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<document ENCODING="UTF-8" ID-DOI="https://doi.org/10.5962/bhl.title.542" ID-GBIF-Dataset="57ea6fdb-2533-44e5-b60b-736758a88d19" ID-Zenodo-Dep="3922206" ID-ZooBank="2C6327E1-5560-4DB4-B9CA-76A0FA03D975" checkinTime="1593218702860" checkinUser="admin" docAuthor="Linnaeus, Carolus" docDate="1758" docId="D973217B3CB9FBCF521896556F08032D" docLanguage="de" docName="Linnaeus1758Treatments.xml" docOrigin="Stockholm: Laurentius Salvius" docSource="https://archive.org/download/mobot31753000798865/mobot31753000798865.pdf" docTitle="Helix contorta Linnaeus, 1758, spec. nov." docType="treatment" docUuid="05F28933-94A8-4172-AE68-91E80F7A3706" docUuidSource="ZooBank" docVersion="13" lastPageNumber="770" masterDocId="3F896AFC21C459639D0D71D9EDB8875C" masterDocTitle="Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis" pageNumber="770" updateTime="1643600433100" updateUser="ExternalLinkService">
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<mods:title>Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis</mods:title>
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<mods:namePart>Linnaeus, Carolus</mods:namePart>
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<mods:location>
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</mods:location>
<mods:classification>book</mods:classification>
<mods:identifier type="ZooBank">2C6327E1-5560-4DB4-B9CA-76A0FA03D975</mods:identifier>
<mods:identifier type="DOI">https://doi.org/10.5962/bhl.title.542</mods:identifier>
<mods:identifier type="Zenodo-Dep">3922206</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3912520" ID-GBIF-Taxon="164807817" ID-Zenodo-Dep="3912520" LSID="urn:lsid:zoobank.org:act:05F28933-94A8-4172-AE68-91E80F7A3706" httpUri="http://treatment.plazi.org/id/D973217B3CB9FBCF521896556F08032D" lastPageNumber="770" pageNumber="770" sourcePageUrl="https://www.biodiversitylibrary.org/page/727681">
<subSubSection type="nomenclature">
<paragraph pageNumber="770">
<taxonomicName LSID="urn:lsid:zoobank.org:act:05F28933-94A8-4172-AE68-91E80F7A3706" authorityName="Linnaeus" authorityYear="1758" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" phylum="Bryozoa" rank="species" species="contorta" status="spec. nov." zbkClass="Vermes" zbkKingdom="Animalia" zbkOrder="Testacea">Helix contorta</taxonomicName>
[
<taxonomicNameLabel rank="species">spec. nov.</taxonomicNameLabel>
]
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="770">H. testa subumbilicata plana utrinque aequali, apertura lineari arcuata.</paragraph>
<paragraph pageNumber="770">
<emphasis italics="true">Fn. svec.</emphasis>
1309.
</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageNumber="770">
<emphasis italics="true">Habitat in</emphasis>
Europae
<emphasis italics="true">stagnis supra plantas.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.9.e68558" ID-PMC="PMC8408096" ID-Pensoft-Pub="1314-2828-9-e68558" ID-Pensoft-UUID="3F537781399057B984E912F3CACE85A8" ID-PubMed="34539199" ModsDocID="1314-2828-9-e68558" checkinTime="1629846852017" checkinUser="pensoft" docAuthor="Nastasi, Louis F. &amp; Deans, Andrew R." docDate="2021" docId="D9734D66951D5254AC756C53E1E950E1" docLanguage="en" docName="BiodivDatJour 9: e68558" docOrigin="Biodiversity Data Journal 9" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/BDJ.9.e68558" docTitle="Synergus beutenmulleri Lobato-Vila &amp; Pujade-Villar 2021" docType="treatment" docVersion="2" id="3F537781399057B984E912F3CACE85A8" lastPageNumber="68558" masterDocId="3F537781399057B984E912F3CACE85A8" masterDocTitle="Catalogue of Rose Gall, Herb Gall, and Inquiline Gall Wasps (Hymenoptera: Cynipidae) of the United States, Canada and Mexico" masterLastPageNumber="68558" masterPageNumber="68558" pageNumber="68558" updateTime="1668128778028" updateUser="ExternalLinkService">
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<treatment LSID="urn:lsid:plazi:treatment:D9734D66951D5254AC756C53E1E950E1" httpUri="http://treatment.plazi.org/id/D9734D66951D5254AC756C53E1E950E1" lastPageNumber="68558" pageId="0" pageNumber="68558">
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<taxonomicName LSID="D9734D66-951D-5254-AC75-6C53E1E950E1" authority="Lobato-Vila &amp; Pujade-Villar, 2021" authorityName="Lobato-Vila &amp; Pujade-Villar" authorityYear="2021" class="Insecta" family="Cynipidae" genus="Synergus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synergus beutenmulleri" order="Hymenoptera" pageId="0" pageNumber="68558" phylum="Arthropoda" rank="species" species="beutenmulleri">Synergus beutenmulleri Lobato-Vila &amp; Pujade-Villar, 2021</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="68558">Ecological interactions</paragraph>
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<subSubSection pageId="0" pageNumber="68558" type="feeds on">
<paragraph pageId="0" pageNumber="68558">Feeds on</paragraph>
<paragraph pageId="0" pageNumber="68558">
Inquiline of: galls of
<taxonomicName authorityName="Hartig" authorityYear="1840" class="Insecta" family="Cynipidae" genus="Andricus" kingdom="Animalia" lsidName="Andricus" order="Hymenoptera" pageId="0" pageNumber="68558" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="68558">Andricus</emphasis>
</taxonomicName>
sp.
<taxonomicName lsidName="Q. potosina" pageId="0" pageNumber="68558" rank="species" species="potosina">
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<subSubSection pageId="0" pageNumber="68558" type="distribution">
<paragraph pageId="0" pageNumber="68558">Distribution</paragraph>
<paragraph pageId="0" pageNumber="68558">Mexico: Aguascalientes</paragraph>
</subSubSection>
</treatment>
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<treatment id="D97387A89D31FF96FF4488D7FA71889D" ID-DOI="http://doi.org/10.5281/zenodo.6017415" ID-GBIF-Taxon="131876628" ID-Zenodo-Dep="6017415" LSID="urn:lsid:plazi:treatment:D97387A89D31FF96FF4488D7FA71889D" httpUri="http://treatment.plazi.org/id/D97387A89D31FF96FF4488D7FA71889D" lastPageNumber="256" pageId="1" pageNumber="256">
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Key to the Neotropical species of
<taxonomicName id="96DA4D3D9D31FF96FDE688D6FD258A22" ID-CoL="36BZ" authorityName="Wirth &amp; Grogan" authorityYear="1988" box="[565,712,1264,1290]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="63AEEAAC9D31FF96FDE688D6FD258A22" bold="true" box="[565,712,1264,1290]" italics="true" pageId="1" pageNumber="256">Austrohelea</emphasis>
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<keyLead id="EA2B958B9D31FF96FF448911FA768A65" box="[151,1435,1335,1357]" pageId="1" pageNumber="256">1 Males............................................................................................... 2</keyLead>
</paragraph>
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<keyLead id="EA2B958B9D31FF96FF448973FA768A42" box="[151,1435,1364,1386]" pageId="1" pageNumber="256">- Females............................................................................................. 4</keyLead>
</paragraph>
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<keyStep id="EA2E2E1B9D31FF96FF448957FA768932" pageId="1" pageNumber="256">
<paragraph id="516536BE9D31FF96FF448957FA718AEA" blockId="1.[151,1436,1335,1973]" pageId="1" pageNumber="256">
<keyLead id="EA2B958B9D31FF96FF448957FA718AEA" pageId="1" pageNumber="256">
2 Wing membrane mostly pale, lightly infuscated between costa-radius and radius-medius (
<figureCitation id="C9E12A3B9D31FF96FBED8957FB6B8AAF" box="[1086,1158,1393,1415]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="1" pageNumber="256">Fig. 23</figureCitation>
); hind tibia with 3 large subapical spines (
<figureCitation id="C9E12A3B9D31FF96FEBB89A9FE5D8A8D" box="[360,432,1423,1445]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="1" pageNumber="256">Fig. 22</figureCitation>
); aedeagus heavily sclerotized with broad, long basal arms with rounded apices (
<figureCitation id="C9E12A3B9D31FF96FB6789A9FAE98A8D" box="[1204,1284,1423,1445]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="1" pageNumber="256">Figs. 24</figureCitation>
,
<figureCitation id="C9E12A3B9D31FF96FADD89A9FAC48A8D" box="[1294,1321,1423,1445]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="1" pageNumber="256">34</figureCitation>
); basal apodemes of parameres very broad with shallow obtuse angle between anterior, posterior arms (
<figureCitation id="C9E12A3B9D31FF96FB86898AFB488AEA" box="[1109,1189,1452,1474]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="1" pageNumber="256">Figs. 24</figureCitation>
,
<figureCitation id="C9E12A3B9D31FF96FB7C898AFB258AEA" box="[1199,1224,1452,1474]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="1" pageNumber="256">33</figureCitation>
)...
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<emphasis id="63AEEAAC9D31FF96FAC5898BFAB28AEA" box="[1302,1375,1453,1474]" italics="true" pageId="1" pageNumber="256">spinosa</emphasis>
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<emphasis id="63AEEAAC9D31FF96FAB5898AFA718AEA" bold="true" box="[1382,1436,1452,1474]" pageId="1" pageNumber="256">n. sp.</emphasis>
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<keyLead id="EA2B958B9D31FF96FF4489ECFA768932" pageId="1" pageNumber="256">- Wing mostly or entirely infuscated; hind tibia without large subapical spines; aedeagus lightly or heavily sclerotized with relatively short basal arms with blunt apices; basal apodemes of parameres slender to moderately broad with deep acute angle between anterior, posterior arms.......................................................................... 3</keyLead>
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<paragraph id="516536BE9D31FF96FF448A07FA71895A" blockId="1.[151,1436,1335,1973]" pageId="1" pageNumber="256">
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3 Wing membrane uniformly lightly infuscated (
<figureCitation id="C9E12A3B9D31FF96FD7F8A07FD1A891F" box="[684,759,1569,1591]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="1" pageNumber="256">Fig. 14</figureCitation>
); aedeagus with very shallow basal arch, basal arms slender slightly curved, distal portion lightly sclerotized with rounded apex (
<figureCitation id="C9E12A3B9D31FF96FCF98A19FC97897D" box="[810,890,1599,1621]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="1" pageNumber="256">Figs. 15</figureCitation>
,
<figureCitation id="C9E12A3B9D31FF96FC568A19FC4D897D" box="[901,928,1599,1621]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="1" pageNumber="256">32</figureCitation>
); basal apodemes of parameres slender, anterior arm twice as long as posterior arm (
<figureCitation id="C9E12A3B9D31FF96FDDE8A7AFDB0895A" box="[525,605,1628,1650]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="1" pageNumber="256">Figs. 15</figureCitation>
,
<figureCitation id="C9E12A3B9D31FF96FDB48A7AFD92895A" box="[615,639,1628,1650]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="1" pageNumber="256">31</figureCitation>
)......................................................
<taxonomicName id="96DA4D3D9D31FF96FACE8A7BFA8D895A" ID-CoL="9HTKS" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[1309,1376,1629,1650]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D31FF96FACE8A7BFAC1895A" box="[1309,1324,1629,1650]" italics="true" pageId="1" pageNumber="256">A</emphasis>
.
<emphasis id="63AEEAAC9D31FF96FAEA8A7BFA8D895A" box="[1337,1376,1629,1650]" italics="true" pageId="1" pageNumber="256">sirii</emphasis>
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<emphasis id="63AEEAAC9D31FF96FAB58A7AFA71895A" bold="true" box="[1382,1436,1628,1650]" pageId="1" pageNumber="256">n. sp.</emphasis>
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<paragraph id="516536BE9D31FF96FF448A5CFA7189E2" blockId="1.[151,1436,1335,1973]" pageId="1" pageNumber="256">
<keyLead id="EA2B958B9D31FF96FF448A5CFA7189E2" pageId="1" pageNumber="256">
- Wing membrane mostly darkly infuscated (
<figureCitation id="C9E12A3B9D31FF96FDAD8A5FFD5A89A7" box="[638,695,1657,1679]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="1" pageNumber="256">Fig. 3</figureCitation>
); aedeagus with deep triangular basal arch, basal arms broad greatly recurved with blunt apices, distal portion heavily sclerotized (
<figureCitation id="C9E12A3B9D31FF96FD058AB1FCF78985" box="[726,794,1687,1709]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="1" pageNumber="256">Figs. 4</figureCitation>
,
<figureCitation id="C9E12A3B9D31FF96FCF78AB1FCD28985" box="[804,831,1687,1709]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="1" pageNumber="256">30</figureCitation>
); basal apodemes of parameres with broad posterior arm that is twice as long as anterior arm (
<figureCitation id="C9E12A3B9D31FF96FDD18A92FDAB89E2" box="[514,582,1716,1738]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="1" pageNumber="256">Figs. 4</figureCitation>
,
<figureCitation id="C9E12A3B9D31FF96FD838A92FD8489E2" box="[592,617,1716,1738]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="1" pageNumber="256">29</figureCitation>
)..........................................
<taxonomicName id="96DA4D3D9D31FF96FBA28A93FA7189E2" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Blanton" authorityYear="Wirth &amp; Blanton" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[1137,1436,1716,1738]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D31FF96FBA28A93FB6D89E2" box="[1137,1152,1717,1738]" italics="true" pageId="1" pageNumber="256">A</emphasis>
.
<emphasis id="63AEEAAC9D31FF96FB5F8A93FB0E89E2" box="[1164,1251,1717,1738]" italics="true" pageId="1" pageNumber="256">shannoni</emphasis>
(Wirth &amp; Blanton)
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<keyStep id="EA2E2E1B9D31FF96FF448AF7FA768875" pageId="1" pageNumber="256">
<paragraph id="516536BE9D31FF96FF448AF7FA71880A" blockId="1.[151,1436,1335,1973]" pageId="1" pageNumber="256">
<keyLead id="EA2B958B9D31FF96FF448AF7FA71880A" pageId="1" pageNumber="256">
4 Small species (wing length
<quantity id="96229B5B9D31FF96FE3A8AF7FDAF89CF" box="[489,578,1745,1767]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="256" unit="mm" value="1.16">1.16 mm</quantity>
); wing membrane darkly infuscated with dense, coarse macrotrichia (
<figureCitation id="C9E12A3B9D31FF96FB098AF7FACC89CF" box="[1242,1313,1745,1767]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="1" pageNumber="256">Fig. 18</figureCitation>
); palpal segment 3 with large apical sensory pit (
<figureCitation id="C9E12A3B9D31FF96FD908AC9FD67882D" box="[579,650,1775,1797]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="1" pageNumber="256">Fig. 17</figureCitation>
); spermathecae pyriform, unequal, with short, broad, truncate necks (
<figureCitation id="C9E12A3B9D31FF96FAF78AC9FA87882D" box="[1316,1386,1775,1797]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="1" pageNumber="256">Fig. 19</figureCitation>
).............................................................................................
<taxonomicName id="96DA4D3D9D31FF96FACD8B2BFA8D880A" ID-CoL="9HTKS" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[1310,1376,1805,1826]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D31FF96FACD8B2BFAC0880A" box="[1310,1325,1805,1826]" italics="true" pageId="1" pageNumber="256">A</emphasis>
.
