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<document id="7B8F2FD0577D93B0E02CAA54FF6B3981" ID-CLB-Dataset="298715" ID-DOI="10.11646/zootaxa.5471.2.1" ID-GBIF-Dataset="3a469638-dc21-45a3-80f6-cb307fbf5941" ID-ISSN="1175-5326" ID-Zenodo-Dep="12189904" ID-ZooBank="BAA047F4-2D65-4E52-8798-D426E3B26DA1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1718892848818" checkinUser="plazi" docAuthor="Kosterin, Oleg E., Onishko, Vladimir V., Ilyina, Elena V., Chepurnov, Grigory Yu. &amp; Blinov, Alexander G." docDate="2024" docId="DB0F87E19102F10AA7D3F8F47E277CEF" docLanguage="en" docName="zootaxa.5471.2.1.pdf" docOrigin="Zootaxa 5471 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenagrion pulchellum" docType="treatment" docVersion="5" lastPageNumber="171" masterDocId="2736FF999111F11EA744F92E7A657E46" masterDocTitle="The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus" masterLastPageNumber="190" masterPageNumber="151" pageNumber="170" updateTime="1718988639371" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="07F9884BC6142DFAE7E5BFA50B6FC95E">The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus</mods:title>
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<mods:namePart id="FEB28BE3E5E129B33B7D30A0F13C65D1">Kosterin, Oleg E.</mods:namePart>
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<mods:affiliation id="8FCD7FE9D1F220272A7159AB760DFFEA">Institute of Cytology &amp; Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:namePart id="B4BDB82E672669BE249DB94CBC288A46">Onishko, Vladimir V.</mods:namePart>
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<mods:affiliation id="CE2F072B00157FF5EE2E781AB0D33AA2">Moscow Zoo, Department of Herpetology, Bolshaya Gruzinskaya Str. 1, Moscow 123242 Russia.</mods:affiliation>
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<mods:namePart id="21FA079656E6FB062F51C4AA44585898">Ilyina, Elena V.</mods:namePart>
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<mods:affiliation id="05A5B2195FBE909D40773DD306DC8DD4">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, Dagestanskiy State Nature Reserve, Gagarina Street 120, Makhachkala, Republic of Dagestan, Russia.</mods:affiliation>
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<mods:namePart id="CEEA1E8A7A5551493359917BA42519A3">Chepurnov, Grigory Yu.</mods:namePart>
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<mods:affiliation id="E6B9D018B67938BDD61A0ABD3044274C">Institute of Cytology &amp; Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. &amp; Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:namePart id="31EFC7831D44626A13C5013F07271529">Blinov, Alexander G.</mods:namePart>
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<mods:affiliation id="FBDA4B80815ED4171423295C52CA2D9D">Institute of Cytology &amp; Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. &amp; Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<treatment id="DB0F87E19102F10AA7D3F8F47E277CEF" ID-DOI="http://doi.org/10.5281/zenodo.12190721" ID-GBIF-Taxon="233309489" ID-Zenodo-Dep="12190721" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF" httpUri="http://treatment.plazi.org/id/DB0F87E19102F10AA7D3F8F47E277CEF" lastPageId="20" lastPageNumber="171" pageId="19" pageNumber="170">
<subSubSection id="1BBC657C9102F10DA7D3F8F478917FB2" box="[151,756,474,501]" pageId="19" pageNumber="170" type="nomenclature">
<paragraph id="531936F79102F10DA7D3F8F478917FB2" blockId="19.[151,756,474,537]" box="[151,756,474,501]" pageId="19" pageNumber="170">
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<emphasis id="61D2EAE59102F10DA7D3F8F478917FB2" bold="true" box="[151,756,474,501]" pageId="19" pageNumber="170">
3.
<taxonomicName id="94A64D749102F10DA7F1F8F578917FB2" ID-CoL="WTXZ" authority="(Vander Linden, 1825)" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[181,756,474,501]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59102F10DA7F1F8F57BB17FB3" bold="true" box="[181,468,474,501]" italics="true" pageId="19" pageNumber="170">Coenagrion pulchellum</emphasis>
(Vander Linden, 1825)
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</emphasis>
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<figureCitation id="CB9D2A729102F10DA7D3F8D17B6C7C5F" box="[151,265,511,537]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="19" pageNumber="170">Figs 2o–p</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA650F8D17B2A7C5F" box="[276,335,511,537]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="19" pageNumber="170">3o–p</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA61EF8D17BEA7C5F" box="[346,399,511,537]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="19" pageNumber="170">4e–f</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA6DEF8D07BB17C5F" box="[410,468,510,537]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="19" pageNumber="170">5b–c</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA69BF8D17B937C5F" box="[479,502,511,537]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="19" pageNumber="170">7j</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA546F8D178247C5F" box="[514,577,511,537]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">8e–m</figureCitation>
</paragraph>
<paragraph id="531936F79102F10DA7D3FB697E107CB7" blockId="19.[151,1437,582,2013]" pageId="19" pageNumber="170">
<emphasis id="61D2EAE59102F10DA7D3FB697B367C26" bold="true" box="[151,339,582,609]" pageId="19" pageNumber="170">Reported from:</emphasis>
throughout Krasnodarskiy Kray,
<collectingRegion id="9162F8159102F10DA58BFB6979437C27" box="[719,806,583,609]" country="Russia" name="Adygeya" pageId="19" pageNumber="170">Adygea</collectingRegion>
and
<collectingRegion id="9162F8159102F10DA41BFB697E247C27" box="[863,1089,583,609]" country="Russia" name="Kabardino-Balkariya" pageId="19" pageNumber="170">Kabardino-Balkaria</collectingRegion>
; scarce unprecise reports from Stavropol’skiy Kray, Karachay-Cherkressia and
<collectingRegion id="9162F8159102F10DA58DFB4579087CC3" box="[713,877,619,645]" country="Russia" name="North Ossetia" pageId="19" pageNumber="170">North Ossetia</collectingRegion>
; not reported from
<collectingRegion id="9162F8159102F10DA31DFB457EAC7CC3" box="[1113,1225,619,645]" country="Russia" name="Chechnya" pageId="19" pageNumber="170">Chechnya</collectingRegion>
and
<collectingRegion id="9162F8159102F10DA24EFB457F107CC3" box="[1290,1397,619,645]" country="Russia" name="Dagestan" pageId="19" pageNumber="170">Dagestan</collectingRegion>
, as summarised by
<bibRefCitation id="37374B069102F10DA608FBA1786E7CEF" author="Skvortsov, V. E." box="[332,523,655,681]" pageId="19" pageNumber="170" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov (2010</bibRefCitation>
; for
<collectingRegion id="9162F8159102F10DA504FBA178F27CEF" box="[576,663,655,681]" country="Russia" name="Adygeya" pageId="19" pageNumber="170">Adygea</collectingRegion>
see also
<bibRefCitation id="37374B069102F10DA446FBA17E3B7CEF" author="Shapovalov, M. I. &amp; Korotkov, E. A." box="[770,1118,655,681]" pageId="19" pageNumber="170" pagination="341 - 349" refId="ref29460" refString="Shapovalov, M. I. &amp; Korotkov, E. A. (2019) Update of the Odonata fauna of Georgia, southern Caucasus ecoregion. Russian Entomological Journal, 28 (4), 341 - 349. https: // doi. org / 10.15298 / rusentj. 28.4.01" type="journal article" year="2019">Shapovalov &amp; Korotkov 2019</bibRefCitation>
). Kamysh-Burun, for which the species was reported in the paper on dragonflies of
<collectingRegion id="9162F8159102F10DA460FB9D79E97C8B" box="[804,908,691,717]" country="Russia" name="Dagestan" pageId="19" pageNumber="170">Dagestan</collectingRegion>
by
<bibRefCitation id="37374B069102F10DA4F8FB9D7ECB7C8B" author="Artobolevskij, G. V." box="[956,1198,691,717]" pageId="19" pageNumber="170" pagination="225 - 240" refId="ref27170" refString="Artobolevskij, G. V. (1929) Les Odonates du Daghestan. Revue Russe d'Entomologie, 23 (3 - 4), 225 - 240. [in Russian, with French title]" type="journal article" year="1929">Artobolevskij (1929)</bibRefCitation>
, was in
<collectingRegion id="9162F8159102F10DA250FB9D7F197C8B" box="[1300,1404,691,717]" country="Russia" name="Dagestan" pageId="19" pageNumber="170">Dagestan</collectingRegion>
in Artobolevskij’s time but presently is a part of Neftekumsk Town in Stavropol’skiy Kray.
</paragraph>
<paragraph id="531936F79102F10DA783FBD47B267B13" blockId="19.[151,1437,582,2013]" pageId="19" pageNumber="170">
<emphasis id="61D2EAE59102F10DA783FBD478397D53" bold="true" box="[199,604,762,789]" pageId="19" pageNumber="170">
Specimens.
<materialsCitation id="E3CE3CAA9102F10DA618FBD47B7C7D1B" ID-GBIF-Occurrence="4897992318" collectingDate="2019-06-24" elevation="5" latitude="37.261" location="Dzhiga River" longLatPrecision="70" longitude="37.262" municipality="Anapa Municipality" pageId="19" pageNumber="170" specimenCount="1" specimenCount-female="1">
Krasnodarskiy Kray:
<specimenCount id="45A0FD7E9102F10DA537FBD578ED7D53" box="[627,648,763,789]" count="1" pageId="19" pageNumber="170" type="female">♀</specimenCount>
(dark morph),
<collectingMunicipality id="B37DAC8D9102F10DA40DFBD57E517D53" box="[841,1076,763,789]" pageId="19" pageNumber="170">Anapa Municipality</collectingMunicipality>
, the
<location id="5679602C9102F10DA33CFBD57F737D53" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA33CFBD57F737D53" box="[1144,1302,763,789]" latitude="37.261" longLatPrecision="70" longitude="37.262" municipality="Anapa Municipality" name="Dzhiga River" pageId="19" pageNumber="170">Dzhiga River</location>
lowermost reaches and mouth
<locationDeviation id="42AB2FD59102F10DA635FA3178867D7F" box="[369,739,799,825]" location="Suvorov-Cherkesskiy" pageId="19" pageNumber="170">1 km W of Suvorov-Cherkesskiy</locationDeviation>
settlement,
<geoCoordinate id="369250309102F10DA428FA3079DC7D7F" box="[876,953,798,825]" degrees="45.060" direction="north" orientation="latitude" pageId="19" pageNumber="170" precision="55" value="45.06">45.060</geoCoordinate>
–45.064 N, 37.261–
<geoCoordinate id="369250309102F10DA3DAFA317F6B7D7F" box="[1182,1294,799,825]" degrees="37.262" direction="east" orientation="longitude" pageId="19" pageNumber="170" precision="55" value="37.262">37.262 E</geoCoordinate>
,
<elevation id="D88BD1C49102F10DA25EFA317FF87D7F" box="[1306,1437,799,825]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="19" pageNumber="170" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<quantity id="945E9B129102F10DA25EFA317F047D7F" box="[1306,1377,799,825]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="19" pageNumber="170" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
a.s.l.
</elevation>
<date id="271810379102F10DA7D3FA6D7B7C7D1B" box="[151,281,835,861]" pageId="19" pageNumber="170" value="2019-06-24">
<collectingDate id="375CE9DF9102F10DA7D3FA6D7B7C7D1B" box="[151,281,835,861]" pageId="19" pageNumber="170" value="2019-06-24">24.06.2019</collectingDate>
</date>
</materialsCitation>
</emphasis>
.
<materialsCitation id="E3CE3CAA9102F10DA660FA6C7FFD7DC7" ID-GBIF-Occurrence="4897992353" collectingDate="2019-07-03" county="Zelenchuk District" elevation="1013" latitude="41.536" location="Karachay-Cherkes Republic" longLatPrecision="69" longitude="41.54" municipality="Dausuz village" pageId="19" pageNumber="170" specimenCount="1" specimenCount-female="1">
<emphasis id="61D2EAE59102F10DA660FA6C78E17D1A" bold="true" box="[292,644,834,860]" pageId="19" pageNumber="170">
<location id="5679602C9102F10DA660FA6C781A7D1A" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA660FA6C781A7D1A" box="[292,639,834,860]" name="Karachay-Cherkes Republic" pageId="19" pageNumber="170">Karachay-Cherkes Republic</location>
:
</emphasis>
<specimenCount id="45A0FD7E9102F10DA5C8FA6D78C47D1B" box="[652,673,835,861]" count="1" pageId="19" pageNumber="170" type="female">♀</specimenCount>
,
<collectingCounty id="BA784E7B9102F10DA5EBFA6D79E17D1B" box="[687,900,835,861]" pageId="19" pageNumber="170">Zelenchuk District</collectingCounty>
,
<collectingMunicipality id="B37DAC8D9102F10DA4D4FA6D7E5D7D1B" box="[912,1080,835,861]" pageId="19" pageNumber="170">Dausuz village</collectingMunicipality>
W environs, S banks of a large water reserve at the
<location id="5679602C9102F10DA637FA4978F17DC7" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA637FA4978F17DC7" box="[371,660,871,897]" county="Zelenchuk District" latitude="41.536" longLatPrecision="69" longitude="41.54" municipality="Dausuz village" name="Bolzhoy Zelenchuk River" pageId="19" pageNumber="170">Bolzhoy Zelenchuk River</location>
,
<geoCoordinate id="369250309102F10DA5D9FA49788C7DC7" box="[669,745,871,897]" degrees="43.793" direction="north" orientation="latitude" pageId="19" pageNumber="170" precision="55" value="43.793">43.793</geoCoordinate>
–43.796 N, 41.536–
<geoCoordinate id="369250309102F10DA48AFA487E5E7DC7" box="[974,1083,870,897]" degrees="41.540" direction="east" orientation="longitude" pageId="19" pageNumber="170" precision="55" value="41.54">41.540 E</geoCoordinate>
,
<elevation id="D88BD1C49102F10DA301FA497F7F7DC7" box="[1093,1306,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.013" metricValueMax="1.014" metricValueMin="1.012" pageId="19" pageNumber="170" unit="m" value="1013.0" valueMax="1014.0" valueMin="1012.0">
<quantity id="945E9B129102F10DA301FA497EBA7DC7" box="[1093,1247,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.013" metricValueMax="1.014" metricValueMin="1.012" pageId="19" pageNumber="170" unit="m" value="1013.0" valueMax="1014.0" valueMin="1012.0">1012–1014 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379102F10DA261FA497FFD7DC7" box="[1317,1432,871,897]" pageId="19" pageNumber="170" value="2019-07-03">
<collectingDate id="375CE9DF9102F10DA261FA497FFD7DC7" box="[1317,1432,871,897]" pageId="19" pageNumber="170" value="2019-07-03">3.07.2019</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="E3CE3CAA9102F10DA7D3FAA478C97DAB" ID-GBIF-Occurrence="4897992387" collectingDate="2023-05-30" county="Kumtorkala District" elevation="8" latitude="47.223" location="Republic" longLatPrecision="66" longitude="47.246" municipality="Shura-Ozen' River" pageId="19" pageNumber="170" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
<emphasis id="61D2EAE59102F10DA7D3FAA47BC07DE3" bold="true" box="[151,421,906,933]" pageId="19" pageNumber="170">
<collectingRegion id="9162F8159102F10DA7D3FAA47BC57DE3" box="[151,416,906,933]" country="Russia" name="Dagestan" pageId="19" pageNumber="170">Republic of Dagestan</collectingRegion>
:
</emphasis>
<specimenCount id="45A0FD7E9102F10DA6EAFAA57BA67DE3" box="[430,451,907,933]" count="1" pageId="19" pageNumber="170" type="male">♂</specimenCount>
(176989333, 181375745 (
<figureCitation id="CB9D2A729102F10DA5B0FAA579297DE3" box="[756,844,907,933]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig. 8m</figureCitation>
)—both refer to the only male specimen collected),
<collectingCounty id="BA784E7B9102F10DA7D3FA817B197D8F" box="[151,380,943,969]" pageId="19" pageNumber="170">Kumtorkala District</collectingCounty>
, the
<collectingMunicipality id="B37DAC8D9102F10DA6F7FA8178E37D8F" box="[435,646,943,969]" pageId="19" pageNumber="170">Shura-Ozen’ River</collectingMunicipality>
at
<location id="5679602C9102F10DA5EFFA8179EE7D8F" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA5EFFA8179EE7D8F" box="[683,907,943,969]" county="Kumtorkala District" latitude="47.223" longLatPrecision="66" longitude="47.246" municipality="Shura-Ozen' River" name="Kumtorkala Station" pageId="19" pageNumber="170" stateProvince="Dagestan">Kumtorkala Station</location>
, next to
<location id="5679602C9102F10DA4AAFA817EDB7D8F" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA4AAFA817EDB7D8F" box="[1006,1214,943,969]" county="Kumtorkala District" latitude="47.223" longLatPrecision="66" longitude="47.246" municipality="Shura-Ozen' River" name="Sarykum Barkhan" pageId="19" pageNumber="170" stateProvince="Dagestan">Sarykum Barkhan</location>
,
<geoCoordinate id="369250309102F10DA38EFA817F727D8F" box="[1226,1303,943,969]" degrees="42.997" direction="north" orientation="latitude" pageId="19" pageNumber="170" precision="55" value="42.997">42.997</geoCoordinate>
–43.006 N, 47.223–
<geoCoordinate id="369250309102F10DA7B6FAFD7B017DAB" box="[242,356,979,1005]" degrees="47.246" direction="east" orientation="longitude" pageId="19" pageNumber="170" precision="55" value="47.246">47.246 E</geoCoordinate>
,
<elevation id="D88BD1C49102F10DA637FAFC787F7DAB" box="[371,538,978,1005]" metricMagnitude="1" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="19" pageNumber="170" unit="m" value="72.5" valueMax="80.0" valueMin="65.0">
<quantity id="945E9B129102F10DA637FAFC7BBF7DAB" box="[371,474,978,1005]" metricMagnitude="1" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="19" pageNumber="170" unit="m" value="72.5" valueMax="80.0" valueMin="65.0">65–80 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379102F10DA56EFAFC78C97DAB" box="[554,684,978,1005]" pageId="19" pageNumber="170" value="2023-05-30">
<collectingDate id="375CE9DF9102F10DA56EFAFC78C97DAB" box="[554,684,978,1005]" pageId="19" pageNumber="170" value="2023-05-30">30.05.2023</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="E3CE3CAA9102F10DA5FEFAFD78287A73" ID-GBIF-Occurrence="4897992343" collectingDate="2021-06-01" collectorName="E. Ilyina" county="Makhachkala" elevation="26" latitude="47.491" location="Sulak River" longLatPrecision="66" longitude="47.497" municipality="Muraveynik Protected Area" pageId="19" pageNumber="170" specimenCount="1" specimenCount-female="1" stateProvince="Dagestan">
<specimenCount id="45A0FD7E9102F10DA5FEFAFD78AA7DAB" box="[698,719,979,1005]" count="1" pageId="19" pageNumber="170" type="male">♂</specimenCount>
(
<figureCitation id="CB9D2A729102F10DA5A5FAFD79527DAB" box="[737,823,979,1005]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="19" pageNumber="170">Figs 4f</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA403FAFC79007DAB" box="[839,869,978,1005]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="19" pageNumber="170">5b</figureCitation>
),
<specimenCount id="45A0FD7E9102F10DA439FAFD79F77DAB" box="[893,914,979,1005]" count="1" pageId="19" pageNumber="170" type="female">♀</specimenCount>
(
<figureCitation id="CB9D2A729102F10DA4E1FAFD79987DAB" box="[933,1021,979,1005]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="19" pageNumber="170">Fig. 7k</figureCitation>
),
<collectingCounty id="BA784E7B9102F10DA350FAFD7ECE7DAB" box="[1044,1195,979,1005]" pageId="19" pageNumber="170">Makhachkala</collectingCounty>
N env.,
<collectingMunicipality id="B37DAC8D9102F10DA257FAFD7B257A57" pageId="19" pageNumber="170">Muraveynik Protected Area</collectingMunicipality>
at the
<location id="5679602C9102F10DA6D1FAD978787A57" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA6D1FAD978787A57" box="[405,541,1015,1041]" county="Makhachkala" latitude="47.491" longLatPrecision="66" longitude="47.497" municipality="Muraveynik Protected Area" name="Sulak River" pageId="19" pageNumber="170" stateProvince="Dagestan">Sulak River</location>
mouth, pools among coastal dunes,
<geoCoordinate id="369250309102F10DA483FAD97E717A57" box="[967,1044,1015,1041]" degrees="43.204" direction="north" orientation="latitude" pageId="19" pageNumber="170" precision="55" value="43.204">43.204</geoCoordinate>
–43.211 N, 47.491–
<geoCoordinate id="369250309102F10DA3BBFAD97F147A57" box="[1279,1393,1015,1041]" degrees="47.497" direction="east" orientation="longitude" pageId="19" pageNumber="170" precision="55" value="47.497">47.497 E</geoCoordinate>
,
<quantity id="945E9B129102F10DA2C4FAD97AC87A73" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="170" unit="m" value="26.0">
<elevation id="D88BD1C49102F10DA2C4FAD97AC87A73" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="170" unit="m" value="26.0">26 m</elevation>
</quantity>
below s.l.,
<date id="271810379102F10DA671FD357BCD7A73" box="[309,424,1051,1077]" pageId="19" pageNumber="170" value="2021-06-01">
<collectingDate id="375CE9DF9102F10DA671FD357BCD7A73" box="[309,424,1051,1077]" pageId="19" pageNumber="170" value="2021-06-01">1.06.2021</collectingDate>
</date>
,
<collectorName id="FE5353219102F10DA6F1FD35787D7A73" box="[437,536,1051,1077]" pageId="19" pageNumber="170">E. Ilyina</collectorName>
leg.
</materialsCitation>
;
<specimenCount id="45A0FD7E9102F10DA51FFD3578157A73" box="[603,624,1051,1077]" count="1" pageId="19" pageNumber="170" type="female">♀</specimenCount>
, the same place,
<date id="271810379102F10DA47EFD3579C87A73" box="[826,941,1051,1077]" pageId="19" pageNumber="170" value="2021-06-08">8.06.2021</date>
, E. Ilyina leg.;
<materialsCitation id="E3CE3CAA9102F10DA325FD3579C67AAF" ID-GBIF-Occurrence="4897992371" collectingDate="2023-05-23" collectorName="O. Kosterin &amp; E. Ilyina" elevation="26" latitude="47.486" location="Krivaya Balka River" longLatPrecision="66" longitude="47.49" municipality="Makhachkala" pageId="19" pageNumber="170" specimenCount="8" specimenCount-female="8" stateProvince="Dagestan">
<specimenCount id="45A0FD7E9102F10DA325FD357EC77A73" box="[1121,1186,1051,1077]" count="4" pageId="19" pageNumber="170" type="male">4 ♂♂</specimenCount>
(
<figureCitation id="CB9D2A729102F10DA3F7FD357F697A73" box="[1203,1292,1051,1077]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="19" pageNumber="170">Figs 2o</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA25EFD357F5D7A73" box="[1306,1336,1051,1077]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="19" pageNumber="170">3o</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA203FD357F067A73" box="[1351,1379,1051,1077]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="19" pageNumber="170">4e</figureCitation>
,
<figureCitation id="CB9D2A729102F10DA235FD347FEB7A73" box="[1393,1422,1050,1077]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="19" pageNumber="170">5c</figureCitation>
),
<specimenCount id="45A0FD7E9102F10DA7D3FD117ABD7A1F" box="[151,216,1087,1113]" count="8" pageId="19" pageNumber="170" type="female">8 ♀♀</specimenCount>
(166632124, 166725873, 166726404, 166726653 (
<figureCitation id="CB9D2A729102F10DA461FD11791E7A1F" box="[805,891,1087,1113]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig. 8k</figureCitation>
), 166727248, 166727929, 166728450 (
<figureCitation id="CB9D2A729102F10DA27AFD117FEA7A1F" box="[1342,1423,1087,1113]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig. 8f</figureCitation>
), 166728814, 166728940, 166733552, 166733553 (
<figureCitation id="CB9D2A729102F10DA5F9FD4D79637A3B" box="[701,774,1123,1149]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig.8l</figureCitation>
), 166733810, 166734141, 166734669 (
<figureCitation id="CB9D2A729102F10DA3F1FD4D7F607A3B" box="[1205,1285,1123,1149]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig. 8g</figureCitation>
), 166734952, 166735309, 166735310, 166737122, 166832245, 166835373, 167024132, 167024135),
<collectingMunicipality id="B37DAC8D9102F10DA3C5FDA97F7D7AE7" box="[1153,1304,1159,1185]" pageId="19" pageNumber="170">Makhachkala</collectingMunicipality>
N suburbs, pools between dunes at the
<location id="5679602C9102F10DA69CFD8578A07A83" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA69CFD8578A07A83" box="[472,709,1195,1221]" latitude="47.486" longLatPrecision="66" longitude="47.49" municipality="Makhachkala" name="Krivaya Balka River" pageId="19" pageNumber="170" stateProvince="Dagestan">Krivaya Balka River</location>
left bank in its lowermost reaches,
<geoCoordinate id="369250309102F10DA320FD857ED47A83" box="[1124,1201,1195,1221]" degrees="43.199" direction="north" orientation="latitude" pageId="19" pageNumber="170" precision="55" value="43.199">43.199</geoCoordinate>
–43.200 N, 47.486–
<geoCoordinate id="369250309102F10DA7D3FDE17B627AAF" box="[151,263,1231,1257]" degrees="47.490" direction="east" orientation="longitude" pageId="19" pageNumber="170" precision="55" value="47.49">47.490 E</geoCoordinate>
,
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<elevation id="D88BD1C49102F10DA650FDE17B2A7AAF" box="[276,335,1231,1257]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="170" unit="m" value="26.0">26 m</elevation>
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below s.l.,
<date id="271810379102F10DA690FDE078307AAF" box="[468,597,1230,1257]" pageId="19" pageNumber="170" value="2023-05-23">
<collectingDate id="375CE9DF9102F10DA690FDE078307AAF" box="[468,597,1230,1257]" pageId="19" pageNumber="170" value="2023-05-23">23.05.2023</collectingDate>
</date>
,
<collectorName id="FE5353219102F10DA526FDE178837AAF" box="[610,742,1231,1257]" pageId="19" pageNumber="170">O. Kosterin</collectorName>
&amp;
<collectorName id="FE5353219102F10DA449FDE1790B7AAF" box="[781,878,1231,1257]" pageId="19" pageNumber="170">E. Ilyina</collectorName>
leg.
