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<document id="B04DE275B7F574109995A5E521B6B978" ID-DOI="10.11646/zootaxa.3709.3.6" ID-GBIF-Dataset="84532554-175c-45bf-b83f-7cebff1bf25f" ID-ISSN="1175-5326" ID-Zenodo-Dep="217457" ID-ZooBank="0E6D2049-59E5-4134-B393-233897D3EA4B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460673791363" checkinUser="plazi" docAuthor="Tong, Yanfeng &amp; Li, Shuqiang" docDate="2013" docId="8F1B87C5D957FFF929D520E5FA9F5865" docLanguage="en" docName="zt03709p284.pdf" docOrigin="Zootaxa 3709 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trilacuna hamata Tong &amp; Li, 2013, sp. nov." docType="treatment" docVersion="6" lastPageNumber="278" masterDocId="7322FFBDD956FFF82942207DFFE25E4E" masterDocTitle="The first goblin spiders of the genus Trilacuna from Vietnam (Araneae, Oonopidae)" masterLastPageNumber="284" masterPageNumber="277" pageNumber="278" updateTime="1698349130180" updateUser="plazi">
<mods:mods id="104C28CC5AA5ADD8FE5FD781EEF2D548" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BA98779CB2D894E593377DDB34472AB9">
<mods:title id="D859BA492868A117BC6A077C1B49F7F4">The first goblin spiders of the genus Trilacuna from Vietnam (Araneae, Oonopidae)</mods:title>
</mods:titleInfo>
<mods:name id="48AE108F08C1A8E0CCD2EF12B88D9C21" type="personal">
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<mods:namePart id="F13C485B0865D4151A8C4F1A858E046F">Tong, Yanfeng</mods:namePart>
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<mods:namePart id="E81598417EA6D277D0EE0B3D0505BAD7">Li, Shuqiang</mods:namePart>
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<mods:title id="4D5976BBB873C95E9D696248FECA4F34">Zootaxa</mods:title>
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<mods:date id="E03A31DD42A54E50A94A030FC7518D42">2013</mods:date>
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<mods:number id="9E3B3835E15FD7CDE2A4FAB4F76CFBEA">3709</mods:number>
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<mods:identifier id="8325538AFB614BD95697F39F68EB8B3E" type="DOI">10.11646/zootaxa.3709.3.6</mods:identifier>
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<treatment id="8F1B87C5D957FFF929D520E5FA9F5865" ID-DOI="http://doi.org/10.5281/zenodo.6145090" ID-GBIF-Taxon="119578997" ID-Zenodo-Dep="6145090" LSID="urn:lsid:plazi:treatment:8F1B87C5D957FFF929D520E5FA9F5865" httpUri="http://treatment.plazi.org/id/8F1B87C5D957FFF929D520E5FA9F5865" lastPageNumber="278" pageId="1" pageNumber="278">
<subSubSection id="4FA86558D957FFF929D520E5FEBF5E9D" pageId="1" pageNumber="278" type="nomenclature">
<paragraph id="070D36D3D957FFF929D520E5FE375EFC" blockId="1.[151,469,152,211]" box="[151,469,152,178]" pageId="1" pageNumber="278">
<heading id="5C4581BFD957FFF929D520E5FE375EFC" bold="true" box="[151,469,152,178]" fontSize="11" level="1" pageId="1" pageNumber="278" reason="1">
<emphasis id="35C6EAC1D957FFF929D520E5FE375EFC" bold="true" box="[151,469,152,178]" pageId="1" pageNumber="278">
<taxonomicName id="C0B24D50D957FFF929D520E5FE925EFC" ID-CoL="7D66K" box="[151,368,152,178]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="278" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="35C6EAC1D957FFF929D520E5FE925EFC" bold="true" box="[151,368,152,178]" italics="true" pageId="1" pageNumber="278">Trilacuna hamata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EF557BAD957FFF9283A20E5FE375EFC" box="[376,469,152,178]" pageId="1" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="070D36D3D957FFF929D520C7FEBF5E9D" blockId="1.[151,469,152,211]" box="[151,349,186,211]" pageId="1" pageNumber="278">
(
<figureCitation id="9F892A56D957FFF929DD20C7FEE15E9D" box="[159,259,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1915,1937]" captionStartId-1="3.[151,250,1917,1939]" captionStartId-2="4.[151,250,1750,1772]" captionTargetBox-0="[204,1389,194,1894]" captionTargetBox-1="[252,1342,183,1873]" captionTargetBox-2="[151,1436,193,1728]" captionTargetId-0="figure@2.[204,1390,193,1894]" captionTargetId-1="figure@3.[236,1351,175,1914]" captionTargetId-2="figure@4.[151,1436,193,1729]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." captionText-1="FIGURE 2. Trilacuna hamata sp. nov., male. A, B, left palp, prolateral and retrolateral views; C – H, left embolus, prolateral (C, F), retrolateral (D, G) and dorsal (E, H) views. Abbreviations: dh = dorsal hairs; ffp = filiform, curved projection; mlo = membranous lobe; nb = narrow branch; rdh = retrolateral-dorsally hairs; sso = slightly sclerotized outgrowth. Scales: A, B, 0.2 mm; C – H, 0.1 mm." captionText-2="FIGURE 3. Trilacuna hamata sp. nov., female. A – D, prosoma, dorsal, lateral, frontal and ventral views; E, abdomen, anterior view; F, abdomen, ventral view; G, labium and endites, ventral view; H, abdomen, lateral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A – G, 0.2 mm; H, 0.4 mm." httpUri-0="https://zenodo.org/record/217458/files/figure.png" httpUri-1="https://zenodo.org/record/217459/files/figure.png" httpUri-2="https://zenodo.org/record/217460/files/figure.png" pageId="1" pageNumber="278">Figs 1–3</figureCitation>
,
<figureCitation id="9F892A56D957FFF9284D20C6FEB75E9D" box="[271,341,187,211]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">4A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4FA86558D957FFF929D5217FFA835F71" pageId="1" pageNumber="278" type="materials_examined">
<paragraph id="070D36D3D957FFF929D5217FFA835F71" blockId="1.[151,1437,258,1579]" pageId="1" pageNumber="278">
<emphasis id="35C6EAC1D957FFF929D5217FFEA75F55" bold="true" box="[151,325,258,283]" pageId="1" pageNumber="278">
<typeStatus id="D8098871D957FFF929D5217FFF305F55" box="[151,210,258,283]" pageId="1" pageNumber="278">Type</typeStatus>
material.
