diff --git a/data/DC/A5/0A/DCA50A963FBB5746B34316561BB29571.xml b/data/DC/A5/0A/DCA50A963FBB5746B34316561BB29571.xml new file mode 100644 index 00000000000..e837494c1e6 --- /dev/null +++ b/data/DC/A5/0A/DCA50A963FBB5746B34316561BB29571.xml @@ -0,0 +1,4255 @@ + + + +Cryptantha acrimuricata (Boraginaceae), a distinctive new taxon of series Muricatae + + + +Author + +Simpson, Michael G. +0000-0002-6197-2132 +Department of Biology, San Diego State University, San Diego, California 92182, USA + + + +Author + +Simpson, Lee M. +Department of Biology, San Diego State University, San Diego, California 92182, USA + + + +Author + +André, James M. +https://orcid.org/0009-0000-0433-7493 +Granite Mountains Desert Research Center, University of California, Riverside, PO Box 101, Kelso, California 92309, USA + +text + + +PhytoKeys + + +2024 + +2024-12-30 + + +250 + + +193 +213 + + + +journal article +10.3897/phytokeys.250.138635 + + + + + +Cryptantha acrimuricata +J. M. André, L. M. Simpson, & M. G. Simpson + +sp. nov. + + + + +Type. + + + +United States +• +California +, +San Bernardino County +, +New York Mountains +: +Mojave Natl. Preseve +; +Caruthers Canyon +, +2.3 road miles north of junction New York Mtns Rd, at base of steep ascent to Giant Ledge Mine (historic), along closed dirt road paralleling perennially moist drainage, California, San Bernardino Co. +, + +35.24875 +, +- 115.299217 + +, + +1768 m + +elevation, + +pinyon-juniper-oak woodland, among granitic outcrops, associates: + +Pinus monophylla + +, + +Quercus turbinella + +, + +Garrya flavescens + +, + +Arctostaphylos pungens + +, + +Bouteoua gracilis + +, + +Yucca baccata + +, + +Brickellia californica + +, + +Opuntia phaeacantha + +, + +Dudleya saxosa aloides + + +, + +J. M. André +40457 + +, + +10 May 2019 + +( + +holotype +: +RSA 0633491 + +; + +isotypes +: +ASU + +, + +GMDRC + +, + +SBBG + +, + +SD + +, + +SDSU + +, + +UC + +, + +UCR + +) + +. + + + + +Diagnosis. + + + +Cryptantha acrimuricata + +resembles + +C. muricata + +and varieties in having ovoid fruiting calyces with nutlets that are typically four per fruit, homomorphic, ovate in shape, with margins thickened and bearing tuberculate to muricate processes, with a median dorsal ridge, and with surfaces densely tuberculate to muricate. + +Cryptantha acrimuricata + +differs from + +C. muricata + +and varieties in having dense whitish-strigose stems, with spreading trichomes absent to sparsely present, and in having nutlets with relatively large and densely spaced, conic, often whitish, muricate processes with relatively wide bases and sharp apices, surfaces between the processes often shiny. + + + + +Description. + + +(Figs +1 +, +2 +). +Plants +annual herbs, +10–35 cm +tall, mostly gray-green. +Root +a taproot, brown to red-purple, sometimes staining paper. +Stems +generally erect to ascending, some plants with a dominant, erect primary axis bearing several short to elongate, ascending, secondary branches, others with numerous, subequal, inclined to ascending branches arising from plant base; stem surface light green, vestiture strigose with dense, white, tapered, antrorsely appressed, basally white-pustulate trichomes, ca. +0.5–0.7 mm +long; hispid vestiture scattered to absent, if present with few, white, tapered, spreading (horizontal to inclined), basally white-pustulate trichomes, ca. +1 mm +long. +Leaves +mostly gray-green, sessile, linear to narrowly oblanceolate, midrib grooved adaxially, thickened abaxially, apex rounded to obtuse, both surfaces and margin appressed to ascending hirsute, trichomes white pustulate, especially in basal leaves; basal-most leaf pair apparently opposite, perhaps remnants of cotyledons, all others alternate-spiral; basal leaves +2–5 cm +× ca. +2–3 mm +, in a loose rosette withering at anthesis, cauline leaves +1–2 cm +× ca. +1 mm +, extending to base of cymule inflorescence units, slightly reduced upwards. +Inflorescence +cymules mostly in clusters of 2–3 (bifurcate or trifurcate), occasionally solitary, rarely up to 5, a terminal flower / fruit typically at junction of cymule bases; bracts occasionally at base of cymules, these linear to narrowly oblong; cymules elongated in fruit, 2–12 flowers per unit, lower fruits not touching at maturity. +Flowers +mostly ebracteate, pedicellate, pedicels in fruit more elongate, ascending, thickened apically, +0.5–1 mm +long, surface densely hirsute, trichomes horizontal to ascending. +Calyx +symmetric, ca. +3 mm +long at anthesis, sepals distinct; fruiting calyx ovoid, base truncate, apex acuminate, +3.5–5 mm +long, sepals lance-ovate to lanceolate, apices erect; abaxial surface densely hirsute ascending along margins, marginal surfaces hirsute appressed, midrib strongly thickened, hispid, the trichomes stout, straight, tapered, pustulate, inclined, horizontal, or reflexed; adaxial surface glabrous basally, appressed short hirsute on surface of extreme apex. +Corolla +rotate, inconspicuous, white, fornices white, tube as long as calyx, limb +1–2 mm +wide, not showy, appendages present ca. 1 / 3 from base of corolla tube