</materialsCitation>
;
<specimenCount id="45A0FD7E9102F10DA4F5FDE179AD7AAF" box="[945,968,1231,1257]" count="1" pageId="19" pageNumber="170" type="male">♂</specimenCount>
,
<specimenCount id="45A0FD7E9102F10DA491FDE1798F7AAF" box="[981,1002,1231,1257]" count="1" pageId="19" pageNumber="170" type="female">♀</specimenCount>
(176988555, 176988662, 176988774, 176988928, 176989100, 176989203, 178699404 (
<figureCitation id="CB9D2A729102F10DA58EFDDD797E7B4B" box="[714,795,1267,1293]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig. 8e</figureCitation>
), 178699406 (
<figureCitation id="CB9D2A729102F10DA4FBFDDD7E697B4B" box="[959,1036,1267,1293]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig. 8j</figureCitation>
), 178699407 (
<figureCitation id="CB9D2A729102F10DA3EBFDDD7F677B4B" box="[1199,1282,1267,1293]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="19" pageNumber="170">Fig. 8h</figureCitation>
), 178699412, 181376006, 181376072, 181381216, 181381373), the same place but a broader longitude range of
<geoCoordinate id="369250309102F10DA3B0FC397F247B77" box="[1268,1345,1303,1329]" degrees="47.486" direction="north" orientation="latitude" pageId="19" pageNumber="170" precision="55" value="47.486">47.486</geoCoordinate>
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–
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</geoCoordinate>
E,,
<date id="271810379102F10DA7F9FC147B5A7B13" box="[189,319,1338,1365]" pageId="19" pageNumber="170" value="2023-05-30">30.05.2023</date>
.
</paragraph>
<paragraph id="531936F79102F10DA783FC707F347B87" blockId="19.[151,1437,582,2013]" pageId="19" pageNumber="170">
<emphasis id="61D2EAE59102F10DA783FC7079A47B3F" bold="true" box="[199,961,1374,1401]" pageId="19" pageNumber="170">Additional photographic registrations. Krasnodarskiy Kray</emphasis>
:
<materialsCitation id="E3CE3CAA9102F10DA498FC717EEC7BDB" ID-GBIF-Occurrence="4897992364" collectingDate="2018-06-01" latitude="44.8" location="Sukko village" longLatPrecision="67" longitude="37.439" municipality="Sukko River" pageId="19" pageNumber="170" specimenCount="1" specimenCount-female="1">
<specimenCount id="45A0FD7E9102F10DA498FC7179947B3F" box="[988,1009,1375,1401]" count="1" pageId="19" pageNumber="170" type="female">♀</specimenCount>
(dark morph) (37520705), Anapa Municipality,
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E end, the
<collectingMunicipality id="B37DAC8D9102F10DA52DFCAD78997BDB" box="[617,764,1411,1437]" pageId="19" pageNumber="170">Sukko River</collectingMunicipality>
,
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,
<geoCoordinate id="369250309102F10DA4D6FCAD7E607BDB" box="[914,1029,1411,1437]" degrees="37.439" direction="east" orientation="longitude" pageId="19" pageNumber="170" precision="55" value="37.439">37.439 E</geoCoordinate>
,
<date id="271810379102F10DA352FCAD7EEC7BDB" box="[1046,1161,1411,1437]" pageId="19" pageNumber="170" value="2018-06-01">
<collectingDate id="375CE9DF9102F10DA352FCAD7EEC7BDB" box="[1046,1161,1411,1437]" pageId="19" pageNumber="170" value="2018-06-01">1.06.2018</collectingDate>
</date>
</materialsCitation>
.
<emphasis id="61D2EAE59102F10DA3DCFCAC7B677B86" bold="true" pageId="19" pageNumber="170">Kabardino-Balkarian Republic</emphasis>
:
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<specimenCount id="45A0FD7E9102F10DA655FC897B437B87" box="[273,294,1447,1473]" count="1" pageId="19" pageNumber="170" type="male">♂</specimenCount>
(37681836),
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<location id="5679602C9102F10DA6FAFC8978007B87" LSID="urn:lsid:plazi:treatment:DB0F87E19102F10AA7D3F8F47E277CEF:5679602C9102F10DA6FAFC8978007B87" box="[446,613,1447,1473]" latitude="43.654" longLatPrecision="68" longitude="44.1" name="Mayskiy Town" pageId="19" pageNumber="170">Mayskiy Town</location>
NE
</collectorName>
margin, an inundated quarry,
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,
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,
<date id="271810379102F10DA39DFC887F287B87" box="[1241,1357,1446,1473]" pageId="19" pageNumber="170" value="2019-05-01">
<collectingDate id="375CE9DF9102F10DA39DFC887F287B87" box="[1241,1357,1446,1473]" pageId="19" pageNumber="170" value="2019-05-01">1.05.2019</collectingDate>
</date>
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.
</paragraph>
<paragraph id="531936F79102F10DA783FCE478DD78FB" blockId="19.[151,1437,582,2013]" pageId="19" pageNumber="170">
<emphasis id="61D2EAE59102F10DA783FCE47B317BA2" bold="true" box="[199,340,1482,1508]" pageId="19" pageNumber="170">Characters.</emphasis>
Male cercus large: trunk very robust, in lateral view (
<figureCitation id="CB9D2A729102F10DA4F2FCE57E417BA3" box="[950,1060,1483,1509]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="19" pageNumber="170">Fig. 3o–q</figureCitation>
) with horizontal to subhorizontal upper surface and apex processed and terminated as an acute angle, but with a rounded tip; in dorsal view (
<figureCitation id="CB9D2A729102F10DA22BFCC17AB5786B" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="19" pageNumber="170">Fig. 2o–q</figureCitation>
) rounded with a very large bulge as a blunt rounded angle, with bulges of the opposite cerci contacting each other. Tubercle disposed right below trunk apex, well seen in lateral view (
<figureCitation id="CB9D2A729102F10DA49BFF197E287817" box="[991,1101,1591,1617]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="19" pageNumber="170">Fig. 3o–q</figureCitation>
) and protruding behind to the level of paraproct spine. Tooth disposed at ca 2/3 of trunk length (
<figureCitation id="CB9D2A729102F10DA428FF7579BC7833" box="[876,985,1627,1653]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="19" pageNumber="170">Fig. 2o–q</figureCitation>
) (while almost at middle in
<taxonomicName id="94A64D749102F10DA24BFF757FF37832" authorityName="Lohmann" authorityYear="1983" box="[1295,1430,1627,1653]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE59102F10DA24BFF757FF37832" box="[1295,1430,1627,1653]" italics="true" pageId="19" pageNumber="170">C. persicum</emphasis>
</taxonomicName>
, see
<figureCitation id="CB9D2A729102F10DA787FF517B7378DF" box="[195,278,1663,1689]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="19" pageNumber="170">Fig. 2n</figureCitation>
). Process long and straight, seen only in caudal view (
<figureCitation id="CB9D2A729102F10DA438FF51798378DF" box="[892,998,1663,1689]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="19" pageNumber="170">Fig. 4e–f</figureCitation>
), very narrowly triangular with a blunt apex; terminating at paraproct prominence level.
</paragraph>
<paragraph id="531936F79102F10DA783FFE97882790B" blockId="19.[151,1437,582,2013]" pageId="19" pageNumber="170">
Paparoct spine very thin, directed a bit more than 45° upwards, prominence long, protruding caudad not much less than spine, incision between them very deep and broad (
<figureCitation id="CB9D2A729102F10DA41FFFC579A87943" box="[859,973,1771,1797]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="19" pageNumber="170">Fig. 3o–q</figureCitation>
), broader than in
<taxonomicName id="94A64D749102F10DA3D9FFC57F4D7942" authorityName="Lohmann" authorityYear="1983" box="[1181,1320,1771,1797]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE59102F10DA3D9FFC57F4D7942" box="[1181,1320,1771,1797]" italics="true" pageId="19" pageNumber="170">C. persicum</emphasis>
</taxonomicName>
, in which the prominence is directed caudad (
<figureCitation id="CB9D2A729102F10DA569FE2178E5796F" box="[557,640,1807,1833]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="19" pageNumber="170">Fig. 3n</figureCitation>
) rather than ventrocaudad as in
<taxonomicName id="94A64D749102F10DA4B6FE217EF4796F" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1010,1169,1807,1833]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59102F10DA4B6FE217EF4796F" box="[1010,1169,1807,1833]" italics="true" pageId="19" pageNumber="170">C. pulchellum</emphasis>
</taxonomicName>
(Fig. 30o–q); the same difference is shown in
<bibRefCitation id="37374B069102F10DA6D2FE1D78F4790B" author="Schneider, T. &amp; Ikemeyer, D. &amp; Ferreira, S. &amp; Muller, O." box="[406,657,1843,1869]" pageId="19" pageNumber="170" pagination="561 - 573" refId="ref29181" refString="Schneider, T., Ikemeyer, D., Ferreira, S. &amp; Muller, O. (2016) Rediscovery and redescription of Coenagrion persicum (Lohmann 1993) with description of the female, and some notes on habitat selection (Odonata: Coenagrionidae). Zootaxa, 4103 (6), 561 - 573. https: // doi. org / 10.11646 / zootaxa. 4103.6.6" type="journal article" year="2016">
Schneider
<emphasis id="61D2EAE59102F10DA549FE1D7820790B" box="[525,581,1843,1869]" italics="true" pageId="19" pageNumber="170">et al.</emphasis>
(2016
</bibRefCitation>
: fig. 2).
</paragraph>
<paragraph id="531936F79102F10DA783FE797F0F7937" blockId="19.[151,1437,582,2013]" box="[199,1386,1878,1905]" pageId="19" pageNumber="170">
Male pronotum hind margin tripartite, with the three lobes separated by two deep depressions (
<figureCitation id="CB9D2A729102F10DA3AAFE797F387937" box="[1262,1373,1878,1905]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="19" pageNumber="170">Fig. 5b–c</figureCitation>
).
</paragraph>
<paragraph id="531936F79102F10DA783FE5578A6799B" blockId="19.[151,1437,582,2013]" pageId="19" pageNumber="170">
Female pronotum festoon-like, with three sharp lobes, a rounded triangular central and two broadly rounded lateral ones, separated by deep narrow skewed triangular depressions (
<figureCitation id="CB9D2A729102F10DA485FEB17E7579FF" box="[961,1040,1951,1977]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="19" pageNumber="170">Fig. 7j</figureCitation>
), the latter not as sinuous as in
<taxonomicName id="94A64D749102F10DA2C7FEB17A9B799A" authorityName="Lohmann" authorityYear="1983" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE59102F10DA2C7FEB17A9B799A" italics="true" pageId="19" pageNumber="170">C. persicum</emphasis>
</taxonomicName>
(
<figureCitation id="CB9D2A729102F10DA649FEED7B3C799B" box="[269,345,1987,2013]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="19" pageNumber="170">Fig. 7i</figureCitation>
, see also
<bibRefCitation id="37374B069102F10DA680FEED78D2799B" author="Schneider, T. &amp; Ikemeyer, D. &amp; Ferreira, S. &amp; Muller, O." box="[452,695,1987,2013]" pageId="19" pageNumber="170" pagination="561 - 573" refId="ref29181" refString="Schneider, T., Ikemeyer, D., Ferreira, S. &amp; Muller, O. (2016) Rediscovery and redescription of Coenagrion persicum (Lohmann 1993) with description of the female, and some notes on habitat selection (Odonata: Coenagrionidae). Zootaxa, 4103 (6), 561 - 573. https: // doi. org / 10.11646 / zootaxa. 4103.6.6" type="journal article" year="2016">
Schneider
<emphasis id="61D2EAE59102F10DA57FFEED7816799B" box="[571,627,1987,2013]" italics="true" pageId="19" pageNumber="170">et al.</emphasis>
2016
</bibRefCitation>
).
</paragraph>
<paragraph id="531936F79105F10AA783F9B978AC7F23" blockId="20.[151,1437,151,681]" pageId="20" pageNumber="171">
The Caucasian males of
<taxonomicName id="94A64D749105F10AA6A9F9B978EB7EF7" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[493,654,151,177]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59105F10AA6A9F9B978EB7EF7" box="[493,654,151,177]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
</taxonomicName>
represent more melanic versions of the maculation pattern of this very variable species (
<figureCitation id="CB9D2A729105F10AA6D1F995786D7E93" box="[405,520,187,213]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8j–m</figureCitation>
), like specimens from West Siberia which were once described as
<taxonomicName id="94A64D749105F10AA3BAF9957BDE7EBF" authority="Belyshev, 1964" authorityName="Belyshev" authorityYear="1964" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="pulchellum" subSpecies="sibiricum">
<emphasis id="61D2EAE59105F10AA3BAF9957B647EBF" italics="true" pageId="20" pageNumber="171">C. pulchellum sibiricum</emphasis>
<bibRefCitation id="37374B069105F10AA64FF9F17BDE7EBF" author="Belyshev, B. F." box="[267,443,223,249]" pageId="20" pageNumber="171" pagination="4 - 70" refId="ref27359" refString="Belyshev, B. F. (1964) [Dragonfly fauna of the upper Ob basin]. In: Skalon, V. N. (Ed.), [Dragonflies of Siberia]. Trudy Vostochno- Sibirskogo Filiala Sibirskogo Otdeleniya Akademii Nauk SSSR (Ser. Biol.), Nauka, Leningrad, 40, 4 - 70. [in Russian]" type="journal article" year="1964">Belyshev, 1964</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="37374B069105F10AA689F9F1781B7EBF" author="Belyshev, B. F." box="[461,638,223,249]" pageId="20" pageNumber="171" pagination="4 - 70" refId="ref27359" refString="Belyshev, B. F. (1964) [Dragonfly fauna of the upper Ob basin]. In: Skalon, V. N. (Ed.), [Dragonflies of Siberia]. Trudy Vostochno- Sibirskogo Filiala Sibirskogo Otdeleniya Akademii Nauk SSSR (Ser. Biol.), Nauka, Leningrad, 40, 4 - 70. [in Russian]" type="journal article" year="1964">Belyshev 1964</bibRefCitation>
). In males, the antehumeral blue stripe is entire (
<figureCitation id="CB9D2A729105F10AA3FAF9F17F4D7EBF" box="[1214,1320,223,249]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8j–k</figureCitation>
) or, more frequently, interrupted (
<figureCitation id="CB9D2A729105F10AA6DBF82D7B8E7F5B" box="[415,491,259,285]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8l</figureCitation>
). In the most melanic specimen from the Shura-Ozen’ River it is reduced to tiny blue dots in the fore synthorax part (
<figureCitation id="CB9D2A729105F10AA6B3F80978367F07" box="[503,595,295,321]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">
Fig.
<quantity id="945E9B129105F10AA568F80978367F07" box="[556,595,295,321]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="171" unit="m" value="8.0">8m</quantity>
</figureCitation>
). Male S9 with a posterior variable black bar, entire (
<figureCitation id="CB9D2A729105F10AA3ECF8097F727F07" box="[1192,1303,295,321]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8l–m</figureCitation>
) or more or less bipartite (
<figureCitation id="CB9D2A729105F10AA671F8657BEC7F23" box="[309,393,331,357]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8k</figureCitation>
); S10 dorsally mostly black.
</paragraph>
<paragraph id="531936F79105F10AA783F841795C7FB3" blockId="20.[151,1437,151,681]" pageId="20" pageNumber="171">
Female dimorphic for the dark (
<figureCitation id="CB9D2A729105F10AA56DF84178107FCF" box="[553,629,367,393]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8f</figureCitation>
) and spotty (
<figureCitation id="CB9D2A729105F10AA441F84179EE7FCF" box="[773,907,367,393]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8e–h, g</figureCitation>
) morphs as elsewhere in Europe (
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Dijkstra
<emphasis id="61D2EAE59105F10AA221F8417FF87FCF" box="[1381,1437,367,393]" italics="true" pageId="20" pageNumber="171">et al.</emphasis>
2020
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) but in both morphs, S8 is always blue (
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) or largely blue (
<figureCitation id="CB9D2A729105F10AA4E3F8BD7E487FEB" box="[935,1069,403,429]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8e, f–g</figureCitation>
), with some bipartite (
<figureCitation id="CB9D2A729105F10AA26CF8BD7FF17FEB" box="[1320,1428,403,429]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8e–g</figureCitation>
) or paired black (
<figureCitation id="CB9D2A729105F10AA616F8997BC57F97" box="[338,416,439,465]" captionStart="FIGURE 8" captionStartId="18.[151,250,1401,1425]" captionTargetBox="[189,1398,181,1377]" captionTargetId="figure-24@18.[189,1398,181,1377]" captionTargetPageId="18" captionText="FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189926" httpUri="https://zenodo.org/record/12189926/files/figure.png" pageId="20" pageNumber="171">Fig. 8f</figureCitation>
) mark(s) at most. We never met females with completely dark S8 dorsum, as depicted for both female morphs of this species by
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Dijkstra
<emphasis id="61D2EAE59105F10AA5ECF8F578857FB3" box="[680,736,475,501]" italics="true" pageId="20" pageNumber="171">et al.</emphasis>
(2020)
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.
</paragraph>
</subSubSection>
<subSubSection id="1BBC657C9105F10AA783F8D07E277CEF" pageId="20" pageNumber="171" type="distribution">
<paragraph id="531936F79105F10AA783F8D07E277CEF" blockId="20.[151,1437,151,681]" pageId="20" pageNumber="171">
<emphasis id="61D2EAE59105F10AA783F8D07B937C5E" bold="true" box="[199,502,510,537]" pageId="20" pageNumber="171">Distribution and habitat.</emphasis>
The species is seemingly widespread but rare in the Caucasus at low elevations, always found by solitary individuals with a notable exception of huge populations existing at pools between coastal sandy dunes in the northern environs of Makhachkala, the capital of
<collectingRegion id="9162F8159105F10AA43BFB69798E7C27" box="[895,1003,583,609]" country="Russia" name="Dagestan" pageId="20" pageNumber="171">Dagestan</collectingRegion>
. In the latter locality, the only congener,
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<emphasis id="61D2EAE59105F10AA7D3FB457B7C7CC3" box="[151,281,619,645]" italics="true" pageId="20" pageNumber="171">C. scitulum</emphasis>
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was also found as solitary individuals. Maybe more large coastal populations of
<taxonomicName id="94A64D749105F10AA3FBFB457F3B7CC3" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1215,1374,619,645]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59105F10AA3FBFB457F3B7CC3" box="[1215,1374,619,645]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
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exist elsewhere, as the Azov and Caspian coasts of the Caucasus remain poorly explored.
</paragraph>
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</treatment>
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4.
<taxonomicName id="94A64D749105F10AA7F1FBDE78EA7D4C" ID-CoL="WTXX" authority="(Bartenef, 1929)" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[181,655,752,778]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="ponticum">
<emphasis id="61D2EAE59105F10AA7F1FBDE7BD87D4C" bold="true" box="[181,445,752,778]" italics="true" pageId="20" pageNumber="171">Coenagrion ponticum</emphasis>
(
<bibRefCitation id="37374B069105F10AA68AFBDE78E37D4C" author="Bartenef, A. N." box="[462,646,752,778]" pageId="20" pageNumber="171" pagination="54 - 68" refId="ref27207" refString="Bartenef, A. N. (1929) Neue Arten und Varietaten der Odonata des West Kaukasus. Zoologischer Anzeiger, 85, 54 - 68." type="journal article" year="1929">Bartenef, 1929</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BBC657C9105F10BA7D3FA3A7F3779E5" lastPageId="21" lastPageNumber="172" pageId="20" pageNumber="171" type="description">
<paragraph id="531936F79105F10AA7D3FA3A787B7D68" blockId="20.[151,655,752,815]" box="[151,542,788,815]" pageId="20" pageNumber="171">
<figureCitation id="CB9D2A729105F10AA7D3FA3A7B6E7D68" box="[151,267,788,814]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Figs 2l–m</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA652FA3A7B377D68" box="[278,338,788,814]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">3l–m</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA61AFA3A7BF27D68" box="[350,407,788,814]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="20" pageNumber="171">4c–d</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA6E6FA3A7BBE7D69" box="[418,475,788,815]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="20" pageNumber="171">5d–e</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA6A2FA3A78607D68" box="[486,517,788,814]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="20" pageNumber="171">7h</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA554FA3A787B7D68" box="[528,542,788,814]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">9</figureCitation>
</paragraph>
<paragraph id="531936F79105F10AA7D3FA727B3C7DF8" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
<emphasis id="61D2EAE59105F10AA7D3FA727B627D30" bold="true" box="[151,263,860,886]" pageId="20" pageNumber="171">Reported</emphasis>
(as
<taxonomicName id="94A64D749105F10AA670FA727B957D30" authority="Kosterin &amp; Onishko &amp; Ilyina &amp; Chepurnov &amp; Blinov, 2024" authorityName="Kosterin &amp; Onishko &amp; Ilyina &amp; Chepurnov &amp; Blinov" authorityYear="2024" box="[308,496,860,886]" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="ponticum" status="sp. nov.">
<emphasis id="61D2EAE59105F10AA670FA727B957D30" box="[308,496,860,886]" italics="true" pageId="20" pageNumber="171">Agrion ponticum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AE1579E9105F10AA6B3FA7378577D30" box="[503,562,860,887]" pageId="20" pageNumber="171" rank="species">sp. n.</taxonomicNameLabel>
)
<emphasis id="61D2EAE59105F10AA505FA7278E07D30" bold="true" box="[577,645,860,886]" pageId="20" pageNumber="171">from:</emphasis>
just three locations in Krasnodarskiy Kray: the environs of Sochy (five localities), Tuapse and Adler (one locality in each), all by the species’ author, A.N. Bartenev (
<bibRefCitation id="37374B069105F10AA3E8FAAE7F347DDC" author="Bartenef, A. N." box="[1196,1361,896,922]" pageId="20" pageNumber="171" pagination="54 - 68" refId="ref27207" refString="Bartenef, A. N. (1929) Neue Arten und Varietaten der Odonata des West Kaukasus. Zoologischer Anzeiger, 85, 54 - 68." type="journal article" year="1929">Bartenef 1929</bibRefCitation>
;
<bibRefCitation id="37374B069105F10AA218FAAE7FF27DDC" author="Bartenef, A. N." box="[1372,1431,896,922]" pageId="20" pageNumber="171" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">1930</bibRefCitation>
;
<bibRefCitation id="37374B069105F10AA7D3FA8A7B287DF8" author="Skvortsov, V. E." box="[151,333,932,958]" pageId="20" pageNumber="171" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
).
</paragraph>
<paragraph id="531936F79105F10AA783FAE6795A7A34" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
<emphasis id="61D2EAE59105F10AA783FAE6782C7DA4" bold="true" box="[199,585,968,994]" pageId="20" pageNumber="171">
Specimens.
