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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.1.e977" ID-PMC="PMC3964716" ID-Pensoft-Pub="1314-2828--977" ID-PubMed="24723771" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1314-2828--e977" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Faunistic diversity of spiders (Araneae) in Galichitsa mountain (FYR Macedonia)" checkinTime="1553126911408" checkinUser="pensoft" docAuthor="Deltshev, Christo, Komnenov, Marjan, Blagoev, Gergin, Georgiev, Teodor, Lazarov, Stoyan, Stojkoska, Emilija &amp; Naumova, Maria" docDate="2013" docId="CF0744F12381826EC54A7DA4B4D7EBBB" docLanguage="en" docName="BiodivDatJour 1: e977" docOrigin="Biodiversity Data Journal 1" docSource="http://dx.doi.org/10.3897/BDJ.1.e977" docTitle="Crustulina guttata Wider 1834" docType="treatment" docVersion="5" lastPageNumber="977" masterDocId="230DFFA78F42A155FFC3FFA3301DFF8B" masterDocTitle="Faunistic diversity of spiders (Araneae) in Galichitsa mountain (FYR Macedonia)" masterLastPageNumber="977" masterPageNumber="977" pageNumber="977" updateTime="1668122337209" updateUser="ExternalLinkService">
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<mods:title>Faunistic diversity of spiders (Araneae) in Galichitsa mountain (FYR Macedonia)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Deltshev, Christo</mods:namePart>
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<mods:namePart>Komnenov, Marjan</mods:namePart>
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<mods:namePart>Blagoev, Gergin</mods:namePart>
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<mods:namePart>Georgiev, Teodor</mods:namePart>
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<mods:namePart>Lazarov, Stoyan</mods:namePart>
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<mods:namePart>Stojkoska, Emilija</mods:namePart>
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<mods:namePart>Naumova, Maria</mods:namePart>
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<mods:title>Biodiversity Data Journal</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>1</mods:number>
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<mods:start>977</mods:start>
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<taxonomicName class="Arachnida" family="Theridiidae" genus="Crustulina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crustulina" order="Araneae" pageId="0" pageNumber="977" phylum="Arthropoda" rank="genus">Crustulina</taxonomicName>
<taxonomicName lsidName="" order="Araneae" pageId="0" pageNumber="977" rank="order">Araneae</taxonomicName>
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<taxonomicName kingdom="Animalia" lsidName="" pageId="0" pageNumber="977" rank="kingdom">Animalia</taxonomicName>
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<taxonomicName authority="Wider, 1834" authorityName="Wider" authorityYear="1834" class="Arachnida" family="Theridiidae" genus="Crustulina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crustulina guttata" order="Araneae" pageId="0" pageNumber="977" phylum="Arthropoda" rank="species" species="guttata">Crustulina guttata (Wider, 1834)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="977" type="materials_examined">
<paragraph pageId="0" pageNumber="977">Materials</paragraph>
<paragraph pageId="0" pageNumber="977">Type status: Other material</paragraph>
<paragraph pageId="0" pageNumber="977">
Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="977">C. Deltshev &amp; E. Stojkoska</collectorName>
; sex:
<specimenType pageId="0" pageNumber="977">2 females</specimenType>
; Location: country:
<collectingCountry pageId="0" pageNumber="977">FYR of Macedonia</collectingCountry>
; locality:
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; verbatimElevation:
<elevation pageId="0" pageNumber="977">719 m</elevation>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="977">
<date value="2005-08-31">31-08-2005</date>
</collectingDate>
</paragraph>
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<paragraph pageId="0" pageNumber="977">Distribution</paragraph>
<paragraph pageId="0" pageNumber="977">Palearctic.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="977" type="notes">
<paragraph pageId="0" pageNumber="977">Notes</paragraph>
<paragraph pageId="0" pageNumber="977">
Previously recorded from unspecified locality between Ohrid and Resen (
<bibRefCitation author="Drensky, P." journalOrPublisher="Spisanie na Bulgarska Akademiya na Naukite" pageId="0" pageNumber="977" pagination="1 - 76" title="Spinnen (Araneae) aus Mittel und Sued-West Mazedonien" volume="39" year="1929">Drensky 1929</bibRefCitation>
,
<bibRefCitation author="Drensky, P" journalOrPublisher="Spisanie na Bulgarska Akademiya na Naukite" pageId="0" pageNumber="977" pagination="1 - 223" title="Katalog der echten Spinnen (Araneae) der Balkanhalbinsel" volume="32" year="1936">Drensky 1936</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<document ID-DOI="http://dx.doi.org/10.3897/jhr.87.72842" ID-GBIF-Dataset="3e3ca27b-44ad-4ea2-af6b-516245293ccf" ID-Pensoft-Pub="1314-2607-87-323" ID-Pensoft-UUID="D03A96052A8550F9918BB08ACA344FB9" ID-Zenodo-Dep="5811493" ID-ZooBank="E343379ED04447ABA1ED47B3F01F3E59" ModsDocID="1314-2607-87-323" checkinTime="1640348264987" checkinUser="pensoft" docAuthor="Talamas, Elijah J., Bremer, Jonathan S., Moore, Matthew R., Bon, Marie-Claude, Lahey, Zachary, Roberts, Cheryl G., Combee, Lynn A., McGathey, Natalie, van Noort, Simon, Timokhov, Alexander V., Hougardy, Evelyne &amp; Hogg, Brian" docDate="2021" docId="CF079475702851A2B2160CC06D07308C" docLanguage="en" docName="JourHymenoptRes 87: 323-480" docOrigin="Journal of Hymenoptera Research 87" docPubDate="2021-12-23" docSource="http://dx.doi.org/10.3897/jhr.87.72842" docTitle="Gryon crassifemoratum Mineo 1990" docType="treatment" docVersion="4" id="D03A96052A8550F9918BB08ACA344FB9" lastPageNumber="323" masterDocId="D03A96052A8550F9918BB08ACA344FB9" masterDocTitle="A maximalist approach to the systematics of a biological control agent: Gryon aetherium Talamas, sp. nov. (Hymenoptera, Scelionidae)" masterLastPageNumber="480" masterPageNumber="323" pageNumber="323" updateTime="1678757109313" updateUser="pensoft">
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<mods:title>A maximalist approach to the systematics of a biological control agent: Gryon aetherium Talamas, sp. nov. (Hymenoptera, Scelionidae)</mods:title>
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<mods:namePart>Talamas, Elijah J.</mods:namePart>
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<mods:affiliation>Florida Department of Agriculture and Consumer Services, Gainesville, FL, USA</mods:affiliation>
<mods:nameIdentifier type="email">elijah.talamas@fdacs.gov</mods:nameIdentifier>
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<mods:namePart>Bremer, Jonathan S.</mods:namePart>
<mods:affiliation>Florida Department of Agriculture and Consumer Services, Gainesville, FL, USA</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moore, Matthew R.</mods:namePart>
<mods:affiliation>Florida Department of Agriculture and Consumer Services, Gainesville, FL, USA</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bon, Marie-Claude</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5914-1682</mods:nameIdentifier>
<mods:affiliation>USDA-ARS-EBCL, Montpellier, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lahey, Zachary</mods:namePart>
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<mods:affiliation>Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, Columbus, OH, USA</mods:affiliation>
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<mods:namePart>Roberts, Cheryl G.</mods:namePart>
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<mods:namePart>Combee, Lynn A.</mods:namePart>
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<mods:namePart>Timokhov, Alexander V.</mods:namePart>
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<mods:affiliation>Lomonosov Moscow State University, Moscow, Russia</mods:affiliation>
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<mods:namePart>Hougardy, Evelyne</mods:namePart>
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<mods:affiliation>USDA-ARS-ISPH, Albany, CA, USA</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart>Hogg, Brian</mods:namePart>
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<taxonomicName ID-CoL="6L5RK" LSID="CF079475-7028-51A2-B216-0CC06D07308C" authority="Mineo" authorityName="Mineo" authorityYear="1990" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon crassifemoratum" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="crassifemoratum">Gryon crassifemoratum Mineo</taxonomicName>
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<taxonomicName authorityName="Mineo" authorityYear="1990" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon crassifemoratum" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="crassifemoratum">Gryon crassifemoratum</taxonomicName>
Mineo, 1990a: 181 (original description. Misspelled
<emphasis italics="true" pageId="0" pageNumber="323">crasifemaratum</emphasis>
in description, abstract; correct spelling (G. Mineo) in title); Johnson, 1992: 381 (cataloged, type information).
</paragraph>
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<subSubSection pageId="0" pageNumber="323" type="comments">
<paragraph pageId="0" pageNumber="323">Comments.</paragraph>
<paragraph pageId="0" pageNumber="323">
The original description for this species is woefully insufficient. We leave it in
<taxonomicName authorityName="Haliday" authorityYear="1833" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="323">Gryon</emphasis>
</taxonomicName>
based on its placement in the
<taxonomicName authorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" authorityYear="2021" baseAuthorityName="Talamas &amp; Bremer &amp; Moore &amp; Bon &amp; Lahey &amp; Roberts &amp; Combee &amp; McGathey &amp; van Noort &amp; Timokhov &amp; Hougardy &amp; Hogg" baseAuthorityYear="2021" class="Insecta" family="Scelionidae" genus="Gryon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryon myrmecophilum" order="Hymenoptera" pageId="0" pageNumber="323" phylum="Arthropoda" rank="species" species="myrmecophilum">
<emphasis italics="true" pageId="0" pageNumber="323">Gryon myrmecophilum</emphasis>
</taxonomicName>
species group (Mineo 1990).
</paragraph>
</subSubSection>
</treatment>
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<mods:identifier id="C0089970FF9376C548BBAD31DAD46F43" type="ZooBank">0C5E3F32-FB0B-4AF6-AD4F-9D96A1BA670B</mods:identifier>
</mods:mods>
<treatment id="CF079C1BFF8BFFD0FF5E283D9539FD5D" ID-DOI="http://doi.org/10.5281/zenodo.5618392" ID-GBIF-Taxon="127663037" ID-Zenodo-Dep="5618392" LSID="urn:lsid:plazi:treatment:CF079C1BFF8BFFD0FF5E283D9539FD5D" httpUri="http://treatment.plazi.org/id/CF079C1BFF8BFFD0FF5E283D9539FD5D" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
<subSubSection id="0FB47E86FF8BFFD3FF5E283D9345FC04" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="47112D0DFF8BFFD3FF5E283D9345FCE5" blockId="20.[151,544,858,917]" box="[151,544,858,884]" pageId="20" pageNumber="21">
<heading id="1C599A61FF8BFFD3FF5E283D9345FCE5" bold="true" box="[151,544,858,884]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="75DAF11FFF8BFFD3FF5E283D9345FCE5" bold="true" box="[151,544,858,884]" pageId="20" pageNumber="21">
<taxonomicName id="80AE568EFF8BFFD3FF5E283D90D9FCE5" ID-CoL="7TVP2" box="[151,444,858,884]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF8BFFD3FF5E283D90D9FCE5" bold="true" box="[151,444,858,884]" italics="true" pageId="20" pageNumber="21">Chiromantes eurymerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF8BFFD3FE0B283D9345FCE5" box="[450,544,858,884]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="47112D0DFF8BFFD3FF5E281A9345FC04" blockId="20.[151,544,858,917]" box="[151,544,893,917]" pageId="20" pageNumber="21">
(
<figureCitation id="DF953188FF8BFFD3FF56281A9180FC04" box="[159,229,893,917]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="20" pageNumber="21">
Figs
<date id="33100BCDFF8BFFD3FF1E281A9180FC04" box="[215,229,893,917]" pageId="20" pageNumber="21">3</date>
</figureCitation>
I, J, 4D, 5E, 6E, 7E, 9E, 14)
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8BFFD3FF5E28D99245FA17" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="47112D0DFF8BFFD3FF5E28D9929AFAAF" blockId="20.[151,1436,958,2026]" pageId="20" pageNumber="21">
<emphasis id="75DAF11FFF8BFFD3FF5E28D9901AFC46" bold="true" box="[151,383,958,983]" pageId="20" pageNumber="21">Material Examined</emphasis>
.
<typeStatus id="981593AFFF8BFFD3FE4428D99340FC47" box="[397,549,958,982]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
: NMNS-7028-002, male (18.7×
<quantity id="805680E8FF8BFFD3FC4228D99291FC47" box="[907,1012,958,982]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="20" pageNumber="21" unit="mm" value="14.8">14.8 mm</quantity>
), west coast of Lanyu Island,
<collectingCountry id="3FB96D9DFF8BFFD3FA8B28D994F2FC47" box="[1346,1431,958,982]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
, calcareous rock terraces, spray zone, C.D. Schubart &amp; H.-C. Liu, 20
<date id="33100BCDFF8BFFD3FC602885954EFC6B" box="[937,1067,994,1018]" pageId="20" pageNumber="21" value="1999-09-21">21.09.1999</date>
.
<typeStatus id="981593AFFF8BFFD3FBFF288595B0FC6B" box="[1078,1237,994,1018]" pageId="20" pageNumber="21" type="paratype">PARATYPES</typeStatus>
: ZRC 2000.1823, Pingtung County, Hsiang Chiaowan,
<collectingCountry id="3FB96D9DFF8BFFD3FDF02F6193EBFB8F" box="[569,654,1030,1054]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
, H.-C. Liu,
<date id="33100BCDFF8BFFD3FCDC2F6192F3FB8F" box="[789,918,1030,1054]" pageId="20" pageNumber="21" value="1999-08-25">25.08.1999</date>
, male (18.2×14.0 mm),
<specimenCount id="51A8E684FF8BFFD3FB6F2F619477FB8F" box="[1190,1298,1030,1054]" pageId="20" pageNumber="21" type="female">2 females</specimenCount>
(17.0×13.2, 19.9×15.0 mm).—ZRC 2000.1822, male (13.3×
<quantity id="805680E8FF8BFFD3FD662F4D927DFBD3" box="[687,792,1066,1090]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="20" pageNumber="21" unit="mm" value="10.2">10.2 mm</quantity>
), Pingtung County, Hsiang Chiaowan,
<collectingCountry id="3FB96D9DFF8BFFD3FB002F4D947BFBD3" box="[1225,1310,1066,1090]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
, H.-C. Liu, 5.07.1999.—QM-W29171, male (20.0×
<quantity id="805680E8FF8BFFD3FD922F2993A3FBF7" box="[603,710,1102,1126]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="20" pageNumber="21" unit="mm" value="15.4">15.4 mm</quantity>
), west coast of Lanyu Island,
<collectingCountry id="3FB96D9DFF8BFFD3FBE82F299513FBF7" box="[1057,1142,1102,1126]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
, calcareous rock terraces, spray zone, C.D. Schubart &amp; H.-C. Liu, 20
<date id="33100BCDFF8BFFD3FD012F15922CFB1B" box="[712,841,1138,1162]" pageId="20" pageNumber="21" value="1999-09-21">21.09.1999</date>
. NMMBA-3504b, male (19.3×
<quantity id="805680E8FF8BFFD3FB082F159456FB1B" box="[1217,1331,1138,1162]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="20" pageNumber="21" unit="mm" value="14.8">14.8 mm</quantity>
), female (22.6×
<quantity id="805680E8FF8BFFD3FF282FF1902FFB3F" box="[225,330,1174,1198]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="20" pageNumber="21" unit="mm" value="16.7">16.7 mm</quantity>
), Hsiang-Chiau-Wan, Kenting, Pingtung County,
<collectingCountry id="3FB96D9DFF8BFFD3FCB42FF192B7FB3F" box="[893,978,1174,1198]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
, C.C. Li,
<date id="33100BCDFF8BFFD3FB8A2FF195A0FB3F" box="[1091,1221,1174,1198]" pageId="20" pageNumber="21" value="2012-08-23">23.08.2012</date>
.—NMNS,
<specimenCount id="51A8E684FF8BFFD3FA8D2FF194FEFB3F" box="[1348,1435,1174,1198]" pageId="20" pageNumber="21" type="male">2 males</specimenCount>
(17.5×13.4, 16.7×
<quantity id="805680E8FF8BFFD3FEAD2FDD90AAFB43" box="[356,463,1210,1234]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="20" pageNumber="21" unit="mm" value="12.3">12.3 mm</quantity>
), Siatanzai, Kenting, Pingtung County,
<collectingCountry id="3FB96D9DFF8BFFD3FC532FDD928AFB43" box="[922,1007,1210,1234]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
,
<geoCoordinate id="229A4BCAFF8BFFD3FC342FDD95FBFB43" box="[1021,1182,1210,1234]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="21.926832">21°5536.6”N</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF8BFFD3FB642FDD9432FB43" box="[1197,1367,1210,1234]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-120.73939">120°4421.8”S</geoCoordinate>
, J.-H. Lee &amp; W.-J. Wang,
<date id="33100BCDFF8BFFD3FE4C2FB99361FB67" box="[389,516,1246,1270]" pageId="20" pageNumber="21" value="2012-08-17">17.08.2012</date>
.—ZRC 2012.0956, male (15.4×
<quantity id="805680E8FF8BFFD3FCB62FB99288FB67" box="[895,1005,1246,1270]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="20" pageNumber="21" unit="mm" value="12.3">12.3 mm</quantity>
), ovigerous female (18.0×
<quantity id="805680E8FF8BFFD3FAEB2FB994F5FB67" box="[1314,1424,1246,1270]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="20" pageNumber="21" unit="mm" value="13.4">13.4 mm</quantity>
), Siatanzai, Kenting, Pingtung County,
<geoCoordinate id="229A4BCAFF8BFFD3FDBC2E659270FA8B" box="[629,789,1282,1306]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="21.926832">21°5536.6”N</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF8BFFD3FCE72E6592BCFA8B" box="[814,985,1282,1306]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-120.73939">120°4421.8”S</geoCoordinate>
,
<collectingCountry id="3FB96D9DFF8BFFD3FC382E659523FA8B" box="[1009,1094,1282,1306]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
, J.-H. Lee &amp; W.-J. Wang,
<date id="33100BCDFF8BFFD3FF5E2E41907CFAAF" box="[151,281,1318,1342]" pageId="20" pageNumber="21" value="2012-08-31">31.08.2012</date>
.—NMNS, female (18.8×
<quantity id="805680E8FF8BFFD3FDFA2E4193F9FAAF" box="[563,668,1318,1342]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="20" pageNumber="21" unit="mm" value="13.6">13.6 mm</quantity>
), same data as ZRC 2012.0956.
</paragraph>
<paragraph id="47112D0DFF8BFFD3FF0E2E2D9245FA17" blockId="20.[151,1436,958,2026]" pageId="20" pageNumber="21">
NON-TYPE: ZRC 2012.0957, dried female exuvium (24.8×
<quantity id="805680E8FF8BFFD3FCA32E2D92B4FAF3" box="[874,977,1354,1378]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="20" pageNumber="21" unit="mm" value="18.7">18.7 mm</quantity>
), Dingtanzai, near nuclear power station, Kenting, Pingtung County,
<collectingCountry id="3FB96D9DFF8BFFD3FE002E09937BFA17" box="[457,542,1390,1414]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
, P.K.L. Ng,
<date id="33100BCDFF8BFFD3FD612E099279FA17" box="[680,796,1390,1414]" pageId="20" pageNumber="21" value="2012-04-10">4.10.2012</date>
.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8BFFD3FF0E2EF5958AF97F" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="47112D0DFF8BFFD3FF0E2EF5958AF97F" blockId="20.[151,1436,958,2026]" pageId="20" pageNumber="21">
<emphasis id="75DAF11FFF8BFFD3FF0E2EF5905FFA3A" bold="true" box="[199,314,1426,1451]" pageId="20" pageNumber="21">Diagnosis</emphasis>
. Carapace transversely subovate, c. 1.3 times broader than long; dorsal carapace, lateral branchial regions markedly swollen; exorbital tooth with outer margin broadly convex marking widest point of carapace; front c. 0.650.7 times carapace width, margin broadly convex in frontal view, but medial part relatively straight in dorsal view, appears smooth but microscopically granular, with pair of low lateral swellings behind margin; supraorbital margin entire, semicircular; dorsal surface of carpus of cheliped covered in small granules; walking legs relatively short; merus of P4 c. 2.0 times longer than wide; that of P5 similar at 2.02.1 times longer; propodus of P5 also 2.12.2 times; male abdomen (
<figureCitation id="DF953188FF8BFFD3FDA12D0D93DDF913" box="[616,696,1642,1666]" captionStart="FIGURE 14" captionStartId="22.[151,250,1903,1926]" captionTargetBox="[195,1402,207,1854]" captionTargetId="figure@22.[169,1429,193,1875]" captionTargetPageId="22" captionText="FIGURE 14. Chiromantes eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, left G 2; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1. All structures denuded. Scales: A F = 1.0 mm; G, H = 0.5 mm." httpUri="https://zenodo.org/record/283295/files/figure.png" pageId="20" pageNumber="21">Fig. 14</figureCitation>
C) moderately broad; somite 6 with distolateral margins relatively straight, strongly divergent, margins convex overall; somite 3 width 3.1 times basal width of telson; G1 (
<figureCitation id="DF953188FF8BFFD3FAFA2DE994E1F937" box="[1331,1412,1678,1702]" captionStart="FIGURE 14" captionStartId="22.[151,250,1903,1926]" captionTargetBox="[195,1402,207,1854]" captionTargetId="figure@22.[169,1429,193,1875]" captionTargetPageId="22" captionText="FIGURE 14. Chiromantes eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, left G 2; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1. All structures denuded. Scales: A F = 1.0 mm; G, H = 0.5 mm." httpUri="https://zenodo.org/record/283295/files/figure.png" pageId="20" pageNumber="21">Fig. 14</figureCitation>
E, G, H) markedly stout, with margins subparallel to obtusely angled subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process short, not projecting past line of gonopod shaft, moderately tapering.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8BFFD2FF0E2D9D90C2F94E" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="47112D0DFF8BFFD2FF0E2D9D9396FE65" blockId="20.[151,1436,958,2026]" lastBlockId="21.[151,1437,151,2012]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="75DAF11FFF8BFFD3FF0E2D9D9034F882" bold="true" box="[199,337,1786,1811]" pageId="20" pageNumber="21">Description</emphasis>
. Carapace transversely subovate, c. 1.3 times (range 1.291.32) broader than long; surface bare, lacking setal tufts; covered with fine, but distinct low granules becoming arranged into short striae posteriorly; mesogastric regions well defined; lateral carapace surface generally without obvious discrete oblique striae, except for strong, concave epibranchial sulcus beginning just behind exorbital tooth. Dorsal carapace, lateral branchial regions markedly swollen. Exorbital tooth triangular, pointed, outer margin broadly convex marking widest point of carapace, margin not constricted behind (sometimes slightly flattened) such that entire lateral border appears convex, no trace of second anterolateral tooth. Postfrontal lobes well demarcated but relatively low, rounded, separated by broad grooves, median lobes similar in width to lateral lobes. Front c. 0.650.7 times carapace width (
<quantity id="805680E8FF8AFFD2FF572BDB918BFF45" box="[158,238,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.651" pageId="21" pageNumber="22" unit="in" value="0.65">0.65 in</quantity>
<typeStatus id="981593AFFF8AFFD2FF3C2BDB903EFF45" box="[245,347,188,212]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
), markedly deflexed, margin broadly convex in frontal view, but medial part relatively straight in dorsal view; laterally triangular, bluntly pointed; frontal margin emarginate, microscopically granular but appearing relatively smooth; surface relatively flat behind, but with pair of low lateral swellings behind frontal margin. Supraorbital margin entire, semicircular but oblique medial section somewhat straight; secondary rim formed either side making a smooth, broadly triangular, deflexed central plate behind ocular peduncle. Eye not extending beyond exorbital tooth. Frontal plate protruding as a shelf; medial septum broad, largely covering anterior half of anterior half of epistome; antennae, antennules much reduced in size, lodged under overhanging front; basal antennular segment not much swollen; antennal, antennular basal segments adjacent, not separated by septum. Antennal flagellum very short, entering orbit.
</paragraph>
<paragraph id="47112D0DFF8AFFD2FF0E2A9895FDFD15" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">Third maxillipeds with ischium, merus subequal in length; ischium with shallow longitudinal, curved, median sulcus. Inner margin of merus, ischium with long setae, proximal outer margin of ischium, base of exopod with dense setae; outer margin of merus with short scattered setae only. Exopod slender, hidden behind ishium, merus except near base, tip reaching half length of outer margin of merus, flagellum long, slender.</paragraph>
<paragraph id="47112D0DFF8AFFD2FF0E29E893F0FB9E" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
Chelipeds subequal, robust. Merus trihedral; posterior border broadly convex emarginate, minutely granular, without indication of subdistal spine or lobe; inner anterior border minutely granular, broadly triangular; outer surface broadly convex, with fine transverse striae but appearing almost smooth. Carpus subquadrilateral, inner angle moderately produced, apically granular; outer margin, dorsal surface covered in small granules. Palm dorsal surface slightly striated along margin, with scattering of moderate sized granules, otherwise smooth, punctate. Palm outer surface broadly rounded, smooth; no indication of subventral longitudinal ridge. Inner face of palm of males smooth except for curved projecting crest of large tubercles behind gape (
<figureCitation id="DF953188FF8AFFD2FBED2800950DFCEE" box="[1060,1128,871,895]" captionStart="FIGURE 6" captionStartId="9.[151,250,1537,1560]" captionTargetBox="[151,1435,193,1515]" captionTargetId="figure@9.[151,1436,193,1516]" captionTargetPageId="9" captionText="FIGURE 6. Inner views of chelae. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283287/files/figure.png" pageId="21" pageNumber="22">Fig. 6</figureCitation>
E). Cutting margin of fixed finger typically with 3 or 4 blunt teeth, proximal 3 similar in size, distal-most larger, rounded, conical, placed medially; dactylus with 2 teeth proximally, distal-most being large, third much smaller blunt tooth slightly distal of medial tooth of ventral finger. Dorsal surface of dactylus smooth. Fingers with tips chitinous; adult males with narrow but distinct gape when fingers closed.
</paragraph>
<paragraph id="47112D0DFF8AFFD2FF0E2F7B947DFB76" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">Walking legs relatively short, broad, flattened; second, third pairs sub-equal, longer than other walking legs. Without setae except for scattering of short dark bristles on dactyli, ventral face of propodi. Dorsal margins of meri with blunt subdistal shoulder, but otherwise unarmed; outer surfaces of meri with minutely granular transverse striae. Merus of P4 c. 2.0 times longer than wide; that of P5 similar at 2.02.1 times longer. Carpi with 2 accessory carinae on outer surface but not strongly marked. Propodus of P4 c. 2.12.2 times longer than wide; that of P5 also 2.12.2 times. Dactyli 0.8 times length of propodi, slightly recurved, terminating in acute chitinous tip.</paragraph>
<paragraph id="47112D0DFF8AFFD2FF0E2F939446FA51" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
Thoracic sternites (
<figureCitation id="DF953188FF8AFFD2FE562F939087FA9D" box="[415,482,1268,1292]" captionStart="FIGURE 9" captionStartId="12.[151,250,1576,1599]" captionTargetBox="[151,1435,325,1554]" captionTargetId="figure@12.[151,1436,325,1555]" captionTargetPageId="12" captionText="FIGURE 9. Ventral views of thoracic sternum and anterior parts of male abdomens. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283290/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
E) smooth, mostly bare of setae except for a few scattered bristles; abdominal cavity reaching to thick transverse setal fringe at the junction of sternites 3, 4. Male abdomen (
<figureCitation id="DF953188FF8AFFD2FB6A2E709592FABE" box="[1187,1271,1303,1327]" captionStart="FIGURE 14" captionStartId="22.[151,250,1903,1926]" captionTargetBox="[195,1402,207,1854]" captionTargetId="figure@22.[169,1429,193,1875]" captionTargetPageId="22" captionText="FIGURE 14. Chiromantes eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, left G 2; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1. All structures denuded. Scales: A F = 1.0 mm; G, H = 0.5 mm." httpUri="https://zenodo.org/record/283295/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
C) moderately broad; telson broadly rounded apically, 1.2 times longer than wide, markedly longer than somite 6 (1.41.5 times); somite 6 c. 2.3 times wider than long, distolateral margins relatively straight, strongly divergent, margins convex overall; somites 35 trapezoid, lateral margins of somites 4, 5 relatively straight, lateral margins of somite 3 convex; somite 3 width 3.1 times basal width of telson; somites 1, 2 transversely, longitudinally narrow.
</paragraph>
<paragraph id="47112D0DFF8AFFD2FF0E2EAB920FF9BD" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
G1 (
<figureCitation id="DF953188FF8AFFD2FF332EAB9029FA75" box="[250,332,1484,1508]" captionStart="FIGURE 14" captionStartId="22.[151,250,1903,1926]" captionTargetBox="[195,1402,207,1854]" captionTargetId="figure@22.[169,1429,193,1875]" captionTargetPageId="22" captionText="FIGURE 14. Chiromantes eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, left G 2; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1. All structures denuded. Scales: A F = 1.0 mm; G, H = 0.5 mm." httpUri="https://zenodo.org/record/283295/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
E, G, H) markedly stout, relatively straight, with margins subparallel to obtusely angled subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process short, not projecting past line of gonopod shaft, moderately tapering. G2 short, as for the genus (
<figureCitation id="DF953188FF8AFFD2FD302D73922FF9BD" box="[761,842,1556,1580]" captionStart="FIGURE 14" captionStartId="22.[151,250,1903,1926]" captionTargetBox="[195,1402,207,1854]" captionTargetId="figure@22.[169,1429,193,1875]" captionTargetPageId="22" captionText="FIGURE 14. Chiromantes eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, left G 2; G, ventral view of distal part of left G 1; H, dorsal view of distal part of left G 1. All structures denuded. Scales: A F = 1.0 mm; G, H = 0.5 mm." httpUri="https://zenodo.org/record/283295/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
F).
</paragraph>
<paragraph id="47112D0DFF8AFFD2FF0E2D5F9397F9E5" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
<emphasis id="75DAF11FFF8AFFD2FF0E2D5F904CF9C1" bold="true" box="[199,297,1592,1616]" pageId="21" pageNumber="22">Females</emphasis>
. Chelipeds slightly smaller, less robust, lacking raised granular row on inner face of chela. Vulval morphology without clear species-specific characters.
</paragraph>
<paragraph id="47112D0DFF8AFFD2FF0E2D1890C2F94E" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
<emphasis id="75DAF11FFF8AFFD2FF0E2D189003F909" bold="true" box="[199,358,1663,1688]" pageId="21" pageNumber="22">Colour in life</emphasis>
. Carapaces of adult males and females are generally pale grey and white to yellowish-brown to yellowish-grey; with chelae of both sexes, notably dactylus, white, without obvious pink or red; eyes blue to bluish-green (
<figureCitation id="DF953188FF8AFFD2FEFB2DA09011F94E" box="[306,372,1735,1759]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="21" pageNumber="22">
Fig.
<date id="33100BCDFF8AFFD2FEAF2DA09011F94E" box="[358,372,1735,1759]" pageId="21" pageNumber="22">3</date>
</figureCitation>
I, J).
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8AFFD0FF0E2D8B901BFE5E" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="discussion">
<paragraph id="47112D0DFF8AFFD2FF0E2D8B953DF84D" blockId="21.[151,1437,151,2012]" pageId="21" pageNumber="22">
<emphasis id="75DAF11FFF8AFFD2FF0E2D8B9051F895" bold="true" box="[199,308,1772,1796]" pageId="21" pageNumber="22">Remarks</emphasis>
.
<taxonomicName id="80AE568EFF8AFFD2FE882D8B9328F895" box="[321,589,1772,1796]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF8AFFD2FE882D8B9328F895" box="[321,589,1772,1796]" italics="true" pageId="21" pageNumber="22">Chiromantes eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF8AFFD2FD9D2D8B93C8F895" bold="true" box="[596,685,1772,1796]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="6EE94C64FF8AFFD2FD9D2D8B93C8F895" box="[596,685,1772,1796]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is separated from its sympatric Taiwanese congener,
<taxonomicName id="80AE568EFF8AFFD2FB362D8A94FEF895" box="[1279,1435,1772,1796]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF8AFFD2FB362D8A94FEF895" box="[1279,1435,1772,1796]" italics="true" pageId="21" pageNumber="22">C. leptomerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF8AFFD2FF5E2C779195F8B9" box="[151,240,1808,1832]" pageId="21" pageNumber="22" rank="species">
<emphasis id="75DAF11FFF8AFFD2FF5E2C779195F8B9" bold="true" box="[151,240,1808,1832]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
, by its obviously broader legs, and by its stouter G1 with its much shortened tip, a unique character within the species-group, and which easily separates it from all other species described here. Numerous other features that distinguish
<taxonomicName id="80AE568EFF8AFFD2FED42C3F90DDF8E1" box="[285,440,1880,1904]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="75DAF11FFF8AFFD2FED42C3F90DDF8E1" box="[285,440,1880,1904]" italics="true" pageId="21" pageNumber="22">C. eurymerus</emphasis>
</taxonomicName>
from other related species are given in
<tableCitation id="0A2C18B6FF8AFFD2FC5C2C309288F8FE" box="[917,1005,1879,1903]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[159,1428,329,1954]" captionTargetPageId="5" captionText="TABLE 1. Summary of major differences between species of the Chiromantes obtusifrons complex." httpUri="http://table.plazi.org/id/13D17D85FF9AFFC2FF5E2BF095A0FF3E" pageId="21" pageNumber="22" tableUuid="13D17D85FF9AFFC2FF5E2BF095A0FF3E">Table 1</tableCitation>
.