<emphasis id="63AEEAAC9D31FF96FAEA8B2BFA8D880A" box="[1337,1376,1805,1826]" italics="true" pageId="1" pageNumber="256">sirii</emphasis>
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<emphasis id="63AEEAAC9D31FF96FAB58B2AFA71880A" bold="true" box="[1382,1436,1804,1826]" pageId="1" pageNumber="256">n. sp.</emphasis>
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<paragraph id="516536BE9D31FF96FF448B0CFA768875" blockId="1.[151,1436,1335,1973]" pageId="1" pageNumber="256">
<keyLead id="EA2B958B9D31FF96FF448B0CFA768875" pageId="1" pageNumber="256">
- Large species (wing length 1.57‾
<quantity id="96229B5B9D31FF96FDF18B0FFD918817" box="[546,636,1833,1855]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="256" unit="mm" value="1.85">1.85 mm</quantity>
); wing membrane darkly or lightly infuscated with sparse to moderately dense thin macrotrichia; palpal segment 3 without sensory pit; spermathecae ovoid, subequal.................................. 5
</keyLead>
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<keyStep id="EA2E2E1B9D31FF96FF448B42FA71889D" pageId="1" pageNumber="256">
<paragraph id="516536BE9D31FF96FF448B42FA718852" blockId="1.[151,1436,1335,1973]" box="[151,1436,1892,1914]" pageId="1" pageNumber="256">
<keyLead id="EA2B958B9D31FF96FF448B42FA718852" box="[151,1436,1892,1914]" pageId="1" pageNumber="256">
5 Hind femur greatly swollen; spermathecae with short broad necks (
<figureCitation id="C9E12A3B9D31FF96FCB68B42FC408852" box="[869,941,1892,1914]" captionStart="FIGURES 25 – 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 – 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 – 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="1" pageNumber="256">Fig. 28</figureCitation>
)...........................
<taxonomicName id="96DA4D3D9D31FF96FB2F8B43FAB28852" ID-CoL="9HWY2" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[1276,1375,1893,1914]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D31FF96FB2F8B43FAE68852" box="[1276,1291,1893,1914]" italics="true" pageId="1" pageNumber="256">A</emphasis>
.
<emphasis id="63AEEAAC9D31FF96FAC58B43FAB28852" box="[1302,1375,1893,1914]" italics="true" pageId="1" pageNumber="256">spinosa</emphasis>
</taxonomicName>
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<emphasis id="63AEEAAC9D31FF96FAB58B42FA718852" bold="true" box="[1382,1436,1892,1914]" pageId="1" pageNumber="256">n. sp.</emphasis>
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</keyLead>
</paragraph>
<paragraph id="516536BE9D31FF96FF448BA4FA71889D" blockId="1.[151,1436,1335,1973]" pageId="1" pageNumber="256">
<keyLead id="EA2B958B9D31FF96FF448BA4FA71889D" pageId="1" pageNumber="256">
- Hind femur slightly swollen (
<figureCitation id="C9E12A3B9D31FF96FE288BA7FDDB88BF" box="[507,566,1921,1943]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[201,1376,211,1917]" captionTargetId="figure@4.[179,1397,193,1949]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Austrohelea shannoni (Wirth &amp; Blanton). Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8 – 10 and spermathecae." httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="1" pageNumber="256">Fig. 6</figureCitation>
); spermathecae with moderately long slender necks (
<figureCitation id="C9E12A3B9D31FF96FBE08BA7FB9488BF" box="[1075,1145,1921,1943]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[201,1376,211,1917]" captionTargetId="figure@4.[179,1397,193,1949]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Austrohelea shannoni (Wirth &amp; Blanton). Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8 – 10 and spermathecae." httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="1" pageNumber="256">Fig. 12</figureCitation>
)...................................................................................................
<taxonomicName id="96DA4D3D9D31FF96FBA28BB9FA71889D" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Blanton" authorityYear="Wirth &amp; Blanton" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[1137,1436,1951,1973]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D31FF96FBA28BB9FB6D889C" box="[1137,1152,1951,1972]" italics="true" pageId="1" pageNumber="256">A</emphasis>
.
<emphasis id="63AEEAAC9D31FF96FB5F8BB9FB0E889C" box="[1164,1251,1951,1972]" italics="true" pageId="1" pageNumber="256">shannoni</emphasis>
(Wirth &amp; Blanton)
</taxonomicName>
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</paragraph>
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</subSubSection>
</treatment>
</document>

401
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<mods:title id="FF68FC0CAECFA547B5DEA5AD2F03AFD2">The Neotropical species of the predaceous midge genus Austrohelea Wirth &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="D324A1A251DF6379DD263537A3C87A59">Spinelli, Gustavo R.</mods:namePart>
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<mods:namePart id="6163C57D4A01A3CDBE81E6B4B0F7CA74">Grogan, Willliam L.</mods:namePart>
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<mods:title id="259B4353C12E7A9C6E7FEAC64A67B90B">Zootaxa</mods:title>
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<mods:part id="CF905134D96AC89F37261FBAF2307512">
<mods:date id="B383957DEC3E127CA9245C9CC4B19054">2017</mods:date>
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<mods:number id="9C9182EAA345B0A7625042082B5F1B06">4276</mods:number>
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<treatment id="D97387A89D32FF92FF448CB1FB028CF9" ID-DOI="http://doi.org/10.5281/zenodo.6017417" ID-GBIF-Taxon="131876631" ID-Zenodo-Dep="6017417" LSID="urn:lsid:plazi:treatment:D97387A89D32FF92FF448CB1FB028CF9" httpUri="http://treatment.plazi.org/id/D97387A89D32FF92FF448CB1FB028CF9" lastPageId="5" lastPageNumber="260" pageId="2" pageNumber="257">
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<paragraph id="516536BE9D32FF95FF448CB1FD798F9A" blockId="2.[151,660,151,211]" box="[151,660,151,178]" pageId="2" pageNumber="257">
<heading id="0A2D81D29D32FF95FF448CB1FD798F9A" bold="true" box="[151,660,151,178]" fontSize="11" level="1" pageId="2" pageNumber="257" reason="1">
<taxonomicName id="96DA4D3D9D32FF95FF448CB1FD798F9A" authority="(Wirth &amp; Blanton)" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[151,660,151,178]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D32FF95FF448CB1FD798F9A" bold="true" box="[151,660,151,178]" pageId="2" pageNumber="257">
<emphasis id="63AEEAAC9D32FF95FF448CB1FE4E8F99" bold="true" box="[151,419,151,177]" italics="true" pageId="2" pageNumber="257">Austrohelea shannoni</emphasis>
(Wirth &amp; Blanton)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="516536BE9D32FF95FF448C9CFE9A8FFB" blockId="2.[151,660,151,211]" box="[151,375,186,211]" pageId="2" pageNumber="257">
(
<figureCitation id="C9E12A3B9D32FF95FF4C8C9CFF048FFB" box="[159,233,186,211]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="2" pageNumber="257">Figs. 1</figureCitation>
̄12, 29̄30)
</paragraph>
</subSubSection>
<subSubSection id="19C065359D32FF95FF448D24FD458EF8" pageId="2" pageNumber="257" type="reference_group">
<paragraph id="516536BE9D32FF95FF448D24FE6A8E1F" blockId="2.[151,1436,258,464]" pageId="2" pageNumber="257">
<treatmentCitationGroup id="71CA11909D32FF95FF448D24FE6A8E1F" pageId="2" pageNumber="257">
<taxonomicName id="96DA4D3D9D32FF95FF448D24FCCF8E30" ID-CoL="8CWD5" authority="Wirth &amp; Blanton, 1972: 175" authorityName="Wirth &amp; Blanton" authorityPageNumber="175" authorityYear="1972" box="[151,802,258,280]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="shannoni" subGenus="Isthmohelea">
<emphasis id="63AEEAAC9D32FF95FF448D24FEE48E30" box="[151,265,258,280]" italics="true" pageId="2" pageNumber="257">Monohelea</emphasis>
(
<emphasis id="63AEEAAC9D32FF95FEC48D24FE7E8E30" box="[279,403,258,280]" italics="true" pageId="2" pageNumber="257">Isthmohelea</emphasis>
)
<emphasis id="63AEEAAC9D32FF95FE718D24FE128E30" box="[418,511,258,280]" italics="true" pageId="2" pageNumber="257">shannoni</emphasis>
<treatmentCitation id="D07B10AF9D32FF95FDD68D24FCCF8E30" author="Wirth" box="[517,802,258,280]" page="175" pageId="2" pageNumber="257" year="1972">
<bibRefCitation id="354B4B4F9D32FF95FDD68D24FD1F8E30" author="Wirth" box="[517,754,258,280]" pageId="2" pageNumber="257" refString="Wirth, W. W. &amp; Blanton, F. S. (1972) A new Patagonian biting midge of the genus Monohelea, Isthmohelea (Diptera: Ceratopogonidae). Florida Entomologist, 55, 173 - 176." type="journal article" year="1972">Wirth &amp; Blanton, 1972</bibRefCitation>
: 175
</treatmentCitation>
</taxonomicName>
(female;
<collectingCountry id="29CD762E9D32FF95FC508D24FC008E30" box="[899,1005,258,280]" name="Argentina" pageId="2" pageNumber="257">Argentina</collectingCountry>
);
<treatmentCitation id="D07B10AF9D32FF95FC2E8D24FB7A8E30" author="Wirth" box="[1021,1175,258,280]" page="40" pageId="2" pageNumber="257" year="1974">
<bibRefCitation id="354B4B4F9D32FF95FC2E8D24FB998E30" author="Wirth" box="[1021,1140,258,280]" pageId="2" pageNumber="257" refString="Wirth, W. W. (1974) Family Ceratopogonidae. In: Papavero, N. (Ed.), A Catalog of the Diptera of the Americas South of the United States, Fasc. 14, pp. 1 - 89." type="book chapter" year="1974">Wirth 1974</bibRefCitation>
: 40
</treatmentCitation>
(in New World catalogue south of the
<collectingCountry id="29CD762E9D32FF95FE978D07FE968E1F" box="[324,379,289,311]" name="United States of America" pageId="2" pageNumber="257">USA</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="516536BE9D32FF95FF448D66FD458EF8" blockId="2.[151,1436,258,464]" pageId="2" pageNumber="257">
<treatmentCitationGroup id="71CA11909D32FF95FF448D66FD458EF8" pageId="2" pageNumber="257">
<taxonomicName id="96DA4D3D9D32FF95FF448D66FD708E7E" authority=": Wirth &amp; Grogan 1988: 23" authorityName="Wirth &amp; Grogan" authorityPageNumber="23" authorityYear="1988" box="[151,669,320,342]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D32FF95FF448D66FE9A8E7E" box="[151,375,320,342]" italics="true" pageId="2" pageNumber="257">Austrohelea shannoni</emphasis>
:
<treatmentCitation id="D07B10AF9D32FF95FE5B8D66FD708E7E" author="Wirth" box="[392,669,320,342]" page="23" pageId="2" pageNumber="257" year="1988">
<bibRefCitation id="354B4B4F9D32FF95FE5B8D66FD9B8E7E" author="Wirth" box="[392,630,320,342]" pageId="2" pageNumber="257" refString="Wirth, W. W. &amp; Grogan, W. L. Jr. (1988) The predaceous midges of the World (Diptera: Ceratopogonidae; Tribe Ceratopogonini). In: Flora and Fauna Handbook 4. E. J. Brill, New York, pp. 1 - 160." type="journal article" year="1988">Wirth &amp; Grogan 1988</bibRefCitation>
: 23
</treatmentCitation>
</taxonomicName>
(combination; figs. female, male);
<treatmentCitation id="D07B10AF9D32FF95FBC58D66FAC08E7E" author="Spinelli" box="[1046,1325,320,342]" page="39" pageId="2" pageNumber="257" year="1993">
<bibRefCitation id="354B4B4F9D32FF95FBC58D66FAEA8E7E" author="Spinelli" box="[1046,1287,320,342]" pageId="2" pageNumber="257" refString="Spinelli, G. R. &amp; Wirth, W. W. (1993) Los Ceratopogonidae de la Argentina (Insecta: Diptera). In: Castellanos, Z. A. de (Director), Fauna de agua dulce de la Republica Argentina, 38 (Fasc. 3), 1 - 124." type="journal article" year="1993">Spinelli &amp; Wirth 1993</bibRefCitation>
: 39
</treatmentCitation>
(in list of
<taxonomicName id="96DA4D3D9D32FF95FF148D78FE998E5C" ID-CoL="7WQ" authorityName="Newman" authorityYear="1834" box="[199,372,350,372]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
of
<collectingCountry id="29CD762E9D32FF95FE768D78FDE28E5C" box="[421,527,350,372]" name="Argentina" pageId="2" pageNumber="257">Argentina</collectingCountry>
);
<treatmentCitation id="D07B10AF9D32FF95FDFB8D78FCBF8E5C" author="Borkent" box="[552,850,350,372]" page="91" pageId="2" pageNumber="257" year="1997">
<bibRefCitation id="354B4B4F9D32FF95FDFB8D78FCCA8E5C" author="Borkent" box="[552,807,350,372]" pageId="2" pageNumber="257" refString="Borkent, A. &amp; Wirth, W. W. (1997) World species of biting midges (Diptera: Ceratopogonidae). Bulletin of the American Museum of Natural History, 233, 1 - 257." type="journal article" year="1997">Borkent &amp; Wirth 1997</bibRefCitation>
: 91
</treatmentCitation>
(in World catalogue);
<treatmentCitation id="D07B10AF9D32FF95FB858D78FACC8E5C" author="Spinelli" box="[1110,1313,350,372]" page="325" pageId="2" pageNumber="257" year="1998">
<bibRefCitation id="354B4B4F9D32FF95FB858D78FB048E5C" author="Spinelli" box="[1110,1257,350,372]" pageId="2" pageNumber="257" refString="Spinelli, G. R. (1998) Ceratopogonidae. Capitulo 30. In: Morrone, J. J. &amp; Coscaron, S. (Directors), Biodiversidad de Artropodos argentinos. Una perspectiva biotaxonomica. Ediciones Sur, La Plata, pp. 314 - 326." type="book chapter" year="1998">Spinelli 1998</bibRefCitation>
: 325
</treatmentCitation>
(in list of
<taxonomicName id="96DA4D3D9D32FF95FF148D5BFE998EBB" ID-CoL="7WQ" authorityName="Newman" authorityYear="1834" box="[199,372,381,403]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
of
<collectingCountry id="29CD762E9D32FF95FE488D5BFDE88EBB" box="[411,517,381,403]" name="Argentina" pageId="2" pageNumber="257">Argentina</collectingCountry>
);
<treatmentCitation id="D07B10AF9D32FF95FDCB8D5BFCAE8EBB" author="Borkent" box="[536,835,381,403]" page="44" pageId="2" pageNumber="257" year="2000">
<bibRefCitation id="354B4B4F9D32FF95FDCB8D5BFCF38EBB" author="Borkent" box="[536,798,381,403]" pageId="2" pageNumber="257" refString="Borkent, A. &amp; Spinelli, G. R. (2000) Catalog of the New World biting midges south of the United States of America (Diptera: Ceratopogonidae). Contributions on Entomology, International, 4, 1 - 107." type="journal article" year="2000">Borkent &amp; Spinelli 2000</bibRefCitation>
: 44
</treatmentCitation>
(in New World catalogue south of the
<collectingCountry id="29CD762E9D32FF95FB318D5BFAF48EBB" box="[1250,1305,381,403]" name="United States of America" pageId="2" pageNumber="257">USA</collectingCountry>
);
<treatmentCitation id="D07B10AF9D32FF95FAFE8D5BFE918E9A" author="Borkent" page="77" pageId="2" pageNumber="257" year="2007">
<bibRefCitation id="354B4B4F9D32FF95FAFE8D5BFEB88E9A" author="Borkent" pageId="2" pageNumber="257" refString="Borkent, A. &amp; Spinelli, G. R. (2007) Neotropical Ceratopogonidae (Diptera: Insecta). In: Adis, J., Arias, R., Rueda-Delgado, G. &amp; Wantzen, K. M. (Eds.), Aquatic Biodiversity in Latin America (ABLA). Fol. 4. Pensoft, Sofia-Moscow, pp. 1 - 198." type="journal article" year="2007">Borkent &amp; Spinelli 2007</bibRefCitation>
: 77
</treatmentCitation>
(in Neotropical catalogue);
<treatmentCitation id="D07B10AF9D32FF95FD7B8DBAFC368E9A" author="Spinelli" box="[680,987,412,434]" page="205" pageId="2" pageNumber="257" year="2009">
<bibRefCitation id="354B4B4F9D32FF95FD7B8DBAFC458E9A" author="Spinelli" box="[680,936,412,434]" pageId="2" pageNumber="257" refString="Spinelli, G. R. &amp; Marino, P. I. (2009) Estado actual del conocimiento de la familia Ceratopogonidae en la Patagonia (Diptera: Nematocera). Revista de la Sociedad Entomologica Argentina, 68, 201 - 208." type="journal article" year="2009">Spinelli &amp; Marino 2009</bibRefCitation>
: 205
</treatmentCitation>
(in list of
<taxonomicName id="96DA4D3D9D32FF95FB808DBAFAED8E9A" ID-CoL="7WQ" authorityName="Newman" authorityYear="1834" box="[1107,1280,412,434]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
of Patagonia);
<treatmentCitation id="D07B10AF9D32FF95FF148D9CFE698EF8" author="Borkent" box="[199,388,442,464]" page="115" pageId="2" pageNumber="257" year="2016">
<bibRefCitation id="354B4B4F9D32FF95FF148D9CFEB88EF8" author="Borkent" box="[199,341,442,464]" pageId="2" pageNumber="257" refString="Borkent, A. (2016) World species of biting midges (Diptera: Ceratopogonidae). Available from: http: // wwx. inhs. illinois. edu / files / 1114 / 2384 / 5200 / CeratopogonidaeCatalog. pdf (accessed 16 February 2016)" type="journal article" year="2016">Borkent 2016</bibRefCitation>
: 115
</treatmentCitation>
(in online World catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="19C065359D32FF95FF448DDBFD568DE2" pageId="2" pageNumber="257" type="diagnosis">
<paragraph id="516536BE9D32FF95FF448DDBFD568DE2" blockId="2.[151,1437,509,2013]" pageId="2" pageNumber="257">
<emphasis id="63AEEAAC9D32FF95FF448DDBFEFC8D3E" bold="true" box="[151,273,509,534]" pageId="2" pageNumber="257">Diagnosis.</emphasis>
A large, dark brown Neotropical species. Males lacking a sensory pit on palpal segment 3; legs slender; posterior margin of tergite 9 slightly notched, with short, stout widely spaced apicolateral processes and triangular cerci; gonocoxite tapering distally with basomesal tubercle; gonostylus as long as gonocoxite; parameres with anterior arm of basal apodeme slender, posterior arm stout, longer, forming an acute angle between both arms; basal arch of aedeagus V-shaped. Females lacking a sensory pit on palpal segment 3, with slender legs and unequal, ovoid spermathecae with moderately long necks.