<materialsCitation id="E3CE3CAA9105F10AA616FAE67EF97A6C" ID-GBIF-Occurrence="4897992363" collectingDate="2019-06-27" elevation="101" location="Goryachiy Klyuch Municipality" municipality="Fanagoriyskoe village" pageId="20" pageNumber="171" specimenCount="3" specimenCount-female="3">
Krasnodarskiy Kray:
<specimenCount id="45A0FD7E9105F10AA513FAE678C67DA5" box="[599,675,968,995]" count="22" pageId="20" pageNumber="171" type="male">22 ♂♂</specimenCount>
(
<figureCitation id="CB9D2A729105F10AA5F5FAE679407DA4" box="[689,805,968,994]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Figs 2l–m</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA46BFAE6790E7DA4" box="[815,875,968,994]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">3l–m</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA432FAE679CA7DA4" box="[886,943,968,994]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="20" pageNumber="171">4c–d</figureCitation>
,
<figureCitation id="CB9D2A729105F10AA4FDFAE679947DA5" box="[953,1009,968,995]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="20" pageNumber="171">5d–e</figureCitation>
) (108924869, 108924870, 108924871 (
<figureCitation id="CB9D2A729105F10AA7DBFAC27A9D7A40" box="[159,248,1004,1030]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9a</figureCitation>
), 37680876 (
<figureCitation id="CB9D2A729105F10AA6DEFAC27B917A40" box="[410,500,1004,1030]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9b</figureCitation>
), 37680891),
<specimenCount id="45A0FD7E9105F10AA5D9FAC278867A41" box="[669,739,1004,1031]" count="3" pageId="20" pageNumber="171" type="female">3 ♀♀</specimenCount>
(
<figureCitation id="CB9D2A729105F10AA5BEFAC279367A40" box="[762,851,1004,1030]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="20" pageNumber="171">Fig. 7h</figureCitation>
),
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,
<collectingMunicipality id="B37DAC8D9105F10AA3B3FAC27A867A6C" pageId="20" pageNumber="171">Fanagoriyskoe village</collectingMunicipality>
NW environs, ponds, 44.515–516 N, 39.108–109 E,
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a.s.l.
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,
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;
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, the
<collectingMunicipality id="B37DAC8D9105F10AA61FFD1A7B887A08" box="[347,493,1076,1102]" pageId="20" pageNumber="171">Chepsi River</collectingMunicipality>
banks
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–44.469 N, 39.055–
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a.s.l.
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<collectingDate id="375CE9DF9105F10AA61BFD767B857A34" box="[351,480,1112,1138]" pageId="20" pageNumber="171" value="2019-06-27">27.06.2019</collectingDate>
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.
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&amp;
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leg.
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<paragraph id="531936F79105F10AA783FD5279497B28" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
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:
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1 teneral
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(37681071 (
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)),
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,
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1 teneral
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(37681035), the same place,
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(37680916, 37680932, 37680994, 37681023), the same place,
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centre, a pond,
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, 39.125-
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a.s.l.
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</materialsCitation>
; 2 teneral
<specimenCount id="45A0FD7E9105F10AA484FC23798F7B61" box="[960,1002,1293,1319]" count="1" pageId="20" pageNumber="171" type="male">♂♂</specimenCount>
(75178528, 75178636), 2 teneral
<specimenCount id="45A0FD7E9105F10AA236FC237FF97B61" box="[1394,1436,1293,1319]" count="1" pageId="20" pageNumber="171" type="female">♀♀</specimenCount>
(75178417, 75179228), the same place,
<date id="271810379105F10AA52AFC1E78957B0C" box="[622,752,1328,1354]" pageId="20" pageNumber="171" value="2021-04-23">23.04.2021</date>
;
<specimenCount id="45A0FD7E9105F10AA5BBFC1E79267B0D" box="[767,835,1328,1355]" count="3" pageId="20" pageNumber="171" type="female">3 ♀♀</specimenCount>
(75178494, 75178577, 75178395), the same place,
<date id="271810379105F10AA7D3FC7A7B7C7B28" box="[151,281,1364,1390]" pageId="20" pageNumber="171" value="2021-04-24">24.04.2021</date>
;
<specimenCount id="45A0FD7E9105F10AA660FC7A7B2B7B29" box="[292,334,1364,1391]" count="1" pageId="20" pageNumber="171" type="male">1 ♂</specimenCount>
(108928403), the same place,
<date id="271810379105F10AA5E2FC7A794D7B28" box="[678,808,1364,1390]" pageId="20" pageNumber="171" value="2019-06-27">27.06.2019</date>
.
</paragraph>
<paragraph id="531936F79105F10AA783FC567F437BD4" blockId="20.[151,1437,860,2038]" box="[199,1318,1400,1427]" pageId="20" pageNumber="171">
<emphasis id="61D2EAE59105F10AA783FC567B317BD4" bold="true" box="[199,340,1400,1426]" pageId="20" pageNumber="171">Characters.</emphasis>
The male appendages resemble those of
<taxonomicName id="94A64D749105F10AA465FC5679DB7BD4" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[801,958,1400,1426]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59105F10AA465FC5679DB7BD4" box="[801,958,1400,1426]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
</taxonomicName>
, with the following differences:
</paragraph>
<paragraph id="531936F79105F10AA783FCB27E6B782C" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
Male cercus trunk much shorter and less processed, in lateral view (
<figureCitation id="CB9D2A729105F10AA482FCB27E537BF0" box="[966,1078,1436,1462]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">Fig. 3l–m</figureCitation>
) not raised, with slanting upper surface and blunter apex, so that tubercle is entirely seen in dorsal view (and is remarkably narrow); trunk in dorsal view (
<figureCitation id="CB9D2A729105F10AA7A4FCCA7B377BB8" box="[224,338,1508,1534]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2l–m</figureCitation>
) somewhat expanding posteriorly while of almost an even width in
<taxonomicName id="94A64D749105F10AA32EFCCA7F6C7BB8" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1130,1289,1508,1534]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59105F10AA32EFCCA7F6C7BB8" box="[1130,1289,1508,1534]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
</taxonomicName>
(
<figureCitation id="CB9D2A729105F10AA258FCCA7FEA7BB8" box="[1308,1423,1508,1534]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2o–q</figureCitation>
); prominence well expressed but broadly rounded, those of the two cerci also contact each other (see in dorsal view,
<figureCitation id="CB9D2A729105F10AA7D3FF027B637800" box="[151,262,1580,1606]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2l–m</figureCitation>
) as in
<taxonomicName id="94A64D749105F10AA60BFF027B897800" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[335,492,1580,1606]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59105F10AA60BFF027B897800" box="[335,492,1580,1606]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
</taxonomicName>
(
<figureCitation id="CB9D2A729105F10AA6B8FF0278097800" box="[508,620,1580,1606]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2o–q</figureCitation>
). Process nearly straight as in
<taxonomicName id="94A64D749105F10AA4FBFF027E397800" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[959,1116,1580,1606]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59105F10AA4FBFF027E397800" box="[959,1116,1580,1606]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
</taxonomicName>
but slenderer and somewhat shorter (
<figureCitation id="CB9D2A729105F10AA7B0FF7E7B06782C" box="[244,355,1616,1642]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="20" pageNumber="171">Fig. 4c–d</figureCitation>
), slightly not reaching the level of the paraproct prominence.
</paragraph>
<paragraph id="531936F79105F10AA783FF5A7A847890" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
Male paraproct spine directed more behind than up and protrudes caudad much farer than tubercle; incision between spine and prominence as deep but not as broad, and prominence as attenuated as in
<taxonomicName id="94A64D749105F10AA3FDFFB67F3278F4" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1209,1367,1688,1714]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59105F10AA3FDFFB67F3278F4" box="[1209,1367,1688,1714]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
</taxonomicName>
. (
<figureCitation id="CB9D2A729105F10AA22BFFB67AB07890" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">Fig. 3l–m</figureCitation>
).
</paragraph>
<paragraph id="531936F79105F10AA783FFCE7BB07904" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
Male pronotum hind margin is generally as in
<taxonomicName id="94A64D749105F10AA596FFCE795C78BC" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[722,825,1760,1786]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE59105F10AA596FFCE795C78BC" box="[722,825,1760,1786]" italics="true" pageId="20" pageNumber="171">C. puella</emphasis>
</taxonomicName>
(
<figureCitation id="CB9D2A729105F10AA40DFFCE79D278BD" box="[841,951,1760,1787]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="20" pageNumber="171">Fig. 5d–e</figureCitation>
), female pronotum as well but with a more prominent, roundish triangular central lobe and very slightly deeper and somewhat asymmetrical incisions between it and lateral lobes (
<figureCitation id="CB9D2A729105F10AA631FE067BAC7904" box="[373,457,1832,1858]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="20" pageNumber="171">Fig. 7h</figureCitation>
).
</paragraph>
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The black maculation development in males varies greatly, including in the same population in Fanagoriyskoe village. It is best reflected by the relative middorsal length of the posterior black spot in S3, which varies, quite continuously, ca from 20 to 70% of the S3 length, usually ca 50% (
<figureCitation id="CB9D2A729105F10AA4EDFEBA799879E8" box="[937,1021,1940,1966]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9a</figureCitation>
). Along with its regular long lateral projections, as a rule broad in this species, it forms a cenral tooth (
<figureCitation id="CB9D2A729105F10AA43FFE9679AB7994" box="[891,974,1976,2002]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9b</figureCitation>
), in some specimens produced into short, but of variable length, middorsal line.
</paragraph>
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<paragraph id="531936F79104F10BA7D3FF9D78187955" blockId="21.[151,1436,1715,1811]" pageId="21" pageNumber="172">
<emphasis id="61D2EAE59104F10BA7D3FF9D7B77788D" bold="true" box="[151,274,1715,1739]" pageId="21" pageNumber="172">FIGURE 9.</emphasis>
Males (a–b) and female (c) of
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in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale.
</paragraph>
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Male S9 with a posterior black bar of variable shape, which usually has two (at sides, most frequent version) (
<figureCitation id="CB9D2A729104F10BA7DBFEA77B6979E5" box="[159,268,1929,1955]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="21" pageNumber="172">Fig. 9a, b</figureCitation>
), one (central) or three (all above mentioned) anterior projections. S10 black above, blue at sides.
</paragraph>
</subSubSection>
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<emphasis id="61D2EAE59104F10BA783FE837B917981" bold="true" box="[199,500,1965,1991]" pageId="21" pageNumber="172">Distribution and habitat.</emphasis>
In
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found only in Krasnodarskiy Kray, both at the SE part of the Black Sea coast as well as inland, behind the Main Caucasus Range, in Goryachiy Klyuch District. All records were at ponds situated below
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a.s.l. and surrounded by forest vegetation. The earliest registered date of imaginal emerging was
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. No other
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spp.
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were found at these localities. Outside the region considered, the species ranges along the Black Sea coast of
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and has been reported from
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, Northern
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(
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) and NE
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(east of
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) (
<bibRefCitation id="37374B069107F108A68DF82D78FE7F5B" author="Boudot, J. - P. &amp; Borisov, S. &amp; De Knijf, G. &amp; Van Grunsven, R. &amp; Schroter, A. &amp; Kalkman, V. J." box="[457,667,259,285]" pageId="22" pageNumber="173" pagination="3 - 248" refId="ref27573" refString="Boudot, J. - P., Borisov, S., De Knijf, G., Van Grunsven, R., Schroter, A. &amp; Kalkman, V. J. (2021) Atlas of the dragonflies and damselflies of West and Central Asia. Brachytron, 22, Supplement, 3 - 248." type="journal article" year="2021">Boudot et al. 2021</bibRefCitation>
)
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="61D2EAE59106F109A7D3F9B978A27EF7" bold="true" box="[151,711,151,178]" pageId="23" pageNumber="174">
<figureCitation id="CB9D2A729106F109A7D3F9B97AC27EF7" box="[151,167,151,177]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="23" pageNumber="174">6</figureCitation>
.
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<emphasis id="61D2EAE59106F109A7F1F9B97BAF7EF4" bold="true" box="[181,458,151,178]" italics="true" pageId="23" pageNumber="174">Coenagrion lunulatum</emphasis>
(
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)
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<paragraph id="531936F79106F109A7D3F9F47F5B7F5E" blockId="23.[151,1437,217,496]" pageId="23" pageNumber="174">
<emphasis id="61D2EAE59106F109A7D3F9F47B627EB2" bold="true" box="[151,263,218,244]" pageId="23" pageNumber="174">Reported</emphasis>
(as
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<emphasis id="61D2EAE59106F109A67CF9F47B867EB2" box="[312,483,218,244]" italics="true" pageId="23" pageNumber="174">Agrion vernale</emphasis>
Hag.
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)
<emphasis id="61D2EAE59106F109A576F9F778137EB5" bold="true" box="[562,630,217,243]" pageId="23" pageNumber="174">from:</emphasis>
Krasnodarskiy Kray, Sochi Municipality, “Achischgho Mt, lake No 6” (one of the Khmelevskie Lakes, for explanation see
<bibRefCitation id="37374B069106F109A59BF9D079FE7F5E" author="Kosterin, O. E." box="[735,923,254,280]" pageId="23" pageNumber="174" pagination="89 - 126" refId="ref28414" refString="Kosterin, O. E. (2023) Reconsideration of the three Odonata taxa described by A. N. Bartenev from the same place in West Caucasus. Odonatologica, 52 (1 / 2), 89 - 126. https: // doi. org / 10.60024 / odon. v 52 i 1 - 2. a 7" type="journal article" year="2023">Kosterin (2023))</bibRefCitation>
(
<bibRefCitation id="37374B069106F109A4EDF9D07E1D7F5E" author="Bartenef, A. N." box="[937,1144,254,280]" pageId="23" pageNumber="174" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">Bartenef 1930: 71</bibRefCitation>
;
<bibRefCitation id="37374B069106F109A3C0F9D07F537F5E" author="Skvortsov, V. E." box="[1156,1334,254,280]" pageId="23" pageNumber="174" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BBC657C9106F109A783F80F7A857FB6" pageId="23" pageNumber="174" type="distribution">
<paragraph id="531936F79106F109A783F80F7A857FB6" blockId="23.[151,1437,217,496]" pageId="23" pageNumber="174">
<emphasis id="61D2EAE59106F109A783F80F7B947F7D" bold="true" box="[199,497,289,316]" pageId="23" pageNumber="174">Distribution and habitat.</emphasis>
<specimenCount id="45A0FD7E9106F109A6B3F80C78147F7A" box="[503,625,290,316]" count="2" pageId="23" pageNumber="174" type="male">Two males</specimenCount>
were collected by A.N.
<bibRefCitation id="37374B069106F109A43BF80C7E547F7A" author="Bartenef, A. N." box="[895,1073,290,316]" pageId="23" pageNumber="174" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">Bartenef (1930)</bibRefCitation>
on
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at ‘lake No. 6’ of the Khmelevskie Lakes situated at
<quantity id="945E9B129106F109A561F86878E67F26" box="[549,643,326,353]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="23" pageNumber="174" unit="m" value="1875.0">1,875 m</quantity>
a.s.l. in the subalpine zone of West Caucasus (for explanations of the lakes see
<bibRefCitation id="37374B069106F109A646F8447BC37FC2" author="Kosterin, O. E." box="[258,422,362,388]" pageId="23" pageNumber="174" pagination="89 - 126" refId="ref28414" refString="Kosterin, O. E. (2023) Reconsideration of the three Odonata taxa described by A. N. Bartenev from the same place in West Caucasus. Odonatologica, 52 (1 / 2), 89 - 126. https: // doi. org / 10.60024 / odon. v 52 i 1 - 2. a 7" type="journal article" year="2023">Kosterin 2023</bibRefCitation>
). That place was visited by OEK on
<date id="271810379106F109A47FF84479DF7FC2" box="[827,954,362,388]" pageId="23" pageNumber="174" value="2008-10-07">10.07.2008</date>
(
<bibRefCitation id="37374B069106F109A483F8447E0F7FC2" author="Kosterin, O. E." box="[967,1130,362,388]" pageId="23" pageNumber="174" pagination="89 - 126" refId="ref28414" refString="Kosterin, O. E. (2023) Reconsideration of the three Odonata taxa described by A. N. Bartenev from the same place in West Caucasus. Odonatologica, 52 (1 / 2), 89 - 126. https: // doi. org / 10.60024 / odon. v 52 i 1 - 2. a 7" type="journal article" year="2023">Kosterin 2023</bibRefCitation>
) but this species, as well as any its congener, was not found. No other data for this boreal species exist from the Russian part of the Caucasus, although several records exist from the eastern
<collectingCountry id="2BB176679106F109A5EAF89C789A7F8A" box="[686,767,434,460]" name="Turkey" pageId="23" pageNumber="174">Turkey</collectingCountry>
,
<collectingCountry id="2BB176679106F109A44FF89C790D7F8A" box="[779,872,434,460]" name="Georgia" pageId="23" pageNumber="174">Georgia</collectingCountry>
,
<collectingCountry id="2BB176679106F109A436F89C79B27F8A" box="[882,983,434,460]" name="Armenia" pageId="23" pageNumber="174">Armenia</collectingCountry>
,
<collectingCountry id="2BB176679106F109A4A6F89C7E387F8A" box="[994,1117,434,460]" name="Azerbaijan" pageId="23" pageNumber="174">Azerbaijan</collectingCountry>
and NW
<collectingCountry id="2BB176679106F109A38FF89C7E9D7F8A" box="[1227,1272,434,460]" name="Iran" pageId="23" pageNumber="174">Iran</collectingCountry>
(
<bibRefCitation id="37374B069106F109A243F89C7AB67FB6" author="Boudot, J. - P. &amp; Borisov, S. &amp; De Knijf, G. &amp; Van Grunsven, R. &amp; Schroter, A. &amp; Kalkman, V. J." pageId="23" pageNumber="174" pagination="3 - 248" refId="ref27573" refString="Boudot, J. - P., Borisov, S., De Knijf, G., Van Grunsven, R., Schroter, A. &amp; Kalkman, V. J. (2021) Atlas of the dragonflies and damselflies of West and Central Asia. Brachytron, 22, Supplement, 3 - 248." type="journal article" year="2021">
Boudot
<emphasis id="61D2EAE59106F109A227F89D7FF87F8A" box="[1379,1437,434,460]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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550
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7.
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<emphasis id="61D2EAE59106F109A7F1FB197BD67C14" bold="true" box="[181,435,567,594]" italics="true" pageId="23" pageNumber="174">Coenagrion scitulum</emphasis>
(
<bibRefCitation id="37374B069106F109A687FB1978127C17" author="Rambur, J. P." box="[451,631,567,593]" pageId="23" pageNumber="174" refId="ref29148" refString="Rambur, J. P. (1842) Histoire naturelle des insects: Nevropteres. Ouvrage accompagne de Planches. Nouvelles Suites a Buffon. Paris, Roret, 534 pp." type="book" year="1842">Rambur, 1842</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BBC657C9106F109A7D3FB757F5179C0" pageId="23" pageNumber="174" type="description">
<paragraph id="531936F79106F109A7D3FB757B6B7C33" blockId="23.[151,639,567,629]" box="[151,270,603,629]" pageId="23" pageNumber="174">
<figureCitation id="CB9D2A729106F109A7D3FB757B6B7C33" box="[151,270,603,629]" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="23" pageNumber="174">Fig. 11e–k</figureCitation>
</paragraph>
<paragraph id="531936F79106F109A7D3FB8D7EBE7D43" blockId="23.[151,1437,675,2034]" pageId="23" pageNumber="174">
<emphasis id="61D2EAE59106F109A7D3FB8D7B357CFB" bold="true" box="[151,336,675,701]" pageId="23" pageNumber="174">Reported from:</emphasis>
two localities in Krasnodarskiy Kray: the Pslukh River in Sochi Municipality (
<bibRefCitation id="37374B069106F109A3F0FB8D7F027CFB" author="Skvortsov, V. E." box="[1204,1383,675,701]" pageId="23" pageNumber="174" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
) and Lake Krugloe in Novorossiysk Municipality (
<bibRefCitation id="37374B069106F109A5DAFBE979B27CA7" author="Kosterin, O. E. &amp; Solovyev, V. I." box="[670,983,711,737]" pageId="23" pageNumber="174" pagination="1 - 43" refId="ref28515" refString="Kosterin, O. E. &amp; Solovyev, V. I. (2017) Odonata found in mid-summer 2015 and 2016 at the north-westernmost Black Sea Coast of the Caucasus, with the first record of Cordulegaster picta Selys, 1854 in Russian Federation. International Dragonfly Fund Report, 117, 1 - 43." type="journal article" year="2017">Kosterin &amp; Solovyev 2017</bibRefCitation>
); Voskhod Estate of Maykop in
<collectingRegion id="9162F8159106F109A201FBE97FF97CA7" box="[1349,1436,711,737]" country="Russia" name="Adygeya" pageId="23" pageNumber="174">Adygea</collectingRegion>
Republic (
<bibRefCitation id="37374B069106F109A64FFBC578017D43" author="Shapovalov, M. I. &amp; Korotkov, E. A." box="[267,612,747,773]" pageId="23" pageNumber="174" pagination="341 - 349" refId="ref29460" refString="Shapovalov, M. I. &amp; Korotkov, E. A. (2019) Update of the Odonata fauna of Georgia, southern Caucasus ecoregion. Russian Entomological Journal, 28 (4), 341 - 349. https: // doi. org / 10.15298 / rusentj. 28.4.01" type="journal article" year="2019">Shapovalov &amp; Korotkov 2019</bibRefCitation>
); unprecisely for Stavropol’skiy Kray (
<bibRefCitation id="37374B069106F109A35CFBC57EAA7D43" author="Skvortsov, V. E." box="[1048,1231,747,773]" pageId="23" pageNumber="174" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
).
</paragraph>
<paragraph id="531936F79106F109A783FA2179F87823" blockId="23.[151,1437,675,2034]" pageId="23" pageNumber="174">
<emphasis id="61D2EAE59106F109A783FA21783B7D6F" bold="true" box="[199,606,783,809]" pageId="23" pageNumber="174">
Specimens.
<materialsCitation id="E3CE3CAA9106F109A61CFA217B837D37" ID-GBIF-Occurrence="4897992341" collectingDate="2019-06-24" collectorName="Anapa Municipality, E" elevation="170" location="Lake Zmeinoe" municipality="Supsekh" pageId="23" pageNumber="174" specimenCount="2" specimenCount-male="2">
Krasnodarskiy Kray:
<specimenCount id="45A0FD7E9106F109A52EFA2178CB7D6C" box="[618,686,783,810]" count="2" pageId="23" pageNumber="174" type="male">2 ♂♂</specimenCount>
(37681939, 37681905, 42847761, 108801024, 108802758 (
<figureCitation id="CB9D2A729106F109A22BFA217AA57D0B" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="23" pageNumber="174">Fig. 11e</figureCitation>
)),
<collectorName id="FE5353219106F109A79BFA1D7BBA7D08" box="[223,479,819,846]" pageId="23" pageNumber="174">Anapa Municipality, E</collectorName>
suburbs of
<collectingMunicipality id="B37DAC8D9106F109A522FA1D78A07D0B" box="[614,709,819,845]" pageId="23" pageNumber="174">Supsekh</collectingMunicipality>
settlement,
<location id="5679602C9106F109A414FA1D79917D0B" LSID="urn:lsid:plazi:treatment:DB0F87E19106F109A7D3FB1978B779B7:5679602C9106F109A414FA1D79917D0B" box="[848,1012,819,845]" municipality="Supsekh" name="Lake Zmeinoe" pageId="23" pageNumber="174">Lake Zmeinoe</location>
env., 44.853–857 N, 37.391–393 E,
<elevation id="D88BD1C49106F109A7D3FA797B337D37" box="[151,342,855,882]" metricMagnitude="2" metricUnit="m" metricValue="1.6949999999999998" metricValueMax="1.89" metricValueMin="1.5" pageId="23" pageNumber="174" unit="m" value="169.5" valueMax="189.0" valueMin="150.0">
<quantity id="945E9B129106F109A7D3FA797B7D7D34" box="[151,280,855,882]" metricMagnitude="2" metricUnit="m" metricValue="1.6949999999999998" metricValueMax="1.89" metricValueMin="1.5" pageId="23" pageNumber="174" unit="m" value="169.5" valueMax="189.0" valueMin="150.0">150–189 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379106F109A620FA797B837D37" box="[356,486,855,881]" pageId="23" pageNumber="174" value="2019-06-24">
<collectingDate id="375CE9DF9106F109A620FA797B837D37" box="[356,486,855,881]" pageId="23" pageNumber="174" value="2019-06-24">24.06.2019</collectingDate>
</date>
</materialsCitation>
</emphasis>
;
<materialsCitation id="E3CE3CAA9106F109A6B7FA767E8A7DD3" ID-GBIF-Occurrence="4897992366" collectingDate="2016-10-07" collectorName="O. Kosterin" elevation="70" latitude="44.6768" location="Lake Krugloe" longLatPrecision="6" longitude="37.5973" municipality="Novorossiysk Municipality" pageId="23" pageNumber="174" specimenCount="1" specimenCount-male="1">
<specimenCount id="45A0FD7E9106F109A6B7FA76786E7D34" box="[499,523,856,882]" count="1" pageId="23" pageNumber="174" type="male">♂</specimenCount>
.
