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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.2.e1557" ID-PMC="PMC4206778" ID-Pensoft-Pub="1314-2828--1557" ID-PubMed="25349524" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1314-2828--e1557" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Brazilian Trichoptera Checklist II" checkinTime="1553126666643" checkinUser="pensoft" docAuthor="Paprocki, Henrique &amp; Franca, Diogo" docDate="2014" docId="BE212615AD816824E588A7DD7ED1A3C3" docLanguage="en" docName="BiodivDatJour 2: e1557" docOrigin="Biodiversity Data Journal 2" docSource="http://dx.doi.org/10.3897/BDJ.2.e1557" docTitle="Mortoniella paraunota Blahnik &amp; Holzenthal 2011" docType="treatment" docVersion="3" lastPageNumber="1557" masterDocId="FFB01F7FBE1A5405C0068548FFA3FFAE" masterDocTitle="Brazilian Trichoptera Checklist II" masterLastPageNumber="1557" masterPageNumber="1557" pageNumber="1557" updateTime="1668122726353" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Brazilian Trichoptera Checklist II</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paprocki, Henrique</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Franca, Diogo</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1557</mods:start>
<mods:end>1557</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.2.e1557</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.2.e1557</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--1557</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:BE212615AD816824E588A7DD7ED1A3C3" httpUri="http://treatment.plazi.org/id/BE212615AD816824E588A7DD7ED1A3C3" lastPageNumber="1557" pageId="0" pageNumber="1557">
<subSubSection pageId="0" pageNumber="1557" type="nomenclature">
<paragraph pageId="0" pageNumber="1557">
<taxonomicName authority="Blahnik &amp; Holzenthal, 2011" authorityName="Blahnik &amp; Holzenthal" authorityYear="2011" class="Insecta" family="Glossosomatidae" genus="Mortoniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mortoniella paraunota" order="Trichoptera" pageId="0" pageNumber="1557" phylum="Arthropoda" rank="species" species="paraunota">Mortoniella paraunota Blahnik &amp; Holzenthal, 2011</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1557" type="distribution">
<paragraph pageId="0" pageNumber="1557">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1557">Santa Catarina</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1557" type="notes">
<paragraph pageId="0" pageNumber="1557">Notes</paragraph>
<paragraph pageId="0" pageNumber="1557">
<bibRefCitation author="Blahnik, R. J." journalOrPublisher="Zootaxa" pageId="0" pageNumber="1557" pagination="1 - 75" title="Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae)" volume="2851" year="2011">Blahnik and Holzenthal 2011</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1038.55258" ID-GBIF-Dataset="d0e1b800-7531-48cf-967c-3f8b3fcc3a12" ID-PMC="PMC9341518" ID-Pensoft-Pub="1313-2970-1038-105" ID-Pensoft-UUID="8F8AF721FC8751DAAFF28FF3B81BA7A4" ID-PubMed="36051890" ID-ZooBank="8FDAC6A330AB4D339C009189A44FD8EE" ModsDocID="1313-2970-1038-105" checkinTime="1621457762112" checkinUser="pensoft" docAuthor="Li, Yang, Achterberg, Cornelis van &amp; Chen, Xue-xin" docDate="2020" docId="BE215305939153B1A5E26EE2B84FE25C" docLanguage="en" docName="ZooKeys 1038: 105-178" docOrigin="ZooKeys 1038" docPubDate="2020-05-19" docSource="http://dx.doi.org/10.3897/zookeys.1038.55258" docTitle="Parallobracon prolatus Li &amp; Achterberg &amp; Chen 2020, sp. nov." docType="treatment" docUuid="2E01D4E3-D4CE-48D3-82E7-3A22D5E957F1" docUuidSource="ZooBank" docVersion="4" id="8F8AF721FC8751DAAFF28FF3B81BA7A4" lastPageNumber="105" masterDocId="8F8AF721FC8751DAAFF28FF3B81BA7A4" masterDocTitle="A new genus and eight newly recorded genera of Braconinae Nees (Hymenoptera, Braconidae) from China, with descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="105" pageNumber="105" updateTime="1668150272162" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new genus and eight newly recorded genera of Braconinae Nees (Hymenoptera, Braconidae) from China, with descriptions of fourteen new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Yang</mods:namePart>
<mods:affiliation>State Key Lab of Rice Biology, Zhejiang University, Hangzhou 310058, China &amp; Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Achterberg, Cornelis van</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation>Zhejiang Provincial Key Laboratory of Biology of Crop Pathogens and Insects, Zhejiang University, Hangzhou 310058, China &amp; Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Xue-xin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9109-8853</mods:nameIdentifier>
<mods:affiliation>State Key Lab of Rice Biology, Zhejiang University, Hangzhou 310058, China &amp; Ministry of Agriculture Key Lab of Molecular Biology of Crop Pathogens and Insects, Zhejiang University, Hangzhou 310058, China &amp; Zhejiang Provincial Key Laboratory of Biology of Crop Pathogens and Insects, Zhejiang University, Hangzhou 310058, China &amp; Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, China</mods:affiliation>
<mods:nameIdentifier type="email">xxchen@zju.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-05-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1038</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>105</mods:start>
<mods:end>178</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1038.55258</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1038.55258</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1038-105</mods:identifier>
<mods:identifier type="ZooBank">8FDAC6A330AB4D339C009189A44FD8EE</mods:identifier>
<mods:identifier type="Pensoft-UUID">8F8AF721FC8751DAAFF28FF3B81BA7A4</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182238057" LSID="urn:lsid:zoobank.org:act:2E01D4E3-D4CE-48D3-82E7-3A22D5E957F1" httpUri="http://treatment.plazi.org/id/BE215305939153B1A5E26EE2B84FE25C" lastPageNumber="105" pageId="0" pageNumber="105">
<subSubSection pageId="0" pageNumber="105" type="nomenclature">
<paragraph pageId="0" pageNumber="105">
<taxonomicName LSID="http://zoobank.org/2E01D4E3-D4CE-48D3-82E7-3A22D5E957F1" authority="Li &amp; Achterberg &amp; Chen, 2020" authorityName="Li &amp; Achterberg &amp; Chen" authorityYear="2020" class="Insecta" family="Braconidae" genus="Parallobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parallobracon prolatus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="prolatus" status="sp. nov.">Parallobracon prolatus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="105">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Parallobracon prolatus gen. et sp. nov., ♀, holotype, habitus, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure27" httpUri="https://binary.pensoft.net/fig/544876" pageId="0" pageNumber="105">Figures 27</figureCitation>
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="materials_examined">
<paragraph pageId="0" pageNumber="105">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="105">
<materialsCitation collectingDate="1935-06-26" country="China" location="Zhu Ruzuo" municipality="Hangzhou" specimenCount="♀" stateProvince="Zhejiang Prov." typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount>♀</specimenCount>
,
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Zhejiang">Zhejiang Prov.</collectingRegion>
,
<collectingMunicipality>Hangzhou</collectingMunicipality>
,
<collectingDate value="1935-06-26">26.VI.1935</collectingDate>
,
<location LSID="urn:lsid:plazi:treatment:BE215305939153B1A5E26EE2B84FE25C:17E8401C63DC7BBCB22C7436CA2257ED" country="China" municipality="Hangzhou" name="Zhu Ruzuo" stateProvince="Zhejiang Prov.">Zhu Ruzuo</location>
(ZJUH)
</materialsCitation>
.