<taxonomicName id="80AE568EFF8AFFD2FC372C3F9470F8E1" box="[1022,1301,1880,1904]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF8AFFD2FC372C3F9470F8E1" box="[1022,1301,1880,1904]" italics="true" pageId="21" pageNumber="22">Chiromantes leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF8AFFD2FAE92C3F941BF8E1" bold="true" box="[1312,1406,1880,1904]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="6EE94C64FF8AFFD2FAE92C3F941BF8E1" box="[1312,1406,1880,1904]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="80AE568EFF8AFFD2FEC62C1A90D5F805" box="[271,432,1916,1940]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="obtusifrons" status="sp. nov.">
<emphasis id="75DAF11FFF8AFFD2FEC62C1A90D5F805" box="[271,432,1916,1940]" italics="true" pageId="21" pageNumber="22">C. obtusifrons</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF8AFFD2FE722C1B937DF805" bold="true" box="[443,536,1916,1940]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="6EE94C64FF8AFFD2FE722C1B937DF805" box="[443,536,1916,1940]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having the outer margin of the exorbital tooth broadly convex, and in not having the carapace laterally constricted behind the exorbital tooth. It differs from that species by having a less conspicuously granular carapace and claws, and by the stouter G1 already mentioned.
</paragraph>
<caption id="13D17D85FF89FFD1FF5E2C0893EDF87A" httpUri="https://zenodo.org/record/283295/files/figure.png" pageId="22" pageNumber="23" targetBox="[195,1402,207,1854]" targetPageId="22">
<paragraph id="47112D0DFF89FFD1FF5E2C0893EDF87A" blockId="22.[151,1436,1903,2027]" pageId="22" pageNumber="23">
<emphasis id="75DAF11FFF89FFD1FF5E2C089040F814" bold="true" box="[151,293,1903,1926]" pageId="22" pageNumber="23">FIGURE 14.</emphasis>
<taxonomicName id="80AE568EFF89FFD1FEF92C08934DF817" box="[304,552,1903,1926]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF89FFD1FEF92C08934DF817" box="[304,552,1903,1926]" italics="true" pageId="22" pageNumber="23">Chiromantes eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF89FFD1FDFD2C1793ECF814" bold="true" box="[564,649,1904,1925]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="6EE94C64FF89FFD1FDFD2C1793ECF814" box="[564,649,1904,1925]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS-7028-002, holotype male (18.7×14.8 mm), Taiwan. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 36, telson); D, ventral view of left G1; E, dorsal view of left G1; F, left G2; G, ventral view of distal part of left G1; H, dorsal view of distal part of left G1. All structures denuded. Scales: AF = 1.0 mm; G, H = 0.5 mm.
</paragraph>
</caption>
<paragraph id="47112D0DFF88FFD0FF0E2BF0901BFE5E" blockId="23.[151,1436,151,716]" pageId="23" pageNumber="24">
Both species also appear to be distinctively coloured. In
<taxonomicName id="80AE568EFF88FFD0FC902BFF929DFF21" box="[857,1016,152,176]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="75DAF11FFF88FFD0FC902BFF929DFF21" box="[857,1016,152,176]" italics="true" pageId="23" pageNumber="24">C. leptomerus</emphasis>
</taxonomicName>
, corneas are typically light green to yellowish green (
<figureCitation id="DF953188FF88FFD0FE932BDB90FEFF45" box="[346,411,188,212]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="23" pageNumber="24">Fig. 3</figureCitation>
G, H), whereas in
<taxonomicName id="80AE568EFF88FFD0FDA32BDA9267FF45" box="[618,770,189,212]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="75DAF11FFF88FFD0FDA32BDA9267FF45" box="[618,770,189,212]" italics="true" pageId="23" pageNumber="24">C. eurymerus</emphasis>
</taxonomicName>
, the eyes are blue to bluish-green (
<figureCitation id="DF953188FF88FFD0FB582BDB95B1FF45" box="[1169,1236,188,212]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="23" pageNumber="24">Fig. 3</figureCitation>
J). The colours of the carapace and chelae also appear different, although the reliability of these characters will need to be tested with more specimens. The carapace background colour of adult males of
<taxonomicName id="80AE568EFF88FFD0FC662A629535FE8D" box="[943,1104,260,284]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="75DAF11FFF88FFD0FC662A629535FE8D" box="[943,1104,260,284]" italics="true" pageId="23" pageNumber="24">C. leptomerus</emphasis>
</taxonomicName>
varies from pale brown to a darker grey-green (
<figureCitation id="DF953188FF88FFD0FEBA2A4090D2FEAE" box="[371,439,295,319]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="23" pageNumber="24">Fig. 3</figureCitation>
DH); whereas in
<taxonomicName id="80AE568EFF88FFD0FD472A4F9242FED1" box="[654,807,296,320]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="75DAF11FFF88FFD0FD472A4F9242FED1" box="[654,807,296,320]" italics="true" pageId="23" pageNumber="24">C. eurymerus</emphasis>
</taxonomicName>
adult males (and females) are generally pale grey and white to yellowish-brown to yellowish-grey (
<figureCitation id="DF953188FF88FFD0FD682A2B9383FEF5" box="[673,742,332,356]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="23" pageNumber="24">
Fig.
<date id="33100BCDFF88FFD0FD112A2B9383FEF5" box="[728,742,332,356]" pageId="23" pageNumber="24">3</date>
</figureCitation>
I, J). Finally, in
<taxonomicName id="80AE568EFF88FFD0FC6D2A2A9521FEF5" box="[932,1092,332,356]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="75DAF11FFF88FFD0FC6D2A2A9521FEF5" box="[932,1092,332,356]" italics="true" pageId="23" pageNumber="24">C. leptomerus</emphasis>
</taxonomicName>
the chelae of adult males are often reddish-pink to purplish red overall, especially on the dorsal surfaces and dactyli (
<figureCitation id="DF953188FF88FFD0FB422A0895AAFE16" box="[1163,1231,367,391]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="23" pageNumber="24">Fig. 3</figureCitation>
H); whereas in
<taxonomicName id="80AE568EFF88FFD0FA4B2A179068FE3D" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="75DAF11FFF88FFD0FA4B2A179068FE3D" italics="true" pageId="23" pageNumber="24">C. eurymerus</emphasis>
</taxonomicName>
the chelae of both sexes, and notably the dactyli are yellowish to off-white, without any obvious pink or red tinging (
<figureCitation id="DF953188FF88FFD0FEE92AD09007FE5E" box="[288,354,439,463]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="23" pageNumber="24">Fig. 3</figureCitation>
J).
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF88FFD0FF0E2ABB93C1FDAD" pageId="23" pageNumber="24" type="etymology">
<paragraph id="47112D0DFF88FFD0FF0E2ABB93C1FDAD" blockId="23.[151,1436,151,716]" pageId="23" pageNumber="24">
<emphasis id="75DAF11FFF88FFD0FF0E2ABB9023FE65" bold="true" box="[199,326,476,500]" pageId="23" pageNumber="24">Etymology</emphasis>
. Derived from the Greek
<emphasis id="75DAF11FFF88FFD0FDB32ABA93C9FE65" box="[634,684,477,500]" italics="true" pageId="23" pageNumber="24">eury</emphasis>
meaning “broad”, and
<emphasis id="75DAF11FFF88FFD0FC772ABA9567FE65" box="[958,1026,477,500]" italics="true" pageId="23" pageNumber="24">meros</emphasis>
meaning “thigh”, and refers to the relatively shorter and stouter legs of this species compared to the other sympatric Taiwanese species,
<taxonomicName id="80AE568EFF88FFD0FB36296794FEFD89" box="[1279,1435,512,536]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF88FFD0FB36296794FEFD89" box="[1279,1435,512,536]" italics="true" pageId="23" pageNumber="24">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF88FFD0FF5E29439195FDAD" bold="true" box="[151,240,548,572]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="6EE94C64FF88FFD0FF5E29439195FDAD" box="[151,240,548,572]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It is used here as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF88FFD0FF0E29209539FD5D" pageId="23" pageNumber="24" type="distribution">
<paragraph id="47112D0DFF88FFD0FF0E2920933EFD15" blockId="23.[151,1436,151,716]" pageId="23" pageNumber="24">
<emphasis id="75DAF11FFF88FFD0FF0E2920903CFDF1" bold="true" box="[199,345,583,608]" pageId="23" pageNumber="24">Distribution</emphasis>
. So far only known from
<collectingCountry id="3FB96D9DFF88FFD0FD512920938BFDCE" box="[664,750,583,607]" name="Taiwan" pageId="23" pageNumber="24">Taiwan</collectingCountry>
. We are pleased to thank Peter Castro for his careful and constructive criticism of the manuscript.
</paragraph>
<paragraph id="47112D0DFF88FFD0FF0E29F79539FD5D" blockId="23.[151,1436,151,716]" pageId="23" pageNumber="24">
<emphasis id="75DAF11FFF88FFD0FF0E29F79041FD39" bold="true" box="[199,292,656,680]" pageId="23" pageNumber="24">Ecology</emphasis>
. Supratidal; calcareous rock terraces, spray zone. Found comingled with
<taxonomicName id="80AE568EFF88FFD0FB9029F79593FD39" box="[1113,1270,656,680]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF88FFD0FB9029F79593FD39" box="[1113,1270,656,680]" italics="true" pageId="23" pageNumber="24">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF88FFD0FB3529F79433FD39" bold="true" box="[1276,1366,656,680]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="6EE94C64FF88FFD0FB3529F79433FD39" box="[1276,1366,656,680]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the same shore habitat; the nature and extent of niche separation has not been determined.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,394 @@
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<mods:title id="BF7D74EFB7D37995BAC075B916EAC058">A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island (Indian Ocean), Guam and Taiwan</mods:title>
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<subSubSection id="0FB47E86FF8DFFD5FF5E2BF0921FFE87" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="47112D0DFF8DFFD5FF5E2BF09340FF20" blockId="18.[151,655,151,210]" box="[151,549,151,177]" pageId="18" pageNumber="19">
<heading id="1C599A61FF8DFFD5FF5E2BF09340FF20" bold="true" box="[151,549,151,177]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="75DAF11FFF8DFFD5FF5E2BF09340FF20" bold="true" box="[151,549,151,177]" pageId="18" pageNumber="19">
<taxonomicName id="80AE568EFF8DFFD5FF5E2BF090A5FF20" ID-CoL="7TVP4" box="[151,448,151,177]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF8DFFD5FF5E2BF090A5FF20" bold="true" box="[151,448,151,177]" italics="true" pageId="18" pageNumber="19">Chiromantes leptomerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF8DFFD5FE0E2BF09340FF20" box="[455,549,151,177]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="47112D0DFF8DFFD5FF5E2BDD93EAFF43" blockId="18.[151,655,151,210]" box="[151,655,186,210]" pageId="18" pageNumber="19">
(
<figureCitation id="DF953188FF8DFFD5FF562BDD9186FF43" box="[159,227,186,210]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="18" pageNumber="19">Figs 3</figureCitation>
DF, 4C, 5D, 6D, 7D, 8C, 9D, 13FJ)
</paragraph>
<paragraph id="47112D0DFF8DFFD5FF5E2A67921FFE87" blockId="18.[151,890,256,278]" box="[151,890,256,278]" pageId="18" pageNumber="19">
<taxonomicName id="80AE568EFF8DFFD5FF5E2A6790F5FE87" box="[151,400,256,278]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF8DFFD5FF5E2A6790F5FE87" box="[151,400,256,278]" italics="true" pageId="18" pageNumber="19">Chiromantes obtusifrons</emphasis>
</taxonomicName>
—Lin
<emphasis id="75DAF11FFF8DFFD5FE1A2A669363FE87" box="[467,518,256,278]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2011: 45 (with colour photographs).
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8DFFD5FF5E2A5893FCFA19" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="47112D0DFF8DFFD5FF5E2A589525FC95" blockId="18.[151,1437,319,2023]" pageId="18" pageNumber="19">
<emphasis id="75DAF11FFF8DFFD5FF5E2A58901AFEC9" bold="true" box="[151,383,319,344]" pageId="18" pageNumber="19">Material Examined</emphasis>
.
<typeStatus id="981593AFFF8DFFD5FE442A589340FEC6" box="[397,549,319,343]" pageId="18" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
: NMNS-7028-001, male (16.8×
<quantity id="805680E8FF8DFFD5FC412A589294FEC6" box="[904,1009,319,343]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="19" unit="mm" value="12.7">12.7 mm</quantity>
), Pingtung County, Hsiang Chiaowan,
<collectingCountry id="3FB96D9DFF8DFFD5FF5E2A049189FEEA" box="[151,236,355,379]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
, H.-C. Liu,
<date id="33100BCDFF8DFFD5FEB22A049098FEEA" box="[379,509,355,379]" pageId="18" pageNumber="19" value="1999-08-25">25.08.1999</date>
.
<typeStatus id="981593AFFF8DFFD5FDC22A0493CFFEEA" box="[523,682,355,379]" pageId="18" pageNumber="19" type="paratype">PARATYPES</typeStatus>
: ZRC 2000.1860,
<specimenCount id="51A8E684FF8DFFD5FC492A0492BCFEEA" box="[896,985,355,379]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(13.4×
<quantity id="805680E8FF8DFFD5FBE72A0495F2FEEA" box="[1070,1175,355,379]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="18" pageNumber="19" unit="mm" value="10.3">10.3 mm</quantity>
, 16.4×
<quantity id="805680E8FF8DFFD5FB2E2A049437FEEA" box="[1255,1362,355,379]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="19" unit="mm" value="12.7">12.7 mm</quantity>
), west coast of Lanyu Island,
<collectingCountry id="3FB96D9DFF8DFFD5FE182AE09343FE0E" box="[465,550,391,415]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
, calcareous rock terraces, spray zone, C.D. Schubart &amp; H.-C. Liu, 20
<date id="33100BCDFF8DFFD5FF082ACC9027FE52" box="[193,322,427,451]" pageId="18" pageNumber="19" value="1999-09-21">21.09.1999</date>
.—QM-W25699, male (16.2×
<quantity id="805680E8FF8DFFD5FD5E2ACC9260FE52" box="[663,773,427,451]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="18" pageNumber="19" unit="mm" value="12.4">12.4 mm</quantity>
), female (18.1×
<quantity id="805680E8FF8DFFD5FC022ACC955CFE52" box="[971,1081,427,451]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="18" pageNumber="19" unit="mm" value="13.4">13.4 mm</quantity>
), data as for ZRC 2000.1860. NMMBA-3504a,
<specimenCount id="51A8E684FF8DFFD5FEA12AA890A2FE76" box="[360,455,463,487]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(23.0×
<quantity id="805680E8FF8DFFD5FDEB2AA893EAFE76" box="[546,655,463,487]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="18" pageNumber="19" unit="mm" value="18.3">18.3 mm</quantity>
, 16.7×13.0 mm), Hsiang-Chiau-Wan, Kenting, Pingtung County,
<collectingCountry id="3FB96D9DFF8DFFD5FF5E2A959189FD9B" box="[151,236,498,522]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
, C.C. Li,
<date id="33100BCDFF8DFFD5FEA62A95908AFD9B" box="[367,495,498,522]" pageId="18" pageNumber="19" value="2012-08-23">23.08.2012</date>
.—ZRC 2012.0958,
<specimenCount id="51A8E684FF8DFFD5FD282A959231FD9B" box="[737,852,498,522]" pageId="18" pageNumber="19" type="female">2 females</specimenCount>
(21.0×15.7, 20.0×
<quantity id="805680E8FF8DFFD5FBFC2A9595C1FD9B" box="[1077,1188,498,522]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="18" pageNumber="19" unit="mm" value="14.9">14.9 mm</quantity>
), Siatanzai, Kenting, Pingtung County,
<collectingCountry id="3FB96D9DFF8DFFD5FEA7297190A6FDBF" box="[366,451,534,558]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
,
<geoCoordinate id="229A4BCAFF8DFFD5FE1D29719317FDBF" box="[468,626,534,558]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="21.932165">21°5555.8”N</geoCoordinate>
<geoCoordinate id="229A4BCAFF8DFFD5FDB62971924CFDBF" box="[639,809,534,558]" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="-120.74372">120°4437.4”S</geoCoordinate>
, J.-H. Lee &amp; W.-J. Wang, 6.08.2012.—NMNS, male (17.1×13.0 mm), female (20.5×
<quantity id="805680E8FF8DFFD5FDCF295D9316FDC3" box="[518,627,570,594]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="18" pageNumber="19" unit="mm" value="15.6">15.6 mm</quantity>
), Siatanzai, Kenting, Pingtung County,
<geoCoordinate id="229A4BCAFF8DFFD5FB8D295D9587FDC3" box="[1092,1250,570,594]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="21.926832">21°5536.6”N</geoCoordinate>
<geoCoordinate id="229A4BCAFF8DFFD5FB27295D94FDFDC3" box="[1262,1432,570,594]" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="-120.73939">120°4421.8”S</geoCoordinate>
,
<collectingCountry id="3FB96D9DFF8DFFD5FF5E29399189FDE7" box="[151,236,606,630]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
, J.-H. Lee &amp; W.-J. Wang,
<date id="33100BCDFF8DFFD5FDFF293993D0FDE7" box="[566,693,606,630]" pageId="18" pageNumber="19" value="2012-08-17">17.08.2012</date>
.—ZRC 2012.0959, male (18.1×
<quantity id="805680E8FF8DFFD5FBFD293995C0FDE7" box="[1076,1189,606,630]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="18" pageNumber="19" unit="mm" value="13.9">13.9 mm</quantity>
), Siatanzai, Kenting, Pingtung County,
<geoCoordinate id="229A4BCAFF8DFFD5FEAA29E69364FD08" box="[355,513,641,665]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="21.926832">21°5536.6”N</geoCoordinate>
<geoCoordinate id="229A4BCAFF8DFFD5FDC029E693D6FD08" box="[521,691,641,665]" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="-120.73939">120°4421.8”S</geoCoordinate>
,
<collectingCountry id="3FB96D9DFF8DFFD5FD0929E69270FD08" box="[704,789,641,665]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
, J.-H. Lee &amp; W.-J. Wang,
<date id="33100BCDFF8DFFD5FBF629E695DAFD08" box="[1087,1215,641,665]" pageId="18" pageNumber="19" value="2012-08-31">31.08.2012</date>
.—ZRC 2012.0960, male (17.8×
<quantity id="805680E8FF8DFFD5FEEB29C290EAFD2C" box="[290,399,677,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="18" pageNumber="19" unit="mm" value="13.9">13.9 mm</quantity>
), ovigerous female (18.8×
<quantity id="805680E8FF8DFFD5FD0929C2924BFD2C" box="[704,814,677,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="18" pageNumber="19" unit="mm" value="14.2">14.2 mm</quantity>
), Siatanzai, Kenting, Pingtung County,
<geoCoordinate id="229A4BCAFF8DFFD5FB3729C294F9FD2C" box="[1278,1436,677,701]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="21.926832">21°5536.6”N</geoCoordinate>
<geoCoordinate id="229A4BCAFF8DFFD5FF5E29AE9024FD70" box="[151,321,713,737]" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="-120.73939">120°4421.8”S</geoCoordinate>
,
<collectingCountry id="3FB96D9DFF8DFFD5FE8529AE90C4FD70" box="[332,417,713,737]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
, J.-H. Lee &amp; W.-J. Wang,
<date id="33100BCDFF8DFFD5FD0E29AE9222FD70" box="[711,839,713,737]" pageId="18" pageNumber="19" value="2012-08-31">31.08.2012</date>
.—ZRC 2012.0961, female (22.3×
<quantity id="805680E8FF8DFFD5FB0129AE9455FD70" box="[1224,1328,713,737]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="18" pageNumber="19" unit="mm" value="16.6">16.6 mm</quantity>
), Hsiang- Chiau-Wan, Kenting, Pingtung County,
<collectingCountry id="3FB96D9DFF8DFFD5FD9F298B93CEFC95" box="[598,683,748,772]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
, P.K.L. Ng &amp; C.-W. Lin,
<date id="33100BCDFF8DFFD5FC01298B955EFC95" box="[968,1083,748,772]" pageId="18" pageNumber="19" value="2012-01-10">1.10.2012</date>
.
</paragraph>
<paragraph id="47112D0DFF8DFFD5FF0E287793FCFA19" blockId="18.[151,1437,319,2023]" pageId="18" pageNumber="19">
NON-TYPE: RUMF-ZC-1351, male (16.1×
<quantity id="805680E8FF8DFFD5FD6528779271FCB9" box="[684,788,784,808]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="18" pageNumber="19" unit="mm" value="12.4">12.4 mm</quantity>
), Gazuda Beach, Hateruma I., Ryukyu Is.,
<collectingCountry id="3FB96D9DFF8DFFD5FB2328769449FCB9" box="[1258,1324,785,808]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita, 2.05.2009.—RUMF-ZC-1386,
<specimenCount id="51A8E684FF8DFFD5FE3D2853932EFCDD" box="[500,587,820,844]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(15.5×11.1, 13.2×
<quantity id="805680E8FF8DFFD5FCD228539212FCDD" box="[795,887,820,844]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="18" pageNumber="19" unit="mm" value="9.9">9.9 mm</quantity>
), beside Hateruma Port, Hateruma I., Ryukyu I.,
<collectingCountry id="3FB96D9DFF8DFFD5FF5E283F91BCFCFE" box="[151,217,856,879]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita,
<date id="33100BCDFF8DFFD5FE95283090BFFCFE" box="[348,474,855,879]" pageId="18" pageNumber="19" value="2009-04-19">19.04.2009</date>
.—RUMF-ZC-2075, male (16.9×13.0 mm), Dannu Beach, Yonagun Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FF3D281B9053FC02" box="[244,310,892,915]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita,
<date id="33100BCDFF8DFFD5FE66281C934AFC02" box="[431,559,891,915]" pageId="18" pageNumber="19" value="2007-11-15">15.11.2007</date>
.—RUMF-ZC-2076, male (19.7×
<quantity id="805680E8FF8DFFD5FC57281C9562FC02" box="[926,1031,891,915]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="18" pageNumber="19" unit="mm" value="14.6">14.6 mm</quantity>
), Honba Coast, Minamidaito Island,
<collectingCountry id="3FB96D9DFF8DFFD5FF5E28C791BCFC26" box="[151,217,928,951]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita, 2.09.2008.—RUMF-ZC-2077, male (18.5×
<quantity id="805680E8FF8DFFD5FCFA28F892FEFC26" box="[819,923,927,951]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="18" pageNumber="19" unit="mm" value="13.9">13.9 mm</quantity>
), Honba Coast, Minamidaito Island,
<collectingCountry id="3FB96D9DFF8DFFD5FAFE28C7941CFC26" box="[1335,1401,928,951]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita, 2.09.2008.—RUMF-ZC-2078, male (10.9×
<quantity id="805680E8FF8DFFD5FD0D28A4927BFC4A" box="[708,798,963,987]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="18" pageNumber="19" unit="mm" value="8.2">8.2 mm</quantity>
), Ubama Beach, Yonaguni Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FA9F28A394FDFC4A" box="[1366,1432,964,987]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita,
<date id="33100BCDFF8DFFD5FECE288090E1FC6E" box="[263,388,999,1023]" pageId="18" pageNumber="19" value="2007-11-16">16.11.2007</date>
.—RUMF-ZC-2079, male (17.2×
<quantity id="805680E8FF8DFFD5FD3028809207FC6E" box="[761,866,999,1023]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="18" pageNumber="19" unit="mm" value="12.8">12.8 mm</quantity>
), Dannu Beach, Yonaguni Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FF5E2F6C91BCFBB3" box="[151,217,1035,1058]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita,
<date id="33100BCDFF8DFFD5FE972F6D90BEFBB3" box="[350,475,1034,1058]" pageId="18" pageNumber="19" value="2007-11-15">15.11.2007</date>
.—RUMF-ZC-2080, male (12.5×
<quantity id="805680E8FF8DFFD5FC9D2F6D92D6FBB3" box="[852,947,1034,1058]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="18" pageNumber="19" unit="mm" value="9.5">9.5 mm</quantity>
), Dannu Beach, Yonaguni Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FF3C2F489052FBD7" box="[245,311,1071,1094]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita,
<date id="33100BCDFF8DFFD5FE7C2F499351FBD7" box="[437,564,1070,1094]" pageId="18" pageNumber="19" value="2007-11-15">15.11.2007</date>
.—RUMF-ZC-2082, female (16.9×
<quantity id="805680E8FF8DFFD5FC752F499540FBD7" box="[956,1061,1070,1094]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="19" unit="mm" value="12.9">12.9 mm</quantity>
), Shimoji Island, Miyako Group, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FE9A2F3490F0FBFB" box="[339,405,1107,1130]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita,
<date id="33100BCDFF8DFFD5FDD92F3593F5FBFB" box="[528,656,1106,1130]" pageId="18" pageNumber="19" value="2012-09-23">23.09.2012</date>
.—RUMF-ZC-2083, female (15.6×
<quantity id="805680E8FF8DFFD5FBD12F35951AFBFB" box="[1048,1151,1106,1130]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="18" pageNumber="19" unit="mm" value="12.1">12.1 mm</quantity>
), Shimoji Island, Miyako Group, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FE7F2F10909DFB1F" box="[438,504,1143,1166]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita,
<date id="33100BCDFF8DFFD5FDB62F11939BFB1F" box="[639,766,1142,1166]" pageId="18" pageNumber="19" value="2012-09-23">23.09.2012</date>
.—RUMF-ZC-2089, female (18.0×
<quantity id="805680E8FF8DFFD5FB592F11959EFB1F" box="[1168,1275,1142,1166]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="18" pageNumber="19" unit="mm" value="13.1">13.1 mm</quantity>
), Giza Banta, Okinawa Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FDD12FFD933FFB20" box="[536,602,1178,1201]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, T. Maenosono, 2007.—RUMF-ZC-2090,
<specimenCount id="51A8E684FF8DFFD5FB892FFE95FCFB20" box="[1088,1177,1177,1201]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(16.0×12.2, 14.1×11.0 mm), female (14.8×
<quantity id="805680E8FF8DFFD5FEB72FDA908EFB44" box="[382,491,1213,1237]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="18" pageNumber="19" unit="mm" value="11.6">11.6 mm</quantity>
), Kori Island, off Nakijin Village of Okinawa Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FAFA2FD99410FB44" box="[1331,1397,1214,1237]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, T. Maenosono, 9.06.2010.—RUMF-ZC-2104,
<specimenCount id="51A8E684FF8DFFD5FD4A2F8693BFFB68" box="[643,730,1249,1273]" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(17.5×13.7, 17.4×
<quantity id="805680E8FF8DFFD5FC632F869577FB68" box="[938,1042,1249,1273]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="19" unit="mm" value="12.9">12.9 mm</quantity>
), female (15.7×
<quantity id="805680E8FF8DFFD5FB0A2F86944FFB68" box="[1219,1322,1249,1273]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="18" pageNumber="19" unit="mm" value="12.1">12.1 mm</quantity>
), Muigah, Miyako Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FDDE2E62933CFA8D" box="[535,601,1285,1308]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita, 4.05.2005.—ZRC,
<specimenCount id="51A8E684FF8DFFD5FC772E63957CFA8D" box="[958,1049,1284,1308]" pageId="18" pageNumber="19" type="male">6 males</specimenCount>
(largest 18.9×
<quantity id="805680E8FF8DFFD5FB0C2E639455FA8D" box="[1221,1328,1284,1308]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="18" pageNumber="19" unit="mm" value="14.5">14.5 mm</quantity>
, smallest 14.3×
<quantity id="805680E8FF8DFFD5FF1F2E4F905BFAD1" box="[214,318,1320,1344]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="18" pageNumber="19" unit="mm" value="10.8">10.8 mm</quantity>
),
<specimenCount id="51A8E684FF8DFFD5FE9B2E4F90DBFAD1" box="[338,446,1320,1344]" pageId="18" pageNumber="19" type="female">5 females</specimenCount>
(largest 15.2×
<quantity id="805680E8FF8DFFD5FDA92E4F93A0FAD1" box="[608,709,1320,1344]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="18" pageNumber="19" unit="mm" value="11.3">11.3 mm</quantity>
, smallest 8.3×
<quantity id="805680E8FF8DFFD5FCAF2E4F92DAFAD1" box="[870,959,1320,1344]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="18" pageNumber="19" unit="mm" value="7.1">7.1 mm</quantity>
), Muigah, Miyako Island, Ryukyu Islands,
<collectingCountry id="3FB96D9DFF8DFFD5FF5E2E2A91BCFAF5" box="[151,217,1357,1380]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, Y. Fujita, 4.07.2005.—RUMF-ZC-819, female (15.7×12.0 mm), Giza Banda, Yaese Town, Okinawa I., Ryukyu Is.,
<collectingCountry id="3FB96D9DFF8DFFD5FED42E16903AFA19" box="[285,351,1393,1416]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
, T. Maenosono,
<date id="33100BCDFF8DFFD5FDDC2E1793F3FA19" box="[533,662,1392,1416]" pageId="18" pageNumber="19" value="2007-05-22">22.05.2007</date>
.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8DFFD5FF0E2EF4943FF97C" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="47112D0DFF8DFFD5FF0E2EF4943FF97C" blockId="18.[151,1437,319,2023]" pageId="18" pageNumber="19">
<emphasis id="75DAF11FFF8DFFD5FF0E2EF4905FFA3D" bold="true" box="[199,314,1427,1452]" pageId="18" pageNumber="19">Diagnosis</emphasis>
. Carapace transversely subovate, c. 1.3 times broader than long; dorsal carapace, lateral branchial regions markedly swollen; exorbital tooth moderately oblique, forming slight angle posteriorly marking widest point of carapace, most prominent in larger specimens; front c. 0.6 times carapace width, margin slightly to broadly concave in dorsal view, appears smooth but microscopically granular, with pair of prominent lateral swellings behind margin; supraorbital margin entire, semicircular; dorsal surface of cheliped carpus conspicuously granular; walking legs relatively short; P4 merus c. 2.3 times longer than wide; P5 merus similar, 2.22.3 times longer; propodus P4 propodus c. 2.8 times longer than wide; P5 propodus 2.4 times; male abdomen (
<figureCitation id="DF953188FF8DFFD5FB642D0D9598F913" box="[1197,1277,1642,1666]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
F) moderately broad; somite 6 with divergent lateral margins, margins relatively evenly convex; somite 3 width 2.8 times basal width of telson; G1 (
<figureCitation id="DF953188FF8DFFD5FE482DD690B5F958" box="[385,464,1713,1737]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
GJ) relatively slender, weakly tapering to broadly convex subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process long, with dorsal margin concave, apex slightly flanged.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8DFFD4FF0E2D9F946EF8B7" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="47112D0DFF8DFFD4FF0E2D9F93F0FE5F" blockId="18.[151,1437,319,2023]" lastBlockId="19.[151,1437,151,1973]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="75DAF11FFF8DFFD5FF0E2D9F9034F880" bold="true" box="[199,337,1784,1809]" pageId="18" pageNumber="19">Description</emphasis>
. Carapace transversely subovate, c. 1.3 times (range 1.261.35) broader than long; surface bare, lacking setal tufts; covered with fine, but distinct low granules becoming arranged into short striae posteriorly; mesogastric regions well defined; lateral carapace surface generally without obvious discrete oblique striae, except for strong, concave epibranchial sulcus beginning just behind exorbital tooth. Dorsal carapace, lateral branchial regions markedly swollen. Exorbital tooth triangular, pointed, outer margin long, moderately oblique, forming slight angle posteriorly marking widest point of carapace, most prominent in larger specimens; margin constricted behind, but no trace of second anterolateral tooth. Postfrontal lobes strongly demarcated, separated by broad grooves, median lobes similar in width to lateral lobes. Front c. 0.6 times carapace width (
<quantity id="805680E8FF8CFFD4FB032BF0947BFF3E" box="[1226,1310,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" pageId="19" pageNumber="20" unit="in" value="0.66">0.66 in</quantity>
<typeStatus id="981593AFFF8CFFD4FAE32BF094F5FF3E" box="[1322,1424,151,175]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
), markedly deflexed, margin slightly to broadly concave in dorsal view; laterally triangular, bluntly pointed; frontal margin emarginate, microscopically granular (
<figureCitation id="DF953188FF8CFFD4FD732BB89266FF66" box="[698,771,223,247]" captionStart="FIGURE 8" captionStartId="11.[151,250,1137,1160]" captionTargetBox="[151,1436,409,1116]" captionTargetId="figure@11.[151,1436,409,1116]" captionTargetPageId="11" captionText="FIGURE 8. Frontal margins (ventral views). A, Chiromantes obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; B, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; C, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; D, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283289/files/figure.png" pageId="19" pageNumber="20">Fig. 8</figureCitation>
C); surface somewhat concave behind but with pair of prominent lateral swellings behind frontal margin. Supraorbital margin entire, semicircular; secondary rim formed either side making a smooth, broadly triangular, deflexed central plate behind ocular peduncle. Eye not extending beyond exorbital tooth. Frontal plate protruding as a shelf; medial septum broad, largely covering anterior half of anterior half of epistome; antennae, antennules much reduced in size, lodged under overhanging front; basal antennular segment not much swollen; antennal, antennular basal segments adjacent, not separated by septum. Antennal flagellum very short, entering orbit.