<collectingMunicipality id="B37DAC8D9106F109A55CFA7979297D37" box="[536,844,855,881]" pageId="23" pageNumber="174">Novorossiysk Municipality</collectingMunicipality>
,
<location id="5679602C9106F109A41DFA7979937D37" LSID="urn:lsid:plazi:treatment:DB0F87E19106F109A7D3FB1978B779B7:5679602C9106F109A41DFA7979937D37" box="[857,1014,855,881]" latitude="44.6768" longLatPrecision="6" longitude="37.5973" municipality="Novorossiysk Municipality" name="Lake Krugloe" pageId="23" pageNumber="174">Lake Krugloe</location>
(remnants of a small lake in a lower part of intermontane valley),
<geoCoordinate id="369250309106F109A6ABFA5578127DD0" box="[495,631,891,918]" degrees="44.6768" direction="north" orientation="latitude" pageId="23" pageNumber="174" precision="5" value="44.6768">44.6768° N</geoCoordinate>
,
<geoCoordinate id="369250309106F109A5C2FA55796F7DD0" box="[646,778,891,918]" degrees="37.5973" direction="east" orientation="longitude" pageId="23" pageNumber="174" precision="5" value="37.5973">37.5973° E</geoCoordinate>
,
<elevation id="D88BD1C49106F109A45EFA5579F27DD3" box="[794,919,891,918]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="174" unit="m" value="70.0">
<quantity id="945E9B129106F109A45EFA5579327DD0" box="[794,855,891,918]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="174" unit="m" value="70.0">70 m</quantity>
a.s.l.
</elevation>
<date id="271810379106F109A4E5FA557E477DD3" box="[929,1058,891,917]" pageId="23" pageNumber="174" value="2016-10-07">
<collectingDate id="375CE9DF9106F109A4E5FA557E477DD3" box="[929,1058,891,917]" pageId="23" pageNumber="174" value="2016-10-07">10.07.2016</collectingDate>
</date>
,
<collectorName id="FE5353219106F109A375FA557EDD7DD3" box="[1073,1208,891,917]" pageId="23" pageNumber="174">O. Kosterin</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="E3CE3CAA9106F109A3BBFA527B7C7D9B" ID-GBIF-Occurrence="4897992370" collectingDate="2019-06-27" elevation="101" location="Fanagoriyskoe village" municipality="Goryachiy Klyuch Municipality" pageId="23" pageNumber="174" specimenCount="1" specimenCount-male="1">
<specimenCount id="45A0FD7E9106F109A3BBFA527F737DD0" box="[1279,1302,892,918]" count="1" pageId="23" pageNumber="174" type="male">♂</specimenCount>
,
<collectingMunicipality id="B37DAC8D9106F109A262FA557BE77DFF" pageId="23" pageNumber="174">Goryachiy Klyuch Municipality</collectingMunicipality>
,
<location id="5679602C9106F109A6CFFAB178E77DFF" LSID="urn:lsid:plazi:treatment:DB0F87E19106F109A7D3FB1978B779B7:5679602C9106F109A6CFFAB178E77DFF" box="[395,642,927,953]" municipality="Goryachiy Klyuch Municipality" name="Fanagoriyskoe village" pageId="23" pageNumber="174">Fanagoriyskoe village</location>
NW environs, ponds, 44.515–516 N, 39.108–109 E,
<elevation id="D88BD1C49106F109A399FAB17FF27DFF" box="[1245,1431,927,954]" metricMagnitude="2" metricUnit="m" metricValue="1.015" metricValueMax="1.02" metricValueMin="1.01" pageId="23" pageNumber="174" unit="m" value="101.5" valueMax="102.0" valueMin="101.0">
<quantity id="945E9B129106F109A399FAB17F3E7DFC" box="[1245,1371,927,954]" metricMagnitude="2" metricUnit="m" metricValue="1.015" metricValueMax="1.02" metricValueMin="1.01" pageId="23" pageNumber="174" unit="m" value="101.5" valueMax="102.0" valueMin="101.0">101–102 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379106F109A7D3FAED7B7C7D9B" box="[151,281,963,989]" pageId="23" pageNumber="174" value="2019-06-27">
<collectingDate id="375CE9DF9106F109A7D3FAED7B7C7D9B" box="[151,281,963,989]" pageId="23" pageNumber="174" value="2019-06-27">27.06.2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="E3CE3CAA9106F109A660FAEA795A7A47" ID-GBIF-Occurrence="4897992308" collectingDate="2019-06-27" elevation="82" latitude="44.556" location="Goryachiy Klyuch Municipality" longLatPrecision="67" longitude="39.108" pageId="23" pageNumber="174" specimenCount="1" specimenCount-female="1">
<specimenCount id="45A0FD7E9106F109A660FAEA7B5C7D98" box="[292,313,964,990]" count="1" pageId="23" pageNumber="174" type="female">♀</specimenCount>
(blue morph),
<location id="5679602C9106F109A6A6FAED79297D9B" LSID="urn:lsid:plazi:treatment:DB0F87E19106F109A7D3FB1978B779B7:5679602C9106F109A6A6FAED79297D9B" box="[482,844,963,989]" latitude="44.556" longLatPrecision="67" longitude="39.108" name="Goryachiy Klyuch Municipality" pageId="23" pageNumber="174">Goryachiy Klyuch Municipality</location>
, an overgrown pond
<locationDeviation id="42AB2FD59106F109A37CFAED7A837A47" location="Bezymyannoe village" pageId="23" pageNumber="174">1.5 km WNW of Bezymyannoe village</locationDeviation>
,
<geoCoordinate id="369250309106F109A7B2FAC97B0B7A44" box="[246,366,999,1026]" degrees="44.556" direction="north" orientation="latitude" pageId="23" pageNumber="174" precision="55" value="44.556">44.556 N</geoCoordinate>
,
<geoCoordinate id="369250309106F109A63AFAC97B947A44" box="[382,497,999,1026]" degrees="39.108" direction="east" orientation="longitude" pageId="23" pageNumber="174" precision="55" value="39.108">39.108 E</geoCoordinate>
,
<elevation id="D88BD1C49106F109A546FAC978C97A47" box="[514,684,999,1026]" metricMagnitude="1" metricUnit="m" metricValue="8.15" metricValueMax="8.3" metricValueMin="8.0" pageId="23" pageNumber="174" unit="m" value="81.5" valueMax="83.0" valueMin="80.0">
<quantity id="945E9B129106F109A546FAC9780F7A44" box="[514,618,999,1026]" metricMagnitude="1" metricUnit="m" metricValue="8.15" metricValueMax="8.3" metricValueMin="8.0" pageId="23" pageNumber="174" unit="m" value="81.5" valueMax="83.0" valueMin="80.0">80–83 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379106F109A5F9FAC9795A7A47" box="[701,831,999,1025]" pageId="23" pageNumber="174" value="2019-06-27">
<collectingDate id="375CE9DF9106F109A5F9FAC9795A7A47" box="[701,831,999,1025]" pageId="23" pageNumber="174" value="2019-06-27">27.06.2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="E3CE3CAA9106F109A40BFAC67ECA7A63" ID-GBIF-Occurrence="4897992306" collectingDate="2019-06-30" elevation="208" location="Apsheronsk District" pageId="23" pageNumber="174" specimenCount="1" specimenCount-male="1">
<specimenCount id="45A0FD7E9106F109A40BFAC679037A44" box="[847,870,1000,1026]" count="1" pageId="23" pageNumber="174" type="male">♂</specimenCount>
,
<location id="5679602C9106F109A432FAC97E3B7A47" LSID="urn:lsid:plazi:treatment:DB0F87E19106F109A7D3FB1978B779B7:5679602C9106F109A432FAC97E3B7A47" box="[886,1118,999,1025]" name="Apsheronsk District" pageId="23" pageNumber="174">Apsheronsk District</location>
,
<locationDeviation id="42AB2FD59106F109A32AFAC97B777A63" location="Tsurevskiy Settlement" pageId="23" pageNumber="174">
<quantity id="945E9B129106F109A32AFAC97EA77A47" box="[1134,1218,999,1026]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="174" unit="km" value="0.5">0.5 km</quantity>
NE of Tsurevskiy Settlement
</locationDeviation>
, a big former quarry, 44.414–416 N, 39.786–789 E,
<elevation id="D88BD1C49106F109A423FD257E447A63" box="[871,1057,1035,1062]" metricMagnitude="2" metricUnit="m" metricValue="2.075" metricValueMax="2.08" metricValueMin="2.07" pageId="23" pageNumber="174" unit="m" value="207.5" valueMax="208.0" valueMin="207.0">
<quantity id="945E9B129106F109A423FD2579807A60" box="[871,997,1035,1062]" metricMagnitude="2" metricUnit="m" metricValue="2.075" metricValueMax="2.08" metricValueMin="2.07" pageId="23" pageNumber="174" unit="m" value="207.5" valueMax="208.0" valueMin="207.0">207–208 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379106F109A369FD257ECA7A63" box="[1069,1199,1035,1061]" pageId="23" pageNumber="174" value="2019-06-30">
<collectingDate id="375CE9DF9106F109A369FD257ECA7A63" box="[1069,1199,1035,1061]" pageId="23" pageNumber="174" value="2019-06-30">30.06.2019</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="E3CE3CAA9106F109A3FDFD25796C7A28" ID-GBIF-Occurrence="4897992338" collectingDate="2019-07-05" elevation="610" location="Lake Maloe" municipality="Prikubanskiy District" pageId="23" pageNumber="174" specimenCount="1" specimenCount-female="1">
<emphasis id="61D2EAE59106F109A3FDFD257B697A0F" bold="true" pageId="23" pageNumber="174">Karachay-Cherkes Republic:</emphasis>
<specimenCount id="45A0FD7E9106F109A652FD1E7B4E7A0C" box="[278,299,1072,1098]" count="1" pageId="23" pageNumber="174" type="female">♀</specimenCount>
(blue morph),
<collectingMunicipality id="B37DAC8D9106F109A698FD0178B57A0F" box="[476,720,1071,1097]" pageId="23" pageNumber="174">Prikubanskiy District</collectingMunicipality>
,
<locationDeviation id="42AB2FD59106F109A59AFD017EE07A0F" box="[734,1157,1071,1097]" location="Kavkazskiy village" pageId="23" pageNumber="174">3.3 km WSW of Kavkazskiy village</locationDeviation>
,
<location id="5679602C9106F109A3D7FD017F787A0F" LSID="urn:lsid:plazi:treatment:DB0F87E19106F109A7D3FB1978B779B7:5679602C9106F109A3D7FD017F787A0F" box="[1171,1309,1071,1097]" municipality="Prikubanskiy District" name="Lake Maloe" pageId="23" pageNumber="174">Lake Maloe</location>
NW bank, 44.251–254 N, 42.195–200 E,
<elevation id="D88BD1C49106F109A546FD7D78ED7A2B" box="[514,648,1107,1134]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="23" pageNumber="174" unit="m" value="610.0">
<quantity id="945E9B129106F109A546FD7D782F7A28" box="[514,586,1107,1134]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="23" pageNumber="174" unit="m" value="610.0">610 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379106F109A5D1FD7D796C7A28" box="[661,777,1107,1134]" pageId="23" pageNumber="174" value="2019-07-05">
<collectingDate id="375CE9DF9106F109A5D1FD7D796C7A28" box="[661,777,1107,1134]" pageId="23" pageNumber="174" value="2019-07-05">5.07.2019</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="45A0FD7E9106F109A451FD7D79307A28" box="[789,853,1107,1134]" count="3" pageId="23" pageNumber="174" type="male">3 ♂♂</specimenCount>
(42847508),
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<collectingRegion id="9162F8159106F109A51FFD9179057A9F" box="[603,864,1215,1241]" country="Russia" name="Dagestan" pageId="23" pageNumber="174">Republic of Dagestan</collectingRegion>
:
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environs, the
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et
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leg.
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(blue morph) (99613386, 99613479),
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, the former second pond (a boggy meadow),
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, the
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upper valley,
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a.s.l.
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Additional photographic registrations:
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Krasnodarskiy Kray:
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(blue morph) (42847863, 108804470 (
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settlement,
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environs, a
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Fig.
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<emphasis id="61D2EAE59106F109A783FEA17B9D79EF" bold="true" box="[199,504,1935,1961]" pageId="23" pageNumber="174">Distribution and habitat.</emphasis>
Widespread at lower elevations of the Caucasus but never numerous. Breeds in lakes, rarely pools. These damselflies more readily disperse apart from water than their other congeners, occurring at forest margins or even among steppe vegetation.
</paragraph>
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5.
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<emphasis id="61D2EAE59107F108A7F1F84A7BD67F38" bold="true" box="[181,435,356,382]" italics="true" pageId="22" pageNumber="173">Coenagrion ornatum</emphasis>
(
<bibRefCitation id="37374B069107F108A687F84A782A7F38" author="Selys Lonchamps, E." box="[451,591,356,382]" pageId="22" pageNumber="173" refId="ref29427" refString="Selys Lonchamps, E. (de) (1850) Revue des Odonates ou Libellules d'Europe. C. Murquardt, Bruxelles et Leipzig &amp; Roret, Paris, 408 pp." type="book" year="1850">Selys, 1850</bibRefCitation>
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<paragraph id="531936F79107F108A7D3F8A67B597FE4" blockId="22.[151,599,356,418]" box="[151,316,392,418]" pageId="22" pageNumber="173">
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,
<figureCitation id="CB9D2A729107F108A7BCF8A67B597FE4" box="[248,316,392,418]" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="22" pageNumber="173">11a–d</figureCitation>
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<paragraph id="531936F79107F108A7D3F8FE79197C74" blockId="22.[151,1437,464,1319]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A7D3F8FE7B357FAC" bold="true" box="[151,336,464,490]" pageId="22" pageNumber="173">Reported from:</emphasis>
Krasnodarskiy Kray, environs of Gryaznukhinskaya Stanitsa (sub.
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Selys
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) (
<bibRefCitation id="37374B069107F108A27EF8FE7AB67C48" author="Bartenef, A. N." pageId="22" pageNumber="173" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">Bartenef 1930</bibRefCitation>
), the Pslukh River in Sochi Municipality (
<bibRefCitation id="37374B069107F108A58EF8DA79E37C48" author="Skvortsov, V. E." box="[714,902,500,526]" pageId="22" pageNumber="173" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
); Kellermesskaya and Khanskaya Stanitsas in
<collectingRegion id="9162F8159107F108A7D3FB367A8B7C74" box="[151,238,536,562]" country="Russia" name="Adygeya" pageId="22" pageNumber="173">Adygea</collectingRegion>
Republic (
<bibRefCitation id="37374B069107F108A62DFB3678687C74" author="Bartenef, A. N." box="[361,525,536,562]" pageId="22" pageNumber="173" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">Bartenef 1930</bibRefCitation>
;
<bibRefCitation id="37374B069107F108A55DFB3679157C74" author="Shapovalov, M. I. &amp; Korotkov, E. A." box="[537,880,536,562]" pageId="22" pageNumber="173" pagination="341 - 349" refId="ref29460" refString="Shapovalov, M. I. &amp; Korotkov, E. A. (2019) Update of the Odonata fauna of Georgia, southern Caucasus ecoregion. Russian Entomological Journal, 28 (4), 341 - 349. https: // doi. org / 10.15298 / rusentj. 28.4.01" type="journal article" year="2019">Shapovalov &amp; Korotkov 2019</bibRefCitation>
).
</paragraph>
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<emphasis id="61D2EAE59107F108A783FB127B297C10" bold="true" box="[199,332,572,598]" pageId="22" pageNumber="173">Specimens.</emphasis>
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<emphasis id="61D2EAE59107F108A61DFB1278D67C10" bold="true" box="[345,691,572,598]" pageId="22" pageNumber="173">
<collectingRegion id="9162F8159107F108A61DFB1278D67C10" box="[345,691,572,598]" name="Karachay-Cherkes Republic" pageId="22" pageNumber="173">Karachay-Cherkes Republic</collectingRegion>
</emphasis>
:
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(37680843, 37680856),
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,
<collectingCountry id="2BB176679107F108A36EFB127E1D7C10" box="[1066,1144,572,598]" name="Russia" pageId="22" pageNumber="173">Russia</collectingCountry>
, N
<collectingCounty id="BA784E7B9107F108A3EDFB127F737C10" box="[1193,1302,572,598]" pageId="22" pageNumber="173">Caucasus</collectingCounty>
,
<collectingMunicipality id="B37DAC8D9107F108A262FB127A8B7C3C" pageId="22" pageNumber="173">Zelenchuk District</collectingMunicipality>
,
<location id="5679602C9107F108A7BDFB4F7BC57C3C" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A7BDFB4F7BC57C3C" box="[249,416,608,635]" country="Russia" county="Caucasus" latitude="1.012" longLatPrecision="78" longitude="-1.014" municipality="Zelenchuk District" name="Dausuz village" pageId="22" pageNumber="173" stateProvince="Karachay-Cherkes Republic">Dausuz village</location>
W environs, S banks of a large water reserve at the
<location id="5679602C9107F108A4ADFB4E7F697C3C" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A4ADFB4E7F697C3C" box="[1001,1292,608,634]" country="Russia" county="Caucasus" latitude="1.012" longLatPrecision="78" longitude="-1.014" municipality="Zelenchuk District" name="Bolzhoy Zelenchuk River" pageId="22" pageNumber="173" stateProvince="Karachay-Cherkes Republic">Bolzhoy Zelenchuk River</location>
, 43.793–796 N, 41.536–540 E,
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a.s.l.
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,
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.
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<emphasis id="61D2EAE59107F108A5BCFBAA79E07CD8" bold="true" box="[760,901,644,670]" pageId="22" pageNumber="173">Republic of</emphasis>
<emphasis id="61D2EAE59107F108A4D5FBAA7E6C7CD8" bold="true" box="[913,1033,644,670]" pageId="22" pageNumber="173">Dagestan:</emphasis>
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<specimenCount id="45A0FD7E9107F108A350FBAB7E4E7CD9" box="[1044,1067,645,671]" count="1" pageId="22" pageNumber="173" type="male">♂</specimenCount>
,
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(178696749, 180155503, 180155777, 1180155989, 180156091, 80156210, 180156435, 180157387, 180159191 (
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), 180160258, 180160572, 180160723, 180160831, 180160934),
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,
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S environs,
<location id="5679602C9107F108A3D9FBE27A867D4C" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A3D9FBE27A867D4C" country="Russia" county="Buynaksk District" latitude="47.1157" longLatPrecision="6" longitude="47.1169" municipality="Buynaksk" name="Verkhnee Kazanishche village" pageId="22" pageNumber="173" stateProvince="Dagestan">Verkhnee Kazanishche village</location>
environs, a swampy, reedy brook on a gentle left slope of the
<location id="5679602C9107F108A4FFFBDE7EAF7D4C" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A4FFFBDE7EAF7D4C" box="[955,1226,752,778]" country="Russia" county="Buynaksk District" latitude="47.1157" longLatPrecision="6" longitude="47.1169" municipality="Buynaksk" name="Akpeozen' River valley" pageId="22" pageNumber="173" stateProvince="Dagestan">Akpeozen’ River valley</location>
,
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–42.7992 N, 47.1157–
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,
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a.s.l.
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,
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<collectingDate id="375CE9DF9107F108A5CCFA3A796F7D69" box="[648,778,788,815]" pageId="22" pageNumber="173" value="2023-05-29">29.05.2023</collectingDate>
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;
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<specimenCount id="45A0FD7E9107F108A45DFA3A79397D69" box="[793,860,788,815]" count="6" pageId="22" pageNumber="173" type="male">6 ♂♂</specimenCount>
(
<figureCitation id="CB9D2A729107F108A42BFA3A79887D68" box="[879,1005,788,814]" captionStart="FIGURE 10" captionStartId="22.[151,250,1641,1665]" captionTargetBox="[163,1425,1352,1618]" captionTargetId="figure-399@22.[151,1436,1342,1618]" captionTargetPageId="22" captionText="FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189930" httpUri="https://zenodo.org/record/12189930/files/figure.png" pageId="22" pageNumber="173">Fig. 10d–e</figureCitation>
) (178695634, 178695635 (
<figureCitation id="CB9D2A729107F108A26FFA3A7FEB7D68" box="[1323,1422,788,814]" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="22" pageNumber="173">Fig. 11b</figureCitation>
), 178695636, 180231099, 180231185, 180643335, 180643481, 180643723, 180643834, 180643926, 180644391, 180644477, 180644677, 180644769, 180644838, 180644922, 180645019, 180645088, 180950128, 180951966, 180952452, 180952577, 180952701, 180952786, 180952885, 180953036, 180953178, 180953270),
<collectingCounty id="BA784E7B9107F108A26FFAAE7A8B7DF8" pageId="22" pageNumber="173">Tabasaran District</collectingCounty>
, the
<collectingMunicipality id="B37DAC8D9107F108A666FA8A7B9D7DF8" box="[290,504,932,958]" pageId="22" pageNumber="173">Rubas River valley</collectingMunicipality>
<locationDeviation id="42AB2FD59107F108A6BBFA8A79217DF8" box="[511,836,932,959]" location="Sirtych village" pageId="22" pageNumber="173">
<quantity id="945E9B129107F108A6BBFA8A78287DF8" box="[511,589,932,959]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="173" unit="km" value="2.5">2.5 km</quantity>
NE of Sirtych village
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, a swampy
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,
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–41.8527 N, 48.2396–
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,
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<quantity id="945E9B129107F108A6F4FAE678547DA5" box="[432,561,968,995]" metricMagnitude="2" metricUnit="m" metricValue="1.04" metricValueMax="1.05" metricValueMin="1.03" pageId="22" pageNumber="173" unit="m" value="104.0" valueMax="105.0" valueMin="103.0">103–105 m</quantity>
a.s.l.
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,
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<collectingDate id="375CE9DF9107F108A53AFAE679657DA5" box="[638,768,968,995]" pageId="22" pageNumber="173" value="2023-05-28">28.05.2023</collectingDate>
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;
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<specimenCount id="45A0FD7E9107F108A448FAE679357DA5" box="[780,848,968,995]" count="4" pageId="22" pageNumber="173" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="45A0FD7E9107F108A41AFAE679C57DA5" box="[862,928,968,995]" count="3" pageId="22" pageNumber="173" type="female">3 ♀♀</specimenCount>
,
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,
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,
<location id="5679602C9107F108A7D3FAC27BAA7A40" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A7D3FAC27BAA7A40" box="[151,463,1004,1030]" country="Russia" county="Magaramkent District" latitude="48.547" longLatPrecision="66" longitude="48.548" municipality="Samur Delta Forest" name="Bil' - Bil' Kazmalyar village" pageId="22" pageNumber="173" stateProvince="Dagestan">Bil’-Bil’ Kazmalyar village</location>
, a brook and a pond,
<geoCoordinate id="369250309107F108A581FAC279777A40" box="[709,786,1004,1030]" degrees="41.818" direction="north" orientation="latitude" pageId="22" pageNumber="173" precision="55" value="41.818">41.818</geoCoordinate>
–41.819 N, 48.547–
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a.s.l.
</elevation>
,
<date id="271810379107F108A261FAC27FFD7A40" box="[1317,1432,1004,1030]" pageId="22" pageNumber="173" value="2022-06-04">
<collectingDate id="375CE9DF9107F108A261FAC27FFD7A40" box="[1317,1432,1004,1030]" pageId="22" pageNumber="173" value="2022-06-04">4.06.2022</collectingDate>
</date>
,
<collectorName id="FE5353219107F108A7D3FD3F7A9F7A6C" box="[151,250,1040,1067]" pageId="22" pageNumber="173">E. Ilyina</collectorName>
leg
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;
<specimenCount id="45A0FD7E9107F108A67CFD3E7B1E7A6D" box="[312,379,1040,1067]" count="2" pageId="22" pageNumber="173" type="male">2 ♂♂</specimenCount>
, the same place,
<date id="271810379107F108A501FD3E78DD7A6D" box="[581,696,1040,1067]" pageId="22" pageNumber="173" value="2023-06-05">5.06.2023</date>
, E. Ilyina leg.;
<specimenCount id="45A0FD7E9107F108A429FD3E79F97A6D" box="[877,924,1040,1067]" count="2" pageId="22" pageNumber="173" type="male">2 ♂</specimenCount>
, the same place,
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, E. Ilyina leg.;
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(178693662 (
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), 178693663, 181380130, 181380245, 181380516, 181380589, 181380955, 181381118, 182680720, 182680856, 182681001, 182681109, 182681639, 182681821, 182682133, 182682287, 182683094, 182683374, 182683704, 182683883, 182683987, 182684102, 182771447, 182771534, 182771582), the same place,
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), 98116240, 98963192, 98963230, 98963408, 98963476, 98963562, 98963695, 98963732, 98964853, 98964859, 98964915, 98964939, 98964993, 98965076, 98968166, 98968201),
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left arm between
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<paragraph id="531936F79107F108A7D3FF477995788F" blockId="22.[151,1436,1641,1738]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A7D3FF477B4678C4" bold="true" box="[151,291,1641,1666]" pageId="22" pageNumber="173">FIGURE 10.</emphasis>
Male appendages of
<taxonomicName id="94A64D749107F108A543FF4478BA78C7" baseAuthorityName="Selys" baseAuthorityYear="1850" box="[519,735,1642,1665]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ornatum">
<emphasis id="61D2EAE59107F108A543FF4478BA78C7" box="[519,735,1642,1665]" italics="true" pageId="22" pageNumber="173">Coenagrion ornatum</emphasis>
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specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm.
