Add updates up until 2024-12-16 14:45:06

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@ -1,49 +1,50 @@
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<treatment id="03A47546FFF67C6E657C1C52FE22D114" ID-DOI="http://doi.org/10.5281/zenodo.13759603" ID-Zenodo-Dep="13759603" LSID="urn:lsid:plazi:treatment:03A47546FFF67C6E657C1C52FE22D114" httpUri="http://treatment.plazi.org/id/03A47546FFF67C6E657C1C52FE22D114" lastPageId="28" lastPageNumber="29" pageId="23" pageNumber="24">
<treatment id="03A47546FFF67C6E657C1C52FE22D114" ID-DOI="http://doi.org/10.5281/zenodo.13759603" ID-GBIF-Taxon="241065175" ID-Zenodo-Dep="13759603" LSID="urn:lsid:plazi:treatment:03A47546FFF67C6E657C1C52FE22D114" httpUri="http://treatment.plazi.org/id/03A47546FFF67C6E657C1C52FE22D114" lastPageId="28" lastPageNumber="29" pageId="23" pageNumber="24" scope_class="Insecta" scope_family="Trigonidiidae" scope_infraOrder="Gryllidea" scope_order="Orthoptera" scope_subFamily="Nemobiinae" scope_subOrder="Ensifera" scope_superFamily="Grylloidea" scope_tribe="Nemobiini">
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@ -217,7 +218,7 @@ must take female gender also.
<tableCitation id="C68FF1EBFFF67C6565A719E0FCACD4EB" box="[690,793,1684,1711]" captionStart="Table 1" captionStartId="9.[189,259,265,291]" captionTargetId="graphics-72@9.[189,1400,571,1843]" captionTargetPageId="9" captionText="Table 1. Genus Bobilla Otte &amp; Alexander, 1983. Summary of morphological differences between adults of the two New Zealand species, Bobilla nigrova (Swan, 1972) and Bobilla bigelowi (Swan, 1972). All abbreviations used here are explained in the Material and methods section. All counts and measurements are median values, followed by minimum and maximum values in brackets. N indicates the sample size." httpUri="http://table.plazi.org/id/DF7294D8FFE87C7B67A81E7DFAC2D3C8" pageId="23" pageNumber="24" tableUuid="DF7294D8FFE87C7B67A81E7DFAC2D3C8">Table S1</tableCitation>
)
</paragraph>
<materialsCitation id="3B65CE0DFFF67C6567FF19B7FC24D560" collectingDate="1970-05-21" collectionCode="NZAC" collectorName="D. I. Swan" country="New Zealand" elevation="14" latitude="-43.52318" location="Mid Canterbury" longLatPrecision="1" longitude="172.58565" municipality="Ilam" pageId="23" pageNumber="24" specimenCode="NZAC 03016349" specimenCount="1" specimenCount-male="1" stateProvince="Canterbury" typeStatus="holotype">
<materialsCitation id="3B65CE0DFFF67C6567FF19B7FC24D560" ID-GBIF-Occurrence="4939245383" collectingDate="1970-05-21" collectionCode="NZAC" collectorName="D. I. Swan" country="New Zealand" elevation="14" latitude="-43.52318" location="Mid Canterbury" longLatPrecision="1" longitude="172.58565" municipality="Ilam" pageId="23" pageNumber="24" specimenCode="NZAC 03016349" specimenCount="1" specimenCount-male="1" stateProvince="Canterbury" typeStatus="holotype">
<paragraph id="8BB2C450FFF67C6567FF19B7FEE9D499" blockId="23.[189,1398,1731,1828]" box="[234,348,1731,1757]" pageId="23" pageNumber="24">
<emphasis id="B9791842FFF67C6567FF19B7FEE9D499" bold="true" box="[234,348,1731,1757]" pageId="23" pageNumber="24">
<typeStatus id="54B67AF2FFF67C6567FF19B7FEE9D499" box="[234,348,1731,1757]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
@ -251,7 +252,7 @@ leg.; in grassland;
.
</paragraph>
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<materialsCitation id="3B65CE0DFFF67C6567FF183FFB6FD5CC" collectingDate="1970-05-21" collectionCode="NZAC" collectorName="D. I. Swan" country="New Zealand" elevation="14" latitude="-43.52318" location="NEW ZEALAND" longLatPrecision="1" longitude="172.58565" municipality="Mid" pageId="23" pageNumber="24" specimenCode="NZAC 03016448" specimenCount="66" specimenCount-female="41" specimenCount-male="25" stateProvince="Canterbury" typeStatus="paratype">
<materialsCitation id="3B65CE0DFFF67C6567FF183FFB6FD5CC" ID-GBIF-Occurrence="4939245327" collectingDate="1970-05-21" collectionCode="NZAC" collectorName="D. I. Swan" country="New Zealand" elevation="14" latitude="-43.52318" location="NEW ZEALAND" longLatPrecision="1" longitude="172.58565" municipality="Mid" pageId="23" pageNumber="24" specimenCode="NZAC 03016448" specimenCount="66" specimenCount-female="41" specimenCount-male="25" stateProvince="Canterbury" typeStatus="paratype">
<paragraph id="8BB2C450FFF67C6567FF183FFED2D521" blockId="23.[189,1398,1867,1964]" box="[234,359,1867,1893]" pageId="23" pageNumber="24">
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<typeStatus id="54B67AF2FFF67C6567FF183FFED2D521" box="[234,359,1867,1893]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
@ -266,7 +267,7 @@ leg.; in grassland;
, adults; same data as for holotype;
<specimenCode id="DBAB6C2BFFF67C656319181AFB6FD5CC" box="[1036,1242,1902,1928]" collectionCode="NZAC" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="23" pageNumber="24" type="Other institutional type">NZAC 03016448</specimenCode>
<materialsCitation id="3B65CE0DFFF67C6563E3181AFB72D5EF" collectingDate="1970-08-28" collectionCode="NZAC" collectorName="D. I. Swan" country="New Zealand" elevation="14" latitude="-43.52318" location="Ilam" longLatPrecision="1" longitude="172.58565" municipality="Mid" pageId="23" pageNumber="24" specimenCode="NZAC 03016361" specimenCount="1" specimenCount-male="1" stateProvince="Canterbury" typeStatus="paratype">
<materialsCitation id="3B65CE0DFFF67C6563E3181AFB72D5EF" ID-GBIF-Occurrence="4939245396" collectingDate="1970-08-28" collectionCode="NZAC" collectorName="D. I. Swan" country="New Zealand" elevation="14" latitude="-43.52318" location="Ilam" longLatPrecision="1" longitude="172.58565" municipality="Mid" pageId="23" pageNumber="24" specimenCode="NZAC 03016361" specimenCount="1" specimenCount-male="1" stateProvince="Canterbury" typeStatus="paratype">
<specimenCount id="9D0B0FD9FFF67C6563E3181AFA93D5CD" box="[1270,1318,1902,1929]" count="1" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
, adult; same data as for holotype;
<date id="FFB3E290FFF67C6566ED18E5FD2FD5EF" box="[504,666,1937,1963]" pageId="23" pageNumber="24" value="1970-08-28">
@ -284,7 +285,7 @@ leg.; in grassland;
<emphasis id="B9791842FFF97C6A67FF1E7CFE13D366" bold="true" box="[234,422,264,290]" pageId="24" pageNumber="25">Other material</emphasis>
</paragraph>
<paragraph id="8BB2C450FFF97C6A67A81E5FFD7BD500" blockId="24.[189,1399,264,1861]" pageId="24" pageNumber="25">
<materialsCitation id="3B65CE0DFFF97C6A67A81E5FFAA7D323" collectingDate="2024-04-14" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="200" latitude="-45.86207" location="Dunedin" longLatPrecision="1" longitude="170.53995" municipality="Dunedin" pageId="24" pageNumber="25" specimenCode="OR0409, OR0410, OR0411" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<materialsCitation id="3B65CE0DFFF97C6A67A81E5FFAA7D323" ID-GBIF-Occurrence="4939245442" collectingDate="2024-04-14" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="200" latitude="-45.86207" location="Dunedin" longLatPrecision="1" longitude="170.53995" municipality="Dunedin" pageId="24" pageNumber="25" specimenCode="OR0409, OR0410, OR0411" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<collectingCountry id="F31A84C0FFF97C6A67A81E5FFE26D301" box="[189,403,299,325]" name="New Zealand" pageId="24" pageNumber="25">NEW ZEALAND</collectingCountry>
<emphasis id="B9791842FFF97C6A66A01E5EFDDDD300" bold="true" box="[437,616,298,324]" pageId="24" pageNumber="25">
@ -319,7 +320,7 @@ leg.; grassland; trowel and jar;
<specimenCode id="DBAB6C2BFFF97C6A63301E39FAA7D323" box="[1061,1298,333,359]" pageId="24" pageNumber="25">OR0409 to OR0411</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A62391E39FC8AD3E8" collectingDate="2024-04-15" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="Central Otago" elevation="270" latitude="-45.83906" location="McGregors Hill" longLatPrecision="1" longitude="170.56956" municipality="Dunedin" pageId="24" pageNumber="25" specimenCode="OR0412, OR0413, OR0414, OR0415" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<materialsCitation id="3B65CE0DFFF97C6A62391E39FC8AD3E8" ID-GBIF-Occurrence="4939245368" collectingDate="2024-04-15" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Central Otago" elevation="270" latitude="-45.83906" location="McGregors Hill" longLatPrecision="1" longitude="170.56956" municipality="Dunedin" pageId="24" pageNumber="25" specimenCode="OR0412, OR0413, OR0414, OR0415" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="9D0B0FD9FFF97C6A62391E39FAC4D323" box="[1324,1393,333,359]" count="2" pageId="24" pageNumber="25" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D0B0FD9FFF97C6A67A81E1BFEB6D3CD" box="[189,259,367,393]" count="2" pageId="24" pageNumber="25" type="female">2 ♀♀</specimenCount>
@ -349,7 +350,7 @@ leg.; grassland;
<specimenCode id="DBAB6C2BFFF97C6A65521EE5FC89D3E8" box="[583,828,401,428]" pageId="24" pageNumber="25">OR0412 to OR0415</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A645D1EE6FE43D3AB" collectingDate="2024-01-31" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="Central Otago" elevation="160" latitude="-45.83906" location="Clutha Gold Trail" longLatPrecision="1" longitude="170.56956" municipality="Dunedin" pageId="24" pageNumber="25" specimenCode="OR0125, OR0126, OR0127, OR0128, OR0129, OR0130, OR0131, OR0132" specimenCount="8" specimenCount-female="3" specimenCount-male="5">
<materialsCitation id="3B65CE0DFFF97C6A645D1EE6FE43D3AB" ID-GBIF-Occurrence="4939245342" collectingDate="2024-01-31" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Central Otago" elevation="160" latitude="-45.83906" location="Clutha Gold Trail" longLatPrecision="1" longitude="170.56956" municipality="Dunedin" pageId="24" pageNumber="25" specimenCode="OR0125, OR0126, OR0127, OR0128, OR0129, OR0130, OR0131, OR0132" specimenCount="8" specimenCount-female="3" specimenCount-male="5">
<emphasis id="B9791842FFF97C6A64741EE5FBD5D3EF" bold="true" box="[865,1120,401,427]" pageId="24" pageNumber="25">
<collectingCounty id="62D3BCDCFFF97C6A64741EE5FBA2D3EF" box="[865,1047,401,427]" pageId="24" pageNumber="25">Central Otago</collectingCounty>
@ -383,7 +384,7 @@ leg.; grassland;
<specimenCode id="DBAB6C2BFFF97C6A66131EA1FE43D3AB" box="[262,502,469,496]" pageId="24" pageNumber="25">OR0125 to OR0132</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A65061EA1FAB2D056" collectingDate="2021-07-01" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="Pillar Range" elevation="750" latitude="-45.44488" location="Big Hut" longLatPrecision="1" longitude="170.09697" municipality="Rock" pageId="24" pageNumber="25" specimenCode="OR0478" specimenCount="1" specimenCount-male="1">
<materialsCitation id="3B65CE0DFFF97C6A65061EA1FAB2D056" ID-GBIF-Occurrence="4939245319" collectingDate="2021-07-01" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Pillar Range" elevation="750" latitude="-45.44488" location="Big Hut" longLatPrecision="1" longitude="170.09697" municipality="Rock" pageId="24" pageNumber="25" specimenCode="OR0478" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0B0FD9FFF97C6A65061EA1FDF1D3B4" box="[531,580,469,496]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
;
<location id="8ED2928BFFF97C6A65471EA1FD07D3B4" LSID="urn:lsid:plazi:treatment:03A47546FFF67C6E657C1C52FE22D114:8ED2928BFFF97C6A65471EA1FD07D3B4" box="[594,690,469,496]" county="Pillar Range" latitude="-45.44488" longLatPrecision="1" longitude="170.09697" municipality="Rock" name="Big Hut" pageId="24" pageNumber="25">Big Hut</location>
@ -413,7 +414,7 @@ leg.; mixed grass and snow;
<specimenCode id="DBAB6C2BFFF97C6A63B11E8CFAB2D056" box="[1188,1287,504,530]" collectionCode="MPN" pageId="24" pageNumber="25">OR0478</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A623D1E8CFD5ED012" collectingDate="2021-03-18" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" elevation="140" latitude="-45.24063" location="Earnscleugh Tailings" longLatPrecision="1" longitude="169.36047" municipality="Alexandra" pageId="24" pageNumber="25" specimenCode="OR0043, OR0044" specimenCount="2" specimenCount-female="2">
<materialsCitation id="3B65CE0DFFF97C6A623D1E8CFD5ED012" ID-GBIF-Occurrence="4939245331" collectingDate="2021-03-18" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="140" latitude="-45.24063" location="Earnscleugh Tailings" longLatPrecision="1" longitude="169.36047" municipality="Alexandra" pageId="24" pageNumber="25" specimenCode="OR0043, OR0044" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D0B0FD9FFF97C6A623D1E8CFADAD056" box="[1320,1391,504,530]" count="2" pageId="24" pageNumber="25" type="female">2 ♀♀</specimenCount>
;
<location id="8ED2928BFFF97C6A67A81D6DFE00D077" LSID="urn:lsid:plazi:treatment:03A47546FFF67C6E657C1C52FE22D114:8ED2928BFFF97C6A67A81D6DFE00D077" box="[189,437,537,563]" latitude="-45.24063" longLatPrecision="1" longitude="169.36047" municipality="Alexandra" name="Earnscleugh Tailings" pageId="24" pageNumber="25">Earnscleugh Tailings</location>
@ -443,7 +444,7 @@ leg.; dirt track;
<specimenCode id="DBAB6C2BFFF97C6A65951D48FD52D012" box="[640,743,572,598]" collectionCode="MPN" pageId="24" pageNumber="25">OR0044</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A65E01D48FAABD03C" collectingDate="2022-05-01" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" elevation="600" latitude="-45.24063" location="Lake Ohau" longLatPrecision="1" longitude="169.36047" municipality="Alexandra" pageId="24" pageNumber="25" specimenCode="OR110" specimenCount="1" specimenCount-female="1" stateProvince="Mackenzie">
<materialsCitation id="3B65CE0DFFF97C6A65E01D48FAABD03C" ID-GBIF-Occurrence="4939245418" collectingDate="2022-05-01" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="600" latitude="-45.24063" location="Lake Ohau" longLatPrecision="1" longitude="169.36047" municipality="Alexandra" pageId="24" pageNumber="25" specimenCode="OR110" specimenCount="1" specimenCount-female="1" stateProvince="Mackenzie">
<emphasis id="B9791842FFF97C6A641B1D4FFC50D011" bold="true" box="[782,997,571,597]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A641B1D4FFC20D011" box="[782,917,571,597]" pageId="24" pageNumber="25">Mackenzie</collectingRegion>
@ -475,7 +476,7 @@ leg.; grassland;
<specimenCode id="DBAB6C2BFFF97C6A63DF1D2AFAABD03C" box="[1226,1310,606,632]" collectionCode="MPN" pageId="24" pageNumber="25">OR110</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A622A1D2AFCE7D0F8" collectingDate="2023-02-24" collectingDateMax="2023-02-26" collectingDateMin="2023-02-24" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" elevation="620" latitude="-44.25812" location="Lake Ohau" longLatPrecision="1" longitude="169.81464" pageId="24" pageNumber="25" specimenCode="OR0108, OR0109, OR0111, OR0112" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<materialsCitation id="3B65CE0DFFF97C6A622A1D2AFCE7D0F8" ID-GBIF-Occurrence="4939245451" collectingDate="2023-02-24" collectingDateMax="2023-02-26" collectingDateMin="2023-02-24" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="620" latitude="-44.25812" location="Lake Ohau" longLatPrecision="1" longitude="169.81464" pageId="24" pageNumber="25" specimenCode="OR0108, OR0109, OR0111, OR0112" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="9D0B0FD9FFF97C6A622A1D2AFAC4D03C" box="[1343,1393,606,632]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0B0FD9FFF97C6A67A81DF4FEB5D0DE" box="[189,256,640,666]" count="3" pageId="24" pageNumber="25" type="female">3 ♀♀</specimenCount>
@ -509,7 +510,7 @@ leg.; grassland;
<specimenCode id="DBAB6C2BFFF97C6A65E51DD6FCE7D0F8" box="[752,850,674,700]" collectionCode="MPN" pageId="24" pageNumber="25">OR0112</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A64661DD6FB8BD09B" collectingDate="2021-04-04" collectionCode="MPN" collectorName="D. Hegg" elevation="720" latitude="-44.00842" location="Tekapo" longLatPrecision="1" longitude="170.49794" pageId="24" pageNumber="25" specimenCode="OR0045" specimenCount="1" specimenCount-male="1">
<materialsCitation id="3B65CE0DFFF97C6A64661DD6FB8BD09B" ID-GBIF-Occurrence="4939245386" collectingDate="2021-04-04" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="720" latitude="-44.00842" location="Tekapo" longLatPrecision="1" longitude="170.49794" pageId="24" pageNumber="25" specimenCode="OR0045" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0B0FD9FFF97C6A64661DD6FC12D0F9" box="[883,935,674,701]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
;
<location id="8ED2928BFFF97C6A64A21DD6FBA4D0F8" LSID="urn:lsid:plazi:treatment:03A47546FFF67C6E657C1C52FE22D114:8ED2928BFFF97C6A64A21DD6FBA4D0F8" box="[951,1041,674,700]" latitude="-44.00842" longLatPrecision="1" longitude="170.49794" name="Tekapo" pageId="24" pageNumber="25">Tekapo</location>
@ -533,7 +534,7 @@ leg.; in short grass; jar;
<specimenCode id="DBAB6C2BFFF97C6A64CE1DB0FB8BD09B" box="[987,1086,708,735]" collectionCode="MPN" pageId="24" pageNumber="25">OR0045</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A63481DB0FE1ED167" collectingDate="2022-02-17" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="South Canterbury" elevation="700" latitude="-44.05623" location="Patersons Terrace" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0134" specimenCount="1" specimenCount-male="1">
<materialsCitation id="3B65CE0DFFF97C6A63481DB0FE1ED167" ID-GBIF-Occurrence="4939245452" collectingDate="2022-02-17" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="South Canterbury" elevation="700" latitude="-44.05623" location="Patersons Terrace" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0134" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0B0FD9FFF97C6A63481DB0FB3BD09B" box="[1117,1166,708,735]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
;
<location id="8ED2928BFFF97C6A638B1DB1FAC6D09B" LSID="urn:lsid:plazi:treatment:03A47546FFF67C6E657C1C52FE22D114:8ED2928BFFF97C6A638B1DB1FAC6D09B" box="[1182,1395,709,735]" county="South Canterbury" latitude="-44.05623" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" name="Patersons Terrace" pageId="24" pageNumber="25">Patersons Terrace</location>
@ -561,7 +562,7 @@ leg.; on 4WD track;
<specimenCode id="DBAB6C2BFFF97C6A66551C7DFE12D167" box="[320,423,777,803]" collectionCode="MPN" pageId="24" pageNumber="25">OR0134</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A66A21C7DFAD5D101" collectingDate="2021-04-04" collectionCode="MPN" collectorName="D. Hegg" county="South Canterbury" elevation="300" latitude="-44.05623" location="Fairlie" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0451, OR0452, OR0453, OR0454, OR0455" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="South Canterbury">
<materialsCitation id="3B65CE0DFFF97C6A66A21C7DFAD5D101" ID-GBIF-Occurrence="4939245403" collectingDate="2021-04-04" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="South Canterbury" elevation="300" latitude="-44.05623" location="Fairlie" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0451, OR0452, OR0453, OR0454, OR0455" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="South Canterbury">
<emphasis id="B9791842FFF97C6A66C41C7CFCB5D166" bold="true" box="[465,768,776,802]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A66C41C7CFD0FD166" box="[465,698,776,802]" pageId="24" pageNumber="25">South Canterbury</collectingRegion>
@ -593,7 +594,7 @@ leg.; in short grass in picnic area; jar;