in vertical line with anthers. +Androecium +of five stamens, attached at slightly different levels ca. 2 / 3 from base of corolla tube between and below fornices; anthers ca. +0.4 mm +long, ellipsoid, dithecal, introrsely dehiscent, dorsifixed; filaments filiform, ca. +0.1 mm +long. +Gynoecium +four-lobed, lobes ca. +0.3 mm +long, globose, style gynobasic, ca. +0.7 mm +long, protruding ca. +0.4 mm +beyond ovary; stigma minute, discoid. +Gynobase +narrowly pyramidal, extending nearly to nutlet apices; style extending +0.1–0.3 mm +beyond nutlet apices. +Nutlets +4, erect, homomorphic, ca. +2–2.4 mm +× +1.7 mm +(at widest width), whitish to tan, sometimes dark brown-mottled, surface typically shiny, appearing varnished, ovate, abaxially flattened to low convex, adaxially shallowly 2 - planed-convex, base truncate, margins sharp-angled with a thickened, usually whitish rim appearing “ beaded ” / ” toothed ” from marginal tuberculate to muricate protuberances, nutlet apex acute-rounded (blunt), surface irregularly transversely ridged, papillate and densely white tuberculate to muricate, these processes generally widely conic, densely spaced on surface, especially abaxially; spinal ridge present, sometimes obscure; attachment scar edges abutted at upper two-thirds, strongly bifurcate in lower one-third, delimiting a narrowly triangular, basal areole. + + + + + + + +Cryptantha acrimuricata + +. Images of collection: +M. G. Simpson 4186 +, 14 April 2019, + +SDSU +22965 + +A +herbarium specimen. Note dominant primary stem terminating in bifurcate cymules +B – E, H, I +field photographs +B +primary stem, showing cauline leaves and lateral branches +C +close-up of primary stem, the vestiture primarily strigose plus scattered spreading-hirsute +D +extreme close-up of primary stem, showing predominantly whitish strigose vestiture (trichomes antrorsely oriented) +E +close-up of cymule tip, showing relatively small corolla, this one slightly <2 mm wide +F, G +flower removed and dehydrated +F +corolla, showing fornices, anthers adnate to corolla tube, and basal appendages +G +gynoecium, showing 4 - lobed ovary, gynobasic style, and discoid stigma +H +cymule of primary stem +I +close-up of mature fruiting calyces. Note ovoid shape and spreading-hispid, pustulate trichomes of thickened sepal midribs. + + + + + + + + +Cryptantha acrimuricata + +. Images from holotype specimen: +J. M. André 40457 +, 10 May 2019, + +RSA +0633491 + +A +nutlet, in (left to right) dorsal, ventral, and lateral views. Note ovate shape (dorsal and ventral outlines); truncate base; thickened, “ toothed ” or “ beaded ” margins; whitish, conic surface tubercles; and narrow ventral groove, which is widely bifurcate at the base, delimiting small areole +B +fruiting calyx, showing appressed marginal trichomes and spreading hispid along midrib +C +fruit opened, showing prominent style, protruding well beyond nutlets +D +herbarium specimen. + + + + + +Distribution and habitat. + + + +Cryptantha acrimuricata + +occurs in the eastern part of the Peninsular Ranges, in the eastern and central Transverse Ranges, in higher elevations of the Mohave and Sonoran Deserts, and in scattered regions of the Sierra +Nevada +, of +California +and +Arizona +in the +United States +and Baja +California +in Mexico (Fig. +3 +). Its habitat ranges from sandy to coarsely gravelly substrates, usually granitic, sometimes limestone, sometimes adjacent to boulders, of chaparral, desert scrub / wash, pinyon-juniper woodland, or pine / oak woodland vegetation. The documented elevation range is +180–2621 m +(average +1270 m +). + + + + + + +Distribution maps of + +Cryptantha acrimuricata + +from specimens cited here, plus of the three varieties of + +C. muricata + +from preliminary data, in two scales +A +showing overall distribution and +B +in a larger scale map highlighting the Transverse Ranges and northern Peninsular Ranges. See text for distribution summaries of the taxa. Arrow indicates location of holotype / isotypes specimens of + +Cryptantha acrimuricata + +. Map data @ 2024 Google, INEGI. + + + + + +Phenology. + +Based on data from available specimens, the species typically flowers April – July, rarely in the fall following warm-seasonal rains (J. M. André, personal observation). + + + +Rarity and conservation status. + + + +Cryptantha acrimuricata + +has been somewhat sparsely collected compared to the close relative + +Cryptantha muricata + +. Where encountered, populations of + +C. acrimuricata + +tend to be relatively isolated and restricted in areal extent, with few total numbers of individual plants. Though we are not aware of any imminent threats to existing populations, based upon its fairly limited distribution and small populations, this taxon may warrant conservation status. + + + + +Etymology. + + +The specific epithet + +acrimuricata + +means “ sharply muricate, ” from the Latin +acri +-, sharp, and + +muricata + +(derived from Latin +murex +, conch), hard-pointed. The name refers to the stout, widely-based, and sharply pointed conic tubercles, shaped like miniature