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<subSubSection id="1BBC657C9107F108A783FE3F7F557999" pageId="22" pageNumber="173" type="distribution">
<paragraph id="531936F79107F108A783FE3F79427916" blockId="22.[151,1437,1809,2016]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A783FE3F7B96796D" bold="true" box="[199,499,1809,1835]" pageId="22" pageNumber="173">Distribution and habitat.</emphasis>
Widespread but local in the Caucasus, although the populations found were rather large. The species invariably occurs at small slow streams.
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<paragraph id="531936F79107F108A783FE777F557999" blockId="22.[151,1437,1809,2016]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A783FE777B317935" bold="true" box="[199,340,1881,1907]" pageId="22" pageNumber="173">Characters.</emphasis>
The characters are unmistakable for this species (
<bibRefCitation id="37374B069107F108A4CCFE777E507935" author="Kalkman, V. J." box="[904,1077,1881,1907]" pageId="22" pageNumber="173" pagination="3 - 82" refId="ref28232" refString="Kalkman, V. J. (2006) Key to the dragonflies of Turkey including species known from Greece, Bulgaria, Lebanon, Syria, the Trans-Caucasus and Iran. Brachytron, 10 (1), 3 - 82." type="journal article" year="2006">Kalkman 2006</bibRefCitation>
;
<bibRefCitation id="37374B069107F108A304FE777E937935" author="Skvortsov, V. E." box="[1088,1270,1881,1907]" pageId="22" pageNumber="173" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
;
<bibRefCitation id="37374B069107F108A245FE777AB479D1" author="Dijkstra, K. - D. &amp; Schroter, A. &amp; Lewington, R." pageId="22" pageNumber="173" refId="ref27712" refString="Dijkstra, K. - D., Schroter, A. &amp; Lewington, R. (2020) Field Guide to the Dragonflies of Britain and Europe, including western Turkey and north-western Africa. 2 nd Edition. Bloomsbury Publishing, London, Oxford etc., 336 pp." type="book" year="2020">
Dijkstra
<emphasis id="61D2EAE59107F108A227FE747FF87935" box="[1379,1437,1881,1907]" italics="true" pageId="22" pageNumber="173">et al.</emphasis>
2020
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) and deserve no attention beyond a particular problem with the position of the male cerci (see ‘Discussion’). In our male specimens, the cerci are more or less divergent to a variable extent (
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); in specimens from the Rubas River valley the divergence is on average less expressed, with the least divergent case shown in
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<treatment id="DB0F87E19117F118A7D3F9B97E6D7D76" ID-DOI="http://doi.org/10.5281/zenodo.12206836" ID-GBIF-Taxon="233309476" ID-Zenodo-Dep="12206836" LSID="urn:lsid:plazi:treatment:DB0F87E19117F118A7D3F9B97E6D7D76" httpUri="http://treatment.plazi.org/id/DB0F87E19117F118A7D3F9B97E6D7D76" lastPageNumber="157" pageId="6" pageNumber="157">
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<heading id="0851819B9117F118A7D3F9B979967EF7" bold="true" box="[151,1011,151,177]" fontSize="11" level="1" pageId="6" pageNumber="157" reason="1">
<emphasis id="61D2EAE59117F118A7D3F9B979967EF7" bold="true" box="[151,1011,151,177]" pageId="6" pageNumber="157">
<taxonomicName id="94A64D749117F118A7D3F9B9780D7EF7" ID-CoL="WTXW" authority="Lohmann, 1983" authorityName="Lohmann" authorityYear="1983" box="[151,616,151,177]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE59117F118A7D3F9B97BF87EF7" bold="true" box="[151,413,151,177]" italics="true" pageId="6" pageNumber="157">Coenagrion persicum</emphasis>
Lohmann, 1983
</taxonomicName>
(see
<bibRefCitation id="37374B069117F118A5E2F9B979807EF7" author="Kosterin, O. E. &amp; Ahmadi, A." box="[678,997,151,177]" pageId="6" pageNumber="157" pagination="1 - 65" refId="ref28477" refString="Kosterin, O. E. &amp; Ahmadi, A. (2018) Odonata observed in Central Zagros, Iran, in late May 2017. International Dragonfly Fund Report, 117, 1 - 65." type="journal article" year="2018">Kosterin &amp; Ahmadi 2018</bibRefCitation>
):
</emphasis>
</heading>
</paragraph>
<paragraph id="531936F79117F118A7D3F9F47B5D7CAE" blockId="6.[151,1436,217,816]" pageId="6" pageNumber="157">
<materialsCitation id="E3CE3CAA9117F118A7D3F9F478967F5E" ID-GBIF-Occurrence="4897992321" collectingDate="2017-05-29" country="Iran" county="Tafresh Shakhrestan" elevation="2057" location="Shilanevar valley" municipality="Tafresh City" pageId="6" pageNumber="157" specimenCount="1" specimenCount-male="1" stateProvince="Markazi">
<specimenCount id="45A0FD7E9117F118A7D3F9F47ACB7EB2" box="[151,174,218,244]" count="1" pageId="6" pageNumber="157" type="male">♂</specimenCount>
,
<collectingCountry id="2BB176679117F118A7FEF9F47A8F7EB2" box="[186,234,218,244]" name="Iran" pageId="6" pageNumber="157">Iran</collectingCountry>
,
<collectingRegion id="9162F8159117F118A7B2F9F77BFB7EB2" box="[246,414,217,244]" country="Iran" name="Markazi" pageId="6" pageNumber="157">Markazi Ostan</collectingRegion>
,
<collectingCounty id="BA784E7B9117F118A6EDF9F778F57EB5" box="[425,656,217,243]" pageId="6" pageNumber="157">Tafresh Shakhrestan</collectingCounty>
,
<collectingMunicipality id="B37DAC8D9117F118A5DFF9F779437EB5" box="[667,806,217,243]" pageId="6" pageNumber="157">Tafresh City</collectingMunicipality>
env.,
<location id="5679602C9117F118A42DF9F77E487EB5" LSID="urn:lsid:plazi:treatment:DB0F87E19117F118A7D3F9B97E6D7D76:5679602C9117F118A42DF9F77E487EB5" box="[873,1069,217,243]" country="Iran" county="Tafresh Shakhrestan" municipality="Tafresh City" name="Shilanevar valley" pageId="6" pageNumber="157" stateProvince="Markazi">Shilanevar valley</location>
, the lower pond, 34.6608–6654° N 50.0323–0402° E,
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<quantity id="945E9B129117F118A6CDF9D378407F5E" box="[393,549,253,280]" metricMagnitude="3" metricUnit="m" metricValue="2.0565" metricValueMax="2.077" metricValueMin="2.036" pageId="6" pageNumber="157" unit="m" value="2056.5" valueMax="2077.0" valueMin="2036.0">2036–2077 m</quantity>
a.s.l.
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,
<date id="271810379117F118A535F9D378967F5E" box="[625,755,253,280]" pageId="6" pageNumber="157" value="2017-05-29">
<collectingDate id="375CE9DF9117F118A535F9D378967F5E" box="[625,755,253,280]" pageId="6" pageNumber="157" value="2017-05-29">29.05.2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="E3CE3CAA9117F118A5BBF9D07FFD7F7A" ID-GBIF-Occurrence="4897992329" collectingDate="2017-05-30" collectorName="Lake Kavir-e Meighan S" country="Iran" elevation="1658" location="Arak Shakhtrestan" pageId="6" pageNumber="157" specimenCount="1" specimenCount-female="1" stateProvince="Markazi">
<specimenCount id="45A0FD7E9117F118A5BBF9D079717F5E" box="[767,788,254,280]" count="1" pageId="6" pageNumber="157" type="female">♀</specimenCount>
(‘dark’ morph),
<collectingCountry id="2BB176679117F118A497F9D07E667F5E" box="[979,1027,254,280]" name="Iran" pageId="6" pageNumber="157">Iran</collectingCountry>
,
<collectingRegion id="9162F8159117F118A34BF9D37EDC7F51" box="[1039,1209,253,279]" country="Iran" name="Markazi" pageId="6" pageNumber="157">Markazi Ostan</collectingRegion>
,
<location id="5679602C9117F118A380F9D37FFC7F51" LSID="urn:lsid:plazi:treatment:DB0F87E19117F118A7D3F9B97E6D7D76:5679602C9117F118A380F9D37FFC7F51" box="[1220,1433,253,279]" country="Iran" name="Arak Shakhtrestan" pageId="6" pageNumber="157" stateProvince="Markazi">Arak Shakhtrestan</location>
,
<collectorName id="FE5353219117F118A7D3F80F7BCF7F7D" box="[151,426,289,315]" pageId="6" pageNumber="157">Lake Kavir-e Meighan S</collectorName>
bank, a rush swamp, 34.1530–1536° N 49.8112–8131° E,
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<quantity id="945E9B129117F118A371F80F7EAA7F7D" box="[1077,1231,289,316]" metricMagnitude="3" metricUnit="m" metricValue="1.658" metricValueMax="1.659" metricValueMin="1.657" pageId="6" pageNumber="157" unit="m" value="1658.0" valueMax="1659.0" valueMin="1657.0">1657–1659 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379117F118A252F80F7FFD7F7A" box="[1302,1432,289,316]" pageId="6" pageNumber="157" value="2017-05-30">
<collectingDate id="375CE9DF9117F118A252F80F7FFD7F7A" box="[1302,1432,289,316]" pageId="6" pageNumber="157" value="2017-05-30">30.05.2017</collectingDate>
</date>
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;
<materialsCitation id="E3CE3CAA9117F118A7D3F86B7F757FC5" ID-GBIF-Occurrence="4897992312" collectingDate="2017-05-19" country="Iran" county="Shazand Shakhrestan" elevation="1804" latitude="49.3627" location="Gavigodar village" longLatPrecision="6" longitude="49.3657" municipality="Garehchay River" pageId="6" pageNumber="157" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Markazi">
<specimenCount id="45A0FD7E9117F118A7D3F86B7ABD7F19" box="[151,216,325,351]" count="2" pageId="6" pageNumber="157" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="45A0FD7E9117F118A7A0F86B7B467F19" box="[228,291,325,351]" count="4" pageId="6" pageNumber="157" type="female">4 ♀♀</specimenCount>
(3 ‘blue morph’, 1 ‘dark morph’) (
<figureCitation id="CB9D2A729117F118A5E8F86B789C7F19" box="[684,761,325,351]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="6" pageNumber="157">Fig. 7j</figureCitation>
),
<collectingCountry id="2BB176679117F118A44AF86B795B7F19" box="[782,830,325,351]" name="Iran" pageId="6" pageNumber="157">Iran</collectingCountry>
,
<collectingRegion id="9162F8159117F118A40DF86B79947F19" box="[841,1009,325,351]" country="Iran" name="Markazi" pageId="6" pageNumber="157">Markazi Ostan</collectingRegion>
,
<collectingCounty id="BA784E7B9117F118A4B9F86B7E8A7F19" box="[1021,1263,325,351]" pageId="6" pageNumber="157">Shazand Shakhrestan</collectingCounty>
, the
<collectingMunicipality id="B37DAC8D9117F118A267F86B7AB07FC4" pageId="6" pageNumber="157">Garehchay River</collectingMunicipality>
at
<location id="5679602C9117F118A7B2F8467BA77FC4" LSID="urn:lsid:plazi:treatment:DB0F87E19117F118A7D3F9B97E6D7D76:5679602C9117F118A7B2F8467BA77FC4" box="[246,450,360,386]" country="Iran" county="Shazand Shakhrestan" latitude="49.3627" longLatPrecision="6" longitude="49.3657" municipality="Garehchay River" name="Gavigodar village" pageId="6" pageNumber="157" stateProvince="Markazi">Gavigodar village</location>
,
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–34.1111° N 49.3627–
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,
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a.s.l.
</elevation>
,
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<collectingDate id="375CE9DF9117F118A3CAF8467F757FC5" box="[1166,1296,360,387]" pageId="6" pageNumber="157" value="2017-05-19">19.05.2017</collectingDate>
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;
<materialsCitation id="E3CE3CAA9117F118A25EF8477BD97F8D" ID-GBIF-Occurrence="4897992358" collectingDate="2017-05-20" country="Iran" elevation="1857" latitude="49.3606" location="Bolagh" longLatPrecision="6" longitude="49.3618" municipality="Shazand Shakhrestan" pageId="6" pageNumber="157" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Markazi">
<specimenCount id="45A0FD7E9117F118A25EF8477F547FC5" box="[1306,1329,361,387]" count="1" pageId="6" pageNumber="157" type="male">♂</specimenCount>
,
<specimenCount id="45A0FD7E9117F118A279F8477F377FC5" box="[1341,1362,361,387]" count="1" pageId="6" pageNumber="157" type="female">♀</specimenCount>
(‘dark morph),
<collectingCountry id="2BB176679117F118A7BCF8A37B4D7FE1" box="[248,296,397,423]" name="Iran" pageId="6" pageNumber="157">Iran</collectingCountry>
,
<collectingRegion id="9162F8159117F118A677F8A27BB97FE0" box="[307,476,396,422]" country="Iran" name="Markazi" pageId="6" pageNumber="157">Markazi Ostan</collectingRegion>
,
<collectingMunicipality id="B37DAC8D9117F118A6ACF8A278BF7FE0" box="[488,730,396,422]" pageId="6" pageNumber="157">Shazand Shakhrestan</collectingMunicipality>
,
<location id="5679602C9117F118A5A1F8A279537FE0" LSID="urn:lsid:plazi:treatment:DB0F87E19117F118A7D3F9B97E6D7D76:5679602C9117F118A5A1F8A279537FE0" box="[741,822,396,422]" country="Iran" latitude="49.3606" longLatPrecision="6" longitude="49.3618" municipality="Shazand Shakhrestan" name="Bolagh" pageId="6" pageNumber="157" stateProvince="Markazi">Bolagh</location>
spring, 33.9875–9880° N
<geoCoordinate id="369250309117F118A324F8A27ED87FE0" box="[1120,1213,396,422]" degrees="49.3606" direction="north" orientation="latitude" pageId="6" pageNumber="157" precision="5" value="49.3606">49.3606</geoCoordinate>
–
<geoCoordinate id="369250309117F118A38FF8A27F2F7FE1" box="[1227,1354,396,423]" degrees="49.3618" direction="east" orientation="longitude" pageId="6" pageNumber="157" precision="5" value="49.3618">49.3618° E</geoCoordinate>
,
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<quantity id="945E9B129117F118A212F8A27A8B7F8C" metricMagnitude="3" metricUnit="m" metricValue="1.857" metricValueMax="1.861" metricValueMin="1.853" pageId="6" pageNumber="157" unit="m" value="1857.0" valueMax="1861.0" valueMin="1853.0">1853– 1861 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379117F118A67EF89E7BD97F8D" box="[314,444,432,459]" pageId="6" pageNumber="157" value="2017-05-20">
<collectingDate id="375CE9DF9117F118A67EF89E7BD97F8D" box="[314,444,432,459]" pageId="6" pageNumber="157" value="2017-05-20">20.05.2017</collectingDate>
</date>
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;
<materialsCitation id="E3CE3CAA9117F118A68CF89E79977FA8" ID-GBIF-Occurrence="4897992335" collectingDate="2017-05-20" country="Iran" county="Shazand Shakhrestan" elevation="2158" latitude="49.343" location="Kaedan Chashmah" longLatPrecision="6" longitude="49.3434" municipality="Astaneh Town" pageId="6" pageNumber="157" specimenCount="1" specimenCount-male="1" stateProvince="Markazi">
<specimenCount id="45A0FD7E9117F118A68CF89E7BBA7F8C" box="[456,479,432,458]" count="1" pageId="6" pageNumber="157" type="male">♂</specimenCount>
,
<collectingCountry id="2BB176679117F118A6A9F89E78787F8C" box="[493,541,432,458]" name="Iran" pageId="6" pageNumber="157">Iran</collectingCountry>
,
<collectingRegion id="9162F8159117F118A56EF89E78B67F8C" box="[554,723,432,458]" country="Iran" name="Markazi" pageId="6" pageNumber="157">Markazi Ostan</collectingRegion>
,
<collectingCounty id="BA784E7B9117F118A5A4F89E79B17F8C" box="[736,980,432,458]" pageId="6" pageNumber="157">Shazand Shakhrestan</collectingCounty>
,
<collectingMunicipality id="B37DAC8D9117F118A49BF89E7EE77F8C" box="[991,1154,432,458]" pageId="6" pageNumber="157">Astaneh Town</collectingMunicipality>
env.,
<location id="5679602C9117F118A38CF89E7FF97F8C" LSID="urn:lsid:plazi:treatment:DB0F87E19117F118A7D3F9B97E6D7D76:5679602C9117F118A38CF89E7FF97F8C" box="[1224,1436,432,458]" country="Iran" county="Shazand Shakhrestan" latitude="49.343" longLatPrecision="6" longitude="49.3434" municipality="Astaneh Town" name="Kaedan Chashmah" pageId="6" pageNumber="157" stateProvince="Markazi">Kaedan Chashmah</location>
spring,
<geoCoordinate id="369250309117F118A7AFF8FD7B227FA8" box="[235,327,467,494]" degrees="33.9175" direction="north" orientation="latitude" pageId="6" pageNumber="157" precision="5" value="33.9175">33.9175</geoCoordinate>
–33.9177° N 49.3430–
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,
<elevation id="D88BD1C49117F118A596F8FD79017FA8" box="[722,868,467,494]" metricMagnitude="3" metricUnit="m" metricValue="2.158" pageId="6" pageNumber="157" unit="m" value="2158.0">
<quantity id="945E9B129117F118A596F8FD79427FA8" box="[722,807,467,494]" metricMagnitude="3" metricUnit="m" metricValue="2.158" pageId="6" pageNumber="157" unit="m" value="2158.0">2158 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379117F118A434F8FD79977FA8" box="[880,1010,467,494]" pageId="6" pageNumber="157" value="2017-05-20">
<collectingDate id="375CE9DF9117F118A434F8FD79977FA8" box="[880,1010,467,494]" pageId="6" pageNumber="157" value="2017-05-20">20.05.2017</collectingDate>
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;
<materialsCitation id="E3CE3CAA9117F118A4B9F8FA78797C70" ID-GBIF-Occurrence="4897992342" collectingDate="2017-05-20" country="Iran" elevation="1998" location="Emarat village" municipality="Shazand Shakhrestan" pageId="6" pageNumber="157" specimenCount="1" specimenCount-male="1" stateProvince="Markazi">
<specimenCount id="45A0FD7E9117F118A4B9F8FA7E777FA8" box="[1021,1042,468,494]" count="1" pageId="6" pageNumber="157" type="male">♂</specimenCount>
(
<figureCitation id="CB9D2A729117F118A365F8FA7E1C7FA8" box="[1057,1145,468,494]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="6" pageNumber="157">Figs 2n</figureCitation>
,
<figureCitation id="CB9D2A729117F118A3C0F8FA7EC77FA8" box="[1156,1186,468,494]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="6" pageNumber="157">3n</figureCitation>
,
<figureCitation id="CB9D2A729117F118A3EAF8FA7EA37FA8" box="[1198,1222,468,494]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="6" pageNumber="157">4l</figureCitation>
,
<figureCitation id="CB9D2A729117F118A396F8FD7E8B7FA8" box="[1234,1262,467,494]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="6" pageNumber="157">5a</figureCitation>
),
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,
<collectingRegion id="9162F8159117F118A27BF8FA7ABE7C57" country="Iran" name="Markazi" pageId="6" pageNumber="157">Markazi Ostan</collectingRegion>
,
<collectingMunicipality id="B37DAC8D9117F118A7A2F8D97BB37C57" box="[230,470,503,529]" pageId="6" pageNumber="157">Shazand Shakhrestan</collectingMunicipality>
,
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env., a spring and swampy reservoir, 33.8637–8667° N 49.5775–5795° E,
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<quantity id="945E9B129117F118A7F2FB357B347C70" box="[182,337,539,566]" metricMagnitude="3" metricUnit="m" metricValue="1.998" metricValueMax="2.001" metricValueMin="1.9949999999999999" pageId="6" pageNumber="157" unit="m" value="1998.0" valueMax="2001.0" valueMin="1995.0">1995–2001 m</quantity>
a.s.l.
</elevation>
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<collectingDate id="375CE9DF9117F118A6DEFB3578797C70" box="[410,540,539,566]" pageId="6" pageNumber="157" value="2017-05-20">20.05.2017</collectingDate>
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;
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<specimenCount id="45A0FD7E9117F118A563FB32785B7C70" box="[551,574,540,566]" count="1" pageId="6" pageNumber="157" type="male">♂</specimenCount>
,
<collectingCountry id="2BB176679117F118A50EFB32781F7C70" box="[586,634,540,566]" name="Iran" pageId="6" pageNumber="157">Iran</collectingCountry>
,
<collectingRegion id="9162F8159117F118A5C1FB3579487C73" box="[645,813,539,565]" country="Iran" name="Markazi" pageId="6" pageNumber="157">Markazi Ostan</collectingRegion>
,
<collectingMunicipality id="B37DAC8D9117F118A47CFB357E4F7C73" box="[824,1066,539,565]" pageId="6" pageNumber="157">Shazand Shakhrestan</collectingMunicipality>
, the
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,
<locationDeviation id="42AB2FD59117F118A25AFB357B187C1F" location="Hendudar village" pageId="6" pageNumber="157">1.8 km SW of Hendudar village</locationDeviation>
,
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–33.7665° N 49.2178–
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,
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<quantity id="945E9B129117F118A432FB117E777C1F" box="[886,1042,575,601]" metricMagnitude="3" metricUnit="m" metricValue="1.9949999999999999" metricValueMax="1.9969999999999999" metricValueMin="1.9929999999999999" pageId="6" pageNumber="157" unit="m" value="1995.0" valueMax="1997.0" valueMin="1993.0">1993–1997 m</quantity>
a.s.l.
