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- - -Antarctic Kinorhyncha: Seven new species from the Antarctic Peninsula + + + +Antarctic Kinorhyncha: Seven new species from the Antarctic Peninsula - - -Author + + +Author -Sørensen, Martin V. -4143D650-12FC-4914-93F5-2C39339A7156 -Natural History Museum of Denmark, University of Copenhagen, 2100 Copenhagen, Denmark. -mvsorensen@snm.ku.dk +Sørensen, Martin V. +4143D650-12FC-4914-93F5-2C39339A7156 +Natural History Museum of Denmark, University of Copenhagen, 2100 Copenhagen, Denmark. +mvsorensen@snm.ku.dk - - -Author + + +Author -Macheriotou, Lara -1037C143-8562-41A3-8149-BC3DE4323845 -Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. -lara.macheriotou@ugent.be +Macheriotou, Lara +1037C143-8562-41A3-8149-BC3DE4323845 +Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. +lara.macheriotou@ugent.be - - -Author + + +Author -Braeckman, Ulrike -6C48F169-ECC8-4F98-A9CF-9B2A664DA903 -Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. & Ulrike. -Ulrike.Braeckman@ugent.be +Braeckman, Ulrike +6C48F169-ECC8-4F98-A9CF-9B2A664DA903 +Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. +Ulrike.Braeckman@ugent.be - - -Author + + +Author -Smith, Craig R. -F004E0AC-049A-48D6-9369-EBE18ECEF53C -Department of Oceanography, University of Hawai’i at Mānoa, Honolulu HI 96822, Hawaii, USA. -craigsmi@hawaii.edu +Smith, Craig R. +F004E0AC-049A-48D6-9369-EBE18ECEF53C +Department of Oceanography, University of Hawai’i at Mānoa, Honolulu HI 96822, Hawaii, USA. +craigsmi@hawaii.edu - - -Author + + +Author -Ingels, Jeroen -F6107F36-C67F-4CB8-A6C0-DFBF66D3C464 -Coastal and Marine Laboratory, Florida State University, St. Teresa FL 32346, Florida, USA. & National Institute for Water and Atmospheric Research, Wellington 6021, New Zealand. -jeroen.ingels@niwa.co.nz +Ingels, Jeroen +F6107F36-C67F-4CB8-A6C0-DFBF66D3C464 +Coastal and Marine Laboratory, Florida State University, St. Teresa FL 32346, Florida, USA. & National Institute for Water and Atmospheric Research, Wellington 6021, New Zealand. +jeroen.ingels@niwa.co.nz -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2025 - -2025-07-01 + +2025 + +2025-07-01 - -1000 + +1000 - -1 + +1 - -1 -102 + +1 +102 - -https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2947/13307 + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2947/13307 -journal article -10.5852/ejt.2025.1000.2947 -2118-9773 -15808384 -98563124-EFCC-4542-B5AB-E14C0C3978DD +journal article +10.5852/ejt.2025.1000.2947 +2118-9773 +15808384 +98563124-EFCC-4542-B5AB-E14C0C3978DD - + @@ -134,21 +134,19 @@ sublateral positions. Material examined - + + Holotype - - - ANTARCTICA • ♂ -(mounted for LM in -Fluoromount G -on HS slide); -Antarctic Peninsula -, CRS 1706; +(mounted for LM in Fluoromount G on HS slide); +Antarctic Peninsula +, +CRS 1706 +; 64°50.47′ S , 62°35.12′ W @@ -161,7 +159,11 @@ b.s.l. 1 Dec. 2015 -; FjordEco1; soft sediment; +; +FjordEco1 +; +soft sediment +; NHMD 1784249 . @@ -172,7 +174,7 @@ b.s.l. - + ANTARCTICA – @@ -180,11 +182,7 @@ b.s.l. • 1♀ -(mounted as -holotype -); -CRS 1769 -; +(mounted as holotype); CRS 1769; 64°52.37′ S , 62°25.27′ W @@ -197,17 +195,17 @@ b.s.l. 5 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; USNM 1740027 • - + 1 ♂ -(mounted as -holotype -); -CRS 1773 -; +(mounted as holotype); CRS 1773; 64°52.35′ S , 62°25.88′ W @@ -220,17 +218,17 @@ b.s.l. 6 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; NHMD 1784250 • - + 1 ♂ -(mounted as -holotype -); -CRS 1776 -; +(mounted as holotype); CRS 1776; 64°52.53′ S , 62°33.90′ W @@ -243,17 +241,17 @@ b.s.l. 7 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; USNM 1740028 • - + 3 ♀♀ -(mounted as -holotype -); -CRS 1793 -; +(mounted as holotype); CRS 1793; 64°39.53′ S , 62°55.03′ W @@ -266,7 +264,11 @@ b.s.l. 11 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; NHMD 1784251 , 1784284 @@ -274,15 +276,11 @@ b.s.l. 1784303 • - + 2 ♂♂ , 1 ♀ -(mounted as -holotype -); -CRS 1809 -; +(mounted as holotype); CRS 1809; 64°39.59′ S , 62°55.09′ W @@ -295,7 +293,11 @@ b.s.l. 15 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; NHMD 1784304 to 1784306 @@ -305,8 +307,8 @@ to Additional material - - + + ANTARCTICA – @@ -318,7 +320,7 @@ to MVS • - + 1 ♂ (mounted for SEM); CRS 1776; 64°52.53′ S @@ -333,10 +335,15 @@ b.s.l. 7 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; MVS • + 4 ♂♂ , 1 ♀ @@ -345,19 +352,20 @@ b.s.l. , 62°55.03′ W ; -701 m -b.s.l.; - - - + +701 m + +b.s.l.; 11 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; soft sediment; MVS • - + 3 ♂♂ , 1 ♀ @@ -374,11 +382,15 @@ b.s.l. 15 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; MVS • - + 4 ♂♂ , 5 ♀♀ @@ -395,7 +407,11 @@ b.s.l. 21 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; MVS . @@ -1173,7 +1189,7 @@ SEGMENT 11. With lateral terminal spines. Females with thin lateral terminal acc ). - + Distribution @@ -1185,9 +1201,11 @@ Antarctic Peninsula: Gerlache Strait and Andvord Bay MBA and IBB, b.s.l. See Fig. 1 for geographic overview of stations and -Table 1 +Table 1 for station and specimen information. + + Polymorphism @@ -1222,5 +1240,280 @@ USNM 1740027 from stn 1769) had a set of small acicular spines in ventromedial p in all samples – small acicular spines in lateral accessory positions on segment 7. The seven remaining specimens also appeared to follow the common pattern but had, due to their condition or mounting orientation, characters that could not be confirmed visually. + + + + +Diagnostic remarks on + +Polacanthoderes grzelakae + +sp. nov. + + + + +With the addition of + +P. grzelakae + +sp. nov. +, + +Polacanthoderes + +now accommodates three species that all are restricted to the Antarctic continent ( +Sørensen 2008a +; + +Yamasaki +et al. +2022 + +). The genus and its first described species, + +Polacanthoderes martinezi +Sørensen, 2008 + +, was described from the South +Shetland Islands +( +Sørensen 2008a +), and a phylogenetic analysis supported that + +Polacanthoderes + +represents a separate evolutionary linage within + +Echinoderidae ( +Sørensen 2008b +) + +. One of the characters that makes + +Polacanthoderes + +stand out from other echinoderids is the numerous small acicular spines in rather unusual positions, i.e., subdorsal, laterodorsal, midlateral, and ventromedial ( +Sørensen 2008a +). + + +Some years later, + +Yamasaki +et al. +(2022) + +described the second species of the genus, + +P. shiraseae + +. The species was described from Lützow-Holm Bay, Cape Damley, and near Totten Glacier, i.e., from Antarctic areas that geographically are pretty much opposite to the Antarctic Peninsula and South +Shetland Island +. Besides contributing with a new species, + +Yamasaki +et al. +(2022) + +also provided a redescription of + +P. martinezi + +and, in addition, shed light on the exceptionally high level of morphological variation within the two species (see following section for further discussion of polymorphism in species of + +Polacanthoderes + +). In a group like +Echinoderidae +, where much of the taxonomy traditionally has been based on the presence and position of spines and tubes, such variation can obviously lead to some taxonomic challenges. Yet, + +Yamasaki +et al. +(2022) + +explained how + +P. martinezi + +and + +P. shiraseae + +fairly easily could be distinguished by the conspicuously stronger lateroventral spines on segments 8 and +9 in + +P. shiraseae + +, and by the position of short acicular spines on segment 7, which appear in lateral accessory positions in + +P. martinezi + +and in sublateral positions in + +P. shiraseae + +. The latter character might seem like a very subtle alteration, open for subjective interpretation, but when observed the difference is in fact very distinct, and since both species otherwise have a series of short lateral accessory acicular spines on segments 6 to 9, it is easy to detect the sublateral displacement of the spines on segment +7 in + +P. shiraseae + +. + + +The new species, + +P. grzelakae + +sp. nov. +, very clearly shares most characters with + +P. shiraseae + +, including the strong lateroventral spines on segments 8 and 9, and the sublateral short acicular spines on segment 7. Thus, + +P. grzelakae + +is easily distinguished from + +P. martinezi + +. The characters that separate + +P. grzelakae + +from both congeners are the presence of subdorsal and ventrolateral tubes on segment 2. In the present study, a total of +38 specimens +of + +Polacanthoderes + +were examined. Out of these, +four specimens +had no tubes on segment 2 and were thus identified as + +P. shiraseae + +. Thirty other specimens had both subdorsal and ventrolateral tubes on segment 2, and were assigned to + +P. grzelakae + +. Of the remaining +four specimens +, three had only subdorsal tubes on segment 2, whereas a single specimen had only ventrolateral tubes. + + +In light of the known morphological variation within species of + +Polacanthoderes + +(and without access to molecular barcoding data), it is obviously not straightforward to determine whether + +P. grzelakae + +sp. nov. +is a distinct species or a morphological variation of + +P. shiraseae + +. However, a strong argument favours the first option. Recently, + +Anguas-Escalante +et al. +(2023) + +demonstrated with molecular barcoding that the presence or absence of tubes on segment +2 in +otherwise very similar species of + +Echinoderes + +actually should be seen as species diagnostic. In + +P. grzelakae + +the species diagnostic trait is exactly expressed as the presence of tubes on segment 2, and these tubes are consistently present within the +type +series of the species, whereas the morphological variation we observe is mostly expressed in the presence/absence of short acicular spines in the dorsal series on segments 4, 5, and 8 (see below for further discussion of this). + + + + +Based on these arguments, we propose + +P. grzelakae + +sp. nov. +as a new species showing close resemblance with + +P. shiraseae + +, but distinguished by the presence of subdorsal and ventrolateral tubes on segment 2. This proposal seems fair, but it is admittedly challenged by the four remaining specimens, + +Polacanthoderes +sp. 1 + +and +P. +sp. 2, that show variation in their tubes on segment 2. With tubes only in either subdorsal or ventrolateral positions on segment 2, the +four specimens +fall in between the two species, and the number of potential explanations are too numerous to allow a final conclusion. The two morphotypes could represent the bridging between + +P. shiraseae + +and + +P. grzelakae + +, suggesting that they are all conspecific; they could represent another two distinct species; or they could be results of hybridisation between + +P. shiraseae + +and + +P. grzelakae + +. It would require molecular barcoding to solve this question, and any other conclusion at this point would be nothing but speculation. However, based on the arguments above, we find it justified to consider + +P. grzelakae + +as a new, easily distinguishable species of + +Polacanthoderes + +. + + + + +The distribution pattern of glandular cell outlets +type +1 in + +P. grzelakae + +sp. nov. +follows the MD Seg. 1–3, PD 4–9 pattern (see table with summary of species with this pattern described up to +2020 in + +Sørensen +et al. +2020 + +). The same pattern is present in both + +Polacanthoderes + +congeners ( +Sørensen 2008a +; + +Yamasaki +et al. +2022 + +). + + \ No newline at end of file diff --git a/data/03/AE/40/03AE4066FFB6D45C1772FC84FEEAFE43.xml b/data/03/AE/40/03AE4066FFB6D45C1772FC84FEEAFE43.xml index 0a26c74e7fe..3d89d4bf567 100644 --- a/data/03/AE/40/03AE4066FFB6D45C1772FC84FEEAFE43.xml +++ b/data/03/AE/40/03AE4066FFB6D45C1772FC84FEEAFE43.xml @@ -1,83 +1,83 @@ - - - -Antarctic Kinorhyncha: Seven new species from the Antarctic Peninsula + + + +Antarctic Kinorhyncha: Seven new species from the Antarctic Peninsula - - -Author + + +Author -Sørensen, Martin V. -4143D650-12FC-4914-93F5-2C39339A7156 -Natural History Museum of Denmark, University of Copenhagen, 2100 Copenhagen, Denmark. -mvsorensen@snm.ku.dk +Sørensen, Martin V. +4143D650-12FC-4914-93F5-2C39339A7156 +Natural History Museum of Denmark, University of Copenhagen, 2100 Copenhagen, Denmark. +mvsorensen@snm.ku.dk - - -Author + + +Author -Macheriotou, Lara -1037C143-8562-41A3-8149-BC3DE4323845 -Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. -lara.macheriotou@ugent.be +Macheriotou, Lara +1037C143-8562-41A3-8149-BC3DE4323845 +Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. +lara.macheriotou@ugent.be - - -Author + + +Author -Braeckman, Ulrike -6C48F169-ECC8-4F98-A9CF-9B2A664DA903 -Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. & Ulrike. -Ulrike.Braeckman@ugent.be +Braeckman, Ulrike +6C48F169-ECC8-4F98-A9CF-9B2A664DA903 +Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. +Ulrike.Braeckman@ugent.be - - -Author + + +Author -Smith, Craig R. -F004E0AC-049A-48D6-9369-EBE18ECEF53C -Department of Oceanography, University of Hawai’i at Mānoa, Honolulu HI 96822, Hawaii, USA. -craigsmi@hawaii.edu +Smith, Craig R. +F004E0AC-049A-48D6-9369-EBE18ECEF53C +Department of Oceanography, University of Hawai’i at Mānoa, Honolulu HI 96822, Hawaii, USA. +craigsmi@hawaii.edu - - -Author + + +Author -Ingels, Jeroen -F6107F36-C67F-4CB8-A6C0-DFBF66D3C464 -Coastal and Marine Laboratory, Florida State University, St. Teresa FL 32346, Florida, USA. & National Institute for Water and Atmospheric Research, Wellington 6021, New Zealand. -jeroen.ingels@niwa.co.nz +Ingels, Jeroen +F6107F36-C67F-4CB8-A6C0-DFBF66D3C464 +Coastal and Marine Laboratory, Florida State University, St. Teresa FL 32346, Florida, USA. & National Institute for Water and Atmospheric Research, Wellington 6021, New Zealand. +jeroen.ingels@niwa.co.nz -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2025 - -2025-07-01 + +2025 + +2025-07-01 - -1000 + +1000 - -1 + +1 - -1 -102 + +1 +102 - -https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2947/13307 + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2947/13307 -journal article -10.5852/ejt.2025.1000.2947 -2118-9773 -15808384 -98563124-EFCC-4542-B5AB-E14C0C3978DD +journal article +10.5852/ejt.2025.1000.2947 +2118-9773 +15808384 +98563124-EFCC-4542-B5AB-E14C0C3978DD @@ -98,7 +98,7 @@ urn:lsid:zoobank.org:act: Figs 2–4 , -Tables 5–6 +Tables 5–6 @@ -138,20 +138,16 @@ and only the second member of the genus recorded from the Southern Hemisphere. Material examined - + + Holotype - - - ANTARCTICA • ♂ -(mounted for LM in -Fluoromount G -on HS slide); -Antarctic Peninsula +(mounted for LM in Fluoromount G on HS slide); +Antarctic Peninsula , CRS 1702; 64°51.15′ S , @@ -165,7 +161,11 @@ b.s.l. 30 Nov. 2015 -; FjordEco1; soft sediment; +; +FjordEco +1; +soft sediment +; NHMD 1784659 . @@ -174,17 +174,17 @@ b.s.l. Additional material - + ANTARCTICA – Antarctic Peninsula • -1 juv. +1 juv. (mounted for LM in -Fluoromount G -on HS slide); CRS 1706; +Fluoromount +G on HS slide); CRS 1706; 64°50.47′ S , 62°35.12′ W @@ -197,12 +197,16 @@ b.s.l. 1 Dec. 2015 -; FjordEco1; soft sediment; +; +FjordEco1 +; +soft sediment +; NHMD 1784660 • - -1 juv. + +1 juv. (mounted for SEM); CRS 1793; 64°39.53′ S , @@ -216,14 +220,18 @@ b.s.l. 11 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; MVS • - + 3 ♂♂ , -2 juv. +2 juv. (mounted for SEM); CRS 1832; 64°39.30′ S , @@ -237,7 +245,11 @@ b.s.l. 21 Apr. 2016 -; FjordEco2; soft sediment; +; +FjordEco2 +; +soft sediment +; MVS . @@ -257,12 +269,12 @@ GENERAL. Adults with head, neck and eleven trunk segments ( , 4A–D ). An overview of measurements and dimensions is given in -Table 5 +Table 5 . Distributions of cuticular structures, i.e., sensory spots and spines, are summarized in -Table 6 +Table 6 . Only males and juveniles were available for examination, so female dimorphism remains unknown for this species. - + Table 4. Number of identified adult specimens per species and station. @@ -707,7 +719,7 @@ NECK. Consists of 16 placids with condyles ( Fig. 4D ). All placids are about 12 µm in length, but differ in width. The midventral placid is the broadest, 15 µm wide. It is flanked by two pairs of narrow placids, each measuring 7 µm in width. From the two pairs of narrow, ventromedial placids, the placids alternate between broad (width: 13 µm) and narrow (width: 7 µm) ones towards the narrow middorsal placid. All condyles form structures with a narrow posterior part that broadens anteriorly, which makes them resemble small mushrooms. The midventral placid has three condyles, whereas other broad placids have two condyles, and narrow ones have a single. - + Table 5. Measurements from light microscopy of @@ -901,7 +913,7 @@ Neuhaus 4D–G ). - + Table 6. Summary of nature and location of sensory spots, tubes, and spines arranged by series in @@ -1203,7 +1215,7 @@ ventromedial positions. The tergal plate is covered with minute, triangular hair ). - + Distribution @@ -1218,9 +1230,11 @@ Antarctic Peninsula: Gerlache Strait and Andvord Bay MBA, . See Fig. 1 for geographic overview of stations and -Table 1 +Table 1 for station and specimen information. + + Diagnostic remarks diff --git a/data/03/B5/61/03B5612FFA02FFCD0FD7AACFD771ED43.xml b/data/03/B5/61/03B5612FFA02FFCD0FD7AACFD771ED43.xml new file mode 100644 index 00000000000..90b625ea59d --- /dev/null +++ b/data/03/B5/61/03B5612FFA02FFCD0FD7AACFD771ED43.xml @@ -0,0 +1,232 @@ + + + +The first leucospid wasp from the fossil record (Hymenoptera: Leucospidae) + + + +Author + +Engel, Michael S. +Department of Ecology and Evolutionary Biology, and Division of Entomology, Natural History Museum, University of Kansas, Lawrence, KS 66045 - 7523, USA + +text + + +Journal of Natural History + + +2001 + +2001-12-30 + + +36 + + +4 + + +435 +441 + + + +journal article +310840 +10.1080/00222930110059682 +e428b0ca-62b9-46db-b886-fa483a87a5cc +1464-5262 +10083332 + + + + + + +Leucospis glaesaria + +sp. nov. + + + + + +( +figures 1-6 +) + + + + + + +Holotype +. + +Female +( +figure 1 +), early +Miocene +amber, +Dominican Republic +, +AMNH +, +DR-10-186 9 +(specific mine unknown). Labelled`Holotype, + +Leucospis glaesaria +Engel. + +’ Deposited in the amber fossil collection, Department of Entomology, American Museum of Natural History, New York, USA + +. + + + + +Diagnosis. +The new species is distinctive for the combination of sparse pubescence, metallic cupreous integument on the mesosoma and metasoma, absence of a semicircular depression separating the lower tooth of the mandible ( +figure 3 +), a strong premarginal carina on the pronotum, carinate borders on the dorsellum, contiguous punctures on the mesosoma and metasoma ( +figure 2 +), a truncate metatibial apex ( +figure 5 +), long outer metatibial spur ( +figure 5 +), a long metabarsitarsus ( +figure 5 +), absence of a medial carina on the propodeum, a laterally conspicuous occipital carina, eight teeth on the ventral margin of the metafemur ( +figure 5 +), and a relatively short ovipositor ( +figure 4 +). It is most similar to + +L. speifera + +but differs by the absence of distinct interspaces between punctures, the cupreous body coloration, a depression along the posterior margin of the scutellum, absence of a medial carina on the propodeum and the sparse pubescence. + + + + +FIGS 1, 2. Photomicrographs of female holotype of + +Leucospis glaesaria + +. (1) Lateral habitus. (2) Detailed lateral view showing surface sculpturing. + + + + +FIGS 3-5. Female holotype of + +Leucospis glaesaria + +; setae and sculpturing omitted. (3) Face, frontal view; antennae omitted except for the position of one scape. (4) Metasomal apex. (5) Hind leg. Scale bars=1 mm; ep, epimerum; stippled outline indicates impunctate area on metacoxa. + + + + +Description. +Female: Total body length (exclusive of antennae) +11.6 mm +; forewing length +11.3 mm +. Dorsal head width +3 mm +, length +0.9 mm +; frontal head width +3 mm +, length +2.2 mm +. Width of scrobes +0.6 mm +. Clypeal length +0.5 mm +, width +0.5 mm +. Malar length +0.4 mm +. Compound eye length +1.5 mm +, width +1.2 mm +; upper interorbital distance +1.5 mm +; lower interorbital distance +1.4 mm +. Interocellar distance (POL of BoucÏek) +0.4 mm +; ocellocular distance (OOL of BoucÏek) +0.4 mm +. Scape length +0.7 mm +, width +0.2 mm +; pedicel length +0.2 mm +; flagellum length +4 mm +. Intertegular distance +3 mm +. Metasoma length +6.1 mm +. Body generally metallic cupreous except as follows: mandibular apex black; head just above lower tangent of compound eyes to vertex dark, metallic green; gena dark, metallic green; apices of claws black; forewing somewhat infuscate on anterior third beyond stigma, otherwise very weakly infumate. + + + +FIG. 6. Photomicrograph of female holotype of + +Leucospis glaesaria + +, dorsal habitus. + + + +Head narrower than mesoscutum, dorsally about 3.3 times as broad as long; occipital carina bordering lateral ocelli posteriorly, sharp, running down till just below centre of compound eyes. Scape minutely punctured, punctures separated by about a puncture width, integument between smooth. Upper carinate margin of scrobes less than one-half diameter of median ocellus from median ocellus, sharp along all margins ( +figure 3 +). Face with small, contiguous punctures; punctures slightly larger on vertex. Interantennal area weakly convex, with weak median carina. Gena punctured as on face. Head with exceedingly sparse, minute setae. + + +Pronotum, mesoscutum and scutellum with coarse, regular punctures, punctures contiguous ( +figure 6 +); those along posterior margin of scutellum elongate posteriorly, approximately two to three times longer than wide. Pronotum with strong, premarginal carina. Posterior margin of scutellum slightly depressed and forming a short lamella that slightly overhangs anterior margin of dorsellum. Dorsellum with two rows of strong alveolae, integument within smooth, lateral and posterior margins strongly carinate. Metanotum with single row of alveolae along posterior margin, anterior half with small, contiguous punctures. Propodeum two times wider than dorsellum, with coarse, contiguous punctures, medial carina absent. Mesopleuron above depression with coarse, contiguous punctures ( +figure 2 +); depression and preepisternum smooth with sparse, minute punctures. Metacoxa with coarse, contiguous punctures except large impunctate area on dorsal surface ( +figure 5 +), depression in posterior-dorsal surface impunctate; lateral surface with smaller punctures than those of dorsal surface, punctures separated by width or less, integument between smooth. Metafemur not very stout; ventral margin with broad basal tooth followed by seven smaller teeth ( +figure 5 +). Metatibia apex truncate; outer metatibial spur long; metabasitarsus dorsally longer than breadth of metatibia ( +figure 5 +). Mesosoma with sparse, minute setae except propodeum without pubescence. + + +Metasoma hardly longer than combined lengths of head and mesosoma. Anterior third of T1 (of gaster) impunctate, middle third with coarse, contiguous punctures, punctured area bordered posteriorly by short band of impunctate integument, posterior margin with a short band of coarse, contiguous punctures; T1 with premarginal constriction at point of posterior impunctate band. Remaining terga uniformly punctured, punctures coarse and contiguous. Ovipositor relatively short ( +figure 4 +). Metasoma with exceedingly sparse, minute setae. + +Male: Unknown. + + + +Etymology. +The specific epithet is taken from the Latin; +glaesarius +, meaning`of amber’. + + + + +Preservation. +The specimen is magni fi cently preserved in tact and without mould ( +figures 1, 2 +, +6 +). There is essentially no damage to the specimen and only the position of certain body parts obscure specific views (e.g. the left forewing is positioned such that it makes viewing of the left hind leg difficult, although not impossible). + + + + \ No newline at end of file diff --git a/data/03/E8/4E/03E84E71F7469433FFF180CEFDE9F9DE.xml b/data/03/E8/4E/03E84E71F7469433FFF180CEFC24F90F.xml similarity index 67% rename from data/03/E8/4E/03E84E71F7469433FFF180CEFDE9F9DE.xml rename to data/03/E8/4E/03E84E71F7469433FFF180CEFC24F90F.xml index 36bd6b3d2a3..3845b18cc4a 100644 --- a/data/03/E8/4E/03E84E71F7469433FFF180CEFDE9F9DE.xml +++ b/data/03/E8/4E/03E84E71F7469433FFF180CEFC24F90F.xml @@ -1,64 +1,64 @@ - - - -New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco + + + +New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco - - -Author + + +Author -Saidoun, Imane +Saidoun, Imane - - -Author + + +Author -El Mouden, Mohamed Amin +El Mouden, Mohamed Amin - - -Author + + +Author -Boussaa, Samia +Boussaa, Samia - - -Author + + +Author -Belqat, Boutaïna +Belqat, Boutaïna -text - - -Journal of Insect Biodiversity and Systematics +text + + +Journal of Insect Biodiversity and Systematics - -2025 - -2025-01-01 + +2025 + +2025-01-01 - -11 + +11 - -1 + +1 - -151 -169 + +151 +169 - -https://doi.org/10.61186/jibs.11.1.151 + +https://doi.org/10.61186/jibs.11.1.151 -journal article -10.61186/jibs.11.1.151 -2423-8112 -15810256 -36CE1DF6-3801-4A65-B784-5641542CD443 +journal article +10.61186/jibs.11.1.151 +2423-8112 +15810256 +36CE1DF6-3801-4A65-B784-5641542CD443 - + @@ -76,7 +76,7 @@ Eaton, 1893 - + Material examined. Middle Atlas: @@ -104,6 +104,8 @@ Middle Atlas: PO . + + General distribution. @@ -119,6 +121,8 @@ is a European, West-Siberian, and Caucasian species ( Ježek et al., 2023 ). + + Habitat. This species ( @@ -132,5 +136,13 @@ This species ( )) in the heart of the National Park of Tazekka, a protected area. + + +Comment. +Male genitalia have been illustrated by +Withers (1989:62) +. + + \ No newline at end of file diff --git a/data/03/E8/4E/03E84E71F7479432FFF18119FF2BF93C.xml b/data/03/E8/4E/03E84E71F7479432FFF18119FBD8F96C.xml