</paragraph>
<paragraph id="47112D0DFF8CFFD4FF0E2ABE95FDFDCC" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">Third maxillipeds with ischium, merus subequal in length; ischium with shallow longitudinal, curved, median sulcus. Inner margin of merus, ischium with long setae, proximal outer margin of ischium, base of exopod with dense setae; outer margin of merus with short scattered setae only. Exopod slender, hidden behind ishium, merus except near base, tip reaching half length of outer margin of merus, flagellum long, slender.</paragraph>
<paragraph id="47112D0DFF8CFFD4FF0E290E90DFFC78" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">
Chelipeds subequal, robust. Merus trihedral; posterior border broadly convex emarginate, minutely granular, without indication of subdistal spine or lobe; inner anterior border minutely granular, broadly triangular; outer surface broadly convex, with fine transverse striae but appearing almost smooth. Carpus subquadrilateral, inner angle moderately produced, apically granular; outer margin, dorsal surface conspicuously granular. Palm dorsal surface slightly striated along superior margin, with scattering of moderate sized granules, otherwise smooth, punctate. Palm outer surface broadly rounded, smooth; no indication of subventral longitudinal ridge. Palm inner face of males smooth except for curved projecting crest of large tubercles behind gape (
<figureCitation id="DF953188FF8CFFD4FBAE282695CFFCC8" box="[1127,1194,833,857]" captionStart="FIGURE 6" captionStartId="9.[151,250,1537,1560]" captionTargetBox="[151,1435,193,1515]" captionTargetId="figure@9.[151,1436,193,1516]" captionTargetPageId="9" captionText="FIGURE 6. Inner views of chelae. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283287/files/figure.png" pageId="19" pageNumber="20">Fig. 6</figureCitation>
D). Cutting margin of fixed finger with 3 or 4 blunt teeth, proximal 3 similar in size, distalmost larger, rounded, conical, placed medially; dactylus with 2 teeth proximally, distalmost large, third much smaller blunt tooth slightly distal of medial tooth of ventral finger. Dorsal surface of dactylus smooth. Fingers with chitinous tips; adult males with narrow but distinct gape when fingers closed.
</paragraph>
<paragraph id="47112D0DFF8CFFD4FF0E28929458FB50" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">Walking legs relatively short, broad, flattened; second, third pairs sub-equal, longer than other walking legs. Without setae except for scattering of short dark bristles on dactyli, ventral face of propodi. Dorsal margins of meri with blunt subdistal shoulder, but otherwise unarmed; outer surfaces of meri with minutely granular transverse striae. P4 merus c. 2.3 times longer than wide; P5 merus similar, 2.22.3 times longer. Carpi with 2 accessory carinae on outer surface but not strongly marked. P4 propodus c. 2.8 times longer than wide; P5 propodus 2.4 times longer than wide. Dactyli 0.8 times length of propodi, slightly recurved, terminating in acute chitinous tip.</paragraph>
<paragraph id="47112D0DFF8CFFD4FF0E2FAA92CDFA08" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">
Thoracic sternites (
<figureCitation id="DF953188FF8CFFD4FE6B2FAA9083FB74" box="[418,486,1229,1253]" captionStart="FIGURE 9" captionStartId="12.[151,250,1576,1599]" captionTargetBox="[151,1435,325,1554]" captionTargetId="figure@12.[151,1436,325,1555]" captionTargetPageId="12" captionText="FIGURE 9. Ventral views of thoracic sternum and anterior parts of male abdomens. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283290/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
D) smooth, mostly bare of setae except for few scattered bristles; abdominal cavity reaching to thick transverse setal fringe at junction of sternites 3, 4. Male abdomen (
<figureCitation id="DF953188FF8CFFD4FB922F9695C9FA98" box="[1115,1196,1265,1289]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="19" pageNumber="20">Fig. 13</figureCitation>
F) moderately broad; telson broadly rounded apically, 1.1 times longer than wide, only slightly longer than somite 6; somite 6 c. 1.8 times wider than long, lateral margins divergent with margins relatively evenly convex; somites 35 trapezoidal, lateral margins of somites 4, 5 relatively straight, lateral margins of somite 3 convex; somite 3 width 2.8 times basal width of telson; somites 1, 2 transversely, longitudinally narrow.
</paragraph>
<paragraph id="47112D0DFF8CFFD4FF0E2EC29345F997" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">
G1 (
<figureCitation id="DF953188FF8CFFD4FF342EC2902AFA2C" box="[253,335,1445,1469]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="19" pageNumber="20">Fig. 13</figureCitation>
GJ) relatively slender, almost straight, weakly tapering to broadly convex subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process long, with dorsal margin concave, apex slightly flanged. G2 short, as for the genus.
</paragraph>
<paragraph id="47112D0DFF8CFFD4FF0E2D759397F9DC" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">
<emphasis id="75DAF11FFF8CFFD4FF0E2D75904CF9BB" bold="true" box="[199,297,1554,1578]" pageId="19" pageNumber="20">Females</emphasis>
. Chelipeds slightly smaller, less robust, lacking raised granular row on inner face of chela. Vulval morphology without clear species-specific characters.
</paragraph>
<paragraph id="47112D0DFF8CFFD4FF0E2D3E946EF8B7" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">
<emphasis id="75DAF11FFF8CFFD4FF0E2D3E9002F9E3" bold="true" box="[199,359,1625,1650]" pageId="19" pageNumber="20">Colour in life</emphasis>
. (
<figureCitation id="DF953188FF8CFFD4FEB72D3E90A5F9E0" box="[382,448,1625,1649]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="19" pageNumber="20">Fig. 3</figureCitation>
DH). Carapace background colour of adult males varying from pale brown to a darker grey-green; females and smaller males mottled greyish white, sometimes yellowish. Upper surface of carapace and legs with some course blotches and streaks; a distinctive pattern of large dirty yellow to green blotches across anterior part of carapace. Chelae of adult males reddish-pink to purplish red overall, especially on dorsal surfaces and dactylus; smaller males and females with palm whitish to whitish-pink, dactylus always clearly tinged with red to pink in part or whole. Ocular peduncles similar to carapace; corneas light green to yellowish green.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8CFFD3FF0E2C559429FDAD" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
<paragraph id="47112D0DFF8CFFD4FF0E2C55924BF824" blockId="19.[151,1437,151,1973]" pageId="19" pageNumber="20">
<emphasis id="75DAF11FFF8CFFD4FF0E2C559051F8DB" bold="true" box="[199,308,1842,1866]" pageId="19" pageNumber="20">Remarks</emphasis>
. Easily separated from its sympatric Taiwanese congener,
<taxonomicName id="80AE568EFF8CFFD4FC072C5695B9F8D8" box="[974,1244,1841,1865]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="75DAF11FFF8CFFD4FC072C5695B9F8D8" box="[974,1244,1841,1865]" italics="true" pageId="19" pageNumber="20">Chiromantes eurymerus</emphasis>
</taxonomicName>
, by its relatively narrower legs, but also by its more slender G1 with its markedly elongated tip, and other differences discussed further under that species account. The numerous features that distinguish this species from all other presently known species in this sibling complex are given in
<tableCitation id="0A2C18B6FF8CFFD4FD1D2CFA924DF824" box="[724,808,1949,1973]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[159,1428,329,1954]" captionTargetPageId="5" captionText="TABLE 1. Summary of major differences between species of the Chiromantes obtusifrons complex." httpUri="http://table.plazi.org/id/13D17D85FF9AFFC2FF5E2BF095A0FF3E" pageId="19" pageNumber="20" tableUuid="13D17D85FF9AFFC2FF5E2BF095A0FF3E">Table 1</tableCitation>
.
</paragraph>
<paragraph id="47112D0DFF8BFFD3FF0E2BF09429FDAD" blockId="20.[151,1436,151,788]" pageId="20" pageNumber="21">
Specimens from the Ryukyu Is. seem to show some minor variability in the proportions of the leg segments, with the meri of some individuals appearing a little broader than typical, and almost approaching the
<taxonomicName id="80AE568EFF8BFFD3FACC2BDA94F9FF45" box="[1285,1436,189,212]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="eurymerus">
<emphasis id="75DAF11FFF8BFFD3FACC2BDA94F9FF45" box="[1285,1436,189,212]" italics="true" pageId="20" pageNumber="21">C. eurymerus</emphasis>
</taxonomicName>
condition. However, in all other characters, including the form of the thoracic sternum and G1, they appear to be typical
<taxonomicName id="80AE568EFF8BFFD3FF242A6290E9FE8D" box="[237,396,260,284]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="75DAF11FFF8BFFD3FF242A6290E9FE8D" box="[237,396,260,284]" italics="true" pageId="20" pageNumber="21">C. leptomerus</emphasis>
</taxonomicName>
. Their live colours are also similar. Perhaps this could indicate some incipient population separation taking place between
<collectingCountry id="3FB96D9DFF8BFFD3FDC42A40933AFEAE" box="[525,607,295,319]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
and
<collectingCountry id="3FB96D9DFF8BFFD3FD532A4F93B8FEAE" box="[666,733,296,319]" name="Japan" pageId="20" pageNumber="21">Japan</collectingCountry>
, as has been observed for the mitten crab
<taxonomicName id="80AE568EFF8BFFD3FB0C2A4F907AFEF5" class="Malacostraca" family="Varunidae" genus="Eriocheir" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="75DAF11FFF8BFFD3FB0C2A4F94F9FED1" box="[1221,1436,296,320]" italics="true" pageId="20" pageNumber="21">Eriocheir japonica</emphasis>
(Varunidae)
</taxonomicName>
, with crabs from Okinawa proving to be genetically distinct from Taiwanese and Chinese populations (Xu
<emphasis id="75DAF11FFF8BFFD3FF042A16906FFE19" box="[205,266,368,392]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2009). Twin species-pairs on either side of the deep-water strait between
<collectingCountry id="3FB96D9DFF8BFFD3FBBE2A0895ACFE16" box="[1143,1225,367,391]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
and the Yaeyama Islands has now been observed for several crabs including
<taxonomicName id="80AE568EFF8BFFD3FCB02AF395A1FE3D" box="[889,1220,404,428]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ceratophora">
<emphasis id="75DAF11FFF8BFFD3FCB02AF395A1FE3D" box="[889,1220,404,428]" italics="true" pageId="20" pageNumber="21">Tmethypocoelis ceratophora</emphasis>
</taxonomicName>
and
<taxonomicName id="80AE568EFF8BFFD3FAC52AF2907FFE5E" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="choreutes">
<emphasis id="75DAF11FFF8BFFD3FAC52AF294F9FE3D" box="[1292,1436,404,428]" italics="true" pageId="20" pageNumber="21">T. choreutes</emphasis>
(Dotillidae)
</taxonomicName>
(Davie &amp; Kosuge 1995), and
<taxonomicName id="80AE568EFF8BFFD3FDA52ADE9256FE41" box="[620,819,441,464]" class="Malacostraca" family="Mictyridae" genus="Mictyris" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="75DAF11FFF8BFFD3FDA52ADE9256FE41" box="[620,819,441,464]" italics="true" pageId="20" pageNumber="21">Mictyris guinotae</emphasis>
</taxonomicName>
and
<taxonomicName id="80AE568EFF8BFFD3FCA32ADE95DFFE5E" box="[874,1210,439,464]" class="Malacostraca" family="Mictyridae" genus="Mictyris" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brevidactylus">
<emphasis id="75DAF11FFF8BFFD3FCA32ADE9540FE41" box="[874,1061,440,464]" italics="true" pageId="20" pageNumber="21">M. brevidactylus</emphasis>
(Mictyridae)
</taxonomicName>
(Davie
<emphasis id="75DAF11FFF8BFFD3FADD2ADE9428FE41" box="[1300,1357,440,464]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2010). While we cannot discern any clear morphological grounds for separating the Japanese and Taiwanese populations of
<taxonomicName id="80AE568EFF8BFFD3FF7529679004FD89" box="[188,353,512,536]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="75DAF11FFF8BFFD3FF7529679004FD89" box="[188,353,512,536]" italics="true" pageId="20" pageNumber="21">C. leptomerus</emphasis>
</taxonomicName>
for the moment, it will be interesting if future genetic studies may show some level of evolutionary divergence reflecting that observed for other populations and species between these two areas.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8BFFD3FF0E292F9319FD36" pageId="20" pageNumber="21" type="etymology">
<paragraph id="47112D0DFF8BFFD3FF0E292F9319FD36" blockId="20.[151,1436,151,788]" pageId="20" pageNumber="21">
<emphasis id="75DAF11FFF8BFFD3FF0E292F9023FDF1" bold="true" box="[199,326,584,608]" pageId="20" pageNumber="21">Etymology</emphasis>
. Derived from the Greek
<emphasis id="75DAF11FFF8BFFD3FDB4292F93DAFDF1" box="[637,703,584,608]" italics="true" pageId="20" pageNumber="21">leptos</emphasis>
meaning “thin” and
<emphasis id="75DAF11FFF8BFFD3FC72292E929BFDF1" box="[955,1022,585,608]" italics="true" pageId="20" pageNumber="21">meros</emphasis>
meaning “thigh”, and refers to the relatively more slender legs of this species compared to the other sympatric Taiwanese species,
<taxonomicName id="80AE568EFF8BFFD3FB13290A9417FD15" box="[1242,1394,621,644]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF8BFFD3FB13290A9417FD15" box="[1242,1394,621,644]" italics="true" pageId="20" pageNumber="21">C. eurymerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF8BFFD3FAB3290B91ADFD39" pageId="20" pageNumber="21" rank="species">
<emphasis id="75DAF11FFF8BFFD3FAB3290B91ADFD39" bold="true" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
It is used here as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8BFFD3FF0E29D49539FC85" pageId="20" pageNumber="21" type="distribution">
<paragraph id="47112D0DFF8BFFD3FF0E29D49536FD5D" blockId="20.[151,1436,151,788]" box="[199,1107,691,716]" pageId="20" pageNumber="21">
<emphasis id="75DAF11FFF8BFFD3FF0E29D4903CFD5D" bold="true" box="[199,345,691,716]" pageId="20" pageNumber="21">Distribution</emphasis>
. Only known from
<collectingCountry id="3FB96D9DFF8BFFD3FDFE29D393ECFD5D" box="[567,649,692,716]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
and the Ryukyu Islands, southern
<collectingCountry id="3FB96D9DFF8BFFD3FBC229D2952BFD5D" box="[1035,1102,693,716]" name="Japan" pageId="20" pageNumber="21">Japan</collectingCountry>
.
</paragraph>
<paragraph id="47112D0DFF8BFFD3FF0E29BF9539FC85" blockId="20.[151,1436,151,788]" pageId="20" pageNumber="21">
<emphasis id="75DAF11FFF8BFFD3FF0E29BF9041FD61" bold="true" box="[199,292,728,752]" pageId="20" pageNumber="21">Ecology</emphasis>
. Supratidal; calcareous rock terraces, spray zone. Found comingled with
<taxonomicName id="80AE568EFF8BFFD3FB9429BF9591FD61" box="[1117,1268,728,752]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF8BFFD3FB9429BF9591FD61" box="[1117,1268,728,752]" italics="true" pageId="20" pageNumber="21">C. eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF8BFFD3FB3529BF9430FD61" bold="true" box="[1276,1365,728,752]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="6EE94C64FF8BFFD3FB3529BF9430FD61" box="[1276,1365,728,752]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the same shore habitat; the nature and extent of niche separation has not been determined.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="828FAA85227999A166B7FC39820DBD0A">A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island (Indian Ocean), Guam and Taiwan</mods:title>
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<mods:title id="79F2C565E5A3F5449E107297D4DF5142">Zootaxa</mods:title>
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<mods:part id="A0C67D6AE8A33C38F7308E300CDD9E44">
<mods:date id="C393C671F92F6A6DD9512D5E467CF443">2013</mods:date>
<mods:detail id="547CB16D3D8FEBC6BE19A5F0BFAB966F" type="volume">
<mods:number id="6B43A327E1489E38FE1A11D62C4ABB72">3609</mods:number>
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<mods:start id="BF7C36A08E3D787D56AFB5E550A06F67">1</mods:start>
<mods:end id="4B4AC64220090D76F4AC5130CE8A543E">25</mods:end>
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<mods:identifier id="94587B951B1159BC4ED3D9A42F76D6F2" type="DOI">10.11646/zootaxa.3609.1.1</mods:identifier>
<mods:identifier id="AB8F82CB59888D1453DE83073FDAABCF" type="GBIF-Dataset">de6f3817-2480-4937-9530-b8f5a03a2068</mods:identifier>
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<mods:identifier id="6A07C475E06834A1389181DA2A19A690" type="Zenodo-Dep">283281</mods:identifier>
<mods:identifier id="90019F8DAC03793320614A653C4D2DEE" type="ZooBank">0C5E3F32-FB0B-4AF6-AD4F-9D96A1BA670B</mods:identifier>
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<treatment id="CF079C1BFF92FFD7FF5E2D3D90E4F870" ID-DOI="http://doi.org/10.5281/zenodo.5618388" ID-GBIF-Taxon="127663035" ID-Zenodo-Dep="5618388" LSID="urn:lsid:plazi:treatment:CF079C1BFF92FFD7FF5E2D3D90E4F870" httpUri="http://treatment.plazi.org/id/CF079C1BFF92FFD7FF5E2D3D90E4F870" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<subSubSection id="0FB47E86FF92FFCAFF5E2D3D9283F94A" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="47112D0DFF92FFCAFF5E2D3D90B3F9E5" blockId="13.[151,665,1626,1685]" box="[151,470,1626,1652]" pageId="13" pageNumber="14">
<heading id="1C599A61FF92FFCAFF5E2D3D90B3F9E5" bold="true" box="[151,470,1626,1652]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="75DAF11FFF92FFCAFF5E2D3D90B3F9E5" bold="true" box="[151,470,1626,1652]" pageId="13" pageNumber="14">
<taxonomicName id="80AE568EFF92FFCAFF5E2D3D9017F9E5" ID-CoL="7TVP9" box="[151,370,1626,1652]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF92FFCAFF5E2D3D9017F9E5" bold="true" box="[151,370,1626,1652]" italics="true" pageId="13" pageNumber="14">Chiromantes silus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF92FFCAFEB02D3D90B3F9E5" box="[377,470,1626,1652]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="47112D0DFF92FFCAFF5E2D1A93FCF904" blockId="13.[151,665,1626,1685]" box="[151,665,1661,1685]" pageId="13" pageNumber="14">
(
<figureCitation id="DF953188FF92FFCAFF562D1A9186F904" box="[159,227,1661,1685]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="13" pageNumber="14">Figs 3</figureCitation>
AC, 4B, 5C, 6C, 7C, 8B, 9C, 13AE)
</paragraph>
<paragraph id="47112D0DFF92FFCAFF5E2DA29283F94A" blockId="13.[151,998,1733,1755]" box="[151,998,1733,1755]" pageId="13" pageNumber="14">
<taxonomicName id="80AE568EFF92FFCAFF5E2DA290F5F94A" box="[151,400,1733,1755]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF92FFCAFF5E2DA290F5F94A" box="[151,400,1733,1755]" italics="true" pageId="13" pageNumber="14">Chiromantes obtusifrons</emphasis>
</taxonomicName>
—Paulay
<emphasis id="75DAF11FFF92FFCAFE3D2DA1934DF94A" box="[500,552,1733,1755]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2003: 508.—Paulay &amp; Starmer 2011: 2, 11.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF92FFC9FF5E2C60902AFF66" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="47112D0DFF92FFC9FF5E2C60902AFF66" blockId="13.[151,1437,1799,2005]" lastBlockId="14.[151,1436,151,247]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="75DAF11FFF92FFCAFF5E2C6090E7F8B1" bold="true" box="[151,386,1799,1824]" pageId="13" pageNumber="14">Material Examined</emphasis>
.
<typeStatus id="981593AFFF92FFCAFE5D2C6F9349F8B1" box="[404,556,1800,1824]" pageId="13" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
: ZRC 2012.0787, male (16.3×
<quantity id="805680E8FF92FFCAFC452C6F929DF8B1" box="[908,1016,1800,1824]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="13" pageNumber="14" unit="mm" value="12.8">12.8 mm</quantity>
), Tanguisson,
<collectingCountry id="3FB96D9DFF92FFCAFB562C6F9582F8B1" box="[1183,1255,1800,1824]" name="Guam" pageId="13" pageNumber="14">Guam</collectingCountry>
,
<quantity id="805680E8FF92FFCAFB3C2C6F943EF8B1" box="[1269,1371,1800,1824]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="14" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">1020 m</quantity>
from shore,
<quantity id="805680E8FF92FFCAFF2A2C4B904EF8D5" box="[227,299,1836,1860]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="13" pageNumber="14" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">58 m</quantity>
elevation, under limestone rocks, G. Paulay, 0
<date id="33100BCDFF92FFCAFCF12C4B92CEF8D5" box="[824,939,1836,1860]" pageId="13" pageNumber="14" value="1997-05-10">5.10.1997</date>
.
<typeStatus id="981593AFFF92FFCAFC7E2C4B9533F8D5" box="[951,1110,1836,1860]" pageId="13" pageNumber="14" type="paratype">PARATYPES</typeStatus>
: ZRC 2012.0788, data as for
<typeStatus id="981593AFFF92FFCAFF5E2C379199F8F9" box="[151,252,1872,1896]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, male (15.1×
<quantity id="805680E8FF92FFCAFE472C379092F8F9" box="[398,503,1872,1896]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="13" pageNumber="14" unit="mm" value="11.3">11.3 mm</quantity>
), female (14.9×
<quantity id="805680E8FF92FFCAFD6E2C37926AF8F9" box="[679,783,1872,1896]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="13" pageNumber="14" unit="mm" value="11.2">11.2 mm</quantity>
).—UFUF-122,
<specimenCount id="51A8E684FF92FFCAFC0B2C37957FF8F9" box="[962,1050,1872,1896]" pageId="13" pageNumber="14" type="male">3 males</specimenCount>
(16.3×13.2, 15.9×11.9, 13.6×
<quantity id="805680E8FF92FFCAFAA22C3791A2F81D" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="13" pageNumber="14" unit="mm" value="10.5">10.5 mm</quantity>
), Tanguisson,
<quantity id="805680E8FF92FFCAFEA22C1390B5F81D" box="[363,464,1908,1932]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="14" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">1020 m</quantity>
from shore,
<quantity id="805680E8FF92FFCAFDAF2C1393F6F81D" box="[614,659,1908,1932]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1.0">1 m</quantity>
from forest, under rocks, G. Paulay,
<date id="33100BCDFF92FFCAFBF42C1395D8F81D" box="[1085,1213,1908,1932]" pageId="13" pageNumber="14" value="1997-05-10">05.10.1997</date>
.—ZRC 2001.0742, male (15.7×
<quantity id="805680E8FF92FFCAFED42CFF90E3F821" box="[285,390,1944,1968]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="13" pageNumber="14" unit="mm" value="11.7">11.7 mm</quantity>
), Haputo Beach, P.K.L. Ng, 3.08.2001.—ZRC 2001.0743, ovigerous female (15.2×
<quantity id="805680E8FF92FFCAFAE72CFF94F1F821" box="[1326,1428,1944,1968]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="13" pageNumber="14" unit="mm" value="11.6">11.6 mm</quantity>
) (with zoea 1), Haputo Beach, P.K.L. Ng,
<date id="33100BCDFF92FFCAFD442CDB9269F845" box="[653,780,1980,2004]" pageId="13" pageNumber="14" value="2001-04-08">04.08.2001</date>
.—ZRC 2000.0573, female (15.2×
<quantity id="805680E8FF92FFCAFB572CDB9469F845" box="[1182,1292,1980,2004]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="13" pageNumber="14" unit="mm" value="11.2">11.2 mm</quantity>
), Pago Bay, outside of University of
<collectingCountry id="3FB96D9DFF91FFC9FE6F2BF0908EFF3E" box="[422,491,151,175]" name="Guam" pageId="14" pageNumber="15">Guam</collectingCountry>
Marine Laboratory, P.K.L. Ng &amp; C.H. Wang, 15
<date id="33100BCDFF91FFC9FBDA2BF095F0FF3E" box="[1043,1173,151,175]" pageId="14" pageNumber="15" value="2000-04-18">18.04.2000</date>
.—UFUF-90,
<specimenCount id="51A8E684FF91FFC9FAF92BF094F9FF3E" box="[1328,1436,151,175]" pageId="14" pageNumber="15" type="female">2 females</specimenCount>
(19.1×13.6, 19.0×
<quantity id="805680E8FF91FFC9FEA82BDB90ACFF45" box="[353,457,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="14" pageNumber="15" unit="mm" value="13.7">13.7 mm</quantity>
), Tagachang, near shore, karst rubble, under
<quantity id="805680E8FF91FFC9FC772BDB9292FF45" box="[958,1015,188,212]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="m" value="15.0">15 m</quantity>
overhang, G. Paulay,
<date id="33100BCDFF91FFC9FB252BDB9408FF45" box="[1260,1389,188,212]" pageId="14" pageNumber="15" value="1996-09-29">29.09.1996</date>
. All localities
<collectingCountry id="3FB96D9DFF91FFC9FECA2BB8902EFF66" box="[259,331,223,247]" name="Guam" pageId="14" pageNumber="15">Guam</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="13D17D85FF91FFC9FF5E2C319341F843" httpUri="https://zenodo.org/record/283292/files/figure.png" pageId="14" pageNumber="15" targetBox="[161,1420,309,1846]" targetPageId="14">
<paragraph id="47112D0DFF91FFC9FF5E2C319341F843" blockId="14.[151,1437,1878,2002]" pageId="14" pageNumber="15">
<emphasis id="75DAF11FFF91FFC9FF5E2C319046F8FA" bold="true" box="[151,291,1878,1901]" pageId="14" pageNumber="15">FIGURE 11.</emphasis>
<taxonomicName id="80AE568EFF91FFC9FEE72C3193A1F8FD" authority="(Dana, 1851)" authorityName="Dana" authorityYear="1851" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[302,708,1878,1900]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF91FFC9FEE72C319349F8FD" box="[302,556,1878,1900]" italics="true" pageId="14" pageNumber="15">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, ZRC 2002.0220, male (19.7×14.7 mm), Hawaiian Is.. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 36, telson); D, ventral view of left G1; E, dorsal view of left G1; F, ventral view of distal part of left G1; G, dorsal view of distal part of left G1. All structures denuded. Scales: AE = 1.0 mm; F, G = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="0FB47E86FF90FFC8FF0E2BF09446FE65" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="47112D0DFF90FFC8FF0E2BF09446FE65" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
<emphasis id="75DAF11FFF90FFC8FF0E2BF0905FFF21" bold="true" box="[199,314,151,176]" pageId="15" pageNumber="16">Diagnosis</emphasis>
. Carapace transversely subovate, c. 1.3 times broader than long; dorsal carapace, lateral branchial regions markedly swollen; exorbital tooth weakly oblique, forming rounded obtuse angle posteriorly marking widest point of carapace, especially in larger specimens; front c. 0.65 times carapace width, margin straight or slightly concave in dorsal view, appearing smooth but microscopically granular, with pair of low lateral swellings behind margin; supraorbital margin entire, semicircular; dorsal surface of cheliped carpus conspicuously granular; walking legs relatively short; P4 merus c.2.3 times longer than wide; P5 merus similar, 2.4 times longer; P4 propodus c.2.6 times longer than wide; P5 merus c. 2.3 times; male abdomen (
<figureCitation id="DF953188FF90FFC8FBC02A08950AFE16" box="[1033,1135,367,391]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="15" pageNumber="16">Fig. 13A</figureCitation>
) moderately broad, somite 6 with lateral margins broadly divergent with margins relatively evenly convex, somite 3 width 3.2 times basal width of telson; G1 (
<figureCitation id="DF953188FF90FFC8FE492AD09361FE5E" box="[384,516,439,463]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="15" pageNumber="16">Fig. 13BE</figureCitation>
) relatively slender, weakly tapering to broadly convex subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process long, with dorsal margin concave, apically pointed.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF90FFD7FF0E2A9895E9F9E8" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="47112D0DFF90FFC8FF0E2A9891B6FBC9" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
<emphasis id="75DAF11FFF90FFC8FF0E2A989034FD89" bold="true" box="[199,337,511,536]" pageId="15" pageNumber="16">Description</emphasis>
. Carapace transversely subovate, c. 1.3 (range 1.271.36) times broader than long; surface bare, lacking setal tufts; covered with fine, but distinct low granules arranged into short striae that become longer posteriorly; mesogastric regions well defined; lateral carapace surface generally without obvious, discrete oblique striae except for strong, concave epibranchial sulcus beginning just posterior to exorbital tooth. Dorsal carapace, lateral branchial regions markedly swollen. Exorbital tooth triangular, pointed, outer margin long, weakly oblique, forming rounded obtuse angle posteriorly marking widest point of carapace, especially in larger specimens; margin markedly constricted posteriorly, but no trace of second anterolateral tooth. Postfrontal lobes strongly demarcated, separated by broad grooves, median lobes similar in width to lateral lobes. Front c. 0.65 times carapace width, markedly deflexed, margin straight or slightly concave in dorsal view; laterally triangular, bluntly pointed; frontal margin emarginated, appearing smooth but microscopically granular (
<figureCitation id="DF953188FF90FFC8FC652823928AFCCD" box="[940,1007,836,860]" captionStart="FIGURE 8" captionStartId="11.[151,250,1137,1160]" captionTargetBox="[151,1436,409,1116]" captionTargetId="figure@11.[151,1436,409,1116]" captionTargetPageId="11" captionText="FIGURE 8. Frontal margins (ventral views). A, Chiromantes obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; B, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; C, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; D, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283289/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
B); frontal surface somewhat concave medially but with pair of low lateral swellings posterior to frontal margin. Supraorbital margin entire, semicircular but oblique medial section somewhat straight; secondary rim formed either side making smooth, broadly triangular, deflexed central plate posterior to ocular peduncle. Eye not extending beyond exorbital tooth. Frontal plate protruding as shelf; medial septum broad, largely covering anterior half of anterior half of epistome; antennae, antennules much reduced in size, lodged under overhanging front; basal antennular segment not much swollen; antennal, antennular basal segments adjacent, not separated by septum. Antennal flagellum very short, entering orbit.
</paragraph>
<paragraph id="47112D0DFF90FFC8FF0E2F0395FDFB76" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">Third maxilliped ischium, merus subequal in length; ischium with shallow longitudinal, curved, median sulcus. Inner margin of merus, ischium with long setae, proximal outer margin of ischium, base of exopod with dense setae; outer margin of merus with short scattered setae only. Exopod slender, hidden behind ishium, merus except near base, tip reaching half length of outer margin of merus, flagellum long, slender.</paragraph>
<paragraph id="47112D0DFF90FFC8FF0E2F939451F9E5" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">
Chelipeds subequal, robust. Merus trihedral; posterior border broadly convex emarginate, minutely granular, without indication of subdistal spine or lobe; inner anterior border armed with small pointed granules, broadly triangular; outer surface broadly convex, with fine transverse striae but appearing almost smooth. Carpus subquadrilateral, inner angle moderately produced, apically granular; outer margin, dorsal surface conspicuously granular. Palm dorsal surface slightly striated along superior margin, with scattering of moderate sized granules, otherwise smooth, punctate. Palm outer surface broadly rounded, smooth; no indication of subventral longitudinal ridge. Palm inner face of males smooth except for curved, projecting crest armed with large tubercles posterior to gape (
<figureCitation id="DF953188FF90FFC8FF562E889184F996" box="[159,225,1519,1543]" captionStart="FIGURE 6" captionStartId="9.[151,250,1537,1560]" captionTargetBox="[151,1435,193,1515]" captionTargetId="figure@9.[151,1436,193,1516]" captionTargetPageId="9" captionText="FIGURE 6. Inner views of chelae. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283287/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
C), but which can vary in prominence among specimens. Cutting margin of fixed finger with 3 or 4 blunt teeth, proximal 3 similar in size, distalmost larger, rounded, conical, placed medially; dactylus with 2 teeth proximally, distalmost large, third much smaller blunt tooth slightly distal of medial tooth of ventral finger. Dorsal surface of dactylus smooth. Fingers with chitinous tips; adult males with narrow but distinct gape when fingers closed.
</paragraph>
<paragraph id="47112D0DFF90FFC8FF0E2D1895D2F8DD" blockId="15.[151,1437,151,2012]" pageId="15" pageNumber="16">Walking legs relatively short, broad, flattened; second, third pairs sub-equal, longer than other walking legs. No setae except for scattering of short dark bristles on dactyli, ventral face of propodi. Dorsal margins of meri with blunt subdistal shoulder, otherwise unarmed; outer surfaces of meri with minutely granular transverse striae. P4 merus c. 2.3 times longer than wide; P5 merus similar, c. 2.4 times longer. Carpi with 2 accessory carinae on outer surface but not strongly marked. P4 propodus c. 2.6 times longer than wide; P5 propodus 2.3 times longer than wide. Dactyli 0.8 times length of propodi, slightly recurved, terminating in acute chitinous tip.</paragraph>
<paragraph id="47112D0DFF90FFD7FF0E2C309573FF45" blockId="15.[151,1437,151,2012]" lastBlockId="16.[151,1436,151,212]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Thoracic sternites (
<figureCitation id="DF953188FF90FFC8FE6B2C309083F8FE" box="[418,486,1879,1903]" captionStart="FIGURE 9" captionStartId="12.[151,250,1576,1599]" captionTargetBox="[151,1435,325,1554]" captionTargetId="figure@12.[151,1436,325,1555]" captionTargetPageId="12" captionText="FIGURE 9. Ventral views of thoracic sternum and anterior parts of male abdomens. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283290/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
C) smooth, mostly bare of setae except for few scattered bristles; abdominal cavity reaching to thick transverse setal fringe at junction of sternites 3, 4. Male abdomen (
<figureCitation id="DF953188FF90FFC8FB9E2C1B95DBF805" box="[1111,1214,1916,1940]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="15" pageNumber="16">Fig. 13A</figureCitation>
) moderately broad; telson broadly rounded apically, 1.1 times longer than wide, only slightly longer than somite 6; somite 6 c. 1.9 times wider than long, lateral margins broadly divergent with margins relatively evenly convex; somites 35 trapezoidal, lateral margins of somites 4, 5 relatively straight, lateral margins of somite 3 convex; somite 3 width 3.2 times basal width of telson; somites 1, 2 transversely, longitudinally narrow.
