Add updates up until 2024-06-21 15:47:58

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<mods:title id="ABD32F9AAB477844321C54E1ECDDCDE4">Review of the ground beetle subgenus Micronialoe from Korea, with a description of a new species (Coleoptera: Carabidae: Pterostichus)</mods:title>
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<mods:namePart id="2955EDD4684FACABE13744CE746D6A9D">Kim, Dooyoung</mods:namePart>
<mods:affiliation id="170CBF907264ECF406C3413C3125656B">Department of Biomedical Convergence Science and Technology, School of Industrial Technology Advances, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<mods:namePart id="1D4BC9FAEBDF408D43B07ECFC6C6615E">Suh, Sang Jae</mods:namePart>
<mods:affiliation id="BAA855E3EC27A65D0611AB1656B3D9DD">Department of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea &amp; Institute of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<mods:date id="3554601BF23D59B51C497A2BB7E37219">2024</mods:date>
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<treatment id="135E87AA5217FFEE2997F92AFF08494F" ID-DOI="http://doi.org/10.5281/zenodo.12190874" ID-GBIF-Taxon="233318009" ID-Zenodo-Dep="12190874" LSID="urn:lsid:plazi:treatment:135E87AA5217FFEE2997F92AFF08494F" httpUri="http://treatment.plazi.org/id/135E87AA5217FFEE2997F92AFF08494F" lastPageId="10" lastPageNumber="461" pageId="8" pageNumber="459">
<subSubSection id="D3ED65375217FFEC2997F92AFDF24B6B" pageId="8" pageNumber="459" type="nomenclature">
<paragraph id="9B4836BC5217FFEC2997F92AFC3F4B46" blockId="8.[151,977,1770,1868]" box="[151,977,1770,1797]" pageId="8" pageNumber="459">
<heading id="C00081D05217FFEC2997F92AFC3F4B46" bold="true" box="[151,977,1770,1797]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
<taxonomicName id="5CF74D3F5217FFEC2997F92AFC3F4B46" ID-CoL="4Q3X5" authority="Park, Kwon &amp; Lafer, 1996" authorityName="Park, Kwon &amp; Lafer" authorityYear="1996" box="[151,977,1770,1797]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="chogyesanus" subGenus="Micronialoe">
<emphasis id="A983EAAE5217FFEC2997F92AFC3F4B46" bold="true" box="[151,977,1770,1797]" pageId="8" pageNumber="459">
<emphasis id="A983EAAE5217FFEC2997F92AFEC34B47" bold="true" box="[151,301,1770,1797]" italics="true" pageId="8" pageNumber="459">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5217FFEC283EF92AFE3C4B47" bold="true" box="[318,466,1770,1797]" italics="true" pageId="8" pageNumber="459">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5217FFEC28E4F92AFD914B47" bold="true" box="[484,639,1770,1797]" italics="true" pageId="8" pageNumber="459">chogyesanus</emphasis>
<bibRefCitation id="FF664B4D5217FFEC2B86F92AFC3F4B46" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[646,977,1770,1796]" pageId="8" pageNumber="459" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9B4836BC5217FFEC2997F8CFFDF24B6B" blockId="8.[151,977,1770,1868]" box="[151,540,1806,1833]" pageId="8" pageNumber="459">Korean name: 송광길쭉먼지벌레</paragraph>
</subSubSection>
<subSubSection id="D3ED65375217FFEC2997F8F2FEE64B0E" box="[151,264,1842,1868]" pageId="8" pageNumber="459" type="description">
<paragraph id="9B4836BC5217FFEC2997F8F2FEE64B0E" blockId="8.[151,977,1770,1868]" box="[151,264,1842,1868]" pageId="8" pageNumber="459">
(
<figureCitation id="03CC2A395217FFEC299FF8F2FF114B0E" box="[159,255,1842,1868]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="8" pageNumber="459">Figure 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375217FFEC2997F8B7FBA04BB9" pageId="8" pageNumber="459" type="reference_group">
<paragraph id="9B4836BC5217FFEC2997F8B7FC9D4BCC" blockId="8.[151,883,1910,1935]" box="[151,883,1910,1935]" pageId="8" pageNumber="459">
<treatmentCitationGroup id="BBE711925217FFEC2997F8B7FC9D4BCC" box="[151,883,1910,1935]" pageId="8" pageNumber="459">
<treatmentCitation id="1A5610AD5217FFEC2997F8B7FC804BCC" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[151,878,1910,1935]" page="75" pageId="8" pageNumber="459" year="1996">
<taxonomicName id="5CF74D3F5217FFEC2997F8B7FC804BCC" ID-CoL="4Q3X5" authority="Park, Kwon &amp; Lafer, 1996: 75" authorityName="Park, Kwon &amp; Lafer" authorityPageNumber="75" authorityYear="1996" box="[151,878,1910,1935]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="chogyesanus" subGenus="Micronialoe">
<emphasis id="A983EAAE5217FFEC2997F8B7FEFB4BCD" box="[151,277,1911,1935]" italics="true" pageId="8" pageNumber="459">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5217FFEC2824F8B7FE4E4BCD" box="[292,416,1911,1935]" italics="true" pageId="8" pageNumber="459">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5217FFEC28AFF8B7FDDE4BCD" box="[431,560,1911,1935]" italics="true" pageId="8" pageNumber="459">chogyesanus</emphasis>
<bibRefCitation id="FF664B4D5217FFEC2B37F8B7FC804BCC" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[567,878,1910,1935]" pageId="8" pageNumber="459" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996: 75</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9B4836BC5217FFEC2997F87CFBA04BB9" blockId="8.[151,1436,1980,2044]" pageId="8" pageNumber="459">
<emphasis id="A983EAAE5217FFEC2997F87CFE7A4B94" bold="true" box="[151,404,1980,2006]" pageId="8" pageNumber="459">Specimen examined.</emphasis>
<materialsCitation id="2B9F3CE15217FFEC28A4F87DFBA64BB9" ID-GBIF-Occurrence="4898139303" collectingDate="2022-09-03" collectorName="Dooyoung Kim" country="South Korea" county="Gurye-gun" elevation="580" latitude="35.132164" location="Jungdae-ri" longLatPrecision="1" longitude="127.6035" municipality="Ganjeon-myeon" pageId="8" pageNumber="459" specimenCount="5" specimenCount-male="5" stateProvince="Jeonnam">
<collectingCountry id="E3E0762C5217FFEC28A4F87DFE064B95" box="[420,488,1981,2007]" name="South Korea" pageId="8" pageNumber="459">Korea</collectingCountry>
<specimenCount id="8DF1FD355217FFEC2B11F87DFDD94B9A" box="[529,567,1981,2008]" count="5" pageId="8" pageNumber="459" type="male">5♂</specimenCount>
;
<collectingRegion id="5933F85E5217FFEC2B4CF87DFD084B95" box="[588,742,1981,2007]" country="South Korea" name="Jeonnam" pageId="8" pageNumber="459">Jeollanam-do</collectingRegion>
,
<collectingCounty id="72294E305217FFEC2BF9F87DFC9B4B95" box="[761,885,1981,2007]" pageId="8" pageNumber="459">Gurye-gun</collectingCounty>
,
<collectingMunicipality id="7B2CACC65217FFEC2A89F87DFBAE4B95" box="[905,1088,1981,2007]" pageId="8" pageNumber="459">Ganjeon-myeon</collectingMunicipality>
,
<location id="9E2860675217FFEC2D55F87DFB214B95" LSID="urn:lsid:plazi:treatment:135E87AA5217FFEE2997F92AFF08494F:9E2860675217FFEC2D55F87DFB214B95" box="[1109,1231,1981,2007]" country="South Korea" county="Gurye-gun" latitude="35.132164" longLatPrecision="1" longitude="127.6035" municipality="Ganjeon-myeon" name="Jungdae-ri" pageId="8" pageNumber="459" stateProvince="Jeonnam">Jungdae-ri</location>
;
<location id="9E2860675217FFEC2DE2F87DFA764B95" LSID="urn:lsid:plazi:treatment:135E87AA5217FFEE2997F92AFF08494F:9E2860675217FFEC2DE2F87DFA764B95" box="[1250,1432,1981,2007]" country="South Korea" county="Gurye-gun" latitude="35.132164" longLatPrecision="1" longitude="127.6035" municipality="Ganjeon-myeon" name="Mt. Baegunsan" pageId="8" pageNumber="459" stateProvince="Jeonnam">Mt. Baegunsan</location>
;
<geoCoordinate id="FEC3507B5217FFEC2997F821FEDE4BBE" box="[151,304,2017,2044]" degrees="35" direction="north" minutes="07" orientation="latitude" pageId="8" pageNumber="459" precision="1" seconds="55.8" value="35.132164">35°07'55.8&quot;N</geoCoordinate>
<geoCoordinate id="FEC3507B5217FFEC2837F821FE304BB9" box="[311,478,2017,2043]" degrees="127" direction="east" minutes="36" orientation="longitude" pageId="8" pageNumber="459" precision="1" seconds="12.6" value="127.6035">127°36'12.6&quot;E</geoCoordinate>
; alt. ca
<quantity id="5C0F9B595217FFEC2B33F821FD934BB9" box="[563,637,2017,2044]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="8" pageNumber="459" unit="m" value="580.0">
<elevation id="10DAD18F5217FFEC2B33F821FD934BB9" box="[563,637,2017,2044]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="8" pageNumber="459" unit="m" value="580.0">580 m</elevation>
</quantity>
;
<date id="EF49107C5217FFEC2B89F821FCFC4BB9" box="[649,786,2017,2043]" pageId="8" pageNumber="459" value="2022-09-03">
<collectingDate id="FF0DE9945217FFEC2B89F821FCFC4BB9" box="[649,786,2017,2043]" pageId="8" pageNumber="459" value="2022-09-03">3 Sep. 2022</collectingDate>
</date>
;
<collectorName id="3602536A5217FFEC2A1DF821FC224BB9" box="[797,972,2017,2043]" pageId="8" pageNumber="459">Dooyoung Kim</collectorName>
leg.; KNU
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="CF8866345216FFED2997FA78FE944A22" ID-DOI="http://doi.org/10.5281/zenodo.12190405" ID-Zenodo-Dep="12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="9" pageNumber="460" startId="9.[151,250,1464,1488]" targetBox="[161,1436,190,1431]" targetPageId="9" targetType="figure">
<paragraph id="9B4836BC5216FFED2997FA78FE944A22" blockId="9.[151,1437,1464,1632]" pageId="9" pageNumber="460">
<emphasis id="A983EAAE5216FFED2997FA78FEFB4992" bold="true" box="[151,277,1464,1488]" pageId="9" pageNumber="460">FIGURE 8.</emphasis>
Male habitus and genitalia of
<taxonomicName id="5CF74D3F5216FFED2B54FA78FB3A4992" authority="Park, Kwon &amp; Lafer." authorityName="Park, Kwon &amp; Lafer." box="[596,1236,1464,1488]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="chogyesanus" subGenus="Micronialoe">
<emphasis id="A983EAAE5216FFED2B54FA78FD3C4992" box="[596,722,1464,1488]" italics="true" pageId="9" pageNumber="460">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5216FFED2BE2FA78FCB04992" box="[738,862,1464,1488]" italics="true" pageId="9" pageNumber="460">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5216FFED2A6FFA78FC1E4992" box="[879,1008,1464,1488]" italics="true" pageId="9" pageNumber="460">chogyesanus</emphasis>
Park, Kwon &amp; Lafer.
</taxonomicName>
<emphasis id="A983EAAE5216FFED2DDBFA78FB114992" bold="true" box="[1243,1279,1464,1488]" pageId="9" pageNumber="460">(A)</emphasis>
habitus, dorsal view;
<emphasis id="A983EAAE5216FFED29D5FA1CFF1949B6" bold="true" box="[213,247,1500,1524]" pageId="9" pageNumber="460">(B)</emphasis>
habitus, ventral view;
<emphasis id="A983EAAE5216FFED28DBFA1CFE1049B6" bold="true" box="[475,510,1500,1524]" pageId="9" pageNumber="460">(C)</emphasis>
head and pronotum, dorsal view;
<emphasis id="A983EAAE5216FFED2A53FA1CFC9949B6" bold="true" box="[851,887,1500,1524]" pageId="9" pageNumber="460">(D)</emphasis>
sternite VII;
<emphasis id="A983EAAE5216FFED2AFBFA1CFBF049B6" bold="true" box="[1019,1054,1500,1524]" pageId="9" pageNumber="460">(E)</emphasis>
median lobe of aedeagus, right lateral view;
<emphasis id="A983EAAE5216FFED29D6F9C0FF194A5A" bold="true" box="[214,247,1536,1560]" pageId="9" pageNumber="460">(F)</emphasis>
median lobe of aedeagus, dorsal view;
<emphasis id="A983EAAE5216FFED2B88F9C0FD434A5A" bold="true" box="[648,685,1536,1560]" pageId="9" pageNumber="460">(G)</emphasis>
median lobe of aedeagus, ventral view;
<emphasis id="A983EAAE5216FFED2D47F9C0FB824A5A" bold="true" box="[1095,1132,1536,1560]" pageId="9" pageNumber="460">(H)</emphasis>
median lobe of aedeagus, left lateral view;
<emphasis id="A983EAAE5216FFED281BF9E4FED84A7E" bold="true" box="[283,310,1572,1596]" pageId="9" pageNumber="460">(I)</emphasis>
left paramere;
<emphasis id="A983EAAE5216FFED28D0F9E4FE004A7E" bold="true" box="[464,494,1572,1596]" pageId="9" pageNumber="460">(J)</emphasis>
right paramere. Scale bars: 1 mm
<emphasis id="A983EAAE5216FFED2A4AF9E4FC954A7E" bold="true" box="[842,891,1572,1596]" pageId="9" pageNumber="460">AC</emphasis>
; 0.5 mm
<emphasis id="A983EAAE5216FFED2ADCF9E4FC004A7E" bold="true" box="[988,1006,1572,1596]" pageId="9" pageNumber="460">D</emphasis>
; 0.3 mm
<emphasis id="A983EAAE5216FFED2D4FF9E4FB6F4A7E" bold="true" box="[1103,1153,1572,1596]" pageId="9" pageNumber="460">EH</emphasis>
; 0.2 mm
<emphasis id="A983EAAE5216FFED2DE1F9E4FAEB4A7E" bold="true" box="[1249,1285,1572,1596]" pageId="9" pageNumber="460">IJ</emphasis>
. Abbreviation: CS: C-shaped sclerite.