</emphasis>
<specimenCount id="11B4FD5AD957FFF9280E217EFE7A5F55" box="[332,408,258,283]" pageId="1" pageNumber="278" type="male">1 male</specimenCount>
<typeStatus id="D8098871D957FFF928E2217FFDE05F55" box="[416,514,258,283]" pageId="1" pageNumber="278" type="holotype">holotype</typeStatus>
(
<collectionCode id="61A3AE16D957FFF92B53217EFD865F55" box="[529,612,259,283]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="278">IZCAS</collectionCode>
Ar 19419), Tam Dao National Park, Vinh Phuc,
<collectingCountry id="7FA57643D957FFF92DC8217FFB0E5F55" box="[1162,1260,258,283]" name="Vietnam" pageId="1" pageNumber="278">Vietnam</collectingCountry>
, pitfall traps, in natural forest, Pham Dinh-Sac,
<date id="730C1013D957FFF928BA215AFD465F71" box="[504,676,295,319]" pageId="1" pageNumber="278" value="2007-11-01" valueMax="2007-11-30" valueMin="2007-11-01">1–30. XI. 2007</date>
;
<specimenCount id="11B4FD5AD957FFF92BF2215AFCF35F71" box="[688,785,294,319]" pageId="1" pageNumber="278" type="female">1 female</specimenCount>
<typeStatus id="D8098871D957FFF92A5A215AFC9A5F71" box="[792,888,295,319]" pageId="1" pageNumber="278" type="paratype">paratype</typeStatus>
(
<collectionCode id="61A3AE16D957FFF92AC5215AFC385F71" box="[903,986,295,319]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="278">IZCAS</collectionCode>
Ar 19420), same data as
<typeStatus id="D8098871D957FFF92DB5215BFABF5F71" box="[1271,1373,294,319]" pageId="1" pageNumber="278" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="4FA86558D957FFF929852137FCE45FC9" pageId="1" pageNumber="278" type="etymology">
<paragraph id="070D36D3D957FFF929852137FCE45FC9" blockId="1.[151,1437,258,1579]" pageId="1" pageNumber="278">
<emphasis id="35C6EAC1D957FFF929852137FEAC5F2D" bold="true" box="[199,334,330,355]" pageId="1" pageNumber="278">Etymology.</emphasis>
The specific name is Latin, meaning “hook-shaped” and refers to the shape of the strongly curved outgrowth of the male embolus (
<figureCitation id="9F892A56D957FFF92B462113FDA85FC9" box="[516,586,366,391]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1939]" captionTargetBox="[252,1342,183,1873]" captionTargetId="figure@3.[236,1351,175,1914]" captionTargetPageId="3" captionText="FIGURE 2. Trilacuna hamata sp. nov., male. A, B, left palp, prolateral and retrolateral views; C – H, left embolus, prolateral (C, F), retrolateral (D, G) and dorsal (E, H) views. Abbreviations: dh = dorsal hairs; ffp = filiform, curved projection; mlo = membranous lobe; nb = narrow branch; rdh = retrolateral-dorsally hairs; sso = slightly sclerotized outgrowth. Scales: A, B, 0.2 mm; C – H, 0.1 mm." httpUri="https://zenodo.org/record/217459/files/figure.png" pageId="1" pageNumber="278">Figs 2</figureCitation>
A–H); adjective.