</paragraph>
</subSubSection>
<subSubSection id="19C065359D32FF92FF148EF3FB918EB8" lastPageId="5" lastPageNumber="260" pageId="2" pageNumber="257" type="description">
<paragraph id="516536BE9D32FF95FF148EF3FDC08ACD" blockId="2.[151,1437,509,2013]" pageId="2" pageNumber="257">
<emphasis id="63AEEAAC9D32FF95FF148EF3FEE68DC6" bold="true" box="[199,267,725,750]" pageId="2" pageNumber="257">Male.</emphasis>
Head (
<figureCitation id="C9E12A3B9D32FF95FEA18EF3FE508DC6" box="[370,445,725,750]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="2" pageNumber="257">Fig. 1</figureCitation>
) dark brown. Eyes narrowly contiguous on extreme lower portions; with fine interommatidial spicules. Antennal flagellum (
<figureCitation id="C9E12A3B9D32FF95FD728EDFFD098C3A" box="[673,740,761,786]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="2" pageNumber="257">Fig. 2</figureCitation>
) with flagellomeres 2̄8 barrel-shaped, 9̄10 vasiform, 11̄13 elongate, 11 longest; plume well developed, moderately dense, extending to apex of flagellomere 12; flagellomere 12 with single subapical sensillum coeloconicum; antennal ratio 0.73̄0.75 (0.74, n = 4). Clypeus with 3̄5 pairs of setae. Palpus moderately long; segment 5 extending well beyond apex of proboscis; segment 3 elongate with a few capitate sensilla on distal 1/3 of mesal surface; palpal ratio 3.00̄3.39 (3.20, n = 5). Thorax dark brown. Legs similar to those of female except for the tarsal claws, which are shorter, equal, without basal inner teeth, apices bifid; hind tibial ratio 11.00̄11.70 (11.53, n = 5); hind tarsal ratio 1.73̄1.82 (1.79, n = 5). Wing (
<figureCitation id="C9E12A3B9D32FF95FB078FF7FAFB8CC2" box="[1236,1302,977,1002]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="2" pageNumber="257">Fig. 3</figureCitation>
) membrane infuscated brown, darkest on proximal section from base to apex of costa, radius and beyond r-m crossvein, slightly lighter on distal sections of cells r3, m1 and m2; veins dark brown; wing length 1.66̄1.98 (1.79, n = 4) mm; width 0.54̄0.59 (0.56, n = 4) mm; costal ratio 0.62̄0.64 (0.63, n = 4). Abdomen brown. Genitalia (
<figureCitation id="C9E12A3B9D32FF95FB308866FAD88B70" box="[1251,1333,1088,1113]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,264,1997,2020]" captionTargetBox="[229,1390,246,1920]" captionTargetId="figure@3.[168,1451,185,1999]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Austrohelea shannoni (Wirth &amp; Blanton). Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia." httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="2" pageNumber="257">Figs. 4</figureCitation>
,
<figureCitation id="C9E12A3B9D32FF95FA978866FA898B70" box="[1348,1380,1088,1112]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="2" pageNumber="257">29</figureCitation>
̄30). Tergite 9 tapering distally, posterior margin with shallow median notch; apicolateral processes short, stout widely spaced with a single small apical seta; cercus slender, triangular, folded ventrally; sternite 9 moderately long, twice as broad as long with shallow, concave posteromedian excavation. Gonocoxite stout, tapering distally, with mesobasal tubercle, surface with sparse coarse elongate setae; gonostylus as long as gonocoxite, nearly straight, distal half broad, tip pointed. Parameres (
<figureCitation id="C9E12A3B9D32FF95FDB888D2FD508A24" box="[619,701,1268,1293]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="2" pageNumber="257">Fig. 29</figureCitation>
) separate, moderately sclerotized, somewhat boomerang-shaped; basal apodemes with slender anterior arm, posterior arm much longer, broader, apex truncate; an acute angle formed between anterior and posterior arms; distal portions elongate, slightly curved, apices divergent, slightly pointed. Aedeagus (
<figureCitation id="C9E12A3B9D32FF95FEAC8946FE388A50" box="[383,469,1376,1401]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="2" pageNumber="257">Fig. 30</figureCitation>
) Y-shaped, slightly broader than long; basal arch V-shaped, extending 1/3 of total aedeagal length; basal arms heavily sclerotized, recurved 120˚, apices truncate; main body triangular, heavily sclerotized, tapering abruptly, apex sharply pointed, with a moderately long, lightly sclerotized distal process with short, slender ventrally recurved tip.
</paragraph>
<paragraph id="516536BE9D32FF92FF1489D6FB918EB8" blockId="2.[151,1437,509,2013]" lastBlockId="5.[151,1437,151,977]" lastPageId="5" lastPageNumber="260" pageId="2" pageNumber="257">
<emphasis id="63AEEAAC9D32FF95FF1489D6FEC88921" bold="true" box="[199,293,1520,1545]" pageId="2" pageNumber="257">Female.</emphasis>
Head (
<figureCitation id="C9E12A3B9D32FF95FE5389D6FE2A8921" box="[384,455,1520,1545]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[201,1376,211,1917]" captionTargetId="figure@4.[179,1397,193,1949]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Austrohelea shannoni (Wirth &amp; Blanton). Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8 – 10 and spermathecae." httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="2" pageNumber="257">Fig. 5</figureCitation>
) dark brown. Eyes barely contiguous, with fine interommatidial spicules. Antennal pedicel very dark brown, flagellum dark brown; flagellomeres 2̄8 short, slightly vasiform, 9̄13 elongate cylindrical, 13 longest; flagellomeres 8, 12 with single subapical sensillum coeloconicum; antennal ratio 1.23̄1.42 (1.32, n = 5). Clypeus with 3̄5 pairs of setae. Palpus brown; segment 3 slightly swollen, with a few capitate sensilla on distal 1/3 of mesal surface; palpal ratio 2.53̄3.12 (2.89, n = 5). Mandible with 8 large teeth (n =7). Thorax (
<figureCitation id="C9E12A3B9D32FF95FF2B8A82FEA58995" box="[248,328,1700,1725]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[201,1376,211,1917]" captionTargetId="figure@4.[179,1397,193,1949]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Austrohelea shannoni (Wirth &amp; Blanton). Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8 – 10 and spermathecae." httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="2" pageNumber="257">Figs. 6</figureCitation>
̄7). Scutum, post-scutellum dark brown, humeral areas and scutellum slightly paler (more evident in specimens cleared with KOH); scutellum with 6̄7 large, 10̄12 smaller setae. Legs dark brown; fore, mid femora, tibiae slender; hind femur, tibia more massive with more numerous, long stout setae, especially on tibia; hind tibial ratio 8.10̄9.05 (8.57, n = 6); hind tibial comb with 5 spines; tarsi (
<figureCitation id="C9E12A3B9D32FF95FBDD8B36FB6F8800" box="[1038,1154,1808,1833]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[201,1376,211,1917]" captionTargetId="figure@4.[179,1397,193,1949]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Austrohelea shannoni (Wirth &amp; Blanton). Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8 – 10 and spermathecae." httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="2" pageNumber="257">Figs. 8-10</figureCitation>
) dark brown, tarsomeres 1̄2 slightly paler in some specimens; tarsomeres 1 with basal, apical ventral spines and 1̄5 similar spines along ventral surface on fore, mid legs; tarsomeres 2̄3 of fore, mid legs with apical spine; hind tarsomeres 1̄2 with two rows of ventral palisade setae; hind tarsal ratio 1.85̄1.95 (1.90, n = 6); tarsomere 4 of fore, mid legs short, cordiform, tarsomere 4 of hind leg elongate, cylindrical; claws small, equal-size on fore, mid legs, hind leg with single long claw, slightly longer than tarsomere 5 with a stout basal tooth about 1/3 length of claw. Wing (
<figureCitation id="C9E12A3B9D32FF95FAED8BE2FA6088F4" box="[1342,1421,1988,2013]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[201,1376,211,1917]" captionTargetId="figure@4.[179,1397,193,1949]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Austrohelea shannoni (Wirth &amp; Blanton). Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8 – 10 and spermathecae." httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="2" pageNumber="257">Fig. 11</figureCitation>
) membrane infuscated, darkest on distal 1/3, along veins and around radial cells; two well developed radial cells, 2nd broader, twice as long as 1st; macrotrichia most numerous on distal ½ of cell r3, less numerous in m1, m2, cua1 except near margin, sparse in anal cell; wing length 1.57̄1.85 (1.71, n = 6) mm, width 0.63̄0.75 (0.68, n = 6) mm; costal ratio 0.65̄0.67 (0.66, n = 7). Halter pale. Abdomen dark brown (
<figureCitation id="C9E12A3B9D35FF92FCA78D2AFC2A8E0D" box="[884,967,268,293]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[201,1376,211,1917]" captionTargetId="figure@4.[179,1397,193,1949]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Austrohelea shannoni (Wirth &amp; Blanton). Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8 – 10 and spermathecae." httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="5" pageNumber="260">Fig. 12</figureCitation>
). Sternite 9 with V-shaped posteromedian excavation; sternite 8 divided mesally, each half tapered; sternite 10 with two pairs of setae. Two ovoid, subequal spermathecae with moderately long, slender necks, measuring 60̄75 (65, n = 4) µm by 37̄42 (40, n = 4) µm, and 50̄68 (57, n = 4) µm by 33̄39 (37, n = 4) µm; a rudimentary third spermatheca present.
</paragraph>
</subSubSection>
<caption id="05A566369D33FF94FF448BEBFAD888CC" box="[151,1333,1997,2020]" httpUri="https://zenodo.org/record/806117/files/figure.png" pageId="3" pageNumber="258" targetBox="[229,1390,246,1920]" targetPageId="3">
<paragraph id="516536BE9D33FF94FF448BEBFAD888CC" blockId="3.[151,1333,1997,2020]" box="[151,1333,1997,2020]" pageId="3" pageNumber="258">
<emphasis id="63AEEAAC9D33FF94FF448BEBFED788CB" bold="true" box="[151,314,1997,2020]" pageId="3" pageNumber="258">FIGURES 1–4.</emphasis>
<taxonomicName id="96DA4D3D9D33FF94FE938BE8FD0F88CC" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[320,738,1998,2020]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D33FF94FE938BE8FDF088CC" box="[320,541,1998,2020]" italics="true" pageId="3" pageNumber="258">Austrohelea shannoni</emphasis>
(Wirth &amp; Blanton)
</taxonomicName>
. Male. 1. head. 2. antennal flagellum. 3. wing. 4. genitalia.
</paragraph>
</caption>
<caption id="05A566369D34FF93FF448B9CFC3988C7" httpUri="https://zenodo.org/record/806119/files/figure.png" pageId="4" pageNumber="259" targetBox="[201,1376,211,1917]" targetPageId="4">
<paragraph id="516536BE9D34FF93FF448B9CFC3988C7" blockId="4.[151,1435,1978,2031]" pageId="4" pageNumber="259">
<emphasis id="63AEEAAC9D34FF93FF448B9CFEAB88F8" bold="true" box="[151,326,1978,2001]" pageId="4" pageNumber="259">FIGURES 5–12.</emphasis>
<taxonomicName id="96DA4D3D9D34FF93FE9E8B9DFD0388F9" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[333,750,1979,2001]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D34FF93FE9E8B9DFDC788F9" box="[333,554,1979,2001]" italics="true" pageId="4" pageNumber="259">Austrohelea shannoni</emphasis>
(Wirth &amp; Blanton)
</taxonomicName>
. Female. 5. head. 6. thorax and legs. 7. thorax. 8. fore tarsus. 9. mid tarsus. 10. hind tarsus. 11. wing. 12. abdominal segments 8–10 and spermathecae.
</paragraph>
</caption>
<subSubSection id="19C065359D35FF92FF148DBAFF118D41" pageId="5" pageNumber="260" type="materials_examined">
<paragraph id="516536BE9D35FF92FF148DBAFF118D41" blockId="5.[151,1437,151,977]" pageId="5" pageNumber="260">
<materialsCitation id="E1B23CE39D35FF92FF148DBAFE3D8EF1" ID-GBIF-Occurrence="1563642362" collectingDate="1984-11-28" collectionCode="MLPA" collectorName="G. Spinelli" country="Argentina" location="Mascardi" municipality="Material" pageId="5" pageNumber="260" specimenCount="1" specimenCount-female="1" stateProvince="Rio Negro">
<emphasis id="63AEEAAC9D35FF92FF148DBAFE598E9D" bold="true" box="[199,436,412,437]" pageId="5" pageNumber="260">
<collectingMunicipality id="B101ACC49D35FF92FF148DBAFEC28E9D" box="[199,303,412,437]" pageId="5" pageNumber="260">Material</collectingMunicipality>
examined.