</paragraph>
</caption>
<subSubSection id="D3ED65375216FFEE29C7F972FB364C97" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460" type="diagnosis">
<paragraph id="9B4836BC5216FFED29C7F972FD994B7A" blockId="9.[151,1437,1714,2029]" pageId="9" pageNumber="460">
<emphasis id="A983EAAE5216FFED29C7F972FEAC4A8E" bold="true" box="[199,322,1714,1740]" pageId="9" pageNumber="460">Diagnosis.</emphasis>
This species is easily distinguished from known species of the subgenus by: i) pronotum with rudimentary or without outer basal fovea, ii) posteromedian area of male sternite VII not depressed, iii) median lobe of aedeagus slender, without ventral carina; and iv) internal sac of aedeagus with one sclerite, which is C-shaped. Other significant characters are as follows.
</paragraph>
<paragraph id="9B4836BC5216FFED29C7F883FD6B4BE6" blockId="9.[151,1437,1714,2029]" pageId="9" pageNumber="460">Pronotum with weak or rather distinct sinuation on lateral margin before basal angle. Posteromedian area with microsculpture consisting of widely transverse to isodiametric meshes. Median lobe of aedeagus with apical orifice on dorsal side; apex rounded in dorsal view.</paragraph>
<paragraph id="9B4836BC5216FFEE29C7F86EFB364C97" blockId="9.[151,1437,1714,2029]" lastBlockId="10.[151,1437,151,1293]" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460">
This species may be remotely related to the consubgeners by having a peculiar morphology of the median lobe of the aedeagus.
<taxonomicName id="5CF74D3F5216FFED288CF812FDFB4BAE" authorityName="Park, Kwon &amp; Lafer" authorityYear="1996" box="[396,533,2002,2028]" pageId="9" pageNumber="463" rank="subGenus" subGenus="Micronialoe">
<emphasis id="A983EAAE5216FFED288CF812FDFB4BAE" box="[396,533,2002,2028]" italics="true" pageId="9" pageNumber="460">Micronialoe</emphasis>
</taxonomicName>
is included in the clade of
<taxonomicName id="5CF74D3F5216FFED2A46F812FC3F4BAE" authorityName="Bonelli" authorityYear="1810" box="[838,977,2002,2028]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="genus">
<emphasis id="A983EAAE5216FFED2A46F812FC3F4BAE" box="[838,977,2002,2028]" italics="true" pageId="9" pageNumber="460">Pterostichus</emphasis>
</taxonomicName>
which possesses the apical orifice of the median lobe bent to the left (
<bibRefCitation id="FF664B4D5215FFEE28D6FF57FCFE4CF3" author="Sasakawa, K. &amp; Kubota, K." box="[470,784,151,177]" pageId="10" pageNumber="461" pagination="100 - 109" refId="ref7918" refString="Sasakawa, K. &amp; Kubota, K. (2007) Phylogeny and genital evolution of carabid beetles in the genus Pterostichus and its allied genera (Coleoptera: Carabidae) inferred from two nuclear gene sequences. Annals of the Entomological Society of America, 100 (2), 100 - 109. https: // doi. org / 10.1603 / 0013 - 8746 (2007) 100 [100: PAGEOC] 2.0. CO; 2" type="journal article" year="2007">Sasakawa and Kubota 2007</bibRefCitation>
;
<bibRefCitation id="FF664B4D5215FFEE2A1BFF57FC144CF3" author="Schmidt, J. &amp; Opgenoorth, L. &amp; Holl, S. &amp; Bastrop, R." box="[795,1018,151,177]" pageId="10" pageNumber="461" pagination="45482" refId="ref8000" refString="Schmidt, J., Opgenoorth, L., Holl, S. &amp; Bastrop, R. (2012) Into the Himalayan Exile: The Phylogeography of the Ground Beetle Ethira clade Supports the Tibetan Origin of Forest-Dwelling Himalayan Species Groups. PLoS ONE, 7 (9), e 45482. https: // doi. org / 10.1371 / journal. pone. 0045482" type="journal article" year="2012">
Schmidt
<emphasis id="A983EAAE5215FFEE2A7FFF57FC594CF3" box="[895,951,151,177]" italics="true" pageId="10" pageNumber="461">et al.</emphasis>
2012
</bibRefCitation>
).
<taxonomicName id="5CF74D3F5215FFEE2D0CFF57FB584CF3" box="[1036,1206,151,177]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="chogyesanus">
<emphasis id="A983EAAE5215FFEE2D0CFF57FB584CF3" box="[1036,1206,151,177]" italics="true" pageId="10" pageNumber="461">P. chogyesanus</emphasis>
</taxonomicName>
has an apical orifice located on the dorsal side of the median lobe which may be a reversal from the apomorphic state.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375215FFEE29C7FF1EFDCB481B" pageId="10" pageNumber="461" type="description">
<paragraph id="9B4836BC5215FFEE29C7FF1EFC6B4CBB" blockId="10.[151,1437,151,1293]" box="[199,901,222,249]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FF1EFE9B4CBA" bold="true" box="[199,373,222,248]" pageId="10" pageNumber="461">Redescription.</emphasis>
Body length:
<quantity id="5C0F9B595215FFEE2B14FF1EFD734CBB" box="[532,669,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" metricValueMax="7.5" metricValueMin="5.9" pageId="10" pageNumber="461" unit="mm" value="6.7" valueMax="7.5" valueMin="5.9">5.97.5 mm</quantity>
, width:
<quantity id="5C0F9B595215FFEE2BF7FF1FFC6F4CBB" box="[759,897,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="10" pageNumber="461" unit="mm" value="2.35" valueMax="2.6" valueMin="2.1">2.12.6 mm</quantity>
.
</paragraph>
<paragraph id="9B4836BC5215FFEE29C7FEC3FAD44D03" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
Coloration brownish-black or black and shiny; antennae, legs, and lateral margin of both pronotum and elytra dark brown; palpi yellowish brown, with apices distinctly paler; ventral side brownish black (
<figureCitation id="03CC2A395215FFEE2DAEFEE7FAC24D03" box="[1198,1324,295,321]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Figs 8A, B</figureCitation>
).
</paragraph>
<paragraph id="9B4836BC5215FFEE29C7FE8BFB9D4DCB" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FE8BFEEA4D27" box="[199,260,331,357]" italics="true" pageId="10" pageNumber="461">Head</emphasis>
(
<figureCitation id="03CC2A395215FFEE2814FE8BFE804D27" box="[276,366,331,357]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Fig. 8C</figureCitation>
). Eye moderately convex; tempus oblique, shorter than in previous species; frontal impression moderately depressed, tilted outward, and clearly ends before anterior supraorbital pore.
</paragraph>
<paragraph id="9B4836BC5215FFEE29C7FE53FE4F4E8F" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FE53FED84DEE" box="[199,310,403,428]" italics="true" pageId="10" pageNumber="461">Pronotum</emphasis>
(
<figureCitation id="03CC2A395215FFEE284CFE53FE494DEF" box="[332,423,403,429]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Fig. 8C</figureCitation>
) widest before 2/5 from apex; apical angle narrowly rounded, moderately protruded anteriorly; lateral margin with weak or clear sinuation before basal angle; anterolateral seta before widest point, 1/4 to 3/10 from apex; basal angle obtuse; anterior transverse impression faint; posterior transverse impression more or less distinct; with one pair of clear basal foveae; inner foveae deep and distinct, linear, almost parallel or slightly tilted outward anteriorly, with distinct punctures; outer foveae rudimentary or absent; area between inner foveae weakly rugose longitudinally; area between inner fovea and outer fovea almost smooth.
<collectionCode id="FDE6AE795215FFEE2DA0FD87FB244E23" box="[1184,1226,583,609]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="461">PW</collectionCode>
/HW = 1.501.57, mean 1.52 (
<quantity id="5C0F9B595215FFEE281AFDAAFE874EC7" box="[282,361,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="4.0132" pageId="10" pageNumber="461" unit="in" value="1.58">1.58 in</quantity>
<typeStatus id="444C881E5215FFEE286FFDABFE384EC7" box="[367,470,619,645]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
);
<collectionCode id="FDE6AE795215FFEE28E8FDABFDFF4EC7" box="[488,529,619,645]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="461">PW</collectionCode>
/
<collectionCode id="FDE6AE795215FFEE2B1AFDABFDD44EC7" box="[538,570,619,645]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="461" type="Museum">PL</collectionCode>
= 1.111.16, mean 1.14 (
<quantity id="5C0F9B595215FFEE2A56FDABFC4B4EC7" box="[854,933,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.8701999999999996" pageId="10" pageNumber="461" unit="in" value="1.13">1.13 in</quantity>
<typeStatus id="444C881E5215FFEE2AABFDABFBFC4EC7" box="[939,1042,619,645]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
);
<collectionCode id="FDE6AE795215FFEE2D24FDABFBA04EC7" box="[1060,1102,619,645]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="461">PW</collectionCode>
/PbW = 1.241.25, mean 1.25 (
<quantity id="5C0F9B595215FFEE299FFD4FFF014EEB" box="[159,239,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="3.1496" pageId="10" pageNumber="461" unit="in" value="1.24">1.24 in</quantity>
<typeStatus id="444C881E5215FFEE29F7FD4FFEB04EEB" box="[247,350,655,681]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
);
<collectionCode id="FDE6AE795215FFEE2872FD4FFE724EEB" box="[370,412,655,681]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="461">PW</collectionCode>
/PaW = 1.451.47, mean 1.46 (
<quantity id="5C0F9B595215FFEE2BFEFD4FFCA04EEB" box="[766,846,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="3.5814" pageId="10" pageNumber="461" unit="in" value="1.41">1.41 in</quantity>
<typeStatus id="444C881E5215FFEE2A55FD4FFC524EEB" box="[853,956,655,681]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
); PbW/PaW = 1.161.18, mean 1.17 (
<quantity id="5C0F9B595215FFEE2C6AFD4FFF434E8F" metricMagnitude="-2" metricUnit="m" metricValue="2.8956" pageId="10" pageNumber="461" unit="in" value="1.14">1.14 in</quantity>
<typeStatus id="444C881E5215FFEE29B4FD73FEF64E8F" box="[180,280,691,717]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
) in
<specimenCount id="8DF1FD355215FFEE2843FD73FE734E8F" box="[323,413,691,717]" count="4" pageId="10" pageNumber="461" type="male">4 males</specimenCount>
.
</paragraph>
<paragraph id="9B4836BC5215FFEE29C7FD17FC614F1F" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FD17FEE34EB3" box="[199,269,727,753]" italics="true" pageId="10" pageNumber="461">Elytra</emphasis>
widest near 2/5 from base; striae deep, punctate from base to about basal two-thirds; marginal umbilicate series composed of three groups, 5 pores in anterior group, 1 pore in medial group, and 8 pores (rarely 9 pores) in posterior group. EW/
<collectionCode id="FDE6AE795215FFEE2887FCDFFE5F4F7B" box="[391,433,799,825]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="461">PW</collectionCode>
=1.151.23, mean 1.18 (
<quantity id="5C0F9B595215FFEE2BCEFCDFFCF14F7B" box="[718,799,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="3.0987999999999998" pageId="10" pageNumber="461" unit="in" value="1.22">1.22 in</quantity>
<typeStatus id="444C881E5215FFEE2A27FCDFFC604F7B" box="[807,910,799,825]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
); EL/
<collectionCode id="FDE6AE795215FFEE2ACBFCDFFC054F7B" box="[971,1003,799,825]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="461" type="Museum">PL</collectionCode>
= 2.022.09 (
<quantity id="5C0F9B595215FFEE2D8CFCDFFB334F7B" box="[1164,1245,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="5.5118" pageId="10" pageNumber="461" unit="in" value="2.17">2.17 in</quantity>
<typeStatus id="444C881E5215FFEE2DE5FCDFFAA54F7B" box="[1253,1355,799,825]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
), mean 2.06; EL/EW = 1.471.59, mean 1.53 (
<quantity id="5C0F9B595215FFEE2B4CFC82FD754F1F" box="[588,667,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="4.0132" pageId="10" pageNumber="461" unit="in" value="1.58">1.58 in</quantity>
<typeStatus id="444C881E5215FFEE2BA2FC83FCE84F1F" box="[674,774,835,861]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
) in
<specimenCount id="8DF1FD355215FFEE2A31FC83FC654F1F" box="[817,907,835,861]" count="4" pageId="10" pageNumber="461" type="male">4 males</specimenCount>
.