</elevation>
,
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<collectingDate id="375CE9DF9117F118A31AFB117E857C1C" box="[1118,1248,575,602]" pageId="6" pageNumber="157" value="2017-05-20">20.05.2017</collectingDate>
</date>
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;
<materialsCitation id="E3CE3CAA9117F118A3A8FB1178797CE7" ID-GBIF-Occurrence="4897992369" collectingDate="2017-05-25" collectorName="O. Kosterin" country="Iran" county="Borujerd Shakhrestan" elevation="1763" latitude="48.9398" location="Hayan village" longLatPrecision="6" longitude="48.9402" municipality="Borujerd Shakhrestan" pageId="6" pageNumber="157" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Lorestan">
<specimenCount id="45A0FD7E9117F118A3A8FB117F4A7C1F" box="[1260,1327,575,601]" count="2" pageId="6" pageNumber="157" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="45A0FD7E9117F118A278FB117F347C1F" box="[1340,1361,575,601]" count="1" pageId="6" pageNumber="157" type="female">♀</specimenCount>
(‘blue morph’,
<collectingCountry id="2BB176679117F118A7BDFB4D7B4C7C3B" box="[249,297,611,637]" name="Iran" pageId="6" pageNumber="157">Iran</collectingCountry>
,
<collectingRegion id="9162F8159117F118A671FB4D7B867C3B" box="[309,483,611,637]" country="Iran" name="Lorestan" pageId="6" pageNumber="157">Lorestan Ostan</collectingRegion>
,
<collectingCounty id="BA784E7B9117F118A6ABFB4C78837C3A" box="[495,742,610,636]" pageId="6" pageNumber="157">Borujerd Shakhrestan</collectingCounty>
, Goldargilan village,
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–33.9654° N 48.7501–
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,
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<quantity id="945E9B129117F118A7F1FBA87B2A7CE7" box="[181,335,646,673]" metricMagnitude="3" metricUnit="m" metricValue="1.763" metricValueMax="1.781" metricValueMin="1.7449999999999999" pageId="6" pageNumber="157" unit="m" value="1763.0" valueMax="1781.0" valueMin="1745.0">1745–1781 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379117F118A6D2FBA8787D7CE7" box="[406,536,646,673]" pageId="6" pageNumber="157" value="2017-05-25">
<collectingDate id="375CE9DF9117F118A6D2FBA8787D7CE7" box="[406,536,646,673]" pageId="6" pageNumber="157" value="2017-05-25">25.05.2017</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="E3CE3CAA9117F118A566FBA97B517CAE" ID-GBIF-Occurrence="4897992356" collectingDate="2017-05-25" collectorName="O. Kosterin" country="Iran" county="Borujerd Shakhrestan" elevation="1763" latitude="48.9398" location="Hayan village" longLatPrecision="6" longitude="48.9402" municipality="Borujerd Shakhrestan" pageId="6" pageNumber="157" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Lorestan">
<specimenCount id="45A0FD7E9117F118A566FBA9785C7CE7" box="[546,569,647,673]" count="1" pageId="6" pageNumber="157" type="male">♂</specimenCount>
,
<specimenCount id="45A0FD7E9117F118A500FBA9783C7CE7" box="[580,601,647,673]" count="1" pageId="6" pageNumber="157" type="female">♀</specimenCount>
)’dark morph’),
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,
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,
<collectingMunicipality id="B37DAC8D9117F118A346FBA87E927CE6" box="[1026,1271,646,672]" pageId="6" pageNumber="157">Borujerd Shakhrestan</collectingMunicipality>
,
<location id="5679602C9117F118A245FBA97FF97CE6" LSID="urn:lsid:plazi:treatment:DB0F87E19117F118A7D3F9B97E6D7D76:5679602C9117F118A245FBA97FF97CE6" box="[1281,1436,646,673]" country="Iran" county="Borujerd Shakhrestan" latitude="48.9398" longLatPrecision="6" longitude="48.9402" municipality="Borujerd Shakhrestan" name="Hayan village" pageId="6" pageNumber="157" stateProvince="Lorestan">Hayan village</location>
env.,
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sipring,
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–33.7920° N 48.9398–
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,
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<quantity id="945E9B129117F118A343FB847EC17C83" box="[1031,1188,682,709]" metricMagnitude="3" metricUnit="m" metricValue="1.8225" metricValueMax="1.829" metricValueMin="1.816" pageId="6" pageNumber="157" unit="m" value="1822.5" valueMax="1829.0" valueMin="1816.0">1816–1829 m</quantity>
a.s.l.
</elevation>
,
<date id="271810379117F118A3B5FB847F0A7C83" box="[1265,1391,682,709]" pageId="6" pageNumber="157" value="2017-05-25">
<collectingDate id="375CE9DF9117F118A3B5FB847F0A7C83" box="[1265,1391,682,709]" pageId="6" pageNumber="157" value="2017-05-25">25.05.2017</collectingDate>
</date>
(
<collectorName id="FE5353219117F118A2C5FB847A927CAE" pageId="6" pageNumber="157">O. Kosterin</collectorName>
leg.)
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.
</paragraph>
</subSubSection>
<subSubSection id="1BBC657C9117F118A783FBDF7E6D7D76" pageId="6" pageNumber="157" type="description">
<paragraph id="531936F79117F118A783FBDF7E6D7D76" blockId="6.[151,1436,217,816]" pageId="6" pageNumber="157">
Charlotte Hartong, the Curator of
<taxonomicName id="94A64D749117F118A51FFBDF78DF7D4D" box="[603,698,753,779]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Odonata" pageId="6" pageNumber="157" phylum="Tracheophyta" rank="order">Odonata</taxonomicName>
collection in Naturalis Biodiversity Centre, Leiden (further on RMNH), kindly provided photographs of the male appendages of
<taxonomicName id="94A64D749117F118A43EFA387E677D69" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[890,1026,790,816]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="syriacum">
<emphasis id="61D2EAE59117F118A43EFA387E677D69" box="[890,1026,790,816]" italics="true" pageId="6" pageNumber="157">C. syriacum</emphasis>
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1.
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<emphasis id="61D2EAE59119F116A7F1FF1F7844780A" bold="true" box="[181,545,1585,1612]" italics="true" pageId="8" pageNumber="159">Coenagrion australocaspicum</emphasis>
<bibRefCitation id="37374B069119F116A56CFF1F7905780D" author="Dumont, H. J. &amp; Heidari, H." box="[552,864,1585,1611]" pageId="8" pageNumber="159" pagination="63 - 78" refId="ref27800" refString="Dumont, H. J. &amp; Heidari, H. (1996) On a collection of spring Odonata from Iran, with the description of Coenagrion australocaspicum n. sp. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 132, 63 - 78." type="journal article" year="1996">Dumont &amp; Heidari, 1996</bibRefCitation>
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<figureCitation id="CB9D2A729119F116A7D3FF7B7B637829" box="[151,262,1621,1647]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="8" pageNumber="159">Figs 2a–c</figureCitation>
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<figureCitation id="CB9D2A729119F116A655FF7B7B2C7829" box="[273,329,1621,1647]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="8" pageNumber="159">3a–c</figureCitation>
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<figureCitation id="CB9D2A729119F116A610FF7B7BE87829" box="[340,397,1621,1647]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="8" pageNumber="159">4a–b</figureCitation>
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<figureCitation id="CB9D2A729119F116A6DDFF7B7BD77836" box="[409,434,1621,1648]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="8" pageNumber="159">5f</figureCitation>
,
<figureCitation id="CB9D2A729119F116A6FAFF7B7BAB7829" box="[446,462,1621,1647]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="8" pageNumber="159">6</figureCitation>
,
<figureCitation id="CB9D2A729119F116A69EFF787B907836" box="[474,501,1622,1648]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="8" pageNumber="159">7a</figureCitation>
</paragraph>
<paragraph id="531936F79119F116A7D3FFB37B9B789D" blockId="8.[151,1436,1693,2008]" pageId="8" pageNumber="159">
<emphasis id="61D2EAE59119F116A7D3FFB37B6278F1" bold="true" box="[151,263,1693,1719]" pageId="8" pageNumber="159">Reported</emphasis>
(as
<emphasis id="61D2EAE59119F116A672FFB0787E78F1" box="[310,539,1693,1720]" italics="true" pageId="8" pageNumber="159">
<taxonomicName id="94A64D749119F116A672FFB07BD978FE" authorityName="Kirby" authorityYear="1890" box="[310,444,1694,1720]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">Coenagrion</taxonomicName>
(
<taxonomicName id="94A64D749119F116A68FFFB3787678F1" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[459,531,1693,1719]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="puella">puella</taxonomicName>
)
</emphasis>
sp2)
<emphasis id="61D2EAE59119F116A51EFFB378FB78F1" bold="true" box="[602,670,1693,1719]" pageId="8" pageNumber="159">from:</emphasis>
the Samurskiy Zakaznik Stationary environs, Magaramkent District of
<collectingRegion id="9162F8159119F116A7F2FFEC7B7B789A" box="[182,286,1730,1756]" country="Russia" name="Dagestan" pageId="8" pageNumber="159">Dagestan</collectingRegion>
(
<bibRefCitation id="37374B069119F116A668FFEF7B97789D" author="Ilyina, E. V. &amp; Onishko, V. V. &amp; Kosterin, O. E." box="[300,498,1729,1755]" pageId="8" pageNumber="159" pagination="31 - 50" refId="ref28161" refString="Ilyina, E. V., Onishko, V. V. &amp; Kosterin, O. E. (2022) Strekozy (Odonata) klastera &quot; Del'ta Samura &quot; natsional'nogo parka &quot; Samurskiy &quot; [Dragonflies (Odonata) of the Samur Delta Cluster of Samurskiy National Park']. Trydy Gosudarstvennogo Prirodnogo Zapvednika &quot; Dagestanskiy &quot;, 18, 31 - 50. [in Russian]" type="journal article" year="2022">
Ilyina
<emphasis id="61D2EAE59119F116A632FFEC7BCA789D" box="[374,431,1729,1755]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
<paragraph id="531936F79119F112A783FFCB7BD17E93" blockId="8.[151,1436,1693,2008]" lastBlockId="12.[151,1437,150,2013]" lastPageId="12" lastPageNumber="163" pageId="8" pageNumber="159">
<emphasis id="61D2EAE59119F116A783FFCB7B8F78B9" bold="true" box="[199,490,1765,1791]" pageId="8" pageNumber="159">
Specimens.
<materialsCitation id="E3CE3CAA9119F116A61EFFCB791B7965" ID-GBIF-Occurrence="4897992383" collectingDate="2023-06-08" collectorName="E. Ilyina" county="Babayurt District" elevation="10" latitude="43.612" location="Khamamatyurt village" longLatPrecision="68" longitude="46.455" municipality="Terek River" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
<collectingRegion id="9162F8159119F116A61EFFCB780378B9" box="[346,614,1765,1791]" country="Russia" name="Dagestan" pageId="8" pageNumber="159">
Republic of
<emphasis id="61D2EAE59119F116A6BCFFCB780378B9" bold="true" box="[504,614,1765,1791]" pageId="8" pageNumber="159">Dagestan</emphasis>
</collectingRegion>
:
<specimenCount id="45A0FD7E9119F116A538FFC878F67946" box="[636,659,1766,1792]" count="1" pageId="8" pageNumber="159" type="male">♂</specimenCount>
,
<collectingCounty id="BA784E7B9119F116A5E2FFCB791478B9" box="[678,881,1765,1791]" pageId="8" pageNumber="159">Babayurt District</collectingCounty>
, the
<collectingMunicipality id="B37DAC8D9119F116A4F6FFCB7E5B78B9" box="[946,1086,1765,1791]" pageId="8" pageNumber="159">Terek River</collectingMunicipality>
right bank at
<location id="5679602C9119F116A3B0FFCB7A867965" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C9119F116A3B0FFCB7A867965" county="Babayurt District" latitude="43.612" longLatPrecision="68" longitude="46.455" municipality="Terek River" name="Khamamatyurt village" pageId="8" pageNumber="159" stateProvince="Dagestan">Khamamatyurt village</location>
,
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,
<geoCoordinate id="369250309119F116A62DFE277BBD7962" box="[361,472,1801,1828]" degrees="46.455" direction="east" orientation="longitude" pageId="8" pageNumber="159" precision="55" value="46.455">46.455 E</geoCoordinate>
,
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a.s.l.
</elevation>
,
<date id="271810379119F116A52EFE2778B97965" box="[618,732,1801,1827]" pageId="8" pageNumber="159" value="2023-06-08">
<collectingDate id="375CE9DF9119F116A52EFE2778B97965" box="[618,732,1801,1827]" pageId="8" pageNumber="159" value="2023-06-08">8.06.2023</collectingDate>
</date>
,
<collectorName id="FE5353219119F116A5ADFE24792F7965" box="[745,842,1801,1828]" pageId="8" pageNumber="159">E. Ilyina</collectorName>
leg.
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</emphasis>
;
<materialsCitation id="E3CE3CAA9119F116A4CFFE247B57792D" ID-GBIF-Occurrence="4897992349" collectingDate="2021-05-03" collectorName="E. Ilyina" county="Kumtorkala District" elevation="73" latitude="47.223" location="Kumtorkala Station" longLatPrecision="66" longitude="47.246" municipality="Shura-Ozen' River" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
<specimenCount id="45A0FD7E9119F116A4CFFE2479C77962" box="[907,930,1802,1828]" count="1" pageId="8" pageNumber="159" type="male">♂</specimenCount>
,
<collectingCounty id="BA784E7B9119F116A4EBFE277EF67965" box="[943,1171,1801,1827]" pageId="8" pageNumber="159">Kumtorkala District</collectingCounty>
, the
<collectingMunicipality id="B37DAC8D9119F116A38DFE277FF97965" box="[1225,1436,1801,1827]" pageId="8" pageNumber="159">Shura-Ozen’ River</collectingMunicipality>
at
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, next to
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,
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–43.006 N, 47.223–
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,
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a.s.l.
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,
<date id="271810379119F116A261FE037FFD790E" box="[1317,1432,1837,1864]" pageId="8" pageNumber="159" value="2021-05-03">
<collectingDate id="375CE9DF9119F116A261FE037FFD790E" box="[1317,1432,1837,1864]" pageId="8" pageNumber="159" value="2021-05-03">3.05.2021</collectingDate>
</date>
,
<collectorName id="FE5353219119F116A7D3FE7C7A9E792D" box="[151,251,1873,1900]" pageId="8" pageNumber="159">E. Ilyina</collectorName>
leg.
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;
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(
<figureCitation id="CB9D2A729119F116A62CFE7F7BA5792D" box="[360,448,1873,1899]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="8" pageNumber="159">Figs 2c</figureCitation>
,
<figureCitation id="CB9D2A729119F116A694FE7F7B8F792D" box="[464,490,1873,1899]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="8" pageNumber="159">3c</figureCitation>
) (196433669, 196426386), the same place,
<date id="271810379119F116A4A8FE7F7E0B792D" box="[1004,1134,1873,1899]" pageId="8" pageNumber="159" value="2021-06-29">29.06.2021</date>
;
<materialsCitation id="E3CE3CAA9119F116A338FE7C7B7C79F2" ID-GBIF-Occurrence="4897992379" collectingDate="2023-05-25" elevation="17" latitude="41.9048" location="Derbent District" longLatPrecision="6" longitude="48.4452" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
<specimenCount id="45A0FD7E9119F116A338FE7C7EF4792A" box="[1148,1169,1874,1900]" count="1" pageId="8" pageNumber="159" type="male">♂</specimenCount>
(teneral) (196427366),
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,
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<quantity id="945E9B129119F116A61FFE5B7BCF79C9" box="[347,426,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="159" unit="km" value="3.5">3.5 km</quantity>
NE of Belidzhi village
</locationDeviation>
, thickets at a swamp,
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,
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,
<quantity id="945E9B129119F116A3FAFE5B7F4479D6" box="[1214,1313,1909,1936]" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="8" pageNumber="159" unit="m" value="17.0" valueMax="18.0" valueMin="16.0">
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</quantity>
below s.l.,
<date id="271810379119F116A7D3FEB77B7C79F2" box="[151,281,1945,1972]" pageId="8" pageNumber="159" value="2023-05-25">
<collectingDate id="375CE9DF9119F116A7D3FEB77B7C79F2" box="[151,281,1945,1972]" pageId="8" pageNumber="159" value="2023-05-25">25.05.2023</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="E3CE3CAA9119F115A661FEB77F2278DC" ID-GBIF-Occurrence="4897992339" collectingDate="2021-06-24" county="Magaramkent District" elevation="16" lastPageId="11" lastPageNumber="162" latitude="48.5539" location="Samurskiy Zakaznik Stationary" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" pageId="8" pageNumber="159" specimenCount="13" specimenCount-female="3" specimenCount-male="10" stateProvince="Primorskiy">
<specimenCount id="45A0FD7E9119F116A661FEB77B1679F2" box="[293,371,1945,1972]" count="10" pageId="8" pageNumber="159" type="male">10 ♂♂</specimenCount>
(
<figureCitation id="CB9D2A729119F116A6C7FEB77B9379F5" box="[387,502,1945,1971]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="8" pageNumber="159">Figs 2a–b</figureCitation>
,
<figureCitation id="CB9D2A729119F116A546FEB7785E79F5" box="[514,571,1945,1971]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="8" pageNumber="159">3a–b</figureCitation>
,
<figureCitation id="CB9D2A729119F116A503FEB778E579F5" box="[583,640,1945,1971]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="8" pageNumber="159">4a–b</figureCitation>
,
<figureCitation id="CB9D2A729119F116A5C9FEB778C279F2" box="[653,679,1945,1972]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="8" pageNumber="159">5f</figureCitation>
),
<specimenCount id="45A0FD7E9119F116A5F9FEB7789879F2" box="[701,765,1945,1972]" count="3" pageId="8" pageNumber="159" type="female">3 ♀♀</specimenCount>
(
<figureCitation id="CB9D2A729119F116A449FEB7793879F5" box="[781,861,1945,1971]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="8" pageNumber="159">Fig. 7a</figureCitation>
) (196425390, 196425415, 196425436, 196425479 (
<figureCitation id="CB9D2A729119F116A7DBFE937A9C7991" box="[159,249,1981,2007]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="8" pageNumber="159">
Fig.
<quantity id="945E9B129119F116A79FFE937A9C7991" box="[219,249,1981,2007]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="8" pageNumber="159" unit="g" value="6.0">6g</quantity>
</figureCitation>
), 196425508, 196425543, 196425590, 196425634, 196425653, 196425691, 196425722, 196425755, 196425780, 196425808, 196425854, 196425868, 196425900),
<collectingCounty id="BA784E7B911AF115A42DFF167E007814" box="[873,1125,1592,1618]" pageId="11" pageNumber="162">Magaramkent District</collectingCounty>
,
<collectingMunicipality id="B37DAC8D911AF115A336FF177F317815" box="[1138,1364,1592,1619]" pageId="11" pageNumber="162">Samur Delta Forest</collectingMunicipality>
,
<locationDeviation id="42AB2FD5911AF115A225FF177B0E7830" location="Primorskiy" pageId="11" pageNumber="162">
2 km NW of
<collectingRegion id="9162F815911AF115A7ABFF727B0E7830" box="[239,363,1628,1654]" country="Russia" name="Primorskiy" pageId="11" pageNumber="162">Primorskiy</collectingRegion>
</locationDeviation>
village, environs of
<location id="5679602C911AF115A51FFF7279A47830" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911AF115A51FFF7279A47830" box="[603,961,1628,1654]" county="Magaramkent District" latitude="48.5539" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" name="Samurskiy Zakaznik Stationary" pageId="11" pageNumber="162" stateProvince="Primorskiy">Samurskiy Zakaznik Stationary</location>
, a swampy overgrown estuary of a forest rivulet and a nearby forest road,
<geoCoordinate id="36925030911AF115A556FFAE780878DD" box="[530,621,1664,1691]" degrees="41.8619" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="41.8619">41.8619</geoCoordinate>
–41.8650 N, 48.5539–
<geoCoordinate id="36925030911AF115A430FFAE799678DD" box="[884,1011,1664,1691]" degrees="48.5592" direction="east" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="48.5592">48.5592 E</geoCoordinate>
,
<quantity id="945E9B12911AF115A345FFAE7E5878DD" box="[1025,1085,1664,1691]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="162" unit="m" value="16.0">
<elevation id="D88BD1C4911AF115A345FFAE7E5878DD" box="[1025,1085,1664,1691]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="162" unit="m" value="16.0">16 m</elevation>
</quantity>
below s.l.,
<date id="27181037911AF115A381FFAE7F2278DC" box="[1221,1351,1664,1690]" pageId="11" pageNumber="162" value="2021-06-24">
<collectingDate id="375CE9DF911AF115A381FFAE7F2278DC" box="[1221,1351,1664,1690]" pageId="11" pageNumber="162" value="2021-06-24">24.06.2021</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="45A0FD7E911AF115A210FFAF7FF278DD" box="[1364,1431,1665,1691]" count="8" pageId="11" pageNumber="162" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="45A0FD7E911AF115A7D3FF8B7ABC78F9" box="[151,217,1701,1727]" count="7" pageId="11" pageNumber="162" type="female">7 ♀♀</specimenCount>
(196427413, 196427432, 196427528, 196427547 (
<figureCitation id="CB9D2A72911AF115A46DFF8A791B78F8" box="[809,894,1700,1726]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6h</figureCitation>
), 196427578, 196432921 (
<figureCitation id="CB9D2A72911AF115A3F1FF8A7F6278F8" box="[1205,1287,1700,1726]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6f</figureCitation>
); 196503424 (
<figureCitation id="CB9D2A72911AF115A7DBFFE67A9178A4" box="[159,244,1736,1762]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6c</figureCitation>
)), the same place,
<geoCoordinate id="36925030911AF115A693FFE6785778A5" box="[471,562,1736,1763]" degrees="41.8634" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="41.8634">41.8634</geoCoordinate>
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–
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</geoCoordinate>
N,
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<geoCoordinate id="36925030911AF115A46BFFE679F278A5" box="[815,919,1736,1763]" degrees="48.5563" direction="west" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="-48.5563">
–
<geoCoordinate id="36925030911AF115A479FFE679F278A5" box="[829,919,1736,1763]" degrees="48.5563" direction="east" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="48.5563">48.5563</geoCoordinate>
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E,
<quantity id="945E9B12911AF115A489FFE77E5178A5" box="[973,1076,1737,1763]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="11" pageNumber="162" unit="m" value="19.0" valueMax="20.0" valueMin="18.0">18–20 m</quantity>
below s.l.,
<date id="27181037911AF115A385FFE67F2678A5" box="[1217,1347,1736,1763]" pageId="11" pageNumber="162" value="2023-05-26">26.05.2023</date>
;
<specimenCount id="45A0FD7E911AF115A216FFE77FF278A5" box="[1362,1431,1737,1763]" count="4" pageId="11" pageNumber="162" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="45A0FD7E911AF115A7D3FFC37ABC7941" box="[151,217,1773,1799]" count="2" pageId="11" pageNumber="162" type="female">2 ♀♀</specimenCount>
(196430322 (
<figureCitation id="CB9D2A72911AF115A63FFFC27BAC7940" box="[379,457,1772,1798]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6i</figureCitation>
), 196430351, 196430424, 196430513, 196430593, 196430622 (
<figureCitation id="CB9D2A72911AF115A3EFFFC27F657940" box="[1195,1280,1772,1798]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6d</figureCitation>
), 196430665, 196430702, 196430744, 196430788, 196430823, 196430891, 196431009, 196431069, 196431104, 196431208, 196431263, 196431305, 196431337, 196431381, 196433323 (
<figureCitation id="CB9D2A72911AF115A43FFE1A79B17908" box="[891,980,1844,1870]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6a</figureCitation>
), 196431926, 196431963, 196432017, 196432083, 196432115, 196432155), the same place,
<date id="27181037911AF115A5AFFE7679087935" box="[747,877,1880,1907]" pageId="11" pageNumber="162" value="2023-05-27">27.05.2023</date>
;
<materialsCitation id="E3CE3CAA911AF115A432FE777B517998" ID-GBIF-Occurrence="4897992344" collectingDate="2022-06-28" collectorName="E. Ilyina" county="Magaramkent District" elevation="16" latitude="48.5494" location="Samurskiy Zakaznik Stationary" longLatPrecision="6" longitude="48.5674" municipality="Samur Delta Forest" pageId="11" pageNumber="162" specimenCount="2" specimenCount-male="2" stateProvince="Primorskiy">
<specimenCount id="45A0FD7E911AF115A432FE7779D07935" box="[886,949,1881,1907]" count="2" pageId="11" pageNumber="162" type="male">2 ♂♂</specimenCount>
,
<collectingCounty id="BA784E7B911AF115A4FBFE767ED37934" box="[959,1206,1880,1906]" pageId="11" pageNumber="162">Magaramkent District</collectingCounty>
,
<collectingMunicipality id="B37DAC8D911AF115A3FBFE777FFD7935" box="[1215,1432,1880,1907]" pageId="11" pageNumber="162">Samur Delta Forest</collectingMunicipality>
,
<locationDeviation id="42AB2FD5911AF115A7D3FE537BCA79D0" box="[151,431,1916,1943]" location="Primorskiy" pageId="11" pageNumber="162">
<quantity id="945E9B12911AF115A7D3FE537AB779D0" box="[151,210,1916,1943]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="162" unit="km" value="3.0">3 km</quantity>
NW of
<collectingRegion id="9162F815911AF115A677FE527BCA79D0" box="[307,431,1916,1942]" country="Russia" name="Primorskiy" pageId="11" pageNumber="162">Primorskiy</collectingRegion>
</locationDeviation>
village, environs of
<location id="5679602C911AF115A5DBFE527E6079D0" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911AF115A5DBFE527E6079D0" box="[671,1029,1916,1942]" county="Magaramkent District" latitude="48.5494" longLatPrecision="6" longitude="48.5674" municipality="Samur Delta Forest" name="Samurskiy Zakaznik Stationary" pageId="11" pageNumber="162" stateProvince="Primorskiy">Samurskiy Zakaznik Stationary</location>
(broadly the same place as above), a coastal lake and nearby meadows,
<geoCoordinate id="36925030911AF115A506FE8E78F879FD" box="[578,669,1952,1979]" degrees="41.8524" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="41.8524">41.8524</geoCoordinate>
–41.8669 N, 48.5494–
<geoCoordinate id="36925030911AF115A4E2FE8E7E4079FD" box="[934,1061,1952,1979]" degrees="48.5674" direction="east" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="48.5674">48.5674 E</geoCoordinate>
,
<quantity id="945E9B12911AF115A371FE8F7EE879FD" box="[1077,1165,1953,1979]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="2.9" metricValueMin="0.2" pageId="11" pageNumber="162" unit="m" value="15.5" valueMax="29.0" valueMin="2.0">
<elevation id="D88BD1C4911AF115A371FE8F7EE879FD" box="[1077,1165,1953,1979]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="2.9" metricValueMin="0.2" pageId="11" pageNumber="162" unit="m" value="15.5" valueMax="29.0" valueMin="2.0">2–29 m</elevation>
</quantity>
below s.l.,
<date id="27181037911AF115A253FE8E7FFD79FC" box="[1303,1432,1952,1978]" pageId="11" pageNumber="162" value="2022-06-28">
<collectingDate id="375CE9DF911AF115A253FE8E7FFD79FC" box="[1303,1432,1952,1978]" pageId="11" pageNumber="162" value="2022-06-28">28.06.2022</collectingDate>
</date>
,
<collectorName id="FE535321911AF115A7D3FEEB7A997998" box="[151,252,1988,2015]" pageId="11" pageNumber="162">E. Ilyina</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="E3CE3CAA911AF112A600FEEB7BD57E93" ID-GBIF-Occurrence="4897992324" collectingDate="2023-05-26" elevation="15" lastPageId="12" lastPageNumber="163" latitude="41.871" location="Samur Delta forest" longLatPrecision="6" longitude="48.5382" municipality="Magaramkent District" pageId="11" pageNumber="162" specimenCount="15" specimenCount-female="1" specimenCount-male="14" stateProvince="Primorskiy">
<specimenCount id="45A0FD7E911AF115A600FEEB7BFD7999" box="[324,408,1989,2015]" count="14" pageId="11" pageNumber="162" type="male">14 ♂♂</specimenCount>
,
<specimenCount id="45A0FD7E911AF115A6ECFEEB7BD87999" box="[424,445,1989,2015]" count="1" pageId="11" pageNumber="162" type="female">♀</specimenCount>
(196432912, 196432913, 196432914, 196432915, 196432917 (
<figureCitation id="CB9D2A72911AF115A3E2FEEA7E9B7998" box="[1190,1278,1988,2014]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6b</figureCitation>
), 196432918, 196432920),
<collectingMunicipality id="B37DAC8D911DF112A668F9B978437EF7" box="[300,550,151,177]" pageId="12" pageNumber="163">Magaramkent District</collectingMunicipality>
,
<location id="5679602C911DF112A574F9B979627EF7" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A574F9B979627EF7" box="[560,775,151,177]" latitude="41.871" longLatPrecision="6" longitude="48.5382" municipality="Magaramkent District" name="Samur Delta forest" pageId="12" pageNumber="163" stateProvince="Primorskiy">Samur Delta forest</location>
, at the 700 years old plane tree,
<geoCoordinate id="36925030911DF112A32AF9B97E8B7EF7" box="[1134,1262,151,177]" degrees="41.8710" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="5" value="41.871">41.8710 N</geoCoordinate>
,
<geoCoordinate id="36925030911DF112A3BDF9B87F117EF7" box="[1273,1396,150,177]" degrees="48.5382" direction="east" orientation="longitude" pageId="12" pageNumber="163" precision="5" value="48.5382">48.5382 E</geoCoordinate>
,
<quantity id="945E9B12911DF112A2C4F9B87AC87E93" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="15.0">
<elevation id="D88BD1C4911DF112A2C4F9B87AC87E93" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="15.0">15 m</elevation>
</quantity>
below s.l.,
<date id="27181037911DF112A66AF9947BD57E93" box="[302,432,186,213]" pageId="12" pageNumber="163" value="2023-05-26">
<collectingDate id="375CE9DF911DF112A66AF9947BD57E93" box="[302,432,186,213]" pageId="12" pageNumber="163" value="2023-05-26">26.05.2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="07D9667F9118F117A7D3FCA379CE79A2" ID-DOI="http://doi.org/10.5281/zenodo.12189910" ID-Zenodo-Dep="12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="9" pageNumber="160" startId="9.[151,250,1421,1445]" targetBox="[381,1200,187,1391]" targetPageId="9" targetType="figure">
<paragraph id="531936F79118F117A7D3FCA379CE79A2" blockId="9.[151,1437,1421,2021]" pageId="9" pageNumber="160">
<emphasis id="61D2EAE59118F117A7D3FCA37B717BE3" bold="true" box="[151,276,1421,1445]" pageId="9" pageNumber="160">FIGURE 2.</emphasis>
Male appendages of
<taxonomicName id="94A64D749118F117A6B7FCA378FB7BE2" box="[499,670,1421,1444]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="61D2EAE59118F117A6B7FCA378097BE2" box="[499,620,1421,1444]" italics="true" pageId="9" pageNumber="160">Coenagrion</emphasis>
spp.
