ADADE0FA74" blockId="17.[151,1436,1498,2026]" box="[199,410,1498,1522]" pageId="17" pageNumber="18">MALE. Unknown.</paragraph>
<paragraph id="572836E1FFC5B900B9E2E189AF39F9BC" blockId="17.[151,1436,1498,2026]" pageId="17" pageNumber="18">
<emphasis id="65E3EAF3FFC5B900B9E2E189AD79F990" bold="true" box="[199,259,1534,1558]" pageId="17" pageNumber="18">Nest.</emphasis>
A nest (Nest#: 2008-SL-P-1) collected in Thuan Chau, Son La was examined (
<tableCitation id="1A15035AFFC5B900BD90E189A977F990" box="[1205,1293,1534,1558]" captionStart="TABLE 1" captionStartId="5.[151,235,157,178]" captionTargetBox="[165,1417,218,1135]" captionTargetPageId="5" captionText="TABLE 1. Biological data of examined nests of Polistes species from subgenus Polistella." httpUri="http://table.plazi.org/id/03E86669FFD1B914B9B2E4EAA857FF35" pageId="17" pageNumber="18" tableUuid="03E86669FFD1B914B9B2E4EAA857FF35">Table 1</tableCitation>
); one of its resident females is designated as the
<typeStatus id="882C8843FFC5B900BB10E255AEEDF9BC" box="[565,663,1570,1594]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
of this species.
</paragraph>
<paragraph id="572836E1FFC5B900B9E2E231AE34F8F8" blockId="17.[151,1436,1498,2026]" pageId="17" pageNumber="18">
The nest had only two pupae as immatures, and was attached to a thin branch of a shurub about
<quantity id="906F9B04FFC5B900BC35E231A92BF9D8" box="[1296,1361,1606,1630]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="m" value="1.5">1.5 m</quantity>
above the ground. The nest characters are as follows: Petiole single, terminal,
<quantity id="906F9B04FFC5B900BAE6E21DA861F904" box="[963,1051,1642,1666]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="mm" value="3.5">3.5 mm</quantity>
long, 2 ×
<quantity id="906F9B04FFC5B900BDB4E21DA892F904" box="[1169,1256,1642,1666]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="18" unit="mm" value="2.5">2.5 mm</quantity>
thick, with thin core of plant fibers, enlarged strictly with adult salivary secretion, black and lustrous; salivary coating extended onto substrate and largely onto back of comb. Comb rather tough, pliable in texture, uniformly grayish brown, fan shape in ventral view, ventral surface convex, dorsal surface deeply concave. Cells arranged in regular rows and round at open end, but partly irregularly arranged, where some cells are pentagonal; each cell gradually expanded towards open end,
<quantity id="906F9B04FFC5B900B849E369ADB8F8B0" box="[364,450,1822,1846]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="18" unit="mm" value="2.5">2.5 mm</quantity>
(range 2.33.0 mm; n=10) wide at bottom and
<quantity id="906F9B04FFC5B900BAFCE369A855F8B0" box="[985,1071,1822,1846]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="17" pageNumber="18" unit="mm" value="4.1">4.1 mm</quantity>
(range
<quantity id="906F9B04FFC5B900BDA0E369A975F8B0" box="[1157,1295,1822,1846]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.6" metricValueMin="3.9" pageId="17" pageNumber="18" unit="mm" value="4.25" valueMax="4.6" valueMin="3.9">3.94.6 mm</quantity>
; n=10) wide at open end,
<quantity id="906F9B04FFC5B900B80AE335ADECF8DC" box="[303,406,1858,1882]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="17" pageNumber="18" unit="mm" value="12.6">12.6 mm</quantity>
(range
<quantity id="906F9B04FFC5B900B8D7E335AEE0F8DC" box="[498,666,1858,1882]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" metricValueMax="1.34" metricValueMin="1.1199999999999999" pageId="17" pageNumber="18" unit="mm" value="12.3" valueMax="13.4" valueMin="11.2">11.213.4 mm</quantity>
; n=10) deep; cell wall about
<quantity id="906F9B04FFC5B900BAD4E335A822F8DC" box="[1009,1112,1858,1882]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="mm" value="0.02">0.02 mm</quantity>
thick. Cocoon caps slightly domed beyond rim of cell, pale yellow.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC5B903B9E2E3FDAF1DFE9A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="572836E1FFC5B900B9E2E3FDA9E6F86C" blockId="17.[151,1436,1498,2026]" pageId="17" pageNumber="18">
<emphasis id="65E3EAF3FFC5B900B9E2E3FDAD4AF824" bold="true" box="[199,304,1930,1954]" pageId="17" pageNumber="18">Remark.</emphasis>
Among species of subgenus
<emphasis id="65E3EAF3FFC5B900BB59E3FDAE9DF824" box="[636,743,1930,1954]" italics="true" pageId="17" pageNumber="18">Polistella</emphasis>
occurring in
<collectingCountry id="2F807671FFC5B900BAA5E3FDAF98F824" box="[896,994,1930,1954]" name="Vietnam" pageId="17" pageNumber="18">Vietnam</collectingCountry>
, this species is most similar to
<taxonomicName id="90974D62FFC5B900BC66E3FDAC98F840" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFC5B900BC66E3FDAC98F840" italics="true" pageId="17" pageNumber="18">Polistes lepcha</emphasis>
</taxonomicName>
, but can be easily distinguished from the latter by having the clypeus in lateral view less swollen and smoothly passing into the supraclypeal area (
<figureCitation id="CFAC2A64FFC5B900BBB0E3A5AE9DF86C" box="[661,743,2002,2026]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="17" pageNumber="18">Fig. 56</figureCitation>
) (in
<taxonomicName id="90974D62FFC5B900BA3BE3A4AFFCF86C" box="[798,902,2002,2026]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFC5B900BA3BE3A4AFFCF86C" box="[798,902,2002,2026]" italics="true" pageId="17" pageNumber="18">P. lepcha</emphasis>
</taxonomicName>
, slightly concave in the supraclypeal area; Fig.
</paragraph>
<paragraph id="572836E1FFC6B903B9B2E4E0AF1DFE9A" blockId="18.[151,1436,151,356]" pageId="18" pageNumber="19">
16), propodeal striae weaker, the border between posterior and lateral faces of the propodeum less sharp, and the second metasomal sturnum in lateral view with the anteroventral corner angled, not produced anteriorly (
<figureCitation id="CFAC2A64FFC6B903BC64E4CBA9EEFF52" box="[1345,1428,188,212]" captionStart="FIGURES 54 60" captionStartId="17.[151,269,1330,1354]" captionTargetBox="[209,1396,191,1305]" captionTargetId="figure@17.[191,1396,191,1305]" captionTargetPageId="17" captionText="FIGURES 54 60. Polistes (Polistella) gilvus sp. nov., Ƥ. 54. Ocelli, eye and posterior margin of vertex. 55 56. Head. 55. Frontal view. 56. Lateral view. 57. Right antenna. 58. Mesosoma, lateral view. 59. Jugal lobe of left hind wing. 60. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201099/files/figure.png" pageId="18" pageNumber="19">Fig. 60</figureCitation>
) (in
<taxonomicName id="90974D62FFC6B903B998E496AD59FF7E" box="[189,291,224,248]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFC6B903B998E496AD59FF7E" box="[189,291,224,248]" italics="true" pageId="18" pageNumber="19">P. lepcha</emphasis>
</taxonomicName>
sharply angled, slightly produced anteriorly, Fig. 20). The nest cocoon caps are also different in color, namely pale yellow in this species while pale green in
<emphasis id="65E3EAF3FFC6B903BBDFE572AF26FE9A" box="[762,860,260,284]" italics="true" pageId="18" pageNumber="19">P. l e p c h a</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC6B903B9E2E55FAF79FEE2" pageId="18" pageNumber="19" type="etymology">
<paragraph id="572836E1FFC6B903B9E2E55FAF79FEE2" blockId="18.[151,1436,151,356]" pageId="18" pageNumber="19">
<emphasis id="65E3EAF3FFC6B903B9E2E55FAD34FEC6" bold="true" box="[199,334,296,320]" pageId="18" pageNumber="19">Etymology.</emphasis>
The specific name,
<taxonomicName id="90974D62FFC6B903BB0AE55FAE08FEC6" box="[559,626,296,320]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gilvus">
<emphasis id="65E3EAF3FFC6B903BB0AE55FAE08FEC6" box="[559,626,296,320]" italics="true" pageId="18" pageNumber="19">gilvus</emphasis>
</taxonomicName>
, is a Latin adjective and refers to the pale-yellow cocoon caps.
<emphasis id="65E3EAF3FFC6B903B9E2E53CAD1AFEE2" bold="true" box="[199,352,331,356]" pageId="18" pageNumber="19">Distribution.</emphasis>
Known only from northern
<collectingCountry id="2F807671FFC6B903BBB9E53BAE85FEE2" box="[668,767,332,356]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="28A565182CB5D4CBCD8CA3E9B31374B0">Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species</mods:title>
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<mods:identifier id="D64A1C4AD9FE28AECAB08BB59A9BDE2F" type="CLB-Dataset">46173</mods:identifier>
<mods:identifier id="F6B7D1C6BAB31429C693A606496E8F35" type="DOI">10.5281/zenodo.201094</mods:identifier>
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<treatment id="DF3E87F7FFC6B905B9B2E5C0AD73F888" ID-DOI="http://doi.org/10.5281/zenodo.5489178" ID-GBIF-Taxon="119419232" ID-Zenodo-Dep="5489178" LSID="urn:lsid:plazi:treatment:DF3E87F7FFC6B905B9B2E5C0AD73F888" httpUri="http://treatment.plazi.org/id/DF3E87F7FFC6B905B9B2E5C0AD73F888" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="1F8D656AFFC6B903B9B2E5C0AD5DFE74" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="572836E1FFC6B903B9B2E5C0AFB8FE57" blockId="18.[151,962,439,498]" box="[151,962,439,465]" pageId="18" pageNumber="19">
<heading id="0C60818DFFC6B903B9B2E5C0AFB8FE57" bold="true" box="[151,962,439,465]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="65E3EAF3FFC6B903B9B2E5C0AFB8FE57" bold="true" box="[151,962,439,465]" pageId="18" pageNumber="19">
<taxonomicName id="90974D62FFC6B903B9B2E5C0AF22FE57" authority="Nguyen, Kojima &amp; Saito" authorityName="Nguyen, Kojima &amp; Saito" box="[151,856,439,465]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="clandestinus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFC6B903B9B2E5C0AC8EFE57" bold="true" box="[151,244,439,465]" italics="true" pageId="18" pageNumber="19">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC6B903B821E5C0AD0EFE57" bold="true" box="[260,372,439,465]" italics="true" pageId="18" pageNumber="19">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC6B903B8A3E5C0AE67FE57" bold="true" box="[390,541,439,465]" italics="true" pageId="18" pageNumber="19">clandestinus</emphasis>
Nguyen, Kojima
<emphasis id="65E3EAF3FFC6B903BBDCE5C0AF6AFE57" bold="true" box="[761,784,439,465]" italics="true" pageId="18" pageNumber="19">&amp;</emphasis>
Saito
</taxonomicName>
,
<taxonomicNameLabel id="7ED05788FFC6B903BA40E5C0AFB8FE57" box="[869,962,439,465]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="572836E1FFC6B903B9B2E5ADAD5DFE74" blockId="18.[151,962,439,498]" box="[151,295,474,498]" pageId="18" pageNumber="19">
(
<figureCitation id="CFAC2A64FFC6B903B9BAE5ADAD64FE74" box="[159,286,474,498]" captionStart-0="FIGURES 61 67" captionStart-1="FIGURES 68 74" captionStartId-0="19.[151,269,1617,1641]" captionStartId-1="20.[151,269,1290,1314]" captionTargetBox-0="[191,1396,411,1592]" captionTargetBox-1="[191,1396,195,1266]" captionTargetId-0="figure@19.[191,1396,411,1592]" captionTargetId-1="figure@20.[191,1396,195,1266]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." captionText-1="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri-0="https://zenodo.org/record/201100/files/figure.png" httpUri-1="https://zenodo.org/record/201101/files/figure.png" pageId="18" pageNumber="19">Figs 6174</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC6B903B9B2E654AFB4FD4C" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="572836E1FFC6B903B9B2E654AFB4FD4C" blockId="18.[151,1437,546,2011]" pageId="18" pageNumber="19">
<emphasis id="65E3EAF3FFC6B903B9B2E654AD38FDBD" bold="true" box="[151,322,546,571]" pageId="18" pageNumber="19">
<typeStatus id="882C8843FFC6B903B9B2E654ACAAFDBD" box="[151,208,547,571]" pageId="18" pageNumber="19">Type</typeStatus>
material.
</emphasis>
<typeStatus id="882C8843FFC6B903B86EE655AD99FDBC" box="[331,483,546,570]" pageId="18" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
: Ƥ (
<collectionCode id="3186AE24FFC6B903BB2BE655AE22FDBC" box="[526,600,546,570]" pageId="18" pageNumber="19">IUNH</collectionCode>
, on long-term loan from the
<collectionCode id="3186AE24FFC6B903BABFE655AFA4FDBC" box="[922,990,546,570]" pageId="18" pageNumber="19">IEBR</collectionCode>
), labeled “
<collectingCountry id="2F807671FFC6B903BD77E655A8A2FDBC" box="[1106,1240,546,570]" name="Vietnam" pageId="18" pageNumber="19">VIETNAM</collectingCountry>
, Sa Pa NP office, Lao Cai,
<quantity id="906F9B04FFC6B903B825E631AD22FDD8" box="[256,344,582,606]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="m" value="1600.0">1600 m</quantity>
, L.P.T. Nguyen, F. Saito &amp; J. Kojima,
<date id="23291021FFC6B903BA2AE631AFE1FDD8" box="[783,923,582,606]" pageId="18" pageNumber="19" value="2006-08-22">22.viii.2006</date>
”.
<typeStatus id="882C8843FFC6B903BA97E631A82BFDD8" box="[946,1105,582,606]" pageId="18" pageNumber="19" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="2F807671FFC6B903BD79E631A8C5FDD8" box="[1116,1215,582,606]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
:
<emphasis id="65E3EAF3FFC6B903BDEFE630A951FDD9" bold="true" box="[1226,1323,582,607]" pageId="18" pageNumber="19">Lao Cai</emphasis>
: 5 3, 3 Ƥ (
<collectionCode id="3186AE24FFC6B903B9BAE61DAC99FD04" box="[159,227,618,642]" pageId="18" pageNumber="19">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFC6B903B9CBE61CAD4DFD04" box="[238,311,619,642]" pageId="18" pageNumber="19">IUNH</collectionCode>
,
<collectionCode id="3186AE24FFC6B903B864E61DADE1FD04" LSID="urn:lsid:biocol.org:col:34925" box="[321,411,618,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="19">AMNH</collectionCode>
), same data as
<typeStatus id="882C8843FFC6B903BB64E61DAEDDFD04" box="[577,679,618,642]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
; 3 3, 3 Ƥ (
<collectionCode id="3186AE24FFC6B903BA3BE61DAF18FD04" box="[798,866,618,642]" pageId="18" pageNumber="19">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFC6B903BA48E61DAFCDFD04" box="[877,951,618,642]" pageId="18" pageNumber="19">IUNH</collectionCode>
), alt.
<quantity id="906F9B04FFC6B903BAD6E61DA830FD04" box="[1011,1098,618,642]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="m" value="1600.0">1600 m</quantity>
, Sa Pa Town,
<date id="23291021FFC6B903BDCDE61DA91FFD04" box="[1256,1381,618,642]" pageId="18" pageNumber="19" value="2009-07-08">8.VII.2009</date>
, HP; 1 Ƥ, 2 3 (
<collectionCode id="3186AE24FFC6B903B832E6F9AD26FD20" box="[279,348,654,678]" pageId="18" pageNumber="19">IEBR</collectionCode>
), Xin Chai, Sa Pa, alt.
<quantity id="906F9B04FFC6B903BB53E6F9AEA8FD20" box="[630,722,654,678]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="18" pageNumber="19" unit="m" value="1350.0">1350 m</quantity>
,
<date id="23291021FFC6B903BBC6E6F9AF1AFD20" box="[739,864,654,678]" pageId="18" pageNumber="19" value="2009-07-06">6.VII.2009</date>
, HP; 1 Ƥ (
<collectionCode id="3186AE24FFC6B903BAC2E6F9A856FD20" box="[999,1068,654,678]" pageId="18" pageNumber="19">IEBR</collectionCode>
), Hoang Lien NP, alt.
<quantity id="906F9B04FFC6B903BC1EE6F9A9EDFD20" box="[1339,1431,654,678]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="18" pageNumber="19" unit="m" value="1660.0">1660 m</quantity>
,
<date id="23291021FFC6B903B9B2E6C5AD6EFD4C" box="[151,276,690,714]" pageId="18" pageNumber="19" value="2009-07-06">6.VII.2009</date>
, HP; 1 Ƥ (
<collectionCode id="3186AE24FFC6B903B8ADE6C5ADB7FD4C" box="[392,461,690,714]" pageId="18" pageNumber="19">IEBR</collectionCode>
), Cat Cat, Sa Pa, alt.
<quantity id="906F9B04FFC6B903BB9FE6C5AF6BFD4C" box="[698,785,690,714]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="18" pageNumber="19" unit="m" value="1450.0">1450 m</quantity>
,
<date id="23291021FFC6B903BA38E6C5AFE0FD4C" box="[797,922,690,714]" pageId="18" pageNumber="19" value="2009-07-07">7.VII.2009</date>
, HP.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC6B903B9E2E6A1A96FFCF8" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="572836E1FFC6B903B9E2E6A1A96FFCF8" blockId="18.[151,1437,546,2011]" pageId="18" pageNumber="19">
<emphasis id="65E3EAF3FFC6B903B9E2E6A1AD3BFD69" bold="true" box="[199,321,726,751]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
This species can be distinguished from other known species of subgenus
<emphasis id="65E3EAF3FFC6B903BD56E6A1A8A4FD68" box="[1139,1246,726,750]" italics="true" pageId="18" pageNumber="19">Polistella</emphasis>
by the following combination of characters: scutellum entirely flat, in lateral view dorsal margin smoothly passing into either mesoscutum and metanotum; posterior face of propodeum with shallow median excavation about 0.3 × the maximum width of propodeum, in lateral view slope down in nearly straight line; metasomal sternum
<quantity id="906F9B04FFC6B903BDC1E735A96AFCDC" box="[1252,1296,834,858]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="2.0">2 in</quantity>
lateral view basally abruptly swollen ventrally, then ventral margin broadly rounded; male clypeus as wide as high.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC6B905B9E2E7FDA82FFA68" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph id="572836E1FFC6B903B9E2E7FDADEAFC40" blockId="18.[151,1437,546,2011]" pageId="18" pageNumber="19">
<emphasis id="65E3EAF3FFC6B903B9E2E7FDAD23FC25" bold="true" box="[199,345,906,931]" pageId="18" pageNumber="19">Description.</emphasis>
FEMALE. Body length
<quantity id="906F9B04FFC6B903BB49E7FDAEAFFC24" box="[620,725,906,930]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="18" pageNumber="19" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
(
<typeStatus id="882C8843FFC6B903BBC6E7FDAF31FC24" box="[739,843,906,930]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFC6B903BABFE7FDAF89FC24" box="[922,1011,906,930]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="18" pageNumber="19" unit="mm" value="9.5">9.5 mm</quantity>
); fore wing length
<quantity id="906F9B04FFC6B903BDE0E7FDA92BFC24" box="[1221,1361,906,930]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="18" pageNumber="19" unit="mm" value="10.75" valueMax="11.5" valueMin="10.0">1011.5 mm</quantity>
(
<typeStatus id="882C8843FFC6B903BC7AE7FDACB1FC41" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFC6B903B83EE7D9ADFEFC40" box="[283,388,942,966]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="19" unit="mm" value="10.5">10.5 mm</quantity>
).
</paragraph>
<paragraph id="572836E1FFC6B903B9E2E7A5A859FA8C" blockId="18.[151,1437,546,2011]" pageId="18" pageNumber="19">
Structure as in
<taxonomicName id="90974D62FFC6B903B84AE7A4AE09FC6D" box="[367,627,979,1003]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFC6B903B84AE7A4AE09FC6D" box="[367,627,979,1003]" italics="true" pageId="18" pageNumber="19">Polistes reliciniclypeus</emphasis>
</taxonomicName>
, but differing as follows: POD:OOD=1:2.1 (
<figureCitation id="CFAC2A64FFC6B903BD4EE7A5A8C4FC6C" box="[1131,1214,978,1002]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 61</figureCitation>
); inner eye margins in frontal view about 1.1 × further apart from each other at vertex than at clypeus (
<figureCitation id="CFAC2A64FFC6B903BD11E781A8F3FB88" box="[1076,1161,1014,1038]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 62</figureCitation>
); anterior tentorial pit as far from eye margin as from antennal socket; clypeus in frontal view about 1.1 × as wide as high (
<figureCitation id="CFAC2A64FFC6B903BDF6E06DA95CFBB4" box="[1235,1318,1050,1074]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 62</figureCitation>
), in lateral view convex, smoothly curved anteriorly (
<figureCitation id="CFAC2A64FFC6B903BB5CE049AEB4FBD0" box="[633,718,1086,1110]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 63</figureCitation>
); antennal flagellomere 1 nearly 3 × as long as its apical width (
<figureCitation id="CFAC2A64FFC6B903B9BAE015AC82FBFC" box="[159,248,1122,1146]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 64</figureCitation>
); scutellum entirely flat, in lateral view dorsal margin smoothly passing into either mesoscutum and metanotum (
<figureCitation id="CFAC2A64FFC6B903B801E0F1AD02FB18" box="[292,376,1158,1182]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 65</figureCitation>
); jugal lobe of hind wing rounded (
<figureCitation id="CFAC2A64FFC6B903BA21E0F1AF23FB18" box="[772,857,1158,1182]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 66</figureCitation>
); posterior face of propodeum with shallow median excavation about 0.3 × maximum width of propodeum, in lateral view slope down in nearly straight line (
<figureCitation id="CFAC2A64FFC6B903BC67E0DDA9EEFB44" box="[1346,1428,1194,1218]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 65</figureCitation>
) (slightly convex in
<taxonomicName id="90974D62FFC6B903B854E0A7AE4BFB60" box="[369,561,1230,1255]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFC6B903B854E0A7AE4BFB60" box="[369,561,1230,1255]" italics="true" pageId="18" pageNumber="19">P. reliciniclypeus</emphasis>
</taxonomicName>
;
<figureCitation id="CFAC2A64FFC6B903BB62E0B9AEE0FB60" box="[583,666,1230,1254]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="18" pageNumber="19">Fig. 31</figureCitation>
); metasomal sternum
<quantity id="906F9B04FFC6B903BAB7E0B9AFC4FB60" box="[914,958,1230,1254]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="2.0">2 in</quantity>
lateral view basally abruptly swollen ventrally in anterior half to two-third, then ventral margin broadly rounded (
<figureCitation id="CFAC2A64FFC6B903BAE7E085A86DFA8C" box="[962,1047,1266,1290]" captionStart="FIGURES 61 67" captionStartId="19.[151,269,1617,1641]" captionTargetBox="[191,1396,411,1592]" captionTargetId="figure@19.[191,1396,411,1592]" captionTargetPageId="19" captionText="FIGURES 61 67. Polistes (Polistella) clandestinus sp. nov., Ƥ. 61. Ocelli, eye and posterior margin of vertex. 62 63. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="18" pageNumber="19">Fig. 67</figureCitation>
).
</paragraph>
<paragraph id="572836E1FFC6B903B9E2E161AF25F9AC" blockId="18.[151,1437,546,2011]" pageId="18" pageNumber="19">
Body surface sculpture as in
<taxonomicName id="90974D62FFC6B903BB31E16FAEACFAA8" box="[532,726,1302,1327]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFC6B903BB31E16FAEACFAA8" box="[532,726,1302,1327]" italics="true" pageId="18" pageNumber="19">P. reliciniclypeus</emphasis>
</taxonomicName>
, but dorsal half of clypeus covered with fine pale pubescence and ventral half with scattered deep large punctures, each baering sharply pointed golden bristle; mandible with scattered shallow punctures; frons and vertex to level of posterior ocelli with moderately close fine punctures; vertex behind posterior ocelli and gena with sparse fine punctures (punctures in ventral third of gena scaratted and deep), with space between punctures smooth; mesoscutum with dense fine punctures in addition to scattered deep flat-bottomed punctures; scutellum and metanotum with scattered moderately deep punctures; ventral metapleuron smooth, without punctures; propodeum with distinct transverse striae, and punctures between striae on lateral faces, interpuncture spaces weakly raised to interconnect striae.
</paragraph>
<paragraph id="572836E1FFC6B903B9E2E241AC9CF8CC" blockId="18.[151,1437,546,2011]" pageId="18" pageNumber="19">
<emphasis id="65E3EAF3FFC6B903B9E2E241AD77F9C8" box="[199,269,1590,1614]" italics="true" pageId="18" pageNumber="19">Color.</emphasis>
Black, with following parts yellow:clypeus except black basal and lateral margins, broad band along inner eye margin extending from bottom of frons to middle of eye emargination, line in dorsal part of gena along posterodorsal margin of eye, most ventral part of gena, narrow band along pronotal carina with branches running along posterodosal margin of pronotum, scutellum except for anterior margin, paired anterolateral spots on metanotum, scrobal spot on mesepisternum, paired lateral oval spot on posterior face of propodeum, propodeal valvula, narrow apical band on metasomal terga 15, entire visible part of tergum 4. Legs black, dorsal surface of femora and ventral surface of tibiae, all tarsus largely reddish brown. Wings semitransparent, pale brown; veins dark brown.