</emphasis>
<collectingCountry id="29CD762E9D35FF92FE138DBBFD8B8E9D" box="[448,614,413,437]" name="Argentina" pageId="5" pageNumber="260">ARGENTINA</collectingCountry>
,
<collectingRegion id="931EF85C9D35FF92FDA58DBBFD1E8E9D" box="[630,755,413,437]" country="Argentina" name="Rio Negro" pageId="5" pageNumber="260">Río Negro</collectingRegion>
,
<quantity id="96229B5B9D35FF92FCD18DBAFCAD8E9D" box="[770,832,412,437]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="260" unit="km" value="6.0">6 km</quantity>
oeste, extremo sur lago
<location id="540560659D35FF92FBBB8DBAFB3E8E9D" LSID="urn:lsid:plazi:treatment:D97387A89D32FF92FF448CB1FB028CF9:540560659D35FF92FBBB8DBAFB3E8E9D" box="[1128,1235,412,437]" country="Argentina" municipality="Material" name="Mascardi" pageId="5" pageNumber="260" stateProvince="Rio Negro">Mascardi</location>
,
<collectingDate id="3520E9969D35FF92FB318DBBFA988E9D" box="[1250,1397,413,437]" pageId="5" pageNumber="260" value="1984-11-28">
<date id="2564107E9D35FF92FB318DBBFA988E9D" box="[1250,1397,413,437]" pageId="5" pageNumber="260" value="1984-11-28">28-XI- 1984</date>
</collectingDate>
,
<collectorName id="FC2F53689D35FF92FA578DBBFF1C8EF1" pageId="5" pageNumber="260">G. Spinelli</collectorName>
,
<specimenCount id="47DCFD379D35FF92FF2C8DE7FE898EF1" box="[255,356,448,473]" pageId="5" pageNumber="260" type="female">1 female</specimenCount>
(
<collectionCode id="37CBAE7B9D35FF92FEA58DE7FE258EF1" box="[374,456,449,473]" pageId="5" pageNumber="260">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1B23CE39D35FF92FE0D8DE6FAF28EF1" ID-GBIF-Occurrence="1563642359" box="[478,1311,448,473]" collectingDate="1984-11-29" collectionCode="MLPA" collectorName="G. Spinelli" country="Argentina" location="Llao-Llao" municipality="Escondido" pageId="5" pageNumber="260" specimenCount="1" specimenCount-female="1" stateProvince="Rio Negro">
lago
<collectingMunicipality id="B101ACC49D35FF92FDCA8DE6FD628EF1" box="[537,655,448,473]" pageId="5" pageNumber="260">Escondido</collectingMunicipality>
(
<location id="540560659D35FF92FD728DE6FCFA8EF1" LSID="urn:lsid:plazi:treatment:D97387A89D32FF92FF448CB1FB028CF9:540560659D35FF92FD728DE6FCFA8EF1" box="[673,791,448,473]" country="Argentina" municipality="Escondido" name="Llao-Llao" pageId="5" pageNumber="260" stateProvince="Rio Negro">Llao-Llao</location>
),
<collectingDate id="3520E9969D35FF92FCFE8DE7FC588EF1" box="[813,949,449,473]" pageId="5" pageNumber="260" value="1984-11-29">
<date id="2564107E9D35FF92FCFE8DE7FC588EF1" box="[813,949,449,473]" pageId="5" pageNumber="260" value="1984-11-29">29-XI-1984</date>
</collectingDate>
,
<collectorName id="FC2F53689D35FF92FC108DE7FBD28EF1" box="[963,1087,448,473]" pageId="5" pageNumber="260">G. Spinelli</collectorName>
,
<specimenCount id="47DCFD379D35FF92FB9D8DE7FB5F8EF1" box="[1102,1202,448,473]" pageId="5" pageNumber="260" type="female">1 female</specimenCount>
(
<collectionCode id="37CBAE7B9D35FF92FB168DE7FAFA8EF1" box="[1221,1303,449,473]" pageId="5" pageNumber="260">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E1B23CE39D35FF92FAFF8DE6FBF58ED5" ID-GBIF-Occurrence="1563642361" collectingDate="1993-03-01" collectionCode="MLPA" collectorName="G. Spinelli" country="Argentina" location="Lapataia" pageId="5" pageNumber="260" specimenCount="2" specimenCount-female="2" stateProvince="Tierra del Fuego">
<collectingRegion id="931EF85C9D35FF92FAFF8DE6FF0D8ED5" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="5" pageNumber="260">Tierra del Fuego</collectingRegion>
, camino a
<location id="540560659D35FF92FE8F8DC2FE518ED5" LSID="urn:lsid:plazi:treatment:D97387A89D32FF92FF448CB1FB028CF9:540560659D35FF92FE8F8DC2FE518ED5" box="[348,444,484,509]" country="Argentina" name="Lapataia" pageId="5" pageNumber="260" stateProvince="Tierra del Fuego">Lapataia</location>
(turbera),
<collectingDate id="3520E9969D35FF92FDE58DC3FD418ED5" box="[566,684,485,509]" pageId="5" pageNumber="260" value="1993-03-01">
<date id="2564107E9D35FF92FDE58DC3FD418ED5" box="[566,684,485,509]" pageId="5" pageNumber="260" value="1993-03-01">1-III-1993</date>
</collectingDate>
,
<collectorName id="FC2F53689D35FF92FD698DC3FCD98ED5" box="[698,820,484,509]" pageId="5" pageNumber="260">G. Spinelli</collectorName>
,
<specimenCount id="47DCFD379D35FF92FC938DC3FC438ED5" box="[832,942,484,509]" pageId="5" pageNumber="260" type="female">2 females</specimenCount>
(
<collectionCode id="37CBAE7B9D35FF92FC6D8DC3FBFD8ED5" box="[958,1040,485,509]" pageId="5" pageNumber="260">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1B23CE39D35FF92FBF78DC3FD818D09" ID-GBIF-Occurrence="1563642357" collectingDate="1993-03-02" collectionCode="MLPA" collectorName="G. Spinelli" country="Argentina" elevation="170" location="rio Tristen" pageId="5" pageNumber="260" specimenCount="1" specimenCount-female="1" stateProvince="Tierra del Fuego">
<quantity id="96229B5B9D35FF92FBF78DC3FBB38ED5" box="[1060,1118,484,509]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="260" unit="km" value="1.0">1 km</quantity>
norte
<location id="540560659D35FF92FB7B8DC3FACC8ED5" LSID="urn:lsid:plazi:treatment:D97387A89D32FF92FF448CB1FB028CF9:540560659D35FF92FB7B8DC3FACC8ED5" box="[1192,1313,484,509]" country="Argentina" name="rio Tristen" pageId="5" pageNumber="260" stateProvince="Tierra del Fuego">río Tristen</location>
,
<elevation id="DAF7D18D9D35FF92FAFE8DC3FA958ED4" box="[1325,1400,485,509]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="260" unit="m" value="170.0">
<quantity id="96229B5B9D35FF92FAFE8DC3FA958ED4" box="[1325,1400,485,509]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="260" unit="m" value="170.0">170 m</quantity>
</elevation>
,
<collectingDate id="3520E9969D35FF92FA568DC3FF1A8D09" pageId="5" pageNumber="260" value="1993-03-02">
<date id="2564107E9D35FF92FA568DC3FF1A8D09" pageId="5" pageNumber="260" value="1993-03-02">2- III-1993</date>
</collectingDate>
,
<collectorName id="FC2F53689D35FF92FEDB8E2FFE6B8D09" box="[264,390,520,545]" pageId="5" pageNumber="260">G. Spinelli</collectorName>
,
<specimenCount id="47DCFD379D35FF92FE458E2FFE108D09" box="[406,509,520,545]" pageId="5" pageNumber="260" type="female">1 female</specimenCount>
(
<collectionCode id="37CBAE7B9D35FF92FDC28E2FFD8E8D09" box="[529,611,521,545]" pageId="5" pageNumber="260">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1B23CE39D35FF92FDAF8E2EFADE8D09" ID-GBIF-Occurrence="1563642366" box="[636,1331,520,545]" collectingDate="1993-03-03" collectionCode="MLPA" collectorName="Estancia La Vega &amp; G. Spinelli" country="Argentina" location="Estancia La Vega" pageId="5" pageNumber="260" specimenCount="1" specimenCount-male="1" stateProvince="Tierra del Fuego">
<collectorName id="FC2F53689D35FF92FDAF8E2EFCA18D09" box="[636,844,520,545]" pageId="5" pageNumber="260">
<location id="540560659D35FF92FDAF8E2EFCA18D09" LSID="urn:lsid:plazi:treatment:D97387A89D32FF92FF448CB1FB028CF9:540560659D35FF92FDAF8E2EFCA18D09" box="[636,844,520,545]" country="Argentina" name="Estancia La Vega" pageId="5" pageNumber="260" stateProvince="Tierra del Fuego">Estancia La Vega</location>
</collectorName>
,
<collectingDate id="3520E9969D35FF92FC8E8E2FFC3E8D09" box="[861,979,521,545]" pageId="5" pageNumber="260" value="1993-03-03">
<date id="2564107E9D35FF92FC8E8E2FFC3E8D09" box="[861,979,521,545]" pageId="5" pageNumber="260" value="1993-03-03">3-III-1993</date>
</collectingDate>
,
<collectorName id="FC2F53689D35FF92FC368E2FFB8E8D09" box="[997,1123,520,545]" pageId="5" pageNumber="260">G. Spinelli</collectorName>
,
<specimenCount id="47DCFD379D35FF92FBA78E2FFB298D09" box="[1140,1220,520,545]" pageId="5" pageNumber="260" type="male">1 male</specimenCount>
(
<collectionCode id="37CBAE7B9D35FF92FB0A8E2FFAC68D09" box="[1241,1323,521,545]" pageId="5" pageNumber="260">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E1B23CE39D35FF92FA908E2FFF148D41" ID-GBIF-Occurrence="1563642358" collectingDate="1980-02-25" collectionCode="MLPA" collectorName="D. Lanfranco" country="Chile" location="Bahia Scourfield" municipality="Isla Wollaston" pageId="5" pageNumber="260" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Magallanes">
<collectingCountry id="29CD762E9D35FF92FA908E2FFA758D09" box="[1347,1432,521,545]" name="Chile" pageId="5" pageNumber="260">CHILE</collectingCountry>
,
<collectingRegion id="931EF85C9D35FF92FF448E0AFEF68D6D" box="[151,283,556,581]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="5" pageNumber="260">Magallanes</collectingRegion>
,
<collectingMunicipality id="B101ACC49D35FF92FEFF8E0AFE3B8D6D" box="[300,470,556,581]" pageId="5" pageNumber="260">Isla Wollaston</collectingMunicipality>
,
<location id="540560659D35FF92FE358E0AFD418D6D" LSID="urn:lsid:plazi:treatment:D97387A89D32FF92FF448CB1FB028CF9:540560659D35FF92FE358E0AFD418D6D" box="[486,684,556,581]" country="Chile" municipality="Isla Wollaston" name="Bahia Scourfield" pageId="5" pageNumber="260" stateProvince="Magallanes">Bahia Scourfield</location>
, 17/
<collectingDate id="3520E9969D35FF92FD318E0AFCB38D6D" box="[738,862,556,581]" pageId="5" pageNumber="260" value="1980-02-25">
<date id="2564107E9D35FF92FD318E0AFCB38D6D" box="[738,862,556,581]" pageId="5" pageNumber="260" value="1980-02-25">25-II-1980</date>
</collectingDate>
,
<collectorName id="FC2F53689D35FF92FCBC8E0BFBE08D6D" box="[879,1037,556,581]" pageId="5" pageNumber="260">D. Lanfranco</collectorName>
,
<collectingMethod id="889B4EA99D35FF92FBCD8E0AFB5F8D6C" box="[1054,1202,556,581]" pageId="5" pageNumber="260">Malaise trap</collectingMethod>
,
<specimenCount id="47DCFD379D35FF92FB178E0BFACE8D6D" box="[1220,1315,556,581]" pageId="5" pageNumber="260" type="male">4 males</specimenCount>
,
<specimenCount id="47DCFD379D35FF92FAE68E0BFA718D6D" box="[1333,1436,556,581]" pageId="5" pageNumber="260" type="female">1 female</specimenCount>
(
<collectionCode id="37CBAE7B9D35FF92FF4D8E77FF1D8D41" box="[158,240,593,617]" pageId="5" pageNumber="260">MLPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19C065359D35FF92FF148E52FD338D99" pageId="5" pageNumber="260" type="distribution">
<paragraph id="516536BE9D35FF92FF148E52FD338D99" blockId="5.[151,1437,151,977]" pageId="5" pageNumber="260">
<emphasis id="63AEEAAC9D35FF92FF148E52FE8D8DA5" bold="true" box="[199,352,628,653]" pageId="5" pageNumber="260">Distribution.</emphasis>
This species is known from
<taxonomicName id="96DA4D3D9D35FF92FD7C8E52FCDD8DA4" authorityName="Blume" authorityYear="1850" box="[687,816,628,652]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="260" phylum="Tracheophyta" rank="genus">
<emphasis id="63AEEAAC9D35FF92FD7C8E52FCDD8DA4" box="[687,816,628,652]" italics="true" pageId="5" pageNumber="260">Nothofagus</emphasis>
</taxonomicName>
forests of
<collectingRegion id="931EF85C9D35FF92FC7D8E53FBCB8DA5" box="[942,1062,629,653]" country="Argentina" name="Rio Negro" pageId="5" pageNumber="260">Río Negro</collectingRegion>
and
<collectingRegion id="931EF85C9D35FF92FBB38E52FAF28DA5" box="[1120,1311,628,653]" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="5" pageNumber="260">Tierra del Fuego</collectingRegion>
provinces,
<collectingCountry id="29CD762E9D35FF92FF448EBEFEE48D99" box="[151,265,664,689]" name="Argentina" pageId="5" pageNumber="260">Argentina</collectingCountry>
, and on Wollaston Island,
<collectingCountry id="29CD762E9D35FF92FDE28EBEFD808D99" box="[561,621,664,689]" name="Chile" pageId="5" pageNumber="260">Chile</collectingCountry>
(
<figureCitation id="C9E12A3B9D35FF92FDAE8EBEFD3C8D99" box="[637,721,664,689]" captionStart="FIGURE 35" captionStartId="13.[151,250,1234,1256]" captionTargetBox="[267,1325,252,1118]" captionTargetId="figure@13.[205,1396,196,1194]" captionTargetPageId="13" captionText="FIGURE 35. Map of Argentine and Chilean Patagonia with distribution records of: Austrohelea shannoni (Wirth &amp; Blanton), green circles. Austrohelea sirii n. sp., yellow triangles. Austrohelea spinosa n. sp., red diamonds." httpUri="https://zenodo.org/record/806131/files/figure.png" pageId="5" pageNumber="260">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="19C065359D35FF92FF148E9AFB028CF9" pageId="5" pageNumber="260" type="discussion">
<paragraph id="516536BE9D35FF92FF148E9AFB028CF9" blockId="5.[151,1437,151,977]" pageId="5" pageNumber="260">
<emphasis id="63AEEAAC9D35FF92FF148E9AFEA18DFD" bold="true" box="[199,332,700,725]" pageId="5" pageNumber="260">Discussion.</emphasis>
This species was originally described by
<bibRefCitation id="354B4B4F9D35FF92FCFB8E9AFBD48DFD" author="Wirth" box="[808,1081,700,725]" pageId="5" pageNumber="260" refString="Wirth, W. W. &amp; Blanton, F. S. (1972) A new Patagonian biting midge of the genus Monohelea, Isthmohelea (Diptera: Ceratopogonidae). Florida Entomologist, 55, 173 - 176." type="journal article" year="1972">Wirth &amp; Blanton (1972)</bibRefCitation>
on the basis of a single female from Bariloche,
<collectingRegion id="931EF85C9D35FF92FE838EC7FDC28DD1" box="[336,559,736,761]" country="Argentina" name="Rio Negro" pageId="5" pageNumber="260">Río Negro province</collectingRegion>
in northwestern
<collectingCountry id="29CD762E9D35FF92FD3E8EC6FCB18DD1" box="[749,860,736,761]" name="Argentina" pageId="5" pageNumber="260">Argentine</collectingCountry>
Patagonia. In their book on the predaceous midges of the World,
<bibRefCitation id="354B4B4F9D35FF92FEE08F22FDD28C35" author="Wirth" box="[307,575,772,797]" pageId="5" pageNumber="260" refString="Wirth, W. W. &amp; Grogan, W. L. Jr. (1988) The predaceous midges of the World (Diptera: Ceratopogonidae; Tribe Ceratopogonini). In: Flora and Fauna Handbook 4. E. J. Brill, New York, pp. 1 - 160." type="journal article" year="1988">Wirth &amp; Grogan (1988)</bibRefCitation>
provided a generic diagnosis, the first description and data on males as well as illustrations of both sexes based on specimens from Wollaston Island, which is located at the extreme southeastern-most tip of
<collectingCountry id="29CD762E9D35FF92FE7F8F6AFE048C4D" box="[428,489,844,869]" name="Chile" pageId="5" pageNumber="260">Chile</collectingCountry>
south of the Beagle channel. We examined specimens from Bariloche and the Island of
<collectingRegion id="931EF85C9D35FF92FF668F56FE9C8CA1" box="[181,369,880,905]" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="5" pageNumber="260">Tierra del Fuego</collectingRegion>
,
<collectingCountry id="29CD762E9D35FF92FEAF8F56FE038CA1" box="[380,494,880,905]" name="Argentina" pageId="5" pageNumber="260">Argentina</collectingCountry>
, and also from Wollaston Island,
<collectingCountry id="29CD762E9D35FF92FCB08F56FC4F8CA1" box="[867,930,880,905]" name="Chile" pageId="5" pageNumber="260">Chile</collectingCountry>
, and therefore, we confirm the validity of the sex association assumed by
<bibRefCitation id="354B4B4F9D35FF92FE0D8FB2FD198C85" author="Wirth" box="[478,756,916,941]" pageId="5" pageNumber="260" refString="Wirth, W. W. &amp; Grogan, W. L. Jr. (1988) The predaceous midges of the World (Diptera: Ceratopogonidae; Tribe Ceratopogonini). In: Flora and Fauna Handbook 4. E. J. Brill, New York, pp. 1 - 160." type="journal article" year="1988">Wirth &amp; Grogan (1988)</bibRefCitation>
.