</paragraph>
<caption id="13D17D85FF8FFFD7FF5E2FA69522FA85" httpUri="https://zenodo.org/record/283293/files/figure.png" pageId="16" pageNumber="17" targetBox="[174,1420,278,1178]" targetPageId="16">
<paragraph id="47112D0DFF8FFFD7FF5E2FA69522FA85" blockId="16.[151,1436,1217,1300]" pageId="16" pageNumber="17">
<emphasis id="75DAF11FFF8FFFD7FF5E2FA6907AFB46" bold="true" box="[151,287,1217,1240]" pageId="16" pageNumber="17">FIGURE 12.</emphasis>
<emphasis id="75DAF11FFF8FFFD7FEED2FA6934EFB46" box="[292,555,1217,1239]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="80AE568EFF8FFFD7FEED2FA690C3FB46" box="[292,422,1217,1239]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Chiromantes</taxonomicName>
<taxonomicName id="80AE568EFF8FFFD7FE642FA5934EFB46" box="[429,555,1217,1239]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">C. garfunkel</taxonomicName>
</emphasis>
<taxonomicNameLabel id="6EE94C64FF8FFFD7FDF92FA593E5FB46" box="[560,640,1218,1239]" pageId="16" pageNumber="17" rank="species">
<emphasis id="75DAF11FFF8FFFD7FDF92FA593E5FB46" bold="true" box="[560,640,1218,1239]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I. A, male abdomen (somites 36, telson); B, ventral view of left G1; C, dorsal view of left G1; D, ventral view of distal part of left G1; E, dorsal view of distal part of left G1. All structures denuded. Scales: AC = 1.0 mm; D, E = 0.5 mm.
</paragraph>
</caption>
<paragraph id="47112D0DFF8FFFD7FF0E2E2690F0FA33" blockId="16.[151,1436,1345,2017]" pageId="16" pageNumber="17">
G1 (
<figureCitation id="DF953188FF8FFFD7FF352E2690E6FAC8" box="[252,387,1345,1369]" captionStart="FIGURE 13. A E" captionStartId="17.[151,250,1920,1943]" captionTargetBox="[221,1357,181,1885]" captionTargetId="figure@17.[198,1376,173,1893]" captionTargetPageId="17" captionText="FIGURE 13. A E, Chiromantes silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; F J, C. leptomerus sp. nov., NMNS- 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan. A, F, male abdomen (somites 3 6, telson); B, G, ventral view of left G 1; C, H, dorsal view of left G 1; D, I, ventral view of distal part of left G 1; E, J, dorsal view of distal part of left G 1. All structures denuded. Scales: A C, F H, = 1.0 mm; D, E, I, J = 0.5 mm." httpUri="https://zenodo.org/record/283294/files/figure.png" pageId="16" pageNumber="17">Fig. 13BE</figureCitation>
) relatively slender, almost straight, weakly tapering to broadly convex subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process long, with dorsal margin concave, apically pointed. G2 short, as for the genus.
</paragraph>
<paragraph id="47112D0DFF8FFFD7FF0E2EC99397FA78" blockId="16.[151,1436,1345,2017]" pageId="16" pageNumber="17">
<emphasis id="75DAF11FFF8FFFD7FF0E2EC9904CFA57" bold="true" box="[199,297,1454,1478]" pageId="16" pageNumber="17">Females</emphasis>
. Chelipeds slightly smaller, less robust, lacking raised granular row on inner face of chela. Vulval morphology without clear species-specific characters.
</paragraph>
<paragraph id="47112D0DFF8FFFD7FF0E2E9295E9F9E8" blockId="16.[151,1436,1345,2017]" pageId="16" pageNumber="17">
<emphasis id="75DAF11FFF8FFFD7FF0E2E929001F99F" bold="true" box="[199,356,1525,1550]" pageId="16" pageNumber="17">Colour in life</emphasis>
. Colour photographs taken from fresh specimens are shown in
<figureCitation id="DF953188FF8FFFD7FBED2E929506F99C" box="[1060,1123,1525,1549]" captionStart="FIGURE 3" captionStartId="3.[151,250,1854,1877]" captionTargetBox="[174,1416,179,1840]" captionTargetId="figure@3.[174,1418,179,1840]" captionTargetPageId="3" captionText="FIGURE 3. Colours in life. A C, Chiromantes silus sp. nov., Guam; A, ZRC 2000.0573, female (15.2 × 11.2 mm); B, C, ZRC 2001.0742, male (15.7 × 11.7 mm). D H, Chiromantes leptomerus sp. nov., D, RUMF-ZC- 2104, male (17.5 × 13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC- 2076, male (19.7 × 14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1 × 13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. I J, Chiromantes eurymerus sp. nov., I, NMNS, male (17.5 × 13.4 mm); J, NMNS, male (16.7 × 12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H. - C. Liu; D E by Y. Fujita; F J by H. - T. Shih." httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
AC. Background colour of carapace typically maroon, with coarse paler orange to yellow-green blotching. Legs speckled with some indication of transverse banding. Legs and chelipeds dorsally reddish brown to dark orange, becoming paler orange in frontal and ventral view. Ocular peduncles speckled similar to carapace; corneas darker.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8FFFD7FF0E2DE1932CF898" pageId="16" pageNumber="17" type="discussion">
<paragraph id="47112D0DFF8FFFD7FF0E2DE1932CF898" blockId="16.[151,1436,1345,2017]" pageId="16" pageNumber="17">
<emphasis id="75DAF11FFF8FFFD7FF0E2DE19051F90F" bold="true" box="[199,308,1670,1694]" pageId="16" pageNumber="17">Remarks</emphasis>
. The numerous features that distinguish this species from all others in the genus are given in
<tableCitation id="0A2C18B6FF8FFFD7FA8A2DE294F3F90C" box="[1347,1430,1669,1693]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[159,1428,329,1954]" captionTargetPageId="5" captionText="TABLE 1. Summary of major differences between species of the Chiromantes obtusifrons complex." httpUri="http://table.plazi.org/id/13D17D85FF9AFFC2FF5E2BF095A0FF3E" pageId="16" pageNumber="17" tableUuid="13D17D85FF9AFFC2FF5E2BF095A0FF3E">Table 1</tableCitation>
. The greater marked constriction of the carapace behind the epibranchial teeth, the shape of the tip of the G1, the relatively straight frontal margin, the abdominal proportions, and the colour of the live crabs are all important characters to help identify this species.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8FFFD7FF0E2C719346F8C3" pageId="16" pageNumber="17" type="etymology">
<paragraph id="47112D0DFF8FFFD7FF0E2C719346F8C3" blockId="16.[151,1436,1345,2017]" pageId="16" pageNumber="17">
<emphasis id="75DAF11FFF8FFFD7FF0E2C719023F8BF" bold="true" box="[199,326,1814,1838]" pageId="16" pageNumber="17">Etymology</emphasis>
. The name
<taxonomicName id="80AE568EFF8FFFD7FE072C729367F8BC" box="[462,514,1813,1837]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="silus">
<emphasis id="75DAF11FFF8FFFD7FE072C729367F8BC" box="[462,514,1813,1837]" italics="true" pageId="16" pageNumber="17">silus</emphasis>
</taxonomicName>
is Latin for “pug-nosed” in reference to the broad, flattened frontal plate that the new species shares with congeners.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF8FFFD7FF0E2C3A90E4F870" pageId="16" pageNumber="17" type="distribution">
<paragraph id="47112D0DFF8FFFD7FF0E2C3A93A2F8E4" blockId="16.[151,1436,1345,2017]" box="[199,711,1885,1910]" pageId="16" pageNumber="17">
<emphasis id="75DAF11FFF8FFFD7FF0E2C3A9005F8E7" bold="true" box="[199,352,1885,1910]" pageId="16" pageNumber="17">Distribution.</emphasis>
So far only known from
<collectingCountry id="3FB96D9DFF8FFFD7FDB22C3A93A6F8E4" box="[635,707,1885,1909]" name="Guam" pageId="16" pageNumber="17">Guam</collectingCountry>
.
</paragraph>
<paragraph id="47112D0DFF8FFFD7FF0E2CE590E4F870" blockId="16.[151,1436,1345,2017]" pageId="16" pageNumber="17">
<emphasis id="75DAF11FFF8FFFD7FF0E2CE59041F80B" bold="true" box="[199,292,1922,1946]" pageId="16" pageNumber="17">Ecology</emphasis>
. Reported to live “under rocks and in karstic crevices in sparsely vegetated coastal platforms, up to ca.
<quantity id="805680E8FF8FFFD7FF5E2CC291B4F82C" box="[151,209,1957,1981]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="m" value="50.0">50 m</quantity>
inland and ca.
<quantity id="805680E8FF8FFFD7FEB72CC290D2F82C" box="[382,439,1957,1981]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="m" value="10.0">10 m</quantity>
elevation” (Paulay &amp; Starmer 2011: 11); and this is confirmed by personal observations of the second author.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="47112D0DFF99FFC1FF5E2E929373F99E" blockId="6.[151,601,1525,1584]" box="[151,534,1525,1551]" pageId="6" pageNumber="7">
<heading id="1C599A61FF99FFC1FF5E2E929373F99E" bold="true" box="[151,534,1525,1551]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="75DAF11FFF99FFC1FF5E2E929373F99E" bold="true" box="[151,534,1525,1551]" pageId="6" pageNumber="7">
<taxonomicName id="80AE568EFF99FFC1FF5E2E9290D7F99E" ID-CoL="7TVP3" box="[151,434,1525,1551]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF99FFC1FF5E2E9290D7F99E" bold="true" box="[151,434,1525,1551]" italics="true" pageId="6" pageNumber="7">Chiromantes garfunkel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF99FFC1FE702E929373F99E" box="[441,534,1525,1551]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="47112D0DFF99FFC1FF5E2D7F933CF9A1" blockId="6.[151,601,1525,1584]" box="[151,601,1560,1584]" pageId="6" pageNumber="7">
(
<figureCitation id="DF953188FF99FFC1FF562D7F918DF9A1" box="[159,232,1560,1584]" captionStart="FIGURE 2" captionStartId="2.[151,246,1309,1332]" captionTargetBox="[151,1435,153,1287]" captionTargetId="figure@2.[151,1436,153,1288]" captionTargetPageId="2" captionText="FIGURE 2. Colours in life. Chiromantes garfunkel sp. nov. A, ZRC 2012.0782, paratype female (11.1 × 8.3 mm), Flying Fish Cove; B D, QM-W 29172, holotype male (17.0 × 13.0 mm), Greta Beach; E, ZRC 2012.0783, paratype ovig. female (11.4 × 8.6 mm), Flying Fish Cove; F, ZRC 2012.0786, paratype female (11.6 × 8.6 mm), Lily Beach. All from Christmas I .. Photographs by H. H. Tan." httpUri="https://zenodo.org/record/283283/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
,
<figureCitation id="DF953188FF99FFC1FF3A2D7F9067F9A1" box="[243,258,1560,1584]" captionStart="FIGURE 4" captionStartId="7.[151,250,1841,1864]" captionTargetBox="[151,1435,717,1822]" captionTargetId="figure@7.[151,1436,715,1823]" captionTargetPageId="7" captionText="FIGURE 4. Overall dorsal views. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; C, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; D, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; E, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283285/files/figure.png" pageId="6" pageNumber="7">4</figureCitation>
E, 5F, 6F, 7F, 8D, 9F, 10B, 12)
</paragraph>
<paragraph id="47112D0DFF99FFC1FF5E2D0693A2F9E4" blockId="6.[151,1437,1632,1746]" box="[151,711,1632,1654]" pageId="6" pageNumber="7">
<taxonomicName id="80AE568EFF99FFC1FF5E2D069366F9E7" box="[151,515,1632,1654]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusifrons" subGenus="Holometopus">
<emphasis id="75DAF11FFF99FFC1FF5E2D06918BF9E7" box="[151,238,1633,1654]" italics="true" pageId="6" pageNumber="7">Sesarma</emphasis>
(
<emphasis id="75DAF11FFF99FFC1FF342D0790E1F9E7" box="[253,388,1632,1654]" italics="true" pageId="6" pageNumber="7">Holometopus</emphasis>
)
<emphasis id="75DAF11FFF99FFC1FE5B2D079366F9E7" box="[402,515,1632,1654]" italics="true" pageId="6" pageNumber="7">obtusifrons</emphasis>
</taxonomicName>
Balss 1934: 229.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF99FFC1FF5E2D18933CF940" pageId="6" pageNumber="7" type="description">
<paragraph id="47112D0DFF99FFC1FF5E2D189362F922" blockId="6.[151,1437,1632,1746]" pageId="6" pageNumber="7">
<taxonomicName id="80AE568EFF99FFC1FF5E2D189000F904" box="[151,357,1663,1685]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF99FFC1FF5E2D189000F904" box="[151,357,1663,1685]" italics="true" pageId="6" pageNumber="7">Sesarma obtusifrons</emphasis>
</taxonomicName>
—Gibson-Hill 1947: 44.—Tweedie 1947: 33.—George 1978: 13 (unnumbered pages).—Hicks
<emphasis id="75DAF11FFF99FFC1FAE32DE79438F904" box="[1322,1373,1663,1685]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
1984: 22, 65 including colour picture.
</paragraph>
<paragraph id="47112D0DFF99FFC1FF5E2DDB933CF940" blockId="6.[151,1437,1632,1746]" box="[151,601,1724,1746]" pageId="6" pageNumber="7">
<taxonomicName id="80AE568EFF99FFC1FF5E2DDB90F5F943" box="[151,400,1724,1746]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF99FFC1FF5E2DDB90F5F943" box="[151,400,1724,1746]" italics="true" pageId="6" pageNumber="7">Chiromantes obtusifrons</emphasis>
</taxonomicName>
—Davie 2002: 221.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF99FFC0FF5E2D99942BFD36" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="47112D0DFF99FFC0FF5E2D99942BFD36" blockId="6.[151,1436,1790,2031]" lastBlockId="7.[151,1436,151,679]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="75DAF11FFF99FFC1FF5E2D9990E6F886" bold="true" box="[151,387,1790,1815]" pageId="6" pageNumber="7">Material Examined</emphasis>
.
<typeStatus id="981593AFFF99FFC1FE5C2D999348F887" box="[405,557,1790,1814]" pageId="6" pageNumber="7" type="holotype">HOLOTYPE</typeStatus>
: QM-W29172, male (17.0×13.0 mm), Greta Beach,
<collectingCountry id="3FB96D9DFF99FFC1FB5A2D999433F887" box="[1171,1366,1790,1814]" name="Christmas Island" pageId="6" pageNumber="7">Christmas Island</collectingCountry>
, H.H. Tan, 0
<date id="33100BCDFF99FFC1FF202C459038F8AB" box="[233,349,1826,1850]" pageId="6" pageNumber="7" value="2007-08-12">8.12.2007</date>
.
<typeStatus id="981593AFFF99FFC1FEBD2C459376F8AB" box="[372,531,1826,1850]" pageId="6" pageNumber="7" type="paratype">PARATYPES</typeStatus>
: ZRC 2012.0776, female (13.3×12.0 mm), station CI-33, Greta Beach,
<geoCoordinate id="229A4BCAFF99FFC1FF5E2C219047F8CF" box="[151,290,1862,1886]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-10.502117">10°30.127'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF99FFC1FEFB2C2190A8F8CF" box="[306,461,1862,1886]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="105.67458">105°40.475'E</geoCoordinate>
,
<date id="33100BCDFF99FFC1FE142C219339F8CF" box="[477,604,1862,1886]" pageId="6" pageNumber="7" value="2011-03-27">27.03.2011</date>
.—ZRC 2012.0777, male (13.9×
<quantity id="805680E8FF99FFC1FC1C2C219526F8CF" box="[981,1091,1862,1886]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="6" pageNumber="7" unit="mm" value="10.6">10.6 mm</quantity>
), station CI-12, Greta Beach,
<geoCoordinate id="229A4BCAFF99FFC1FF5E2C0D9047F813" box="[151,290,1898,1922]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-10.502117">10°30.127'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF99FFC1FEFB2C0D90A8F813" box="[306,461,1898,1922]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="105.67458">105°40.475'E</geoCoordinate>
, limestone cliff, sandy beach, limestone bedrock,
<date id="33100BCDFF99FFC1FBE92C0D95C5F813" box="[1056,1184,1898,1922]" pageId="6" pageNumber="7" value="2010-01-24">24.01.2010</date>
.—ZRC 2012.0778,
<specimenCount id="51A8E684FF99FFC1FA472C0D91BDF837" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
(13.5×10.2, 15.2×11.6, 18.2×14.7, 17.9×14.0 mm),
<specimenCount id="51A8E684FF99FFC1FCF82CE992C5F837" box="[817,928,1934,1958]" pageId="6" pageNumber="7" type="female">4 females</specimenCount>
(13.0×9.6, 14.3×10.8, 14.7×11.2, 17.5×
<quantity id="805680E8FF99FFC1FAA32CE991A2F85B" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="6" pageNumber="7" unit="mm" value="13.6">13.6 mm</quantity>
), station CI-06, The Grotto, Waterfall Road,
<geoCoordinate id="229A4BCAFF99FFC1FD332CD592E0F85B" box="[762,901,1970,1994]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-10.4230995">10°25.386'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF99FFC1FC532CD59550F85B" box="[922,1077,1970,1994]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="105.70212">105°42.127'E</geoCoordinate>
, weathered cave, tidal sump,
<date id="33100BCDFF99FFC1FF5E2CB19072F87F" box="[151,279,2006,2030]" pageId="6" pageNumber="7" value="2010-01-23">23.01.2010</date>
.—ZRC 2012.0779,
<specimenCount id="51A8E684FF99FFC1FE322CB19337F87F" box="[507,594,2006,2030]" pageId="6" pageNumber="7" type="male">2 males</specimenCount>
(10.8×9.8, 14.5×
<quantity id="805680E8FF99FFC1FCDF2CB1921AF87F" box="[790,895,2006,2030]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="6" pageNumber="7" unit="mm" value="13.2">13.2 mm</quantity>
), station CI-07, Waterfall Bay,
<collectingCountry id="3FB96D9DFF99FFC1FB172CB194F9F87F" box="[1246,1436,2006,2030]" name="Christmas Island" pageId="6" pageNumber="7">Christmas Island</collectingCountry>
Resort,
<geoCoordinate id="229A4BCAFF98FFC0FF3E2BF09010FF3E" box="[247,373,151,175]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="9" value="-10.459">10°27.54'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FE402BF09373FF3E" box="[393,534,151,175]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="9" value="105.705">105°42.30'E</geoCoordinate>
, freshwater stream, sandy beach, limestone base rock,
<date id="33100BCDFF98FFC0FB0C2BF09420FF3E" box="[1221,1349,151,175]" pageId="7" pageNumber="8" value="2010-01-23">23.01.2010</date>
.—ZRC 2012.0780, female (12.1×
<quantity id="805680E8FF98FFC0FE722BDB9372FF45" box="[443,535,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="8" unit="mm" value="9.1">9.1 mm</quantity>
), station CI-04, Merrial Beach,
<geoCoordinate id="229A4BCAFF98FFC0FC482BDB956EFF45" box="[897,1035,188,212]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.47425">10°28.455'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FBD02BDB95D1FF45" box="[1049,1204,188,212]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.55918">105°33.551'E</geoCoordinate>
, sandy beach, beach forest, limestone base rock,
<date id="33100BCDFF98FFC0FE2A2BB89307FF66" box="[483,610,223,247]" pageId="7" pageNumber="8" value="2011-03-20">20.03.2011</date>
.—ZRC 2012.0781, male (13.5×
<quantity id="805680E8FF98FFC0FC102BB89522FF66" box="[985,1095,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="7" pageNumber="8" unit="mm" value="10.2">10.2 mm</quantity>
), station CI-09, Ethel Beach,
<geoCoordinate id="229A4BCAFF98FFC0FF5E2A639047FE8D" box="[151,290,260,284]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.463417">10°27.805'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FEE42A6390ADFE8D" box="[301,456,260,284]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.70738">105°42.443'E</geoCoordinate>
, sandy beach, beach forest, limestone base rock,
<date id="33100BCDFF98FFC0FC242A639509FE8D" box="[1005,1132,260,284]" pageId="7" pageNumber="8" value="2011-03-21">21.03.2011</date>
.—ZRC 2012.0782, female (11.1×
<quantity id="805680E8FF98FFC0FF162A40902EFEAE" box="[223,331,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="7" pageNumber="8" unit="mm" value="8.3">8.3 mm</quantity>
), station CI-01, Flying Fish Cove,
<geoCoordinate id="229A4BCAFF98FFC0FCF12A4092A6FEAE" box="[824,963,295,319]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.43025">10°25.815'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FC292A40951EFEAE" box="[992,1147,295,319]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.66967">105°40.180'E</geoCoordinate>
, rocky, gravel beach,
<date id="33100BCDFF98FFC0FF5E2A2B9072FEF5" box="[151,279,332,356]" pageId="7" pageNumber="8" value="2010-01-22">22.01.2010</date>
.—ZRC 2012.0783, 1 ovig. female (11.4×
<quantity id="805680E8FF98FFC0FCCF2A2B9202FEF5" box="[774,871,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="7" pageNumber="8" unit="mm" value="8.6">8.6 mm</quantity>
), station CI-01, Flying Fish Cove,
<geoCoordinate id="229A4BCAFF98FFC0FAC52A2B94F2FEF5" box="[1292,1431,332,356]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.43025">10°25.815'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FF5E2A089057FE16" box="[151,306,367,391]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.66967">105°40.180'E</geoCoordinate>
, rocky, gravel beach,
<date id="33100BCDFF98FFC0FDF62A0893DAFE16" box="[575,703,367,391]" pageId="7" pageNumber="8" value="2010-01-22">22.01.2010</date>
.—ZRC 2012.0784, male (14.0×
<quantity id="805680E8FF98FFC0FBF42A0895C9FE16" box="[1085,1196,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="7" pageNumber="8" unit="mm" value="10.8">10.8 mm</quantity>
), Flying Fish Cove,
<geoCoordinate id="229A4BCAFF98FFC0FF5E2AF39047FE3D" box="[151,290,404,428]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.43025">10°25.815'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FEE42AF390ADFE3D" box="[301,456,404,428]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.66967">105°40.180'E</geoCoordinate>
, rocky, gravel beach,
<date id="33100BCDFF98FFC0FD732AF3925FFE3D" box="[698,826,404,428]" pageId="7" pageNumber="8" value="2010-01-22">22.01.2010</date>
.—ZRC 2012.0785, female (10.8×
<quantity id="805680E8FF98FFC0FB712AF39476FE3D" box="[1208,1299,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="7" pageNumber="8" unit="mm" value="8.2">8.2 mm</quantity>
), station CI- 0 2, Lily Beach,
<geoCoordinate id="229A4BCAFF98FFC0FE9D2AD090B8FE5E" box="[340,477,439,463]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.46685">10°28.011'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FE252AD093E2FE5E" box="[492,647,439,463]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.711464">105°42.688'E</geoCoordinate>
, sandy beach, beach forest, limestone base rock,
<date id="33100BCDFF98FFC0FB0D2AD09421FE5E" box="[1220,1348,439,463]" pageId="7" pageNumber="8" value="2010-01-22">22.01.2010</date>
.—ZRC 2012.0786, female (11.6×
<quantity id="805680E8FF98FFC0FE092ABB9345FE65" box="[448,544,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="7" pageNumber="8" unit="mm" value="8.6">8.6 mm</quantity>
), station CI-02, Lily Beach,
<geoCoordinate id="229A4BCAFF98FFC0FCBC2ABB929BFE65" box="[885,1022,476,500]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.46685">10°28.011'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FBC62ABB95CFFE65" box="[1039,1194,476,500]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.711464">105°42.688'E</geoCoordinate>
, sandy beach, beach forest, limestone base rock,
<date id="33100BCDFF98FFC0FE122A989338FD86" box="[475,605,511,535]" pageId="7" pageNumber="8" value="2010-01-22">22.01.2010</date>
.—QM-W29173, male (18.2×
<quantity id="805680E8FF98FFC0FC642A98957DFD86" box="[941,1048,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="7" pageNumber="8" unit="mm" value="13.8">13.8 mm</quantity>
),
<specimenCount id="51A8E684FF98FFC0FBE42A9895F9FD86" box="[1069,1180,511,535]" pageId="7" pageNumber="8" type="female">3 females</specimenCount>
(12.8×9.8, 16.5×12.7, 16.6×
<quantity id="805680E8FF98FFC0FF112943902CFDAD" box="[216,329,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="8" unit="mm" value="12.7">12.7 mm</quantity>
),
<date id="33100BCDFF98FFC0FEAC29439080FDAD" box="[357,485,548,572]" pageId="7" pageNumber="8" value="2011-03-31">31.03.2011</date>
, The Blowholes,
<geoCoordinate id="229A4BCAFF98FFC0FD0D2943922AFDAD" box="[708,847,548,572]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.51435">10°30.861'S</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF98FFC0FCAC29439575FDAD" box="[869,1040,548,572]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="105.622635">105°37.358' E</geoCoordinate>
, under limestone rocks adjacent vegetation at back of limestone terrace, P. Davie.—ZRC 1965.7.29.153, male (13.7×
<quantity id="805680E8FF98FFC0FB90292095A1FDCE" box="[1113,1220,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="7" pageNumber="8" unit="mm" value="10.4">10.4 mm</quantity>
), 1932, no specific location.—ZRC 1965.7.29.154-163,
<specimenCount id="51A8E684FF98FFC0FDF5290B93F2FD15" box="[572,663,620,644]" pageId="7" pageNumber="8" type="male">2 males</specimenCount>
(larger 11.5×
<quantity id="805680E8FF98FFC0FCFF290B92F1FD15" box="[822,916,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="7" pageNumber="8" unit="mm" value="8.8">8.8 mm</quantity>
),
<specimenCount id="51A8E684FF98FFC0FC62290A957EFD15" box="[939,1051,620,644]" pageId="7" pageNumber="8" type="female">7 females</specimenCount>
(largest 20.4×
<quantity id="805680E8FF98FFC0FB0F290B9457FD15" box="[1222,1330,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="7" pageNumber="8" unit="mm" value="16.2">16.2 mm</quantity>
), M.W.F. Tweedie, shore terrace along east and north coasts, and Isabel Beach, 0 2.03.1932. All localities Christmas I.
</paragraph>
</subSubSection>
<caption id="13D17D85FF98FFC0FF5E2C56928AF832" httpUri="https://zenodo.org/record/283285/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1435,717,1822]" targetPageId="7">
<paragraph id="47112D0DFF98FFC0FF5E2C56928AF832" blockId="7.[151,1436,1841,1955]" pageId="7" pageNumber="8">
<emphasis id="75DAF11FFF98FFC0FF5E2C569071F8D6" bold="true" box="[151,276,1841,1864]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
Overall dorsal views. A,
<taxonomicName id="80AE568EFF98FFC0FDD62C5692CEF8D6" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[543,939,1841,1863]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF98FFC0FDD62C56927CF8D6" box="[543,793,1841,1863]" italics="true" pageId="7" pageNumber="8">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, UFUF-14837, male (17.7×13.2 mm), Hawaiian Is.; B,
<taxonomicName id="80AE568EFF98FFC0FF122C36904DF8F7" box="[219,296,1872,1894]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF98FFC0FF122C36904DF8F7" box="[219,296,1872,1894]" italics="true" pageId="7" pageNumber="8">C. silus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF98FFC0FEE72C36901AF8F7" bold="true" box="[302,383,1873,1894]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="6EE94C64FF98FFC0FEE72C36901AF8F7" box="[302,383,1873,1894]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZRC 2012.0787, holotype male (16.3×12.8 mm), Guam; C,
<taxonomicName id="80AE568EFF98FFC0FC382C3695E5F8F7" box="[1009,1152,1872,1894]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF98FFC0FC382C3695E5F8F7" box="[1009,1152,1872,1894]" italics="true" pageId="7" pageNumber="8">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF98FFC0FB4F2C3695B2F8F7" bold="true" box="[1158,1239,1873,1894]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="6EE94C64FF98FFC0FB4F2C3695B2F8F7" box="[1158,1239,1873,1894]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS 7028-001, holotype male (16.8×12.7 mm), Taiwan; D,
<taxonomicName id="80AE568EFF98FFC0FD9F2C0893BAF814" box="[598,735,1903,1925]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF98FFC0FD9F2C0893BAF814" box="[598,735,1903,1925]" italics="true" pageId="7" pageNumber="8">C. eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF98FFC0FD2C2C089250F815" bold="true" box="[741,821,1903,1924]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="6EE94C64FF98FFC0FD2C2C089250F815" box="[741,821,1903,1924]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS-7028-002, holotype male (18.7×14.8 mm), Taiwan; E,
<taxonomicName id="80AE568EFF98FFC0FF7A2CE99054F832" box="[179,305,1933,1955]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF98FFC0FF7A2CE99054F832" box="[179,305,1933,1955]" italics="true" pageId="7" pageNumber="8">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF98FFC0FEF12CE990EDF832" bold="true" box="[312,392,1934,1955]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="6EE94C64FF98FFC0FEF12CE990EDF832" box="[312,392,1934,1955]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I.
</paragraph>
</caption>
<caption id="13D17D85FF97FFCFFF5E2E12933EF996" httpUri="https://zenodo.org/record/283286/files/figure.png" pageId="8" pageNumber="9" targetBox="[151,1435,187,1364]" targetPageId="8">
<paragraph id="47112D0DFF97FFCFFF5E2E12933EF996" blockId="8.[151,1436,1397,1543]" pageId="8" pageNumber="9">
<emphasis id="75DAF11FFF97FFCFFF5E2E129076FA1A" bold="true" box="[151,275,1397,1420]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
Outer views of chelae. A,
<taxonomicName id="80AE568EFF97FFCFFDEF2E1192CAFA1D" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[550,943,1398,1420]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF97FFCFFDEF2E119245FA1D" box="[550,800,1398,1420]" italics="true" pageId="8" pageNumber="9">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, UFUF-14837, male (17.7×13.2 mm), Hawaiian Is.; B,
<taxonomicName id="80AE568EFF97FFCFFF132EF29093FA3B" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[218,502,1428,1451]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF97FFCFFF132EF2900DFA3A" box="[218,360,1429,1451]" italics="true" pageId="8" pageNumber="9">C. obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, ZRC-2002.0220, male (19.7×14.7 mm), Hawaiian Is.; C,
<taxonomicName id="80AE568EFF97FFCFFB8D2EF295EAFA3A" box="[1092,1167,1429,1451]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF97FFCFFB8D2EF295EAFA3A" box="[1092,1167,1429,1451]" italics="true" pageId="8" pageNumber="9">C. silus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF97FFCFFB5F2EF29583FA3B" bold="true" box="[1174,1254,1429,1450]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="6EE94C64FF97FFCFFB5F2EF29583FA3B" box="[1174,1254,1429,1450]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZRC 2012.0787, holotype male (16.3×12.8 mm), Guam; D,
<taxonomicName id="80AE568EFF97FFCFFD842ED393B9FA58" box="[589,732,1459,1481]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF97FFCFFD842ED393B9FA58" box="[589,732,1459,1481]" italics="true" pageId="8" pageNumber="9">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF97FFCFFD2A2ED39251FA58" bold="true" box="[739,820,1460,1481]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="6EE94C64FF97FFCFFD2A2ED39251FA58" box="[739,820,1460,1481]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS 7028-001, holotype male (16.8×12.7 mm), Taiwan; E,
<taxonomicName id="80AE568EFF97FFCFFF7D2EB4905BFA79" box="[180,318,1491,1512]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF97FFCFFF7D2EB4905BFA79" box="[180,318,1491,1512]" italics="true" pageId="8" pageNumber="9">C. eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF97FFCFFE8C2EB590F3FA76" bold="true" box="[325,406,1490,1511]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="6EE94C64FF97FFCFFE8C2EB590F3FA76" box="[325,406,1490,1511]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS-7028-002, holotype male (18.7×14.8 mm), Taiwan; F,
<taxonomicName id="80AE568EFF97FFCFFBEF2EB495C3FA79" box="[1062,1190,1490,1512]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF97FFCFFBEF2EB495C3FA79" box="[1062,1190,1490,1512]" italics="true" pageId="8" pageNumber="9">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF97FFCFFB642EB5959AFA76" bold="true" box="[1197,1279,1490,1511]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="6EE94C64FF97FFCFFB642EB5959AFA76" box="[1197,1279,1490,1511]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I.
</paragraph>
</caption>
<subSubSection id="0FB47E86FF97FFCFFF0E2D309098F846" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="47112D0DFF97FFCFFF0E2D309098F846" blockId="8.[151,1436,1623,2007]" pageId="8" pageNumber="9">
<emphasis id="75DAF11FFF97FFCFFF0E2D30905BF9E1" bold="true" box="[199,318,1623,1648]" pageId="8" pageNumber="9">Diagnosis</emphasis>
. Carapace transversely subovate, c. 1.3 times broader than long; dorsal carapace, lateral branchial regions prominently swollen; exorbital tooth at widest point only projecting slightly more than exorbital tooth; front c. 0.6 times carapace width, margin broadly convex in dorsal view, appearing smooth but microscopically granular, with pair of weak lateral swellings behind margin; supraorbital margin entire, straight, obliquely sloping posteriorly; outer surface of cheliped carpus moderately granular; walking legs relatively short; P4 merus c. 2.3 times longer than wide; P5 merus relatively narrower than P4 merus, c. 2.6 times longer; P4 propodus c. 2.8 times longer than wide; P5 propodus c. 2.7 times longer; male abdomen (
<figureCitation id="DF953188FF97FFCFFAA42C4891D6F8FA" captionStart="FIGURE 12" captionStartId="16.[151,250,1217,1240]" captionTargetBox="[174,1420,278,1178]" captionTargetId="figure@16.[151,1435,265,1196]" captionTargetPageId="16" captionText="FIGURE 12. Chiromantes C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I. A, male abdomen (somites 3 6, telson); B, ventral view of left G 1; C, dorsal view of left G 1; D, ventral view of distal part of left G 1; E, dorsal view of distal part of left G 1. All structures denuded. Scales: A C = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/283293/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
A) moderately broad; somite 6 with lateral margins diverging, slightly concave over distal half, more subparallel over proximal half; somite 3 width 3.0 times basal width of telson; G1 (
<figureCitation id="DF953188FF97FFCFFB592C109583F81E" box="[1168,1254,1911,1935]" captionStart="FIGURE 12" captionStartId="16.[151,250,1217,1240]" captionTargetBox="[174,1420,278,1178]" captionTargetId="figure@16.[151,1435,265,1196]" captionTargetPageId="16" captionText="FIGURE 12. Chiromantes C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I. A, male abdomen (somites 3 6, telson); B, ventral view of left G 1; C, dorsal view of left G 1; D, ventral view of distal part of left G 1; E, dorsal view of distal part of left G 1. All structures denuded. Scales: A C = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/283293/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
BE) relatively slender, weakly tapering to prominent subdistal shoulder; terminal process long, with slender narrow apex, dorsal margin slightly sinuous.