</taxonomicName>
of the
<taxonomicName id="94A64D749118F117A5ADFCA3794D7BE3" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[745,808,1421,1445]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE59118F117A5ADFCA3794D7BE3" box="[745,808,1421,1445]" italics="true" pageId="9" pageNumber="160">puella</emphasis>
</taxonomicName>
-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—
<taxonomicName id="94A64D749118F117A63FFC9F78357B8F" authorityName="Dumont &amp; Heidari" authorityYear="1996" box="[379,592,1457,1481]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE59118F117A63FFC9F78357B8F" box="[379,592,1457,1481]" italics="true" pageId="9" pageNumber="160">C. australocaspicum</emphasis>
</taxonomicName>
(b—with the appendages expanded); d-k—
<taxonomicName id="94A64D749118F117A35AFC9F7E1B7B8F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1054,1150,1457,1481]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE59118F117A35AFC9F7E1B7B8F" box="[1054,1150,1457,1481]" italics="true" pageId="9" pageNumber="160">C. puella</emphasis>
</taxonomicName>
; l–m—
<taxonomicName id="94A64D749118F117A394FC9F7F2A7B8F" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[1232,1359,1457,1481]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="ponticum">
<emphasis id="61D2EAE59118F117A394FC9F7F2A7B8F" box="[1232,1359,1457,1481]" italics="true" pageId="9" pageNumber="160">C. ponticum</emphasis>
</taxonomicName>
(males with a more (c) and less (d) expanded black abdominal maculation); n—
<taxonomicName id="94A64D749118F117A4E6FCFB7E457BAB" authorityName="Lohmann" authorityYear="1983" box="[930,1056,1493,1517]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE59118F117A4E6FCFB7E457BAB" box="[930,1056,1493,1517]" italics="true" pageId="9" pageNumber="160">C. persicum</emphasis>
</taxonomicName>
; o–p—
<taxonomicName id="94A64D749118F117A335FCFB7F667BAB" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1137,1283,1493,1517]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE59118F117A335FCFB7F667BAB" box="[1137,1283,1493,1517]" italics="true" pageId="9" pageNumber="160">C. pulchellum</emphasis>
</taxonomicName>
; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.
</paragraph>
</caption>
<caption id="07D9667F911BF114A7D3FCB67BA779A9" ID-DOI="http://doi.org/10.5281/zenodo.12189912" ID-Zenodo-Dep="12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1432,1456]" targetBox="[393,1180,188,1398]" targetPageId="10" targetType="figure">
<paragraph id="531936F7911BF114A7D3FCB67BA779A9" blockId="10.[151,1437,1432,2032]" pageId="10" pageNumber="161">
<emphasis id="61D2EAE5911BF114A7D3FCB67B757BF6" bold="true" box="[151,272,1432,1456]" pageId="10" pageNumber="161">FIGURE 3.</emphasis>
Male appendages of
<taxonomicName id="94A64D74911BF114A69AFCB678E07BE9" box="[478,645,1432,1455]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="61D2EAE5911BF114A69AFCB678327BE9" box="[478,599,1432,1455]" italics="true" pageId="10" pageNumber="161">Coenagrion</emphasis>
spp.
</taxonomicName>
of the
<taxonomicName id="94A64D74911BF114A587FCB679677BF6" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[707,770,1432,1456]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911BF114A587FCB679677BF6" box="[707,770,1432,1456]" italics="true" pageId="10" pageNumber="161">puella</emphasis>
</taxonomicName>
-group in the broad sense in lateral view: a–c—
<taxonomicName id="94A64D74911BF114A389FCB67FF97BF6" authorityName="Dumont &amp; Heidari" authorityYear="1996" box="[1229,1436,1432,1456]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE5911BF114A389FCB67FF97BF6" box="[1229,1436,1432,1456]" italics="true" pageId="10" pageNumber="161">C. australocaspicum</emphasis>
</taxonomicName>
(b—with the appendages expanded); d–k—
<taxonomicName id="94A64D74911BF114A513FC9278D07B92" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[599,693,1468,1492]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911BF114A513FC9278D07B92" box="[599,693,1468,1492]" italics="true" pageId="10" pageNumber="161">C. puella</emphasis>
</taxonomicName>
; l–m—
<taxonomicName id="94A64D74911BF114A441FC9279E77B92" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[773,898,1468,1492]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="ponticum">
<emphasis id="61D2EAE5911BF114A441FC9279E77B92" box="[773,898,1468,1492]" italics="true" pageId="10" pageNumber="161">C. ponticum</emphasis>
</taxonomicName>
(males with a more (c) and less (d) expanded black abdominal maculation); n—
<taxonomicName id="94A64D74911BF114A6FFFCCE785D7BBE" authorityName="Lohmann" authorityYear="1983" box="[443,568,1504,1528]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE5911BF114A6FFFCCE785D7BBE" box="[443,568,1504,1528]" italics="true" pageId="10" pageNumber="161">C. persicum</emphasis>
</taxonomicName>
; o–q—
<taxonomicName id="94A64D74911BF114A5CCFCCE797C7BBE" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[648,793,1504,1528]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE5911BF114A5CCFCCE797C7BBE" box="[648,793,1504,1528]" italics="true" pageId="10" pageNumber="161">C. pulchellum</emphasis>
</taxonomicName>
; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.
</paragraph>
</caption>
<caption id="07D9667F911AF115A7D3FD30789A7BA0" ID-DOI="http://doi.org/10.5281/zenodo.12189914" ID-Zenodo-Dep="12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1054,1078]" targetBox="[189,1398,181,1021]" targetPageId="11" targetType="figure">
<paragraph id="531936F7911AF115A7D3FD30789A7BA0" blockId="11.[151,1437,1054,1511]" pageId="11" pageNumber="162">
<emphasis id="61D2EAE5911AF115A7D3FD307B7C7A70" bold="true" box="[151,281,1054,1078]" pageId="11" pageNumber="162">FIGURE 4.</emphasis>
Male appendages of
<taxonomicName id="94A64D74911AF115A548FD3178D97A70" box="[524,700,1055,1078]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="61D2EAE5911AF115A548FD3178E07A70" box="[524,645,1055,1078]" italics="true" pageId="11" pageNumber="162">Coenagrion</emphasis>
spp.
</taxonomicName>
of the
<taxonomicName id="94A64D74911AF115A451FD3079317A70" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[789,852,1054,1078]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911AF115A451FD3079317A70" box="[789,852,1054,1078]" italics="true" pageId="11" pageNumber="162">puella</emphasis>
</taxonomicName>
-group in the broad sense in posterior view: a–b—
<taxonomicName id="94A64D74911AF115A2C1FD317B2E7A1C" authorityName="Dumont &amp; Heidari" authorityYear="1996" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE5911AF115A2C1FD317B2E7A1C" italics="true" pageId="11" pageNumber="162">C. australocaspicum</emphasis>
</taxonomicName>
(b—with the appendages expanded); c–d—
<taxonomicName id="94A64D74911AF115A466FD6D79C77A1C" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[802,930,1091,1114]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="ponticum">
<emphasis id="61D2EAE5911AF115A466FD6D79C77A1C" box="[802,930,1091,1114]" italics="true" pageId="11" pageNumber="162">C. ponticum</emphasis>
</taxonomicName>
(males with a more (c) and less (d) expanded black abdominal maculation); e-f—
<taxonomicName id="94A64D74911AF115A548FD4978FB7A38" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[524,670,1126,1150]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE5911AF115A548FD4978FB7A38" box="[524,670,1126,1150]" italics="true" pageId="11" pageNumber="162">C. pulchellum</emphasis>
</taxonomicName>
; g–k—
<taxonomicName id="94A64D74911AF115A5AAFD49792A7A38" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[750,847,1126,1150]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911AF115A5AAFD49792A7A38" box="[750,847,1126,1150]" italics="true" pageId="11" pageNumber="162">C. puella</emphasis>
</taxonomicName>
; l—
<taxonomicName id="94A64D74911AF115A4C4FD49799B7A38" authorityName="Lohmann" authorityYear="1983" box="[896,1022,1127,1150]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE5911AF115A4C4FD49799B7A38" box="[896,1022,1127,1150]" italics="true" pageId="11" pageNumber="162">C. persicum</emphasis>
</taxonomicName>
; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="531936F7911DF112A783F9F079137CEF" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
<emphasis id="61D2EAE5911DF112A783F9F0797D7EBE" bold="true" box="[199,792,222,248]" pageId="12" pageNumber="163">
Additional photographic registrations.
<materialsCitation id="E3CE3CAA911DF112A5D5F9F07BB27F23" ID-GBIF-Occurrence="4897992307" collectingDate="2021-06-24" county="Magaramkent District" elevation="17" latitude="48.5576" location="Samurskiy Zakaznik Stationary" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" pageId="12" pageNumber="163" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Dagestan">
<collectingRegion id="9162F815911DF112A5D5F9F079EE7EBF" box="[657,907,222,249]" country="Russia" name="Dagestan" pageId="12" pageNumber="163">
Republic of
<emphasis id="61D2EAE5911DF112A459F9F179EE7EBF" bold="true" box="[797,907,223,249]" pageId="12" pageNumber="163">Dagestan</emphasis>
</collectingRegion>
:
<specimenCount id="45A0FD7E911DF112A4D3F9F179A67EBF" box="[919,963,223,249]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
,
<specimenCount id="45A0FD7E911DF112A489F9F179927EBF" box="[973,1015,223,249]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
(196426531, 196426558, 196426592, 196426638),
<collectingCounty id="BA784E7B911DF112A66AF82D784C7F5B" box="[302,553,259,285]" pageId="12" pageNumber="163">Magaramkent District</collectingCounty>
,
<collectingMunicipality id="B37DAC8D911DF112A571F82D79707F5B" box="[565,789,259,285]" pageId="12" pageNumber="163">Samur Delta Forest</collectingMunicipality>
,
<locationDeviation id="42AB2FD5911DF112A466F82D7E527F5B" box="[802,1079,259,285]" location="Primorskiy" pageId="12" pageNumber="163">2 km NW of Primorskiy</locationDeviation>
village, environs of
<location id="5679602C911DF112A261F82D7B1C7F07" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A261F82D7B1C7F07" county="Magaramkent District" latitude="48.5576" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" name="Samurskiy Zakaznik Stationary" pageId="12" pageNumber="163" stateProvince="Dagestan">Samurskiy Zakaznik Stationary</location>
, a swampy overgrown estuary of a forest rivulet,
<geoCoordinate id="36925030911DF112A4D0F809798A7F07" box="[916,1007,295,321]" degrees="41.8619" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="5" value="41.8619">41.8619</geoCoordinate>
–41.8630 N, 48.5576–
<geoCoordinate id="36925030911DF112A3A9F8087F0D7F07" box="[1261,1384,294,321]" degrees="48.5592" direction="east" orientation="longitude" pageId="12" pageNumber="163" precision="5" value="48.5592">48.5592 E</geoCoordinate>
,
<quantity id="945E9B12911DF112A236F8097AB47F23" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="12" pageNumber="163" unit="m" value="17.0" valueMax="18.0" valueMin="16.0">
<elevation id="D88BD1C4911DF112A236F8097AB47F23" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="12" pageNumber="163" unit="m" value="17.0" valueMax="18.0" valueMin="16.0">16– 18 m</elevation>
</quantity>
below s.l.,
<date id="27181037911DF112A611F8657BB27F23" box="[341,471,331,357]" pageId="12" pageNumber="163" value="2021-06-24">
<collectingDate id="375CE9DF911DF112A611F8657BB27F23" box="[341,471,331,357]" pageId="12" pageNumber="163" value="2021-06-24">24.06.2021</collectingDate>
</date>
</materialsCitation>
</emphasis>
;
<specimenCount id="45A0FD7E911DF112A6A7F865786A7F23" box="[483,527,331,357]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
,
<specimenCount id="45A0FD7E911DF112A558F86578237F23" box="[540,582,331,357]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
(196425929, 196425961, 196426001, 196426083, 196426117, 196426145, 196426182 (
<figureCitation id="CB9D2A72911DF112A669F8417BE47FCF" box="[301,385,367,393]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6j</figureCitation>
), 196426298, 196426322, 196432235, 196432236, 196432237, 196432239, 196432240, 196432241, 196432243, 196432244, 196432245), the same place,
<date id="27181037911DF112A4CDF8BD7E6E7FEB" box="[905,1035,403,429]" pageId="12" pageNumber="163" value="2021-06-26">26.06.2021</date>
;
<specimenCount id="45A0FD7E911DF112A353F8BD7E267FEB" box="[1047,1091,403,429]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
,
<specimenCount id="45A0FD7E911DF112A30BF8BD7E1C7FEB" box="[1103,1145,403,429]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
(196431432, 196431467, 196431500, 196431519, 196431595, 196431732, 196431756, 196431799 (
<figureCitation id="CB9D2A72911DF112A4AFF8997E587F97" box="[1003,1085,439,465]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6e</figureCitation>
)), the same place,
<date id="27181037911DF112A252F8987FFD7F97" box="[1302,1432,438,465]" pageId="12" pageNumber="163" value="2023-05-28">28.05.2023</date>
;
<materialsCitation id="E3CE3CAA911DF112A7D3F8F57E6E7C5F" ID-GBIF-Occurrence="4897992317" collectingDate="2021-06-26" elevation="29" latitude="41.8668" location="Samur Delta Forest" longLatPrecision="41" longitude="48.558" municipality="Magaramkent District" pageId="12" pageNumber="163" specimenCount="1" stateProvince="Primorskiy">
copula (196432247,
<figureCitation id="CB9D2A72911DF112A6C0F8F57BBC7FB3" box="[388,473,475,501]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6k</figureCitation>
)),
<collectingMunicipality id="B37DAC8D911DF112A6BFF8F578927FB3" box="[507,759,475,501]" pageId="12" pageNumber="163">Magaramkent District</collectingMunicipality>
,
<location id="5679602C911DF112A441F8F5798D7FB3" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A441F8F5798D7FB3" box="[773,1000,475,501]" latitude="41.8668" longLatPrecision="41" longitude="48.558" municipality="Magaramkent District" name="Samur Delta Forest" pageId="12" pageNumber="163" stateProvince="Primorskiy">Samur Delta Forest</location>
,
<locationDeviation id="42AB2FD5911DF112A4B2F8F57F777FB3" box="[1014,1298,475,501]" location="Primorskiy" pageId="12" pageNumber="163">
<quantity id="945E9B12911DF112A4B2F8F57E567FB3" box="[1014,1075,475,501]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="163" unit="km" value="3.0">3 km</quantity>
NW of
<collectingRegion id="9162F815911DF112A3D2F8F57F777FB3" box="[1174,1298,475,501]" country="Russia" name="Primorskiy" pageId="12" pageNumber="163">Primorskiy</collectingRegion>
</locationDeviation>
village, the SE bank of a coastal lake,
<geoCoordinate id="36925030911DF112A680F8D178207C5F" box="[452,581,511,537]" degrees="41.8668" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="5" value="41.8668">41.8668 N</geoCoordinate>
,
<geoCoordinate id="36925030911DF112A515F8D078DA7C5F" box="[593,703,510,537]" degrees="48.558" direction="east" orientation="longitude" pageId="12" pageNumber="163" precision="55" value="48.558">48.558 E</geoCoordinate>
,
<quantity id="945E9B12911DF112A588F8D179637C5F" box="[716,774,511,537]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="163" unit="m" value="29.0">
<elevation id="D88BD1C4911DF112A588F8D179637C5F" box="[716,774,511,537]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="163" unit="m" value="29.0">29 m</elevation>
</quantity>
below s.l.,
<date id="27181037911DF112A4CDF8D17E6E7C5F" box="[905,1035,511,537]" pageId="12" pageNumber="163" value="2021-06-26">
<collectingDate id="375CE9DF911DF112A4CDF8D17E6E7C5F" box="[905,1035,511,537]" pageId="12" pageNumber="163" value="2021-06-26">26.06.2021</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="E3CE3CAA911DF112A353F8D179177CEF" ID-GBIF-Occurrence="4897992336" collectingDate="2021-06-26" elevation="20" location="Samur Delta Forest" municipality="Magaramkent District" pageId="12" pageNumber="163" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Primorskiy">
<specimenCount id="45A0FD7E911DF112A353F8D17E267C5F" box="[1047,1091,511,537]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
,
<specimenCount id="45A0FD7E911DF112A30BF8D17E1C7C5F" box="[1103,1145,511,537]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
(196427626, 196428813, 196428891, 196428956, 196428980, 196429008, 196429029, 196429061, 196429094, 196429172, 196429235, 196429278, 196429336, 196429394, 196429463, 196429541, 196429578, 196429724, 196429748),
<collectingMunicipality id="B37DAC8D911DF112A245FB697A8B7CC3" pageId="12" pageNumber="163">Magaramkent District</collectingMunicipality>
,
<location id="5679602C911DF112A7BEFB457BB27CC3" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A7BEFB457BB27CC3" box="[250,471,619,645]" municipality="Magaramkent District" name="Samur Delta Forest" pageId="12" pageNumber="163" stateProvince="Primorskiy">Samur Delta Forest</location>
,
<locationDeviation id="42AB2FD5911DF112A6A7FB44796D7CC3" box="[483,776,618,645]" location="Primorskiy" pageId="12" pageNumber="163">
<quantity id="945E9B12911DF112A6A7FB4478547CC3" box="[483,561,618,645]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="163" unit="km" value="3.5">3.5 km</quantity>
NW of
<collectingRegion id="9162F815911DF112A5C8FB45796D7CC3" box="[652,776,619,645]" country="Russia" name="Primorskiy" pageId="12" pageNumber="163">Primorskiy</collectingRegion>
</locationDeviation>
village, bushy meadows at the NW bank of a coastal lake, 41.8709–8755 N, 48.5409–5459 E,
<quantity id="945E9B12911DF112A573FBA178157CEF" box="[567,624,655,681]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="163" unit="m" value="20.0">
<elevation id="D88BD1C4911DF112A573FBA178157CEF" box="[567,624,655,681]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="163" unit="m" value="20.0">20 m</elevation>
</quantity>
below s.l.,
<date id="27181037911DF112A5B4FBA179177CEF" box="[752,882,655,681]" pageId="12" pageNumber="163" value="2021-06-26">
<collectingDate id="375CE9DF911DF112A5B4FBA179177CEF" box="[752,882,655,681]" pageId="12" pageNumber="163" value="2021-06-26">26.06.2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="531936F7911DF112A783FB9C79017DAB" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
<emphasis id="61D2EAE5911DF112A783FB9C7B317C8A" bold="true" box="[199,340,690,716]" pageId="12" pageNumber="163">Characters.</emphasis>
Male cercus trunk in dorsal view (
<figureCitation id="CB9D2A72911DF112A595FB9D795F7C8B" box="[721,826,691,717]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Fig. 2a–c</figureCitation>
) evenly thick, with a very gentle convection in place of bulge (so their bases do not touch each other), rather long. In lateral view (
<figureCitation id="CB9D2A72911DF112A48FFBF97E517CB7" box="[971,1076,727,753]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">Fig. 3a–c</figureCitation>
) it is smoothly rounded, without an expressed apex. Tubercle in dorsal view (
<figureCitation id="CB9D2A72911DF112A5CBFBD5789F7D53" box="[655,762,763,789]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Fig. 2a–c</figureCitation>
) of the same width as trunk and looks like its continuation, rather elongate, in lateral view (
<figureCitation id="CB9D2A72911DF112A6B1FA31783B7D7F" box="[501,606,799,825]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">Fig. 3a–c</figureCitation>
) always well seen, never hidden behind paraprocts. (It should be noted that the specimen shown in
<figureCitation id="CB9D2A72911DF112A6E4FA6D7B837D1B" box="[416,486,835,861]" captionStart="FIGURE 1" captionStartId="7.[151,250,790,814]" captionTargetBox="[201,1386,187,756]" captionTargetId="figure-19@7.[201,1386,179,764]" captionTargetPageId="7" captionText="FIGURE 1. Explanation of terminology (see the text) used for morphological details of male terminalia of Coenagrion spp. of the puella-group in the broad sense used in this paper, on examples of a male specimen of C. australocaspicum with an unusual expanded position of the cerci in lateral view (left, the same as Fig. 3a) and a male of C. persicum in dorsal view (right, the same as Fig. 2n)." figureDoi="http://doi.org/10.5281/zenodo.12189908" httpUri="https://zenodo.org/record/12189908/files/figure.png" pageId="12" pageNumber="163">Figs 1</figureCitation>
(left), 2b and 3b has dead with abnormally expanded cerci, while
<figureCitation id="CB9D2A72911DF112A395FA6D7F527D1B" box="[1233,1335,835,861]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Figs 2a,c</figureCitation>
and
<figureCitation id="CB9D2A72911DF112A22AFA6D7FF97D1B" box="[1390,1436,835,861]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">3a,c</figureCitation>
show their normal downward position. Nevertheless, the above described characters are well seen in
<figureCitation id="CB9D2A72911DF112A3BCFA497F2E7DC7" box="[1272,1355,871,897]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Figs 2b</figureCitation>
and
<figureCitation id="CB9D2A72911DF112A2C4FA497FF97DC7" box="[1408,1436,871,897]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">3b</figureCitation>
as well.) Tooth small, normally not seen in dorsal view unless cerci are unusually expanded as in
<figureCitation id="CB9D2A72911DF112A398FAA57F4A7DE3" box="[1244,1327,907,933]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Fig. 2b</figureCitation>
, disposed at ca 3/4 of the trunk length. Process broadly triangular, somewhat hooked inside apically but rather blunt, reaching about the middle of the height of the paraproct base (
<figureCitation id="CB9D2A72911DF112A5ACFAFD793D7DAB" box="[744,856,979,1005]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="12" pageNumber="163">Fig. 4a–b</figureCitation>
).