</paragraph>
<paragraph id="9B4836BC5215FFEE29C7FCA7FCD84FE7" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FCA7FEA34FC3" box="[199,333,871,897]" italics="true" pageId="10" pageNumber="461">Ventral side</emphasis>
(
<figureCitation id="03CC2A395215FFEE285EFCA7FE0E4FC3" box="[350,480,871,897]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Figs 8B, D</figureCitation>
). Posteromedian area of male sternite VII normal, with microsculpture consisting of widely transverse or isodiametric meshes in male (
<figureCitation id="03CC2A395215FFEE2BCFFC4BFCC74FE7" box="[719,809,907,933]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Fig. 8D</figureCitation>
).
</paragraph>
<paragraph id="9B4836BC5215FFEE29C7FC6FFDCB481B" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FC6FFE804F8B" box="[199,366,943,969]" italics="true" pageId="10" pageNumber="461">Male genitalia</emphasis>
(
<figureCitation id="03CC2A395215FFEE287FFC6FFE184F8B" box="[383,502,943,969]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Figs 8EJ</figureCitation>
). Median lobe of aedeagus (
<figureCitation id="03CC2A395215FFEE2A3CFC6FFC564F8B" box="[828,952,943,969]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Figs 8EH</figureCitation>
) slender, distinctly curved in lateral view, almost linear in dorsal view; apical orifice on dorsal side, not bent left; terminal lamella wide and long; apex rounded in dorsal view; ventral carina absent; sclerotized area in internal sac with only C-shaped sclerite on median area (CS); left paramere (
<figureCitation id="03CC2A395215FFEE28B2FBDBFE134877" box="[434,509,1051,1077]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Fig. 8I</figureCitation>
) wide; right paramere (
<figureCitation id="03CC2A395215FFEE2A03FBDBFCBE4877" box="[771,848,1051,1077]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EH; 0.2 mm IJ. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Fig. 8J</figureCitation>
) slender, weakly to distinctly curved, apical part less developed than in previous species.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375215FFEE29C7FBA3FBD548E3" pageId="10" pageNumber="461" type="materials_examined">
<paragraph id="9B4836BC5215FFEE29C7FBA3FBD548E3" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
<materialsCitation id="2B9F3CE15215FFEE29C7FBA3FBD548E3" ID-GBIF-Occurrence="4898139305" country="South Korea" location="Mt. Jogyesan" municipality="Suncheon-si" pageId="10" pageNumber="461" specimenCount="1" typeStatus="holotype">
<emphasis id="A983EAAE5215FFEE29C7FBA3FE8B483E" bold="true" box="[199,357,1122,1149]" pageId="10" pageNumber="461">
<typeStatus id="444C881E5215FFEE29C7FBA3FEEC483F" box="[199,258,1123,1149]" pageId="10" pageNumber="461">Type</typeStatus>
locality.
</emphasis>
<location id="9E2860675215FFEE286AFBA3FDEF483F" LSID="urn:lsid:plazi:treatment:135E87AA5217FFEE2997F92AFF08494F:9E2860675215FFEE286AFBA3FDEF483F" box="[362,513,1123,1149]" country="South Korea" municipality="Suncheon-si" name="Mt. Jogyesan" pageId="10" pageNumber="461">Mt. Jogyesan</location>
,
<collectingMunicipality id="7B2CACC65215FFEE2B0CFBA3FD77483F" box="[524,665,1123,1149]" pageId="10" pageNumber="461">Suncheon-si</collectingMunicipality>
,
<collectingCountry id="E3E0762C5215FFEE2BA3FBA3FD05483F" box="[675,747,1123,1149]" name="South Korea" pageId="10" pageNumber="461">Korea</collectingCountry>
. (The Korean name ‘송광’ in the original description suggests that Songgwangsa Temple in the western part of the mountain is the exact locality.)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375215FFEE29C7FB6BFBA44887" box="[199,1098,1195,1221]" pageId="10" pageNumber="461" type="biology_ecology">
<paragraph id="9B4836BC5215FFEE29C7FB6BFBA44887" blockId="10.[151,1437,151,1293]" box="[199,1098,1195,1221]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FB6BFEC24887" bold="true" box="[199,300,1195,1221]" pageId="10" pageNumber="461">Ecology.</emphasis>
The adult specimens were observed beneath rocks in the humid valley.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375215FFEE29C7FB0FFF08494F" pageId="10" pageNumber="461" type="discussion">
<paragraph id="9B4836BC5215FFEE29C7FB0FFF08494F" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="461">
<emphasis id="A983EAAE5215FFEE29C7FB0FFEDA48AB" bold="true" box="[199,308,1231,1257]" pageId="10" pageNumber="461">Remarks</emphasis>
. This species was only known from its
<typeStatus id="444C881E5215FFEE2A0EFB0FFCD048AB" box="[782,830,1231,1257]" pageId="10" pageNumber="461">type</typeStatus>
locality, and Mt. Beagunsan is a new distributional record.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,273 @@
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<mods:title id="FC39E4B066333E24D0472149EAE28756">Review of the ground beetle subgenus Micronialoe from Korea, with a description of a new species (Coleoptera: Carabidae: Pterostichus)</mods:title>
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<mods:affiliation id="33587A96AFA8175243DC4D0436B8E83F">Department of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea &amp; Institute of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<treatment id="135E87AA5218FFEC2997FDA6FCC04CBB" ID-DOI="http://doi.org/10.5281/zenodo.12190872" ID-GBIF-Taxon="233318008" ID-Zenodo-Dep="12190872" LSID="urn:lsid:plazi:treatment:135E87AA5218FFEC2997FDA6FCC04CBB" httpUri="http://treatment.plazi.org/id/135E87AA5218FFEC2997FDA6FCC04CBB" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<subSubSection id="D3ED65375218FFE32997FDA6FDF24EE4" pageId="7" pageNumber="458" type="nomenclature">
<paragraph id="9B4836BC5218FFE32997FDA6FC274EC2" blockId="7.[151,969,614,713]" box="[151,969,614,641]" pageId="7" pageNumber="458">
<heading id="C00081D05218FFE32997FDA6FC274EC2" bold="true" box="[151,969,614,641]" fontSize="11" level="1" pageId="7" pageNumber="458" reason="1">
<taxonomicName id="5CF74D3F5218FFE32997FDA6FC274EC2" ID-CoL="4Q3X4" authority="Park, Kwon &amp; Lafer, 1996" authorityName="Park, Kwon &amp; Lafer" authorityYear="1996" box="[151,969,614,641]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="bifoveolatus" subGenus="Micronialoe">
<emphasis id="A983EAAE5218FFE32997FDA6FC274EC2" bold="true" box="[151,969,614,641]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE32997FDA6FEC34EC3" bold="true" box="[151,301,614,641]" italics="true" pageId="7" pageNumber="458">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5218FFE3283EFDA6FE3C4EC3" bold="true" box="[318,466,614,641]" italics="true" pageId="7" pageNumber="458">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5218FFE328E4FDA6FD994EC3" bold="true" box="[484,631,614,641]" italics="true" pageId="7" pageNumber="458">bifoveolatus</emphasis>
<bibRefCitation id="FF664B4D5218FFE32B7EFDA7FC274EC2" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[638,969,614,641]" pageId="7" pageNumber="458" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9B4836BC5218FFE32997FD4BFDF24EE4" blockId="7.[151,969,614,713]" box="[151,540,651,678]" pageId="7" pageNumber="458">Korean name: Ī마길쭉먼지벌레</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE32997FD6FFEE64E8B" box="[151,264,687,713]" pageId="7" pageNumber="458" type="description">
<paragraph id="9B4836BC5218FFE32997FD6FFEE64E8B" blockId="7.[151,969,614,713]" box="[151,264,687,713]" pageId="7" pageNumber="458">
(
<figureCitation id="03CC2A395218FFE3299FFD6FFEEE4E8B" box="[159,256,687,713]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figure 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE32997FD33FB7C4F3A" pageId="7" pageNumber="458" type="reference_group">
<paragraph id="9B4836BC5218FFE32997FD33FC834F49" blockId="7.[151,877,755,779]" box="[151,877,755,779]" pageId="7" pageNumber="458">
<treatmentCitationGroup id="BBE711925218FFE32997FD33FC834F49" box="[151,877,755,779]" pageId="7" pageNumber="458">
<treatmentCitation id="1A5610AD5218FFE32997FD33FC874F49" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[151,873,755,779]" page="74" pageId="7" pageNumber="458" year="1996">
<taxonomicName id="5CF74D3F5218FFE32997FD33FC874F49" ID-CoL="4Q3X4" authority="Park, Kwon &amp; Lafer, 1996: 74" authorityName="Park, Kwon &amp; Lafer" authorityPageNumber="74" authorityYear="1996" box="[151,873,755,779]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="bifoveolatus" subGenus="Micronialoe">
<emphasis id="A983EAAE5218FFE32997FD33FEFB4F49" box="[151,277,755,779]" italics="true" pageId="7" pageNumber="458">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5218FFE32824FD33FE4E4F49" box="[292,416,755,779]" italics="true" pageId="7" pageNumber="458">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5218FFE328AFFD33FDC54F49" box="[431,555,755,779]" italics="true" pageId="7" pageNumber="458">bifoveolatus</emphasis>
<bibRefCitation id="FF664B4D5218FFE32B31FD33FC874F49" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[561,873,755,779]" pageId="7" pageNumber="458" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996: 74</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="9B4836BC5218FFE32997FCF9FB7C4F3A" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE32997FCF9FE6F4F11" bold="true" box="[151,385,825,851]" pageId="7" pageNumber="458">Specimen examined</emphasis>
.
<materialsCitation id="2B9F3CE15218FFE3288CFCFAFB654F3A" ID-GBIF-Occurrence="4898139313" collectingDate="2022-08-05" collectorName="Dooyoung Kim" country="South Korea" county="Sancheong-gun" elevation="530" latitude="35.37964" location="Honggye-ri" longLatPrecision="1" longitude="127.853806" municipality="Samjang-myeon" pageId="7" pageNumber="458" specimenCount="1" specimenCount-male="1" stateProvince="Gyeongnam">
<collectingCountry id="E3E0762C5218FFE3288CFCFAFE3E4F16" box="[396,464,826,852]" name="South Korea" pageId="7" pageNumber="458">Korea</collectingCountry>
<specimenCount id="8DF1FD355218FFE328E2FCF9FDE94F11" box="[482,519,825,851]" count="1" pageId="7" pageNumber="458" type="male">1♂</specimenCount>
;
<collectingRegion id="5933F85E5218FFE32B11FCF9FD1B4F11" box="[529,757,825,851]" country="South Korea" name="Gyeongnam" pageId="7" pageNumber="458">Gyeongsangnam-do</collectingRegion>
,
<collectingCounty id="72294E305218FFE32BFEFCF9FC414F11" box="[766,943,825,851]" pageId="7" pageNumber="458">Sancheong-gun</collectingCounty>
,
<collectingMunicipality id="7B2CACC65218FFE32AB7FCF9FB9F4F11" box="[951,1137,825,851]" pageId="7" pageNumber="458">Samjang-myeon</collectingMunicipality>
,
<location id="9E2860675218FFE32D79FCF9FB134F11" LSID="urn:lsid:plazi:treatment:135E87AA5218FFEC2997FDA6FCC04CBB:9E2860675218FFE32D79FCF9FB134F11" box="[1145,1277,825,851]" country="South Korea" county="Sancheong-gun" latitude="35.37964" longLatPrecision="1" longitude="127.853806" municipality="Samjang-myeon" name="Honggye-ri" pageId="7" pageNumber="458" stateProvince="Gyeongnam">Honggye-ri</location>
,
<location id="9E2860675218FFE32C05FCF9FA724F11" LSID="urn:lsid:plazi:treatment:135E87AA5218FFEC2997FDA6FCC04CBB:9E2860675218FFE32C05FCF9FA724F11" box="[1285,1436,825,851]" country="South Korea" county="Sancheong-gun" latitude="35.37964" longLatPrecision="1" longitude="127.853806" municipality="Samjang-myeon" name="Bammeorijae" pageId="7" pageNumber="458" stateProvince="Gyeongnam">Bammeorijae</location>
pass;
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<geoCoordinate id="FEC3507B5218FFE32876FC9DFDF34F3A" box="[374,541,861,888]" degrees="127" direction="east" minutes="51" orientation="longitude" pageId="7" pageNumber="458" precision="1" seconds="13.7" value="127.853806">127°51'13.7&quot;E</geoCoordinate>
; alt. ca
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;
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<collectingDate id="FF0DE9945218FFE32BC8FC9DFCB84F35" box="[712,854,861,888]" pageId="7" pageNumber="458" value="2022-08-05">5 Aug. 2022</collectingDate>
</date>
;
<collectorName id="3602536A5218FFE32A60FC9EFBFE4F35" box="[864,1040,861,888]" pageId="7" pageNumber="458">Dooyoung Kim</collectorName>
leg.; KNU
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE329C7FC41FD1D4941" pageId="7" pageNumber="458" type="diagnosis">
<paragraph id="9B4836BC5218FFE329C7FC41FABB4FA1" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FC41FEAC4FD9" bold="true" box="[199,322,897,923]" pageId="7" pageNumber="458">Diagnosis.</emphasis>
This species is easily distinguished from known species of the subgenus by: i) posteromedian area of male sternite VII not rugose and asymmetrically depressed, ii) median lobe of aedeagus with rounded apex in dorsal view, and iii) internal sac of aedeagus with two sclerotized areas. Other significant characters are as follows.