</paragraph>
</subSubSection>
<caption id="13D17D85FF96FFCEFF5E2D66933EF903" httpUri="https://zenodo.org/record/283287/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1435,193,1515]" targetPageId="9">
<paragraph id="47112D0DFF96FFCEFF5E2D66933EF903" blockId="9.[151,1436,1537,1682]" pageId="9" pageNumber="10">
<emphasis id="75DAF11FFF96FFCEFF5E2D669076F986" bold="true" box="[151,275,1537,1560]" pageId="9" pageNumber="10">FIGURE 6.</emphasis>
Inner views of chelae. A,
<taxonomicName id="80AE568EFF96FFCEFDED2D6692CBF986" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[548,942,1537,1559]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF96FFCEFDED2D66927BF986" box="[548,798,1537,1559]" italics="true" pageId="9" pageNumber="10">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, UFUF-14837, male (17.7×13.2 mm), Hawaiian Is.; B,
<taxonomicName id="80AE568EFF96FFCEFF132D469093F9A7" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[218,502,1568,1590]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF96FFCEFF132D46900DF9A7" box="[218,360,1568,1590]" italics="true" pageId="9" pageNumber="10">C. obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, ZRC 2002.0220, male (19.7×14.7 mm), Hawaiian Is.; C,
<taxonomicName id="80AE568EFF96FFCEFB8A2D4695EAF9A7" box="[1091,1167,1568,1590]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF96FFCEFB8A2D4695EAF9A7" box="[1091,1167,1568,1590]" italics="true" pageId="9" pageNumber="10">C. silus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF96FFCEFB5C2D469580F9A7" bold="true" box="[1173,1253,1569,1590]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="6EE94C64FF96FFCEFB5C2D469580F9A7" box="[1173,1253,1569,1590]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZRC 2012.0787, holotype male (16.3×12.8 mm), Guam; D,
<taxonomicName id="80AE568EFF96FFCEFD842D5893B9F9C4" box="[589,732,1599,1621]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF96FFCEFD842D5893B9F9C4" box="[589,732,1599,1621]" italics="true" pageId="9" pageNumber="10">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF96FFCEFD2A2D589251F9C5" bold="true" box="[739,820,1599,1620]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="6EE94C64FF96FFCEFD2A2D589251F9C5" box="[739,820,1599,1620]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS 7028-001, holotype male (16.8×12.7 mm), Taiwan; E,
<taxonomicName id="80AE568EFF96FFCEFF7D2D39905BF9E2" box="[180,318,1630,1651]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF96FFCEFF7D2D39905BF9E2" box="[180,318,1630,1651]" italics="true" pageId="9" pageNumber="10">C. eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF96FFCEFE8C2D3990F3F9E2" bold="true" box="[325,406,1630,1651]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="6EE94C64FF96FFCEFE8C2D3990F3F9E2" box="[325,406,1630,1651]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS-7028-002, holotype male (18.7×14.8 mm), Taiwan; F,
<taxonomicName id="80AE568EFF96FFCEFBEF2D3995C3F9E2" box="[1062,1190,1629,1651]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF96FFCEFBEF2D3995C3F9E2" box="[1062,1190,1629,1651]" italics="true" pageId="9" pageNumber="10">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF96FFCEFB642D39959AF9E2" bold="true" box="[1197,1279,1630,1651]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="6EE94C64FF96FFCEFB642D39959AF9E2" box="[1197,1279,1630,1651]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I.
</paragraph>
</caption>
<subSubSection id="0FB47E86FF96FFCBFF0E2DD99001FE8D" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
<paragraph id="47112D0DFF96FFCDFF0E2DD99293FE3D" blockId="9.[151,1436,1726,2004]" lastBlockId="10.[151,1436,151,428]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="75DAF11FFF96FFCEFF0E2DD9903CF946" bold="true" box="[199,345,1726,1751]" pageId="9" pageNumber="10">Description.</emphasis>
Carapace transversely subovate, c. 1.3 (range 1.261.35) times broader than long; surface bare, lacking setal tufts; covered with fine, but distinct low granules becoming arranged into short striae posteriorly; mesogastric regions well defined; lateral carapace surface generally without obvious discrete oblique striae, except for strong, concave epibranchial sulcus beginning just behind exorbital tooth. Dorsal carapace, lateral branchial regions relatively weakly convex, not prominently swollen (
<figureCitation id="DF953188FF96FFCEFCF32C2892EFF8F6" box="[826,906,1871,1895]" captionStart="FIGURE 10" captionStartId="13.[151,250,593,616]" captionTargetBox="[151,1436,193,571]" captionTargetId="figure@13.[151,1436,193,572]" captionTargetPageId="13" captionText="FIGURE 10. Lateral carapace views. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283291/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
A, B). Exorbital tooth triangular, pointed, outer margin long, slightly convex, at widest point only projecting slightly more than exorbital tooth; margin constricted behind, but no trace of second anterolateral tooth. Postfrontal lobes strongly demarcated, separated by broad grooves, median lobes similar in width to lateral lobes. Front c. 0.6 times carapace width (
<quantity id="805680E8FF96FFCEFB032CDC947BF842" box="[1226,1310,1979,2003]" metricMagnitude="-2" metricUnit="m" metricValue="1.6002" pageId="9" pageNumber="10" unit="in" value="0.63">0.63 in</quantity>
<typeStatus id="981593AFFF96FFCEFAE32CDC94F5F842" box="[1322,1424,1979,2003]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
), markedly deflexed, margin broadly convex in dorsal view; laterally triangular, bluntly pointed; frontal margin emarginated, minutely granular (
<figureCitation id="DF953188FF95FFCDFDCF2BDB932CFF45" box="[518,585,188,212]" captionStart="FIGURE 8" captionStartId="11.[151,250,1137,1160]" captionTargetBox="[151,1436,409,1116]" captionTargetId="figure@11.[151,1436,409,1116]" captionTargetPageId="11" captionText="FIGURE 8. Frontal margins (ventral views). A, Chiromantes obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; B, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; C, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; D, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283289/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
D); surface somewhat concave behind frontal margin, but with pair of weak lateral swellings. Supraorbital margin entire, straight, obliquely sloping posteriorly at about 45° angle; secondary rim sloping obliquely inwards, downwards to form smooth, broadly triangular, deflexed plate behind ocular peduncle. Eye not extending beyond exorbital tooth. Frontal plate protruding as shelf; medial septum broad, largely covering anterior half of anterior half of epistome; antennae, antennules much reduced in size, lodged under overhanging front; basal antennular segment not much swollen; antennal, antennular basal segments adjacent, not separated by septum. Antennal flagellum conspicuously short, entering orbit.
</paragraph>
<caption id="13D17D85FF95FFCDFF5E2D6492BFF905" httpUri="https://zenodo.org/record/283288/files/figure.png" pageId="10" pageNumber="11" targetBox="[151,1435,481,1517]" targetPageId="10">
<paragraph id="47112D0DFF95FFCDFF5E2D6492BFF905" blockId="10.[151,1436,1539,1684]" pageId="10" pageNumber="11">
<emphasis id="75DAF11FFF95FFCDFF5E2D649076F988" bold="true" box="[151,275,1539,1562]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
Last walking legs. A,
<taxonomicName id="80AE568EFF95FFCDFE352D6492E2F988" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[508,903,1539,1561]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF95FFCDFE352D649393F988" box="[508,758,1539,1561]" italics="true" pageId="10" pageNumber="11">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, UFUF-14837, male (17.7×13.2 mm), Hawaiian Is.; B,
<taxonomicName id="80AE568EFF95FFCDFF7F2D4490BCF9A9" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[182,473,1570,1592]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF95FFCDFF7F2D449023F9A9" box="[182,326,1570,1592]" italics="true" pageId="10" pageNumber="11">C. obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, ZRC 2002.0220, male (19.7×14.7 mm), Hawaiian Is.; C,
<taxonomicName id="80AE568EFF95FFCDFBF02D4495E3F9A9" box="[1081,1158,1570,1592]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF95FFCDFBF02D4495E3F9A9" box="[1081,1158,1570,1592]" italics="true" pageId="10" pageNumber="11">C. silus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF95FFCDFB462D459584F9A6" bold="true" box="[1167,1249,1570,1591]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="6EE94C64FF95FFCDFB462D459584F9A6" box="[1167,1249,1570,1591]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZRC 2012.0787, holotype male (16.3×12.8 mm), Guam; D,
<taxonomicName id="80AE568EFF95FFCDFD842D2693B9F9C7" box="[589,732,1600,1622]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF95FFCDFD842D2693B9F9C7" box="[589,732,1600,1622]" italics="true" pageId="10" pageNumber="11">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF95FFCDFD2A2D269251F9C7" bold="true" box="[739,820,1601,1622]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="6EE94C64FF95FFCDFD2A2D269251F9C7" box="[739,820,1601,1622]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS 7028-001, holotype male (16.8×12.7 mm), Taiwan; E,
<taxonomicName id="80AE568EFF95FFCDFF7D2D07905BF9E4" box="[180,318,1632,1653]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF95FFCDFF7D2D07905BF9E4" box="[180,318,1632,1653]" italics="true" pageId="10" pageNumber="11">C. eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF95FFCDFE8C2D0790F3F9E4" bold="true" box="[325,406,1632,1653]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="6EE94C64FF95FFCDFE8C2D0790F3F9E4" box="[325,406,1632,1653]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS-7028-002, holotype male (18.7×14.8 mm), Taiwan; F,
<taxonomicName id="80AE568EFF95FFCDFBEF2D0795C3F9E4" box="[1062,1190,1631,1653]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF95FFCDFBEF2D0795C3F9E4" box="[1062,1190,1631,1653]" italics="true" pageId="10" pageNumber="11">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF95FFCDFB642D07959AF9E4" bold="true" box="[1197,1279,1632,1653]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="6EE94C64FF95FFCDFB642D07959AF9E4" box="[1197,1279,1632,1653]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I.. A, B, E, right legs; C, D, F, left legs.
</paragraph>
</caption>
<paragraph id="47112D0DFF95FFCDFF0E2DDC9575F8AE" blockId="10.[151,1436,1723,2035]" pageId="10" pageNumber="11">Third maxilliped ischium, merus subequal in length; ischium with shallow longitudinal, curved, median sulcus. Merus, ischium inner margins with long setae, ischium proximal outer margin, base of exopod with dense setae; merus outer margin with short scattered setae only. Exopod slender, hidden behind ishium, merus except near base, tip reaching half length of outer margin of merus, flagellum long, slender.</paragraph>
<paragraph id="47112D0DFF95FFCCFF0E2C2C9415FEF5" blockId="10.[151,1436,1723,2035]" lastBlockId="11.[151,1436,151,357]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Chelipeds subequal, robust. Merus trihedral; posterior border broadly convex, emarginated, minutely granular, without indication of subdistal spine or lobe; inner anterior border minutely granular, broadly triangular; outer surface broadly convex, with fine transverse striae but appearing almost smooth. Carpus subquadrilateral, inner angle moderately produced, apically granular; outer margin, dorsal surface moderately granular. Palm dorsal surface slightly striated along superior margin, with scattering of moderate sized granules, otherwise smooth, punctate. Palm outer surface broadly rounded, smooth; no indication of subventral longitudinal ridge. Inner face of palm in males smooth except for curved projecting crest of large tubercles behind gape (
<figureCitation id="DF953188FF94FFCCFB482BDB95A0FF45" box="[1153,1221,188,212]" captionStart="FIGURE 6" captionStartId="9.[151,250,1537,1560]" captionTargetBox="[151,1435,193,1515]" captionTargetId="figure@9.[151,1436,193,1516]" captionTargetPageId="9" captionText="FIGURE 6. Inner views of chelae. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283287/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
F), but prominence variable. Cutting margin of fixed finger with 3 or 4 blunt teeth (
<figureCitation id="DF953188FF94FFCCFC492BB892A0FF66" box="[896,965,223,247]" captionStart="FIGURE 5" captionStartId="8.[151,250,1397,1420]" captionTargetBox="[151,1435,187,1364]" captionTargetId="figure@8.[151,1436,187,1365]" captionTargetPageId="8" captionText="FIGURE 5. Outer views of chelae. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC- 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283286/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
F), proximal 3 similar in size, distalmost larger, rounded, conical, placed medially; dactylus with 2 proximal teeth, distalmost large, third much smaller blunt tooth slightly distal of medial tooth of ventral finger; teeth on both fingers can vary in prominence. Dorsal surface of dactylus smooth. Fingers with chitinous tips; adult males with narrow but distinct gape when fingers closed.
</paragraph>
<caption id="13D17D85FF94FFCCFF5E2F16902AFB72" httpUri="https://zenodo.org/record/283289/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1436,409,1116]" targetPageId="11">
<paragraph id="47112D0DFF94FFCCFF5E2F16902AFB72" blockId="11.[151,1436,1137,1251]" pageId="11" pageNumber="12">
<emphasis id="75DAF11FFF94FFCCFF5E2F169073FB16" bold="true" box="[151,278,1137,1160]" pageId="11" pageNumber="12">FIGURE 8.</emphasis>
Frontal margins (ventral views). A,
<taxonomicName id="80AE568EFF94FFCCFD552F159548FB19" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[668,1069,1138,1160]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF94FFCCFD552F1592FDFB19" box="[668,920,1138,1160]" italics="true" pageId="11" pageNumber="12">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, ZRC 2002.0220, male (19.7×14.7 mm), Hawaiian Is.; B,
<taxonomicName id="80AE568EFF94FFCCFE4C2FF690B7FB37" box="[389,466,1168,1190]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF94FFCCFE4C2FF690B7FB37" box="[389,466,1168,1190]" italics="true" pageId="11" pageNumber="12">C. silus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF94FFCCFE122FF69348FB37" bold="true" box="[475,557,1169,1190]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="6EE94C64FF94FFCCFE122FF69348FB37" box="[475,557,1169,1190]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZRC 2012.0787, holotype male (16.3×12.8 mm), Guam; C,
<taxonomicName id="80AE568EFF94FFCCFB622FF6945EFB37" box="[1195,1339,1168,1190]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF94FFCCFB622FF6945EFB37" box="[1195,1339,1168,1190]" italics="true" pageId="11" pageNumber="12">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF94FFCCFA8A2FF694F0FB37" bold="true" box="[1347,1429,1169,1190]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="6EE94C64FF94FFCCFA8A2FF694F0FB37" box="[1347,1429,1169,1190]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS 7028-001, holotype male (16.8×12.7 mm), Taiwan; D,
<taxonomicName id="80AE568EFF94FFCCFCD02FD792FDFB54" box="[793,920,1199,1221]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF94FFCCFCD02FD792FDFB54" box="[793,920,1199,1221]" italics="true" pageId="11" pageNumber="12">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF94FFCCFC692FC89294FB55" bold="true" box="[928,1009,1199,1220]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="6EE94C64FF94FFCCFC692FC89294FB55" box="[928,1009,1199,1220]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I.
</paragraph>
</caption>
<paragraph id="47112D0DFF94FFCCFF0E2E779414FA4D" blockId="11.[151,1437,1296,2004]" pageId="11" pageNumber="12">Walking legs relatively short, broad, flattened; second, third pairs sub-equal, longer than other walking legs. Without setae except for scattering of short dark bristles on dactyli, ventral face of propodi. Dorsal margins of meri with blunt subdistal shoulder, otherwise unarmed; outer surfaces of meri with minutely granular transverse striae. P4 merus c. 2.3 times longer than wide; P5 merus relatively narrower than P4 merus, c. 2.6 times longer. Carpi with 2 accessory carinae on outer surface but not strongly marked. P4 propodus c.2.8 times longer than wide; P5 propodus c. 2.7 times. Dactyli 0.8 times length of propodi, slightly recurved, terminating in acute chitinous tip.</paragraph>
<paragraph id="47112D0DFF94FFCCFF0E2E8F95A1F925" blockId="11.[151,1437,1296,2004]" pageId="11" pageNumber="12">
Thoracic sternites (
<figureCitation id="DF953188FF94FFCCFE6B2E8F9082F991" box="[418,487,1512,1536]" captionStart="FIGURE 9" captionStartId="12.[151,250,1576,1599]" captionTargetBox="[151,1435,325,1554]" captionTargetId="figure@12.[151,1436,325,1555]" captionTargetPageId="12" captionText="FIGURE 9. Ventral views of thoracic sternum and anterior parts of male abdomens. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283290/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
F) smooth, mostly bare of setae except for few scattered bristles; abdominal cavity reaching to thick transverse setal fringe at junction of sternites 3, 4. Male abdomen (
<figureCitation id="DF953188FF94FFCCFB9E2D6B95CDF9B5" box="[1111,1192,1548,1572]" captionStart="FIGURE 12" captionStartId="16.[151,250,1217,1240]" captionTargetBox="[174,1420,278,1178]" captionTargetId="figure@16.[151,1435,265,1196]" captionTargetPageId="16" captionText="FIGURE 12. Chiromantes C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I. A, male abdomen (somites 3 6, telson); B, ventral view of left G 1; C, dorsal view of left G 1; D, ventral view of distal part of left G 1; E, dorsal view of distal part of left G 1. All structures denuded. Scales: A C = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/283293/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
A) moderately broad; telson broadly rounded apically, 1.1 times longer than wide, slightly longer than somite 6; somite 6 c. 1.8 times wider than long, lateral margins divergent, slightly concave over distal half, but more subparallel over proximal half; somites 35 trapezoidal, lateral margins of somites 4, 5 relatively straight, lateral margins of somite 3 convex; somite 3 width 3.0 times basal width of telson; somites 1, 2 transversely, longitudinally narrow.
</paragraph>
<paragraph id="47112D0DFF94FFCCFF0E2DA790F0F8B0" blockId="11.[151,1437,1296,2004]" pageId="11" pageNumber="12">
G1 (
<figureCitation id="DF953188FF94FFCCFF302DA7902CF949" box="[249,329,1728,1752]" captionStart="FIGURE 12" captionStartId="16.[151,250,1217,1240]" captionTargetBox="[174,1420,278,1178]" captionTargetId="figure@16.[151,1435,265,1196]" captionTargetPageId="16" captionText="FIGURE 12. Chiromantes C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I. A, male abdomen (somites 3 6, telson); B, ventral view of left G 1; C, dorsal view of left G 1; D, ventral view of distal part of left G 1; E, dorsal view of distal part of left G 1. All structures denuded. Scales: A C = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/283293/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
BE) relatively slender, straight, weakly tapering to prominent subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process long, with slender narrow apex, dorsal margin slightly sinuous. G2 short, as for the genus.
</paragraph>
<paragraph id="47112D0DFF94FFCCFF0E2C4A9397F8F9" blockId="11.[151,1437,1296,2004]" pageId="11" pageNumber="12">
<emphasis id="75DAF11FFF94FFCCFF0E2C4A904CF8D4" bold="true" box="[199,297,1837,1861]" pageId="11" pageNumber="12">Females</emphasis>
. Chelipeds slightly smaller, less robust, lacking raised granular row on inner face of chela. Vulval morphology without clear species-specific characters.
</paragraph>
<paragraph id="47112D0DFF94FFCBFF0E2C139001FE8D" blockId="11.[151,1437,1296,2004]" lastBlockId="12.[151,1436,151,284]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="75DAF11FFF94FFCCFF0E2C139009F81C" bold="true" box="[199,364,1908,1933]" pageId="11" pageNumber="12">Colour in life</emphasis>
. Colour photographs taken from fresh specimens are shown in
<figureCitation id="DF953188FF94FFCCFB9B2C1395FFF81D" box="[1106,1178,1908,1932]" captionStart="FIGURE 2" captionStartId="2.[151,246,1309,1332]" captionTargetBox="[151,1435,153,1287]" captionTargetId="figure@2.[151,1436,153,1288]" captionTargetPageId="2" captionText="FIGURE 2. Colours in life. Chiromantes garfunkel sp. nov. A, ZRC 2012.0782, paratype female (11.1 × 8.3 mm), Flying Fish Cove; B D, QM-W 29172, holotype male (17.0 × 13.0 mm), Greta Beach; E, ZRC 2012.0783, paratype ovig. female (11.4 × 8.6 mm), Flying Fish Cove; F, ZRC 2012.0786, paratype female (11.6 × 8.6 mm), Lily Beach. All from Christmas I .. Photographs by H. H. Tan." httpUri="https://zenodo.org/record/283283/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
. Carapace and legs of adults rich, dark purple to maroon; without prominent speckling or splotching. Legs uniform in colour, without transverse banding. Chelipeds relatively uniform in colour, not clearly darker dorsally; off-white to white porcelain in adults, markedly contrasting with purple carapace. Ocular peduncles same as carapace in colour; corneas bright yellow (
<figureCitation id="DF953188FF93FFCBFF3F2BDB9058FF45" box="[246,317,188,212]" captionStart="FIGURE 2" captionStartId="2.[151,246,1309,1332]" captionTargetBox="[151,1435,153,1287]" captionTargetId="figure@2.[151,1436,153,1288]" captionTargetPageId="2" captionText="FIGURE 2. Colours in life. Chiromantes garfunkel sp. nov. A, ZRC 2012.0782, paratype female (11.1 × 8.3 mm), Flying Fish Cove; B D, QM-W 29172, holotype male (17.0 × 13.0 mm), Greta Beach; E, ZRC 2012.0783, paratype ovig. female (11.4 × 8.6 mm), Flying Fish Cove; F, ZRC 2012.0786, paratype female (11.6 × 8.6 mm), Lily Beach. All from Christmas I .. Photographs by H. H. Tan." httpUri="https://zenodo.org/record/283283/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
AD). Female specimens from more exposed supralittoral areas behind beaches can be paler in coloration, with the carapace and legs beige with streaks of purple and brown (
<figureCitation id="DF953188FF93FFCBFBCA2BB89523FF66" box="[1027,1094,223,247]" captionStart="FIGURE 2" captionStartId="2.[151,246,1309,1332]" captionTargetBox="[151,1435,153,1287]" captionTargetId="figure@2.[151,1436,153,1288]" captionTargetPageId="2" captionText="FIGURE 2. Colours in life. Chiromantes garfunkel sp. nov. A, ZRC 2012.0782, paratype female (11.1 × 8.3 mm), Flying Fish Cove; B D, QM-W 29172, holotype male (17.0 × 13.0 mm), Greta Beach; E, ZRC 2012.0783, paratype ovig. female (11.4 × 8.6 mm), Flying Fish Cove; F, ZRC 2012.0786, paratype female (11.6 × 8.6 mm), Lily Beach. All from Christmas I .. Photographs by H. H. Tan." httpUri="https://zenodo.org/record/283283/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
E, F). Their eyes, however, are still bright yellow.
</paragraph>
</subSubSection>
<caption id="13D17D85FF93FFCBFF5E2D4F951AF928" httpUri="https://zenodo.org/record/283290/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1435,325,1554]" targetPageId="12">
<paragraph id="47112D0DFF93FFCBFF5E2D4F951AF928" blockId="12.[151,1436,1576,1721]" pageId="12" pageNumber="13">
<emphasis id="75DAF11FFF93FFCBFF5E2D4F9077F9AF" bold="true" box="[151,274,1576,1599]" pageId="12" pageNumber="13">FIGURE 9.</emphasis>
Ventral views of thoracic sternum and anterior parts of male abdomens. A,
<taxonomicName id="80AE568EFF93FFCBFBD82D4F94FDF9AF" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[1041,1432,1576,1598]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF93FFCBFBD82D4F95F6F9AF" box="[1041,1171,1576,1598]" italics="true" pageId="12" pageNumber="13">Chiromantes</emphasis>
<emphasis id="75DAF11FFF93FFCBFB502D4F946FF9AF" box="[1177,1290,1576,1598]" italics="true" pageId="12" pageNumber="13">obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, UFUF-14837, male (17.7×13.2 mm), Hawaiian Is.; B,
<taxonomicName id="80AE568EFF93FFCBFD192D2F9296F9CC" authority="(Dana, 1851)" authorityName="Dana" authorityYear="1851" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[720,1011,1607,1629]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF93FFCBFD192D2F9204F9CC" box="[720,865,1607,1629]" italics="true" pageId="12" pageNumber="13">C. obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, ZRC 2002.0220, male (19.7×14.7 mm), Hawaiian Is.; C,
<taxonomicName id="80AE568EFF93FFCBFE8D2D0090F5F9ED" box="[324,400,1638,1660]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF93FFCBFE8D2D0090F5F9ED" box="[324,400,1638,1660]" italics="true" pageId="12" pageNumber="13">C. silus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF93FFCBFE5E2D01908DF9EA" bold="true" box="[407,488,1638,1659]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="6EE94C64FF93FFCBFE5E2D01908DF9EA" box="[407,488,1638,1659]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZRC 2012.0787, holotype male (16.3×12.8 mm), Guam; D,
<taxonomicName id="80AE568EFF93FFCBFB972D009589F9ED" box="[1118,1260,1638,1660]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF93FFCBFB972D009589F9ED" box="[1118,1260,1638,1660]" italics="true" pageId="12" pageNumber="13">C. leptomerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF93FFCBFB3D2D019420F9EA" bold="true" box="[1268,1349,1638,1659]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="6EE94C64FF93FFCBFB3D2D019420F9EA" box="[1268,1349,1638,1659]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS 7028-001, holotype male (16.8×12.7 mm), Taiwan; E,
<taxonomicName id="80AE568EFF93FFCBFD1C2DE29204F90A" box="[725,865,1669,1691]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF93FFCBFD1C2DE29204F90A" box="[725,865,1669,1691]" italics="true" pageId="12" pageNumber="13">C. eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF93FFCBFCA32DE292DBF90B" bold="true" box="[874,958,1669,1690]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="6EE94C64FF93FFCBFCA32DE292DBF90B" box="[874,958,1669,1690]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMNS-7028-002, holotype male (18.7×14.8 mm), Taiwan; F,
<taxonomicName id="80AE568EFF93FFCBFE8D2DC390A7F928" box="[324,450,1699,1721]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF93FFCBFE8D2DC390A7F928" box="[324,450,1699,1721]" italics="true" pageId="12" pageNumber="13">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF93FFCBFE012DC3937CF928" bold="true" box="[456,537,1700,1721]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="6EE94C64FF93FFCBFE012DC3937CF928" box="[456,537,1700,1721]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I.
</paragraph>
</caption>
<subSubSection id="0FB47E86FF93FFCAFF0E2D869454FBA3" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
<paragraph id="47112D0DFF93FFCBFF0E2D8692E2F865" blockId="12.[151,1437,1760,2037]" pageId="12" pageNumber="13">
<emphasis id="75DAF11FFF93FFCBFF0E2D869051F968" bold="true" box="[199,308,1761,1785]" pageId="12" pageNumber="13">Remarks</emphasis>
. This species was first recorded from
<collectingCountry id="3FB96D9DFF93FFCBFD242D8792C8F969" box="[749,941,1760,1784]" name="Christmas Island" pageId="12" pageNumber="13">Christmas Island</collectingCountry>
by Balss (1934) as
<taxonomicName id="80AE568EFF93FFCBFB5D2D86941EF968" box="[1172,1403,1761,1785]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF93FFCBFB5D2D86941EF968" box="[1172,1403,1761,1785]" italics="true" pageId="12" pageNumber="13">Sesarma obtusifrons</emphasis>
</taxonomicName>
. It was subsequently listed by Gibson-Hill (1947a, b) and Tweedie (1947) who reported on comparisons with specimens from the Hawaiian Islands made by F.A. Chace (in litt. to M. Tweedie). Chace noted (Tweedie, 1947: 34) that the Christmas I. specimens had more slender legs but felt that it was within the possible range of variation. It was pictured by Hicks
<emphasis id="75DAF11FFF93FFCBFE752C159092F818" box="[444,503,1905,1929]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(1984: 65), who gave a short account and gave it the name “Yellow-eyed Crab” (
<taxonomicName id="80AE568EFF93FFCBFF692CF290E3F83C" box="[160,390,1941,1965]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF93FFCBFF692CF290E3F83C" box="[160,390,1941,1965]" italics="true" pageId="12" pageNumber="13">Sesarma obtusifrons</emphasis>
</taxonomicName>
) because “Bright yellow eyes peering from a pock-mark in the seacliff are all you normally see of this shy little crab”. They further commented: “Yellow-eyed crabs live high on the seacliff, beyond tidal or salt spray influences, and have been recorded all around the coast.”
</paragraph>
<caption id="13D17D85FF92FFCAFF5E29369518FD17" httpUri="https://zenodo.org/record/283291/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1436,193,571]" targetPageId="13">
<paragraph id="47112D0DFF92FFCAFF5E29369518FD17" blockId="13.[151,1436,593,646]" pageId="13" pageNumber="14">
<emphasis id="75DAF11FFF92FFCAFF5E29369043FDF6" bold="true" box="[151,294,593,616]" pageId="13" pageNumber="14">FIGURE 10.</emphasis>
Lateral carapace views. A,
<taxonomicName id="80AE568EFF92FFCAFDAB2936929BFDF6" authority="(Dana, 1851)" authorityName="Dana" authorityYear="1851" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[610,1022,593,615]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF92FFCAFDAB29369207FDF6" box="[610,866,593,615]" italics="true" pageId="13" pageNumber="14">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
, UFUF-14837, male (17.7×13.2 mm), Hawaiian Is.; B,
<taxonomicName id="80AE568EFF92FFCAFE8B291690A5FD17" box="[322,448,624,646]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF92FFCAFE8B291690A5FD17" box="[322,448,624,646]" italics="true" pageId="13" pageNumber="14">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF92FFCAFE0F29179372FD14" bold="true" box="[454,535,624,645]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="6EE94C64FF92FFCAFE0F29179372FD14" box="[454,535,624,645]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, QM-W29172, holotype male (17.0×13.0 mm), Christmas I.
</paragraph>
</caption>
<paragraph id="47112D0DFF92FFCAFF0E29D59454FBA3" blockId="13.[151,1437,690,1544]" pageId="13" pageNumber="14">
Tweedie (1947) considered the record of
<taxonomicName id="80AE568EFF92FFCAFD5F29D4921EFD5A" box="[662,891,691,715]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF92FFCAFD5F29D4921EFD5A" box="[662,891,691,715]" italics="true" pageId="13" pageNumber="14">Sesarma obtusifrons</emphasis>
</taxonomicName>
by De Man (1895, 1898) from Atjeh, Sumatera,
<collectingCountry id="3FB96D9DFF92FFCAFF5E29B09061FD7E" box="[151,260,727,751]" name="Indonesia" pageId="13" pageNumber="14">Indonesia</collectingCountry>
as being conspecific with his Christmas I. samples. We disagree and consider that the identity of these specimens must remain unresolved at this time. From De Mans figure (1898: pl. 29, fig. 31) the abdomen appears significantly narrower than in
<taxonomicName id="80AE568EFF92FFCAFE3C284793E1FCA6" box="[501,644,799,823]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="garfunkel">
<emphasis id="75DAF11FFF92FFCAFE3C284793E1FCA6" box="[501,644,799,823]" italics="true" pageId="13" pageNumber="14">C. garfunkel</emphasis>
</taxonomicName>
and the legs are relatively shorter, both characters that are useful in separating other species in the group. Furthermore, the live colouration was well described by De Man and this does not agree with the colour of the Christmas I. species. De Man (1895: 161165) described the colour as “The surface of the carapace has an olive-green base color [purple in
<taxonomicName id="80AE568EFF92FFCAFCBF28EC9560FC32" box="[886,1029,907,931]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="garfunkel">
<emphasis id="75DAF11FFF92FFCAFCBF28EC9560FC32" box="[886,1029,907,931]" italics="true" pageId="13" pageNumber="14">C. garfunkel</emphasis>
</taxonomicName>
] and transmits some symmetrically arranged dark brown patches of irregular shape. The lower frontal border, the orbital margins, and the lateral margins are yellowish, as are the depressions of the hepatic regions immediately behind the external orbital angle. The claws are yellowish, turning reddish-yellow dorsally, the legs are brownish in color with purple transverse bands.The numerous features that distinguish this species from its closest congeners are given in
<tableCitation id="0A2C18B6FF92FFCAFB1E2F7D944EFBA3" box="[1239,1323,1050,1074]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[159,1428,329,1954]" captionTargetPageId="5" captionText="TABLE 1. Summary of major differences between species of the Chiromantes obtusifrons complex." httpUri="http://table.plazi.org/id/13D17D85FF9AFFC2FF5E2BF095A0FF3E" pageId="13" pageNumber="14" tableUuid="13D17D85FF9AFFC2FF5E2BF095A0FF3E">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF92FFCAFF0E2F589557FB0E" pageId="13" pageNumber="14" type="etymology">
<paragraph id="47112D0DFF92FFCAFF0E2F589557FB0E" blockId="13.[151,1437,690,1544]" pageId="13" pageNumber="14">
<emphasis id="75DAF11FFF92FFCAFF0E2F589023FBC6" bold="true" box="[199,326,1087,1111]" pageId="13" pageNumber="14">Etymology</emphasis>
. Named for Art
<taxonomicName id="80AE568EFF92FFCAFDCD2F58931DFBC6" box="[516,632,1087,1111]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="17" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">Garfunkel</taxonomicName>
, who sang “Bright Eyes” for the soundtrack of the 1978 animated film
<emphasis id="75DAF11FFF92FFCAFF5E2F049033FBEA" box="[151,342,1123,1147]" italics="true" pageId="13" pageNumber="14">Watership Down</emphasis>
. Like the rabbit to which the song alludes, the new crab species also lives in holes, and is remarkable for its brightly shining eyes. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF92FFCAFF0E2FCD92E3F996" pageId="13" pageNumber="14" type="distribution">
<paragraph id="47112D0DFF92FFCAFF0E2FCD9290FB53" blockId="13.[151,1437,690,1544]" box="[199,1013,1194,1219]" pageId="13" pageNumber="14">
<emphasis id="75DAF11FFF92FFCAFF0E2FCD903CFB52" bold="true" box="[199,345,1194,1219]" pageId="13" pageNumber="14">Distribution</emphasis>
. Only known from
<collectingCountry id="3FB96D9DFF92FFCAFDFE2FCD9393FB53" box="[567,758,1194,1218]" name="Christmas Island" pageId="13" pageNumber="14">Christmas Island</collectingCountry>
, eastern Indian Ocean.