mountain peaks, as contrasted with the more cylindrical or rounded tubercles of the varieties of + +Cryptantha muricata + +. + + + + +Suggested common name. + + +We suggest Sharp-Pointed Prickly + +Cryptantha + +as a common name. + + + +<typeStatus id="32B8EF05FAAC7AB913F36864ECA6DEF8" type="paratype">Paratypes</typeStatus> + + +Arranged by country, then alphabetically by collector; * = estimated from label data. + + +Mexico +• +Baja California +. +San Rafael +, +Sierra San Pedro Mártir +; +west side of Cerro Blanco +; +3 k SSW of San Rafael, Baja California +, + +31.09333 +, +- 115.65167 + +, + +1550 m + +elevation, + +west facing slope, granite rock and gravel, associates: + +Adenostoma fasciculatum + +, + +Arctostaphylos + +, + +Chorizanthe fimbriata + +, + +Eriogonum fasciculatum + +, + +Lotus rigidus + +, + +Lupinus concinnus + +, + +Mimulus pilosus + +, + +Rhus ovata + +, + +Salvia apiana + +, + +Swertia albomarginata + +, and + +Trichostemma parishii + + +, + +M. A. Baker +12915 with +Robert Johnson + +, + +12 May 1998 + +( + +ASU +0014635 + +!, + +BCMEX +11466 + +!) + +; • + +Cerro San Luis +, +Baja California +, + +29.3167 +, +- 114.1167 + +, + +1300 m + +elevation, +open gravelly place, north side, gravelly +, + +R. Moran +10296 with +Jim Henrickson + +, + +2 March 1963 + +( + +RSA +165004 + +!, + +SD +54579 + +, + +UC +1235600 + +) + +; • + +Sierra Juarez +, + +5 km +W of + +La Rumorosa +, +Baja California +, +32.55, - 116.1 +, + +1325 m + +elevation, +colony under piñon, granitic area, associates: Piñon +, + +R. Moran +24108 + +, + +15 May 1977 + +( + +CAS +612817 + +!, + +SD +97068 + +!) + +; • + +Sierra Juarez +, + +1 km +NW of + +Tres Pozos +, +Baja California +, + +32.37917 +, +- 116.075 + +, + +1350 m + +elevation, +under shrubs along arroyo +, + +R. Moran +27383 + +, + +26 May 1979 + +( + +RSA +292842 + +!, + +SD +103648 + +!) + +; • + +Sierra Juarez +, + +1.5 km +NW of + +El Mezquite +, +Baja California +, + +32.35833 +, +- 116.0667 + +, + +1400 m + +elevation, +semishade of piñones, piñon-juniper-pine wood, associates: piñon-juniper-pine wood +, + +R. Moran +27437 + +, + +27 May 1979 + +( + +POM +369185 + +!, + +SD +103689 + +) + +; • + +Sierra Juarez +, +arroyo Agua Grande +, + +10 km +south of La Rumorosa + +, +Baja California +, + +32.4625 +, +- 116.0333 + +, + +1225 m + +elevation, +in semishade, among rocks +, + +R. Moran +30751 + +, + +16 May 1982 + +( + +SD +110999 + +!) + +; • + +San Salvador +: +Bridge over branch of Rio San Carlos near Rancho San Salvador +, +Baja California +, + +31.8333 +, +- 116.0667 + +, + +1000 m + +elevation, + +R. F. Thorne +60086 with +Dave Charlton + +, + +19 April 1985 + +( + +RSA +346809 + +!) + +; • + +Canon de Guadalupe +, +Baja California +, +32.15, - 115.8 +, + +450 m + +elevation, +dry slopes along canyon above resort area, sandy areas and rocky +, + +R. F. Thorne +61750 with +Steve Boyd + +, +etc. +, + +23 March 1986 + +( + +RSA +349247 + +!) + +; + + +1.5 mi +NW + +Village of Valle Trinidad +, +Baja California +, +31.423618 *, - 115.739987 * +, + +989 m + +elevation *, +granitic hillside +, + +I. L. Wiggins +16061 A + +, + +3 April 1960 + +( + +DS +506948 + +!) + +; + + + +United States +• +New York Mtns +, +[above Brant Siding] in canyon above old mine site +, +California +, +San Bernardino Co. +, + +35.27667 +, +- 115.34556 + +, + +1364 m + +elevation, +granite, talus, gravel +, + +J. M. André +4153 with +G. L. Clifton + +, + +7 May 2003 + +( + +GMDRC +1194 + +!, + +UCR +-164170 + +!) + +; • + +New York Mountains +: +growing in bottom of main drainage, just above main parking area / camp. Caruthers Canyon +, +Mojave National Preserve +, +California +, +San Bernardino Co. +, + +35.224817 +, +- 115.303467 + +, + +1675 m + +elevation, + +at base of boulders in creek bottom, granite, alluvium, gravel, associates: + +Salix exigua + +, + +Baccharis sergiloides + +, + +Carex alma + +, + +Ericameria cuneata + +, + +Pinus monophylla + + +, + +J. M. André +9554 with +T. La Doux + +, + +G. L. Clifton + +, + +5 May 2008 + +( + +GMDRC +2793 + +!, + +SDSU +18622 + +!) + +; • + +Joshua Tree National Park +, +Little San Bernardino Mountains +: + +Long Canyon, up side canyon approx. +400 m +. west of main canyon drainage + +, +California +, +Riverside Co. +, + +34.063517 +, +- 116.443033 + +, + +1172 m + +elevation, + +south-facing slope, decomposed granite, alluvium, associates: + +Juniperus californica + +, + +Ephedra nevadensis + +, + +Coleogyne ramosissima + +, + +Mentzelia involucrata + +, + +Calycoseris parryi + +, + +Eriophyllum confertiflorum + + +, + +J. M. André +21088 with +T. La Doux + +, + +R. B. Kelley + +, + +4 May 2011 + +( + +GMDRC +5506 + +!, + +GMDRC +5507 + +!, + +UCR +0005646 + +!) + +; • + +Hualapai Mountains +: +along dry creekbed at Moss Wash trailhead, just below Wild Cow Campground, 4.0 mi. south of Hualapai Mtn Ranger Sta. +, +Arizona +, +Mohave Co. +, + +35.064183 +, +- 113.8672 + +, + +1836 m + +elevation, + +yellow pine - oak woodland, granite, alluvium, gravel, associates: + +Pinus ponderosa + +, + +Quercus turbinella + +, + +Q. gambelii + +, + +Q. chysolepis + +, + +Ptelea trilobata + +, + +Symphoricarpos rotundifolius + +, + +Prunus virginiana + +, + +Eriogonum davidsonii + + +, + +J. M. André +30870 + +, + +29 May 2014 + +( + +ASU +0307471 + +!, + +GMDRC +6806 + +!) + +; • + +Cerbat Mountains +: + +west side of range c. +12 mi +north of Kingman, +0.3 mi +along faint dirt road north of Mineral Park Road, +2.7 mi +east of Hwy 93 + +, +Arizona +, +Mohave Co. +, + +35.35557 +, +- 114.17613 + +, + +1166 m + +elevation, + +upper alluvial fan, mixed, alluvium, gravel, associates: + +Juniperus californica + +, + +Coleogyne ramosissima + +, + +Larrea tridentata + +, + +Salvia dorrii + +, + +Acamptopappus sphaerocephala + +, + +Krameria erecta + +, + +Camissoniopsis pallida + + +, + +J. M. André +32653 + +, + +24 April 2015 + +( + +GMDRC +7523 + +!, + +UCR +-275600 + +!) + +; • + +Granite Mountains +; +UC GM Desert Research Center +; + +upper Granite Cove in wash +0.1 mile +south of Granite Cove Spring, +0.2 mile +north of Staples cabin + +, +California +, +San Bernardino Co. +, + +34.786 +, +- 115.65863 + +, + +1338 m + +elevation, + +under granite boulder, sand, associates: + +Acacia greggii + +, + +Ambrosia eriocentra + +, + +Prunus fasciculata + +, + +Eriophyllum wallacei + +, + +Plagiobothrys arizonicus + +, + +Descurainia pinnata + + +, + +J. M. André +33209 + +, + +4 April 2015 + +( + +GMDRC +7681 + +!, + +RSA +0087609 + +!, + +UCR +0004260 + +!) + +; • + +Peninsular Ranges +: + +along north side of Hwy 74 approx +1 mile +east of Pinyon Flats, south facing slopes of Sugarloaf Mtn + +, +California +, +Riverside Co. +, + +33.583617 +, +- 116.437467 + +, + +1210 m + +elevation, + +granitic gravelly soils among boulders, granite, alluvium, gravel, associates: + +Juniperus californica + +, + +Adenostoma sparsifolium + +, + +Arctostaphylos glauca + +, + +Cercocarpus betuloides + +, + +Mentzelia veatchiana + +, + +Phacelia fremontii + +, + +Saltugilia latimeri + + +, + +J. M. André +41103 + +, + +5 April 2019 + +( + +GMDRC +11665 + +!) + +; • + +Peninsular Ranges +: + +southeastern end of San Jacinto Mountains, along Hwy 74, +12.6 miles +SW of Palm Desert (Hwy 111), +1.2 miles +NNE of Sugarloaf Mtn + +, +California +, +Riverside Co. +, + +33.603767 +, +- 116.4199 + +, + +1081 m + +elevation, + +granitic boulders, granite, alluvium, gravel, associates: + +Pinus monophylla + +, + +Rhus ovata + +, + +Ericameria linearifolia + +, + +Encelia actoni + +, + +Phacelia distans + +, + +Chaenactis fremontii + +, + +Acmispon argophyllus + +, + +Cryptantha lepida + + +, + +J. M. André +42565 + +, + +14 April 2020 + +( + +GMDRC +12668 + +!) + +; • + +Peninsular Ranges +; +Anza Bench region +: +Hills north of Warner Springs, Indian Flats Campground of Cleveland National Forest along Road 9 S 05 +, +California +, +San Diego Co. +, + +33.349 +, +- 116.661 + +, + +1097 m + +elevation, + +slopes and shallow draws bordered by + +Quercus agrifolia + +, thin soils over granitic bedrock with extensive boulder outcrops and on sandy benches, associates: + +Adenostoma sparsifolium + +, + +Ceanothus greggii + +, + +C. perplexans + +, + +Adenostoma fasciculatum + +, + +Gutierrezia sarothrae + +, + +Muhlenbergia rigens + +, + +Lonicera subspicata + +, + +Rosa californica + +and + +Rhus trilobata + + +, + +S. Boyd +11034 with +LeRoy Gross + +, + +4 May 2004 + +( + +RSA +726843 + +!, + +UCR +0004357 + +) + +; • + +Kernville +, +California +, +Kern Co. +, +35.755333 *, - 118.425998 * +, + +812 m + +elevation *, + +T. S. Brandegee +s. n. + +, + +13 May 1892 + +( + +UC +79312 + +!) + +; + + +Canon wash, +1 mi +w of Morengo + +, +California +, +San Bernardino Co. +, +34.047071 *, - 116.621009 * +, + +898 m + +elevation *, + +J. R. Bruff +99 a + +, + +2 April 1928 + +( + +RSA +699336 + +!) + +; • + +Local landmark: +Cerbat Mountains +, +Chloride Quad +, +Arizona +, +Mohave Co. +, + +35.464472 +, +- 114.179333 + +, + +1584 m + +elevation, +Rocky Hillside; Slope Aspect: North, Slope Position: Lower Third; Vertical Slope Shape: Smooth; Horizontal Slope Shape: Smooth +, + +G. L. Clifton +14473 + +, + +28 April 1986 + +( + +PUA +32315 + +(Card # 64080)!) + +; • + + +Whitewater Canyon Rd. (exit from Interstate- 10 ca. +2.5 mi. +W of jct I- 10 with State 62); near start of Angeles [Pacific] Crest Trail + +, +California +, +Riverside Co. +, +33.99178 *, - 116.663304 * +, + +610 m + +elevation, + +riparian woodland (through creosote bush scrub) along rocky sandy stream channel joining the main Whitewater wash, rocky, sandy, associates: + +Encelia farinosa + +, + +Baccharis + +, + +Quercus + + +, + +C. Davidson +5578 with +B. Gustafson + +, + +7 April 1977 + +( + +RSA +499703 + +!) + +; • + +Near Dos Palmas Spring +, +San Jacinto Mts +, +California +, +Riverside Co. +, +33.61841 *, - 116.424376 * +, + +914 m + +elevation, + +H. & M. Dearing +4851 + +, + +29 March 1942 + +( + +SBBG +6266 + +!) + +; • + +Rincon Mountains +, +Chimenea Canyon above Madrona Ranger Station +, +Arizona +, +Pima Co. +, +32.183772 *, - 110.593461 * +, + +1219 m + +elevation, + +riparian forest in desert