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FC2EFD9748DA" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">Pronotum slightly sinuate on lateral margin before basal angle; with two pairs of basal foveae. Posteromedian area of male sternite VII with microsculpture consisting of irregular transverse meshes. Median lobe of aedeagus stout, left lateral margin below apical orifice distinctly convex; ventral carina well developed, widely rounded; sclerotized areas in internal sac consist of followings: i) C-shaped sclerite (CS) on median area and ii) wide triangular shaped ventral sclerite (VS) on apical area.</paragraph>
<paragraph id="9B4836BC5218FFE329C7FB61FD1D4941" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
This species may be closely allied to
<taxonomicName id="5CF74D3F5218FFE32B72FB62FC5E48FE" authority="Morita, 2005" authorityName="Morita" authorityYear="2005" box="[626,944,1185,1212]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="461" phylum="Arthropoda" rank="species" species="kaniei" subGenus="Micronialoe">
<emphasis id="A983EAAE5218FFE32B72FB62FD6D48F9" box="[626,643,1186,1211]" italics="true" pageId="7" pageNumber="458">P</emphasis>
. (
<emphasis id="A983EAAE5218FFE32B9CFB62FD5D48F9" box="[668,691,1186,1211]" italics="true" pageId="7" pageNumber="458">M</emphasis>
.)
<emphasis id="A983EAAE5218FFE32BCCFB61FCFE48F9" box="[716,784,1185,1211]" italics="true" pageId="7" pageNumber="458">kaniei</emphasis>
<bibRefCitation id="FF664B4D5218FFE32A19FB61FC5E48FE" author="Morita, S." box="[793,944,1185,1212]" pageId="7" pageNumber="458" pagination="9 - 12" refId="ref7740" refString="Morita, S. (2005) A New Pterostichus (Coleoptera: Carabidae) from the Islands of Tsushima, West Japan. Entomological Review of Japan, 60 (1), 9 - 12." type="journal article" year="2005">Morita, 2005</bibRefCitation>
</taxonomicName>
, based on their common genital characters: i) median lobe of aedeagus stout, ii) apex moderately rounded in dorsal view, iii) left lateral margin below apical orifice convex, and iv) ventral carina widely rounded.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFEC29C7FACDFB014C97" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458" type="description">
<paragraph id="9B4836BC5218FFE329C7FACDFC6B496A" blockId="7.[151,1437,825,2040]" box="[199,901,1293,1320]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FACDFE9B4965" bold="true" box="[199,373,1293,1319]" pageId="7" pageNumber="458">Redescription.</emphasis>
Body length:
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, width:
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.
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FAF1FF5449D1" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
Coloration reddish black and shiny, elytra slightly iridescent; antennae, legs, and lateral margin of both pronotum and elytra reddish brown; palpi yellowish brown, with apices distinctly paler; ventral side reddish brown (
<figureCitation id="03CC2A395218FFE32C3AFA95FF4249D1" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7A, B</figureCitation>
).
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FA5DFC43499E" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FA5DFEEA49F5" box="[199,260,1437,1463]" italics="true" pageId="7" pageNumber="458">Head</emphasis>
(
<figureCitation id="03CC2A395218FFE32818FA5DFE9B49F5" box="[280,373,1437,1463]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Fig. 7C</figureCitation>
). Eye weakly convex; tempus well developed, moderately convex, as long as eye; frontal impression deep, moderately curved toward anterior supraorbital pore.
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FA26FCB54AF1" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FA26FED849BD" box="[199,310,1510,1535]" italics="true" pageId="7" pageNumber="458">Pronotum</emphasis>
(
<figureCitation id="03CC2A395218FFE32848FA25FE6149BD" box="[328,399,1509,1535]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Fig. 7</figureCitation>
) widest near 2/5 from apex; apical angle narrowly rounded, slightly protruded anteriorly; lateral margin slightly sinuate before basal angle; basal angle obtuse; with two pairs of basal foveae; inner foveae deep and distinct, linear, almost parallel, with distinct punctures; outer foveae almost linear, slightly tilted outward; basal surface almost smooth medially; area between inner fovea and outer fovea with or without distinct punctures.
<materialsCitation id="2B9F3CE15218FFE32997F9B5FDF64ACD" ID-GBIF-Occurrence="4898139302" box="[151,536,1653,1680]" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32997F9B5FF2F4ACD" box="[151,193,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/HW = 1.61 (
<quantity id="5C0F9B595218FFE32854F9B5FE4D4ACD" box="[340,419,1653,1680]" metricMagnitude="-2" metricUnit="m" metricValue="4.0386" pageId="7" pageNumber="458" unit="in" value="1.59">1.59 in</quantity>
<typeStatus id="444C881E5218FFE328A9F9B5FDE14ACD" box="[425,527,1653,1679]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
,
<materialsCitation id="2B9F3CE15218FFE32B22F9B5FC7A4ACD" ID-GBIF-Occurrence="4898139312" box="[546,916,1653,1680]" collectionCode="PW, PL" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32B22F9B5FDA54ACD" box="[546,587,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/
<collectionCode id="FDE6AE795218FFE32B54F9B5FD9A4ACD" box="[596,628,1653,1679]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="458" type="Museum">PL</collectionCode>
= 1.11 (
<quantity id="5C0F9B595218FFE32BD0F9B5FCF14ACD" box="[720,799,1653,1679]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="7" pageNumber="458" unit="in" value="1.2">1.20 in</quantity>
<typeStatus id="444C881E5218FFE32A25F9B5FC624ACD" box="[805,908,1653,1679]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2B9F3CE15218FFE32A9FF9B5FAC44ACD" ID-GBIF-Occurrence="4898139307" box="[927,1322,1653,1680]" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32A9FF9B5FC274ACD" box="[927,969,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/PbW = 1.19 (
<quantity id="5C0F9B595218FFE32D65F9B5FB5A4ACD" box="[1125,1204,1653,1679]" metricMagnitude="-2" metricUnit="m" metricValue="2.9463999999999997" pageId="7" pageNumber="458" unit="in" value="1.16">1.16 in</quantity>
<typeStatus id="444C881E5218FFE32DBBF9B5FACC4ACD" box="[1211,1314,1653,1679]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2B9F3CE15218FFE32C34F9B5FE5D4AF1" ID-GBIF-Occurrence="4898139315" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32C34F9B5FAB04ACD" box="[1332,1374,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/PaW = 1.46 (
<quantity id="5C0F9B595218FFE329EEF959FED34AF1" box="[238,317,1689,1715]" metricMagnitude="-2" metricUnit="m" metricValue="3.5814" pageId="7" pageNumber="458" unit="in" value="1.41">1.41 in</quantity>
<typeStatus id="444C881E5218FFE32844F959FE454AF1" box="[324,427,1689,1715]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
; PbW/PaW = 1.23 (
<quantity id="5C0F9B595218FFE32B92F959FD0F4AF1" box="[658,737,1689,1715]" metricMagnitude="-2" metricUnit="m" metricValue="3.0734" pageId="7" pageNumber="458" unit="in" value="1.21">1.21 in</quantity>
<typeStatus id="444C881E5218FFE32BE8F959FCA14AF1" box="[744,847,1689,1715]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="9B4836BC5218FFE329C7F97DFA8E4B5D" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7F97DFEE34A95" box="[199,269,1725,1751]" italics="true" pageId="7" pageNumber="458">Elytra</emphasis>
<materialsCitation id="2B9F3CE15218FFE32816F97DFD8C4B5D" ID-GBIF-Occurrence="4898139306" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
widest before 2/5 from base; striae deep, punctate from base to basal half; marginal umbilicate series composed of three groups, humeral group with 5 pores, medial group with 1 pore, and apical group with 7 or 8 pores. EW/
<collectionCode id="FDE6AE795218FFE32815F8C5FED14B5D" box="[277,319,1797,1823]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
= 1.19 (
<quantity id="5C0F9B595218FFE3289CF8C5FE024B5D" box="[412,492,1797,1823]" metricMagnitude="-2" metricUnit="m" metricValue="3.0225999999999997" pageId="7" pageNumber="458" unit="in" value="1.19">1.19 in</quantity>
<typeStatus id="444C881E5218FFE328F3F8C5FDB44B5D" box="[499,602,1797,1823]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2B9F3CE15218FFE32B6DF8C5FC374B5D" ID-GBIF-Occurrence="4898139316" box="[621,985,1797,1824]" collectionCode="PL" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
EL/
<collectionCode id="FDE6AE795218FFE32B96F8C5FD584B5D" box="[662,694,1797,1823]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="458" type="Museum">PL</collectionCode>
= 2.02 (
<quantity id="5C0F9B595218FFE32A14F8C5FC8D4B5D" box="[788,867,1797,1823]" metricMagnitude="-2" metricUnit="m" metricValue="5.740399999999999" pageId="7" pageNumber="458" unit="in" value="2.26">2.26 in</quantity>
<typeStatus id="444C881E5218FFE32A6AF8C5FC3F4B5D" box="[874,977,1797,1823]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
; EL/EW = 1.55 (
<quantity id="5C0F9B595218FFE32D97F8C5FB084B5D" box="[1175,1254,1797,1824]" metricMagnitude="-2" metricUnit="m" metricValue="4.0132" pageId="7" pageNumber="458" unit="in" value="1.58">1.58 in</quantity>
<typeStatus id="444C881E5218FFE32DEDF8C5FABA4B5D" box="[1261,1364,1797,1823]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="9B4836BC5218FFE329C7F8E9FC464B25" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7F8E9FEA04B01" box="[199,334,1833,1859]" italics="true" pageId="7" pageNumber="458">Ventral side</emphasis>
(
<figureCitation id="03CC2A395218FFE3285EF8E9FE014B01" box="[350,495,1833,1860]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7B, 7D</figureCitation>
). Posteromedian area of male sternite VII (
<figureCitation id="03CC2A395218FFE32ADBF8E9FBDB4B01" box="[987,1077,1833,1859]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Fig. 7D</figureCitation>
) asymmetrically depressed, not rugose, with microsculpture consisting of irregular transverse meshes.
</paragraph>
<paragraph id="9B4836BC5218FFEC29C7F8B1FB014C97" blockId="7.[151,1437,825,2040]" lastBlockId="8.[151,1437,151,249]" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7F8B1FE844BC9" box="[199,362,1905,1931]" italics="true" pageId="7" pageNumber="458">Male genitalia</emphasis>
(
<figureCitation id="03CC2A395218FFE32878F8B1FE1B4BC9" box="[376,501,1905,1931]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7EK</figureCitation>
). Median lobe of aedeagus (
<figureCitation id="03CC2A395218FFE32A29F8B1FC764BC9" box="[809,920,1905,1931]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7EI</figureCitation>
) stout, distinctly curved in lateral view, almost linear in dorsal view; left lateral margin below apical orifice distinctly convex; apical orifice bent to left; terminal lamella very short; apex moderately rounded in dorsal view; ventral carina well developed, widely rounded, clearly visible in right dorsolateral view; internal sac with two sclerotized areas: i) C-shaped sclerite (CS) on median area and ii) wide triangular shaped ventral sclerite (VS) on apical area; left paramere (
<figureCitation id="03CC2A395217FFEC2D1CFF57FB844CF3" box="[1052,1130,151,177]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="8" pageNumber="459">Fig. 7J</figureCitation>
) wide; right paramere (
<figureCitation id="03CC2A395217FFEC2C6FFF57FF554C97" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="8" pageNumber="459">Fig. 7K</figureCitation>
) slender, distinctly curved, ventral margin forming obtuse angle, dorsal margin widely rounded.