<specimenCode id="DBAB6C2BFFF97C6A637F1C5EFAE9D101" box="[1130,1372,810,837]" pageId="24" pageNumber="25">OR0451 to OR0455</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A627D1C5FFC66D1CD" collectingDate="2023-07-28" collectionCode="MPN" collectorName="D. Hegg" county="South Canterbury" elevation="630" latitude="-44.05623" location="Glenrock Stream" longLatPrecision="1" longitude="170.42581" municipality="Rakaia River" pageId="24" pageNumber="25" specimenCode="OR0136" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Mid Canterbury">
<materialsCitation id="3B65CE0DFFF97C6A627D1C5FFC66D1CD" ID-GBIF-Occurrence="4939245429" collectingDate="2023-07-28" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="South Canterbury" elevation="630" latitude="-44.05623" location="Glenrock Stream" longLatPrecision="1" longitude="170.42581" municipality="Rakaia River" pageId="24" pageNumber="25" specimenCode="OR0136" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Mid Canterbury">
<emphasis id="B9791842FFF97C6A67A81C38FE60D122" bold="true" box="[189,469,844,870]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A67A81C38FE3ED122" box="[189,395,844,870]" pageId="24" pageNumber="25">Mid Canterbury</collectingRegion>
@ -625,7 +626,7 @@ leg.; grassy 4WD track; jar;
<specimenCode id="DBAB6C2BFFF97C6A647D1C1BFC7AD1CD" box="[872,975,879,905]" collectionCode="MPN" pageId="24" pageNumber="25">OR0136</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A64CE1C1BFE17D189" collectingDate="2021-04-03" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="South Canterbury" elevation="520" latitude="-44.05623" location="Boyle River" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0046" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Buller">
<materialsCitation id="3B65CE0DFFF97C6A64CE1C1BFE17D189" ID-GBIF-Occurrence="4939245393" collectingDate="2021-04-03" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="South Canterbury" elevation="520" latitude="-44.05623" location="Boyle River" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0046" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Buller">
<emphasis id="B9791842FFF97C6A64E61C1BFB30D1CC" bold="true" box="[1011,1157,878,905]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A64E61C1BFBF4D1CD" box="[1011,1089,879,905]" pageId="24" pageNumber="25">Buller</collectingRegion>
@ -657,7 +658,7 @@ leg.; short grass on road verge;
<specimenCode id="DBAB6C2BFFF97C6A66221CC7FE2BD189" box="[311,414,947,973]" collectionCode="MPN" pageId="24" pageNumber="25">OR0046</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A66BC1CC0FB9BD1AB" collectingDate="2023-03-12" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="South Canterbury" elevation="270" latitude="-44.05623" location="Kahutara River" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0133" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Kaikoura">
<materialsCitation id="3B65CE0DFFF97C6A66BC1CC0FB9BD1AB" ID-GBIF-Occurrence="4939245405" collectingDate="2023-03-12" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="South Canterbury" elevation="270" latitude="-44.05623" location="Kahutara River" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0133" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Kaikoura">
<emphasis id="B9791842FFF97C6A66AA1CC7FDCBD189" bold="true" box="[447,638,947,973]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A66AA1CC7FD8DD189" box="[447,568,947,973]" pageId="24" pageNumber="25">Kaikōura</collectingRegion>
@ -689,15 +690,13 @@ leg.; on gravel road;
<specimenCode id="DBAB6C2BFFF97C6A64D61CA1FB9FD1AB" box="[963,1066,981,1007]" collectionCode="MPN" pageId="24" pageNumber="25">OR0133</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A632C1CA2FAA9D1AB" box="[1081,1308,981,1008]" collectingDate="2023-03-12" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="South Canterbury" elevation="270" latitude="-44.05623" location="Kahutara River" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0133" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Kaikoura">
<materialsCitation id="3B65CE0DFFF97C6A632C1CA2FDCFD670" ID-GBIF-Occurrence="4939245322" collectingDate="2023-03-12" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="South Canterbury" elevation="270" latitude="-44.05623" location="Kahutara River" longLatPrecision="1" longitude="170.42581" municipality="Tekapo" pageId="24" pageNumber="25" specimenCode="OR0133" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Kaikoura">
<emphasis id="B9791842FFF97C6A63461CA1FAA9D1AB" bold="true" box="[1107,1308,981,1007]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A63461CA1FB61D1AB" box="[1107,1236,981,1007]" pageId="24" pageNumber="25">Wanganui</collectingRegion>
(WI)
</emphasis>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A622B1CA1FDCFD670" collectingDate="2023-03-17" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" elevation="150" latitude="-39.94103" location="McPhersons Bush" longLatPrecision="1" longitude="175.33578" pageId="24" pageNumber="25" specimenCode="OR0147" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0B0FD9FFF97C6A622B1CA1FAC4D1B4" box="[1342,1393,981,1008]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
;
<location id="8ED2928BFFF97C6A67A81C83FE23D655" LSID="urn:lsid:plazi:treatment:03A47546FFF67C6E657C1C52FE22D114:8ED2928BFFF97C6A67A81C83FE23D655" box="[189,406,1015,1041]" latitude="-39.94103" longLatPrecision="1" longitude="175.33578" name="McPhersons Bush" pageId="24" pageNumber="25">McPhersons Bush</location>
@ -723,7 +722,7 @@ leg.; tall grass on road verge;
<specimenCode id="DBAB6C2BFFF97C6A651A1B6EFDC3D670" box="[527,630,1050,1076]" collectionCode="MPN" pageId="24" pageNumber="25">OR0147</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A65941B6EFE92D63C" collectingDate="2024-03-14" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" elevation="720" latitude="-39.94103" location="Waipawa Forks Hut" longLatPrecision="1" longitude="175.33578" pageId="24" pageNumber="25" specimenCode="OR0310, OR0311" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rangitikei">
<materialsCitation id="3B65CE0DFFF97C6A65941B6EFE92D63C" ID-GBIF-Occurrence="4939245361" collectingDate="2024-03-14" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="720" latitude="-39.94103" location="Waipawa Forks Hut" longLatPrecision="1" longitude="175.33578" pageId="24" pageNumber="25" specimenCode="OR0310, OR0311" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rangitikei">
<emphasis id="B9791842FFF97C6A65821B6DFCE0D677" bold="true" box="[663,853,1049,1075]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A65821B6DFCACD677" box="[663,793,1049,1075]" pageId="24" pageNumber="25">Rangitikei</collectingRegion>
@ -759,7 +758,7 @@ leg.; in lawn outside hut;
<specimenCode id="DBAB6C2BFFF97C6A67A81B2AFE96D63C" box="[189,291,1118,1144]" collectionCode="MPN" pageId="24" pageNumber="25">OR0311</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A66251B2AFB82D6DE" collectingDate="2023-05-13" collectionCode="MPN" collectorName="D. Hegg" elevation="650" latitude="-39.94103" location="Taupo" longLatPrecision="1" longitude="175.33578" pageId="24" pageNumber="25" specimenCode="OR0121, OR0122" specimenCount="2" specimenCount-female="2" specimenCount-male="1" stateProvince="Rangitikei">
<materialsCitation id="3B65CE0DFFF97C6A66251B2AFB82D6DE" ID-GBIF-Occurrence="4939245460" collectingDate="2023-05-13" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="650" latitude="-39.94103" location="Taupo" longLatPrecision="1" longitude="175.33578" pageId="24" pageNumber="25" specimenCode="OR0121, OR0122" specimenCount="2" specimenCount-female="2" specimenCount-male="1" stateProvince="Rangitikei">
<emphasis id="B9791842FFF97C6A665D1B2AFD2BD633" bold="true" box="[328,670,1117,1144]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A665D1B2AFE7FD63C" box="[328,458,1118,1144]" pageId="24" pageNumber="25">Rangitikei</collectingRegion>
@ -793,7 +792,7 @@ leg.; in road pull-out; jar;
<specimenCode id="DBAB6C2BFFF97C6A64DE1BF4FB87D6DE" box="[971,1074,1152,1178]" collectionCode="MPN" pageId="24" pageNumber="25">OR0122</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A632A1BF4FBA2D69A" collectingDate="2022-03-29" collectingDateMax="2022-03-30" collectingDateMin="2022-03-29" collectionCode="MPN" collectorName="D. Hegg" county="Bay of Plenty" elevation="50" latitude="-39.94103" location="Manganui Road" longLatPrecision="1" longitude="175.33578" municipality="Awakino" pageId="24" pageNumber="25" specimenCode="OR0113, OR0114, OR0115, OR0116" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Waikato">
<materialsCitation id="3B65CE0DFFF97C6A632A1BF4FBA2D69A" ID-GBIF-Occurrence="4939245463" collectingDate="2022-03-29" collectingDateMax="2022-03-30" collectingDateMin="2022-03-29" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Bay of Plenty" elevation="50" latitude="-39.94103" location="Manganui Road" longLatPrecision="1" longitude="175.33578" municipality="Awakino" pageId="24" pageNumber="25" specimenCode="OR0113, OR0114, OR0115, OR0116" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Waikato">
<emphasis id="B9791842FFF97C6A63431BF4FAA5D6DE" bold="true" box="[1110,1296,1152,1178]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A63431BF4FB75D6DE" box="[1110,1216,1152,1178]" pageId="24" pageNumber="25">Waikato</collectingRegion>
@ -827,7 +826,7 @@ leg.; tall grass on road verge; trowel and jar;
<specimenCode id="DBAB6C2BFFF97C6A64371BB0FBA6D69A" box="[802,1043,1220,1247]" pageId="24" pageNumber="25">OR0113 to OR0116</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A63351BB1FD81D767" collectingDate="2023-04-15" collectingDateMax="2022-03-30" collectingDateMin="2022-03-29" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" county="Bay of Plenty" elevation="540" latitude="-39.94103" location="Hurunui Hut" longLatPrecision="1" longitude="175.33578" municipality="Kaimai Range" pageId="24" pageNumber="25" specimenCode="OR0124" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Bay of Plenty">
<materialsCitation id="3B65CE0DFFF97C6A63351BB1FD81D767" ID-GBIF-Occurrence="4939245400" collectingDate="2023-04-15" collectingDateMax="2022-03-30" collectingDateMin="2022-03-29" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Bay of Plenty" elevation="540" latitude="-39.94103" location="Hurunui Hut" longLatPrecision="1" longitude="175.33578" municipality="Kaimai Range" pageId="24" pageNumber="25" specimenCode="OR0124" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Bay of Plenty">
<emphasis id="B9791842FFF97C6A63221BB0FA91D69A" bold="true" box="[1079,1316,1220,1246]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A63221BB0FB56D69A" box="[1079,1251,1220,1246]" pageId="24" pageNumber="25">Bay of Plenty</collectingRegion>
@ -861,7 +860,7 @@ leg.; in grass;
<specimenCode id="DBAB6C2BFFF97C6A66C51A7DFD81D767" box="[464,564,1289,1315]" collectionCode="MPN" pageId="24" pageNumber="25">OR0124</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A65581A7DFAEBD701" collectingDate="2021-04-18" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" elevation="550" latitude="-38.27295" location="Little Manganuku Saddle" longLatPrecision="1" longitude="177.38828" pageId="24" pageNumber="25" specimenCode="OR0443, OR0444, OR0445, OR0446, OR0447" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<materialsCitation id="3B65CE0DFFF97C6A65581A7DFAEBD701" ID-GBIF-Occurrence="4939245414" collectingDate="2021-04-18" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="550" latitude="-38.27295" location="Little Manganuku Saddle" longLatPrecision="1" longitude="177.38828" pageId="24" pageNumber="25" specimenCode="OR0443, OR0444, OR0445, OR0446, OR0447" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<specimenCount id="9D0B0FD9FFF97C6A65581A7DFD27D767" box="[589,658,1289,1315]" count="2" pageId="24" pageNumber="25" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D0B0FD9FFF97C6A658A1A7DFD57D767" box="[671,738,1289,1315]" count="3" pageId="24" pageNumber="25" type="female">3 ♀♀</specimenCount>
@ -889,7 +888,7 @@ leg.; on walking track;
<specimenCode id="DBAB6C2BFFF97C6A63701A5FFAEED701" box="[1125,1371,1323,1349]" pageId="24" pageNumber="25">OR0443 to OR0447</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A627D1A5FFDE7D7CD" collectingDate="2024-04-19" collectionCode="MPN" collectorName="D. Hegg" elevation="1020" latitude="-38.27295" location="Hikurangi" longLatPrecision="1" longitude="177.38828" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0416" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Gisborne">
<materialsCitation id="3B65CE0DFFF97C6A627D1A5FFDE7D7CD" ID-GBIF-Occurrence="4939245323" collectingDate="2024-04-19" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="1020" latitude="-38.27295" location="Hikurangi" longLatPrecision="1" longitude="177.38828" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0416" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Gisborne">
<emphasis id="B9791842FFF97C6A67A81A38FEC0D722" bold="true" box="[189,373,1356,1382]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A67A81A38FE84D722" box="[189,305,1356,1382]" pageId="24" pageNumber="25">Gisborne</collectingRegion>
@ -919,7 +918,7 @@ leg.; on 4WD track; jar;
<specimenCode id="DBAB6C2BFFF97C6A66F31A1BFDFBD7CD" box="[486,590,1391,1417]" collectionCode="MPN" pageId="24" pageNumber="25">OR0416</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A654D1A1BFB0FD7EF" collectingDate="2024-03-25" collectionCode="MPN" collectorName="D. Hegg" elevation="100" latitude="-38.27295" location="Coromandel" longLatPrecision="1" longitude="177.38828" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0343" specimenCount="1" specimenCount-female="3" specimenCount-male="1" stateProvince="Bay of Plenty">
<materialsCitation id="3B65CE0DFFF97C6A654D1A1BFB0FD7EF" ID-GBIF-Occurrence="4939245434" collectingDate="2024-03-25" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="100" latitude="-38.27295" location="Coromandel" longLatPrecision="1" longitude="177.38828" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0343" specimenCount="1" specimenCount-female="3" specimenCount-male="1" stateProvince="Bay of Plenty">
<emphasis id="B9791842FFF97C6A657B1A1BFB82D7CC" bold="true" box="[622,1079,1390,1417]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A657B1A1BFCA0D7CD" box="[622,789,1391,1417]" pageId="24" pageNumber="25">Bay of Plenty</collectingRegion>
@ -951,7 +950,7 @@ leg.; in lawn; jar;
<specimenCode id="DBAB6C2BFFF97C6A635A1AE5FB02D7EF" box="[1103,1207,1425,1451]" collectionCode="MPN" pageId="24" pageNumber="25">OR0343</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A63D61AE6FD3FD7AB" collectingDate="2021-04-13" collectionCode="MPN" collectorName="D. Hegg" elevation="10" latitude="-38.27295" location="Coromandel" longLatPrecision="1" longitude="177.38828" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0481" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Coromandel">
<materialsCitation id="3B65CE0DFFF97C6A63D61AE6FD3FD7AB" ID-GBIF-Occurrence="4939245347" collectingDate="2021-04-13" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="10" latitude="-38.27295" location="Coromandel" longLatPrecision="1" longitude="177.38828" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0481" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Coromandel">
<emphasis id="B9791842FFF97C6A63CE1AE5FF4FD789" bold="true" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A63CE1AE5FAC2D7EF" box="[1243,1399,1425,1451]" pageId="24" pageNumber="25">Coromandel</collectingRegion>
@ -981,7 +980,7 @@ leg.; short grass in picnic area; jar;
<specimenCode id="DBAB6C2BFFF97C6A65331AA1FD3FD7AB" box="[550,650,1493,1519]" collectionCode="MPN" pageId="24" pageNumber="25">OR0481</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A65B61AA1FAC2D7AB" box="[675,1399,1493,1520]" collectingDate="2024-04-06" collectionCode="MPN" collectorName="D. Hegg" elevation="10" latitude="-36.76217" location="Auckland" longLatPrecision="1" longitude="175.49142" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0398" specimenCount="1" specimenCount-male="1" stateProvince="Coromandel">
<materialsCitation id="3B65CE0DFFF97C6A65B61AA1FAC2D7AB" ID-GBIF-Occurrence="4939245374" box="[675,1399,1493,1520]" collectingDate="2024-04-06" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="10" latitude="-36.76217" location="Auckland" longLatPrecision="1" longitude="175.49142" municipality="Bay of Plenty" pageId="24" pageNumber="25" specimenCode="OR0398" specimenCount="1" specimenCount-male="1" stateProvince="Coromandel">
<specimenCount id="9D0B0FD9FFF97C6A65B61AA1FD66D7B4" box="[675,723,1493,1520]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
; same data as for preceding;
<date id="FFB3E290FFF97C6A633F1AA1FB0DD7AB" box="[1066,1208,1493,1519]" pageId="24" pageNumber="25" value="2024-04-06">
@ -992,7 +991,7 @@ leg.; short grass in picnic area; jar;
<specimenCode id="DBAB6C2BFFF97C6A62191AA1FAC6D7AB" box="[1292,1395,1493,1519]" collectionCode="MPN" pageId="24" pageNumber="25">OR0398</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A67A81A8CFBD2D470" collectingDate="2022-03-28" collectionCode="MPN" collectorName="D. Hegg" elevation="40" latitude="-36.76217" location="Avondale Motor Park" longLatPrecision="1" longitude="175.49142" municipality="Auckland" pageId="24" pageNumber="25" specimenCode="OR0123" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Auckland">
<materialsCitation id="3B65CE0DFFF97C6A67A81A8CFBD2D470" ID-GBIF-Occurrence="4939245358" collectingDate="2022-03-28" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="40" latitude="-36.76217" location="Avondale Motor Park" longLatPrecision="1" longitude="175.49142" municipality="Auckland" pageId="24" pageNumber="25" specimenCode="OR0123" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Auckland">
<emphasis id="B9791842FFF97C6A67C31A83FE2FD455" bold="true" box="[214,410,1527,1553]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A67C31A83FEE5D455" box="[214,336,1527,1553]" pageId="24" pageNumber="25">Auckland</collectingRegion>
@ -1024,7 +1023,7 @@ leg.; on sparsely vegetated ground; jar;
<specimenCode id="DBAB6C2BFFF97C6A6311196EFBD2D470" box="[1028,1127,1562,1588]" collectionCode="MPN" pageId="24" pageNumber="25">OR0123</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A6390196EFE0DD43C" collectingDate="2021-05-02" collectionCode="MPN" collectorName="Northland" elevation="20" latitude="-36.48248" location="Kaipara Harbour" longLatPrecision="1" longitude="174.45068" pageId="24" pageNumber="25" specimenCode="OR0475, OR0476, OR0477" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<materialsCitation id="3B65CE0DFFF97C6A6390196EFE0DD43C" ID-GBIF-Occurrence="4939245337" collectingDate="2021-05-02" collectionCode="MPN" collectorName="Northland" country="New Zealand" elevation="20" latitude="-36.48248" location="Kaipara Harbour" longLatPrecision="1" longitude="174.45068" pageId="24" pageNumber="25" specimenCode="OR0475, OR0476, OR0477" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D0B0FD9FFF97C6A6390196EFB7ED470" box="[1157,1227,1562,1588]" count="2" pageId="24" pageNumber="25" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D0B0FD9FFF97C6A63CF196EFABCD470" box="[1242,1289,1562,1588]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
@ -1048,7 +1047,7 @@ a.s.l.