scrub, associates: + +Quercus + +, + +Juglans + +, + +Fraxinus + +, + +Baccharis + +, + +Ambrosia + + +, + +M. Fishbein +2013 + +, + +15 April 1994 + +( + +ARIZ +315946 + +!) + +; • + +South fork of San Joaquin river +, +California +, +Madera Co. +, +37.437614 *, - 119.239026 * +, + +2621 m + +elevation, + +H. M. Hall +650 B with +H. P. Chandler + +, + +July 1900 + +( + +UC +79530 + +!) + +; + +0.5 west of Peg Leg monument along Henderson Canyon Road +, +Anza Borrego Desert State Park +, +California +, +San Diego Co. +, + +33.3004 +, +- 116.314 + +, + +180 m + +elevation *, + +sandy wash, sandy wash, overlain with dry silty ash from 2003 fire above Coyote Canyon, associates: + +Larrea tridentata + +, + +Atriplex lentiformis + +, + +Phacelia brachyloba + +, + +Melilotus indica + +, + +Palafoxia arida var. arida + +, + +Geraea canescens + +, + +Abronia villosa var. villosa + +, + +Antirrhynum coulterianum + +, + +Calycoseris parryi + +, + +Chaenactis artemisiifolia + + +, + +L. Hendrickson +53 with +L. Louise Jee + +, + +9 April 2004 + +( + +SD +161807 + +!) + +; • + +San Felipe Valley +. +Cigarettes Hills +. +San Felipe Wildlife Area +, + +California Department of Fish & Game property. 1.8 air miles W of SW intersection of State Hwy. 78 and County Rd S- 2 and. +7 mile +N of hwy 78, on south side of prominent hill (2765) + +, +California +, +San Diego Co. +, + +33.0963 +, +- 116.5064 + +, + +769 m + +elevation, + +decomposing pegmatite dike, coarse gravel and sandy soil, associates: + +Agave deserti + +, + +Juniperus californica + +, + +Prunus fremontii + +, + +Ziziphus parryi + +, + +Ericameria brachylepsis + +, + +Eriogonum fasciculatum var polifolium + + +, + +L. Hendrickson +3928 + +, + +10 May 2009 + +( + +BSCA +1213 + +, + +SD +214894 + +!) + +; + + +5 mi. +se. of Nevada-Arizona border along Lime Kiln Canyon BLM rd. 242 + +, +Arizona +, +Mohave Co. +, +36.680548 *, - 114.009796 * +, + +1000 m + +elevation, + +mixed desert shrub community, gravelly limestone soils, associates: + +Quercus + +, + +Juniperus + + +, + +L. C. Higgins +25212 + +, + +6 April 2004 + +( + +BRYV +0155702 + +, + +HSC +202913 + +!, + +KHD +00031415 + +) + +; • + +Lime Kiln Canyon +, +Virgin Mountains +, +Arizona +, +Mohave Co. +, + +36.646315 +, +- 114.019191 + +, + +1250 m + +elevation, +Pinyon, juniper, oak community, limestone soil +., + +L. C. Higgins +25402 with +G. Green + +, + +4 May 2004 + +( +BRYV 0155701 +, + +RENO +81684 + +!, + +UNM +0026372 + +) + +; • + +Arizona +, +Skull valley +, Arizona, +Yavapai Co. +, +34.504684 *, - 112.687251 * +, + +1311 m + +elevation, + +M. E. Jones +7028 + +, + +28 April 1903 + +( + +UC +78720 + +!) + +; • + +Arizona +, +Skull valley +, Arizona, +Yavapai Co. +, +34.504684 *, - 112.687251 * +, + +1311 m + +elevation, + +M. E. Jones +s. n. + +, + +1 May 1903 + +( + +POM +71271 + +!) + +; • + +Arizona +, +Skull valley +, Arizona, +Yavapai Co. +, +34.504684 *, - 112.687251 * +, + +1311 m + +elevation, + +M. E. Jones +s. n. + +, + +1 May 1903 + +( + +POM +71274 + +!) + +; • + +Pinal Mts. above road to Madera Peak on road to Signal Peak and Pinal Peak +, +Arizona +, +Gila Co. +, +33.2822 *, - 110.821 * +, + +2384 m + +elevation *, +upper south facing slopes +, + +D. Keil +4628 + +, + +12 May 1969 + +( + +ASU +69306 + +!) + +; • + +Little San Bernardino Mountains +, +ridge along Upper East Deception Canyon, Joshua Tree National Park +, +California +, +Riverside Co. +, + +33.97936 +, +- 116.30985 + +, + +1319 m + +elevation, + +associates: + +Yucca schidigera + +, + +Juniperus californica + +, + +Eriogonum wrightii nodusum + +, + +Peucephyllum schotii + + +, + +T. LaDoux +3724 with +E. Babich + +, + +N. Pietrasiak + +, + +21 April 2008 + +( + +JOTR +00914 + +!) + +; + + +2 miles +east of Banner on San Felipe Wash + +, +California +, +San Diego Co. +, +33.083288 *, - 116.516252 * +, + +773 m + +elevation, + +K. McCully +92 + +, + +17 May 1925 + +( + +POM +97231 + +!) + +; • + +Traverse Ranges +; +San Bernardino Mountain region +: +Whitewater Canyon +; +Along Whitewater River drainage and adjacent slopes below Whitewater Visitor Center (Whitewater Trout Farm) in the vicinity of Bonnie Bell +, +California +, +Riverside Co. +, + +33.95477 +, +- 116.64248 + +, + +525 m + +elevation, +riparian woodland along water course to desert sage scrub on adjacent flats and slopes +, + +O. Mistretta +4627 with +Duncan Bell + +, + +Jill Beckman + +, + +Joy England + +, + +Katie Kane + +, + +Jamie Hall + +, + +Chris McDonald + +, + +Lucila Reccia + +, + +Lonnie Rodriguez + +, + +April Sall + +, + +Tracy Tennant + +, + +12 April 2010 + +( + +RSA +772061 + +!) + +; • + +Traverse Ranges +; +San Bernardino Mountain region +: +Whitewater Canyon +; +Along Whitewater River drainage and adjacent slopes below Whitewater Visitor Center (Whitewater Trout Farm) +, +California +, +Riverside Co. +, + +33.97538 +, +- 116.65119 + +, + +600 m + +elevation, +riparian woodland along water course to desert sage scrub on adjacent flats and slopes +, + +O. Mistretta +4702 a with +Duncan Bell + +, + +Jill Beckman + +, + +Joy England + +, + +Katie