</paragraph>
<paragraph id="531936F7911DF112A783FAD97E237A3B" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
Paraproct spine very long, directed strictly caudad in lateral view (
<figureCitation id="CB9D2A72911DF112A4E9FAD97E727A57" box="[941,1047,1015,1041]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">Fig. 3a–c</figureCitation>
) where is ca 5/6 as long as S10 (as measured along the line of its dorsal side); spines diverging and slightly hooked in at tips dorsal view. Prominence weakly expressed as resembling a rounded right angle, incision between spine and prominense not expressed. S
<quantity id="945E9B12911DF112A2C4FD117AC87A3B" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="163" unit="in" value="10.0">10 in</quantity>
caudal view looks more flattened above (
<figureCitation id="CB9D2A72911DF112A5C5FD4D78887A3B" box="[641,749,1123,1149]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="12" pageNumber="163">Fig. 4a–b</figureCitation>
) as compared to other species.
</paragraph>
<paragraph id="531936F7911DF112A783FDA97BD07A83" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
Male pronotum hind margin almost straight, with negligible, very shallow lateral depressions and a very slight median incision (
<figureCitation id="CB9D2A72911DF112A61DFD857BCD7A83" box="[345,424,1194,1221]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="12" pageNumber="163">Fig. 5f</figureCitation>
).
</paragraph>
<paragraph id="531936F7911DF112A783FDE178CE7B77" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
Male abdomen is somewhat relatively longer than in
<taxonomicName id="94A64D74911DF112A456FDE179127AAF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[786,887,1231,1257]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911DF112A456FDE179127AAF" box="[786,887,1231,1257]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
</taxonomicName>
, coloured as in that species but S9 blue with (
<figureCitation id="CB9D2A72911DF112A22BFDE17ABF7B4B" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6a, 6c</figureCitation>
) or without (
<figureCitation id="CB9D2A72911DF112A62DFDDD7BDC7B4B" box="[361,441,1267,1293]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6b</figureCitation>
) a pair of black dots or (rarely) black triangular marks (
<figureCitation id="CB9D2A72911DF112A359FDDD7E0B7B4B" box="[1053,1134,1267,1293]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6c</figureCitation>
), but never with a posterior black bar. S10 black dorsally and blue laterally.
</paragraph>
<paragraph id="531936F7911DF112A783FC15797B7BDB" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
Female pronotum hind lobe margin almost straight when seen flat, perpendicularly to its surface, that is anteriodorsally with respect to the insect body axis, (
<figureCitation id="CB9D2A72911DF112A599FC7179487B3F" box="[733,813,1375,1401]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="12" pageNumber="163">Fig. 7a</figureCitation>
, below), but is slightly wavy as can be seen in the dorsal view with respect to the insect body axis (
<figureCitation id="CB9D2A72911DF112A529FCAD78DA7BDB" box="[621,703,1411,1437]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="12" pageNumber="163">Fig. 7a</figureCitation>
, above).
</paragraph>
<paragraph id="531936F7911DF112A783FC897F3F7937" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
As usual in the genus, females are polymorphic for the coloration pattern. Most frequent is the dark morph which resembles the corresponding morph in
<taxonomicName id="94A64D74911DF112A508FCE578D67BA3" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[588,691,1483,1509]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911DF112A508FCE578D67BA3" box="[588,691,1483,1509]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
</taxonomicName>
, with most of the abdomen dorsum black, with only expression of pale rings at S8–10 varying to complete disappearance (
<figureCitation id="CB9D2A72911DF112A44EFCC1791C784F" box="[778,889,1519,1545]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6d–e</figureCitation>
). Less frequent is a spotty morph, also paralleled by the analogous morph of
<taxonomicName id="94A64D74911DF112A68DFF3D784A786B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[457,559,1555,1581]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911DF112A68DFF3D784A786B" box="[457,559,1555,1581]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
</taxonomicName>
, with anterior parts of most abdominal segments pale (
<figureCitation id="CB9D2A72911DF112A3D1FF3D7F64786B" box="[1173,1281,1555,1581]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6f–h</figureCitation>
). However, in these morphs, the pale ground colour occupies a larger area on S8 than in
<taxonomicName id="94A64D74911DF112A49FFF197E277817" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[987,1090,1591,1617]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911DF112A49FFF197E277817" box="[987,1090,1591,1617]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
</taxonomicName>
, varying from a large bipartite anterior pale spots (
<figureCitation id="CB9D2A72911DF112A6C4FF757B827833" box="[384,487,1627,1653]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6f–j</figureCitation>
) to, rather infrequently, S8 being pale with a black streak (
<figureCitation id="CB9D2A72911DF112A3E0FF757E9E7833" box="[1188,1275,1627,1653]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6h</figureCitation>
), often of the horseshoe shape—this version resembles females of rather
<taxonomicName id="94A64D74911DF112A47FFF5179BC78DF" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[827,985,1663,1689]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE5911DF112A47FFF5179BC78DF" box="[827,985,1663,1689]" italics="true" pageId="12" pageNumber="163">C. pulchellum</emphasis>
</taxonomicName>
than
<taxonomicName id="94A64D74911DF112A35FFF517EE678DF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1051,1155,1663,1689]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911DF112A35FFF517EE678DF" box="[1051,1155,1663,1689]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
</taxonomicName>
. The extreme version of the spotty morph are extra-pale females the maculation of which rather resembles males of
<emphasis id="61D2EAE5911DF112A3CBFF8D7FF978FB" box="[1167,1436,1699,1725]" italics="true" pageId="12" pageNumber="163">Enallahma cyathigerum</emphasis>
(
<bibRefCitation id="37374B06911DF112A7DBFFE97B0878A7" author="Charpentier, T." box="[159,365,1735,1761]" pageId="12" pageNumber="163" refId="ref27675" refString="Charpentier, T. (de) (1840) Libellulinae europaeae descripta ac depictae. L. Voss, Lipsiae, 181 pp. https: // doi. org / 10.5962 / t. 173037" type="book" year="1840">Charpentier, 1840</bibRefCitation>
): with a black arrowhead mark on S2, black spots in the posterior parts of S2–7, producing one to three processes in the anterior direction, a paired black mark on S8 and S9–10 black at dorsum (
<figureCitation id="CB9D2A72911DF112A3F8FFC57F6D7943" box="[1212,1288,1771,1797]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6i</figureCitation>
). The ground pale colour varies (
<figureCitation id="CB9D2A72911DF112A634FE217BBE796F" box="[368,475,1807,1833]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6d–i</figureCitation>
), at least in part with age, from pale greyish (
<figureCitation id="CB9D2A72911DF112A49BFE217E57796F" box="[991,1074,1807,1833]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6h</figureCitation>
), sometime pale orange, to blue (
<figureCitation id="CB9D2A72911DF112A7DBFE1D7B69790B" box="[159,268,1843,1869]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6e, g</figureCitation>
), often this is grey but with the anteriodorsal part of the abdominal segments blue (
<figureCitation id="CB9D2A72911DF112A3F5FE1D7E9A790B" box="[1201,1279,1843,1869]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6i</figureCitation>
). However, in mature blue females, the antehumeral stripes are usually green, as well as the eye lower part (
<figureCitation id="CB9D2A72911DF112A3E9FE797F2B7937" box="[1197,1358,1879,1905]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6e, g, j–k</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1BBC657C911DF10EA783FE547F207CEF" lastPageId="16" lastPageNumber="167" pageId="12" pageNumber="163" type="distribution">
<paragraph id="531936F7911DF10EA783FE547EC27F23" blockId="12.[151,1437,150,2013]" lastBlockId="16.[151,1437,151,681]" lastPageId="16" lastPageNumber="167" pageId="12" pageNumber="163">
<emphasis id="61D2EAE5911DF112A783FE547B9C79D2" bold="true" box="[199,505,1914,1941]" pageId="12" pageNumber="163">Distribution and habitat.</emphasis>
Found in the Russian Caucasus only in
<collectingRegion id="9162F815911DF112A49EFE557E2079D3" box="[986,1093,1915,1941]" country="Russia" name="Dagestan" pageId="12" pageNumber="163">Dagestan</collectingRegion>
, in the coastal plains of both southern (Samur Delta Forest) and northern (Khamamatyurt)
<collectingRegion id="9162F815911DF112A47BFEB179C279FF" box="[831,935,1951,1977]" country="Russia" name="Dagestan" pageId="12" pageNumber="163">Dagestan</collectingRegion>
and in the foothills near the famous Sarykum Barkhan (the Shura-Ozen’ River). However, the Khamamatyurt locality is situated at the right bank of the Terek River which serves there as the border with Chechen Republic, where the species certainly occurs as well. The latter is the northermost known locality of this species in the world, of
<geoCoordinate id="369250309101F10EA43BF99579AB7E93" box="[895,974,187,213]" degrees="43.6" direction="north" orientation="latitude" pageId="16" pageNumber="167" precision="5555" value="43.6">43.6 N</geoCoordinate>
. Beyond
<collectingCountry id="2BB176679101F10EA37EF9957EED7E93" box="[1082,1160,187,213]" name="Russia" pageId="16" pageNumber="167">Russia</collectingCountry>
, the species is known to occur along the southern coast of the Caspian Sea in
<collectingCountry id="2BB176679101F10EA5B8F9F1794C7EBF" box="[764,809,223,249]" name="Iran" pageId="16" pageNumber="167">Iran</collectingCountry>
and in the coastal Lankoran and Kachmaz Districts of
<collectingCountry id="2BB176679101F10EA7D3F82D7B777F5B" box="[151,274,259,285]" name="Azerbaijan" pageId="16" pageNumber="167">Azerbaijan</collectingCountry>
(
<bibRefCitation id="37374B069101F10EA665F82D78597F5B" author="Dumont, H. J. &amp; Heidari, H." box="[289,572,259,285]" pageId="16" pageNumber="167" pagination="63 - 78" refId="ref27800" refString="Dumont, H. J. &amp; Heidari, H. (1996) On a collection of spring Odonata from Iran, with the description of Coenagrion australocaspicum n. sp. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 132, 63 - 78." type="journal article" year="1996">Dumont &amp; Heidari 1996</bibRefCitation>
;
<bibRefCitation id="37374B069101F10EA50DF82D788C7F5B" author="Dumont, H. J." box="[585,745,259,285]" pageId="16" pageNumber="167" pagination="87 - 92" refId="ref27765" refString="Dumont, H. J. (2004) Dragonflies from Azerbaijan. Zoology in the Middle East, 31, 87 - 92. https: // doi. org / 10.1080 / 09397140.2004.10638027" type="journal article" year="2004">Dumont 2004</bibRefCitation>
;
<bibRefCitation id="37374B069101F10EA5B1F82D79AE7F5B" author="Boudot, J. - P. &amp; Borisov, S. &amp; De Knijf, G. &amp; Van Grunsven, R. &amp; Schroter, A. &amp; Kalkman, V. J." box="[757,971,259,285]" pageId="16" pageNumber="167" pagination="3 - 248" refId="ref27573" refString="Boudot, J. - P., Borisov, S., De Knijf, G., Van Grunsven, R., Schroter, A. &amp; Kalkman, V. J. (2021) Atlas of the dragonflies and damselflies of West and Central Asia. Brachytron, 22, Supplement, 3 - 248." type="journal article" year="2021">
Boudot
<emphasis id="61D2EAE59101F10EA40BF82D79EC7F5B" box="[847,905,259,285]" italics="true" pageId="16" pageNumber="167">et al.</emphasis>
2021
</bibRefCitation>
;
<bibRefCitation id="37374B069101F10EA49CF82D7EFD7F5B" author="Snegovaya, N. Y." box="[984,1176,259,285]" pageId="16" pageNumber="167" pagination="1 - 33" refId="ref29587" refString="Snegovaya, N. Y. (2024) Odonata collected in 2023 in Azerbaijan, including the confirmation of Cordulia aenea Linnaeus, 1758 for Azerbaijan. International Dragonfly Fund Report, 186, 1 - 33." type="journal article" year="2024">Snegovaya 2024</bibRefCitation>
). In the latter district it was near Şollar (
<bibRefCitation id="37374B069101F10EA612F8097B937F07" author="Dumont, H. J." box="[342,502,295,321]" pageId="16" pageNumber="167" pagination="87 - 92" refId="ref27765" refString="Dumont, H. J. (2004) Dragonflies from Azerbaijan. Zoology in the Middle East, 31, 87 - 92. https: // doi. org / 10.1080 / 09397140.2004.10638027" type="journal article" year="2004">Dumont 2004</bibRefCitation>
) and Narban (
<bibRefCitation id="37374B069101F10EA5DDF809793D7F07" author="Snegovaya, N. Y." box="[665,856,295,321]" pageId="16" pageNumber="167" pagination="1 - 33" refId="ref29587" refString="Snegovaya, N. Y. (2024) Odonata collected in 2023 in Azerbaijan, including the confirmation of Cordulia aenea Linnaeus, 1758 for Azerbaijan. International Dragonfly Fund Report, 186, 1 - 33." type="journal article" year="2024">Snegovaya 2024</bibRefCitation>
) villages, the latter being situated near the Russian border, just
<quantity id="945E9B129101F10EA659F8657B017F23" box="[285,356,331,357]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="167" unit="km" value="12.0">12 km</quantity>
south-east from our closest findings of this species in Samur Delta Forest.
</paragraph>
<caption id="07D9667F911CF113A7D3FF0C7B9B79AF" ID-DOI="http://doi.org/10.5281/zenodo.12189916" ID-Zenodo-Dep="12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,250,1570,1594]" targetBox="[151,1436,181,1541]" targetPageId="13" targetType="figure">
<paragraph id="531936F7911CF113A7D3FF0C7B9B79AF" blockId="13.[151,1437,1570,2026]" pageId="13" pageNumber="164">
<emphasis id="61D2EAE5911CF113A7D3FF0C7B72787C" bold="true" box="[151,279,1570,1594]" pageId="13" pageNumber="164">FIGURE 5.</emphasis>
Male pronotum of
<taxonomicName id="94A64D74911CF113A6A9FF0C78FE787F" box="[493,667,1570,1593]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="61D2EAE5911CF113A6A9FF0C7803787F" box="[493,614,1570,1593]" italics="true" pageId="13" pageNumber="164">Coenagrion</emphasis>
spp.
</taxonomicName>
of the
<taxonomicName id="94A64D74911CF113A5AAFF0C7948787C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[750,813,1570,1594]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911CF113A5AAFF0C7948787C" box="[750,813,1570,1594]" italics="true" pageId="13" pageNumber="164">puella</emphasis>
</taxonomicName>
-group in the broad sense in dorsal view: a—
<taxonomicName id="94A64D74911CF113A253FF0C7FF0787C" authorityName="Lohmann" authorityYear="1983" box="[1303,1429,1570,1594]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE5911CF113A253FF0C7FF0787C" box="[1303,1429,1570,1594]" italics="true" pageId="13" pageNumber="164">C. persicum</emphasis>
</taxonomicName>
; b–c—
<taxonomicName id="94A64D74911CF113A791FF687B027818" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[213,359,1606,1630]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE5911CF113A791FF687B027818" box="[213,359,1606,1630]" italics="true" pageId="13" pageNumber="164">C. pulchellum</emphasis>
</taxonomicName>
; d–e—
<taxonomicName id="94A64D74911CF113A6F0FF6878567818" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[436,563,1606,1630]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="ponticum">
<emphasis id="61D2EAE5911CF113A6F0FF6878567818" box="[436,563,1606,1630]" italics="true" pageId="13" pageNumber="164">C. ponticum</emphasis>
</taxonomicName>
(males with a more (d) and less (e) expanded black abdominal maculation); f—
<taxonomicName id="94A64D74911CF113A2C1FF687B2E78C4" authorityName="Dumont &amp; Heidari" authorityYear="1996" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE5911CF113A2C1FF687B2E78C4" italics="true" pageId="13" pageNumber="164">C. australocaspicum</emphasis>
</taxonomicName>
; g–l—
<taxonomicName id="94A64D74911CF113A6D0FF447B9778C4" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[404,498,1642,1666]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911CF113A6D0FF447B9778C4" box="[404,498,1642,1666]" italics="true" pageId="13" pageNumber="164">C. puella</emphasis>
</taxonomicName>
; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, KarachayCherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.
</paragraph>
</caption>
<caption id="07D9667F911FF110A7D3FFFF7ECA793F" ID-DOI="http://doi.org/10.5281/zenodo.12189920" ID-Zenodo-Dep="12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,250,1745,1769]" targetBox="[151,1436,181,1721]" targetPageId="14" targetType="figure">
<paragraph id="531936F7911FF110A7D3FFFF7ECA793F" blockId="14.[151,1437,1745,1913]" pageId="14" pageNumber="165">
<emphasis id="61D2EAE5911FF110A7D3FFFF7B7778AF" bold="true" box="[151,274,1745,1769]" pageId="14" pageNumber="165">FIGURE 6.</emphasis>
Males (a–c), females (d–i) and copulae (j–k) of
<taxonomicName id="94A64D74911FF110A5BFFFFC7E4B78AF" authorityName="Dumont &amp; Heidari" authorityYear="1996" box="[763,1070,1745,1769]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE5911FF110A5BFFFFC7E4B78AF" box="[763,1070,1745,1769]" italics="true" pageId="14" pageNumber="165">Coenagrion australocaspicum</emphasis>
</taxonomicName>
in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.
</paragraph>
</caption>
<caption id="07D9667F911EF111A7D3FF33794979A3" ID-DOI="http://doi.org/10.5281/zenodo.12189924" ID-Zenodo-Dep="12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="15" pageNumber="166" startId="15.[151,250,1565,1589]" targetBox="[447,1140,184,1541]" targetPageId="15" targetType="figure">
<paragraph id="531936F7911EF111A7D3FF33794979A3" blockId="15.[151,1437,1565,2022]" pageId="15" pageNumber="166">
<emphasis id="61D2EAE5911EF111A7D3FF337B717873" bold="true" box="[151,276,1565,1589]" pageId="15" pageNumber="166">FIGURE 7.</emphasis>
Female pronotum of
<taxonomicName id="94A64D74911EF111A6B0FF3378FA7873" box="[500,671,1565,1589]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="61D2EAE5911EF111A6B0FF3378087872" box="[500,621,1565,1588]" italics="true" pageId="15" pageNumber="166">Coenagrion</emphasis>
spp.
</taxonomicName>
of the
<taxonomicName id="94A64D74911EF111A5ACFF3379427873" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[744,807,1565,1589]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911EF111A5ACFF3379427873" box="[744,807,1565,1589]" italics="true" pageId="15" pageNumber="166">puella</emphasis>
</taxonomicName>
-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—
<taxonomicName id="94A64D74911EF111A7BEFF4B7BB5783B" authorityName="Dumont &amp; Heidari" authorityYear="1996" box="[250,464,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE5911EF111A7BEFF4B7BB5783B" box="[250,464,1637,1661]" italics="true" pageId="15" pageNumber="166">C. australocaspicum</emphasis>
</taxonomicName>
; b–g—
<taxonomicName id="94A64D74911EF111A565FF4B78E4783B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[545,641,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE5911EF111A565FF4B78E4783B" box="[545,641,1637,1661]" italics="true" pageId="15" pageNumber="166">C. puella</emphasis>
</taxonomicName>
; h—
<taxonomicName id="94A64D74911EF111A5FDFF4B795C783B" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[697,825,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="ponticum">
<emphasis id="61D2EAE5911EF111A5FDFF4B795C783B" box="[697,825,1637,1661]" italics="true" pageId="15" pageNumber="166">C. ponticum</emphasis>
</taxonomicName>
; i—
<taxonomicName id="94A64D74911EF111A42FFF4B798C783B" authorityName="Lohmann" authorityYear="1983" box="[875,1001,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="61D2EAE5911EF111A42FFF4B798C783B" box="[875,1001,1637,1661]" italics="true" pageId="15" pageNumber="166">C. persicum</emphasis>
</taxonomicName>
; j—
<taxonomicName id="94A64D74911EF111A35FFF4B7ECB783B" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1051,1198,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="61D2EAE5911EF111A35FFF4B7ECB783B" box="[1051,1198,1637,1661]" italics="true" pageId="15" pageNumber="166">C. pulchellum</emphasis>
</taxonomicName>
; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="531936F79101F10EA783F8417F207CEF" blockId="16.[151,1437,151,681]" pageId="16" pageNumber="167">
These damselflies were invariably found near slow flowing overgrown brooks. They often concentrate at forest roads and margins (and also around an old plane tree surrounded by a meadow in dozen metres apart from water). A very abundant population was found in the Samur Delta Forest, composed of tall oak and hornbeam trees woven with many lianas, which grow, below the conventional sea level at the Caspian Sea coast, in the semidesert zone because the delta of the large Samur River forms a number of fast and cold arms locally called ‘Karasu’. In that forest, this species seems to nearly segregate with the visually very similar
<taxonomicName id="94A64D749101F10EA4BFFB0D7E017C7B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1019,1124,547,573]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE59101F10EA4BFFB0D7E017C7B" box="[1019,1124,547,573]" italics="true" pageId="16" pageNumber="167">C. puella</emphasis>
</taxonomicName>
, with just
<specimenCount id="45A0FD7E9101F10EA3A4FB0D7F197C7B" box="[1248,1404,547,573]" count="1" pageId="16" pageNumber="167" type="generic">one specimen</specimenCount>
of
<taxonomicName id="94A64D749101F10EA7D3FB697A9B7C27" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,254,583,609]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE59101F10EA7D3FB697A9B7C27" box="[151,254,583,609]" italics="true" pageId="16" pageNumber="167">C. puella</emphasis>
</taxonomicName>
found (on
<date id="271810379101F10EA63FFB687B9E7C27" box="[379,507,582,609]" pageId="16" pageNumber="167" value="2023-05-26">26.05.2023</date>
) among dozens of
<taxonomicName id="94A64D749101F10EA595FB6979DC7C27" authorityName="Dumont &amp; Heidari" authorityYear="1996" box="[721,953,583,609]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE59101F10EA595FB6979DC7C27" box="[721,953,583,609]" italics="true" pageId="16" pageNumber="167">C. australocaspicum</emphasis>
</taxonomicName>
at the Samurskiy Zakaznik Stationary, and none among numerous representatives of
<taxonomicName id="94A64D749101F10EA537FB4579397CC3" authorityName="Dumont &amp; Heidari" authorityYear="1996" box="[627,860,619,645]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="australocaspicum">
<emphasis id="61D2EAE59101F10EA537FB4579397CC3" box="[627,860,619,645]" italics="true" pageId="16" pageNumber="167">C. australocaspicum</emphasis>
</taxonomicName>
at the famous old plane tree, while only
<taxonomicName id="94A64D749101F10EA270FB457FF97CC3" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1332,1436,619,645]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="61D2EAE59101F10EA270FB457FF97CC3" box="[1332,1436,619,645]" italics="true" pageId="16" pageNumber="167">C. puella</emphasis>
</taxonomicName>
was present at a brook in Bil’-Bil’ Kazmalyar village just
<quantity id="945E9B129101F10EA45BFBA0793D7CEF" box="[799,856,654,681]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="167" unit="km" value="5.0">5 km</quantity>
SSE of the Samurskiy Zakaznik Stationary.
</paragraph>
</subSubSection>
</treatment>
</document>

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data/DB/28/A2/DB28A21D921DFF82FF18FF59FA14FA35.xml
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@ -1,83 +1,85 @@
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