<materialsCitation collectingDate="1934-06-17" country="China" location="Zhu Ruzuo" municipality="Hangzhou" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Zhejiang Prov." typeStatus="Paratype">
<typeStatus>Paratype</typeStatus>
.
<specimenCount type="female">1♀</specimenCount>
<specimenCount type="male">1♂</specimenCount>
,
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Zhejiang">Zhejiang Prov.</collectingRegion>
,
<collectingMunicipality>Hangzhou</collectingMunicipality>
,
<collectingDate value="1934-06-17">17.VI.1934</collectingDate>
,
<location LSID="urn:lsid:plazi:treatment:BE215305939153B1A5E26EE2B84FE25C:983D4D48D08E7F4A37BE7DBA602C1EB8" country="China" municipality="Hangzhou" name="Zhu Ruzuo" stateProvince="Zhejiang Prov.">Zhu Ruzuo</location>
(ZJUH)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="diagnosis">
<paragraph pageId="0" pageNumber="105">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="105">
T II of the new species is similar to that of
<taxonomicName authorityName="Belokobylskij" authorityYear="2000" class="Insecta" family="Braconidae" genus="Cyanopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyanopterus oriens" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="oriens">
<emphasis italics="true" pageId="0" pageNumber="105">Cyanopterus oriens</emphasis>
</taxonomicName>
Belokobylskij, 2000, from Far East Russia (e.g., size and shape of medio-basal area and with pair of diverging depressions), but can be separated by having first discal cell of fore wing parallel-sided and vein 1-SR+M straight (widened basally and vein 1-SR+M weakly bent in
<taxonomicName lsidName="C. oriens" pageId="0" pageNumber="105" rank="species" species="oriens">
<emphasis italics="true" pageId="0" pageNumber="105">C. oriens</emphasis>
</taxonomicName>
), vein 1r-m of hind wing ca. 5
<normalizedToken originalValue="×">x</normalizedToken>
longer than vein 2-SC+R (ca. equal in
<taxonomicName lsidName="C. oriens" pageId="0" pageNumber="105" rank="species" species="oriens">
<emphasis italics="true" pageId="0" pageNumber="105">C. oriens</emphasis>
</taxonomicName>
), vein 1-SR of fore wing distinctly oblique and pointing basad of vein cu-a (less oblique and pointing to vein cu-a in
<taxonomicName lsidName="C. oriens" pageId="0" pageNumber="105" rank="species" species="oriens">
<emphasis italics="true" pageId="0" pageNumber="105">C. oriens</emphasis>
</taxonomicName>
), second submarginal cell of fore wing widened distally (parallel-sided in
<taxonomicName lsidName="C. oriens" pageId="0" pageNumber="105" rank="species" species="oriens">
<emphasis italics="true" pageId="0" pageNumber="105">C. oriens</emphasis>
</taxonomicName>
) and apex of scapus strongly protruding ventrally (slightly protruding in
<taxonomicName lsidName="C. oriens" pageId="0" pageNumber="105" rank="species" species="oriens">
<emphasis italics="true" pageId="0" pageNumber="105">C. oriens</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="description">
<paragraph pageId="0" pageNumber="105">Description.</paragraph>
<paragraph pageId="0" pageNumber="105">Holotype, ♀, length of body 7.6 mm, of fore wing 7.7 mm, of ovipositor sheath 4.9 mm.</paragraph>
<caption doi="10.3897/zookeys.1038.55258.figure27" httpUri="https://binary.pensoft.net/fig/544876" pageId="0" pageNumber="105" start="Figure 27" startId="F27">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 27.</emphasis>
<taxonomicName authorityName="Li &amp; Achterberg &amp; Chen" authorityYear="2020" class="Insecta" family="Braconidae" genus="Parallobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parallobracon prolatus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="prolatus">
<emphasis italics="true" pageId="0" pageNumber="105">Parallobracon prolatus</emphasis>
</taxonomicName>
gen. et sp. nov., ♀, holotype, habitus, lateral view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105" start="Figure 28" startId="F28">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 28.</emphasis>
<taxonomicName authorityName="Li &amp; Achterberg &amp; Chen" authorityYear="2020" class="Insecta" family="Braconidae" genus="Parallobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parallobracon prolatus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="prolatus">
<emphasis italics="true" pageId="0" pageNumber="105">Parallobracon prolatus</emphasis>
</taxonomicName>
gen. et sp. nov., ♀, holotype (
<emphasis bold="true" pageId="0" pageNumber="105">a-j</emphasis>
), paratype (
<emphasis bold="true" pageId="0" pageNumber="105">k</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="105">a</emphasis>
fore wing
<emphasis bold="true" pageId="0" pageNumber="105">b</emphasis>
hind wing
<emphasis bold="true" pageId="0" pageNumber="105">c</emphasis>
mesosoma, lateral view
<emphasis bold="true" pageId="0" pageNumber="105">d</emphasis>
mesosoma, dorsal view
<emphasis bold="true" pageId="0" pageNumber="105">e</emphasis>
metasoma, dorsal view
<emphasis bold="true" pageId="0" pageNumber="105">f</emphasis>
hind leg, lateral view
<emphasis bold="true" pageId="0" pageNumber="105">g</emphasis>
head, anterior view
<emphasis bold="true" pageId="0" pageNumber="105">h</emphasis>
head, dorsal view
<emphasis bold="true" pageId="0" pageNumber="105">i</emphasis>
head, lateral view
<emphasis bold="true" pageId="0" pageNumber="105">j</emphasis>
first metasomal tergite, dorsal view
<emphasis bold="true" pageId="0" pageNumber="105">k</emphasis>
apex of ovipositor, lateral view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Head</emphasis>
.