</paragraph>
</subSubSection>
<subSubSection id="4FA86558D957FFF9298521EFFB945FBD" pageId="1" pageNumber="278" type="diagnosis">
<paragraph id="070D36D3D957FFF9298521EFFB945FBD" blockId="1.[151,1437,258,1579]" pageId="1" pageNumber="278">
<emphasis id="35C6EAC1D957FFF9298521EFFEA35FE5" bold="true" box="[199,321,402,427]" pageId="1" pageNumber="278">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="C0B24D50D957FFF92BDB21E9FBE65FE5" authority="Grismado &amp; Piacentini" authorityName="Grismado &amp; Piacentini" box="[665,1028,402,427]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="278" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="35C6EAC1D957FFF92BDB21E9FD145FE5" box="[665,758,402,427]" italics="true" pageId="1" pageNumber="278">T. loebli</emphasis>
Grismado &amp; Piacentini
</taxonomicName>
, but can be distinguished from it by the smooth sternum (
<figureCitation id="9F892A56D957FFF928C621CBFE275F81" box="[388,453,438,463]" captionStart="FIGURE 3" captionStartId="4.[151,250,1750,1772]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@4.[151,1436,193,1729]" captionTargetPageId="4" captionText="FIGURE 3. Trilacuna hamata sp. nov., female. A – D, prosoma, dorsal, lateral, frontal and ventral views; E, abdomen, anterior view; F, abdomen, ventral view; G, labium and endites, ventral view; H, abdomen, lateral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A – G, 0.2 mm; H, 0.4 mm." httpUri="https://zenodo.org/record/217460/files/figure.png" pageId="1" pageNumber="278">Fig. 3</figureCitation>
D), the fold on the anterior margin of the postepigastric scutum in females (
<figureCitation id="9F892A56D957FFF92C5421CBFA765F81" box="[1302,1428,438,463]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Figs 4A, C</figureCitation>
) and the nearly rectangular, sponge-like sclerite (rss) in the female genitalia (
<figureCitation id="9F892A56D957FFF92AA821A7FB8B5FBD" box="[1002,1129,474,499]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Figs 4B, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4FA86558D957FFF929852183FA9F5865" pageId="1" pageNumber="278" type="description">
<paragraph id="070D36D3D957FFF929852183FEA35A41" blockId="1.[151,1437,258,1579]" pageId="1" pageNumber="278">
<emphasis id="35C6EAC1D957FFF929852183FEBB5C59" bold="true" box="[199,345,510,535]" pageId="1" pageNumber="278">Description.</emphasis>
Male. Completely chitinized. Body yellow-brown, chelicerae and sternum lighter, legs yellow. Habitus as in
<figureCitation id="9F892A56D957FFF92877225FFE9A5C75" box="[309,376,546,571]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">Figs 1</figureCitation>
A, B. Body length 1.72; carapace 0.81 length, 0.65 width; abdomen 0.85 length, 0.53 width. Carapace broadly oval, slightly elevated, dorsal surface smooth, with three rows of short hairs between eyes and posterior border of dorsal elevation of carapace; sides granulated; lateral margin rebordered, with a row of short, finely hairs and small blunt denticles.
<emphasis id="35C6EAC1D957FFF92B1522F2FD6E5CE8" box="[599,652,655,678]" italics="true" pageId="1" pageNumber="278">Eyes</emphasis>
six, well developed, arranged in a compact group, about subequal, roundish (
<figureCitation id="9F892A56D957FFF9284822CFFEB35C85" box="[266,337,690,715]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">Figs 1</figureCitation>
C, D); posterior row recurved from above, straight from front (
<figureCitation id="9F892A56D957FFF92D6F22CFFB915C85" box="[1069,1139,690,715]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">Figs 1</figureCitation>
C, F).
<emphasis id="35C6EAC1D957FFF92D8122CFFAFF5C85" box="[1219,1309,690,715]" italics="true" pageId="1" pageNumber="278">Clypeus</emphasis>
sinuous in frontal view, anterior lateral eyes separated from edge of carapace by about 1.5 times their diameter, with needlelike setae.
<emphasis id="35C6EAC1D957FFF9285F2287FE425D5D" box="[285,416,762,787]" italics="true" pageId="1" pageNumber="278">Mouthparts</emphasis>
: Chelicerae straight, proximal region with one hair with elevated hair base; labium rectangular, not fused to sternum, anterior margin deeply incised (ldi) (
<figureCitation id="9F892A56D957FFF92AA32363FBC45D79" box="[993,1062,798,823]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">
Fig.
<date id="730C1013D957FFF92D582362FBC45D79" box="[1050,1062,799,823]" pageId="1" pageNumber="278">1</date>
</figureCitation>
I); endites slender, distally only slightly branched (sdb) (
<figureCitation id="9F892A56D957FFF928EA233FFE0A5D15" box="[424,488,834,859]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">
Fig.
<date id="730C1013D957FFF9289E233EFE0A5D15" box="[476,488,835,859]" pageId="1" pageNumber="278">1</date>
</figureCitation>
I).
<emphasis id="35C6EAC1D957FFF92B49233EFD855D14" box="[523,615,835,858]" italics="true" pageId="1" pageNumber="278">Sternum</emphasis>
surface smooth with small pits on radial furrows between coxae, anterior margin with transverse groove; setae sparse, light, needle-like, evenly scattered (
<figureCitation id="9F892A56D957FFF92D5D231BFBBD5D31" box="[1055,1119,870,895]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">Fig. 1</figureCitation>
E).
<emphasis id="35C6EAC1D957FFF92DC8231BFB165D31" box="[1162,1268,870,895]" italics="true" pageId="1" pageNumber="278">Abdomen</emphasis>
: Ovoid, dorsal scutum not fused to epigastric scutum, postepigastric and epigastric scutum fused, apodemes absent, posterior spiracles not connected by groove (
<figureCitation id="9F892A56D957FFF92B6823D3FD8C5D89" box="[554,622,942,967]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">Figs 1</figureCitation>
G, H), booklung covers (boc) large, ovoid, surface smooth. Leg
<emphasis id="35C6EAC1D957FFF92C0E23D2FA7E5D88" box="[1356,1436,943,966]" italics="true" pageId="1" pageNumber="278">spines:</emphasis>
metatarsi I, II with 4 pairs of ventral spines each, tarsi I, II with 2 pairs of ventral spines each; legs III and IV without spines.