</paragraph>
<paragraph id="572836E1FFC6B903B9E2E320AFCEF8E8" blockId="18.[151,1437,546,2011]" box="[199,948,1878,1903]" pageId="18" pageNumber="19">
MALE. Body length 9.010.0 mm; fore wing length 9.0
<quantity id="906F9B04FFC6B903BA6EE321AFCBF8E8" box="[843,945,1878,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="19" unit="mm" value="10.5">10.5 mm</quantity>
.
</paragraph>
<paragraph id="572836E1FFC6B902B9E2E30DA87FFEE2" blockId="18.[151,1437,546,2011]" lastBlockId="19.[151,1436,151,356]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Structure as in female, but head in frontal view about 1.5 × as wide as high, eye strongly swollen laterally, inner eye margins about 1.4 × as further apart from each other at vertex as at clypeus (
<figureCitation id="CFAC2A64FFC6B903BD78E3E9A8C8F830" box="[1117,1202,1950,1974]" captionStart="FIGURES 68 74" captionStartId="20.[151,269,1290,1314]" captionTargetBox="[191,1396,195,1266]" captionTargetId="figure@20.[191,1396,195,1266]" captionTargetPageId="20" captionText="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="18" pageNumber="19">Fig. 68</figureCitation>
); gena narrow, about 0.3 × as wide as eye in lateral view, with weakly raised blunt ridge running along posterior margin of eye; clypeus in frontal view about 1.2 × as wide as high (
<figureCitation id="CFAC2A64FFC7B902BB84E4E0AE8DFF29" box="[673,759,151,175]" captionStart="FIGURES 68 74" captionStartId="20.[151,269,1290,1314]" captionTargetBox="[191,1396,195,1266]" captionTargetId="figure@20.[191,1396,195,1266]" captionTargetPageId="20" captionText="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="19" pageNumber="20">Fig. 68</figureCitation>
), barely produced medioventrally, with apical margin very weakly and broadly rounded, in lateral view slightly convex dorsally, slightly bent backwards ventrally, and covered with dense short golden hairs; antennal scape short and thick, about 2.8 × as long as its maximum width; flagellum slender, flagellomere 1 as long as length of flagellomeres 2 and 3 combined, flagellomeres 2 and 3 each longer than wide; terminal flagellomere elongate, slightly curved, about 2 × as long as its basal width (
<figureCitation id="CFAC2A64FFC7B902BC1CE550A9EAFEB9" box="[1337,1424,295,319]" captionStart="FIGURES 68 74" captionStartId="20.[151,269,1290,1314]" captionTargetBox="[191,1396,195,1266]" captionTargetId="figure@20.[191,1396,195,1266]" captionTargetPageId="20" captionText="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="19" pageNumber="20">Fig. 69</figureCitation>
). Metasomal sternum 7 depressed medially (
<figureCitation id="CFAC2A64FFC7B902BB52E53BAEB1FEE2" box="[631,715,332,356]" captionStart="FIGURES 68 74" captionStartId="20.[151,269,1290,1314]" captionTargetBox="[191,1396,195,1266]" captionTargetId="figure@20.[191,1396,195,1266]" captionTargetPageId="20" captionText="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="19" pageNumber="20">Fig. 70</figureCitation>
), without tubercle (
<figureCitation id="CFAC2A64FFC7B902BA80E53BAF82FEE2" box="[933,1016,332,356]" captionStart="FIGURES 68 74" captionStartId="20.[151,269,1290,1314]" captionTargetBox="[191,1396,195,1266]" captionTargetId="figure@20.[191,1396,195,1266]" captionTargetPageId="20" captionText="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="19" pageNumber="20">Fig. 71</figureCitation>
).
</paragraph>
<caption id="03E86669FFC7B902B9B2E226AF6AF92B" httpUri="https://zenodo.org/record/201100/files/figure.png" pageId="19" pageNumber="20" targetBox="[191,1396,411,1592]" targetPageId="19">
<paragraph id="572836E1FFC7B902B9B2E226AF6AF92B" blockId="19.[151,1437,1617,1709]" pageId="19" pageNumber="20">
<emphasis id="65E3EAF3FFC7B902B9B2E226AD24F9EF" bold="true" box="[151,350,1617,1641]" pageId="19" pageNumber="20">FIGURES 6167.</emphasis>
<taxonomicName id="90974D62FFC7B902B843E226AEB2F9EE" box="[358,712,1617,1640]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="clandestinus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFC7B902B843E226ADC1F9EE" box="[358,443,1617,1640]" italics="true" pageId="19" pageNumber="20">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC7B902B8E9E226AE48F9EE" box="[460,562,1617,1640]" italics="true" pageId="19" pageNumber="20">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC7B902BB66E226AEB2F9EE" box="[579,712,1617,1640]" italics="true" pageId="19" pageNumber="20">clandestinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFC7B902BBF5E226AF5CF9EF" box="[720,806,1617,1641]" pageId="19" pageNumber="20" rank="species">
<emphasis id="65E3EAF3FFC7B902BBF5E226AF5CF9EF" bold="true" box="[720,806,1617,1641]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
, Ƥ. 61. Ocelli, eye and posterior margin of vertex. 6263. Head. 62. Frontal view. 63. Lateral view. 64. Right antenna. 65. Mesosoma, lateral view. 66. Jugal lobe of left hind wing. 67. Metasomal segments 1 and 2, lateral view. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="572836E1FFC7B902B9E2E376AFB3F803" blockId="19.[151,1436,1793,2033]" pageId="19" pageNumber="20">
<emphasis id="65E3EAF3FFC7B902B9E2E376AD77F89F" box="[199,269,1793,1817]" italics="true" pageId="19" pageNumber="20">Color.</emphasis>
Similar to female, but with more abundant yellow markings as follows: clypeus entirely, mandible except for small black basal spot, ventral half of frons, narrow short band on gena along posterior margin of eye, antennal scape beneath, paired anterolateral spots on metanotum. Antennal flagellum reddish-brown to dark yellow beneath. Paired lateral yellow spots on propodeum much reduced in size.
</paragraph>
<paragraph id="572836E1FFC7B902B9E2E3E6ACBEF877" blockId="19.[151,1436,1793,2033]" pageId="19" pageNumber="20">
<emphasis id="65E3EAF3FFC7B902B9E2E3E6AD42F82F" box="[199,312,1937,1961]" italics="true" pageId="19" pageNumber="20">Genitalia.</emphasis>
Similar to that of
<taxonomicName id="90974D62FFC7B902BB28E3E6AF69F82F" box="[525,787,1937,1961]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="reliciniclypeus" status="sp. nov.">
<emphasis id="65E3EAF3FFC7B902BB28E3E6AF69F82F" box="[525,787,1937,1961]" italics="true" pageId="19" pageNumber="20">Polistes reliciniclypeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFC7B902BA3EE3E5AF0DF82C" box="[795,887,1938,1962]" pageId="19" pageNumber="20" rank="species">
<emphasis id="65E3EAF3FFC7B902BA3EE3E5AF0DF82C" bold="true" box="[795,887,1938,1962]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
except: the digitus distinctly swollen near base, narrowed in apical half (
<figureCitation id="CFAC2A64FFC7B902B883E3C2AD80F84B" box="[422,506,1973,1997]" captionStart="FIGURES 68 74" captionStartId="20.[151,269,1290,1314]" captionTargetBox="[191,1396,195,1266]" captionTargetId="figure@20.[191,1396,195,1266]" captionTargetPageId="20" captionText="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="19" pageNumber="20">Fig. 72</figureCitation>
); aedeagus with proxomoventral corner prominently produced proximally (
<figureCitation id="CFAC2A64FFC7B902BC64E3C2ACCDF877" captionStart="FIGURES 68 74" captionStartId="20.[151,269,1290,1314]" captionTargetBox="[191,1396,195,1266]" captionTargetId="figure@20.[191,1396,195,1266]" captionTargetPageId="20" captionText="FIGURES 68 74. Polistes (Polistella) clandestinus sp. nov., 3. 68. Head, frontal view. 69. Left antenna. 70 71. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 73 74. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="19" pageNumber="20">Figs 73, 74</figureCitation>
).
</paragraph>
<caption id="03E86669FFC0B905B9B2E17DAF2DFAE7" httpUri="https://zenodo.org/record/201101/files/figure.png" pageId="20" pageNumber="21" targetBox="[191,1396,195,1266]" targetPageId="20">
<paragraph id="572836E1FFC0B905B9B2E17DAF2DFAE7" blockId="20.[151,1437,1289,1377]" pageId="20" pageNumber="21">
<emphasis id="65E3EAF3FFC0B905B9B2E17DAD19FAA4" bold="true" box="[151,355,1290,1314]" pageId="20" pageNumber="21">FIGURES 6874.</emphasis>
<taxonomicName id="90974D62FFC0B905B84BE17DAEA3FAA7" box="[366,729,1290,1313]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="clandestinus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFC0B905B84BE17DADB9FAA7" box="[366,451,1290,1313]" italics="true" pageId="20" pageNumber="21">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFC0B905B8FDE17DAE44FAA7" box="[472,574,1290,1313]" italics="true" pageId="20" pageNumber="21">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFC0B905BB76E17DAEA3FAA7" box="[595,729,1290,1313]" italics="true" pageId="20" pageNumber="21">clandestinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFC0B905BBC1E17DAF45FAA4" box="[740,831,1290,1314]" pageId="20" pageNumber="21" rank="species">
<emphasis id="65E3EAF3FFC0B905BBC1E17DAF45FAA4" bold="true" box="[740,831,1290,1314]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
, 3. 68. Head, frontal view. 69. Left antenna. 7071. Terminal sterna. 70. Ventral view. 71. Apical part, lateral view. 72. Inner aspect of paramere with digitus and volsella. 7374. Aedeagus. 73. Ventral view. 74. Lateral view. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="572836E1FFC0B905B9E2E1C4A82FFA68" blockId="20.[151,1437,1458,1806]" pageId="20" pageNumber="21">
<emphasis id="65E3EAF3FFC0B905B9E2E1C4AD79FA4D" bold="true" box="[199,259,1459,1483]" pageId="20" pageNumber="21">Nest.</emphasis>
Although we were unable to collect the nest of this species, we saw frequent traffic of the wasps coming in and out through a small hole respectively of an electric meter and a window frame.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC0B905B9E2E18CAE51F940" pageId="20" pageNumber="21" type="discussion">
<paragraph id="572836E1FFC0B905B9E2E18CAE51F940" blockId="20.[151,1437,1458,1806]" pageId="20" pageNumber="21">
<emphasis id="65E3EAF3FFC0B905B9E2E18CAD4AF995" bold="true" box="[199,304,1531,1555]" pageId="20" pageNumber="21">Remark.</emphasis>
Among species of subgenus
<emphasis id="65E3EAF3FFC0B905BB59E18DAE9DF994" box="[636,743,1530,1554]" italics="true" pageId="20" pageNumber="21">Polistella</emphasis>
occurring in
<collectingCountry id="2F807671FFC0B905BAA5E18DAF98F994" box="[896,994,1530,1554]" name="Vietnam" pageId="20" pageNumber="21">Vietnam</collectingCountry>
, this species is most similar to
<taxonomicName id="90974D62FFC0B905BC66E18DAC94F9B0" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFC0B905BC66E18DAC94F9B0" italics="true" pageId="20" pageNumber="21">Polistes dawnae</emphasis>
</taxonomicName>
, but it can be easily distinguished from the later by having the deeper median excavation of the propodeum, the metasomal tergum 1 with the anterodorsal corner between the anterior and dorsal faces more angulated, the second metasomal sternum in lateral view swollen ventrally in anterior half to two-third (one-third in
<taxonomicName id="90974D62FFC0B905BCA2E210AC94F924" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFC0B905BCA2E210AC94F924" italics="true" pageId="20" pageNumber="21">P. dawnae</emphasis>
</taxonomicName>
), the paired lateral yellow spots on propodeum small (large in
<taxonomicName id="90974D62FFC0B905BA8EE2FCA867F924" box="[939,1053,1674,1698]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFC0B905BA8EE2FCA867F924" box="[939,1053,1674,1698]" italics="true" pageId="20" pageNumber="21">P. dawnae</emphasis>
</taxonomicName>
), and the male clypeus wider than high (as wide as high in
<taxonomicName id="90974D62FFC0B905B88DE2D8AE61F940" box="[424,539,1710,1734]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFC0B905B88DE2D8AE61F940" box="[424,539,1710,1734]" italics="true" pageId="20" pageNumber="21">P. dawnae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC0B905B9E2E2A4AD73F888" pageId="20" pageNumber="21" type="etymology">
<paragraph id="572836E1FFC0B905B9E2E2A4AD73F888" blockId="20.[151,1437,1458,1806]" pageId="20" pageNumber="21">
<emphasis id="65E3EAF3FFC0B905B9E2E2A4AD34F96D" bold="true" box="[199,334,1747,1771]" pageId="20" pageNumber="21">Etymology.</emphasis>
The specific name,
<taxonomicName id="90974D62FFC0B905BB17E2A5AEC4F96C" box="[562,702,1746,1770]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="clandestinus">
<emphasis id="65E3EAF3FFC0B905BB17E2A5AEC4F96C" box="[562,702,1746,1770]" italics="true" pageId="20" pageNumber="21">clandestinus</emphasis>
</taxonomicName>
, is a Latin adjective and refers to nesting of this species in a hidden place.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0D2E672F89705D4C0496615EEC7C8B73">Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species</mods:title>
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<mods:namePart id="1ED189252CC072BB02F3E219AC1B38D6">Nguyen, Lien Thi Phuong</mods:namePart>
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<mods:namePart id="EC33D36A459210FA090B4E3224AF5ED8">Saito, Fuki</mods:namePart>
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<mods:date id="7A5C8222A19AE7927FA53E7508102C5B">2011</mods:date>
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<treatment id="DF3E87F7FFD2B919B9B2E4E0AF98F927" ID-DOI="http://doi.org/10.5281/zenodo.5489168" ID-GBIF-Taxon="119419233" ID-Zenodo-Dep="5489168" LSID="urn:lsid:plazi:treatment:DF3E87F7FFD2B919B9B2E4E0AF98F927" httpUri="http://treatment.plazi.org/id/DF3E87F7FFD2B919B9B2E4E0AF98F927" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="1F8D656AFFD2B917B9B2E4E0AD2AFF54" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="572836E1FFD2B917B9B2E4E0AEEEFF37" blockId="6.[151,660,151,210]" box="[151,660,151,177]" pageId="6" pageNumber="7">
<heading id="0C60818DFFD2B917B9B2E4E0AEEEFF37" bold="true" box="[151,660,151,177]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="90974D62FFD2B917B9B2E4E0AEEEFF37" ID-CoL="4KYDM" authority="Cameron 1900" authorityName="Cameron" authorityYear="1900" box="[151,660,151,177]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lepcha" subGenus="Polistella">
<emphasis id="65E3EAF3FFD2B917B9B2E4E0AEEEFF37" bold="true" box="[151,660,151,177]" pageId="6" pageNumber="7">
<emphasis id="65E3EAF3FFD2B917B9B2E4E0AC8EFF37" bold="true" box="[151,244,151,177]" italics="true" pageId="6" pageNumber="7">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFD2B917B821E4E0AD0EFF37" bold="true" box="[260,372,151,177]" italics="true" pageId="6" pageNumber="7">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFD2B917B8A3E4E0ADAFFF37" bold="true" box="[390,469,151,177]" italics="true" pageId="6" pageNumber="7">lepcha</emphasis>
<bibRefCitation id="33064B10FFD2B917B8F8E4E0AEEEFF37" author="Cameron" box="[477,660,151,177]" pageId="6" pageNumber="7" refString="Cameron, P. (1900) Description of new genera and species of Hymenoptera. A. New species of Vespidae from India and Japan. Annals and Magazine of Natural History, ser. 7, 6, 410 - 419, 495 - 506, 530 - 537." type="journal article" year="1900">
<collectingCountry id="2F807671FFD2B917B8F8E4E0AE28FF37" box="[477,594,151,177]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
1900
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="572836E1FFD2B917B9B2E4CDAD2AFF54" blockId="6.[151,660,151,210]" box="[151,336,186,210]" pageId="6" pageNumber="7">(Figs 1426, 86)</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD2B917B9B2E575AEF5FE42" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="572836E1FFD2B917B9B2E575A84EFEB1" blockId="6.[151,1311,258,311]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="778711CFFFD2B917B9B2E575A84EFEB1" pageId="6" pageNumber="7">
<taxonomicName id="90974D62FFD2B917B9B2E575AE5FFE9E" ID-CoL="6VSPV" authority="Cameron 1900: 415" authorityName="Cameron" authorityPageNumber="415" authorityYear="1900" box="[151,549,258,280]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="khasianus">
<emphasis id="65E3EAF3FFD2B917B9B2E575AD28FE9E" box="[151,338,258,280]" italics="true" pageId="6" pageNumber="7">Polistes khasianus</emphasis>
<treatmentCitation id="D63610F0FFD2B917B87CE574AE5FFE9E" author="Cameron" box="[345,549,258,280]" page="415" pageId="6" pageNumber="7" year="1900">
<bibRefCitation id="33064B10FFD2B917B87CE574AD89FE9E" author="Cameron" box="[345,499,259,280]" pageId="6" pageNumber="7" refString="Cameron, P. (1900) Description of new genera and species of Hymenoptera. A. New species of Vespidae from India and Japan. Annals and Magazine of Natural History, ser. 7, 6, 410 - 419, 495 - 506, 530 - 537." type="journal article" year="1900">
<collectingCountry id="2F807671FFD2B917B87CE574ADCCFE9E" box="[345,438,259,280]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
1900
</bibRefCitation>
: 415
</treatmentCitation>
</taxonomicName>
, Ƥ, &quot;Khasia Hills, Coll. Rothney&quot;, nom. praeocc., non
<treatmentCitation id="D63610F0FFD2B917BD6AE574A961FE9E" author="Cameron" box="[1103,1307,259,280]" page="413" pageId="6" pageNumber="7" year="1900">
<bibRefCitation id="33064B10FFD2B917BD6AE574A892FE9E" author="Cameron" box="[1103,1256,259,280]" pageId="6" pageNumber="7" refString="Cameron, P. (1900) Description of new genera and species of Hymenoptera. A. New species of Vespidae from India and Japan. Annals and Magazine of Natural History, ser. 7, 6, 410 - 419, 495 - 506, 530 - 537." type="journal article" year="1900">
<collectingCountry id="2F807671FFD2B917BD6AE574A8D6FE9E" box="[1103,1196,259,280]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
1900
</bibRefCitation>
: 413
</treatmentCitation>
.
<taxonomicName id="90974D62FFD2B917B9B2E556AE79FEB1" ID-CoL="4KYDM" authority="Cameron 1900: 506" authorityName="Cameron" authorityPageNumber="506" authorityYear="1900" box="[151,515,289,311]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFD2B917B9B2E556AD4BFEB1" box="[151,305,289,311]" italics="true" pageId="6" pageNumber="7">Polistes lepcha</emphasis>
<treatmentCitation id="D63610F0FFD2B917B812E555AE79FEB1" author="Cameron" box="[311,515,289,311]" page="506" pageId="6" pageNumber="7" year="1900">
<bibRefCitation id="33064B10FFD2B917B812E555ADAAFEB1" author="Cameron" box="[311,464,290,311]" pageId="6" pageNumber="7" refString="Cameron, P. (1900) Description of new genera and species of Hymenoptera. A. New species of Vespidae from India and Japan. Annals and Magazine of Natural History, ser. 7, 6, 410 - 419, 495 - 506, 530 - 537." type="journal article" year="1900">
<collectingCountry id="2F807671FFD2B917B812E555ADEEFEB1" box="[311,404,290,311]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
1900
</bibRefCitation>
: 506
</treatmentCitation>
</taxonomicName>
. Replacement name for
<taxonomicName id="90974D62FFD2B917BBDDE556A84AFEB1" ID-CoL="6VSPV" authority="Cameron 1900: 415" authorityName="Cameron" authorityPageNumber="415" authorityYear="1900" box="[760,1072,289,311]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="khasianus">
<emphasis id="65E3EAF3FFD2B917BBDDE556AF27FEB1" box="[760,861,289,311]" italics="true" pageId="6" pageNumber="7">khasianus</emphasis>
<treatmentCitation id="D63610F0FFD2B917BA41E555A84AFEB1" author="Cameron" box="[868,1072,289,311]" page="415" pageId="6" pageNumber="7" year="1900">
<bibRefCitation id="33064B10FFD2B917BA41E555AF87FEB1" author="Cameron" box="[868,1021,290,311]" pageId="6" pageNumber="7" refString="Cameron, P. (1900) Description of new genera and species of Hymenoptera. A. New species of Vespidae from India and Japan. Annals and Magazine of Natural History, ser. 7, 6, 410 - 419, 495 - 506, 530 - 537." type="journal article" year="1900">
<collectingCountry id="2F807671FFD2B917BA41E555AFBBFEB1" box="[868,961,290,311]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
1900
</bibRefCitation>
: 415
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="572836E1FFD2B917B9B2E513AEF5FE42" blockId="6.[151,1436,356,1964]" pageId="6" pageNumber="7">
<bibRefCitation id="33064B10FFD2B917B9B2E513AD35FEFA" author="Cameron" box="[151,335,356,380]" pageId="6" pageNumber="7" refString="Cameron, P. (1900) Description of new genera and species of Hymenoptera. A. New species of Vespidae from India and Japan. Annals and Magazine of Natural History, ser. 7, 6, 410 - 419, 495 - 506, 530 - 537." type="journal article" year="1900">
<collectingCountry id="2F807671FFD2B917B9B2E513AC84FEFA" box="[151,254,356,380]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
(1900)
</bibRefCitation>
described this species based only on the female. The male characters except for the genitalia were described by
<bibRefCitation id="33064B10FFD2B917B808E5FFAE53FE19" author="Das" box="[301,553,391,415]" pageId="6" pageNumber="7" refString="Das, B. P. &amp; Gupta, V. K. (1989) The Social Wasps of India and the Adjacent Countries (Hymenoptera: Vespidae). Oriental Insects Monograph 11. The Association for the Study of Oriental Insects, Gainesville, 292 pp." type="book" year="1989">Das and Gupta (1989)</bibRefCitation>
. Followings are detailed descriptions of the characters based on the specimens from
<collectingCountry id="2F807671FFD2B917B9F1E5DBAD4CFE42" box="[212,310,428,452]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
, and the nest is also described.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD2B917B9E2E5B8A9ECFDA9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="572836E1FFD2B917B9E2E5B8A9ECFDA9" blockId="6.[151,1436,356,1964]" pageId="6" pageNumber="7">
<emphasis id="65E3EAF3FFD2B917B9E2E5B8ADCAFE6E" bold="true" box="[199,432,463,488]" pageId="6" pageNumber="7">Material examined.</emphasis>
<collectingCountry id="2F807671FFD2B917B893E5A7AE41FE6E" box="[438,571,464,488]" name="Vietnam" pageId="6" pageNumber="7">VIETNAM</collectingCountry>
: Binh Lu, Phong Tho, Lai Chau, 22°
<geoCoordinate id="32A35026FFD2B917BAF4E5B8A862FE61" box="[977,1048,463,487]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="5555" value="21.5">21.5N</geoCoordinate>
103°46E,
<date id="23291021FFD2B917BDABE5B8A959FE61" box="[1166,1315,463,487]" pageId="6" pageNumber="7" value="2006-08-22">22.VIII.2006</date>
, LN, FS &amp; JK [
<date id="23291021FFD2B917B9E0E582AD63FD8A" box="[197,281,500,524]" pageId="6" pageNumber="7" value="1962-07-03">7 3, 62</date>
Ƥ (
<collectionCode id="3186AE24FFD2B917B864E583ADFFFD8A" box="[321,389,500,524]" pageId="6" pageNumber="7">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFD2B917B8B5E583ADA0FD8A" box="[400,474,500,524]" pageId="6" pageNumber="7">IUNH</collectionCode>
), alt.
<quantity id="906F9B04FFD2B917BB3DE583AE14FD8A" box="[536,622,500,524]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="7" unit="m" value="1600.0">1600 m</quantity>
, Nest# VN-P-2006-7; 11 Ƥ (
<collectionCode id="3186AE24FFD2B917BA8BE583AF89FD8A" box="[942,1011,500,524]" pageId="6" pageNumber="7">IEBR</collectionCode>
), alt.
<quantity id="906F9B04FFD2B917BD14E583A8FDFD8A" box="[1073,1159,500,524]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="2000.0">2000 m</quantity>
, Nest# VN-P-2006-9];
<date id="23291021FFD2B917BCABE583ACAEFDA9" pageId="6" pageNumber="7" value="1919-06-03">6 3, 19</date>
Ƥ (
<collectionCode id="3186AE24FFD2B917B9DFE660AD44FDA9" box="[250,318,535,559]" pageId="6" pageNumber="7">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFD2B917B86CE660ADE9FDA9" box="[329,403,535,559]" pageId="6" pageNumber="7">IUNH</collectionCode>
), Sa Pa NP (Silver Waterfall), alt.