<specimenCode id="DBAB6C2BFFF97C6A67A8192AFE00D43C" box="[189,437,1630,1657]" pageId="24" pageNumber="25">OR0475 to OR0477</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A66D6192AFC53D4DF" collectingDate="2022-03-23" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" elevation="100" latitude="-36.48248" location="Kerikeri" longLatPrecision="1" longitude="174.45068" pageId="24" pageNumber="25" specimenCode="OR0135" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Northland">
<materialsCitation id="3B65CE0DFFF97C6A66D6192AFC53D4DF" ID-GBIF-Occurrence="4939245407" collectingDate="2022-03-23" collectingMethod="insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="100" latitude="-36.48248" location="Kerikeri" longLatPrecision="1" longitude="174.45068" pageId="24" pageNumber="25" specimenCode="OR0135" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Northland">
<emphasis id="B9791842FFF97C6A66C9192AFD12D433" bold="true" box="[476,679,1629,1656]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A66C9192AFDEBD43C" box="[476,606,1630,1656]" pageId="24" pageNumber="25">Northland</collectingRegion>
@ -1080,7 +1079,7 @@ leg.; in tall grass;
<specimenCode id="DBAB6C2BFFF97C6A649619F4FC53D4DF" box="[899,998,1664,1691]" collectionCode="MPN" pageId="24" pageNumber="25">OR0135</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A631619F4FDA5D49B" collectingDate="2024-03-28" collectionCode="MPN" collectorName="D. Hegg" country="Switzerland" county="Chatham Islands" elevation="30" latitude="-34.82342" location="Maitai Bay" longLatPrecision="1" longitude="173.40619" municipality="Karikari Peninsula" pageId="24" pageNumber="25" specimenCode="OR0357, OR0358" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<materialsCitation id="3B65CE0DFFF97C6A631619F4FDA5D49B" ID-GBIF-Occurrence="4939245301" collectingDate="2024-03-28" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Chatham Islands" elevation="30" latitude="-34.82342" location="Maitai Bay" longLatPrecision="1" longitude="173.40619" municipality="Karikari Peninsula" pageId="24" pageNumber="25" specimenCode="OR0357, OR0358" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D0B0FD9FFF97C6A631619F4FB86D4DE" box="[1027,1075,1664,1690]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D0B0FD9FFF97C6A635719F4FBC4D4DE" box="[1090,1137,1664,1690]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
@ -1110,7 +1109,7 @@ leg.; in grass; trowel and jar;
<specimenCode id="DBAB6C2BFFF97C6A66B019B0FDB9D49B" box="[421,524,1732,1759]" collectionCode="MPN" pageId="24" pageNumber="25">OR0358</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A650219B1FAA1D544" collectingDate="2024-02-15" collectionCode="MPN" collectorName="D. Hegg" country="Switzerland" county="Chatham Islands" elevation="50" latitude="-34.82342" location="Canister Cove" longLatPrecision="1" longitude="173.40619" municipality="Pitt Island" pageId="24" pageNumber="25" specimenCode="OR0137" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Chatham Islands">
<materialsCitation id="3B65CE0DFFF97C6A650219B1FAA1D544" ID-GBIF-Occurrence="4939245351" collectingDate="2024-02-15" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Chatham Islands" elevation="50" latitude="-34.82342" location="Canister Cove" longLatPrecision="1" longitude="173.40619" municipality="Pitt Island" pageId="24" pageNumber="25" specimenCode="OR0137" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Chatham Islands">
<emphasis id="B9791842FFF97C6A653819B0FCFFD49A" bold="true" box="[557,842,1732,1758]" pageId="24" pageNumber="25">
<collectingRegion id="49C90AB2FFF97C6A653819B0FCB1D49A" box="[557,772,1732,1758]" pageId="24" pageNumber="25">Chatham Islands</collectingRegion>
@ -1142,7 +1141,7 @@ leg.; short ground cover; jar;
<specimenCode id="DBAB6C2BFFF97C6A63A41992FAA1D544" box="[1201,1300,1766,1792]" collectionCode="MPN" pageId="24" pageNumber="25">OR0137</specimenCode>
</materialsCitation>
<materialsCitation id="3B65CE0DFFF97C6A62381993FD7FD500" collectingDate="2024-02-16" collectionCode="MPN" collectorName="D. Hegg &amp; S. Purdie" country="Switzerland" elevation="120" latitude="-44.3235" location="Pitt Island" longLatPrecision="6" longitude="-176.2337" pageId="24" pageNumber="25" specimenCode="OR0138, OR0139, OR0140, OR0141, OR0142, OR0143, OR0144, OR0145, OR0146" specimenCount="9" specimenCount-female="2" specimenCount-male="7">
<materialsCitation id="3B65CE0DFFF97C6A62381993FD7FD500" ID-GBIF-Occurrence="4939245426" collectingDate="2024-02-16" collectionCode="MPN" collectorName="D. Hegg &amp; S. Purdie" country="New Zealand" elevation="120" latitude="-44.3235" location="Pitt Island" longLatPrecision="6" longitude="-176.2337" pageId="24" pageNumber="25" specimenCode="OR0138, OR0139, OR0140, OR0141, OR0142, OR0143, OR0144, OR0145, OR0146" specimenCount="9" specimenCount-female="2" specimenCount-male="7">
<specimenCount id="9D0B0FD9FFF97C6A62381993FAC4D545" box="[1325,1393,1767,1793]" count="7" pageId="24" pageNumber="25" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D0B0FD9FFF97C6A67A8187DFEB7D567" box="[189,258,1801,1827]" count="2" pageId="24" pageNumber="25" type="female">2 ♀♀</specimenCount>

View file

@ -1,355 +0,0 @@
<document id="C231E6CCF38CD4BD233AB213DA694830" ID-DOI="10.1093/zoolinnean/zlad190" ID-ISSN="0024-4082" ID-Zenodo-Dep="14422893" ID-ZooBank="83066B9-A377-4D41-813D-74BD818762F6XXXXX" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734029032058" checkinUser="plazi" docAuthor="Hsiao, Yun &amp; Oberprieler, Rolf G." docDate="2024" docId="965ED847FF81FF81FEB8F99DA8F427E6" docLanguage="en" docName="zlad190.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (2)" docSource="https://doi.org/10.1093/zoolinnean/zlad190" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Zimmiodes papuanus Hsiao &amp; Oberprieler, 2024, sp.nov." docType="treatment" docVersion="2" lastPageNumber="19" masterDocId="6A67A03FFF90FF93FF99FF8CA9672410" masterDocTitle="An integrative taxonomic and phylogenetic approach reveals a new genus of Australasian Cycas-pollinating weevils (Coleoptera: Curculionidae: Cossoninae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="18" updateTime="1734029438208" updateUser="ExternalLinkService">
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<mods:title id="408C6DE7D7958C1D20C63ADB5BAB3297">An integrative taxonomic and phylogenetic approach reveals a new genus of Australasian Cycas-pollinating weevils (Coleoptera: Curculionidae: Cossoninae)</mods:title>
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<mods:namePart id="6E9E8A1FE566B53C38A459F3D69565A7">Hsiao, Yun</mods:namePart>
<mods:affiliation id="82BDE160D71D0B9390599C6FF5182369">Institute of Ecology and Evolutionary Biology, National Taiwan University, Taipei, Taiwan &amp; Australian National Insect Collection, CSIRO, GPO Box 1700, Canberra, ACT 2601, Australia &amp; Division of Ecology and Evolution, Research School of Biology, The Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="E00596576A5BE10B5350A13FF9EB2F52">Oberprieler, Rolf G.</mods:namePart>
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<mods:title id="7EF1623733F1A201DAD1178F905129D1">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="6CD26E373168064C8AD931731154DAA9">2024</mods:date>
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<mods:title id="55578438BEEDCE3BF11CBA00F5079C55">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="CB2B022D8EDBE076F7D1F87A43635E1C">2024-01-04</mods:number>
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<mods:number id="1E1AA0EC46102D890FA6B1EA80B1B57F">202</mods:number>
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<mods:url id="3EFB38BBE692F1E9888734612457E488">https://doi.org/10.1093/zoolinnean/zlad190</mods:url>
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<subSubSection id="56ED3ADAFF81FF82FEB8F99DAB2D223C" box="[289,586,1553,1580]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="1E486951FF81FF82FEB8F99DAB2D223C" blockId="17.[289,586,1553,1580]" box="[289,586,1553,1580]" pageId="17" pageNumber="18">
<heading id="4500DE3DFF81FF82FEB8F99DAB2D223C" bold="true" box="[289,586,1553,1580]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<emphasis id="2C83B543FF81FF82FEB8F99DAB2D223C" bold="true" box="[289,586,1553,1580]" pageId="17" pageNumber="18">
<taxonomicName id="D9F712D2FF81FF82FEB8F99DA89D223C" authorityName="Hsiao &amp; Oberprieler" authorityYear="2024" box="[289,506,1553,1580]" class="Insecta" family="Curculionidae" genus="Zimmiodes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="papuanus" status="sp. nov.">
<emphasis id="2C83B543FF81FF82FEB8F99DA89D223C" bold="true" box="[289,506,1553,1580]" italics="true" pageId="17" pageNumber="18">Zimmiodes papuanus</emphasis>
</taxonomicName>
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</emphasis>
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<subSubSection id="56ED3ADAFF81FF81FF12F9CDA8F427E6" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="1E486951FF81FF82FF12F9CDAB86224B" blockId="17.[139,737,1601,1627]" box="[139,737,1601,1627]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FF12F9CDAB86224B" bold="true" box="[139,737,1601,1627]" pageId="17" pageNumber="18">
(
<figureCitation id="86CC75D4FF81FF82FF0FF9CDA864224B" box="[150,259,1601,1627]" captionStart="Figure 8" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[239,1364,146,1798]" captionTargetId="figure-8@10.[237,1366,144,1808]" captionTargetPageId="10" captionText="Figure 8. Habitus of Zimmiodes, dorsal view. A, Zimmiodes australis, male, holotype.B, Z. australis, female, paratype. C, Zimmiodes papuanus, male, holotype.D, Z. papuanus, female, paratype. Scale bars: 1.00 mm." pageId="17" pageNumber="18">Figs 8C, D</figureCitation>
,
<figureCitation id="86CC75D4FF81FF82FE94F9CDA82D224B" box="[269,330,1601,1627]" captionStart="Figure 9" captionStartId="11.[113,178,1836,1860]" captionTargetBox="[255,1316,146,1806]" captionTargetId="figure-6@11.[253,1319,144,1808]" captionTargetPageId="11" captionText="Figure 9. Habitus of Zimmiodes, lateral view. A, Zimmiodes australis, male, holotype. B, Z. australis, female, paratype. C, Zimmiodes papuanus, male, holotype. D, Z. papuanus, female, paratype. Scale bars: 1.00 mm." pageId="17" pageNumber="18">9C, D</figureCitation>
,
<figureCitation id="86CC75D4FF81FF82FECDF9CDA8FA224B" box="[340,413,1601,1627]" captionStart="Figure 10" captionStartId="12.[129,194,1836,1860]" captionTargetBox="[308,1284,146,1806]" captionTargetId="figure-8@12.[306,1296,144,1808]" captionTargetPageId="12" captionText="Figure 10. Diagnostic differences of antennae and forelegs of Zimmiodes australis and Zimmiodes papuanus. A, Z. australis, right antenna. B, Z. papuanus, right antenna. C, Z. australis, foreleg of male.D, Z. papuanus, foreleg of male.Scale bars: 0.20 mm." pageId="17" pageNumber="18">10B, D</figureCitation>
,
<figureCitation id="86CC75D4FF81FF82FE3EF9CDAB61224B" box="[423,518,1601,1627]" captionStart="Figure 11" captionStartId="13.[115,178,1449,1473]" captionTargetBox="[114,1457,145,1417]" captionTargetId="figure-341@13.[112,1459,144,1422]" captionTargetPageId="13" captionText="Figure 11.Terminalia ofZimmiodes. A, Zimmiodes australis, sternite VIII,male.B,Zimmiodes papuanus, sternite VIII,male.C,Z.australis, sternite VIII,female.D, Z.papuanus, sternite VIII, female.E,Z. australis, spiculumgastrale.F,Z. papuanus, spiculum gastrale.Scale bars:0.20 mm." pageId="17" pageNumber="18">11B, D, F</figureCitation>
,
<figureCitation id="86CC75D4FF81FF82FD89F9CDAB3C224B" box="[528,603,1601,1627]" captionStart="Figure 12" captionStartId="14.[129,194,1119,1143]" captionTargetBox="[130,1473,145,1090]" captionTargetId="figure-498@14.[128,1475,144,1092]" captionTargetPageId="14" captionText="Figure 12. Male genitalia (aedeagus) of Zimmiodes. A, Zimmiodes australis, dorsal view. B, Z. australis, ventral view.C, Z. australis, lateral view. D, Zimmiodes papuanus, dorsal view.E, Z. papuanus, ventral view.F, Z. papuanus, lateral view. Scale bar: 0.20 mm." pageId="17" pageNumber="18">12DF</figureCitation>
,
<figureCitation id="86CC75D4FF81FF82FDFCF9CDABD7224B" box="[613,688,1601,1627]" captionStart="Figure 13" captionStartId="15.[113,178,1836,1860]" captionTargetBox="[183,1384,145,1806]" captionTargetId="figure-7@15.[182,1390,144,1808]" captionTargetPageId="15" captionText="Figure 13. Female genitalia of Zimmiodes. A, Zimmiodes australis, ovipositor.B, Z. australis, spermatheca with gland. C, Zimmiodes papuanus, ovipositor.D, Z. papuanus, spermatheca with gland.Scale bars: 0.20 mm." pageId="17" pageNumber="18">13C, D</figureCitation>
,
<figureCitation id="86CC75D4FF81FF82FD23F9CDABB1224B" box="[698,726,1601,1627]" captionStart="Figure 14" captionStartId="16.[129,194,1236,1260]" captionTargetBox="[129,1473,144,1208]" captionTargetId="figure-393@16.[129,1473,144,1208]" captionTargetPageId="16" captionText="Figure 14. Geographical distribution of Zimmiodes in Australia and Papua New Guinea." pageId="17" pageNumber="18">14</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="1E486951FF81FF82FFE8F90EA87022A9" blockId="17.[113,762,1666,1721]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FFE8F90EA9D62289" box="[113,177,1666,1689]" italics="true" pageId="17" pageNumber="18">LSID:</emphasis>
<uri id="6A666553FF81FF82FF25F90EA87022A9" pageId="17" pageNumber="18">
urn:lsid:zoobank.org:act:
<uuid id="6A515384FF81FF82FE25F90EA87022A9" pageId="17" pageNumber="18">0A0B8FB6-6DE9-406F-88F3- 7E9940C88B15</uuid>
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<paragraph id="1E486951FF81FF82FEE2F955A89622E2" blockId="17.[379,497,1752,1778]" box="[379,497,1752,1778]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FEE2F955A89622E2" box="[379,497,1752,1778]" italics="true" pageId="17" pageNumber="18">Type locality</emphasis>
</paragraph>
<paragraph id="1E486951FF81FF82FFE8F973ABE42307" blockId="17.[113,643,1791,1815]" box="[113,643,1791,1815]" pageId="17" pageNumber="18">
Erap, Markham Valley,
<collectingRegion id="DC33A7B3FF81FF82FEC2F973A8C82307" box="[347,431,1791,1815]" country="Papua New Guinea" name="Morobe" pageId="17" pageNumber="18">Morobe</collectingRegion>
,
<collectingCountry id="66E029C1FF81FF82FE20F973ABE72307" box="[441,640,1791,1815]" name="Papua New Guinea" pageId="17" pageNumber="18">Papua New Guinea</collectingCountry>
.
</paragraph>
<paragraph id="1E486951FF81FF82FEECF8BBA8902340" blockId="17.[373,503,1846,1872]" box="[373,503,1846,1872]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FEECF8BBA8902340" box="[373,503,1846,1872]" italics="true" pageId="17" pageNumber="18">Type material</emphasis>
</paragraph>
<paragraph id="1E486951FF81FF82FFE8F8D1AAA124D7" blockId="17.[113,762,1885,1972]" lastBlockId="17.[811,1459,144,293]" pageId="17" pageNumber="18">
<materialsCitation id="AE9F630CFF81FF82FFE8F8D1AA2324B8" collectingDate="1956-08-17" collectionCode="NEW, COM, INST, COLL" collectorName="J. H. Ardley &amp; From" country="Guinea-Bissau" location="Markham Valley" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="C14CD7F3FF81FF82FFE8F8D1A9B22365" box="[113,213,1885,1909]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
,
<specimenCount id="08F1A2D8FF81FF82FF7DF8ECA9992364" box="[228,254,1888,1908]" count="1" pageId="17" pageNumber="18" type="male"></specimenCount>
:
<collectionCode id="78E6F194FF81FF82FE8DF8D1A8362365" box="[276,337,1885,1909]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="17" pageNumber="18">NEW</collectionCode>
<collectingCountry id="66E029C1FF81FF82FEC5F8D1A8A72365" box="[348,448,1885,1909]" name="Guinea-Bissau" pageId="17" pageNumber="18">GUINEA</collectingCountry>
/
<location id="1B283F8AFF81FF82FE78F8D1ABE92365" LSID="urn:lsid:plazi:treatment:965ED847FF81FF81FEB8F99DA8F427E6:1B283F8AFF81FF82FE78F8D1ABE92365" box="[481,654,1885,1909]" country="Guinea-Bissau" name="Markham Valley" pageId="17" pageNumber="18">Markham Valley</location>
/ Erap. /
<date id="6A494F91FF81FF82FFE8F8F0A9932384" box="[113,244,1916,1940]" pageId="17" pageNumber="18" value="1956-08-17">
<collectingDate id="7A0DB679FF81FF82FFE8F8F0A9932384" box="[113,244,1916,1940]" pageId="17" pageNumber="18" value="1956-08-17">17.VIII.1956</collectingDate>
</date>
W. /
<collectorName id="B3020C87FF81FF82FEAEF8F1A8CA2384" box="[311,429,1916,1940]" pageId="17" pageNumber="18">J.H. Ardley</collectorName>
//
<collectorName id="B3020C87FF81FF82FE41F8F1AB772384" box="[472,528,1917,1940]" pageId="17" pageNumber="18">From</collectorName>
<specimenCount id="08F1A2D8FF81FF82FD83F8F3AB532383" box="[538,564,1919,1939]" count="1" pageId="17" pageNumber="18" type="male"></specimenCount>
cone / of
<taxonomicName id="D9F712D2FF81FF82FD35F8F1AB822384" box="[684,741,1917,1940]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="2C83B543FF81FF82FD35F8F1AB822384" box="[684,741,1917,1940]" italics="true" pageId="17" pageNumber="18">Cycas</emphasis>
</taxonomicName>
/
<emphasis id="2C83B543FF81FF82FFE8F810A9A123A4" box="[113,198,1948,1972]" italics="true" pageId="17" pageNumber="18">circinalis</emphasis>
/
<taxonomicName id="D9F712D2FF81FF82FF46F810A85223A3" box="[223,309,1948,1971]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="papuana">
<emphasis id="2C83B543FF81FF82FF46F810A85223A3" box="[223,309,1948,1971]" italics="true" pageId="17" pageNumber="18">papuana</emphasis>
</taxonomicName>
//
<collectionCode id="78E6F194FF81FF82FEC3F810A8FD23A4" box="[346,410,1948,1972]" country="Sri Lanka" httpUri="http://grbio.org/cool/zeq7-2jnj" name="Colombo Museum" pageId="17" pageNumber="18">COM</collectionCode>
.