<taxonomicName id="96DA4D3D9D35FF92FCD28FB2FC158C84" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[769,1016,916,940]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D35FF92FCD28FB2FC158C84" box="[769,1016,916,940]" italics="true" pageId="5" pageNumber="260">Austrohelea shannoni</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="96DA4D3D9D35FF92FB0B8FB3FABD8C84" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[1240,1360,917,940]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D35FF92FB0B8FB3FB048C84" box="[1240,1257,917,940]" italics="true" pageId="5" pageNumber="260">A</emphasis>
.
<emphasis id="63AEEAAC9D35FF92FB298FB3FABD8C84" box="[1274,1360,917,940]" italics="true" pageId="5" pageNumber="260">spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D35FF92FA898FB2FA718C85" box="[1370,1436,916,941]" pageId="5" pageNumber="260" rank="species">
<emphasis id="63AEEAAC9D35FF92FA898FB2FA718C85" bold="true" box="[1370,1436,916,941]" pageId="5" pageNumber="260">n. sp.</emphasis>
</taxonomicNameLabel>
Characters to distinguish both species are in the key and the discussion section of the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>

287
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<mods:title id="16B1E37253974670D31DBF08BD29B38E">The Neotropical species of the predaceous midge genus Austrohelea Wirth &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
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<subSubSection id="19C065359D35FF92FF448805FE688B77" pageId="5" pageNumber="260" type="nomenclature">
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<heading id="0A2D81D29D35FF92FF448805FDEC8B16" bold="true" box="[151,513,1059,1086]" fontSize="11" level="1" pageId="5" pageNumber="260" reason="1">
<emphasis id="63AEEAAC9D35FF92FF448805FDEC8B16" bold="true" box="[151,513,1059,1086]" pageId="5" pageNumber="260">
<taxonomicName id="96DA4D3D9D35FF92FF448805FE8D8B15" ID-CoL="9HTKS" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[151,352,1059,1085]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D35FF92FF448805FE8D8B15" bold="true" box="[151,352,1059,1085]" italics="true" pageId="5" pageNumber="260">Austrohelea sirii</emphasis>
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<taxonomicNameLabel id="789D57D79D35FF92FEB48802FDEC8B16" box="[359,513,1060,1086]" pageId="5" pageNumber="260" rank="species">New Species</taxonomicNameLabel>
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<paragraph id="516536BE9D35FF92FF448860FE688B77" blockId="5.[151,513,1059,1119]" box="[151,389,1094,1119]" pageId="5" pageNumber="260">
(
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̄19, 31̄32)
</paragraph>
</subSubSection>
<subSubSection id="19C065359D35FF92FF4488A8FAB78AA8" pageId="5" pageNumber="260" type="diagnosis">
<paragraph id="516536BE9D35FF92FF4488A8FAB78AA8" blockId="5.[151,1437,1166,2028]" pageId="5" pageNumber="260">
<emphasis id="63AEEAAC9D35FF92FF4488A8FEFC8B8F" bold="true" box="[151,273,1166,1191]" pageId="5" pageNumber="260">Diagnosis.</emphasis>
A small (male wing length
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, female wing length
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), brown species. Both sexes with sensory pit on third palpal segment and slender legs. Males with posterior margin of tergite 9 deeply concave with elongate, cylindrical, slightly divergent apicolateral processes; gonocoxite short, stout, without basomesal tubercle; gonostylus longer than gonocoxite; parameres with moderately slender basal apodeme with an elongate, curved anterior arm; aedeagus triangular with slender, slightly curved basal arms, very low basal arch, and lightly sclerotized distal portion with rounded apex. Females with wing darkly infuscated with dense, coarse macrotrichia in all cells beyond r-m crossvein; and unequal size, pyriform spermathecae with short, broad, truncate necks.
</paragraph>
</subSubSection>
<subSubSection id="19C065359D35FF9FFF1489ADFBC98B31" lastPageId="8" lastPageNumber="263" pageId="5" pageNumber="260" type="materials_examined">
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<emphasis id="63AEEAAC9D35FF92FF1489ADFE6F8A8C" bold="true" box="[199,386,1419,1444]" pageId="5" pageNumber="260">
<typeStatus id="8E61881C9D35FF92FF1489ADFED98A8C" box="[199,308,1419,1444]" pageId="5" pageNumber="260" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Head (
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) brown. Eyes narrowly contiguous with sparse interommatidial spicules. Antennal pedicel dark brown; flagellomeres 2̄8 globular, 9̄10 vasiform, 11̄13 elongate, 11 longest; plume sparse, well developed, extending beyond flagellomere 11; flagellomere 12 with single subapical sensillum coeloconicum; antennal ratio 0.81. Clypeus with 3 pairs of setae. Palpus moderately long, segments 4̄5 extending beyond proboscis; segment 3 moderately broad with small apicomesal pit; palpal ratio 2.33. Thorax brown except for pale scutellum; scutellum with 6 large, 8 smaller setae. Legs similar to those of female except for tarsal claws, which are shorter, equal size, without basal inner teeth, apices bifid; hind tibial ratio 9.00; hind tarsal ratio 1.68. Wing (
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) membrane slightly infuscated, surface with dense macrotrichia along distal margin of cell r3, sparse above and below distal portion of vein M1 and apex of cell m1; veins brown, base of M2 thin, joined to M1; radial cells well developed, 2nd broader, nearly twice as long as 1st; wing length
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, width
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; costal ratio 0.57. Halter pale. Genitalia (
<figureCitation id="C9E12A3B9D35FF92FE078ADDFDDC883C" box="[468,561,1787,1812]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="5" pageNumber="260">Figs. 15</figureCitation>
,
<figureCitation id="C9E12A3B9D35FF92FDEE8ADAFDB6883C" box="[573,603,1788,1812]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="5" pageNumber="260">31</figureCitation>
̄32). Tergite 9 tapering gradually distally, posterior margin narrow, deeply concave, apicolateral processes elongate, cylindrical, slightly divergent with a single apical seta; cercus slender; sternite 9 3.5 x broader than long with narrow, moderately deep, concave posteromedian excavation. Gonocoxite short, stout, 1.25 x longer than broad, apex broad, sinuate; gonostylus 1.2 x longer than gonocoxite, broadest on proximal 1/5, tapering gradually distally, distal 2/3 slightly curved, apex rounded. Parameres (
<figureCitation id="C9E12A3B9D35FF92FB008BADFACA888C" box="[1235,1319,1931,1956]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="5" pageNumber="260">Fig. 31</figureCitation>
) separate, moderately slender, somewhat boomerang-shaped; basal apodeme with elongate anterior arm, apical 1/3 curved, posterior arm shorter with slightly expanded blunt apex; distal portions moderately short, curved distally, apices pointed, divergent. Aedeagus (
<figureCitation id="C9E12A3B9D38FF9FFDD18CB1FDB68F98" box="[514,603,151,176]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="8" pageNumber="263">Fig. 32</figureCitation>
) triangular; basal arch very low, slightly concave; basal arm heavily sclerotized, slender, nearly straight, laterally directed with blunt apex; distal portion barely discernible, lightly sclerotized, tapered distally, apex rounded.
</paragraph>
<caption id="05A566369D36FF91FF448BF0FC3988C5" box="[151,980,2006,2029]" httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="6" pageNumber="261" targetBox="[201,1416,202,1972]" targetPageId="6">
<paragraph id="516536BE9D36FF91FF448BF0FC3988C5" blockId="6.[151,980,2006,2029]" box="[151,980,2006,2029]" pageId="6" pageNumber="261">
<emphasis id="63AEEAAC9D36FF91FF448BF0FEBE88C4" bold="true" box="[151,339,2006,2029]" pageId="6" pageNumber="261">FIGURES 13–15.</emphasis>
<taxonomicName id="96DA4D3D9D36FF91FE898BF1FDEF88C5" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[346,514,2007,2029]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D36FF91FE898BF1FDEF88C5" box="[346,514,2007,2029]" italics="true" pageId="6" pageNumber="261">Austrohelea sirii</emphasis>
</taxonomicName>
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<emphasis id="63AEEAAC9D36FF91FDDB8BF1FDAC88C4" bold="true" box="[520,577,2007,2028]" pageId="6" pageNumber="261">n. sp.</emphasis>
</taxonomicNameLabel>
Male. 13. head. 14. wing. 15. genitalia.
</paragraph>
</caption>
<caption id="05A566369D37FF90FF448B9AFEBE88DA" httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="7" pageNumber="262" targetBox="[205,1391,209,1933]" targetPageId="7">
<paragraph id="516536BE9D37FF90FF448B9AFEBE88DA" blockId="7.[151,1436,1980,2034]" pageId="7" pageNumber="262">
<emphasis id="63AEEAAC9D37FF90FF448B9AFEB988FA" bold="true" box="[151,340,1980,2003]" pageId="7" pageNumber="262">FIGURES 16–19.</emphasis>
<taxonomicName id="96DA4D3D9D37FF90FE888B9BFDE988FB" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[347,516,1981,2003]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D37FF90FE888B9BFDE988FB" box="[347,516,1981,2003]" italics="true" pageId="7" pageNumber="262">Austrohelea sirii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D37FF90FDD88B98FDA888FB" box="[523,581,1982,2003]" pageId="7" pageNumber="262" rank="species">
<emphasis id="63AEEAAC9D37FF90FDD88B98FDA888FB" bold="true" box="[523,581,1982,2003]" pageId="7" pageNumber="262">n. sp.</emphasis>
</taxonomicNameLabel>
Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7–9 and spermathecae.
</paragraph>
</caption>
<paragraph id="516536BE9D38FF9FFF148D22FB128CA0" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
<emphasis id="63AEEAAC9D38FF9FFF148D22FE678E35" bold="true" box="[199,394,260,285]" pageId="8" pageNumber="263">
<typeStatus id="8E61881C9D38FF9FFF148D22FEC18E35" box="[199,300,260,285]" pageId="8" pageNumber="263" type="allotype">Allotype</typeStatus>
female.
</emphasis>
Head (
<figureCitation id="C9E12A3B9D38FF9FFE0E8D22FDC28E35" box="[477,559,260,285]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 16</figureCitation>
) mostly dark brown. Eyes barely contiguous on extreme lower portions, with sparse interommatidial spicules. Antennal pedicel dark brown, flagellum light brown; flagellomeres 2-8 (
<figureCitation id="C9E12A3B9D38FF9FFAE88D01FA638E68" box="[1339,1422,295,320]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 17</figureCitation>
), moderately short, barrel-shaped, 9-13 elongate, cylindrical, 13 longest; flagellomeres 8, 12 with single subapical sensillum coeloconicum; antennal ratio 1.19. Clypeus with 3 pairs of setae. Palpus (
<figureCitation id="C9E12A3B9D38FF9FFBE88D49FB618EA0" box="[1083,1164,367,392]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 17</figureCitation>
) light brown; segment 3 moderately stout, with broad, shallow, subapicomesal sensory pit; palpal ratio 2.16. Mandible with 8 coarse teeth. Thorax. Scutum uniformly brown; scutellum pale with 6-7 large, 8 smaller setae. Femora, tibiae slender, uniformly brown (mid tibiae, tarsi missing), tarsi slightly paler; tarsomere 1 of fore, hind legs with ventral basal, apical spines; hind tarsomeres 1̄2 with two rows of ventral palisade setae, apparently with moderately numerous large setae (only insertions remain); hind tibial comb with 6 spines; hind tibial ratio 7.60; hind tarsal ratio 1.80; tarsomere 4 of fore leg short, cordiform, tarsomere 4 of hind leg elongate, cylindrical; claws small, equal size on fore leg, hind leg with single long claw that is slightly longer than tarsomere 5 with a stout basal tooth about 1/3 as long as claw. Wing (
<figureCitation id="C9E12A3B9D38FF9FFEA88EA9FE218D80" box="[379,460,655,680]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 18</figureCitation>
) membrane infuscated, particularly in radial sector, distal 2/3 of cell r3 and along veins; two well developed radial cells, 2nd broader, slightly longer than 1st; macrotrichia numerous on distal 2/3 of membrane, particularly dense on distal half of cell r3, and margins of m1, m2, about 40 on distal ½ of anal cell; wing length
<quantity id="96229B5B9D38FF9FFF378F22FEA68C34" box="[228,331,772,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="8" pageNumber="263" unit="mm" value="1.16">1.16 mm</quantity>
, width
<quantity id="96229B5B9D38FF9FFE4D8F23FDEB8C34" box="[414,518,773,797]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="8" pageNumber="263" unit="mm" value="0.47">0.47 mm</quantity>
; costal ratio 0.60. Halter missing. Abdomen (
<figureCitation id="C9E12A3B9D38FF9FFBD58F22FBB58C35" box="[1030,1112,772,797]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 19</figureCitation>
) dark brown. Sternite 8 with V-shaped posteromedian cleft; sternite 9 halves moderately broad with rounded apices. Sternite 10 with two pairs of setae. Two unequal spermathecae; the larger pyriform with short, broad truncate neck, measuring 75 µm by 42 µm; the smaller partially collapsed with short narrow neck; a rudimentary third spermatheca present.