<quantity id="906F9B04FFD2B917BA2BE660AF1EFDA9" box="[782,868,535,559]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="2000.0">2000 m</quantity>
, Nest# VN-P-
<date id="23291021FFD2B917BD20E660A829FDA9" box="[1029,1107,535,559]" pageId="6" pageNumber="7" value="2006-08-03">2006-3</date>
,
<date id="23291021FFD2B917BD78E660A88BFDA9" box="[1117,1265,535,559]" pageId="6" pageNumber="7" value="2006-08-20">20.VIII.2006</date>
, LN, FS &amp; JK.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD2B919B9E2E64CAEF8F9FB" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="572836E1FFD2B917B9E2E64CA835FDD2" blockId="6.[151,1436,356,1964]" box="[199,1103,571,596]" pageId="6" pageNumber="7">
<emphasis id="65E3EAF3FFD2B917B9E2E64CAD23FDD2" bold="true" box="[199,345,571,596]" pageId="6" pageNumber="7">Description.</emphasis>
FEMALE. Body length
<quantity id="906F9B04FFD2B917BB4BE64BAE8DFDD2" box="[622,759,572,596]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="6" pageNumber="7" unit="mm" value="10.75" valueMax="11.0" valueMin="10.5">10.511mm</quantity>
; fore wing length
<quantity id="906F9B04FFD2B917BAE7E64BA835FDD2" box="[962,1103,572,596]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.35" metricValueMin="1.1" pageId="6" pageNumber="7" unit="mm" value="12.25" valueMax="13.5" valueMin="11.0">1113.5 mm</quantity>
</paragraph>
<paragraph id="572836E1FFD2B917B9E2E628A8CFFB82" blockId="6.[151,1436,356,1964]" pageId="6" pageNumber="7">Head in frontal view about 1.2 × as wide as high (Fig. 14); in dorsal view weakly swollen laterally just behind eyes, then narrowed posteriorly, with posterior margin shallowly and broadly emarginated. Vertex slightly raised in area among ocelli, slightly slope down behind posterior ocelli towards occpital carina; POD:OOD=1:2.2; POD about 1.4 × Od (Fig. 15). Gena in lateral view about 0.8 × as wide as eye (Fig. 16); occipital carina fine, evanescent in ventral one third of gena. Inner eye margins weakly convergent ventrally, in frontal view of head about 1.1 × further apart from each other at vertex than at clypeus. Antennal sockets closer to inner eye margin than to each other. Anterior tentorial pit slightly further apart from ventral margin of antennal socket than from inner eye margin. Clypeus in frontal view as wide as high (Fig. 14); in lateral view weakly convex (Fig. 16); length of lateral margin of clypeus lying along inner eye margin shorter than diameter of antennal socket. Malar space longer than diameter of antennal socket. Antenna (Fig. 17) with scape slightly more than 3 × as long as its maximum width; flagellomere 1 slightly more than 2.5 × as long as its apical width, longer than length of flagellomeres 2 and 3 combined; flagellomere 2 as long as wide; terminal flagellomere bullet shaped, as long as its basal width.</paragraph>
<paragraph id="572836E1FFD2B917B9E2E067A8D1FA86" blockId="6.[151,1436,356,1964]" pageId="6" pageNumber="7">Pronotal carina sharply raised, produced dorsally into thin lamella in dorsal part, slightly sinuate backward on lateral sides, reaching ventral corner of pronotum. Mesoscutum weakly convex (Fig. 18), about as long as wide between tegulae; anterior margin broadly rounded. Scutellum convex, slightly depressed medially. Metanotum weakly convex, strongly depressed along anterior margin. Propodeum short; posterior face with shallow wide median excavation, of which width is about 0.4 × maximum width of propodeum; border between posterior and lateral faces bluntly angled, marked with slightly raised carina; propodeal orifice about 2 × as long as wide (measured at widest part), somewhat narrowed in dorsal half. Jugal lobe of hind wing rounded (Fig. 19).</paragraph>
<paragraph id="572836E1FFD2B917B9E2E17BAE40FA16" blockId="6.[151,1436,356,1964]" pageId="6" pageNumber="7">
Metasomal tergum 1 distinctly shorter than its apical width, in lateral view abruptly swollen dorsally just behind basal slit for reception of propodeal suspensory ligament; corner between anterior and dorsal face bluntly angled (Fig. 20). Sternum
<quantity id="906F9B04FFD2B917B89BE123AD93FAEA" box="[446,489,1364,1388]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="7" unit="in" value="2.0">2 in</quantity>
lateral view with dorsal margin slightly longer than its ventral margin; anteroventral corner in nearly right-angle (Fig. 20).
</paragraph>
<paragraph id="572836E1FFD2B917B9E2E1EBA838F97E" blockId="6.[151,1436,356,1964]" pageId="6" pageNumber="7">Clypeus with scattered large punctures, each bearing sharply pointed golden bristle. Mandible with several deep punctures. Frons densely covered with coarse flat-bottomed punctures. Vertex and gena with sparse strong punctures; punctures around ocelli strong and sparse; ventral one third of gena with sparse strong puncture. Pronotum with dense deep punctures; spaces between punctures very narrow, slightly raised to form reticulation. Mesocutum with coarse dense flat-bottomed punctures; punctures on scutellum and metanotum dense, smaller than those on mesoscutum. Mesepisternum densely with coarse punctures in posterodorsal part, barely punctured in anteroventral part; border between them distinct. Dorsal metapleuron with striae and sparse punctures. Propodeum with strong sharply-raised transverse striae; lateral face with strong punctures between striae, interpuncture spaces weakly raised to interconnect striae. Metasomal segments densely covered with minute punctures in addition to scattered small punctures (ill-defined on terga; more or less well-defined on sterna).</paragraph>
<paragraph id="572836E1FFD2B917B9E2E373A8B0F80E" blockId="6.[151,1436,356,1964]" pageId="6" pageNumber="7">
<emphasis id="65E3EAF3FFD2B917B9E2E373AD77F89A" box="[199,269,1796,1820]" italics="true" pageId="6" pageNumber="7">Color.</emphasis>
Black, with following parts orange-yellow: clypeus except black margins, line on gena along posterodorsal margin of eye, narrow band along pronotal carina with branches running along posterior margin of pronotum, scutellum, apical bands on terga 1 and 5 (wide), orange-yellow. Legs black. Wings pale brown, weakly infuscate, slightly more strongly infuscate along anterior margin of forewing; veins dark brown.
</paragraph>
<paragraph id="572836E1FFD2B917B9E2E3E3AFE7F82A" blockId="6.[151,1436,356,1964]" box="[199,925,1940,1964]" pageId="6" pageNumber="7">
MALE. Body length
<quantity id="906F9B04FFD2B917B891E3E3AE47F82A" box="[436,573,1940,1964]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="6" pageNumber="7" unit="mm" value="10.75" valueMax="11.0" valueMin="10.5">10.511mm</quantity>
; fore wing length
<quantity id="906F9B04FFD2B917BA2CE3E3AFE0F82A" box="[777,922,1940,1964]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" pageId="6" pageNumber="7" unit="mm" value="11.75" valueMax="12.0" valueMin="11.5">11.512 mm</quantity>
.
</paragraph>
<paragraph id="572836E1FFD3B916B9B2E301A8B6F870" blockId="7.[151,1436,1910,2038]" pageId="7" pageNumber="8">
<emphasis id="65E3EAF3FFD3B916B9B2E301AD25F808" bold="true" box="[151,351,1910,1934]" pageId="7" pageNumber="8">FIGURES 1426.</emphasis>
<taxonomicName id="90974D62FFD3B916B843E300AEF3F808" box="[358,649,1911,1934]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lepcha" subGenus="Polistella">
<emphasis id="65E3EAF3FFD3B916B843E300ADC0F808" box="[358,442,1911,1934]" italics="true" pageId="7" pageNumber="8">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFD3B916B8EEE300AE4BF808" box="[459,561,1911,1934]" italics="true" pageId="7" pageNumber="8">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFD3B916BB67E300AEF3F808" box="[578,649,1911,1934]" italics="true" pageId="7" pageNumber="8">lepcha</emphasis>
</taxonomicName>
. 1420. Ƥ.
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, 16, 21. Head. 14, 21. Frontal view. 16. Lateral view. 15. Ocelli, eye and posterior margin of vertex. 17. Right antenna. 18. mesosoma, lateral view. 19. Jugal lobe of right hind wing. 20. Metasomal segments 1 and 2, lateral view. 22. Left antenna. 2324. Terminal sterna. 23. Ventral view. 24. Apical part, lateral view. 2526. Aedeagus. 25. Ventral view. 26. Lateral view. Scale
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.
</paragraph>
<paragraph id="572836E1FFDCB919B9E2E4E0A86DFE2A" blockId="8.[151,1436,151,1077]" pageId="8" pageNumber="9">Structure as in female except: head proportionally smaller, more transverse, about 1.5 × as wide as high in frontal view (Fig. 21); eye strongly swollen laterally, inner eye margins about 1.4 × further apart from each other at vertex than at clypeus; gena narrow, in lateral view about 0.6 × as wide as eye, with weakly raised blunt ridge running along posterior margin of eye; clypeus in frontal view with apical margin slightly rounded, about 1.1 × as wide as high, covered with dense short golden hairs; antenna (Fig. 22) slender than in female; scape short, about 3 × as long as its maximum width; flagellomere 1 as long as length of flagellomeres 2 and 3 combined; flagellomeres 2 and 3 each longer than wide; terminal flagellomere elongate, slightly curved, about twice as long as its basal width. Metasomal sternum 7 depressed medially, without tubercle (Figs 23, 24).</paragraph>
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<emphasis id="65E3EAF3FFDCB919B9E2E5CFAD77FE56" box="[199,269,440,464]" italics="true" pageId="8" pageNumber="9">Color.</emphasis>
Similar to female, but antennal flagellomeres 5-11 reddish-brown to dark yellow beneath.
</paragraph>
<paragraph id="572836E1FFDCB919B9E2E5ABAE38FDBA" blockId="8.[151,1436,151,1077]" pageId="8" pageNumber="9">
<emphasis id="65E3EAF3FFDCB919B9E2E5ABAD42FE72" box="[199,312,476,500]" italics="true" pageId="8" pageNumber="9">Genitalia.</emphasis>
Similar to that of
<taxonomicName id="90974D62FFDCB919BB27E5ABAEC2FE72" box="[514,696,476,500]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFDCB919BB27E5ABAEC2FE72" box="[514,696,476,500]" italics="true" pageId="8" pageNumber="9">Polistes dawnae</emphasis>
</taxonomicName>
, but ventral serration of penis valves coarse and strong, especially in proximal half of penis valves (Fig. 25); penis valves in lateral view with dorsal margin and proximoventral margins both nearly straight (Fig. 26).
</paragraph>
<paragraph id="572836E1FFDCB919B9E2E63FAEA7FD69" blockId="8.[151,1436,151,1077]" pageId="8" pageNumber="9">
<emphasis id="65E3EAF3FFDCB919B9E2E63FAD79FDE6" bold="true" box="[199,259,584,608]" pageId="8" pageNumber="9">Nest.</emphasis>
The following three nests were examined: Nest# VN-P-2006-3, Silver Waterfall (
<geoCoordinate id="32A35026FFDCB919BDBDE630A977FDD9" box="[1176,1293,583,607]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="92" value="22.358334">22°21.5'N</geoCoordinate>
,
<geoCoordinate id="32A35026FFDCB919BC3DE630A9E2FDD9" box="[1304,1432,583,607]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="92" value="103.775">103°46.5'E</geoCoordinate>
, alt.
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), Sa Pa, Lao Cai,
<date id="23291021FFDCB919BB06E61BAEC2FD02" box="[547,696,620,644]" pageId="8" pageNumber="9" value="2006-08-20">20.VIII.2006</date>
, LN, FS &amp; JK; Nest# VN-P-2006-7 at alt.
<quantity id="906F9B04FFDCB919BDB2E61BA896FD02" box="[1175,1260,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="m" value="1600.0">1600 m</quantity>
and Nest# VN- P-2006-9 at alt.
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, Binh Lu, Phong Tho, Lai Chau,
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,
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,
<date id="23291021FFDCB919BD30E6F8A8D0FD21" box="[1045,1194,655,679]" pageId="8" pageNumber="9" value="2006-08-22">22.VIII.2006</date>
, LN, FS &amp; JK (
<tableCitation id="1A15035AFFDCB919BC79E6F8ACD2FD4A" captionStart="TABLE 2" captionStartId="8.[151,241,1129,1153]" captionTargetBox="[151,1412,1325,1428]" captionTargetPageId="8" captionText="TABLE 2. Measurements of nests of Polistes lepcha." httpUri="http://table.plazi.org/id/03E86669FFDCB919B9B2E01EAEA9FB07" pageId="8" pageNumber="9" tableUuid="03E86669FFDCB919B9B2E01EAEA9FB07">Table 2</tableCitation>
). They were all collected along the Hoang Lien Pass road, where the nests were on the cliffs along the road and attached directly to rocks
<quantity id="906F9B04FFDCB919B89DE6A0AE50FD69" box="[440,554,727,751]" metricMagnitude="0" metricUnit="m" metricValue="2.65" metricValueMax="4.0" metricValueMin="1.3" pageId="8" pageNumber="9" unit="m" value="2.65" valueMax="4.0" valueMin="1.3">1.3 to 4 m</quantity>
above the road.
</paragraph>
<paragraph id="572836E1FFDCB919B9E2E68AAD18FBB3" blockId="8.[151,1436,151,1077]" pageId="8" pageNumber="9">
Nest structural characters are as follows. Petiole single, terminal, with thin central core of plant fibers (in VN- P-2006-7 petiole with thick irregular central core of plant fibers,
<tableCitation id="1A15035AFFDCB919BAB4E768AF93FCB1" box="[913,1001,799,823]" captionStart="TABLE 2" captionStartId="8.[151,241,1129,1153]" captionTargetBox="[151,1412,1325,1428]" captionTargetPageId="8" captionText="TABLE 2. Measurements of nests of Polistes lepcha." httpUri="http://table.plazi.org/id/03E86669FFDCB919B9B2E01EAEA9FB07" pageId="8" pageNumber="9" tableUuid="03E86669FFDCB919B9B2E01EAEA9FB07">Table 2</tableCitation>
), enlarged strictly with adult salivary secretion, black and lustrous; salivary coating extending onto substrate and back of comb. Comb tough, pliable in texture, dark brownish gray, expanded from the petiole to form fan-shaped in ventral view, with ventral surface convex and dorsal surface deeply concave. Cells generally arranged in regular rows and hexagonal at open end, but partly irregularly arranged and some cells pentagonal; outer free margins of peripheral cells round; each cell expanded towards open end, with width at opening more than 1.5 × larger than that at bottom (
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); cell wall about
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thick. Cocoon cap slightly domed beyond rim of cell, pale green when spun, weathered with time to become pale gray.
</paragraph>
<caption id="03E86669FFDCB919B9B2E01EAEA9FB07" ID-Table-UUID="03E86669FFDCB919B9B2E01EAEA9FB07" box="[151,723,1129,1153]" httpUri="http://table.plazi.org/id/03E86669FFDCB919B9B2E01EAEA9FB07" pageId="8" pageNumber="9" targetBox="[151,1412,1325,1428]" targetIsTable="true" targetPageId="8">
<paragraph id="572836E1FFDCB919B9B2E01EAEA9FB07" blockId="8.[151,1395,1129,1304]" box="[151,723,1129,1153]" pageId="8" pageNumber="9">
<emphasis id="65E3EAF3FFDCB919B9B2E01EAD71FB07" bold="true" box="[151,267,1129,1153]" pageId="8" pageNumber="9">TABLE 2.</emphasis>
Measurements of nests of
<taxonomicName id="90974D62FFDCB919BB0FE01DAEB7FB07" box="[554,717,1130,1153]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFDCB919BB0FE01DAEB7FB07" box="[554,717,1130,1153]" italics="true" pageId="8" pageNumber="9">Polistes lepcha</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="572836E1FFDCB919B9EEE0E8A919FB34" blockId="8.[151,1395,1129,1304]" box="[203,1379,1182,1202]" pageId="8" pageNumber="9">Nest code Petiole Petiole Cell width at open Cell width at bottom Cell depth (mm)</paragraph>
<paragraph id="572836E1FFDCB919B858E0CAAE55FB69" blockId="8.[151,1395,1129,1304]" pageId="8" pageNumber="9">length thickness end(mm) (n=10) (mm) (n=10) (n=10) (mm) (mm)</paragraph>
<paragraph id="572836E1FFDCB919BBA3E172AEF8F9FB" blockId="8.[151,1395,1129,1304]" lastBlockId="8.[151,1436,1493,1698]" pageId="8" pageNumber="9">
Mean Range Mean Range Mean Range
<emphasis id="65E3EAF3FFDCB919B9E2E1A2AD4AFA6B" bold="true" box="[199,304,1493,1517]" pageId="8" pageNumber="9">Remark.</emphasis>
This species is distinguished from other species of subgenus
<emphasis id="65E3EAF3FFDCB919BAC5E1A2A831FA6B" box="[992,1099,1493,1517]" italics="true" pageId="8" pageNumber="9">Polistella</emphasis>
occurring in nothern
<collectingCountry id="2F807671FFDCB919BC18E1A2A9E6FA6B" box="[1341,1436,1493,1517]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
in having the combination of the following characters: propodeum with strongly and sharply raised transverse striae and strong punctures between the striae on lateral faces; border between the posterior and lateral faces of the propodeum well-defined and weakly carinate; anteriorly truncate second metasomal sternum; penis valves with strong serration along their ventral margins.
</paragraph>
</subSubSection>
<paragraph id="572836E1FFDCB919B9EEE15AA9FEFA12" pageId="8" pageNumber="9">
<table id="2597C441FFDC46EEB9B2E15AA9FEFA12" box="[151,1412,1325,1428]" gridcols="9" gridrows="3" pageId="8" pageNumber="9">
<tr id="E9A734A3FFDC46EEB9B2E15AA9FEFAC7" box="[151,1412,1325,1345]" gridrow="0" pageId="8" pageNumber="9">
<th id="AA765DDFFFDC46EEB9B2E15AAD3DFAC7" box="[151,327,1325,1345]" gridcol="0" gridrow="0" pageId="8" pageNumber="9">VN-P-2006-3</th>
<th id="AA765DDFFFDC46EEB8A9E15AADD2FAC7" box="[396,424,1325,1345]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">3.5</th>
<th id="AA765DDFFFDC46EEB8CBE15AAE47FAC7" box="[494,573,1325,1345]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">1.5 x 2.1</th>
<th id="AA765DDFFFDC46EEBBB7E15AAED4FAC7" box="[658,686,1325,1345]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">5.1</th>
<th id="AA765DDFFFDC46EEBA2DE15AAF37FAC7" box="[776,845,1325,1345]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">4.85.4</th>
<th id="AA765DDFFFDC46EEBA94E15AAFB7FAC7" box="[945,973,1325,1345]" gridcol="5" gridrow="0" pageId="8" pageNumber="9">3.2</th>
<th id="AA765DDFFFDC46EEBD3AE15AA819FAC7" box="[1055,1123,1325,1345]" gridcol="6" gridrow="0" pageId="8" pageNumber="9">3.03.6</th>
<th id="AA765DDFFFDC46EEBD9BE15AA89CFAC7" box="[1214,1254,1325,1345]" gridcol="7" gridrow="0" pageId="8" pageNumber="9">17.0</th>
<th id="AA765DDFFFDC46EEBC0CE15AA9FEFAC7" box="[1321,1412,1325,1345]" gridcol="8" gridrow="0" pageId="8" pageNumber="9">16.019.1</th>
</tr>
<tr id="E9A734A3FFDC46EEB9B2E121A9FEFAEC" box="[151,1412,1366,1386]" gridrow="1" pageId="8" pageNumber="9">
<th id="AA765DDFFFDC46EEB9B2E121AD3DFAEC" box="[151,327,1366,1386]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">VN-P-2006-7</th>
<td id="AA765DDFFFDC46EEB8A9E121ADD2FAEC" box="[396,424,1366,1386]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">3.3</td>
<td id="AA765DDFFFDC46EEB8CBE121AE47FAEC" box="[494,573,1366,1386]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">4.2 x 5.1</td>
<td id="AA765DDFFFDC46EEBBB7E121AED4FAEC" box="[658,686,1366,1386]" gridcol="3" gridrow="1" pageId="8" pageNumber="9">5.2</td>
<td id="AA765DDFFFDC46EEBA2DE121AF37FAEC" box="[776,845,1366,1386]" gridcol="4" gridrow="1" pageId="8" pageNumber="9">5.05.6</td>
<td id="AA765DDFFFDC46EEBA94E121AFB7FAEC" box="[945,973,1366,1386]" gridcol="5" gridrow="1" pageId="8" pageNumber="9">3.1</td>
<td id="AA765DDFFFDC46EEBD3AE121A819FAEC" box="[1055,1123,1366,1386]" gridcol="6" gridrow="1" pageId="8" pageNumber="9">2.93.7</td>
<td id="AA765DDFFFDC46EEBD9BE121A89CFAEC" box="[1214,1254,1366,1386]" gridcol="7" gridrow="1" pageId="8" pageNumber="9">17.2</td>
<td id="AA765DDFFFDC46EEBC0CE121A9FEFAEC" box="[1321,1412,1366,1386]" gridcol="8" gridrow="1" pageId="8" pageNumber="9">15.618.4</td>
</tr>
<tr id="E9A734A3FFDC46EEB9B2E1F7A9FEFA12" box="[151,1412,1408,1428]" gridrow="2" pageId="8" pageNumber="9">
<th id="AA765DDFFFDC46EEB9B2E1F7AD3DFA12" box="[151,327,1408,1428]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">VN-P-2006-9</th>
<td id="AA765DDFFFDC46EEB8A9E1F7ADD2FA12" box="[396,424,1408,1428]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">3.2</td>
<td id="AA765DDFFFDC46EEB8CBE1F7AE47FA12" box="[494,573,1408,1428]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">1.7 x 1.8</td>
<td id="AA765DDFFFDC46EEBBB7E1F7AED4FA12" box="[658,686,1408,1428]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">5.0</td>
<td id="AA765DDFFFDC46EEBA2DE1F7AF37FA12" box="[776,845,1408,1428]" gridcol="4" gridrow="2" pageId="8" pageNumber="9">4.75.3</td>
<td id="AA765DDFFFDC46EEBA94E1F7AFB7FA12" box="[945,973,1408,1428]" gridcol="5" gridrow="2" pageId="8" pageNumber="9">3.0</td>
<td id="AA765DDFFFDC46EEBD3AE1F7A819FA12" box="[1055,1123,1408,1428]" gridcol="6" gridrow="2" pageId="8" pageNumber="9">2.63.4</td>
<td id="AA765DDFFFDC46EEBD9BE1F7A89CFA12" box="[1214,1254,1408,1428]" gridcol="7" gridrow="2" pageId="8" pageNumber="9">17.6</td>
<td id="AA765DDFFFDC46EEBC0CE1F7A9FEFA12" box="[1321,1412,1408,1428]" gridcol="8" gridrow="2" pageId="8" pageNumber="9">16.619.1</td>
</tr>
</table>
</paragraph>
<subSubSection id="1F8D656AFFDCB919B9E2E2FFAF98F927" box="[199,994,1672,1698]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="572836E1FFDCB919B9E2E2FFAF98F927" blockId="8.[151,1436,1493,1698]" box="[199,994,1672,1698]" pageId="8" pageNumber="9">
<emphasis id="65E3EAF3FFDCB919B9E2E2FFAD1AF927" bold="true" box="[199,352,1672,1697]" pageId="8" pageNumber="9">Distribution.</emphasis>
<collectingCountry id="2F807671FFDCB919B842E2FEADDFF927" box="[359,421,1673,1697]" name="India" pageId="8" pageNumber="9">India</collectingCountry>
: Haryana, Meghalaya, Manipur; northern
<collectingCountry id="2F807671FFDCB919BA5EE2FEAFA4F927" box="[891,990,1673,1697]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="311EABA5097B28499E35B3A9E7A01BB8" type="DOI">10.5281/zenodo.201094</mods:identifier>
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<treatment id="DF3E87F7FFD6B914B9B2E152AF2FF8D5" ID-DOI="http://doi.org/10.5281/zenodo.5489166" ID-GBIF-Taxon="119419230" ID-Zenodo-Dep="5489166" LSID="urn:lsid:plazi:treatment:DF3E87F7FFD6B914B9B2E152AF2FF8D5" httpUri="http://treatment.plazi.org/id/DF3E87F7FFD6B914B9B2E152AF2FF8D5" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="1F8D656AFFD6B913B9B2E152AD38FAD9" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="572836E1FFD6B913B9B2E152AEA8FAB9" blockId="2.[151,722,1317,1375]" box="[151,722,1317,1343]" pageId="2" pageNumber="3">
<heading id="0C60818DFFD6B913B9B2E152AEA8FAB9" bold="true" box="[151,722,1317,1343]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<taxonomicName id="90974D62FFD6B913B9B2E152AEA8FAB9" ID-CoL="77SQ5" authority="Dover &amp; Rao 1922" authorityName="Dover &amp; Rao" authorityYear="1922" box="[151,722,1317,1343]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dawnae" subGenus="Polistella">
<emphasis id="65E3EAF3FFD6B913B9B2E152AEA8FAB9" bold="true" box="[151,722,1317,1343]" pageId="2" pageNumber="3">
<emphasis id="65E3EAF3FFD6B913B9B2E152AC8EFAB9" bold="true" box="[151,244,1317,1343]" italics="true" pageId="2" pageNumber="3">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFD6B913B821E152AD0EFAB9" bold="true" box="[260,372,1317,1343]" italics="true" pageId="2" pageNumber="3">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFD6B913B8A3E152AD99FAB9" bold="true" box="[390,483,1317,1343]" italics="true" pageId="2" pageNumber="3">dawnae</emphasis>
<bibRefCitation id="33064B10FFD6B913B8CFE152AEA8FAB9" author="Dover" box="[490,722,1317,1343]" pageId="2" pageNumber="3" refString="Dover, C. &amp; Rao, S. (1922) A note on the diplopterous wasps in the collection of the Indian Museum. Journal and Proceedings, Asiatic Society of Bengal, 18 (4), 236 - 249." type="journal article" year="1922">Dover &amp; Rao 1922</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="572836E1FFD6B913B9B2E130AD38FAD9" blockId="2.[151,722,1317,1375]" box="[151,322,1351,1375]" pageId="2" pageNumber="3">(Figs 113, 85)</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD6B913B9B2E1E7AEA1F8AF" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="572836E1FFD6B913B9B2E1E7ADF3FA42" blockId="2.[151,1436,1424,1476]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="778711CFFFD6B913B9B2E1E7ADF3FA42" pageId="2" pageNumber="3">
<taxonomicName id="90974D62FFD6B913B9B2E1E7AE4CFA23" ID-CoL="77SQ5" authority="Dover &amp; Rao 1922: 248" authorityName="Dover &amp; Rao" authorityPageNumber="248" authorityYear="1922" box="[151,566,1424,1446]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFD6B913B9B2E1E7AD46FA20" box="[151,316,1424,1446]" italics="true" pageId="2" pageNumber="3">Polistes dawnae</emphasis>
<treatmentCitation id="D63610F0FFD6B913B864E1E6AE4CFA23" author="Dover" box="[321,566,1424,1446]" page="248" pageId="2" pageNumber="3" year="1922">
<bibRefCitation id="33064B10FFD6B913B864E1E6AE7EFA23" author="Dover" box="[321,516,1424,1446]" pageId="2" pageNumber="3" refString="Dover, C. &amp; Rao, S. (1922) A note on the diplopterous wasps in the collection of the Indian Museum. Journal and Proceedings, Asiatic Society of Bengal, 18 (4), 236 - 249." type="journal article" year="1922">Dover &amp; Rao 1922</bibRefCitation>
: 248
</treatmentCitation>
</taxonomicName>
, Ƥ,
<typeStatus id="882C8843FFD6B913BB7BE1E7AECCFA20" box="[606,694,1424,1446]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
Dawna Hills [
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,
<geoCoordinate id="32A35026FFD6B913BA8CE1E7AF86FA20" box="[937,1020,1424,1446]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="98.25">98°15'E</geoCoordinate>
],
<collectingCountry id="2F807671FFD6B913BD2BE1E6A829FA20" box="[1038,1107,1425,1446]" name="Myanmar" pageId="2" pageNumber="3">Burma</collectingCountry>
[
<collectingCountry id="2F807671FFD6B913BD7AE1E7A8B3FA20" box="[1119,1225,1424,1446]" name="Myanmar" pageId="2" pageNumber="3">Myanmar</collectingCountry>
], [Zoological Survey of
<collectingCountry id="2F807671FFD6B913B9C7E1D9AD63FA42" box="[226,281,1454,1476]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
, Kolikata].