</emphasis>
Antenna incomplete, only remaining with scapus and pedicellus; malar suture with dense short setae (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28i</figureCitation>
); clypeus height: inter-tentorial distance: tentorio-ocular distance = 3: 6: 7; clypeus with sparse short setae; eye weakly emarginated (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28g</figureCitation>
); face weakly convex medially and weakly granulate, with some sparse punctures laterally (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28g</figureCitation>
); eye height: shortest distance between eyes: head width = 17: 20: 40; frons largely smooth, weakly concave behind antennal sockets, with a strong median groove (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28h</figureCitation>
); vertex smooth, but with some sparse short setae; minimum distance between posterior ocelli: minimum diameter of elliptical posterior ocellus: minimum distance between posterior ocellus and eye = 1: 1: 2; temples largely glabrous except for a few short setae, and subparallel behind eyes (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28h</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Mesosoma</emphasis>
.
</emphasis>
Length of mesosoma 1.9
<normalizedToken originalValue="×">x</normalizedToken>
its height (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28c</figureCitation>
); notauli impressed anteriorly half (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28d</figureCitation>
); mesoscutum smooth, with a few sparse setae (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28d</figureCitation>
); scutellar sulcus narrow, moderately deep, and with crenulae (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28d</figureCitation>
); scutellum with sparse short setae posteriorly; metanotum strongly convex medially (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28d</figureCitation>
); propodeum smooth, without longitudinal carinae or groove, with sparse setae medially, and with dense long setae laterally (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28d</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Wings</emphasis>
.
</emphasis>
Fore wing (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28a</figureCitation>
): SR1: 3-SR: r = 21: 13: 5; 1-SR+M more or less straight, and 1.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than 1-M; 2-SR: 3-SR: r-m = 14: 26: 13; angle between 1-SR and C+SC+R ca. 75°; m-cu straight; 2-SR+M rather short; cu-a slightly postfurcal. Hind wing (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28b</figureCitation>
): SC+R1: 2-SC+R: 1r-m = 17: 3: 13.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Legs</emphasis>
.
</emphasis>
Length of fore femur: tibia: tarsus = 27: 31: 45; length of hind femur: tibia: basitarsus = 37: 53: 21; length of femur, tibia and basitarsus of hind leg 3.5, 6.6 and 5.3
<normalizedToken originalValue="×">x</normalizedToken>
their maximum width, respectively (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28f</figureCitation>
); hind tibial spurs 0.35 and 0.40
<normalizedToken originalValue="×">x</normalizedToken>
as long as hind basitarsus.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Metasoma</emphasis>
.
</emphasis>
Length of T I 1.1
<normalizedToken originalValue="×">x</normalizedToken>
its apical width, median area convex, anteriorly half smooth and posteriorly half coarsely sculptured (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28j</figureCitation>
); lateral grooves of T I sparsely crenulate (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28j</figureCitation>
); T II largely smooth except medially (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28e</figureCitation>
); triangular medio-basal area of T II large and smooth, with a few short oblique carinae connected laterally, and acute apically, but not attached with medio-longitudinal carina; antero-lateral areas of T II developed and smooth, anterior grooves moderately wide and sparsely crenulate (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28e</figureCitation>
); second suture deep and crenulate, wide and weakly curved medially, narrow laterally (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28e</figureCitation>
); T III with antero-lateral areas; T III-V with crenulate transverse subposterior groove (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28e</figureCitation>
); T III-VII largely smooth, and with some spare short setae posteriorly; hypopygium acute apically, not reaching level of apex of metasoma; ovipositor sheath 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as fore wing.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Colour</emphasis>
.
</emphasis>
Head and mesosoma largely reddish yellow (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Parallobracon prolatus gen. et sp. nov., ♀, holotype, habitus, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure27" httpUri="https://binary.pensoft.net/fig/544876" pageId="0" pageNumber="105">27</figureCitation>
); antenna, eye, mandible apically, stemmaticum, propodeum medially and posteriorly, blackish brown (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28d, g, h</figureCitation>
); fore legs (but claws dark brown) reddish yellow, middle and hind legs dark brown (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28f</figureCitation>
); metasoma and ovipositor sheath black brown (Figs
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Parallobracon prolatus gen. et sp. nov., ♀, holotype, habitus, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure27" httpUri="https://binary.pensoft.net/fig/544876" pageId="0" pageNumber="105">27</figureCitation>
,
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28e</figureCitation>
); wing membrane greyish brown, pterostigma and veins dark brown (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Parallobracon prolatus gen. et sp. nov., ♀, holotype (a-j), paratype (k) a fore wing b hind wing c mesosoma, lateral view d mesosoma, dorsal view e metasoma, dorsal view f hind leg, lateral view g head, anterior view h head, dorsal view i head, lateral view j first metasomal tergite, dorsal view k apex of ovipositor, lateral view." figureDoi="10.3897/zookeys.1038.55258.figure28" httpUri="https://binary.pensoft.net/fig/544877" pageId="0" pageNumber="105">28a, b</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Variation</emphasis>
.
</emphasis>
Length of body of female 7.6-7.7 mm, of fore wing of female 7.7 mm, and of ovipositor sheath 4.2-4.9 mm; antenna of paratype female incomplete, with 30 antennomeres remaining; third antennomere 1.7
<normalizedToken originalValue="×">x</normalizedToken>
longer than its maximum width, 1.2 and 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than fourth and fifth, respectively, the latter 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; ovipositor sheath 0.5-0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as fore wing.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Male</emphasis>
.