<collectionCode id="78E6F194FF81FF82FE3AF810A8B923A4" box="[419,478,1948,1972]" pageId="17" pageNumber="18">INST</collectionCode>
. ENT. /
<collectionCode id="78E6F194FF81FF82FDAFF810AB1A23A4" box="[566,637,1948,1972]" pageId="17" pageNumber="18">COLL</collectionCode>
. NO. 14956 //
</materialsCitation>
<materialsCitation id="AE9F630CFF81FF82FCD3FF1CAAA524D7" collectionCode="ANIC" collectorName="Hsiao &amp; Oberprieler" country="Guinea-Bissau" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<typeStatus id="C14CD7F3FF81FF82FCD3FF1CAAB224B8" box="[842,981,144,168]" pageId="17" pageNumber="18" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="D9F712D2FF81FF82FC76FF1CADA624B8" authorityName="Hsiao &amp; Oberprieler" authorityYear="2024" box="[1007,1217,144,168]" class="Insecta" family="Curculionidae" genus="Zimmiodes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="2C83B543FF81FF82FC76FF1CADA624B8" box="[1007,1217,144,168]" italics="true" pageId="17" pageNumber="18">Zimmiodes papuanus</emphasis>
</taxonomicName>
/
<collectorName id="B3020C87FF81FF82FB40FF1CAC7224B8" box="[1241,1301,144,168]" pageId="17" pageNumber="18">Hsiao</collectorName>
&amp;
<collectorName id="B3020C87FF81FF82FAACFF1CACC824B8" box="[1333,1455,144,168]" pageId="17" pageNumber="18">Oberprieler</collectorName>
, 2024 (
<collectionCode id="78E6F194FF81FF82FCEBFF23AADD24D7" box="[882,954,175,199]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="17" pageNumber="18">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="1E486951FF81FF82FCDCFF43AAE12535" blockId="17.[811,1459,144,293]" pageId="17" pageNumber="18">
<materialsCitation id="AE9F630CFF81FF82FCDCFF43AAE42535" collectionCode="ANIC" collectorName="Hsiao &amp; Oberprieler" pageId="17" pageNumber="18" specimenCount="8" specimenCount-female="6" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="C14CD7F3FF81FF82FCDCFF43AACF24F7" box="[837,936,207,231]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
(all labelled
<typeStatus id="C14CD7F3FF81FF82FBAAFF43ADD424F7" box="[1075,1203,207,231]" pageId="17" pageNumber="18" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="D9F712D2FF81FF82FB56FF43ACC624F6" authorityName="Hsiao &amp; Oberprieler" authorityYear="2024" box="[1231,1441,207,231]" class="Insecta" family="Curculionidae" genus="Zimmiodes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="2C83B543FF81FF82FB56FF43ACC624F6" box="[1231,1441,207,231]" italics="true" pageId="17" pageNumber="18">Zimmiodes papuanus</emphasis>
</taxonomicName>
/
<collectorName id="B3020C87FF81FF82FCB5FF62AA0F2516" box="[812,872,238,262]" pageId="17" pageNumber="18">Hsiao</collectorName>
&amp;
<collectorName id="B3020C87FF81FF82FC1EFF62AD662516" box="[903,1025,238,262]" pageId="17" pageNumber="18">Oberprieler</collectorName>
, 2024):
<specimenCount id="08F1A2D8FF81FF82FBC4FF62ADC42515" box="[1117,1187,238,262]" count="2" pageId="17" pageNumber="18" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="08F1A2D8FF81FF82FB36FF62AD912515" box="[1199,1270,238,262]" count="6" pageId="17" pageNumber="18" type="female">6 ♀♀</specimenCount>
, same as
<typeStatus id="C14CD7F3FF81FF82FACFFF62ACD52516" box="[1366,1458,238,262]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(
<collectionCode id="78E6F194FF81FF82FCAAFE81AA1C2535" box="[819,891,269,293]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="17" pageNumber="18">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="1E486951FF81FF82FBB0FEC9ADD3254F" blockId="17.[810,1459,325,419]" box="[1065,1204,325,351]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FBB0FEC9ADD3254F" box="[1065,1204,325,351]" italics="true" pageId="17" pageNumber="18">Other material</emphasis>
</paragraph>
<paragraph id="1E486951FF81FF82FCB3FEE7ADD925B3" blockId="17.[810,1459,325,419]" pageId="17" pageNumber="18">
<materialsCitation id="AE9F630CFF81FF82FCB3FEE7ADDE25B3" collectingDate="1955-07-27" collectionCode="ANIC" collectorName="Hoogland" location="Ramu Valley" municipality="Faita" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Madang">
<collectingRegion id="DC33A7B3FF81FF82FCB3FEE7AAB42593" box="[810,979,363,387]" country="Papua New Guinea" name="Madang" pageId="17" pageNumber="18">Madang District</collectingRegion>
,
<collectingMunicipality id="FE2CF32BFF81FF82FC45FEE7AD692593" box="[988,1038,363,387]" pageId="17" pageNumber="18">Faita</collectingMunicipality>
airstrip,
<location id="1B283F8AFF81FF82FBFFFEE0AD802593" LSID="urn:lsid:plazi:treatment:965ED847FF81FF81FEB8F99DA8F427E6:1B283F8AFF81FF82FBFFFEE0AD802593" box="[1126,1255,363,387]" municipality="Faita" name="Ramu Valley" pageId="17" pageNumber="18" stateProvince="Madang">Ramu Valley</location>
,
<date id="6A494F91FF81FF82FB69FEE7AC162593" box="[1264,1393,363,387]" pageId="17" pageNumber="18" value="1955-07-27">
<collectingDate id="7A0DB679FF81FF82FB69FEE7AC162593" box="[1264,1393,363,387]" pageId="17" pageNumber="18" value="1955-07-27">27 July 1955</collectingDate>
</date>
,
<taxonomicName id="D9F712D2FF81FF82FAE3FEE0AA2025B2" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2C83B543FF81FF82FAE3FEE0ACD42593" box="[1402,1459,364,387]" italics="true" pageId="17" pageNumber="18">Cycas</emphasis>
sp.
</taxonomicName>
,
<collectorName id="B3020C87FF81FF82FCC9FE07AAD025B3" box="[848,951,395,419]" pageId="17" pageNumber="18">Hoogland</collectorName>
leg. (
<specimenCount id="08F1A2D8FF81FF82FC69FE07AD7B25B2" box="[1008,1052,395,419]" count="1" pageId="17" pageNumber="18" type="male">1 ♂</specimenCount>
, 3 exx.,
<collectionCode id="78E6F194FF81FF82FBF4FE07ADD525B3" box="[1133,1202,395,419]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="17" pageNumber="18">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="1E486951FF81FF82FBD9FE4FADFA25CC" blockId="17.[1088,1181,451,476]" box="[1088,1181,451,476]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FBD9FE4FADFA25CC" box="[1088,1181,451,476]" italics="true" pageId="17" pageNumber="18">Diagnosis</emphasis>
</paragraph>
<paragraph id="1E486951FF81FF82FCB3FE65ADD12630" blockId="17.[810,1459,489,544]" pageId="17" pageNumber="18">
The species is readily distinguishable from
<taxonomicName id="D9F712D2FF81FF82FB70FE65AC302611" baseAuthorityName="Carter" baseAuthorityYear="1932" box="[1257,1367,489,513]" class="Insecta" family="Tenebrionidae" genus="Ulomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="2C83B543FF81FF82FB70FE65AC302611" box="[1257,1367,489,513]" italics="true" pageId="17" pageNumber="18">Z. australis</emphasis>
</taxonomicName>
as specified in the diagnosis of the latter above.
</paragraph>
<paragraph id="1E486951FF81FF82FBA1FDCCADC32649" blockId="17.[1080,1188,576,601]" box="[1080,1188,576,601]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FBA1FDCCADC32649" box="[1080,1188,576,601]" italics="true" pageId="17" pageNumber="18">Description</emphasis>
</paragraph>
<paragraph id="1E486951FF81FF82FCB3FDEAAD5926AD" blockId="17.[810,1459,614,701]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FCB3FDEAAAA2266E" box="[810,965,614,638]" italics="true" pageId="17" pageNumber="18">Shape and size:</emphasis>
Body slender and elongate (
<figureCitation id="86CC75D4FF81FF82FB76FDEAAC3C266D" box="[1263,1371,614,638]" captionStart="Figure 8" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[239,1364,146,1798]" captionTargetId="figure-8@10.[237,1366,144,1808]" captionTargetPageId="10" captionText="Figure 8. Habitus of Zimmiodes, dorsal view. A, Zimmiodes australis, male, holotype.B, Z. australis, female, paratype. C, Zimmiodes papuanus, male, holotype.D, Z. papuanus, female, paratype. Scale bars: 1.00 mm." pageId="17" pageNumber="18">Fig. 8C, D</figureCitation>
), length
<quantity id="D90FC4B4FF81FF82FCB5FD09AACF268D" box="[812,936,645,669]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.4" metricValueMin="2.6" pageId="17" pageNumber="18" unit="mm" value="3.0" valueMax="3.4" valueMin="2.6">2.63.4 mm</quantity>
in both sexes, width ~0.33× length, weakly convex in lateral view (
<figureCitation id="86CC75D4FF81FF82FC5FFD29AD4926AC" box="[966,1070,677,701]" captionStart="Figure 9" captionStartId="11.[113,178,1836,1860]" captionTargetBox="[255,1316,146,1806]" captionTargetId="figure-6@11.[253,1319,144,1808]" captionTargetPageId="11" captionText="Figure 9. Habitus of Zimmiodes, lateral view. A, Zimmiodes australis, male, holotype. B, Z. australis, female, paratype. C, Zimmiodes papuanus, male, holotype. D, Z. papuanus, female, paratype. Scale bars: 1.00 mm." pageId="17" pageNumber="18">Fig. 9C, D</figureCitation>
).
</paragraph>
<paragraph id="1E486951FF81FF82FCB3FD52AD732705" blockId="17.[810,1459,734,789]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FCB3FD52AA9D26E6" box="[810,1018,734,758]" italics="true" pageId="17" pageNumber="18">Colour and vestiture:</emphasis>
Body and legs reddish to blackish brown, semilustrous, glabrous.
</paragraph>
<paragraph id="1E486951FF81FF82FCB3FCBBAA8B2079" blockId="17.[810,1460,823,1129]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FCB3FCBBAA00275F" box="[810,871,823,847]" italics="true" pageId="17" pageNumber="18">Head:</emphasis>
Rostrum: moderately long, as long as pronotum, robust, slightly thinner in female, moderately ventrally curved, dorsoventrally flattened, slightly broadened apically in dorsal view, coarsely punctate dorsally, punctures finer in distal half. Eyes: subcircular in outline, strongly convex and protruding. Antennae: inserted close to base of rostrum in male and female (
<figureCitation id="86CC75D4FF81FF82FCACFC7FAAC7201A" box="[821,928,1011,1035]" captionStart="Figure 8" captionStartId="10.[129,194,1836,1860]" captionTargetBox="[239,1364,146,1798]" captionTargetId="figure-8@10.[237,1366,144,1808]" captionTargetPageId="10" captionText="Figure 8. Habitus of Zimmiodes, dorsal view. A, Zimmiodes australis, male, holotype.B, Z. australis, female, paratype. C, Zimmiodes papuanus, male, holotype.D, Z. papuanus, female, paratype. Scale bars: 1.00 mm." pageId="17" pageNumber="18">Figs 8C, D</figureCitation>
,
<figureCitation id="86CC75D4FF81FF82FC35FC7FAA8E201A" box="[940,1001,1011,1035]" captionStart="Figure 9" captionStartId="11.[113,178,1836,1860]" captionTargetBox="[255,1316,146,1806]" captionTargetId="figure-6@11.[253,1319,144,1808]" captionTargetPageId="11" captionText="Figure 9. Habitus of Zimmiodes, lateral view. A, Zimmiodes australis, male, holotype. B, Z. australis, female, paratype. C, Zimmiodes papuanus, male, holotype. D, Z. papuanus, female, paratype. Scale bars: 1.00 mm." pageId="17" pageNumber="18">9C, D</figureCitation>
); scapes reaching eye; funicles with segment 1 ~1.0×, 1.6×, 1.6×, and 1.6× as long as segments 25, respectively; clubs stout, ~1.8× as long as wide, densely and finely pubescent (
<figureCitation id="86CC75D4FF81FF82FC11FBDDAABC2079" box="[904,987,1105,1129]" captionStart="Figure 10" captionStartId="12.[129,194,1836,1860]" captionTargetBox="[308,1284,146,1806]" captionTargetId="figure-8@12.[306,1296,144,1808]" captionTargetPageId="12" captionText="Figure 10. Diagnostic differences of antennae and forelegs of Zimmiodes australis and Zimmiodes papuanus. A, Z. australis, right antenna. B, Z. papuanus, right antenna. C, Z. australis, foreleg of male.D, Z. papuanus, foreleg of male.Scale bars: 0.20 mm." pageId="17" pageNumber="18">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="1E486951FF81FF82FCB3FB06AC4921EB" blockId="17.[810,1460,1162,1531]" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FCB3FB06AA1F20B2" box="[810,888,1162,1186]" italics="true" pageId="17" pageNumber="18">Thorax:</emphasis>
Pronotum: roundly trapezoidal, apex ~0.6× narrower than base; anterior and posterior margins subtruncate, lateral margins mostly rounded, distinctly narrowed anteriorly; disc weakly and evenly convex; surface densely and distinctly punctate. Scutellar shield: roundly pentagonal. Elytra: ~2.22.4× as long as pronotum, jointly ~0.5× as broad as long, slightly broader than base of pronotum; humeri broadly rounded, slightly protruding; surface slightly evenly convex, deeply and coarsely punctate in rows, forming distinct striae. Legs: femora ventrally without subapical tooth, profemora of male distinctly obtusely protruding inwards (
<figureCitation id="86CC75D4FF81FF82FBFAFA4FADDA21CB" box="[1123,1213,1475,1499]" captionStart="Figure 10" captionStartId="12.[129,194,1836,1860]" captionTargetBox="[308,1284,146,1806]" captionTargetId="figure-8@12.[306,1296,144,1808]" captionTargetPageId="12" captionText="Figure 10. Diagnostic differences of antennae and forelegs of Zimmiodes australis and Zimmiodes papuanus. A, Z. australis, right antenna. B, Z. papuanus, right antenna. C, Z. australis, foreleg of male.D, Z. papuanus, foreleg of male.Scale bars: 0.20 mm." pageId="17" pageNumber="18">Fig. 10D</figureCitation>
); tibiae with premucro smaller than uncus; tarsi with claws free, divergent.
</paragraph>
<paragraph id="1E486951FF81FF81FCB3F9AEAB9A2593" blockId="17.[810,1460,1570,1970]" lastBlockId="18.[129,778,144,387]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="2C83B543FF81FF82FCB3F9AEAAFA222A" box="[810,925,1570,1594]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="D9F712D2FF81FF82FCB3F9AEAAFF222A" box="[810,920,1570,1594]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
:
</emphasis>
Sternite VIII of male subtrapezoidal, plates sclerotized, slanting, apical margin rounded, sparsely covered with short setae subapically, basal margin strongly sclerotized (
<figureCitation id="86CC75D4FF81FF82FA14F9EDAA352288" captionStart="Figure 11" captionStartId="13.[115,178,1449,1473]" captionTargetBox="[114,1457,145,1417]" captionTargetId="figure-341@13.[112,1459,144,1422]" captionTargetPageId="13" captionText="Figure 11.Terminalia ofZimmiodes. A, Zimmiodes australis, sternite VIII,male.B,Zimmiodes papuanus, sternite VIII,male.C,Z.australis, sternite VIII,female.D, Z.papuanus, sternite VIII, female.E,Z. australis, spiculumgastrale.F,Z. papuanus, spiculum gastrale.Scale bars:0.20 mm." pageId="17" pageNumber="18">Fig. 11B</figureCitation>
), of female abruptly angled laterally, with apical margin densely covered with short setae (
<figureCitation id="86CC75D4FF81FF82FB14F913AD8022A7" box="[1165,1255,1695,1719]" captionStart="Figure 11" captionStartId="13.[115,178,1449,1473]" captionTargetBox="[114,1457,145,1417]" captionTargetId="figure-341@13.[112,1459,144,1422]" captionTargetPageId="13" captionText="Figure 11.Terminalia ofZimmiodes. A, Zimmiodes australis, sternite VIII,male.B,Zimmiodes papuanus, sternite VIII,male.C,Z.australis, sternite VIII,female.D, Z.papuanus, sternite VIII, female.E,Z. australis, spiculumgastrale.F,Z. papuanus, spiculum gastrale.Scale bars:0.20 mm." pageId="17" pageNumber="18">Fig. 11D</figureCitation>
); spiculum gastrale strongly curved apically, base lightly sclerotized (
<figureCitation id="86CC75D4FF81FF82FADFF933ACC722C7" box="[1350,1440,1727,1751]" captionStart="Figure 11" captionStartId="13.[115,178,1449,1473]" captionTargetBox="[114,1457,145,1417]" captionTargetId="figure-341@13.[112,1459,144,1422]" captionTargetPageId="13" captionText="Figure 11.Terminalia ofZimmiodes. A, Zimmiodes australis, sternite VIII,male.B,Zimmiodes papuanus, sternite VIII,male.C,Z.australis, sternite VIII,female.D, Z.papuanus, sternite VIII, female.E,Z. australis, spiculumgastrale.F,Z. papuanus, spiculum gastrale.Scale bars:0.20 mm." pageId="17" pageNumber="18">Fig. 11F</figureCitation>
); tegmen with complete ring, without parameroid lobes, manubrium very short; penis elongate (~2.83.1× as long as wide), subparallel-sided, slightly narrowing in apical half, moderately narrowed apically, dorsum strongly sclerotized, forming a median groove roundly open in apical quarter, apical margin bluntly rounded, apical opening with a pair of lanceolate orificial sclerites, temones slender, ~1.2× as long as body of penis, anteriorly confluent, joined ventrally to body of penis (
<figureCitation id="86CC75D4FF82FF81FD3DFF1CA9E824D7" captionStart="Figure 12" captionStartId="14.[129,194,1119,1143]" captionTargetBox="[130,1473,145,1090]" captionTargetId="figure-498@14.[128,1475,144,1092]" captionTargetPageId="14" captionText="Figure 12. Male genitalia (aedeagus) of Zimmiodes. A, Zimmiodes australis, dorsal view. B, Z. australis, ventral view.C, Z. australis, lateral view. D, Zimmiodes papuanus, dorsal view.E, Z. papuanus, ventral view.F, Z. papuanus, lateral view. Scale bar: 0.20 mm." pageId="18" pageNumber="19">Fig. 12D F</figureCitation>
); endophallus long, extending below body of penis, membranous, medially with a sleeve of dense, strongly developed denticles, apically with a pair of slender sclerites, bent basally; ovipositor with gonocoxites slender, elongate, apically bluntly rounded, styli short, subcylindrical, bluntly rounded and setose apically (
<figureCitation id="86CC75D4FF82FF81FF7AFEC0A8582574" box="[227,319,332,356]" captionStart="Figure 13" captionStartId="15.[113,178,1836,1860]" captionTargetBox="[183,1384,145,1806]" captionTargetId="figure-7@15.[182,1390,144,1808]" captionTargetPageId="15" captionText="Figure 13. Female genitalia of Zimmiodes. A, Zimmiodes australis, ovipositor.B, Z. australis, spermatheca with gland. C, Zimmiodes papuanus, ovipositor.D, Z. papuanus, spermatheca with gland.Scale bars: 0.20 mm." pageId="18" pageNumber="19">Fig. 13C</figureCitation>
); spermatheca evenly narrow, right-angled, gland elongate, narrowing apicad, rounded apically (
<figureCitation id="86CC75D4FF82FF81FD0CFEE7AB8B2593" box="[661,748,363,387]" captionStart="Figure 13" captionStartId="15.[113,178,1836,1860]" captionTargetBox="[183,1384,145,1806]" captionTargetId="figure-7@15.[182,1390,144,1808]" captionTargetPageId="15" captionText="Figure 13. Female genitalia of Zimmiodes. A, Zimmiodes australis, ovipositor.B, Z. australis, spermatheca with gland. C, Zimmiodes papuanus, ovipositor.D, Z. papuanus, spermatheca with gland.Scale bars: 0.20 mm." pageId="18" pageNumber="19">Fig. 13D</figureCitation>
).