Kane + +, + +Jamie Hall + +, + +Chris McDonald + +, + +Lucila Reccia + +, + +Lonnie Rodriguez + +, + +April Sall + +, + +Tracy Tennant + +, + +12 April 2010 + +( + +RSA +771786 + +!) + +; • + + +Northwest slope of Santa + +Rosa +Mtns. + + +, +California +, +Riverside Co. +, +33.555223 *, - 116.487153 * +, + +2134 m + +elevation, +dry banks, northwest slope +, + +P. A. Munz +15086 + +, + +30 May 1937 + +( + +BRYV +0155683 + +, + +RSA +0079225 + +, + +UC +662529 + +!) + +; • + +Pines to Palms Highway # 74 +, + +15 miles +west of + +Palm Desert +, +California +, +Riverside Co. +, +33.581968 *, - 116.459557 * +, + +1219 m + +elevation, + +D. Myrick +833 + +, + +30 April 1964 + +( + +SBBG +28428 + +!) + +; • + +Colorado Desert +, +California +, +San Diego Co. +, +33.05 *, - 116.12 * +, m elevation, + +C. R. Orcutt +s. n. + +, + +April 1889 + +( + +UC +280510 + +!) + +; + +Walker Canyon Ecological Preserve +: +between Boulevard and Jacumba on the north side of Interstate 8, along arroyo bottom near stream +, +California +, +San Diego Co. +, + +32.66306 +, +- 116.22333 + +, + +915 m + +elevation, + +Chaparral / desert transition, granitic substrates, associates: + +Quercus cornelius mulleri + +, + +Adenostoma sparsifolium + +, + +Rhus ovata + +, + +Juniperus californica + +, + +Cylindropuntia ganderi + + +, + +J. P. Rebman +8530 with +Jeannie Gregory + +, + +17 April 2003 + +( + +SD +159509 + +, + +UC +1787644 + +!, + +UCR +0004378 + +) + +; • + +San Jacinto Mountains +, +along the trail toward Spitler Pk from the Hurkey Crk camp to Bonita Vista Rd +, +California +, +Riverside Co. +, +33.6961 *, - 116.6392 * +, + +1707 m + +elevation, +along trail; chaparral & some yellow pine forest +, + +A. C. Sanders +6537 + +, + +1 June 1986 + +( + +DES +00030658 + +, + +OBI +131801 + +, + +SBBG +87639 + +!, + +UCR +0004196 + +, + +UNLV +23020 + +) + +; • + +Along FS road 22 S 82, east base of the Needles +, + +approx. +3 miles +from + +Pyles Camp. +, +California +, +Tulare Co. +, +36.10939 *, - 118.484532 * +, + +2419 m + +elevation, +granitic sandy soils +, + +J. R. Shevock +5114 + +, + +18 June 1976 + +( + +LOB +100364 + +!) + +; • + +SW Imperial Co, near SE boundary of San Diego Co. +, +In-Ko-Pah Mountains, access road to Valley of the Moon +, +California +, +Imperial Co. +, +32.639288 *, - 116.100194 * +, + +1031 m + +elevation, + +transition area from scrub oak-yucca to diverse high desert scrub, steep N-facing bouldery drainage, rocky slopes, steep N-facing bouldery drainage; granitic outcrops and soils, mostly coarse particle size;, associates: + +C. pterocarya var. cycloptera + +, + +C. p. var. purpusii + +, + +C. clevelandii + +, + +C. intermedia + +, + +Microsersis douglasii + +, + +Gilia caruifolia + +, + +Phacelia distans + +, + +Lotus scoparius + +, + +Quercus cornelius-muelleri + +, + +Yucca schidigera + +, + +Nolina parryi + +, + +Rhus ovata + +, + +Simmondsia chinensis + +, + +Acacia greggii + +, + +Encelia actoni + +, + +Salvia apiana + +, + +Cylindropuntia ganderi + +, + +C. wolfii + +, + +Ericameria brachylepsis + +, and + +Lupinus excubitus var. medius + + +, + +D. Silverman +3964 with +Jon Rebman + +, + +23 April 2001 + +( + +JEPS +103093 + +!, + +SBBG +116573 + +!) + +; • + +Wash adjacent to and just west of jeep road +, +Rodriguez Canyon +, +ca. 2.0 miles southwest of Granite Mtn. peak. +, +California +, +San Diego Co. +, + +33.03111 +, +- 116.50361 + +, + +946 m + +elevation, + +wash adjacent to exposed, sparse desert scrub, sandy, gravelly soil, associates: + +Prunus fremontii + +, + +Rhus ovata + +, + +Baccharis sergiloides + +, + +Quercus +sp. + + +, + +M. G. Simpson +2790 with +K. Hasenstab + +, + +M. Silveira + +, + +and +L. Simpson + +, + +7 April 2007 + +( + +SDSU +19296 + +!) + +; • + +Wash adjacent to and just west of jeep road +, +Rodriguez Canyon +, +ca. 2.0 miles southwest of Granite Mtn. peak. +, +California +, +San Diego Co. +, + +33.03111 +, +- 116.50361 + +, + +946 m + +elevation, + +wash adjacent to exposed, sparse desert scrub, sandy, gravelly soil, associates: + +Prunus fremontii + +, + +Rhus ovata + +, + +Baccharis sergiloides + +, + +Quercus +sp. + + +, + +M. G. Simpson +2794 with +K. Hasenstab + +, + +M. Silveira + +, + +and +L. Simpson + +, + +7 April 2007 + +( + +SDSU +17573 + +!) + +; • + +Pacific Coast Trail +, + +ca. +0.1 mile +north of parking area at Hwy 74. + +, +California +, +Riverside Co. +, + +33.56416 +, +- 116.57673 + +, + +1499 m + +elevation, + +beneath + +Adenostoma fasciculatum + +. Adjacent to open + +Adenostoma sparsifolium + +- + +Adenostoma fasciculatum + +- + +Ceanothus greggii + +scrub, coarse sand, associates: + +Adenostoma sparsifolium + +, + +Adenostoma fasciculatum + +, + +Ceanothus greggii + + +, + +M. G. Simpson +3080 with +Lori Simpson + +, + +26 April 2009 + +( + +SDSU +21885 + +!) + +; • + +Pacific Coast Trail +, + +ca. +0.6 mile +north of parking area at Hwy 74. + +, +California +, +Riverside Co. +, + +33.56977 +, +- 116.57572 + +, + +1526 m + +elevation, + +side of trail. Open + +Adenostoma sparsifolium + +- + +A. fasciculatum + +- + +Ceanothus