</paragraph>
<paragraph id="47112D0DFF92FFCAFF0E2FA892E3F996" blockId="13.[151,1437,690,1544]" pageId="13" pageNumber="14">
<emphasis id="75DAF11FFF92FFCAFF0E2FA8904EFB76" bold="true" box="[199,299,1231,1255]" pageId="13" pageNumber="14">Ecology.</emphasis>
Gibson-Hill (1947b: 44) noted that: “Yet another species,
<taxonomicName id="80AE568EFF92FFCAFC012FA895CBFB76" box="[968,1198,1231,1255]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF92FFCAFC012FA895CBFB76" box="[968,1198,1231,1255]" italics="true" pageId="13" pageNumber="14">Sesarma obtusifrons</emphasis>
</taxonomicName>
, is fairly plentiful in the pock-marks and crevices at the top of the sea cliff. It is particularly common in Flying Fish Cove and behind Greta and West White Beaches. It also occurs up on the shore terrace proper, at the base of the inland cliff, and during the rainy season can occasionally be found in the open. Females with spawn (a crab with a carapace width of
<quantity id="805680E8FF92FFCAFF7E2E389062FAE6" box="[183,263,1375,1399]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="14" unit="mm" value="18.0">18 mm</quantity>
carrying about 5500 eggs) are most frequent from January to April. The spawn is taken to the sea and deposited, at low tide, on the edges of the rock pools on the uncovered portions of the fringing reef.” George (1978) largely repeated the earlier notes made by Gibson-Hill (1947a, b). Hicks
<emphasis id="75DAF11FFF92FFCAFC2E2ECF9547FA2E" box="[999,1058,1447,1471]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(1984: 65) stated: “Yellow-eyed crabs live high on the seacliff, beyond tidal or salt spray influences, and have been recorded all around the coast.” Our personal observations agree entirely with these earlier authors.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,505 @@
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<paragraph id="47112D0DFF9EFFC6FF5E2A67937FFE8B" blockId="1.[151,654,222,282]" box="[151,538,256,282]" pageId="1" pageNumber="2">
<heading id="1C599A61FF9EFFC6FF5E2A67937FFE8B" bold="true" box="[151,538,256,282]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="75DAF11FFF9EFFC6FF5E2A67937FFE8B" bold="true" box="[151,538,256,282]" pageId="1" pageNumber="2">
Family
<taxonomicName id="80AE568EFF9EFFC6FF3F2A67937FFE8B" ID-CoL="7NL25" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[246,538,256,282]" class="Malacostraca" family="Sesarmidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Sesarmidae Dana, 1851</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="47112D0DFF9EFFC6FF5E2A209300FEF3" blockId="1.[151,769,327,387]" box="[151,613,327,354]" pageId="1" pageNumber="2">
<taxonomicName id="80AE568EFF9EFFC6FF5E2A209300FEF3" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[151,613,327,354]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9EFFC6FF5E2A2090A1FEF0" bold="true" box="[151,452,327,353]" italics="true" pageId="1" pageNumber="2">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
</paragraph>
<paragraph id="47112D0DFF9EFFC6FF5E2A0D9264FE13" blockId="1.[151,769,327,387]" box="[151,769,362,387]" pageId="1" pageNumber="2">
(
<figureCitation id="DF953188FF9EFFC6FF562A0D9189FE13" box="[159,236,362,386]" captionStart="FIGURE 1" captionStartId="1.[151,250,1544,1567]" captionTargetBox="[151,1435,806,1520]" captionTargetId="figure@1.[151,1436,806,1520]" captionTargetPageId="1" captionText="FIGURE 1. Colours in life. Chiromantes obtusifrons (Dana, 1851). A C, UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; D, UFUF- 14837, female (15.7 × 11.6 mm), Hawaiian Is. Photographs by G. Paulay." httpUri="https://zenodo.org/record/283282/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
,
<figureCitation id="DF953188FF9EFFC6FF332A0D906CFE13" box="[250,265,362,386]" captionStart="FIGURE 4" captionStartId="7.[151,250,1841,1864]" captionTargetBox="[151,1435,717,1822]" captionTargetId="figure@7.[151,1436,715,1823]" captionTargetPageId="7" captionText="FIGURE 4. Overall dorsal views. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; C, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; D, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; E, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283285/files/figure.png" pageId="1" pageNumber="2">4</figureCitation>
A, 5A, B, 6A, B, 7A, B, 8A, 9A, B, 10A, 11)
</paragraph>
<paragraph id="47112D0DFF9EFFC6FF5E2AD492CDFE76" blockId="1.[151,1436,434,610]" pageId="1" pageNumber="2">
<taxonomicName id="80AE568EFF9EFFC6FF5E2AD49379FE59" authority="Dana, 1851: 250" authorityName="Dana" authorityPageNumber="250" authorityYear="1851" box="[151,540,434,456]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9EFFC6FF5E2AD49003FE59" box="[151,358,434,456]" italics="true" pageId="1" pageNumber="2">Sesarma obtusifrons</emphasis>
Dana, 1851: 250
</taxonomicName>
; 1852: 355, pl. 22, fig. 9.—H. Milne Edwards 1853: 183.—De Man 1887: 644.—Lenz 1901: 472.—Titcomb
<emphasis id="75DAF11FFF9EFFC6FE612AB590B6FE76" box="[424,467,465,487]" italics="true" pageId="1" pageNumber="2">et al</emphasis>
. 1979: 367.—Godwin &amp; Bolick 2006: 39, 49.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF9EFFC6FF5E2A969063FDF3" pageId="1" pageNumber="2" type="description">
<paragraph id="47112D0DFF9EFFC6FF5E2A9690ACFDB5" blockId="1.[151,1436,434,610]" pageId="1" pageNumber="2">
<taxonomicName id="80AE568EFF9EFFC6FF5E2A96936DFD97" box="[151,520,496,518]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="obtusifrons" subGenus="Holometopus">
<emphasis id="75DAF11FFF9EFFC6FF5E2A96918BFD97" box="[151,238,497,518]" italics="true" pageId="1" pageNumber="2">Sesarma</emphasis>
(
<emphasis id="75DAF11FFF9EFFC6FF362A9790E3FD97" box="[255,390,496,518]" italics="true" pageId="1" pageNumber="2">Holometopus</emphasis>
)
<emphasis id="75DAF11FFF9EFFC6FE5E2A97936DFD97" box="[407,520,496,518]" italics="true" pageId="1" pageNumber="2">obtusifrons</emphasis>
</taxonomicName>
—Rathbun 1906: 840; 1907: 35.—Tesch 1917: 179.—Edmondson 1946: 306, fig. 183e; 1959: 185, figs. 13c, 17c.
</paragraph>
<paragraph id="47112D0DFF9EFFC6FF5E294A9063FDF3" blockId="1.[151,1436,434,610]" pageId="1" pageNumber="2">
<taxonomicName id="80AE568EFF9EFFC6FF5E294A90F6FDD2" box="[151,403,557,579]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9EFFC6FF5E294A90F6FDD2" box="[151,403,557,579]" italics="true" pageId="1" pageNumber="2">Chiromantes obtusifrons</emphasis>
</taxonomicName>
—Ng &amp; Liu 1999: 230.—Castro 2011: 120. “
<taxonomicName id="80AE568EFF9EFFC6FCBC294A9292FDD2" box="[885,1015,557,579]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="75DAF11FFF9EFFC6FCBC294A9292FDD2" box="[885,1015,557,579]" italics="true" pageId="1" pageNumber="2">Chiromantes</emphasis>
</taxonomicName>
<taxonomicName id="80AE568EFF9EFFC6FBC5294A9518FDD2" box="[1036,1149,557,579]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9EFFC6FBC5294A9518FDD2" box="[1036,1149,557,579]" italics="true" pageId="1" pageNumber="2">obtusifrons</emphasis>
</taxonomicName>
—Ng
<emphasis id="75DAF11FFF9EFFC6FB7729499590FDD2" box="[1214,1269,557,579]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
2008: 220, 224 (part).
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF9EFFC6FF5E29E9955FFD7F" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="47112D0DFF9EFFC6FF5E29E9955FFD7F" blockId="1.[151,1436,654,751]" pageId="1" pageNumber="2">
<emphasis id="75DAF11FFF9EFFC6FF5E29E990E7FD36" bold="true" box="[151,386,654,679]" pageId="1" pageNumber="2">Material Examined</emphasis>
. ZRC 2002.0220,
<specimenCount id="51A8E684FF9EFFC6FD9529E993D1FD37" box="[604,692,654,678]" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
(16.8×12.9, 19.7×
<quantity id="805680E8FF9EFFC6FC4229E99293FD37" box="[907,1014,654,678]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="2" unit="mm" value="14.7">14.7 mm</quantity>
), Malaikahana, Oahu, Hawaiian Is., C.M. Cooke
<emphasis id="75DAF11FFF9EFFC6FEE229D39001FD5A" box="[299,356,691,715]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
, 0 8.07.1916 [ex. BPBM-266].—UFUF-14837, male (17.7×
<quantity id="805680E8FF9EFFC6FBC029D59511FD5B" box="[1033,1140,690,714]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="1" pageNumber="2" unit="mm" value="13.2">13.2 mm</quantity>
), female (15.7×
<quantity id="805680E8FF9EFFC6FAEF29D594F5FD5B" box="[1318,1424,690,714]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="2" unit="mm" value="11.6">11.6 mm</quantity>
),
<geoCoordinate id="229A4BCAFF9EFFC6FF5E29B19071FD7F" box="[151,276,726,750]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="5" value="21.2833">21.2833°N</geoCoordinate>
,
<geoCoordinate id="229A4BCAFF9EFFC6FED629B190FCFD7F" box="[287,409,726,750]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="55" value="157.667">157.667°E</geoCoordinate>
, Coco Head, Oahu, Hawaiian Is., G. Paulay
<emphasis id="75DAF11FFF9EFFC6FC5829BF92ACFD7F" box="[913,969,726,751]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
, 10.2006.
</paragraph>
</subSubSection>
<caption id="13D17D85FF9EFFC6FF5E2D6F92BDF9AC" httpUri="https://zenodo.org/record/283282/files/figure.png" pageId="1" pageNumber="2" targetBox="[151,1435,806,1520]" targetPageId="1">
<paragraph id="47112D0DFF9EFFC6FF5E2D6F92BDF9AC" blockId="1.[151,1436,1544,1597]" pageId="1" pageNumber="2">
<emphasis id="75DAF11FFF9EFFC6FF5E2D6F9068F98F" bold="true" box="[151,269,1544,1567]" pageId="1" pageNumber="2">FIGURE 1</emphasis>
. Colours in life.
<taxonomicName id="80AE568EFF9EFFC6FE742D6E922CF98E" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[445,841,1545,1567]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9EFFC6FE742D6E93D2F98E" box="[445,695,1545,1567]" italics="true" pageId="1" pageNumber="2">Chiromantes obtusifrons</emphasis>
(Dana, 1851)
</taxonomicName>
. AC, UFUF-14837, male (17.7×13.2 mm), Hawaiian Is.; D, UFUF-14837, female (15.7×11.6 mm), Hawaiian Is. Photographs by G. Paulay.
</paragraph>
</caption>
<subSubSection id="0FB47E86FF9EFFC3FF0E2D0E92E8FABE" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="47112D0DFF9EFFC6FF0E2D0E919FF87B" blockId="1.[151,1436,1641,2026]" pageId="1" pageNumber="2">
<emphasis id="75DAF11FFF9EFFC6FF0E2D0E905FF913" bold="true" box="[199,314,1641,1666]" pageId="1" pageNumber="2">Diagnosis</emphasis>
. Carapace transversely rectangular, c. 1.3 times broader than long; dorsal carapace, lateral branchial regions weakly convex, not prominently swollen; exorbital tooth with outer margin broadly convex so greatest carapace width clearly posterior to exorbital tooth; front c. 0.65 times carapace width, margin broadly convex in frontal view with medial part relatively straight in dorsal view, beaded with row of small but distinct granules, with pair of low lateral swellings behind margin; supraorbital margin entire, semicircular; dorsal surface of cheliped carpus covered in small but conspicuous granules; walking legs relatively short; P4 merus c. 2.2 times longer than wide; P5 merus c. 2.3 times longer; P5 propodus c. 2.2 times longer; male abdomen moderately broad; somite 6 with distolateral margins strongly divergent, relatively straight over distal two-thirds; somite 3 width 2.9 times basal width of telson; G1 (
<figureCitation id="DF953188FF9EFFC6FE0E2CED9370F833" box="[455,533,1930,1954]" captionStart="FIGURE 11" captionStartId="14.[151,250,1878,1901]" captionTargetBox="[161,1420,309,1846]" captionTargetId="figure@14.[151,1436,301,1857]" captionTargetPageId="14" captionText="FIGURE 11. Chiromantes obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is .. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, ventral view of distal part of left G 1; G, dorsal view of distal part of left G 1. All structures denuded. Scales: A E = 1.0 mm; F, G = 0.5 mm." httpUri="https://zenodo.org/record/283292/files/figure.png" pageId="1" pageNumber="2">Fig. 11</figureCitation>
DG) relatively slender, weakly tapering to obtusely angled subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process long, with slender narrow apex, dorsal margin slightly concave.
</paragraph>
<caption id="13D17D85FF9DFFC5FF5E2E7A9432FAE0" httpUri="https://zenodo.org/record/283283/files/figure.png" pageId="2" pageNumber="3" targetBox="[151,1435,153,1287]" targetPageId="2">
<paragraph id="47112D0DFF9DFFC5FF5E2E7A9432FAE0" blockId="2.[151,1436,1309,1393]" pageId="2" pageNumber="3">
<emphasis id="75DAF11FFF9DFFC5FF5E2E7A906DFAA2" bold="true" box="[151,264,1309,1332]" pageId="2" pageNumber="3">FIGURE 2</emphasis>
. Colours in life.
<taxonomicName id="80AE568EFF9DFFC5FE652E7993EEFAA5" box="[428,651,1310,1332]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF9DFFC5FE652E7993EEFAA5" box="[428,651,1310,1332]" italics="true" pageId="2" pageNumber="3">Chiromantes garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF9DFFC5FD5B2E7993BAFAA2" bold="true" box="[658,735,1310,1331]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="6EE94C64FF9DFFC5FD5B2E7993BAFAA2" box="[658,735,1310,1331]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, ZRC 2012.0782, paratype female (11.1×8.3 mm), Flying Fish Cove; BD, QM-W29172, holotype male (17.0×13.0 mm), Greta Beach; E, ZRC 2012.0783, paratype ovig. female (11.4×8.6 mm), Flying Fish Cove; F, ZRC 2012.0786, paratype female (11.6×8.6 mm), Lily Beach. All from Christmas I.. Photographs by H. H. Tan.
</paragraph>
</caption>
<paragraph id="47112D0DFF9DFFC5FF0E2EF19513F87E" blockId="2.[151,1436,1430,2031]" pageId="2" pageNumber="3">
<emphasis id="75DAF11FFF9DFFC5FF0E2EF19011FA3E" bold="true" box="[199,372,1430,1455]" pageId="2" pageNumber="3">Redescription.</emphasis>
Carapace transversely rectangular, c. 1.3 times (range 1.301.34) broader than long; surface bare, lacking setal tufts; anterior half conspicuously granular, granules become minute posterior, arranged into inconspicuous striae; mesogastric regions well defined; lateral carapace surface generally without obvious discrete oblique striae, except for strong, concave epibranchial sulcus beginning just behind exorbital tooth. Dorsal carapace, lateral branchial regions weakly convex. Exorbital tooth triangular, pointed, outer margin broadly convex so that greatest carapace width clearly posterior to exorbital tooth, margin not constricted behind (except on one side of larger male, but probably result of damage) so that entire lateral border appears convex, no trace of second anterolateral tooth. Postfrontal lobes well demarcated but relatively low, rounded, separated by broad grooves, median lobes similar in width to lateral lobes. Front c. 0.65 times carapace width, markedly deflexed, margin broadly convex in frontal view, but medial part relatively straight in dorsal view; laterally triangular, bluntly pointed; frontal margin emarginated, beaded with row of small but distinct granules; surface concave behind rim, but with pair of low lateral swellings behind frontal margin. Supraorbital margin entire, semicircular but oblique medial section somewhat straight; secondary rim formed either side making smooth, broadly triangular, deflexed central plate behind ocular peduncle. Eye not extending beyond exorbital tooth. Frontal plate protruding as shelf; medial septum broad, largely covering anterior half of anterior half of epistome; antennae, antennules much reduced in size, lodged under overhanging front; basal antennular segment not much swollen; antennal, antennular basal segments adjacent, not separated by septum. Antennal flagellum distinctively short, entering orbit.
</paragraph>
<caption id="13D17D85FF9CFFC4FF5E2C59956AF87F" httpUri="https://zenodo.org/record/283284/files/figure.png" pageId="3" pageNumber="4" targetBox="[174,1416,179,1840]" targetPageId="3">
<paragraph id="47112D0DFF9CFFC4FF5E2C59956AF87F" blockId="3.[151,1436,1854,2030]" pageId="3" pageNumber="4">
<emphasis id="75DAF11FFF9CFFC4FF5E2C599076F8C5" bold="true" box="[151,275,1854,1877]" pageId="3" pageNumber="4">FIGURE 3.</emphasis>
Colours in life. AC,
<taxonomicName id="80AE568EFF9CFFC4FE3E2C5893CBF8C4" box="[503,686,1855,1877]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF9CFFC4FE3E2C5893CBF8C4" box="[503,686,1855,1877]" italics="true" pageId="3" pageNumber="4">Chiromantes silus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF9CFFC4FD7C2C589263F8C5" bold="true" box="[693,774,1855,1876]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="6EE94C64FF9CFFC4FD7C2C589263F8C5" box="[693,774,1855,1876]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Guam; A, ZRC 2000.0573, female (15.2×11.2 mm); B, C, ZRC 2001.0742, male (15.7×11.7 mm). DH,
<taxonomicName id="80AE568EFF9CFFC4FD9D2C399236F8E5" box="[596,851,1886,1908]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="75DAF11FFF9CFFC4FD9D2C399236F8E5" box="[596,851,1886,1908]" italics="true" pageId="3" pageNumber="4">Chiromantes leptomerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF9CFFC4FC962C3992D3F8E2" box="[863,950,1886,1907]" pageId="3" pageNumber="4" rank="species">
<emphasis id="75DAF11FFF9CFFC4FC962C3992CAF8E2" bold="true" box="[863,943,1886,1907]" pageId="3" pageNumber="4">sp. nov</emphasis>
.
</taxonomicNameLabel>
, D, RUMF-ZC-2104, male (17.5×13.7 mm), Muigah, Miyako Island, Ryukyu Islands, Japan; E, RUMF-ZC-2076, male (19.7×14.6 mm), Honba Coast, Minamidaito Island, Japan; F, male, southern Taiwan, specimen not kept; H, NMNS, male (17.1×13.0 mm), Siatanzai, Kenting, Pingtung County, Taiwan. IJ,
<taxonomicName id="80AE568EFF9CFFC4FED42CDD9377F841" box="[285,530,1978,2000]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="eurymerus" status="sp. nov.">
<emphasis id="75DAF11FFF9CFFC4FED42CDD9377F841" box="[285,530,1978,2000]" italics="true" pageId="3" pageNumber="4">Chiromantes eurymerus</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF9CFFC4FDD32CDD9309F85E" bold="true" box="[538,620,1978,1999]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="6EE94C64FF9CFFC4FDD32CDD9309F85E" box="[538,620,1978,1999]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, I, NMNS, male (17.5×13.4 mm); J, NMNS, male (16.7×12.3 mm), Siatanzai, Kenting, Taiwan. Photographs: B, C by H.-C. Liu; DE by Y. Fujita; FJ by H.-T. Shih.
</paragraph>
</caption>
<paragraph id="47112D0DFF9BFFC3FF0E2BF0952DFE8D" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">Third maxilliped ischium subequal to merus in length; ischium with shallow longitudinal, curved, median sulcus. Inner margins of merus, ischium with long setae, proximal outer margin of ischium, base of exopod with dense setae; outer margin of merus with short scattered setae only. Exopod slender, hidden behind ischium, merus except near base, tip reaching half length of outer margin of merus, flagellum long, slender.</paragraph>
<paragraph id="47112D0DFF9BFFC3FF0E2A40950DFD36" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
Chelipeds subequal, robust. Merus trihedral; posterior border broadly convex, emarginated, minutely granular, without indication of subdistal spine or lobe; inner anterior border with conspicuous row of pointed granules, broadly triangular; outer surface broadly convex, with fine transverse striae but appearing almost smooth. Carpus subquadrilateral, inner angle moderately produced, apically granular; outer margin, dorsal surface covered in small but conspicuous granules. Dorsal surface of palm slightly striated along superior margin, with scattering of moderate sized granules, otherwise smooth, punctate. Palm outer surface broadly rounded, smooth; no indication of subventral longitudinal ridge. Inner face of palm of males smooth except for curved projecting crest of large tubercles posterior to gape (
<figureCitation id="DF953188FF9BFFC3FEA8294390C6FDAD" box="[353,419,548,572]" captionStart="FIGURE 6" captionStartId="9.[151,250,1537,1560]" captionTargetBox="[151,1435,193,1515]" captionTargetId="figure@9.[151,1436,193,1516]" captionTargetPageId="9" captionText="FIGURE 6. Inner views of chelae. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283287/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
A, B). Cutting margin of fixed finger typically with 3 or 4 blunt teeth, proximal 3 similar in size, distal-most larger, rounded, conical, placed medially; dactylus with 2 teeth proximally, distalmost large, third much smaller, blunt tooth slightly distal of medial tooth of ventral finger. Dorsal surface of dactylus smooth. Fingers with chitinous tips; adult males with narrow but distinct gape when fingers closed.
</paragraph>
<paragraph id="47112D0DFF9BFFC3FF0E29D3904CFC35" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">Walking legs relatively short, broad, flattened; second, third pairs sub-equal, longer than other walking legs. Without setae except for scattering of short dark bristles on dactyli, ventral face of propodi. Dorsal margin of meri with blunt subdistal shoulder, otherwise unarmed; outer surfaces of meri with minutely granular transverse striae. P4 merus c. 2.2 times longer than wide; P5 merus c. 2.3 times longer. Carpi with 2 well-developed accessory carinae on outer surface. Dorsal surfaces of carpi, propodi conspicuously granular. P4 propodus c. 2.1 times longer than wide; P5 propodus c. 2.2 times. Dactyli 0.8 times length of propodi, slightly recurved, terminating in acute chitinous tip.</paragraph>
<paragraph id="47112D0DFF9BFFC3FF0E28C8906CFB78" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
Thoracic sternites (
<figureCitation id="DF953188FF9BFFC3FE5428C890BAFC56" box="[413,479,943,967]" captionStart="FIGURE 9" captionStartId="12.[151,250,1576,1599]" captionTargetBox="[151,1435,325,1554]" captionTargetId="figure@12.[151,1436,325,1555]" captionTargetPageId="12" captionText="FIGURE 9. Ventral views of thoracic sternum and anterior parts of male abdomens. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is.; C, C. silus sp. nov., ZRC 2012.0787, holotype male (16.3 × 12.8 mm), Guam; D, C. leptomerus sp. nov., NMNS 7028 - 001, holotype male (16.8 × 12.7 mm), Taiwan; E, C. eurymerus sp. nov., NMNS- 7028 - 002, holotype male (18.7 × 14.8 mm), Taiwan; F, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283290/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
A, B) smooth, mostly bare of setae except for few scattered bristles; abdominal cavity reaching to thick transverse setal fringe at junction of sternites 3, 4. Male abdomen (
<figureCitation id="DF953188FF9BFFC3FB9028B395CCFC7D" box="[1113,1193,980,1004]" captionStart="FIGURE 11" captionStartId="14.[151,250,1878,1901]" captionTargetBox="[161,1420,309,1846]" captionTargetId="figure@14.[151,1436,301,1857]" captionTargetPageId="14" captionText="FIGURE 11. Chiromantes obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is .. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, ventral view of distal part of left G 1; G, dorsal view of distal part of left G 1. All structures denuded. Scales: A E = 1.0 mm; F, G = 0.5 mm." httpUri="https://zenodo.org/record/283292/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
C) moderately broad; telson broadly rounded apically, 1.1 times longer than wide, slightly longer than somite 6 (1.15 times); somite 6 c. 2.0 times wider than long, distolateral margins strongly divergent, relatively straight over distal two-thirds; somites 35 trapezoidal, lateral margins of somites 4, 5 relatively straight, lateral margins of somite 3 convex; somite 3 width 2.9 times basal width of telson; somites 1, 2 transversely narrower, longitudinally narrow. G1 (
<figureCitation id="DF953188FF9BFFC3FAC12F03943FFBED" box="[1288,1370,1124,1148]" captionStart="FIGURE 11" captionStartId="14.[151,250,1878,1901]" captionTargetBox="[161,1420,309,1846]" captionTargetId="figure@14.[151,1436,301,1857]" captionTargetPageId="14" captionText="FIGURE 11. Chiromantes obtusifrons (Dana, 1851), ZRC 2002.0220, male (19.7 × 14.7 mm), Hawaiian Is .. A, left third maxilliped; B, left side of anterior thoracic sternum; C, male abdomen (somites 3 6, telson); D, ventral view of left G 1; E, dorsal view of left G 1; F, ventral view of distal part of left G 1; G, dorsal view of distal part of left G 1. All structures denuded. Scales: A E = 1.0 mm; F, G = 0.5 mm." httpUri="https://zenodo.org/record/283292/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
DG) relatively slender, almost straight, weakly tapering to obtusely angled subdistal shoulder; distally slender, strongly bent to 45° angle; terminal process long, with slender narrow apex, dorsal margin slightly concave. G2 short, as for the genus.
</paragraph>
<paragraph id="47112D0DFF9BFFC3FF0E2F9392E8FABE" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<emphasis id="75DAF11FFF9BFFC3FF0E2F93904CFA9D" bold="true" box="[199,297,1268,1292]" pageId="4" pageNumber="5">Females</emphasis>
. Chelipeds slightly smaller, less robust than in males; lacking raised granular row on inner face of chela. Vulval morphology without clear species-specific characters.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF9BFFC3FF0E2E5C9334FA75" pageId="4" pageNumber="5" type="description">
<paragraph id="47112D0DFF9BFFC3FF0E2E5C9334FA75" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<emphasis id="75DAF11FFF9BFFC3FF0E2E5C9001FAC5" bold="true" box="[199,356,1339,1364]" pageId="4" pageNumber="5">Colour in life</emphasis>
. Colour photographs taken from fresh specimens are given in
<figureCitation id="DF953188FF9BFFC3FBD22E5B9539FAC5" box="[1051,1116,1340,1364]" captionStart="FIGURE 1" captionStartId="1.[151,250,1544,1567]" captionTargetBox="[151,1435,806,1520]" captionTargetId="figure@1.[151,1436,806,1520]" captionTargetPageId="1" captionText="FIGURE 1. Colours in life. Chiromantes obtusifrons (Dana, 1851). A C, UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; D, UFUF- 14837, female (15.7 × 11.6 mm), Hawaiian Is. Photographs by G. Paulay." httpUri="https://zenodo.org/record/283282/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
. Background colour of carapace, legs fawn to bluish gray, covered in fine darker speckling with scattering of slightly larger irregular spots. Legs with darker, broad transverse bands that are more noticeable on the carpi and meri. Chelipeds dorsally similar in colour to carapace and legs but becoming greyish white in frontal and ventral view. Ocular peduncles and corneas pale, similar to carapace in colour.
</paragraph>
</subSubSection>
<subSubSection id="0FB47E86FF9BFFC3FF0E2E979212F805" pageId="4" pageNumber="5" type="discussion">
<paragraph id="47112D0DFF9BFFC3FF0E2E9795E5F8B6" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<emphasis id="75DAF11FFF9BFFC3FF0E2E979051F999" bold="true" box="[199,308,1520,1544]" pageId="4" pageNumber="5">Remarks</emphasis>
.
<taxonomicName id="80AE568EFF9BFFC3FE842E979309F999" box="[333,620,1520,1544]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9BFFC3FE842E979309F999" box="[333,620,1520,1544]" italics="true" pageId="4" pageNumber="5">Chiromantes obtusifrons</emphasis>
</taxonomicName>
was originally described by Dana (1851) from Maui, Sandwich Archipelago [= Hawaiian Is.], and since then it has been reported from a number of Indo-West Pacific localities stretching as far west as
<collectingCountry id="3FB96D9DFF9BFFC3FE792D50930AF9DE" box="[432,623,1591,1615]" name="Christmas Island" pageId="4" pageNumber="5">Christmas Island</collectingCountry>
in the Indian Ocean (Davie 2002). Like most of Danas specimens, the
<typeStatus id="981593AFFF9BFFC3FF5E2D3A91A2F9E4" box="[151,199,1629,1653]" pageId="4" pageNumber="5">type</typeStatus>
material is now considered lost, and so, to be certain of its identity we obtained topotypic material from the Hawaiian Is. for redescription and refiguring. Although we have concluded that
<taxonomicName id="80AE568EFF9BFFC3FBED2DE795A6F909" box="[1060,1219,1664,1688]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" sensu="lato" species="obtusifrons">
<emphasis id="75DAF11FFF9BFFC3FBED2DE795A6F909" box="[1060,1219,1664,1688]" italics="true" pageId="4" pageNumber="5">C. obtusifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF9BFFC3FB022DE7945EF906" box="[1227,1339,1663,1687]" pageId="4" pageNumber="5" sensu="lato">sensu lato</taxonomicNameLabel>
consists of at least five sibling species, we feel confident that each can be reliably recognised based on differences in the morphological characters given here for the Hawaiian populations. We therefore do not consider it necessary to designate a
<typeStatus id="981593AFFF9BFFC3FEEA2D8A9019F894" box="[291,380,1773,1797]" pageId="4" pageNumber="5" type="neotype">neotype</typeStatus>
at this time, especially because it appears that most of the species have relatively narrow distributions, and this will make it unlikely that there will be future taxonomic confusion.
</paragraph>
<paragraph id="47112D0DFF9BFFC3FF0E2C539212F805" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<taxonomicName id="80AE568EFF9BFFC3FF0E2C5390B8F8DD" box="[199,477,1844,1868]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9BFFC3FF0E2C5390B8F8DD" box="[199,477,1844,1868]" italics="true" pageId="4" pageNumber="5">Chiromantes obtusifrons</emphasis>
</taxonomicName>
can be identified using the characters given in
<tableCitation id="0A2C18B6FF9BFFC3FBC02C53953AF8DD" box="[1033,1119,1844,1868]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[159,1428,329,1954]" captionTargetPageId="5" captionText="TABLE 1. Summary of major differences between species of the Chiromantes obtusifrons complex." httpUri="http://table.plazi.org/id/13D17D85FF9AFFC2FF5E2BF095A0FF3E" pageId="4" pageNumber="5" tableUuid="13D17D85FF9AFFC2FF5E2BF095A0FF3E">Table 1</tableCitation>
. Of all the members in the species-complex it has the most conspicuously granular carapace (over anterior half), frontal margin and claws; and the carapace is more flattened and less convex longitudinally.
</paragraph>
</subSubSection>
<caption id="13D17D85FF9AFFC2FF5E2BF095A0FF3E" ID-Table-UUID="13D17D85FF9AFFC2FF5E2BF095A0FF3E" box="[151,1221,151,175]" httpUri="http://table.plazi.org/id/13D17D85FF9AFFC2FF5E2BF095A0FF3E" pageId="5" pageNumber="6" targetBox="[159,1428,329,1954]" targetIsTable="true" targetPageId="5">
<paragraph id="47112D0DFF9AFFC2FF5E2BF095A0FF3E" blockId="5.[151,1221,151,175]" box="[151,1221,151,175]" pageId="5" pageNumber="6">
<emphasis id="75DAF11FFF9AFFC2FF5E2BF09066FF3E" bold="true" box="[151,259,151,175]" pageId="5" pageNumber="6">TABLE 1</emphasis>
. Summary of major differences between species of the
<taxonomicName id="80AE568EFF9AFFC2FC9D2BFF953EFF3E" box="[852,1115,152,175]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF9AFFC2FC9D2BFF953EFF3E" box="[852,1115,152,175]" italics="true" pageId="5" pageNumber="6">Chiromantes obtusifrons</emphasis>
</taxonomicName>
complex.