</paragraph>
<paragraph id="1E486951FF82FF81FEF2FE28AB7825AD" blockId="18.[128,777,419,513]" box="[363,543,419,445]" pageId="18" pageNumber="19">
<emphasis id="2C83B543FF82FF81FEF2FE28AB7825AD" box="[363,543,419,445]" italics="true" pageId="18" pageNumber="19">Derivation of name</emphasis>
</paragraph>
<paragraph id="1E486951FF82FF81FF18FE45ABA92611" blockId="18.[128,777,419,513]" pageId="18" pageNumber="19">
The specific epithet
<taxonomicName id="D9F712D2FF82FF81FED0FE46A8CF25F1" authorityName="Hsiao &amp; Oberprieler" authorityYear="2024" box="[329,424,458,481]" class="Insecta" family="Curculionidae" genus="Zimmiodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="2C83B543FF82FF81FED0FE46A8CF25F1" box="[329,424,458,481]" italics="true" pageId="18" pageNumber="19">papuanus</emphasis>
</taxonomicName>
refers to the Papuan region of New
<collectingCountry id="66E029C1FF82FF81FF19FE65A9AA2611" box="[128,205,489,513]" name="Guinea-Bissau" pageId="18" pageNumber="19">Guinea</collectingCountry>
, the only known area of occurrence of this species.
</paragraph>
<paragraph id="1E486951FF82FF81FE12FDACA899262A" blockId="18.[395,510,544,570]" box="[395,510,544,570]" pageId="18" pageNumber="19">
<emphasis id="2C83B543FF82FF81FE12FDACA899262A" box="[395,510,544,570]" italics="true" pageId="18" pageNumber="19">Distribution</emphasis>
</paragraph>
<paragraph id="1E486951FF82FF81FF18FDCBA801266E" blockId="18.[129,778,583,638]" pageId="18" pageNumber="19">
This species is known only from the eastern regions of
<collectingCountry id="66E029C1FF82FF81FD52FDCBA866266E" name="Papua New Guinea" pageId="18" pageNumber="19">Papua New Guinea</collectingCountry>
(
<figureCitation id="86CC75D4FF82FF81FE89FDEAA832266E" box="[272,341,614,638]" captionStart="Figure 14" captionStartId="16.[129,194,1236,1260]" captionTargetBox="[129,1473,144,1208]" captionTargetId="figure-393@16.[129,1473,144,1208]" captionTargetPageId="16" captionText="Figure 14. Geographical distribution of Zimmiodes in Australia and Papua New Guinea." pageId="18" pageNumber="19">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="1E486951FF82FF81FEE4FD11AB6A26A7" blockId="18.[381,525,669,695]" box="[381,525,669,695]" pageId="18" pageNumber="19">
<emphasis id="2C83B543FF82FF81FEE4FD11AB6A26A7" box="[381,525,669,695]" italics="true" pageId="18" pageNumber="19">Natural history</emphasis>
</paragraph>
<paragraph id="1E486951FF82FF81FF18FD48A8F427E6" blockId="18.[127,780,708,1014]" pageId="18" pageNumber="19">
Based on the label data, the species was recorded to be associated with
<taxonomicName id="D9F712D2FF82FF81FF2AFD68A82026EB" box="[179,327,740,763]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="papuana">
<emphasis id="2C83B543FF82FF81FF2AFD68A82026EB" box="[179,327,740,763]" italics="true" pageId="18" pageNumber="19">Cycas papuana</emphasis>
</taxonomicName>
. However, the
<taxonomicName id="D9F712D2FF82FF81FE47FD68AB0326EB" box="[478,612,739,763]" pageId="18" pageNumber="19">
<emphasis id="2C83B543FF82FF81FE47FD68AB7026EB" box="[478,535,740,763]" italics="true" pageId="18" pageNumber="19">Cycas</emphasis>
species
</taxonomicName>
occurring at the
<typeStatus id="C14CD7F3FF82FF81FF19FC8FA9C9270B" box="[128,174,771,795]" pageId="18" pageNumber="19">type</typeStatus>
locality (i.e.
<collectingRegion id="DC33A7B3FF82FF81FEA7FC8FA8F1270B" box="[318,406,771,795]" country="Papua New Guinea" name="Madang" pageId="18" pageNumber="19">Madang</collectingRegion>
) is apparently
<taxonomicName id="D9F712D2FF82FF81FDA0FC8FAA6E270B" box="[569,777,771,795]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="schumanniana">
<emphasis id="2C83B543FF82FF81FDA0FC8FAA6E270B" box="[569,777,771,795]" italics="true" pageId="18" pageNumber="19">Cycas schumanniana</emphasis>
</taxonomicName>
rather than
<taxonomicName id="D9F712D2FF82FF81FF65FCAEA8F4272A" box="[252,403,802,826]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="papuana">
<emphasis id="2C83B543FF82FF81FF65FCAEA8F4272A" box="[252,403,802,826]" italics="true" pageId="18" pageNumber="19">Cycas papuana</emphasis>
</taxonomicName>
, which is distributed in western regions of
<collectingCountry id="66E029C1FF82FF81FF47FCCEA8CC2749" box="[222,427,833,857]" name="Papua New Guinea" pageId="18" pageNumber="19">Papua New Guinea</collectingCountry>
, Indonesian Papua and Prince of
<collectingCountry id="66E029C1FF82FF81FF18FCEDA8612769" box="[129,262,865,889]" name="Wales Island" pageId="18" pageNumber="19">Wales Island</collectingCountry>
of
<collectingRegion id="DC33A7B3FF82FF81FEA8FCEDA8C82769" box="[305,431,865,889]" country="Australia" name="Queensland" pageId="18" pageNumber="19">Queensland</collectingRegion>
,
<collectingCountry id="66E029C1FF82FF81FE24FCEDAB7F2769" box="[445,536,865,889]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
(
<bibRefCitation id="7A6614A0FF82FF81FDB7FCEDAB9B2769" author="Calonje M &amp; Stevenson DW &amp; Osborne R" box="[558,764,865,889]" pageId="18" pageNumber="19" refId="ref11232" refString="Calonje M, Stevenson DW, Osborne R. The World List of Cycads, online edn. 2013 - 2023. http: // www. cycadlist. org (25 July 2023, date last accessed)." type="url" year="2013">
Calonje
<emphasis id="2C83B543FF82FF81FD10FCEDABD92769" box="[649,702,865,889]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2013
</bibRefCitation>
2023). A species of
<taxonomicName id="D9F712D2FF82FF81FED6FC0CA8A42788" box="[335,451,896,920]" class="Insecta" family="Curculionidae" genus="Tychiosoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="2C83B543FF82FF81FED6FC0CA8A42788" box="[335,451,896,920]" italics="true" pageId="18" pageNumber="19">Tychiosoma</emphasis>
</taxonomicName>
(of the
<taxonomicName id="D9F712D2FF82FF81FD8FFC0CAB122788" box="[534,629,896,920]" class="Insecta" family="Curculionidae" genus="Tychiodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="2C83B543FF82FF81FD8FFC0CAB122788" box="[534,629,896,920]" italics="true" pageId="18" pageNumber="19">Tychiodes</emphasis>
</taxonomicName>
group of weevils; Supporting Information, Appendix S4C) co-exists with
<taxonomicName id="D9F712D2FF82FF81FD6FFC2CA98727C6" authorityName="Hsiao &amp; Oberprieler" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Zimmiodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="2C83B543FF82FF81FD6FFC2CA98727C6" italics="true" pageId="18" pageNumber="19">Z. papuanus</emphasis>
</taxonomicName>
at these localities (several specimens examined, from the same collecting event).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,140 +0,0 @@
<document id="A315BE78977F47C92904B19E99AD9BB8" ID-DOI="10.1093/zoolinnean/zlad188" ID-ISSN="0024-4082" ID-Zenodo-Dep="14422812" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734028749482" checkinUser="plazi" docAuthor="Santos, Rodolfo Otávio, Wilkinson, Mark, Graziella, Ribeiro, Couto, Carvalho, Alberto B. &amp; Zaher, Hussam" docDate="2024" docId="DF1E2454FF8A9E14FB92FD9133D15611" docLanguage="en" docName="zlad188.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (2)" docSource="https://doi.org/10.1093/zoolinnean/zlad188" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Ymboirana Santos, Wilkinson, Graziella, Ribeiro, Carvalho &amp; Zaher, 2024, gen. nov." docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="23275C2CFF889E16FF84FFA8304D503F" masterDocTitle="Ŋe first fossil record of an aquatic caecilian (Gymnophiona: Typhlonectidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="3" updateTime="1734029593310" updateUser="ExternalLinkService">
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<mods:title id="7426E789B4F5F63ADF14B05C9D26F4FF">Ŋe first fossil record of an aquatic caecilian (Gymnophiona: Typhlonectidae)</mods:title>
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<heading id="0C40222EFF8A9E14FB92FD9134A8526B" bold="true" box="[1046,1253,569,596]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="65C34950FF8A9E14FB92FD9134A8526B" bold="true" box="[1046,1253,569,596]" pageId="2" pageNumber="3">
<taxonomicName id="90B7EEC1FF8A9E14FB92FD9134C8526B" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[1046,1157,569,596]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="65C34950FF8A9E14FB92FD9134C8526B" bold="true" box="[1046,1157,569,596]" italics="true" pageId="2" pageNumber="3">Ymboirana</emphasis>
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<taxonomicNameLabel id="7EF0F42BFF8A9E14FB0EFD9234A8526B" box="[1162,1253,570,596]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<emphasis id="65C34950FF8A9E14FCBDFDD2346B52AD" box="[825,1062,634,658]" italics="true" pageId="2" pageNumber="3">ZooBank registration:</emphasis>
urn:lsid:zoobank.org:act:1B16342C* 6788*4962*BF69*E970F3F93A56
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<emphasis id="65C34950FF8A9E14FCBDFD7C33D252D4" box="[825,927,724,747]" italics="true" pageId="2" pageNumber="3">Diagnosis:</emphasis>
Ŋe new taxon is recognized as a caecilian because it has a pseudangular with a distinctive retroarticular process, an elongate vertebral column comprised of amphicoelous vertebrae bearing well*developed anterior basapophyseal processes, and lacks limbs and girdles.
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<emphasis id="65C34950FF8A9E14FBB2FCF834EC5357" box="[1078,1185,848,872]" italics="true" pageId="2" pageNumber="3">Ymboirana</emphasis>
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differs from rhinatrematids in lacking sagiưal crests on the parietals; from caeciliids in having all teeth roughly similar in size; from siphonopids and scolecomorphids in having an inner mandibular tooth row; from the herpelid
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<emphasis id="65C34950FF8A9E14FC72FC66340C53D9" box="[1014,1089,974,998]" italics="true" pageId="2" pageNumber="3">Herpele</emphasis>
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by having a smooth dorsal surface of the parietal (i.e. without ornamentations in the form of foramina for neurovascular canals) and from chikilids, dermophiids, grandisoniids, ichthyophiids, and the herpelid
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<emphasis id="65C34950FF8A9E14FAC7FB83358F547C" box="[1347,1474,1067,1091]" italics="true" pageId="2" pageNumber="3">Boulengerula</emphasis>
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by having only monocuspid teeth.
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<emphasis id="65C34950FF8A9E14FB36FBE33550545C" box="[1202,1309,1099,1123]" italics="true" pageId="2" pageNumber="3">Ymboirana</emphasis>
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shares with all typhlonectids, except
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<emphasis id="65C34950FF8A9E14FBA5FBC234FF54BD" box="[1057,1202,1130,1154]" italics="true" pageId="2" pageNumber="3">Chthonerpeton</emphasis>
</taxonomicName>
, the presence of elongate trunk vertebrae and well*developed ventral ridges on the basapophyseal processes. Unique characteristics include the following: pseudangular dorsally recurved and posteriorly pro* jected, bearing a poorly developed processus condyloides and a fossa leading to the canalis primordialis located far from (i.e. not immediately adjacent to) the articular surface.
</paragraph>
<paragraph id="57089542FF8A9E14FCBDFAC8358E5547" blockId="2.[825,1475,1376,1400]" box="[825,1475,1376,1400]" pageId="2" pageNumber="3">
<emphasis id="65C34950FF8A9E14FCBDFAC834755548" box="[825,1080,1376,1400]" italics="true" pageId="2" pageNumber="3">
Content:
<taxonomicName id="90B7EEC1FF8A9E14FC16FAC834755548" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[914,1080,1376,1400]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="acrux" status="sp. nov.">Ymboirana acrux</taxonomicName>
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,
<taxonomicNameLabel id="7EF0F42BFF8A9E14FBC4FAC834C65548" box="[1088,1163,1376,1399]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
, by monotypy and designation.
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<emphasis id="65C34950FF8A9E14FCBDFA3133E0558E" box="[825,941,1433,1457]" italics="true" pageId="2" pageNumber="3">Etomologo:</emphasis>
From the Tupi
<emphasis id="65C34950FF8A9E14FBDDFA323425558E" box="[1113,1128,1434,1457]" italics="true" pageId="2" pageNumber="3">Y</emphasis>
(water),
<emphasis id="65C34950FF8A9E14FB49FA3134B2558E" box="[1229,1279,1433,1457]" italics="true" pageId="2" pageNumber="3">mboî</emphasis>
(snake), and
<emphasis id="65C34950FF8A9E14FA10FA32358F558E" box="[1428,1474,1434,1457]" italics="true" pageId="2" pageNumber="3">rana</emphasis>
(similar to). Caecilians are ossen misidentified as snakes due to their overall similarity, and typhlonectids are distin* guished from other caecilians by their aquatic or semi*aquatic lifestyles.
</paragraph>
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<mods:identifier id="73A955219168B665CFAF0337E1E93B41" type="Zenodo-Dep">14422812</mods:identifier>
</mods:mods>
<treatment id="DF1E2454FF8A9E1DFC7BF9E5350854FE" LSID="urn:lsid:plazi:treatment:DF1E2454FF8A9E1DFC7BF9E5350854FE" httpUri="http://treatment.plazi.org/id/DF1E2454FF8A9E1DFC7BF9E5350854FE" lastPageId="11" lastPageNumber="12" pageId="2" pageNumber="3">
<subSubSection id="1FADC6C9FF8A9E14FC7BF9E534E656FA" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="57089542FF8A9E14FC7BF9E534B15657" blockId="2.[1023,1276,1613,1640]" box="[1023,1276,1613,1640]" pageId="2" pageNumber="3">
<heading id="0C40222EFF8A9E14FC7BF9E534B15657" bold="true" box="[1023,1276,1613,1640]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="65C34950FF8A9E14FC7BF9E534B15657" bold="true" box="[1023,1276,1613,1640]" pageId="2" pageNumber="3">
<taxonomicName id="90B7EEC1FF8A9E14FC7BF9E534E65657" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[1023,1195,1613,1640]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="acrux" status="sp. nov.">
<emphasis id="65C34950FF8A9E14FC7BF9E534E65657" bold="true" box="[1023,1195,1613,1640]" italics="true" pageId="2" pageNumber="3">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF8A9E14FB34F9E634B15657" box="[1200,1276,1614,1640]" pageId="2" pageNumber="3" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="57089542FF8A9E14FCBDF92634E656FA" blockId="2.[825,1474,1678,1733]" pageId="2" pageNumber="3">
<emphasis id="65C34950FF8A9E14FCBDF926346A5699" box="[825,1063,1678,1702]" italics="true" pageId="2" pageNumber="3">ZooBank registration:</emphasis>
urn:lsid:zoobank.org:act:D6487148* 6E74*436A*83C9*C89AC455B4BB
</paragraph>
</subSubSection>
<subSubSection id="1FADC6C9FF8A9E12FCBDF94035635791" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="57089542FF8A9E14FCBDF940346456C0" blockId="2.[825,1065,1767,1791]" box="[825,1065,1767,1791]" pageId="2" pageNumber="3">
<emphasis id="65C34950FF8A9E14FCBDF94033D256C0" box="[825,927,1768,1791]" italics="true" pageId="2" pageNumber="3">Diagnosis:</emphasis>
As for genus.