greggii + +scrub., coarse sand, associates: + +Adenostoma sparsifolium + +, + +Adenostoma fasciculatum + +, + +Ceanothus greggii + + +, + +M. G. Simpson +3085 with +Lori Simpson + +, + +26 April 2009 + +( + +SDSU +21647 + +!) + +; • + +Anza Borrego Desert State Park +. +Wash of Bitter Creek Canyon, ca. 1.0 mile north of Grapevine Mountain peak. +, +California +, +San Diego Co. +, + +33.13444 +, +- 116.46798 + +, + +739 m + +elevation, + +along base of canyon wall rocks of wash, open scrub of desert wash, gravelly sand, associates: + +Agave deserti + +, + +Chenopodium +sp. + +, + +Encelia farinosa + +, + +Opuntia basilaris + +, + +Senegalia greggii + + +, + +M. G. Simpson +3790 with +Amanda Everett + +, + +Andy Siekkinen + +, + +Lee Ripma + +, + +Matt Newcomb + +, + +15 February 2014 + +( + +SDSU +20519 + +!) + +; • + +Anza Borrego Desert State Park +, + +Cool Canyon, ca. +1,000 m +west-southwest of parking area at end of dirt road, ca. +1.8 km +east-southeast of Granite Mountain peak. + +, +California +, +San Diego Co. +, + +33.04362 +, +- 116.44427 + +, + +869 m + +elevation, + +desert succulent scrub, brown, sandy loam, associates: + +Juniperus californica + +, + +Cylindropuntia ganderi + +, + +Prunus fremontii + +, + +Encelia farinosa + +, + +Bahiopsis parishii + + +, + +M. G. Simpson +3795 with +SDSU Taxonomy of California Plants Class + +, + +29 March 2014 + +( + +SDSU +23021 + +!) + +; • + +Anza Borrego Desert State Park +, +Bitter Creek Canyon, ca. 2.3 kilometers southwest of Grapevine Canyon Road (dirt). +, +California +, +San Diego Co. +, + +33.13402 +, +- 116.47022 + +, + +745 m + +elevation, + +wash of canyon, desert succulent scrub, silty, coarse sand, associates: + +Prosopis glandulosa + +, + +Encelia farinosa + +, + +Chenopodium murale + +, + +Bromus rubens + + +, + +M. G. Simpson +3800 with +Makenzie E. Mabry + +, + +3 April 2014 + +( + +SDSU +23020 + +!) + +; • + +Anza Borrego Desert State Park +, +Culp Valley, along hiking trail at camping area. +, +California +, +San Diego Co. +, + +33.22376 +, +- 116.45312 + +, + +1033 m + +elevation, + +flat, coarse sand / fine gravel, organic material beneath, associates: + +Rhus ovata + +, + +Ziziphus parryi var. parryi + +, + +Cylindropuntia ganderi + +scrub + +, + +M. G. Simpson +3856 with +Makenzie Mabry +& +Andy Siekkinen + +, + +1 April 2015 + +( + +SDSU +23271 + +!) + +; • + +Culp Valley +, + +ca. +0.1 mile +along dirt road, to parking lot at left, then 10 meters west of northern end of lot. + +, +California +, +San Diego Co. +, + +33.22209 +, +- 116.45791 + +, + +1014 m + +elevation, + +desert transtion vegetation, brownish, gravelly sand, growing adjacent to boulder, associates: + +Prunus fremontii + +, + +Ziziphus parryi var. parryi + + +, + +M. G. Simpson +4186 with +Jepson 2019 workshop members + +, + +14 April 2019 + +( + +SDSU +22965 + +!) + +; • + +San Gabriel Mountains +, + +east side of Cabin Canyon, c. +200 m +north of canyon mouth, west of Aliso Cyn. Rd., +0.5 mi +west of Angeles Forest Hwy + +, +California +, +Los Angeles Co. +, + +34.4169444 +, +- 118.0925 + +, + +1183 m + +elevation, + +shallow canyon bottom with flowing stream; chaparral dominant, associates: +Ceanothus leu +., +Adenostoma f +., +Quercus wis +., +Prunus ili +., +Eriodictyon tri +., with +Layia gla +., +Phacelia fre +., +Calyptridium mon +., +Penstemon cen +., +Malacothrix gla +., +Camissoniopsis hir +., +Bromus tec +., +Antirrhinum cou +., +Cirsium occ +., +Chaenactis gla +., etc + +, + +R. G. Swinney +14663 + +, + +12 May 2011 + +( + +RSA +0091168 + +!, + +SDSU +22957 + +!) + +; • + +San Gabriel Mountains +, + +0.3–0.5 miles +south of Aliso Canyon Road, +2 miles +west of Angeles Forest Hwy., nameless cyn. To the east of Beartrap Cyn. + +, +California +, +Los Angeles Co. +, + +34.42778 +, +- 118.11639 + +, + +1066 m + +elevation, + +Station Fire burn of 2009, shallow ravine and adj. slopes, chaparral, associates: +Adenostoma f +., +Prunus i +., +Lonicera su +., +Rhus ar +., +Cercocarpus b +., also w / +Opuntia ba +., +Malacothamnus fr +., +Lupinus bi +., +Camissonia s. +, + +Uropappus + +, +Mentzelia co. +, +Delphinium pat. +, +Phacelia fr. +, +Leptosiphon a +., etc + +, + +R. G. Swinney +14845 + +, + +26 May 2011 + +( + +RSA +0092147 + +!, + +SDSU +22955 + +!) + +; • + +San Gabriel Mountains +: +Colby Ranch (Camp Colby), S end of Inspiration Canyon tr., in a N-flowing tributary of upper Coldwater Cyn. +, +California +, +Los Angeles Co. +, + +34.29861 +, +- 118.11361 + +, + +1226 m + +elevation, + +Station Fire burn of ‘ 09, canyon wih slopes; chaparral, coarse granite and fine brown loam, associates: + +Ceanothus + +, + +Eriodictyon + +, + +Quercus + +, + +Pinus + +, + +Nicotiana + +, + +Eschscholzia + +, + +Phacelia + +, + +Adenostoma + +, + +Mimulus + +, + +Chaenactis + +, + +Keckiella + +, + +Calyptridium + +, + +Salvia + +, + +Galium + +, + +Penstemon + +, + +Argemone +, +Erhendorferia + +, + +Dendromecon + +, + +Eriogonum + +, + +Mentzelia + +, + +Solanum + + +, + +R. G. Swinney +15440 + +, + +7 July 2011 + +( + +RSA +0091783 + +!, + +SDSU +22954 + +!, + +UCR +0004166 + +!) + +; • + +San Gabriel Mountains +: +Colby Ranch (Camp Colby), S end of Inspiration Canyon tr., in a N-flowing tributary of upper Coldwater Cyn. +, +California +, +Los Angeles Co. +, + +34.29861 +, +- 118.11361 + +, + +1226 m + +elevation, + +Station Fire burn of ‘ 09, canyon wih slopes; chaparral, coarse granite and fine brown loam, associates: + +Ceanothus + +, + +Eriodictyon + +, + +Quercus + +, + +Pinus + +, + +Nicotiana + +, + +Eschscholzia + +, + +Phacelia + +, + +Adenostoma + +, + +Mimulus + +, + +Chaenactis + +, + +Keckiella + +, + +Calyptridium + +, + +Salvia + +, + +Galium + +, + +Penstemon + +, + +Argemone +, +Erhendorferia + +, + +Dendromecon + +, + +Eriogonum + +, + +Mentzelia + +, + +Solanum + + +, + +R. G. Swinney +15442 + +, + +7 July 2011 + +( + +RSA +0091784 + +, + +SDSU +22946 + +!, + +UCR +0039547 + +) + +; • + +San Gabriel Mountains +: +Strawberry Peak summit (Chilao Flat) +., +California +, +Los Angeles Co. +, + +34.28389 +, +- 118.12028 + +, + +1869 m + +elevation, + +Station Fire burn of 2009, entire summit burned; resprouting chaparral, associates: + +Eriodictyon trichocalyx + +, + +E. parryi + +, + +Quercus wislizeni + +, + +Q. chrysolepis + +, + +Penstemon centranthifolius + +, + +Eriastrum densifolium + +, + +Ericameria nausiosa + +, + +Camissoniopsis hirtella + +, + +Bromus tectorum + +, + +B. carinatus + +, + +Eriophyllum confertiflorum + +, + +Cryptantha muricata + +, + +Garrya flavescens + +, + +Tauschia parishii + +, + +Eriogonum saxatile + +, + +Orobanche fasciculata + +, + +Sidotheca trilobata + +, etc + +, + +R. G. Swinney +15527 + +, + +21 July 2011 + +( + +RSA +0089514 + +, + +SDSU +22959 + +!, + +UCR +0004165 + +) + +; • + +San Gabriel Mts. +, + +n. e. boundary ridge top of San Dimas Experimental Forest, c. +0.14 miles +s. w. of summit at far s. e. end of ridge, 1.7 air miles w. of Cow Cyn. Saddle at Glendora Ridge Rd. to 1.8 land miles ESE along SDEF boundary ridge to ridge top at +5595 ft. +( +1706 m +.), and east via level ridge connection to ridge top at +5560 ft. +, 04 air miles NNE of Sunset Pk. summit. This is the highest elev. within SDEF, Mt. Baldy Quad + +, +California +, +Los Angeles Co. +, + +34.209722 +, +- 117.695278 + +, + +1695 m + +elevation, + +Ridge top, chaparral, Big Cone Spruce / + +Quercus + +woodland, associates: + +Pseudotsuga macrocarpa + +, + +Quercus wislizeni subsp. frutescens + +, + +Q. chrysolepis + +, + +Ceanothus crassifolius + +, + +Eriogonum fasciculatum subsp. foliolosum + +, + +Bromus tectorum + + +, + +R. G. Swinney +18319 + +, + +4 May 2016 + +( + +SDSU +23973 + +!) + +; • + +North of Azusa, WCA’s Azusa Foothills Open Space +, +ridgetop overlooking mouth of San Gabriel Cyn., within 50 meters of Vasquez Ranch Flagpole, last burned 2014 - Cobly Fire, 0.55 air miles n. of jct. of Sierra Madre Ave. and Hilltop Dr., Azusa Quad 7.5 ’ +, +California +, +Los Angeles Co. +, + +34.156355 +, +- 117.904528 + +, + +450 m + +elevation, + +ridge top, coastal sage scrub and chaparral surrounding previously slightly-disturbed area, granitic-loam, associates: + +Malosma laurina + +, + +Bromus diandrus + +, + +Encelia farinosa + +, + +Artemisia californica + +, + +Hirschfeldia incana + +, + +Malacothrix saxatilis var. tenuifolia + + +, + +R. G. Swinney +24723 A + +, + +5 May 2021 + +( + +SDSU +25595 + +!, + +UCR +0164808 + +) + +; • + +San Bernardino Mountains +: +unnamed ridgetop (7,582 ’) and adjacent slope just W of Heart Bar Peak and SE of Cienaga Seca Creek +., +California +, +San Bernardino Co. +, + +34.1667 +, +- 116.775 + +, + +2222 m + +elevation, + +ridgetop recently bulldozed for firebreak; adjacent N and W slopes w / + +Quercus chrysolepis + +and + +Pinus jeffreyi + +, etc., + +Quercus chrysolepis + +and + +Pinus jeffreyi + +, etc. + +, + +S. D. White +13324 with +Marissa Caringella + +, + +24 July 2009 + +( + +RSA +752962 + +!, + +UC +1949977 + +, + +UCR +0004334 + +, + +UCSB +029502 + +) + +; • + +Peninsular Ranges +: + +Santa + +Rosa +Mountains + + +, +California +, +Riverside Co. +, +33.539 *, - 116.457 * +, + +2310 m + +elevation, + +R. S. Woglum +3072 + +, + +12 June 1941 + +( + +RSA +611564 + +!) + +; • + +San Bernardino Mountains +: +Unnamed canyon and ridgeline immediately SW of Cienega Seca (Blue Sky Meadow) +, +California +, +San Bernardino Co. +, + +34.1833 +, +- 116.725 + +, + +2476 m + +elevation, +yellow pine forest with somes areas of pinyon / juniper woodland +, + +J. M. Wood +948 with +Dustin Ray + +, + +18 June 2009 + +( + +RSA +752329 + +!) + +; • + +San Bernardino Mtns. +, +Santa Ana River, unnamed canyon and ridgeline E of Wildhorse Canyon, N of Hwy 38 +, +California +, +San Bernardino Co. +, +34.17083, - 116.8 +, + +2100 m + +elevation, +dry canyon wash, riparian woodland and adjacent uplands +, + +J. M. Wood +1013 with +Dustin Ray + +, + +30 June 2009 + +( + +RSA +750733 + +!) + +; • + +Borrego Palm Canyon +, +California +, +San Diego Co. +, +33.276836 *, - 116.426169 * +, + +322 m + +elevation, + +F. Youngberg +103 + +, + +11 April 1935 + +( + +POM +209514 + +!) + +. + + + + \ No newline at end of file