</emphasis>
Length of body of male 7.8 mm, of fore wing of male 7.9 mm; antenna of male with 44 antennomeres; Length of mesosoma 1.7
<normalizedToken originalValue="×">x</normalizedToken>
its height; fore wing vein cu-a interstitial; length of T I 1.0
<normalizedToken originalValue="×">x</normalizedToken>
its apical width; other characters as in female.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="biology_ecology">
<paragraph pageId="0" pageNumber="105">Biology.</paragraph>
<paragraph pageId="0" pageNumber="105">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="distribution">
<paragraph pageId="0" pageNumber="105">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="105">China (Zhejiang).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="etymology">
<paragraph pageId="0" pageNumber="105">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="105">
Named after the long and slender scapus which strong elongated ventrally:
<emphasis italics="true" pageId="0" pageNumber="105">prolatus</emphasis>
is Latin for elongated.
</paragraph>
</subSubSection>
</treatment>
</document>

134
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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.53.33425" ID-GBIF-Dataset="8d11df69-1cba-4e11-9e7a-feb1502f0d55" ID-PMC="PMC6551344" ID-Pensoft-Pub="1314-4049-53-23" ID-PubMed="31191113" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-53-23" ModsDocOrigin="MycoKeys 53" ModsDocTitle="A regional study of the genus Phyllopsora (Ramalinaceae) in Asia and Melanesia" checkinTime="1559196956708" checkinUser="pensoft" docAuthor="Kistenich, Sonja, Bendiksby, Mika, Vairappan, Charles S., Weerakoon, Gothamie, Wijesundara, Siril, Wolseley, Patricia A. &amp; Timdal, Einar" docDate="2019" docId="BE218BAEEB977AA29575579E1F486FB3" docLanguage="en" docName="MycoKeys 53: 23-72" docOrigin="MycoKeys 53" docSource="http://dx.doi.org/10.3897/mycokeys.53.33425" docTitle="Phyllopsora confusa Swinscow &amp; Krog" docType="treatment" docVersion="4" lastPageNumber="23" masterDocId="7A11FB47FFCE0E346802FFBD522F5F1E" masterDocTitle="A regional study of the genus Phyllopsora (Ramalinaceae) in Asia and Melanesia" masterLastPageNumber="72" masterPageNumber="23" pageNumber="23" updateTime="1668136241517" updateUser="ExternalLinkService">
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<mods:title>A regional study of the genus Phyllopsora (Ramalinaceae) in Asia and Melanesia</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kistenich, Sonja</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bendiksby, Mika</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vairappan, Charles S.</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Weerakoon, Gothamie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wijesundara, Siril</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wolseley, Patricia A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Timdal, Einar</mods:namePart>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
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<mods:start>23</mods:start>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.53.33425</mods:identifier>
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<treatment ID-GBIF-Taxon="157322373" LSID="urn:lsid:plazi:treatment:BE218BAEEB977AA29575579E1F486FB3" httpUri="http://treatment.plazi.org/id/BE218BAEEB977AA29575579E1F486FB3" lastPageNumber="23" pageId="0" pageNumber="23">
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<taxonomicName authority="Swinscow &amp; Krog" class="Lecanoromycetes" family="Ramalinaceae" genus="Phyllopsora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phyllopsora confusa" order="Lecanorales" pageId="0" pageNumber="23" phylum="Ascomycota" rank="species" species="confusa">Phyllopsora confusa Swinscow &amp; Krog</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="description">
<paragraph pageId="0" pageNumber="23">Description.</paragraph>
<paragraph pageId="0" pageNumber="23">
<bibRefCitation author="Swinscow, TDV" journalOrPublisher="The Lichenologist" pageId="0" pageNumber="23" pagination="203 - 247" title="The genus Phyllopsora, with a report on East African species." url="https://doi.org/10.1017/S0024282981000315" volume="13" year="1981">Swinscow and Krog (1981)</bibRefCitation>
,
<bibRefCitation author="Timdal, E" journalOrPublisher="Mycotaxon" pageId="0" pageNumber="23" pagination="57 - 90" title="Further studies on African species of the lichen genus Phyllopsora (Lecanorales)." volume="77" year="2001">Timdal and Krog (2001)</bibRefCitation>
,
<bibRefCitation pageId="0" pageNumber="23">Elix (2009)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="distribution">
<paragraph pageId="0" pageNumber="23">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="23">
Pantropical (
<bibRefCitation pageId="0" pageNumber="23">Brako 1991</bibRefCitation>
;
<bibRefCitation author="Timdal, E" journalOrPublisher="Mycotaxon" pageId="0" pageNumber="23" pagination="57 - 90" title="Further studies on African species of the lichen genus Phyllopsora (Lecanorales)." volume="77" year="2001">Timdal and Krog 2001</bibRefCitation>
;
<bibRefCitation pageId="0" pageNumber="23">Elix 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="remarks">
<paragraph pageId="0" pageNumber="23">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="23">
This species is characterized by the small, lacinulate squamules lacking lichen substances (Fig. 4A), but may be difficult to separate from, for example,
<taxonomicName lsidName="P. foliata" pageId="0" pageNumber="23" rank="species" species="foliata">P. foliata</taxonomicName>
and
<taxonomicName lsidName="P. mediocris" pageId="0" pageNumber="23" rank="species" species="mediocris">P. mediocris</taxonomicName>
. It is also possible that some of the specimens we have left undetermined belong in this species. We have sequenced nine specimens from Asia and Melanesia (Table 2), in addition to the holotype from Kenya, and those specimens make up the core in our concept of this species. The accessions of
<taxonomicName lsidName="P. confusa" pageId="0" pageNumber="23" rank="species" species="confusa">P. confusa</taxonomicName>
form a strongly supported clade with
<taxonomicName lsidName="P. loekoesii" pageId="0" pageNumber="23" rank="species" species="loekoesii">P. loekoesii</taxonomicName>
in the phylogenetic tree (Fig. 1). The two specimens from Papua New Guinea (6360 and 6361) fall in between the
<taxonomicName lsidName="P. confusa" pageId="0" pageNumber="23" rank="species" species="confusa">P. confusa</taxonomicName>
and
<taxonomicName lsidName="P. loekoesii" pageId="0" pageNumber="23" rank="species" species="loekoesii">P. loekoesii</taxonomicName>
clade (Suppl. material 1: Fig. S1) and show an intermediate morphology. Further specimens are needed to investigate the possible synonymy of these two species.
<taxonomicName class="Lecanoromycetes" family="Ramalinaceae" genus="Phyllopsora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phyllopsora confusa" order="Lecanorales" pageId="0" pageNumber="23" phylum="Ascomycota" rank="species" species="confusa">Phyllopsora confusa</taxonomicName>
is new to Indonesia, Japan, Malaysia, Taiwan, and Thailand.