</paragraph>
<paragraph id="57089542FF8A9E12FCBDF8883151576A" blockId="2.[825,1474,1824,1974]" lastBlockId="4.[129,777,1728,1877]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<emphasis id="65C34950FF8A9E14FCBDF88833D35707" box="[825,926,1824,1848]" italics="true" pageId="2" pageNumber="3">Holotope:</emphasis>
DGM 1462*R (
<figureCitation id="CF8C89C7FF8A9E14FBDCF88834FA5707" box="[1112,1207,1824,1848]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStart-6="Figure 8" captionStart-7="Figure 9" captionStartId-0="3.[114,179,1836,1860]" captionStartId-1="4.[129,194,1627,1651]" captionStartId-2="5.[115,180,1628,1652]" captionStartId-3="6.[130,195,1627,1651]" captionStartId-4="7.[115,180,1627,1651]" captionStartId-5="8.[129,194,1626,1650]" captionStartId-6="9.[115,180,1627,1651]" captionStartId-7="10.[129,194,1629,1653]" captionTargetBox-0="[238,1336,144,1808]" captionTargetBox-1="[129,1473,144,1600]" captionTargetBox-2="[114,1458,144,1600]" captionTargetBox-3="[129,1473,144,1600]" captionTargetBox-4="[114,1458,144,1600]" captionTargetBox-5="[129,1473,142,1598]" captionTargetBox-6="[114,1458,144,1600]" captionTargetBox-7="[129,1473,145,1601]" captionTargetId-0="figure-8@3.[238,1336,144,1808]" captionTargetId-1="figure-140@4.[129,1473,144,1600]" captionTargetId-2="figure-107@5.[114,1458,144,1600]" captionTargetId-3="figure-131@6.[129,1473,144,1600]" captionTargetId-4="figure-181@7.[114,1458,144,1600]" captionTargetId-5="figure-120@8.[129,1473,142,1598]" captionTargetId-6="figure-148@9.[114,1458,144,1600]" captionTargetId-7="figure-150@10.[129,1473,145,1601]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="Figure 2. Fossil slabs containing the holotype (DGM 1462*R) of Ymboirana acrux gen. et sp. nov. (A1, B1). CT*scans of the part (A2) and counterpart (B2) of the holotype.Scale bar = 1 cm." captionText-1="Figure 3. CT*scans of the two skull parts of Ymboirana acrux gen. et sp. nov. Top part in dorsal (A) and ventral (B) views. Boưom part in dorsal (C) and ventral (D) views. See Appendix for abbreviations.Scale bar = 1 mm." captionText-2="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." captionText-3="Figure 5. Lower jaw of Ymboirana acrux gen. et sp. nov. Mandibles in dorsal (A) and ventral (B) views.Less pseudangular in lateral (C), posterior oblique (D), and dorsal (E) views. Doưed lines indicate the region of the contact between the pseudangular and pseudodentary.See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for A, B, and the lower one is for CE." captionText-4="Figure 6. Isolated tooth crowns found near the skull of Ymboirana acruxgen.et sp. nov.. Scale bar = 1 mm." captionText-5="Figure 7. Vertebral column of Ymboirana acrux gen. et sp. nov. (A). Articulated trunk vertebrae from the posterior portion of the body in dorsal (B) and lateral (C) views. See Appendix for abbreviations. Upper scale bar = 1 cm. Lower scale bar = 1 mm; upper scale bar is for A, and the lower one is for B, C." captionText-6="Figure 8. Postcranial elements of Ymboirana acrux gen. et sp. nov.. Atlas in dorsal (A) and ventral (B) views. Ventral part of atlas (i.e. atlantal centrum and cotyles) in anterior (C), posterior (D), and oblique less lateral (E) views. Second vertebra in dorsal (F) and ventral (G) views.See Appendix for abbreviations.Scale bar = 1 mm." captionText-7="Figure 9. Postcranial elements of Ymboirana acrux gen. et sp.nov.. Trunk vertebra in dorsal (A), ventral (B), and oblique less lateral (C) views. Ribs (D, E) in lateral view. See Appendix for abbreviations. Scale bar = 1 mm." pageId="2" pageNumber="3">Figs 29</figureCitation>
,
<figureCitation id="CF8C89C7FF8A9E14FB4CF88834B95707" box="[1224,1268,1824,1848]" captionStart="Figure 12" captionStartId="15.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-115@15.[114,1458,144,1600]" captionTargetPageId="15" captionText="Figure 12. Paired parietals of four different caecilian species, all in dorsal view.Ymboirana acrux gen.et sp. nov.(A), Rhinatrema bivitatum (B), Tophlonectes natans (C), and Herpele squalostoma (D). See Appendix for abbreviations. Scale bar = 1 mm." pageId="2" pageNumber="3">12A</figureCitation>
,
<figureCitation id="CF8C89C7FF8A9E14FA82F888357F5707" box="[1286,1330,1824,1848]" captionStart="Figure 13" captionStartId="16.[130,195,1627,1651]" captionTargetBox="[129,1473,144,1600]" captionTargetId="figure-149@16.[129,1473,144,1600]" captionTargetPageId="16" captionText="Figure 13. Less stapes of six different caecilian species, all in lateral view.Fragment tentatively identified as the stapes of Ymboirana acrux gen. et sp.nov.(A), Potomotophlus kaupii (B), Tophlonectes natans (C), Caecilia sp. (D), Siphonops paulensis (E), and Rhinatrema bivitatum (F). See Appendix for abbreviations.Scale bar = 1 mm." pageId="2" pageNumber="3">13A</figureCitation>
,
<figureCitation id="CF8C89C7FF8A9E14FAC0F888353C5707" box="[1348,1393,1824,1848]" captionStart="Figure 14" captionStartId="17.[112,177,932,956]" captionTargetBox="[114,1458,143,904]" captionTargetId="figure-536@17.[114,1458,143,904]" captionTargetPageId="17" captionText="Figure 14. Less pseudangulars of four different caecilian species. Ymboirana acrux gen. et sp. nov. in lateral (A1) and dorsal (A2) views. Rhinatrema bivitatum in lateral (B1) and dorsal (B2) views. Scolecomorphus ulugurensis in lateral (C1) and dorsal (C2) views.Tophlonectes natans in lateral (D1) and dorsal (D2) views. See Appendix for abbreviations. Scale bar = 1 mm." pageId="2" pageNumber="3">14A</figureCitation>
,
<figureCitation id="CF8C89C7FF8A9E14FA06F88835FE5707" box="[1410,1459,1824,1848]" captionStart="Figure 15" captionStartId="18.[129,194,1899,1923]" captionTargetBox="[129,1473,1111,1872]" captionTargetId="figure-568@18.[129,1473,1111,1872]" captionTargetPageId="18" captionText="Figure 15. Atlases of four different typhlonectid caecilians. Ymboirana acrux gen. et sp. nov. in anterior (A1) and dorsal (A2) views. Chthonerpeton indistinctum in anterior (B1) and dorsal (B2) views. Potomotophlus kaupii in anterior (C1) and dorsal (C2) views. Nectocaecilia petersii in anterior (D1) and dorsal (D2) views. See Appendix for abbreviations.Scale bar = 1 mm." pageId="2" pageNumber="3">15A</figureCitation>
<figureCitation id="CF8C89C7FF8A9E14FA30F88835F05707" box="[1460,1469,1824,1848]" captionStart="Figure 1" captionStartId="1.[113,178,1960,1984]" captionTargetBox="[116,1456,1173,1930]" captionTargetId="figure-558@1.[114,1458,1171,1932]" captionTargetPageId="1" captionText="Figure 1. Map of the Taubaté Basin in Southeastern Brazil, showing the type locality for Ymboirana acrux gen. et sp.nov.." pageId="2" pageNumber="3">1</figureCitation>
, A
<figureCitation id="CF8C89C7FF8A9E14FCC8F8E833155767" box="[844,856,1856,1880]" captionStart="Figure 2" captionStartId="3.[114,179,1836,1860]" captionTargetBox="[238,1336,144,1808]" captionTargetId="figure-8@3.[238,1336,144,1808]" captionTargetPageId="3" captionText="Figure 2. Fossil slabs containing the holotype (DGM 1462*R) of Ymboirana acrux gen. et sp. nov. (A1, B1). CT*scans of the part (A2) and counterpart (B2) of the holotype.Scale bar = 1 cm." pageId="2" pageNumber="3">2</figureCitation>
; Supporting Information,
<figureCitation id="CF8C89C7FF8A9E14FBFCF8E834855767" box="[1144,1224,1856,1880]" captionStart="Figure 1" captionStartId="1.[113,178,1960,1984]" captionTargetBox="[116,1456,1173,1930]" captionTargetId="figure-558@1.[114,1458,1171,1932]" captionTargetPageId="1" captionText="Figure 1. Map of the Taubaté Basin in Southeastern Brazil, showing the type locality for Ymboirana acrux gen. et sp.nov.." pageId="2" pageNumber="3">Figs S1</figureCitation>
, S
<figureCitation id="CF8C89C7FF8A9E14FB66F8E834A25767" box="[1250,1263,1856,1880]" captionStart="Figure 2" captionStartId="3.[114,179,1836,1860]" captionTargetBox="[238,1336,144,1808]" captionTargetId="figure-8@3.[238,1336,144,1808]" captionTargetPageId="3" captionText="Figure 2. Fossil slabs containing the holotype (DGM 1462*R) of Ymboirana acrux gen. et sp. nov. (A1, B1). CT*scans of the part (A2) and counterpart (B2) of the holotype.Scale bar = 1 cm." pageId="2" pageNumber="3">2</figureCitation>
), housed at Museu de Ciências da Terra (formerly Divisão de Geologia e Mineralogia of the Departamento Nacional de Produção Mineral),
<collectingRegion id="95735BA0FF8A9E14FC2DF83634065789" box="[937,1099,1950,1974]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="3">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="2FA0D5D2FF8A9E14FBDDF83634D75789" box="[1113,1178,1950,1974]" name="Brazil" pageId="2" pageNumber="3">Brazil</collectingCountry>
. The specimen consists of part and counterpart preserving the anterior portion of a cae* cilian skeleton (with an estimated length of around
<quantity id="904F38A7FF8C9E12FD32F97732B056C8" box="[694,765,1759,1783]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="cm" value="20.0">20 cm</quantity>
), including skull, mandibles, and the anteriormost portion of the vertebral column, including the atlas, 66 trunk vertebrae, and some ribs.
</paragraph>
<caption id="03C8C5CAFF8B9E15FFF6F884321A575F" pageId="3" pageNumber="4" startId="3.[114,179,1836,1860]" targetBox="[238,1336,144,1808]" targetPageId="3" targetType="figure">
<paragraph id="57089542FF8B9E15FFF6F884321A575F" blockId="3.[114,1409,1836,1888]" pageId="3" pageNumber="4">
<emphasis id="65C34950FF8B9E15FFF6F8843087577B" bold="true" box="[114,202,1836,1860]" pageId="3" pageNumber="4">Figure 2.</emphasis>
Fossil slabs containing the holotype (DGM 1462*R) of
<taxonomicName id="90B7EEC1FF8B9E15FD5EF884333B577B" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[730,886,1836,1860]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov.">
<emphasis id="65C34950FF8B9E15FD5EF884333B577B" box="[730,886,1836,1860]" italics="true" pageId="3" pageNumber="4">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF8B9E15FCFEF885344E577B" box="[890,1027,1837,1861]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(A1, B1). CT*scans of the part (A2) and counterpart (B2) of the holotype. Scale bar = 1 cm.
</paragraph>
</caption>
<caption id="03C8C5CAFF8C9E12FF05F9F3331056B0" pageId="4" pageNumber="5" startId="4.[129,194,1627,1651]" targetBox="[129,1473,144,1600]" targetPageId="4" targetType="figure">
<paragraph id="57089542FF8C9E12FF05F9F3331056B0" blockId="4.[129,1468,1627,1680]" pageId="4" pageNumber="5">
<emphasis id="65C34950FF8C9E12FF05F9F33095564B" bold="true" box="[129,216,1627,1652]" pageId="4" pageNumber="5">Figure 3.</emphasis>
CT*scans of the two skull parts of
<taxonomicName id="90B7EEC1FF8C9E12FD98F9F432F5564C" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[540,696,1628,1652]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov.">
<emphasis id="65C34950FF8C9E12FD98F9F432F5564C" box="[540,696,1628,1652]" italics="true" pageId="4" pageNumber="5">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF8C9E12FD38F9F43308564C" box="[700,837,1628,1652]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Top part in dorsal (A) and ventral (B) views. Boưom part in dorsal (C) and ventral (D) views. See Appendix for abbreviations. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="57089542FF8C9E12FF05F8DF349A5728" blockId="4.[129,778,1910,1965]" lastBlockId="4.[825,1475,1727,1815]" pageId="4" pageNumber="5">
<emphasis id="65C34950FF8C9E12FF05F8DF31C957B1" box="[129,388,1910,1934]" italics="true" pageId="4" pageNumber="5">Tope localito and horizon:</emphasis>
<materialsCitation id="E7DF9F1FFF8C9E12FE0EF8DE349E5728" country="Brazil" county="Tremembe" location="Outcrop" longitude="-45.898613" municipality="Sul" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Paraiba">
<location id="5268C399FF8C9E12FE0EF8DE31AF57B1" LSID="urn:lsid:plazi:treatment:DF1E2454FF8A9E1DFC7BF9E5350854FE:5268C399FF8C9E12FE0EF8DE31AF57B1" box="[394,482,1910,1934]" country="Brazil" county="Tremembe" longitude="-45.898613" municipality="Sul" name="Outcrop" pageId="4" pageNumber="5" stateProvince="Paraiba">Outcrop</location>
located on the right bank of the river
<collectingRegion id="95735BA0FF8C9E12FF64F83D31665792" box="[224,299,1941,1965]" country="Brazil" name="Paraiba" pageId="4" pageNumber="5">Paraíba</collectingRegion>
do
<collectingMunicipality id="B76C0F38FF8C9E12FED1F83D313A5792" box="[341,375,1941,1965]" pageId="4" pageNumber="5">Sul</collectingMunicipality>
,
<quantity id="904F38A7FF8C9E12FE07F83D31865792" box="[387,459,1941,1965]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="5" unit="km" value="1.6">1.6 km</quantity>
north*east of the municipality of
<collectingCounty id="BE69EDCEFF8C9E12FCD3F968338456E7" box="[855,969,1728,1752]" pageId="4" pageNumber="5">Tremembé</collectingCounty>
,
<collectingRegion id="95735BA0FF8C9E12FC51F968347356E7" box="[981,1086,1728,1752]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="5">São Paulo</collectingRegion>
,
<collectingCountry id="2FA0D5D2FF8C9E12FBCEF96834C856E7" box="[1098,1157,1728,1752]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
(22º94
<emphasis id="65C34950FF8C9E12FB52F917349156E7" box="[1238,1244,1727,1752]" italics="true" pageId="4" pageNumber="5">ʹ</emphasis>
50
<emphasis id="65C34950FF8C9E12FB72F917354C56E7" box="[1270,1281,1727,1752]" italics="true" pageId="4" pageNumber="5">ʹʹ</emphasis>
S;
<geoCoordinate id="3283F385FF8C9E12FAA3F96835FB56E7" box="[1319,1462,1727,1752]" degrees="45" direction="west" minutes="53" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="55" value="-45.898613">
45º53
<emphasis id="65C34950FF8C9E12FAE1F917352656E7" box="[1381,1387,1727,1752]" italics="true" pageId="4" pageNumber="5">ʹ</emphasis>
55
<emphasis id="65C34950FF8C9E12FA01F91735DD56E7" box="[1413,1424,1727,1752]" italics="true" pageId="4" pageNumber="5">ʹʹ</emphasis>
W
</geoCoordinate>
). Ŋese deposits are assigned to the Oligocene age Tremembé Formation, and lie within Taubaté Basin
</materialsCitation>
.
</paragraph>
<paragraph id="57089542FF8C9E12FCBDF89035635791" blockId="4.[825,1474,1848,1966]" pageId="4" pageNumber="5">
<emphasis id="65C34950FF8C9E12FCBDF89033E4576F" box="[825,937,1848,1872]" italics="true" pageId="4" pageNumber="5">Etomologo:</emphasis>
<materialsCitation id="E7DF9F1FFF8C9E12FC37F89035675791" location="Star of Magellan. Nis" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Sao Paulo">
Ŋe epithet is a noun in apposition and refers to
<taxonomicName id="90B7EEC1FF8C9E12FCBDF8FF33F45750" authority="Crucis" authorityName="Crucis" box="[825,953,1879,1903]" class="Insecta" family="Vespidae" genus="Alpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="65C34950FF8C9E12FCBDF8FF33F45750" box="[825,953,1879,1903]" italics="true" pageId="4" pageNumber="5">Alpha Crucis</emphasis>
</taxonomicName>
, also known as
<location id="5268C399FF8C9E12FBDEF8F035755750" LSID="urn:lsid:plazi:treatment:DF1E2454FF8A9E1DFC7BF9E5350854FE:5268C399FF8C9E12FBDEF8F035755750" box="[1114,1336,1879,1904]" name="Star of Magellan. Nis" pageId="4" pageNumber="5" stateProvince="Sao Paulo">Star of Magellan. Ŋis</location>
star (actually a multiple star system) represents, in the Brazilian national flag, the state of
<collectingRegion id="95735BA0FF8C9E12FC2AF83E345E5791" box="[942,1043,1942,1966]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="5">São Paulo</collectingRegion>
, where the fossil was found
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="03C8C5CAFF8D9E13FFF7F9F4346456F7" pageId="5" pageNumber="6" startId="5.[115,180,1628,1652]" targetBox="[114,1458,144,1600]" targetPageId="5" targetType="figure">
<paragraph id="57089542FF8D9E13FFF7F9F4346456F7" blockId="5.[113,1425,1628,1736]" pageId="5" pageNumber="6">
<emphasis id="65C34950FF8D9E13FFF7F9F43086564B" bold="true" box="[115,203,1628,1652]" pageId="5" pageNumber="6">Figure 4.</emphasis>
Skull bones of
<taxonomicName id="90B7EEC1FF8D9E13FEDDF9F431B8564B" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[345,501,1628,1652]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov.">
<emphasis id="65C34950FF8D9E13FEDDF9F431B8564B" box="[345,501,1628,1652]" italics="true" pageId="5" pageNumber="6">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF8D9E13FE7EF9F532CE564B" box="[506,643,1629,1653]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Collapsed fragments of the posterolateral region of the
<taxonomicName id="90B7EEC1FF8D9E13FB08F9F534AD564B" box="[1164,1248,1628,1652]" class="Amphibia" family="Caeciliidae" genus="Oscaecilia" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="15" phylum="Chordata" rank="species" species="basale">os basale</taxonomicName>
below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the
<taxonomicName id="90B7EEC1FF8D9E13FB21F9D134B456AF" box="[1189,1273,1656,1680]" class="Amphibia" family="Caeciliidae" genus="Oscaecilia" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="15" phylum="Chordata" rank="species" species="basale">os basale</taxonomicName>
in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations. Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G.
</paragraph>
</caption>
<subSubSection id="1FADC6C9FF8D9E1DFEF2F95F350854FE" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6" type="description">
<paragraph id="57089542FF8D9E13FEF2F95F31BB572E" blockId="5.[374,502,1783,1809]" box="[374,502,1783,1809]" pageId="5" pageNumber="6">
<emphasis id="65C34950FF8D9E13FEF2F95F31BB572E" bold="true" box="[374,502,1783,1809]" pageId="5" pageNumber="6">Description</emphasis>
</paragraph>
<paragraph id="57089542FF8D9E13FE1AF88A31835703" blockId="5.[113,763,1826,1982]" box="[414,462,1826,1852]" pageId="5" pageNumber="6">
<emphasis id="65C34950FF8D9E13FE1AF88A31835703" box="[414,462,1826,1852]" italics="true" pageId="5" pageNumber="6">Skull</emphasis>
</paragraph>
<paragraph id="57089542FF8D9E10FFF5F8E031A95712" blockId="5.[113,763,1826,1982]" lastBlockId="6.[129,778,1750,1837]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Ŋe skull is severely damaged and strongly dorsoventrally compressed (
<figureCitation id="CF8C89C7FF8D9E13FF7AF8C0317457BF" box="[254,313,1896,1920]" captionStart="Figure 3" captionStartId="4.[129,194,1627,1651]" captionTargetBox="[129,1473,144,1600]" captionTargetId="figure-140@4.[129,1473,144,1600]" captionTargetPageId="4" captionText="Figure 3. CT*scans of the two skull parts of Ymboirana acrux gen. et sp. nov. Top part in dorsal (A) and ventral (B) views. Boưom part in dorsal (C) and ventral (D) views. See Appendix for abbreviations.Scale bar = 1 mm." pageId="5" pageNumber="6">Fig. 3</figureCitation>
). Most of the bones are broken into short or indistinct fragments; however, both parietals and the pos* terior portion of the less pseudangular are well preserved. Some fragments are tentatively assigned to the nasopremaxilla, maxillopalatine, frontals, squamosals, quadrates, stapes,
<taxonomicName id="90B7EEC1FF8D9E13FA18F8B033275770" class="Amphibia" family="Caeciliidae" genus="Oscaecilia" kingdom="Animalia" order="Gymnophiona" pageId="5" pageNumber="15" phylum="Chordata" rank="species" species="basale">os basale</taxonomicName>
, and pseudodentaries. Many tooth crowns are also pre* sent, most of them disarticulated and a few still aưached to their pedicels and to their corresponding jaw bones. Given that the skull bones may be difficult to distinguish due to the damaged condition of the fossil, we also provide a figure of the skull in the Supporting Information (
<figureCitation id="CF8C89C7FF8E9E10FE0EF95E319C5731" box="[394,465,1782,1806]" captionStart="Figure 2" captionStartId="3.[114,179,1836,1860]" captionTargetBox="[238,1336,144,1808]" captionTargetId="figure-8@3.[238,1336,144,1808]" captionTargetPageId="3" captionText="Figure 2. Fossil slabs containing the holotype (DGM 1462*R) of Ymboirana acrux gen. et sp. nov. (A1, B1). CT*scans of the part (A2) and counterpart (B2) of the holotype.Scale bar = 1 cm." pageId="6" pageNumber="7">Fig. S2</figureCitation>
) with an outline line drawing and using lighter shades of colours.