</paragraph>
</caption>
<subSubSection id="0FB47E86FF9AFFC1FEA22BB79077FB58" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="47112D0DFF9AFFC2FEA22BB79431FF77" blockId="5.[159,1428,208,308]" box="[363,1364,208,230]" pageId="5" pageNumber="6">
<emphasis id="75DAF11FFF9AFFC2FEA22BB79431FF77" bold="true" box="[363,1364,208,230]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="80AE568EFF9AFFC2FEA22BB7909BFF77" box="[363,510,208,230]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="obtusifrons">C. obtusifrons</taxonomicName>
<taxonomicName id="80AE568EFF9AFFC2FDF82BB793D3FF77" box="[561,694,208,230]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="garfunkel">C. garfunkel</taxonomicName>
<taxonomicName id="80AE568EFF9AFFC2FCC72BB7923EFF77" box="[782,859,208,230]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silus">C. silus</taxonomicName>
<taxonomicName id="80AE568EFF9AFFC2FC222BB79519FF77" box="[1003,1148,208,230]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leptomerus">C. leptomerus</taxonomicName>
<taxonomicName id="80AE568EFF9AFFC2FB0E2BB79431FF77" box="[1223,1364,208,230]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="eurymerus">C. eurymerus</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="47112D0DFF9AFFC2FF562B9B94F1FE83" blockId="5.[159,1428,208,308]" box="[159,1428,252,274]" pageId="5" pageNumber="6">Carapace dorsal Weakly convex Prominently convex Prominently convex Prominently convex Prominently convex</paragraph>
<paragraph id="47112D0DFF9AFFC2FF562A7A93FEFEA5" blockId="5.[159,1428,208,308]" box="[159,667,285,308]" pageId="5" pageNumber="6">
surface (
<figureCitation id="DF953188FF9AFFC2FEBB2A7A90DEFEA5" box="[370,443,285,308]" captionStart="FIGURE 10" captionStartId="13.[151,250,593,616]" captionTargetBox="[151,1436,193,571]" captionTargetId="figure@13.[151,1436,193,572]" captionTargetPageId="13" captionText="FIGURE 10. Lateral carapace views. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283291/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
A) (
<figureCitation id="DF953188FF9AFFC2FDF02A7A93E7FEA5" box="[569,642,285,308]" captionStart="FIGURE 10" captionStartId="13.[151,250,593,616]" captionTargetBox="[151,1436,193,571]" captionTargetId="figure@13.[151,1436,193,572]" captionTargetPageId="13" captionText="FIGURE 10. Lateral carapace views. A, Chiromantes obtusifrons (Dana, 1851), UFUF- 14837, male (17.7 × 13.2 mm), Hawaiian Is.; B, C. garfunkel sp. nov., QM-W 29172, holotype male (17.0 × 13.0 mm), Christmas I." httpUri="https://zenodo.org/record/283291/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
B)
</paragraph>
<paragraph id="47112D0DFF9AFFC2FF562A2D9425F8D9" pageId="5" pageNumber="6">
<table id="35AEDFADFF9A0038FF562A2E94F1F833" box="[159,1428,329,1954]" gridcols="6" gridrows="11" pageId="5" pageNumber="6">
<tr id="F99E2F4FFF9A0038FF562A2E94F1FE74" box="[159,1428,329,485]" gridrow="0" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF562A2E9030FE74" box="[159,341,329,485]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">Carapace granulation</th>
<th id="BA4F4633FF9A0038FEA22A2E937BFE74" box="[363,542,329,485]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">Anterior half conspicuously granular,granules become minute posteriorly</th>
<th id="BA4F4633FF9A0038FDF82A2E9394FE74" box="[561,753,329,485]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">Covered with fine, but distinct low granules becoming arranged into short striae posteriorly</th>
<th id="BA4F4633FF9A0038FCC72A2E92B3FE74" box="[782,982,329,485]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">Fine, but distinct low granules arranged into short striae that become longer posteriorly</th>
<th id="BA4F4633FF9A0038FC232A2E95CBFE74" box="[1002,1198,329,485]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">Fine, but distinct low granules becoming arranged into short striae posteriorly</th>
<th id="BA4F4633FF9A0038FB0E2A2E94F1FE74" box="[1223,1428,329,485]" gridcol="5" gridrow="0" pageId="5" pageNumber="6">Fine, but distinct low granules becoming arranged into short striae posteriorly</th>
</tr>
<tr id="F99E2F4FFF9A0038FF562A9C94F1FDE4" box="[159,1428,507,629]" gridrow="1" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF562A9C9030FDE4" box="[159,341,507,629]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">Exorbital tooth outer margin</th>
<td id="BA4F4633FF9A0038FEA22A9C937BFDE4" box="[363,542,507,629]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">Broadly convex</td>
<td id="BA4F4633FF9A0038FDF82A9C9394FDE4" box="[561,753,507,629]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">Long, slightly convex</td>
<td id="BA4F4633FF9A0038FCC72A9C92B3FDE4" box="[782,982,507,629]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">Long, weakly oblique, forming rounded obtuse angle posteriorly</td>
<td id="BA4F4633FF9A0038FC232A9C95CBFDE4" box="[1002,1198,507,629]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">Long, moderately oblique, forming slight angle posteriorly</td>
<td id="BA4F4633FF9A0038FB0E2A9C94F1FDE4" box="[1223,1428,507,629]" gridcol="5" gridrow="1" pageId="5" pageNumber="6">Broadly convex</td>
</tr>
<tr id="F99E2F4FFF9A0038FF5629EC94F1FD72" box="[159,1428,651,739]" gridrow="2" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF5629EC9030FD72" box="[159,341,651,739]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">Constriction posterior to epibranchial tooth</th>
<td id="BA4F4633FF9A0038FEA229EC937BFD72" box="[363,542,651,739]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">Absent</td>
<td id="BA4F4633FF9A0038FDF829EC9394FD72" box="[561,753,651,739]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">Present</td>
<td id="BA4F4633FF9A0038FCC729EC92B3FD72" box="[782,982,651,739]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">Markedly constricted</td>
<td id="BA4F4633FF9A0038FC2329EC95CBFD72" box="[1002,1198,651,739]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">Present</td>
<td id="BA4F4633FF9A0038FB0E29EC94F1FD72" box="[1223,1428,651,739]" gridcol="5" gridrow="2" pageId="5" pageNumber="6">Absent</td>
</tr>
<tr id="F99E2F4FFF9A0038FF56299D94F1FCC3" box="[159,1428,762,850]" gridrow="3" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF56299D9030FCC3" box="[159,341,762,850]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">Postfrontal lobes</th>
<td id="BA4F4633FF9A0038FEA2299D937BFCC3" box="[363,542,762,850]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">Relatively low, rounded</td>
<td id="BA4F4633FF9A0038FDF8299D9394FCC3" box="[561,753,762,850]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">Strongly demarcated</td>
<td id="BA4F4633FF9A0038FCC7299D92B3FCC3" box="[782,982,762,850]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">Strongly demarcated</td>
<td id="BA4F4633FF9A0038FC23299D95CBFCC3" box="[1002,1198,762,850]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">Strongly demarcated</td>
<td id="BA4F4633FF9A0038FB0E299D94F1FCC3" box="[1223,1428,762,850]" gridcol="5" gridrow="3" pageId="5" pageNumber="6">Well demarcated but relatively low, rounded</td>
</tr>
<tr id="F99E2F4FFF9A0038FF56280F94F1FC50" box="[159,1428,872,961]" gridrow="4" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF56280F9030FC50" box="[159,341,872,961]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Frontal width/ carapace width ratio</th>
<td id="BA4F4633FF9A0038FEA2280F937BFC50" box="[363,542,872,961]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">c. 0.65 times</td>
<td id="BA4F4633FF9A0038FDF8280F9394FC50" box="[561,753,872,961]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">c. 0.6 times</td>
<td id="BA4F4633FF9A0038FCC7280F92B3FC50" box="[782,982,872,961]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">c. 0.65</td>
<td id="BA4F4633FF9A0038FC23280F95CBFC50" box="[1002,1198,872,961]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">c. 0.65</td>
<td id="BA4F4633FF9A0038FB0E280F94F1FC50" box="[1223,1428,872,961]" gridcol="5" gridrow="4" pageId="5" pageNumber="6">c. 0.650.7</td>
</tr>
<tr id="F99E2F4FFF9A0038FF5628B094F1FBE3" box="[159,1428,983,1138]" gridrow="5" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF5628B09030FBE3" box="[159,341,983,1138]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">Shape of frontal border</th>
<td id="BA4F4633FF9A0038FEA228B0937BFBE3" box="[363,542,983,1138]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">Broadly convex in frontal view, but medial part relatively straight in dorsal view</td>
<td id="BA4F4633FF9A0038FDF828B09394FBE3" box="[561,753,983,1138]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">Broadly convex in dorsal view</td>
<td id="BA4F4633FF9A0038FCC728B092B3FBE3" box="[782,982,983,1138]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">Margin straight or slightly concave in dorsal view</td>
<td id="BA4F4633FF9A0038FC2328B095CBFBE3" box="[1002,1198,983,1138]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">Margin slightly to broadly concave in dorsal view</td>
<td id="BA4F4633FF9A0038FB0E28B094F1FBE3" box="[1223,1428,983,1138]" gridcol="5" gridrow="5" pageId="5" pageNumber="6">Margin broadly convex in frontal view, but medial part relatively straight in dorsal view</td>
</tr>
<tr id="F99E2F4FFF9A0038FF562FEF94F1FA93" box="[159,1428,1160,1282]" gridrow="6" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF562FEF9030FA93" box="[159,341,1160,1282]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">Granulation of frontal margin</th>
<td id="BA4F4633FF9A0038FEA22FEF937BFA93" box="[363,542,1160,1282]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">Beaded with row of small but distinct granules</td>
<td id="BA4F4633FF9A0038FDF82FEF9394FA93" box="[561,753,1160,1282]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">Minutely granular</td>
<td id="BA4F4633FF9A0038FCC72FEF92B3FA93" box="[782,982,1160,1282]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">Appearing smooth but microscopically granular</td>
<td id="BA4F4633FF9A0038FC232FEF95CBFA93" box="[1002,1198,1160,1282]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">Microscopically granular</td>
<td id="BA4F4633FF9A0038FB0E2FEF94F1FA93" box="[1223,1428,1160,1282]" gridcol="5" gridrow="6" pageId="5" pageNumber="6">Microscopically granular but appearing relatively smooth</td>
</tr>
<tr id="F99E2F4FFF9A0038FF562E7E94F1FAC1" box="[159,1428,1305,1360]" gridrow="7" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF562E7E9030FAC1" box="[159,341,1305,1360]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">Post-frontal lateral swellings</th>
<td id="BA4F4633FF9A0038FEA22E7E937BFAC1" box="[363,542,1305,1360]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">Low</td>
<td id="BA4F4633FF9A0038FDF82E7E9394FAC1" box="[561,753,1305,1360]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">Low</td>
<td id="BA4F4633FF9A0038FCC72E7E92B3FAC1" box="[782,982,1305,1360]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">Low</td>
<td id="BA4F4633FF9A0038FC232E7E95CBFAC1" box="[1002,1198,1305,1360]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">Prominent</td>
<td id="BA4F4633FF9A0038FB0E2E7E94F1FAC1" box="[1223,1428,1305,1360]" gridcol="5" gridrow="7" pageId="5" pageNumber="6">Low</td>
</tr>
<tr id="F99E2F4FFF9A0038FF562E0194F1FA71" box="[159,1428,1382,1504]" gridrow="8" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF562E019030FA71" box="[159,341,1382,1504]" gridcol="0" gridrow="8" pageId="5" pageNumber="6">Supraorbital margin</th>
<td id="BA4F4633FF9A0038FEA22E01937BFA71" box="[363,542,1382,1504]" gridcol="1" gridrow="8" pageId="5" pageNumber="6">Semicircular but oblique medial section somewhat straight</td>
<td id="BA4F4633FF9A0038FDF82E019394FA71" box="[561,753,1382,1504]" gridcol="2" gridrow="8" pageId="5" pageNumber="6">Medially straight</td>
<td id="BA4F4633FF9A0038FCC72E0192B3FA71" box="[782,982,1382,1504]" gridcol="3" gridrow="8" pageId="5" pageNumber="6">Semicircular but oblique medial section somewhat straight</td>
<td id="BA4F4633FF9A0038FC232E0195CBFA71" box="[1002,1198,1382,1504]" gridcol="4" gridrow="8" pageId="5" pageNumber="6">Semicircular</td>
<td id="BA4F4633FF9A0038FB0E2E0194F1FA71" box="[1223,1428,1382,1504]" gridcol="5" gridrow="8" pageId="5" pageNumber="6">Semicircular but oblique medial section somewhat straight</td>
</tr>
<tr id="F99E2F4FFF9A0038FF562E9194F1F900" box="[159,1428,1526,1681]" gridrow="9" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF562E919030F900" box="[159,341,1526,1681]" gridcol="0" gridrow="9" pageId="5" pageNumber="6">Cheliped merus</th>
<td id="BA4F4633FF9A0038FEA22E91937BF900" box="[363,542,1526,1681]" gridcol="1" gridrow="9" pageId="5" pageNumber="6">Inner anterior border with conspicous row of pointed granules</td>
<td id="BA4F4633FF9A0038FDF82E919394F900" box="[561,753,1526,1681]" gridcol="2" gridrow="9" pageId="5" pageNumber="6">Minutely granular</td>
<td id="BA4F4633FF9A0038FCC72E9192B3F900" box="[782,982,1526,1681]" gridcol="3" gridrow="9" pageId="5" pageNumber="6">Inner anterior border armed with small pointed granules</td>
<td id="BA4F4633FF9A0038FC232E9195CBF900" box="[1002,1198,1526,1681]" gridcol="4" gridrow="9" pageId="5" pageNumber="6">Minutely granular</td>
<td id="BA4F4633FF9A0038FB0E2E9194F1F900" box="[1223,1428,1526,1681]" gridcol="5" gridrow="9" pageId="5" pageNumber="6">Minutely granular</td>
</tr>
<tr id="F99E2F4FFF9A0038FF562DC094F1F833" box="[159,1428,1703,1954]" gridrow="10" pageId="5" pageNumber="6">
<th id="BA4F4633FF9A0038FF562DC09030F833" box="[159,341,1703,1954]" gridcol="0" gridrow="10" pageId="5" pageNumber="6">Cheliped carpus outer margin and dorsal surface P4 merus and propodus length/ breadth ratios, respectively</th>
<td id="BA4F4633FF9A0038FEA22DC0937BF833" box="[363,542,1703,1954]" gridcol="1" gridrow="10" pageId="5" pageNumber="6">Covered with small but conspicuous granules 2.2; 2.1 times longer</td>
<td id="BA4F4633FF9A0038FDF82DC09394F833" box="[561,753,1703,1954]" gridcol="2" gridrow="10" pageId="5" pageNumber="6">Moderately granular 2.3; 2.8</td>
<td id="BA4F4633FF9A0038FCC72DC092B3F833" box="[782,982,1703,1954]" gridcol="3" gridrow="10" pageId="5" pageNumber="6">Conspicuously granular 2.3; 2.6</td>
<td id="BA4F4633FF9A0038FC232DC095CBF833" box="[1002,1198,1703,1954]" gridcol="4" gridrow="10" pageId="5" pageNumber="6">Conspicuously granular 2.3; 2.8</td>
<td id="BA4F4633FF9A0038FB0E2DC094F1F833" box="[1223,1428,1703,1954]" gridcol="5" gridrow="10" pageId="5" pageNumber="6">Covered with small granules 2.0; 2.12.2</td>
</tr>
</table>
</paragraph>
<paragraph id="47112D0DFF9AFFC2FB9E2CD994F1F842" blockId="5.[1111,1428,1983,2003]" box="[1111,1428,1981,2003]" pageId="5" pageNumber="6">
<emphasis id="75DAF11FFF9AFFC2FB9E2CD994F1F842" box="[1111,1428,1981,2003]" italics="true" pageId="5" pageNumber="6">...... continued on the next page</emphasis>
</paragraph>
<caption id="13D17D85FF99FFC1FF562BC290EAFF2A" ID-Table-UUID="13D17D85FF99FFC1FF562BC290EAFF2A" box="[159,399,165,187]" httpUri="http://table.plazi.org/id/13D17D85FF99FFC1FF562BC290EAFF2A" pageId="6" pageNumber="7" targetBox="[159,1414,248,999]" targetIsTable="true" targetPageId="6">
<paragraph id="47112D0DFF99FFC1FF562BC290EAFF2A" blockId="6.[159,1364,165,231]" box="[159,399,165,187]" pageId="6" pageNumber="7">
<emphasis id="75DAF11FFF99FFC1FF562BC2906BFF2B" bold="true" box="[159,270,165,186]" pageId="6" pageNumber="7">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="47112D0DFF99FFC1FEA22BB69077FB58" blockId="6.[159,1364,165,231]" lastBlockId="6.[151,1436,1057,1441]" pageId="6" pageNumber="7">
<emphasis id="75DAF11FFF99FFC1FEA22BB69431FF76" bold="true" box="[363,1364,209,231]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="80AE568EFF99FFC1FEA22BB6909BFF76" box="[363,510,209,231]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusifrons">C. obtusifrons</taxonomicName>
<taxonomicName id="80AE568EFF99FFC1FDF82BB693D3FF76" box="[561,694,209,231]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="garfunkel">C. garfunkel</taxonomicName>
<taxonomicName id="80AE568EFF99FFC1FCC72BB6923EFF76" box="[782,859,209,231]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="silus">C. silus</taxonomicName>
<taxonomicName id="80AE568EFF99FFC1FC222BB69519FF76" box="[1003,1148,209,231]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="leptomerus">C. leptomerus</taxonomicName>
<taxonomicName id="80AE568EFF99FFC1FB0E2BB69431FF76" box="[1223,1364,209,231]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="eurymerus">C. eurymerus</taxonomicName>
</emphasis>
We have included previous reports of
<taxonomicName id="80AE568EFF99FFC1FDA62F459269FBA8" box="[623,780,1057,1081]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF99FFC1FDA62F459269FBA8" box="[623,780,1057,1081]" italics="true" pageId="6" pageNumber="7">C. obtusifrons</emphasis>
</taxonomicName>
by other authors under the synonymies of our new species as appropriate, however the record of De Man (1895, 1898) from the southern coast of Sumatra must remain
<emphasis id="75DAF11FFF99FFC1FF5E2F0D9054FB10" box="[151,305,1129,1153]" italics="true" pageId="6" pageNumber="7">incertae sedis</emphasis>
until his material can be reexamined. It occurs relatively close to Christmas I., but we believe it to be different from
<taxonomicName id="80AE568EFF99FFC1FE8F2FE990BDFB34" box="[326,472,1165,1189]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="garfunkel" status="sp. nov.">
<emphasis id="75DAF11FFF99FFC1FE8F2FE990BDFB34" box="[326,472,1165,1189]" italics="true" pageId="6" pageNumber="7">C. garfunkel</emphasis>
</taxonomicName>
<emphasis id="75DAF11FFF99FFC1FE2C2FEA9321FB34" bold="true" box="[485,580,1165,1189]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="6EE94C64FF99FFC1FE2C2FEA9321FB34" box="[485,580,1165,1189]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and probably represent a yet undescribed species (see Remarks for
<taxonomicName id="80AE568EFF99FFC1FA4B2FE99067FB58" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="garfunkel">
<emphasis id="75DAF11FFF99FFC1FA4B2FE99067FB58" italics="true" pageId="6" pageNumber="7">C. garfunkel</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<paragraph id="47112D0DFF99FFC1FF562B9F959BFC76" pageId="6" pageNumber="7">
<table id="35AEDFADFF990038FF562B9F94E3FC76" box="[159,1414,248,999]" gridcols="6" gridrows="9" pageId="6" pageNumber="7">
<tr id="F99E2F4FFF990038FF562B9F94E3FEF8" box="[159,1414,248,361]" gridrow="0" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF562B9F903FFEF8" box="[159,346,248,361]" gridcol="0" gridrow="0" pageId="6" pageNumber="7">P5 merus and propodus length/ breadth ratios, respectively</th>
<th id="BA4F4633FF990038FEA22B9F937BFEF8" box="[363,542,248,361]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">2.3; 2.2 times longer</th>
<th id="BA4F4633FF990038FDF82B9F9393FEF8" box="[561,758,248,361]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">2.6; 2.7</th>
<th id="BA4F4633FF990038FCC72B9F92A3FEF8" box="[782,966,248,361]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">2.4; 2.3</th>
<th id="BA4F4633FF990038FC222B9F95C7FEF8" box="[1003,1186,248,361]" gridcol="4" gridrow="0" pageId="6" pageNumber="7">2.22.3; 2.4</th>
<th id="BA4F4633FF990038FB0E2B9F94E3FEF8" box="[1223,1414,248,361]" gridcol="5" gridrow="0" pageId="6" pageNumber="7">2.02.1; 2.12.2</th>
</tr>
<tr id="F99E2F4FFF990038FF562A1A94E3FE41" box="[159,1414,381,464]" gridrow="1" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF562A1A903FFE41" box="[159,346,381,464]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">Pereiopods 25 carpus accessory ridges</th>
<td id="BA4F4633FF990038FEA22A1A937BFE41" box="[363,542,381,464]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">Conspicuous</td>
<td id="BA4F4633FF990038FDF82A1A9393FE41" box="[561,758,381,464]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">Not strongly marked</td>
<td id="BA4F4633FF990038FCC72A1A92A3FE41" box="[782,966,381,464]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">Not strongly marked</td>
<td id="BA4F4633FF990038FC222A1A95C7FE41" box="[1003,1186,381,464]" gridcol="4" gridrow="1" pageId="6" pageNumber="7">Not strongly marked</td>
<td id="BA4F4633FF990038FB0E2A1A94E3FE41" box="[1223,1414,381,464]" gridcol="5" gridrow="1" pageId="6" pageNumber="7">Not strongly marked</td>
</tr>
<tr id="F99E2F4FFF990038FF562A8494E3FD89" box="[159,1414,483,536]" gridrow="2" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF562A84903FFD89" box="[159,346,483,536]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">Telson length/ breadth ratio</th>
<td id="BA4F4633FF990038FEA22A84937BFD89" box="[363,542,483,536]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">1.1</td>
<td id="BA4F4633FF990038FDF82A849393FD89" box="[561,758,483,536]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">1.1</td>
<td id="BA4F4633FF990038FCC72A8492A3FD89" box="[782,966,483,536]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">1.1</td>
<td id="BA4F4633FF990038FC222A8495C7FD89" box="[1003,1186,483,536]" gridcol="4" gridrow="2" pageId="6" pageNumber="7">1.1</td>
<td id="BA4F4633FF990038FB0E2A8494E3FD89" box="[1223,1414,483,536]" gridcol="5" gridrow="2" pageId="6" pageNumber="7">1.2</td>
</tr>
<tr id="F99E2F4FFF990038FF56294C94E3FDF1" box="[159,1414,555,608]" gridrow="3" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF56294C903FFDF1" box="[159,346,555,608]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">Telson length/ somite 6 length ratio</th>
<td id="BA4F4633FF990038FEA2294C937BFDF1" box="[363,542,555,608]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">1.15</td>
<td id="BA4F4633FF990038FDF8294C9393FDF1" box="[561,758,555,608]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">1.05</td>
<td id="BA4F4633FF990038FCC7294C92A3FDF1" box="[782,966,555,608]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">1.1</td>
<td id="BA4F4633FF990038FC22294C95C7FDF1" box="[1003,1186,555,608]" gridcol="4" gridrow="3" pageId="6" pageNumber="7">1.06</td>
<td id="BA4F4633FF990038FB0E294C94E3FDF1" box="[1223,1414,555,608]" gridcol="5" gridrow="3" pageId="6" pageNumber="7">1.41.5</td>
</tr>
<tr id="F99E2F4FFF990038FF56291494E3FD39" box="[159,1414,627,680]" gridrow="4" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF562914903FFD39" box="[159,346,627,680]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">Somite 6 length/ width ratio</th>
<td id="BA4F4633FF990038FEA22914937BFD39" box="[363,542,627,680]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">2.0</td>
<td id="BA4F4633FF990038FDF829149393FD39" box="[561,758,627,680]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">1.8</td>
<td id="BA4F4633FF990038FCC7291492A3FD39" box="[782,966,627,680]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">1.9</td>
<td id="BA4F4633FF990038FC22291495C7FD39" box="[1003,1186,627,680]" gridcol="4" gridrow="4" pageId="6" pageNumber="7">1.8</td>
<td id="BA4F4633FF990038FB0E291494E3FD39" box="[1223,1414,627,680]" gridcol="5" gridrow="4" pageId="6" pageNumber="7">2.3</td>
</tr>
<tr id="F99E2F4FFF990038FF5629DC94E3FCDD" box="[159,1414,699,844]" gridrow="5" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF5629DC903FFCDD" box="[159,346,699,844]" gridcol="0" gridrow="5" pageId="6" pageNumber="7">Somite 6 lateral margin shape</th>
<td id="BA4F4633FF990038FEA229DC937BFCDD" box="[363,542,699,844]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">Distolateral margins strongly divergent,relatively straight over distal two-thirds</td>
<td id="BA4F4633FF990038FDF829DC9393FCDD" box="[561,758,699,844]" gridcol="2" gridrow="5" pageId="6" pageNumber="7">Divergent and slightly concave over distal half, but more subparallel over proximal half</td>
<td id="BA4F4633FF990038FCC729DC92A3FCDD" box="[782,966,699,844]" gridcol="3" gridrow="5" pageId="6" pageNumber="7">Broadly divergent with margins relatively evenly convex</td>
<td id="BA4F4633FF990038FC2229DC95C7FCDD" box="[1003,1186,699,844]" gridcol="4" gridrow="5" pageId="6" pageNumber="7">Lateral margins divergent with margins relatively evenly convex</td>
<td id="BA4F4633FF990038FB0E29DC94E3FCDD" box="[1223,1414,699,844]" gridcol="5" gridrow="5" pageId="6" pageNumber="7">Distolateral margins relatively straight, strongly divergent, margins convex overall</td>
</tr>
<tr id="F99E2F4FFF990038FF56280794E3FC05" box="[159,1414,864,916]" gridrow="6" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF562807903FFC05" box="[159,346,864,916]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">Telson width/ somite 3 width ratio</th>
<td id="BA4F4633FF990038FEA22807937BFC05" box="[363,542,864,916]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">2.9</td>
<td id="BA4F4633FF990038FDF828079393FC05" box="[561,758,864,916]" gridcol="2" gridrow="6" pageId="6" pageNumber="7">3.0</td>
<td id="BA4F4633FF990038FCC7280792A3FC05" box="[782,966,864,916]" gridcol="3" gridrow="6" pageId="6" pageNumber="7">3.2</td>
<td id="BA4F4633FF990038FC22280795C7FC05" box="[1003,1186,864,916]" gridcol="4" gridrow="6" pageId="6" pageNumber="7">2.8</td>
<td id="BA4F4633FF990038FB0E280794E3FC05" box="[1223,1414,864,916]" gridcol="5" gridrow="6" pageId="6" pageNumber="7">3.1</td>
</tr>
<tr id="F99E2F4FFF990038FF5628CF94E3FC2F" box="[159,1414,936,958]" gridrow="7" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF5628CF903FFC2F" box="[159,346,936,958]" gridcol="0" gridrow="7" pageId="6" pageNumber="7">G1shaft</th>
<td id="BA4F4633FF990038FEA228CF937BFC2F" box="[363,542,936,958]" gridcol="1" gridrow="7" pageId="6" pageNumber="7">Slender</td>
<td id="BA4F4633FF990038FDF828CF9393FC2F" box="[561,758,936,958]" gridcol="2" gridrow="7" pageId="6" pageNumber="7">Slender</td>
<td id="BA4F4633FF990038FCC728CF92A3FC2F" box="[782,966,936,958]" gridcol="3" gridrow="7" pageId="6" pageNumber="7">Slender</td>
<td id="BA4F4633FF990038FC2228CF95C7FC2F" box="[1003,1186,936,958]" gridcol="4" gridrow="7" pageId="6" pageNumber="7">Slender</td>
<td id="BA4F4633FF990038FB0E28CF94E3FC2F" box="[1223,1414,936,958]" gridcol="5" gridrow="7" pageId="6" pageNumber="7">Broad,stocky</td>
</tr>
<tr id="F99E2F4FFF990038FF5628B694E3FC76" box="[159,1414,977,999]" gridrow="8" pageId="6" pageNumber="7">
<th id="BA4F4633FF990038FF5628B6903FFC76" box="[159,346,977,999]" gridcol="0" gridrow="8" pageId="6" pageNumber="7">G1 tip</th>
<td id="BA4F4633FF990038FEA228B6937BFC76" box="[363,542,977,999]" gridcol="1" gridrow="8" pageId="6" pageNumber="7">Long, slender</td>
<td id="BA4F4633FF990038FDF828B69393FC76" box="[561,758,977,999]" gridcol="2" gridrow="8" pageId="6" pageNumber="7">Long, slender</td>
<td id="BA4F4633FF990038FCC728B692A3FC76" box="[782,966,977,999]" gridcol="3" gridrow="8" pageId="6" pageNumber="7">Long, slender</td>
<td id="BA4F4633FF990038FC2228B695C7FC76" box="[1003,1186,977,999]" gridcol="4" gridrow="8" pageId="6" pageNumber="7">Long, slender</td>
<td id="BA4F4633FF990038FB0E28B694E3FC76" box="[1223,1414,977,999]" gridcol="5" gridrow="8" pageId="6" pageNumber="7">Short</td>
</tr>
</table>
</paragraph>
<subSubSection id="0FB47E86FF99FFC1FF0E2FB29349FA30" pageId="6" pageNumber="7" type="distribution">
<paragraph id="47112D0DFF99FFC1FF0E2FB2924CFB7C" blockId="6.[151,1436,1057,1441]" box="[199,809,1237,1262]" pageId="6" pageNumber="7">
<emphasis id="75DAF11FFF99FFC1FF0E2FB2903CFB7F" bold="true" box="[199,345,1237,1262]" pageId="6" pageNumber="7">Distribution</emphasis>
. Only known from the Hawaiian Islands.