</paragraph>
<paragraph id="516536BE9D38FF9FFF148FB2FBC98B31" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
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<emphasis id="63AEEAAC9D38FF9FFF148FB2FE988C85" bold="true" box="[199,373,916,941]" pageId="8" pageNumber="263">
<typeStatus id="8E61881C9D38FF9FFF148FB2FEEF8C85" box="[199,258,916,941]" pageId="8" pageNumber="263">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8E61881C9D38FF9FFEA88FB2FE0E8C85" box="[379,483,916,941]" pageId="8" pageNumber="263" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="29CD762E9D38FF9FFDFD8FB3FD398C85" box="[558,724,917,941]" name="Argentina" pageId="8" pageNumber="263">ARGENTINA</collectingCountry>
,
<collectingRegion id="931EF85C9D38FF9FFD338FB3FCA58C84" box="[736,840,917,940]" country="Argentina" name="Neuquen" pageId="8" pageNumber="263">Neuquen</collectingRegion>
,
<collectingMunicipality id="B101ACC49D38FF9FFC808FB3FB5F8C85" box="[851,1202,916,941]" pageId="8" pageNumber="263">Parque Nacional Nahuel Huapi</collectingMunicipality>
, mallin
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</collectorName>
,
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,
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,
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,
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<date id="2564107E9D38FF9FFD838F9FFD5E8CF9" box="[592,691,953,977]" pageId="8" pageNumber="263" value="2007-01-07">7-I-2007</date>
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/
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<date id="2564107E9D38FF9FFD688F9FFD008CF9" box="[699,749,953,977]" bridgedPair="-" pageId="8" pageNumber="263">4-II-</date>
</collectingDate>
2 0 0 7,
<collectorName id="FC2F53689D38FF9FFCEA8F9FFC4B8CF9" box="[825,934,953,977]" pageId="8" pageNumber="263">A. Garré</collectorName>
̄
<collectorName id="FC2F53689D38FF9FFC638F9FFB658CF9" box="[944,1160,952,977]" pageId="8" pageNumber="263">F. Montes de Oca</collectorName>
,
<collectingMethod id="889B4EA99D38FF9FFB4A8F9EFAC78CF8" box="[1177,1322,952,977]" pageId="8" pageNumber="263">Malaise trap</collectingMethod>
(
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)
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;
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<typeStatus id="8E61881C9D38FF9FFF448FFAFF1C8CDD" box="[151,241,988,1013]" pageId="8" pageNumber="263" type="allotype">allotype</typeStatus>
female,
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,
<collectingMunicipality id="B101ACC49D38FF9FFE0A8FFBFCA18CDD" box="[473,844,988,1013]" pageId="8" pageNumber="263">Parque Nacional Nahuel Huapi</collectingMunicipality>
, laguna
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,
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,
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,
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,
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<date id="2564107E9D38FF9FFF218826FEB08B31" box="[242,349,1024,1049]" pageId="8" pageNumber="263" value="2007-02-05">5-II-2007</date>
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/
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<date id="2564107E9D38FF9FFEB48826FE4F8B31" box="[359,418,1024,1049]" bridgedPair="-" pageId="8" pageNumber="263">3-III-</date>
</collectingDate>
2 0 0 7,
<collectorName id="FC2F53689D38FF9FFE348827FDA28B31" box="[487,591,1025,1049]" pageId="8" pageNumber="263">A. Garré</collectorName>
̄
<collectorName id="FC2F53689D38FF9FFD8A8827FCCC8B31" box="[601,801,1024,1049]" pageId="8" pageNumber="263">F. Montes de Oca</collectorName>
,
<collectingMethod id="889B4EA99D38FF9FFCFF8826FC5A8B30" box="[812,951,1024,1049]" pageId="8" pageNumber="263">Malaise trap</collectingMethod>
(
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="19C065359D38FF9FFF148802FCC98BD9" pageId="8" pageNumber="263" type="distribution">
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<emphasis id="63AEEAAC9D38FF9FFF148802FE8D8B15" bold="true" box="[199,352,1060,1085]" pageId="8" pageNumber="263">Distribution.</emphasis>
Known only from the
<typeStatus id="8E61881C9D38FF9FFD8C8803FD628B15" box="[607,655,1061,1085]" pageId="8" pageNumber="263">type</typeStatus>
locality, the
<taxonomicName id="96DA4D3D9D38FF9FFCCC8802FC4D8B14" authorityName="Blume" authorityYear="1850" box="[799,928,1060,1084]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="263" phylum="Tracheophyta" rank="genus">
<emphasis id="63AEEAAC9D38FF9FFCCC8802FC4D8B14" box="[799,928,1060,1084]" italics="true" pageId="8" pageNumber="263">Nothofagus</emphasis>
</taxonomicName>
forests of the Nahuel Huapi National Park of
<collectingRegion id="931EF85C9D38FF9FFF44886FFE858B49" box="[151,360,1096,1121]" country="Argentina" name="Neuquen" pageId="8" pageNumber="263">Neuquen province</collectingRegion>
,
<collectingCountry id="29CD762E9D38FF9FFEA0886EFE0F8B49" box="[371,482,1096,1121]" name="Argentina" pageId="8" pageNumber="263">Argentina</collectingCountry>
(
<figureCitation id="C9E12A3B9D38FF9FFE21886EFDA88B49" box="[498,581,1096,1121]" captionStart="FIGURE 35" captionStartId="13.[151,250,1234,1256]" captionTargetBox="[267,1325,252,1118]" captionTargetId="figure@13.[205,1396,196,1194]" captionTargetPageId="13" captionText="FIGURE 35. Map of Argentine and Chilean Patagonia with distribution records of: Austrohelea shannoni (Wirth &amp; Blanton), green circles. Austrohelea sirii n. sp., yellow triangles. Austrohelea spinosa n. sp., red diamonds." httpUri="https://zenodo.org/record/806131/files/figure.png" pageId="8" pageNumber="263">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="516536BE9D38FF9FFF14884AFCC98BD9" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
<emphasis id="63AEEAAC9D38FF9FFF14884AFDBE8BAD" bold="true" box="[199,595,1132,1157]" pageId="8" pageNumber="263">Derivation of the specific epithet.</emphasis>
We are pleased to name this new species after our colleague Augusto Siri, at the Instituto de Limnología “Dr. Raúl A. Ringuelet”, La Plata,
<collectingCountry id="29CD762E9D38FF9FFC7E88B6FBF28B81" box="[941,1055,1168,1193]" name="Argentina" pageId="8" pageNumber="263">Argentina</collectingCountry>
, in recognition of his important contributions to the taxonomy of
<taxonomicName id="96DA4D3D9D38FF9FFDF28892FD2C8BE5" authorityName="Newman" authorityYear="1834" box="[545,705,1204,1229]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
in the Neotropical Region, and for his valuable collections of
<taxonomicName id="96DA4D3D9D38FF9FFF4488FEFEBB8BD9" authorityName="Newman" authorityYear="1834" box="[151,342,1240,1265]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
from
<collectingCountry id="29CD762E9D38FF9FFE4888FEFDE78BD9" box="[411,522,1240,1265]" name="Argentina" pageId="8" pageNumber="263">Argentina</collectingCountry>
during the past 10 years.
</paragraph>
</subSubSection>
<subSubSection id="19C065359D38FF9FFF1488DAFB8F891D" pageId="8" pageNumber="263" type="discussion">
<paragraph id="516536BE9D38FF9FFF1488DAFAB68A8D" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
<emphasis id="63AEEAAC9D38FF9FFF1488DAFEA18A3D" bold="true" box="[199,332,1276,1301]" pageId="8" pageNumber="263">Discussion.</emphasis>
This new species is readily distinguished from its Neotropical congeners by its small size and the well-defined sensory pit on palpal segment 3. The male gonocoxite is very short, stout and lacks a basomesal tubercle; the gonostylus is much longer than the gonocoxite; and, the aedeagus is small, lightly sclerotized with a very low slightly concave basal arch, the apex is rounded and lacks a distal extension. The female wing membrane has dense, coarse macrotrichia and the spermathecae are pyriform, unequal, with short, broad truncate necks.
</paragraph>
<paragraph id="516536BE9D38FF9FFF148997FB8F891D" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
Two other species from Australasia also have a sensory pit on palpal segment 3. In his original description of
<taxonomicName id="96DA4D3D9D38FF9FFF4489F3FEFC8AC4" authorityName="Macfie" authorityYear="1932" baseAuthorityName="Macfie" baseAuthorityYear="1932" box="[151,273,1493,1516]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="tonnoiri">
<emphasis id="63AEEAAC9D38FF9FFF4489F3FF458AC4" box="[151,168,1493,1516]" italics="true" pageId="8" pageNumber="263">A</emphasis>
.
<emphasis id="63AEEAAC9D38FF9FFF6489F3FEFC8AC4" box="[183,273,1493,1516]" italics="true" pageId="8" pageNumber="263">tonnoiri</emphasis>
</taxonomicName>
from
<collectingCountry id="29CD762E9D38FF9FFE8B89F3FE1F8AC5" box="[344,498,1492,1517]" name="New Zealand" pageId="8" pageNumber="263">New Zealand</collectingCountry>
,
<bibRefCitation id="354B4B4F9D38FF9FFE2D89F2FD4D8AC5" author="Macfie" box="[510,672,1492,1517]" pageId="8" pageNumber="263" refString="Macfie, J. W. S. (1932) New Zealand biting midges (Diptera, Ceratopogonidae). Annals of Tropical Medicine and Parasitology, 26, 23 - 53." type="journal article" year="1932">Macfie (1932)</bibRefCitation>
stated “…third (palpal segment) not inflated, bearing sensory hairs on a depressed area on apical third” and
<bibRefCitation id="354B4B4F9D38FF9FFDBA89DEFCC78939" author="Tokunaga" box="[617,810,1528,1553]" pageId="8" pageNumber="263" refString="Tokunaga, M. (1964) Insects of Campbell Island. Diptera: Ceratopogonidae. Pacific Insects Monograph, 7, 289 - 291." type="journal article" year="1964">Tokunaga (1964)</bibRefCitation>
noted for
<taxonomicName id="96DA4D3D9D38FF9FFC7189DFFB8F8938" authorityName="Tokunaga" authorityYear="1964" baseAuthorityName="Tokunaga" baseAuthorityYear="1964" box="[930,1122,1528,1552]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="campbellensis">
<emphasis id="63AEEAAC9D38FF9FFC7189DFFC5E8938" box="[930,947,1529,1552]" italics="true" pageId="8" pageNumber="263">A</emphasis>
.
<emphasis id="63AEEAAC9D38FF9FFC1089DEFB8F8938" box="[963,1122,1528,1552]" italics="true" pageId="8" pageNumber="263">campbellensis</emphasis>
</taxonomicName>
from Campbell Island, that palpal segment 3 is “…slightly thickened, with very small sensory pore on apical 1/3.”
</paragraph>
</subSubSection>
</treatment>
</document>

342
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<mods:title id="0F3896ED4F0BD0C44B2520280371B38E">The Neotropical species of the predaceous midge genus Austrohelea Wirth &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
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<subSubSection id="19C065359D38FF9FFF448AA1FE6889EB" pageId="8" pageNumber="263" type="nomenclature">
<paragraph id="516536BE9D38FF9FFF448AA1FDC0898A" blockId="8.[151,557,1671,1732]" box="[151,557,1671,1698]" pageId="8" pageNumber="263">
<heading id="0A2D81D29D38FF9FFF448AA1FDC0898A" bold="true" box="[151,557,1671,1698]" fontSize="11" level="1" pageId="8" pageNumber="263" reason="1">
<emphasis id="63AEEAAC9D38FF9FFF448AA1FDC0898A" bold="true" box="[151,557,1671,1698]" pageId="8" pageNumber="263">
<taxonomicName id="96DA4D3D9D38FF9FFF448AA1FE618989" ID-CoL="9HWY2" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[151,396,1671,1697]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D38FF9FFF448AA1FE618989" bold="true" box="[151,396,1671,1697]" italics="true" pageId="8" pageNumber="263">Austrohelea spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D38FF9FFE408AAEFDC0898A" box="[403,557,1672,1698]" pageId="8" pageNumber="263" rank="species">New Species</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="516536BE9D38FF9FFF448A8DFE6889EB" blockId="8.[151,557,1671,1732]" box="[151,389,1707,1732]" pageId="8" pageNumber="263">
(
<figureCitation id="C9E12A3B9D38FF9FFF4C8A8DFF1389EB" box="[159,254,1707,1732]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="8" pageNumber="263">Figs. 20</figureCitation>
̄28, 33̄34)
</paragraph>
</subSubSection>
<subSubSection id="19C065359D38FF9FFF448AD5FCEF88CC" pageId="8" pageNumber="263" type="diagnosis">
<paragraph id="516536BE9D38FF9FFF448AD5FCEF88CC" blockId="8.[151,1437,1779,2020]" pageId="8" pageNumber="263">
<emphasis id="63AEEAAC9D38FF9FFF448AD5FEFC8824" bold="true" box="[151,273,1779,1804]" pageId="8" pageNumber="263">Diagnosis.</emphasis>
A large (male wing length 1.50̄
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, female wing length 1.57̄
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), dark brown species. Males with hind tibia with 3 long, stout, black subapical spines; tergite 9 with narrow shallow posterior excavation; gonocoxite short, stout, tapering slightly distally with basomesal tubercle; gonostylus longer than gonocoxite; parameres with very broad basal apodemes that form an obtuse angle between the anterior and posterior arms; and the aedeagus is very large, heavily sclerotized with long basal arms, a U-shaped basal arch and lightly sclerotized apical extension. Females with palpal segment 3 lacking sensory pit, hind femur greatly swollen, and subequal, ovoid spermathecae with moderately long broad necks.
</paragraph>
</subSubSection>
<caption id="05A566369D39FF9EFF448B9CFF1888D8" httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="9" pageNumber="264" targetBox="[260,1331,213,1923]" targetPageId="9">
<paragraph id="516536BE9D39FF9EFF448B9CFF1888D8" blockId="9.[151,1436,1978,2032]" pageId="9" pageNumber="264">
<emphasis id="63AEEAAC9D39FF9EFF448B9CFEBB88F8" bold="true" box="[151,342,1978,2001]" pageId="9" pageNumber="264">FIGURES 20–24.</emphasis>
<taxonomicName id="96DA4D3D9D39FF9EFE8D8B9DFDC188F9" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[350,556,1979,2001]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D39FF9EFE8D8B9DFDC188F9" box="[350,556,1979,2001]" italics="true" pageId="9" pageNumber="264">Austrohelea spinosa</emphasis>
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<emphasis id="63AEEAAC9D39FF9EFDE58B9DFD9C88F8" bold="true" box="[566,625,1979,2000]" pageId="9" pageNumber="264">n. sp.</emphasis>
</taxonomicNameLabel>
Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia.
</paragraph>
</caption>
<caption id="05A566369D3AFF9DFF448B95FE6188C0" httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="10" pageNumber="265" targetBox="[203,1371,219,1911]" targetPageId="10">
<paragraph id="516536BE9D3AFF9DFF448B95FE6188C0" blockId="10.[151,1436,1971,2024]" pageId="10" pageNumber="265">
<emphasis id="63AEEAAC9D3AFF9DFF448B95FEB888E1" bold="true" box="[151,341,1971,1994]" pageId="10" pageNumber="265">FIGURES 25–28.</emphasis>
<taxonomicName id="96DA4D3D9D3AFF9DFE8F8B92FDC488E2" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[348,553,1972,1994]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D3AFF9DFE8F8B92FDC488E2" box="[348,553,1972,1994]" italics="true" pageId="10" pageNumber="265">Austrohelea spinosa</emphasis>
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<emphasis id="63AEEAAC9D3AFF9DFDE28B92FD8688E1" bold="true" box="[561,619,1972,1993]" pageId="10" pageNumber="265">n. sp.</emphasis>
</taxonomicNameLabel>
Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7–10 and spermathecae.