similarity index 67% rename from data/03/E8/4E/03E84E71F7479432FFF18119FF2BF93C.xml rename to data/03/E8/4E/03E84E71F7479432FFF18119FBD8F96C.xml index b590dc4d013..f0688e8dd72 100644 --- a/data/03/E8/4E/03E84E71F7479432FFF18119FF2BF93C.xml +++ b/data/03/E8/4E/03E84E71F7479432FFF18119FBD8F96C.xml @@ -1,64 +1,64 @@ - - - -New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco + + + +New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco - - -Author + + +Author -Saidoun, Imane +Saidoun, Imane - - -Author + + +Author -El Mouden, Mohamed Amin +El Mouden, Mohamed Amin - - -Author + + +Author -Boussaa, Samia +Boussaa, Samia - - -Author + + +Author -Belqat, Boutaïna +Belqat, Boutaïna -text - - -Journal of Insect Biodiversity and Systematics +text + + +Journal of Insect Biodiversity and Systematics - -2025 - -2025-01-01 + +2025 + +2025-01-01 - -11 + +11 - -1 + +1 - -151 -169 + +151 +169 - -https://doi.org/10.61186/jibs.11.1.151 + +https://doi.org/10.61186/jibs.11.1.151 -journal article -10.61186/jibs.11.1.151 -2423-8112 -15810256 -36CE1DF6-3801-4A65-B784-5641542CD443 +journal article +10.61186/jibs.11.1.151 +2423-8112 +15810256 +36CE1DF6-3801-4A65-B784-5641542CD443 - + @@ -76,7 +76,7 @@ - + Material examined. @@ -102,6 +102,8 @@ . + + General distribution. This widespread species occurs in the Holarctic region ( @@ -126,6 +128,8 @@ in the High Atlas ( Wagner, 1987 ). + + Habitat. We collected specimens by sweep-netting the abundant vegetation, mostly @@ -147,5 +151,13 @@ We collected specimens by sweep-netting the abundant vegetation, mostly ). The watercourse, which was shallow and shaded, flowed slowly over a stony substrate with scattered boulders. + + +Comment. +A detailed description of the adult was given by +Ježek (1990) +. + + \ No newline at end of file diff --git a/data/03/E8/4E/03E84E71F7479433FFF18349FEA9FEB1.xml b/data/03/E8/4E/03E84E71F7479433FFF18349FB19FEE7.xml similarity index 59% rename from data/03/E8/4E/03E84E71F7479433FFF18349FEA9FEB1.xml rename to data/03/E8/4E/03E84E71F7479433FFF18349FB19FEE7.xml index a1babbfaa53..e47909c418c 100644 --- a/data/03/E8/4E/03E84E71F7479433FFF18349FEA9FEB1.xml +++ b/data/03/E8/4E/03E84E71F7479433FFF18349FB19FEE7.xml @@ -1,64 +1,64 @@ - - - -New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco + + + +New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco - - -Author + + +Author -Saidoun, Imane +Saidoun, Imane - - -Author + + +Author -El Mouden, Mohamed Amin +El Mouden, Mohamed Amin - - -Author + + +Author -Boussaa, Samia +Boussaa, Samia - - -Author + + +Author -Belqat, Boutaïna +Belqat, Boutaïna -text - - -Journal of Insect Biodiversity and Systematics +text + + +Journal of Insect Biodiversity and Systematics - -2025 - -2025-01-01 + +2025 + +2025-01-01 - -11 + +11 - -1 + +1 - -151 -169 + +151 +169 - -https://doi.org/10.61186/jibs.11.1.151 + +https://doi.org/10.61186/jibs.11.1.151 -journal article -10.61186/jibs.11.1.151 -2423-8112 -15810256 -36CE1DF6-3801-4A65-B784-5641542CD443 +journal article +10.61186/jibs.11.1.151 +2423-8112 +15810256 +36CE1DF6-3801-4A65-B784-5641542CD443 - + @@ -72,7 +72,7 @@ - + Material examined. Rif: @@ -85,6 +85,8 @@ Rif: 19.V.2022 , sweep net, coll. Group Belqat, UAE- FST. + + General distribution. @@ -100,6 +102,8 @@ is a rare European species (Vaillant, 1974; Oboňa et al., 2023b ). + + Habitat. This species occupies disparate Moroccan habitats, similar to those in Europe, having been found in the forested lotic habitat of the cascade Chrafate ( @@ -109,5 +113,13 @@ This species occupies disparate Moroccan habitats, similar to those in Europe, h ). + + +Comment. +Detailed descriptions of the adult were provided by +Ježek & Goutner (1993) +. + + \ No newline at end of file diff --git a/data/03/E8/4E/03E84E71F749943CFFF1811EFD58F95E.xml b/data/03/E8/4E/03E84E71F749943CFFF1811EFD58F95E.xml new file mode 100644 index 00000000000..c0acf5a8744 --- /dev/null +++ b/data/03/E8/4E/03E84E71F749943CFFF1811EFD58F95E.xml @@ -0,0 +1,140 @@ + + + +New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco + + + +Author + +Saidoun, Imane + + + +Author + +El Mouden, Mohamed Amin + + + +Author + +Boussaa, Samia + + + +Author + +Belqat, Boutaïna + +text + + +Journal of Insect Biodiversity and Systematics + + +2025 + +2025-01-01 + + +11 + + +1 + + +151 +169 + + + + +https://doi.org/10.61186/jibs.11.1.151 + +journal article +10.61186/jibs.11.1.151 +2423-8112 +15810256 +36CE1DF6-3801-4A65-B784-5641542CD443 + + + + + + + +Clytocerus +( +Boreoclytocerus +) + +sp. +** + + + + + + +Material examined. + +Rif: +1♂ +, Oued Belwazen, + +28.IV.2022 + +, +sweep net +, coll. +Saidoun +, FST- +UAE + +. + + + + +General distribution. +This genus has been found throughout Europe ( +Ježek et al., 2008 +; +Ježek et al., 2017 +; +Ježek et al., 2018a +), In North Africa, the genus was previously known only from +Algeria +( +Wagner, 1987 +) based on the species C. +kabylicus +Wagner, 1987 +( +Afzan & Belqat, 2016 +). + + + + +Habitat. +The specimens were captured ( +Figs 19, 20 +) in a forested mountain environment with streams having strong current, submerged vegetation (e.g., algae and mosses), coarse substrate, and abundant riparian vegetation such as + +Rubus ulmifolius + +( +Figs 7, 8 +). + + + + +Comment. +This species remains unnamed because of an issue during the mounting procedure. Consequently, the shape of the hypandrium, which is a crucial morphological characteristic for distinguishing the species, was not visible. + + + + \ No newline at end of file diff --git a/data/03/E8/4E/03E84E71F74A943FFFF181FFFBACF964.xml b/data/03/E8/4E/03E84E71F74A943FFFF181FFFC67F88A.xml similarity index 60% rename from data/03/E8/4E/03E84E71F74A943FFFF181FFFBACF964.xml rename to data/03/E8/4E/03E84E71F74A943FFFF181FFFC67F88A.xml index 86f75571fa3..4d253f8c36d 100644 --- a/data/03/E8/4E/03E84E71F74A943FFFF181FFFBACF964.xml +++ b/data/03/E8/4E/03E84E71F74A943FFFF181FFFC67F88A.xml @@ -1,64 +1,64 @@ - - - -New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco + + + +New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco - - -Author + + +Author -Saidoun, Imane +Saidoun, Imane - - -Author + + +Author -El Mouden, Mohamed Amin +El Mouden, Mohamed Amin - - -Author + + +Author -Boussaa, Samia +Boussaa, Samia - - -Author + + +Author -Belqat, Boutaïna +Belqat, Boutaïna -text - - -Journal of Insect Biodiversity and Systematics +text + + +Journal of Insect Biodiversity and Systematics - -2025 - -2025-01-01 + +2025 + +2025-01-01 - -11 + +11 - -1 + +1 - -151 -169 + +151 +169 - -https://doi.org/10.61186/jibs.11.1.151 + +https://doi.org/10.61186/jibs.11.1.151 -journal article -10.61186/jibs.11.1.151 -2423-8112 -15810256 -36CE1DF6-3801-4A65-B784-5641542CD443 +journal article +10.61186/jibs.11.1.151 +2423-8112 +15810256 +36CE1DF6-3801-4A65-B784-5641542CD443 - + @@ -76,7 +76,7 @@ - + Material examined. Beni Snassen: @@ -89,6 +89,8 @@ Beni Snassen: 11.XII.2018 , sweep net, coll. Group Belqat, UAE- FST. + + General distribution. This species is common and widely distributed in Europe ( @@ -103,6 +105,8 @@ This species is common and widely distributed in Europe ( Afzan & Belqat, 2016 ). + + Habitat. The adults are found in damp places from lowlands to mountains. We collected the species in a forest of @@ -122,5 +126,13 @@ and ). + + +Comment. +A description of the adult was given by +Ježek (1985) +. + + \ No newline at end of file diff --git a/data/03/E8/4E/03E84E71F74A943FFFF1876BFCF9FAAC.xml b/data/03/E8/4E/03E84E71F74A943FFFF1876BFB0AFAD2.xml similarity index 70% rename from data/03/E8/4E/03E84E71F74A943FFFF1876BFCF9FAAC.xml rename to data/03/E8/4E/03E84E71F74A943FFFF1876BFB0AFAD2.xml index e041e0a9cbe..c62b1485a15 100644 --- a/data/03/E8/4E/03E84E71F74A943FFFF1876BFCF9FAAC.xml +++ b/data/03/E8/4E/03E84E71F74A943FFFF1876BFB0AFAD2.xml @@ -1,64 +1,64 @@ - - - -New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco + + + +New records of genera and species of moth flies (Diptera, Psychodidae: Psychodinae) from Morocco - - -Author + + +Author -Saidoun, Imane +Saidoun, Imane - - -Author + + +Author -El Mouden, Mohamed Amin +El Mouden, Mohamed Amin - - -Author + + +Author -Boussaa, Samia +Boussaa, Samia - - -Author + + +Author -Belqat, Boutaïna +Belqat, Boutaïna -text - - -Journal of Insect Biodiversity and Systematics +text + + +Journal of Insect Biodiversity and Systematics - -2025 - -2025-01-01 + +2025 + +2025-01-01 - -11 + +11 - -1 + +1 - -151 -169 + +151 +169 - -https://doi.org/10.61186/jibs.11.1.151 + +https://doi.org/10.61186/jibs.11.1.151 -journal article -10.61186/jibs.11.1.151 -2423-8112 -15810256 -36CE1DF6-3801-4A65-B784-5641542CD443 +journal article +10.61186/jibs.11.1.151 +2423-8112 +15810256 +36CE1DF6-3801-4A65-B784-5641542CD443 - + @@ -76,7 +76,7 @@ - + Material examined. Rif: @@ -85,6 +85,8 @@ Rif: 24.V.2013 , reared, coll. Saidoun, UAE-FST. + + General distribution. Brazil @@ -108,6 +110,8 @@ Rif: Gibernau & Albre, 2022 ). + + Habitat. We found the species in a forest where the waterfall Cascade Chrafate is located. Abundant vegetation included @@ -151,5 +155,13 @@ wide, but the sections wetted at the time of capture were 2.6 wide. The water was cold (12.7 ºC), slightly mineralized (395 μS), with low salinity (0.2), a pH near neutral (7.5), and a strong to torrential current. Sunlight at the time of collection was medium. The substrate consisted of gravel, with scattered boulders and abundant riparian vegetation, submerged and floating mosses, and algae. Substrate, richer in mosses than algae, was removed from rocks, along with the soil, and transported to the laboratory to rear the aquatic stages. + + +Comment. +A detailed description of the adult has been given by +Cordeiro et al. (2011) +. + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406906DB2FC51BFBAA0E1502CA.xml b/data/03/FB/55/03FB55406906DB2FC51BFBAA0E1502CA.xml new file mode 100644 index 00000000000..8fedf30ebee --- /dev/null +++ b/data/03/FB/55/03FB55406906DB2FC51BFBAA0E1502CA.xml @@ -0,0 +1,185 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos tritaleum +Quate, 1996 + + + + + + +Figs 1 +, +23 + + + + + + + +Bryopharsos tritaleum +Quate, 1996: 41 + + +. +Type +locality: +Costa Rica +, +Heredia +, Estación Biológica La Selva (INBio). + + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows; wing 2.2 times as long as wide; ejaculatory apodeme ovoid in ventral view; gonocoxal apodeme with a spine-shaped projection ( +Fig. 23A +); surstyli with three tenacula ( +Fig. 23B +); aedeagus straight with a pointed apex ( +Fig. 23A +). This species is similar to + +B. asymmetricum + +sp. nov. +They can be easily differentiated by the number of facet rows in the eye bridge (four in + +B. tritaleum + +; five in + +B. asymmetricum + +), and the length of the ejaculatory apodeme (about as long as the aedeagus in + +B. tritaleum + +; shorter than the aedeagus in + +B. asymmetricum + +). + + +Female + +Unknown. + + + + +Material examined + + +None. + + + + +Distribution + + + +Costa Rica +( +Quate 1996 +; +Bravo & Araújo 2019 +) ( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406908DB2FC6B0FE130E15012D.xml b/data/03/FB/55/03FB55406908DB2FC6B0FE130E15012D.xml new file mode 100644 index 00000000000..1bec80b4342 --- /dev/null +++ b/data/03/FB/55/03FB55406908DB2FC6B0FE130E15012D.xml @@ -0,0 +1,291 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos tetracanthus +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +635E31CD-1373-4E14-B29E-381911CCAE3C + + + + + +Figs 1 +, +22 + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows ( +Fig. 22A +); wing 2.4 times as long as wide; ejaculatory apodeme cylindrical and hour-glass-shaped ( +Fig. 22B +), about the same length as the aedeagus ( +Fig. 22D +); gonocoxal apodeme fused; surstyli with three apical tenacula ( +Fig. 22B–C +); aedeagus digitiform, curved and evenly tapering towards apex ( +Fig. 22D +). This species shares the same number of apical tenacula in the surstyli (three tenacula) with + +B. tritaleum + +but it can be easily differentiated by the distribution of the apical tenacula in the surstyli (closely together at the apex in + +B. tetracanthus + +sp. nov. +, 2 apical tenacula separated from 1 basal tenaculum in + +B. tritaleum + +), and the shape of the aedeagus (digitiform and curved in + +B. tetracanthus + +, digitiform and straight in + +B. tritaleum + +). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name + +ʻ +tetracanthus + +ʼ derives from the Greek word ʻτετρα-ʼ (ʻtetra-ʼ) as a prefix, meaning four, and the Greek word ʻἄκαΝΘΟςʼ (ʻákanthosʼ), meaning spine. The specific name makes reference to the four spines located in between the apical tenacula in the surstyli. Specific name to be treated as a name in apposition. + + + + + +Type material + + + + +Holotype + + + + +COLOMBIA +– + +Magdalena + +• +♂ +; +Sierra Nevada de Santa Maria +, + +El Ramo + +; alt. + +2400 m + +; + +10–24 May 2000 + +; +I. Uribe +leg.; +Malaise trap +; +LACM +, LACM-ENT-279397. + + + + + + +Description + + +MEASUREMENTS. In mm (n = 1). Wing length: 1.92, wing width: 0.80; head length: 0.39, head width: 0.46; antennal segments: scape: 0.08, pedicel: 0.06, flagellomeres 1–4: 0.11; palpal segment 1: 0.05, palpal segment 2: 0.06, palpal segment 3: 0.06, palpal segment 4: 0.07. + + +Holotype +male + + + +HEAD ( +Fig. 22A +). A little wider than long; eye bridge contiguous, with four rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; the frontal patch of alveoli not divided, trapezoidal with the lower margin with a concavity in the middle. Antennal scape about the same length as pedicel, cylindrical; pedicel spherical, about the same length as scape; flagellomeres asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomeres absent in examined material, the maximum number of flagellomeres present is seven; ascoids absent in examined material. Palpal segments cylindrical, palpal segment 4 apically pointed, palpal proportions: 1.0:1.2:1.2:1.4; labium without any strong sclerite; labella not bulbous with 3–4 setae on outer margin. + + + +Fig. 22. + +Bryopharsos tetracanthus +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279397). +A +. Head. +B +. Terminalia (ventral view). +C +. Epandrium and surstyli. +D +. Terminalia (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of gonocoxal lobe; par = paramere; surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about 2.4 times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R +4 +; fork of R +2+3 +at the same level as M +1+2 +and joining R +4 +; fork of M +1+2 +normally sclerotized; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + + +TERMINALIA ( +Fig. 22B–D +). Hypandrium is a distinct band that connects the gonocoxites, stripe-like; gonocoxites cylindrical, shorter than gonostyli, gonostyli digitiform, with rounded apex; gonocoxal apodeme anteriorly projected with rectangular anterior margin; gonocoxal lobes with four setae on each side, spine of the gonocoxal lobes present; aedeagus digitiform and curved, evenly narrowing towards the apex, apex pointed, ending beyond the apex of the paramere, paramere digitiform, rounded at apex; ejaculatory apodeme with anterior margin rounded, shorter than the length of the aedeagus; epandrium plate-like, square and resembling an inverted U ( +Fig. 22C +); hypoproct elongated, almost thumb-like, about three times as long as wide, and covered in small setulae, epiproct shorter than hypoproct, about as wide as long; surstyli conical, tapering towards the apex and curved ventrally, with three close-together apical tenacula, tenacula with rounded apex, with additional 4 spiniform projections in-between the tenacula ( +Fig. 22B–D +). + + + + + +Distribution + + + +Only known from the +type +locality in +Colombia +( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB5540690BDB21C6ACFEEF09E90495.xml b/data/03/FB/55/03FB5540690BDB21C6ACFEEF09E90495.xml new file mode 100644 index 00000000000..7ecfb289a10 --- /dev/null +++ b/data/03/FB/55/03FB5540690BDB21C6ACFEEF09E90495.xml @@ -0,0 +1,420 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos septenacula +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +D9D297C4-05C9-4217-9EB0-9E0355EB9FB2 + + + + + +Figs 1 +, +20–21 + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows ( +Fig. 20A +); wing 2.1 times as long as wide; ejaculatory apodeme oval, about the same length as the aedeagus ( +Figs 20B +, +21 +); gonocoxal apodeme with spine-like anterior projection; surstyli with seven tenacula ( +Figs 20B +, +21 +); aedeagus digitiform and evenly tapering towards apex, and ending before the apex of paramere ( +Fig. 21 +), paramere shaped as an inverted J ( +Fig. 21B +). This species shares the same number of facet rows in the eye bridge as + +B. uncinatum + +, + +B. tritaleum + +, + +B. clavigum + +, + +B. amazonensis + +, and + +B. claviformosum + +, but it can be easily differentiated by the number of tenacula on the surstyli (seven in + +B. septenacula + +sp. nov. +, five in + +B. amazonensis + +and + +B. claviformosum + +, one in + +B. uncinatum + +, and four in + +B. tritaleum + +). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name derives from the Latin words ʻ +septem +ʼ meaning seven, and ʻ +tenaculum +ʼ (plural ʻ +tenacula +ʼ), the so-called stiff flattened setae on the inner apical surface of the surstyli of many +Psychodidae +. The name makes reference to the number of tenacula present in the species. Specific name to be treated as name in apposition. + + + + +Fig. 20. + +Bryopharsos septenacula +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (MECN, ZFMK-TIS-2637106). +A +. Head. +B +. Terminalia (ventral view). All scale bars are in millimeters. + + + + +Fig. 21. + +Bryopharsos septenacula +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (MECN, ZFMK-TIS-2637106). Terminalia (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of gonocoxal lobe; hyp = hypandrium; par = paramere; surs = surstyli; ten = tenacula. Scale bar in millimeters. + + + + + +Type material + + + + +Holotype + + + + +ECUADOR +– + +Pichincha + +• +♂ +; +Pedro Vicente Maldonado +, +Parroquia Pedro Vicente Maldonado +, near +San Pancracio +, roadway to +Pachijal +; +0.11561° N +, +78.95805° E +; alt. + +750 m + +; + +1–9 Feb. 2022 + +; +Isabel Kilian +leg.; +MECN +, ZFMK-TIS-2637106. + + + + +Paratypes + + + + +ECUADOR +– + +Pichincha + +• +5 ♂♂ +; same data as for holotype; +ZFMK +, ZFMK-TIS-2637088, +ZFMK- +TIS-2637153, ZFMK-TIS-2637157, ZFMK-TIS-2637174, ZFMK-TIS-2637170 + +• + +1 ♂ +; same data as for holotype; +MECN +, ZFMK-TIS-2637178 + +• + +1 ♂ +; same data as for holotype; +0.11862° N +, +78.95805° E +; alt. + +770 m + +; + +30 Dec. 2021 + +– + +5 Jan. 2022 + +; +ZFMK +, ZFMK-TIS-2636933 + +. – + + +Esmeraldas + +• +1 ♂ +; +Parroquia San Francisco del Cabo +, canton +Bunche +; +0.64562° N +, +80.0253° E +; alt. + +46 m + +; + +30 Dec. 2021 + +– + +5 Jan. 2022 + +; +ZFMK +, ZFMK-TIS-2636988 + +• + +2 ♂♂ +; same data as for preceding; + +25–28 Jan. 2020 + +; +ZFMK +, +ZFMK- +TIS-2629871, ZFMK-TIS-2629873 + +. + + + + + +Description + + +MEASUREMENTS. In mm (n = 5). Wing length: 1.70 (1.80–1.50), wing width: 0.79 (0.80–0.72); head length: 0.40 (0.40), head width: 0.35 (0.35); antennal segments: scape: 0.10 (0.11–0.08), pedicel: 0.06 (0.06–0.05), flagellomeres 1–7: 0.1 (0.11–0.10); palpal segment 1: 0.05 (0.5), palpal segment 2: 0.07 (0.07), palpal segment 3: 0.07 (0.07), palpal segment 4: 0.07 (0.07). + + +Holotype +male + + + +HEAD ( +Fig. 20A +). A little wider than long; eye bridge contiguous, with four rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; the frontal patch of alveoli not divided, trapezoidal with upper and lower margins straight. Antennal scape about two times the length of the pedicel, almost cylindrical; pedicel spherical, smaller than scape; flagellomeres asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomeres absent in examined material, the maximum number of flagellomeres present seven; ascoids rectangular and broad, about the same length, and about two times as wide as flagellomere carrying them. Palpal segments cylindrical, palpal segment 4 with pointed apex, palpal proportions: 1.0:1.5:1.5:1.5; labium without any strong sclerite; labella not bulbous with 3–4 setae on outer margin. + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about 2.1 times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R 4; fork of R 2+3 at the same level as M 1+2 and joining R +4 +; fork of M +1+2 +weak; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + +TERMINALIA ( +Figs 20B +, +21 +). Hypandrium is a distinct band that connects the gonocoxites, plate-like; gonocoxites about the same length as gonostyli, gonostyli slightly incurved, with rounded-blunt apex; aedeagus digitiform, evenly narrowing towards the apex, apex rounded, ending beyond the level of the paramere, paramere resembling an inverted ʻJʼ ( +Fig. 21 +), apex pointed; ejaculatory apodeme oval, about the same length as the aedeagus, anterior margin rounded; gonocoxal apodemes projected anteriorly and fused, resembling an inverted and wide ʻUʼ, spine of the gonocoxal lobes present; epandrium narrow, rectangular, about three times as wide long; hypoproct digitiform, shorter than epandrium and covered in small setulae, epiproct shorter than hypoproct; surstyli conical, slightly tapering towards the apex and curved ventrally, with seven apical tenacula, tenacula with rounded apex. + + + + + +Distribution + + + +Only known from the +type +locality in +Ecuador +( +Fig. 1 +). + + + +DNA barcodes + + + +Eleven specimens +were successfully sequenced ZFMK-TIS-2629871, ZFMK-TIS-2629873, ZFMK- TIS-2636933, ZFMK-TIS-2636988, ZFMK-TIS-2637088, ZFMK-TIS-2637106, ZFMK-TIS-2637153, ZFMK-TIS-2637157, ZFMK-TIS-2637170, ZFMK-TIS-2637174, and ZFMK-TIS-2637178. The maximum intraspecific uncorrected pairwise distance for COI sequences was 0.46% or 2 bp. + + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB5540690DDB25C6A0FAA90E1504E3.xml b/data/03/FB/55/03FB5540690DDB25C6A0FAA90E1504E3.xml new file mode 100644 index 00000000000..77be4d72a82 --- /dev/null +++ b/data/03/FB/55/03FB5540690DDB25C6A0FAA90E1504E3.xml @@ -0,0 +1,178 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos paulistensis +Bravo & Araújo, 2019 + + + + + + +Figs 1 +, +19 + + + + + + + +Bryopharsos paulistensis +Bravo & Araújo, 2019: 365 + + +. +Type +locality: +Brazil +, +São Paulo +, Sete Barras (MZFS). + + + + + + +Diagnosis + + + +Male + + +Eye bridge of five facet rows ( +Fig. 19A +); wing two times as long as wide; ejaculatory apodeme subrectangular in ventral view ( +Fig. 19B +); gonocoxal apodeme with anterior projection as a lobe ( +Fig. 19B +); surstyli with one tenaculum ( +Fig. 19C +); aedeagus long and expanded, hemispheric basally and narrowly pointed at the apex ( +Fig. 19B +). This species is similar to + +B. uncinatum + +, both can be easily differentiated by the number of facet rows in the eye bridge (four in + +B. uncinatum + +, five in + +B. paulistensis + +). + + +Female + +Unknown. + + + + +Material examined + + +None. + + + + +Distribution + + + +Brazil +( +Bravo & Araújo 2019 +) ( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406921DB17C6A4FBE60E1503B3.xml b/data/03/FB/55/03FB55406921DB17C6A4FBE60E1503B3.xml new file mode 100644 index 00000000000..faa40041c05 --- /dev/null +++ b/data/03/FB/55/03FB55406921DB17C6A4FBE60E1503B3.xml @@ -0,0 +1,328 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos bitenacula +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +D6D6DE95-B8A7-430B-B4B0-AD99DA55F54F + + + + + +Figs 1 +, +8–9 + + + + + +Diagnosis + + + +Male + + +Eye bridge with three facet rows ( +Fig. 8A +); wing 2.1 times as long as wide; ejaculatory apodeme ovoid, with anterior margin straight, shorter than the aedeagus; gonocoxal apodeme projecting anteriorly; surstyli with two apical tenacula; aedeagus digitiform, with rounded apex, paramere digitiform and longer than aedeagus. This species shares the same number of apical tenacula in the surstyli as + +B. asymmetricum + +sp. nov. +but it can be easily differentiated by the shape of the paramere (digitiform with pointed apex in + +B. asymmetricum + +, digitiform and with rounded apex in + +B. bitenacula + +sp. nov. +), and the length of the paramere (shorter than the aedeagus in + +B. bitenacula + +, longer than the aedeagus in + +B. asymmetricum + +). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name derives from the Latin word ʻ +bi +ʼ meaning two, and ʻ +tenacula +ʼ. It makes reference to the two apical tenacula present in the surstyli. To be treated as a noun in apposition. + + + + + +Type material + + + + +Holotype + + + + +PERU +– + +Cuzco + +• +♂ +; + +26 km +West of Pilcopata + +; +13.055° N +, +71.546667° E +; alt. + +1500 m + +; + +24 Jul.–2 Aug. 1997 + +; +L.W. Quate +leg.; +Malaise trap +; +Cloud Forest +; +Mounted in Euparal +; +LACM +, LACM-ENT-279272. + + + + +Paratypes + + + + +PERU +– + +Cuzco + +• +12 ♂♂ +; same data as for holotype; +LACM +, LACM-ENT-279266, LACM-ENT-279267, LACM-ENT-279268, LACM-ENT-279269, LACM-ENT-279270, LACM-ENT-279273, +LACM- +ENT-279274, LACM-ENT-279275, LACM-ENT-279277, LACM-ENT-279278, LACM-ENT-279290, LACM-ENT-279291 + +• + +1 ♂ +, same data as for holotype; + +25 Jul.–3 Aug. 1997 + +; +LACM +, LACM-ENT-279381 + +. + + + + +Fig. 8. + +Bryopharsos bitenacula +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279272). +A +. Head. +B +. Terminalia. +C +. Surstyli. Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of the gonocoxal lobes; par = paramere; surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + + +Fig. 9. + +Bryopharsos bitenacula +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279272). +A +. Terminalia (ventral view). +B +. Surstyli (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of the gonocoxal lobes; par = paramere; surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + + + +Description + + +MEASUREMENTS. In mm (n = 8). Wing length: 2.38 (2.24–2.45), wing width: 1.23 (1.17–1.22); head length: 0.43 (0.42–0.50), head width: 0.54 (0.52–0.68); antennal segments: scape: 0.10 (0.10–0.10), pedicel: 0.08 (0.07–0.08), flagellomere 1: 0.12 (0.12–0.13), flagellomeres 2–12: 0.15 (0.14–0.15), flagellomere 13: 0.06 (0.06–0.06), flagellomere 14: 0.08 (0.08–0.08); palpal segment 1: 0.04 (0.04–0.04), palpal segment 2: 0.08 (0.07–0.08), palpal segment 3: 0.07 (0.07–0.07), palpal segment 4: 0.06 (0.06–0.06). + + +Holotype +male + + + +HEAD ( +Fig. 8A +). A little wider than long; eye bridge contiguous, with five rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; frontal patch of alveoli not divided, rectangular with the lower margin straight, upper margin convex. Antennal scape slightly longer than the pedicel, cylindrical; pedicel spherical; 14 flagellomeres asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomere with terminal apiculus; ascoids about the same length as the flagellomere carrying them, and about two times wider than the width of flagellomere carrying them. Palpal segments cylindrical, palpal segment 4 apically pointed, palpal proportions: 1.0:1.8:1.6:1.4; labium without any strong sclerite; labella not bulbous with 3–4 setae on outer margin. + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about 2.1 times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R 4; fork of R 2+3 basal to the level of M 1+2 and joining R +4 +; fork of M +1+2 +normally sclerotized; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + + +TERMINALIA ( +Figs 8B–C +, +9 +). Hypandrium is a distinct band that connects the gonocoxites, plate-like; gonocoxites are cylindrical, about the same length as gonostyli, gonostyli digitiform; gonocoxal apodeme fused; gonocoxal lobes without anterior projections, with 3–5 setae on each side, with gonocoxal spine; aedeagus digitiform with rounded apex, ending beyond the apex of the paramere, paramere digitiform, with rounded apex, about ¾ the length of the aedeagus; ejaculatory apodeme with anterior margin straight, shorter than the aedeagus; epandrium rectangular, about two times as wide as long; hypoproct V-shaped, and covered in small setulae, epiproct broader and shorter than hypoproct; surstyli conical narrowing towards the apex, curved ventrally, with two apical tenacula, tenacula with rounded apex. + + + + + +Distribution + + + +Only known from the +type +locality in +Peru +( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406924DB08C555FEEF0FCB06E6.xml b/data/03/FB/55/03FB55406924DB08C555FEEF0FCB06E6.xml new file mode 100644 index 00000000000..f0952457f0c --- /dev/null +++ b/data/03/FB/55/03FB55406924DB08C555FEEF0FCB06E6.xml @@ -0,0 +1,336 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos bifidum +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +E3E364DD-E83D-4ADB-8A1F-CAB198C11DF1 + + + + + +Figs 1 +, +6–7 + + + + + +Diagnosis + + + +Male + + +Eye bridge with three facet rows ( +Fig. 6A +); wing 1.9 times as long as wide; ejaculatory apodeme ovoid, with anterior margin straight, slightly longer than the aedeagus; gonocoxal apodeme fused; surstyli with a single apical tenaculum; aedeagus digitiform, curved, with rounded apex. This species shares the same number of apical tenacula in the surstyli but it can be easily differentiated by having bifurcated gonostyli (no other known + +Bryopharsos +species + +has bifurcated gonostyli). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name + +ʻ +bifidum + +ʼ derives from Latin ʻ +bifidus +ʼ meaning split into two parts, referring to the bifurcate gonostyli. Name to be treated as a noun in apposition. + + + + + +Type material + + + + +Holotype + + + + +VENEZUELA +– + +Aragua + +• +♂ +; + +19 km +N of Maracay + +; + +14–17 Oct. 1993 + +; alt. + +1280 m + +; +L.W. Quate +leg.; +Malaise trap +; +Mounted in Euparal +; +LACM +, LACM-ENT-279295. + + + + +Paratype + + + + +VENEZUELA +– + +Choroní + +• +1 ♂ +; + +22 km +S of Choroní + +; + +17 Oct. 1993 + +; alt. + +1000 m + +.; +L.W. Quate +leg.; +Malaise Trap +; +Mounted in Euparal +; +LACM +, LACM-ENT-279294 + +. + + + + + +Description + + +MEASUREMENTS. In mm (n = 2). Wing length: 2.34 (2.24–2.45), wing width: 1.19 (1.17–1.22); head length: 0.60 (0.60–0.60), head width: 0.79 (0.71–0.86); antennal segments: scape: 0.10 (0.10–0.10), pedicel: 0.08 (0.07–0.08), flagellomere 1: 0.14 (0.13–0.14), flagellomeres 2–12: 0.15 (0.14–0.15), flagellomere 13: 0.06 (0.06–0.06), flagellomere 14: 0.08 (0.08–0.08); palpal segment 1: 0.05 (0.05–0,05), palpal segment 2: 0.05 (0.05–0.05), palpal segment 3: 0.05 (0.05–0.05), palpal segment 4: 0.04 (0.04–0.04). + + +Holotype +male + + + +HEAD ( +Fig. 6A +). A little wider than long; eye bridge contiguous, with three rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; frontal patch of alveoli not divided, triangular with the lower margin rounded. Antennal scape slightly longer the length of the pedicel, cylindrical; pedicel spherical; 14 flagellomeres asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomere with terminal apiculus; ascoids absent in examined material. Palpal segments cylindrical, palpal segment 4 apically pointed, palpal proportions: 1.0:1.0:1.0:0.8; labium without any strong sclerite; labella not bulbous with 3–4 setae on outer margin. + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about 2.1 times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R +4 +; fork of R +2+3 +basal to the level of M +1+2 +and joining R +4 +; fork of M +1+2 +weakly sclerotized; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + + + +Fig. 6. + +Bryopharsos bifidum +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279294). +A +. Head. +B +. Terminalia. +C +. Surstyli. +D +. Terminalia. Abbreviations: eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; surs = surstyli. All scale bars are in millimeters. + + + + +Fig. 7. + +Bryopharsos bifidum +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279294). +A +. Terminalia (ventral view). +B +. Surstyli (ventral view). Abbreviations: eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; surs = surstyli; ten = tenacula.All scale bars are in millimeters. + + + +TERMINALIA ( +Figs 6B–D +, +7 +). Hypandrium is a distinct band that connects the gonocoxites, plate-like; gonocoxites cylindrical, about the same length as gonostyli; gonostyli bifurcated, bifurcation occurs around half, both rami have rounded apex, mesal ramus digitiform, lateral ramus ovate; gonocoxal apodeme without anterior projections; gonocoxal lobes with six setae on each side; aedeagus digitiform, curved, apex rounded, ending beyond the apex of the paramere, paramere digitiform, almost thumb-like, with rounded apex; ejaculatory apodeme with rounded margin, about the same length as the aedeagus; epandrium plate-like, about two times as wide as long; hypoproct tongue-shaped, and covered in small setulae, epiproct broader and shorter than hypoproct; surstyli conical, abruptly narrowing towards the apex, curved ventrally, with a single apical tenaculum, tenaculum with rounded apex. + + + + + +Distribution + + + +Only known from the +type +locality in +Venezuela +( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + + +Remarks + + + +Both specimens presented a label with L.W. Quate’s handwriting reading “ + +Bryopharsos bifidum + +”, this specific name was never published nor was the description of the species, so we decided to keep the name chosen by Quate as it is fitting for the species. + + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406928DB0CC683FC420C9A04AE.xml b/data/03/FB/55/03FB55406928DB0CC683FC420C9A04AE.xml new file mode 100644 index 00000000000..ebb60b6c7f9 --- /dev/null +++ b/data/03/FB/55/03FB55406928DB0CC683FC420C9A04AE.xml @@ -0,0 +1,356 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos asymmetricum +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +40B57D7B-F290-4EF5-A80D-D7E78C86F1EC + + + + + +Figs 1 +, +4–5 + + + + + +Diagnosis + + + +Male + + +Eye bridge with five facet rows ( +Fig. 4A +); wing 2.2 times as long as wide; ejaculatory apodeme convex in ventral view; gonocoxal apodeme with spine projection; surstyli with 2 or 3 tenacula; aedeagus digitiform and evenly tapering towards apex ( +Fig. 4C +). This species is similar to + +B. tritaleum + +, but they can be differentiated by the number of tenacula (three in + +B. tritaleum + +, some specimens with three tenacula on one surstylus in + +B. asymmetricum + +sp. nov. +but the remaining with two tenacula), the number of facet rows in the eye bridge (four in + +B. tritaleum + +, five in + +B. asymmetricum + +), and the length of the ejaculatory apodeme (about as long as the aedeagus in + +B. tritaleum + +, shorter than the aedeagus in + +B. asymmetricum + +). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name + +ʻ +asymmetricum + +ʼ derives from the Greek word ʻσυμμετρικόςʼ (symetrikós) with the Greek prefix ʻἀ-ʼ (without), and it referrs to the asymmetrical number of apical tenacula on the surstyli. Specific name to be treated as an adjective. + + + + + +Type material + + + + +Holotype + + + + +ECUADOR +– + +Pichincha + +• +♂ +; +Pedro Vicente Maldonado +, +Parroquia Pedro Vicente Maldonado +, near +San Pancracio +, roadway to +Pachijal +; +0.11561° N +, +78.95805° E +; alt. + +750 m + +; + +1–9 Feb. 2022 + +; +Isabel Kilian +leg.; +MECN +, ZFMK-TIS-2637130. + + + + +Paratypes + + + + +ECUADOR +– + +Pichincha + +• +9 ♂♂ +; same data as for holotype; +ZFMK +, ZFMK-TIS-2629868, +ZFMK- +TIS-2629902, ZFMK-TIS-2636934, ZFMK-TIS-2637062, ZFMK-TIS-2637064, ZFMK-TIS-2637068 + +, + + + +Fig. 4. + +Bryopharsos asymmetricum +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (MECN, ZFMK-TIS-2637130). +A +. Head. +B +. Antennal segments. +C +. Terminalia (ventral view). Abbreviations: aed = aedeagus; asc = ascoids; eja = ejaculatory apodeme; fglm = flagellomere; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of the gonocoxal lobes; par = paramere; surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + +ZFMK-TIS-2637077, ZFMK-TIS-2637115, ZFMK-TIS-2637127 • + +1 ♂ +; same data as for holotype; +MECN +, ZFMK-TIS-2637149 + +. + + + + + +Description + + + +MEASUREMENTS. In mm (n = 6). Wing length: 1.90 (1.95–1.80), wing width: 0.85 (0.90–0.80); head length: 0.50 (0.50–0.45), head width: 0.40 (0.47–0.38); antennal segments ( +Fig. 4B +): scape: 0.11 (0.11–0.10), pedicel: 0.06 (0.06–0.05), flagellomeres 1–5: 0.1 (0.12–0.11); palpal segment 1: 0.05 (0.5–0.05), palpal segment 2: 0.07 (0.08–0.06), palpal segment 3: 0.07 (0.08–0.07), palpal segment 4: 0.08 (0.08–0.07). + + + +Holotype +male + + + +HEAD ( +Fig. 4A +). Slightly longer than wide; eye bridge contiguous with five rows of facets, interocular suture absent ( +Fig. 4A +); post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; the frontal patch of alveoli not divided, upper margin with a concavity in the middle, lower margin rounded. Antennal scape about two times as long as pedicel, almost cylindrical; pedicel spherical, smaller than scape; flagellomeres asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomeres absent in examined material, the maximum number of flagellomeres present five; ascoids rectangular and broad, about the same length, and about two times as wide as flagellomere carrying them. Palpal segments cylindrical, palpal segment 4 with pointed apex, palpal proportions: 1.0:1.5:1.5:1.6; labium without any strong sclerite; labella bulbous with seven three setae on outer margin and two setae on inner margin. + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about 2.2 times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R +4 +; fork of R +2+3 +at the same level as M +1+2 +and joining R +4 +; fork of M +1+2 +weak; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + + +TERMINALIA ( +Figs 4C +, +5 +). Hypandrium is a distinct band that connects the gonocoxites, broad and plate-like; gonocoxites about the same length as gonostyli, gonostyli slightly incurved, with rounded-blunt apex; aedeagus digitiform, apex rounded, with an additional digitiform paramere, paramere evenly narrowing towards the apex, longer than aedeagus; ejaculatory apodeme shorter than aedeagus, basal margin convex; gonocoxal apodemes projected anteriorly as a trapezoidal plate, fused; epandrium narrow, about three times wider than long, with posterior margin concave; hypoproct tongue-shaped, longer than epandrium and covered in small setulae, epiproct shorter than hypoproct; surstyli evenly tapering towards the apex and curved, one with three apical tenacula, the other with two tenacula, tenacula with rounded apex. + + + + + +Distribution + + + +Only known from the +type +locality in +Ecuador +( +Fig. 1 +). + + + +DNA barcodes + + + +Eleven