</paragraph>
</subSubSection>
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<paragraph id="9B4836BC5217FFEC29C7FF1FFCC04CBB" blockId="8.[151,1437,151,249]" box="[199,814,222,249]" pageId="8" pageNumber="459">
<materialsCitation id="2B9F3CE15217FFEC29C7FF1FFCC44CBB" ID-GBIF-Occurrence="4898139304" box="[199,810,222,249]" country="South Korea" location="Mt. Jirisan" pageId="8" pageNumber="459" specimenCount="1" stateProvince="Gyeongnam" typeStatus="holotype">
<emphasis id="A983EAAE5217FFEC29C7FF1FFE884CBA" bold="true" box="[199,358,222,249]" pageId="8" pageNumber="459">
<typeStatus id="444C881E5217FFEC29C7FF1FFEEC4CBB" box="[199,258,223,249]" pageId="8" pageNumber="459">Type</typeStatus>
locality.
</emphasis>
<location id="9E2860675217FFEC286CFF1FFE094CBB" LSID="urn:lsid:plazi:treatment:135E87AA5218FFEC2997FDA6FCC04CBB:9E2860675217FFEC286CFF1FFE094CBB" box="[364,487,223,249]" country="South Korea" name="Mt. Jirisan" pageId="8" pageNumber="459" stateProvince="Gyeongnam">Mt. Jirisan</location>
,
<collectingRegion id="5933F85E5217FFEC28F2FF1FFD384CBB" box="[498,726,223,249]" country="South Korea" name="Gyeongnam" pageId="8" pageNumber="459">Gyeongsangnam-do</collectingRegion>
,
<collectingCountry id="E3E0762C5217FFEC2BE2FF1FFCC44CBB" box="[738,810,223,249]" name="South Korea" pageId="8" pageNumber="459">Korea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="AF33D120FD21B8D24A33A46CA33105D5">Review of the ground beetle subgenus Micronialoe from Korea, with a description of a new species (Coleoptera: Carabidae: Pterostichus)</mods:title>
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<mods:namePart id="F007AD529EFC8BBFFA7C2865838C0B8F">Kim, Dooyoung</mods:namePart>
<mods:affiliation id="081E0497F20706DC2BB8B9BEBDABACAE">Department of Biomedical Convergence Science and Technology, School of Industrial Technology Advances, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<mods:namePart id="7974061B98A5BF99A0C0AE30C799AAF1">Suh, Sang Jae</mods:namePart>
<mods:affiliation id="48BA535A6445292BA818B246D76954E8">Department of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea &amp; Institute of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<emphasis id="A983EAAE521CFFE72997F9BAFEC34AD7" bold="true" box="[151,301,1658,1685]" italics="true" pageId="3" pageNumber="454">Pterostichus</emphasis>
(
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)
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Kim &amp; Suh
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https://zoobank.org/
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(
<figureCitation id="03CC2A39521CFFE7299FF927FEC94B43" box="[159,295,1767,1793]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="2.[151,250,1973,1997]" captionStartId-1="4.[151,250,1399,1423]" captionStartId-2="5.[151,250,755,779]" captionStartId-3="5.[151,250,1877,1901]" captionStartId-4="6.[151,250,1951,1975]" captionTargetBox-0="[155,1436,1248,1941]" captionTargetBox-1="[151,1436,189,1365]" captionTargetBox-2="[151,1436,181,732]" captionTargetBox-3="[161,1426,850,1841]" captionTargetBox-4="[151,1436,1208,1927]" captionTargetId-0="figure-445@2.[151,1436,1240,1950]" captionTargetId-1="figure-35@4.[151,1436,181,1375]" captionTargetId-2="figure-18@5.[151,1436,181,732]" captionTargetId-3="figure-42@5.[151,1436,840,1854]" captionTargetId-4="figure-499@6.[151,1436,1208,1927]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 2. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) habitus, dorsal view; (B) ditto, ventral view; C head and pronotum, dorsal view. Scale bars: 1 mm." captionText-1="FIGURE 3. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) mouthpart, ventral view; (B) ditto, lateral view; (C) metepisternum; (D) sternite VII. Scale bars: 0.3 mm." captionText-2="FIGURE 4. Microsculpture of Pterostichus (Micronialoe) spinulosus sp. nov. (A) head; (B) pronotum; (C) elytron. Scale bars: 0.1 mm." captionText-3="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." captionText-4="FIGURE 6. Everted internal sac of Pterostichus (Micronialoe) spinulosus sp. nov. (A) left lateral view; (B) right lateral view. Scale bar: 0.3 mm. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi-0="http://doi.org/10.5281/zenodo.12190391" figureDoi-1="http://doi.org/10.5281/zenodo.12190393" figureDoi-2="http://doi.org/10.5281/zenodo.12190397" figureDoi-3="http://doi.org/10.5281/zenodo.12190399" figureDoi-4="http://doi.org/10.5281/zenodo.12190401" httpUri-0="https://zenodo.org/record/12190391/files/figure.png" httpUri-1="https://zenodo.org/record/12190393/files/figure.png" httpUri-2="https://zenodo.org/record/12190397/files/figure.png" httpUri-3="https://zenodo.org/record/12190399/files/figure.png" httpUri-4="https://zenodo.org/record/12190401/files/figure.png" pageId="3" pageNumber="454">Figures 26</figureCitation>
)
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<emphasis id="A983EAAE521CFFE72997F8EBFEA64B06" bold="true" box="[151,328,1834,1861]" pageId="3" pageNumber="454">Type materials</emphasis>
.
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:
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;
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leg.; KNU
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: •
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same data as for holotype
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<location id="9E286067521CFFE72D2CF8B3FB414BCF" LSID="urn:lsid:plazi:treatment:135E87AA521CFFE32997F9BAFD1F4E5B:9E286067521CFFE72D2CF8B3FB414BCF" box="[1068,1199,1907,1933]" country="South Korea" county="Mt. Biseulsan" latitude="35.725224" longLatPrecision="1" longitude="128.54254" municipality="Eastern" name="Jeongdae-ri" pageId="3" pageNumber="454">Jeongdae-ri</location>
,
<collectingMunicipality id="7B2CACC6521CFFE72DBEF8B3FAFE4BCF" box="[1214,1296,1907,1933]" pageId="3" pageNumber="454">Eastern</collectingMunicipality>
part of
<collectingCounty id="72294E30521CFFE72C75F8B3FEEB4BF3" pageId="3" pageNumber="454">Mt. Biseulsan</collectingCounty>
;
<geoCoordinate id="FEC3507B521CFFE7280FF856FE4B4BF3" box="[271,421,1942,1969]" degrees="35" direction="north" minutes="43" orientation="latitude" pageId="3" pageNumber="454" precision="1" seconds="30.8" value="35.725224">35°43'30.8&quot;N</geoCoordinate>
<geoCoordinate id="FEC3507B521CFFE728AAF857FDA04BF3" box="[426,590,1943,1969]" degrees="128" direction="east" minutes="32" orientation="longitude" pageId="3" pageNumber="454" precision="1" seconds="33.1" value="128.54254">128°32'33.1&quot;E</geoCoordinate>
; alt. ca
<quantity id="5C0F9B59521CFFE72B9EF856FD0B4BF3" box="[670,741,1942,1969]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="454" unit="m" value="580.0">
<elevation id="10DAD18F521CFFE72B9EF856FD0B4BF3" box="[670,741,1942,1969]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="454" unit="m" value="580.0">580 m</elevation>
</quantity>
;
<date id="EF49107C521CFFE72BEFF857FC6C4BF3" box="[751,898,1943,1969]" pageId="3" pageNumber="454" value="2021-06-26">
<collectingDate id="FF0DE994521CFFE72BEFF857FC6C4BF3" box="[751,898,1943,1969]" pageId="3" pageNumber="454" value="2021-06-26">26 June 2021</collectingDate>
</date>
;
<collectorName id="3602536A521CFFE72A8CF857FBD94BF3" box="[908,1079,1943,1969]" pageId="3" pageNumber="454">Dooyoung Kim</collectorName>
leg.; KNU
</materialsCitation>
<materialsCitation id="2B9F3CE1521CFFE72DC2F857FED14B97" ID-GBIF-Occurrence="4898139308" collectingDate="2021-10-29" collectorName="Dooyoung Kim" country="South Korea" county="Mt. Biseulsan" elevation="500" latitude="35.73003" location="Daegu-si" longLatPrecision="1" longitude="128.5518" municipality="Eastern" pageId="3" pageNumber="454" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
<specimenCount id="8DF1FD35521CFFE72DC2F857FB054BF3" box="[1218,1259,1943,1969]" count="4" pageId="3" pageNumber="454" type="male">4♂</specimenCount>
<specimenCount id="8DF1FD35521CFFE72DEBF857FAE84BF3" box="[1259,1286,1943,1969]" count="1" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
same locality as for holotype
</materialsCitation>
;
<date id="EF49107C521CFFE72849F87BFE304B97" box="[329,478,1979,2005]" pageId="3" pageNumber="454" value="2021-05-16">16 May 2021</date>
; Dooyoung Kim leg.; KNU •
<materialsCitation id="2B9F3CE1521CFFE72A22F87BFAE44BBB" ID-GBIF-Occurrence="4898139314" collectingDate="2021-07-01" collectorName="Dooyoung Kim" country="South Korea" county="Mt. Choejeongsan" elevation="800" latitude="35.754" location="Daegu-si" longLatPrecision="1" longitude="128.59988" municipality="Ju-ri" pageId="3" pageNumber="454" specimenCount="35" specimenCount-female="12" specimenCount-male="23" typeStatus="paratype">
<specimenCount id="8DF1FD35521CFFE72A22F87BFCB84B97" box="[802,854,1979,2005]" count="23" pageId="3" pageNumber="454" type="male">23♂</specimenCount>
<specimenCount id="8DF1FD35521CFFE72A56F87BFC6D4B97" box="[854,899,1979,2005]" count="12" pageId="3" pageNumber="454" type="female">12♀</specimenCount>
<location id="9E286067521CFFE72A89F87BFC014B97" LSID="urn:lsid:plazi:treatment:135E87AA521CFFE32997F9BAFD1F4E5B:9E286067521CFFE72A89F87BFC014B97" box="[905,1007,1979,2005]" country="South Korea" county="Mt. Choejeongsan" latitude="35.754" longLatPrecision="1" longitude="128.59988" municipality="Ju-ri" name="Daegu-si" pageId="3" pageNumber="454">Daegu-si</location>
,
<location id="9E286067521CFFE72AF9F87BFB784B97" LSID="urn:lsid:plazi:treatment:135E87AA521CFFE32997F9BAFD1F4E5B:9E286067521CFFE72AF9F87BFB784B97" box="[1017,1174,1979,2005]" country="South Korea" county="Mt. Choejeongsan" latitude="35.754" longLatPrecision="1" longitude="128.59988" municipality="Ju-ri" name="Dalseong-gun" pageId="3" pageNumber="454">Dalseong-gun</location>
,
<location id="9E286067521CFFE72DA0F87BFAB64B97" LSID="urn:lsid:plazi:treatment:135E87AA521CFFE32997F9BAFD1F4E5B:9E286067521CFFE72DA0F87BFAB64B97" box="[1184,1368,1979,2005]" country="South Korea" county="Mt. Choejeongsan" latitude="35.754" longLatPrecision="1" longitude="128.59988" municipality="Ju-ri" name="Gachang-myeon" pageId="3" pageNumber="454">Gachang-myeon</location>
,
<collectingMunicipality id="7B2CACC6521CFFE72C63F87BFA764B97" box="[1379,1432,1979,2005]" pageId="3" pageNumber="454">Ju-ri</collectingMunicipality>
,
<collectingCounty id="72294E30521CFFE72997F81FFE8A4BBB" box="[151,356,2015,2041]" pageId="3" pageNumber="454">Mt. Choejeongsan</collectingCounty>
;
<geoCoordinate id="FEC3507B521CFFE7286FF81EFDEB4BBB" box="[367,517,2014,2041]" degrees="35" direction="north" minutes="45" orientation="latitude" pageId="3" pageNumber="454" precision="1" seconds="14.4" value="35.754">35°45'14.4&quot;N</geoCoordinate>
<geoCoordinate id="FEC3507B521CFFE72B0CF81EFD5E4BBB" box="[524,688,2014,2041]" degrees="128" direction="east" minutes="35" orientation="longitude" pageId="3" pageNumber="454" precision="1" seconds="59.6" value="128.59988">128°35'59.6&quot;E</geoCoordinate>
; alt. ca
<quantity id="5C0F9B59521CFFE72A03F81FFCA54BBB" box="[771,843,2015,2041]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="454" unit="m" value="800.0">
<elevation id="10DAD18F521CFFE72A03F81FFCA54BBB" box="[771,843,2015,2041]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="454" unit="m" value="800.0">800 m</elevation>
</quantity>
;
<date id="EF49107C521CFFE72A57F81FFC344BBB" box="[855,986,2015,2041]" pageId="3" pageNumber="454" value="2021-07-01">
<collectingDate id="FF0DE994521CFFE72A57F81FFC344BBB" box="[855,986,2015,2041]" pageId="3" pageNumber="454" value="2021-07-01">1 July 2021</collectingDate>
</date>
;
<collectorName id="3602536A521CFFE72AE5F81FFB7F4BBB" box="[997,1169,2015,2041]" pageId="3" pageNumber="454">Dooyoung Kim</collectorName>
leg.; KNU
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="CF886634521BFFE02997FAB7FD4049F0" ID-DOI="http://doi.org/10.5281/zenodo.12190393" ID-Zenodo-Dep="12190393" httpUri="https://zenodo.org/record/12190393/files/figure.png" pageId="4" pageNumber="455" startId="4.[151,250,1399,1423]" targetBox="[151,1436,189,1365]" targetPageId="4" targetType="figure">
<paragraph id="9B4836BC521BFFE02997FAB7FD4049F0" blockId="4.[151,1436,1398,1459]" pageId="4" pageNumber="455">
<emphasis id="A983EAAE521BFFE02997FAB7FEF849CD" bold="true" box="[151,278,1399,1423]" pageId="4" pageNumber="455">FIGURE 3.</emphasis>
<taxonomicName id="5CF74D3F521BFFE0281FFAB7FD4849CD" authority="Kim &amp; Suh, 2024" authorityName="Kim &amp; Suh" authorityYear="2024" box="[287,678,1399,1423]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="spinulosus" status="sp. nov." subGenus="Micronialoe">
<emphasis id="A983EAAE521BFFE0281FFAB7FE7349CD" box="[287,413,1399,1423]" italics="true" pageId="4" pageNumber="455">Pterostichus</emphasis>
(
<emphasis id="A983EAAE521BFFE028AEFAB7FDC449CD" box="[430,554,1399,1423]" italics="true" pageId="4" pageNumber="455">Micronialoe</emphasis>
)
<emphasis id="A983EAAE521BFFE02B3BFAB7FD4849CD" box="[571,678,1399,1423]" italics="true" pageId="4" pageNumber="455">spinulosus</emphasis>
</taxonomicName>
<emphasis id="A983EAAE521BFFE02BAFFAB7FCED49CD" bold="true" box="[687,771,1399,1423]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="B2B057D5521BFFE02BAFFAB7FCED49CD" box="[687,771,1399,1423]" pageId="4" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="A983EAAE521BFFE02A53FAB6FC9849CC" bold="true" box="[851,886,1398,1422]" pageId="4" pageNumber="455">(A)</emphasis>
mouthpart, ventral view;
<emphasis id="A983EAAE521BFFE02D87FAB6FB4749CC" bold="true" box="[1159,1193,1398,1422]" pageId="4" pageNumber="455">(B)</emphasis>
ditto, lateral view;
<emphasis id="A983EAAE521BFFE02C79FAB6FA7249CC" bold="true" box="[1401,1436,1398,1422]" pageId="4" pageNumber="455">(C)</emphasis>
metepisternum;
<emphasis id="A983EAAE521BFFE0283BFA5AFEB149F0" bold="true" box="[315,351,1434,1458]" pageId="4" pageNumber="455">(D)</emphasis>
sternite VII. Scale bars: 0.3 mm.