</treatmentCitationGroup>
</paragraph>
<paragraph id="572836E1FFD6B913B9B2E186A8CBF9F3" blockId="2.[151,1436,1521,2013]" pageId="2" pageNumber="3">
<taxonomicName id="90974D62FFD6B913B9B2E186AD37F98F" ID-CoL="77SQ5" box="[151,333,1521,1545]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFD6B913B9B2E186AD37F98F" box="[151,333,1521,1545]" italics="true" pageId="2" pageNumber="3">Polistes dawnae</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="33064B10FFD6B913BB39E185AF79F98F" author="Dover" box="[540,771,1521,1545]" pageId="2" pageNumber="3" refString="Dover, C. &amp; Rao, S. (1922) A note on the diplopterous wasps in the collection of the Indian Museum. Journal and Proceedings, Asiatic Society of Bengal, 18 (4), 236 - 249." type="journal article" year="1922">Dover &amp; Rao (1922)</bibRefCitation>
based on a single female specimen (
<typeStatus id="882C8843FFD6B913BDB9E186A97BF98F" box="[1180,1281,1521,1545]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
) from Dawna Hills [
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,
<geoCoordinate id="32A35026FFD6B913B862E262ADDFF9AB" box="[327,421,1557,1581]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="98.25">98°15'E</geoCoordinate>
],
<collectingCountry id="2F807671FFD6B913B89DE261AE7EF9AB" box="[440,516,1558,1581]" name="Myanmar" pageId="2" pageNumber="3">Burma</collectingCountry>
[
<collectingCountry id="2F807671FFD6B913BB34E261AEFCF9A8" box="[529,646,1558,1582]" name="Myanmar" pageId="2" pageNumber="3">Myanmar</collectingCountry>
], which is housed in the Zoological Survey of
<collectingCountry id="2F807671FFD6B913BDAEE262A8BDF9AB" box="[1163,1223,1557,1581]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
, Kolikata.
<bibRefCitation id="33064B10FFD6B913BC64E261AD4BF9D7" author="Das" pageId="2" pageNumber="3" refString="Das, B. P. &amp; Gupta, V. K. (1989) The Social Wasps of India and the Adjacent Countries (Hymenoptera: Vespidae). Oriental Insects Monograph 11. The Association for the Study of Oriental Insects, Gainesville, 292 pp." type="book" year="1989">Das and Gupta (1989)</bibRefCitation>
gave a detailed description of the characters of the
<typeStatus id="882C8843FFD6B913BABCE24EAF84F9D7" box="[921,1022,1593,1617]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
, and no further information on this species has been added since then. The male and a nest of this species are herewith described.
</paragraph>
<paragraph id="572836E1FFD6B913B9E2E2F6AEA1F8AF" blockId="2.[151,1436,1521,2013]" pageId="2" pageNumber="3">
The female specimens we examined well agreed well with the description by
<bibRefCitation id="33064B10FFD6B913BD01E2F5A967F91F" author="Das" box="[1060,1309,1665,1689]" pageId="2" pageNumber="3" refString="Das, B. P. &amp; Gupta, V. K. (1989) The Social Wasps of India and the Adjacent Countries (Hymenoptera: Vespidae). Oriental Insects Monograph 11. The Association for the Study of Oriental Insects, Gainesville, 292 pp." type="book" year="1989">Das and Gupta (1989)</bibRefCitation>
. One of the seven females collected at the Muong Nhe NP Station was the only adult found on the nest described here. The nest was made in a cut bamboo and had no immatures in it. The other six females and a male were collected in flight near the nest site and may have emerged from the nest. As described below, the male is similar to the female both in structure and coloration except for some characters.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD6B913B9E2E342AFA7F813" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="572836E1FFD6B913B9E2E342AFA7F813" blockId="2.[151,1436,1521,2013]" pageId="2" pageNumber="3">
<emphasis id="65E3EAF3FFD6B913B9E2E342ADCAF8C8" bold="true" box="[199,432,1845,1870]" pageId="2" pageNumber="3">Material examined.</emphasis>
1 Ƥ (
<collectionCode id="3186AE24FFD6B913B8CEE342AE57F8CB" box="[491,557,1845,1869]" httpUri="http://grbio.org/cool/zeys-ppzt" name="NSM-PV, National Science Museum" pageId="2" pageNumber="3">NSM</collectionCode>
), Thung Man Pass, Phong Tho, Lai Chau, alt.
<quantity id="906F9B04FFD6B913BD08E342A8FEF8CB" box="[1069,1156,1845,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="1200.0">1200 m</quantity>
,
<date id="23291021FFD6B913BDAAE342A88DF8CB" box="[1167,1271,1845,1869]" pageId="2" pageNumber="3" value="1995-05-09">9.V.1995</date>
, A. Shinohara; 7 Ƥ, 1 3 (
<collectionCode id="3186AE24FFD6B913B827E32EAD3CF8F7" box="[258,326,1881,1905]" pageId="2" pageNumber="3">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFD6B913B874E32EADE6F8F7" box="[337,412,1881,1905]" pageId="2" pageNumber="3">IUNH</collectionCode>
), Muong Nhe NP Station, Dien Bien,
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,
<geoCoordinate id="32A35026FFD6B913BAE5E32EA850F8F7" box="[960,1066,1881,1905]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="102.4">102°24'E</geoCoordinate>
,
<date id="23291021FFD6B913BD13E32EA8BAF8F7" box="[1078,1216,1881,1905]" pageId="2" pageNumber="3" value="2009-07-23">23.VII.2009</date>
, LN, HP &amp; JK; 1 3 (
<collectionCode id="3186AE24FFD6B913B9BAE30AAC9EF813" box="[159,228,1917,1941]" pageId="2" pageNumber="3">IEBR</collectionCode>
), Pa Co, Mai Chau, Hoa Binh, alt.
<quantity id="906F9B04FFD6B913BB4FE30AAEBBF813" box="[618,705,1917,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="1000.0">1000 m</quantity>
,
<date id="23291021FFD6B913BBE8E30AAF1BF813" box="[717,865,1917,1941]" pageId="2" pageNumber="3" value="2005-08-23">23.VIII.2005</date>
, LN &amp; JK.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD6B914B9E2E3D6AD16F8B6" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
<paragraph id="572836E1FFD6B913B9E2E3D6AE1EF85B" blockId="2.[151,1436,1521,2013]" pageId="2" pageNumber="3">
<emphasis id="65E3EAF3FFD6B913B9E2E3D6AD23F83C" bold="true" box="[199,345,1953,1978]" pageId="2" pageNumber="3">Description.</emphasis>
MALE [female characters in square brackets]. Body length about
<quantity id="906F9B04FFD6B913BD7AE3D6A8CBF83F" box="[1119,1201,1953,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="mm" value="10.0">10 mm</quantity>
[
<quantity id="906F9B04FFD6B913BDE7E3D6A923F83F" box="[1218,1369,1953,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="2" pageNumber="3" unit="mm" value="10.75" valueMax="11.0" valueMin="10.5">10.511 mm</quantity>
]; fore wing length about
<quantity id="906F9B04FFD6B913B842E3B2ADCDF85B" box="[359,439,1989,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="mm" value="10.0">10 mm</quantity>
[
<quantity id="906F9B04FFD6B913B8E1E3B2AE20F85B" box="[452,602,1989,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="2" pageNumber="3" unit="mm" value="10.75" valueMax="11.0" valueMin="10.5">10.511 mm</quantity>
].
</paragraph>
<paragraph id="572836E1FFD7B912B9B2E32AAF55F85B" blockId="3.[151,1436,1885,2013]" pageId="3" pageNumber="4">
<emphasis id="65E3EAF3FFD7B912B9B2E32AAD2AF8F3" bold="true" box="[151,336,1885,1909]" pageId="3" pageNumber="4">FIGURES 113.</emphasis>
<taxonomicName id="90974D62FFD7B912B873E329AEF9F8F3" box="[342,643,1886,1909]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dawnae" subGenus="Polistella">
<emphasis id="65E3EAF3FFD7B912B873E329ADD0F8F3" box="[342,426,1886,1909]" italics="true" pageId="3" pageNumber="4">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFD7B912B89FE329AE5AF8F3" box="[442,544,1886,1909]" italics="true" pageId="3" pageNumber="4">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFD7B912BB15E329AEF9F8F3" box="[560,643,1886,1909]" italics="true" pageId="3" pageNumber="4">dawnae</emphasis>
</taxonomicName>
. 15. Ƥ. 613. 3. 1, 3, 67. Head. 1, 6. Frontal view. 3, 7. Lateral view. 2. Ocelli, eye and posterior margin of vertex. 4, 8. Right antenna. 5. Metasomal segments 1 and 2, lateral view. 910. Terminal sterna. 9. Ventral view. 10. Apical part, lateral view. 11. Inner aspect of paramere with digitus and volsella. 12 13. Aedeagus. 12. Ventral view. 13. Lateral view. Scale
<quantity id="906F9B04FFD7B912BBCFE3B1AF51F85B" box="[746,811,1990,2013]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="572836E1FFD0B915B9E2E4E0AF43FDD9" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="5">Head in frontal view (Fig. 6) about 1.4 × [about 1.2 × (Fig. 1)] as wide as high. Vertex slightly raised in area among ocelli; POD:OOD=1:1.5 [1:1.9 (Fig. 2)]; POD about 1.7 × [about 1.6 ×] Od. Eye much more strongly swollen laterally than in female; inner eye margins about 1.4 × [about 1.1 ×] further apart from each other at vertex than at clypeus. Gena narrow [more or less wide], in lateral view (Fig. 7) about 0.3 × [about 0.8 × (Fig. 3)] as wide as eye; weakly raised blunt ridge present, running along posterior margin of eye [absent]. Clypeus in frontal view (Fig. 6) about as wide as high, apical margin slightly produced, broadly rounded [pointed apically (Fig. 1)]; in lateral view (Fig. 7) slightly convex dorsally, slightly concave ventrally [weakly swollen anteriorly in dorsal half (Fig. 3)]; surface covered with dense short golden hairs [upper half of clypeus covered with fine pale pubescence and lower half with scattered deep large punctures each bearing sharply pointed golden bristle]. Antenna (Fig. 8) with slenderer flagellum than in female (Fig. 4); scape short and thick, about 2.8 × [about 3.0 ×] as long as its maximum width; flagellomere 1 as long as [slightly longer than] length of flagellomeres 2 and 3 combined; flagellomeres 2 and 3 each longer than wide [as long as wide]; terminal flagellomere slightly curved [bullet shaped], about 2.5 × longer than [as long as] its basal width.</paragraph>
<paragraph id="572836E1FFD0B915B9E2E61AAF44FCDA" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="5">Mesosoma as in female. Pronotal carina sharply raised, produced into thin lamella in dorsal part, slightly sinuate on lateral sides, reaching ventral corner of pronotum. Mesoscutum weakly convex, about as long as wide between tegulae; anterior margin broadly rounded. Disc of scutellum flat, in profile at the same level as mesoscutum. Metanotum weakly convex, strongly depressed along anterior margin. Propodeum short; posterior face widely (about half the maximum width of propodeum) and shallowly excavated medially, more or less smoothly passing into lateral faces; propodeal orifice elongate, about 1.8 × as long as wide (measured at widest part), somewhat narrowed in dorsal half. Jugal lobe of hind wing large, rounded.</paragraph>
<paragraph id="572836E1FFD0B915B9E2E71EAF80FB89" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="5">
Metasoma except terminal two segments as in female (Fig. 5). Tergum 1 distinctly shorter than its apical width, in lateral view abruptly swollen dorsally just behind basal slit for reception of propodeal suspensory ligament; corner between anterior and dorsal faces bluntly angled. Sternum
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lateral view strongly swollen ventrally in anterior one-third, then ventral margin bluntly angled into nearly straight line parallel to ventral margin of the tergum. Metasomal sternum 7 depressed medially, without tubercle (Figs 9, 10).
</paragraph>
<paragraph id="572836E1FFD0B915B9E2E06BAEE1FA1A" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="5">Body surface sculpture as in female, except for densely-haired clypeus. Mandible with scattered shallow punctures. Frons and vertex up to level of posterior ocelli with close fine punctures. Vertex behind posterior ocelli and gena with sparse fine punctures; spaces between punctures smooth; punctures in ventral one third of gena scaratted and deep. Pronotum with dense deep punctures; spaces between punctures very narrow, slightly raised to form reticulation. Mesoscutum with scattered deep flat-bottomed punctures and dense fine punctures between flat-bottomed punctures. Scutellum and metanotum with scattered deep punctures. Mesepisternum with coarse dense welldefined punctures in posterodorsal part; punctures in dorsal margin similar to those on pronotum. Dorsal metapleuron with week striation; ventral metapleuron with shallow large punctures. Propodeum with distinct transverse striation and punctures between striae on lateral faces; interpuncture spaces weakly raised to interconnect striae. Metasomal segements densely covered with minute punctures in addition to scattered small punctures (ill-defined on terga; more or less well-defined on sterna).</paragraph>
<paragraph id="572836E1FFD0B915B9E2E1DFAE0FF882" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="5">
<emphasis id="65E3EAF3FFD0B915B9E2E1DFAD77FA46" box="[199,269,1448,1472]" italics="true" pageId="4" pageNumber="5">Color.</emphasis>
Similar to female [body black, with following yellow markings: large apical spot on clypeus, narrow band along inner eye margin extending from bottom of frons to middle of eye emargination, narrow short band on gena along posterodorsal margin of eye, narrow band along pronotal carina, paired anterolateral spots on metanotum, scrobal spot on mesepisternum, paired large lateral spots on propodeum, propodeal valvula, metasomal tergum 1 except for basal triangular dark-brown spot, and apical margin of tergum 4; legs black], but more extensively marked with yellow as follows: clypeus entirely, mandible except for small basal black spot, ventral half of frons, narrow long band on gene along posterior margin of eye, antennal scape beneath, paired anterolateral spots (sometimes united to form narrow transverse band) on metanotum, spot on propodeal valvula, ventral face of fore coxa and spot on mid-coxa; antennal flagellum reddish brown to dark yellow beneath; paired lateral yellow spots on propodeum much reduced in size.
</paragraph>
<paragraph id="572836E1FFD0B915B9E2E367AEE4F85A" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="5">
<emphasis id="65E3EAF3FFD0B915B9E2E367AD0CF8AE" box="[199,374,1808,1832]" italics="true" pageId="4" pageNumber="5">Male genitalia.</emphasis>
Lamina volsellaris desclerotized and fused with cuspis. Digitus in inner aspect of paramere (Fig. 11) slightly more than 3.5 × as long as wide (measured at widest part). Aedeagus (Figs 1213): penis valve slightly longer than basal apodeme, in ventral view swollen laterally in proximal one fourth and distinctly produced laterally near proximal margin, in lateral view slightly thickened in proximal one fourth and with dorsal margin weakly and smoothly sinuate, with proxomoventral corner produced into small tooth (Fig. 13); ventral margin of penis valve finely serrated along entire length.
</paragraph>
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<paragraph id="572836E1FFD1B914B9B2E4EAA857FF35" blockId="5.[151,1069,156,179]" box="[151,1069,156,179]" pageId="5" pageNumber="6">
<emphasis id="65E3EAF3FFD1B914B9B2E4EAAD7EFF34" bold="true" box="[151,260,157,178]" pageId="5" pageNumber="6">TABLE 1.</emphasis>
Biological data of examined nests of
<taxonomicName id="90974D62FFD1B914BBA6E4EAAEA9FF35" box="[643,723,157,179]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="65E3EAF3FFD1B914BBA6E4EAAEA9FF35" box="[643,723,157,179]" italics="true" pageId="5" pageNumber="6">Polistes</emphasis>
</taxonomicName>
species from subgenus
<emphasis id="65E3EAF3FFD1B914BAE0E4EAA85CFF35" box="[965,1062,157,179]" italics="true" pageId="5" pageNumber="6">Polistella</emphasis>
.
</paragraph>
</caption>
<paragraph id="572836E1FFD1B914B980E4A3A903FBE9" pageId="5" pageNumber="6">
<table id="2597C441FFD146EEB980E4ADA9F3FBE9" box="[165,1417,218,1135]" gridcols="6" gridrows="8" pageId="5" pageNumber="6">
<tr id="E9A734A3FFD146EEB980E4ADA9F3FEC1" box="[165,1417,218,327]" gridrow="0" pageId="5" pageNumber="6">
<th id="AA765DDFFFD146EEB980E4ADAD03FEC1" box="[165,377,218,327]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">Species and nest code</th>
<th id="AA765DDFFFD146EEB8EFE4ADAE38FEC1" box="[458,578,218,327]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">Substrate</th>
<th id="AA765DDFFFD146EEBB41E4ADAEA1FEC1" box="[612,731,218,327]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">Date of collection</th>
<th id="AA765DDFFFD146EEBA26E4ADAF39FEC1" box="[771,835,218,327]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">Height from ground (m)</th>
<th id="AA765DDFFFD146EEBA57E4ADAFE1FEC1" box="[882,923,218,327]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">cells</th>
<th id="AA765DDFFFD146EEBAE4E4ADA9F3FEC1" box="[961,1417,218,327]" gridcol="5" gridrow="0" pageId="5" pageNumber="6">Number of Number of adults collected eggs larvae pupae empty female male cells</th>
</tr>
<tr id="E9A734A3FFD146EEB980E514A9F3FEF0" box="[165,1417,355,374]" gridrow="1" pageId="5" pageNumber="6" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="AA765DDFFFD146EEB980E514AD03FEF0" box="[165,377,355,374]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">
<taxonomicName id="90974D62FFD1B914B980E514AD4AFEF0" box="[165,304,355,374]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFD1B914B980E514AD4AFEF0" box="[165,304,355,374]" italics="true" pageId="5" pageNumber="6">Polistes lepcha</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="E9A734A3FFD146EEB980E5E6A9F3FE23" box="[165,1417,401,421]" gridrow="2" pageId="5" pageNumber="6">
<th id="AA765DDFFFD146EEB980E5E6AD03FE23" box="[165,377,401,421]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">VN-P-2006-3</th>
<td id="AA765DDFFFD146EEB8EFE5E6AE38FE23" box="[458,578,401,421]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">Rock</td>
<td id="AA765DDFFFD146EEBB41E5E6AEA1FE23" box="[612,731,401,421]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">20.VIII.2006</td>
<td id="AA765DDFFFD146EEBA26E5E6AF39FE23" box="[771,835,401,421]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">1.3</td>
<td id="AA765DDFFFD146EEBA57E5E6AFE1FE23" box="[882,923,401,421]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">229</td>
<td id="AA765DDFFFD146EEBAE4E5E6A9F3FE23" box="[961,1417,401,421]" gridcol="5" gridrow="2" pageId="5" pageNumber="6">29 32 41 127 13 0</td>
</tr>
<tr id="E9A734A3FFD146EEB980E5C9A9F3FE54" box="[165,1417,446,466]" gridrow="3" pageId="5" pageNumber="6">
<th id="AA765DDFFFD146EEB980E5C9AD03FE54" box="[165,377,446,466]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">VN-P-2006-7</th>
<td id="AA765DDFFFD146EEB8EFE5C9AE38FE54" box="[458,578,446,466]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">Rock</td>
<td id="AA765DDFFFD146EEBB41E5C9AEA1FE54" box="[612,731,446,466]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">22.VIII.2006</td>
<td id="AA765DDFFFD146EEBA26E5C9AF39FE54" box="[771,835,446,466]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">4.0</td>
<td id="AA765DDFFFD146EEBA57E5C9AFE1FE54" box="[882,923,446,466]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">390</td>
<td id="AA765DDFFFD146EEBAE4E5C9A9F3FE54" box="[961,1417,446,466]" gridcol="5" gridrow="3" pageId="5" pageNumber="6">62 53 15 260 38 17</td>
</tr>
<tr id="E9A734A3FFD146EEB980E599A9F3FD84" box="[165,1417,494,514]" gridrow="4" pageId="5" pageNumber="6">
<th id="AA765DDFFFD146EEB980E599AD03FD84" box="[165,377,494,514]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">VN-P-2006-9</th>
<td id="AA765DDFFFD146EEB8EFE599AE38FD84" box="[458,578,494,514]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">Rock</td>
<td id="AA765DDFFFD146EEBB41E599AEA1FD84" box="[612,731,494,514]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">22.VIII.2006</td>
<td id="AA765DDFFFD146EEBA26E599AF39FD84" box="[771,835,494,514]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">1.6</td>
<td id="AA765DDFFFD146EEBA57E599AFE1FD84" box="[882,923,494,514]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">161</td>
<td id="AA765DDFFFD146EEBAE4E599A9F3FD84" box="[961,1417,494,514]" gridcol="5" gridrow="4" pageId="5" pageNumber="6">29 25 19 88 11 0</td>
</tr>
<tr id="E9A734A3FFD146EEB980E669A9F3FDB7" box="[165,1417,542,561]" gridrow="5" pageId="5" pageNumber="6" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="AA765DDFFFD146EEB980E669AD03FDB7" box="[165,377,542,561]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">
<taxonomicName id="90974D62FFD1B914B980E669AD40FDB7" box="[165,314,542,561]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFD1B914B980E669AD40FDB7" box="[165,314,542,561]" italics="true" pageId="5" pageNumber="6">Polistes dawnae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="E9A734A3FFD146EEB980E63CA9F3FD20" box="[165,1417,587,678]" gridrow="6" pageId="5" pageNumber="6">
<th id="AA765DDFFFD146EEB980E63CAD03FD20" box="[165,377,587,678]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">
2009-DB-P-01
<taxonomicName id="90974D62FFD1B914B980E6E4AD03FD20" box="[165,377,659,678]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFD1B914B980E6E4AD03FD20" box="[165,377,659,678]" italics="true" pageId="5" pageNumber="6">Polistes reliciniclypeus</emphasis>
</taxonomicName>
</th>
<td id="AA765DDFFFD146EEB8EFE63CAE38FD20" box="[458,578,587,678]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">Inside cut bamboo</td>
<td id="AA765DDFFFD146EEBB41E63CAEA1FD20" box="[612,731,587,678]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">21.VII. 2009</td>
<td id="AA765DDFFFD146EEBA26E63CAF39FD20" box="[771,835,587,678]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">1.3</td>
<td id="AA765DDFFFD146EEBA57E63CAFE1FD20" box="[882,923,587,678]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">247</td>
<td id="AA765DDFFFD146EEBAE4E63CA9F3FD20" box="[961,1417,587,678]" gridcol="5" gridrow="6" pageId="5" pageNumber="6">0 0 0 247 1 0</td>
</tr>
<tr id="E9A734A3FFD146EEB980E6B7A9F3FBE9" box="[165,1417,704,1135]" gridrow="7" pageId="5" pageNumber="6">
<th id="AA765DDFFFD146EEB980E6B7AD03FBE9" box="[165,377,704,1135]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">
VN-P-2006-4 VN-P-2006-10 VN-P-2006-11 VN-P-2006-12 VN-P-2006-13 2008-LC-P-02 2008-LC-P-03 2008-HB-P-03 2009-Sapa-P-01
<taxonomicName id="90974D62FFD1B914B980E045ACAFFBE9" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gilvus">
<emphasis id="65E3EAF3FFD1B914B980E045AD53FBC3" box="[165,297,1074,1093]" italics="true" pageId="5" pageNumber="6">Polistes gilvus</emphasis>
2008
</taxonomicName>
-SL-P-1
</th>
<td id="AA765DDFFFD146EEB8EFE6B7AE38FBE9" box="[458,578,704,1135]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">Palm leaf Rock Rock Rock Rock unrecorded unrecorded unrecorded Rock Shrub branch</td>
<td id="AA765DDFFFD146EEBB41E6B7AEA1FBE9" box="[612,731,704,1135]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">21.VIII.2006 22.VIII.2006 22.VIII.2006 22.VIII.2006 22.VIII.2006 25.VI.2008 26.VI.2008 11.VI.2008 5.VII.2009 7.VI.2008</td>
<td id="AA765DDFFFD146EEBA26E6B7AF39FBE9" box="[771,835,704,1135]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">0.5 - - - - - - - - 1.2</td>
<td id="AA765DDFFFD146EEBA57E6B7AFE1FBE9" box="[882,923,704,1135]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">287 245 107 68 580 55 19 63 53 126</td>
<td id="AA765DDFFFD146EEBAE4E6B7A9F3FBE9" box="[961,1417,704,1135]" gridcol="5" gridrow="7" pageId="5" pageNumber="6">5 14 0 268 3 2 0 44 32 169 0 14 22 34 15 36 3 3 2 20 3 43 3 57 13 55 455 15 15 0 0 0 55 4 0 0 0 0 19 2 0 0 0 0 63 5 0 0 19 0 34 6 0 0 0 2 124 7 0</td>
</tr>
</table>
</paragraph>
<paragraph id="572836E1FFD1B914B9E2E08CAF69FADD" blockId="5.[151,1437,1275,1876]" pageId="5" pageNumber="6">
<emphasis id="65E3EAF3FFD1B914B9E2E08CAD79FA95" bold="true" box="[199,259,1275,1299]" pageId="5" pageNumber="6">Nest.</emphasis>
A nest (Nest# 2009-DB-P-01) collected in Muong Nhe, Dien Bien on
<date id="23291021FFD1B914BD3DE08CA8D2FA95" box="[1048,1192,1275,1299]" pageId="5" pageNumber="6" value="2009-07-21">21 July 2009</date>
was examined (
<tableCitation id="1A15035AFFD1B914BC79E08CACDEFAB1" captionStart="TABLE 1" captionStartId="5.[151,235,157,178]" captionTargetBox="[165,1417,218,1135]" captionTargetPageId="5" captionText="TABLE 1. Biological data of examined nests of Polistes species from subgenus Polistella." httpUri="http://table.plazi.org/id/03E86669FFD1B914B9B2E4EAA857FF35" pageId="5" pageNumber="6" tableUuid="03E86669FFD1B914B9B2E4EAA857FF35">Table 1</tableCitation>
). As mentioned above, the nest was collected, together with a single female, from the inside of a cut bamboo used for the wall of the kitchen at the Muong Nhe NP Station.