</paragraph>
<caption id="03C8C5CAFF8E9E10FF06F9F3341E5693" pageId="6" pageNumber="7" startId="6.[130,195,1627,1651]" targetBox="[129,1473,144,1600]" targetPageId="6" targetType="figure">
<paragraph id="57089542FF8E9E10FF06F9F3341E5693" blockId="6.[129,1460,1627,1708]" pageId="6" pageNumber="7">
<emphasis id="65C34950FF8E9E10FF06F9F33097564B" bold="true" box="[130,218,1627,1652]" pageId="6" pageNumber="7">Figure 5.</emphasis>
Lower jaw of
<taxonomicName id="90B7EEC1FF8E9E10FED9F9F431B4564C" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[349,505,1628,1652]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov.">
<emphasis id="65C34950FF8E9E10FED9F9F431B4564C" box="[349,505,1628,1652]" italics="true" pageId="6" pageNumber="7">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF8E9E10FE79F9F432CB564C" box="[509,646,1628,1652]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Mandibles in dorsal (A) and ventral (B) views. Less pseudangular in lateral (C), posterior oblique (D), and dorsal (E) views. Doưed lines indicate the region of the contact between the pseudangular and pseudodentary. See Appendix for abbreviations. Scale bars = 1 mm; upper scale bar is for A, B, and the lower one is for CE.
</paragraph>
</caption>
<paragraph id="57089542FF8E9E10FF05F8E634665773" blockId="6.[129,778,1870,1988]" lastBlockId="6.[825,1474,1750,1868]" pageId="6" pageNumber="7">
<emphasis id="65C34950FF8E9E10FF05F8E631605759" box="[129,301,1870,1894]" italics="true" pageId="6" pageNumber="7">Nasopremaxillae:</emphasis>
Fragments of the presumed nasopremaxillae are present, identified primarily on the basis of their position. Due to their poor preservation, few diagnostic features can be ob* served. However, based on the relative position of the fragments in relation to the lower jaw, we interpret the nasopremaxillae as forming an anteriorly projected snout and, thus, the mouth to be subterminal. Several other fragments were also preserved but lack diagnostic features.
</paragraph>
<paragraph id="57089542FF8E9E11FCBDF8C531995706" blockId="6.[825,1474,1901,1988]" lastBlockId="7.[113,763,1700,1849]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="65C34950FF8E9E10FCBDF8C533A957BA" box="[825,996,1901,1925]" italics="true" pageId="6" pageNumber="7">Maxillopalatines:</emphasis>
Both maxillopalatines are partially pre* served though severely damaged. Ŋe outer tooth rows of both maxillopalatines are present but incompletely preserved, so we could recognize only 12 pedicels. A fragment tentatively inter* preted as a portion of the less vomeropalatine tooth row is lo* cated medially and bears seven pedicels. Several other fragments, possibly belonging to the maxillopalatines, are present but do not exhibit any diagnostic features.
</paragraph>
<caption id="03C8C5CAFF8F9E11FFF7F9F33419564C" box="[115,1108,1627,1652]" pageId="7" pageNumber="8" startId="7.[115,180,1627,1651]" targetBox="[114,1458,144,1600]" targetPageId="7" targetType="figure">
<paragraph id="57089542FF8F9E11FFF7F9F33419564C" blockId="7.[115,1108,1627,1652]" box="[115,1108,1627,1652]" pageId="7" pageNumber="8">
<emphasis id="65C34950FF8F9E11FFF7F9F33086564B" bold="true" box="[115,203,1627,1652]" pageId="7" pageNumber="8">Figure 6.</emphasis>
Isolated tooth crowns found near the skull of
<taxonomicName id="90B7EEC1FF8F9E11FDFDF9F43358564C" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[633,789,1628,1652]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov">
<emphasis id="65C34950FF8F9E11FDFDF9F43358564C" box="[633,789,1628,1652]" italics="true" pageId="7" pageNumber="8">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF8F9E11FC93F9F433D6564C" box="[791,923,1628,1652]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="57089542FF8F9E11FFF5F8F2355D5767" blockId="7.[113,762,1882,2000]" lastBlockId="7.[810,1459,1699,1880]" pageId="7" pageNumber="8">
<emphasis id="65C34950FF8F9E11FFF5F8F23080574D" box="[113,205,1882,1906]" italics="true" pageId="7" pageNumber="8">Parietals:</emphasis>
Ŋese are the best*preserved skull bones in the spe* cimen and are almost complete, except for a short dorsolateral portion missing from the less. In dorsal view, except for their anterolateral pointed corners, both parietals exhibit a roughly trapezoidal shape (
<figureCitation id="CF8C89C7FF8F9E11FC69F90B34765683" box="[1005,1083,1699,1724]" captionStart="Figure 4" captionStartId="5.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-107@5.[114,1458,144,1600]" captionTargetPageId="5" captionText="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." pageId="7" pageNumber="8">Figs 4A</figureCitation>
,
<figureCitation id="CF8C89C7FF8F9E11FBC3F90C343F5683" box="[1095,1138,1700,1724]" captionStart="Figure 12" captionStartId="15.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-115@15.[114,1458,144,1600]" captionTargetPageId="15" captionText="Figure 12. Paired parietals of four different caecilian species, all in dorsal view.Ymboirana acrux gen.et sp. nov.(A), Rhinatrema bivitatum (B), Tophlonectes natans (C), and Herpele squalostoma (D). See Appendix for abbreviations. Scale bar = 1 mm." pageId="7" pageNumber="8">12A</figureCitation>
). Ŋe dorsal surface is smooth, lacking ornamentations in the form of small neurovascular for* amina that are common in some caecilians. Ŋe parietals are also flat medially but have a pronounced ventrally oriented slope at their posterolateral margins. Collapsed fragments of the
<taxonomicName id="90B7EEC1FF8F9E11FA18F88933255767" class="Amphibia" family="Caeciliidae" genus="Oscaecilia" kingdom="Animalia" order="Gymnophiona" pageId="7" pageNumber="15" phylum="Chordata" rank="species" species="basale">os basale</taxonomicName>
cover the ventral surfaces of the parietals.
</paragraph>
<paragraph id="57089542FF8F9E1EFCAEF8D132E2572D" blockId="7.[810,1459,1913,2000]" lastBlockId="8.[129,777,1754,1810]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="65C34950FF8F9E11FCAEF8D133CC57AE" box="[810,897,1913,1937]" italics="true" pageId="7" pageNumber="8">Frontals:</emphasis>
Located immediately anterior to both parietals, lie two poorly preserved bones, tentatively identified as parts of the frontals. Ŋey are broken and show an irregular shape, and similar to the parietals, the preserved portions of the dorsal sur* face of both frontals lack small neurovascular foramina.
</paragraph>
<caption id="03C8C5CAFF809E1EFF05F9F231085694" pageId="8" pageNumber="9" startId="8.[129,194,1626,1650]" targetBox="[129,1473,142,1598]" targetPageId="8" targetType="figure">
<paragraph id="57089542FF809E1EFF05F9F231085694" blockId="8.[129,1471,1626,1707]" pageId="8" pageNumber="9">
<emphasis id="65C34950FF809E1EFF05F9F23095564C" bold="true" box="[129,216,1626,1651]" pageId="8" pageNumber="9">Figure 7.</emphasis>
Vertebral column of
<taxonomicName id="90B7EEC1FF809E1EFE19F9F23274564D" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[413,569,1626,1650]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov.">
<emphasis id="65C34950FF809E1EFE19F9F23274564D" box="[413,569,1626,1650]" italics="true" pageId="8" pageNumber="9">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF809E1EFDBAF9F3328A564D" box="[574,711,1627,1651]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(A). Articulated trunk vertebrae from the posterior portion of the body in dorsal (B) and lateral (C) views. See Appendix for abbreviations. Upper scale bar = 1 cm. Lower scale bar = 1 mm; upper scale bar is for A, and the lower one is for B, C.
</paragraph>
</caption>
<paragraph id="57089542FF809E1EFF05F89B33475796" blockId="8.[129,778,1843,1961]" pageId="8" pageNumber="9">
<emphasis id="65C34950FF809E1EFF05F89B30B65774" box="[129,251,1843,1867]" italics="true" pageId="8" pageNumber="9">Squamosals:</emphasis>
Some fragments are tentatively assigned to the less squamosal (
<figureCitation id="CF8C89C7FF809E1EFF79F8FA31085755" box="[253,325,1874,1898]" captionStart="Figure 4" captionStartId="5.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-107@5.[114,1458,144,1600]" captionTargetPageId="5" captionText="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." pageId="8" pageNumber="9">Fig. 4F</figureCitation>
). Ŋey are located close to the less quadrate and above the less parietal. Ŋe squamosal is roughly rectangular and bears a postorbital process with a slightly concave margin.
</paragraph>
<paragraph id="57089542FF809E1EFCBDF97333DB572D" blockId="8.[825,1474,1754,1810]" pageId="8" pageNumber="9">Ŋe only evidence of ornamentation is a single neurovascular foramen.</paragraph>
<paragraph id="57089542FF809E1FFCBDF89B326E574E" blockId="8.[825,1475,1843,1961]" lastBlockId="9.[113,763,1756,1905]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="65C34950FF809E1EFCBDF89B33EB5774" box="[825,934,1843,1867]" italics="true" pageId="8" pageNumber="9">Quadrates:</emphasis>
Both quadrates are preserved but broken (
<figureCitation id="CF8C89C7FF809E1EFAD6F89B35C15774" box="[1362,1420,1843,1867]" captionStart="Figure 3" captionStartId="4.[129,194,1627,1651]" captionTargetBox="[129,1473,144,1600]" captionTargetId="figure-140@4.[129,1473,144,1600]" captionTargetPageId="4" captionText="Figure 3. CT*scans of the two skull parts of Ymboirana acrux gen. et sp. nov. Top part in dorsal (A) and ventral (B) views. Boưom part in dorsal (C) and ventral (D) views. See Appendix for abbreviations.Scale bar = 1 mm." pageId="8" pageNumber="9">Figs 3</figureCitation>
,
<figureCitation id="CF8C89C7FF809E1EFA12F89B35FC5774" box="[1430,1457,1843,1867]" captionStart="Figure 4" captionStartId="5.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-107@5.[114,1458,144,1600]" captionTargetPageId="5" captionText="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." pageId="8" pageNumber="9">4B</figureCitation>
). Portions of the less quadrate are located just above the articular facet of the pseudangular and not so widely separated from the maxillopalatines. Ŋe right quadrate is preserved far forward, next to fragments interpreted as parts of the pseudodentary. Due to the damaged condition of the squamosals, the degree of dorsal exposure of the quadrates cannot be determined. Ŋe pterygoid process of the less quadrate is present and elongate. A distinctive otic process is absent from both quadrates.
</paragraph>
<caption id="03C8C5CAFF819E1FFFF7F9F332655694" pageId="9" pageNumber="10" startId="9.[115,180,1627,1651]" targetBox="[114,1458,144,1600]" targetPageId="9" targetType="figure">
<paragraph id="57089542FF819E1FFFF7F9F332655694" blockId="9.[113,1454,1627,1708]" pageId="9" pageNumber="10">
<emphasis id="65C34950FF819E1FFFF7F9F33087564B" bold="true" box="[115,202,1627,1652]" pageId="9" pageNumber="10">Figure 8.</emphasis>
Postcranial elements of
<taxonomicName id="90B7EEC1FF819E1FFE2AF9F43207564C" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[430,586,1628,1652]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov">
<emphasis id="65C34950FF819E1FFE2AF9F43207564C" box="[430,586,1628,1652]" italics="true" pageId="9" pageNumber="10">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF819E1FFDCAF9F4329F564C" box="[590,722,1628,1652]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. Atlas in dorsal (A) and ventral (B) views. Ventral part of atlas (i.e. atlantal centrum and cotyles) in anterior (C), posterior (D), and oblique less lateral (E) views. Second vertebra in dorsal (F) and ventral (G) views. See Appendix for abbreviations. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="57089542FF819E1FFFF5F83B34C2570D" blockId="9.[113,763,1938,1993]" lastBlockId="9.[810,1459,1756,1842]" pageId="9" pageNumber="10">
<emphasis id="65C34950FF819E1FFFF5F83B30FA5795" box="[113,183,1939,1962]" italics="true" pageId="9" pageNumber="10">Stapes:</emphasis>
A small fragment located on the posterodorsal portion of the skull, close to the less quadrate, is tentatively interpreted as the less stapes (
<figureCitation id="CF8C89C7FF819E1FFC47F974345F56CB" box="[963,1042,1756,1780]" captionStart="Figure 4" captionStartId="5.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-107@5.[114,1458,144,1600]" captionTargetPageId="5" captionText="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." pageId="9" pageNumber="10">Figs 4G</figureCitation>
,
<figureCitation id="CF8C89C7FF819E1FFB98F974340A56CB" box="[1052,1095,1756,1780]" captionStart="Figure 13" captionStartId="16.[130,195,1627,1651]" captionTargetBox="[129,1473,144,1600]" captionTargetId="figure-149@16.[129,1473,144,1600]" captionTargetPageId="16" captionText="Figure 13. Less stapes of six different caecilian species, all in lateral view.Fragment tentatively identified as the stapes of Ymboirana acrux gen. et sp.nov.(A), Potomotophlus kaupii (B), Tophlonectes natans (C), Caecilia sp. (D), Siphonops paulensis (E), and Rhinatrema bivitatum (F). See Appendix for abbreviations.Scale bar = 1 mm." pageId="9" pageNumber="10">13A</figureCitation>
). Ŋis fragment is composed of two different portions (i.e. a short and slender stapedial style and a small footplate) and is imperforate.
</paragraph>
<paragraph id="57089542FF819E1CFCAEF8FC33C15708" blockId="9.[810,1459,1875,1993]" lastBlockId="10.[825,1474,1729,1847]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="65C34950FF819E1FFCAEF8FC33DD5754" box="[810,912,1875,1899]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="90B7EEC1FF819E1FFCAEF8FC33C65754" box="[810,907,1875,1899]" class="Amphibia" family="Caeciliidae" genus="Oscaecilia" kingdom="Animalia" order="Gymnophiona" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="basale">Os basale</taxonomicName>
:
</emphasis>
Several small broken parts, scaưered about the skull but mainly at its posterior portion appear to represent different regions of the
<taxonomicName id="90B7EEC1FF819E1FFC42F83B346A5795" box="[966,1063,1938,1962]" class="Amphibia" family="Caeciliidae" genus="Oscaecilia" kingdom="Animalia" order="Gymnophiona" pageId="9" pageNumber="15" phylum="Chordata" rank="species" species="basale">os basale</taxonomicName>
. One of these fragments, located just behind the less pseudangular and still articulated with the atlas, corresponds to a posterior portion of the
<taxonomicName id="90B7EEC1FF829E1CFDA4F96A323656E6" box="[544,635,1729,1753]" class="Amphibia" family="Caeciliidae" genus="Oscaecilia" kingdom="Animalia" order="Gymnophiona" pageId="10" pageNumber="15" phylum="Chordata" rank="species" species="basale">os basale</taxonomicName>
, including the less occipital condyle (
<figureCitation id="CF8C89C7FF829E1CFEEDF948319C56C6" box="[361,465,1760,1785]" captionStart="Figure 4" captionStartId="5.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-107@5.[114,1458,144,1600]" captionTargetPageId="5" captionText="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." pageId="10" pageNumber="11">Fig. 4CE</figureCitation>
; Supporting Information,
<figureCitation id="CF8C89C7FF829E1CFD67F94830D65727" captionStart="Figure 1" captionStartId="1.[113,178,1960,1984]" captionTargetBox="[116,1456,1173,1930]" captionTargetId="figure-558@1.[114,1458,1171,1932]" captionTargetPageId="1" captionText="Figure 1. Map of the Taubaté Basin in Southeastern Brazil, showing the type locality for Ymboirana acrux gen. et sp.nov.." pageId="10" pageNumber="11">Fig. S1</figureCitation>
). A jugular foramen is present just anterior to the condyle. Ŋe region of the otic capsule is located anterior to the jugular foramen, but its external wall is badly damaged, whereas parts of the internal walls of the otic capsule are beưer preserved and bear the perilymphatic foramen and other foramina for branches of the auditory nerve. A large fragment preserved collapsed on the ventral surface of the right parietal bears a carotid for* amen (
<figureCitation id="CF8C89C7FF829E1CFC00F948338256C6" box="[900,975,1760,1785]" captionStart="Figure 4" captionStartId="5.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-107@5.[114,1458,144,1600]" captionTargetPageId="5" captionText="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." pageId="10" pageNumber="11">Fig. 4A</figureCitation>
), and lateral to this foramen, a lip of bone with a distinctive articular surface is interpreted as the basipterygoid process.