</paragraph>
<paragraph id="070D36D3D957FFF929852467FE095B45" blockId="1.[151,1437,258,1579]" pageId="1" pageNumber="278">
Male genitalia:
<emphasis id="35C6EAC1D957FFF928352461FDD95A7D" box="[375,571,1052,1075]" italics="true" pageId="1" pageNumber="278">Epigastric region</emphasis>
(
<figureCitation id="9F892A56D957FFF92B092467FD685A7D" box="[587,650,1050,1075]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1937]" captionTargetBox="[204,1389,194,1894]" captionTargetId="figure@2.[204,1390,193,1894]" captionTargetPageId="2" captionText="FIGURE 1. Trilacuna hamata sp. nov., male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C – H, 0.2 mm; I, 0.1 mm." httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="1" pageNumber="278">Fig. 1</figureCitation>
G) with sperm pore small, oval, rebordered, situated between anterior and posterior spiracles.
<emphasis id="35C6EAC1D957FFF928E72443FE385A19" box="[421,474,1086,1111]" italics="true" pageId="1" pageNumber="278">Palp</emphasis>
(
<figureCitation id="9F892A56D957FFF928A92443FDD15A19" box="[491,563,1086,1111]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1939]" captionTargetBox="[252,1342,183,1873]" captionTargetId="figure@3.[236,1351,175,1914]" captionTargetPageId="3" captionText="FIGURE 2. Trilacuna hamata sp. nov., male. A, B, left palp, prolateral and retrolateral views; C – H, left embolus, prolateral (C, F), retrolateral (D, G) and dorsal (E, H) views. Abbreviations: dh = dorsal hairs; ffp = filiform, curved projection; mlo = membranous lobe; nb = narrow branch; rdh = retrolateral-dorsally hairs; sso = slightly sclerotized outgrowth. Scales: A, B, 0.2 mm; C – H, 0.1 mm." httpUri="https://zenodo.org/record/217459/files/figure.png" pageId="1" pageNumber="278">Figs 2</figureCitation>
A–H): pale-orange. Femur slightly enlarged (
<figureCitation id="9F892A56D957FFF92D772443FB9C5A19" box="[1077,1150,1086,1111]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1939]" captionTargetBox="[252,1342,183,1873]" captionTargetId="figure@3.[236,1351,175,1914]" captionTargetPageId="3" captionText="FIGURE 2. Trilacuna hamata sp. nov., male. A, B, left palp, prolateral and retrolateral views; C – H, left embolus, prolateral (C, F), retrolateral (D, G) and dorsal (E, H) views. Abbreviations: dh = dorsal hairs; ffp = filiform, curved projection; mlo = membranous lobe; nb = narrow branch; rdh = retrolateral-dorsally hairs; sso = slightly sclerotized outgrowth. Scales: A, B, 0.2 mm; C – H, 0.1 mm." httpUri="https://zenodo.org/record/217459/files/figure.png" pageId="1" pageNumber="278">Figs 2</figureCitation>
A, B), attached to patella basally. Cymbium not fused with bulb, with scattered setae. Bulb pear shaped, about twice as long as cymbium, stout, tapering apically. Embolus system (
<figureCitation id="9F892A56D957FFF92B3024FBFD585AD1" box="[626,698,1158,1183]" captionStart="FIGURE 2" captionStartId="3.[151,250,1917,1939]" captionTargetBox="[252,1342,183,1873]" captionTargetId="figure@3.[236,1351,175,1914]" captionTargetPageId="3" captionText="FIGURE 2. Trilacuna hamata sp. nov., male. A, B, left palp, prolateral and retrolateral views; C – H, left embolus, prolateral (C, F), retrolateral (D, G) and dorsal (E, H) views. Abbreviations: dh = dorsal hairs; ffp = filiform, curved projection; mlo = membranous lobe; nb = narrow branch; rdh = retrolateral-dorsally hairs; sso = slightly sclerotized outgrowth. Scales: A, B, 0.2 mm; C – H, 0.1 mm." httpUri="https://zenodo.org/record/217459/files/figure.png" pageId="1" pageNumber="278">Figs 2</figureCitation>
C–H) complicated, with two slightly sclerotized, strongly curved outgrowth (sso), retrolateral-dorsally bordered by numerous hairs (rdh); with a filiform, long and mesially curved projection (ffp), framed by some dorsal hairs (dh) and a narrow branch (nb); base of the embolus system with a long, membranous lobe (mlo).
</paragraph>
<paragraph id="070D36D3D957FFF92985256BFE1C5BD5" blockId="1.[151,1437,258,1579]" pageId="1" pageNumber="278">
Female: As in male except as noted. Slightly larger than male. Body length 1.85; carapace 0.83 length, 0.65 width; abdomen 0.99 length, 0.61 width. Postepigastric scutum long, semicircular, not fused to epigastric scutum. Posterior spiracles connected by groove; with two strongly sclerotized, recurved arches (sar) anterior to the posterior spiracles (
<figureCitation id="9F892A56D957FFF9283025FFFE135BD5" box="[370,497,1410,1435]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Figs 4A, C</figureCitation>
).
</paragraph>
<paragraph id="070D36D3D957FFF9298525DBFA9F5865" blockId="1.[151,1437,258,1579]" pageId="1" pageNumber="278">
Female genitalia: Ventral view (
<figureCitation id="9F892A56D957FFF92B6925DBFD485BF1" box="[555,682,1446,1471]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Figs 4A, C</figureCitation>
): Posterior margin of epigastric scutum with two dark brown knobs (dkn); anterior margin of postepigastric scutum with a fold (fo); surface without external features except for a rugose surface around the epigastric furrow. Dorsal view (
<figureCitation id="9F892A56D957FFF92A062593FC295849" box="[836,971,1518,1543]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Figs 4B, D</figureCitation>
): with a nearly rectangular, sponge-like sclerite (rss), covered by a narrow, stick-like sclerite (sls), extending anteriorly, with very short apodemes (apo).