</paragraph>
</caption>
<subSubSection id="D3ED6537521BFFE029C7F9C5FAB74BB1" pageId="4" pageNumber="455" type="diagnosis">
<paragraph id="9B4836BC521BFFE029C7F9C5FB1F4A25" blockId="4.[151,1437,1541,2036]" pageId="4" pageNumber="455">
<emphasis id="A983EAAE521BFFE029C7F9C5FEAC4A5D" bold="true" box="[199,322,1541,1567]" pageId="4" pageNumber="455">Diagnosis.</emphasis>
The new species is easily distinguished from all known species of the subgenus by: i) posteromedian area of male sternite VII distinctly rugose transversely; ii) median lobe of aedeagus truncate at apex in dorsal view; iii) internal sac of aedeagus with three sclerotized areas. Other significant characters are as follows.
</paragraph>
<paragraph id="9B4836BC521BFFE029C7F9B2FD834B7D" blockId="4.[151,1437,1541,2036]" pageId="4" pageNumber="455">Pronotum with sinuation on lateral margin faint or absent before basal angle; with two pairs of basal foveae. Posteromedian area of male sternite VII flat or partially depressed, with microsculpture consisting of irregular transverse meshes. Median lobe of aedeagus stout; left lateral margin below apical orifice almost linear; ventral carina present, narrowly rounded; sclerotized areas of internal sac consist of followings: i) C-shaped sclerite (CS) on median area; ii) small ventral sclerite (VS) on subapical area; iii) spiny patch (SP) consisting of numerous sclerotized compact spines on dorsal side.</paragraph>
<paragraph id="9B4836BC521BFFE029C7F889FAB74BB1" blockId="4.[151,1437,1541,2036]" pageId="4" pageNumber="455">
This species seems to be very similar to
<taxonomicName id="5CF74D3F521BFFE02B96F88AFC9A4B21" box="[662,884,1865,1891]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="bifoveolatus" subGenus="Micronialoe">
<emphasis id="A983EAAE521BFFE02B96F88AFD454B21" box="[662,683,1866,1891]" italics="true" pageId="4" pageNumber="455">P.</emphasis>
(
<emphasis id="A983EAAE521BFFE02BBCF88AFD3D4B21" box="[700,723,1866,1891]" italics="true" pageId="4" pageNumber="455">M</emphasis>
.)
<emphasis id="A983EAAE521BFFE02BECF889FC9A4B21" box="[748,884,1865,1891]" italics="true" pageId="4" pageNumber="455">bifoveolatus</emphasis>
</taxonomicName>
and
<taxonomicName id="5CF74D3F521BFFE02AADF88AFBA54B21" authorityName="Morita" authorityYear="2005" box="[941,1099,1865,1892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="461" phylum="Arthropoda" rank="species" species="kaniei" subGenus="Micronialoe">
<emphasis id="A983EAAE521BFFE02AADF88AFC504B21" box="[941,958,1866,1891]" italics="true" pageId="4" pageNumber="455">P</emphasis>
. (
<emphasis id="A983EAAE521BFFE02AD7F88AFC004B21" box="[983,1006,1866,1891]" italics="true" pageId="4" pageNumber="455">M</emphasis>
.)
<emphasis id="A983EAAE521BFFE02D07F889FBA54B21" box="[1031,1099,1865,1891]" italics="true" pageId="4" pageNumber="455">kaniei</emphasis>
</taxonomicName>
in their external morphology. Nevertheless, the new species can be easily distinguished from both by having the median lobe of the aedeagus with a truncate apex (
<figureCitation id="03CC2A39521BFFE02839F851FE7F4BEE" box="[313,401,1937,1964]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="4" pageNumber="455">Fig. 5D</figureCitation>
), whereas the others have a rounded apex in the dorsal view (
<figureCitation id="03CC2A39521BFFE02D33F851FB664BEE" box="[1075,1160,1937,1964]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="4" pageNumber="455">Fig. 7H</figureCitation>
;
<bibRefCitation id="FF664B4D521BFFE02D96F851FAA84BE9" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[1174,1350,1937,1964]" pageId="4" pageNumber="455" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">
Park
<emphasis id="A983EAAE521BFFE02DCFF852FB114BE9" box="[1231,1279,1937,1963]" italics="true" pageId="4" pageNumber="455">et al</emphasis>
. 1996
</bibRefCitation>
;
<bibRefCitation id="FF664B4D521BFFE02C51F851FF3D4B92" author="Morita, S." pageId="4" pageNumber="455" pagination="9 - 12" refId="ref7740" refString="Morita, S. (2005) A New Pterostichus (Coleoptera: Carabidae) from the Islands of Tsushima, West Japan. Entomological Review of Japan, 60 (1), 9 - 12." type="journal article" year="2005">Morita 2005</bibRefCitation>
). The new species has three sclerotized areas in the internal sac of the aedeagus (
<figureCitation id="03CC2A39521BFFE02D59F875FB384B92" box="[1113,1238,1973,2000]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="4" pageNumber="455">Figs 5AE</figureCitation>
), whereas
<taxonomicName id="5CF74D3F521BFFE02C4CF876FEF14BB1" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="bifoveolatus" subGenus="Micronialoe">
<emphasis id="A983EAAE521BFFE02C4CF876FAB34B8D" box="[1356,1373,1974,1999]" italics="true" pageId="4" pageNumber="455">P</emphasis>
. (
<emphasis id="A983EAAE521BFFE02C75F876FA624B8D" box="[1397,1420,1974,1999]" italics="true" pageId="4" pageNumber="455">M</emphasis>
.)
<emphasis id="A983EAAE521BFFE02997F819FEF14BB1" box="[151,287,2009,2035]" italics="true" pageId="4" pageNumber="455">bifoveolatus</emphasis>
</taxonomicName>
has two sclerotized areas (the feature not known for
<taxonomicName id="5CF74D3F521BFFE02A6BF81AFBEB4BB1" authorityName="Morita" authorityYear="2005" box="[875,1029,2009,2036]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="461" phylum="Arthropoda" rank="species" species="kaniei" subGenus="Micronialoe">
<emphasis id="A983EAAE521BFFE02A6BF81AFC924BB1" box="[875,892,2010,2035]" italics="true" pageId="4" pageNumber="455">P</emphasis>
. (
<emphasis id="A983EAAE521BFFE02A93F81AFC444BB1" box="[915,938,2010,2035]" italics="true" pageId="4" pageNumber="455">M</emphasis>
.)
<emphasis id="A983EAAE521BFFE02AC1F819FBEB4BB1" box="[961,1029,2009,2035]" italics="true" pageId="4" pageNumber="455">kaniei</emphasis>
</taxonomicName>
) (
<figureCitation id="03CC2A39521BFFE02D1DF819FB604BB6" box="[1053,1166,2009,2036]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="4" pageNumber="455">Figs 7EI</figureCitation>
;
<bibRefCitation id="FF664B4D521BFFE02D99F819FAA34BB1" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[1177,1357,2009,2036]" pageId="4" pageNumber="455" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">
Park
<emphasis id="A983EAAE521BFFE02DD3F81AFAEA4BB1" box="[1235,1284,2009,2035]" italics="true" pageId="4" pageNumber="455">et al</emphasis>
. 1996
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="CF886634521AFFE12997FD33FF024F6D" ID-DOI="http://doi.org/10.5281/zenodo.12190397" ID-Zenodo-Dep="12190397" httpUri="https://zenodo.org/record/12190397/files/figure.png" pageId="5" pageNumber="456" startId="5.[151,250,755,779]" targetBox="[151,1436,181,732]" targetPageId="5" targetType="figure">
<paragraph id="9B4836BC521AFFE12997FD33FF024F6D" blockId="5.[151,1437,755,816]" pageId="5" pageNumber="456">
<emphasis id="A983EAAE521AFFE12997FD33FEFD4F49" bold="true" box="[151,275,755,779]" pageId="5" pageNumber="456">FIGURE 4.</emphasis>
Microsculpture of
<taxonomicName id="5CF74D3F521AFFE128D5FD33FCB94F49" authority="Kim &amp; Suh, 2024" authorityName="Kim &amp; Suh" authorityYear="2024" box="[469,855,755,780]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="spinulosus" status="sp. nov." subGenus="Micronialoe">
<emphasis id="A983EAAE521AFFE128D5FD33FDBD4F49" box="[469,595,755,779]" italics="true" pageId="5" pageNumber="456">Pterostichus</emphasis>
(
<emphasis id="A983EAAE521AFFE12B61FD33FD334F49" box="[609,733,755,779]" italics="true" pageId="5" pageNumber="456">Micronialoe</emphasis>
)
<emphasis id="A983EAAE521AFFE12BECFD33FCB94F49" box="[748,855,755,779]" italics="true" pageId="5" pageNumber="456">spinulosus</emphasis>
</taxonomicName>
<emphasis id="A983EAAE521AFFE12A5DFD34FC394F49" bold="true" box="[861,983,755,780]" pageId="5" pageNumber="456">
<taxonomicNameLabel id="B2B057D5521AFFE12A5DFD34FC434F4E" box="[861,941,756,780]" pageId="5" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
(A)
</emphasis>
head;
<emphasis id="A983EAAE521AFFE12D19FD33FBD54F49" bold="true" box="[1049,1083,755,779]" pageId="5" pageNumber="456">(B)</emphasis>
pronotum;
<emphasis id="A983EAAE521AFFE12DB1FD33FB3A4F49" bold="true" box="[1201,1236,755,779]" pageId="5" pageNumber="456">(C)</emphasis>
elytron. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="CF886634521AFFE12997F895FD694BB8" ID-DOI="http://doi.org/10.5281/zenodo.12190399" ID-Zenodo-Dep="12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="5" pageNumber="456" startId="5.[151,250,1877,1901]" targetBox="[161,1426,850,1841]" targetPageId="5" targetType="figure">
<paragraph id="9B4836BC521AFFE12997F895FD694BB8" blockId="5.[151,1437,1876,2043]" pageId="5" pageNumber="456">
<emphasis id="A983EAAE521AFFE12997F895FEFA4B2F" bold="true" box="[151,276,1877,1901]" pageId="5" pageNumber="456">FIGURE 5.</emphasis>
Male and female genitalia of
<taxonomicName id="5CF74D3F521AFFE12B4CF895FC3F4B2F" authority="Kim &amp; Suh, 2024" authorityName="Kim &amp; Suh" authorityYear="2024" box="[588,977,1877,1901]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="spinulosus" status="sp. nov." subGenus="Micronialoe">
<emphasis id="A983EAAE521AFFE12B4CF895FD244B2F" box="[588,714,1877,1901]" italics="true" pageId="5" pageNumber="456">Pterostichus</emphasis>
(
<emphasis id="A983EAAE521AFFE12BD9F895FCBB4B2F" box="[729,853,1877,1901]" italics="true" pageId="5" pageNumber="456">Micronialoe</emphasis>
)
<emphasis id="A983EAAE521AFFE12A66F895FC3F4B2F" box="[870,977,1877,1901]" italics="true" pageId="5" pageNumber="456">spinulosus</emphasis>
</taxonomicName>
<emphasis id="A983EAAE521AFFE12AD8F895FBC44B2F" bold="true" box="[984,1066,1877,1901]" pageId="5" pageNumber="456">
<taxonomicNameLabel id="B2B057D5521AFFE12AD8F895FBC44B2F" box="[984,1066,1877,1901]" pageId="5" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male
<emphasis id="A983EAAE521AFFE12D71F895FB4A4B2F" bold="true" box="[1137,1188,1877,1901]" pageId="5" pageNumber="456">AH</emphasis>
, female (
<emphasis id="A983EAAE521AFFE12C07F895FAFF4B2F" bold="true" box="[1287,1297,1877,1901]" pageId="5" pageNumber="456">I</emphasis>
).