</paragraph>
<caption pageId="0" pageNumber="23">
<paragraph pageId="0" pageNumber="23">
Figure 4. Species of
<taxonomicName class="Lecanoromycetes" family="Ramalinaceae" genus="Phyllopsora" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phyllopsora" order="Lecanorales" pageId="0" pageNumber="23" phylum="Ascomycota" rank="genus">Phyllopsora</taxonomicName>
occurring in Asia and Melanesia. A
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</mods:role>
<mods:namePart>Kassie, Abeje</mods:namePart>
<mods:affiliation>Ethiopian Biodiversity Institute, Addis Ababa, Ethiopia &amp; Addis Ababa University, Addis Ababa, Ethiopia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Boissinot, Stephane</mods:namePart>
<mods:affiliation>New York University Abu Dhabi, Saadiyat Island, Abu Dhabi, UAE</mods:affiliation>
<mods:nameIdentifier type="email">stephane.boissinot@nyu.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1016</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>77</mods:start>
<mods:end>141</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1016.59699</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1016.59699</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1016-77</mods:identifier>
<mods:identifier type="ZooBank">A462DE7344284857973CBCFD17294D15</mods:identifier>
<mods:identifier type="Pensoft-UUID">D7F1A3CBC5AC592DADE3787D4EC62BF9</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="177383271" LSID="urn:lsid:zoobank.org:act:CCF36B05-F7FC-436B-8858-765B6F682127" httpUri="http://treatment.plazi.org/id/BE21FD2A9D415C909D56E7446862297C" lastPageNumber="77" pageId="0" pageNumber="77">
<subSubSection pageId="0" pageNumber="77" type="nomenclature">
<paragraph pageId="0" pageNumber="77">
<taxonomicName LSID="http://zoobank.org/CCF36B05-F7FC-436B-8858-765B6F682127" authority="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot, 2021" authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka" status="sp. nov.">Ptychadena beka</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="77">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="type material">
<paragraph pageId="0" pageNumber="77">Type material.</paragraph>
<paragraph pageId="0" pageNumber="77">
<emphasis bold="true" pageId="0" pageNumber="77">
<emphasis italics="true" pageId="0" pageNumber="77">Holotype</emphasis>
.
</emphasis>
Adult male (SB291) collected by S. Goutte and J. Reyes-Velasco on 5 June 2018 southwest of Nekemte (
<geoCoordinate degrees="8.9950" direction="north" orientation="latitude" precision="5" value="8.995">8.9950°N</geoCoordinate>
,
<geoCoordinate degrees="36.4955" direction="east" orientation="longitude" precision="5" value="36.4955">36.4955°E</geoCoordinate>
, 2213 m a.s.l.).
<emphasis bold="true" pageId="0" pageNumber="77">
<emphasis italics="true" pageId="0" pageNumber="77">Paratypes</emphasis>
.
</emphasis>
one male (15-5) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot on 3 August 2015 west of Holeta Genet (
<geoCoordinate degrees="9.0508" direction="north" orientation="latitude" precision="5" value="9.0508">9.0508°N</geoCoordinate>
,
<geoCoordinate degrees="38.4312" direction="east" orientation="longitude" precision="5" value="38.4312">38.4312°E</geoCoordinate>
, 2433 m a.s.l.), one male (15-283) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot on 14 August 2015 southwest of Nekemte (
<geoCoordinate degrees="8.9742" direction="north" orientation="latitude" precision="5" value="8.9742">8.9742°N</geoCoordinate>
,
<geoCoordinate degrees="36.4906" direction="east" orientation="longitude" precision="5" value="36.4906">36.4906°E</geoCoordinate>
, 2243 m a.s.l.), one female (16-1) collected by J. Reyes-Velasco and S. Boissinot on 4 July 2016 west of Holeta Genet (
<geoCoordinate degrees="9.05078" direction="north" orientation="latitude" precision="1" value="9.05078">9.05078°N</geoCoordinate>
,
<geoCoordinate degrees="38.4312" direction="east" orientation="longitude" precision="5" value="38.4312">38.4312°E</geoCoordinate>
, 2386 m a.s.l.), one female (SB268) collected by S. Goutte and J. Reyes-Velasco on 3 June 2018 northwest of Bonga (
<geoCoordinate degrees="7.5085" direction="north" orientation="latitude" precision="5" value="7.5085">7.5085°N</geoCoordinate>
,
<geoCoordinate degrees="36.0637" direction="east" orientation="longitude" precision="5" value="36.0637">36.0637°E</geoCoordinate>
, 2038 m a.s.l.), 3 females (SB270, SB272 and SB276) collected by S. Goutte and J. Reyes-Velasco on 3 June 2018 south of Ambo (8.9671-8.9922°N, 37.7951-37.8471°E, 1945-2170 m a.s.l.), one female (SB282) collected by S. Goutte and J. Reyes-Velasco on 4 June 2018 southwest of Nekemte (
<geoCoordinate degrees="8.9950" direction="north" orientation="latitude" precision="5" value="8.995">8.9950°N</geoCoordinate>
,
<geoCoordinate degrees="36.4955" direction="east" orientation="longitude" precision="5" value="36.4955">36.4955°E</geoCoordinate>
, 2213 m a.s.l.), one female (SB287) and one male (SB292) collected by S. Goutte and J. Reyes-Velasco on 5 June 2018 southwest of Nekemte (
<geoCoordinate degrees="8.9950" direction="north" orientation="latitude" precision="5" value="8.995">8.9950°N</geoCoordinate>
,
<geoCoordinate degrees="36.4955" direction="east" orientation="longitude" precision="5" value="36.4955">36.4955°E</geoCoordinate>
, 2213 m a.s.l.), one male (SB387) collected by S. Goutte and J. Reyes-Velasco on 11 June 2018 south of
<normalizedToken originalValue="Gech’a">Gech'a</normalizedToken>
(
<geoCoordinate degrees="7.5544" direction="north" orientation="latitude" precision="5" value="7.5544">7.5544°N</geoCoordinate>
,
<geoCoordinate degrees="35.4148" direction="east" orientation="longitude" precision="5" value="35.4148">35.4148°E</geoCoordinate>
, 1936 m a.s.l.), one male (SB471) collected by S. Goutte and Y. Bourgeois on 21 June 2018 east of Iteya (
<geoCoordinate degrees="8.1275" direction="north" orientation="latitude" precision="5" value="8.1275">8.1275°N</geoCoordinate>