specimens +were successfully sequenced: ZFMK-TIS-2629868, ZFMK-TIS-2629902, ZFMK- TIS-2636934, ZFMK-TIS-2637062, ZFMK-TIS-2637064, ZFMK-TIS-2637068, ZFMK-TIS-2637077, ZFMK-TIS-2637115, ZFMK-TIS-2637127, ZFMK-TIS-2637130, ZFMK-TIS-2637149. All eleven obtained sequences are identical. + + + + + +Remarks + + + +The +holotype +and paragypes ZFMK-TIS-2629868, ZFMK-TIS-2637115, and ZFMK-TIS-2637149 have three tenacula in one surstylus and two in the other, while the +paratypes +ZFMK-TIS-2629902 and ZFMK- TIS-2637127 only present two tenacula on both surstyli (no empty alveoli for a missing tenaculum is distinguishable), which leads us to believe this character is variable. The presence of one tenaculum in one surstylus and two on the other has also been recorded in the +holotype +of + +B. paulistensis + +( +Bravo & Araujo 2019 +: fig. 24). Furthermore, the spine in the gonocoxal apodeme is present either on the right side or on the left side, which is also a variable character inside the examined material. Nonetheless, the COI sequence of all specimens is identical, supporting our hypothesis that all the specimens belong to the same species. + + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB5540692CDB01C6A9F8A208550645.xml b/data/03/FB/55/03FB5540692CDB01C6A9F8A208550645.xml new file mode 100644 index 00000000000..4e7cac779ac --- /dev/null +++ b/data/03/FB/55/03FB5540692CDB01C6A9F8A208550645.xml @@ -0,0 +1,397 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos amazonensis +Bravo & Araújo, 2019 + + + + + + +Figs 1–3 + + + + + + + +Bryopharsos amazonensis +Bravo & Araújo, 2019: 368 + + +. Type locality: +Brazil +, +15 km +SE of Rio Branco, EMBRAPA ( +holotype +male, MZFS). + + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows ( +Fig. 2A +); wing 1.8 times as long as wide ( +Fig. 2B +); ejaculatory apodeme paddle-shaped in ventral view; surstyli with five tenacula of equal length; aedeagus long, triangular, and with a pointed apex ( +Figs 2C +, +3A–B +). According to +Bravo & Araújo (2019) +, + +B. amazonensis + +is morphologically similar to + +B. clavigum + +and + +B. claviformosum + +, but it can be differentiated by the number of tenacula (four in + +B. clavigum + +; five in + +B. amazonensis + +and + +B. claviformosum + +) and the length of the tenacula (four long and one short in + +B. claviformosum + +( +Fig. 3C–D +), five of equal length in + +B. amazonensis + +; four of the same length in + +B. clavigum + +). + + +Female + +Unknown. + + + +Fig. 1. +Species distribution map. Species dots do not represent the actual localities; jitter was added for visualization as the species’ real distributions overlap. + + + + +Fig. 2. + +Bryopharsos amazonensis +Bravo & Araújo, 2019 + +, ♂ (ZFMK-TIS-2628286). +A +. Head. +B +. Wing. +C +. Terminalia (ventral view). Abbreviations: aed = aedeagus; gns = gonostylus; gnx = gonocoxite; surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + + + +Material examined + + + + +ECUADOR +– + +Pichincha + +• +9 ♂♂ +; +Pedro Vicente Maldonado +, +Parroquia Pedro Vicente Maldonado +, near +San Pancracio +, roadway to +Pachijal +; +0.11862° N +, +78.95802° E +; alt. + +750 m + +; + +25–28 Jan. 2022 + +; +Isabel Kilian +leg.; +ZFMK +, ZFMK-TIS-2628286, ZFMK-TIS-2628287, ZFMK-TIS-2628288, +ZFMK- +TIS-2628289, ZFMK-TIS-2628298, ZFMK-TIS-2628299, ZFMK-TIS-2628303, ZFMK-TIS-2628304, ZFMK-TIS-2628307 + +• + +2 ♂♂ +; same data as for preceding; +0.11561° N +, +78.95805° E +; + +1–9 Feb. 2022 + +; +ZFMK +, ZFMK-TIS-2636932, ZFMK-TIS-2636935 + +• + +6 ♂♂ +; same data as for preceding; +MECN +, +ZFMK- +TIS-2636989, ZFMK-TIS-2637097, ZFMK-TIS-2637102, ZFMK-TIS-2637110, ZFMK-TIS-2637122, ZFMK-TIS-2637145 + +• + +8 ♂♂ +; same data as for preceding; +MECN +, ZFMK-TIS-2637067, +ZFMK- +TIS-2637069, ZFMK-TIS-2627070, ZFMK-TIS-2627072, ZFMK-TIS-2627074, ZFMK-TIS-2627075, ZFMK-TIS-2627076, ZFMK-TIS-2627078 + +. + + + +COLOMBIA +– + +Cauca + +• +5 ♂♂ +Guapí +, +Gorgona Island +, alta + +El Mirador + +; alt. + +180 m + +; + +4–24 Mar. 2000 + +; +R. Doque +leg.; +LACM +, +LACM +ENT 279390 +, +LACM +ENT 279391 +, +LACM +ENT 279392 +, +LACM +ENT 279399 +, +LACM +ENT 279398 + +. + + + + + +Distribution + + + +Brazil +( +Bravo & Araújo 2019 +), +Colombia +(this publication, new record), and +Ecuador +(this publication, new record) ( +Fig. 1 +). + + + +Fig. 3. +SEM pictures of male terminalia (ventral view). +A–B +. + +Bryopharsos amazonensis +Bravo & Araújo, 2019 + +. +C–D +. + +Bryopharsos claviformosum +Quate, 1996 + +. Colors: blue = paramere; cyan = gonocoxal lobes; green = hypandrium; orange = gonostyli; red = aedeagus; yellow = gonocoxites. All scale bars are in millimeters. + + + + +DNA barcodes + + + +22 specimens +were successfully sequenced: ZFMK-TIS-2628286, ZFMK-TIS-2628288, ZFMK- TIS-2628289, ZFMK-TIS-2628298, ZFMK-TIS-2628303, ZFMK-TIS-2628304, ZFMK-TIS-2628307, ZFMK-TIS-2636932, ZFMK-TIS-2636935, ZFMK-TIS-2636989, ZFMK-TIS-2637067, ZFMK- TIS-2637069, ZFMK-TIS-2627070, ZFMK-TIS-2627072, ZFMK-TIS-2627074, ZFMK-TIS-2627075, ZFMK-TIS-2627076, ZFMK-TIS-2627078, ZFMK-TIS-2637097, ZFMK-TIS-2637102, ZFMK- TIS-2637110, ZFMK-TIS-2637122. The maximum intraspecific uncorrected pairwise distance for COI sequences was 0.37% or 3 bp. + + + + + +Remarks + + + +Three specimens +(ZFMK-TIS-2636935, ZFMK-TIS-2636989, ZMFK-TIS-2637097) have a small morphological variation as they present a spine in the gonocoxal apodeme, while the remaining specimens do not present the spine. The molecular evidence and the remaining morphological characters suggest all specimens belong to the same species. The original differentiation of + +B. amazonensis + +and + +B. claviformosum + +by +Bravo & Araújo (2019) +based on the difference in tenacula lengths (five of the same length in + +B. amazonensis + +; four of equal length and one shorter in + +B. claviformosum + +) appears to be a good morphological character to separate them. This is further supported by molecular data in this study. + + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406930DB24C566FB5C085F0032.xml b/data/03/FB/55/03FB55406930DB24C566FB5C085F0032.xml new file mode 100644 index 00000000000..c3a9dd84c40 --- /dev/null +++ b/data/03/FB/55/03FB55406930DB24C566FB5C085F0032.xml @@ -0,0 +1,565 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos palpiculum +Quate, 1996 + + + + + + +Figs 1 +, +17–18 + + + + + + + +Bryopharsos palpiculum +Quate, 1996: 41 + + +. +Type +locality: +Costa Rica +, +Heredia +, Estación Biológica La Selva (INBio). + + + + + + +Diagnosis + + + +Male + + +Eye bridge with five facet rows ( +Fig. 17A +); wing 2.4 times as long as wide; ejaculatory apodeme subcircular in ventral view; gonocoxal apodeme with a spine-shaped projection; surstyli with four tenacula ( +Fig. 17C +); aedeagus long and straight, narrowing at the apex ( +Fig. 17B +). This species shares five face rows with + +B. paulistensis + +but they can be easily differentiated by the number of tenacula on the surstyli (four in + +B. palpiculum + +, one in + +B. paulistensis + +). + + +Female +(adapted from +Quate 1999 +) + + +Subgenital plate with slender constriction before small apical lobes, triangular in shape before constriction; membranous plate indistinct; genital chamber faint, without distinct structure, appears as a spherical structure partially divided apically (see +Quate 1999: 434–435 +, fig. 7c). + + + + + +Material examined + + + + +COSTA RICA +– + +Heredia + +• +6 ♂♂ +; +Puerto Viejo de Sarapaqui +, +Estación Biológica La Selva +; +0.11862° N +, +78.95802° E +; alt. + +50–100 m + +; + +15 Dec. 1993 + +; +LACM +, +INBIO +CRI001470484 +, +INBIO +CRI001470634 +, +INBIO +CRI001470595 +, +INBIO +CRI001470134 +, +INBIO +CRI001470343 +, +INBIO +CRI001470240 + +. + +ECUADOR +– + +Pichincha + +• +1 ♂ +; +Pedro Vicente Maldonado +, near +San Pancracio +, roadway to +Pachijal +; +0.11862° N +, +78.95805° E +; alt. + +770 m + +; + +25–28 Jan. 2020 + +; +Isabel Kilian +leg.; +MECN +, ZFMK-TIS-2628311 + +• + +1 ♀ +; same data as for preceding; +ZFMK +, ZFMK-TIS-2629904 + +• + +2 ♂♂ +; same data as for preceding; +0.1156° N +, +78.9580° E +; alt. + +750 m + +; + +1–9 Feb. 2022 + +; +ZFMK +, ZFMK-TIS-2637073, ZFMK-TIS-2637080 + +• + +1 ♂ +; +MECN +, ZFMK-TIS-2637071 + +. + + + +NICARAGUA +– + +Rio San Juan + +• +5 ♂♂ +SE of +San Carlos +; +10.9666° N +, +84.3333° E +; alt. + +30 m + +; + +6–10 Feb. 2000 + +; +L.W. Quate +leg.; +Malaise trap +; lowland rain forest; +LACM +LACM-ENT-279365, +LACM- +ENT-279366, LACM-ENT-279367, LACM-ENT-279368, LACM-ENT-279369 + +. + + + +PANAMA +– + +Canal Zone + +• +1 ♂ +; +Barro Colorado Island +; +9.1500° N +, +79.8500° E +; + +11–18 Aug. 1993 + +; +J. Pickering +leg.; +LACM +, LACM-ENT-279380 + +• + +1 ♂ +; same collection data as for preceding; + +28 Jul.–4Aug. 1993 + +; +LACM + +• + +1 ♂ +; same data as for preceding; + +31 Jan.–7 Feb. 1996 + +; +LACM +, LACM-ENT-279374 + +• + +1 ♂ +; same data as for preceding; + +6–13 Oct. 1996 + +; +LACM +, LACM-ENT-279373 + +• + +1 ♂ +; same data as for preceding; + +23–30 Oct. 1996 + +; +LACM +, LACM-ENT-279372 + +• + +1 ♂ +; same data as for preceding; + +18–27 Dec. 1996 + +; +LACM +, LACM-ENT-279370 + +• + +1 ♂ +; same data as for preceding; 1996; +LACM +, LACM-ENT-279378. – + +Guna Yala + +(previously known as +San Blas +) + +• + +1 ♂ +; +Nusagandi Reserve +; +9.3333° N +, +79.0000° E +; + +12–19 Feb. 1994 + +; +J. Pickering +leg; +LACM +, LACM-ENT-279377 + +• + +3 ♂♂ +; same data as for preceding; + +24 Jun.–4 Jul. 1993 + +; +LACM +, LACM-ENT-279371; LACM-ENT-279379; LACM-ENT-279376 + +. + + + + +Fig. 17. + +Bryopharsos palpiculum +Quate, 1996 + +, ♂ (LACM, INBIO CRI001470240). +A +. Head. +B +. Terminalia (ventral view). +C +. Surstyli. All scale bars are in millimeters. + + + + + +Redescription + + + +Female + +HEAD. About as long as wide, eye bridge contiguous with four facet rows; interocular suture absent; five enlarged post-ocular alveoli on each lateral margin; frontal patch of alveoli undivided, triangular with rounded vertices. Antennal scape about 1.5 times as long as the pedicel, cylindrical; pedicel spherical, shorter than scape; flagellomeres smaller than male flagellomeres, asymmetrical-podiform, apical flagellomeres missing in examined material; palpal segments cylindrical, apical palpal segment with pointed apex; palpal proportions: 1.0:1.2:1.2:1.5. + +THORAX. Without allurement organs. Wing. 2.3 times as long as wide, wing membrane brown-hyaline, alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R +4 +; fork of R +2+3 +at the same level as M +1+2 +and joining R +4 +; fork of M +1+2 +weak; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. Tergite nine with apical lobes ( +Fig. 18A +), broad and digitiform, about the same length as tergite nine; subgenital plate membranous except for apical lobes which are sclerotized, basal margin round, with a constriction before apical lobes; genital chamber membranous, barely visible; cerci short, about the same length as the subgenital plate. + + + +Fig. 18. + +Bryopharsos palpiculum +Quate, 1996 + +, (ZFMK-TIS-2629904). ♀ +A +. Terminalia (ventral view). +B +. Egg. All scale bars are in millimeters. + + + +EGG ( +Fig. 18B +). Length 0.336 ± +0.020 mm +. Width 0.14 ± +0.032 mm +(n = 8). No exochorion sculptures where observed along the long axis of the egg. No aeropiles were observed in the posterior pole. The anterior pole presents a cylindrical projection of about +0.02 mm +, with a truncated apex, without exochorion sculptures. + + + + + +Distribution + + + +Costa Rica +, +Panama +( +Quate 1996 +; +Bravo & Araújo 2019 +), and +Ecuador +(this publication, new record) ( +Fig. 1 +). + + + +DNA barcodes + + + +Five specimens +were successfully sequenced: ZFMK-TIS-2628311, ZFMK-TIS-2629904, ZFMK- TIS-2637071, ZFMK-TIS-2637073, ZFMK-TIS-2637080. The maximum intraspecific uncorrected pairwise distance for COI sequences was 1.07% or 7 bp. + + + + + +Remarks + + + +Quate (1996) +originally described + +B. palpiculum + +based on male specimens. Later, he ( +Quate 1999 +) described the females collected in Nusagandi Reserve ( +Panama +) appealing to the co-occurrence of these female specimens with males of + +B. palpiculum + +. These females were the first known and reported female specimens of + +Bryopharsos + +. The female description is relatively brief and the illustration only shows the genital chamber and the subgenital plate. + + +In this study, the association between the examined male and female specimens was done using the DNA barcodes (see section above). Our specimen differs from that on the drawings provided by +Quate (1999) +, i.e., in specimen ZFMK-TIS-2637073 the paramere looks shorter than in Quate’s drawing while in specimen ZFMK-TIS-2637080, the paramere is strongly curved (although the shape/position likely changed during the mounting process). In specimen ZFMK-TIS-2637071, the base of the paramere looks broader than on Quate’s drawing, but the general morphology is similar. The variation can be explained by the angle in which the terminalia were prepared on the microscope slides, but when comparing with other specimens deposited in other collections there is no doubt they all belong to the same species. + + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406933DB19C6AFFBDC0E15015F.xml b/data/03/FB/55/03FB55406933DB19C6AFFBDC0E15015F.xml new file mode 100644 index 00000000000..317db174ce8 --- /dev/null +++ b/data/03/FB/55/03FB55406933DB19C6AFFBDC0E15015F.xml @@ -0,0 +1,318 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos insperatum +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +EE933B1F-738A-48DE-9693-4129341979F4 + + + + + +Figs 1 +, +15–16 + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows ( +Fig. 15A +); wing 2.2 times as long as wide; ejaculatory apodeme cylindrical and hour-glass-shaped ( +Fig. 15B +), about the same length as the aedeagus; gonocoxal apodeme fused; surstyli with one tenaculum ( +Figs 15C–D +, +16B +); aedeagus digitiform, out curved and evenly tapering towards apex ( +Fig. 16 A +). This species shares the same number of apical tenaculum in the surstyli (one apical tenaculum) with + +B. uncinatum + +and + +B. paulistensis + +. Still, it can be easily differentiated by the number of rows of facets in the eye bridge (three in + +B. insperatum + +sp nov., four in + +B. uncinatum + +, and five in + +B. paulistensis + +). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name + +ʻ +insperatum + +ʼ derives from the Latin word ʻ +insperatus +ʼ (neuter + +ʻ +insperatum + +ʼ), meaning unexpected, referring to unexpectedly finding a new species while looking in the collections. Name to be treated as an adjective. + + + + + +Type material + + + + +Holotype + + + + +COSTA RICA +– + +Heredia + +• +♂ +; +Limon +Rs. Biol. Hitoy Cerere +, +Rio Cerrere +, sidestream; +9.806667° N +, +83.0175° E +; alt. + +100–200 m + +; + +17–26 Feb. 1999 + +; +L.W. Quate +leg.; +Malaise trap +; +LACM +, +INBIO +CRI001473066 +. + + + + + + +Description + + +MEASUREMENTS. In mm (n = 1). Wing length: 2.87, wing width: 1.29; head length: 0.52, head width: 0.61; antennal segments: scape: 0.14, pedicel: 0.09, flagellomere 1: 0.18, flagellomeres 2–12: 0.20; palpal segment 1: 0.07, palpal segment 2: 0.10, palpal segment 3: 0.10, palpal segment 4: 0.20. + + +Holotype +male + + + +HEAD ( +Fig. 15A +). A little wider than long; eye bridge contiguous, with four rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; the frontal patch of alveoli not divided, triangular with lower margin straight. Antennal scape about two times the length of the pedicel, cylindrical; pedicel spherical, smaller than scape; flagellomeres asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomeres absent in examined material, the maximum number of flagellomeres present is 12; ascoids rectangular and broad, about the same length, and about two times as wide as flagellomere carrying them. Palpal segments cylindrical, palpal segment 4 apically pointed, with bilobed apex, palpal proportions: 1.0:1.4:1.4:2.8; labium without any strong sclerite; labella not bulbous with 3–4 setae on outer margin. + + + +Fig. 15. + +Bryopharsos insperatum +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM, INBIO CRI001473066). +A +. Head. +B +. Terminalia (ventral view). +C +. Surstylus. +D +. Surstylus (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; par = paramere, surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + + +Fig. 16. + +Bryopharsos insperatum +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM, INBIO CRI001473066). +A +. Terminalia (ventral view). +B +. Surstyli. Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; par = paramere; surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about 2.2 times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R 4; fork of R 2+3 at the basal to the level of M +1+2 +and joining R +4 +; fork of M +1+2 +weakly sclerotized, almost appears as it is not joining; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + + +TERMINALIA ( +Figs 15B–D +, +16 +). Hypandrium is a distinct band that connects the gonocoxites, plate-like; gonocoxites cylindrical, longer than gonostyli, gonostyli conical, with rounded-blunt apex; gonocoxal apodeme not anteriorly projected; gonocoxal lobes posteriorly projected on each side, each lobe with five setae; aedeagus digitiform and out curved, evenly narrowing towards the apex, apex rounded, and ending beyond the apex of the paramere, paramere digitiform ( +Fig. 16A +), broader than aedeagus, with apex rounded; ejaculatory apodeme with anterior margin round ( +Fig. 16A +), about the same length of the aedeagus; epandrium not discernable in examined material, but appears U-shaped; hypoproct tongue-shaped ( +Fig. 15D +), and covered in small setulae, epiproct shorter than hypoproct; surstyli conical, slightly tapering towards the apex and curved ventrally, with one apical tenaculum, tenaculum with rounded apex. + + + + + +Distribution + + + +Only known from the +type +locality in +Costa Rica +( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406935DB1AC559FE760E1506DC.xml b/data/03/FB/55/03FB55406935DB1AC559FE760E1506DC.xml new file mode 100644 index 00000000000..55a943da82c --- /dev/null +++ b/data/03/FB/55/03FB55406935DB1AC559FE760E1506DC.xml @@ -0,0 +1,314 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos gorgona +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +7BE7FB52-C67C-417B-86AF-CCA712ADB470 + + + + + +Figs 1 +, +14 + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows ( +Fig. 14A +); wing 2.5 times as long as wide; ejaculatory apodeme short, with anterior margin straight, shorter than the aedeagus ( +Fig. 14B, D +); gonocoxal apodeme projecting anteriorly; surstyli with six apical tenacula ( +Fig. 14B–C +); aedeagus digitiform, with rounded apex, paramere digitiform and longer than aedeagus. This species shares the same number of apical tenacula in the surstyli as + +B. gorgona + +sp. nov. +but it can be easily differentiated by the length of the hypandrium (hypandrium length is longer than aedeagal width in + +B. gorgona + +, hypandrium length is shorter than aedeagal width in + +B. chuspi + +sp. nov. +), and the shape of the epandrium (rectangular in + +B. gorgona + +, U-shaped in + +B. chuspi + +). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name + +ʻ +gorgona + +ʼ derives from the name of the +type +locality, Gorgona Island. To be treated as a name in apposition. + + + + + +Type material + + + + +Holotype + + + + +COLOMBIA +– + +Cauca + +• +♂ +; +Guapí +, +Gorgona Island +, alta + +El Mirador + +; +2.9689° N +, +78.1856° E +; alt. + +180 m + +; + +4–24 Mar. 2000 + +; +R. Doque +leg.; +LACM +, LACM-ENT-279394. + + + + +Paratypes + + + + +COLOMBIA +– + +Cauca + +• +2 ♂♂ +; same data as for holotype; +LACM +, +LACM +–ENT-279395, +LACM- +ENT-279388 + +. + + + + + +Description + + +MEASUREMENTS. In mm (n = 3). Wing length: 2.27 (2.204–2.55), wing width: 0.90 (0.85–0.96); head length: 0.40 (0.40–0.40), head width: 0.45 (0.45–0.45); antennal segments: scape: 0.10 (0.10–0.10), pedicel: 0.06 (0.06–0.06), flagellomere 1: 0.11 (0.11–0.12), flagellomeres 2–9: 0.12 (0.12–0.12), palpal segment 1: 0.05 (0.05–0.06), palpal segment 2: 0.07 (0.07–0.07), palpal segment 3: 0.07 (0.07–0.07), palpal segment 4: 0.08 (0.08–0.08). + + +Holotype +male + + + +HEAD ( +Fig. 14A +). A little wider than long; eye bridge contiguous, with four rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; the frontal patch of alveoli not divided, rectangular with the lower and upper margins straight. Antennal scape is about two times as long as the pedicel, cylindrical; pedicel spherical; flagellomeres are asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomeres absent, the maximum number of flagellomeres present in examined material: 9; ascoids absent in examined material. Palpal segments cylindrical, palpal segment 4 apically pointed, palpal proportions: 1.0:1.3:1.3:1.5; labium without any strong sclerite; labella not bulbous with 3–4 setae on the outer margin. + + + +Fig. 14. + +Bryopharsos gorgona +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279394). +A +. Head. +B +. Terminalia (ventral view). +C +. Surstyli (ventral view). +D +. Terminalia (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of the gonocoxal lobes; par = paramere; surs = surstyli; ten = tenacula. All scale bars are in millimeters. + + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about two times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R +4 +; fork of R +2+3 +basal to the level of M +1+2 +and joining R + +4 +; fork of M +1+2 +weakly sclerotized; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + +TERMINALIA ( +Fig. 14B–D +). Hypandrium is a distinct band that connects the gonocoxites, plate-like; gonocoxites are cylindrical, about two-thirds the length of the gonostyli, gonostyli digitiform; gonocoxal lobes without anterior projection, fused, and with 4–5 setae on each side; aedeagus digitiform with rounded apex, ending slightly after the apex of the paramere, paramere digitiform, evenly tapering towards apex, with pointed apex; ejaculatory apodeme with anterior margin straight, about the same length of the aedeagus; epandrium rectangular; hypoproct V-shaped, and covered in small setulae, epiproct broader and shorter than hypoproct; surstyli conical narrowing towards the apex, with six apical tenacula, tenacula with rounded apex. + + + + + +Distribution + + + +Only known from the +type +locality in +Colombia +( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406939DB10C51BFDF20E6F0364.xml b/data/03/FB/55/03FB55406939DB10C51BFDF20E6F0364.xml new file mode 100644 index 00000000000..27d8263847d --- /dev/null +++ b/data/03/FB/55/03FB55406939DB10C51BFDF20E6F0364.xml @@ -0,0 +1,255 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos clavigum +Quate, 1996 + + + + + + +Figs 1 +, +12 + + + + + + + +Bryopharsos clavigum +Quate, 1996: 41 + + +. +Type +locality: +Costa Rica +, +Heredia +, Estación Biológica La Selva (INBio). + + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows ( +Fig. 12A +); wing 2.5 times as long as wide; ejaculatory apodeme subcircular in ventral view ( +Fig. 12B +); gonocoxal apodeme with spine ( +Fig. 12B +); surstyli with four tenacula ( +Fig. 12C +). This species is morphologically similar to + +B. amazonensis + +and + +B. claviformosum + +but they can be differentiated by the number of tenacula present in the surstyli (four in + +B. clavigum + +, five in + +B. amazonensis + +and + +B. claviformosum + +). + + +Female + +Unknown. + + + + +Material examined + + + + +COSTA RICA +– + +Heredia + +• +4 ♂♂ +; +Puerto Viejo de Sarapaqui +, +Estación Biológica La Selva +; +0.11862° N +, +78.95802° E +; alt. + +50–100 m + +; + +15 Dec. 1993 + +; leg.; +LACM +, LACM-ENT-279401, LACM-ENT-279402, LACM-ENT-279403, +INBIO +CRI001470316 + +. + + + +ECUADOR +– + +Pichincha + +• +1 ♂ +; +Pedro Vicente Maldonado +, near +San Pancracio +, roadway to +Pachijal +, +0.11862° N +, +78.95805° E +; alt. + +770 m + +; + +25–28 Jan. 2020 + +; +Isabel Kilian +leg.; +MECN +, ZFMK-TIS-2628309 + +. + + + + + +Distribution + + + +Costa Rica +( +Quate 1996 +; +Bravo & Araújo 2019 +) and +Ecuador +(this publication, new record) ( +Fig. 1 +). + + + +DNA barcodes + + +DNA barcoding of the examined material was unsuccessful. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB55406939DB1CC555F9620E150470.xml b/data/03/FB/55/03FB55406939DB1CC555F9620E150470.xml new file mode 100644 index 00000000000..23f35d4ddf0 --- /dev/null +++ b/data/03/FB/55/03FB55406939DB1CC555F9620E150470.xml @@ -0,0 +1,298 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos curvum +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +8D75061B-8145-4250-9E15-E1AA5C0BF549 + + + + + +Figs 1 +, +13 + + + + + +Diagnosis + + + +Male +Eye bridge with three facet rows ( +Fig. 13A +); wing 1.9 times as long as wide; ejaculatory apodeme cylindrical and hour-glass-shaped, about the same length as the aedeagus ( +Fig. 13B, D +); aedeagus digitiform, outcurved and evenly tapering towards apex ( +Fig. 13B, D +); gonocoxal apodeme fused ( +Fig. 13B, D +); surstyli with one tenaculum ( +Fig. 13C +). This species shares the same number of apical tenacula in the surstyli with + +B. uncinatum + +and + +B. paulistensis + +. Still, it can be easily differentiated by the number of rows of facets in the eye bridge (three in + +B. curvum + +sp. nov. +, four in + +B. uncinatum + +, and five in + +B. paulistensis + +). + + + +Fig. 12. + +Bryopharsos clavigum +Quate, 1996 + +, ♂ (LACM-ENT-279403). +A +. Head. +B +. Terminalia (ventral view). +C +. Surstyli (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; par = paramere; surs = surstylus; ten = tenacula. All scale bars are in millimeters. + + + +Female + +Unknown. + + + + +Etymology + + + +Specific name is derived from Latin ʻ +curvus +ʼ (neuter ʻcurvumʼ), making reference to the outcurved gonocoxites, as well as the curved aedeagus and paramere. Specific name to be treated as a noun in apposition. + + + + + +Type material + + + + +Holotype + + + + +COLOMBIA +– + +Magdalena + +• +♂ +; +Sierra Nevada de Santa Marta +, + +El Ramo + +; alt. + +2400 m + +; + +10–24 Jun. 2000 + +; +I. Uribe +leg.; +Malaise trap +; +LACM +, LACM-ENT-279396. + + + + + + +Description + + +MEASUREMENTS. In mm (n = 1). Wing length: 2.47, wing width: 1.28; head length: 0.50, head width: 0.60; antennal segments: scape: 0.10, pedicel: 0.06, flagellomeres 1–4: 0.11 (0.11–0.12); palpal segment 1: 0.04, palpal segment 2: 0.8, palpal segment 3: 0.07, palpal segment 4: 0.05. + + +Holotype +male + + + +HEAD ( +Fig. 13A +). A little wider than long; eye bridge contiguous, with three rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; the frontal patch of alveoli not divided, triangular with lower margin straight. Antennal scape about two times the length of the pedicel, slightly broader than wide; pedicel spherical, smaller than scape; flagellomeres asymmetrical and nodiform, with scattered setae on the basal half surface, apical flagellomeres absent in examined material, the maximum number of flagellomeres present is four; ascoids absent in examined material. Palpal segments cylindrical, palpal segment 4 with pointed apex, palpal proportions: 1.0:2.0:1.7:1.2; labium without strong sclerites; labella not bulbous with 3–4 setae on outer margin. + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about 1.9 times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R +4 +; fork of R +2+3 +at the same level as M +1+2 +and joining R +4 +; fork of M +1+2 +weak; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + + +TERMINALIA ( +Fig. 13B–D +). Hypandrium is a distinct band that connects the gonocoxites, plate-like; gonocoxites cylindrical, shorter than gonostyli, gonostyli digitiform and outcurved, with rounded-blunt apex; aedeagus digitiform and outcurved, evenly narrowing towards the apex, apex pointed, ending at the level of the apex of the paramere, paramere digitiform and out curved, paramere evenly narrowing towards the apex, apex pointed; ejaculatory apodeme almost cylindrical, waisted and resembling a broad hour-glass shape, about the same length of the aedeagus; gonocoxal apodemes fused; epandrium like a narrow rectangle, about four times wider than long; hypoproct tongue-shaped, longer than epandrium and covered in small setulae, epiproct shorter than hypoproct; surstyli conical, evenly tapering towards the apex, with one apical tenaculum, tenaculum with rounded apex. + + + + + +Distribution + + + +Only known from the +type +locality in +Colombia +( +Fig. 1 +). + + + +Fig. 13. + +Bryopharsos curvum +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279396). +A +. Head. +B +. Terminalia (ventral view). +C +. Surstyli (ventral view). +D +. Terminalia (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; par = paramere; surs = surstylus; ten = tenacula. All scale bars are in millimeters. + + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB5540693ADB10C578FD080C2E04F4.xml b/data/03/FB/55/03FB5540693ADB10C578FD080C2E04F4.xml new file mode 100644 index 00000000000..0c58ad80965 --- /dev/null +++ b/data/03/FB/55/03FB5540693ADB10C578FD080C2E04F4.xml @@ -0,0 +1,268 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos claviformosum +Quate, 1996 + + + + + + +Figs 1 +, +3C–D + + + + + + + +Bryopharsos claviformosum +Quate, 1996: 41 + + +. +Type +locality: +Costa Rica +, +Heredia +, Estación Biológica La Selva (INBio). + + + + + +Bryopharsos claviformosus +– + + +Bravo & Araújo 2019: 370 + +(lapsus calami in identification key). + + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows; wing 2.5 times as long as wide; ejaculatory apodeme rectangular in ventral view; gonocoxal apodeme with a spine; surstyli with five tenacula. This species is similar to + +B. amazonensis + +and + +B. clavigum + +but it can be easily differentiated by the number of tenacula (four in + +B. clavigum + +, five in + +B. amazonensis + +and + +B. claviformosum + +) and the length of the tenacula (equal length in + +B. amazonensis + +and + +B. clavigum + +, four long and one shorter in + +B. claviformosum + +). + + +Female + +Unknown. + + + + +Material examined + + + + +ECUADOR +– + +Pichincha + +• +2 ♂♂ +; +Pedro Vicente Maldonado +, near +San Pancracio +, roadway to +Pachijal +; +0.1156° N +, +78.9580° E +; alt. + +750 m + +; + +1–9 Feb. 2022 + +; +Isabel Kilian +leg.; +ZFMK +, ZFMK-TIS-2637066, ZFMK-TIS-2637079 + +• + +1 ♂ +same data as for preceding; +0.115611° N +, +78.95805° E +; +MECN +, +ZFMK- +TIS-2637137 + +. + + + + + +Distribution + + + +Costa Rica +( +Quate 1996 +; +Bravo & Araújo 2019 +) and +Ecuador +(this publication, new record) ( +Fig. 1 +). + + + +DNA barcodes + + + +Three specimens +were successfully sequenced: ZFMK-TIS-2637066, ZFMK-TIS-2637079, and ZFMK- TIS-2637137. The maximum intraspecific uncorrected pairwise distance for COI sequences was 0.31% or 2 bp. + + + + + +Remarks + + + +The original description of the male by +Quate (1996) +is rather short and incomplete, with some important characters missing. Nonetheless, the drawings and the general description are enough to distinguish the males of this species. In the SEM pictures ( +Fig. 3C–D +), the aedeagus seems curved, with a rounded apex and the paramere seems digitiform and slender, while in the original description, the aedeagus looks straight and tapering towards the apex, while the paramere seems broad. We must point out that for our specimens prepared on permanent slides, the perspective affects the perceived shape of the aedeagus and paramere, making the aedeagus look straight and tapering towards the apex, and the paramere looks digitiform. + + + + \ No newline at end of file diff --git a/data/03/FB/55/03FB5540693EDB13C553F9280E150790.xml b/data/03/FB/55/03FB5540693EDB13C553F9280E150790.xml new file mode 100644 index 00000000000..78da0073d79 --- /dev/null +++ b/data/03/FB/55/03FB5540693EDB13C553F9280E150790.xml @@ -0,0 +1,313 @@ + + + +Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species + + + +Author + +Jaume-Schinkel, Santiago +CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +santijaumes@hotmail.com + + + +Author + +Kilian, Isabel C. +B1ED76B4-2C0C-4621-9F46-F602954DDAC6 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Department of Agroecology and Organic Farming, Institute of Crop Science and Resource Conservation, Faculty of Agriculture, University of Bonn, Auf dem Hügel 6, D- 53121 Bonn, Germany. +i.kilian@leibniz-lib.de + + + +Author + +Pazmiño-Palomino, Alex +3BA927BD-71D8-4917-8269-A987B30350D4 +Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. & Museo de Entomología, Facultad de Recursos Naturales, Escuela Superior Politécnica de Chimborazo, Panamericana Sur Km 1.5, Riobamba, Ecuador. +marabuntalex@gmail.com + + + +Author + +Mengual, Ximo +A509310D-B567-4830-B8A4-BCB139BB8768 +Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany. & Instituto Nacional de Biodiversidad, División de Entomología. Rumipamba N 341 y Av. Los Shyris, 170506 Quito, Ecuador. +x.mengual@leibniz-lib.de + +text + + +European Journal of Taxonomy + + +2025 + +2025-07-03 + + +1001 + + +1 + + +1 +51 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2951/13319 + +journal article +10.5852/ejt.2025.1001.2951 +2118-9773 +1726F665-E94D-45CD-83DC-3102CE2C1C50 + + + + + + +Bryopharsos chuspi +Jaume-Schinkel + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +DCC6BE41-DA48-451D-81C3-052088A67BDD + + + + + +Figs 1 +, +10–11 + + + + + +Diagnosis + + + +Male + + +Eye bridge with four facet rows ( +Fig. 10A +); wing 2.8 times as long as wide; ejaculatory apodeme about the same length as the aedeagus, with anterior margin rounded, gonocoxal apodeme projecting anteriorly; surstyli with six apical tenacula; aedeagus digitiform, tapering towards the apex, paramere digitiform, tapering towards the apex and longer than aedeagus. This species shares the same number of apical tenacula in the surstyli as + +B. gorgona + +sp. nov. +Still, it can be easily differentiated by the length of the hypandrium (hypandrium length is shorter than aedeagal width in + +B. chuspi + +sp. nov. +, hypandrium length is longer than aedeagal width) + +B. gorgona + +) and the shape of the epandrium (U-shaped in + +B. chuspi + +, rectangular in + +B. gorgona + +). + + +Female + +Unknown. + + + + +Etymology + + + +The specific name + +ʻ +chuspi + +ʼ derives from the Quechuan word ʻchuspiʼ meaning fly. Species name to be treated as a name in apposition. + + + + + +Type material + + + + +Holotype + + + + +PERU +– + +Cuzco + +• +♂ +; +26 km +W of +Pilcopata +; alt. + +1500 m + +; + +24 Jul.–2 Aug. 1997 + +; +L.W. Quate +leg.; +LACM +, LACM-ENT-279279. + + + + +Paratypes + + + + +PERU +– + +Cuzco + +• +12 ♂♂ +; same data as for holotype; +LACM +, LACM-ENT-279286, LACM-ENT-279285, LACM-ENT-279284, LACM-ENT-279276, LACM-ENT-279281, LACM-ENT-279282, +LACM- +ENT-279283, LACM-ENT-279280, LACM-ENT-279202, LACM-ENT-279289, LACM-ENT-279288, LACM-ENT-279287 + +. + + + + + +Description + + +MEASUREMENTS. In mm (n = 13). Wing length: 2.76 (2.50–3.00), wing width: 0.94 (0.90–1.00); head length: 0.44 (0.42–0.45), head width: 0.46 (0.46–0.48); antennal segments: scape: 0.12 (0.11–0.13), pedicel: 0.07 (0.07–0.08), flagellomere 1: 0.12 (0.11–0.12), flagellomeres 2–13: 0.14 (0.14–0.15), flagellomere 14: 0.16 (0.16–0.16); palpal segment 1: 0.06 (0.05–0.06), palpal segment 2: 0.06 (0.06–0.07), palpal segment 3: 0.08 (0.08–0.08), palpal segment 4: 0.11 (0.10–0.12). + + +Holotype +male + + + +HEAD ( +Fig. 10A +). A little wider than long; eye bridge contiguous, with four rows of facets, interocular suture absent; post-ocular alveoli not enlarged and non-distinguishable from the remaining alveoli on the head; frontal patch of alveoli not divided, rectangular with concave lateral margins, lower margin straight, upper margin partially divided in the middle. Antennal scape is about two times as long as the pedicel, cylindrical; the pedicel is spherical; flagellomeres are asymmetrical and nodiform, with scattered setae on the basal half surface, flagellomere 14 with terminal apiculus being two times as long as the elongated basal part of the flagellomere; ascoids about the same length as the flagellomere carrying them, and about two times as wide as the width of flagellomere. Palpal segments cylindrical, palpal segment 4 apically pointed, palpal proportions: 1.0:1.0:1.3:1.8; labium without any strong sclerite; labella not bulbous with 3–4 setae on outer margin. + + +THORAX. Without allurement organs; all coxae with a stripe of three to five rows of alveoli. Wing length about two times its width; wing membrane brown-hyaline; alveoli distributed uniformly on wing membrane; subcostal vein short ending beyond the origin of R +4 +; fork of R +2+3 +basal to the level of M +1+2 +and joining R +4 +; fork of M +1+2 +weakly sclerotized; R +5 +ending at the wing apex; CuA +2 +ending at wing margin. + + +TERMINALIA ( +Figs 10B–C +, +11 +). Hypandrium is a distinct band that connects the gonocoxites, plate-like; gonocoxites are cylindrical, about two-thirds the length of the gonostyli, gonostyli digitiform; gonocoxal apodeme fused; gonocoxal lobe with 5–6 setae on each side, spine of the gonocoxal lobes present; aedeagus digitiform, with rounded apex, ending at about the same level as the paramere, paramere digitiform, tapering towards apex, with pointed apex; ejaculatory apodeme with anterior margin rounded, about the same length as the aedeagus; epandrium U-shaped; hypoproct U-shaped, and covered in small setulae, epiproct broader and shorter than hypoproct; surstyli conical narrowing towards the apex, with six apical tenacula, tenacula with rounded apex. + + + + +Fig. 10. + +Bryopharsos chuspi +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279279). +A +. Head. +B +. Terminalia (ventral view). +C +. Surstyli. Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of the gonocoxal lobes; surs = surstylus; ten = tenacula. All scale bars are in millimeters. + + + + +Fig. 11. + +Bryopharsos chuspi +Jaume-Schinkel + +sp. nov. +, holotype, ♂ (LACM-ENT-279279). +A +. Terminalia (ventral view). +B +. Surstyli (ventral view). Abbreviations: aed = aedeagus; eja = ejaculatory apodeme; gns = gonostylus; gnx = gonocoxite; gnx lb = gonocoxal lobes; gnx sp = spine of the gonocoxal lobes; hyp = hypandrium; par = paramere; surs = surstylus; ten = tenacula. All scale bars are in millimeters. + + + + + +Distribution + + + +Only known from the +type +locality in +Peru +( +Fig. 1 +). + + + +DNA barcodes + + +No specimens were available for DNA extraction. + + + \ No newline at end of file