<emphasis id="A983EAAE521AFFE12C27F894FAA44B2E" bold="true" box="[1319,1354,1876,1900]" pageId="5" pageNumber="456">(A)</emphasis>
median lobe of aedeagus, right lateral view;
<emphasis id="A983EAAE521AFFE12B10F8B8FDDC4BD2" bold="true" box="[528,562,1912,1936]" pageId="5" pageNumber="456">(B)</emphasis>
ditto, dorsolateral view;
<emphasis id="A983EAAE521AFFE12A34F8B8FCB94BD2" bold="true" box="[820,855,1912,1936]" pageId="5" pageNumber="456">(C)</emphasis>
ditto, dorsal view;
<emphasis id="A983EAAE521AFFE12D20F8B8FBAA4BD2" bold="true" box="[1056,1092,1912,1936]" pageId="5" pageNumber="456">(D)</emphasis>
ditto, ventral view;
<emphasis id="A983EAAE521AFFE12C15F8B8FAD94BD2" bold="true" box="[1301,1335,1912,1936]" pageId="5" pageNumber="456">(E)</emphasis>
ditto, left lateral view;
<emphasis id="A983EAAE521AFFE1281AF85BFED54BF1" bold="true" box="[282,315,1947,1971]" pageId="5" pageNumber="456">(F)</emphasis>
left paramere;
<emphasis id="A983EAAE521AFFE128D3F85BFE164BF1" bold="true" box="[467,504,1947,1971]" pageId="5" pageNumber="456">(G)</emphasis>
right paramere;
<emphasis id="A983EAAE521AFFE12B9EF85BFD2D4BF1" bold="true" box="[670,707,1947,1971]" pageId="5" pageNumber="456">(H)</emphasis>
male sternite IX;
<emphasis id="A983EAAE521AFFE12A77F85BFC674BF1" bold="true" box="[887,905,1947,1971]" pageId="5" pageNumber="456">(I</emphasis>
) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm
<emphasis id="A983EAAE521AFFE12B9BF87EFD254B95" bold="true" box="[667,715,1982,2007]" pageId="5" pageNumber="456">AE</emphasis>
,
<emphasis id="A983EAAE521AFFE12BD6F87FFD044B95" bold="true" box="[726,746,1983,2007]" pageId="5" pageNumber="456">H</emphasis>
; 0.2 mm
<emphasis id="A983EAAE521AFFE12A47F87FFCB84B95" bold="true" box="[839,854,1983,2007]" pageId="5" pageNumber="456">F</emphasis>
,
<emphasis id="A983EAAE521AFFE12A62F87EFC984B94" bold="true" box="[866,886,1982,2006]" pageId="5" pageNumber="456">G</emphasis>
, 0.1 mm
<emphasis id="A983EAAE521AFFE12AD3F87FFC334B95" bold="true" box="[979,989,1983,2007]" pageId="5" pageNumber="456">I</emphasis>
. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina.
</paragraph>
</caption>
<subSubSection id="D3ED65375219FFE329C7FF56FE5A4D5F" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457" type="description">
<paragraph id="9B4836BC5219FFE229C7FF56FC874CF3" blockId="6.[151,1437,150,1185]" box="[199,873,150,177]" pageId="6" pageNumber="457">
<emphasis id="A983EAAE5219FFE229C7FF56FEB74CF2" bold="true" box="[199,345,150,176]" pageId="6" pageNumber="457">Description.</emphasis>
Body length:
<quantity id="5C0F9B595219FFE228F8FF56FD6F4CF3" box="[504,641,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.3" metricValueMin="7.2" pageId="6" pageNumber="457" unit="mm" value="7.75" valueMax="8.3" valueMin="7.2">7.28.3 mm</quantity>
, width: 2.73.0 mm.
</paragraph>
<paragraph id="9B4836BC5219FFE229C7FF7AFB094CBA" blockId="6.[151,1437,150,1185]" pageId="6" pageNumber="457">
Coloration black or brownish-black and shiny; antennae, legs, and lateral margin of pronotum and elytra dark brown; palpi yellowish brown, with apices distinctly paler; ventral side overall black (
<figureCitation id="03CC2A395219FFE22D5BFF1EFB374CBA" box="[1115,1241,222,248]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1997]" captionTargetBox="[155,1436,1248,1941]" captionTargetId="figure-445@2.[151,1436,1240,1950]" captionTargetPageId="2" captionText="FIGURE 2. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) habitus, dorsal view; (B) ditto, ventral view; C head and pronotum, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190391" httpUri="https://zenodo.org/record/12190391/files/figure.png" pageId="6" pageNumber="457">Figs 2A, B</figureCitation>
).
</paragraph>
<paragraph id="9B4836BC5219FFE229C7FEC2FC114D03" blockId="6.[151,1437,150,1185]" pageId="6" pageNumber="457">
<emphasis id="A983EAAE5219FFE229C7FEC2FEEA4D5E" box="[199,260,258,284]" italics="true" pageId="6" pageNumber="457">Head</emphasis>
(
<figureCitation id="03CC2A395219FFE22813FEC2FE814D5E" box="[275,367,258,284]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1997]" captionTargetBox="[155,1436,1248,1941]" captionTargetId="figure-445@2.[151,1436,1240,1950]" captionTargetPageId="2" captionText="FIGURE 2. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) habitus, dorsal view; (B) ditto, ventral view; C head and pronotum, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190391" httpUri="https://zenodo.org/record/12190391/files/figure.png" pageId="6" pageNumber="457">Figs 2C</figureCitation>
,
<figureCitation id="03CC2A395219FFE2287BFEC2FE214D5E" box="[379,463,258,284]" captionStart="FIGURE 3" captionStartId="4.[151,250,1399,1423]" captionTargetBox="[151,1436,189,1365]" captionTargetId="figure-35@4.[151,1436,181,1375]" captionTargetPageId="4" captionText="FIGURE 3. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) mouthpart, ventral view; (B) ditto, lateral view; (C) metepisternum; (D) sternite VII. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.12190393" httpUri="https://zenodo.org/record/12190393/files/figure.png" pageId="6" pageNumber="457">3A, 3B</figureCitation>
,
<figureCitation id="03CC2A395219FFE228DBFEC2FDEC4D5E" box="[475,514,258,284]" captionStart="FIGURE 4" captionStartId="5.[151,250,755,779]" captionTargetBox="[151,1436,181,732]" captionTargetId="figure-18@5.[151,1436,181,732]" captionTargetPageId="5" captionText="FIGURE 4. Microsculpture of Pterostichus (Micronialoe) spinulosus sp. nov. (A) head; (B) pronotum; (C) elytron. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190397" httpUri="https://zenodo.org/record/12190397/files/figure.png" pageId="6" pageNumber="457">4A</figureCitation>
). Eye weakly convex; tempus well developed, moderately convex, as long as eye; frontal impression deep, moderately curved toward anterior supraorbital pore.
</paragraph>
<paragraph id="9B4836BC5219FFE229C7FE8BFBBE4EC6" blockId="6.[151,1437,150,1185]" pageId="6" pageNumber="457">
<emphasis id="A983EAAE5219FFE229C7FE8BFED84D26" box="[199,310,331,356]" italics="true" pageId="6" pageNumber="457">Pronotum</emphasis>
(
<figureCitation id="03CC2A395219FFE2284AFE8AFE444D26" box="[330,426,330,356]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1997]" captionTargetBox="[155,1436,1248,1941]" captionTargetId="figure-445@2.[151,1436,1240,1950]" captionTargetPageId="2" captionText="FIGURE 2. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) habitus, dorsal view; (B) ditto, ventral view; C head and pronotum, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190391" httpUri="https://zenodo.org/record/12190391/files/figure.png" pageId="6" pageNumber="457">Figs 2C</figureCitation>
,
<figureCitation id="03CC2A395219FFE228BBFE8AFE334D26" box="[443,477,330,356]" captionStart="FIGURE 4" captionStartId="5.[151,250,755,779]" captionTargetBox="[151,1436,181,732]" captionTargetId="figure-18@5.[151,1436,181,732]" captionTargetPageId="5" captionText="FIGURE 4. Microsculpture of Pterostichus (Micronialoe) spinulosus sp. nov. (A) head; (B) pronotum; (C) elytron. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190397" httpUri="https://zenodo.org/record/12190397/files/figure.png" pageId="6" pageNumber="457">4B</figureCitation>
) widest near 2/5 from apex; apical angle narrowly rounded, slightly to distinctly protruded anteriorly; lateral margin with faint or without sinuation before basal angle; anterolateral seta before widest point, near 3/10 from apex; basal angle obtuse; anterior transverse impression faint or absent; posterior transverse impression slightly or moderately distinct; with two pairs of basal foveae; inner foveae deep, linear, almost parallel or slightly diverging anteriorly; outer foveae diverging, mostly clearly visible, but sometimes weakly depressed; area between inner foveae with or without sparse punctures and rugose longitudinally, sometimes almost smooth; area between inner fovea and outer fovea usually smooth, rarely with faint punctures near basal border. PW/HW = 1.601.68, mean 1.64; PW/PL = 1.121.19, mean 1.15; PW/PbW = 1.151.21, mean 1.18; PW/PaW = 1.441.55, mean 1.48; PbW/PaW = 1.201.35, mean
<quantity id="5C0F9B595219FFE22BEEFDAAFCD24EC6" box="[750,828,618,644]" metricMagnitude="-2" metricUnit="m" metricValue="3.2004" pageId="6" pageNumber="457" unit="in" value="1.26">1.26 in</quantity>
<specimenCount id="8DF1FD355219FFE22A43FDAAFC4B4EC6" box="[835,933,618,644]" count="11" pageId="6" pageNumber="457" type="male">11 males</specimenCount>
and
<specimenCount id="8DF1FD355219FFE22ADCFDAAFBA24EC6" box="[988,1100,618,644]" count="4" pageId="6" pageNumber="457" type="female">4 females</specimenCount>
.
</paragraph>
<paragraph id="9B4836BC5219FFE229C7FD4EFC814F56" blockId="6.[151,1437,150,1185]" pageId="6" pageNumber="457">
<emphasis id="A983EAAE5219FFE229C7FD4EFEE34EEA" box="[199,269,654,680]" italics="true" pageId="6" pageNumber="457">Elytra</emphasis>
(
<figureCitation id="03CC2A395219FFE2281AFD4EFE804EEA" box="[282,366,654,680]" captionStart="FIGURE 4" captionStartId="5.[151,250,755,779]" captionTargetBox="[151,1436,181,732]" captionTargetId="figure-18@5.[151,1436,181,732]" captionTargetPageId="5" captionText="FIGURE 4. Microsculpture of Pterostichus (Micronialoe) spinulosus sp. nov. (A) head; (B) pronotum; (C) elytron. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190397" httpUri="https://zenodo.org/record/12190397/files/figure.png" pageId="6" pageNumber="457">Fig. 4C</figureCitation>
) widest at 3/10 to 2/5 from base; striae deep, punctate from base to basal half; marginal umbilicate series composed of three groups, humeral group with 5 pores (rarely 6 pores), medial group with 1 pore (rarely 2 pores), and apical group with 7 pores (rarely 8 pores). EW/PW =1.111.21, mean 1.17; EL/PL = 1.992.10, mean 2.05; EL/EW = 1.451.57, mean
<quantity id="5C0F9B595219FFE22B0BFD3AFDB74F56" box="[523,601,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="3.8862" pageId="6" pageNumber="457" unit="in" value="1.53">1.53 in</quantity>
<specimenCount id="8DF1FD355219FFE22B60FD3AFD2A4F56" box="[608,708,762,788]" count="10" pageId="6" pageNumber="457" type="male">10 males</specimenCount>
and
<specimenCount id="8DF1FD355219FFE22BFBFD3AFC854F56" box="[763,875,762,788]" count="4" pageId="6" pageNumber="457" type="female">4 females</specimenCount>
.