,
<geoCoordinate degrees="39.2732" direction="east" orientation="longitude" precision="5" value="39.2732">39.2732°E</geoCoordinate>
, 2138 m a.s.l.), one female (SB566) collected by S. Goutte and Y. Bourgeois north of Gohatsion (
<geoCoordinate degrees="10.01967" direction="north" orientation="latitude" precision="1" value="10.01967">10.01967°N</geoCoordinate>
,
<geoCoordinate degrees="38.2494" direction="east" orientation="longitude" precision="5" value="38.2494">38.2494°E</geoCoordinate>
, 2448 m a.s.l.), one male (SB576) collected by S. Goutte and Y. Bourgeois on 7 July 2018 north of Gebre Guracha (
<geoCoordinate degrees="9.8664" direction="north" orientation="latitude" precision="5" value="9.8664">9.8664°N</geoCoordinate>
,
<geoCoordinate degrees="38.3758" direction="east" orientation="longitude" precision="5" value="38.3758">38.3758°E</geoCoordinate>
, 2596 m a.s.l.), two males (SB614, SB617) collected by S. Goutte and Y. Bourgeois on 14 July 2018 east of Holeta Genet (
<geoCoordinate degrees="9.0682" direction="north" orientation="latitude" precision="5" value="9.0682">9.0682°N</geoCoordinate>
,
<geoCoordinate degrees="38.5214" direction="east" orientation="longitude" precision="5" value="38.5214">38.5214°E</geoCoordinate>
, 2397 m a.s.l.). All specimens are deposited at ZNHM.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="diagnosis">
<paragraph pageId="0" pageNumber="77">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="77">
A medium-sized species (male (9) SVL 37.9
<normalizedToken originalValue="±">+/-</normalizedToken>
2.6 mm, female (8) SVL 43.2
<normalizedToken originalValue="±">+/-</normalizedToken>
2.9 mm) of the
<taxonomicName baseAuthorityName="Ahl" baseAuthorityYear="1924" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena neumanni" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="neumanni">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena neumanni</emphasis>
</taxonomicName>
species group (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Ptychadena beka sp. nov. A live male holotype (SB 291; left) and female paratype (SB 270, right) B dorsal and ventral views of the same individuals (male left, female right) after euthanasia and before fixation." figureDoi="10.3897/zookeys.1016.59699.figure18" httpUri="https://binary.pensoft.net/fig/508434" pageId="0" pageNumber="77">18</figureCitation>
), distinguished by the following combination of characters: (1) relatively short hind limbs (male TL/SVL 0.51
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03, female TL/SVL 0.57
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03), (2) tympanum with a light vertical bar, (3) medium or wide vertebral stripe always present, (4) vocal sacs of most males are bicolored, from light to dark grey posteriorly and from yellow to cream anteriorly, rarely, they are light grey, (5) forearms of adult males not thickened.
</paragraph>
<caption doi="10.3897/zookeys.1016.59699.figure18" httpUri="https://binary.pensoft.net/fig/508434" pageId="0" pageNumber="77" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="77">
<emphasis bold="true" pageId="0" pageNumber="77">Figure 18.</emphasis>
<taxonomicName authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena beka</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="77">A</emphasis>
live male holotype (SB291; left) and female paratype (SB270, right)
<emphasis bold="true" pageId="0" pageNumber="77">B</emphasis>
dorsal and ventral views of the same individuals (male left, female right) after euthanasia and before fixation.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="comparisons">
<paragraph pageId="0" pageNumber="77">Comparison.</paragraph>
<paragraph pageId="0" pageNumber="77">
<taxonomicName authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena beka</emphasis>
</taxonomicName>
sp. nov. is smaller than
<taxonomicName lsidName="P. cooperi" pageId="0" pageNumber="77" rank="species" species="cooperi">
<emphasis italics="true" pageId="0" pageNumber="77">P. cooperi</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. goweri" pageId="0" pageNumber="77" rank="species" species="goweri">
<emphasis italics="true" pageId="0" pageNumber="77">P. goweri</emphasis>
</taxonomicName>
and larger than
<taxonomicName lsidName="P. nana" pageId="0" pageNumber="77" rank="species" species="nana">
<emphasis italics="true" pageId="0" pageNumber="77">P. nana</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. robeensis" pageId="0" pageNumber="77" rank="species" species="robeensis">
<emphasis italics="true" pageId="0" pageNumber="77">P. robeensis</emphasis>
</taxonomicName>
sp. nov. Hand, feet, tibias, thighs and snout are of similar dimensions than
<taxonomicName lsidName="P. amharensis" pageId="0" pageNumber="77" rank="species" species="amharensis">
<emphasis italics="true" pageId="0" pageNumber="77">P. amharensis</emphasis>
</taxonomicName>
and shorter than those of
<taxonomicName lsidName="P. delphina" pageId="0" pageNumber="77" rank="species" species="delphina">
<emphasis italics="true" pageId="0" pageNumber="77">P. delphina</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. goweri" pageId="0" pageNumber="77" rank="species" species="goweri">
<emphasis italics="true" pageId="0" pageNumber="77">P. goweri</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena beka</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="P. amharensis" pageId="0" pageNumber="77" rank="species" species="amharensis">
<emphasis italics="true" pageId="0" pageNumber="77">P. amharensis</emphasis>
</taxonomicName>
by a wider inter-orbital distance. It can be distinguished from
<taxonomicName lsidName="P. erlangeri" pageId="0" pageNumber="77" rank="species" species="erlangeri">
<emphasis italics="true" pageId="0" pageNumber="77">P. erlangeri</emphasis>
</taxonomicName>
by a larger tympanum, the light stripe or blotch on the tympanum and the bicolored vocal sacs in adult males.