</paragraph>
</caption>
<subSubSection id="19C065359D3BFF9CFF148CB1FE6F883A" pageId="11" pageNumber="266" type="description">
<paragraph id="516536BE9D3BFF9CFF148CB1FE678B31" blockId="11.[151,1437,151,1990]" pageId="11" pageNumber="266">
<emphasis id="63AEEAAC9D3BFF9CFF148CB1FEE68F98" bold="true" box="[199,267,151,176]" pageId="11" pageNumber="266">Male.</emphasis>
Head (
<figureCitation id="C9E12A3B9D3BFF9CFE888CB1FE468F98" box="[347,427,151,176]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 20</figureCitation>
) dark brown. Eyes barely contiguous with fine interommatidial spicules. Antennal pedicel very dark brown; flagellomeres 2̄8 globular, 9̄10 slightly elongate, cylindrical, 11̄13 elongate, 11 longest; plume sparse, well developed, extending beyond apex of flagellomere 11; flagellomere 12 with single subapical sensillum coeloconicum; antennal ratio 0.72̄0.77 (0.75, n = 3). Clypeus with 2̄3 pairs of setae. Palpus slender, extending beyond apex of proboscis; segment 3 without apical pit; palpal ratio 3.64̄4.02 (3.78, n = 3). Thorax. Dark brown; scutellum slightly paler with 9̄11 large, 18̄25 smaller setae. Legs (
<figureCitation id="C9E12A3B9D3BFF9CFBE08D6AFB688E4C" box="[1075,1157,332,357]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 21</figureCitation>
) dark brown, fore femur slightly paler; fore, mid femora stout, hind femur greatly swollen; hind tibia broader than fore, mid tibiae, hind tibia (
<figureCitation id="C9E12A3B9D3BFF9CFF4C8DB2FF188E84" box="[159,245,404,429]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 22</figureCitation>
) with 3 long, black subapical spines and moderately numerous coarse setae on ventral surface (only insertions remain on most specimens); hind tibial comb with 5 long spines; hind tarsomere 1 with large basal, apical spines; hind tibial ratio 7.90̄8.57 (8.16, n = 3); hind tarsal ratio 1.82̄2.00 (1.89, n = 3); tarsomeres 4 cylindrical; claws short, equal size, without basal inner teeth, apices bifid. Wing (
<figureCitation id="C9E12A3B9D3BFF9CFBE18DD9FB698D30" box="[1074,1156,511,536]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 23</figureCitation>
) membrane mostly pale, lightly infuscated on costa, radius and base of medius and on membrane between these veins; veins M1, M2, CuA1, CuA2 and A infuscated; base of M2 obsolete; macrotrichia present on costa, radius and a few along anterior margin of cell r3; wing length 1.50̄1.60 (1.65, n = 3) mm, width 0.50̄0.54 (0.53, n = 3) mm; costal ratio 0.58̄0.62 (0.60, n = 3). Halter pale. Abdomen. Brown. Genitalia (
<figureCitation id="C9E12A3B9D3BFF9CFD1A8EBEFCC58D99" box="[713,808,664,689]" captionStart="FIGURES 20 – 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 – 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Figs. 24</figureCitation>
,
<figureCitation id="C9E12A3B9D3BFF9CFCE68EBFFCB98D99" box="[821,852,665,689]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="11" pageNumber="266">33</figureCitation>
̄34). Tergite 9 tapering gradually distally, posterior margin straight, apicolateral processes cylindrical, moderately long with a single apical seta, cercus hyaline, slender with 3-4 setae; sternite 9 2.2 x broader than long with narrow shallow, rounded posteromedian excavation. Gonocoxite short, stout, tapering slightly distally, with a small basomesal tubercle; gonostylus 1.3 x longer than gonocoxite, nearly straight, apex broad, spoon-shaped, tip pointed. Parameres (
<figureCitation id="C9E12A3B9D3BFF9CFBE88F0EFB7C8C69" box="[1083,1169,808,833]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="11" pageNumber="266">Fig. 33</figureCitation>
) separate, boomerangshaped; basal apodeme very broad, forming an obtuse angle between anterior and posterior arms; distal portion broadest proximally, curved, tapered distally to mid length, apices recurved, with widely divergent pointed tips. Aedeagus (
<figureCitation id="C9E12A3B9D3BFF9CFEC48FB2FE868C85" box="[279,363,916,941]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="11" pageNumber="266">Fig. 34</figureCitation>
) triangular, heavily sclerotized; basal arch U-shaped, extending 1/3 of total aedeagus length; basal arm broad, recurved 90˚ near base, distal portion long, tapered apically, apex rounded; distal portion tapering abruptly distally to a narrow sharply pointed tip, with a lightly sclerotized underlying apical process with a bulbous apex and rounded tip.
</paragraph>
<paragraph id="516536BE9D3BFF9CFF148802FE6F883A" blockId="11.[151,1437,151,1990]" pageId="11" pageNumber="266">
<emphasis id="63AEEAAC9D3BFF9CFF148802FEC88B15" bold="true" box="[199,293,1060,1085]" pageId="11" pageNumber="266">Female.</emphasis>
Head (
<figureCitation id="C9E12A3B9D3BFF9CFEAD8802FE3E8B15" box="[382,467,1060,1085]" captionStart="FIGURES 25 – 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 – 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 – 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 25</figureCitation>
) dark brown. Eyes barely contiguous ventrally, with fine interommatidial spicules. Antennal pedicel very dark brown; flagellum dark brown, flagellomeres 2̄8 (
<figureCitation id="C9E12A3B9D3BFF9CFBC8886EFB828B49" box="[1051,1135,1096,1121]" captionStart="FIGURES 25 – 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 – 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 – 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 26</figureCitation>
) moderately short, barrelshaped, 9̄13 elongate, cylindrical; flagellomeres 8, 12 with single subapical sensillum coeloconicum; antennal ratio 1.36̄1.47 (1.39, n = 5). Clypeus with 2̄4 pairs of setae. Palpus (
<figureCitation id="C9E12A3B9D3BFF9CFC6088B6FBE98B81" box="[947,1028,1168,1193]" captionStart="FIGURES 25 – 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 – 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 – 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 26</figureCitation>
) dark brown; segment 3 moderately slender with a few capitate sensilla on distal 1/3 of mesal surface; palpal ratio 3.00 (n = 5). Mandible with 8̄11 (9, n = 5) large teeth. Thorax. Scutum dark brown, humeral areas and scutellum paler; scutellum with 6̄7 large, 10̄12 smaller setae. Legs dark brown, hind tibia and femur darkest; fore, mid femora moderately swollen, hind femur greatly swollen; hind tibia without subapical spines, stouter than fore, mid tibiae; hind tibial comb with 5 large spines; hind tibial ratio 5.93̄7.17 (6.42, n = 5); tarsi dark brown, darkest on hind leg; tarsomeres 1 with large basal, apical ventral spines, 2 additional similar spines along ventral surface of mid leg, tarsomeres 2̄3 of fore, mid legs with apical spine; tarsomeres 1̄2 of hind leg with two rows of ventral palisade setae; hind tarsal ratio 1.76̄1.88 (1.83, n = 5); tarsomere 4 of fore, mid legs short, cordiform, tarsomere 4 of hind leg elongate, cylindrical; claws small, equal on fore, mid legs, hind leg with single long claw that is slightly longer than tarsomere 5 with a stout basal tooth about 1/3 as long as claw. Wing (
<figureCitation id="C9E12A3B9D3BFF9CFD5589DEFD3A8939" box="[646,727,1528,1553]" captionStart="FIGURES 25 – 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 – 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 – 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 27</figureCitation>
) membrane infuscated, particularly dark along veins and in and around radial cells; two well developed radial cells, 2nd broader, twice as long as1st; macrotrichia fairly numerous on distal ½ of cell r3, apices of m1, m2, a few in cua1 and anal cell; wing length 1.57̄1.68 (1.63, n = 5) mm, width 0.65̄0.68 (0.67, n = 5) mm; costal ratio 0.64̄0.66 (0.65, n = 5). Halter pale. Abdomen (
<figureCitation id="C9E12A3B9D3BFF9CFB788A4CFB1289AA" box="[1195,1279,1642,1667]" captionStart="FIGURES 25 – 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 – 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 – 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 28</figureCitation>
) dark brown. Sternite 8 with moderately deep V-shaped posteromedian excavation; sternite 9 separated, each half tapered distally with hook-like apices; sternite 10 with two pairs of setae. Two ovoid, subequal size spermathecae with moderately long, broad necks, measuring 55̄62 (59, n = 5) µm by 35̄42 (39, n = 5) µm; a small rudimentary third spermatheca present.
</paragraph>
</subSubSection>
<subSubSection id="19C065359D3BFF9CFF148B3BFA9188EE" pageId="11" pageNumber="266" type="materials_examined">
<paragraph id="516536BE9D3BFF9CFF148B3BFA9188EE" blockId="11.[151,1437,151,1990]" pageId="11" pageNumber="266">
<materialsCitation id="E1B23CE39D3BFF9CFF148B3BFED48872" ID-GBIF-Occurrence="1563642360" collectingDate="2004-01-15" collectionCode="MLPA" collectorName="Villa La Angostura &amp; G. Spinelli" country="Argentina" location="Neuquen" pageId="11" pageNumber="266" specimenCount="2" stateProvince="Neuquen" typeStatus="holotype">
<emphasis id="63AEEAAC9D3BFF9CFF148B3BFE95881E" bold="true" box="[199,376,1821,1846]" pageId="11" pageNumber="266">
<typeStatus id="8E61881C9D3BFF9CFF148B3BFEEF881E" box="[199,258,1821,1846]" pageId="11" pageNumber="266">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8E61881C9D3BFF9CFE508B3BFE06881E" box="[387,491,1821,1846]" pageId="11" pageNumber="266" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="29CD762E9D3BFF9CFDED8B38FD09881E" box="[574,740,1822,1846]" name="Argentina" pageId="11" pageNumber="266">ARGENTINA</collectingCountry>
,
<collectingRegion id="931EF85C9D3BFF9CFD218B39FCB7881E" box="[754,858,1823,1846]" country="Argentina" name="Neuquen" pageId="11" pageNumber="266">Neuquen</collectingRegion>
,
<collectorName id="FC2F53689D3BFF9CFCBA8B3BFBA7881E" box="[873,1098,1821,1846]" pageId="11" pageNumber="266">Villa La Angostura</collectorName>
, 11/
<collectingDate id="3520E9969D3BFF9CFBA88B3BFB03881E" box="[1147,1262,1821,1846]" pageId="11" pageNumber="266" value="2004-01-15">
<date id="2564107E9D3BFF9CFBA88B3BFB03881E" box="[1147,1262,1821,1846]" pageId="11" pageNumber="266" value="2004-01-15">15-I-2004</date>
</collectingDate>
,
<collectorName id="FC2F53689D3BFF9CFB2E8B38FA94881E" box="[1277,1401,1821,1846]" pageId="11" pageNumber="266">G. Spinelli</collectorName>
, at light (
<collectionCode id="37CBAE7B9D3BFF9CFF0C8B64FEDC8872" box="[223,305,1858,1882]" pageId="11" pageNumber="266">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1B23CE39D3BFF9CFE998B64FA748856" ID-GBIF-Occurrence="1563642355" collectingDate="2006-12-14" collectingDateMax="2007-01-08" collectingDateMin="2006-12-14" collectionCode="MLPA" collectorName="La Cortadera &amp; A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="769" latitude="-41.086945" location="La Cortadera" longLatPrecision="19" longitude="-71.80723" municipality="Parque Nacional Nahuel Huapi" pageId="11" pageNumber="266" specimenCount="1" stateProvince="Rio Negro" typeStatus="allotype">
<typeStatus id="8E61881C9D3BFF9CFE998B64FE498873" box="[330,420,1858,1883]" pageId="11" pageNumber="266" type="allotype">allotype</typeStatus>
female,
<collectingCountry id="29CD762E9D3BFF9CFDC28B64FD6E8873" box="[529,643,1858,1883]" name="Argentina" pageId="11" pageNumber="266">Argentina</collectingCountry>
,
<collectingRegion id="931EF85C9D3BFF9CFD468B64FCFE8873" box="[661,787,1858,1883]" country="Argentina" name="Rio Negro" pageId="11" pageNumber="266">Río Negro</collectingRegion>
,
<collectingMunicipality id="B101ACC49D3BFF9CFCF78B65FB7B8873" box="[804,1174,1858,1883]" pageId="11" pageNumber="266">Parque Nacional Nahuel Huapi</collectingMunicipality>
, mallín
<collectorName id="FC2F53689D3BFF9CFB2F8B65FA758873" box="[1276,1432,1858,1883]" pageId="11" pageNumber="266">
<location id="540560659D3BFF9CFB2F8B65FA758873" LSID="urn:lsid:plazi:treatment:D97387A89D38FF9AFF448AA1FCC38960:540560659D3BFF9CFB2F8B65FA758873" box="[1276,1432,1858,1883]" country="Argentina" latitude="-41.086945" longLatPrecision="19" longitude="-71.80723" municipality="Parque Nacional Nahuel Huapi" name="La Cortadera" pageId="11" pageNumber="266" stateProvince="Rio Negro">La Cortadera</location>
</collectorName>
,
<geoCoordinate id="34EE50799D3BFF9CFF448B43FEF48856" box="[151,281,1893,1918]" direction="south" orientation="latitude" pageId="11" pageNumber="266" precision="15" value="-41.086945">41°05'13&quot;S</geoCoordinate>
,
<geoCoordinate id="34EE50799D3BFF9CFEFF8B43FE5B8856" box="[300,438,1893,1918]" direction="west" orientation="longitude" pageId="11" pageNumber="266" precision="15" value="-71.80723">71°48'26&quot;W</geoCoordinate>
,
<elevation id="DAF7D18D9D3BFF9CFE1A8B43FDE68856" box="[457,523,1893,1918]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="11" pageNumber="266" unit="m" value="769.0">
<quantity id="96229B5B9D3BFF9CFE1A8B43FDE68856" box="[457,523,1893,1918]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="11" pageNumber="266" unit="m" value="769.0">769m</quantity>
</elevation>
,
<collectingDate id="3520E9969D3BFF9CFDCD8B43FD428856" box="[542,687,1893,1918]" pageId="11" pageNumber="266" value="2006-12-14">
<date id="2564107E9D3BFF9CFDCD8B43FD428856" box="[542,687,1893,1918]" pageId="11" pageNumber="266" value="2006-12-14">14-XII-2006</date>
</collectingDate>
/
<collectingDate id="3520E9969D3BFF9CFD648B43FCF58856" box="[695,792,1893,1918]" pageId="11" pageNumber="266" value="2007-01-08">
<date id="2564107E9D3BFF9CFD648B43FCF58856" box="[695,792,1893,1918]" pageId="11" pageNumber="266" value="2007-01-08">8-I-2007</date>
</collectingDate>
,
<collectorName id="FC2F53689D3BFF9CFCF88B40FC768856" box="[811,923,1894,1918]" pageId="11" pageNumber="266">A. Garré</collectorName>
̄
<collectorName id="FC2F53689D3BFF9CFC778B40FB6E8856" box="[932,1155,1893,1918]" pageId="11" pageNumber="266">F. Montes de Oca</collectorName>
,
<collectingMethod id="889B4EA99D3BFF9CFB458B43FAC48856" box="[1174,1321,1893,1918]" pageId="11" pageNumber="266">Malaise trap</collectingMethod>
(
<collectionCode id="37CBAE7B9D3BFF9CFAEC8B40FA7C8856" box="[1343,1425,1894,1918]" pageId="11" pageNumber="266">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E1B23CE39D3BFF9CFF448BADFB6A888A" ID-GBIF-Occurrence="1563642356" box="[151,1159,1930,1955]" collectingDate="2004-01-15" collectionCode="MLPA" collectorName="Villa La Angostura &amp; G. Spinelli" country="Argentina" location="Neuquen" pageId="11" pageNumber="266" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Neuquen" typeStatus="paratype">
<typeStatus id="8E61881C9D3BFF9CFF448BADFEEA888B" box="[151,263,1931,1955]" pageId="11" pageNumber="266" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="47DCFD379D3BFF9CFEC78BACFE9D888B" box="[276,368,1930,1955]" pageId="11" pageNumber="266" type="male">2 males</specimenCount>
,
<specimenCount id="47DCFD379D3BFF9CFEAE8BACFE02888B" box="[381,495,1930,1955]" pageId="11" pageNumber="266" type="female">4 females</specimenCount>
, as follows:
<specimenCount id="47DCFD379D3BFF9CFD518BACFD22888B" box="[642,719,1930,1955]" pageId="11" pageNumber="266" type="male">1 male</specimenCount>
with same data as holotype (
<collectionCode id="37CBAE7B9D3BFF9CFBFE8BACFB92888A" box="[1069,1151,1930,1954]" pageId="11" pageNumber="266">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1B23CE39D3BFF9CFB478BACFA9588EE" ID-GBIF-Occurrence="1563642363" collectingDate="2006-12-14" collectingDateMax="2007-01-08" collectingDateMin="2006-12-14" collectionCode="MLPA, BMNH, USNM, FSCA" collectorName="La Cortadera &amp; A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="769" latitude="-41.086945" location="La Cortadera" longLatPrecision="19" longitude="-71.80723" municipality="Parque Nacional Nahuel Huapi" pageId="11" pageNumber="266" specimenCount="10" specimenCount-female="8" specimenCount-male="2" stateProvince="Rio Negro" typeStatus="paratype">
<specimenCount id="47DCFD379D3BFF9CFB478BACFB08888B" box="[1172,1253,1930,1955]" pageId="11" pageNumber="266" type="male">1 male</specimenCount>
,
<specimenCount id="47DCFD379D3BFF9CFB208BACFA8C888B" box="[1267,1377,1930,1955]" pageId="11" pageNumber="266" type="female">4 females</specimenCount>
with same data as allotype (
<specimenCount id="47DCFD379D3BFF9CFE4B8B88FE0988EE" box="[408,484,1965,1990]" pageId="11" pageNumber="266" type="male">1 male</specimenCount>
,
<specimenCount id="47DCFD379D3BFF9CFE258B88FDBA88EE" box="[502,599,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFDA88B88FD2488EE" box="[635,713,1966,1990]" pageId="11" pageNumber="266">MLPA</collectionCode>
,
<specimenCount id="47DCFD379D3BFF9CFD068B88FCD888EE" box="[725,821,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFC898B89FC5C88EE" box="[858,945,1967,1990]" pageId="11" pageNumber="266">BMNH</collectionCode>
,
<specimenCount id="47DCFD379D3BFF9CFC688B88FBF088EE" box="[955,1053,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFB928B88FB7888EE" box="[1089,1173,1966,1990]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="266">USNM</collectionCode>
,
<specimenCount id="47DCFD379D3BFF9CFB4C8B88FAED88EE" box="[1183,1280,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFAF68B88FA9D88EE" box="[1317,1392,1966,1990]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="11" pageNumber="266">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="05A566369D3CFF9BFF448ADBFBD4881A" httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="12" pageNumber="267" targetBox="[175,1421,225,1689]" targetPageId="12">
<paragraph id="516536BE9D3CFF9BFF448ADBFBD4881A" blockId="12.[151,1437,1789,1842]" pageId="12" pageNumber="267">
<emphasis id="63AEEAAC9D3CFF9BFF448ADBFEB9883B" bold="true" box="[151,340,1789,1812]" pageId="12" pageNumber="267">FIGURES 29–34.</emphasis>
<taxonomicName id="96DA4D3D9D3CFF9BFE898ADBFD11883B" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[346,764,1789,1811]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3CFF9BFE898ADBFDDA883B" box="[346,567,1789,1811]" italics="true" pageId="12" pageNumber="267">Austrohelea shannoni</emphasis>
(Wirth &amp; Blanton)
</taxonomicName>
. Male. 29. parameres. 30. aedeagus.