</paragraph>
<caption id="03C8C5CAFF829E1CFF05F9F5331556AE" pageId="10" pageNumber="11" startId="10.[129,194,1629,1653]" targetBox="[129,1473,145,1601]" targetPageId="10" targetType="figure">
<paragraph id="57089542FF829E1CFF05F9F5331556AE" blockId="10.[129,1457,1629,1681]" pageId="10" pageNumber="11">
<emphasis id="65C34950FF829E1CFF05F9F53094564A" bold="true" box="[129,217,1629,1653]" pageId="10" pageNumber="11">Figure 9.</emphasis>
Postcranial elements of
<taxonomicName id="90B7EEC1FF829E1CFE38F9F53215564A" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[444,600,1629,1653]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="acrux" status="gen. et sp. nov">
<emphasis id="65C34950FF829E1CFE38F9F53215564A" box="[444,600,1629,1653]" italics="true" pageId="10" pageNumber="11">Ymboirana acrux</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF0F42BFF829E1CFDD9F9F632AD564A" box="[605,736,1630,1654]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. Trunk vertebra in dorsal (A), ventral (B), and oblique less lateral (C) views. Ribs (D, E) in lateral view. See Appendix for abbreviations. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="57089542FF829E1CFBC9F8FF34E2574F" blockId="10.[1101,1199,1879,1904]" box="[1101,1199,1879,1904]" pageId="10" pageNumber="11">
<emphasis id="65C34950FF829E1CFBC9F8FF34E2574F" box="[1101,1199,1879,1904]" italics="true" pageId="10" pageNumber="11">Lower jaw</emphasis>
</paragraph>
<paragraph id="57089542FF829E1DFCBDF8D532145200" blockId="10.[825,1474,1917,1973]" lastBlockId="11.[113,763,144,575]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="65C34950FF829E1CFCBDF8D5338057AA" box="[825,973,1917,1941]" italics="true" pageId="10" pageNumber="11">Pseudangulars:</emphasis>
Only the well*preserved posteriormost portion of the less pseudangular is discernable (
<figureCitation id="CF8C89C7FF829E1CFB48F8343555578A" box="[1228,1304,1948,1973]" captionStart="Figure 4" captionStartId="5.[115,180,1628,1652]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-107@5.[114,1458,144,1600]" captionTargetPageId="5" captionText="Figure 4. Skull bones of Ymboirana acrux gen. et sp. nov. Collapsed fragments of the posterolateral region of the os basale below the right parietal in ventral oblique view (A). Less quadrate and pseudangular in lateral view (B). Less occipital region of the os basale in lateral (C), posterior (D), and dorsal (E) views. Fragments of the less squamosal in lateral view (F). Fragment tentatively assigned to the stapes (G). See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for AF, and the lower one is for G." pageId="10" pageNumber="11">Figs 4B</figureCitation>
,
<figureCitation id="CF8C89C7FF829E1CFAA7F8343512578B" box="[1315,1375,1948,1972]" captionStart="Figure 5" captionStartId="6.[130,195,1627,1651]" captionTargetBox="[129,1473,144,1600]" captionTargetId="figure-131@6.[129,1473,144,1600]" captionTargetPageId="6" captionText="Figure 5. Lower jaw of Ymboirana acrux gen. et sp. nov. Mandibles in dorsal (A) and ventral (B) views.Less pseudangular in lateral (C), posterior oblique (D), and dorsal (E) views. Doưed lines indicate the region of the contact between the pseudangular and pseudodentary.See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for A, B, and the lower one is for CE." pageId="10" pageNumber="11">5CE</figureCitation>
,
<figureCitation id="CF8C89C7FF829E1CFAEFF83535D8578A" box="[1387,1429,1949,1973]" captionStart="Figure 14" captionStartId="17.[112,177,932,956]" captionTargetBox="[114,1458,143,904]" captionTargetId="figure-536@17.[114,1458,143,904]" captionTargetPageId="17" captionText="Figure 14. Less pseudangulars of four different caecilian species. Ymboirana acrux gen. et sp. nov. in lateral (A1) and dorsal (A2) views. Rhinatrema bivitatum in lateral (B1) and dorsal (B2) views. Scolecomorphus ulugurensis in lateral (C1) and dorsal (C2) views.Tophlonectes natans in lateral (D1) and dorsal (D2) views. See Appendix for abbreviations. Scale bar = 1 mm." pageId="10" pageNumber="11">14A</figureCitation>
). In lateral view, a recurved retroarticular process is present, bearing a well*developed lateral crest. Ŋe sutural surface that contacts the pseudodentary is excavated in the anterolateral margin. Ŋe processus condyloides is limited to a weakly developed process. In dorsal view, the articular surface is anteriorly positioned, concave, and surrounded posteriorly by the short processus condyloides. Ŋere are no signs of a preserved fossa or a canalis primordialis immediately anterior to the articular surface. Ŋe distinct processus internus is present, anteromedially oriented, and shows a roughly quadrangular outline. Ŋe retroarticular process curves medially mainly along its posterior portion. In medial view, two foramina of the ramus intermandibularis are present, one located at the posterior base of the processus internus and the other excavated anteriorly to it.
</paragraph>
<paragraph id="57089542FF839E1DFFF5FDC932BF538D" blockId="11.[113,763,609,946]" pageId="11" pageNumber="12">
<emphasis id="65C34950FF839E1DFFF5FDC931585246" box="[113,277,609,633]" italics="true" pageId="11" pageNumber="12">Pseudodentaries:</emphasis>
Both pseudodentaries are partially preserved, including the region of the mandibular symphysis (
<figureCitation id="CF8C89C7FF839E1DFD00FD2832A452A8" box="[644,745,640,664]" captionStart="Figure 5" captionStartId="6.[130,195,1627,1651]" captionTargetBox="[129,1473,144,1600]" captionTargetId="figure-131@6.[129,1473,144,1600]" captionTargetPageId="6" captionText="Figure 5. Lower jaw of Ymboirana acrux gen. et sp. nov. Mandibles in dorsal (A) and ventral (B) views.Less pseudangular in lateral (C), posterior oblique (D), and dorsal (E) views. Doưed lines indicate the region of the contact between the pseudangular and pseudodentary.See Appendix for abbreviations.Scale bars = 1 mm; upper scale bar is for A, B, and the lower one is for CE." pageId="11" pageNumber="12">Fig. 5A, B</figureCitation>
). Whereas the less one is represented by its anteriormost portion, in the right one the posterior portion is also preserved. Pedicels are present in the dorsal surface of the right pseudodentary, al* though the exact number is difficult to estimate. Ŋe pedicels are arranged in two rows, with definitive inner mandibular (i.e. splenial) pedicels being limited to the symphyseal region (based on the preserved pedicels, at least four are present on the right pseudodentary). Ŋe outer tooth row is also represented only by damaged pedicels, and we are able to recognize just 16 of them.
</paragraph>
<paragraph id="57089542FF839E1DFE0DFC7A31AF53D4" blockId="11.[393,482,978,1003]" box="[393,482,978,1003]" pageId="11" pageNumber="12">
<emphasis id="65C34950FF839E1DFE0DFC7A31AF53D4" box="[393,482,978,1003]" italics="true" pageId="11" pageNumber="12">Dentition</emphasis>
</paragraph>
<paragraph id="57089542FF839E1DFFF5FC5031025576" blockId="11.[113,763,1016,1353]" pageId="11" pageNumber="12">
We recognized at least 48 scaưered tooth crowns near the skull (
<figureCitation id="CF8C89C7FF839E1DFFF8FBBF30F55410" box="[124,184,1047,1071]" captionStart="Figure 6" captionStartId="7.[115,180,1627,1651]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-181@7.[114,1458,144,1600]" captionTargetPageId="7" captionText="Figure 6. Isolated tooth crowns found near the skull of Ymboirana acruxgen.et sp. nov.. Scale bar = 1 mm." pageId="11" pageNumber="12">Fig. 6</figureCitation>
). Some teeth were also preserved still aưached to the pedicels. Most of them are similar in size (around
<quantity id="904F38A7FF839E1DFD0FFB9E32905471" box="[651,733,1078,1102]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="mm" value="0.5">0.5 mm</quantity>
in length) and shape: slender, conical, recurved, and monocusped. Based on the size of the preserved pedicels, the teeth of the inner rows are a liưle smaller than those of the outer rows. Ŋe apex is pointed but lateral flanges are seemingly absent. Due to the dam* aged condition of the fossil, it was not possible to verify the exact total number of teeth; however, the pedicel arrangement allows confirmation of the ancestral caecilian double tooth rows in the upper and lower jaws.
</paragraph>
<paragraph id="57089542FF839E1DFEFFFAC231BD55BC" blockId="11.[113,763,1386,1573]" box="[379,496,1386,1411]" pageId="11" pageNumber="12">
<emphasis id="65C34950FF839E1DFEFFFAC231BD55BC" box="[379,496,1386,1411]" italics="true" pageId="11" pageNumber="12">Postcranium</emphasis>
</paragraph>
<paragraph id="57089542FF839E1DFFF5FA2732B7561A" blockId="11.[113,763,1386,1573]" pageId="11" pageNumber="12">
Postcranial elements assigned to
<taxonomicName id="90B7EEC1FF839E1DFE40FA2732205598" authorityName="Santos &amp; Wilkinson &amp; Graziella &amp; Ribeiro &amp; Carvalho &amp; Zaher" authorityYear="2024" box="[452,621,1423,1447]" class="Amphibia" family="Typhlonectidae" genus="Ymboirana" kingdom="Animalia" order="Gymnophiona" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acrux">
<emphasis id="65C34950FF839E1DFE40FA2732205598" box="[452,621,1423,1447]" italics="true" pageId="11" pageNumber="12">Ymboirana acrux</emphasis>
</taxonomicName>
comprise the fragmented atlas followed by 66 subsequent vertebrae (most of them still articulated, although in the median portion of the vertebral column they are scaưered) and some ribs (
<figureCitation id="CF8C89C7FF839E1DFD08FA453288563A" box="[652,709,1517,1541]" captionStart="Figure 7" captionStartId="8.[129,194,1626,1650]" captionTargetBox="[129,1473,142,1598]" captionTargetId="figure-120@8.[129,1473,142,1598]" captionTargetPageId="8" captionText="Figure 7. Vertebral column of Ymboirana acrux gen. et sp. nov. (A). Articulated trunk vertebrae from the posterior portion of the body in dorsal (B) and lateral (C) views. See Appendix for abbreviations. Upper scale bar = 1 cm. Lower scale bar = 1 mm; upper scale bar is for A, and the lower one is for B, C." pageId="11" pageNumber="12">Fig. 7</figureCitation>
). An unknown number of posteriormost vertebrae are not preserved.
</paragraph>
<paragraph id="57089542FF839E1DFFF5F9EE33E150D9" blockId="11.[113,763,1606,1974]" lastBlockId="11.[810,1460,144,230]" pageId="11" pageNumber="12">
<emphasis id="65C34950FF839E1DFFF5F9EE30E65661" box="[113,171,1606,1630]" italics="true" pageId="11" pageNumber="12">Atlas:</emphasis>
Broken elements interpreted as the atlantal centrum and the neural arch were preserved separately in the two slabs (
<figureCitation id="CF8C89C7FF839E1DFD54F9CD30E156A2" captionStart="Figure 8" captionStartId="9.[115,180,1627,1651]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-148@9.[114,1458,144,1600]" captionTargetPageId="9" captionText="Figure 8. Postcranial elements of Ymboirana acrux gen. et sp. nov.. Atlas in dorsal (A) and ventral (B) views. Ventral part of atlas (i.e. atlantal centrum and cotyles) in anterior (C), posterior (D), and oblique less lateral (E) views. Second vertebra in dorsal (F) and ventral (G) views.See Appendix for abbreviations.Scale bar = 1 mm." pageId="11" pageNumber="12">Figs 8AE</figureCitation>
,
<figureCitation id="CF8C89C7FF839E1DFF3EF92C30A756A3" box="[186,234,1668,1692]" captionStart="Figure 15" captionStartId="18.[129,194,1899,1923]" captionTargetBox="[129,1473,1111,1872]" captionTargetId="figure-568@18.[129,1473,1111,1872]" captionTargetPageId="18" captionText="Figure 15. Atlases of four different typhlonectid caecilians. Ymboirana acrux gen. et sp. nov. in anterior (A1) and dorsal (A2) views. Chthonerpeton indistinctum in anterior (B1) and dorsal (B2) views. Potomotophlus kaupii in anterior (C1) and dorsal (C2) views. Nectocaecilia petersii in anterior (D1) and dorsal (D2) views. See Appendix for abbreviations.Scale bar = 1 mm." pageId="11" pageNumber="12">15A</figureCitation>
<figureCitation id="CF8C89C7FF839E1DFF6EF92C30BE56A3" box="[234,243,1668,1692]" captionStart="Figure 1" captionStartId="1.[113,178,1960,1984]" captionTargetBox="[116,1456,1173,1930]" captionTargetId="figure-558@1.[114,1458,1171,1932]" captionTargetPageId="1" captionText="Figure 1. Map of the Taubaté Basin in Southeastern Brazil, showing the type locality for Ymboirana acrux gen. et sp.nov.." pageId="11" pageNumber="12">1</figureCitation>
, A
<figureCitation id="CF8C89C7FF839E1DFE96F92D315356A2" box="[274,286,1669,1693]" captionStart="Figure 2" captionStartId="3.[114,179,1836,1860]" captionTargetBox="[238,1336,144,1808]" captionTargetId="figure-8@3.[238,1336,144,1808]" captionTargetPageId="3" captionText="Figure 2. Fossil slabs containing the holotype (DGM 1462*R) of Ymboirana acrux gen. et sp. nov. (A1, B1). CT*scans of the part (A2) and counterpart (B2) of the holotype.Scale bar = 1 cm." pageId="11" pageNumber="12">2</figureCitation>
). Both atlantal cotyles are present and fused anteroventrally but with a pronounced median notch between them. Ŋe spinal nerve foramen is preserved only in the pos* terior surface of the less cotyle and is oriented posterolaterally. Ŋe centrum has a length considerably shorter than the fol* lowing vertebrae, and its ventral surface is smooth, bearing a medially positioned fossa and no hypapophyseal keel. Ŋe pos* terior cotyle has a roughly oval outline and the notochordal fossa is deep. Only a few portions of the neural arch were preserved. Two separated and anteriorly directed projections are present and are interpreted as the anterior bases of the lateral pedicles. Ŋe dorsal surface of the atlas is smooth, with no evidence of a neural spine.
</paragraph>
<paragraph id="57089542FF839E1DFCAEFEA0339F51C4" blockId="11.[810,1460,264,507]" pageId="11" pageNumber="12">
<emphasis id="65C34950FF839E1DFCAEFEA0339E511F" box="[810,979,264,288]" italics="true" pageId="11" pageNumber="12">Second vertebra:</emphasis>
Ŋe second vertebra is preserved still ar* ticulated to the atlas, but severely damaged (
<figureCitation id="CF8C89C7FF839E1DFA8CFE8F353F5100" box="[1288,1394,295,319]" captionStart="Figure 8" captionStartId="9.[115,180,1627,1651]" captionTargetBox="[114,1458,144,1600]" captionTargetId="figure-148@9.[114,1458,144,1600]" captionTargetPageId="9" captionText="Figure 8. Postcranial elements of Ymboirana acrux gen. et sp. nov.. Atlas in dorsal (A) and ventral (B) views. Ventral part of atlas (i.e. atlantal centrum and cotyles) in anterior (C), posterior (D), and oblique less lateral (E) views. Second vertebra in dorsal (F) and ventral (G) views.See Appendix for abbreviations.Scale bar = 1 mm." pageId="11" pageNumber="12">Fig. 8F, G</figureCitation>
). Ŋe neural arch is completely collapsed over the centrum. Ŋe two prezygapophyses are preserved and exhibit a flat articular surface. On the ventral surface of the centrum, there is a well*developed hypapophyseal keel, but basapophyseal processes seem to be ab* sent. No rib*like elements can be confidently identified near the second vertebra.
</paragraph>
<paragraph id="57089542FF839E1DFCAEFDB434355392" blockId="11.[810,1459,540,941]" pageId="11" pageNumber="12">
<emphasis id="65C34950FF839E1DFCAEFDB4339C520B" box="[810,977,540,564]" italics="true" pageId="11" pageNumber="12">Trunk vertebrae:</emphasis>
Ŋe exact number of preserved trunk verte* brae (
<figureCitation id="CF8C89C7FF839E1DFCEEFD9433E7526C" box="[874,938,572,596]" captionStart="Figure 7" captionStartId="8.[129,194,1626,1650]" captionTargetBox="[129,1473,142,1598]" captionTargetId="figure-120@8.[129,1473,142,1598]" captionTargetPageId="8" captionText="Figure 7. Vertebral column of Ymboirana acrux gen. et sp. nov. (A). Articulated trunk vertebrae from the posterior portion of the body in dorsal (B) and lateral (C) views. See Appendix for abbreviations. Upper scale bar = 1 cm. Lower scale bar = 1 mm; upper scale bar is for A, and the lower one is for B, C." pageId="11" pageNumber="12">Figs 7</figureCitation>
,
<figureCitation id="CF8C89C7FF839E1DFC3DFD9433BB526B" box="[953,1014,572,596]" captionStart="Figure 9" captionStartId="10.[129,194,1629,1653]" captionTargetBox="[129,1473,145,1601]" captionTargetId="figure-150@10.[129,1473,145,1601]" captionTargetPageId="10" captionText="Figure 9. Postcranial elements of Ymboirana acrux gen. et sp.nov.. Trunk vertebra in dorsal (A), ventral (B), and oblique less lateral (C) views. Ribs (D, E) in lateral view. See Appendix for abbreviations. Scale bar = 1 mm." pageId="11" pageNumber="12">9AC</figureCitation>
) cannot be determined because of exten* sive fragmentation and disarticulation. All of these vertebrae show diagenetic dorsoventral compression. Ŋey are elongate anteroposteriorly and have medially constricted amphicoelous centra, well*developed basapophyseal processes projecting anteroventrally, a pronounced hypapophyseal keel, and low neural arches. Poorly developed neural spines, limited to faint ridges, are present in some, but not all, of the posteriormost pre* served vertebrae. Basapophyseal processes bear well*developed and convex ventral ridges, and this feature is particularly pro* nounced in some vertebrae preserved at the posterior region of the
<typeStatus id="880C2BE0FF839E1DFCD4FC3C33E15393" box="[848,940,916,940]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
(
<figureCitation id="CF8C89C7FF839E1DFC39FC3C34695393" box="[957,1060,916,940]" captionStart="Figure 7" captionStartId="8.[129,194,1626,1650]" captionTargetBox="[129,1473,142,1598]" captionTargetId="figure-120@8.[129,1473,142,1598]" captionTargetPageId="8" captionText="Figure 7. Vertebral column of Ymboirana acrux gen. et sp. nov. (A). Articulated trunk vertebrae from the posterior portion of the body in dorsal (B) and lateral (C) views. See Appendix for abbreviations. Upper scale bar = 1 cm. Lower scale bar = 1 mm; upper scale bar is for A, and the lower one is for B, C." pageId="11" pageNumber="12">Figs 7B, C</figureCitation>
,
<figureCitation id="CF8C89C7FF839E1DFBABFC3C34255393" box="[1071,1128,916,940]" captionStart="Figure 9" captionStartId="10.[129,194,1629,1653]" captionTargetBox="[129,1473,145,1601]" captionTargetId="figure-150@10.[129,1473,145,1601]" captionTargetPageId="10" captionText="Figure 9. Postcranial elements of Ymboirana acrux gen. et sp.nov.. Trunk vertebra in dorsal (A), ventral (B), and oblique less lateral (C) views. Ribs (D, E) in lateral view. See Appendix for abbreviations. Scale bar = 1 mm." pageId="11" pageNumber="12">9B, C</figureCitation>
).
</paragraph>
<paragraph id="57089542FF839E1DFCAEFC66350854FE" blockId="11.[810,1460,974,1218]" pageId="11" pageNumber="12">
<emphasis id="65C34950FF839E1DFCAEFC66331153D9" box="[810,860,974,998]" italics="true" pageId="11" pageNumber="12">Ribs:</emphasis>
Preserved ribs are hard to distinguish. Some are still articu* lated with vertebrae, but some are isolated and are relatively well preserved (
<figureCitation id="CF8C89C7FF839E1DFC26FBA43440541B" box="[930,1037,1036,1060]" captionStart="Figure 9" captionStartId="10.[129,194,1629,1653]" captionTargetBox="[129,1473,145,1601]" captionTargetId="figure-150@10.[129,1473,145,1601]" captionTargetPageId="10" captionText="Figure 9. Postcranial elements of Ymboirana acrux gen. et sp.nov.. Trunk vertebra in dorsal (A), ventral (B), and oblique less lateral (C) views. Ribs (D, E) in lateral view. See Appendix for abbreviations. Scale bar = 1 mm." pageId="11" pageNumber="12">Fig. 9D, E</figureCitation>
). Ŋey have a relatively straight outline along their entire length, lacking pronounced posterior flexion. Ŋe ribs are dorsoventrally broad anteriorly, but they gradually narrow distally. Ŋe tuberculum and capitulum have straight or more rounded tips, whereas the distal tips are pointed. Ŋe capit* ulum is very elongate anteriorly in the preserved ribs.
</paragraph>
</subSubSection>
</treatment>
</document>