</paragraph>
<paragraph id="572836E1FFD1B914B9E2E110AD16F8B6" blockId="5.[151,1437,1275,1876]" pageId="5" pageNumber="6">
The nest characters other than the nesting site are as follows. Single combed (
<figureCitation id="CFAC2A64FFD1B914BD51E110A8B7FAF9" box="[1140,1229,1383,1407]" captionStart="FIGURES 85 88" captionStartId="26.[151,269,1501,1525]" captionTargetBox="[346,1242,198,1469]" captionTargetId="figure@26.[346,1243,194,1469]" captionTargetPageId="26" captionText="FIGURES 85 88. Nests of Polistes species of subgenus Polistella occurring in northern Vietnam. 85. P. dawnae (Nest # 2009 - DB-P- 01). The thick and thin arrows indicate the position of the primary petiole and the subsidiary petioles, respectively. 86. P. l e p c h a (Nest # VN-P- 2006 - 3). 87. P. reliciniclypeus sp. nov. (Nest # VN-P- 2006 - 13). 88. P. gilvus sp. nov. (Nest # 2008 - SL-P- 1)." httpUri="https://zenodo.org/record/201103/files/figure.png" pageId="5" pageNumber="6">Fig. 85</figureCitation>
); comb vertically elongate to fit the space inside the bamboo, about
<quantity id="906F9B04FFD1B914BBE0E1FCAF5BFA25" box="[709,801,1419,1443]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="6" unit="mm" value="170.0">170 mm</quantity>
long and about
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wide, tough paper-like in texture, uniformly grayish brown. Four nest petioles present (indicated by arrows in
<figureCitation id="CFAC2A64FFD1B914BD24E1D8A82FFA41" box="[1025,1109,1455,1479]" captionStart="FIGURES 85 88" captionStartId="26.[151,269,1501,1525]" captionTargetBox="[346,1242,198,1469]" captionTargetId="figure@26.[346,1243,194,1469]" captionTargetPageId="26" captionText="FIGURES 85 88. Nests of Polistes species of subgenus Polistella occurring in northern Vietnam. 85. P. dawnae (Nest # 2009 - DB-P- 01). The thick and thin arrows indicate the position of the primary petiole and the subsidiary petioles, respectively. 86. P. l e p c h a (Nest # VN-P- 2006 - 3). 87. P. reliciniclypeus sp. nov. (Nest # VN-P- 2006 - 13). 88. P. gilvus sp. nov. (Nest # 2008 - SL-P- 1)." httpUri="https://zenodo.org/record/201103/files/figure.png" pageId="5" pageNumber="6">Fig. 85</figureCitation>
); primary petiole (lost while collecting the nest) situated near mid-length of comb, first two cells having been constructed on its tip; other three petioles subsidiary, constructed to connect bottom of preexisting cell and substrate, made exclusively with adult salivary secretion, dark brown and lustrous; one of the subsidiary petioles situated about
<quantity id="906F9B04FFD1B914BDB2E26CA892F9B5" box="[1175,1256,1563,1587]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="6" unit="mm" value="34.0">34 mm</quantity>
lower from the primary petiole,
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long and 0.96 ×
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thick; remaining petioles respectively about
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and
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above the primary petiole, and
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long and 0.94 ×
<quantity id="906F9B04FFD1B914BA29E214AF19F9FD" box="[780,867,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="6" unit="mm" value="1.6">1.6 mm</quantity>
thick, and
<quantity id="906F9B04FFD1B914BAC6E214A840F9FD" box="[995,1082,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="6" unit="mm" value="4.2">4.2 mm</quantity>
long and 0.43 ×
<quantity id="906F9B04FFD1B914BDDBE214A92FF9FD" box="[1278,1365,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="6" unit="mm" value="1.1">1.1 mm</quantity>
thick. Cells generally arranged in regular rows, hexagonal at open end, but partly irregularly arranged, where some cells are pentagonal; outer free margins of peripheral cells rounded; cell gradually expanded towards open end,
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(range 2.93.6; n=10) wide at bottom and
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(range 4.0
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; n=10) wide at open end,
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(range
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; n=10) deep; cell wall about
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thick; cocoon cap slightly domed beyond level of cell opening, pale gray.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD1B914B9E2E34DAF2FF8D5" box="[199,853,1850,1876]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="572836E1FFD1B914B9E2E34DAF2FF8D5" blockId="5.[151,1437,1275,1876]" box="[199,853,1850,1876]" pageId="5" pageNumber="6">
<emphasis id="65E3EAF3FFD1B914B9E2E34DAD1AF8D5" bold="true" box="[199,352,1850,1875]" pageId="5" pageNumber="6">Distribution.</emphasis>
Northern
<collectingCountry id="2F807671FFD1B914B8F6E34BAE39F8D2" box="[467,579,1852,1876]" name="Myanmar" pageId="5" pageNumber="6">Myanmar</collectingCountry>
; Northwestern
<collectingCountry id="2F807671FFD1B914BBCBE34CAF2AF8D5" box="[750,848,1851,1875]" name="Vietnam" pageId="5" pageNumber="6">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="71F2F291003C7DA8F45CAED2F88C21FE">Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species</mods:title>
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<mods:namePart id="6E07BCB05189EAA364D7A706CB277975">Nguyen, Lien Thi Phuong</mods:namePart>
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<paragraph id="572836E1FFD9B91CB9B2E6F8AF1CFD2F" blockId="13.[151,870,655,714]" box="[151,870,655,681]" pageId="13" pageNumber="14">
<heading id="0C60818DFFD9B91CB9B2E6F8AF1CFD2F" bold="true" box="[151,870,655,681]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="65E3EAF3FFD9B91CB9B2E6F8AF1CFD2F" bold="true" box="[151,870,655,681]" pageId="13" pageNumber="14">
<taxonomicName id="90974D62FFD9B91CB9B2E6F8AE86FD2F" authority="Nguyen, Kojima &amp; Saito" authorityName="Nguyen, Kojima &amp; Saito" box="[151,764,655,681]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="paco" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFD9B91CB9B2E6F8AC8EFD2F" bold="true" box="[151,244,655,681]" italics="true" pageId="13" pageNumber="14">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFD9B91CB821E6F8AD0EFD2F" bold="true" box="[260,372,655,681]" italics="true" pageId="13" pageNumber="14">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFD9B91CB8A3E6F8ADC5FD2F" bold="true" box="[390,447,655,681]" italics="true" pageId="13" pageNumber="14">paco</emphasis>
Nguyen, Kojima &amp; Saito
</taxonomicName>
,
<taxonomicNameLabel id="7ED05788FFD9B91CBA2CE6F8AF1CFD2F" box="[777,870,655,681]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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(
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)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9B2E68CAFE2FCB0" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="572836E1FFD9B91CB9B2E68CAFE2FCB0" blockId="13.[151,1437,762,1831]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9B2E68CAD3CFC95" bold="true" box="[151,326,762,787]" pageId="13" pageNumber="14">
<typeStatus id="882C8843FFD9B91CB9B2E68CACAAFC95" box="[151,208,763,787]" pageId="13" pageNumber="14">Type</typeStatus>
material.
</emphasis>
<typeStatus id="882C8843FFD9B91CB876E68DAD91FC94" box="[339,491,762,786]" pageId="13" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
: Ƥ (
<collectionCode id="3186AE24FFD9B91CBB3BE68DAE12FC94" box="[542,616,762,786]" pageId="13" pageNumber="14">IUNH</collectionCode>
, on long-term loan from the
<collectionCode id="3186AE24FFD9B91CBAE5E68DA87EFC94" box="[960,1028,762,786]" pageId="13" pageNumber="14">IEBR</collectionCode>
), labeled “
<collectingCountry id="2F807671FFD9B91CBDA4E68DA97DFC94" box="[1153,1287,762,786]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
, Pa Co, Mai Chau, Hoa Binh,
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, L.T.P. Nguyen &amp; J. Kojima,
<date id="23291021FFD9B91CBBD8E769AFF3FCB0" box="[765,905,798,822]" pageId="13" pageNumber="14" value="2005-08-23">23.viii.2005</date>
”.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9E2E735A828FC6C" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="572836E1FFD9B91CB9E2E735A828FC6C" blockId="13.[151,1437,762,1831]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E735AD3BFCDD" bold="true" box="[199,321,834,859]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
This species can be distinguished from all other known species of subgenus
<emphasis id="65E3EAF3FFD9B91CBDFEE734A93DFCDD" box="[1243,1351,835,859]" italics="true" pageId="13" pageNumber="14">Polistella</emphasis>
by the following combination of characters: female clypeus convex; sctuellum strongly depressed medially; propodeum with weak striae; metasomal tergum 1 short and thick, about 0.8 × as long as its apical width, in lateral view abruptly swollen dorsally at base, then broadly convex dorsally; metasomal sternum
<quantity id="906F9B04FFD9B91CBD51E7D9A8D9FC40" box="[1140,1187,942,966]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="14" unit="in" value="2.0">2 in</quantity>
lateral view abruptly swollen ventrally in anterior one-third, then ventral margin rounded into straight line.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9E2E781ADE0F9C8" pageId="13" pageNumber="14" type="description">
<paragraph id="572836E1FFD9B91CB9E2E781A894FB88" blockId="13.[151,1437,762,1831]" box="[199,1262,1014,1039]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E781AD23FB89" bold="true" box="[199,345,1014,1039]" pageId="13" pageNumber="14">Description.</emphasis>
FEMALE. Body length about
<quantity id="906F9B04FFD9B91CBB96E781AF76FB88" box="[691,780,1014,1038]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="14" unit="mm" value="9.5">9.5 mm</quantity>
; fore wing length about
<quantity id="906F9B04FFD9B91CBD39E781A808FB88" box="[1052,1138,1014,1038]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="14" unit="mm" value="9.5">9.5 mm</quantity>
(
<typeStatus id="882C8843FFD9B91CBDA4E781A89CFB88" box="[1153,1254,1014,1038]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="572836E1FFD9B91CB9E2E06DAF9AFA1C" blockId="13.[151,1437,762,1831]" pageId="13" pageNumber="14">
Structure as in
<taxonomicName id="90974D62FFD9B91CB853E06CAE06FBB5" box="[374,636,1051,1075]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFD9B91CB853E06CAE06FBB5" box="[374,636,1051,1075]" italics="true" pageId="13" pageNumber="14">Polistes reliciniclypeus</emphasis>
</taxonomicName>
, but differing as follows: POD:OOD=1:2; POD about 1.3 × Od (
<figureCitation id="CFAC2A64FFD9B91CBC4BE06DACCDFBD0" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 42</figureCitation>
); inner eye margins less strongly convergent ventrally, about 1.1 × further apart from each other at vertex than at clypeus (
<figureCitation id="CFAC2A64FFD9B91CB9D8E015AD2BFBFC" box="[253,337,1122,1146]" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 43</figureCitation>
); interantennal space strongly raised; clypeus in frontal view as wide as high (
<figureCitation id="CFAC2A64FFD9B91CBDEBE015A958FBFC" box="[1230,1314,1122,1146]" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 43</figureCitation>
), in lateral view distinctly convex (
<figureCitation id="CFAC2A64FFD9B91CB88AE0F1AE7DFB18" box="[431,519,1158,1182]" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 44</figureCitation>
), lateral margin lying along inner eye margin slightly shorter than diameter of antennal socket; gena wider, in lateral view nearly as wide as eye (
<figureCitation id="CFAC2A64FFD9B91CBAA1E0DDAFA3FB44" box="[900,985,1194,1218]" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 44</figureCitation>
); antennal flagellomere 1 about 2.5 × as long as its apical width (
<figureCitation id="CFAC2A64FFD9B91CB882E0B9AD81FB60" box="[423,507,1230,1254]" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 45</figureCitation>
); scutellum strongly depressed medially; propodeal orifice elongate, about 1.9 × as long as wide (measured at widest part), somewhat narrowed in dorsal half; metasomal tergum 1 about 0.8 × as long as its apical width, in lateral view abruptly swollen dorsally just behind basal slit for reception of propodeal suspensory ligament, then dorsal face broadly convex; corner between anterior and dorsal faces rounded (
<figureCitation id="CFAC2A64FFD9B91CBC10E14DA9F5FAD4" box="[1333,1423,1338,1362]" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 46</figureCitation>
); metasomal sternum
<quantity id="906F9B04FFD9B91CB85EE129ADD2FAF0" box="[379,424,1374,1398]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="14" unit="in" value="2.0">2 in</quantity>
lateral view basally abruptly swollen ventrally in anterior one-third, then ventral margin broadly rounded into straight line in posterior half of the sternum (
<figureCitation id="CFAC2A64FFD9B91CBA5AE1F5AFAEFA1C" box="[895,980,1410,1434]" captionStart="FIGURES 42 46" captionStartId="14.[151,269,1173,1197]" captionTargetBox="[191,1396,189,1149]" captionTargetId="figure@14.[191,1396,189,1149]" captionTargetPageId="14" captionText="FIGURES 42 46. Polistes (Polistella) paco sp. nov., Ƥ. 42. Ocelli, eye and posterior margin of vertex. 43 44. Head. 43. Frontal view. 44. Lateral view. 45. Right antenna. 46. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201097/files/figure.png" pageId="13" pageNumber="14">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="572836E1FFD9B91CB9E2E1D1AE0EF980" blockId="13.[151,1437,762,1831]" pageId="13" pageNumber="14">
Body surface sculpture as in
<taxonomicName id="90974D62FFD9B91CBB2CE1DFAEB2FA38" box="[521,712,1446,1471]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFD9B91CBB2CE1DFAEB2FA38" box="[521,712,1446,1471]" italics="true" pageId="13" pageNumber="14">P. reliciniclypeus</emphasis>
</taxonomicName>
, but clypeus except margins with scattered deep large punctures, each of which bears a golden sharply pointed bristle, apical margin of metanotum smooth and without punctures, propodeum with weak transverse striation.
</paragraph>
<paragraph id="572836E1FFD9B91CB9E2E264AE66F9AC" blockId="13.[151,1437,762,1831]" box="[199,540,1554,1579]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E264AD77F9AD" box="[199,269,1555,1579]" italics="true" pageId="13" pageNumber="14">Color.</emphasis>
As in
<taxonomicName id="90974D62FFD9B91CB873E263AE6FF9AD" box="[342,533,1555,1579]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFD9B91CB873E263AE6FF9AD" box="[342,533,1555,1579]" italics="true" pageId="13" pageNumber="14">P. reliciniclypeus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="572836E1FFD9B91CB9E2E240ADE0F9C8" blockId="13.[151,1437,762,1831]" box="[199,410,1590,1615]" pageId="13" pageNumber="14">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9E2E22CAF24F958" pageId="13" pageNumber="14" type="discussion">
<paragraph id="572836E1FFD9B91CB9E2E22CAF24F958" blockId="13.[151,1437,762,1831]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E22CAD4AF9F5" bold="true" box="[199,304,1627,1651]" pageId="13" pageNumber="14">Remark.</emphasis>
This species is most similar to
<taxonomicName id="90974D62FFD9B91CBBB9E22BAF27F9F5" box="[668,861,1627,1651]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFD9B91CBBB9E22BAF27F9F5" box="[668,861,1627,1651]" italics="true" pageId="13" pageNumber="14">P. reliciniclypeus</emphasis>
</taxonomicName>
, but can be easily distinguished from the latter by having the female clypeus in lateral view evenly curved anteriorly, not prominently swollen anteriorly in upper part, the scutellum strongly depressed medially, the second metasomal sternum with the anteroventral corner broadly rounded, and the striation on propodeum much weaker.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9E2E29CA96CF884" box="[199,1302,1770,1795]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="572836E1FFD9B91CB9E2E29CA96CF884" blockId="13.[151,1437,762,1831]" box="[199,1302,1770,1795]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E29CAD34F885" bold="true" box="[199,334,1771,1795]" pageId="13" pageNumber="14">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="882C8843FFD9B91CBB93E29CAE9CF885" box="[694,742,1771,1795]" pageId="13" pageNumber="14">type</typeStatus>
locality, Pa Co in Hoa Binh Province of
<collectingCountry id="2F807671FFD9B91CBD95E29DA969F884" box="[1200,1299,1770,1794]" name="Vietnam" pageId="13" pageNumber="14">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9E2E379AFBEF8A0" box="[199,964,1806,1831]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="572836E1FFD9B91CB9E2E379AFBEF8A0" blockId="13.[151,1437,762,1831]" box="[199,964,1806,1831]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E379AD1AF8A1" bold="true" box="[199,352,1806,1831]" pageId="13" pageNumber="14">Distribution.</emphasis>
Known only from the
<typeStatus id="882C8843FFD9B91CBB45E378AEEAF8A1" box="[608,656,1807,1831]" pageId="13" pageNumber="14">type</typeStatus>
locality, northern
<collectingCountry id="2F807671FFD9B91CBA78E379AFBAF8A0" box="[861,960,1806,1830]" name="Vietnam" pageId="13" pageNumber="14">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,233 @@
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<mods:title id="180F27B83327B1EDFC860122BD6A124F">Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species</mods:title>
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<treatment id="DF3E87F7FFDAB901B9B2E153AFD7FBFA" ID-DOI="http://doi.org/10.5281/zenodo.5489174" ID-GBIF-Taxon="119419228" ID-Zenodo-Dep="5489174" LSID="urn:lsid:plazi:treatment:DF3E87F7FFDAB901B9B2E153AFD7FBFA" httpUri="http://treatment.plazi.org/id/DF3E87F7FFDAB901B9B2E153AFD7FBFA" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="1F8D656AFFDAB91FB9B2E153AD5DFAD9" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="572836E1FFDAB91FB9B2E153AFACFAB8" blockId="14.[151,982,1316,1375]" box="[151,982,1316,1342]" pageId="14" pageNumber="15">
<heading id="0C60818DFFDAB91FB9B2E153AFACFAB8" bold="true" box="[151,982,1316,1342]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="65E3EAF3FFDAB91FB9B2E153AFACFAB8" bold="true" box="[151,982,1316,1342]" pageId="14" pageNumber="15">
<taxonomicName id="90974D62FFDAB91FB9B2E153AF16FAB8" authority="Nguyen, Kojima &amp; Saito" authorityName="Nguyen, Kojima &amp; Saito" box="[151,876,1316,1342]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="curcipunctum" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFDAB91FB9B2E153AC8EFAB8" bold="true" box="[151,244,1316,1342]" italics="true" pageId="14" pageNumber="15">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFDAB91FB821E153AD0EFAB8" bold="true" box="[260,372,1316,1342]" italics="true" pageId="14" pageNumber="15">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFDAB91FB8A3E153AE55FAB8" bold="true" box="[390,559,1316,1342]" italics="true" pageId="14" pageNumber="15">curcipunctum</emphasis>
Nguyen, Kojima &amp; Saito
</taxonomicName>
,
<taxonomicNameLabel id="7ED05788FFDAB91FBA5CE153AFACFAB8" box="[889,982,1316,1342]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="572836E1FFDAB91FB9B2E130AD5DFAD9" blockId="14.[151,982,1316,1375]" box="[151,295,1351,1375]" pageId="14" pageNumber="15">
(
<figureCitation id="CFAC2A64FFDAB91FB9BAE130AD65FAD9" box="[159,287,1351,1375]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="14" pageNumber="15">Figs 4753</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFDAB91FB9B2E1F8AD02F9B1" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="572836E1FFDAB91FB9B2E1F8AD02F9B1" blockId="14.[151,1436,1422,2023]" pageId="14" pageNumber="15">
<emphasis id="65E3EAF3FFDAB91FB9B2E1F8AD3DFA21" bold="true" box="[151,327,1422,1447]" pageId="14" pageNumber="15">
<typeStatus id="882C8843FFDAB91FB9B2E1F8ACA8FA21" box="[151,210,1423,1447]" pageId="14" pageNumber="15">Type</typeStatus>
material.
</emphasis>
<typeStatus id="882C8843FFDAB91FB874E1F8AD93FA21" box="[337,489,1423,1447]" pageId="14" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
: Ƥ (
<collectionCode id="3186AE24FFDAB91FBB3CE1F8AE1BFA21" box="[537,609,1423,1447]" pageId="14" pageNumber="15">IUNH</collectionCode>
on long-term loan from the
<collectionCode id="3186AE24FFDAB91FBA8BE1F8AF88FA21" box="[942,1010,1423,1447]" pageId="14" pageNumber="15">IEBR</collectionCode>
), labeled “
<collectingCountry id="2F807671FFDAB91FBD49E1F8A888FA21" box="[1132,1266,1423,1447]" name="Vietnam" pageId="14" pageNumber="15">VIETNAM</collectingCountry>
, Tam Dao NP, Vinh Phuc,
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, L.T.P. Nguyen &amp; J. Kojima,
<date id="23291021FFDAB91FBB8AE1C4AF41FA4D" box="[687,827,1459,1483]" pageId="14" pageNumber="15" value="2005-08-19">19.viii.2005</date>
”.
<typeStatus id="882C8843FFDAB91FBA74E1C4AF8AFA4D" box="[849,1008,1459,1483]" pageId="14" pageNumber="15" type="paratype">PARATYPES</typeStatus>
: 1 Ƥ (
<collectionCode id="3186AE24FFDAB91FBD15E1C4A80FFA4D" box="[1072,1141,1459,1483]" pageId="14" pageNumber="15">IEBR</collectionCode>
), same data as
<typeStatus id="882C8843FFDAB91FBC38E1C4A9F9FA4D" box="[1309,1411,1459,1483]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
; 1 Ƥ (
<collectionCode id="3186AE24FFDAB91FB999E1A0AD62FA69" LSID="urn:lsid:biocol.org:col:34925" box="[188,280,1495,1519]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="15">AMNH</collectionCode>
), Huong Son, Ha Tinh,
<geoCoordinate id="32A35026FFDAB91FBB1CE1A0AEECFA69" box="[569,662,1495,1519]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="18.35">18°21'N</geoCoordinate>
<geoCoordinate id="32A35026FFDAB91FBB87E1A0AF76FA69" box="[674,780,1495,1519]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="106.25">106°15'E</geoCoordinate>
, alt.
<quantity id="906F9B04FFDAB91FBA6EE1A0AFE2FA69" box="[843,920,1495,1519]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="15" unit="m" value="900.0">900 m</quantity>
, Malaise Trap,
<date id="23291021FFDAB91FBD71E1A0A8B3FA69" box="[1108,1225,1495,1519]" pageId="14" pageNumber="15" value="1998-05-18">18.V.1998</date>
, Carpenter, Long, Grimaldi, Herman and Silva; 3 Ƥ (
<collectionCode id="3186AE24FFDAB91FBB6CE18CAEF7F995" box="[585,653,1531,1555]" pageId="14" pageNumber="15">IEBR</collectionCode>
,
<collectionCode id="3186AE24FFDAB91FBB85E18BAE93F995" box="[672,745,1532,1555]" pageId="14" pageNumber="15">IUNH</collectionCode>
,
<collectionCode id="3186AE24FFDAB91FBBD8E18CAF2DF995" LSID="urn:lsid:biocol.org:col:34925" box="[765,855,1531,1555]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="15">AMNH</collectionCode>
), Lo Xo, Phuoc Son, Quang Nam, alt.