<taxonomicName id="96DA4D3D9D3CFF9BFBA18ADBFAF6883C" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[1138,1307,1789,1812]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D3CFF9BFBA18ADBFAF6883C" box="[1138,1307,1789,1812]" italics="true" pageId="12" pageNumber="267">Austrohelea sirii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3CFF9BFAF18AD8FAB6883B" box="[1314,1371,1790,1811]" pageId="12" pageNumber="267" rank="species">
<emphasis id="63AEEAAC9D3CFF9BFAF18AD8FAB6883B" bold="true" box="[1314,1371,1790,1811]" pageId="12" pageNumber="267">n. sp.</emphasis>
</taxonomicNameLabel>
Male. 31. parameres. 32. aedeagus.
<taxonomicName id="96DA4D3D9D3CFF9BFE128B3AFD63881A" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[449,654,1820,1842]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D3CFF9BFE128B3AFD63881A" box="[449,654,1820,1842]" italics="true" pageId="12" pageNumber="267">Austrohelea spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3CFF9BFD478B3BFD20881A" box="[660,717,1821,1842]" pageId="12" pageNumber="267" rank="species">
<emphasis id="63AEEAAC9D3CFF9BFD478B3BFD20881A" bold="true" box="[660,717,1821,1842]" pageId="12" pageNumber="267">n. sp.</emphasis>
</taxonomicNameLabel>
Male. 33. parameres. 34. aedeagus.
</paragraph>
</caption>
<subSubSection id="19C065359D3CFF9BFF148B78FCE488CC" pageId="12" pageNumber="267" type="distribution">
<paragraph id="516536BE9D3CFF9BFF148B78FE1C88B4" blockId="12.[151,1436,1886,2020]" pageId="12" pageNumber="267">
<emphasis id="63AEEAAC9D3CFF9BFF148B78FE8D885F" bold="true" box="[199,352,1886,1911]" pageId="12" pageNumber="267">Distribution.</emphasis>
Known only from
<taxonomicName id="96DA4D3D9D3CFF9BFDE68B79FD5B885F" authorityName="Blume" authorityYear="1850" box="[565,694,1887,1911]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="267" phylum="Tracheophyta" rank="genus">
<emphasis id="63AEEAAC9D3CFF9BFDE68B79FD5B885F" box="[565,694,1887,1911]" italics="true" pageId="12" pageNumber="267">Nothofagus</emphasis>
</taxonomicName>
forests of Nahuel Huapi National Park in
<collectingRegion id="931EF85C9D3CFF9BFB5F8B46FB1C885F" box="[1164,1265,1888,1911]" country="Argentina" name="Neuquen" pageId="12" pageNumber="267">Neuquen</collectingRegion>
and
<collectingRegion id="931EF85C9D3CFF9BFAF48B79FA718850" box="[1319,1436,1887,1912]" country="Argentina" name="Rio Negro" pageId="12" pageNumber="267">Río Negro</collectingRegion>
provinces,
<collectingCountry id="29CD762E9D3CFF9BFEC18BA5FE6C88B4" box="[274,385,1923,1948]" name="Argentina" pageId="12" pageNumber="267">Argentina</collectingCountry>
(
<figureCitation id="C9E12A3B9D3CFF9BFE438BA5FE0988B4" box="[400,484,1923,1948]" captionStart="FIGURE 35" captionStartId="13.[151,250,1234,1256]" captionTargetBox="[267,1325,252,1118]" captionTargetId="figure@13.[205,1396,196,1194]" captionTargetPageId="13" captionText="FIGURE 35. Map of Argentine and Chilean Patagonia with distribution records of: Austrohelea shannoni (Wirth &amp; Blanton), green circles. Austrohelea sirii n. sp., yellow triangles. Austrohelea spinosa n. sp., red diamonds." httpUri="https://zenodo.org/record/806131/files/figure.png" pageId="12" pageNumber="267">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="516536BE9D3CFF9BFF148B80FCE488CC" blockId="12.[151,1436,1886,2020]" pageId="12" pageNumber="267">
<emphasis id="63AEEAAC9D3CFF9BFF148B80FDB78897" bold="true" box="[199,602,1958,1983]" pageId="12" pageNumber="267">Derivation of the specific epithet.</emphasis>
The specific name of this new species is Latin, in reference to the long, coarse, subapical spine-like setae of the male hind tibia.
</paragraph>
</subSubSection>
<caption id="05A566369D3DFF9AFF4488F4FB9C8A2F" httpUri="https://zenodo.org/record/806131/files/figure.png" pageId="13" pageNumber="268" targetBox="[267,1325,252,1118]" targetPageId="13">
<paragraph id="516536BE9D3DFF9AFF4488F4FB9C8A2F" blockId="13.[151,1436,1234,1287]" pageId="13" pageNumber="268">
<emphasis id="63AEEAAC9D3DFF9AFF4488F4FECD8BCF" bold="true" box="[151,288,1234,1256]" pageId="13" pageNumber="268">FIGURE 35.</emphasis>
Map of Argentine and Chilean Patagonia with distribution records of:
<taxonomicName id="96DA4D3D9D3DFF9AFC2788F4FA758BC0" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[1012,1432,1234,1256]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3DFF9AFC2788F4FB3F8BC0" box="[1012,1234,1234,1256]" italics="true" pageId="13" pageNumber="268">Austrohelea shannoni</emphasis>
(Wirth &amp; Blanton)
</taxonomicName>
, green circles.
<taxonomicName id="96DA4D3D9D3DFF9AFEF788D7FE208A2F" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[292,461,1265,1287]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D3DFF9AFEF788D7FE208A2F" box="[292,461,1265,1287]" italics="true" pageId="13" pageNumber="268">Austrohelea sirii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3DFF9AFE0088D7FDE18A2E" box="[467,524,1265,1286]" pageId="13" pageNumber="268" rank="species">
<emphasis id="63AEEAAC9D3DFF9AFE0088D7FDE18A2E" bold="true" box="[467,524,1265,1286]" pageId="13" pageNumber="268">n. sp.</emphasis>
</taxonomicNameLabel>
, yellow triangles.
<taxonomicName id="96DA4D3D9D3DFF9AFD1488D7FC798A2F" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[711,916,1265,1287]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D3DFF9AFD1488D7FC798A2F" box="[711,916,1265,1287]" italics="true" pageId="13" pageNumber="268">Austrohelea spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3DFF9AFC4888D7FC398A2E" box="[923,980,1265,1286]" pageId="13" pageNumber="268" rank="species">
<emphasis id="63AEEAAC9D3DFF9AFC4888D7FC398A2E" bold="true" box="[923,980,1265,1286]" pageId="13" pageNumber="268">n. sp.</emphasis>
</taxonomicNameLabel>
, red diamonds.
</paragraph>
</caption>
<subSubSection id="19C065359D3DFF9AFF148915FCC38960" pageId="13" pageNumber="268" type="discussion">
<paragraph id="516536BE9D3DFF9AFF148915FE428928" blockId="13.[151,1436,1331,1608]" pageId="13" pageNumber="268">
<emphasis id="63AEEAAC9D3DFF9AFF148915FEA18A64" bold="true" box="[199,332,1331,1356]" pageId="13" pageNumber="268">Discussion.</emphasis>
Males of
<taxonomicName id="96DA4D3D9D3DFF9AFE188912FDA98A63" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[459,580,1332,1355]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="63AEEAAC9D3DFF9AFE188912FE318A63" box="[459,476,1332,1355]" italics="true" pageId="13" pageNumber="268">A</emphasis>
.
<emphasis id="63AEEAAC9D3DFF9AFE3C8912FDA98A63" box="[495,580,1332,1355]" italics="true" pageId="13" pageNumber="268">spinosa</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="96DA4D3D9D3DFF9AFC768912FBDD8A64" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[933,1072,1332,1356]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3DFF9AFC768912FC5B8A63" box="[933,950,1332,1355]" italics="true" pageId="13" pageNumber="268">A</emphasis>
.
<emphasis id="63AEEAAC9D3DFF9AFC1A8912FBDD8A64" box="[969,1072,1332,1356]" italics="true" pageId="13" pageNumber="268">shannoni</emphasis>
</taxonomicName>
as they have similar genitalia, however; this new species differs from this previously described congener by the very long stout, black, subapical spines on the hind tibia of males; the posterior margin of tergite 9 is straight and the apicolateral processes are slender and much longer; the gonostylus is longer than the gonocoxite; the parameres are very broad and their basal apodemes form a shallow obtuse angle between the anterior and posterior arms; and the basal arch of the aedeagus is broader and U-shaped.
</paragraph>
<paragraph id="516536BE9D3DFF9AFF148A2DFCC38960" blockId="13.[151,1436,1331,1608]" pageId="13" pageNumber="268">
Females of this new species can be distinguished from those of
<taxonomicName id="96DA4D3D9D3DFF9AFC648A2AFBAD890C" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[951,1088,1548,1572]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3DFF9AFC648A2AFC25890B" box="[951,968,1548,1571]" italics="true" pageId="13" pageNumber="268">A</emphasis>
.
<emphasis id="63AEEAAC9D3DFF9AFC098A2AFBAD890C" box="[986,1088,1548,1572]" italics="true" pageId="13" pageNumber="268">shannoni</emphasis>
</taxonomicName>
by their greatly swollen hind femora and subequal size spermathecae with shorter necks.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName LSID="http://zoobank.org/8781E88B-CC9F-46AB-A12D-757917CF1418" authority="Sharkey" authorityName="Sharkey" authorityYear="2021" class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes mikeiviei" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mikeiviei" status="sp. nov.">Aleiodes mikeiviei Sharkey</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 400" captionStartId="F400" captionText="Figure 400. Aleiodes mikeiviei, holotype." figureDoi="10.3897/zookeys.1013.55600.figure400" httpUri="https://binary.pensoft.net/fig/503855" pageId="0" pageNumber="1">Figure 400</figureCitation>
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<subSubSection pageId="0" pageNumber="1" type="diagnostics">
<paragraph pageId="0" pageNumber="1">Diagnostics.</paragraph>
<paragraph pageId="0" pageNumber="1">BOLD:ACS9594. Consensus barcode. AGTTTTATATTTTATGTTTGGGATATGAGCTGGTATGATTGGGCTATCAATAAGATTAATTATTCGATTAGAATTAAGAATTAG---AGGGAGAATTTTAAAGAATGATCAAATTTATAATGGTATAGTTACATTACATGCTTTTGTAATAATTTTTTTTATAGTTATACCTATTATAATTGGAGGGTTTGGAAATTGATTGGTACCATTAATATTAGGGGCCCCTGATATAGCTTTTCCACGAATGAATAATATAAGGTTTTGATTATTAATTCCATCTTTTTTTTTATTATTAATTAGGGGGGTTATTAATGCAGGAGTTGGGACTGGATGGACAATGTATCCTCCTTTATCTTTATTAATTGGTCATAATGGTATTTCAGTTGATATATCTATTTTTTCTTTACATTTAGCTGGGGCTTCTTCAATTATAGGGGCTATTAATTTTATCACTACAATTTTTAATATAAAGTTAAAAGGGATTAAGTTAGATCAAATTTCTTTATTAATTTGATCTATTTTAATTACTACAATTTTGTTATTACTTTCATTACCTGTTTTAGCTGGTGCTATTACTATATTATTGACTGATCGTAATTTAAATACTAGATTTTTTGATTTTTCAGGGGGAGGAGACCCAG--------------------.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype ♀">
<paragraph pageId="0" pageNumber="1">Holotype ♀.</paragraph>
<paragraph pageId="0" pageNumber="1">
Guanacaste, Sector Pitilla, Bullas,
<geoCoordinate degrees="10.98670" direction="north" orientation="latitude" precision="1" value="10.9867">10.98670</geoCoordinate>
,
<geoCoordinate degrees="85.38503" direction="west" orientation="longitude" precision="1" value="-85.38503">-85.38503</geoCoordinate>
, 440 meters, caterpillar collection date: 31/vii/2014, wasp eclosion date: 24/viii/2014. Depository: CNC.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Host data</emphasis>
.
</emphasis>
arctJanzen01 Janzen71434 (
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Erebidae</taxonomicName>
) feeding on algae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Host caterpillar and holotype wasp voucher codes</emphasis>
.
</emphasis>
14-SRNP-71445, DHJPAR0056334.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="paratypes">
<paragraph pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="1">None.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharkey" authorityYear="2021" class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes mikeiviei" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mikeiviei">
<emphasis italics="true" pageId="0" pageNumber="1">Aleiodes mikeiviei</emphasis>
</taxonomicName>
is named in honor of Mike
<normalizedToken originalValue="Ivie’s">Ivie's</normalizedToken>
long-appreciated contributions to publicity for ACG, GDFCF, and now, BioAlfa.
</paragraph>
<caption doi="10.3897/zookeys.1013.55600.figure400" httpUri="https://binary.pensoft.net/fig/503855" pageId="0" pageNumber="1" start="Figure 400" startId="F400">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 400.</emphasis>
<taxonomicName authorityName="Sharkey" authorityYear="2021" class="Insecta" family="Braconidae" genus="Aleiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleiodes mikeiviei" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mikeiviei">
<emphasis italics="true" pageId="0" pageNumber="1">Aleiodes mikeiviei</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

50
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12.
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,
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<paragraph id="7E1372B229C1635C55C90891006A16E6">Wien. Zool. Bot. Ges. 1862,721.</paragraph>
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<paragraph id="47654076FF72CD04477FACC4253DD657">Sciotel, (Beccari); [[ worker ]].</paragraph>
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