</paragraph>
</subSubSection>
</treatment>
</document>

216
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<subSubSection id="4FA86558D957FFF929D52603FEDC58F4" pageId="1" pageNumber="278" type="nomenclature">
<paragraph id="070D36D3D957FFF929D52603FE3558D6" blockId="1.[151,471,1662,1722]" box="[151,471,1662,1688]" pageId="1" pageNumber="278">
<heading id="5C4581BFD957FFF929D52603FE3558D6" bold="true" box="[151,471,1662,1688]" fontSize="11" level="1" pageId="1" pageNumber="278" reason="1">
<emphasis id="35C6EAC1D957FFF929D52603FE3558D6" bold="true" box="[151,471,1662,1688]" pageId="1" pageNumber="278">
<taxonomicName id="C0B24D50D957FFF929D52603FE9058D6" ID-CoL="7CTSR" box="[151,370,1662,1688]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="278" phylum="Arthropoda" rank="species" species="sinuosa" status="sp. nov.">
<emphasis id="35C6EAC1D957FFF929D52603FE9058D6" bold="true" box="[151,370,1662,1688]" italics="true" pageId="1" pageNumber="278">Trilacuna sinuosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EF557BAD957FFF928382603FE3558D6" box="[378,471,1662,1688]" pageId="1" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="070D36D3D957FFF929D526DCFEDC58F4" blockId="1.[151,471,1662,1722]" box="[151,318,1697,1722]" pageId="1" pageNumber="278">
(
<figureCitation id="9F892A56D957FFF929DD26DCFF0758F4" box="[159,229,1697,1722]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Figs 4</figureCitation>
E–H, 5)
</paragraph>
</subSubSection>
<subSubSection id="4FA86558D957FFF929D52694FA845968" pageId="1" pageNumber="278" type="materials_examined">
<paragraph id="070D36D3D957FFF929D52694FA845968" blockId="1.[151,1436,1769,2010]" pageId="1" pageNumber="278">
<emphasis id="35C6EAC1D957FFF929D52694FEAA594C" bold="true" box="[151,328,1769,1794]" pageId="1" pageNumber="278">
<typeStatus id="D8098871D957FFF929D52694FF30594C" box="[151,210,1769,1794]" pageId="1" pageNumber="278">Type</typeStatus>
material.
</emphasis>
<specimenCount id="11B4FD5AD957FFF928132697FE57594C" box="[337,437,1769,1794]" pageId="1" pageNumber="278" type="female">1 female</specimenCount>
<typeStatus id="D8098871D957FFF928822694FDC0594C" box="[448,546,1769,1794]" pageId="1" pageNumber="278" type="holotype">holotype</typeStatus>
(
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Ar 19421), Tam Dao National Park, Vinh Phuc,
<collectingCountry id="7FA57643D957FFF92D812694FAC7594C" box="[1219,1317,1769,1794]" name="Vietnam" pageId="1" pageNumber="278">Vietnam</collectingCountry>
, leaf litter sieving in disturbed forest (21˚30.516′N, 105˚34.263′E, altitude ca. 509 meters), Pham Dinh-Sac,
<date id="730C1013D957FFF92D9C2773FA805968" box="[1246,1378,1806,1830]" pageId="1" pageNumber="278" value="2007-05-17">17. V. 2007</date>
.
</paragraph>
</subSubSection>
<subSubSection id="4FA86558D957FFF92985274CFD6E5920" pageId="1" pageNumber="278" type="etymology">
<paragraph id="070D36D3D957FFF92985274CFD6E5920" blockId="1.[151,1436,1769,2010]" pageId="1" pageNumber="278">
<emphasis id="35C6EAC1D957FFF92985274CFEAC5904" bold="true" box="[199,334,1841,1866]" pageId="1" pageNumber="278">Etymology.</emphasis>
The specific name is Latin, meaning “curving”, and refers to the strongly winding tube in the female epigastric region (
<figureCitation id="9F892A56D957FFF928F72728FE1A5920" box="[437,504,1877,1902]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Fig. 4</figureCitation>
F); adjective.
</paragraph>
</subSubSection>
<subSubSection id="4FA86558D957FFF929852704FAE45994" pageId="1" pageNumber="278" type="diagnosis">
<paragraph id="070D36D3D957FFF929852704FAE45994" blockId="1.[151,1436,1769,2010]" pageId="1" pageNumber="278">
<emphasis id="35C6EAC1D957FFF929852704FEA359DC" bold="true" box="[199,321,1913,1938]" pageId="1" pageNumber="278">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="C0B24D50D957FFF92BD52707FBD059DC" authority="Grismado &amp; Piacentini" authorityName="Grismado &amp; Piacentini" box="[663,1074,1913,1938]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="278" phylum="Arthropoda" rank="species" species="mahanadi">
<emphasis id="35C6EAC1D957FFF92BD52707FCC659DC" box="[663,804,1913,1938]" italics="true" pageId="1" pageNumber="278">T. mahanadi</emphasis>
Grismado &amp; Piacentini
</taxonomicName>
, but can be distinguished by the numerous granulates on the sides of the carapace (
<figureCitation id="9F892A56D957FFF92BAB27E0FCD159F8" box="[745,819,1949,1974]" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="1" pageNumber="278">Figs 5</figureCitation>
A–E), the longer postepigastric scutum (
<figureCitation id="9F892A56D957FFF92C4F27E0FAB159F8" box="[1293,1363,1949,1974]" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="1" pageNumber="278">Fig. 5</figureCitation>
J), the strongly winding tube (swt) and straight, narrow sclerite (sns) in the female genital area (
<figureCitation id="9F892A56D957FFF92D3927BCFB205994" box="[1147,1218,1985,2010]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="1" pageNumber="278">Figs 4</figureCitation>
E–H).