<quantity id="906F9B04FFDAB91FBC63E18CA9EDF995" box="[1350,1431,1531,1555]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="m" value="700.0">700 m</quantity>
,
<date id="23291021FFDAB91FB9B2E268AD58F9B1" box="[151,290,1567,1591]" pageId="14" pageNumber="15" value="2004-07-28">28.VII.2004</date>
, ISD-c.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFDAB91FB9E2E234AF5DF8B5" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="572836E1FFDAB91FB9E2E234AF5DF8B5" blockId="14.[151,1436,1422,2023]" pageId="14" pageNumber="15">
<emphasis id="65E3EAF3FFDAB91FB9E2E234AD3BF9DA" bold="true" box="[199,321,1603,1628]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
This species can be distinguished from other known species of subgenus
<emphasis id="65E3EAF3FFDAB91FBD56E234A8A4F9DD" box="[1139,1246,1603,1627]" italics="true" pageId="14" pageNumber="15">Polistella</emphasis>
by the following combination of characters: body surface with very coarse and large punctures; posterior face of propodeum with wide median excavation (width about 0.7 × maximum width of propodeum); border between posterior and lateral faces of propodeum bluntly angulate; metasomal tergum 1 short and robust, about 0.7 × as long as its apical width, in lateral view abruptly swollen dorsally at base, then dorsal face weakly convex, corner between anterior and dorsal faces broadly rounded; metasomal sternum
<quantity id="906F9B04FFDAB91FBB8FE280AEACF889" box="[682,726,1783,1807]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="15" unit="in" value="2.0">2 in</quantity>
lateral view with square outline, basally abruptly swollen ventrally, then ventral margin bluntly angled into straight line.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFDAB901B9E2E349ADE0FC6A" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="572836E1FFDAB91FB9E2E349AD87F8FD" blockId="14.[151,1436,1422,2023]" pageId="14" pageNumber="15">
<emphasis id="65E3EAF3FFDAB91FB9E2E349AD23F8D1" bold="true" box="[199,345,1854,1879]" pageId="14" pageNumber="15">Description.</emphasis>
FEMALE. Body length
<quantity id="906F9B04FFDAB91FBB5CE348AF64F8D1" box="[633,798,1855,1879]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="14" pageNumber="15" unit="mm" value="12.0" valueMax="12.5" valueMin="11.5">11.512.5 mm</quantity>
(
<typeStatus id="882C8843FFDAB91FBA15E348AFE2F8D1" box="[816,920,1855,1879]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFDAB91FBAC9E348A822F8D1" box="[1004,1112,1855,1879]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="15" unit="mm" value="12.5">12.5 mm</quantity>
); forewing length
<quantity id="906F9B04FFDAB91FBC08E348ACB9F8FD" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="14" pageNumber="15" unit="mm" value="12.0" valueMax="12.5" valueMin="11.5">11.512.5 mm</quantity>
(
<typeStatus id="882C8843FFDAB91FB9F4E314AD43F8FD" box="[209,313,1891,1915]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFDAB91FB8ADE314AD8AF8FD" box="[392,496,1891,1915]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="15" unit="mm" value="12.5">12.5 mm</quantity>
).
</paragraph>
<paragraph id="572836E1FFDAB901B9E2E3F0A94EFE9B" blockId="14.[151,1436,1422,2023]" lastBlockId="16.[151,1436,151,1149]" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
Structure as in
<taxonomicName id="90974D62FFDAB91FB853E3F0AE06F819" box="[374,636,1927,1951]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFDAB91FB853E3F0AE06F819" box="[374,636,1927,1951]" italics="true" pageId="14" pageNumber="15">Polistes reliciniclypeus</emphasis>
</taxonomicName>
, but differing as follows: POD:OOD=1:2; POD about 1.3 × Od (
<figureCitation id="CFAC2A64FFDAB91FBC4BE3F0ACCDF845" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="14" pageNumber="15">Fig. 47</figureCitation>
); inner eye margins in frontal view less strongly convergent ventrally, about 1.1 × further apart from each other at vertex than at clypeus (
<figureCitation id="CFAC2A64FFDAB91FB8E0E3B8AE66F861" box="[453,540,1999,2023]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="14" pageNumber="15">Fig. 48</figureCitation>
); clypeus in frontal view slightly wider than high (
<figureCitation id="CFAC2A64FFDAB91FBD4BE3B8A8BFF861" box="[1134,1221,1999,2023]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="14" pageNumber="15">Fig. 48</figureCitation>
); lateral margin of clypeus touching inner eye margin by length longer than malar space and antennal socket diameter; gena proportionally narrower, in lateral view about 0.6 × as wide as eye (
<figureCitation id="CFAC2A64FFDBB91EBAAAE4CBAF9EFF52" box="[911,996,188,212]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="15" pageNumber="16">Fig. 49</figureCitation>
); antennal scape slightly more than 3 × as long as its maximum width (
<figureCitation id="CFAC2A64FFDBB91EB8D9E4A8AE28FF71" box="[508,594,223,247]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="15" pageNumber="16">Fig. 50</figureCitation>
); flagellomere 1 slightly greater than 2.5 × as long as its maximum width; flagellomere 2 longer than wide; terminal flagellomere about 1.3 × as long as its basal width; mesoscutum weakly convex; scutellum strongly convex (
<figureCitation id="CFAC2A64FFDBB91EBB10E550AEF3FEB9" box="[565,649,295,319]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="15" pageNumber="16">Fig. 51</figureCitation>
), with median part weakly depressed or nearly flat; posterior face of propodeum with shallow wide median excavation, of which width is about 0.7 × maximum width of propodeum, border between posterior and lateral faces bluntly angulate; propodeal orifice elongate, about 2 × as long as wide (measured at widest part); jugal lobe of hind wing rounded (
<figureCitation id="CFAC2A64FFDBB91EBA65E5E3AFECFE2A" box="[832,918,404,428]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="15" pageNumber="16">Fig. 52</figureCitation>
); metasomal tergum 1 short and robust, about 0.7 × as long as its apical width, in lateral view abruptly swollen dorsally just behind basal slit for reception of propodeal suspensory ligament, then dorsal face weakly convex; corner between anterior and dorsal faces broadly rounded (
<figureCitation id="CFAC2A64FFC4B901B827E4A8AD2FFF71" box="[258,341,223,247]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="16" pageNumber="17">Fig. 53</figureCitation>
); metasomal sternum
<quantity id="906F9B04FFC4B901BB6EE4A8AE0CFF71" box="[587,630,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
lateral view with square outline (
<figureCitation id="CFAC2A64FFC4B901BACEE4A8A844FF71" box="[1003,1086,223,247]" captionStart="FIGURES 47 53" captionStartId="15.[151,269,1923,1947]" captionTargetBox="[191,1396,483,1895]" captionTargetId="figure@15.[191,1396,483,1895]" captionTargetPageId="15" captionText="FIGURES 47 53. Polistes (Polistella) curcipunctum sp. nov.,. 47. Ocelli, eye and posterior margin of vertex. 48 49. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="16" pageNumber="17">Fig. 53</figureCitation>
), basally abruptly swollen ventrally, then ventral margin bluntly angled into straight line nearly parallel to ventral margin of the tergum.
</paragraph>
<caption id="03E86669FFDBB91EB9B2E3F4AF75F85C" httpUri="https://zenodo.org/record/201098/files/figure.png" pageId="15" pageNumber="16" targetBox="[191,1396,483,1895]" targetPageId="15">
<paragraph id="572836E1FFDBB91EB9B2E3F4AF75F85C" blockId="15.[151,1437,1923,2010]" pageId="15" pageNumber="16">
<emphasis id="65E3EAF3FFDBB91EB9B2E3F4AD27F81D" bold="true" box="[151,349,1923,1947]" pageId="15" pageNumber="16">FIGURES 4753.</emphasis>
<taxonomicName id="90974D62FFDBB91EB846E3F4AEB4F81C" box="[355,718,1923,1946]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="curcipunctum" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFDBB91EB846E3F4ADC2F81C" box="[355,440,1923,1946]" italics="true" pageId="15" pageNumber="16">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFDBB91EB8E3E3F4AE56F81C" box="[454,556,1923,1946]" italics="true" pageId="15" pageNumber="16">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFDBB91EBB19E3F4AEB4F81C" box="[572,718,1923,1946]" italics="true" pageId="15" pageNumber="16">curcipunctum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFDBB91EBBF1E3F4AF52F81D" box="[724,808,1923,1947]" pageId="15" pageNumber="16" rank="species">
<emphasis id="65E3EAF3FFDBB91EBBF1E3F4AF52F81D" bold="true" box="[724,808,1923,1947]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
,. 47. Ocelli, eye and posterior margin of vertex. 4849. Head. 48. Frontal view. 49. Lateral view. 50. Right antenna. 51. Mesosoma, lateral view. 52. Jugal lobe of left hind wing. 53. Metasomal segments 1 and 2, lateral view. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="572836E1FFC4B901B9E2E550ADDBFD21" blockId="16.[151,1436,151,1149]" pageId="16" pageNumber="17">
Body surface with much coarser punctures than in
<emphasis id="65E3EAF3FFC4B901BA04E55EAF8DFEC6" box="[801,1015,296,320]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="90974D62FFC4B901BA04E55EAF89FEC6" box="[801,1011,296,320]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="reliciniclypeus">P. reliciniclypeus</taxonomicName>
.
</emphasis>
Clypeus densely covered with very coarse large punctures, each bearing sharply pointed golden bristle; basal one-third to half of clypeus covered with fine pale-yellow pubescence. Mandible with scattered deep punctures, punctures denser at basal margin. Frons and vertex anterior to level of posterior ocelli densely covered with very coarse deep punctures; punctures around ocelli less coarse; vertex behind posterior ocelli and gena densely covered with very coarse punctures. Pronotum with punctures similar to those in
<taxonomicName id="90974D62FFC4B901B8CDE5AAAED0FE72" box="[488,682,476,500]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="reliciniclypeus">
<emphasis id="65E3EAF3FFC4B901B8CDE5AAAED0FE72" box="[488,682,476,500]" italics="true" pageId="16" pageNumber="17">P. reliciniclypeus</emphasis>
</taxonomicName>
(dense and deep; spaces between punctures very narrow, slightly raised to form reticulation). Scutellum, metanotum and posterodorsal part of mesepisternum with punctures similar to those on pronotum. Dorsal metapleuron with striae and strong punctures; ventral metapleuron with dense coarse punctures. Propodeum with distinct transverse striae with coarse punctures between striae; lateral face with welldefined punctures between striae, interpuncture spaces weakly raised to interconnect striae. Metasomal terga with dense coarse punctures.
</paragraph>
<paragraph id="572836E1FFC4B901B9E2E6C3AC9CFC41" blockId="16.[151,1436,151,1149]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9E2E6C3AD77FD4A" box="[199,269,692,716]" italics="true" pageId="16" pageNumber="17">Color.</emphasis>
Body black, with following parts orange-yellow: apical one to to two-thirds of clypeus, narrow band along inner eye margin extending from bottom of frons to middle of eye emargination, narrow short band on gena along posterodorsal margin of eye, narrow band along pronotal carina, paired spots on propodeal valvula; reddish brown as follows: mandible except black basal margin and teeth, ventral side of antennal scape, pedicel and basal part of flagellomere 1, sometimes posterolateral spots on scutellum and narrow apical band on metasomal tergum 2. Legs black; dorsal surface of femora, ventral surface of tibiae and tarsomeres 1 largely reddish brown. Wings yellowish-brown, weakly infuscate, slightly more strongly infuscate along anterior margin of forewing; veins dark brown.
</paragraph>
<paragraph id="572836E1FFC4B901B9E2E7A3ADE0FC6A" blockId="16.[151,1436,151,1149]" box="[199,410,980,1004]" pageId="16" pageNumber="17">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC4B901B9E2E78FADABFBB2" pageId="16" pageNumber="17" type="discussion">
<paragraph id="572836E1FFC4B901B9E2E78FADABFBB2" blockId="16.[151,1436,151,1149]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9E2E78FAD4AFB96" bold="true" box="[199,304,1016,1040]" pageId="16" pageNumber="17">Remark.</emphasis>
Among the species of subgenus
<emphasis id="65E3EAF3FFC4B901BB8FE78FAF6CFB96" box="[682,790,1016,1040]" italics="true" pageId="16" pageNumber="17">Polistella</emphasis>
occurring in
<collectingCountry id="2F807671FFC4B901BA94E780A869FB89" box="[945,1043,1015,1039]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
, this species can be recognized by extremely corase punctures.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC4B901B9E2E037A8A9FBDE" box="[199,1235,1088,1112]" pageId="16" pageNumber="17" type="etymology">
<paragraph id="572836E1FFC4B901B9E2E037A8A9FBDE" blockId="16.[151,1436,151,1149]" box="[199,1235,1088,1112]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9E2E037AD34FBDE" bold="true" box="[199,334,1088,1112]" pageId="16" pageNumber="17">Etymology.</emphasis>
The specific name refer to the coarse (
<emphasis id="65E3EAF3FFC4B901BA27E036AF36FBDE" box="[770,844,1089,1112]" italics="true" pageId="16" pageNumber="17">curcus</emphasis>
in Latin) punctures in this species.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFC4B901B9E2E014AFD7FBFA" box="[199,941,1123,1149]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="572836E1FFC4B901B9E2E014AFD7FBFA" blockId="16.[151,1436,151,1149]" box="[199,941,1123,1149]" pageId="16" pageNumber="17">
<emphasis id="65E3EAF3FFC4B901B9E2E014AD1AFBFA" bold="true" box="[199,352,1123,1148]" pageId="16" pageNumber="17">Distribution.</emphasis>
Known only from
<typeStatus id="882C8843FFC4B901BB12E012AE1DFBFB" box="[567,615,1125,1149]" pageId="16" pageNumber="17">type</typeStatus>
localities, northern
<collectingCountry id="2F807671FFC4B901BA62E013AFD0FBFA" box="[839,938,1124,1148]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,521 @@
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<mods:title id="96C82F762E127B552A83D484B78557FD">Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species</mods:title>
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<subSubSection id="1F8D656AFFDCB919B9B2E282AD2AF8A9" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="572836E1FFDCB919B9B2E282AFA7F888" blockId="8.[151,989,1780,1839]" box="[151,989,1780,1807]" pageId="8" pageNumber="9">
<heading id="0C60818DFFDCB919B9B2E282AFA7F888" bold="true" box="[151,989,1780,1807]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="65E3EAF3FFDCB919B9B2E282AFA7F888" bold="true" box="[151,989,1780,1807]" pageId="8" pageNumber="9">
<taxonomicName id="90974D62FFDCB919B9B2E282AF08F888" authority="Nguyen, Kojima &amp; Saito" authorityName="Nguyen, Kojima &amp; Saito" box="[151,882,1780,1807]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="reliciniclypeus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFDCB919B9B2E282AC8EF889" bold="true" box="[151,244,1781,1807]" italics="true" pageId="8" pageNumber="9">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFDCB919B821E282AD0EF889" bold="true" box="[260,372,1781,1807]" italics="true" pageId="8" pageNumber="9">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFDCB919B8A3E282AE4CF889" bold="true" box="[390,566,1781,1807]" italics="true" pageId="8" pageNumber="9">reliciniclypeus</emphasis>
Nguyen, Kojima &amp; Saito
</taxonomicName>
,
<taxonomicNameLabel id="7ED05788FFDCB919BA5AE283AFA7F888" box="[895,989,1780,1806]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="572836E1FFDCB919B9B2E360AD2AF8A9" blockId="8.[151,989,1780,1839]" box="[151,336,1815,1839]" pageId="8" pageNumber="9">
(
<figureCitation id="CFAC2A64FFDCB919B9BAE360AD5AF8A9" box="[159,288,1815,1839]" captionStart-0="FIGURES 27 33" captionStart-1="FIGURES 34 41" captionStartId-0="10.[151,269,1754,1778]" captionStartId-1="11.[151,269,1879,1903]" captionTargetBox-0="[201,1396,411,1729]" captionTargetBox-1="[191,1396,591,1864]" captionTargetId-0="figure@10.[191,1396,411,1729]" captionTargetId-1="figure@11.[191,1396,591,1873]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." captionText-1="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri-0="https://zenodo.org/record/201095/files/figure.png" httpUri-1="https://zenodo.org/record/201096/files/figure.png" pageId="8" pageNumber="9">Figs 2741</figureCitation>
,
<figureCitation id="CFAC2A64FFDCB919B80EE360AD32F8A9" box="[299,328,1815,1839]" captionStart="FIGURES 85 88" captionStartId="26.[151,269,1501,1525]" captionTargetBox="[346,1242,198,1469]" captionTargetId="figure@26.[346,1243,194,1469]" captionTargetPageId="26" captionText="FIGURES 85 88. Nests of Polistes species of subgenus Polistella occurring in northern Vietnam. 85. P. dawnae (Nest # 2009 - DB-P- 01). The thick and thin arrows indicate the position of the primary petiole and the subsidiary petioles, respectively. 86. P. l e p c h a (Nest # VN-P- 2006 - 3). 87. P. reliciniclypeus sp. nov. (Nest # VN-P- 2006 - 13). 88. P. gilvus sp. nov. (Nest # 2008 - SL-P- 1)." httpUri="https://zenodo.org/record/201103/files/figure.png" pageId="8" pageNumber="9">87</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFDCB918B9B2E317AE40FD02" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="572836E1FFDCB918B9B2E317AE40FD02" blockId="8.[151,1436,1887,2020]" lastBlockId="9.[151,1437,151,2012]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="65E3EAF3FFDCB919B9B2E317AD3FF8FE" bold="true" box="[151,325,1887,1912]" pageId="8" pageNumber="9">
<typeStatus id="882C8843FFDCB919B9B2E317ACA8F8FE" box="[151,210,1888,1912]" pageId="8" pageNumber="9">Type</typeStatus>
material.
</emphasis>
<typeStatus id="882C8843FFDCB919B869E328AD9EF8F1" box="[332,484,1887,1911]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
: Ƥ (deposited in the
<collectionCode id="3186AE24FFDCB919BBEEE317AF6EF8F1" box="[715,788,1888,1911]" pageId="8" pageNumber="9">IUNH</collectionCode>
, on long-term loan from the
<collectionCode id="3186AE24FFDCB919BD72E328A8E1F8F1" box="[1111,1179,1887,1911]" pageId="8" pageNumber="9">IEBR</collectionCode>
), labeled “
<collectingCountry id="2F807671FFDCB919BC34E328A9E2F8F1" box="[1297,1432,1887,1911]" name="Vietnam" pageId="8" pageNumber="9">VIETNAM</collectingCountry>
, Binh Lu, Phong Tho, Lai Chau,
<geoCoordinate id="32A35026FFDCB919B8DBE3F4AE20F81D" box="[510,602,1923,1947]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="22.35">22°21'N</geoCoordinate>
<geoCoordinate id="32A35026FFDCB919BB44E3F4AE9AF81D" box="[609,736,1923,1947]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="92" value="103.725">103°43.5'E</geoCoordinate>
, alt.
<quantity id="906F9B04FFDCB919BA31E3F4AF11F81D" box="[788,875,1923,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="m" value="1600.0">1600 m</quantity>
, Nest#: VN-P-
<date id="23291021FFDCB919BD33E3F4A808F81D" box="[1046,1138,1923,1947]" pageId="8" pageNumber="9" value="2006" valueMax="2013">2006-13</date>
, L.T.P. Nguyen, F. Saito &amp; J. Kojima,
<date id="23291021FFDCB919B830E3D0ADDBF839" box="[277,417,1959,1983]" pageId="8" pageNumber="9" value="2006-08-22">22.viii.2006</date>
”.
<typeStatus id="882C8843FFDCB919B89DE3D0AE28F839" box="[440,594,1959,1983]" pageId="8" pageNumber="9" type="paratype">PARATYPES</typeStatus>
(
<collectionCode id="3186AE24FFDCB919BB46E3D0AED1F839" box="[611,683,1959,1983]" pageId="8" pageNumber="9">IUNH</collectionCode>
and
<collectionCode id="3186AE24FFDCB919BBC1E3DFAF47F846" LSID="urn:lsid:biocol.org:col:34925" box="[740,829,1960,1984]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="9">AMNH</collectionCode>
, on long-term loan from the
<collectionCode id="3186AE24FFDCB919BDA2E3D0A8B6F839" box="[1159,1228,1959,1983]" pageId="8" pageNumber="9">IEBR</collectionCode>
):
<collectingCountry id="2F807671FFDCB919BDC5E3DFA91BF846" box="[1248,1377,1960,1984]" name="Vietnam" pageId="8" pageNumber="9">VIETNAM</collectingCountry>
:
<emphasis id="65E3EAF3FFDCB919BC56E3D0ACA2F862" bold="true" pageId="8" pageNumber="9">Lai Chau</emphasis>
: Binh Lu, Phong Tho,
<geoCoordinate id="32A35026FFDCB919B8C1E3BCAE3AF865" box="[484,576,1995,2019]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="22.35">22°21'N</geoCoordinate>
<geoCoordinate id="32A35026FFDCB919BB6CE3BCAEB2F865" box="[585,712,1995,2019]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="92" value="103.725">103°43.5'E</geoCoordinate>
, alt.
<quantity id="906F9B04FFDCB919BA24E3BCAF23F865" box="[769,857,1995,2019]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="m" value="1600.0">1600 m</quantity>
, LN, FS &amp; JK,
<date id="23291021FFDCB919BD37E3BCA8D7F865" box="[1042,1197,1995,2019]" pageId="8" pageNumber="9" value="2006-08-22">22. VIII.2006</date>
[15 3, Nest# VN-P-
<date id="23291021FFDDB918B9B2E4E0AC8DFF29" box="[151,247,151,175]" pageId="9" pageNumber="10" value="2006" valueMax="2010">2006-10</date>
; 3 3, 3 Ƥ, Nest# VN-P-
<date id="23291021FFDDB918BB23E4E0AE1BFF29" box="[518,609,151,175]" pageId="9" pageNumber="10" value="2006" valueMax="2011">2006-11</date>
; 3 Ƥ, Nest# VN-P-
<date id="23291021FFDDB918BA1CE4E0AFE2FF29" box="[825,920,151,175]" pageId="9" pageNumber="10" value="2006" valueMax="2012">2006-12</date>
]; 15 3, 11 Ƥ, same data as
<typeStatus id="882C8843FFDDB918BDE1E4E0A950FF29" box="[1220,1322,151,175]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
;
<emphasis id="65E3EAF3FFDDB918BC11E4EFA9EEFF36" bold="true" box="[1332,1428,151,176]" pageId="9" pageNumber="10">Lao Cai</emphasis>
: 2 3, 3 Ƥ, Sa Pa (City, NP office),
<geoCoordinate id="32A35026FFDDB918BB28E4CBAE10FF52" box="[525,618,188,212]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="22.333334">22°20'N</geoCoordinate>
<geoCoordinate id="32A35026FFDDB918BB54E4CBAEA0FF52" box="[625,730,188,212]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="103.833336">103°50'E</geoCoordinate>
, alt.
<quantity id="906F9B04FFDDB918BA2AE4CBAF1FFF52" box="[783,869,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="m" value="1600.0">1600 m</quantity>
, Nest# VN-P-
<date id="23291021FFDDB918BD22E4CBA82CFF52" box="[1031,1110,188,212]" pageId="9" pageNumber="10" value="2006-08-04">2006-4</date>
,
<date id="23291021FFDDB918BD45E4CBA88FFF52" box="[1120,1269,188,212]" pageId="9" pageNumber="10" value="2006-08-21">21.VIII.2006</date>
, LN, FS &amp; JK; 5 Ƥ, Xin Chai, Sa Pa, alt.
<quantity id="906F9B04FFDDB918B896E4A8AE73FF71" box="[435,521,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="9" pageNumber="10" unit="m" value="1350.0">1350 m</quantity>
,
<date id="23291021FFDDB918BB31E4A8AEE8FF71" box="[532,658,223,247]" pageId="9" pageNumber="10" value="2009-07-06">6.VII.2009</date>
, HP; 4 Ƥ, Cat Cat, Sa Pa, alt.
<quantity id="906F9B04FFDDB918BAFCE4A8A84AFF71" box="[985,1072,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="9" pageNumber="10" unit="m" value="1450.0">1450 m</quantity>
,
<date id="23291021FFDDB918BD1EE4A8A8C2FF71" box="[1083,1208,223,247]" pageId="9" pageNumber="10" value="2009-07-07">7.VII.2009</date>
, HP; 2 3, Ta Pin, Sa Pa, alt.
<quantity id="906F9B04FFDDB918B9C9E573AD39FE9A" box="[236,323,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="m" value="1600.0">1600 m</quantity>
,
<date id="23291021FFDDB918B875E573AD9FFE9A" box="[336,485,260,284]" pageId="9" pageNumber="10" value="2006-08-20">20.VIII.2006</date>
, LN, FS &amp; JK; 1 Ƥ, Ban Khoang, Sa Pa, alt.
<quantity id="906F9B04FFDDB918BAC7E573A840FE9A" box="[994,1082,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="m" value="1800.0">1800 m</quantity>
,
<date id="23291021FFDDB918BD63E573A8A6FE9A" box="[1094,1244,260,284]" pageId="9" pageNumber="10" value="2006-08-20">20.VIII.2006</date>
, LN, FS &amp; JK; 1 3, 3 Ƥ, Sa Pa NP (Silver Waterfall),
<geoCoordinate id="32A35026FFDDB918BB0AE550AEDAFEB9" box="[559,672,295,319]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="22.358334">22°21.5'N</geoCoordinate>
<geoCoordinate id="32A35026FFDDB918BB82E550AF5CFEB9" box="[679,806,295,319]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="92" value="103.775">103°46.5'E</geoCoordinate>
, alt.