</paragraph>
<paragraph id="9B4836BC5219FFE229C7FCDEFD514FC2" blockId="6.[151,1437,150,1185]" pageId="6" pageNumber="457">
<emphasis id="A983EAAE5219FFE229C7FCDEFEA44F7A" box="[199,330,798,824]" italics="true" pageId="6" pageNumber="457">Ventral side</emphasis>
(
<figureCitation id="03CC2A395219FFE22857FCDEFE5F4F7A" box="[343,433,798,824]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1997]" captionTargetBox="[155,1436,1248,1941]" captionTargetId="figure-445@2.[151,1436,1240,1950]" captionTargetPageId="2" captionText="FIGURE 2. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) habitus, dorsal view; (B) ditto, ventral view; C head and pronotum, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190391" httpUri="https://zenodo.org/record/12190391/files/figure.png" pageId="6" pageNumber="457">Figs 2B</figureCitation>
,
<figureCitation id="03CC2A395219FFE228BAFCDEFDE34F7A" box="[442,525,798,824]" captionStart="FIGURE 3" captionStartId="4.[151,250,1399,1423]" captionTargetBox="[151,1436,189,1365]" captionTargetId="figure-35@4.[151,1436,181,1375]" captionTargetPageId="4" captionText="FIGURE 3. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) mouthpart, ventral view; (B) ditto, lateral view; (C) metepisternum; (D) sternite VII. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.12190393" httpUri="https://zenodo.org/record/12190393/files/figure.png" pageId="6" pageNumber="457">3C, 3D</figureCitation>
). Posteromedian area of sternite VII flat or partially depressed, slightly to distinctly rugose, with microsculpture consisting of irregular transverse lines in male (
<figureCitation id="03CC2A395219FFE22AE7FC82FBAF4F1E" box="[999,1089,834,860]" captionStart="FIGURE 3" captionStartId="4.[151,250,1399,1423]" captionTargetBox="[151,1436,189,1365]" captionTargetId="figure-35@4.[151,1436,181,1375]" captionTargetPageId="4" captionText="FIGURE 3. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) mouthpart, ventral view; (B) ditto, lateral view; (C) metepisternum; (D) sternite VII. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.12190393" httpUri="https://zenodo.org/record/12190393/files/figure.png" pageId="6" pageNumber="457">Fig. 3D</figureCitation>
); not depressed, smooth, rarely transversely rugose near apical margin in female.
</paragraph>
<paragraph id="9B4836BC5219FFE229C7FC4AFB4348E3" blockId="6.[151,1437,150,1185]" pageId="6" pageNumber="457">
<emphasis id="A983EAAE5219FFE229C7FC4AFE854FE6" box="[199,363,906,932]" italics="true" pageId="6" pageNumber="457">Male genitalia</emphasis>
(
<figureCitation id="03CC2A395219FFE22879FC4AFE184FE7" box="[377,502,906,933]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="6" pageNumber="457">Figs 5AH</figureCitation>
,
<figureCitation id="03CC2A395219FFE22B04FC4AFDFA4FE6" box="[516,532,906,932]" captionStart="FIGURE 6" captionStartId="6.[151,250,1951,1975]" captionTargetBox="[151,1436,1208,1927]" captionTargetId="figure-499@6.[151,1436,1208,1927]" captionTargetPageId="6" captionText="FIGURE 6. Everted internal sac of Pterostichus (Micronialoe) spinulosus sp. nov. (A) left lateral view; (B) right lateral view. Scale bar: 0.3 mm. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190401" httpUri="https://zenodo.org/record/12190401/files/figure.png" pageId="6" pageNumber="457">6</figureCitation>
). Median lobe of aedeagus (
<figureCitation id="03CC2A395219FFE22A4FFC4AFC244FE7" box="[847,970,906,933]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="6" pageNumber="457">Figs 5AE</figureCitation>
) stout, well curved in lateral view, almost linear in dorsal view, ventral side distinctly curved behind basal bulb, and slightly curved apically; left lateral margin below apical orifice almost linear; apical orifice bent to left; terminal lamella very short; apex truncate in dorsal view; ventral carina weak to distinct, clearly visible in right dorsolateral view; internal sac with three sclerotized areas: i) C-shaped sclerite (CS) on median area, ii) small ventral sclerite (VS) on subapical area, and iii) spiny patch (SP) consisting of numerous sclerotized compact spines on dorsal side; everted internal sac (
<figureCitation id="03CC2A395219FFE22C51FBFEFA7A481A" box="[1361,1428,1086,1112]" captionStart="FIGURE 6" captionStartId="6.[151,250,1951,1975]" captionTargetBox="[151,1436,1208,1927]" captionTargetId="figure-499@6.[151,1436,1208,1927]" captionTargetPageId="6" captionText="FIGURE 6. Everted internal sac of Pterostichus (Micronialoe) spinulosus sp. nov. (A) left lateral view; (B) right lateral view. Scale bar: 0.3 mm. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190401" httpUri="https://zenodo.org/record/12190401/files/figure.png" pageId="6" pageNumber="457">Fig. 6</figureCitation>
) curved ventrally; left paramere (
<figureCitation id="03CC2A395219FFE22B05FBA2FDB6483F" box="[517,600,1122,1149]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="6" pageNumber="457">Fig. 5F</figureCitation>
) wide; right paramere (
<figureCitation id="03CC2A395219FFE22A64FBA2FC52483E" box="[868,956,1122,1149]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="6" pageNumber="457">Fig. 5G</figureCitation>
) slender, distinctly curved, ventral margin forming almost right angle and dorsal margin forming obtuse angle; sternite IX as in
<figureCitation id="03CC2A395219FFE22D50FB46FB4B48E3" box="[1104,1189,1158,1185]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="6" pageNumber="457">Fig. 5H</figureCitation>
.
</paragraph>
<caption id="CF8866345219FFE22997F85FFAE24B99" ID-DOI="http://doi.org/10.5281/zenodo.12190401" ID-Zenodo-Dep="12190401" httpUri="https://zenodo.org/record/12190401/files/figure.png" pageId="6" pageNumber="457" startId="6.[151,250,1951,1975]" targetBox="[151,1436,1208,1927]" targetPageId="6" targetType="figure">
<paragraph id="9B4836BC5219FFE22997F85FFAE24B99" blockId="6.[151,1437,1951,2011]" pageId="6" pageNumber="457">
<emphasis id="A983EAAE5219FFE22997F85FFEFD4BF5" bold="true" box="[151,275,1951,1975]" pageId="6" pageNumber="457">FIGURE 6.</emphasis>
Everted internal sac of
<taxonomicName id="5CF74D3F5219FFE22B07F85FFC644BF5" authority="Kim &amp; Suh, 2024" authorityName="Kim &amp; Suh" authorityYear="2024" box="[519,906,1951,1975]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="spinulosus" status="sp. nov." subGenus="Micronialoe">
<emphasis id="A983EAAE5219FFE22B07F85FFD6B4BF5" box="[519,645,1951,1975]" italics="true" pageId="6" pageNumber="457">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5219FFE22B94F85FFCFE4BF5" box="[660,784,1951,1975]" italics="true" pageId="6" pageNumber="457">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5219FFE22A1FF85FFC644BF5" box="[799,906,1951,1975]" italics="true" pageId="6" pageNumber="457">spinulosus</emphasis>
</taxonomicName>
<emphasis id="A983EAAE5219FFE22A91F85FFBE34BF5" bold="true" box="[913,1037,1951,1975]" pageId="6" pageNumber="457">
<taxonomicNameLabel id="B2B057D55219FFE22A91F85FFC0C4BF5" box="[913,994,1951,1975]" pageId="6" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
(A)
</emphasis>
left lateral view;
<emphasis id="A983EAAE5219FFE22DC1F85FFB0D4BF5" bold="true" box="[1217,1251,1951,1975]" pageId="6" pageNumber="457">(B)</emphasis>
right lateral view. Scale bar: 0.3 mm. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina.
</paragraph>
</caption>
<paragraph id="9B4836BC5218FFE329C7FF57FE5A4D5F" blockId="7.[151,1437,150,537]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FF57FE674CF3" box="[199,393,151,177]" italics="true" pageId="7" pageNumber="458">Female genitalia</emphasis>
(
<figureCitation id="03CC2A395218FFE3289AFF57FE054CF3" box="[410,491,150,177]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male AH, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm AE, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="7" pageNumber="458">Fig. 5I</figureCitation>
). Laterotergite IX with several setae along apical margin; basal stylomere with or without one small preapical spine; apical stylomere subtriangular, inner margin slightly curved, outer ventral margin bisinuate, with two thin nematiform setae subapically and three ensiform setae, of which one on dorsolateral and two on ventrolateral side.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE329C7FEE7FB6B4D03" box="[199,1157,294,321]" pageId="7" pageNumber="458" type="materials_examined">
<paragraph id="9B4836BC5218FFE329C7FEE7FB6B4D03" blockId="7.[151,1437,150,537]" box="[199,1157,294,321]" pageId="7" pageNumber="458">
<materialsCitation id="2B9F3CE15218FFE329C7FEE7FB6F4D03" ID-GBIF-Occurrence="4898139310" box="[199,1153,294,321]" country="South Korea" county="Daegu-si" location="Eastern" municipality="Mt. Choejeongsan" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<emphasis id="A983EAAE5218FFE329C7FEE7FE884D02" bold="true" box="[199,358,294,321]" pageId="7" pageNumber="458">
<typeStatus id="444C881E5218FFE329C7FEE7FEEC4D03" box="[199,258,295,321]" pageId="7" pageNumber="458">Type</typeStatus>
locality.
</emphasis>
<location id="9E2860675218FFE3286CFEE7FE2E4D03" LSID="urn:lsid:plazi:treatment:135E87AA521CFFE32997F9BAFD1F4E5B:9E2860675218FFE3286CFEE7FE2E4D03" box="[364,448,295,321]" country="South Korea" county="Daegu-si" municipality="Mt. Choejeongsan" name="Eastern" pageId="7" pageNumber="458">Eastern</location>
part of
<location id="9E2860675218FFE32B18FEE7FD5A4D03" LSID="urn:lsid:plazi:treatment:135E87AA521CFFE32997F9BAFD1F4E5B:9E2860675218FFE32B18FEE7FD5A4D03" box="[536,692,295,321]" country="South Korea" county="Daegu-si" municipality="Mt. Choejeongsan" name="Mt. Biseulsan" pageId="7" pageNumber="458">Mt. Biseulsan</location>
and
<collectingMunicipality id="7B2CACC65218FFE32BEAFEE7FC554D03" box="[746,955,295,321]" pageId="7" pageNumber="458">Mt. Choejeongsan</collectingMunicipality>
,
<collectingCounty id="72294E305218FFE32AC5FEE7FBC34D03" box="[965,1069,295,321]" pageId="7" pageNumber="458">Daegu-si</collectingCounty>
,
<collectingCountry id="E3E0762C5218FFE32D39FEE7FB6F4D03" box="[1081,1153,295,321]" name="South Korea" pageId="7" pageNumber="458">Korea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE329C7FE8BFE964DCB" pageId="7" pageNumber="458" type="etymology">
<paragraph id="9B4836BC5218FFE329C7FE8BFE964DCB" blockId="7.[151,1437,150,537]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FE8BFEA04D27" bold="true" box="[199,334,331,357]" pageId="7" pageNumber="458">Etymology.</emphasis>
The specific name refers to the characteristic of the new species having a spiny patch in the internal sac of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE329C7FE53FD1F4E5B" pageId="7" pageNumber="458" type="biology_ecology">
<paragraph id="9B4836BC5218FFE329C7FE53FD1F4E5B" blockId="7.[151,1437,150,537]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FE53FEC24DEF" bold="true" box="[199,300,403,429]" pageId="7" pageNumber="458">Ecology.</emphasis>
The adult specimens were observed beneath rocks in the forest valley or under stacked leaves on the side of the valley (
<figureCitation id="03CC2A395218FFE32865FE77FE484D93" box="[357,422,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1441,1465]" captionTargetBox="[151,1436,453,1417]" captionTargetId="figure-113@1.[151,1436,453,1417]" captionTargetPageId="1" captionText="FIGURE 1. Habitat of Pterostichus (Micronialoe) spinulosus sp. nov. in Mt. Biseulsan, Daegu, South Korea." figureDoi="http://doi.org/10.5281/zenodo.12190389" httpUri="https://zenodo.org/record/12190389/files/figure.png" pageId="7" pageNumber="458">Fig. 1</figureCitation>
). Older individuals with relatively poor conditions tended to be collected in June and fresh individuals tended to be collected in October at Mt. Biseulsan. Immature adults with a color of integument not fully darkened were collected in July at Mt. Choejeongsan.
</paragraph>
</subSubSection>
</treatment>
</document>