<taxonomicName authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena beka</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="P. levenorum" pageId="0" pageNumber="77" rank="species" species="levenorum">
<emphasis italics="true" pageId="0" pageNumber="77">P. levenorum</emphasis>
</taxonomicName>
by longer thighs, larger tympanum and longer snout. The head is wider and the tibias are longer than in
<taxonomicName lsidName="P. doro" pageId="0" pageNumber="77" rank="species" species="doro">
<emphasis italics="true" pageId="0" pageNumber="77">P. doro</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="description">
<paragraph pageId="0" pageNumber="77">Description of the holotype.</paragraph>
<paragraph pageId="0" pageNumber="77">
A medium-sized (SVL 36.3 mm) adult male (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Ptychadena beka sp. nov. A live male holotype (SB 291; left) and female paratype (SB 270, right) B dorsal and ventral views of the same individuals (male left, female right) after euthanasia and before fixation." figureDoi="10.3897/zookeys.1016.59699.figure18" httpUri="https://binary.pensoft.net/fig/508434" pageId="0" pageNumber="77">18</figureCitation>
). Head slightly wider than long. Snout projecting beyond the lower jaw. Interorbital distance 0.62
<normalizedToken originalValue="×">x</normalizedToken>
the eye diameter. Internarial distance 1.12
<normalizedToken originalValue="×">x</normalizedToken>
interorbital distance. Tympanum 0.67
<normalizedToken originalValue="×">x</normalizedToken>
eye diameter. Finger tips not expanded but rounded, with very small subarticular tubercles. Finger formula: I&lt;II&lt;IV&lt;III. Hand free of webbing, palmar tubercle absent. Nuptial pads light grey. Hindlimbs moderately elongated, with tibia length 0.51
<normalizedToken originalValue="×">x</normalizedToken>
snout-vent length. Foot as long as thigh and shorter than tibia (FL/TL0.97). Toe tips rounded. Subarticular tubercles extremely small. Inner metatarsal tubercle present, external absent. Toe formula: I&lt;II&lt;V&lt;III&lt;IV. Foot webbing formula: Ie(1), IIi/e(1-1), IIIi/e(1-2), IVi/e(2-2), Vi(2). Two light brown, continuous lateral ridges, two continuous and four interrupted dorsal ridges. No vertebral nor sacral ridges. No warts on the body. Small and round light brown warts on tibias and thighs.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="coloration of the holotype in life">
<paragraph pageId="0" pageNumber="77">Coloration of the holotype in life.</paragraph>
<paragraph pageId="0" pageNumber="77">Dorsal ground color brown with a few small, elongated dark brown blotches symmetrically distributed on the dorsolateral ridges. Wide light brown vertebral stripe, doubled with a thin, clearer line from the tip of the snout to the vent. Dark brown canthal stripe from the tip of the snout to the back of the jaw. Faint light vertical stripe on the otherwise brown tympanum. Upper and lower lip cream with irregular brown markings.</paragraph>
<paragraph pageId="0" pageNumber="77">Iris bicolored, with upper third light silver-grey, and the lower two thirds dark brown. Small irregular dark olive brown blotches on greyish flanks. Throat and chest cream, ventrum, ventral sides of the thighs and of the tibias light yellow. Two small symmetrical dark brown blotches on the antero-ventral side of the shoulders. Thin, light longitudinal stripe on the tibias. Few white round warts on the tibias. Hind limbs brown with dark olive bars over the thighs, tibias, and feet. A few very small round white dots around the groin. Back of the thighs dark brown irregularly molted with yellow. Vocal sacs dark grey anteriorly, yellow posteriorly.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="coloration of the holotype in preservative">
<paragraph pageId="0" pageNumber="77">Coloration of the holotype in preservative.</paragraph>
<paragraph pageId="0" pageNumber="77">Dorsal ground color grey with a few oval dark brown blotches symmetrically distributed on the dorsolateral ridges. Wide light brown vertebral stripe, doubled with a lighter thin line from the tip of the snout to the vent. Dark brown canthal stripe from the tip of the snout to the back of the jaw. Faint vertical stripe on the otherwise brown tympanum. Upper lip and lower lip cream heavily blotched with grey. Flanks brown with a few black blotches underlined with a cream bar. Throat, ventrum, ventral side of the thighs and tibias cream, with the chest and throat dusted with grey. Two symmetrical dark brown blotches on the ventral side of the shoulders. Faint, thin longitudinal stripe on the tibias. Faint brown bars on the thigh and tibia. A few small round white spots forming a line on the posterior side of tibia and foot. Back of thighs molted light and dark brown. Vocal sacs dark grey anteriorly, cream posteriorly. Nuptial pads light grey.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="variation">
<paragraph pageId="0" pageNumber="77">Variations.</paragraph>
<paragraph pageId="0" pageNumber="77">In life, background color varies from yellowish brown to greyish brown. Dorsal ridges vary in number and definition. In all examined specimens, median ridges were continuous from the back of the head to the groin. Postpalpebral fold usually interrupted in the middle of the back, sometimes continuing on the lower back by a ridge or multiple aligned warts. Short latero-dorsal fold almost always present, sometimes fractioned. Lateral ridges, briefly interrupted in a few specimens, most often continuous, from orange-brown to cream. All specimens examined had small dark brown or black blotches distributed symmetrically on the dorsal ridges, in very few specimens, those blotches were very small or barely visible. All examined individuals have a wide, generally around a lighter thin stripe. The light blotch on the tympanum is generally small and more conspicuous in some specimens than in others. The thin cream longitudinal stripe on the tibia may be more or less conspicuous and extended to half of the thigh in some individuals. Tibias, thighs and feet posteriorly barred with more or less defined brown or light brown markings. Yellow and brown marbling on the posterior side of the thighs more or less contrasted and almost absent in some individuals. Ventrum and throat uniformly white to light yellow. Vocal sacs grey, bicolored cream to yellow and light grey to grey. Small warts over the back and flanks in ca. 10% of adult males examined.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="etymology">
<paragraph pageId="0" pageNumber="77">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="77">
The specific name corresponds to the translation of the word
<normalizedToken originalValue="“enough”">&quot;enough&quot;</normalizedToken>
or
<normalizedToken originalValue="“that’s">&quot;that's</normalizedToken>
it&quot; in Amharic (በቃ), in reference to the controversial taxonomic history of the group that we hope has come to an end. It is an invariable noun used in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="habitat">
<paragraph pageId="0" pageNumber="77">Habitat, distribution, and natural history.</paragraph>
<paragraph pageId="0" pageNumber="77">
<taxonomicName authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena beka</emphasis>
</taxonomicName>
sp. nov. has a wide distribution range extending both sides of the Great Rift Valley, although most records were made west of the GRV (6.14-10.01°N, 35.41-39.27°E). It is restricted to moderate altitudes, from 1695 m to 2596 m a.s.l. (based on 106 barcoded individuals). In the north, its range is limited by the Blue Nile valley and the northernmost specimens have been found just north of Gohatsion. The westernmost population has been found in
<normalizedToken originalValue="Gech’a">Gech'a</normalizedToken>
. Two populations have been found southeast of the GRV, near Iteya and south of Irba Muda. GPS coordinates for all examined specimens are given in Suppl. material 3: Table S1.