</paragraph>
</subSubSection>
<caption id="53CD665BD954FFFA29D52706FED759A3" httpUri="https://zenodo.org/record/217458/files/figure.png" pageId="2" pageNumber="279" targetBox="[204,1389,194,1894]" targetPageId="2">
<paragraph id="070D36D3D954FFFA29D52706FED759A3" blockId="2.[151,1436,1915,2029]" pageId="2" pageNumber="279">
<emphasis id="35C6EAC1D954FFFA29D52706FEF759DE" bold="true" box="[151,277,1915,1937]" pageId="2" pageNumber="279">FIGURE 1.</emphasis>
<taxonomicName id="C0B24D50D954FFFA285E2706FE3159DF" box="[284,467,1915,1937]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="35C6EAC1D954FFFA285E2706FE3159DF" box="[284,467,1915,1937]" italics="true" pageId="2" pageNumber="279">Trilacuna hamata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EF557BAD954FFFA28992706FDCF59DE" box="[475,557,1915,1936]" pageId="2" pageNumber="279" rank="species">
<emphasis id="35C6EAC1D954FFFA28992706FDCF59DE" bold="true" box="[475,557,1915,1936]" pageId="2" pageNumber="279">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. A, habitus, dorsal view; B, habitus, lateral view; C, D, E, F, prosoma, dorsal, lateral, ventral and frontal views; G, abdomen, ventral view; H, abdomen, anterior view; I, labium and endites, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision; sdb = slightly distal branch. Scales: A, B, 0.4 mm; C–H, 0.2 mm; I, 0.1 mm.
</paragraph>
</caption>
<caption id="53CD665BD955FFFB29D52700FEBF59A1" httpUri="https://zenodo.org/record/217459/files/figure.png" pageId="3" pageNumber="280" targetBox="[252,1342,183,1873]" targetPageId="3">
<paragraph id="070D36D3D955FFFB29D52700FEBF59A1" blockId="3.[151,1436,1917,2031]" pageId="3" pageNumber="280">
<emphasis id="35C6EAC1D955FFFB29D52700FEF659DC" bold="true" box="[151,276,1917,1939]" pageId="3" pageNumber="280">FIGURE 2.</emphasis>
<taxonomicName id="C0B24D50D955FFFB285E2700FE3359DD" box="[284,465,1917,1939]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="35C6EAC1D955FFFB285E2700FE3359DD" box="[284,465,1917,1939]" italics="true" pageId="3" pageNumber="280">Trilacuna hamata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EF557BAD955FFFB289B2703FDC959DD" box="[473,555,1918,1939]" pageId="3" pageNumber="280" rank="species">
<emphasis id="35C6EAC1D955FFFB289B2703FDC959DD" bold="true" box="[473,555,1918,1939]" pageId="3" pageNumber="280">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. A, B, left palp, prolateral and retrolateral views; C–H, left embolus, prolateral (C, F), retrolateral (D, G) and dorsal (E, H) views. Abbreviations: dh = dorsal hairs; ffp = filiform, curved projection; mlo = membranous lobe; nb = narrow branch; rdh = retrolateral-dorsally hairs; sso = slightly sclerotized outgrowth. Scales: A, B, 0.2 mm; C–H, 0.1 mm.
</paragraph>
</caption>
<caption id="53CD665BD952FFFC29D526ABFCB05964" httpUri="https://zenodo.org/record/217460/files/figure.png" pageId="4" pageNumber="281" targetBox="[151,1436,193,1728]" targetPageId="4">
<paragraph id="070D36D3D952FFFC29D526ABFCB05964" blockId="4.[151,1436,1750,1834]" pageId="4" pageNumber="281">
<emphasis id="35C6EAC1D952FFFC29D526ABFEF058A2" bold="true" box="[151,274,1750,1772]" pageId="4" pageNumber="281">FIGURE 3.</emphasis>
<taxonomicName id="C0B24D50D952FFFC285B26ABFE2C58A2" box="[281,462,1750,1772]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="35C6EAC1D952FFFC285B26ABFE2C58A2" box="[281,462,1750,1772]" italics="true" pageId="4" pageNumber="281">Trilacuna hamata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EF557BAD952FFFC289626AAFDC758A2" box="[468,549,1751,1772]" pageId="4" pageNumber="281" rank="species">
<emphasis id="35C6EAC1D952FFFC289626AAFDC758A2" bold="true" box="[468,549,1751,1772]" pageId="4" pageNumber="281">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. A–D, prosoma, dorsal, lateral, frontal and ventral views; E, abdomen, anterior view; F, abdomen, ventral view; G, labium and endites, ventral view; H, abdomen, lateral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A–G, 0.2 mm; H, 0.4 mm.