<quantity id="906F9B04FFDDB918BA7EE550AFC8FEB9" box="[859,946,295,319]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="m" value="2000.0">2000 m</quantity>
,
<date id="23291021FFDDB918BA98E550A828FEB9" box="[957,1106,295,319]" pageId="9" pageNumber="10" value="2006-08-21">21.VIII.2006</date>
, LN, FS &amp; JK; 3 Ƥ, Xeomity, Sa Pa, alt.
<quantity id="906F9B04FFDDB918B834E53BAD13FEE2" box="[273,361,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="10" unit="m" value="1700.0">1700 m</quantity>
,
<date id="23291021FFDDB918B853E53BAD89FEE2" box="[374,499,332,356]" pageId="9" pageNumber="10" value="2009-07-09">9.VII.2009</date>
, HP; 1 3, Sa Pa town, alt.
<quantity id="906F9B04FFDDB918BA02E53BAF05FEE2" box="[807,895,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="m" value="1600.0">1600 m</quantity>
,
<date id="23291021FFDDB918BAA9E53BA870FEE2" box="[908,1034,332,356]" pageId="9" pageNumber="10" value="2009-07-08">8.VII.2009</date>
, HP, 1 3, Sa Pa, alt.
<quantity id="906F9B04FFDDB918BDDCE53BA9EDFEE2" box="[1273,1431,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="9" pageNumber="10" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">15002000 m</quantity>
,
<date id="23291021FFDDB918B9B2E518AD41FE01" box="[151,315,367,391]" pageId="9" pageNumber="10" value="2003-05-17" valueMax="2003-05-18" valueMin="2003-05-17">17-18.V. 2003</date>
, ISD-c; 6 Ƥ, Sa Pa, alt.
<quantity id="906F9B04FFDDB918BB76E518AED6FE01" box="[595,684,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="m" value="1600.0">1600 m</quantity>
, Nest# Sapa-P-01,
<date id="23291021FFDDB918BAAFE518A87DFE01" box="[906,1031,367,391]" pageId="9" pageNumber="10" value="2009-07-05">5.VII.2009</date>
, HP; 4 Ƥ, Ban Pho, Bac Ha, Nest# 2008-LC-P-02,
<date id="23291021FFDDB918B86EE5E3ADB6FE2A" box="[331,460,404,428]" pageId="9" pageNumber="10" value="2008-06-25">25.VI.2008</date>
, LN &amp; HP; 2 Ƥ, Nam Lung, Bac Ha, Nest# 2008-LC-P-03,
<date id="23291021FFDDB918BDB5E5E3A968FE2A" box="[1168,1298,404,428]" pageId="9" pageNumber="10" value="2008-06-26">26.VI.2008</date>
, LN &amp; HP;
<emphasis id="65E3EAF3FFDDB918B9B2E5C0AD76FE56" bold="true" box="[151,268,439,464]" pageId="9" pageNumber="10">Dien Bien</emphasis>
: 2 Ƥ, Ban Bua, Muong Fang, alt.
<quantity id="906F9B04FFDDB918BBAFE5C0AE9BFE49" box="[650,737,439,463]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="1000.0">1000 m</quantity>
,
<date id="23291021FFDDB918BBC8E5C0AF14FE49" box="[749,878,439,463]" pageId="9" pageNumber="10" value="2001-06-14">14.VI.2001</date>
, ISD-c; 1 3, Tuan Giao, alt.
<quantity id="906F9B04FFDDB918BD96E5C0A973FE49" box="[1203,1289,439,463]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="9" pageNumber="10" unit="m" value="1168.0">1168 m</quantity>
,
<date id="23291021FFDDB918BC30E5C0A9E2FE49" box="[1301,1432,439,463]" pageId="9" pageNumber="10" value="2009-06-25">25.VI.2009</date>
, LN, HP &amp; JK; 1 3, Co Ma, Thuan Chau, Son La, alt.
<quantity id="906F9B04FFDDB918BBD8E5ABAF2FFE72" box="[765,853,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="10" unit="m" value="1400.0">1400 m</quantity>
,
<date id="23291021FFDDB918BA47E5ABAFB3FE72" box="[866,969,476,500]" pageId="9" pageNumber="10" value="2009-05-04">4.V.2009</date>
, HP;
<emphasis id="65E3EAF3FFDDB918BD2CE5ABA807FE72" bold="true" box="[1033,1149,475,500]" pageId="9" pageNumber="10">Hoa Binh</emphasis>
: 13 Ƥ, Pa Co, Mai Chau,
<geoCoordinate id="32A35026FFDDB918B9B2E588AD72FD91" box="[151,264,511,535]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="20.741667">20°44.5'N</geoCoordinate>
<geoCoordinate id="32A35026FFDDB918B82AE588ADF4FD91" box="[271,398,511,535]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="92" value="104.89167">104°53.5'E</geoCoordinate>
, alt.
<quantity id="906F9B04FFDDB918B8E7E588AE63FD91" box="[450,537,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="9" pageNumber="10" unit="m" value="1450.0">1450 m</quantity>
, Nest # VN-P-
<date id="23291021FFDDB918BBE7E588AF67FD91" box="[706,797,511,535]" pageId="9" pageNumber="10" value="2006" valueMax="2017">2006-17</date>
,
<date id="23291021FFDDB918BA0DE588AFC7FD91" box="[808,957,511,535]" pageId="9" pageNumber="10" value="2006-08-27">27.VIII.2006</date>
, LN, FS &amp; JK; 2 3, 6 Ƥ, Pa Co, Mai Chau, alt.
<quantity id="906F9B04FFDDB918B9E4E653AD6DFDBA" box="[193,279,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="1000.0">1000 m</quantity>
,
<date id="23291021FFDDB918B806E653ADC2FDBA" box="[291,440,548,572]" pageId="9" pageNumber="10" value="2006-08-23">23.VIII.2006</date>
, LN &amp; JK; Pa Co, Mai Chau [1 Ƥ, alt.
<quantity id="906F9B04FFDDB918BA4CE653AF82FDBA" box="[873,1016,548,572]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="9" pageNumber="10" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">9001000 m</quantity>
,
<date id="23291021FFDDB918BD21E653A8FFFDBA" box="[1028,1157,548,572]" pageId="9" pageNumber="10" value="2001-06-27">27.VI.2001</date>
, ISD-c; 1 Ƥ, alt.
<quantity id="906F9B04FFDDB918BC64E653A9EDFDBA" box="[1345,1431,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="10" unit="m" value="1100.0">1100 m</quantity>
,
<date id="23291021FFDDB918B9B2E630AD76FDD9" box="[151,268,583,607]" pageId="9" pageNumber="10" value="2002-05-20">20.V.2002</date>
, V. T. Hoang; 5 Ƥ, alt.
<quantity id="906F9B04FFDDB918BB35E630AE1DFDD9" box="[528,615,583,607]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="10" unit="m" value="1400.0">1400 m</quantity>
, Nest# 2008-HB-P-03,
<date id="23291021FFDDB918BA40E63FAF9AFDD9" box="[869,992,584,607]" pageId="9" pageNumber="10" value="2008-06-11">11.VI.2008</date>
, LN &amp; HP; 1 3, 2 Ƥ, Pa Co, Mai Chau, alt.
<quantity id="906F9B04FFDDB918B9E4E61BAD6DFD02" box="[193,279,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="1000.0">1000 m</quantity>
,
<date id="23291021FFDDB918B806E61BADC3FD02" box="[291,441,620,644]" pageId="9" pageNumber="10" value="2005-08-23">23.VIII.2005</date>
, LN &amp; J K.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFDDB918B9E2E6F8ACAAFC07" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="572836E1FFDDB918B9E2E6F8ACAAFC07" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
<emphasis id="65E3EAF3FFDDB918B9E2E6F8AD3BFD2E" bold="true" box="[199,321,655,680]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
This species can be distinguished from all other known species of subgenus
<emphasis id="65E3EAF3FFDDB918BDFEE6E7A93DFD2E" box="[1243,1351,656,680]" italics="true" pageId="9" pageNumber="10">Polistella</emphasis>
by the following combination of characters: female clypeus convex, produced ventrally into blunt angle, in lateral view prominently swollen anteriorly in dorsal one-third; male clypeus bent backward apically; metasomal tergum 1 short and thick, about 0.9 × as long as its apical width, in lateral view abruptly swollen dorsally at base, with corner between anterior and dorsal faces bluntly angled; metasomal sternum
<quantity id="906F9B04FFDDB918BA87E768AFB7FCB1" box="[930,973,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="10" unit="in" value="2.0">2 in</quantity>
lateral view strongly swollen ventrally in anterior half to two thirds, then ventral margin bluntly angled into straight line parallel to ventral margin of the tergum.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFDDB91CB9E2E7FCAE43FEB9" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10" type="description">
<paragraph id="572836E1FFDDB918B9E2E7FCADCEFC41" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
<emphasis id="65E3EAF3FFDDB918B9E2E7FCAD23FC22" bold="true" box="[199,345,907,932]" pageId="9" pageNumber="10">Description.</emphasis>
FEMALE. Body length
<quantity id="906F9B04FFDDB918BB5FE7FBAF64FC22" box="[634,798,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.15" metricValueMin="1.05" pageId="9" pageNumber="10" unit="mm" value="11.0" valueMax="11.5" valueMin="10.5">10.511.5 mm</quantity>
(
<typeStatus id="882C8843FFDDB918BA15E7FBAFE2FC22" box="[816,920,908,932]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFDDB918BAC8E7FBA838FC22" box="[1005,1090,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="10" unit="mm" value="11.0">11 mm</quantity>
); fore wing length
<quantity id="906F9B04FFDDB918BC04E7FBA9E6FC22" box="[1313,1436,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="9" pageNumber="10" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
(
<typeStatus id="882C8843FFDDB918B9BAE7D8AD7DFC41" box="[159,263,943,967]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
: about
<quantity id="906F9B04FFDDB918B873E7D8ADD2FC41" box="[342,424,943,967]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="10" unit="mm" value="11.0">11 mm</quantity>
).
</paragraph>
<paragraph id="572836E1FFDDB918B9E2E7A3AE6BFA1A" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
Head in frontal view about 1.2 × as wide as high (
<figureCitation id="CFAC2A64FFDDB918BBD2E7A3AF36FC6A" box="[759,844,980,1004]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 27</figureCitation>
); in dorsal view weakly swollen laterally just behind eyes, then narrowed posteriorly, with posterior margin shallowly and broadly emarginate (
<figureCitation id="CFAC2A64FFDDB918BDAAE780A899FB89" box="[1167,1251,1015,1039]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 28</figureCitation>
). Vertex slightly raised in area among ocelli, slightly slope down behind posterior ocelli towards occpital carina; POD:OOD = about 1:1.9; POD about 1.5 × Od. Gena in lateral view about 0.8 × as wide as eye (
<figureCitation id="CFAC2A64FFDDB918BAD7E037A83CFBDE" box="[1010,1094,1088,1112]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 29</figureCitation>
); occipital carina fine, evanescent in ventral one third of gena. Inner eye margins weakly convergent ventrally, in frontal view about 1.2 × further apart from each other at vertex than at clypeus (
<figureCitation id="CFAC2A64FFDDB918BB91E0FFAF73FB26" box="[692,777,1160,1184]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 27</figureCitation>
). Antennal sockets closer to inner eye margin than to each other; anterior tentorial pit slightly further from antennal socket than from inner eye margin; interantennal space weakly raised. Clypeus in frontal view as wide as high, produced ventrally into blunt angle; in lateral view convex, prominently swollen anteriorly in upper one third (
<figureCitation id="CFAC2A64FFDDB918BBF4E083AF5FFA8A" box="[721,805,1268,1292]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 29</figureCitation>
); lateral margin of clypeus lying along inner eye margin about as long as diameter of antennal socket and shorter than length of malar space. Antenna (
<figureCitation id="CFAC2A64FFDDB918BD90E16FA973FAB6" box="[1205,1289,1304,1328]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 30</figureCitation>
): scape more than 3 × as long as its maximum width; flagellomere 1 slightly less than 3 × as long as its maximum width, longer than length of flagellomeres 2 and 3 combined; flagellomere 2 as long as wide; terminal flagellomere bulletshaped, as long as its basal width.
</paragraph>
<paragraph id="572836E1FFDDB918B9E2E1DFAF1BF91E" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
Pronotal carina sharply raised, produced dorsally into thin lamella in dorsal part, slightly sinuate backward on lateral sides, reaching ventral corner of pronotum. Mesoscutum weakly convex (
<figureCitation id="CFAC2A64FFDDB918BD11E1BBA8F3FA62" box="[1076,1161,1484,1508]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 31</figureCitation>
), about as long as wide between tegulae; anterior margin broadly rounded. Scutellum strongly convex except nearly flat median part. Metanotum weakly convex, strongly depressed along anterior margin. Propodeum short; posterior face widely (about half the maximum width of propodeum) and shallowly excavated medially, more or less smoothly passing into lateral faces; propodeal orifice elongate, about 1.8 × as long as wide (measured at widest part), somewhat narrowed in dorsal half. Jugal lobe of hind wing rounded (
<figureCitation id="CFAC2A64FFDDB918BA25E2F7AF2EF91E" box="[768,852,1664,1688]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 32</figureCitation>
).
</paragraph>
<paragraph id="572836E1FFDDB918B9E2E2D3A81AF8AF" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
Metasomal tergum 1 short and thick, about 0.9 × as long as its apical width, in lateral view abruptly swollen dorsally just behind basal slit for reception of propodeal suspensory ligament; corner between anterior and dorsal faces bluntly angled (
<figureCitation id="CFAC2A64FFDDB918B8ABE29BAD9FF882" box="[398,485,1772,1796]" captionStart="FIGURES 27 33" captionStartId="10.[151,269,1754,1778]" captionTargetBox="[201,1396,411,1729]" captionTargetId="figure@10.[191,1396,411,1729]" captionTargetPageId="10" captionText="FIGURES 27 33. Polistes (Polistella) reliciniclypeus sp. nov., Ƥ. 27 29. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="9" pageNumber="10">Fig. 33</figureCitation>
). Sternum
<quantity id="906F9B04FFDDB918BB46E29BAEEAF882" box="[611,656,1772,1796]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="10" unit="in" value="2.0">2 in</quantity>
lateral view strongly swollen ventrally in anterior half, then ventral margin bluntly angled into nearly straight line parallel to ventral margin of the tergum.
</paragraph>
<paragraph id="572836E1FFDDB91BB9E2E343AF5AFEE2" blockId="9.[151,1437,151,2012]" lastBlockId="10.[151,1436,151,357]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">Clypeus with scattered large punctures, each bearing sharply pointed golden bristle. Mandible with several deep punctures. Frons densely covered with coarse punctures. Vertex and gena with sparse punctures less coarse than those on frons; punctures around ocelli sparse strong; ventral one third of gena with coarse punctures. Pronotum with dense coarse flat-bottomed punctures; spaces between punctures very narrow, slightly raised to form reticulation. Mesocutum densely with coarse flat-bottomed punctures; punctures on scutellum and metanotum dense coarse but smaller than those on mesoscutum. Mesepisternum with dense coarse well-defined punctures in posterodorsal part (punctures in dorsal margin similar to those on pronotum), barely punctured in anteroventral part; border between posterodorsal and anteroventral parts distinct. Dorsal metapleuron with striae; ventral metapleuron with sparse strong punctures. Propodeum with strong transverse striae; lateral face with sparse ill-defined punctures. Metasomal segements densely covered with minute punctures in addition to scattered small punctures (ill-defined on terga; more or less well-defined on sterna).</paragraph>
<caption id="03E86669FFDEB91BB9B2E2ADAD97F8B0" httpUri="https://zenodo.org/record/201095/files/figure.png" pageId="10" pageNumber="11" targetBox="[201,1396,411,1729]" targetPageId="10">
<paragraph id="572836E1FFDEB91BB9B2E2ADAD97F8B0" blockId="10.[151,1436,1754,1846]" pageId="10" pageNumber="11">
<emphasis id="65E3EAF3FFDEB91BB9B2E2ADAD25F974" bold="true" box="[151,351,1754,1778]" pageId="10" pageNumber="11">FIGURES 2733.</emphasis>
<taxonomicName id="90974D62FFDEB91BB843E2ADAEA4F977" box="[358,734,1754,1777]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="reliciniclypeus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFDEB91BB843E2ADADC0F977" box="[358,442,1754,1777]" italics="true" pageId="10" pageNumber="11">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFDEB91BB8EEE2ADAE4BF977" box="[459,561,1754,1777]" italics="true" pageId="10" pageNumber="11">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFDEB91BBB67E2ADAEA4F977" box="[578,734,1754,1777]" italics="true" pageId="10" pageNumber="11">reliciniclypeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFDEB91BBBC3E2ADAF47F974" box="[742,829,1754,1778]" pageId="10" pageNumber="11" rank="species">
<emphasis id="65E3EAF3FFDEB91BBBC3E2ADAF47F974" bold="true" box="[742,829,1754,1778]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
, Ƥ. 2729. Head. 27. Frontal view. 28. Dorsal view. 29. Lateral view. 30. Left antenna. 31. Jugal lobe of left hind wing. 32. Mesosoma, lateral view. 33. Metasomal segments 1 and 2, lateral view. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="572836E1FFDEB91AB9E2E311AE89FE2A" blockId="10.[151,1437,1894,2026]" lastBlockId="11.[151,1436,151,536]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="65E3EAF3FFDEB91BB9E2E311AD77F8F8" box="[199,269,1894,1918]" italics="true" pageId="10" pageNumber="11">Color.</emphasis>
Black; following parts yellow to orange-yellow: clypeus except black margins, narrow band along inner eye margin extending from bottom of frons to middle of eye emargination, narrow short band along eye margin in dorsal part of gena, narrow band along pronotal carina with branches extending along posterodorsal margin of pronotum, anterior transverse band on scutellum (often most of scutellar disc, sometimes reduced to paired anterolateral spots), paired anterolateral spots on metanotum (sometimes united into narrow to wide anterior band), spot on propodeal valvula, narrow subapical band on tergum 1 (often narrowly interrupted medially), wide apical band with irregular anterior margin on tergum 6; following parts reddish brown: mandible except black basal margin and teeth, antennal scape beneath, pedicel, basal part of flagellomere 1, apical flagellomeres beneath, pronotum ventrolaterally, tegulae, ill-defined wide bands on tergum 1 (in front of yellow band), narrow apical bands on terga 2, 3 and 5 and sterna 25, entire visible part of metasomal segment 6. Legs black, dorsal surface of femora and ventral surface of tibiae largely reddish brown. Wings yellowish-brown, weakly infuscate, slightly more strongly infuscate along anterior margin of fore wing; veins dark brown.
</paragraph>
<paragraph id="572836E1FFDFB91AB9E2E5CFAFA8FE49" blockId="11.[151,1436,151,536]" box="[199,978,439,464]" pageId="11" pageNumber="12">
MALE. Body length 9.0 mm
<quantity id="906F9B04FFDFB91ABB3CE5C0AE0BFE49" box="[537,625,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="12" unit="mm" value="9.5">9.5 mm</quantity>
; forewing length
<quantity id="906F9B04FFDFB91ABA10E5C0AFB4FE49" box="[821,974,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.05" metricValueMin="0.95" pageId="11" pageNumber="12" unit="mm" value="10.0" valueMax="10.5" valueMin="9.5">9.510.5 mm</quantity>
.
</paragraph>
<paragraph id="572836E1FFDFB91DB9E2E5ABAD1BFE2A" blockId="11.[151,1436,151,536]" lastBlockId="12.[151,1437,151,1509]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Structure as in female, but differing from the latter as follows: head proportionally smaller, transverse, about 1.4 × as wide as high in frontal view (
<figureCitation id="CFAC2A64FFDFB91ABBB4E588AE92FD91" box="[657,744,511,535]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="11" pageNumber="12">Fig. 34</figureCitation>
); eye strongly swollen laterally; inner eye margins strongly convergent, about 1.5 × further apart from each other at vertex than at clypeus; gena narrow, in lateral view about 0.4 × as wide as eye (
<figureCitation id="CFAC2A64FFD8B91DB8B1E4CBAD93FF52" box="[404,489,188,212]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 35</figureCitation>
), with weakly raised blunt ridge running along posterior margin of eye; clypeus in frontal view about 1.1 × as wide as high (
<figureCitation id="CFAC2A64FFD8B91DBBAAE4A8AE9DFF71" box="[655,743,223,247]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 34</figureCitation>
), only slightly produced ventrally, evenly and very weakly rounded apically, in lateral view weakly convex dorsally, bent backward ventrally (
<figureCitation id="CFAC2A64FFD8B91DBD16E573A8FCFE9A" box="[1075,1158,260,284]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 35</figureCitation>
). Antenna slenderer than in female (
<figureCitation id="CFAC2A64FFD8B91DB831E550AD11FEB9" box="[276,363,295,319]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 36</figureCitation>
); scape short, about 2.8 × as long as its maximum width; flagellomere 1 as long as length of flagellomeres 2 and 3 combined; flagellomeres 2 and 3 each longer than wide; terminal flagellomere elongate, slightly curved, about 2 × as long as its basal width. Metasomal sternum 7 depressed medially (
<figureCitation id="CFAC2A64FFD8B91DBDF9E518A94BFE01" box="[1244,1329,367,391]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 37</figureCitation>
), without tubercle (
<figureCitation id="CFAC2A64FFD8B91DB825E5E3AD2EFE2A" box="[256,340,404,428]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 38</figureCitation>
).
</paragraph>
<caption id="03E86669FFDFB91AB9B2E320A857F833" httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="11" pageNumber="12" targetBox="[191,1396,591,1864]" targetPageId="11">
<paragraph id="572836E1FFDFB91AB9B2E320A857F833" blockId="11.[151,1436,1879,1973]" pageId="11" pageNumber="12">
<emphasis id="65E3EAF3FFDFB91AB9B2E320AD24F8E9" bold="true" box="[151,350,1879,1903]" pageId="11" pageNumber="12">FIGURES 3441.</emphasis>
<taxonomicName id="90974D62FFDFB91AB840E32FAEA7F8E9" box="[357,733,1880,1903]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="reliciniclypeus" status="sp. nov." subGenus="Polistella">
<emphasis id="65E3EAF3FFDFB91AB840E32FADC0F8E9" box="[357,442,1880,1903]" italics="true" pageId="11" pageNumber="12">Polistes</emphasis>
(
<emphasis id="65E3EAF3FFDFB91AB8EFE32FAE4AF8E9" box="[458,560,1880,1903]" italics="true" pageId="11" pageNumber="12">Polistella</emphasis>
)
<emphasis id="65E3EAF3FFDFB91ABB64E32FAEA7F8E9" box="[577,733,1880,1903]" italics="true" pageId="11" pageNumber="12">reliciniclypeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFDFB91ABBC1E320AF40F8E9" box="[740,826,1879,1903]" pageId="11" pageNumber="12" rank="species">
<emphasis id="65E3EAF3FFDFB91ABBC1E320AF40F8E9" bold="true" box="[740,826,1879,1903]" pageId="11" pageNumber="12">sp. nov.</emphasis>
</taxonomicNameLabel>
, 3. 3435. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 3738. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 4041. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1mm.
</paragraph>
</caption>
<paragraph id="572836E1FFD8B91DB9E2E5C0ACAFFE72" blockId="12.[151,1437,151,1509]" pageId="12" pageNumber="13">Body surface sculpture as in female, but clypeus without large punctures and densely covered with long golden hairs.</paragraph>
<paragraph id="572836E1FFD8B91DB9E2E677ADF8FD69" blockId="12.[151,1437,151,1509]" pageId="12" pageNumber="13">
<emphasis id="65E3EAF3FFD8B91DB9E2E677AD77FD9E" box="[199,269,512,536]" italics="true" pageId="12" pageNumber="13">Color.</emphasis>
Similar to female, but reddish-brown markings on head and mesosoma reduced, while more extensively marked with yellow as follows: clypeus entirely, mandible except for small basal black spot, ventral half of frons except for ill-defined black band connecting anterior tentorial pit and antennal socket and interantennal black spot, narrow long band on gena along posterodorsal margin of eye, antennal scape beneath, disc of scutellum (sometimes reduced into wide anterior band), paired anterolateral spots (sometimes narrow transverse band) on metanotum, spot on propodeal valvula, ventral face of fore coxa and spot on mid-coxa; antennal flagellum reddish-brown to dark yellow beneath.
</paragraph>
<paragraph id="572836E1FFD8B91DB9E2E68BAD51FBB2" blockId="12.[151,1437,151,1509]" pageId="12" pageNumber="13">
<emphasis id="65E3EAF3FFD8B91DB9E2E68BAD42FC92" box="[199,312,764,788]" italics="true" pageId="12" pageNumber="13">Genitalia.</emphasis>
Generally similar to that of
<taxonomicName id="90974D62FFD8B91DBB54E68BAF52FC92" box="[625,808,764,788]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="65E3EAF3FFD8B91DBB54E68BAF52FC92" box="[625,808,764,788]" italics="true" pageId="12" pageNumber="13">Polistes dawnae</emphasis>
</taxonomicName>
. Lamina volsellaris desclerotized and fused with cuspis. Digitus in inner aspect of paramere (
<figureCitation id="CFAC2A64FFD8B91DBB1EE768AEF7FCB1" box="[571,653,799,823]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 39</figureCitation>
) slightly more than 3.5 × as long as wide (measured at widest part), distinctly swollen near base, gradually narrowed apically to mid-length, then slightly swollen towards the rounded apex. Aedeagus (
<figureCitation id="CFAC2A64FFD8B91DB873E710ADA2FCF9" box="[342,472,871,895]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Figs 40, 41</figureCitation>
): penis valves slightly longer than basal apodeme, in ventral view narrowest near midlength, distally club-shaped, weakly swollen proximally from mid-length, strongly swollen and produced proximolaterally in about proximal one fourth and distinctly produced laterally near proximal margins, in lateral view slightly thickened in proximal one fourth and with dorsal margin weakly and smoothly sinuate, with proximoventral corner produced into abtuse angle (
<figureCitation id="CFAC2A64FFD8B91DBBF8E780AF4BFB89" box="[733,817,1015,1039]" captionStart="FIGURES 34 41" captionStartId="11.[151,269,1879,1903]" captionTargetBox="[191,1396,591,1864]" captionTargetId="figure@11.[191,1396,591,1873]" captionTargetPageId="11" captionText="FIGURES 34 41. Polistes (Polistella) reliciniclypeus sp. nov., 3. 34 35. Head. 34. Frontal view. 35. Lateral view. 36. Left antenna. 37 38. Terminal sterna. 37. Ventral view. 38. Apical part, lateral view. 39. Inner aspect of paramere with digitus and volsella. 40 41. Aedeagus. 40. Ventral view. 41. Lateral view. Scale 1 mm." httpUri="https://zenodo.org/record/201096/files/figure.png" pageId="12" pageNumber="13">Fig. 41</figureCitation>
); ventral margins of penis valves finely serrated along entire length.