<taxonomicName authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena beka</emphasis>
</taxonomicName>
sp. nov. is found in syntopy with
<taxonomicName lsidName="P. erlangeri" pageId="0" pageNumber="77" rank="species" species="erlangeri">
<emphasis italics="true" pageId="0" pageNumber="77">P. erlangeri</emphasis>
</taxonomicName>
at the higher end of its altitudinal range, notably near Fitche, Holeta, between Ambo and Wonchi, and possibly near Assela. Within the genus,
<taxonomicName lsidName="P. beka" pageId="0" pageNumber="77" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">P. beka</emphasis>
</taxonomicName>
sp. nov. is also sympatric with
<taxonomicName lsidName="P. neumanni" pageId="0" pageNumber="77" rank="species" species="neumanni">
<emphasis italics="true" pageId="0" pageNumber="77">P. neumanni</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. delphina" pageId="0" pageNumber="77" rank="species" species="delphina">
<emphasis italics="true" pageId="0" pageNumber="77">P. delphina</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="P. doro" pageId="0" pageNumber="77" rank="species" species="doro">
<emphasis italics="true" pageId="0" pageNumber="77">P. doro</emphasis>
</taxonomicName>
in the west.
</paragraph>
<paragraph pageId="0" pageNumber="77">Males are found calling at in shallow roadside puddles or agricultural fields. Males can be found vocalizing very close from one another, sometimes in important numbers. Calling activity depends on rainfall and is highest during rainy months. Calling usually starts after 22:00, and sometimes as late as 02:00 in dry weather, and ceases before dawn. Numerous, small bicolored eggs are laid in the same water body.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="advertisement call">
<paragraph pageId="0" pageNumber="77">Advertisement call.</paragraph>
<paragraph pageId="0" pageNumber="77">
The call of
<taxonomicName authorityName="Goutte &amp; Reyes-Velasco * &amp; Freilich &amp; Kassie &amp; Boissinot" authorityYear="2021" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychadena beka" order="Anura" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">Ptychadena beka</emphasis>
</taxonomicName>
sp. nov. (5 males, 128 calls) is composed of a single pulsed note of 447
<normalizedToken originalValue="±">+/-</normalizedToken>
112 ms in duration, containing 31
<normalizedToken originalValue="±">+/-</normalizedToken>
10.6 pulses. Within calls, pulses are grouped by 3-5 pulses, with the first and the last pulses of each pulse group notably lower in amplitude than the other pulses. Low-amplitude single pulses are sometimes present between pulse groups. Amplitude increases gradually during the call, peaking at 266
<normalizedToken originalValue="±">+/-</normalizedToken>
78 ms, after which it drops. Call dominant frequency is 2491
<normalizedToken originalValue="±">+/-</normalizedToken>
129 Hz.
</paragraph>
<paragraph pageId="0" pageNumber="77">
The call of
<taxonomicName lsidName="P. beka" pageId="0" pageNumber="77" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">P. beka</emphasis>
</taxonomicName>
sp. nov. can be distinguished from those of
<taxonomicName lsidName="P. cooperi" pageId="0" pageNumber="77" rank="species" species="cooperi">
<emphasis italics="true" pageId="0" pageNumber="77">P. cooperi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. amharensis" pageId="0" pageNumber="77" rank="species" species="amharensis">
<emphasis italics="true" pageId="0" pageNumber="77">P. amharensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. doro" pageId="0" pageNumber="77" rank="species" species="doro">
<emphasis italics="true" pageId="0" pageNumber="77">P. doro</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName lsidName="P. neumanni" pageId="0" pageNumber="77" rank="species" species="neumanni">
<emphasis italics="true" pageId="0" pageNumber="77">P. neumanni</emphasis>
</taxonomicName>
(type A and B) by the distinguishable pulses composing the calls. It is also distinct from the calls of
<taxonomicName lsidName="P. delphina" pageId="0" pageNumber="77" rank="species" species="delphina">
<emphasis italics="true" pageId="0" pageNumber="77">P. delphina</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. robeensis" pageId="0" pageNumber="77" rank="species" species="robeensis">
<emphasis italics="true" pageId="0" pageNumber="77">P. robeensis</emphasis>
</taxonomicName>
sp. nov. by its short inter-pulse intervals. The call of
<taxonomicName lsidName="P. beka" pageId="0" pageNumber="77" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">P. beka</emphasis>
</taxonomicName>
sp. nov. differs from the calls of
<taxonomicName lsidName="P. delphina" pageId="0" pageNumber="77" rank="species" species="delphina">
<emphasis italics="true" pageId="0" pageNumber="77">P. delphina</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. erlangeri" pageId="0" pageNumber="77" rank="species" species="erlangeri">
<emphasis italics="true" pageId="0" pageNumber="77">P. erlangeri</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. levenorum" pageId="0" pageNumber="77" rank="species" species="levenorum">
<emphasis italics="true" pageId="0" pageNumber="77">P. levenorum</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="P. goweri" pageId="0" pageNumber="77" rank="species" species="goweri">
<emphasis italics="true" pageId="0" pageNumber="77">P. goweri</emphasis>
</taxonomicName>
by its higher dominant frequency. Finally, the call of
<taxonomicName lsidName="P. beka" pageId="0" pageNumber="77" rank="species" species="beka">
<emphasis italics="true" pageId="0" pageNumber="77">P. beka</emphasis>
</taxonomicName>
sp. nov. differs from the call of
<taxonomicName lsidName="P. nana" pageId="0" pageNumber="77" rank="species" species="nana">
<emphasis italics="true" pageId="0" pageNumber="77">P. nana</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. erlangeri" pageId="0" pageNumber="77" rank="species" species="erlangeri">
<emphasis italics="true" pageId="0" pageNumber="77">P. erlangeri</emphasis>
</taxonomicName>
by its longer duration.
</paragraph>
</subSubSection>
</treatment>
</document>