</paragraph>
</caption>
<caption id="53CD665BD953FFFD29D52738FF24599B" httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="5" pageNumber="282" targetBox="[168,1406,212,1750]" targetPageId="5">
<paragraph id="070D36D3D953FFFD29D52738FF24599B" blockId="5.[151,1436,1860,2005]" pageId="5" pageNumber="282">
<emphasis id="35C6EAC1D953FFFD29D52738FEF05914" bold="true" box="[151,274,1861,1883]" pageId="5" pageNumber="282">FIGURE 4.</emphasis>
A–D,
<taxonomicName id="C0B24D50D953FFFD28142739FDE85914" box="[342,522,1860,1882]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="35C6EAC1D953FFFD28142739FDE85914" box="[342,522,1860,1882]" italics="true" pageId="5" pageNumber="282">Trilacuna hamata</emphasis>
</taxonomicName>
<emphasis id="35C6EAC1D953FFFD2B522738FD825914" bold="true" box="[528,608,1861,1882]" pageId="5" pageNumber="282">
<taxonomicNameLabel id="2EF557BAD953FFFD2B522738FD825914" box="[528,608,1861,1882]" pageId="5" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, E–H,
<taxonomicName id="C0B24D50D953FFFD2BEA2739FCBC5915" box="[680,862,1860,1883]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="sinuosa">
<emphasis id="35C6EAC1D953FFFD2BEA2739FCBC5915" box="[680,862,1860,1883]" italics="true" pageId="5" pageNumber="282">Trilacuna sinuosa</emphasis>
</taxonomicName>
s
<emphasis id="35C6EAC1D953FFFD2A2C2738FC565914" bold="true" box="[878,948,1861,1882]" pageId="5" pageNumber="282">p. nov.</emphasis>
A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm.
</paragraph>
</caption>
<caption id="53CD665BD950FFFE29D52708FA665986" httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="6" pageNumber="283" targetBox="[151,1436,196,1887]" targetPageId="6">
<paragraph id="070D36D3D950FFFE29D52708FA665986" blockId="6.[151,1436,1909,1992]" pageId="6" pageNumber="283">
<emphasis id="35C6EAC1D950FFFE29D52708FEF159C5" bold="true" box="[151,275,1909,1931]" pageId="6" pageNumber="283">FIGURE 5.</emphasis>
<taxonomicName id="C0B24D50D950FFFE285B2708FE2C59C5" box="[281,462,1909,1931]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="sinuosa" status="sp. nov.">
<emphasis id="35C6EAC1D950FFFE285B2708FE2C59C5" box="[281,462,1909,1931]" italics="true" pageId="6" pageNumber="283">Trilacuna sinuosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EF557BAD950FFFE2897270BFDC759C5" box="[469,549,1910,1931]" pageId="6" pageNumber="283" rank="species">
<emphasis id="35C6EAC1D950FFFE2897270BFDC759C5" bold="true" box="[469,549,1910,1931]" pageId="6" pageNumber="283">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. A, habitus, dorsal view; B, habitus, lateral view; C–F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C–G, I, 0.2 mm.
</paragraph>
</caption>
<subSubSection id="4FA86558D951FFFF298520EAFE675DCE" pageId="7" pageNumber="284" type="description">
<paragraph id="070D36D3D951FFFF298520EAFD995CE6" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="284">
<emphasis id="35C6EAC1D951FFFF298520EAFEBB5EFE" bold="true" box="[199,345,151,176]" pageId="7" pageNumber="284">Description.</emphasis>
Female. Completely chitinized. Body yellow-brown,chelicerae and sternum lighter, legs yellow. Habitus as in
<figureCitation id="9F892A56D951FFFF287720C1FE9E5E9B" box="[309,380,188,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="7" pageNumber="284">Figs 5</figureCitation>
A, B. Body length 2.07; carapace 0.96 length, 0.75 width; abdomen 1.19 length, 0.71 width. Carapace broadly oval, slightly elevated, dorsal surface smooth, with three rows of short hairs between eyes and posterior border of elevation of carapace; sides with numerous granulates; lateral margin rebordered, with a row of short, finely hairs and small blunt denticles.
<emphasis id="35C6EAC1D951FFFF2BD02154FD255F0E" box="[658,711,297,320]" italics="true" pageId="7" pageNumber="284">Eyes</emphasis>
six (
<figureCitation id="9F892A56D951FFFF2A43215AFCAA5F0E" box="[769,840,295,320]" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="7" pageNumber="284">Figs 5</figureCitation>
C–E), well developed, arranged in a compact group, about subequal, roundish; posterior row recurved from above, straight from front.
<emphasis id="35C6EAC1D951FFFF2D712131FB6F5F2B" box="[1075,1165,332,357]" italics="true" pageId="7" pageNumber="284">Clypeus</emphasis>
sinuous in frontal view, anterior lateral eyes separated from edge of carapace by about twice their diameter, with needle-like setae (
<figureCitation id="9F892A56D951FFFF2C2C2112FF475FE3" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="7" pageNumber="284">Fig. 5</figureCitation>
E).
<emphasis id="35C6EAC1D951FFFF298D21E9FEB95FE3" box="[207,347,404,429]" italics="true" pageId="7" pageNumber="284">Mouthparts:</emphasis>
Chelicerae straight, proximal region with one hair with elevated hair base; labium rectangular, not fused to sternum, anterior margin deeply incised (ldi) (
<figureCitation id="9F892A56D951FFFF2A6521CAFC885F9E" box="[807,874,439,464]" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="7" pageNumber="284">Fig. 5</figureCitation>
F); endites slender, distally not branched.