</paragraph>
<paragraph id="572836E1FFD8B91DB9E2E037ACC5FB6E" blockId="12.[151,1437,151,1509]" pageId="12" pageNumber="13">
<emphasis id="65E3EAF3FFD8B91DB9E2E037AD79FBDE" bold="true" box="[199,259,1088,1112]" pageId="12" pageNumber="13">Nest.</emphasis>
The following nine nests were examined: Nest# VN-P-2006-4, Sa Pa, Lao Cai, alt.
<quantity id="906F9B04FFD8B91DBC13E037A9E2FBDE" box="[1334,1432,1088,1112]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="13" unit="m" value="1600.0">1600 m</quantity>
,
<date id="23291021FFD8B91DB9B2E013AD56FBFA" box="[151,300,1124,1148]" pageId="12" pageNumber="13" value="2006-08-21">21.VIII.2006</date>
, LN, FS &amp; JK; Nest# 2009-Sapa-P-01, Sa Pa, Lao Cai, alt.
<quantity id="906F9B04FFD8B91DBAC5E013A843FBFA" box="[992,1081,1124,1148]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="13" unit="m" value="1600.0">1600 m</quantity>
,
<date id="23291021FFD8B91DBD62E013A8BEFBFA" box="[1095,1220,1124,1148]" pageId="12" pageNumber="13" value="2009-07-05">5.VII.2009</date>
, HP; Nest# VN-P-
<date id="23291021FFD8B91DB9B2E0FFAC8EFB26" box="[151,244,1160,1184]" pageId="12" pageNumber="13" value="2006" valueMax="2010">2006-10</date>
to -13, Phong Tho, Lai Chau, alt.
<quantity id="906F9B04FFD8B91DBBA5E0FFAEA2FB26" box="[640,728,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="13" unit="m" value="1600.0">1600 m</quantity>
,
<date id="23291021FFD8B91DBBC0E0FFAF01FB26" box="[741,891,1160,1184]" pageId="12" pageNumber="13" value="2006-08-22">22.VIII.2006</date>
, LN, FS &amp; JK; Nest# 2008-LC-P-02 &amp; 0 3, Bac Ha, Lao Cai, 25 &amp;
<date id="23291021FFD8B91DB850E0DBAD8DFB42" box="[373,503,1196,1220]" pageId="12" pageNumber="13" value="2008-06-26">26.VI.2008</date>
, LN &amp; HP; Nest# 2008-HB-P-03, Mai Chau, Hoa Binh, alt.
<quantity id="906F9B04FFD8B91DBD95E0DBA973FB42" box="[1200,1289,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="13" unit="m" value="1400.0">1400 m</quantity>
,
<date id="23291021FFD8B91DBC33E0DBA9EDFB42" box="[1302,1431,1196,1220]" pageId="12" pageNumber="13" value="2008-06-11">11.VI.2008</date>
, HP.
</paragraph>
<paragraph id="572836E1FFD8B91DB9E2E082A82FFAFE" blockId="12.[151,1437,151,1509]" pageId="12" pageNumber="13">
Nest VN-P-2006-4 was found under the leaf of a palm about
<quantity id="906F9B04FFD8B91DBA53E083AFCCFA8A" box="[886,950,1268,1292]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="m" value="0.5">0.5 m</quantity>
above the ground. Five nests (VN-P-
<date id="23291021FFD8B91DBC7EE083ACC9FAB6" pageId="12" pageNumber="13" value="2006" valueMax="2010">2006- 10</date>
to 13 and 2009-Sapa-P-01) were found on the cliffs along the Hoang Lien Pass road, and they all were directly attached to rocks
<quantity id="906F9B04FFD8B91DB87EE14BADCDFAD2" box="[347,439,1340,1364]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="12" pageNumber="13" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1 to 2 m</quantity>
above the road. The two nests collected in Bac Ha and one in Mai Chau were at the early nest-founding stage and had a comb comprised of only empty shallow cells (
<tableCitation id="1A15035AFFD8B91DBAD4E117A83EFAFE" box="[1009,1092,1376,1400]" captionStart="TABLE 1" captionStartId="5.[151,235,157,178]" captionTargetBox="[165,1417,218,1135]" captionTargetPageId="5" captionText="TABLE 1. Biological data of examined nests of Polistes species from subgenus Polistella." httpUri="http://table.plazi.org/id/03E86669FFD1B914B9B2E4EAA857FF35" pageId="12" pageNumber="13" tableUuid="03E86669FFD1B914B9B2E4EAA857FF35">Table 1</tableCitation>
).
</paragraph>
<paragraph id="572836E1FFD8B91DB9E2E1F2A9F6F9E6" blockId="12.[151,1437,151,1509]" lastBlockId="12.[203,1420,1612,1734]" pageId="12" pageNumber="13">
Nest structural characters are as follows (for measurements see
<tableCitation id="1A15035AFFD8B91DBAAFE1F3AFA7FA1A" box="[906,989,1412,1436]" captionStart="TABLE 3" captionStartId="12.[151,231,1558,1579]" captionTargetBox="[203,1420,1755,1981]" captionTargetPageId="12" captionText="TABLE 3. Measuments of nests of Polistes reliciniclypeus sp. nov." httpUri="http://table.plazi.org/id/03E86669FFD8B91DB9B2E261AF66F9AA" pageId="12" pageNumber="13" tableUuid="03E86669FFD8B91DB9B2E261AF66F9AA">Table 3</tableCitation>
). Petiole single, terminal, with thin central core of plant fibers, enlarged strictly with adult salivary secretion, black and lustrous; salivary coating extended onto substrate and largely onto back of comb. Comb tough, pliable in texture, dark brownish gray, expanded from Nest coder Petiolate Petiole Cell width at open end Cell width at bottom Cell depth (mm) (N=10)
</paragraph>
<caption id="03E86669FFD8B91DB9B2E261AF66F9AA" ID-Table-UUID="03E86669FFD8B91DB9B2E261AF66F9AA" box="[151,796,1558,1580]" httpUri="http://table.plazi.org/id/03E86669FFD8B91DB9B2E261AF66F9AA" pageId="12" pageNumber="13" targetBox="[203,1420,1755,1981]" targetIsTable="true" targetPageId="12">
<paragraph id="572836E1FFD8B91DB9B2E261AF66F9AA" blockId="12.[151,796,1558,1580]" box="[151,796,1558,1580]" pageId="12" pageNumber="13">
<emphasis id="65E3EAF3FFD8B91DB9B2E261AC84F9AD" bold="true" box="[151,254,1558,1579]" pageId="12" pageNumber="13">TABLE 3.</emphasis>
Measuments of nests of
<taxonomicName id="90974D62FFD8B91DB8C8E260AEB1F9AA" box="[493,715,1559,1580]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="reliciniclypeus" status="sp. nov.">
<emphasis id="65E3EAF3FFD8B91DB8C8E260AEB1F9AA" box="[493,715,1559,1580]" italics="true" pageId="12" pageNumber="13">Polistes reliciniclypeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ED05788FFD8B91DBBF4E260AF66F9AA" box="[721,796,1559,1580]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="572836E1FFD8B91DB881E21DA839F9F8" blockId="12.[203,1420,1612,1734]" box="[420,1091,1642,1662]" pageId="12" pageNumber="13">length thickness (mm) (n=10) (mm) (n=10)</paragraph>
<paragraph id="572836E1FFD8B91DB882E2FEAE2CF91B" blockId="12.[203,1420,1612,1734]" box="[423,598,1673,1693]" pageId="12" pageNumber="13">(mm) (mm)</paragraph>
<paragraph id="572836E1FFD8B91CBB85E2C4AE43FEB9" blockId="12.[203,1420,1612,1734]" lastBlockId="13.[151,1436,151,572]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Mean Range Mean Range Mean Range the petiole to become fan-shaped in ventral view; ventral surface convex; dorsal surface deeply concave. Nest cells generally arranged regularly and hexagonal at open end, but partly irregularly arranged, where some cells are pentagonal; outer free margins of peripheral cells rounded; cell expanded towards open end, with width at open end about 1.5 × larger than that at bottom (
<tableCitation id="1A15035AFFD9B91CBB66E573AEE3FE9A" box="[579,665,260,284]" captionStart="TABLE 3" captionStartId="12.[151,231,1558,1579]" captionTargetBox="[203,1420,1755,1981]" captionTargetPageId="12" captionText="TABLE 3. Measuments of nests of Polistes reliciniclypeus sp. nov." httpUri="http://table.plazi.org/id/03E86669FFD8B91DB9B2E261AF66F9AA" pageId="13" pageNumber="14" tableUuid="03E86669FFD8B91DB9B2E261AF66F9AA">Table 3</tableCitation>
); cell wall about
<quantity id="906F9B04FFD9B91CBA7FE573AFC4FE9A" box="[858,958,260,284]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="mm" value="0.03">0.03 mm</quantity>
thick. Cocoon caps pale gray, domed, projecting
<quantity id="906F9B04FFD9B91CB9CFE550AD26FEB9" box="[234,348,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="14" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1 to 2 mm</quantity>
beyond rim of cell.
</paragraph>
</subSubSection>
<paragraph id="572836E1FFD8B91DB9EEE2ACA9F9F83B" pageId="12" pageNumber="13">
<table id="2597C441FFD846EEB9EEE2ACA9F6F83B" box="[203,1420,1755,1981]" gridcols="9" gridrows="6" pageId="12" pageNumber="13">
<tr id="E9A734A3FFD846EEB9EEE2ACA9F6F969" box="[203,1420,1755,1775]" gridrow="0" pageId="12" pageNumber="13">
<th id="AA765DDFFFD846EEB9EEE2ACAD24F969" box="[203,350,1755,1775]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">VN-P-2006-4</th>
<th id="AA765DDFFFD846EEB897E2ACADB4F969" box="[434,462,1755,1775]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">5.1</th>
<th id="AA765DDFFFD846EEBB30E2ACAE1FF969" box="[533,613,1755,1775]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">1.1 x 1.4</th>
<th id="AA765DDFFFD846EEBB8EE2ACAEBDF969" box="[683,711,1755,1775]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">4.0</th>
<th id="AA765DDFFFD846EEBA37E2ACAF2CF969" box="[786,854,1755,1775]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">3.54.5</th>
<th id="AA765DDFFFD846EEBA91E2ACAFAAF969" box="[948,976,1755,1775]" gridcol="5" gridrow="0" pageId="12" pageNumber="13">2.8</th>
<th id="AA765DDFFFD846EEBD06E2ACA81DF969" box="[1059,1127,1755,1775]" gridcol="6" gridrow="0" pageId="12" pageNumber="13">2.53.2</th>
<th id="AA765DDFFFD846EEBDE7E2ACA890F969" box="[1218,1258,1755,1775]" gridcol="7" gridrow="0" pageId="12" pageNumber="13">16.3</th>
<th id="AA765DDFFFD846EEBC0DE2ACA9F6F969" box="[1320,1420,1755,1775]" gridcol="8" gridrow="0" pageId="12" pageNumber="13">15.117.2</th>
</tr>
<tr id="E9A734A3FFD846EEB9EEE373A9F6F89E" box="[203,1420,1796,1816]" gridrow="1" pageId="12" pageNumber="13">
<th id="AA765DDFFFD846EEB9EEE373AD24F89E" box="[203,350,1796,1816]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">VN-P-2006-10</th>
<td id="AA765DDFFFD846EEB897E373ADB4F89E" box="[434,462,1796,1816]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">3.8</td>
<td id="AA765DDFFFD846EEBB30E373AE1FF89E" box="[533,613,1796,1816]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">1.4 x 1.5</td>
<td id="AA765DDFFFD846EEBB8EE373AEBDF89E" box="[683,711,1796,1816]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">4.1</td>
<td id="AA765DDFFFD846EEBA37E373AF2CF89E" box="[786,854,1796,1816]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">3.94.3</td>
<td id="AA765DDFFFD846EEBA91E373AFAAF89E" box="[948,976,1796,1816]" gridcol="5" gridrow="1" pageId="12" pageNumber="13">3.1</td>
<td id="AA765DDFFFD846EEBD06E373A81DF89E" box="[1059,1127,1796,1816]" gridcol="6" gridrow="1" pageId="12" pageNumber="13">2.73.9</td>
<td id="AA765DDFFFD846EEBDE7E373A890F89E" box="[1218,1258,1796,1816]" gridcol="7" gridrow="1" pageId="12" pageNumber="13">16.7</td>
<td id="AA765DDFFFD846EEBC0DE373A9F6F89E" box="[1320,1420,1796,1816]" gridcol="8" gridrow="1" pageId="12" pageNumber="13">15.617.9</td>
</tr>
<tr id="E9A734A3FFD846EEB9EEE35AA9F6F8C7" box="[203,1420,1837,1857]" gridrow="2" pageId="12" pageNumber="13">
<th id="AA765DDFFFD846EEB9EEE35AAD24F8C7" box="[203,350,1837,1857]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">VN-P-2006-11</th>
<td id="AA765DDFFFD846EEB897E35AADB4F8C7" box="[434,462,1837,1857]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">1.9</td>
<td id="AA765DDFFFD846EEBB30E35AAE1FF8C7" box="[533,613,1837,1857]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">1.6 x 2.0</td>
<td id="AA765DDFFFD846EEBB8EE35AAEBDF8C7" box="[683,711,1837,1857]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">4.7</td>
<td id="AA765DDFFFD846EEBA37E35AAF2CF8C7" box="[786,854,1837,1857]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">4.35.2</td>
<td id="AA765DDFFFD846EEBA91E35AAFAAF8C7" box="[948,976,1837,1857]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">3.0</td>
<td id="AA765DDFFFD846EEBD06E35AA81DF8C7" box="[1059,1127,1837,1857]" gridcol="6" gridrow="2" pageId="12" pageNumber="13">2.53.7</td>
<td id="AA765DDFFFD846EEBDE7E35AA890F8C7" box="[1218,1258,1837,1857]" gridcol="7" gridrow="2" pageId="12" pageNumber="13">16.4</td>
<td id="AA765DDFFFD846EEBC0DE35AA9F6F8C7" box="[1320,1420,1837,1857]" gridcol="8" gridrow="2" pageId="12" pageNumber="13">15.317.7</td>
</tr>
<tr id="E9A734A3FFD846EEB9EEE321A9F6F8EC" box="[203,1420,1878,1898]" gridrow="3" pageId="12" pageNumber="13">
<th id="AA765DDFFFD846EEB9EEE321AD24F8EC" box="[203,350,1878,1898]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">VN-P-2006-12</th>
<td id="AA765DDFFFD846EEB897E321ADB4F8EC" box="[434,462,1878,1898]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">-</td>
<td id="AA765DDFFFD846EEBB30E321AE1FF8EC" box="[533,613,1878,1898]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">-</td>
<td id="AA765DDFFFD846EEBB8EE321AEBDF8EC" box="[683,711,1878,1898]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">3.9</td>
<td id="AA765DDFFFD846EEBA37E321AF2CF8EC" box="[786,854,1878,1898]" gridcol="4" gridrow="3" pageId="12" pageNumber="13">3.74.2</td>
<td id="AA765DDFFFD846EEBA91E321AFAAF8EC" box="[948,976,1878,1898]" gridcol="5" gridrow="3" pageId="12" pageNumber="13">2.7</td>
<td id="AA765DDFFFD846EEBD06E321A81DF8EC" box="[1059,1127,1878,1898]" gridcol="6" gridrow="3" pageId="12" pageNumber="13">2.23.4</td>
<td id="AA765DDFFFD846EEBDE7E321A890F8EC" box="[1218,1258,1878,1898]" gridcol="7" gridrow="3" pageId="12" pageNumber="13">17.5</td>
<td id="AA765DDFFFD846EEBC0DE321A9F6F8EC" box="[1320,1420,1878,1898]" gridcol="8" gridrow="3" pageId="12" pageNumber="13">15.919.6</td>
</tr>
<tr id="E9A734A3FFD846EEB9EEE308A9F6F815" box="[203,1420,1919,1939]" gridrow="4" pageId="12" pageNumber="13">
<th id="AA765DDFFFD846EEB9EEE308AD24F815" box="[203,350,1919,1939]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">VN-P-2006-13</th>
<td id="AA765DDFFFD846EEB897E308ADB4F815" box="[434,462,1919,1939]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">4.4</td>
<td id="AA765DDFFFD846EEBB30E308AE1FF815" box="[533,613,1919,1939]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">1.6 x 1.9</td>
<td id="AA765DDFFFD846EEBB8EE308AEBDF815" box="[683,711,1919,1939]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">4.4</td>
<td id="AA765DDFFFD846EEBA37E308AF2CF815" box="[786,854,1919,1939]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">4.05.0</td>
<td id="AA765DDFFFD846EEBA91E308AFAAF815" box="[948,976,1919,1939]" gridcol="5" gridrow="4" pageId="12" pageNumber="13">2.8</td>
<td id="AA765DDFFFD846EEBD06E308A81DF815" box="[1059,1127,1919,1939]" gridcol="6" gridrow="4" pageId="12" pageNumber="13">2.43.4</td>
<td id="AA765DDFFFD846EEBDE7E308A890F815" box="[1218,1258,1919,1939]" gridcol="7" gridrow="4" pageId="12" pageNumber="13">17.6</td>
<td id="AA765DDFFFD846EEBC0DE308A9F6F815" box="[1320,1420,1919,1939]" gridcol="8" gridrow="4" pageId="12" pageNumber="13">16.219.2</td>
</tr>
<tr id="E9A734A3FFD846EEB9EEE3DEA9F6F83B" box="[203,1420,1961,1981]" gridrow="5" pageId="12" pageNumber="13">
<th id="AA765DDFFFD846EEB9EEE3DEAD24F83B" box="[203,350,1961,1981]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">2009-Sapa-P-01</th>
<td id="AA765DDFFFD846EEB897E3DEADB4F83B" box="[434,462,1961,1981]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">5.7</td>
<td id="AA765DDFFFD846EEBB30E3DEAE1FF83B" box="[533,613,1961,1981]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">1.2 x 1.8</td>
<td id="AA765DDFFFD846EEBB8EE3DEAEBDF83B" box="[683,711,1961,1981]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">4.0</td>
<td id="AA765DDFFFD846EEBA37E3DEAF2CF83B" box="[786,854,1961,1981]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">3.64.4</td>
<td id="AA765DDFFFD846EEBA91E3DEAFAAF83B" box="[948,976,1961,1981]" gridcol="5" gridrow="5" pageId="12" pageNumber="13">2.8</td>
<td id="AA765DDFFFD846EEBD06E3DEA81DF83B" box="[1059,1127,1961,1981]" gridcol="6" gridrow="5" pageId="12" pageNumber="13">2.43.3</td>
<td id="AA765DDFFFD846EEBDE7E3DEA890F83B" box="[1218,1258,1961,1981]" gridcol="7" gridrow="5" pageId="12" pageNumber="13">16.6</td>
<td id="AA765DDFFFD846EEBC0DE3DEA9F6F83B" box="[1320,1420,1961,1981]" gridcol="8" gridrow="5" pageId="12" pageNumber="13">16.317.0</td>
</tr>
</table>
</paragraph>
<subSubSection id="1F8D656AFFD9B91CB9E2E53BAEECFE49" pageId="13" pageNumber="14" type="discussion">
<paragraph id="572836E1FFD9B91CB9E2E53BAEECFE49" blockId="13.[151,1436,151,572]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E53BAD4AFEE2" bold="true" box="[199,304,332,356]" pageId="13" pageNumber="14">Remark.</emphasis>
Along the Hoang Lien Pass road this species and
<emphasis id="65E3EAF3FFD9B91CBA41E53AAFBCFEE2" box="[868,966,332,356]" italics="true" pageId="13" pageNumber="14">P. l e p c h a</emphasis>
nested on the cliffs. While the nests of
<taxonomicName id="90974D62FFD9B91CBCA2E53AAC98FE0E" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lepcha">
<emphasis id="65E3EAF3FFD9B91CBCA2E53AAC98FE0E" italics="true" pageId="13" pageNumber="14">P. lepcha</emphasis>
</taxonomicName>
, having pale-green cocoon caps, were rather conspicuous to us, it seemed to us that the nests of this new species are well camouflaged. Two nests collected in Bac Ha and one in Mai Chau suggested that this species commonly founds a nest by multiple females.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9E2E5ABADE9FD91" pageId="13" pageNumber="14" type="etymology">
<paragraph id="572836E1FFD9B91CB9E2E5ABADE9FD91" blockId="13.[151,1436,151,572]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E5ABAD34FE72" bold="true" box="[199,334,476,500]" pageId="13" pageNumber="14">Etymology.</emphasis>
The specific name refers to the male clypeus bent back (
<emphasis id="65E3EAF3FFD9B91CBAE9E5ABA857FE72" box="[972,1069,476,500]" italics="true" pageId="13" pageNumber="14">relicinus</emphasis>
in Latin) ventrally and is treated as a noun in aposition.
</paragraph>
</subSubSection>
<subSubSection id="1F8D656AFFD9B91CB9E2E654AF79FDBA" box="[199,771,547,572]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="572836E1FFD9B91CB9E2E654AF79FDBA" blockId="13.[151,1436,151,572]" box="[199,771,547,572]" pageId="13" pageNumber="14">
<emphasis id="65E3EAF3FFD9B91CB9E2E654AD1AFDBA" bold="true" box="[199,352,547,572]" pageId="13" pageNumber="14">Distribution.</emphasis>
Known only from northern
<collectingCountry id="2F807671FFD9B91CBBB9E653AE85FDBA" box="[668,767,548,572]" name="Vietnam" pageId="13" pageNumber="14">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName authority="Boutique, Bull. Jard. Bot. Etat Bruxelles 21: 97, 1951" authorityName="Boutique, Bull. Jard. Bot. Etat Bruxelles 21: 97" authorityYear="1951" class="Magnoliopsida" family="Annonaceae" genus="Toussaintia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Toussaintia" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
Toussaintia Boutique, Bull. Jard. Bot.
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Bruxelles 21: 97, 1951
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</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName genus="Tousaintia" lsidName="Tousaintia congolensis" pageId="0" pageNumber="1" rank="species" species="congolensis">
<emphasis italics="true" pageId="0" pageNumber="1">Tousaintia congolensis</emphasis>
</taxonomicName>
Boutique.
</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Same as species.</paragraph>
<paragraph pageId="0" pageNumber="1">
A small genus of four species (
<bibRefCitation DOI="https://doi.org/10.2982/0012-8317(2005)94[165:ANTAFT]2.0.CO;2" author="Luke, Q" journalOrPublisher="TAXON" pageId="0" pageNumber="1" refId="B186" refString="Luke, Q, Deroin, T, 2005. A new Toussaintia (Annonaceae) from Tanzania. Journal of East African Natural History 94: 165-174. https://doi.org/10.2982/0012-8317(2005)94[165:ANTAFT]2.0.CO;2" title="A new Toussaintia (Annonaceae) from Tanzania. Journal of East African Natural History 94: 165 - 174." url="https://doi.org/10.2982/0012-8317(2005)94[165:ANTAFT]2.0.CO;2" year="2005">Luke and Deroin 2005</bibRefCitation>
) distributed in central and east Africa, one species occurs in Cameroon, not endemic.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Boutique, Bull. Jard. Bot. Etat Bruxelles 21: 97" authorityYear="1951" class="Magnoliopsida" family="Annonaceae" genus="Toussaintia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Toussaintia" order="Magnoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Toussaintia</emphasis>
</taxonomicName>
is a genus of lianas or scandent shrubs, characterized by the presence of an androgynophore (elongated receptacle on which the stamens and carpels are inserted). All species are rare across their range.
</paragraph>
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<paragraph pageId="0" pageNumber="1">Taxonomy.</paragraph>
<paragraph pageId="0" pageNumber="1">
There is no revision available for this genus yet, but
<bibRefCitation DOI="https://doi.org/10.2982/0012-8317(2005)94[165:ANTAFT]2.0.CO;2" author="Luke, Q" journalOrPublisher="TAXON" pageId="0" pageNumber="1" refId="B186" refString="Luke, Q, Deroin, T, 2005. A new Toussaintia (Annonaceae) from Tanzania. Journal of East African Natural History 94: 165-174. https://doi.org/10.2982/0012-8317(2005)94[165:ANTAFT]2.0.CO;2" title="A new Toussaintia (Annonaceae) from Tanzania. Journal of East African Natural History 94: 165 - 174." url="https://doi.org/10.2982/0012-8317(2005)94[165:ANTAFT]2.0.CO;2" year="2005">Luke and Deroin (2005)</bibRefCitation>
provide a key to all four species.
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