</paragraph>
<paragraph id="47112D0DFF99FFC1FF0E2F9D9349FA30" blockId="6.[151,1436,1057,1441]" pageId="6" pageNumber="7">
<emphasis id="75DAF11FFF99FFC1FF0E2F9D904EFA83" bold="true" box="[199,299,1274,1298]" pageId="6" pageNumber="7">Ecology.</emphasis>
Edmondson (1959) noted that
<taxonomicName id="80AE568EFF99FFC1FD442F9D9249FA80" box="[653,812,1273,1297]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="75DAF11FFF99FFC1FD442F9D9249FA80" box="[653,812,1273,1297]" italics="true" pageId="6" pageNumber="7">C. obtusifrons</emphasis>
</taxonomicName>
was found intertidally and “even above the high water mark”, and Paulay &amp; Starmer (2011: 11) reported finding it living in a supratidal boulder field on Oahu. Paulay &amp; Starmer (2011: 11) nevertheless also suggested that this species may not be as terrestrial as
<taxonomicName id="80AE568EFF99FFC1FB6E2E269417FAC8" box="[1191,1394,1345,1369]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="silus" status="sp. nov.">
<emphasis id="75DAF11FFF99FFC1FB6E2E269417FAC8" box="[1191,1394,1345,1369]" italics="true" pageId="6" pageNumber="7">Chiromantes silus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EE94C64FF99FFC1FAB32E2591ADFAEC" pageId="6" pageNumber="7" rank="species">
<emphasis id="75DAF11FFF99FFC1FAB32E2591ADFAEC" bold="true" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
(see below), which they had observed in
<collectingCountry id="3FB96D9DFF99FFC1FD552E029278FAEC" box="[668,797,1381,1405]" name="Micronesia" pageId="6" pageNumber="7">Micronesia</collectingCountry>
, but indicated that additional observations on its ecology in the Hawaiian Is. would be useful.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="2C4D913CA1FDF914DDEDB62AF00DC0D9" ID-DOI="http://dx.doi.org/10.3897/zookeys.424.7388" ID-GBIF-Dataset="bb867cb3-6bab-4bb4-9c0e-e260f98b7425" ID-PMC="PMC4106100" ID-Pensoft-Pub="1313-2970-424-19" ID-PubMed="25061395" ID-ZooBank="38EAC4B788344054B9AC9747AC476543" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-424-19" ModsDocOrigin="ZooKeys 424" ModsDocTitle="Systematic and biogeographical study of Protura (Hexapoda) in Russian Far East: new data on high endemism of the group" checkinTime="1451245639639" checkinUser="pensoft" docAuthor="Bu, Yun, Potapov, Mikhail B. &amp; Yin, Wen Ying" docDate="2014" docId="CF07AAB7A693CE19FF34E40668C56F83" docLanguage="en" docName="ZooKeys 424: 19-57" docOrigin="ZooKeys 424" docSource="http://dx.doi.org/10.3897/zookeys.424.7388" docTitle="Baculentulus samchonri Imdadate &amp; Szeptycki 1976" docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="1667FFD97D393E4B7E41FFBAFFBCFFA2" masterDocTitle="Systematic and biogeographical study of Protura (Hexapoda) in Russian Far East: new data on high endemism of the group" masterLastPageNumber="57" masterPageNumber="19" pageNumber="35" updateTime="1701317129163" updateUser="plazi">
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<mods:title id="3A8AD5E125A983360C976CEFDE1A62C4">Systematic and biogeographical study of Protura (Hexapoda) in Russian Far East: new data on high endemism of the group</mods:title>
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<paragraph id="B842DEFB5D423CE5151D25FB5CD44BDA" pageId="16" pageNumber="35">Taxon classification Animalia Protura Berberentulidae</paragraph>
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Baculentulus samchonri
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<paragraph id="EB1E542B1FB0233E0D5993DC9BD32A58" pageId="16" pageNumber="35">Material examined.</paragraph>
<paragraph id="260746D91406DE95B05EC746044279AF" pageId="16" pageNumber="35">1 female, 1 male, locality 5, 16-IX-2011, 12 females, 7 males, locality 7, 22-IX-2011, coll. Y. Bu, C. W. Huang, M. Potapov &amp; V. Alpatov.</paragraph>
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<paragraph id="8375CF8930636D9A08E75AD5C958EF50" pageId="16" pageNumber="35">Distribution.</paragraph>
<paragraph id="94C617A5D15DEB1E43A5A0BE7DC83011" pageId="16" pageNumber="35">Korea; Russia (Far East: Primorsky Krai). New for Russia.</paragraph>
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<emphasis id="75DA84432735FFEEFF42F945FE9B59D4" box="[224,317,1721,1747]" italics="true" pageId="13" pageNumber="14">alaemon</emphasis>
]
<treatmentCitation id="C60F7E402735FFEEFEECF945FC9059D4" box="[334,822,1721,1748]" pageId="13" pageNumber="14">
<taxonomicName id="80AE23D22735FFEEFEECF945FC9059D4" ID-CoL="1401b564-3306-4d83-ae6a-503aaa79fdef" authority="Nobili, 1899: 241" authorityName="Nobili" authorityPageNumber="241" authorityYear="1899" box="[334,822,1721,1748]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ustulatus" subGenus="Eupalaemon">
(
<emphasis id="75DA84432735FFEEFEFAF945FE4259D4" box="[344,484,1721,1747]" italics="true" pageId="13" pageNumber="14">Eupalaemon</emphasis>
)
<emphasis id="75DA84432735FFEEFE57F945FDF959D4" box="[501,607,1721,1747]" italics="true" pageId="13" pageNumber="14">ustulatus</emphasis>
<bibRefCitation id="233F25A02735FFEEFDC4F945FC9059D4" author="Nobili" box="[614,822,1721,1747]" firstAuthor="Nobili" pageId="13" pageNumber="14" pagination="230 - 282" refId="ref11537" refString="Nobili G. 1899. Contribuzioni alla conoscenza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali del Museo civico di Storia naturale di Genova, Series 2 20 (40): 230 - 282." type="journal article" year="1899">Nobili, 1899: 241</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(Rigo, SE
<collectingRegion id="856A96B32735FFEEFC1AF946FBA359D3" box="[952,1029,1722,1748]" country="Indonesia" name="Papua Barat" pageId="13" pageNumber="14">Papua</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="471158512735FFEEFF1FF907FB815812" blockId="13.[189,1063,1786,1813]" box="[189,1063,1786,1813]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="67BE7F7F2735FFEEFF1FF907FB815812" box="[189,1063,1786,1813]" pageId="13" pageNumber="14">
<taxonomicName id="80AE23D22735FFEEFF1FF907FE035812" ID-CoL="1401b564-3306-4d83-ae6a-503aaa79fdef" authorityName="Nobili" authorityYear="1899" box="[189,421,1787,1813]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ustulatus">
<emphasis id="75DA84432735FFEEFF1FF907FE035812" box="[189,421,1787,1813]" italics="true" pageId="13" pageNumber="14">Palaemon ustulatus</emphasis>
</taxonomicName>
<emphasis id="75DA84432735FFEEFE0FF907FE1A5812" bold="true" box="[429,444,1787,1813]" pageId="13" pageNumber="14"></emphasis>
<treatmentCitation id="C60F7E402735FFEEFE61F906FD055812" author="De Man J. G." box="[451,675,1786,1813]" page="431" pageId="13" pageNumber="14" year="1915">
<bibRefCitation id="233F25A02735FFEEFE61F906FD055812" author="de Man" box="[451,675,1786,1813]" firstAuthor="de Man" pageId="13" pageNumber="14" pagination="385 - 458" refId="ref10791" refString="De Man J. G. 1915. Zur Fauna von Nord-Neuguinea. Nach den Sammlungen von Dr. P. N. van Kampen und K. Gjellerup in den Jahren 1910 - 1911. Macrura. Zoologische Jahrbucher - Abteilung fur Systematik, Geographie und Biologie der Thiere 38: 385 - 458." type="journal article" year="1915">de Man 1915: 431</bibRefCitation>
</treatmentCitation>
, pl. 29, fig. 32 (Rigo, SE
<collectingRegion id="856A96B32735FFEEFC6CF907FBBD5812" box="[974,1051,1787,1813]" country="Indonesia" name="Papua Barat" pageId="13" pageNumber="14">Papua</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0FB40BDA2735FFECFF1FF8C0FD285ECF" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="471158512735FFEEFF1FF8C0FE0C5851" blockId="13.[189,426,1852,1878]" box="[189,426,1852,1878]" pageId="13" pageNumber="14">
<heading id="1C59EF3D2735FFEEFF1FF8C0FE0C5851" bold="true" box="[189,426,1852,1878]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="75DA84432735FFEEFF1FF8C0FE0C5851" bold="true" box="[189,426,1852,1878]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="F7C6520C2735FFEEFF48F897FE3458CB" ID-GBIF-Occurrence="2613744301" collectionCode="MSNG" country="Papua New Guinea" latitude="-9.8767" location="Rigo District" longLatPrecision="7" longitude="147.7938" pageId="13" pageNumber="14" specimenCode="MSNG VII-106" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="471158512735FFEEFF48F897FEFA5882" blockId="13.[189,1399,1899,1996]" box="[234,348,1899,1925]" pageId="13" pageNumber="14">
<emphasis id="75DA84432735FFEEFF48F897FEFA5882" bold="true" box="[234,348,1899,1925]" pageId="13" pageNumber="14">
<typeStatus id="9815E6F32735FFEEFF48F897FEFA5882" box="[234,348,1899,1925]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="471158512735FFEEFF1FF872FE3458CB" blockId="13.[189,1399,1899,1996]" pageId="13" pageNumber="14">
<collectingCountry id="3FB918C12735FFEEFF1FF872FE4158AF" box="[189,487,1934,1960]" name="Papua New Guinea" pageId="13" pageNumber="14">PAPUA NEW GUINEA</collectingCountry>
:
<specimenCount id="51A893D82735FFEEFE54F872FD8E58AF" box="[502,552,1934,1960]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
,
<location id="42710E8A2735FFEEFD9BF872FD7F58AF" LSID="urn:lsid:plazi:treatment:CF07E9472735FFF2FDBDF9A8FE845E4E:42710E8A2735FFEEFD9BF872FD7F58AF" box="[569,729,1934,1960]" country="Papua New Guinea" latitude="-9.8767" longLatPrecision="7" longitude="147.7938" name="Rigo District" pageId="13" pageNumber="14">Rigo District</location>
,
<geoCoordinate id="229A3E962735FFEEFD4BF872FCC458AF" box="[745,866,1934,1960]" degrees="9.8767" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="5" value="-9.8767">9.8767° S</geoCoordinate>
,
<geoCoordinate id="229A3E962735FFEEFCD6F872FBAB58AF" box="[884,1037,1934,1960]" degrees="147.7938" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="5" value="147.7938">147.7938° E</geoCoordinate>
, unknown date,
<quantity id="8056F5B42735FFEEFB78F872FAE158AF" box="[1242,1351,1934,1961]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="13" pageNumber="14" unit="mm" value="23.5">23.5 mm</quantity>
CL (
<specimenCode id="1708F02A2735FFEEFF67F84DFE2058CB" box="[197,390,1969,1996]" pageId="13" pageNumber="14">MSNG VII-106</specimenCode>
).
</paragraph>
</materialsCitation>
<caption id="13D108D92736FFEDFF1FF856FAC558EF" ID-DOI="http://doi.org/10.5281/zenodo.3824737" ID-Zenodo-Dep="3824737" httpUri="https://zenodo.org/record/3824737/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1962,1988]" targetBox="[215,1375,265,1928]" targetPageId="14">
<paragraph id="471158512736FFEDFF1FF856FAC558EF" blockId="14.[189,1398,1962,2024]" pageId="14" pageNumber="15">
<emphasis id="75DA84432736FFEDFF1FF856FE8A58C3" bold="true" box="[189,300,1962,1988]" pageId="14" pageNumber="15">Fig. 4. A</emphasis>
. Live coloration of
<taxonomicName id="80AE23D22736FFEDFD82F856FCEE58C3" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" box="[544,840,1962,1988]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="australe">
<emphasis id="75DA84432736FFEDFD82F856FCEE58C3" box="[544,840,1962,1988]" italics="true" pageId="14" pageNumber="15">Macrobrachium australe</emphasis>
</taxonomicName>
(Guérin-Méneville, 1838 in Guérin-Méneville 18291838) (photo: E. Vigneux).
<emphasis id="75DA84432736FFEDFDE8F832FDF858EF" bold="true" box="[586,606,1998,2024]" pageId="14" pageNumber="15">B</emphasis>
. Live coloration of
<taxonomicName id="80AE23D22736FFEDFCE5F832FB3758EF" authority="(Nobili, 1899)" baseAuthorityName="Nobili" baseAuthorityYear="1899" box="[839,1169,1997,2024]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ustulatum">
<emphasis id="75DA84432736FFEDFCE5F832FC4458E0" box="[839,994,1997,2023]" italics="true" pageId="14" pageNumber="15">M. ustulatum</emphasis>
(
<bibRefCitation id="233F25A02736FFEDFC50F831FB2F58EF" author="Nobili" box="[1010,1161,1997,2024]" firstAuthor="Nobili" pageId="14" pageNumber="15" pagination="230 - 282" refId="ref11537" refString="Nobili G. 1899. Contribuzioni alla conoscenza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali del Museo civico di Storia naturale di Genova, Series 2 20 (40): 230 - 282." type="journal article" year="1899">Nobili, 1899</bibRefCitation>
)
</taxonomicName>
(photo: P. Keith).
</paragraph>
</caption>
<paragraph id="471158512737FFECFF48FEF7FE005E22" blockId="15.[189,1399,267,364]" box="[234,422,267,293]" pageId="15" pageNumber="16">
<emphasis id="75DA84432737FFECFF48FEF7FE005E22" bold="true" box="[234,422,267,293]" pageId="15" pageNumber="16">Other material</emphasis>
</paragraph>
<paragraph id="471158512737FFECFF1FFED2FB2D5E6C" blockId="15.[189,1399,267,364]" pageId="15" pageNumber="16">
<collectingCountry id="3FB918C12737FFECFF1FFED2FEEA5E4F" box="[189,332,302,328]" name="Vanuatu" pageId="15" pageNumber="16">VANUATU</collectingCountry>
:
<specimenCount id="51A893D82737FFECFEF8FED2FE2D5E4E" box="[346,395,302,329]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Espiritu Santo Island, Ora Jourdain River,
<geoCoordinate id="229A3E962737FFECFC02FED2FB835E4F" box="[928,1061,302,329]" degrees="15.1666" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="5" value="-15.1666">15.1666° S</geoCoordinate>
,
<geoCoordinate id="229A3E962737FFECFB95FED2FB765E4E" box="[1079,1232,302,329]" degrees="166.8999" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="5" value="166.8999">166.8999° E</geoCoordinate>
,
<date id="33107E912737FFECFB7DFED2FAD75E4F" box="[1247,1393,302,328]" pageId="15" pageNumber="16" value="1994-10-03">3 Oct. 1994</date>
,
<quantity id="8056F5B42737FFECFF1FFEADFE805E6B" box="[189,294,337,364]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="15" pageNumber="16" unit="mm" value="20.7">20.7 mm</quantity>
CL (MNHN-IU-2013-13201);
<specimenCount id="51A893D82737FFECFD37FEADFD655E6B" box="[661,707,337,364]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
,
<quantity id="8056F5B42737FFECFD72FEADFC855E6B" box="[720,803,337,364]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="16" unit="mm" value="22.0">22 mm</quantity>
CL (MNHN-IU-2013-13202).
</paragraph>
<paragraph id="471158512737FFECFF1FFE77FD285ECF" blockId="15.[189,1399,395,457]" pageId="15" pageNumber="16">
<collectingCountry id="3FB918C12737FFECFF1FFE77FE455EA2" box="[189,483,395,421]" name="Palau" pageId="15" pageNumber="16">REPUBLIC OF PALAU</collectingCountry>
:
<specimenCount id="51A893D82737FFECFE4CFE77FDBA5EA1" box="[494,540,395,422]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Palau Island, Negmeskang River,
<geoCoordinate id="229A3E962737FFECFC15FE77FB965EA1" box="[951,1072,395,422]" degrees="7.5413" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="5" value="7.5413">7.5413° N</geoCoordinate>
,
<geoCoordinate id="229A3E962737FFECFB9DFE77FB735EA1" box="[1087,1237,395,422]" degrees="134.5778" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="5" value="134.5778">134.5778° E</geoCoordinate>
,
<date id="33107E912737FFECFB43FE77FAD45EA2" box="[1249,1394,395,422]" pageId="15" pageNumber="16" value="2011-03-01">1 Mar. 2011</date>
,
<quantity id="8056F5B42737FFECFF1FFE52FE805ECE" box="[189,294,430,457]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="15" pageNumber="16" unit="mm" value="20.7">20.7 mm</quantity>
CL (MNHN-IU-2013-13203).
</paragraph>
</subSubSection>
<subSubSection id="0FB40BDA2737FFECFF1FFE0CFC315AA1" pageId="15" pageNumber="16" type="description">
<paragraph id="471158512737FFECFF1FFE0CFEE85D0D" blockId="15.[189,334,496,522]" box="[189,334,496,522]" pageId="15" pageNumber="16">
<heading id="1C59EF3D2737FFECFF1FFE0CFEE85D0D" bold="true" box="[189,334,496,522]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="75DA84432737FFECFF1FFE0CFEE85D0D" bold="true" box="[189,334,496,522]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="471158512737FFECFF1FFDE3FC2B5DA5" blockId="15.[189,1398,542,674]" pageId="15" pageNumber="16">
ROSTRUM. Moderately long, reaching end of scaphocerite (
<figureCitation id="DF9544D42737FFECFCC7FDE2FC655D3E" box="[869,963,542,569]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5A</figureCitation>
); tip horizontal; dorsal margin nearly straight, faintly convex, with 1012 teeth, including 3 teeth on carapace; ventral margin with 34 teeth; teeth large, generally evenly spaced, with gap near anterior end of dorsal series; sometimes distance between 1
<superScript id="B0DBF5192737FFECFE97FD7BFEE65D92" attach="left" box="[309,320,647,661]" fontSize="6" pageId="15" pageNumber="16">st</superScript>
and 2
<superScript id="B0DBF5192737FFECFE2AFD7AFE3F5D93" attach="left" box="[392,409,646,660]" fontSize="6" pageId="15" pageNumber="16">nd</superScript>
teeth larger than between 2
<superScript id="B0DBF5192737FFECFD7FFD7AFD485D93" attach="left" box="[733,750,646,660]" fontSize="6" pageId="15" pageNumber="16">nd</superScript>
and 3
<superScript id="B0DBF5192737FFECFC94FD7AFCE25D93" attach="left" box="[822,836,646,660]" fontSize="6" pageId="15" pageNumber="16">rd</superScript>
teeth.
</paragraph>
<paragraph id="471158512737FFECFF1FFD3EFE295C6E" blockId="15.[189,1399,706,873]" pageId="15" pageNumber="16">
GENERAL CEPHALON. Eyes well developed; cornea as long as, but broader than peduncle; stylocerite distinctly pointed, reaching beyond basal segment; inferior orbit bluntly angular (
<figureCitation id="DF9544D42737FFECFB18FD19FA865C07" box="[1210,1312,741,768]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5A</figureCitation>
); postantennular carapace margin straight or slightly concave; protective setation absent on carapace; epistome lobes narrowly separated, strongly produced anteroventrally (
<figureCitation id="DF9544D42737FFECFC16FCD7FBB05C41" box="[948,1046,811,838]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5B</figureCitation>
); TS4 with rounded median process (
<figureCitation id="DF9544D42737FFECFE87FCB2FE245C6E" box="[293,386,846,873]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="471158512737FFECFF1FFC75FB3B5A6A" blockId="15.[189,1399,904,1389]" pageId="15" pageNumber="16">
SECOND CHELIPEDS (fully developed
<specimenCount id="51A893D82737FFECFDD6FC75FD2C5CA4" box="[628,650,905,931]" pageId="15" pageNumber="16" type="male"></specimenCount>
). Unequal sizes, non-isomorphic. Major P2 (
<figureCitation id="DF9544D42737FFECFB60FC74FA825CA4" box="[1218,1316,904,931]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5D</figureCitation>
) long, subcylindrical, with small part of merus extending beyond scaphocerite; carpus 0.680.73 CHL, 11.12 PL. Fingers slender 0.460.58 PL; dactylus shorter than FF. Cutting edges (
<figureCitation id="DF9544D42737FFECFB03FC33FB5D5CED" box="[1185,1275,975,1002]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5E</figureCitation>
) with 45 conical teeth (3
<superScript id="B0DBF5192737FFECFEDBFC0CFE215CF9" attach="left" box="[377,391,1008,1022]" fontSize="6" pageId="15" pageNumber="16">rd</superScript>
largest) followed by 610 evenly spaced, very small, blunt teeth, not along entire length; teeth diminishing in size distally; CEFF with teeth arrangement similar to that of dactylus; basally armed with ridge formed by 2 small teeth and 34 larger, conical teeth followed by 46 evenly spaced, small, blunt teeth, not along entire length; teeth diminishing in size distally. Largest tooth of FF lies just after largest tooth of CED when fingers closed. Cheliped with numerous spiniform setae, larger on inner edge, smaller and denser on outer edge. Numerous long setae present on manus and outer edge of fingers. Merus 0.540.61 CAL, ischium 0.640.85 MEL. Minor P2 (
<figureCitation id="DF9544D42737FFECFB9CFB39FB325BE7" box="[1086,1172,1221,1248]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5F</figureCitation>
) with entire carpus beyond scaphocerite. Carpus 0.670.75 CHL, 1.011.25 PL. Fingers slender 0.490.66 PL. CEFF with 1114 teeth, gradually diminishing in size distally (
<figureCitation id="DF9544D42737FFECFCADFAF0FCCB5A21" box="[783,877,1291,1318]" captionStart="Fig" captionStartId="16.[189,232,1858,1884]" captionTargetBox="[189,1399,268,1819]" captionTargetId="figure@16.[176,1411,214,1834]" captionTargetPageId="16" captionText="Fig. 5. Macrobrachium ustulatus (Nobili, 1899). AB, E. MNHN-IU-2013-13202. A. Cephalothorax. B. Epistome. E. Major second pereiopod finger. C, G. MNHN-IU-2013-13201. C. Fourth thoracic sternite. G. Minor second pereiopod finger. D, F. MNHN-IU-2013-13203. D. Major second pereiopod. F. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="15" pageNumber="16">Fig. 5G</figureCitation>
). CED with 1315 teeth, slightly smaller and more numerous than those of major P2, similar teeth arrangement to that of dactylus. Fingers, palm and carpus densely packed with velvety setae. MEL 0.620.72 CAL, IL 0.741.05 MEL.
</paragraph>
<paragraph id="471158512737FFECFF1FFA70FC315AA1" blockId="15.[189,919,1420,1446]" box="[189,919,1420,1446]" pageId="15" pageNumber="16">ABDOMEN. Smooth, inter-uropodal sclerite with preanal carina.</paragraph>
</subSubSection>
<subSubSection id="0FB40BDA2737FFECFF1FFA35FB9C5932" pageId="15" pageNumber="16" type="distribution">
<paragraph id="471158512737FFECFF1FFA35FEF05AE4" blockId="15.[189,342,1481,1507]" box="[189,342,1481,1507]" pageId="15" pageNumber="16">
<heading id="1C59EF3D2737FFECFF1FFA35FEF05AE4" bold="true" box="[189,342,1481,1507]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="75DA84432737FFECFF1FFA35FEF05AE4" bold="true" box="[189,342,1481,1507]" pageId="15" pageNumber="16">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="471158512737FFECFF1FFA04FB9C5932" blockId="15.[189,1398,1528,1590]" pageId="15" pageNumber="16">
Syntopic with
<taxonomicName id="80AE23D22737FFECFECFFA05FE515915" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1838" box="[365,503,1528,1554]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="australe">
<emphasis id="75DA84432737FFECFECFFA05FE515915" box="[365,503,1528,1554]" italics="true" pageId="15" pageNumber="16">M. australe</emphasis>
</taxonomicName>
in the Northwest Pacific Ocean (
<collectingCountry id="3FB918C12737FFECFC28FA04FC435915" box="[906,997,1528,1554]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
,
<collectingCountry id="3FB918C12737FFECFC50FA04FBDF5915" box="[1010,1145,1528,1554]" name="Philippines" pageId="15" pageNumber="16">Philippines</collectingCountry>
,
<collectingCountry id="3FB918C12737FFECFB2AFA05FAD45915" box="[1160,1394,1528,1555]" name="Papua New Guinea" pageId="15" pageNumber="16">Papua New Guinea</collectingCountry>
,
<collectingCountry id="3FB918C12737FFECFF1FF9E7FE895932" box="[189,303,1563,1589]" name="Indonesia" pageId="15" pageNumber="16">Indonesia</collectingCountry>
and
<collectingCountry id="3FB918C12737FFECFECAF9E7FE0D5931" box="[360,427,1563,1590]" name="Palau" pageId="15" pageNumber="16">Palau</collectingCountry>
) and in the Southwest Pacific Ocean (
<collectingCountry id="3FB918C12737FFECFCCAF9E7FC6A5932" box="[872,972,1563,1589]" name="Vanuatu" pageId="15" pageNumber="16">Vanuatu</collectingCountry>
) (
<figureCitation id="DF9544D42737FFECFC46F9E7FB8F5932" box="[996,1065,1563,1589]" captionStart="Fig" captionStartId="2.[189,232,1774,1800]" captionTargetBox="[189,1398,1283,1741]" captionTargetId="figure@2.[189,1398,1266,1762]" captionTargetPageId="2" captionText="Fig. 1. Map of the Indo-Pacific showing localities where Macrobrachium australe (Guérin-Méneville, 1838 in Guérin-Méneville 18291838) (black area) and M. ustulatum (Nobili, 1899) (red area) were collected and/or recorded. Capitalized locality names correspond to the 7 localities sampled for this study. Non-capitalized locality names correspond to the localities reported from the literature. Stars shows the type localities of the synonyms of M. australe (black stars) and M. ustulatum (red star)." figureDoi="http://doi.org/10.5281/zenodo.3824731" httpUri="https://zenodo.org/record/3824731/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0FB40BDA2737FFECFF1FF9A5FCA65853" pageId="15" pageNumber="16" type="biology_ecology">
<paragraph id="471158512737FFECFF1FF9A5FEBB5974" blockId="15.[189,285,1625,1651]" box="[189,285,1625,1651]" pageId="15" pageNumber="16">
<heading id="1C59EF3D2737FFECFF1FF9A5FEBB5974" bold="true" box="[189,285,1625,1651]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="75DA84432737FFECFF1FF9A5FEBB5974" bold="true" box="[189,285,1625,1651]" pageId="15" pageNumber="16">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="471158512737FFECFF1FF97BFC1B59C2" blockId="15.[189,1399,1671,1733]" pageId="15" pageNumber="16">Lotic habitats in the middle course of rivers, clear and oxygenated waters, sometimes above waterfalls. Rocky substrates, boulders, pebbles and pools in countercurrents.</paragraph>
<paragraph id="471158512737FFECFF1FF914FED55804" blockId="15.[189,371,1768,1795]" box="[189,371,1768,1795]" pageId="15" pageNumber="16">
<heading id="1C59EF3D2737FFECFF1FF914FED55804" bold="true" box="[189,371,1768,1795]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="75DA84432737FFECFF1FF914FED55804" bold="true" box="[189,371,1768,1795]" pageId="15" pageNumber="16">Color patterns</emphasis>
</heading>
</paragraph>
<paragraph id="471158512737FFECFF1FF8EBFCA65853" blockId="15.[189,1399,1815,1877]" pageId="15" pageNumber="16">
Body colors (
<figureCitation id="DF9544D42737FFECFEC6F8EBFE655836" box="[356,451,1815,1841]" captionStart="Fig" captionStartId="14.[189,232,1962,1988]" captionTargetBox="[215,1375,265,1928]" captionTargetId="figure@14.[185,1401,239,1936]" captionTargetPageId="14" captionText="Fig. 4. A. Live coloration of Macrobrachium australe (Guérin-Méneville, 1838 in Guérin-Méneville 18291838) (photo: E. Vigneux). B. Live coloration of M. ustulatum (Nobili, 1899) (photo: P. Keith)." figureDoi="http://doi.org/10.5281/zenodo.3824737" httpUri="https://zenodo.org/record/3824737/files/figure.png" pageId="15" pageNumber="16">Fig. 4B</figureCitation>
) tend to match the substrate, varying from black to brownish red, with three reddish stripes on each side of the cephalothorax.
</paragraph>
</subSubSection>
<subSubSection id="0FB40BDA2737FFF2FF1FF884FE845E4E" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="discussion">
<paragraph id="471158512737FFECFF1FF884FE895895" blockId="15.[189,303,1912,1938]" box="[189,303,1912,1938]" pageId="15" pageNumber="16">
<heading id="1C59EF3D2737FFECFF1FF884FE895895" bold="true" box="[189,303,1912,1938]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="75DA84432737FFECFF1FF884FE895895" bold="true" box="[189,303,1912,1938]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="471158512737FFECFF1FF85AFAD058E3" blockId="15.[189,1399,1958,2020]" pageId="15" pageNumber="16">
Some specimens originally assigned to
<taxonomicName id="80AE23D22737FFECFD07F85BFCB758C7" authorityName="von Martens" authorityYear="1868" box="[677,785,1958,1984]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="75DA84432737FFECFD07F85BFCB758C7" box="[677,785,1958,1984]" italics="true" pageId="15" pageNumber="16">P. dispar</emphasis>
</taxonomicName>
may correspond to
<taxonomicName id="80AE23D22737FFECFBAFF85BFB0B58C7" baseAuthorityName="Nobili" baseAuthorityYear="1899" box="[1037,1197,1958,1984]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ustulatum">
<emphasis id="75DA84432737FFECFBAFF85BFB0B58C7" box="[1037,1197,1958,1984]" italics="true" pageId="15" pageNumber="16">M. ustulatum</emphasis>
</taxonomicName>
. This applies to
<bibRefCitation id="233F25A02737FFECFF1FF835FE3C58E3" author="de Man" box="[189,410,1993,2020]" firstAuthor="de Man" pageId="15" pageNumber="16" pagination="265 - 527" refId="ref10613" refString="De Man J. G. 1892. Decapoden des Indischen Archipels. In: Weber M. (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien: 265 - 527. E. J. Brill, Leiden, the Netherlands. https: // doi. org / 10.5962 / bhl. title. 10298" type="book chapter" year="1892">de Man 1892: 435</bibRefCitation>
, pl. 26, fig. 44d (Salayar Island,
<collectingCountry id="3FB918C12737FFECFC82F835FC3E58E3" box="[800,920,1993,2020]" name="Indonesia" pageId="15" pageNumber="16">Indonesia</collectingCountry>
);
<bibRefCitation id="233F25A02737FFECFC0FF835FB2158E3" author="de Man" box="[941,1159,1993,2020]" firstAuthor="de Man" pageId="15" pageNumber="16" pagination="284 - 310" refId="ref10672" refString="De Man J. G. 1893. Report on the Podophthalmous Crustacea, collected in the year 1891 by Dr H. Ten Kate in some islands of the Malay Archipelago. Notes from the Leyden Museum 15: 284 - 310." type="journal article" year="1893">de Man 1893: 304</bibRefCitation>
(Flores,
<collectingCountry id="3FB918C12737FFECFB50F835FACC58E3" box="[1266,1386,1993,2020]" name="Indonesia" pageId="15" pageNumber="16">Indonesia</collectingCountry>
);
</paragraph>
<caption id="13D108D92728FFF3FF1FF8BEFB0558C0" ID-DOI="http://doi.org/10.5281/zenodo.3824739" ID-Zenodo-Dep="3824739" httpUri="https://zenodo.org/record/3824739/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1858,1884]" targetBox="[189,1399,268,1819]" targetPageId="16">
<paragraph id="471158512728FFF3FF1FF8BEFB0558C0" blockId="16.[189,1399,1858,1991]" pageId="16" pageNumber="17">
<emphasis id="75DA84432728FFF3FF1FF8BEFEAC585A" bold="true" box="[189,266,1858,1885]" pageId="16" pageNumber="17">Fig. 5.</emphasis>
<taxonomicName id="80AE23D22728FFF3FEB2F8BFFD56585A" authority="(Nobili, 1899)" baseAuthorityName="Nobili" baseAuthorityYear="1899" box="[272,752,1858,1885]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ustulatus">
<emphasis id="75DA84432728FFF3FEB2F8BFFD99585A" box="[272,575,1859,1885]" italics="true" pageId="16" pageNumber="17">Macrobrachium ustulatus</emphasis>
(
<bibRefCitation id="233F25A02728FFF3FDECF8BEFD41585A" author="Nobili" box="[590,743,1858,1885]" firstAuthor="Nobili" pageId="16" pageNumber="17" pagination="230 - 282" refId="ref11537" refString="Nobili G. 1899. Contribuzioni alla conoscenza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali del Museo civico di Storia naturale di Genova, Series 2 20 (40): 230 - 282." type="journal article" year="1899">Nobili, 1899</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="75DA84432728FFF3FCB2F8BEFCCC585A" bold="true" box="[784,874,1858,1885]" pageId="16" pageNumber="17">AB, E</emphasis>
. MNHN-IU-2013-13202.
<emphasis id="75DA84432728FFF3FB3DF8BEFB12585B" bold="true" box="[1183,1204,1858,1884]" pageId="16" pageNumber="17">A</emphasis>
. Cephalothorax.
<emphasis id="75DA84432728FFF3FF1FF89AFF775887" bold="true" box="[189,209,1894,1920]" pageId="16" pageNumber="17">B</emphasis>
. Epistome.
<emphasis id="75DA84432728FFF3FEC0F89AFED05887" bold="true" box="[354,374,1894,1920]" pageId="16" pageNumber="17">E</emphasis>
. Major second pereiopod finger.
<emphasis id="75DA84432728FFF3FC82F89AFCFB5887" bold="true" box="[800,861,1894,1920]" pageId="16" pageNumber="17">C, G</emphasis>
. MNHN-IU-2013-13201.
<emphasis id="75DA84432728FFF3FB3BF89AFB085887" bold="true" box="[1177,1198,1894,1920]" pageId="16" pageNumber="17">C</emphasis>
. Fourth thoracic sternite.
<emphasis id="75DA84432728FFF3FE82F875FE9158A4" bold="true" box="[288,311,1929,1955]" pageId="16" pageNumber="17">G</emphasis>
. Minor second pereiopod finger.
<emphasis id="75DA84432728FFF3FD68F875FD5958A4" bold="true" box="[714,767,1929,1955]" pageId="16" pageNumber="17">D, F</emphasis>
. MNHN-IU-2013-13203.
<emphasis id="75DA84432728FFF3FB91F875FBEE58A4" bold="true" box="[1075,1096,1929,1955]" pageId="16" pageNumber="17">D</emphasis>
. Major second pereiopod.
<emphasis id="75DA84432728FFF3FF1FF850FF6958C1" bold="true" box="[189,207,1964,1990]" pageId="16" pageNumber="17">F</emphasis>
. Minor second pereiopod. Scale bars: A, E, G = 2 mm; BC = 1 mm; D, F = 4 mm.
</paragraph>
</caption>
<paragraph id="471158512729FFF2FF1FFEF7FE845E4E" blockId="17.[189,1398,267,329]" pageId="17" pageNumber="18">
<bibRefCitation id="233F25A02729FFF2FF1FFEF7FE3A5E22" author="de Man" box="[189,412,267,294]" firstAuthor="de Man" pageId="17" pageNumber="18" pagination="467 - 929" refId="ref10716" refString="De Man J. G. 1902. Die von Herrn Professor Kukenthal in indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal W. (ed.) Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo: im Auftrage der Senckenbergischen Naturforschenden Gesellschaft: 467 - 929. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main. https: // doi. org / 10.5962 / bhl. title. 10409" type="book chapter" year="1902">de Man 1902: 766</bibRefCitation>
(Halmahera,
<collectingCountry id="3FB918C12729FFF2FDE1FEF7FD1D5E21" box="[579,699,267,294]" name="Indonesia" pageId="17" pageNumber="18">Indonesia</collectingCountry>
); and
<bibRefCitation id="233F25A02729FFF2FCA4FEF7FC4E5E21" author="de Man" box="[774,1000,267,294]" firstAuthor="de Man" pageId="17" pageNumber="18" pagination="385 - 458" refId="ref10791" refString="De Man J. G. 1915. Zur Fauna von Nord-Neuguinea. Nach den Sammlungen von Dr. P. N. van Kampen und K. Gjellerup in den Jahren 1910 - 1911. Macrura. Zoologische Jahrbucher - Abteilung fur Systematik, Geographie und Biologie der Thiere 38: 385 - 458." type="journal article" year="1915">de Man 1915: 435</bibRefCitation>
, pl. 29, figs 1414a (
<collectingCountry id="3FB918C12729FFF2FB49FEF7FEB25E4E" name="Papua New Guinea" pageId="17" pageNumber="18">Papua New Guinea</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>