<emphasis id="35C6EAC1D951FFFF2C7D21C5FA795F81" box="[1343,1435,440,463]" italics="true" pageId="7" pageNumber="284">Sternum</emphasis>
(
<figureCitation id="9F892A56D951FFFF29DD21A1FF015FBB" box="[159,227,476,501]" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="7" pageNumber="284">Fig. 5</figureCitation>
F) without radial furrows between coxae, surface finely reticulate; setae sparse, light, needle-like, evenly scattered.
<emphasis id="35C6EAC1D951FFFF284B2182FE915C56" box="[265,371,511,536]" italics="true" pageId="7" pageNumber="284">Abdomen</emphasis>
: Ovoid, dorsal scutum not fused to epigastric scutum, postepigastric and epigastric scutum not fused, apodemes present, posterior spiracles connected by groove (
<figureCitation id="9F892A56D951FFFF2AE02259FC0C5C72" box="[930,1006,548,573]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="7" pageNumber="284">Figs 4</figureCitation>
E,
<date id="730C1013D951FFFF2D522259FBC85C73" box="[1040,1066,548,573]" pageId="7" pageNumber="284">5I</date>
); with two strongly sclerotized, recurved arches (sar) anterior to the posterior spiracles (
<figureCitation id="9F892A56D951FFFF2A69223AFC915C2E" box="[811,883,583,608]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="7" pageNumber="284">Fig. 4</figureCitation>
E); booklung covers (boc) large, ovoid, surface smooth (
<figureCitation id="9F892A56D951FFFF29BA2211FEDD5CCB" box="[248,319,620,645]" captionStart="FIGURE 5" captionStartId="6.[151,250,1909,1931]" captionTargetBox="[151,1436,196,1887]" captionTargetId="figure@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURE 5. Trilacuna sinuosa sp. nov., female. A, habitus, dorsal view; B, habitus, lateral view; C – F, prosoma, dorsal, lateral, frontal and ventral views; G, abdomen, anterior view; H, abdomen, lateral view; I, genital area, ventral view; J, abdomen, ventral view. Abbreviations: boc = booklung covers; ldi = labium deep incision. Scales: A, B, H, J, 0.4 mm; C – G, I, 0.2 mm." httpUri="https://zenodo.org/record/217462/files/figure.png" pageId="7" pageNumber="284">Figs 5</figureCitation>
H, J). Leg
<emphasis id="35C6EAC1D951FFFF28F52210FDE75CCA" box="[439,517,621,644]" italics="true" pageId="7" pageNumber="284">spines:</emphasis>
metatarsi I, II with 4 pairs of ventral spines each, tarsi I, II with 2 pairs of ventral spines each; legs III and IV without spines.
</paragraph>
<paragraph id="070D36D3D951FFFF298522C9FE4A5D12" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="284">
Female genitalia: Ventral view (
<figureCitation id="9F892A56D951FFFF2B7722C9FD9D5C82" box="[565,639,692,717]" captionStart="FIGURE 4. A – D" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[168,1406,212,1750]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 4. A – D, Trilacuna hamata sp. nov., E – H, Trilacuna sinuosa s p. nov. A, C, E, G, female genital area, ventral view; B, D, F, H, female genital area, dorsal view. Abbreviations: amp = anterior margin of postepigastic scutum; apo = apodemes; dkn = dark brown knobs; fo = fold; pfo = procurved fold of uterus externus; rss = rectangular, sponge-like sclerite; sar = strongly sclerotized, recurved arches; sls = stick-like sclerite; sns = straight, narrow sclerite; swt = strongly winding tube. Scales: 0.2 mm." httpUri="https://zenodo.org/record/217461/files/figure.png" pageId="7" pageNumber="284">Figs 4</figureCitation>
E, G): Uterus externus appearing as a procurved fold (pfo), ending in two dark brown knobs (dkn); anterior margin of postepigastic scutum (amp) recurved at middle; surface without external features except for a rugose surface around the epigastric furrow. Dorsal view (
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F, H): with a strongly winding tube (swt) running posteriorly, then turning backwards, ending in a straight, narrow sclerite (sns); apodemes present (apo).
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<paragraph id="070D36D3D951FFFF2985231AFE675DCE" blockId="7.[151,1437,151,896]" box="[199,389,871,896]" pageId="7" pageNumber="284">Male: Unknown.</paragraph>
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82
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<taxonomicName id="B861312215FBA129F2990AE58147FA51" genus="Ichneumonoidea" lsidName="Ichneumonoidea apterus" pageId="0" pageNumber="8151" rank="species" species="apterus">apterus</taxonomicName>
Wollaston, 1858
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Fahringer, 1930
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<paragraph id="0B31CCFEA9E0D2A90619156AC4DCAF96" pageId="0" pageNumber="8151">England</paragraph>
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<paragraph id="76CE92AB2BCD79EE780EC95AC57EDA6C" pageId="0" pageNumber="8151">Notes</paragraph>
<paragraph id="F06F27FE68C0BF603D0BF67E6C4FEBCF" pageId="0" pageNumber="8151">
Recorded as new to Britain by
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.
<bibRefCitation id="4A9E63CCD3CE332AE204FB33ABD90D9A" author="Huddleston, T." editor="Fitton, M. G." journalOrPublisher="Handbooks for the Identification of British Insects 11: ix + 159 pp." pageId="0" pageNumber="8151" title="Kloet and Hincks. A check list of British insects. Part 4: Hymenoptera" year="1978">Huddleston (1978)</bibRefCitation>
listed this as a species in need of confirmation, although, being wingless, it is straightforward to identify. There are additional British specimens in BMNH, World Museum Liverpool, Manchester Museum and Leeds City Museum.
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